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<document id="FE31CDA6F6607C4EA2C5E7756B29AC2C" ID-DOI="10.3897/zookeys.1216.130270" ID-ZooBank="1221A319-03DC-4157-A7F9-F5A18E20E1FF" ID-publisher-id="130270" URI-arpha="DE3D6390-FB9F-566D-B398-5D22FE3B56DF" URI-zoobank="http://zoobank.org/1221A319-03DC-4157-A7F9-F5A18E20E1FF" article-type="research-article" checkinTime="1729868654848" checkinUser="pensoft" docAuthor="Yetchom Fondjo, Jeanne Agrippine, Nzoko Fiemapong, Armand Richard, Tindo, Maurice, Duressa, Tarekegn Fite, Ivković, Slobodan &amp; Husemann, Martin" docDate="2024" docId="0A2940B8308D547389D9645345E729B7" docLanguage="en" docName="ZooKeys 1216: 219-264" docOrigin="ZooKeys 1216" docSource="https://zookeys.pensoft.net/article/130270/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Pteropera verrucigena Karsch 1891" docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="264" masterDocId="DE3D6390FB9F566DB3985D22FE3B56DF" masterDocTitle="Taxonomic review of the grasshopper genus Pteropera Karsch, 1891 (Orthoptera, Acrididea, Catantopinae) with description of three new species and a preliminary phylogeny of the Cameroonian species" masterLastPageNumber="264" masterPageNumber="219" pageNumber="219" updateTime="1729868729188" updateUser="pensoft">
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<mods:title id="E85300B5DD9A36039BBF092FA3AA95C2">Taxonomic review of the grasshopper genus Pteropera Karsch, 1891 (Orthoptera, Acrididea, Catantopinae) with description of three new species and a preliminary phylogeny of the Cameroonian species</mods:title>
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<mods:namePart id="406879FC924D84A3A517A6BAE924BC6E">Yetchom Fondjo, Jeanne Agrippine</mods:namePart>
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<mods:affiliation id="4A5330A5A0598AF6BCB07C2905E8179C">Zoology Unit, Laboratory of Biology and Physiology of Animal Organisms, Graduate School in Fundamental and Applied Sciences, University of Douala, Douala, Cameroon &amp; Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<mods:namePart id="F3342B563449FBA2CC4E3B5061DBD26F">Nzoko Fiemapong, Armand Richard</mods:namePart>
<mods:nameIdentifier id="63DB139CF99AB685B4B2515F0ADB0E59" type="ORCID">0000-0002-8759-8142</mods:nameIdentifier>
<mods:affiliation id="F87601EBBCD24C6A23BCAC61E9F9C927">University of Neuchâtel, Neuchâtel, Switzerland</mods:affiliation>
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<mods:namePart id="059995EBC47A5CF8D45322C2A70B2F22">Tindo, Maurice</mods:namePart>
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<mods:affiliation id="92550CACA2D1E7BC1A529135C2DCBBC9">Zoology Unit, Laboratory of Biology and Physiology of Animal Organisms, Graduate School in Fundamental and Applied Sciences, University of Douala, Douala, Cameroon</mods:affiliation>
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<mods:namePart id="5879470C12AC1D98C2C44D92F4EAF454">Duressa, Tarekegn Fite</mods:namePart>
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<mods:affiliation id="7FF70154DE44FCE941830A416625733C">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany &amp; School of Plant Sciences, College of Agriculture and Environmental Sciences, Haramaya University, Dire Dawa, Ethiopia</mods:affiliation>
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<mods:namePart id="D14D1B1A775CDB1F0335031E2E0C6BE6">Ivković, Slobodan</mods:namePart>
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<mods:affiliation id="0C9F0B32404CABFE22ED6724DDC372F1">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<mods:namePart id="603116E9A6BFD7E25ED775C3FEA01546">Husemann, Martin</mods:namePart>
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<mods:affiliation id="C0F0A1CC49C8F8FC224206519E80641C">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<mods:date id="336834036649291B766025E0A232E75E">2024</mods:date>
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<mods:number id="E23B4C73528CC63CEF8B65B7805E0569">1216</mods:number>
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<treatment id="0A2940B8308D547389D9645345E729B7" ID-arpha="0A2940B8-308D-5473-89D9-645345E729B7" LSID="urn:lsid:plazi:treatment:0A2940B8308D547389D9645345E729B7" httpUri="http://treatment.plazi.org/id/0A2940B8308D547389D9645345E729B7">
<subSubSection id="20CC036B80A237A2B8361B97F0D45552" type="nomenclature">
<paragraph id="C8CFCC4AFE0CA2AF018B98546294E31F">
<taxonomicName id="6695E2282164B47AE0212B5A43B8FF7C" ID-CoL="4Q26K" authority="Karsch, 1891" authorityName="Karsch" authorityYear="1891" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="verrucigena">
<emphasis id="CF98CAF2993B3A92C044539E82BF4B04" italics="true">Pteropera verrucigena</emphasis>
<bibRefCitation id="D124F03886DB170349A39975795C7007" DOI="10.1002/mmnd.18910360116" author="Karsch F" issue="1" journalOrPublisher="Berliner Entomologische Zeitschrift" pagination="175-196" refId="B10" refString="Karsch F (1891) Verzeichniss der von Herrn Dr. Paul Preuss in Kamerun erbeuteten Acridiodeen. Berliner Entomologische Zeitschrift 36 (1): 175 - 196. https: // doi. org / 10.1002 / mmnd. 18910360116" title="Verzeichniss der von Herrn Dr. Paul Preuss in Kamerun erbeuteten Acridiodeen." volume="36" year="1891">Karsch, 1891</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="6B2BAED3B69DD5681B3F1F7B7E5BD200" type="description">
<paragraph id="3268FF972ED8E57386B684BF6B4584EE">
<figureCitation id="21243BFA89E3789E3B4014501F370FBF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Images of holotypes, allotypes, letotype and neallotype of Pteropera species in lateral view A P. spleniata (holotype ♂) B P. spleniata (holotype ♀) C P. teocchii (holotype ♂) D P. teocchii (allotype ♀) E P. thibaudi (holotype ♂) F P. thibaudi (allotype ♀) G P. verrucigena (lectotype ♂) H P. verrucigena (♀) I P. villiersi (holotype ♂) J P. villiersi (allotype ♀) K P. uniformis (neallotype ♂)." figureDoi="10.3897/zookeys.1216.130270.figure6" httpUri="https://binary.pensoft.net/fig/1160994">Figs 6 G, H</figureCitation>
,
<figureCitation id="C45C7AC260B49EFF981B5C9033E74B4B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Images of holotypes, allotypes, neallotype and paratype of Pteropera species in dorsal view A P. teocchii (holotype ♂) B P. teocchii (allotype ♀) C P. thibaudi (holotype ♂) D P. thibaudi (allotype ♀) E P. verrucigena (holotype ♂) F P. verrucigena (allotype ♀) G P. villiersi (holotype ♂) H P. villiersi (allotype ♀) I P. uniformis (neallotype ♂) J P. femorata (♀)." figureDoi="10.3897/zookeys.1216.130270.figure11" httpUri="https://binary.pensoft.net/fig/1160999">11 E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERFCK" type="type material">
<paragraph id="2D23AC4A93D57987B7746CF5E1A9C368">
<heading id="3F7117737F689CC2F47E83BFA7E55445" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="BA9426985473DD7923ECB1B337ECE908">
<materialsCitation id="2CEEB0C3E5A7A9183B46A93D220A6B30" collectionCode="MfN, DORSA" collectorName="Dr. Paul Preuss" country="Cameroon" latitude="4.666933" location="Barombi Station" longLatPrecision="1" longitude="9.383317" specimenCode="BA 000175, S 01" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="A0CDA27D54D9D567C573CD4ED2E1AC5E" bold="true">
<emphasis id="F0AFA20BB1C2D6E092022A321C85284E" bold="true" italics="true">
<typeStatus id="FE422A326F0BB7FA4660E397E9BFD5AC" type="lectotype">Lectotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="44B29DFB1FBF2EAB7FE59BF0164AE6CA" name="Cameroon">Cameroon</collectingCountry>
<specimenCount id="CE63F4414E7178F362B7637E18200E3B" count="1" type="male"></specimenCount>
;
<location id="D91984C70B9C4E27BAD567AD3E035D1D" LSID="urn:lsid:plazi:treatment:0A2940B8308D547389D9645345E729B7:D91984C70B9C4E27BAD567AD3E035D1D" country="Cameroon" latitude="4.666933" longLatPrecision="1" longitude="9.383317" name="Barombi Station">Barombi Station</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6FCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.383317,4.666933]}">
<geoCoordinate id="898D80E89CC9A15C33C58261966C903D" degrees="4" direction="north" minutes="40.016" orientation="latitude" precision="1" value="4.666933">4 ° 40.016 ' N</geoCoordinate>
,
<geoCoordinate id="8CE9A5AF6B120EA4AD93B7E5B4195692" degrees="9" direction="east" minutes="22.999" orientation="longitude" precision="1" value="9.383317">9 ° 22.999 ' E</geoCoordinate>
</named-content>
;
<collectorName id="72BEEB67DDA882F632BB672B2807A4FE">Dr. Paul Preuss</collectorName>
leg.;
<named-content id="E140D71E99D59776BA0D3C25A05FD2F2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-f%C3%BCr-naturkunde-berlin" xlink_title="Museum für Naturkunde Berlin, Germany">
<collectionCode id="94C6A559C4DA00050204B5B2DF0DC565" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" type="Museum">MfN</collectionCode>
</named-content>
,
<specimenCode id="579D1CBEA7EF26739C845A0869031DE9" collectionCode="MfN" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" type="Museum">
<accessionNumber id="1512FB6571A0087885C620065C085312" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/BA000175" type="EnaNcbi">BA 000175</accessionNumber>
</specimenCode>
<specimenCode id="C4B1FA28EA745109CECEAED00FD88A2D" collectionCode="MfN" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" type="Museum">S 01</specimenCode>
-
<collectionCode id="D42264FCD72E3B54BFF2E084E33A766E">DORSA</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIGCK" type="material">
<paragraph id="F0BA64877BD0299AC0EB1FECBBA0D98D">
<heading id="8F428BDF68C32ABEF7CA24A11F55AAB6" reason="title">Other material examined.</heading>
</paragraph>
<paragraph id="5312F32CA49AC934D4EB72DECAF14E9B">
<materialsCitation id="83B8A4227813DF811194EDDF977E14AE" collectingDate="2017-04-03" collectionCode="SMNK" collectorName="J. A. Yetchom Fondjo" country="Cameroon" latitude="4.9541664" location="Sohock" longLatPrecision="1" longitude="10.247216" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<collectingCountry id="053E5ADBD4241A4E8FF3CB8310E93557" name="Cameroon">Cameroon</collectingCountry>
<specimenCount id="67201C9F261BC12083A77A62A0AC309F" count="2" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="93F61AE657A03A3907B7CD06B065AA2F" count="1" type="female">1 ♀</specimenCount>
;
<location id="96D38F6F638444702A7E1B4818A5391A" LSID="urn:lsid:plazi:treatment:0A2940B8308D547389D9645345E729B7:96D38F6F638444702A7E1B4818A5391A" country="Cameroon" latitude="4.9541664" longLatPrecision="1" longitude="10.247216" name="Sohock">Sohock</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERGCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[10.247217,4.954167]}">
<geoCoordinate id="D1C9EAEFE1FBAAE842C70FF32B9B0373" degrees="4" direction="north" minutes="57.250" orientation="latitude" precision="1" value="4.9541664">4 ° 57.250 ' N</geoCoordinate>
,
<geoCoordinate id="3ADBC2D4BD8E786DDD823A7F2A6A50D9" degrees="10" direction="east" minutes="14.833" orientation="longitude" precision="1" value="10.247216">10 ° 14.833 ' E</geoCoordinate>
</named-content>
;
<date id="48FA5F003682C732742DC814DFE9B65D" value="2017-04-03">
<collectingDate id="27DDAA3BCB9B7B294A23A7C8DD6C2445" value="2017-04-03">3 Apr. 2017</collectingDate>
</date>
;
<collectorName id="F5D15EBE1CA76E4FC9B3508393E0126F">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="E05BDADF85DF208E4453533DF3CABBA7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="B60C743A4F223F60ECE900309F33EF13" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1GCK" type="redescription">
<paragraph id="5060D69CD9D16F5C82F23648E035D963">
<heading id="30E35A1FA2BA2A8BF9AE7A602063CAA3" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="72F2CB9D640A3229BAB5867B4EBD4BC1">
<emphasis id="FFC95CFDAA12DAA043957D6F9A8C6219" bold="true">Male</emphasis>
: medium size, integument rugous; head conical and oblique; fastigium of vertex short with obtuse apex; eyes prominent and globose; antenna, filiform longer than head and pronotum combined; pronotum without lateral carinae and with straight median carina, crossed by three sulci, its anterior and posterior margins rounded and incised in the middle; pale basal band of lateral lobes of pronotum narrowed in front of second transverse furrow but not interrupted; longitudinal median band of pronotum disc dark and less wider than adjacent clear bands; prozona longer than metazona; prosternal tubercle conical; anterior margin of mesosternum broadly projected medially; mesosternal space open and longer than it is wide; elytra vestigial or lobiform; median pale spot on inner area of hind femora absent; outer area of hind femora with three pale spots; incipient spots along medio-superior margin at level of outer spots present; dorsal carina of hind femora finely toothed; lower outer areas of hind femora dark, wine-colored; hind tibiae wine-colored; distal half of hind tibiae widened, basal ring present; supra-anal plate subconical, with two digital tubercles near lateral margins; subgenital plate short conical, gradually tapering towards rounded apex; cerci conical, curved inward and without preapical lobule.
<emphasis id="D0FAC2D7BF0F95CC3EBE96DB384221E6" bold="true" italics="true">Epiphallus</emphasis>
(Fig.
<figureCitation id="822990B54880CC6C4DBE9CC2CF505632" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Internal genitalia of P. carnapi (A D), P. descampsi (E H), P. karschi zenkeri (I L), P. uniformis (M P) and P. verrucigena (O T)." figureDoi="10.3897/zookeys.1216.130270.figure15" httpUri="https://binary.pensoft.net/fig/1161003">15 Q</figureCitation>
): bridge narrow, short and arched; ancorae small, close together and with acute apex; lophi short, broad and lobiform; lateral plates broad and rounded; anterior and posterior projections short.
<emphasis id="5E6599D510AE5607FB68096F6C56E0E0" bold="true" italics="true">Phallic complex</emphasis>
(Fig.
<figureCitation id="4BF10BBA9190F6A9A1DC46CC42F7629D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Internal genitalia of P. carnapi (A D), P. descampsi (E H), P. karschi zenkeri (I L), P. uniformis (M P) and P. verrucigena (O T)." figureDoi="10.3897/zookeys.1216.130270.figure15" httpUri="https://binary.pensoft.net/fig/1161003">15 R T</figureCitation>
): dorsal arch of the cingulum V-shaped, strongly open not overlying the endophallic sclerites; latero-ventral sclerite broad, subtriangular; zygoma wide; apodemes of the cingulum long, reaching the apex of the endophallic apodemes; lower ectophallic sheath not enveloping the base of the rami; upper ectophallic sheath globular, sloping forward.
</paragraph>
<paragraph id="2336D5E6F167C44447CE0A0E6256E03C">
<emphasis id="BAE7F2F8701B053940FAD06F5EDC1E1D" bold="true">Female</emphasis>
: Similar to the male but larger; supra-anal plate conical with a transverse groove in the middle field; posterior edge of the subgenital plate projecting; cercus conical with angular apex; dorsal valves of the ovipositor weakly toothed; base of the spermathecal duct widened well before it opened into the copulatory bursa; spermatheca ampulla relatively thin; distal, recurrent trunk of the spermatheca lateral diverticulum&gt; 3 × longer than the proximal trunk.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECICK" type="remarks">
<paragraph id="3F2369DFBD6CE1FB1DD9D27BA271469F">
<heading id="19D00FC82F0E638597C9A140ACBC676B" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="967EC07691DD281247A55D0F8E973311">
<taxonomicName id="BABD5A0328E6FCB19DADCE1A210FE4A8" authorityName="Karsch" authorityYear="1891" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="verrucigena">
<emphasis id="04242C7DE31ECA94DCAAAB0ED3412B1B" italics="true">Pteropera verrucigena</emphasis>
</taxonomicName>
was originally described from Barombi station (southwest
<collectingCountry id="430C48EAB9FCBED5221DE93E28AEA099" name="Cameroon">Cameroon</collectingCountry>
) with some
<typeStatus id="4D75C275CACF69F2AEFA3882BCF39B38" type="paratype">paratypes</typeStatus>
recorded between Kumba-Mamfé (Southwest
<collectingCountry id="3FE7044658DB2263D9666E88DE6888F5" name="Cameroon">Cameroon</collectingCountry>
) by
<bibRefCitation id="74628DCB9CAB71F480E11D074DCE876D" DOI="10.1002/mmnd.18910360116" author="Karsch F" firstAuthor="Karsch" issue="1" journalOrPublisher="Berliner Entomologische Zeitschrift" pagination="175-196" refId="B10" refString="Karsch F (1891) Verzeichniss der von Herrn Dr. Paul Preuss in Kamerun erbeuteten Acridiodeen. Berliner Entomologische Zeitschrift 36 (1): 175196. https://doi.org/10.1002/mmnd.18910360116" title="Verzeichniss der von Herrn Dr. Paul Preuss in Kamerun erbeuteten Acridiodeen." volume="36" year="1891">Karsch (1891)</bibRefCitation>
. We now additionally recorded the species from Sohock (Littoral
<collectingCountry id="4B09C500C4B50664AF767248A550CB0E" name="Cameroon">Cameroon</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXICK" type="distribution">
<paragraph id="BA66E41501B672340A0367F57660BC96">
<heading id="246780C9074AA63AA45459C1D42AF3A0" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="74F63374058CE24327D9B1A8DB2D6467">
<collectingCountry id="4C4DB020C6DA47CA77FF49DCFE5BE734" name="Cameroon">Cameroon</collectingCountry>
(Fig.
<figureCitation id="ED367869822C7C397123992295D99311" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of Pteropera species in African rainforests." figureDoi="10.3897/zookeys.1216.130270.figure18" httpUri="https://binary.pensoft.net/fig/1161006">18 B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,462 @@
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<mods:title id="FF499296B30C16B79F779D7703A139D6">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<mods:namePart id="E6D5CEFC6A3B236AB61B87CC00D56255">Li, Jinglin</mods:namePart>
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<mods:affiliation id="C0CC02B8DCFAC0C21E555E69A0D4B412">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:namePart id="D9071B4869949F658DCE1FB023BF90CC">Li, Shuqiang</mods:namePart>
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<mods:affiliation id="FE6CFA51C96CC55D9AC6864DB45AE9A1">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="AA795F9F686A741593482E1253599F04">Yao, Zhiyuan</mods:namePart>
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<mods:affiliation id="7B914FE008967F1CCDAC26FBA91369E1">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<treatment id="0A3BBAAB9E695516BEF769327FD252EF" ID-DOI="http://doi.org/10.5281/zenodo.13992768" ID-Zenodo-Dep="13992768" ID-arpha="0A3BBAAB-9E69-5516-BEF7-69327FD252EF" ID-zoobank="https://zoobank.org/1E1B4058-113E-48F6-9734-39D486FF0FB9" LSID="urn:lsid:zoobank.org:act:1E1B4058-113E-48F6-9734-39D486FF0FB9" LSID-ZBK="urn:lsid:zoobank.org:act:1E1B4058-113E-48F6-9734-39D486FF0FB9" httpUri="http://treatment.plazi.org/id/0A3BBAAB9E695516BEF769327FD252EF">
<subSubSection id="EC44C1510DC78300FE096E619BD98307" type="nomenclature">
<paragraph id="10B3CEB996F609267F2C9B15D9C093AD">
<taxonomicName id="AD35EF2090B505422092699F332A251F" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="19BB750051F2787B8241C1D54B72FE82" italics="true">Belisana yongcong</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="2F77201A2CD2C9DAF7D623D4205E50CF" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8C82330A88A974FB1DBFBEB87A15CB52" type="description">
<paragraph id="C6B0B038F7BDF96EDC4CB18DEC7BFF4F">
<figureCitation id="FB66DB2DE27896DDDB90F987FB144474" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">Figs 14</figureCitation>
,
<figureCitation id="85EFE8915E35B05F86B683888F89205E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15</figureCitation>
,
<figureCitation id="FF798F433EB20134093C44E729BD6529" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A, B</figureCitation>
,
<figureCitation id="7AB73F6F7FE1CB6BEEDBA3C9483B343E" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIUAI" type="type material">
<paragraph id="2852AC0BDA970E99EBD3A48C1D72AF45">
<heading id="E7FF8DAEFC7A298BBD08C720D330B10D" reason="title">Type material.</heading>
</paragraph>
<paragraph id="4758EE97AB6B21E475D506E73B440741">
<materialsCitation id="F45C9D06D45587356B4B54B4EBEAE47D" collectingDate="2007-05-23" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Liping County" elevation="544" latitude="25.9838" location="Yongcong Town" longLatPrecision="1" longitude="109.11632" municipality="Qiandongnan" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="DA1AE2D9F5C47576E92E16794F4E3E4F" bold="true">
<emphasis id="789EE1AA866047525F61295B42E07356" bold="true" italics="true">
<typeStatus id="4A32BB7F200C0ADCD7FCDAE057C2FEFE" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="C5BAD92999ED82D53554177305C46306" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="02A841C688F6884B75BDA97C0BA43399" count="1" type="male"></specimenCount>
;
<collectingRegion id="E7A415FBD59AE649FE901C537883C5CA" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingMunicipality id="47B0D637F7A8503FA002140ABCBBE7F4">Qiandongnan</collectingMunicipality>
,
<collectingCounty id="7A242C435108A7C03CCC78A37D5BE0AA">Liping County</collectingCounty>
,
<location id="EB2AE286881D54155DEF9E4DED4D6F26" LSID="urn:lsid:plazi:treatment:0A3BBAAB9E695516BEF769327FD252EF:EB2AE286881D54155DEF9E4DED4D6F26" country="China" county="Liping County" latitude="25.9838" longLatPrecision="1" longitude="109.11632" municipality="Qiandongnan" name="Yongcong Town" stateProvince="Guizhou">Yongcong Town</location>
,
<location id="865ADA5EBEE851B6512972810C23D965" LSID="urn:lsid:plazi:treatment:0A3BBAAB9E695516BEF769327FD252EF:865ADA5EBEE851B6512972810C23D965" country="China" county="Liping County" latitude="25.9838" longLatPrecision="1" longitude="109.11632" municipality="Qiandongnan" name="Guantuan Village" stateProvince="Guizhou">Guantuan Village</location>
,
<location id="BF8C054371C50E66C6369D2FCD072114" LSID="urn:lsid:plazi:treatment:0A3BBAAB9E695516BEF769327FD252EF:BF8C054371C50E66C6369D2FCD072114" country="China" county="Liping County" latitude="25.9838" longLatPrecision="1" longitude="109.11632" municipality="Qiandongnan" name="Guantuan Cave" stateProvince="Guizhou">Guantuan Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWUAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[109.116317,25.983800]}">
<geoCoordinate id="B86A2E2008464241EB7DA67097B7BEE7" degrees="25" direction="north" minutes="59.028" orientation="latitude" precision="1" value="25.9838">25 ° 59.028 ' N</geoCoordinate>
,
<geoCoordinate id="48A2C2B0417111DA6187A45B3EEA0F73" degrees="109" direction="east" minutes="06.979" orientation="longitude" precision="1" value="109.11632">109 ° 06.979 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="C49EF18FBA2208B71701CF7BF32AF52A" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" unit="m" value="544.0">
<elevation id="C21C47D780BC360D6B4642E96C21B9FB" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" unit="m" value="544.0">544 m</elevation>
</quantity>
;
<date id="306F01121AF2CB9037808461EAE42D36" value="2007-05-23">
<collectingDate id="5FF1B5AF758B52B6D171B30232CC9162" value="2007-05-23">23 May 2007</collectingDate>
</date>
;
<collectorName id="DEC8AA90D68552DB74EEF9CA54B8D379">J. Liu</collectorName>
&amp;
<collectorName id="F2E25EFFD31DC0F3E294C1B502CEAB20">Y. Lin</collectorName>
leg.;
<named-content id="CB00DD673C18A0F47B468E078B1C9723" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="2AD9B85A9AB47E195BB37E248731BD27" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="C9ACA715A216F6FBABB360BB390475CE">Ar</collectorName>
45218
</materialsCitation>
.
<materialsCitation id="2B6091AC6E11E02201EFA3F4193B8F12" collectingDate="2007-05-23" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Liping County" elevation="544" latitude="25.9838" location="Yongcong Town" longLatPrecision="1" longitude="109.11632" municipality="Qiandongnan" specimenCount="2" specimenCount-male="2" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="7510BD6BDB755EDA44E2AD0548AD8561" bold="true">
<emphasis id="8B2E86802D988A119B8E85C365620EF7" bold="true" italics="true">
<typeStatus id="91D205DFDE6E9F2448014A3AB8429CF5" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="FF0A5CDE61F5661EA546D1502609902D" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="F49EE154DF5CDEA6FFDFBB1D44949FE7" count="2" type="male">2 ♂</specimenCount>
; same data as for holotype;
<named-content id="0F1CE3D3E6C546DA57DDB370BF2573DD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="812EA57D8F8C79D252050EFAAB5F46BD" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4521920
</materialsCitation>
<materialsCitation id="2F067B092FA1149DDB020120AFDEDA63" collectingDate="2007-05-23" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Liping County" elevation="544" latitude="25.9838" location="Yongcong Town" longLatPrecision="1" longitude="109.11632" municipality="Qiandongnan" specimenCount="5" specimenCount-female="5" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="C6DB5BB0B6C7F5E54802BAB3CD86A0C4" count="5" type="female">5 ♀</specimenCount>
; same data as for holotype;
<named-content id="E76714BD7A29F3DA7CF7F5120F4122E3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="C4D81800021B380120F6DE07E3972388" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4522125
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELVAI" type="diagnosis">
<paragraph id="2795AAE6A9C75A6DAD9B328A29587844">
<heading id="46461715F0470051C71609C0F407FA48" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="1CFD74FD1D9C2068E2D8D6BFC6ECC7FF">
The new species resembles
<taxonomicName id="119DB947A3C309EC2BE0C6376338C3C9" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nayong" status="sp. nov.">
<emphasis id="8CE892AD515DC0DD6481B8EF6BF7C783" italics="true">B. nayong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F2E04392E5C5A21F3EBB0381855EA557" rank="species">sp. nov.</taxonomicNameLabel>
(Figs
<figureCitation id="1E4E5F4D178CEEED3BF48E7B533FA19C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8</figureCitation>
,
<figureCitation id="622E8C4924F57F1F2FEBA42E81FFF31E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9</figureCitation>
,
<figureCitation id="882F96A6636A30D0825B55571B56B964" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 G, H</figureCitation>
,
<figureCitation id="9BE98CBDF53663D5472270A074CF9E2B" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 G, H</figureCitation>
) by having similar bulbal apophysis (hooked; Fig.
<figureCitation id="CD13860494FC9DE9E9BA303F206C7449" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 C</figureCitation>
) and vulva (anterior arch ridge-shaped, pore plates curved, long elliptic and 8 × longer than wide; Figs
<figureCitation id="2843106302A76D2509D6A8683A91BB3E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 B</figureCitation>
,
<figureCitation id="E37200AAE4079D63407FF236AF767967" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 D</figureCitation>
), but can be distinguished by procursus with blunt ventro-subdistal membranous process (arrow
<quantity id="FC26D145C072BC099FAB6894D94BCECF" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="B504101C84DF2279B4E2E219D224FD8A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 C</figureCitation>
,
<figureCitation id="1048AE44E8C248B3E147744C5FA9C013" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A</figureCitation>
vs pointed), by male cheliceral distal apophyses pointing outwards (
<abbrev id="ABBRID0EAXAI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="985480961DECC2C058705294C8C462F2" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 D</figureCitation>
vs inwards), and by epigyne with lateral pockets (
<abbrev id="ABBRID0EIXAI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="12F7DED755AB284476EEC0F42E0E86E5" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 A</figureCitation>
,
<figureCitation id="99D785FC437874EB4789144A48EBB7FA" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 C</figureCitation>
vs postero-median).
</paragraph>
<caption id="28AA904B691A0D2EE386C1AF712DC176" ID-DOI="10.3897/zookeys.1216.132561.figure14" ID-arpha="5F656357-BA10-542A-A335-57B3191C9436" httpUri="https://binary.pensoft.net/fig/1161059" startId="F14">
<paragraph id="1000EBE2F9E6775235D01D47D53E983F">
<label id="3B89632B84BA412D5D18A1B9509517FC">Figure 14.</label>
</paragraph>
<paragraph id="119691F24EF2F295C83927554B3F91DB">
<taxonomicName id="6251AB9AD8060D9770E5C751DA2DC841" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="BC795235696A52F38F63B839D693D40F" italics="true">Belisana yongcong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5263C2A0D46BE346B4A5664D768FD913" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="3A78C3A7EF0C0122E49050D13A5AF9B7" bold="true">A, B</emphasis>
palp (
<emphasis id="2C195939DEBE43C6239FAA68D591F92F" bold="true">A</emphasis>
prolateral view
<emphasis id="256038764CA43EEE36C9327C574F5A53" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="B87181E1868D6F5BE58EBD6FF5095234" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="5E24673EDCA1E140F592EF9E28714883" bold="true">C</emphasis>
prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis
<emphasis id="628ED961383680AA06F76F74113ACBFD" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb,
<abbrev id="ABBRID0ETYAI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0EXYAI" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="B9EAEFC6EE36273436A9A18B9EC74B3B" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="9FD3B9ADEDF68CD8F9B26BDD1DF16BFF" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="7651A7CFFDBC6874C1FC2E457EFAD1F5" ID-DOI="10.3897/zookeys.1216.132561.figure15" ID-arpha="13B34B84-B654-5554-97FE-D90C46E63039" httpUri="https://binary.pensoft.net/fig/1161060" startId="F15">
<paragraph id="50CA1F7E56785CAF3A0888BA33615A91">
<label id="898016993234B2ADC3E9471297C68BBB">Figure 15.</label>
</paragraph>
<paragraph id="231D4D7E57CCA6DAB44E7239BABB5DDE">
<taxonomicName id="F858B08340DE99DD97D3262095ED53E2" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="759FE0722EAD5701D79221F189FECCDF" italics="true">Belisana yongcong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13F136D813C64C52063452865CF4618F" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="AEB485E1234EC40535987DD72D332449" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="0B8C49BFC86698285101FDD2E7BDBE63" bold="true">A, B, G, H</emphasis>
)
<emphasis id="7977D7ECDE80976D9FE01F22699C6CE7" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="80D0921F800C9D5C5D1FCAD2291403FA" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="5132D8632D8539333A5FDB635DB1CBD7" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="BA36CE362838B858F7896EA09FE92DC0" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="70D98B52164825BE43BAE5EB152813A2" bold="true">E H</emphasis>
habitus (
<emphasis id="64AA3482369D6CB93768104216572FFB" bold="true">E, G</emphasis>
dorsal view
<emphasis id="92BC6861DDDC6FF65339E9BFE9D1E836" bold="true">F</emphasis>
lateral view
<emphasis id="D5646FE0BA9695ADEBC4D934AC6F5AD2" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EL1AI" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EP1AI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0ET1AI" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EX1AI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0E21AI" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0E61AI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 (
<emphasis id="34C6D7C15A5A44C4B3B096A4FD4FB9E7" bold="true">A D</emphasis>
); 0.40 (
<emphasis id="35E54E177E58ED3BEC13A5D17B05822C" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EM2AI" type="description">
<paragraph id="C384EF4ECEC16358488FAF32C0F4C13B">
<heading id="8A10232D2C633604F7D2BF36883F1883" reason="title">Description.</heading>
</paragraph>
<paragraph id="DE5B6C3F53F45E32DE9CEEEA548329B3">
<emphasis id="D728FB96CF1C79A88577399BAB555851" bold="true">Male</emphasis>
(
<emphasis id="4DA5FEF8BB3FC7D6FC823153943788FF" bold="true">
<typeStatus id="5F8A5E677AA8C481B863FE6CDD74E923" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.73 (1.82 with clypeus), prosoma 0.83 long, 0.67 wide, opisthosoma 0.90 long, 0.66 wide. Leg I: 14.76 (3.60, 0.34, 4.05, 5.58, 1.56), leg II: 12.45 (3.36, 0.32, 3.08, 4.60, 1.09), leg III: 8.21 (2.38, 0.26, 1.98, 2.84, 0.75), leg IV: 10.64 (3.20, 0.26, 2.66, 3.72, 0.80); tibia I
<abbrev id="ABBRID0EW2AI" xlink_title="length/diameter">L / d</abbrev>
: 46. Eye interdistances and diameters:
<abbrev id="ABBRID0E12AI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E52AI" xlink_title="posterior median eye">PME</abbrev>
0.06,
<abbrev id="ABBRID0EC3AI" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EG3AI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EK3AI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.54 / 0.49. Habitus as in Fig.
<figureCitation id="9C6755D0F305F25A70374BECBF8653D1" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, with brownish radiating marks; clypeus and sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0ES3AI" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="A05E4F44D4B9288EF0C454F6A6B95C5F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.32;
<abbrev id="ABBRID0E13AI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="9C7C5A2D0F97175A200EBAA24975AD18" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 D</figureCitation>
). Palp as in Fig.
<figureCitation id="2A58551CC2EAB6FE9AB573CA332553D5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
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Fig.
<figureCitation id="8BFB77AA75AB499CD665B24B2EA1E911" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="AA74845FFA4F8E06B46F0A67D0774BAD" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="F30DC805ACC49C5DC440F2A555DE929F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 B</figureCitation>
); procursus simple proximally but complex distally, with ventro-subdistal membranous lamella (arrow
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Figs
<figureCitation id="61058A9F1C98FB55DDD07EB362C6A8C5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 C</figureCitation>
,
<figureCitation id="4C6D044C21FE8D3ABE024E6E55FFA5FF" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A</figureCitation>
) bearing proximally sclerotized part, prolatero-distal membranous lamella (arrow
<quantity id="7A4CC4CD8249149FE8EBAD45F7B8DE93" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="0A279E94052738304385CC4611B9EF4A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 C</figureCitation>
,
<figureCitation id="4E03A66A1D2063D9BA459669FB11CCB7" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A</figureCitation>
) bearing proximally sclerotized part, sclerotized dorso-subdistal apophysis (arrow
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Figs
<figureCitation id="68B863C592A29446A24CE71B0EFDB0BA" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 C</figureCitation>
,
<figureCitation id="5B8D912C0A1C514C5510FAC81E62C922" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="9C88EB8227F36B0FD8333870C2A81287" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14 D</figureCitation>
,
<figureCitation id="A5C22BA9DB46BF04B5ED50138E4113B3" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 B</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EO5AI" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="D033D622A7CEF254B7B33DA5C740207C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="446A9E10F5859E521C49293E8897C421" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 C</figureCitation>
). Retrolateral trichobothria on tibia I at 10 % proximally; legs with short vertical setae on metatarsi; tarsus I with 20 distinct pseudosegments.
</paragraph>
<paragraph id="AC16706B4577CEA243C59A093FE2F7D3">
<emphasis id="2BE1311248F5C67F2C62B9108F06706B" bold="true">Female</emphasis>
(
<emphasis id="27FFAB6D56C589F6A701092DA847D6F0" bold="true">
<typeStatus id="8C7A2ABD09A330240C16D10E37753BA3" type="paratype">paratype</typeStatus>
</emphasis>
,
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<collectionCode id="A8AC9C5B037B390B1C9951C5F9263804" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45221): Similar to male, habitus as in Fig.
<figureCitation id="F5458D05EE807C2C01B66EDB139C47E3" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 G, H</figureCitation>
. Total length 2.17 (2.29 with clypeus), prosoma 0.87 long, 0.76 wide, opisthosoma 1.30 long, 1.08 wide; tibia I: 3.65; tibia I
<abbrev id="ABBRID0EI6AI" xlink_title="length/diameter">L / d</abbrev>
: 47. Eye interdistances and diameters:
<abbrev id="ABBRID0EM6AI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EQ6AI" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EU6AI" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EY6AI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E36AI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.59 / 0.45. Epigyne simple and flat, posteriorly curved, with pair of lateral pockets 0.35 apart (
<abbrev id="ABBRID0EAABI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="17649A7F9536611652566ADD793577F5" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 A</figureCitation>
,
<figureCitation id="AC8C6306480D20D1B34225E3D211E1D3" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 C</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0EMABI" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="F4835899F898D9E9E32B8B0FE54CF921" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 B</figureCitation>
,
<figureCitation id="B211FB56E6AEDA995D5467F3D70587B2" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 D</figureCitation>
) and pair of curved, long elliptic pore plates (9 × longer than wide,
<abbrev id="ABBRID0EYABI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="D05F7DBE1FF48939432F919E1CC4C87B" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15 B</figureCitation>
,
<figureCitation id="6A0F1794CEA46D96A4FBDB3C4910EE81" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEBBI" type="variation">
<paragraph id="3ED59D51287080444CCCC47E38A02939">
<heading id="4B6FE540057CD67AA10B95901B9E9777" reason="title">Variation.</heading>
</paragraph>
<paragraph id="DD48DE64B5AA070513ADB2802248652E">
Tibia I in
<specimenCount id="BAF72977FCD93D3C11CFC5174AD92088" count="1" type="male">one male</specimenCount>
<typeStatus id="1D395A079AC9B42EF93E8FBB32C37232" type="paratype">paratype</typeStatus>
(
<named-content id="5188D2E6711D25D6BA8750183D97C887" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="110D1581519D6F7623C58BFFD36E5EDA" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45219): 5.32 (leg I missing in
<named-content id="8B41E652133493D584A9A3BBF8347965" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="691B5AD71C96E69C3C7ED0A82A64C3A8" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45220). Tibia I in the other
<specimenCount id="CA90D9E641578E9A0EAE977C327A6EF1" count="4" type="female">four female</specimenCount>
<typeStatus id="03873553D8179C5B55E895771A25A974" type="paratype">paratypes</typeStatus>
(
<named-content id="7F0E175122BBA16B3EDFC35CDE602FD9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="4E4A0D877A15273E27D838A23D63598D" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4522225): 3.76, 3.85, 3.96, 4.16.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWBBI" type="habitat">
<paragraph id="70C6274EE2EED3908D97A32E1B6E5D48">
<heading id="7FCD3065B55F97AEE261C2B1DD0342C0" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="D7B707704703EF5EC72D9FB3F9E5F16B">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2BBI" type="distribution">
<paragraph id="66124BB45953CF2D50BC2B418CAD4473">
<heading id="0D694D3CA342C1858376AE16463395CE" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="7D59A0F4AA036FF30BE19D0E7F770F10">
<collectingCountry id="7F7BC44E4B77FB932647FEB139E81B79" name="China">China</collectingCountry>
(
<collectingRegion id="BD0C17938584A71879BFE3D3A228FB45" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="663EDD33147525D954EB7EFCF466CD93">type</typeStatus>
locality; Fig.
<figureCitation id="1E7C05C3E9872644E583E2786270CD14" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFCBI" type="etymology">
<paragraph id="19F10BA43FD171EB838E08F45EFD5D03">
<heading id="73EC9B34C7D3EB9704339566F8457FFC" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="64EAD874C0B143B813A4687881A7F143">
The specific name refers to the
<typeStatus id="7D282D033C536A093585A1A3898A2C72">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="84D68E1ED77E052FD871EAB701E815C2">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<mods:title id="04D10418A9A2C441D8598EADACB92867">ZooKeys</mods:title>
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<mods:part id="AE3C742CD1DEADE4AB2878783B941A71">
<mods:date id="98B52139874E3B7D8152C8C45EE5BE3E">2024</mods:date>
<mods:detail id="6F36D3477A1CA7904FCBCDBB3EA77A77" type="pubDate">
<mods:number id="DCCEF1B6D5B3C67672B49DF34F4F273A">2024-10-25</mods:number>
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<mods:detail id="D2DD72FE43E508E65801FCA4E35F2035" type="volume">
<mods:number id="2E7ED584EDB80FECB8A598C123164D9E">1216</mods:number>
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<mods:classification id="C59A98341A97EE5095AB23C68A920B24">journal article</mods:classification>
<mods:identifier id="F37ED10FB54D34BC659EF26BE1C51E70" type="DOI">10.3897/zookeys.1216.132561</mods:identifier>
<mods:identifier id="91094D33B8E8304F210B3B0A36870BC0" type="ZooBank">0E173F72-36D3-4B97-BC43-F096067A3D1E</mods:identifier>
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<treatment id="0B2A9B44F3CE5520B3C980275FF5193E" ID-DOI="http://doi.org/10.5281/zenodo.13992750" ID-Zenodo-Dep="13992750" ID-arpha="0B2A9B44-F3CE-5520-B3C9-80275FF5193E" ID-zoobank="https://zoobank.org/938091C8-867C-4607-9EB1-9B4BE301F78F" LSID="urn:lsid:zoobank.org:act:938091C8-867C-4607-9EB1-9B4BE301F78F" LSID-ZBK="urn:lsid:zoobank.org:act:938091C8-867C-4607-9EB1-9B4BE301F78F" httpUri="http://treatment.plazi.org/id/0B2A9B44F3CE5520B3C980275FF5193E">
<subSubSection id="2E4E14613FCAF337F1038AC986280A59" type="nomenclature">
<paragraph id="87C05D6B278F07CA2674B316B430B319">
<taxonomicName id="49357C3A9733D9C4934687854EC50A87" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bijie" status="sp. nov.">
<emphasis id="084837C095F12E6BF092DB523158E7FA" italics="true">Belisana bijie</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="0A6B66A847CC12D33504E81F04E1BCA0" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B3896655300196DD60DA5DA3FF44A6A5" type="description">
<paragraph id="2D3BB423748FB27C4E7170AF8D1B2EE7">
<figureCitation id="DCEFA8C07E02884CCF265535945A6FF1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">Figs 2</figureCitation>
,
<figureCitation id="77A21D94322DE572B014FD6922214BB5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3</figureCitation>
,
<figureCitation id="35DB366E08AACA2B8961555A83345D3D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 A, B</figureCitation>
,
<figureCitation id="E65AE5856311591872B929E611FCE100" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENWAE" type="type material">
<paragraph id="EF1F79347CEF253D66E60F3B6B4C4B0B">
<heading id="2E7B585EC63E0F3F641AA82E468B87E0" reason="title">Type material.</heading>
</paragraph>
<paragraph id="09EDF7C9166812D5BF01DECB28CAC809">
<materialsCitation id="AA25CBB4593D50F4EF3E41320C88FEFE" collectingDate="2011-11-18" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zha &amp; Ar" country="China" county="Bijie" elevation="1920" latitude="27.19155" location="Salaxi Town" longLatPrecision="1" longitude="105.06416" municipality="Qixingguan District" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="24E21304588EC3EA0D7953D2F01E327F" bold="true">
<emphasis id="6246F199235953B5274A197BEF86BD27" bold="true" italics="true">
<typeStatus id="9DC1F6BB3137856E5D3A87213CC2F4EE" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="BE410C3C7EFE951BA465EBB33EC1C063" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="8B4F0A79717F4897D8507B180DD65728" count="1" type="male"></specimenCount>
;
<collectingRegion id="A768103DB57AFFFA427FD5145D9C8D1E" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingCounty id="F92AB64588C6179583F60FE3EC5F4260">Bijie</collectingCounty>
,
<collectingMunicipality id="6E46ADACEF5BDFFC681A387B16640E3F">Qixingguan District</collectingMunicipality>
,
<location id="6DE12F28F7572A309368A61263B4E4AD" LSID="urn:lsid:plazi:treatment:0B2A9B44F3CE5520B3C980275FF5193E:6DE12F28F7572A309368A61263B4E4AD" country="China" county="Bijie" latitude="27.19155" longLatPrecision="1" longitude="105.06416" municipality="Qixingguan District" name="Salaxi Town" stateProvince="Guizhou">Salaxi Town</location>
,
<location id="71AECA72BAA7FDBAD690C125A783AB5B" LSID="urn:lsid:plazi:treatment:0B2A9B44F3CE5520B3C980275FF5193E:71AECA72BAA7FDBAD690C125A783AB5B" country="China" county="Bijie" latitude="27.19155" longLatPrecision="1" longitude="105.06416" municipality="Qixingguan District" name="Shuanglongdao Cave" stateProvince="Guizhou">Shuanglongdao Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2WAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[105.064167,27.191550]}">
<geoCoordinate id="C7182B085106167A23BD5602164BBF0F" degrees="27" direction="north" minutes="11.493" orientation="latitude" precision="1" value="27.19155">27 ° 11.493 ' N</geoCoordinate>
,
<geoCoordinate id="720E19524258FBFC80D5EF8BE6BE13A1" degrees="105" direction="east" minutes="03.850" orientation="longitude" precision="1" value="105.06416">105 ° 03.850 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="1A78B3CFC881A98B74E4EF105B7DEFDC" metricMagnitude="3" metricUnit="m" metricValue="1.92" unit="m" value="1920.0">
<elevation id="AFCA3A4D896D5686D6E78430F4656701" metricMagnitude="3" metricUnit="m" metricValue="1.92" unit="m" value="1920.0">1920 m</elevation>
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;
<date id="6807CC1DFF7A2D9F4D557BED4FA99C37" value="2011-11-18">
<collectingDate id="4030FC23ABD33F08AB8A7FAE909A4099" value="2011-11-18">18 Nov. 2011</collectingDate>
</date>
;
<collectorName id="D3FAA3AB77D8EF322DD01774084CA1E3">Z. Chen</collectorName>
&amp;
<collectorName id="863672E9F2F6A810EFDB1BE4F5751B67">Z. Zha</collectorName>
leg.;
<named-content id="6B91BB61E1EEBB6EF19AEE0687A246B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="E91A8570326FD1B142ABC3DC9DFF63FF" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="707D3F4AD2BE282509DBDB0225B70D00">Ar</collectorName>
45175
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.
