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<mods:mods id="0C5B25CE606483C9263761BA7096E2E0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E59005F7957716FE4271F8A2FA45A21F">Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae)</mods:title>
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<mods:namePart id="4FA75C7AC26072636EAB4C5E86EADD96">Chen, Rong</mods:namePart>
<mods:affiliation id="F1B5753285D73C235692FBB510D17061">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, China</mods:affiliation>
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<mods:namePart id="C2F666AAC46E2A2250B26808930037D9">Zhao, Yi</mods:namePart>
<mods:affiliation id="A95D81597389AEE868851E576B905DD8">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, China &amp; Guangdong Academy of Forestry, 233 Guangshan 1 st Road, Guangzhou 510520, China</mods:affiliation>
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<mods:namePart id="AB194A5832B4432604DE39F74B7F7CA6">Golovatch, Sergei</mods:namePart>
<mods:affiliation id="200DE13975BD8C2DCF86080A14A56611">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart id="D37DD67D55E59CC4B986943F8FFF5617">Liu, Wei-Xin</mods:namePart>
<mods:affiliation id="3C7B4E471F00873F3FD1F026CC7A8539">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, China</mods:affiliation>
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<treatment id="1E81187968E0580FA7AE94DD1EAC22E7" ID-DOI="http://doi.org/10.5281/zenodo.14513059" ID-Zenodo-Dep="14513059" ID-arpha="1E811879-68E0-580F-A7AE-94DD1EAC22E7" ID-zoobank="https://zoobank.org/A91F4660-B1BD-46C3-BE71-FF0FEAB0427E" LSID="urn:lsid:plazi:treatment:1E81187968E0580FA7AE94DD1EAC22E7" LSID-ZBK="urn:lsid:zoobank.org:act:A91F4660-B1BD-46C3-BE71-FF0FEAB0427E" httpUri="http://treatment.plazi.org/id/1E81187968E0580FA7AE94DD1EAC22E7">
<subSubSection id="E974031A9DD2F2DE1EF198269804973C" type="nomenclature">
<paragraph id="A06DB24FCE7A2CB14596047E1F59D840">
<heading id="9477A980D705F78E42E152EA60118449" reason="label">3.3. 2.</heading>
<taxonomicName id="9BC7A31B5EC3F963AC7DBF90C66144C6" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="tiankeng" status="sp. nov.">
<emphasis id="85BA0614BCDC5D4BFCBFDDBB17313DBF" italics="true">Skleroprotopus tiankeng</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3A2DFDA6CF6B2A2A6D89C1D8664E5F32" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C9CAA9AA414CCD1EC9615B7101FD380D" type="description">
<paragraph id="E08C044275DC823551A5E7C29EC79E2A">
<figureCitation id="E2B8809B01C73DD26658D8DCC5DF2E14" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov." figureDoi="10.3897/asp.82.e136751.figure2" httpUri="https://binary.pensoft.net/fig/1201329">Figures 2 E</figureCitation>
,
<figureCitation id="5D979FA2FA2C37ADD8537A0E89BDAF94" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm." figureDoi="10.3897/asp.82.e136751.figure3" httpUri="https://binary.pensoft.net/fig/1201330">3 C</figureCitation>
,
<figureCitation id="A1C54A00ED67F29790E5D2317C85E676" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EELBG" type="material">
<paragraph id="7516498CA19E4D6BAFA51E0D9D0B9B75">
<heading id="2E356082D495F717C12FF4DFB3837B7B" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="3B624251732D7C4F2822932F54244FF0">
<materialsCitation id="D30B3A2F47A85DD762916799B67F2C76" collectingDate="2017-09-06" collectionCode="SCAU" collectorName="Tian Mingyi &amp; Huang Sunbin &amp; Wang Dianmei &amp; Yin Haomin" country="China" county="Hanzhong City" elevation="850" latitude="32.827198" location="Cave Xigoutiankeng Dong" longLatPrecision="1" longitude="107.02716" municipality="Nanzheng District" specimenCount="1" stateProvince="Shaanxi" typeStatus="holotype">
<typeStatus id="5B32EABF75688F2C634E2460E8BD6515" type="holotype">Holotype</typeStatus>
