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<document id="0C48CB8061C5C717FDC189F0013B7B4C" ENCODING="UTF-8" ID-GBIF-Dataset="1e2bcc49-86c8-4612-a4eb-d9bdce9785ef" ModsDocAuthor="Smith, F." ModsDocDate="1858" ModsDocID="8127" ModsDocOrigin="http://antbase.org/ants/publications/8127/8127.pdf" ModsDocTitle="Catalogue of the hymenopterous insects in the collection of the British Museum. Part VI. Formicidae." checkinTime="1252707217356" checkinUser="donat" docAuthor="Smith, F." docDate="1858" docId="2D610243A37770953A06494D3B9DF863" docLanguage="en" docName="8127.xml" docOrigin="London: British Museum" docSource="http://antbase.org/ants/publications/8127/8127.pdf" docTitle="Plectroctena" docType="treatment" docVersion="17" lastPageNumber="101" masterDocId="913D7CCA2C271185BA478FF390B73E80" masterDocTitle="Catalogue of the hymenopterous insects in the collection of the British Museum. Part VI. Formicidae." pageNumber="101" updateTime="1701312656745" updateUser="plazi">
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Genus 5.
<taxonomicName id="9B6692D97B59A955A5F8B883FC55F23E" ID-CoL="6SC9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24844" class="Insecta" family="Formicidae" genus="Plectroctena" kingdom="Animalia" lsidName-HNS="Plectroctena Smith" order="Hymenoptera" pageId="101" pageNumber="101" phylum="Arthropoda" rank="genus">PLECTROCTENA</taxonomicName>
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<paragraph id="943E99E8EB6332936841BA72F0E5993B" pageId="101" pageNumber="101">Head subquadrate; antennae shorter than the head and thorax; the flagellum clavate, inserted very forward on the face near its anterior margin on each side of a central protuberance, which is advanced over and conceals the clypeus; the anterior margin of the latter widely emarginate; eyes small and ovate, not prominent, situated very forward on the sides of the head near the base of the mandibles; mandibles elongate, very slightly bent and crossing each other, their inner margin with a blunt tooth towards their base; the labial palpi 4-jointed; the maxillary palpi 3- jointed. Thorax oblong; the calcar at the apex of all the tibiae, pectinate within. Abdomen oblong, with a deep strangulation between the first and second segments; the node subglobose.</paragraph>
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This subgenus of
<taxonomicName id="6B4556CAE961A2D23D2FD6B6873899AF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24845" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" lsidName-HNS="Ponera Latreille" order="Hymenoptera" pageId="101" pageNumber="101" phylum="Arthropoda" rank="genus">Ponera</taxonomicName>
differs in such remarkable peculiarities, and will not assimilate with any divisional group yet proposed, that it must stand isolated for the present: the form of the mandibles appears to indicate some peculiar economy; the small advanced eyes are also remarkable: the characters, being drawn from a worker ant, will doubtless require hereafter some additions and modifications. The beautiful pectination of the tibial spurs is common to several of the Poneridae, but is sometimes only to be observed on the anterior legs, as in
<taxonomicName id="F933F4CB7502517CC223769B1BACC114" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2493" class="Insecta" family="Formicidae" genus="Ectatomma" kingdom="Animalia" lsidName-HNS="Ectatomma Smith" order="Hymenoptera" pageId="101" pageNumber="101" phylum="Arthropoda" rank="genus">Ectatomma</taxonomicName>
.
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</treatment>
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,
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<paragraph id="60C9D098B482B3DF788CC895CADCDFC3" pageNumber="13">AUM 18946, 4, AUM 22836, 2, MCNG 9721, 1, MCNG 24200, 1, MCNG 37452, 1, MCNG 38972, 1, MCNG 42665, 1, UF 32382, 2, USNM 258213, 1, USNM 258215, 1,;</paragraph>
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18.
<taxonomicName id="9D60BC4FC9D6B5391C26BCF0D4FE2A6F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145576" class="Insecta" family="Formicidae" genus="Cryptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Cryptocerus complanatus Guerin-Meneville" order="Hymenoptera" pageId="191" pageNumber="191" phylum="Arthropoda" rank="species" species="complanatus">Cryptocerus complanatus</taxonomicName>
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<taxonomicName id="A09B49711BFDBD2CAC20A314444BD67C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145576" class="Insecta" family="Formicidae" genus="Cryptocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Cryptocerus complanatus Guerin-Meneville" order="Hymenoptera" pageId="191" pageNumber="191" phylum="Arthropoda" rank="species" species="complanatus">Cryptocerus complanatus, Guer.</taxonomicName>
Icon. Reg. Anim. 424 [[worker]].
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<paragraph id="C6142584E80CEAABDA4CAE1020EAB375" pageId="191" pageNumber="191">Smith, Mon. Crypt. Trans. Ent. Soc. Lond. 2nd ser. ii.221.20.</paragraph>
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<paragraph id="3A94FF0DE4A33923CF26C61067438342" pageId="191" pageNumber="191">Hab. Cayenne.</paragraph>
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</mods:role>
<mods:namePart id="D68E1CA4B4177129B8B4FDD4D5E595BD">Lin, Pengcheng</mods:namePart>
</mods:name>
<mods:name id="450845774A2757461962BA63EF4A1C1B" type="personal">
<mods:role id="DD34DF8A41FEFC7D473C92C9621C2632">
<mods:roleTerm id="A703CC3C2F064A7A6B9D91F2D5A7DF28">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="544C3EA0DDC054FC809B1591408A54B5">Jiang, Yueliang</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="64E8D986928950E9A2CFEEFEAD908C77">Liu, Huanzhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="B1AF58A1D0C25B802D67F34533C828A8">text</mods:typeOfResource>
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<mods:title id="BE9A572BE94C721C316F59D5B3A95758">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="81D85C82817BC039BF60D2DEDB42D097">
<mods:date id="A91DD70AC11D610640BDD6D288D269D1">2018</mods:date>
<mods:detail id="3DCB268CF25C283829859616D21D8176" type="pubDate">
<mods:number id="6E98716CE6587A219EF93A53BAD1B70B">2018-12-19</mods:number>
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<mods:number id="1124CEE2221DEAC6DD1F911D9100AEB4">4532</mods:number>
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<mods:identifier id="623A2FC210CD15D2E27DBA48880C2F6D" type="DOI">10.11646/zootaxa.4532.3.3</mods:identifier>
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<treatment id="2D6187EBFF963F39FF2F2183BAD5FE72" ID-DOI="http://doi.org/10.5281/zenodo.5967919" ID-GBIF-Taxon="154521377" ID-Zenodo-Dep="5967919" LSID="urn:lsid:plazi:treatment:2D6187EBFF963F39FF2F2183BAD5FE72" httpUri="http://treatment.plazi.org/id/2D6187EBFF963F39FF2F2183BAD5FE72" lastPageId="8" lastPageNumber="375" pageId="4" pageNumber="371">
<subSubSection id="EDD26576FF963F35FF2F2183BB3FFA29" box="[151,527,1427,1454]" pageId="4" pageNumber="371" type="nomenclature">
<paragraph id="A57736FDFF963F35FF2F2183BB3FFA29" blockId="4.[151,527,1427,1487]" box="[151,527,1427,1454]" pageId="4" pageNumber="371">
<heading id="FE3F8191FF963F35FF2F2183BB3FFA29" bold="true" box="[151,527,1427,1454]" fontSize="11" level="1" pageId="4" pageNumber="371" reason="1">
<emphasis id="97BCEAEFFF963F35FF2F2183BB3FFA29" bold="true" box="[151,527,1427,1454]" pageId="4" pageNumber="371">
<taxonomicName id="62C84D7EFF963F35FF2F2183B840FA2A" authority="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu, 2018" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[151,368,1427,1453]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="97BCEAEFFF963F35FF2F2183B840FA2A" bold="true" box="[151,368,1427,1453]" italics="true" pageId="4" pageNumber="371">Garra motuoensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="8C8F5794FF963F35FEC72184BB3FFA29" box="[383,527,1428,1454]" pageId="4" pageNumber="371" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF963F35FF2F21A6B822FA48" box="[151,274,1462,1487]" pageId="4" pageNumber="371" type="description">
<paragraph id="A57736FDFF963F35FF2F21A6B822FA48" blockId="4.[151,527,1427,1487]" box="[151,274,1462,1487]" pageId="4" pageNumber="371">
(
<figureCitation id="3DF32A78FF963F35FF2721A6B838FA48" box="[159,264,1462,1487]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="6.[151,250,1052,1074]" captionStartId-1="6.[151,250,1614,1636]" captionTargetBox-0="[151,1436,195,1028]" captionTargetBox-1="[151,1436,1155,1578]" captionTargetId-0="figure@6.[151,1436,195,1028]" captionTargetId-1="figure@6.[151,1436,1155,1578]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionText-0="FIGURE 1. Garra motuoensis, holotype, IHB 20161470, 162.5 mm SL; China: Tibet: Xigong River, a tributary of the Yarlung Tsangpo River." captionText-1="FIGURE 2. Garra motuoensis, holotype, IHB 20161470, 162.5 mm SL; China: Tibet: Xigong River, a tributary of the Yarlung Tsangpo River; dorsal and ventral views of head." httpUri-0="https://zenodo.org/record/2615431/files/figure.png" httpUri-1="https://zenodo.org/record/2615433/files/figure.png" pageId="4" pageNumber="371">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF963F35FF2F21EEBA71F94C" pageId="4" pageNumber="371" type="materials_examined">
<paragraph id="A57736FDFF963F35FF2F21EEBB47F9D8" blockId="4.[151,1437,1534,2027]" pageId="4" pageNumber="371">
<materialsCitation id="15A03CA0FF963F35FF2F21EEBB47F9D8" ID-GBIF-Occurrence="2119291559" collectingDate="2016-10" collectionCode="IHB" collectorName="Zheng Gong &amp; Jian Wang &amp; Huaming Hu" country="China" county="Motuo County" elevation="680" latitude="29.278889" location="Yarlung Tsangpo River" longLatPrecision="20" longitude="95.248886" municipality="Xigong River" pageId="4" pageNumber="371" specimenCode="IHB 20161470, 162.5" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="97BCEAEFFF963F35FF2F21EEB83BF990" bold="true" box="[151,267,1534,1559]" pageId="4" pageNumber="371">
<typeStatus id="7A73885FFF963F35FF2F21EEB836F990" box="[151,262,1534,1559]" pageId="4" pageNumber="371" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="F56E9E86FF963F35FEAE21EFB8F5F990" box="[278,453,1534,1559]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="4" pageNumber="371">
IHB
<quantity id="62309B18FF963F35FEEA21EEB8F5F990" box="[338,453,1534,1559]" metricMagnitude="7" metricUnit="m" metricValue="2.01614701625" pageId="4" pageNumber="371" unit="mm" value="2.01614701625E10">20161470</quantity>
</specimenCode>
,
<specimenCode id="F56E9E86FF963F35FE6C21EEBB23F990" box="[468,531,1534,1559]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="4" pageNumber="371">162.5</specimenCode>
mm SL;
<collectingCountry id="DDDF766DFF963F35FD3121EEBBE0F990" box="[649,720,1534,1559]" name="China" pageId="4" pageNumber="371">China</collectingCountry>
:
<collectingRegion id="670CF81FFF963F35FD6721EEBD26F990" box="[735,1046,1534,1559]" country="China" name="Xizang" pageId="4" pageNumber="371">Tibet Autonomous Region</collectingRegion>
:
<collectingCounty id="4C164E71FF963F35FB9D21EFBDE2F990" box="[1061,1234,1535,1559]" pageId="4" pageNumber="371">Motuo County</collectingCounty>
:
<collectingMunicipality id="4513AC87FF963F35FB5921EEBCB0F990" box="[1249,1408,1534,1559]" pageId="4" pageNumber="371">Xigong River</collectingMunicipality>
, a tributary of the
<location id="A0176026FF963F35FEFD2232BB7AF9BC" LSID="urn:lsid:plazi:treatment:2D6187EBFF963F39FF2F2183BAD5FE72:A0176026FF963F35FEFD2232BB7AF9BC" box="[325,586,1570,1595]" country="China" county="Motuo County" latitude="29.278889" longLatPrecision="20" longitude="95.248886" municipality="Xigong River" name="Yarlung Tsangpo River" pageId="4" pageNumber="371" stateProvince="Xizang">Yarlung Tsangpo River</location>
,
<geoCoordinate id="C0FC503AFF963F35FDEC2232BBEBF9BC" box="[596,731,1570,1595]" direction="north" orientation="latitude" pageId="4" pageNumber="371" precision="15" value="29.278889">29°16'44&quot;N</geoCoordinate>
,
<geoCoordinate id="C0FC503AFF963F35FD5E2232BA59F9BC" box="[742,873,1570,1595]" direction="east" orientation="longitude" pageId="4" pageNumber="371" precision="15" value="95.248886">95°14'56&quot;E</geoCoordinate>
,
<elevation id="2EE5D1CEFF963F35FCCC2232BD5FF9BC" box="[884,1135,1570,1595]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="371" unit="m" value="680.0">
<quantity id="62309B18FF963F35FCCC2232BA8BF9BD" box="[884,955,1570,1595]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="371" unit="m" value="680.0">680 m</quantity>
above sea level
</elevation>
; collected by
<collectorName id="083D532BFF963F35FAB12232BCA8F9BC" box="[1289,1432,1570,1595]" pageId="4" pageNumber="371">Zheng Gong</collectorName>
,
<collectorName id="083D532BFF963F35FF2F2256B83FF9D8" box="[151,271,1606,1631]" pageId="4" pageNumber="371">Jian Wang</collectorName>
, &amp;
<collectorName id="083D532BFF963F35FE8E2256B8FBF9D9" box="[310,459,1606,1631]" pageId="4" pageNumber="371">Huaming Hu</collectorName>
,
<date id="D176103DFF963F35FE6E2256BB43F9D8" box="[470,627,1606,1631]" pageId="4" pageNumber="371" value="2016-10">
<collectingDate id="C132E9D5FF963F35FE6E2256BB43F9D8" box="[470,627,1606,1631]" pageId="4" pageNumber="371" value="2016-10">October 2016</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="A57736FDFF963F35FF7F227ABA71F94C" blockId="4.[151,1437,1534,2027]" pageId="4" pageNumber="371">
<materialsCitation id="15A03CA0FF963F35FF7F227ABA80F904" ID-GBIF-Occurrence="2119289675" box="[199,944,1642,1667]" collectingDate="2016-04" collectionCode="IHB" pageId="4" pageNumber="371" specimenCode="IHB 20161471" specimenCount="1" typeStatus="paratype">
<emphasis id="97BCEAEFFF963F35FF7F227AB876F904" bold="true" box="[199,326,1642,1667]" pageId="4" pageNumber="371">
<typeStatus id="7A73885FFF963F35FF7F227AB871F904" box="[199,321,1642,1667]" pageId="4" pageNumber="371" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="F56E9E86FF963F35FEF4227BB8C6F904" box="[332,502,1642,1667]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="4" pageNumber="371">IHB 20161471</specimenCode>
,
<specimenCount id="B3CEFD74FF963F35FDB9227BBBB2F904" box="[513,642,1642,1667]" pageId="4" pageNumber="371" type="generic">1 specimen</specimenCount>
,
<quantity id="62309B18FF963F35FD35227BBBCFF905" box="[653,767,1643,1667]" metricMagnitude="-1" metricUnit="m" metricValue="1.208" pageId="4" pageNumber="371" unit="mm" value="120.8">120.8 mm</quantity>
SL,
<date id="D176103DFF963F35FC8B227ABA80F904" box="[819,944,1642,1667]" pageId="4" pageNumber="371" value="2016-04">
<collectingDate id="C132E9D5FF963F35FC8B227ABA80F904" box="[819,944,1642,1667]" pageId="4" pageNumber="371" value="2016-04">April 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="15A03CA0FF963F35FC04227BB83DF920" ID-GBIF-Occurrence="2119287133" collectingDate="2016-06" collectionCode="IHB" pageId="4" pageNumber="371" specimenCode="IHB 20161472" specimenCount="1" typeStatus="paratype">
<specimenCode id="F56E9E86FF963F35FC04227BBD56F904" box="[956,1126,1642,1667]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="4" pageNumber="371">IHB 20161472</specimenCode>
,
<specimenCount id="B3CEFD74FF963F35FBC8227BBDC2F904" box="[1136,1266,1642,1667]" pageId="4" pageNumber="371" type="generic">1 specimen</specimenCount>
,
<quantity id="62309B18FF963F35FB44227ABC5EF905" box="[1276,1390,1642,1667]" metricMagnitude="-1" metricUnit="m" metricValue="1.541" pageId="4" pageNumber="371" unit="mm" value="154.1">154.1 mm</quantity>
SL,
<date id="D176103DFF963F35FF2F229FB83DF920" box="[151,269,1678,1703]" pageId="4" pageNumber="371" value="2016-06">
<collectingDate id="C132E9D5FF963F35FF2F229FB83DF920" box="[151,269,1678,1703]" pageId="4" pageNumber="371" value="2016-06">June 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="15A03CA0FF963F35FEA1229FBA46F920" ID-GBIF-Occurrence="2119288962" box="[281,886,1678,1703]" collectingDate="2016-07" collectionCode="IHB" pageId="4" pageNumber="371" specimenCode="IHB 20161473" specimenCount="1" typeStatus="paratype">
<specimenCode id="F56E9E86FF963F35FEA1229FB8F3F920" box="[281,451,1678,1703]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="4" pageNumber="371">IHB 20161473</specimenCode>
,
<specimenCount id="B3CEFD74FF963F35FE77229FBB61F920" box="[463,593,1678,1703]" pageId="4" pageNumber="371" type="generic">1 specimen</specimenCount>
,
<quantity id="62309B18FF963F35FDE5229EBBFFF921" box="[605,719,1678,1703]" metricMagnitude="-1" metricUnit="m" metricValue="1.544" pageId="4" pageNumber="371" unit="mm" value="154.4">154.4 mm</quantity>
SL,
<date id="D176103DFF963F35FCBD229EBA46F920" box="[773,886,1678,1703]" pageId="4" pageNumber="371" value="2016-07">
<collectingDate id="C132E9D5FF963F35FCBD229EBA46F920" box="[773,886,1678,1703]" pageId="4" pageNumber="371" value="2016-07">July 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="15A03CA0FF963F35FC3A229FB8C6F94C" ID-GBIF-Occurrence="2119291890" collectingDate="2017-11" collectionCode="IHB" pageId="4" pageNumber="371" specimenCode="IHB 20171474-20171475" specimenCount="2" typeStatus="paratype">
<specimenCode id="F56E9E86FF963F35FC3A229FBD9BF920" box="[898,1195,1678,1703]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="4" pageNumber="371">IHB 20171474–20171475</specimenCode>
,
<specimenCount id="B3CEFD74FF963F35FB0F229FBC73F920" box="[1207,1347,1678,1703]" pageId="4" pageNumber="371" type="generic">2 specimens</specimenCount>
,
<quantity id="62309B18FF963F35FAF7229FB839F94D" metricMagnitude="-1" metricUnit="m" metricValue="1.3505" metricValueMax="1.403" metricValueMin="1.298" pageId="4" pageNumber="371" unit="mm" value="135.05" valueMax="140.3" valueMin="129.8">129.8– 140.3 mm</quantity>
SL,
<date id="D176103DFF963F35FE8622A2B8C6F94C" box="[318,502,1714,1739]" pageId="4" pageNumber="371" value="2017-11">
<collectingDate id="C132E9D5FF963F35FE8622A2B8C6F94C" box="[318,502,1714,1739]" pageId="4" pageNumber="371" value="2017-11">November 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="15A03CA0FF963F35FDBA22A2BA0DF94C" ID-GBIF-Occurrence="2119298357" box="[514,829,1714,1739]" collectingDate="2016-10" collectionCode="IHB" collectorName="Zheng Gong &amp; Jian Wang &amp; Huaming Hu" country="China" county="Motuo County" elevation="680" latitude="29.278889" location="Yarlung Tsangpo River" longLatPrecision="20" longitude="95.248886" municipality="Xigong River" pageId="4" pageNumber="371" specimenCount="1" stateProvince="Xizang" typeStatus="paratype">other data same as holotype</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF963F35FF7F22C6B818F86D" pageId="4" pageNumber="371" type="diagnosis">
<paragraph id="A57736FDFF963F35FF7F22C6B818F86D" blockId="4.[151,1437,1534,2027]" pageId="4" pageNumber="371">
<emphasis id="97BCEAEFFF963F35FF7F22C6B871F968" bold="true" box="[199,321,1750,1775]" pageId="4" pageNumber="371">Diagnosis.</emphasis>
<taxonomicName id="62C84D7EFF963F35FEF522C7BB2EF969" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[333,542,1751,1774]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="371" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF963F35FEF522C7BB2EF969" box="[333,542,1751,1774]" italics="true" pageId="4" pageNumber="371">Garra motuoensis</emphasis>
</taxonomicName>
, a member of the proboscis species group, is distinguished from the other members of this group by the following combination of characters: a prominent, quadrate, and slightly bilobed proboscis covered with 6–20 small to middle-sized unicuspid tubercles; some middle-sized unicuspid tubercles on the transverse lobe of the snout and several small unicuspid tubercles on the lateral surface of snout; anus slightly closer to anal-fin origin than to pelvic-fin origin (distance from anus to anal fin 36–46% of pelvic-anal distance); 36–37 lateral-line scales; 12 circumpeduncular scales; and other morphometric features: body depth 16.7–22.5% SL, caudal-peduncle length 16.8–20.3% SL, snout length 42–49% HL, eye diameter 15–18% HL, and disc length 42–51% HL.
