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<document id="528AA256B46F8F4FCDC5011803C3F1A7" ID-DOI="10.11646/phytotaxa.312.1.8" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725577053966" checkinUser="felipe" docAuthor="Wilding, Nicholas" docDate="2017" docId="BF1287B4FF99FFCAF6CFFEAE2A78FCF6" docLanguage="en" docName="phytotaxa.312.1.8.pdf" docOrigin="Phytotaxa 312 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.312.1.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Entosthodon heddersonii N. Wilding 2017, sp. nov." docType="treatment" docVersion="1" lastPageNumber="105" masterDocId="432BFFCCFF98FFC8F647FF832800FFB7" masterDocTitle="Three new species of Entosthodon Schwägr. (Bryopsida, Funariaceae) from sub-Saharan Africa" masterLastPageNumber="110" masterPageNumber="103" pageNumber="104" updateTime="1725586680830" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5C26D053FFBAD34201EA58E790350A75">Three new species of Entosthodon Schwägr. (Bryopsida, Funariaceae) from sub-Saharan Africa</mods:title>
</mods:titleInfo>
<mods:name id="2B962EDA9325C8CF61B9C7E1D52D7A2C" type="personal">
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<mods:roleTerm id="6242B1F41E1E808D3A9CC5A1678BDC6C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A9A0C5F2F33E367B4CD57A6C48FE49B">Wilding, Nicholas</mods:namePart>
<mods:affiliation id="112C8FE4D5FE2EBB29438E9FB9D2B679">Université de La Réunion, UMR PVBMT, Pôle de Protection des Plantes, Pôle Forêt, Saint-Pierre, Ile de La Réunion. &amp; Department of Biological Sciences, University of Cape Town, Private Bag X 3, 7701 Rondebosch, South Africa.</mods:affiliation>
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<mods:title id="EAE5483980486F4E200DF144A24A003F">Phytotaxa</mods:title>
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<mods:part id="4B45A973567C33D69448C35FCD7D55CD">
<mods:date id="4AA32A9650147C46DDE198579399C072">2017</mods:date>
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<mods:number id="9E1B55A7837AC388345D90A7B4A918E8">2017-07-04</mods:number>
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<mods:number id="3C208A31516DE9C27D2B439BE419FDF7">312</mods:number>
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<mods:url id="253534C2A82790CAD46C421FD4A95C1E">http://dx.doi.org/10.11646/phytotaxa.312.1.8</mods:url>
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<mods:classification id="2BD45763575983E42C09D6D38CBAE348">journal article</mods:classification>
<mods:identifier id="36C3727E113881AE53EADB61D61D2B94" type="DOI">10.11646/phytotaxa.312.1.8</mods:identifier>
<mods:identifier id="B546A775B41ADC23625304009DE2E221" type="ISSN">1179-3163</mods:identifier>
</mods:mods>
<treatment id="BF1287B4FF99FFCAF6CFFEAE2A78FCF6" LSID="urn:lsid:plazi:treatment:BF1287B4FF99FFCAF6CFFEAE2A78FCF6" httpUri="http://treatment.plazi.org/id/BF1287B4FF99FFCAF6CFFEAE2A78FCF6" lastPageId="2" lastPageNumber="105" pageId="1" pageNumber="104">
<subSubSection id="7FA16529FF99FFC9F6CFFEAE2AF2FEFF" box="[136,754,301,328]" pageId="1" pageNumber="104" type="nomenclature">
<paragraph id="370436A2FF99FFC9F6CFFEAE2AF2FEFF" blockId="1.[136,754,301,328]" box="[136,754,301,328]" pageId="1" pageNumber="104">
<heading id="6C4C81CEFF99FFC9F6CFFEAE2AF2FEFF" box="[136,754,301,328]" fontSize="11" level="2" pageId="1" pageNumber="104" reason="3">
<taxonomicName id="F0BB4D21FF99FFC9F6CFFEAE2A31FEFF" ID-CoL="9W5ZR" authority="N. Wilding" authorityName="N. Wilding" authorityYear="2017" box="[136,561,301,328]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="1" pageNumber="104" phylum="Bryophyta" rank="species" species="heddersonii" status="sp. nov.">
<emphasis id="05CFEAB0FF99FFC9F6CFFEAE29AAFEFF" bold="true" box="[136,426,301,328]" italics="true" pageId="1" pageNumber="104">Entosthodon heddersonii</emphasis>
N. Wilding
</taxonomicName>
,
<emphasis id="05CFEAB0FF99FFC9F47BFEAC2A91FEFF" box="[572,657,303,328]" italics="true" pageId="1" pageNumber="104">
<taxonomicNameLabel id="1EFC57CBFF99FFC9F47BFEAC2A91FEFF" box="[572,657,303,328]" pageId="1" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="AF802A27FF99FFC9F4E1FEAC2AE7FEFF" box="[678,743,303,328]" captionStart="FIGURE 1" captionStartId="1.[136,229,1997,2018]" captionTargetBox="[273,1320,488,1962]" captionTargetId="figure-82@1.[258,1330,478,1969]" captionTargetPageId="1" captionText="FIGURE 1. Entosthodon heddersonii. 1. Habit, dry; 24. Leaves; 5. Leaf apex; 6. Leaf cross-section; 7. Capsule mouth &amp; peristome (teeth missing in image); 8. Operculum; 9. Spores.All from Holotype. Scale bars: A (24) = 500 μm; B (5, 7) = 100 μm; C (1) = 1 mm; D (6) = 100 μm; E (9) = 50 μm; F (8) = 100 μm." pageId="1" pageNumber="104">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF99FFC9F6CFFED62BE2FE39" pageId="1" pageNumber="104" type="diagnosis">
<paragraph id="370436A2FF99FFC9F6CFFED62BE2FE39" blockId="1.[136,1452,341,470]" pageId="1" pageNumber="104">
Diagnosis:—Differing from all other species of the genus
<taxonomicName id="F0BB4D21FF99FFC9F4EAFED62B26FEDD" authorityName="Schwagrichen" authorityYear="1823" box="[685,806,341,362]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="1" pageNumber="104" phylum="Bryophyta" rank="genus">
<emphasis id="05CFEAB0FF99FFC9F4EAFED62B26FEDD" box="[685,806,341,362]" italics="true" pageId="1" pageNumber="104">Entosthodon</emphasis>
</taxonomicName>
by its combination of narrowly oblong-obovate, aristate leaves, that are bordered and toothed in the upper ⅔ and its zygomorphic, peristomate capsules.
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF99FFCAF6CFFE1E2B75FEDA" lastPageId="2" lastPageNumber="105" pageId="1" pageNumber="104" type="materials_examined">
<paragraph id="370436A2FF99FFC9F6CFFE1E2B05FE61" blockId="1.[136,1452,341,470]" pageId="1" pageNumber="104">
<materialsCitation id="87D33CFFFF99FFC9F6CFFE1E2B00FE61" collectingDate="1988-06-09" collectionCode="TYPE, T, R, H, V, BOL, EGR" country="Tanzania" county="Lukwangule Plateau" elevation="2360" location="South Uluguru Mts." municipality="East" pageId="1" pageNumber="104" specimenCount="2" typeStatus="holotype">
<collectionCode id="51AAAE67FF99FFC9F6CFFE1E28C5FE05" box="[136,197,413,434]" pageId="1" pageNumber="104">TYPE</collectionCode>
:—
<collectingCountry id="4FAC7632FF99FFC9F6A6FE1E2960FE05" box="[225,352,413,434]" name="Tanzania" pageId="1" pageNumber="104">TANZANIA</collectingCountry>
.