male (
<named-content id="6DB53820314471F6B6168513D4172C6B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Zoological Collection of the South China Agricultural University">
<collectionCode id="F72BC7CF77014A7A404A17F6A04E1DE2">SCAU</collectionCode>
</named-content>
),
<collectingCountry id="803A2560C56E480D49B58461E66FFB91" name="China">China</collectingCountry>
,
<collectingRegion id="975E0138EA4369E8596492A35F872ED5" country="China" name="Shaanxi">Shaanxi Province</collectingRegion>
,
<collectingCounty id="84E286EDA41E2952CF82D4DA1D197863">Hanzhong City</collectingCounty>
,
<collectingMunicipality id="1928FF29F456007E9E4D55B37F67135D">Nanzheng District</collectingMunicipality>
,
<location id="FA780FEB2752913620287BC77DEFD9CC" LSID="urn:lsid:plazi:treatment:1E81187968E0580FA7AE94DD1EAC22E7:FA780FEB2752913620287BC77DEFD9CC" country="China" county="Hanzhong City" latitude="32.827198" longLatPrecision="1" longitude="107.02716" municipality="Nanzheng District" name="Cave Xigoutiankeng Dong" stateProvince="Shaanxi">Cave Xigoutiankeng Dong</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERLBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.027158,32.827197]}">
<geoCoordinate id="50585E65AE2E639E2ED3AA1985DF86F0" degrees="32" direction="north" minutes="49" orientation="latitude" precision="1" seconds="37.91" value="32.827198">32 ° 49 ′ 37.91 ′′ N</geoCoordinate>
<geoCoordinate id="BAF86C93DCE4F10C9E24C766A5127E10" degrees="107" direction="east" minutes="1" orientation="longitude" precision="1" seconds="37.77" value="107.02716">107 ° 1 ′ 37.77 ′′ E</geoCoordinate>
</named-content>
,
<quantity id="7CDCAFF456E9BC7AE39A58E1F3C13C88" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">
<elevation id="546EC19B7D2EDC2089F5F9AB62FD6D45" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<date id="68FCFE3BF8B13A0D3B1F735458BD34B4" value="2017-09-06">
<collectingDate id="3A3976AA0B8A3B1825247D7F360278DB" value="2017-09-06">6. IX. 2017</collectingDate>
</date>
,
<collectorName id="E493C2A1C763E65B6AA00D4A1937F5B7">Tian Mingyi</collectorName>
,
<collectorName id="60A86EF8896D1096449E3FC4AAA158C6">Huang Sunbin</collectorName>
,
<collectorName id="DA4B9BFC4E7DCA3D956D354BCC226356">Wang Dianmei</collectorName>
and
<collectorName id="26ABA785F6B414EBDFFDE71D3709252C">Yin Haomin</collectorName>
leg.
</materialsCitation>
<typeStatus id="11F53876947DD1BF6005F118CF064470" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="C8261F8937F1FE3D6CC7F4C8C566DD33" count="4" type="male">4 males</specimenCount>
,
<specimenCount id="D2CA9ED31B0014D9B05BECD1A3D084D3" count="8" type="female">8 females</specimenCount>
(
<named-content id="F9315BF16D6484B15E87D340292829F6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Zoological Collection of the South China Agricultural University">
<collectionCode id="8A3F364102022336065B89A39576636B">SCAU</collectionCode>
</named-content>
), same data as for
<typeStatus id="B3D1AF8FAAD9328FBD17F3FCE4189D7A" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1LBG" type="diagnosis">
<paragraph id="44080B7435F9AC54F9A9CA9D855410AC">
<heading id="7E7D90464DE56768BA42D846F1F28281" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="91819F46341AC3EA87F23AB545004B14">
Differs from congeners mostly based on the following combination of characters: (1) penis much longer than coxae 2 (Fig.
<figureCitation id="DF60D2C11C58820B094F7E800B68D84A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 I</figureCitation>
); (2) male leg 7 with a very small coxal process, lower than the telopodite (Fig.
<figureCitation id="9C5AB9BDFDF994F73E28022A13828352" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 E</figureCitation>
); (3) anterior gonopod (Fig.