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF973F39FF7F2487BC73FF70" lastPageId="8" lastPageNumber="375" pageId="5" pageNumber="372" type="description">
<paragraph id="A57736FDFF973F34FF7F2487B876FE2A" blockId="5.[151,1437,151,429]" pageId="5" pageNumber="372">
<emphasis id="97BCEAEFFF973F34FF7F2487B869FF37" bold="true" box="[199,345,151,176]" pageId="5" pageNumber="372">Description.</emphasis>
Morphometric data of
<specimenCount id="B3CEFD74FF973F34FDCF2487BBF3FF37" box="[631,707,151,176]" pageId="5" pageNumber="372" type="generic" typeStatus="type">6 type</specimenCount>
specimens in
<tableCitation id="E84A0346FF973F34FCCF2487BAE2FF37" box="[887,978,151,176]" captionStart="TABLE 3" captionStartId="5.[151,239,472,495]" captionText="TABLE 3. Morphometric data of holotype and five paratypes of Garra motuoensis. Ranges include values of holotype (n=6). SD, standard deviation." pageId="5" pageNumber="372">Table 3</tableCitation>
. Body elongate; cylindrical anteriorly, laterally compressed posteriorly. Head moderately large; head length greater than width, head width greater than depth; interorbital region slightly convex. Snout moderately rounded with transverse lobe covered with some middle-sized prominent unicuspid tubercles, demarcated posteriorly by deep transverse groove; several small unicuspid tubercles on lateral surface. Proboscis prominent, quadrate, and slightly bilobed, each lobe slightly elevated; anterodorsal region of proboscis covered with 6–20 irregularly arranged small to middle-sized unicuspid tubercles; tubercles more conspicuous in larger individuals. Eyes moderately large, situated dorsolaterally behind middle of head.
</paragraph>
<caption id="F1B76675FF973F34FF2F25C8B8EBFD88" pageId="5" pageNumber="372">
<paragraph id="A57736FDFF973F34FF2F25C8B8EBFD88" blockId="5.[151,1436,472,527]" pageId="5" pageNumber="372">
<emphasis id="97BCEAEFFF973F34FF2F25C8B83AFE68" bold="true" box="[151,266,472,495]" pageId="5" pageNumber="372">TABLE 3.</emphasis>
Morphometric data of holotype and five paratypes of
<taxonomicName id="62C84D7EFF973F34FCEF25C8BD2AFE68" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[855,1050,472,495]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="372" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF973F34FCEF25C8BD2AFE68" box="[855,1050,472,495]" italics="true" pageId="5" pageNumber="372">Garra motuoensis</emphasis>
</taxonomicName>
. Ranges include values of holotype (n=6). SD, standard deviation.
</paragraph>
</caption>
<paragraph id="A57736FDFF973F34FDD1263ABC3DF9F4" pageId="5" pageNumber="372">
<table id="D7C8C45DFF97C0CEFF25263ABC29F9F4" box="[157,1305,554,1651]" colsContinueIn="9.[159,1304,738,1835]" gridcols="5" gridrows="27" pageId="5" pageNumber="372">
<tr id="1BF834BFFF97C0CEFF25263ABC29FDC7" box="[157,1305,554,576]" gridrow="0" pageId="5" pageNumber="372" rowspan-0="1">
<th id="58295DC3FF97C0CEFDD1263ABBF7FDC7" box="[617,711,554,576]" gridcol="1" gridrow="0" pageId="5" pageNumber="372">Holotype</th>
<th id="58295DC3FF97C0CEFCF7263ABAFDFDC7" box="[847,973,554,576]" gridcol="2" gridrow="0" pageId="5" pageNumber="372">Range</th>
<th id="58295DC3FF97C0CEFB8D263ABD5FFDC7" box="[1077,1135,554,576]" gridcol="3" gridrow="0" pageId="5" pageNumber="372">Mean</th>
<th id="58295DC3FF97C0CEFB55263ABC29FDC7" box="[1261,1305,554,576]" gridcol="4" gridrow="0" pageId="5" pageNumber="372">SD</th>
</tr>
<tr id="1BF834BFFF97C0CEFF252644BC29FDED" box="[157,1305,596,618]" gridrow="1" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF252644B8FFFDED" box="[157,463,596,618]" gridcol="0" gridrow="1" pageId="5" pageNumber="372">Standard length (SL in mm)</th>
<td id="58295DC3FF97C0CEFDD12644BBF7FDED" box="[617,711,596,618]" gridcol="1" gridrow="1" pageId="5" pageNumber="372">162.5</td>
<td id="58295DC3FF97C0CEFCF72644BAFDFDED" box="[847,973,596,618]" gridcol="2" gridrow="1" pageId="5" pageNumber="372">118.9–162.5</td>
<td id="58295DC3FF97C0CEFB8D2644BD5FFDED" box="[1077,1135,596,618]" gridcol="3" gridrow="1" pageId="5" pageNumber="372">142.0</td>
<td id="58295DC3FF97C0CEFB552644BC29FDED" box="[1261,1305,596,618]" gridcol="4" gridrow="1" pageId="5" pageNumber="372">15.8</td>
</tr>
<tr id="1BF834BFFF97C0CEFF25266DBC29FD14" box="[157,1305,637,659]" gridrow="2" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25266DBC29FD14" box="[157,1305,637,659]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="5" pageNumber="372">In percent of SL</th>
</tr>
<tr id="1BF834BFFF97C0CEFF2526B6BC29FD3B" box="[157,1305,678,700]" gridrow="3" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2526B6B8FFFD3B" box="[157,463,678,700]" gridcol="0" gridrow="3" pageId="5" pageNumber="372">Body depth</th>
<td id="58295DC3FF97C0CEFDD126B6BBF7FD3B" box="[617,711,678,700]" gridcol="1" gridrow="3" pageId="5" pageNumber="372">17.6</td>
<td id="58295DC3FF97C0CEFCF726B6BAFDFD3B" box="[847,973,678,700]" gridcol="2" gridrow="3" pageId="5" pageNumber="372">16.7–22.5</td>
<td id="58295DC3FF97C0CEFB8D26B6BD5FFD3B" box="[1077,1135,678,700]" gridcol="3" gridrow="3" pageId="5" pageNumber="372">19.0</td>
<td id="58295DC3FF97C0CEFB5526B6BC29FD3B" box="[1261,1305,678,700]" gridcol="4" gridrow="3" pageId="5" pageNumber="372">2.5</td>
</tr>
<tr id="1BF834BFFF97C0CEFF2526C0BC29FD61" box="[157,1305,720,742]" gridrow="4" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2526C0B8FFFD61" box="[157,463,720,742]" gridcol="0" gridrow="4" pageId="5" pageNumber="372">Head length</th>
<td id="58295DC3FF97C0CEFDD126C0BBF7FD61" box="[617,711,720,742]" gridcol="1" gridrow="4" pageId="5" pageNumber="372">21.2</td>
<td id="58295DC3FF97C0CEFCF726C0BAFDFD61" box="[847,973,720,742]" gridcol="2" gridrow="4" pageId="5" pageNumber="372">21.2–23.8</td>
<td id="58295DC3FF97C0CEFB8D26C0BD5FFD61" box="[1077,1135,720,742]" gridcol="3" gridrow="4" pageId="5" pageNumber="372">22.4</td>
<td id="58295DC3FF97C0CEFB5526C0BC29FD61" box="[1261,1305,720,742]" gridcol="4" gridrow="4" pageId="5" pageNumber="372">0.9</td>
</tr>
<tr id="1BF834BFFF97C0CEFF2526E9BC29FC88" box="[157,1305,761,783]" gridrow="5" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2526E9B8FFFC88" box="[157,463,761,783]" gridcol="0" gridrow="5" pageId="5" pageNumber="372">Head height</th>
<td id="58295DC3FF97C0CEFDD126E9BBF7FC88" box="[617,711,761,783]" gridcol="1" gridrow="5" pageId="5" pageNumber="372">13.4</td>
<td id="58295DC3FF97C0CEFCF726E9BAFDFC88" box="[847,973,761,783]" gridcol="2" gridrow="5" pageId="5" pageNumber="372">13.4–15.1</td>
<td id="58295DC3FF97C0CEFB8D26E9BD5FFC88" box="[1077,1135,761,783]" gridcol="3" gridrow="5" pageId="5" pageNumber="372">14.1</td>
<td id="58295DC3FF97C0CEFB5526E9BC29FC88" box="[1261,1305,761,783]" gridcol="4" gridrow="5" pageId="5" pageNumber="372">0.7</td>
</tr>
<tr id="1BF834BFFF97C0CEFF252732BC29FCBF" box="[157,1305,802,824]" gridrow="6" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF252732B8FFFCBF" box="[157,463,802,824]" gridcol="0" gridrow="6" pageId="5" pageNumber="372">Head width</th>
<td id="58295DC3FF97C0CEFDD12732BBF7FCBF" box="[617,711,802,824]" gridcol="1" gridrow="6" pageId="5" pageNumber="372">17.3</td>
<td id="58295DC3FF97C0CEFCF72732BAFDFCBF" box="[847,973,802,824]" gridcol="2" gridrow="6" pageId="5" pageNumber="372">16.0–17.6</td>
<td id="58295DC3FF97C0CEFB8D2732BD5FFCBF" box="[1077,1135,802,824]" gridcol="3" gridrow="6" pageId="5" pageNumber="372">17.1</td>
<td id="58295DC3FF97C0CEFB552732BC29FCBF" box="[1261,1305,802,824]" gridcol="4" gridrow="6" pageId="5" pageNumber="372">0.6</td>
</tr>
<tr id="1BF834BFFF97C0CEFF25275CBC29FCE5" box="[157,1305,844,866]" gridrow="7" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25275CB8FFFCE5" box="[157,463,844,866]" gridcol="0" gridrow="7" pageId="5" pageNumber="372">Dorsal-fin length</th>
<td id="58295DC3FF97C0CEFDD1275CBBF7FCE5" box="[617,711,844,866]" gridcol="1" gridrow="7" pageId="5" pageNumber="372">23.3</td>
<td id="58295DC3FF97C0CEFCF7275CBAFDFCE5" box="[847,973,844,866]" gridcol="2" gridrow="7" pageId="5" pageNumber="372">22.6–24.4</td>
<td id="58295DC3FF97C0CEFB8D275CBD5FFCE5" box="[1077,1135,844,866]" gridcol="3" gridrow="7" pageId="5" pageNumber="372">23.4</td>
<td id="58295DC3FF97C0CEFB55275CBC29FCE5" box="[1261,1305,844,866]" gridcol="4" gridrow="7" pageId="5" pageNumber="372">0.6</td>
</tr>
<tr id="1BF834BFFF97C0CEFF252765BC29FC0C" box="[157,1305,885,907]" gridrow="8" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF252765B8FFFC0C" box="[157,463,885,907]" gridcol="0" gridrow="8" pageId="5" pageNumber="372">Pectoral-fin length</th>
<td id="58295DC3FF97C0CEFDD12765BBF7FC0C" box="[617,711,885,907]" gridcol="1" gridrow="8" pageId="5" pageNumber="372">20.0</td>
<td id="58295DC3FF97C0CEFCF72765BAFDFC0C" box="[847,973,885,907]" gridcol="2" gridrow="8" pageId="5" pageNumber="372">19.6–21.6</td>
<td id="58295DC3FF97C0CEFB8D2765BD5FFC0C" box="[1077,1135,885,907]" gridcol="3" gridrow="8" pageId="5" pageNumber="372">20.7</td>
<td id="58295DC3FF97C0CEFB552765BC29FC0C" box="[1261,1305,885,907]" gridcol="4" gridrow="8" pageId="5" pageNumber="372">0.9</td>
</tr>
<tr id="1BF834BFFF97C0CEFF25278EBC29FC33" box="[157,1305,926,948]" gridrow="9" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25278EB8FFFC33" box="[157,463,926,948]" gridcol="0" gridrow="9" pageId="5" pageNumber="372">Pelvic-fin length</th>
<td id="58295DC3FF97C0CEFDD1278EBBF7FC33" box="[617,711,926,948]" gridcol="1" gridrow="9" pageId="5" pageNumber="372">19.0</td>
<td id="58295DC3FF97C0CEFCF7278EBAFDFC33" box="[847,973,926,948]" gridcol="2" gridrow="9" pageId="5" pageNumber="372">19.0–21.0</td>
<td id="58295DC3FF97C0CEFB8D278EBD5FFC33" box="[1077,1135,926,948]" gridcol="3" gridrow="9" pageId="5" pageNumber="372">19.8</td>
<td id="58295DC3FF97C0CEFB55278EBC29FC33" box="[1261,1305,926,948]" gridcol="4" gridrow="9" pageId="5" pageNumber="372">0.8</td>
</tr>
<tr id="1BF834BFFF97C0CEFF2527D8BC29FC59" box="[157,1305,968,990]" gridrow="10" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2527D8B8FFFC59" box="[157,463,968,990]" gridcol="0" gridrow="10" pageId="5" pageNumber="372">Anal-fin length</th>
<td id="58295DC3FF97C0CEFDD127D8BBF7FC59" box="[617,711,968,990]" gridcol="1" gridrow="10" pageId="5" pageNumber="372">18.5</td>
<td id="58295DC3FF97C0CEFCF727D8BAFDFC59" box="[847,973,968,990]" gridcol="2" gridrow="10" pageId="5" pageNumber="372">18.5–20.2</td>
<td id="58295DC3FF97C0CEFB8D27D8BD5FFC59" box="[1077,1135,968,990]" gridcol="3" gridrow="10" pageId="5" pageNumber="372">19.3</td>
<td id="58295DC3FF97C0CEFB5527D8BC29FC59" box="[1261,1305,968,990]" gridcol="4" gridrow="10" pageId="5" pageNumber="372">0.6</td>
</tr>
<tr id="1BF834BFFF97C0CEFF2527E1BC29FB80" box="[157,1305,1009,1031]" gridrow="11" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2527E1B8FFFB80" box="[157,463,1009,1031]" gridcol="0" gridrow="11" pageId="5" pageNumber="372">Caudal-peduncle length</th>
<td id="58295DC3FF97C0CEFDD127E1BBF7FB80" box="[617,711,1009,1031]" gridcol="1" gridrow="11" pageId="5" pageNumber="372">16.8</td>
<td id="58295DC3FF97C0CEFCF727E1BAFDFB80" box="[847,973,1009,1031]" gridcol="2" gridrow="11" pageId="5" pageNumber="372">16.8–20.3</td>
<td id="58295DC3FF97C0CEFB8D27E1BD5FFB80" box="[1077,1135,1009,1031]" gridcol="3" gridrow="11" pageId="5" pageNumber="372">18.6</td>
<td id="58295DC3FF97C0CEFB5527E1BC29FB80" box="[1261,1305,1009,1031]" gridcol="4" gridrow="11" pageId="5" pageNumber="372">1.2</td>
</tr>
<tr id="1BF834BFFF97C0CEFF25200ABC29FBB7" box="[157,1305,1050,1072]" gridrow="12" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25200AB8FFFBB7" box="[157,463,1050,1072]" gridcol="0" gridrow="12" pageId="5" pageNumber="372">Caudal-peduncle depth</th>
<td id="58295DC3FF97C0CEFDD1200ABBF7FBB7" box="[617,711,1050,1072]" gridcol="1" gridrow="12" pageId="5" pageNumber="372">11.3</td>
<td id="58295DC3FF97C0CEFCF7200ABAFDFBB7" box="[847,973,1050,1072]" gridcol="2" gridrow="12" pageId="5" pageNumber="372">11.0–12.9</td>
<td id="58295DC3FF97C0CEFB8D200ABD5FFBB7" box="[1077,1135,1050,1072]" gridcol="3" gridrow="12" pageId="5" pageNumber="372">11.8</td>
<td id="58295DC3FF97C0CEFB55200ABC29FBB7" box="[1261,1305,1050,1072]" gridcol="4" gridrow="12" pageId="5" pageNumber="372">0.6</td>
</tr>
<tr id="1BF834BFFF97C0CEFF252054BC29FBDD" box="[157,1305,1092,1114]" gridrow="13" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF252054B8FFFBDD" box="[157,463,1092,1114]" gridcol="0" gridrow="13" pageId="5" pageNumber="372">Predorsal length</th>
<td id="58295DC3FF97C0CEFDD12054BBF7FBDD" box="[617,711,1092,1114]" gridcol="1" gridrow="13" pageId="5" pageNumber="372">43.4</td>
<td id="58295DC3FF97C0CEFCF72054BAFDFBDD" box="[847,973,1092,1114]" gridcol="2" gridrow="13" pageId="5" pageNumber="372">43.4–46.8</td>
<td id="58295DC3FF97C0CEFB8D2054BD5FFBDD" box="[1077,1135,1092,1114]" gridcol="3" gridrow="13" pageId="5" pageNumber="372">44.5</td>
<td id="58295DC3FF97C0CEFB552054BC29FBDD" box="[1261,1305,1092,1114]" gridcol="4" gridrow="13" pageId="5" pageNumber="372">1.2</td>
</tr>
<tr id="1BF834BFFF97C0CEFF25207DBC29FB04" box="[157,1305,1133,1155]" gridrow="14" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25207DB8FFFB04" box="[157,463,1133,1155]" gridcol="0" gridrow="14" pageId="5" pageNumber="372">Prepectoral length</th>
<td id="58295DC3FF97C0CEFDD1207DBBF7FB04" box="[617,711,1133,1155]" gridcol="1" gridrow="14" pageId="5" pageNumber="372">19.3</td>
<td id="58295DC3FF97C0CEFCF7207DBAFDFB04" box="[847,973,1133,1155]" gridcol="2" gridrow="14" pageId="5" pageNumber="372">18.7–20.2</td>
<td id="58295DC3FF97C0CEFB8D207DBD5FFB04" box="[1077,1135,1133,1155]" gridcol="3" gridrow="14" pageId="5" pageNumber="372">19.4</td>
<td id="58295DC3FF97C0CEFB55207DBC29FB04" box="[1261,1305,1133,1155]" gridcol="4" gridrow="14" pageId="5" pageNumber="372">0.6</td>
</tr>
<tr id="1BF834BFFF97C0CEFF252086BC29FB2B" box="[157,1305,1174,1196]" gridrow="15" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF252086B8FFFB2B" box="[157,463,1174,1196]" gridcol="0" gridrow="15" pageId="5" pageNumber="372">Prepelvic length</th>
<td id="58295DC3FF97C0CEFDD12086BBF7FB2B" box="[617,711,1174,1196]" gridcol="1" gridrow="15" pageId="5" pageNumber="372">48.6</td>
<td id="58295DC3FF97C0CEFCF72086BAFDFB2B" box="[847,973,1174,1196]" gridcol="2" gridrow="15" pageId="5" pageNumber="372">48.6–52.0</td>
<td id="58295DC3FF97C0CEFB8D2086BD5FFB2B" box="[1077,1135,1174,1196]" gridcol="3" gridrow="15" pageId="5" pageNumber="372">50.6</td>
<td id="58295DC3FF97C0CEFB552086BC29FB2B" box="[1261,1305,1174,1196]" gridcol="4" gridrow="15" pageId="5" pageNumber="372">1.1</td>
</tr>
<tr id="1BF834BFFF97C0CEFF2520D0BC29FB51" box="[157,1305,1216,1238]" gridrow="16" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2520D0B8FFFB51" box="[157,463,1216,1238]" gridcol="0" gridrow="16" pageId="5" pageNumber="372">Preanal length</th>