<location id="32646079FF99FFC9F72CFE1E2A33FE05" LSID="urn:lsid:plazi:treatment:BF1287B4FF99FFCAF6CFFEAE2A78FCF6:32646079FF99FFC9F72CFE1E2A33FE05" box="[363,563,413,434]" country="Tanzania" county="Lukwangule Plateau" municipality="East" name="South Uluguru Mts." pageId="1" pageNumber="104">South Uluguru Mts.</location>
:
<collectingMunicipality id="D760ACD8FF99FFC9F479FE1E2A67FE05" box="[574,615,413,434]" pageId="1" pageNumber="104">East</collectingMunicipality>
edge of
<collectingCounty id="DE654E2EFF99FFC9F485FE1E2B8CFE05" box="[706,908,413,434]" pageId="1" pageNumber="104">Lukwangule Plateau</collectingCounty>
,
<quantity id="F0439B47FF99FFC9F5DFFE1E2C21FE05" box="[920,1057,413,434]" metricMagnitude="3" metricUnit="m" metricValue="2.36" metricValueMax="2.37" metricValueMin="2.35" pageId="1" pageNumber="104" unit="m" value="2360.0" valueMax="2370.0" valueMin="2350.0">
<elevation id="BC96D191FF99FFC9F5DFFE1E2C21FE05" box="[920,1057,413,434]" metricMagnitude="3" metricUnit="m" metricValue="2.36" metricValueMax="2.37" metricValueMin="2.35" pageId="1" pageNumber="104" unit="m" value="2360.0" valueMax="2370.0" valueMin="2350.0">23502370 m</elevation>
</quantity>
,
<date id="43051062FF99FFC9F26AFE1E2CA8FE05" box="[1069,1192,413,434]" pageId="1" pageNumber="104" value="1988-06-09">
<collectingDate id="5341E98AFF99FFC9F26AFE1E2CA8FE05" box="[1069,1192,413,434]" pageId="1" pageNumber="104" value="1988-06-09">9 June 1988</collectingDate>
</date>
,
<emphasis id="05CFEAB0FF99FFC9F2F3FE1E29C0FE61" italics="true" pageId="1" pageNumber="104">
<collectionCode id="51AAAE67FF99FFC9F2F3FE1E2CC3FE05" box="[1204,1219,413,434]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="104">T</collectionCode>
. Pócs,
<collectionCode id="51AAAE67FF99FFC9F34CFE1E2D1CFE05" box="[1291,1308,413,434]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="104">R</collectionCode>
. Ochyra &amp;
<collectionCode id="51AAAE67FF99FFC9F3D3FE1E2DA8FE05" box="[1428,1448,413,434]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="1" pageNumber="104" type="Herbarium">H</collectionCode>
. Bednarek-Ochyra 88108/
<collectionCode id="51AAAE67FF99FFC9F7F5FE4229C0FE61" box="[434,448,449,470]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="104" type="Museum">V</collectionCode>
</emphasis>
(
<collectionCode id="51AAAE67FF99FFC9F78AFE422A01FE61" box="[461,513,449,470]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15865" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="1" pageNumber="104" type="Herbarium">BOL</collectionCode>
,
<typeStatus id="E8008800FF99FFC9F44DFE422A66FE61" box="[522,614,449,470]" pageId="1" pageNumber="104" type="holotype">holotype</typeStatus>
!;
<typeStatus id="E8008800FF99FFC9F435FE422AB7FE61" box="[626,695,449,470]" pageId="1" pageNumber="104" type="isotype">isotype</typeStatus>
<collectionCode id="51AAAE67FF99FFC9F4F9FE422AF6FE61" box="[702,758,449,470]" country="Hungary" httpUri="http://grbio.org/cool/39sm-y0v8" name="Eszterházy Károly College" pageId="1" pageNumber="104" type="Herbarium">EGR</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<caption id="63C4662AFF99FFC9F6CFF84E2A0CF79D" pageId="1" pageNumber="104" startId="1.[136,229,1997,2018]" targetBox="[273,1320,488,1962]" targetPageId="1" targetType="figure">
<paragraph id="370436A2FF99FFC9F6CFF84E2A0CF79D" blockId="1.[136,1452,1997,2090]" pageId="1" pageNumber="104">
<emphasis id="05CFEAB0FF99FFC9F6CFF84E28FDF855" bold="true" box="[136,253,1997,2018]" pageId="1" pageNumber="104">FIGURE 1.</emphasis>
<emphasis id="05CFEAB0FF99FFC9F746F84E29F8F855" box="[257,504,1997,2018]" italics="true" pageId="1" pageNumber="104">
<taxonomicName id="F0BB4D21FF99FFC9F746F84E29F3F855" authorityName="N. Wilding" authorityYear="2017" box="[257,499,1997,2018]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="1" pageNumber="104" phylum="Bryophyta" rank="species" species="heddersonii">Entosthodon heddersonii</taxonomicName>
.
</emphasis>
1. Habit, dry; 24. Leaves; 5. Leaf apex; 6. Leaf cross-section; 7. Capsule mouth &amp; peristome (teeth missing in image); 8. Operculum; 9. Spores. All from Holotype. Scale bars: A (24) = 500 μm; B (5, 7) = 100 μm; C (1) = 1 mm; D (6) = 100 μm; E (9) = 50 μm; F (8) = 100 μm.
</paragraph>
</caption>
<paragraph id="370436A2FF9AFFCAF6CFFF232B75FEDA" blockId="2.[136,1452,160,833]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6CFFF2328D3FF0E" bold="true" box="[136,211,160,185]" pageId="2" pageNumber="105">Plants</emphasis>
small, light-green.
<emphasis id="05CFEAB0FF9AFFCAF7EDFF2329F2FF0E" bold="true" box="[426,498,160,185]" pageId="2" pageNumber="105">Stems</emphasis>
reddish-brown, to
<quantity id="F0439B47FF9AFFCAF484FF232B03FF0E" box="[707,771,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="105" unit="mm" value="3.0">3 mm</quantity>
high, branching multiple times by sub-perigonial innovation; in cross-section with 12 layers of thick-walled cortical cells, a hyalodermis and a central strand; rhizoids cerise.
<emphasis id="05CFEAB0FF9AFFCAF6CFFF6B28DAFEB6" bold="true" box="[136,218,232,257]" pageId="2" pageNumber="105">Leaves</emphasis>
erect-spreading, contorted when dry, narrowly oblong-obovate, (0.75)1.21.6(1.9) ×
<quantity id="F0439B47FF9AFFCAF2F3FF6B2D40FEB6" box="[1204,1344,232,257]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="2" pageNumber="105" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.40.6 mm</quantity>
, plane to slightly recurved, short-acuminate, serrate in upper ⅔, aristate, arista to 300 μm;
<emphasis id="05CFEAB0FF9AFFCAF261FE8F2D23FE92" bold="true" box="[1062,1315,268,293]" pageId="2" pageNumber="105">cells of upper lamina</emphasis>
quadrate to oblong-hexagonal, 3763 × 1525 μm;
<emphasis id="05CFEAB0FF9AFFCAF402FEB32ABDFEFE" bold="true" box="[581,701,304,329]" pageId="2" pageNumber="105">basal cells</emphasis>
50125 × 2030 μm;
<emphasis id="05CFEAB0FF9AFFCAF5F0FEB32C5EFEFE" bold="true" box="[951,1118,304,329]" pageId="2" pageNumber="105">marginal cells</emphasis>
longer and narrower, thicker-walled, forming a border one cell wide;
<emphasis id="05CFEAB0FF9AFFCAF40EFED72A92FEDA" bold="true" box="[585,658,340,365]" pageId="2" pageNumber="105">costae</emphasis>
ending below apex.
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9AFFCAF6FAFEFB2C77FD3A" pageId="2" pageNumber="105" type="description">
<paragraph id="370436A2FF9AFFCAF6FAFEFB2C77FD3A" blockId="2.[136,1452,160,833]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6FAFEFB293DFE26" bold="true" box="[189,317,376,401]" pageId="2" pageNumber="105">Polyoicous</emphasis>
.
<emphasis id="05CFEAB0FF9AFFCAF70FFEFB2988FE26" bold="true" box="[328,392,376,401]" pageId="2" pageNumber="105">Setae</emphasis>
<quantity id="F0439B47FF9AFFCAF7CCFEFB29F3FE26" box="[395,499,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="2" pageNumber="105" unit="mm" value="9.0" valueMax="13.0" valueMin="5.0">513mm</quantity>
long, straight to curved, pale-yellow to reddish-brown.
<emphasis id="05CFEAB0FF9AFFCAF20BFEFB2CB7FE26" bold="true" box="[1100,1207,376,401]" pageId="2" pageNumber="105">Capsules</emphasis>
inclined to horizontal, gibbous, zygomorphic, oblong-obovoid, 1.42.5 ×
<quantity id="F0439B47FF9AFFCAF4FAFE1F2B30FE02" box="[701,816,412,437]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="105" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
, weakly constricted below the mouth when dry, yellow to reddish-brown at maturity, with a well differentiated neck ca. ½ the total length of the capsule;
<emphasis id="05CFEAB0FF9AFFCAF2E5FE432CF0FE6E" bold="true" box="[1186,1264,448,473]" pageId="2" pageNumber="105">mouth</emphasis>
slightly oblique, ca. ¾ the diameter of the capsule;
<emphasis id="05CFEAB0FF9AFFCAF45BFE672AD1FE4A" bold="true" box="[540,721,484,509]" pageId="2" pageNumber="105">exothecial cells</emphasis>
1525 × 37 μm, in cross-section with thick, strongly cuneate anticlinal walls, 35 rows of oblate cells at mouth.
<emphasis id="05CFEAB0FF9AFFCAF48FFD8B2B38FD96" bold="true" box="[712,824,520,545]" pageId="2" pageNumber="105">Opercula</emphasis>
plano-convex, cells twisted anti-clockwise from above.
<emphasis id="05CFEAB0FF9AFFCAF6CFFDAF2900FDF2" bold="true" box="[136,256,556,581]" pageId="2" pageNumber="105">Peristome</emphasis>
double;
<emphasis id="05CFEAB0FF9AFFCAF722FDAF2A17FDF2" bold="true" box="[357,535,556,581]" pageId="2" pageNumber="105">exostome teeth</emphasis>
straight, regular in shape, orange, tapered to an acute apex, to 100 μm high, ca. 20 μm wide at base, smooth, striate, trabeculate;
<emphasis id="05CFEAB0FF9AFFCAF4F2FDD32B78FDDE" bold="true" box="[693,888,592,617]" pageId="2" pageNumber="105">endostome teeth</emphasis>
smooth, hyaline, to 40 μm high and 30 μm wide.
<emphasis id="05CFEAB0FF9AFFCAF6CFFDF728D8FD3A" bold="true" box="[136,216,628,653]" pageId="2" pageNumber="105">Spores</emphasis>
3035 μm, tetrahedral, verrucate-lirate to reticulate.
<emphasis id="05CFEAB0FF9AFFCAF56DFDF72BA2FD3A" bold="true" box="[810,930,628,653]" pageId="2" pageNumber="105">Calyptrae</emphasis>
cucullate, rostrate.
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9AFFCAF6FAFD1B2A78FCF6" pageId="2" pageNumber="105" type="etymology">
<paragraph id="370436A2FF9AFFCAF6FAFD1B2C47FD62" blockId="2.[136,1452,160,833]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6FAFD1B2948FD06" bold="true" box="[189,328,664,689]" pageId="2" pageNumber="105">Etymology:</emphasis>
The species is named in honour of Terry A.J. Hedderson, for his interest in, and extensive collecting of,
<taxonomicName id="F0BB4D21FF9AFFCAF6EAFD3F293BFD62" authorityName="Schwagrichen" authorityYear="1823" box="[173,315,700,725]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="genus">
<emphasis id="05CFEAB0FF9AFFCAF6EAFD3F293BFD62" box="[173,315,700,725]" italics="true" pageId="2" pageNumber="105">Entosthodon</emphasis>
</taxonomicName>
in southern Africa and for his contribution towards African bryology.