<figureCitation id="612BF64318F040CC96E155B58E512B1E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 G, H</figureCitation>
) with a large and foot-shaped like coxal process, carrying a broadened membranous lobe, outer margin with a triangular protrusion, and a relatively thick flagellum. — In addition, this new species differs from all other species analyzed in a&gt; 10.3 % uncorrected
<emphasis id="4F0FF37518A3D13906B9B785C382F861" italics="true">p</emphasis>
- distance of the
<abbrev id="ABBRID0EOMBG" xlink_title="Protein-coding gene sequences">COI</abbrev>
barcoding gene.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESMBG" type="etymology">
<paragraph id="8CB7E0540549EC71D0E779E8E28C9987">
<heading id="C3DE7A6082968C7C0592E69B82DA638B" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="7D2B159C902286C40D73183B5C6A2A5F">
The species is named after its
<typeStatus id="59BF3A09132A17ED8E69BEF4E0673F1D">type</typeStatus>
locality that is the Hanzhong sinkhole group, a geological wonder composed of multiple heavenly pits. “ Tiankeng ” in Chinese “ 天坑 ”, a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXMBG" type="description">
<paragraph id="F9037E56BADBB10DBA99FF83093F431E">
<heading id="9091345A61E97B8474A638F82EFA6A43" reason="title">Description.</heading>
</paragraph>
<paragraph id="9FADE27AB35474521C6208D14FC68333">
Length of both sexes ca 29.0–
<quantity id="929C9E5F7E3FAB1D5F9180C6709142E4" metricMagnitude="-2" metricUnit="m" metricValue="3.8899999999999997" unit="mm" value="38.9">38.9 mm</quantity>
, width
<quantity id="71A9FC31A998ADB603ED1C358E73D392" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9–2.2 mm</quantity>
, midbody height
<quantity id="44E370B18CE9B2A394EFEDAAE71B53F7" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8–2.1 mm</quantity>
, body with 49–63 podous + 1 apodous rings + telson. Natural coloration generally yellow to brownish. Antennae and legs light yellowish (Fig.
<figureCitation id="0ED225B04EE617D952C4914B9085C9E2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov." figureDoi="10.3897/asp.82.e136751.figure2" httpUri="https://binary.pensoft.net/fig/1201329">2 E</figureCitation>
). Eye patches transparent, subtriangular, arranged in 4–6 irregular rows, altogether about 18–27 (males) or 15–40 (females) ommatidia per eye patch.
</paragraph>
<caption id="F754B0DAB9651EE90F9744675650A4FA" ID-DOI="10.3897/asp.82.e136751.figure2" ID-arpha="92342405-B89B-5064-91C4-84083B192E3F" httpUri="https://binary.pensoft.net/fig/1201329" startId="F2">
<paragraph id="CA0EFE7ED15B647E5EB178E1EBA4AC4B">
<label id="0A7F3F61A5B436009B728CAF7A2E9E1F">Figure 2.</label>
</paragraph>
<paragraph id="7590B68AAE9669815FD4347B35F64269">
Pictures of live animals, photos by Prof. Mingyi Tian from
<named-content id="0431831B9846052AAB849731C1A621A0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Zoological Collection of the South China Agricultural University">SCAU</named-content>
.
<emphasis id="01C7134B9DEC18DF3A2FE7AE86C2AD0D" bold="true">A</emphasis>
<taxonomicName id="77352919CAD36BD314113CD0AA27CDAA" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="change" status="sp. nov.">
<emphasis id="6F75EFB065F0860C9ED0EC7477A31096" italics="true">Skleroprotopus change</emphasis>
</taxonomicName>
<emphasis id="D307B627D1688EC07E1FFFF6E00419C1" bold="true">
<taxonomicNameLabel id="DFB89FA160081E63052CD256C5FD481D" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="7B0D6A596B46ADC3BA5072A595F2B6EB" bold="true">B</emphasis>
<taxonomicName id="DD5960BB89D23B6AB7497EF508ED78B8" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="megistus" status="sp. nov.">
<emphasis id="6FDA33736620F791F630049A77D38F45" italics="true">Skleroprotopus megistus</emphasis>
</taxonomicName>
<emphasis id="F71B29FC9BC7749C9E134C71492F39C8" bold="true">
<taxonomicNameLabel id="C10EA198B6014D6E7C1828F51200A4F1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="D004456A5E395C69A50CAA7B618E973C" bold="true">C</emphasis>
<taxonomicName id="439C5AF2F12CFF1D8A369A57DDE6B380" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="longissimus" status="sp. nov.">
<emphasis id="EFD346A4289A40ED5AC6D0A83A63EB97" italics="true">Skleroprotopus longissimus</emphasis>