<td id="58295DC3FF97C0CEFDD120D0BBF7FB51" box="[617,711,1216,1238]" gridcol="1" gridrow="16" pageId="5" pageNumber="372">75.3</td>
<td id="58295DC3FF97C0CEFCF720D0BAFDFB51" box="[847,973,1216,1238]" gridcol="2" gridrow="16" pageId="5" pageNumber="372">72.7–75.4</td>
<td id="58295DC3FF97C0CEFB8D20D0BD5FFB51" box="[1077,1135,1216,1238]" gridcol="3" gridrow="16" pageId="5" pageNumber="372">74.1</td>
<td id="58295DC3FF97C0CEFB5520D0BC29FB51" box="[1261,1305,1216,1238]" gridcol="4" gridrow="16" pageId="5" pageNumber="372">1.1</td>
</tr>
<tr id="1BF834BFFF97C0CEFF2520F9BC29FB78" box="[157,1305,1257,1279]" gridrow="17" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2520F9BC29FB78" box="[157,1305,1257,1279]" colspan="5" colspanRight="4" gridcol="0" gridrow="17" pageId="5" pageNumber="372">In percent of HL</th>
</tr>
<tr id="1BF834BFFF97C0CEFF252102BC29FAAF" box="[157,1305,1298,1320]" gridrow="18" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF252102B8FFFAAF" box="[157,463,1298,1320]" gridcol="0" gridrow="18" pageId="5" pageNumber="372">Snout length</th>
<td id="58295DC3FF97C0CEFDD12102BBF7FAAF" box="[617,711,1298,1320]" gridcol="1" gridrow="18" pageId="5" pageNumber="372">48</td>
<td id="58295DC3FF97C0CEFCF72102BAFDFAAF" box="[847,973,1298,1320]" gridcol="2" gridrow="18" pageId="5" pageNumber="372">42–49</td>
<td id="58295DC3FF97C0CEFB8D2102BD5FFAAF" box="[1077,1135,1298,1320]" gridcol="3" gridrow="18" pageId="5" pageNumber="372">46.0</td>
<td id="58295DC3FF97C0CEFB552102BC29FAAF" box="[1261,1305,1298,1320]" gridcol="4" gridrow="18" pageId="5" pageNumber="372">2.9</td>
</tr>
<tr id="1BF834BFFF97C0CEFF25212CBC29FAD5" box="[157,1305,1340,1362]" gridrow="19" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25212CB8FFFAD5" box="[157,463,1340,1362]" gridcol="0" gridrow="19" pageId="5" pageNumber="372">Eye diameter</th>
<td id="58295DC3FF97C0CEFDD1212CBBF7FAD5" box="[617,711,1340,1362]" gridcol="1" gridrow="19" pageId="5" pageNumber="372">16</td>
<td id="58295DC3FF97C0CEFCF7212CBAFDFAD5" box="[847,973,1340,1362]" gridcol="2" gridrow="19" pageId="5" pageNumber="372">15–18</td>
<td id="58295DC3FF97C0CEFB8D212CBD5FFAD5" box="[1077,1135,1340,1362]" gridcol="3" gridrow="19" pageId="5" pageNumber="372">16.7</td>
<td id="58295DC3FF97C0CEFB55212CBC29FAD5" box="[1261,1305,1340,1362]" gridcol="4" gridrow="19" pageId="5" pageNumber="372">1.1</td>
</tr>
<tr id="1BF834BFFF97C0CEFF252175BC29FAFC" box="[157,1305,1381,1403]" gridrow="20" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF252175B8FFFAFC" box="[157,463,1381,1403]" gridcol="0" gridrow="20" pageId="5" pageNumber="372">Interorbital distance</th>
<td id="58295DC3FF97C0CEFDD12175BBF7FAFC" box="[617,711,1381,1403]" gridcol="1" gridrow="20" pageId="5" pageNumber="372">49</td>
<td id="58295DC3FF97C0CEFCF72175BAFDFAFC" box="[847,973,1381,1403]" gridcol="2" gridrow="20" pageId="5" pageNumber="372">47–49</td>
<td id="58295DC3FF97C0CEFB8D2175BD5FFAFC" box="[1077,1135,1381,1403]" gridcol="3" gridrow="20" pageId="5" pageNumber="372">47.9</td>
<td id="58295DC3FF97C0CEFB552175BC29FAFC" box="[1261,1305,1381,1403]" gridcol="4" gridrow="20" pageId="5" pageNumber="372">0.8</td>
</tr>
<tr id="1BF834BFFF97C0CEFF25219EBC29FA23" box="[157,1305,1422,1444]" gridrow="21" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25219EB8FFFA23" box="[157,463,1422,1444]" gridcol="0" gridrow="21" pageId="5" pageNumber="372">Disc length</th>
<td id="58295DC3FF97C0CEFDD1219EBBF7FA23" box="[617,711,1422,1444]" gridcol="1" gridrow="21" pageId="5" pageNumber="372">48</td>
<td id="58295DC3FF97C0CEFCF7219EBAFDFA23" box="[847,973,1422,1444]" gridcol="2" gridrow="21" pageId="5" pageNumber="372">42–51</td>
<td id="58295DC3FF97C0CEFB8D219EBD5FFA23" box="[1077,1135,1422,1444]" gridcol="3" gridrow="21" pageId="5" pageNumber="372">46.6</td>
<td id="58295DC3FF97C0CEFB55219EBC29FA23" box="[1261,1305,1422,1444]" gridcol="4" gridrow="21" pageId="5" pageNumber="372">3.7</td>
</tr>
<tr id="1BF834BFFF97C0CEFF2521A8BC29FA49" box="[157,1305,1464,1486]" gridrow="22" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2521A8B8FFFA49" box="[157,463,1464,1486]" gridcol="0" gridrow="22" pageId="5" pageNumber="372">Disc width</th>
<td id="58295DC3FF97C0CEFDD121A8BBF7FA49" box="[617,711,1464,1486]" gridcol="1" gridrow="22" pageId="5" pageNumber="372">69</td>
<td id="58295DC3FF97C0CEFCF721A8BAFDFA49" box="[847,973,1464,1486]" gridcol="2" gridrow="22" pageId="5" pageNumber="372">60–75</td>
<td id="58295DC3FF97C0CEFB8D21A8BD5FFA49" box="[1077,1135,1464,1486]" gridcol="3" gridrow="22" pageId="5" pageNumber="372">67.6</td>
<td id="58295DC3FF97C0CEFB5521A8BC29FA49" box="[1261,1305,1464,1486]" gridcol="4" gridrow="22" pageId="5" pageNumber="372">5.1</td>
</tr>
<tr id="1BF834BFFF97C0CEFF2521F1BC29FA70" box="[157,1305,1505,1527]" gridrow="23" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF2521F1B8FFFA70" box="[157,463,1505,1527]" gridcol="0" gridrow="23" pageId="5" pageNumber="372">Callous-pad length</th>
<td id="58295DC3FF97C0CEFDD121F1BBF7FA70" box="[617,711,1505,1527]" gridcol="1" gridrow="23" pageId="5" pageNumber="372">33</td>
<td id="58295DC3FF97C0CEFCF721F1BAFDFA70" box="[847,973,1505,1527]" gridcol="2" gridrow="23" pageId="5" pageNumber="372">28–34</td>
<td id="58295DC3FF97C0CEFB8D21F1BD5FFA70" box="[1077,1135,1505,1527]" gridcol="3" gridrow="23" pageId="5" pageNumber="372">32.2</td>
<td id="58295DC3FF97C0CEFB5521F1BC29FA70" box="[1261,1305,1505,1527]" gridcol="4" gridrow="23" pageId="5" pageNumber="372">2.3</td>
</tr>
<tr id="1BF834BFFF97C0CEFF25221ABC29F9A7" box="[157,1305,1546,1568]" gridrow="24" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25221AB8FFF9A7" box="[157,463,1546,1568]" gridcol="0" gridrow="24" pageId="5" pageNumber="372">Callous-pad width</th>
<td id="58295DC3FF97C0CEFDD1221ABBF7F9A7" box="[617,711,1546,1568]" gridcol="1" gridrow="24" pageId="5" pageNumber="372">48</td>
<td id="58295DC3FF97C0CEFCF7221ABAFDF9A7" box="[847,973,1546,1568]" gridcol="2" gridrow="24" pageId="5" pageNumber="372">42–48</td>
<td id="58295DC3FF97C0CEFB8D221ABD5FF9A7" box="[1077,1135,1546,1568]" gridcol="3" gridrow="24" pageId="5" pageNumber="372">44.7</td>
<td id="58295DC3FF97C0CEFB55221ABC29F9A7" box="[1261,1305,1546,1568]" gridcol="4" gridrow="24" pageId="5" pageNumber="372">2.5</td>
</tr>
<tr id="1BF834BFFF97C0CEFF252224BC29F9CD" box="[157,1305,1588,1610]" gridrow="25" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF252224BC29F9CD" box="[157,1305,1588,1610]" colspan="5" colspanRight="4" gridcol="0" gridrow="25" pageId="5" pageNumber="372">In percent of pelvic-anal distance</th>
</tr>
<tr id="1BF834BFFF97C0CEFF25224DBC29F9F4" box="[157,1305,1629,1651]" gridrow="26" pageId="5" pageNumber="372">
<th id="58295DC3FF97C0CEFF25224DB8FFF9F4" box="[157,463,1629,1651]" gridcol="0" gridrow="26" pageId="5" pageNumber="372">Distance from anus to anal fin</th>
<td id="58295DC3FF97C0CEFDD1224DBBF7F9F4" box="[617,711,1629,1651]" gridcol="1" gridrow="26" pageId="5" pageNumber="372">37</td>
<td id="58295DC3FF97C0CEFCF7224DBAFDF9F4" box="[847,973,1629,1651]" gridcol="2" gridrow="26" pageId="5" pageNumber="372">36–46</td>
<td id="58295DC3FF97C0CEFB8D224DBD5FF9F4" box="[1077,1135,1629,1651]" gridcol="3" gridrow="26" pageId="5" pageNumber="372">40.9</td>
<td id="58295DC3FF97C0CEFB55224DBC29F9F4" box="[1261,1305,1629,1651]" gridcol="4" gridrow="26" pageId="5" pageNumber="372">3.8</td>
</tr>
</table>
</paragraph>
<paragraph id="A57736FDFF973F34FF7F22A4BB6BF822" blockId="5.[151,1437,1716,1957]" pageId="5" pageNumber="372">Barbels in two pairs; rostral barbels situated antero-laterally, shorter than eye diameter; maxillary barbels shorter than rostral barbel. Rostral cap with crenulated margin and numerous tiny papillae, separated from upper jaw by deep groove, posteriorly continuous with lower lip on each side. Edge of upper and lower jaws covered with thin horny sheath. Mental adhesive disc large and elliptical; disc length shorter than width, slightly shorter than half of head length; anteromedian fold densely covered with tiny papillae, separated from lower jaw by deep groove and posteriorly bordered by shallow notch from central callous-pad; lateral and posterior margins free; lateroposterior flap with densely arranged tiny papillae.</paragraph>
<caption id="F1B76675FF943F37FF2F200CB801FBD6" httpUri="https://zenodo.org/record/2615431/files/figure.png" pageId="6" pageNumber="373" startId="6.[151,250,1052,1074]" targetBox="[151,1436,195,1028]" targetPageId="6">
<paragraph id="A57736FDFF943F37FF2F200CB801FBD6" blockId="6.[151,1435,1052,1105]" pageId="6" pageNumber="373">
<emphasis id="97BCEAEFFF943F37FF2F200CB822FBB5" bold="true" box="[151,274,1052,1074]" pageId="6" pageNumber="373">FIGURE 1.</emphasis>
<taxonomicName id="62C84D7EFF943F37FEA0200DB8E0FBB4" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[280,464,1053,1075]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF943F37FEA0200DB8E0FBB4" box="[280,464,1053,1075]" italics="true" pageId="6" pageNumber="373">Garra motuoensis</emphasis>
</taxonomicName>
, holotype, IHB 20161470, 162.5 mm SL; China: Tibet: Xigong River, a tributary of the Yarlung Tsangpo River.
</paragraph>
</caption>
<caption id="F1B76675FF943F37FF2F225EBBB8F904" httpUri="https://zenodo.org/record/2615433/files/figure.png" pageId="6" pageNumber="373" startId="6.[151,250,1614,1636]" targetBox="[151,1436,1155,1578]" targetPageId="6">
<paragraph id="A57736FDFF943F37FF2F225EBBB8F904" blockId="6.[151,1435,1614,1667]" pageId="6" pageNumber="373">
<emphasis id="97BCEAEFFF943F37FF2F225EB822F9E3" bold="true" box="[151,274,1614,1636]" pageId="6" pageNumber="373">FIGURE 2.</emphasis>
<taxonomicName id="62C84D7EFF943F37FEA0225FB8E0F9E2" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[280,464,1615,1637]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="373" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF943F37FEA0225FB8E0F9E2" box="[280,464,1615,1637]" italics="true" pageId="6" pageNumber="373">Garra motuoensis</emphasis>
</taxonomicName>
, holotype, IHB 20161470, 162.5 mm SL; China: Tibet: Xigong River, a tributary of the Yarlung Tsangpo River; dorsal and ventral views of head.
</paragraph>
</caption>
<paragraph id="A57736FDFF943F37FF7F22A0B847F826" blockId="6.[151,1437,1712,2025]" pageId="6" pageNumber="373">Dorsal fin with 3 simple and 8½ branched rays; distal margin slightly concave; first branched ray longest, nearly equal to head length; origin closer to snout tip than to caudal-fin base. Pectoral fin with 1 simple and 13 (1) or 14 (5*) branched rays; length shorter than head length; distal tip not reaching pelvic-fin origin and dorsal-fin origin. Pelvic fin with 1 simple and 8 branched rays; length nearly equal to head length; distal tip reaching beyond anus; origin closer to caudal-fin base than to snout tip. Anal fin with 3 simple and 5½ branched rays; distal tip not reaching to base of caudal fin; origin closer to caudal-fin base than to pelvic-fin origin. Caudal fin forked; lower lobe slightly longer.</paragraph>
<paragraph id="A57736FDFF943F36FF7F23BCBB92FE9B" blockId="6.[151,1437,1712,2025]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="374" pageId="6" pageNumber="373">
Lateral line complete and horizontal; with 34 (2*) or 35 (4) scales plus 2 scales on the caudal-fin base. Transverse scale rows above lateral line 3½, between lateral line and pelvic-fin origin 2½ (5*) or 3 (1). Circumpeduncular scales 12. Predorsal scales 10 (2*) or 11 (4); smaller than flank scales. Chest and belly scaled; scales on chest smaller than on belly. One long axillary scale at base of each pelvic fin. Anus slightly closer to analfin origin than to pelvic-fin origin (see
<figureCitation id="3DF32A78FF953F36FDDB24CFBB98FF7F" box="[611,680,223,248]" captionStart="FIGURE 1" captionStartId="6.[151,250,1052,1074]" captionTargetBox="[151,1436,195,1028]" captionTargetId="figure@6.[151,1436,195,1028]" captionTargetPageId="6" captionText="FIGURE 1. Garra motuoensis, holotype, IHB 20161470, 162.5 mm SL; China: Tibet: Xigong River, a tributary of the Yarlung Tsangpo River." httpUri="https://zenodo.org/record/2615431/files/figure.png" pageId="7" pageNumber="374">Fig. 1</figureCitation>
, distance from anus to anal fin 36–46% of pelvic-anal distance); separated from anal-fin origin by 5 scale rows.
</paragraph>
<caption id="F1B76675FF953F36FF2F2057BDB7FBDA" box="[151,1159,1095,1117]" httpUri="https://zenodo.org/record/2615435/files/figure.png" pageId="7" pageNumber="374" startId="7.[151,250,1095,1117]" targetBox="[219,1355,347,1071]" targetPageId="7">
<paragraph id="A57736FDFF953F36FF2F2057BDB7FBDA" blockId="7.[151,1159,1095,1117]" box="[151,1159,1095,1117]" pageId="7" pageNumber="374">
<emphasis id="97BCEAEFFF953F36FF2F2057B822FBDA" bold="true" box="[151,274,1095,1117]" pageId="7" pageNumber="374">FIGURE 3.</emphasis>
Map showing the sampling sites of
<taxonomicName id="62C84D7EFF953F36FD392058BA08FBDA" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[641,824,1096,1117]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="374" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF953F36FD392058BA08FBDA" box="[641,824,1096,1117]" italics="true" pageId="7" pageNumber="374">Garra motuoensis</emphasis>
</taxonomicName>
(▲) and
<taxonomicName id="62C84D7EFF953F36FC222058BD6DFBDA" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[922,1117,1096,1117]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="374" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF953F36FC222058BD6DFBDA" box="[922,1117,1096,1117]" italics="true" pageId="7" pageNumber="374">Garra yajiangensis</emphasis>
</taxonomicName>
(?).
</paragraph>
</caption>
<caption id="F1B76675FF953F36FF2F23C7BA23F86A" box="[151,787,2007,2029]" httpUri="https://zenodo.org/record/2615437/files/figure.png" pageId="7" pageNumber="374" startId="7.[151,250,2007,2029]" targetBox="[186,1399,1169,1978]" targetPageId="7">
<paragraph id="A57736FDFF953F36FF2F23C7BA23F86A" blockId="7.[151,787,2007,2029]" box="[151,787,2007,2029]" pageId="7" pageNumber="374">
<emphasis id="97BCEAEFFF953F36FF2F23C7B822F86A" bold="true" box="[151,274,2007,2029]" pageId="7" pageNumber="374">FIGURE 4.</emphasis>
Habitat of
<taxonomicName id="62C84D7EFF953F36FE3D23C8BB0DF86A" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[389,573,2008,2029]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="374" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF953F36FE3D23C8BB0DF86A" box="[389,573,2008,2029]" italics="true" pageId="7" pageNumber="374">Garra motuoensis</emphasis>
</taxonomicName>
in the Xigong River.