<materialsCitation id="BF382D659A60330C28F7DF656DDCDD79" collectingDate="2011-11-18" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zha &amp; Ar" country="China" county="Bijie" elevation="1920" latitude="27.19155" location="Salaxi Town" longLatPrecision="1" longitude="105.06416" municipality="Qixingguan District" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="E227FBDBBF6F38485B0A82BDDEA06930" bold="true">
<emphasis id="FD602CD7BE8E66C613E7CC6F2999F94D" bold="true" italics="true">
<typeStatus id="7578B305AFC8C64AE02639E83C96E0BD" type="paratype">Paratype</typeStatus>
</emphasis>
:
<collectingCountry id="23E20EAA1ABC91880312B4A58DC84F72" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="4BE0AC51340E736259A24E2AFCDE9CAB" count="1" type="female">1 ♀</specimenCount>
; same data as for holotype;
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<collectionCode id="9403D7E956B2BCE47136C213814FC5F6" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
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- Ar 45176
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.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMXAE" type="diagnosis">
<paragraph id="B5D915F373254A61C6C4C659C679C667">
<heading id="2774BC1F5F8967DB60BD7F94A09F0DB4" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="F0A479FD450DA1C41045E4A4EA7C5E97">
The new species resembles
<taxonomicName id="36A0A9B36EC36CE93FFC68859B8F8139" authority="Wang, Yao &amp; Zhang, 2024" authorityName="Wang, Yao &amp; Zhang" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="wangchengi">
<emphasis id="C62BB8A66F910A60377C5B1E58E9F894" italics="true">B. wangchengi</emphasis>
<bibRefCitation id="7FBB05AB7383F89A649F4806BFE563D1" DOI="10.3897/zookeys.1209.127951" author="Zhang L &amp; Wang Y &amp; Li S &amp; Zhang X &amp; Yao Z" journalOrPublisher="ZooKeys" pagination="315-330" refId="B23" refString="Zhang L, Wang Y, Li S, Zhang X, Yao Z (2024 a) Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China. ZooKeys 1209: 315 - 330. https: // doi. org / 10.3897 / zookeys. 1209.127951" title="Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China." volume="1209" year="2024">Wang, Yao &amp; Zhang, 2024</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="263996EE0B9E6ED602DD5A476BBA98DD" DOI="10.3897/BDJ.12.e125111" author="Wang B &amp; Yao Z &amp; Zhang X" etAl="et al." firstAuthor="Wang" page="2" refId="B8" refString="Wang B, Yao Z, Zhang X (2024) A new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from Guizhou Province, south-western China. Biodiversity Data Journal 12: e 125111 [17]. https://doi.org/10.3897/BDJ.12.e125111" year="2024">Wang et al. 2024: 2</bibRefCitation>
, figs 1 A D, 2 A H, 3 A D) by having similar male chelicerae (tips of distal apophyses widely separated and pointing inwards; Fig.
<figureCitation id="142E973ABCBBE1DB07A43183DDB75C07" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 D</figureCitation>
), bulbal apophysis (hooked; Fig.
<figureCitation id="0810CC6075204FE3733D60CC7921E67C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 C</figureCitation>
), and epigyne (epigynal pockets on lateral part of epigynal plate; Figs
<figureCitation id="91C6E0ACFE29AADFB19FEBD95DA7B2CD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 A</figureCitation>
,
<figureCitation id="52CA4C10B7E085637651A0B058623FF2" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 A</figureCitation>
), but can be distinguished by procursus with retrolatero-subdistal membranous process (arrow
<quantity id="8E2FFB9BE806719ED6531919CAC313FC" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="D7472637C123BBCCDF852EB5D1D84665" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 D</figureCitation>
,
<figureCitation id="0BA657D340A974388C1DA2FE436A53D1" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 B</figureCitation>
vs absent), by vulval pore plates with nearly angular sclerites (arrow in Figs
<figureCitation id="112584047E383BEFB61FC2EF7A732E5F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 B</figureCitation>
,
<figureCitation id="E93A0AEF51EF197A6669B94740B204C0" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 B</figureCitation>
vs blunt), and by dorsal shield of prosoma with lateral brown bands (Fig.
<figureCitation id="A9546BCD70AA5A8FB27087B4D9699781" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 E, G</figureCitation>
vs absent, but with median radiating marks).
</paragraph>
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<paragraph id="5D7AC8FAD9870F8BA238D4CE2AF40839">
<label id="81DDF1F6F99C3536835BD3B706A33052">Figure 2.</label>
</paragraph>
<paragraph id="A8FA2DC7AFB0836DB2A0F3880F133987">
<taxonomicName id="FDEDD6D18F2AC9D77EE59A752832D415" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bijie" status="sp. nov.">
<emphasis id="8CEFCD2034894352E784DC65DEDC4737" italics="true">Belisana bijie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43419AF37EC00B4FE12CD74A53434821" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="7A61E3CCB211AF758C9E7D2FD2EB66CA" bold="true">A, B</emphasis>
palp (
<emphasis id="7BDF14A9F79DC0FDDF8C0FA36D8DF81E" bold="true">A</emphasis>
prolateral view
<emphasis id="8E37A490D7CF3B3B016B7408723FEA8C" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="AC29B5F4590992644379F68DDD9F2E0C" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="636A11C55E558BF8977FADC4B850F4AB" bold="true">C</emphasis>
prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella
<emphasis id="EFFC6FA901D3DD749CFFEA3D0F3A0EB0" bold="true">D</emphasis>
retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb,
<abbrev id="ABBRID0EE1AE" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0EI1AE" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.20 (
<emphasis id="302FC03D26012B279AA1097BA9DC0482" bold="true">A, B</emphasis>
); 0.05 (
<emphasis id="7624631AF6AE6EC4AAABB0974233845A" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="6A9F724BE1DA27F87898885E46C346B8" ID-DOI="10.3897/zookeys.1216.132561.figure3" ID-arpha="564960EC-56BD-5B90-92EC-C07A0E9B40EB" httpUri="https://binary.pensoft.net/fig/1161048" startId="F3">
<paragraph id="7ADB9D964CEE11D0A5DFFAEC18560F5E">
<label id="E6A16D114A76F782F2F1F63A84E53F55">Figure 3.</label>
</paragraph>
<paragraph id="4FE2D0218329E0B5C9CE814A3907F1E8">
<taxonomicName id="432FEF47537CC137442A95583231CEE6" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bijie" status="sp. nov.">
<emphasis id="38DCA431DFAA05A343A34EB908F94C50" italics="true">Belisana bijie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5614F4BD74E4F986F717AF8EF95169B0" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="B2622630D13A555AE4C7B30F759314FA" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="636B62BC094F2EE2E89F7A0BB65C52F0" bold="true">A, B, G, H</emphasis>
)
<emphasis id="893A616040EF44B299D0B61BE7BD6BF7" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="2A3812FFE8EC21EFD78A8427779ED832" bold="true">B</emphasis>
vulva, dorsal view, arrow points at lateral sclerite
<emphasis id="3275C6198A6120816134B5CA2C7B207E" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="64FA75F09F9CC46C01895A214B13C2D8" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="F88380F55574685C90AC6B77F019E47B" bold="true">E H</emphasis>
habitus (
<emphasis id="C3F95940AC8DBF8BC91B1B00911AAE92" bold="true">E, G</emphasis>
dorsal view
<emphasis id="080A6374787B98BF661226A2D91B0922" bold="true">F</emphasis>
lateral view
<emphasis id="4EB1FEFAD51BEB1150EB5E5A9D577138" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0E32AE" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EA3AE" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EE3AE" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EI3AE" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EM3AE" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0EQ3AE" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 (
<emphasis id="7E477A1F26DC6930E0D0939A0C9802D3" bold="true">A D</emphasis>
); 0.40 (
<emphasis id="73E41DBDE89ECD868554A634CDD5F6D5" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E43AE" type="description">
<paragraph id="156932FDB13A6317C5D15A3B60915DB1">
<heading id="A3230485D78B12C9FFEB7400056F3E4D" reason="title">Description.</heading>
</paragraph>
<paragraph id="B5FA338E28C5CAF529C2A5DAB7AD0689">
<emphasis id="411C7F05A101A7ECB884612B39195B52" bold="true">Male</emphasis>
(
<emphasis id="A07ADB5E5CEC14382BEEDEC275079F2D" bold="true">
<typeStatus id="DE88FB0F39ACAA00D287876F3A1F38DD" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.95 (2.08 with clypeus), prosoma 0.70 long, 0.82 wide, opisthosoma 1.25 long, 0.90 wide. Leg I: 19.96 (5.13, 0.38, 5.05, 7.52, 1.88), leg II: 13.25 (3.52, 0.37, 3.32, 4.70, 1.34), leg III: 7.81 (2.25, 0.30, 1.90, 2.59, 0.77), leg IV: 9.83 (3.09, 0.30, 2.45, 3.16, 0.83); tibia I
<abbrev id="ABBRID0EH4AE" xlink_title="length/diameter">L / d</abbrev>
: 73. Eye interdistances and diameters:
<abbrev id="ABBRID0EL4AE" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EP4AE" xlink_title="posterior median eye">PME</abbrev>
0.11,
<abbrev id="ABBRID0ET4AE" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0EX4AE" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E24AE" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.63 / 0.56. Habitus as in Fig.
<figureCitation id="D889CC69D2A1AB7395FD30A73133FA4D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, with brown lateral bands; clypeus brown; sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, with black spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0ED5AE" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="0901F49D7CD5D844AD6CF6458FF42343" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.31;
<abbrev id="ABBRID0EL5AE" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="35805FDCE584BA98C0CB925AE37D8DB2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 D</figureCitation>
). Palp as in Fig.
<figureCitation id="708CF7DE316646F2A7DFE2B0D4EB55B2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 A, B</figureCitation>
; trochanter with ventral apophysis (2 × longer than wide, arrow
<quantity id="EEC73625E97B05E566A0FFD599E5031E" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="6EB8B7AECC90406BEF9AD057C764007F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="3C0DFB79C8CFCD20E25B32EBC9BDF40B" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="A3099002F1C53F276D8A9FB2A151802D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 B</figureCitation>
); procursus simple proximally but complex distally, with prolatero-subdistal sclerite (arrow
<quantity id="F0C946F6CDECB83ABD8C29D7B3EF91AC" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="C7B4E63467FF2C1FBED429E984AD73B5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 C</figureCitation>
,
<figureCitation id="A84A644BA468793E76DF26B511D2341A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 A</figureCitation>
), prolatero-distal membranous lamella (arrow
<quantity id="C5A01B882EB1AD2FBDCC4D0EC2D146BF" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="AE625F394CD8BF454E248BAE962E5624" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 C</figureCitation>
,
<figureCitation id="7AD476DBEE7BEA4FAE0967A9E01CE44B" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 A</figureCitation>
) bearing proximally sclerotized part, sclerotized dorso-subdistal apophysis (arrow
<quantity id="6C7725ADE89F580528DDF7062CC82933" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="706F1AF63D084C904B47AE3D5788564C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 D</figureCitation>
,
<figureCitation id="2FD350ACCD2AFC02B6C495F8A564C9C3" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 B</figureCitation>
), retrolatero-distal spine (arrow
<quantity id="0F5B279F63CEF11503A41FF5B4F9E206" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="6A2BF197FF74021FC5A904F84A5C1305" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 D</figureCitation>
,
<figureCitation id="94A08EC5414516C28FA060F696434A04" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 B</figureCitation>
), retrolatero-subdistal membranous process (arrow
<quantity id="CE874A94F0196DDC21E90EC27DC50076" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="5F3E3DE1CE095C7FAE11F7138FE4F109" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 D</figureCitation>
,
<figureCitation id="20CAA2C631A9A40E6D2D123FE5C69E9A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 B</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="0AA4210AE0182243BFF067A93DDCEDDA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Belisana bijie sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine, arrow 3 points at retrolatero-subdistal membranous process). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure2" httpUri="https://binary.pensoft.net/fig/1161047">2 D</figureCitation>
,
<figureCitation id="E61451D6E101B110970ADD563E941BE3" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 B</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EQAAG" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="3185BDFC61B5C6D13ACD4BA57EFF2584" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="797638600BC52BA0FED1C965927DBCFF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 C</figureCitation>
). Retrolateral trichobothria on tibia I at 9 % proximally; legs with short vertical setae on metatarsi; tarsus I with 17 distinct pseudosegments.
</paragraph>
<paragraph id="479F557244E27C9B6F56C2B4F3D3E36B">
<emphasis id="1C8D805583A8CB5149D1507F58375706" bold="true">Female</emphasis>
(
<emphasis id="7A3F84605CADB4027BC704DF48050E67" bold="true">
<typeStatus id="3EB42B12B23F1E19B8375649B573F7DF" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="43BE0BCA273C49E27951DFAD2E6FB1EA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="D579A6876C1A4C9137497A197E299C20" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45176): Similar to male, habitus as in Fig.
<figureCitation id="29EEFA7392C0FC3B3CE20401BBEA8A91" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 G, H</figureCitation>
. Total length 1.96 (2.09 with clypeus), prosoma 0.66 long, 0.83 wide, opisthosoma 1.30 long, 1.01 wide; tibia I: 3.40; tibia I
<abbrev id="ABBRID0EKBAG" xlink_title="length/diameter">L / d</abbrev>
: 49. Eye interdistances and diameters:
<abbrev id="ABBRID0EOBAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0ESBAG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0EWBAG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0E1BAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E5BAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.60 / 0.53. Dorsal shield of prosoma with distinct postero-median marks; clypeus yellowish. Epigyne simple and flat, posteriorly curved, with pair of lateral pockets 0.31 apart (
<abbrev id="ABBRID0ECCAG" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="76997EA5361C2D7DB2F5E6460DC2B55F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 A</figureCitation>
,
<figureCitation id="7B07B28CC157847738623E375A258410" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 A</figureCitation>
). Vulva with ridge-shaped anterior arch bearing pair of nearly angular sclerites (arrow in Figs
<figureCitation id="4593E71EB8B3A02DD05BB01D1F28A836" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 B</figureCitation>
,
<figureCitation id="B174400F9387D3D3E511076EB7F0C51B" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 B</figureCitation>
), and pair of nearly triangular pore plates (4 × longer than wide,
<abbrev id="ABBRID0EWCAG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="B974BE3A977ACA8B297FCFD8B9B5FF10" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Belisana bijie sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure3" httpUri="https://binary.pensoft.net/fig/1161048">3 B</figureCitation>
,
<figureCitation id="7AF5AD17AE9BDC6D928EC8AD3526655A" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECDAG" type="habitat">
<paragraph id="ECEA7C114B5A41E64FC781604E2B948D">
<heading id="635C534FEB0F6F7940CDD73CBF588E00" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="7E298F934B4946F7AFA014D038F9DFF1">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EHDAG" type="distribution">
<paragraph id="4304D58C6EB9B03729AFF53B428D6FF4">
<heading id="328F32697403E2695BCE4821E9368785" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="45A18F8ABE584264EB28FDC8FDB14EBB">
<collectingCountry id="3B39D31FE7EDF0D165267DACC41ED086" name="China">China</collectingCountry>
(
<collectingRegion id="367D3C6AA66BA077D63A60A04F3F2828" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="64BB20684C701C90D7A43C5DB2B7A4B4">type</typeStatus>
locality; Fig.
<figureCitation id="877888ECF034F53D3E6518AC9E23CE33" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERDAG" type="etymology">
<paragraph id="7EC01BD04A8971EF6E5034A3221C2DA5">
<heading id="5BD2DA1E51A397A3DF6CBDB4F94FC819" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="5CA6AE69735A1754C4FDC29EE85C9B64">
The specific name refers to the
<typeStatus id="A14B4A53FE9CD95150338764CF8C3E5B">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,516 @@
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<mods:title id="10A32933E9200A8E15EECE5DD5AE4F04">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<subSubSection id="4BE0F466222574375A89686BC58E88EA" type="nomenclature">
<paragraph id="8D8FF7548020976BED0269C24BC2BDC2">
<taxonomicName id="8F5887B0A9459F2D9E99DCEB9ABE0B9C" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="qixingguan" status="sp. nov.">
<emphasis id="F18DF5EF8E4B04F2A7C7296343990DC5" italics="true">Belisana qixingguan</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="47773CCF236D1A72962BFD0EBB04D75E" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="A984ADF8F0855BF6C2E48AF3B7FB4FFA" type="description">
<paragraph id="7FD2B647E68F7B153AC20DF314CCE64B">
<figureCitation id="77A75590255E3781B568C0A0864413DA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana qixingguan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure10" httpUri="https://binary.pensoft.net/fig/1161055">Figs 10</figureCitation>
,
<figureCitation id="A518CCA3C1C689B372FB159969716340" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11</figureCitation>
,
<figureCitation id="43E258772B0558B7B9BD815DCDB26F6F" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 I, J</figureCitation>
,
<figureCitation id="8A2C03C9C70EC5A7538E6AA5CB4FBBC5" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 I, J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2VBG" type="type material">
<paragraph id="F68CFDD2101B6A3A60D73B77E9C6CD52">
<heading id="F59658B147F8B185E36D367FDF63BEFF" reason="title">Type material.</heading>
</paragraph>
<paragraph id="0F811D04B6A14B51703D05E46CE8224A">
<materialsCitation id="3CF234E41ABB0AE8330524E92B0E18FD" collectingDate="2007-04-29" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Bijie" elevation="1580" latitude="27.231733" location="Changchun Town" longLatPrecision="1" longitude="105.17329" municipality="Qixingguan District" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="0C3EEFCC8B86379587E10CC1CCD78AC9" bold="true">
<emphasis id="FCBFDC6BD4E7E76C170AE381B61340F0" bold="true" italics="true">
<typeStatus id="9099C28A5031ECC5F2CF62AD40AD1700" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="7FA800A607AB0DFC504CFA36E839041E" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="FBEEB6CA25C07E703A08CF4099AA0DFD" count="1" type="male"></specimenCount>
;
<collectingRegion id="824AAA7F95AED50BCA1CF362360B2311" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingCounty id="4FF610C1BCE5FC16F825A0E234B0401A">Bijie</collectingCounty>
,
<collectingMunicipality id="C88F5BC043A487E3BC96A433FF975B73">Qixingguan District</collectingMunicipality>
,
<location id="3095BE36A82ABBB3CBE717C74D65D609" LSID="urn:lsid:plazi:treatment:12EF7C27F2705B43803B632567E519EA:3095BE36A82ABBB3CBE717C74D65D609" country="China" county="Bijie" latitude="27.231733" longLatPrecision="1" longitude="105.17329" municipality="Qixingguan District" name="Changchun Town" stateProvince="Guizhou">Changchun Town</location>
,
<location id="A39583794F8B4D5D8888A09910F81599" LSID="urn:lsid:plazi:treatment:12EF7C27F2705B43803B632567E519EA:A39583794F8B4D5D8888A09910F81599" country="China" county="Bijie" latitude="27.231733" longLatPrecision="1" longitude="105.17329" municipality="Qixingguan District" name="Changchun Village" stateProvince="Guizhou">Changchun Village</location>
,
<location id="1E7FB5A280668C619655BF1692D37642" LSID="urn:lsid:plazi:treatment:12EF7C27F2705B43803B632567E519EA:1E7FB5A280668C619655BF1692D37642" country="China" county="Bijie" latitude="27.231733" longLatPrecision="1" longitude="105.17329" municipality="Qixingguan District" name="Changchun Cave" stateProvince="Guizhou">Changchun Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJWBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[105.173283,27.231733]}">
<geoCoordinate id="1177AF3F5B9B13966C5BEE16D99A9DC8" degrees="27" direction="north" minutes="13.904" orientation="latitude" precision="1" value="27.231733">27 ° 13.904 ' N</geoCoordinate>
,
<geoCoordinate id="7519EF29B916E71A67F6B7BC9E11EC81" degrees="105" direction="east" minutes="10.397" orientation="longitude" precision="1" value="105.17329">105 ° 10.397 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="8DE5900A6F0FD6B281BCA13ACCF1CE78" metricMagnitude="3" metricUnit="m" metricValue="1.58" unit="m" value="1580.0">
<elevation id="73A70D900C3825477D2922021005A2E9" metricMagnitude="3" metricUnit="m" metricValue="1.58" unit="m" value="1580.0">1580 m</elevation>
</quantity>
;
<date id="0A7FF7DF861948369A1FF11C580BC31A" value="2007-04-29">
<collectingDate id="D4D66558DC36730421BFCCB8CAE11CBF" value="2007-04-29">29 Apr. 2007</collectingDate>
</date>
;
<collectorName id="3B3790DE007A31F3F23DB8BE1C835F35">J. Liu</collectorName>
&amp;
<collectorName id="99CFBAD905853545FBDD0AD51B394031">Y. Lin</collectorName>
leg.;
<named-content id="EFADE4117EB464D53D27845721449A01" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="78732DFBA435D702514D8EE279DF2392" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="9B3E43B5A6508A07F91E5A9AE39EB2CE">Ar</collectorName>
45206
</materialsCitation>
.
<materialsCitation id="B34EA793DFC4067552EE3329C9A6AA73" collectingDate="2007-04-29" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Bijie" elevation="1580" latitude="27.231733" location="Changchun Town" longLatPrecision="1" longitude="105.17329" municipality="Qixingguan District" specimenCount="7" specimenCount-female="7" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="7CB8AE993A506D0192BE908B4872D35C" bold="true">
<emphasis id="28BFBB98303305AD17D200D9FB434219" bold="true" italics="true">
<typeStatus id="A24BA74A932F453BF4D99B7D53B94AE6" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="142E7A836AA14D1E5B107C73C3DAFA48" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="9EFA6B5C1F1BA47C07783987CCC16E07" count="7" type="female">7 ♀</specimenCount>
; same data as for holotype;
<named-content id="49590CD795822F6C6EFEFC9737D522EE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="14FC130292DC6E1C1D9A270A9C2B515D" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4520713
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1WBG" type="diagnosis">
<paragraph id="3664BC1FB0D71BC6AFE1E62670C08AE2">
<heading id="AC908BE684F79BC8D0E392DE7EC32F39" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="56F5C9F8B850E1491A45CD91C1DC4032">
The new species resembles
<taxonomicName id="BCE8720F2D44594CC8D8EF95A7DA5942" authority="Zhang, Li &amp; Yao, 2024" authorityName="Zhang, Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tongi">
<emphasis id="7BD3BBCBD26BC22B3DD1D5D33E372D2A" italics="true">B. tongi</emphasis>
<bibRefCitation id="04C917E882C42EB52B8C76E6231D9472" DOI="10.3897/zookeys.1209.127951" author="Zhang L &amp; Wang Y &amp; Li S &amp; Zhang X &amp; Yao Z" journalOrPublisher="ZooKeys" pagination="315-330" refId="B23" refString="Zhang L, Wang Y, Li S, Zhang X, Yao Z (2024 a) Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China. ZooKeys 1209: 315 - 330. https: // doi. org / 10.3897 / zookeys. 1209.127951" title="Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China." volume="1209" year="2024">Zhang, Li &amp; Yao, 2024</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="BDD85767F6CE3823811ECB3778AAA1B2" DOI="10.3897/zookeys.1202.121633" author="Zhang L &amp; Wu Z &amp; Li S &amp; Yao Z" journalOrPublisher="ZooKeys" pagination="255-286" refId="B24" refString="Zhang L, Wu Z, Li S, Yao Z (2024 b) Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China. ZooKeys 1202: 255286. https://doi.org/10.3897/zookeys.1202.121633" title="Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China." volume="1202">Zhang et al. 2024 b</bibRefCitation>
: 273, figs 12 A D, 13 A H, 18 K, L, 21 C, D) by having similar distal part of procursus (distal membranous process nearly half-round; arrow
<quantity id="D45792238AB1CA898B86C2020A4006E4" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="0F6ED6DAC3E09BE83AF49E97177479C4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana qixingguan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure10" httpUri="https://binary.pensoft.net/fig/1161055">10 C</figureCitation>
,
<figureCitation id="BC681C3FFF09CE94716E53B163ABEA7D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 I</figureCitation>
), bulbal apophysis (hooked; Fig.
<figureCitation id="CBEB8F3E4CD0B4FF14D389B50345C99F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 C</figureCitation>
), and epigyne (epigynal pockets on postero-lateral part of epigynal plate, epigynal plate posteriorly curved; Figs
<figureCitation id="B3C44B469EB0FF33AEDA316780D744E3" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 A</figureCitation>
,
<figureCitation id="0AF49FD7E39EE61DA8500EABF9B57838" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 I</figureCitation>
), but can be distinguished by procursus without distal membranous lamella (Figs
<figureCitation id="C21A3E1A9673A7C2FA5FFEB86A123228" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana qixingguan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure10" httpUri="https://binary.pensoft.net/fig/1161055">10 C</figureCitation>
,
<figureCitation id="0579D090EEB046FA2D4E35D4B63188BA" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 I</figureCitation>
vs present), by male cheliceral distal apophyses pointing downwards (
<abbrev id="ABBRID0ELYBG" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="0F3EEDFCC07E70EC36EB77068B59C6A4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 D</figureCitation>
vs outwards), by vulva without teeth (Figs
<figureCitation id="5F1188271EDDF9380D3554D3D47EC45B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 B</figureCitation>
,
<figureCitation id="3ADECD4874FAB5D021C813F003AE1406" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 J</figureCitation>
vs present), and by vulval pore plates medially narrow and posteriorly strongly widened (
<abbrev id="ABBRID0E2YBG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="8189BED129B822775B98C894AF0B940C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 B</figureCitation>
,
<figureCitation id="7B6395C3A62040D8F54995F298CAAC1C" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 J</figureCitation>
vs nearly triangular).
</paragraph>
<caption id="9B5DFAED35ADB0B69EBD6C6E91D6CBD8" ID-DOI="10.3897/zookeys.1216.132561.figure10" ID-arpha="260422A0-CD22-57F9-AD75-ED54DAAA7E3E" httpUri="https://binary.pensoft.net/fig/1161055" startId="F10">
<paragraph id="D5EDE9965AA1F657D1E31A8D85F9BCE9">
<label id="3FDD3F357C6EADBAF3DAE82CD06A974D">Figure 10.</label>
</paragraph>
<paragraph id="B6AAEB9BF8A180992C89854622A0654D">
<taxonomicName id="99131F73999E938BD8C5EB6C4E211C6C" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="qixingguan" status="sp. nov.">
<emphasis id="A3C51A90489062AACB7BC11B2AC032F9" italics="true">Belisana qixingguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F5172640ED61FC856D743174E3FD48A8" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="A287B53D423F67E9BE14DEB47F555EA2" bold="true">A, B</emphasis>
palp (
<emphasis id="9F883156668B70F89B20F3B461215420" bold="true">A</emphasis>
prolateral view
<emphasis id="8E0DBF19AA5DEEF313CDFC07588ED2A1" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="60CA49DD67E55A6C3A69399DB4F92E4B" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="DDA77F67CA99BD12D5E1311BA0802FC9" bold="true">C</emphasis>
prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine
<emphasis id="8097FCAD304115D0874CFA920C576D43" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb,
<abbrev id="ABBRID0EG1BG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus,
<abbrev id="ABBRID0EK1BG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="738DAA87A9A2DA7D3162FAAC7AEB5AD7" bold="true">A, B</emphasis>
); 0.05 (
<emphasis id="E88C47F420460EC2FA441B8CB0B50ACA" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="9692E847D761041245DD560928072C22" ID-DOI="10.3897/zookeys.1216.132561.figure11" ID-arpha="224C330D-23DC-514D-B441-1579D51D328B" httpUri="https://binary.pensoft.net/fig/1161056" startId="F11">
<paragraph id="F3E984024CD600C0E12488413F6B3C13">
<label id="BA49A5FD6A3FE31D58BEA41ADF8E0CAF">Figure 11.</label>
</paragraph>
<paragraph id="1A9324D109725C4149BEBA66C769DEA5">
<taxonomicName id="4DAF2E499EB3FE3AFDE7177B94C94259" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="qixingguan" status="sp. nov.">
<emphasis id="53A8CD7DD86BBF7FAEBD23615C610122" italics="true">Belisana qixingguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9297C77819B49C22EEA41056F85B95B" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="DC7FA6FA11CA266EFF996F57761C82E4" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="BCBC24C8074319732AE928730AD5069D" bold="true">A, B, G, H</emphasis>
)
<emphasis id="558D6050A3923477FF4D9FAB90B39E26" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="DC610D40138B344828B3B0A2B233274B" bold="true">B</emphasis>
vulva, dorsal view, arrow points at sclerotized protrusion
<emphasis id="5A8B07BD75FC1ABE6058D7D2809D8F8F" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="9691F1C643758E13144100D05B8EBF7F" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="2687CBA0ABD1134374890D44728B2F98" bold="true">E H</emphasis>
habitus (
<emphasis id="A3465279FED49C29A3A1060D0C611684" bold="true">E, G</emphasis>
dorsal view
<emphasis id="ADA451BD15A1A293A9188B716793DCA7" bold="true">F</emphasis>
lateral view
<emphasis id="0B110AD3D0FBAE1EEF8FE7A36597B024" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0E52BG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EC3BG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EG3BG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EK3BG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EO3BG" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0ES3BG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 (
<emphasis id="8B65B4467E9AD9C674125F0268119C3D" bold="true">A D</emphasis>
); 0.50 (
<emphasis id="1B6E1C31DEF3D684029B5EEC7F18A921" bold="true">E H</emphasis>
).
</paragraph>
</caption>
<caption id="28CA07FED871ACCD7761913A3FDA310E" ID-DOI="10.3897/zookeys.1216.132561.figure12" ID-arpha="0362F4E8-2CE5-5D21-8101-691DDF1E5704" httpUri="https://binary.pensoft.net/fig/1161057" startId="F12">
<paragraph id="4A22800E499AF1F5049A5B4996BCD487">
<label id="AFBFF4FF64C065FA55D8BEF96167FF9C">Figure 12.</label>
</paragraph>
<paragraph id="B1A3AC35CA0F619F7BCADEBA67E18230">
<taxonomicName id="F6BC6E83FDD0C3D75CDFF6303EE41DF7" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="xiuwen" status="sp. nov.">
<emphasis id="78BEABB7DBD2C943FBDB8C8C4BCA7F33" italics="true">Belisana xiuwen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D9018846963D7AFF6A409AB0EB04441" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="4C0C68D8A70EC038D8CA5738F2A2AB84" bold="true">A, B</emphasis>
palp (
<emphasis id="42934D34F513624971B9D011F9B3DB68" bold="true">A</emphasis>
prolateral view
<emphasis id="3755469C3018A5E803A0B9E658BDC62C" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="4503AE52E8B5BCEAEDD658269409F533" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="92D486F396FCF628C2F12A7459FBCFA3" bold="true">C</emphasis>
prolateral view, arrow 1 points at distal spine
<emphasis id="0EB5ABC1156DE0CFA920E79EFED5E36A" bold="true">D</emphasis>
retrolateral view). Abbreviations: f = flap,
<abbrev id="ABBRID0E54BG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="A1B455BE08FAD0E65CAA156FC0540495" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="C7EDC222E5BA4E9B0D8C34D48C9240D0" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="8FAC603D7BAE9DC8F5884C24453F7590" ID-DOI="10.3897/zookeys.1216.132561.figure13" ID-arpha="40ED5424-C87F-5DC8-902A-EA3095488026" httpUri="https://binary.pensoft.net/fig/1161058" startId="F13">
<paragraph id="E2E092A9E2C6F893F483A72A50BCFE6C">
<label id="55F7BAA1EF7C950079B29188D97EFDAB">Figure 13.</label>
</paragraph>
<paragraph id="C7EFA09D44C87237695F4A73EF4BD7ED">
<taxonomicName id="D80A83FE593901ADC684C439A281C417" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="xiuwen" status="sp. nov.">
<emphasis id="FE71041350D0603A19A83739B34ADA5D" italics="true">Belisana xiuwen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11886DD5222BD12481010343917AEBCD" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="F15C49152D07A09D682F4A64E3794118" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="B936D4A4ED4EDA7EB9F5953884C4F9DD" bold="true">A, B, G, H</emphasis>
)
<emphasis id="34611D8D5EA65A65BE0C0C7697C80800" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="55D1848EF9641916A451F7AC4B5E5A9A" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="01547679F25CBAA64052BB974A2D4EE5" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="588D476F6FF9752DE8F1B178D0727189" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="0D26C4550D70780BA2FFC4CD9799C9A0" bold="true">E H</emphasis>
habitus (
<emphasis id="F7DEF8744BAB9B37C71ED35EE01FCBA8" bold="true">E, G</emphasis>
dorsal view
<emphasis id="2325E3A13CCB94BAC3641BF0F98A70D2" bold="true">F</emphasis>
lateral view
<emphasis id="CE1B6C983BF426D32A5F06526269AD43" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0ES6BG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EW6BG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0E16BG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0E56BG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EDAAI" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0EHAAI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.05 (
<emphasis id="D8A7CC9C90955EAAA58F4D71495EE55B" bold="true">A D</emphasis>
); 0.50 (
<emphasis id="14A6348AB8CC06B0A3A5375D5385B1B8" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EUAAI" type="description">
<paragraph id="56A34C0F43AC630723157245626D9757">
<heading id="FFA1805C42081184C8584F26853E2A94" reason="title">Description.</heading>
</paragraph>
<paragraph id="D51153D6C1F84FBE1FC427F41223695A">
<emphasis id="14EB2D887BCEA37E6F5CA36E6495B3FB" bold="true">Male</emphasis>
(
<emphasis id="8B483EE77EC0CBE2A0968D696BC51BDE" bold="true">
<typeStatus id="43EE7C582B26CC877924E2637E2F4C62" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.78 (1.89 with clypeus), prosoma 0.68 long, 0.75 wide, opisthosoma 1.10 long, 0.77 wide. Leg I: 14.43 (3.80, 0.31, 3.72, 5.00, 1.60), leg II: (3.00, 0.31, 2.66, 3.75, ), leg III missing, leg IV: 7.17 (2.13, 0.27, 1.66, 2.41, 0.70); tibia I
<abbrev id="ABBRID0E5AAI" xlink_title="length/diameter">L / d</abbrev>
: 50. Eye interdistances and diameters:
<abbrev id="ABBRID0ECBAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EGBAI" xlink_title="posterior median eye">PME</abbrev>
0.14,
<abbrev id="ABBRID0EKBAI" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EOBAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0ESBAI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.55 / 0.48. Habitus as in Fig.
<figureCitation id="B2FF2091D3DB17426204FA1F2DACEB6F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, with brown radiating marks; ocular area with brown median stripe; clypeus brown; sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, with black spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0E1BAI" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="CB2BA50E9533C22C5ECCF9A1DAE90B81" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.29;
<abbrev id="ABBRID0ECCAI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="6A9971A2FC6F2B4FD6A05F824217FEE2" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 D</figureCitation>
). Palp as in Fig.
<figureCitation id="DF3A5A6BDA7FCE510BD3896F6107FB46" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana qixingguan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure10" httpUri="https://binary.pensoft.net/fig/1161055">10 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="55C65A989FACE7B8F1222325E83C564A" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="FB655227490125EA24D9D72BF149AD4B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana qixingguan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure10" httpUri="https://binary.pensoft.net/fig/1161055">10 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="A8EDDDB98E2E101D13D7CBFA245299B0" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="4B5545EC1FB694323F079886965BB61A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana qixingguan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure10" httpUri="https://binary.pensoft.net/fig/1161055">10 B</figureCitation>
); procursus simple, with distal membranous process (arrow
<quantity id="21B87F5818B22FB495E5FD9DC4D7FDCA" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="DEFB04F64464CB49EABCB2C765D6AA11" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana qixingguan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure10" httpUri="https://binary.pensoft.net/fig/1161055">10 C</figureCitation>
,
<figureCitation id="76B0B4F0C3AB2E544746408F7ABA1E3A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 I</figureCitation>
) and subdistal spine (arrow
<quantity id="D38F6394FF685814DC307509A76B0C29" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="968AEAE639CF6446DDFCB1137E721BAC" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana qixingguan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal membranous process, arrow 2 points at subdistal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure10" httpUri="https://binary.pensoft.net/fig/1161055">10 C</figureCitation>
,
<figureCitation id="3900C71B5AB2806208801BB40C07C645" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 I</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EGDAI" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="12A0C7460100EB9A98AF96311EDB8E1E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="5D3E7CA56C9287E412FF6CDC6EC1E5D3" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 C</figureCitation>
). Retrolateral trichobothria on tibia I at 10 % proximally; legs with short vertical setae on metatarsi; tarsus I with 15 distinct pseudosegments.
</paragraph>
<paragraph id="1E2DF51AC4987A705A15B97DD92BC31A">
<emphasis id="4C6999DB9EBFA4F2E849BAC815034362" bold="true">Female</emphasis>
(
<emphasis id="176B06D1C976095EB4F6B8EDD25165F7" bold="true">
<typeStatus id="2CD5F86343914403520868EEF4DFEA90" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="092FD8F26341C7732A7C87684B283682" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="E30A3BFB3F7EF476B8CA29299BB04D1C" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45207): Similar to male, habitus as in Fig.
<figureCitation id="4C32BCA4B21216B63CAE1683B777DEDB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 G, H</figureCitation>
. Total length 2.04 (2.15 with clypeus), prosoma 0.78 long, 0.81 wide, opisthosoma 1.26 long, 1.00 wide; tibia I: 3.08; tibia I
<abbrev id="ABBRID0EAEAI" xlink_title="length/diameter">L / d</abbrev>
: 39. Eye interdistances and diameters:
<abbrev id="ABBRID0EEEAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EIEAI" xlink_title="posterior median eye">PME</abbrev>
0.12,
<abbrev id="ABBRID0EMEAI" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EQEAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EUEAI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.55 / 0.46. Epigyne simple and flat, posteriorly curved, with pair of postero-lateral pockets 0.28 apart (
<abbrev id="ABBRID0EYEAI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="BB52B97CB7407FB56C095B440CA0B671" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 A</figureCitation>
,
<figureCitation id="26D97FDAD7408522612EB724018AA1DB" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 I</figureCitation>
). Vulva with curved anterior arch (
<abbrev id="ABBRID0EEFAI" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="D904134F274EF3014EDAC30B95F85C10" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 B</figureCitation>
,
<figureCitation id="742229A65EE8A090C068678B9CA1B0EA" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 J</figureCitation>
), pair of medially narrow and posteriorly strongly widened pore plates (
<abbrev id="ABBRID0EQFAI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="FAE54DB2A961AFC47B368B83110E5F4D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 B</figureCitation>
,
<figureCitation id="9149D2505F031E462F348ABE8D476D8C" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 J</figureCitation>
), and pair of sclerotized protrusions (arrow in Figs
<figureCitation id="577A220FF56EA7388104B391B36E3732" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana qixingguan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sclerotized protrusion C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure11" httpUri="https://binary.pensoft.net/fig/1161056">11 B</figureCitation>
,
<figureCitation id="79F6751523CCF1F3F65CD02DFF85745D" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEGAI" type="variation">
<paragraph id="6D10F9A237A8C6346592B259E7EF6181">
<heading id="C252C41B751EE65E7CF5BD53E9D8EDCB" reason="title">Variation.</heading>
</paragraph>
<paragraph id="08C3C3CEC71B8D70FB86F09506178835">
Tibia I in the other
<specimenCount id="5DB6F860002BC6D7D751C155AB09CACC" count="6" type="female">six female</specimenCount>
<typeStatus id="4CB750A8FBA0366060701B21CD57C15C" type="paratype">paratypes</typeStatus>
(
<named-content id="0797C40B1C146D9F57DC031AC7074DA4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="D7161CC25DD572FCCDAE34AE37076921" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4520813): 2.893.09.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOGAI" type="habitat">
<paragraph id="4CF3DBA2AD9A07C09238EA09BA34B3BA">
<heading id="3E05799D899D04FA315A3A148BF660C0" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="2353D9A6B26CAD2AB4A1641CB696C7DD">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0ETGAI" type="distribution">
<paragraph id="3D02107FB6B2E311A29E0A05932AD5B9">
<heading id="44CEDB6BA5A7DF0C565F7FB7102BAB49" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="0352EDEF7C1D9E64C555575BBC2E1C53">
<collectingCountry id="57B3CC735D06CBEB5DB3F76C99DCAEB1" name="China">China</collectingCountry>
(
<collectingRegion id="16D9458FCE4DE621F611E97A3C61D111" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="E993E6668DE388845AF6F058DFD6E326">type</typeStatus>
locality; Fig.
