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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.153.52920" ID-GBIF-Dataset="3fba4968-0ad6-433b-b8bc-468497567f14" ID-PMC="PMC7381485" ID-Pensoft-Pub="1314-2003-153-111" ID-Pensoft-UUID="475172DBEA915C52B9909334D3DB5478" ID-PubMed="32765183" ModsDocID="1314-2003-153-111" checkinTime="1594919048947" checkinUser="pensoft" docAuthor="Bakalin, Vadim A., Klimova, Ksenia G. &amp; Nguyen, Van Sinh" docDate="2020" docId="B5C02B09C85D5B4796816ECE1F494680" docLanguage="en" docName="PhytoKeys 153: 111-154" docOrigin="PhytoKeys 153" docSource="http://dx.doi.org/10.3897/phytokeys.153.52920" docTitle="Calypogeia asakawana S. Hatt. ex Inoue, J. Jap. Bot. 39 (4): 107. 1964" docType="treatment" docVersion="4" id="475172DBEA915C52B9909334D3DB5478" lastPageNumber="111" masterDocId="475172DBEA915C52B9909334D3DB5478" masterDocTitle="A review of Calypogeia (Marchantiophyta) in the eastern Sino-Himalaya and Meta-Himalaya based mostly on types" masterLastPageNumber="154" masterPageNumber="111" pageNumber="111" updateTime="1668139873509" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Calypogeia (Marchantiophyta) in the eastern Sino-Himalaya and Meta-Himalaya based mostly on types</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bakalin, Vadim A.</mods:namePart>
<mods:affiliation>Botanical Garden-Institute, Vladivostok, Russia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-7897-4305</mods:nameIdentifier>
<mods:nameIdentifier type="email">vabakalin@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klimova, Ksenia G.</mods:namePart>
<mods:affiliation>Botanical Garden-Institute, Vladivostok, Russia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3229-1880</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Van Sinh</mods:namePart>
<mods:affiliation>Institute of Ecology and Biological Resources, Graduate University of Science and Technology, Vietnam Academy of Science and Technology, Ha Noi, Vietnam</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>153</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>111</mods:start>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.153.52920</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.153.52920</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-153-111</mods:identifier>
<mods:identifier type="Pensoft-UUID">475172DBEA915C52B9909334D3DB5478</mods:identifier>
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<treatment ID-GBIF-Taxon="165119435" LSID="urn:lsid:plazi:treatment:B5C02B09C85D5B4796816ECE1F494680" httpUri="http://treatment.plazi.org/id/B5C02B09C85D5B4796816ECE1F494680" lastPageNumber="111" pageId="0" pageNumber="111">
<subSubSection pageId="0" pageNumber="111" type="nomenclature">
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<taxonomicName LSID="B5C02B09-C85D-5B47-9681-6ECE1F494680" authority="S. Hatt. ex Inoue, J. Jap. Bot. 39 (4): 107. 1964." class="Jungermanniopsida" family="Calypogeiaceae" genus="Calypogeia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calypogeia asakawana" order="Jungermanniales" pageId="0" pageNumber="111" phylum="Marchantiophyta" rank="species" species="asakawana">Calypogeia asakawana S. Hatt. ex Inoue, J. Jap. Bot. 39 (4): 107. 1964.</taxonomicName>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Calypogeia asakawana S. Hatt. ex Inoue: A plant habit, fragment, ventral view B, C underleaves D Leaf Calypogeia ceylanica S. Hatt. et Mizut.: E, F leaves H underleaf J leaf middle cells Calypogeia cuspidata (Steph.) Steph. G plant habit, fragment, ventral view I underleaf L leaf M leaf middle cells Calypogeia decurrens (Steph.) Steph.: K plant habit, fragment, ventral view. Scale bars: 2 mm (G); 1 mm (E, F, K); 500 µm (A, D, L); 200 µm (H, I); 100 µm (B, C); 50 µm (J, M). A from Holotype TNS- 174359; B, C, D from authentic material of C. okamurana Steph. nom. herb., G 00067726; E, F, H, J Isotype G 00064248, G, I, L, M from Lectotype G 00069713; K from Isotype G 00060745." figureDoi="10.3897/phytokeys.153.52920.figure9" httpUri="https://binary.pensoft.net/fig/432970" pageId="0" pageNumber="111">Figure 9A-D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="reference_group">
