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<figureCitation id="13102A28FF89FF9006ACFC183CDD59F4" box="[159,237,901,927]" captionStart="FIGURE 1" captionStartId="2.[152,255,1219,1243]" captionTargetBox="[231,1354,191,1182]" captionTargetId="figure-28@2.[220,1368,179,1192]" captionTargetPageId="2" captionText="FIGURE 1. Bodies of D. (E.) meiacanthus sp. nov. A. male; B. female; Scale bar: 10 mm." pageId="1" pageNumber="398">Figs. 1</figureCitation>
,
<figureCitation id="13102A28FF89FF9006C8FC183D3A59F4" box="[251,266,901,927]" captionStart="FIGURE 2" captionStartId="2.[151,250,1904,1928]" captionTargetBox="[230,1358,1385,1866]" captionTargetId="figure-53@2.[217,1370,1373,1879]" captionTargetPageId="2" captionText="FIGURE 2. Genitalia of D. (E.) meiacanthus sp. nov. A. dorsal view; B. lateral view; C. ventral view." pageId="1" pageNumber="398">2</figureCitation>
)
</paragraph>
<paragraph id="8B9436ADFF89FF9006A4FC343E2359AA" blockId="1.[151,907,865,963]" box="[151,531,937,963]" pageId="1" pageNumber="398">
<emphasis id="B95FEABFFF89FF9006A4FC343D7859A8" bold="true" box="[151,328,937,963]" pageId="1" pageNumber="398">Chinese name:</emphasis>
小NJ矛÷ƶ兰DZ
</paragraph>
<paragraph id="8B9436ADFF89FF9006A4FC6C3DF55E38" blockId="1.[151,1437,1009,2007]" pageId="1" pageNumber="398">
<emphasis id="B95FEABFFF89FF9006A4FC6C3DA95E60" bold="true" box="[151,409,1009,1035]" pageId="1" pageNumber="398">Specimens examined.</emphasis>
<materialsCitation id="3B433CF0FF89FF900791FC6C3DF15E44" collectingDate="2024-06-22" collectionCode="SNNU" collectorName="Jinhong Dai" country="China" county="Male" location="Daweishan National Forest Park" municipality="Pingbian" pageId="1" pageNumber="398" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B95FEABFFF89FF900791FC6C3E265E60" bold="true" box="[418,534,1009,1035]" pageId="1" pageNumber="398">
<typeStatus id="5490880FFF89FF900791FC6C3E215E60" box="[418,529,1009,1035]" pageId="1" pageNumber="398" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F33C763DFF89FF90042CFC6C3E565E60" box="[543,614,1009,1035]" name="China" pageId="1" pageNumber="398">China</collectingCountry>
:
<collectingCounty id="62F54E21FF89FF900440FC6C3E9F5E60" box="[627,687,1009,1035]" pageId="1" pageNumber="398">Male</collectingCounty>
,
<collectingRegion id="49EFF84FFF89FF90048FFC6F3F255E67" box="[700,789,1010,1036]" country="China" name="Yunnan" pageId="1" pageNumber="398">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BF0ACD7FF89FF900511FC6C3FB85E60" box="[802,904,1009,1035]" pageId="1" pageNumber="398">Pingbian</collectingMunicipality>
,
<location id="8EF46076FF89FF9005A5FC6C393B5E60" LSID="urn:lsid:plazi:treatment:038287BBFF89FF9206A4FCFC3D745856:8EF46076FF89FF9005A5FC6C393B5E60" box="[918,1291,1009,1036]" country="China" county="Male" municipality="Pingbian" name="Daweishan National Forest Park" pageId="1" pageNumber="398" stateProvince="Yunnan">
<location id="8EF46076FF89FF9005A5FC6C38FC5E67" LSID="urn:lsid:plazi:treatment:038287BBFF89FF9206A4FCFC3D745856:8EF46076FF89FF9005A5FC6C38FC5E67" box="[918,1228,1009,1036]" country="China" county="Male" municipality="Pingbian" name="Daweishan National Forest" pageId="1" pageNumber="398" stateProvince="Yunnan">Daweishan National Forest</location>
Park
</location>
,
<date id="FF95106DFF89FF90032AFC6C39A85E60" box="[1305,1432,1009,1035]" pageId="1" pageNumber="398" value="2024-06-22">
<collectingDate id="EFD1E985FF89FF90032AFC6C39A85E60" box="[1305,1432,1009,1035]" pageId="1" pageNumber="398" value="2024-06-22">22-vi-2024</collectingDate>
</date>
,
<collectorName id="26DE537BFF89FF9006A4FB883D2F5E44" box="[151,287,1045,1071]" pageId="1" pageNumber="398">Jinhong Dai</collectorName>
coll. (
<collectionCode id="ED3AAE68FF89FF900755FB883D895E44" box="[358,441,1045,1071]" pageId="1" pageNumber="398">SNNU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B433CF0FF89FF9007F8FB883DF15E38" collectingDate="2024-06-22" collectionCode="SNNU" collectorName="Jinhong Dai" country="China" county="Pingbian" location="Daweishan National Forest" municipality="Daweishan National Forest Park" pageId="1" pageNumber="398" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B95FEABFFF89FF9007F8FB883E7A5E44" bold="true" box="[459,586,1045,1071]" pageId="1" pageNumber="398">
<typeStatus id="5490880FFF89FF9007F8FB883E755E44" box="[459,581,1045,1071]" pageId="1" pageNumber="398" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D2DFD24FF89FF900462FB883EF15E44" box="[593,705,1045,1071]" count="2" pageId="1" pageNumber="398" type="female">2 females</specimenCount>
.
<collectingRegion id="49EFF84FFF89FF9004F9FB8B3F135E5B" box="[714,803,1046,1072]" country="China" name="Yunnan" pageId="1" pageNumber="398">Yunnan</collectingRegion>
,
<collectingCounty id="62F54E21FF89FF90051DFB883FA45E44" box="[814,916,1045,1071]" pageId="1" pageNumber="398">Pingbian</collectingCounty>
,
<collectingMunicipality id="6BF0ACD7FF89FF9005ACFB88393D5E44" box="[927,1293,1045,1072]" pageId="1" pageNumber="398">
<location id="8EF46076FF89FF9005ACFB8838E05E5B" LSID="urn:lsid:plazi:treatment:038287BBFF89FF9206A4FCFC3D745856:8EF46076FF89FF9005ACFB8838E05E5B" box="[927,1232,1045,1072]" country="China" county="Pingbian" municipality="Daweishan National Forest Park" name="Daweishan National Forest" pageId="1" pageNumber="398" stateProvince="Yunnan">Daweishan National Forest</location>
Park
</collectingMunicipality>
,
<date id="FF95106DFF89FF90032AFB8839A85E44" box="[1305,1432,1045,1071]" pageId="1" pageNumber="398" value="2024-06-22">
<collectingDate id="EFD1E985FF89FF90032AFB8839A85E44" box="[1305,1432,1045,1071]" pageId="1" pageNumber="398" value="2024-06-22">22-vi-2024</collectingDate>
</date>
,
<collectorName id="26DE537BFF89FF9006A4FBA43D2F5E38" box="[151,287,1081,1107]" pageId="1" pageNumber="398">Jinhong Dai</collectorName>
coll. (
<collectionCode id="ED3AAE68FF89FF900755FBA43D895E38" box="[358,441,1081,1107]" pageId="1" pageNumber="398">SNNU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3316526FF89FF9006F4FBC03D2E5EF0" pageId="1" pageNumber="398" type="etymology">
<paragraph id="8B9436ADFF89FF9006F4FBC03D2E5EF0" blockId="1.[151,1437,1009,2007]" pageId="1" pageNumber="398">
<emphasis id="B95FEABFFF89FF9006F4FBC03D7E5E1C" bold="true" box="[199,334,1117,1143]" pageId="1" pageNumber="398">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C2B4D2EFF89FF90040EFBC03EE75E1C" authorityName="Ma &amp; Dai" authorityYear="2025" box="[573,727,1117,1143]" class="Insecta" family="Gryllidae" genus="Duolandrevus" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="398" phylum="Arthropoda" rank="species" species="meiacanthus">
<emphasis id="B95FEABFFF89FF90047AFBC03EE75E1C" box="[585,727,1117,1143]" italics="true" pageId="1" pageNumber="398">meiacanthus</emphasis>
</taxonomicName>
” refers this species having epiphallic median lobe shaped as a small spine.
</paragraph>
</subSubSection>
<subSubSection id="C3316526FF89FF9206F4FB383DE65A92" lastPageId="3" lastPageNumber="400" pageId="1" pageNumber="398" type="description">
<paragraph id="8B9436ADFF89FF9006F4FB383E0B5CDC" blockId="1.[151,1437,1009,2007]" pageId="1" pageNumber="398">
<emphasis id="B95FEABFFF89FF9006F4FB383E345ED4" bold="true" box="[199,516,1189,1215]" pageId="1" pageNumber="398">
Description (
<figureCitation id="13102A28FF89FF900751FB383D985ED4" box="[354,424,1189,1215]" captionStart="FIGURE 1" captionStartId="2.[152,255,1219,1243]" captionTargetBox="[231,1354,191,1182]" captionTargetId="figure-28@2.[220,1368,179,1192]" captionTargetPageId="2" captionText="FIGURE 1. Bodies of D. (E.) meiacanthus sp. nov. A. male; B. female; Scale bar: 10 mm." pageId="1" pageNumber="398">Fig. 1</figureCitation>
). Male.
</emphasis>
Body sized medium for this genus.
<emphasis id="B95FEABFFF89FF900590FB383FD75ED4" box="[931,999,1189,1215]" italics="true" pageId="1" pageNumber="398">Head.</emphasis>
Head rounded and slightly wider than the pronotum. Vertex broad and flattened, weakly inclined. Frontal rostrum slightly convex in frontal view, equal to the width of antennal scape. Antennal scape transversely broad, about 1.5 times as wide as length. Eyes slightly protruding, not extending beyond the head; ocellus three, lateral ocelli ovoid and distinctly larger than the median ocellus. Postclypeus narrowly banded, and slightly narrower in the middle than laterally; anteclypeus broader, apical narrowing; labrum shaped as a shield. Maxillary palpus five, the third and fourth segments equal in length, and the apical of the fifth segment distinctly widened; labial palpus three, the third segment being as long as the first two and widening at the end.
<emphasis id="B95FEABFFF89FF900420FA3C3E5B5FD0" box="[531,619,1441,1467]" italics="true" pageId="1" pageNumber="398">Thorax.</emphasis>
Pronotum broad, anterior margin slightly narrower than posterior margin and slightly concave.
<emphasis id="B95FEABFFF89FF9007BDFA5B3E405FB4" box="[398,624,1477,1503]" italics="true" pageId="1" pageNumber="398">Tegmina and wings.</emphasis>
Forewings reaching the fifth abdominal segment, oblique veins six, the inner two slightly curved; chord veins three, second and third chord veins basally connected. Diagonal vein straight, proximally curved and connected with oblique veins. Mirror irregularly shaped and surrounded by several large wing cells. The length of cercus equal to half of the abdomen. Both inner and outer tympanum ovoid and equal in size.
<emphasis id="B95FEABFFF89FF9006E3F9CB3D3D5C04" box="[208,269,1622,1647]" italics="true" pageId="1" pageNumber="398">Legs.</emphasis>
Posterior femur well developed, slightly shorter than the abdomen. Posterior tibiae equipped with spines on the one half of proximal portion (inside: 3-4; outside: 5) and spurs on distal part (inside: 4; outside: 4); basitarsus armed with two rows of small spines.
</paragraph>
<paragraph id="8B9436ADFF89FF9006F4F95C39385D2C" blockId="1.[151,1437,1009,2007]" pageId="1" pageNumber="398">
<emphasis id="B95FEABFFF89FF9006F4F95C3D095CB0" box="[199,313,1729,1755]" italics="true" pageId="1" pageNumber="398">Genitalia.</emphasis>
In dorsal view, basal of the epiphallus broader, and the posterior margin of the middle lobe armed with short protuberances. Lateral lobe barrow and length, distinctly enlarged subterminal and curved medially. In lateral view, the lateral lobe of epiphallus slightly curved dorsally, protuberance of middle lobe distinctly shorter than lateral lobe. In ventral view, the ectoparamere broader and about half as long as the lateral lobes.
</paragraph>
<paragraph id="8B9436ADFF89FF9006F4F8CC3F4B5DFB" blockId="1.[151,1437,1009,2007]" pageId="1" pageNumber="398">
<emphasis id="B95FEABFFF89FF9006F4F8CC3D155D00" bold="true" box="[199,293,1873,1899]" pageId="1" pageNumber="398">Female.</emphasis>
Similar to male, but larger than males and with shorter forewings covering only the first abdominal segment. Apical of ovipositor pointed, slightly longer than cercus.
</paragraph>
<paragraph id="8B9436ADFF89FF9006F4F8043E715DBC" blockId="1.[151,1437,1009,2007]" pageId="1" pageNumber="398">
<emphasis id="B95FEABFFF89FF9006F4F8043D7F5DD8" bold="true" box="[199,335,1945,1971]" pageId="1" pageNumber="398">Coloration.</emphasis>
Body dark brown. Head and pronotum black, ocellus white; legs dark reddish brown, and the inner posterior femur and forewings brown.
</paragraph>
<caption id="DF546625FF8AFF9306ABFB5E38065EB1" box="[152,1078,1219,1243]" pageId="2" pageNumber="399" startId="2.[152,255,1219,1243]" targetBox="[231,1354,191,1182]" targetPageId="2" targetType="figure">
<paragraph id="8B9436ADFF8AFF9306ABFB5E38065EB1" blockId="2.[152,1078,1219,1243]" box="[152,1078,1219,1243]" pageId="2" pageNumber="399">
<emphasis id="B95FEABFFF8AFF9306ABFB5E3D295EB0" bold="true" box="[152,281,1219,1243]" pageId="2" pageNumber="399">FIGURE 1.</emphasis>
Bodies of
<taxonomicName id="4C2B4D2EFF8AFF9307B4FB593E635EB0" authorityName="Dai &amp; Wang &amp; Wang &amp; Ma" authorityYear="2025" box="[391,595,1219,1243]" class="Insecta" family="Gryllidae" genus="Duolandrevus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="meiacanthus" status="sp. nov." subGenus="Eulandrevus">
<emphasis id="B95FEABFFF8AFF9307B4FB593D905EB0" box="[391,416,1220,1243]" italics="true" pageId="2" pageNumber="399">D.</emphasis>
(
<emphasis id="B95FEABFFF8AFF93079DFB593DF55EB0" box="[430,453,1220,1243]" italics="true" pageId="2" pageNumber="399">E.</emphasis>
)
<emphasis id="B95FEABFFF8AFF9307E0FB5E3E635EB0" box="[467,595,1219,1243]" italics="true" pageId="2" pageNumber="399">meiacanthus</emphasis>
</taxonomicName>
<emphasis id="B95FEABFFF8AFF93046AFB5E3E9A5EB0" bold="true" box="[601,682,1219,1243]" pageId="2" pageNumber="399">
<taxonomicNameLabel id="A26C57C4FF8AFF93046AFB5E3E9A5EB0" box="[601,682,1219,1243]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. male; B. female; Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="DF546625FF8AFF9306A4F8ED38955DE3" box="[151,1189,1904,1929]" pageId="2" pageNumber="399" startId="2.[151,250,1904,1928]" targetBox="[230,1358,1385,1866]" targetPageId="2" targetType="figure">
<paragraph id="8B9436ADFF8AFF9306A4F8ED38955DE3" blockId="2.[151,1189,1904,1929]" box="[151,1189,1904,1929]" pageId="2" pageNumber="399">
<emphasis id="B95FEABFFF8AFF9306A4F8ED3D235DE3" bold="true" box="[151,275,1904,1928]" pageId="2" pageNumber="399">FIGURE 2.</emphasis>
Genitalia of
<taxonomicName id="4C2B4D2EFF8AFF9307ABF8EC3E545DE3" authorityName="Dai &amp; Wang &amp; Wang &amp; Ma" authorityYear="2025" box="[408,612,1904,1929]" class="Insecta" family="Gryllidae" genus="Duolandrevus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="399" phylum="Arthropoda" rank="species" species="meiacanthus" status="sp. nov." subGenus="Eulandrevus">
<emphasis id="B95FEABFFF8AFF9307ABF8EC3D9A5DE3" box="[408,426,1905,1928]" italics="true" pageId="2" pageNumber="399">D</emphasis>
. (
<emphasis id="B95FEABFFF8AFF93078CF8EC3DE65DE3" box="[447,470,1905,1928]" italics="true" pageId="2" pageNumber="399">E.</emphasis>
)
<emphasis id="B95FEABFFF8AFF9307D7F8ED3E545DE3" box="[484,612,1904,1928]" italics="true" pageId="2" pageNumber="399">meiacanthus</emphasis>
</taxonomicName>
<emphasis id="B95FEABFFF8AFF930459F8EC3E8B5DE2" bold="true" box="[618,699,1905,1929]" pageId="2" pageNumber="399">
<taxonomicNameLabel id="A26C57C4FF8AFF930459F8EC3E8B5DE2" box="[618,699,1905,1929]" pageId="2" pageNumber="399" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. dorsal view; B. lateral view; C. ventral view.
</paragraph>
</caption>
<paragraph id="8B9436ADFF8BFF9206F4FF0A3DE65A92" blockId="3.[151,1437,150,573]" pageId="3" pageNumber="400">
<emphasis id="B95FEABFFF8BFF9206F4FF0A3E685ADB" bold="true" box="[199,600,150,177]" pageId="3" pageNumber="400">Measurements (mm). Male (n=1)</emphasis>
: BL 18.09, HL 3.5, HW 4.6, PL 3.05, PW 4.83, FWL 7.39, HFL 10.86;
<emphasis id="B95FEABFFF8BFF9206A4FF263CC95ABE" bold="true" box="[151,249,187,213]" pageId="3" pageNumber="400">Females</emphasis>
(
<emphasis id="B95FEABFFF8BFF92073BFF273D065ABF" bold="true" box="[264,310,186,212]" pageId="3" pageNumber="400">n=2</emphasis>
). BL 18.04±0.17, HL 3.44±0.08, HW 5.14±0.02, PL 3.24±0.06, PW 5.01±0.16, FWL 2.32±0.24, HFL 12.20±0.08, OL 10.39±0.11.
</paragraph>
</subSubSection>
<subSubSection id="C3316526FF8BFF9206F4FE9E3D745856" pageId="3" pageNumber="400" type="discussion">
<paragraph id="8B9436ADFF8BFF9206F4FE9E3D745856" blockId="3.[151,1437,150,573]" pageId="3" pageNumber="400">
<emphasis id="B95FEABFFF8BFF9206F4FE9E3D0B5B76" bold="true" box="[199,315,259,285]" pageId="3" pageNumber="400">Remarks.</emphasis>
This new species is very similar to
<taxonomicName id="4C2B4D2EFF8BFF9204F9FE9E396B5B76" authority="(Gorochov, 2001)" baseAuthorityName="Gorochov" baseAuthorityYear="2001" box="[714,1371,259,285]" class="Insecta" family="Gryllidae" genus="Duolandrevus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="paradoxus" subGenus="Eulandrevus">
<emphasis id="B95FEABFFF8BFF9204F9FE9E3F585B76" box="[714,872,259,285]" italics="true" pageId="3" pageNumber="400">Duolandrevus</emphasis>
(
<emphasis id="B95FEABFFF8BFF92054BFE9E38345B76" box="[888,1028,259,285]" italics="true" pageId="3" pageNumber="400">Eulandrevus</emphasis>
)
<emphasis id="B95FEABFFF8BFF920226FE9E38BB5B76" box="[1045,1163,259,285]" italics="true" pageId="3" pageNumber="400">paradoxus</emphasis>
(
<bibRefCitation id="EFBA4B5CFF8BFF9202AAFE9E39635B76" author="Gorochov, A. V." box="[1177,1363,259,285]" pageId="3" pageNumber="400" refId="ref1910" refString="Gorochov, A. V. (2001) A new species of Landrevinae from Thailand (Orthoptera: Gryllidae). Zoosystematica Rossica, 10 (1), 36." type="journal volume" year="2001">Gorochov, 2001</bibRefCitation>
)
</taxonomicName>
. Both species have dark brown bodies, the male forewings extending to the 5th abdominal tergite, and long narrow lateral lobes of epiphallus. However, the new species is distinguished from
<taxonomicName id="4C2B4D2EFF8BFF920583FED6384F5B0E" baseAuthorityName="Gorochov" baseAuthorityYear="2001" box="[944,1151,331,357]" class="Insecta" family="Gryllidae" genus="Duolandrevus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="paradoxus" subGenus="Eulandrevus">
<emphasis id="B95FEABFFF8BFF920583FED63FFB5B0F" box="[944,971,331,356]" italics="true" pageId="3" pageNumber="400">D.</emphasis>
(
<emphasis id="B95FEABFFF8BFF9205EDFED63FC65B0F" box="[990,1014,331,356]" italics="true" pageId="3" pageNumber="400">E.</emphasis>
)
<emphasis id="B95FEABFFF8BFF92023AFED6384F5B0E" box="[1033,1151,331,357]" italics="true" pageId="3" pageNumber="400">paradoxus</emphasis>
</taxonomicName>
by the wider and longer basal of the epiphallus. In addition, the lateral lobe of the epiphallus of the new species are significantly longer than those of
<taxonomicName id="4C2B4D2EFF8BFF9206C5FE0E3D8E5BC6" baseAuthorityName="Gorochov" baseAuthorityYear="2001" box="[246,446,403,429]" class="Insecta" family="Gryllidae" genus="Duolandrevus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="paradoxus" subGenus="Eulandrevus">
<emphasis id="B95FEABFFF8BFF9206C5FE0E3D215BC7" box="[246,273,403,428]" italics="true" pageId="3" pageNumber="400">D.</emphasis>
(
<emphasis id="B95FEABFFF8BFF920713FE0E3D095BC7" box="[288,313,403,428]" italics="true" pageId="3" pageNumber="400">E.</emphasis>
)
<emphasis id="B95FEABFFF8BFF92077BFE0E3D8E5BC6" box="[328,446,403,429]" italics="true" pageId="3" pageNumber="400">paradoxus</emphasis>
</taxonomicName>
. Finally, the length of the posterior marginal protuberance of the middle lobe is markedly different in the two species of epiphallus. The protuberance of the new species is about 1/5 as long as the lateral lobe, whereas the posterior marginal protuberance of
<taxonomicName id="4C2B4D2EFF8BFF9204D9FE463F835B9E" baseAuthorityName="Gorochov" baseAuthorityYear="2001" box="[746,947,475,501]" class="Insecta" family="Gryllidae" genus="Duolandrevus" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="400" phylum="Arthropoda" rank="species" species="paradoxus" subGenus="Eulandrevus">
<emphasis id="B95FEABFFF8BFF9204D9FE463F355B9F" box="[746,773,475,500]" italics="true" pageId="3" pageNumber="400">D.</emphasis>
(
<emphasis id="B95FEABFFF8BFF920526FE463F1D5B9F" box="[789,813,475,500]" italics="true" pageId="3" pageNumber="400">E.</emphasis>
)
<emphasis id="B95FEABFFF8BFF92050EFE463F835B9E" box="[829,947,475,501]" italics="true" pageId="3" pageNumber="400">paradoxus</emphasis>
</taxonomicName>
extends to the expansion of the lateral lobe, exceeding the length of the lateral lobe by half. Therefore, we consider this species should be treated as a species new to science.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="0730C1AA9F80B4B473FB7EB320D06A72">A new species of Microhyla (Amphibia: Anura: Microhylidae), with an extended description of Microhyla nanapollexa from Vietnam</mods:title>
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<emphasis id="B9542E38FFFCAE5AFF27E33D7BE68BCF" bold="true" box="[151,434,1390,1417]" italics="true" pageId="5" pageNumber="352">Microhyla nanapollexa</emphasis>
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(
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,
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;
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)
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<emphasis id="B9542E38FFFCAE5AFF27E3867BCD8BA9" bold="true" box="[151,409,1493,1519]" pageId="5" pageNumber="352">Specimens examined.</emphasis>
(n = 2),
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male
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female
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A.6270 (Field number
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), collected by C.
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<emphasis id="B9542E38FFFCAE5AFCF2E3A9792E8855" box="[834,890,1529,1555]" italics="true" pageId="5" pageNumber="352">et al.</emphasis>
in
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April
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in the forest of
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,
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,
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(
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, at an elevation of
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a.s.l.
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.
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<emphasis id="B9542E38FFFCAE5AFF77E012782E881D" bold="true" box="[199,634,1601,1627]" pageId="5" pageNumber="352">Description of specimens examined.</emphasis>
Morphological characters of the specimens from
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,
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, match the description of
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: Size small (SVL
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in male;
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in female); habitus slender, head longer than wide (HL/HW
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male,
<quantity id="4CD85FCFFFFCAE5AFCDEE0DA79EA88E5" box="[878,958,1673,1700]" metricMagnitude="-2" metricUnit="m" metricValue="2.921" pageId="5" pageNumber="352" unit="in" value="1.15">1.15 in</quantity>
female); snout round, slightly protruding beyond margin of lower jaw, longer than diameter of eye (SL/ED
<quantity id="4CD85FCFFFFCAE5AFC32E0FE79858881" box="[898,977,1709,1736]" metricMagnitude="-2" metricUnit="m" metricValue="3.9624" pageId="5" pageNumber="352" unit="in" value="1.56">1.56 in</quantity>
male;
<quantity id="4CD85FCFFFFCAE5AFBAFE0FE7E3A8881" box="[1055,1134,1709,1736]" metricMagnitude="-2" metricUnit="m" metricValue="4.191" pageId="5" pageNumber="352" unit="in" value="1.65">1.65 in</quantity>
female); eyes protuberant, pupil round; dorsal surface of head flat; canthus rostralis round, distinct; loreal region oblique, flat; nostril round, lateral, on canthus rostralis, closer to tip of snout than to eye (NED/SL
<quantity id="4CD85FCFFFFCAE5AFC75E0A67E418949" box="[965,1045,1781,1807]" metricMagnitude="-2" metricUnit="m" metricValue="1.524" pageId="5" pageNumber="352" unit="in" value="0.6">0.60 in</quantity>
male,
<quantity id="4CD85FCFFFFCAE5AFBD3E0A67EE68949" box="[1123,1202,1781,1808]" metricMagnitude="-2" metricUnit="m" metricValue="1.4224" pageId="5" pageNumber="352" unit="in" value="0.56">0.56 in</quantity>
female); interorbital distance wider than internarial distance (IOD/IND
<quantity id="4CD85FCFFFFCAE5AFD61E14A794B8975" box="[721,799,1817,1843]" metricMagnitude="-2" metricUnit="m" metricValue="4.0894" pageId="5" pageNumber="352" unit="in" value="1.61">1.61 in</quantity>
male,
<quantity id="4CD85FCFFFFCAE5AFCDBE14A79ED8975" box="[875,953,1817,1843]" metricMagnitude="-2" metricUnit="m" metricValue="4.1148" pageId="5" pageNumber="352" unit="in" value="1.62">1.62 in</quantity>
female) and wider than upper eyelid width (IOD/UEW
<quantity id="4CD85FCFFFFCAE5AFE91E16E7B3B8911" box="[289,367,1853,1880]" metricMagnitude="-2" metricUnit="m" metricValue="4.699" pageId="5" pageNumber="352" unit="in" value="1.85">1.85 in</quantity>
male and female); tympanum hidden; vomerine teeth absent, tongue round posteriorly, free for approximately two thirds its length; slit like openings to a median vocal sac.
</paragraph>
<paragraph id="8B9FF22AFFFCAE59FF77E1D578C38F24" blockId="5.[151,1437,1493,2021]" lastBlockId="6.[151,1437,152,354]" lastPageId="6" lastPageNumber="353" pageId="5" pageNumber="352">
Forelimbs relatively short, shorter than snout vent length (FLL/SVL
<quantity id="4CD85FCFFFFCAE5AFC73E1D47E4489D8" box="[963,1040,1926,1951]" metricMagnitude="-2" metricUnit="m" metricValue="1.8796" pageId="5" pageNumber="352" unit="in" value="0.74">0.74 in</quantity>
male,
<quantity id="4CD85FCFFFFCAE5AFBEAE1D47EFC89D8" box="[1114,1192,1926,1951]" metricMagnitude="-2" metricUnit="m" metricValue="1.9812" pageId="5" pageNumber="352" unit="in" value="0.78">0.78 in</quantity>
female); hand shorter than half of forelimb length (HAL/FLL
<quantity id="4CD85FCFFFFCAE5AFDFBE1F978C38984" box="[587,663,1962,1986]" metricMagnitude="-2" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="352" unit="in" value="0.4">0.40 in</quantity>
male,
<quantity id="4CD85FCFFFFCAE5AFD6FE1F9797F8984" box="[735,811,1962,1986]" metricMagnitude="-3" metricUnit="m" metricValue="9.652" pageId="5" pageNumber="352" unit="in" value="0.38">0.38 in</quantity>
female); fingers slender, free of webbing, dorsoventrally flattened, fingers with weak skin fringes on finger II and III; nuptial pad absent on male finger I; partially free nub just proximal to finger II (1FLO/2FLO
<quantity id="4CD85FCFFFFFAE59FDE1E6CB78CB8EF6" box="[593,671,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="353" unit="in" value="0.2">0.20 in</quantity>
male,
<quantity id="4CD85FCFFFFFAE59FD5BE6CB796D8EF6" box="[747,825,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="5.334" pageId="6" pageNumber="353" unit="in" value="0.21">0.21 in</quantity>
female); finger II slightly longer than finger IV length (2FLI/4FLI
<quantity id="4CD85FCFFFFFAE59FE91E6EF7B248E92" box="[289,368,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.6924" pageId="6" pageNumber="353" unit="in" value="1.06">1.06 in</quantity>
male,
<quantity id="4CD85FCFFFFFAE59FE0FE6EF785B8E92" box="[447,527,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.6416" pageId="6" pageNumber="353" unit="in" value="1.04">1.04 in</quantity>
female), and much shorter than the third (2FLI/3FLI
<quantity id="4CD85FCFFFFFAE59FBC6E6EF7E928E92" box="[1142,1222,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="353" unit="in" value="0.6">0.60 in</quantity>
male and female); relative finger lengths: I&lt;II≤IV&lt;III; tip of finger I round, not enlarged; narrow peripheral grooves present on fingers II, III and IV; grooves on dorsal surface of disks present in fingers II, III, IV, absent in finger I; relative finger disk widths II&lt;IV&lt;III; subarticular tubercles on fingers distinct, round, formula 1:1:2:1 (given for fingers I:II:III:IV, respectively); two round, flat palmar tubercle.
</paragraph>
<caption id="DF5FA2A2FFFFAE59FF27E5077AB68DD6" pageId="6" pageNumber="353" startId="6.[151,250,852,876]" targetBox="[151,1435,378,828]" targetPageId="6" targetType="figure">
<paragraph id="8B9FF22AFFFFAE59FF27E5077AB68DD6" blockId="6.[151,1437,852,912]" pageId="6" pageNumber="353">
<emphasis id="B9542E38FFFFAE59FF27E5077B478D2A" bold="true" box="[151,275,852,876]" pageId="6" pageNumber="353">FIGURE 2.</emphasis>
<taxonomicName id="4C2089A9FFFFAE59FEAAE50778538D2A" authorityName="Bain &amp; Nguyen" authorityYear="2004" box="[282,519,852,876]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="353" phylum="Chordata" rank="species" species="nanapollexa">
<emphasis id="B9542E38FFFFAE59FEAAE50778538D2A" box="[282,519,852,876]" italics="true" pageId="6" pageNumber="353">Microhyla nanapollexa</emphasis>
</taxonomicName>
dorsolateral view (A) and ventral view (B) in life (IEBR A.6270, female). Photos by C.V. Hoang.
</paragraph>
</caption>
<paragraph id="8B9FF22AFFFFAE59FF77E58E7B948AA8" blockId="6.[151,1437,989,2017]" pageId="6" pageNumber="353">
Hindlimbs slender, twice as long as snout-vent length (HLL/SVL
<quantity id="4CD85FCFFFFFAE59FC0BE58D7E5F8DB3" box="[955,1035,989,1014]" metricMagnitude="-2" metricUnit="m" metricValue="4.8768" pageId="6" pageNumber="353" unit="in" value="1.92">1.92 in</quantity>
male,
<quantity id="4CD85FCFFFFFAE59FBEBE58D7EFF8DB3" box="[1115,1195,989,1014]" metricMagnitude="-2" metricUnit="m" metricValue="5.2578" pageId="6" pageNumber="353" unit="in" value="2.07">2.07 in</quantity>
female), tibia longer than half of snout-vent length (TL/SVL
<quantity id="4CD85FCFFFFFAE59FDE5E25278F68A5F" box="[597,674,1025,1049]" metricMagnitude="-2" metricUnit="m" metricValue="1.6002" pageId="6" pageNumber="353" unit="in" value="0.63">0.63 in</quantity>
male,
<quantity id="4CD85FCFFFFFAE59FD5DE252796E8A5F" box="[749,826,1025,1049]" metricMagnitude="-2" metricUnit="m" metricValue="1.7272" pageId="6" pageNumber="353" unit="in" value="0.68">0.68 in</quantity>
female); tibiotarsal articulation reaching beyond snout when limb adpressed along body; foot longer than tibia (FL/TL
<quantity id="4CD85FCFFFFFAE59FCD9E27779E38A7A" box="[873,951,1060,1084]" metricMagnitude="-2" metricUnit="m" metricValue="3.6322" pageId="6" pageNumber="353" unit="in" value="1.43">1.43 in</quantity>
male,
<quantity id="4CD85FCFFFFFAE59FBB4E2777E068A7A" box="[1028,1106,1060,1084]" metricMagnitude="-2" metricUnit="m" metricValue="3.5052" pageId="6" pageNumber="353" unit="in" value="1.38">1.38 in</quantity>
female); relative toe lengths: I&lt;II&lt;V&lt;III&lt;IV; tarsus smooth, inner tarsal fold absent; tips of all toes slightly dilated, forming truncated disks, disks of all toes with peripheral grooves, dorsal surface of toe disks IIV with short median longitudinal grooves; relative toe disk widths I&lt;V&lt;II&lt;III=IV; webbing formula I12II12½III2½2½IV2½1V; subarticular tubercles on toes distinct, round, formula 1:1:2:3:2 (for toes I:II:III:IV:V, respectively); inner metatarsal tubercle oval; outer metatarsal tubercle absent.
</paragraph>
<paragraph id="8B9FF22AFFFFAE59FF77E2AA7B738B3A" blockId="6.[151,1437,989,2017]" pageId="6" pageNumber="353">Skin on dorsum relatively smooth with low, round tubercles concentrated on the outer edge, flanks, below and posterior to eye; dorsolateral folds distinct in adult male; cloaca unmodified, directed posteriorly, at upper level of thighs; supratympanic fold indistinct; lateral side of head smooth; ventral surface of body and limbs smooth, vent area smooth.</paragraph>
<paragraph id="8B9FF22AFFFFAE59FF77E3D57AE188DA" blockId="6.[151,1437,989,2017]" pageId="6" pageNumber="353">
Coloration in life. Dorsal surface greyish brown to light brown with a dark brown or brown spear-shape marking, running from the posterior eye to the groin; black spear-shape marking border, or just two black spots on either side of the spear-shape marking; a black broken streak below dorsolateral folds; black streak-shape marking present on the groin region in adult male; a light brown streak from posterior corner of eye to arm insertion; dorsolateral surfaces of arm and fingers dark brown with a small black bar near the wrist; dorsal surfaces of thigh, tibia, tarsus and toes brown with dark brown cross bars on each hindlimb; throat and chest dark grey with intense small white mottling, belly and limbs ventrally lighter with white marbling mixed with black spots; pupil black, iris copper (
<figureCitation id="131BEEAFFFFFAE59FADFE00D7AFC88DA" captionStart="FIGURE 2" captionStartId="6.[151,250,852,876]" captionTargetBox="[151,1435,378,828]" captionTargetId="figure-139@6.[151,1435,378,828]" captionTargetPageId="6" captionText="FIGURE 2. Microhyla nanapollexa dorsolateral view (A) and ventral view (B) in life (IEBR A.6270, female). Photos by C.V. Hoang." pageId="6" pageNumber="353">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B9FF22AFFFFAE59FF77E0F57E9A894E" blockId="6.[151,1437,989,2017]" pageId="6" pageNumber="353">
Coloration in preservative. In ethanol, dorsal coloration changed to dark greyish brown, ventral surface of chest, belly, and limbs greyish beige; dorsal pattern, dark spots on dorsum and stripes on dorsal surface of limbs unchanged, dark brown pattern changed to dark grey; pupil white, iris completely black (
<figureCitation id="131BEEAFFFFFAE59FBCCE0BD7E94894E" box="[1148,1216,1774,1800]" captionStart="FIGURE 3" captionStartId="7.[151,250,1810,1834]" captionTargetBox="[156,1431,530,1782]" captionTargetId="figure-143@7.[151,1435,525,1786]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal view (A) and ventral view (B) of the female specimen (IEBR A.6270) of Microhyla nanapollexa in preservative. Scale bar = 7 mm. Photos by C.V. Hoang." pageId="6" pageNumber="353">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B9FF22AFFFFAE58FF77E1417B0B8F23" blockId="6.[151,1437,989,2017]" lastBlockId="7.[151,1436,151,429]" lastPageId="7" lastPageNumber="354" pageId="6" pageNumber="353">
<emphasis id="B9542E38FFFFAE59FF77E1417BE1896A" bold="true" box="[199,437,1810,1836]" pageId="6" pageNumber="353">Extended diagnosis.</emphasis>
<taxonomicName id="4C2089A9FFFFAE59FE0BE14178EB896A" authorityName="Bain &amp; Nguyen" authorityYear="2004" box="[443,703,1810,1836]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="353" phylum="Chordata" rank="species" species="nanapollexa">
<emphasis id="B9542E38FFFFAE59FE0BE14178EB896A" box="[443,703,1810,1836]" italics="true" pageId="6" pageNumber="353">Microhyla nanapollexa</emphasis>
</taxonomicName>
<materialsCitation id="3B48F877FFFFAE58FD76E1417B0F8F23" collectionCode="SVL, II, III, IV, V" lastPageId="7" lastPageNumber="354" pageId="6" pageNumber="353" specimenCode="II1-2, III2, IV 2" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
is characterized by having a small body size (
<collectionCode id="ED316AEFFFFFAE59FB0EE1417EA0896A" box="[1214,1268,1810,1836]" pageId="6" pageNumber="353">SVL</collectionCode>
<quantity id="4CD85FCFFFFFAE59FB4AE1417F39896B" box="[1274,1389,1810,1837]" metricMagnitude="-2" metricUnit="m" metricValue="1.543" pageId="6" pageNumber="353" unit="mm" value="15.43">15.43 mm</quantity>
, n =
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;
<quantity id="4CD85FCFFFFFAE59FF42E1657BE78917" box="[242,435,1846,1873]" metricMagnitude="-2" metricUnit="m" metricValue="1.6324999999999998" metricValueMax="1.6629999999999998" metricValueMin="1.6019999999999999" pageId="6" pageNumber="353" unit="mm" value="16.325" valueMax="16.63" valueMin="16.02">16.0216.63 mm</quantity>
, n=
<specimenCount id="9D2639A3FFFFAE59FE6DE16578198916" box="[477,589,1846,1872]" count="2" pageId="6" pageNumber="353" type="female">2 females</specimenCount>
); body habitus slender; snout round; skin on dorsum relatively smooth with low, round tubercles concentrated on the outer edge, flanks, below and posterior to eye; venter smooth; vertebral skin ridge and dorsomedial stripe absent; superciliary tubercles absent; supratympanic fold indistinct; fingers slender, free of webbing, with weak dermal fringes on fingers
<collectionCode id="ED316AEFFFFFAE59FCC7E1F079DE89FB" box="[887,906,1955,1981]" pageId="6" pageNumber="353">II</collectionCode>
and
<collectionCode id="ED316AEFFFFFAE59FC78E1F079B389FB" box="[968,999,1955,1981]" pageId="6" pageNumber="353">III</collectionCode>
; nuptial pad absent on male finger I; partially free nub just proximal to finger
<collectionCode id="ED316AEFFFFFAE59FDD4E194782E89A7" box="[612,634,1991,2017]" pageId="6" pageNumber="353">II</collectionCode>
; grooves on dorsal surface of disks present in fingers
<collectionCode id="ED316AEFFFFFAE59FB67E1947EBE89A7" box="[1239,1258,1991,2017]" pageId="6" pageNumber="353">II</collectionCode>
<collectionCode id="ED316AEFFFFFAE59FB49E1947F4389A7" box="[1273,1303,1991,2017]" pageId="6" pageNumber="353">IV</collectionCode>
; two round, flat palmar tubercles; hindlimbs long, tibiotarsal articulation of adpressed limb reaching well beyond snout; toe webbing formula I12
<specimenCode id="DB865A51FFFEAE58FE3DE6EF7B938E91" box="[397,455,188,215]" collectionCode="II" pageId="7" pageNumber="354">II12</specimenCode>
½
<specimenCode id="DB865A51FFFEAE58FE6DE6EF785C8E91" box="[477,520,188,215]" collectionCode="III" pageId="7" pageNumber="354">III2</specimenCode>
½
<specimenCode id="DB865A51FFFEAE58FDE1E6EF78D18E91" box="[593,645,188,215]" collectionCode="IV" pageId="7" pageNumber="354">IV 2</specimenCode>
½1
<collectionCode id="ED316AEFFFFEAE58FD06E6EF78988E91" box="[694,716,188,215]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="354" type="Museum">V</collectionCode>
; inner metatarsal tubercle oval; outer metatarsal tubercle absent; dorsal surface greyish brown to light brown with a dark brown or brown spear shape marking, running from the posterior eye to the groin; a black broken streak below dorsolateral folds; black streak shape marking absent or present on the groin region; belly and limbs ventrally lighter with white marble mixed with black spots; pupil black, iris copper in life
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33AA1A1FFFEAE58FF77E73D7F068FEB" pageId="7" pageNumber="354" type="distribution">
<paragraph id="8B9FF22AFFFEAE58FF77E73D7F068FEB" blockId="7.[151,1436,151,429]" pageId="7" pageNumber="354">
<emphasis id="B9542E38FFFEAE58FF77E73D7B358FCE" bold="true" box="[199,353,366,392]" pageId="7" pageNumber="354">Distribution.</emphasis>
<taxonomicName id="4C2089A9FFFEAE58FED7E73C783F8FCF" authorityName="Bain &amp; Nguyen" authorityYear="2004" box="[359,619,367,393]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="354" phylum="Chordata" rank="species" species="nanapollexa">
<emphasis id="B9542E38FFFEAE58FED7E73C783F8FCF" box="[359,619,367,393]" italics="true" pageId="7" pageNumber="354">Microhyla nanapollexa</emphasis>
</taxonomicName>
is currently known from the
<typeStatus id="549B4C88FFFEAE58FC1EE73C798A8FCF" box="[942,990,367,393]" pageId="7" pageNumber="354">type</typeStatus>
locality in Tra My District,
<collectingRegion id="49E43CC8FFFEAE58FAA5E73C7AAF8FEB" country="Vietnam" name="Quang Nam" pageId="7" pageNumber="354">Quang Nam Province</collectingRegion>
and Kon Plong District,
<collectingRegion id="49E43CC8FFFEAE58FDA3E7C078BE8FEB" box="[531,746,403,429]" country="Vietnam" name="Kon Tum" pageId="7" pageNumber="354">Kon Tum Province</collectingRegion>
,
<collectingCountry id="F337B2BAFFFEAE58FD45E7C079008FEB" box="[757,852,403,429]" name="Vietnam" pageId="7" pageNumber="354">Vietnam</collectingCountry>
at elevations between 1260 and
<quantity id="4CD85FCFFFFEAE58FB72E7C07F438FEB" box="[1218,1303,403,429]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="7" pageNumber="354" unit="m" value="1480.0">1480 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,161 @@
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<mods:title id="EAE609044917B126DF97BADC1FE6C526">Taxonomic redescription and molecular confirmation of Lutjanus rufolineatus (Acanthuriformes: Lutjanidae) from the Andaman Islands, India</mods:title>
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<mods:name id="276BA8B5F9ABABB2FAB870D4FEE5B2FF" type="personal">
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<mods:namePart id="23B3352730E15E25746887B9055DD5F6">Ps, Fahmeeda Parveen</mods:namePart>
<mods:affiliation id="1D3319CD42A8191AC6EA758B5A0985A4">Department of Ocean Studies and Marine Biology, Pondicherry University, Brookshabad Campus, Port Blair- 744112, Andaman &amp; Nicobar Islands, India.</mods:affiliation>
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<mods:namePart id="486F5A1A66AF91934A0066552F94FCE1">Venu, Sasidharan</mods:namePart>
<mods:affiliation id="BC86F8DFE2571D6F7D06781B016B1326">Department of Ocean Studies and Marine Biology, Pondicherry University, Brookshabad Campus, Port Blair- 744112, Andaman &amp; Nicobar Islands, India. &amp; Department of Marine Biology Microbiology and Biochemistry, School of Marine Sciences, Lakeside campus, Cochin University of Science and Technology, Fine Arts Avenue, Kochi- 682016, Ernakulam, Kerala.</mods:affiliation>
</mods:name>
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<mods:namePart id="0F59DD30C42FECE5B07F4147F0AC055A">Eranhottu, Shibin</mods:namePart>
<mods:affiliation id="7EB3EF0385990BC1B25ECD697DF2207A">Department of Ocean Studies and Marine Biology, Pondicherry University, Brookshabad Campus, Port Blair- 744112, Andaman &amp; Nicobar Islands, India.</mods:affiliation>
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<mods:name id="97AC9537B909DCBC52E875EEA679D254" type="personal">
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<figureCitation id="132F2A3B310DFF82FF25BFD8D0D525C9" box="[159,246,1231,1257]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Figs 10</figureCitation>
,
<figureCitation id="132F2A3B310DFF82FEB8BFD8D13D25C9" box="[258,286,1231,1257]" captionStart="FIGURE 13" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[169,1415,195,1904]" captionTargetId="figure-33@22.[157,1430,181,1919]" captionTargetPageId="22" captionText="FIGURE 13. Distributions of species of the Homoplectra japonica Species Group in Japan." pageId="19" pageNumber="322">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E6535310DFF82FF2DBE04D16724FB" pageId="19" pageNumber="322" type="reference_group">
<paragraph id="8BAB36BE310DFF82FF2DBE04D1AC2469" blockId="19.[151,1437,1299,1354]" pageId="19" pageNumber="322">
<treatmentCitationGroup id="AB041190310DFF82FF2DBE04D1AC2469" pageId="19" pageNumber="322">
<taxonomicName id="4C144D3D310DFF82FF2DBE04D156240B" ID-CoL="36JDB" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[151,373,1299,1323]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="322" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC310DFF82FF2DBE04D156240B" box="[151,373,1299,1323]" italics="true" pageId="19" pageNumber="322">Diplectrona japonica</emphasis>
</taxonomicName>
nec
<bibRefCitation id="EF854B4F310DFF82FE0ABE04D212240B" author="Banks, N." box="[432,561,1299,1323]" pageId="19" pageNumber="322" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
:
<treatmentCitation id="0AB510AF310DFF82FDFABE03D3BB240B" author="Inazu, K. &amp; Nishida, A." box="[576,920,1299,1323]" page="191" pageId="19" pageNumber="322" year="2011">
<bibRefCitation id="EF854B4F310DFF82FDFABE03D360240B" author="Inazu, K. &amp; Nishida, A." box="[576,835,1299,1323]" pageId="19" pageNumber="322" pagination="169 - 212" refId="ref17188" refString="Inazu, K. &amp; Nishida, A. (2011) Trichoptera. In: Editorial office of Hyogo Fresh Water Biology (Ed.), Hyogo no kawa no ikimono zukan, Hyogo Rikusui-seibutsu Kenkyu-kai, Himeji, Hyogo, pp. 169 - 212 [in Japanese]." type="book chapter" year="2011">Inazu and Nishida 2011</bibRefCitation>
, p. 191
</treatmentCitation>
(in part);
<treatmentCitation id="0AB510AF310DFF82FBB4BE03D4E6240B" author="Inazu, K." box="[1038,1221,1299,1323]" page="8" pageId="19" pageNumber="322" year="2012">
<bibRefCitation id="EF854B4F310DFF82FBB4BE03D4A4240B" author="Inazu, K." box="[1038,1159,1299,1323]" pageId="19" pageNumber="322" pagination="1 - 14" refId="ref17125" refString="Inazu, K. (2012) Collection record of caddisflies in the Kishida River, Hyogo Prefecture. Hyogo Freshwater Biology, 63, 1 - 14 [in Japanese]." type="journal article" year="2012">Inazu 2012</bibRefCitation>
, p. 8
</treatmentCitation>
;
<treatmentCitation id="0AB510AF310DFF82FB6FBE03D5BA240B" author="Inazu, K." box="[1237,1433,1299,1323]" page="72" pageId="19" pageNumber="322" year="2013">
<bibRefCitation id="EF854B4F310DFF82FB6FBE03D56D240B" author="Inazu, K." box="[1237,1358,1299,1323]" pageId="19" pageNumber="322" pagination="65 - 78" refId="ref17158" refString="Inazu, K. (2013) Collection records of caddisflies in the Yada River. Hyogo Freshwater Biology, 64, 65 - 78 [in Japanese]." type="journal article" year="2013">Inazu 2013</bibRefCitation>
, p. 72
</treatmentCitation>
.
<emphasis id="B960EAAC310DFF82FF7DBE25D1AC2469" box="[199,399,1330,1354]" inLineHeading="true" pageId="19" pageNumber="322" reason="1">
<emphasis id="B960EAAC310DFF82FF7DBE25D1AA246A" bold="true" box="[199,393,1330,1354]" pageId="19" pageNumber="322">Misidentifications</emphasis>
.
</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAB36BE310DFF82FF2DBE6FD16724FB" blockId="19.[151,1437,1400,1499]" pageId="19" pageNumber="322">
<emphasis id="B960EAAC310DFF82FF2DBE6FD10324B2" bold="true" box="[151,288,1400,1426]" pageId="19" pageNumber="322">Diagonosis.</emphasis>
This species is similar to
<taxonomicName id="4C144D3D310DFF82FDECBE6ED33A24B2" authorityName="Nozaki" authorityYear="2021" baseAuthorityName="Ulmer" baseAuthorityYear="1907" box="[598,793,1400,1426]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="322" phylum="Arthropoda" rank="species" species="albomargnata">
<emphasis id="B960EAAC310DFF82FDECBE6ED33A24B2" box="[598,793,1400,1426]" italics="true" pageId="19" pageNumber="322">H. albomargnata</emphasis>
</taxonomicName>
in having a long scape of each antenna, but both male and female are distinguishable from those of the latter by the character states given in the diagnosis of
<taxonomicName id="4C144D3D310DFF82FA3BBE8AD11F24FA" baseAuthorityName="Ulmer" baseAuthorityYear="1907" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="322" phylum="Arthropoda" rank="species" species="albomarginata">
<emphasis id="B960EAAC310DFF82FA3BBE8AD11F24FA" italics="true" pageId="19" pageNumber="322">H. albomarginata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6535310DFF86FF2DBD3BD23020F1" lastPageId="23" lastPageNumber="326" pageId="19" pageNumber="322" type="description">
<paragraph id="8BAB36BE310DFF82FF2DBD3BD10B2766" blockId="19.[151,296,1580,1606]" box="[151,296,1580,1606]" pageId="19" pageNumber="322">
<emphasis id="B960EAAC310DFF82FF2DBD3BD10B2766" bold="true" box="[151,296,1580,1606]" pageId="19" pageNumber="322">Description</emphasis>
</paragraph>
<paragraph id="8BAB36BE310DFF82FF2DBD63D19B27DA" blockId="19.[151,1436,1652,2039]" pageId="19" pageNumber="322">
<emphasis id="B960EAAC310DFF82FF2DBD63D0C027AE" bold="true" box="[151,227,1652,1678]" pageId="19" pageNumber="322">Adult.</emphasis>
General morphology and coloration similar to those of
<taxonomicName id="4C144D3D310DFF82FCDEBD62D40E27AE" baseAuthorityName="Ulmer" baseAuthorityYear="1907" box="[868,1069,1652,1678]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="322" phylum="Arthropoda" rank="species" species="albomarginata">
<emphasis id="B960EAAC310DFF82FCDEBD62D40E27AE" box="[868,1069,1652,1678]" italics="true" pageId="19" pageNumber="322">H. albomarginata</emphasis>
</taxonomicName>
including having long scape on each antenna, each scape 34 times as long as its pedicel, approximately same length as basal segment of flagellum, with long setae dorsally. Forewings each 9.0
<quantity id="4CEC9B5B310DFF82FD2CBDABD2C827F7" box="[662,747,1724,1751]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="19" pageNumber="322" unit="mm" value="9.8">9.8 mm</quantity>
long in male (n = 10), 10.012.0 mm long in female (n = 10),
<quantity id="4CEC9B5B310DFF82FF2DBDF7D0CE27DB" box="[151,237,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="19" pageNumber="322" unit="mm" value="9.3">9.3 mm</quantity>
in male
<typeStatus id="54AF881C310DFF82FEF4BDF7D19727DA" box="[334,436,1760,1786]" pageId="19" pageNumber="322" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BAB36BE310DFF85FF7DBC13D5152061" blockId="19.[151,1436,1652,2039]" lastBlockId="20.[151,1436,151,321]" lastPageId="20" lastPageNumber="323" pageId="19" pageNumber="322">
<emphasis id="B960EAAC310DFF82FF7DBC13D152263E" bold="true" box="[199,369,1796,1822]" pageId="19" pageNumber="322">Male genitalia</emphasis>
(
<figureCitation id="132F2A3B310DFF82FEC4BC13D207263E" box="[382,548,1796,1822]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Figs 10A10F</figureCitation>
). Ventrolateral sides of synsclerotized segment IX triangular anteriorly in lateral aspect (
<figureCitation id="132F2A3B310DFF82FF54BC3FD17B2662" box="[238,344,1832,1858]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Fig. 10A</figureCitation>
); posteroventral lobe almost square with median apical concavity and acute apicolateral corners in ventral aspect (
<figureCitation id="132F2A3B310DFF82FEE5BC5BD1E42646" box="[351,455,1868,1894]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Fig. 10C</figureCitation>
); dorsal part of segment IX trapezoidal in dorsal aspect (
<figureCitation id="132F2A3B310DFF82FBFFBC5BD48F2646" box="[1093,1196,1868,1894]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Fig. 10B</figureCitation>
); fused with segment X laterally (
<figureCitation id="132F2A3B310DFF82FEA4BC67D1A826AA" box="[286,395,1904,1930]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Figs 10A</figureCitation>
). Segment X very wide in dorsal aspect, with pair of fin-like dorsolateral projections, pair of posterolateral setose areas protruding posterad (
<figureCitation id="132F2A3B310DFF82FD1FBC83D368268E" box="[677,843,1940,1966]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Figs 10A, 10B</figureCitation>
); pair of posterior processes lacking. Genital chamber with or without pair of short slender inner processes (
<figureCitation id="132F2A3B310DFF82FCB8BCAFD34F26F2" box="[770,876,1976,2002]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Fig. 10A</figureCitation>
), processes rarely long (
<figureCitation id="132F2A3B310DFF82FB3EBCAFD4C926F2" box="[1156,1258,1976,2002]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Fig. 10F</figureCitation>
); pair of lateral strips weakly sclerotized (
<figureCitation id="132F2A3B310DFF82FE00BCCBD20126D6" box="[442,546,2012,2038]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="19" pageNumber="322">Fig. 10A</figureCitation>
). Inferior appendages without distal segment, long, finger-like and very slightly upturned in lateral aspect (
<figureCitation id="132F2A3B310AFF85FE7ABB80D20B2191" box="[448,552,151,177]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="20" pageNumber="323">Fig. 10A</figureCitation>
), extending beyond apex of segment X, each with apex round, with many short spine-like setae subapicomesally (
<figureCitation id="132F2A3B310AFF85FDAFBBACD25E21F5" box="[533,637,187,213]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="20" pageNumber="323">Figs 10A</figureCitation>
inset, 10C). Phallotheca strongly bent posteroventrad, with long dorsal process acute apically (
<figureCitation id="132F2A3B310AFF85FE26BBC8D26721D9" box="[412,580,223,249]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="20" pageNumber="323">Figs 10D, 10E</figureCitation>
); pair of ventral processes 2/3 to 3/4 length as dorsal process, acute apically, slightly thicker subapically (
<figureCitation id="132F2A3B310AFF85FE6DBA14D2A3203D" box="[471,640,259,285]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="20" pageNumber="323">Figs 10D, 10E</figureCitation>
). Aedeagus evenly curved ventrad, but apex curved dorsad (
<figureCitation id="132F2A3B310AFF85FA9CBA14D5AC203D" box="[1318,1423,259,285]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="20" pageNumber="323">Fig. 10D</figureCitation>
); head short, less than 1/10 as long as stem; stem half-pipe-like, with broad lateral flanges (
<figureCitation id="132F2A3B310AFF85FB38BA30D5092061" box="[1154,1322,295,321]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="20" pageNumber="323">Figs 10D, 10E</figureCitation>
).
</paragraph>
<caption id="DF6B6636310AFF85FF2DBDD1D17C266A" pageId="20" pageNumber="323" startId="20.[151,250,1734,1758]" targetBox="[157,1429,351,1705]" targetPageId="20" targetType="figure">
<paragraph id="8BAB36BE310AFF85FF2DBDD1D17C266A" blockId="20.[151,1436,1734,1866]" pageId="20" pageNumber="323">
<emphasis id="B960EAAC310AFF85FF2DBDD1D2EA27FE" bold="true" box="[151,713,1734,1758]" pageId="20" pageNumber="323">
FIGURE 10. Adults of
<taxonomicName id="4C144D3D310AFF85FE20BDD1D24E27FE" authorityName="Nozaki" authorityYear="2025" box="[410,621,1734,1758]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="20" pageNumber="323" phylum="Arthropoda" rank="species" species="inazui" status="sp. nov.">
<emphasis id="B960EAAC310AFF85FE20BDD1D24E27FE" bold="true" box="[410,621,1734,1758]" italics="true" pageId="20" pageNumber="323">Homoplectra inazui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357D7310AFF85FDCCBDD1D2EA27FE" box="[630,713,1734,1758]" pageId="20" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral.
</paragraph>
</caption>
<caption id="DF6B6636310BFF84FF2DBD69D201264D" pageId="21" pageNumber="324" startId="21.[151,264,1662,1686]" targetBox="[157,1429,187,1632]" targetPageId="21" targetType="figure">
<paragraph id="8BAB36BE310BFF84FF2DBD69D201264D" blockId="21.[151,1437,1662,1902]" pageId="21" pageNumber="324">
<emphasis id="B960EAAC310BFF84FF2DBD69D4C427B6" bold="true" box="[151,1255,1662,1686]" pageId="21" pageNumber="324">
FIGURES 1112. Larva and pupa of
<taxonomicName id="4C144D3D310BFF84FD94BD69D2C827B6" box="[558,747,1662,1686]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="21" pageNumber="324" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B960EAAC310BFF84FD94BD69D29427B6" bold="true" box="[558,695,1662,1686]" italics="true" pageId="21" pageNumber="324">Homoplectra</emphasis>
spp.
</taxonomicName>
11A11E,
<taxonomicName id="4C144D3D310BFF84FCE5BD69D4C127B6" authority="(Banks 1906)" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[863,1250,1662,1686]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="21" pageNumber="324" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC310BFF84FCE5BD69D46B27B6" bold="true" box="[863,1096,1662,1686]" italics="true" pageId="21" pageNumber="324">Homoplectra japonica</emphasis>
(
<bibRefCitation id="EF854B4F310BFF84FBEDBD69D4F927B6" author="Banks, N." box="[1111,1242,1662,1686]" pageId="21" pageNumber="324" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
)
</taxonomicName>
:
</emphasis>
11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width.
<emphasis id="B960EAAC310BFF84FBBEBDFDD10E2606" bold="true" pageId="21" pageNumber="324">
12A12C,
<taxonomicName id="4C144D3D310BFF84FBCEBDFDD10B2606" authority="(Ulmer 1907)" baseAuthorityName="Ulmer" baseAuthorityYear="1907" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="21" pageNumber="324" phylum="Arthropoda" rank="species" species="albomarginata">
<emphasis id="B960EAAC310BFF84FBCEBDFDD5BF2622" bold="true" box="[1140,1436,1770,1794]" italics="true" pageId="21" pageNumber="324">Homoplectra albomarginata</emphasis>
(
<bibRefCitation id="EF854B4F310BFF84FF24BC19D1032606" author="Ulmer, G." box="[158,288,1806,1830]" pageId="21" pageNumber="324" pagination="1 - 102" refId="ref18285" refString="Ulmer, G. (1907) Trichopteren. Part. 1. Collection of Zoologiques du Baron Edm. de Selys Longchamps, 6 (1), 1 - 102." type="journal article" year="1907">Ulmer 1907</bibRefCitation>
)
</taxonomicName>
:
</emphasis>
12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same). Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII.
</paragraph>
</caption>
<caption id="DF6B66363108FF87FF2DBC80D4432690" box="[151,1120,1943,1968]" pageId="22" pageNumber="325" startId="22.[151,250,1943,1967]" targetBox="[169,1415,195,1904]" targetPageId="22" targetType="figure">
<paragraph id="8BAB36BE3108FF87FF2DBC80D4432690" blockId="22.[151,1120,1943,1968]" box="[151,1120,1943,1968]" pageId="22" pageNumber="325">
FIGURE 13. Distributions of species of the
<taxonomicName id="4C144D3D3108FF87FDD0BC80D370268F" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[618,851,1943,1967]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="22" pageNumber="325" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3108FF87FDD0BC80D370268F" bold="true" box="[618,851,1943,1967]" italics="true" pageId="22" pageNumber="325">Homoplectra japonica</emphasis>
</taxonomicName>
Species Group in Japan.
</paragraph>
</caption>
<paragraph id="8BAB36BE3109FF86FF7DBB80D432208D" blockId="23.[151,1437,150,1257]" pageId="23" pageNumber="326">
<emphasis id="B960EAAC3109FF86FF7DBB80D1AD2190" bold="true" box="[199,398,150,177]" pageId="23" pageNumber="326">Female genitalia</emphasis>
(
<figureCitation id="132F2A3B3109FF86FE24BB80D2632191" box="[414,576,151,177]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="23" pageNumber="326">Figs 10G10I</figureCitation>
). Lateral lobes of segment VIII widely separated from each other in ventral aspect (
<figureCitation id="132F2A3B3109FF86FF51BBACD16421F5" box="[235,327,187,213]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="23" pageNumber="326">Fig. 10I</figureCitation>
), their mesal margins weakly concave. Segment IX quadrate in lateral aspect (
<figureCitation id="132F2A3B3109FF86FB12BBACD52C21F5" box="[1192,1295,187,213]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="23" pageNumber="326">Fig. 10G</figureCitation>
), pentagonal in dorsal aspect (
<figureCitation id="132F2A3B3109FF86FEE0BBC8D1E721D9" box="[346,452,223,249]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="23" pageNumber="326">Fig. 10H</figureCitation>
), with transverse sulcus dorsally on posterior 1/31/4; pair of sclerotized ribs forming angular ridges on vulvar scale basolaterally in ventral aspect (
<figureCitation id="132F2A3B3109FF86FCFFBA14D382203D" box="[837,929,259,285]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="23" pageNumber="326">Fig. 10I</figureCitation>
); mesal lobe weakly sclerotized, semicircular in lateral aspect (
<figureCitation id="132F2A3B3109FF86FEDEBA30D1F22061" box="[356,465,295,321]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="23" pageNumber="326">Fig. 10G</figureCitation>
). Vulvar scale large, tongue-like, membranous posteriorly; with pair of deep holes basolaterally in ventral aspect. Vaginal apparatus bulb-shaped anteriorly in dorsal aspect (
<figureCitation id="132F2A3B3109FF86FB27BA5CD52B2045" box="[1181,1288,331,357]" captionStart="FIGURE 10" captionStartId="20.[151,250,1734,1758]" captionTargetBox="[157,1429,351,1705]" captionTargetId="figure-119@20.[151,1435,345,1711]" captionTargetPageId="20" captionText="FIGURE 10. Adults of Homoplectra inazui sp. nov. 10A10E, male genitalia, type locality (Yabu-shi, Hyogo): 10A, left lateral, apicomesal part of inferior appendage enlarged; 10B, dorsal; 10C, ventral; 10D, phallic apparatus, left lateral; 10E, same, dorsal. 10F, male genitalia, variation (Hidaka-cho, Hyogo), lateral. 10G10I, female genitalia, type locality: 10G, lateral, 10H, dorsal; 10I, ventral." pageId="23" pageNumber="326">Fig. 10H</figureCitation>
), rectangular posteriorly; posterodorsal part mostly semi-membranous; longitudinal dorsal ridges divergent anteriorly, opening of duct of spermatheca completely surrounded by dark oval mark in dorsal aspect.
</paragraph>
<paragraph id="8BAB36BE3109FF86FF7DBAA0D23020F1" blockId="23.[151,1437,150,1257]" box="[199,531,439,465]" pageId="23" pageNumber="326">
<emphasis id="B960EAAC3109FF86FF7DBAA0D1B520F1" bold="true" box="[199,406,439,465]" pageId="23" pageNumber="326">Immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353109FF86FF7DBACCD1912531" pageId="23" pageNumber="326" type="materials_examined">
<paragraph id="8BAB36BE3109FF86FF7DBACCD2B22339" blockId="23.[151,1437,150,1257]" pageId="23" pageNumber="326">
<materialsCitation id="3B7C3CE33109FF86FF7DBACCD2B22339" collectingDate="2006-05-14" collectionCode="KPM-NK" collectorName="K. Inazu" county="Male" elevation="600" latitude="35.4315" location="Yado" longLatPrecision="6" longitude="134.5819" municipality="Uwano" pageId="23" pageNumber="326" specimenCode="TN00690" specimenCount="1" stateProvince="Hyogo" typeStatus="holotype">
<emphasis id="B960EAAC3109FF86FF7DBACCD11820D5" bold="true" box="[199,315,475,501]" pageId="23" pageNumber="326">
<typeStatus id="54AF881C3109FF86FF7DBACCD11520D5" box="[199,310,475,501]" pageId="23" pageNumber="326" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62CA4E323109FF86FEF8BACCD15F20D5" box="[322,380,475,501]" pageId="23" pageNumber="326">Male</collectingCounty>
(in alcohol):
<collectingMunicipality id="6BCFACC43109FF86FDAEBACCD24B20D5" box="[532,616,475,501]" pageId="23" pageNumber="326">Uwano</collectingMunicipality>
,
<location id="8ECB60653109FF86FDC8BACCD28C20D5" LSID="urn:lsid:plazi:treatment:03BD87A8310DFF86FF2DBFBDD2D025C9:8ECB60653109FF86FDC8BACCD28C20D5" box="[626,687,475,501]" county="Male" latitude="35.4315" longLatPrecision="6" longitude="134.5819" municipality="Uwano" name="Yado" pageId="23" pageNumber="326" stateProvince="Hyogo">Yado</location>
,
<location id="8ECB60653109FF86FD00BACCD36420D5" LSID="urn:lsid:plazi:treatment:03BD87A8310DFF86FF2DBFBDD2D025C9:8ECB60653109FF86FD00BACCD36420D5" box="[698,839,475,501]" county="Male" latitude="35.4315" longLatPrecision="6" longitude="134.5819" municipality="Uwano" name="Muraoka-ku" pageId="23" pageNumber="326" stateProvince="Hyogo">Muraoka-ku</location>
,
<location id="8ECB60653109FF86FCE9BACCD3E520D5" LSID="urn:lsid:plazi:treatment:03BD87A8310DFF86FF2DBFBDD2D025C9:8ECB60653109FF86FCE9BACCD3E520D5" box="[851,966,475,501]" county="Male" latitude="35.4315" longLatPrecision="6" longitude="134.5819" municipality="Uwano" name="Kami-cho" pageId="23" pageNumber="326" stateProvince="Hyogo">Kami-cho</location>
,
<collectingRegion id="49D0F85C3109FF86FC6BBACCD40320D5" box="[977,1056,475,501]" country="Japan" name="Hyogo" pageId="23" pageNumber="326">Hyogo</collectingRegion>
,
<geoCoordinate id="EE2050793109FF86FB96BACDD48A20D5" box="[1068,1193,474,501]" degrees="35.4315" direction="north" orientation="latitude" pageId="23" pageNumber="326" precision="5" value="35.4315">35.4315°N</geoCoordinate>
,
<geoCoordinate id="EE2050793109FF86FB0EBACDD51E20D5" box="[1204,1341,474,501]" degrees="134.5819" direction="east" orientation="longitude" pageId="23" pageNumber="326" precision="5" value="134.5819">134.5819°E</geoCoordinate>
, alt.
<quantity id="4CEC9B5B3109FF86FAC8BACCD08C2339" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="326" unit="m" value="600.0">
<elevation id="0039D18D3109FF86FAC8BACCD08C2339" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="326" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="FFAA107E3109FF86FF01BAE8D10F2339" box="[187,300,511,537]" pageId="23" pageNumber="326" value="2006-05-14">
<collectingDate id="EFEEE9963109FF86FF01BAE8D10F2339" box="[187,300,511,537]" pageId="23" pageNumber="326" value="2006-05-14">14.v.2006</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FE8DBAE8D1B42339" box="[311,407,511,537]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FE12BAE8D2352339" box="[424,534,511,537]" pageId="23" pageNumber="326">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53109FF86FDA7BAE8D2A92339" box="[541,650,511,537]" collectionCode="TN" pageId="23" pageNumber="326">TN00690</specimenCode>
)
</materialsCitation>
</paragraph>
<paragraph id="8BAB36BE3109FF86FF7DB934D1372389" blockId="23.[151,1437,150,1257]" pageId="23" pageNumber="326">
<materialsCitation id="3B7C3CE33109FF86FF7DB934D1332341" collectingDate="2004-05-05" collectionCode="KPM-NK" collectorName="K. Inazu" county="Male" elevation="600" latitude="35.4315" location="Yado" longLatPrecision="6" longitude="134.5819" municipality="Uwano" pageId="23" pageNumber="326" specimenCode="TN00700, TN00710" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hyogo" typeStatus="paratype">
<emphasis id="B960EAAC3109FF86FF7DB934D165231D" bold="true" box="[199,326,547,573]" pageId="23" pageNumber="326">
<typeStatus id="54AF881C3109FF86FF7DB934D162231D" box="[199,321,547,573]" pageId="23" pageNumber="326" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D12FD373109FF86FEF4B934D18B231D" box="[334,424,547,573]" count="3" pageId="23" pageNumber="326" type="male">3 males</specimenCount>
,
<specimenCount id="9D12FD373109FF86FE0EB934D207231D" box="[436,548,547,573]" count="2" pageId="23" pageNumber="326" type="female">2 females</specimenCount>
, same locality as holotype,
<date id="FFAA107E3109FF86FCE6B935D39C231D" box="[860,959,546,573]" pageId="23" pageNumber="326" value="2004-05-05">
<collectingDate id="EFEEE9963109FF86FCE6B935D39C231D" box="[860,959,546,573]" pageId="23" pageNumber="326" value="2004-05-05">5.v.2004</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FC76B934D40E231D" box="[972,1069,547,573]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FB84B934D48F231D" box="[1086,1196,547,573]" pageId="23" pageNumber="326">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53109FF86FB0EB934D501231D" box="[1204,1314,547,573]" collectionCode="TN" pageId="23" pageNumber="326">TN00700</specimenCode>
,
<collectionCode id="ED05AE7B3109FF86FA94B934D5BF231D" box="[1326,1436,547,573]" pageId="23" pageNumber="326">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53109FF86FF2DB950D12B2341" box="[151,264,583,609]" collectionCode="TN" pageId="23" pageNumber="326">TN00710</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33109FF86FEA4B950D13323A5" collectingDate="2005-05-28" collectionCode="KPM-NK" collectorName="Soryuno-taki &amp; Itashino &amp; Muraoka-ku &amp; Kami-cho &amp; K. Inazu" elevation="600" pageId="23" pageNumber="326" specimenCode="TN00720" specimenCount="3" specimenCount-male="3" stateProvince="Hyogo" typeStatus="paratype">
<specimenCount id="9D12FD373109FF86FEA4B950D1582341" box="[286,379,583,609]" count="3" pageId="23" pageNumber="326" type="male">3 males</specimenCount>
,
<collectorName id="26E153683109FF86FE30B950D23C2341" box="[394,543,583,609]" pageId="23" pageNumber="326">Soryuno-taki</collectorName>
,
<collectorName id="26E153683109FF86FD94B950D2A82341" box="[558,651,583,609]" pageId="23" pageNumber="326">Itashino</collectorName>
,
<collectorName id="26E153683109FF86FD23B950D3052341" box="[665,806,583,609]" pageId="23" pageNumber="326">Muraoka-ku</collectorName>
,
<collectorName id="26E153683109FF86FC8FB950D38B2341" box="[821,936,583,609]" pageId="23" pageNumber="326">Kami-cho</collectorName>
, alt.
<quantity id="4CEC9B5B3109FF86FC5FB950D4042341" box="[997,1063,583,609]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="326" unit="m" value="600.0">
<elevation id="0039D18D3109FF86FC5FB950D4042341" box="[997,1063,583,609]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="326" unit="m" value="600.0">600m</elevation>
</quantity>
,
<date id="FFAA107E3109FF86FB8CB951D4852341" box="[1078,1190,582,609]" pageId="23" pageNumber="326" value="2005-05-28">
<collectingDate id="EFEEE9963109FF86FB8CB951D4852341" box="[1078,1190,582,609]" pageId="23" pageNumber="326" value="2005-05-28">28.v.2005</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FB0CB950D5392341" box="[1206,1306,583,609]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FA94B950D5BF2341" box="[1326,1436,583,609]" pageId="23" pageNumber="326">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53109FF86FF2DB97CD12B23A5" box="[151,264,619,645]" collectionCode="TN" pageId="23" pageNumber="326">TN00720</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33109FF86FEA7B97CD1332389" collectingDate="2005-05-28" collectionCode="KPM-NK" collectorName="Torokawa-daki &amp; Itashino &amp; Muraoka-ku &amp; Kami-cho &amp; K. Inazu" elevation="900" pageId="23" pageNumber="326" specimenCode="TN00730" specimenCount="1" specimenCount-male="1" stateProvince="Hyogo" typeStatus="paratype">
<specimenCount id="9D12FD373109FF86FEA7B97CD14F23A5" box="[285,364,619,645]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
,
<collectorName id="26E153683109FF86FEC3B97CD20623A5" box="[377,549,619,645]" pageId="23" pageNumber="326">Torokawa-daki</collectorName>
,
<collectorName id="26E153683109FF86FD88B97CD2AC23A5" box="[562,655,619,645]" pageId="23" pageNumber="326">Itashino</collectorName>
,
<collectorName id="26E153683109FF86FD26B97CD30A23A5" box="[668,809,619,645]" pageId="23" pageNumber="326">Muraoka-ku</collectorName>
,
<collectorName id="26E153683109FF86FC8DB97CD38923A5" box="[823,938,619,645]" pageId="23" pageNumber="326">Kami-cho</collectorName>
,
<date id="FFAA107E3109FF86FC0DB97DD40423A5" box="[951,1063,618,645]" pageId="23" pageNumber="326" value="2005-05-28">
<collectingDate id="EFEEE9963109FF86FC0DB97DD40423A5" box="[951,1063,618,645]" pageId="23" pageNumber="326" value="2005-05-28">28.v.2005</collectingDate>
</date>
, alt.
<quantity id="4CEC9B5B3109FF86FBDBB97CD48F23A5" box="[1121,1196,619,645]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="326" unit="m" value="900.0">
<elevation id="0039D18D3109FF86FBDBB97CD48F23A5" box="[1121,1196,619,645]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="326" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<collectorName id="26E153683109FF86FB03B97CD53823A5" box="[1209,1307,619,645]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FA94B97CD5BF23A5" box="[1326,1436,619,645]" pageId="23" pageNumber="326">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53109FF86FF2DB998D12B2389" box="[151,264,655,681]" collectionCode="TN" pageId="23" pageNumber="326">TN00730</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAB36BE3109FF86FF7DB9A5D1912531" blockId="23.[151,1437,150,1257]" pageId="23" pageNumber="326">
<emphasis id="B960EAAC3109FF86FF7DB9A5D34E23ED" bold="true" box="[199,877,690,717]" pageId="23" pageNumber="326">
Other specimens examined.
<materialsCitation id="3B7C3CE33109FF86FDA4B9A5D34B23ED" box="[542,872,690,717]" country="Japan" location="Hyogo" pageId="23" pageNumber="326" specimenCount="1" stateProvince="Hyogo">
<collectingCountry id="F303762E3109FF86FDA4B9A5D25F23EC" box="[542,636,690,716]" name="Japan" pageId="23" pageNumber="326">JAPAN</collectingCountry>
: HONSHU:
<collectingRegion id="49D0F85C3109FF86FCACB9A4D34B23ED" box="[790,872,691,717]" country="Japan" name="Hyogo" pageId="23" pageNumber="326">Hyogo</collectingRegion>
</materialsCitation>
:
</emphasis>
<specimenCount id="9D12FD373109FF86FCCDB9A4D3FE23ED" box="[887,989,691,717]" count="1" pageId="23" pageNumber="326" type="female">1 female</specimenCount>
, Hattanno-taki, Niiya, Ojiro-ku, Kami-cho, alt.
<quantity id="4CEC9B5B3109FF86FF41B9C1D16623D1" box="[251,325,726,753]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="326" unit="m" value="750.0">750 m</quantity>
,
<date id="FFAA107E3109FF86FEE9B9C0D19C23D1" box="[339,447,727,753]" pageId="23" pageNumber="326" value="2006-06-04">4.vi.2006</date>
, K. Inazu;
<materialsCitation id="3B7C3CE33109FF86FD84B9C0D1302235" collectingDate="2004-05-09" collectionCode="KI" collectorName="Myoken-zan &amp; Ishihara &amp; Yoka-cho &amp; Yabu-shi &amp; K. Inazu" country="Japan" elevation="700" pageId="23" pageNumber="326" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373109FF86FD84B9C0D2AD23D1" box="[574,654,727,753]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
,
<collectorName id="26E153683109FF86FD21B9C0D30823D1" box="[667,811,727,753]" pageId="23" pageNumber="326">Myoken-zan</collectorName>
,
<collectorName id="26E153683109FF86FC82B9C0D3B623D1" box="[824,917,727,753]" pageId="23" pageNumber="326">Ishihara</collectorName>
,
<collectorName id="26E153683109FF86FC1AB9C0D42C23D1" box="[928,1039,727,753]" pageId="23" pageNumber="326">Yoka-cho</collectorName>
,
<collectorName id="26E153683109FF86FBA0B9C0D4A223D1" box="[1050,1153,727,753]" pageId="23" pageNumber="326">Yabu-shi</collectorName>
, alt.
<quantity id="4CEC9B5B3109FF86FB03B9C0D52723D1" box="[1209,1284,727,753]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="326" unit="m" value="700.0">
<elevation id="0039D18D3109FF86FB03B9C0D52723D1" box="[1209,1284,727,753]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="326" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="FFAA107E3109FF86FAABB9C0D55723D1" box="[1297,1396,727,753]" pageId="23" pageNumber="326" value="2004-05-09">
<collectingDate id="EFEEE9963109FF86FAABB9C0D55723D1" box="[1297,1396,727,753]" pageId="23" pageNumber="326" value="2004-05-09">9.v.2004</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FA3BB9C0D0F62235" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FF53B9ECD1292235" box="[233,266,763,789]" pageId="23" pageNumber="326">KI</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D12FD373109FF86FE9BB9ECD1512235" box="[289,370,763,789]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
, Betsugu, Sekinomiya, Yabu-shi, alt.
<specimenCount id="9D12FD373109FF86FC9FB9ECD3552235" box="[805,886,763,789]" count="700" pageId="23" pageNumber="326" type="generic">
<quantity id="4CEC9B5B3109FF86FC9FB9ECD3512235" box="[805,882,763,789]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="326" unit="m" value="700.0">700 m</quantity>
<materialsCitation id="3B7C3CE33109FF86FCC8B9ECD3552235" box="[882,886,763,789]" collectingDate="2007-05-03" collectionCode="KI" collectorName="Inazu" country="Japan" pageId="23" pageNumber="326" specimenCount="1">.</materialsCitation>
</specimenCount>
<date id="FFAA107E3109FF86FC3AB9ECD3C12235" box="[896,994,763,789]" pageId="23" pageNumber="326" value="2007-05-03">
<collectingDate id="EFEEE9963109FF86FC3AB9ECD3C12235" box="[896,994,763,789]" pageId="23" pageNumber="326" value="2007-05-03">3.v.2007</collectingDate>
</date>
, K.
<collectorName id="26E153683109FF86FBACB9ECD4772235" box="[1046,1108,763,789]" pageId="23" pageNumber="326">Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FBD2B9ECD4AA2235" box="[1128,1161,763,789]" pageId="23" pageNumber="326">KI</collectionCode>
);
<materialsCitation id="3B7C3CE33109FF86FB1AB9ECD18C2219" collectingDate="2010-05-05" collectionCode="KI" collectorName="K. Inazu" country="Japan" pageId="23" pageNumber="326" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373109FF86FB1AB9ECD4D22235" box="[1184,1265,763,789]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
, same locality,
<date id="FFAA107E3109FF86FF2DB809D0DA2219" box="[151,249,798,825]" pageId="23" pageNumber="326" value="2010-05-05">
<collectingDate id="EFEEE9963109FF86FF2DB809D0DA2219" box="[151,249,798,825]" pageId="23" pageNumber="326" value="2010-05-05">5.v.2010</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FEB0B808D14C2219" box="[266,367,799,825]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FE3EB808D1862219" box="[388,421,799,825]" pageId="23" pageNumber="326">KI</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D12FD373109FF86FE05B808D2042219" box="[447,551,799,825]" count="1" pageId="23" pageNumber="326" type="female">1 female</specimenCount>
, Hachibuse-kogen, Sekimiya-cho, Yabu-shi,
<date id="FFAA107E3109FF86FB88B808D4812219" box="[1074,1186,799,825]" pageId="23" pageNumber="326" value="2009-05-18">18.v.2009</date>
, K. Inazu;
<materialsCitation id="3B7C3CE33109FF86FA91B808D42D227D" collectingDate="2007-05-11" collectionCode="KI" collectorName="Takenocho-mihara &amp; Toyooka-shi &amp; K. Inazu" country="Japan" elevation="420" pageId="23" pageNumber="326" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="9D12FD373109FF86FA91B808D55D2219" box="[1323,1406,799,825]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
,
<specimenCount id="9D12FD373109FF86FA34B808D0D2227D" count="2" pageId="23" pageNumber="326" type="female">2 females</specimenCount>
,
<collectorName id="26E153683109FF86FF46B854D1F2227D" box="[252,465,835,861]" pageId="23" pageNumber="326">Takenocho-mihara</collectorName>
,
<collectorName id="26E153683109FF86FE66B854D248227D" box="[476,619,835,861]" pageId="23" pageNumber="326">Toyooka-shi</collectorName>
, alt.
<quantity id="4CEC9B5B3109FF86FD1AB854D2C9227D" box="[672,746,835,861]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="23" pageNumber="326" unit="m" value="420.0">
<elevation id="0039D18D3109FF86FD1AB854D2C9227D" box="[672,746,835,861]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="23" pageNumber="326" unit="m" value="420.0">420 m</elevation>
</quantity>
,
<date id="FFAA107E3109FF86FD4CB854D346227D" box="[758,869,835,861]" pageId="23" pageNumber="326" value="2007-05-11">
<collectingDate id="EFEEE9963109FF86FD4CB854D346227D" box="[758,869,835,861]" pageId="23" pageNumber="326" value="2007-05-11">11.v.2007</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FCC8B854D3F1227D" box="[882,978,835,861]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FC59B854D427227D" box="[995,1028,835,861]" pageId="23" pageNumber="326">KI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33109FF86FBA0B854D11522A1" collectingDate="2007-06-01" collectionCode="KI" collectorName="K. Inazu" country="Japan" elevation="420" pageId="23" pageNumber="326" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D12FD373109FF86FBA0B854D45D227D" box="[1050,1150,835,861]" count="1" pageId="23" pageNumber="326" type="female">1 female</specimenCount>
, same locality,
<date id="FFAA107E3109FF86FA91B854D5B4227D" box="[1323,1431,835,861]" pageId="23" pageNumber="326" value="2007-06-01">
<collectingDate id="EFEEE9963109FF86FA91B854D5B4227D" box="[1323,1431,835,861]" pageId="23" pageNumber="326" value="2007-06-01">1.vi.2007</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FF2DB870D0DA22A1" box="[151,249,871,897]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FEB6B870D10E22A1" box="[268,301,871,897]" pageId="23" pageNumber="326">KI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33109FF86FEF9B870D34022A1" box="[323,867,871,897]" collectingDate="2013-05-14" collectionCode="KI" collectorName="K. Inazu" country="Japan" elevation="420" pageId="23" pageNumber="326" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373109FF86FEF9B870D1B022A1" box="[323,403,871,897]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
, same locality,
<date id="FFAA107E3109FF86FDFFB870D29522A1" box="[581,694,871,897]" pageId="23" pageNumber="326" value="2013-05-14">
<collectingDate id="EFEEE9963109FF86FDFFB870D29522A1" box="[581,694,871,897]" pageId="23" pageNumber="326" value="2013-05-14">14.v.2013</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FD79B870D30522A1" box="[707,806,871,897]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FC82B870D37A22A1" box="[824,857,871,897]" pageId="23" pageNumber="326">KI</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D12FD373109FF86FCCAB870D3F522A1" box="[880,982,871,897]" count="1" pageId="23" pageNumber="326" type="female">1 female</specimenCount>
, Mihara-oku, Takeno-cho, Toyooka-shi,
<date id="FFAA107E3109FF86FF2DB89CD1242285" box="[151,263,907,933]" pageId="23" pageNumber="326" value="2009-05-29">29.v.2009</date>
, K. Inazu;
<specimenCount id="9D12FD373109FF86FEC5B89CD1EF2285" box="[383,460,907,933]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
,
<specimenCount id="9D12FD373109FF86FE6CB89CD21A2285" box="[470,569,907,933]" count="1" pageId="23" pageNumber="326" type="female">1 female</specimenCount>
, Inab, Hidakacho, Toyooka-shi, alt.
<quantity id="4CEC9B5B3109FF86FC73B89CD4322285" box="[969,1041,907,933]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="23" pageNumber="326" unit="m" value="600.0">600 m</quantity>
,
<date id="FFAA107E3109FF86FBA6B89CD4B72285" box="[1052,1172,907,933]" pageId="23" pageNumber="326" value="2003-04-29">29.iv.2003</date>
, K. Inazu;
<materialsCitation id="3B7C3CE33109FF86FAB1B89CD23522E9" collectingDate="2004-04-25" collectionCode="KI" collectorName="K. Inazu" country="Japan" elevation="420" pageId="23" pageNumber="326" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373109FF86FAB1B89CD57B2285" box="[1291,1368,907,933]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
, same locality,
<date id="FFAA107E3109FF86FF4DB8B9D15322E9" box="[247,368,942,969]" pageId="23" pageNumber="326" value="2004-04-25">
<collectingDate id="EFEEE9963109FF86FF4DB8B9D15322E9" box="[247,368,942,969]" pageId="23" pageNumber="326" value="2004-04-25">25.iv.2004</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FEC1B8B8D1F822E9" box="[379,475,943,969]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FE51B8B8D22F22E9" box="[491,524,943,969]" pageId="23" pageNumber="326">KI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33109FF86FD9BB8B8D51C22E9" box="[545,1343,942,969]" collectingDate="2004-05-30" collectionCode="KI" collectorName="Sugawara &amp; Shinonsen-cho &amp; K. Inazu" country="Japan" elevation="500" pageId="23" pageNumber="326" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373109FF86FD9BB8B8D24D22E9" box="[545,622,943,969]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
,
<collectorName id="26E153683109FF86FDC0B8B8D2C822E9" box="[634,747,943,969]" pageId="23" pageNumber="326">Sugawara</collectorName>
,
<collectorName id="26E153683109FF86FD4CB8B8D3BC22E9" box="[758,927,943,969]" pageId="23" pageNumber="326">Shinonsen-cho</collectorName>
, alt.