</paragraph>
<paragraph id="370436A2FF9AFFCAF6FAFD632A78FCF6" blockId="2.[136,1452,160,833]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6FAFD6329F2FD4E" bold="true" box="[189,498,736,761]" pageId="2" pageNumber="105">Habitat and distribution:</emphasis>
The species is known only from the
<typeStatus id="E8008800FF9AFFCAF5EDFD632BDAFD4E" box="[938,986,736,761]" pageId="2" pageNumber="105">type</typeStatus>
collection in the Uluguru Mountains of
<collectingCountry id="4FAC7632FF9AFFCAF6CFFC8728F0FCAA" box="[136,240,772,797]" name="Tanzania" pageId="2" pageNumber="105">Tanzania</collectingCountry>
. The
<typeStatus id="E8008800FF9AFFCAF76AFC87295DFCAA" box="[301,349,772,797]" pageId="2" pageNumber="105">type</typeStatus>
of
<taxonomicName id="F0BB4D21FF9AFFCAF7C4FC872A26FCAA" authorityName="N. Wilding" authorityYear="2017" box="[387,550,772,797]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="heddersonii">
<emphasis id="05CFEAB0FF9AFFCAF7C4FC872A26FCAA" box="[387,550,772,797]" italics="true" pageId="2" pageNumber="105">E. heddersonii</emphasis>
</taxonomicName>
was collected at an elevation of ca.
<quantity id="F0439B47FF9AFFCAF5FBFC872C11FCAA" box="[956,1041,772,797]" metricMagnitude="3" metricUnit="m" metricValue="2.36" pageId="2" pageNumber="105" unit="m" value="2360.0">2360 m</quantity>
along a rocky, peaty stream bank on the eastern edge of the Lukwangule Plateau.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,296 @@
<document id="DC51D0655D53E1714EE18467F39FC7F4" ID-DOI="10.11646/phytotaxa.312.1.8" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725577053966" checkinUser="felipe" docAuthor="Wilding, Nicholas" docDate="2017" docId="BF1287B4FF9AFFCAF6CFFCED2D71F79A" docLanguage="en" docName="phytotaxa.312.1.8.pdf" docOrigin="Phytotaxa 312 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.312.1.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Entosthodon pocsii N. Wilding 2017, sp. nov." docType="treatment" docVersion="1" lastPageNumber="105" masterDocId="432BFFCCFF98FFC8F647FF832800FFB7" masterDocTitle="Three new species of Entosthodon Schwägr. (Bryopsida, Funariaceae) from sub-Saharan Africa" masterLastPageNumber="110" masterPageNumber="103" pageNumber="105" updateTime="1725586680830" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8180F1E2DB882208BA3C6EDA739A5F18">Three new species of Entosthodon Schwägr. (Bryopsida, Funariaceae) from sub-Saharan Africa</mods:title>
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<mods:namePart id="1088BF250BE2667C5366B88FB10768ED">Wilding, Nicholas</mods:namePart>
<mods:affiliation id="E27A3370387A327A6844572C2B619F15">Université de La Réunion, UMR PVBMT, Pôle de Protection des Plantes, Pôle Forêt, Saint-Pierre, Ile de La Réunion. &amp; Department of Biological Sciences, University of Cape Town, Private Bag X 3, 7701 Rondebosch, South Africa.</mods:affiliation>
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<mods:title id="83714C8D3BC716DA6E974E374A401B43">Phytotaxa</mods:title>
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<mods:date id="D767A25F3539D0A208BBF8E7688F518B">2017</mods:date>
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<mods:identifier id="8C0DD69D6971385FB3C1580B37BEF292" type="ISSN">1179-3163</mods:identifier>
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<treatment id="BF1287B4FF9AFFCAF6CFFCED2D71F79A" LSID="urn:lsid:plazi:treatment:BF1287B4FF9AFFCAF6CFFCED2D71F79A" httpUri="http://treatment.plazi.org/id/BF1287B4FF9AFFCAF6CFFCED2D71F79A" lastPageNumber="105" pageId="2" pageNumber="105">
<subSubSection id="7FA16529FF9AFFCAF6CFFCED2AA7FC3E" box="[136,679,878,905]" pageId="2" pageNumber="105" type="nomenclature">
<paragraph id="370436A2FF9AFFCAF6CFFCED2AA7FC3E" blockId="2.[136,679,878,905]" box="[136,679,878,905]" pageId="2" pageNumber="105">
<heading id="6C4C81CEFF9AFFCAF6CFFCED2AA7FC3E" box="[136,679,878,905]" fontSize="11" level="2" pageId="2" pageNumber="105" reason="3">
<taxonomicName id="F0BB4D21FF9AFFCAF6CFFCED29EAFC3E" ID-CoL="9W5FD" authority="N. Wilding" authorityName="N. Wilding" authorityYear="2017" box="[136,490,878,905]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="pocsii" status="sp. nov.">
<emphasis id="05CFEAB0FF9AFFCAF6CFFCED2966FC3F" bold="true" box="[136,358,878,905]" italics="true" pageId="2" pageNumber="105">Entosthodon pocsii</emphasis>
N. Wilding
</taxonomicName>
,
<emphasis id="05CFEAB0FF9AFFCAF7B0FCF32A4CFC3E" box="[503,588,880,905]" italics="true" pageId="2" pageNumber="105">
<taxonomicNameLabel id="1EFC57CBFF9AFFCAF7B0FCF32A4CFC3E" box="[503,588,880,905]" pageId="2" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="AF802A27FF9AFFCAF41CFCF32A9EFC3E" box="[603,670,880,905]" captionStart="FIGURE 2" captionStartId="3.[136,229,1802,1823]" captionTargetBox="[236,1360,198,1766]" captionTargetId="figure-12@3.[227,1361,190,1777]" captionTargetPageId="3" captionText="FIGURE 2. Entosthodon pocsii. 1. Habit, dry (S. Iversen, E. Persson, B. Petterson &amp; T. Pócs, 87145/D, BOL); 25. Leaves (2 as for 1; 3—5 S. Iversen, E. Persson, B. Petterson &amp; T. Pócs, 87145/N, BOL); 6, 7. Leaf apex (6 as for 1; 7 as for 3); 8. Leaf cross-section (as for 3); 9. Capsule, dry (T. Pócs, D. Harrison &amp; J.M. Mushy 90130/UV, (BOL, EGR); 10. Capsule mouth &amp; peristome (as for 3); 11. Operculum (as for 1); 12. Spores (as for 1). Scale bars: A (25) = 500 μm; B (10) = 100 μm; C (1, 9) = 1 mm; D (8) = 100 μm; E (12) = 50 μm; F (11) = 100 μm; G (6, 7) = 100 μm." pageId="2" pageNumber="105">Fig. 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9AFFCAF6CFFC152A85FAA2" pageId="2" pageNumber="105" type="diagnosis">
<paragraph id="370436A2FF9AFFCAF6CFFC152C7BFC78" blockId="2.[136,1452,918,1047]" pageId="2" pageNumber="105">
Diagnosis:—Differing from all other species of the genus
<taxonomicName id="F0BB4D21FF9AFFCAF4FCFC152B34FC1C" authorityName="Schwagrichen" authorityYear="1823" box="[699,820,918,939]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="genus">
<emphasis id="05CFEAB0FF9AFFCAF4FCFC152B34FC1C" box="[699,820,918,939]" italics="true" pageId="2" pageNumber="105">Entosthodon</emphasis>
</taxonomicName>
by its concave, ovate to oblong-obovate, short acuminate, entire, leaves and its ± upright, radially symmetric, peristomate capsules and finely verrucate-lirate spores.
</paragraph>
<paragraph id="370436A2FF9AFFCAF6CFFC5D2BFEFBA0" blockId="2.[136,1452,918,1047]" pageId="2" pageNumber="105">
<materialsCitation id="87D33CFFFF9AFFCAF6CFFC5D2BFAFBA0" collectingDate="1987-05-26" collectionCode="S, E, B, T, BOL, EGR" country="Tanzania" elevation="2500" location="Kipengere" municipality="Type" pageId="2" pageNumber="105" specimenCount="2" stateProvince="Njombe" typeStatus="holotype">
<collectingMunicipality id="D760ACD8FF9AFFCAF6CFFC5D28B9FC44" box="[136,185,990,1011]" pageId="2" pageNumber="105">Type</collectingMunicipality>
:—
<collectingCountry id="4FAC7632FF9AFFCAF692FC5D2954FC44" box="[213,340,990,1011]" name="Tanzania" pageId="2" pageNumber="105">TANZANIA</collectingCountry>
.
