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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.101.1133" ID-GBIF-Dataset="75c91bde-91db-46c1-bafd-9eaa41db4380" ID-PMC="PMC3118702" ID-Pensoft-Pub="1313-2970-101-1" ID-PubMed="21747669" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-101-1" ModsDocOrigin="ZooKeys 101" ModsDocTitle="Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species" checkinTime="1451250351161" checkinUser="pensoft" docAuthor="Breure, Abraham S. H." docDate="2011" docId="2DD418E31ECD6A61B23807CBE5F15983" docLanguage="en" docName="ZooKeys 101: 1-50" docOrigin="ZooKeys 101" docSource="http://dx.doi.org/10.3897/zookeys.101.1133" docTitle="Thaumastus nystianus subsp. nigricans Cousin 1887" docType="treatment" docVersion="4" lastPageNumber="21" masterDocId="FFE6FFCFFF94FFD6FF9DFFFFFFD0FF80" masterDocTitle="Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species" masterLastPageNumber="50" masterPageNumber="1" pageNumber="21" updateTime="1668149715901" updateUser="ExternalLinkService">
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<mods:title>Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Breure, Abraham S. H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>101</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>50</mods:end>
</mods:extent>
</mods:part>
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<mods:url>http://dx.doi.org/10.3897/zookeys.101.1133</mods:url>
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<mods:identifier type="Pensoft-Pub">1313-2970-101-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152029444" LSID="urn:lsid:plazi:treatment:2DD418E31ECD6A61B23807CBE5F15983" httpUri="http://treatment.plazi.org/id/2DD418E31ECD6A61B23807CBE5F15983" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection pageId="20" pageNumber="21" type="nomenclature">
<paragraph pageId="20" pageNumber="21">
<taxonomicName LSID="http://species-id.net/wiki/Thaumastus_nystianus_nigricans" authority="Cousin, 1887" authorityName="Cousin" authorityYear="1887" class="Gastropoda" family="Megaspiridae" genus="Thaumastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thaumastus nystianus" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="subSpecies" species="nystianus" subSpecies="nigricans">Thaumastus nystianus nigricans Cousin, 1887</taxonomicName>
Figs 12B, 12ii
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="reference_group">
<paragraph pageId="20" pageNumber="21">
<taxonomicName class="Gastropoda" family="Megaspiridae" genus="Thaumastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thaumastus nystianus" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="nystianus">Thaumastus nystianus</taxonomicName>
var. nigricans
<bibRefCitation author="Cousin, A" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="28" pageNumber="29" pagination="187 - 287" title="Faune malacologique de la republique de l'Equateur." volume="12" year="1887">Cousin 1887</bibRefCitation>
: 220.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="type locality">
<paragraph pageId="20" pageNumber="21">Type locality.</paragraph>
<paragraph pageId="20" pageNumber="21">
[Ecuador, Prov. Pichincha] &quot;les bords du chemin qui conduit de Pomasqui
<normalizedToken originalValue="à">a</normalizedToken>
Chilguiltina&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="label">
<paragraph pageId="20" pageNumber="21">Label.</paragraph>
<paragraph pageId="20" pageNumber="21">
&quot;chemin de Pomasqui
<normalizedToken originalValue="à">a</normalizedToken>
Chilguiltina&quot; in
<normalizedToken originalValue="Dautzenberg’s">Dautzenberg's</normalizedToken>
handwriting; see remarks.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="dimensions">
<paragraph pageId="20" pageNumber="21">Dimensions.</paragraph>
<paragraph pageId="20" pageNumber="21">Not given. Figured specimen H 37.3, D 17.7, W 6.3.</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="type material">
<paragraph pageId="20" pageNumber="21">Type material.</paragraph>
<paragraph pageId="20" pageNumber="21">RBINS/MT2365, 13 possible syntypes, ex Cousin (Dautzenberg coll.).</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="remarks">
<paragraph pageId="20" pageNumber="21">Remarks.</paragraph>
<paragraph pageId="20" pageNumber="21">
The material is not accompanied by an original Cousin label, but it originates from Cousin; the specimens are treated herein as possible syntypes. This taxon is now considered a junior subjective synonym of
<taxonomicName class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus nystianus" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="nystianus">Bulimus nystianus</taxonomicName>
Pfeiffer, 1853 (syn. n.).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="21" type="current systematic position">
<paragraph pageId="20" pageNumber="21">Current systematic position.</paragraph>
<paragraph pageId="20" pageNumber="21">
<taxonomicName family="Bulimulidae" lsidName="" pageId="20" pageNumber="21" rank="family">Bulimulidae</taxonomicName>
,
<taxonomicName class="Gastropoda" family="Bulimulidae" genus="Drymaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymaeus (Drymaeus) nystianus" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="nystianus" subGenus="Drymaeus">Drymaeus (Drymaeus) nystianus</taxonomicName>
(Pfeiffer, 1853).
