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<document id="0532F0468C96E2B9395867BB267791B3" ENCODING="Cp1252" ID-GBIF-Dataset="54f50fd3-f456-4d76-b185-c2bfcc2963e9" ModsDocID="20017" checkinTime="1252781355209" checkinUser="donat" docAuthor="Wilson, E. O." docDate="2003" docId="B4F712295DB7BB7081BD7285ED6EE9FD" docLanguage="en" docName="20017_tristis_group_gg1.xml" docOrigin="Cambridge, MA: Harvard University Press" docSource="http://atbi.biosci.ohio-state.edu/HymOnline/reference-full.html?id=20017" docTitle="Pheidole rhinoceros" docType="treatment" docVersion="12" lastPageNumber="738" masterDocId="E2C3EA16EFBE44B5493D3E8099DC2AEA" masterDocTitle="Pheidole in the New World. A dominant, hyperdiverse ant genus." pageNumber="738" updateTime="1701311097043" updateUser="plazi">
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<mods:title id="F0DABC50C00561F0D0A201267B78B575">Pheidole in the New World. A dominant, hyperdiverse ant genus.</mods:title>
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<treatment id="B4F712295DB7BB7081BD7285ED6EE9FD" ID-DOI="http://doi.org/10.5281/zenodo.6275541" ID-GBIF-Taxon="100119430" ID-Zenodo-Dep="6275541" LSID="urn:lsid:plazi:treatment:B4F712295DB7BB7081BD7285ED6EE9FD" httpUri="http://treatment.plazi.org/id/B4F712295DB7BB7081BD7285ED6EE9FD" lastPageNumber="738" pageNumber="738">
<subSubSection id="036B63503029F167245F7324B657E0DE" type="nomenclature">
<paragraph id="37700CBCC1F1E3B7DC2E470A6541515C" pageNumber="738">
<taxonomicName id="354402B7134D00A587251688C7281408" ID-CoL="4FYK4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33965" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole rhinoceros Forel" lsidName-HNS="Pheidole rhinoceros Forel" order="Hymenoptera" pageNumber="738" phylum="Arthropoda" rank="species" species="rhinoceros">Pheidole rhinoceros</taxonomicName>
Forel
</paragraph>
</subSubSection>
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<paragraph id="8837C4CF40A29E958E58D1E135630524" pageNumber="738">
<taxonomicName id="BFF84BE49AC36A54686386F8A3225556" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33965" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole rhinoceros Forel" lsidName-HNS="Pheidole rhinoceros Forel" order="Hymenoptera" pageNumber="738" phylum="Arthropoda" rank="species" species="rhinoceros">Pheidole rhinoceros</taxonomicName>
f'orel 1899e: 73.
</paragraph>
</subSubSection>
<subSubSection id="E408606E8CFD4665497971BB4A280E58" type="nomenclature">
<paragraph id="3C90ABFC7BD7A1E88C3EEBE39B4CD4B0" pageNumber="738">Types Mus. Hist. Nat. Geneve.</paragraph>
</subSubSection>
<subSubSection id="FBDD7033AAF3261F21197F2E984410EF" type="etymology">
<paragraph id="8FB3F19C8ABA1BCBA4C8CC41B8B04040" pageNumber="738">Etymology Species name inspired by the hornlike extension of the midclypeus.</paragraph>
</subSubSection>
<subSubSection id="23588AECFF011B277C30428064A6C015" type="diagnosis">
<paragraph id="0B74499CE7C1F78ECE79A9BB6DC725FE" pageNumber="738">Diagnosis Similar in various ways to the species listed in the heading above, differing as follows.</paragraph>
</subSubSection>
<subSubSection id="14A234F674C217BD7669BEC82FD0749D" type="description">
<paragraph id="EDD7A0760D5AABC1440763FC41B6BEC7" pageNumber="738">Major: middle carina of clypeus raised into a right-angular hornlike protrusion, from which radiate carinulae traveling obliquely downward toward the anterior clypeal border; carinae originating on borders of frontal lobe curve inward toward the midline; posterior half of dorsal surface and all of pronotum smooth and shiny; propodeal spines slender, and vertical on the propodeal basal face. Minor: propodeal spines short, slender, and vertical on the propodeal basal face; head entirely smooth. Longino (1997) notes that considerable variation exist in the sculpturing of the minors in at least Honduras and Costa Rica. At elevations of about 500 m or higher, minors tend to have more extensive foveolation on the pronotum and mesopleuron than do minors at lower elevations. In Costa Rica at least, the two forms are nowhere sympatric but in Braulio Carrillo National Park approach within 300 m elevation and about 5 km distance. The two forms may represent distinct species, but Longino treats them as conspecifics, and this more conservative arrangement appears appropriate for the time being.</paragraph>
<paragraph id="27802A9ED5227182277B268CE04609C2" pageNumber="738">Measurements (mm) Lectotype major: HW 1.26, HL 1.40, SL 0.60, EL 0.16, PW 0.70. Paralectotype minor: HW 0.60, HL 0.64, SL 0.58, EL 0.10, PW 0.38. color Major and minor: concolorous light to medium brown.</paragraph>
</subSubSection>
<subSubSection id="8067492416BBF8F61E3A19D07B407FA5" type="distribution">
<paragraph id="796383861D8E98F8FB7349680E0442E8" pageNumber="738">
Range
<taxonomicName id="68C3BD06479585C064BC43492CB57408" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33965" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole rhinoceros Forel" lsidName-HNS="Pheidole rhinoceros Forel" order="Hymenoptera" pageNumber="738" phylum="Arthropoda" rank="species" species="rhinoceros">P. rhinoceros</taxonomicName>
is at least locally common in upland forests to as high as 1100 m, from Honduras to Colombia.