<collectingRegion id="F57FF840FF9AFFCAF727FC5D2A03FC44" box="[352,515,990,1011]" country="Tanzania" name="Njombe" pageId="2" pageNumber="105">Njombe District</collectingRegion>
:
<location id="32646079FF9AFFCAF448FC5D2A73FC44" LSID="urn:lsid:plazi:treatment:BF1287B4FF9AFFCAF6CFFCED2D71F79A:32646079FF9AFFCAF448FC5D2A73FC44" box="[527,627,990,1011]" country="Tanzania" municipality="Type" name="Kipengere" pageId="2" pageNumber="105" stateProvince="Njombe">Kipengere</location>
range,
<locationDeviation id="26B62F80FF9AFFCAF486FC5D2C5EFC44" box="[705,1118,990,1011]" firstLocation="Mts. Ishinga" pageId="2" pageNumber="105" secondLocation="Sanshashi Hill">between Mts. Ishinga and Sanshashi Hill</locationDeviation>
,
<quantity id="F0439B47FF9AFFCAF22CFC5D2CF5FC44" box="[1131,1269,990,1011]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="2" pageNumber="105" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">
<elevation id="BC96D191FF9AFFCAF22CFC5D2CF5FC44" box="[1131,1269,990,1011]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="2" pageNumber="105" unit="m" value="2500.0" valueMax="2600.0" valueMin="2400.0">24002600 m</elevation>
</quantity>
,
<date id="43051062FF9AFFCAF344FC5D2D8CFC44" box="[1283,1420,990,1011]" pageId="2" pageNumber="105" value="1987-05-26">
<collectingDate id="5341E98AFF9AFFCAF344FC5D2D8CFC44" box="[1283,1420,990,1011]" pageId="2" pageNumber="105" value="1987-05-26">26 May 1987</collectingDate>
</date>
,
<emphasis id="05CFEAB0FF9AFFCAF3DEFC5D2AB9FBA0" italics="true" pageId="2" pageNumber="105">
<collectionCode id="51AAAE67FF9AFFCAF3DEFC5D2DA8FC44" box="[1433,1448,990,1011]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="2" pageNumber="105" type="Herbarium">S</collectionCode>
. Iversen,
<collectionCode id="51AAAE67FF9AFFCAF757FB812922FBA0" box="[272,290,1026,1047]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="2" pageNumber="105" type="Herbarium">E</collectionCode>
. Persson,
<collectionCode id="51AAAE67FF9AFFCAF7C3FB812995FBA0" box="[388,405,1026,1047]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="2" pageNumber="105" type="Herbarium">B</collectionCode>
. Petterson &amp;
<collectionCode id="51AAAE67FF9AFFCAF45EFB812A28FBA0" box="[537,552,1026,1047]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="105">T</collectionCode>
. Pócs 87145/D
</emphasis>
(
<collectionCode id="51AAAE67FF9AFFCAF481FB812AFAFBA0" box="[710,762,1026,1047]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15865" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="2" pageNumber="105" type="Herbarium">BOL</collectionCode>
,
<typeStatus id="E8008800FF9AFFCAF543FB812B60FBA0" box="[772,864,1026,1047]" pageId="2" pageNumber="105" type="holotype">holotype</typeStatus>
!;
<typeStatus id="E8008800FF9AFFCAF52BFB812BB1FBA0" box="[876,945,1026,1047]" pageId="2" pageNumber="105" type="isotype">isotype</typeStatus>
<collectionCode id="51AAAE67FF9AFFCAF5FFFB812BF0FBA0" box="[952,1008,1026,1047]" country="Hungary" httpUri="http://grbio.org/cool/39sm-y0v8" name="Eszterházy Károly College" pageId="2" pageNumber="105" type="Herbarium">EGR</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="370436A2FF9AFFCAF6CFFBCB2A85FAA2" blockId="2.[136,1452,1096,2093]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6CFFBCB28D3FBD6" bold="true" box="[136,211,1096,1121]" pageId="2" pageNumber="105">Plants</emphasis>
small, light-green.
<emphasis id="05CFEAB0FF9AFFCAF7EDFBCB29F2FBD6" bold="true" box="[426,498,1096,1121]" pageId="2" pageNumber="105">Stems</emphasis>
reddish-brown, to
<quantity id="F0439B47FF9AFFCAF484FBCB2B03FBD6" box="[707,771,1096,1121]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="105" unit="mm" value="5.0">5 mm</quantity>
high, branching multiple times by sub-perigonial innovation; in cross-section with 12 layers of thick-walled cortical cells, a hyalodermis and a central strand; rhizoids cerise.
<emphasis id="05CFEAB0FF9AFFCAF6CFFB1328DAFB1E" bold="true" box="[136,218,1168,1193]" pageId="2" pageNumber="105">Leaves</emphasis>
erect-spreading, little contorted when dry, ovate to oblong-obovate, 1.22 ×
<quantity id="F0439B47FF9AFFCAF226FB132CEDFB1E" box="[1121,1261,1168,1193]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="2" pageNumber="105" unit="mm" value="0.65" valueMax="0.9" valueMin="0.4">0.40.9 mm</quantity>
, concave, short-acuminate, entire, aristate or not, arista if present to 400 μm;
<emphasis id="05CFEAB0FF9AFFCAF50DFB372C49FB7A" bold="true" box="[842,1097,1204,1229]" pageId="2" pageNumber="105">cells of upper lamina</emphasis>
quadrate to oblong-hexagonal, 2355 × (10)1530(35) μm;
<emphasis id="05CFEAB0FF9AFFCAF78AFB5B2A46FB46" bold="true" box="[461,582,1240,1265]" pageId="2" pageNumber="105">basal cells</emphasis>
62150(188) × (15)2035(43) μm;
<emphasis id="05CFEAB0FF9AFFCAF599FB5B2C86FB46" bold="true" box="[990,1158,1240,1265]" pageId="2" pageNumber="105">marginal cells</emphasis>
longer and narrower near apex;
<emphasis id="05CFEAB0FF9AFFCAF68BFB7F2915FAA2" bold="true" box="[204,277,1276,1301]" pageId="2" pageNumber="105">costae</emphasis>
ending below apex to excurrent.
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9AFFCAF6FAFAA32D58F9EE" pageId="2" pageNumber="105" type="description">
<paragraph id="370436A2FF9AFFCAF6FAFAA32D58F9EE" blockId="2.[136,1452,1096,2093]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6FAFAA3293DFA8E" bold="true" box="[189,317,1312,1337]" pageId="2" pageNumber="105">Polyoicous</emphasis>
.
<emphasis id="05CFEAB0FF9AFFCAF708FAA3298FFA8E" bold="true" box="[335,399,1312,1337]" pageId="2" pageNumber="105">Setae</emphasis>
<quantity id="F0439B47FF9AFFCAF7DEFAA329FAFA8E" box="[409,506,1312,1337]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="105" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
long, straight, pale-yellow to reddish-brown.
<emphasis id="05CFEAB0FF9AFFCAF255FAA32C7DFA8E" bold="true" box="[1042,1149,1312,1337]" pageId="2" pageNumber="105">Capsules</emphasis>
erect to inclined, radially symmetric or with slight bend in neck, oblong-pyriform, 1.21.7 ×
<quantity id="F0439B47FF9AFFCAF5DDFAC72C26FAEA" box="[922,1062,1348,1373]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="2" pageNumber="105" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
, weakly constricted below mouth when dry, yellow to reddish-brown at maturity, with a well differentiated neck ca. ½ total length of capsule;
<emphasis id="05CFEAB0FF9AFFCAF319FAEB2DACFA36" bold="true" box="[1374,1452,1384,1409]" pageId="2" pageNumber="105">mouth</emphasis>
transverse, ca. ⅔ diameter of capsule;
<emphasis id="05CFEAB0FF9AFFCAF478FA0F2AF2FA12" bold="true" box="[575,754,1420,1445]" pageId="2" pageNumber="105">exothecial cells</emphasis>
(30)4068 × 1020 μm, in cross-section with thick, strongly cuneate anticlinal walls, 36 rows of oblate cells at mouth.
<emphasis id="05CFEAB0FF9AFFCAF564FA332B93FA7E" bold="true" box="[803,915,1456,1481]" pageId="2" pageNumber="105">Opercula</emphasis>
plano-convex, cells not or slightly twisted anti-clockwise from above.
<emphasis id="05CFEAB0FF9AFFCAF7CEFA572A01FA5A" bold="true" box="[393,513,1492,1517]" pageId="2" pageNumber="105">Peristome</emphasis>
double and rudimentary;
<emphasis id="05CFEAB0FF9AFFCAF55AFA572BCCFA5A" bold="true" box="[797,972,1492,1517]" pageId="2" pageNumber="105">exostome teeth</emphasis>
straight, irregular in shape, orange, tapered to an acute or rounded apex, sometimes becoming hyaline in upper ⅓, to 175 μm high and 60 μm wide at base, finely papillose to verrucate-lirate, trabeculate, weakly appendiculate;
<emphasis id="05CFEAB0FF9AFFCAF513F99F2C16F982" bold="true" box="[852,1046,1564,1589]" pageId="2" pageNumber="105">endostome teeth</emphasis>
rudimentary, smooth, hyaline, to 40 μm high and 22 μm wide.
<emphasis id="05CFEAB0FF9AFFCAF7E8F9C329FFF9EE" bold="true" box="[431,511,1600,1625]" pageId="2" pageNumber="105">Spores</emphasis>
2738 μm, tetrahedral, finely papillose-lirate.
<emphasis id="05CFEAB0FF9AFFCAF24DF9C32C82F9EE" bold="true" box="[1034,1154,1600,1625]" pageId="2" pageNumber="105">Calyptrae</emphasis>
cucullate, rostrate.
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9AFFCAF6FAF9E729C5F972" pageId="2" pageNumber="105" type="materials_examined">
<paragraph id="370436A2FF9AFFCAF6FAF9E729C5F972" blockId="2.[136,1452,1096,2093]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6FAF9E72A43F9CA" bold="true" box="[189,579,1636,1661]" pageId="2" pageNumber="105">Additional specimens examined:</emphasis>
<materialsCitation id="87D33CFFFF9AFFCAF40DF9E72998F916" collectingDate="1990-06-09" collectionCode="N, P, T, J, M, UV, BOL, EGR" collectorName="Pocs, D &amp; Mushy" country="Tanzania" location="Arusha" pageId="2" pageNumber="105" specimenCount="1" stateProvince="Arusha">
<collectingCountry id="4FAC7632FF9AFFCAF40DF9E72ADCF9CA" box="[586,732,1636,1661]" name="Tanzania" pageId="2" pageNumber="105">TANZANIA</collectingCountry>
.
<collectingRegion id="F57FF840FF9AFFCAF4A1F9E72B37F9CA" box="[742,823,1636,1661]" country="Tanzania" name="Arusha" pageId="2" pageNumber="105">Arusha</collectingRegion>
<collectionCode id="51AAAE67FF9AFFCAF578F9E72B57F9CA" box="[831,855,1636,1661]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="2" pageNumber="105" type="Herbarium">N</collectionCode>
.
