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<document id="D4EBBBE99E8DB91429CF6B8D78FA2459" ID-CLB-Dataset="28566" ID-DOI="http://dx.doi.org/10.3897/zookeys.822.29835" ID-GBIF-Dataset="acba2a19-0345-444f-ae2c-49273bbccfeb" ID-PMC="PMC6374355" ID-Pensoft-Pub="1313-2970-822-141" ID-PubMed="30774504" ID-ZooBank="6FEB97EC0DB54270BAD796379C87C2EB" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-822-141" ModsDocOrigin="ZooKeys 822" ModsDocTitle="Revision of the ant-eating spider genus Mallinus Simon, 1893 (Araneae, Zodariidae)" checkinTime="1549492742523" checkinUser="pensoft" docAuthor="Haddad, Charles R., Henrard, Arnaud &amp; Jocque, Rudy" docDate="2019" docId="90D72C44E760655FFF843FB5DCAF59AA" docLanguage="en" docName="ZooKeys 822: 141-158" docOrigin="ZooKeys 822" docSource="http://dx.doi.org/10.3897/zookeys.822.29835" docTitle="Mallinus nitidiventris Simon 1893" docType="treatment" docVersion="6" lastPageNumber="150" masterDocId="FFBDA823DE2EE60EFFE6FF5432414715" masterDocTitle="Revision of the ant-eating spider genus Mallinus Simon, 1893 (Araneae, Zodariidae)" masterLastPageNumber="158" masterPageNumber="141" pageNumber="147" updateTime="1701374799160" updateUser="ExternalLinkService">
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<mods:title id="C7589F1BACD0CE698960A2A99E312CAB">Revision of the ant-eating spider genus Mallinus Simon, 1893 (Araneae, Zodariidae)</mods:title>
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<mods:namePart id="B9CA6901208B4D496F1E374BDDDFF4EB">Haddad, Charles R.</mods:namePart>
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<mods:namePart id="EF4DA1EC49139F9C8320316F7BF0BE90">Jocque, Rudy</mods:namePart>
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<treatment id="90D72C44E760655FFF843FB5DCAF59AA" ID-GBIF-Taxon="154127141" LSID="urn:lsid:plazi:treatment:90D72C44E760655FFF843FB5DCAF59AA" httpUri="http://treatment.plazi.org/id/90D72C44E760655FFF843FB5DCAF59AA" lastPageId="9" lastPageNumber="150" pageId="6" pageNumber="147">
<subSubSection id="01D49B494803B9EDC3549FA143D11B6D" pageId="6" pageNumber="147" type="nomenclature">
<paragraph id="40BC4CFD585A7B6104C1A5C26A3522AF" pageId="6" pageNumber="147">
<taxonomicName id="2752BD1B84B194C04FE4822BF40E0CA7" ID-CoL="3XNPY" authority="Simon, 1893" authorityName="Simon" authorityYear="1893" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris Simon, 1893</taxonomicName>
Figs 1-4, 5-19, 20-34, 35-46, 47-49, 50-53, 54-57
</paragraph>
</subSubSection>
<subSubSection id="EEB9CA1D9D8CF3A90B80F1DC6ABC4036" pageId="6" pageNumber="147" type="reference_group">
<paragraph id="318B066F2AEB2D1CCC925DC6802E3967" pageId="6" pageNumber="147">
<taxonomicName id="DC74CF4DE7A7B82F97B2104EBF3093A2" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="6" pageNumber="147" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
Simon, 1893: 436, figs 409-410 (juv.);
<bibRefCitation id="62F3C218EFA332D1331198808925C4C6" author="Jocque, R" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="15" pageNumber="156" pagination="1 - 160" title="A generic revision of the spider family Zodariidae (Araneae)." url="http://digitallibrary.amnh.org/handle/2246/894" volume="201" year="1991">
<normalizedToken id="52EFDFE3C72D29A09BA422BD3E2B0BAB" originalValue="Jocqué">Jocque</normalizedToken>
1991
</bibRefCitation>
: 136, figs 334-335 (juv.).