<figureCitation id="EBBA728DB4C7DC0D9E58DFADBE0F0056" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4GAI" type="etymology">
<paragraph id="1FD8DA10042E9C0322E00216601CD9E2">
<heading id="812ECBA8C69EAC9242758AA61B4D96DA" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="098A2CABA530A753E5A6DFBB39B5B1EC">
The specific name refers to the
<typeStatus id="FBBACAD7A8794A24E54B33C472E69C44">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,427 @@
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<mods:title id="A908ED7DBD2BFC4DDD183020C9719533">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<mods:namePart id="DB243622B285324D34FF4845383D890E">Wang, Bing</mods:namePart>
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<mods:namePart id="A478AE13E5E678D24D10F87EC6B87B5F">Li, Jinglin</mods:namePart>
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<mods:affiliation id="1BAD83412FB3D1067EFE5019539581A3">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:namePart id="CEE0C4AF2EA28E2CBA2554B1870752A7">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="2F3E923B87A200DA519A2A56A5F73A23" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="E3DAA1DB2BC73A9E608433F817504476">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="9533AB1A38BF2F5445347A8B7070447E">Yao, Zhiyuan</mods:namePart>
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<treatment id="2266C4A0AF5E516FA4335C0A6547F024" ID-DOI="http://doi.org/10.5281/zenodo.13992772" ID-Zenodo-Dep="13992772" ID-arpha="2266C4A0-AF5E-516F-A433-5C0A6547F024" ID-zoobank="https://zoobank.org/D2E3F0DB-7736-46DA-ADC9-9F1FC1CF3ACD" LSID="urn:lsid:zoobank.org:act:D2E3F0DB-7736-46DA-ADC9-9F1FC1CF3ACD" LSID-ZBK="urn:lsid:zoobank.org:act:D2E3F0DB-7736-46DA-ADC9-9F1FC1CF3ACD" httpUri="http://treatment.plazi.org/id/2266C4A0AF5E516FA4335C0A6547F024">
<subSubSection id="19D47E6713B5703BD44B2632006C2977" type="nomenclature">
<paragraph id="1A7BCC9AB31959F0E4C3609247D02C51">
<taxonomicName id="ECEA6C3D8A177A893498C7402075D517" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhouxi" status="sp. nov.">
<emphasis id="C06F74C656D8736DFC5319AA9A22ADCE" italics="true">Belisana zhouxi</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="8FAE9EEDABA0D4369FF067FC02CD68D3" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="74FA357F7F0D1FD79D540559E984A70B" type="description">
<paragraph id="DB3DCC55B7CA72EC0C86666C1D4C6A83">
<figureCitation id="53EC31CAFDF9EFE54868EAE7DE0B208B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">Figs 18</figureCitation>
,
<figureCitation id="6F16F58340F5B146E25FD99980F2D42F" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19</figureCitation>
,
<figureCitation id="EE3AFDC177814088AF4D2E6336627FE6" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 E, F</figureCitation>
,
<figureCitation id="7642D0496E942C627F81AED315D2691F" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EONBI" type="type material">
<paragraph id="ABEFDAB3ABE0552AC3E8C8836C430DE7">
<heading id="68A125B4FD37065A6DAE6D6CF19FC497" reason="title">Type material.</heading>
</paragraph>
<paragraph id="60C84D77FD90F81727CD7D65531324AC">
<materialsCitation id="A3DF33489D30CC0950E6A4890FFBB274" collectingDate="2007-05-25" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Kaili" elevation="665" latitude="26.488" location="Hebian Cave" longLatPrecision="1" longitude="107.924034" municipality="Zhouxi Town" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="203A9A4F6DA65553A9EAFAD8874F445A" bold="true">
<emphasis id="D6A33275E0960533444E538AB9BB9F5C" bold="true" italics="true">
<typeStatus id="421B845DD0E1AFC4015833C15DF01FF0" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="B8A586CBD7C48325F155233A14E948A0" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="8670FCD0DB3D4E73FEB6FBE1B37FE794" count="1" type="male"></specimenCount>
;
<collectingRegion id="3120C6C829440D6F9D393857141D8137" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingCounty id="ED95508C779BA7F2EF6D70AF9F01D759">Kaili</collectingCounty>
,
<collectingMunicipality id="220A18E192FF56A436216547E5B2B246">Zhouxi Town</collectingMunicipality>
,
<location id="2F8B432866E49CA24F32B0DF5F13F733" LSID="urn:lsid:plazi:treatment:2266C4A0AF5E516FA4335C0A6547F024:2F8B432866E49CA24F32B0DF5F13F733" country="China" county="Kaili" latitude="26.488" longLatPrecision="1" longitude="107.924034" municipality="Zhouxi Town" name="Hebian Cave" stateProvince="Guizhou">Hebian Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3NBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.924033,26.488000]}">
<geoCoordinate id="B888FF1C74FFB7E06BD263AA81FDA7C6" degrees="26" direction="north" minutes="29.280" orientation="latitude" precision="1" value="26.488">26 ° 29.280 ' N</geoCoordinate>
,
<geoCoordinate id="F85E7C914793C4FECCE3F26E08694D9F" degrees="107" direction="east" minutes="55.442" orientation="longitude" precision="1" value="107.924034">107 ° 55.442 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="0DB7655BFFB64F9315907A1F14418F3C" metricMagnitude="2" metricUnit="m" metricValue="6.65" unit="m" value="665.0">
<elevation id="A99EB81376CF431D46EEE52D783236EA" metricMagnitude="2" metricUnit="m" metricValue="6.65" unit="m" value="665.0">665 m</elevation>
</quantity>
;
<date id="85D33FDEE592FEB9DB2957F7BD112804" value="2007-05-25">
<collectingDate id="08A046E38735087D5AA6BD82E44701EC" value="2007-05-25">25 May 2007</collectingDate>
</date>
;
<collectorName id="59AFE3C4495959760BDCAB4EA7F0039D">J. Liu</collectorName>
&amp;
<collectorName id="AB18548579ABC4ECB22398F8FD87CCC0">Y. Lin</collectorName>
leg.;
<named-content id="0E42DD3C34AE225C9320D7113DD009F1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="2BB78DB6CDCD67D9F94E2BEBB23F4F3C" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="A8514A4CC5E21CCBB522E58800BAD6C2">Ar</collectorName>
45226
</materialsCitation>
.
<materialsCitation id="9503A914EC241B9F10172D7562679D65" collectingDate="2007-05-25" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Kaili" elevation="665" latitude="26.488" location="Hebian Cave" longLatPrecision="1" longitude="107.924034" municipality="Zhouxi Town" specimenCount="2" specimenCount-female="2" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="4080F5B8F92D15E44999DACE30774B89" bold="true">
<emphasis id="97CDC75EA1933FD5BA5864182FC4C349" bold="true" italics="true">
<typeStatus id="F2F6376C2291D945188F471B14003AF3" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="E7715A7443F27EADFED0DF0590C44068" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="A26A5AF22F684EBB99F8DD5A932D9254" count="2" type="female">2 ♀</specimenCount>
; same data as for holotype;
<named-content id="DA4EF56F5B622C5ACC9F5C75A325D527" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="3D55ACFB3E2FBC1BD08F5A5D4ACB5DB8" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4522728
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENOBI" type="diagnosis">
<paragraph id="317D5B8B05EE2AF32643712C04361FDC">
<heading id="B7FEEB910DE880E95F18BD53B827C541" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="668F0DD80A81DA17E9EA1A8A5D421BB4">
The new species resembles
<taxonomicName id="0B322CD129AC56FE15FA4D490F6D73B7" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="majiang" status="sp. nov.">
<emphasis id="8DD56C0ECB70DF617B7EB0036FDD5926" italics="true">B. majiang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0D670F9F6051B4BDBFA4950FB6B0985" rank="species">sp. nov.</taxonomicNameLabel>
(Figs
<figureCitation id="D1A45E33D9FF1F103EB12FAFBF56F244" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6</figureCitation>
,
<figureCitation id="B227900B95B61FF58A42CD5817EF04A7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7</figureCitation>
,
<figureCitation id="5130AAA22C74898C60AD28E84AC18149" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E, F</figureCitation>
,
<figureCitation id="8773B7A80AFA2FC60F612A49998B2840" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 E, F</figureCitation>
) by having similar male chelicerae (tips of distal apophyses pointing downwards; Fig.
<figureCitation id="0EDEF18F32ACC2CF28CC0DFDE7B41B11" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 D</figureCitation>
), bulbal apophysis (hooked; Fig.
<figureCitation id="CFDCE4D3DC9C6FEB8D58A197BADEF599" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 C</figureCitation>
), and epigyne (epigynal pockets on antero-lateral part of epigynal plate, epigynal plate posteriorly curved; Figs
<figureCitation id="4D9801F90ACDC461470C924BDB1C37E3" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 A</figureCitation>
,
<figureCitation id="EADCB82351F21374646C8DD689103DFC" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 G</figureCitation>
), but can be distinguished by procursus without ventro-subdistal membranous process and dorso-distal spine (Figs
<figureCitation id="58456ADA60624AF1802F41216081119B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">18 C</figureCitation>
,
<figureCitation id="D73155784DC54DE4B1294DDCE9212FE2" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 E</figureCitation>
vs present) and by vulval pore plates elliptic (2 × longer than wide,
<abbrev id="ABBRID0EGQBI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="62509FE1CDF960EDB676E69F4C15FBEF" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 B</figureCitation>
,
<figureCitation id="86BAD81018AB2A49A0D047293E8C5A10" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 H</figureCitation>
vs 3 ×).
</paragraph>
<caption id="8A0FE059DFD46DCAF4ABF5F33514A6D3" ID-DOI="10.3897/zookeys.1216.132561.figure18" ID-arpha="0CE0D328-06A7-5E37-A984-FB9ECC9B71D0" httpUri="https://binary.pensoft.net/fig/1161063" startId="F18">
<paragraph id="753E5EC5B3E99B2674C98A4D89E2FAAE">
<label id="DC890672C3E95A4098D892250BAEF359">Figure 18.</label>
</paragraph>
<paragraph id="AD40EF11A5CDC90559D55BBE18792280">
<taxonomicName id="04218372FCF3DD8CE99226551CB41525" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhouxi" status="sp. nov.">
<emphasis id="EA3FF17CAF597FB9A725D483651B911B" italics="true">Belisana zhouxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FB370B70DC8BBB24F359E2ED615F7749" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="B55D017011FA9D7E6E85B4CFD34F62FB" bold="true">A, B</emphasis>
palp (
<emphasis id="31B65B4386DA311F21DF5D641D96E21A" bold="true">A</emphasis>
prolateral view
<emphasis id="5192672C2FAA7EA489501446C030E393" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="7C623368136BF83437F6FCBA69C26A1D" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="5AD4B21A5373BE22E173260B9F257DCB" bold="true">C</emphasis>
prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process
<emphasis id="F885CB70EE6DA05B52218568E1ECA39C" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb,
<abbrev id="ABBRID0ERRBI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0EVRBI" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.05 (
<emphasis id="939549D4FDD3DC94438512FB2C8F9420" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="6649CFDBBC111AA232521C8D80640035" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="7FC5339D68C6D591A694C3BF9DB766D1" ID-DOI="10.3897/zookeys.1216.132561.figure19" ID-arpha="3A77E230-1967-5EB5-80BE-3CFE006592CD" httpUri="https://binary.pensoft.net/fig/1161064" startId="F19">
<paragraph id="E9AD90BF165B27E5AF45325F234248E9">
<label id="08D272CEEC5EBE9082956D9825B6BC00">Figure 19.</label>
</paragraph>
<paragraph id="7C4D7B9D795F9295C24339F0B7E1AF3F">
<taxonomicName id="241BCE84EC267A4B9E614D2158AEB4B5" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhouxi" status="sp. nov.">
<emphasis id="F61E7BD374A567C9887F09996F279168" italics="true">Belisana zhouxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAF2605DA7B4FB1CD94FC0960BDCFC77" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="1A2E5232B39BA940326A22AF039B0185" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="DCDFDD6ED062307405DED8E8AF2E36A7" bold="true">A, B, G, H</emphasis>
)
<emphasis id="466DA4EE7F2B8DC8CAA52BDD8282FBFA" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="828A4C018E379BE831842C870BCDE5EE" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="7B3DBF4D964B677E5517F3C4CA232774" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="B4DD044F2823B9C0548099723BDF3376" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="9248E1F3F45AB8FCB8C36DC4785CFB94" bold="true">E H</emphasis>
habitus (
<emphasis id="E59F2D1A8135E5F4584CE00878F4E870" bold="true">E, G</emphasis>
dorsal view
<emphasis id="A812E33D725C3F28EB65B9151FCA8AAA" bold="true">F</emphasis>
lateral view
<emphasis id="3FFDB277ADDB526BA4E541AE7EC59154" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EJTBI" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0ENTBI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0ERTBI" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EVTBI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EZTBI" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0E4TBI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.05 (
<emphasis id="27C3B10C14A68B02DBE212CD99A98169" bold="true">A D</emphasis>
); 0.20 (
<emphasis id="08F9F1E3B0BD22C81033E0ABAB2455E4" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EKUBI" type="description">
<paragraph id="2AD26951C3B94F7B2DE4F1BC080F6488">
<heading id="092E770D4F5428ABF731D063D5BB499A" reason="title">Description.</heading>
</paragraph>
<paragraph id="01ACD6A7300A3055E1EBC858601CE29A">
<emphasis id="1AB2F78A0E1026A5FFBB4E4A584F18B8" bold="true">Male</emphasis>
(
<emphasis id="2EB9A6CFAD4C0CE4C6942255BDB3206F" bold="true">
<typeStatus id="F65201C3863AE22B62F1ED01FE20D43B" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.17 (1.27 with clypeus), prosoma 0.46 long, 0.50 wide, opisthosoma 0.71 long, 0.61 wide. Leg I: 6.72 (1.84, 0.21, 1.66, 2.13, 0.88), leg II: 4.56 (1.36, 0.19, 1.06, 1.36, 0.59), leg III: 3.29 (0.96, 0.19, 0.66, 1.01, 0.47), leg IV: 4.69 (1.44, 0.18, 1.16, 1.42, 0.49); tibia I
<abbrev id="ABBRID0EUUBI" xlink_title="length/diameter">L / d</abbrev>
: 27. Eye interdistances and diameters:
<abbrev id="ABBRID0EYUBI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E3UBI" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EAVBI" xlink_title="posterior median eye">PME</abbrev>
0.06,
<abbrev id="ABBRID0EEVBI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EIVBI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.41 / 0.33. Habitus as in Fig.
<figureCitation id="95FF33F2A87E27604D45FDFD11E29048" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, without marks; clypeus and sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0EQVBI" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="233065DC7BCC08952CAFE08346C37F00" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.19;
<abbrev id="ABBRID0EYVBI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="D1067554A454734303A0A848A4127480" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 D</figureCitation>
). Palp as in Fig.
<figureCitation id="0D7848B72A0F3D3884947E41F01436CA" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">18 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="54CE65C98381972F3A64ACC91A96E507" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="BE872033A04244E9F72B54B8C0237A65" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">18 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="3B8B7667D66AC75B45803A178C7AC267" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="B3D0AF8D30BA4F38734B6BAA4F78F5FD" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">18 B</figureCitation>
); procursus simple proximally but complex distally, with prolatero-distal membranous lamella (arrow
<quantity id="5E993643833CCB9F0AC0C76D846604A5" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="BA860EC19DF8BE12B12D49A1DB9067A6" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">18 C</figureCitation>
,
<figureCitation id="865B907D8C4B357CD9454DFFD1370301" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 E</figureCitation>
) bearing proximally slightly sclerotized part, dorso-subdistal membranous process (arrow
<quantity id="599CEFFFED08112DB0A2C680F10D24FD" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="FA5E6F35F3275990920AA7B6C68B943D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">18 C</figureCitation>
,
<figureCitation id="EE9FD87D68EDA3F1D8B67943AD7BD0BB" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 E</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="C640C410AF8357AF597261C2DBD38D5B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">18 D</figureCitation>
,
<figureCitation id="7BE417B6761A0804AA9C217E5995E065" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 F</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EEXBI" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="6CF6F62B755AAB81952B4C1583459175" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="40E20B4EC6D91F1701DCE1791830BE3E" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 C</figureCitation>
). Retrolateral trichobothria on tibia I at 7 % proximally; legs with short vertical setae on metatarsi; tarsus I with 14 distinct pseudosegments.
</paragraph>
<paragraph id="8E7E1D55D599AF8789423B3F182FD28E">
<emphasis id="3140D23B4141AD642B7325AC3F1C9391" bold="true">Female</emphasis>
(
<emphasis id="AF2EE089B8CF971A36673C3B9E29C966" bold="true">
<typeStatus id="0626209BDC5EE1547D00CFB45ADDA63B" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="2ADDB60F285D8C59F11BF639DEF8E1ED" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="78C29B802978A7F984B9AED0439F2994" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45227): Similar to male, habitus as in Fig.
<figureCitation id="FC632948950EE215D5A69F2F36280E44" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 G, H</figureCitation>
. Total length 1.21 (1.31 with clypeus), prosoma 0.49 long, 0.57 wide, opisthosoma 0.72 long, 0.63 wide; tibia I: 1.22; tibia I
<abbrev id="ABBRID0E5XBI" xlink_title="length/diameter">L / d</abbrev>
: 22. Eye interdistances and diameters:
<abbrev id="ABBRID0ECYBI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EGYBI" xlink_title="posterior median eye">PME</abbrev>
0.07,
<abbrev id="ABBRID0EKYBI" xlink_title="posterior median eye">PME</abbrev>
0.06,
<abbrev id="ABBRID0EOYBI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0ESYBI" xlink_title="anterior lateral eye">ALE</abbrev>
0.01. Sternum width / length: 0.43 / 0.32. Epigyne simple and flat, posteriorly curved, with pair of antero-lateral pockets 0.16 apart (
<abbrev id="ABBRID0EWYBI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="DDD6CC5C7B31DB92E87AE82307F5E399" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 A</figureCitation>
,
<figureCitation id="20ED3B3B3AD0C9A9546B3588D6C5CFCF" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 G</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0ECZBI" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="DEA61EF8561FBE6E671A41C7B82CEA04" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 B</figureCitation>
,
<figureCitation id="B7FE1D5F686618EC55DA4556FD3648FF" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 H</figureCitation>
) and pair of elliptic pore plates (2 × longer than wide,
<abbrev id="ABBRID0EOZBI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="E756AFEBDFFB1F6C7E14E6B8F34C44ED" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19 B</figureCitation>
,
<figureCitation id="DB1C300CEC5F2457A0E416E4C8866F9D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1ZBI" type="variation">
<paragraph id="203B864C182EA3DAB8F8EAAC67632A58">
<heading id="E3FFD952B79E68698F6300BD990ECE95" reason="title">Variation.</heading>
</paragraph>
<paragraph id="314973DFD27F0C91D5FE6594380E1420">
Unknown. Leg I missing in another female
<typeStatus id="0936CC474DD6EC06A6351D13F94EF957" type="paratype">paratype</typeStatus>
(
<named-content id="49F3FF13ECA32E661404C2E98398989B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="6E5D7F9EACB7A750DA53579A2691CBA0" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45228).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE1BI" type="habitat">
<paragraph id="87210767692DEEFAB4140CEC5CE49653">
<heading id="5ED5FC4C4DFE2FBCAB529D8024CED947" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="8A05D722C13D43303C1E45CA42ACB3B5">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ1BI" type="distribution">
<paragraph id="72902625E31885113D91E2887FC894B8">
<heading id="0442F85B8ADEFE8D02D6631A79C8301A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="340C7FE1955EA4DC6EE000AAB0B27888">
<collectingCountry id="C4E6F7EF291F1B1D8439ED9D0B9808B8" name="China">China</collectingCountry>
(
<collectingRegion id="EA72A96398C58F28CF56EE78F02B00C0" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="8B74623BC2F61919808B75FA3D4A7B19">type</typeStatus>
locality; Fig.
<figureCitation id="C92FA595A48557DBAB8956C2187EA75D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ET1BI" type="etymology">
<paragraph id="38ECC0F341A8B2DF181AA87460B7449E">
<heading id="300E740811F0FAF40D0ABC55D3870AE5" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="1FA16AEF974CEE7068E0A58ECF5FE459">
The specific name refers to the
<typeStatus id="6514A8F6AF06D38DFB63E1B21CDC48E5">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,290 @@
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<paragraph id="30B1081C903D332EDCBC121714FF9663">
<taxonomicName id="901B7F37B79D933112C42BD2EC0E4763" ID-CoL="5WGZ3" authority="Tong &amp; Li, 2007" authorityName="Tong &amp; Li" authorityYear="2007" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhangi">
<emphasis id="E973C50599D4887A9A63DC38D3B961EC" italics="true">Belisana zhangi</emphasis>
<bibRefCitation id="E0A80E7AFAEE275EFCBD0D72E7181550" author="Tong Y &amp; Li S" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="505-507" refId="B7" refString="Tong Y, Li S (2007) A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China. Acta Zootaxonomica Sinica 32: 505 - 507." title="A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China." volume="32" year="2007">Tong &amp; Li, 2007</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="800196E272E16B68456ECAE65551714B" type="description">
<paragraph id="057E95FF8E061003C7CB322AD9A57535">
<figureCitation id="29AE708AE285BB837495E18E82C7166D" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana zhangi Tong &amp; Li, 2007, male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure16" httpUri="https://binary.pensoft.net/fig/1161061">Figs 16</figureCitation>
,
<figureCitation id="4E3313DDC7216EFEA84C0E21CC7FDB9C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana zhangi Tong &amp; Li, 2007, male (C F) and female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure17" httpUri="https://binary.pensoft.net/fig/1161062">17</figureCitation>
,
<figureCitation id="F7FE31FE844C14B6FF5A7C9ECBB93C88" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 C, D</figureCitation>
,
<figureCitation id="B3443801EEFBADBA1025AF9F231F50B3" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="40C898A3BA125CA5CCBA40B9C59D86E9" type="reference_group">
<paragraph id="E08101DAC6D790C69584C974DF71903C">
<treatmentCitationGroup id="3CF573C8DB7A1E0F60703BA73666804D">
<treatmentCitation id="7EECCFABAB759932F358A86C9B783256" author="Tong Y &amp; Li S" page="505" year="2007">
<taxonomicName id="4A6DF5208350F0CC387E1ACDB2A25B6E" ID-CoL="5WGZ3" authority="Tong and Li 2007: 505" authorityName="Tong and Li" authorityPageNumber="505" authorityYear="2007" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhangi">
<emphasis id="78C3FD4A928AA409DDFE1FB63EDC8BF1" italics="true">Belisana zhangi</emphasis>
<comment id="7F5A1C8E38F1AA91B31F333069DB7202">
<bibRefCitation id="736BE57E15419FBD9E074B7BBE68BA98" author="Tong Y &amp; Li S" firstAuthor="Tong" journalOrPublisher="Acta Zootaxonomica Sinica" page="505" pagination="505-507" refId="B7" refString="Tong Y, Li S (2007) A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China. Acta Zootaxonomica Sinica 32: 505507." title="A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China." volume="32" year="2007">Tong and Li 2007: 505</bibRefCitation>
, figs 16.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQEBI" type="material">
<paragraph id="737B60B167168ED8B3647023BCC2922F">
<heading id="46FF8600AD348303E498FAD3EA34244A" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="74FEF72C87B10CE7B6D3CDC8DFD1CC47">
<materialsCitation id="248F3E11F56135394F5CE188341A4D72" collectingDate="2005-05-03" collectionCode="IZCAS" country="China" county="Puding County" latitude="26.24725" location="Huachu Town" longLatPrecision="1" longitude="105.62535" municipality="Anshun" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou">
<emphasis id="57146117AB7B9B758971FE724001C6A4" bold="true">
<collectingCountry id="9D8525E3FFED92796BB045DFEE33ED07" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="B6A86657733B78D576FB0FB5310F3958" count="1" type="male">1 ♂</specimenCount>
;
<collectingRegion id="D1F77919AF9396BE037C3C2393F559C6" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingMunicipality id="329905B7D77CF87DA5C4CF1FEEA24CE8">Anshun</collectingMunicipality>
,
<collectingCounty id="42338A4C4904B2C300A672ECBA194B66">Puding County</collectingCounty>
,
<location id="28CD508AA477ACA493DD0D14F7538F6D" LSID="urn:lsid:plazi:treatment:257E481B38D65DC5AAD15ABABE86F051:28CD508AA477ACA493DD0D14F7538F6D" country="China" county="Puding County" latitude="26.24725" longLatPrecision="1" longitude="105.62535" municipality="Anshun" name="Huachu Town" stateProvince="Guizhou">Huachu Town</location>
,
<location id="18B21BB5FF3F1CAD1715C44EE8019C76" LSID="urn:lsid:plazi:treatment:257E481B38D65DC5AAD15ABABE86F051:18B21BB5FF3F1CAD1715C44EE8019C76" country="China" county="Puding County" latitude="26.24725" longLatPrecision="1" longitude="105.62535" municipality="Anshun" name="Jinqian Cave" stateProvince="Guizhou">Jinqian Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2EBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[105.625350,26.247250]}">
<geoCoordinate id="964C2A91B5A282D41CA65F833DBCAF29" degrees="26" direction="north" minutes="14.835" orientation="latitude" precision="1" value="26.24725">26 ° 14.835 ' N</geoCoordinate>
,
<geoCoordinate id="DD58E1314636FE049501A66477CE26F3" degrees="105" direction="east" minutes="37.521" orientation="longitude" precision="1" value="105.62535">105 ° 37.521 ' E</geoCoordinate>
</named-content>
;
<date id="25A986FD91ED082C338D3DB943295A88" value="2005-05-03">
<collectingDate id="6E75B524423F1C09832772153FCAE381" value="2005-05-03">3 May 2005</collectingDate>
</date>
; collector unknown;
<named-content id="5233F0E27D2F544A91C0B1D5998E2358" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="9C6DB19E2283B259259B9401196170D6" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45177
</materialsCitation>
<materialsCitation id="7FCF0D1715787A07A0258337BD998C5F" collectingDate="2005-05-03" collectionCode="IZCAS" country="China" county="Puding County" latitude="26.24725" location="Huachu Town" longLatPrecision="1" longitude="105.62535" municipality="Anshun" specimenCount="3" specimenCount-female="3" stateProvince="Guizhou">
<specimenCount id="E6DB4CF6B3F91C2E2F83A2D32010B7A7" count="3" type="female">3 ♀</specimenCount>
; same data as for preceding;
<named-content id="068694B56457E2BC66DC25C7D54D656C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="CA4A6815AF22ED9B61D5EEDED3A81E2B" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4517880
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIFBI" type="diagnosis">
<paragraph id="3096124DF2308B9920E770F3229C1C3D">
<heading id="2692A94A3B249F6A2E13D88D4C85AC13" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="675F08EAC28CB5F32A3972AA00909D23">
The species resembles
<taxonomicName id="FCC09BF07E26D71F763A283CBDAF7E74" authority="Yang &amp; Yao, 2023" authorityName="Yang &amp; Yao" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yuhaoi">
<emphasis id="5FF1538B4CDF271B88696FAB4EF1EC97" italics="true">B. yuhaoi</emphasis>
<bibRefCitation id="295653DE8A44641F3DDD4267331DD6D3" DOI="10.3897/zookeys.1186.105736" author="Yang L &amp; Zhao F &amp; He Q &amp; Yao Z" journalOrPublisher="ZooKeys" pagination="175-184" refId="B10" refString="Yang L, Zhao F, He Q, Yao Z (2023 a) A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China. ZooKeys 1186: 175 - 184. https: // doi. org / 10.3897 / zookeys. 1186.105736" title="A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China." volume="1186" year="2023">Yang &amp; Yao, 2023</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="15C2EEBA3C0506D593B11E2999F70B04" DOI="10.3897/zookeys.1186.105736" author="Yang L &amp; Zhao F &amp; He Q &amp; Yao Z" journalOrPublisher="ZooKeys" pagination="175-184" refId="B10" refString="Yang L, Zhao F, He Q, Yao Z (2023 a) A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China. ZooKeys 1186: 175184. https://doi.org/10.3897/zookeys.1186.105736" title="A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China." volume="1186">Yang et al. 2023 a</bibRefCitation>
: 178, figs 2 A, B, 3 A D, 4 A H) by having similar bulbal apophysis (hooked; Fig.
<figureCitation id="41B2EC2AA9230E19DE5DAF046BCBEE60" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana zhangi Tong &amp; Li, 2007, male (C F) and female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure17" httpUri="https://binary.pensoft.net/fig/1161062">17 C</figureCitation>
) and epigyne (epigynal pockets on posterior part of epigynal plate, epigynal plate posteriorly curved; Figs
<figureCitation id="A4D1BFC911EBFC59FA68FE3830508718" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana zhangi Tong &amp; Li, 2007, male (C F) and female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure17" httpUri="https://binary.pensoft.net/fig/1161062">17 A</figureCitation>
,
<figureCitation id="D034FB1FBCCFFA73F008F84C9C91E081" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 E</figureCitation>
), but can be distinguished by procursus with nearly angular ventro-subdistal membranous lamella (arrow
<quantity id="F326370F5527D410BBDDBBEC41E95A19" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="C39E944F359DED599E31E5F125A7EBFF" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana zhangi Tong &amp; Li, 2007, male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure16" httpUri="https://binary.pensoft.net/fig/1161061">16 C</figureCitation>
,
<figureCitation id="4F8754F6907043C5316457310E6C8ED1" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 C</figureCitation>
vs nearly round) and curved retrolateral membranous flap (f in Figs
<figureCitation id="3A5905FDDAEB5228FD4E14A70CBBE3FA" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana zhangi Tong &amp; Li, 2007, male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure16" httpUri="https://binary.pensoft.net/fig/1161061">16 D</figureCitation>
,
<figureCitation id="F177FD02C8657BD1F6FD9FA4F9AFE5DD" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 D</figureCitation>
vs angular), by male cheliceral distal apophyses pointing inwards (
<abbrev id="ABBRID0EZGBI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="BA738202DF1F82883B814071D4CEE5BD" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana zhangi Tong &amp; Li, 2007, male (C F) and female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure17" httpUri="https://binary.pensoft.net/fig/1161062">17 D</figureCitation>
vs downwards), and by vulval pore plates long elliptic (
<abbrev id="ABBRID0EBHBI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="6C8CFE7BC8FCF964C8921B50E42B886F" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana zhangi Tong &amp; Li, 2007, male (C F) and female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure17" httpUri="https://binary.pensoft.net/fig/1161062">17 B</figureCitation>
,
<figureCitation id="FA02A62CFB0A03C1F9ADEB9F86F8E939" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 F</figureCitation>
vs nearly triangular).
</paragraph>
<caption id="F1CD1F032DD2125B79ABF5AE4892CE6E" ID-DOI="10.3897/zookeys.1216.132561.figure16" ID-arpha="82CD42A0-43E5-5F7A-B665-839CF4B66E6E" httpUri="https://binary.pensoft.net/fig/1161061" startId="F16">
<paragraph id="70D9509CA55B00847D979F4D3CC446F5">
<label id="9E6BFF529C8F2B7BC39006781F78AEE6">Figure 16.</label>
</paragraph>
<paragraph id="DDFD6851B5939E122B9F4F35E4554125">
<taxonomicName id="47C2C5D0BC67A24E10D3CCADE2C005C7" authority="Tong &amp; Li, 2007" authorityName="Tong &amp; Li" authorityYear="2007" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhangi">
<emphasis id="CF1CE060AF8066BAC84C586FD1DA61FE" italics="true">Belisana zhangi</emphasis>
<bibRefCitation id="3A81689A09F2697EDD5C4055F24F3CA4" author="Tong Y &amp; Li S" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="505-507" refId="B7" refString="Tong Y, Li S (2007) A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China. Acta Zootaxonomica Sinica 32: 505 - 507." title="A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China." volume="32" year="2007">Tong &amp; Li, 2007</bibRefCitation>
</taxonomicName>
, male
<emphasis id="73A9B853AD5BE9748672747716424EC4" bold="true">A, B</emphasis>
palp (
<emphasis id="478E75E43F21B9EFCD106237EF891A1D" bold="true">A</emphasis>
prolateral view
<emphasis id="61D5352F67EE2619DFC3512F081FC4CF" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="3480E419CFA500F06CB9BD62B517C417" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="B97755BB88AD991DA5D37C6437EE0339" bold="true">C</emphasis>
prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella
<emphasis id="C35650159BE1C5C3124C98C1CB8AFBB0" bold="true">D</emphasis>
retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine). Abbreviations: b = bulb,
<abbrev id="ABBRID0EMIBI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0EQIBI" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="E602A249D50C78EB08AF76E5CC5408D5" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="03239834B74B27807E832C8672E316B7" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="057F5A5E9B1EA66F8393B83AC4F31CC3" ID-DOI="10.3897/zookeys.1216.132561.figure17" ID-arpha="6344F0F5-EACA-5BB1-9AEE-8459AB72CD48" httpUri="https://binary.pensoft.net/fig/1161062" startId="F17">
<paragraph id="9F34C127F38B0F4EB528B98CF1E9BBFD">
<label id="4D2B27162554CF1B085F07220DE6AF1A">Figure 17.</label>
</paragraph>
<paragraph id="D474C4ED8860437E8BD4D0584810ED71">
<taxonomicName id="5296E7C69211EC774F2765AA410EF441" authority="Tong &amp; Li, 2007" authorityName="Tong &amp; Li" authorityYear="2007" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhangi">
<emphasis id="3F6FCC19535ADC31D4CA3F79FDB46632" italics="true">Belisana zhangi</emphasis>
<bibRefCitation id="854B105EB4A07814929CC3A0695E31CF" author="Tong Y &amp; Li S" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="505-507" refId="B7" refString="Tong Y, Li S (2007) A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China. Acta Zootaxonomica Sinica 32: 505 - 507." title="A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China." volume="32" year="2007">Tong &amp; Li, 2007</bibRefCitation>
</taxonomicName>
, male (
<emphasis id="79491A1A237EB0A977B0F44AAA421CD2" bold="true">C F</emphasis>
) and female (
<emphasis id="6655798682453A25E80F7ACE7588810E" bold="true">A, B, G, H</emphasis>
)
<emphasis id="DAB7AFC22161D86D3EE9EAD7BCE90941" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="5E6FAD97B189A3CAE8D30774D0591CB3" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="11C23F752CBDC26BB91A2C9ED192CACE" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="1D99687F9668784BD2032FFB0BC50764" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="C956BA5A8B2AD049BF5501391B3103C0" bold="true">E H</emphasis>
habitus (
<emphasis id="B21BCF5F01A82D5DC5E7DF5C44AECD27" bold="true">E, G</emphasis>
dorsal view
<emphasis id="174479BDD06C8A7B66B495D4846F1846" bold="true">F</emphasis>
lateral view
<emphasis id="8887D21D6FC54F460DAFE33988EA8840" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EEKBI" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EIKBI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EMKBI" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EQKBI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EUKBI" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0EYKBI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 (
<emphasis id="C233A78A461EAFB51E14C1B5BA2BD6B2" bold="true">A D</emphasis>
); 0.40 (
<emphasis id="94BDA7C5462AA9A97C225469AF142503" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EFLBI" type="habitat">
<paragraph id="1AA6D220CCE0EC42980D2B4A2EB50D31">
<heading id="BD80035216B861C6EB459ED0C221C220" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="DD5F35B5BBB09F0C28B7808B9101773E">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EKLBI" type="distribution">
<paragraph id="D6DC16841A77E40C21E822060A3F9201">
<heading id="6D4AF32404FA8F5A1F5C92F6ACF4299F" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="AB1425A4B9338312ECF70AEBA6496A25">
<collectingCountry id="412B7CB3BF3FF4BB3C759E8B9259B2CA" name="China">China</collectingCountry>
(
<collectingRegion id="089E95A37479C088E498DB44C83A05D2" country="China" name="Guangxi">Guangxi</collectingRegion>
,
<typeStatus id="79A5BFE1576E41687C64A9B939497C38">type</typeStatus>
locality;
<collectingRegion id="D741AE44569FADA3076E855682E4B429" country="China" name="Guizhou">Guizhou</collectingRegion>
, Fig.
<figureCitation id="7497D366FB017155FBDA73874D85ADBC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,602 @@
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<mods:title id="97EFB830BEDE07DBCE6C1CC2C7814EBA">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<subSubSection id="77CE9241A469B6548F21A6329B041005" type="nomenclature">
<paragraph id="3A54A5EF8F29858EFF36C000819152A1">
<taxonomicName id="992BFB6087F41037081F993C07A1789B" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zunyi" status="sp. nov.">
<emphasis id="DF6CB0201820BB70ADBA0EA3914D0940" italics="true">Belisana zunyi</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="EDCD33249BE9D08B531F3CEEFDE26A81" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2021C1F08DED31D11CF25FC3309D5CFC" type="description">
<paragraph id="50A8459BDE969B192605941CA263B902">
<figureCitation id="8DD0EDCEEF1878590D50224E6CC7D2A0" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">Figs 20</figureCitation>
,
<figureCitation id="2AD6D109C7DE418CD96669E8AFB6E5A8" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21</figureCitation>
,
<figureCitation id="C336247E2657C0B8C7834B69A20AD6C5" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 G, H</figureCitation>
,
<figureCitation id="26F59929D2B3F906C12E43A9204C8A2B" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 I, J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES3BI" type="type material">
<paragraph id="FE95CD31E9AD88FEDA5D8A8BFC97D73A">
<heading id="914750C39498083DA455953370CF4B05" reason="title">Type material.</heading>
</paragraph>
<paragraph id="D876F550196EA6FD45042D8337A1AF72">
<materialsCitation id="05ECAD79093612671F6A5F05501BA7C9" collectingDate="2007-05-11" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Tongzi County" elevation="1008" latitude="28.144867" location="Leishanguan Town" longLatPrecision="1" longitude="106.789635" municipality="Zunyi" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="375D4FDB7E107A14BE4EDDC02E15B649" bold="true">
<emphasis id="819BE060A6C375F4F07A5B5FA8207FBE" bold="true" italics="true">
<typeStatus id="5D567B0FAEC4A744C3AF2D1DCC60A59E" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="1D5D59EB31D4AE55406D48084F5322C7" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="5DFFB0EE4163B7BC2B992D23A34844E2" count="1" type="male"></specimenCount>
;
<collectingRegion id="EB76C86145C8638540FD74865FB24017" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingMunicipality id="B04008D5425FBE0B830BF18F5948F7A1">Zunyi</collectingMunicipality>
,
<collectingCounty id="696C10AB02DB95E804F7F308EE8C99F5">Tongzi County</collectingCounty>
,
<location id="B895465072693E6F0A1B021EF1BF81E8" LSID="urn:lsid:plazi:treatment:2EC26726DC0E5E33A3A6A4ED91089BAC:B895465072693E6F0A1B021EF1BF81E8" country="China" county="Tongzi County" latitude="28.144867" longLatPrecision="1" longitude="106.789635" municipality="Zunyi" name="Leishanguan Town" stateProvince="Guizhou">Leishanguan Town</location>
,
<location id="A3841EE4931088C0A9B240B651EAA902" LSID="urn:lsid:plazi:treatment:2EC26726DC0E5E33A3A6A4ED91089BAC:A3841EE4931088C0A9B240B651EAA902" country="China" county="Tongzi County" latitude="28.144867" longLatPrecision="1" longitude="106.789635" municipality="Zunyi" name="Yangliuping Village" stateProvince="Guizhou">Yangliuping Village</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EA4BI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.789633,28.144867]}">
<geoCoordinate id="DB48419EAB1A45D4043455D197174A92" degrees="28" direction="north" minutes="08.692" orientation="latitude" precision="1" value="28.144867">28 ° 08.692 ' N</geoCoordinate>
,
<geoCoordinate id="EB48D43F1C5AE5F406E87540B714B09F" degrees="106" direction="east" minutes="47.378" orientation="longitude" precision="1" value="106.789635">106 ° 47.378 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="A11F0959001FCA6864D3670FEEF11114" metricMagnitude="3" metricUnit="m" metricValue="1.008" unit="m" value="1008.0">
<elevation id="468FB5A412784E5E6745F3753D9F144F" metricMagnitude="3" metricUnit="m" metricValue="1.008" unit="m" value="1008.0">1008 m</elevation>
</quantity>
;
<date id="6C593BCC3F57C06AB2DA71DF7A49E2B6" value="2007-05-11">
<collectingDate id="FBBEFF4A6CF6E95FA17ECC32B116B130" value="2007-05-11">11 May 2007</collectingDate>
</date>
;
<collectorName id="6CDC223B20461A97DA53C17D8BA6663D">J. Liu</collectorName>
&amp;
<collectorName id="0F62998D9E6883434EC693F1C6C27560">Y. Lin</collectorName>
leg.;
<named-content id="841D9D7F7F44AFF0DAF1A166C5473789" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="23FE2874DF5661216AC86B738CC6524D" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="3A757C8A8F99761D3931029C6D0BBA40">Ar</collectorName>
45229
</materialsCitation>
.
<materialsCitation id="10301A45861915565ECF6961764D5956" collectingDate="2007-05-11" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Tongzi County" elevation="1008" latitude="28.144867" location="Leishanguan Town" longLatPrecision="1" longitude="106.789635" municipality="Zunyi" specimenCount="4" specimenCount-male="4" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="0DC57CF744D48D818D54B68B9FA483FB" bold="true">
<emphasis id="E1F9F73F8ECCCD794A4716779F4BAFF3" bold="true" italics="true">
<typeStatus id="1B4963381538C5E12ADB04A39232B34B" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="80D65C1FDC04EEC6BEFA3F6C7B7FCD11" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="2613469EB51E954F020D4E9A203E8DA3" count="4" type="male">4 ♂</specimenCount>
; same data as for holotype;
<named-content id="4ACFC165323BB96722BC4C86A0671889" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="7001F097A899C552DE375B151331E1C7" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4523033
</materialsCitation>
<materialsCitation id="9FBFC442EDEA166C5B88E183F24FD167" collectingDate="2007-05-11" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Tongzi County" elevation="1008" latitude="28.144867" location="Leishanguan Town" longLatPrecision="1" longitude="106.789635" municipality="Zunyi" specimenCount="13" specimenCount-female="13" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="544C8D971FFB7C418CA346F4D5FC99FD" count="13" type="female">13 ♀</specimenCount>
; same data as for holotype;
<named-content id="CD4FBFE70CC9920AB9637ABEF1A66261" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="2527998E3CB98A7A84DDA724F19323DA" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4523446
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EV4BI" type="diagnosis">
<paragraph id="5C0B91CDB00FBC2FCEA92C690B28A100">
<heading id="AF2A31A0568EEBF2215C902EFF06545C" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="320E4B34A32573364CC368C423916C0B">
The new species resembles
<taxonomicName id="6B735C7C66FDC96BE5F1C5A3303C38DA" authority="Tong &amp; Li, 2007" authorityName="Tong &amp; Li" authorityYear="2007" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhangi">
<emphasis id="660B97D88C148824094722D4DDC22FD1" italics="true">B. zhangi</emphasis>
<bibRefCitation id="53587EEA254FC23C7B9CC7A02D02AC25" author="Tong Y &amp; Li S" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="505-507" refId="B7" refString="Tong Y, Li S (2007) A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China. Acta Zootaxonomica Sinica 32: 505 - 507." title="A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China." volume="32" year="2007">Tong &amp; Li, 2007</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="AEF4F4E27ADC8B2825EFC06563A03569" author="Tong Y &amp; Li S" firstAuthor="Tong" journalOrPublisher="Acta Zootaxonomica Sinica" page="505" pagination="505-507" refId="B7" refString="Tong Y, Li S (2007) A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China. Acta Zootaxonomica Sinica 32: 505507." title="A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China." volume="32" year="2007">Tong and Li 2007: 505</bibRefCitation>
, figs 16) by having similar bulbal apophysis (hooked; Fig.