<collectionCode id="51AAAE67FF9AFFCAF51CF9E72B6DF9CA" box="[859,877,1636,1661]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="105" type="Herbarium">P</collectionCode>
.:
<date id="43051062FF9AFFCAF53AF9E72C0AF9CA" box="[893,1034,1636,1661]" pageId="2" pageNumber="105" value="1990-06-09">
<collectingDate id="5341E98AFF9AFFCAF53AF9E72C0AF9CA" box="[893,1034,1636,1661]" pageId="2" pageNumber="105" value="1990-06-09">9 June 1990</collectingDate>
</date>
,
<emphasis id="05CFEAB0FF9AFFCAF252F9E728FBF916" italics="true" pageId="2" pageNumber="105">
<collectionCode id="51AAAE67FF9AFFCAF252F9E72C26F9CA" box="[1045,1062,1636,1661]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="105">T</collectionCode>
.
<collectorName id="9A4E5374FF9AFFCAF275F9E72C8FF9CA" box="[1074,1167,1636,1661]" pageId="2" pageNumber="105">Pócs, D</collectorName>
. Harrison &amp;
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.
<collectionCode id="51AAAE67FF9AFFCAF37DF9E72D56F9CA" box="[1338,1366,1636,1661]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="2" pageNumber="105" type="Herbarium">M</collectionCode>
.
<collectorName id="9A4E5374FF9AFFCAF326F9E72DACF9CA" box="[1377,1452,1636,1661]" pageId="2" pageNumber="105">Mushy</collectorName>
90130/
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</emphasis>
(
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,
<collectionCode id="51AAAE67FF9AFFCAF714F90B2990F916" box="[339,400,1672,1697]" country="Hungary" httpUri="http://grbio.org/cool/39sm-y0v8" name="Eszterházy Károly College" pageId="2" pageNumber="105" type="Herbarium">EGR</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="87D33CFFFF9AFFCAF7E4F90B29C1F972" collectingDate="1987-05-26" collectionCode="S, E, B, T, N, BOL, EGR" collectorName="Iversen &amp; Persson &amp; Petterson &amp; Pocs" country="Tanzania" location="Kipengere Range" pageId="2" pageNumber="105" specimenCount="1" stateProvince="Njombe">
<collectingRegion id="F57FF840FF9AFFCAF7E4F90B2A5DF916" box="[419,605,1672,1697]" country="Tanzania" name="Njombe" pageId="2" pageNumber="105">Njombe District</collectingRegion>
:
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,
<date id="43051062FF9AFFCAF57CF90B2BD5F916" box="[827,981,1672,1697]" pageId="2" pageNumber="105" value="1987-05-26">
<collectingDate id="5341E98AFF9AFFCAF57CF90B2BD5F916" box="[827,981,1672,1697]" pageId="2" pageNumber="105" value="1987-05-26">26 May 1987</collectingDate>
</date>
,
<emphasis id="05CFEAB0FF9AFFCAF5A7F90B2926F972" italics="true" pageId="2" pageNumber="105">
<collectionCode id="51AAAE67FF9AFFCAF5A7F90B2BF1F916" box="[992,1009,1672,1697]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="2" pageNumber="105" type="Herbarium">S</collectionCode>
.
<collectorName id="9A4E5374FF9AFFCAF5BAF90B2C52F916" box="[1021,1106,1672,1697]" pageId="2" pageNumber="105">Iversen</collectorName>
,
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.
<collectorName id="9A4E5374FF9AFFCAF239F90B2CDBF916" box="[1150,1243,1672,1697]" pageId="2" pageNumber="105">Persson</collectorName>
,
<collectionCode id="51AAAE67FF9AFFCAF2A0F90B2CFBF916" box="[1255,1275,1672,1697]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="2" pageNumber="105" type="Herbarium">B</collectionCode>
.
<collectorName id="9A4E5374FF9AFFCAF341F90B2D71F916" box="[1286,1393,1672,1697]" pageId="2" pageNumber="105">Petterson</collectorName>
&amp;
<collectionCode id="51AAAE67FF9AFFCAF3D0F90B2DA8F916" box="[1431,1448,1672,1697]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="105">T</collectionCode>
.
<collectorName id="9A4E5374FF9AFFCAF6CFF92F28BEF972" box="[136,190,1708,1733]" pageId="2" pageNumber="105">Pócs</collectorName>
87145/
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</emphasis>
(
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,
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9AFFCAF6FAF9532D71F79A" pageId="2" pageNumber="105" type="etymology">
<paragraph id="370436A2FF9AFFCAF6FAF9532B10F8BA" blockId="2.[136,1452,1096,2093]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6FAF9532948F95E" bold="true" box="[189,328,1744,1769]" pageId="2" pageNumber="105">Etymology:</emphasis>
The species is named in honour of Tamas Pócs for his immense work on the African bryophyte flora and attention to even the smallest species of
<taxonomicName id="F0BB4D21FF9AFFCAF43CF9772B09F8BA" authorityName="Schwagrichen" authorityYear="1823" box="[635,777,1780,1805]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="genus">
<emphasis id="05CFEAB0FF9AFFCAF43CF9772B09F8BA" box="[635,777,1780,1805]" italics="true" pageId="2" pageNumber="105">Entosthodon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="370436A2FF9AFFCAF6FAF89B2C74F8E2" blockId="2.[136,1452,1096,2093]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6FAF89B29F2F886" bold="true" box="[189,498,1816,1841]" pageId="2" pageNumber="105">Habitat and Distribution:</emphasis>
The species is known from only three collections in
<collectingCountry id="4FAC7632FF9AFFCAF20DF89B2CB2F886" box="[1098,1202,1816,1841]" name="Tanzania" pageId="2" pageNumber="105">Tanzania</collectingCountry>
, two in the Kipengere Range and one from
<collectingRegion id="F57FF840FF9AFFCAF736F8BF29C2F8E2" box="[369,450,1852,1877]" country="Tanzania" name="Arusha" pageId="2" pageNumber="105">Arusha</collectingRegion>
National Park, where it occurs at high elevations ca.
<quantity id="F0439B47FF9AFFCAF25FF8BF2C70F8E2" box="[1048,1136,1852,1877]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="105" unit="m" value="2700.0">2700 m</quantity>
.
</paragraph>
<paragraph id="370436A2FF9AFFCAF6FAF8E32999F852" blockId="2.[136,1452,1096,2093]" pageId="2" pageNumber="105">
<emphasis id="05CFEAB0FF9AFFCAF6FAF8E328FFF8CE" bold="true" box="[189,255,1888,1913]" pageId="2" pageNumber="105">Note:</emphasis>
Phylogenetic analyses (Wilding
<emphasis id="05CFEAB0FF9AFFCAF42BF8E32AA4F8CE" box="[620,676,1888,1913]" italics="true" pageId="2" pageNumber="105">et al.</emphasis>
in prep.) place
<taxonomicName id="F0BB4D21FF9AFFCAF508F8E32BB0F8CE" authorityName="N. Wilding" authorityYear="2017" box="[847,944,1888,1913]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="pocsii">
<emphasis id="05CFEAB0FF9AFFCAF508F8E32BB0F8CE" box="[847,944,1888,1913]" italics="true" pageId="2" pageNumber="105">E. pocsii</emphasis>
</taxonomicName>
sister to
<taxonomicName id="F0BB4D21FF9AFFCAF254F8E3293BF82A" authority="Brotherus (1913: 737) Kis (1985: 220)" authorityName="Brotherus (1913: 737) Kis" authorityYear="1985" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="pertenellus">
<emphasis id="05CFEAB0FF9AFFCAF254F8E32CAEF8CE" box="[1043,1198,1888,1913]" italics="true" pageId="2" pageNumber="105">E. pertenellus</emphasis>
<bibRefCitation id="532A4B53FF9AFFCAF2F3F8E32DABF8CE" author="Brotherus, V. F." box="[1204,1451,1888,1913]" pageId="2" pageNumber="105" refId="ref3849" refString="Brotherus, V. F. (1913) Forschungsreise nach Deutsch-Ostafrika und Sudafrika. Denkschriften der Kaiserlichen Akademie der Wissenschaften, Wien. Mathematisch-naturwissenschaftliche Klasse 88: 737." type="journal volume" year="1913">Brotherus (1913: 737)</bibRefCitation>
<bibRefCitation id="532A4B53FF9AFFCAF6CFF807293BF82A" author="Kis, G." box="[136,315,1924,1949]" pageId="2" pageNumber="105" refId="ref4060" refString="Kis, G. (1985) Mosses of South-East Tropical Africa: An annotated list with distributionnal data. A. Berczik, ed., Vacratot: Institute of Ecology and Botany of the Hungarian Acadamy of Sciences, Hungary, 170 pp." type="book" year="1985">Kis (1985: 220)</bibRefCitation>
</taxonomicName>
another African endemic (see:
<bibRefCitation id="532A4B53FF9AFFCAF4DCF8072B7BF82A" author="Wilding, N. &amp; Ah-Peng, C. &amp; Hedderson, T. A." box="[667,891,1924,1949]" pageId="2" pageNumber="105" pagination="457 - 460" refId="ref4560" refString="Wilding, N., Ah-Peng, C. &amp; Hedderson, T. A. (2013) Out of obscurity: On the identity of Entosthodon Pertenellus (Broth.) Kis (Funariaceae). Polish Botanical Journal 58 (2): 457 - 460. https: // doi. org / 10.2478 / pbj- 2013 - 0046" type="journal article" year="2013">
Wilding
<emphasis id="05CFEAB0FF9AFFCAF4BAF8072B36F82A" box="[765,822,1924,1949]" italics="true" pageId="2" pageNumber="105">et al.</emphasis>
2013
</bibRefCitation>
). The two species share a similar habitat and both possess a leaf architecture that can be highly variable among populations.