</taxonomicName>
<emphasis id="0E2BC2741FE36CE83107D5DF3297AF25" bold="true">
<taxonomicNameLabel id="F2AC7BF79000AFCDEDD0ADE94D364609" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="33E37A0E47FBF3930F102A538BEC6E8C" bold="true">D</emphasis>
<taxonomicName id="24B62CE105F749C6683E7A403DAD8091" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="genjudi" status="sp. nov.">
<emphasis id="29C38214658416208C46D7DD30C352B6" italics="true">Skleroprotopus genjudi</emphasis>
</taxonomicName>
<emphasis id="E4DCBBA7597B6D2339601E764CBEB296" bold="true">
<taxonomicNameLabel id="C23EE3D84CA64BE7F2452F6BC5C7E09A" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="A9711B1F3B0AAFF6FCA8504FEA26A8B9" bold="true">E</emphasis>
<taxonomicName id="B8BA71A16FA30267016E900DF2AC1E97" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="tiankeng" status="sp. nov.">
<emphasis id="7EA16E60C896580472F96922606D59F8" italics="true">Skleroprotopus tiankeng</emphasis>
</taxonomicName>
<emphasis id="0321703A857CDC2F4E3E7657ED69B5AA" bold="true">
<taxonomicNameLabel id="E91898A05C5DE245DC1E93EE4B53A7EF" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="504AB531DED88D07DEBFB959FEA54BEF" bold="true">F</emphasis>
<taxonomicName id="57CE207D4257DAF3BD1DCF2E8E932660" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="laiyuanensis" status="sp. nov.">
<emphasis id="0B1E044BF1730C4F5FFBBE52CE0B1F33" italics="true">Skleroprotopus laiyuanensis</emphasis>
</taxonomicName>
<emphasis id="4BA925A97EDF9086E609722ED0FBA490" bold="true">
<taxonomicNameLabel id="C68B343DF6114C94D947FEE29496DDCD" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="432FE7476082490F1D5F8F31A0B2A883" bold="true">G</emphasis>
<taxonomicName id="53A5725F1C3770F52B42F59094DA3019" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="incisodentatus" status="sp. nov.">
<emphasis id="153AE9CE27EBB39F38B7C9272F0A2326" italics="true">Skleroprotopus incisodentatus</emphasis>
</taxonomicName>
<emphasis id="1F6869A65CC642BDE4A67B6E8A851BC7" bold="true">
<taxonomicNameLabel id="FA6CD7CC7FA9611444A428CEC6A87C85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</caption>
<paragraph id="FC87E0E8648C81311FF88F49B609B0C4">
Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 + 2 supra-labral and 8 + 8 labral setae. Antennae long, reaching behind to anterior of ring 4 when stretched dorsally (Fig.
<figureCitation id="0B76F7C5E037DB0336E12B5138D3866B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm." figureDoi="10.3897/asp.82.e136751.figure3" httpUri="https://binary.pensoft.net/fig/1201330">3 C</figureCitation>
). In length, antennomeres 3&gt; 5&gt; 2&gt; 4 ≈ 6&gt; 1 ≈ 7 (Fig.
<figureCitation id="E96BD5CE6B2621EC03225A8D69C1D53E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 C</figureCitation>
). Antennomeres 5, 6 and 7 each with a distal corolla of sensilla basiconica. Mandibular stipes enlarged, with two small lobes in males, well-rounded in females. Gnathochilarium with at least 3–8 setae on each lamella lingualis (
<emphasis id="AD3AF4081EE62FFF6BB244DA433DB07A" bold="true">ll</emphasis>
) in males; promentum (
<emphasis id="9B71ED8526D1B5A58E7FA8B4D342FA04" bold="true">pr</emphasis>
) columnar in males, anteriorly swollen and tubular in shape (Fig.
<figureCitation id="93A66CEF51331249955B85345EB11803" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 B</figureCitation>
), vs a broad conical
<emphasis id="44698B8FAC28B25CB95FA8FD9F550A55" bold="true">pr</emphasis>
in females (Fig.
<figureCitation id="9EE4518181EA9BED2BEE3B129B42CF1B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 A</figureCitation>
).
</paragraph>
<caption id="CCE33FCFF4D1AF95F9EA167EABF636F9" ID-DOI="10.3897/asp.82.e136751.figure3" ID-arpha="D7C8E060-091F-51A1-904E-452ADBBC9093" httpUri="https://binary.pensoft.net/fig/1201330" startId="F3">
<paragraph id="765142D7ECFA7ECBDBE9E0F835D9496E">
<label id="C30DF3957CAC3497D1E7B50919EDE31D">Figure 3.</label>
</paragraph>
<paragraph id="1C7DC4337D299EA57E274B80AC310840">
Habitus of some
<taxonomicName id="210A23A9639E6FE45AC138BD4FCAD230">
<emphasis id="309F70DDE2C934A131836E4553FB16C0" italics="true">Skleroprotopus</emphasis>
species
</taxonomicName>
in lateral view.
