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<document id="B63070610B5CDE83D3E204019BF75F49" ID-DOI="10.3897/phytokeys.254.136085" ID-publisher-id="136085" URI-arpha="914EC270-213E-56F6-86F1-32C95EF6B8CB" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1743434091246" checkinUser="pensoft" docAuthor="Siti-Munirah, Mat Yunoh &amp; Mohamad Alias, Shakri" docDate="2025" docId="5A527BBD3783531E8BDE11ED7CC76107" docLanguage="en" docName="PhytoKeys 254: 175-188" docOrigin="PhytoKeys 254" docSource="https://phytokeys.pensoft.net/article/136085/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Thismia aliasii Siti-Munirah 2025, sp. nov." docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="188" masterDocId="914EC270213E56F686F132C95EF6B8CB" masterDocTitle="Thismia aliasii (Thismiaceae), a new species from Terengganu, Peninsular Malaysia" masterLastPageNumber="188" masterPageNumber="175" pageNumber="175" updateTime="1744128412911" updateUser="admin">
<mods:mods id="72F5FA4770794F8D659B67D39B15EABD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="55351B567C55C6968732DD6B65624341">
<mods:title id="E2DE696E163BB6938C12310D804AA304">Thismia aliasii (Thismiaceae), a new species from Terengganu, Peninsular Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="19915FC6F720C5574A416C0F61A61431" type="personal">
<mods:role id="8944E3F47B263FB16817C5CBE19443F0">
<mods:roleTerm id="1C6FB3053FD476F8949DE66956671F4D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="39970DDF4016FCE6F2450D1BF29AFD21">Siti-Munirah, Mat Yunoh</mods:namePart>
<mods:nameIdentifier id="F38CABD3927194319CB0F85646106D26" type="ORCID">0000-0002-5062-9988</mods:nameIdentifier>
<mods:affiliation id="78466845E0F165AF673ED02F88D0CE6A">Forest Research Institute Malaysia, 52109 Kepong, Selangor, Malaysia</mods:affiliation>
</mods:name>
<mods:name id="C7063394E1326EB19F7E650D9C97CB3D" type="personal">
<mods:role id="3EB299AB658972E87770620FD77308A7">
<mods:roleTerm id="0121B3C7E89BA912A90A019BE9992687">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97805FF923D840D069C27543F62EFDD7">Mohamad Alias, Shakri</mods:namePart>
<mods:affiliation id="C7F16FC77A43556911CF6D023F727605">Jabatan Perhutanan Negeri Terengganu, 21500 Kuala Terengganu, Terengganu, Malaysia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="ADAFDF78F3AE967938960E8D4FA10561">text</mods:typeOfResource>
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<mods:titleInfo id="0CAE05A8E9B3D6AA90FE19BE793AB147">
<mods:title id="3FAF40E5258E9441DDC8180412DC7FC8">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="7AC24FE387A3E905C70D30A95680411E">
<mods:date id="248D29AD2F4ACF744E1E43ED97968B66">2025</mods:date>
<mods:detail id="56DC3E3EB0CF465AA1EBBB2B781EE5B9" type="pubDate">
<mods:number id="2E783605240DD9B9B23A5AD324C01B73">2025-03-31</mods:number>
</mods:detail>
<mods:detail id="EFEB60732C7DA76823ABC4EFA7BF7A4F" type="volume">
<mods:number id="9015E701177C2BE3322C33B1DB35D2F8">254</mods:number>
</mods:detail>
<mods:extent id="DC06757C4957F395E9E4D431E6BB2634" unit="page">
<mods:start id="888BA28A8B75505D78DA99EAB74BB40B">175</mods:start>
<mods:end id="E4EA132710C8D3E7B21E41AD4FFD152B">188</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9424016CF321C28AF5D2858EF3421A23">journal article</mods:classification>
<mods:identifier id="B6E6FF69B793C861230C51FA513A11DF" type="DOI">10.3897/phytokeys.254.136085</mods:identifier>
</mods:mods>
<treatment id="5A527BBD3783531E8BDE11ED7CC76107" ID-arpha="5A527BBD-3783-531E-8BDE-11ED7CC76107" ID-ipni="urn:lsid:ipni.org:names:77359488-1" LSID="urn:lsid:plazi:treatment:5A527BBD3783531E8BDE11ED7CC76107" LSID-IPNI="urn:lsid:ipni.org:names:77359488-1" httpUri="http://treatment.plazi.org/id/5A527BBD3783531E8BDE11ED7CC76107">
<subSubSection id="C21CAFF1A04E1FA636BCC71C276E6541" type="nomenclature">
<paragraph id="CF98BB03B1C63F18B2A1B26123A871C2">
<taxonomicName id="C0CCD92D889FA6D24530808DB5E95864" authority="Siti-Munirah" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii" status="sp. nov.">
<emphasis id="33F5FF6BB3E35E4D8A5E47318F3E200C" italics="true">Thismia aliasii</emphasis>
Siti-Munirah
</taxonomicName>
<taxonomicNameLabel id="9E9EC46849C30421BFE863AB8F1DB8A6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="AC34CFD81991E5779B84BB35DF69CF9C" type="description">
<paragraph id="17DAFBF509BB4029A09ED91D103A1400">
<figureCitation id="F82D83F2F24BA1689D0876DBF9768AEE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Thismia aliasii A plant with flower and roots B top view of flower showing tepals and annulus C bract (abaxial view) D leaf (abaxial view) E style and stigma F stamens (outer view) G stamens (inner view) H fruiting plant. All drawn by Mohamad Aidil Noordin from spirit material, FRI 91119." figureDoi="10.3897/phytokeys.254.136085.figure4" httpUri="https://binary.pensoft.net/fig/1295839">Figs 4</figureCitation>
,
<figureCitation id="52E402F6394D3C3AE099BD27C3DCC579" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Thismia aliasii A flowering plant (over-brightened by a camera flash) B anthetic flower, top view B 1 annulus, top view B 2 tip of appendage of outer tepal B 3 tip of appendage of inner tepal C distal portion of flower, lateral view C 1 flower, top view D floral tube (with its apex removed), side view D 1 outer surface of floral tube (showing tiny glands) E inner surface of flower tube, pistil and ovary E 1 longitudinal section of ovary and pistil with stigma F fruiting plant F 1 seeds in capsule F 2 seeds. Photos by Siti-Munirah (A, C 1: FRI 79119; A 1, B E: FRI 79167; E 1, F 1, F 2: FRI 91119, spirit material) and Mohamad Alias (F: FRI 91119)." figureDoi="10.3897/phytokeys.254.136085.figure5" httpUri="https://binary.pensoft.net/fig/1295840">5</figureCitation>
,
<figureCitation id="F35AAB21D27520902207BA81D3187AD0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Thismia aliasii A longitudinally dissected stamen tube showing the margins of individual connectives abaxially raised into conspicuous ribs (spirit material) A 1 stamens, view from inside B, C stamens, view from outside D, E stamen tube, oblique view from below and view from below (showing apical appendages and longitudinal ribs). Yellow arrows: filiform apical appendages; red arrows: acute hook-like subapical appendage; orange arrows: anthers. All photos by Siti-Munirah (A: FRI 91119; A 1, B E: FRI 79167)." figureDoi="10.3897/phytokeys.254.136085.figure6" httpUri="https://binary.pensoft.net/fig/1295841">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4LAE" type="diagnosis">
<paragraph id="0B92DA14966563E2C07696BB9DFD1CEE">
<heading id="F6F21706ECFE90C397CC6B93652C67A3" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="11598721F4472F116336B337CA0BB394">
<taxonomicName id="B8A32F3D194B95A21279B2798CC865E5" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="363F3449D94C5717099F1F6AA3936960" italics="true">Thismia aliasii</emphasis>
</taxonomicName>
is very similar to the species of the
<emphasis id="7155D510ABC001101BC266743AB4E95F" italics="true">
T. subsection
<taxonomicName id="46B36EFEA961D13E0FC245F40A5126F9" rank="subSection" subSection="Odoardoa">Odoardoa</taxonomicName>
</emphasis>
, as the tepals are the same in shape and size. However, the tepal appendages of the new species are of unequal length, the inner ones are longer than the outer ones, while the tepal appendages of the other species are of equal length. In addition, in the new species the margins of the individual connectives are raised abaxially into the conspicuous rib, whereas connectives are almost flat abaxially in the rest of the species.