<taxonomicName id="F0BB4D21FF9AFFCAF58CF82B2C2DF876" authorityName="N. Wilding" authorityYear="2017" box="[971,1069,1960,1985]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="pocsii">
<emphasis id="05CFEAB0FF9AFFCAF58CF82B2C2DF876" box="[971,1069,1960,1985]" italics="true" pageId="2" pageNumber="105">E. pocsii</emphasis>
</taxonomicName>
differs from
<taxonomicName id="F0BB4D21FF9AFFCAF285F82B2D5EF876" authorityName="Brotherus (1913: 737) Kis" authorityYear="1985" box="[1218,1374,1960,1985]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="pertenellus">
E.
<emphasis id="05CFEAB0FF9AFFCAF2A5F82B2D5EF876" box="[1250,1374,1960,1985]" italics="true" pageId="2" pageNumber="105">pertenellus</emphasis>
</taxonomicName>
by the presence of a peristome.
</paragraph>
<paragraph id="370436A2FF9AFFCAF6FAF8732D71F79A" blockId="2.[136,1452,1096,2093]" pageId="2" pageNumber="105">
Macroscopically, the species may also be confused with
<taxonomicName id="F0BB4D21FF9AFFCAF576F8732DA8F7BE" authority="Hornschuch (1820: 59) Muller (1848: 126)" authorityName="Hornschuch (1820: 59) Muller" authorityYear="1848" box="[817,1448,2032,2057]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="bergianus">
<emphasis id="05CFEAB0FF9AFFCAF576F8732BBFF7BE" box="[817,959,2032,2057]" italics="true" pageId="2" pageNumber="105">E. bergianus</emphasis>
<bibRefCitation id="532A4B53FF9AFFCAF581F8732CC8F7BE" author="Hornschuch, C. F." box="[966,1224,2032,2057]" pageId="2" pageNumber="105" refId="ref4040" refString="Hornschuch, C. F. (1820) Musci Frondosi Exotici. Horae Physicae Berolinenses 12: 59." type="journal volume" year="1820">Hornschuch (1820: 59)</bibRefCitation>
<bibRefCitation id="532A4B53FF9AFFCAF288F8732DA8F7BE" author="Muller, C." box="[1231,1448,2032,2057]" pageId="2" pageNumber="105" refId="ref4433" refString="Muller, C. (1848) Funarioideae. In: Synopsis Muscorum Frondosorum omnium hucusque Cognitorum 1. A. Foerstner, Berlin, 812 pp." type="book" year="1848">Müller (1848: 126)</bibRefCitation>
</taxonomicName>
. However, their spore ornamentations differ and
<taxonomicName id="F0BB4D21FF9AFFCAF4E6F7972B2FF79A" authorityName="Hornschuch (1820: 59) Muller" authorityYear="1848" box="[673,815,2068,2093]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="bergianus">
<emphasis id="05CFEAB0FF9AFFCAF4E6F7972B2FF79A" box="[673,815,2068,2093]" italics="true" pageId="2" pageNumber="105">E. bergianus</emphasis>
</taxonomicName>
is generally larger and the costa is never excurrent.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,255 @@
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<taxonomicName id="F0BB4D21FF9CFFCCF6CFFF1E2A41FF0F" ID-CoL="9VM73" authority="N. Wilding" authorityName="N. Wilding" authorityYear="2017" box="[136,577,157,184]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="species" species="zygolimbatus" status="sp. nov.">
<emphasis id="05CFEAB0FF9CFFCCF6CFFF1E29BAFF0F" bold="true" box="[136,442,157,184]" italics="true" pageId="4" pageNumber="107">Entosthodon zygolimbatus</emphasis>
N. Wilding
</taxonomicName>
,
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<taxonomicNameLabel id="1EFC57CBFF9CFFCCF40CFF1C2AA0FF0F" box="[587,672,159,184]" pageId="4" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="AF802A27FF9CFFCCF4E8FF1C2AF1FF0F" box="[687,753,159,184]" captionStart="FIGURE 3" captionStartId="4.[136,229,1945,1966]" captionTargetBox="[227,1361,358,1920]" captionTargetId="figure-70@4.[227,1361,353,1920]" captionTargetPageId="4" captionText="FIGURE 3. Entosthodon zygolimbatus. 1. Habit, dry; 24. Leaves; 5. Leaf apex; 6. Leaf cross-section; 7. Capsule mouth &amp; peristome (teeth missing in image); 8. Spores. All from: T. Pócs, R. Ochyra &amp; H. BednarekOchyra 88108/M, BOL. Scale bars: A (24) = 500 μm; B (5, 7) = 100 μm; C (1) = 1 mm; D (6) = 100 μm; E (8) = 50 μm." pageId="4" pageNumber="107">Fig. 3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9CFFCDF6CFFF452CC4FEDA" lastPageId="5" lastPageNumber="108" pageId="4" pageNumber="107" type="diagnosis">
<paragraph id="370436A2FF9CFFCCF6CFFF452A6CFF48" blockId="4.[136,1452,198,327]" pageId="4" pageNumber="107">
Diagnosis:—Differing from all other species of the genus
<taxonomicName id="F0BB4D21FF9CFFCCF4F3FF452B2DFF6C" authorityName="Schwagrichen" authorityYear="1823" box="[692,813,198,219]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="genus">
<emphasis id="05CFEAB0FF9CFFCCF4F3FF452B2DFF6C" box="[692,813,198,219]" italics="true" pageId="4" pageNumber="107">Entosthodon</emphasis>
</taxonomicName>
by its combination of zygomorphic capsules, strongly limbate leaf margins and coarsely verrucate-lirate spores.
</paragraph>
<paragraph id="370436A2FF9CFFCCF6CFFE8D29EFFEF0" blockId="4.[136,1452,198,327]" pageId="4" pageNumber="107">
<materialsCitation id="87D33CFFFF9CFFCCF6CFFE8D29EBFEF0" collectingDate="1980-10-14" collectionCode="B, K, PRE, MO" country="South Africa" county="Type" elevation="2010" location="Verlorenvallei" municipality="Dullstroom" pageId="4" pageNumber="107" specimenCount="2" stateProvince="Mpumalanga" typeStatus="holotype">
<collectingCounty id="DE654E2EFF9CFFCCF6CFFE8D28BAFE94" box="[136,186,270,291]" pageId="4" pageNumber="107">Type</collectingCounty>
:—
<collectingCountry id="4FAC7632FF9CFFCCF692FE8D2988FE94" box="[213,392,270,291]" name="South Africa" pageId="4" pageNumber="107">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="F57FF840FF9CFFCCF7D3FE8D2A17FE94" box="[404,535,270,291]" country="South Africa" name="Mpumalanga" pageId="4" pageNumber="107">Mpumalanga</collectingRegion>
:
<collectingMunicipality id="D760ACD8FF9CFFCCF465FE8D2A92FE94" box="[546,658,270,291]" pageId="4" pageNumber="107">Dullstroom</collectingMunicipality>
,
<quantity id="F0439B47FF9CFFCCF4DAFE8D2ADDFE94" box="[669,733,270,291]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="107" unit="km" value="14.0">14 km</quantity>
north on
<location id="32646079FF9CFFCCF507FE8D2BC8FE94" LSID="urn:lsid:plazi:treatment:BF1287B4FF9CFFCDF6CFFF1E28CFFB1E:32646079FF9CFFCCF507FE8D2BC8FE94" box="[832,968,270,291]" country="South Africa" county="Type" municipality="Dullstroom" name="Verlorenvallei" pageId="4" pageNumber="107" stateProvince="Mpumalanga">Verlorenvallei</location>
farm,
<quantity id="F0439B47FF9CFFCCF24BFE8D2C59FE94" box="[1036,1113,270,291]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="4" pageNumber="107" unit="m" value="2010.0">
<elevation id="BC96D191FF9CFFCCF24BFE8D2C59FE94" box="[1036,1113,270,291]" metricMagnitude="3" metricUnit="m" metricValue="2.01" pageId="4" pageNumber="107" unit="m" value="2010.0">2010 m</elevation>
</quantity>
,
<date id="43051062FF9CFFCCF222FE8D2D0EFE94" box="[1125,1294,270,291]" pageId="4" pageNumber="107" value="1980-10-14">
<collectingDate id="5341E98AFF9CFFCCF222FE8D2D0EFE94" box="[1125,1294,270,291]" pageId="4" pageNumber="107" value="1980-10-14">14 October 1980</collectingDate>
</date>
,
<emphasis id="05CFEAB0FF9CFFCCF35DFE8D2DABFE94" box="[1306,1451,270,291]" italics="true" pageId="4" pageNumber="107">
<collectionCode id="51AAAE67FF9CFFCCF35DFE8D2D2BFE94" box="[1306,1323,270,291]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="4" pageNumber="107" type="Herbarium">B</collectionCode>
.