<figureCitation id="B27ECE5532C2981D17884D93A26DE23E" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 C</figureCitation>
) and vulva (anterior arch ridge-shaped, pore plates long elliptic and 6 × longer than wide; Figs
<figureCitation id="98F46345A521945F73C72278653E7CFB" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 B</figureCitation>
,
<figureCitation id="A87BFEEB78B39B51446646B767DCDD64" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 J</figureCitation>
), but can be distinguished by procursus with dorso-subdistal membranous process (arrow
<quantity id="99A7F0982638AFA46F0F3D665E51E64E" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="0B7F836393F781D8E0E183B1F1D7076D" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 C</figureCitation>
,
<figureCitation id="505D175532F51163383A48C63DF9C91B" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 G</figureCitation>
vs sclerotized apophysis) and without ventro-subdistal membranous lamella (Figs
<figureCitation id="4585DBD6C8507A492970B66EA02A1D31" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 C</figureCitation>
,
<figureCitation id="55825F626F6A0A483CC1BD74C65A0F36" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 G</figureCitation>
vs present), by tips of male cheliceral distal apophyses close to each other (
<abbrev id="ABBRID0EG6BI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="30BA66E521881FB9E3F046E949D03F08" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 D</figureCitation>
vs widely separated), and by epigyne with median pockets (
<abbrev id="ABBRID0EO6BI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="E7ADD21E753C71A3ABEBDE288CE9DB73" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 A</figureCitation>
,
<figureCitation id="48F944AF066CECBF6480EA587145BF5D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 I</figureCitation>
vs postero-median).
</paragraph>
<caption id="51AE15F9DBAAC51E09C880680D763633" ID-DOI="10.3897/zookeys.1216.132561.figure20" ID-arpha="0CFD1263-2751-584F-A5B6-76D2C4403367" httpUri="https://binary.pensoft.net/fig/1161065" startId="F20">
<paragraph id="07AFA81D2227FFCD8D54D11CE8199233">
<label id="CA873071FA7ACDE25870326427705E39">Figure 20.</label>
</paragraph>
<paragraph id="B0A9F9DB36622E9B8D72A6A1BB6F9F24">
<taxonomicName id="94E7460992A25E149D428CC0A70EE030" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zunyi" status="sp. nov.">
<emphasis id="F019E10242C1770B00C3F62DF782ABAE" italics="true">Belisana zunyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2FA1EE442F1400977B1CC4EABDB86F6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="B673A156FE0364F7C22106E000EF4E1E" bold="true">A, B</emphasis>
palp (
<emphasis id="F72D5D0899F0394C76B307289294AE53" bold="true">A</emphasis>
prolateral view
<emphasis id="AD63A13BC02F69065060B270EAFE1980" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="30EB7A19D155DFEF1EDECAF783E8A436" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="4E6857658CF5EEAC25E49CE485813ABF" bold="true">C</emphasis>
prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process
<emphasis id="DA04F58AFC357E71624894983453A558" bold="true">D</emphasis>
retrolateral view). Abbreviations: f = flap,
<abbrev id="ABBRID0EZACI" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="301F6FACBFC51CF6D9DD720458874BCF" bold="true">A, B</emphasis>
); 0.03 (
<emphasis id="F6B192802331D7717B10822CCFFD45A7" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="29074BDFFD7C24C233458352C494C79B" ID-DOI="10.3897/zookeys.1216.132561.figure21" ID-arpha="3544F1AC-1033-5B2D-A3F9-B1582ED10C1E" httpUri="https://binary.pensoft.net/fig/1161066" startId="F21">
<paragraph id="E928D3B000858838D7908625E3592BA4">
<label id="AD412DDC59E265B3543552F3784E5165">Figure 21.</label>
</paragraph>
<paragraph id="0DBD6A7BE79AAA404DDC7237167A526E">
<taxonomicName id="5507407EAD29C91A626F7B749BE221CE" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zunyi" status="sp. nov.">
<emphasis id="9A66382BBA56371A386D572FD9FDDFF0" italics="true">Belisana zunyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A7A8D530426DC8C26AA156466912BC5" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="4B5F566FBF3ECFE38772EC8E81F67CA1" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="0136F374DC704B5A36E65A4C42C0C65D" bold="true">A, B, G, H</emphasis>
)
<emphasis id="923AF8F42653171E449AE3400C60DDBE" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="6068E753D4DBA9B31A3595B88B87BBE6" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="6D08C3D858C90B7207516B5B1935DF69" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="B3589DC5BA2B407D89E9A697DEFF0F54" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="60E34178E26032935C4DEA7659A14C5A" bold="true">E H</emphasis>
habitus (
<emphasis id="07C8B00F0DF19D71817AA772496BE883" bold="true">E, G</emphasis>
dorsal view
<emphasis id="C682085FF3A1D0872B1C418AC1A7BBBE" bold="true">F</emphasis>
lateral view
<emphasis id="79388464264B808A58EB0E561740EC63" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0ENCCI" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0ERCCI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EVCCI" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EZCCI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0E4CCI" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0EBDCI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 (
<emphasis id="7785E9CF6BB26DF7AFEBAA2C203DE937" bold="true">A D</emphasis>
); 0.30 (
<emphasis id="1F2541C0767E80331038C3FA190D36B7" bold="true">E H</emphasis>
).
</paragraph>
</caption>
<caption id="6CA8FB98EDD34A4F491EFB8C21D685AF" ID-DOI="10.3897/zookeys.1216.132561.figure22" ID-arpha="C680BFAB-A54E-5B2F-9269-CA347C546826" httpUri="https://binary.pensoft.net/fig/1161067" startId="F22">
<paragraph id="25612CC47D600EB823E0A10308A87516">
<label id="7DD787DE858A2888BE8FF3C7591A4061">Figure 22.</label>
</paragraph>
<paragraph id="3A8F438E2C509D8DB19F3F7A8509026E">
Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species)
<emphasis id="8B9B3ED291B547DFE56CB134835F8F8F" bold="true">A, B</emphasis>
<taxonomicName id="504BEE1EC558F76731421688F2B8F3F5" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bijie" status="sp. nov.">
<emphasis id="7122B0EE7528C125BBA0C038C1FD4E50" italics="true">Belisana bijie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4BDB0EA3F84A2E681E362F8086F19CB" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="272DB13357DDBE314C5AF614AD4E2ED7" bold="true">C, D</emphasis>
<taxonomicName id="2A56C34780F84297E894FAA6015A0464" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="liupanshui" status="sp. nov.">
<emphasis id="B03488AC3081BE2462550286A169DFEF" italics="true">B. liupanshui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A56B79AF0C3B7420C28B20BA85913C98" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="FB7A58D9C9FAF955EFCB70D976AD9EB8" bold="true">E, F</emphasis>
<taxonomicName id="4C5F5B4076733DE86742C942F75908F5" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="majiang" status="sp. nov.">
<emphasis id="93675FAD168B23ED418A6F50F5E46084" italics="true">B. majiang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="23E6DC90E6F6CEC6C5850EBC34598E62" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="A6DD73CD44C1A8FE92DFC77A9E6EEDFE" bold="true">G, H</emphasis>
<taxonomicName id="9FBB383677697DDEFC2740970652800D" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nayong" status="sp. nov.">
<emphasis id="3B6DEA296B624E5D248DD9D160133642" italics="true">B. nayong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF118E670FE457A3E1F734FA6031C24A" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F3EDAE376F5064F16E4C7A53D77E7496" bold="true">I, J</emphasis>
<taxonomicName id="97BBA4A95197D6BF427B1FF2FA2D4381" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="qixingguan" status="sp. nov.">
<emphasis id="5892A7462DE370EC4E8F621B249F86BB" italics="true">B. qixingguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5387B467702E034EB62E89DD34167524" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="0C9A3B81EE848F0A05C6E7D5761858A3" bold="true">K, L</emphasis>
<taxonomicName id="ED0B74AE9280AB2D7CE6541E676B8940" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="xiuwen" status="sp. nov.">
<emphasis id="7B88BD5D0CD243AD5528E0351B1C1B8D" italics="true">B. xiuwen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E892E5D33794F6F828E9F0246727D052" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviation: f = flap. Scale bars: 0.10.
</paragraph>
</caption>
<caption id="9F948EC86416C38A332E0F3EFD4F9D4B" ID-DOI="10.3897/zookeys.1216.132561.figure23" ID-arpha="28FFF4C8-FB0E-57DD-8A0C-948DAB19E5F1" httpUri="https://binary.pensoft.net/fig/1161068" startId="F23">
<paragraph id="22AC45082EAB0F39286B6C9FE9331D08">
<label id="8C367B2CFBDCD355DFF78965F191AA79">Figure 23.</label>
</paragraph>
<paragraph id="1A087E00363AE3B1885B259A6C8DAEF3">
Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species)
<emphasis id="7D3435BD42B1054FBD89B18E5142FA5B" bold="true">A, B</emphasis>
<taxonomicName id="A9ED41A293D469BC4A4DB2DA9D2E71C3" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="588FF30D11DB26233D04F79D807244B1" italics="true">Belisana yongcong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3BC9E361D76448413B54E96EA3770C7E" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="FC6025AF3DA9E9DB27F66DC19C9868C5" bold="true">C, D</emphasis>
<taxonomicName id="0E57FD420D1C248709DD060066989705" authority="Tong &amp; Li, 2007" authorityName="Tong &amp; Li" authorityYear="2007" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhangi">
<emphasis id="6D749ECF7DD83970B5F02DD8A89A0D6C" italics="true">B. zhangi</emphasis>
<bibRefCitation id="4BA371DEE32FD07474E71341608AFF08" author="Tong Y &amp; Li S" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="505-507" refId="B7" refString="Tong Y, Li S (2007) A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China. Acta Zootaxonomica Sinica 32: 505 - 507." title="A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China." volume="32" year="2007">Tong &amp; Li, 2007</bibRefCitation>
</taxonomicName>
<emphasis id="2DC865A058BC4926889291CED1668EDB" bold="true">E, F</emphasis>
<taxonomicName id="3BC1DEF9DA48DFEA24060076EF56D5BA" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhouxi" status="sp. nov.">
<emphasis id="9201ADA40B27185A22C554D152191E6C" italics="true">B. zhouxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F931EA401CEF9A18B67FCFA2055D7011" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="EA4E348BDE21A0DF8B91A80E25B192BE" bold="true">G, H</emphasis>
<taxonomicName id="4ED5286C9FF8477DA785261F40A3D7E2" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zunyi" status="sp. nov.">
<emphasis id="6ACE55D0E5F7064882B0AEFAB1A6E994" italics="true">B. zunyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33DE3DDA22258F9AC9407FF2D093AD83" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviation: f = flap. Scale bars: 0.10.
</paragraph>
</caption>
<caption id="D323BE9D9F9AB8836FF6031409A5787E" ID-DOI="10.3897/zookeys.1216.132561.figure24" ID-arpha="0CD23808-6495-57CE-817A-84581C1499A1" httpUri="https://binary.pensoft.net/fig/1161069" startId="F24">
<paragraph id="FDA5B47619B7EAB53B41B7DAD5C3C302">
<label id="96944E87FADE6179BE6DAF56B7A06604">Figure 24.</label>
</paragraph>
<paragraph id="E923D593F668277453B7E5E81F6BCA44">
Female genitalia in ventral and dorsal views
<emphasis id="803A0335E392D2BABC8A74C600EE8EE2" bold="true">A, B</emphasis>
<taxonomicName id="AA7325C48920DD16D6C112D50D3596DD" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="bijie" status="sp. nov.">
<emphasis id="917B22EAE94EC6E9DE51AD972976E158" italics="true">Belisana bijie</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D461E314C23899786A2363EB284652B1" rank="species">sp. nov.</taxonomicNameLabel>
, arrow points at lateral sclerite
<emphasis id="340A93739B3E81A88D095B130A08CD37" bold="true">C, D</emphasis>
<taxonomicName id="B955AD2DBB75418512DEA76E3EA8E0A0" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="liupanshui" status="sp. nov.">
<emphasis id="1001B66FB1BFCE3D1E9E6EF6AC536A9A" italics="true">B. liupanshui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0DE33279AFB8AC4F1DA66922DD58DA93" rank="species">sp. nov.</taxonomicNameLabel>
, arrow points at lateral sclerite
<emphasis id="BFDCC844C3DF60672BD67BDA1C854756" bold="true">E, F</emphasis>
<taxonomicName id="908F21A1FCBD89DD62FDBF83204B296E" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="majiang" status="sp. nov.">
<emphasis id="299FCDDE3EAF12C5FC237D6817CA31D2" italics="true">B. majiang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30C4A1F4553006EB91B1FC138932F977" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="AE4ECC7CA52FA1913FA10034050FF29E" bold="true">G, H</emphasis>
<taxonomicName id="379ECFB82538722F87ACB25CAE344ABB" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nayong" status="sp. nov">
<emphasis id="632E7A79D82570E68A27DC17D59FCEE0" italics="true">B. nayong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1E7BCFBE0427F72FD0966DB8A4864F07" rank="species">sp. nov</taxonomicNameLabel>
<emphasis id="DF7506A14E4EA5FC49DCA4B3217322E7" bold="true">I, J</emphasis>
<taxonomicName id="804F86BE3C93FE48C07E3CFC9EE92170" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="qixingguan" status="sp. nov.">
<emphasis id="4929004556026ED615311F041EAC8AF8" italics="true">B. qixingguan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A151079F56D0BA9D2E953C920BBC515B" rank="species">sp. nov.</taxonomicNameLabel>
, arrow points at sclerotized protrusion. Abbreviations:
<abbrev id="ABBRID0ETKCI" xlink_title="anterior arch">aa</abbrev>
= anterior arch,
<abbrev id="ABBRID0EXKCI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0E2KCI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10.
</paragraph>
</caption>
<caption id="D6472C7F18F8633CC8EA9865D7DFF702" ID-DOI="10.3897/zookeys.1216.132561.figure25" ID-arpha="0C3BCBB5-1366-5FAD-8E74-FF7C0410794E" httpUri="https://binary.pensoft.net/fig/1161070" startId="F25">
<paragraph id="F5DBB44BD0EFC6D16E9E1688DB14A3B2">
<label id="577F92B17A9EC393273CC23003FF63AF">Figure 25.</label>
</paragraph>
<paragraph id="D9E3F47EA3EF2CBE74DB828BAA1F4717">
Female genitalia in ventral and dorsal views
<emphasis id="CDF4A6AF2609FD4101E8974D870A53EC" bold="true">A, B</emphasis>
<taxonomicName id="0774CB704BF3192B9CFF98CFD9FAB360" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="xiuwen" status="sp. nov.">
<emphasis id="ED7456D7373A16102C583254D85B8811" italics="true">Belisana xiuwen</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4B5E117D47D4F94054C80543F2940D27" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="862FFA4C74D7D35F173F21743973B209" bold="true">C, D</emphasis>
<taxonomicName id="E2C095D7FA00DEFD8BE24554935B5A7D" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="704A2E7F45F249A49EF0BC8DBD991850" italics="true">B. yongcong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4566006D1E06B390E8157FE7DABC2BB" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="3A469736DE315090D5FB880FCA9B5796" bold="true">E, F</emphasis>
<taxonomicName id="5F7D08499DBAAEBA00C297358B16811B" authority="Tong &amp; Li, 2007" authorityName="Tong &amp; Li" authorityYear="2007" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhangi">
<emphasis id="0A74ECCE6C1AF44F3F04B70696966CF4" italics="true">B. zhangi</emphasis>
<bibRefCitation id="05762AA316DE73FD58ADA75BC59F3B18" author="Tong Y &amp; Li S" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="505-507" refId="B7" refString="Tong Y, Li S (2007) A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China. Acta Zootaxonomica Sinica 32: 505 - 507." title="A new six-eyed pholcid spider (Araneae, Pholcidae) from Karst Tiankeng of Leye County, Guangxi, China." volume="32" year="2007">Tong &amp; Li, 2007</bibRefCitation>
</taxonomicName>
<emphasis id="7B517D9A0DD311384D0A199C7DB09141" bold="true">G, H</emphasis>
<taxonomicName id="39272CE8512FCDFC8DADAC4EF42A6C7F" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhouxi" status="sp. nov.">
<emphasis id="D7984DC8BFFD9F871A8B9F52011C854D" italics="true">B. zhouxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="03A930C53189611CDA1B02F8428AC44D" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="27B907DAD93BAE50645FBB3ABD7A829D" bold="true">I, J</emphasis>
<taxonomicName id="9A6186EDC59B0A566844CF1F61DA3093" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zunyi" status="sp. nov.">
<emphasis id="F6A64463FD65E6FA182172EF7FC5CDEA" italics="true">B. zunyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AA6DC53C96981C3C601CECD68358FC3" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations:
<abbrev id="ABBRID0ENNCI" xlink_title="anterior arch">aa</abbrev>
= anterior arch,
<abbrev id="ABBRID0ERNCI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EVNCI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E5NCI" type="description">
<paragraph id="81C88B32441535A96FEC4DE5449AD08E">
<heading id="1EB87A50359A297CDE96CC3848A41F85" reason="title">Description.</heading>
</paragraph>
<paragraph id="435B064CCE8F8E6C8CBC32CBD771ED4E">
<emphasis id="BE949437FFD099D36A17FBA6171DD906" bold="true">Male</emphasis>
(
<emphasis id="0B994931E98FE16A68CD8A32BDEF94AC" bold="true">
<typeStatus id="3197039F6459EEAD30FB9F1B9463968A" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.62 (1.72 with clypeus), prosoma 0.60 long, 0.66 wide, opisthosoma 1.02 long, 0.78 wide. Leg I: 13.53 (3.60, 0.29, 3.50, 4.60, 1.54), leg II: 8.40 (2.25, 0.30, 2.08, 2.81, 0.96), leg III: 5.69 (1.64, 0.25, 1.34, 1.86, 0.60), leg IV: 7.72 (2.12, 0.24, 2.00, 2.56, 0.80); tibia I
<abbrev id="ABBRID0EIOCI" xlink_title="length/diameter">L / d</abbrev>
: 43. Eye interdistances and diameters:
<abbrev id="ABBRID0EMOCI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EQOCI" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EUOCI" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EYOCI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E3OCI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.53 / 0.47. Habitus as in Fig.
<figureCitation id="982994B9C7574EA4A473CEE7BA940645" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, without marks; clypeus and sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0EEPCI" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="407FF34D1273D2DCBF149F839202A371" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.04;
<abbrev id="ABBRID0EMPCI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="D75D4425721AC38FB2BABF7C34DC8F3D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 D</figureCitation>
). Palp as in Fig.
<figureCitation id="B1FF1B4A32C09B322DB8D91ECB10DF06" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="D779B50E20A8D6FB3C864DEEA6E9F815" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="580DEE5FE7EAF93D624D0ED083E810A1" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="9F7531FF077D9C747F735A371ADA0160" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="5B12F58FD667238FC709278D126A48BF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 B</figureCitation>
); procursus simple proximally but complex distally, with prolatero-distal membranous lamella (arrow
<quantity id="5BDE69DCD32F6BF886669CDD26F1E31B" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="830B04F3B94D3DE4925F11A606C806CC" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 C</figureCitation>
,
<figureCitation id="5912BAC4DA480E3550E5A219CE1FCCB3" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 G</figureCitation>
) bearing proximally sclerotized part, dorso-distal spine (arrow
<quantity id="51977879B9C0153104E899C10847A0A2" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="679F8875E88B9EC25491D203606FE812" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 C</figureCitation>
,
<figureCitation id="6B2B85B7F5B25B2A3C296562F1CC4E85" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 G</figureCitation>
), dorso-subdistal membranous process (arrow
<quantity id="3428CBB65525DFBE5235BF49E9692649" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="ECA014572773B4F15D903CBF880A5061" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 C</figureCitation>
,
<figureCitation id="293554C612BCEDF53FA0117C416FC690" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 G</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="C575BFDBCE827BF53429F6F3B37D9F43" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Belisana zunyi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-distal spine, arrow 3 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.03 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure20" httpUri="https://binary.pensoft.net/fig/1161065">20 D</figureCitation>
,
<figureCitation id="0FE7EAFFB316F2D1503C89C44D4DA951" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 H</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EARCI" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="ED8EF5345B90D555649E5D2FAEA445E8" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="14C7B4D040B61E7377C1E915DB4DFAC6" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 C</figureCitation>
). Retrolateral trichobothria on tibia I at 10 % proximally; legs with short vertical setae on metatarsi; tarsus I with 15 distinct pseudosegments.
</paragraph>
<paragraph id="842827BDEE5396FDAD96B3AA34C61C5A">
<emphasis id="F291BFE32E74A81722A8BDF493588CA3" bold="true">Female</emphasis>
(
<emphasis id="DF50B227B287A770C9561F2B8BEDF174" bold="true">
<typeStatus id="B842046E9281F93C86E667D0C7CDDF97" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="B7A91CDAFF8FC1AB0CBD405201A6ADDC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="91604F33218E0B1F1C7B51BD1338548A" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45234): Similar to male, habitus as in Fig.
<figureCitation id="E2FB53EAFE1BBCA0B874F6BF22783CFC" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 G, H</figureCitation>
. Total length 1.57 (1.66 with clypeus), prosoma 0.60 long, 0.68 wide, opisthosoma 0.97 long, 0.78 wide; tibia I: 2.33; tibia I
<abbrev id="ABBRID0E1RCI" xlink_title="length/diameter">L / d</abbrev>
: 35. Eye interdistances and diameters:
<abbrev id="ABBRID0E5RCI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0ECSCI" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EGSCI" xlink_title="posterior median eye">PME</abbrev>
0.06,
<abbrev id="ABBRID0EKSCI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EOSCI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.50 / 0.46. Epigyne simple and flat, posteriorly straight, with pair of median pockets 0.06 apart (
<abbrev id="ABBRID0ESSCI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="2398D90B3B8608911DA6869206D5E6E9" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 A</figureCitation>
,
<figureCitation id="8FDD79823406D979C1CE89F3FFD28B42" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 I</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0E5SCI" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="FD6A2C9737CCB60B685D6736B6E2B8F5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 B</figureCitation>
,
<figureCitation id="24385175F470CA0B783F255C10454D59" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 J</figureCitation>
) and pair of long elliptic pore plates (6 × longer than wide,
<abbrev id="ABBRID0EKTCI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="8306604DAEDED2AFCA365A63B7D4B7DB" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Belisana zunyi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure21" httpUri="https://binary.pensoft.net/fig/1161066">21 B</figureCitation>
,
<figureCitation id="E262F291799A60BCA6FDF9E9763E51DA" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWTCI" type="variation">
<paragraph id="81A8466D623E5338E178A39B5F586A34">
<heading id="243827D5C09BF19D80C4A87AC875399F" reason="title">Variation.</heading>
</paragraph>
<paragraph id="68F627EF9F2172D83AAAFB453037F7F6">
Tibia I in
<specimenCount id="61B7E717787BC5CC36F9EADB5192FA90" count="4" type="male">four male</specimenCount>
<typeStatus id="587EBA59FAEE34CCA4E87F3B90472173" type="paratype">paratypes</typeStatus>
(
<named-content id="2BC938773FB22AB5D2788DB1E016E359" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="C537805E6FF5FD37A0DFADC1982B0BE2" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4523045233): 3.12, 3.20, 3.28, 3.70. Tibia I in the other
<specimenCount id="A44AA134091DFA8AD506D1290B151FD9" count="12" type="female">12 female</specimenCount>
<typeStatus id="2D4D9E4536EA13D18BACA3DA4E2A98ED" type="paratype">paratypes</typeStatus>
(
<named-content id="7AC045212EC511B66A1B499AB3F9844F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="B66511990F2DD27910FCBBACD2EBA75C" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4523546): 2.312.50.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEUCI" type="habitat">
<paragraph id="F758358A5B12144B90EAE4DEB15F50DE">
<heading id="01FB5FA550AA55A7AF18B98D72AF49A1" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="01C182D29263DB4056B80CC6A461A1B2">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJUCI" type="distribution">
<paragraph id="653F55BAF50D8AC435822187316DA003">
<heading id="B0DA3A8C61EC86F1769C046E8C516805" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="2EA450C8A69204CF7A5A86D809779278">
<collectingCountry id="D66B23D79F3B815FED11BDCFFC5F8312" name="China">China</collectingCountry>
(
<collectingRegion id="C3B0AA508FA195BCB562822D0ACD4AFE" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="EA8B372A8794BA82A4D8AFB92F91A8B9">type</typeStatus>
locality; Fig.
<figureCitation id="217275A8554431473BC074EC4365B85A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETUCI" type="etymology">
<paragraph id="5C2B1C077A79EB09A5E6C8FBBEE4E5D9">
<heading id="E83F58BB68A09FD07E7B722C83C11953" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="E9082DB4D2F2AAA050850B06F57151D4">
The specific name refers to the
<typeStatus id="F984B44B9A77B703F0E07BA1FDFF1657">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,352 @@
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<subSubSection id="40599FAA928F17D382F63FF1ED98310E" type="nomenclature">
<paragraph id="B9F9F6B58B615DD163C518CB05D7A888">
<taxonomicName id="65C86AF0480A27CE9952E127FF3127D4" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="xiuwen" status="sp. nov.">
<emphasis id="A7C682696016BD274878D20BF57BD0E9" italics="true">Belisana xiuwen</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="8A978E4B5AF2310E5F56355D7CDFFE42" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="23295CE82F67AC45690A1665B6AF169F" type="description">
<paragraph id="F6D37D67DAD43C4F54296F100994853D">
<figureCitation id="81B29DF28083CEF5A2ABF9BA9D1A4278" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Belisana xiuwen sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal spine D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure12" httpUri="https://binary.pensoft.net/fig/1161057">Figs 12</figureCitation>
,
<figureCitation id="DEB83BDED53E29405408F9C75C390795" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13</figureCitation>
,
<figureCitation id="6BD066A7E7B197815D096CB89225D320" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 K, L</figureCitation>
,
<figureCitation id="55C2F8B3034528FC338D27E6C714DFEE" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3IAI" type="type material">
<paragraph id="E461D529E37C43A0991E0B745A0F6A01">
<heading id="9BA28BF5E0C973E805FD52FBEE9BCE94" reason="title">Type material.</heading>
</paragraph>
<paragraph id="F21B811ED7073A6C2CC65DD4AD0B5B6C">
<materialsCitation id="F64597C4B4077DF34F085C8B45F831AC" collectingDate="2007-04-20" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Xiuwen County" elevation="1035" latitude="27.1092" location="Liutong Town" longLatPrecision="1" longitude="106.49458" municipality="Guiyang" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="CC43A27A26B547697B3C0BD29C1DBB3A" bold="true">
<emphasis id="F201EC09936F347A9417FE5890AD75FD" bold="true" italics="true">
<typeStatus id="19F021BB041167148699220B4990C6B6" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="66699D1DFEA9B8AE4E6A9F9C9AE361B3" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="7CB305F91EE68B0E5544CAE989115E2C" count="1" type="male"></specimenCount>
;
<collectingRegion id="C5EC20DE54B38FB8C61269A3C4AFF520" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingMunicipality id="EAE8EC30360058EEB5AA459ECAC9229C">Guiyang</collectingMunicipality>
,
<collectingCounty id="325D370BE822089065295FCA62EB5690">Xiuwen County</collectingCounty>
,
<location id="B47EB3E65F70D529666C190446B09C8D" LSID="urn:lsid:plazi:treatment:39A5A39107205634AF79DE26FEAC4E47:B47EB3E65F70D529666C190446B09C8D" country="China" county="Xiuwen County" latitude="27.1092" longLatPrecision="1" longitude="106.49458" municipality="Guiyang" name="Liutong Town" stateProvince="Guizhou">Liutong Town</location>
,
<location id="C859F3E6B0E1ECA681D907C929355C57" LSID="urn:lsid:plazi:treatment:39A5A39107205634AF79DE26FEAC4E47:C859F3E6B0E1ECA681D907C929355C57" country="China" county="Xiuwen County" latitude="27.1092" longLatPrecision="1" longitude="106.49458" municipality="Guiyang" name="Daxing Village" stateProvince="Guizhou">Daxing Village</location>
,
<location id="ABB97282D1F37E2A1E6A3AB84CF09EFE" LSID="urn:lsid:plazi:treatment:39A5A39107205634AF79DE26FEAC4E47:ABB97282D1F37E2A1E6A3AB84CF09EFE" country="China" county="Xiuwen County" latitude="27.1092" longLatPrecision="1" longitude="106.49458" municipality="Guiyang" name="Duobing Cave" stateProvince="Guizhou">Duobing Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKJAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.494583,27.109200]}">
<geoCoordinate id="31E24E56199CFE968A3034B548DCC98E" degrees="27" direction="north" minutes="06.552" orientation="latitude" precision="1" value="27.1092">27 ° 06.552 ' N</geoCoordinate>
,
<geoCoordinate id="8F496B764B3A7B1CA1476FB6DFEB31A7" degrees="106" direction="east" minutes="29.675" orientation="longitude" precision="1" value="106.49458">106 ° 29.675 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="0A4C4FECE566D4568B622DCF5D86A0E1" metricMagnitude="3" metricUnit="m" metricValue="1.035" unit="m" value="1035.0">
<elevation id="158CCCAC9C0E8EF34CFB88FB54E38C08" metricMagnitude="3" metricUnit="m" metricValue="1.035" unit="m" value="1035.0">1035 m</elevation>
</quantity>
;
<date id="E3D6531A8045B5B81ABEFB65D71CF6D7" value="2007-04-20">
<collectingDate id="64239FE04320EBD8A258F1BF0C8117C2" value="2007-04-20">20 Apr. 2007</collectingDate>
</date>
;
<collectorName id="34DCC83577B8FA936F5D01E6A13E7E97">J. Liu</collectorName>
&amp;
<collectorName id="74478360CEB4B620398F8F9612E90EE8">Y. Lin</collectorName>
leg.;
<named-content id="C592FA8FB3450D03D16005142DC8C79D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="8190B9C94DA3D1D8539141E973FC2AEB" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="149DFC2423FFCA10DBA2B11BFCF6E278">Ar</collectorName>
45214
</materialsCitation>
.
<materialsCitation id="B135EC9830953188E676FF0162D10E1A" collectingDate="2007-04-20" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Xiuwen County" elevation="1035" latitude="27.1092" location="Liutong Town" longLatPrecision="1" longitude="106.49458" municipality="Guiyang" specimenCount="3" specimenCount-female="3" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="156400AC8CFC9402315F10A9E8CCBA6D" bold="true">
<emphasis id="5F4BC3E98847694091DAC95AF8DAD00E" bold="true" italics="true">
<typeStatus id="BA478355B312BD101A0C416E72CF9301" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="AB2CD5CD12C1DDD1CB84E6F96C2A7F72" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="DD0E3461161A6C76A2043F03886054CD" count="3" type="female">3 ♀</specimenCount>
; same data as for holotype;
<named-content id="8CBE928A57991BF53DBABB2E8D8A1321" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="CCB8BAF1967DF8D034D71C0C55403A63" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4521517
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2JAI" type="diagnosis">
<paragraph id="DB3F6EE8E6CF075052856ACD6325BC99">
<heading id="0C628AE9404F1BE6113DA5EB5C36248E" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="97BD84A9681601328F207E2025353B6E">
The new species resembles
<taxonomicName id="8531CB5335B3AAABDEE31DC3653B5A5E" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="C4863770B5C9BB39C30D58C938E909EF" italics="true">B. yongcong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="85BDA780570BE55E8F72A3C8A6D0E4A3" rank="species">sp. nov.</taxonomicNameLabel>
(Figs
<figureCitation id="5570A6CB5FD4D9B39C70C527D9BD0618" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14</figureCitation>
,
<figureCitation id="6EDE951489592EF114A81B872BD2CB2E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15</figureCitation>
,
<figureCitation id="5A1576030E9AA0A59BEDC4914451892F" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A, B</figureCitation>
,
<figureCitation id="ABDBED8665F2FF739DCA47D4584510CA" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 C, D</figureCitation>
) by having similar male chelicerae (tips of distal apophyses widely separated and pointing inwards; Fig.
<figureCitation id="34A55EAE093844245F326AB8F01C0BDA" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 D</figureCitation>
), bulbal apophysis (hooked; Fig.
<figureCitation id="DCB114D4A55CC8C7A7A03EAD8E6C5AFD" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 C</figureCitation>
), and epigyne (epigynal pockets on lateral part of epigynal plate, epigynal plate posteriorly curved; Figs
<figureCitation id="19A13828E26BC53A0873F48AD9811A16" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 A</figureCitation>
,
<figureCitation id="7A554FAC6185A719A976657A851AE1B1" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 A</figureCitation>
), but can be distinguished by procursus with distal spine (arrow
<quantity id="FD8048A09DD54B99EF6A53C80D6C7EA6" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="DD174A1B5532A990B53366A8495D3219" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Belisana xiuwen sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal spine D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure12" httpUri="https://binary.pensoft.net/fig/1161057">12 C</figureCitation>
,
<figureCitation id="B48A52448C4AE41D40F5A9F0B765D7B6" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 K</figureCitation>
vs absent) and without ventro-subdistal membranous lamella, prolatero-distal membranous lamella, and sclerotized dorso-subdistal apophysis (Figs
<figureCitation id="92FA81F5F8F2EAF797BFC29323A0854D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Belisana xiuwen sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal spine D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure12" httpUri="https://binary.pensoft.net/fig/1161057">12 C</figureCitation>
,
<figureCitation id="C32E3D8054922C987F706EA4F226E2F7" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 K</figureCitation>
vs present), and by vulval pore plates long elliptic (3 × longer than wide,
<abbrev id="ABBRID0E3LAI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="4B8CE32DA3578A04C333FAD061D4CB86" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 B</figureCitation>
,
<figureCitation id="C4C8E2B1BC08825AAD0D11AA9D6BC9AA" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 B</figureCitation>
vs curved and 9 ×).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIMAI" type="description">
<paragraph id="5C6879DD2C4FC7973BB21D92AA167CC7">
<heading id="C048B283FFDCFFA1F4EA080B3909E785" reason="title">Description.</heading>
</paragraph>
<paragraph id="B64A8D0957F0E3EFE3C831532BCA0D1E">
<emphasis id="051218FF1484270B9F1368F75A33924D" bold="true">Male</emphasis>
(
<emphasis id="7100E9EE54FFE56F9D471165BE24BF8A" bold="true">
<typeStatus id="F2D9DDBDC02AE62D63A8BF461F3E876F" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.35 (1.48 with clypeus), prosoma 0.57 long, 0.59 wide, opisthosoma 0.78 long, 0.60 wide. Leg I: 10.06 (2.48, 0.21, 2.73, 3.36, 1.28), leg II: 6.94 (2.00, 0.22, 1.74, 2.18, 0.80), leg III: 5.08 (1.50, 0.20, 1.14, 1.64, 0.60), leg IV: 6.72 (1.94, 0.20, 1.72, 2.15, 0.71); tibia I
<abbrev id="ABBRID0ESMAI" xlink_title="length/diameter">L / d</abbrev>
: 46. Eye interdistances and diameters:
<abbrev id="ABBRID0EWMAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E1MAI" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0E5MAI" xlink_title="posterior median eye">PME</abbrev>
0.05,
<abbrev id="ABBRID0ECNAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EGNAI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.45 / 0.40. Habitus as in Fig.
<figureCitation id="B3BF2443A738B5487CEFE34D28EA473A" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, without marks; clypeus and sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Eye pigments indistinct. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0EONAI" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="AC6D72B1E17D3F813E9E74AFEF57108C" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.30;
<abbrev id="ABBRID0EWNAI" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="7F6D4D9A255B8BAFABAFF8CE5DB6FDEB" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 D</figureCitation>
). Palp as in Fig.
<figureCitation id="A89E8B7FC6878327CE5D2CBD3A307FD4" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Belisana xiuwen sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal spine D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure12" httpUri="https://binary.pensoft.net/fig/1161057">12 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="A2120FD3C18D7AE6AFE69754501DB1FE" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="6DE3B2064FAE939330F2D67C3F582415" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Belisana xiuwen sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal spine D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure12" httpUri="https://binary.pensoft.net/fig/1161057">12 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="63239A4BEDA8B5C1C77E8141C7C9CB63" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="CA45BEADFED68EBAD50959D866195477" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Belisana xiuwen sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal spine D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure12" httpUri="https://binary.pensoft.net/fig/1161057">12 B</figureCitation>
); procursus simple, with distal spine (arrow
<quantity id="FD238DCCF65D672C7A5607D4F1C51DB4" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="50858E067EA97E790CF51AA6C124268F" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Belisana xiuwen sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal spine D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure12" httpUri="https://binary.pensoft.net/fig/1161057">12 C</figureCitation>
,
<figureCitation id="2515B2E083B6B1CFE18A14FCAC563102" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 K</figureCitation>
) and retrolateral membranous flap (f in Figs
<figureCitation id="41FE362ACC96E5963AD2581DEF1F8C25" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Belisana xiuwen sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at distal spine D retrolateral view). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure12" httpUri="https://binary.pensoft.net/fig/1161057">12 D</figureCitation>
,
<figureCitation id="44E8CE6CA230A0E0CDBF920F380E5A56" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 L</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0E1OAI" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="EAC17FF33FA3952ED5FDE34D4F91C271" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="3830DBB171B086E76E0626CEE4D4EF3E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 C</figureCitation>
). Retrolateral trichobothria on tibia I at 6 % proximally; legs with short vertical setae on metatarsi; tarsus I with 15 distinct pseudosegments.
</paragraph>
<paragraph id="7DFBFB1A1258C68CE6B4B067861762AE">
<emphasis id="FDEDAE14D293D9290720692B854DE9E5" bold="true">Female</emphasis>
(
<emphasis id="00EC6482F90DBC7F0C9771A602542514" bold="true">
<typeStatus id="BB43CEB4CF29B7128FECAAEE950EB309" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="A984BD2A98F2176AD53C12C932711F1F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="1011C030D91843DBD59E690449344B8C" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45215): Similar to male, habitus as in Fig.
<figureCitation id="95D9BB773503367A195F44D1F04216F5" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 G, H</figureCitation>
. Total length 1.75 (1.85 with clypeus), prosoma 0.56 long, 0.59 wide, opisthosoma 1.19 long, 0.80 wide; tibia I: 2.19; tibia I
<abbrev id="ABBRID0EUPAI" xlink_title="length/diameter">L / d</abbrev>
: 44. Eye interdistances and diameters:
<abbrev id="ABBRID0EYPAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E3PAI" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EAQAI" xlink_title="posterior median eye">PME</abbrev>
0.05,
<abbrev id="ABBRID0EEQAI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EIQAI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.44 / 0.38. Epigyne simple and flat, posteriorly curved, with pair of lateral pockets 0.33 apart (
<abbrev id="ABBRID0EMQAI" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="02D01ED90862401FCD93373436965A53" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 A</figureCitation>
,
<figureCitation id="B6E78FFAAC00B560625DDFDD98DB5921" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 A</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0EYQAI" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="211727097B24A4BED8F1A1B6D36A0E23" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 B</figureCitation>
,
<figureCitation id="6D650421045AADEFC2CEEF936684E0DB" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 B</figureCitation>
) and pair of long elliptic pore plates (3 × longer than wide,
<abbrev id="ABBRID0EERAI" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="F9B53F3B12B27C4F5BF6ED84B5CE38AF" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Belisana xiuwen sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure13" httpUri="https://binary.pensoft.net/fig/1161058">13 B</figureCitation>
,
<figureCitation id="3DFDD21693E115DE0CE3AF2FE9DCC50B" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQRAI" type="variation">
<paragraph id="6812C3F754F2278A692B3173096D5ABE">
<heading id="9DB1EFFA5A951B961DE0D4891B6DEB46" reason="title">Variation.</heading>
</paragraph>
<paragraph id="1A8BCD0ADCEF3983DB2E1B8D691902D6">
Tibia I in the other
<specimenCount id="D4B537B4CCAFFE63AF563D9BAEEB2D44" count="2" type="female">two female</specimenCount>
<typeStatus id="EE8B7F5A1FA301078C9A5BE1580FEAF2" type="paratype">paratypes</typeStatus>
(
<named-content id="472CA0EDF2FC0E0803ABB3138670752A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="42639D91F6739AA3A9A8D295C097F162" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4521617): 2.10, 2.18.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1RAI" type="habitat">
<paragraph id="7A60867E05F7094B6179FB123B50AE35">
<heading id="E8CE29918B0EB225EF7A4C97B1C7A3FC" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="0ECE072CE134F7D94F957F672A037B6F">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0E6RAI" type="distribution">
<paragraph id="3D720B4FFCA86A62A8AA3B58DC4A8D77">
<heading id="04F00A0AA29A375C9B6F15E85B6E0BA8" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="5FE053B9495762764FEDBAD2064E0D33">
<collectingCountry id="B9E85F68C3816FCA8BD81C6085E009A7" name="China">China</collectingCountry>
(
<collectingRegion id="9A8DBD03C72D0D91EB95AD45CDBBB8DF" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="0E908206141FA6285094B21B2711E361">type</typeStatus>
locality; Fig.