<quantity id="4CEC9B5B3109FF86FC6EB8B9D43E22E9" box="[980,1053,942,969]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="326" unit="m" value="500.0">
<elevation id="0039D18D3109FF86FC6EB8B9D43E22E9" box="[980,1053,942,969]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="326" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="FFAA107E3109FF86FB92B8B8D4BA22E9" box="[1064,1177,943,969]" pageId="23" pageNumber="326" value="2004-05-30">
<collectingDate id="EFEEE9963109FF86FB92B8B8D4BA22E9" box="[1064,1177,943,969]" pageId="23" pageNumber="326" value="2004-05-30">30.v.2004</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FB1EB8B8D52722E9" box="[1188,1284,943,969]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FAAFB8B8D51522E9" box="[1301,1334,943,969]" pageId="23" pageNumber="326">KI</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D12FD373109FF86FAF0B8B8D5BB22E9" box="[1354,1432,943,969]" count="1" pageId="23" pageNumber="326" type="male">1 male</specimenCount>
,
<specimenCount id="9D12FD373109FF86FF2DB8C4D12522CD" box="[151,262,979,1005]" count="3" pageId="23" pageNumber="326" type="female">3 females</specimenCount>
, Koke-daki, Sugawara, Shinonsen-cho, alt.
<quantity id="4CEC9B5B3109FF86FD4BB8C4D31822CD" box="[753,827,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="326" unit="m" value="800.0">800 m</quantity>
,
<date id="FFAA107E3109FF86FCFDB8C4D39022CD" box="[839,947,979,1005]" pageId="23" pageNumber="326" value="2003-06-08">8.vi.2003</date>
, K. Inazu;
<materialsCitation id="3B7C3CE33109FF86FB94B8C4D18C2531" collectingDate="2004-05-30" collectionCode="KI" collectorName="K. Inazu" country="Japan" elevation="500" pageId="23" pageNumber="326" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="9D12FD373109FF86FB94B8C4D4A422CD" box="[1070,1159,979,1005]" count="3" pageId="23" pageNumber="326" type="male">3 males</specimenCount>
,
<specimenCount id="9D12FD373109FF86FB29B8C4D4D422CD" box="[1171,1271,979,1005]" count="1" pageId="23" pageNumber="326" type="female">1 female</specimenCount>
, same locality,
<date id="FFAA107E3109FF86FF2DB8E0D12B2531" box="[151,264,1015,1041]" pageId="23" pageNumber="326" value="2004-05-30">
<collectingDate id="EFEEE9963109FF86FF2DB8E0D12B2531" box="[151,264,1015,1041]" pageId="23" pageNumber="326" value="2004-05-30">30.v.2004</collectingDate>
</date>
,
<collectorName id="26E153683109FF86FEA9B8E0D1572531" box="[275,372,1015,1041]" pageId="23" pageNumber="326">K. Inazu</collectorName>
(
<collectionCode id="ED05AE7B3109FF86FE3EB8E0D1852531" box="[388,422,1015,1041]" pageId="23" pageNumber="326">KI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353109FF86FF7DBF0CD1B32579" pageId="23" pageNumber="326" type="etymology">
<paragraph id="8BAB36BE3109FF86FF7DBF0CD1B32579" blockId="23.[151,1437,150,1257]" pageId="23" pageNumber="326">
<emphasis id="B960EAAC3109FF86FF7DBF0CD16D2515" bold="true" box="[199,334,1051,1077]" pageId="23" pageNumber="326">Etymology.</emphasis>
This species is dedicated to Mr Kazuyuki Inazu, who provided me with many valuable specimens including this species.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353109FF86FF7DBF75D203255D" box="[199,544,1122,1149]" pageId="23" pageNumber="326" type="distribution">
<paragraph id="8BAB36BE3109FF86FF7DBF75D203255D" blockId="23.[151,1437,150,1257]" box="[199,544,1122,1149]" pageId="23" pageNumber="326">
<emphasis id="B960EAAC3109FF86FF7DBF75D142255C" bold="true" box="[199,353,1122,1148]" pageId="23" pageNumber="326">Distribution.</emphasis>
Central Honshu.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353109FF86FF7DBF91D2D025C9" pageId="23" pageNumber="326" type="biology_ecology">
<paragraph id="8BAB36BE3109FF86FF7DBF91D10D25E5" blockId="23.[151,1437,150,1257]" pageId="23" pageNumber="326">
<emphasis id="B960EAAC3109FF86FF7DBF91D1042580" bold="true" box="[199,295,1158,1184]" pageId="23" pageNumber="326">Biology.</emphasis>
<typeStatus id="54AF881C3109FF86FE94BF90D1452581" box="[302,358,1159,1185]" pageId="23" pageNumber="326">Type</typeStatus>
series specimens of this species were collected near small streams or falls in mountainous areas (
<bibRefCitation id="EF854B4F3109FF86FF25BFBCD10025E5" author="Inazu, K." box="[159,291,1195,1221]" pageId="23" pageNumber="326" pagination="65 - 78" refId="ref17158" refString="Inazu, K. (2013) Collection records of caddisflies in the Yada River. Hyogo Freshwater Biology, 64, 65 - 78 [in Japanese]." type="journal article" year="2013">Inazu 2013</bibRefCitation>
).
</paragraph>
<paragraph id="8BAB36BE3109FF86FF7DBFD8D2D025C9" blockId="23.[151,1437,150,1257]" box="[199,755,1231,1257]" pageId="23" pageNumber="326">
<emphasis id="B960EAAC3109FF86FF7DBFD8D1A025C9" bold="true" box="[199,387,1231,1257]" pageId="23" pageNumber="326">Japanese name.</emphasis>
Inazu-kimadara-shima-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,627 @@
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<mods:title id="E01C8F97D2B8CDDD5CD7EB9C8283FD7E">Redescriptions of Homoplectra japonica (Banks 1906) and H. albomarginata (Ulmer 1907) (Trichoptera, Hydropsychidae), with descriptions of five new similar species from Japan</mods:title>
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<mods:namePart id="4C13A90762A7D70C20678E9A3C069447">Nozaki, Takao</mods:namePart>
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<mods:date id="90C1BE7C73415F63B002628A97882148">2025</mods:date>
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<paragraph id="8BAB36BE3111FF9EFF2DBFE5D220242D" blockId="15.[151,515,1266,1329]" box="[151,515,1266,1293]" pageId="15" pageNumber="318">
<heading id="D0E381D23111FF9EFF2DBFE5D220242D" bold="true" box="[151,515,1266,1293]" fontSize="11" level="1" pageId="15" pageNumber="318" reason="1">
<emphasis id="B960EAAC3111FF9EFF2DBFE5D220242D" bold="true" box="[151,515,1266,1293]" pageId="15" pageNumber="318">
<taxonomicName id="4C144D3D3111FF9EFF2DBFE5D1BC242D" authorityName="Nozaki" authorityYear="2025" box="[151,415,1266,1293]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="318" phylum="Arthropoda" rank="species" species="bicornis" status="sp. nov.">
<emphasis id="B960EAAC3111FF9EFF2DBFE5D1BC242D" bold="true" box="[151,415,1266,1293]" italics="true" pageId="15" pageNumber="318">Homoplectra bicornis</emphasis>
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<taxonomicNameLabel id="A25357D73111FF9EFE1CBFE4D220242D" box="[422,515,1267,1293]" pageId="15" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BAB36BE3111FF9EFF2DBE00D1362411" blockId="15.[151,515,1266,1329]" box="[151,277,1303,1329]" pageId="15" pageNumber="318">
(
<figureCitation id="132F2A3B3111FF9EFF25BE00D0C02411" box="[159,227,1303,1329]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8</figureCitation>
,
<figureCitation id="132F2A3B3111FF9EFF4ABE00D12F2411" box="[240,268,1303,1329]" captionStart="FIGURE 13" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[169,1415,195,1904]" captionTargetId="figure-33@22.[157,1430,181,1919]" captionTargetPageId="22" captionText="FIGURE 13. Distributions of species of the Homoplectra japonica Species Group in Japan." pageId="15" pageNumber="318">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E65353111FF9EFF2DBE49D56C24E1" pageId="15" pageNumber="318" type="diagnosis">
<paragraph id="8BAB36BE3111FF9EFF2DBE49D56C24E1" blockId="15.[151,1436,1374,1473]" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF2DBE49D1312458" bold="true" box="[151,274,1374,1400]" pageId="15" pageNumber="318">Diagnosis.</emphasis>
The male of this species is easily distinguishable from those of other congeneric Japanese species by the pair of posterodorsal horn-like lobes of the segment IX (
<figureCitation id="132F2A3B3111FF9EFC88BE94D3AA24BD" box="[818,905,1411,1437]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8B</figureCitation>
). The female is also easily distinguishable from those of known Japanese species by the bottle-like sclerotization of the ventral side of segment X (
<figureCitation id="132F2A3B3111FF9EFB52BEB0D56124E1" box="[1256,1346,1447,1473]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8H</figureCitation>
).
</paragraph>
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<subSubSection id="C30E65353111FF80FF2DBD05D230208D" lastPageId="17" lastPageNumber="320" pageId="15" pageNumber="318" type="description">
<paragraph id="8BAB36BE3111FF9EFF2DBD05D10B270C" blockId="15.[151,296,1554,1580]" box="[151,296,1554,1580]" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF2DBD05D10B270C" bold="true" box="[151,296,1554,1580]" pageId="15" pageNumber="318">Description</emphasis>
</paragraph>
<paragraph id="8BAB36BE3111FF9EFF2DBD4DD1AD279D" blockId="15.[151,1437,1626,2013]" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF2DBD4DD0C02754" bold="true" box="[151,227,1626,1652]" pageId="15" pageNumber="318">Adult.</emphasis>
General morphology and coloration similar to those of
<taxonomicName id="4C144D3D3111FF9EFCE1BD4CD47A2754" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[859,1113,1627,1653]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="15" pageNumber="318" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3111FF9EFCE1BD4CD47A2754" box="[859,1113,1627,1653]" italics="true" pageId="15" pageNumber="318">Homoplectra japonica</emphasis>
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, yellow markings on female wings often indistinct. Forewings each 6.09.0 mm long in male (n = 10), 7.510.0 mm long in female (n = 3),
<quantity id="4CEC9B5B3111FF9EFAC3BD68D0E0279D" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="15" pageNumber="318" unit="mm" value="7.2">7.2 mm</quantity>
in male
<typeStatus id="54AF881C3111FF9EFE9EBDB4D1A9279D" box="[292,394,1699,1725]" pageId="15" pageNumber="318" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BAB36BE3111FF81FF7DBDD0D114208D" blockId="15.[151,1437,1626,2013]" lastBlockId="16.[151,1437,151,429]" lastPageId="16" lastPageNumber="319" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF7DBDD0D15027C0" bold="true" box="[199,371,1734,1761]" pageId="15" pageNumber="318">Male genitalia</emphasis>
(8A8E). Ventrolateral sides of synsclerotized segment IX with triangular anterior projection in lateral aspect (
<figureCitation id="132F2A3B3111FF9EFEE0BDFCD1962625" box="[346,437,1771,1797]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8A</figureCitation>
); posteroventral lobe triangular and tiny or lacking (
<figureCitation id="132F2A3B3111FF9EFBB3BDFCD4402625" box="[1033,1123,1771,1797]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8C</figureCitation>
); dorsal part of segment IX rectangular in dorsal aspect (
<figureCitation id="132F2A3B3111FF9EFE5FBC18D2632609" box="[485,576,1807,1833]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8B</figureCitation>
), with pair of straight, horn-like lobes posterolaterally; broadly fused with segment X laterally (
<figureCitation id="132F2A3B3111FF9EFEC5BC24D1FA266D" box="[383,473,1843,1869]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8A</figureCitation>
). Segment X tapering to acute apex in dorsal aspect (
<figureCitation id="132F2A3B3111FF9EFB9EBC24D458266D" box="[1060,1147,1843,1869]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8B</figureCitation>
), bilobed posteriorly; pair of posterolateral setose areas protruding posteriorly (
<figureCitation id="132F2A3B3111FF9EFD65BC40D3142651" box="[735,823,1879,1905]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8A</figureCitation>
); pair of posterior processes of segment X (p.p.X) long, spine-like, directed posterodorsad, each fused with lateral strip of genital chamber (l.s.) near posteroventral corner of segment X (
<figureCitation id="132F2A3B3111FF9EFEF8BC88D1BD2699" box="[322,414,1951,1977]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8A</figureCitation>
). Genital chamber with pair of sclerotized lateral strips curved dorsad near posteroventral corner of segment X (
<figureCitation id="132F2A3B3111FF9EFE31BCD4D1C726FD" box="[395,484,1987,2013]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="15" pageNumber="318">Fig. 8A</figureCitation>
); pair of inner processes (i.p.) long, spine-like, each arising from near anterior part of lateral strip, slightly curved ventrad, approximately as long as posterior processes of segment X (
<figureCitation id="132F2A3B310EFF81FB62BB80D5102191" box="[1240,1331,151,177]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8A</figureCitation>
). Inferior appendages each without distal segment, slightly clavate in lateral aspect (
<figureCitation id="132F2A3B310EFF81FC68BBACD40821F5" box="[978,1067,187,213]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8A</figureCitation>
), extending slightly beyond apex of segment X; each weakly convex subapicomesally, with 510 spine-like setae (
<figureCitation id="132F2A3B310EFF81FB80BBC8D4B621D9" box="[1082,1173,223,249]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Figs 8A</figureCitation>
inset, 8B, 8C). Phallic apparatus arcuate in lateral aspect (
<figureCitation id="132F2A3B310EFF81FD9DBA14D2A0203D" box="[551,643,259,285]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8D</figureCitation>
); dorsal process of phallotheca long and narrowly triangular in dorsal aspect (
<figureCitation id="132F2A3B310EFF81FF57BA30D1662061" box="[237,325,295,321]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8E</figureCitation>
); pair of lateral processes of phallotheca 1/5 as long as dorsal process, usually acute apically (
<figureCitation id="132F2A3B310EFF81FAD5BA30D0992045" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8D</figureCitation>
) but rarely blunt; pair of ventral processes longer than dorsal process, each with apical half weakly sinuated in dorsal aspect (
<figureCitation id="132F2A3B310EFF81FE81BA78D1B020A9" box="[315,403,367,393]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8E</figureCitation>
). Aedeagal stem half-pipe-like, moderately curved ventrad; head about 1/7 as long as stem (
<figureCitation id="132F2A3B310EFF81FF25BA84D108208D" box="[159,299,403,429]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Figs 8D, 8E</figureCitation>
).
</paragraph>
<caption id="DF6B6636310EFF81FF2DBD91D3A6262A" pageId="16" pageNumber="319" startId="16.[151,250,1670,1694]" targetBox="[157,1430,459,1641]" targetPageId="16" targetType="figure">
<paragraph id="8BAB36BE310EFF81FF2DBD91D3A6262A" blockId="16.[151,1437,1670,1803]" pageId="16" pageNumber="319">
<emphasis id="B960EAAC310EFF81FF2DBD91D2E327BF" bold="true" box="[151,704,1670,1695]" pageId="16" pageNumber="319">
FIGURE 8. Adults of
<taxonomicName id="4C144D3D310EFF81FE3EBD91D24B27BE" authorityName="Nozaki" authorityYear="2025" box="[388,616,1670,1694]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="bicornis" status="sp. nov.">
<emphasis id="B960EAAC310EFF81FE3EBD91D24B27BE" bold="true" box="[388,616,1670,1694]" italics="true" pageId="16" pageNumber="319">Homoplectra bicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357D7310EFF81FDD5BD90D2E327BF" box="[623,704,1671,1695]" pageId="16" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral. Arrow in Fig. 8H see text.
</paragraph>
</caption>
<paragraph id="8BAB36BE310EFF81FF7DBC4AD52426BB" blockId="16.[151,1437,1885,2019]" pageId="16" pageNumber="319">
<emphasis id="B960EAAC310EFF81FF7DBC4AD2B52657" bold="true" box="[199,662,1885,1911]" pageId="16" pageNumber="319">Geographic variation of male genitalia</emphasis>
(
<figureCitation id="132F2A3B310EFF81FD1FBC4AD2D82657" box="[677,763,1885,1911]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8F</figureCitation>
). In males collected from
<collectingRegion id="49D0F85C310EFF81FB9BBC4AD53A2657" box="[1057,1305,1885,1911]" country="Japan" name="Yamanashi" pageId="16" pageNumber="319">Yamanashi Prefecture</collectingRegion>
, each horn-like posterolateral lobe of segment IX in dorsal aspect is apically round and directed mesad (
<figureCitation id="132F2A3B310EFF81FB1EBC96D4D926BB" box="[1188,1274,1921,1947]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE310EFF80FF7DBCB2D3DF20A9" blockId="16.[151,1437,1885,2019]" lastBlockId="17.[151,1437,151,1293]" lastPageId="17" lastPageNumber="320" pageId="16" pageNumber="319">
<emphasis id="B960EAAC310EFF81FF7DBCB2D1AF269F" bold="true" box="[199,396,1957,1983]" pageId="16" pageNumber="319">Female genitalia</emphasis>
(
<figureCitation id="132F2A3B310EFF81FE26BCB2D205269F" box="[412,550,1957,1983]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8G8H</figureCitation>
). Lateral lobes of sternum VIII cleft from base (
<figureCitation id="132F2A3B310EFF81FBFBBCB2D4AC269F" box="[1089,1167,1957,1983]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8I</figureCitation>
). Segment IX obliquely rectangular in lateral aspect (
<figureCitation id="132F2A3B310EFF81FE68BCDED20B26C3" box="[466,552,1993,2019]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8G</figureCitation>
), subpentagonal posteriorly in dorsal aspect (
<figureCitation id="132F2A3B310EFF81FBAFBCDED44E26C3" box="[1045,1133,1993,2019]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="16" pageNumber="319">Fig. 8H</figureCitation>
), tergum widely sclerotized ventrally; each mesal lobe sclerotized, bean-shaped in lateral aspect (
<figureCitation id="132F2A3B310FFF80FC22BB80D3D32191" box="[920,1008,151,177]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="17" pageNumber="320">Fig. 8G</figureCitation>
), forming large crevice between it and segment IX; with pair of sclerotized ribs on vulvar scale basolaterally in ventral aspect (
<figureCitation id="132F2A3B310FFF80FB28BBACD4C021F5" box="[1170,1251,187,213]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="17" pageNumber="320">Fig. 8I</figureCitation>
), weakly ridged near anteromesal margins. Vulvar scale large, membranous apically (
<figureCitation id="132F2A3B310FFF80FC17BBC8D3DE21D9" box="[941,1021,223,249]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="17" pageNumber="320">Fig. 8I</figureCitation>
). Segment X setose, subrectangular, longitudinally short in lateral aspect (
<figureCitation id="132F2A3B310FFF80FD85BA14D2B9203D" box="[575,666,259,285]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="17" pageNumber="320">Fig. 8G</figureCitation>
); ventral side sclerotized, with shape of large bottle in dorsal aspect (marked with an arrow in
<figureCitation id="132F2A3B310FFF80FE79BA30D23D2061" box="[451,542,295,321]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="17" pageNumber="320">Fig. 8H</figureCitation>
). Vaginal apparatus slender pentagonal in dorsal aspect (
<figureCitation id="132F2A3B310FFF80FB1FBA30D5232061" box="[1189,1280,295,321]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="17" pageNumber="320">Fig. 8H</figureCitation>
), with pair of falcate sclerites posterodorsally, with pair of dorsal longitudinal ridges, and with dark arch mark between ridges anterodorsally, mostly surrounding opening of duct of spermatheca (
<figureCitation id="132F2A3B310FFF80FC2FBA78D3CC20A9" box="[917,1007,367,393]" captionStart="FIGURE 8" captionStartId="16.[151,250,1670,1694]" captionTargetBox="[157,1430,459,1641]" captionTargetId="figure-158@16.[151,1435,453,1647]" captionTargetPageId="16" captionText="FIGURE 8. Adults of Homoplectra bicornis sp. nov. 8A8E, male genitalia, type locality (Shizuoka-shi, Shizuoka): 8A, left lateral, apicomesal part of inferior appendage enlarged; 8B, dorsal; 8C, ventral; 8D, phallic apparatus, left lateral; 8E, same, dorsal. 8F, posterodorsal part of male segment IX, variation (Minami-alps-shi, Yamanashi), dorsal. 8G8I, female genitalia, type locality: 8G, left lateral, 8H, dorsal; 8I, ventral.Arrow in Fig. 8H see text." pageId="17" pageNumber="320">Fig. 8H</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE310FFF80FF7DBA84D230208D" blockId="17.[151,1437,151,1293]" box="[199,531,403,429]" pageId="17" pageNumber="320">
<emphasis id="B960EAAC310FFF80FF7DBA84D1B5208D" bold="true" box="[199,406,403,429]" pageId="17" pageNumber="320">Immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E6535310FFF80FF7DBAA0D2BD2515" pageId="17" pageNumber="320" type="materials_examined">
<paragraph id="8BAB36BE310FFF80FF7DBAA0D26C20D5" blockId="17.[151,1437,151,1293]" pageId="17" pageNumber="320">
<materialsCitation id="3B7C3CE3310FFF80FF7DBAA0D26C20D5" collectingDate="2006-05-05" collectionCode="T" county="Male" elevation="400" latitude="35.227" location="Aoi-ku" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" pageId="17" pageNumber="320" specimenCode="KPM-NK 91358" specimenCount="1" stateProvince="Shizuoka" typeStatus="holotype">
<emphasis id="B960EAAC310FFF80FF7DBAA0D11820F1" bold="true" box="[199,315,439,465]" pageId="17" pageNumber="320">
<typeStatus id="54AF881C310FFF80FF7DBAA0D11520F1" box="[199,310,439,465]" pageId="17" pageNumber="320" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62CA4E32310FFF80FEFDBAA0D1A220F1" box="[327,385,439,465]" pageId="17" pageNumber="320">Male</collectingCounty>
(pinned):
<collectingMunicipality id="6BCFACC4310FFF80FE45BAA0D24720F1" box="[511,612,439,465]" pageId="17" pageNumber="320">Nyujima</collectingMunicipality>
,
<location id="8ECB6065310FFF80FDC8BAA0D2E720F1" LSID="urn:lsid:plazi:treatment:03BD87A83111FF80FF2DBFE5D2DA242D:8ECB6065310FFF80FDC8BAA0D2E720F1" box="[626,708,439,465]" county="Male" latitude="35.227" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" name="Aoi-ku" pageId="17" pageNumber="320" stateProvince="Shizuoka">Aoi-ku</location>
,
<location id="8ECB6065310FFF80FD6FBAA0D34920F1" LSID="urn:lsid:plazi:treatment:03BD87A83111FF80FF2DBFE5D2DA242D:8ECB6065310FFF80FD6FBAA0D34920F1" box="[725,874,439,465]" county="Male" latitude="35.227" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" name="Shizuoka-shi" pageId="17" pageNumber="320" stateProvince="Shizuoka">Shizuoka-shi</location>
,
<collectingRegion id="49D0F85C310FFF80FCC0BAA0D3C620F1" box="[890,997,439,465]" country="Japan" name="Shizuoka" pageId="17" pageNumber="320">Shizuoka</collectingRegion>
,
<geoCoordinate id="EE205079310FFF80FC4FBAA1D45120F1" box="[1013,1138,438,465]" degrees="35.2270" direction="north" orientation="latitude" pageId="17" pageNumber="320" precision="5" value="35.227">35.2270°N</geoCoordinate>
,
<geoCoordinate id="EE205079310FFF80FB38BAA1D52820F1" box="[1154,1291,438,465]" degrees="138.3456" direction="east" orientation="longitude" pageId="17" pageNumber="320" precision="5" value="138.3456">138.3456°E</geoCoordinate>
, alt.
<quantity id="4CEC9B5B310FFF80FAF3BAA0D5B420F1" box="[1353,1431,439,465]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="320" unit="m" value="400.0">
<elevation id="0039D18D310FFF80FAF3BAA0D5B420F1" box="[1353,1431,439,465]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="320" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="FFAA107E310FFF80FF2DBACDD0D920D5" box="[151,250,474,501]" pageId="17" pageNumber="320" value="2006-05-05">
<collectingDate id="EFEEE996310FFF80FF2DBACDD0D920D5" box="[151,250,474,501]" pageId="17" pageNumber="320" value="2006-05-05">5.v.2006</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FEBFBACCD13420D5" box="[261,279,475,501]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
. Nozaki (
<specimenCode id="DBB29EC5310FFF80FE39BACCD26720D5" box="[387,580,474,501]" collectionCode="KPM-NK" pageId="17" pageNumber="320">KPM-NK 91358</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAB36BE310FFF80FF7DBAE8D3ED2341" blockId="17.[151,1437,151,1293]" pageId="17" pageNumber="320">
<materialsCitation id="3B7C3CE3310FFF80FF7DBAE8D3CE2339" box="[199,1005,510,537]" collectingDate="2006-05-05" collectionCode="T" county="Male" elevation="400" latitude="35.227" location="Aoi-ku" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" pageId="17" pageNumber="320" specimenCode="KPM-NK 91359" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka" typeStatus="paratype">
<emphasis id="B960EAAC310FFF80FF7DBAE8D1652339" bold="true" box="[199,326,511,537]" pageId="17" pageNumber="320">
<typeStatus id="54AF881C310FFF80FF7DBAE8D1622339" box="[199,321,511,537]" pageId="17" pageNumber="320" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D12FD37310FFF80FEF5BAE8D1BF2339" box="[335,412,511,537]" count="1" pageId="17" pageNumber="320" type="male">1 male</specimenCount>
(pinned), same data as holotype (
<specimenCode id="DBB29EC5310FFF80FC99BAE8D3C62339" box="[803,997,510,537]" collectionCode="KPM-NK" pageId="17" pageNumber="320">KPM-NK 91359</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE3310FFF80FC40BAE8D230231D" collectingDate="1997-05-11" collectionCode="T" county="Male" elevation="400" latitude="35.227" location="Aoi-ku" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" pageId="17" pageNumber="320" specimenCount="9" specimenCount-female="1" specimenCount-male="8" stateProvince="Shizuoka" typeStatus="paratype">
<specimenCount id="9D12FD37310FFF80FC40BAE8D4762339" box="[1018,1109,511,537]" count="8" pageId="17" pageNumber="320" type="male">8 males</specimenCount>
,
<specimenCount id="9D12FD37310FFF80FBD9BAE8D4E52339" box="[1123,1222,511,537]" count="1" pageId="17" pageNumber="320" type="female">1 female</specimenCount>
(in alcohol), same locality as holotype,
<date id="FFAA107E310FFF80FE3FB934D1D7231D" box="[389,500,547,573]" pageId="17" pageNumber="320" value="1997-05-11">
<collectingDate id="EFEEE996310FFF80FE3FB934D1D7231D" box="[389,500,547,573]" pageId="17" pageNumber="320" value="1997-05-11">11.v.1997</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FDBBB934D230231D" box="[513,531,547,573]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FDA5B934D474231D" box="[543,1111,546,573]" collectionCode="KPM-NK" pageId="17" pageNumber="320" specimenCode="TN00640, KPM-NKTN00650" specimenCount="1" stateProvince="Shizuoka" typeStatus="paratype">
Hattori (
<collectionCode id="ED05AE7B310FFF80FD3AB934D2CD231D" box="[640,750,547,573]" pageId="17" pageNumber="320">KPM-NK</collectionCode>
<specimenCode id="DBB29EC5310FFF80FD4CB934D346231D" box="[758,869,547,573]" collectionCode="TN" pageId="17" pageNumber="320">TN00640</specimenCode>
,
<specimenCode id="DBB29EC5310FFF80FCCBB935D473231D" box="[881,1104,546,573]" collectionCode="KPM-NKTN" pageId="17" pageNumber="320">KPM-NKTN00650</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE3310FFF80FBDEB934D2622341" collectingDate="2002-05-03" collectionCode="T" county="Male" elevation="400" latitude="35.227" location="Aoi-ku" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" pageId="17" pageNumber="320" specimenCount="7" specimenCount-female="1" specimenCount-male="6" stateProvince="Shizuoka" typeStatus="paratype">
<specimenCount id="9D12FD37310FFF80FBDEB934D49C231D" box="[1124,1215,547,573]" count="6" pageId="17" pageNumber="320" type="male">6 males</specimenCount>
,
<specimenCount id="9D12FD37310FFF80FB71B934D50D231D" box="[1227,1326,547,573]" count="1" pageId="17" pageNumber="320" type="female">1 female</specimenCount>
(pinned), same locality as holotype,
<date id="FFAA107E310FFF80FE7BB950D2072341" box="[449,548,583,609]" pageId="17" pageNumber="320" value="2002-05-03">
<collectingDate id="EFEEE996310FFF80FE7BB950D2072341" box="[449,548,583,609]" pageId="17" pageNumber="320" value="2002-05-03">3.v.2002</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FD95B950D2622341" box="[559,577,583,609]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
, Hattori (SPMN-IS-5980459810).
</paragraph>
<paragraph id="8BAB36BE310FFF80FF7DB97DD2BD2515" blockId="17.[151,1437,151,1293]" pageId="17" pageNumber="320">
<materialsCitation id="3B7C3CE3310FFF80FF7DB97DD20D2389" collectingDate="1997-07-06" collectionCode="T" collectorName="Heisei-kyo &amp; Takao &amp; Minami-alpsshi" country="Japan" elevation="1300" location="Yamanashi" pageId="17" pageNumber="320" specimenCount="4" specimenCount-male="4" stateProvince="Yamanashi">
<emphasis id="B960EAAC310FFF80FF7DB97DD38823A4" bold="true" box="[199,939,618,644]" pageId="17" pageNumber="320">
<typeStatus id="54AF881C310FFF80FF7DB97DD23123A4" box="[199,530,618,644]" isOtherTypeStatus="true" pageId="17" pageNumber="320">Other specimens examined</typeStatus>
.
<collectingCountry id="F303762E310FFF80FD98B97DD2A323A4" box="[546,640,618,644]" name="Japan" pageId="17" pageNumber="320">JAPAN</collectingCountry>
: HONSHU:
<collectingRegion id="49D0F85C310FFF80FCA7B97DD38523A4" box="[797,934,618,644]" country="Japan" name="Yamanashi" pageId="17" pageNumber="320">Yamanashi</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD37310FFF80FC0FB97CD43123A5" box="[949,1042,619,645]" count="4" pageId="17" pageNumber="320" type="male">4 males</specimenCount>
,
<collectorName id="26E15368310FFF80FB98B97CD48323A5" box="[1058,1184,619,645]" pageId="17" pageNumber="320">Heisei-kyo</collectorName>
,
<collectorName id="26E15368310FFF80FB14B97CD4D623A5" box="[1198,1269,619,645]" pageId="17" pageNumber="320">Takao</collectorName>
,
<collectorName id="26E15368310FFF80FABEB97CD0992389" pageId="17" pageNumber="320">Minami-alpsshi</collectorName>
, alt.
<quantity id="4CEC9B5B310FFF80FF48B998D1B32389" box="[242,400,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="17" pageNumber="320" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">
<elevation id="0039D18D310FFF80FF48B998D1B32389" box="[242,400,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="17" pageNumber="320" unit="m" value="1300.0" valueMax="1400.0" valueMin="1200.0">12001400 m</elevation>
</quantity>
,
<date id="FFAA107E310FFF80FE26B998D2332389" box="[412,528,655,681]" pageId="17" pageNumber="320" value="1997-07-06">
<collectingDate id="EFEEE996310FFF80FE26B998D2332389" box="[412,528,655,681]" pageId="17" pageNumber="320" value="1997-07-06">6.vii.1997</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FDA6B998D20D2389" box="[540,558,655,681]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FD80B998D5622389" box="[570,1345,655,681]" collectingDate="1996-06-09" collectionCode="SPMN, T" collectorName="Hattori &amp; Fujikawa-cho" country="Japan" location="Oyana-gawa" pageId="17" pageNumber="320" specimenCount="1" specimenCount-female="1">
<collectorName id="26E15368310FFF80FD80B998D2AA2389" box="[570,649,655,681]" pageId="17" pageNumber="320">Hattori</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FD20B998D2CE2389" box="[666,749,655,681]" pageId="17" pageNumber="320">SPMN</collectionCode>
);
<specimenCount id="9D12FD37310FFF80FCBAB998D3462389" box="[768,869,655,681]" count="1" pageId="17" pageNumber="320" type="female">1 female</specimenCount>
,
<location id="8ECB6065310FFF80FCCBB998D4222389" LSID="urn:lsid:plazi:treatment:03BD87A83111FF80FF2DBFE5D2DA242D:8ECB6065310FFF80FCCBB998D4222389" box="[881,1025,655,681]" country="Japan" name="Oyana-gawa" pageId="17" pageNumber="320">Oyana-gawa</location>
,
<collectorName id="26E15368310FFF80FBB4B998D4882389" box="[1038,1195,655,681]" pageId="17" pageNumber="320">Fujikawa-cho</collectorName>
,
<date id="FFAA107E310FFF80FB0DB998D5002389" box="[1207,1315,655,681]" pageId="17" pageNumber="320" value="1996-06-09">
<collectingDate id="EFEEE996310FFF80FB0DB998D5002389" box="[1207,1315,655,681]" pageId="17" pageNumber="320" value="1996-06-09">9.vi.1996</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FA95B998D5622389" box="[1327,1345,655,681]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FAF7B998D3C823ED" collectingDate="1997-05-05" collectionCode="SPMN, T" collectorName="Hattori" country="Japan" county="Male" elevation="400" latitude="35.227" location="Aoi-ku" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" pageId="17" pageNumber="320" specimenCount="9" specimenCount-female="1" specimenCount-male="8" stateProvince="Shizuoka">
<collectorName id="26E15368310FFF80FAF7B998D5BF2389" box="[1357,1436,655,681]" pageId="17" pageNumber="320">Hattori</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FF25B9A4D0D123ED" box="[159,242,691,717]" pageId="17" pageNumber="320">SPMN</collectionCode>
).
<emphasis id="B960EAAC310FFF80FEB8B9A5D15923EC" bold="true" box="[258,378,690,716]" pageId="17" pageNumber="320">
<collectingRegion id="49D0F85C310FFF80FEB8B9A5D15623EC" box="[258,373,690,716]" country="Japan" name="Shizuoka" pageId="17" pageNumber="320">Shizuoka</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD37310FFF80FEC5B9A4D1F523ED" box="[383,470,691,717]" count="8" pageId="17" pageNumber="320" type="male">8 males</specimenCount>
,
<specimenCount id="9D12FD37310FFF80FE5AB9A4D26123ED" box="[480,578,691,717]" count="1" pageId="17" pageNumber="320" type="female">1 female</specimenCount>
, same locality as holotype,
<date id="FFAA107E310FFF80FCD7B9A5D3F323ED" box="[877,976,690,717]" pageId="17" pageNumber="320" value="1997-05-05">
<collectingDate id="EFEEE996310FFF80FCD7B9A5D3F323ED" box="[877,976,690,717]" pageId="17" pageNumber="320" value="1997-05-05">5.v.1997</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FC63B9A4D3C823ED" box="[985,1003,691,717]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FC4EB9A4D18323D1" collectingDate="1995-05-05" collectionCode="SPMN, T" collectorName="Hattori" country="Japan" county="Male" elevation="400" latitude="35.227" location="Aoi-ku" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" pageId="17" pageNumber="320" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka">
<collectorName id="26E15368310FFF80FC4EB9A4D46023ED" box="[1012,1091,691,717]" pageId="17" pageNumber="320">Hattori</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FBEAB9A4D48023ED" box="[1104,1187,691,717]" pageId="17" pageNumber="320">SPMN</collectionCode>
);
<specimenCount id="9D12FD37310FFF80FB0EB9A4D52323ED" box="[1204,1280,691,717]" count="1" pageId="17" pageNumber="320" type="male">1 male</specimenCount>
, same locality as holotype,
<date id="FFAA107E310FFF80FE98B9C1D1A623D1" box="[290,389,726,753]" pageId="17" pageNumber="320" value="1995-05-05">
<collectingDate id="EFEEE996310FFF80FE98B9C1D1A623D1" box="[290,389,726,753]" pageId="17" pageNumber="320" value="1995-05-05">5.v.1995</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FE34B9C0D18323D1" box="[398,416,727,753]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FE10B9C0D4C523D1" box="[426,1254,727,753]" collectingDate="1997-05-11" collectionCode="SPMN, T" collectorName="Hattori" country="Japan" county="Male" elevation="400" latitude="35.227" location="Aoi-ku" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" pageId="17" pageNumber="320" specimenCount="8" specimenCount-female="1" specimenCount-male="7" stateProvince="Shizuoka">
<collectorName id="26E15368310FFF80FE10B9C0D1DA23D1" box="[426,505,727,753]" pageId="17" pageNumber="320">Hattori</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FDBCB9C0D27A23D1" box="[518,601,727,753]" pageId="17" pageNumber="320">SPMN</collectionCode>
);
<specimenCount id="9D12FD37310FFF80FDD0B9C0D2E123D1" box="[618,706,727,753]" count="7" pageId="17" pageNumber="320" type="male">7 males</specimenCount>
,
<specimenCount id="9D12FD37310FFF80FD71B9C0D30D23D1" box="[715,814,727,753]" count="1" pageId="17" pageNumber="320" type="female">1 female</specimenCount>
, same locality as holotype,
<date id="FFAA107E310FFF80FBE1B9C0D4E923D1" box="[1115,1226,727,753]" pageId="17" pageNumber="320" value="1997-05-11">
<collectingDate id="EFEEE996310FFF80FBE1B9C0D4E923D1" box="[1115,1226,727,753]" pageId="17" pageNumber="320" value="1997-05-11">11.v.1997</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FB6EB9C0D4C523D1" box="[1236,1254,727,753]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FB55B9C0D26B2235" collectingDate="2006-05-09" collectionCode="T" collectorName="Hattori" country="Japan" county="Male" elevation="400" latitude="35.227" location="Aoi-ku" longLatPrecision="6" longitude="138.3456" municipality="Nyujima" pageId="17" pageNumber="320" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka">
<collectorName id="26E15368310FFF80FB55B9C0D56123D1" box="[1263,1346,727,753]" pageId="17" pageNumber="320">Hattori</collectorName>
;
<specimenCount id="9D12FD37310FFF80FAF1B9C0D5BB23D1" box="[1355,1432,727,753]" count="1" pageId="17" pageNumber="320" type="male">1 male</specimenCount>
, same locality as holotype,
<date id="FFAA107E310FFF80FE7DB9ECD2092235" box="[455,554,763,789]" pageId="17" pageNumber="320" value="2006-05-09">
<collectingDate id="EFEEE996310FFF80FE7DB9ECD2092235" box="[455,554,763,789]" pageId="17" pageNumber="320" value="2006-05-09">9.v.2006</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FD8CB9ECD26B2235" box="[566,584,763,789]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FDEEB9ECD41E2219" collectingDate="1990-05-06" collectionCode="TT, SPMN, T" collectorName="Torii &amp; Utogi &amp; Aoi-ku &amp; Shizuoka-shi" country="Japan" elevation="700" location="Aoi-ku" municipality="Kamikouchi-zawa" pageId="17" pageNumber="320" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<collectorName id="26E15368310FFF80FDEEB9ECD2A92235" box="[596,650,763,789]" pageId="17" pageNumber="320">Torii</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FD21B9ECD2E12235" box="[667,706,763,789]" pageId="17" pageNumber="320">TT</collectionCode>
);
<specimenCount id="9D12FD37310FFF80FD62B9ECD3102235" box="[728,819,763,789]" count="2" pageId="17" pageNumber="320" type="male">2 males</specimenCount>
,
<specimenCount id="9D12FD37310FFF80FC85B9ECD3862235" box="[831,933,763,789]" count="1" pageId="17" pageNumber="320" type="female">1 female</specimenCount>
,
<collectorName id="26E15368310FFF80FC08B9ECD3D62235" box="[946,1013,763,789]" pageId="17" pageNumber="320">Utogi</collectorName>
, alt. ca.
<quantity id="4CEC9B5B310FFF80FBEFB9ECD4BC2235" box="[1109,1183,763,789]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="320" unit="m" value="700.0">
<elevation id="0039D18D310FFF80FBEFB9ECD4BC2235" box="[1109,1183,763,789]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="320" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<collectorName id="26E15368310FFF80FB11B9ECD4DE2235" box="[1195,1277,763,789]" pageId="17" pageNumber="320">Aoi-ku</collectorName>
,
<collectorName id="26E15368310FFF80FAB0B9ECD5BF2235" box="[1290,1436,763,789]" pageId="17" pageNumber="320">Shizuoka-shi</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FF25B808D0D12219" box="[159,242,799,825]" pageId="17" pageNumber="320">SPMN</collectionCode>
);
<specimenCount id="9D12FD37310FFF80FEBEB808D1712219" box="[260,338,799,825]" count="1" pageId="17" pageNumber="320" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6BCFACC4310FFF80FEE7B808D20E2219" box="[349,557,799,825]" pageId="17" pageNumber="320">Kamikouchi-zawa</collectingMunicipality>
, alt.
<quantity id="4CEC9B5B310FFF80FDDBB808D2942219" box="[609,695,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="320" unit="m" value="1000.0">
<elevation id="0039D18D310FFF80FDDBB808D2942219" box="[609,695,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="320" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<location id="8ECB6065310FFF80FD7BB808D3302219" LSID="urn:lsid:plazi:treatment:03BD87A83111FF80FF2DBFE5D2DA242D:8ECB6065310FFF80FD7BB808D3302219" box="[705,787,799,825]" country="Japan" municipality="Kamikouchi-zawa" name="Aoi-ku" pageId="17" pageNumber="320">Aoi-ku</location>
,
<collectorName id="26E15368310FFF80FCA4B808D3902219" box="[798,947,799,825]" pageId="17" pageNumber="320">Shizuoka-shi</collectorName>
,
<date id="FFAA107E310FFF80FC04B808D4032219" box="[958,1056,799,825]" pageId="17" pageNumber="320" value="1990-05-06">
<collectingDate id="EFEEE996310FFF80FC04B808D4032219" box="[958,1056,799,825]" pageId="17" pageNumber="320" value="1990-05-06">6.v.1990</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FB91B808D41E2219" box="[1067,1085,799,825]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FBFDB808D1B0227D" collectingDate="1990-06-20" collectionCode="SPMN, T" collectorName="Hattori" country="Japan" elevation="700" location="Aoi-ku" municipality="Kamikouchi-zawa" pageId="17" pageNumber="320" specimenCount="1" specimenCount-male="1">
<collectorName id="26E15368310FFF80FBFDB808D4B52219" box="[1095,1174,799,825]" pageId="17" pageNumber="320">Hattori</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FB1FB808D4DB2219" box="[1189,1272,799,825]" pageId="17" pageNumber="320">SPMN</collectionCode>
);
<specimenCount id="9D12FD37310FFF80FAB0B808D57B2219" box="[1290,1368,799,825]" count="1" pageId="17" pageNumber="320" type="male">1 male</specimenCount>
, same locality,
<date id="FFAA107E310FFF80FF43B854D157227D" box="[249,372,835,861]" pageId="17" pageNumber="320" value="1990-06-20">
<collectingDate id="EFEEE996310FFF80FF43B854D157227D" box="[249,372,835,861]" pageId="17" pageNumber="320" value="1990-06-20">20.vi.1990</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FE3BB854D1B0227D" box="[385,403,835,861]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FE1AB854D3D6227D" box="[416,1013,835,861]" collectingDate="1991-06-09" collectionCode="SPMN, T" collectorName="Hattori" country="Japan" elevation="700" location="Aoi-ku" municipality="Kamikouchi-zawa" pageId="17" pageNumber="320" specimenCount="1" specimenCount-male="1">
<collectorName id="26E15368310FFF80FE1AB854D1CC227D" box="[416,495,835,861]" pageId="17" pageNumber="320">Hattori</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FDBAB854D270227D" box="[512,595,835,861]" pageId="17" pageNumber="320">SPMN</collectionCode>
);
<specimenCount id="9D12FD37310FFF80FDD2B854D29B227D" box="[616,696,835,861]" count="1" pageId="17" pageNumber="320" type="male">1 male</specimenCount>
, same locality,
<date id="FFAA107E310FFF80FCD0B854D3F6227D" box="[874,981,835,861]" pageId="17" pageNumber="320" value="1991-06-09">
<collectingDate id="EFEEE996310FFF80FCD0B854D3F6227D" box="[874,981,835,861]" pageId="17" pageNumber="320" value="1991-06-09">9.vi.1991</collectingDate>
</date>
,
<collectionCode id="ED05AE7B310FFF80FC59B854D3D6227D" box="[995,1013,835,861]" name="Tavera, Department of Geology and Geophysics" pageId="17" pageNumber="320">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3310FFF80FBB8B854D28722A1" collectingDate="1989-06-18" collectionCode="SPMN, T" collectorName="Hattori &amp; Aoi-ku &amp; Shizuoka-shi" country="Japan" location="Hatanagi" pageId="17" pageNumber="320" specimenCount="8" specimenCount-female="1" specimenCount-male="7">
<collectorName id="26E15368310FFF80FBB8B854D472227D" box="[1026,1105,835,861]" pageId="17" pageNumber="320">Hattori</collectorName>
(
<collectionCode id="ED05AE7B310FFF80FBD8B854D496227D" box="[1122,1205,835,861]" pageId="17" pageNumber="320">SPMN</collectionCode>
);
<specimenCount id="9D12FD37310FFF80FB70B854D506227D" box="[1226,1317,835,861]" count="7" pageId="17" pageNumber="320" type="male">7 males</specimenCount>
,
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</paragraph>
</subSubSection>
<subSubSection id="C30E6535310FFF80FF7DBF28D2FD255D" pageId="17" pageNumber="320" type="etymology">
<paragraph id="8BAB36BE310FFF80FF7DBF28D2FD255D" blockId="17.[151,1437,151,1293]" pageId="17" pageNumber="320">
<emphasis id="B960EAAC310FFF80FF7DBF28D16D2579" bold="true" box="[199,334,1087,1113]" pageId="17" pageNumber="320">Etymology.</emphasis>
The species epithet (Latin adjective, two-horned) refers to the pair of horn-like posterolateral projections of the male segment IX in dorsal aspect.
</paragraph>
</subSubSection>
<subSubSection id="C30E6535310FFF80FF7DBF91D2032581" box="[199,544,1158,1185]" pageId="17" pageNumber="320" type="distribution">
<paragraph id="8BAB36BE310FFF80FF7DBF91D2032581" blockId="17.[151,1437,151,1293]" box="[199,544,1158,1185]" pageId="17" pageNumber="320">
<emphasis id="B960EAAC310FFF80FF7DBF91D1422580" bold="true" box="[199,353,1158,1184]" pageId="17" pageNumber="320">Distribution.</emphasis>
Central Honshu.
</paragraph>
</subSubSection>
<subSubSection id="C30E6535310FFF80FF7DBFBDD2DA242D" pageId="17" pageNumber="320" type="biology_ecology">
<paragraph id="8BAB36BE310FFF80FF7DBFBDD14E25C9" blockId="17.[151,1437,151,1293]" pageId="17" pageNumber="320">
<emphasis id="B960EAAC310FFF80FF7DBFBDD10425E4" bold="true" box="[199,295,1194,1220]" pageId="17" pageNumber="320">Biology.</emphasis>
The
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and a
<typeStatus id="54AF881C310FFF80FDB6BFBCD24F25E5" box="[524,620,1195,1221]" pageId="17" pageNumber="320" type="paratype">paratype</typeStatus>
male of this species were flying in the day time around a small waterfall at the type locality.
</paragraph>
<paragraph id="8BAB36BE310FFF80FF7DBFE4D2DA242D" blockId="17.[151,1437,151,1293]" box="[199,761,1267,1293]" pageId="17" pageNumber="320">
<emphasis id="B960EAAC310FFF80FF7DBFE4D1A0242D" bold="true" box="[199,387,1267,1293]" pageId="17" pageNumber="320">Japanese name.</emphasis>
Tsuno-kimadara-shima-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,844 @@
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<mods:classification id="AAF4E8BA5FD2086DF242EDAFA75E367C">journal article</mods:classification>
<mods:identifier id="4A318E767B4315E762615F6427B13D9A" type="DOI">10.11646/zootaxa.5566.2.4</mods:identifier>
<mods:identifier id="7510B5D475DC42A67824C9C475F68984" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BD87A83113FF9EFF2DB935D3292581" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581" httpUri="http://treatment.plazi.org/id/03BD87A83113FF9EFF2DB935D3292581" lastPageId="15" lastPageNumber="318" pageId="13" pageNumber="316">
<subSubSection id="C30E65353113FF9CFF2DB935D221231D" box="[151,514,546,573]" pageId="13" pageNumber="316" type="nomenclature">
<paragraph id="8BAB36BE3113FF9CFF2DB935D221231D" blockId="13.[151,514,546,609]" box="[151,514,546,573]" pageId="13" pageNumber="316">
<heading id="D0E381D23113FF9CFF2DB935D221231D" bold="true" box="[151,514,546,573]" fontSize="11" level="1" pageId="13" pageNumber="316" reason="1">
<emphasis id="B960EAAC3113FF9CFF2DB935D221231D" bold="true" box="[151,514,546,573]" pageId="13" pageNumber="316">
<taxonomicName id="4C144D3D3113FF9CFF2DB935D1BE231D" authorityName="Nozaki" authorityYear="2025" box="[151,413,546,573]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="shikoku" status="sp. nov.">
<emphasis id="B960EAAC3113FF9CFF2DB935D1BE231D" bold="true" box="[151,413,546,573]" italics="true" pageId="13" pageNumber="316">Homoplectra shikoku</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357D73113FF9CFE1EB934D221231D" box="[420,514,547,573]" pageId="13" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E65353113FF9CFF2DB950D13A2341" box="[151,281,583,609]" pageId="13" pageNumber="316" type="description">
<paragraph id="8BAB36BE3113FF9CFF2DB950D13A2341" blockId="13.[151,514,546,609]" box="[151,281,583,609]" pageId="13" pageNumber="316">
(
<figureCitation id="132F2A3B3113FF9CFF25B950D0CB2341" box="[159,232,583,609]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Figs 7</figureCitation>
,
<figureCitation id="132F2A3B3113FF9CFF4EB950D1332341" box="[244,272,583,609]" captionStart="FIGURE 13" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[169,1415,195,1904]" captionTargetId="figure-33@22.[157,1430,181,1919]" captionTargetPageId="22" captionText="FIGURE 13. Distributions of species of the Homoplectra japonica Species Group in Japan." pageId="13" pageNumber="316">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E65353113FF9CFF2DB99CD57B23E2" pageId="13" pageNumber="316" type="reference_group">
<paragraph id="8BAB36BE3113FF9CFF2DB99CD33C2383" blockId="13.[151,1368,651,706]" box="[151,799,651,675]" pageId="13" pageNumber="316">
<treatmentCitationGroup id="AB0411903113FF9CFF2DB99CD33C2383" box="[151,799,651,675]" pageId="13" pageNumber="316">
<taxonomicName id="4C144D3D3113FF9CFF18B99CD1A52383" ID-CoL="36JDB" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[162,390,651,675]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3113FF9CFF18B99CD1382383" box="[162,283,651,675]" italics="true" pageId="13" pageNumber="316">Diplectrona</emphasis>
<emphasis id="B960EAAC3113FF9CFE97B99BD1A52383" box="[301,390,652,675]" italics="true" pageId="13" pageNumber="316">japonica</emphasis>
</taxonomicName>
Complex:
<treatmentCitation id="0AB510AF3113FF9CFE4CB99BD3382383" author="Yamamoto, E. &amp; Ito, T." box="[502,795,651,675]" page="13" pageId="13" pageNumber="316" year="2014">
<bibRefCitation id="EF854B4F3113FF9CFE4CB99BD2FF2383" author="Yamamoto, E. &amp; Ito, T." box="[502,732,651,675]" pageId="13" pageNumber="316" pagination="6 - 21" refId="ref18649" refString="Yamamoto, E. &amp; Ito, T. (2014) Caddisflies (Trichoptera) collected from Izugataniyama, Kumakogen-cho, Shikoku, Japan in 2012 - 2013. Shikokukogera, 14, 6 - 21 [in Japanese]." type="journal article" year="2014">Yamamoto &amp; Ito 2014</bibRefCitation>
, p. 13
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAB36BE3113FF9CFF2DB9BDD57B23E2" blockId="13.[151,1368,651,706]" box="[151,1368,682,706]" pageId="13" pageNumber="316">
Diplectroninae Gen.
<taxonomicName id="4C144D3D3113FF9CFED1B9BDD1E723E1" ID-CoL="36JDB" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[363,452,682,705]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3113FF9CFED1B9BDD1E723E1" box="[363,452,682,705]" italics="true" pageId="13" pageNumber="316">japonica</emphasis>
</taxonomicName>
Species Complex:
<bibRefCitation id="EF854B4F3113FF9CFD32B9BDD32F23E2" author="Nozaki, T." box="[648,780,682,706]" pageId="13" pageNumber="316" pagination="456 - 461" refId="ref17951" refString="Nozaki, T. (2016) Trichoptera. In: Maruyama, H. &amp; Hanada, S. (Eds), A Field Guide to Japanese Aquatic Insects: Adults of Mayflies, Stoneflies and Caddisflies, Zenkoku Noson Kyoiku Kyokai, Tokyo, pp. 69 - 87, 294 - 410, 429 - 442, 449 - 451, 456 - 461 [in Japanese]." type="book chapter" year="2016">Nozaki 2016</bibRefCitation>
, pp 75, 348, female (photo);
<bibRefCitation id="EF854B4F3113FF9CFB8BB9BDD53723E2" author="Yamamoto, E. &amp; Nozaki, T. &amp; Kuhara, N. &amp; Ito, T." box="[1073,1300,682,706]" pageId="13" pageNumber="316" pagination="67 - 76" refId="ref18691" refString="Yamamoto, E., Nozaki, T., Kuhara, N. &amp; Ito, T. (2021) Caddisflies (Trichoptera) collected from a mountain stream, Namakusa-dani, in Shikoku, western Japan. In: Yamamoto Institute of Forest Biology (Ed.), Odamiyama no suisei-konchu to Matsubara- izumi no shizen. Ecosystem Conservation Society of Ehime, Uchiko-cho, Uchiko-cho, Ehime, pp. 67 - 76." type="book chapter" year="2021">
Yamamoto
<emphasis id="B960EAAC3113FF9CFB1CB9BCD4F123E2" box="[1190,1234,682,706]" italics="true" pageId="13" pageNumber="316">et al</emphasis>
. 2021
</bibRefCitation>
, p. 73.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353113FF9CFF2DB9E7D102229E" pageId="13" pageNumber="316" type="diagnosis">
<paragraph id="8BAB36BE3113FF9CFF2DB9E7D5502272" blockId="13.[151,1437,752,958]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF2DB9E7D131222A" bold="true" box="[151,274,752,778]" pageId="13" pageNumber="316">Diagnosis.</emphasis>
The male of this species is somewhat similar to those of
<taxonomicName id="4C144D3D3113FF9CFC18B9E6D404222A" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[930,1063,753,778]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3113FF9CFC18B9E6D404222A" box="[930,1063,753,778]" italics="true" pageId="13" pageNumber="316">H. japonica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C144D3D3113FF9CFBDAB9E6D538222A" authorityName="Nozaki" authorityYear="2025" box="[1120,1307,752,778]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="flagelliformis" status="sp. nov.">
<emphasis id="B960EAAC3113FF9CFBDAB9E6D538222A" box="[1120,1307,752,778]" italics="true" pageId="13" pageNumber="316">H. flagelliformis</emphasis>
</taxonomicName>
<emphasis id="B960EAAC3113FF9CFA99B9E7D55D222A" bold="true" box="[1315,1406,752,778]" pageId="13" pageNumber="316">
<taxonomicNameLabel id="A25357D73113FF9CFA99B9E7D55D222A" box="[1315,1406,752,778]" pageId="13" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having a short antennal scape and in the shape of the phallic apparatus; but is easily distinguishable from those two species by bearing long inner processes of the genital chamber and posterior processes of segment X (
<figureCitation id="132F2A3B3113FF9CFAB7B82FD5452272" box="[1293,1382,824,850]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE3113FF9CFF7DB84BD102229E" blockId="13.[151,1437,752,958]" pageId="13" pageNumber="316">
The female of this species is similar to that of
<taxonomicName id="4C144D3D3113FF9CFD7EB84AD35E2256" authorityName="Nozaki" authorityYear="2025" box="[708,893,860,886]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="flagelliformis" status="sp. nov.">
<emphasis id="B960EAAC3113FF9CFD7EB84AD35E2256" box="[708,893,860,886]" italics="true" pageId="13" pageNumber="316">H. flagelliformis</emphasis>
</taxonomicName>
<emphasis id="B960EAAC3113FF9CFC39B84BD3F82256" bold="true" box="[899,987,860,886]" pageId="13" pageNumber="316">
<taxonomicNameLabel id="A25357D73113FF9CFC39B84BD3F82256" box="[899,987,860,886]" pageId="13" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having a trapezoidal basoventral lobe on the vulvar scale, but they can be distinguished from each other by the character states given in the diagnosis for that species.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353113FF9CFF2DBF07D23026D6" pageId="13" pageNumber="316" type="description">
<paragraph id="8BAB36BE3113FF9CFF2DBF07D10B250A" blockId="13.[151,296,1040,1066]" box="[151,296,1040,1066]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF2DBF07D10B250A" bold="true" box="[151,296,1040,1066]" pageId="13" pageNumber="316">Description</emphasis>
</paragraph>
<paragraph id="8BAB36BE3113FF9CFF2DBF4FD250259A" blockId="13.[151,1437,1112,2038]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF2DBF4FD0F82552" bold="true" box="[151,219,1112,1138]" pageId="13" pageNumber="316">Adult</emphasis>
(
<figureCitation id="132F2A3B3113FF9CFF53BF4FD1612552" box="[233,322,1112,1138]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7A</figureCitation>
). General coloration of body and wings, and length of antennal scapes similar to those of
<taxonomicName id="4C144D3D3113FF9CFAA9BF4ED5B62552" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1299,1429,1113,1138]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3113FF9CFAA9BF4ED5B62552" box="[1299,1429,1113,1138]" italics="true" pageId="13" pageNumber="316">H. japonica</emphasis>
</taxonomicName>
. Yellow markings on female wings rarely lacking. Forewings each 9.011.0 mm in male (n = 10), 9.813.0 mm in female (n = 10),
<quantity id="4CEC9B5B3113FF9CFEE9BFB7D18A259B" box="[339,425,1184,1211]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="13" pageNumber="316" unit="mm" value="9.5">9.5 mm</quantity>
in male
<typeStatus id="54AF881C3113FF9CFDB0BFB7D253259A" box="[522,624,1184,1210]" pageId="13" pageNumber="316" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BAB36BE3113FF9CFF7DBFD3D35F2792" blockId="13.[151,1437,1112,2038]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF7DBFD3D15025FE" bold="true" box="[199,371,1220,1246]" pageId="13" pageNumber="316">Male genitalia</emphasis>
(
<figureCitation id="132F2A3B3113FF9CFE38BFD3D23225FE" box="[386,529,1220,1246]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Figs 7C7H</figureCitation>
). Ventrolateral sides of synsclerotized segment IX subtriangular in lateral aspect (
<figureCitation id="132F2A3B3113FF9CFF25BFFFD0DA2422" box="[159,249,1256,1282]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7C</figureCitation>
); posteroventral lobe trapezoidal in ventral aspect (
<figureCitation id="132F2A3B3113FF9CFCFABFFFD3BB2422" box="[832,920,1256,1282]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7E</figureCitation>
), usually with central shallow notch apically; dorsal part of segment IX triangular with round apex in dorsal aspect (
<figureCitation id="132F2A3B3113FF9CFC76BE1BD40A2406" box="[972,1065,1292,1318]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7D</figureCitation>
), broadly fused with segment X laterally (
<figureCitation id="132F2A3B3113FF9CFEBFBE27D143246A" box="[261,352,1328,1354]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7C</figureCitation>
). Segment X bilobed in dorsal aspect (
<figureCitation id="132F2A3B3113FF9CFC9EBE27D3A2246A" box="[804,897,1328,1354]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7D</figureCitation>
), pair of posterolateral setose areas thumb-like in lateral aspect (
<figureCitation id="132F2A3B3113FF9CFEE0BE43D197244E" box="[346,436,1364,1390]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7C</figureCitation>
); each with ventrolateral margin sclerotized; segment X with pair of posterior processes (p.p.X) having variable length, even in same locality (
<figureCitation id="132F2A3B3113FF9CFCB3BE6FD39524B2" box="[777,950,1400,1426]" captionStart-0="FIGURE 7" captionStart-1="FIGURES 12" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="14.[151,250,1696,1720]" captionStartId-1="3.[151,264,1626,1650]" captionStartId-2="4.[151,250,1613,1637]" captionStartId-3="7.[151,250,1588,1612]" captionStartId-4="8.[151,250,1567,1591]" captionStartId-5="11.[151,250,1872,1896]" captionTargetBox-0="[157,1429,187,1666]" captionTargetBox-1="[157,1429,351,1596]" captionTargetBox-2="[157,1429,185,1583]" captionTargetBox-3="[157,1429,187,1558]" captionTargetBox-4="[166,1431,191,1536]" captionTargetBox-5="[165,1420,498,1834]" captionTargetId-0="figure-33@14.[151,1435,181,1672]" captionTargetId-1="figure-102@3.[151,1435,345,1602]" captionTargetId-2="figure-33@4.[151,1435,181,1589]" captionTargetId-3="figure-19@7.[151,1435,181,1564]" captionTargetId-4="figure-33@8.[151,1435,181,1543]" captionTargetId-5="figure-95@11.[151,1435,491,1849]" captionTargetPageId-0="14" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="11" captionText-0="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." captionText-1="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." captionText-2="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." captionText-3="FIGURE 4. Homoplectra albomarginata (Ulmer 1907). 4A4F: male genitalia (Sakauchi-sakamoto, Ibigawa-cho, Gifu): 4A, left lateral, apicomesal part of inferior appendage enlarged; 4B, dorsal; 4C, ventral; 4D, inner process, lateral strip and posterior processe of segment X, left lateral, variation; 4E14E2, phallotheca, left lateral, variations; 4F, aedeagus, left lateral. 4G4J, female genitalia (Sakauchi-sakamoto, Ibigawa-cho, Gifu): 4G, left lateral; 4H, dorsal; 4I, ventral; 4J14J3, lobes formed by sclerotized ribs on vulvar scale, variations, ventral. Arrows on Figs 4H and 4I see text." captionText-4="FIGURE 5. Homoplectra albomarginata (Ulmer 1907). Variations of male and female genitalia. 5A: male genitalia, left lateral, inferior appendage omitted: 5A1, Ozu, Ibigawa-cho, Gifu; 5A2, Tsurumi, Ibigawa-cho, Gifu. 5B5F: male and female genitalia: 5B, Fudo-daki, Tsukechi-cho, Nakatsugawa-shi, Gifu; 5C, Ookaya-gawa, Kamiooka, Shimada-shi, Shizuoka; 5D, Mennoki-toge, Tsugu, Shitara-cho, Aichi; 5E, Miyazuma-kyo, Suizawa-cho, Yokkaichi-shi, Mie; 5F, Ombara, Kamisaibara, Kagamino-cho, Okayama. Abbreviations: D = male segments IX &amp; X, dorsal; L = same, right lateral; ph.a = phallic apparatus, left lateral; p.l. = posteroventral lobe of segment IX, ventral; v.s. = vulvar scale, ventral." captionText-5="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="13" pageNumber="316">Figs 7H17H3</figureCitation>
), with few apical spine-like setae. Genital chamber with pair of weakly sclerotized lateral strips (l.s.); with pair of long spine-like inner processes (i.p.) evenly curved ventrad (
<figureCitation id="132F2A3B3113FF9CFEEABED7D18924FA" box="[336,426,1472,1498]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7C</figureCitation>
). Inferior appendages each without distal segment, long finger-like in lateral aspect (
<figureCitation id="132F2A3B3113FF9CFAD5BED7D09824DE" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7C</figureCitation>
), extending beyond apex of segment X, weakly bent upward in apical 1/3; subapicomesal apex weakly bulging mesally and with more than 10 spine-like setae (
<figureCitation id="132F2A3B3113FF9CFD01BD1FD3362702" box="[699,789,1544,1570]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Figs 7C</figureCitation>
inset, 7D, 7E). Phallic apparatus arcuate in lateral aspect (
<figureCitation id="132F2A3B3113FF9CFF25BD3BD0D12766" box="[159,242,1580,1606]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7F</figureCitation>
), dorsal process of phallotheca acicular in lateral and dorsal aspects (
<figureCitation id="132F2A3B3113FF9CFC64BD3BD4432766" box="[990,1120,1580,1607]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Figs 7F, 7G</figureCitation>
), gently curved ventrad (
<figureCitation id="132F2A3B3113FF9CFAD5BD3BD09B274A" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7F</figureCitation>
); pair of ventral processes of phallotheca approximately 2/3 as long as dorsal process (
<figureCitation id="132F2A3B3113FF9CFBC7BD47D4F1274A" box="[1149,1234,1616,1642]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7F</figureCitation>
), but occasionally only slightly shorter than dorsal process. Aedeagal stem thick, half-pipe-like (
<figureCitation id="132F2A3B3113FF9CFC46BD63D47627AE" box="[1020,1109,1652,1678]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7G</figureCitation>
), moderately curved ventrad, with long head, approximately 1/4 as long as basal stem (
<figureCitation id="132F2A3B3113FF9CFCA3BD8FD34C2792" box="[793,879,1688,1714]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7F</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE3113FF9CFF7DBDABD10826F2" blockId="13.[151,1437,1112,2038]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF7DBDABD1B027F6" bold="true" box="[199,403,1724,1750]" pageId="13" pageNumber="316">Female genitalia</emphasis>
(
<figureCitation id="132F2A3B3113FF9CFE13BDABD21327F7" box="[425,560,1724,1751]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Figs 7I7K</figureCitation>
). Lateral lobes of sternum VIII cleft from base, mesal margins convex, posteromesal margin evenly curved laterad in ventral aspect (
<figureCitation id="132F2A3B3113FF9CFCDEBDF7D39C27DB" box="[868,959,1760,1787]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7K</figureCitation>
). Segment IX subrectangular with round posterolateral corners in dorsal aspect (
<figureCitation id="132F2A3B3113FF9CFDE9BC13D286263E" box="[595,677,1796,1822]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7J</figureCitation>
), obliquely S-shaped in lateral aspect (
<figureCitation id="132F2A3B3113FF9CFBE4BC13D48D263E" box="[1118,1198,1796,1822]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7I</figureCitation>
); pair of mesal lobes sclerotized, semicircular in lateral aspect (
<figureCitation id="132F2A3B3113FF9CFDD7BC3FD2982662" box="[621,699,1832,1858]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7I</figureCitation>
); pair of sclerotized ribs forming wide trapezoidal lobes on vulvar scale basoventrally (
<figureCitation id="132F2A3B3113FF9CFEC5BC5BD1F42647" box="[383,471,1868,1895]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7K</figureCitation>
). Vulvar scale tongue-like in ventral aspect, membranous apically (
<figureCitation id="132F2A3B3113FF9CFB6DBC5BD50D2647" box="[1239,1326,1868,1895]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7K</figureCitation>
). Vaginal apparatus long triangular in dorsal aspect (
<figureCitation id="132F2A3B3113FF9CFD36BC67D2C226AA" box="[652,737,1904,1930]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="13" pageNumber="316">Fig. 7J</figureCitation>
); with pair of sclerotized round bulges posterolaterally, and with dark M-shaped mark between pair of longitudinal ridges anteriorly, partially surrounding opening of duct of spermatheca.
</paragraph>
<paragraph id="8BAB36BE3113FF9CFF7DBCCBD23026D6" blockId="13.[151,1437,1112,2038]" box="[199,531,2012,2038]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF7DBCCBD1B526D6" bold="true" box="[199,406,2012,2038]" pageId="13" pageNumber="316">Immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="DF6B66363110FF9FFF2DBDB7D44B2668" pageId="14" pageNumber="317" startId="14.[151,250,1696,1720]" targetBox="[157,1429,187,1666]" targetPageId="14" targetType="figure">
<paragraph id="8BAB36BE3110FF9FFF2DBDB7D44B2668" blockId="14.[151,1437,1695,1864]" pageId="14" pageNumber="317">
<emphasis id="B960EAAC3110FF9FFF2DBDB7D2992798" bold="true" box="[151,698,1695,1720]" pageId="14" pageNumber="317">
FIGURE 7. Adults of
<taxonomicName id="4C144D3D3110FF9FFE3BBD88D2402797" authorityName="Nozaki" authorityYear="2025" box="[385,611,1695,1719]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="species" species="shikoku" status="sp. nov.">
<emphasis id="B960EAAC3110FF9FFE3BBD88D2402797" bold="true" box="[385,611,1695,1719]" italics="true" pageId="14" pageNumber="317">Homoplectra shikoku</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357D73110FF9FFDD0BDB7D2992798" box="[618,698,1696,1720]" pageId="14" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral.
</paragraph>
</caption>
<subSubSection id="C30E65353110FF9EFF7DBC8DD5792285" lastPageId="15" lastPageNumber="318" pageId="14" pageNumber="317" type="materials_examined">
<paragraph id="8BAB36BE3110FF9FFF7DBC8DD5BE2694" blockId="14.[151,1437,1946,2009]" box="[199,1437,1946,1973]" pageId="14" pageNumber="317">
<materialsCitation id="3B7C3CE33110FF9FFF7DBC8DD5BE2694" box="[199,1437,1946,1973]" county="Male" latitude="33.5628" location="Nishidani" longLatPrecision="6" longitude="132.9358" municipality="Izugataniyama" pageId="14" pageNumber="317" specimenCount="1" stateProvince="Ehime" typeStatus="holotype">
<emphasis id="B960EAAC3110FF9FFF7DBC8DD1182694" bold="true" box="[199,315,1946,1972]" pageId="14" pageNumber="317">
<typeStatus id="54AF881C3110FF9FFF7DBC8DD1152694" box="[199,310,1946,1972]" pageId="14" pageNumber="317" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62CA4E323110FF9FFEF8BC8DD15F2694" box="[322,380,1946,1972]" pageId="14" pageNumber="317">Male</collectingCounty>
(in alcohol):
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,
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,
<location id="8ECB60653110FF9FFC85BC8DD3DE2694" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653110FF9FFC85BC8DD3DE2694" box="[831,1021,1946,1972]" county="Male" latitude="33.5628" longLatPrecision="6" longitude="132.9358" municipality="Izugataniyama" name="Kumakogen-cho" pageId="14" pageNumber="317" stateProvince="Ehime">Kumakogen-cho</location>
,
<collectingRegion id="49D0F85C3110FF9FFBBDBC8DD4702694" box="[1031,1107,1946,1972]" country="Japan" name="Ehime" pageId="14" pageNumber="317">Ehime</collectingRegion>
,
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,
<geoCoordinate id="EE2050793110FF9FFB5CBC8DD54D2695" box="[1254,1390,1946,1973]" degrees="132.9358" direction="east" orientation="longitude" pageId="14" pageNumber="317" precision="5" value="132.9358">132.9358°E</geoCoordinate>
, alt.
</materialsCitation>
</paragraph>
<paragraph id="8BAB36BE3110FF9FFF2DBCA9D32C26F8" blockId="14.[151,1437,1946,2009]" box="[151,783,1982,2009]" pageId="14" pageNumber="317">
<materialsCitation id="3B7C3CE33110FF9FFF2DBCA9D32F26F8" box="[151,780,1982,2009]" collectingDate="2020-05-20" collectionCode="KPM-NK" collectorName="E. Yamamoto" elevation="1370" pageId="14" pageNumber="317" specimenCode="TN00610" specimenCount="1">
<quantity id="4CEC9B5B3110FF9FFF2DBCA9D0CD26F9" box="[151,238,1982,2009]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="14" pageNumber="317" unit="m" value="1370.0">
<elevation id="0039D18D3110FF9FFF2DBCA9D0CD26F9" box="[151,238,1982,2009]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="14" pageNumber="317" unit="m" value="1370.0">1370 m</elevation>
</quantity>
,
<date id="FFAA107E3110FF9FFF40BCA9D14926F8" box="[250,362,1982,2008]" pageId="14" pageNumber="317" value="2020-05-20">
<collectingDate id="EFEEE9963110FF9FFF40BCA9D14926F8" box="[250,362,1982,2008]" pageId="14" pageNumber="317" value="2020-05-20">20.v.2020</collectingDate>
</date>
,
<collectorName id="26E153683110FF9FFECCBCA8D22D26F9" box="[374,526,1983,2009]" pageId="14" pageNumber="317">E. Yamamoto</collectorName>
(
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<specimenCode id="DBB29EC53110FF9FFD29BCA9D32726F8" box="[659,772,1982,2008]" collectionCode="TN" pageId="14" pageNumber="317">TN00610</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAB36BE3111FF9EFF7DBB80D0CD21D9" blockId="15.[151,1437,150,1185]" pageId="15" pageNumber="318">
<materialsCitation id="3B7C3CE33111FF9EFF7DBB80D5142191" box="[199,1335,150,177]" collectionCode="KPM-NK" county="Male" latitude="33.5628" location="Nishidani" longLatPrecision="6" longitude="132.9358" municipality="Izugataniyama" pageId="15" pageNumber="318" specimenCode="TN00620, TN00630" specimenCount="23" specimenCount-female="8" specimenCount-male="15" stateProvince="Ehime" typeStatus="paratype">
<emphasis id="B960EAAC3111FF9EFF7DBB80D1652191" bold="true" box="[199,326,151,177]" pageId="15" pageNumber="318">
<typeStatus id="54AF881C3111FF9EFF7DBB80D1622191" box="[199,321,151,177]" pageId="15" pageNumber="318" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D12FD373111FF9EFEF5BB81D19B2191" box="[335,440,150,177]" count="15" pageId="15" pageNumber="318" type="male">15 males</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFE7CBB80D2142191" box="[454,567,151,177]" count="8" pageId="15" pageNumber="318" type="female">8 females</specimenCount>
, same data as holotype (
<collectionCode id="ED05AE7B3111FF9EFCEFBB80D3E02191" box="[853,963,151,177]" pageId="15" pageNumber="318">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53111FF9EFC76BB80D4192191" box="[972,1082,151,177]" collectionCode="TN" pageId="15" pageNumber="318">TN00620</specimenCode>
,
<collectionCode id="ED05AE7B3111FF9EFBFDBB80D4962191" box="[1095,1205,151,177]" pageId="15" pageNumber="318">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53111FF9EFB05BB80D50D2191" box="[1215,1326,151,177]" collectionCode="TN" pageId="15" pageNumber="318">TN00630</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B7C3CE33111FF9EFAFEBB80D3FC21F5" collectingDate="2011-05-25" collectionCode="T" elevation="1310" location="Kumakogen-cho" municipality="Nishidani" pageId="15" pageNumber="318" specimenCount="2" specimenCount-male="2" stateProvince="Ehime" typeStatus="paratype">
<specimenCount id="9D12FD373111FF9EFAFEBB80D5BF2191" box="[1348,1436,151,177]" count="2" pageId="15" pageNumber="318" type="male">2 males</specimenCount>
(pinned),
<collectingMunicipality id="6BCFACC43111FF9EFEB3BBACD15A21F5" box="[265,377,187,213]" pageId="15" pageNumber="318">Nishidani</collectingMunicipality>
,
<location id="8ECB60653111FF9EFE33BBACD26421F5" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFE33BBACD26421F5" box="[393,583,187,213]" municipality="Nishidani" name="Kumakogen-cho" pageId="15" pageNumber="318" stateProvince="Ehime">Kumakogen-cho</location>
,
<collectingRegion id="49D0F85C3111FF9EFDECBBACD28121F5" box="[598,674,187,213]" country="Japan" name="Ehime" pageId="15" pageNumber="318">Ehime</collectingRegion>
, alt.
<quantity id="4CEC9B5B3111FF9EFD5BBBACD31E21F5" box="[737,829,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="15" pageNumber="318" unit="m" value="1310.0">
<elevation id="0039D18D3111FF9EFD5BBBACD31E21F5" box="[737,829,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="15" pageNumber="318" unit="m" value="1310.0">1310 m</elevation>
</quantity>
,
<date id="FFAA107E3111FF9EFCF4BBADD39E21F5" box="[846,957,186,213]" pageId="15" pageNumber="318" value="2011-05-25">
<collectingDate id="EFEEE9963111FF9EFCF4BBADD39E21F5" box="[846,957,186,213]" pageId="15" pageNumber="318" value="2011-05-25">25.v.2011</collectingDate>
</date>
,
<collectionCode id="ED05AE7B3111FF9EFC77BBACD3FC21F5" box="[973,991,187,213]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
. Hattori (SPMN-IS-59802, SPMN-IS-59803).
</paragraph>
<paragraph id="8BAB36BE3111FF9EFF7DBA15D5792285" blockId="15.[151,1437,150,1185]" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF7DBA15D3B6203C" bold="true" box="[199,917,258,284]" pageId="15" pageNumber="318">
<materialsCitation id="3B7C3CE33111FF9EFF7DBA15D3B3203C" box="[199,912,258,284]" country="Japan" location="Tokushima" pageId="15" pageNumber="318" specimenCount="1" stateProvince="Tokushima">
<typeStatus id="54AF881C3111FF9EFF7DBA15D227203C" box="[199,516,258,284]" isOtherTypeStatus="true" pageId="15" pageNumber="318">Other specimens examined</typeStatus>
.
<collectingCountry id="F303762E3111FF9EFDB7BA15D248203C" box="[525,619,258,284]" name="Japan" pageId="15" pageNumber="318">JAPAN</collectingCountry>
: SHIKOKU:
<collectingRegion id="49D0F85C3111FF9EFCBDBA15D3B3203C" box="[775,912,258,284]" country="Japan" name="Tokushima" pageId="15" pageNumber="318">Tokushima</collectingRegion>
</materialsCitation>
:
</emphasis>
<materialsCitation id="3B7C3CE33111FF9EFC23BA14D1BA2061" collectingDate="2009-05-06" collectionCode="T" country="Japan" county="Konose-kyo" location="Naka-cho" municipality="Kitokitagawa" pageId="15" pageNumber="318" specimenCount="11" specimenCount-female="5" specimenCount-male="6">
<specimenCount id="9D12FD373111FF9EFC23BA14D3CC203D" box="[921,1007,259,285]" count="6" pageId="15" pageNumber="318" type="male">6 males</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFC4DBA15D440203D" box="[1015,1123,258,285]" count="5" pageId="15" pageNumber="318" type="female">5 females</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFBD1BA14D4D4203D" box="[1131,1271,259,285]" pageId="15" pageNumber="318">Konose-kyo</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFB44BA14D5BB203D" box="[1278,1432,259,285]" pageId="15" pageNumber="318">Kitokitagawa</collectingMunicipality>
,
<location id="8ECB60653111FF9EFF2DBA30D1242061" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFF2DBA30D1242061" box="[151,263,295,321]" country="Japan" county="Konose-kyo" municipality="Kitokitagawa" name="Naka-cho" pageId="15" pageNumber="318">Naka-cho</location>
,
<date id="FFAA107E3111FF9EFEACBA30D15A2061" box="[278,377,295,321]" pageId="15" pageNumber="318" value="2009-05-06">
<collectingDate id="EFEEE9963111FF9EFEACBA30D15A2061" box="[278,377,295,321]" pageId="15" pageNumber="318" value="2009-05-06">6.v.2009</collectingDate>
</date>
,
<collectionCode id="ED05AE7B3111FF9EFE3DBA30D1BA2061" box="[391,409,295,321]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFE1DBA30D2692061" box="[423,586,295,321]" collectionCode="SPMN" country="Japan" location="Torii" pageId="15" pageNumber="318" specimenCount="1">
<location id="8ECB60653111FF9EFE1DBA30D1FE2061" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFE1DBA30D1FE2061" box="[423,477,295,321]" country="Japan" name="Torii" pageId="15" pageNumber="318">Torii</location>
(
<collectionCode id="ED05AE7B3111FF9EFE55BA30D2612061" box="[495,578,295,321]" pageId="15" pageNumber="318">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFDE3BA30D5BB2061" box="[601,1432,295,321]" collectingDate="2011-06-08" collectionCode="T" country="Japan" elevation="1355" location="Miyoshi-shi" municipality="Higashiiya-sugeoi" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFDE3BA30D2892061" box="[601,682,295,321]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6BCFACC43111FF9EFD03BA30D3AB2061" box="[697,904,295,321]" pageId="15" pageNumber="318">Higashiiya-sugeoi</collectingMunicipality>
,
<location id="8ECB60653111FF9EFC2CBA30D4032061" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFC2CBA30D4032061" box="[918,1056,295,321]" country="Japan" municipality="Higashiiya-sugeoi" name="Miyoshi-shi" pageId="15" pageNumber="318">Miyoshi-shi</location>
, alt.
<quantity id="4CEC9B5B3111FF9EFBE7BA30D4DE2061" box="[1117,1277,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.355" metricValueMax="1.38" metricValueMin="1.33" pageId="15" pageNumber="318" unit="m" value="1355.0" valueMax="1380.0" valueMin="1330.0">
<elevation id="0039D18D3111FF9EFBE7BA30D4DE2061" box="[1117,1277,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.355" metricValueMax="1.38" metricValueMin="1.33" pageId="15" pageNumber="318" unit="m" value="1355.0" valueMax="1380.0" valueMin="1330.0">13301380 m</elevation>
</quantity>
,
<date id="FFAA107E3111FF9EFAB6BA30D55B2061" box="[1292,1400,295,321]" pageId="15" pageNumber="318" value="2011-06-08">
<collectingDate id="EFEEE9963111FF9EFAB6BA30D55B2061" box="[1292,1400,295,321]" pageId="15" pageNumber="318" value="2011-06-08">8.vi.2011</collectingDate>
</date>
,
<collectionCode id="ED05AE7B3111FF9EFA3CBA30D5BB2061" box="[1414,1432,295,321]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFF2DBA5CD1832044" box="[151,416,330,357]" collectionCode="KPM" country="Japan" location="Nozaki" pageId="15" pageNumber="318" specimenCount="1" stateProvince="Ehime">
<location id="8ECB60653111FF9EFF2DBA5CD0CB2045" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFF2DBA5CD0CB2045" box="[151,232,331,357]" country="Japan" name="Nozaki" pageId="15" pageNumber="318" stateProvince="Ehime">Nozaki</location>
(
<collectionCode id="ED05AE7B3111FF9EFF43BA5CD1182045" box="[249,315,331,357]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
).
<collectingRegion id="49D0F85C3111FF9EFEF4BA5DD1832044" box="[334,416,330,356]" country="Japan" name="Ehime" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFEF4BA5DD1832044" bold="true" box="[334,416,330,356]" pageId="15" pageNumber="318">Ehime</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="B960EAAC3111FF9EFE1ABA5DD1862044" bold="true" box="[416,421,330,356]" pageId="15" pageNumber="318">:</emphasis>
<specimenCount id="9D12FD373111FF9EFE14BA5CD22B2045" box="[430,520,331,357]" count="7" pageId="15" pageNumber="318" type="male">7 males</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFDAEBA5CD2A72045" box="[532,644,331,357]" count="4" pageId="15" pageNumber="318" type="female">4 females</specimenCount>
,
<typeStatus id="54AF881C3111FF9EFD2ABA5CD2E32045" box="[656,704,331,357]" pageId="15" pageNumber="318">type</typeStatus>
locality,
<date id="FFAA107E3111FF9EFC93BA5CD3C22045" box="[809,993,331,357]" pageId="15" pageNumber="318" value="2013-05-11" valueMax="2013-06-10" valueMin="2013-05-11">11.v10.vi.2013</date>
, E. Yamamoto;
<specimenCount id="9D12FD373111FF9EFB2DBA5CD4DC2045" box="[1175,1279,331,357]" count="10" pageId="15" pageNumber="318" type="male">10 males</specimenCount>
,
<typeStatus id="54AF881C3111FF9EFAB1BA5CD5182045" box="[1291,1339,331,357]" pageId="15" pageNumber="318">type</typeStatus>
locality,
<date id="FFAA107E3111FF9EFF2DBA78D11120A9" box="[151,306,367,393]" pageId="15" pageNumber="318" value="2018-05-21" valueMax="2018-05-31" valueMin="2018-05-21">2131.v.2018</date>
, E. Yamamoto;
<materialsCitation id="3B7C3CE33111FF9EFE4BBA78D4FC20A9" box="[497,1247,367,393]" collectingDate="2007-05-10" collectorName="E. Yamamoto" country="Japan" location="Kumakogen-cho" municipality="Yurano-mori" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFE4BBA78D26020A9" box="[497,579,367,393]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6BCFACC43111FF9EFDEBBA78D2C020A9" box="[593,739,367,393]" pageId="15" pageNumber="318">Yurano-mori</collectingMunicipality>
,
<location id="8ECB60653111FF9EFD49BA78D39220A9" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFD49BA78D39220A9" box="[755,945,367,393]" country="Japan" municipality="Yurano-mori" name="Kumakogen-cho" pageId="15" pageNumber="318">Kumakogen-cho</location>
,
<date id="FFAA107E3111FF9EFC7ABA78D41320A9" box="[960,1072,367,393]" pageId="15" pageNumber="318" value="2007-05-10">
<collectingDate id="EFEEE9963111FF9EFC7ABA78D41320A9" box="[960,1072,367,393]" pageId="15" pageNumber="318" value="2007-05-10">10.v.2007</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFBFABA78D4FC20A9" box="[1088,1247,367,393]" pageId="15" pageNumber="318">E. Yamamoto</collectorName>
</materialsCitation>
;
<specimenCount id="9D12FD373111FF9EFB55BA78D56F20A9" box="[1263,1356,367,393]" count="4" pageId="15" pageNumber="318" type="male">4 males</specimenCount>
, same,
<date id="FFAA107E3111FF9EFF2DBA84D124208D" box="[151,263,403,429]" pageId="15" pageNumber="318" value="2007-05-21">21.v.2007</date>
, E. Yamamoto;
<materialsCitation id="3B7C3CE33111FF9EFE01BA84D4B5208D" box="[443,1174,403,429]" collectingDate="1996-07-20" collectionCode="T" country="Japan" location="Kumakogen-cho" municipality="Tsuchigoya" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1" stateProvince="Wakayama">
<specimenCount id="9D12FD373111FF9EFE01BA84D229208D" box="[443,522,403,429]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<collectingMunicipality id="6BCFACC43111FF9EFDAFBA84D2BA208D" box="[533,665,403,429]" pageId="15" pageNumber="318">Tsuchigoya</collectingMunicipality>
,
<collectingRegion id="49D0F85C3111FF9EFD1EBA84D300208D" box="[676,803,403,429]" country="Japan" name="Wakayama" pageId="15" pageNumber="318">Wakayama</collectingRegion>
,
<location id="8ECB60653111FF9EFC94BA84D3CF208D" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFC94BA84D3CF208D" box="[814,1004,403,429]" country="Japan" municipality="Tsuchigoya" name="Kumakogen-cho" pageId="15" pageNumber="318" stateProvince="Wakayama">Kumakogen-cho</location>
,
<date id="FFAA107E3111FF9EFC4DBA84D45A208D" box="[1015,1145,403,429]" pageId="15" pageNumber="318" value="1996-07-20">
<collectingDate id="EFEEE9963111FF9EFC4DBA84D45A208D" box="[1015,1145,403,429]" pageId="15" pageNumber="318" value="1996-07-20">20.vii.1996</collectingDate>
</date>
,
<collectionCode id="ED05AE7B3111FF9EFB3EBA84D4B5208D" box="[1156,1174,403,429]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFB18BA84D510208D" box="[1186,1331,403,429]" collectionCode="KPM" country="Japan" location="Befu" pageId="15" pageNumber="318" specimenCount="1">
<location id="8ECB60653111FF9EFB18BA84D4FB208D" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFB18BA84D4FB208D" box="[1186,1240,403,429]" country="Japan" name="Befu" pageId="15" pageNumber="318">Befu</location>
(
<collectionCode id="ED05AE7B3111FF9EFB53BA84D508208D" box="[1257,1323,403,429]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFA84BA84D30220F1" collectingDate="1999-05-21" collectionCode="T" collectorName="A. Ohkawa" country="Japan" location="Kumakogen-cho" pageId="15" pageNumber="318" specimenCount="2" specimenCount-male="2" stateProvince="Wakayama">
<specimenCount id="9D12FD373111FF9EFA84BA84D5BB208D" box="[1342,1432,403,429]" count="2" pageId="15" pageNumber="318" type="male">2 males</specimenCount>
,
<collectingRegion id="49D0F85C3111FF9EFF2DBAA0D13520F1" box="[151,278,439,465]" country="Japan" name="Wakayama" pageId="15" pageNumber="318">Wakayama</collectingRegion>
,
<location id="8ECB60653111FF9EFE98BAA0D1C320F1" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFE98BAA0D1C320F1" box="[290,480,439,465]" country="Japan" name="Kumakogen-cho" pageId="15" pageNumber="318" stateProvince="Wakayama">Kumakogen-cho</location>
,
<date id="FFAA107E3111FF9EFE56BAA0D27F20F1" box="[492,604,439,465]" pageId="15" pageNumber="318" value="1999-05-21">
<collectingDate id="EFEEE9963111FF9EFE56BAA0D27F20F1" box="[492,604,439,465]" pageId="15" pageNumber="318" value="1999-05-21">21.v.1999</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFDD2BAA0D2CA20F1" box="[616,745,439,465]" pageId="15" pageNumber="318">A. Ohkawa</collectorName>
&amp;
<collectionCode id="ED05AE7B3111FF9EFCB5BAA0D30220F1" box="[783,801,439,465]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFC94BAA0D37320F1" box="[814,848,439,465]" country="Japan" location="Ito" pageId="15" pageNumber="318" specimenCount="1">
<location id="8ECB60653111FF9EFC94BAA0D37320F1" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFC94BAA0D37320F1" box="[814,848,439,465]" country="Japan" name="Ito" pageId="15" pageNumber="318">Ito</location>
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFCE7BAA0D19220D5" collectingDate="2011-06-09" collectionCode="KPM" collectorName="K. Nio" country="Japan" location="Kumakogen-cho" municipality="Banjo-dani" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1" stateProvince="Wakayama">
<specimenCount id="9D12FD373111FF9EFCE7BAA0D39420F1" box="[861,951,439,465]" count="1" pageId="15" pageNumber="318" type="male">1 males</specimenCount>
,
<collectingMunicipality id="6BCFACC43111FF9EFC7EBAA0D46020F1" box="[964,1091,439,465]" pageId="15" pageNumber="318">Banjo-dani</collectingMunicipality>
,
<collectingRegion id="49D0F85C3111FF9EFBEABAA0D4EC20F1" box="[1104,1231,439,465]" country="Japan" name="Wakayama" pageId="15" pageNumber="318">Wakayama</collectingRegion>
,
<location id="8ECB60653111FF9EFB61BAA0D5BA20F1" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFB61BAA0D5BA20F1" box="[1243,1433,439,465]" country="Japan" municipality="Banjo-dani" name="Kumakogen-cho" pageId="15" pageNumber="318" stateProvince="Wakayama">Kumakogen-cho</location>
,
<date id="FFAA107E3111FF9EFF2DBACCD12120D5" box="[151,258,475,501]" pageId="15" pageNumber="318" value="2011-06-09">
<collectingDate id="EFEEE9963111FF9EFF2DBACCD12120D5" box="[151,258,475,501]" pageId="15" pageNumber="318" value="2011-06-09">9.vi.2011</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFEB7BACCD17B20D5" box="[269,344,475,501]" pageId="15" pageNumber="318">K. Nio</collectorName>
(
<collectionCode id="ED05AE7B3111FF9EFEDDBACCD18A20D5" box="[359,425,475,501]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFE01BACCD5A720D5" box="[443,1412,474,501]" collectingDate="2013-06-06" collectingDateMax="2013-06-10" collectingDateMin="2013-06-06" collectorName="E. Yamamoto" country="Japan" elevation="1500" location="Uchiko-cho" municipality="Mt. Kasatori" pageId="15" pageNumber="318" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="9D12FD373111FF9EFE01BACCD23020D5" box="[443,531,475,501]" count="2" pageId="15" pageNumber="318" type="male">2 males</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFDA7BACCD2A820D5" box="[541,651,475,501]" count="2" pageId="15" pageNumber="318" type="female">2 females</specimenCount>
,
<collectingMunicipality id="6BCFACC43111FF9EFD2FBACCD30620D5" box="[661,805,475,501]" pageId="15" pageNumber="318">Mt. Kasatori</collectingMunicipality>
,
<location id="8ECB60653111FF9EFC95BACCD39520D5" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFC95BACCD39520D5" box="[815,950,475,501]" country="Japan" municipality="Mt. Kasatori" name="Uchiko-cho" pageId="15" pageNumber="318">Uchiko-cho</location>
, alt.