</paragraph>
</caption>
<paragraph id="A57736FDFF9A3F39FF7F2487BC73FF70" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="375">
<emphasis id="97BCEAEFFF9A3F39FF7F2487B87FFF37" bold="true" box="[199,335,151,176]" pageId="8" pageNumber="375">Coloration.</emphasis>
In preserved specimens, head, dorsum, and flank dark gray. Mouth, chest, and belly faint yellow or slightly brown. Dorsal, pectoral, pelvic, anal, and caudal fins gray; pectoral fin darker than other fins. Interradial membranes of all fins covered with conspicuous black pigment. Smaller individuals paler than larger ones.
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF9A3F39FF7F2514B802FEC7" pageId="8" pageNumber="375" type="distribution">
<paragraph id="A57736FDFF9A3F39FF7F2514B802FEC7" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="375">
<emphasis id="97BCEAEFFF9A3F39FF7F2514B850FE9A" bold="true" box="[199,352,260,285]" pageId="8" pageNumber="375">Distribution.</emphasis>
Known from the Xigong River, a tributary of the lower Yarlung Tsangpo River in southeastern
<collectingRegion id="670CF81FFF9A3F39FF2F2537B9E1FEC7" box="[151,209,295,320]" country="China" name="Xizang" pageId="8" pageNumber="375">Tibet</collectingRegion>
(
<figureCitation id="3DF32A78FF9A3F39FF582537B815FEC7" box="[224,293,295,320]" captionStart="FIGURE 3" captionStartId="7.[151,250,1095,1117]" captionTargetBox="[219,1355,347,1071]" captionTargetId="figure@7.[208,1379,337,1081]" captionTargetPageId="7" captionText="FIGURE 3. Map showing the sampling sites of Garra motuoensis (▲) and Garra yajiangensis (?)." httpUri="https://zenodo.org/record/2615435/files/figure.png" pageId="8" pageNumber="375">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF9A3F39FF7F255CBAE8FE57" pageId="8" pageNumber="375" type="biology_ecology">
<paragraph id="A57736FDFF9A3F39FF7F255CBAE8FE57" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="375">
<emphasis id="97BCEAEFFF9A3F39FF7F255CB81BFEE2" bold="true" box="[199,299,332,357]" pageId="8" pageNumber="375">Ecology.</emphasis>
This new species was caught from swift-flowing waters with numerous rocks in the Xigong River, where water is very clear (
<figureCitation id="3DF32A78FF9A3F39FE07257FBB35FE0F" box="[447,517,367,392]" captionStart="FIGURE 4" captionStartId="7.[151,250,2007,2029]" captionTargetBox="[186,1399,1169,1978]" captionTargetId="figure@7.[186,1399,1169,1978]" captionTargetPageId="7" captionText="FIGURE 4. Habitat of Garra motuoensis in the Xigong River." httpUri="https://zenodo.org/record/2615437/files/figure.png" pageId="8" pageNumber="375">Fig. 4</figureCitation>
). Macroscopic visual examination of the gonads suggests that its spawning period occurs in May when water temperature reaches 14°C. The eggs are yellow and demersal. Examinations of intestinal contents show that this species mainly feeds on periphytic algae.
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF9A3F39FF7F25CCBAD5FE72" box="[199,997,476,501]" pageId="8" pageNumber="375" type="etymology">
<paragraph id="A57736FDFF9A3F39FF7F25CCBAD5FE72" blockId="8.[151,1436,151,501]" box="[199,997,476,501]" pageId="8" pageNumber="375">
<emphasis id="97BCEAEFFF9A3F39FF7F25CCB87EFE72" bold="true" box="[199,334,476,501]" pageId="8" pageNumber="375">Etymology.</emphasis>
Named after its
<typeStatus id="7A73885FFF9A3F39FDB025CDBB08FE72" box="[520,568,477,501]" pageId="8" pageNumber="375">type</typeStatus>
locality, Motuo County. An adjective.
</paragraph>
</subSubSection>
</treatment>
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<mods:namePart id="8FCA5052EF23A9675112AA8E4C131624">Lin, Pengcheng</mods:namePart>
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<mods:name id="F1E62A8E07149C83586074AE7F0F5D09" type="personal">
<mods:role id="7C147E4BBFEB2FB47D13AAFD42B2F0A3">
<mods:roleTerm id="6097EC3A6B044D0453BD91CF70CF61F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9CE70F63AF8B22A43C03DD89FD015BBB">Jiang, Yueliang</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="BE85FCECA9A4ABFD68AE55921A8EF776">Liu, Huanzhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="DEBD5AFC11CBFD47737AA26241880E26">text</mods:typeOfResource>
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<mods:title id="9187830347729161C1BDE3DD9503F1BE">Zootaxa</mods:title>
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<mods:part id="D171B511773D024578D939B3A24DEFD7">
<mods:date id="ACD9B35E245EC6E99CE1E2E6B702C053">2018</mods:date>
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<mods:number id="7D0D47A7B54A766E881184B8832E9321">2018-12-19</mods:number>
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<mods:number id="83C751F5058F8071B00438A227FA0A89">4532</mods:number>
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<mods:classification id="9440918A9355448269F650F465774514">journal article</mods:classification>
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<mods:identifier id="7B64E50B22C7A507D2FAEF5190CCF73B" type="DOI">10.11646/zootaxa.4532.3.3</mods:identifier>
<mods:identifier id="912F9DD0BB074316FCF9AC3AB13DB2A7" type="GBIF-Dataset">9ac702b0-ddd3-4f52-a0b9-386ffbf2e4d5</mods:identifier>
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<treatment id="2D6187EBFF9A3F3DFF2F2657BDE5FF7F" ID-DOI="http://doi.org/10.5281/zenodo.5967922" ID-GBIF-Taxon="154521376" ID-Zenodo-Dep="5967922" LSID="urn:lsid:plazi:treatment:2D6187EBFF9A3F3DFF2F2657BDE5FF7F" httpUri="http://treatment.plazi.org/id/2D6187EBFF9A3F3DFF2F2657BDE5FF7F" lastPageId="12" lastPageNumber="378" pageId="8" pageNumber="375">
<subSubSection id="EDD26576FF9A3F39FF2F2657BB2BFDE5" box="[151,539,583,610]" pageId="8" pageNumber="375" type="nomenclature">
<paragraph id="A57736FDFF9A3F39FF2F2657BB2BFDE5" blockId="8.[151,539,583,643]" box="[151,539,583,610]" pageId="8" pageNumber="375">
<heading id="FE3F8191FF9A3F39FF2F2657BB2BFDE5" bold="true" box="[151,539,583,610]" fontSize="11" level="1" pageId="8" pageNumber="375" reason="1">
<emphasis id="97BCEAEFFF9A3F39FF2F2657BB2BFDE5" bold="true" box="[151,539,583,610]" pageId="8" pageNumber="375">
<taxonomicName id="62C84D7EFF9A3F39FF2F2657B84CFDE6" authority="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu, 2018" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[151,380,583,609]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="375" phylum="Chordata" rank="species" species="yajiangensis" status="sp. nov.">
<emphasis id="97BCEAEFFF9A3F39FF2F2657B84CFDE6" bold="true" box="[151,380,583,609]" italics="true" pageId="8" pageNumber="375">Garra yajiangensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="8C8F5794FF9A3F39FE342658BB2BFDE5" box="[396,539,584,610]" pageId="8" pageNumber="375" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF9A3F39FF2F267AB821FD04" box="[151,273,618,643]" pageId="8" pageNumber="375" type="description">
<paragraph id="A57736FDFF9A3F39FF2F267AB821FD04" blockId="8.[151,539,583,643]" box="[151,273,618,643]" pageId="8" pageNumber="375">
(
<figureCitation id="3DF32A78FF9A3F39FF27267AB839FD04" box="[159,265,618,643]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="8.[151,250,1899,1921]" captionStartId-1="10.[151,250,1018,1040]" captionTargetBox-0="[151,1436,1164,1878]" captionTargetBox-1="[150,1436,517,997]" captionTargetId-0="figure@8.[151,1436,1164,1878]" captionTargetId-1="figure@10.[150,1436,517,997]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionText-0="FIGURE 5. Garra yajiangensis, holotype, IHB 20161577, 165.0 mm SL; China: Tibet: lower Yarlung Tsangpo River." captionText-1="FIGURE 6. Garra yajiangensis, holotype, IHB 20161577, 165.0 mm SL; China: Tibet: lower Yarlung Tsangpo River; dorsal and ventral views of head." httpUri-0="https://zenodo.org/record/2615439/files/figure.png" httpUri-1="https://zenodo.org/record/2615443/files/figure.png" pageId="8" pageNumber="375">Figs. 5–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF9A3F39FF2F26A2BC29FCB0" pageId="8" pageNumber="375" type="materials_examined">
<paragraph id="A57736FDFF9A3F39FF2F26A2B858FC94" blockId="8.[151,1436,690,1111]" pageId="8" pageNumber="375">
<materialsCitation id="15A03CA0FF9A3F39FF2F26A2B858FC94" ID-GBIF-Occurrence="2119295057" collectingDate="2016-10" collectionCode="IHB" collectorName="Zheng Gong &amp; Jian Wang &amp; Huaming Hu" country="China" county="Motuo County" elevation="708" latitude="29.322779" location="lower Yarlung Tsangpo River" longLatPrecision="20" longitude="95.30361" pageId="8" pageNumber="375" specimenCode="IHB 20161577" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="97BCEAEFFF9A3F39FF2F26A2B83BFD4C" bold="true" box="[151,267,690,715]" pageId="8" pageNumber="375">
<typeStatus id="7A73885FFF9A3F39FF2F26A2B836FD4C" box="[151,262,690,715]" pageId="8" pageNumber="375" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="F56E9E86FF9A3F39FEA026A3B8F8FD4C" box="[280,456,690,715]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="8" pageNumber="375">IHB 20161577</specimenCode>
, 165.0 mm SL;
<collectingCountry id="DDDF766DFF9A3F39FD2C26A2BBECFD4C" box="[660,732,690,715]" name="China" pageId="8" pageNumber="375">China</collectingCountry>
:
<collectingRegion id="670CF81FFF9A3F39FD5426A2BD16FD4C" box="[748,1062,690,715]" country="China" name="Xizang" pageId="8" pageNumber="375">Tibet Autonomous Region</collectingRegion>
:
<collectingCounty id="4C164E71FF9A3F39FB8F26A3BDD6FD4C" box="[1079,1254,691,715]" pageId="8" pageNumber="375">Motuo County</collectingCounty>
:
<location id="A0176026FF9A3F39FB4E26A2B80CFD68" LSID="urn:lsid:plazi:treatment:2D6187EBFF9A3F3DFF2F2657BDE5FF7F:A0176026FF9A3F39FB4E26A2B80CFD68" country="China" county="Motuo County" latitude="29.322779" longLatPrecision="20" longitude="95.30361" name="lower Yarlung Tsangpo River" pageId="8" pageNumber="375" stateProvince="Xizang">lower Yarlung Tsangpo River</location>
,
<geoCoordinate id="C0FC503AFF9A3F39FEFF26C7B8FDFD68" box="[327,461,727,751]" direction="north" orientation="latitude" pageId="8" pageNumber="375" precision="15" value="29.322779">29°19'22&quot;N</geoCoordinate>
,
<geoCoordinate id="C0FC503AFF9A3F39FE6026C6BB6BFD68" box="[472,603,726,751]" direction="east" orientation="longitude" pageId="8" pageNumber="375" precision="15" value="95.30361">95°18'13&quot;E</geoCoordinate>
,
<elevation id="2EE5D1CEFF9A3F39FDDF26C7BA52FD68" box="[615,866,726,751]" metricMagnitude="2" metricUnit="m" metricValue="7.08" pageId="8" pageNumber="375" unit="m" value="708.0">
<quantity id="62309B18FF9A3F39FDDF26C7BB9DFD69" box="[615,685,727,751]" metricMagnitude="2" metricUnit="m" metricValue="7.08" pageId="8" pageNumber="375" unit="m" value="708.0">708 m</quantity>
above sea level
</elevation>
; collected by
<collectorName id="083D532BFF9A3F39FC4326C6BDBBFD68" box="[1019,1163,726,751]" pageId="8" pageNumber="375">Zheng Gong</collectorName>
,
<collectorName id="083D532BFF9A3F39FB2D26C6BC3DFD68" box="[1173,1293,726,751]" pageId="8" pageNumber="375">Jian Wang</collectorName>
, &amp;
<collectorName id="083D532BFF9A3F39FA8C26C6B98CFC95" pageId="8" pageNumber="375">Huaming Hu</collectorName>
,
<date id="D176103DFF9A3F39FF7F26EAB854FC94" box="[199,356,762,787]" pageId="8" pageNumber="375" value="2016-10">
<collectingDate id="C132E9D5FF9A3F39FF7F26EAB854FC94" box="[199,356,762,787]" pageId="8" pageNumber="375" value="2016-10">October 2016</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="A57736FDFF9A3F39FF7F270EBC29FCB0" blockId="8.[151,1436,690,1111]" box="[199,1305,798,823]" pageId="8" pageNumber="375">
<materialsCitation id="15A03CA0FF9A3F39FF7F270EBC25FCB0" ID-GBIF-Occurrence="2119294077" box="[199,1301,798,823]" collectingDate="2016-10" collectionCode="IHB" collectorName="Zheng Gong &amp; Jian Wang &amp; Huaming Hu" country="China" county="Motuo County" elevation="708" latitude="29.322779" location="lower Yarlung Tsangpo River" longLatPrecision="20" longitude="95.30361" pageId="8" pageNumber="375" specimenCode="IHB 20161578-20161592" specimenCount="15" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="97BCEAEFFF9A3F39FF7F270EB876FCB0" bold="true" box="[199,326,798,823]" pageId="8" pageNumber="375">
<typeStatus id="7A73885FFF9A3F39FF7F270EB871FCB0" box="[199,321,798,823]" pageId="8" pageNumber="375" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="F56E9E86FF9A3F39FEF4270FBB45FCB0" box="[332,629,798,823]" collectionCode="IHB" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35160" lsid="urn:lsid:biocol.org:col:35160" name="Institute of Hydrobiology, Chinese Academy of Sciences" pageId="8" pageNumber="375">IHB 20161578–20161592</specimenCode>
,
<specimenCount id="B3CEFD74FF9A3F39FD38270EBA2BFCB0" box="[640,795,798,823]" pageId="8" pageNumber="375" type="generic">15 specimens</specimenCount>
,
<quantity id="62309B18FF9A3F39FC9E270EBAD4FCB1" box="[806,996,798,823]" metricMagnitude="-1" metricUnit="m" metricValue="1.5044999999999997" metricValueMax="1.751" metricValueMin="1.258" pageId="8" pageNumber="375" unit="mm" value="150.45" valueMax="175.1" valueMin="125.8">125.8–175.1 mm</quantity>
SL; same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF9A3F39FF7F2752BB3CFBD1" pageId="8" pageNumber="375" type="diagnosis">
<paragraph id="A57736FDFF9A3F39FF7F2752BB3CFBD1" blockId="8.[151,1436,690,1111]" pageId="8" pageNumber="375">
<emphasis id="97BCEAEFFF9A3F39FF7F2752B871FCDC" bold="true" box="[199,321,834,859]" pageId="8" pageNumber="375">Diagnosis.</emphasis>
<taxonomicName id="62C84D7EFF9A3F39FEF42753BB1AFCDD" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[332,554,835,858]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="375" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF9A3F39FEF42753BB1AFCDD" box="[332,554,835,858]" italics="true" pageId="8" pageNumber="375">Garra yajiangensis</emphasis>
</taxonomicName>
, a member of the proboscis species group, is distinguished from the other members of this group by the following combination of characters: a quadrate and slightly bilobed proboscis covered with 2–7 middle-sized unicuspid tubercles, including two large-sized tubercles on the anterior margin of each lobe; some small unicuspid tubercles on the transverse lobe and the lateral surface of snout; anus situated substantially closer to the anal-fin origin than to the pelvic-fin origin (distance from anus to anal fin 19–24% of pelvic-anal distance); 34–36 lateral-line scales; 12 circumpeduncular scales; and morphometric features: body depth 17.4–19.5% SL, caudal-peduncle length 15.7–19.2% SL, snout length 43–49% HL, eye diameter 13–18% HL, and disc length 49–58% HL.
</paragraph>
</subSubSection>
<caption id="F1B76675FF9A3F39FF2F237BBC77F806" box="[151,1351,1899,1921]" httpUri="https://zenodo.org/record/2615439/files/figure.png" pageId="8" pageNumber="375" startId="8.[151,250,1899,1921]" targetBox="[151,1436,1164,1878]" targetPageId="8">
<paragraph id="A57736FDFF9A3F39FF2F237BBC77F806" blockId="8.[151,1351,1899,1921]" box="[151,1351,1899,1921]" pageId="8" pageNumber="375">
<emphasis id="97BCEAEFFF9A3F39FF2F237BB822F806" bold="true" box="[151,274,1899,1921]" pageId="8" pageNumber="375">FIGURE 5.</emphasis>
<taxonomicName id="62C84D7EFF9A3F39FEA1237CB8ECF806" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[281,476,1900,1921]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="375" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF9A3F39FEA1237CB8ECF806" box="[281,476,1900,1921]" italics="true" pageId="8" pageNumber="375">Garra yajiangensis</emphasis>
</taxonomicName>
, holotype, IHB 20161577, 165.0 mm SL; China: Tibet: lower Yarlung Tsangpo River.
</paragraph>
</caption>
<subSubSection id="EDD26576FF9A3F3AFF7F23BDB84DFE9A" lastPageId="11" lastPageNumber="378" pageId="8" pageNumber="375" type="description">
<paragraph id="A57736FDFF9A3F38FF7F23BDBB0BFEE2" blockId="8.[151,1436,1965,2026]" lastBlockId="9.[151,1437,151,608]" lastPageId="9" lastPageNumber="376" pageId="8" pageNumber="375">
<emphasis id="97BCEAEFFF9A3F39FF7F23BDB869F841" bold="true" box="[199,345,1965,1990]" pageId="8" pageNumber="375">Description.</emphasis>
Morphometric data of
<specimenCount id="B3CEFD74FF9A3F39FDCB23BDBBFBF840" box="[627,715,1965,1991]" pageId="8" pageNumber="375" type="generic" typeStatus="type">16 type</specimenCount>
specimens in
<tableCitation id="E84A0346FF9A3F39FCC423BDBAE8F841" box="[892,984,1965,1990]" captionStart="TABLE 4" captionStartId="9.[151,239,656,679]" captionText="TABLE 4. Morphometric data of holotype and 15 paratypes of Garra yajiangensis Ranges include values of holotype (n=16). SD, standard deviation." pageId="8" pageNumber="375">Table 4</tableCitation>
. Body elongate; cylindrical anteriorly, laterally compressed posteriorly. Head moderately large; head length greater than width, head width greater than depth; interorbital region slightly convex. Snout rounded with transverse lobe covered with some small unicuspid tubercles, demarcated posteriorly by deep transverse groove; some small unicuspid tubercles on lateral surface. Proboscis quadrate, moderately prominent, and slightly bilobed, each lobe slightly protruding; anterodorsal region of proboscis covered with 2–7 middle-sized unicuspid tubercles, including two large-sized tubercles on anterior margin of each lobe; tubercles more conspicuous in larger individuals. Eyes moderately large, situated dorsolaterally behind middle of head.