</paragraph>
</subSubSection>
<subSubSection id="2993AB02F381B108534C5C58308A2BDF" pageId="6" pageNumber="147" type="materials_examined">
<paragraph id="1EDFCD5A6FD7C876FDF322910CF329B4" pageId="6" pageNumber="147">Type material.</paragraph>
<paragraph id="D00A5CCFA7AF787704EBCAD7DB5FACE4" pageId="6" pageNumber="147">
Subadult ♂ lectotype and subadult ♀ paralectotype: SOUTH AFRICA: Western Cape: Matjiesfontein,
<geoCoordinate id="43E67677D9528965176A89F61E3A3410" direction="south" orientation="latitude" precision="925" value="-33.25">33°15'S</geoCoordinate>
,
<geoCoordinate id="7EF0D10F67EF6C9427E44E7FE19B3AB8" direction="east" orientation="longitude" precision="925" value="20.666666">20°40'E</geoCoordinate>
, MNHN AR 3280 - examined by
<bibRefCitation id="2EA4EDCF141E5FCDFBC67B250613C0FB" author="Jocque, R" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="15" pageNumber="156" pagination="1 - 160" title="A generic revision of the spider family Zodariidae (Araneae)." url="http://digitallibrary.amnh.org/handle/2246/894" volume="201" year="1991">
<normalizedToken id="4C0D01B39468656A65412602BF896D11" originalValue="Jocqué">Jocque</normalizedToken>
(1991)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7577D4C259C6854AC774D88C06660EA4" pageId="6" pageNumber="147" type="materials_examined">
<paragraph id="02324360C3BBC5E929BC6D127632F139" pageId="6" pageNumber="147">Other material examined.</paragraph>
<paragraph id="90C994A73E035D289EA1A76C669E930D" pageId="6" pageNumber="147">
SOUTH AFRICA: Eastern Cape: Aberdeen district, Farm Bokvlei,
<geoCoordinate id="D56EEC93827DF7EB808FCEFB76F816CB" direction="south" orientation="latitude" precision="92" value="-32.43">32°25.8'S</geoCoordinate>
,
<geoCoordinate id="1444BDE6B108CF1D1537AE081E3F116B" direction="east" orientation="longitude" precision="92" value="23.35">23°21.0'E</geoCoordinate>
, 14.XII.2007, leg. D.H. Jacobs (pitfall traps, karoo), 2♂ (NCA 2008/4662); Aberdeen district, Farm De Pannen,
<geoCoordinate id="2BBF8252DCB492C7D64A2580CB93965B" direction="south" orientation="latitude" precision="92" value="-32.69">32°41.4'S</geoCoordinate>
,
<geoCoordinate id="816F02DEFF3157E1E22EB6CCF3D69F15" direction="east" orientation="longitude" precision="92" value="23.43">23°25.8'E</geoCoordinate>
, 14.XII.2007, leg. D.H. Jacobs (pitfall traps), 1 imm. 4♂ (NCA 2011/604); Aberdeen district, Farm Juriesfontein,
<geoCoordinate id="B63D88EED5A011BDFD4EE313D72D4EF1" direction="south" orientation="latitude" precision="92" value="-32.53">32°31.8'S</geoCoordinate>
,
<geoCoordinate id="55B281395BF2BB9865D8883BAE732D68" direction="east" orientation="longitude" precision="92" value="23.43">23°25.8'E</geoCoordinate>
, 11.XII.2007, leg. D.H. Jacobs (pitfall traps), 1♂ (NCA 2008/4665); Aberdeen district, Farm Nuwejaarsfontein,
<geoCoordinate id="236E0A1515B54EC65836BFC060A6AB63" direction="south" orientation="latitude" precision="92" value="-32.95">32°57.0'S</geoCoordinate>
,
<geoCoordinate id="FBD8EA196958D26AD9335C59A540E0A0" direction="east" orientation="longitude" precision="92" value="24.39">24°23.4'E</geoCoordinate>
, 14.XII.2007, leg. D.H. Jacobs (pitfall traps, karoo), 2 imm. 7♂ 1♀ (NCA 2008/2612). Free State: Luckhoff district, Farm Bankfontein,
<geoCoordinate id="7D02AC0792E111C89AD76375B88AAE2D" direction="south" orientation="latitude" precision="1" value="-30.083">30°04.980'S</geoCoordinate>
,
<geoCoordinate id="ECBCDD1EF31AC62710FA1F9006E3B2F6" direction="east" orientation="longitude" precision="1" value="24.902834">24°54.170'E</geoCoordinate>
, 22.I.2015, leg. C. Haddad (base of grass tussocks, wetland margin), 1♂ (NCA 2015/1655); Same locality,
<geoCoordinate id="10E1AAABC02F414D0A86A85573F7BF66" direction="south" orientation="latitude" precision="1" value="-30.073683">30°04.421'S</geoCoordinate>
,
<geoCoordinate id="EED45BFF5CC5E584646FFD9422EDB51F" direction="east" orientation="longitude" precision="1" value="24.883617">24°53.017'E</geoCoordinate>