<paragraph pageId="0" pageNumber="111">
=
<taxonomicName class="Jungermanniopsida" family="Calypogeiaceae" genus="Calypogeia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calypogeia okamurana" order="Jungermanniales" pageId="0" pageNumber="111" phylum="Marchantiophyta" rank="species" species="okamurana">Calypogeia okamurana</taxonomicName>
Steph. ex Bonner, Index Hepaticarum 3: 501, 1963 (nom. inval., Art. 38.1(a), no description). Authentic material: Japan. Iyo: Tokonabe Mt., 30 March 1913, S. Okamura no. 383 (original material, probably scheduled as the type: G [G00067726!], the specimen in all ways is similar to
<taxonomicName family="Calypogeiaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="C. asakawana" order="Jungermanniales" pageId="0" pageNumber="111" rank="species" species="asakawana">C. asakawana</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="materials_examined">
<paragraph pageId="0" pageNumber="111">Type.</paragraph>
<paragraph pageId="0" pageNumber="111">Japan. Tokyo: Asakawa Experimental forest, 12 June 1954, U. Mizushima, no. 5 (holotype: TNS [TNS-174359!]; isotype NICH [NICH-55582!]).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="111" type="remarks">
<paragraph pageId="0" pageNumber="111">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="111">
The species is regarded as Japanese endemic (known in Honshu only, cf.
<bibRefCitation author="Yamada, K" journalOrPublisher="The Journal of the Hattori Botanical Laboratory" pageId="0" pageNumber="111" pagination="1 - 106" refId="B77" refString="Yamada, K, Iwatsuki, Z, 2006. Catalog of the hepatics of Japan. The Journal of the Hattori Botanical Laboratory 99: 1 - 106" title="Catalog of the hepatics of Japan." volume="99" year="2006">Yamada and Iwatsuki 2006</bibRefCitation>
) and is characterized morphologically by small, deeply divided, bifid and spreading underleaves (slightly wider than stem) and rounded leaf apices.
</paragraph>
<paragraph pageId="0" pageNumber="111">
The description based on the holotype is as follows: plants pale brownish in herbarium, prostrate, translucent, slightly glistening, 1.1-1.5 mm wide and 8-20 mm long, forming loose mats; rhizoids rather numerous, originating as several unclear fascicles near underleaf bases and obliquely to erect spreading, attaching plants to the substratum; stem brownish (in the herbarium), 100-160
<normalizedToken originalValue="µm">µm</normalizedToken>
in diameter, branching not seen; leaves subhorizontally inserted, dorsally insertion line subtransverse to loosely arcuate, ventrally decurrent for 1/2-2/3 of stem width, obliquely lingulate, to obliquely ovate-lingulate, slightly convex to planar, slightly undulate along margin, laterally spreading, 650-700
<normalizedToken originalValue="×">x</normalizedToken>
400-550
<normalizedToken originalValue="µm">µm</normalizedToken>
, leaf apex rounded; underleaves loosely sinuately to transversely inserted, shortly decurrent (up 1/3 of stem width), 120-170
<normalizedToken originalValue="×">x</normalizedToken>
200-220
<normalizedToken originalValue="µm">µm</normalizedToken>
, bilobed by U-shaped sinus descending to 2/3 of leaf length, undivided zone 1-2 cells, lobes in the base 3-5 cells wide, midleaf cells oblong, thin-walled, 32-62
<normalizedToken originalValue="×">x</normalizedToken>
20-33
<normalizedToken originalValue="µm">µm</normalizedToken>
, trigones vestigial, cuticle virtually smooth; cells along leaf margin subquadrate to oblong, 20-38
<normalizedToken originalValue="µm">µm</normalizedToken>
, thin-walled, with very small concave trigones, cuticle smooth.