<quantity id="4CEC9B5B3111FF9EFC53BACDD41C20D5" box="[1001,1087,474,501]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="318" unit="m" value="1500.0">
<elevation id="0039D18D3111FF9EFC53BACDD41C20D5" box="[1001,1087,474,501]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="318" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FFAA107E3111FF9EFBF3BACCD4C320D5" box="[1097,1248,475,501]" pageId="15" pageNumber="318" value="2013-06-06" valueMax="2013-06-10" valueMin="2013-06-06">
<collectingDate id="EFEEE9963111FF9EFBF3BACCD4C320D5" box="[1097,1248,475,501]" pageId="15" pageNumber="318" value="2013-06-06" valueMax="2013-06-10" valueMin="2013-06-06">610.vi.2013</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFB50BACCD5A720D5" box="[1258,1412,475,501]" pageId="15" pageNumber="318">E. Yamamoto</collectorName>
</materialsCitation>
;
<specimenCount id="9D12FD373111FF9EFA34BACCD0F32339" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
, same,
<date id="FFAA107E3111FF9EFE98BAE8D1E52339" box="[290,454,511,537]" pageId="15" pageNumber="318" value="2020-06-16" valueMax="2020-06-20" valueMin="2020-06-16">1620.vi.2020</date>
, E. Yamamoto;
<materialsCitation id="3B7C3CE33111FF9EFDCCBAE8D5162339" box="[630,1333,511,537]" collectingDate="1992-05-08" collectionCode="KPM" collectorName="E. Yamamoto" country="Japan" location="Uchiko-cho" municipality="Oda-cho" pageId="15" pageNumber="318" specimenCount="4" specimenCount-male="4">
<specimenCount id="9D12FD373111FF9EFDCCBAE8D2ED2339" box="[630,718,511,537]" count="4" pageId="15" pageNumber="318" type="male">4 males</specimenCount>
,
<collectingMunicipality id="6BCFACC43111FF9EFD63BAE8D31F2339" box="[729,828,511,537]" pageId="15" pageNumber="318">Oda-cho</collectingMunicipality>
,
<location id="8ECB60653111FF9EFCFCBAE8D3EE2339" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFCFCBAE8D3EE2339" box="[838,973,511,537]" country="Japan" municipality="Oda-cho" name="Uchiko-cho" pageId="15" pageNumber="318">Uchiko-cho</location>
,
<date id="FFAA107E3111FF9EFC62BAE8D4182339" box="[984,1083,511,537]" pageId="15" pageNumber="318" value="1992-05-08">
<collectingDate id="EFEEE9963111FF9EFC62BAE8D4182339" box="[984,1083,511,537]" pageId="15" pageNumber="318" value="1992-05-08">8.v.1992</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFBFFBAE8D4FF2339" box="[1093,1244,511,537]" pageId="15" pageNumber="318">E. Yamamoto</collectorName>
(
<collectionCode id="ED05AE7B3111FF9EFB56BAE8D50D2339" box="[1260,1326,511,537]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFAFABAE8D301231D" collectingDate="1986-05-04" collectionCode="KPM" collectorName="E. Yamamaoto" country="Japan" location="Uchiko-cho" municipality="Odamiyama" pageId="15" pageNumber="318" specimenCount="4" specimenCount-male="4">
<specimenCount id="9D12FD373111FF9EFAFABAE8D5BB2339" box="[1344,1432,511,537]" count="4" pageId="15" pageNumber="318" type="male">4 males</specimenCount>
,
<collectingMunicipality id="6BCFACC43111FF9EFF2DB934D100231D" box="[151,291,547,573]" pageId="15" pageNumber="318">Odamiyama</collectingMunicipality>
,
<location id="8ECB60653111FF9EFE96B934D190231D" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFE96B934D190231D" box="[300,435,547,573]" country="Japan" municipality="Odamiyama" name="Uchiko-cho" pageId="15" pageNumber="318">Uchiko-cho</location>
,
<date id="FFAA107E3111FF9EFE06B934D23C231D" box="[444,543,547,573]" pageId="15" pageNumber="318" value="1986-05-04">
<collectingDate id="EFEEE9963111FF9EFE06B934D23C231D" box="[444,543,547,573]" pageId="15" pageNumber="318" value="1986-05-04">4.v.1986</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFD92B934D2E9231D" box="[552,714,547,573]" pageId="15" pageNumber="318">E.Yamamaoto</collectorName>
(
<collectionCode id="ED05AE7B3111FF9EFD63B934D338231D" box="[729,795,547,573]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFC96B934D5BB231D" box="[812,1432,547,573]" collectingDate="1990-04-28" collectionCode="KPM" collectorName="Yamamoto" country="Japan" location="Uchiko-cho" municipality="Odamiyama" pageId="15" pageNumber="318" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D12FD373111FF9EFC96B934D3A0231D" box="[812,899,547,573]" count="2" pageId="15" pageNumber="318" type="male">2 males</specimenCount>
, same locality,
<date id="FFAA107E3111FF9EFB93B934D482231D" box="[1065,1185,547,573]" pageId="15" pageNumber="318" value="1990-04-28">
<collectingDate id="EFEEE9963111FF9EFB93B934D482231D" box="[1065,1185,547,573]" pageId="15" pageNumber="318" value="1990-04-28">28.iv.1990</collectingDate>
</date>
, E.
<collectorName id="26E153683111FF9EFB7DB934D562231D" box="[1223,1345,547,573]" pageId="15" pageNumber="318">Yamamoto</collectorName>
(
<collectionCode id="ED05AE7B3111FF9EFAF4B934D5B3231D" box="[1358,1424,547,573]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFF2DB950D3242341" box="[151,775,583,609]" collectingDate="1991-05-20" collectionCode="KPM" collectorName="Yamamoto" country="Japan" location="Uchiko-cho" municipality="Odamiyama" pageId="15" pageNumber="318" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D12FD373111FF9EFF2DB950D0D32341" box="[151,240,583,609]" count="2" pageId="15" pageNumber="318" type="male">2 males</specimenCount>
, same locality,
<date id="FFAA107E3111FF9EFE20B950D2282341" box="[410,523,583,609]" pageId="15" pageNumber="318" value="1991-05-20">
<collectingDate id="EFEEE9963111FF9EFE20B950D2282341" box="[410,523,583,609]" pageId="15" pageNumber="318" value="1991-05-20">20.v.1991</collectingDate>
</date>
, E.
<collectorName id="26E153683111FF9EFD89B950D28E2341" box="[563,685,583,609]" pageId="15" pageNumber="318">Yamamoto</collectorName>
(
<collectionCode id="ED05AE7B3111FF9EFD07B950D2DC2341" box="[701,767,583,609]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFCABB950D5552341" box="[785,1398,583,609]" collectingDate="2002-05-13" collectionCode="KPM" collectorName="Yamamoto" country="Japan" location="Uchiko-cho" municipality="Odamiyama" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFCABB950D37C2341" box="[785,863,583,609]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
, same locality,
<date id="FFAA107E3111FF9EFBB3B950D4592341" box="[1033,1146,583,609]" pageId="15" pageNumber="318" value="2002-05-13">
<collectingDate id="EFEEE9963111FF9EFBB3B950D4592341" box="[1033,1146,583,609]" pageId="15" pageNumber="318" value="2002-05-13">13.v.2002</collectingDate>
</date>
, E.
<collectorName id="26E153683111FF9EFB18B950D53F2341" box="[1186,1308,583,609]" pageId="15" pageNumber="318">Yamamoto</collectorName>
(
<collectionCode id="ED05AE7B3111FF9EFA96B950D54D2341" box="[1324,1390,583,609]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFA3AB950D44523A5" collectingDate="2020-05-23" collectorName="E. Yamamoto" country="Japan" county="Namakusa-dani" location="Uchiko-cho" municipality="Odamiyama" pageId="15" pageNumber="318" specimenCount="38" specimenCount-female="6" specimenCount-male="32">
<specimenCount id="9D12FD373111FF9EFA3AB950D0F823A5" count="32" pageId="15" pageNumber="318" type="male">32 males</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFF5DB97CD17423A5" box="[231,343,619,645]" count="6" pageId="15" pageNumber="318" type="female">6 females</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFED9B97CD23523A5" box="[355,534,619,645]" pageId="15" pageNumber="318">Namakusa-dani</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFD98B97CD28D23A5" box="[546,686,619,645]" pageId="15" pageNumber="318">Odamiyama</collectingMunicipality>
,
<location id="8ECB60653111FF9EFD00B97CD36223A5" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFD00B97CD36223A5" box="[698,833,619,645]" country="Japan" county="Namakusa-dani" municipality="Odamiyama" name="Uchiko-cho" pageId="15" pageNumber="318">Uchiko-cho</location>
,
<date id="FFAA107E3111FF9EFCF7B97CD39D23A5" box="[845,958,619,645]" pageId="15" pageNumber="318" value="2020-05-23">
<collectingDate id="EFEEE9963111FF9EFCF7B97CD39D23A5" box="[845,958,619,645]" pageId="15" pageNumber="318" value="2020-05-23">23.v.2020</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFC70B97CD44523A5" box="[970,1126,619,645]" pageId="15" pageNumber="318">E. Yamamoto</collectorName>
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFBC9B97CD3392389" collectingDate="2020-05-17" collectorName="E. Yamamoto" country="Japan" county="Hondani" location="Uchiko-cho" municipality="Odamiyama" pageId="15" pageNumber="318" specimenCount="19" specimenCount-female="7" specimenCount-male="12">
<specimenCount id="9D12FD373111FF9EFBC9B97CD4F923A5" box="[1139,1242,619,645]" count="12" pageId="15" pageNumber="318" type="male">12 males</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFB5DB97CD57423A5" box="[1255,1367,619,645]" count="7" pageId="15" pageNumber="318" type="female">7 females</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFAD9B97CD0E92389" pageId="15" pageNumber="318">Hondani</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFF6CB998D1412389" box="[214,354,655,681]" pageId="15" pageNumber="318">Odamiyama</collectingMunicipality>
,
<location id="8ECB60653111FF9EFED4B998D1D62389" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFED4B998D1D62389" box="[366,501,655,681]" country="Japan" county="Hondani" municipality="Odamiyama" name="Uchiko-cho" pageId="15" pageNumber="318">Uchiko-cho</location>
,
<date id="FFAA107E3111FF9EFDBBB998D2512389" box="[513,626,655,681]" pageId="15" pageNumber="318" value="2020-05-17">
<collectingDate id="EFEEE9963111FF9EFDBBB998D2512389" box="[513,626,655,681]" pageId="15" pageNumber="318" value="2020-05-17">17.v.2020</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFDC4B998D3392389" box="[638,794,655,681]" pageId="15" pageNumber="318">E. Yamamoto</collectorName>
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFC9DB998D1CA23ED" collectingDate="2020-05-21" collectorName="E. Yamamoto" country="Japan" county="Koya-yama" location="Uchiko-cho" municipality="Odamiyama" pageId="15" pageNumber="318" specimenCount="16" specimenCount-female="3" specimenCount-male="13">
<specimenCount id="9D12FD373111FF9EFC9DB998D3AC2389" box="[807,911,655,681]" count="13" pageId="15" pageNumber="318" type="male">13 males</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFC21B998D4282389" box="[923,1035,655,681]" count="3" pageId="15" pageNumber="318" type="female">3 females</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFBADB998D4BE2389" box="[1047,1181,655,681]" pageId="15" pageNumber="318">Koya-yama</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFB13B998D5162389" box="[1193,1333,655,681]" pageId="15" pageNumber="318">Odamiyama</collectingMunicipality>
,
<location id="8ECB60653111FF9EFAFAB998D0E123ED" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFAFAB998D0E123ED" country="Japan" county="Koya-yama" municipality="Odamiyama" name="Uchiko-cho" pageId="15" pageNumber="318">Uchiko-cho</location>
,
<date id="FFAA107E3111FF9EFF75B9A4D16323ED" box="[207,320,691,717]" pageId="15" pageNumber="318" value="2020-05-21">
<collectingDate id="EFEEE9963111FF9EFF75B9A4D16323ED" box="[207,320,691,717]" pageId="15" pageNumber="318" value="2020-05-21">21.v.2020</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFEF6B9A4D1CA23ED" box="[332,489,691,717]" pageId="15" pageNumber="318">E. Yamamoto</collectorName>
</materialsCitation>
;
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<collectingDate id="EFEEE9963111FF9EFAB1B9A4D5B423ED" box="[1291,1431,691,717]" pageId="15" pageNumber="318" value="2018-05-01" valueMax="2018-05-20" valueMin="2018-05-01">120.v.2018</collectingDate>
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<emphasis id="B960EAAC3111FF9EFEFBB9C1D1AE23D0" bold="true" box="[321,397,726,752]" pageId="15" pageNumber="318">Kochi</emphasis>
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<emphasis id="B960EAAC3111FF9EFE37B9C1D1B123D0" bold="true" box="[397,402,726,752]" pageId="15" pageNumber="318">:</emphasis>
<materialsCitation id="3B7C3CE33111FF9EFE21B9C0D45A23D1" box="[411,1145,727,753]" collectingDate="1999-06-20" collectionCode="T" country="Japan" county="Nishikuma-rindo" location="Kami-shi" municipality="Monobe-cho" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFE21B9C0D1C823D1" box="[411,491,727,753]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFE43B9C0D29823D1" box="[505,699,727,753]" pageId="15" pageNumber="318">Nishikuma-rindo</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFD72B9C0D37923D1" box="[712,858,727,753]" pageId="15" pageNumber="318">Monobe-cho</collectingMunicipality>
,
<location id="8ECB60653111FF9EFCDDB9C0D3F123D1" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFCDDB9C0D3F123D1" box="[871,978,727,753]" country="Japan" county="Nishikuma-rindo" municipality="Monobe-cho" name="Kami-shi" pageId="15" pageNumber="318">Kami-shi</location>
,
<date id="FFAA107E3111FF9EFC5AB9C0D47923D1" box="[992,1114,727,753]" pageId="15" pageNumber="318" value="1999-06-20">
<collectingDate id="EFEEE9963111FF9EFC5AB9C0D47923D1" box="[992,1114,727,753]" pageId="15" pageNumber="318" value="1999-06-20">20.vi.1999</collectingDate>
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<collectionCode id="ED05AE7B3111FF9EFBDDB9C0D45A23D1" box="[1127,1145,727,753]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFB3CB9C0D53A23D1" box="[1158,1305,727,753]" collectionCode="KPM" country="Japan" location="Befu" pageId="15" pageNumber="318" specimenCount="1">
<location id="8ECB60653111FF9EFB3CB9C0D49F23D1" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFB3CB9C0D49F23D1" box="[1158,1212,727,753]" country="Japan" name="Befu" pageId="15" pageNumber="318">Befu</location>
(
<collectionCode id="ED05AE7B3111FF9EFB75B9C0D53223D1" box="[1231,1297,727,753]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="15" pageNumber="318" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D12FD373111FF9EFA9CB9C0D5A223D1" box="[1318,1409,727,753]" count="3" pageId="15" pageNumber="318" type="male">3 males</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFA34B9C0D0C52235" count="1" pageId="15" pageNumber="318" type="female">1 female</specimenCount>
, same,
<date id="FFAA107E3111FF9EFE83B9ECD1892235" box="[313,426,763,789]" pageId="15" pageNumber="318" value="2001-05-12">12.v.2001</date>
, M. Takai;
<specimenCount id="9D12FD373111FF9EFD92B9ECD2AE2235" box="[552,653,763,789]" count="1" pageId="15" pageNumber="318" type="female">1 female</specimenCount>
, same,
<date id="FFAA107E3111FF9EFD65B9EDD3612235" box="[735,834,762,789]" pageId="15" pageNumber="318" value="2004-05-05">5.v.2004</date>
, M. Takai;
<materialsCitation id="3B7C3CE33111FF9EFC7BB9ECD1762219" collectingDate="2004-07-03" collectionCode="T" country="Japan" county="Shiraga-toge" location="Kami-shi" municipality="Monobe-cho" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFC7BB9ECD42C2235" box="[961,1039,763,789]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFBA0B9ECD48F2235" box="[1050,1196,763,789]" pageId="15" pageNumber="318">Shiraga-toge</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFB0DB9ECD56A2235" box="[1207,1353,763,789]" pageId="15" pageNumber="318">Monobe-cho</collectingMunicipality>
,
<location id="8ECB60653111FF9EFAEFB9ECD0992219" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFAEFB9ECD0992219" country="Japan" county="Shiraga-toge" municipality="Monobe-cho" name="Kami-shi" pageId="15" pageNumber="318">Kami-shi</location>
,
<date id="FFAA107E3111FF9EFF7FB808D11A2219" box="[197,313,799,825]" pageId="15" pageNumber="318" value="2004-07-03">
<collectingDate id="EFEEE9963111FF9EFF7FB808D11A2219" box="[197,313,799,825]" pageId="15" pageNumber="318" value="2004-07-03">3.vii.2004</collectingDate>
</date>
,
<collectionCode id="ED05AE7B3111FF9EFEF9B808D1762219" box="[323,341,799,825]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFEE4B808D1BC2219" box="[350,415,799,825]" country="Japan" location="Takai" pageId="15" pageNumber="318" specimenCount="1">
<location id="8ECB60653111FF9EFEE4B808D1BC2219" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFEE4B808D1BC2219" box="[350,415,799,825]" country="Japan" name="Takai" pageId="15" pageNumber="318">Takai</location>
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFE13B808D4782219" box="[425,1115,798,825]" collectingDate="2002-05-05" collectorName="M. Takai" country="Japan" county="Befu-kyo" location="Kami-shi" municipality="Monobe-cho" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFE13B808D1D52219" box="[425,502,799,825]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFDBAB808D24E2219" box="[512,621,799,825]" pageId="15" pageNumber="318">Befu-kyo</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFDCDB808D32A2219" box="[631,777,799,825]" pageId="15" pageNumber="318">Monobe-cho</collectingMunicipality>
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<location id="8ECB60653111FF9EFCA9B808D35D2219" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFCA9B808D35D2219" box="[787,894,799,825]" country="Japan" county="Befu-kyo" municipality="Monobe-cho" name="Kami-shi" pageId="15" pageNumber="318">Kami-shi</location>
,
<date id="FFAA107E3111FF9EFC32B809D3C92219" box="[904,1002,798,825]" pageId="15" pageNumber="318" value="2002-05-05">
<collectingDate id="EFEEE9963111FF9EFC32B809D3C92219" box="[904,1002,798,825]" pageId="15" pageNumber="318" value="2002-05-05">5.v.2002</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFC4FB808D4782219" box="[1013,1115,799,825]" pageId="15" pageNumber="318">M. Takai</collectorName>
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFBDFB808D193227D" collectingDate="2002-05-11" collectorName="M. Takai" country="Japan" location="Tsuno-cho" municipality="Tengu-ike" pageId="15" pageNumber="318" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D12FD373111FF9EFBDFB808D4EB2219" box="[1125,1224,799,825]" count="1" pageId="15" pageNumber="318" type="female">1 female</specimenCount>
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<collectingMunicipality id="6BCFACC43111FF9EFB6BB808D5662219" box="[1233,1349,799,825]" pageId="15" pageNumber="318">Tengu-ike</collectingMunicipality>
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<location id="8ECB60653111FF9EFAF5B808D0E1227D" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFAF5B808D0E1227D" country="Japan" municipality="Tengu-ike" name="Tsuno-cho" pageId="15" pageNumber="318">Tsuno-cho</location>
,
<date id="FFAA107E3111FF9EFF74B854D11D227D" box="[206,318,835,861]" pageId="15" pageNumber="318" value="2002-05-11">
<collectingDate id="EFEEE9963111FF9EFF74B854D11D227D" box="[206,318,835,861]" pageId="15" pageNumber="318" value="2002-05-11">11.v.2002</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFEF3B854D193227D" box="[329,432,835,861]" pageId="15" pageNumber="318">M. Takai</collectorName>
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFE06B854D453227D" box="[444,1136,835,861]" collectingDate="2011-05-24" collectionCode="T" country="Japan" elevation="910" location="Tsuno-cho" municipality="Funato" pageId="15" pageNumber="318" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFE06B854D229227D" box="[444,522,835,861]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<specimenCount id="9D12FD373111FF9EFDAFB854D259227D" box="[533,634,835,861]" count="1" pageId="15" pageNumber="318" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BCFACC43111FF9EFD3FB854D2F5227D" box="[645,726,835,861]" pageId="15" pageNumber="318">Funato</collectingMunicipality>
,
<location id="8ECB60653111FF9EFD5AB854D37A227D" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFD5AB854D37A227D" box="[736,857,835,861]" country="Japan" municipality="Funato" name="Tsuno-cho" pageId="15" pageNumber="318">Tsuno-cho</location>
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<quantity id="4CEC9B5B3111FF9EFC34B854D3F4227D" box="[910,983,835,861]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="15" pageNumber="318" unit="m" value="910.0">
<elevation id="0039D18D3111FF9EFC34B854D3F4227D" box="[910,983,835,861]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="15" pageNumber="318" unit="m" value="910.0">910 m</elevation>
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,
<date id="FFAA107E3111FF9EFC59B854D470227D" box="[995,1107,835,861]" pageId="15" pageNumber="318" value="2011-05-24">
<collectingDate id="EFEEE9963111FF9EFC59B854D470227D" box="[995,1107,835,861]" pageId="15" pageNumber="318" value="2011-05-24">24.v.2011</collectingDate>
</date>
,
<collectionCode id="ED05AE7B3111FF9EFBE4B854D453227D" box="[1118,1136,835,861]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFBC1B854D517227D" box="[1147,1332,835,861]" collectionCode="SPMN" country="Japan" location="Hattori" pageId="15" pageNumber="318" specimenCount="1">
<location id="8ECB60653111FF9EFBC1B854D4E9227D" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFBC1B854D4E9227D" box="[1147,1226,835,861]" country="Japan" name="Hattori" pageId="15" pageNumber="318">Hattori</location>
(
<collectionCode id="ED05AE7B3111FF9EFB63B854D50F227D" box="[1241,1324,835,861]" pageId="15" pageNumber="318">SPMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFA85B854D38B22A1" collectingDate="1999-05-25" collectionCode="T" collectorName="A. Ohkawa" country="Japan" county="Nano-gawa" location="Ino-cho" municipality="Terakawa" pageId="15" pageNumber="318" specimenCount="8" specimenCount-female="2" specimenCount-male="6">
<specimenCount id="9D12FD373111FF9EFA85B854D5BB227D" box="[1343,1432,835,861]" count="6" pageId="15" pageNumber="318" type="male">6 males</specimenCount>
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<specimenCount id="9D12FD373111FF9EFF2DB870D12622A1" box="[151,261,871,897]" count="2" pageId="15" pageNumber="318" type="female">2 females</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFEAAB870D1B722A1" box="[272,404,871,897]" pageId="15" pageNumber="318">Nano-gawa</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFE24B870D22E22A1" box="[414,525,871,897]" pageId="15" pageNumber="318">Terakawa</collectingMunicipality>
,
<location id="8ECB60653111FF9EFDADB870D25222A1" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFDADB870D25222A1" box="[535,625,871,897]" country="Japan" county="Nano-gawa" municipality="Terakawa" name="Ino-cho" pageId="15" pageNumber="318">Ino-cho</location>
,
<date id="FFAA107E3111FF9EFDC6B871D2CE22A1" box="[636,749,870,897]" pageId="15" pageNumber="318" value="1999-05-25">
<collectingDate id="EFEEE9963111FF9EFDC6B871D2CE22A1" box="[636,749,870,897]" pageId="15" pageNumber="318" value="1999-05-25">25.v.1999</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFD4CB870D35722A1" box="[758,884,871,897]" pageId="15" pageNumber="318">A. Ohkawa</collectorName>
&amp;
<collectionCode id="ED05AE7B3111FF9EFC2CB870D38B22A1" box="[918,936,871,897]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFC09B870D3F622A1" box="[947,981,871,897]" country="Japan" location="Ito" pageId="15" pageNumber="318" specimenCount="1">
<location id="8ECB60653111FF9EFC09B870D3F622A1" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFC09B870D3F622A1" box="[947,981,871,897]" country="Japan" name="Ito" pageId="15" pageNumber="318">Ito</location>
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFC5AB870D18F2285" collectingDate="2011-06-09" collectionCode="T" country="Japan" county="Mt. Iyofuji" elevation="1540" location="Ino-cho" municipality="Nakanoka" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFC5AB870D40E22A1" box="[992,1069,871,897]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFB82B870D49622A1" box="[1080,1205,871,897]" pageId="15" pageNumber="318">Mt. Iyofuji</collectingCounty>
,
<collectingMunicipality id="6BCFACC43111FF9EFB05B870D51022A1" box="[1215,1331,871,897]" pageId="15" pageNumber="318">Nakanoka</collectingMunicipality>
,
<location id="8ECB60653111FF9EFA84B870D5BB22A1" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFA84B870D5BB22A1" box="[1342,1432,871,897]" country="Japan" county="Mt. Iyofuji" municipality="Nakanoka" name="Ino-cho" pageId="15" pageNumber="318">Ino-cho</location>
, alt.
<quantity id="4CEC9B5B3111FF9EFF7BB89DD13B2285" box="[193,280,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="15" pageNumber="318" unit="m" value="1540.0">
<elevation id="0039D18D3111FF9EFF7BB89DD13B2285" box="[193,280,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="15" pageNumber="318" unit="m" value="1540.0">1540 m</elevation>
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,
<date id="FFAA107E3111FF9EFE9EB89CD1AC2285" box="[292,399,907,933]" pageId="15" pageNumber="318" value="2011-06-09">
<collectingDate id="EFEEE9963111FF9EFE9EB89CD1AC2285" box="[292,399,907,933]" pageId="15" pageNumber="318" value="2011-06-09">9.vi.2011</collectingDate>
</date>
,
<collectionCode id="ED05AE7B3111FF9EFE20B89CD18F2285" box="[410,428,907,933]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="318">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE33111FF9EFE0DB89CD2582285" box="[439,635,907,933]" collectorName="K. Nio" country="Japan" location="Nozaki" pageId="15" pageNumber="318" specimenCount="1">
<location id="8ECB60653111FF9EFE0DB89CD22B2285" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFE0DB89CD22B2285" box="[439,520,907,933]" country="Japan" name="Nozaki" pageId="15" pageNumber="318">Nozaki</location>
&amp;
<collectorName id="26E153683111FF9EFD96B89CD2582285" box="[556,635,907,933]" pageId="15" pageNumber="318">K. Nio</collectorName>
</materialsCitation>
;
<materialsCitation id="3B7C3CE33111FF9EFD3DB89CD5752285" box="[647,1366,906,933]" collectingDate="2005-04-30" collectorName="M. Takai" country="Japan" county="Kuroson" location="Shimanto-shi" municipality="Nishitosa" pageId="15" pageNumber="318" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D12FD373111FF9EFD3DB89CD2F62285" box="[647,725,907,933]" count="1" pageId="15" pageNumber="318" type="male">1 male</specimenCount>
,
<collectingCounty id="62CA4E323111FF9EFD5AB89CD3602285" box="[736,835,907,933]" pageId="15" pageNumber="318">Kuroson</collectingCounty>
,
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,
<location id="8ECB60653111FF9EFC7DB89CD47C2285" LSID="urn:lsid:plazi:treatment:03BD87A83113FF9EFF2DB935D3292581:8ECB60653111FF9EFC7DB89CD47C2285" box="[967,1119,907,933]" country="Japan" county="Kuroson" municipality="Nishitosa" name="Shimanto-shi" pageId="15" pageNumber="318">Shimanto-shi</location>
,
<date id="FFAA107E3111FF9EFBD0B89DD4C02285" box="[1130,1251,906,933]" pageId="15" pageNumber="318" value="2005-04-30">
<collectingDate id="EFEEE9963111FF9EFBD0B89DD4C02285" box="[1130,1251,906,933]" pageId="15" pageNumber="318" value="2005-04-30">30.iv.2005</collectingDate>
</date>
,
<collectorName id="26E153683111FF9EFB54B89CD5752285" box="[1262,1366,907,933]" pageId="15" pageNumber="318">M. Takai</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353111FF9EFF7DB8B8D13422CD" pageId="15" pageNumber="318" type="etymology">
<paragraph id="8BAB36BE3111FF9EFF7DB8B8D13422CD" blockId="15.[151,1437,150,1185]" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF7DB8B8D16D22E9" bold="true" box="[199,334,943,969]" pageId="15" pageNumber="318">Etymology.</emphasis>
The specific epithet refers to a major island in
<collectingCountry id="F303762E3111FF9EFCE1B8B8D3BD22E9" box="[859,926,943,969]" name="Japan" pageId="15" pageNumber="318">Japan</collectingCountry>
, named Shikoku, where this species is widely distributed.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353111FF9EFF7DB8E1D1EF2531" box="[199,460,1014,1041]" pageId="15" pageNumber="318" type="distribution">
<paragraph id="8BAB36BE3111FF9EFF7DB8E1D1EF2531" blockId="15.[151,1437,150,1185]" box="[199,460,1014,1041]" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF7DB8E1D1422530" bold="true" box="[199,353,1014,1040]" pageId="15" pageNumber="318">Distribution.</emphasis>
Shikoku.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353111FF9EFF7DBF0DD3292581" pageId="15" pageNumber="318" type="biology_ecology">
<paragraph id="8BAB36BE3111FF9EFF7DBF0DD41A255D" blockId="15.[151,1437,150,1185]" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF7DBF0DD1042514" bold="true" box="[199,295,1050,1076]" pageId="15" pageNumber="318">Biology.</emphasis>
Many adults of this species, including a mating pair, were found resting on leaves of vegetation along spring seeps or small flows at the
<typeStatus id="54AF881C3111FF9EFD9CBF28D2752579" box="[550,598,1087,1113]" pageId="15" pageNumber="318">type</typeStatus>
locality in the daytime on
<date id="FFAA107E3111FF9EFC29BF28D40D2579" box="[915,1070,1087,1113]" pageId="15" pageNumber="318" value="2020-05-20">20 May 2020</date>
(
<figureCitation id="132F2A3B3111FF9EFB85BF28D4F22579" box="[1087,1233,1087,1113]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="15" pageNumber="318">Figs 7A, 7B</figureCitation>
), and were easily collected with an insect net (Yamamoto personal communication on
<date id="FFAA107E3111FF9EFC2DBF74D40E255D" box="[919,1069,1123,1149]" pageId="15" pageNumber="318" value="2024-07-13">13 July 2024</date>
).
</paragraph>
<paragraph id="8BAB36BE3111FF9EFF7DBF90D3292581" blockId="15.[151,1437,150,1185]" box="[199,778,1159,1185]" pageId="15" pageNumber="318">
<emphasis id="B960EAAC3111FF9EFF7DBF90D1A02581" bold="true" box="[199,387,1159,1185]" pageId="15" pageNumber="318">Japanese name.</emphasis>
Shikoku-kimadara-shima-tobikera
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,462 @@
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<mods:title id="90992710A84AF7099DE55D5C797F37EF">Redescriptions of Homoplectra japonica (Banks 1906) and H. albomarginata (Ulmer 1907) (Trichoptera, Hydropsychidae), with descriptions of five new similar species from Japan</mods:title>
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<paragraph id="8BAB36BE3115FF9AFF2DBA2AD2672077" blockId="11.[151,580,317,379]" box="[151,580,317,344]" pageId="11" pageNumber="314">
<heading id="D0E381D23115FF9AFF2DBA2AD2672077" bold="true" box="[151,580,317,344]" fontSize="11" level="1" pageId="11" pageNumber="314" reason="1">
<emphasis id="B960EAAC3115FF9AFF2DBA2AD2672077" bold="true" box="[151,580,317,344]" pageId="11" pageNumber="314">
<taxonomicName id="4C144D3D3115FF9AFF2DBA2AD1C32078" authorityName="Nozaki" authorityYear="2025" box="[151,480,317,344]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="species" species="flagelliformis" status="sp. nov.">
<emphasis id="B960EAAC3115FF9AFF2DBA2AD1C32078" bold="true" box="[151,480,317,344]" italics="true" pageId="11" pageNumber="314">Homoplectra flagelliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357D73115FF9AFE5DBA2AD2672077" box="[487,580,317,343]" pageId="11" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E65353115FF9AFF2DBA76D13A205B" box="[151,281,353,379]" pageId="11" pageNumber="314" type="description">
<paragraph id="8BAB36BE3115FF9AFF2DBA76D13A205B" blockId="11.[151,580,317,379]" box="[151,281,353,379]" pageId="11" pageNumber="314">
(
<figureCitation id="132F2A3B3115FF9AFF25BA76D0CB205B" box="[159,232,353,379]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="11" pageNumber="314">Figs 6</figureCitation>
,
<figureCitation id="132F2A3B3115FF9AFF4EBA76D133205B" box="[244,272,353,379]" captionStart="FIGURE 13" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[169,1415,195,1904]" captionTargetId="figure-33@22.[157,1430,181,1919]" captionTargetPageId="22" captionText="FIGURE 13. Distributions of species of the Homoplectra japonica Species Group in Japan." pageId="11" pageNumber="314">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E65353115FF9AFF2DBA88D39B20F6" pageId="11" pageNumber="314" type="reference_group">
<paragraph id="8BAB36BE3115FF9AFF2DBA88D40F2097" blockId="11.[151,1068,415,470]" box="[151,1068,415,439]" pageId="11" pageNumber="314">
<treatmentCitationGroup id="AB0411903115FF9AFF2DBA88D40F2097" box="[151,1068,415,439]" pageId="11" pageNumber="314">
<taxonomicName id="4C144D3D3115FF9AFF2DBA88D1532097" ID-CoL="36JDB" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[151,368,415,439]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3115FF9AFF2DBA88D1532097" box="[151,368,415,439]" italics="true" pageId="11" pageNumber="314">Diplectrona japonica</emphasis>
</taxonomicName>
nec
<bibRefCitation id="EF854B4F3115FF9AFE25BA88D2382097" author="Banks, N." box="[415,539,415,439]" pageId="11" pageNumber="314" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
:
<treatmentCitation id="0AB510AF3115FF9AFD9FBAB7D3472097" author="Kawase, N. &amp; Hayashi, M." box="[549,868,415,439]" page="85" pageId="11" pageNumber="314" year="2010">
<bibRefCitation id="EF854B4F3115FF9AFD9FBAB7D3072097" author="Kawase, N. &amp; Hayashi, M." box="[549,804,415,439]" pageId="11" pageNumber="314" pagination="83 - 86" refId="ref17291" refString="Kawase, N. &amp; Hayashi, M. (2010) Records of caddisflies from Mt. Sentsu-zan, Okuizumio-cho, Shimane Prefecture, Japan. Bulletin of Hoshizaki Green Foundation, 13, 83 - 86 [in Japanese with English abstract]." type="journal article" year="2010">Kawase &amp; Hayashi 2010</bibRefCitation>
, p. 85
</treatmentCitation>
.
<emphasis id="B960EAAC3115FF9AFCD4BA88D4052097" bold="true" box="[878,1062,415,439]" pageId="11" pageNumber="314">Misidentification</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAB36BE3115FF9AFF2DBAA9D39B20F6" blockId="11.[151,1068,415,470]" box="[151,952,446,470]" pageId="11" pageNumber="314">
Diplectroninae Gen.
<taxonomicName id="4C144D3D3115FF9AFED1BAA9D1E720F5" ID-CoL="36JDB" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[363,452,446,469]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3115FF9AFED1BAA9D1E720F5" box="[363,452,446,469]" italics="true" pageId="11" pageNumber="314">japonica</emphasis>
</taxonomicName>
Species Complex:
<bibRefCitation id="EF854B4F3115FF9AFD32BAA9D32E20F6" author="Nojima, K." box="[648,781,446,470]" pageId="11" pageNumber="314" pagination="107 - 131" refId="ref17913" refString="Nojima, K. (2017) Caddisflies (Insecta: Trichoptera) in Okayama Prefecture, western Japan. Biology of Inland Waters, 32, 107 - 131 [in Japanese with English abstract]." type="journal article" year="2017">Nojima 2017</bibRefCitation>
(in part), p. 119.
</paragraph>
</subSubSection>
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<paragraph id="8BAB36BE3115FF9AFF2DBC47D57726D9" blockId="11.[151,1436,1872,2041]" pageId="11" pageNumber="314">
<emphasis id="B960EAAC3115FF9AFF2DBC47D2CF2649" bold="true" box="[151,748,1872,1897]" pageId="11" pageNumber="314">
FIGURE 6. Adults of
<taxonomicName id="4C144D3D3115FF9AFEC7BC47D2BB2648" authorityName="Nozaki" authorityYear="2025" box="[381,664,1872,1896]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="species" species="flagelliformis" status="sp. nov.">
<emphasis id="B960EAAC3115FF9AFEC7BC47D2BB2648" bold="true" box="[381,664,1872,1896]" italics="true" pageId="11" pageNumber="314">Homoplectra flagelliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25357D73115FF9AFD27BC46D2CF2649" box="[669,748,1873,1897]" pageId="11" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral.
</paragraph>
</caption>
<subSubSection id="C30E65353112FF9DFF2DBB81D2D920D5" pageId="12" pageNumber="315" type="diagnosis">
<paragraph id="8BAB36BE3112FF9DFF2DBB81D23D2045" blockId="12.[151,1437,150,501]" pageId="12" pageNumber="315">
<emphasis id="B960EAAC3112FF9DFF2DBB81D1312190" bold="true" box="[151,274,150,176]" pageId="12" pageNumber="315">Diagnosis.</emphasis>
The male of this species is very similar to that of
<taxonomicName id="4C144D3D3112FF9DFC85BB80D3E02190" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[831,963,151,176]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3112FF9DFC85BB80D3E02190" box="[831,963,151,176]" italics="true" pageId="12" pageNumber="315">H. japonica</emphasis>
</taxonomicName>
in the short antennal scapes and the shape of the phallic apparatus; but can be distinguished by the shape of the ventral processes of the phallotheca and aedeagus: In this species, the ventral processes of the phallotheca are almost straight apically (
<figureCitation id="132F2A3B3112FF9DFB11BBC8D51B21D9" box="[1195,1336,223,249]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Figs 6D, 6H</figureCitation>
), and the stem of the aedeagus bears a pair of round lobes apically in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFC61BA14D415203D" box="[987,1078,259,285]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6D</figureCitation>
); but in
<taxonomicName id="4C144D3D3112FF9DFB28BA14D534203C" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1170,1303,259,284]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3112FF9DFB28BA14D534203C" box="[1170,1303,259,284]" italics="true" pageId="12" pageNumber="315">H. japonica</emphasis>
</taxonomicName>
, the ventral processes of phallotheca curve dotsomesad apically (
<figureCitation id="132F2A3B3112FF9DFD5DBA30D3522061" box="[743,881,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="12" pageNumber="315">Figs 3D, 3E</figureCitation>
), and the posterior margin of the aedeagal stem is truncate in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFE0DBA5CD2312045" box="[439,530,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="12" pageNumber="315">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE3112FF9DFF7DBA78D2D920D5" blockId="12.[151,1437,150,501]" pageId="12" pageNumber="315">
The female of this species is similar to that of
<taxonomicName id="4C144D3D3112FF9DFD7ABA78D38C20A9" authorityName="Nozaki" authorityYear="2025" box="[704,943,367,393]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="shikoku" status="sp. nov.">
<emphasis id="B960EAAC3112FF9DFD7ABA78D38C20A9" box="[704,943,367,393]" italics="true" pageId="12" pageNumber="315">Homoplectra shikoku</emphasis>
</taxonomicName>
<emphasis id="B960EAAC3112FF9DFC0FBA78D42E20A9" bold="true" box="[949,1037,367,393]" pageId="12" pageNumber="315">
<taxonomicNameLabel id="A25357D73112FF9DFC0FBA78D42E20A9" box="[949,1037,367,393]" pageId="12" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in having a trapezoidal basoventral lobe on the vulvar scale but these species can be easily distinguished from each other by the shape of the lateral lobes of segment VIII: The mesal margin of each lateral lobe is concave in the middle in this species (
<figureCitation id="132F2A3B3112FF9DFAB0BAA0D54220F1" box="[1290,1377,439,465]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6K</figureCitation>
), but it is smoothly convex in
<taxonomicName id="4C144D3D3112FF9DFE11BACCD20720D5" authorityName="Nozaki" authorityYear="2025" box="[427,548,475,501]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="shikoku" status="sp. nov.">
<emphasis id="B960EAAC3112FF9DFE11BACCD20720D5" box="[427,548,475,501]" italics="true" pageId="12" pageNumber="315">H. shikoku</emphasis>
</taxonomicName>
<emphasis id="B960EAAC3112FF9DFD90BACCD2A720D5" bold="true" box="[554,644,475,501]" pageId="12" pageNumber="315">
<taxonomicNameLabel id="A25357D73112FF9DFD90BACCD2A720D5" box="[554,644,475,501]" pageId="12" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132F2A3B3112FF9DFD29BACCD2CA20D5" box="[659,745,475,501]" captionStart="FIGURE 7" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[157,1429,187,1666]" captionTargetId="figure-33@14.[151,1435,181,1672]" captionTargetPageId="14" captionText="FIGURE 7. Adults of Homoplectra shikoku sp. nov. 7A, male, habitus, type locality (Kumakogen-cho, Ehime, photographed by E. Yamamoto). 7B, a spring seep at the type locality where many adults were collected (photographed by E. Yamamoto). 7C7G, male genitalia, type locality: 7C, left lateral, apicomesal part of inferior appendage enlarged; 7D, dorsal; 7E, ventral; 7F, phallic apparatus, left lateral; 7G, same, dorsal. 7H17H3, posterior process of male segment X, variations (Uchiko-cho, Ehime), left lateral. 7I7K, female genitalia, type locality: 7I, left lateral, 7J, dorsal; 7K, ventral." pageId="12" pageNumber="315">Fig. 7K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E65353112FF9DFF2DB951D2302625" pageId="12" pageNumber="315" type="description">
<paragraph id="8BAB36BE3112FF9DFF2DB951D10B2340" blockId="12.[151,296,582,608]" box="[151,296,582,608]" pageId="12" pageNumber="315">
<emphasis id="B960EAAC3112FF9DFF2DB951D10B2340" bold="true" box="[151,296,582,608]" pageId="12" pageNumber="315">Description</emphasis>
</paragraph>
<paragraph id="8BAB36BE3112FF9DFF2DB999D25023D1" blockId="12.[151,1437,654,2013]" pageId="12" pageNumber="315">
<emphasis id="B960EAAC3112FF9DFF2DB999D0C02388" bold="true" box="[151,227,654,680]" pageId="12" pageNumber="315">Adult.</emphasis>
Length of antennal scape and general coloration of body and wings very similar to those of
<taxonomicName id="4C144D3D3112FF9DFAB5B998D5B62388" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1295,1429,655,680]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC3112FF9DFAB5B998D5B62388" box="[1295,1429,655,680]" italics="true" pageId="12" pageNumber="315">H. japonica</emphasis>
</taxonomicName>
. Wings occasionally uniformly dark brown in female. Forewings each
<quantity id="4CEC9B5B3112FF9DFC1DB9A5D40423ED" box="[935,1063,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" pageId="12" pageNumber="315" unit="mm" value="9.25" valueMax="10.0" valueMin="8.5">8.510 mm</quantity>
in male (n = 10),
<quantity id="4CEC9B5B3112FF9DFB48B9A5D5A323ED" box="[1266,1408,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.3" metricValueMin="1.05" pageId="12" pageNumber="315" unit="mm" value="11.75" valueMax="13.0" valueMin="10.5">10.513 mm</quantity>
in female (n = 10),
<quantity id="4CEC9B5B3112FF9DFEE9B9C0D18A23D1" box="[339,425,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="12" pageNumber="315" unit="mm" value="9.7">9.7 mm</quantity>
in male
<typeStatus id="54AF881C3112FF9DFDB0B9C0D25323D1" box="[522,624,727,753]" pageId="12" pageNumber="315" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BAB36BE3112FF9DFF7DB9ECD16125C9" blockId="12.[151,1437,654,2013]" pageId="12" pageNumber="315">
<emphasis id="B960EAAC3112FF9DFF7DB9ECD1502234" bold="true" box="[199,371,762,789]" pageId="12" pageNumber="315">Male genitalia</emphasis>
(
<figureCitation id="132F2A3B3112FF9DFE38B9ECD2372235" box="[386,532,763,789]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Figs 6A6H</figureCitation>
). Ventrolateral sides of synsclerotized segment IX protruding anterad and round on each side in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFE4DB808D2702219" box="[503,595,799,825]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6A</figureCitation>
); posteroventral lobe long and rectangular in ventral aspect (
<figureCitation id="132F2A3B3112FF9DFAAAB808D5482219" box="[1296,1387,799,825]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6C</figureCitation>
), its apex weakly bilobed in ventral aspect. Dorsal part of segment IX subtriangular, apically round in dorsal aspect (
<figureCitation id="132F2A3B3112FF9DFF25B870D0D522A1" box="[159,246,871,897]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6B</figureCitation>
), fused with segment X laterally (
<figureCitation id="132F2A3B3112FF9DFDD4B870D2E422A1" box="[622,711,871,897]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6A</figureCitation>
). Segment X bilobed in dorsal aspect (
<figureCitation id="132F2A3B3112FF9DFBC8B870D4EA22A1" box="[1138,1225,871,897]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6B</figureCitation>
), with pair of large posterolateral setose areas (
<figureCitation id="132F2A3B3112FF9DFE69B89CD2132285" box="[467,560,907,933]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6A</figureCitation>
); each ventrolateral margin sclerotized, recessed into segment IX (
<figureCitation id="132F2A3B3112FF9DFA89B89CD5AC2285" box="[1331,1423,907,933]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6A</figureCitation>
). Inferior appendages each without distal segment, long and finger-like in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFBDDB8B8D4E322E9" box="[1127,1216,943,969]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6A</figureCitation>
), extending beyond apex of segment X, slightly curved upward apically; slightly convex subapicomesally, with about 10 spine-like setae (
<figureCitation id="132F2A3B3112FF9DFF67B8E0D1492531" box="[221,362,1015,1041]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Figs 6A6C</figureCitation>
). In phallotheca, long dorsal process thin, whip-like in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFBD1B8E0D4E72531" box="[1131,1220,1015,1041]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6D</figureCitation>
), gradually curving posteroventrad (specimens macerated by KOH solution often strongly curved or recurved apically as in
<figureCitation id="132F2A3B3112FF9DFA83BF0CD5AD2515" box="[1337,1422,1051,1077]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6F</figureCitation>
); broad basally and tapering to acute apex in dorsal aspect (
<figureCitation id="132F2A3B3112FF9DFCA9BF28D34A2579" box="[787,873,1087,1113]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6E</figureCitation>
); lateral processes absent; pair of ventral processes in lateral and dorsal aspect (
<figureCitation id="132F2A3B3112FF9DFE64BF74D24A255D" box="[478,617,1123,1149]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Figs 6D6F</figureCitation>
) slender, almost straight, approximately half as long as dorsal process. Aedeagus evenly curved ventrad in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFD68BF90D30E2581" box="[722,813,1159,1185]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6D</figureCitation>
); stem half-pipe-like, with pair of posterior lobes, each lobe oval in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFE73BFBCD20625E5" box="[457,549,1195,1221]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Figs 6D</figureCitation>
) and concave mesally (
<figureCitation id="132F2A3B3112FF9DFC8BBFBCD3AA25E5" box="[817,905,1195,1221]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6E</figureCitation>
); with head approximately 1/4 as long as basal stem (
<figureCitation id="132F2A3B3112FF9DFF61BFD8D11525C9" box="[219,310,1231,1257]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6D</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE3112FF9DFF7DBFE4D0E3270D" blockId="12.[151,1437,654,2013]" pageId="12" pageNumber="315">
<emphasis id="B960EAAC3112FF9DFF7DBFE4D1B3242C" bold="true" box="[199,400,1266,1293]" pageId="12" pageNumber="315">Female genitalia</emphasis>
(
<figureCitation id="132F2A3B3112FF9DFE18BFE4D20B242D" box="[418,552,1267,1293]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Figs 6I6L</figureCitation>
). Lateral lobes of sternum VIII widely separated from each other in ventral aspect (
<figureCitation id="132F2A3B3112FF9DFF56BE00D1612411" box="[236,322,1303,1329]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6K</figureCitation>
), each with mesal margin concave in middle. Segment IX oblique rectangular in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFAD5BE00D0932475" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6I</figureCitation>
), subpentagonal with round lateral and apical margins in dorsal aspect (
<figureCitation id="132F2A3B3112FF9DFC63BE2CD4092475" box="[985,1066,1339,1365]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6J</figureCitation>
); pair of mesal lobes sclerotized, semicircular in lateral aspect (
<figureCitation id="132F2A3B3112FF9DFE41BE48D26D2459" box="[507,590,1375,1401]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6I</figureCitation>
); pair of sclerotized ribs forming wide trapezoidal lobe on vulvar scale basoventrally (
<figureCitation id="132F2A3B3112FF9DFE87BE94D1B124BD" box="[317,402,1411,1437]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6K</figureCitation>
). Vulvar scale large, membranous and convex apically. Vaginal apparatus long pentagonal in dorsal aspect (
<figureCitation id="132F2A3B3112FF9DFE82BEB0D1AB24E1" box="[312,392,1447,1473]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6J</figureCitation>
), tapering to duct of bursa copulatrix; posterodorsal part semimembranous, with complicated wrinkles, with pair of longitudinal ridges dorsally, opening of duct of spermatheca situated between ridges near middle, with dark inverted V-shaped mark dorsally, partially surrounding opening of duct of spermatheca (
<figureCitation id="132F2A3B3112FF9DFAD5BEF8D097270D" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6J</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE3112FF9DFF7DBD21D32627C1" blockId="12.[151,1437,654,2013]" pageId="12" pageNumber="315">
<emphasis id="B960EAAC3112FF9DFF7DBD21D3102770" bold="true" box="[199,819,1590,1617]" pageId="12" pageNumber="315">Geographic variations of male and female genitalia</emphasis>
(
<figureCitation id="132F2A3B3112FF9DFCFEBD20D4262771" box="[836,1029,1591,1617]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Figs 6G, 6H, 6L</figureCitation>
). In males collected from
<collectingRegion id="49D0F85C3112FF9DFA8BBD20D5BF2771" box="[1329,1436,1591,1617]" country="Japan" name="Okayama" pageId="12" pageNumber="315">Okayama</collectingRegion>
and
<collectingRegion id="49D0F85C3112FF9DFF72BD4CD11D2755" box="[200,318,1627,1653]" country="Japan" name="Hiroshima" pageId="12" pageNumber="315">Hiroshima</collectingRegion>
Prefectures, the posteroventral lobe of segment IX is short (
<figureCitation id="132F2A3B3112FF9DFC57BD4CD46A2755" box="[1005,1097,1627,1653]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6G</figureCitation>
). The ventral processes of the phallotheca are approximately the same length as the dorsal process of the phallotheca in specimens collected from
<collectingRegion id="49D0F85C3112FF9DFF2DBDB4D1A5279D" box="[151,390,1699,1725]" country="Japan" name="Hiroshima" pageId="12" pageNumber="315">Hiroshima Prefecture</collectingRegion>
(
<figureCitation id="132F2A3B3112FF9DFE2EBDB4D1CF279D" box="[404,492,1699,1725]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6H</figureCitation>
). In females collected from
<collectingRegion id="49D0F85C3112FF9DFCA4BDB4D433279D" box="[798,1040,1699,1725]" country="Japan" name="Hiroshima" pageId="12" pageNumber="315">Hiroshima Prefecture</collectingRegion>
, the posterior margin of basoventral lobe on the vulvar scale is shallowly concave (
<figureCitation id="132F2A3B3112FF9DFD1BBDD0D2DB27C1" box="[673,760,1735,1761]" captionStart="FIGURE 6" captionStartId="11.[151,250,1872,1896]" captionTargetBox="[165,1420,498,1834]" captionTargetId="figure-95@11.[151,1435,491,1849]" captionTargetPageId="11" captionText="FIGURE 6. Adults of Homoplectra flagelliformis sp. nov. 6A6F, male genitalia, type locality (Okuizumo-cho, Shimane): 6A, left lateral, apicomesal part of inferior appendage, enlarged; 6B, dorsal, 6C, ventral; 6D, phallic apparatus, left lateral; 6E, same, dorsal; 6F, phallotheca, treated by KOH, left lateral. 6G &amp; 6H, variations of male genitalia (Hatsukaichi-shi, Hiroshima): 6G, posteroventral lobe of segment IX, ventral; 6H, phallotheca, left lateral. 6I6K, female genitalia, type locality: 6I, left lateral; 6J, dorsal; 6K, ventral. 6L, lobe formed by sclerotized ribs on vulvar scale, variation (Hatsukaichi-shi, Hiroshima), ventral." pageId="12" pageNumber="315">Fig. 6L</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE3112FF9DFF7DBDFCD2302625" blockId="12.[151,1437,654,2013]" box="[199,531,1771,1797]" pageId="12" pageNumber="315">
<emphasis id="B960EAAC3112FF9DFF7DBDFCD1B52625" bold="true" box="[199,406,1771,1797]" pageId="12" pageNumber="315">Immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353112FF9CFF7DBC18D41B21D9" lastPageId="13" lastPageNumber="316" pageId="12" pageNumber="315" type="materials_examined">
<paragraph id="8BAB36BE3112FF9DFF7DBC18D312266D" blockId="12.[151,1437,654,2013]" pageId="12" pageNumber="315">
<materialsCitation id="3B7C3CE33112FF9DFF7DBC18D312266D" collectingDate="2007-05-16" collectingDateMax="2007-05-28" collectingDateMin="2007-05-16" collectionCode="KPM-NK" collectorName="M. Hayashi" county="Male" elevation="761" latitude="35.1693" location="Okuizumo-cho" longLatPrecision="6" longitude="133.1751" municipality="Takezaki" pageId="12" pageNumber="315" specimenCode="TN00580" specimenCount="1" stateProvince="Shimane" typeStatus="holotype">
<emphasis id="B960EAAC3112FF9DFF7DBC18D1182609" bold="true" box="[199,315,1807,1833]" pageId="12" pageNumber="315">
<typeStatus id="54AF881C3112FF9DFF7DBC18D1152609" box="[199,310,1807,1833]" pageId="12" pageNumber="315" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62CA4E323112FF9DFEFCBC18D1A32609" box="[326,384,1807,1833]" pageId="12" pageNumber="315">Male</collectingCounty>
(in alcohol):
<collectingMunicipality id="6BCFACC43112FF9DFD98BC18D2A92609" box="[546,650,1807,1833]" pageId="12" pageNumber="315">Takezaki</collectingMunicipality>
,
<location id="8ECB60653112FF9DFD22BC18D3602609" LSID="urn:lsid:plazi:treatment:03BD87A83115FF9CFF2DBA2AD32F20F1:8ECB60653112FF9DFD22BC18D3602609" box="[664,835,1807,1833]" county="Male" latitude="35.1693" longLatPrecision="6" longitude="133.1751" municipality="Takezaki" name="Okuizumo-cho" pageId="12" pageNumber="315" stateProvince="Shimane">Okuizumo-cho</location>
,
<collectingRegion id="49D0F85C3112FF9DFCE8BC18D3942609" box="[850,951,1807,1833]" country="Japan" name="Shimane" pageId="12" pageNumber="315">Shimane</collectingRegion>
,
<geoCoordinate id="EE2050793112FF9DFC7FBC19D4612609" box="[965,1090,1806,1833]" degrees="35.1693" direction="north" orientation="latitude" pageId="12" pageNumber="315" precision="5" value="35.1693">35.1693°N</geoCoordinate>
,
<geoCoordinate id="EE2050793112FF9DFBEBBC19D4FA2609" box="[1105,1241,1806,1833]" degrees="133.1751" direction="east" orientation="longitude" pageId="12" pageNumber="315" precision="5" value="133.1751">133.1751°E</geoCoordinate>
, alt.
<quantity id="4CEC9B5B3112FF9DFAACBC18D5402609" box="[1302,1379,1807,1833]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="12" pageNumber="315" unit="m" value="761.0">
<elevation id="0039D18D3112FF9DFAACBC18D5402609" box="[1302,1379,1807,1833]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="12" pageNumber="315" unit="m" value="761.0">761 m</elevation>
</quantity>
,
<date id="FFAA107E3112FF9DFAC8BC18D12B266D" pageId="12" pageNumber="315" value="2007-05-16" valueMax="2007-05-28" valueMin="2007-05-16">
<collectingDate id="EFEEE9963112FF9DFAC8BC18D12B266D" pageId="12" pageNumber="315" value="2007-05-16" valueMax="2007-05-28" valueMin="2007-05-16">16 28.v.2007</collectingDate>
</date>
,
<collectorName id="26E153683112FF9DFEA9BC24D1BA266D" box="[275,409,1843,1869]" pageId="12" pageNumber="315">M. Hayashi</collectorName>
,
<collectingMethod id="52554EA93112FF9DFE1EBC24D213266D" box="[420,560,1843,1869]" pageId="12" pageNumber="315">Malaise trap</collectingMethod>
(
<collectionCode id="ED05AE7B3112FF9DFDFABC24D28D266D" box="[576,686,1843,1869]" pageId="12" pageNumber="315">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53112FF9DFD0EBC25D305266D" box="[692,806,1842,1869]" collectionCode="TN" pageId="12" pageNumber="315">TN00580</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAB36BE3112FF9DFF7DBC40D13726B5" blockId="12.[151,1437,654,2013]" pageId="12" pageNumber="315">
<materialsCitation id="3B7C3CE33112FF9DFF7DBC40D13326B5" collectingDate="2007-05-16" collectingDateMax="2007-05-28" collectingDateMin="2007-05-16" collectionCode="KPM-NK" collectorName="M. Hayashi" county="Male" elevation="761" latitude="35.1693" location="Okuizumo-cho" longLatPrecision="6" longitude="133.1751" municipality="Takezaki" pageId="12" pageNumber="315" specimenCode="TN00590, TN00600" specimenCount="15" specimenCount-female="5" specimenCount-male="10" stateProvince="Shimane" typeStatus="paratype">
<emphasis id="B960EAAC3112FF9DFF7DBC40D1652651" bold="true" box="[199,326,1879,1905]" pageId="12" pageNumber="315">
<typeStatus id="54AF881C3112FF9DFF7DBC40D1622651" box="[199,321,1879,1905]" pageId="12" pageNumber="315" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D12FD373112FF9DFEECBC40D1E52651" box="[342,454,1879,1905]" count="10" pageId="12" pageNumber="315" type="male">10 males</specimenCount>
,
<specimenCount id="9D12FD373112FF9DFE61BC41D2732651" box="[475,592,1878,1905]" count="5" pageId="12" pageNumber="315" type="female">5 females</specimenCount>
(in alcohol), same data as holotype (
<collectionCode id="ED05AE7B3112FF9DFB97BC40D4B82651" box="[1069,1179,1879,1905]" pageId="12" pageNumber="315">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53112FF9DFB11BC41D5392651" box="[1195,1306,1878,1905]" collectionCode="TN" pageId="12" pageNumber="315">TN00590</specimenCode>
,
<collectionCode id="ED05AE7B3112FF9DFA94BC40D5BF2651" box="[1326,1436,1879,1905]" pageId="12" pageNumber="315">KPM-NK</collectionCode>
<specimenCode id="DBB29EC53112FF9DFF2DBC6CD12B26B5" box="[151,264,1915,1941]" collectionCode="TN" pageId="12" pageNumber="315">TN00600</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAB36BE3112FF9CFF7DBC89D41B21D9" blockId="12.[151,1437,654,2013]" lastBlockId="13.[151,1437,150,465]" lastPageId="13" lastPageNumber="316" pageId="12" pageNumber="315">
<materialsCitation id="3B7C3CE33112FF9DFF7DBC89D21226FD" collectingDate="1993-05-25" collectorName="Funadani-gawa &amp; Kofu-cho &amp; N. Kuhara" country="Japan" elevation="750" location="Tottori" pageId="12" pageNumber="315" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Tottori">
<emphasis id="B960EAAC3112FF9DFF7DBC89D34B2698" bold="true" box="[199,872,1950,1976]" pageId="12" pageNumber="315">
<typeStatus id="54AF881C3112FF9DFF7DBC89D2282698" box="[199,523,1950,1976]" isOtherTypeStatus="true" pageId="12" pageNumber="315">Other specimens examined</typeStatus>
.
<collectingCountry id="F303762E3112FF9DFDA2BC89D2552698" box="[536,630,1950,1976]" name="Japan" pageId="12" pageNumber="315">JAPAN</collectingCountry>
: HONSHU:
<collectingRegion id="49D0F85C3112FF9DFCB6BC89D3402698" box="[780,867,1950,1976]" country="Japan" name="Tottori" pageId="12" pageNumber="315">Tottori</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD373112FF9DFCD5BC88D39D2699" box="[879,958,1951,1977]" count="1" pageId="12" pageNumber="315" type="male">1 male</specimenCount>
,
<specimenCount id="9D12FD373112FF9DFC70BC88D41A2699" box="[970,1081,1951,1977]" count="2" pageId="12" pageNumber="315" type="female">2 females</specimenCount>
,
<collectorName id="26E153683112FF9DFBFFBC88D4D62699" box="[1093,1269,1951,1977]" pageId="12" pageNumber="315">Funadani-gawa</collectorName>
,
<collectorName id="26E153683112FF9DFABABC88D54D2699" box="[1280,1390,1951,1977]" pageId="12" pageNumber="315">Kofu-cho</collectorName>
, alt.
<quantity id="4CEC9B5B3112FF9DFF2DBCD5D0C326FD" box="[151,224,1986,2013]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="315" unit="m" value="750.0">
<elevation id="0039D18D3112FF9DFF2DBCD5D0C326FD" box="[151,224,1986,2013]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="315" unit="m" value="750.0">750 m</elevation>
</quantity>
,
<date id="FFAA107E3112FF9DFF56BCD5D17E26FD" box="[236,349,1986,2013]" pageId="12" pageNumber="315" value="1993-05-25">
<collectingDate id="EFEEE9963112FF9DFF56BCD5D17E26FD" box="[236,349,1986,2013]" pageId="12" pageNumber="315" value="1993-05-25">25.v.1993</collectingDate>
</date>
,
<collectorName id="26E153683112FF9DFED2BCD4D1FE26FD" box="[360,477,1987,2013]" pageId="12" pageNumber="315">N. Kuhara</collectorName>
(KuN)
</materialsCitation>
.
<materialsCitation id="3B7C3CE33112FF9CFD81BCD5D0E82191" collectingDate="2007-05-16" collectingDateMax="2007-05-28" collectingDateMin="2007-05-16" collectionCode="NK" collectorName="M. Hayashi" country="Japan" county="Male" elevation="761" lastPageId="13" lastPageNumber="316" latitude="35.1693" location="Okuizumo-cho" longLatPrecision="6" longitude="133.1751" municipality="Takezaki" pageId="12" pageNumber="315" specimenCount="64" specimenCount-female="19" specimenCount-male="45" stateProvince="Shimane">
<emphasis id="B960EAAC3112FF9DFD81BCD5D28E26FC" bold="true" box="[571,685,1986,2012]" pageId="12" pageNumber="315">
<collectingRegion id="49D0F85C3112FF9DFD81BCD5D28B26FC" box="[571,680,1986,2012]" country="Japan" name="Shimane" pageId="12" pageNumber="315">Shimane</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD373112FF9DFD0EBCD5D33826FD" box="[692,795,1986,2013]" count="25" pageId="12" pageNumber="315" type="male">25 males</specimenCount>
,
<specimenCount id="9D12FD373112FF9DFC9CBCD4D38726FD" box="[806,932,1987,2013]" count="12" pageId="12" pageNumber="315" type="female">12 females</specimenCount>
, same data as holotype (
<specimenCount id="9D12FD373112FF9DFB0EBCD4D53E26FD" box="[1204,1309,1987,2013]" count="20" pageId="12" pageNumber="315" type="male">20 males</specimenCount>
,
<specimenCount id="9D12FD373112FF9DFA92BCD4D5BA26FD" box="[1320,1433,1987,2013]" count="7" pageId="12" pageNumber="315" type="female">7 females</specimenCount>
:
<collectionCode id="ED05AE7B3113FF9CFF2DBB80D09C2191" box="[151,191,151,177]" pageId="13" pageNumber="316">NK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CE33113FF9CFF6DBB81D4792191" box="[215,1114,150,177]" collectingDate="2016-05-08" collectionCode="KN" collectorName="Oosaooino &amp; Niimi-shi &amp; K. Nojima" country="Japan" location="Okayama" pageId="13" pageNumber="316" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Okayama">
<emphasis id="B960EAAC3113FF9CFF6DBB81D1762190" bold="true" box="[215,341,150,176]" pageId="13" pageNumber="316">
<collectingRegion id="49D0F85C3113FF9CFF6DBB81D1732190" box="[215,336,150,176]" country="Japan" name="Okayama" pageId="13" pageNumber="316">Okayama</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD373113FF9CFEE6BB81D1962191" box="[348,437,150,177]" count="5" pageId="13" pageNumber="316" type="male">5 males</specimenCount>
,
<specimenCount id="9D12FD373113FF9CFE7ABB80D2072191" box="[448,548,151,177]" count="1" pageId="13" pageNumber="316" type="female">1 female</specimenCount>
,
<collectorName id="26E153683113FF9CFD95BB80D2882191" box="[559,683,151,177]" pageId="13" pageNumber="316">Oosaooino</collectorName>
,
<collectorName id="26E153683113FF9CFD0DBB80D3062191" box="[695,805,151,177]" pageId="13" pageNumber="316">Niimi-shi</collectorName>
,
<date id="FFAA107E3113FF9CFC8ABB80D3B02191" box="[816,915,151,177]" pageId="13" pageNumber="316" value="2016-05-08">
<collectingDate id="EFEEE9963113FF9CFC8ABB80D3B02191" box="[816,915,151,177]" pageId="13" pageNumber="316" value="2016-05-08">8.v.2016</collectingDate>
</date>
,
<collectorName id="26E153683113FF9CFC24BB80D4372191" box="[926,1044,151,177]" pageId="13" pageNumber="316">K. Nojima</collectorName>
(
<collectionCode id="ED05AE7B3113FF9CFB9FBB80D4712191" box="[1061,1106,151,177]" pageId="13" pageNumber="316">KN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33113FF9CFBDFBB80D24521F5" collectingDate="2016-05-05" collectionCode="KN" collectorName="Hiruzenkamifukuda &amp; Maniwa-shi" country="Japan" pageId="13" pageNumber="316" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="9D12FD373113FF9CFBDFBB80D49D2191" box="[1125,1214,151,177]" count="3" pageId="13" pageNumber="316" type="male">3 males</specimenCount>
,
<specimenCount id="9D12FD373113FF9CFB70BB80D50D2191" box="[1226,1326,151,177]" count="1" pageId="13" pageNumber="316" type="female">1 female</specimenCount>
,
<collectorName id="26E153683113FF9CFA83BB80D10321F5" pageId="13" pageNumber="316">Hiruzenkamifukuda</collectorName>
,
<collectorName id="26E153683113FF9CFE96BBACD19621F5" box="[300,437,187,213]" pageId="13" pageNumber="316">Maniwa-shi</collectorName>
,
<date id="FFAA107E3113FF9CFE7ABBADD20221F5" box="[448,545,186,213]" pageId="13" pageNumber="316" value="2016-05-05">
<collectingDate id="EFEEE9963113FF9CFE7ABBADD20221F5" box="[448,545,186,213]" pageId="13" pageNumber="316" value="2016-05-05">5.v.2016</collectingDate>
</date>
(
<collectionCode id="ED05AE7B3113FF9CFD8BBBACD27D21F5" box="[561,606,187,213]" pageId="13" pageNumber="316">KN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CE33113FF9CFDC8BBADD10821D9" collectingDate="1976-05-11" collectionCode="KPM" collectorName="Yoshiwa &amp; Hatsukaichi-shi &amp; K. Baba" country="Japan" location="Hiroshima" pageId="13" pageNumber="316" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Hiroshima">
<emphasis id="B960EAAC3113FF9CFDC8BBADD2D921F4" bold="true" box="[626,762,186,212]" pageId="13" pageNumber="316">
<collectingRegion id="49D0F85C3113FF9CFDC8BBADD2D621F4" box="[626,757,186,212]" country="Japan" name="Hiroshima" pageId="13" pageNumber="316">Hiroshima</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD373113FF9CFCB8BBACD37221F5" box="[770,849,187,213]" count="1" pageId="13" pageNumber="316" type="male">1 male</specimenCount>
,
<specimenCount id="9D12FD373113FF9CFCE7BBACD3EE21F5" box="[861,973,187,213]" count="2" pageId="13" pageNumber="316" type="female">2 females</specimenCount>
,
<collectorName id="26E153683113FF9CFC6DBBACD41821F5" box="[983,1083,187,213]" pageId="13" pageNumber="316">Yoshiwa</collectorName>
,
<collectorName id="26E153683113FF9CFBFDBBACD4D921F5" box="[1095,1274,187,213]" pageId="13" pageNumber="316">Hatsukaichi-shi</collectorName>
,
<date id="FFAA107E3113FF9CFABFBBACD55721F5" box="[1285,1396,187,213]" pageId="13" pageNumber="316" value="1976-05-11">
<collectingDate id="EFEEE9963113FF9CFABFBBACD55721F5" box="[1285,1396,187,213]" pageId="13" pageNumber="316" value="1976-05-11">11.v.1976</collectingDate>
</date>
,
<collectorName id="26E153683113FF9CFA3BBBACD0F221D9" pageId="13" pageNumber="316">K. Baba</collectorName>
(
<collectionCode id="ED05AE7B3113FF9CFF5BBBC8D10021D9" box="[225,291,223,249]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="13" pageNumber="316" type="Herbarium">KPM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7C3CE33113FF9CFE8CBBC8D41721D9" box="[310,1076,222,249]" collectingDate="2005-05-11" collectorName="Hosomi-dani &amp; Yoshiwa &amp; I. Mori" country="Japan" location="Hatsukaichi-shi" pageId="13" pageNumber="316" specimenCount="3" specimenCount-male="3">
<specimenCount id="9D12FD373113FF9CFE8CBBC8D1AC21D9" box="[310,399,223,249]" count="3" pageId="13" pageNumber="316" type="male">3 males</specimenCount>
,
<collectorName id="26E153683113FF9CFE20BBC8D20C21D9" box="[410,559,223,249]" pageId="13" pageNumber="316">Hosomi-dani</collectorName>
,
<collectorName id="26E153683113FF9CFD83BBC8D2BE21D9" box="[569,669,223,249]" pageId="13" pageNumber="316">Yoshiwa</collectorName>
,
<location id="8ECB60653113FF9CFD12BBC8D37821D9" LSID="urn:lsid:plazi:treatment:03BD87A83115FF9CFF2DBA2AD32F20F1:8ECB60653113FF9CFD12BBC8D37821D9" box="[680,859,223,249]" country="Japan" name="Hatsukaichi-shi" pageId="13" pageNumber="316">Hatsukaichi-shi</location>
,
<date id="FFAA107E3113FF9CFCDCBBC9D3F621D9" box="[870,981,222,249]" pageId="13" pageNumber="316" value="2005-05-11">
<collectingDate id="EFEEE9963113FF9CFCDCBBC9D3F621D9" box="[870,981,222,249]" pageId="13" pageNumber="316" value="2005-05-11">11.v.2005</collectingDate>
</date>
,
<collectorName id="26E153683113FF9CFC5BBBC8D41721D9" box="[993,1076,223,249]" pageId="13" pageNumber="316">I. Mori</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353113FF9CFF7DBA14D1032061" pageId="13" pageNumber="316" type="etymology">
<paragraph id="8BAB36BE3113FF9CFF7DBA14D1032061" blockId="13.[151,1437,150,465]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF7DBA14D16D203D" bold="true" box="[199,334,259,285]" pageId="13" pageNumber="316">Etymology.</emphasis>
The species epithet (Latin adjective, whip-like) refers to the whip-like dorsal process of the phallotheca.
</paragraph>
</subSubSection>
<subSubSection id="C30E65353113FF9CFF7DBA5DD3202045" box="[199,771,330,357]" pageId="13" pageNumber="316" type="distribution">
<paragraph id="8BAB36BE3113FF9CFF7DBA5DD3202045" blockId="13.[151,1437,150,465]" box="[199,771,330,357]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF7DBA5DD17A2044" bold="true" box="[199,345,330,356]" pageId="13" pageNumber="316">Distribution</emphasis>
. Western Honshu (Chugoku District).
</paragraph>
</subSubSection>
<subSubSection id="C30E65353113FF9CFF7DBA79D32F20F1" pageId="13" pageNumber="316" type="biology_ecology">
<paragraph id="8BAB36BE3113FF9CFF7DBA79D24E208D" blockId="13.[151,1437,150,465]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF7DBA79D10420A8" bold="true" box="[199,295,366,392]" pageId="13" pageNumber="316">Biology.</emphasis>
Many adults of this species were collected in May using a Malaise trap over a small spring flow at the
<typeStatus id="54AF881C3113FF9CFF2DBA84D0E4208D" box="[151,199,403,429]" pageId="13" pageNumber="316">type</typeStatus>
locality (
<bibRefCitation id="EF854B4F3113FF9CFE89BA84D241208D" author="Kawase, N. &amp; Hayashi, M." box="[307,610,403,429]" pageId="13" pageNumber="316" pagination="83 - 86" refId="ref17291" refString="Kawase, N. &amp; Hayashi, M. (2010) Records of caddisflies from Mt. Sentsu-zan, Okuizumio-cho, Shimane Prefecture, Japan. Bulletin of Hoshizaki Green Foundation, 13, 83 - 86 [in Japanese with English abstract]." type="journal article" year="2010">Kawase and Hayashi 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8BAB36BE3113FF9CFF7DBAA0D32F20F1" blockId="13.[151,1437,150,465]" box="[199,780,439,465]" pageId="13" pageNumber="316">
<emphasis id="B960EAAC3113FF9CFF7DBAA0D1A020F1" bold="true" box="[199,387,439,465]" pageId="13" pageNumber="316">Japanese name.</emphasis>
Muchio-kimadara-shima-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,647 @@
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<treatment id="03BD87A8311CFF94FF2DBACCD2E52699" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699" httpUri="http://treatment.plazi.org/id/03BD87A8311CFF94FF2DBACCD2E52699" lastPageId="5" lastPageNumber="308" pageId="2" pageNumber="305">
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<taxonomicName id="4C144D3D311CFF93FF2DBACCD27320D5" authority="(Banks 1906)" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[151,592,475,502]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311CFF93FF2DBACCD27320D5" bold="true" box="[151,592,475,502]" pageId="2" pageNumber="305">
<emphasis id="B960EAAC311CFF93FF2DBACCD18620D5" bold="true" box="[151,421,475,502]" italics="true" pageId="2" pageNumber="305">Homoplectra japonica</emphasis>
(
<bibRefCitation id="EF854B4F311CFF93FE0EBACCD26B20D5" author="Banks, N." box="[436,584,475,501]" pageId="2" pageNumber="305" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E6535311CFF93FF2DBAE8D1CD2339" box="[151,494,511,537]" pageId="2" pageNumber="305" type="description">
<paragraph id="8BAB36BE311CFF93FF2DBAE8D1CD2339" blockId="2.[151,592,475,537]" box="[151,494,511,537]" pageId="2" pageNumber="305">
(
<figureCitation id="132F2A3B311CFF93FF25BAE8D10A2339" box="[159,297,511,537]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Figs 1A1E</figureCitation>
,
<figureCitation id="132F2A3B311CFF93FE8FBAE8D1672339" box="[309,324,511,537]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">3</figureCitation>
,
<figureCitation id="132F2A3B311CFF93FEEBBAE8D19E2339" box="[337,445,511,537]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="2" pageNumber="305">11A11E</figureCitation>
,
<figureCitation id="132F2A3B311CFF93FE72BAE8D1C72339" box="[456,484,511,537]" captionStart="FIGURE 13" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[169,1415,195,1904]" captionTargetId="figure-33@22.[157,1430,181,1919]" captionTargetPageId="22" captionText="FIGURE 13. Distributions of species of the Homoplectra japonica Species Group in Japan." pageId="2" pageNumber="305">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E6535311CFF93FF2DB953D4682398" pageId="2" pageNumber="305" type="reference_group">
<paragraph id="8BAB36BE311CFF93FF2DB953D39E237C" blockId="2.[151,1099,579,696]" box="[151,957,579,604]" pageId="2" pageNumber="305">
<taxonomicName id="4C144D3D311CFF93FF2DB953D1D0237B" authority="Banks 1906" authorityName="Banks" authorityYear="1906" box="[151,499,579,604]" class="Insecta" family="Hydropsychidae" genus="Arctopsyche" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311CFF93FF2DB953D150237B" box="[151,371,580,604]" italics="true" pageId="2" pageNumber="305">Arctopsyche japonica</emphasis>
<bibRefCitation id="EF854B4F311CFF93FEC3B953D1D0237B" author="Banks, N." box="[377,499,579,604]" pageId="2" pageNumber="305" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
</taxonomicName>
, pp 111, 113, color pattern of male forewing.
</paragraph>
<paragraph id="8BAB36BE311CFF93FF2DB975D415235A" blockId="2.[151,1099,579,696]" box="[151,1078,610,634]" pageId="2" pageNumber="305">
<taxonomicName id="4C144D3D311CFF93FF2DB975D1E9235A" ID-CoL="36JDB" authority="(Banks)" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[151,458,610,634]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311CFF93FF2DB975D153235A" box="[151,368,610,634]" italics="true" pageId="2" pageNumber="305">Diplectrona japonica</emphasis>
(Banks)
</taxonomicName>
:
<bibRefCitation id="EF854B4F311CFF93FE6EB975D273235A" author="Ulmer, G." box="[468,592,610,634]" pageId="2" pageNumber="305" pagination="1 - 102" refId="ref18285" refString="Ulmer, G. (1907) Trichopteren. Part. 1. Collection of Zoologiques du Baron Edm. de Selys Longchamps, 6 (1), 1 - 102." type="journal article" year="1907">Ulmer 1907</bibRefCitation>
, pp 7274, figs 114, 115, changed combination.
</paragraph>
<paragraph id="8BAB36BE311CFF93FF2DB996D46023B9" blockId="2.[151,1099,579,696]" box="[151,1091,641,665]" pageId="2" pageNumber="305">
<taxonomicName id="4C144D3D311CFF93FF2DB996D22023B9" ID-CoL="36JDB" authority="(Banks 1906)" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[151,515,641,665]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311CFF93FF2DB996D15323B9" box="[151,368,641,665]" italics="true" pageId="2" pageNumber="305">Diplectrona japonica</emphasis>
(
<bibRefCitation id="EF854B4F311CFF93FEC7B996D1D823B9" author="Banks, N." box="[381,507,641,665]" pageId="2" pageNumber="305" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="EF854B4F311CFF93FDB7B996D32923B9" author="Gyotoku, N. &amp; Nozaki, T." box="[525,778,641,665]" pageId="2" pageNumber="305" pagination="71 - 77" refId="ref16998" refString="Gyotoku, N. &amp; Nozaki, T. (1991) A list of Trichoptera of Fukuoka Prefecture, northern Kyushu, Japan 1. Kita Kyushu no Kontyu, 38, 71 - 77 + pl. 9 [in Japanese]." type="journal article" year="1991">Gyotoku &amp; Nozaki 1991</bibRefCitation>
, pp 7475, pl. 9, male (photo).
</paragraph>
<paragraph id="8BAB36BE311CFF93FF2DB9B7D4682398" blockId="2.[151,1099,579,696]" box="[151,1099,671,696]" pageId="2" pageNumber="305">
<taxonomicName id="4C144D3D311CFF93FF2DB9B7D2332397" authority="(Banks 1906)" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[151,528,671,696]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311CFF93FF2DB9B7D15F2397" box="[151,380,672,696]" italics="true" pageId="2" pageNumber="305">Homoplectra japonica</emphasis>
(
<bibRefCitation id="EF854B4F311CFF93FE33B9B7D2242397" author="Banks, N." box="[393,519,671,696]" pageId="2" pageNumber="305" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="EF854B4F311CFF93FDA3B9B7D2BE2398" author="Nozaki, T." box="[537,669,672,696]" pageId="2" pageNumber="305" pagination="245 - 258" refId="ref18077" refString="Nozaki, T. (2021) The genus Diplectrona Westwood (Trichoptera: Hydropsychidae) in the Japanese main islands. Zootaxa, 5082 (3), 245 - 258. https: // doi. org / 10.11646 / zootaxa. 5082.3.3" type="journal article" year="2021">Nozaki 2021</bibRefCitation>
, pp 254, 255, male, changed combination.
</paragraph>
</subSubSection>
<subSubSection id="C30E6535311CFF93FF2DB9F2D2BC2548" pageId="2" pageNumber="305" type="diagnosis">
<paragraph id="8BAB36BE311CFF93FF2DB9F2D3932204" blockId="2.[151,1437,741,1128]" pageId="2" pageNumber="305">
<emphasis id="B960EAAC311CFF93FF2DB9F2D13123DF" bold="true" box="[151,274,741,767]" pageId="2" pageNumber="305">Diagnosis.</emphasis>
The male of this species is very similar to that of
<taxonomicName id="4C144D3D311CFF93FC9EB9F1D4702220" authorityName="Nozaki" authorityYear="2025" box="[804,1107,742,768]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="flagelliformis" status="sp. nov.">
<emphasis id="B960EAAC311CFF93FC9EB9F1D4702220" box="[804,1107,742,768]" italics="true" pageId="2" pageNumber="305">Homoplectra flagelliformis</emphasis>
</taxonomicName>
<emphasis id="B960EAAC311CFF93FBEDB9F1D48C2220" bold="true" box="[1111,1199,742,768]" pageId="2" pageNumber="305">
<taxonomicNameLabel id="A25357D7311CFF93FBEDB9F1D48C2220" box="[1111,1199,742,768]" pageId="2" pageNumber="305" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but is distinguishable from the latter by the characters given in the diagnosis for that species.