, 6-8.IV.2015, leg. C. Haddad (hand collecting, Nama Karoo veld), 1♂ 1♀ (NCA 2015/1818), 2♂ 1♀ (S.E.M. preparations); Same locality,
<geoCoordinate id="A73365ABDA7AF840AB611215E488FFBE" direction="south" orientation="latitude" precision="1" value="-30.073683">30°04.421'S</geoCoordinate>
,
<geoCoordinate id="E2BB13206CE89C51C1206333B03F13C2" direction="east" orientation="longitude" precision="1" value="24.883966">24°53.038'E</geoCoordinate>
, 26.III.2017, leg. C. Haddad &amp; R. Booysen (hand collecting, Nama Karoo veld), 3 imm. 5♂ (NCA 2017/1447); Same locality,
<geoCoordinate id="8568038FD20F00A8DA9BFA56CD43A074" direction="south" orientation="latitude" precision="1" value="-30.0829">30°04.974'S</geoCoordinate>
,
<geoCoordinate id="9CBCF732F53B10B76BB7F537907B61F5" direction="east" orientation="longitude" precision="1" value="24.90495">24°54.297'E</geoCoordinate>
, 2-6.IV.2015, leg. University of the Free State students (pitfall traps, Nama Karoo veld), 1♂ (NCA 2015/2398); Same locality, Research camp,
<geoCoordinate id="BE46A9D6515B335FD5C7F1FFD47BC052" direction="south" orientation="latitude" precision="1" value="-30.073683">30°04.421'S</geoCoordinate>
,
<geoCoordinate id="4B58B7C9D7CBE571EDC0FC399064A8B5" direction="east" orientation="longitude" precision="1" value="24.88355">24°53.013'E</geoCoordinate>
, 1185 m a.s.l., 24-26.XI.2015, leg. C. Haddad &amp; R. Booysen (night collecting), 1♂ (NCA 2015/2231). Northern Cape: Pofadder,
<geoCoordinate id="4EEFCA0F60943CCCAF1DD79632C24DA6" direction="south" orientation="latitude" precision="925" value="-29.366667">29°22'S</geoCoordinate>
,
<geoCoordinate id="EAD586F6E921A104B076C583A553BB61" direction="east" orientation="longitude" precision="925" value="19.116667">19°07'E</geoCoordinate>
, 15.X.2006, leg. L. Spangenberg (pitfall traps), 1♀ (MRAC 222264); Tswalu Game Reserve, man-made dam,
<geoCoordinate id="88F003F5266446A22ECC229A1ED06D40" direction="south" orientation="latitude" precision="925" value="-27.25">27°15'S</geoCoordinate>
,
<geoCoordinate id="45C8B5E42F7239FAC48B78E127995136" direction="east" orientation="longitude" precision="925" value="22.45">22°27'E</geoCoordinate>
, 1176 m a.s.l., 6.II.2005, leg. R.
<normalizedToken id="49EC7F4D133F35D92A058374F63FD967" originalValue="Jocqué">Jocque</normalizedToken>
(savanna shrubland, by hand), 1♂ (MRAC 216253). Western Cape: Beaufort West district, Farm Eerste Water,
<geoCoordinate id="49441C8845920692213CE3DF766AE665" direction="south" orientation="latitude" precision="92" value="-32.69">32°41.4'S</geoCoordinate>
,
<geoCoordinate id="0298A66FFEC2481D56F0E38389EC63C3" direction="east" orientation="longitude" precision="92" value="22.96">22°57.6'E</geoCoordinate>
, 6.XII.2007, leg. D.H. Jacobs (pitfall traps, karoo), 1 imm. (NCA 2008/2610); Prince Albert, Tierberg,
<geoCoordinate id="7C658A2657ACB20D925CFAE5245113D0" direction="south" orientation="latitude" precision="925" value="-33.216667">33°13'S</geoCoordinate>
,
<geoCoordinate id="3E1291BF72125AB252A6B7D30DD15269" direction="east" orientation="longitude" precision="925" value="22.033333">22°02'E</geoCoordinate>
, 23.IX.1989, leg. R. Dean (old lands), 1♂ (NCA 91/1299), 1♀ (NCA 91/1300).
</paragraph>
</subSubSection>
<subSubSection id="A7C9000602AA5E60569B145FC9228DE3" lastPageId="9" lastPageNumber="150" pageId="6" pageNumber="147" type="unconfirmed record">
<paragraph id="F04E782E3A2CFB385929599D86AE7410" pageId="6" pageNumber="147">Unconfirmed record.</paragraph>
<paragraph id="1C6059E7190D476B3F2A0535EB6C2F4F" pageId="6" pageNumber="147">
SOUTH AFRICA: Northern Cape: Goepag Nature Reserve,
<geoCoordinate id="6F5315A904B056F775ED0035C329D9A6" direction="south" orientation="latitude" precision="1" value="-29.6651">29°39.906'S</geoCoordinate>
,
<geoCoordinate id="32EAA3E3C18AF3ADAF7FC24DF5054AD2" direction="east" orientation="longitude" precision="1" value="17.9973">17°59.838'E</geoCoordinate>
, 14-16.VII.2017, leg. R. Booysen, Z. Mbo &amp; R. Christiaan (pitfall traps, Nama Karoo veld), 1 imm. (NCA 2017/1189).