<collectionCode id="51AAAE67FF9CFFCCF369FE8D2D42FE94" box="[1326,1346,270,291]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="4" pageNumber="107" type="Herbarium">K</collectionCode>
. Drews 19
</emphasis>
(
<collectionCode id="51AAAE67FF9CFFCCF683FEB128F4FEF0" box="[196,244,306,327]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="4" pageNumber="107" type="Herbarium">PRE</collectionCode>
,
<typeStatus id="E8008800FF9CFFCCF6BAFEB12959FEF0" box="[253,345,306,327]" pageId="4" pageNumber="107" type="holotype">holotype</typeStatus>
!;
<typeStatus id="E8008800FF9CFFCCF721FEB129ABFEF0" box="[358,427,306,327]" pageId="4" pageNumber="107" type="isotype">isotype</typeStatus>
<collectionCode id="51AAAE67FF9CFFCCF7F6FEB129DFFEF0" box="[433,479,306,327]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="4" pageNumber="107" type="Herbarium">MO</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<caption id="63C4662AFF9CFFCCF6CFF81A2B03F841" pageId="4" pageNumber="107" startId="4.[136,229,1945,1966]" targetBox="[227,1361,358,1920]" targetPageId="4" targetType="figure">
<paragraph id="370436A2FF9CFFCCF6CFF81A2B03F841" blockId="4.[136,1452,1945,2038]" pageId="4" pageNumber="107">
<emphasis id="05CFEAB0FF9CFFCCF6CFF81A28FFF819" bold="true" box="[136,255,1945,1966]" pageId="4" pageNumber="107">FIGURE 3.</emphasis>
<emphasis id="05CFEAB0FF9CFFCCF741F81A2A0DF819" box="[262,525,1945,1966]" italics="true" pageId="4" pageNumber="107">
<taxonomicName id="F0BB4D21FF9CFFCCF741F81A2A09F819" authorityName="N. Wilding" authorityYear="2017" box="[262,521,1945,1966]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="species" species="zygolimbatus">Entosthodon zygolimbatus</taxonomicName>
.
</emphasis>
1. Habit, dry; 24. Leaves; 5. Leaf apex; 6. Leaf cross-section; 7. Capsule mouth &amp; peristome (teeth missing in image); 8. Spores. All from:
<emphasis id="05CFEAB0FF9CFFCCF404F83E2C40F865" box="[579,1088,1981,2002]" italics="true" pageId="4" pageNumber="107">T. Pócs, R. Ochyra &amp; H. BednarekOchyra 88108/M</emphasis>
, BOL. Scale bars: A (24) = 500 μm; B (5, 7) = 100 μm; C (1) = 1 mm; D (6) = 100 μm; E (8) = 50 μm.
</paragraph>
</caption>
<paragraph id="370436A2FF9DFFCDF6CFFF232CC4FEDA" blockId="5.[136,1452,160,1193]" pageId="5" pageNumber="108">
<emphasis id="05CFEAB0FF9DFFCDF6CFFF2328D3FF0E" bold="true" box="[136,211,160,185]" pageId="5" pageNumber="108">Plants</emphasis>
medium, light-green.
<emphasis id="05CFEAB0FF9DFFCDF7A6FF232A29FF0E" bold="true" box="[481,553,160,185]" pageId="5" pageNumber="108">Stems</emphasis>
reddish-brown, to
<quantity id="F0439B47FF9DFFCDF556FF232B59FF0E" box="[785,857,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="108" unit="mm" value="5.0">5 mm</quantity>
high, branching multiple times by sub-perigonial innovation; in cross-section with 12 layers of thick-walled cortical cells, a hyalodermis and a central strand; rhizoids cerise.
<emphasis id="05CFEAB0FF9DFFCDF69BFF6B292EFEB6" bold="true" box="[220,302,232,257]" pageId="5" pageNumber="108">Leaves</emphasis>
erect-spreading, contorted when dry, oblong-obovate to spathulate, 2.23 ×
<quantity id="F0439B47FF9DFFCDF2F6FF6B2D3EFEB6" box="[1201,1342,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="5" pageNumber="108" unit="mm" value="1.3" valueMax="1.5" valueMin="1.1">1.11.5 mm</quantity>
, plane to slightly concave, short-acuminate to apiculate, entire to weakly serrate in upper ⅓, arista absent;
<emphasis id="05CFEAB0FF9DFFCDF2F1FE8F2DABFE92" bold="true" box="[1206,1451,268,293]" pageId="5" pageNumber="108">cells of upper lamina</emphasis>
rectangular or rhomboidal to oblong-hexagonal, 4563 × 2538 μm;
<emphasis id="05CFEAB0FF9DFFCDF5CEFEB32C00FEFE" bold="true" box="[905,1024,304,329]" pageId="5" pageNumber="108">basal cells</emphasis>
112225 × 2538 μm;
<emphasis id="05CFEAB0FF9DFFCDF342FEB32DABFEFE" bold="true" box="[1285,1451,304,329]" pageId="5" pageNumber="108">marginal cells</emphasis>
frequently narrower, thicker-walled, forming a border 12 cells wide;
<emphasis id="05CFEAB0FF9DFFCDF5DFFED72BE1FEDA" bold="true" box="[920,993,340,365]" pageId="5" pageNumber="108">costae</emphasis>
ending below apex.
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9DFFCDF6FAFEFB2B0EFDDE" pageId="5" pageNumber="108" type="description">
<paragraph id="370436A2FF9DFFCDF6FAFEFB2B0EFDDE" blockId="5.[136,1452,160,1193]" pageId="5" pageNumber="108">
<emphasis id="05CFEAB0FF9DFFCDF6FAFEFB293DFE26" bold="true" box="[189,317,376,401]" pageId="5" pageNumber="108">Polyoicous</emphasis>
.
<emphasis id="05CFEAB0FF9DFFCDF70CFEFB298BFE26" bold="true" box="[331,395,376,401]" pageId="5" pageNumber="108">Setae</emphasis>
<quantity id="F0439B47FF9DFFCDF7D5FEFB29FEFE26" box="[402,510,376,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="5" pageNumber="108" unit="mm" value="12.0" valueMax="16.0" valueMin="8.0">816 mm</quantity>
long, straight to slightly curved, reddish-brown.
<emphasis id="05CFEAB0FF9DFFCDF263FEFB2C8FFE26" bold="true" box="[1060,1167,376,401]" pageId="5" pageNumber="108">Capsules</emphasis>
erect to inclined, weakly gibbous, zygomorphic, oblong-pyriform, 1.52.1 ×
<quantity id="F0439B47FF9DFFCDF493FE1F2B5EFE02" box="[724,862,412,437]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="5" pageNumber="108" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
, weakly constricted below mouth when dry, yellow to reddish-brown at maturity, with a well differentiated neck ca. ½ total length of capsule;
<emphasis id="05CFEAB0FF9DFFCDF23DFE432CC8FE6E" bold="true" box="[1146,1224,448,473]" pageId="5" pageNumber="108">mouth</emphasis>
slightly oblique, ca. ¾ diameter of capsule;
<emphasis id="05CFEAB0FF9DFFCDF7D0FE672A4BFE4A" bold="true" box="[407,587,484,509]" pageId="5" pageNumber="108">exothecial cells</emphasis>
3253 × 1523 μm, in cross-section with thick, strongly cuneate anticlinal walls, 45 rows of oblate cells at mouth.
<emphasis id="05CFEAB0FF9DFFCDF40AFD8B2ABDFD96" bold="true" box="[589,701,520,545]" pageId="5" pageNumber="108">Opercula</emphasis>
not seen.
<emphasis id="05CFEAB0FF9DFFCDF56CFD8B2BA3FD96" bold="true" box="[811,931,520,545]" pageId="5" pageNumber="108">Peristome</emphasis>
single (possibly double, see below);
<emphasis id="05CFEAB0FF9DFFCDF37AFD8B28C3FDF2" bold="true" pageId="5" pageNumber="108">exostome teeth</emphasis>
straight, regular in shape, orange, to 230 μm high, ca. 60 μm wide at base, striate, trabeculate.
<emphasis id="05CFEAB0FF9DFFCDF29AFDAF2D2DFDF2" bold="true" box="[1245,1325,556,581]" pageId="5" pageNumber="108">Spores</emphasis>
2528 μm, tetrahedral, coarsely verrucate-lirate.
<emphasis id="05CFEAB0FF9DFFCDF46CFDD32AA3FDDE" bold="true" box="[555,675,592,617]" pageId="5" pageNumber="108">Calyptrae</emphasis>
not seen.
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9DFFCDF6FAFDF729D9FD62" pageId="5" pageNumber="108" type="materials_examined">
<paragraph id="370436A2FF9DFFCDF6FAFDF729D9FD62" blockId="5.[136,1452,160,1193]" pageId="5" pageNumber="108">
<emphasis id="05CFEAB0FF9DFFCDF6FAFDF72A43FD3A" bold="true" box="[189,579,628,653]" pageId="5" pageNumber="108">Additional specimens examined:</emphasis>
<collectingCountry id="4FAC7632FF9DFFCDF40BFDF72ADEFD3A" box="[588,734,628,653]" name="Tanzania" pageId="5" pageNumber="108">TANZANIA</collectingCountry>
.