</paragraph>
<paragraph id="A57736FDFF9B3F38FF7F257FBB6BFDE7" blockId="9.[151,1437,151,608]" pageId="9" pageNumber="376">Barbels in two pairs; rostral barbels situated antero-laterally, shorter than eye diameter; maxillary barbels shorter than rostral barbels. Rostral cap with crenulated margin and numerous tiny papillae, separated from upper jaw by deep groove, posteriorly continuous with lower lip on each side. Edge of upper and lower jaws covered with thin horny sheath. Mental adhesive disc large, elliptical; disc length shorter than width, slightly longer than half head length; anteromedian fold densely covered by tiny papillae, separated from lower jaw by deep groove, posteriorly bordered by shallow notch from central callous-pad; lateral and posterior margins free; lateroposterior flap with densely arranged tiny papillae.</paragraph>
<caption id="F1B76675FF9B3F38FF2F2680B8D8FD40" pageId="9" pageNumber="376">
<paragraph id="A57736FDFF9B3F38FF2F2680B8D8FD40" blockId="9.[151,1436,656,711]" pageId="9" pageNumber="376">
<emphasis id="97BCEAEFFF9B3F38FF2F2680B83BFD20" bold="true" box="[151,267,656,679]" pageId="9" pageNumber="376">TABLE 4.</emphasis>
Morphometric data of holotype and 15 paratypes of
<taxonomicName id="62C84D7EFF9B3F38FCF52680BD44FD20" authority="Ranges" authorityName="Ranges" box="[845,1140,656,679]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="376" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF9B3F38FCF52680BD2EFD20" box="[845,1054,656,679]" italics="true" pageId="9" pageNumber="376">Garra yajiangensis</emphasis>
Ranges
</taxonomicName>
include values of holotype (n=16). SD, standard deviation.
</paragraph>
</caption>
<paragraph id="A57736FDFF9B3F38FDD226F2BC3CF8AC" pageId="9" pageNumber="376">
<table id="D7C8C45DFF9BC0CEFF2726F2BC28F8AC" box="[159,1304,738,1835]" colsContinueFrom="5.[157,1305,554,1651]" gridcols="5" gridrows="27" pageId="9" pageNumber="376">
<tr id="1BF834BFFF9BC0CEFF2726F2BC28FD7F" box="[159,1304,738,760]" gridrow="0" pageId="9" pageNumber="376" rowspan-0="1">
<th id="58295DC3FF9BC0CEFDD226F2BBF8FD7F" box="[618,712,738,760]" gridcol="1" gridrow="0" pageId="9" pageNumber="376">Holotype</th>
<th id="58295DC3FF9BC0CEFCF726F2BAFEFD7F" box="[847,974,738,760]" gridcol="2" gridrow="0" pageId="9" pageNumber="376">Range</th>
<th id="58295DC3FF9BC0CEFB8C26F2BD5EFD7F" box="[1076,1134,738,760]" gridcol="3" gridrow="0" pageId="9" pageNumber="376">Mean</th>
<th id="58295DC3FF9BC0CEFB5426F2BC28FD7F" box="[1260,1304,738,760]" gridcol="4" gridrow="0" pageId="9" pageNumber="376">SD</th>
</tr>
<tr id="1BF834BFFF9BC0CEFF27271CBC28FCA5" box="[159,1304,780,802]" gridrow="1" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF27271CB8E1FCA5" box="[159,465,780,802]" gridcol="0" gridrow="1" pageId="9" pageNumber="376">Standard length (SL in mm)</th>
<td id="58295DC3FF9BC0CEFDD2271CBBF8FCA5" box="[618,712,780,802]" gridcol="1" gridrow="1" pageId="9" pageNumber="376">165.0</td>
<td id="58295DC3FF9BC0CEFCF7271CBAFEFCA5" box="[847,974,780,802]" gridcol="2" gridrow="1" pageId="9" pageNumber="376">125.8–175.1</td>
<td id="58295DC3FF9BC0CEFB8C271CBD5EFCA5" box="[1076,1134,780,802]" gridcol="3" gridrow="1" pageId="9" pageNumber="376">151.4</td>
<td id="58295DC3FF9BC0CEFB54271CBC28FCA5" box="[1260,1304,780,802]" gridcol="4" gridrow="1" pageId="9" pageNumber="376">16.3</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272725BC28FCCC" box="[159,1304,821,843]" gridrow="2" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272725BC28FCCC" box="[159,1304,821,843]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="9" pageNumber="376">In percent of SL</th>
</tr>
<tr id="1BF834BFFF9BC0CEFF27274EBC28FCF3" box="[159,1304,862,884]" gridrow="3" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF27274EB8E1FCF3" box="[159,465,862,884]" gridcol="0" gridrow="3" pageId="9" pageNumber="376">Body depth</th>
<td id="58295DC3FF9BC0CEFDD2274EBBF8FCF3" box="[618,712,862,884]" gridcol="1" gridrow="3" pageId="9" pageNumber="376">18.6</td>
<td id="58295DC3FF9BC0CEFCF7274EBAFEFCF3" box="[847,974,862,884]" gridcol="2" gridrow="3" pageId="9" pageNumber="376">17.4–19.5</td>
<td id="58295DC3FF9BC0CEFB8C274EBD5EFCF3" box="[1076,1134,862,884]" gridcol="3" gridrow="3" pageId="9" pageNumber="376">18.4</td>
<td id="58295DC3FF9BC0CEFB54274EBC28FCF3" box="[1260,1304,862,884]" gridcol="4" gridrow="3" pageId="9" pageNumber="376">0.6</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272798BC28FC19" box="[159,1304,904,926]" gridrow="4" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272798B8E1FC19" box="[159,465,904,926]" gridcol="0" gridrow="4" pageId="9" pageNumber="376">Head length</th>
<td id="58295DC3FF9BC0CEFDD22798BBF8FC19" box="[618,712,904,926]" gridcol="1" gridrow="4" pageId="9" pageNumber="376">23.8</td>
<td id="58295DC3FF9BC0CEFCF72798BAFEFC19" box="[847,974,904,926]" gridcol="2" gridrow="4" pageId="9" pageNumber="376">22.2–25.4</td>
<td id="58295DC3FF9BC0CEFB8C2798BD5EFC19" box="[1076,1134,904,926]" gridcol="3" gridrow="4" pageId="9" pageNumber="376">23.6</td>
<td id="58295DC3FF9BC0CEFB542798BC28FC19" box="[1260,1304,904,926]" gridcol="4" gridrow="4" pageId="9" pageNumber="376">0.8</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2727A1BC28FC40" box="[159,1304,945,967]" gridrow="5" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2727A1B8E1FC40" box="[159,465,945,967]" gridcol="0" gridrow="5" pageId="9" pageNumber="376">Head height</th>
<td id="58295DC3FF9BC0CEFDD227A1BBF8FC40" box="[618,712,945,967]" gridcol="1" gridrow="5" pageId="9" pageNumber="376">14.0</td>
<td id="58295DC3FF9BC0CEFCF727A1BAFEFC40" box="[847,974,945,967]" gridcol="2" gridrow="5" pageId="9" pageNumber="376">13.9–15.5</td>
<td id="58295DC3FF9BC0CEFB8C27A1BD5EFC40" box="[1076,1134,945,967]" gridcol="3" gridrow="5" pageId="9" pageNumber="376">14.4</td>
<td id="58295DC3FF9BC0CEFB5427A1BC28FC40" box="[1260,1304,945,967]" gridcol="4" gridrow="5" pageId="9" pageNumber="376">0.4</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2727CABC28FC77" box="[159,1304,986,1008]" gridrow="6" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2727CAB8E1FC77" box="[159,465,986,1008]" gridcol="0" gridrow="6" pageId="9" pageNumber="376">Head width</th>
<td id="58295DC3FF9BC0CEFDD227CABBF8FC77" box="[618,712,986,1008]" gridcol="1" gridrow="6" pageId="9" pageNumber="376">16.9</td>
<td id="58295DC3FF9BC0CEFCF727CABAFEFC77" box="[847,974,986,1008]" gridcol="2" gridrow="6" pageId="9" pageNumber="376">15.8–22.4</td>
<td id="58295DC3FF9BC0CEFB8C27CABD5EFC77" box="[1076,1134,986,1008]" gridcol="3" gridrow="6" pageId="9" pageNumber="376">17.7</td>
<td id="58295DC3FF9BC0CEFB5427CABC28FC77" box="[1260,1304,986,1008]" gridcol="4" gridrow="6" pageId="9" pageNumber="376">1.5</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272014BC28FB9D" box="[159,1304,1028,1050]" gridrow="7" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272014B8E1FB9D" box="[159,465,1028,1050]" gridcol="0" gridrow="7" pageId="9" pageNumber="376">Dorsal-fin length</th>
<td id="58295DC3FF9BC0CEFDD22014BBF8FB9D" box="[618,712,1028,1050]" gridcol="1" gridrow="7" pageId="9" pageNumber="376">22.8</td>
<td id="58295DC3FF9BC0CEFCF72014BAFEFB9D" box="[847,974,1028,1050]" gridcol="2" gridrow="7" pageId="9" pageNumber="376">22.1–25.7</td>
<td id="58295DC3FF9BC0CEFB8C2014BD5EFB9D" box="[1076,1134,1028,1050]" gridcol="3" gridrow="7" pageId="9" pageNumber="376">23.6</td>
<td id="58295DC3FF9BC0CEFB542014BC28FB9D" box="[1260,1304,1028,1050]" gridcol="4" gridrow="7" pageId="9" pageNumber="376">0.8</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF27203DBC28FBC4" box="[159,1304,1069,1091]" gridrow="8" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF27203DB8E1FBC4" box="[159,465,1069,1091]" gridcol="0" gridrow="8" pageId="9" pageNumber="376">Pectoral-fin length</th>
<td id="58295DC3FF9BC0CEFDD2203DBBF8FBC4" box="[618,712,1069,1091]" gridcol="1" gridrow="8" pageId="9" pageNumber="376">21.7</td>
<td id="58295DC3FF9BC0CEFCF7203DBAFEFBC4" box="[847,974,1069,1091]" gridcol="2" gridrow="8" pageId="9" pageNumber="376">21.3–24.2</td>
<td id="58295DC3FF9BC0CEFB8C203DBD5EFBC4" box="[1076,1134,1069,1091]" gridcol="3" gridrow="8" pageId="9" pageNumber="376">22.8</td>
<td id="58295DC3FF9BC0CEFB54203DBC28FBC4" box="[1260,1304,1069,1091]" gridcol="4" gridrow="8" pageId="9" pageNumber="376">0.9</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272046BC28FBEB" box="[159,1304,1110,1132]" gridrow="9" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272046B8E1FBEB" box="[159,465,1110,1132]" gridcol="0" gridrow="9" pageId="9" pageNumber="376">Pelvic-fin length</th>
<td id="58295DC3FF9BC0CEFDD22046BBF8FBEB" box="[618,712,1110,1132]" gridcol="1" gridrow="9" pageId="9" pageNumber="376">21.8</td>
<td id="58295DC3FF9BC0CEFCF72046BAFEFBEB" box="[847,974,1110,1132]" gridcol="2" gridrow="9" pageId="9" pageNumber="376">20.0–23.9</td>
<td id="58295DC3FF9BC0CEFB8C2046BD5EFBEB" box="[1076,1134,1110,1132]" gridcol="3" gridrow="9" pageId="9" pageNumber="376">21.9</td>
<td id="58295DC3FF9BC0CEFB542046BC28FBEB" box="[1260,1304,1110,1132]" gridcol="4" gridrow="9" pageId="9" pageNumber="376">1.1</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272090BC28FB11" box="[159,1304,1152,1174]" gridrow="10" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272090B8E1FB11" box="[159,465,1152,1174]" gridcol="0" gridrow="10" pageId="9" pageNumber="376">Anal-fin length</th>
<td id="58295DC3FF9BC0CEFDD22090BBF8FB11" box="[618,712,1152,1174]" gridcol="1" gridrow="10" pageId="9" pageNumber="376">19.9</td>
<td id="58295DC3FF9BC0CEFCF72090BAFEFB11" box="[847,974,1152,1174]" gridcol="2" gridrow="10" pageId="9" pageNumber="376">18.6–21.4</td>
<td id="58295DC3FF9BC0CEFB8C2090BD5EFB11" box="[1076,1134,1152,1174]" gridcol="3" gridrow="10" pageId="9" pageNumber="376">20.0</td>
<td id="58295DC3FF9BC0CEFB542090BC28FB11" box="[1260,1304,1152,1174]" gridcol="4" gridrow="10" pageId="9" pageNumber="376">0.8</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2720B9BC28FB38" box="[159,1304,1193,1215]" gridrow="11" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2720B9B8E1FB38" box="[159,465,1193,1215]" gridcol="0" gridrow="11" pageId="9" pageNumber="376">Caudal-peduncle length</th>
<td id="58295DC3FF9BC0CEFDD220B9BBF8FB38" box="[618,712,1193,1215]" gridcol="1" gridrow="11" pageId="9" pageNumber="376">18.6</td>
<td id="58295DC3FF9BC0CEFCF720B9BAFEFB38" box="[847,974,1193,1215]" gridcol="2" gridrow="11" pageId="9" pageNumber="376">15.7–19.2</td>
<td id="58295DC3FF9BC0CEFB8C20B9BD5EFB38" box="[1076,1134,1193,1215]" gridcol="3" gridrow="11" pageId="9" pageNumber="376">17.4</td>
<td id="58295DC3FF9BC0CEFB5420B9BC28FB38" box="[1260,1304,1193,1215]" gridcol="4" gridrow="11" pageId="9" pageNumber="376">0.9</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2720C2BC28FB6F" box="[159,1304,1234,1256]" gridrow="12" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2720C2B8E1FB6F" box="[159,465,1234,1256]" gridcol="0" gridrow="12" pageId="9" pageNumber="376">Caudal-peduncle depth</th>
<td id="58295DC3FF9BC0CEFDD220C2BBF8FB6F" box="[618,712,1234,1256]" gridcol="1" gridrow="12" pageId="9" pageNumber="376">10.5</td>
<td id="58295DC3FF9BC0CEFCF720C2BAFEFB6F" box="[847,974,1234,1256]" gridcol="2" gridrow="12" pageId="9" pageNumber="376">10.4–11.5</td>
<td id="58295DC3FF9BC0CEFB8C20C2BD5EFB6F" box="[1076,1134,1234,1256]" gridcol="3" gridrow="12" pageId="9" pageNumber="376">11.0</td>
<td id="58295DC3FF9BC0CEFB5420C2BC28FB6F" box="[1260,1304,1234,1256]" gridcol="4" gridrow="12" pageId="9" pageNumber="376">0.3</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2720ECBC28FA95" box="[159,1304,1276,1298]" gridrow="13" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2720ECB8E1FA95" box="[159,465,1276,1298]" gridcol="0" gridrow="13" pageId="9" pageNumber="376">Predorsal length</th>
<td id="58295DC3FF9BC0CEFDD220ECBBF8FA95" box="[618,712,1276,1298]" gridcol="1" gridrow="13" pageId="9" pageNumber="376">43.2</td>
<td id="58295DC3FF9BC0CEFCF720ECBAFEFA95" box="[847,974,1276,1298]" gridcol="2" gridrow="13" pageId="9" pageNumber="376">43.2–47.5</td>
<td id="58295DC3FF9BC0CEFB8C20ECBD5EFA95" box="[1076,1134,1276,1298]" gridcol="3" gridrow="13" pageId="9" pageNumber="376">44.9</td>
<td id="58295DC3FF9BC0CEFB5420ECBC28FA95" box="[1260,1304,1276,1298]" gridcol="4" gridrow="13" pageId="9" pageNumber="376">1.1</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272135BC28FABC" box="[159,1304,1317,1339]" gridrow="14" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272135B8E1FABC" box="[159,465,1317,1339]" gridcol="0" gridrow="14" pageId="9" pageNumber="376">Prepectoral length</th>
<td id="58295DC3FF9BC0CEFDD22135BBF8FABC" box="[618,712,1317,1339]" gridcol="1" gridrow="14" pageId="9" pageNumber="376">21.7</td>
<td id="58295DC3FF9BC0CEFCF72135BAFEFABC" box="[847,974,1317,1339]" gridcol="2" gridrow="14" pageId="9" pageNumber="376">19.1–23.0</td>
<td id="58295DC3FF9BC0CEFB8C2135BD5EFABC" box="[1076,1134,1317,1339]" gridcol="3" gridrow="14" pageId="9" pageNumber="376">21.4</td>
<td id="58295DC3FF9BC0CEFB542135BC28FABC" box="[1260,1304,1317,1339]" gridcol="4" gridrow="14" pageId="9" pageNumber="376">1.0</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF27215EBC28FAE3" box="[159,1304,1358,1380]" gridrow="15" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF27215EB8E1FAE3" box="[159,465,1358,1380]" gridcol="0" gridrow="15" pageId="9" pageNumber="376">Prepelvic length</th>
<td id="58295DC3FF9BC0CEFDD2215EBBF8FAE3" box="[618,712,1358,1380]" gridcol="1" gridrow="15" pageId="9" pageNumber="376">49.0</td>
<td id="58295DC3FF9BC0CEFCF7215EBAFEFAE3" box="[847,974,1358,1380]" gridcol="2" gridrow="15" pageId="9" pageNumber="376">49.0–52.4</td>
<td id="58295DC3FF9BC0CEFB8C215EBD5EFAE3" box="[1076,1134,1358,1380]" gridcol="3" gridrow="15" pageId="9" pageNumber="376">50.6</td>
<td id="58295DC3FF9BC0CEFB54215EBC28FAE3" box="[1260,1304,1358,1380]" gridcol="4" gridrow="15" pageId="9" pageNumber="376">1.2</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272168BC28FA09" box="[159,1304,1400,1422]" gridrow="16" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272168B8E1FA09" box="[159,465,1400,1422]" gridcol="0" gridrow="16" pageId="9" pageNumber="376">Preanal length</th>
<td id="58295DC3FF9BC0CEFDD22168BBF8FA09" box="[618,712,1400,1422]" gridcol="1" gridrow="16" pageId="9" pageNumber="376">73.4</td>
<td id="58295DC3FF9BC0CEFCF72168BAFEFA09" box="[847,974,1400,1422]" gridcol="2" gridrow="16" pageId="9" pageNumber="376">73.3–77.4</td>
<td id="58295DC3FF9BC0CEFB8C2168BD5EFA09" box="[1076,1134,1400,1422]" gridcol="3" gridrow="16" pageId="9" pageNumber="376">75.6</td>
<td id="58295DC3FF9BC0CEFB542168BC28FA09" box="[1260,1304,1400,1422]" gridcol="4" gridrow="16" pageId="9" pageNumber="376">1.4</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2721B1BC28FA30" box="[159,1304,1441,1463]" gridrow="17" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2721B1BC28FA30" box="[159,1304,1441,1463]" colspan="5" colspanRight="4" gridcol="0" gridrow="17" pageId="9" pageNumber="376">In percent of HL</th>
</tr>
<tr id="1BF834BFFF9BC0CEFF2721DABC28FA67" box="[159,1304,1482,1504]" gridrow="18" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2721DAB8E1FA67" box="[159,465,1482,1504]" gridcol="0" gridrow="18" pageId="9" pageNumber="376">Snout length</th>
<td id="58295DC3FF9BC0CEFDD221DABBF8FA67" box="[618,712,1482,1504]" gridcol="1" gridrow="18" pageId="9" pageNumber="376">47</td>
<td id="58295DC3FF9BC0CEFCF721DABAFEFA67" box="[847,974,1482,1504]" gridcol="2" gridrow="18" pageId="9" pageNumber="376">43–49</td>
<td id="58295DC3FF9BC0CEFB8C21DABD5EFA67" box="[1076,1134,1482,1504]" gridcol="3" gridrow="18" pageId="9" pageNumber="376">46.4</td>
<td id="58295DC3FF9BC0CEFB5421DABC28FA67" box="[1260,1304,1482,1504]" gridcol="4" gridrow="18" pageId="9" pageNumber="376">1.4</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2721E4BC28F98D" box="[159,1304,1524,1546]" gridrow="19" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2721E4B8E1F98D" box="[159,465,1524,1546]" gridcol="0" gridrow="19" pageId="9" pageNumber="376">Eye diameter</th>
<td id="58295DC3FF9BC0CEFDD221E4BBF8F98D" box="[618,712,1524,1546]" gridcol="1" gridrow="19" pageId="9" pageNumber="376">13</td>
<td id="58295DC3FF9BC0CEFCF721E4BAFEF98D" box="[847,974,1524,1546]" gridcol="2" gridrow="19" pageId="9" pageNumber="376">13–18</td>
<td id="58295DC3FF9BC0CEFB8C21E4BD5EF98D" box="[1076,1134,1524,1546]" gridcol="3" gridrow="19" pageId="9" pageNumber="376">15.3</td>