</paragraph>
<paragraph id="128054B92601F6CAE0AA11E1580E53AB" pageId="6" pageNumber="147">
Female (Bankfontein, NCA 2015/1818). Measurements: CL 1.32, CW 0.89, AL 1.39, AW 1.36, AH 1.65, TL 2.60 (2.13-2.70), SL 0.62, SW 0.63,
<normalizedToken id="DCDB66AE779A7D5C546A8B780859AB56" originalValue="AME–AME">AME-AME</normalizedToken>
0.06,
<normalizedToken id="6C39F59D374CBEDD3A8DC7CAC7FD52E9" originalValue="AME–ALE">AME-ALE</normalizedToken>
0.03,
<normalizedToken id="17AE613FB5986E4DAF05945B87F39135" originalValue="ALE–ALE">ALE-ALE</normalizedToken>
0.25,
<normalizedToken id="1F464B11EF3621FA55A80D6C48D39507" originalValue="PME–PME">PME-PME</normalizedToken>
0.09,
<normalizedToken id="877387669BB5A8B30C7EBCC3ACD57368" originalValue="PME–PLE">PME-PLE</normalizedToken>
0.09,
<normalizedToken id="E44E12BA0892CF79DEFAB88A4088E56F" originalValue="PLE–PLE">PLE-PLE</normalizedToken>
0.31, MOQAW 0.21, MOQPW 0.22, MOQL 0.25.
</paragraph>
<paragraph id="36BE8F9042D121DDF301F92F547451FD" pageId="6" pageNumber="147">Length of leg segments: I 0.70 + 0.29 + 0.54 + 0.67 + 0.41 = 2.61; II 0.71 + 0.30 + 0.53 + 0.73 + 0.45 = 2.72; III 0.71 + 0.32 + 0.53 + 0.78 + 0.45 = 2.79; IV 0.92 + 0.32 + 0.68 + 1.00 + 0.48 = 3.40.</paragraph>
<paragraph id="EBB358005D2D07A163B8DF6B81771C24" lastPageId="7" lastPageNumber="148" pageId="6" pageNumber="147">
Colour: carapace orange-brown, with faint black mottling and striae (Figs 1, 2); chelicerae orange; endites yellow, slightly darker retrolaterally at midpoint; labium orange, cream distally; sternum orange, cream along anterior margin; leg femora orange-brown; patellae yellow proximally, orange brown distally; tibiae and metatarsi light brown proximally, yellow-brown distally; tarsi yellow; palps yellow-brown; abdomen black dorsally, grey ventrally along midline, with large white patches laterally, fused
<pageBreakToken id="705C531A5E86922DA7439DE9DEA42884" pageId="7" pageNumber="148" start="start">narrowly</pageBreakToken>
in ring around anterior of abdomen; spinnerets creamy-yellow. Eyes: AME diameter 1.1 times ALE diameter; AME separated by distance equal to 0.76 times their diameter; AME separated from ALE by 0.4 times AME diameter; clypeus height 4.2 times AME diameter at AME, 4.16 times ALE diameter at ALE; PME and PLE equal in diameter; PME separated by distance equal to 1.15 times their diameter; PME separated from PLE by distance slightly less than 1.21 times PME diameter; CW:PERW = 2.06:1. Legs spineless, covered in short erect setae and incised setae. Abdomen slightly longer than carapace, higher than long or broad, with shiny scutum covering most of dorsum (Figs 1, 2); dorsum sparsely covered in short straight setae, denser on posterior slope and venter. Epigyne as in genus description (Figs 43, 47, 50, 51). Other characters as in genus description.