</paragraph>
<caption id="86085DA3B18C5AE12C84BF466DA2325D" ID-DOI="10.3897/phytokeys.254.136085.figure4" ID-arpha="310CF7ED-B857-5239-977B-6DCC3E63FFD0" httpUri="https://binary.pensoft.net/fig/1295839" startId="F4">
<paragraph id="EA96746D20ADBBA64DD4E40257D1AC7C">
<label id="FBCEF3B966765679DD617A937B723EA9">Figure 4.</label>
</paragraph>
<paragraph id="272631F70141AFB3DC93B97E9245F8D9">
<taxonomicName id="66C18E9CDF4B9F43F92A84188190008F" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="259DF82C1AA72DB27B5D461FD0060F7F" italics="true">Thismia aliasii</emphasis>
</taxonomicName>
<emphasis id="D09CA53B3ADDCB25C85101DFD80CDB60" bold="true">A</emphasis>
plant with flower and roots
<emphasis id="D130DD19E69DB7BFB898BB89DF556E40" bold="true">B</emphasis>
top view of flower showing tepals and annulus
<emphasis id="B7A376C11B6CE2D779D9B21A0FC23A11" bold="true">C</emphasis>
bract (abaxial view)
<emphasis id="C838524EF1DF6C6B416201B0EE44E47E" bold="true">D</emphasis>
leaf (abaxial view)
<emphasis id="BBC5E9171A14274AF247EAE4D86404B4" bold="true">E</emphasis>
style and stigma
<emphasis id="5CBF22C1E657F5978433503376224A6E" bold="true">F</emphasis>
stamens (outer view)
<emphasis id="7B693F1D6FB3C0279DBF4DA8AD3991B7" bold="true">G</emphasis>
stamens (inner view)
<emphasis id="41A8F925D54C98BB4C8AE8970C11D6BC" bold="true">H</emphasis>
fruiting plant. All drawn by Mohamad Aidil Noordin from spirit material, FRI 91119.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EDOAE" type="material">
<paragraph id="631879A73AAD62CD7B2F7C939779D478">
<heading id="30F4311D1765A60105298F8AC59E34A2" reason="title">Type.</heading>
</paragraph>
<paragraph id="395E6E92452FA8207EA4CD0648824067">
<materialsCitation id="E406B933CD3B0F82F5AF6B06229D665A" collectingDate="2023-07-26" collectionCode="KEP" collectorName="Siti-Munirah" country="Malaysia" county="Hulu Terengganu District" elevation="640" latitude="4.659222" location="Chemerong Forest Eco Park" longLatPrecision="1" longitude="102.98295" municipality="Hutan Simpan Pasir Raja" specimenCode="SC 13201" specimenCount="1" stateProvince="Terengganu" typeStatus="holotype">
<collectingCountry id="4C285160D651C59E9D9BF13B794DFB99" name="Malaysia">Malaysia</collectingCountry>
. • Peninsular Malaysia:
<collectingRegion id="CAD5FEC3D11C0883D4ED69C09F79F2E3" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
,
<collectingCounty id="08A4479A6080FEA26463433E568CE144">Hulu Terengganu District</collectingCounty>
,
<collectingMunicipality id="8394242CC7395017D98DBA9DAB6363B1">Hutan Simpan Pasir Raja</collectingMunicipality>
,
<location id="6F602236526D72B23090A2E207232B58" LSID="urn:lsid:plazi:treatment:5A527BBD3783531E8BDE11ED7CC76107:6F602236526D72B23090A2E207232B58" country="Malaysia" county="Hulu Terengganu District" latitude="4.659222" longLatPrecision="1" longitude="102.98295" municipality="Hutan Simpan Pasir Raja" name="Chemerong Forest Eco Park" stateProvince="Terengganu">Chemerong Forest Eco Park</location>
,
<location id="F8F84AC796FF646B14C21F547CBBC016" LSID="urn:lsid:plazi:treatment:5A527BBD3783531E8BDE11ED7CC76107:F8F84AC796FF646B14C21F547CBBC016" country="Malaysia" county="Hulu Terengganu District" latitude="4.659222" longLatPrecision="1" longitude="102.98295" municipality="Hutan Simpan Pasir Raja" name="Gunung Chemerong" stateProvince="Terengganu">Gunung Chemerong</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMOAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[102.982944,4.659222]}">
<geoCoordinate id="90E49AD07C3908710EC5C4526D0AC91F" degrees="4" direction="north" minutes="39" orientation="latitude" precision="1" seconds="33.2" value="4.659222">4°39'33.2&quot;N</geoCoordinate>
,
<geoCoordinate id="DBA37EE14310087705B633A1F6C8A523" degrees="102" direction="east" minutes="58" orientation="longitude" precision="1" seconds="58.6" value="102.98295">102°58'58.6&quot;E</geoCoordinate>
</named-content>
, elev. ca
<quantity id="8DC32FC4B813A5771F0B9DE334EFF9FD" metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">
<elevation id="36B2F4A50D0C01E68F0F4F17031E526B" metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<date id="4B3EEC40CBBE6ABD69E62A9A7806F087" value="2023-07-26">
<collectingDate id="F6FADB8C3981BA0C34CC3F9C967E79EC" value="2023-07-26">26 July 2023</collectingDate>
</date>
,
<emphasis id="28B18E1F2126B8812297A72BDECDAB9B" italics="true">
<collectorName id="5FF69D15232EE19E57E38BB142E57416">Siti-Munirah</collectorName>
, FRI 79119
</emphasis>
(
<typeStatus id="D71BF7B295815C62F0D0DA0D8B916F72" type="holotype">holotype</typeStatus>
<abbrev id="ABBRID0ETOAE" xlink_title="Kepong Herbarium">
<collectionCode id="724FF899A33127304DA3630DB8A708A8" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" type="Herbarium">KEP</collectionCode>
</abbrev>
!, spirit collection, barcode no.
<specimenCode id="16F9ADAC58ED640DC0D63640C0F982BD" collectionCode="SC" country="USA" lsid="urn:lsid:biocol.org:col:15455" name="Salem College" type="Herbarium">SC 13201</specimenCode>
)
</materialsCitation>
.
</paragraph>
<caption id="ED89CC7FF5E3653517FE9647F0AC2CD3" ID-DOI="10.3897/phytokeys.254.136085.figure5" ID-arpha="2272C47F-3778-5B81-B03D-54820BEC0F7C" httpUri="https://binary.pensoft.net/fig/1295840" startId="F5">
<paragraph id="B07F6E687E1B770EAA0AA83389CD15D3">
<label id="3BD1566E3F5C45BD89CE3F0A77D6C42A">Figure 5.</label>
</paragraph>
<paragraph id="34E9ACAAD2238D294D9506898BEBDB5D">
<taxonomicName id="CBA48E2A343E3F266D575EC76C196A15" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="60E49A7633DBFDDE191F1539A7C47FEC" italics="true">Thismia aliasii</emphasis>
</taxonomicName>
<emphasis id="25D0F6C61BBFCC15D3B928C46B1D7EF5" bold="true">A</emphasis>
flowering plant (over-brightened by a camera flash)
<emphasis id="3F1058CE453E731A7A3E433C3C074D76" bold="true">B</emphasis>
anthetic flower, top view
<emphasis id="0A90A463B62B4B6CC2B09437B4D2C2F8" bold="true">B 1</emphasis>
annulus, top view
<emphasis id="356D797689C10898E339E478B2371BE8" bold="true">B 2</emphasis>
tip of appendage of outer tepal
<emphasis id="5589CD390104B15BC2B68668AA8B03DF" bold="true">B 3</emphasis>
tip of appendage of inner tepal
<emphasis id="91FC47757F0168348CB11CFABA42FD99" bold="true">C</emphasis>
distal portion of flower, lateral view
<emphasis id="19E5EF5C059B375A1D455C7054AD6404" bold="true">C 1</emphasis>
flower, top view
<emphasis id="F2782557840F73D1B11A07610F62979C" bold="true">D</emphasis>
floral tube (with its apex removed), side view
<emphasis id="27CF3EAA1669EB84502618A85E90F8FF" bold="true">D 1</emphasis>
outer surface of floral tube (showing tiny glands)
<emphasis id="626E7CC05B25CCC8CE9366A4C190F899" bold="true">E</emphasis>
inner surface of flower tube, pistil and ovary
<emphasis id="FD7B5D9A74445BDA5B2549AD640A765E" bold="true">E 1</emphasis>
longitudinal section of ovary and pistil with stigma
<emphasis id="D8799DC14D023AFA9C935AA6C3AC094A" bold="true">F</emphasis>
fruiting plant
<emphasis id="6F30D517C86C0988453B6599D20DB932" bold="true">F 1</emphasis>
seeds in capsule
<emphasis id="7D4DD6A05C2432CCA9C4A1AE00EEADDA" bold="true">F 2</emphasis>
seeds. Photos by Siti-Munirah (
<emphasis id="1C584D7323680D93BD9B4F23918F68CB" bold="true">A, C 1</emphasis>
: FRI 79119;
<emphasis id="2ADFC565BB474DA7905E06EA108F44D2" bold="true">A 1, B E</emphasis>
: FRI 79167;
<emphasis id="76E592998226450141E606AEF5024CCA" bold="true">E 1, F 1</emphasis>
,
<emphasis id="041267705F1D723BDBCE326938E84FD6" bold="true">F 2</emphasis>
: FRI 91119, spirit material) and Mohamad Alias (
<emphasis id="90CAEB2B6475EE26758A79A76C280220" bold="true">F</emphasis>
: FRI 91119).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EVQAE" type="description">
<paragraph id="B6ABB823F5907DF3CD4CC7BE47DC21B8">
<heading id="73D29C5110A34BC4F27F80E021EBEF06" reason="title">Description.</heading>
</paragraph>
<paragraph id="C94621928C473C30F56BD763D01F1B65">
Achlorophyllous herb, up to ca.
<quantity id="9AE67AC2BE7F2D627BC60AF9DD468101" metricMagnitude="-1" metricUnit="m" metricValue="1.1" unit="cm" value="11.0">11 cm</quantity>
tall, mostly glabrous (where not stated otherwise).
<emphasis id="52CD86D19F0A204BF41E0D228793D089" bold="true" italics="true">Roots</emphasis>
vermiform, unbranched, ca.
<quantity id="8BB0B751926D7A7EF19F4D075F9028AE" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
in diameter, light brown.
<emphasis id="0CF1874904A16313F4C24F3CAE77D821" bold="true" italics="true">Stem</emphasis>
erect, up to
<quantity id="55CD5210C53F59043CB018F6FD89B013" metricMagnitude="-2" metricUnit="m" metricValue="5.6" unit="mm" value="56.0">56 mm</quantity>
long,
<quantity id="010ED93F331D7B55D4A494AE91C782CE" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" unit="mm" value="1.9" valueMax="2.0" valueMin="1.8">1.82 mm</quantity>
in diameter, white-cream to light brown, bearing 12 flowers.
<emphasis id="92E07760C875CAA9FABF5CE363E2FF15" bold="true" italics="true">Leaves</emphasis>
up to 5, spirally arranged (arranged denser at stem base and looser at apex), triangular to narrowly triangular (shorter at stem base and longer and narrower at stem apex), scale-like, apex pointed, margin almost entire to slightly irregularly serrate, up to
<quantity id="868892E091EA13A056EE8633332FD405" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
long, ca.
<quantity id="9A966ED750EAEEE3DB036CE080A96408" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
wide at base, colored similar to stem.
<emphasis id="6C1215361DE27F5061A7E94F6FE4D912" bold="true" italics="true">Flowers</emphasis>
terminal and solitary, or in 2 - flowered terminal inflorescence, actinomorphic, ca.