<emphasis id="8726E511307B1BF83BC6939A754525C7" bold="true">A</emphasis>
<taxonomicName id="58A4D7484875CF2629FDC67E83917CCB" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="penglai" status="sp. nov.">
<emphasis id="976465EAD18E99D147F59D9546E910F8" italics="true">Skleroprotopus penglai</emphasis>
</taxonomicName>
<emphasis id="A13350E530D049359B7BC65C07CD92BF" bold="true">
<taxonomicNameLabel id="DB726A7679630E89659A77DA7E232C9B" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="9DDA26ACF6352D4A514C0BA614A60094" bold="true">B</emphasis>
<taxonomicName id="1EC00E37B707E6E31336321FCAA307F5" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="longiflagellatus" status="sp. nov.">
<emphasis id="4E85F1652A941AC75ECD3F5DCAE390C6" italics="true">Skleroprotopus longiflagellatus</emphasis>
</taxonomicName>
<emphasis id="447ECD6A556D6F0EA969233CBA6FFFC4" bold="true">
<taxonomicNameLabel id="C9A2B72E15520C5CC1B0FE94C7E88831" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="EAB47C6D916D8001D75298E8131A5FD0" bold="true">C</emphasis>
<taxonomicName id="E58F27CFCF75A3097C283DDEC4B74FDA" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="tiankeng" status="sp. nov.">
<emphasis id="AB1B2789D0C2F37DA015424CDC534E9F" italics="true">Skleroprotopus tiankeng</emphasis>
</taxonomicName>
<emphasis id="C9D441BB576976807A45C43908B5FE6A" bold="true">
<taxonomicNameLabel id="E81FA0DCF95D086C62C85E20426EEBA6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="71531F892E4DEAF6B287BE6D871D04E0" bold="true">D</emphasis>
<taxonomicName id="E028B382C7C84939AE2E869A7089407A" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="ampullaceus" status="sp. nov.">
<emphasis id="E59760CA45E13D6795016ED3DA7AEA9A" italics="true">Skleroprotopus ampullaceus</emphasis>
</taxonomicName>
<emphasis id="EF6D680E9CFB63655DD6F21A0EA71B36" bold="true">
<taxonomicNameLabel id="1F4330D71A4EB158657D125A0411502A" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="CD56F03A7E9ACDC7851168737E5CFE20" bold="true">E</emphasis>
<taxonomicName id="EA6EB8E00338D570A0C46ECFB86F35AD" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="conicus" status="sp. nov.">
<emphasis id="120E3A01404F5AFB6360F3905599221F" italics="true">Skleroprotopus conicus</emphasis>
</taxonomicName>
<emphasis id="0F39B07FD3485A74D8D291044587412B" bold="true">
<taxonomicNameLabel id="416B6A446B0936918E14790AEDF5DE71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="DFE31BDA2E878D7A9BFE583B3C206449" bold="true">F</emphasis>
<taxonomicName id="80105650BA35F1AD3F96A2EDFB802F0C" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="yutiantianae" status="sp. nov.">
<emphasis id="57A7D1683A78C58E72507EA60EB57B6C" italics="true">Skleroprotopus yutiantianae</emphasis>
</taxonomicName>
<emphasis id="CA99F447BFB3E12E772D6619C9FB4355" bold="true">
<taxonomicNameLabel id="412816367BC3F44165E087EA67CFF771" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B127334759C1882B482B241A696B0E65" bold="true">G</emphasis>
<taxonomicName id="DB1A5C2A74A31DAA421D2248ABFF33D1" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="multistriatus" status="sp. nov.">
<emphasis id="41609084FA50DD6E567C8B460E1C3779" italics="true">Skleroprotopus multistriatus</emphasis>
</taxonomicName>
<emphasis id="6E3A3E4493363C1AC6B7FEE84ACED7C7" bold="true">
<taxonomicNameLabel id="9C4F669DE140BF7DE1DA21712E8DBE8D" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="C2870A3616699E4B3E94529E19A224C6" bold="true">H</emphasis>
<taxonomicName id="DCCBE995D9A1542DBAAE6A914FAFFAE2" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="change" status="sp. nov.">
<emphasis id="F0400664A3A4754493443C4CADE032D2" italics="true">Skleroprotopus change</emphasis>
</taxonomicName>
<emphasis id="AF9B24565B66B8AFEB9A6965AB0F63B4" bold="true">
<taxonomicNameLabel id="7CC488DEEE409B8FB2E61645BB4BCF86" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars
</emphasis>
: 5.0 mm.