</paragraph>
<caption doi="10.3897/phytokeys.153.52920.figure9" httpUri="https://binary.pensoft.net/fig/432970" pageId="0" pageNumber="111" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="111">
<emphasis bold="true" pageId="0" pageNumber="111">Figure 9.</emphasis>
<taxonomicName class="Jungermanniopsida" family="Calypogeiaceae" genus="Calypogeia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calypogeia asakawana" order="Jungermanniales" pageId="0" pageNumber="111" phylum="Marchantiophyta" rank="species" species="asakawana">
<emphasis italics="true" pageId="0" pageNumber="111">Calypogeia asakawana</emphasis>
</taxonomicName>
S.Hatt. ex Inoue:
<emphasis bold="true" pageId="0" pageNumber="111">A</emphasis>
plant habit, fragment, ventral view
<emphasis bold="true" pageId="0" pageNumber="111">B, C</emphasis>
underleaves
<emphasis bold="true" pageId="0" pageNumber="111">D</emphasis>
Leaf
<taxonomicName class="Jungermanniopsida" family="Calypogeiaceae" genus="Calypogeia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calypogeia ceylanica" order="Jungermanniales" pageId="0" pageNumber="111" phylum="Marchantiophyta" rank="species" species="ceylanica">
<emphasis italics="true" pageId="0" pageNumber="111">Calypogeia ceylanica</emphasis>
</taxonomicName>
S.Hatt. et Mizut.:
<emphasis bold="true" pageId="0" pageNumber="111">E, F</emphasis>
leaves
<emphasis bold="true" pageId="0" pageNumber="111">H</emphasis>
underleaf
<emphasis bold="true" pageId="0" pageNumber="111">J</emphasis>
leaf middle cells
<taxonomicName class="Jungermanniopsida" family="Calypogeiaceae" genus="Calypogeia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calypogeia cuspidata" order="Jungermanniales" pageId="0" pageNumber="111" phylum="Marchantiophyta" rank="species" species="cuspidata">
<emphasis italics="true" pageId="0" pageNumber="111">Calypogeia cuspidata</emphasis>
</taxonomicName>
(Steph.) Steph.
<emphasis bold="true" pageId="0" pageNumber="111">G</emphasis>
plant habit, fragment, ventral view
<emphasis bold="true" pageId="0" pageNumber="111">I</emphasis>
underleaf
<emphasis bold="true" pageId="0" pageNumber="111">L</emphasis>
leaf
<emphasis bold="true" pageId="0" pageNumber="111">M</emphasis>
leaf middle cells
<taxonomicName class="Jungermanniopsida" family="Calypogeiaceae" genus="Calypogeia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calypogeia decurrens" order="Jungermanniales" pageId="0" pageNumber="111" phylum="Marchantiophyta" rank="species" species="decurrens">
<emphasis italics="true" pageId="0" pageNumber="111">Calypogeia decurrens</emphasis>
</taxonomicName>
(Steph.) Steph.:
<emphasis bold="true" pageId="0" pageNumber="111">K</emphasis>
plant habit, fragment, ventral view. Scale bars: 2 mm (
<emphasis bold="true" pageId="0" pageNumber="111">G</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="111">E, F, K</emphasis>
); 500
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="111">A, D, L</emphasis>
); 200
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="111">H, I</emphasis>
); 100
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="111">B, C</emphasis>
); 50
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="111">J, M</emphasis>
).