</paragraph>
<paragraph id="8BAB36BE311CFF93FF7DB839D09C2294" blockId="2.[151,1437,741,1128]" pageId="2" pageNumber="305">
The female is easily distinguishable from those of other Japanese species by the shape of a pair of large round basoventral lobes on the vulvar scale (
<figureCitation id="132F2A3B311CFF93FDF6B845D2F2224C" box="[588,721,850,876]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Figs 3H, 3I</figureCitation>
). Furthermore, female segment IX of this species has a pair of unique ventrolateral sclerotized plates that are oblong in dorsal and lateral aspects (marked with arrows in
<figureCitation id="132F2A3B311CFF93FA86B861D0922294" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Figs 3G, 3I</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE311CFF93FF7DB8A9D2BC2548" blockId="2.[151,1437,741,1128]" pageId="2" pageNumber="305">
The larva is distinguishable from known Japanese larvae by the shape of anterior margin of the frontoclypeal apotome:Asymmetrical and sinuous in this species (
<figureCitation id="132F2A3B311CFF93FD62B8F5D35F22DC" box="[728,892,994,1020]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="2" pageNumber="305">Figs 11A, 11B</figureCitation>
); but symmetrical in
<taxonomicName id="4C144D3D311CFF93FBD9B8F5D0EC2500" authority="Nozaki 2019" authorityName="Nozaki" authorityYear="2019" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="crassa">
<emphasis id="B960EAAC311CFF93FBD9B8F5D56522DC" box="[1123,1350,994,1020]" italics="true" pageId="2" pageNumber="305">Homoplectra crassa</emphasis>
<bibRefCitation id="EF854B4F311CFF93FAF1B8F5D0EC2500" author="Nozaki, T." pageId="2" pageNumber="305" pagination="329 - 344" refId="ref18024" refString="Nozaki, T. (2019) Description of five new species Homoplectra Ross (Trichoptera, Hydropsychidae) from Japan with reassignment of Homoplectra tohokuensis (Kobayashi). Zootaxa, 4608 (2), 329 - 344. https: // doi. org / 10.11646 / zootaxa. 4608.2.7" type="journal article" year="2019">Nozaki 2019</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C144D3D311CFF93FEBCBF11D29D2500" authority="Nozaki 2019" authorityName="Nozaki" authorityYear="2019" box="[262,702,1030,1056]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="tohokuensis">
<emphasis id="B960EAAC311CFF93FEBCBF11D2052500" box="[262,550,1030,1056]" italics="true" pageId="2" pageNumber="305">Homoplectra tohokuensis</emphasis>
<bibRefCitation id="EF854B4F311CFF93FD97BF11D29D2500" author="Nozaki, T." box="[557,702,1030,1056]" pageId="2" pageNumber="305" pagination="329 - 344" refId="ref18024" refString="Nozaki, T. (2019) Description of five new species Homoplectra Ross (Trichoptera, Hydropsychidae) from Japan with reassignment of Homoplectra tohokuensis (Kobayashi). Zootaxa, 4608 (2), 329 - 344. https: // doi. org / 10.11646 / zootaxa. 4608.2.7" type="journal article" year="2019">Nozaki 2019</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF854B4F311CFF93FD77BF11D3412500" author="Nozaki, T." box="[717,866,1030,1056]" pageId="2" pageNumber="305" pagination="329 - 344" refId="ref18024" refString="Nozaki, T. (2019) Description of five new species Homoplectra Ross (Trichoptera, Hydropsychidae) from Japan with reassignment of Homoplectra tohokuensis (Kobayashi). Zootaxa, 4608 (2), 329 - 344. https: // doi. org / 10.11646 / zootaxa. 4608.2.7" type="journal article" year="2019">Nozaki 2019</bibRefCitation>
, figs 7A7L, 8), bearing a small median triangular process in
<taxonomicName id="4C144D3D311CFF93FEB5BF3DD2B52564" authority="Nozaki 2019" authorityName="Nozaki" authorityYear="2019" box="[271,662,1066,1092]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B960EAAC311CFF93FEB5BF3DD1DC2564" box="[271,511,1066,1092]" italics="true" pageId="2" pageNumber="305">Homoplectra gracilis</emphasis>
<bibRefCitation id="EF854B4F311CFF93FDBCBF3DD2B52564" author="Nozaki, T." box="[518,662,1066,1092]" pageId="2" pageNumber="305" pagination="329 - 344" refId="ref18024" refString="Nozaki, T. (2019) Description of five new species Homoplectra Ross (Trichoptera, Hydropsychidae) from Japan with reassignment of Homoplectra tohokuensis (Kobayashi). Zootaxa, 4608 (2), 329 - 344. https: // doi. org / 10.11646 / zootaxa. 4608.2.7" type="journal article" year="2019">Nozaki 2019</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF854B4F311CFF93FD1CBF3DD3482564" author="Kochi, K. &amp; Higashida, S. &amp; Iijima, R. &amp; Nozaki, T." box="[678,875,1066,1092]" pageId="2" pageNumber="305" pagination="284 - 296" refId="ref17475" refString="Kochi, K., Higashida, S., Iijima, R. &amp; Nozaki, T. (2023) Morphology and biology of Homoplectra gracilis Nozaki 2019 in a headwater stream in Yata Hills, Japan. Aquatic Insects, 44, 284 - 296. https: // doi. org / 10.1080 / 01650424.2023.2212647" type="journal article" year="2023">
<collectingCountry id="F303762E311CFF93FD1CBF3DD2C92564" box="[678,746,1066,1092]" name="India" pageId="2" pageNumber="305">Kochi</collectingCountry>
<emphasis id="B960EAAC311CFF93FD4BBF3CD3012564" box="[753,802,1066,1092]" italics="true" pageId="2" pageNumber="305">et al</emphasis>
. 2023
</bibRefCitation>
, figs
<quantity id="4CEC9B5B311CFF93FC1DBF3DD3E62564" box="[935,965,1066,1092]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="2" pageNumber="305" unit="g" value="3.0">3g</quantity>
, 3i), and asymmetrical and evenly convex mesally in
<taxonomicName id="4C144D3D311CFF93FEA8BF58D1FA2548" baseAuthorityName="Ulmer" baseAuthorityYear="1907" box="[274,473,1102,1128]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="albomarginata">
<emphasis id="B960EAAC311CFF93FEA8BF58D1FA2548" box="[274,473,1102,1128]" italics="true" pageId="2" pageNumber="305">H. albomarginata</emphasis>
</taxonomicName>
(
<figureCitation id="132F2A3B311CFF93FE53BF59D2B12548" box="[489,658,1102,1128]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="2" pageNumber="305">Figs 12A, 12B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E6535311CFF94FF2DBFAED47225E5" lastPageId="5" lastPageNumber="308" pageId="2" pageNumber="305" type="description">
<paragraph id="8BAB36BE311CFF93FF2DBFAED10B25F3" blockId="2.[151,296,1209,1235]" box="[151,296,1209,1235]" pageId="2" pageNumber="305">
<emphasis id="B960EAAC311CFF93FF2DBFAED10B25F3" bold="true" box="[151,296,1209,1235]" pageId="2" pageNumber="305">Description</emphasis>
</paragraph>
<paragraph id="8BAB36BE311CFF93FF2DBE16D1442788" blockId="2.[151,1437,1281,2028]" pageId="2" pageNumber="305">
<emphasis id="B960EAAC311CFF93FF2DBE16D0F8243B" bold="true" box="[151,219,1281,1307]" pageId="2" pageNumber="305">Adult</emphasis>
(
<figureCitation id="132F2A3B311CFF93FF53BE15D150243C" box="[233,371,1282,1308]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Figs 1A1E</figureCitation>
). Head and thorax mostly dark brown dorsally, but dorsal setal warts of head, pro-, and mesonota with golden hair; face light brown in male, dark brown in female; compound eyes slightly larger in male than those in female (
<figureCitation id="132F2A3B311CFF93FEAABE5DD1BA2444" box="[272,409,1354,1380]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Figs 1C, 1E</figureCitation>
). Legs mostly pale yellow to light brown; but middle and hind coxae of male and all coxae of female dark brown. Abdomen mostly dark brown, but pleural membranes pale yellow. Antennae light brown in male, dark brown in female, shorter than forewings; scapes thick, approximately as long as wide, with long hair-like setae dorsally (
<figureCitation id="132F2A3B311CFF93FEFABEA1D1EB24F0" box="[320,456,1462,1488]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Figs 1C, 1E</figureCitation>
); pedicels shorter than scapes, basal segment of each flagellum longer than scape (
<figureCitation id="132F2A3B311CFF93FAD0BEA1D0C524D4" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Figs 1C, 1E</figureCitation>
). Wing venation similar in male and female. Forewings each
<quantity id="4CEC9B5B311CFF93FC31BECDD43224D4" box="[907,1041,1498,1525]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.5" metricValueMin="6.5" pageId="2" pageNumber="305" unit="mm" value="8.0" valueMax="9.5" valueMin="6.5">6.59.5 mm</quantity>
long in male (n = 11), 9.0
<quantity id="4CEC9B5B311CFF93FA86BECDD5BE24D4" box="[1340,1437,1498,1525]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="305" unit="mm" value="11.5">11.5 mm</quantity>
long in female (n = 10),
<quantity id="4CEC9B5B311CFF93FE18BEE9D1D42738" box="[418,503,1534,1561]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="305" unit="mm" value="6.5">6.5 mm</quantity>
in male
<typeStatus id="54AF881C311CFF93FDEFBEE9D29C2738" box="[597,703,1534,1560]" pageId="2" pageNumber="305" type="lectotype">lectotype</typeStatus>
; yellow, with broad dark stripes (
<figureCitation id="132F2A3B311CFF93FB94BEE9D4A52738" box="[1070,1158,1534,1560]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Fig. 1A</figureCitation>
); hind wings pale yellow with dark stripes (
<figureCitation id="132F2A3B311CFF93FEE4BD35D195271C" box="[350,438,1570,1596]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Fig. 1A</figureCitation>
); venation of both wings as in
<figureCitation id="132F2A3B311CFF93FCB3BD35D35F271C" box="[777,892,1570,1596]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Figure 1B</figureCitation>
: with forks I, II, III, IV, and V in forewings and I, II, III, and V in hind wings; discoidal and medial cells closed in both wings. Tibial spurs 2-4-4. Abdominal sternum V with pair of finger-like processes associated with scent glands near anterolateral margins, slightly shorter than sternum (
<figureCitation id="132F2A3B311CFF93FEBBBD99D1792788" box="[257,346,1678,1704]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="2" pageNumber="305">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE311CFF92FF7DBDA5D3A52061" blockId="2.[151,1437,1281,2028]" lastBlockId="3.[151,1436,151,321]" lastPageId="3" lastPageNumber="306" pageId="2" pageNumber="305">
<emphasis id="B960EAAC311CFF93FF7DBDA5D15627EB" bold="true" box="[199,373,1713,1740]" pageId="2" pageNumber="305">Male genitalia</emphasis>
(
<figureCitation id="132F2A3B311CFF93FE3CBDA5D23627EC" box="[390,533,1714,1740]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Figs 3A3E</figureCitation>
). Ventrolateral sides of synsclerotized segment IX (IX) subtriangular in lateral aspect (
<figureCitation id="132F2A3B311CFF93FF57BDC1D16B27D0" box="[237,328,1750,1776]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Fig. 3A</figureCitation>
); posteroventral lobe (p.l.) trapezoidal in ventral aspect, with shallow median concavity (
<figureCitation id="132F2A3B311CFF93FA8FBDC1D5AD27D0" box="[1333,1422,1750,1776]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Fig. 3C</figureCitation>
); dorsal part of segment IX triangular with round and setose apex in dorsal aspect (
<figureCitation id="132F2A3B311CFF93FB8FBDEDD4AD2634" box="[1077,1166,1786,1812]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Fig. 3B</figureCitation>
), fused with segment X (X) laterally (
<figureCitation id="132F2A3B311CFF93FE8BBC09D1AF2618" box="[305,396,1822,1848]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Fig. 3A</figureCitation>
). Segment X bilobed in dorsal aspect (
<figureCitation id="132F2A3B311CFF93FCFABC09D3B42618" box="[832,919,1822,1848]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Fig. 3B</figureCitation>
), with pair of large posterolateral setose areas (
<figureCitation id="132F2A3B311CFF93FF25BC55D108267C" box="[159,299,1858,1884]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Figs 3A, 3B</figureCitation>
); each ventrolateral margin sclerotized, recessed into segment IX (
<figureCitation id="132F2A3B311CFF93FBAEBC55D44D267C" box="[1044,1134,1858,1884]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Fig. 3A</figureCitation>
). Inferior appendages (i.a.) each without distal segment, long and finger-like in lateral aspect, dorsal margin weakly concave at apical 1/
<quantity id="4CEC9B5B311CFF93FAD5BC71D5BF26A0" box="[1391,1436,1894,1921]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="305" unit="in" value="5.0">5 in</quantity>
lateral aspect, extending beyond apex of segment X (
<figureCitation id="132F2A3B311CFF93FD43BC9DD3772684" box="[761,852,1930,1956]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Fig. 3A</figureCitation>
); apex slightly enlarged mesally and truncate, with several short spine-like setae (
<figureCitation id="132F2A3B311CFF93FE53BCB9D25426E8" box="[489,631,1966,1992]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Figs 3A3C</figureCitation>
). Basal plate of inferior appendages (b.p.i.a) sclerotized, subrectangular in ventral aspect (
<figureCitation id="132F2A3B311CFF93FED8BCC5D19F26CC" box="[354,444,2002,2028]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="2" pageNumber="305">Fig. 3C</figureCitation>
). Phallotheca with dorsal process (d.p.p.), acicular in dorsal and lateral aspects, weakly curved ventrad (
<figureCitation id="132F2A3B311DFF92FEE9BB80D1C72191" box="[339,484,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="3" pageNumber="306">Figs 3D, 3E</figureCitation>
); with pair of ventral processes (v.p.p), long, approximately same length as dorsal process of phallotheca, each apex acute, curved dorsad and slightly mesad (
<figureCitation id="132F2A3B311DFF92FC40BBACD4AA21F5" box="[1018,1161,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="3" pageNumber="306">Figs 3D, 3E</figureCitation>
). Aedeagus (ae.) arising from membranous endotheca (end.), evenly curved ventrad, with long head approximately 1/4 as long as basal stem (
<figureCitation id="132F2A3B311DFF92FF25BA14D0DB203D" box="[159,248,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="3" pageNumber="306">Fig. 3D</figureCitation>
); stem with lateral flanges, half-pipe-like, narrow at apical 3/4 and near base in dorsal aspect (
<figureCitation id="132F2A3B311DFF92FAB8BA14D574203D" box="[1282,1367,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="3" pageNumber="306">Fig. 3E</figureCitation>
), with pair of apical swellings in dorsal and ventral aspects (
<figureCitation id="132F2A3B311DFF92FD55BA30D35A2061" box="[751,889,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="3" pageNumber="306">Figs 3C, 3E</figureCitation>
).
</paragraph>
<caption id="DF6B6636311DFF92FF2DBD4DD29E2606" pageId="3" pageNumber="306" startId="3.[151,264,1626,1650]" targetBox="[157,1429,351,1596]" targetPageId="3" targetType="figure">
<paragraph id="8BAB36BE311DFF92FF2DBD4DD29E2606" blockId="3.[151,1437,1626,1830]" pageId="3" pageNumber="306">
<emphasis id="B960EAAC311DFF92FF2DBD4DD4732752" bold="true" box="[151,1104,1626,1650]" pageId="3" pageNumber="306">
FIGURES 12. Adults of
<taxonomicName id="4C144D3D311DFF92FE17BD4DD25F2752" box="[429,636,1626,1650]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="306" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B960EAAC311DFF92FE17BD4DD2152752" bold="true" box="[429,566,1626,1650]" italics="true" pageId="3" pageNumber="306">Homoplectra</emphasis>
spp. 1
</taxonomicName>
A1E,
<taxonomicName id="4C144D3D311DFF92FD72BD4DD4682752" authority="(Banks 1906)" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[712,1099,1626,1650]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="306" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311DFF92FD72BD4DD3922752" bold="true" box="[712,945,1626,1650]" italics="true" pageId="3" pageNumber="306">Homoplectra japonica</emphasis>
(
<bibRefCitation id="EF854B4F311DFF92FC7ABD4DD4602752" author="Banks, N." box="[960,1091,1626,1650]" pageId="3" pageNumber="306" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
)
</taxonomicName>
:
</emphasis>
1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same).
<emphasis id="B960EAAC311DFF92FCA8BDB5D50F279A" bold="true" box="[786,1324,1698,1722]" pageId="3" pageNumber="306">
2A2C,
<taxonomicName id="4C144D3D311DFF92FCD2BDB5D50B279A" authority="(Ulmer 1907)" baseAuthorityName="Ulmer" baseAuthorityYear="1907" box="[872,1320,1698,1722]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="306" phylum="Arthropoda" rank="species" species="albomarginata">
<emphasis id="B960EAAC311DFF92FCD2BDB5D4B3279A" bold="true" box="[872,1168,1698,1722]" italics="true" pageId="3" pageNumber="306">Homoplectra albomarginata</emphasis>
(
<bibRefCitation id="EF854B4F311DFF92FB27BDB5D503279A" author="Ulmer, G." box="[1181,1312,1698,1722]" pageId="3" pageNumber="306" pagination="1 - 102" refId="ref18285" refString="Ulmer, G. (1907) Trichopteren. Part. 1. Collection of Zoologiques du Baron Edm. de Selys Longchamps, 6 (1), 1 - 102." type="journal article" year="1907">Ulmer 1907</bibRefCitation>
)
</taxonomicName>
:
</emphasis>
2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm.
</paragraph>
</caption>
<caption id="DF6B6636311AFF95FF2DBD5AD17427F1" pageId="4" pageNumber="307" startId="4.[151,250,1613,1637]" targetBox="[157,1429,185,1583]" targetPageId="4" targetType="figure">
<paragraph id="8BAB36BE311AFF95FF2DBD5AD17427F1" blockId="4.[151,1437,1613,1745]" pageId="4" pageNumber="307">
<emphasis id="B960EAAC311AFF95FF2DBD5AD3352745" bold="true" box="[151,790,1613,1637]" pageId="4" pageNumber="307">
FIGURE 3. Adults of
<taxonomicName id="4C144D3D311AFF95FE31BD5AD3322745" authority="(Banks 1906)" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[395,785,1613,1637]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311AFF95FE31BD5AD2542745" bold="true" box="[395,631,1613,1637]" italics="true" pageId="4" pageNumber="307">Homoplectra japonica</emphasis>
(
<bibRefCitation id="EF854B4F311AFF95FD3DBD5AD32A2745" author="Banks, N." box="[647,777,1613,1637]" pageId="4" pageNumber="307" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks 1906</bibRefCitation>
)
</taxonomicName>
.
</emphasis>
3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text.
</paragraph>
</caption>
<paragraph id="8BAB36BE311AFF94FF7DBC34D3FD2061" blockId="4.[151,1437,1827,2034]" lastBlockId="5.[151,1437,151,1977]" lastPageId="5" lastPageNumber="308" pageId="4" pageNumber="307">
<emphasis id="B960EAAC311AFF95FF7DBC34D1AD261D" bold="true" box="[199,398,1827,1853]" pageId="4" pageNumber="307">Female genitalia</emphasis>
(
<figureCitation id="132F2A3B311AFF95FE25BC34D202261D" box="[415,545,1827,1853]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Figs 3F3I</figureCitation>
). Sternum VIII widely separated into pair of lateral lobes (l.l.) ventrally, each lobe bean-shaped in ventral aspect (
<figureCitation id="132F2A3B311AFF95FD8FBC50D2B12641" box="[565,658,1863,1889]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Fig. 3H</figureCitation>
). Segment IX (IX) obliquely rectangular in lateral aspect (
<figureCitation id="132F2A3B311AFF95FA8DBC50D5AC2641" box="[1335,1423,1863,1889]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Fig. 3F</figureCitation>
), posterodorsal part trapezoidal in dorsal aspect (
<figureCitation id="132F2A3B311AFF95FD1ABC7CD2DA26A5" box="[672,761,1899,1925]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Fig. 3G</figureCitation>
); tergum with pair of ventrolateral sclerotized plates (marked with arrows in
<figureCitation id="132F2A3B311AFF95FE87BC98D19F2689" box="[317,444,1935,1961]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Figs 3G, 3I</figureCitation>
), weakly constricted at mid-length in lateral aspect (
<figureCitation id="132F2A3B311AFF95FC45BC98D46F2689" box="[1023,1100,1935,1961]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Fig. 3I</figureCitation>
), oblong in dorsal aspect (
<figureCitation id="132F2A3B311AFF95FAD5BC98D09E26ED" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Fig. 3G</figureCitation>
); each mesal lobe (m.l.IX) sclerotized, bean-shape in lateral aspect (
<figureCitation id="132F2A3B311AFF95FC0DBCA4D41026ED" box="[951,1075,1971,1997]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Figs 3F, 3I</figureCitation>
), forming large crevice between it and ventrolateral plate in ventral aspect (marked with arrow and asterisk in
<figureCitation id="132F2A3B311AFF95FC40BCC0D47026D1" box="[1018,1107,2007,2033]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="4" pageNumber="307">Fig. 3H</figureCitation>
); pair of sclerotized ribs (s.r.) forming large round plate-like lobes on vulvar scale (v.s.) basoventrally (
<figureCitation id="132F2A3B311BFF94FC68BB80D4752191" box="[978,1110,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="5" pageNumber="308">Figs 3H, 3I</figureCitation>
). Vulvar scale large, tongue-like in ventral aspect (
<figureCitation id="132F2A3B311BFF94FE2EBBACD1CC21F5" box="[404,495,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="5" pageNumber="308">Fig. 3H</figureCitation>
), semimembranous apically. Segment X slender in lateral aspect (
<figureCitation id="132F2A3B311BFF94FB61BBACD51121F5" box="[1243,1330,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="5" pageNumber="308">Fig. 3F</figureCitation>
). Vaginal apparatus (v.a.) long-pentagonal in dorsal aspect (
<figureCitation id="132F2A3B311BFF94FD76BBC8D30521D9" box="[716,806,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1613,1637]" captionTargetBox="[157,1429,185,1583]" captionTargetId="figure-33@4.[151,1435,181,1589]" captionTargetPageId="4" captionText="FIGURE 3. Adults of Homoplectra japonica (Banks 1906). 3A3E, male genitalia, (Mt. Hiko, Fukuoka): 3A, left lateral, apicomesal part of inferior appendage enlarged; 3B, dorsal; 3C, ventral; 3D phallic apparatus, left lateral; 3E, same, dorsal. 3F3H, female genitalia (Mt. Hiko, Fukuoka): 3F, left lateral; 3G, dorsal; 3H, ventral; 3I, left ventrolateral. Arrows on Figs 3G, 3H and 3I see text." pageId="5" pageNumber="308">Fig. 3G</figureCitation>
), tapering to duct of bursa copulatrix (d.b.c.); with pair of peanut shape sclerites posterodorsally, with pair of longitudinal ridges dorsally, and with dark arched marking anterodorsally, partially surrounding opening of duct of spermatheca (d.s.).
</paragraph>
<paragraph id="8BAB36BE311BFF94FF7DBA5DD27E2341" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="308">
<emphasis id="B960EAAC311BFF94FF7DBA5DD1B72045" bold="true" box="[199,404,330,357]" pageId="5" pageNumber="308">Final instar larva</emphasis>
(
<figureCitation id="132F2A3B311BFF94FE18BA5CD25E2045" box="[418,637,331,357]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="5" pageNumber="308">Figs 11A, 11B, 11F</figureCitation>
). Head
<quantity id="4CEC9B5B311BFF94FD6ABA5CD3072045" box="[720,804,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="308" unit="mm" value="1.6">1.6 mm</quantity>
wide (n = 1), approximately as long as wide (
<figureCitation id="132F2A3B311BFF94FAA2BA5CD5562045" box="[1304,1397,331,357]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="5" pageNumber="308">Fig. 11F</figureCitation>
, hl, hw); frontoclypeal apotome asymmetrical in dorsal aspect (
<figureCitation id="132F2A3B311BFF94FC9DBA78D3AB20A9" box="[807,904,367,393]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="5" pageNumber="308">Fig. 11B</figureCitation>
inset), maximum width
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, posterior angle about 90°, with anterior margin sinuous, left side recessed, very weakly concave in middle, with most secondary setae clavate. Mandibles each with 5 teeth. Pronotum with transverse sulcus on posterior 1/4 (
<figureCitation id="132F2A3B311BFF94FB6FBAA0D51C20F1" box="[1237,1343,439,465]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="5" pageNumber="308">Fig. 11A</figureCitation>
). Meso- and metathorax and abdominal segments bearing gills, with one pair of ventral tufts of gills on mesothorax and abdominal segment VII, with two pairs of subventral tufts of gills on metathorax and each of abdominal segments I to VI, with 13 sublateral conical gills on each of abdominal segments III to VII. Secondary setae on thoracic nota and abdominal segments mostly clavate.
</paragraph>
<paragraph id="8BAB36BE311BFF94FF7DB97CD4C92219" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="308">
<emphasis id="B960EAAC311BFF94FF7DB97CD12623A5" bold="true" box="[199,261,619,645]" pageId="5" pageNumber="308">Pupa</emphasis>
(
<figureCitation id="132F2A3B311BFF94FEADB97CD19D23A5" box="[279,446,619,645]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="5" pageNumber="308">Figs 11C11E</figureCitation>
). Body length
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(n = 1). Mandibles slender, with 3 (right) or 4 (left) apical and subapical teeth, each mandible with stout mesal plate-like projection on apical 1/3. Middle tarsi bearing sparse hair-like setae. Abdominal segments with dorsal hook plates anterior on II to VIII, posterior on III and IV (
<figureCitation id="132F2A3B311BFF94FA97B9A4D0E923D1" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="5" pageNumber="308">Figs 11C, 11D</figureCitation>
). Abdominal segments bearing gills, with one pair of ventral tufts of gills on each of abdominal segments II to VI, with lateral conical gills on each of abdominal segments III to VII. Anal processes sclerotized, ventral surfaces covered by tiny spines; bifurcated, each apex acute, mesal branch longer than outer one (
<figureCitation id="132F2A3B311BFF94FBC3B808D4FE2219" box="[1145,1245,799,825]" captionStart="FIGURES 1112" captionStartId="21.[151,264,1662,1686]" captionTargetBox="[157,1429,187,1632]" captionTargetId="figure-19@21.[151,1435,181,1638]" captionTargetPageId="21" captionText="FIGURES 1112. Larva and pupa of Homoplectra spp. 11A11E, Homoplectra japonica (Banks 1906): 11A, larva, head, pro-, meso-, and metanota, dorsal (Mt. Hiko, Fukuoka); 11B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 11C, pupa, body, dorsal (same); 11D, pupa, right hook plates, dorsal (same); 11E, pupa, anal processes, ventral (same); 11F, larval head of the dorsal aspect showing measurement points of length and width. 12A12C, Homoplectra albomarginata (Ulmer 1907): 12A, larva, head, dorsal (Ibigawa-cho, Gifu); 12B, larva, frontoclypeal apotome, dorsal, secondary setae enlarged (same); 12C, pupa, anal processes, ventral (same).Abbreviations: a = anterior, hl = head length, hw = head width, p = posterior, IIVIII = abdominal segments IIVIII." pageId="5" pageNumber="308">Fig. 11E</figureCitation>
).
</paragraph>
<paragraph id="8BAB36BE311BFF94FF7DB854D29222E9" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="308">
<materialsCitation id="3B7C3CE3311BFF94FF7DB854D29222E9" collectionCode="MCZ, HIST" collectorName="Collection N. Banks &amp; Det. H. H. Ross" country="Japan" determinerName="Det. H. H. Ross" location="Japan" pageId="5" pageNumber="308" specimenCount="1" typeStatus="lectotype">
<emphasis id="B960EAAC311BFF94FF7DB854D164227D" bold="true" box="[199,327,835,861]" pageId="5" pageNumber="308">
<typeStatus id="54AF881C311BFF94FF7DB854D161227D" box="[199,322,835,861]" pageId="5" pageNumber="308" type="lectotype">Lectotype</typeStatus>
:
</emphasis>
Male (here designated, pinned:
<collectionCode id="ED05AE7B311BFF94FD6CB854D33A227D" box="[726,793,835,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="308" type="Museum">MCZ</collectionCode>
:Ent:11833): labeled “type”, “Hikosan, Buzen,
<collectingCountry id="F303762E311BFF94FAF3B854D5AD227D" box="[1353,1422,835,861]" name="Japan" pageId="5" pageNumber="308">Japan</collectingCountry>
”, “
<collectorName id="26E15368311BFF94FF1BB870D1B722A1" box="[161,404,871,897]" pageId="5" pageNumber="308">Collection N. Banks</collectorName>
”, “
<taxonomicName id="4C144D3D311BFF94FE00B870D2DC22A1" authority="Banks" authorityName="Banks" authorityYear="1906" box="[442,767,871,897]" class="Insecta" family="Hydropsychidae" genus="Arctopsyche" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311BFF94FE00B870D29322A0" box="[442,688,871,897]" italics="true" pageId="5" pageNumber="308">Arctopsyche japonica</emphasis>
Banks
</taxonomicName>
/type”, “Type 11833”, “
<taxonomicName id="4C144D3D311BFF94FBA8B870D4B822A1" box="[1042,1179,871,897]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
:
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<emphasis id="B960EAAC311BFF94FB13B870D5BF22A0" box="[1193,1436,871,897]" italics="true" pageId="5" pageNumber="308">Diplectrona japonica</emphasis>
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wings from Type/
<determinerName id="7E77DA7A311BFF94FED9B89CD22C2285" box="[355,527,907,933]" pageId="5" pageNumber="308">
<collectorName id="26E15368311BFF94FED9B89CD22C2285" box="[355,527,907,933]" pageId="5" pageNumber="308">Det. H.H. Ross</collectorName>
</determinerName>
, 19/ILL. NAT.
<collectionCode id="ED05AE7B311BFF94FD05B89CD2DC2285" box="[703,767,907,933]" pageId="5" pageNumber="308">HIST</collectionCode>
. SUR (right wings in slide)” (https://mczbase.mcz.harvard. edu/guid/
<collectionCode id="ED05AE7B311BFF94FEBDB8B8D16422E9" box="[263,327,943,969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="308" type="Museum">MCZ</collectionCode>
:Ent:11833;
<bibRefCitation id="EF854B4F311BFF94FE72B8B8D27822E9" author="Nozaki, T." box="[456,603,943,969]" pageId="5" pageNumber="308" pagination="245 - 258" refId="ref18077" refString="Nozaki, T. (2021) The genus Diplectrona Westwood (Trichoptera: Hydropsychidae) in the Japanese main islands. Zootaxa, 5082 (3), 245 - 258. https: // doi. org / 10.11646 / zootaxa. 5082.3.3" type="journal article" year="2021">Nozaki 2021</bibRefCitation>
, fig. 5).
</materialsCitation>
</paragraph>
<paragraph id="8BAB36BE311BFF94FF7DB8C5D47225E5" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="308">
<emphasis id="B960EAAC311BFF94FF7DB8C5D1E922CC" bold="true" box="[199,458,978,1004]" pageId="5" pageNumber="308">Specimens examined.</emphasis>
<materialsCitation id="3B7C3CE3311BFF94FE6FB8C5D1A12579" collectingDate="1935-05-31" collectingDateMax="1994-06-09" collectingDateMin="1935-05-31" collectionCode="KPM, T" collectorName="Yamauchi" country="Japan" county="Nankai" location="Yamaguchi-shi" municipality="Atojifukushimo" pageId="5" pageNumber="308" specimenCount="33" specimenCount-female="14" specimenCount-male="19" stateProvince="Yamaguchi">
<emphasis id="B960EAAC311BFF94FE6FB8C5D37D22CC" bold="true" box="[469,862,978,1004]" pageId="5" pageNumber="308">
<collectingCountry id="F303762E311BFF94FE6FB8C5D21022CC" box="[469,563,978,1004]" name="Japan" pageId="5" pageNumber="308">JAPAN</collectingCountry>
: HONSHU:
<collectingRegion id="49D0F85C311BFF94FD75B8C5D37A22CC" box="[719,857,978,1004]" country="Japan" name="Yamaguchi" pageId="5" pageNumber="308">Yamaguchi</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD37311BFF94FCD3B8C4D39822CD" box="[873,955,979,1005]" count="1" pageId="5" pageNumber="308" type="male">1 male</specimenCount>
,
<collectingCounty id="62CA4E32311BFF94FC70B8C4D43D22CD" box="[970,1054,979,1005]" pageId="5" pageNumber="308">Nankai</collectingCounty>
,
<collectingMunicipality id="6BCFACC4311BFF94FB91B8C4D4FC22CD" box="[1067,1247,979,1005]" pageId="5" pageNumber="308">Atojifukushimo</collectingMunicipality>
,
<location id="8ECB6065311BFF94FB57B8C4D5BB22CD" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FB57B8C4D5BB22CD" box="[1261,1432,979,1005]" country="Japan" county="Nankai" municipality="Atojifukushimo" name="Yamaguchi-shi" pageId="5" pageNumber="308" stateProvince="Yamaguchi">Yamaguchi-shi</location>
,
<date id="FFAA107E311BFF94FF2DB8E0D1242531" box="[151,263,1015,1041]" pageId="5" pageNumber="308" value="1993-05-17">
<collectingDate id="EFEEE996311BFF94FF2DB8E0D1242531" box="[151,263,1015,1041]" pageId="5" pageNumber="308" value="1993-05-17">17.v.1993</collectingDate>
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, N.
<location id="8ECB6065311BFF94FEFBB8E0D1B82531" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FEFBB8E0D1B82531" box="[321,411,1015,1041]" country="Japan" county="Nankai" municipality="Atojifukushimo" name="Kuhara." pageId="5" pageNumber="308" stateProvince="Yamaguchi">Kuhara.</location>
<emphasis id="B960EAAC311BFF94FE1DB8E1D28A2531" bold="true" box="[423,681,1014,1041]" pageId="5" pageNumber="308">
KYUSHU:
<collectingRegion id="49D0F85C311BFF94FD8EB8E0D2872531" box="[564,676,1015,1041]" country="Japan" name="Fukuoka" pageId="5" pageNumber="308">Fukuoka</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD37311BFF94FD0FB8E0D33C2531" box="[693,799,1015,1041]" count="1" pageId="5" pageNumber="308" type="female">1 female</specimenCount>
,
<location id="8ECB6065311BFF94FC8AB8E0D3ED2531" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FC8AB8E0D3ED2531" box="[816,974,1015,1041]" country="Japan" county="Nankai" municipality="Atojifukushimo" name="Mt. Hiko-san" pageId="5" pageNumber="308" stateProvince="Yamaguchi">Mt. Hiko-san</location>
,
<location id="8ECB6065311BFF94FC64B8E0D4502531" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FC64B8E0D4502531" box="[990,1139,1015,1041]" country="Japan" county="Nankai" municipality="Atojifukushimo" name="Soeda-machi" pageId="5" pageNumber="308" stateProvince="Yamaguchi">Soeda-machi</location>
,
<date id="FFAA107E311BFF94FB3EB8E1D4D72531" box="[1156,1268,1014,1041]" pageId="5" pageNumber="308" value="1935-05-31">
<collectingDate id="EFEEE996311BFF94FB3EB8E1D4D72531" box="[1156,1268,1014,1041]" pageId="5" pageNumber="308" value="1935-05-31">31.v.1935</collectingDate>
</date>
, K.
<collectorName id="26E15368311BFF94FA96B8E0D5BF2531" box="[1324,1436,1015,1041]" pageId="5" pageNumber="308">Yamauchi</collectorName>
(
<collectionCode id="ED05AE7B311BFF94FF1ABF0CD0C12515" box="[160,226,1051,1077]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="5" pageNumber="308" type="Herbarium">KPM</collectionCode>
);
<specimenCount id="9D12FD37311BFF94FF4EBF0CD17A2515" box="[244,345,1051,1077]" count="11" pageId="5" pageNumber="308" type="male">11 males</specimenCount>
,
<specimenCount id="9D12FD37311BFF94FED9BF0CD1F22515" box="[355,465,1051,1077]" count="9" pageId="5" pageNumber="308" type="female">9 females</specimenCount>
, same locality,
<date id="FFAA107E311BFF94FDC3BF0CD2C92515" box="[633,746,1051,1077]" pageId="5" pageNumber="308" value="1993-05-19">
<collectingDate id="EFEEE996311BFF94FDC3BF0CD2C92515" box="[633,746,1051,1077]" pageId="5" pageNumber="308" value="1993-05-19">19.v.1993</collectingDate>
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, N.
<location id="8ECB6065311BFF94FCAFBF0CD3442515" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FCAFBF0CD3442515" box="[789,871,1051,1077]" country="Japan" county="Nankai" municipality="Atojifukushimo" name="Kuhara" pageId="5" pageNumber="308" stateProvince="Yamaguchi">Kuhara</location>
(
<specimenCount id="9D12FD37311BFF94FCCCBF0CD3EC2515" box="[886,975,1051,1077]" count="6" pageId="5" pageNumber="308" type="male">6 males</specimenCount>
,
<specimenCount id="9D12FD37311BFF94FC63BF0CD46A2515" box="[985,1097,1051,1077]" count="3" pageId="5" pageNumber="308" type="female">3 females</specimenCount>
: KuN);
<specimenCount id="9D12FD37311BFF94FB25BF0CD4CF2515" box="[1183,1260,1051,1077]" count="1" pageId="5" pageNumber="308" type="male">1 male</specimenCount>
,
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, same locality,
<date id="FFAA107E311BFF94FF42BF28D1472579" box="[248,356,1087,1113]" pageId="5" pageNumber="308" value="1994-06-09">
<collectingDate id="EFEEE996311BFF94FF42BF28D1472579" box="[248,356,1087,1113]" pageId="5" pageNumber="308" value="1994-06-09">9.vi.1994</collectingDate>
</date>
,
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.
<materialsCitation id="3B7C3CE3311BFF94FE34BF28D08A255D" collectingDate="1996-05-22" collectingDateMax="1994-06-09" collectingDateMin="1935-05-31" collectionCode="SPMN, T" collectorName="Yamauchi" country="Japan" county="Nankai" location="Nozaki" municipality="Atojifukushimo" pageId="5" pageNumber="308" specimenCount="3" specimenCount-larva="1" specimenCount-male="2" stateProvince="Yamaguchi">
<location id="8ECB6065311BFF94FE34BF28D1FC2579" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FE34BF28D1FC2579" box="[398,479,1087,1113]" country="Japan" name="Nozaki" pageId="5" pageNumber="308">Nozaki</location>
(
<specimenCount id="9D12FD37311BFF94FE4BBF28D2632579" box="[497,576,1087,1113]" count="1" pageId="5" pageNumber="308" type="male">1 male</specimenCount>
:
<collectionCode id="ED05AE7B311BFF94FDF1BF28D2BF2579" box="[587,668,1087,1113]" pageId="5" pageNumber="308">SPMN</collectionCode>
);
<specimenCount id="9D12FD37311BFF94FD0ABF28D2DF2579" box="[688,764,1087,1113]" count="1" pageId="5" pageNumber="308" type="male">1 male</specimenCount>
pupa with larval exuviae,
<specimenCount id="9D12FD37311BFF94FB93BF28D45A2579" box="[1065,1145,1087,1113]" count="1" pageId="5" pageNumber="308" type="larva">1 larva</specimenCount>
, same locality,
<date id="FFAA107E311BFF94FA9DBF28D5BB2579" box="[1319,1432,1087,1113]" pageId="5" pageNumber="308" value="1996-05-22">
<collectingDate id="EFEEE996311BFF94FA9DBF28D5BB2579" box="[1319,1432,1087,1113]" pageId="5" pageNumber="308" value="1996-05-22">22.v.1996</collectingDate>
</date>
,
<collectionCode id="ED05AE7B311BFF94FF2DBF74D08A255D" box="[151,169,1123,1149]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="308">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3311BFF94FF0EBF74D4522581" collectingDate="1969-05-18" collectingDateMax="2004-04-17" collectingDateMin="1969-05-18" collectionCode="SPMN, KPM, T" collectorName="N. Gyotoku &amp; Dobaru &amp; Kokuraminami-ku" country="Japan" county="Hattori" elevation="200" location="Asakura-shi" municipality="Mt. Kosho" pageId="5" pageNumber="308" specimenCount="2" specimenCount-male="1" specimenCount-pupa="1">
<collectingCounty id="62CA4E32311BFF94FF0EBF74D120255D" box="[180,259,1123,1149]" pageId="5" pageNumber="308">Hattori</collectingCounty>
(
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);
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,
<collectingMunicipality id="6BCFACC4311BFF94FE75BF74D269255D" box="[463,586,1123,1149]" pageId="5" pageNumber="308">Mt. Kosho</collectingMunicipality>
,
<location id="8ECB6065311BFF94FDE9BF74D2FE255D" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FDE9BF74D2FE255D" box="[595,733,1123,1149]" country="Japan" county="Hattori" municipality="Mt. Kosho" name="Asakura-shi" pageId="5" pageNumber="308">Asakura-shi</location>
,
<date id="FFAA107E311BFF94FD52BF74D37B255D" box="[744,856,1123,1149]" pageId="5" pageNumber="308" value="1969-05-18">
<collectingDate id="EFEEE996311BFF94FD52BF74D37B255D" box="[744,856,1123,1149]" pageId="5" pageNumber="308" value="1969-05-18">18.v.1969</collectingDate>
</date>
,
<collectorName id="26E15368311BFF94FCD9BF74D3C4255D" box="[867,999,1123,1149]" pageId="5" pageNumber="308">N. Gyotoku</collectorName>
(
<collectionCode id="ED05AE7B311BFF94FC4DBF74D41A255D" box="[1015,1081,1123,1149]" country="Japan" lsid="urn:lsid:biocol.org:col:15118" name="Kanagawa Prefectural Museum of Natural History" pageId="5" pageNumber="308" type="Herbarium">KPM</collectionCode>
);
<specimenCount id="9D12FD37311BFF94FBF1BF74D4BA255D" box="[1099,1177,1123,1149]" count="1" pageId="5" pageNumber="308" type="pupa">1 pupa</specimenCount>
, 2 pupal remnants with larval exuviae,
<collectorName id="26E15368311BFF94FEF2BF90D1BF2581" box="[328,412,1159,1185]" pageId="5" pageNumber="308">Dobaru</collectorName>
(alt.
<quantity id="4CEC9B5B311BFF94FE61BF90D20B2581" box="[475,552,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="308" unit="m" value="200.0">
<elevation id="0039D18D311BFF94FE61BF90D20B2581" box="[475,552,1159,1185]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="308" unit="m" value="200.0">200 m</elevation>
</quantity>
),
<collectorName id="26E15368311BFF94FD84BF90D3332581" box="[574,784,1159,1185]" pageId="5" pageNumber="308">Kokuraminami-ku</collectorName>
,
<location id="8ECB6065311BFF94FCA7BF90D3E82581" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FCA7BF90D3E82581" box="[797,971,1159,1185]" country="Japan" county="Hattori" municipality="Mt. Kosho" name="Kitakyushu-shi" pageId="5" pageNumber="308">Kitakyushu-shi</location>
,
<date id="FFAA107E311BFF94FC63BF90D4712581" box="[985,1106,1159,1185]" pageId="5" pageNumber="308" value="2004-04-17">
<collectingDate id="EFEEE996311BFF94FC63BF90D4712581" box="[985,1106,1159,1185]" pageId="5" pageNumber="308" value="2004-04-17">17.iv.2004</collectingDate>
</date>
,
<collectionCode id="ED05AE7B311BFF94FBE5BF90D4522581" box="[1119,1137,1159,1185]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="308">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3311BFF94FBC4BF90D3AA25E5" collectingDate="2004-04-18" collectionCode="SPMN, T" collectorName="Shimizubakuen" country="Japan" county="Hattori" elevation="450" location="Kusu-machi" municipality="Mori" pageId="5" pageNumber="308" specimenCount="1" specimenCount-male="1" stateProvince="Oita">
<collectingCounty id="62CA4E32311BFF94FBC4BF90D4EE2581" box="[1150,1229,1159,1185]" pageId="5" pageNumber="308">Hattori</collectingCounty>
(
<collectionCode id="ED05AE7B311BFF94FB65BF90D5112581" box="[1247,1330,1159,1185]" pageId="5" pageNumber="308">SPMN</collectionCode>
).
<emphasis id="B960EAAC311BFF94FAFDBF91D5A62580" bold="true" box="[1351,1413,1158,1184]" pageId="5" pageNumber="308">
<collectingRegion id="49D0F85C311BFF94FAFDBF91D5A32580" box="[1351,1408,1158,1184]" country="Japan" name="Oita" pageId="5" pageNumber="308">Oita</collectingRegion>
:
</emphasis>
<specimenCount id="9D12FD37311BFF94FA34BF90D0F325E5" count="1" pageId="5" pageNumber="308" type="male">1 male</specimenCount>
,
<collectorName id="26E15368311BFF94FF66BFBCD1AE25E5" box="[220,397,1195,1221]" pageId="5" pageNumber="308">Shimizubakuen</collectorName>
, alt.
<quantity id="4CEC9B5B311BFF94FE78BFBDD22825E5" box="[450,523,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="308" unit="m" value="450.0">
<elevation id="0039D18D311BFF94FE78BFBDD22825E5" box="[450,523,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="308" unit="m" value="450.0">450 m</elevation>
</quantity>
,
<collectingMunicipality id="6BCFACC4311BFF94FDADBFBCD27125E5" box="[535,594,1195,1221]" pageId="5" pageNumber="308">Mori</collectingMunicipality>
,
<location id="8ECB6065311BFF94FDE7BFBCD2CA25E5" LSID="urn:lsid:plazi:treatment:03BD87A8311CFF94FF2DBACCD2E52699:8ECB6065311BFF94FDE7BFBCD2CA25E5" box="[605,745,1195,1221]" country="Japan" county="Hattori" municipality="Mori" name="Kusu-machi" pageId="5" pageNumber="308" stateProvince="Oita">Kusu-machi</location>
,
<date id="FFAA107E311BFF94FD49BFBCD34F25E5" box="[755,876,1195,1221]" pageId="5" pageNumber="308" value="2004-04-18">
<collectingDate id="EFEEE996311BFF94FD49BFBCD34F25E5" box="[755,876,1195,1221]" pageId="5" pageNumber="308" value="2004-04-18">18.iv.2004</collectingDate>
</date>
,
<collectionCode id="ED05AE7B311BFF94FCCDBFBCD3AA25E5" box="[887,905,1195,1221]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="308">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B7C3CE3311BFF94FC2EBFBCD46E25E5" box="[916,1101,1195,1221]" collectionCode="SPMN" collectorName="Hattori" country="Japan" pageId="5" pageNumber="308" specimenCount="1">
<collectorName id="26E15368311BFF94FC2EBFBCD3C025E5" box="[916,995,1195,1221]" pageId="5" pageNumber="308">Hattori</collectorName>
(
<collectionCode id="ED05AE7B311BFF94FC48BFBCD46625E5" box="[1010,1093,1195,1221]" pageId="5" pageNumber="308">SPMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E6535311BFF94FF7DBFD9D30A25C9" box="[199,809,1230,1257]" pageId="5" pageNumber="308" type="distribution">
<paragraph id="8BAB36BE311BFF94FF7DBFD9D30A25C9" blockId="5.[151,1437,151,1977]" box="[199,809,1230,1257]" pageId="5" pageNumber="308">
<emphasis id="B960EAAC311BFF94FF7DBFD9D14225C8" bold="true" box="[199,353,1230,1256]" pageId="5" pageNumber="308">Distribution.</emphasis>
Westernmost Honshu, northern Kyushu.
</paragraph>
</subSubSection>
<subSubSection id="C30E6535311BFF94FF7DBFE5D2912475" pageId="5" pageNumber="308" type="biology_ecology">
<paragraph id="8BAB36BE311BFF94FF7DBFE5D0FB2411" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="308">
<emphasis id="B960EAAC311BFF94FF7DBFE5D104242C" bold="true" box="[199,295,1266,1292]" pageId="5" pageNumber="308">Biology.</emphasis>
Larval and pupal specimens of this species were collected from small streams in hilly and mountainous areas.
</paragraph>
<paragraph id="8BAB36BE311BFF94FF7DBE2CD2912475" blockId="5.[151,1437,151,1977]" box="[199,690,1339,1365]" pageId="5" pageNumber="308">
<emphasis id="B960EAAC311BFF94FF7DBE2CD1A02475" bold="true" box="[199,387,1339,1365]" pageId="5" pageNumber="308">Japanese name.</emphasis>
Kimadara-shima-tobikera.
</paragraph>
</subSubSection>
<subSubSection id="C30E6535311BFF94FF7DBE48D2E52699" pageId="5" pageNumber="308" type="discussion">
<paragraph id="8BAB36BE311BFF94FF7DBE48D0F82625" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="308">
<emphasis id="B960EAAC311BFF94FF7DBE48D1182459" bold="true" box="[199,315,1375,1401]" pageId="5" pageNumber="308">Remarks.</emphasis>
<bibRefCitation id="EF854B4F311BFF94FEF8BE48D1F92459" author="Banks, N." box="[322,474,1375,1401]" pageId="5" pageNumber="308" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks (1906)</bibRefCitation>
described
<taxonomicName id="4C144D3D311BFF94FDEEBE48D2F52458" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[596,726,1375,1401]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311BFF94FDEEBE48D2F52458" box="[596,726,1375,1401]" italics="true" pageId="5" pageNumber="308">A. japonica</emphasis>
</taxonomicName>
from
<collectingRegion id="49D0F85C311BFF94FCA1BE48D3A32459" box="[795,896,1375,1401]" country="Japan" name="Fukuoka" pageId="5" pageNumber="308">Fukuoka</collectingRegion>
, Kyushu and
<collectingRegion id="49D0F85C311BFF94FBA3BE48D46C2459" box="[1049,1103,1375,1401]" country="Japan" name="Gifu" pageId="5" pageNumber="308">Gifu</collectingRegion>
, central Honshu as “Hikosan, Buzen [now
<collectingRegion id="49D0F85C311BFF94FE9FBE94D1AE24BD" box="[293,397,1411,1437]" country="Japan" name="Fukuoka" pageId="5" pageNumber="308">Fukuoka</collectingRegion>
],
<collectingCountry id="F303762E311BFF94FE27BE94D1C324BD" box="[413,480,1411,1437]" name="Japan" pageId="5" pageNumber="308">Japan</collectingCountry>
, 28 April; also from
<collectingRegion id="49D0F85C311BFF94FD79BE94D2D824BD" box="[707,763,1411,1437]" country="Japan" name="Gifu" pageId="5" pageNumber="308">Gifu</collectingRegion>
”, but no
<typeStatus id="54AF881C311BFF94FCE6BE94D39D24BD" box="[860,958,1411,1437]" pageId="5" pageNumber="308" type="holotype">holotype</typeStatus>
was designated.
<bibRefCitation id="EF854B4F311BFF94FBC1BE94D53E24BD" author="Nozaki, T." box="[1147,1309,1411,1437]" pageId="5" pageNumber="308" pagination="245 - 258" refId="ref18077" refString="Nozaki, T. (2021) The genus Diplectrona Westwood (Trichoptera: Hydropsychidae) in the Japanese main islands. Zootaxa, 5082 (3), 245 - 258. https: // doi. org / 10.11646 / zootaxa. 5082.3.3" type="journal article" year="2021">Nozaki (2021)</bibRefCitation>
transferred this species to the genus
<taxonomicName id="4C144D3D311BFF94FE09BEB0D26424E1" authorityName="Ross" authorityYear="1938" box="[435,583,1447,1473]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAAC311BFF94FE09BEB0D26424E1" box="[435,583,1447,1473]" italics="true" pageId="5" pageNumber="308">Homoplectra</emphasis>
</taxonomicName>
based on examination of photographs of a
<typeStatus id="54AF881C311BFF94FB8EBEB0D4A824E1" box="[1076,1163,1447,1473]" pageId="5" pageNumber="308" type="syntype">syntype</typeStatus>
male (MCZ:Ent:11833) collected from
<collectingRegion id="49D0F85C311BFF94FEFEBEDCD18524C5" box="[324,422,1483,1509]" country="Japan" name="Fukuoka" pageId="5" pageNumber="308">Fukuoka</collectingRegion>
and deposited in the Banks collection of MCZ. In this study, I examined photographs of another specimen also deposited in the Banks collection as a
<typeStatus id="54AF881C311BFF94FCEDBEF8D38D2729" box="[855,942,1519,1545]" pageId="5" pageNumber="308" type="syntype">syntype</typeStatus>
(MCZ:Ent:648191) kindly provided Dr. C. W. Farnum (MCZ). Since this specimen had a handwritten label “
<collectingRegion id="49D0F85C311BFF94FC32BD04D39D270D" box="[904,958,1555,1581]" country="Japan" name="Gifu" pageId="5" pageNumber="308">Gifu</collectingRegion>
<collectingCountry id="F303762E311BFF94FC7CBD04D42B270D" box="[966,1032,1555,1581]" name="Japan" pageId="5" pageNumber="308">Japan</collectingCountry>
” (https://mczbase.mcz.harvard.edu/ guid/MCZ:Ent:648191), the record from
<collectingRegion id="49D0F85C311BFF94FDD9BD20D2B52771" box="[611,662,1591,1617]" country="Japan" name="Gifu" pageId="5" pageNumber="308">Gifu</collectingRegion>
in the original description by
<bibRefCitation id="EF854B4F311BFF94FC5DBD20D45D2771" author="Banks, N." box="[999,1150,1591,1617]" pageId="5" pageNumber="308" pagination="106 - 113" refId="ref16877" refString="Banks, N. (1906) New Trichoptera from Japan. Proceedings of the Entomological Society of Washington, 7, 106 - 113." type="journal article" year="1906">Banks (1906)</bibRefCitation>
must have been based on it. Although I could not examine its genital morphology because of mold covered the abdomen, the length of the scape of left antenna (right antenna missing) was apparently longer than its width. Since the
<typeStatus id="54AF881C311BFF94FB26BD68D4D027B9" box="[1180,1267,1663,1689]" pageId="5" pageNumber="308" type="syntype">syntype</typeStatus>
male collected from
<collectingRegion id="49D0F85C311BFF94FF6EBDB4D115279D" box="[212,310,1699,1725]" country="Japan" name="Fukuoka" pageId="5" pageNumber="308">Fukuoka</collectingRegion>
has a short scape (
<bibRefCitation id="EF854B4F311BFF94FDB9BDB4D2B4279D" author="Nozaki, T." box="[515,663,1699,1725]" pageId="5" pageNumber="308" pagination="245 - 258" refId="ref18077" refString="Nozaki, T. (2021) The genus Diplectrona Westwood (Trichoptera: Hydropsychidae) in the Japanese main islands. Zootaxa, 5082 (3), 245 - 258. https: // doi. org / 10.11646 / zootaxa. 5082.3.3" type="journal article" year="2021">Nozaki 2021</bibRefCitation>
, fig. 5A), these
<specimenCount id="9D12FD37311BFF94FCFFBDB4D3C9279D" box="[837,1002,1699,1725]" count="2" pageId="5" pageNumber="308" type="generic">two specimens</specimenCount>
must belong to different species. Thus, I designate here the male collected from
<collectingRegion id="49D0F85C311BFF94FDDDBDD0D2EA27C1" box="[615,713,1735,1761]" country="Japan" name="Fukuoka" pageId="5" pageNumber="308">Fukuoka</collectingRegion>
(MCZ:Ent:11833) as the
<typeStatus id="54AF881C311BFF94FC54BDD0D47627C1" box="[1006,1109,1735,1761]" pageId="5" pageNumber="308" type="lectotype">lectotype</typeStatus>
to fix the status of the taxon name.
</paragraph>
<paragraph id="8BAB36BE311BFF94FF7DBC18D2E52699" blockId="5.[151,1437,151,1977]" pageId="5" pageNumber="308">
As a result of the
<typeStatus id="54AF881C311BFF94FE12BC18D22C2609" box="[424,527,1807,1833]" pageId="5" pageNumber="308" type="lectotype">lectotype</typeStatus>
designation of
<taxonomicName id="4C144D3D311BFF94FD76BC18D3712608" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[716,850,1807,1833]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311BFF94FD76BC18D3712608" box="[716,850,1807,1833]" italics="true" pageId="5" pageNumber="308">A. japonica</emphasis>
</taxonomicName>
, another specimen (MCZ:Ent:648191) becomes a
<typeStatus id="54AF881C311BFF94FF2DBC24D111266D" box="[151,306,1843,1869]" pageId="5" pageNumber="308" type="paralectotype">paralectotype</typeStatus>
. Although I could not identify this specimen at the species level,
<taxonomicName id="4C144D3D311BFF94FBEFBC24D501266D" baseAuthorityName="Ulmer" baseAuthorityYear="1907" box="[1109,1314,1843,1869]" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="albomarginata">
<emphasis id="B960EAAC311BFF94FBEFBC24D501266D" box="[1109,1314,1843,1869]" italics="true" pageId="5" pageNumber="308">H. albomarginata</emphasis>
</taxonomicName>
is widely distributed in
<collectingRegion id="49D0F85C311BFF94FE8EBC40D1C12651" box="[308,482,1879,1905]" country="Japan" name="Gifu" pageId="5" pageNumber="308">Gifu Prefecture</collectingRegion>
and the scape of each antenna (
<figureCitation id="132F2A3B311BFF94FCF0BC40D3F72651" box="[842,980,1879,1905]" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="5" pageNumber="308">Figs 2B, 2C</figureCitation>
) is longer than that of
<taxonomicName id="4C144D3D311BFF94FB6CBC40D5782650" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1238,1371,1879,1904]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311BFF94FB6CBC40D5782650" box="[1238,1371,1879,1904]" italics="true" pageId="5" pageNumber="308">H. japonica</emphasis>
</taxonomicName>
(
<figureCitation id="132F2A3B311BFF94FAD0BC40D0CB26B5" captionStart="FIGURES 12" captionStartId="3.[151,264,1626,1650]" captionTargetBox="[157,1429,351,1596]" captionTargetId="figure-102@3.[151,1435,345,1602]" captionTargetPageId="3" captionText="FIGURES 12. Adults of Homoplectra spp. 1A1E, Homoplectra japonica (Banks 1906): 1A, habitus, dorsal (Mt. Kosho, Fukuoka); 1B, male right wings, dorsal (Mt. Hiko, Fukuoka); 1C, male, head and pro- and mesonota, dorsal (same); 1D, male sternum V, left lateral (same); 1E, female, head, dorsal (same). 2A2C, Homoplectra albomarginata (Ulmer 1907): 2A12A3, right wings, dorsal (Ibigawa-cho, Gifu): 2A1, male; 2A2, female; 2A3, female with indistinct markings; 2B, male, head and pro- and mesonota, dorsal (same); 2C, female, head, dorsal (same). Abbreviations: I, II, III, IV, V = apical forks I, II, III, IV, V; DC = discoidal cell; MC = median cell. Scales: 5 mm." pageId="5" pageNumber="308">Figs 1C, 1E</figureCitation>
).
<bibRefCitation id="EF854B4F311BFF94FF44BC6CD1BB26B5" author="Ulmer, G." box="[254,408,1915,1941]" pageId="5" pageNumber="308" pagination="1 - 102" refId="ref18285" refString="Ulmer, G. (1907) Trichopteren. Part. 1. Collection of Zoologiques du Baron Edm. de Selys Longchamps, 6 (1), 1 - 102." type="journal article" year="1907">Ulmer (1907)</bibRefCitation>
recorded
<taxonomicName id="4C144D3D311BFF94FDB3BC6CD2AD26B4" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[521,654,1915,1940]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311BFF94FDB3BC6CD2AD26B4" box="[521,654,1915,1940]" italics="true" pageId="5" pageNumber="308">H. japonica</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C144D3D311BFF94FD05BC6CD36726B4" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[703,836,1915,1940]" class="Insecta" family="Hydropsychidae" genus="Diplectrona" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B960EAAC311BFF94FD05BC6CD36726B4" box="[703,836,1915,1940]" italics="true" pageId="5" pageNumber="308">D. japonica</emphasis>
</taxonomicName>
), also from
<collectingRegion id="49D0F85C311BFF94FC75BC6CD42526B5" box="[975,1030,1915,1941]" country="Japan" name="Gifu" pageId="5" pageNumber="308">Gifu</collectingRegion>
; but it was a misidentification of
<taxonomicName id="4C144D3D311BFF94FA3BBC6CD11F2699" baseAuthorityName="Ulmer" baseAuthorityYear="1907" class="Insecta" family="Hydropsychidae" genus="Homoplectra" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="albomarginata">
<emphasis id="B960EAAC311BFF94FA3BBC6CD11F2699" italics="true" pageId="5" pageNumber="308">H. albomarginata</emphasis>
</taxonomicName>
(see the Remarks for that species).
</paragraph>
</subSubSection>
</treatment>
</document>

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<bibRefCitation id="EFD44089FFA6FC42FED4FEB95C3BFE93" author="Pons, L. N. &amp; Calcinai, B. &amp; Gates, R. D." box="[353,572,255,280]" pageId="18" pageNumber="261" refId="ref19800" refString="Pons, L. N., Calcinai, B. &amp; Gates, R. D. (2017) Who's there? - First morphological and DNA barcoding catalogue of the shallow Hawaiian sponge fauna. PLoS ONE, 12, e 0189357. https: // doi. org / 10.1371 / journal. pone. 0189357" type="journal volume" year="2017">
Pons
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, 2017: 46
</bibRefCitation>
, Fig 22;
<bibRefCitation id="EFD44089FFA6FC42FD2EFEB95D9AFE9C" author="Vicente, J. &amp; Osberg, A. &amp; Marty, M. J. &amp; Rice, K. &amp; Toonen, R. J." box="[667,925,256,280]" pageId="18" pageNumber="261" pagination="109 - 122" refId="ref21450" refString="Vicente, J., Osberg, A., Marty, M. J., Rice, K. &amp; Toonen, R. J. (2020) Influence of sponge palatability on the feeding preferences of the endemic Hawaiian tiger cowrie for indigenous and introduced sponges. Marine Ecology Progress Series, 647, 109 - 122." type="journal article" year="2020">
Vicente
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, 2020: 111
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; Vicente et al; 2022b:
<figureCitation id="137E21FDFFA6FC42FB64FEB95B0CFE9C" box="[1233,1291,256,280]" captionStart="FIGURE 4" captionStartId="9.[151,250,1876,1900]" captionTargetBox="[162,1425,428,1841]" captionTargetId="figure-146@9.[151,1436,417,1852]" captionTargetPageId="9" captionText="FIGURE 4. In situ growth variations of Haliclona (Reniera) kahoe sp. nov. a, holotype BPBM C1539; b, paratype BPBM C1538; c, paratype BPBM C1551; d, paratype BPBM C1537; e, paratype BPBM C1540; f, paratype BPBM C1570; g, paratype BPBM C1554; h, BPBM C1553; i, paratype BPBM C1552. Scale bars: a-f, 1 cm; g, 0.5 cm; h-i, 1 cm." pageId="18" pageNumber="261">Fig 4</figureCitation>
,
<tableCitation id="C6C708C3FFA6FC42FAA2FEB95B8DFE93" box="[1303,1418,255,280]" captionStart="TABLE 4" captionStartId="17.[152,244,151,175]" captionText="TABLE 4. Spicule measurements of oxeas and sigmas for specimens of Haliclona (Soestella) caerulea. Measurements are expressed as minimummean (±1 standard deviation)maximum. N=50 for oxeas and N=10 for sigmas. Measurements for the holotype were provided from Hechtel (1965)" pageId="18" pageNumber="261">Table S4-S</tableCitation>
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;
<bibRefCitation id="EFD44089FFA6FC42FF72FE9D5FADFEB8" author="Vicente, J. &amp; Timmers, M. A. &amp; Webb, M. K. &amp; Bahr, K. D. &amp; Jury, C. P. &amp; Toonen, R. J." box="[199,426,290,316]" pageId="18" pageNumber="261" refId="ref21574" refString="Vicente, J., Timmers, M. A., Webb, M. K., Bahr, K. D., Jury, C. P. &amp; Toonen, R. J. (2022 a) Ecological succession of the sponge cryptofauna in Hawaiian reefs add new insights to detritus production by pioneering species. Scientific Reports, 12, 15093. https: // doi. org / 10.1038 / s 41598 - 022 - 18856 - 8" type="journal volume" year="2022">
Vicente
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: Suppl.
<tableCitation id="C6C708C3FFA6FC42FDB6FE9B5C7AFEB8" box="[515,637,290,316]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="7.[152,244,471,495]" captionStartId-1="11.[151,243,151,175]" captionText-0="TABLE 1. Spicule measurements of oxeas and sigmas for Haliclona (Gellius) loe sp. nov.. Measurements are expressed as minimummean (±1 standard deviation)maximum. N=50 for oxeas and N=10 for sigmas." captionText-1="TABLE 2. Spicule measurements of oxeas for Haliclona (Reniera) kahoe holotype (h) and paratypes (p). Measurements are expressed as minimummean (±1 standard deviation)maximum. N=50." pageId="18" pageNumber="261">Table S1-2</tableCitation>
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<mods:namePart id="9E791108D349C81552E3ABAD4E67523F">García-Prieto, Luis</mods:namePart>
<mods:affiliation id="E068C255431F04AF0B2DE8361105377A">Instituto de Biología, Universidad Nacional Autónoma de México, Ciudad Universitaria, Ciudad de México C. P. 04510, México.</mods:affiliation>
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<mods:namePart id="23EE16FF60A37B2FED3030562F30BC0C">Canché-Pool, Elsy B.</mods:namePart>
<mods:affiliation id="55BB4BC0069FA12DCD189EF9515337A9">Centro de Investigaciones Regionales Dr. Hideyo Noguchi, Universidad Autónoma de Yucatán, Avenida Itzáes, Mérida 97000, Yucatán, México.</mods:affiliation>
</mods:name>
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<mods:namePart id="424FAC673ABBCC3B06C6FA15B4A60397">Robles, María Del Rosario</mods:namePart>
<mods:affiliation id="606FF9649AE148C41247623598DF1378">Centro de Estudios Parasitológicos y de Vectores (CEPAVE), Universidad Nacional de La Plata, La Plata, Buenos Aires, Argentina.</mods:affiliation>
</mods:name>
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<mods:namePart id="107960DB4D3290DB522C2E2999769684">Ávila-López, Mariana B.</mods:namePart>
<mods:affiliation id="72D94E5D8D1265005EDC2DD63D1EAC65">Comisión Intersecretarial de Bioseguridad de los Organismos Genéticamente Modificados (CIBIOGEM-CONAHCYT), Ciudad de México, México.</mods:affiliation>
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<mods:namePart id="84142D7A48EBA39FCC101713050CAC50">Chan-Casanova, Anyela Jackelin</mods:namePart>
<mods:affiliation id="DC351FFF1BD5F8E8888FF71B4E8830A8">Facultad de Medicina Veterinaria y Zootecnia, Universidad Autónoma de Yucatán, Mérida Yucatán, México.</mods:affiliation>
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<mods:namePart id="EB4F3F5549802C451AABA5614541B19C">Panti-May, Jesús Alonso</mods:namePart>
<mods:affiliation id="8FD3678A415A6321E2AC2E8EA87FEE33">Centro de Investigaciones Regionales Dr. Hideyo Noguchi, Universidad Autónoma de Yucatán, Avenida Itzáes, Mérida 97000, Yucatán, México.</mods:affiliation>
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<mods:title id="0F2ADC1C62DDB296A172D79FD27F207F">Zootaxa</mods:title>
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<mods:date id="39A9924569E777506FDD5789F68DBD69">2025</mods:date>
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<mods:number id="5301EC23610578A60057E9A254D026E7">5566</mods:number>
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<mods:url id="A243F40A8D554DB1DC5C87101271435D">https://doi.org/10.11646/zootaxa.5566.2.5</mods:url>
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<mods:identifier id="A3614012A8D7D472FC84BDFCEBBB2506" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C3664F578AB1597ED665495725A69B35" type="ZooBank">9A17BF2C-EB04-4215-9783-46E0C45BB126</mods:identifier>
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<treatment id="03ED5E372C21FF9BA8FEACB2938AFD78" LSID="urn:lsid:plazi:treatment:03ED5E372C21FF9BA8FEACB2938AFD78" httpUri="http://treatment.plazi.org/id/03ED5E372C21FF9BA8FEACB2938AFD78" lastPageId="11" lastPageNumber="340" pageId="8" pageNumber="337">
<subSubSection id="C35EBCAA2C21FF98A8FEACB2902FFB60" pageId="8" pageNumber="337" type="nomenclature">
<paragraph id="8BFBEF212C21FF98A8FEACB29003FB29" blockId="8.[151,549,1158,1185]" box="[151,549,1158,1185]" pageId="8" pageNumber="337">
<heading id="D0B3584D2C21FF98A8FEACB29003FB29" bold="true" box="[151,549,1158,1185]" fontSize="11" level="1" pageId="8" pageNumber="337" reason="1">
<taxonomicName id="4C4494A22C21FF98A8FEACB29003FB29" authority="Chitwood" authorityName="Chitwood" box="[151,549,1158,1185]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="8" pageNumber="337" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C21FF98A8FEACB29003FB29" bold="true" box="[151,549,1158,1185]" pageId="8" pageNumber="337">
<emphasis id="B93033332C21FF98A8FEACB29382FB28" bold="true" box="[151,420,1158,1185]" italics="true" pageId="8" pageNumber="337">Bidigiticauda vivipara</emphasis>
Chitwood
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFBEF212C21FF98A8FEACFA902FFB60" blockId="8.[151,1437,1230,2013]" box="[151,521,1230,1257]" pageId="8" pageNumber="337">
<emphasis id="B93033332C21FF98A8FEACFA937DFB61" bold="true" box="[151,347,1230,1256]" pageId="8" pageNumber="337">Site of infection:</emphasis>
Small intestine
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</subSubSection>
<subSubSection id="C35EBCAA2C21FF98A8AEACC7910BFA14" pageId="8" pageNumber="337" type="biology_ecology">
<paragraph id="8BFBEF212C21FF98A8AEACC793D0FA85" blockId="8.[151,1437,1230,2013]" box="[199,502,1267,1293]" pageId="8" pageNumber="337">
<emphasis id="B93033332C21FF98A8AEACC7932EFA84" bold="true" box="[199,264,1267,1293]" pageId="8" pageNumber="337">Host:</emphasis>
<taxonomicName id="4C4494A22C21FF98A979ACC793D0FA85" authorityName="Merriam" authorityYear="1898" box="[272,502,1267,1293]" class="Mammalia" family="Phyllostomidae" genus="Glossophaga" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="337" phylum="Chordata" rank="species" species="mutica">
<emphasis id="B93033332C21FF98A979ACC793D0FA85" box="[272,502,1267,1293]" italics="true" pageId="8" pageNumber="337">Glossophaga mutica</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BFBEF212C21FF98A8AEAD22974DFAB8" blockId="8.[151,1437,1230,2013]" box="[199,1387,1302,1329]" pageId="8" pageNumber="337">
<emphasis id="B93033332C21FF98A8AEAD229315FAB9" bold="true" box="[199,307,1302,1328]" pageId="8" pageNumber="337">Locality:</emphasis>
<materialsCitation id="3B2CE57C2C21FF98A950AD23974DFAB8" box="[313,1387,1302,1329]" collectionCode="MASL" latitude="21.661444" location="Islas Marias" longLatPrecision="1" longitude="-106.616196" pageId="8" pageNumber="337" specimenCount="1" stateProvince="Nayarit">
Campamento Bugambilias (
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,
<geoCoordinate id="EE7089E62C21FF98AB4EAD2291C6FAB8" box="[807,992,1302,1329]" degrees="106" direction="west" minutes="36" orientation="longitude" pageId="8" pageNumber="337" precision="1" seconds="58.3" value="-106.616196">106°3658.3” W</geoCoordinate>
, 164
<collectionCode id="ED5577E42C21FF98AC75AD239648FAB8" box="[1052,1134,1303,1329]" pageId="8" pageNumber="337">MASL</collectionCode>
),
<location id="8E9BB9FA2C21FF98ACE8AD23972BFAB8" LSID="urn:lsid:plazi:treatment:03ED5E372C21FF9BA8FEACB2938AFD78:8E9BB9FA2C21FF98ACE8AD23972BFAB8" box="[1153,1293,1303,1329]" latitude="21.661444" longLatPrecision="1" longitude="-106.616196" name="Islas Marias" pageId="8" pageNumber="337" stateProvince="Nayarit">Islas Marías</location>
,
<collectingRegion id="498021C32C21FF98AD7EAD23974DFAB8" box="[1303,1387,1303,1329]" country="Mexico" name="Nayarit" pageId="8" pageNumber="337">Nayarit</collectingRegion>
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</paragraph>
<paragraph id="8BFBEF212C21FF98A8AEAD0F90C4FADC" blockId="8.[151,1437,1230,2013]" box="[199,738,1338,1365]" pageId="8" pageNumber="337">
<emphasis id="B93033332C21FF98A8AEAD0F93D3FADD" bold="true" box="[199,501,1338,1365]" pageId="8" pageNumber="337">Prevalence and intensity:</emphasis>
33.3% (1/3) and 131
</paragraph>
<paragraph id="8BFBEF212C21FF98A8AEAD6A9041FAF0" blockId="8.[151,1437,1230,2013]" box="[199,615,1374,1401]" pageId="8" pageNumber="337">
<emphasis id="B93033332C21FF98A8AEAD6A93E1FAF1" bold="true" box="[199,455,1374,1400]" pageId="8" pageNumber="337">Specimens deposited:</emphasis>
CNHE 12131
</paragraph>
<paragraph id="8BFBEF212C21FF98A8AEADB6910BFA14" blockId="8.[151,1437,1230,2013]" box="[199,813,1410,1437]" pageId="8" pageNumber="337">
<emphasis id="B93033332C21FF98A8AEADB6900EFA14" bold="true" box="[199,552,1410,1437]" pageId="8" pageNumber="337">GenBank accession numbers:</emphasis>
PQ476176, PQ476177
</paragraph>
</subSubSection>
<subSubSection id="C35EBCAA2C21FF9AA8AEAD9290EEF90A" lastPageId="10" lastPageNumber="339" pageId="8" pageNumber="337" type="description">
<paragraph id="8BFBEF212C21FF98A8AEAD9290EDF9A4" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="337">
<emphasis id="B93033332C21FF98A8AEAD929351FA49" bold="true" box="[199,375,1446,1472]" pageId="8" pageNumber="337">Redescription:</emphasis>
Based on
<specimenCount id="9D4224A82C21FF98A99AAD93907DFA48" box="[499,603,1447,1473]" count="11" pageId="8" pageNumber="337" type="male">11 males</specimenCount>
,
<specimenCount id="9D4224A82C21FF98AA00AD9390C1FA48" box="[617,743,1447,1473]" count="11" pageId="8" pageNumber="337" type="female">11 females</specimenCount>
, and six third-stage larvae. Nematodes with cephalic vesicle (
<figureCitation id="137FF3A42C21FF98A8F6ADFF9306FA6C" box="[159,288,1483,1509]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figures 4A</figureCitation>
,
<figureCitation id="137FF3A42C21FF98A945ADFE9377FA6C" box="[300,337,1482,1509]" captionStart="FIGURE 5" captionStartId="10.[151,250,1211,1235]" captionTargetBox="[165,1421,181,1187]" captionTargetId="figure-25@10.[151,1435,181,1187]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electron micrographs of Bidigiticauda vivipara. A) Female anterior extremity, lateral view. B) Female head, apical view. C) Male posterior extremity, lateral view. D) Male caudal bursa, ventral view. E) Magnified image of the dorsal lip of the genital cone.F) Female posterior extremity, dorsal view.Abbreviations:dl, dorsolateral papillae; vl, ventrolateral papillae; a, amphids; p7, papillae 7. Scale bars: A, F, 50 = μm; B, D = 40 μm; C = 100 μm; E = 8 μm" pageId="8" pageNumber="337">5A</figureCitation>
), usually wider than long (
<figureCitation id="137FF3A42C21FF98AA14ADFF90D3FA6C" box="[637,757,1483,1509]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figure 4A</figureCitation>
). Presence of four double cephalic papillae, two dorsolateral and two ventrolateral, as well as two amphids (
<figureCitation id="137FF3A42C21FF98AACDADDB913CF980" box="[676,794,1518,1545]" captionStart="FIGURE 5" captionStartId="10.[151,250,1211,1235]" captionTargetBox="[165,1421,181,1187]" captionTargetId="figure-25@10.[151,1435,181,1187]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electron micrographs of Bidigiticauda vivipara. A) Female anterior extremity, lateral view. B) Female head, apical view. C) Male posterior extremity, lateral view. D) Male caudal bursa, ventral view. E) Magnified image of the dorsal lip of the genital cone.F) Female posterior extremity, dorsal view.Abbreviations:dl, dorsolateral papillae; vl, ventrolateral papillae; a, amphids; p7, papillae 7. Scale bars: A, F, 50 = μm; B, D = 40 μm; C = 100 μm; E = 8 μm" pageId="8" pageNumber="337">Figure 5B</figureCitation>
). Esophageal tooth present (
<figureCitation id="137FF3A42C21FF98AC3CADDB96EBF980" box="[1109,1229,1519,1545]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figure 4A</figureCitation>
). Male and female lack longitudinal ridges (
<figureCitation id="137FF3A42C21FF98A9C7AE279069F9A4" box="[430,591,1555,1581]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figures 4B, C</figureCitation>
,
<figureCitation id="137FF3A42C21FF98AA33AE269098F9A4" box="[602,702,1554,1581]" captionStart="FIGURE 5" captionStartId="10.[151,250,1211,1235]" captionTargetBox="[165,1421,181,1187]" captionTargetId="figure-25@10.[151,1435,181,1187]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electron micrographs of Bidigiticauda vivipara. A) Female anterior extremity, lateral view. B) Female head, apical view. C) Male posterior extremity, lateral view. D) Male caudal bursa, ventral view. E) Magnified image of the dorsal lip of the genital cone.F) Female posterior extremity, dorsal view.Abbreviations:dl, dorsolateral papillae; vl, ventrolateral papillae; a, amphids; p7, papillae 7. Scale bars: A, F, 50 = μm; B, D = 40 μm; C = 100 μm; E = 8 μm" pageId="8" pageNumber="337">5A, C, F</figureCitation>
).
</paragraph>
<paragraph id="8BFBEF212C21FF98A8AEAE03974FF8F8" blockId="8.[151,1437,1230,2013]" pageId="8" pageNumber="337">
Males: Body length 3746 (34804250) and width at midbody 124 (100140). Cephalic vesicle 103 (85118) long and 117 (95135) wide. Nerve ring, excretory pore and deirids situated at 199 (175220), 306 (235368), and 301 (232368) from anterior end, respectively. Esophagus 319 (280370) long. Caudal bursa subsymmetrical (
<figureCitation id="137FF3A42C21FF98A8F6AE979304F934" box="[159,290,1699,1725]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figures 4D</figureCitation>
,
<figureCitation id="137FF3A42C21FF98A947AE969372F934" box="[302,340,1698,1725]" captionStart="FIGURE 5" captionStartId="10.[151,250,1211,1235]" captionTargetBox="[165,1421,181,1187]" captionTargetId="figure-25@10.[151,1435,181,1187]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electron micrographs of Bidigiticauda vivipara. A) Female anterior extremity, lateral view. B) Female head, apical view. C) Male posterior extremity, lateral view. D) Male caudal bursa, ventral view. E) Magnified image of the dorsal lip of the genital cone.F) Female posterior extremity, dorsal view.Abbreviations:dl, dorsolateral papillae; vl, ventrolateral papillae; a, amphids; p7, papillae 7. Scale bars: A, F, 50 = μm; B, D = 40 μm; C = 100 μm; E = 8 μm" pageId="8" pageNumber="337">5D</figureCitation>
). Rays 2, 3, and 4 arising independently, ray 4 slightly separated from ray 3. Rays 5 and 6 long, originating from common trunk (
<figureCitation id="137FF3A42C21FF98AA6EAEF39059F968" box="[519,639,1735,1761]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figure 4D</figureCitation>
). Dorsal ray long, divided at its distal third into 2 branches (
<figureCitation id="137FF3A42C21FF98AD71AEF397B6F968" box="[1304,1424,1735,1761]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figure 4D</figureCitation>
). In
<specimenCount id="9D4224A82C21FF98A8DFAEDF9379F88C" box="[182,351,1771,1797]" count="2" pageId="8" pageNumber="337" type="generic">two specimens</specimenCount>
, bursal rays presented variations: right rays 4 and 5 fused (
<figureCitation id="137FF3A42C21FF98AB9DAEDF964FF88C" box="[1012,1129,1771,1797]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figure 4E</figureCitation>
), left ray 2 and 3 fused and dorsal rays right branch long (
<figureCitation id="137FF3A42C21FF98A987AF3B9047F8A0" box="[494,609,1807,1833]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figure 4F</figureCitation>
). Spicules subequal, alate, 182 (160200) long with sharp tips enclosed in a spatulate membrane (
<figureCitation id="137FF3A42C21FF98A9FFAF07902BF8C4" box="[406,525,1843,1869]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figure 4H</figureCitation>
). Ratio spicule length/body length: 4.8% (45.3%). Gubernaculum absent. Genital cone with dorsal lip more developed than ventral lip (
<figureCitation id="137FF3A42C21FF98AA87AF639140F8F8" box="[750,870,1878,1905]" captionStart="FIGURE 5" captionStartId="10.[151,250,1211,1235]" captionTargetBox="[165,1421,181,1187]" captionTargetId="figure-25@10.[151,1435,181,1187]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electron micrographs of Bidigiticauda vivipara. A) Female anterior extremity, lateral view. B) Female head, apical view. C) Male posterior extremity, lateral view. D) Male caudal bursa, ventral view. E) Magnified image of the dorsal lip of the genital cone.F) Female posterior extremity, dorsal view.Abbreviations:dl, dorsolateral papillae; vl, ventrolateral papillae; a, amphids; p7, papillae 7. Scale bars: A, F, 50 = μm; B, D = 40 μm; C = 100 μm; E = 8 μm" pageId="8" pageNumber="337">Figure 5D</figureCitation>
); papillae 7 situated on dorsal lip (
<figureCitation id="137FF3A42C21FF98AC81AF63977AF8F8" box="[1256,1372,1878,1905]" captionStart="FIGURE 5" captionStartId="10.[151,250,1211,1235]" captionTargetBox="[165,1421,181,1187]" captionTargetId="figure-25@10.[151,1435,181,1187]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electron micrographs of Bidigiticauda vivipara. A) Female anterior extremity, lateral view. B) Female head, apical view. C) Male posterior extremity, lateral view. D) Male caudal bursa, ventral view. E) Magnified image of the dorsal lip of the genital cone.F) Female posterior extremity, dorsal view.Abbreviations:dl, dorsolateral papillae; vl, ventrolateral papillae; a, amphids; p7, papillae 7. Scale bars: A, F, 50 = μm; B, D = 40 μm; C = 100 μm; E = 8 μm" pageId="8" pageNumber="337">Figure 5E</figureCitation>
).
</paragraph>
<paragraph id="8BFBEF212C21FF9AA8AEAF4F91F3FA7D" blockId="8.[151,1437,1230,2013]" lastBlockId="10.[151,1437,1425,2028]" lastPageId="10" lastPageNumber="339" pageId="8" pageNumber="337">
Females: Body length 6330 (54807670) and width at midbody 186 (170200). Cephalic vesicle 111 (90130) long and 145 (115170) wide. Nerve ring, excretory pore, and deirids situated at 201 (140245), 291 (228365), and 290 (220380) from anterior end, respectively. Esophagus 315 (240360) long (
<figureCitation id="137FF3A42C21FF98AC7DAFF796AAF854" box="[1044,1164,1987,2013]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="8" pageNumber="337">Figure 4A</figureCitation>
). Viviparous. Didelphic. Vulva situated at 3093 (25003750) from caudal extremity (
<figureCitation id="137FF3A42C23FF9AAB2EADA59193FA22" box="[839,949,1425,1451]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="10" pageNumber="339">Figure 4I</figureCitation>
). Ratio distance vulva-posterior extremity/ body length 48.9% (39.355.6%). Anus 245 (180300) from caudal extremity (
<figureCitation id="137FF3A42C23FF9AAC40AD819683FA46" box="[1065,1189,1461,1487]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="10" pageNumber="339">Figure 4G</figureCitation>
). Tail elongated, subcylindrical with two small terminal digitiform processes (
<figureCitation id="137FF3A42C23FF9AAB73ADED91BAFA7A" box="[794,924,1497,1523]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="10" pageNumber="339">Figures 4G</figureCitation>
,
<figureCitation id="137FF3A42C23FF9AABCEADED91EEFA7D" box="[935,968,1497,1524]" captionStart="FIGURE 5" captionStartId="10.[151,250,1211,1235]" captionTargetBox="[165,1421,181,1187]" captionTargetId="figure-25@10.[151,1435,181,1187]" captionTargetPageId="10" captionText="FIGURE 5. Scanning electron micrographs of Bidigiticauda vivipara. A) Female anterior extremity, lateral view. B) Female head, apical view. C) Male posterior extremity, lateral view. D) Male caudal bursa, ventral view. E) Magnified image of the dorsal lip of the genital cone.F) Female posterior extremity, dorsal view.Abbreviations:dl, dorsolateral papillae; vl, ventrolateral papillae; a, amphids; p7, papillae 7. Scale bars: A, F, 50 = μm; B, D = 40 μm; C = 100 μm; E = 8 μm" pageId="10" pageNumber="339">5F</figureCitation>
).
</paragraph>
<caption id="DF3BBFA92C20FF99A8FEAF419746F870" pageId="9" pageNumber="338" startId="9.[151,250,1909,1933]" targetBox="[202,1385,185,1882]" targetPageId="9" targetType="figure">
<paragraph id="8BFBEF212C20FF99A8FEAF419746F870" blockId="9.[151,1437,1909,2041]" pageId="9" pageNumber="338">
<emphasis id="B93033332C20FF99A8FEAF419333F804" bold="true" box="[151,277,1909,1933]" pageId="9" pageNumber="338">FIGURE 4.</emphasis>
<taxonomicName id="4C4494A22C20FF99A977AF419022F804" box="[286,516,1909,1933]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="9" pageNumber="338" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C20FF99A977AF419022F804" box="[286,516,1909,1933]" italics="true" pageId="9" pageNumber="338">Bidigiticauda vivipara</emphasis>
</taxonomicName>
. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view. Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm.
</paragraph>
</caption>
<caption id="DF3BBFA92C23FF9AA8FEAC8F96B4FAC9" pageId="10" pageNumber="339" startId="10.[151,250,1211,1235]" targetBox="[165,1421,181,1187]" targetPageId="10" targetType="figure">
<paragraph id="8BFBEF212C23FF9AA8FEAC8F96B4FAC9" blockId="10.[151,1437,1211,1344]" pageId="10" pageNumber="339">
<emphasis id="B93033332C23FF9AA8FEAC8F9332FB5A" bold="true" box="[151,276,1211,1235]" pageId="10" pageNumber="339">FIGURE 5.</emphasis>
Scanning electron micrographs of
<taxonomicName id="4C4494A22C23FF9AAA16AC8F9143FB5A" box="[639,869,1211,1235]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="339" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C23FF9AAA16AC8F9143FB5A" box="[639,869,1211,1235]" italics="true" pageId="10" pageNumber="339">Bidigiticauda vivipara</emphasis>
</taxonomicName>
. A) Female anterior extremity, lateral view. B) Female head, apical view. C) Male posterior extremity, lateral view. D) Male caudal bursa, ventral view. E) Magnified image of the dorsal lip of the genital cone. F) Female posterior extremity, dorsal view.Abbreviations: dl, dorsolateral papillae; vl, ventrolateral papillae; a, amphids; p7, papillae 7. Scale bars: A, F, 50 = μm; B, D = 40 μm; C = 100 μm; E = 8 μm
</paragraph>
</caption>
<paragraph id="8BFBEF212C23FF9AA8AEADC990EEF90A" blockId="10.[151,1437,1425,2028]" pageId="10" pageNumber="339">
Third-stage larvae: Body length 538 (510608) and width 35 (3537). Esophagus 176 (165200) long. Anterior end showing rod-like structure, 11 (1012) long. Nerve ring and excretory pore situated at 76 (7080) and 91 (70 110) from apex, respectively (
<figureCitation id="137FF3A42C23FF9AA980AE71907EF9D6" box="[489,600,1605,1631]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="10" pageNumber="339">Figure 4J</figureCitation>
). Genital primordium and anus located 185 (138512205) and 48 (4550) from posterior extremity, respectively (
<figureCitation id="137FF3A42C23FF9AAA25AE5D909CF90A" box="[588,698,1641,1667]" captionStart="FIGURE 4" captionStartId="9.[151,250,1909,1933]" captionTargetBox="[202,1385,185,1882]" captionTargetId="figure-18@9.[198,1389,181,1885]" captionTargetPageId="9" captionText="FIGURE 4. Bidigiticauda vivipara. A) Female anterior extremity, ventral view. B) Female transverse body section, at midbody. C) Male transverse body section, at mid-body. DF) Male caudal bursae, ventral views. G) Female posterior extremity, lateral view. H) Right spicule, lateral view. I) Vulvar aperture, ventral view. J) Third-stage larvae, lateral view.Abbreviations: Et, esophageal tooth; Nr, nerve ring; Ep, excretory pore; De, deirid; Gp, genital primordium; An, anus. Scale bars = 100 μm." pageId="10" pageNumber="339">Figure 4J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35EBCAA2C23FF9BA8AEAEB9938AFD78" lastPageId="11" lastPageNumber="340" pageId="10" pageNumber="339" type="discussion">
<paragraph id="8BFBEF212C23FF9BA8AEAEB99742FE00" blockId="10.[151,1437,1425,2028]" lastBlockId="11.[151,1437,151,753]" lastPageId="11" lastPageNumber="340" pageId="10" pageNumber="339">
<emphasis id="B93033332C23FF9AA8AEAEB99312F92E" bold="true" box="[199,308,1677,1703]" pageId="10" pageNumber="339">Remarks</emphasis>
: The bursal pattern and dorsal lobe of the male specimens from
<collectingRegion id="498021C32C23FF9AAB8FAEB9961CF92E" box="[998,1082,1677,1703]" country="Mexico" name="Nayarit" pageId="10" pageNumber="339">Nayarit</collectingRegion>
agrees with the brief description of
<taxonomicName id="4C4494A22C23FF9AA8D0AE86931FF942" box="[185,313,1714,1740]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="339" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C23FF9AA8D0AE8692ECF942" box="[185,202,1714,1739]" italics="true" pageId="10" pageNumber="339">B</emphasis>
.
<emphasis id="B93033332C23FF9AA8B5AE86931FF942" box="[220,313,1714,1739]" italics="true" pageId="10" pageNumber="339">vivipara</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EFD592D02C23FF9AA9B3AE8590BBF942" author="Chitwood, B. G." box="[474,669,1713,1739]" pageId="10" pageNumber="339" pagination="51 - 66" refId="ref8856" refString="Chitwood, B. G. (1938) Some nematodes from the caves of Yucatan. Publications of the Carnegie Institution of Washington, 491, 51 - 66." type="journal article" year="1938">Chitwood (1938)</bibRefCitation>
and the redescription by Caballero-Deloya (1971). In
<collectingRegion id="498021C32C23FF9AAD74AE859753F942" box="[1309,1397,1713,1739]" country="Mexico" name="Mexico" pageId="10" pageNumber="339">Mexico</collectingRegion>
,
<taxonomicName id="4C4494A22C23FF9AADEDAE8692D2F966" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="339" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C23FF9AADEDAE8697B3F942" box="[1412,1429,1714,1739]" italics="true" pageId="10" pageNumber="339">B</emphasis>
.
<emphasis id="B93033332C23FF9AA8FEAEE292D2F966" box="[151,244,1750,1775]" italics="true" pageId="10" pageNumber="339">vivipara</emphasis>
</taxonomicName>
has been reported from
<taxonomicName id="4C4494A22C23FF9AAA65AEE29097F966" box="[524,689,1750,1776]" class="Enoplea" family="Capillariidae" genus="Aonchotheca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="339" phylum="Nematoda" rank="species" species="jamaicensis">
<emphasis id="B93033332C23FF9AAA65AEE2903BF979" box="[524,541,1750,1776]" italics="true" pageId="10" pageNumber="339">A</emphasis>
.
<emphasis id="B93033332C23FF9AAA44AEE29097F966" box="[557,689,1750,1775]" italics="true" pageId="10" pageNumber="339">jamaicensis</emphasis>
</taxonomicName>
in
<collectingRegion id="498021C32C23FF9AAABFAEE29116F979" box="[726,816,1750,1776]" country="Mexico" name="Yucatan" pageId="10" pageNumber="339">Yucatan</collectingRegion>
(
<bibRefCitation id="EFD592D02C23FF9AAB29AEE191D3F966" author="Chitwood, B. G." box="[832,1013,1749,1775]" pageId="10" pageNumber="339" pagination="51 - 66" refId="ref8856" refString="Chitwood, B. G. (1938) Some nematodes from the caves of Yucatan. Publications of the Carnegie Institution of Washington, 491, 51 - 66." type="journal article" year="1938">Chitwood 1938</bibRefCitation>
),
<taxonomicName id="4C4494A22C23FF9AAC63AEE296AFF966" box="[1034,1161,1749,1776]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B93033332C23FF9AAC63AEE2963DF979" box="[1034,1051,1750,1776]" italics="true" pageId="10" pageNumber="339">A</emphasis>
.
