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Questionable status. Reported from Greece by
<bibRefCitation id="EEDAB0348B275DDA9EA62097472CBBA7" pageId="0" pageNumber="20997">Akoumianaki and Hughes (1997)</bibRefCitation>
, also found by
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(unpublished data), but
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is a taxonomically confused species.
<bibRefCitation id="A884F17D532E7495C6CD20E368342044" author="Riser, Nathan W" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="0" pageNumber="20997" pagination="514 - 225" title="Podarkeaberrans Webster &amp; Benedict, 1887 - resolution, with description of two new species in the genus Microphthalmus (Annelida: Polychaeta)" volume="113" year="2000">Riser (2000)</bibRefCitation>
revised the species and found the slides of the type material to be comprised of two different species. He assigned specimens previously assigned to
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to three species:
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,
<taxonomicName id="D750721C44F1AE1E7322491B50232086" class="Polychaeta" family="Microphthalmidae" genus="Microphthalmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microphthalmus pettiboneae" order="Phyllodocida" pageId="0" pageNumber="20997" phylum="Annelida" rank="species" species="pettiboneae">Microphthalmus pettiboneae</taxonomicName>
Riser, 2000 and
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Riser, 2000. However, the species re-described as
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by Riser does not resemble previously available descriptions of the species, as those were mostly copied from a re-description by
<bibRefCitation id="2EE7E9078353897BD40B6D43D85EAE57" author="Southern, Rowland" journalOrPublisher="Proceedings of the Royal Irish Academy" pageId="0" pageNumber="20997" pagination="1 - 160" title="Clare Island Survey. Archiannelida and Polychaeta" volume="31" year="1914">Southern (1914)</bibRefCitation>
, which was based on the wrong microscope slide. The specimen on this slide was re-described as
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by Riser, causing confusion in the application of the name
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. All specimens reported under this name would need to be re-investigated to clarify their identity.
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Tribe
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Newton and Thayer, 1995: 297 [stem: Silphotel-]. Type genus:
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Broun, 1895.
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.531.6097" ID-GBIF-Dataset="70702787-a6a8-4af7-9702-738f344a5cbe" ID-PMC="PMC4668911" ID-Pensoft-Pub="1313-2970-531-1" ID-PubMed="26692803" ID-ZBK="C845838EC9124BD8AB4E07980F91959E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-531-1" ModsDocOrigin="ZooKeys 531" ModsDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" checkinTime="1451243834596" checkinUser="pensoft" docAuthor="Vermeulen, Jaap J., Liew, Thor-Seng &amp; Schilthuizen, Menno" docDate="2015" docId="2DF4B7547F1732295642312A1CE2983F" docLanguage="en" docName="ZooKeys 531: 1-139" docOrigin="ZooKeys 531" docSource="http://dx.doi.org/10.3897/zookeys.531.6097" docTitle="Philalanka malimgunung Vermeulen, Liew &amp; Schilthuizen, sp. n." docType="treatment" docUuid="33E06180-04E1-4A2B-8C80-41DB4216F799" docUuidSource="ZooBank" docVersion="10" lastPageNumber="60" masterDocId="1527DF5A6645C540FFF1FF85AF4AE358" masterDocTitle="Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species" masterLastPageNumber="139" masterPageNumber="1" pageNumber="60" updateTime="1668160884494" updateUser="ExternalLinkService">
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<mods:title>Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liew, Thor-Seng</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>531</mods:number>
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<mods:start>1</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.531.6097</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.531.6097</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-531-1</mods:identifier>
<mods:identifier type="ZBK">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
<mods:identifier type="ZooBank">C845838EC9124BD8AB4E07980F91959E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="144057521" LSID="urn:lsid:zoobank.org:act:33E06180-04E1-4A2B-8C80-41DB4216F799" httpUri="http://treatment.plazi.org/id/2DF4B7547F1732295642312A1CE2983F" lastPageNumber="60" pageId="59" pageNumber="60">
<subSubSection pageId="59" pageNumber="60" type="multiple">
<paragraph pageId="59" pageNumber="60">Taxon classification Animalia Stylommatophora Endodontidae</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="nomenclature">
<paragraph pageId="59" pageNumber="60">
<taxonomicName LSID="http://zoobank.org/33E06180-04E1-4A2B-8C80-41DB4216F799" authority="Vermeulen, Liew &amp; Schilthuizen" class="Gastropoda" family="Charopidae" genus="Philalanka" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philalanka malimgunung" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="malimgunung">Philalanka malimgunung Vermeulen, Liew &amp; Schilthuizen</taxonomicName>
<taxonomicNameLabel pageId="59" pageNumber="60">sp. n.</taxonomicNameLabel>
Figure 49
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="holotype">
<paragraph pageId="59" pageNumber="60">
<materialsCitation collectingDate="2015-01-01" collectingDateMax="2015-12-31" collectingDateMin="2015-01-01" collectionCode="RMNH" country="Malaysia" httpUri="http://data.biodiversitydata.nl/naturalis/specimen/RMNH.5003942" location="West Coast Province" municipality="Sabah" specimenCode="RMNH.5003942" specimenCount="1" typeStatus="holotype">
<typeStatus>Holotype</typeStatus>
.