</paragraph>
</subSubSection>
<subSubSection id="867F2F045827BA049C0C7AACEEED3A5F" type="biology_ecology">
<paragraph id="AA33F4C154C6A3B8EFED84DF20E9E111" pageNumber="738">biology In Costa Rica Longino (1997) found the species nesting in pieces of dead wood on the ground and, in one instance, beneath the loose bark of a 1-meter-wide tree trunk in an old treefall. At Guanacaste Conservation Area, in montane rainforest, he observed workers tending homopterans under loose bark.</paragraph>
</subSubSection>
<subSubSection id="B3D7498574F4C572EAFE710164405928" type="materials_examined">
<paragraph id="D83BDA23EE09BD7A9BB2BB942920B1CD" pageNumber="738">figure Upper: major (side view, El Volcan Chiriqui, Panama, 900 m; full-face view, Cerro Campana, Panama, 800-950 m); compared with lectotype major. Lower: minor (Cerro Campana, Panama); compared with paralectotype. (Type locality: Bugaba, Panama.) Scale bars = 1 mm.</paragraph>
</subSubSection>
</treatment>
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<paragraph pageNumber="560">
<bibCitation author="Linnaeus" pagination="7" title="Hedysarum junceum" volumeTitle="Dec. Prima Pl. Horti Upsal." year="1762">
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<taxonomicName authority="Linnaeus" authorityName="Linnaeus" authorityYear="1762" class="Magnoliopsida" family="Fabaceae" genus="Hedysarum" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="junceum">
<emphasis bold="true" pageNumber="560">Hedysarum junceum</emphasis>
Linnaeus
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filius,
<emphasis italics="true" pageNumber="560">
<volumeTitle>Dec. Prima Pl. Horti Upsal.</volumeTitle>
</emphasis>
: 7. 1762
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</bibCitation>
.
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<paragraph pageNumber="560">
<typeStatus>Lectotype</typeStatus>
(Ali in
<emphasis italics="true" pageNumber="560">Biologia (Lahore)</emphasis>
12: 40. 1966): [icon] &quot;
<taxonomicName authority="in Linnaeus" authorityName="Linnaeus" authorityYear="1762" class="Magnoliopsida" family="Fabaceae" genus="Hedysarum" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="junceum">
<emphasis italics="true" pageNumber="560">Hedysarum junceum</emphasis>
</taxonomicName>
&quot; in Linnaeus filius,
<volumeTitle>Dec. Prima Pl. Horti Upsal.</volumeTitle>
: 7, t. 4. 1762.
</paragraph>
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<paragraph pageNumber="560">
Current name:
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<emphasis bold="true" italics="true" pageNumber="560">Lespedeza juncea</emphasis>
</taxonomicName>
(L. f.) Pers. (
<taxonomicName class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
:
<taxonomicName authorityName="Rudd" authorityYear="1968" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="subFamily" subFamily="Faboideae">Faboideae</taxonomicName>
).
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Type status:
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. Occurrence: recordedBy:
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; sex:
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; Location: country:
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; stateProvince: Tyrol; verbatimLocality: Zemmgrund; verbatimElevation:
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; habitat: alpine meadow / dwarf shrub community
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Type status:
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; stateProvince: Tyrol; verbatimLocality: Zemmgrund; verbatimElevation:
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<paragraph id="6C9B08239C7AF57A3C8F4D7288F75594" pageId="0" pageNumber="1115">Distribution</paragraph>
<paragraph id="192A8A4CD144BDAEEE389C247FC5AEB5" pageId="0" pageNumber="1115">
A species with boreal-alpine distribution (
<bibRefCitation id="03EE2203CEA1846F7DDFA0974D105244" author="Gusenleitner, Fritz" journalOrPublisher="Ber. nat. - med. Ver. Innsbruck" pageId="0" pageNumber="1115" pagination="199 - 221" title="Angaben zur Kenntnis der Bienengattung Andrena in Nordtirol (Oesterreich)" volume="72" year="1985">Gusenleitner 1985</bibRefCitation>
).
</paragraph>
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<subSubSection id="99D526C61F4E2D5A2B652C3589B88A15" pageId="0" pageNumber="1115" type="notes">
<paragraph id="D2536671CCB54A3E53327C9D9482B863" pageId="0" pageNumber="1115">Notes</paragraph>
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The species is oligolectic on
<taxonomicName id="29647919F24C4DCBFE06CB474A9D994F" family="Ericaceae" lsidName="Apoidea" pageId="0" pageNumber="1115" rank="family">Ericaceae</taxonomicName>
(
<bibRefCitation id="51FA8C238A626041FC7D2999F6DC6F19" author="Gusenleitner, F" editor="Schuster, R." journalOrPublisher="Oesterreichische Akademie der Wissenschaften, Wien" pageId="0" pageNumber="1115" title="Checklisten der Fauna Oesterreichs, No. 6" year="2012">Gusenleitner et al. 2012</bibRefCitation>
).
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Fabricius, 1793
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<emphasis italics="true">Dentes Primores</emphasis>
superiores IV: intermediis remotis.
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solitarii, approximati.
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plures, sublobati: antici longiores acutiores.
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