Add updates up until 2025-01-08 18:54:06
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<document id="BD0E7D72476247929CBF21A07B188A94" ID-DOI="10.11646/zootaxa.5538.1.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="14611362" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1736286526445" checkinUser="plazi" docAuthor="López-García, Margarita M., Domínguez-Trujillo, Mariela & Cisneros-Heredia, Diego F." docDate="2024" docId="03F75E2B24410534FF30F023C6AAFBC4" docLanguage="en" docName="zootaxa.5538.1.10.pdf" docOrigin="Zootaxa 5538 (1)" docSource="https://doi.org/10.11646/zootaxa.5538.1.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Totoia Ocampo 2003" docType="key" docVersion="3" lastPageNumber="99" masterDocId="FFCE265324450530FFA7F33DC337FFBE" masterDocTitle="New species of Totoia Ocampo (Coleoptera: Hybosoridae: Anaidinae) from Ecuador" masterLastPageNumber="100" masterPageNumber="95" pageNumber="99" updateTime="1736362443542" updateUser="guilherme" zenodo-license-document="CLOSED">
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<mods:titleInfo id="69E2FF20EFFB7FF4594CEB9295DE5856">
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<mods:title id="79222BF45F094EF258DADAD032AE0E56">New species of Totoia Ocampo (Coleoptera: Hybosoridae: Anaidinae) from Ecuador</mods:title>
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<mods:name id="06C1FF771DFCB7D90D142EFC85CCA7CA" type="personal">
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<mods:roleTerm id="D278381E8DF2CC495D2C2716113CD591">Author</mods:roleTerm>
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<mods:namePart id="1176EEDC673B4585B0505AE0057BD640">López-García, Margarita M.</mods:namePart>
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<mods:affiliation id="48070FA84286A1AF2EB362A9E273743F">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoología Terrestre, Museo de Zoología, Quito 170901, Ecuador</mods:affiliation>
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<mods:namePart id="93236B5CC346E64596846B2B776684D1">Domínguez-Trujillo, Mariela</mods:namePart>
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<mods:nameIdentifier id="6CA75F255D7FEC97A19E55E2C615BC77" type="ORCID">0000-0003-1203-3021</mods:nameIdentifier>
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<mods:affiliation id="8CF38DBA20E2E4934EC8E0C134C7FF97">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoología Terrestre, Museo de Zoología, Quito 170901, Ecuador</mods:affiliation>
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<mods:nameIdentifier id="9D160ACA7DC3EF7293FCDA26A45F2184" type="email">alejandradominguez227@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0D58340723D1A175334E8B1482980523">Cisneros-Heredia, Diego F.</mods:namePart>
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<mods:nameIdentifier id="E99EF9857862F04C6B28A81536B6562D" type="ORCID">0000-0002-6132-2738</mods:nameIdentifier>
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<mods:affiliation id="3D622E1C1BA1785CDD3D04D0899D39BE">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoología Terrestre, Museo de Zoología, Quito 170901, Ecuador & Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biológicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoología Terrestre, Museo de Zoología, Quito 170901, Ecuador</mods:affiliation>
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<mods:nameIdentifier id="15BC3C78D613FB0B9755A626E10672C2" type="email">dcisneros@usfq.edu.ec</mods:nameIdentifier>
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<mods:title id="EFE91C6A98F39D2E9E588C7E44B26B15">Zootaxa</mods:title>
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<mods:date id="FDD63A397614C5D317CDE5FCC7F13E1C">2024</mods:date>
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<mods:number id="98BF9666AD3E48A8E1233BFC72CE7559">2024-11-13</mods:number>
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<mods:number id="BC742ECE604B95DFBB8AE052D44DA575">5538</mods:number>
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<mods:url id="21B83561FDCA878CF98456B2ACAB27B9">https://doi.org/10.11646/zootaxa.5538.1.10</mods:url>
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<mods:identifier id="B65BE576B690B2A05E15FB8674EF7EFF" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F75E2B24410534FF30F023C6AAFBC4" LSID="urn:lsid:plazi:treatment:03F75E2B24410534FF30F023C6AAFBC4" httpUri="http://treatment.plazi.org/id/03F75E2B24410534FF30F023C6AAFBC4" lastPageNumber="99" pageId="4" pageNumber="99">