<collectingRegion id="F57FF840FF9DFFCDF4AEFDF72B5BFD3A" box="[745,859,628,653]" country="Tanzania" name="Morogoro" pageId="5" pageNumber="108">Morogoro</collectingRegion>
Districtrict: South-Uluguru Mts., in gorge of Mgeta River Falls at East edge of Lukwangule Plateau,
<date id="43051062FF9DFFCDF4F1FD1B2B44FD06" box="[694,836,664,689]" pageId="5" pageNumber="108" value="1988-06-09">9 June 1988</date>
,
<emphasis id="05CFEAB0FF9DFFCDF517FD1B2DACFD06" box="[848,1452,664,689]" italics="true" pageId="5" pageNumber="108">
<collectionCode id="51AAAE67FF9DFFCDF517FD1B2B61FD06" box="[848,865,664,689]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="108">T</collectionCode>
. Pócs,
<collectionCode id="51AAAE67FF9DFFCDF5F4FD1B2BC7FD06" box="[947,967,664,689]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="108">R</collectionCode>
. Ochyra &amp;
<collectionCode id="51AAAE67FF9DFFCDF209FD1B2C66FD06" box="[1102,1126,664,689]" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" pageId="5" pageNumber="108" type="Herbarium">H</collectionCode>
. BednarekOchyra 88108/
<collectionCode id="51AAAE67FF9DFFCDF3D3FD1B2DACFD06" box="[1428,1452,664,689]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="5" pageNumber="108" type="Herbarium">M</collectionCode>
</emphasis>
(
<collectionCode id="51AAAE67FF9DFFCDF6D7FD3F28CCFD62" box="[144,204,700,725]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15865" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="5" pageNumber="108" type="Herbarium">BOL</collectionCode>
,
<collectionCode id="51AAAE67FF9DFFCDF690FD3F28EAFD62" box="[215,234,700,725]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="5" pageNumber="108" type="Herbarium">E</collectionCode>
,
<collectionCode id="51AAAE67FF9DFFCDF6B0FD3F2932FD62" box="[247,306,700,725]" country="Hungary" httpUri="http://grbio.org/cool/39sm-y0v8" name="Eszterházy Károly College" pageId="5" pageNumber="108" type="Herbarium">EGR</collectionCode>
,
<collectionCode id="51AAAE67FF9DFFCDF77AFD3F296DFD62" box="[317,365,700,725]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="5" pageNumber="108" type="Herbarium">MO</collectionCode>
,
<collectionCode id="51AAAE67FF9DFFCDF73FFD3F29AEFD62" box="[376,430,700,725]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="5" pageNumber="108" type="Herbarium">PRE</collectionCode>
,
<collectionCode id="51AAAE67FF9DFFCDF7FEFD3F29CBFD62" box="[441,459,700,725]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="5" pageNumber="108" type="Herbarium">S</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="7FA16529FF9DFFCDF6FAFD6328CFFB1E" pageId="5" pageNumber="108" type="etymology">
<paragraph id="370436A2FF9DFFCDF6FAFD632DACFD4E" blockId="5.[136,1452,160,1193]" box="[189,1452,736,761]" pageId="5" pageNumber="108">
<emphasis id="05CFEAB0FF9DFFCDF6FAFD632948FD4E" bold="true" box="[189,328,736,761]" pageId="5" pageNumber="108">Etymology:</emphasis>
The species epithet is derived from the presence of zygomorphic capsules and limbate leaf margins.
</paragraph>
<paragraph id="370436A2FF9DFFCDF6FAFC872D57FB8A" blockId="5.[136,1452,160,1193]" pageId="5" pageNumber="108">
<emphasis id="05CFEAB0FF9DFFCDF6FAFC8728FFFCAA" bold="true" box="[189,255,772,797]" pageId="5" pageNumber="108">Note:</emphasis>
The strongly limbate, dentate leaves and zygomorphic, gibbous capsules clearly distinguish
<taxonomicName id="F0BB4D21FF9DFFCDF2BDFC872DACFCAA" authorityName="N. Wilding" authorityYear="2017" box="[1274,1452,772,797]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="5" pageNumber="108" phylum="Bryophyta" rank="species" species="zygolimbatus">
<emphasis id="05CFEAB0FF9DFFCDF2BDFC872DACFCAA" box="[1274,1452,772,797]" italics="true" pageId="5" pageNumber="108">E. zygolimbatus</emphasis>
</taxonomicName>
from other African species. The peristome morphology of this species, in particular exostome shape and ornamentation, and whether or not an endostome is present, remains slightly obscure because of a lack of well-preserved capsules in the only two known collections. The species is quite possibly a close relative of the Andean-Australasian species
<taxonomicName id="F0BB4D21FF9DFFCDF3D4FCF32A97FC1A" authority="Hedwig (1801: 73) Muller (1848: 122)" authorityName="Hedwig (1801: 73) Muller" authorityYear="1848" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="5" pageNumber="108" phylum="Bryophyta" rank="species" species="radians">
<emphasis id="05CFEAB0FF9DFFCDF3D4FCF328DEFC1A" italics="true" pageId="5" pageNumber="108">E. radians</emphasis>
<bibRefCitation id="532A4B53FF9DFFCDF6A3FC1729B7FC1A" author="Hedwig, J." box="[228,439,916,941]" pageId="5" pageNumber="108" refId="ref4018" refString="Hedwig, J. (1801) Species Muscorum Frondosorum. Ed. F. Schwaegrichen, Leipzig, 352 pp." type="book" year="1801">Hedwig (1801: 73)</bibRefCitation>
<bibRefCitation id="532A4B53FF9DFFCDF7F9FC172A97FC1A" author="Muller, C." box="[446,663,916,941]" pageId="5" pageNumber="108" refId="ref4433" refString="Muller, C. (1848) Funarioideae. In: Synopsis Muscorum Frondosorum omnium hucusque Cognitorum 1. A. Foerstner, Berlin, 812 pp." type="book" year="1848">Müller (1848: 122)</bibRefCitation>
</taxonomicName>
. The two species share a similar overall morphology, of which the only major difference is in the ornamentation of their spores (baculate-insulate in
<taxonomicName id="F0BB4D21FF9DFFCDF598FC3B2C53FC66" authorityName="Hedwig (1801: 73) Muller" authorityYear="1848" box="[991,1107,952,977]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="5" pageNumber="108" phylum="Bryophyta" rank="species" species="radians">
<emphasis id="05CFEAB0FF9DFFCDF598FC3B2C53FC66" box="[991,1107,952,977]" italics="true" pageId="5" pageNumber="108">E. radians</emphasis>
</taxonomicName>
vs. coarsely verrucate-lirate in
<taxonomicName id="F0BB4D21FF9DFFCDF6CFFC5F293BFC42" authorityName="N. Wilding" authorityYear="2017" box="[136,315,988,1013]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="5" pageNumber="108" phylum="Bryophyta" rank="species" species="zygolimbatus">
<emphasis id="05CFEAB0FF9DFFCDF6CFFC5F293BFC42" box="[136,315,988,1013]" italics="true" pageId="5" pageNumber="108">E. zygolimbatus</emphasis>
</taxonomicName>
), the two species also share similar high elevation habitats.
<taxonomicName id="F0BB4D21FF9DFFCDF593FC5F2C86FC42" authorityName="N. Wilding" authorityYear="2017" box="[980,1158,988,1013]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="5" pageNumber="108" phylum="Bryophyta" rank="species" species="zygolimbatus">
<emphasis id="05CFEAB0FF9DFFCDF593FC5F2C86FC42" box="[980,1158,988,1013]" italics="true" pageId="5" pageNumber="108">E. zygolimbatus</emphasis>
</taxonomicName>
can also be confused with
<taxonomicName id="F0BB4D21FF9DFFCDF6CFFB832AC9FBAE" authority="Muller (1845: 697) Muller (1848: 126)" authorityName="Muller (1845: 697) Muller" authorityYear="1848" box="[136,713,1024,1049]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="5" pageNumber="108" phylum="Bryophyta" rank="species" species="curvipes">
<emphasis id="05CFEAB0FF9DFFCDF6CFFB832907FBAE" box="[136,263,1024,1049]" italics="true" pageId="5" pageNumber="108">E. curvipes</emphasis>
<bibRefCitation id="532A4B53FF9DFFCDF749FB8329E6FBAE" author="Muller, C." box="[270,486,1024,1049]" pageId="5" pageNumber="108" refId="ref4416" refString="Muller, C. (1845) einer Flora de Aequinoctial-Gegenden. Linnaea 18: 697." type="journal volume" year="1845">Müller (1845: 697)</bibRefCitation>
<bibRefCitation id="532A4B53FF9DFFCDF7A9FB832AC9FBAE" author="Muller, C." box="[494,713,1024,1049]" pageId="5" pageNumber="108" refId="ref4433" refString="Muller, C. (1848) Funarioideae. In: Synopsis Muscorum Frondosorum omnium hucusque Cognitorum 1. A. Foerstner, Berlin, 812 pp." type="book" year="1848">Müller (1848: 126)</bibRefCitation>
</taxonomicName>
, however, the latter differs in having leaves with a weaker border of 12 cells wide, significantly longer peristome teeth (to 450 μm) and highly-thickened exothecial cell walls.
</paragraph>
<paragraph id="370436A2FF9DFFCDF6FAFBCB28CFFB1E" blockId="5.[136,1452,160,1193]" pageId="5" pageNumber="108">
<emphasis id="05CFEAB0FF9DFFCDF6FAFBCB29E8FBD6" bold="true" box="[189,488,1096,1121]" pageId="5" pageNumber="108">Habitat and distribution:</emphasis>
The species is known from only two localities, one in
<collectingCountry id="4FAC7632FF9DFFCDF27EFBCB2CC6FBD6" box="[1081,1222,1096,1121]" name="South Africa" pageId="5" pageNumber="108">South Africa</collectingCountry>
and one in
<collectingCountry id="4FAC7632FF9DFFCDF307FBCB2DA8FBD6" box="[1344,1448,1096,1121]" name="Tanzania" pageId="5" pageNumber="108">Tanzania</collectingCountry>
.
<taxonomicName id="F0BB4D21FF9DFFCDF6CFFBEF29B6FB32" authorityName="N. Wilding" authorityYear="2017" box="[136,438,1132,1157]" class="Bryopsida" family="Funariaceae" genus="Entosthodon" kingdom="Plantae" order="Funariales" pageId="5" pageNumber="108" phylum="Bryophyta" rank="species" species="zygolimbatus">
<emphasis id="05CFEAB0FF9DFFCDF6CFFBEF29B6FB32" box="[136,438,1132,1157]" italics="true" pageId="5" pageNumber="108">Entosthodon zygolimbatus</emphasis>
</taxonomicName>
is recorded from elevations between
<quantity id="F0439B47FF9DFFCDF530FBEF2BD1FB32" box="[887,977,1132,1157]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="108" unit="m" value="2000.0">2000 m</quantity>
and
<quantity id="F0439B47FF9DFFCDF256FBEF2C6BFB32" box="[1041,1131,1132,1157]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="108" unit="m" value="2400.0">2400 m</quantity>
in tropical and subtropical zones.
</paragraph>
</subSubSection>
</treatment>
</document>