<emphasis bold="true" pageId="0" pageNumber="111">A</emphasis>
from Holotype TNS-174359;
<emphasis bold="true" pageId="0" pageNumber="111">B, C, D</emphasis>
from authentic material of
<taxonomicName lsidName="C. okamurana" pageId="0" pageNumber="111" rank="species" species="okamurana">
<emphasis italics="true" pageId="0" pageNumber="111">C. okamurana</emphasis>
</taxonomicName>
Steph. nom. herb., G00067726;
<emphasis bold="true" pageId="0" pageNumber="111">E, F, H, J</emphasis>
Isotype G00064248,
<emphasis bold="true" pageId="0" pageNumber="111">G, I, L, M</emphasis>
from Lectotype G00069713;
<emphasis bold="true" pageId="0" pageNumber="111">K</emphasis>
from Isotype G00060745.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/dez.65.21000" ID-GBIF-Dataset="03cce405-5887-4474-b040-35de18e8d020" ID-GBIF-Taxon="182223666" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="77D7AEC9294554E5BBDCABA37901022A" ID-Zenodo-Dep="1166397" ID-ZooBank="73DEE0F32BB04A21B4455E168FE50F54" LSID="urn:lsid:zoobank.org:act:003616EF-AF92-4481-864C-A91512EC417E" ModsDocID="1860-1324-65-1-1" checkinTime="1621372489409" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="B5C044608FCEF65CA145FF6A28D6B93E" docLanguage="en" docName="DeutEntomolZeit 65(1): 1-63" docOrigin="Deutsche Entomologische Zeitschrift 65 (1)" docPubDate="2018-01-18" docSource="http://dx.doi.org/10.3897/dez.65.21000" docTitle="Mecyclothorax octavius Liebherr 2018, sp. n." docType="treatment" docUuid="003616EF-AF92-4481-864C-A91512EC417E" docUuidSource="ZooBank" docVersion="3" id="77D7AEC9294554E5BBDCABA37901022A" lastPageId="29" lastPageNumber="30" masterDocId="77D7AEC9294554E5BBDCABA37901022A" masterDocTitle="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel" masterLastPageNumber="63" masterPageNumber="1" pageId="27" pageNumber="28" updateTime="1643465225333" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liebherr, James K.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
<mods:affiliation>Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
<mods:nameIdentifier type="email">jkl5@cornell.edu</mods:nameIdentifier>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-01-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>65</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>1</mods:start>
<mods:end>63</mods:end>
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<mods:url>http://dx.doi.org/10.3897/dez.65.21000</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.65.21000</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-1-1</mods:identifier>
<mods:identifier type="ZooBank">73DEE0F32BB04A21B4455E168FE50F54</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1166397</mods:identifier>
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<treatment ID-GBIF-Taxon="182223666" LSID="urn:lsid:zoobank.org:act:003616EF-AF92-4481-864C-A91512EC417E" httpUri="http://treatment.plazi.org/id/B5C044608FCEF65CA145FF6A28D6B93E" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<subSubSection pageId="27" pageNumber="28" type="nomenclature">
<paragraph pageId="27" pageNumber="28">
8.
<taxonomicName LSID="http://zoobank.org/003616EF-AF92-4481-864C-A91512EC417E" authority="Liebherr" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax octavius" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="octavius" status="sp. n.">Mecyclothorax octavius Liebherr</taxonomicName>
<taxonomicNameLabel pageId="27" pageNumber="28">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 16" captionStartId="F17" captionText="Figure 16. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. najtae." figureDoi="10.3897/dez.65.21000.figure16" httpUri="https://binary.pensoft.net/fig/179972" pageId="27" pageNumber="28">Figures 16C</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="27" pageNumber="28">, 17E-F</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F19" captionText="Figure 18. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. manautei; F, M. paniensis; G, M. mouensis." figureDoi="10.3897/dez.65.21000.figure18" httpUri="https://binary.pensoft.net/fig/179974" pageId="27" pageNumber="28">, 18C</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F20" captionText="Figure 19. Geographical distributions of Mecyclothorax (Phacothorax) spp." figureDoi="10.3897/dez.65.21000.figure19" httpUri="https://binary.pensoft.net/fig/179975" pageId="27" pageNumber="28">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="diagnosis">