<td id="58295DC3FF9BC0CEFB5421E4BC28F98D" box="[1260,1304,1524,1546]" gridcol="4" gridrow="19" pageId="9" pageNumber="376">1.3</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF27220DBC28F9B4" box="[159,1304,1565,1587]" gridrow="20" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF27220DB8E1F9B4" box="[159,465,1565,1587]" gridcol="0" gridrow="20" pageId="9" pageNumber="376">Interorbital distance</th>
<td id="58295DC3FF9BC0CEFDD2220DBBF8F9B4" box="[618,712,1565,1587]" gridcol="1" gridrow="20" pageId="9" pageNumber="376">42</td>
<td id="58295DC3FF9BC0CEFCF7220DBAFEF9B4" box="[847,974,1565,1587]" gridcol="2" gridrow="20" pageId="9" pageNumber="376">42–46</td>
<td id="58295DC3FF9BC0CEFB8C220DBD5EF9B4" box="[1076,1134,1565,1587]" gridcol="3" gridrow="20" pageId="9" pageNumber="376">44.0</td>
<td id="58295DC3FF9BC0CEFB54220DBC28F9B4" box="[1260,1304,1565,1587]" gridcol="4" gridrow="20" pageId="9" pageNumber="376">1.2</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272256BC28F9DB" box="[159,1304,1606,1628]" gridrow="21" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272256B8E1F9DB" box="[159,465,1606,1628]" gridcol="0" gridrow="21" pageId="9" pageNumber="376">Disc length</th>
<td id="58295DC3FF9BC0CEFDD22256BBF8F9DB" box="[618,712,1606,1628]" gridcol="1" gridrow="21" pageId="9" pageNumber="376">52</td>
<td id="58295DC3FF9BC0CEFCF72256BAFEF9DB" box="[847,974,1606,1628]" gridcol="2" gridrow="21" pageId="9" pageNumber="376">49–58</td>
<td id="58295DC3FF9BC0CEFB8C2256BD5EF9DB" box="[1076,1134,1606,1628]" gridcol="3" gridrow="21" pageId="9" pageNumber="376">54.4</td>
<td id="58295DC3FF9BC0CEFB542256BC28F9DB" box="[1260,1304,1606,1628]" gridcol="4" gridrow="21" pageId="9" pageNumber="376">2.3</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272260BC28F901" box="[159,1304,1648,1670]" gridrow="22" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272260B8E1F901" box="[159,465,1648,1670]" gridcol="0" gridrow="22" pageId="9" pageNumber="376">Disc width</th>
<td id="58295DC3FF9BC0CEFDD22260BBF8F901" box="[618,712,1648,1670]" gridcol="1" gridrow="22" pageId="9" pageNumber="376">67</td>
<td id="58295DC3FF9BC0CEFCF72260BAFEF901" box="[847,974,1648,1670]" gridcol="2" gridrow="22" pageId="9" pageNumber="376">66–77</td>
<td id="58295DC3FF9BC0CEFB8C2260BD5EF901" box="[1076,1134,1648,1670]" gridcol="3" gridrow="22" pageId="9" pageNumber="376">70.3.</td>
<td id="58295DC3FF9BC0CEFB542260BC28F901" box="[1260,1304,1648,1670]" gridcol="4" gridrow="22" pageId="9" pageNumber="376">3.0</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF272289BC28F928" box="[159,1304,1689,1711]" gridrow="23" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272289B8E1F928" box="[159,465,1689,1711]" gridcol="0" gridrow="23" pageId="9" pageNumber="376">Callous-pad length</th>
<td id="58295DC3FF9BC0CEFDD22289BBF8F928" box="[618,712,1689,1711]" gridcol="1" gridrow="23" pageId="9" pageNumber="376">35</td>
<td id="58295DC3FF9BC0CEFCF72289BAFEF928" box="[847,974,1689,1711]" gridcol="2" gridrow="23" pageId="9" pageNumber="376">33–40</td>
<td id="58295DC3FF9BC0CEFB8C2289BD5EF928" box="[1076,1134,1689,1711]" gridcol="3" gridrow="23" pageId="9" pageNumber="376">37.0</td>
<td id="58295DC3FF9BC0CEFB542289BC28F928" box="[1260,1304,1689,1711]" gridcol="4" gridrow="23" pageId="9" pageNumber="376">2.0</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2722D2BC28F95F" box="[159,1304,1730,1752]" gridrow="24" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2722D2B8E1F95F" box="[159,465,1730,1752]" gridcol="0" gridrow="24" pageId="9" pageNumber="376">Callous-pad width</th>
<td id="58295DC3FF9BC0CEFDD222D2BBF8F95F" box="[618,712,1730,1752]" gridcol="1" gridrow="24" pageId="9" pageNumber="376">45</td>
<td id="58295DC3FF9BC0CEFCF722D2BAFEF95F" box="[847,974,1730,1752]" gridcol="2" gridrow="24" pageId="9" pageNumber="376">44–50</td>
<td id="58295DC3FF9BC0CEFB8C22D2BD5EF95F" box="[1076,1134,1730,1752]" gridcol="3" gridrow="24" pageId="9" pageNumber="376">47.1</td>
<td id="58295DC3FF9BC0CEFB5422D2BC28F95F" box="[1260,1304,1730,1752]" gridcol="4" gridrow="24" pageId="9" pageNumber="376">2.0</td>
</tr>
<tr id="1BF834BFFF9BC0CEFF2722FCBC28F885" box="[159,1304,1772,1794]" gridrow="25" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF2722FCBC28F885" box="[159,1304,1772,1794]" colspan="5" colspanRight="4" gridcol="0" gridrow="25" pageId="9" pageNumber="376">In percent of pelvic-anal distance</th>
</tr>
<tr id="1BF834BFFF9BC0CEFF272305BC28F8AC" box="[159,1304,1813,1835]" gridrow="26" pageId="9" pageNumber="376">
<th id="58295DC3FF9BC0CEFF272305B8E1F8AC" box="[159,465,1813,1835]" gridcol="0" gridrow="26" pageId="9" pageNumber="376">Distance from anus to anal fin</th>
<td id="58295DC3FF9BC0CEFDD22305BBF8F8AC" box="[618,712,1813,1835]" gridcol="1" gridrow="26" pageId="9" pageNumber="376">20</td>
<td id="58295DC3FF9BC0CEFCF72305BAFEF8AC" box="[847,974,1813,1835]" gridcol="2" gridrow="26" pageId="9" pageNumber="376">19–24</td>
<td id="58295DC3FF9BC0CEFB8C2305BD5EF8AC" box="[1076,1134,1813,1835]" gridcol="3" gridrow="26" pageId="9" pageNumber="376">22.0</td>
<td id="58295DC3FF9BC0CEFB542305BC28F8AC" box="[1260,1304,1813,1835]" gridcol="4" gridrow="26" pageId="9" pageNumber="376">1.5</td>
</tr>
</table>
</paragraph>
<paragraph id="A57736FDFF9B3F3BFF7F237FBA67FF7F" blockId="9.[151,1437,1903,2037]" lastBlockId="10.[151,1437,151,464]" lastPageId="10" lastPageNumber="377" pageId="9" pageNumber="376">Dorsal fin with 3 simple and 8½ branched rays; distal margin slightly concave; first branched ray longest, almost equal to head length; origin closer to snout tip than to caudal-fin base. Pectoral fin with 1 simple and 13 (10), or 14 (6*) branched rays; length shorter than head length; distal tip not reaching pelvic-fin origin, reaching vertically to dorsal-fin origin. Pelvic fin with 1 simple and 8 branched rays; length shorter than head length, equal to pectoral-fin length; distal tip reaching beyond anus; origin closer to caudal-fin base than to snout tip. Anal fin with 3 simple and 5½ branched rays; distal tip reaching base of caudal fin; origin closer to caudal-fin base than to pelvic-fin origin. Caudal fin forked; lower lobe slightly longer.</paragraph>
<paragraph id="A57736FDFF983F3BFF7F2514BA25FE48" blockId="10.[151,1437,151,464]" pageId="10" pageNumber="377">
Lateral line complete and horizontal; with 32 (3), 33 (10), or 34 (3*) scales plus 2 scales on the caudal-fin base. Transverse scale rows above lateral line 4 (8) or 4½ (8*), between lateral line and pelvic-fin origin 2½ (3*) or 3 (13). Circumpeduncular scales 12. Predorsal scales 9 (7*), 10 (8), or 11 (1); smaller than flank scales. Chest and belly scaled; scales on chest smaller than on belly. One long axillary scale at base of pelvic fin. Anus situated very closer to anal-fin origin than to pelvic-fin origin (see
<figureCitation id="3DF32A78FF983F3BFD4B2584BA07FE2A" box="[755,823,404,429]" captionStart="FIGURE 5" captionStartId="8.[151,250,1899,1921]" captionTargetBox="[151,1436,1164,1878]" captionTargetId="figure@8.[151,1436,1164,1878]" captionTargetPageId="8" captionText="FIGURE 5. Garra yajiangensis, holotype, IHB 20161577, 165.0 mm SL; China: Tibet: lower Yarlung Tsangpo River." httpUri="https://zenodo.org/record/2615439/files/figure.png" pageId="10" pageNumber="377">Fig. 5</figureCitation>
, distance from anus to anal fin 19–24% of pelvic-anal distance), separated from anal-fin origin by 3 scale rows.
</paragraph>
<caption id="F1B76675FF983F3BFF2F27EAB892FBA8" httpUri="https://zenodo.org/record/2615443/files/figure.png" pageId="10" pageNumber="377" startId="10.[151,250,1018,1040]" targetBox="[150,1436,517,997]" targetPageId="10">
<paragraph id="A57736FDFF983F3BFF2F27EAB892FBA8" blockId="10.[151,1436,1018,1071]" pageId="10" pageNumber="377">
<emphasis id="97BCEAEFFF983F3BFF2F27EAB823FB97" bold="true" box="[151,275,1018,1040]" pageId="10" pageNumber="377">FIGURE 6.</emphasis>
<taxonomicName id="62C84D7EFF983F3BFEA327EBB8D0FB97" box="[283,480,1019,1040]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="377" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF983F3BFEA327EBB8D0FB97" box="[283,480,1019,1040]" italics="true" pageId="10" pageNumber="377">Garra yajiangensis</emphasis>
</taxonomicName>
, holotype, IHB 20161577, 165.0 mm SL; China: Tibet: lower Yarlung Tsangpo River; dorsal and ventral views of head.
</paragraph>
</caption>
<caption id="F1B76675FF983F3BFF2F23DFBA8FF862" box="[151,959,1999,2022]" httpUri="https://zenodo.org/record/2615445/files/figure.png" pageId="10" pageNumber="377" startId="10.[151,250,1999,2021]" targetBox="[150,1436,1121,1978]" targetPageId="10">
<paragraph id="A57736FDFF983F3BFF2F23DFBA8FF862" blockId="10.[151,959,1999,2022]" box="[151,959,1999,2022]" pageId="10" pageNumber="377">
<emphasis id="97BCEAEFFF983F3BFF2F23DFB822F862" bold="true" box="[151,274,1999,2021]" pageId="10" pageNumber="377">FIGURE 7.</emphasis>
Habitat of
<taxonomicName id="62C84D7EFF983F3BFE3D23C0BB7AF861" box="[389,586,2000,2022]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="377" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF983F3BFE3D23C0BB7AF861" box="[389,586,2000,2022]" italics="true" pageId="10" pageNumber="377">Garra yajiangensis</emphasis>
</taxonomicName>
in the lower Yarlung Tsangpo River.
</paragraph>
</caption>
<paragraph id="A57736FDFF993F3AFF7F2487B84DFE9A" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="378">
<emphasis id="97BCEAEFFF993F3AFF7F2487B87FFF37" bold="true" box="[199,335,151,176]" pageId="11" pageNumber="378">Coloration.</emphasis>
In preserved specimens, head, dorsum, and side of head pale brown or gray. Mouth, chest, and belly faint yellow or light brown. Dorsal, pectoral, pelvic, and anal fins gray or grayish white; pectoral fin darker than other fins. A gray submarginal band on distal half of dorsal fin. Lower lobe and distal margin of upper lobe of caudal fin dark gray.
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF993F3AFF7F2537B802FEE3" pageId="11" pageNumber="378" type="distribution">
<paragraph id="A57736FDFF993F3AFF7F2537B802FEE3" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="378">
<emphasis id="97BCEAEFFF993F3AFF7F2537B850FEC7" bold="true" box="[199,352,295,320]" pageId="11" pageNumber="378">Distribution.</emphasis>
Known only from the lower Yarlung Tsangpo River drainage in Motuo County, southeastern
<collectingRegion id="670CF81FFF993F3AFF2F255CB9E1FEE2" box="[151,209,332,357]" country="China" name="Xizang" pageId="11" pageNumber="378">Tibet</collectingRegion>
(
<figureCitation id="3DF32A78FF993F3AFF58255CB814FEE3" box="[224,292,332,357]" captionStart="FIGURE 3" captionStartId="7.[151,250,1095,1117]" captionTargetBox="[219,1355,347,1071]" captionTargetId="figure@7.[208,1379,337,1081]" captionTargetPageId="7" captionText="FIGURE 3. Map showing the sampling sites of Garra motuoensis (▲) and Garra yajiangensis (?)." httpUri="https://zenodo.org/record/2615435/files/figure.png" pageId="11" pageNumber="378">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF993F3AFF7F257FB887FD9F" pageId="11" pageNumber="378" type="biology_ecology">
<paragraph id="A57736FDFF993F3AFF7F257FB887FD9F" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="378">
<emphasis id="97BCEAEFFF993F3AFF7F257FB81BFE0F" bold="true" box="[199,299,367,392]" pageId="11" pageNumber="378">Ecology.</emphasis>
This new species was primarily caught in the mainstem of the Yarlung Tsangpo (occasionally found in the tributaries), where it inhabits swift-flowing waters with numerous rocks (
<figureCitation id="3DF32A78FF993F3AFBB12584BD7DFE2B" box="[1033,1101,404,429]" captionStart="FIGURE 7" captionStartId="10.[151,250,1999,2021]" captionTargetBox="[150,1436,1121,1978]" captionTargetId="figure@10.[150,1436,1121,1978]" captionTargetPageId="10" captionText="FIGURE 7. Habitat of Garra yajiangensis in the lower Yarlung Tsangpo River." httpUri="https://zenodo.org/record/2615445/files/figure.png" pageId="11" pageNumber="378">Fig. 7</figureCitation>
). The spawning period occurs from March to May when water temperature ranges from 12–17°C according to macroscopic visual examination of the gonads. The eggs are yellow and demersal. Examinations of intestinal contents show that this species mainly feeds on periphytic algae.
</paragraph>
</subSubSection>
<subSubSection id="EDD26576FF993F3DFF7F2634BDE5FF7F" lastPageId="12" lastPageNumber="379" pageId="11" pageNumber="378" type="etymology">
<paragraph id="A57736FDFF993F3AFF7F2634B81DFDE7" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="378">
<emphasis id="97BCEAEFFF993F3AFF7F2634B87EFDBA" bold="true" box="[199,334,548,573]" pageId="11" pageNumber="378">Etymology.</emphasis>
Named after its
<typeStatus id="7A73885FFF993F3AFDA82635BB70FDBA" box="[528,576,549,573]" pageId="11" pageNumber="378">type</typeStatus>
locality, Yajiang (the Chinese abbreviation of the Yarlung Tsangpo River). An adjective.
</paragraph>
<paragraph id="A57736FDFF993F3AFF7F267CBB80FCDB" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="378">
<emphasis id="97BCEAEFFF993F3AFF7F267CBB69FD02" bold="true" box="[199,601,620,645]" pageId="11" pageNumber="378">Molecular phylogenetic analysis.</emphasis>
After alignment, 1137 bps of Cyt
<emphasis id="97BCEAEFFF993F3AFBBC267CBD22FD03" box="[1028,1042,620,644]" italics="true" pageId="11" pageNumber="378">b</emphasis>
gene sequences belonging to 86 individuals of 40 species of
<taxonomicName id="62C84D7EFF993F3AFE662680BB14FD20" authorityName="Hamilton" authorityYear="1822" box="[478,548,656,679]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="genus">
<emphasis id="97BCEAEFFF993F3AFE662680BB14FD20" box="[478,548,656,679]" italics="true" pageId="11" pageNumber="378">Garra</emphasis>
</taxonomicName>
and two species of
<taxonomicName id="62C84D7EFF993F3AFCB52680BA63FD2F" box="[781,851,656,680]" class="Actinopterygii" family="Cyprinidae" genus="Labeo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="genus">
<emphasis id="97BCEAEFFF993F3AFCB52680BA63FD2F" box="[781,851,656,680]" italics="true" pageId="11" pageNumber="378">Labeo</emphasis>
</taxonomicName>
were used for analysis. Of all 1137 bps, 182 were variable sites and 151 were parsimony informative sites. Based on the K2P model, paired genetic divergences between these two new species and their congeners are given in
<tableCitation id="E84A0346FF993F3AFC2A26C7BADBFD77" box="[914,1003,727,752]" captionStart="TABLE 5" captionStartId="11.[151,239,904,927]" captionText="TABLE 5. The K2P genetic divergences (in %) and S.E. (in %, parenthetical) between G. motuoensis, G. yajiangensis, and sampled congeners known from the Brahmaputra and adjacent river drainages." pageId="11" pageNumber="378">Table 5</tableCitation>
. The genetic divergences between
<taxonomicName id="62C84D7EFF993F3AFA3C26C8B825FC93" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF993F3AFA3C26C8B825FC93" italics="true" pageId="11" pageNumber="378">G. motuoensis</emphasis>
</taxonomicName>
and its sampled congeners known from the Brahmaputra and the adjacent river drainages ranged from 2.6% (vs.
<taxonomicName id="62C84D7EFF993F3AFEBF2730B8BFFCBF" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[263,399,799,824]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="97BCEAEFFF993F3AFEBF2730B82DFCB0" box="[263,285,800,823]" italics="true" pageId="11" pageNumber="378">G.</emphasis>
cf.
<emphasis id="97BCEAEFFF993F3AFEF12730B8BFFCBF" box="[329,399,800,824]" italics="true" pageId="11" pageNumber="378">gotyla</emphasis>
</taxonomicName>
A) to 19.8% (vs.
<taxonomicName id="62C84D7EFF993F3AFDEC2730BBEEFCBF" authorityName="Kullander &amp; Fang" authorityYear="2004" box="[596,734,800,824]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="poecilura">
<emphasis id="97BCEAEFFF993F3AFDEC2730BBEEFCBF" box="[596,734,800,824]" italics="true" pageId="11" pageNumber="378">G. poecilura</emphasis>
</taxonomicName>
); between
<taxonomicName id="62C84D7EFF993F3AFCE12730BD33FCB0" box="[857,1027,800,823]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF993F3AFCE12730BD33FCB0" box="[857,1027,800,823]" italics="true" pageId="11" pageNumber="378">G. yajiangensis</emphasis>
</taxonomicName>
and its congeners ranged from 6.3% (vs.
<taxonomicName id="62C84D7EFF993F3AFF7F2754B87FFCDB" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[199,335,836,861]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="97BCEAEFFF993F3AFF7F2754B9EFFCDC" box="[199,223,836,859]" italics="true" pageId="11" pageNumber="378">G.</emphasis>
cf.