<collectingCountry>Malaysia</collectingCountry>
,
<collectingMunicipality>Sabah</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:2DF4B7547F1732295642312A1CE2983F:39DEFB746E67DBCCBD4C3557857450C1" country="Malaysia" municipality="Sabah" name="West Coast Province">West Coast Province</location>
,
<location LSID="urn:lsid:plazi:treatment:2DF4B7547F1732295642312A1CE2983F:37CD92810CED3614A77EBF2F972CC5E3" country="Malaysia" municipality="Sabah" name="Kinabalu N. P.">Kinabalu N.P.</location>
,
<location LSID="urn:lsid:plazi:treatment:2DF4B7547F1732295642312A1CE2983F:380C531D985B2C76CDC02DFDD2EAB812" country="Malaysia" municipality="Sabah" name="summit trail">summit trail</location>
(
<specimenCode>RMNH.5003942</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="materials_examined">
<paragraph pageId="59" pageNumber="60">Examined material from Sabah.</paragraph>
<paragraph pageId="59" pageNumber="60">West Coast Province. Kinabalu N.P., Summit trail between 3119 and 4080 m (leg. T.S. Liew, BOR/MOL 4192, BOR/MOL 4193, BOR/MOL 4195, BOR/MOL 4196,V 14336, V 14337, V 14338).</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="description">
<paragraph pageId="59" pageNumber="60">Description.</paragraph>
<paragraph pageId="59" pageNumber="60">Shell minute, very thin, hardly translucent, pale yellow-brown, yellow-corneous below, (low-)conical with approx. flat sides; apex rounded. Surface with a silky luster. Whorls convex, rounded (angular in juveniles), suture somewhat impressed. Protoconch whorls moderately convex; with inconspicuous, very fine (just visible at 40 times magnification) and dense spiral striation. Teleoconch with very fine (just visible at 40 times magnification), dense spiral striation above the periphery, which is continuous on the top whorls but more patchy on the last whorl; with a similar but somewhat more widely spaced striation below the periphery; spiral striation somewhat subordinate to the rather distinct, irregularly spaced, slightly raised growth lines, which are less conspicuous towards the umbilicus. Umbilicus open, rather narrow. Dimensions: Height up to 1.4 mm; width up to 1.9 mm; diameters of the first three whorls 0.45-0.50 mm, 0.85-1.00 mm, 1.40-1.55 mm respectively; number of whorls up to c. 3 1/2; height aperture up to 0.8 mm; width aperture up to 0.9 mm.</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="habitat in sabah and distribution">
<paragraph pageId="59" pageNumber="60">Habitat in Sabah and distribution.</paragraph>
<paragraph pageId="59" pageNumber="60">Subalpine vegetation on granodiorite soil. Alt. 3200-4100 m. Sabah: Mount Kinabalu (summit trail area). Endemic to Sabah.</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="cross diagnosis">
<paragraph pageId="59" pageNumber="60">Cross diagnosis.</paragraph>
<paragraph pageId="59" pageNumber="60">
Most similar in size and shape to
<taxonomicName class="Gastropoda" family="Charopidae" genus="Philalanka" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philalanka anomphala" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="anomphala">Philalanka anomphala</taxonomicName>
, it differs by having an open umbilicus, and by lacking the peripheral thread. It also differs from the extralimital species listed under
<taxonomicName class="Gastropoda" family="Charopidae" genus="Philalanka" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philalanka rugulosa" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="rugulosa">Philalanka rugulosa</taxonomicName>
by the lack of a peripheral thread.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="etymology">
<paragraph pageId="59" pageNumber="60">Etymology.</paragraph>
<paragraph pageId="59" pageNumber="60">
The species is named in recognition of generations of mountain guides ('Malim
<normalizedToken originalValue="Gunung" pageId="59" pageNumber="60">Gunung'</normalizedToken>