<figureCitation id="6511F07120F5EDFBB6DACF7FE4027077" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJSAI" type="etymology">
<paragraph id="6649C42AA80E579E01F26164DDA1DF8F">
<heading id="367EB650DFBED7D7BE348EF47B3555C2" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="93F2C563C2BCDED1ADC8B04E7E93D6A6">
The specific name refers to the
<typeStatus id="156D30225C2F99C79DDBBEE4C13A56A1">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,367 @@
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<mods:title id="58FFDD7591788C86EBBE9C80B2221E84">Taxonomic review of the grasshopper genus Pteropera Karsch, 1891 (Orthoptera, Acrididea, Catantopinae) with description of three new species and a preliminary phylogeny of the Cameroonian species</mods:title>
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<mods:namePart id="266444B8AAD240AF8154F66FB7767769">Ivković, Slobodan</mods:namePart>
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<mods:affiliation id="A2F28155EF68997A5823A0A5913035B6">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<mods:namePart id="FB1ADE6AD72C2E789E8C388B99AD3555">Husemann, Martin</mods:namePart>
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<mods:affiliation id="34BB2FDD39959A333A4746F594FC38E4">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<mods:date id="7676BE10EDF2C4650B1EE63421100D39">2024</mods:date>
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<treatment id="5D23573267F55BF18ED3BEC76DDB2983" ID-arpha="5D235732-67F5-5BF1-8ED3-BEC76DDB2983" LSID="urn:lsid:plazi:treatment:5D23573267F55BF18ED3BEC76DDB2983" httpUri="http://treatment.plazi.org/id/5D23573267F55BF18ED3BEC76DDB2983">
<subSubSection id="C9A285B02E5BB2E7A9D35515A024771D" type="nomenclature">
<paragraph id="72790EC84B9B41A861D18C62B2201548">
<taxonomicName id="56562D1356798ECB729EBADA83E88CAE" ID-CoL="4Q25W" authority="Ramme, 1929" authorityName="Ramme" authorityYear="1929" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="carnapi">
<emphasis id="7D831F21694D02834C7ADA62E4224AA6" italics="true">Pteropera carnapi</emphasis>
<bibRefCitation id="02AF9A6CA78FCFC7B341C6FC63864E4D" DOI="10.1002/mmnz.4830150202" author="Ramme W" journalOrPublisher="Mitteilungen aus dem zoologischen Museum in Berlin" pagination="247-492" refId="B23" refString="Ramme W (1929) Afrikanische Acrididae. Revisionen und Beschreibungen wenig bekannter und neuer Gattungen und Arten. Mitteilungen aus dem zoologischen Museum in Berlin 15: 247 - 492. https: // doi. org / 10.1002 / mmnz. 4830150202" title="Afrikanische Acrididae. Revisionen und Beschreibungen wenig bekannter und neuer Gattungen und Arten." volume="15" year="1929">Ramme, 1929</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="91C2146F2AACD648A0CBA3C65422B1C1" type="description">
<paragraph id="87A101C4CD384A87BE7AF94B833900B4">
<figureCitation id="ED8BEF694D5CCDAE9C6C358D05B78571" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Images of holotypes and allotypes of Pteropera species in lateral view A P. carnapi (holotype ♂) B P. carnapi (paratype ♀) C P. congoensis (holotype ♂) D P. congoensis (allotype ♀) E P. cornici (holotype ♂) F P. cornici (allotype ♀) G P. descampsi (holotype ♂) H P. descampsi (allotype ♀) I P. descarpentriesi (holotype ♂) J P. descarpentriesi (allotype ♀)." figureDoi="10.3897/zookeys.1216.130270.figure3" httpUri="https://binary.pensoft.net/fig/1160991">Figs 3 A, B</figureCitation>
,
<figureCitation id="5DCA62A9D49F0F8491A9C5EE6BF903BD" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Images of holotypes, allotypes and paratype of Pteropera species in dorsal view A P. carnapi (holotype ♂) B P. carnapi (paratype ♀) C P. congoensis (holotype ♂) D P. congoensis (allotype ♀) E P. cornici (holotype ♂) F P. cornici (allotype ♀) G P. descampsi (holotype ♂) H P. descampsi (allotype ♀) I P. descarpentriesi (holotype ♂) J P. descarpentriesi (allotype ♀)." figureDoi="10.3897/zookeys.1216.130270.figure8" httpUri="https://binary.pensoft.net/fig/1160996">8 A, B</figureCitation>
</paragraph>
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<subSubSection id="0AB5BD9F59A4A5FE963A868453BA35FF" type="reference_group">
<paragraph id="16BB4C7C89FA7CB3F6A387EBA3782BB7">
<treatmentCitationGroup id="51834FF38CD452830801E7D8D3BBF5ED">
<treatmentCitation id="14EDF607B1601A83E9483D6E4D0109CF" author="Sjöstedt Y" page="28" year="1931">
<taxonomicName id="DD4C19E2D74766CB53BD55A4DE1991A3" ID-CoL="4Q25T" authority="Sjostedt 1931: 28" authorityName="Sjostedt" authorityPageNumber="28" authorityYear="1931" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="biloloca">
<emphasis id="FD24BE2EDC68C36B57783D5D65C8A045" italics="true">Pteropera biloloca</emphasis>
<comment id="D603ECAFFC3AE7CAA808FF004A2BCAFE">
<bibRefCitation id="7388FFBEBEAC1E004B1D03248F5AB380" author="Sjöstedt Y" firstAuthor="Sjostedt" issue="15" journalOrPublisher="Arkiv för Zoologi (A)" page="28" pagination="1-64" refId="B27" refString="Sjöstedt Y (1931) Acridoidea aus Kongo und anderen Teilen von Afrika. Arkiv för Zoologi (A) 22 (15): 164." title="Acridoidea aus Kongo und anderen Teilen von Afrika." volume="22" year="1931">Sjöstedt 1931: 28</bibRefCitation>
;
<treatmentCitation id="51AD9398AC97E982E6F0E9F082F38B1D" author="Dirsh VM" page="339" year="1965">
<bibRefCitation id="7CEC03FD8F553E70829431C46F4E0FEB" author="Dirsh VM" firstAuthor="Dirsh" journalOrPublisher="Anti-Locust Research Centre and Cambridge University Press, London" page="339" refId="B3" refString="Dirsh VM (1965) The African genera of Acridoidea. Anti-Locust Research Centre and Cambridge University Press, London, 579 pp." title="The African genera of Acridoidea." year="1965">Dirsh 1965: 339</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
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</paragraph>
<paragraph id="BE0D1A9DB7A4C79FB902C0C6F8BC373E">
<treatmentCitationGroup id="8E5B33868989FCA72C6277B1C0EA4225">
<treatmentCitation id="AA5A0B0D7DE615143DAFD63C859F79FF" author="Ramme W" page="364" year="1929">
<taxonomicName id="8255A218972CE088EC9A1DCC2877794D" ID-CoL="4Q25W" authority="Ramme 1929: 364" authorityName="Ramme" authorityPageNumber="364" authorityYear="1929" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="carnapi">
<emphasis id="771C84186B182F267419CE0A867D265A" italics="true">Pteropera carnapi</emphasis>
<comment id="D51CFA234CBEB1C6412466F320C481BD">
<bibRefCitation id="8277A0ABDD889E1FAEDC263DA321B302" DOI="10.1002/mmnz.4830150202" author="Ramme W" firstAuthor="Ramme" journalOrPublisher="Mitteilungen aus dem zoologischen Museum in Berlin" page="364" pagination="247-492" refId="B23" refString="Ramme W (1929) Afrikanische Acrididae. Revisionen und Beschreibungen wenig bekannter und neuer Gattungen und Arten. Mitteilungen aus dem zoologischen Museum in Berlin 15: 247492. https://doi.org/10.1002/mmnz.4830150202" title="Afrikanische Acrididae. Revisionen und Beschreibungen wenig bekannter und neuer Gattungen und Arten." volume="15" year="1929">Ramme 1929: 364</bibRefCitation>
;
<treatmentCitation id="1042AF5B87D3EE0327BF90F84F2F29AE" author="Donskoff M" page="59" year="1981">
<bibRefCitation id="C700FD28C84277A43ABA08209F356C91" DOI="10.1080/21686351.1981.12278263" author="Donskoff M" firstAuthor="Donskoff" issue="1" journalOrPublisher=")" page="59" pagination="33-88" refId="B4" refString="Donskoff M (1981) Les acridiens de la forêt africaine. I. Révision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Société entomologique de France (N. S.) 17 (1): 3388. [51 figs] https://doi.org/10.1080/21686351.1981.12278263" title="Les acridiens de la forêt africaine. I. Révision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Société entomologique de France (N. S." volume="17" year="1981">Donskoff 1981: 59</bibRefCitation>
</treatmentCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
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</paragraph>
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<paragraph id="FB610B8AF89D73F17B8F16585B9583E2">
<heading id="4028FC6E830E6162F699AD9708A80A16" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="C2361A068A47C9D81BE3ABE000BC2C8E">
<materialsCitation id="4000AE1222898F812D38FE1563C60005" collectingDate="1887-06" collectionCode="V, MfN, DORSA" collectorName="Carnap" country="Cameroon" latitude="3.84805" location="Yaounde" longLatPrecision="15" longitude="11.501945" specimenCode="BA 000180, S 01" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="18320611A3817C0B8016D32A9EAA4362" bold="true">
<emphasis id="8840AD400357D308A26009A0976B97EB" bold="true" italics="true">
<typeStatus id="20E3D43A249894407028872AC26449B2" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="1025D666C1DE2BAEC65A08C25FA1CBD8" name="Cameroon">Cameroon</collectingCountry>
<specimenCount id="32542D44E2980340F4EB4C7B10FF113E" count="1" type="male"></specimenCount>
;
<location id="5B668A3F39D4BA276D986329EF67A9B0" LSID="urn:lsid:plazi:treatment:5D23573267F55BF18ED3BEC76DDB2983:5B668A3F39D4BA276D986329EF67A9B0" country="Cameroon" latitude="3.84805" longLatPrecision="15" longitude="11.501945" name="Yaounde">Yaoundé</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EV1AK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[11.501944,3.848050]}">
<geoCoordinate id="A71EA899FBD675A941AC97FD7E4E1CDA" degrees="3" direction="north" minutes="50.883" orientation="latitude" precision="1" value="3.84805">3 ° 50.883 ' N</geoCoordinate>
,
<geoCoordinate id="B204B9EFBABD20EA8A70CBF47EC1C238" degrees="11" direction="east" minutes="30" orientation="longitude" precision="15" seconds="7" value="11.501945">11 ° 30 ' 7 &quot; E</geoCoordinate>
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;
<date id="42C0B3ECD39A53BEA64CCAE25A0B4910" value="1887-06">
<collectingDate id="69F40BED136AFF69007C90549CCE3FF0" value="1887-06">Jun. 1887</collectingDate>
</date>
;
<collectionCode id="BEE151559525038A438615CC3C9918D5" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.
<collectorName id="25A61F1B298B1A1FEBAA7C791D2220D0">Carnap</collectorName>
leg.;
<named-content id="E4C1F0843F19ECA89771A98F41DC5E45" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-f%C3%BCr-naturkunde-berlin" xlink_title="Museum für Naturkunde Berlin, Germany">
<collectionCode id="419822445A8D220871A773EEAEF7D774" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" type="Museum">MfN</collectionCode>
</named-content>
,
<specimenCode id="A0D5ECBCB47F27839537102CE1E1AADE" collectionCode="MfN" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" type="Museum">
<accessionNumber id="0AFB925748AA269D943F2D254EAA609D" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/BA000180" type="EnaNcbi">BA 000180</accessionNumber>
</specimenCode>
<specimenCode id="2F62E4B294F3F7FD674593D5D3939A52" collectionCode="MfN" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" type="Museum">S 01</specimenCode>
-
<collectionCode id="D4B232E71B0289CA90558D9F3EA2F03B">DORSA</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E51AK" type="material">
<paragraph id="2EB2D3853317CC492D0B4C4653A7531E">
<heading id="AC4BEC5EFFCA0FBD2F3A274B0C011210" reason="title">Other material examined.</heading>
</paragraph>
<paragraph id="2826507C956AFFB99B922CA4BE74E964">
<collectingCountry id="BD12D5197ED591490C065ADA59B7BC53" name="Cameroon">Cameroon</collectingCountry>
<materialsCitation id="3332EEDA522AFC3CA65157CBAD0530D0" collectingDate="2020-06-15" collectionCode="SMNK" collectorName="J. A. Yetchom Fondjo" country="Cameroon" latitude="3.7086167" location="Ongot" longLatPrecision="1" longitude="11.252784" specimenCount="4" specimenCount-female="4">
<specimenCount id="D30637EF901C6781D07D74ACB44E85DB" count="4" type="female">4 ♀♀</specimenCount>
;
<location id="85F8C3429107003A7EF721CD3D74E15C" LSID="urn:lsid:plazi:treatment:5D23573267F55BF18ED3BEC76DDB2983:85F8C3429107003A7EF721CD3D74E15C" country="Cameroon" latitude="3.7086167" longLatPrecision="1" longitude="11.252784" name="Ongot">Ongot</location>
;
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<geoCoordinate id="B29B0D4523573E9D7060C171D9218503" degrees="3" direction="north" minutes="42.517" orientation="latitude" precision="1" value="3.7086167">3 ° 42.517 ' N</geoCoordinate>
,
<geoCoordinate id="C22FA14F87BFE831CB64FC6363F967F5" degrees="11" direction="east" minutes="15.167" orientation="longitude" precision="1" value="11.252784">11 ° 15.167 ' E</geoCoordinate>
</named-content>
;
<date id="2E98444FEF060705590DAEFA4AD664FA" value="2020-06-15">
<collectingDate id="4359C0D540C14047B01AD9D3ED371232" value="2020-06-15">15 Jun. 2020</collectingDate>
</date>
;
<collectorName id="8D2994E15A3D8D834C8F9070F5FFC26C">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="F5CD7A29EA799817BA55FBFF5A350230" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="A6AF1ED2CE7015147FB1572FF91B2BC3" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
.
<collectingCountry id="61F8A135215019B66E374A71E1DE1E33" name="Cameroon">Cameroon</collectingCountry>
</materialsCitation>
<materialsCitation id="865EA1853088682DDDE2466E71D499F3" collectingDate="2022-01-07" collectionCode="SMNK" collectorName="J. A. Yetchom Fondjo" country="Cameroon" elevation="731" latitude="4.4500165" location="Iboti" longLatPrecision="1" longitude="10.450033" municipality="Ebo forest" specimenCount="2" specimenCount-male="2">
<specimenCount id="D4145B3B6D59BD5B55F1B9B2D0B9A07C" count="2" type="male">2 ♂♂</specimenCount>
;
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,
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;
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<geoCoordinate id="AAACAAF4B2F8310825901D04BA4286D0" degrees="4" direction="north" minutes="27.001" orientation="latitude" precision="1" value="4.4500165">4 ° 27.001 ' N</geoCoordinate>
,
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,
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<quantity id="31FA303223327A182B6360151BB2AD42" metricMagnitude="2" metricUnit="m" metricValue="7.31" unit="m" value="731.0">731 m</quantity>
a. s. l.
</elevation>
;
<date id="679C597E72CE53D5C051E747A0496F2F" value="2022-01-07">
<collectingDate id="BE82BB80410E511873AF3149F5B74C36" value="2022-01-07">7 Jan. 2022</collectingDate>
</date>
;
<collectorName id="1D7103638CC22288861CA4B5E36CBCA5">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="56848235361274059A48752ACBE6002D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="1ED8B1690388D2FE3160E7992D99294A" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
.
<collectingCountry id="5CD129B8727D3677FFB3289B265FBC9B" name="Cameroon">Cameroon</collectingCountry>
</materialsCitation>
<materialsCitation id="70C2C442D2BC2485E0FE04634C5201FB" collectingDate="2022-06-12" collectionCode="R, SMNK" collectorName="Nzoko Fiemapong" country="Cameroon" elevation="731" latitude="3.3560667" location="Deng-Deng National Park" longLatPrecision="1" longitude="12.743584" specimenCount="13" specimenCount-female="4" specimenCount-male="9">
<specimenCount id="1774DD50CCA9B62F70E0A344E131A787" count="9" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="9708579E5C201577B310432370E81EFB" count="4" type="female">4 ♀♀</specimenCount>
;
<location id="5950A0DDE5680AF73AD6C88688DEFBE6" LSID="urn:lsid:plazi:treatment:5D23573267F55BF18ED3BEC76DDB2983:5950A0DDE5680AF73AD6C88688DEFBE6" country="Cameroon" latitude="3.3560667" longLatPrecision="1" longitude="12.743584" name="Deng-Deng National Park">Deng-Deng National Park</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E62AK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[12.743583,3.356067]}">
<geoCoordinate id="EB00CC5778D30D2DF38F415D0FB0AE2B" degrees="3" direction="north" minutes="21.364" orientation="latitude" precision="1" value="3.3560667">3 ° 21.364 ' N</geoCoordinate>
,
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<quantity id="425BEC96FD424258FE408A2DF675B2AE" metricMagnitude="2" metricUnit="m" metricValue="7.31" unit="m" value="731.0">731 m</quantity>
a. s. l.
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;
<date id="ADA83F4BB866E8A5C997634E304A88EB" value="2022-06-12">
<collectingDate id="F2B270F21E11AC8DDCEDCF2784574A73" value="2022-06-12">12 Jun. 2022</collectingDate>
</date>
; A.
<collectionCode id="B38A0C063C7C305E548F127FA9349FE4" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectorName id="1A621B862BBCA8375E09B3306407D90D">Nzoko Fiemapong</collectorName>
leg.;
<named-content id="1BFF481F5C000F4A1E55A26D9FA2C7AC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="42929DAF4336C3E1AFD28ED631AC910B" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<caption id="E7EA238DFC674C8E662D7B1DF5DCFD3E" ID-DOI="10.3897/zookeys.1216.130270.figure3" ID-arpha="A1D52247-85FD-5989-9E93-193BDD4B6F7A" httpUri="https://binary.pensoft.net/fig/1160991" startId="F3">
<paragraph id="7140C1B0F7F0336472E0E2D014A2B1F1">
<label id="2840563B04CA61CEB9B9A890A063D8EA">Figure 3.</label>
</paragraph>
<paragraph id="3932EFC0ABE96D4F12CF3182FFE478A4">
Images of holotypes and allotypes of
<taxonomicName id="B59B9A73CA14E3E91184E1F85E547154">
<emphasis id="10741244EDDB43F06B3D9C0250F4C670" italics="true">Pteropera</emphasis>
species
</taxonomicName>
in lateral view
<emphasis id="42ED53C2EB13AE6895F9DBD0BBBE6875" bold="true">A</emphasis>
<taxonomicName id="3E67C83B0F56428E802393FBB4DA4645" authorityName="Ramme" authorityYear="1929" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="carnapi">
<emphasis id="82B977395B7FF74DA04024B23FB2D91E" italics="true">P. carnapi</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="08035F17AC623F0592F13DB37DC99E64" bold="true">B</emphasis>
<taxonomicName id="275DD1AA311258F63D3FCA43C711D5B0" authorityName="Ramme" authorityYear="1929" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="carnapi">
<emphasis id="A899C2E7795041248EA65FC1839E9C6D" italics="true">P. carnapi</emphasis>
</taxonomicName>
(paratype ♀)
<emphasis id="629D754C6473333BFFC13FC68D6DC5A2" bold="true">C</emphasis>
<taxonomicName id="936D8E04B08F4B3D34E7CF1D062F3EEA" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="congoensis">
<emphasis id="3AA27015E4076D290601BF812135CAE7" italics="true">P. congoensis</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="5F6EA771888FE2E4E0DE48F05A37A491" bold="true">D</emphasis>
<taxonomicName id="5C644D34A9BE8CC4E7537F4062435316" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="congoensis">
<emphasis id="D7EC525C893ACEAD4B3049BBC604A798" italics="true">P. congoensis</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="C834BBC116A1FC5B02D954BF3CF4D4BF" bold="true">E</emphasis>
<taxonomicName id="5874D54C27691EFA870D65C025AD933C" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="cornici">
<emphasis id="762B08F7EBB5EF6AE3C504AFB4BA217C" italics="true">P. cornici</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="A772B2E927A6C5411C34CAA60D0EBF32" bold="true">F</emphasis>
<taxonomicName id="8178098197C56E082351C5E3EB8CD3EB" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="cornici">
<emphasis id="6CAECF3F412A4D2C121E160FF280533C" italics="true">P. cornici</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="D29B2B3D236BBB77553721A9EA5DD627" bold="true">G</emphasis>
<taxonomicName id="3E406B2D3B3A8B0523DAD3785AC5F372" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="5BB834813F8ACAF9FED3D55246630E2F" italics="true">P. descampsi</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="303B482C52CE0AB9A977CFCBD4E4A270" bold="true">H</emphasis>
<taxonomicName id="41C967891401AAFAD083D1C63E99C4A6" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="79543355EC93391C7DCE0BC0BF493E0D" italics="true">P. descampsi</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="FB93A3BFD2C16915FFA18C6A7ECC92C5" bold="true">I</emphasis>
<taxonomicName id="9BAD56A6D70DAA69F11987F2C56C2596" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="E13AD73AC4017494473D62E051DCE2F8" italics="true">P. descarpentriesi</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="E823A5886A354B8EB08B36EC197977AD" bold="true">J</emphasis>
<taxonomicName id="B709CD682E52679ECB769BA701104C82" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="30437BA81475D2128198B90DE44B07ED" italics="true">P. descarpentriesi</emphasis>
</taxonomicName>
(allotype ♀).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E5ABK" type="redescription">
<paragraph id="4EC42927DA12245A24AC490A723ABF43">
<heading id="D9D5DA69478993BFD1F9A733CABAF5BA" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="B6DA882F4A91A75817F6494952533945">
Frontal ridge raised above the median ocelli, prominent between the antennae; head and pronotum with well contrasted pale and dark colors; subocellar facial spot large; lower margin of elytra shiny black; meso- and metathoracic episternites yellow in their center; front legs, ventral area of hind femora yellow-green; dorsal area of the body and upper areas of hind femora more or less dark brown; hind tibiae bluish-green; male pallium and supra-anal plate raised; male cerci bilobate, with inner lobe equal to or longer than the outer lobe.
<emphasis id="2CC031E565FD0B04029E0CBB3FE5C785" bold="true" italics="true">Epiphallus</emphasis>
(Fig.
<figureCitation id="80A4AB3B085C873F5215D0B22CC5D331" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Internal genitalia of P. carnapi (A D), P. descampsi (E H), P. karschi zenkeri (I L), P. uniformis (M P) and P. verrucigena (O T)." figureDoi="10.3897/zookeys.1216.130270.figure15" httpUri="https://binary.pensoft.net/fig/1161003">15 A</figureCitation>
): Ancorae small and closer to each other; bridge arched; anterior projections prominent, triangular.
<emphasis id="9381454A58FEC39DA08885109F80CCE1" bold="true" italics="true">Phallic complex</emphasis>
(Fig.
<figureCitation id="C5CE9D16B2B6C2E40FA72CEE0122A6A8" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Internal genitalia of P. carnapi (A D), P. descampsi (E H), P. karschi zenkeri (I L), P. uniformis (M P) and P. verrucigena (O T)." figureDoi="10.3897/zookeys.1216.130270.figure15" httpUri="https://binary.pensoft.net/fig/1161003">15 B D</figureCitation>
): dorsal arch of cingulum U-shaped, apodemes slender, clearly overhanging or exceeding level of separation of endophallic valves, with strongly incurved apex; upper ectophallic sheath very long in profile; lower ectophallic sheath not covering or enveloping the base of the rami; rami not bent; aedeagus valves in the form of a thin blade; latero-ventral sclerite triangular.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EACBK" type="distribution">
<paragraph id="650FCDCB17753ED3E6266A8E0FD68D33">
<heading id="03D938D3AC7EE1C1C2DB38241B770A81" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="AEBCCD292E7F71DA50889762F36B8E2B">
<collectingCountry id="6EC1886551AD7C57D8D820A372326480" name="Cameroon">Cameroon</collectingCountry>
;
<collectingCountry id="820D349067CBC6D7B50798F40DE322AA" name="Central African Republic">Central African Republic</collectingCountry>
;
<collectingCountry id="A0B50536992B11D0ADF1448C2895102E" name="Gabon">Gabon</collectingCountry>
;
<collectingCountry id="4A87A0483359FCA3ACB8E603DDA4CAEE" name="Democratic Republic of the Congo">Congo</collectingCountry>
(Fig.
<figureCitation id="3D7F10245A64391754ADCD6818AE11C8" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of Pteropera species in African rainforests." figureDoi="10.3897/zookeys.1216.130270.figure16" httpUri="https://binary.pensoft.net/fig/1161004">16 B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,487 @@
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<mods:title id="05C84E17363334E914931AA1A8B90F00">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<subSubSection id="F99472D6DC35D1B36B7B2D370A59931D" type="nomenclature">
<paragraph id="9CE48018DF74CE96B592A5B18910E15C">
<taxonomicName id="248A46F32D44D61F1E8730FD44FEB16D" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="majiang" status="sp. nov.">
<emphasis id="EB9472508EBA2DC840828BED2C586509" italics="true">Belisana majiang</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="1FB91F58298D8F756AA0079C907A6B66" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="221C99AB08A1B0808EEFA78D0B1F873B" type="description">
<paragraph id="CB318CC3BD7E478635B7F43D4F1DB95F">
<figureCitation id="DEF1CE2E2CC40E0A09AF0AA664D12582" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">Figs 6</figureCitation>
,
<figureCitation id="494C5776AF24E71C877C1201AD3F0FF3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7</figureCitation>
,
<figureCitation id="98559904883A820BCB45C91F71A72B7D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E, F</figureCitation>
,
<figureCitation id="4F98EEC532024F6544DCE9371F8D8ABB" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAVAG" type="type material">
<paragraph id="D9275CEBEE5D5D43B5D4221DA7DD2F15">
<heading id="9432A10A4EEEB92DB48A8DC15DA6F785" reason="title">Type material.</heading>
</paragraph>
<paragraph id="2239B8BE553DBFE6ECD538F6781B1532">
<materialsCitation id="98B77ADFA4664CAD207F75EFCE5E2652" collectingDate="2011-11-28" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zha &amp; Ar" country="China" county="Majiang County" elevation="1056" latitude="26.504284" location="Xingshan Town" longLatPrecision="1" longitude="107.51572" municipality="Kaili" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="6FF89EE200C84E108C9E3FBC242A888A" bold="true">
<emphasis id="2E40F58534AC393D3F94504EB11C0510" bold="true" italics="true">
<typeStatus id="F2A540F3BE3C8BCFACFDEF66CE1446CE" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="185C621E817F58830FEA0C6E872A472B" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="058669F935D2051C3E675BB435E27B1C" count="1" type="male"></specimenCount>
;
<collectingRegion id="4EABC678D788052571DD8490DC86F507" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingMunicipality id="7E38A290ED6B8AD997768490EE8E711E">Kaili</collectingMunicipality>
,
<collectingCounty id="96DC936D2EB56A503932E92A5F0581FB">Majiang County</collectingCounty>
,
<location id="B81DDC8F55203B0551F6FF3FDD35004C" LSID="urn:lsid:plazi:treatment:6422EB5754B05143A2195D60B09B121B:B81DDC8F55203B0551F6FF3FDD35004C" country="China" county="Majiang County" latitude="26.504284" longLatPrecision="1" longitude="107.51572" municipality="Kaili" name="Xingshan Town" stateProvince="Guizhou">Xingshan Town</location>
,
<location id="606EDFCEF5A3558AA872240D44E943F9" LSID="urn:lsid:plazi:treatment:6422EB5754B05143A2195D60B09B121B:606EDFCEF5A3558AA872240D44E943F9" country="China" county="Majiang County" latitude="26.504284" longLatPrecision="1" longitude="107.51572" municipality="Kaili" name="Gubin Village" stateProvince="Guizhou">Gubin Village</location>
,
<location id="D3253ABE49133482B8B2C426F6A48EB9" LSID="urn:lsid:plazi:treatment:6422EB5754B05143A2195D60B09B121B:D3253ABE49133482B8B2C426F6A48EB9" country="China" county="Majiang County" latitude="26.504284" longLatPrecision="1" longitude="107.51572" municipality="Kaili" name="Guazhutou Cave" stateProvince="Guizhou">Guazhutou Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOVAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[107.515717,26.504283]}">
<geoCoordinate id="094C68EC7D884E7D48A758FEE75BB600" degrees="26" direction="north" minutes="30.257" orientation="latitude" precision="1" value="26.504284">26 ° 30.257 ' N</geoCoordinate>
,
<geoCoordinate id="C5A956A29FC7EBD1D0959B0F7DE78E8F" degrees="107" direction="east" minutes="30.943" orientation="longitude" precision="1" value="107.51572">107 ° 30.943 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="18E393D611E0F079253F793DC1A5FA8E" metricMagnitude="3" metricUnit="m" metricValue="1.0559999999999998" unit="m" value="1056.0">
<elevation id="AF74E8A888EF98E75CA579B2ED4FBC34" metricMagnitude="3" metricUnit="m" metricValue="1.0559999999999998" unit="m" value="1056.0">1056 m</elevation>
</quantity>
;
<date id="628FD04BFC343B54400B693737679B7F" value="2011-11-28">
<collectingDate id="3F0DA8D8DF14C39ACE9B1E213A5E54FD" value="2011-11-28">28 Nov. 2011</collectingDate>
</date>
;
<collectorName id="2C7D2C41E3E64ACF50FA6B4E9ED968B8">Z. Chen</collectorName>
&amp;
<collectorName id="F8521B5501E61C9ADED6F7855A3AE059">Z. Zha</collectorName>
leg.;
<named-content id="2FCDA13344BCE0576740970C3BAD8E4D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="F50736CAA24FB96517B8CF1F0ECB166E" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="D1C4C0D18D49016C6150E9683991C2A9">Ar</collectorName>
45194
</materialsCitation>
.
<materialsCitation id="96D9A6E51DC0AA7AE2BF3C35BF9BBC67" collectingDate="2011-11-28" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zha &amp; Ar" country="China" county="Majiang County" elevation="1056" latitude="26.504284" location="Xingshan Town" longLatPrecision="1" longitude="107.51572" municipality="Kaili" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="345EA2F9741FF0B7D2248454863FF9AD" bold="true">
<emphasis id="5D1CC63FB8ADFDFE905B53D5AC180B67" bold="true" italics="true">
<typeStatus id="2BE987A8A1ECBC135D9864FF545776F5" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="64BB56F3E2BC0DD642A43F10DE80C6CB" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="57534AAD5FCA01EFAF6F32C4BECEC728" count="1" type="male">1 ♂</specimenCount>
; same data as for holotype;
<named-content id="6DA01412D6F6180130893BD06F9CD4E0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="AADB3E9A73C0B3609C10D1BF42D22B40" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45195
</materialsCitation>
<materialsCitation id="3093922D215699E78B6F93C5D2BA681C" collectingDate="2011-11-28" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zha &amp; Ar" country="China" county="Majiang County" elevation="1056" latitude="26.504284" location="Xingshan Town" longLatPrecision="1" longitude="107.51572" municipality="Kaili" specimenCount="2" specimenCount-female="2" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="CB0B4D9A1FE0393F7EA953EE53F431D9" count="2" type="female">2 ♀</specimenCount>
; same data as for holotype;
<named-content id="8F4E7B36BD6FFC489EEF75C08202FC3E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
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</named-content>
- Ar 4519697
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDWAG" type="diagnosis">
<paragraph id="168D113BAC32A432ADA43A2801DA6078">
<heading id="440C26E039B445706579A65BAD47DAC8" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="88756C724AFF51A4C5C4438FFB8AA83B">
The new species resembles
<taxonomicName id="854AB72B0ECCC7074E920E872438F0B9" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="zhouxi" status="sp. nov.">
<emphasis id="2CEA63601FAEC26D38837E1717BA8027" italics="true">B. zhouxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F83CE601ACE4096D589CE0DCA1CB96A5" rank="species">sp. nov.</taxonomicNameLabel>
(Figs
<figureCitation id="436CA3FD985D26865A6E3C094AC38B89" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Belisana zhouxi sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrow 2 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure18" httpUri="https://binary.pensoft.net/fig/1161063">18</figureCitation>
,
<figureCitation id="94A59D1AE7BEA37C4DC80070918F5EE9" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Belisana zhouxi sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.20 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure19" httpUri="https://binary.pensoft.net/fig/1161064">19</figureCitation>
,
<figureCitation id="EB17BF4E6B81AB5D9C0BD16C03D9C300" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 E, F</figureCitation>
,
<figureCitation id="C0AD19B125D5536B99343124701C1490" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 G, H</figureCitation>
) by having similar male chelicerae (tips of distal apophyses pointing downwards; Fig.
<figureCitation id="7897DCA3301FB8567FE42EB82BF5B0E2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 D</figureCitation>
), bulbal apophysis (hooked; Fig.
<figureCitation id="A01365F5228A1A3F40745B302B33F083" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 C</figureCitation>
), and epigyne (epigynal pockets on antero-lateral part of epigynal plate, epigynal plate posteriorly curved; Figs
<figureCitation id="BA6DA5A7B098D7C94191937E30088956" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 A</figureCitation>
,
<figureCitation id="5E6A0454A412F033E97830795917E254" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 E</figureCitation>
), but can be distinguished by procursus with distinct ventro-subdistal membranous process and dorso-distal spine (arrows 1,
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Figs
<figureCitation id="EB9332787DC303209930AB9BEB4C7E65" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="4D7101A5A40383B15B7205C6253BA266" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
vs absent) and by vulval pore plates long elliptic (3 × longer than wide,
<abbrev id="ABBRID0E3XAG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="A4833B00A8740FD9C3D6EB1BC92AF457" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 B</figureCitation>
,
<figureCitation id="16885E088BB7F2A181674921CB76416A" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 F</figureCitation>
vs 2 ×); also distinguished from
<taxonomicName id="BC6B31E0F8EF04FF76D625C97367CDDC" authority="Zhang, Chen &amp; Zhu, 2008" authorityName="Zhang, Chen &amp; Zhu" authorityYear="2008" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tongle">
<emphasis id="E80F58F39DB79B361492C83F0CFA8731" italics="true">B. tongle</emphasis>
<bibRefCitation id="9A4B9D22C1CA7F94E517B6C60A820106" author="Zhang Y &amp; Chen H &amp; Zhu M" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="654-656" refId="B21" refString="Zhang Y, Chen H, Zhu M (2008) A new troglophilous Belisana spider from Guangxi, China (Araneae, Pholcidae). Acta Zootaxonomica Sinica 33: 654 - 656." title="A new troglophilous Belisana spider from Guangxi, China (Araneae, Pholcidae)." volume="33" year="2008">Zhang, Chen &amp; Zhu, 2008</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="8CE4ADFCC175FE191A56F9A30712B5EA" author="Zhang Y &amp; Chen H &amp; Zhu M" etAl="et al." firstAuthor="Zhang" journalOrPublisher="Acta Zootaxonomica Sinica" page="654" pagination="654-656" refId="B21" refString="Zhang Y, Chen H, Zhu M (2008) A new troglophilous Belisana spider from Guangxi, China (Araneae, Pholcidae). Acta Zootaxonomica Sinica 33: 654656." title="A new troglophilous Belisana spider from Guangxi, China (Araneae, Pholcidae)." volume="33" year="2008">Zhang et al. 2008: 654</bibRefCitation>
, figs 15) by procursus with dorso-subdistal membranous process (arrow
<quantity id="EB616897A3BE8FF0714A04E09B48CEFF" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
Figs
<figureCitation id="BEA779E74799068CA79DB5515B603D74" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="93D0D8538295B6FDE4A8C735DCA551C8" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
vs absent), by prolatero-distal membranous lamella of procursus without rectangular sclerite (arrow
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Figs
<figureCitation id="D6017B32311573E31B156B447A6D499D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="6EAE9575B1DE780B076BD62D4BCC92F3" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
vs present), by ventro-subdistal membranous process of procursus 3 × longer than wide (arrow
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Figs
<figureCitation id="83EE7AAE867BE3EEE5C9614F248FA9E0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="2F44653E7366035C5912D1E93CD82E99" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
vs 8 ×), and by dorso-distal spine of procursus 6 × longer than wide (arrow
<quantity id="25CE56A2266B81ED55488B7F910A23AF" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="0D31E8C515972971D73623FA755C33BB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="94B11CCCED7F241875AC75D6E007A8E3" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
vs 12 ×).
</paragraph>
<caption id="3204799C26DBDC0FB7C266FF934AB127" ID-DOI="10.3897/zookeys.1216.132561.figure6" ID-arpha="ED60C96D-7076-5C27-B1D9-F8195BFB3E26" httpUri="https://binary.pensoft.net/fig/1161051" startId="F6">
<paragraph id="B6B27A5C5AE21CAB2ABD6A954B48D56B">
<label id="DB3EBCC0559F75EE5196AE1B2442420C">Figure 6.</label>
</paragraph>
<paragraph id="13B60FCB62BA6AD7245836E139D459C7">
<taxonomicName id="1CBAC17717B7C78A3959AAC314759B91" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="majiang" status="sp. nov.">
<emphasis id="361E352A6AFDC43228D221BAAB37538A" italics="true">Belisana majiang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="14F8773C97A3E027DB443F8024B0CBD5" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="BDAD7CC7A4626C3E15DEE75C6F387CC5" bold="true">A, B</emphasis>
palp (
<emphasis id="B918E44350C844F8FA128FB46C1F382A" bold="true">A</emphasis>
prolateral view
<emphasis id="5FD1166C9C58AD227851F9603F99C0D3" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="18997609FAAADB96C05641D63A372D36" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="62A1B26C8E84636F41F0471A916B2355" bold="true">C</emphasis>
prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process
<emphasis id="B173CF17A136F0E2CE7A748055DFF04D" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb,
<abbrev id="ABBRID0EW1AG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0E11AG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="C793EAC59B757A8D80BC4E03935146CB" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="C136BE80EBD2CE8EE702B095E685B3D6" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="C7850E2DFD0B61E64C210D8CF2BFD39C" ID-DOI="10.3897/zookeys.1216.132561.figure7" ID-arpha="FC7070CB-316A-58C2-8B2D-3BF6CE2DF030" httpUri="https://binary.pensoft.net/fig/1161052" startId="F7">
<paragraph id="64735B3EC6FBABF10D76E960BA123645">
<label id="8AFB17BDF21995F973B191D120C807A7">Figure 7.</label>
</paragraph>
<paragraph id="AA24C595894D64A16F17230A90807051">
<taxonomicName id="0C37D9FEC1B29559D6A280AF5DDCFB8F" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="majiang" status="sp. nov.">
<emphasis id="3768048624FC296891D892CC5C1E9C51" italics="true">Belisana majiang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B588055174D4A486D2A46841A5137A00" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="3B4C72209D45F7BC3B560CE26E725099" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="4EF47B717379EDF71C6A2F39016A9950" bold="true">A, B, G, H</emphasis>
)
<emphasis id="1049682F8F1816B896F1BAA8C0FCFBA5" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="ADB622B0CDC6666ED3A2477278714EED" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="4F43A6FCA9B2FA1560E5244ECF790386" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="9100D36CBA497F6E951817076E62E6A9" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="63B352C1B607F91FD29F613CFD522F95" bold="true">E H</emphasis>
habitus (
<emphasis id="01824C14E73A0734E3986EB1A8A125E5" bold="true">E, G</emphasis>
dorsal view
<emphasis id="EA975BCA9DC1E72B670A2A5459EFEA0B" bold="true">F</emphasis>
lateral view
<emphasis id="116080EE5E7C798A952085483D3F515C" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EO3AG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0ES3AG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EW3AG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0E13AG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0E53AG" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0EC4AG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.05 (
<emphasis id="1E33F8468A394D2A4F0AF3DF285822E8" bold="true">A D</emphasis>
); 0.30 (
<emphasis id="DB7C489DFF74F16E6B88EC1322171A39" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EP4AG" type="description">
<paragraph id="EFF1FB9EF577C6FB1F8689AB01F10AA3">
<heading id="EA8BEDE75A1D1E1A7E6B68F661980681" reason="title">Description.</heading>
</paragraph>
<paragraph id="182B826126ED7D2F16C8839EF184785E">
<emphasis id="37B2DB53C4C58B20466E62DEA82ECCAF" bold="true">Male</emphasis>
(
<emphasis id="B47733DF518BAB51CEBD17FCFC7BE4C6" bold="true">
<typeStatus id="4E256ACC88A596D4DF377D050CD535EF" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.35 (1.43 with clypeus), prosoma 0.52 long, 0.59 wide, opisthosoma 0.83 long, 0.64 wide. Leg I missing, leg II: 6.13 (1.78, 0.22, 1.50, 1.90, 0.73), leg III: 4.19 (1.28, 0.20, 0.95, 1.33, 0.43), leg IV: 5.87 (1.72, 0.22, 1.48, 1.80, 0.65). Eye interdistances and diameters:
<abbrev id="ABBRID0EZ4AG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E44AG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EB5AG" xlink_title="posterior median eye">PME</abbrev>
0.06,
<abbrev id="ABBRID0EF5AG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EJ5AG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.44 / 0.38. Habitus as in Fig.
<figureCitation id="CF5D22490B70DF48C70670DEF1D03846" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, without marks; clypeus and sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0ER5AG" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="D1D642B63C8BCF7B9486913CCE68070E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.19;
<abbrev id="ABBRID0EZ5AG" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="22FFF53DDCBCBF1EAC64E2ACD9D57C88" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 D</figureCitation>
). Palp as in Fig.
<figureCitation id="FA7ADB715C319D66CBE50F09EB599743" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 A, B</figureCitation>
; trochanter with ventral apophysis (1.5 × longer than wide, arrow
<quantity id="DD316DE8743F97FEB7CE8BA51927FA1C" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="3BC1652D8FCE10EE04EF098FD235CFBE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="D57FE8D1A80013FC3487EC94D2F0344B" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="EC3FD5AA19E062C68A61BEC8C884083B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 B</figureCitation>
); procursus simple proximally but complex distally, with ventro-subdistal membranous process (arrow
<quantity id="A3A4A868511F28F81E8736A38EAB7B83" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="C78EBBDC1391425C96DF5A1EDB7FE349" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="34D599D708EE2B200CFE74392F9FDE5B" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
), prolatero-distal membranous lamella (arrow
<quantity id="C940CB9ADE35537D2BD5E563D46457A8" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="C609EE544138D8F6D014EAD5BD85B9AE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="1AB6612A0E8FB4A89EDE8C4D10C5BF0D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
) bearing proximally slightly sclerotized part, dorso-distal spine (arrow
<quantity id="606B51606968563D5A2FAFF4A1FA8CF5" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="0E4882072B19B70FFFA2D4AB2B728D75" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="E4BBDA4405A828CA1D421515E59BD641" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
), dorso-subdistal membranous process (arrow
<quantity id="CEF9121AFD7F92B542B00DB9CA6E518F" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
Figs
<figureCitation id="69D975DB23A31BFCA58F5A918B52DDA8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 C</figureCitation>
,
<figureCitation id="B002E13C8A4BF041D7F916DB0EFC7C85" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 E</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="952EFB47EAD8C4530CC37EE50D392D29" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana majiang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure6" httpUri="https://binary.pensoft.net/fig/1161051">6 D</figureCitation>
,
<figureCitation id="B82EC171F7DF1F998D6578D279005962" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 F</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EVABG" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="1608EF5F02074EEDC544AC5FCC98A0D6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="E37144298A699F5FA6881B3CFA9FE8CE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 C</figureCitation>
).
</paragraph>
<paragraph id="A5BA5C7F373C1AB04FFEA9A34F855215">
<emphasis id="AAB03885A58DB746DC409E2C9FC8AA1D" bold="true">Female</emphasis>
(
<emphasis id="97126A80EE59228F05162C61D9D9DCBA" bold="true">
<typeStatus id="006890CF2E99DF5567E6D23FF86BD7BF" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="E1EB1C43027A1E29AB656ED93E95F13E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="D21482D35D8F2607C3E2227578C46F49" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45196): Similar to male, habitus as in Fig.
<figureCitation id="73B8F6F9D7B5E3893E638DF7247857C1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 G, H</figureCitation>
. Total length 1.48 (1.58 with clypeus), prosoma 0.48 long, 0.56 wide, opisthosoma 1.00 long, 0.84 wide. Eye interdistances and diameters:
<abbrev id="ABBRID0EPBBG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0ETBBG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EXBBG" xlink_title="posterior median eye">PME</abbrev>
0.06,
<abbrev id="ABBRID0E2BBG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E6BBG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.43 / 0.39. Epigyne simple and flat, posteriorly curved, with pair of antero-lateral pockets 0.14 apart (
<abbrev id="ABBRID0EDCBG" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="075533F98E1763CE896FA79109D77170" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 A</figureCitation>
,
<figureCitation id="89070562CD2CD08BA473C6EDBA28B8D9" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 E</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0EPCBG" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="7412DB92E41F26669C69ACC5124F7487" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 B</figureCitation>
,
<figureCitation id="F0485D96498E534C5E442127A5DCD26D" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 F</figureCitation>
) and pair of long elliptic pore plates (3 × longer than wide,
<abbrev id="ABBRID0E2CBG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="89A2C82FDE5889326C6D10F8068C6CBD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana majiang sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure7" httpUri="https://binary.pensoft.net/fig/1161052">7 B</figureCitation>
,
<figureCitation id="19E68B5AB6031A21618DEB7FED50873B" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHDBG" type="variation">
<paragraph id="235EF4846548A2D02FCAB07E8C8C1879">
<heading id="BA61059E155F0007460EE7768B34ED42" reason="title">Variation.</heading>
</paragraph>
<paragraph id="85625EDEDCBA992268F7A2D89E48E23C">
Unknown. Legs I missing in male
<typeStatus id="19CD3CF6815ABF41C404C0952C136221" type="paratype">paratype</typeStatus>
(
<named-content id="8F4CE5828D990E15E494D2DF7A46E7CC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="D79C01B962CEA9005CF5227E2709C9B1" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45195) and another female
<typeStatus id="D98C11B3B6541979293B41CADB99EDE6" type="paratype">paratype</typeStatus>
(
<named-content id="A08A98D1AE1286AE0AFBA70F031ED53E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="628DD564CF034E09490C1649D9B94B3E" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45197).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVDBG" type="habitat">
<paragraph id="60AF5F86F8E3F240E781F3DBDBAC15B1">
<heading id="F4AD46BFB0C3051DAD163C20C470FDF9" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="3139617FFFD450A6690B2DE5AC9AE70E">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0E1DBG" type="distribution">
<paragraph id="406C2D291F8A4349A6106B0E7891E61F">
<heading id="44F96672151DF793B591EDEA8A233E8B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="7F70098EF27FB2E11FDCF0375F8D3152">
<collectingCountry id="D1B0F9D6413CF92E67FA56699F9C7466" name="China">China</collectingCountry>
(
<collectingRegion id="64D29E1F0FC2ECC26FB9131B9FDF0FFC" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="597B54023DE21E90881A3F64AEC9453D">type</typeStatus>
locality; Fig.