<paragraph pageId="28" pageNumber="29">
<pageBreakToken pageId="28" pageNumber="29" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="28" pageNumber="29">
This species (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F17" captionText="Figure 16. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. najtae." figureDoi="10.3897/dez.65.21000.figure16" httpUri="https://binary.pensoft.net/fig/179972" pageId="28" pageNumber="29">16C</figureCitation>
) exhibits superficial similarity to
<taxonomicName genus="M." lsidName="M. fleutiauxi" pageId="28" pageNumber="29" rank="species" species="fleutiauxi">
<emphasis italics="true" pageId="28" pageNumber="29">M. fleutiauxi</emphasis>
</taxonomicName>
and
<taxonomicName genus="M." lsidName="M. jeanneli" pageId="28" pageNumber="29" rank="species" species="jeanneli">
<emphasis italics="true" pageId="28" pageNumber="29">M. jeanneli</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 9" captionStartId="F10" captionText="Figure 9. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli." figureDoi="10.3897/dez.65.21000.figure9" httpUri="https://binary.pensoft.net/fig/179965" pageId="28" pageNumber="29">9D-E</figureCitation>
), but differs in: 1, larger body size, standardized body length 5.8 mm; 2, presence of both anterior and posterior supraorbital setae; 3, an unmargined and depressed pronotal median base with sinuate lateral margins anterad the obtusely rounded hind angles; and 4, eyes of very small diameter, i.e. ocular lobe ratio = 0.71 versus ratios of 0.89-0.94 for individuals of the former two species. The male aedeagus is completely different, exhibiting a flagellum and flagellar sheath, and a dorsoventrally expanded apex (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="28" pageNumber="29">17E</figureCitation>
). Chaetotaxy +/+//+/-//+/2/+/+.
</paragraph>
</subSubSection>
<subSubSection lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph pageId="28" pageNumber="29">Description</paragraph>
<paragraph lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
(n = 1). Head capsule elongate, distinctly broader at the small, moderately convex eyes, ocular lobe gradually curving to meet gena well behind eye posterior margin; 20 ommatidia along horizontal diameter of eye; ocular ratio 1.45, ocular lobe ratio 0.71, EyL/EyD = 2.4; frontal grooves deep, sinuously convergent to just posterad clypeus, extended deeply onto clypeus; mandibles moderately elongate, mandibular ratio 1.8; ligular lateroanterior margin rounded to ligular seta, the two setae separated by one to two setal diameters; paraglossae thin, extended twice as far beyond ligular margin as their basal length to margin, apex observably spiculate (100
<normalizedToken originalValue="×">x</normalizedToken>
); antennae elongate, antennomere 9 length 2.3
<normalizedToken originalValue="×">x</normalizedToken>
maximal breadth; antennomere 3 with glabrous except for apical
<pageBreakToken pageId="29" pageNumber="30" start="start">ring</pageBreakToken>
of setae. Pronotum transverse, vase-shaped, median base depressed relative to broadly convex disc, lateral margins evenly curved until joined to long median peduncular collar, pronotal lateral margin obtusely concave at the juncture (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F17" captionText="Figure 16. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. najtae." figureDoi="10.3897/dez.65.21000.figure16" httpUri="https://binary.pensoft.net/fig/179972" pageId="29" pageNumber="30">16C</figureCitation>
); MPW/BPW = 2.4, MPW/PL = 1.1; front angles narrowly protruded, obtuse, pronotal apex constricted, APW/BPW = 1.35; median base trapezoidally depressed relative to convex disc, 8-10 small punctures present each side, lateral marginal bead continued along anterior half of basal peduncle, terminated at base of peduncle, median base unmargined posteriorly; median longitudinal impression very finely incised on disc, extended briefly anterad broad and shallow anterior transverse impression; anterior transverse impression very broad, very shallow, indistinctly traceable to front angles; proepisternum separated from prosternum by a deep, sinuous groove both anteriorly and ventrally; prosternum