<emphasis id="97BCEAEFFF993F3AFEB12754B87FFCDB" box="[265,335,836,860]" italics="true" pageId="11" pageNumber="378">gotyla</emphasis>
</taxonomicName>
B) to 16.1% (vs.
<taxonomicName id="62C84D7EFF993F3AFDAE2754BB90FCDB" authorityName="Kullander &amp; Fang" authorityYear="2004" box="[534,672,836,860]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="poecilura">
<emphasis id="97BCEAEFFF993F3AFDAE2754BB90FCDB" box="[534,672,836,860]" italics="true" pageId="11" pageNumber="378">G. poecilura</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="F1B76675FF993F3AFF2F2798BD3CFC38" pageId="11" pageNumber="378">
<paragraph id="A57736FDFF993F3AFF2F2798BD3CFC38" blockId="11.[151,1436,904,959]" pageId="11" pageNumber="378">
<emphasis id="97BCEAEFFF993F3AFF2F2798B83CFC18" bold="true" box="[151,268,904,927]" pageId="11" pageNumber="378">TABLE 5.</emphasis>
The K2P genetic divergences (in %) and S.E. (in %, parenthetical) between
<taxonomicName id="62C84D7EFF993F3AFBF42798BDD3FC18" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[1100,1251,904,927]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF993F3AFBF42798BDD3FC18" box="[1100,1251,904,927]" italics="true" pageId="11" pageNumber="378">G. motuoensis</emphasis>
</taxonomicName>
,
<taxonomicName id="62C84D7EFF993F3AFB4A2798BCA5FC18" box="[1266,1429,904,927]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF993F3AFB4A2798BCA5FC18" box="[1266,1429,904,927]" italics="true" pageId="11" pageNumber="378">G. yajiangensis</emphasis>
</taxonomicName>
, and sampled congeners known from the Brahmaputra and adjacent river drainages.
</paragraph>
</caption>
<paragraph id="A57736FDFF993F3AFDF327CBBD6AF8AC" pageId="11" pageNumber="378">
<table id="D7C8C45DFF99C0CEFF2727CBBDA2F8AC" box="[159,1170,987,1835]" gridcols="3" gridrows="21" pageId="11" pageNumber="378">
<tr id="1BF834BFFF99C0CEFF2727CBBDA2FC77" box="[159,1170,987,1008]" gridrow="0" pageId="11" pageNumber="378" rowspan-0="1">
<th id="58295DC3FF99C0CEFDF327CBBBE9FC77" box="[587,729,987,1008]" gridcol="1" gridrow="0" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFDF327CBBBE9FC77" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[587,729,987,1008]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF993F3AFDF327CBBBE9FC77" box="[587,729,987,1008]" italics="true" pageId="11" pageNumber="378">G. motuoensis</emphasis>
</taxonomicName>
</th>
<th id="58295DC3FF99C0CEFC4F27CBBDA2FC77" box="[1015,1170,987,1008]" gridcol="2" gridrow="0" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFC4F27CBBDA2FC77" box="[1015,1170,987,1008]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF993F3AFC4F27CBBDA2FC77" box="[1015,1170,987,1008]" italics="true" pageId="11" pageNumber="378">G. yajiangensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="1BF834BFFF99C0CEFF272014BDA2FB9D" box="[159,1170,1028,1050]" gridrow="1" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272014B868FB9D" box="[159,344,1028,1050]" gridcol="0" gridrow="1" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF272014B81DFB9D" authorityName="Gong &amp; Freyhof &amp; Wang &amp; Liu &amp; Liu &amp; Lin &amp; Jiang &amp; Liu" authorityYear="2018" box="[159,301,1028,1050]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="motuoensis">
<emphasis id="97BCEAEFFF993F3AFF272014B81DFB9D" box="[159,301,1028,1050]" italics="true" pageId="11" pageNumber="378">G. motuoensis</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF32014BBE9FB9D" box="[587,729,1028,1050]" gridcol="1" gridrow="1" pageId="11" pageNumber="378">–</td>
<td id="58295DC3FF99C0CEFC4F2014BDA2FB9D" box="[1015,1170,1028,1050]" gridcol="2" gridrow="1" pageId="11" pageNumber="378">10.1(1.0)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF27203DBDA2FBC4" box="[159,1170,1069,1091]" gridrow="2" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF27203DB868FBC4" box="[159,344,1069,1091]" gridcol="0" gridrow="2" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF27203EB80AFBC4" box="[159,314,1070,1091]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="yajiangensis">
<emphasis id="97BCEAEFFF993F3AFF27203EB80AFBC4" box="[159,314,1070,1091]" italics="true" pageId="11" pageNumber="378">G. yajiangensis</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF3203DBBE9FBC4" box="[587,729,1069,1091]" gridcol="1" gridrow="2" pageId="11" pageNumber="378">10.1 (1.0)</td>
<td id="58295DC3FF99C0CEFC4F203DBDA2FBC4" box="[1015,1170,1069,1091]" gridcol="2" gridrow="2" pageId="11" pageNumber="378">–</td>
</tr>
<tr id="1BF834BFFF99C0CEFF272046BDA2FBEB" box="[159,1170,1110,1132]" gridrow="3" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272046B868FBEB" box="[159,344,1110,1132]" gridcol="0" gridrow="3" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF272047B82CFBEB" authorityName="Zhang" authorityYear="2002" box="[159,284,1110,1132]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="bispinosa">
<emphasis id="97BCEAEFFF993F3AFF272047B82CFBEB" box="[159,284,1110,1132]" italics="true" pageId="11" pageNumber="378">G. bispinosa</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF32046BBE9FBEB" box="[587,729,1110,1132]" gridcol="1" gridrow="3" pageId="11" pageNumber="378">6.0 (0.8)</td>
<td id="58295DC3FF99C0CEFC4F2046BDA2FBEB" box="[1015,1170,1110,1132]" gridcol="2" gridrow="3" pageId="11" pageNumber="378">10.8 (1.1)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF272090BDA2FB11" box="[159,1170,1152,1174]" gridrow="4" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272090B868FB11" box="[159,344,1152,1174]" gridcol="0" gridrow="4" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF272091B87DFB11" authorityName="Hora" authorityYear="1921" box="[159,333,1152,1174]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="97BCEAEFFF993F3AFF272091B985FB11" box="[159,181,1153,1174]" italics="true" pageId="11" pageNumber="378">G.</emphasis>
cf.
<emphasis id="97BCEAEFFF993F3AFF632090B87DFB11" box="[219,333,1152,1174]" italics="true" pageId="11" pageNumber="378">annandalei</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF32090BBE9FB11" box="[587,729,1152,1174]" gridcol="1" gridrow="4" pageId="11" pageNumber="378">12.4 (1.1)</td>
<td id="58295DC3FF99C0CEFC4F2090BDA2FB11" box="[1015,1170,1152,1174]" gridcol="2" gridrow="4" pageId="11" pageNumber="378">10.6 (1.0)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF2720B9BDA2FB38" box="[159,1170,1193,1215]" gridrow="5" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF2720B9B868FB38" box="[159,344,1193,1215]" gridcol="0" gridrow="5" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF2720BAB82AFB38" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[159,282,1193,1215]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="97BCEAEFFF993F3AFF2720BAB985FB38" box="[159,181,1194,1215]" italics="true" pageId="11" pageNumber="378">G.</emphasis>
cf.
<emphasis id="97BCEAEFFF993F3AFF6320B9B82AFB38" box="[219,282,1193,1215]" italics="true" pageId="11" pageNumber="378">gotyla</emphasis>
</taxonomicName>
A
</th>
<td id="58295DC3FF99C0CEFDF320B9BBE9FB38" box="[587,729,1193,1215]" gridcol="1" gridrow="5" pageId="11" pageNumber="378">2.6 (0.5)</td>
<td id="58295DC3FF99C0CEFC4F20B9BDA2FB38" box="[1015,1170,1193,1215]" gridcol="2" gridrow="5" pageId="11" pageNumber="378">10.2 (1.0)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF2720C2BDA2FB6F" box="[159,1170,1234,1256]" gridrow="6" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF2720C2B868FB6F" box="[159,344,1234,1256]" gridcol="0" gridrow="6" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF2720C3B82AFB6F" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[159,282,1234,1256]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="gotyla">
<emphasis id="97BCEAEFFF993F3AFF2720C3B985FB6F" box="[159,181,1235,1256]" italics="true" pageId="11" pageNumber="378">G.</emphasis>
cf.
<emphasis id="97BCEAEFFF993F3AFF6320C2B82AFB6F" box="[219,282,1234,1256]" italics="true" pageId="11" pageNumber="378">gotyla</emphasis>
</taxonomicName>
B
</th>
<td id="58295DC3FF99C0CEFDF320C2BBE9FB6F" box="[587,729,1234,1256]" gridcol="1" gridrow="6" pageId="11" pageNumber="378">9.8 (1.0)</td>
<td id="58295DC3FF99C0CEFC4F20C2BDA2FB6F" box="[1015,1170,1234,1256]" gridcol="2" gridrow="6" pageId="11" pageNumber="378">6.3 (0.7)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF2720ECBDA2FA95" box="[159,1170,1276,1298]" gridrow="7" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF2720ECB868FA95" box="[159,344,1276,1298]" gridcol="0" gridrow="7" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF2720ECB827FA95" authorityName="Hora" authorityYear="1921" box="[159,279,1276,1298]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="kempi">
<emphasis id="97BCEAEFFF993F3AFF2720ECB985FA96" box="[159,181,1276,1297]" italics="true" pageId="11" pageNumber="378">G.</emphasis>
cf.
<emphasis id="97BCEAEFFF993F3AFF6320ECB827FA95" box="[219,279,1276,1298]" italics="true" pageId="11" pageNumber="378">kempi</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF320ECBBE9FA95" box="[587,729,1276,1298]" gridcol="1" gridrow="7" pageId="11" pageNumber="378">15.3 (1.2)</td>
<td id="58295DC3FF99C0CEFC4F20ECBDA2FA95" box="[1015,1170,1276,1298]" gridcol="2" gridrow="7" pageId="11" pageNumber="378">13.1 (1.1)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF272135BDA2FABC" box="[159,1170,1317,1339]" gridrow="8" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272135B868FABC" box="[159,344,1317,1339]" gridcol="0" gridrow="8" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF272136B836FABC" authorityName="Deng, Cao &amp; Zhang" authorityYear="2018" box="[159,262,1317,1339]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="dengba">
<emphasis id="97BCEAEFFF993F3AFF272136B836FABC" box="[159,262,1317,1339]" italics="true" pageId="11" pageNumber="378">G. dengba</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF32135BBE9FABC" box="[587,729,1317,1339]" gridcol="1" gridrow="8" pageId="11" pageNumber="378">14.0 (1.3)</td>
<td id="58295DC3FF99C0CEFC4F2135BDA2FABC" box="[1015,1170,1317,1339]" gridcol="2" gridrow="8" pageId="11" pageNumber="378">11.7 (1.0)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF27215EBDA2FAE3" box="[159,1170,1358,1380]" gridrow="9" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF27215EB868FAE3" box="[159,344,1358,1380]" gridcol="0" gridrow="9" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF27215FB804FAE3" baseAuthorityName="Chen, Pan, Xiao &amp; Yang" baseAuthorityYear="2012" box="[159,308,1358,1380]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="dulongensis">
<emphasis id="97BCEAEFFF993F3AFF27215FB804FAE3" box="[159,308,1358,1380]" italics="true" pageId="11" pageNumber="378">G. dulongensis</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF3215EBBE9FAE3" box="[587,729,1358,1380]" gridcol="1" gridrow="9" pageId="11" pageNumber="378">12.7 (1.2)</td>
<td id="58295DC3FF99C0CEFC4F215EBDA2FAE3" box="[1015,1170,1358,1380]" gridcol="2" gridrow="9" pageId="11" pageNumber="378">11.1 (1.0)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF272168BDA2FA09" box="[159,1170,1400,1422]" gridrow="10" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272168B868FA09" box="[159,344,1400,1422]" gridcol="0" gridrow="10" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF272169B83BFA09" authorityName="Kullander &amp; Fang" authorityYear="2004" box="[159,267,1400,1422]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="flavatra">
<emphasis id="97BCEAEFFF993F3AFF272169B83BFA09" box="[159,267,1400,1422]" italics="true" pageId="11" pageNumber="378">G. flavatra</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF32168BBE9FA09" box="[587,729,1400,1422]" gridcol="1" gridrow="10" pageId="11" pageNumber="378">16.6 (1.4)</td>
<td id="58295DC3FF99C0CEFC4F2168BDA2FA09" box="[1015,1170,1400,1422]" gridcol="2" gridrow="10" pageId="11" pageNumber="378">14.7 (1.2)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF2721B1BDA2FA30" box="[159,1170,1441,1463]" gridrow="11" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF2721B1B868FA30" box="[159,344,1441,1463]" gridcol="0" gridrow="11" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF2721B2B9C7FA30" authorityName="Hora" authorityYear="1921" box="[159,247,1441,1463]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="kempi">
<emphasis id="97BCEAEFFF993F3AFF2721B2B9C7FA30" box="[159,247,1441,1463]" italics="true" pageId="11" pageNumber="378">G. kempi</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF321B1BBE9FA30" box="[587,729,1441,1463]" gridcol="1" gridrow="11" pageId="11" pageNumber="378">12.2 (1.2)</td>
<td id="58295DC3FF99C0CEFC4F21B1BDA2FA30" box="[1015,1170,1441,1463]" gridcol="2" gridrow="11" pageId="11" pageNumber="378">10.9 (1.0)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF2721DABDA2FA67" box="[159,1170,1482,1504]" gridrow="12" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF2721DAB868FA67" box="[159,344,1482,1504]" gridcol="0" gridrow="12" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF2721DBB876FA67" baseAuthorityName="McClelland" baseAuthorityYear="1842" box="[159,326,1482,1504]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="lissorhynchus">
<emphasis id="97BCEAEFFF993F3AFF2721DBB876FA67" box="[159,326,1482,1504]" italics="true" pageId="11" pageNumber="378">G. lissorhynchus</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF321DABBE9FA67" box="[587,729,1482,1504]" gridcol="1" gridrow="12" pageId="11" pageNumber="378">16.0 (1.5)</td>
<td id="58295DC3FF99C0CEFC4F21DABDA2FA67" box="[1015,1170,1482,1504]" gridcol="2" gridrow="12" pageId="11" pageNumber="378">15.0 (1.3)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF2721E4BDA2F98D" box="[159,1170,1524,1546]" gridrow="13" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF2721E4B868F98D" box="[159,344,1524,1546]" gridcol="0" gridrow="13" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF2721E4B868F98D" authorityName="Yu, Wang, Xiong &amp; He" authorityYear="2016" box="[159,344,1524,1546]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="longchuanensis">
<emphasis id="97BCEAEFFF993F3AFF2721E4B868F98D" box="[159,344,1524,1546]" italics="true" pageId="11" pageNumber="378">G. longchuanensis</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF321E4BBE9F98D" box="[587,729,1524,1546]" gridcol="1" gridrow="13" pageId="11" pageNumber="378">9.5 (1.1)</td>
<td id="58295DC3FF99C0CEFC4F21E4BDA2F98D" box="[1015,1170,1524,1546]" gridcol="2" gridrow="13" pageId="11" pageNumber="378">6.8 (0.7)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF27220DBDA2F9B4" box="[159,1170,1565,1587]" gridrow="14" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF27220DB868F9B4" box="[159,344,1565,1587]" gridcol="0" gridrow="14" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF27220EB9CBF9B4" baseAuthorityName="McClelland" baseAuthorityYear="1838" box="[159,251,1566,1587]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="nasuta">
<emphasis id="97BCEAEFFF993F3AFF27220EB9CBF9B4" box="[159,251,1566,1587]" italics="true" pageId="11" pageNumber="378">G. nasuta</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF3220DBBE9F9B4" box="[587,729,1565,1587]" gridcol="1" gridrow="14" pageId="11" pageNumber="378">11.5 (1.2)</td>
<td id="58295DC3FF99C0CEFC4F220DBDA2F9B4" box="[1015,1170,1565,1587]" gridcol="2" gridrow="14" pageId="11" pageNumber="378">11.1 (1.0)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF272256BDA2F9DB" box="[159,1170,1606,1628]" gridrow="15" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272256B868F9DB" box="[159,344,1606,1628]" gridcol="0" gridrow="15" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF272257B82CF9DB" authorityName="Kullander &amp; Fang" authorityYear="2004" box="[159,284,1606,1628]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="poecilura">
<emphasis id="97BCEAEFFF993F3AFF272257B82CF9DB" box="[159,284,1606,1628]" italics="true" pageId="11" pageNumber="378">G. poecilura</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF32256BBE9F9DB" box="[587,729,1606,1628]" gridcol="1" gridrow="15" pageId="11" pageNumber="378">19.8 (1.5)</td>
<td id="58295DC3FF99C0CEFC4F2256BDA2F9DB" box="[1015,1170,1606,1628]" gridcol="2" gridrow="15" pageId="11" pageNumber="378">16.1 (1.4)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF272260BDA2F901" box="[159,1170,1648,1670]" gridrow="16" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272260B868F901" box="[159,344,1648,1670]" gridcol="0" gridrow="16" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF272261B819F901" authorityName="Wu &amp; Yao" authorityYear="1977" box="[159,297,1649,1670]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="qiaojiensis">
<emphasis id="97BCEAEFFF993F3AFF272261B819F901" box="[159,297,1649,1670]" italics="true" pageId="11" pageNumber="378">G. qiaojiensis</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF32260BBE9F901" box="[587,729,1648,1670]" gridcol="1" gridrow="16" pageId="11" pageNumber="378">7.3 (0.8)</td>
<td id="58295DC3FF99C0CEFC4F2260BDA2F901" box="[1015,1170,1648,1670]" gridcol="2" gridrow="16" pageId="11" pageNumber="378">9.5 (0.9)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF272289BDA2F928" box="[159,1170,1689,1711]" gridrow="17" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272289B868F928" box="[159,344,1689,1711]" gridcol="0" gridrow="17" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF27228AB80EF928" authorityName="Zhang" authorityYear="2006" box="[159,318,1689,1711]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="rotundinasus">
<emphasis id="97BCEAEFFF993F3AFF27228AB80EF928" box="[159,318,1689,1711]" italics="true" pageId="11" pageNumber="378">G. rotundinasus</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF32289BBE9F928" box="[587,729,1689,1711]" gridcol="1" gridrow="17" pageId="11" pageNumber="378">9.3 (0.9)</td>
<td id="58295DC3FF99C0CEFC4F2289BDA2F928" box="[1015,1170,1689,1711]" gridcol="2" gridrow="17" pageId="11" pageNumber="378">7.4 (0.8)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF2722D2BDA2F95F" box="[159,1170,1730,1752]" gridrow="18" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF2722D2B868F95F" box="[159,344,1730,1752]" gridcol="0" gridrow="18" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF2722D3B830F95F" authorityName="Kullander &amp; Fang" authorityYear="2004" box="[159,256,1730,1752]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="spilota">
<emphasis id="97BCEAEFFF993F3AFF2722D3B830F95F" box="[159,256,1730,1752]" italics="true" pageId="11" pageNumber="378">G. spilota</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF322D2BBE9F95F" box="[587,729,1730,1752]" gridcol="1" gridrow="18" pageId="11" pageNumber="378">12.6 (1.2)</td>
<td id="58295DC3FF99C0CEFC4F22D2BDA2F95F" box="[1015,1170,1730,1752]" gridcol="2" gridrow="18" pageId="11" pageNumber="378">12.1 (1.1)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF2722FCBDA2F885" box="[159,1170,1772,1794]" gridrow="19" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF2722FCB868F885" box="[159,344,1772,1794]" gridcol="0" gridrow="19" pageId="11" pageNumber="378">
<taxonomicName id="62C84D7EFF993F3AFF2722FCB811F885" authorityName="Sun, Li, Zhou &amp; Li" authorityYear="2018" box="[159,289,1772,1794]" class="Actinopterygii" family="Cyprinidae" genus="Garra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="378" phylum="Chordata" rank="species" species="surgifrons">
<emphasis id="97BCEAEFFF993F3AFF2722FCB811F885" box="[159,289,1772,1794]" italics="true" pageId="11" pageNumber="378">G. surgifrons</emphasis>
</taxonomicName>
</th>
<td id="58295DC3FF99C0CEFDF322FCBBE9F885" box="[587,729,1772,1794]" gridcol="1" gridrow="19" pageId="11" pageNumber="378">10.2 (1.1)</td>
<td id="58295DC3FF99C0CEFC4F22FCBDA2F885" box="[1015,1170,1772,1794]" gridcol="2" gridrow="19" pageId="11" pageNumber="378">10.2 (1.0)</td>
</tr>
<tr id="1BF834BFFF99C0CEFF272305BDA2F8AC" box="[159,1170,1813,1835]" gridrow="20" pageId="11" pageNumber="378">
<th id="58295DC3FF99C0CEFF272305B868F8AC" box="[159,344,1813,1835]" gridcol="0" gridrow="20" pageId="11" pageNumber="378">
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<emphasis id="97BCEAEFFF993F3AFF272306B867F8AC" box="[159,343,1813,1835]" italics="true" pageId="11" pageNumber="378">G. tengchongensis</emphasis>
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<td id="58295DC3FF99C0CEFDF32305BBE9F8AC" box="[587,729,1813,1835]" gridcol="1" gridrow="20" pageId="11" pageNumber="378">13.4 (1.3)</td>
<td id="58295DC3FF99C0CEFC4F2305BDA2F8AC" box="[1015,1170,1813,1835]" gridcol="2" gridrow="20" pageId="11" pageNumber="378">12.8 (1.1)</td>
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Phylogenetic trees from two inference methods (BI and ML) showed identical topologies for the most part, although some shallow nodes varied. Only the Bayesian Inference tree is shown, with node support values displayed from the two inference methods (
<figureCitation id="3DF32A78FF993F3AFD3323A4BBE1F84B" box="[651,721,1972,1997]" captionStart="FIGURE 8" captionStartId="15.[151,250,1951,1973]" captionTargetBox="[209,1397,205,1915]" captionTargetId="figure@15.[183,1401,195,1932]" captionTargetPageId="15" captionText="FIGURE 8. Phylogenetic relationships of G. motuoensis, G. yajiangensis, and 38 Garra species using Bayesian Inference of Cyt b sequences. Numbers on nodes before and after slash (/) represent posterior probabilities in BI tree and bootstrap values in ML tree, respectively (above 0.5/50 are shown). Garra motuoensis and G. yajiangensis are labeled with black rectangles." httpUri="https://zenodo.org/record/2615447/files/figure.png" pageId="11" pageNumber="378">Fig. 8</figureCitation>
). Topology of the phylogenetic tree indicated that all sampled individuals of
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<emphasis id="97BCEAEFFF993F3AFEF823C8B8D1F877" box="[320,481,2008,2032]" italics="true" pageId="11" pageNumber="378">G. motuoensis</emphasis>
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grouped into a monophyletic clade with high probabilities (0.99/100), which was sister to
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<emphasis id="97BCEAEFFF9E3F3DFF402488B821FF28" box="[248,273,152,175]" italics="true" pageId="12" pageNumber="379">G.</emphasis>
cf.