<figureCitation id="CC3E45B1765357F79DA1316B75B3EB2B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEEBG" type="etymology">
<paragraph id="5EC93315142189C43C2B2CAA3DDFC861">
<heading id="28B2EB5A05648F2511B709F72FDA3A68" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="7A69EFA716AC4F500B0C9735FEBF2E34">
The specific name refers to the
<typeStatus id="9463989EF39B7272F22C3F6BFF635A1E">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,382 @@
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<paragraph id="F4DDCE549C8F4743309473F67FDDBC0E">
<taxonomicName id="E16908A6B220BE632863C8A74847B0ED" authority="Yetchom &amp; Husemann" authorityName="Yetchom Fondjo &amp; Nzoko Fiemapong &amp; Tindo &amp; Duressa &amp; Ivković &amp; Husemann" authorityYear="2024" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="kennei" status="sp. nov.">
<emphasis id="4998F9FAF91CD64DF6D2ED7434C95525" italics="true">Pteropera kennei</emphasis>
Yetchom &amp; Husemann
</taxonomicName>
<taxonomicNameLabel id="D47F4F8936A823B7CEBD7BBB8993203F" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="4EEC943C557A8BCE3C1BD608BCA77794" type="description">
<paragraph id="219C94650E09510B83137CC983510F58">
<figureCitation id="F6559005AACE80C6CF5EC9FD5D662F68" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pteropera kennei sp. nov. A habitus image of a male under natural conditions B male frontal view C female frontal view D male lateral view E female lateral view F male dorsal view G female dorsal view H epiphallus dorsal view I phallic complex lateral view J phallic complex dorsal view K phallic complex ventral view L female subgenital plate M female spermatheca." figureDoi="10.3897/zookeys.1216.130270.figure12" httpUri="https://binary.pensoft.net/fig/1161000">Fig. 12 A M</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPKCK" type="type material">
<paragraph id="B6FB50456273063CBFA15F639D730AD8">
<heading id="0612EE7F84607202101002FB1B07A3F0" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="B81A1F69223AC658F6EA23EA5CA5D590">
<materialsCitation id="BE162AE8B5139B5CB9EE23C53C7089E1" collectingDate="2022-04-10" collectionCode="SMNK" collectorName="J. A. Yetchom Fondjo" country="Cameroon" elevation="606" latitude="3.3741333" location="Somalomo" longLatPrecision="1" longitude="12.733167" municipality="Dja River" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="F8A556F83C3DA487A83A63423A0F2A62" bold="true">
<emphasis id="F8CA6D1EF3B3CE1640DBCACEC47A94D8" bold="true" italics="true">
<typeStatus id="7F17F2EF4BE7C418386E19E090CD5DDC" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="A6F9AFF44C6AB55C9866313206B3B154" name="Cameroon">Cameroon</collectingCountry>
<specimenCount id="43A32666D428517384D9D1E82356A9A4" count="1" type="male"></specimenCount>
;
<location id="C25F52EE68D6CC67ED68C4F22306D962" LSID="urn:lsid:plazi:treatment:649D939286565941890E7162AD71835F:C25F52EE68D6CC67ED68C4F22306D962" country="Cameroon" latitude="3.3741333" longLatPrecision="1" longitude="12.733167" municipality="Dja River" name="Somalomo">Somalomo</location>
, in the forest along the
<collectingMunicipality id="4F612B4F64308D8682B4E1C477730BE6">Dja River</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4KCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[12.733167,3.374133]}">
<geoCoordinate id="4CF997FB024AC6F915299A26AFB2B1D4" degrees="3" direction="north" minutes="22.448" orientation="latitude" precision="1" value="3.3741333">3 ° 22.448 ' N</geoCoordinate>
,
<geoCoordinate id="4CF7D68095D8F03BB8BA55BBA81F6C0E" degrees="12" direction="east" minutes="43.990" orientation="longitude" precision="1" value="12.733167">12 ° 43.990 ' E</geoCoordinate>
</named-content>
,
<elevation id="C4E105C5884E4F23E72EE3D19ED370EA" metricMagnitude="2" metricUnit="m" metricValue="6.06" unit="m" value="606.0">
<quantity id="B7327542FCE2A34F5749FBABDB7A4914" metricMagnitude="2" metricUnit="m" metricValue="6.06" unit="m" value="606.0">606 m</quantity>
a. s. l.
</elevation>
;
<date id="AE631DE5B24A23C92FD6382054BE00E4" value="2022-04-10">
<collectingDate id="7CE4DB733C6EB54336A09B0C4EC54B7C" value="2022-04-10">10 Apr. 2022</collectingDate>
</date>
;
<collectorName id="0E80148C4EBDA3D543FD6BF89F825F9B">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="C87B46C4BE704591A14C161F1E859365" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="1DE652CF4F160DCD2B66CF4EC91252AB" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
,
<named-content id="0B3428E4C5A0F88B49BE634B241F1D1F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="8CBE83A1AC09190ACC7EB34A4C8CF19A" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
- ORTH- 0000001
</materialsCitation>
.
<materialsCitation id="278383C904355DDC60879017C7CEF3D0" country="Cameroon" location="Cameroon" specimenCount="1" typeStatus="paratype">
<emphasis id="7D4C9770FF77C155A02573468EA8A3CD" bold="true">
<emphasis id="091A3E7D05E355D18BB673A00C83C2BD" bold="true" italics="true">
<typeStatus id="D7F6EF7144210A59D374185CB3D383F8" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="04C54BE7010FFFDF2D8345A04933DECC" name="Cameroon">Cameroon</collectingCountry>
</materialsCitation>
<materialsCitation id="6CF732BACB6418971FA0C60B5A6375E1" collectingDate="2022-04-10" collectingDateMax="2022-04-11" collectingDateMin="2022-04-10" collectionCode="R, SMNK, MNHN" collectorName="J. A. Yetchom Fondjo &amp; Nzoko-Fiemapong" country="Cameroon" elevation="606" latitude="3.3741333" location="Somalomo" longLatPrecision="1" longitude="12.733167" municipality="Dja River" specimenCount="19" specimenCount-female="3" specimenCount-male="16" typeStatus="paratype">
<specimenCount id="D4CAE5CCF30EA8DE1978C312D43BD70C" count="16" type="male">16 ♂♂</specimenCount>
,
<specimenCount id="882DABDCD5CE6D3A1CF943BC68144143" count="3" type="female">3 ♀♀</specimenCount>
;
<location id="9BF74D13119FE803DCB8AC4E6A780148" LSID="urn:lsid:plazi:treatment:649D939286565941890E7162AD71835F:9BF74D13119FE803DCB8AC4E6A780148" country="Cameroon" latitude="3.3741333" longLatPrecision="1" longitude="12.733167" municipality="Dja River" name="Somalomo">Somalomo</location>
, in the forest along the
<collectingMunicipality id="321362FA20B5B62160DBADD51F016AD9">Dja River</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERLCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[12.733167,3.374133]}">
<geoCoordinate id="2EF03A3818808F81F68204B10205D951" degrees="3" direction="north" minutes="22.448" orientation="latitude" precision="1" value="3.3741333">3 ° 22.448 ' N</geoCoordinate>
,
<geoCoordinate id="9B56163B1E082345A93116BDB2B6F28E" degrees="12" direction="east" minutes="43.990" orientation="longitude" precision="1" value="12.733167">12 ° 43.990 ' E</geoCoordinate>
</named-content>
,
<elevation id="5A33CD00BF19F569EBDF218C83D355F3" metricMagnitude="2" metricUnit="m" metricValue="6.06" unit="m" value="606.0">
<quantity id="E757C8FBE470B947BBFC10E866FC82AB" metricMagnitude="2" metricUnit="m" metricValue="6.06" unit="m" value="606.0">606 m</quantity>
a. s. l.
</elevation>
;
<date id="5F5CE5EA38D61C1433346E8514FC96AF" value="2022-04-10" valueMax="2022-04-11" valueMin="2022-04-10">
<collectingDate id="BC45D907B6277FE2442045337B40BA54" value="2022-04-10" valueMax="2022-04-11" valueMin="2022-04-10">1011 Apr. 2022</collectingDate>
</date>
;
<collectorName id="4BE02A9559F758BB7240DA4BFA31226B">J. A. Yetchom Fondjo</collectorName>
&amp; A.
<collectionCode id="11DC0CEA86F669B81E90E01CFFA56AD2" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectorName id="1248E67147F1E8E739D960CF9689D3E8">Nzoko-Fiemapong</collectorName>
leg.;
<named-content id="D2940E14B0F448692AD8A187CA0CEBFF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="72744F0CE66A045C44EDC1E50C8CC3CB" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
,
<named-content id="B0725B287B9EC772AC851E0813404366" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum National d´Histoire Naturelle Paris, France">
<collectionCode id="C3837C6F2D2E75B17E1F0B9BCFA8979E" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
.
<collectingCountry id="2E1105E77E730536A0F4E9F8A67DFC85" name="Cameroon">Cameroon</collectingCountry>
</materialsCitation>
<materialsCitation id="56DE161D2A5058A3B19BFB2A7DA78EAA" collectingDate="2022-06-12" collectionCode="R, SMNK" collectorName="Nzoko-Fiemapong" country="Cameroon" elevation="661" latitude="3.3560667" location="Deng-Deng National Park" longLatPrecision="1" longitude="12.743584" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="316964EFA618D11585D30DD4920C9B29" count="1" type="male">1 ♂</specimenCount>
;
<location id="3798BD6F7522F1DB76EA264200E17D1D" LSID="urn:lsid:plazi:treatment:649D939286565941890E7162AD71835F:3798BD6F7522F1DB76EA264200E17D1D" country="Cameroon" latitude="3.3560667" longLatPrecision="1" longitude="12.743584" name="Deng-Deng National Park">Deng-Deng National Park</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBMCK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[12.743583,3.356067]}">
<geoCoordinate id="DD5A3CD767CFE46CB7148D63ACC3D758" degrees="3" direction="north" minutes="21.364" orientation="latitude" precision="1" value="3.3560667">3 ° 21.364 ' N</geoCoordinate>
,
<geoCoordinate id="28DC994EBDE47B5E15A4603A28088066" degrees="12" direction="east" minutes="44.615" orientation="longitude" precision="1" value="12.743584">12 ° 44.615 ' E</geoCoordinate>
</named-content>
,
<elevation id="4D4CAD41DDA3814908CCE91897E644B4" metricMagnitude="2" metricUnit="m" metricValue="6.609999999999999" unit="m" value="661.0">
<quantity id="1525FE80072B19CE9685D59255D95A08" metricMagnitude="2" metricUnit="m" metricValue="6.609999999999999" unit="m" value="661.0">661 m</quantity>
a. s. l.
</elevation>
;
<date id="167E3684DEB7FED872983CEA9407F5B3" value="2022-06-12">
<collectingDate id="4E3879EBF1CEF9F96389D0769B244ABB" value="2022-06-12">12 Jun. 2022</collectingDate>
</date>
; A.
<collectionCode id="105C20A187A14CCF6E79ADF2B09AF966" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectorName id="D5C90E1BB01C2466F74FD63968F34E86">Nzoko-Fiemapong</collectorName>
leg.;
<named-content id="32AE339AF716BAD2DAAB0247A0F85C91" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="900A278AD920BCFA998F6F4ED8E41030" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKMCK" type="diagnosis">
<paragraph id="5C5850EC2D33B99CAEE9AC0065055BDE">
<heading id="7983E806C67F5A1A62C0BA0C2063B280" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="8C66858CCDF43247E4BEDC5E17FC2A1C">
<taxonomicName id="CF7757FD7753699E311E5794A7F579B8" authorityName="Yetchom Fondjo &amp; Nzoko Fiemapong &amp; Tindo &amp; Duressa &amp; Ivković &amp; Husemann" authorityYear="2024" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="kennei" status="sp. nov.">
<emphasis id="70C34758255ECFDE57F42016DC7342DD" italics="true">Pteropera kennei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE5DE16C317282FD3CDECE8FA4D7F6CC" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="C5D11A4274DB4644AE9FE0154CD9373B" authority="Bruner, 1920" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="5C437B1AFDA8B348A684CDCFFCFB3DCB" italics="true">P. uniformis</emphasis>
Bruner, 1920
</taxonomicName>
, from
<collectingCountry id="6FAE00D09E8656E5265DA9F2596CFCBA" name="Cameroon">Cameroon</collectingCountry>
in terms of its general coloration, a dark longitudinal band and contiguous pale bands on the pronotum disc and the outer area of hind femora without pale spots. However, the new species can easily be distinguished from
<taxonomicName id="19E579525698BFA881C5267BBF0F5FB4" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="BD5A6F34D2F0129905852A832AB1EF6E" italics="true">P. uniformis</emphasis>
</taxonomicName>
(Figs
<figureCitation id="CF272C6100A463841DEBA8E7D5FCC4B1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Images of holotypes, allotypes, letotype and neallotype of Pteropera species in lateral view A P. spleniata (holotype ♂) B P. spleniata (holotype ♀) C P. teocchii (holotype ♂) D P. teocchii (allotype ♀) E P. thibaudi (holotype ♂) F P. thibaudi (allotype ♀) G P. verrucigena (lectotype ♂) H P. verrucigena (♀) I P. villiersi (holotype ♂) J P. villiersi (allotype ♀) K P. uniformis (neallotype ♂)." figureDoi="10.3897/zookeys.1216.130270.figure6" httpUri="https://binary.pensoft.net/fig/1160994">6 K</figureCitation>
,
<figureCitation id="AEB29D1D7E63E3DA21375A0A59679179" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Images of holotypes, allotypes, neallotype and paratype of Pteropera species in dorsal view A P. teocchii (holotype ♂) B P. teocchii (allotype ♀) C P. thibaudi (holotype ♂) D P. thibaudi (allotype ♀) E P. verrucigena (holotype ♂) F P. verrucigena (allotype ♀) G P. villiersi (holotype ♂) H P. villiersi (allotype ♀) I P. uniformis (neallotype ♂) J P. femorata (♀)." figureDoi="10.3897/zookeys.1216.130270.figure11" httpUri="https://binary.pensoft.net/fig/1160999">11 I</figureCitation>
) by its lateral lobes of pronotum without a pale basal band (present in
<taxonomicName id="E3D958EAEEDA91B2D0E3CF89995E9279" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="42794883952154271DCEDB424241E6F9" italics="true">P. uniformis</emphasis>
</taxonomicName>
as well as in all other
<taxonomicName id="9AF95FD6FF52290E59B9D81D18C0A4AF">
<emphasis id="71141036FA1EF4321227C8719210B681" italics="true">Pteropera</emphasis>
species
</taxonomicName>
); its more or less pale green coloration on the hind femora (inner and outer sides of hind femora greenish yellow in
<taxonomicName id="ACC19DF84118A5B1AF39CE9D598D09DD" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="A227A17CB444A53D75C054917B13848E" italics="true">P. uniformis</emphasis>
</taxonomicName>
); male genitalia differ by its closed dorsal arch of cingulum (strongly open in
<taxonomicName id="98F3B4F16BFF9811378E8032FC5B6A31" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="661E2DFF1FB952AEB40D87275566305F" italics="true">P. uniformis</emphasis>
</taxonomicName>
); aedeagus horizontal apically, in line with valves (anteriorly sloping in
<taxonomicName id="72DC3E9443346880A0CF4A2B2118FF51" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="9420B2C34424CCFD31C307650181C7FE" italics="true">P. uniformis</emphasis>
</taxonomicName>
); female genitalia differ by egg-guide being slender (broad in
<taxonomicName id="A156C5DED003AAC366425432F7C71919" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="D0AEBD2796C88EDD0A4D673D839BA810" italics="true">P. uniformis</emphasis>
</taxonomicName>
); and basivalvar sclerites forming an obtuse angle (acute angle in
<taxonomicName id="1E1210AD735B32720C1CEF411907A246" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="D0FFAB2AC84F4A60399C5FE50C538AFC" italics="true">P. uniformis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="9C58C5142F820D74D99846FB0164E40C">
The new species is also similar to
<taxonomicName id="2D30D6FC79D6F1D60069358F8900B24A" authority="Donskoff, 1981" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="91918483AB3FC804EE1F56616D80B815" italics="true">Pteropera descampsi</emphasis>
<bibRefCitation id="C8E2C527E2453D8970414AFC998B0C8E" DOI="10.1080/21686351.1981.12278263" author="Donskoff M" issue="1" journalOrPublisher=")" pagination="33-88" refId="B4" refString="Donskoff M (1981) Les acridiens de la foret africaine. I. Revision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Societe entomologique de France (N. S.) 17 (1): 33 - 88. [51 figs] https: // doi. org / 10.1080 / 21686351.1981. 12278263" title="Les acridiens de la forêt africaine. I. Révision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Société entomologique de France (N. S." volume="17" year="1981">Donskoff, 1981</bibRefCitation>
</taxonomicName>
, from which it can be distinguished by the following characteristics: a pale basal band on the lateral lobes of the pronotum is absent but present in
<taxonomicName id="18F57FE47097229497B477BD5D7600A5" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="25B288C98F6048A8E2DEE14BD53E3375" italics="true">P. descampsi</emphasis>
</taxonomicName>
; bilobed male cerci, whereas male cerci are short in
<taxonomicName id="03F57C33E0B59CF064E2FD0B060634F8" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="66DC06D869026311AC5AA3086101DA95" italics="true">P. descampsi</emphasis>
</taxonomicName>
; the pallium and subgenital plate in males are slightly raised, whereas the apex of the subgenital plate is truncated in
<taxonomicName id="48A6E3A8DC586FCABDBCFB1B055D6CBC" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="1671B2B51D17114ED7DC816FDF6D4C7C" italics="true">P. descampsi</emphasis>
</taxonomicName>
; the apex of the aedeagus is horizontal, in line with valves, whereas the aedeagus is curved upwards, and the apex is divergent pallets in
<taxonomicName id="C0B4458038A38DBA3F9FE6815513CDD5" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="FB6C7114367570883B21971A9AA128D2" italics="true">P. descampsi</emphasis>
</taxonomicName>
; the dorsal arc of the cingulum is closed, whereas it is strongly open in
<taxonomicName id="FE1AC78680DA29495A308AFB897B0A73" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="64C2634C45A84034945546E7C24DA582" italics="true">P. descampsi</emphasis>
</taxonomicName>
; and the basivalvar sclerites of the female subgenital plate are described as an obtuse angle (acute angle in
<taxonomicName id="3EF47F2EEC84B33E7FC52155933CFE74" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="E39D18C45DC84292926571884C0906E1" italics="true">P. descampsi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERSCK" type="description">
<paragraph id="497601E0A1E12BC1EAEDDA2087E335B3">
<heading id="8CF78A96A76B32AF6D1F15FF5C19111D" reason="title">Description.</heading>
</paragraph>
<paragraph id="05E2EDB44C7FEB7D0A733E6427CAF532">
<emphasis id="017AF77CC5E7B2058C90BA0AF39A7196" bold="true">Male</emphasis>
: General coloration brown with pale green; body and legs with inconspicuous hairs, moderately rugous dorsally, smooth ventrally; eyes prominent; the large subocellar facial spot interrupted at the facial furrow, sometimes extending to the cheeks; antennae thin, filiform, longer than head and pronotum together; pronotum dark brown; dark longitudinal median band on pronotum disc present, wider than adjacent pale bands; basal pale bands on lateral lobes of pronotum absent; two incipient pale spots on the anterior margin of lateral lobes of pronotum; posterior margins of pronotum with or without incipient pale spots; median carina present and crossed dorsally by three sulci; lateral carinae absent; prozona longer than metazona; prosternal process short conical, compressed at its base; tegmina lobiform, only slightly reaching the third abdominal segment, lower half shiny black and upper half pale ochreous; mesosternal interspace open, ~ 1.3 × longer than wide; meso- and metathoracic episternites dark brown; front and middle legs, inner and outer sides of hind femora pale green, with the apical third gradually darkening toward the knee; knee dark orange; dorsal basal lobes of hind femora longer than ventral ones; upper margins of hind femora with fine teeth; hind tibiae dark green, basal ring absent; external apical spines of hind tibiae absent; male cerci bilobed, the inner lobe being twice shorter than the outer; subgenital plate obtuse to rounded in dorsal view; pallium and supra-anal plate of male slightly raised.
<emphasis id="BCA0605879AC6AF0514C5666E0FE7D25" bold="true" italics="true">Epiphallus</emphasis>
(Fig.
<figureCitation id="C423DBC74A006FF030677831C57E8869" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pteropera kennei sp. nov. A habitus image of a male under natural conditions B male frontal view C female frontal view D male lateral view E female lateral view F male dorsal view G female dorsal view H epiphallus dorsal view I phallic complex lateral view J phallic complex dorsal view K phallic complex ventral view L female subgenital plate M female spermatheca." figureDoi="10.3897/zookeys.1216.130270.figure12" httpUri="https://binary.pensoft.net/fig/1161000">12 H</figureCitation>
): small, bridge narrow, arched; lateral margins parallel; ancorae small, internally directed; lophi slender.
<emphasis id="CA80030CCBCF51EAB20F79577532F947" bold="true" italics="true">Phallic complex</emphasis>
(Fig.
<figureCitation id="70E085EE875FB84E80C56B199F7A7A35" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pteropera kennei sp. nov. A habitus image of a male under natural conditions B male frontal view C female frontal view D male lateral view E female lateral view F male dorsal view G female dorsal view H epiphallus dorsal view I phallic complex lateral view J phallic complex dorsal view K phallic complex ventral view L female subgenital plate M female spermatheca." figureDoi="10.3897/zookeys.1216.130270.figure12" httpUri="https://binary.pensoft.net/fig/1161000">12 I K</figureCitation>
): aedeagus small, short, curved; membranous apex of aedeagus, outside sheaths without sclerites, never filiform, and without ridge-like expansion; membranous apex of aedeagus outside sheaths, without sclerites, enlarged into a broad transverse lamina, never angular, never rolled up on itself; the dorsal arch of cingulum V-shaped, its apex acute, curved inwards; rami not bent, its lower part short; zygoma reduced; latero-ventral sclerites narrow; upper ectophallic sheath tight, short, slightly curved, with acute apex; upper eadeagus valve widened into a transverse blade; lower ectophallic sheath small, not enveloping the base of rami.
</paragraph>
<paragraph id="13598352E2323C7C16FF3B0D121BA4C7">
<emphasis id="41D7FA3B27297B34E1D455CD1FD62D5C" bold="true">Female</emphasis>
: As male, but larger; cerci short conical; valves of ovipositor narrow,&gt; 3.5 × longer than wide in coalescence position; subgenital plate (Fig.
<figureCitation id="AA61F5500DCA5C79984E0A21BF3123D1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pteropera kennei sp. nov. A habitus image of a male under natural conditions B male frontal view C female frontal view D male lateral view E female lateral view F male dorsal view G female dorsal view H epiphallus dorsal view I phallic complex lateral view J phallic complex dorsal view K phallic complex ventral view L female subgenital plate M female spermatheca." figureDoi="10.3897/zookeys.1216.130270.figure12" httpUri="https://binary.pensoft.net/fig/1161000">12 L</figureCitation>
) pentagonal, elongated, with truncated posterior margins; anterior apodemes narrow and short; egg-guide thin and long; ventral pockets of the vaginal floor large; copulatory bursa almost straight, gradually narrowing towards the front; bottom of the copulatory bursa close to the arc of the basivalvar sclerites; copulatory bursa above the basivalvar sclerites with a thick ventral gutter and membranous roof; each basivalvar sclerite barely curved, forming an obtuse angle; internal sclerite of the copulatory bursa short; the recurrent distal trunk of the lateral spermathecal diverticulum 3 × longer than the proximal trunk; the base of the spermathecal duct opening at the apex of the bursa; spermathecal ampulla narrowed at the apex; spermathecal duct very long; axial diverticle of the spermatheca almost as long as the lateral diverticulum (Fig.
<figureCitation id="1571C5A788CEA0FFF03DD765904857A1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pteropera kennei sp. nov. A habitus image of a male under natural conditions B male frontal view C female frontal view D male lateral view E female lateral view F male dorsal view G female dorsal view H epiphallus dorsal view I phallic complex lateral view J phallic complex dorsal view K phallic complex ventral view L female subgenital plate M female spermatheca." figureDoi="10.3897/zookeys.1216.130270.figure12" httpUri="https://binary.pensoft.net/fig/1161000">12 M</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBUCK" type="description">
<paragraph id="2100BF7B8342C8D885D4BA27FE1F64BE">
<heading id="B35DE2CEBC89D98F43CED550A5BC3B99" reason="title">Measurements.</heading>
</paragraph>
<paragraph id="D7F4DF9E96D99AA786C11E3B95031830">
Males (mm) (
<emphasis id="F9435A74FBE083F328E2C7C815710A0E" italics="true">n</emphasis>
= 20): total length of body 11.8119.83; length of pronotum 3.124.32; length of hind femur 11.2312.53; length of elytra 3.244.27. Females (mm) (
<emphasis id="DE781070100CC588F7E9D02EA8829B7A" italics="true">n</emphasis>
= 5): total length of body 21.0925.39; length of pronotum 4.595.37; length of hind femur 13.6215.12; length of elytra 3.654.88; length of ovipositor 1.973.27. Detailed information is shown in Table
<tableCitation id="10BC595C9F072FF2EAEB0CC2119FBB35" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements in millimeters (mm) of the examined Pteropera species currently known from Cameroon.">1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPUCK" type="etymology">
<paragraph id="414381051EFEE33F2FA3C6E3A736E9E3">
<heading id="9BA4C463E3342F25220E9B9FDBA4530D" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="DAA9DB804FFC57B5669D1A6E23FFD211">
The species was named in honor of Professor Martin Kenne in recognition of his work and scientific contribution to the biodiversity of insects in
<collectingCountry id="2B046D8995BC658A4BFFF409F1B596C3" name="Cameroon">Cameroon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUUCK" type="habitat">
<paragraph id="EAE251388158A8924208406B77ADEA33">
<heading id="FDA4B2338CB65241D2C41B7676ABA1A6" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="319F3FDA8E67FC55E30EE7B7A5D9D19F">
Dense evergreen forest in the
<collectingCountry id="A2F79E29246544BF7D8CF2A0E4EEF32D" name="Democratic Republic of the Congo">Congo</collectingCountry>
Basin, Dja Biosphere Reserve, south
<collectingCountry id="AE6E46D44426C6F5A00EF40DF5BCCAEF" name="Cameroon">Cameroon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZUCK" type="distribution">
<paragraph id="7BF466A13918D44EC6786102BC7BDA7B">
<heading id="D2B4DDCA70EC9A62002C3D97132D1896" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="768C206AAF23E256EE6FABDB32E0FA6D">
<collectingCountry id="933DCC34D75BD592837284EFFDBAACEF" name="Cameroon">Cameroon</collectingCountry>
, Somalomo in the Dja Biosphere Reserve and Deng-Deng National Park (Fig.
<figureCitation id="1BDC1E18169028F35C9E1232EC600BD1" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of Pteropera species in African rainforests." figureDoi="10.3897/zookeys.1216.130270.figure17" httpUri="https://binary.pensoft.net/fig/1161005">17 B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,454 @@
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<mods:title id="D69D249B54402B04A004C35D428DBC29">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<subSubSection id="4899A9AC8A444D31ADA8FEC7D7D0BEB1" type="nomenclature">
<paragraph id="6DC84B9D4CCC7C5950D8522441A22D9A">
<taxonomicName id="C9B5BBD4BCC4B98A318381F463949790" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="liupanshui" status="sp. nov.">
<emphasis id="FDBF38DE3F752B5194A9DC75A97B0674" italics="true">Belisana liupanshui</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="84CA1448E504A4A134B0FBC4B95EA91A" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8F551FBB422E51D000B0C240B6CEEAAC" type="description">
<paragraph id="0D8D78DD6E65B118869FE3DAD24CC101">
<figureCitation id="1525D0FE0A877FC21F20B328D022E914" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">Figs 4</figureCitation>
,
<figureCitation id="C1236654CEDCCEA5359B4400DFC92022" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5</figureCitation>
,
<figureCitation id="9FE620DDBE51BFAE775593B27C6717EE" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 C, D</figureCitation>
,
<figureCitation id="1F07B1031E0D61BFB7D38AC1642CA099" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQFAG" type="type material">
<paragraph id="71A98CA76B7A03B4488E8544A9C7C6FE">
<heading id="2608CB5B734A8713F706E42A2962CE81" reason="title">Type material.</heading>
</paragraph>
<paragraph id="4E5752E24DD77B0D41AD8CBED375838F">
<materialsCitation id="CC50D20051049D93145524957EFD9B33" collectingDate="2007-04-13" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Pan County" elevation="1468" latitude="25.77545" location="Biyun Cave" longLatPrecision="1" longitude="104.63797" municipality="Liupanshui" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="6B04D45260A8436AD4DF6C6AF78A486A" bold="true">
<emphasis id="D276D23952883B5403234F57BEF5B8AE" bold="true" italics="true">
<typeStatus id="EA9AA53BC3D3F77F11ED37DEF836E536" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="D8E1ABF83339A02D8775DF9E65472D1F" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="FE156E73BCBDDB8FC426691A64F3F0E2" count="1" type="male"></specimenCount>
;
<collectingRegion id="28D48CA47F65DE6DE008D2FBB0CAB2D9" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingMunicipality id="75271D86FBCE91833B0B8240A1B0B59C">Liupanshui</collectingMunicipality>
,
<collectingCounty id="D229C65691F23584A61A2D98B1CD92C3">Pan County</collectingCounty>
,
<location id="691C3578037771BB65B0A7A48387E40D" LSID="urn:lsid:plazi:treatment:712BAFCCAD3F56E596F490BDE5164A50:691C3578037771BB65B0A7A48387E40D" country="China" county="Pan County" latitude="25.77545" longLatPrecision="1" longitude="104.63797" municipality="Liupanshui" name="Biyun Cave" stateProvince="Guizhou">Biyun Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5FAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[104.637967,25.775450]}">
<geoCoordinate id="024997FBFE95AFA324EEF83C575791BF" degrees="25" direction="north" minutes="46.527" orientation="latitude" precision="1" value="25.77545">25 ° 46.527 ' N</geoCoordinate>
,
<geoCoordinate id="8FF73DB1D586E13EBC3DD947F46A1C99" degrees="104" direction="east" minutes="38.278" orientation="longitude" precision="1" value="104.63797">104 ° 38.278 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="6F9888C94E21D647F97963E540A531F1" metricMagnitude="3" metricUnit="m" metricValue="1.468" unit="m" value="1468.0">
<elevation id="02C708B385B014D2F272161FC38C08FD" metricMagnitude="3" metricUnit="m" metricValue="1.468" unit="m" value="1468.0">1468 m</elevation>
</quantity>
;
<date id="B01A95B04622E0209C224FF31BE7AFA3" value="2007-04-13">
<collectingDate id="EF2F1A4F84552A3F5527BB2B24328223" value="2007-04-13">13 Apr. 2007</collectingDate>
</date>
;
<collectorName id="B35AAD0C9D76797C79C45D405153257F">J. Liu</collectorName>
&amp;
<collectorName id="10C42A5BD7863DFF8C95BBD845F15689">Y. Lin</collectorName>
leg.;
<named-content id="8EB989026258B8EC429B5406A7A287BA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="0E40420702B321A2ADCD8532F0FFBC5D" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="1EEAA0A501370AF24CD92AA4281153F1">Ar</collectorName>
45181
</materialsCitation>
.
<materialsCitation id="C26F8C60045453602DB5882323B0EADF" collectingDate="2007-04-13" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Pan County" elevation="1468" latitude="25.77545" location="Biyun Cave" longLatPrecision="1" longitude="104.63797" municipality="Liupanshui" specimenCount="5" specimenCount-male="5" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="C71D53FA3D7E87109A24B6E2693A147F" bold="true">
<emphasis id="B5B3B29B85C7F6989631C2B5009D4B4A" bold="true" italics="true">
<typeStatus id="003871FCC53317FD68BABD4A44B517D0" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="7F206B3EFFF60F8E2D18EF27C0216B4B" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="B1F54AE602EB3307D1D42028BB020400" count="5" type="male">5 ♂</specimenCount>
; same data as for holotype;
<named-content id="E20278925E1C10AEA98AECF25CD3CCDF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="4E4438925AD47187D5BCC47448C2DD0E" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4518286
</materialsCitation>
<materialsCitation id="821E1533BFEE681E68A6D404F8096497" collectingDate="2007-04-13" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Pan County" elevation="1468" latitude="25.77545" location="Biyun Cave" longLatPrecision="1" longitude="104.63797" municipality="Liupanshui" specimenCount="7" specimenCount-female="7" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="D9E8A453007C697E7510FB43BAB4D990" count="7" type="female">7 ♀</specimenCount>
; same data as for holotype;
<named-content id="02AD61A242C299FB32D3183F42D04F32" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="1123D24E4AC11AB3F01DCE9A3CE64CD3" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4518793
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETGAG" type="diagnosis">
<paragraph id="AADA308AB34FCF14BE6964AD9069D11B">
<heading id="066688C3CE4C1A69550DBA48671E334B" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="8A918459E692D7CD2F9A719DBE774F7A">
The new species resembles
<taxonomicName id="2DF17F95FEC28E4D52125495159BDCD8" authority="Zhang, Li &amp; Yao, 2024" authorityName="Zhang, Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="jiuxiang">
<emphasis id="FBED8F1D917220A16B8819EE0D48C230" italics="true">B. jiuxiang</emphasis>
<bibRefCitation id="B604F2E90649F2B0C3EAE8BF3F68125E" DOI="10.3897/zookeys.1209.127951" author="Zhang L &amp; Wang Y &amp; Li S &amp; Zhang X &amp; Yao Z" journalOrPublisher="ZooKeys" pagination="315-330" refId="B23" refString="Zhang L, Wang Y, Li S, Zhang X, Yao Z (2024 a) Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China. ZooKeys 1209: 315 - 330. https: // doi. org / 10.3897 / zookeys. 1209.127951" title="Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China." volume="1209" year="2024">Zhang, Li &amp; Yao, 2024</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="889317E359071D85C9556B614895DFC3" DOI="10.3897/zookeys.1202.121633" author="Zhang L &amp; Wu Z &amp; Li S &amp; Yao Z" journalOrPublisher="ZooKeys" pagination="255-286" refId="B24" refString="Zhang L, Wu Z, Li S, Yao Z (2024 b) Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China. ZooKeys 1202: 255286. https://doi.org/10.3897/zookeys.1202.121633" title="Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China." volume="1202">Zhang et al. 2024 b</bibRefCitation>
: 261, figs 4 A D, 5 A H, 18 C, D, 20 C, D) by having similar male chelicerae (distal apophyses directed towards frontally, but tips pointing inwards; Fig.
<figureCitation id="C1E76D883037462ADC5E63AE85A7C7ED" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 D</figureCitation>
) and epigyne (epigynal pockets on median part of epigynal plate, epigynal plate posteriorly straight; Figs
<figureCitation id="1CCC0B638415EE2489C53AB3EA4E9395" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 A</figureCitation>
,
<figureCitation id="048A643D210A7A4106A77EE7749BB6A4" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 C</figureCitation>
), but can be distinguished by procursus with prolatero-subdistal and subdistal membranous processes (arrows 1,
<quantity id="EFC1B96E26ED66D32F953D195EBAEFF7" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="A61E636B14D349E202879F14D1979035" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 C</figureCitation>
,
<figureCitation id="829268CCC07B7F07CCA145765BB7873A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 C</figureCitation>
vs absent) and nearly rectangle dorso-subdistal membranous process (arrow
<quantity id="15331C3EE5D4377ECFA265C6A05948C6" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
Figs
<figureCitation id="D815795C1AF5E77D420047816EE50DCF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 C</figureCitation>
,
<figureCitation id="BDDF15F772F2B942A9B2866938FBA02D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 C</figureCitation>
vs angular), by bulbal apophysis with angular subdistal apophysis (arrow in Fig.
<figureCitation id="FCC5048EC3AC3E82C856077108E5349B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 C</figureCitation>
vs absent), and by vulval pore plates anteriorly narrow and posteriorly wide (
<abbrev id="ABBRID0EIIAG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="89F96CA3F4FB65C749D4F2EE28038359" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 B</figureCitation>
,
<figureCitation id="B1395F1BDD43F16287260CC6742E35F8" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 D</figureCitation>
vs nearly quadrilateral).
</paragraph>
<caption id="B8CD2AA21BE39117F222BAF9CEE43D59" ID-DOI="10.3897/zookeys.1216.132561.figure4" ID-arpha="E702CAC1-FD7E-5170-A72E-478483A649AD" httpUri="https://binary.pensoft.net/fig/1161049" startId="F4">
<paragraph id="E7FDF7BB1D596074B96B3627A6C0E990">
<label id="1BEECAA0CB22C21302CFE85A56B667D6">Figure 4.</label>
</paragraph>
<paragraph id="8141F3CCDA81B87B83EF5CB7B1AA2EA3">
<taxonomicName id="ED3B19467C7783AC0477CB4DFA73F456" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="liupanshui" status="sp. nov.">
<emphasis id="170FAC3DA57CE760402451C0087FB786" italics="true">Belisana liupanshui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="981678FCF51F35BB4AAD87D0E4F58BAA" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="924D3000F4EE5D46B985E1735B7D1F13" bold="true">A, B</emphasis>
palp (
<emphasis id="977ED521C733295D56E17CC3E13E0789" bold="true">A</emphasis>
prolateral view
<emphasis id="B03F53BBBE3856AEDD34A691302096F8" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="99F685D6758BCF3C6B8EDECC0B7DFAA5" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="34A5E06B405FF334A7152ED2496A1973" bold="true">C</emphasis>
prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process
<emphasis id="1A6C4039CE812ED29AB421C39F161066" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb,
<abbrev id="ABBRID0ETJAG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0EXJAG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="F6E6DF9004037792886BC5A3D5E5DDDE" bold="true">A, B</emphasis>
); 0.05 (
<emphasis id="80524C6525D1477A0A249F53E2CCD69E" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="14AAB5F5B4D402D80156C7D881C84F1E" ID-DOI="10.3897/zookeys.1216.132561.figure5" ID-arpha="A6AE6C75-0E8A-5AB3-A8EB-DA1C1B109061" httpUri="https://binary.pensoft.net/fig/1161050" startId="F5">
<paragraph id="4DCEA1B8B359F3B716B5A7A06B01D3EF">
<label id="7B81DCE26EDE2607DDCE48F26BB0BE7A">Figure 5.</label>
</paragraph>
<paragraph id="F62ACF19167A1F35418C1BA68B7FE08F">
<taxonomicName id="62D96D6BF326781E1B82714A34AED650" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="liupanshui" status="sp. nov.">
<emphasis id="215CA534E117FD7F4D41BBE067235225" italics="true">Belisana liupanshui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D76997A5AB773FAC9EACBC88DFCA3C64" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="4C30E13DA29C55796F3EFA682D19B62D" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="F4C2E307FD8AE9420423B91E4555D742" bold="true">A, B, G, H</emphasis>
)
<emphasis id="19FC24403C0AAE792CF9E1F87E2A3740" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="BEE69759B826CE6F79DFA3A41EE5B285" bold="true">B</emphasis>
vulva, dorsal view, arrow points at lateral sclerite
<emphasis id="B064F72DFA904979F05FD4C4FF043A04" bold="true">C</emphasis>
bulb, prolateral view, arrow points at subdistal apophysis
<emphasis id="B76B7E910E79AF7FCFED00A295F60DEE" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="FEBA3338C93DC6402F5ED4A4A539FF0F" bold="true">E H</emphasis>
habitus (
<emphasis id="1AAA879AD37049CD4DB418075CEF4708" bold="true">E, G</emphasis>
dorsal view
<emphasis id="5FD123114608C14066B9CEBD77F01D6C" bold="true">F</emphasis>
lateral view
<emphasis id="CF847B9DF70D6BC4A9F827F6B6098B92" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0ELLAG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EPLAG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0ETLAG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EXLAG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0E2LAG" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0E6LAG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 (
<emphasis id="7319FBE3D7A5469EE8FF22AD2CE99C97" bold="true">A D</emphasis>
); 0.50 (
<emphasis id="211A3D98D3815FFA0EA0B28A5DF69953" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EMMAG" type="description">
<paragraph id="32B7742ADBDA310DDE93DA3E03AE3EDC">
<heading id="D3C6FC3500CFBFFFF4D016D0E826FFE6" reason="title">Description.</heading>
</paragraph>
<paragraph id="E9D8FFE9E55F9419A8B030C1F60FA481">
<emphasis id="1D8FF137C7624D2888B7F706A756364F" bold="true">Male</emphasis>
(
<emphasis id="E7E1BC355F42AB366160DED72EAC2C5B" bold="true">
<typeStatus id="9A95CB83E71FEDA60EAB1B0C2966A140" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.71 (1.79 with clypeus), prosoma 0.63 long, 0.67 wide, opisthosoma 1.08 long, 0.81 wide. Leg I: (4.04, 0.29, 4.50, 5.96, ), leg II: 11.08 (2.80, 0.30, 2.97, 3.92, 1.09), leg III: 7.98 (2.22, 0.25, 1.98, 2.72, 0.81), leg IV: 10.20 (3.00, 0.26, 2.63, 3.45, 0.86); tibia I
<abbrev id="ABBRID0EWMAG" xlink_title="length/diameter">L / d</abbrev>
: 63. Eye interdistances and diameters:
<abbrev id="ABBRID0E1MAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E5MAG" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0ECNAG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EGNAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EKNAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.51 / 0.44. Habitus as in Fig.
<figureCitation id="83AC0C15117BFBDAA33D5A8A5DD0166D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, with brownish radiating marks; clypeus brownish; sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0ESNAG" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="CCD45BB41D1C6955C9DE335E97501F01" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.17;
<abbrev id="ABBRID0E1NAG" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="633D8A8038538CC08F3B89C5B6EF7540" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 D</figureCitation>
). Palp as in Fig.
<figureCitation id="2027EF5C5B05DC274EAB7E10E970D985" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="ED61317051377362D15AE33FF60739BE" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="8DFE1A7840E2D130FFD03DDC9BB6971F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="A99075542A46DBCBB98ECE11B4F68ABE" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="1FC710F1273E69881D76B675DB57A3C3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 B</figureCitation>
); procursus simple proximally but complex distally, with prolatero-subdistal membranous process (arrow
<quantity id="20C05033E6A069AA8E69110DC6672EC0" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="AB4DF5AF4FDC8723191B8A5918846E63" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 C</figureCitation>
,
<figureCitation id="B45C78A2D9910258014B95F363AD801A" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 C</figureCitation>
), sclerotized distal apophysis (arrow
<quantity id="50A0588F042C96DA0DC21E56C91468FE" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="753B3EB368E5C5869153302AD1DA504A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 C</figureCitation>
,
<figureCitation id="AA3FB579DC3FAD559DD856751F3A3A8D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 C</figureCitation>
), subdistal membranous process (arrow
<quantity id="2D5370EEE6139AA3A78127FDE037B1A3" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="79092CA5A8787BB3E924086B1BBA9B5C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 C</figureCitation>
,
<figureCitation id="8768EFD28E3086A31F16FB165A73EC38" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 C</figureCitation>
), dorso-subdistal membranous process (arrow
<quantity id="35E53E836435ACD82E0F9FA97B032F00" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
Figs
<figureCitation id="F37D97A2ECDAA923B62D820D5F44BB6C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 C</figureCitation>
,
<figureCitation id="AF2B340EAA2C0CD5D46D4963BC667159" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 C</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="E4844B1FCB236E707BBD5EC93E4A72C5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana liupanshui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal membranous process, arrow 2 points at sclerotized distal apophysis, arrow 3 points at subdistal membranous process, arrow 4 points at dorso-subdistal membranous process D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure4" httpUri="https://binary.pensoft.net/fig/1161049">4 D</figureCitation>
,
<figureCitation id="EEA71220253C5C57477E2CCF99009D01" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 D</figureCitation>
); bulb with hooked apophysis bearing angular subdistal apophysis (arrow in Fig.