<emphasis id="B93033332C23FF9AAC43AEE196AFF966" box="[1066,1161,1749,1775]" italics="true" pageId="10" pageNumber="339">lituratus</emphasis>
</taxonomicName>
in
<collectingRegion id="498021C32C23FF9AACC6AEE19732F966" box="[1199,1300,1749,1775]" country="Mexico" name="Guerrero" pageId="10" pageNumber="339">Guerrero</collectingRegion>
(Caballero-Deloya 1971), and
<taxonomicName id="4C4494A22C23FF9AA900AECE902CF89A" box="[361,522,1786,1812]" class="Enoplea" family="Capillariidae" genus="Aonchotheca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="339" phylum="Nematoda" rank="species" species="jamaicensis">
<emphasis id="B93033332C23FF9AA900AECE935CF89D" box="[361,378,1786,1812]" italics="true" pageId="10" pageNumber="339">A</emphasis>
.
<emphasis id="B93033332C23FF9AA9EFAECE902CF89A" box="[390,522,1786,1811]" italics="true" pageId="10" pageNumber="339">jamaicensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4494A22C23FF9AAA54AECE909CF89A" box="[573,698,1785,1812]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="10" pageNumber="339" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B93033332C23FF9AAA54AECE909CF89A" box="[573,698,1785,1812]" italics="true" pageId="10" pageNumber="339">A. lituratus</emphasis>
</taxonomicName>
in
<collectingRegion id="498021C32C23FF9AAAB3AECD9112F89A" box="[730,820,1785,1811]" country="Mexico" name="Chiapas" pageId="10" pageNumber="339">Chiapas</collectingRegion>
(
<bibRefCitation id="EFD592D02C23FF9AAB28AECD9600F89A" author="Ubelaker, J. E. &amp; Specian, R. D. &amp; Duszynski, D. W." box="[833,1062,1785,1811]" pageId="10" pageNumber="339" pagination="7 - 56" refId="ref12318" refString="Ubelaker, J. E., Specian, R. D. &amp; Duszynski, D. W. (1977) Endoparasites. In: Baker, R. J., Jones, J. K. &amp; Carter, D. C. (Eds.), Biology of bats of the New World family Phyllostomatidae. Part II. Texas Tech University, Texas, pp. 7 - 56." type="book chapter" year="1977">
Ubelaker
<emphasis id="B93033332C23FF9AABC6AECE91C3F89A" box="[943,997,1785,1811]" italics="true" pageId="10" pageNumber="339">et al.</emphasis>
1977
</bibRefCitation>
). However, the present specimens were smaller than those reported by Chitwood, (1938) and Caballero-Deloya (1971) (male body length 34804250 vs 5120 and vs 1038410554; female body length 54807670 vs 80208060 and vs 1193211967). Similarly, the spicule length of our specimens was smaller than those described by Chitwood and Caballero-Deloya (160200 vs 256270 and vs 235252). Nevertheless, the ratio spicule length/body length was similar between the
<collectingRegion id="498021C32C23FF9AAD71AFBD974AF82A" box="[1304,1388,1929,1955]" country="Mexico" name="Nayarit" pageId="10" pageNumber="339">Nayarit</collectingRegion>
and
<collectingRegion id="498021C32C23FF9AA8FEAF9A92D7F841" box="[151,241,1966,1992]" country="Mexico" name="Yucatan" pageId="10" pageNumber="339">Yucatan</collectingRegion>
specimens (45.3 vs 5). In contrast, this ratio was longer compared to Caballero-Deloyas material (45.3 vs 2.22.4). The difference in size among these
<taxonomicName id="4C4494A22C23FF9AAACEAFE69104F862" box="[679,802,2002,2028]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="10" pageNumber="339" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C23FF9AAACEAFE6909EF862" box="[679,696,2002,2027]" italics="true" pageId="10" pageNumber="339">B</emphasis>
.
<emphasis id="B93033332C23FF9AAAACAFE69104F862" box="[709,802,2002,2027]" italics="true" pageId="10" pageNumber="339">vivipara</emphasis>
</taxonomicName>
isolates may be attributed to their occurrence in different host species (
<taxonomicName id="4C4494A22C22FF9BA947A8A393F7FF39" box="[302,465,151,177]" class="Enoplea" family="Capillariidae" genus="Aonchotheca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="jamaicensis">
<emphasis id="B93033332C22FF9BA947A8A39319FF38" box="[302,319,151,177]" italics="true" pageId="11" pageNumber="340">A</emphasis>
.
<emphasis id="B93033332C22FF9BA924A8A393F7FF39" box="[333,465,151,176]" italics="true" pageId="11" pageNumber="340">jamaicensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4494A22C22FF9BA989A8A39078FF38" box="[480,606,151,177]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="lituratus">
<emphasis id="B93033332C22FF9BA989A8A393D7FF38" box="[480,497,151,177]" italics="true" pageId="11" pageNumber="340">A</emphasis>
.
<emphasis id="B93033332C22FF9BA996A8A39078FF38" box="[511,606,151,177]" italics="true" pageId="11" pageNumber="340">lituratus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C4494A22C22FF9BAAF5A8A3912CFF39" authorityName="Merriam" authorityYear="1898" box="[668,778,151,177]" class="Mammalia" family="Phyllostomidae" genus="Glossophaga" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="mutica">
<emphasis id="B93033332C22FF9BAAF5A8A39096FF38" box="[668,688,151,177]" italics="true" pageId="11" pageNumber="340">G</emphasis>
.
<emphasis id="B93033332C22FF9BAAD7A8A3912CFF39" box="[702,778,151,176]" italics="true" pageId="11" pageNumber="340">mutica</emphasis>
</taxonomicName>
), as well as the fact that each morphological description is based on nematodes isolated from a single individual host. Furthermore, Chitwoods original description was based on
<specimenCount id="9D4224A82C22FF9BA8D5A8EB9302FF70" box="[188,292,223,249]" count="1" pageId="11" pageNumber="340" type="male">one male</specimenCount>
and
<specimenCount id="9D4224A82C22FF9BA936A8EB93CBFF70" box="[351,493,223,249]" count="2" pageId="11" pageNumber="340" type="female">two females</specimenCount>
, whereas Caballero-Deloya redescribed the species without specifying the number of specimens measured, although he collected
<specimenCount id="9D4224A82C22FF9BAAC7A937916EFE94" box="[686,840,259,285]" count="49" pageId="11" pageNumber="340" type="generic">49 specimens</specimenCount>
(Caballero-Deloya 1971). Based on these data, it is difficult to determine at which point along this morphological and morphometrical continuum a
<taxonomicName id="4C4494A22C22FF9BACA6A913976AFEC9" box="[1231,1356,295,321]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C22FF9BACA6A91396C6FEC9" box="[1231,1248,295,320]" italics="true" pageId="11" pageNumber="340">B</emphasis>
.
<emphasis id="B93033332C22FF9BAC86A913976AFEC9" box="[1263,1356,295,320]" italics="true" pageId="11" pageNumber="340">vivipara</emphasis>
</taxonomicName>
isolate may be considered a different species. Further studies incorporating morphological and molecular data from several host species and geographical locations are necessary to investigate whether
<taxonomicName id="4C4494A22C22FF9BAB9AA95B9649FE01" box="[1011,1135,367,393]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C22FF9BAB9AA95B9622FE01" box="[1011,1028,367,392]" italics="true" pageId="11" pageNumber="340">B</emphasis>
.
<emphasis id="B93033332C22FF9BAC7BA95B9649FE01" box="[1042,1135,367,392]" italics="true" pageId="11" pageNumber="340">vivipara</emphasis>
</taxonomicName>
is a species complex.
</paragraph>
<paragraph id="8BFBEF212C22FF9BA8AEA9A792C6FD20" blockId="11.[151,1437,151,753]" pageId="11" pageNumber="340">
The genus
<taxonomicName id="4C4494A22C22FF9BA929A9A793FEFE24" authorityName="Chitwood" authorityYear="1938" box="[320,472,403,429]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="genus">
<emphasis id="B93033332C22FF9BA929A9A793FEFE24" box="[320,472,403,429]" italics="true" pageId="11" pageNumber="340">Bidigiticauda</emphasis>
</taxonomicName>
contains two other species:
<taxonomicName id="4C4494A22C22FF9BAB79A9A7970CFE24" authority="(Freitas &amp; Dobbin)" baseAuthorityName="Freitas &amp; Dobbin" box="[784,1322,403,429]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="embryophilum">
<emphasis id="B93033332C22FF9BAB79A9A79676FE24" box="[784,1104,403,429]" italics="true" pageId="11" pageNumber="340">Bidigiticauda embryophilum</emphasis>
(Freitas &amp; Dobbin)
</taxonomicName>
described from
<taxonomicName id="4C4494A22C22FF9BA8BCA983900EFE58" authority="(Pallas)" baseAuthorityName="Pallas" baseAuthorityYear="1766" box="[213,552,439,465]" class="Mammalia" family="Phyllostomidae" genus="Glossophaga" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="soricina">
<emphasis id="B93033332C22FF9BA8BCA98393EDFE59" box="[213,459,439,465]" italics="true" pageId="11" pageNumber="340">Glossophaga soricina</emphasis>
(Pallas)
</taxonomicName>
in
<collectingCountry id="F353AFB12C22FF9BAA22A98390B4FE58" box="[587,658,439,465]" name="Brazil" pageId="11" pageNumber="340">Brazil</collectingCountry>
, and
<taxonomicName id="4C4494A22C22FF9BAAA4A98393F2FE7C" authority="de Oliveira Simoes, Fraga-Neto, Vilar Maldonado Junior &amp; Vilela" authorityName="de Oliveira Simoes, Fraga-Neto, Vilar Maldonado Junior &amp; Vilela" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="serrafreirei">
<emphasis id="B93033332C22FF9BAAA4A98391CAFE58" box="[717,1004,439,465]" italics="true" pageId="11" pageNumber="340">Bidigiticauda serrafreirei</emphasis>
de Oliveira Simões, Fraga-Neto, Vilar Maldonado Júnior &amp; Vilela
</taxonomicName>
described from
<taxonomicName id="4C4494A22C22FF9BAAFCA9EF91F5FE7C" authority="(Spix)" baseAuthorityName="Spix" baseAuthorityYear="1823" box="[661,979,475,501]" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="planirostris">
<emphasis id="B93033332C22FF9BAAFCA9EF91A6FE7C" box="[661,896,475,501]" italics="true" pageId="11" pageNumber="340">Artibeus planirostris</emphasis>
(Spix)
</taxonomicName>
, in
<collectingCountry id="F353AFB12C22FF9BAC6BA9EF966CFE7C" box="[1026,1098,475,501]" name="Brazil" pageId="11" pageNumber="340">Brazil</collectingCountry>
.
<taxonomicName id="4C4494A22C22FF9BAC31A9EF97BAFE7C" box="[1112,1436,475,501]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="embryophilum">
<emphasis id="B93033332C22FF9BAC31A9EF97BAFE7C" box="[1112,1436,475,501]" italics="true" pageId="11" pageNumber="340">Bidigiticauda embryophilum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4494A22C22FF9BA94FA9CB9382FD91" box="[294,420,511,537]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C22FF9BA94FA9CB9311FD91" box="[294,311,511,536]" italics="true" pageId="11" pageNumber="340">B</emphasis>
.
<emphasis id="B93033332C22FF9BA92EA9CB9382FD91" box="[327,420,511,536]" italics="true" pageId="11" pageNumber="340">vivipara</emphasis>
</taxonomicName>
by having rays 5 and 6 arising independently, the branches of the dorsal ray not reaching the edge of the caudal bursa, and deirids situated posterior to the esophagus-intestinal junction (
<bibRefCitation id="EFD592D02C22FF9BACD1AA1797B6FDB4" author="Vicente, J. J. &amp; Rodrigues, H. O. &amp; Gomes, D. C. &amp; Pinto, R." box="[1208,1424,547,573]" pageId="11" pageNumber="340" pagination="1 - 452" refId="ref12395" refString="Vicente, J. J., Rodrigues, H. O., Gomes, D. C. &amp; Pinto, R. M (1997). Nematoides do Brasil. Parte 625 V: nematoides de mamiferos. Revista Brasileira de Zoologia, 14, 1 - 452." type="journal article" year="1997">
Vicente
<emphasis id="B93033332C22FF9BAD7FAA179760FDB4" box="[1302,1350,547,573]" italics="true" pageId="11" pageNumber="340">et al</emphasis>
.,1997
</bibRefCitation>
).
<taxonomicName id="4C4494A22C22FF9BA8FEAA739391FDE8" box="[151,439,583,609]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="serrafreirei">
<emphasis id="B93033332C22FF9BA8FEAA739391FDE8" box="[151,439,583,609]" italics="true" pageId="11" pageNumber="340">Bidigiticauda serrafreirei</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C4494A22C22FF9BAA84AA73914CFDE9" box="[749,874,583,609]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C22FF9BAA84AA7390D8FDE9" box="[749,766,583,608]" italics="true" pageId="11" pageNumber="340">B</emphasis>
.
<emphasis id="B93033332C22FF9BAB64AA73914CFDE9" box="[781,874,583,608]" italics="true" pageId="11" pageNumber="340">vivipara</emphasis>
</taxonomicName>
by the bifurcation of the dorsal ray in the middle of the trunk and the distance of the excretory pore from the anterior end that is longer (de Oliveira Simões
<emphasis id="B93033332C22FF9BAD35AA5F97A8FD0C" box="[1372,1422,619,645]" italics="true" pageId="11" pageNumber="340">et al</emphasis>
., 2019).
</paragraph>
<paragraph id="8BFBEF212C22FF9BA8AEAA87938AFD78" blockId="11.[151,1437,151,753]" pageId="11" pageNumber="340">
The new record of
<taxonomicName id="4C4494A22C22FF9BA9C3AA87900FFD45" box="[426,553,691,717]" class="Chromadorea" family="Molineidae" genus="Bidigiticauda" kingdom="Animalia" order="Rhabditida" pageId="11" pageNumber="340" phylum="Nematoda" rank="species" species="vivipara">
<emphasis id="B93033332C22FF9BA9C3AA87939DFD45" box="[426,443,691,716]" italics="true" pageId="11" pageNumber="340">B</emphasis>
.
<emphasis id="B93033332C22FF9BA9A5AA87900FFD45" box="[460,553,691,716]" italics="true" pageId="11" pageNumber="340">vivipara</emphasis>
</taxonomicName>
from
<taxonomicName id="4C4494A22C22FF9BAA1CAA8790C1FD45" authorityName="Merriam" authorityYear="1898" box="[629,743,691,717]" class="Mammalia" family="Phyllostomidae" genus="Glossophaga" kingdom="Animalia" order="Chiroptera" pageId="11" pageNumber="340" phylum="Chordata" rank="species" species="mutica">
<emphasis id="B93033332C22FF9BAA1CAA8790AFFD44" box="[629,649,691,717]" italics="true" pageId="11" pageNumber="340">G</emphasis>
.
<emphasis id="B93033332C22FF9BAAF2AA8790C1FD45" box="[667,743,691,716]" italics="true" pageId="11" pageNumber="340">mutica</emphasis>
</taxonomicName>
in
<collectingRegion id="498021C32C22FF9BAB78AA879143FD44" box="[785,869,691,717]" country="Mexico" name="Nayarit" pageId="11" pageNumber="340">Nayarit</collectingRegion>
expands the host and geographical range of this nematode in the country.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,257 @@
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<mods:classification id="DFA25ABCCFCBA53B286F89B6E3BA39FE">journal article</mods:classification>
<mods:identifier id="AF62D9B5588245398943B6465D0C6FAC" type="DOI">10.11646/zootaxa.5566.2.5</mods:identifier>
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<mods:identifier id="5C53035B9A7E0B5EC1946981972DD497" type="ZooBank">9A17BF2C-EB04-4215-9783-46E0C45BB126</mods:identifier>
</mods:mods>
<treatment id="03ED5E372C2AFF94A8FEAEDE9032FD44" LSID="urn:lsid:plazi:treatment:03ED5E372C2AFF94A8FEAEDE9032FD44" httpUri="http://treatment.plazi.org/id/03ED5E372C2AFF94A8FEAEDE9032FD44" lastPageId="4" lastPageNumber="333" pageId="3" pageNumber="332">
<subSubSection id="C35EBCAA2C2AFF93A8FEAEDE902FF8C5" pageId="3" pageNumber="332" type="nomenclature">
<paragraph id="8BFBEF212C2AFF93A8FEAEDE9130F88D" blockId="3.[151,790,1770,1797]" box="[151,790,1770,1797]" pageId="3" pageNumber="332">
<heading id="D0B3584D2C2AFF93A8FEAEDE9130F88D" bold="true" box="[151,790,1770,1797]" fontSize="11" level="1" pageId="3" pageNumber="332" reason="1">
<taxonomicName id="4C4494A22C2AFF93A8FEAEDE9130F88D" authority="Cacique, Cruces &amp; Chero" authorityName="Cacique, Cruces &amp; Chero" box="[151,790,1770,1797]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Plagiorchiida" pageId="3" pageNumber="332" phylum="Platyhelminthes" rank="species" species="hastati">
<emphasis id="B93033332C2AFF93A8FEAEDE9130F88D" bold="true" box="[151,790,1770,1797]" pageId="3" pageNumber="332">
<emphasis id="B93033332C2AFF93A8FEAEDE9368F88D" bold="true" box="[151,334,1770,1796]" italics="true" pageId="3" pageNumber="332">Anenterotrema</emphasis>
cf.
<emphasis id="B93033332C2AFF93A912AEDE93EBF88C" bold="true" box="[379,461,1770,1797]" italics="true" pageId="3" pageNumber="332">hastati</emphasis>
Cacique, Cruces &amp; Chero
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFBEF212C2AFF93A8FEAF06902FF8C5" blockId="3.[151,1358,1842,2012]" box="[151,521,1842,1868]" pageId="3" pageNumber="332">
<emphasis id="B93033332C2AFF93A8FEAF06937DF8C5" bold="true" box="[151,347,1842,1868]" pageId="3" pageNumber="332">Site of infection:</emphasis>
Small intestine
</paragraph>
</subSubSection>
<subSubSection id="C35EBCAA2C2AFF93A8AEAF62907AF855" pageId="3" pageNumber="332" type="biology_ecology">
<paragraph id="8BFBEF212C2AFF93A8AEAF629042F8F9" blockId="3.[151,1358,1842,2012]" box="[199,612,1878,1904]" pageId="3" pageNumber="332">
<emphasis id="B93033332C2AFF93A8AEAF629344F8F9" bold="true" box="[199,354,1878,1904]" pageId="3" pageNumber="332">Host species:</emphasis>
<taxonomicName id="4C4494A22C2AFF93A900AF639042F8F9" box="[361,612,1878,1904]" class="Mammalia" family="Vespertilionidae" genus="Bauerus" kingdom="Animalia" order="Chiroptera" pageId="3" pageNumber="332" phylum="Chordata" rank="species" species="dubiaquercus">
<emphasis id="B93033332C2AFF93A900AF639042F8F9" box="[361,612,1878,1904]" italics="true" pageId="3" pageNumber="332">Bauerus dubiaquercus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BFBEF212C2AFF93A8AEAF4E90EBF830" blockId="3.[151,1358,1842,2012]" pageId="3" pageNumber="332">
<emphasis id="B93033332C2AFF93A8AEAF4E9315F81D" bold="true" box="[199,307,1914,1940]" pageId="3" pageNumber="332">Locality:</emphasis>
<materialsCitation id="3B2CE57C2C2AFF93A950AF4E90EBF830" collectionCode="MASL" latitude="18.53161" location="Prevalence" longLatPrecision="1" longitude="-95.133804" municipality="Tuxtlas" pageId="3" pageNumber="332" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
Ejido Adolfo Ruiz Cortines (
<geoCoordinate id="EE7089E62C2AFF93AA15AF4E9103F81C" box="[636,805,1914,1941]" degrees="18" direction="north" minutes="31" orientation="latitude" pageId="3" pageNumber="332" precision="1" seconds="53.8" value="18.53161">18°3153.8” N</geoCoordinate>
,
<geoCoordinate id="EE7089E62C2AFF93AB59AF4E91FAF81C" box="[816,988,1914,1941]" degrees="95" direction="west" minutes="08" orientation="longitude" pageId="3" pageNumber="332" precision="1" seconds="01.7" value="-95.133804">95°0801.7” W</geoCoordinate>
, 1061
<collectionCode id="ED5577E42C2AFF93AC4CAF4E9651F81D" box="[1061,1143,1914,1940]" pageId="3" pageNumber="332">MASL</collectionCode>
),
<collectingMunicipality id="6B9F755B2C2AFF93ACE3AF4E96C6F81D" box="[1162,1248,1914,1940]" pageId="3" pageNumber="332">Tuxtlas</collectingMunicipality>
,
<collectingRegion id="498021C32C2AFF93AC82AF4F9768F81C" box="[1259,1358,1915,1941]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="332">Veracruz</collectingRegion>
<emphasis id="B93033332C2AFF93A8AEAFAA93D3F831" bold="true" box="[199,501,1950,1976]" pageId="3" pageNumber="332">
<location id="8E9BB9FA2C2AFF93A8AEAFAA936FF831" LSID="urn:lsid:plazi:treatment:03ED5E372C2AFF94A8FEAEDE9032FD44:8E9BB9FA2C2AFF93A8AEAFAA936FF831" box="[199,329,1950,1976]" latitude="18.53161" longLatPrecision="1" longitude="-95.133804" municipality="Tuxtlas" name="Prevalence" pageId="3" pageNumber="332" stateProvince="Veracruz de Ignacio de la Llave">Prevalence</location>
and intensity:
</emphasis>
50% (1/2) and 151
</materialsCitation>
</paragraph>
<paragraph id="8BFBEF212C2AFF93A8AEAFF6907AF855" blockId="3.[151,1358,1842,2012]" box="[199,604,1986,2012]" pageId="3" pageNumber="332">
<emphasis id="B93033332C2AFF93A8AEAFF6939AF855" bold="true" box="[199,444,1986,2012]" pageId="3" pageNumber="332">Specimen deposited:</emphasis>
CNHE 12134
</paragraph>
</subSubSection>
<subSubSection id="C35EBCAA2C2DFF94A8AEA8A29032FD44" pageId="4" pageNumber="333" type="discussion">
<paragraph id="8BFBEF212C2DFF94A8AEA8A29777FEEC" blockId="4.[151,1437,150,717]" pageId="4" pageNumber="333">
<emphasis id="B93033332C2DFF94A8AEA8A29374FF39" bold="true" box="[199,338,150,176]" pageId="4" pageNumber="333">Comments:</emphasis>
Based on five trematodes. Body pyriform, minute, 225 (165330) long and 123 (105151) wide (
<figureCitation id="137FF3A42C2DFF94A8F6A88F9331FF5C" box="[159,279,187,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,1874,1898]" captionTargetBox="[168,1419,717,1834]" captionTargetId="figure-165@5.[151,1436,701,1851]" captionTargetPageId="5" captionText="FIGURE 2. Helminths of bats from Nayarit and Veracruz, Mexico. A) Adult of Anenterotrema cf. hastati, ventral view. B) Adult of Urotrema scabridum, ventral view. C) Scolex of Vampirolepis macroti, apical view. D) Scolex of Vampirolepis sp. (1), apical view. E) Scolex of Vampirolepis sp. (2), lateral view. F) Male posterior extremity of Capillariidae gen. sp., ventral view. G) Esophagus-intestine junction and vulva of female Capillariidae gen. sp., lateral view. Scale bars: A, C, D, E = 100 μm; B = 1000 μm; F = 10 μm; G = 65 μm." pageId="4" pageNumber="333">Figure 2A</figureCitation>
). Tegument unarmed. Oral sucker spherical, subterminal, 50 (5051) long by 52 (5056) wide. Ventral sucker in middle third of body, 55 (5060) long by 69 (6274) wide. Pharynx, oesophagus, and intestinal caeca not observed. Gonads in anterior half of hindbody. Testes postero-lateral to ventral sucker; right testis 42 long by 70 wide, left testis 55 long by 62 wide. Ovary median, post-testicular, 32 long by 53 wide. Vitellaria in hindbody, concentrated in lateral region, occupying almost entire left testis. Eggs 27 (2530) long by 13 (1115) wide.
</paragraph>
<paragraph id="8BFBEF212C2DFF94A8AEA95B9032FD44" blockId="4.[151,1437,150,717]" pageId="4" pageNumber="333">
The closest species in morphology is
<taxonomicName id="4C4494A22C2DFF94AAF8A95B9125FE00" box="[657,771,367,393]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="hastati">
<emphasis id="B93033332C2DFF94AAF8A95B9084FE00" box="[657,674,367,393]" italics="true" pageId="4" pageNumber="333">A</emphasis>
.
<emphasis id="B93033332C2DFF94AADEA95B9125FE00" box="[695,771,367,393]" italics="true" pageId="4" pageNumber="333">hastati</emphasis>
</taxonomicName>
described from
<taxonomicName id="4C4494A22C2DFF94ABA4A95B9714FE00" authority="(Pallas)" baseAuthorityName="Pallas" baseAuthorityYear="1767" box="[973,1330,367,393]" class="Mammalia" family="Phyllostomidae" genus="Phyllostomus" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="hastatus">
<emphasis id="B93033332C2DFF94ABA4A95B96E8FE00" box="[973,1230,367,393]" italics="true" pageId="4" pageNumber="333">Phyllostomus hastatus</emphasis>
(Pallas)
</taxonomicName>
in
<collectingCountry id="F353AFB12C2DFF94AD0BA95B97BFFE00" box="[1378,1433,367,393]" name="Peru" pageId="4" pageNumber="333">Peru</collectingCountry>
. However, we were unable to observe the characteristic unicellular glands located anterolateral to cirrus sac due to the poor condition of preservation of our specimens. The remaining species,
<taxonomicName id="4C4494A22C2DFF94AC09A983933AFE7C" authority="(Travassos)" baseAuthorityName="Travassos" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="liliputianum">
<emphasis id="B93033332C2DFF94AC09A98397BAFE58" box="[1120,1436,439,465]" italics="true" pageId="4" pageNumber="333">Anenterotrema liliputianum</emphasis>
(Travassos)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94A94FA9EF908AFE7C" authority="(Stunkard)" baseAuthorityName="Stunkard" box="[294,684,475,501]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="auritum">
<emphasis id="B93033332C2DFF94A94FA9EF900DFE7D" box="[294,555,475,501]" italics="true" pageId="4" pageNumber="333">Anenterotrema auritum</emphasis>
(Stunkard)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94AADFA9EF96BEFE7C" authority="(Freitas)" baseAuthorityName="Freitas" box="[694,1176,475,501]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="eduardocaballeroi">
<emphasis id="B93033332C2DFF94AADFA9EF9616FE7C" box="[694,1072,475,501]" italics="true" pageId="4" pageNumber="333">Anenterotrema eduardocaballeroi</emphasis>
(Freitas)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94ACCBA9EF933CFD90" authority="(Caballero)" baseAuthorityName="Caballero" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="freitasi">
<emphasis id="B93033332C2DFF94ACCBA9EF97BAFE7C" box="[1186,1436,475,501]" italics="true" pageId="4" pageNumber="333">Anenterotrema freitasi</emphasis>
(Caballero)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94A945A9CB914EFD90" authority="(Marshall &amp; Miller)" baseAuthorityName="Marshall &amp; Miller" box="[300,872,511,537]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="mesolecitha">
<emphasis id="B93033332C2DFF94A945A9CB9041FD90" box="[300,615,511,537]" italics="true" pageId="4" pageNumber="333">Anenterotrema mesolecitha</emphasis>
(Marshall &amp; Miller)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94AB12A9CB968AFD91" box="[891,1196,511,537]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="iannaconei">
<emphasis id="B93033332C2DFF94AB12A9CB968AFD91" box="[891,1196,511,537]" italics="true" pageId="4" pageNumber="333">Anenterotrema iannaconei</emphasis>
</taxonomicName>
(Achatz, Cardenaz- Callirgos &amp; Tkach),
<taxonomicName id="4C4494A22C2DFF94A912AA1797BEFDB4" authority="(Fernandes, Santos, Melo, Achatz, Greiman, Bonilla &amp; Tkach)" baseAuthorityName="Fernandes, Santos, Melo, Achatz, Greiman, Bonilla &amp; Tkach" box="[379,1432,547,573]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="megacetabulum">
<emphasis id="B93033332C2DFF94A912AA1790FEFDB4" box="[379,728,547,573]" italics="true" pageId="4" pageNumber="333">Anenterotrema megacetabulum</emphasis>
(Fernandes, Santos, Melo, Achatz, Greiman, Bonilla &amp; Tkach)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94A8FEAA739157FDE8" authority="(Cacique, Cruces &amp; Chero)" baseAuthorityName="Cacique, Cruces &amp; Chero" box="[151,881,583,609]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="paramegacetabulum">
<emphasis id="B93033332C2DFF94A8FEAA73900AFDE8" box="[151,556,583,609]" italics="true" pageId="4" pageNumber="333">Anenterotrema paramegacetabulum</emphasis>
(Cacique, Cruces &amp; Chero)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94AB17AA7392CEFD0C" authority="(Cacique, Cruces &amp; Chero)" baseAuthorityName="Cacique, Cruces &amp; Chero" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="kawsayense">
<emphasis id="B93033332C2DFF94AB17AA739697FDE8" box="[894,1201,583,609]" italics="true" pageId="4" pageNumber="333">Anenterotrema kawsayense</emphasis>
(Cacique, Cruces &amp; Chero)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94A89AAA5F912BFD0C" authority="(Caballero &amp; Grocott)" baseAuthorityName="Caballero &amp; Grocott" box="[243,781,619,645]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="stunkardi">
<emphasis id="B93033332C2DFF94A89AAA5F902FFD0C" box="[243,521,619,645]" italics="true" pageId="4" pageNumber="333">Anenterotrema stunkardi</emphasis>
(Caballero &amp; Grocott)
</taxonomicName>
, and
<taxonomicName id="4C4494A22C2DFF94AB21AA5F97BEFD0C" authority="(Cacique, Cruces &amp; Chero)" baseAuthorityName="Cacique, Cruces &amp; Chero" box="[840,1432,619,645]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="peruense">
<emphasis id="B93033332C2DFF94AB21AA5F967CFD0D" box="[840,1114,619,645]" italics="true" pageId="4" pageNumber="333">Anenterotrema peruense</emphasis>
(Cacique, Cruces &amp; Chero)
</taxonomicName>
, differ in shape and size of oral sucker, body length, and position of gonads (
<bibRefCitation id="EFD592D02C2DFF94AB90AABB96FBFD20" author="Cacique, E. R. &amp; Bello, R. &amp; Zamora, H. &amp; Mares, K. &amp; Saez, G. &amp; Severino, R. &amp; Cruces, C. &amp; Chero, J. D." box="[1017,1245,655,681]" pageId="4" pageNumber="333" pagination="531 - 542" refId="ref8693" refString="Cacique, E. R., Bello, R., Zamora, H., Mares, K., Saez, G., Severino, R., Cruces, C. &amp; Chero, J. D. (2023) Four new species of Anenterotrema Stunkard, 1938 (Digenea: Dicrocoellidae) infecting phyllostomid bats in the Peruvian Amazon. Systematic Parasitology, 100, 531 - 542. https: // doi. org / 10.1007 / s 11230 - 023 - 10103 - 6" type="journal article" year="2023">
Cacique
<emphasis id="B93033332C2DFF94AC37AABB96BEFD20" box="[1118,1176,655,681]" italics="true" pageId="4" pageNumber="333">et al.</emphasis>
2023
</bibRefCitation>
). This is the first record of
<taxonomicName id="4C4494A22C2DFF94A96DAA8793B5FD44" box="[260,403,691,717]" class="Trematoda" family="Anenterotrematidae" genus="Anenterotrema" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="hastati">
<emphasis id="B93033332C2DFF94A96DAA879333FD44" box="[260,277,691,717]" italics="true" pageId="4" pageNumber="333">A</emphasis>
. cf.
<emphasis id="B93033332C2DFF94A92EAA8793B5FD44" box="[327,403,691,717]" italics="true" pageId="4" pageNumber="333">hastati</emphasis>
</taxonomicName>
in
<collectingCountry id="F353AFB12C2DFF94A9DEAA879036FD44" box="[439,528,691,717]" name="Mexico" pageId="4" pageNumber="333">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,226 @@
<document id="FD79CCC94A39FAA9FD40E757AF36424A" ID-DOI="10.11646/zootaxa.5566.2.5" ID-ISSN="1175-5326" ID-ZooBank="9A17BF2C-EB04-4215-9783-46E0C45BB126" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737370187565" checkinUser="plazi" docAuthor="Moguel-Chin, Wilson I., Digiani, María Celina, Macswiney, M. Cristina, Ortega, Jorge, Ospina-Garcés, Sandra M., García-Prieto, Luis, Canché-Pool, Elsy B., Robles, María Del Rosario, Ávila-López, Mariana B., Chan-Casanova, Anyela Jackelin &amp; Panti-May, Jesús Alonso" docDate="2025" docId="03ED5E372C2CFF95A8FEA8A3911AFD2C" docLanguage="en" docName="zootaxa.5566.2.5.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vampirolepis macroti Zdzitowiecki &amp; Rutkowska" docType="treatment" docVersion="1" lastPageNumber="334" masterDocId="FFD4264F2C29FF90A869A8349226FF89" masterDocTitle="Intestinal helminths of bats in the States of Nayarit and Veracruz, Mexico, with redescription of Bidigiticauda vivipara Chitwood" masterLastPageNumber="346" masterPageNumber="329" pageNumber="334" updateTime="1737370323016" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="5D1591D9DE9DC6BFF4BAC56C32E4B29E">Intestinal helminths of bats in the States of Nayarit and Veracruz, Mexico, with redescription of Bidigiticauda vivipara Chitwood</mods:title>
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<mods:namePart id="9BB146839C57D85AD0EBF5E134E063D5">Moguel-Chin, Wilson I.</mods:namePart>
<mods:affiliation id="F7ACC89CC29C41C51BD486F8330B0BB8">Facultad de Medicina Veterinaria y Zootecnia, Universidad Autónoma de Yucatán, Mérida Yucatán, México.</mods:affiliation>
</mods:name>
<mods:name id="B94111F27955F3D7D375D9DE0BD7FB3D" type="personal">
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</mods:role>
<mods:namePart id="4BDFC2092112F9579CD554D549A70008">Digiani, María Celina</mods:namePart>
<mods:affiliation id="F55313454416322A049CF27B5439FB7E">CONICET. División Zoología Invertebrados, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Pase del Bosque, Buenos Aires, Argentina.</mods:affiliation>
</mods:name>
<mods:name id="CE8AD62DF8F9250DC2D7B23BF948C888" type="personal">
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</mods:role>
<mods:namePart id="961C2A944E17286AA5B7D7AC2BEEB10D">Macswiney, M. Cristina</mods:namePart>
<mods:affiliation id="624129BD5B9546663F46FD98245A5EE4">Centro de Investigaciones Tropicales, Universidad Veracruzana, Xalapa, Veracruz, México.</mods:affiliation>
</mods:name>
<mods:name id="14E206E4402E5949F1E698759710C87E" type="personal">
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</mods:role>
<mods:namePart id="2A75516CA8F508F75AF10195A8084261">Ortega, Jorge</mods:namePart>
<mods:affiliation id="C39DF5A11C6323D6EFD425C10C11D748">Escuela Nacional de Ciencias Biológicas, Instituto Politécnico Nacional, Ciudad de México, México.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="04C4C184D47F993FA2E7548D6DB0A097">Ospina-Garcés, Sandra M.</mods:namePart>
<mods:affiliation id="64BCD8C25B373238D117E4083437EBC9">Centro de Investigaciones Tropicales, Universidad Veracruzana, Xalapa, Veracruz, México. &amp; Centro de Investigación en Biodiversidad y Conservación, Universidad Autónoma del Estado de Morelos, Cuernavaca, Morelos, México.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="5D12C0F742FB368A8DB7D320F1B367F9">García-Prieto, Luis</mods:namePart>
<mods:affiliation id="CFDCD419BFBBC3807AF8B7E671A13C16">Instituto de Biología, Universidad Nacional Autónoma de México, Ciudad Universitaria, Ciudad de México C. P. 04510, México.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="FD731855706625561DB2ED19FFB97EDA">Canché-Pool, Elsy B.</mods:namePart>
<mods:affiliation id="2874DB0E9C208BC89AC3B118CC100189">Centro de Investigaciones Regionales Dr. Hideyo Noguchi, Universidad Autónoma de Yucatán, Avenida Itzáes, Mérida 97000, Yucatán, México.</mods:affiliation>
</mods:name>
<mods:name id="38A44920C6B29300AD101E1054F50F55" type="personal">
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<mods:namePart id="108284AD25215240016FE81F05F1C94C">Robles, María Del Rosario</mods:namePart>
<mods:affiliation id="F59932271843423DE884E7B44B5314A7">Centro de Estudios Parasitológicos y de Vectores (CEPAVE), Universidad Nacional de La Plata, La Plata, Buenos Aires, Argentina.</mods:affiliation>
</mods:name>
<mods:name id="0DED040D189DEF960C725CBBA4461546" type="personal">
<mods:role id="8DE330F14299B7B7C9D7EA182D15D259">
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</mods:role>
<mods:namePart id="7401EC6B936485985771FC805AC79D6B">Ávila-López, Mariana B.</mods:namePart>
<mods:affiliation id="AB0B13999811313C28C9210F740C2E47">Comisión Intersecretarial de Bioseguridad de los Organismos Genéticamente Modificados (CIBIOGEM-CONAHCYT), Ciudad de México, México.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="7B1D6A7A9C4AD0D237B6CDAF9AF3C080">Chan-Casanova, Anyela Jackelin</mods:namePart>
<mods:affiliation id="E7EDF742C7307122763BF72F439C250D">Facultad de Medicina Veterinaria y Zootecnia, Universidad Autónoma de Yucatán, Mérida Yucatán, México.</mods:affiliation>
</mods:name>
<mods:name id="05962EEA307FF80C126A580EF771FF34" type="personal">
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</mods:role>
<mods:namePart id="D46F063E2B5C9245CBAF9D33C4855E88">Panti-May, Jesús Alonso</mods:namePart>
<mods:affiliation id="F235C7075E25866962FA37C7892278EC">Centro de Investigaciones Regionales Dr. Hideyo Noguchi, Universidad Autónoma de Yucatán, Avenida Itzáes, Mérida 97000, Yucatán, México.</mods:affiliation>
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<mods:date id="788BC22BFA434B53D7C921067B0363F8">2025</mods:date>
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<mods:number id="B2E01956C523C8A9EA75C1A96B74E7EC">5566</mods:number>
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<mods:url id="0954FD9B3FCE037DC0A7D6225E5DEDFA">https://doi.org/10.11646/zootaxa.5566.2.5</mods:url>
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<mods:identifier id="59A0D344DE0AA38101ED3F7EAD0FA3AF" type="DOI">10.11646/zootaxa.5566.2.5</mods:identifier>
<mods:identifier id="10C5C62086BF315C323BA478A1F18DFF" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="03ED5E372C2CFF95A8FEA8A3911AFD2C" LSID="urn:lsid:plazi:treatment:03ED5E372C2CFF95A8FEA8A3911AFD2C" httpUri="http://treatment.plazi.org/id/03ED5E372C2CFF95A8FEA8A3911AFD2C" lastPageNumber="334" pageId="5" pageNumber="334">
<subSubSection id="C35EBCAA2C2CFF95A8FEA8A3902FFF7C" pageId="5" pageNumber="334" type="nomenclature">
<paragraph id="8BFBEF212C2CFF95A8FEA8A390D1FF38" blockId="5.[151,759,151,178]" box="[151,759,151,178]" pageId="5" pageNumber="334">
<heading id="D0B3584D2C2CFF95A8FEA8A390D1FF38" bold="true" box="[151,759,151,178]" fontSize="11" level="1" pageId="5" pageNumber="334" reason="1">
<taxonomicName id="4C4494A22C2CFF95A8FEA8A390D1FF38" authority="Zdzitowiecki &amp; Rutkowska" authorityName="Zdzitowiecki &amp; Rutkowska" box="[151,759,151,178]" class="Cestoda" family="Hymenolepididae" genus="Vampirolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="5" pageNumber="334" phylum="Platyhelminthes" rank="species" species="macroti">
<emphasis id="B93033332C2CFF95A8FEA8A390D1FF38" bold="true" box="[151,759,151,178]" pageId="5" pageNumber="334">
<emphasis id="B93033332C2CFF95A8FEA8A393BCFF38" bold="true" box="[151,410,151,178]" italics="true" pageId="5" pageNumber="334">Vampirolepis macroti</emphasis>
Zdzitowiecki &amp; Rutkowska
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFBEF212C2CFF95A8FEA8EF902FFF7C" blockId="5.[151,1437,219,678]" box="[151,521,219,245]" pageId="5" pageNumber="334">
<emphasis id="B93033332C2CFF95A8FEA8EF937DFF7C" bold="true" box="[151,347,219,245]" pageId="5" pageNumber="334">Site of infection:</emphasis>
Small intestine
</paragraph>
</subSubSection>
<subSubSection id="C35EBCAA2C2CFF95A8AEA8CB90BAFE20" pageId="5" pageNumber="334" type="biology_ecology">
<paragraph id="8BFBEF212C2CFF95A8AEA8CB902DFE90" blockId="5.[151,1437,219,678]" box="[199,523,255,281]" pageId="5" pageNumber="334">
<emphasis id="B93033332C2CFF95A8AEA8CB932EFE90" bold="true" box="[199,264,255,281]" pageId="5" pageNumber="334">Host:</emphasis>
<taxonomicName id="4C4494A22C2CFF95A979A934902DFE90" box="[272,523,255,281]" class="Mammalia" family="Vespertilionidae" genus="Bauerus" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="334" phylum="Chordata" rank="species" species="dubiaquercus">
<emphasis id="B93033332C2CFF95A979A934902DFE90" box="[272,523,255,281]" italics="true" pageId="5" pageNumber="334">Bauerus dubiaquercus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BFBEF212C2CFF95A8AEA9179768FEB7" blockId="5.[151,1437,219,678]" box="[199,1358,291,318]" pageId="5" pageNumber="334">
<emphasis id="B93033332C2CFF95A8AEA9179315FEB4" bold="true" box="[199,307,291,317]" pageId="5" pageNumber="334">Locality:</emphasis>
<materialsCitation id="3B2CE57C2C2CFF95A950A9179768FEB7" box="[313,1358,291,318]" collectionCode="MASL" latitude="18.53161" location="Tuxtlas" longLatPrecision="1" longitude="-95.133804" pageId="5" pageNumber="334" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
Ejido Adolfo Ruiz Cortines (
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,
<geoCoordinate id="EE7089E62C2CFF95AB59A91791FAFEB7" box="[816,988,291,318]" degrees="95" direction="west" minutes="08" orientation="longitude" pageId="5" pageNumber="334" precision="1" seconds="01.7" value="-95.133804">95°0801.7” W</geoCoordinate>
, 1061
<collectionCode id="ED5577E42C2CFF95AC4CA9179651FEB4" box="[1061,1143,291,317]" pageId="5" pageNumber="334">MASL</collectionCode>
),
<location id="8E9BB9FA2C2CFF95ACE3A91796C6FEB4" LSID="urn:lsid:plazi:treatment:03ED5E372C2CFF95A8FEA8A3911AFD2C:8E9BB9FA2C2CFF95ACE3A91796C6FEB4" box="[1162,1248,291,317]" latitude="18.53161" longLatPrecision="1" longitude="-95.133804" name="Tuxtlas" pageId="5" pageNumber="334" stateProvince="Veracruz de Ignacio de la Llave">Tuxtlas</location>
,
<collectingRegion id="498021C32C2CFF95AC82A9109768FEB7" box="[1259,1358,292,318]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="5" pageNumber="334">Veracruz</collectingRegion>
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</paragraph>
<paragraph id="8BFBEF212C2CFF95A8AEA9739097FEE8" blockId="5.[151,1437,219,678]" box="[199,689,327,354]" pageId="5" pageNumber="334">
<emphasis id="B93033332C2CFF95A8AEA97393D3FEE8" bold="true" box="[199,501,327,353]" pageId="5" pageNumber="334">Prevalence and intensity:</emphasis>
50% (1/2) and 4
</paragraph>
<paragraph id="8BFBEF212C2CFF95A8AEA95F907AFE0F" blockId="5.[151,1437,219,678]" box="[199,604,363,390]" pageId="5" pageNumber="334">
<emphasis id="B93033332C2CFF95A8AEA95F939AFE0C" bold="true" box="[199,444,363,389]" pageId="5" pageNumber="334">Specimen deposited:</emphasis>
CNHE 12135
</paragraph>
<paragraph id="8BFBEF212C2CFF95A8AEA9BB90BAFE20" blockId="5.[151,1437,219,678]" box="[199,668,399,425]" pageId="5" pageNumber="334">
<emphasis id="B93033332C2CFF95A8AEA9BB903BFE20" bold="true" box="[199,541,399,425]" pageId="5" pageNumber="334">GenBank accession number:</emphasis>
PQ476180
</paragraph>
</subSubSection>
<subSubSection id="C35EBCAA2C2CFF95A8AEA987911AFD2C" pageId="5" pageNumber="334" type="discussion">
<paragraph id="8BFBEF212C2CFF95A8AEA9879093FDD4" blockId="5.[151,1437,219,678]" pageId="5" pageNumber="334">
<emphasis id="B93033332C2CFF95A8AEA9879374FE44" bold="true" box="[199,338,435,461]" pageId="5" pageNumber="334">Comments:</emphasis>
Based on two incomplete specimens. Scolex
<quantity id="4CBC42C42C2CFF95AB1DA98791E6FE44" box="[884,960,435,461]" metricMagnitude="0" metricUnit="m" metricValue="6.603999999999999" pageId="5" pageNumber="334" unit="in" value="260.0">260 in</quantity>
diameter. Suckers unarmed,
<quantity id="4CBC42C42C2CFF95AD71A98797BAFE44" box="[1304,1436,435,461]" metricMagnitude="0" metricUnit="m" metricValue="2.6416" metricValueMax="2.7432" metricValueMin="2.54" pageId="5" pageNumber="334" unit="in" value="104.0" valueMax="108.0" valueMin="100.0">100108 in</quantity>
diameter. Rostellum broad,
<quantity id="4CBC42C42C2CFF95A9BDA9E3901FFE78" box="[468,569,471,498]" metricMagnitude="0" metricUnit="m" metricValue="1.8796" metricValueMax="1.905" metricValueMin="1.8542" pageId="5" pageNumber="334" unit="in" value="74.0" valueMax="75.0" valueMin="73.0">7375 in</quantity>
diameter, armed with 2829 hooks in single row (
<figureCitation id="137FF3A42C2CFF95ACE9A9E396DFFE78" box="[1152,1273,471,497]" captionStart="FIGURE 2" captionStartId="5.[151,250,1874,1898]" captionTargetBox="[168,1419,717,1834]" captionTargetId="figure-165@5.[151,1436,701,1851]" captionTargetPageId="5" captionText="FIGURE 2. Helminths of bats from Nayarit and Veracruz, Mexico. A) Adult of Anenterotrema cf. hastati, ventral view. B) Adult of Urotrema scabridum, ventral view. C) Scolex of Vampirolepis macroti, apical view. D) Scolex of Vampirolepis sp. (1), apical view. E) Scolex of Vampirolepis sp. (2), lateral view. F) Male posterior extremity of Capillariidae gen. sp., ventral view. G) Esophagus-intestine junction and vulva of female Capillariidae gen. sp., lateral view. Scale bars: A, C, D, E = 100 μm; B = 1000 μm; F = 10 μm; G = 65 μm." pageId="5" pageNumber="334">Figure 2C</figureCitation>
). Hook length 2632. Although no mature proglottids were found, the number and length of hooks observed in the scolex of our specimens conformed to the original description of
<taxonomicName id="4C4494A22C2CFF95AA8CAA14917AFDB0" box="[741,860,544,570]" class="Cestoda" family="Hymenolepididae" genus="Vampirolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="5" pageNumber="334" phylum="Platyhelminthes" rank="species" species="macroti">
<emphasis id="B93033332C2CFF95AA8CAA1490D0FDB0" box="[741,758,544,569]" italics="true" pageId="5" pageNumber="334">V</emphasis>
.
<emphasis id="B93033332C2CFF95AB6FAA14917AFDB0" box="[774,860,544,569]" italics="true" pageId="5" pageNumber="334">macroti</emphasis>
</taxonomicName>
from
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<emphasis id="B93033332C2CFF95ABCDAA14919DFDB0" box="[932,955,544,569]" italics="true" pageId="5" pageNumber="334">M</emphasis>
.
<emphasis id="B93033332C2CFF95ABA2AA2B9669FDB0" box="[971,1103,543,569]" italics="true" pageId="5" pageNumber="334">waterhousii</emphasis>
</taxonomicName>
in
<collectingCountry id="F353AFB12C2CFF95AC1FAA2B9697FDB0" box="[1142,1201,543,569]" name="Cuba" pageId="5" pageNumber="334">Cuba</collectingCountry>
(
<emphasis id="B93033332C2CFF95ACAAAA1496FCFDB0" box="[1219,1242,544,569]" italics="true" pageId="5" pageNumber="334">vs</emphasis>
. 2934 hooks of 2830 long) (
<bibRefCitation id="EFD592D02C2CFF95A947AA77908EFDD4" author="Zdzitowiecki, K. &amp; Rutkowska, M. A." box="[302,680,579,605]" pageId="5" pageNumber="334" pagination="187 - 200" refId="ref12496" refString="Zdzitowiecki, K. &amp; Rutkowska, M. A. (1980) The helminth fauna of bats (Chiroptera) from Cuba. II. A review of cestodes with description of four new species and a key to Hymenolepididae of American bats. Acta Parasitologica Polonica, 15, 187 - 200." type="journal article" year="1980">Zdzitowiecki &amp; Rutkowska 1980</bibRefCitation>
).
</paragraph>
<paragraph id="8BFBEF212C2CFF95A8AEAA53911AFD2C" blockId="5.[151,1437,219,678]" pageId="5" pageNumber="334">
<taxonomicName id="4C4494A22C2CFF95A8AEAA53939EFD08" box="[199,440,615,641]" class="Cestoda" family="Hymenolepididae" genus="Vampirolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="5" pageNumber="334" phylum="Platyhelminthes" rank="species" species="macroti">
<emphasis id="B93033332C2CFF95A8AEAA53939EFD08" box="[199,440,615,641]" italics="true" pageId="5" pageNumber="334">Vampirolepis macroti</emphasis>
</taxonomicName>
has been reported from
<taxonomicName id="4C4494A22C2CFF95AAA7AA5C915EFD08" authorityName="Gray" authorityYear="1843" box="[718,888,615,642]" class="Mammalia" family="Phyllostomidae" genus="Macrotus" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="334" phylum="Chordata" rank="species" species="waterhousii">
<emphasis id="B93033332C2CFF95AAA7AA5C90C3FD08" box="[718,741,616,641]" italics="true" pageId="5" pageNumber="334">M</emphasis>
.
<emphasis id="B93033332C2CFF95AA9DAA53915EFD08" box="[756,888,615,641]" italics="true" pageId="5" pageNumber="334">waterhousii</emphasis>
</taxonomicName>
in
<collectingRegion id="498021C32C2CFF95ABF4AA53963FFD08" box="[925,1049,615,641]" country="Mexico" name="Michoacan de Ocampo" pageId="5" pageNumber="334">Michoacán</collectingRegion>
(
<bibRefCitation id="EFD592D02C2CFF95AC40AA5397B7FD08" author="Luviano-Hernandez, J. &amp; Tafolla-Venegas, D. &amp; Melendez-Herrera, E. &amp; Fuentes-Farias, A. L." box="[1065,1425,615,641]" pageId="5" pageNumber="334" pagination="1315 - 1321" refId="ref10590" refString="Luviano-Hernandez, J., Tafolla-Venegas, D., Melendez-Herrera, E. &amp; Fuentes-Farias, A. L. (2018) Helmintos del murcielago Macrotus waterhousii (Chiroptera: Phyllostomidae) en dos refugios de selva baja caducifolia en el municipio de Huetamo, Michoacan, Mexico. Revista Mexicana de Biodiversidad, 89, 1315 - 1321." type="journal article" year="2018">
Luviano-Hernández
<emphasis id="B93033332C2CFF95AD7AAA5C976AFD08" box="[1299,1356,615,641]" italics="true" pageId="5" pageNumber="334">et al.</emphasis>
2018
</bibRefCitation>
). This is the second record of
<taxonomicName id="4C4494A22C2CFF95A9BDAAB8906CFD2C" box="[468,586,652,678]" class="Cestoda" family="Hymenolepididae" genus="Vampirolepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclophyllidea" pageId="5" pageNumber="334" phylum="Platyhelminthes" rank="species" species="macroti">
<emphasis id="B93033332C2CFF95A9BDAAB893C3FD2C" box="[468,485,652,677]" italics="true" pageId="5" pageNumber="334">V</emphasis>
.
<emphasis id="B93033332C2CFF95A99DAAB8906CFD2C" box="[500,586,652,677]" italics="true" pageId="5" pageNumber="334">macroti</emphasis>
</taxonomicName>
in bats from
<collectingCountry id="F353AFB12C2CFF95AAB6AABF911EFD2C" box="[735,824,651,677]" name="Mexico" pageId="5" pageNumber="334">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,300 @@
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<mods:title id="8B5CB58450BCF95751AEFF13217596D5">Intestinal helminths of bats in the States of Nayarit and Veracruz, Mexico, with redescription of Bidigiticauda vivipara Chitwood</mods:title>
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<mods:namePart id="474B959DC17A48548BC9A5CD82205B28">Moguel-Chin, Wilson I.</mods:namePart>
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<mods:namePart id="962D5D5C7BFDF0FD1870B648EACCA601">Digiani, María Celina</mods:namePart>
<mods:affiliation id="CE29756B67AD3712E097FA452826D206">CONICET. División Zoología Invertebrados, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Pase del Bosque, Buenos Aires, Argentina.</mods:affiliation>
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<mods:namePart id="CAB39F9BB2FD14F9DF24F7BD1CBFF4A1">Macswiney, M. Cristina</mods:namePart>
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<mods:namePart id="AABB60F274515B9C71BE08CECDE85E2D">Ortega, Jorge</mods:namePart>
<mods:affiliation id="5E1F88AFF6FD87EF969BF453C12EB25A">Escuela Nacional de Ciencias Biológicas, Instituto Politécnico Nacional, Ciudad de México, México.</mods:affiliation>
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<mods:affiliation id="F4519AC231F0255CFE18E64BD88737F9">Centro de Investigaciones Tropicales, Universidad Veracruzana, Xalapa, Veracruz, México. &amp; Centro de Investigación en Biodiversidad y Conservación, Universidad Autónoma del Estado de Morelos, Cuernavaca, Morelos, México.</mods:affiliation>
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<mods:affiliation id="F8466D0533D3AEEECADCBA8B5F402125">Instituto de Biología, Universidad Nacional Autónoma de México, Ciudad Universitaria, Ciudad de México C. P. 04510, México.</mods:affiliation>
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<mods:namePart id="337C2F192D6F951FCB77B07897AE1B5C">Canché-Pool, Elsy B.</mods:namePart>
<mods:affiliation id="888A14D760A64DF37AB2DE8E1EB02A0A">Centro de Investigaciones Regionales Dr. Hideyo Noguchi, Universidad Autónoma de Yucatán, Avenida Itzáes, Mérida 97000, Yucatán, México.</mods:affiliation>
</mods:name>
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<mods:affiliation id="67901874E170CE5E7C07AF1577E2ECEE">Centro de Estudios Parasitológicos y de Vectores (CEPAVE), Universidad Nacional de La Plata, La Plata, Buenos Aires, Argentina.</mods:affiliation>
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<mods:namePart id="CC3FD9B4671AFBCF7F7FD122BF6C9D4D">Ávila-López, Mariana B.</mods:namePart>
<mods:affiliation id="9A42793DA23F7C374C56FF3A937DA694">Comisión Intersecretarial de Bioseguridad de los Organismos Genéticamente Modificados (CIBIOGEM-CONAHCYT), Ciudad de México, México.</mods:affiliation>
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<mods:namePart id="BE32DD789B94BAD826174E5CE1B216D1">Chan-Casanova, Anyela Jackelin</mods:namePart>
<mods:affiliation id="6C00839E702FD604F5B68807B1FDB80A">Facultad de Medicina Veterinaria y Zootecnia, Universidad Autónoma de Yucatán, Mérida Yucatán, México.</mods:affiliation>
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<mods:namePart id="4F042ECCF3EE15EFB8BB7C35591251CE">Panti-May, Jesús Alonso</mods:namePart>
<mods:affiliation id="46FEDA21D970A7961F072E6423E4FE8B">Centro de Investigaciones Regionales Dr. Hideyo Noguchi, Universidad Autónoma de Yucatán, Avenida Itzáes, Mérida 97000, Yucatán, México.</mods:affiliation>
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<treatment id="03ED5E372C2DFF94A8FEAB9B9163F9D8" LSID="urn:lsid:plazi:treatment:03ED5E372C2DFF94A8FEAB9B9163F9D8" httpUri="http://treatment.plazi.org/id/03ED5E372C2DFF94A8FEAB9B9163F9D8" lastPageNumber="333" pageId="4" pageNumber="333">
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<paragraph id="8BFBEF212C2DFF94A8FEAB9B93CCFC40" blockId="4.[151,490,942,969]" box="[151,490,942,969]" pageId="4" pageNumber="333">
<heading id="D0B3584D2C2DFF94A8FEAB9B93CCFC40" bold="true" box="[151,490,942,969]" fontSize="11" level="1" pageId="4" pageNumber="333" reason="1">
<taxonomicName id="4C4494A22C2DFF94A8FEAB9B93CCFC40" ID-CoL="84DZC" authority="Braun" authorityName="Braun" authorityYear="1900" box="[151,490,942,969]" class="Trematoda" family="Pleurogenidae" genus="Urotrema" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="scabridum">
<emphasis id="B93033332C2DFF94A8FEAB9B93CCFC40" bold="true" box="[151,490,942,969]" pageId="4" pageNumber="333">
<emphasis id="B93033332C2DFF94A8FEAB9B93B5FC40" bold="true" box="[151,403,942,969]" italics="true" pageId="4" pageNumber="333">Urotrema scabridum</emphasis>
Braun
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</paragraph>
<paragraph id="8BFBEF212C2DFF94A8FEABC2902FFB98" blockId="4.[151,1437,1014,1617]" box="[151,521,1014,1041]" pageId="4" pageNumber="333">
<emphasis id="B93033332C2DFF94A8FEABC2937DFB99" bold="true" box="[151,347,1014,1040]" pageId="4" pageNumber="333">Site of infection:</emphasis>
Small intestine
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</subSubSection>
<subSubSection id="C35EBCAA2C2DFF94A8AEAC2F907AFB28" pageId="4" pageNumber="333" type="biology_ecology">
<paragraph id="8BFBEF212C2DFF94A8AEAC2F93C3FBBC" blockId="4.[151,1437,1014,1617]" box="[199,485,1051,1077]" pageId="4" pageNumber="333">
<emphasis id="B93033332C2DFF94A8AEAC2F932EFBBC" bold="true" box="[199,264,1051,1077]" pageId="4" pageNumber="333">Host:</emphasis>
<taxonomicName id="4C4494A22C2DFF94A979AC2F93C3FBBC" box="[272,485,1051,1077]" class="Mammalia" family="Vespertilionidae" genus="Rhogeessa" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="parvula">
<emphasis id="B93033332C2DFF94A979AC2F93C3FBBC" box="[272,485,1051,1077]" italics="true" pageId="4" pageNumber="333">Rhogeessa parvula</emphasis>
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<paragraph id="8BFBEF212C2DFF94A8AEAC0A96AFFBD0" blockId="4.[151,1437,1014,1617]" box="[199,1161,1086,1113]" pageId="4" pageNumber="333">
<emphasis id="B93033332C2DFF94A8AEAC0A9315FBD1" bold="true" box="[199,307,1086,1112]" pageId="4" pageNumber="333">Locality:</emphasis>
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Cleofas (
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,
<geoCoordinate id="EE7089E62C2DFF94AA3AAC0B912AFBD0" box="[595,780,1087,1113]" degrees="106" direction="west" minutes="14" orientation="longitude" pageId="4" pageNumber="333" precision="1" seconds="41.6" value="-106.24489">106°1441.6” W</geoCoordinate>
, 98
<collectionCode id="ED5577E42C2DFF94AB53AC0B91AAFBD0" box="[826,908,1087,1113]" pageId="4" pageNumber="333">MASL</collectionCode>
),
<location id="8E9BB9FA2C2DFF94ABF6AC0B960CFBD0" LSID="urn:lsid:plazi:treatment:03ED5E372C2DFF94A8FEAB9B9163F9D8:8E9BB9FA2C2DFF94ABF6AC0B960CFBD0" box="[927,1066,1087,1113]" latitude="21.32203" longLatPrecision="1" longitude="-106.24489" name="Islas Marias" pageId="4" pageNumber="333" stateProvince="Nayarit">Islas Marías</location>
,
<collectingRegion id="498021C32C2DFF94AC5CAC0B96AFFBD0" box="[1077,1161,1087,1113]" country="Mexico" name="Nayarit" pageId="4" pageNumber="333">Nayarit</collectingRegion>
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</paragraph>
<paragraph id="8BFBEF212C2DFF94A8AEAC5790E0FBF4" blockId="4.[151,1437,1014,1617]" box="[199,710,1122,1149]" pageId="4" pageNumber="333">
<emphasis id="B93033332C2DFF94A8AEAC5793D3FBF5" bold="true" box="[199,501,1122,1149]" pageId="4" pageNumber="333">Prevalence and intensity:</emphasis>
33.3% (1/3) and 1
</paragraph>
<paragraph id="8BFBEF212C2DFF94A8AEACB2907AFB28" blockId="4.[151,1437,1014,1617]" box="[199,604,1158,1185]" pageId="4" pageNumber="333">
<emphasis id="B93033332C2DFF94A8AEACB2939AFB29" bold="true" box="[199,444,1158,1184]" pageId="4" pageNumber="333">Specimen deposited:</emphasis>
CNHE 12133
</paragraph>
</subSubSection>
<subSubSection id="C35EBCAA2C2DFF94A8AEAC9E9163F9D8" pageId="4" pageNumber="333" type="discussion">
<paragraph id="8BFBEF212C2DFF94A8AEAC9E9387FAB8" blockId="4.[151,1437,1014,1617]" pageId="4" pageNumber="333">
<emphasis id="B93033332C2DFF94A8AEAC9E9374FB4D" bold="true" box="[199,338,1194,1220]" pageId="4" pageNumber="333">Comments:</emphasis>
Based on a single specimen. Body elongate (3360 long and 485 wide), covered with fine spines (
<figureCitation id="137FF3A42C2DFF94A8F6ACFB9332FB60" box="[159,276,1231,1257]" captionStart="FIGURE 2" captionStartId="5.[151,250,1874,1898]" captionTargetBox="[168,1419,717,1834]" captionTargetId="figure-165@5.[151,1436,701,1851]" captionTargetPageId="5" captionText="FIGURE 2. Helminths of bats from Nayarit and Veracruz, Mexico. A) Adult of Anenterotrema cf. hastati, ventral view. B) Adult of Urotrema scabridum, ventral view. C) Scolex of Vampirolepis macroti, apical view. D) Scolex of Vampirolepis sp. (1), apical view. E) Scolex of Vampirolepis sp. (2), lateral view. F) Male posterior extremity of Capillariidae gen. sp., ventral view. G) Esophagus-intestine junction and vulva of female Capillariidae gen. sp., lateral view. Scale bars: A, C, D, E = 100 μm; B = 1000 μm; F = 10 μm; G = 65 μm." pageId="4" pageNumber="333">Figure 2B</figureCitation>
). Oral sucker 145 long by 100 wide, ventral sucker 200 long by 140 wide. Pharynx 60 long by 50 wide. Ovary 250 long by 190 wide. Anterior testis 260 long by 270 wide, posterior testis 250 long by 225 wide. Eggs 1520 long by 10 wide.
</paragraph>
<paragraph id="8BFBEF212C2DFF94A8AEAD0F9163F9D8" blockId="4.[151,1437,1014,1617]" pageId="4" pageNumber="333">
Morphologically, the specimen fits the original description of
<taxonomicName id="4C4494A22C2DFF94ABE0AD0F96E2FADC" authority="(Braun 1900)" baseAuthorityName="Braun" baseAuthorityYear="1900" box="[905,1220,1339,1365]" class="Trematoda" family="Pleurogenidae" genus="Urotrema" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="scabridum">
<emphasis id="B93033332C2DFF94ABE0AD0F9605FADC" box="[905,1059,1339,1365]" italics="true" pageId="4" pageNumber="333">U. scabridum</emphasis>
(
<bibRefCitation id="EFD592D02C2DFF94AC5AAD0F969DFADC" author="Braun, M." box="[1075,1211,1339,1365]" pageId="4" pageNumber="333" pagination="217 - 236" refId="ref8493" refString="Braun, M. (1900) Trematoden der chiropterans. Annalen des K. K. Natur- historischen Hofmuseums, 15, 217 - 236." type="journal article" year="1900">Braun 1900</bibRefCitation>
)
</taxonomicName>
and recent records (
<bibRefCitation id="EFD592D02C2DFF94A8F6AD6B93CCFAF0" author="Martinez-Salazar, E. A. &amp; Medina-Rodriguez, A. J. &amp; Rosas-Valdez, R. &amp; Del Real-Monroy, M. &amp; Falcon-Ordaz, J." box="[159,490,1375,1401]" pageId="4" pageNumber="333" pagination="181 - 192" refId="ref10783" refString="Martinez-Salazar, E. A., Medina-Rodriguez, A. J., Rosas-Valdez, R., Del Real-Monroy, M. &amp; Falcon-Ordaz, J. (2020) First record of Urotrema scabridum (Platyhelminthes), and new redords of helminths of Tadarida brasiliensis from Mexican Plateau. Therya, 11, 181 - 192. https: // doi. org / 10.12933 / therya- 20 - 788" type="journal article" year="2020">
Martínez-Salazar
<emphasis id="B93033332C2DFF94A903AD6B9382FAF0" box="[362,420,1375,1401]" italics="true" pageId="4" pageNumber="333">et al.</emphasis>
2020
</bibRefCitation>
).
<taxonomicName id="4C4494A22C2DFF94A997AD6B90CFFAF0" baseAuthorityName="Braun" baseAuthorityYear="1900" box="[510,745,1375,1401]" class="Trematoda" family="Pleurogenidae" genus="Urotrema" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="scabridum">
<emphasis id="B93033332C2DFF94A997AD6B90CFFAF0" box="[510,745,1375,1401]" italics="true" pageId="4" pageNumber="333">Urotrema scabridum</emphasis>
</taxonomicName>
has been recorded in several bat species in
<collectingCountry id="F353AFB12C2DFF94AC8AAD6B971DFAF0" box="[1251,1339,1375,1401]" name="Mexico" pageId="4" pageNumber="333">Mexico</collectingCountry>
, such as
<taxonomicName id="4C4494A22C2DFF94A8FEADB790BFFA14" authority="(Geoffroy Saint-Hilaire)" baseAuthorityName="Geoffroy Saint-Hilaire" box="[151,665,1411,1437]" class="Mammalia" family="Molossidae" genus="Tadarida" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="B93033332C2DFF94A8FEADB793A2FA14" box="[151,388,1411,1437]" italics="true" pageId="4" pageNumber="333">Tadarida brasiliensis</emphasis>
(Geoffroy Saint-Hilaire)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94AACFADB7960DFA14" authority="Smith" authorityName="Smith" authorityYear="1972" box="[678,1067,1411,1437]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="mesoamericanus">
<emphasis id="B93033332C2DFF94AACFADB791C4FA15" box="[678,994,1411,1436]" italics="true" pageId="4" pageNumber="333">Pteronotus mesoamericanus</emphasis>
Smith
</taxonomicName>
(syn.
<taxonomicName id="4C4494A22C2DFF94AC07ADB797B6FA14" authority="Gray" authorityName="Gray" box="[1134,1424,1411,1437]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="parnellii">
<emphasis id="B93033332C2DFF94AC07ADB79768FA14" box="[1134,1358,1411,1437]" italics="true" pageId="4" pageNumber="333">Pteronotus parnellii</emphasis>
Gray
</taxonomicName>
),
<taxonomicName id="4C4494A22C2DFF94A8FEAD939322FA48" authorityName="Peters" authorityYear="1867" box="[151,260,1447,1473]" class="Mammalia" family="Emballonuridae" genus="Balantiopteryx" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="plicata">
<emphasis id="B93033332C2DFF94A8FEAD93928EFA49" box="[151,168,1447,1472]" italics="true" pageId="4" pageNumber="333">B</emphasis>
.
<emphasis id="B93033332C2DFF94A8DFAD939322FA48" box="[182,260,1447,1473]" italics="true" pageId="4" pageNumber="333">plicata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94A978AD93906EFA48" authority="(Thomas)" baseAuthorityName="Thomas" baseAuthorityYear="1892" box="[273,584,1447,1473]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="fulvus">
<emphasis id="B93033332C2DFF94A978AD9393F2FA48" box="[273,468,1447,1473]" italics="true" pageId="4" pageNumber="333">Pteronotus fulvus</emphasis>
(Thomas)
</taxonomicName>
(syn.
<taxonomicName id="4C4494A22C2DFF94AAE2AD9391ACFA48" authority="Gray" authorityName="Gray" authorityYear="1838" box="[651,906,1447,1473]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="davyi">
<emphasis id="B93033332C2DFF94AAE2AD93916EFA48" box="[651,840,1447,1473]" italics="true" pageId="4" pageNumber="333">Pteronotus davyi</emphasis>
Gray
</taxonomicName>
),
<taxonomicName id="4C4494A22C2DFF94ABF4AD9396F9FA48" authority="(Dobson)" baseAuthorityName="Dobson" baseAuthorityYear="1878" box="[925,1247,1447,1473]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="psilotis">
<emphasis id="B93033332C2DFF94ABF4AD939648FA48" box="[925,1134,1447,1473]" italics="true" pageId="4" pageNumber="333">Pteronotus psilotis</emphasis>
(Dobson)
</taxonomicName>
(syn.
<taxonomicName id="4C4494A22C2DFF94AD4BAD93935EFA6C" authority="Wagner" authorityName="Wagner" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="personatus">
<emphasis id="B93033332C2DFF94AD4BAD939334FA6D" italics="true" pageId="4" pageNumber="333">Pteronotus personatus</emphasis>
Wagner
</taxonomicName>
),
<taxonomicName id="4C4494A22C2DFF94A9E6ADFF90AAFA6C" authority="(Allen)" baseAuthorityName="Allen" box="[399,652,1483,1509]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="velifer">
<emphasis id="B93033332C2DFF94A9E6ADFF9008FA6C" box="[399,558,1483,1509]" italics="true" pageId="4" pageNumber="333">Myotis velifer</emphasis>
(Allen)
</taxonomicName>
,
<taxonomicName id="4C4494A22C2DFF94AAF5ADFF963AFA6C" authority="(Peters)" baseAuthorityName="Peters" baseAuthorityYear="1864" box="[668,1052,1483,1509]" class="Mammalia" family="Mormoopidae" genus="Mormoops" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="megalophylla">
<emphasis id="B93033332C2DFF94AAF5ADFF919FFA6C" box="[668,953,1483,1509]" italics="true" pageId="4" pageNumber="333">Mormoops megalophylla</emphasis>
(Peters)
</taxonomicName>
in
<collectingRegion id="498021C32C2DFF94AC21ADFF9683FA6C" box="[1096,1189,1483,1509]" country="Mexico" name="Morelos" pageId="4" pageNumber="333">Morelos</collectingRegion>
(see the checklist of
<bibRefCitation id="EFD592D02C2DFF94A8FEADDB9355F980" author="Jimenez, F. A. &amp; Caspeta-Mandujano, J. M. &amp; Ramirez-Chavez, S. B. &amp; Ramirez-Diaz, S. E. &amp; Juarez-Urbina, M. G. &amp; Peralta-Ramirez, J. L. &amp; Guerrero, J. A." box="[151,371,1519,1545]" pageId="4" pageNumber="333" pagination="1 - 28" refId="ref10020" refString="Jimenez, F. A., Caspeta-Mandujano, J. M., Ramirez-Chavez, S. B., Ramirez-Diaz, S. E., Juarez-Urbina, M. G., Peralta-Ramirez, J. L. &amp; Guerrero, J. A. (2017) Checklist of helminths of bats from Mexico and Central America. Journal of Parasite Biodiversity, 7, 1 - 28. https: // doi. org / 10.13014 / K 2 ZG 6 QDX" type="journal article" year="2017">
Jiménez
<emphasis id="B93033332C2DFF94A892ADDB9315F980" box="[251,307,1519,1545]" italics="true" pageId="4" pageNumber="333">et al.</emphasis>
2017
</bibRefCitation>
and references therein),
<taxonomicName id="4C4494A22C2DFF94AAE2ADDB910CF980" box="[651,810,1519,1545]" class="Mammalia" family="Molossidae" genus="Tadarida" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="B93033332C2DFF94AAE2ADDB90BDF981" box="[651,667,1519,1544]" italics="true" pageId="4" pageNumber="333">T</emphasis>
.
<emphasis id="B93033332C2DFF94AAC0ADDB910CF980" box="[681,810,1519,1545]" italics="true" pageId="4" pageNumber="333">brasiliensis</emphasis>
</taxonomicName>
in
<collectingRegion id="498021C32C2DFF94AB26ADDB9192F980" box="[847,948,1519,1545]" country="Mexico" name="Durango" pageId="4" pageNumber="333">Durango</collectingRegion>
,
<collectingCountry id="F353AFB12C2DFF94ABA9ADDB9630F980" box="[960,1046,1519,1545]" name="Mexico" pageId="4" pageNumber="333">Mexico</collectingCountry>
State,
<collectingRegion id="498021C32C2DFF94AC0AADDB96D4F980" box="[1123,1266,1519,1545]" country="Mexico" name="Nuevo Leon" pageId="4" pageNumber="333">Nuevo León</collectingRegion>
,
<collectingRegion id="498021C32C2DFF94AC97ADDB974AF980" box="[1278,1388,1519,1545]" country="Mexico" name="Zacatecas" pageId="4" pageNumber="333">Zacatecas</collectingRegion>
and
<collectingRegion id="498021C32C2DFF94A8FEAE2792C0F9A4" box="[151,230,1555,1581]" country="Mexico" name="Puebla" pageId="4" pageNumber="333">Puebla</collectingRegion>
, and
<taxonomicName id="4C4494A22C2DFF94A94AAE279060F9A4" authority="Miller" authorityName="Miller" authorityYear="1902" box="[291,582,1555,1581]" class="Mammalia" family="Natalidae" genus="Natalus" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="mexicanus">
<emphasis id="B93033332C2DFF94A94AAE2793DEF9A5" box="[291,504,1555,1581]" italics="true" pageId="4" pageNumber="333">Natalus mexicanus</emphasis>
Miller
</taxonomicName>
in
<collectingRegion id="498021C32C2DFF94AA02AE2790DFF9A4" box="[619,761,1555,1581]" country="Mexico" name="Distrito Federal" pageId="4" pageNumber="333">Mexico City</collectingRegion>
(
<bibRefCitation id="EFD592D02C2DFF94AB60AE2791CCF9A4" author="Jimenez, F. A. &amp; Caspeta-Mandujano, J. M. &amp; Ramirez-Chavez, S. B. &amp; Ramirez-Diaz, S. E. &amp; Juarez-Urbina, M. G. &amp; Peralta-Ramirez, J. L. &amp; Guerrero, J. A." box="[777,1002,1555,1581]" pageId="4" pageNumber="333" pagination="1 - 28" refId="ref10020" refString="Jimenez, F. A., Caspeta-Mandujano, J. M., Ramirez-Chavez, S. B., Ramirez-Diaz, S. E., Juarez-Urbina, M. G., Peralta-Ramirez, J. L. &amp; Guerrero, J. A. (2017) Checklist of helminths of bats from Mexico and Central America. Journal of Parasite Biodiversity, 7, 1 - 28. https: // doi. org / 10.13014 / K 2 ZG 6 QDX" type="journal article" year="2017">
Jiménez
<emphasis id="B93033332C2DFF94AB07AE279181F9A4" box="[878,935,1555,1581]" italics="true" pageId="4" pageNumber="333">et al.</emphasis>
2017
</bibRefCitation>
;
<bibRefCitation id="EFD592D02C2DFF94AB9EAE27971BF9A4" author="Martinez-Salazar, E. A. &amp; Medina-Rodriguez, A. J. &amp; Rosas-Valdez, R. &amp; Del Real-Monroy, M. &amp; Falcon-Ordaz, J." box="[1015,1341,1555,1581]" pageId="4" pageNumber="333" pagination="181 - 192" refId="ref10783" refString="Martinez-Salazar, E. A., Medina-Rodriguez, A. J., Rosas-Valdez, R., Del Real-Monroy, M. &amp; Falcon-Ordaz, J. (2020) First record of Urotrema scabridum (Platyhelminthes), and new redords of helminths of Tadarida brasiliensis from Mexican Plateau. Therya, 11, 181 - 192. https: // doi. org / 10.12933 / therya- 20 - 788" type="journal article" year="2020">
Martínez-Salazar
<emphasis id="B93033332C2DFF94ACD6AE2796DEF9A4" box="[1215,1272,1555,1581]" italics="true" pageId="4" pageNumber="333">et al.</emphasis>
2020
</bibRefCitation>
). This is the first record of
<taxonomicName id="4C4494A22C2DFF94A908AE0393DCF9D8" baseAuthorityName="Braun" baseAuthorityYear="1900" box="[353,506,1591,1617]" class="Trematoda" family="Pleurogenidae" genus="Urotrema" kingdom="Animalia" order="Plagiorchiida" pageId="4" pageNumber="333" phylum="Platyhelminthes" rank="species" species="scabridum">
<emphasis id="B93033332C2DFF94A908AE039353F9D9" box="[353,373,1591,1616]" italics="true" pageId="4" pageNumber="333">U</emphasis>
.
<emphasis id="B93033332C2DFF94A9EDAE0393DCF9D8" box="[388,506,1591,1617]" italics="true" pageId="4" pageNumber="333">scabridum</emphasis>
</taxonomicName>
in
<collectingRegion id="498021C32C2DFF94AA77AE039054F9D8" box="[542,626,1591,1617]" country="Mexico" name="Nayarit" pageId="4" pageNumber="333">Nayarit</collectingRegion>
and for
<taxonomicName id="4C4494A22C2DFF94AAB9AE039118F9D9" authorityName="Merriam" authorityYear="1898" box="[720,830,1591,1617]" class="Mammalia" family="Phyllostomidae" genus="Glossophaga" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="333" phylum="Chordata" rank="species" species="mutica">
<emphasis id="B93033332C2DFF94AAB9AE0390C2F9D8" box="[720,740,1591,1617]" italics="true" pageId="4" pageNumber="333">G</emphasis>
.
<emphasis id="B93033332C2DFF94AA9BAE039118F9D9" box="[754,830,1591,1616]" italics="true" pageId="4" pageNumber="333">mutica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,242 @@
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<mods:title id="FA0C06A7AB6FA9BF391CA64DFAA84FB5">Contributions to the genus Bolbochromus Boucomont, 1909 (Coleoptera: Geotrupidae: Bolboceratinae) from China, with the description of a new species</mods:title>
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<taxonomicName id="47E34D19F741191D0EA5F9407215F971" ID-CoL="7SPCP" authority="Krikken &amp; Li, 2013" authorityName="Krikken &amp; Li" authorityYear="2013" box="[151,891,1746,1773]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="389" phylum="Arthropoda" rank="species" species="sinensis" subGenus="Bolbochromus">
<emphasis id="B297EA88F741191D0EA5F9407215F971" bold="true" box="[151,891,1746,1773]" pageId="8" pageNumber="389">
<emphasis id="B297EA88F741191D0EA5F9407027F971" bold="true" box="[151,329,1746,1773]" italics="true" pageId="8" pageNumber="389">Bolbochromus</emphasis>
(
<emphasis id="B297EA88F741191D0F68F9407362F971" bold="true" box="[346,524,1746,1773]" italics="true" pageId="8" pageNumber="389">Bolbochromus</emphasis>
)
<emphasis id="B297EA88F741191D0C2FF9417313F971" bold="true" box="[541,637,1747,1773]" italics="true" pageId="8" pageNumber="389">sinensis</emphasis>
<bibRefCitation id="E4724B6BF741191D0CB6F9407215F971" author="Krikken, J. &amp; Li, C. - L." box="[644,891,1746,1773]" pageId="8" pageNumber="389" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Krikken &amp; Li, 2013</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F741191D0EA5F96570ADF88E" box="[151,451,1783,1810]" pageId="8" pageNumber="389" type="description">
<paragraph id="805C369AF741191D0EA5F96570ADF88E" blockId="8.[151,891,1746,1810]" box="[151,451,1783,1810]" pageId="8" pageNumber="389">
(
<figureCitation id="18D82A1FF741191D0EADF9657048F88D" box="[159,294,1783,1809]" captionStart="FIGURE 3" captionStartId="4.[151,250,1732,1756]" captionTargetBox="[154,1416,201,1690]" captionTargetId="figure-23@4.[151,1435,181,1708]" captionTargetPageId="4" captionText="FIGURE 3. Habitus of Bolbochromus sinensis Krikken &amp; Li, 2013. AB, male (Nanchang, Jiangxi); CD, female (Lishui, Zhejiang); A, C, dorsal view; B, D, ventral view." pageId="8" pageNumber="389">Figs. 3AD</figureCitation>
,
<figureCitation id="18D82A1FF741191D0F03F9657001F88D" box="[305,367,1783,1809]" captionStart="FIGURE 4" captionStartId="6.[151,250,1908,1932]" captionTargetBox="[176,1407,595,1866]" captionTargetId="figure-216@6.[151,1435,561,1888]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Bolbochromus species in oblique view. AB, Bolbochromus sunfengyii Gao &amp; Yang, new species; A, holotype; B, female paratype; CD, Bolbochromus nomurai Li &amp; Krikken, 2013; C, male (Xishuangbanna, Yunnan); D, female (Xishuangbanna, Yunnan); EH, Bolbochromus sinensis Krikken &amp; Li, 2013; E, male (Nanchang, Jiangxi); H, female (Lishui, Zhejiang); not to scale." pageId="8" pageNumber="389">4EF</figureCitation>
,
<figureCitation id="18D82A1FF741191D0F48F96570D4F88E" box="[378,442,1783,1810]" captionStart="FIGURE 5" captionStartId="7.[151,250,1193,1217]" captionTargetBox="[161,1424,262,1154]" captionTargetId="figure-17@7.[157,1431,235,1170]" captionTargetPageId="7" captionText="FIGURE 5. Aedeagus of Bolbochromus species.AC, A*C*, Bolbochromus sunfengyii Gao &amp; Yang, new species (holotype), images denoted by an asterisk are enlarged version from AC; DF, Bolbochromus sinensis Krikken &amp; Li, 2013 (Nanchang, Jiangxi); GI, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan); A, A*, D, G, dorsal view; B, B*, E, H, lateral view; C, C*, F, I, ventral view; scale bar 0.2 mm." pageId="8" pageNumber="389">5DF</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F741191D0EA5F8A672FAF8D0" box="[151,916,1844,1869]" pageId="8" pageNumber="389" type="reference_group">
<paragraph id="805C369AF741191D0EA5F8A672FAF8D0" blockId="8.[151,916,1844,1869]" box="[151,916,1844,1869]" pageId="8" pageNumber="389">
<treatmentCitationGroup id="A0F311B4F741191D0EA5F8A672FAF8D0" box="[151,916,1844,1869]" pageId="8" pageNumber="389">
<treatmentCitation id="0142108BF741191D0EA5F8A673E9F8D0" author="Krikken, J. &amp; Li, C. - L." box="[151,647,1844,1869]" page="514" pageId="8" pageNumber="389" year="2013">
<taxonomicName id="47E34D19F741191D0EA5F8A673E9F8D0" ID-CoL="7SPCP" authority="Krikken &amp; Li, 2013: 514" authorityName="Krikken &amp; Li" authorityPageNumber="514" authorityYear="2013" box="[151,647,1844,1869]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="389" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F741191D0EA5F8A670EEF8D0" box="[151,384,1844,1868]" italics="true" pageId="8" pageNumber="389">Bolbochromus sinensis</emphasis>
<bibRefCitation id="E4724B6BF741191D0FB5F8A673E9F8D0" author="Krikken, J. &amp; Li, C. - L." box="[391,647,1844,1869]" pageId="8" pageNumber="389" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Krikken &amp; Li, 2013: 514</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0142108BF741191D0CA3F8A672FEF8D0" author="Nikolajev, G. V. &amp; Kral, D." box="[657,912,1844,1868]" page="35" pageId="8" pageNumber="389" year="2016">
<bibRefCitation id="E4724B6BF741191D0CA3F8A672FEF8D0" author="Nikolajev, G. V. &amp; Kral, D." box="[657,912,1844,1868]" pageId="8" pageNumber="389" pagination="33 - 36" refId="ref4906" refString="Nikolajev, G. V., Kral, D. &amp; Bezdӗk. A. (2016) Bolboceratinae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera, volume 3. Koninklijke Brill NV, Leiden, pp. 33 - 36. https: // doi. org / 10.1163 / 9789004309142" type="book chapter" year="2016">
Nikolajev
<emphasis id="B297EA88F741191D0CC9F8A77240F8D0" box="[763,814,1844,1868]" italics="true" pageId="8" pageNumber="389">et al.</emphasis>
2016: 35
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F741191C0EA5F8E1735AFF65" lastPageId="9" lastPageNumber="390" pageId="8" pageNumber="389" type="materials_examined">
<paragraph id="805C369AF741191D0EA5F8E17266F811" blockId="8.[151,1436,1907,2040]" box="[151,776,1907,1933]" pageId="8" pageNumber="389">
<materialsCitation id="308B3CC7F741191D0EA5F8E1726AF811" box="[151,772,1907,1933]" country="China" location="Shaowu" municipality="Tachulan" pageId="8" pageNumber="389" specimenCount="1" typeStatus="holotype">
<emphasis id="B297EA88F741191D0EA5F8E17058F811" bold="true" box="[151,310,1907,1933]" pageId="8" pageNumber="389">
<typeStatus id="5F588838F741191D0EA5F8E171BCF811" box="[151,210,1907,1933]" pageId="8" pageNumber="389">Type</typeStatus>
locality.
</emphasis>
“FUKIEN: S
<collectingCountry id="F8F4760AF741191D0FE7F8E17375F811" box="[469,539,1907,1933]" name="China" pageId="8" pageNumber="389">China</collectingCountry>
,
<location id="853C6041F741191D0C14F8E173EBF811" LSID="urn:lsid:plazi:treatment:084A878CF741191C0EA5F94070CDFE15:853C6041F741191D0C14F8E173EBF811" box="[550,645,1907,1933]" country="China" municipality="Tachulan" name="Shaowu" pageId="8" pageNumber="389">Shaowu</location>
:
<collectingMunicipality id="6038ACE0F741191D0CBCF8E17394F811" box="[654,762,1907,1933]" pageId="8" pageNumber="389">Tachulan</collectingMunicipality>
</materialsCitation>
,
</paragraph>
<paragraph id="805C369AF741191D0EF5F8057354F82D" blockId="8.[151,1436,1907,2040]" box="[199,570,1942,1969]" pageId="8" pageNumber="389">
<emphasis id="B297EA88F741191D0EF5F8057081F82C" bold="true" box="[199,495,1942,1969]" pageId="8" pageNumber="389">
<typeStatus id="5F588838F741191D0EF5F805706CF82D" box="[199,258,1943,1969]" pageId="8" pageNumber="389">Type</typeStatus>
material examined.
</emphasis>
None.
</paragraph>
<paragraph id="805C369AF741191D0EF5F82B74F2F86B" blockId="8.[151,1436,1907,2040]" pageId="8" pageNumber="389">
<emphasis id="B297EA88F741191D0EF5F82B7354F84F" bold="true" box="[199,570,1977,2003]" pageId="8" pageNumber="389">Additional material examined</emphasis>
(
<specimenCount id="96E5FD13F741191D0C60F8287318F848" box="[594,630,1978,2004]" count="1" pageId="8" pageNumber="389" type="male">1♂</specimenCount>
,
<specimenCount id="96E5FD13F741191D0CBBF82873D1F848" box="[649,703,1978,2004]" count="3" pageId="8" pageNumber="389" type="female">3♀♀</specimenCount>
).