<quantity id="007DBB0995254CAF59D2F564F4760561" metricMagnitude="-2" metricUnit="m" metricValue="5.8" unit="mm" value="58.0">58 mm</quantity>
long (including ovary, floral tube, tepals and tepal appendages when erected).
<emphasis id="4E9F122211CF320BDCE294EA8457B595" bold="true" italics="true">Involucral bracts</emphasis>
3, similar to upper leaves, triangular to narrowly triangular, scale-like, acute, with entire margin,
<quantity id="5B1A7F175982BC0702F970988CFF0A23" metricMagnitude="-3" metricUnit="m" metricValue="8.0" unit="mm" value="8.0">8 mm</quantity>
long, ca.
<quantity id="14CE92E1D71001EC9A1107968691CFB4" metricMagnitude="-3" metricUnit="m" metricValue="2.5" unit="mm" value="2.5">2.5 mm</quantity>
wide at base, white-brownish / light brown.
<emphasis id="F3ED09E66190CAB6689BB58D10852471" bold="true" italics="true">Pedicel</emphasis>
to
<quantity id="48DED273C8397905BA6735F2EA441BEB" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
long at anthesis, to ca.
<quantity id="40B76637DABA7BB0D6B163FD1AF88C1D" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
long after anthesis, white-brownish.
<emphasis id="FB0BCE60311BF7521606B7AFBBF87316" bold="true" italics="true">Floral tube</emphasis>
obovoid funnel-shaped,
<quantity id="6B9F17E6D8C00DA67B057C648163D4E2" metricMagnitude="-2" metricUnit="m" metricValue="2.0" unit="mm" value="20.0">20 mm</quantity>
long, ca.
<quantity id="28C684C17BF212D3C49260019B3DB40B" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
wide at base, ca.
<quantity id="157290B0F951A1812B254FA6500FEC6E" metricMagnitude="-3" metricUnit="m" metricValue="8.0" unit="mm" value="8.0">8 mm</quantity>
wide at middle, ca.
<quantity id="49C40AC359FE7A7E4E3A138ABC938BE7" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
wide distally;
<emphasis id="C516F480FFFA59B021E73461802CD507" bold="true" italics="true">outer surface</emphasis>
with minute glands, orange to sepia-brown, with 12 darker longitudinal ribs;
<emphasis id="B959CFE40857E7DA72EA427DC35A9422" bold="true" italics="true">inner surface</emphasis>
smooth or rough, almost similar color to the outer surface, with transverse bars.
<emphasis id="C8EE899BE3CDCF24A04D8768040CA830" bold="true" italics="true">Tepals</emphasis>
6, free, spreading, triangular, apex acute,
<quantity id="87F374B39B1B0F426780DC24C40EFB6D" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
long, ca.
<quantity id="CD5623A8D384AA0DF91B69F59F5A9174" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
wide (ca.
<quantity id="2EA4675615533CE1ECE37100FAEFA41A" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
at base), smooth, color similar to flower tube (dark brown at apex, yellow-orange at back), each apically bearing a tentacle-like appendage
<quantity id="7EC0ACC0E668D921C75C26D68AD8AA71" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
wide and narrowing towards the apex;
<emphasis id="DFC5018A530CE208FA0E392250E6B340" bold="true" italics="true">appendage of outer tepal</emphasis>
up to
<quantity id="9D9149D6C10205EA7DD96AAD83E8D0A5" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
long, dark brown;
<emphasis id="C51C11325730EDA4F1795BCBEAD153F3" bold="true" italics="true">appendage of inner tepal</emphasis>
ca.
<quantity id="7A20323178A42AB00C7CF26037941338" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" unit="mm" value="29.0" valueMax="32.0" valueMin="26.0">2632 mm</quantity>
long, brownish.
<emphasis id="B34FF6111C9C0F1B02F58D92864FA613" bold="true" italics="true">Annulus</emphasis>
moderately raised, hexagonal in outline, ca.
<quantity id="C0BD52E45B26AF5EE52A85D01ADC560B" metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
in diameter, with ring width ca
<quantity id="5711784546E03DB420798DDEE6098DD8" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
, brown-orange, aperture ca.
<quantity id="101DB0AACF6575675B2CD51EE1665746" metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
in diameter.
<emphasis id="FB5992444870B7BE226D53D2762D61EA" bold="true" italics="true">Stamens</emphasis>
6, pendulous from annulus;
<emphasis id="44C674C0CC921DF44159FF12849F422F" bold="true" italics="true">filaments</emphasis>
laterally pale orange, otherwise translucent white;
<emphasis id="0AF1297E4F0CDA24587E2A8D14ACB2BE" bold="true" italics="true">connectives</emphasis>
broad and flattened, fused laterally into a tube, shortly papillose, translucent white, with hairs around thecae, with margins abaxially raised to form conspicuous ribs along the sutures between the connectives, ribs distally slightly protruding beyond the apex of supraconnective;
<emphasis id="607AB584451B83CFE5896548DA670B6E" bold="true" italics="true">interstaminal glands</emphasis>
conspicuous, placed between bases of lateral appendages;
<emphasis id="60D05F42E08F066AF6788C3A465B762C" bold="true" italics="true">supraconnective</emphasis>
with 3 filiform apical appendages (central appendage ca.
<quantity id="70845ED442B3DD5635292F406927E257" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
long and paired appendages ca.
<quantity id="E8FAD15C16403AB315F9F75944271323" metricMagnitude="-3" metricUnit="m" metricValue="2.4" unit="mm" value="2.4">2.4 mm</quantity>
long) and with 2 acute hook-like subapical appendages ca.
<quantity id="BC567913DBBF051A551C43BE9E37AD96" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.6 mm</quantity>
long positioned adaxially between the apical appendages and the lateral appendage;
<emphasis id="3FD139D04FC95CAF962708F380961D50" bold="true" italics="true">lateral appendage</emphasis>
skirt-like, projecting towards the floral tube and not reaching the supraconnective apex, translucent white and brown on sides, lateral margin wavy.
<emphasis id="4CA5EAA2AC97F0593730BDFCB039C4CA" bold="true" italics="true">Ovary</emphasis>
red, pale cream proximally and blackish distally; unilocular;
<emphasis id="B91D32A6528EE9FCDFDE0C04FCB43EC0" bold="true" italics="true">placentas</emphasis>
3, free, column-like, arising at ovary base; ovules numerous.
<emphasis id="743309851E1EEBA2D42206D50E8B6F25" bold="true" italics="true">Style</emphasis>
dark blackish-orange, ca.
<quantity id="BBD7FD22F6D86911469819818A94DDA3" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
long; stigma ca.
<quantity id="6B66733D32CFEFAA2CAA2764D327F803" metricMagnitude="-3" metricUnit="m" metricValue="1.4" unit="mm" value="1.4">1.4 mm</quantity>
long, papillose, 3 - lobed, erect, lobes ± rectangular, bifurcate at the apex, black-greenish.
<emphasis id="6BB6AD713D9C793765A83BC93AFD44F9" bold="true" italics="true">Fruit</emphasis>
dehiscent, cup-shaped, ca.
<quantity id="1A477F0C66C24DCD4A3080DFEA9A8FF1" metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
high, ca.
<quantity id="277921A8B7DFA0FC429F3DE2746E13A4" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
in diameter, pale brownish, darker in the upper part.
<emphasis id="12A6100D31D5FC7046D5E8A2863CAC36" bold="true" italics="true">Seeds</emphasis>
long fusiform ca.
<quantity id="015E854AD97812EB3F3045AC060EE3BF" metricMagnitude="-4" metricUnit="m" metricValue="7.5" unit="mm" value="0.75">0.75 mm</quantity>
long, ca.
<quantity id="E2381C13E0D74FDE00D17A0951F0C215" metricMagnitude="-4" metricUnit="m" metricValue="2.5" unit="mm" value="0.25">0.25 mm</quantity>
wide.