with anteapical impression that is continuous ventrally; prosternal process narrowly and deeply depressed between procoxae, depression extended anteriorly half the distance to prothoracic anterior margin, the depression broadest at anterior terminus. Elytra ellipsoid, broadest just anterad midlength; MEW/EL = 0.72; basal groove absent from very narrowly pedunculate elytral base, the very narrow lateral marginal depressions converging on posterior margin of pronotum; striae 1-7 very shallow though traceable, stria 8 deep, diverging from stria 9 just posterad anterior series of lateral elytral setae; all striae except 8 absent from elytral apex; apical elytral setae positioned only slightly nearer to suture than subapical seta; elytra appressed and conjoined at apex, the upraised sutural margin forming an elongate isosceles triangle connected to the narrowly beaded elytral apical margin. Pterothoracic mesepisternal anterior surface with eight broad punctures, the largest anterior puncture separated from the remainder by a vertical ridge; mesosternal-mesepisternal suture incomplete, obsolete to absent anteriorly (as in Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Thoracic structures of Mecyclothorax spp. A-B. Lateral view of elytral epipleuron and meso-and metapleurites: A, M. laterorectus; B, M. fleutiauxi. C-D. Metathoracic flight wing: C, M. punctipennis, showing fully developed flight wing folded under elytron; D, M. lissus, stenopterous wing rudiment that extends beyond metanotum, rudimentary wing venation present at base of strap. Abbreviations include: mms, mesepisternal-mesosternal suture; msp, mesepimeron; mst, mesepisternum; mtp, metepimeron; mtt, metepisternum." figureDoi="10.3897/dez.65.21000.figure3" httpUri="https://binary.pensoft.net/fig/179959" pageId="29" pageNumber="30">3B</figureCitation>
); metepisternum very short, maximum width/lateral length = 1.46 (difficult to measure due to fusion of metepisternum and metepimeron). Abdomen with first and second ventrites fused laterally, not suture visible; suture between second and third ventrites reduced though traceable laterally; ventrites 2-6 with broad, shallow, linear plaques near lateral margin. Microsculpture of frons reduced, an indistinct transverse mesh on a glossy surface, microsculpture on vertex an evident transverse mesh with some isodiametric sculpticells; pronotal disc and base with fine, elongate transverse mesh, sculpticell breadth 3-4
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length, surface iridescent; elytra with evident transverse mesh, sculpticell breadth 2-3
<normalizedToken originalValue="×length">xlength</normalizedToken>
; elytral apex with elongate transverse mesh, sculpticell breadth 3-4
<normalizedToken originalValue="×">x</normalizedToken>
length where traceable.
</paragraph>
<paragraph pageId="29" pageNumber="30">
<emphasis italics="true" pageId="29" pageNumber="30">Male genitalia</emphasis>
(n = 1). Antecostal margin of abdominal mediotergite IX angulate, not extended (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="29" pageNumber="30">17F</figureCitation>
); right paramere elongate, slightly expanded dorsally in basal half, narrowly extended apically (Fig.
<figureCitation captionStart="Figure 18" captionStartId="F19" captionText="Figure 18. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. manautei; F, M. paniensis; G, M. mouensis." figureDoi="10.3897/dez.65.21000.figure18" httpUri="https://binary.pensoft.net/fig/179974" pageId="29" pageNumber="30">18C</figureCitation>
), with 13 setae along the apical 2/3 of the ventral margin, those setae complementing two elongate apical setae; left paramere moderately broad basally, evenly narrowed to a moderately extended, narrow apex, two small setae present apically; aedeagal median lobe gracile, tubular, straight at midlength, with apex broadly expanded dorsoventrally, dorsally to am obtuse point, ventrally in a tightly rounded and projected margin; aedeagal median lobe internal sac with flagellum and flagellar sheath, apparently unilobate based on interpretation of uneverted unique type male (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="29" pageNumber="30">17E</figureCitation>
), a diffuse spicular patch apicad flagellar apparatus, no structure present at position of dorsal plate.