</paragraph>
<caption id="82C9BA0F86579854AA60FC19768ACFFE" pageId="7" pageNumber="148">
<paragraph id="B3B7AB93FB14A8F200AC957440492C66" pageId="7" pageNumber="148">
Figures 1-4. Somatic morphology of
<taxonomicName id="155FEDA5B58F77BF20E8664AB608B706" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
female (1, 3) and male (2, 4) from Bankfontein, Free State (NCA 2015/1818) 1, 2 habitus, dorsal view 3, 4 same, lateral view. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="AFC821F7BAF735FCD5B3CCEE29695836" pageId="7" pageNumber="148">
Male (Bankfontein, NCA 2015/1818). Measurements: CL 1.33, CW 0.88, AL 0.98, AW 0.94, AH 0.98, TL 2.34 (2.23-2.72), SL 0.60, SW 0.59,
<normalizedToken id="D0C66F02BEB80EA6CBBC952C53835E6A" originalValue="AME–AME">AME-AME</normalizedToken>
0.06,
<normalizedToken id="3BC22F2DE4048EF64850AA96B3C2407B" originalValue="AME–ALE">AME-ALE</normalizedToken>
0.03,
<normalizedToken id="18F45C8979EF9EC5B78D9F315C64BEEE" originalValue="ALE–ALE">ALE-ALE</normalizedToken>
0.25,
<normalizedToken id="282F74B921D04E0C16BC77F6B1E18080" originalValue="PME–PME">PME-PME</normalizedToken>
0.08,
<normalizedToken id="B16C0C00653FAEF226E9CD793A8CF325" originalValue="PME–PLE">PME-PLE</normalizedToken>
0.08,
<normalizedToken id="C5AD4130C0CC6423E9BF9D686AE74681" originalValue="PLE–PLE">PLE-PLE</normalizedToken>
0.29, MOQAW 0.21, MOQPW 0.19, MOQL 0.22.
</paragraph>
<paragraph id="E5CD870B4FDC62D8A18829459E0B4286" pageId="8" pageNumber="149">
<pageBreakToken id="2A5421C8BA5D703FAB71FA8F12ECA935" pageId="8" pageNumber="149" start="start">Length</pageBreakToken>
of leg segments: I 0.79 + 0.29 + 0.65 + 0.81 + 0.46 = 3.00; II 0.83 + 0.30 + 0.63 + 0.84 + 0.46 = 3.06; III 0.81 + 0.33 + 0.57 + 0.90 + 0.47 = 3.08; IV 1.03 + 0.33 + 0.75 + 1.15 + 0.53 = 3.79.
</paragraph>
<paragraph id="78C6A0D5EFFF1B9B0464D2F2B81AEDAD" lastPageId="9" lastPageNumber="150" pageId="8" pageNumber="149">
Morphology and colouration similar to female (Figs 3, 4), except for the following: AME diameter equals 1.16 times ALE diameter; AME separated by distance 0.67 times their diameter; AME separated from ALE by distance 0.38 times AME diameter; clypeus height 3.85 times AME diameter at AME, 4.0 times ALE diameter at ALE; PME diameter equals 0.89 times PLE diameter; PME separated by distance 1.25 times their diameter; PME separated from PLE by distance 1.25 times PME diameter;
<pageBreakToken id="F35F738A3110B1672A28C219CCCA6CA7" pageId="9" pageNumber="150" start="start">CW</pageBreakToken>
:PERW = 2.17:1. Abdomen relatively smaller than female (Figs 3, 4), shorter than carapace, as high as long, slightly longer than broad, with conspicuous circumferential folds. Palp as in genus description (Figs 44, 48, 49, 52, 53).
</paragraph>
<caption id="DF96A24A7BA15AB1A53C2C9B11787154" pageId="9" pageNumber="150">
<paragraph id="DAC6BC3A0606012531758F343BF47D91" pageId="9" pageNumber="150">
Figures 5-19. Scanning electron microscope photographs of
<taxonomicName id="BBFCD74285BB96B2864BCADED26A5438" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
male (5-10, 12-19) and female (11) 5 carapace, dorsal view 6 detail of carapace integument 7 same, detail of carapace setae 8 same, detail of carapace pore 9 fovea 10 eye region, dorsal view 11 endites, labium and anterior end of sternum 12 distal end of chelicera, arrow indicating posterior groove of fang 13 distal end of endite, black arrow indicating distal endite scale, white arrow indicating prolateral field of flattened denticles 14 prolateral margin of endite, showing detail of flattened denticles 15 femur I, dorsal view, incised setae and femoral organ (black arrow) 16-19 femoral organ on legs
<normalizedToken id="D823B882213BD159C36A7E8AFB1C7FD5" originalValue="I–IV">I-IV</normalizedToken>
, respectively.
</paragraph>
</caption>
<caption id="B863C5707114B9D63C3AC5D63EE9A892" pageId="9" pageNumber="150">
<paragraph id="42474ECCDBC5FC23BD8EDC94A548FA2A" pageId="9" pageNumber="150">
Figures 20-34. Scanning electron microscope photographs of
<taxonomicName id="99BA36574A97CF033FC4A6D99A4F535A" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
female (20, 21, 28, 29, 33, 34) and male (22-27, 30-32) 20 patella IV, retrolateral view 21 same, detail of lyriform organ 22 metatarsus I, incised setae and dorsal fields of short chemosensory setae (black arrows) 23 same, detail of short chemosensory setae 24 metatarsus I, trichobothrium 25 metatarsus III, arrow indicating metatarsal stopper 26 metatarsus III, arrows indicating thickened ventral terminal setae 27 same, metatarsus IV 28 tarsus I, paired claws in distal view 29 same, prolateral distal view, black arrow indicating chemosensory seta, white arrow indicating subdistal suture of tarsus 30 same, trichobothrium 31 same, chemosensory seta 32 same, tarsal organ 33 palpal tarsus and claw 34 palpal patella, lyriform organ.