</paragraph>
<caption id="D2C8118736235AE693F17DFD9ADD804F" ID-DOI="10.3897/phytokeys.254.136085.figure6" ID-arpha="1F38F81F-52D7-5B67-83B5-E25336B67E1E" httpUri="https://binary.pensoft.net/fig/1295841" startId="F6">
<paragraph id="C05588B9E09C180415B99AE4B12C9C74">
<label id="C95258542420FF1FBB1C2FEA746775A3">Figure 6.</label>
</paragraph>
<paragraph id="55B5B25A8FA2D29B8328822811A44345">
<taxonomicName id="CBEF616D7FFC4804787B547EFAEEB530" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="F50A930F484E0BC19F1B7F4732B6B3BA" italics="true">Thismia aliasii</emphasis>
</taxonomicName>
<emphasis id="D82F4AD4CB49EC878327BD4D65E7CAEE" bold="true">A</emphasis>
longitudinally dissected stamen tube showing the margins of individual connectives abaxially raised into conspicuous ribs (spirit material)
<emphasis id="5DD17A1A5DDC600503D80A9FC8966CF7" bold="true">A 1</emphasis>
stamens, view from inside
<emphasis id="77A1FF6F32088EED779D0661B65D7A8F" bold="true">B, C</emphasis>
stamens, view from outside
<emphasis id="88BF0AA56DF4EDC273CF636AE2E0D819" bold="true">D, E</emphasis>
stamen tube, oblique view from below and view from below (showing apical appendages and longitudinal ribs). Yellow arrows: filiform apical appendages; red arrows: acute hook-like subapical appendage; orange arrows: anthers. All photos by Siti-Munirah (
<emphasis id="D7DD86918A8FA89C3C5509B872E0B16B" bold="true">A</emphasis>
: FRI 91119;
<emphasis id="CF02F32524474FA626B053C0EC3C3F48" bold="true">A 1, B E</emphasis>
: FRI 79167).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EMUAE" type="material">
<paragraph id="B608268FFA30E99176AF7911DBE93FAB">
<heading id="3906B2C5FA96CCB5A42129D787C10E71" reason="title">Additional specimens examined.</heading>
</paragraph>
<paragraph id="9ACAC0E8419B1E43D5144530A85E5936">
<materialsCitation id="0340E7D7AA2BD317BD22FFC757B5D667" collectingDate="2019-10-03" collectingDateMax="2023-07-26" collectingDateMin="2019-10-03" collectionCode="KEP" collectorName="Mohamad Alias" country="Malaysia" county="Hulu Terengganu District" elevation="642" location="Chemerong Forest Eco Park" municipality="Pasir Raja FR" specimenCode="SC 13202" specimenCount="1" stateProvince="Terengganu">
<collectingCountry id="5A5BECD9954A424B68772BBC189B5ED8" name="Malaysia">Malaysia</collectingCountry>
. • Peninsular Malaysia:
<collectingRegion id="734FC827E213552A247ADA5B674A4F68" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
,
<collectingCounty id="1AABBDBD7F0FCD2F60A3A5A97A7E39AE">Hulu Terengganu District</collectingCounty>
,
<collectingMunicipality id="55858228AB567BA4C6DF0128BDDA1E54">
Pasir Raja
<abbrev id="ABBRID0ESUAE" xlink_title="Forest Reserve">FR</abbrev>
</collectingMunicipality>
,
<location id="B13546E33D9D68082D0CDA236C46D338" LSID="urn:lsid:plazi:treatment:5A527BBD3783531E8BDE11ED7CC76107:B13546E33D9D68082D0CDA236C46D338" country="Malaysia" county="Hulu Terengganu District" municipality="Pasir Raja FR" name="Chemerong Forest Eco Park" stateProvince="Terengganu">Chemerong Forest Eco Park</location>
,
<location id="681E2EB45056833A87E39A840A86755B" LSID="urn:lsid:plazi:treatment:5A527BBD3783531E8BDE11ED7CC76107:681E2EB45056833A87E39A840A86755B" country="Malaysia" county="Hulu Terengganu District" municipality="Pasir Raja FR" name="Gunung Chemerong" stateProvince="Terengganu">Gunung Chemerong</location>
, elev. ca
<quantity id="05F590263FC7937C3320804D545631CF" metricMagnitude="2" metricUnit="m" metricValue="6.42" unit="m" value="642.0">
<elevation id="95F3C36BE6A9B3CD9315B6C048AD18AA" metricMagnitude="2" metricUnit="m" metricValue="6.42" unit="m" value="642.0">642 m</elevation>
</quantity>
,
<date id="D3A6ADCB8B4E809B56CA0549FF5CC45B" value="2019-10-03">
<collectingDate id="0B42869BA63F1B839E6CAE27295FD118" value="2019-10-03">3 October 2019</collectingDate>
</date>
,
<collectorName id="D282178D1CA4A9AF06A92092CBE81F0D">Mohamad Alias</collectorName>
, FRI 91119 (
<abbrev id="ABBRID0EWUAE" xlink_title="Kepong Herbarium">
<collectionCode id="17633ECAED5AD817D16FB4D8C9DA2EBA" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" type="Herbarium">KEP</collectionCode>
</abbrev>
spirit collection, barcode no.
<specimenCode id="C353AC17CA4A498D810B220E5677480F" collectionCode="SC" country="USA" lsid="urn:lsid:biocol.org:col:15455" name="Salem College" type="Herbarium">SC 13202</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9D50CB19BC303EFDEC06F9FD1C87423D" collectingDate="2023-07-26" collectingDateMax="2023-07-26" collectingDateMin="2019-10-03" collectionCode="KEP" collectorName="Siti-Munirah" country="Malaysia" county="Hulu Terengganu District" elevation="640" location="Chemerong Forest Eco Park" municipality="Pasir Raja FR" specimenCode="SC 13203" specimenCount="1" stateProvince="Terengganu">
elev. ca
<quantity id="F03E8965BFF15A4E5BA27E95CF63E5FA" metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">
<elevation id="ACE336E70412B30BA372D81A7D7C161E" metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<date id="1902D34E961B40F23631688AD6FCD36B" value="2023-07-26">
<collectingDate id="4E06E8B80AA9A9B6CF649705BC7B3660" value="2023-07-26">26 July 2023</collectingDate>
</date>
,
<collectorName id="EE518E122653106B0A9CD72E2F1C5B99">Siti-Munirah</collectorName>
, FRI 79167 (
<abbrev id="ABBRID0E1UAE" xlink_title="Kepong Herbarium">
<collectionCode id="EF114E77D34E29938EEDAF41502AF32D" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" type="Herbarium">KEP</collectionCode>
</abbrev>
spirit collection, barcode no.
<specimenCode id="69B483C19B2BADB3EE877804ACE01ACD" collectionCode="SC" country="USA" lsid="urn:lsid:biocol.org:col:15455" name="Salem College" type="Herbarium">SC 13203</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5UAE" type="distribution">
<paragraph id="A37B820B0F5332ED528234D70C628DE9">
<heading id="3FAA7E1AFAD96339AD81A9928AC9794E" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="06E67DEA21DC277A16CC1CFA54A532EF">
Endemic to
<collectingRegion id="C70762956E673A8C3F38A57B9CFD5ECD" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
, Peninsular
<collectingCountry id="C76B68D1CC16CA82A42BB8F0359D65D5" name="Malaysia">Malaysia</collectingCountry>
. Currently only known from the type locality, Gunung Chemerong (Figs
<figureCitation id="0209687212E75A05C9590A0C57831A89" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Landscapes of Gn. Chemerong and the species of Thismia found there A the Chemerong waterfall with a height of 370 m B the Chemerong River not far from the habitat of T. aliasii C Thismia aliasii, an individual discovered by Mohamad Alias in 2019 (FRI 91119) D Thismia aseroe (FRI 79116) found on the trail at Hutan Lipur Chemerong. Photos by Siti-Munirah (A, B, D) and Mohamad Alias (C)." figureDoi="10.3897/phytokeys.254.136085.figure2" httpUri="https://binary.pensoft.net/fig/1295837">2</figureCitation>
,
<figureCitation id="5926B2C63996F588C44C83F991195CB3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Gunung Chemerong (black circle) in Terengganu, the only known locality of Thismia aliasii." figureDoi="10.3897/phytokeys.254.136085.figure3" httpUri="https://binary.pensoft.net/fig/1295838">3</figureCitation>
,
<figureCitation id="8B3ACE7CE2F9686261706AD0753978E8" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitat and habit of Thismia aliasii A plants in situ B plant with flower bud C plant with anthetic flower. All photos by Siti-Munirah." figureDoi="10.3897/phytokeys.254.136085.figure7" httpUri="https://binary.pensoft.net/fig/1295842">7</figureCitation>
).
</paragraph>
<caption id="C620B6422893466016699ED12FBDF6A6" ID-DOI="10.3897/phytokeys.254.136085.figure7" ID-arpha="5FD8FF82-3171-549D-B2B6-9D3C2B2E026B" httpUri="https://binary.pensoft.net/fig/1295842" startId="F7">
<paragraph id="1DED6EFA429FBFC8DF839E28345CBAA9">
<label id="4271D5631BFEDF72EDB7211FABF85462">Figure 7.</label>
</paragraph>
<paragraph id="5E7D09DDCD1904C336FC19D6F4500294">
Habitat and habit of
<taxonomicName id="02A1D5847285DCC12914077ACAD8EACE" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="E9AB7F6FE89E580362A1BF5561D8093A" italics="true">Thismia aliasii</emphasis>
</taxonomicName>
<emphasis id="58E4F72680B7EA95387D095AB7DBFAFF" bold="true">A</emphasis>
plants in situ
<emphasis id="36A88B5CE1AFAC5D34216982A5F4554C" bold="true">B</emphasis>
plant with flower bud
<emphasis id="28BDF28A6C4B3EBF6C9C162C7B2BF723" bold="true">C</emphasis>
plant with anthetic flower. All photos by Siti-Munirah.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EOWAE" type="ecology">
<paragraph id="CB408AB68678758FC14ACD1BE818C46B">
<heading id="DA24865350851E573F0C366249297C80" reason="title">Ecology.</heading>
</paragraph>
<paragraph id="3A3A74AF1EADD7BA78B9A8B136FEBD41">
The species inhabits moist shady areas of upper hill dipterocarp forest on moist soil at elevation of
<quantity id="68CED801C263B6A30E6B28911CBD4115" metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">640 m</quantity>
a. s. l. Flowering and fruiting recorded in July and October. The population was found in moist, shaded areas next to the main hiking trail to the summit of Gunung Chemerong Berembun Langsir at altitudes of