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A. Similarly
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, all sampled individuals of
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clustered into a monophyletic clade with high probabilities (1/99), which was the closest to
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<emphasis id="97BCEAEFFF9E3F3DFCA324ACBA03FF54" box="[795,819,188,211]" italics="true" pageId="12" pageNumber="379">G.</emphasis>
cf.
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B. Both
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and
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<emphasis id="97BCEAEFFF9E3F3DFB5724ACBCACFF53" box="[1263,1436,188,212]" italics="true" pageId="12" pageNumber="379">G. yajiangensis</emphasis>
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were independent from their sister taxon with high probabilities (1/100 and 0.9/66, respectively).
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<paragraph id="A5773F74FFA3FFFFDED1BA64C957FBAB" blockId="0.[136,1079,1048,1075]" box="[136,1079,1048,1075]" pageId="0" pageNumber="300">Thanks to Dr. Ian Turner (Royal Botanical Gardens, Kew) for pointing out the error.</paragraph>
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75
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<paragraph pageId="0" pageNumber="10084">Notes</paragraph>
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<bibRefCitation author="Anagnostidis, K" journalOrPublisher="Institute of Systematic Botany and Phytogeography of the Aristotle University of Thessaloniki, Thessaloniki" pageId="0" pageNumber="10084" title="Investigations of the Cyanophyceae in some thermal springs in Greece" year="1961">Anagnostidis 1961</bibRefCitation>
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<taxonomicName id="928F0F59176F89FBAD6528D762311103" ID-CoL="3F6FF" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Rubiaceae" genus="Galium" kingdom="Plantae" order="Gentianales" phylum="Tracheophyta" rank="species" species="saxatile" status="spec. nov.">Galium saxatile</taxonomicName>
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<paragraph id="016AF811E3B0BABE108D9754CC12FA24" pageNumber="106">8. Galium foliis senis obovatis obtusis, caule ramosissimo procumbente.</paragraph>
<paragraph id="F5FF4DC5C0C195EB36C47F860AFB57A6" pageNumber="106">
Galium caule ramosissimo, foliis quinis obverse ovatis.
<emphasis id="4C0BD517D587D786AF6CC85FE1FFCC53" italics="true">Hort. cliff. 34. Roy. lugdb. 257.</emphasis>
</paragraph>
<paragraph id="A594CD76D5875668C82279ADEF24EBE4" pageNumber="106">
Gallium saxatile supinum, molliore folio.
<emphasis id="7D2FD89331DE9BE544C2EF62270A1396" italics="true">Juss. act. 1714. p.492. t.15.</emphasis>
</paragraph>
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<paragraph id="7884E51BD83A73D9D6C2F6A3F606F639" pageNumber="106">
<emphasis id="AA17760C8010FE0CFEEB84E9A108E8D0" italics="true">Habitat in</emphasis>
Hispaniae
<emphasis id="52D2CE0D50B7FE707D07AB7DA6916683" italics="true">maritimis lapidosis.</emphasis>
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<taxonomicName authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Ranunculaceae" genus="Helleborus" kingdom="Plantae" order="Ranunculales" phylum="Tracheophyta" rank="species" species="hyemalis">
<emphasis bold="true" pageNumber="566">Helleborus hyemalis</emphasis>
Linnaeus
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,
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<emphasis italics="true" pageNumber="566">Species Plantarum</emphasis>
1
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: 557. 1753
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.
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<paragraph pageNumber="566">&quot;Habitat in Lombardia, Italia, Apenninis.&quot; RCN: 4106.</paragraph>
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<paragraph pageNumber="566">
<materialsCitation ID-GBIF-Occurrence="2611885550" collectionCode="BM" specimenCode="BM-000628873" typeStatus="lectotype">
<typeStatus>Lectotype</typeStatus>
(Sell in Jonsell &amp; Jarvis in
<emphasis italics="true" pageNumber="566">Nordic J. Bot.</emphasis>
14: 161. 1994): Herb. Clifford: 227,
<taxonomicName authorityName="Linnaeus" authorityYear="1754" class="Magnoliopsida" family="Ranunculaceae" genus="Helleborus" kingdom="Plantae" order="Ranunculales" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageNumber="566">Helleborus</emphasis>
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4 (BM-000628873)
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.
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<paragraph pageNumber="566">
Current name:
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<emphasis bold="true" italics="true" pageNumber="566">Eranthis hyemalis</emphasis>
(L.) Salisb.
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(
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<emphasis italics="true" pageNumber="566">Note:</emphasis>
Although Riedl (in Rechinger,
<emphasis italics="true" pageNumber="566">Fl. Iranica</emphasis>
171: 34. 1992) indicated unspecified material at LINN as type, this does not distinguish between sheets 718.1 and 718.2. As they are evidently not part of a single gathering, Art. 9.15 does not apply.
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<taxonomicName LSID="2D61DFD1-3A97-57E0-AD58-904BEFFD9632" authority="Wollaston, 1869" authorityName="Wollaston" authorityYear="1869" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachyphloeosoma" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">Trachyphloeosoma Wollaston, 1869</taxonomicName>
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<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Wollaston" authorityYear="1869" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachyphloeosoma" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">Trachyphloeosoma</taxonomicName>
Wollaston, 1869: 414 (original description);
<bibRefCitation author="Zimmerman, EC" journalOrPublisher="The Coleopterists Bulletin" pageId="0" pageNumber="93" pagination="27 - 31" publicationUrl="https://www.jstor.org/stable/3999054" refId="B16" refString="Zimmerman, EC, 1956. On Trachyphloeosoma and a new species from Hawaii (Coleoptera. Curculionidae). The Coleopterists Bulletin 10: 27 - 31, https://www.jstor.org/stable/3999054" title="On Trachyphloeosoma and a new species from Hawaii (Coleoptera. Curculionidae)." url="https://www.jstor.org/stable/3999054" volume="10" year="1956">Zimmerman 1956</bibRefCitation>
: 27 (review of genus);
<bibRefCitation author="Alonso-Zarazaga, MA" journalOrPublisher="Entomopraxis, Barcelona SCP" pageId="0" pageNumber="93" refId="B1" refString="Alonso-Zarazaga, MA, Lyal, CHC, 1999. A world catalogue of families and genera of Curculionoidea (Insecta: Coleoptera) (excluding Scolytidae and Platypodidae). Entomopraxis, Barcelona SCP" title="A world catalogue of families and genera of Curculionoidea (Insecta: Coleoptera) (excluding Scolytidae and Platypodidae)." year="1999">Alonso-Zarazaga and Lyal 1999</bibRefCitation>
: 183 (catalogue);
<bibRefCitation author="Borovec, R" journalOrPublisher="Klapalekiana" pageId="0" pageNumber="93" pagination="1 - 97" refId="B4" refString="Borovec, R, 2009. Revision of the Palaearctic supraspecific taxa of the tribe Trachyphloeini (Coleoptera: Curculionidae: Entiminae). Klapalekiana 45: 1 - 97" title="Revision of the Palaearctic supraspecific taxa of the tribe Trachyphloeini (Coleoptera: Curculionidae: Entiminae)." volume="45" year="2009">Borovec 2009</bibRefCitation>
: 52 (redescription of genus);
<bibRefCitation author="Borovec, R" journalOrPublisher="Studies and Reports, Taxonomical Series" pageId="0" pageNumber="93" pagination="1 - 39" refId="B6" refString="Borovec, R, 2014. Study on Trachyphloeini of Oriental region (Coleoptera: Curculionidae: Entiminae). Studies and Reports, Taxonomical Series 10: 1 - 39" title="Study on Trachyphloeini of Oriental region (Coleoptera: Curculionidae: Entiminae)." volume="10" year="2014">Borovec 2014</bibRefCitation>
: 11 (revision of genus);
<bibRefCitation author="Morimoto, K" journalOrPublisher="Fukuoka, Touka Shobo" pageId="0" pageNumber="93" refId="B8" refString="Morimoto, K, 2015. [new taxa] In: Morimoto K, Nakamura T, Kanno K. Curculionidae: Entiminae (Part 2) (Coleoptera). The Insects of Japan, Volume 4. Fukuoka, Touka Shobo" title="[new taxa] In: Morimoto K, Nakamura T, Kanno K. Curculionidae: Entiminae (Part 2) (Coleoptera). The Insects of Japan, Volume 4." year="2015">Morimoto 2015</bibRefCitation>
: 343 (review of Japanese species);
<bibRefCitation author="Alonso-Zarazaga, MA" journalOrPublisher="Monografias electronicas SEA" pageId="0" pageNumber="93" pagination="1 - 729" publicationUrl="http://sea-entomologia.org/PDF/MeSEA_8_Catalogue_Palaeartic_Curculionoidea.pdf" refId="B2" refString="Alonso-Zarazaga, MA, Barrios, H, Borovec, R, Bouchard, P, Caldara, R, Colonnelli, E, Gueltekin, L, Hlavac, P, Korotyaev, B, Lyal, CHC, Machado, A, Meregalli, M, Pierotti, H, Ren, L, Sanchez-Ruiz, M, Sforzi, A, Silfverberg, H, Skuhrovec, J, Tryzna, M, Velazquez de Castro, AJ, Yunakov, NN, 2017. Cooperative catalogue of palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8: 1 - 729, http://sea-entomologia.org/PDF/MeSEA_8_Catalogue_Palaeartic_Curculionoidea.pdf" title="Cooperative catalogue of palaearctic Coleoptera Curculionoidea." url="http://sea-entomologia.org/PDF/MeSEA_8_Catalogue_Palaeartic_Curculionoidea.pdf" volume="8" year="2017">Alonso-Zarazaga et al. 2017</bibRefCitation>
: 406 (catalogue).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="remarks">
<paragraph pageId="0" pageNumber="93">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="93">
This genus was described by Wollaston based on material from the island of St. Helena. Additional species were described later, and the present number of valid species is five. The genus was redescribed and compared to all other Palaearctic
<taxonomicName lsidName="" pageId="0" pageNumber="93" rank="tribe" tribe="Trachyphloeini">Trachyphloeini</taxonomicName>
by
<bibRefCitation author="Borovec, R" journalOrPublisher="Klapalekiana" pageId="0" pageNumber="93" pagination="1 - 97" refId="B4" refString="Borovec, R, 2009. Revision of the Palaearctic supraspecific taxa of the tribe Trachyphloeini (Coleoptera: Curculionidae: Entiminae). Klapalekiana 45: 1 - 97" title="Revision of the Palaearctic supraspecific taxa of the tribe Trachyphloeini (Coleoptera: Curculionidae: Entiminae)." volume="45" year="2009">Borovec (2009)</bibRefCitation>
, then subsequently revised by
<bibRefCitation author="Borovec, R" journalOrPublisher="Studies and Reports, Taxonomical Series" pageId="0" pageNumber="93" pagination="1 - 39" refId="B6" refString="Borovec, R, 2014. Study on Trachyphloeini of Oriental region (Coleoptera: Curculionidae: Entiminae). Studies and Reports, Taxonomical Series 10: 1 - 39" title="Study on Trachyphloeini of Oriental region (Coleoptera: Curculionidae: Entiminae)." volume="10" year="2014">Borovec (2014)</bibRefCitation>
, based on material from China, Vietnam, Japan, Korea, and the Moluccas.
<bibRefCitation author="Morimoto, K" journalOrPublisher="Fukuoka, Touka Shobo" pageId="0" pageNumber="93" refId="B8" refString="Morimoto, K, 2015. [new taxa] In: Morimoto K, Nakamura T, Kanno K. Curculionidae: Entiminae (Part 2) (Coleoptera). The Insects of Japan, Volume 4. Fukuoka, Touka Shobo" title="[new taxa] In: Morimoto K, Nakamura T, Kanno K. Curculionidae: Entiminae (Part 2) (Coleoptera). The Insects of Japan, Volume 4." year="2015">Morimoto (2015)</bibRefCitation>
, in his monography of Japanese
<taxonomicName authorityName="Schoenherr" authorityYear="1823" lsidName="" pageId="0" pageNumber="93" rank="subFamily" subFamily="Entiminae">Entiminae</taxonomicName>
, surveyed the Japanese species of the genus. China is the most northwestern part of the range of the genus, and
<taxonomicName authorityName="Wollaston" authorityYear="1869" class="Insecta" family="Curculionidae" genus="Trachyphloeosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachyphloeosoma" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="93">Trachyphloeosoma</emphasis>
</taxonomicName>
was first recorded from this country only in 2009 by Borovec, based on one male from Yunnan, and subsequently by
<bibRefCitation author="Borovec, R" journalOrPublisher="Studies and Reports, Taxonomical Series" pageId="0" pageNumber="93" pagination="1 - 39" refId="B6" refString="Borovec, R, 2014. Study on Trachyphloeini of Oriental region (Coleoptera: Curculionidae: Entiminae). Studies and Reports, Taxonomical Series 10: 1 - 39" title="Study on Trachyphloeini of Oriental region (Coleoptera: Curculionidae: Entiminae)." volume="10" year="2014">Borovec (2014)</bibRefCitation>
based on several additional specimens of the same species. Following examination of some newly sifted material from China, and comparing this with
<bibRefCitation author="Morimoto, K" journalOrPublisher="Fukuoka, Touka Shobo" pageId="0" pageNumber="93" refId="B8" refString="Morimoto, K, 2015. [new taxa] In: Morimoto K, Nakamura T, Kanno K. Curculionidae: Entiminae (Part 2) (Coleoptera). The Insects of Japan, Volume 4. Fukuoka, Touka Shobo" title="[new taxa] In: Morimoto K, Nakamura T, Kanno K. Curculionidae: Entiminae (Part 2) (Coleoptera). The Insects of Japan, Volume 4." year="2015">
<normalizedToken originalValue="Morimoto’s">Morimoto's</normalizedToken>
(2015)
</bibRefCitation>
review of the genus from the Japanese islands, we can state that the species collected in China are new to science. We can thus correct the name of the species so far recorded from China. This newly collected material from China was erroneously identified as
<taxonomicName lsidName="T. advena" pageId="0" pageNumber="93" rank="species" species="advena">
<emphasis italics="true" pageId="0" pageNumber="93">T. advena</emphasis>
</taxonomicName>
Zimmerman, 1956 and was listed under this name in the Palaearctic catalogue (
<bibRefCitation author="Alonso-Zarazaga, MA" journalOrPublisher="Monografias electronicas SEA" pageId="0" pageNumber="93" pagination="1 - 729" publicationUrl="http://sea-entomologia.org/PDF/MeSEA_8_Catalogue_Palaeartic_Curculionoidea.pdf" refId="B2" refString="Alonso-Zarazaga, MA, Barrios, H, Borovec, R, Bouchard, P, Caldara, R, Colonnelli, E, Gueltekin, L, Hlavac, P, Korotyaev, B, Lyal, CHC, Machado, A, Meregalli, M, Pierotti, H, Ren, L, Sanchez-Ruiz, M, Sforzi, A, Silfverberg, H, Skuhrovec, J, Tryzna, M, Velazquez de Castro, AJ, Yunakov, NN, 2017. Cooperative catalogue of palaearctic Coleoptera Curculionoidea. Monografias electronicas SEA 8: 1 - 729, http://sea-entomologia.org/PDF/MeSEA_8_Catalogue_Palaeartic_Curculionoidea.pdf" title="Cooperative catalogue of palaearctic Coleoptera Curculionoidea." url="http://sea-entomologia.org/PDF/MeSEA_8_Catalogue_Palaeartic_Curculionoidea.pdf" volume="8" year="2017">Alonso-Zarazaga et al. 2017</bibRefCitation>
). After dissection and thorough examination, we are able to recognise the specimens as distinct from
<taxonomicName lsidName="T. advena" pageId="0" pageNumber="93" rank="species" species="advena">
<emphasis italics="true" pageId="0" pageNumber="93">T. advena</emphasis>
</taxonomicName>
and belonging to three different species, which are described below.
</paragraph>
<caption doi="10.3897/zookeys.974.56059.figure25" httpUri="https://binary.pensoft.net/fig/460958" pageId="0" pageNumber="93" start="Figure 25" startId="F4">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figure 25.</emphasis>
Head of
<taxonomicName authorityName="Ren &amp; Borovec &amp; Zhang" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Rhinodontodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinodontodes alashanensis" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="alashanensis">
<emphasis italics="true" pageId="0" pageNumber="93">Rhinodontodes alashanensis</emphasis>
</taxonomicName>
, sp. nov., dorsal view, arrows indicate the epistome.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>