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="2DD47A9A17A65718AB06FC5EC21ACCED" ID-GBIF-Taxon="182233273" LSID="urn:lsid:plazi:treatment:2DD47A9A17A65718AB06FC5EC21ACCED" httpUri="http://treatment.plazi.org/id/2DD47A9A17A65718AB06FC5EC21ACCED" lastPageNumber="110" pageId="109" pageNumber="110">
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<paragraph id="DF4F64CB1D61096407405656EBCF61FD" pageId="109" pageNumber="110">
†
<taxonomicName id="00845A4DF0D4D466DF8B860D3F94B4F3" ID-CoL="6R94C" LSID="2DD47A9A-17A6-5718-AB06-FC5EC21ACCED" authority="Handmann, 1887" authorityName="Handmann" authorityYear="1887" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Melanopsis (Martinia) vindobonensis subsp. var. var. elongata" order="Sorbeoconcha" pageId="109" pageNumber="110" phylum="Mollusca" rank="variety" species="vindobonensis" status="[invalid]" subGenus="Martinia" subSpecies="var." variety="elongata">Melanopsis (Martinia) vindobonensis var. elongata Handmann, 1887</taxonomicName>
<taxonomicNameLabel id="DD17AD2A986C2E26B4B2D1E53A168434" pageId="109" pageNumber="110">[invalid]</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C160C34D1441950A663095F94B6190AD" pageId="109" pageNumber="110" type="original source">
<paragraph id="38EB1A7AC1DEF9C9C596A86B2E2582A2" pageId="109" pageNumber="110">Original source.</paragraph>
<paragraph id="46BB38D839C971DF4D50A11BC65B0283" pageId="109" pageNumber="110">
<bibRefCitation id="5A50F66E670FAD6C1DB0557AF4C20A5B" author="Handmann, R" journalOrPublisher="Aschendorff, Muenster" pageId="313" pageNumber="314" refId="B215" refString="Handmann, R, 1887. Die fossile Conchylienfauna von Leobersdorf im Tertiaerbecken von Wien. Aschendorff, Muenster" title="Die fossile Conchylienfauna von Leobersdorf im Tertiaerbecken von Wien." year="1887">Handmann 1887</bibRefCitation>
: 28, pl. 6, figs 3-4.
</paragraph>
</subSubSection>
<subSubSection id="85E3FED4F1DFF445A95CAD88781B25E5" pageId="109" pageNumber="110" type="type horizon">
<paragraph id="3C776E49409A791C221FEFF742550B43" pageId="109" pageNumber="110">Type horizon.</paragraph>
<paragraph id="60D4208388EE8B29905DF662915C0059" pageId="109" pageNumber="110">Pannonian, zone B-D, late Miocene.</paragraph>
</subSubSection>
<subSubSection id="A30606B358AD1FDEB7D52DB7AD6645F1" pageId="109" pageNumber="110" type="type locality">
<paragraph id="A5E6419674720785EC65E24EC8D8EF40" pageId="109" pageNumber="110">Type locality.</paragraph>
<paragraph id="2AC9CB91E84DE0EBC1FA1A4720362162" pageId="109" pageNumber="110">
<normalizedToken id="1238F33828DC7D25DE8A3196A5602297" originalValue="“Leobersdorf”">&quot;Leobersdorf&quot;</normalizedToken>
, Austria.