</paragraph>
</caption>
<caption id="6BC36C51C19057E5BB88D09C047247BD" ID-DOI="10.3897/asp.82.e136751.figure4" ID-arpha="0DFE8BD2-9A18-577E-A145-65690D111BFE" httpUri="https://binary.pensoft.net/fig/1201331" startId="F4">
<paragraph id="28C301055442D9A686F4774F67F60FAA">
<label id="86D52F7347CA2ACC82B50B527EABDC6F">Figure 4.</label>
</paragraph>
<paragraph id="3021689C8886B504380A06A39AA99843">
<taxonomicName id="475D338AD0A6887994F1926F1A00C6DA" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="yutiantianae" status="sp. nov.">
<emphasis id="10F5C3D394803DE29C2EE2B7380EB813" italics="true">Skleroprotopus yutiantianae</emphasis>
</taxonomicName>
<emphasis id="D6B18F51FBF936E17F1A4A24E611D187" bold="true">
<taxonomicNameLabel id="1F9C329AC251E067CFBBA1701B94E58D" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis id="F3F4DDD491524E6A207AA50A7FE5757A" bold="true">A</emphasis>
gnathochilarium;
<emphasis id="D589474381204FF37FABE1681C01A75C" bold="true">B</emphasis>
head and collum, dorsal view;
<emphasis id="5EE0DA21F288C5506C3E9A8E7356C724" bold="true">C</emphasis>
left antenna, lateral view;
<emphasis id="098ED8257E721A90A46E1BD060E67278" bold="true">D</emphasis>
legs 1, anterior view;
<emphasis id="BDF0CA1FD6933A050FC0763D0B754522" bold="true">E</emphasis>
legs 7, anterior view;
<emphasis id="0AA209534FF083FA8ED0D15AE5F5B4CF" bold="true">F</emphasis>
legs 2 with penis, posterior view;
<emphasis id="C43E51AE4278845B450A7EA10BD34458" bold="true">G</emphasis>
posterior gonopods, anterior view;
<emphasis id="981EDAAD4016975B93E21D16787C8E03" bold="true">H</emphasis>
anterior gonopods, anterior view;
<emphasis id="12FCDD9EAA2DB310D4D52B6A087847D6" bold="true">I</emphasis>
anterior gonopods, posterior view.
<emphasis id="0D0C33C15D4441FE9EAA35241F859DB7" bold="true">Abbreviations</emphasis>
: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; dl, dividing line; f, flagellum; ll, lamella lingualis; ml, membranous lobe; pn, penis; pr, promentum; t 1 – t 5, telopoditomeres 1–5; te, telopodite.
<emphasis id="80934AD0CA1E9D11D21F1BD2B5AE111D" bold="true">Scale bars</emphasis>
: 0.2 mm.
</paragraph>
</caption>
<caption id="ABF3B039BF776F6DFC561CFEE4E670D2" ID-DOI="10.3897/asp.82.e136751.figure5" ID-arpha="DAE82C5E-E8E6-54E1-800F-A341615393B9" httpUri="https://binary.pensoft.net/fig/1201332" startId="F5">
<paragraph id="F2074BDAB1D9B5E1DA8CC9FE89D45110">
<label id="A42EC9B14648FBA051A5005EE8E15462">Figure 5.</label>
</paragraph>
<paragraph id="288BEB51A74533284C4B418359F5397E">
<taxonomicName id="D690A4DB6BEBB3EF2C2C82C1A1AE19C2" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="tiankeng" status="sp. nov.">
<emphasis id="2A65B292BB9E30D84D3476E6919AE4DC" italics="true">Skleroprotopus tiankeng</emphasis>
</taxonomicName>
<emphasis id="E0F462D30803DE42D1BB51526705828D" bold="true">
<taxonomicNameLabel id="8514745AF2833E2DF5A5476DD674E82A" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
<emphasis id="98F470E2C0D1290796355B238202132B" bold="true">A</emphasis>
,
<emphasis id="1F5FC4D690916EFB458958B64A9B6BCC" bold="true">J</emphasis>
female paratype,
<emphasis id="686B3B3E01ADB51E3C90CBE3B93E702B" bold="true">B</emphasis>
–
<emphasis id="13F1A3E5CA651EFFDF28AEF5ED8E0D77" bold="true">I</emphasis>
male paratype.