<emphasis id="B297EA88F741191D0CEDF82B72DDF84F" bold="true" box="[735,947,1977,2003]" pageId="8" pageNumber="389">
<materialsCitation id="308B3CC7F741191D0CEDF82B72C1F84F" box="[735,943,1977,2003]" country="China" location="Zhejiang" pageId="8" pageNumber="389" specimenCount="1" stateProvince="Zhejiang" typeStatus="holotype">
<collectingCountry id="F8F4760AF741191D0CEDF82B7242F84F" box="[735,812,1977,2003]" name="China" pageId="8" pageNumber="389">China</collectingCountry>
:
<collectingRegion id="4227F878F741191D0D0DF82B72C1F84F" box="[831,943,1977,2003]" country="China" name="Zhejiang" pageId="8" pageNumber="389">Zhejiang</collectingRegion>
</materialsCitation>
:
</emphasis>
<materialsCitation id="308B3CC7F741191D0DF3F8287083F86B" collectingDate="2024-05-01" collectionCode="CHG" collectorName="Cailiao &amp; Lishui City &amp; Jing-Xiang Luo" country="China" pageId="8" pageNumber="389" specimenCount="1" specimenCount-female="1" stateProvince="Zhejiang" typeStatus="holotype">
<specimenCount id="96E5FD13F741191D0DF3F828728AF848" box="[961,996,1978,2004]" count="1" pageId="8" pageNumber="389" type="female">1♀</specimenCount>
(
<collectionCode id="E6F2AE5FF741191D0DC8F8287554F848" box="[1018,1082,1978,2004]" pageId="8" pageNumber="389">CHG</collectionCode>
),
<collectorName id="2D16534CF741191D0A66F82875CBF848" box="[1108,1189,1978,2004]" pageId="8" pageNumber="389">Cailiao</collectorName>
(Ầ寮),
<collectorName id="2D16534CF741191D0B20F82874F6F848" box="[1298,1432,1978,2004]" pageId="8" pageNumber="389">Lishui City</collectorName>
,
<date id="F45D105AF741191D0EA5F84F7190F86B" box="[151,254,2013,2039]" pageId="8" pageNumber="389" value="2024-05-01">
<collectingDate id="E419E9B2F741191D0EA5F84F7190F86B" box="[151,254,2013,2039]" pageId="8" pageNumber="389" value="2024-05-01">1.V.2024</collectingDate>
</date>
,
<collectorName id="2D16534CF741191D0F38F84F70D4F86B" box="[266,442,2013,2039]" pageId="8" pageNumber="389">Jing-Xiang Luo</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="308B3CC7F741191D0FCBF84F74F6F86B" box="[505,1432,2013,2040]" collectionCode="CTKL" collectorName="Jiangshi &amp; Shaowu City" country="China" location="Fujian" longitude="117.279755" pageId="8" pageNumber="389" specimenCount="1" specimenCount-female="1" stateProvince="Fujian" typeStatus="holotype">
<emphasis id="B297EA88F741191D0FCBF84F7326F86B" bold="true" box="[505,584,2013,2039]" pageId="8" pageNumber="389">
<collectingRegion id="4227F878F741191D0FCBF84F7326F86B" box="[505,584,2013,2039]" country="China" name="Fujian" pageId="8" pageNumber="389">Fujian</collectingRegion>
</emphasis>
:
<specimenCount id="96E5FD13F741191D0C65F84F7317F86B" box="[599,633,2013,2039]" count="1" pageId="8" pageNumber="389" type="female">1♀</specimenCount>
(
<collectionCode id="E6F2AE5FF741191D0CBAF84F73B9F86B" box="[648,727,2013,2039]" pageId="8" pageNumber="389">CTKL</collectionCode>
), 27°25.40”N,
<geoCoordinate id="E5D7505DF741191D0DBAF84F7551F86B" box="[904,1087,2013,2039]" degrees="117" direction="east" minutes="16" orientation="longitude" pageId="8" pageNumber="389" precision="1" seconds="47.11" value="117.279755">117°1647.11”E</geoCoordinate>
,
<collectorName id="2D16534CF741191D0A79F84F75C9F86B" box="[1099,1191,2013,2039]" pageId="8" pageNumber="389">Jiangshi</collectorName>
(将石),
<collectorName id="2D16534CF741191D0B34F84F74F6F86B" box="[1286,1432,2013,2039]" pageId="8" pageNumber="389">Shaowu City</collectorName>
</materialsCitation>
,
</paragraph>
<paragraph id="805C369AF740191C0EA5FF05735AFF65" blockId="9.[151,1437,151,394]" pageId="9" pageNumber="390">
<date id="F45D105AF740191C0EA5FF057062FF2D" box="[151,268,151,177]" pageId="9" pageNumber="390" value="2024-05-12">12.V.2024</date>
, Tao-Kun Liao leg.;
<materialsCitation id="308B3CC7F740191C0FC9FF0571A6FF49" collectionCode="CTKL" latitude="26.146852" location="Fuzhou City" longLatPrecision="1" longitude="119.297935" municipality="Fuzhou Zoo" pageId="9" pageNumber="390" specimenCount="1" specimenCount-female="1">
<specimenCount id="96E5FD13F740191C0FC9FF057370FF2D" box="[507,542,151,177]" count="1" pageId="9" pageNumber="390" type="female">1♀</specimenCount>
(
<collectionCode id="E6F2AE5FF740191C0C1DFF057310FF2D" box="[559,638,151,177]" pageId="9" pageNumber="390">CTKL</collectionCode>
),
<geoCoordinate id="E5D7505DF740191C0CA1FF05725DFF2D" box="[659,819,151,177]" degrees="26" direction="north" minutes="8" orientation="latitude" pageId="9" pageNumber="390" precision="1" seconds="48.67" value="26.146852">26°848.67”N</geoCoordinate>
,
<geoCoordinate id="E5D7505DF740191C0D73FF057294FF2E" box="[833,1018,151,178]" degrees="119" direction="east" minutes="17" orientation="longitude" pageId="9" pageNumber="390" precision="1" seconds="52.56" value="119.297935">119°1752.56”E</geoCoordinate>
,
<collectingMunicipality id="6038ACE0F740191C0A35FF0575FFFF2E" box="[1031,1169,151,178]" pageId="9" pageNumber="390">Fuzhou Zoo</collectingMunicipality>
(DZ州ṅ物园),
<location id="853C6041F740191C0B7AFF0571A6FF49" LSID="urn:lsid:plazi:treatment:084A878CF741191C0EA5F94070CDFE15:853C6041F740191C0B7AFF0571A6FF49" latitude="26.146852" longLatPrecision="1" longitude="119.297935" municipality="Fuzhou Zoo" name="Fuzhou City" pageId="9" pageNumber="390">Fuzhou City</location>
</materialsCitation>
,
<date id="F45D105AF740191C0EE6FF297048FF49" box="[212,294,187,213]" pageId="9" pageNumber="390" value="2024-05">V.2024</date>
, Tao-Kun Liao leg.;
<materialsCitation id="308B3CC7F740191C0C3DFF2971A6FF65" collectionCode="CHG" location="Nanchang City" municipality="Agricultural University" pageId="9" pageNumber="390" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi">
<emphasis id="B297EA88F740191C0C3DFF297308FF49" bold="true" box="[527,614,187,213]" pageId="9" pageNumber="390">
<collectingRegion id="4227F878F740191C0C3DFF297308FF49" box="[527,614,187,213]" country="China" name="Jiangxi" pageId="9" pageNumber="390">Jiangxi</collectingRegion>
</emphasis>
:
<specimenCount id="96E5FD13F740191C0C44FF2973F7FF49" box="[630,665,187,213]" count="1" pageId="9" pageNumber="390" type="male">1♂</specimenCount>
(
<collectionCode id="E6F2AE5FF740191C0C9AFF297386FF49" box="[680,744,187,213]" pageId="9" pageNumber="390">CHG</collectionCode>
),
<collectingRegion id="4227F878F740191C0CCFFF297220FF49" box="[765,846,187,213]" country="China" name="Jiangxi" pageId="9" pageNumber="390">Jiangxi</collectingRegion>
<collectingMunicipality id="6038ACE0F740191C0D61FF297534FF49" box="[851,1114,187,213]" pageId="9" pageNumber="390">Agricultural University</collectingMunicipality>
(江ĦR业kṭ),
<location id="853C6041F740191C0B19FF2971A6FF65" LSID="urn:lsid:plazi:treatment:084A878CF741191C0EA5F94070CDFE15:853C6041F740191C0B19FF2971A6FF65" municipality="Agricultural University" name="Nanchang City" pageId="9" pageNumber="390" stateProvince="Jiangxi">Nanchang City</location>
</materialsCitation>
,
<date id="F45D105AF740191C0EE6FF4D7027FF66" box="[212,329,223,250]" pageId="9" pageNumber="390" value="2018-05-15">15.V.2018</date>
, Ding-Hua Shen leg.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F740191C0EF5FE917579FE82" box="[199,1047,259,286]" pageId="9" pageNumber="390" type="distribution">
<paragraph id="805C369AF740191C0EF5FE917579FE82" blockId="9.[151,1437,151,394]" box="[199,1047,259,286]" pageId="9" pageNumber="390">
<emphasis id="B297EA88F740191C0EF5FE91700FFE81" bold="true" box="[199,353,259,285]" pageId="9" pageNumber="390">Distribution.</emphasis>
<collectingCountry id="F8F4760AF740191C0F55FE9170C0FE81" box="[359,430,259,285]" name="China" pageId="9" pageNumber="390">China</collectingCountry>
:
<collectingRegion id="4227F878F740191C0F8BFE91736AFE81" box="[441,516,259,285]" country="China" name="Fujian" pageId="9" pageNumber="390">Fujian</collectingRegion>
,
<collectingRegion id="4227F878F740191C0C3DFE91730EFE81" box="[527,608,259,285]" country="China" name="Jiangxi" pageId="9" pageNumber="390">Jiangxi</collectingRegion>
<emphasis id="B297EA88F740191C0C55FE91726EFE81" bold="true" box="[615,768,259,285]" pageId="9" pageNumber="390">(new record)</emphasis>
,
<collectingRegion id="4227F878F740191C0D3FFE91721FFE81" box="[781,881,259,285]" country="China" name="Zhejiang" pageId="9" pageNumber="390">Zhejiang</collectingRegion>
<emphasis id="B297EA88F740191C0D4AFE91757FFE81" bold="true" box="[888,1041,259,285]" pageId="9" pageNumber="390">(new record)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F740191C0EF5FEB570CDFE15" pageId="9" pageNumber="390" type="discussion">
<paragraph id="805C369AF740191C0EF5FEB570CDFE15" blockId="9.[151,1437,151,394]" pageId="9" pageNumber="390">
<emphasis id="B297EA88F740191C0EF5FEB57055FEDD" bold="true" box="[199,315,295,321]" pageId="9" pageNumber="390">Remarks.</emphasis>
The identification of the specimens is based on comparison of the aedeagus, and the general external characters. The specimens from
<collectingRegion id="4227F878F740191C0C3BFED97334FEF9" box="[521,602,331,357]" country="China" name="Jiangxi" pageId="9" pageNumber="390">Jiangxi</collectingRegion>
and
<collectingRegion id="4227F878F740191C0CA6FED97396FEF9" box="[660,760,331,357]" country="China" name="Zhejiang" pageId="9" pageNumber="390">Zhejiang</collectingRegion>
Provinces mark the first provincial records for this species from the two provinces.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,437 @@
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<mods:title id="F4226E7963323EC28766E5C63746D498">Contributions to the genus Bolbochromus Boucomont, 1909 (Coleoptera: Geotrupidae: Bolboceratinae) from China, with the description of a new species</mods:title>
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<mods:namePart id="984D0207BEB51336CE0BAC1CF066CD58">Gao, Hao</mods:namePart>
<mods:affiliation id="974C43546714595550E33C6288C8152E">4 A, Hongtangxincheng, Cangshan Dist., Fuzhou, Fujian, 350028.</mods:affiliation>
</mods:name>
<mods:name id="5BFE8597E86581566D31C8CB896FD852" type="personal">
<mods:role id="E072A0D51A3F7D40A247DB713BFD64AB">
<mods:roleTerm id="26B4E369BA7D108940C43021E74507CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DA9993B41BE2E8D0B18665DB1721F540">Geshev, Georgi A.</mods:namePart>
</mods:name>
<mods:name id="7C867BE778EEB3A3B14833FEF77D86A6" type="personal">
<mods:role id="65A25ECC1073C3554463C698605F4D6B">
<mods:roleTerm id="EDEE67B583DD5A1A62580CB366D406CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD210302494F9FE1DD20929B052B045B">Yang, Qiao-Zhi</mods:namePart>
<mods:affiliation id="D7590FE665E39CA9D46DF97CB2D8043C">Department of Life Sciences, Imperial College London, Exhibition Road, London SW 7 2 BX, United Kingdom. &amp; Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="46ED07500ED14E4F195A3CC492DEC5EC">text</mods:typeOfResource>
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<mods:title id="3BCCC8FE63C7949CE916EC38FD23D8C7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F1B650509C837D937D3EE06BBCDA8358">
<mods:date id="490D43A3EE1C928D1216C08B7D7F04E7">2025</mods:date>
<mods:detail id="956008827EBEA99FB36A34E8ABA1ED9E" type="pubDate">
<mods:number id="B8B95244F13FA080A51A691671FAF482">2025-01-08</mods:number>
</mods:detail>
<mods:detail id="D424B523A9A44F1EB806C9610CFD5B16" type="volume">
<mods:number id="847A8BE8616AB0B50E8C09387AFF8F20">5566</mods:number>
</mods:detail>
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<mods:number id="8BCF0C5FB3AE6F22B426DD714BAE42CE">2</mods:number>
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<mods:start id="466B1C312073139C5B583A55777EB955">381</mods:start>
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<mods:location id="21EBDC70CCDE5B182B2A5AA7310C91C9">
<mods:url id="DABB56B7819BDC5046CF14FB00038C50">https://doi.org/10.11646/zootaxa.5566.2.8</mods:url>
</mods:location>
<mods:classification id="E3BFDB875C1F950532A931A88AFEDB6E">journal article</mods:classification>
<mods:identifier id="9EA77AA45C891CBA063FD6DD37B0BACD" type="DOI">10.11646/zootaxa.5566.2.8</mods:identifier>
<mods:identifier id="A863B7FEA99DF1B39CD2B6A30C1B4EEB" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="084A878CF74819100EA5FBA6727CFC1D" LSID="urn:lsid:plazi:treatment:084A878CF74819100EA5FBA6727CFC1D" httpUri="http://treatment.plazi.org/id/084A878CF74819100EA5FBA6727CFC1D" lastPageId="5" lastPageNumber="386" pageId="1" pageNumber="382">
<subSubSection id="C8F96511F74819140EA5FBA672B5FBD2" box="[151,987,1076,1103]" pageId="1" pageNumber="382" type="nomenclature">
<paragraph id="805C369AF74819140EA5FBA672B5FBD2" blockId="1.[151,987,1076,1139]" box="[151,987,1076,1103]" pageId="1" pageNumber="382">
<heading id="DB1481F6F74819140EA5FBA672B5FBD2" bold="true" box="[151,987,1076,1103]" fontSize="11" level="1" pageId="1" pageNumber="382" reason="1">
<emphasis id="B297EA88F74819140EA5FBA672B5FBD2" bold="true" box="[151,987,1076,1103]" pageId="1" pageNumber="382">
<taxonomicName id="47E34D19F74819140EA5FBA67250FBD3" authority="Gao &amp; Yang" authorityName="Gao &amp; Yang" authorityYear="2025" box="[151,830,1076,1103]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="382" phylum="Arthropoda" rank="species" species="sunfengyii" status="sp. nov." subGenus="Bolbochromus">
<emphasis id="B297EA88F74819140EA5FBA67027FBD3" bold="true" box="[151,329,1076,1103]" italics="true" pageId="1" pageNumber="382">Bolbochromus</emphasis>
(
<emphasis id="B297EA88F74819140F68FBA67362FBD3" bold="true" box="[346,524,1076,1103]" italics="true" pageId="1" pageNumber="382">Bolbochromus</emphasis>
)
<emphasis id="B297EA88F74819140C2FFBA673F2FBD2" bold="true" box="[541,668,1076,1102]" italics="true" pageId="1" pageNumber="382">sunfengyii</emphasis>
Gao &amp; Yang
</taxonomicName>
,
<taxonomicNameLabel id="A9A457F3F74819140D7EFBA772B5FBD2" box="[844,987,1076,1103]" pageId="1" pageNumber="382" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74819140EA5FBCA7315FBEE" box="[151,635,1112,1139]" pageId="1" pageNumber="382" type="description">
<paragraph id="805C369AF74819140EA5FBCA7315FBEE" blockId="1.[151,987,1076,1139]" box="[151,635,1112,1139]" pageId="1" pageNumber="382">
(
<figureCitation id="18D82A1FF74819140EADFBCA704BFBEF" box="[159,293,1112,1139]" captionStart="FIGURE 1" captionStartId="2.[151,250,1948,1972]" captionTargetBox="[162,1418,492,1901]" captionTargetId="figure-164@2.[151,1435,450,1922]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Bolbochromus sunfengyii Gao &amp; Yang, new species. AB, holotype; CD, female paratype; A, C, dorsal view; B, D, ventral view." pageId="1" pageNumber="382">Figs. 1AD</figureCitation>
,
<figureCitation id="18D82A1FF74819140F03FBCB7019FBEF" box="[305,375,1113,1139]" captionStart="FIGURE 4" captionStartId="6.[151,250,1908,1932]" captionTargetBox="[176,1407,595,1866]" captionTargetId="figure-216@6.[151,1435,561,1888]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Bolbochromus species in oblique view. AB, Bolbochromus sunfengyii Gao &amp; Yang, new species; A, holotype; B, female paratype; CD, Bolbochromus nomurai Li &amp; Krikken, 2013; C, male (Xishuangbanna, Yunnan); D, female (Xishuangbanna, Yunnan); EH, Bolbochromus sinensis Krikken &amp; Li, 2013; E, male (Nanchang, Jiangxi); H, female (Lishui, Zhejiang); not to scale." pageId="1" pageNumber="382">4AB</figureCitation>
,
<figureCitation id="18D82A1FF74819140FB0FBCA7086FBEF" box="[386,488,1112,1139]" captionStart="FIGURE 5" captionStartId="7.[151,250,1193,1217]" captionTargetBox="[161,1424,262,1154]" captionTargetId="figure-17@7.[157,1431,235,1170]" captionTargetPageId="7" captionText="FIGURE 5. Aedeagus of Bolbochromus species.AC, A*C*, Bolbochromus sunfengyii Gao &amp; Yang, new species (holotype), images denoted by an asterisk are enlarged version from AC; DF, Bolbochromus sinensis Krikken &amp; Li, 2013 (Nanchang, Jiangxi); GI, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan); A, A*, D, G, dorsal view; B, B*, E, H, lateral view; C, C*, F, I, ventral view; scale bar 0.2 mm." pageId="1" pageNumber="382">5AC, A</figureCitation>
*C*, 6AE)
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74819140EA5FB337433FA2A" pageId="1" pageNumber="382" type="materials_examined">
<paragraph id="805C369AF74819140EA5FB337578FB27" blockId="1.[151,1437,1184,2039]" box="[151,1046,1184,1211]" pageId="1" pageNumber="382">
<emphasis id="B297EA88F74819140EA5FB337058FB26" bold="true" box="[151,310,1184,1211]" pageId="1" pageNumber="382">Type locality.</emphasis>
<collectingCountry id="F8F4760AF74819140F0EFB3270ECFB26" box="[316,386,1184,1210]" name="China" pageId="1" pageNumber="382">China</collectingCountry>
,
<collectingRegion id="4227F878F74819140FBEFB337321FB26" box="[396,591,1184,1211]" country="China" name="Yunnan" pageId="1" pageNumber="382">Yunnan Province</collectingRegion>
, Xishuangbanna Prefecture, Gasa Town.
</paragraph>
<paragraph id="805C369AF74819140EF5FB5773E0FAF2" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="382">
<emphasis id="B297EA88F74819140EF5FB577018FB42" bold="true" box="[199,374,1220,1247]" pageId="1" pageNumber="382">Type material</emphasis>
(
<specimenCount id="96E5FD13F74819140FBDFB5770B8FB43" box="[399,470,1221,1247]" count="11" pageId="1" pageNumber="382" type="male">11♂♂</specimenCount>
,
<specimenCount id="96E5FD13F74819140FD9FB57735DFB43" box="[491,563,1221,1247]" count="20" pageId="1" pageNumber="382" type="female">20♀♀</specimenCount>
).
<emphasis id="B297EA88F74819140C62FB5773A9FB43" bold="true" box="[592,711,1221,1247]" pageId="1" pageNumber="382">
<typeStatus id="5F588838F74819140C62FB5773AFFB43" box="[592,705,1221,1247]" pageId="1" pageNumber="382" type="holotype">Holotype</typeStatus>
:
</emphasis>
<materialsCitation id="308B3CC7F74819140CE4FB567210FA9E" collectingDate="2020-09" collectionCode="CAU" country="China" location="Xishuangbanna Prefecture" municipality="Gasa Town" pageId="1" pageNumber="382" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B297EA88F74819140CE4FB567246FB42" bold="true" box="[726,808,1220,1246]" pageId="1" pageNumber="382">
<collectingCountry id="F8F4760AF74819140CE4FB56724DFB42" box="[726,803,1220,1246]" name="China" pageId="1" pageNumber="382">China</collectingCountry>
:
</emphasis>
<specimenCount id="96E5FD13F74819140D0AFB577235FB43" box="[824,859,1221,1247]" count="1" pageId="1" pageNumber="382" type="male">1♂</specimenCount>
(
<collectionCode id="E6F2AE5FF74819140D40FB5772DCFB43" box="[882,946,1221,1247]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="1" pageNumber="382">CAU</collectionCode>
), “
<collectingMunicipality id="6038ACE0F74819140DEBFB57750FFB43" box="[985,1121,1221,1247]" pageId="1" pageNumber="382">Gasa Town</collectingMunicipality>
(Ŕ洒Dz),
<location id="853C6041F74819140ADCFB567063FA9E" LSID="urn:lsid:plazi:treatment:084A878CF74819100EA5FBA6727CFC1D:853C6041F74819140ADCFB567063FA9E" country="China" municipality="Gasa Town" name="Xishuangbanna Prefecture" pageId="1" pageNumber="382" stateProvince="Yunnan">Xishuangbanna Prefecture</location>
,
<collectingRegion id="4227F878F74819140F25FB7B70B7FA9E" box="[279,473,1256,1283]" country="China" name="Yunnan" pageId="1" pageNumber="382">Yunnan Province</collectingRegion>
,
<collectingCountry id="F8F4760AF74819140FD6FB7A7344FA9E" box="[484,554,1256,1282]" name="China" pageId="1" pageNumber="382">China</collectingCountry>
,
<date id="F45D105AF74819140C07FB7B73FAFA9F" box="[565,660,1257,1283]" pageId="1" pageNumber="382" value="2020-09">
<collectingDate id="E419E9B2F74819140C07FB7B73FAFA9F" box="[565,660,1257,1283]" pageId="1" pageNumber="382" value="2020-09">IX.2020</collectingDate>
</date>
, local collector leg.”
</materialsCitation>
.
<materialsCitation id="308B3CC7F74819140DBBFB7B73E4FAF2" collectingDate="2020-08-15" collectingDateMax="2020-08-30" collectingDateMin="2020-08-15" collectionCode="CFYS, CQZY, CMJU, CHG" country="China" location="Xishuangbanna Prefecture" municipality="Gasa Town" pageId="1" pageNumber="382" specimenCount="30" specimenCount-female="20" specimenCount-male="10" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B297EA88F74819140DBBFB7B750DFA9E" bold="true" box="[905,1123,1256,1283]" pageId="1" pageNumber="382">
<typeStatus id="5F588838F74819140DBBFB7B756BFA9F" box="[905,1029,1257,1283]" pageId="1" pageNumber="382" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F8F4760AF74819140A23FB7A7530FA9E" box="[1041,1118,1256,1282]" name="China" pageId="1" pageNumber="382">China</collectingCountry>
:
</emphasis>
<specimenCount id="96E5FD13F74819140A58FB7B75CAFA9F" box="[1130,1188,1257,1283]" count="8" pageId="1" pageNumber="382" type="male">8♂♂</specimenCount>
,
<specimenCount id="96E5FD13F74819140A9DFB7B7598FA9F" box="[1199,1270,1257,1283]" count="17" pageId="1" pageNumber="382" type="female">17♀♀</specimenCount>
(
<collectionCode id="E6F2AE5FF74819140B36FB7B7421FA9F" box="[1284,1359,1257,1283]" pageId="1" pageNumber="382">CFYS</collectionCode>
), same data as the holotype;
<specimenCount id="96E5FD13F74819140FA2FA9F70DAFABB" box="[400,436,1293,1319]" count="1" pageId="1" pageNumber="382" type="male">1♂</specimenCount>
,
<specimenCount id="96E5FD13F74819140FF6FA9F7088FABB" box="[452,486,1293,1319]" count="1" pageId="1" pageNumber="382" type="female">1♀</specimenCount>
(
<collectionCode id="E6F2AE5FF74819140FCAFA9F7324FABB" box="[504,586,1293,1319]" pageId="1" pageNumber="382">CQZY</collectionCode>
), same data as the
<typeStatus id="5F588838F74819140D1AFA9E72E0FABA" box="[808,910,1292,1318]" pageId="1" pageNumber="382" type="holotype">holotype</typeStatus>
;
<specimenCount id="96E5FD13F74819140DAEFA9F72D0FABB" box="[924,958,1293,1319]" count="1" pageId="1" pageNumber="382" type="female">1♀</specimenCount>
(
<collectionCode id="E6F2AE5FF74819140DE2FA9F754EFABB" box="[976,1056,1293,1319]" pageId="1" pageNumber="382">CMJU</collectionCode>
), same data as the
<typeStatus id="5F588838F74819140ACDFA9E740BFABA" box="[1279,1381,1292,1318]" pageId="1" pageNumber="382" type="holotype">holotype</typeStatus>
;
<specimenCount id="96E5FD13F74819140B40FA9F74F9FABB" box="[1394,1431,1293,1319]" count="1" pageId="1" pageNumber="382" type="male">1♂</specimenCount>
, (
<collectionCode id="E6F2AE5FF74819140EADFAA371B1FAD7" box="[159,223,1329,1355]" pageId="1" pageNumber="382">CHG</collectionCode>
), same data as the
<typeStatus id="5F588838F74819140F9EFAA2737CFAD6" box="[428,530,1328,1354]" pageId="1" pageNumber="382" type="holotype">holotype</typeStatus>
;
<specimenCount id="96E5FD13F74819140C2EFAA37351FAD7" box="[540,575,1329,1355]" count="1" pageId="1" pageNumber="382" type="female">1♀</specimenCount>
(
<collectionCode id="E6F2AE5FF74819140C7FFAA373E3FAD7" box="[589,653,1329,1355]" pageId="1" pageNumber="382">CHG</collectionCode>
), “
<collectingMunicipality id="6038ACE0F74819140C98FAA37247FAD7" box="[682,809,1329,1355]" pageId="1" pageNumber="382">Gasa Town</collectingMunicipality>
(Ŕ洒Dz),
<location id="853C6041F74819140D91FAA275A3FAD6" LSID="urn:lsid:plazi:treatment:084A878CF74819100EA5FBA6727CFC1D:853C6041F74819140D91FAA275A3FAD6" box="[931,1229,1328,1354]" country="China" municipality="Gasa Town" name="Xishuangbanna Prefecture" pageId="1" pageNumber="382" stateProvince="Yunnan">Xishuangbanna Prefecture</location>
,
<collectingRegion id="4227F878F74819140AE4FAA374F6FAD6" box="[1238,1432,1328,1355]" country="China" name="Yunnan" pageId="1" pageNumber="382">Yunnan Province</collectingRegion>
,
<collectingCountry id="F8F4760AF74819140EA5FAC671B1FAF3" box="[151,223,1364,1391]" name="China" pageId="1" pageNumber="382">China</collectingCountry>
,
<date id="F45D105AF74819140ED6FAC670CEFAF3" box="[228,416,1364,1391]" pageId="1" pageNumber="382" value="2020-08-15" valueMax="2020-08-30" valueMin="2020-08-15">
<collectingDate id="E419E9B2F74819140ED6FAC670CEFAF3" box="[228,416,1364,1391]" pageId="1" pageNumber="382" value="2020-08-15" valueMax="2020-08-30" valueMin="2020-08-15">1530.VIII.2020</collectingDate>
</date>
, local collector leg.”
</materialsCitation>
.
</paragraph>
<paragraph id="805C369AF74819140EF5FAEA7433FA2A" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="382">
<emphasis id="B297EA88F74819140EF5FAEA735DFA0E" bold="true" box="[199,563,1400,1426]" pageId="1" pageNumber="382">Additional material examined</emphasis>
(
<specimenCount id="96E5FD13F74819140C75FAEB7302FA0F" box="[583,620,1401,1427]" count="2" pageId="1" pageNumber="382" type="male">2♂</specimenCount>
,
<specimenCount id="96E5FD13F74819140C49FAEB73F1FA0F" box="[635,671,1401,1427]" count="2" pageId="1" pageNumber="382" type="female">2♀</specimenCount>
).
<materialsCitation id="308B3CC7F74819140C8BFAEB700DFA2A" collectingDate="2020-05" collectionCode="CBHH" location="Dehong Prefecture" pageId="1" pageNumber="382" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="96E5FD13F74819140C8BFAEB73B3FA0F" box="[697,733,1401,1427]" count="1" pageId="1" pageNumber="382" type="male">1♂</specimenCount>
,
<specimenCount id="96E5FD13F74819140CDFFAEB727EFA0F" box="[749,784,1401,1427]" count="1" pageId="1" pageNumber="382" type="female">1♀</specimenCount>
(
<collectionCode id="E6F2AE5FF74819140D10FAEB721BFA0F" box="[802,885,1401,1427]" pageId="1" pageNumber="382">CBHH</collectionCode>
),
<location id="853C6041F74819140DBEFAEA750AFA0E" LSID="urn:lsid:plazi:treatment:084A878CF74819100EA5FBA6727CFC1D:853C6041F74819140DBEFAEA750AFA0E" box="[908,1124,1400,1426]" name="Dehong Prefecture" pageId="1" pageNumber="382" stateProvince="Yunnan">Dehong Prefecture</location>
,
<collectingRegion id="4227F878F74819140A40FAEB7459FA0E" box="[1138,1335,1400,1427]" country="China" name="Yunnan" pageId="1" pageNumber="382">Yunnan Province</collectingRegion>
,
<date id="F45D105AF74819140B74FAEB74F6FA0F" box="[1350,1432,1401,1427]" pageId="1" pageNumber="382" value="2020-05">
<collectingDate id="E419E9B2F74819140B74FAEB74F6FA0F" box="[1350,1432,1401,1427]" pageId="1" pageNumber="382" value="2020-05">V.2020</collectingDate>
</date>
, local collector leg
</materialsCitation>
;
<materialsCitation id="308B3CC7F74819140F5DFA0F7434FA2A" box="[367,1370,1436,1463]" collectingDate="2019-05" collectionCode="CBHH" location="Diqing Prefecture" pageId="1" pageNumber="382" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="96E5FD13F74819140F5DFA0F70FDFA2B" box="[367,403,1437,1463]" count="1" pageId="1" pageNumber="382" type="male">1♂</specimenCount>
,
<specimenCount id="96E5FD13F74819140F92FA0F70D6FA2B" box="[416,440,1437,1463]" count="1" pageId="1" pageNumber="382" type="female">1♀</specimenCount>
(
<collectionCode id="E6F2AE5FF74819140FF3FA0F7373FA2B" box="[449,541,1437,1463]" pageId="1" pageNumber="382">CBHH</collectionCode>
),
<location id="853C6041F74819140C03FA0E7392FA2A" LSID="urn:lsid:plazi:treatment:084A878CF74819100EA5FBA6727CFC1D:853C6041F74819140C03FA0E7392FA2A" box="[561,764,1436,1462]" name="Diqing Prefecture" pageId="1" pageNumber="382" stateProvince="Yunnan">Diqing Prefecture</location>
,
<collectingRegion id="4227F878F74819140D34FA0F72A6FA2A" box="[774,968,1436,1463]" country="China" name="Yunnan" pageId="1" pageNumber="382">Yunnan Province</collectingRegion>
,
<collectingMethod id="59A24E8DF74819140DE6FA0E754BFA2B" box="[980,1061,1436,1463]" pageId="1" pageNumber="382">by trap</collectingMethod>
,
<date id="F45D105AF74819140A02FA0F75ECFA2B" box="[1072,1154,1437,1463]" pageId="1" pageNumber="382" value="2019-05">
<collectingDate id="E419E9B2F74819140A02FA0F75ECFA2B" box="[1072,1154,1437,1463]" pageId="1" pageNumber="382" value="2019-05">V.2019</collectingDate>
</date>
, local collector leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74819100EF5FA52738BFEF9" lastPageId="5" lastPageNumber="386" pageId="1" pageNumber="382" type="description">
<paragraph id="805C369AF74819160EF5FA52745BF864" blockId="1.[151,1437,1184,2039]" lastBlockId="3.[151,1437,1761,2040]" lastPageId="3" lastPageNumber="384" pageId="1" pageNumber="382">
<emphasis id="B297EA88F74819140EF5FA5273D2FA46" bold="true" box="[199,700,1472,1499]" pageId="1" pageNumber="382">
Description of the
<typeStatus id="5F588838F74819140F93FA537367FA47" box="[417,521,1473,1499]" pageId="1" pageNumber="382" type="holotype">holotype</typeStatus>
, male. General
</emphasis>
(
<figureCitation id="18D82A1FF74819140CF9FA527221FA47" box="[715,847,1472,1499]" captionStart="FIGURE 1" captionStartId="2.[151,250,1948,1972]" captionTargetBox="[162,1418,492,1901]" captionTargetId="figure-164@2.[151,1435,450,1922]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Bolbochromus sunfengyii Gao &amp; Yang, new species. AB, holotype; CD, female paratype; A, C, dorsal view; B, D, ventral view." pageId="1" pageNumber="382">Figs. 1AB</figureCitation>
,
<figureCitation id="18D82A1FF74819140D68FA537210FA47" box="[858,894,1473,1499]" captionStart="FIGURE 4" captionStartId="6.[151,250,1908,1932]" captionTargetBox="[176,1407,595,1866]" captionTargetId="figure-216@6.[151,1435,561,1888]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Bolbochromus species in oblique view. AB, Bolbochromus sunfengyii Gao &amp; Yang, new species; A, holotype; B, female paratype; CD, Bolbochromus nomurai Li &amp; Krikken, 2013; C, male (Xishuangbanna, Yunnan); D, female (Xishuangbanna, Yunnan); EH, Bolbochromus sinensis Krikken &amp; Li, 2013; E, male (Nanchang, Jiangxi); H, female (Lishui, Zhejiang); not to scale." pageId="1" pageNumber="382">4A</figureCitation>
): Body convex, dorsum glossy, maximum width at pronotum posterior to middle.
<emphasis id="B297EA88F74819140C34FA76733DFA62" bold="true" box="[518,595,1508,1534]" pageId="1" pageNumber="382">Color:</emphasis>
Head black. Pronotum with black along anterior and posterior margin, with the posterior widening towards middle; middle of disc more testaceous than the rest of pronotum. Scutellum black. Elytra bicolored testaceous and black. Sutural interstria black along entire length; interstriae 2 to 4 partially black; area from interstria 5 to 9 on the lateral edge of the elytron starts black, intensifying from the distal one-third to the two-thirds, and then decreasing towards lateral edge. Striae basal margins black to elytra apex.
<emphasis id="B297EA88F74819140B32F9E77427F913" bold="true" box="[1280,1353,1653,1679]" pageId="1" pageNumber="382">Head:</emphasis>
Dorsal surface covered with coarse punctures, glabrous. Vertex with distinct, transverse tubercle, rounded apically. Frons along inner margins of eyes barely raised. Clypeus broadly arcuate anteriorly; middle of anterior margin with low tubercle; margin of ocular canthus evenly arcuate; ocular canthus completely bisecting eyes. Antenna testaceous; with 11 antennomeres, antennomere 1 longer than any of 28, antennomere 2 elongated and thick, antennomeres 38 gradually thickening, antennomeres 911 elliptically swollen, inner face of antennomere 9 with glabrous plate.
<emphasis id="B297EA88F74819140EA5F8DF7077F8FB" bold="true" box="[151,281,1869,1895]" pageId="1" pageNumber="382">Pronotum:</emphasis>
Each lateral third with scattered large punctures, central third impunctate except anteriorly; smaller secondary punctures distributed along lateral margins. Disc with distinctly impressed, median longitudinal groove terminating before reaching anterior and posterior margins; entire length with sparse punctures. Anterior triangular flat area with small punctures in front of mid-longitudinal impression. Lateral foveae barely impressed, with several punctures. Anterior angles obtuse. Anterior and lateral margins with border terminating on basal margin at level of humeral umbone of elytra; lateral margins fringed with setae underneath, visible from above.
<emphasis id="B297EA88F74B19170AF9FF047423FF2C" bold="true" box="[1227,1357,150,176]" pageId="2" pageNumber="383">Scutellum:</emphasis>
Shiny, with extremely fine nearly invisible punctures along anterior margin. Distinct border on both sides relative to the elytra.
<emphasis id="B297EA88F74B19170ED4FF4D7055FF65" bold="true" box="[230,315,223,249]" pageId="2" pageNumber="383">Elytra:</emphasis>
Each elytron with 13 punctate striae, all evenly punctate along their lengths; with 7 striae between suture and humeral umbone. Sutural stria running parallel to scutellum and terminating at elytral base, interrupting stria 2 before elytral base; stria 5 ending before elytral base; striae 8 and 9 terminating on humeral umbone, not reaching elytral base; stria 10 extending around and below humeral umbone, longer than striae 8 and 9, not reaching elytral base; stria 11 terminating at elytral mid-length, well before elytral base; stria 12 extending to approximately same length as stria 10 not reaching elytral base. Stria 13 with very deep punctures, situated in reflexed lateral margin of elytron, reaching elytral base. Reflexed lateral margin of elytra with setae of two lengths, long and short setae irregularly alternate, sparsely and evenly scattered between punctures of stria 13, long setae largely located at anterior, short setae more numerous than long setae. Elytral interstriae slightly convex; surface smooth, micropunctures almost invisible. Elytral apices acuminate. Ventral face of epipleuron black, with single row of setae.
<emphasis id="B297EA88F74A19160EEFF8E37070F817" bold="true" box="[221,286,1905,1931]" pageId="3" pageNumber="384">Legs:</emphasis>
Protibia with nine teeth along outer margin, gradually increasing in size from base to apex, with distal three teeth much larger than preceding. Apical mesotibial spurs adjacent to each other, unequal in length, with inner spur shorter than outer spur; apical metatibial spurs subequal in length, spaced apart by basal width of a spur.
<emphasis id="B297EA88F74A19160EA5F84F7064F86B" bold="true" box="[151,266,2013,2039]" pageId="3" pageNumber="384">Aedeagus</emphasis>
(
<figureCitation id="18D82A1FF74A19160F28F84F70D0F86B" box="[282,446,2013,2040]" captionStart="FIGURE 5" captionStartId="7.[151,250,1193,1217]" captionTargetBox="[161,1424,262,1154]" captionTargetId="figure-17@7.[157,1431,235,1170]" captionTargetPageId="7" captionText="FIGURE 5. Aedeagus of Bolbochromus species.AC, A*C*, Bolbochromus sunfengyii Gao &amp; Yang, new species (holotype), images denoted by an asterisk are enlarged version from AC; DF, Bolbochromus sinensis Krikken &amp; Li, 2013 (Nanchang, Jiangxi); GI, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan); A, A*, D, G, dorsal view; B, B*, E, H, lateral view; C, C*, F, I, ventral view; scale bar 0.2 mm." pageId="3" pageNumber="384">Figs. 5AC, A</figureCitation>
*C*): Slender, with a pair of distinct, sclerotized, elongate-convex parameres.
</paragraph>
<caption id="D49C6612F74B19170EA5F80E70F8F844" pageId="2" pageNumber="383" startId="2.[151,250,1948,1972]" targetBox="[162,1418,492,1901]" targetPageId="2" targetType="figure">
<paragraph id="805C369AF74B19170EA5F80E70F8F844" blockId="2.[151,1436,1948,2008]" pageId="2" pageNumber="383">
<emphasis id="B297EA88F74B19170EA5F80E707CF828" bold="true" box="[151,274,1948,1972]" pageId="2" pageNumber="383">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="47E34D19F74B19170FB4F80E7265F829" authority="Gao &amp; Yang" authorityName="Gao &amp; Yang" authorityYear="2025" box="[390,779,1948,1973]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="383" phylum="Arthropoda" rank="species" species="sunfengyii" status="sp. nov.">
<emphasis id="B297EA88F74B19170FB4F80E73E9F829" box="[390,647,1948,1973]" italics="true" pageId="2" pageNumber="383">Bolbochromus sunfengyii</emphasis>
Gao &amp; Yang
</taxonomicName>
,
<emphasis id="B297EA88F74B19170D26F80F72FEF828" bold="true" box="[788,912,1948,1973]" pageId="2" pageNumber="383">
<taxonomicNameLabel id="A9A457F3F74B19170D26F80F72FEF828" box="[788,912,1948,1973]" pageId="2" pageNumber="383" rank="species">new species</taxonomicNameLabel>
</emphasis>
. AB, holotype; CD, female paratype; A, C, dorsal view; B, D, ventral view.
</paragraph>
</caption>
<caption id="D49C6612F74A19160EA5F9C17244F913" pageId="3" pageNumber="384" startId="3.[151,250,1619,1643]" targetBox="[168,1415,192,1567]" targetPageId="3" targetType="figure">
<paragraph id="805C369AF74A19160EA5F9C17244F913" blockId="3.[151,1436,1619,1679]" pageId="3" pageNumber="384">
<emphasis id="B297EA88F74A19160EA5F9C17079F9F7" bold="true" box="[151,279,1619,1643]" pageId="3" pageNumber="384">FIGURE 2.</emphasis>
Habitus of
<taxonomicName id="47E34D19F74A19160FA8F9C1721FF9F7" authority="Li &amp; Krikken, 2013" authorityName="Li &amp; Krikken" authorityYear="2013" box="[410,881,1619,1643]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="384" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="B297EA88F74A19160FA8F9C173E0F9F7" box="[410,654,1619,1643]" italics="true" pageId="3" pageNumber="384">Bolbochromus nomurai</emphasis>
<bibRefCitation id="E4724B6BF74A19160CAAF9C1721FF9F7" author="Krikken, J. &amp; Li, C. - L." box="[664,881,1619,1643]" pageId="3" pageNumber="384" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Li &amp; Krikken, 2013</bibRefCitation>
</taxonomicName>
. AB, male (Xishuangbanna, Yunnan); CD, female (Xishuangbanna, Yunnan); A, C, dorsal view; B, D, ventral view.
</paragraph>
</caption>
<caption id="D49C6612F74D19110EA5F95673EBF89C" pageId="4" pageNumber="385" startId="4.[151,250,1732,1756]" targetBox="[154,1416,201,1690]" targetPageId="4" targetType="figure">
<paragraph id="805C369AF74D19110EA5F95673EBF89C" blockId="4.[151,1436,1732,1792]" pageId="4" pageNumber="385">
<emphasis id="B297EA88F74D19110EA5F956707BF940" bold="true" box="[151,277,1732,1756]" pageId="4" pageNumber="385">FIGURE 3.</emphasis>
Habitus of
<taxonomicName id="47E34D19F74D19110FA6F9567233F940" authority="Krikken &amp; Li, 2013" authorityName="Krikken &amp; Li" authorityYear="2013" box="[404,861,1732,1756]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="385" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F74D19110FA6F95673EEF940" box="[404,640,1732,1756]" italics="true" pageId="4" pageNumber="385">Bolbochromus sinensis</emphasis>
<bibRefCitation id="E4724B6BF74D19110CBBF9567233F940" author="Krikken, J. &amp; Li, C. - L." box="[649,861,1732,1756]" pageId="4" pageNumber="385" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Krikken &amp; Li, 2013</bibRefCitation>
</taxonomicName>
. AB, male (Nanchang, Jiangxi); CD, female (Lishui, Zhejiang); A, C, dorsal view; B, D, ventral view.
</paragraph>
</caption>
<paragraph id="805C369AF74D19110EF5F8C074E2F8F0" blockId="4.[151,1437,1874,2009]" box="[199,1420,1874,1900]" pageId="4" pageNumber="385">
<emphasis id="B297EA88F74D19110EF5F8C07051F8F0" bold="true" box="[199,319,1874,1900]" pageId="4" pageNumber="385">
<typeStatus id="5F588838F74D19110EF5F8C07051F8F0" box="[199,319,1874,1900]" pageId="4" pageNumber="385" type="paratype">Paratypes</typeStatus>
</emphasis>
(females;
<figureCitation id="18D82A1FF74D19110F87F8C07357F8F0" box="[437,569,1874,1900]" captionStart="FIGURE 1" captionStartId="2.[151,250,1948,1972]" captionTargetBox="[162,1418,492,1901]" captionTargetId="figure-164@2.[151,1435,450,1922]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Bolbochromus sunfengyii Gao &amp; Yang, new species. AB, holotype; CD, female paratype; A, C, dorsal view; B, D, ventral view." pageId="4" pageNumber="385">Figs. 1CD</figureCitation>
,
<figureCitation id="18D82A1FF74D19110C76F8C07308F8F0" box="[580,614,1874,1900]" captionStart="FIGURE 4" captionStartId="6.[151,250,1908,1932]" captionTargetBox="[176,1407,595,1866]" captionTargetId="figure-216@6.[151,1435,561,1888]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Bolbochromus species in oblique view. AB, Bolbochromus sunfengyii Gao &amp; Yang, new species; A, holotype; B, female paratype; CD, Bolbochromus nomurai Li &amp; Krikken, 2013; C, male (Xishuangbanna, Yunnan); D, female (Xishuangbanna, Yunnan); EH, Bolbochromus sinensis Krikken &amp; Li, 2013; E, male (Nanchang, Jiangxi); H, female (Lishui, Zhejiang); not to scale." pageId="4" pageNumber="385">4B</figureCitation>
). General characters mainly stable, without distinct sexual dimorphism.
</paragraph>
<paragraph id="805C369AF74D19100EF5F8E47348FE81" blockId="4.[151,1437,1874,2009]" lastBlockId="5.[151,1437,151,897]" lastPageId="5" lastPageNumber="386" pageId="4" pageNumber="385">
<emphasis id="B297EA88F74D19110EF5F8E47024F80C" bold="true" box="[199,330,1910,1936]" pageId="4" pageNumber="385">Variations.</emphasis>
The black marks on the pronotum and elytra vary distinctly. On the pronotum, the anterior black mark varies from barely visible (
<figureCitation id="18D82A1FF74D19110FCCF808733AF828" box="[510,596,1946,1972]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="4" pageNumber="385">Fig. 6A</figureCitation>
) to a triangular area reaching the anterior third (
<figureCitation id="18D82A1FF74D19110A53F80875B8F828" box="[1121,1238,1946,1972]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="4" pageNumber="385">Fig. 6D, E</figureCitation>
), most commonly covering the anterior fifth (
<figureCitation id="18D82A1FF74D19110FF4F82C7373F844" box="[454,541,1982,2008]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="4" pageNumber="385">Fig. 6B</figureCitation>
); the posterior black mark varies from covering the posterior eighth (
<figureCitation id="18D82A1FF74D19110B12F82C7416F844" box="[1312,1400,1982,2008]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="4" pageNumber="385">Fig. 6A</figureCitation>
) to covering the posterior fourth (
<figureCitation id="18D82A1FF74C19100FD8FF0573EAFF2D" box="[490,644,151,177]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="5" pageNumber="386">Fig. 6C, D, E</figureCitation>
), most commonly covering the posterior fifth (
<figureCitation id="18D82A1FF74C19100AA7FF057580FF2D" box="[1173,1262,151,177]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="5" pageNumber="386">Fig. 6B</figureCitation>
). On the elytra, the black marks vary more dramatically, from only covering the sutural interstriae and regions near the basial margins (
<figureCitation id="18D82A1FF74C19100F33FF4D7035FF65" box="[257,347,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="5" pageNumber="386">Fig. 6A</figureCitation>
), to having a pair of black bands in interstriae 34 (
<figureCitation id="18D82A1FF74C19100D90FF4D7575FF65" box="[930,1051,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="5" pageNumber="386">Fig. 6B, C</figureCitation>
), in interstriae 24 and 610 (
<figureCitation id="18D82A1FF74C19100B5DFF4D71D3FE81" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="5" pageNumber="386">Fig. 6D</figureCitation>
), or across the middle (
<figureCitation id="18D82A1FF74C19100FF0FE917374FE81" box="[450,538,259,285]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="5" pageNumber="386">Fig. 6E</figureCitation>
).
</paragraph>
<paragraph id="805C369AF74C19100EF5FEB5738BFEF9" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EF5FEB57010FEDD" bold="true" box="[199,382,295,321]" pageId="5" pageNumber="386">Measurements.</emphasis>
Body length: male 8.0
<quantity id="471B9B7FF74C19100CA8FEB5739AFEDD" box="[666,756,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="5" pageNumber="386" unit="mm" value="9.6">9.6 mm</quantity>
(
<typeStatus id="5F588838F74C19100D37FEB57207FEDD" box="[773,873,295,321]" pageId="5" pageNumber="386" type="holotype">holotype</typeStatus>
<quantity id="471B9B7FF74C19100D46FEB572BFFEDD" box="[884,977,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="5" pageNumber="386" unit="mm" value="9.6">9.6 mm</quantity>
), female 7.0
<quantity id="471B9B7FF74C19100A40FEB575ACFEDD" box="[1138,1218,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="5" pageNumber="386" unit="mm" value="9.3">9.3mm</quantity>
; body width: male 5.0
<quantity id="471B9B7FF74C19100EFBFED87070FEF9" box="[201,286,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="5" pageNumber="386" unit="mm" value="5.9">5.9 mm</quantity>
(
<typeStatus id="5F588838F74C19100F1EFED970FEFEF9" box="[300,400,331,357]" pageId="5" pageNumber="386" type="holotype">holotype</typeStatus>
<quantity id="471B9B7FF74C19100FA5FED8709FFEF9" box="[407,497,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="5" pageNumber="386" unit="mm" value="5.6">5.6 mm</quantity>
), female
<quantity id="471B9B7FF74C19100C65FED8738FFEF9" box="[599,737,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.5" metricValueMin="4.4" pageId="5" pageNumber="386" unit="mm" value="4.95" valueMax="5.5" valueMin="4.4">4.45.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74C19100EF5FEFC728FFDFD" pageId="5" pageNumber="386" type="diagnosis">
<paragraph id="805C369AF74C19100EF5FEFC728FFDFD" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EF5FEFC702CFE14" bold="true" box="[199,322,366,392]" pageId="5" pageNumber="386">Diagnosis.</emphasis>
<taxonomicName id="47E34D19F74C19100F7AFEFD7398FE15" authority="Gao &amp; Yang" authorityName="Gao &amp; Yang" authorityYear="2025" box="[328,758,367,393]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="sunfengyii" status="sp. nov.">
<emphasis id="B297EA88F74C19100F7AFEFD730DFE15" box="[328,611,367,393]" italics="true" pageId="5" pageNumber="386">Bolbochromus sunfengyii</emphasis>
Gao &amp; Yang
</taxonomicName>
,
<emphasis id="B297EA88F74C19100D33FEFD72E7FE14" bold="true" box="[769,905,366,393]" pageId="5" pageNumber="386">
<taxonomicNameLabel id="A9A457F3F74C19100D33FEFD72E7FE14" box="[769,905,366,393]" pageId="5" pageNumber="386" rank="species">new species</taxonomicNameLabel>
</emphasis>
is related to
<taxonomicName id="47E34D19F74C19100A25FEFD75E1FE14" authorityName="Krikken &amp; Li" authorityYear="2013" box="[1047,1167,367,392]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F74C19100A25FEFD75E1FE14" box="[1047,1167,367,392]" italics="true" pageId="5" pageNumber="386">B. sinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="47E34D19F74C19100AF1FEFD7078FE31" authority="Krikken &amp; Li, 2013" authorityName="Krikken &amp; Li" authorityYear="2013" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B297EA88F74C19100AF1FEFD7455FE15" box="[1219,1339,367,393]" italics="true" pageId="5" pageNumber="386">B. pumilus</emphasis>
<bibRefCitation id="E4724B6BF74C19100B72FEFD7078FE31" author="Krikken, J. &amp; Li, C. - L." pageId="5" pageNumber="386" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Krikken &amp; Li, 2013</bibRefCitation>
</taxonomicName>
, but can be distinguished by: larger in body size (body length approximately
<quantity id="471B9B7FF74C19100A48FE0075BFFE31" box="[1146,1233,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="386" unit="mm" value="9.5">9.5 mm</quantity>
in
<taxonomicName id="47E34D19F74C19100AC6FE017403FE30" authorityName="Krikken &amp; Li" authorityYear="2013" box="[1268,1389,403,428]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F74C19100AC6FE017403FE30" box="[1268,1389,403,428]" italics="true" pageId="5" pageNumber="386">B. sinensis</emphasis>
</taxonomicName>
and 6.0 mm in
<taxonomicName id="47E34D19F74C19100F2AFE2570FCFE4D" authorityName="Krikken &amp; Li" authorityYear="2013" box="[280,402,439,465]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B297EA88F74C19100F2AFE2570FCFE4D" box="[280,402,439,465]" italics="true" pageId="5" pageNumber="386">B. pumilus</emphasis>
</taxonomicName>
); interstriae 2 and 4 less convex than adjacent interstriae (interstriae 2-3 less convex than adjacent interstriae in
<taxonomicName id="47E34D19F74C19100FBDFE497369FE68" authorityName="Krikken &amp; Li" authorityYear="2013" box="[399,519,475,500]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F74C19100FBDFE497369FE68" box="[399,519,475,500]" italics="true" pageId="5" pageNumber="386">B. sinensis</emphasis>
</taxonomicName>
; similar in
<taxonomicName id="47E34D19F74C19100CB5FE497268FE69" authorityName="Krikken &amp; Li" authorityYear="2013" box="[647,774,475,501]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="pumilus">B. pumilus</taxonomicName>
); elytron with 8 striae between suture and humeral umbone (7 striae in
<taxonomicName id="47E34D19F74C19100F20FE6D70E7FD84" authorityName="Krikken &amp; Li" authorityYear="2013" box="[274,393,511,536]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F74C19100F20FE6D70E7FD84" box="[274,393,511,536]" italics="true" pageId="5" pageNumber="386">B. sinensis</emphasis>
</taxonomicName>
; 5 striae in
<taxonomicName id="47E34D19F74C19100C3AFE6D7311FD85" authorityName="Krikken &amp; Li" authorityYear="2013" box="[520,639,511,537]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B297EA88F74C19100C3AFE6D7311FD85" box="[520,639,511,537]" italics="true" pageId="5" pageNumber="386">B. pumilus</emphasis>
</taxonomicName>
); elytral striae 2 and 5 not reaching base (similar in
<taxonomicName id="47E34D19F74C19100A88FE6D745FFD84" authorityName="Krikken &amp; Li" authorityYear="2013" box="[1210,1329,511,536]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F74C19100A88FE6D745FFD84" box="[1210,1329,511,536]" italics="true" pageId="5" pageNumber="386">B. sinensis</emphasis>
</taxonomicName>
; all striae reaching base in
<taxonomicName id="47E34D19F74C19100F60FDB170A7FDA1" authorityName="Krikken &amp; Li" authorityYear="2013" box="[338,457,547,573]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B297EA88F74C19100F60FDB170A7FDA1" box="[338,457,547,573]" italics="true" pageId="5" pageNumber="386">B. pumilus</emphasis>
</taxonomicName>
); male genitalia moderately sclerotized (strongly sclerotized, with median lobe aculeate in
<taxonomicName id="47E34D19F74C19100E86FDD57042FDFC" authorityName="Krikken &amp; Li" authorityYear="2013" box="[180,300,583,608]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F74C19100E86FDD57042FDFC" box="[180,300,583,608]" italics="true" pageId="5" pageNumber="386">B. sinensis</emphasis>
</taxonomicName>
; elongate-convex, parameres with rounded tip in
<taxonomicName id="47E34D19F74C19100D6AFDD572BFFDFD" authorityName="Krikken &amp; Li" authorityYear="2013" box="[856,977,583,609]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="pumilus">
<emphasis id="B297EA88F74C19100D6AFDD572BFFDFD" box="[856,977,583,609]" italics="true" pageId="5" pageNumber="386">B. pumilus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74C19100EF5FDF97341FD35" pageId="5" pageNumber="386" type="etymology">
<paragraph id="805C369AF74C19100EF5FDF97341FD35" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EF5FDF97020FD19" bold="true" box="[199,334,619,645]" pageId="5" pageNumber="386">Etymology.</emphasis>
This species is named in honor of Mr. Feng-Yi Sun, who kindly provided us most of the specimens of this new species for this research.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74C19100EF5FD20737BFD51" box="[199,533,690,717]" pageId="5" pageNumber="386" type="distribution">
<paragraph id="805C369AF74C19100EF5FD20737BFD51" blockId="5.[151,1437,151,897]" box="[199,533,690,717]" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EF5FD20700FFD50" bold="true" box="[199,353,690,716]" pageId="5" pageNumber="386">Distribution.</emphasis>
<collectingCountry id="F8F4760AF74C19100F55FD2170C0FD51" box="[359,430,691,717]" name="China" pageId="5" pageNumber="386">China</collectingCountry>
:
<collectingRegion id="4227F878F74C19100F8AFD21737FFD51" box="[440,529,691,717]" country="China" name="Yunnan" pageId="5" pageNumber="386">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74C19100EF5FD45727CFC1D" pageId="5" pageNumber="386" type="discussion">
<paragraph id="805C369AF74C19100EF5FD45727CFC1D" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EF5FD457055FD6D" bold="true" box="[199,315,727,753]" pageId="5" pageNumber="386">Remarks.</emphasis>
Additional to the series of specimens of this new species from Xishuangbanna,
<collectingRegion id="4227F878F74C19100A9EFD45746BFD6D" box="[1196,1285,727,753]" country="China" name="Yunnan" pageId="5" pageNumber="386">Yunnan</collectingRegion>
, a few clearly conspecific specimens from Dehong and Diqing Prefectures,
<collectingRegion id="4227F878F74C19100D0EFD6972FCFC89" box="[828,914,763,789]" country="China" name="Yunnan" pageId="5" pageNumber="386">Yunnan</collectingRegion>
have also been examined. The fauna of the two localities is distinctly different and rarely share common species, especially between Diqing and Xishaungbanna Prefectures. Therefore, the
<specimenCount id="96E5FD13F74C19100FF6FCD17307FCC1" box="[452,617,835,861]" count="2" pageId="5" pageNumber="386" type="generic">two specimens</specimenCount>
are not added to the
<typeStatus id="5F588838F74C19100D7EFCD17212FCC1" box="[844,892,835,861]" pageId="5" pageNumber="386">type</typeStatus>
series; and the distribution in these two localities should be confirmed with additional, reliable specimens.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,394 @@
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<mods:title id="8E5A0A75A54AE3F38D09F1E4F384A53D">Contributions to the genus Bolbochromus Boucomont, 1909 (Coleoptera: Geotrupidae: Bolboceratinae) from China, with the description of a new species</mods:title>
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<mods:affiliation id="20691E727BB2718C90FDA4E06CE5425E">Department of Life Sciences, Imperial College London, Exhibition Road, London SW 7 2 BX, United Kingdom. &amp; Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom.</mods:affiliation>
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<heading id="DB1481F6F74C19100EA5FC5F72EDFC7B" bold="true" box="[151,899,973,1000]" fontSize="11" level="1" pageId="5" pageNumber="386" reason="1">
<taxonomicName id="47E34D19F74C19100EA5FC5F72EDFC7B" ID-CoL="7SPCD" authority="Li &amp; Krikken, 2013" authorityName="Li &amp; Krikken" authorityYear="2013" box="[151,899,973,1000]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="nomurai" subGenus="Bolbochromus">
<emphasis id="B297EA88F74C19100EA5FC5F72EDFC7B" bold="true" box="[151,899,973,1000]" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EA5FC5F7027FC74" bold="true" box="[151,329,973,1000]" italics="true" pageId="5" pageNumber="386">Bolbochromus</emphasis>
(
<emphasis id="B297EA88F74C19100F68FC5F7362FC74" bold="true" box="[346,524,973,1000]" italics="true" pageId="5" pageNumber="386">Bolbochromus</emphasis>
)
<emphasis id="B297EA88F74C19100C2FFC5F73EBFC7B" bold="true" box="[541,645,973,999]" italics="true" pageId="5" pageNumber="386">nomurai</emphasis>
<bibRefCitation id="E4724B6BF74C19100CBEFC5F72EDFC7B" author="Krikken, J. &amp; Li, C. - L." box="[652,899,973,999]" pageId="5" pageNumber="386" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Li &amp; Krikken, 2013</bibRefCitation>
</emphasis>
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<paragraph id="805C369AF74C19100EA5FC63709EFB97" blockId="5.[151,899,973,1036]" box="[151,496,1009,1036]" pageId="5" pageNumber="386">
(
<figureCitation id="18D82A1FF74C19100EADFC637048FB97" box="[159,294,1009,1035]" captionStart="FIGURE 2" captionStartId="3.[151,250,1619,1643]" captionTargetBox="[168,1415,192,1567]" captionTargetId="figure-17@3.[151,1435,178,1595]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Bolbochromus nomurai Li &amp; Krikken, 2013. AB, male (Xishuangbanna, Yunnan); CD, female (Xishuangbanna, Yunnan); A, C, dorsal view; B, D, ventral view." pageId="5" pageNumber="386">Figs. 2AD</figureCitation>
,
<figureCitation id="18D82A1FF74C19100F03FC637019FB97" box="[305,375,1009,1035]" captionStart="FIGURE 4" captionStartId="6.[151,250,1908,1932]" captionTargetBox="[176,1407,595,1866]" captionTargetId="figure-216@6.[151,1435,561,1888]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Bolbochromus species in oblique view. AB, Bolbochromus sunfengyii Gao &amp; Yang, new species; A, holotype; B, female paratype; CD, Bolbochromus nomurai Li &amp; Krikken, 2013; C, male (Xishuangbanna, Yunnan); D, female (Xishuangbanna, Yunnan); EH, Bolbochromus sinensis Krikken &amp; Li, 2013; E, male (Nanchang, Jiangxi); H, female (Lishui, Zhejiang); not to scale." pageId="5" pageNumber="386">4CD</figureCitation>
,
<figureCitation id="18D82A1FF74C19100FB0FC6370D0FB90" box="[386,446,1009,1036]" captionStart="FIGURE 5" captionStartId="7.[151,250,1193,1217]" captionTargetBox="[161,1424,262,1154]" captionTargetId="figure-17@7.[157,1431,235,1170]" captionTargetPageId="7" captionText="FIGURE 5. Aedeagus of Bolbochromus species.AC, A*C*, Bolbochromus sunfengyii Gao &amp; Yang, new species (holotype), images denoted by an asterisk are enlarged version from AC; DF, Bolbochromus sinensis Krikken &amp; Li, 2013 (Nanchang, Jiangxi); GI, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan); A, A*, D, G, dorsal view; B, B*, E, H, lateral view; C, C*, F, I, ventral view; scale bar 0.2 mm." pageId="5" pageNumber="386">5GI</figureCitation>
,
<figureCitation id="18D82A1FF74C19100FF8FC637086FB97" box="[458,488,1009,1035]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="5" pageNumber="386">6F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74C19100EA5FBA47477FBD2" box="[151,1305,1078,1102]" pageId="5" pageNumber="386" type="reference_group">
<paragraph id="805C369AF74C19100EA5FBA47477FBD2" blockId="5.[151,1305,1078,1102]" box="[151,1305,1078,1102]" pageId="5" pageNumber="386">
<treatmentCitationGroup id="A0F311B4F74C19100EA5FBA47477FBD2" box="[151,1305,1078,1102]" pageId="5" pageNumber="386">
<treatmentCitation id="0142108BF74C19100EA5FBA4732BFBD2" author="Li, C. - L. &amp; Yang, P. - S. &amp; Krikken, J. &amp; Wang, C. - C." box="[151,581,1078,1102]" page="47" pageId="5" pageNumber="386" year="2013">
<taxonomicName id="47E34D19F74C19100EA5FBA4732BFBD2" ID-CoL="7SPCD" authority="Li et al., 2013: 47" authorityName="Li" authorityPageNumber="47" authorityYear="2013" box="[151,581,1078,1102]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="B297EA88F74C19100EA5FBA470E8FBD1" box="[151,390,1078,1102]" italics="true" pageId="5" pageNumber="386">Bolbochromus nomurai</emphasis>
<bibRefCitation id="E4724B6BF74C19100FBFFBA4732BFBD2" author="Li, C. - L. &amp; Yang, P. - S. &amp; Krikken, J. &amp; Wang, C. - C." box="[397,581,1078,1102]" pageId="5" pageNumber="386" pagination="39 - 54" refId="ref4838" refString="Li, C. - L., Yang, P. - S., Krikken, J. &amp; Wang, C. - C. (2013) Three new species of Bolbochromus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Southeast Asia. ZooKeys, 290, 39 - 54. https: // doi. org / 10.3897 / zookeys. 290.4696" type="journal article" year="2013">
Li
<emphasis id="B297EA88F74C19100F9BFBA570B3FBD2" box="[425,477,1078,1102]" italics="true" pageId="5" pageNumber="386">et al.</emphasis>
, 2013: 47
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0142108BF74C19100C7DFBA47226FBD2" author="Krikken, J. &amp; Li, C. - L." box="[591,840,1078,1102]" page="500" pageId="5" pageNumber="386" year="2013">
<bibRefCitation id="E4724B6BF74C19100C7DFBA47226FBD2" author="Krikken, J. &amp; Li, C. - L." box="[591,840,1078,1102]" pageId="5" pageNumber="386" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Krikken &amp; Li 2013: 500</bibRefCitation>
</treatmentCitation>
, table 1 (checklist);
<treatmentCitation id="0142108BF74C19100A27FBA4747AFBD2" author="Nikolajev, G. V. &amp; Kral, D." box="[1045,1300,1078,1102]" page="35" pageId="5" pageNumber="386" year="2016">
<bibRefCitation id="E4724B6BF74C19100A27FBA4747AFBD2" author="Nikolajev, G. V. &amp; Kral, D." box="[1045,1300,1078,1102]" pageId="5" pageNumber="386" pagination="33 - 36" refId="ref4906" refString="Nikolajev, G. V., Kral, D. &amp; Bezdӗk. A. (2016) Bolboceratinae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera, volume 3. Koninklijke Brill NV, Leiden, pp. 33 - 36. https: // doi. org / 10.1163 / 9789004309142" type="book chapter" year="2016">
Nikolajev
<emphasis id="B297EA88F74C19100A4DFBA575C2FBD2" box="[1151,1196,1078,1102]" italics="true" pageId="5" pageNumber="386">et al</emphasis>
. 2016: 35
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74C19100EA5FBEE7358FA0E" pageId="5" pageNumber="386" type="materials_examined">
<paragraph id="805C369AF74C19100EA5FBEE7519FB0B" blockId="5.[151,1437,1148,2039]" box="[151,1143,1148,1175]" pageId="5" pageNumber="386">
<materialsCitation id="308B3CC7F74C19100EA5FBEE751DFB0B" box="[151,1139,1148,1175]" country="Vietnam" location="Deo Pha Din" longitude="103.833336" municipality="Northern" pageId="5" pageNumber="386" specimenCount="1" stateProvince="Son La" typeStatus="holotype">
<emphasis id="B297EA88F74C19100EA5FBEE7058FB0A" bold="true" box="[151,310,1148,1174]" pageId="5" pageNumber="386">
<typeStatus id="5F588838F74C19100EA5FBEE71BCFB0A" box="[151,210,1148,1174]" pageId="5" pageNumber="386">Type</typeStatus>
locality.
</emphasis>
<collectingMunicipality id="6038ACE0F74C19100F74FBEE70C0FB0A" box="[326,430,1148,1174]" pageId="5" pageNumber="386">Northern</collectingMunicipality>
<collectingCountry id="F8F4760AF74C19100F86FBEE7379FB0A" box="[436,535,1148,1174]" name="Vietnam" pageId="5" pageNumber="386">Vietnam</collectingCountry>
:
<collectingRegion id="4227F878F74C19100C10FBEE73B1FB0A" box="[546,735,1148,1175]" country="Vietnam" name="Son La" pageId="5" pageNumber="386">Son La Province</collectingRegion>
,
<location id="853C6041F74C19100CD9FBEF7210FB0A" LSID="urn:lsid:plazi:treatment:084A878CF74C19120EA5FC5F73F6F8F2:853C6041F74C19100CD9FBEF7210FB0A" box="[747,894,1148,1175]" country="Vietnam" longitude="103.833336" municipality="Northern" name="Deo Pha Din" pageId="5" pageNumber="386" stateProvince="Son La">Deo Pha Din</location>
, 21°70N,
<geoCoordinate id="E5D7505DF74C19100DCAFBEE7506FB0B" box="[1016,1128,1148,1175]" degrees="103" direction="east" minutes="50" orientation="longitude" pageId="5" pageNumber="386" precision="925" value="103.833336">103°50E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="805C369AF74C19100EF5FB327354FB27" blockId="5.[151,1437,1148,2039]" box="[199,570,1184,1211]" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EF5FB327081FB26" bold="true" box="[199,495,1184,1210]" pageId="5" pageNumber="386">
<typeStatus id="5F588838F74C19100EF5FB32706CFB26" box="[199,258,1184,1210]" pageId="5" pageNumber="386">Type</typeStatus>
material examined.
</emphasis>
None.
</paragraph>
<paragraph id="805C369AF74C19100EF5FB567358FA0E" blockId="5.[151,1437,1148,2039]" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EF5FB56735FFB42" bold="true" box="[199,561,1220,1246]" pageId="5" pageNumber="386">Additional material examined</emphasis>
(
<specimenCount id="96E5FD13F74C19100C71FB567310FB42" box="[579,638,1220,1246]" count="6" pageId="5" pageNumber="386" type="male">6♂♂</specimenCount>
,
<specimenCount id="96E5FD13F74C19100CBEFB5673A8FB42" box="[652,710,1220,1246]" count="2" pageId="5" pageNumber="386" type="female">2♀♀</specimenCount>
, 2 unsexed specimens).
<materialsCitation id="308B3CC7F74C19100DEDFB567257FA9E" collectingDate="2020-09" collectionCode="CFYS" country="China" location="Gasa Town" pageId="5" pageNumber="386" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="96E5FD13F74C19100DEDFB567577FB42" box="[991,1049,1220,1246]" count="3" pageId="5" pageNumber="386" type="male">3♂♂</specimenCount>
,
<specimenCount id="96E5FD13F74C19100A15FB56750EFB42" box="[1063,1120,1220,1246]" count="2" pageId="5" pageNumber="386" type="female">2♀♀</specimenCount>
(
<collectionCode id="E6F2AE5FF74C19100A43FB5675D2FB42" box="[1137,1212,1220,1246]" pageId="5" pageNumber="386">CFYS</collectionCode>
),
<location id="853C6041F74C19100AE0FB56743FFB43" LSID="urn:lsid:plazi:treatment:084A878CF74C19120EA5FC5F73F6F8F2:853C6041F74C19100AE0FB56743FFB43" box="[1234,1361,1220,1247]" country="China" name="Gasa Town" pageId="5" pageNumber="386" stateProvince="Yunnan">Gasa Town</location>
(Ŕ洒 Dz),
<collectingRegion id="4227F878F74C19100EFFFB7B70FDFA9E" box="[205,403,1256,1283]" country="China" name="Yunnan" pageId="5" pageNumber="386">Yunnan Province</collectingRegion>
,
<collectingCountry id="F8F4760AF74C19100F90FB7A7086FA9E" box="[418,488,1256,1282]" name="China" pageId="5" pageNumber="386">China</collectingCountry>
,
<date id="F45D105AF74C19100FC4FB7A733BFA9E" box="[502,597,1256,1282]" pageId="5" pageNumber="386" value="2020-09">
<collectingDate id="E419E9B2F74C19100FC4FB7A733BFA9E" box="[502,597,1256,1282]" pageId="5" pageNumber="386" value="2020-09">IX.2020</collectingDate>
</date>
, local collector leg
</materialsCitation>
.;
<materialsCitation id="308B3CC7F74C19100D7FFB7A7237FABA" collectingDate="2020-05" collectionCode="CQZY" country="China" location="Gasa Town" municipality="Xishuangbanna Prefecture" pageId="5" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="96E5FD13F74C19100D7FFB7A721EFA9E" box="[845,880,1256,1282]" count="1" pageId="5" pageNumber="386" type="male">1♂</specimenCount>
(
<collectionCode id="E6F2AE5FF74C19100DB0FB7A72BAFA9E" box="[898,980,1256,1282]" pageId="5" pageNumber="386">CQZY</collectionCode>
),
<location id="853C6041F74C19100DD8FB7A7505FA9F" LSID="urn:lsid:plazi:treatment:084A878CF74C19120EA5FC5F73F6F8F2:853C6041F74C19100DD8FB7A7505FA9F" box="[1002,1131,1256,1283]" country="China" municipality="Xishuangbanna Prefecture" name="Gasa Town" pageId="5" pageNumber="386" stateProvince="Yunnan">Gasa Town</location>
(Ŕ洒Dz),
<collectingMunicipality id="6038ACE0F74C19100ADCFB7A7063FABA" pageId="5" pageNumber="386">Xishuangbanna Prefecture</collectingMunicipality>
,
<collectingRegion id="4227F878F74C19100F24FA9F70B9FABA" box="[278,471,1292,1319]" country="China" name="Yunnan" pageId="5" pageNumber="386">Yunnan Province</collectingRegion>
,
<collectingCountry id="F8F4760AF74C19100FD0FA9E7346FABA" box="[482,552,1292,1318]" name="China" pageId="5" pageNumber="386">China</collectingCountry>
,
<date id="F45D105AF74C19100C03FA9E73EAFABA" box="[561,644,1292,1318]" pageId="5" pageNumber="386" value="2020-05">
<collectingDate id="E419E9B2F74C19100C03FA9E73EAFABA" box="[561,644,1292,1318]" pageId="5" pageNumber="386" value="2020-05">V.2020</collectingDate>
</date>
, local collector leg
</materialsCitation>
.;
<materialsCitation id="308B3CC7F74C19100D5AFA9E7209FAD6" collectingDate="2022-07" collectionCode="CHG" country="China" location="Puwen Town" municipality="Xishuangbanna Prefecture" pageId="5" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="96E5FD13F74C19100D5AFA9E72E5FABA" box="[872,907,1292,1318]" count="1" pageId="5" pageNumber="386" type="male">1♂</specimenCount>
(
<collectionCode id="E6F2AE5FF74C19100DAAFA9E72B8FABA" box="[920,982,1292,1318]" pageId="5" pageNumber="386">CHG</collectionCode>
)
<location id="853C6041F74C19100DD6FA9F751AFABB" LSID="urn:lsid:plazi:treatment:084A878CF74C19120EA5FC5F73F6F8F2:853C6041F74C19100DD6FA9F751AFABB" box="[996,1140,1293,1319]" country="China" municipality="Xishuangbanna Prefecture" name="Puwen Town" pageId="5" pageNumber="386" stateProvince="Yunnan">Puwen Town</location>
(üĸDz),
<collectingMunicipality id="6038ACE0F74C19100ADCFA9E7063FAD6" pageId="5" pageNumber="386">Xishuangbanna Prefecture</collectingMunicipality>
,
<collectingRegion id="4227F878F74C19100F27FAA370BBFAD6" box="[277,469,1328,1355]" country="China" name="Yunnan" pageId="5" pageNumber="386">Yunnan Province</collectingRegion>
,
<collectingCountry id="F8F4760AF74C19100FECFAA2734AFAD6" box="[478,548,1328,1354]" name="China" pageId="5" pageNumber="386">China</collectingCountry>
,
<date id="F45D105AF74C19100C1EFAA273FAFAD6" box="[556,660,1328,1354]" pageId="5" pageNumber="386" value="2022-07">
<collectingDate id="E419E9B2F74C19100C1EFAA273FAFAD6" box="[556,660,1328,1354]" pageId="5" pageNumber="386" value="2022-07">VII.2022</collectingDate>
</date>
, local collector leg
</materialsCitation>
.;
<materialsCitation id="308B3CC7F74C19100D46FAA273E0FAF2" collectingDate="2019-08" collectionCode="CBHH" country="China" location="Diqing Prefecture" pageId="5" pageNumber="386" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="96E5FD13F74C19100D46FAA272F9FAD6" box="[884,919,1328,1354]" count="1" pageId="5" pageNumber="386" type="male">1♂</specimenCount>
(
<collectionCode id="E6F2AE5FF74C19100D96FAA27299FAD6" box="[932,1015,1328,1354]" pageId="5" pageNumber="386">CBHH</collectionCode>
),
<location id="853C6041F74C19100A3AFAA275BEFAD6" LSID="urn:lsid:plazi:treatment:084A878CF74C19120EA5FC5F73F6F8F2:853C6041F74C19100A3AFAA275BEFAD6" box="[1032,1232,1328,1354]" country="China" name="Diqing Prefecture" pageId="5" pageNumber="386" stateProvince="Yunnan">Diqing Prefecture</location>
,
<collectingRegion id="4227F878F74C19100AEAFAA374F6FAD6" box="[1240,1432,1328,1355]" country="China" name="Yunnan" pageId="5" pageNumber="386">Yunnan Province</collectingRegion>
,
<collectingCountry id="F8F4760AF74C19100EA5FAC671B3FAF2" box="[151,221,1364,1390]" name="China" pageId="5" pageNumber="386">China</collectingCountry>
,
<collectingMethod id="59A24E8DF74C19100EDAFAC67057FAF3" box="[232,313,1364,1391]" pageId="5" pageNumber="386">by trap</collectingMethod>
,
<date id="F45D105AF74C19100F76FAC670DBFAF2" box="[324,437,1364,1390]" pageId="5" pageNumber="386" value="2019-08">
<collectingDate id="E419E9B2F74C19100F76FAC670DBFAF2" box="[324,437,1364,1390]" pageId="5" pageNumber="386" value="2019-08">VIII.2019</collectingDate>
</date>
, local collector leg
</materialsCitation>
.;
<materialsCitation id="308B3CC7F74C19100CACFAC6735CFA0E" collectingDate="2020-05" collectionCode="CBHH" country="China" location="Mangyun" municipality="Dehong Prefecture" pageId="5" pageNumber="386" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="96E5FD13F74C19100CACFAC67246FAF2" box="[670,808,1364,1390]" count="2" pageId="5" pageNumber="386" type="generic">2 specimens</specimenCount>
(
<collectionCode id="E6F2AE5FF74C19100D05FAC672E4FAF2" box="[823,906,1364,1390]" pageId="5" pageNumber="386">CBHH</collectionCode>
),
<location id="853C6041F74C19100DAFFAC77566FAF3" LSID="urn:lsid:plazi:treatment:084A878CF74C19120EA5FC5F73F6F8F2:853C6041F74C19100DAFFAC77566FAF3" box="[925,1032,1365,1391]" country="China" municipality="Dehong Prefecture" name="Mangyun" pageId="5" pageNumber="386" stateProvince="Yunnan">Mangyun</location>
(Ệ允),
<collectingMunicipality id="6038ACE0F74C19100A55FAC67453FAF2" box="[1127,1341,1364,1390]" pageId="5" pageNumber="386">Dehong Prefecture</collectingMunicipality>
,
<collectingRegion id="4227F878F74C19100B75FAC77193FA0E" country="China" name="Yunnan" pageId="5" pageNumber="386">Yunnan Province</collectingRegion>
,
<date id="F45D105AF74C19100F3AFAEA7035FA0E" box="[264,347,1400,1426]" pageId="5" pageNumber="386" value="2020-05">
<collectingDate id="E419E9B2F74C19100F3AFAEA7035FA0E" box="[264,347,1400,1426]" pageId="5" pageNumber="386" value="2020-05">V.2020</collectingDate>
</date>
, local collector leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74C19120EF5FA0E704BF90A" lastPageId="7" lastPageNumber="388" pageId="5" pageNumber="386" type="description">
<paragraph id="805C369AF74C19130EF5FA0E7577FD85" blockId="5.[151,1437,1148,2039]" lastBlockId="6.[151,1437,150,537]" lastPageId="6" lastPageNumber="387" pageId="5" pageNumber="386">
<emphasis id="B297EA88F74C19100EF5FA0E731BFA2A" bold="true" box="[199,629,1436,1462]" pageId="5" pageNumber="386">Description of the females. General</emphasis>
(
<figureCitation id="18D82A1FF74C19100CBAFA0E7261FA2A" box="[648,783,1436,1462]" captionStart="FIGURE 2" captionStartId="3.[151,250,1619,1643]" captionTargetBox="[168,1415,192,1567]" captionTargetId="figure-17@3.[151,1435,178,1595]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Bolbochromus nomurai Li &amp; Krikken, 2013. AB, male (Xishuangbanna, Yunnan); CD, female (Xishuangbanna, Yunnan); A, C, dorsal view; B, D, ventral view." pageId="5" pageNumber="386">Figs. 2CD</figureCitation>
,
<figureCitation id="18D82A1FF74C19100D2FFA0E722EFA2A" box="[797,832,1436,1462]" captionStart="FIGURE 4" captionStartId="6.[151,250,1908,1932]" captionTargetBox="[176,1407,595,1866]" captionTargetId="figure-216@6.[151,1435,561,1888]" captionTargetPageId="6" captionText="FIGURE 4. Habitus of Bolbochromus species in oblique view. AB, Bolbochromus sunfengyii Gao &amp; Yang, new species; A, holotype; B, female paratype; CD, Bolbochromus nomurai Li &amp; Krikken, 2013; C, male (Xishuangbanna, Yunnan); D, female (Xishuangbanna, Yunnan); EH, Bolbochromus sinensis Krikken &amp; Li, 2013; E, male (Nanchang, Jiangxi); H, female (Lishui, Zhejiang); not to scale." pageId="5" pageNumber="386">4D</figureCitation>
): Length
<quantity id="471B9B7FF74C19100D86FA0E7551FA2B" box="[948,1087,1436,1463]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.8" metricValueMin="6.7" pageId="5" pageNumber="386" unit="mm" value="7.25" valueMax="7.8" valueMin="6.7">6.77.8 mm</quantity>
, maximum width at pronotum posterior to middle (3.94.0 mm). Body convex, dorsum glossy.
<emphasis id="B297EA88F74C19100D50FA5272DEFA46" bold="true" box="[866,944,1472,1498]" pageId="5" pageNumber="386">Color:</emphasis>
Head black. Middle of disk more testaceous than the rest of pronotum. Scutellum testaceous. Elytra bicolored testaceous and black. Sutural interstria dark brown along entire length; all striae black.
<emphasis id="B297EA88F74C19100C1EF99A731BF9BE" bold="true" box="[556,629,1544,1570]" pageId="5" pageNumber="386">Head:</emphasis>
Dorsal surface covered with coarse punctures, glabrous. Vertex of head with faint, transverse tubercle. Frons along inner margins of eyes barely raised. Clypeus broadly arcuate anteriorly; middle of anterior margin with low tubercle; margin of ocular canthus evenly arcuate; ocular canthus completely bisecting eyes. Antenna testaceous, with 11 antennomeres, antennomere 1 longer than any of 28, antennomere 2 elongated and thick, antennomeres 38 gradually thickening, antennomeres 911 elliptically swollen, inner face of antennomere 9 with glabrous plate.
<emphasis id="B297EA88F74C19100C61F92E73B8F94A" bold="true" box="[595,726,1724,1750]" pageId="5" pageNumber="386">Pronotum:</emphasis>
Each lateral third with scattered large punctures, central third impunctate except anteriorly; smaller secondary punctures distributed along lateral margins. Disc with distinctly impressed, median longitudinal groove terminating before reaching anterior and posterior margins, entire length with sparse punctures. Anterior triangular flat area with small punctures in front of mid-longitudinal impression. Lateral foveae barely impressed, with several punctures. Anterior angles obtuse. Anterior and lateral margins with border terminating on basal margin at level of humeral umbone of elytra; lateral margins fringed with setae underneath, visible from above.
<emphasis id="B297EA88F74C19100FB5F8067366F832" bold="true" box="[391,520,1940,1966]" pageId="5" pageNumber="386">Scutellum:</emphasis>
Shiny, with extremely fine, nearly invisible punctures along anterior margin. Distinct border on both sides relative to the elytra.
<emphasis id="B297EA88F74C19100CFCF82A724DF84E" bold="true" box="[718,803,1976,2002]" pageId="5" pageNumber="386">Elytra:</emphasis>
Each elytron with 13 punctate striae, all evenly punctate along their lengths; with 7 striae between suture and humeral umbone. Sutural stria runs parallel to scutellum and terminates at elytral base, interrupting stria 2 before elytral base; stria 5 ending before elytral base; striae 8 and 9 terminating on humeral umbone, not reaching elytral base; stria 10 extending around and below humeral umbone, longer than striae 8 and 9, not reaching elytral base; stria 11 terminating at elytral mid-length, well before elytral base; stria 12 reaching elytral base. Stria 13 with very deep punctures, situated in reflexed lateral margin of elytron, reaching elytral base. Reflexed lateral margin of elytra with setae of two lengths, long and short setae irregularly alternate, sparsely and evenly scattered between punctures of stria 13, long setae largely located at anterior, short setae more numerous than long setae. Elytral interstriae slightly convex; surface smooth, micropunctures almost invisible. Elytral apices acuminate. Ventral face of epipleuron black, with single row of setae.
<emphasis id="B297EA88F74F19130AF3FE01746CFE31" bold="true" box="[1217,1282,403,429]" pageId="6" pageNumber="387">Legs:</emphasis>
Protibia with 9 teeth along outer margin, gradually increasing in size from base to apex, with distal 3 teeth much larger than preceding. Apical mesotibial spurs adjacent to each other, unequal in length, with inner spur longer than outer spur; apical metatibial spurs subequal in length, spaced apart by basal width of a spur.
</paragraph>
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<paragraph id="805C369AF74F19130EA5F8E670ECF864" blockId="6.[151,1436,1908,2040]" pageId="6" pageNumber="387">
<emphasis id="B297EA88F74F19130EA5F8E6707AF810" bold="true" box="[151,276,1908,1932]" pageId="6" pageNumber="387">FIGURE 4.</emphasis>
Habitus of
<taxonomicName id="47E34D19F74F19130FBCF8E6731CF810" box="[398,626,1908,1932]" pageId="6" pageNumber="387">
<emphasis id="B297EA88F74F19130FBCF8E6734FF810" box="[398,545,1908,1932]" italics="true" pageId="6" pageNumber="387">Bolbochromus</emphasis>
species
</taxonomicName>
in oblique view. AB,
<taxonomicName id="47E34D19F74F19130D53F8E67580F811" authority="Gao &amp; Yang" authorityName="Gao &amp; Yang" authorityYear="2025" box="[865,1262,1908,1933]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="sunfengyii" status="sp. nov.">
<emphasis id="B297EA88F74F19130D53F8E6750DF811" box="[865,1123,1908,1933]" italics="true" pageId="6" pageNumber="387">Bolbochromus sunfengyii</emphasis>
Gao &amp; Yang
</taxonomicName>
,
<emphasis id="B297EA88F74F19130ACBF8E77419F810" bold="true" box="[1273,1399,1908,1933]" pageId="6" pageNumber="387">
<taxonomicNameLabel id="A9A457F3F74F19130ACBF8E77419F810" box="[1273,1399,1908,1933]" pageId="6" pageNumber="387" rank="species">new species</taxonomicNameLabel>
</emphasis>
; A, holotype; B, female paratype; CD,
<taxonomicName id="47E34D19F74F19130C34F80A72A8F82D" authority="Li &amp; Krikken, 2013" authorityName="Li &amp; Krikken" authorityYear="2013" box="[518,966,1944,1969]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="B297EA88F74F19130C34F80A739AF82C" box="[518,756,1944,1968]" italics="true" pageId="6" pageNumber="387">Bolbochromus nomurai</emphasis>
<bibRefCitation id="E4724B6BF74F19130CC8F80A72A8F82D" author="Krikken, J. &amp; Li, C. - L." box="[762,966,1944,1969]" pageId="6" pageNumber="387" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Li &amp; Krikken, 2013</bibRefCitation>
</taxonomicName>
; C, male (Xishuangbanna, Yunnan); D, female (Xishuangbanna, Yunnan); EH,
<taxonomicName id="47E34D19F74F19130FDBF82E72C9F849" authority="Krikken &amp; Li, 2013" authorityName="Krikken &amp; Li" authorityYear="2013" box="[489,935,1980,2005]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="387" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B297EA88F74F19130FDBF82E73BCF848" box="[489,722,1980,2004]" italics="true" pageId="6" pageNumber="387">Bolbochromus sinensis</emphasis>
<bibRefCitation id="E4724B6BF74F19130CEBF82E72C9F849" author="Krikken, J. &amp; Li, C. - L." box="[729,935,1980,2005]" pageId="6" pageNumber="387" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Krikken &amp; Li, 2013</bibRefCitation>
</taxonomicName>
; E, male (Nanchang, Jiangxi); H, female (Lishui, Zhejiang); not to scale.