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<subSubSection id="C344BCB624410534FF30F023C0F1FC89" box="[151,966,797,824]" pageId="4" pageNumber="99" type="nomenclature">
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<paragraph id="8BE1EF3D24410534FF30F023C0F1FC89" blockId="4.[151,1437,797,1146]" box="[151,966,797,824]" pageId="4" pageNumber="99">
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<heading id="D0A9585124410534FF30F023C0F1FC89" bold="true" box="[151,966,797,824]" fontSize="11" level="1" pageId="4" pageNumber="99" reason="1">
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<emphasis id="B92A332F24410534FF30F023C0F1FC89" bold="true" box="[151,966,797,824]" pageId="4" pageNumber="99">
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Key to the known species of
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||||
<taxonomicName id="4C5E94BE24410534FE5EF023C172FC86" ID-CoL="7XZK" authority="Ocampo" authorityName="Ocampo" authorityYear="2003" box="[505,581,798,824]" class="Insecta" family="Hybosoridae" genus="Totoia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B92A332F24410534FE5EF023C172FC86" bold="true" box="[505,581,798,824]" italics="true" pageId="4" pageNumber="99">Totoia</emphasis>
|
||||
</taxonomicName>
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||||
(modified from Ocampo 2006)
|
||||
</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C344BCB624410534FF30F002C6AAFBC4" pageId="4" pageNumber="99" type="key">
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<keyStep id="30AAF79824410534FF30F002C6ABFCCF" pageId="4" pageNumber="99">
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<paragraph id="8BE1EF3D24410534FF30F002C6ABFCEA" blockId="4.[151,1437,797,1146]" box="[151,1436,831,853]" pageId="4" pageNumber="99">
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||||
<keyLead id="30AF4C0824410534FF30F002C6ABFCEA" box="[151,1436,831,853]" pageId="4" pageNumber="99">1. Pronotum without longitudinal carinae, base with a depressed area; phallobase about three times longer than parameres.... 2</keyLead>
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||||
</paragraph>
|
||||
<paragraph id="8BE1EF3D24410534FF30F061C6ABFCCF" blockId="4.[151,1437,797,1146]" box="[151,1436,860,882]" pageId="4" pageNumber="99">
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||||
<keyLead id="30AF4C0824410534FF30F061C6ABFCCF" box="[151,1436,860,882]" pageId="4" pageNumber="99">1’. Pronotum with six longitudinal carinae; phallobase about 1.2 times longer than parameres........................... 3</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30AAF79824410534FF30F044C6ABFBBA" pageId="4" pageNumber="99">
|
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<paragraph id="8BE1EF3D24410534FF30F044C6AAFC12" blockId="4.[151,1437,797,1146]" pageId="4" pageNumber="99">
|
||||
<keyLead id="30AF4C0824410534FF30F044C6AAFC12" pageId="4" pageNumber="99">
|
||||
2. Surface between discal carinae of elytra with about nine striae of equal width (
|
||||
<figureCitation id="1365F3B824410534FBB5F044C7B6FC30" box="[1042,1153,889,911]" captionStart="FIGURE 1" captionStartId="1.[151,250,1726,1750]" captionTargetBox="[191,1397,344,1702]" captionTargetId="figure-77@1.[191,1397,344,1702]" captionTargetPageId="1" captionText="FIGURE 1. Head and pronotum in frontal view: A. Totoia recki new species; B. Totoia magnifica Ocampo, 2006. Body in dorsal view: C. Totoia recki; D. Totoia magnifica. Body in lateral view: E. Totoia recki; F. Totoia magnifica. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14611366" httpUri="https://zenodo.org/record/14611366/files/figure.png" pageId="4" pageNumber="99">Fig. 1D, F</figureCitation>
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||||
); metatrochanter with small posteriomedial tooth; inner margin of parameres curved (
|
||||
<figureCitation id="1365F3B824410534FCA1F0AAC065FC12" box="[774,850,919,941]" captionStart="FIGURE 2" captionStartId="2.[151,250,1519,1543]" captionTargetBox="[159,1428,818,1489]" captionTargetId="figure-327@2.[158,1430,812,1495]" captionTargetPageId="2" captionText="FIGURE 2. Parameres in dorsal and lateral view: A, C. Totoia recki new species; B, D. T. magnifica Ocampo, 2006. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14611368" httpUri="https://zenodo.org/record/14611368/files/figure.png" pageId="4" pageNumber="99">Fig. 2B</figureCitation>
|
||||
);
|
||||
<collectingCountry id="F349AFAD24410534FCC3F0AAC087FC12" box="[868,944,919,940]" name="Panama" pageId="4" pageNumber="99">Panama</collectingCountry>
|
||||
..................