</paragraph>
</subSubSection>
<subSubSection id="8D643632ACF1895E12F22C2022B2B3F0" pageId="109" pageNumber="110" type="remarks">
<paragraph id="2E6D5D8A524157B5308652AF1E3D90BE" pageId="109" pageNumber="110">Remarks.</paragraph>
<paragraph id="07A4FB3BB614EA18310EBAE9A447FE65" pageId="109" pageNumber="110">
Junior homonym of
<taxonomicName id="04A783C425E9A1194536EE8BD5594385" authorityName="Ferussac" authorityYear="1822" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" lsidName="Melanopsis buccinoidea subsp. elongata" order="Pteropoda" pageId="109" pageNumber="110" phylum="Mollusca" rank="subSpecies" species="buccinoidea" subSpecies="elongata">
<emphasis id="8836B244ECA0AFC6382BEB1D9E833631" italics="true" pageId="109" pageNumber="110">Melanopsis buccinoidea elongata</emphasis>
</taxonomicName>
<normalizedToken id="A14CE191DBF9323B65461E3479DF6EE3" originalValue="Férussac">Ferussac</normalizedToken>
, 1822.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:namePart>Delfim, Fagner Ribeiro</mods:namePart>
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<mods:namePart>Mesquita, Daniel Oliveira</mods:namePart>
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<mods:namePart>Colli, Guarino Rinaldi</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Universidade de Brasilia, 70910 - 900, Brasilia, DF, Brazil</mods:affiliation>
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<mods:role>
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</mods:role>
<mods:namePart>Garda, Adrian Antonio</mods:namePart>
<mods:affiliation>Departamento de Botanica e Zoologia, Universidade Federal do Rio Grande do Norte, 59078 - 900, Natal, RN, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guedes, Thais B.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3318-7193</mods:nameIdentifier>
<mods:affiliation>Departamento de Biologia Animal, Instituto de Biologia, Universidade Estadual de Campinas, 13083 - 862, Campinas, SP, Brazil &amp; University of Gothenburg, Gothenburg Global Biodiversity Center, Department of Biological and Environmental Sciences, Box 461, SE- 405 30, Goeteborg, Sweden</mods:affiliation>
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</mods:name>
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<taxonomicName LSID="2DD4CD9E-8223-5D65-8D4D-F4B658F38DC1" authority="(Rodrigues, 1984)" baseAuthorityName="Rodrigues" baseAuthorityYear="1984" class="Reptilia" family="Tropiduridae" genus="Tropidurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidurus pinima" order="Squamata" pageId="0" pageNumber="599" phylum="Chordata" rank="species" species="Pinima">Tropidurus pinima (Rodrigues, 1984)</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="599" type="description">
<paragraph pageId="0" pageNumber="599">
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lizards recorded in the Caatinga region. 12.1 Tropidurus helenae; 12.2 Tropidurus hispidus; 12.3 Tropidurus jaguaribanus; 12.4 Tropidurus montanus; 12.5 Tropidurus mucujensis; 12.6 Tropidurus oreadicus; 12.7 Tropidurus pinima and 12.8 Tropidurus semitaeniatus. Photograph credits: Daniel Mesquita (12.1, 12.6, 12.8), Adrian Garda (12.2), Daniel Passos (12.3, 12.4, 12.7), Marco A. Freitas (12.5)." figureDoi="10.3897/vz.72.e78828.figure12" httpUri="https://binary.pensoft.net/fig/726820" pageId="0" pageNumber="599">Figs 12.7 and 20</figureCitation>
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<subSubSection pageId="0" pageNumber="599" type="type locality">
<paragraph pageId="0" pageNumber="599">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="599">
Santo
<normalizedToken originalValue="Inácio">Inacio</normalizedToken>
, state of Bahia, Brazil.