</paragraph>
</caption>
<caption id="D49C6612F74E19120EA5FB3B73BFFAB1" pageId="7" pageNumber="388" startId="7.[151,250,1193,1217]" targetBox="[161,1424,262,1154]" targetPageId="7" targetType="figure">
<paragraph id="805C369AF74E19120EA5FB3B73BFFAB1" blockId="7.[151,1436,1193,1326]" pageId="7" pageNumber="388">
<emphasis id="B297EA88F74E19120EA5FB3B707DFB5E" bold="true" box="[151,275,1193,1218]" pageId="7" pageNumber="388">FIGURE 5.</emphasis>
Aedeagus of
<taxonomicName id="47E34D19F74E19120FAEFB3B73F2FB5E" box="[412,668,1193,1218]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B297EA88F74E19120FAEFB3B7341FB5D" box="[412,559,1193,1217]" italics="true" pageId="7" pageNumber="388">Bolbochromus</emphasis>
species. A
</taxonomicName>
C, A*C*,
<taxonomicName id="47E34D19F74E19120D28FB3B75CEFB5E" authority="Gao &amp; Yang" authorityName="Gao &amp; Yang" authorityYear="2025" box="[794,1184,1193,1218]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="sunfengyii" status="sp. nov.">
<emphasis id="B297EA88F74E19120D28FB3B7574FB5D" box="[794,1050,1193,1217]" italics="true" pageId="7" pageNumber="388">Bolbochromus sunfengyii</emphasis>
Gao &amp; Yang
</taxonomicName>
,
<emphasis id="B297EA88F74E19120A98FB387449FB5D" bold="true" box="[1194,1319,1193,1218]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A9A457F3F74E19120A98FB387449FB5D" box="[1194,1319,1193,1218]" pageId="7" pageNumber="388" rank="species">new species</taxonomicNameLabel>
</emphasis>
(holotype), images denoted by an asterisk are enlarged version from AC; DF,
<taxonomicName id="47E34D19F74E19120D6EFB5F7471FB7A" authority="Krikken &amp; Li, 2013" authorityName="Krikken &amp; Li" authorityYear="2013" box="[860,1311,1229,1254]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="sinensis">
Bolbochromus sinensis
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</taxonomicName>
(Nanchang, Jiangxi); GI,
<taxonomicName id="47E34D19F74E19120F1EFB63739EFA96" authority="Li &amp; Krikken, 2013" authorityName="Li &amp; Krikken" authorityYear="2013" box="[300,752,1265,1290]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="B297EA88F74E19120F1EFB637372FA95" box="[300,540,1265,1289]" italics="true" pageId="7" pageNumber="388">Bolbochromus nomurai</emphasis>
<bibRefCitation id="E4724B6BF74E19120C16FB63739EFA96" author="Krikken, J. &amp; Li, C. - L." box="[548,752,1265,1290]" pageId="7" pageNumber="388" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Li &amp; Krikken, 2013</bibRefCitation>
</taxonomicName>
(Xishuangbanna, Yunnan); A, A*, D, G, dorsal view; B, B*, E, H, lateral view; C, C*, F, I, ventral view; scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="805C369AF74E19120EF5FAED722BFA22" blockId="7.[151,1437,1407,1902]" pageId="7" pageNumber="388">
<emphasis id="B297EA88F74E19120EF5FAED7051FA05" bold="true" box="[199,319,1407,1433]" pageId="7" pageNumber="388">Variation.</emphasis>
The brownish-orange marks on the male elytra varies distinctly, which vary from absent (
<figureCitation id="18D82A1FF74E19120B14FA127416FA05" box="[1318,1400,1407,1434]" captionStart="FIGURE 6" captionStartId="8.[151,250,1582,1606]" captionTargetBox="[164,1419,191,1550]" captionTargetId="figure-23@8.[151,1435,178,1562]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Bolbochromus species. demonstrating color variations on the dorsal surface in dorsal view. AE, Bolbochromus sunfengyii Gao &amp; Yang, new species (Xishuangbanna, Yunnan); F, Bolbochromus nomurai Li &amp; Krikken, 2013 (Xishuangbanna, Yunnan)." pageId="7" pageNumber="388">Fig. 6F</figureCitation>
) to having a small brownish-orange band in striae 26 (
<figureCitation id="18D82A1FF74E19120CEFFA367256FA22" box="[733,824,1444,1470]" captionStart="FIGURE 2" captionStartId="3.[151,250,1619,1643]" captionTargetBox="[168,1415,192,1567]" captionTargetId="figure-17@3.[151,1435,178,1595]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of Bolbochromus nomurai Li &amp; Krikken, 2013. AB, male (Xishuangbanna, Yunnan); CD, female (Xishuangbanna, Yunnan); A, C, dorsal view; B, D, ventral view." pageId="7" pageNumber="388">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="805C369AF74E19120EF5FA55758AF9D2" blockId="7.[151,1437,1407,1902]" pageId="7" pageNumber="388">
<emphasis id="B297EA88F74E19120EF5FA5570D9FA7D" bold="true" box="[199,439,1479,1505]" pageId="7" pageNumber="388">Sexual dimorphism.</emphasis>
The two sexes show distinct sexual dimorphism in color pattern on the body. Males have a black body and brownish-orange marks on both sides of the pronotum, while the elytra are with or without oval brownish-orange marks near the base. The female body is generally testaceous, with small black marks near the margins of the pronotum, while the elytra have black or brown marks near the margins and suture.
</paragraph>
<paragraph id="805C369AF74E19120EF5F9CA704BF90A" blockId="7.[151,1437,1407,1902]" pageId="7" pageNumber="388">
<emphasis id="B297EA88F74E19120EF5F9CA7010F9EE" bold="true" box="[199,382,1624,1650]" pageId="7" pageNumber="388">Measurements.</emphasis>
Body length: male
<quantity id="471B9B7FF74E19120C53F9CA7382F9EE" box="[609,748,1624,1650]" metricMagnitude="-3" metricUnit="m" metricValue="7.6499999999999995" metricValueMax="8.2" metricValueMin="7.1" pageId="7" pageNumber="388" unit="mm" value="7.6499999999999995" valueMax="8.2" valueMin="7.1">7.18.2 mm</quantity>
, female
<quantity id="471B9B7FF74E19120D7DF9C572B4F9EE" box="[847,986,1623,1650]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" metricValueMax="7.7" metricValueMin="6.7" pageId="7" pageNumber="388" unit="mm" value="7.2" valueMax="7.7" valueMin="6.7">6.77.7 mm</quantity>
; body width: male 4.0
<quantity id="471B9B7FF74E19120ADBF9CA742DF9EE" box="[1257,1347,1624,1650]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="388" unit="mm" value="4.9">4.9 mm</quantity>
, female
<quantity id="471B9B7FF74E19120EA5F9EE704FF90A" box="[151,289,1660,1686]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="7" pageNumber="388" unit="mm" value="4.05" valueMax="4.3" valueMin="3.8">3.84.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74E19120EF5F90D73ECF926" box="[199,642,1695,1722]" pageId="7" pageNumber="388" type="distribution">
<paragraph id="805C369AF74E19120EF5F90D73ECF926" blockId="7.[151,1437,1407,1902]" box="[199,642,1695,1722]" pageId="7" pageNumber="388">
<emphasis id="B297EA88F74E19120EF5F90D700FF925" bold="true" box="[199,353,1695,1721]" pageId="7" pageNumber="388">Distribution.</emphasis>
<collectingCountry id="F8F4760AF74E19120F55F93270C0F926" box="[359,430,1696,1722]" name="China" pageId="7" pageNumber="388">China</collectingCountry>
:
<collectingRegion id="4227F878F74E19120F8AF932737FF926" box="[440,529,1696,1722]" country="China" name="Yunnan" pageId="7" pageNumber="388">Yunnan</collectingRegion>
;
<collectingCountry id="F8F4760AF74E19120C2EF9327311F926" box="[540,639,1696,1722]" name="Vietnam" pageId="7" pageNumber="388">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8F96511F74E19120EF5F95673F6F8F2" pageId="7" pageNumber="388" type="discussion">
<paragraph id="805C369AF74E19120EF5F95673F6F8F2" blockId="7.[151,1437,1407,1902]" pageId="7" pageNumber="388">
<emphasis id="B297EA88F74E19120EF5F9567055F942" bold="true" box="[199,315,1732,1758]" pageId="7" pageNumber="388">Remarks.</emphasis>
This species is the only species possessing sexual dimorphism in this study, although such phenomenon is seen in some other congeneric species, such as
<taxonomicName id="47E34D19F74E19120CFFF97A759EF89E" authority="Krikken &amp; Li, 2013" authorityName="Krikken &amp; Li" authorityYear="2013" box="[717,1264,1768,1794]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="mindanaicus">
<emphasis id="B297EA88F74E19120CFFF97A7568F89E" box="[717,1030,1768,1794]" italics="true" pageId="7" pageNumber="388">Bolbochromus mindanaicus</emphasis>
<bibRefCitation id="E4724B6BF74E19120A3CF97A759EF89E" author="Krikken, J. &amp; Li, C. - L." box="[1038,1264,1768,1794]" pageId="7" pageNumber="388" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Krikken &amp; Li, 2013</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E4724B6BF74E19120B32F97A71BCF8BA" author="Krikken, J. &amp; Li, C. - L." pageId="7" pageNumber="388" pagination="495 - 519" refId="ref4559" refString="Krikken, J. &amp; Li, C. - L. (2013) Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae). Zootaxa, 3731 (4), 495 - 519. https: // doi. org / 10.11646 / zootaxa. 3731.4.4" type="journal article" year="2013">Krikken &amp; Li 2013</bibRefCitation>
;
<bibRefCitation id="E4724B6BF74E19120ED2F89E70EFF8BA" author="Li, C. - L. &amp; Krikken, J. &amp; Wang, C. - C." box="[224,385,1804,1830]" pageId="7" pageNumber="388" pagination="135 - 152" refId="ref4672" refString="Li, C. - L., Krikken, J. &amp; Wang, C. - C. (2019) Review of the genus Bolbochromus (Coleoptera, Scarabaeoidea, Geotrupidae, Bolboceratinae) in the Philippines. ZooKeys, 842, 135 - 152. https: // doi. org / 10.3897 / zookeys. 842.32315" type="journal article" year="2019">
Li
<emphasis id="B297EA88F74E19120F30F89E705BF8BA" box="[258,309,1804,1830]" italics="true" pageId="7" pageNumber="388">et al</emphasis>
. 2019
</bibRefCitation>
). Additionally, the material from Dehong Prefecture,
<collectingRegion id="4227F878F74E19120DD0F89E7556F8BA" box="[994,1080,1804,1830]" country="China" name="Yunnan" pageId="7" pageNumber="388">Yunnan</collectingRegion>
is unexpected, similar to those discussed in the remarks section of
<taxonomicName id="47E34D19F74E19120C18F8A2723AF8D6" authority="Gao &amp; Yang" authorityName="Gao &amp; Yang" authorityYear="2025" box="[554,852,1840,1866]" class="Insecta" family="Geotrupidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="388" phylum="Arthropoda" rank="species" species="sunfengyii" status="new species">
<emphasis id="B297EA88F74E19120C18F8A273D3F8D6" box="[554,701,1840,1866]" italics="true" pageId="7" pageNumber="388">B. sunfengyii</emphasis>
Gao &amp; Yang
</taxonomicName>
,
<emphasis id="B297EA88F74E19120D52F8A27284F8D5" bold="true" box="[864,1002,1839,1866]" pageId="7" pageNumber="388">
<taxonomicNameLabel id="A9A457F3F74E19120D52F8A27284F8D5" box="[864,1002,1839,1866]" pageId="7" pageNumber="388" rank="species">new species</taxonomicNameLabel>
</emphasis>
. For the same reason, this distribution should be confirmed with additional material.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,360 @@
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<mods:namePart id="8236C5D060A381BC74273DCF1B717F9F">Kochalka, John A.</mods:namePart>
<mods:affiliation id="45BCD542AA8CD8A3AB0A81D40738EF0C">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
</mods:name>
<mods:name id="28A6E75A59B872A0EBF731F222C209C6" type="personal">
<mods:role id="6D06C3D56B4F15E2566366CDDC2338CC">
<mods:roleTerm id="09D9249451FF1C5F762958F3B5CBB6A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="431046ED1BC53BA9D192376B5FA21146">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="675B5E8E304030739105DC9B3A0541A0">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
</mods:name>
<mods:name id="58A5C34775BD9E272065279090CEF71D" type="personal">
<mods:role id="8C31B67C8D4A4866F631B6EBA3049E3C">
<mods:roleTerm id="FFCCC81BC087095240FD6209D9DF7AF7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="398936432E018CEA17D814286828E8D2">Ramírez, Martin</mods:namePart>
<mods:affiliation id="C7F41E06BB52E6ECEE9A9C95E94873A6">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E0B0F6F8564D8E007B599E297D66057F">text</mods:typeOfResource>
<mods:relatedItem id="ABF09E6218EF2413EE6207A94E7CD72E" type="host">
<mods:titleInfo id="83482A5AF5EBB9385A7965CB60432A6C">
<mods:title id="EAAED817ADF4E3696F41EEE60B531595">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="52C6AC37F0281B0234175FA489E7D271">
<mods:date id="C3B8C6DDB24448D245B59F3A8B12128C">2025</mods:date>
<mods:detail id="683500CBFF7758153567A8E4D75CD507" type="pubDate">
<mods:number id="907321F4B700EBC10CB267C5B01A1AF2">2025-01-08</mods:number>
</mods:detail>
<mods:detail id="2D35E512F5F50BE6A4AE0967B8EDA08F" type="volume">
<mods:number id="F32B075031D5BD1D747BCA64E93D731C">5566</mods:number>
</mods:detail>
<mods:detail id="4F49D60B5BC8F939B685680739AAD3EB" type="issue">
<mods:number id="14D80837BEB06DBB488C30F57FB05B00">2</mods:number>
</mods:detail>
<mods:extent id="565F33F5AC69B9C1769E26153010B46F" unit="page">
<mods:start id="2AAFDC5778DDA267F57CBC558EA8A52C">201</mods:start>
<mods:end id="37A5E678B3CED9D7AD0E970B28D63D20">242</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="BC7740490C7C828397EB020355953D92">
<mods:url id="7D9BA6CACC798D40E9180977DCDCC452">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
</mods:location>
<mods:classification id="649BC48F61676E3EDD73CD8BD6073D2B">journal article</mods:classification>
<mods:identifier id="DE6A54C2FF696A131A84160F4340748B" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="030792040077E95B409A3A90EE346768" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5985FBE246F6D6A1626F2A05909415C7" type="ZooBank">5807FF17-467F-48D9-9A9F-ECDFC0161529</mods:identifier>
</mods:mods>
<treatment id="4D76240CFFA0FFEC62EBFD6BFBC8F905" LSID="urn:lsid:plazi:treatment:4D76240CFFA0FFEC62EBFD6BFBC8F905" httpUri="http://treatment.plazi.org/id/4D76240CFFA0FFEC62EBFD6BFBC8F905" lastPageId="38" lastPageNumber="239" pageId="30" pageNumber="231">
<subSubSection id="8DC5C691FFA0FFD462EBFD6BFE3FFD71" box="[151,411,654,681]" pageId="30" pageNumber="231" type="nomenclature">
<paragraph id="C560951AFFA0FFD462EBFD6BFE3FFD71" blockId="30.[151,411,654,717]" box="[151,411,654,681]" pageId="30" pageNumber="231">
<heading id="9E282276FFA0FFD462EBFD6BFE3FFD71" bold="true" box="[151,411,654,681]" fontSize="11" level="1" pageId="30" pageNumber="231" reason="1">
<emphasis id="F7AB4908FFA0FFD462EBFD6BFE3FFD71" bold="true" box="[151,411,654,681]" pageId="30" pageNumber="231">
<taxonomicName id="02DFEE99FFA0FFD462EBFD6BFE9FFD71" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[151,315,654,681]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
<emphasis id="F7AB4908FFA0FFD462EBFD6BFE9FFD71" bold="true" box="[151,315,654,681]" italics="true" pageId="30" pageNumber="231">Josa berlandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EC98F473FFA0FFD4633EFD6BFE3FFD71" box="[322,411,655,681]" pageId="30" pageNumber="231" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA0FFD462EBFD57FE9CFD15" box="[151,312,691,717]" pageId="30" pageNumber="231" type="description">
<paragraph id="C560951AFFA0FFD462EBFD57FE9CFD15" blockId="30.[151,411,654,717]" box="[151,312,691,717]" pageId="30" pageNumber="231">
<figureCitation id="5DE4899FFFA0FFD462EBFD57FE9CFD15" box="[151,312,691,717]" captionStart-0="FIGURE 23" captionStart-1="FIGURE 24" captionStart-2="FIGURE 25" captionStart-3="FIGURE 26" captionStart-4="FIGURE 27" captionStart-5="FIGURE 28" captionStart-6="FIGURE 29" captionStartId-0="31.[151,250,1883,1907]" captionStartId-1="32.[151,250,1879,1903]" captionStartId-2="33.[151,250,1836,1860]" captionStartId-3="34.[151,250,1866,1890]" captionStartId-4="35.[151,250,1779,1803]" captionStartId-5="36.[151,250,1731,1755]" captionStartId-6="37.[151,250,1517,1541]" captionTargetBox-0="[159,1426,193,1860]" captionTargetBox-1="[159,1425,181,1853]" captionTargetBox-2="[151,1436,212,1812]" captionTargetBox-3="[158,1427,185,1840]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetBox-6="[163,1429,193,1485]" captionTargetId-0="figure-17@31.[151,1436,181,1860]" captionTargetId-1="figure-22@32.[151,1436,181,1855]" captionTargetId-2="figure-17@33.[151,1436,181,1812]" captionTargetId-3="figure-22@34.[151,1435,181,1842]" captionTargetId-4="figure-17@35.[151,1435,181,1756]" captionTargetId-5="figure-22@36.[151,1435,181,1708]" captionTargetId-6="figure-17@37.[151,1436,181,1493]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="34" captionTargetPageId-4="35" captionTargetPageId-5="36" captionTargetPageId-6="37" captionText-0="FIGURE 23. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 24. Josa berlandi sp. nov., female (MUSENUV-Ar-2676; vchLAM-255): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-2="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." captionText-3="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." captionText-5="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." captionText-6="FIGURE 29. Geographical records of spiders in South America above 3000 masl, classified by altitude intervals." pageId="30" pageNumber="231">Figures 2329</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA0FFD462EBFD10FB39FC1B" pageId="30" pageNumber="231" type="materials_examined">
<paragraph id="C560951AFFA0FFD462EBFD10FB39FC1B" blockId="30.[151,1437,756,2043]" pageId="30" pageNumber="231">
<emphasis id="F7AB4908FFA0FFD462EBFD10FEECFCD6" bold="true" box="[151,328,756,782]" pageId="30" pageNumber="231">Type material.</emphasis>
<materialsCitation id="75B79F47FFA0FFD4632DFD10FD51FC8F" collectingDate="1975-03-05" collectorName="Quebrada El Chorro &amp; J. Kochalka" country="Colombia" county="Cienaga" elevation="3797" latitude="10.783334" location="Quebrada El Chorro" longLatPrecision="21" longitude="-73.53445" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" pageId="30" pageNumber="231" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="holotype">
<typeStatus id="1A642BB8FFA0FFD4632DFD10FE1AFCD6" box="[337,446,756,782]" pageId="30" pageNumber="231" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="BDC8D58AFFA0FFD463B7FD10FDC0FCD6" box="[459,612,756,782]" name="Colombia" pageId="30" pageNumber="231">COLOMBIA</collectingCountry>
.
<collectingRegion id="071B5BF8FFA0FFD4600DFD10FD56FCD6" box="[625,754,756,782]" country="Colombia" name="Magdalena" pageId="30" pageNumber="231">Magdalena</collectingRegion>
:
<collectingCounty id="2C01ED96FFA0FFD46082FD10FCF9FCD6" box="[766,861,756,782]" pageId="30" pageNumber="231">Ciénaga</collectingCounty>
,
<collectingMunicipality id="25040F60FFA0FFD46117FD11FF79FCEB" pageId="30" pageNumber="231">Parque Nacional Natural Sierra Nevada de Santa Marta</collectingMunicipality>
,
<collectorName id="682AF0CCFFA0FFD46299FCFCFE60FCEA" box="[229,452,792,818]" pageId="30" pageNumber="231">
<location id="C000C3C1FFA0FFD46299FCFCFE60FCEA" LSID="urn:lsid:plazi:treatment:4D76240CFFA0FFEC62EBFD6BFBC8F905:C000C3C1FFA0FFD46299FCFCFE60FCEA" box="[229,452,792,818]" country="Colombia" county="Cienaga" latitude="10.783334" longLatPrecision="21" longitude="-73.53445" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" name="Quebrada El Chorro" pageId="30" pageNumber="231" stateProvince="Magdalena">Quebrada El Chorro</location>
</collectorName>
,
<location id="C000C3C1FFA0FFD463B0FCFDFD3AFCEB" LSID="urn:lsid:plazi:treatment:4D76240CFFA0FFEC62EBFD6BFBC8F905:C000C3C1FFA0FFD463B0FCFDFD3AFCEB" box="[460,670,792,819]" country="Colombia" county="Cienaga" latitude="10.783334" longLatPrecision="21" longitude="-73.53445" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" name="Rama Sur del Lago" pageId="30" pageNumber="231" stateProvince="Magdalena">Rama Sur del Lago</location>
, manual caught,
<quantity id="022738FFFFA0FFD4612EFCFCFC02FCEB" box="[850,934,792,819]" metricMagnitude="3" metricUnit="m" metricValue="3.7969999999999997" pageId="30" pageNumber="231" unit="m" value="3797.0">
<elevation id="4EF27229FFA0FFD4612EFCFCFC02FCEB" box="[850,934,792,819]" metricMagnitude="3" metricUnit="m" metricValue="3.7969999999999997" pageId="30" pageNumber="231" unit="m" value="3797.0">3797 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFA0FFD461C9FCFDFB9BFCEC" box="[949,1087,793,820]" degrees="10" direction="north" minutes="47" orientation="latitude" pageId="30" pageNumber="231" precision="15" seconds="00" value="10.783334">10°4700″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFA0FFD46634FCFDFB7CFCEC" box="[1096,1240,793,820]" degrees="73" direction="west" minutes="32" orientation="longitude" pageId="30" pageNumber="231" precision="15" seconds="04" value="-73.53445">73°3204″W</geoCoordinate>
],
<collectorName id="682AF0CCFFA0FFD4669BFCFDFACCFCEA" box="[1255,1384,792,819]" pageId="30" pageNumber="231">J. Kochalka</collectorName>
leg.,
<date id="B161B3DAFFA0FFD462EBFCD8FEADFC8F" box="[151,265,828,855]" pageId="30" pageNumber="231" value="1975-03-05">
<collectingDate id="A1254A32FFA0FFD462EBFCD8FEADFC8F" box="[151,265,828,855]" pageId="30" pageNumber="231" value="1975-03-05">5.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA0FFD46369FCD8FE9BFC8F" box="[277,319,828,855]" count="1" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2675; vchLAM-253)
</materialsCitation>
.
<materialsCitation id="75B79F47FFA0FFD46083FCD9FB3EFC1B" collectingDate="1975-03-07" collectingDateMax="1975-03-16" collectingDateMin="1975-03-07" collectorName="J. Kochalka" country="Colombia" county="Cienaga" elevation="4785" latitude="10.900499" location="Lake of Rio Tucurinca" longLatPrecision="1" longitude="-73.8885" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" pageId="30" pageNumber="231" specimenCount="10" specimenCount-female="8" specimenCount-male="2" stateProvince="Magdalena" typeStatus="paratype">
<typeStatus id="1A642BB8FFA0FFD46083FCD9FCD4FC8F" box="[767,880,829,855]" pageId="30" pageNumber="231" type="paratype">Paratypes</typeStatus>
: same data as the holotype
<specimenCount id="D3D95E93FFA0FFD466DCFCD8FB6EFC8F" box="[1184,1226,828,855]" count="1" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
<specimenCount id="D3D95E93FFA0FFD466ADFCD8FB5FFC8F" box="[1233,1275,828,855]" count="4" pageId="30" pageNumber="231" type="female">4 ♀</specimenCount>
(MUSENUV- Ar-2674);
<location id="C000C3C1FFA0FFD46370FC84FDADFCA2" LSID="urn:lsid:plazi:treatment:4D76240CFFA0FFEC62EBFD6BFBC8F905:C000C3C1FFA0FFD46370FC84FDADFCA2" box="[268,521,864,890]" latitude="10.900499" longLatPrecision="1" longitude="-73.8885" name="Lake of Rio Tucurinca" pageId="30" pageNumber="231">Lake of Río Tucurinca</location>
, manual caught,
<quantity id="022738FFFFA0FFD460BEFC84FCBCFCA3" box="[706,792,864,891]" metricMagnitude="3" metricUnit="m" metricValue="4.785" pageId="30" pageNumber="231" unit="m" value="4785.0">
<elevation id="4EF27229FFA0FFD460BEFC84FCBCFCA3" box="[706,792,864,891]" metricMagnitude="3" metricUnit="m" metricValue="4.785" pageId="30" pageNumber="231" unit="m" value="4785.0">4785 m</elevation>
</quantity>
,
<collectorName id="682AF0CCFFA0FFD4615FFC85FC02FCA2" box="[803,934,864,891]" pageId="30" pageNumber="231">J. Kochalka</collectorName>
leg.,
<date id="B161B3DAFFA0FFD4619EFC84FBC6FCA3" box="[994,1122,864,891]" pageId="30" pageNumber="231" value="1975-03-16">
<collectingDate id="A1254A32FFA0FFD4619EFC84FBC6FCA3" box="[994,1122,864,891]" pageId="30" pageNumber="231" value="1975-03-16">16.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA0FFD46611FC84FB32FCA3" box="[1133,1174,864,891]" count="1" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2676; vchLAM-255),
<specimenCount id="D3D95E93FFA0FFD46334FC60FED6FC47" box="[328,370,900,927]" count="2" pageId="30" pageNumber="231" type="female">2 ♀</specimenCount>
(MUSENUV-Ar-2677),
<specimenCount id="D3D95E93FFA0FFD460F5FC60FD17FC47" box="[649,691,900,927]" count="1" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
(MACN-Ar 46694; vchLAM-254); norther peak of Norte Grande,
<quantity id="022738FFFFA0FFD462EBFC4CFF4AFC1B" box="[151,238,936,963]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="30" pageNumber="231" unit="m" value="3900.0">
<elevation id="4EF27229FFA0FFD462EBFC4CFF4AFC1B" box="[151,238,936,963]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="30" pageNumber="231" unit="m" value="3900.0">3900 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFA0FFD4637DFC4DFE04FC1C" box="[257,416,937,964]" degrees="10" direction="north" minutes="54" orientation="latitude" pageId="30" pageNumber="231" precision="1" seconds="01.8" value="10.900499">10°5401.8″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFA0FFD463D0FC4DFDF5FC1C" box="[428,593,937,964]" degrees="73" direction="west" minutes="53" orientation="longitude" pageId="30" pageNumber="231" precision="1" seconds="18.6" value="-73.8885">73°5318.6″W</geoCoordinate>
],
<date id="B161B3DAFFA0FFD4601FFC4CFD72FC1B" box="[611,726,936,963]" pageId="30" pageNumber="231" value="1975-03-07">
<collectingDate id="A1254A32FFA0FFD4601FFC4CFD72FC1B" box="[611,726,936,963]" pageId="30" pageNumber="231" value="1975-03-07">7.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA0FFD4609DFC4CFCAFFC1B" box="[737,779,936,963]" count="1" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
(MACN-Ar 46695; vchLAM-256)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA0FFD462BBFC28FA3CFBD2" pageId="30" pageNumber="231" type="etymology">
<paragraph id="C560951AFFA0FFD462BBFC28FA3CFBD2" blockId="30.[151,1437,756,2043]" pageId="30" pageNumber="231">
<emphasis id="F7AB4908FFA0FFD462BBFC28FEEAFC3E" bold="true" box="[199,334,972,998]" pageId="30" pageNumber="231">Etymology.</emphasis>
The specific epithet is a patronymic in honor of the late French arachnologist Lucien Berland, for his many contributions to arachnology including the Ecuadorian
<taxonomicName id="02DFEE99FFA0FFD46138FC14FC4DFBD2" authorityName="Bertkau" authorityYear="1878" box="[836,1001,1008,1034]" class="Arachnida" family="Anyphaenidae" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="231" phylum="Arthropoda" rank="family">Anyphaenidae</taxonomicName>
. It is a noun in possessive substantive.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA0FFD462BBFBF0FC36FB3B" pageId="30" pageNumber="231" type="diagnosis">
<paragraph id="C560951AFFA0FFD462BBFBF0FC36FB3B" blockId="30.[151,1437,756,2043]" pageId="30" pageNumber="231">
<emphasis id="F7AB4908FFA0FFD462BBFBF0FEE6FBF6" bold="true" box="[199,322,1044,1070]" pageId="30" pageNumber="231">Diagnosis.</emphasis>
Males are easily distinguished from other species by the bifid external section of the conductor, with beak-shaped anterior tip, longer than the posterior one, which is sharp and thin, elongated subapical protrusion (
<figureCitation id="5DE4899FFFA0FFD46716FBDCFF6EFBAF" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD462ABFBB8FEAEFBAE" box="[215,266,1116,1142]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD4636BFBB8FE2BFBAE" box="[279,399,1116,1142]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27AB, D</figureCitation>
), median apophysis bifid, laminar, projected, with the anterior tip longer than the posterior one (
<figureCitation id="5DE4899FFFA0FFD462AEFB64FED9FB42" box="[210,381,1152,1179]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">Fig. 27B, F, H</figureCitation>
), tegulum prolaterobasally projected (
<figureCitation id="5DE4899FFFA0FFD4614DFB64FC3BFB43" box="[817,927,1152,1179]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD461D1FB64FC45FB42" box="[941,993,1152,1178]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46192FB64FB87FB42" box="[1006,1059,1152,1178]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27A</figureCitation>
). Females are distinguished by a bilobed median field folded toward the median notch, wide lateral lobes with basally rounded posterior sclerotized projections, and an arch-shaped narrow median notch (
<figureCitation id="5DE4899FFFA0FFD46082FB2CFC22FB3B" box="[766,902,1224,1251]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25EF</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA0FFEC62BBFB08FBC8F905" lastPageId="38" lastPageNumber="239" pageId="30" pageNumber="231" type="description">
<paragraph id="C560951AFFA0FFD462BBFB08FB51F822" blockId="30.[151,1437,756,2043]" pageId="30" pageNumber="231">
<emphasis id="F7AB4908FFA0FFD462BBFB08FEFDFADE" bold="true" box="[199,345,1260,1286]" pageId="30" pageNumber="231">Description.</emphasis>
<emphasis id="F7AB4908FFA0FFD46319FB08FE3BFADE" box="[357,415,1260,1286]" italics="true" pageId="30" pageNumber="231">Male</emphasis>
(
<typeStatus id="1A642BB8FFA0FFD463CFFB08FDBDFADE" box="[435,537,1260,1286]" pageId="30" pageNumber="231" type="holotype">holotype</typeStatus>
, MUSENUV-Ar-2675; vchLAM-253) (
<figureCitation id="5DE4899FFFA0FFD46199FB08FB9BFADE" box="[997,1087,1260,1286]" captionStart="FIGURE 23" captionStartId="31.[151,250,1883,1907]" captionTargetBox="[159,1426,193,1860]" captionTargetId="figure-17@31.[151,1436,181,1860]" captionTargetPageId="31" captionText="FIGURE 23. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="30" pageNumber="231">Fig. 23</figureCitation>
): Total length 4.18. Carapace length 2.35, width 1.73, height 0.89. Clypeus length 0.15. Sternum length 1.20, width 0.83. Legs. Leg I: ti length 1.79/width 0.31, mt 1.45. II: ti 1.72/0.28, mt 1.43. III: ti 1.35/0.24, mt 1.35. IV: ti 1.71/0.30, mt 1.73. Spines: legs III, femur d 1-1-1, p 2ap, r 0-d1-d1; tibia v p1-2-2, p 1-d1-1-0, r 1-d1-1-0; metatarsus v 2-2-0, p d1-0. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 0-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1- 2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. IV=III but tibia v 2-2-2, metatarsus v 2-2-2. Abdomen: length 1.92, width 1.01, spiracleepigastrium 0.67. Color: carapace dark brown, with cervical groves spotted in black and converging in the large black patch on fovea. Lateral region with two poorly defined dark bands on sides, covered by abundant white setae. Ocular area on a black patch, eyes bordered in black rings. Clypeus brown, low with abundant white setae. Chelicerae brown with black stripes and fine setae on frontal edge. Endites, labium, and sternum brown with some dark spots; labium more pigmented. Legs coxae-tarsi brown, femora with dark longitudinal bands, tibia-metatarsi with rings of white setae; tarsi uniformly brown. Articles with abundant white setae. Abdomen dehydrated, with a pattern of faint light spots on a dark gray background, covered by abundant white setae. Venter uniformly dark gray. Palp: tegulum with very developed median hematodochae (
<figureCitation id="5DE4899FFFA0FFD4615EF924FC28F903" box="[802,908,1728,1755]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD461E9F924FC6CF902" box="[917,968,1728,1754]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26A</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD461ADF924FBA2F902" box="[977,1030,1728,1754]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27A</figureCitation>
). Median apophysis very large, bifid, with anterior branch long, laminar, with projected prolateral tip, posterior branch membranous at base and with the external border sub-rounded (
<figureCitation id="5DE4899FFFA0FFD46399F8ECFDD6F8FB" box="[485,626,1800,1827]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25AB</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46002F8ECFD76F8FA" box="[638,722,1800,1826]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26AC</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD460A1F8ECFCD7F8FA" box="[733,883,1800,1827]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27A, B, D, F</figureCitation>
). Conductor small, rounded, with projected base, external section apically wide and bifid, with a swollen, elongated projection; anterior tip of external section large, curved, and longer than posterior; posterior tip short and sharp; internal section ending as a sharp tip, converging with the posterior tip of external section (
<figureCitation id="5DE4899FFFA0FFD4600EF890FCA5F857" box="[626,769,1908,1935]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25AB</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46172F890FCC6F856" box="[782,866,1908,1934]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26AC</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46112F890FBACF856" box="[878,1032,1908,1935]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27A, B, D, F</figureCitation>
). Paramedian apophysis large, cup-shaped, heavily sclerotized, concave, apically acuminated (
<figureCitation id="5DE4899FFFA0FFD46156F87CFC08F86A" box="[810,940,1944,1970]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">Fig. 27C, E</figureCitation>
) Embolus short, tape-shaped at base, coiled, with poorly developed laminar projection, apically filiform; basal process small, flat, retrolateral region pointed in posterior region, prolateral region oval, poorly pronounced external keel (
<figureCitation id="5DE4899FFFA0FFD461ACF804FBFBF823" box="[976,1119,2016,2043]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="30" pageNumber="231">Figs 25AD</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD46615F804FB1AF822" box="[1129,1214,2016,2042]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="30" pageNumber="231">26AC</figureCitation>
,
<figureCitation id="5DE4899FFFA0FFD466B4F804FB4CF822" box="[1224,1256,2016,2042]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="30" pageNumber="231">27</figureCitation>
).
</paragraph>
<caption id="91A0C592FFA1FFD562EBF8BFFDECF840" pageId="31" pageNumber="232" startId="31.[151,250,1883,1907]" targetBox="[159,1426,193,1860]" targetPageId="31" targetType="figure">
<paragraph id="C560951AFFA1FFD562EBF8BFFDECF840" blockId="31.[151,1436,1883,1944]" pageId="31" pageNumber="232">
<emphasis id="F7AB4908FFA1FFD562EBF8BFFEBAF8AB" bold="true" box="[151,286,1883,1907]" pageId="31" pageNumber="232">FIGURE 23.</emphasis>
<taxonomicName id="02DFEE99FFA1FFD5635FF8B8FE08F8AC" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[291,428,1884,1908]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
<emphasis id="F7AB4908FFA1FFD5635FF8B8FE08F8AC" box="[291,428,1884,1908]" italics="true" pageId="31" pageNumber="232">Josa berlandi</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFA1FFD563CDF8B8FDA4F8AC" bold="true" box="[433,512,1884,1908]" pageId="31" pageNumber="232">
<taxonomicNameLabel id="EC98F473FFA1FFD563CDF8B8FDA4F8AC" box="[433,512,1884,1908]" pageId="31" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2675; vchLAM-253):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FF9EFFEA62EBF8B3FD3CF84C" pageId="32" pageNumber="233" startId="32.[151,250,1879,1903]" targetBox="[159,1425,181,1853]" targetPageId="32" targetType="figure">
<paragraph id="C560951AFF9EFFEA62EBF8B3FD3CF84C" blockId="32.[151,1436,1879,1940]" pageId="32" pageNumber="233">
<emphasis id="F7AB4908FF9EFFEA62EBF8B3FE86F8B7" bold="true" box="[151,290,1879,1903]" pageId="32" pageNumber="233">FIGURE 24.</emphasis>
<taxonomicName id="02DFEE99FF9EFFEA6357F8BCFE1CF8B7" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,440,1879,1903]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
<emphasis id="F7AB4908FF9EFFEA6357F8BCFE1CF8B7" box="[299,440,1879,1903]" italics="true" pageId="32" pageNumber="233">Josa berlandi</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FF9EFFEA63BDF8BCFDB0F8A8" bold="true" box="[449,532,1880,1904]" pageId="32" pageNumber="233">
<taxonomicNameLabel id="EC98F473FF9EFFEA63BDF8BCFDB0F8A8" box="[449,532,1880,1904]" pageId="32" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (MUSENUV-Ar-2676; vchLAM-255): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FF9FFFEB62EBF8C8FBB1F80C" pageId="33" pageNumber="234" startId="33.[151,250,1836,1860]" targetBox="[151,1436,212,1812]" targetPageId="33" targetType="figure">
<paragraph id="C560951AFF9FFFEB62EBF8C8FBB1F80C" blockId="33.[151,1436,1836,2005]" pageId="33" pageNumber="234">
<emphasis id="F7AB4908FF9FFFEB62EBF8C8FE80F89C" bold="true" box="[151,292,1836,1860]" pageId="33" pageNumber="234">FIGURE 25.</emphasis>
<taxonomicName id="02DFEE99FF9FFFEB6352F8C9FE19F89C" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[302,445,1836,1860]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
<emphasis id="F7AB4908FF9FFFEB6352F8C9FE19F89C" box="[302,445,1836,1860]" italics="true" pageId="33" pageNumber="234">Josa berlandi</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FF9FFFEB63BBF8C9FDB8F89D" bold="true" box="[455,540,1837,1861]" pageId="33" pageNumber="234">
<taxonomicNameLabel id="EC98F473FF9FFFEB63BBF8C9FDB8F89D" box="[455,540,1837,1861]" pageId="33" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view;
<emphasis id="F7AB4908FF9FFFEB634AF890FE73F854" bold="true" box="[310,471,1908,1932]" pageId="33" pageNumber="234">Abbreviations:</emphasis>
AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum.
</paragraph>
</caption>
<caption id="91A0C592FF9CFFE862EBF8AEFC9DF82A" pageId="34" pageNumber="235" startId="34.[151,250,1866,1890]" targetBox="[158,1427,185,1840]" targetPageId="34" targetType="figure">
<paragraph id="C560951AFF9CFFE862EBF8AEFC9DF82A" blockId="34.[151,1437,1866,2034]" pageId="34" pageNumber="235">
<emphasis id="F7AB4908FF9CFFE862EBF8AEFE80F8BA" bold="true" box="[151,292,1866,1890]" pageId="34" pageNumber="235">FIGURE 26.</emphasis>
<taxonomicName id="02DFEE99FF9CFFE86352F8AFFE19F8BA" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[302,445,1866,1890]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
<emphasis id="F7AB4908FF9CFFE86352F8AFFE19F8BA" box="[302,445,1866,1890]" italics="true" pageId="34" pageNumber="235">Josa berlandi</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FF9CFFE863BBF8AEFDB8F8BA" bold="true" box="[455,540,1866,1890]" pageId="34" pageNumber="235">
<taxonomicNameLabel id="EC98F473FF9CFFE863BBF8AEFDB8F8BA" box="[455,540,1866,1890]" pageId="34" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view.
<emphasis id="F7AB4908FF9CFFE862EBF876FE9CF872" bold="true" box="[151,312,1938,1962]" pageId="34" pageNumber="235">Abbreviations:</emphasis>
AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct.
</paragraph>
</caption>
<caption id="91A0C592FF9DFFE962EBF917FC32F844" pageId="35" pageNumber="236" startId="35.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="35" targetType="figure">
<paragraph id="C560951AFF9DFFE962EBF917FC32F844" blockId="35.[151,1436,1779,1948]" pageId="35" pageNumber="236">
<emphasis id="F7AB4908FF9DFFE962EBF917FE87F8D4" bold="true" box="[151,291,1779,1804]" pageId="35" pageNumber="236">FIGURE 27.</emphasis>
<taxonomicName id="02DFEE99FF9DFFE96357F910FE1CF8D4" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,440,1780,1804]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
<emphasis id="F7AB4908FF9DFFE96357F910FE1CF8D4" box="[299,440,1780,1804]" italics="true" pageId="35" pageNumber="236">Josa berlandi</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FF9DFFE963BDF910FDB0F8D4" bold="true" box="[449,532,1780,1804]" pageId="35" pageNumber="236">
<taxonomicNameLabel id="EC98F473FF9DFFE963BDF910FDB0F8D4" box="[449,532,1780,1804]" pageId="35" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details.
<emphasis id="F7AB4908FF9DFFE96047F8BBFD78F8AF" bold="true" box="[571,732,1887,1911]" pageId="35" pageNumber="236">Abbreviations:</emphasis>
C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum.
</paragraph>
</caption>
<caption id="91A0C592FF9AFFEE62EBF927FAFDF8FC" pageId="36" pageNumber="237" startId="36.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="36" targetType="figure">
<paragraph id="C560951AFF9AFFEE62EBF927FAFDF8FC" blockId="36.[151,1436,1731,1828]" pageId="36" pageNumber="237">
<emphasis id="F7AB4908FF9AFFEE62EBF927FE85F904" bold="true" box="[151,289,1731,1756]" pageId="36" pageNumber="237">FIGURE 28.</emphasis>
<taxonomicName id="02DFEE99FF9AFFEE635BF920FE17F904" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[295,435,1732,1756]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="237" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
<emphasis id="F7AB4908FF9AFFEE635BF920FE17F904" box="[295,435,1732,1756]" italics="true" pageId="36" pageNumber="237">Josa berlandi</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FF9AFFEE63C6F920FDAFF904" bold="true" box="[442,523,1732,1756]" pageId="36" pageNumber="237">
<taxonomicNameLabel id="EC98F473FF9AFFEE63C6F920FDAFF904" box="[442,523,1732,1756]" pageId="36" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view.
<emphasis id="F7AB4908FF9AFFEE618DF903FB36F927" bold="true" box="[1009,1170,1767,1791]" pageId="36" pageNumber="237">Abbreviations:</emphasis>
AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca.
</paragraph>
</caption>
<caption id="91A0C592FF9BFFEF62EBFA09FAB2F9DD" box="[151,1302,1517,1541]" pageId="37" pageNumber="238" startId="37.[151,250,1517,1541]" targetBox="[163,1429,193,1485]" targetPageId="37" targetType="figure">
<paragraph id="C560951AFF9BFFEF62EBFA09FAB2F9DD" blockId="37.[151,1302,1517,1541]" box="[151,1302,1517,1541]" pageId="37" pageNumber="238">
<emphasis id="F7AB4908FF9BFFEF62EBFA09FE84F9DD" bold="true" box="[151,288,1517,1541]" pageId="37" pageNumber="238">FIGURE 29.</emphasis>
Geographical records of spiders in South America above 3000 masl, classified by altitude intervals.
</paragraph>
</caption>
<paragraph id="C560951AFF9BFFEF62BBF9B4FD90F82F" blockId="37.[151,1437,1616,2039]" pageId="37" pageNumber="238">
<emphasis id="F7AB4908FF9BFFEF62BBF9B4FEBFF9B2" box="[199,283,1616,1642]" italics="true" pageId="37" pageNumber="238">Female</emphasis>
(
<typeStatus id="1A642BB8FF9BFFEF6351F9B5FE36F9B3" box="[301,402,1617,1643]" pageId="37" pageNumber="238" type="paratype">paratype</typeStatus>
, MUSENUV-Ar-2676; vchLAM-255). (
<figureCitation id="5DE4899FFF9BFFEF6122F9B4FC12F9B3" box="[862,950,1616,1643]" captionStart="FIGURE 24" captionStartId="32.[151,250,1879,1903]" captionTargetBox="[159,1425,181,1853]" captionTargetId="figure-22@32.[151,1436,181,1855]" captionTargetPageId="32" captionText="FIGURE 24. Josa berlandi sp. nov., female (MUSENUV-Ar-2676; vchLAM-255): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="37" pageNumber="238">Fig. 24</figureCitation>
): Total length 4.09. Carapace length 1.73, width 1.31, height 0.67. Clypeus length 0.08. Sternum length 1.06, width 0.71. Legs. Leg I: ti length 0.97/width 0.23, mt 0.70. II: ti 0.90/0.21, mt 0.69. III: ti 0.71/0.21, mt 0.66. IV: ti 1.12/0.22, mt 1,01. Spines: legs III, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. IIIIV, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v p1- 2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 2.53, width 1.54, spiracleepigastrium 0.80. Color pattern as in male, but dark brown, abdominal and legs spots more defined. Sternum and labium darker. Epigyne: anterior plate sclerotized with accentuated anterior excavations, median excavation long and narrow (
<figureCitation id="5DE4899FFF9BFFEF6036F8A8FD13F8BF" box="[586,695,1868,1895]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="37" pageNumber="238">Figs 25E</figureCitation>
,
<figureCitation id="5DE4899FFF9BFFEF60BAF8A8FD5EF8BE" box="[710,762,1868,1894]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="37" pageNumber="238">26D</figureCitation>
). Median field totally divided in two sub-rounded section interrupted by a large medial depression (
<figureCitation id="5DE4899FFF9BFFEF6002F894FCA0F853" box="[638,772,1904,1931]" captionStart="FIGURE 28" captionStartId="36.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-22@36.[151,1435,181,1708]" captionTargetPageId="36" captionText="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="37" pageNumber="238">
Fig. 28E
<emphasis id="F7AB4908FF9BFFEF609FF895FD55F853" bold="true" box="[739,753,1905,1931]" pageId="37" pageNumber="238"></emphasis>
F
</figureCitation>
). Lateral lobes with sclerotized extensions wide, swollen and apically rounded (
<figureCitation id="5DE4899FFF9BFFEF63E0F870FDA1F877" box="[412,517,1940,1967]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="37" pageNumber="238">Fig. 25E</figureCitation>
). Copulatory ducts short joining with the accessory bulb ducts at medial level. Spermathecae rounded in median position oriented towards lateral region. Ducts of accessory bulbs long, thin, convergent (
<figureCitation id="5DE4899FFF9BFFEF635EF838FE02F82F" box="[290,422,2012,2039]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="37" pageNumber="238">Figs 25EF</figureCitation>
,
<figureCitation id="5DE4899FFF9BFFEF63CDF838FE59F82E" box="[433,509,2012,2038]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="37" pageNumber="238">26EF</figureCitation>
,
<figureCitation id="5DE4899FFF9BFFEF6074F839FD8CF82F" box="[520,552,2013,2039]" captionStart="FIGURE 28" captionStartId="36.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-22@36.[151,1435,181,1708]" captionTargetPageId="36" captionText="FIGURE 28. Josa berlandi sp. nov., female (MACN-Ar 46695; vchLAM-256), SEM images: A Dorsal view; B Apical view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="37" pageNumber="238">28</figureCitation>
).
</paragraph>
<caption id="91A0C592FF98FFEC62EBFA09FAFBF9DD" box="[151,1375,1517,1541]" pageId="38" pageNumber="239" startId="38.[151,250,1517,1541]" targetBox="[163,1429,193,1485]" targetPageId="38" targetType="figure">
<paragraph id="C560951AFF98FFEC62EBFA09FAFBF9DD" blockId="38.[151,1375,1517,1541]" box="[151,1375,1517,1541]" pageId="38" pageNumber="239">
<emphasis id="F7AB4908FF98FFEC62EBFA09FE84F9DD" bold="true" box="[151,288,1517,1541]" pageId="38" pageNumber="239">FIGURE 30.</emphasis>
Geographical records of
<taxonomicName id="02DFEE99FF98FFEC605CFA09FD16F9DD" authorityName="Bertkau" authorityYear="1878" box="[544,690,1517,1541]" class="Arachnida" family="Anyphaenidae" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="239" phylum="Arthropoda" rank="family">Anyphaenidae</taxonomicName>
in South America above 3000 masl, classified by altitude intervals.
</paragraph>
</caption>
<paragraph id="C560951AFF98FFEC62BBF9B3FBC8F905" blockId="38.[151,1437,1623,1758]" pageId="38" pageNumber="239">
<emphasis id="F7AB4908FF98FFEC62BBF9B3FEE5F9A9" bold="true" box="[199,321,1623,1649]" pageId="38" pageNumber="239">Variation:</emphasis>
Males (n=3): total length: 4.084.18; carapace length: 2.102.35, width: 1.591.73; abdomen length: 1.871.93, width: 1.011.25. Females (n=5): total length: 3.894.09; carapace length: 1.732.10, width: 1.311.59; abdomen length: 1.932.53, width: 1.251.54.
<emphasis id="F7AB4908FF98FFEC62BBF927FEC5F905" bold="true" box="[199,353,1731,1757]" pageId="38" pageNumber="239">Distribution.</emphasis>
Only known from Sierra Nevada de Santa Marta
<collectingCountry id="BDC8D58AFF98FFEC61F2F927FC58F905" box="[910,1020,1731,1757]" name="Colombia" pageId="38" pageNumber="239">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFF98FFEC6670F927FBFBF905" box="[1036,1119,1731,1757]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="38" pageNumber="239">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="2453A579842476465CA5D825C28C4076" ID-DOI="10.11646/zootaxa.5566.2.1" ID-ISSN="1175-5326" ID-ZooBank="5807FF17-467F-48D9-9A9F-ECDFC0161529" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737369844893" checkinUser="plazi" docAuthor="Martínez, Leonel, Kochalka, John A., Cabra-Garcia, Jimmy &amp; Ramírez, Martin" docDate="2025" docId="4D76240CFFA8FFD462EBFADFFB5CFDE5" docLanguage="en" docName="zootaxa.5566.2.1.pdf" docOrigin="Zootaxa 5566 (2)" docSource="https://doi.org/10.11646/zootaxa.5566.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Josa olimpica Martínez, Kochalka, Cabra-Garcia &amp; Ramírez, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="231" masterDocId="B14F5C74FFBEFFCA627CFFE4FFA4FFD8" masterDocTitle="Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America" masterLastPageNumber="242" masterPageNumber="201" pageNumber="223" updateTime="1737369942339" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="F304DC65159AA2D3779B8B62F147F71A">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
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<mods:namePart id="9617FB43068345851B2A7F8B434A4293">Martínez, Leonel</mods:namePart>
<mods:affiliation id="078C74968424C998758902EBB7DDE533">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
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<mods:namePart id="EFD4C212C16A57E5AF3706EFB73CDE78">Kochalka, John A.</mods:namePart>
<mods:affiliation id="0EEFF070DF7B860D82EDE979CB074C72">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
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<mods:namePart id="D4DDAA74596A28EE275415F09AE64864">Cabra-Garcia, Jimmy</mods:namePart>
<mods:affiliation id="433DF690456FABE3489EC8C23420CE6A">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
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<mods:namePart id="99FA2B3D5A5D4631D4DB64695A850F6D">Ramírez, Martin</mods:namePart>
<mods:affiliation id="9918628317D77B1452813EA7D580B9E8">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
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<emphasis id="F7AB4908FFA8FFDC62EBFADFFE9AFA8D" bold="true" box="[151,318,1338,1365]" italics="true" pageId="22" pageNumber="223">Josa olimpica</emphasis>
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<figureCitation id="5DE4899FFFA8FFDC62EBFABBFEB0FAA1" box="[151,276,1375,1401]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStart-2="FIGURE 19" captionStart-3="FIGURE 20" captionStart-4="FIGURE 21" captionStart-5="FIGURE 22" captionStartId-0="24.[151,250,1880,1904]" captionStartId-1="25.[151,250,1879,1903]" captionStartId-2="26.[151,250,1851,1875]" captionStartId-3="27.[151,250,1858,1882]" captionStartId-4="28.[151,250,1779,1803]" captionStartId-5="29.[151,250,1731,1755]" captionTargetBox-0="[155,1428,182,1847]" captionTargetBox-1="[158,1429,192,1849]" captionTargetBox-2="[151,1428,212,1819]" captionTargetBox-3="[162,1427,189,1832]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetId-0="figure-22@24.[151,1436,181,1856]" captionTargetId-1="figure-17@25.[151,1436,181,1855]" captionTargetId-2="figure-22@26.[151,1436,181,1828]" captionTargetId-3="figure-17@27.[158,1430,181,1834]" captionTargetId-4="figure-22@28.[151,1435,181,1756]" captionTargetId-5="figure-17@29.[151,1435,181,1708]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="26" captionTargetPageId-3="27" captionTargetPageId-4="28" captionTargetPageId-5="29" captionText-0="FIGURE 17. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 18. Josa olimpica sp. nov., female (MUSENUV-Ar-2683; vchLAM-261): A Habitus, dorsal view; B Carapace; C Habitus, ventral view; D Sternum." captionText-2="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." captionText-3="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." captionText-5="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="22" pageNumber="223">Figs 1722</figureCitation>
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:
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, southern branch of the lake, manual caught,
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, [
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<collectorName id="682AF0CCFFA8FFDC67F6FA2FFEA6F9D1" pageId="22" pageNumber="223">J. Kochalka</collectorName>
leg.,
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.
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: same locality data as the
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;
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(MUSENUV-Ar- 2695)
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,
<materialsCitation id="75B79F47FFA8FFDC6299F9BFFA3CF9AD" box="[229,1432,1627,1653]" collectingDate="1974-02-02" country="Colombia" elevation="4398" location="Tip of Pico Norte Grande de la Cuchilla Cesarguarniaga" pageId="22" pageNumber="223" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
<location id="C000C3C1FFA8FFDC6299F9BFFCF1F9AD" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC6299F9BFFCF1F9AD" box="[229,853,1627,1653]" country="Colombia" name="Tip of Pico Norte Grande de la Cuchilla Cesarguarniaga" pageId="22" pageNumber="223" stateProvince="Magdalena">Tip of Pico Norte Grande de la Cuchilla Cesarguarniaga</location>
,
<quantity id="022738FFFFA8FFDC6123F9BFFC11F9AD" box="[863,949,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="4.398" pageId="22" pageNumber="223" unit="m" value="4398.0">
<elevation id="4EF27229FFA8FFDC6123F9BFFC11F9AD" box="[863,949,1627,1653]" metricMagnitude="3" metricUnit="m" metricValue="4.398" pageId="22" pageNumber="223" unit="m" value="4398.0">4398 m</elevation>
</quantity>
,
<date id="B161B3DAFFA8FFDC61BCF9BFFB8DF9AD" box="[960,1065,1627,1653]" pageId="22" pageNumber="223" value="1974-02-02">
<collectingDate id="A1254A32FFA8FFDC61BCF9BFFB8DF9AD" box="[960,1065,1627,1653]" pageId="22" pageNumber="223" value="1974-02-02">2.II.1974</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA8FFDC6648F9BFFBF9F9AD" box="[1076,1117,1627,1653]" count="1" pageId="22" pageNumber="223" type="male">1 ♂</specimenCount>
<specimenCount id="D3D95E93FFA8FFDC661FF9BFFB28F9AD" box="[1123,1164,1627,1653]" count="3" pageId="22" pageNumber="223" type="female">3 ♀</specimenCount>
(MUSENUV-Ar-2687)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA8FFDC62EBF99BFA3CF941" box="[151,1432,1662,1689]" collectingDate="1975-03-07" country="Colombia" elevation="3992" location="Norte Grande" municipality="Serrania Dunanchucua" pageId="22" pageNumber="223" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
<collectingMunicipality id="25040F60FFA8FFDC62EBF99BFE3CF941" box="[151,408,1663,1689]" pageId="22" pageNumber="223">Serranía Dunanchucua</collectingMunicipality>
, northern peak of
<location id="C000C3C1FFA8FFDC601CF99BFD5FF941" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC601CF99BFD5FF941" box="[608,763,1663,1689]" country="Colombia" municipality="Serrania Dunanchucua" name="Norte Grande" pageId="22" pageNumber="223" stateProvince="Magdalena">Norte Grande</location>
,
<quantity id="022738FFFFA8FFDC6179F99BFCF8F941" box="[773,860,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="3.992" pageId="22" pageNumber="223" unit="m" value="3992.0">
<elevation id="4EF27229FFA8FFDC6179F99BFCF8F941" box="[773,860,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="3.992" pageId="22" pageNumber="223" unit="m" value="3992.0">3992 m</elevation>
</quantity>
,
<date id="B161B3DAFFA8FFDC611BF99AFC7DF941" box="[871,985,1662,1689]" pageId="22" pageNumber="223" value="1975-03-07">
<collectingDate id="A1254A32FFA8FFDC611BF99AFC7DF941" box="[871,985,1662,1689]" pageId="22" pageNumber="223" value="1975-03-07">7.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA8FFDC6198F99BFBA9F941" box="[996,1037,1663,1689]" count="1" pageId="22" pageNumber="223" type="male">1 ♂</specimenCount>
(MACN-Ar 46696; vchLAM-262)
</materialsCitation>
,
<materialsCitation id="75B79F47FFA8FFDC62EBF947FE77F965" box="[151,467,1698,1725]" collectingDate="1975-03-07" country="Colombia" elevation="3992" isTailEnumeration="true" location="Norte Grande" municipality="Serrania Dunanchucua" pageId="22" pageNumber="223" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
<specimenCount id="D3D95E93FFA8FFDC62EBF947FF60F965" box="[151,196,1699,1725]" count="1" pageId="22" pageNumber="223" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2695)
</materialsCitation>
,
<quantity id="022738FFFFA8FFDC639EF947FD98F965" box="[482,572,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="4.236" pageId="22" pageNumber="223" unit="m" value="4236.0">4236 m</quantity>
,
<specimenCount id="D3D95E93FFA8FFDC6037F947FDDDF965" box="[587,633,1699,1725]" count="1" pageId="22" pageNumber="223" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2690);
<materialsCitation id="75B79F47FFA8FFDC61EBF947FD17F939" collectingDate="1974-02-13" collectorName="Quebrada El Chorro" country="Colombia" elevation="3718" location="Rio Sevillano" pageId="22" pageNumber="223" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
<collectorName id="682AF0CCFFA8FFDC61EBF947FB27F965" box="[919,1155,1699,1725]" pageId="22" pageNumber="223">Quebrada El Chorro</collectorName>
, north of
<location id="C000C3C1FFA8FFDC6685F947FA3CF965" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC6685F947FA3CF965" box="[1273,1432,1699,1725]" country="Colombia" name="Rio Sevillano" pageId="22" pageNumber="223" stateProvince="Magdalena">Río Sevillano</location>
,
<quantity id="022738FFFFA8FFDC62EBF923FF4AF939" box="[151,238,1735,1761]" metricMagnitude="3" metricUnit="m" metricValue="3.718" pageId="22" pageNumber="223" unit="m" value="3718.0">
<elevation id="4EF27229FFA8FFDC62EBF923FF4AF939" box="[151,238,1735,1761]" metricMagnitude="3" metricUnit="m" metricValue="3.718" pageId="22" pageNumber="223" unit="m" value="3718.0">3718 m</elevation>
</quantity>
,
<date id="B161B3DAFFA8FFDC6286F923FED5F939" box="[250,369,1735,1761]" pageId="22" pageNumber="223" value="1974-02-13">
<collectingDate id="A1254A32FFA8FFDC6286F923FED5F939" box="[250,369,1735,1761]" pageId="22" pageNumber="223" value="1974-02-13">13.II.1974</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA8FFDC6301F923FE03F939" box="[381,423,1735,1761]" count="1" pageId="22" pageNumber="223" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2692)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA8FFDC60C3F923FE39F8DD" collectingDate="1975-03-16" country="Colombia" elevation="4785" location="Lago del Rio Tucurinca" municipality="Rio Tucurinca" pageId="22" pageNumber="223" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena" typeStatus="paratype">
<collectingMunicipality id="25040F60FFA8FFDC60C3F923FCC6F939" box="[703,866,1735,1761]" pageId="22" pageNumber="223">Río Tucurinca</collectingMunicipality>
,
<location id="C000C3C1FFA8FFDC6110F923FBDDF939" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC6110F923FBDDF939" box="[876,1145,1735,1761]" country="Colombia" municipality="Rio Tucurinca" name="Lago del Rio Tucurinca" pageId="22" pageNumber="223" stateProvince="Magdalena">Lago del Rio Tucurinca</location>
,
<quantity id="022738FFFFA8FFDC66FFF922FB7EF939" box="[1155,1242,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="4.785" pageId="22" pageNumber="223" unit="m" value="4785.0">
<elevation id="4EF27229FFA8FFDC66FFF922FB7EF939" box="[1155,1242,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="4.785" pageId="22" pageNumber="223" unit="m" value="4785.0">4785 m</elevation>
</quantity>
,
<date id="B161B3DAFFA8FFDC669AF922FAC3F939" box="[1254,1383,1734,1761]" pageId="22" pageNumber="223" value="1975-03-16">
<collectingDate id="A1254A32FFA8FFDC669AF922FAC3F939" box="[1254,1383,1734,1761]" pageId="22" pageNumber="223" value="1975-03-16">16.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA8FFDC670EF923FA38F939" box="[1394,1436,1735,1761]" count="1" pageId="22" pageNumber="223" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2693)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA8FFDC63D5F90FFE39F8F1" collectingDate="1974-01-12" country="Colombia" elevation="5275" latitude="10.83325" location="Pico Cristobal Colon" longLatPrecision="1" longitude="-73.66905" municipality="East" pageId="22" pageNumber="223" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena" typeStatus="paratype">
<collectingMunicipality id="25040F60FFA8FFDC63D5F90FFE7DF8DD" box="[425,473,1771,1797]" pageId="22" pageNumber="223">East</collectingMunicipality>
tip of
<location id="C000C3C1FFA8FFDC6054F90FFCBCF8DD" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC6054F90FFCBCF8DD" box="[552,792,1771,1797]" country="Colombia" latitude="10.83325" longLatPrecision="1" longitude="-73.66905" municipality="East" name="Pico Cristobal Colon" pageId="22" pageNumber="223" stateProvince="Magdalena">Pico Cristobal Colón</location>
,
<quantity id="022738FFFFA8FFDC6159F90EFCD9F8DD" box="[805,893,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="5.275" pageId="22" pageNumber="223" unit="m" value="5275.0">
<elevation id="4EF27229FFA8FFDC6159F90EFCD9F8DD" box="[805,893,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="5.275" pageId="22" pageNumber="223" unit="m" value="5275.0">5275 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFA8FFDC61EDF908FB94F8DF" box="[913,1072,1772,1799]" degrees="10" direction="north" minutes="49" orientation="latitude" pageId="22" pageNumber="223" precision="1" seconds="59.7" value="10.83325">10°4959.7″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFA8FFDC6641F908FB47F8DF" box="[1085,1251,1772,1799]" degrees="73" direction="west" minutes="40" orientation="longitude" pageId="22" pageNumber="223" precision="1" seconds="08.6" value="-73.66905">73°4008.6″W</geoCoordinate>
],
<date id="B161B3DAFFA8FFDC668AF90FFAC0F8DD" box="[1270,1380,1771,1797]" pageId="22" pageNumber="223" value="1974-01-12">
<collectingDate id="A1254A32FFA8FFDC668AF90FFAC0F8DD" box="[1270,1380,1771,1797]" pageId="22" pageNumber="223" value="1974-01-12">12.I.1974</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA8FFDC670DF90FFA38F8DD" box="[1393,1436,1771,1797]" count="1" pageId="22" pageNumber="223" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2696)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA8FFDC63DBF8EBFF78F8A9" collectingDate="1974-01-10" country="Colombia" elevation="5500" latitude="10.836194" location="Pico Cristobal Colon" longLatPrecision="1" longitude="-73.686386" municipality="East" pageId="22" pageNumber="223" specimenCount="1" stateProvince="Magdalena" typeStatus="paratype">
<collectingMunicipality id="25040F60FFA8FFDC63DBF8EBFE73F8F1" box="[423,471,1807,1833]" pageId="22" pageNumber="223">East</collectingMunicipality>
tip of NW
<location id="C000C3C1FFA8FFDC602DF8EBFC98F8F1" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC602DF8EBFC98F8F1" box="[593,828,1807,1833]" country="Colombia" latitude="10.836194" longLatPrecision="1" longitude="-73.686386" municipality="East" name="Pico Cristobal Colon" pageId="22" pageNumber="223" stateProvince="Magdalena">Pico Cristobal Colón</location>
, almost at superior peak, under loose rocks, on a nearly vertical cliff, surrounded by glacial ice,
<quantity id="022738FFFFA8FFDC6030F8D6FD06F895" box="[588,674,1842,1869]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="22" pageNumber="223" unit="m" value="5500.0">
<elevation id="4EF27229FFA8FFDC6030F8D6FD06F895" box="[588,674,1842,1869]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="22" pageNumber="223" unit="m" value="5500.0">5500 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFA8FFDC60CFF8D0FCF6F897" box="[691,850,1844,1871]" degrees="10" direction="north" minutes="50" orientation="latitude" pageId="22" pageNumber="223" precision="1" seconds="10.3" value="10.836194">10°5010.3″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFA8FFDC6120F8D0FBA4F897" box="[860,1024,1844,1871]" degrees="73" direction="west" minutes="41" orientation="longitude" pageId="22" pageNumber="223" precision="1" seconds="11.0" value="-73.686386">73°4111.0″W</geoCoordinate>
],
<date id="B161B3DAFFA8FFDC666DF8D7FBDAF895" box="[1041,1150,1843,1869]" pageId="22" pageNumber="223" value="1974-01-10">
<collectingDate id="A1254A32FFA8FFDC666DF8D7FBDAF895" box="[1041,1150,1843,1869]" pageId="22" pageNumber="223" value="1974-01-10">10.I.1974</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA8FFDC66F4F8D7FB73F895" box="[1160,1239,1843,1869]" count="1" pageId="22" pageNumber="223" type="generic">1 imm.</specimenCount>
(MUSENUV-Ar- 2697)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA8FFDC6294F8B3FB58F8A9" box="[232,1276,1878,1905]" collectingDate="1975-03-12" country="Colombia" elevation="4176" location="Rio Tucurinca" municipality="Lago Arucuina" pageId="22" pageNumber="223" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena" typeStatus="paratype">
<collectingMunicipality id="25040F60FFA8FFDC6294F8B3FE37F8A9" box="[232,403,1879,1905]" pageId="22" pageNumber="223">Lago Arucuina</collectingMunicipality>
,
<location id="C000C3C1FFA8FFDC63E3F8B3FDE7F8A9" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC63E3F8B3FDE7F8A9" box="[415,579,1879,1905]" country="Colombia" municipality="Lago Arucuina" name="Rio Tucurinca" pageId="22" pageNumber="223" stateProvince="Magdalena">Río Tucurinca</location>
,
<quantity id="022738FFFFA8FFDC6033F8B3FD03F8A9" box="[591,679,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="4.176" pageId="22" pageNumber="223" unit="m" value="4176.0">
<elevation id="4EF27229FFA8FFDC6033F8B3FD03F8A9" box="[591,679,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="4.176" pageId="22" pageNumber="223" unit="m" value="4176.0">4176 m</elevation>
</quantity>
,
<date id="B161B3DAFFA8FFDC60CFF8B2FC90F8A9" box="[691,820,1878,1905]" pageId="22" pageNumber="223" value="1975-03-12">
<collectingDate id="A1254A32FFA8FFDC60CFF8B2FC90F8A9" box="[691,820,1878,1905]" pageId="22" pageNumber="223" value="1975-03-12">12.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA8FFDC613CF8B3FCCFF8A9" box="[832,875,1879,1905]" count="1" pageId="22" pageNumber="223" type="female">1 ♀</specimenCount>
(MACN-Ar 46697; vchLAM-263)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA8FFDC6774F8B3FC81F84D" collectingDate="1975-03-16" country="Colombia" elevation="4785" location="Lago del Rio Tucurinca" pageId="22" pageNumber="223" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
<location id="C000C3C1FFA8FFDC6774F8B3FEAFF84D" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC6774F8B3FEAFF84D" country="Colombia" name="Lago del Rio Tucurinca" pageId="22" pageNumber="223" stateProvince="Magdalena">Lago del Rio Tucurinca</location>
,
<quantity id="022738FFFFA8FFDC636EF89EFEC2F84D" box="[274,358,1914,1941]" metricMagnitude="3" metricUnit="m" metricValue="4.785" pageId="22" pageNumber="223" unit="m" value="4785.0">
<elevation id="4EF27229FFA8FFDC636EF89EFEC2F84D" box="[274,358,1914,1941]" metricMagnitude="3" metricUnit="m" metricValue="4.785" pageId="22" pageNumber="223" unit="m" value="4785.0">4785 m</elevation>
</quantity>
,
<date id="B161B3DAFFA8FFDC6312F89EFE4AF84D" box="[366,494,1914,1941]" pageId="22" pageNumber="223" value="1975-03-16">
<collectingDate id="A1254A32FFA8FFDC6312F89EFE4AF84D" box="[366,494,1914,1941]" pageId="22" pageNumber="223" value="1975-03-16">16.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA8FFDC638AF89FFDB9F84D" box="[502,541,1915,1941]" count="1" pageId="22" pageNumber="223" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2698)
</materialsCitation>
,
<materialsCitation id="75B79F47FFA8FFDC6151F89FFBF8F84D" box="[813,1116,1915,1941]" collectingDate="1975-03-16" country="Colombia" elevation="4785" isTailEnumeration="true" location="Lago del Rio Tucurinca" pageId="22" pageNumber="223" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
<specimenCount id="D3D95E93FFA8FFDC6151F89FFCF7F84D" box="[813,851,1915,1941]" count="1" pageId="22" pageNumber="223" type="male">1♂</specimenCount>
(MUSENUV-Ar-2699)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA8FFDC6618F89FFE6AF861" country="Colombia" location="South" pageId="22" pageNumber="223" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
<location id="C000C3C1FFA8FFDC6618F89FFB01F84D" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC6618F89FFB01F84D" box="[1124,1189,1915,1941]" country="Colombia" name="South" pageId="22" pageNumber="223" stateProvince="Magdalena">South</location>
slope of Pico Tiaroma,
<specimenCount id="D3D95E93FFA8FFDC62EBF87BFF65F861" box="[151,193,1951,1977]" count="1" pageId="22" pageNumber="223" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2700)
</materialsCitation>
.
<materialsCitation id="75B79F47FFA8FFDD63A4F87BFD9FFF69" collectingDate="1973-01-06" collectorName="J. Kochalka" country="Colombia" county="Valledupar" elevation="4230" lastPageId="23" lastPageNumber="224" latitude="10.797223" location="Donachui" longLatPrecision="1" longitude="-73.62625" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" pageId="22" pageNumber="223" specimenCount="1" specimenCount-female="1" stateProvince="Cesar" typeStatus="paratype">
<emphasis id="F7AB4908FFA8FFDC63A4F87BFDBFF861" box="[472,539,1951,1977]" italics="true" pageId="22" pageNumber="223">
<collectingRegion id="071B5BF8FFA8FFDC63A4F87BFDBFF861" box="[472,539,1951,1977]" country="Colombia" name="Cesar" pageId="22" pageNumber="223">César</collectingRegion>
</emphasis>
:
<collectingCounty id="2C01ED96FFA8FFDC6055F87BFD00F861" box="[553,676,1951,1977]" pageId="22" pageNumber="223">Valledupar</collectingCounty>
,
<collectingMunicipality id="25040F60FFA8FFDC60D3F87BFABAF861" box="[687,1310,1951,1977]" pageId="22" pageNumber="223">Parque Nacional Natural Sierra Nevada de Santa Marta</collectingMunicipality>
,
<location id="C000C3C1FFA8FFDC6754F87BFA3CF861" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC6754F87BFA3CF861" box="[1320,1432,1951,1977]" country="Colombia" county="Valledupar" latitude="10.797223" longLatPrecision="1" longitude="-73.62625" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" name="Donachui" pageId="22" pageNumber="223" stateProvince="Cesar">Donachui</location>
,
<location id="C000C3C1FFA8FFDC62EBF827FE93F805" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA8FFDC62EBF827FE93F805" box="[151,311,1987,2013]" country="Colombia" county="Valledupar" latitude="10.797223" longLatPrecision="1" longitude="-73.62625" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" name="Rio Donachui" pageId="22" pageNumber="223" stateProvince="Cesar">Río Donachui</location>
, top of woody vegetation, manual caught,
<quantity id="022738FFFFA8FFDC6168F827FCCFF805" box="[788,875,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="4.2299999999999995" pageId="22" pageNumber="223" unit="m" value="4230.0">
<elevation id="4EF27229FFA8FFDC6168F827FCCFF805" box="[788,875,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="4.2299999999999995" pageId="22" pageNumber="223" unit="m" value="4230.0">4230 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFA8FFDC6102F820FBB9F807" box="[894,1053,1988,2015]" degrees="10" direction="north" minutes="47" orientation="latitude" pageId="22" pageNumber="223" precision="1" seconds="50.0" value="10.797223">10°4750.0″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFA8FFDC6655F820FB6AF807" box="[1065,1230,1988,2015]" degrees="73" direction="west" minutes="37" orientation="longitude" pageId="22" pageNumber="223" precision="1" seconds="34.5" value="-73.62625">73°3734.5″W</geoCoordinate>
],
<collectorName id="682AF0CCFFA8FFDC669DF827FAC1F805" box="[1249,1381,1987,2013]" pageId="22" pageNumber="223">J. Kochalka</collectorName>
leg.,
<date id="B161B3DAFFA9FFDD62EBFF73FF53FF69" box="[151,247,151,177]" pageId="23" pageNumber="224" value="1973-01-06">
<collectingDate id="A1254A32FFA9FFDD62EBFF73FF53FF69" box="[151,247,151,177]" pageId="23" pageNumber="224" value="1973-01-06">6.I.1973</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA9FFDD637FFF73FE8AFF69" box="[259,302,151,177]" count="1" pageId="23" pageNumber="224" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2685)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA9FFDD6034FF73FE76FF0D" collectingDate="1973-01-02" country="Colombia" elevation="3596" latitude="10.784584" location="Rio Donachui" longLatPrecision="1" longitude="-73.561226" municipality="Meoyaca" pageId="23" pageNumber="224" specimenCount="1" specimenCount-female="1" stateProvince="Cesar" typeStatus="paratype">
<collectingMunicipality id="25040F60FFA9FFDD6034FF73FD16FF69" box="[584,690,151,177]" pageId="23" pageNumber="224">Meoyaca</collectingMunicipality>
,
<location id="C000C3C1FFA9FFDD60C1FF73FCF9FF69" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA9FFDD60C1FF73FCF9FF69" box="[701,861,151,177]" country="Colombia" latitude="10.784584" longLatPrecision="1" longitude="-73.561226" municipality="Meoyaca" name="Rio Donachui" pageId="23" pageNumber="224" stateProvince="Cesar">Río Donachuí</location>
,
<quantity id="022738FFFFA9FFDD6115FF72FC64FF69" box="[873,960,150,177]" metricMagnitude="3" metricUnit="m" metricValue="3.596" pageId="23" pageNumber="224" unit="m" value="3596.0">
<elevation id="4EF27229FFA9FFDD6115FF72FC64FF69" box="[873,960,150,177]" metricMagnitude="3" metricUnit="m" metricValue="3.596" pageId="23" pageNumber="224" unit="m" value="3596.0">3596 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFA9FFDD61A8FF7CFBD7FF6B" box="[980,1139,152,179]" degrees="10" direction="north" minutes="47" orientation="latitude" pageId="23" pageNumber="224" precision="1" seconds="04.5" value="10.784584">10°4704.5″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFA9FFDD6603FF7CFA81FF6B" box="[1151,1317,152,179]" degrees="73" direction="west" minutes="33" orientation="longitude" pageId="23" pageNumber="224" precision="1" seconds="40.4" value="-73.561226">73°3340.4″W</geoCoordinate>
],
<date id="B161B3DAFFA9FFDD6744FF73FA33FF69" box="[1336,1431,151,177]" pageId="23" pageNumber="224" value="1973-01-02">
<collectingDate id="A1254A32FFA9FFDD6744FF73FA33FF69" box="[1336,1431,151,177]" pageId="23" pageNumber="224" value="1973-01-02">2.I.1973</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA9FFDD62EBFF5FFF67FF0D" box="[151,195,187,213]" count="1" pageId="23" pageNumber="224" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2689)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA9FFDD639CFF5FFD4FFEC5" collectingDate="1973-01-09" country="Colombia" county="Naboba" elevation="4800" latitude="10.818889" location="Rio Donachui" longLatPrecision="1" longitude="-73.63295" municipality="Meoyaca" pageId="23" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Cesar" typeStatus="paratype">
same locality as before, but
<collectingCounty id="2C01ED96FFA9FFDD6159FF5FFCD8FF0D" box="[805,892,187,213]" pageId="23" pageNumber="224">Naboba</collectingCounty>
zone, top of
<collectingMunicipality id="25040F60FFA9FFDD6664FF5FFB26FF0D" box="[1048,1154,187,213]" pageId="23" pageNumber="224">Meoyaca</collectingMunicipality>
,
<location id="C000C3C1FFA9FFDD66F3FF5FFA94FF0D" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA9FFDD66F3FF5FFA94FF0D" box="[1167,1328,187,213]" county="Naboba" latitude="10.818889" longLatPrecision="1" longitude="-73.63295" municipality="Meoyaca" name="Rio Donachui" pageId="23" pageNumber="224">Rio Donachuí</location>
, northern extension, under rocks, at the lower edge of the
<location id="C000C3C1FFA9FFDD60A9FF3BFCDFFF21" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA9FFDD60A9FF3BFCDFFF21" box="[725,891,223,249]" county="Naboba" latitude="10.818889" longLatPrecision="1" longitude="-73.63295" municipality="Meoyaca" name="Pico La Reina" pageId="23" pageNumber="224">Pico La Reina</location>
/
<location id="C000C3C1FFA9FFDD61E5FF3BFBDCFF21" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA9FFDD61E5FF3BFBDCFF21" box="[921,1144,223,249]" county="Naboba" latitude="10.818889" longLatPrecision="1" longitude="-73.63295" municipality="Meoyaca" name="Pico Ojeda Glacier" pageId="23" pageNumber="224">Pico Ojeda Glacier</location>
,
<quantity id="022738FFFFA9FFDD66FBFF3BFB46FF21" box="[1159,1250,223,249]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="23" pageNumber="224" unit="m" value="4800.0">
<elevation id="4EF27229FFA9FFDD66FBFF3BFB46FF21" box="[1159,1250,223,249]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="23" pageNumber="224" unit="m" value="4800.0">4800 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFA9FFDD6684FF04FA3CFF23" box="[1272,1432,224,251]" degrees="10" direction="north" minutes="49" orientation="latitude" pageId="23" pageNumber="224" precision="1" seconds="08.0" value="10.818889">10°4908.0″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFA9FFDD62EBFEE0FE99FEC7" box="[151,317,260,287]" degrees="73" direction="west" minutes="37" orientation="longitude" pageId="23" pageNumber="224" precision="1" seconds="58.6" value="-73.63295">73°3758.6″W</geoCoordinate>
],
<date id="B161B3DAFFA9FFDD6332FEE7FE09FEC5" box="[334,429,259,285]" pageId="23" pageNumber="224" value="1973-01-09">
<collectingDate id="A1254A32FFA9FFDD6332FEE7FE09FEC5" box="[334,429,259,285]" pageId="23" pageNumber="224" value="1973-01-09">9.I.1973</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA9FFDD63C4FEE7FE44FEC5" box="[440,480,259,285]" count="1" pageId="23" pageNumber="224" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2691)
</materialsCitation>
;
<materialsCitation id="75B79F47FFA9FFDD6089FEE7FCFEFE99" collectingDate="1973-03-08" collectorName="Lago Mavaria" country="Colombia" county="Naboba" elevation="4008" latitude="10.818889" location="Rio Donachui" longLatPrecision="1" longitude="-73.63295" municipality="Donachui" pageId="23" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Cesar" typeStatus="paratype">
same locality, but
<collectingMunicipality id="25040F60FFA9FFDD61C1FEE7FB89FEC5" box="[957,1069,259,285]" pageId="23" pageNumber="224">Donachuí</collectingMunicipality>
,
<location id="C000C3C1FFA9FFDD664AFEE7FB71FEC5" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA9FFDD664AFEE7FB71FEC5" box="[1078,1237,259,285]" municipality="Donachui" name="Rio Donachui" pageId="23" pageNumber="224">Río Donachuí</location>
, north tributary of
<collectorName id="682AF0CCFFA9FFDD62EBFEC3FE9CFE99" box="[151,312,295,321]" pageId="23" pageNumber="224">Lago Mavaria</collectorName>
,
<quantity id="022738FFFFA9FFDD633FFEC3FE3EFE99" box="[323,410,295,321]" metricMagnitude="3" metricUnit="m" metricValue="4.008" pageId="23" pageNumber="224" unit="m" value="4008.0">
<elevation id="4EF27229FFA9FFDD633FFEC3FE3EFE99" box="[323,410,295,321]" metricMagnitude="3" metricUnit="m" metricValue="4.008" pageId="23" pageNumber="224" unit="m" value="4008.0">4008 m</elevation>
</quantity>
,
<date id="B161B3DAFFA9FFDD63DAFEC3FDBCFE99" box="[422,536,295,321]" pageId="23" pageNumber="224" value="1973-03-08">
<collectingDate id="A1254A32FFA9FFDD63DAFEC3FDBCFE99" box="[422,536,295,321]" pageId="23" pageNumber="224" value="1973-03-08">8.III.1973</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA9FFDD6058FEC3FDEAFE99" box="[548,590,295,321]" count="1" pageId="23" pageNumber="224" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2694)
</materialsCitation>
.