</paragraph>
</caption>
<caption id="4B2C5073AB5C18314A64DD81AC3D7335" pageId="9" pageNumber="150">
<paragraph id="E7F6EDC74B5091C46AF244848A4988D1" pageId="9" pageNumber="150">
Figures 35-46. Scanning electron microscope photographs of
<taxonomicName id="460D5C6B1DC44D5A3A246723B3C38E60" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
male (35, 41, 42, 44-46) and female (36-40, 43) 35 abdomen, ventrolateral view 36 petiole, ventral view 37 ventral abdominal setae 38, 41 spinnerets, ventral view, arrow in 38 indicating sclerite in front of tracheal spiracle 39 detail of setae on sclerite in front of tracheal spiracle 40, 42 detail of anterior lateral spinnerets 43 epigyne, ventral view 44 palp, ventral view 45 detail of palpal claw and thickened distal prolateral seta 46 detail of palpal claw.
</paragraph>
</caption>
<caption id="B72D7354C754D2851C1AB2919AAC0BD6" pageId="9" pageNumber="150">
<paragraph id="46A82344585F5DDB2B2E7FBB411BAD46" pageId="9" pageNumber="150">
Figures 47-49.
<taxonomicName id="F28C2C267486F0FF9D651E0CD9B62C18" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
, female (47) and male (48, 49) genitalic morphology (NCA 2008/2612) 47 epigyne, ventral view 48 palp, retrolateral view 49 same, ventral view. Abbreviations:
<normalizedToken id="4EB98EF91C5AB3180FFF04EAABCF07BF" originalValue="C–conductor">C-conductor</normalizedToken>
;
<normalizedToken id="AC1B7C44DDF3D803D5323AA8A751C5AC" originalValue="E–embolus">E-embolus</normalizedToken>
;
<normalizedToken id="92BEB29BD1E2366364E0FD8FB0E47BF0" originalValue="MA–median">MA-median</normalizedToken>
apophysis. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B61EC0583640265959E20F69BF19A45E" pageId="9" pageNumber="150" type="variation">
<paragraph id="E160F01B2CD1A98C9D07AAC4644887B3" pageId="9" pageNumber="150">Variation.</paragraph>
<paragraph id="5EA56E7E5F318B64323A43B7B1E6439F" pageId="9" pageNumber="150">
Populations from the south-western parts of the
<normalizedToken id="B6382298CB5BDF10315FAEF518F109A2" originalValue="species’">species'</normalizedToken>
range (including the type locality) have a clearly darker carapace and legs, which are wine-red in colour (Figs 54-57). It is plausible that populations in this part of its range may associate with a darker species of model ant, affecting their colouration.
</paragraph>
<caption id="AA152AF0E0278B00323963B6EB67B648" pageId="9" pageNumber="150">
<paragraph id="966181467084491B45308362BE36F745" pageId="9" pageNumber="150">
Figures 50-53.
<taxonomicName id="E241A342887585F9683CE5A840A455B7" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
, female (50, 51) and male (52, 53) genitalic morphology (NCA 2008/2612) 50 epigyne, ventral view 51 same, digested, dorsal view 52 palp, ventral view 53 same, retrolateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="88573C4B16F051DBC1A14C639EB85479" pageId="9" pageNumber="150">
<paragraph id="967F93031EB06F7E25E2739C9D13F4BA" pageId="9" pageNumber="150">
Figures 54-57. Somatic morphology of
<taxonomicName id="C74F5183DC1D80A398CF0D6FC9CA762F" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
female (54, 55) and male (56, 57) from Beaufort West, Western Cape (NCA 91/1300 and NCA 2008/2612, respectively) 54, 56 habitus, dorsal view 55, 57 same, ventral view. Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption id="49EAF37A11F367DB40000D19AE3DEF3F" pageId="9" pageNumber="150">
<paragraph id="7CB816C4F52146077730E80CBAAF3E27" pageId="9" pageNumber="150">
Figures 58-64. Somatic morphology of &quot;
<taxonomicName id="0F306C63D10507F49DE281132D1B3F8E" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">Mallinus</taxonomicName>
&quot; sp. male from north-western Namibia (SMN 40843) 58 habitus, dorsal view 59 same, ventral view 60 carapace, lateral view 61 same, anterior view 62 abdomen, lateral view 63 palp, ventral view 64 same, retrolateral view. Scale bars: 1.0 mm (58, 59); 0.5 mm (60-62); 0.25 mm (63, 64).