<quantity id="445AC43995E1CA6D6C8C193160700A7C" metricMagnitude="2" metricUnit="m" metricValue="6.4" unit="m" value="640.0">640 m</quantity>
above sea level. The species was found a few hundred meters away from the main river.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETWAE" type="etymology">
<paragraph id="4E81607B77DFC541CE6D45BEA4E22D86">
<heading id="2010C9E8823032D798B889C31468FA1D" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="60DFFD6261279F1D0E5FDAEB63F11DAF">
The species is named after Mr. Alias (the second author), a current ranger of the
<collectingRegion id="4B37C865592DF1CEFC864E405CD0353A" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
Forestry Department (
<collectionCode id="0F204BC58CF2E9E8ECC079A45F5DC89E">JPNT</collectionCode>
), who is also a freelance photographer (known as John Sp) and who was the first to discover the species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYWAE" type="conservation status">
<paragraph id="6717C44D376A9C07385CE64441C042F3">
<heading id="372DA03938F013AAC9A99A1357DE22C4" reason="title">Conservation status.</heading>
</paragraph>
<paragraph id="FE4C55B2564AB35CFADC6D3F8368295C">
Since 2019, several surveys have been conducted at an area of 4 hectares, but the species was observed only twice, with a total of 5 individuals recorded. The main threat to the population is the degradation of habitat quality due to hiking activities, as the population is found near the main hiking trail which is heavily used by hikers ascending to the summits of Gunung Chemerong Berembun Langsir. Therefore, according to IUCN Red List (IUCN 2024) this species is assessed as Critically Endangered,
<abbrev id="ABBRID0E5WAE" xlink_title="Critically Endangered">CR</abbrev>
B 2 ab (iii), D. More surveys are needed to determine the population size of the species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECXAE" type="notes">
<paragraph id="D3C8EB79F50BA95F71EC0A68E0D5A539">
<heading id="48BC1146DA4BC5BB6A5CB0557056071A" reason="title">Notes.</heading>
</paragraph>
<paragraph id="633BC398DC051A5A15BAC4785026F508">
<taxonomicName id="BE7755F682C0A664DDF2D8565AAC55DC" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="4C71897F53A73A16A54FC77CA90EA22D" italics="true">Thismia aliasii</emphasis>
</taxonomicName>
is easily recognized within the genus by the combination of the following characters: vermiform roots, almost uniform flower coloration (light to dark orange to sepia-brownish red), inner tepals free from each other, unequal tepal appendages (appendages of the outer tepals being shorter than those of the inner tepals), stamens each with 3 long filiform apical appendages, 2 acute subapical appendages and a lateral appendage, and connectives laterally thickened into conspicuous abaxial interstaminal ribs. Within the infrageneric classification of
<bibRefCitation id="55B3895145E7E959166C176CF618F302" DOI="10.11646/phytotaxa.319.3.2" author="Kumar P &amp; Gale SW &amp; Li J &amp; Bouamanivong S &amp; Fischer GA" etAl="et al." firstAuthor="Kumar" issue="3" journalOrPublisher="Phytotaxa" pagination="225-240" refId="B9" refString="Kumar P, Gale SW, Li J, Bouamanivong S, Fischer GA (2017) Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos and a key to subgeneric classification. Phytotaxa 319 (3): 225240. https://doi.org/10.11646/phytotaxa.319.3.2" title="Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos and a key to subgeneric classification." volume="319" year="2017">Kumar et al. (2017)</bibRefCitation>
,
<taxonomicName id="F89BBA6F226D33E91E4B1F550C34EC24" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="3DB2E46615C71373092350AA2637F88F" italics="true">T. aliasii</emphasis>
</taxonomicName>
is assigned here to
<emphasis id="02AF21371A4D5D42F0F2EF93179264C1" italics="true">
<taxonomicName id="A3F16A6E9B5BB724B8A781DA51E2644F" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="genus">Thismia</taxonomicName>
subgenus
<taxonomicName id="1E3C70C79F9AB06963BFAB72FC4EC3F2" authority="section Thismia" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="section" section="Thismia">
Thismia section
<emphasis id="56E387B16C359A8342C4B88F70792CEE" italics="true">Thismia subsection Odoardoa</emphasis>
</taxonomicName>
</emphasis>
Schltr., as long as it has free tepals equal in shape and size. The new species probably belongs to clade 5 defined by
<bibRefCitation id="1E8B4D4AF9AF40EA31DFA35FD44337C0" DOI="10.1093/botlinnean/boaa017" author="Shepeleva EA &amp; Schelkunov MI &amp; Hroneš M &amp; Sochor M &amp; Dančák M &amp; Merckx VSFT &amp; Kikuchi IABS &amp; Chantanaorrapint S &amp; Suetsugu K &amp; Tsukaya H &amp; Mar SS &amp; Luu HT &amp; Li H-Q &amp; Logacheva MD &amp; Nuraliev MS" etAl="et al." firstAuthor="Shepeleva" issue="3" journalOrPublisher="Botanical Journal of the Linnean Society" pagination="287-315" refId="B14" refString="Shepeleva EA, Schelkunov MI, Hroneš M, Sochor M, Dančák M, Merckx VSFT, Kikuchi IABS, Chantanaorrapint S, Suetsugu K, Tsukaya H, Mar SS, Luu HT, Li H-Q, Logacheva MD, Nuraliev MS (2020) Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits. Botanical Journal of the Linnean Society 193 (3): 287315. https://doi.org/10.1093/botlinnean/boaa017" title="Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits." volume="193" year="2020">Shepeleva et al. (2020)</bibRefCitation>
, which is characterized by free inner tepals and the presence of appendages of the outer and inner tepals. In addition, the fact that the appendages of the inner tepals are much longer than those of the outer tepals make
<taxonomicName id="90B30BD9B86E2DCAAE3AE629A435C807" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="756B3E0F12A70C281A3A9BD8A5A660EC" italics="true">T. aliasii</emphasis>
</taxonomicName>
similar to
<taxonomicName id="1115CA5290DDF2174A5EC678058F7DBD" authority="Becc." authorityName="Becc." class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="neptunis">
<emphasis id="6ADC1FB00DC1604C3635E4206792EA4C" italics="true">T. neptunis</emphasis>
Becc.
</taxonomicName>
, (Sochor at al. 2018), which belongs to
<emphasis id="2D3DD97AF50129E36DE0D720FC97C005" italics="true">
T. subgen.
<taxonomicName id="BBFFE6102CA2F727B71AAA22F8960F8F" authority="Jonker" authorityName="Jonker" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="subSection" section="Thismia" subSection="Brunonithismia">
Thismia sect.
<emphasis id="B766A74E72B5ED66B36A4DAF5963C839" italics="true">Thismia subsect. Brunonithismia</emphasis>
Jonker
</taxonomicName>
</emphasis>
according to
<bibRefCitation id="B9A34541B6A01C43A974256A34E4041E" DOI="10.11646/phytotaxa.319.3.2" author="Kumar P &amp; Gale SW &amp; Li J &amp; Bouamanivong S &amp; Fischer GA" etAl="et al." firstAuthor="Kumar" issue="3" journalOrPublisher="Phytotaxa" pagination="225-240" refId="B9" refString="Kumar P, Gale SW, Li J, Bouamanivong S, Fischer GA (2017) Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos and a key to subgeneric classification. Phytotaxa 319 (3): 225240. https://doi.org/10.11646/phytotaxa.319.3.2" title="Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos and a key to subgeneric classification." volume="319" year="2017">Kumar et al. (2017)</bibRefCitation>
, but was recovered as a member of clade 5 by
<bibRefCitation id="881FAC0D3DC0A7351AE73AD33EE95451" DOI="10.1093/botlinnean/boaa017" author="Shepeleva EA &amp; Schelkunov MI &amp; Hroneš M &amp; Sochor M &amp; Dančák M &amp; Merckx VSFT &amp; Kikuchi IABS &amp; Chantanaorrapint S &amp; Suetsugu K &amp; Tsukaya H &amp; Mar SS &amp; Luu HT &amp; Li H-Q &amp; Logacheva MD &amp; Nuraliev MS" etAl="et al." firstAuthor="Shepeleva" issue="3" journalOrPublisher="Botanical Journal of the Linnean Society" pagination="287-315" refId="B14" refString="Shepeleva EA, Schelkunov MI, Hroneš M, Sochor M, Dančák M, Merckx VSFT, Kikuchi IABS, Chantanaorrapint S, Suetsugu K, Tsukaya H, Mar SS, Luu HT, Li H-Q, Logacheva MD, Nuraliev MS (2020) Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits. Botanical Journal of the Linnean Society 193 (3): 287315. https://doi.org/10.1093/botlinnean/boaa017" title="Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits." volume="193" year="2020">Shepeleva et al. (2020)</bibRefCitation>
.
</paragraph>
<paragraph id="D238FAE1B5545987445C1E04F2FFE258">
The connectives of most
<taxonomicName id="EAA6EBD8C2AD416E707C46E9FB212B5E">
<emphasis id="DBEFD0DBECD0A0EEB7038788A4CB390E" italics="true">Thismia</emphasis>
species
</taxonomicName>
are usually flat and without any processes abaxially, making
<taxonomicName id="E6ADF6E4CE688D9054D2AE3193AA6750" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="3C70F501BC714A10FD6388B0CCCD796F" italics="true">T. aliasii</emphasis>
</taxonomicName>
unique not only among
<emphasis id="695D28F14C69A394737385AADE78C96A" italics="true">
T. subsect.
<taxonomicName id="0C571386BAD19EEBD6145D174E462FC2" rank="subSection" subSection="Odoardoa">Odoardoa</taxonomicName>
</emphasis>
but within the entire genus. The only other exceptions are species from
<emphasis id="BE991E88C66FDF8E36A77B4E8F4CB412" italics="true">T. sect. Geomitra</emphasis>
(Becc.) Kumar &amp; S. W. Gale which have ribs in the center of each stamen but differ by coralliform roots and flower mitre.