|
||||
<taxonomicName id="4C5E94BE24410534FB3FF0AAC6AAFC12" ID-CoL="57KFB" authority="Ocampo" authorityName="Ocampo" authorityYear="2006" box="[1176,1437,918,940]" class="Insecta" family="Hybosoridae" genus="Totoia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="magnifica">
|
||||
<emphasis id="B92A332F24410534FB3FF0AAC6AAFC12" bold="true" box="[1176,1437,918,940]" pageId="4" pageNumber="99">
|
||||
<emphasis id="B92A332F24410534FB3FF0AAC677FC12" bold="true" box="[1176,1344,918,940]" italics="true" pageId="4" pageNumber="99">Totoia magnifica</emphasis>
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||||
Ocampo
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BE1EF3D24410534FF30F089C6ABFBBA" blockId="4.[151,1437,797,1146]" pageId="4" pageNumber="99">
|
||||
<keyLead id="30AF4C0824410534FF30F089C6ABFBBA" pageId="4" pageNumber="99">
|
||||
2’. Surface between discal carinae of elytra with about eight striae, two of them wider and separated from each other by two thinner striae (
|
||||
<figureCitation id="1365F3B824410534FECAF0ECC2E4FC58" box="[365,467,977,999]" captionStart="FIGURE 1" captionStartId="1.[151,250,1726,1750]" captionTargetBox="[191,1397,344,1702]" captionTargetId="figure-77@1.[191,1397,344,1702]" captionTargetPageId="1" captionText="FIGURE 1. Head and pronotum in frontal view: A. Totoia recki new species; B. Totoia magnifica Ocampo, 2006. Body in dorsal view: C. Totoia recki; D. Totoia magnifica. Body in lateral view: E. Totoia recki; F. Totoia magnifica. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14611366" httpUri="https://zenodo.org/record/14611366/files/figure.png" pageId="4" pageNumber="99">Fig. 1C, E</figureCitation>
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||||
); metatrochanter without posteriomedial tooth; inner margin of parameres straight (
|
||||
<figureCitation id="1365F3B824410534FB54F0ECC676FC58" box="[1267,1345,977,999]" captionStart="FIGURE 2" captionStartId="2.[151,250,1519,1543]" captionTargetBox="[159,1428,818,1489]" captionTargetId="figure-327@2.[158,1430,812,1495]" captionTargetPageId="2" captionText="FIGURE 2. Parameres in dorsal and lateral view: A, C. Totoia recki new species; B, D. T. magnifica Ocampo, 2006. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.14611368" httpUri="https://zenodo.org/record/14611368/files/figure.png" pageId="4" pageNumber="99">Fig. 2A</figureCitation>
|
||||
); western
|
||||
<collectingCountry id="F349AFAD24410534FF45F0D2C205FBBB" box="[226,306,1007,1029]" name="Ecuador" pageId="4" pageNumber="99">Ecuador</collectingCountry>
|
||||
...................................