<figureCitation id="6349FB5380D4FC6DF5010B050324D7EC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="1104BAF421D0D6CB8C6DB9F7B4398BE9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 C</figureCitation>
). Retrolateral trichobothria on tibia I at 4 % proximally; legs with short vertical setae on metatarsi.
</paragraph>
<paragraph id="9A6A6D4F55C382BD85448DB17CEFEAEE">
<emphasis id="6072EFC497E005015A72560DF7CB1D25" bold="true">Female</emphasis>
(
<emphasis id="FB5432FE0A08D50E065719DE6C8006D1" bold="true">
<typeStatus id="9DD348416A5A11A712B10516381AF35A" type="paratype">paratype</typeStatus>
</emphasis>
,
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<collectionCode id="6EF17E42322B4B259E8B8375AF6F845D" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45187): Similar to male, habitus as in Fig.
<figureCitation id="1552EE826213FF919B8E05AEE784B7C4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 G, H</figureCitation>
. Total length 2.53 (2.65 with clypeus), prosoma 0.67 long, 0.75 wide, opisthosoma 1.86 long, 1.80 wide; tibia I: 4.05; tibia I
<abbrev id="ABBRID0EMQAG" xlink_title="length/diameter">L / d</abbrev>
: 51. Eye interdistances and diameters:
<abbrev id="ABBRID0EQQAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EUQAG" xlink_title="posterior median eye">PME</abbrev>
0.11,
<abbrev id="ABBRID0EYQAG" xlink_title="posterior median eye">PME</abbrev>
0.07,
<abbrev id="ABBRID0E3QAG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EARAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.52 / 0.46. Epigyne simple and flat, posteriorly straight, with pair of median pockets 0.20 apart (
<abbrev id="ABBRID0EERAG" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="9AA48F5F8BC2A0D3A4A3577C4FB1B322" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 A</figureCitation>
,
<figureCitation id="D1D07C093A5D2E94D4EDCC66C873087A" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 C</figureCitation>
). Vulva with ridge-shaped anterior arch bearing pair of angular lateral sclerites (arrow in Figs
<figureCitation id="2E4DFDECF8984C8A94D5D3B5EB342F03" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 B</figureCitation>
,
<figureCitation id="28B7E9AA7D8BE0D3007837F8677FCC05" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 D</figureCitation>
), and pair of anteriorly narrow and posteriorly wide pore plates (3 × longer than wide,
<abbrev id="ABBRID0EYRAG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="DC35E15C3454A525276B0B5A77C44FB0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana liupanshui sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at lateral sclerite C bulb, prolateral view, arrow points at subdistal apophysis D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.50 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure5" httpUri="https://binary.pensoft.net/fig/1161050">5 B</figureCitation>
,
<figureCitation id="CE8BAE064C33E1F1F120DD8DDA1BE054" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EESAG" type="variation">
<paragraph id="D953FF495054EA2C18F4181221B96C9E">
<heading id="7A92977E0F3922046931AD8075DDEDCD" reason="title">Variation.</heading>
</paragraph>
<paragraph id="B3347591B5C5FCFECA7A37C4C8D03BC2">
Tibia I in
<specimenCount id="355379DF6E0A0C87589B635A6D707B64" count="5" type="male">five male</specimenCount>
<typeStatus id="2C72787974D1B7C820567FB8687097EC" type="paratype">paratypes</typeStatus>
(
<named-content id="2A8B61300CFA028CAE46558A4DFA18F1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="5AD4A990CC1140397D1D2B680678A6AD" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4518286): 4.50, 4.55, 4.62, 4.65, 4.70. Tibia I in the other
<specimenCount id="B8E45A022DBD210E6EF363979100BCF2" count="6" type="female">six female</specimenCount>
<typeStatus id="9A539465CBEE62F01ED09F27B8A17795" type="paratype">paratypes</typeStatus>
(
<named-content id="1D6A30390D720CA8F192BC04C34F48CC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="A13C4BF642A7371CA313E2EC2C25B1B8" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4518893): 3.563.85.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESSAG" type="habitat">
<paragraph id="94CB0BACC8C3E255CD84689C65CAB04F">
<heading id="89255928F357ED47DD7CB2020F6693A8" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="1B54661AADE617FA1D17D88C96912E0E">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EXSAG" type="distribution">
<paragraph id="2AA04B5F688569AEBB9BE71A517C54CD">
<heading id="F926A53C5B19A12B25ED5C952B9A80D3" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="C1150A9463AAB4E93D6D57565D3B5569">
<collectingCountry id="8627D060A86B2A30213E699AF78FDA82" name="China">China</collectingCountry>
(
<collectingRegion id="61F94E39369DE4EB91D27D5129EDF531" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="81F9692D30F6B8106A6213EB0153D41E">type</typeStatus>
locality; Fig.
<figureCitation id="5648E21C7D30CD944ABB67DD805DB007" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBTAG" type="etymology">
<paragraph id="93A7A4340BC561C47F82974678B04245">
<heading id="B5853D0C5D0797D79A851D5CD9D7137B" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="899940863F4CA085FFE4241A6932D052">
The specific name refers to the
<typeStatus id="A00EAE2D9C70CF6DAA74DB3BD8A0F0FB">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,277 @@
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<mods:title id="0C840D2855F6E98C0B454549B6351472">Taxonomic review of the grasshopper genus Pteropera Karsch, 1891 (Orthoptera, Acrididea, Catantopinae) with description of three new species and a preliminary phylogeny of the Cameroonian species</mods:title>
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<mods:namePart id="C5E2D30EA78845C7AE07B48B4EC89CE8">Duressa, Tarekegn Fite</mods:namePart>
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<mods:affiliation id="DEB79BA266B5AF0A601B24CB2F3F176B">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany &amp; School of Plant Sciences, College of Agriculture and Environmental Sciences, Haramaya University, Dire Dawa, Ethiopia</mods:affiliation>
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<treatment id="B4E6187886E45B7EB7929315180E6D04" ID-arpha="B4E61878-86E4-5B7E-B792-9315180E6D04" LSID="urn:lsid:plazi:treatment:B4E6187886E45B7EB7929315180E6D04" httpUri="http://treatment.plazi.org/id/B4E6187886E45B7EB7929315180E6D04">
<subSubSection id="D241E8090BBDFC93671E69CC9801B60C" type="nomenclature">
<paragraph id="24AA9BF0085B91166146A7D97B27F7BF">
<taxonomicName id="AD90172383B12853ECED04BBCA7AFD8C" ID-CoL="4Q25Z" authority="Donskoff, 1981" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="descampsi">
<emphasis id="769CE469138C57B99BD1BCB8A7CB27E5" italics="true">Pteropera descampsi</emphasis>
<bibRefCitation id="C0845508664055D399F9AF434C7ECDAC" DOI="10.1080/21686351.1981.12278263" author="Donskoff M" issue="1" journalOrPublisher=")" pagination="33-88" refId="B4" refString="Donskoff M (1981) Les acridiens de la foret africaine. I. Revision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Societe entomologique de France (N. S.) 17 (1): 33 - 88. [51 figs] https: // doi. org / 10.1080 / 21686351.1981. 12278263" title="Les acridiens de la forêt africaine. I. Révision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Société entomologique de France (N. S." volume="17" year="1981">Donskoff, 1981</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A8EE2C1C16386213F47D78501899AE5C" type="description">
<paragraph id="9022F93E08207D8123ED643AA48DC6BA">
<figureCitation id="57076072665D1C995C75CCBCF87AE97D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Images of holotypes and allotypes of Pteropera species in lateral view A P. carnapi (holotype ♂) B P. carnapi (paratype ♀) C P. congoensis (holotype ♂) D P. congoensis (allotype ♀) E P. cornici (holotype ♂) F P. cornici (allotype ♀) G P. descampsi (holotype ♂) H P. descampsi (allotype ♀) I P. descarpentriesi (holotype ♂) J P. descarpentriesi (allotype ♀)." figureDoi="10.3897/zookeys.1216.130270.figure3" httpUri="https://binary.pensoft.net/fig/1160991">Figs 3 G, H</figureCitation>
,
<figureCitation id="4919CDE78129BD1A68E53B26D5F0DA81" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Images of holotypes, allotypes and paratype of Pteropera species in dorsal view A P. carnapi (holotype ♂) B P. carnapi (paratype ♀) C P. congoensis (holotype ♂) D P. congoensis (allotype ♀) E P. cornici (holotype ♂) F P. cornici (allotype ♀) G P. descampsi (holotype ♂) H P. descampsi (allotype ♀) I P. descarpentriesi (holotype ♂) J P. descarpentriesi (allotype ♀)." figureDoi="10.3897/zookeys.1216.130270.figure8" httpUri="https://binary.pensoft.net/fig/1160996">8 G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXDBK" type="type material">
<paragraph id="94C51A7DD9BA24AC1B033E6E667ABE24">
<heading id="83566C70E3D9557770F5CE422136EF73" reason="title">Type materials examined.</heading>
</paragraph>
<paragraph id="2F8432563608E990AA95B75F7417588D">
<materialsCitation id="38EE568F792C0402312E8BA1182ADD9D" collectingDate="1975-11-02" collectingDateMax="1975-11-04" collectingDateMin="1975-11-02" collectionCode="MNHN" collectorName="M. Descamps" country="Cameroon" latitude="3.8483167" location="Yaounde" longLatPrecision="1" longitude="11.502216" municipality="Ongot" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B87886753ED1ED1736C73B16AE961C7B" bold="true">
<emphasis id="A126A70DD9E5EF237CD364B119FE4F77" bold="true" italics="true">
<typeStatus id="65F6B37E9AE24F6A96384CC2818461EC" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="542341CF3228265C88F31ECE2A59336B" name="Cameroon">Cameroon</collectingCountry>
<specimenCount id="A132331A00C18D02B8B984DB124CE4F1" count="1" type="male"></specimenCount>
;
<location id="62B73B7DB4DA86D291D8D64B0DC6D038" LSID="urn:lsid:plazi:treatment:B4E6187886E45B7EB7929315180E6D04:62B73B7DB4DA86D291D8D64B0DC6D038" country="Cameroon" latitude="3.8483167" longLatPrecision="1" longitude="11.502216" municipality="Ongot" name="Yaounde">Yaoundé</location>
,
<collectingMunicipality id="77F5833AA9FBEFC2F03990AD13D7A9CA">Ongot</collectingMunicipality>
[Onguot];
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFEBK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[11.502217,3.848317]}">
<geoCoordinate id="5E7E05FE3088100EBA1A5B6D978907F6" degrees="3" direction="north" minutes="50.899" orientation="latitude" precision="1" value="3.8483167">3 ° 50.899 ' N</geoCoordinate>
,
<geoCoordinate id="84F88ECD3BE1FF93CE439D1AF56022BF" degrees="11" direction="east" minutes="30.133" orientation="longitude" precision="1" value="11.502216">11 ° 30.133 ' E</geoCoordinate>
</named-content>
;
<date id="4A632655476A1A57AD68E4D0600D6F31" value="1975-11-02" valueMax="1975-11-04" valueMin="1975-11-02">
<collectingDate id="34B7E4529EFC22A0FD7E0D5B5CC77AC7" value="1975-11-02" valueMax="1975-11-04" valueMin="1975-11-02">24 Nov. 1975</collectingDate>
</date>
;
<collectorName id="CCA16E14343E050501322A9617AFD7B4">M. Descamps</collectorName>
leg.;
<named-content id="5E08BCB79597810C598834B3810A9021" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum National d´Histoire Naturelle Paris, France">
<collectionCode id="4A81F01B52FD00F084F8523EAC2B81E2" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
,
<named-content id="FD517CED57AF4253051DEBFEB1D3F012" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum National d´Histoire Naturelle Paris, France">
<collectionCode id="4420BF99943DBF301F483282EF52D0DF" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
- EO-CAELIF 11456.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESEBK" type="material">
<paragraph id="1F403CC95E8A63A7AD3E84A69541E40B">
<heading id="19EE6CA10877D1AF306BCA7FBB8B7611" reason="title">Other material examined.</heading>
</paragraph>
<paragraph id="20E1F6CCB72317F64275F44DD8000A34">
<collectingCountry id="5EE7944240EBD82713640C84AD104E83" name="Cameroon">Cameroon</collectingCountry>
<materialsCitation id="E3F1EF67A4FFB29FFD176E7E9CE6EC74" collectingDate="2020-06-15" collectionCode="R, SMNK" collectorName="Ongot &amp; Yetchom Fondjo &amp; Nzoko Fiemapong" country="Cameroon" latitude="3.8586166" location="Cameroon" longLatPrecision="1" longitude="11.372784" specimenCount="7" specimenCount-female="3" specimenCount-male="4">
<specimenCount id="25C8EBB448BFDA86C96CE1E923732C95" count="4" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="961391C331A533FF099B0CDC4957C18E" count="3" type="female">3 ♀♀</specimenCount>
;
<collectorName id="A0C6B74952B0F0550ACC5B9AC66D55DB">Ongot</collectorName>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2EBK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[11.372783,3.858617]}">
<geoCoordinate id="2FFA20EC4ADC9CB98B6914104D23DBD2" degrees="3" direction="north" minutes="51.517" orientation="latitude" precision="1" value="3.8586166">3 ° 51.517 ' N</geoCoordinate>
,
<geoCoordinate id="2FBED81B6399D4B24C96DBEBF67640A6" degrees="11" direction="east" minutes="22.367" orientation="longitude" precision="1" value="11.372784">11 ° 22.367 ' E</geoCoordinate>
</named-content>
;
<date id="08D5BABA61B5C903A0D15F8DE222C8B8" value="2020-06-15">
<collectingDate id="CF8B889F8C1ACA3D1019B4F214BD309C" value="2020-06-15">15 Jun. 2020</collectingDate>
</date>
; J. A.
<collectorName id="A6C9493D183562D3A5B57C0D33D38A06">Yetchom Fondjo</collectorName>
&amp; A.
<collectionCode id="05E38914F5E17D055E6FB6AFD3006E70" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectorName id="35CEC68FAFCD36037618FBC05AEE20B6">Nzoko Fiemapong</collectorName>
leg.;
<named-content id="2D9841A1AC51E3B232623A59491E2617" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="5D7450647CC0C12721CA80AF45C32908" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
.
<collectingCountry id="123044112A05F0FECE93091B3576B799" name="Cameroon">Cameroon</collectingCountry>
</materialsCitation>
<materialsCitation id="24E469108588A7BEBCC1E3A063ECC878" collectingDate="2021-08-18" collectionCode="SMNK" collectorName="Meyomessala &amp; J. A. Yetchom Fondjo" country="Cameroon" latitude="3.1071835" location="Cameroon" longLatPrecision="1" longitude="12.24505" specimenCount="1" specimenCount-female="1">
<specimenCount id="4D9C76B3B1BB5948956F95EFF185D125" count="1" type="female">1 ♀</specimenCount>
;
<collectorName id="4EAB395A197A2DC7E09B2805188A7343">Meyomessala</collectorName>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHFBK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[12.245050,3.107183]}">
<geoCoordinate id="CA7A07ABCBA677B5208A697459B922F8" degrees="3" direction="north" minutes="6.431" orientation="latitude" precision="1" value="3.1071835">3 ° 6.431 ' N</geoCoordinate>
,
<geoCoordinate id="C60451F618351E1C2DF5F43BE9FAA6D0" degrees="12" direction="east" minutes="14.703" orientation="longitude" precision="1" value="12.24505">12 ° 14.703 ' E</geoCoordinate>
</named-content>
;
<date id="2DDF1BE73FFE6BAEC210F420FA13B552" value="2021-08-18">
<collectingDate id="F460B48B10DDEFD83CE12CB32DFC7957" value="2021-08-18">18 Aug. 2021</collectingDate>
</date>
;
<collectorName id="B6EF02133B803EB752EEA7241E46C69C">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="52AF79C1A284C260DE3D9B1AC572D5AE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="BA67D04B835E84225366E5769C08E078" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
).
<collectingCountry id="0E13B492474C9F94DA1DFF9B0E575D79" name="Cameroon">Cameroon</collectingCountry>
</materialsCitation>
<materialsCitation id="B9AB51D2CE843C066E4E84A0259CA628" collectingDate="2021-12-05" collectionCode="SMNK" collectorName="Ongot &amp; J. A. Yetchom Fondjo" country="Cameroon" latitude="3.8586166" location="Cameroon" longLatPrecision="1" longitude="11.372784" specimenCount="5" specimenCount-female="5">
<specimenCount id="D33B0C975AE85F6425F26D2FFDA6A758" count="5" type="female">5 ♀♀</specimenCount>
;
<collectorName id="517ED1FB23F603BB43D2A9C5F00A057C">Ongot</collectorName>
;
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<geoCoordinate id="7267CE352C4D97A6A7F56A1E65FDCC30" degrees="3" direction="north" minutes="51.517" orientation="latitude" precision="1" value="3.8586166">3 ° 51.517 ' N</geoCoordinate>
,
<geoCoordinate id="A61114968AB58DF964A9CF8B0B88865F" degrees="11" direction="east" minutes="22.367" orientation="longitude" precision="1" value="11.372784">11 ° 22.367 ' E</geoCoordinate>
</named-content>
;
<date id="EACB8D1A273227642ED0B65EAA1C513D" value="2021-12-05">
<collectingDate id="AF4339C2FD90F2B611A199137CEB568C" value="2021-12-05">5 Dec. 2021</collectingDate>
</date>
;
<collectorName id="98884DC84E6F8F8BE0B222DDD8FEDCF0">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="AF8A79325DAF212A11900A759DF02993" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="BBF1B808FE8F495E35612792DB1010E9" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
.
<collectingCountry id="E11D0FA9826287ECE603B16D08EF1B86" name="Cameroon">Cameroon</collectingCountry>
</materialsCitation>
<materialsCitation id="A2554EF28110B3176A6FF6DB2BD00114" collectingDate="2022-03-20" collectionCode="R, SMNK" collectorName="Ongot &amp; Yetchom Fondjo &amp; Nzoko Fiemapong" country="Cameroon" latitude="3.8586166" longLatPrecision="1" longitude="11.372784" specimenCount="8" specimenCount-female="2" specimenCount-male="6">
<specimenCount id="7704CB38DF7DED950FCDCD62C7FE5817" count="6" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="FE5CFFA726B53161D9E84B29E5876A31" count="2" type="female">2 ♀♀</specimenCount>
;
<collectorName id="461B45806E4E4EFE2D503CC37DC7A9F2">Ongot</collectorName>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6FBK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[11.372783,3.858617]}">
<geoCoordinate id="DC9F4B564155C0DB95C0B06E232F6897" degrees="3" direction="north" minutes="51.517" orientation="latitude" precision="1" value="3.8586166">3 ° 51.517 ' N</geoCoordinate>
,
<geoCoordinate id="E1C9ECD5D8E87C7464C0A8487E6CF948" degrees="11" direction="east" minutes="22.367" orientation="longitude" precision="1" value="11.372784">11 ° 22.367 ' E</geoCoordinate>
</named-content>
;
<date id="9E1E2D3C42A7BB5E8F9D0F376C910B57" value="2022-03-20">
<collectingDate id="DD319470BEFC49DB0ED4CCD9EB05CEC9" value="2022-03-20">20 Mar. 2022</collectingDate>
</date>
; J. A.
<collectorName id="B916603A9979F217484127425AE27801">Yetchom Fondjo</collectorName>
&amp; A.
<collectionCode id="31F33A36520EDD90023BE42D3287B72F" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
.
<collectorName id="C938283C431703DAA7CE1C9D16B0518D">Nzoko Fiemapong</collectorName>
leg.;
<named-content id="84E4D322D59B766B1F870498A2AD6F78" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="F690DB14CC82AEB237A6F24091467AFF" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIGBK" type="redescription">
<paragraph id="80B79A34ED0B2161D22E68AD8944D6C2">
<heading id="F60233059C13ABF191369C955E426170" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="1DD924251FC0BA57CB4D71BD07E678A0">
Subocellar facial spot V-shaped, wide; dark median longitudinal band on the pronotum disc not very distinct; the two contiguous pale bands present; lower part of body very pale on living specimen; pronotum disc shiny brownish; pale basal posterior spots on the lateral lobes of the pronotum extending almost to the lower margin; lower half of elytra shiny black, upper half brown; meso- and metathoracic episternites with pale, narrow, median and basal band; front and middle legs pale green; outer and inner sides of hind femora pale, greenish yellow with apical third gradually darkening towards the pregenicular black ring; knees pale brown; hind tibiae green, sometimes very dark; male subgenital plate truncated; male cerci with a small inner preapical lobe.
<emphasis id="AF08F32B03BE020459A682D7161F76B0" bold="true" italics="true">Epiphallus</emphasis>
(15 E): of smaller size, ancorae small, anterior projections and lateral plates wide.
<emphasis id="7D0AA9245266CC6C4D0F3F032EF124E5" bold="true" italics="true">Phallic complex</emphasis>
(Fig.
<figureCitation id="C109FCE0CC06269FC1C09BA56F13ACBE" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Internal genitalia of P. carnapi (A D), P. descampsi (E H), P. karschi zenkeri (I L), P. uniformis (M P) and P. verrucigena (O T)." figureDoi="10.3897/zookeys.1216.130270.figure15" httpUri="https://binary.pensoft.net/fig/1161003">15 F H</figureCitation>
): dorsal arch of cingulum V-shaped, open, apodemes reaching apex of endophallic sclerites; rami bent; upper ectophallic sheath short; lower ectophallic sheath not capping base of rami; latero-ventral sclerite subtriangular.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGHBK" type="distribution">
<paragraph id="4B8937420262ABC6A625E99F34C56D4C">
<heading id="28F8F1BB87704D1F0F37B3014CAE7ED7" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="3C636B72F168BD368CC32A25843F2B4C">
<collectingCountry id="668E548F3A25D660249AB353BA3D3F93" name="Cameroon">Cameroon</collectingCountry>
(Fig.
<figureCitation id="C7A15C4B99D0C7A078AA486987CD5F95" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of Pteropera species in African rainforests." figureDoi="10.3897/zookeys.1216.130270.figure16" httpUri="https://binary.pensoft.net/fig/1161004">16 B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,456 @@
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<mods:title id="A0A9DBCF7672D1B74D4E4C16F625DF37">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<subSubSection id="9A6092B6B4704E7F443E114FF5C74DF6" type="nomenclature">
<paragraph id="C05378EA21433FAA02AAF3DBEFF4F978">
<taxonomicName id="CEE4A511662738B36FD64DC3573BA0CD" authority="Wang, S. Li &amp; Yao" authorityName="Wang, S. Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nayong" status="sp. nov.">
<emphasis id="7FAFAF0E1AFCD3E72A4A09CDF026143E" italics="true">Belisana nayong</emphasis>
Wang, S. Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="DAE10AB82845671B6A423049A678AE56" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3D4C11CF533CEC5F6DC45A280684B739" type="description">
<paragraph id="6A941F788EDA5A0424D454D90307F602">
<figureCitation id="5A45DD59E0809C47415C8FE3B425403F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">Figs 8</figureCitation>
,
<figureCitation id="D0F9F2F99E3155D0068AB5A9F10F3BCD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9</figureCitation>
,
<figureCitation id="1CC30D051141CC6E89FA9756E0006D30" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 G, H</figureCitation>
,
<figureCitation id="1E57EE22B4DB3685918200753EBE4BA8" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDGBG" type="type material">
<paragraph id="894C9E6E17F7F13C1626A181C0A254BB">
<heading id="367C56C855790D953B6FB8CE485E9759" reason="title">Type material.</heading>
</paragraph>
<paragraph id="A7747A6D6B423E1B572FD0A560F32893">
<materialsCitation id="5DAF16F49DE3FB84FD68AF8ABA42C200" collectingDate="2007-04-27" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Nayong County" elevation="1468" latitude="26.8361" location="Laosiba Town" longLatPrecision="1" longitude="105.52037" municipality="Bijie" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="8B0986336BC19A02501F2F6B39F53E24" bold="true">
<emphasis id="F6C9CA462849DDDA88FEE763ADE4BDBA" bold="true" italics="true">
<typeStatus id="224D026F2E950CB1F964538AB0AECE35" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="1CF10E1ADFADB6C5FBB7A82676880582" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="D066EB7ABEDF20D24A4F6E709F0D57C9" count="1" type="male"></specimenCount>
;
<collectingRegion id="C74615C50475E73B2EE1B000DFF0AEF8" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingMunicipality id="56BDE32D6DEEA3B26042B7F972D2B216">Bijie</collectingMunicipality>
,
<collectingCounty id="71CBF7E272F8281C50B05AF0AC099ED8">Nayong County</collectingCounty>
,
<location id="9D476C1BB8249452D43F81627C521BB8" LSID="urn:lsid:plazi:treatment:C614FD4B0E6A5C84A58BBE06859E2EEA:9D476C1BB8249452D43F81627C521BB8" country="China" county="Nayong County" latitude="26.8361" longLatPrecision="1" longitude="105.52037" municipality="Bijie" name="Laosiba Town" stateProvince="Guizhou">Laosiba Town</location>
,
<location id="53603A7BD7BBC14B6DE695EF0D3F80B2" LSID="urn:lsid:plazi:treatment:C614FD4B0E6A5C84A58BBE06859E2EEA:53603A7BD7BBC14B6DE695EF0D3F80B2" country="China" county="Nayong County" latitude="26.8361" longLatPrecision="1" longitude="105.52037" municipality="Bijie" name="Bailong Cave" stateProvince="Guizhou">Bailong Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERGBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[105.520367,26.836100]}">
<geoCoordinate id="B4BC5170D6A2676E819FD0CBD3796973" degrees="26" direction="north" minutes="50.166" orientation="latitude" precision="1" value="26.8361">26 ° 50.166 ' N</geoCoordinate>
,
<geoCoordinate id="E6FADB8188D923F9C3FD1FAD3C3FF04E" degrees="105" direction="east" minutes="31.222" orientation="longitude" precision="1" value="105.52037">105 ° 31.222 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="420A3A206979613ABAFCE339AEF0EFF6" metricMagnitude="3" metricUnit="m" metricValue="1.468" unit="m" value="1468.0">
<elevation id="65744BD9B5FCA43C8D4BF1E1D92839C7" metricMagnitude="3" metricUnit="m" metricValue="1.468" unit="m" value="1468.0">1468 m</elevation>
</quantity>
;
<date id="B2AF8764202A0832DCEABB56822B8EF4" value="2007-04-27">
<collectingDate id="4194D96C7F8C029B89DB50CFA075AFBA" value="2007-04-27">27 Apr. 2007</collectingDate>
</date>
;
<collectorName id="D6DF30CE38B0718BB2559D5312B144B2">J. Liu</collectorName>
&amp;
<collectorName id="C5DA3D4AED15D2514CBFE272F84207F2">Y. Lin</collectorName>
leg.;
<named-content id="D0C6F2037456272CA698C687DC7BA22C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="4FD306B1F95894D5E30CA2A66D7A1DFD" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<collectorName id="5034458D4FABCF283AC3AF3DE5647618">Ar</collectorName>
45198
</materialsCitation>
.
<materialsCitation id="C2EC26BE7BD6982210BF1F0EE5DF8A0B" collectingDate="2007-04-27" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Nayong County" elevation="1468" latitude="26.8361" location="Laosiba Town" longLatPrecision="1" longitude="105.52037" municipality="Bijie" specimenCount="3" specimenCount-male="3" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="D195FF6A511F7FE6C10564F48C09B19A" bold="true">
<emphasis id="8FFEBD52E381934540F84DBC236750A7" bold="true" italics="true">
<typeStatus id="D01841FD44E9FA33CE7D6CF2590B7F55" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="6753FEF59B6F3FE87E7BCF0E122CB827" name="China">China</collectingCountry>
</emphasis>
<specimenCount id="75EB056FE9DF8A06F3F5E6DEF5D8F0FD" count="3" type="male">3 ♂</specimenCount>
; same data as for holotype;
<named-content id="1E3C42DDFD19A35C73EBEFEEE1F47944" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="2CFED1BEDEF33B221A31C5659201BFDC" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45199201
</materialsCitation>
<materialsCitation id="963EE41D9C49BDE9293D4264529B42C8" collectingDate="2007-04-27" collectionCode="IZCAS" collectorName="J. Liu &amp; Y. Lin &amp; Ar" country="China" county="Nayong County" elevation="1468" latitude="26.8361" location="Laosiba Town" longLatPrecision="1" longitude="105.52037" municipality="Bijie" specimenCount="4" specimenCount-female="4" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="0DC125F74372380274C4CF47C7473259" count="4" type="female">4 ♀</specimenCount>
; same data as for holotype;
<named-content id="43AC13CBCD5ADBE6A1BA4F82F007384D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="FF1E056167A95B0A38F0D51D9E55CCC4" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4520205
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGHBG" type="diagnosis">
<paragraph id="C34FA13769A37B2AB1C23B4AA7ACE362">
<heading id="9AD8357803C3DDA3C54C28266D0C29EF" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="63526EEBA744FEEBA5742A09C6442892">
The new species resembles
<taxonomicName id="F2B30EC576C3B8C0272C66857CFA33BB" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongcong" status="sp. nov.">
<emphasis id="95D5CFBDF09BB53F464C708B5F76279A" italics="true">B. yongcong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61CB44649BD80A125955A83D566B2EA7" rank="species">sp. nov.</taxonomicNameLabel>
(Figs
<figureCitation id="970AC675F85D7518D843C783492B5713" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Belisana yongcong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at sclerotized dorso-subdistal apophysis D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure14" httpUri="https://binary.pensoft.net/fig/1161059">14</figureCitation>
,
<figureCitation id="B70CAED2D05922515F8C20CFED791CA2" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Belisana yongcong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure15" httpUri="https://binary.pensoft.net/fig/1161060">15</figureCitation>
,
<figureCitation id="F90A4953C784C3C3A2D61C89F7236597" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana yongcong sp. nov. C, D B. zhangi Tong &amp; Li, 2007 E, F B. zhouxi sp. nov. G, H B. zunyi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure23" httpUri="https://binary.pensoft.net/fig/1161068">23 A, B</figureCitation>
,
<figureCitation id="13AB49BE4BC9A4320AC2A93D69FB2BE2" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Female genitalia in ventral and dorsal views A, B Belisana xiuwen sp. nov. C, D B. yongcong sp. nov. E, F B. zhangi Tong &amp; Li, 2007 G, H B. zhouxi sp. nov. I, J B. zunyi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure25" httpUri="https://binary.pensoft.net/fig/1161070">25 C, D</figureCitation>
) by having similar bulbal apophysis (hooked; Fig.
<figureCitation id="D254DCEF466ECB0AEA61F3745D7EC9B5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 C</figureCitation>
) and vulva (anterior arch ridge-shaped, pore plates curved, long elliptic and 8 × longer than wide; Figs
<figureCitation id="0CE2B765AE9CB6BC6D4191AB67A31EC9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 B</figureCitation>
,
<figureCitation id="157449C7044E9A9836E1695E56C2A292" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 H</figureCitation>
), but can be distinguished by procursus with pointed ventro-subdistal membranous process (arrow
<quantity id="0EA9B98E26B0E2007DC348F458DF2379" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="B4522DA68BF69519699AEB567DF1786D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8 C</figureCitation>
,
<figureCitation id="154DDB58D86E61DB75002FAF52FB2B3E" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 G</figureCitation>
vs blunt), by male cheliceral distal apophyses pointing inwards (
<abbrev id="ABBRID0E2IBG" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="DFC0C378647AA0417AA7DE89831EE4CE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 D</figureCitation>
vs outwards), and by epigyne with postero-median pockets (
<abbrev id="ABBRID0EDJBG" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="60CFEB411722F1C075B9EFABC2F72F9D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 A</figureCitation>
,
<figureCitation id="5E176C5F6D76D1C119D1CBE6EC0B92CF" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 G</figureCitation>
vs lateral).
</paragraph>
<caption id="3A582D7E5DAB624B2419382618C8E0FF" ID-DOI="10.3897/zookeys.1216.132561.figure8" ID-arpha="6752D32F-FCBA-5C26-A6DE-DEA38A220FE6" httpUri="https://binary.pensoft.net/fig/1161053" startId="F8">
<paragraph id="56E32605261E16ED27332692AD3B70FA">
<label id="39E0103F6345F176CAEA544AFFE0A397">Figure 8.</label>
</paragraph>
<paragraph id="91E32FB38DA6C686A1A23B9B599FFC8C">
<taxonomicName id="8A570EE68E422215CD3061B57A4C9431" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nayong" status="sp. nov.">
<emphasis id="3AC034A66E43F7D11DBA3E785F71B021" italics="true">Belisana nayong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57463C18EBCEE2AE33076147636C8981" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="7DBAF663D66E3965C5A993A77F485E11" bold="true">A, B</emphasis>
palp (
<emphasis id="38B30463AD7271D778638B9BA5B25E60" bold="true">A</emphasis>
prolateral view
<emphasis id="08B7ABBE1635CADE60C00990907A9F0B" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="F4D05C2038F777950A644D3C7F6F0B87" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="F8F6F11EF4CD13BE3ABB61604C51C78C" bold="true">C</emphasis>
prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine
<emphasis id="79C8AE2BCC4BAA81B4FA801BC7907E5A" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb,
<abbrev id="ABBRID0EOKBG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0ESKBG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.20 (
<emphasis id="9FCB9CEC7241BF00A9A21A69CC1762EC" bold="true">A, B</emphasis>
); 0.05 (
<emphasis id="6B302EC4DACC5955BCEE76DCCB480357" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="F6D0ECE8FF058CCF19E1605F38DDED2F" ID-DOI="10.3897/zookeys.1216.132561.figure9" ID-arpha="E7E2804A-FFE5-53D5-BCA3-CBB4E1A6AF64" httpUri="https://binary.pensoft.net/fig/1161054" startId="F9">
<paragraph id="28E64412EAF7E34F694D7244330F956F">
<label id="73A32CD52703218E1A9371001FF8A00F">Figure 9.</label>
</paragraph>
<paragraph id="9EF676C9992E427DFC9DAC0337A224D5">
<taxonomicName id="9C6F3C8B340E1BE160692C885B650240" authorityName="Wang &amp; Li &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nayong" status="sp. nov.">
<emphasis id="0BC0AEC99F6C4FEB83ED05C4A719FADF" italics="true">Belisana nayong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EC291DEBC03E83F129D3BE5EF1D8A4D" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="3BBE334542D25E5E3482CACEACE1EAD7" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="022ED0A58426C4FE9C53A56A4764EEE0" bold="true">A, B, G, H</emphasis>
)
<emphasis id="2C9855D296E8F8FD3E174D8CB44D59BA" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="9481BD13CAABA8CB4A5FA534E9CDDBEE" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="580A4C7CD404CBE529B12749792CA6D5" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="611011D4A2401E906EBDE681D3088628" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="12A893B2422BF4893CD400F5D8227BA6" bold="true">E H</emphasis>
habitus (
<emphasis id="940E32527844B10C7960B05E66BB64C8" bold="true">E, G</emphasis>
dorsal view
<emphasis id="D5061C43ED993E1DFB1EB0737D0CC5FA" bold="true">F</emphasis>
lateral view
<emphasis id="42A2B7201DDED7EE8B40EFC74F37574D" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EGMBG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EKMBG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EOMBG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0ESMBG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EWMBG" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0E1MBG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.05 (
<emphasis id="3CD5A0B7D2FD0B5CECF01CE753BB5920" bold="true">A D</emphasis>
); 0.40 (
<emphasis id="6FAE507E739EC7B0A271740116B3FFAD" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EHNBG" type="description">
<paragraph id="731DA977E5925CA345BACDDA8DC5921F">
<heading id="40AE77C06E43BC6FA8C2132CB55F9381" reason="title">Description.</heading>
</paragraph>
<paragraph id="912F6DF156029C28B9DA3CADE2D8A6AD">
<emphasis id="7A357175A3B51B49D3782842D1C34F65" bold="true">Male</emphasis>
(
<emphasis id="BD0FF9167F7DA8DC6E87781BF8D67DCA" bold="true">
<typeStatus id="419EF2EF1F5CEA9A0BC4AA6BC896BE6A" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 2.33 (2.50 with clypeus), prosoma 0.86 long, 0.89 wide, opisthosoma 1.47 long, 1.15 wide. Leg I: 31.51 (8.33, 0.42, 7.82, 13.08, 1.86), leg II missing, leg III: 19.77 (5.08, 0.39, 5.04, 7.88, 1.38), leg IV missing; tibia I
<abbrev id="ABBRID0ERNBG" xlink_title="length/diameter">L / d</abbrev>
: 89. Eye interdistances and diameters:
<abbrev id="ABBRID0EVNBG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EZNBG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0E4NBG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0EBOBG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EFOBG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.68 / 0.64. Habitus as in Fig.
<figureCitation id="5309CB7637D9E5B3AB27642B11C5D94F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, with brown radiating marks; clypeus brown; sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0ENOBG" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="B8A3ABD1B68E43C749A2069EDDE1672C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 D</figureCitation>
) and pair of curved distal apophyses (distance between tips: 0.02;
<abbrev id="ABBRID0EVOBG" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="CD9B72F4E9307F43B7163C4AF88E3DFF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 D</figureCitation>
). Palp as in Fig.
<figureCitation id="EAAE22140BD1D69F02BED6E8A77E70AE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="FED2E0004DCD1E01E26DBC0286D222FD" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="3DC9DCCCDE49484A43F573E3B0E29BE3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8 B</figureCitation>
); femur with retrolatero-proximal protrusion (arrow
<quantity id="EAA0FF616D69065E4EC43D6DB3071DD6" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="5C03D6AB5DE6071046473788B00E85D1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8 B</figureCitation>
); procursus simple proximally but complex distally, with ventro-subdistal membranous process (arrow
<quantity id="6D9F964D4CD897D8C302E83FC708EC6F" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="BBA31A694109241D32DF57CF7C2F40F3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8 C</figureCitation>
,
<figureCitation id="E7ACADA44A44A3E0974EA77E1A2B60B2" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 G</figureCitation>
), prolatero-distal membranous lamella (arrow
<quantity id="0C66E6A2D52F20EA25EACF878E2F26E6" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="67FA3FBCC6DC760837711179C7FE4550" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8 C</figureCitation>
,
<figureCitation id="86181E5DECDC3FA54F9CCBC4DD8623E5" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 G</figureCitation>
) bearing proximally sclerotized part, dorso-distal spine (arrow
<quantity id="067B0EF1ABBAF189FD85422F206926CD" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="CD1557A1CCB4A42CBEE8BEDE2EACDB0F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8 C</figureCitation>
,
<figureCitation id="0C2A38F3AE4B1B9726E963EED6980E03" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 G</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="FE1E193D08B0760487F495C6CBE39F80" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Belisana nayong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous process, arrow 2 points at prolatero-distal membranous lamella, arrow 3 points at dorso-distal spine D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.20 (A, B); 0.05 (C, D)." figureDoi="10.3897/zookeys.1216.132561.figure8" httpUri="https://binary.pensoft.net/fig/1161053">8 D</figureCitation>
,
<figureCitation id="3596EDE2DE997B518880415C1EDADF31" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Procursus in prolateral and retrolateral views (The arrows point at the same structures as those shown in the photos of each species) A, B Belisana bijie sp. nov. C, D B. liupanshui sp. nov. E, F B. majiang sp. nov. G, H B. nayong sp. nov. I, J B. qixingguan sp. nov. K, L B. xiuwen sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure22" httpUri="https://binary.pensoft.net/fig/1161067">22 H</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EJQBG" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="336792D3289205B2ECAC288E7C10A027" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="76B5EF0BE34A547D97FB6939408398F4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 C</figureCitation>
). Retrolateral trichobothria on tibia I at 4 % proximally; legs with short vertical setae on metatarsi; tarsus I with 18 distinct pseudosegments.
</paragraph>
<paragraph id="361811E4AD1DA69ADB45CD38CB423A02">
<emphasis id="53D5DF33A9416B176ED99B64FAF7AD90" bold="true">Female</emphasis>
(
<emphasis id="A18C2C50E725014DE53507A1EF797C64" bold="true">
<typeStatus id="679E70C552869B2F79560E7488449E83" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="29FA33619318A1093B41084C12D69D6C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="584FB8F9E86666B558B65718F0931792" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45202): Similar to male, habitus as in Fig.
<figureCitation id="D6982C100278B73B9D48F8A01883AC26" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 G, H</figureCitation>
. Total length 2.47 (2.67 with clypeus), prosoma 0.81 long, 0.88 wide, opisthosoma 1.66 long, 1.40 wide; tibia I: 4.65; tibia I
<abbrev id="ABBRID0EDRBG" xlink_title="length/diameter">L / d</abbrev>
: 58. Eye interdistances and diameters:
<abbrev id="ABBRID0EHRBG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0ELRBG" xlink_title="posterior median eye">PME</abbrev>
0.11,
<abbrev id="ABBRID0EPRBG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0ETRBG" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EXRBG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.68 / 0.64. Epigyne simple and flat, posteriorly curved, with pair of postero-median pockets 0.02 apart (
<abbrev id="ABBRID0E2RBG" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="B702D213D965482ECA899D5BF55D072D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 A</figureCitation>
,
<figureCitation id="02C84428BCD2C866DEC6566E02CBA469" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 G</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0EHSBG" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="C17932AA5B3EEFD4DF61323C53A643A5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 B</figureCitation>
,
<figureCitation id="BFBED9A89E92AA92F661D921A3CB86E1" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 H</figureCitation>
) and pair of curved, long elliptic pore plates (8 × longer than wide,
<abbrev id="ABBRID0ETSBG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="6442B7FC7CC4FBBF8CBDB908EEBC6C3C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Belisana nayong sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.40 (E H)." figureDoi="10.3897/zookeys.1216.132561.figure9" httpUri="https://binary.pensoft.net/fig/1161054">9 B</figureCitation>
,
<figureCitation id="DFB455A825C0C0084D35E8E56F2D9317" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Female genitalia in ventral and dorsal views A, B Belisana bijie sp. nov., arrow points at lateral sclerite C, D B. liupanshui sp. nov., arrow points at lateral sclerite E, F B. majiang sp. nov. G, H B. nayong sp. nov I, J B. qixingguan sp. nov., arrow points at sclerotized protrusion. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1216.132561.figure24" httpUri="https://binary.pensoft.net/fig/1161069">24 H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6SBG" type="variation">
<paragraph id="82D77F9F2E68F77E6A2F8019A8D7C03B">
<heading id="73207A23994EE23C7EE2170EFBAE0FA4" reason="title">Variation.</heading>
</paragraph>
<paragraph id="2630EFD0F3206B4BD5BCFFBDA4AA9F00">
Tibia I in
<specimenCount id="B7A732BA84233750451D373CC6A8D3EF" count="3" type="male">three male</specimenCount>
<typeStatus id="45E140185FF5AA6D0A54A6BFBC27D3A7" type="paratype">paratypes</typeStatus>
(
<named-content id="89B548D491C1653270AAFAC21E455651" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="7C7C2157C4D20FCB7E5633DEC877F431" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 45199201): 6.15, 6.35, 6.50. Tibia I in the other
<specimenCount id="E083B7607E85E2DFF033F4583118AA26" count="3" type="female">three female</specimenCount>
<typeStatus id="2F8CDEC7E3CDEC360D796945F0F34676" type="paratype">paratypes</typeStatus>
(
<named-content id="CE0D95DFDF4D5CCB71D674FE68979A79" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="66BF9B37F4C73A4606F1EF12E48EB0BE" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
- Ar 4520305): 4.45, 4.50, 4.74.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENTBG" type="habitat">
<paragraph id="CBE29EE49ABFEDBA4A9592CB92F3B842">
<heading id="93DB79FD63B752775945AB8F1AA1D49E" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="862891C33A6D2D81E5F026090B1CCBE1">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0ESTBG" type="distribution">
<paragraph id="4F781A0CF500CC56C84ADAD1EA3F9B86">
<heading id="CE2146D2141C07399423730051372904" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="BA407FBA6AE455A27C7CA86E0F6B5220">
<collectingCountry id="E496F1852747365536A8530316BB1E2E" name="China">China</collectingCountry>
(
<collectingRegion id="78459762188B39BD08BE3144C8A7EF30" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<typeStatus id="1F39BA5FD383607068849C97B90CCC53">type</typeStatus>
locality; Fig.