) who assist tourists and scientists on their way to the peaks of Mount Kinabalu every day, and who have aided scientific discoveries on this mountain since the first documented exploration of Mount Kinabalu in 1851. The species name also serves to honour the guides who lost their lives protecting mountain climbers when Mount Kinabalu was struck by earthquake on June 5th, 2015.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="2DF4D4674AF39CAC183DF3C1215A6687" ID-DOI="http://doi.org/10.5281/zenodo.6216839" ID-GBIF-Taxon="100121216" ID-Zenodo-Dep="6216839" LSID="urn:lsid:zoobank.org:act:AD6BD7CD-1F39-4950-B384-3840A324F0E1" httpUri="http://treatment.plazi.org/id/2DF4D4674AF39CAC183DF3C1215A6687" lastPageNumber="24" pageNumber="24">
<subSubSection id="047F3C96E534F23066076F9B937583B7" type="nomenclature">
<paragraph id="DC05A53EBA455E7F1D02AF2F51557A50" pageNumber="24">
<taxonomicName id="8860115D3FE26589FBBBDA9E634C90DE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253441" class="Insecta" family="Formicidae" genus="Euprenolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Euprenolepis zeta" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="zeta">
<pageStartToken id="82B63F4ADF4C395E4899355914FBFF1E" pageNumber="24">Euprenolepis</pageStartToken>
zeta
</taxonomicName>
,
<taxonomicNameLabel id="C86096ECD6A418C851375A170EEB23F5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F43477E9C8E64B91F9BCF485555CB29D" type="description">
<paragraph id="687F0964FC2B8097A7F869BA60DAA56E" pageNumber="24">(Figure 14A-D)</paragraph>
</subSubSection>
<subSubSection id="CB8B152F5F3ED875E112A2CDBACA4EC4" type="materials_examined">
<paragraph id="A9DD86E558C9A7D1936ED88087DC5668" pageNumber="24">
Holotype worker, MALAYSIA: Borneo, Sabah, Sepilok Forest Reserve, 60 m, ii.1999 (C.
<normalizedToken id="EA4721F88BEB1DE7A541F11FE019D7CF" originalValue="Brühl">Bruehl</normalizedToken>
) (
<collectionCode id="A01A3586FF313EB9B86F5B1A66E99B04" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
); 1 paratype worker, same locality as holotype (
<collectionCode id="794AECFF87DEF62DC0D3E86FF2945F93" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection">ANIC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="70C6831525A10D7EA880BCE016504A9A" type="description">
<paragraph id="5DEF10F57D110B642B1002FCBBB63715" pageNumber="24">Worker. Measurements (n=3): TL: 2.69-3.22; HW: 0.77-0.86; HL: 0.76-0.86; SL: 0.98-1.03; WL: 1.02-1.16; GL: 0.86-1.2. Indices: CI: 99-104; SI: 116-130</paragraph>
<paragraph id="514F2B13C5E2F84945567BAF58CE90E8" pageNumber="24">Head brown, with lighter yellowish-brown mandibles and funiculi; head about as long as broad. Cuticular surface shiny and smooth, with scattered erect setae and sparse pubescence that is thickest in area under the eyes. Posterior margin complete. Scapes surpass posterior margin by about length of the first four funicular segments. Scapes with erect setae and a layer of pubescence, becoming denser distally. Mesosoma brown, smooth and shiny, legs lighter, especially tarsi, which become yellowish-brown. Mesosomal dorsum with scattered erect setae of varying lengths; cuticular surface without pubescence. Pronotum rises at about 45° toward mesonotum; propodeum dorsum rounded, dome-like; declivity short, but steep. Petiole triangular with posterior face longer than anterior face; gaster brown, with scattered erect setae and a sparse layer of pubscence; cuticular surface shiny and weakly reguoreticulate.</paragraph>
</subSubSection>
<subSubSection id="2B9B7EE045E7E0231AED6EAA17D6B3A4" type="etymology">
<paragraph id="35311E5A8043F4710B6D31B23765C568" pageNumber="24">
Etymology: The species epithet is Greek. Zeta is the sixth letter of the Greek alphabet, and this was the sixth and final
<taxonomicNameLabel id="23664CC7E7957A1EAF0A983BA74A84DA" rank="species">new species</taxonomicNameLabel>
discovered during the course of this revision.