</paragraph>
</caption>
<caption id="A41312B39522B095FD04D9759C61CAF5" pageId="9" pageNumber="150">
<paragraph id="7F16AE50F5B2303104B1B259CE3652BE" pageId="9" pageNumber="150">
Figures 65-66. &quot;
<taxonomicName id="FF7101205793BED7C24A6DF6BE9599E8" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">Mallinus</taxonomicName>
&quot; sp., male genitalic morphology (SMN 40843) 65 palp, retrolateral view 66 same, ventral view. Abbreviations:
<normalizedToken id="8BC4A9E880EC65D3B0644214C5913DB3" originalValue="E–embolus">E-embolus</normalizedToken>
;
<normalizedToken id="FFABBED7802FC42A97274A3BD111BD4C" originalValue="MA–median">MA-median</normalizedToken>
apophysis. Scale bars: 0.25 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9989CBEAB0D4323659040A1FB09D17F9" pageId="9" pageNumber="150" type="distribution">
<paragraph id="4569E6338C26970FCE429E0C3E43DE8D" pageId="9" pageNumber="150">Distribution.</paragraph>
<paragraph id="D10955F04437794D6F3B8BAA60ABFDCC" pageId="9" pageNumber="150">Widespread in the western half of South Africa, known from the Eastern Cape, Western Cape, Northern Cape and Free State Provinces (Fig. 67).</paragraph>
<caption id="93950E4EA3B15C6AB305835542F6842C" pageId="9" pageNumber="150">
<paragraph id="9D183ACE4894238E1EFCB5DCC228C016" pageId="9" pageNumber="150">
Figure 67. Distribution of
<taxonomicName id="D342CBBBE5153CA3B80CCB0390262BAC" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
(type locality, red circle; other localities, yellow circles) and &quot;
<taxonomicName id="1110AEBF6020F0EF4210F40DE3959604" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">Mallinus</taxonomicName>
&quot; sp. (red square) in southern Africa.? indicates immature specimen requiring confirmation.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="19F6F19A26879CBA6F215E6ECD9C6C24" pageId="9" pageNumber="150" type="biology_ecology">
<paragraph id="4E267298DEA63FCCCD3AF62FFF6662EB" pageId="9" pageNumber="150">Habitat and biology.</paragraph>
<paragraph id="F87CDE0EA0AA637576CA37EC501224A7" pageId="9" pageNumber="150">
<taxonomicName id="DA0EE4AFC7A1E9E16880D28F974B4B2B" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
is widespread in the semi-arid and arid western half of South Africa, with records in the Nama Karoo and Succulent Karoo biomes, extending into the arid savannas of the southern Kalahari Desert. Specimens collected at Bankfontein in the western Free State Province were all found in Nama Karoo scrubland, either along a hillside or open plains. The substrate at both sites comprised fine Ecca Shale alluvium, siltstone and sandstone gravels that form part of the Ecca Group of the Karoo Supergroup (A. Odendaal and J. Fourie, pers. comm.). Some of the Bankfontein specimens (NCA 2015/1818) were collected during mid-morning (10:00-12:00) foraging in open ground in the vicinity of various ants, including
<taxonomicName id="E868593E1E15083F0C5EEBCC98A8BE51" class="Insecta" family="Formicidae" genus="Anoplolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplolepis custodiens" order="Hymenoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="custodiens">Anoplolepis custodiens</taxonomicName>
F. Smith, 1858,
<taxonomicName id="BDF4DB594F974E59A094BF85CFC56A14" class="Collembola" family="Paronellidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus" order="Entomobryomorpha" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">Camponotus</taxonomicName>
spp.,
<taxonomicName id="9AAC902C46E0052A67208EAFCA09DB41" class="Insecta" family="Formicidae" genus="Messor" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Messor" order="Hymenoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">Messor</taxonomicName>
sp. and
<taxonomicName id="D212C302169DEA3520519F6645DDBD19" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monomorium" order="Hymenoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">Monomorium</taxonomicName>
sp. Of these,
<taxonomicName id="52A3AAD31BEEDEDB8470C33CE3FE9FF2" class="Arachnida" family="Zodariidae" genus="Mallinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mallinus nitidiventris" order="Araneae" pageId="9" pageNumber="150" phylum="Arthropoda" rank="species" species="nitidiventris">Mallinus nitidiventris</taxonomicName>
most closely resembled
<taxonomicName id="15D87446394F9BF10C32056A0378F457" class="Insecta" family="Formicidae" genus="Messor" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Messor" order="Hymenoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">Messor</taxonomicName>
sp. in terms of colouration, although workers of this ant were almost double the body length of the spiders. Only one of these five spiders sampled at this specific site was feeding, a female consuming a
<taxonomicName id="400687803B6B9005088AD082BF927858" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monomorium" order="Hymenoptera" pageId="9" pageNumber="150" phylum="Arthropoda" rank="genus">Monomorium</taxonomicName>
worker ant that measured approximately 2 mm in length, suggesting that this species is myrmecophagous, as are most
<taxonomicName id="5DEC91A6BEEEFB2605F228444BB19B9D" lsidName="" pageId="9" pageNumber="150" rank="subfamily" subfamily="Zodariinae">Zodariinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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<taxonomicName id="75AB0C7BF8354973FCEB70704532DCB6" ID-CoL="5FF6K" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:145979" class="Insecta" family="Formicidae" genus="Aenictus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aenictus luteus moestus Santschi" lsidName-HNS="Aenictus luteus moestus Santschi" order="Hymenoptera" phylum="Arthropoda" rank="variety" species="luteus" variety="moestus">Aenictus luteus Em. v. moestus</taxonomicName>
<taxonomicNameLabel id="533E21C8C93656582214A29C3E8FA612" rank="variety">n. var.</taxonomicNameLabel>
(fig. 18, 19).