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<paragraph pageId="0" pageNumber="599">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="599">
Caatinga endemic species. It is recorded in the the states of Bahia and Pernambuco. It is widespread in the Caatinga and with annual mean temperature 20 to 28°C along two ecoregions (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Lizards from the Caatinga. Summarized data about distribution, ecology, and conservation. Caatinga ecoregions (Silva et al. 2017) - B: Brejos, CCD: Complexo Chapada Diamantina, CIA: Complexo Ibiapaba-Araripe, DSM: Depressao Sertaneja Meridional, DSS: Depressao Sertaneja Setentorial, DSF: Dunas do Sao Francisco, PB: Planalto da Borborema, RC: Raso da Catarina, SFG: Sao Francisco - Gurgueia. Distribution Patterns - WD: widespread (EOO greater than 20,000 km 2, see Material and Methods for details), R: restricted (EOO less than 20,000 km 2). Habitat - O: open areas; F: forested areas; Preferred habitat type: AT: ant hills and termite mounds, B: bromeliads, G: general, LL: leaf litter, P: psammophilous, A: anthropic; S: saxicolous and TB: trees and bushes; Activity - D: diurnal, N: nocturnal and C: crepuscular. Conservation status (IUCN 2021; ICMBio 2018) - Critically Endangered (CR), Data Deficient (DD), Endangered (EN), Least Concern (LC), Near Threatened (NT), Not Evaluated (NE) and Vulnerable (VU). Caatinga endemic species ' * '." httpUri="http://table.plazi.org/id/DCA21B21C4ADC68A0D52968CF9844DE2" pageId="0" pageNumber="599" tableUuid="DCA21B21C4ADC68A0D52968CF9844DE2">1</tableCitation>
; Appendix S3). Distributed in medium to high elevation areas (404-1,154 m a.s.l.), with annual mean temperature 20 to 26°C, and average annual rainfall between 528 and 842 mm.
</paragraph>
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<subSubSection pageId="0" pageNumber="599" type="ecological notes">
<paragraph pageId="0" pageNumber="599">Ecological notes.</paragraph>
<paragraph pageId="0" pageNumber="599">
Terrestrial and diurnal. It is a saxicolous lizard that occurs along open areas in the Caatinga (
<bibRefCitation DOI="https://doi.org/10.2307/1565694" author="Rodrigues, MT" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="599" pagination="513 - 523" refId="B192" refString="Rodrigues, MT, 1996b. Lizards, Snakes, and Amphisbaenians from the Quaternary Sand Dunes of the Middle Rio Sao Francisco, Bahia, Brazil. Journal of Herpetology 30: 513 - 523, DOI: https://doi.org/10.2307/1565694" title="Lizards, Snakes, and Amphisbaenians from the Quaternary Sand Dunes of the Middle Rio Sao Francisco, Bahia, Brazil." url="https://doi.org/10.2307/1565694" volume="30" year="1996 b">Rodrigues 1996b</bibRefCitation>
). Diet based mainly on arthropods, being
<taxonomicName class="Reptilia" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="599" rank="order">Hymenoptera</taxonomicName>
and also Plant material the most representative items (
<bibRefCitation author="Xavier, MA" journalOrPublisher="Herpetology Notes" pageId="0" pageNumber="599" pagination="491 - 492" refId="B274" refString="Xavier, MA, Dias, EJR, 2017. Ecology of endemic and saxicolous lizard Tropidurus pinima (Rodrigues, 1984) from Caatinga domain, Brazil. Herpetology Notes 10: 491 - 492" title="Ecology of endemic and saxicolous lizard Tropidurus pinima (Rodrigues, 1984) from Caatinga domain, Brazil." volume="10" year="2017">Xavier and Dias 2017</bibRefCitation>
). Oviparous, the female usually lays two eggs at a time (
<bibRefCitation author="Xavier, MA" journalOrPublisher="Herpetology Notes" pageId="0" pageNumber="599" pagination="491 - 492" refId="B274" refString="Xavier, MA, Dias, EJR, 2017. Ecology of endemic and saxicolous lizard Tropidurus pinima (Rodrigues, 1984) from Caatinga domain, Brazil. Herpetology Notes 10: 491 - 492" title="Ecology of endemic and saxicolous lizard Tropidurus pinima (Rodrigues, 1984) from Caatinga domain, Brazil." volume="10" year="2017">Xavier and Dias 2017</bibRefCitation>
).
</paragraph>
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</treatment>
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<paragraph pageId="0" pageNumber="11794">Fig. 53</paragraph>
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