</paragraph>
<paragraph id="4E8A8E878206385D7B74785931DE030A">
We summarize
<taxonomicName id="ADD7F68D6420572BE43C3E0C706FB006" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="subSection" subSection="Odoardoa">
<emphasis id="10B86B56208C6756708D686CD26A2581" italics="true">Thismia subsect. Odoardoa</emphasis>
</taxonomicName>
to comprise 23 species based on the above-mentioned characteristics and following the classifications of
<bibRefCitation id="842827420EAE66DC798EB2D8A3F21A4A" DOI="10.11646/phytotaxa.319.3.2" author="Kumar P &amp; Gale SW &amp; Li J &amp; Bouamanivong S &amp; Fischer GA" etAl="et al." firstAuthor="Kumar" issue="3" journalOrPublisher="Phytotaxa" pagination="225-240" refId="B9" refString="Kumar P, Gale SW, Li J, Bouamanivong S, Fischer GA (2017) Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos and a key to subgeneric classification. Phytotaxa 319 (3): 225240. https://doi.org/10.11646/phytotaxa.319.3.2" title="Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos and a key to subgeneric classification." volume="319" year="2017">Kumar et al. (2017)</bibRefCitation>
and
<bibRefCitation id="D82D78E936A9F530BCC48F20847D0E45" DOI="10.1093/botlinnean/boaa017" author="Shepeleva EA &amp; Schelkunov MI &amp; Hroneš M &amp; Sochor M &amp; Dančák M &amp; Merckx VSFT &amp; Kikuchi IABS &amp; Chantanaorrapint S &amp; Suetsugu K &amp; Tsukaya H &amp; Mar SS &amp; Luu HT &amp; Li H-Q &amp; Logacheva MD &amp; Nuraliev MS" etAl="et al." firstAuthor="Shepeleva" issue="3" journalOrPublisher="Botanical Journal of the Linnean Society" pagination="287-315" refId="B14" refString="Shepeleva EA, Schelkunov MI, Hroneš M, Sochor M, Dančák M, Merckx VSFT, Kikuchi IABS, Chantanaorrapint S, Suetsugu K, Tsukaya H, Mar SS, Luu HT, Li H-Q, Logacheva MD, Nuraliev MS (2020) Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits. Botanical Journal of the Linnean Society 193 (3): 287315. https://doi.org/10.1093/botlinnean/boaa017" title="Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits." volume="193" year="2020">Shepeleva et al. (2020)</bibRefCitation>
with additions from
<bibRefCitation id="E605F2A48A4BA14964A923A4F006473D" DOI="10.11646/phytotaxa.280.3.8" author="Chantanaorrapint S &amp; Wai JS &amp; Poopath M &amp; Tetsana N" etAl="et al." firstAuthor="Chantanaorrapint" issue="3" journalOrPublisher="Phytotaxa" pagination="292-296" refId="B2" refString="Chantanaorrapint S, Wai JS, Poopath M, Tetsana N (2016) Thismia claviformis (Thismiaceae), a new species from the Thai-Malay Peninsula. Phytotaxa 280 (3): 292296. https://doi.org/10.11646/phytotaxa.280.3.8" title="Thismia claviformis (Thismiaceae), a new species from the Thai-Malay Peninsula." volume="280" year="2016">Chantanaorrapint et al. (2016)</bibRefCitation>
,
<bibRefCitation id="B84A0F91E078463C70C8CEBAF6DC17B6" DOI="10.5091/plecevo.2018.1387" author="Hroneš M &amp; Rejžek M &amp; Sochor M &amp; Svátek M &amp; Kvasnica J &amp; Egertová Z &amp; Pereira JT &amp; Nilus R &amp; Dančák M" etAl="et al." firstAuthor="Hrones" issue="1" journalOrPublisher="Plant Ecology and Evolution" pagination="110-118" refId="B7" refString="Hroneš M, Rejžek M, Sochor M, Svátek M, Kvasnica J, Egertová Z, Pereira JT, Nilus R, Dančák M (2018) Two new species of Thismia subsect. Odoardoa (Thismiaceae) from Borneo. Plant Ecology and Evolution 151 (1): 110118. https://doi.org/10.5091/plecevo.2018.1387" title="Two new species of Thismia subsect. Odoardoa (Thismiaceae) from Borneo." volume="151" year="2018">Hroneš et al. (2018)</bibRefCitation>
,
<bibRefCitation id="4ADE704F6C44A7FB882B8386F0C24E0D" DOI="10.11646/phytotaxa.360.2.10" author="Nishioka T &amp; Suetsugu K &amp; Repin R &amp; Kitayama K" etAl="et al." firstAuthor="Nishioka" issue="2" journalOrPublisher="Phytotaxa" pagination="174-178" refId="B11" refString="Nishioka T, Suetsugu K, Repin R, Kitayama K (2018) Thismia kinabaluensis (Thismiaceae), a new species from Mt. Kinabalu, Sabah, Borneo. Phytotaxa 360 (2): 174178. https://doi.org/10.11646/phytotaxa.360.2.10" title="Thismia kinabaluensis (Thismiaceae), a new species from Mt. Kinabalu, Sabah, Borneo." volume="360" year="2018">Nishioka et al. (2018)</bibRefCitation>
,
<bibRefCitation id="5467564F1C3BE71D3C9D8694F66A429D" DOI="10.3372/wi.50.50106" author="Dančák M &amp; Hroneš M &amp; Sochor M" etAl="et al." firstAuthor="Dancak" issue="1" journalOrPublisher="Willdenowia" pagination="65-76" refId="B5" refString="Dančák M, Hroneš M, Sochor M (2020) Thismia ornata and T. coronata (Thismiaceae), two new species from Sarawak, Borneo. Willdenowia 50 (1): 6576. https://doi.org/10.3372/wi.50.50106" title="Thismia ornata and T. coronata (Thismiaceae), two new species from Sarawak, Borneo." volume="50" year="2020">Dančák et al. (2020)</bibRefCitation>
,
<bibRefCitation id="86C8C2EFE5EB9D913279F7F5F1D128D3" DOI="10.3897/phytokeys.124.34728" author="Siti-Munirah MY &amp; Dome N" firstAuthor="Siti-Munirah" journalOrPublisher="PhytoKeys" pagination="123-137" refId="B15" refString="Siti-Munirah MY, Dome N (2019) Thismia domei and T. terengganuensis (Thismiaceae), two new species, and T. javanica, a new record from Terengganu, Peninsular Malaysia. PhytoKeys 124: 123137. https://doi.org/10.3897/phytokeys.124.34728" title="Thismia domei and T. terengganuensis (Thismiaceae), two new species, and T. javanica, a new record from Terengganu, Peninsular Malaysia." volume="124" year="2019">Siti-Munirah and Dome (2019)</bibRefCitation>
,
<bibRefCitation id="110D6E63B40ADD3658AB2120D014CB33" DOI="10.3897/phytokeys.242.120967" author="Siti-Munirah MY &amp; Hardy-Adrian C &amp; Mohamad-Shafiq S &amp; Irwan-Syah Z &amp; Hamidi AH" etAl="et al." firstAuthor="Siti-Munirah" journalOrPublisher="PhytoKeys" pagination="229-239" refId="B21" refString="Siti-Munirah MY, Hardy-Adrian C, Mohamad-Shafiq S, Irwan-Syah Z, Hamidi AH (2024) Thismia malayana (Thismiaceae), a new mycoheterotrophic species from Peninsular Malaysia. PhytoKeys 242: 229239. https://doi.org/10.3897/phytokeys.242.120967" title="Thismia malayana (Thismiaceae), a new mycoheterotrophic species from Peninsular Malaysia." volume="242" year="2024">Siti-Munirah et al. (2024)</bibRefCitation>
,
<bibRefCitation id="6E0E6913BEE68781BD5D4A8DB496C8B1" DOI="10.3372/wi.54.54102" author="Ya J-D &amp; Chen H-Y &amp; Zhang W &amp; Zhu R-B &amp; Cai J &amp; Yu W-B" etAl="et al." firstAuthor="Ya" issue="1" journalOrPublisher="Willdenowia" pagination="47-63" refId="B24" refString="Ya J-D, Chen H-Y, Zhang W, Zhu R-B, Cai J, Yu W-B (2024) Phylogenetic and biogeographical analyses of Thismia (Thismiaceae) support T. malipoensis as the eighth species in China. . Willdenowia 54 (1): 4763. https://doi.org/10.3372/wi.54.54102" title="Phylogenetic and biogeographical analyses of Thismia (Thismiaceae) support T. malipoensis as the eighth species in China. -." volume="54" year="2024">Ya et al. (2024)</bibRefCitation>
:
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,345 @@
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<paragraph id="27F9F32E84663C0C55AEFE1BF22EE5E9">
<title id="EB46CD999B3FCD3FCDEA326BEB8BBEB7">
A checklist of
<taxonomicName id="BFA8C9A750B4054448FC8E44DC550AA5" authority="Schltr., 1921" authorityName="Schltr." authorityYear="1921" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="subSection" section="Thismia" subGenus="Thismia" subSection="Odoardoa">
<emphasis id="864EADCC7A16318294659B086CB8257E" italics="true">Thismia subg. Thismia</emphasis>
sect.
<emphasis id="82D30A4AB3F176E18769642ABD455E57" italics="true">Thismia subsect. Odoardoa</emphasis>
Schltr., 1921
</taxonomicName>
.
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<paragraph id="F01CD14DB4522DFA9C8C798B5603D705">
Type species: —
<taxonomicName id="922598ECE47F0E2EEF7E00CE30E74213" authority="Becc., 1878" authorityName="Becc." authorityYear="1878" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aseroe">
<emphasis id="D73B4596660A22EC669C84F55E0CC231" italics="true">Thismia aseroe</emphasis>
Becc., 1878
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C965A7C36E31698B2641FD557CC25915" type="taxon_list">
<paragraph id="18C072A29403E12CF9C3C6A3A4EC43A1">
<title id="E5C7A3292AB1C2C2F2BFD6192736C9FA">Species included:</title>
</paragraph>
<paragraph id="C5E9D794D30492B54804E0F66F115157">
1)
<taxonomicName id="89686E9E6DF33E984634BE6A16D39C3A" authority="Holttum ex Jonker, 1948" authorityName="Jonker" authorityYear="1948" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="6EC0EFFD5FD7F74D49108C351BF48102" italics="true">T. alba</emphasis>
Holttum ex Jonker, 1948
</taxonomicName>
.
</paragraph>
<paragraph id="990B756DAB9D42EF2983C7063901051B">
2)
<taxonomicName id="2812A00C497B1426012B2137C8523983" authority="Siti-Munirah, 2025" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="112D23BC9448BCAF1363FAE634ACA17B" italics="true">T. aliasii</emphasis>
Siti-Munirah, 2025
</taxonomicName>
. (this study)
</paragraph>
<paragraph id="D4473CE817E675B7A598F0A67F127180">
3)
<taxonomicName id="D106774571DC9B46B20AB6869F1A9673" authority="K. Larsen &amp; Aver., 2007" authorityName="K. Larsen &amp; Aver." authorityYear="2007" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="annamensis">
<emphasis id="9A60EFAAE2E20229CC69A87B76C795A6" italics="true">T. annamensis</emphasis>
K. Larsen &amp; Aver., 2007
</taxonomicName>
.