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<paragraph id="4EB743E70BD31853150BB9786DA43F52">D'un jaune terne un peu brunatre. Devant et dessus des segments du gastre plus fonce. Tete noire. Nervures alaires brun roussatre. Le reste comme chez le type.</paragraph>
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<subSubSection id="E49B888277E28CEF6E64ECF693C89DBD" type="materials_examined">
<paragraph id="B3BAB088EA704ADF161DE8D4332DFB1F">Soudan francais: Koulouba (Andrieu), 1 [[ male ]].</paragraph>
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<mods:namePart id="5F9CA71DD1A25AAD9BCF5E38996009D8">Notton, David G.</mods:namePart>
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<mods:title id="35CA804366924EC96CEF23034CD2DB95">Biodiversity Data Journal</mods:title>
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<mods:part id="715EA0D61F97321036EBDA901DD16011">
<mods:date id="16A8A6EDDE06D927D2DA8580C90F9522">2016</mods:date>
<mods:detail id="B9B02C12845E87CC88254BED8F25996F" type="volume">
<mods:number id="16CDBF1662AB243727AE6AF0FBC4C30F">4</mods:number>
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<mods:extent id="1343BA6A29129A82DCC54E30DA9BFCDF" unit="page">
<mods:start id="0066D958FA738F394A1B8CFFBF56B6AA">7991</mods:start>
<mods:end id="3664B1AA97FDCB0FE17359F49C1B305C">7991</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="C94D7BCD2DA647FAA6473BFF253C0A89">
<mods:url id="30011F997EEBCE3DB49CF85B2BFA3108">http://dx.doi.org/10.3897/BDJ.4.e7991</mods:url>
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<mods:classification id="32005027124A9E1DF6BC6CE3AD936662">journal article</mods:classification>
<mods:identifier id="56589D778150B9416DA0BA9B0EF2F15A" type="DOI">http://dx.doi.org/10.3897/BDJ.4.e7991</mods:identifier>
<mods:identifier id="5B32C6CC4241FD0C5121888CE0B92FCE" type="Pensoft-Pub">1314-2828--7991</mods:identifier>
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<paragraph id="773BED27C62C300C19A6BEB536D8F632" pageId="0" pageNumber="7991">
<taxonomicName id="4CB76906DA635FA7E9DD1B4FF666E6E6" ID-CoL="53Y93" ID-ENA="2748242" authority="Walker, 1835" authorityName="Walker" authorityYear="1835" class="Insecta" family="Platygastridae" genus="Synopeas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synopeas myles" order="Hymenoptera" pageId="0" pageNumber="7991" phylum="Arthropoda" rank="species" species="myles">Synopeas myles (Walker, 1835)</taxonomicName>
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<paragraph id="1A5A35D2A612C0ADA7A91AF199645541" pageId="0" pageNumber="7991">
<taxonomicName id="2C5A6E9B66B6FF2B7ECE3123FA61D789" class="Insecta" family="Platygastridae" genus="Platygaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platygaster myles" order="Hymenoptera" pageId="0" pageNumber="7991" phylum="Arthropoda" rank="species" species="myles">Platygaster myles</taxonomicName>
Walker, 1835
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<paragraph id="22B31E31B803BD952084C478A9162DFF" pageId="0" pageNumber="7991">Distribution</paragraph>
<paragraph id="FADE7246A0931C5D5C25D816CE354FA9" pageId="0" pageNumber="7991">England, Scotland, Ireland, Isle of Man</paragraph>
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