<emphasis id="DD8049455A84A4C019B4A4D396B1A1DF" bold="true">A</emphasis>
,
<emphasis id="F283C0BBE90FFDE5AB4E2B05D153B24B" bold="true">B</emphasis>
gnathochilarium;
<emphasis id="116423D7CAAAC6BE94B508326E1F6AFA" bold="true">C</emphasis>
right antenna, lateral view;
<emphasis id="F873402A49DA7525CE8D2900F1E42BBF" bold="true">D</emphasis>
legs 1, anterior view;
<emphasis id="84056804E6451A8FD33F0F8E9654C78E" bold="true">E</emphasis>
legs 7, anterior view;
<emphasis id="E203682888C1DEC74C2F5FD6780D0374" bold="true">F</emphasis>
posterior gonopods, posterior view;
<emphasis id="BFA2E07F83B09405B6C5DC4FEEAD58A0" bold="true">G</emphasis>
anterior gonopods, anterior view;
<emphasis id="578379AFFDAF3889A7CB6D71EA6CC1FF" bold="true">H</emphasis>
anterior gonopods, posterior view;
<emphasis id="01B5778B3210AB56BA5B8E22DBAF3A37" bold="true">I</emphasis>
legs 2 with penis, posterior view;
<emphasis id="84D40233BEB328B99702C77FB8F75F5E" bold="true">J</emphasis>
right vulva, lateral view.
<emphasis id="6739F6D16186B0723E879FB76099DD72" bold="true">Abbreviations</emphasis>
: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite.
<emphasis id="6BA90DFD90D29CE4BC7F5DDE95E7562E" bold="true">Scale bars</emphasis>
: 0.2 mm.
</paragraph>
</caption>
<paragraph id="08910B8BB7D20281CF8C05662CB68887">
Collum with marbled brownish spots and 8–10 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona with 3 irregular subtransverse striae and metazona with 13–24 longitudinal striae laterally (Fig.
<figureCitation id="6DBA4681E9C8D1E2C45BBA936CF1B15C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm." figureDoi="10.3897/asp.82.e136751.figure3" httpUri="https://binary.pensoft.net/fig/1201330">3 C</figureCitation>
). Suture dividing pro- and metazona rather narrow, a nearly regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites.
</paragraph>
<paragraph id="DF539963C8D4A2D7A06491C95F0174A9">Epiproct slightly protruding caudad, posterior margin with 6 + 6 setae. Paraprocts convex, with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae.</paragraph>
<paragraph id="9C67F27A7E1C40C3168B3E2DAA1DF116">
Legs long and slender, about 1.4 × as long as midbody height. Male leg-pair 1 (Fig.
<figureCitation id="88286B06A6775CDD17EC5D75718DACB6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 D</figureCitation>
) hypertrophied, 5 - segmented, strongly curved anteriad. Coxa (
<emphasis id="C57AAA19976A0058EE01BE5890264D96" bold="true">cx</emphasis>
) and telopoditomeres 1 and 2 (
<emphasis id="6302A300074E45537D414D040F4C9A9F" bold="true">t 1</emphasis>
,
<emphasis id="8250C6DADE2A81C3D61010605086870F" bold="true">t 2</emphasis>
) each with few long setae, others mostly short. Telopoditomere 3 (
<emphasis id="379D94739EDD64093C2A7405A71D5F36" bold="true">t 3</emphasis>
) longest, about equivalent in length to the sum of the
<emphasis id="5CD6F2A98668216A64E23B6ED1467F2C" bold="true">cx</emphasis>
and
<emphasis id="46366E5CC09DDBB26A67A93E7EA20FA8" bold="true">t 1–2</emphasis>
; Telopoditomere 4 (
<emphasis id="1D68B15AC71769541E36CB0DFAF4195C" bold="true">t 4</emphasis>
) irregularly shaped, with several long setae subapically. Male leg-pair 2 (Fig.
<figureCitation id="EAA7BF417BE9223AA195673EBB63692A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 I</figureCitation>
) reduced in size; penis (
<emphasis id="D49FBD73B52424DE7F0AD41272F9FBF7" bold="true">pn</emphasis>
) pyramid-shaped, much longer than coxae (
<emphasis id="0CF3A40914B5708171FAD36CC6CAC4AE" bold="true">cx</emphasis>
), distal part with two long setae (Fig.
<figureCitation id="01711167DF11A18811FCCE9D4DB99ABA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 I</figureCitation>
). Male leg-pair 7 (Fig.