</paragraph>
<paragraph id="7447B3C3927AFCC100754A81AC271750">
4)
<taxonomicName id="264BE18232F72AF66AEE645E97D18B0A" authority="Becc., 1878" authorityName="Becc." authorityYear="1878" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aseroe">
<emphasis id="BA268DED0AC221E0116BE17156AF6A9A" italics="true">T. aseroe</emphasis>
Becc., 1878
</taxonomicName>
.
</paragraph>
<paragraph id="1C1EFD5E22D94083653D8A632BFCF845">
5)
<taxonomicName id="2F1B1218ABB735B13CB0C31E22E27F96" authority="M. Hotta, 1967" authorityName="M. Hotta" authorityYear="1967" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="bifida">
<emphasis id="3C05E35C7CE2828A4A6242A71F6BA518" italics="true">T. bifida</emphasis>
M. Hotta, 1967
</taxonomicName>
.
</paragraph>
<paragraph id="AEE058E0F76DBDD4AE7986A60639DD71">
6)
<taxonomicName id="7C98FCDD6C847965012964F04DB0E395" authority="Tsukaya, Suetsugu &amp; Suleiman, 2017" authorityName="Tsukaya, Suetsugu &amp; Suleiman" authorityYear="2017" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="bryndonii">
<emphasis id="FF0FDBA9ABA8CCD64F9E2284FB0F8A30" italics="true">T. bryndonii</emphasis>
Tsukaya, Suetsugu &amp; Suleiman, 2017
</taxonomicName>
.
</paragraph>
<paragraph id="758423DA4C7AF5492B5F99E2CB46AFF1">
7)
<taxonomicName id="09FE7E939A4491F9A01C1D0437025A6F" authority="Ridl., 1895" authorityName="Ridl." authorityYear="1895" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="chrysops">
<emphasis id="9F4BDCECAB116C4252B3A3BFEB76DE59" italics="true">T. chrysops</emphasis>
Ridl., 1895
</taxonomicName>
.
</paragraph>
<paragraph id="381F82AF50E6E7673595FCFA1579F7D2">
8)
<taxonomicName id="F9A031116AFC6478483B896EF5EF6AA1" authority="Chantanaorr. &amp; J. Wai, 2016" authorityName="Chantanaorr. &amp; J. Wai" authorityYear="2016" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="claviformis">
<emphasis id="B34071C5E95AC4588BE7A37FDFE6D7D6" italics="true">T. claviformis</emphasis>
Chantanaorr. &amp; J. Wai, 2016
</taxonomicName>
.
</paragraph>
<paragraph id="658429BAB3BAFFD5FAC18C9CDAC5A1DF">
9)
<taxonomicName id="567D2738614D4AEFD013C9B6A58BFD0B" authority="Hrones, Sochor &amp; Dancak, 2018" authorityName="Hrones, Sochor &amp; Dancak" authorityYear="2018" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="cornuta">
<emphasis id="78C03785762B99FBA8386D34BFE6ED84" italics="true">T. cornuta</emphasis>
<bibRefCitation id="59D1CAF093E64F6509A5E67554B8DD07" DOI="10.11646/phytotaxa.340.1.5" author="Sochor M &amp; Egertová Z &amp; Hroneš M &amp; Dančák M" issue="1" journalOrPublisher="Phytotaxa" pagination="71-78" refId="B22" refString="Sochor M, Egertova Z, Hrones M, Dancak M (2018) Rediscovery of Thismia neptunis (Thismiaceae) after 151 years. Phytotaxa 340 (1): 71-78. https://doi.org/10.11646/phytotaxa.340.1.5" title="Rediscovery of Thismia neptunis (Thismiaceae) after 151 years." volume="340" year="2018">Hroneš, Sochor &amp; Dančák, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="360085AA4992A985A838E86A6AF32B90">
10)
<taxonomicName id="3E5974B6241B780C355E40090BB7FFB6" authority="Siti-Munirah, 2019" authorityName="Siti-Munirah" authorityYear="2019" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="domei">
<emphasis id="D435B514AFFB195420A238247A4A4CDC" italics="true">T. domei</emphasis>
<bibRefCitation id="8F0B77523A8AAFF109A7051EC7D307E1" DOI="10.3897/phytokeys.124.34728" author="Siti-Munirah MY &amp; Dome N" journalOrPublisher="PhytoKeys" pagination="123-137" refId="B15" refString="Siti-Munirah MY, Dome N (2019) Thismia domei and T. terengganuensis (Thismiaceae), two new species, and T. javanica, a new record from Terengganu, Peninsular Malaysia. PhytoKeys 124: 123-137. https://doi.org/10.3897/phytokeys.124.34728" title="Thismia domei and T. terengganuensis (Thismiaceae), two new species, and T. javanica, a new record from Terengganu, Peninsular Malaysia." volume="124" year="2019">Siti-Munirah, 2019</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8CCB5A1721D8963B916D45F604456E27">
11)
<taxonomicName id="D17A7C8DD9501A480555159C4A638BA9" authority="Chantanaorr., 2012" authorityName="Chantanaorr." authorityYear="2012" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="filiformis">
<emphasis id="20A2D3DEF62C3D035ED08F5135D6FDA4" italics="true">T. filiformis</emphasis>
Chantanaorr., 2012
</taxonomicName>
.
</paragraph>
<paragraph id="6867118EE893DA1EC1B558A52D1992C1">
12)
<taxonomicName id="55C8157772A04EF5624C0F8583F81A15" authority="Ridl., 1890" authorityName="Ridl." authorityYear="1890" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="fumida">
<emphasis id="4FAC5E80CB8895E2942DE04EA02F28BB" italics="true">T. fumida</emphasis>
Ridl., 1890
</taxonomicName>
.
</paragraph>
<paragraph id="719D0AD9AEEAD179A90573F06BE57D79">
13)
<taxonomicName id="5E0AA3721179D989CA8FA7A8F2177A7F" authority="Ridl., 1895" authorityName="Ridl." authorityYear="1895" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="grandiflora">
<emphasis id="08675A921DA003D4594DCB47DB2AD572" italics="true">T. grandiflora</emphasis>
Ridl., 1895
</taxonomicName>
.
</paragraph>
<paragraph id="C4B94444C86E8868B70513B3EBA0750F">
14)
<taxonomicName id="C2ECBC39F5756F5F5540E041CC6A67F6" authority="Dancak, Hrones, Kobrlova &amp; Sochor, 2013" authorityName="Dancak, Hrones, Kobrlova &amp; Sochor" authorityYear="2013" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="hexagona">
<emphasis id="C3A9D544D8E5C42F55812E51D90918A8" italics="true">T. hexagona</emphasis>
Dančák, Hroneš, Kobrlová &amp; Sochor, 2013
</taxonomicName>
.
</paragraph>
<paragraph id="85D9F2442B22775755BE355D79336336">
<taxonomicName id="FDDC691FE71C84F4FE8DA2A235E31C79" authority="Tsukaya, Suleiman &amp; H. Okada, 2014" authorityName="Tsukaya, Suleiman &amp; H. Okada" authorityYear="2014" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="variety" species="hexagona" variety="grandiflora">
<emphasis id="13C3F76955F570814373D0A570457045" italics="true">T. hexagona var. grandiflora</emphasis>
Tsukaya, Suleiman &amp; H. Okada, 2014
</taxonomicName>
.
</paragraph>
<paragraph id="E80FA33AF74343ADF60D44F17285B8E9">
15)
<taxonomicName id="52E8E5F151844D123F9FA4DF378E15B5" authority="Sochor &amp; Dancak, 2017" authorityName="Sochor &amp; Dancak" authorityYear="2017" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="inconspicua">
<emphasis id="6EC200CD89799546C26A6E021C69E85D" italics="true">T. inconspicua</emphasis>
Sochor &amp; Dančák, 2017
</taxonomicName>
.
</paragraph>
<paragraph id="AE6C4D27EAF397F037A7E4A642F2D565">
16)
<taxonomicName id="FB00A05B4CF10C72FE78F853220828DF" authority="T. Nishioka &amp; Suetsugu, 2018" authorityName="T. Nishioka &amp; Suetsugu" authorityYear="2018" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="kinabaluensis">
<emphasis id="8C9A533D2870EA9DB42CCB47FF49F34D" italics="true">T. kinabaluensis</emphasis>
T.
<bibRefCitation id="17D0D18FAC8760CC781F6FC9DA9738FD" DOI="10.11646/phytotaxa.360.2.10" author="Nishioka T &amp; Suetsugu K &amp; Repin R &amp; Kitayama K" issue="2" journalOrPublisher="Phytotaxa" pagination="174-178" refId="B11" refString="Nishioka T, Suetsugu K, Repin R, Kitayama K (2018) Thismia kinabaluensis (Thismiaceae), a new species from Mt. Kinabalu, Sabah, Borneo. Phytotaxa 360 (2): 174-178. https://doi.org/10.11646/phytotaxa.360.2.10" title="Thismia kinabaluensis (Thismiaceae), a new species from Mt. Kinabalu, Sabah, Borneo." volume="360" year="2018">Nishioka &amp; Suetsugu, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="D2AEE294B098093BA606D6CA7B6A78BE">
17)
<taxonomicName id="AFA8AE34D5213F8888B83390B0B71080" authority="Jarvie, 1996" authorityName="Jarvie" authorityYear="1996" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="lauriana">
<emphasis id="720404B5D1254C6CE9E4948958FAA0F4" italics="true">T. lauriana</emphasis>
Jarvie, 1996
</taxonomicName>
.