<figureCitation id="96292D064D9F2E3EA850FA94C1508BB3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guizhou, China 1 Belisana bijie sp. nov. 2 B. daji Chen, Zhang &amp; Zhu, 2009 3 B. douqing Chen, Zhang &amp; Zhu, 2009 4 B. galeiformis Zhang &amp; Peng, 2011 5 B. lii Chen, Yu &amp; Guo, 2016 6 B. liupanshui sp. nov. 7 B. majiang sp. nov. 8 B. nayong sp. nov. 9 B. qixingguan sp. nov. 10 B. wangchengi Wang, Yao &amp; Zhang, 2024 11 B. xishui Chen, Zhang &amp; Zhu, 2009 12 B. xiuwen sp. nov. 13 B. yanhe Chen, Zhang &amp; Zhu, 2009 14 B. yongcong sp. nov. 15 B. yuhaoi Yang &amp; Yao, 2023 16 B. zhangi Tong &amp; Li, 2007 17 B. zhouxi sp. nov. 18 B. zunyi sp. nov." figureDoi="10.3897/zookeys.1216.132561.figure1" httpUri="https://binary.pensoft.net/fig/1161046">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3TBG" type="etymology">
<paragraph id="80BAC96A0382E35335CEB4D9E32E23FB">
<heading id="3B6FA41DFE7D37CDA28B270F1C198BAD" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="B21187897D517267C0EB260BCF5469D2">
The specific name refers to the
<typeStatus id="68C963B0A79B30E2770BA50F41864D7A">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="D097520770FC468F1BD5340DA440C46A" ID-DOI="10.3897/zookeys.1216.130270" ID-ZooBank="1221A319-03DC-4157-A7F9-F5A18E20E1FF" ID-publisher-id="130270" URI-arpha="DE3D6390-FB9F-566D-B398-5D22FE3B56DF" URI-zoobank="http://zoobank.org/1221A319-03DC-4157-A7F9-F5A18E20E1FF" article-type="research-article" checkinTime="1729868654848" checkinUser="pensoft" docAuthor="Yetchom Fondjo, Jeanne Agrippine, Nzoko Fiemapong, Armand Richard, Tindo, Maurice, Duressa, Tarekegn Fite, Ivković, Slobodan &amp; Husemann, Martin" docDate="2024" docId="D1FDE9B2026A5D0591A1A486BFB75682" docLanguage="en" docName="ZooKeys 1216: 219-264" docOrigin="ZooKeys 1216" docSource="https://zookeys.pensoft.net/article/130270/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Pteropera augustini Donskoff 1981" docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="264" masterDocId="DE3D6390FB9F566DB3985D22FE3B56DF" masterDocTitle="Taxonomic review of the grasshopper genus Pteropera Karsch, 1891 (Orthoptera, Acrididea, Catantopinae) with description of three new species and a preliminary phylogeny of the Cameroonian species" masterLastPageNumber="264" masterPageNumber="219" pageNumber="219" updateTime="1729868729188" updateUser="pensoft">
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<mods:title id="202ADBC9ECB37AF5F3FE285190B5A193">Taxonomic review of the grasshopper genus Pteropera Karsch, 1891 (Orthoptera, Acrididea, Catantopinae) with description of three new species and a preliminary phylogeny of the Cameroonian species</mods:title>
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<mods:namePart id="41E17700A322D442637989887296F458">Yetchom Fondjo, Jeanne Agrippine</mods:namePart>
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<mods:affiliation id="3F0159E8C22DFD76F18EB54F06D18B65">Zoology Unit, Laboratory of Biology and Physiology of Animal Organisms, Graduate School in Fundamental and Applied Sciences, University of Douala, Douala, Cameroon &amp; Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<mods:namePart id="9CD50D3A7E120FF2502AF7FE53E48246">Nzoko Fiemapong, Armand Richard</mods:namePart>
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<mods:affiliation id="A9C2FAA9A309B6BD09ED5675B6A6CEAB">University of Neuchâtel, Neuchâtel, Switzerland</mods:affiliation>
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<mods:namePart id="06139E8BE63082B60D6289810F8AD1D7">Tindo, Maurice</mods:namePart>
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<mods:affiliation id="6230B5A56AA08AA788B2F62184E65299">Zoology Unit, Laboratory of Biology and Physiology of Animal Organisms, Graduate School in Fundamental and Applied Sciences, University of Douala, Douala, Cameroon</mods:affiliation>
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<mods:namePart id="9E29B75110DE5A35817B61D071DBA692">Duressa, Tarekegn Fite</mods:namePart>
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<mods:affiliation id="9E44DA2C3170D4B8B0D74A37F9C9A237">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany &amp; School of Plant Sciences, College of Agriculture and Environmental Sciences, Haramaya University, Dire Dawa, Ethiopia</mods:affiliation>
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<mods:namePart id="A11EDB99FA0A89C3DD21A38B40D2FD10">Ivković, Slobodan</mods:namePart>
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<mods:affiliation id="CC220E2E7DF21C0F250FE98352DF5382">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<mods:affiliation id="4862B2ECE1E42CB053B0DF17D48E4625">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<paragraph id="623689A4EF1CB105457D3364379DA702">
<taxonomicName id="B39509E4B6CD6A324B1229958F74F906" ID-CoL="4Q25P" authority="Donskoff, 1981" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="augustini">
<emphasis id="A8BD45636FB081DE8A175C607A298E1F" italics="true">Pteropera augustini</emphasis>
<bibRefCitation id="AFCA4F5C42695A94AA704118704393C1" DOI="10.1080/21686351.1981.12278263" author="Donskoff M" issue="1" journalOrPublisher=")" pagination="33-88" refId="B4" refString="Donskoff M (1981) Les acridiens de la foret africaine. I. Revision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Societe entomologique de France (N. S.) 17 (1): 33 - 88. [51 figs] https: // doi. org / 10.1080 / 21686351.1981. 12278263" title="Les acridiens de la forêt africaine. I. Révision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Société entomologique de France (N. S." volume="17" year="1981">Donskoff, 1981</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3420C4C9A0A200F0F320DEC0F5F5E71" type="description">
<paragraph id="C55FA3C507EF33A40A390B9C47B6BFD7">
<figureCitation id="D86F17EB9413F2232EEB578682A2CB2B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Images of holotypes, allotypes, and paratypes of Pteropera species in lateral view A P. augustini (holotype ♂) B P. augustini (allotype ♀) C P. balachowskyi (holotype ♂) D P. balachowskyi (allotype ♀) E P. bertii (holotype ♂) F P. bertii (paratype ♀) G P. brosseti (holotype ♂) H P. brosseti (allotype ♀) I P. basilewskyi (paratype ♀) J P. bredoi (holotype ♀)." figureDoi="10.3897/zookeys.1216.130270.figure2" httpUri="https://binary.pensoft.net/fig/1160990">Figs 2 A, B</figureCitation>
,
<figureCitation id="1AAAFE7A952FE8F4095A4ED6A353FB5C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Images of holotypes, allotypes and paratype of Pteropera species in dorsal view A P. augustini (holotype ♂) B P. augustini (allotype ♀) C P. balachowskyi (holotype ♂) D P. balachowskyi (allotype ♀) E P. bertii (holotype ♂) F P. bertii (paratype ♀) G P. brosseti (holotype ♂) H P. brosseti (allotype ♀) I P. basilewskyi (paratype ♀) J P. bredoi (holotype ♀)." figureDoi="10.3897/zookeys.1216.130270.figure7" httpUri="https://binary.pensoft.net/fig/1160995">7 A, B</figureCitation>
</paragraph>
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<paragraph id="54C15504AF669B4F49AD3DED198288AC">
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<treatmentCitation id="7B8484E95CF6B52F7BFABF6DC144EE0D" author="Donskoff M" page="52" year="1981">
<taxonomicName id="D12AC835C61B6EDF94EB585ACF34A7A3" ID-CoL="4Q25P" authority="Donskoff, 1981: 51 - 52" authorityName="Donskoff" authorityPageNumber="51 - 52" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="augustini">
<emphasis id="DBB34B4B81089E471CADCC16E53DAC34" italics="true">Pteropera augustini</emphasis>
<comment id="832B03B0D96A4E8CF0A46BC2248C8991">
<bibRefCitation id="76D4F58C58D5FBB4623923FF3711D5FF" DOI="10.1080/21686351.1981.12278263" author="Donskoff M" issue="1" journalOrPublisher=")" pagination="33-88" refId="B4" refString="Donskoff M (1981) Les acridiens de la foret africaine. I. Revision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Societe entomologique de France (N. S.) 17 (1): 33 - 88. [51 figs] https: // doi. org / 10.1080 / 21686351.1981. 12278263" title="Les acridiens de la forêt africaine. I. Révision du genre Pteropera Karsch, 1891 (Orthoptera, Acrididae). Annales de la Société entomologique de France (N. S." volume="17" year="1981">Donskoff, 1981: 5152</bibRefCitation>
.
</comment>
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</paragraph>
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<subSubSection id="SECID0EZQAK" type="type material">
<paragraph id="74D0DC9E664CD1C3CB7233FC360592B9">
<heading id="E945FAAD32A048DA55AC5E3738F6979E" reason="title">Type materials examined.</heading>
</paragraph>
<paragraph id="E88549D742672975A60A1AAD1FBACDEA">
<materialsCitation id="95FA7D420EF0A61B157E2782AB9BEDFF" collectingDate="1974-06-11" collectionCode="MNHN" collectorName="M. Donskoff &amp; J. Le Breton" country="Gabon" latitude="0.41028336" location="Near Youmi" longLatPrecision="1" longitude="9.4366665" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="99F9D0FE0C722ADF68523A6A2C03D95D" bold="true" italics="true">
<typeStatus id="3AC405B2232A07A87CFE3FF797E9CFBC" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="53BB540CF85ADC34351EED74F861C697" count="1" type="male"></specimenCount>
;
<collectingCountry id="C4D83D23D915062E0ABB041329EE9D36" name="Gabon">Gabon</collectingCountry>
.
<location id="CDFBC611F4114C0034C77A5E7D22933B" LSID="urn:lsid:plazi:treatment:D1FDE9B2026A5D0591A1A486BFB75682:CDFBC611F4114C0034C77A5E7D22933B" country="Gabon" latitude="0.41028336" longLatPrecision="1" longitude="9.4366665" name="Near Youmi">Near Youmi</location>
, in forest habitat;
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<geoCoordinate id="D270B19859CE9BB1C3FA7B3DCC79FBD1" degrees="0" direction="north" minutes="24.617" orientation="latitude" precision="1" value="0.41028336">0 ° 24.617 ' N</geoCoordinate>
,
<geoCoordinate id="AD77D47C1A008F94104DC32D5379F474" degrees="9" direction="east" minutes="26.200" orientation="longitude" precision="1" value="9.4366665">9 ° 26.200 ' E</geoCoordinate>
</named-content>
;
<date id="814ED5B2729CAE0680D2A242E03A9A2B" value="1974-06-11">
<collectingDate id="03F6776AEA7E330EC9ACB35F3C6B83BF" value="1974-06-11">11 Jun. 1974</collectingDate>
</date>
;
<collectorName id="53C136ED114D2FF0C235324C591A87BE">M. Donskoff</collectorName>
&amp;
<collectorName id="FFF52866EFF8D7C6AA02467E742DF1DC">J. Le Breton</collectorName>
leg.;
<named-content id="06C682C48E466BB8E646262840D27ADE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum National d´Histoire Naturelle Paris, France">
<collectionCode id="D3204205061348CD131C032ABBA449E8" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
,
<named-content id="53A045A046F6CE0DD472C4F1D7D952D2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-national-dhistoire-naturelle-2" xlink_title="Muséum National d´Histoire Naturelle Paris, France">
<collectionCode id="D102F3B26A946D1AEAAAE14E9B72C37A" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" type="">MNHN</collectionCode>
</named-content>
- EO-CAELIF 11462.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EURAK" type="material">
<paragraph id="B33000B5ED51106F8C36D0741B22BAF0">
<heading id="8E4D59576588B4CD5C669A6DC880068A" reason="title">Other material examined.</heading>
</paragraph>
<paragraph id="9850D9D7DA3A25ED57A8AE2555EC3719">
<materialsCitation id="407A14A1A41A09D8FE64C233CB8E4364" collectingDate="2022-04-11" collectionCode="SMNK" collectorName="J. A. Yetchom Fondjo" country="Cameroon" elevation="606" latitude="3.3941667" location="Somalomo" longLatPrecision="1" longitude="12.89305" municipality="Dja Biosphere Reserve" specimenCount="1" specimenCount-female="1">
<collectingCountry id="E4650CF4FE4FC0003C52B9A57B75C06A" name="Cameroon">Cameroon</collectingCountry>
<specimenCount id="2DA9C0AA085019E399D79675B0FF8FE5" count="1" type="female">1 ♀</specimenCount>
, subadult;
<location id="7FA37CDD86DA31F1BBE296D1030A312E" LSID="urn:lsid:plazi:treatment:D1FDE9B2026A5D0591A1A486BFB75682:7FA37CDD86DA31F1BBE296D1030A312E" country="Cameroon" latitude="3.3941667" longLatPrecision="1" longitude="12.89305" municipality="Dja Biosphere Reserve" name="Somalomo">Somalomo</location>
, in the
<collectingMunicipality id="E4501797F2707B5DC994779651ADA2F0">Dja Biosphere Reserve</collectingMunicipality>
, cocoa farm;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4RAK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[12.893050,3.394167]}">
<geoCoordinate id="2F8DDF5332A5557FED75BD87EAB86F94" degrees="3" direction="north" minutes="23.650" orientation="latitude" precision="1" value="3.3941667">3 ° 23.650 ' N</geoCoordinate>
,
<geoCoordinate id="F95F1009F5BA9AA592A40ADFC69E61CF" degrees="12" direction="east" minutes="53.583" orientation="longitude" precision="1" value="12.89305">12 ° 53.583 ' E</geoCoordinate>
</named-content>
,
<elevation id="5E1ED80F507FAB21E985ADDE1BD83536" metricMagnitude="2" metricUnit="m" metricValue="6.06" unit="m" value="606.0">
<quantity id="F01CC4FC8CD6D1841B552826AE3FE37E" metricMagnitude="2" metricUnit="m" metricValue="6.06" unit="m" value="606.0">606 m</quantity>
a. s. l.
</elevation>
;
<date id="F78D5DB959FF8D367DC8C9D5F6A1F5CE" value="2022-04-11">
<collectingDate id="CB7E8A03886763D83A80BC41AC2AF45A" value="2022-04-11">11 Apr. 2022</collectingDate>
</date>
;
<collectorName id="DC4B2C4EEFD917868FA3EAD22667F32C">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="F4108CD737296694355B925AE18F224F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="D5A0008F8B76A34BFDAC25482EA3F154" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
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<subSubSection id="SECID0EGSAK" type="description">
<paragraph id="1D875ECBC9ED1667E955F57834153D85">
<heading id="6E04D48FC817F8C58ABFAF44BD87B235" reason="title">Morphological characteristics.</heading>
</paragraph>
<paragraph id="7CA43EFD60E3BE2CCEF3E1A564B0EECA">Two subocellar facial spots; posterior margin of the pronotum slightly indented; no marked difference between the upper half and lower half of the elytra; shiny black line along the lower margin; metathoracic episternites almost entirely pale; outer area of the hind femora green with pale spots, widely spread on the upper carina, median spot rounded, small, basal spot triangular; lower basal half of the inner area greenish-brown, pale spot rounded and small; only the lower inner area brownish; hind tibiae green.</paragraph>
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<paragraph id="8FA618699FA99C41AD983A66F6B49B14">
<label id="A51202535785A70E8AA405CCAB514568">Figure 2.</label>
</paragraph>
<paragraph id="6A8352ED010996F029DCA36959256631">
Images of holotypes, allotypes, and paratypes of
<taxonomicName id="9AEB776F152A184B73B70EE540D389F5">
<emphasis id="60E5F8AE778BE4ADC14038A903D20A1F" italics="true">Pteropera</emphasis>
species
</taxonomicName>
in lateral view
<emphasis id="C3A069B012C4B5FF3D12961745959FB1" bold="true">A</emphasis>
<taxonomicName id="0AD31600A749A609C4D875BFD77D13BC" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="augustini">
<emphasis id="77F06D3C8FEE25696852EF657E44D1F0" italics="true">P. augustini</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="5E09AC8E32AD42C493BDC9F4D80D507E" bold="true">B</emphasis>
<taxonomicName id="ADF0588F6C0307713F37EF177E53FDB6" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="augustini">
<emphasis id="008C8D5175F52F2F7CB5107001E10349" italics="true">P. augustini</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="E83A1E5BDEA5152D1BCAFA41BEAE1D4B" bold="true">C</emphasis>
<taxonomicName id="6D91577C06194455F6FAA87F72224854" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="balachowskyi">
<emphasis id="4CBEC443930DF57D20AEAF0D24DA7FAA" italics="true">P. balachowskyi</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="1054BC382D8110E79DAB71E3D6000452" bold="true">D</emphasis>
<taxonomicName id="34F066AF282820C909A88DFF5EFE5867" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="balachowskyi">
<emphasis id="767F9615BD3CC15B0DC616E4D7537F25" italics="true">P. balachowskyi</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="3D14C4FC3BE044962357EFF47ECD657B" bold="true">E</emphasis>
<taxonomicName id="720C3377945E48E70A7E10BF2330274D" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="bertii">
<emphasis id="9B523A83A28C93F0819DC8E3D001849D" italics="true">P. bertii</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="658655AE7FEA892FFBE73F480C8E7EF9" bold="true">F</emphasis>
<taxonomicName id="A028F101927262DD2E0D0CFB50182CF2" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="bertii">
<emphasis id="3962336148D66E9000C0268DD5A7032C" italics="true">P. bertii</emphasis>
</taxonomicName>
(paratype ♀)
<emphasis id="BE0F2B1982D6769761E47DF42E029F59" bold="true">G</emphasis>
<taxonomicName id="CAF454D5E88655F0E6230D1188F14823" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="brosseti">
<emphasis id="BF79EB60A64AFF0E8BCBF83434645015" italics="true">P. brosseti</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="A2B245C1509FD7C331FA66C2ADEBB91C" bold="true">H</emphasis>
<taxonomicName id="41F5844721E29C7EA0B8276DD4A12183" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="brosseti">
<emphasis id="B796344120E106CB41FDBAD1AE1DF3AE" italics="true">P. brosseti</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="3EA5C86E1F2F8789E699C8178BA2193A" bold="true">I</emphasis>
<taxonomicName id="FC1B3EA59CC0DA0611FD04158F6D8874" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="F000A20ADB30C6351601AED13F57CF6B" italics="true">P. basilewskyi</emphasis>
</taxonomicName>
(paratype ♀)
<emphasis id="8D3F0825770C92DB9E3519F79FD2EFAE" bold="true">J</emphasis>
<taxonomicName id="14EED123B2C15558498345DDEE64E92F" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="bredoi">
<emphasis id="7E10109E55666C1D91476B5AE2DE0F7D" italics="true">P. bredoi</emphasis>
</taxonomicName>
(holotype ♀).
</paragraph>
</caption>
<paragraph id="E740CB26D13B175AD82D2158644EF890">Female subgenital plate pentagonal; egg-guide short; anterior apodemes narrow, short; medio-dorsal pocket narrow; basivalvar sclerites broad, slightly fused, almost perpendicular to each other; end of the copulatory bursa enlarged in the shape of a bubble; spermatheca ampulla arched, both diverticula of different diameters; recurrent distal trunk of the lateral diverticulum 45 × longer than the proximal trunk.</paragraph>
</subSubSection>
<subSubSection id="SECID0ECXAK" type="remarks">
<paragraph id="586F284AD29CFA6A4EF95971D5846BD1">
<heading id="29C299A5BF95DD6E8E8D8E4EC7127820" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="5E1904320DE1AB1E091C3BFEC361B095">
This species is known by both males and females. The last female subadult stage was collected during this study and represents the first signalization of this species in
<collectingCountry id="5199FE7348BDF0A3666D86E8ADD3C97A" name="Cameroon">Cameroon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHXAK" type="distribution">
<paragraph id="CE8E64313FBD584ADD9E046A7045AA4C">
<heading id="A49B3A49A02C563E25E31493AEAD5896" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="EB286C48626646B9C8D62C8242B03F32">
<collectingCountry id="BAAEF9DBE2F66A967BF4531867227AB2" name="Gabon">Gabon</collectingCountry>
;
<collectingCountry id="6B7D152013B6E7E03413249702137ED8" name="Cameroon">Cameroon</collectingCountry>
(Fig.
<figureCitation id="6998514694D571364CD1DE0EEB1D542A" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution of Pteropera species in African rainforests." figureDoi="10.3897/zookeys.1216.130270.figure16" httpUri="https://binary.pensoft.net/fig/1161004">16 A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,291 @@
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<mods:title id="AEB691280508E6BA421C0B354BA6EED5">Taxonomic review of the grasshopper genus Pteropera Karsch, 1891 (Orthoptera, Acrididea, Catantopinae) with description of three new species and a preliminary phylogeny of the Cameroonian species</mods:title>
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<mods:namePart id="52D26DE391BDE224B2C8E03975DC73A0">Yetchom Fondjo, Jeanne Agrippine</mods:namePart>
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<mods:affiliation id="C7749EE52B3CCEC6211383EF5F2BF57A">Zoology Unit, Laboratory of Biology and Physiology of Animal Organisms, Graduate School in Fundamental and Applied Sciences, University of Douala, Douala, Cameroon &amp; Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<mods:namePart id="026FAA10B2F676C3F5D3EF302FDD4E29">Nzoko Fiemapong, Armand Richard</mods:namePart>
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<mods:affiliation id="A54CB5746D0930C22F1E1DF6A34A0FB1">University of Neuchâtel, Neuchâtel, Switzerland</mods:affiliation>
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<mods:name id="8C57F2B65E3F512397104F833BC796E0" type="personal">
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<mods:namePart id="FB2F13C94D7890694BA2E035850BB6E4">Tindo, Maurice</mods:namePart>
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<mods:affiliation id="0552AFD8D1EB004642359F797487EB5E">Zoology Unit, Laboratory of Biology and Physiology of Animal Organisms, Graduate School in Fundamental and Applied Sciences, University of Douala, Douala, Cameroon</mods:affiliation>
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<mods:namePart id="F9D94F27FDC73362342465A61DC5C239">Duressa, Tarekegn Fite</mods:namePart>
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<mods:affiliation id="D086260540539D488F8F0A1814C4E248">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany &amp; School of Plant Sciences, College of Agriculture and Environmental Sciences, Haramaya University, Dire Dawa, Ethiopia</mods:affiliation>
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<mods:namePart id="F24BB0AFBC8A3437D8292FEBC26FDD4B">Ivković, Slobodan</mods:namePart>
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<mods:namePart id="45CE888A3A4D6D412FA982431CCEEECE">Husemann, Martin</mods:namePart>
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<mods:affiliation id="405B93FCD4D1062266D970ADAAFC322E">Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany</mods:affiliation>
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<treatment id="D9F61B9CB2E3504F966C1A8054E44122" ID-arpha="D9F61B9C-B2E3-504F-966C-1A8054E44122" LSID="urn:lsid:plazi:treatment:D9F61B9CB2E3504F966C1A8054E44122" httpUri="http://treatment.plazi.org/id/D9F61B9CB2E3504F966C1A8054E44122">
<subSubSection id="C2800676F4FD1849C896C57F25CF8947" type="nomenclature">
<paragraph id="559E6B67B936AB01762367A76A550E60">
<taxonomicName id="2A5CC664F30C1490D83025D4B52A5109" ID-CoL="4Q26J" authority="Bruner, 1920" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="28E057A84A9F5173442000D1BEA81D56" italics="true">Pteropera uniformis</emphasis>
Bruner, 1920
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A649667ACFCCAE767A22D4FA7DD10709" type="description">
<paragraph id="D21A501B581277610DFAA983422F22D4">
<figureCitation id="2860632E4CAAA2B5649A519337B8BB98" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Images of holotypes, allotypes, letotype and neallotype of Pteropera species in lateral view A P. spleniata (holotype ♂) B P. spleniata (holotype ♀) C P. teocchii (holotype ♂) D P. teocchii (allotype ♀) E P. thibaudi (holotype ♂) F P. thibaudi (allotype ♀) G P. verrucigena (lectotype ♂) H P. verrucigena (♀) I P. villiersi (holotype ♂) J P. villiersi (allotype ♀) K P. uniformis (neallotype ♂)." figureDoi="10.3897/zookeys.1216.130270.figure6" httpUri="https://binary.pensoft.net/fig/1160994">Figs 6 K</figureCitation>
,
<figureCitation id="F1B94A683E6EF34C398556B442FF2733" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Images of holotypes, allotypes, neallotype and paratype of Pteropera species in dorsal view A P. teocchii (holotype ♂) B P. teocchii (allotype ♀) C P. thibaudi (holotype ♂) D P. thibaudi (allotype ♀) E P. verrucigena (holotype ♂) F P. verrucigena (allotype ♀) G P. villiersi (holotype ♂) H P. villiersi (allotype ♀) I P. uniformis (neallotype ♂) J P. femorata (♀)." figureDoi="10.3897/zookeys.1216.130270.figure11" httpUri="https://binary.pensoft.net/fig/1160999">11 I</figureCitation>
,
<figureCitation id="448F94B407251FB4E453199510AEAD55" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Internal genitalia of P. carnapi (A D), P. descampsi (E H), P. karschi zenkeri (I L), P. uniformis (M P) and P. verrucigena (O T)." figureDoi="10.3897/zookeys.1216.130270.figure15" httpUri="https://binary.pensoft.net/fig/1161003">15 M P</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E12BK" type="type material">
<paragraph id="23EF7D2A04E71825B88E1B99FED1F0FF">
<heading id="69E79DAE06E9B36DDABBFA24CCF56E45" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="DF3631B141277DC5E886375E2CDC4771">
<materialsCitation id="AD6A05629B904BC2C451A3767F3FDAA5" collectingDate="1914-04" collectionCode="ANSP" collectorName="F. H. Hope" country="Cameroon" latitude="2.8465834" location="Batanga" longLatPrecision="1" longitude="9.894983" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="051539089A4AE59E06B1BEAA0FFC62E3" bold="true">
<emphasis id="150BFBD907548DD6161B2822D495BD02" bold="true" italics="true">
<typeStatus id="AC20BF0B43FE388DC49527A18C156876" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="6E5D75DECB4FCA1631B5F40B6A11FB6B" name="Cameroon">Cameroon</collectingCountry>
<specimenCount id="B75FED9B92F628550C7A420A72D1749D" count="1" type="male"></specimenCount>
;
<location id="957D79A4DD8DEE992187DFC5718EAD67" LSID="urn:lsid:plazi:treatment:D9F61B9CB2E3504F966C1A8054E44122:957D79A4DD8DEE992187DFC5718EAD67" country="Cameroon" latitude="2.8465834" longLatPrecision="1" longitude="9.894983" name="Batanga">Batanga</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EI3BK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.894983,2.846583]}">
<geoCoordinate id="8917676F61AFCD88087A96C00D1FBA1D" degrees="2" direction="north" minutes="50.795" orientation="latitude" precision="1" value="2.8465834">2 ° 50.795 ' N</geoCoordinate>
,
<geoCoordinate id="A224B9C48D44A51CCF1F6389C0EF8CB3" degrees="9" direction="east" minutes="53.699" orientation="longitude" precision="1" value="9.894983">9 ° 53.699 ' E</geoCoordinate>
</named-content>
;
<date id="3482C07AC7B33592AC62226B2DAF1D7C" value="1914-04">
<collectingDate id="DB38F9E9D48D43A9D7D210F99CC9FFB1" value="1914-04">Apr. 1914</collectingDate>
</date>
;
<collectorName id="9B05A1D596CC6F8E8A9FA948F6574D17">F. H. Hope</collectorName>
leg.;
<named-content id="E3F64F8AD246A325FEEA4A5136F61EA2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/academy-natural-sciences-philadelphia" xlink_title="Academy of Natural Science of Philadelphia">
<collectionCode id="BE4162FA1516C4D53ACC43E2FDDD7716" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" type="">ANSP</collectionCode>
</named-content>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ES3BK" type="material">
<paragraph id="C994C01D55EE225FF1180D2AC679DF12">
<heading id="D35950577A0985619F6A050394226AB1" reason="title">Other material examined.</heading>
</paragraph>
<paragraph id="F1F5115B163B3E2D8777A39DC59BCAA0">
<collectingCountry id="1B75C56665FABDC549319E319AB07816" name="Cameroon">Cameroon</collectingCountry>
<materialsCitation id="97988319D7909A5042F3F59874DEE92B" collectingDate="2021-12-05" collectionCode="SMNK" collectorName="J. A. Yetchom Fondjo" country="Cameroon" latitude="3.8586166" location="Ongot" longLatPrecision="1" longitude="11.372784" specimenCount="1" specimenCount-male="1">
<specimenCount id="2F42AE26D6CE771E6847D6C595FB125F" count="1" type="male">1 ♂</specimenCount>
;
<location id="D6FBE727776E10F7839D86982596C000" LSID="urn:lsid:plazi:treatment:D9F61B9CB2E3504F966C1A8054E44122:D6FBE727776E10F7839D86982596C000" country="Cameroon" latitude="3.8586166" longLatPrecision="1" longitude="11.372784" name="Ongot">Ongot</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E23BK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[11.372783,3.858617]}">
<geoCoordinate id="F0F80B0FDFF7E430C972348E4CD5252C" degrees="3" direction="north" minutes="51.517" orientation="latitude" precision="1" value="3.8586166">3 ° 51.517 ' N</geoCoordinate>
,
<geoCoordinate id="50F7CD1709CC9CA5A9F16F1C786C4435" degrees="11" direction="east" minutes="22.367" orientation="longitude" precision="1" value="11.372784">11 ° 22.367 ' E</geoCoordinate>
</named-content>
;
<date id="3FA535530A40C7A22D739611FDEC048A" value="2021-12-05">
<collectingDate id="0FBD4C2104CFB41E188C06235416E683" value="2021-12-05">5 Dec. 2021</collectingDate>
</date>
;
<collectorName id="085D8FF2648DD40D7C621155F13B7DE6">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="7ACC567EB8EC23DE67A39F42A87A8759" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="C6B9390464233F171414F23C62D6FBF5" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
.
<collectingCountry id="76A82E00B874776D621A3DC593FB21CF" name="Cameroon">Cameroon</collectingCountry>
</materialsCitation>
<materialsCitation id="4D60E7529919ACD8A7C1A5625832CD32" collectingDate="2022-04-11" collectionCode="SMNK" collectorName="J. A. Yetchom Fondjo" country="Cameroon" latitude="3.3741333" location="Somalomo" longLatPrecision="1" longitude="12.733167" municipality="Dja Biosphere reserve" specimenCount="1" specimenCount-male="1">
<specimenCount id="1F7535C75A2167357FF3E601ADF42563" count="1" type="male">1 ♂</specimenCount>
;
<location id="53F601E45D8AB35B0869D281EA3F9584" LSID="urn:lsid:plazi:treatment:D9F61B9CB2E3504F966C1A8054E44122:53F601E45D8AB35B0869D281EA3F9584" country="Cameroon" latitude="3.3741333" longLatPrecision="1" longitude="12.733167" municipality="Dja Biosphere reserve" name="Somalomo">Somalomo</location>
, in the
<collectingMunicipality id="841CEE0F60B106B97668AF241A9C37D8">Dja Biosphere reserve</collectingMunicipality>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EH4BK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[12.733167,3.374133]}">
<geoCoordinate id="D6CC7A752E13DB1B72DC7859AE4A410B" degrees="3" direction="north" minutes="22.448" orientation="latitude" precision="1" value="3.3741333">3 ° 22.448 ' N</geoCoordinate>
,
<geoCoordinate id="ADDFA586134333E2943EE3FC52145BD2" degrees="12" direction="east" minutes="43.990" orientation="longitude" precision="1" value="12.733167">12 ° 43.990 ' E</geoCoordinate>
</named-content>
;
<date id="B781E23AAA7DC36D3E1EF8452880C52E" value="2022-04-11">
<collectingDate id="697046CEE23B94AB4C765F53F793032D" value="2022-04-11">11 Apr. 2022</collectingDate>
</date>
;
<collectorName id="0F46D27920F1A3ABA58DB1733DBEBF68">J. A. Yetchom Fondjo</collectorName>
leg.;
<named-content id="0824E46FB2CE196034BB1A27BB7C470B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/staatliches-museum-f%C3%BCr-naturkunde-karlsruhe" xlink_title="Staatliches Museum für Naturkunde Karlsruhe, Germany">
<collectionCode id="E47908068C6F0E6E20C7B4AAD7BED4C4" country="Germany" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" type="">SMNK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQ4BK" type="redescription">
<paragraph id="DF45030FC09D0DA6D74A76F05E79F6AF">
<heading id="E7FAFF927ADE42775002E3F16113167E" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="5D1FA4D6F2388890FEB8B93E7D14D798">
Lower side of the body clear; median dark band on pronotum disc narrow, the two contiguous clear bands faintly marked; posterior basal spot on the lateral lobes of pronotum narrow, not reaching the lower edge; metathoracic episternite with a straight, median stripe limited to the base of the segment; lower half of elytra shiny black, upper half brown; front and middle legs pale green. Inner and outer areas of posterior female hind femora pale, greenish yellow, with small pregenicular black ring; knees pale brown; posterior tibiae green, sometimes very dark; male cerci with small internal preapical lobule.
<emphasis id="84A5D16CCF6AF305CA529F57330DFA02" bold="true" italics="true">Epiphallus</emphasis>
(Fig.
<figureCitation id="0036F9E01B9994D4D165AFDFA38F40E7" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Internal genitalia of P. carnapi (A D), P. descampsi (E H), P. karschi zenkeri (I L), P. uniformis (M P) and P. verrucigena (O T)." figureDoi="10.3897/zookeys.1216.130270.figure15" httpUri="https://binary.pensoft.net/fig/1161003">15 M</figureCitation>
): bridge thin, narrow; anterior projections lobiform.
<emphasis id="D157548D0C83792007BB2D49758BB71C" bold="true" italics="true">Phallic complex</emphasis>
(Fig.
<figureCitation id="E8E01ED639A470BA916DED1212552541" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Internal genitalia of P. carnapi (A D), P. descampsi (E H), P. karschi zenkeri (I L), P. uniformis (M P) and P. verrucigena (O T)." figureDoi="10.3897/zookeys.1216.130270.figure15" httpUri="https://binary.pensoft.net/fig/1161003">15 N P</figureCitation>
): dorsal arch of cingulum rounded, almost firm, at the level of the ejaculatory sac, leaving endophallic sclerites almost entirely free, very divergent anteriorly; rami not bent; upper ectophallic sheath long; latero-ventral sclerite in profile, elbowed.
</paragraph>
<caption id="E93B29A6E6937F6EF59DD33DA6429287" ID-DOI="10.3897/zookeys.1216.130270.figure6" ID-arpha="ADAEF289-EDB3-54FB-8A90-ECC67DA861A5" httpUri="https://binary.pensoft.net/fig/1160994" startId="F6">
<paragraph id="79BBEC4A506A7DC06CB485824C9EE74F">
<label id="E066819916476F74A73E9B68A12956E3">Figure 6.</label>
</paragraph>
<paragraph id="6A9972FEE5CD3C9FC7C768D71C294E9A">
Images of holotypes, allotypes, letotype and neallotype of
<taxonomicName id="702E99FB5F32AB453627CFB3F1BCF288">
<emphasis id="EC989824AF03D2180065483DEA1CB9FA" italics="true">Pteropera</emphasis>
species
</taxonomicName>
in lateral view
<emphasis id="0B436B82F3882F7316AA06511A91E3D9" bold="true">A</emphasis>
<taxonomicName id="3AF71756122C0BE202F4AC590E94E434" baseAuthorityName="Karsch" baseAuthorityYear="1896" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="spleniata">
<emphasis id="C4AFC4F5928A455D881870C7145071A3" italics="true">P. spleniata</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="80B6EA8A413DABD0A19795800102E871" bold="true">B</emphasis>
<taxonomicName id="C3A7285F9B7E604FAF592D302E2E3854" baseAuthorityName="Karsch" baseAuthorityYear="1896" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="spleniata">
<emphasis id="99DD1DCAF6DC1E5F067516575052A521" italics="true">P. spleniata</emphasis>
</taxonomicName>
(holotype ♀)
<emphasis id="CBDEC9E545ABE400EC193FAF3A69A40B" bold="true">C</emphasis>
<taxonomicName id="129BEDE1312DB8F55246FF29138BC477" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="teocchii">
<emphasis id="021D2B457C87DE8DCB85BD6051639A53" italics="true">P. teocchii</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="D660B5154847DEEC91F960207A1F7213" bold="true">D</emphasis>
<taxonomicName id="8E3A5EC2BF68183693432DC097531606" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="teocchii">
<emphasis id="08349E38E021C0296A1F73FE07928745" italics="true">P. teocchii</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="152D0425A032224D1A8FA1EB3D175F3A" bold="true">E</emphasis>
<taxonomicName id="FC729BFD9596C4F192CF08EBB83936E7" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="thibaudi">
<emphasis id="1B54D2B83DFD599760E534C2BB6E25A8" italics="true">P. thibaudi</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="FE51A8953014F462A8356D4A4655E652" bold="true">F</emphasis>
<taxonomicName id="E64BF2F439F54820E4CECA0088FAECCF" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="thibaudi">
<emphasis id="B752522EB4B471D2B16D6D9DC154E9A6" italics="true">P. thibaudi</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="B19E89BEA99BAF7F3A654692726A5B6D" bold="true">G</emphasis>
<taxonomicName id="C21CAC55B0B2B13BAF2615A8A98C9D69" authorityName="Karsch" authorityYear="1891" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="verrucigena">
<emphasis id="F8FCCF57DFA42EB759AEADA70FFFC154" italics="true">P. verrucigena</emphasis>
</taxonomicName>
(lectotype ♂)
<emphasis id="B7FEC7A178302BCFD9411DC8245D60D5" bold="true">H</emphasis>
<taxonomicName id="4EB208EECB5E3E704C49D9A04DA3EF02" authorityName="Karsch" authorityYear="1891" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="verrucigena">
<emphasis id="0E2A3A34F98EC753DD6C5CA337804195" italics="true">P. verrucigena</emphasis>
</taxonomicName>
(♀)
<emphasis id="8002288CA48E1B5D4F8E890B335861FB" bold="true">I</emphasis>
<taxonomicName id="E1B29EA70BD89CE31FF6BB17F364FF07" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="villiersi">
<emphasis id="DFF6A5A504EE95A68D1FE1788B02E4C5" italics="true">P. villiersi</emphasis>
</taxonomicName>
(holotype ♂)
<emphasis id="60BEC4E93E5F4EA0567134791A3B0D57" bold="true">J</emphasis>
<taxonomicName id="760A28BC3C798B12614CFD5C1E540306" authorityName="Donskoff" authorityYear="1981" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="villiersi">
<emphasis id="B5409216F4E243B9CC0FC6DD01CFA804" italics="true">P. villiersi</emphasis>
</taxonomicName>
(allotype ♀)
<emphasis id="03E0AC722988746AC5C5D7C1CD2BF34F" bold="true">K</emphasis>
<taxonomicName id="54C2209D63ED377268EA0538026C91A5" authorityName="Bruner" authorityYear="1920" class="Insecta" family="Acrididae" genus="Pteropera" kingdom="Animalia" order="Orthoptera" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="1D787EA3AF1FE431A35B9B29C84F14B9" italics="true">P. uniformis</emphasis>
</taxonomicName>
(neallotype ♂).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EVDCK" type="remarks">
<paragraph id="526E80C03C6869840016B09E82B6AF3A">
<heading id="FFABF0D886C3EADAAEE411F5CD30DF9C" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="9A96DB83E8140CDACD3832162F1A69FE">The juveniles of this species is unknown.</paragraph>
</subSubSection>
<subSubSection id="SECID0E1DCK" type="distribution">
<paragraph id="1793CE9EF1A67FF9A1D47A3577563C6C">
<heading id="4DC2EF2F9D3A86EFAF22AB3C924100CF" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="F1A5729D19BAA5AF99DAA74FE8497B6D">
<collectingCountry id="94E38733368BD0BA32567B4DC1FF7C1A" name="Cameroon">Cameroon</collectingCountry>
(Fig.
<figureCitation id="732C3CC6B2F64E811157961FAA9FD05C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of Pteropera species in African rainforests." figureDoi="10.3897/zookeys.1216.130270.figure18" httpUri="https://binary.pensoft.net/fig/1161006">18 B</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,87 @@
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<mods:title id="985E51E85BC315D9372EC7F1F2077195">Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China</mods:title>
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<mods:namePart id="A6268DF2B2C95B43DAE4ED44E15B33E0">Wang, Bing</mods:namePart>
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<mods:namePart id="6B04140311586931F3C292213EF24F7F">Li, Jinglin</mods:namePart>
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<treatment id="F2FD569EA592550EBE1DAB9FDB88657A" ID-DOI="http://doi.org/10.5281/zenodo.13992748" ID-Zenodo-Dep="13992748" ID-arpha="F2FD569E-A592-550E-BE1D-AB9FDB88657A" LSID="urn:lsid:plazi:treatment:F2FD569EA592550EBE1DAB9FDB88657A" httpUri="http://treatment.plazi.org/id/F2FD569EA592550EBE1DAB9FDB88657A">
<subSubSection id="FA006F1EB501944734CBEDB668228164" type="nomenclature">
<paragraph id="3CE9C435A29A13F3FA20C64F1CCDFF76">
<heading id="191A8E8EAA07CA8F0371FF5160489CF9" reason="label">Genus</heading>
<taxonomicName id="6C4BB4DF2431A40FC15D875E2344B3A9" ID-CoL="398Y" authority="Thorell, 1898" authorityName="Thorell" authorityYear="1898" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="genus">
<emphasis id="28FF9855242780633A3293FAA11124C5" italics="true">Belisana</emphasis>
Thorell, 1898
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECUAE" type="type species">
<paragraph id="9EC1C4766332C0FF12BA514B3392AFEE">
<heading id="C75BD5982DB396903F38D7C2780DBFE1" reason="title">
<typeStatus id="C10573F264643790F8D7C21700BF1C29">Type</typeStatus>
species.
</heading>
</paragraph>
<paragraph id="238F52F0EEE73C156D810F71855A549D">
<taxonomicName id="27C539168D688444E20AB167177203B8" authority="Thorell, 1898" authorityName="Thorell" authorityYear="1898" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tauricornis">
<emphasis id="34DF7C13D30426248CD0EA9E0B8FEB15" italics="true">Belisana tauricornis</emphasis>
Thorell, 1898
</taxonomicName>
.
</paragraph>
</subSubSection>
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