</paragraph>
</subSubSection>
<subSubSection id="8D0A92408A92224F075E7A9169C9CD66" type="materials_examined">
<paragraph id="7288D87E1359E7115B5A824BC27F60F3" pageNumber="24">
Additional material examined: MALAYSIA: Borneo, Sabah, Danum Valley, West Trail P1, 250 m, 17.viii.1997 (C.
<normalizedToken id="F06CA546C56D3B1BA4848D71BA41330C" originalValue="Brühl">Bruehl</normalizedToken>
); Borneo; Kinabalu Nat. Park; Poring Hot Spring, East Ridge; 560 m (C.
<normalizedToken id="6A2A822444DDCB8CC6E54937F7A3B39D" originalValue="Brühl">Bruehl</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection id="3B19F1A7112842C202FC536B2CE7E582" type="discussion">
<paragraph id="7B4E32390158B2173196ACFC0441D9F1" pageNumber="24">
Discussion: This species is most likely to be confused with
<taxonomicName id="2C4C360CAD73261DDFA419FF55D210B5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253438" class="Insecta" family="Formicidae" genus="Euprenolepis" genus.bestMatchDistance="2140" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Euprenolepis thrix" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="thrix">E. thrix</taxonomicName>
, but
<taxonomicName id="12E4334BA14AA4042F796E27A2DF7606" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253441" class="Insecta" family="Formicidae" genus="Euprenolepis" genus.bestMatchDistance="370" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Euprenolepis zeta" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="zeta">E. zeta</taxonomicName>
has far less pubescence on the gaster. As in
<taxonomicName id="BCD3B521FC635C911F1FF00A6FE29850" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253438" class="Insecta" family="Formicidae" genus="Euprenolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Euprenolepis thrix" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="thrix">E. thrix,</taxonomicName>
<taxonomicName id="C5DFD04D79B92FCCD9D9869150262D30" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253441" class="Insecta" family="Formicidae" genus="Euprenolepis" genus.bestMatchDistance="44" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Euprenolepis zeta" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="zeta">E. zeta</taxonomicName>
does have the head wider than long, an unusual trait among
<taxonomicName id="E3FF5823BAD2E238B367A4F392A86584" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24771" class="Insecta" family="Formicidae" genus="Euprenolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Euprenolepis Emery" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="genus">Euprenolepis</taxonomicName>
species. Based on overall morphological similarity, including the head shape and presence of pubescence on the gaster, it is likely that
<taxonomicName id="F4F3BDA2F24FA578C7B11F846A33D22D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253438" class="Insecta" family="Formicidae" genus="Euprenolepis" genus.bestMatchDistance="2202" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Euprenolepis thrix" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="thrix">E. thrix</taxonomicName>
and
<taxonomicName id="B84624D1C458D655F089197CC9484B4D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253441" class="Insecta" family="Formicidae" genus="Euprenolepis" genus.bestMatchDistance="431" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Euprenolepis zeta" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="zeta">E. zeta</taxonomicName>
are close relatives.
</paragraph>
</subSubSection>
</treatment>
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