</paragraph>
<paragraph id="B87F07374D2308BD15DC0050A60FAD28">
18)
<taxonomicName id="DBC283AA7BC1964C5C8821B86FD345F2" authority="Siti-Munirah, Hardy-Adrian, Mohamad-Shafiq &amp; Irwan-Syah, 2024" authorityName="Siti-Munirah, Hardy-Adrian, Mohamad-Shafiq &amp; Irwan-Syah" authorityYear="2024" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="malayana">
<emphasis id="E72E69A0C2E1B99588370BAA5F0ADB45" italics="true">T. malayana</emphasis>
<bibRefCitation id="18663B4FBF15C4AE4D12BA4E0FD77AB3" DOI="10.3897/phytokeys.242.120967" author="Siti-Munirah MY &amp; Hardy-Adrian C &amp; Mohamad-Shafiq S &amp; Irwan-Syah Z &amp; Hamidi AH" journalOrPublisher="PhytoKeys" pagination="229-239" refId="B21" refString="Siti-Munirah MY, Hardy-Adrian C, Mohamad-Shafiq S, Irwan-Syah Z, Hamidi AH (2024) Thismia malayana (Thismiaceae), a new mycoheterotrophic species from Peninsular Malaysia. PhytoKeys 242: 229-239. https://doi.org/10.3897/phytokeys.242.120967" title="Thismia malayana (Thismiaceae), a new mycoheterotrophic species from Peninsular Malaysia." volume="242" year="2024">Siti-Munirah, Hardy-Adrian, Mohamad-Shafiq &amp; Irwan-Syah, 2024</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="5F26AA0C98BCC05F0416F04BF5AFE6A7">
19)
<taxonomicName id="B6E94315431203A3438620C445C4384F" authority="Tsukaya &amp; H. Okada, 2005" authorityName="Tsukaya &amp; H. Okada" authorityYear="2005" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="mullerensis">
<emphasis id="4633C9C7A6F8B747ADD6BFE0765811F4" italics="true">T. mullerensis</emphasis>
Tsukaya &amp; H. Okada, 2005
</taxonomicName>
.
</paragraph>
<paragraph id="2F4001DA350C2BB6ABE11E9F057B6EDF">
20)
<taxonomicName id="8122E7642A66F5332655390ECC33F535" authority="Becc., 1878" authorityName="Becc." authorityYear="1878" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="ophiuris">
<emphasis id="51E880E442F866DE50DB234FE73FBE1E" italics="true">T. ophiuris</emphasis>
Becc., 1878
</taxonomicName>
.
</paragraph>
<paragraph id="F9E58FBDDB75C6A5B5398261FFC32777">
21)
<taxonomicName id="A84A39AEDD0AD467E04ACBD802312920" authority="Dancak, Hrones &amp; Sochor, 2020" authorityName="Dancak, Hrones &amp; Sochor" authorityYear="2020" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="ornata">
<emphasis id="A1A96A4258150258BE9D5D07A0647400" italics="true">T. ornata</emphasis>
<bibRefCitation id="CF5FD1705CFF6BB7DD09F02F0C277A7E" DOI="10.3372/wi.50.50106" author="Dančák M &amp; Hroneš M &amp; Sochor M" issue="1" journalOrPublisher="Willdenowia" pagination="65-76" refId="B5" refString="Dancak M, Hrones M, Sochor M (2020) Thismia ornata and T. coronata (Thismiaceae), two new species from Sarawak, Borneo. Willdenowia 50 (1): 65-76. https://doi.org/10.3372/wi.50.50106" title="Thismia ornata and T. coronata (Thismiaceae), two new species from Sarawak, Borneo." volume="50" year="2020">Dančák, Hroneš &amp; Sochor, 2020</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="DB61F821722EADB1B7F29F071F82090A">
22)
<taxonomicName id="B42EB341CA507093D2FE9C078213779D" authority="Hrones, Dancak &amp; Rejzek, 2018" authorityName="Hrones, Dancak &amp; Rejzek" authorityYear="2018" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="pallida">
<emphasis id="41A017D22C83922781AE345EC5824C0D" italics="true">T. pallida</emphasis>
<bibRefCitation id="689CF7B5CF60D6519A3009925DCA0FCF" DOI="10.5091/plecevo.2018.1387" author="Hroneš M &amp; Rejžek M &amp; Sochor M &amp; Svátek M &amp; Kvasnica J &amp; Egertová Z &amp; Pereira JT &amp; Nilus R &amp; Dančák M" issue="1" journalOrPublisher="Plant Ecology and Evolution" pagination="110-118" refId="B7" refString="Hrones M, Rejzek M, Sochor M, Svatek M, Kvasnica J, Egertova Z, Pereira JT, Nilus R, Dancak M (2018) Two new species of Thismia subsect. Odoardoa (Thismiaceae) from Borneo. Plant Ecology and Evolution 151 (1): 110-118. https://doi.org/10.5091/plecevo.2018.1387" title="Two new species of Thismia subsect. Odoardoa (Thismiaceae) from Borneo." volume="151" year="2018">Hroneš, Dančák &amp; Rejzek, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="5CFF57372B55CFFAEFB86414ADEB4493">
23)
<taxonomicName id="9E2B670DC7E1CF4F53620D8667BCEA5D" authority="Ridl., 1915" authorityName="Ridl." authorityYear="1915" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="racemosa">
<emphasis id="BCD818C03D4A6F576B8A1CAAF214A48E" italics="true">T. racemosa</emphasis>
Ridl., 1915
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="54E755C9351F9C2F815FE81780443DAC" type="discussion">
<paragraph id="78FAA52082A3CDB8BDE30828405FAB9A">
Ten species of the subsection occur in Peninsular
<collectingCountry id="A6AA96B8C78D399E1C289486602D09D5" name="Malaysia">Malaysia</collectingCountry>
(
<taxonomicName id="BC22629DDA386413238A10F337E2BC36" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="17E673E04CB1CBEBB404A62B28CBFB1B" italics="true">T. alba</emphasis>
</taxonomicName>
,
<taxonomicName id="AFC0ED034F137A27C474F708E0CBDA0E" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aseroe">
<emphasis id="3C8AC236C38B33C59D17064F10B8EC8B" italics="true">T. aseroe</emphasis>
</taxonomicName>
,
<taxonomicName id="0E7BDB7B495AE357CA5B27FC59C7193A" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="chrysops">
<emphasis id="6A0BA670825EB95D19612F3B9024ECC7" italics="true">T. chrysops</emphasis>
</taxonomicName>
,
<taxonomicName id="688C303278B2A5FF8D9A980E2C59F0BA" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="domei">
<emphasis id="BC929648A63769584284457858AD105E" italics="true">T. domei</emphasis>
</taxonomicName>
,
<taxonomicName id="8C4AD2CC6E596FCE50130843EBC5678C" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="fumida">
<emphasis id="800DB8E209D358A88F42BA2894D2B8C0" italics="true">T. fumida</emphasis>
</taxonomicName>
,
<taxonomicName id="155D1377C018D077470ECBEEA2021A59" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="grandiflora">
<emphasis id="A6E6D3DB90A870C541720EAD14D02CEC" italics="true">T. grandiflora</emphasis>
</taxonomicName>
,
<emphasis id="B99D917BC2D5346403DF4B8DD3AA93CC" italics="true">
<taxonomicName id="B2E5C75D392A1EB5EB8A45BACF50C267" authorityName="Siti-Munirah, Hardy-Adrian, Mohamad-Shafiq &amp; Irwan-Syah" authorityYear="2024" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="malayana">T. malayana</taxonomicName>
<taxonomicName id="ED80FBBF589B0B21FAA27609A9FF2B5C" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="ornata">T. ornata</taxonomicName>
</emphasis>
,
<taxonomicName id="4C90BDCB4E9D74F1165374B6BAB850A8" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="racemosa">
<emphasis id="AA1632E52B430E71808FFE2C4135315C" italics="true">T. racemosa</emphasis>
</taxonomicName>
and
<taxonomicName id="EA7864E4DF195F9B06B34A2183F687B8" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="3BCEEFA4C9773AF8A1C5171D57F21439" italics="true">T. aliasii</emphasis>
</taxonomicName>
), with five of them (
<taxonomicName id="7F2CE1AFA903BE8ED77B2F09C833E1D6" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="alba">
<emphasis id="ACC23EB450B2B68257BCD75CCD77FE67" italics="true">T. alba</emphasis>
</taxonomicName>
,
<taxonomicName id="454005258F581C1E57D4696285D51C31" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="9A71CC4CF95DB1B8004B1FB8E904F6C6" italics="true">T. aliasii</emphasis>
</taxonomicName>
,
<taxonomicName id="DB90319DE2D3A3501084C1AF5434BE3A" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aseroe">
<emphasis id="898BF9FB8E53E1754A3FBC58D6E0F89B" italics="true">T. aseroe</emphasis>
</taxonomicName>
,
<taxonomicName id="53F17527BCCFBB101595E26BECCEEA6F" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="domei">
<emphasis id="B5C8D6062935AA278E6CE2C24B0FEE21" italics="true">T. domei</emphasis>
</taxonomicName>
and
<taxonomicName id="6EB3614E6CB6982E6FEB699036B6BD08" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="ornata">
<emphasis id="13012687AFA40DB13265381ECC8FBB71" italics="true">T. ornata</emphasis>
</taxonomicName>
) occurring in the state of
<collectingRegion id="BB6F3655E89C3A2BF23BC316AB8D7B63" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
. Whereas most of these species are characterized by tepal appendages equal in size and shape, in
<taxonomicName id="5F39EAC23717B28E3A15565D7AD620E4" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="29357CDD4FCCA33D6BF246F65147B8C9" italics="true">T. aliasii</emphasis>
</taxonomicName>
the appendages of the inner tepals are longer than those of the outer tepals. In addition, the stamen connectives laterally raised to form a conspicuous interstaminal rib are a unique feature of
<taxonomicName id="40F54AA039C1DD372D41FB0A494B214A" authorityName="Siti-Munirah" authorityYear="2025" class="Liliopsida" family="Burmanniaceae" genus="Thismia" kingdom="Plantae" order="Dioscoreales" phylum="Tracheophyta" rank="species" species="aliasii">
<emphasis id="D14C8D74825EF62E7ECF8F93D4B4910B" italics="true">T. aliasii</emphasis>
</taxonomicName>
that is not found in the other species of
<emphasis id="8A8E7A827092015122BFB1923075E128" italics="true">
T. subsect.
<taxonomicName id="23D64022E5B7CCF8030C6CECCA5E252C" rank="subSection" subSection="Odoardoa">Odoardoa</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>