</paragraph>
<paragraph pageId="29" pageNumber="30">
<emphasis italics="true" pageId="29" pageNumber="30">Type</emphasis>
-Holotype male (MNHN): NEW CALEDONIA / Me Maoya, summit / plateau.12Nov2002. / Monteith &amp; Burwell // QM Berlesate 1080 / 21°22
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<geoCoordinate degrees="165" direction="east" minutes="20" orientation="longitude" precision="925" value="165.33333">165°20'E</geoCoordinate>
</normalizedToken>
/ rainforest, 1400 m/ / sieved litter // QUEENSLAND / MUSEUM LOAN / DATE: Nov. 2003 / No. LEN-1686 (green label) //
<emphasis italics="true" pageId="29" pageNumber="30">
New Caledonia
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
</emphasis>
revision / measured specimen 1 / J.K.
<bibRefCitation DOI="https://doi.org/10.3897/dez.63.10241" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="51" pageNumber="52" pagination="211 - 270" refId="B61" refString="Liebherr, JK, 2016. Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63: 211 - 270, DOI: https://doi.org/10.3897/dez.63.10241" title="Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia." url="https://doi.org/10.3897/dez.63.10241" volume="63" year="2016">Liebherr 2016</bibRefCitation>
♂1 // genitalia in polyethylene vial with glycerine // HOLOTYPE /
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
/
<taxonomicName authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax megalovatulus" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="megalovatulus">Mecyclothorax megalovatulus</taxonomicName>
/ J.K.Liebherr 2017 (black-bordered red label).
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="30" type="etymology">
<paragraph pageId="29" pageNumber="30">Etymology.</paragraph>
<paragraph pageId="29" pageNumber="30">
Given that the body form comprises a pair of ovoids (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F17" captionText="Figure 16. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. najtae." figureDoi="10.3897/dez.65.21000.figure16" httpUri="https://binary.pensoft.net/fig/179972" pageId="29" pageNumber="30">16C</figureCitation>
) suggesting the numeral eight, this species is given the epithet
<taxonomicName authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax octavius" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="octavius">Mecyclothorax octavius</taxonomicName>
. As a Latin personal name, denoting the eighth-born son, or a son born in the eighth month, the species name is to be treated as a noun.
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="30" type="distribution">
<paragraph pageId="29" pageNumber="30">Distribution and habitat.</paragraph>
<paragraph pageId="29" pageNumber="30">
The single specimen of this species was collected from sieved litter collected at the 1400 m summit of Mt. Maoya (Fig.
<figureCitation captionStart="Figure 19" captionStartId="F20" captionText="Figure 19. Geographical distributions of Mecyclothorax (Phacothorax) spp." figureDoi="10.3897/dez.65.21000.figure19" httpUri="https://binary.pensoft.net/fig/179975" pageId="29" pageNumber="30">19</figureCitation>
). This species is adelphotaxon to the species triplet
<taxonomicName genus="M." lsidName="M. plurisetosus" pageId="29" pageNumber="30" rank="species" species="plurisetosus">
<emphasis italics="true" pageId="29" pageNumber="30">M. plurisetosus</emphasis>
</taxonomicName>
(
<taxonomicName genus="M." lsidName="M. fleutiauxi" pageId="29" pageNumber="30" rank="species" species="fleutiauxi">
<emphasis italics="true" pageId="29" pageNumber="30">M. fleutiauxi</emphasis>
</taxonomicName>
+
<taxonomicName genus="M." lsidName="M. jeanneli" pageId="29" pageNumber="30" rank="species" species="jeanneli">
<emphasis italics="true" pageId="29" pageNumber="30">M. jeanneli</emphasis>
</taxonomicName>
), all of whom are most commonly found in arboreal microhabitats, supporting transformation of habitat preference to the arboreal realm in their common ancestor.
</paragraph>
</subSubSection>
</treatment>
</document>

45
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1859
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<paragraph id="B838681237F59F99F41D3934EAE968F6" pageNumber="416">Dois machos de Costa Rica, Hamburgfarm, Nevermann(Reichensperger N.° 307).</paragraph>
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