|
||||
<taxonomicName id="4C5E94BE24410534FD42F0D2C6ABFBBA" ID-CoL="CPW23" authority="Lopez-Garcia, Dominguez-Trujillo, & Cisneros-Heredia" authorityName="Lopez-Garcia, Dominguez-Trujillo, & Cisneros-Heredia" authorityYear="2024" box="[741,1436,1006,1028]" class="Insecta" family="Hybosoridae" genus="Totoia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="recki">
|
||||
<emphasis id="B92A332F24410534FD42F0D2C6ABFBBA" bold="true" box="[741,1436,1006,1028]" pageId="4" pageNumber="99">
|
||||
<emphasis id="B92A332F24410534FD42F0D2C06DFBBA" bold="true" box="[741,858,1006,1028]" italics="true" pageId="4" pageNumber="99">Totoia recki</emphasis>
|
||||
López-García, Domínguez-Trujillo, & Cisneros-Heredia
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||||
</emphasis>
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||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="30AAF79824410534FF30F731C6AAFBC4" pageId="4" pageNumber="99">
|
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<paragraph id="8BE1EF3D24410534FF30F731C6AAFB81" blockId="4.[151,1437,797,1146]" pageId="4" pageNumber="99">
|
||||
<keyLead id="30AF4C0824410534FF30F731C6AAFB81" pageId="4" pageNumber="99">
|
||||
3. Humeral carina as long or longer than distance between base of elytron and the second discal carina (figure
|
||||
<quantity id="4CA642D824410534FB79F731C635FB9C" box="[1246,1282,1036,1058]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="99" unit="in" value="9.0">9 in</quantity>
|
||||
Ocampo 2003); male parameres as in figures 74A–B (in Ocampo 2006);
|
||||
<collectingCountry id="F349AFAD24410534FCA5F714C050FB81" box="[770,871,1065,1087]" name="Nicaragua" pageId="4" pageNumber="99">Nicaragua</collectingCountry>
|
||||
,
|
||||
<collectingCountry id="F349AFAD24410534FCD6F714C0EBFB81" box="[881,988,1065,1087]" name="Costa Rica" pageId="4" pageNumber="99">Costa Rica</collectingCountry>
|
||||
, and
|
||||
<collectingCountry id="F349AFAD24410534FBA9F717C76DFB81" box="[1038,1114,1066,1087]" name="Panama" pageId="4" pageNumber="99">Panama</collectingCountry>
|
||||
.....
|
||||
<taxonomicName id="4C5E94BE24410534FB3AF714C6AAFB81" ID-CoL="57KFC" authority="Ocampo" authorityName="Ocampo" authorityYear="2003" box="[1181,1437,1065,1087]" class="Insecta" family="Hybosoridae" genus="Totoia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="splendida">
|
||||
<emphasis id="B92A332F24410534FB3AF714C6AAFB81" bold="true" box="[1181,1437,1065,1087]" pageId="4" pageNumber="99">
|
||||
<emphasis id="B92A332F24410534FB3AF714C677FB81" bold="true" box="[1181,1344,1065,1087]" italics="true" pageId="4" pageNumber="99">Totoia splendida</emphasis>
|
||||
Ocampo
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BE1EF3D24410534FF30F77AC6AAFBC4" blockId="4.[151,1437,797,1146]" pageId="4" pageNumber="99">
|
||||
<keyLead id="30AF4C0824410534FF30F77AC6AAFBC4" pageId="4" pageNumber="99">
|
||||
3’. Humeral carina shorter than distance between base of elytron and the second discal carina (figure
|
||||
<quantity id="4CA642D824410534FB3FF77AC7FDFBE3" box="[1176,1226,1095,1117]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="99" unit="in" value="10.0">10 in</quantity>
|
||||
Ocampo 2003); male genitalia as in figures 74c–d (in Ocampo 2006);
|
||||
<collectingCountry id="F349AFAD24410534FD14F759C1C8FBC7" box="[691,767,1124,1145]" name="Panama" pageId="4" pageNumber="99">Panama</collectingCountry>
|
||||
and
|
||||
<collectingCountry id="F349AFAD24410534FC89F759C0BAFBC4" box="[814,909,1124,1146]" name="Colombia" pageId="4" pageNumber="99">Colombia</collectingCountry>
|
||||
...................
|
||||
<taxonomicName id="4C5E94BE24410534FBDFF759C6AAFBC4" ID-CoL="7CJY2" authority="Ocampo" authorityName="Ocampo" authorityYear="2003" box="[1144,1437,1124,1146]" class="Insecta" family="Hybosoridae" genus="Totoia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="99" phylum="Arthropoda" rank="species" species="brachycarina">
|
||||
<emphasis id="B92A332F24410534FBDFF759C6AAFBC4" bold="true" box="[1144,1437,1124,1146]" pageId="4" pageNumber="99">
|
||||
<emphasis id="B92A332F24410534FBDFF759C677FBC4" bold="true" box="[1144,1344,1124,1146]" italics="true" pageId="4" pageNumber="99">Totoia brachycarina</emphasis>
|
||||
Ocampo
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
Loading…
Reference in a new issue