<figureCitation id="AF3D9909C06809BE69FAA882D131DA8D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 E</figureCitation>
) reduced, with 3 - segmented telopodites, telopoditomere 2 (
<emphasis id="2B9047EEEA26EB7EA6B2B9EE11E65FBA" bold="true">t 2</emphasis>
) largest and setose subapically; coxal process (
<emphasis id="0FA56832A957AD47B6A01AD2E2830C70" bold="true">cp</emphasis>
) small, much lower than telopodite, densely setose at mesal margin.
</paragraph>
<paragraph id="179CFD6E64350A693E7A2D6AD5BCDA72">
Anterior gonopods (Fig.
<figureCitation id="2A7820BDB8FCDB1E9F4E20CEC1FE1084" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 G, H</figureCitation>
) with a large, foot-shaped like coxal process (
<emphasis id="F09F9E0B78EDCC5CFAC1FFAF432BCDE5" bold="true">cp</emphasis>
) bearing with a long row of short villi in anterior view, posteriorly carrying with a broadened, and high membranous lobe (
<emphasis id="3024D5EBF007EE41A4885B19263E8F86" bold="true">ml</emphasis>
), slightly shorter than
<emphasis id="8836E26E7BF2F2E4E5309F0BFB3A99BE" bold="true">cp</emphasis>
, middle outer margin of
<emphasis id="F2F46E333E7A5F1B6D4993F78CDDDB7A" bold="true">ml</emphasis>
with a triangular protrusion. Flagellum (
<emphasis id="97C58C17715528A30787BF33FFF5433B" bold="true">f</emphasis>
) very long and relatively thick, distally spinulose-villose. Telopodite (
<emphasis id="D0227FFB8DA1A6A65984AAE37BDFFB82" bold="true">te</emphasis>
) about half as long as
<emphasis id="8BB4A41A05B256D9E3939D2B0A762273" bold="true">cp</emphasis>
, with long setae apically, laterobasally with a minute remnant (
<emphasis id="1A0C5FED4595CD93AFC5C58CD5240CA6" bold="true">r</emphasis>
) of a second podomere.
</paragraph>
<paragraph id="A0EC451C15BA666939EE23C5F4A5E8DD">
Posterior gonopods (Fig.
<figureCitation id="C96876DC091CFADCBB113498A8440BDA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 F</figureCitation>
) erect and stout, deeply branched at tip, anterior branch smooth, with a pointed tip, higher than a densely setose posterior one.
</paragraph>
<paragraph id="D42650539A51DA20842DC698004B092D">
Vulva (Fig.
<figureCitation id="FF8D5E4AD4F40FEF1792F84D3E7E36E1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure5" httpUri="https://binary.pensoft.net/fig/1201332">5 J</figureCitation>
) mostly wrapped inside a membrane, characteristic in shape, strongly elongated; operculum (
<emphasis id="E7830720A5C9AD457045D705D190199E" bold="true">op</emphasis>
) longer than bursa, with two parallel rows of short setae on anterior and posterior surfaces, and with at least 10 long distal setae.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E22BG" type="remarks">
<paragraph id="86666822D57E973D51E23F588179894F">
<heading id="16ED5CE6C12802285C306A7EDB1F004B" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="4BC75A864306FA9D526A35608249C428">
The geological relic resources of Sinkhole Group at Hanzhong are mainly distributed within Xiaonanhai Town of Nanzheng District, Luojiaba Town of Xixiang County, Chanjiayan Town of Ningqiang County, and Sanyuan Town of Zhenba County, with a total area of about
<quantity id="CA908C1685C8DAC9D067C59F43005910" metricMagnitude="6" metricUnit="m" metricValue="5.019" unit="km" value="5019.0">5019 km</quantity>
<superScript id="56336645B0CB749E2962F8081744497F">2</superScript>
. This new species lives in Xigoutiankeng Dong of Xiaonanhai Town, a cave that is about to be developed to the public. Based on the unpigmented body, transparent eyes, as well as long legs,
<taxonomicName id="45D621AC3F94FF885F9A28052B1D1B95" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="tiankeng" status="sp. nov.">
<emphasis id="26A186F59FE3292643EF4BC0766B4683" italics="true">S. tiankeng</emphasis>
</taxonomicName>
<emphasis id="8A820D582704B50E5DC34E06A66E980A" bold="true">
<taxonomicNameLabel id="590FF3D6EA23E3E291D393929C6FE836" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is considered a troglobite.
</paragraph>
</subSubSection>
</treatment>
</document>