<materialsCitation id="75B79F47FFA9FFDD6119FEC3FA31FEBD" collectingDate="1929-02-02" collectorName="S. W. Paramo &amp; P. Dalington" country="Colombia" county="Cienaga" elevation="3657" location="Sierra Nevada de Santa Marta" municipality="Santa Marta" pageId="23" pageNumber="224" specimenCode="MCZ-IZ 170963, 170964, 170965" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Cesar" typeStatus="paratype">
<collectingCounty id="2C01ED96FFA9FFDD6119FEC3FC60FE99" box="[869,964,295,321]" pageId="23" pageNumber="224">Ciénaga</collectingCounty>
,
<quantity id="022738FFFFA9FFDD61B2FEC2FBA3FE99" box="[974,1031,294,321]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="224" unit="km" value="5.0">5 km</quantity>
N from
<collectingMunicipality id="25040F60FFA9FFDD661BFEC3FB56FE99" box="[1127,1266,295,321]" pageId="23" pageNumber="224">Santa Marta</collectingMunicipality>
,
<location id="C000C3C1FFA9FFDD6681FEC3FEE0FEBD" LSID="urn:lsid:plazi:treatment:4D76240CFFA8FFD462EBFADFFB5CFDE5:C000C3C1FFA9FFDD6681FEC3FEE0FEBD" country="Colombia" county="Cienaga" municipality="Santa Marta" name="Sierra Nevada de Santa Marta" pageId="23" pageNumber="224" stateProvince="Cesar">Sierra Nevada de Santa Marta</location>
,
<collectorName id="682AF0CCFFA9FFDD6333FEAFFE4EFEBD" box="[335,490,331,357]" pageId="23" pageNumber="224">S. W. Páramo</collectorName>
,
<quantity id="022738FFFFA9FFDD6389FEAEFDE2FEBD" box="[501,582,330,357]" metricMagnitude="3" metricUnit="m" metricValue="3.657" pageId="23" pageNumber="224" unit="m" value="3657.0">
<elevation id="4EF27229FFA9FFDD6389FEAEFDE2FEBD" box="[501,582,330,357]" metricMagnitude="3" metricUnit="m" metricValue="3.657" pageId="23" pageNumber="224" unit="m" value="3657.0">3657m</elevation>
</quantity>
,
<collectorName id="682AF0CCFFA9FFDD602DFEAFFD7FFEBD" box="[593,731,331,357]" pageId="23" pageNumber="224">P. Dalington</collectorName>
coll.,
<date id="B161B3DAFFA9FFDD615DFEAFFC2FFEBD" box="[801,907,331,357]" pageId="23" pageNumber="224" value="1929-02-02">
<collectingDate id="A1254A32FFA9FFDD615DFEAFFC2FFEBD" box="[801,907,331,357]" pageId="23" pageNumber="224" value="1929-02-02">2.II.1929</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFA9FFDD61EAFEAFFC66FEBD" box="[918,962,331,357]" count="1" pageId="23" pageNumber="224" type="male">1 ♂</specimenCount>
,
<specimenCount id="D3D95E93FFA9FFDD61B2FEAFFC5CFEBD" box="[974,1016,331,357]" count="2" pageId="23" pageNumber="224" type="female">2 ♀</specimenCount>
(
<specimenCode id="95793D61FFA9FFDD667AFEAFFB63FEBD" box="[1030,1223,331,357]" collectionCode="MCZ-IZ" pageId="23" pageNumber="224">MCZ-IZ 170963</specimenCode>
,
<specimenCode id="95793D61FFA9FFDD66AEFEAFFA8DFEBD" box="[1234,1321,331,357]" collectionCode="MCZ-IZ" pageId="23" pageNumber="224">170964</specimenCode>
,
<specimenCode id="95793D61FFA9FFDD6748FEAEFA29FEBD" box="[1332,1421,330,357]" collectionCode="MCZ-IZ" pageId="23" pageNumber="224">170965</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA9FFDD62BBFE8BFE4FFE75" pageId="23" pageNumber="224" type="etymology">
<paragraph id="C560951AFFA9FFDD62BBFE8BFE4FFE75" blockId="23.[151,1437,150,1869]" pageId="23" pageNumber="224">
<emphasis id="F7AB4908FFA9FFDD62BBFE8BFEEAFE51" bold="true" box="[199,334,367,393]" pageId="23" pageNumber="224">Etymology.</emphasis>
The specific epithet alludes to the high altitudes of the
<typeStatus id="1A642BB8FFA9FFDD61A4FE8BFBACFE51" box="[984,1032,367,393]" pageId="23" pageNumber="224">type</typeStatus>
locality in Sierra
<collectingRegion id="071B5BF8FFA9FFDD66A7FE8BFA95FE51" box="[1243,1329,367,393]" country="United States of America" name="Nevada" pageId="23" pageNumber="224">Nevada</collectingRegion>
de Santa Marta. It is a noun in genitive.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA9FFDD62BBFE52FB6DFD15" pageId="23" pageNumber="224" type="diagnosis">
<paragraph id="C560951AFFA9FFDD62BBFE52FB6DFD15" blockId="23.[151,1437,150,1869]" pageId="23" pageNumber="224">
<emphasis id="F7AB4908FFA9FFDD62BBFE52FEE6FE08" bold="true" box="[199,322,438,464]" pageId="23" pageNumber="224">Diagnosis.</emphasis>
Males can be distinguished from those of other species of
<taxonomicName id="02DFEE99FFA9FFDD6190FE53FBBBFE08" authorityName="Keyserling" authorityYear="1891" box="[1004,1055,439,464]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="F7AB4908FFA9FFDD6190FE53FBBBFE08" box="[1004,1055,439,464]" italics="true" pageId="23" pageNumber="224">Josa</emphasis>
</taxonomicName>
by the following combination of characters: bifid external section of the conductor with anterior branch shorter than posterior, which is long and sharp (
<figureCitation id="5DE4899FFFA9FFDD6298FE1BFED4FDC1" box="[228,368,511,537]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19AB</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD6301FE1BFE75FDC1" box="[381,465,511,537]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20AB</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD63A1FE1BFDDCFDC1" box="[477,632,511,537]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21AB, D, H</figureCitation>
); embolus describing a loop around basal process, with a thin and long laminar projection (
<figureCitation id="5DE4899FFFA9FFDD6307FDC7FE4CFDE5" box="[379,488,547,573]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">Figs 20C</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD638BFDC7FDE7FDE5" box="[503,579,547,573]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21HI</figureCitation>
), and bifid median apophysis with many small denticles on external edge (
<figureCitation id="5DE4899FFFA9FFDD62E3FDA3FEA5FDB9" box="[159,257,583,609]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">Fig. 21F</figureCitation>
, see inset). Females of this species are distinguished from those of other species of
<taxonomicName id="02DFEE99FFA9FFDD66C8FDA3FB43FDB8" authorityName="Keyserling" authorityYear="1891" box="[1204,1255,583,608]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
<emphasis id="F7AB4908FFA9FFDD66C8FDA3FB43FDB8" box="[1204,1255,583,608]" italics="true" pageId="23" pageNumber="224">Josa</emphasis>
</taxonomicName>
by the shape of the posterior sclerotized projections of lateral lobes, which are narrow, handle-like, arched, and with deep rounded anterior depressions (
<figureCitation id="5DE4899FFFA9FFDD63F5FD6BFE56FD71" box="[393,498,655,681]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19E</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD6383FD6BFD96FD71" box="[511,562,655,681]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20D</figureCitation>
), the median field visible in dorsal view (
<figureCitation id="5DE4899FFFA9FFDD6677FD6BFBD0FD71" box="[1035,1140,655,681]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19E</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD66FDFD6BFB6BFD71" box="[1153,1231,655,681]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="23" pageNumber="224">22EF</figureCitation>
), small, elongated spermathecae, laterally positioned at the median region of the epigyne (
<figureCitation id="5DE4899FFFA9FFDD61CBFD57FB9FFD15" box="[951,1083,691,717]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD663AFD57FB36FD15" box="[1094,1170,691,717]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD66E1FD57FB19FD15" box="[1181,1213,691,717]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="23" pageNumber="224">22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA9FFD462BBFD32FA28FF21" lastPageId="30" lastPageNumber="231" pageId="23" pageNumber="224" type="description">
<paragraph id="C560951AFFA9FFDD62BBFD32FCA8FA19" blockId="23.[151,1437,150,1869]" pageId="23" pageNumber="224">
<emphasis id="F7AB4908FFA9FFDD62BBFD32FEFDFD28" bold="true" box="[199,345,726,752]" pageId="23" pageNumber="224">Description.</emphasis>
<emphasis id="F7AB4908FFA9FFDD6319FD33FE3BFD29" box="[357,415,727,753]" italics="true" pageId="23" pageNumber="224">Male</emphasis>
(
<typeStatus id="1A642BB8FFA9FFDD63CFFD33FDBDFD29" box="[435,537,727,753]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
, MUSENUV-Ar-2682; vchLAM-260) (
<figureCitation id="5DE4899FFFA9FFDD6199FD33FB9AFD29" box="[997,1086,727,753]" captionStart="FIGURE 17" captionStartId="24.[151,250,1880,1904]" captionTargetBox="[155,1428,182,1847]" captionTargetId="figure-22@24.[151,1436,181,1856]" captionTargetPageId="24" captionText="FIGURE 17. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="23" pageNumber="224">Fig. 17</figureCitation>
): Total length 8.89. Carapace length 4.35, width 3.29, height 1.07. Clypeus length 0.12. Sternum length 2.31, width 1.58. Legs. Leg I: ti length 3.84/width 0.49, mt 3.20. II: ti 3.82/0.52, mt 3.13. III: ti 3.12/0.45, mt 3.29. IV: ti 3.95/0.54, mt 4.11. Spines: legs III, femur d 1-1-1, p 0-1d2ap, r 0-d1-d1; tibia v 2-2-2; metatarsus v 2-0-0. IIIIV, femur d 1-1-1, p 0-d1-d1, r 0-d1- d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-2-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 4.31, width 2.52, spiracleepigastrium 1.72. Color: carapace brown, with two wide longitudinal discontinuous dark brown stripes, extending to fovea and marginally covered by abundant white setae; eyes bordered by black rings, triangular band emerging from posterior side of PLE and PME. Clypeus brown, low with abundant white setae. Chelicerae brown with black spots and fine setae on frontal edge. Endites light brown, light at anterior side. Labium and sternum dark brown. Legs coxae light brown, femora-tarsi brown with some dark patches, patellae less pigmented. All articles with black dots, stripes, and covered by few white and black setae, abundant white setae between coxae. Abdomen with pattern of black patches on lighter brown background, sub-rounded by thin band; anterior region of dorsum medially with a wide longitudinal dark band extended to submedial region, followed by five wide and very short chevrons, lateral regions with many dots. Abdomen covered with few fine white setae, anterior region with a dense bunch of setae. Venter dark gray with many light patches. Palp: tegulum without tegular lobe. Median apophysis very wide, bifid, with anterior hook-shaped tip, posterior region with a denticulated edge (
<figureCitation id="5DE4899FFFA9FFDD607AFAF3FDCEFAE9" box="[518,618,1303,1329]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">Fig. 21F</figureCitation>
). Conductor large, rounded, external section apically bifid (
<figureCitation id="5DE4899FFFA9FFDD6779FAF3FA33FAE9" box="[1285,1431,1303,1329]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs. 19AB</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD62EBFADFFF4FFA8D" box="[151,235,1339,1365]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20AB</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD6284FADFFED4FA8D" box="[248,368,1339,1365]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21AB, D</figureCitation>
), with posterior tip long, sharp, anterior tip shorter with small apical denticles, dorsal section sharp (
<figureCitation id="5DE4899FFFA9FFDD6298FABBFEE8FAA1" box="[228,332,1375,1401]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">Fig. 21B</figureCitation>
). Paramedian apophysis short, heavily sclerotized, sharp, narrow. Embolus apically filiform, with a narrow laminar projection extending to subapical side; basal process small, reticulate, posterior region rounded, with pronounced external keel (
<figureCitation id="5DE4899FFFA9FFDD6386FA43FDD9FA19" box="[506,637,1447,1473]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD60F5FA43FD71FA19" box="[649,725,1447,1473]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD60A3FA43FD5AFA19" box="[735,766,1447,1473]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFA9FFDD62BBFA2FFDD0F895" blockId="23.[151,1437,150,1869]" pageId="23" pageNumber="224">
<emphasis id="F7AB4908FFA9FFDD62BBFA2FFEBFFA3D" box="[199,283,1483,1509]" italics="true" pageId="23" pageNumber="224">Female</emphasis>
(
<typeStatus id="1A642BB8FFA9FFDD6350FA2FFE35FA3D" box="[300,401,1483,1509]" pageId="23" pageNumber="224" type="paratype">paratype</typeStatus>
, MUSENUV-Ar-2683; vchLAM-261). (
<figureCitation id="5DE4899FFFA9FFDD6126FA2FFC0BFA3D" box="[858,943,1483,1509]" captionStart="FIGURE 18" captionStartId="25.[151,250,1879,1903]" captionTargetBox="[158,1429,192,1849]" captionTargetId="figure-17@25.[151,1436,181,1855]" captionTargetPageId="25" captionText="FIGURE 18. Josa olimpica sp. nov., female (MUSENUV-Ar-2683; vchLAM-261): A Habitus, dorsal view; B Carapace; C Habitus, ventral view; D Sternum." pageId="23" pageNumber="224">Fig. 18</figureCitation>
): Total length 11.18. Carapace length 5.98, width 4.16, height 1.72. Clypeus length 0.28. Sternum length 2.90, width 2.20. Legs. Leg I: ti length 3.43/width 0.70, mt 2.86. II: ti 4.12/0.65, mt 3.26. III: ti 3.24/0.63, mt 3.49. IV: ti 4.39/0.60, mt 3.97. Spines: legs III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; tibia v 2-2-2; metatarsus v 2bas. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v p1-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-2-2, p d1-1-0-1, r d1- 1-0-1, IV=III but femur r 1d. Abdomen: length 6.08, width 3.96, spiracleepigastrium 2.15. Color pattern as in male, but more pigmented. Lateral regions of abdomen with many dots. Epigyne: median field transverse, wide, oval in posterior view (
<figureCitation id="5DE4899FFFA9FFDD63E3F923FD81F939" box="[415,549,1735,1761]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="23" pageNumber="224">Fig. 22EF</figureCitation>
). Lateral lobes with sclerotized extensions wide, rounded, with small, rounded concavities at anterior side. Spermathecae small, oval, lateromedially placed. Copulatory ducts long, thin, coiled along longitudinal axis, oriented in parallel, anterior region rounded (
<figureCitation id="5DE4899FFFA9FFDD61D9F8EBFBAAF8F1" box="[933,1038,1807,1833]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">Figs 20E</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD6666F8EBFBD4F8F1" box="[1050,1136,1807,1833]" captionStart="FIGURE 21" captionStartId="28.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@28.[151,1435,181,1756]" captionTargetPageId="28" captionText="FIGURE 21. Josa olimpica sp. nov., male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="23" pageNumber="224">21AB</figureCitation>
). Ducts of accessory bulbs long, convergent (
<figureCitation id="5DE4899FFFA9FFDD631EF8D7FE41F895" box="[354,485,1843,1869]" captionStart="FIGURE 19" captionStartId="26.[151,250,1851,1875]" captionTargetBox="[151,1428,212,1819]" captionTargetId="figure-22@26.[151,1436,181,1828]" captionTargetPageId="26" captionText="FIGURE 19. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum." pageId="23" pageNumber="224">Figs 19EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD638DF8D7FD99F895" box="[497,573,1843,1869]" captionStart="FIGURE 20" captionStartId="27.[151,250,1858,1882]" captionTargetBox="[162,1427,189,1832]" captionTargetId="figure-17@27.[158,1430,181,1834]" captionTargetPageId="27" captionText="FIGURE 20. Josa olimpica sp. nov., male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="23" pageNumber="224">20EF</figureCitation>
,
<figureCitation id="5DE4899FFFA9FFDD603BF8D7FDC3F895" box="[583,615,1843,1869]" captionStart="FIGURE 22" captionStartId="29.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@29.[151,1435,181,1708]" captionTargetPageId="29" captionText="FIGURE 22. Josa olimpica sp. nov., female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view. Abbreviations: AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="23" pageNumber="224">22</figureCitation>
).
</paragraph>
<caption id="91A0C592FFA6FFD262EBF8BCFD3CF84C" pageId="24" pageNumber="225" startId="24.[151,250,1880,1904]" targetBox="[155,1428,182,1847]" targetPageId="24" targetType="figure">
<paragraph id="C560951AFFA6FFD262EBF8BCFD3CF84C" blockId="24.[151,1436,1880,1940]" pageId="24" pageNumber="225">
<emphasis id="F7AB4908FFA6FFD262EBF8BCFE87F8A8" bold="true" box="[151,291,1880,1904]" pageId="24" pageNumber="225">FIGURE 17.</emphasis>
<taxonomicName id="02DFEE99FFA6FFD26351F8BDFE1BF8A8" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[301,447,1880,1904]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
<emphasis id="F7AB4908FFA6FFD26351F8BDFE1BF8A8" box="[301,447,1880,1904]" italics="true" pageId="24" pageNumber="225">Josa olimpica</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFA6FFD263B4F8BCFDB8F8A8" bold="true" box="[456,540,1880,1904]" pageId="24" pageNumber="225">
<taxonomicNameLabel id="EC98F473FFA6FFD263B4F8BCFDB8F8A8" box="[456,540,1880,1904]" pageId="24" pageNumber="225" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2682; vchLAM-260): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFA7FFD362EBF8B3FE50F84B" pageId="25" pageNumber="226" startId="25.[151,250,1879,1903]" targetBox="[158,1429,192,1849]" targetPageId="25" targetType="figure">
<paragraph id="C560951AFFA7FFD362EBF8B3FE50F84B" blockId="25.[151,1436,1879,1939]" pageId="25" pageNumber="226">
<emphasis id="F7AB4908FFA7FFD362EBF8B3FE86F8B7" bold="true" box="[151,290,1879,1903]" pageId="25" pageNumber="226">FIGURE 18.</emphasis>
<taxonomicName id="02DFEE99FFA7FFD36357F8BCFE1FF8B7" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,443,1879,1903]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
<emphasis id="F7AB4908FFA7FFD36357F8BCFE1FF8B7" box="[299,443,1879,1903]" italics="true" pageId="25" pageNumber="226">Josa olimpica</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFA7FFD363BFF8B3FDB2F8B7" bold="true" box="[451,534,1879,1903]" pageId="25" pageNumber="226">
<taxonomicNameLabel id="EC98F473FFA7FFD363BFF8B3FDB2F8B7" box="[451,534,1879,1903]" pageId="25" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (MUSENUV-Ar-2683; vchLAM-261): A Habitus, dorsal view; B Carapace; C Habitus, ventral view; D Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFA4FFD062EBF8DFFBB2F83C" pageId="26" pageNumber="227" startId="26.[151,250,1851,1875]" targetBox="[151,1428,212,1819]" targetPageId="26" targetType="figure">
<paragraph id="C560951AFFA4FFD062EBF8DFFBB2F83C" blockId="26.[151,1436,1851,2020]" pageId="26" pageNumber="227">
<emphasis id="F7AB4908FFA4FFD062EBF8DFFEBBF88C" bold="true" box="[151,287,1851,1876]" pageId="26" pageNumber="227">FIGURE 19.</emphasis>
<taxonomicName id="02DFEE99FFA4FFD06358F8D8FE15F88C" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[292,433,1852,1876]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
<emphasis id="F7AB4908FFA4FFD06358F8D8FE15F88C" box="[292,433,1852,1876]" italics="true" pageId="26" pageNumber="227">Josa olimpica</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFA4FFD063CAF8D8FDA2F88C" bold="true" box="[438,518,1852,1876]" pageId="26" pageNumber="227">
<taxonomicNameLabel id="EC98F473FFA4FFD063CAF8D8FDA2F88C" box="[438,518,1852,1876]" pageId="26" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Prolateral view; D. Apical view. Female genitalia (MUSENUV-Ar-2683; vchLAM-261): E Ventral view (arrow indicates the anterior excavations); F Dorsal view.
<emphasis id="F7AB4908FFA4FFD0606AF867FD12F843" bold="true" box="[534,694,1923,1947]" pageId="26" pageNumber="227">Abbreviations:</emphasis>
AB, accessory bulb; C, conductor; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; St, subtegulum; SD, spermatic duct; T, tegulum.
</paragraph>
</caption>
<caption id="91A0C592FFA5FFD162EBF8A6FBAFF832" pageId="27" pageNumber="228" startId="27.[151,250,1858,1882]" targetBox="[162,1427,189,1832]" targetPageId="27" targetType="figure">
<paragraph id="C560951AFFA5FFD162EBF8A6FBAFF832" blockId="27.[151,1436,1858,2027]" pageId="27" pageNumber="228">
<emphasis id="F7AB4908FFA5FFD162EBF8A6FEBBF882" bold="true" box="[151,287,1858,1882]" pageId="27" pageNumber="228">FIGURE 20.</emphasis>
<taxonomicName id="02DFEE99FFA5FFD16358F8A7FE15F882" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[292,433,1858,1882]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
<emphasis id="F7AB4908FFA5FFD16358F8A7FE15F882" box="[292,433,1858,1882]" italics="true" pageId="27" pageNumber="228">Josa olimpica</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFA5FFD163CAF8A7FDA2F883" bold="true" box="[438,518,1859,1883]" pageId="27" pageNumber="228">
<taxonomicNameLabel id="EC98F473FFA5FFD163CAF8A7FDA2F883" box="[438,518,1859,1883]" pageId="27" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2682; vchLAM-260) left bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2682; vchLAM-261): D Ventral view; E Dorsal view.
<emphasis id="F7AB4908FFA5FFD166B7F882FAC8F8A6" bold="true" box="[1227,1388,1894,1918]" pageId="27" pageNumber="228">Abbreviations:</emphasis>
AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct.
</paragraph>
</caption>
<caption id="91A0C592FFA2FFD662EBF917FEA8F818" pageId="28" pageNumber="229" startId="28.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="28" targetType="figure">
<paragraph id="C560951AFFA2FFD662EBF917FEA8F818" blockId="28.[151,1437,1779,1984]" pageId="28" pageNumber="229">
<emphasis id="F7AB4908FFA2FFD662EBF917FE85F8D4" bold="true" box="[151,289,1779,1804]" pageId="28" pageNumber="229">FIGURE 21.</emphasis>
<taxonomicName id="02DFEE99FFA2FFD66354F910FE13F8D4" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[296,439,1780,1804]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
<emphasis id="F7AB4908FFA2FFD66354F910FE13F8D4" box="[296,439,1780,1804]" italics="true" pageId="28" pageNumber="229">Josa olimpica</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFA2FFD663C2F910FDABF8D4" bold="true" box="[446,527,1780,1804]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="EC98F473FFA2FFD663C2F910FDABF8D4" box="[446,527,1780,1804]" pageId="28" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MACN-Ar 46696; vchLAM-262) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small denticles); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details.
<emphasis id="F7AB4908FFA2FFD6611DF8BBFBA6F8AF" bold="true" box="[865,1026,1887,1911]" pageId="28" pageNumber="229">Abbreviations:</emphasis>
C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum.
</paragraph>
</caption>
<caption id="91A0C592FFA3FFD762EBF927FED0F890" pageId="29" pageNumber="230" startId="29.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="29" targetType="figure">
<paragraph id="C560951AFFA3FFD762EBF927FED0F890" blockId="29.[151,1437,1731,1864]" pageId="29" pageNumber="230">
<emphasis id="F7AB4908FFA3FFD762EBF927FE87F904" bold="true" box="[151,291,1731,1756]" pageId="29" pageNumber="230">FIGURE 22.</emphasis>
<taxonomicName id="02DFEE99FFA3FFD76350F920FE19F904" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[300,445,1732,1756]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="olimpica" status="sp. nov.">
<emphasis id="F7AB4908FFA3FFD76350F920FE19F904" box="[300,445,1732,1756]" italics="true" pageId="29" pageNumber="230">Josa olimpica</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFA3FFD763BAF920FDBEF904" bold="true" box="[454,538,1732,1756]" pageId="29" pageNumber="230">
<taxonomicNameLabel id="EC98F473FFA3FFD763BAF920FDBEF904" box="[454,538,1732,1756]" pageId="29" pageNumber="230" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (MACN-Ar 46697; vchLAM-263) genitalia, SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Accessory bulb; E Posterior-dorsal view; F Posterior-ventral view.
<emphasis id="F7AB4908FFA3FFD76687F903FA38F927" bold="true" box="[1275,1436,1767,1791]" pageId="29" pageNumber="230">Abbreviations:</emphasis>
AB, accessory bulb; dAB, duct of accessory bulb; CD, copulatory duct; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca.
</paragraph>
</caption>
<paragraph id="C560951AFFA3FFD462BBF87EFA28FF21" blockId="29.[151,1437,1946,2009]" lastBlockId="30.[151,1437,150,573]" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230">
<emphasis id="F7AB4908FFA3FFD762BBF87EFEE5F86C" bold="true" box="[199,321,1946,1972]" pageId="29" pageNumber="230">Variation:</emphasis>
Males (n=5): total length: 7.5111.35; carapace length: 3.754.35, width: 2.723.29; abdomen length: 4.315.38, width: 1.592.62. Females (n=5): total length: 11.1811.35; carapace length: 5.885.98, width: 4.064.16; abdomen length: 5.386.08, width: 3.653.96. The pigmentation and abdominal spots are variable across specimens, the carapace can be darker, and the abdominal spots more defined. The anterior tip of apical region of the external section of the conductor seems broken in some specimens, resulting in an entire and sharp conductor tip.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA0FFD462BBFEE6FB5CFDE5" pageId="30" pageNumber="231" type="distribution">
<paragraph id="C560951AFFA0FFD462BBFEE6FBD7FEC5" blockId="30.[151,1437,150,573]" box="[199,1139,258,285]" pageId="30" pageNumber="231">
<emphasis id="F7AB4908FFA0FFD462BBFEE6FEC5FEC4" bold="true" box="[199,353,258,284]" pageId="30" pageNumber="231">Distribution.</emphasis>
Only known from Sierra Nevada de Santa Marta,
<collectingCountry id="BDC8D58AFFA0FFD461E9FEE7FBA7FEC5" box="[917,1027,259,285]" name="Colombia" pageId="30" pageNumber="231">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFFA0FFD4666FFEE7FBC2FEC5" box="[1043,1126,259,285]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="30" pageNumber="231">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFA0FFD462BBFEC2FB5CFDE5" blockId="30.[151,1437,150,573]" pageId="30" pageNumber="231">
<emphasis id="F7AB4908FFA0FFD462BBFEC2FE68FE99" bold="true" box="[199,460,294,321]" pageId="30" pageNumber="231">Ecological comments.</emphasis>
When the high-altitude specimens were collected at
<quantity id="022738FFFFA0FFD4664EFEC2FB08FE99" box="[1074,1196,294,321]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="30" pageNumber="231" unit="m" value="5500.0">5500 masl</quantity>
, and at several other sites nearby, it was noted that this species was not only present but also abundant and active, a very fast runner on the rocks and snow. The specimens found at
<quantity id="022738FFFFA0FFD460C3FE8AFC91FE51" box="[703,821,366,393]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="30" pageNumber="231" unit="m" value="5500.0">5500 masl</quantity>
were collected from beneath rocks, and the specimen MUSENUV-Ar-2696, found at an altitude of 5275 meters, was caught during the day, near noon, running on the surface of a large rock surrounded by snow and glacial ice; there were no loose rocks for shelter. This species appears to be very abundant in the Páramo of the Sierra
<collectingRegion id="071B5BF8FFA0FFD46164FE3FFCCAFE2D" box="[792,878,475,501]" country="United States of America" name="Nevada" pageId="30" pageNumber="231">Nevada</collectingRegion>
de Santa Marta, which may explain the presence of some specimens at high altitudes. However, systematic field sampling is necessary to determine if there are established populations at altitudes above 5000 meters or if these are merely stochastic occurrences.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,387 @@
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<mods:title id="9C1AE8869BD78E917E6ACFF1AB5BE500">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
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<mods:namePart id="7CE08896FBE11AF0B691AEC5230C117B">Martínez, Leonel</mods:namePart>
<mods:affiliation id="69FE04FE9875997C0CEE1E6439534B62">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
</mods:name>
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<mods:namePart id="7E4B0A74F13DE009DB60E907668E94DC">Kochalka, John A.</mods:namePart>
<mods:affiliation id="C84B1649D51849637777B1B2AA745B22">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
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<mods:affiliation id="29C1AA22E8B72AC78E2EBC4C85B9C1EF">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
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<mods:number id="9CF1CE7481B6B51BD87F611186C79B09">5566</mods:number>
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<mods:url id="EA73082A3168A182C1CFA90E8AF7C6E2">https://doi.org/10.11646/zootaxa.5566.2.1</mods:url>
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<mods:classification id="4F7592730B65788B406F461610CE37D8">journal article</mods:classification>
<mods:identifier id="CBBD1C9638ED652922F09D623B5EA865" type="DOI">10.11646/zootaxa.5566.2.1</mods:identifier>
<mods:identifier id="9CA82574C2584F5F750990502E8352EA" type="ISSN">1175-5326</mods:identifier>
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<treatment id="4D76240CFFB1FFDC62EBFC13FBD7FB31" LSID="urn:lsid:plazi:treatment:4D76240CFFB1FFDC62EBFC13FBD7FB31" httpUri="http://treatment.plazi.org/id/4D76240CFFB1FFDC62EBFC13FBD7FB31" lastPageId="22" lastPageNumber="223" pageId="15" pageNumber="216">
<subSubSection id="8DC5C691FFB1FFC562EBFC13FE0EFBC9" box="[151,426,1014,1041]" pageId="15" pageNumber="216" type="nomenclature">
<paragraph id="C560951AFFB1FFC562EBFC13FE0EFBC9" blockId="15.[151,426,1014,1077]" box="[151,426,1014,1041]" pageId="15" pageNumber="216">
<heading id="9E282276FFB1FFC562EBFC13FE0EFBC9" bold="true" box="[151,426,1014,1041]" fontSize="11" level="1" pageId="15" pageNumber="216" reason="1">
<emphasis id="F7AB4908FFB1FFC562EBFC13FE0EFBC9" bold="true" box="[151,426,1014,1041]" pageId="15" pageNumber="216">
<taxonomicName id="02DFEE99FFB1FFC562EBFC13FEE1FBC9" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[151,325,1014,1041]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFB1FFC562EBFC13FEE1FBC9" bold="true" box="[151,325,1014,1041]" italics="true" pageId="15" pageNumber="216">Josa nadineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EC98F473FFB1FFC56330FC13FE0EFBC9" box="[332,426,1015,1041]" pageId="15" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB1FFC562EBFBFFFE9CFBED" box="[151,312,1051,1077]" pageId="15" pageNumber="216" type="description">
<paragraph id="C560951AFFB1FFC562EBFBFFFE9CFBED" blockId="15.[151,426,1014,1077]" box="[151,312,1051,1077]" pageId="15" pageNumber="216">
<figureCitation id="5DE4899FFFB1FFC562EBFBFFFE9CFBED" box="[151,312,1051,1077]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStart-5="FIGURE 16" captionStartId-0="16.[151,250,1882,1906]" captionStartId-1="17.[151,250,1882,1906]" captionStartId-2="18.[151,250,1836,1860]" captionStartId-3="19.[151,250,1866,1890]" captionStartId-4="20.[151,250,1779,1803]" captionStartId-5="21.[151,250,1731,1755]" captionTargetBox-0="[159,1427,189,1858]" captionTargetBox-1="[159,1423,186,1853]" captionTargetBox-2="[151,1412,182,1812]" captionTargetBox-3="[155,1432,185,1840]" captionTargetBox-4="[151,1435,181,1756]" captionTargetBox-5="[151,1435,181,1708]" captionTargetId-0="figure-22@16.[151,1436,181,1858]" captionTargetId-1="figure-17@17.[151,1436,181,1858]" captionTargetId-2="figure-22@18.[151,1436,181,1812]" captionTargetId-3="figure-17@19.[151,1435,181,1842]" captionTargetId-4="figure-22@20.[151,1435,181,1756]" captionTargetId-5="figure-17@21.[151,1435,181,1708]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionText-0="FIGURE 11. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 12. Josa nadineae sp. nov., female (MUSENUV-Ar-2679; vchLAM-251): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-2="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." captionText-3="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-4="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." captionText-5="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Figures 1116</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB1FFC562EBFBB8FCE6FAAB" pageId="15" pageNumber="216" type="materials_examined">
<paragraph id="C560951AFFB1FFC562EBFBB8FCE6FAAB" blockId="15.[151,1437,1116,2043]" pageId="15" pageNumber="216">
<emphasis id="F7AB4908FFB1FFC562EBFBB8FEE1FBAE" bold="true" box="[151,325,1116,1142]" pageId="15" pageNumber="216">Type material.</emphasis>
<materialsCitation id="75B79F47FFB1FFC56331FBB8FC7FFB66" collectingDate="1975-05-09" collectorName="Loma de la Cebolleta &amp; J. Kochalka" country="Colombia" county="Cienaga" elevation="3505" latitude="10.902499" location="Loma de la Cebolleta" longLatPrecision="21" longitude="-73.924164" municipality="Parque Natural Nacional Sierra Nevada de Santa Marta" pageId="15" pageNumber="216" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="holotype">
<typeStatus id="1A642BB8FFB1FFC56331FBB8FE1EFBAE" box="[333,442,1116,1142]" pageId="15" pageNumber="216" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="BDC8D58AFFB1FFC563B8FBB8FDF9FBAE" box="[452,605,1116,1142]" name="Colombia" pageId="15" pageNumber="216">COLOMBIA</collectingCountry>
.
<collectingRegion id="071B5BF8FFB1FFC56014FBB8FD4DFBAE" box="[616,745,1116,1142]" country="Colombia" name="Magdalena" pageId="15" pageNumber="216">Magdalena</collectingRegion>
:
<collectingCounty id="2C01ED96FFB1FFC56088FBB8FCF7FBAE" box="[756,851,1116,1142]" pageId="15" pageNumber="216">Ciénaga</collectingCounty>
, San Pedro,
<collectingMunicipality id="25040F60FFB1FFC561A3FBB9FEE5FB43" pageId="15" pageNumber="216">Parque Natural Nacional Sierra Nevada de Santa Marta</collectingMunicipality>
,
<collectorName id="682AF0CCFFB1FFC56336FB65FD9CFB42" box="[330,568,1152,1179]" pageId="15" pageNumber="216">
<location id="C000C3C1FFB1FFC56336FB65FD9CFB42" LSID="urn:lsid:plazi:treatment:4D76240CFFB1FFDC62EBFC13FBD7FB31:C000C3C1FFB1FFC56336FB65FD9CFB42" box="[330,568,1152,1179]" country="Colombia" county="Cienaga" latitude="10.902499" longLatPrecision="21" longitude="-73.924164" municipality="Parque Natural Nacional Sierra Nevada de Santa Marta" name="Loma de la Cebolleta" pageId="15" pageNumber="216" stateProvince="Magdalena">Loma de la Cebolleta</location>
</collectorName>
, dead mats of curly grass, manual caught,
<quantity id="022738FFFFB1FFC5667AFB64FBFFFB43" box="[1030,1115,1152,1179]" metricMagnitude="3" metricUnit="m" metricValue="3.505" pageId="15" pageNumber="216" unit="m" value="3505.0">
<elevation id="4EF27229FFB1FFC5667AFB64FBFFFB43" box="[1030,1115,1152,1179]" metricMagnitude="3" metricUnit="m" metricValue="3.505" pageId="15" pageNumber="216" unit="m" value="3505.0">3505 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFB1FFC56610FB65FB52FB44" box="[1132,1270,1153,1180]" degrees="10" direction="north" minutes="54" orientation="latitude" pageId="15" pageNumber="216" precision="15" seconds="09" value="10.902499">10°5409″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFB1FFC5677CFB65FA36FB44" box="[1280,1426,1153,1180]" degrees="73" direction="west" minutes="55" orientation="longitude" pageId="15" pageNumber="216" precision="15" seconds="27" value="-73.924164">73°5527″W</geoCoordinate>
],
<collectorName id="682AF0CCFFB1FFC562EBFB41FE86FB66" box="[151,290,1188,1215]" pageId="15" pageNumber="216">J. Kochalka</collectorName>
leg.,
<date id="B161B3DAFFB1FFC56312FB40FE71FB67" box="[366,469,1188,1215]" pageId="15" pageNumber="216" value="1975-05-09">
<collectingDate id="A1254A32FFB1FFC56312FB40FE71FB67" box="[366,469,1188,1215]" pageId="15" pageNumber="216" value="1975-05-09">9.V.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFB1FFC5639BFB40FDBCFB67" box="[487,536,1188,1215]" count="1" pageId="15" pageNumber="216" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2678; vchLAM-249)
</materialsCitation>
.
<materialsCitation id="75B79F47FFB1FFC56190FB41FC9AFAAB" collectingDate="1975-03-05" collectingDateMax="1975-05-08" collectingDateMin="1975-03-05" collectorName="J. Kochalka &amp; Serrania Dunanchucua &amp; Norte Grande" country="Colombia" county="Parque Natural Nacional Sierra Nevada de Santa Marta" elevation="4008" latitude="10.900499" location="Rio Tucurinca" longLatPrecision="1" longitude="-73.8885" municipality="Lago Mavaria" pageId="15" pageNumber="216" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Magdalena" typeStatus="paratype">
<typeStatus id="1A642BB8FFB1FFC56190FB41FBF9FB67" box="[1004,1117,1189,1215]" pageId="15" pageNumber="216" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="BDC8D58AFFB1FFC56612FB40FAA3FB66" box="[1134,1287,1188,1214]" name="Colombia" pageId="15" pageNumber="216">COLOMBIA</collectingCountry>
.
<collectingRegion id="071B5BF8FFB1FFC56764FB40FA3DFB66" box="[1304,1433,1188,1214]" country="Colombia" name="Magdalena" pageId="15" pageNumber="216">Magdalena</collectingRegion>
: Ciénaga, San Pedro,
<collectingCounty id="2C01ED96FFB1FFC56301FB2DFC46FB3B" box="[381,994,1224,1251]" pageId="15" pageNumber="216">Parque Natural Nacional Sierra Nevada de Santa Marta</collectingCounty>
, north of
<collectingMunicipality id="25040F60FFB1FFC56635FB2DFB4DFB3A" box="[1097,1257,1224,1251]" pageId="15" pageNumber="216">Lago Mavaria</collectingMunicipality>
, manual caught,
<quantity id="022738FFFFB1FFC562EBFB08FF54FADF" box="[151,240,1260,1287]" metricMagnitude="3" metricUnit="m" metricValue="4.008" pageId="15" pageNumber="216" unit="m" value="4008.0">
<elevation id="4EF27229FFB1FFC562EBFB08FF54FADF" box="[151,240,1260,1287]" metricMagnitude="3" metricUnit="m" metricValue="4.008" pageId="15" pageNumber="216" unit="m" value="4008.0">4008 m</elevation>
</quantity>
,
<collectorName id="682AF0CCFFB1FFC56281FB09FE27FADE" box="[253,387,1260,1287]" pageId="15" pageNumber="216">J. Kochalka</collectorName>
leg.,
<date id="B161B3DAFFB1FFC563B9FB08FD8FFADF" box="[453,555,1260,1287]" pageId="15" pageNumber="216" value="1975-05-08">
<collectingDate id="A1254A32FFB1FFC563B9FB08FD8FFADF" box="[453,555,1260,1287]" pageId="15" pageNumber="216" value="1975-05-08">8.V.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFB1FFC56045FB08FDC1FADF" box="[569,613,1260,1287]" count="1" pageId="15" pageNumber="216" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2679; vchLAM-251),
<specimenCount id="D3D95E93FFB1FFC56656FB08FBF1FADF" box="[1066,1109,1260,1287]" count="1" pageId="15" pageNumber="216" type="male">1 ♂</specimenCount>
<specimenCount id="D3D95E93FFB1FFC56622FB08FB2EFADF" box="[1118,1162,1260,1287]" count="1" pageId="15" pageNumber="216" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2680);
<collectorName id="682AF0CCFFB1FFC562EBFAF4FE3EFAF2" box="[151,410,1296,1322]" pageId="15" pageNumber="216">Serranía Dunanchucua</collectorName>
, northern peak of
<collectorName id="682AF0CCFFB1FFC56016FAF5FCA3FAF2" box="[618,775,1296,1323]" pageId="15" pageNumber="216">Norte Grande</collectorName>
,
<quantity id="022738FFFFB1FFC56168FAF4FCC9FAF3" box="[788,877,1296,1323]" metricMagnitude="3" metricUnit="m" metricValue="3.7969999999999997" pageId="15" pageNumber="216" unit="m" value="3797.0">
<elevation id="4EF27229FFB1FFC56168FAF4FCC9FAF3" box="[788,877,1296,1323]" metricMagnitude="3" metricUnit="m" metricValue="3.7969999999999997" pageId="15" pageNumber="216" unit="m" value="3797.0">3797 m</elevation>
</quantity>
,
<date id="B161B3DAFFB1FFC56106FAF4FC48FAF3" box="[890,1004,1296,1323]" pageId="15" pageNumber="216" value="1975-03-05">
<collectingDate id="A1254A32FFB1FFC56106FAF4FC48FAF3" box="[890,1004,1296,1323]" pageId="15" pageNumber="216" value="1975-03-05">5.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFB1FFC56185FAF4FB80FAF3" box="[1017,1060,1296,1323]" count="1" pageId="15" pageNumber="216" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2681);
<quantity id="022738FFFFB1FFC56743FAF4FA33FAF3" box="[1343,1431,1296,1323]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="15" pageNumber="216" unit="m" value="3900.0">
<elevation id="4EF27229FFB1FFC56743FAF4FA33FAF3" box="[1343,1431,1296,1323]" metricMagnitude="3" metricUnit="m" metricValue="3.9" pageId="15" pageNumber="216" unit="m" value="3900.0">3900 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFB1FFC562E2FAD1FE99FA88" box="[158,317,1333,1360]" degrees="10" direction="north" minutes="54" orientation="latitude" pageId="15" pageNumber="216" precision="1" seconds="01.8" value="10.900499">10°5401.8″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFB1FFC56335FAD1FE4AFA88" box="[329,494,1333,1360]" degrees="73" direction="west" minutes="53" orientation="longitude" pageId="15" pageNumber="216" precision="1" seconds="18.6" value="-73.8885">73°5318.6″W</geoCoordinate>
],
<date id="B161B3DAFFB1FFC5607CFAD0FDD7FA97" box="[512,627,1332,1359]" pageId="15" pageNumber="216" value="1975-03-07">
<collectingDate id="A1254A32FFB1FFC5607CFAD0FDD7FA97" box="[512,627,1332,1359]" pageId="15" pageNumber="216" value="1975-03-07">7.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFB1FFC56002FAD0FD0CFA97" box="[638,680,1332,1359]" count="1" pageId="15" pageNumber="216" type="male">1 ♂</specimenCount>
(MACN-Ar 46692; vchLAM-250);
<location id="C000C3C1FFB1FFC5663DFAD0FB40FA96" LSID="urn:lsid:plazi:treatment:4D76240CFFB1FFDC62EBFC13FBD7FB31:C000C3C1FFB1FFC5663DFAD0FB40FA96" box="[1089,1252,1332,1358]" country="Colombia" county="Parque Natural Nacional Sierra Nevada de Santa Marta" latitude="10.900499" longLatPrecision="1" longitude="-73.8885" municipality="Lago Mavaria" name="Rio Tucurinca" pageId="15" pageNumber="216" stateProvince="Magdalena">Río Tucurinca</location>
,
<location id="C000C3C1FFB1FFC56692FAD1FA3CFA96" LSID="urn:lsid:plazi:treatment:4D76240CFFB1FFDC62EBFC13FBD7FB31:C000C3C1FFB1FFC56692FAD1FA3CFA96" box="[1262,1432,1332,1359]" country="Colombia" county="Parque Natural Nacional Sierra Nevada de Santa Marta" latitude="10.900499" longLatPrecision="1" longitude="-73.8885" municipality="Lago Mavaria" name="Lago Arucuina" pageId="15" pageNumber="216" stateProvince="Magdalena">Lago Arucuina</location>
,
<quantity id="022738FFFFB1FFC562EBFABCFF4AFAAB" box="[151,238,1368,1395]" metricMagnitude="3" metricUnit="m" metricValue="4.176" pageId="15" pageNumber="216" unit="m" value="4176.0">
<elevation id="4EF27229FFB1FFC562EBFABCFF4AFAAB" box="[151,238,1368,1395]" metricMagnitude="3" metricUnit="m" metricValue="4.176" pageId="15" pageNumber="216" unit="m" value="4176.0">4176 m</elevation>
</quantity>
,
<date id="B161B3DAFFB1FFC56286FABCFEDFFAAB" box="[250,379,1368,1395]" pageId="15" pageNumber="216" value="1975-03-12">
<collectingDate id="A1254A32FFB1FFC56286FABCFEDFFAAB" box="[250,379,1368,1395]" pageId="15" pageNumber="216" value="1975-03-12">12.III.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFB1FFC563FAFABCFE14FAAB" box="[390,432,1368,1395]" count="1" pageId="15" pageNumber="216" type="female">1 ♀</specimenCount>
(MACN-Ar 46693; vchLAM-252)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB1FFC562BBFA98FDE9FA63" pageId="15" pageNumber="216" type="etymology">
<paragraph id="C560951AFFB1FFC562BBFA98FDE9FA63" blockId="15.[151,1437,1116,2043]" pageId="15" pageNumber="216">
<emphasis id="F7AB4908FFB1FFC562BBFA98FEEAFA4E" bold="true" box="[199,334,1404,1430]" pageId="15" pageNumber="216">Etymology.</emphasis>
The specific name is after the Canadian arachnologist Nadine Dupérré, in recognition of her many contributions to the spiders taxonomy.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB1FFC562BBFA20FEBEF8FA" pageId="15" pageNumber="216" type="diagnosis">
<paragraph id="C560951AFFB1FFC562BBFA20FEBEF8FA" blockId="15.[151,1437,1116,2043]" pageId="15" pageNumber="216">
<emphasis id="F7AB4908FFB1FFC562BBFA20FEE6FA06" bold="true" box="[199,322,1476,1502]" pageId="15" pageNumber="216">Diagnosis.</emphasis>
Males resemble those of
<taxonomicName id="02DFEE99FFB1FFC56013FA21FC79FA06" authority="(Berland, 1913)" baseAuthorityName="Berland" baseAuthorityYear="1913" box="[623,989,1476,1502]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="andesiana">
<emphasis id="F7AB4908FFB1FFC56013FA21FCBBFA06" box="[623,799,1476,1502]" italics="true" pageId="15" pageNumber="216">Josa andesiana</emphasis>
(
<bibRefCitation id="A14EE8EBFFB1FFC5614DFA20FC71FA06" author="Berland, L." box="[817,981,1476,1502]" pageId="15" pageNumber="216" pagination="79 - 119" refId="ref14076" refString="Berland, L. (1913) Araignees. In: Mission du Service geographique de l'armee pour la mesure d'un arc du meridien equatorial en Amerique du Sud (1899 - 1906). Tome 10, fascicule 1. Gauthier-Villars, Paris, pp. 79 - 119, pl. 7 - 12. https: // doi. org / 10.5962 / bhl. title. 980" type="book chapter" year="1913">Berland, 1913</bibRefCitation>
)
</taxonomicName>
by the short and tape-shaped embolus (
<figureCitation id="5DE4899FFFB1FFC562E3FA0CFF4AF9DA" box="[159,238,1512,1538]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="15" pageNumber="216">Fig. 13</figureCitation>
EF, compare with
<bibRefCitation id="A14EE8EBFFB1FFC563AEFA0CFD15F9DA" author="Duperre, N." box="[466,689,1512,1538]" pageId="15" pageNumber="216" pagination="118 - 277" refId="ref14337" refString="Duperre, N. (2023) New light on some historical type specimens - in relation to the South American spider (Araneae) fauna-. New Zealand Journal of Zoology, 50 (1), 118 - 277. https: // doi. org / 10.1080 / 03014223.2022.2123835" type="journal article" year="2023">Dupérré, 2023: 139</bibRefCitation>
, fig. 16AC; Poy. et. al. 2023: 6, fig. 3E). It can be distinguished from the latter species by the following combination of characters: laminar projection of the embolus larger (
<figureCitation id="5DE4899FFFB1FFC56716F9E8FF48F992" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="15" pageNumber="216">Figs 13AD</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC56285F9D4FEE8F992" box="[249,332,1584,1610]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="15" pageNumber="216">14AB</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC56325F9D4FE07F992" box="[345,419,1584,1611]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="15" pageNumber="216">15B, I</figureCitation>
), bifid external section of the conductor with both branches interrupted by a deep groove (
<figureCitation id="5DE4899FFFB1FFC562E3F9B0FE8EF9B7" box="[159,298,1620,1647]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="15" pageNumber="216">Fig. 15AB</figureCitation>
), posterior branch of the conductor protruded, with many small spicules and longer than the anterior branch, which is laminar and apically irregular with small reticulations, bifid median apophysis with small scales on the external edge (
<figureCitation id="5DE4899FFFB1FFC56318F978FE62F96F" box="[356,454,1692,1719]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="15" pageNumber="216">Fig. 15F</figureCitation>
), and pronounced retrolateral basal lobe (
<figureCitation id="5DE4899FFFB1FFC561EFF978FC59F96E" box="[915,1021,1692,1718]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="15" pageNumber="216">Figs 13A</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC56675F978FB99F96E" box="[1033,1085,1692,1718]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="15" pageNumber="216">14A</figureCitation>
). The females are distinguished by the spermathecae converging on the median line, touching each other, the obliquely oriented posterior borders of the epigyne and the copulatory ducts anteriorly arched, widening towards the anterior part of vulva (
<figureCitation id="5DE4899FFFB1FFC56768F900FA3CF926" box="[1300,1432,1764,1790]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="15" pageNumber="216">Figs 13EF</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC562EBF8ECFF46F8FA" box="[151,226,1800,1826]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="15" pageNumber="216">14EF</figureCitation>
,
<figureCitation id="5DE4899FFFB1FFC56292F8ECFEA9F8FA" box="[238,269,1800,1826]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="15" pageNumber="216">16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB1FFDC62BBF8C8FEB1FB1D" lastPageId="22" lastPageNumber="223" pageId="15" pageNumber="216" type="description">
<paragraph id="C560951AFFB1FFDC62BBF8C8FA8EFDB9" blockId="15.[151,1437,1116,2043]" lastBlockId="22.[151,1437,151,1257]" lastPageId="22" lastPageNumber="223" pageId="15" pageNumber="216">
<emphasis id="F7AB4908FFB1FFC562BBF8C8FEFDF89E" bold="true" box="[199,345,1836,1862]" pageId="15" pageNumber="216">Description.</emphasis>
<emphasis id="F7AB4908FFB1FFC56319F8C8FE3BF89E" box="[357,415,1836,1862]" italics="true" pageId="15" pageNumber="216">Male</emphasis>
(
<typeStatus id="1A642BB8FFB1FFC563CFF8C8FDBDF89E" box="[435,537,1836,1862]" pageId="15" pageNumber="216" type="holotype">holotype</typeStatus>
, MUSENUV-Ar-2678; vchLAM-249) (
<figureCitation id="5DE4899FFFB1FFC5619AF8C8FB9AF89E" box="[998,1086,1836,1862]" captionStart="FIGURE 11" captionStartId="16.[151,250,1882,1906]" captionTargetBox="[159,1427,189,1858]" captionTargetId="figure-22@16.[151,1436,181,1858]" captionTargetPageId="16" captionText="FIGURE 11. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="15" pageNumber="216">Fig. 11</figureCitation>
): Total length 5.75. Carapace length 2.95, width 2.13, height 1.10. Clypeus length 0.13. Sternum length 1.45, width 0.98. Legs. Leg I: ti length 1.79/width 0.36, mt 1.39. II: ti 1.70/0.35, mt 1.46. III: ti 1.72/0.38, mt 1.75. IV: ti 2.74/0.44, mt 2.59. Spines: legs III, femur d 1-1-1, p 2ap, r 0-d1-d1; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0; metatarsus v 2-0-0, p d1-0. IIIIV, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-2-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 2.81, width 1.56, spiracleepigastrium 1.01. Color: carapace brown, marginally darker, with two wide black degraded bands, converging and narrow anteriorly. Ocular area dark brown with eyes bordered on a black patch. Lateral regions of carapace upholstered with small fine white setae. Clypeus brown, low with some white setae. Chelicerae brown with fine setae and dark patches on frontal edge. Endites light brown, with faint medial dark patch. Labium brown reddish, anteriorly lighter. Sternum brown with faint black patches. Legs uniformly brown, covered by white setae, mainly between coxae. Abdomen dorsum with a pattern of black spots on a brown background, surrounded by a thin band, extended to medial region; dorsum with an anterior medial dark patch with posterior extensions, followed by irregular spots. Abdomen covered with fine white setae; anterior region with a dense bunch of setae, marginal ring covered by black setae. Venter uniformly brown with light and dark dots, dark patches surrounding the spinnerets base. Palp: tegulum with a basal small lobe, with granulose texture (
<figureCitation id="5DE4899FFFA8FFDC6316FE8BFE73FE51" box="[362,471,367,393]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13A</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC6398FE8BFDB3FE51" box="[484,535,367,393]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14A</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC6058FE8AFDDFFE51" box="[548,635,366,393]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">15A, G</figureCitation>
). Median apophysis long, laminar, ending in a very short hook-shaped tip, with many denticles (
<figureCitation id="5DE4899FFFA8FFDC63CEFE77FDB6FE75" box="[434,530,402,429]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">Fig. 15F</figureCitation>
, see square). Conductor small, rounded, ventral section apically bifid, with many narrow furrows, anterior tip apically laminar with several very small denticles, posterior tip with small granules, posterior tip slightly longer than anterior one; base of the conductor with a sharp, spine-shaped projection (
<figureCitation id="5DE4899FFFA8FFDC674EFE3FFF08FDC1" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">Fig. 15A, B</figureCitation>
). Paramedian apophysis cup-shaped, concave (
<figureCitation id="5DE4899FFFA8FFDC60B4FE1BFCF3FDC1" box="[712,855,511,537]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13AB</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC6118FE1BFC1CFDC1" box="[868,952,511,537]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14AB</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC61BAFE1AFBFBFDC1" box="[966,1119,510,537]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">15AB, D, F</figureCitation>
). Embolus tape-shaped, not describing a loop, very short, with large laminar projection (
<figureCitation id="5DE4899FFFA8FFDC613EFDC7FC47FDE5" box="[834,995,546,573]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">Fig. 15B, G, I</figureCitation>
), apically twisted; basal process small, flat, sclerotized, with poorly pronounced external keel and medial depression (
<figureCitation id="5DE4899FFFA8FFDC667AFDA3FB37FDB9" box="[1030,1171,583,609]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13AD</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC66E3FDA3FB57FDB9" box="[1183,1267,583,609]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14AC</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC6682FDA2FAB9FDB9" box="[1278,1309,582,609]" captionStart="FIGURE 15" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@20.[151,1435,181,1756]" captionTargetPageId="20" captionText="FIGURE 15. Josa nadineae sp. nov., male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum." pageId="22" pageNumber="223">15</figureCitation>
).
</paragraph>
<caption id="91A0C592FFAEFFDA62EBF8BEFD3CF84E" pageId="16" pageNumber="217" startId="16.[151,250,1882,1906]" targetBox="[159,1427,189,1858]" targetPageId="16" targetType="figure">
<paragraph id="C560951AFFAEFFDA62EBF8BEFD3CF84E" blockId="16.[151,1436,1882,1942]" pageId="16" pageNumber="217">
<emphasis id="F7AB4908FFAEFFDA62EBF8BEFE86F8AA" bold="true" box="[151,290,1882,1906]" pageId="16" pageNumber="217">FIGURE 11.</emphasis>
<taxonomicName id="02DFEE99FFAEFFDA6357F8BFFE65F8AA" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,449,1882,1906]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFAEFFDA6357F8BFFE65F8AA" box="[299,449,1882,1906]" italics="true" pageId="16" pageNumber="217">Josa nadineae</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFAEFFDA63B6F8BEFDBAF8AA" bold="true" box="[458,542,1882,1906]" pageId="16" pageNumber="217">
<taxonomicNameLabel id="EC98F473FFAEFFDA63B6F8BEFDBAF8AA" box="[458,542,1882,1906]" pageId="16" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2678; vchLAM-249): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFAFFFDB62EBF8BEFD3CF84E" pageId="17" pageNumber="218" startId="17.[151,250,1882,1906]" targetBox="[159,1423,186,1853]" targetPageId="17" targetType="figure">
<paragraph id="C560951AFFAFFFDB62EBF8BEFD3CF84E" blockId="17.[151,1436,1882,1942]" pageId="17" pageNumber="218">
<emphasis id="F7AB4908FFAFFFDB62EBF8BEFE86F8AA" bold="true" box="[151,290,1882,1906]" pageId="17" pageNumber="218">FIGURE 12.</emphasis>
<taxonomicName id="02DFEE99FFAFFFDB6356F8BFFE1AF8AA" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[298,446,1882,1906]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFAFFFDB6356F8BFFE1AF8AA" box="[298,446,1882,1906]" italics="true" pageId="17" pageNumber="218">Josa nadineae</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFAFFFDB63BAF8BEFDBCF8AA" bold="true" box="[454,536,1882,1906]" pageId="17" pageNumber="218">
<taxonomicNameLabel id="EC98F473FFAFFFDB63BAF8BEFDBCF8AA" box="[454,536,1882,1906]" pageId="17" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (MUSENUV-Ar-2679; vchLAM-251): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFACFFD862EBF8C8FBEEF80C" pageId="18" pageNumber="219" startId="18.[151,250,1836,1860]" targetBox="[151,1412,182,1812]" targetPageId="18" targetType="figure">
<paragraph id="C560951AFFACFFD862EBF8C8FBEEF80C" blockId="18.[151,1436,1836,2005]" pageId="18" pageNumber="219">
<emphasis id="F7AB4908FFACFFD862EBF8C8FE87F89C" bold="true" box="[151,291,1836,1860]" pageId="18" pageNumber="219">FIGURE 13.</emphasis>
<taxonomicName id="02DFEE99FFACFFD86351F8C9FE67F89C" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[301,451,1836,1860]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFACFFD86351F8C9FE67F89C" box="[301,451,1836,1860]" italics="true" pageId="18" pageNumber="219">Josa nadineae</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFACFFD863B1F8C9FD85F89D" bold="true" box="[461,545,1837,1861]" pageId="18" pageNumber="219">
<taxonomicNameLabel id="EC98F473FFACFFD863B1F8C9FD85F89D" box="[461,545,1837,1861]" pageId="18" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view.
<emphasis id="F7AB4908FFACFFD8609CF890FC25F854" bold="true" box="[736,897,1908,1932]" pageId="18" pageNumber="219">Abbreviations:</emphasis>
C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe.
</paragraph>
</caption>
<caption id="91A0C592FFADFFD962EBF8AEFBF9F82A" pageId="19" pageNumber="220" startId="19.[151,250,1866,1890]" targetBox="[155,1432,185,1840]" targetPageId="19" targetType="figure">
<paragraph id="C560951AFFADFFD962EBF8AEFBF9F82A" blockId="19.[151,1436,1866,2035]" pageId="19" pageNumber="220">
<emphasis id="F7AB4908FFADFFD962EBF8AEFE83F8BA" bold="true" box="[151,295,1866,1890]" pageId="19" pageNumber="220">FIGURE 14.</emphasis>
<taxonomicName id="02DFEE99FFADFFD96348F8AFFE68F8BA" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[308,460,1866,1890]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFADFFD96348F8AFFE68F8BA" box="[308,460,1866,1890]" italics="true" pageId="19" pageNumber="220">Josa nadineae</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFADFFD963A5F8AFFD94F8BB" bold="true" box="[473,560,1867,1891]" pageId="19" pageNumber="220">
<taxonomicNameLabel id="EC98F473FFADFFD963A5F8AFFD94F8BB" box="[473,560,1867,1891]" pageId="19" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology.
<emphasis id="F7AB4908FFADFFD96308F876FDB1F872" bold="true" box="[372,533,1938,1962]" pageId="19" pageNumber="220">Abbreviations:</emphasis>
AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct.
</paragraph>
</caption>
<caption id="91A0C592FFAAFFDE62EBF917FD70F818" pageId="20" pageNumber="221" startId="20.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="20" targetType="figure">
<paragraph id="C560951AFFAAFFDE62EBF917FD70F818" blockId="20.[151,1437,1779,1984]" pageId="20" pageNumber="221">
<emphasis id="F7AB4908FFAAFFDE62EBF917FE84F8D4" bold="true" box="[151,288,1779,1804]" pageId="20" pageNumber="221">FIGURE 15.</emphasis>
<taxonomicName id="02DFEE99FFAAFFDE635BF910FE1EF8D4" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[295,442,1780,1804]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFAAFFDE635BF910FE1EF8D4" box="[295,442,1780,1804]" italics="true" pageId="20" pageNumber="221">Josa nadineae</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFAAFFDE63BDF910FDB6F8D4" bold="true" box="[449,530,1780,1804]" pageId="20" pageNumber="221">
<taxonomicNameLabel id="EC98F473FFAAFFDE63BDF910FDB6F8D4" box="[449,530,1780,1804]" pageId="20" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (MACN-Ar 46692; vchLAM-250) copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D. Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of median apophysis, retrolateral view (inset to small scales on external border); G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details.
<emphasis id="F7AB4908FFAAFFDE6638F8BBFB41F8AF" bold="true" box="[1092,1253,1887,1911]" pageId="20" pageNumber="221">Abbreviations:</emphasis>
C, conductor; E, embolus; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe; St, subtegulum.
</paragraph>
</caption>
<caption id="91A0C592FFABFFDF62EBF927FB0AF8FC" pageId="21" pageNumber="222" startId="21.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="21" targetType="figure">
<paragraph id="C560951AFFABFFDF62EBF927FB0AF8FC" blockId="21.[151,1436,1731,1828]" pageId="21" pageNumber="222">
<emphasis id="F7AB4908FFABFFDF62EBF927FE87F904" bold="true" box="[151,291,1731,1756]" pageId="21" pageNumber="222">FIGURE 16.</emphasis>
<taxonomicName id="02DFEE99FFABFFDF6351F920FE66F904" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[301,450,1732,1756]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFABFFDF6351F920FE66F904" box="[301,450,1732,1756]" italics="true" pageId="21" pageNumber="222">Josa nadineae</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFABFFDF63B7F920FDBBF904" bold="true" box="[459,543,1732,1756]" pageId="21" pageNumber="222">
<taxonomicNameLabel id="EC98F473FFABFFDF63B7F920FDBBF904" box="[459,543,1732,1756]" pageId="21" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view.
<emphasis id="F7AB4908FFABFFDF6687F903FA38F927" bold="true" box="[1275,1436,1767,1791]" pageId="21" pageNumber="222">Abbreviations:</emphasis>
dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca.
</paragraph>
</caption>
<paragraph id="C560951AFFA8FFDC62BBFD8FFAF9FBED" blockId="22.[151,1437,151,1257]" pageId="22" pageNumber="223">
<emphasis id="F7AB4908FFA8FFDC62BBFD8FFEBFFD5D" box="[199,283,619,645]" italics="true" pageId="22" pageNumber="223">Female</emphasis>
(
<typeStatus id="1A642BB8FFA8FFDC6351FD8FFE36FD5D" box="[301,402,619,645]" pageId="22" pageNumber="223" type="paratype">paratype</typeStatus>
, MUSENUV-Ar-2679; vchLAM-251). (
<figureCitation id="5DE4899FFFA8FFDC6122FD8FFC12FD5D" box="[862,950,619,645]" captionStart="FIGURE 12" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[159,1423,186,1853]" captionTargetId="figure-17@17.[151,1436,181,1858]" captionTargetPageId="17" captionText="FIGURE 12. Josa nadineae sp. nov., female (MUSENUV-Ar-2679; vchLAM-251): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="22" pageNumber="223">Fig. 12</figureCitation>
): Total length 7.02. Carapace length 3.19, width 2.23, height 1.17. Clypeus length 0.13. Sternum length 1.59, width 1.06. Legs. Leg I: ti length 1.61/width 0.39, mt 1.21. II: ti 1.63/0.34, mt 1.26. III: ti 1.36/0.37, mt 1.56. IV: ti 1.95/0.38, mt 1.50. Spines: legs III, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. IIIIV, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 3.63, width 2.69, spiracleepigastrium 1.53. Color pattern as in male, but more pigmented, abdominal spots more defined. Sternum and labium more pigmented. Legs with defined longitudinal dark bands on dorsal edge, and many dots on ventral one. Epigyne: anterior plate sclerotized with deep and rounded anterior excavations, laterally elongated, median depression small, quadrangular, posteriorly positioned (
<figureCitation id="5DE4899FFFA8FFDC6126FC6FFC66FC7D" box="[858,962,907,933]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13E</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC61B0FC6FFBA5FC7D" box="[972,1025,907,933]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14D</figureCitation>
). Median field transverse, large, oval in posterior view with medial depression (
<figureCitation id="5DE4899FFFA8FFDC60FDFC4BFCACFC11" box="[641,776,943,969]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="22" pageNumber="223">Fig. 16EF</figureCitation>
). Lateral lobes with sclerotized extensions wide, apically rounded (
<figureCitation id="5DE4899FFFA8FFDC637FFC37FE2CFC35" box="[259,392,979,1005]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="22" pageNumber="223">Fig. 16EF</figureCitation>
). Copulatory ducts long, with a very wide posterior coil, laterally expanded in anterior side, joining with the accessory bulb ducts at medial level. Spermathecae small, rounded, in median position, oriented towards median septum (
<figureCitation id="5DE4899FFFA8FFDC63CCFBFFFDB2FBED" box="[432,534,1051,1077]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">Fig. 14E</figureCitation>
). Ducts of accessory bulbs long, thin, convergent (
<figureCitation id="5DE4899FFFA8FFDC6637FBFFFB6BFBED" box="[1099,1231,1051,1077]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="22" pageNumber="223">Figs 13EF</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC66A7FBFFFA82FBED" box="[1243,1318,1051,1077]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="22" pageNumber="223">14EF</figureCitation>
,
<figureCitation id="5DE4899FFFA8FFDC674DFBFFFAF4FBED" box="[1329,1360,1051,1077]" captionStart="FIGURE 16" captionStartId="21.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@21.[151,1435,181,1708]" captionTargetPageId="21" captionText="FIGURE 16. Josa nadineae sp. nov., female (MACN-Ar 46693; vchLAM-252) genitalia SEM images: A Dorsal view; B Anterior-dorsal view; C Fertilization duct; D Pore of bennets gland; E Posterior-dorsal view; F Posterior view. Abbreviations: dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="22" pageNumber="223">16</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFA8FFDC62BBFBDAFEB1FB1D" blockId="22.[151,1437,151,1257]" pageId="22" pageNumber="223">
<emphasis id="F7AB4908FFA8FFDC62BBFBDAFEE5FB80" bold="true" box="[199,321,1086,1112]" pageId="22" pageNumber="223">Variation:</emphasis>
Males (n=3): total length: 5.405.75; carapace length: 2.472.97, width: 1.842.13; abdomen length: 2.662.81, width: 1.561.69. Females (n=4): total length: 7.027.49; carapace length: 3.193.79, width: 2.232.67; abdomen length: 3.633.72, width: 1.252.69. Some specimens present body and abdominal spot patterns more pigmented.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFA8FFDC62BBFB2AFBD7FB31" box="[199,1139,1230,1257]" pageId="22" pageNumber="223" type="distribution">
<paragraph id="C560951AFFA8FFDC62BBFB2AFBD7FB31" blockId="22.[151,1437,151,1257]" box="[199,1139,1230,1257]" pageId="22" pageNumber="223">
<emphasis id="F7AB4908FFA8FFDC62BBFB2AFEC5FB30" bold="true" box="[199,353,1230,1256]" pageId="22" pageNumber="223">Distribution.</emphasis>
Only known from Sierra Nevada de Santa Marta,
<collectingCountry id="BDC8D58AFFA8FFDC61E9FB2BFBA7FB31" box="[917,1027,1231,1257]" name="Colombia" pageId="22" pageNumber="223">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFFA8FFDC666FFB2BFBC2FB31" box="[1043,1126,1231,1257]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="22" pageNumber="223">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,223 @@
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<mods:title id="072FF30F099D9ABC7E97D073BF3FC37F">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
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<mods:namePart id="EC6B14BBA522B2314DA2AC72AF623591">Martínez, Leonel</mods:namePart>
<mods:affiliation id="B4918B6C5EB7D22AE18B30F8B741FEFA">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina &amp; Grupo de Investigación Biodiversidad del Caribe, Departamento de Biología, Universidad del Atlántico, Barranquilla, Colombia &amp; Grupo de Estudios en Sistemática y Conservación, Universidad del Atlántico, Barranquilla, Colombia &amp; Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales, Universidad de Buenos Aires, Ciudad Autónoma de Buenos Aires, Argentina. Argentina</mods:affiliation>
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<mods:namePart id="02C677A31006ECEE4AF02C3953B5A037">Kochalka, John A.</mods:namePart>
<mods:affiliation id="626AA6B3A06863DDB8A9004BA3AD9FD0">Museo Nacional de Historia Natural del Paraguay, Sucursal 1 Campus U. N. A., C. C. 19.004, 111421 CDP, Central XI, San Lorenzo, Paraguay</mods:affiliation>
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<mods:affiliation id="C54F2E77027EE90D11A756E01E7C0D85">Departmento de Biología, Universidad del Valle, Cali, Valle del Cauca, Colombia</mods:affiliation>
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<mods:namePart id="9970DDCC2FF65271C8C0C87775E204C9">Ramírez, Martin</mods:namePart>
<mods:affiliation id="2F1F8D6D8FE400AFA23B69ECC168CE6D">División Aracnología, Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” CONICET, Avenida Ángel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
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<subSubSection id="8DC5C691FFB2FFC662EBFA2FFE39FA3D" box="[151,413,1483,1509]" pageId="12" pageNumber="213" type="nomenclature">
<paragraph id="C560951AFFB2FFC662EBFA2FFE39FA3D" blockId="12.[151,413,1483,1545]" box="[151,413,1483,1509]" pageId="12" pageNumber="213">
<heading id="9E282276FFB2FFC662EBFA2FFE39FA3D" bold="true" box="[151,413,1483,1509]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<emphasis id="F7AB4908FFB2FFC662EBFA2FFE39FA3D" bold="true" box="[151,413,1483,1509]" pageId="12" pageNumber="213">
<taxonomicName id="02DFEE99FFB2FFC662EBFA2FFE9CFA3D" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[151,312,1483,1509]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="samaria" status="sp. nov.">
<emphasis id="F7AB4908FFB2FFC662EBFA2FFE9CFA3D" bold="true" box="[151,312,1483,1509]" italics="true" pageId="12" pageNumber="213">Josa samaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EC98F473FFB2FFC66343FA2FFE39FA3D" box="[319,413,1483,1509]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB2FFC662EBFA0BFE8EF9D1" box="[151,298,1519,1545]" pageId="12" pageNumber="213" type="description">
<paragraph id="C560951AFFB2FFC662EBFA0BFE8EF9D1" blockId="12.[151,413,1483,1545]" box="[151,298,1519,1545]" pageId="12" pageNumber="213">
<figureCitation id="5DE4899FFFB2FFC662EBFA0BFE8EF9D1" box="[151,298,1519,1545]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="13.[151,250,1867,1891]" captionStartId-1="14.[151,250,1785,1809]" captionTargetBox-0="[167,1426,189,1841]" captionTargetBox-1="[151,1417,191,1757]" captionTargetId-0="figure-17@13.[151,1436,181,1844]" captionTargetId-1="figure-22@14.[151,1436,181,1761]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 9. Josa samaria sp. nov., female (MNHP; vchLAM-244): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="12" pageNumber="213">Figures 910</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB2FFC662EBF9D7FABFF993" box="[151,1307,1587,1611]" pageId="12" pageNumber="213" type="reference_group">
<paragraph id="C560951AFFB2FFC662EBF9D7FABFF993" blockId="12.[151,1307,1587,1611]" box="[151,1307,1587,1611]" pageId="12" pageNumber="213">
<treatmentCitationGroup id="E5CFB234FFB2FFC662EBF9D7FABFF993" box="[151,1307,1587,1611]" pageId="12" pageNumber="213">
<treatmentCitation id="447EB30BFFB2FFC662EBF9D7FD8EF993" author="Simon, E." box="[151,554,1587,1611]" page="11" pageId="12" pageNumber="213" year="1897">
<taxonomicName id="02DFEE99FFB2FFC662EBF9D7FD8EF993" authority="Simon, 1897: 11" authorityName="Simon" authorityPageNumber="11" authorityYear="1897" box="[151,554,1587,1611]" class="Arachnida" family="Anyphaenidae" genus="Tomopisthes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFB2FFC662EBF9D7FED8F993" box="[151,380,1587,1611]" italics="true" pageId="12" pageNumber="213">Tomopisthes chazaliae</emphasis>
<materialsCitation id="75B79F47FFB2FFC663FFF9D7FD8EF993" box="[387,554,1587,1611]" collectionCode="MNHP" pageId="12" pageNumber="213" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<bibRefCitation id="A14EE8EBFFB2FFC663FFF9D7FD8EF993" author="Simon, E." box="[387,554,1587,1611]" pageId="12" pageNumber="213" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897: 11</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(only
<specimenCount id="D3D95E93FFB2FFC66010F9D0FD78F993" box="[620,732,1587,1611]" count="1" pageId="12" pageNumber="213" type="female">one female</specimenCount>
<typeStatus id="1A642BB8FFB2FFC6609EF9D7FCCEF993" box="[738,874,1587,1611]" pageId="12" pageNumber="213" type="paralectotype">paralectotype</typeStatus>
MNHP; vchLAM-244, misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB2FFC662EBF99DFCE6F96F" pageId="12" pageNumber="213" type="materials_examined">
<paragraph id="C560951AFFB2FFC662EBF99DFCE6F96F" blockId="12.[151,1436,1657,1935]" pageId="12" pageNumber="213">
<emphasis id="F7AB4908FFB2FFC662EBF99DFEE3F94B" bold="true" box="[151,327,1657,1683]" pageId="12" pageNumber="213">Type Material.</emphasis>
<materialsCitation id="75B79F47FFB2FFC66330F99DFCE6F96F" collectionCode="MNHP" collectorName="De Dalmas" country="Colombia" location="Sierra Nevada de Santa Marta" pageId="12" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena" typeStatus="holotype">
<typeStatus id="1A642BB8FFB2FFC66330F99DFE1DF94B" box="[332,441,1657,1683]" pageId="12" pageNumber="213" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="BDC8D58AFFB2FFC663BDF99DFDFEF94B" box="[449,602,1657,1683]" name="Colombia" pageId="12" pageNumber="213">COLOMBIA</collectingCountry>
.
<collectingRegion id="071B5BF8FFB2FFC6601FF99DFD46F94B" box="[611,738,1657,1683]" country="Colombia" name="Magdalena" pageId="12" pageNumber="213">Magdalena</collectingRegion>
, from
<location id="C000C3C1FFB2FFC66154F99DFBD0F94C" LSID="urn:lsid:plazi:treatment:4D76240CFFB2FFC562EBFA2FFBFAFC7D:C000C3C1FFB2FFC66154F99DFBD0F94C" box="[808,1140,1657,1684]" country="Colombia" name="Sierra Nevada de Santa Marta" pageId="12" pageNumber="213" stateProvince="Magdalena">Sierra Nevada de Santa Marta</location>
,
<collectorName id="682AF0CCFFB2FFC66601F99EFB5FF94B" box="[1149,1275,1657,1684]" pageId="12" pageNumber="213">De Dalmas</collectorName>
,
<specimenCount id="D3D95E93FFB2FFC66779F99DFA8AF94C" box="[1285,1326,1657,1684]" count="1" pageId="12" pageNumber="213" type="female">1 ♀</specimenCount>
, (
<collectionCode id="A3CE0DDFFFB2FFC6673EF99DFA3CF94B" box="[1346,1432,1657,1683]" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="12" pageNumber="213">MNHP</collectionCode>
; vchLAM-244;
<typeStatus id="1A642BB8FFB2FFC6633CF979FE73F96F" box="[320,471,1693,1719]" pageId="12" pageNumber="213" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="02DFEE99FFB2FFC66380F97AFC91F96F" authority="Simon 1897" authorityName="Simon" authorityYear="1897" box="[508,821,1693,1719]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFB2FFC66380F97AFD05F96F" box="[508,673,1693,1719]" italics="true" pageId="12" pageNumber="213">Josa chazaliae</emphasis>
<bibRefCitation id="A14EE8EBFFB2FFC660D5F979FC91F96F" author="Simon, E." box="[681,821,1693,1719]" pageId="12" pageNumber="213" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon 1897</bibRefCitation>
</taxonomicName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB2FFC662BBF925FEB0F927" pageId="12" pageNumber="213" type="etymology">
<paragraph id="C560951AFFB2FFC662BBF925FEB0F927" blockId="12.[151,1436,1657,1935]" pageId="12" pageNumber="213">
<emphasis id="F7AB4908FFB2FFC662BBF925FEEAF903" bold="true" box="[199,334,1729,1755]" pageId="12" pageNumber="213">Etymology.</emphasis>
The specific epithet refers to the demonym for the Santa Marta people “Samario/a”. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB2FFC662BBF8EDFAC4F857" pageId="12" pageNumber="213" type="diagnosis">
<paragraph id="C560951AFFB2FFC662BBF8EDFAC4F857" blockId="12.[151,1436,1657,1935]" pageId="12" pageNumber="213">
<emphasis id="F7AB4908FFB2FFC662BBF8EDFEE6F8FB" bold="true" box="[199,322,1801,1827]" pageId="12" pageNumber="213">Diagnosis.</emphasis>
The female is similar to that of
<taxonomicName id="02DFEE99FFB2FFC660CDF8EEFCF0F8FB" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[689,852,1801,1827]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFB2FFC660CDF8EEFCF0F8FB" box="[689,852,1801,1827]" italics="true" pageId="12" pageNumber="213">Josa nadineae</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFB2FFC66120F8EDFC12F8FB" bold="true" box="[860,950,1801,1827]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="EC98F473FFB2FFC66120F8EDFC12F8FB" box="[860,950,1801,1827]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the posterior quadrangular notch of the epigyne (
<figureCitation id="5DE4899FFFB2FFC6637EF8C9FECDF89F" box="[258,361,1837,1863]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Fig. 13E</figureCitation>
, compare with
<figureCitation id="5DE4899FFFB2FFC66060F8C9FD22F89F" box="[540,646,1837,1863]" captionStart="FIGURE 13" captionStartId="18.[151,250,1836,1860]" captionTargetBox="[151,1412,182,1812]" captionTargetId="figure-22@18.[151,1436,181,1812]" captionTargetPageId="18" captionText="FIGURE 13. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2679; vchLAM-251) genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; LPE, laminar projection of embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Figs 13E</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC660E8F8C9FD62F89F" box="[660,710,1837,1863]" captionStart="FIGURE 14" captionStartId="19.[151,250,1866,1890]" captionTargetBox="[155,1432,185,1840]" captionTargetId="figure-17@19.[151,1435,181,1842]" captionTargetPageId="19" captionText="FIGURE 14. Josa nadineae sp. nov., male (MUSENUV-Ar-2678; vchLAM-249) copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2679; vchLAM-251): D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">14D</figureCitation>
) but can be distinguished by the posterior lateral margins on a straight transversal line (vs. oblique in
<taxonomicName id="02DFEE99FFB2FFC66028F8B6FD53F8B3" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[596,759,1873,1899]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="nadineae" status="sp. nov.">
<emphasis id="F7AB4908FFB2FFC66028F8B6FD53F8B3" box="[596,759,1873,1899]" italics="true" pageId="12" pageNumber="213">Josa nadineae</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFB2FFC66083F8B5FCC7F8B3" bold="true" box="[767,867,1873,1899]" pageId="12" pageNumber="213">
<taxonomicNameLabel id="EC98F473FFB2FFC66083F8B5FCFFF8B3" box="[767,859,1873,1899]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
)
</emphasis>
, and by the copulatory ducts describing two wide posterior coils, and the small spermathecae, anteromedially placed with long fertilization ducts (
<figureCitation id="5DE4899FFFB2FFC666B0F891FAF1F857" box="[1228,1365,1909,1935]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="12" pageNumber="213">Fig. 10AE</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="91A0C592FFB3FFC762EBF8AFFE55F850" pageId="13" pageNumber="214" startId="13.[151,250,1867,1891]" targetBox="[167,1426,189,1841]" targetPageId="13" targetType="figure">
<paragraph id="C560951AFFB3FFC762EBF8AFFE55F850" blockId="13.[151,1436,1867,1928]" pageId="13" pageNumber="214">
<emphasis id="F7AB4908FFB3FFC762EBF8AFFEB0F8BC" bold="true" box="[151,276,1867,1892]" pageId="13" pageNumber="214">FIGURE 9.</emphasis>
<taxonomicName id="02DFEE99FFB3FFC76361F8A8FE02F8BB" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[285,422,1868,1891]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="samaria" status="sp. nov.">
<emphasis id="F7AB4908FFB3FFC76361F8A8FE02F8BB" box="[285,422,1868,1891]" italics="true" pageId="13" pageNumber="214">Josa samaria</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFB3FFC763D2F8A8FDA4F8BC" bold="true" box="[430,512,1868,1892]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="EC98F473FFB3FFC763D2F8A8FDA4F8BC" box="[430,512,1868,1892]" pageId="13" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (MNHP; vchLAM-244): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFB0FFC462EBF91DFDE7F8A5" pageId="14" pageNumber="215" startId="14.[151,250,1785,1809]" targetBox="[151,1417,191,1757]" targetPageId="14" targetType="figure">
<paragraph id="C560951AFFB0FFC462EBF91DFDE7F8A5" blockId="14.[151,1436,1785,1918]" pageId="14" pageNumber="215">
<emphasis id="F7AB4908FFB0FFC462EBF91DFE86F8C9" bold="true" box="[151,290,1785,1809]" pageId="14" pageNumber="215">FIGURE 10.</emphasis>
<taxonomicName id="02DFEE99FFB0FFC46357F91EFE11F8C9" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[299,437,1786,1809]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="samaria" status="sp. nov.">
<emphasis id="F7AB4908FFB0FFC46357F91EFE11F8C9" box="[299,437,1786,1809]" italics="true" pageId="14" pageNumber="215">Josa samaria</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFB0FFC463C2F91EFDB5F8CA" bold="true" box="[446,529,1786,1810]" pageId="14" pageNumber="215">
<taxonomicNameLabel id="EC98F473FFB0FFC463C2F91EFDB5F8CA" box="[446,529,1786,1810]" pageId="14" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology.
<emphasis id="F7AB4908FFB0FFC466B5F8F9FACEF8ED" bold="true" box="[1225,1386,1821,1845]" pageId="14" pageNumber="215">Abbreviations:</emphasis>
AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca.
</paragraph>
</caption>
<subSubSection id="8DC5C691FFB1FFC562BBFF72FE21FC59" pageId="15" pageNumber="216" type="description">
<paragraph id="C560951AFFB1FFC562BBFF72FD5DFC85" blockId="15.[151,1437,150,933]" pageId="15" pageNumber="216">
<emphasis id="F7AB4908FFB1FFC562BBFF72FEFDFF68" bold="true" box="[199,345,150,176]" pageId="15" pageNumber="216">Description.</emphasis>
<emphasis id="F7AB4908FFB1FFC56323FF73FE17FF69" box="[351,435,151,177]" italics="true" pageId="15" pageNumber="216">Female</emphasis>
(
<typeStatus id="1A642BB8FFB1FFC563BDFF73FD83FF69" box="[449,551,151,177]" pageId="15" pageNumber="216" type="holotype">holotype</typeStatus>
, MNHP; vchLAM-244) (
<figureCitation id="5DE4899FFFB1FFC5613EFF73FC22FF69" box="[834,902,151,177]" captionStart="FIGURE 9" captionStartId="13.[151,250,1867,1891]" captionTargetBox="[167,1426,189,1841]" captionTargetId="figure-17@13.[151,1436,181,1844]" captionTargetPageId="13" captionText="FIGURE 9. Josa samaria sp. nov., female (MNHP; vchLAM-244): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="15" pageNumber="216">Fig. 9</figureCitation>
): Total length 7.33. Carapace length 3.41, width 2.36, height 1.35. Clypeus length 0.19. Sternum length 1.79, width 1.31. Legs. Leg I: ti length 1.84/width 0.42, metatarsus (mt) 1.45. II: ti 1.74/0.40, mt 1.39. III: ti 1.59/0.38, mt 1.55. IV: ti 2.17/0.44, mt 2.41. Spines: legs III, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2r-1-2, p 1-d1-1-0 r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. IV=III but metatarsus v 2-2-2. Abdomen: length 4.03, width 2.33, spiracleepigastrium 0.80. Color: carapace brown, with two wide, longitudinal darker band on sides, narrow in cephalic region. Ocular area on black patch with eyes bordered by black rings. Carapace with lateral region covered by fringe of white setae. Clypeus brown, low with abundant white setae. Chelicerae brown with black patches and fine setae on frontal edge. Endites, labium, and sternum dark brown, labium and endites light in apical region. Sternum with a very thin medial pale stripe. Legs coxae-tarsi dark brown, femora and patellae with longitudinal dark bands. All articles covered by white setae, abundant between coxae. Abdomen with dark brown pattern of spots on a gray background, dorsally with white medial band followed by 4-5 faint light chevrons; lateral regions covered by abundant small dots. Abdomen laterally surrounded by abundant white setae, with a thick bunch of setae on anterior region. Venter uniformly covered by dark and light patches. Epigyne: anterior plate sclerotized with deep and rounded anterior excavations, median notch quadrangular, posteriorly positioned (
<figureCitation id="5DE4899FFFB1FFC563E6FD57FD80FD15" box="[410,548,691,717]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Fig. 10A, D</figureCitation>
). Median field transverse, large, oval in posterior view (
<figureCitation id="5DE4899FFFB1FFC566EEFD57FB5DFD15" box="[1170,1273,691,717]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Fig. 10C</figureCitation>
). Lateral lobes with sclerotized extensions wide, forming, posterior extensions rounded at base (
<figureCitation id="5DE4899FFFB1FFC56662FD33FB6EFD29" box="[1054,1226,727,753]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Fig. 10A, CD</figureCitation>
). Copulatory ducts long, forming two basal coils, anteriorly medially expanded, joining with the accessory bulb ducts at medial level. Spermathecae small, rounded, medially positioned in the epigyne, oriented towards longitudinal plane. Ducts of accessory bulbs short, wide, convergent (
<figureCitation id="5DE4899FFFB1FFC56018FCA7FD49FC85" box="[612,749,835,861]" captionStart="FIGURE 10" captionStartId="14.[151,250,1785,1809]" captionTargetBox="[151,1417,191,1757]" captionTargetId="figure-22@14.[151,1436,181,1761]" captionTargetPageId="14" captionText="FIGURE 10. Josa samaria sp. nov., female (MNHP; vchLAM-244) genitalia: A Ventral view (arrow indicates the anterior excavations); B Dorsal view; C Posterior view; D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; CO, copulatory opening; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LB, lateral border; MF, median field; S, spermatheca." pageId="15" pageNumber="216">Fig. 10AE</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFB1FFC562BBFC83FE21FC59" blockId="15.[151,1437,150,933]" box="[199,389,871,897]" pageId="15" pageNumber="216">
<emphasis id="F7AB4908FFB1FFC562BBFC83FEACFC59" box="[199,264,871,897]" italics="true" pageId="15" pageNumber="216">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB1FFC562BBFC6EFBFAFC7D" box="[199,1118,906,933]" pageId="15" pageNumber="216" type="distribution">
<paragraph id="C560951AFFB1FFC562BBFC6EFBFAFC7D" blockId="15.[151,1437,150,933]" box="[199,1118,906,933]" pageId="15" pageNumber="216">
<emphasis id="F7AB4908FFB1FFC562BBFC6EFEC5FC7C" bold="true" box="[199,353,906,932]" pageId="15" pageNumber="216">Distribution.</emphasis>
Only known for Sierra Nevada de Santa Marta,
<collectingCountry id="BDC8D58AFFB1FFC56103FC6FFC49FC7D" box="[895,1005,907,933]" name="Colombia" pageId="15" pageNumber="216">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFFB1FFC56181FC6FFBF4FC7D" box="[1021,1104,907,933]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="15" pageNumber="216">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,494 @@
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<mods:title id="BE06A207442A82F896AC89B701B797EA">Revealing the identity of Josa chazaliae (Simon, 1897) (Araneae: Anyphaenidae): new species and the highest altitude record for a spider in South America</mods:title>
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<heading id="9E282276FFBDFFC962EBFCC3FE58FC99" bold="true" box="[151,508,807,834]" fontSize="11" level="1" pageId="3" pageNumber="204" reason="1">
<taxonomicName id="02DFEE99FFBDFFC962EBFCC3FE58FC99" ID-CoL="6NG93" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[151,508,807,834]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFBDFFC962EBFCC3FE58FC99" bold="true" box="[151,508,807,834]" pageId="3" pageNumber="204">
<emphasis id="F7AB4908FFBDFFC962EBFCC3FEECFC9A" bold="true" box="[151,328,807,834]" italics="true" pageId="3" pageNumber="204">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFBDFFC9632BFCC3FE50FC99" author="Simon, E." box="[343,500,807,833]" pageId="3" pageNumber="204" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFBDFFC962EBFCAFFEB8FCBD" box="[151,284,843,869]" pageId="3" pageNumber="204" type="description">
<paragraph id="C560951AFFBDFFC962EBFCAFFEB8FCBD" blockId="3.[151,508,807,869]" box="[151,284,843,869]" pageId="3" pageNumber="204">
<figureCitation id="5DE4899FFFBDFFC962EBFCAFFEB8FCBD" box="[151,284,843,869]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="4.[151,250,1873,1897]" captionStartId-1="5.[151,250,1825,1849]" captionStartId-2="6.[151,250,1871,1895]" captionStartId-3="7.[151,250,1884,1908]" captionStartId-4="8.[151,250,1859,1883]" captionStartId-5="9.[151,250,1872,1896]" captionStartId-6="10.[151,250,1779,1803]" captionStartId-7="11.[151,250,1731,1755]" captionTargetBox-0="[159,1424,189,1844]" captionTargetBox-1="[158,1435,188,1794]" captionTargetBox-2="[159,1426,189,1841]" captionTargetBox-3="[159,1427,194,1858]" captionTargetBox-4="[159,1428,192,1829]" captionTargetBox-5="[151,1409,185,1846]" captionTargetBox-6="[151,1435,181,1756]" captionTargetBox-7="[151,1435,181,1708]" captionTargetId-0="figure-22@4.[151,1436,181,1849]" captionTargetId-1="figure-17@5.[151,1436,181,1801]" captionTargetId-2="figure-22@6.[151,1436,181,1848]" captionTargetId-3="figure-17@7.[151,1436,181,1860]" captionTargetId-4="figure-22@8.[151,1436,181,1835]" captionTargetId-5="figure-17@9.[151,1435,181,1848]" captionTargetId-6="figure-22@10.[151,1435,181,1756]" captionTargetId-7="figure-17@11.[151,1435,181,1708]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="FIGURE 1. Josa chazaliae (Simon, 1897), female lectotype (MNHN 18396; vchLAM-243):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 2. Josa chazaliae (Simon, 1897), female paralectotype (MNHN; vchLAM-242): A Carapace; B Abdomen, dorsal view; C Sternum; D Abdomen, ventral view; E Epigyne, ventral view (arrow indicates the anterior excavations); F Label of type series." captionText-2="FIGURE 3. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-3="FIGURE 4. Josa chazaliae (Simon, 1897), female (MUSENUV-Ar-2672; vchLAM-247): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-4="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." captionText-5="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." captionText-6="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." captionText-7="FIGURE 8. Josa chazaliae (Simon, 1897), female (MACN-Ar 46691; vchLAM-248) genitalia SEM images: A Dorsal view; B Dorsal view; C Detail of dorsal view; D Accessory bulb; E Posterior-ventral view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="3" pageNumber="204">Figures 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFBDFFC962EBFC74FDAEFBDC" pageId="3" pageNumber="204" type="reference_group">
<paragraph id="C560951AFFBDFFC962EBFC74FE44FC3D" blockId="3.[151,1436,911,1028]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="E5CFB234FFBDFFC962EBFC74FE44FC3D" pageId="3" pageNumber="204">
<treatmentCitation id="447EB30BFFBDFFC962EBFC74FD8CFC70" author="Simon, E." box="[151,552,912,936]" page="11" pageId="3" pageNumber="204" year="1897">
<taxonomicName id="02DFEE99FFBDFFC962EBFC74FD8CFC70" authority="Simon, 1897: 11" authorityName="Simon" authorityPageNumber="11" authorityYear="1897" box="[151,552,912,936]" class="Arachnida" family="Anyphaenidae" genus="Tomopisthes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFBDFFC962EBFC74FED8FC70" box="[151,380,912,936]" italics="true" pageId="3" pageNumber="204">Tomopisthes chazaliae</emphasis>
<materialsCitation id="75B79F47FFBDFFC963FEFC74FD8CFC70" box="[386,552,912,936]" collectionCode="MNHP" collectorName="De Dalmas" country="Colombia" location="Sierra Nevada de Santa Marta" pageId="3" pageNumber="204" specimenCode="MNHP 18396" specimenCount="3" specimenCount-female="2" stateProvince="Magdalena" typeStatus="lectotype">
<bibRefCitation id="A14EE8EBFFBDFFC963FEFC74FD8CFC70" author="Simon, E." box="[386,552,912,936]" pageId="3" pageNumber="204" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897: 11</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(female
<typeStatus id="1A642BB8FFBDFFC960FEFC74FD7BFC70" box="[642,735,912,936]" pageId="3" pageNumber="204" type="lectotype">lectotype</typeStatus>
<specimenCode id="95793D61FFBDFFC96099FC74FCD1FC7F" box="[741,885,911,935]" collectionCode="MNHP" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="3" pageNumber="204">MNHP 18396</specimenCode>
; vchLAM-243 and
<specimenCount id="D3D95E93FFBDFFC96647FC74FB12FC70" box="[1083,1206,912,936]" count="2" pageId="3" pageNumber="204" type="female">two females</specimenCount>
<typeStatus id="1A642BB8FFBDFFC966C0FC74FAEAFC70" box="[1212,1358,912,936]" pageId="3" pageNumber="204" type="paralectotype">paralectotypes</typeStatus>
MNHP vchLAM-242 and vchLAM-244 here designated, from
<location id="C000C3C1FFBDFFC96080FC4AFB94FC1E" LSID="urn:lsid:plazi:treatment:4D76240CFFBDFFC662EBFCC3FBFAFAA1:C000C3C1FFBDFFC96080FC4AFB94FC1E" box="[764,1072,942,966]" country="Colombia" name="Sierra Nevada de Santa Marta" pageId="3" pageNumber="204" stateProvince="Magdalena">Sierra Nevada de Santa Marta</location>
,
<collectingRegion id="071B5BF8FFBDFFC96647FC4AFB0AFC1E" box="[1083,1198,942,966]" country="Colombia" name="Magdalena" pageId="3" pageNumber="204">Magdalena</collectingRegion>
,
<collectingCountry id="BDC8D58AFFBDFFC966C5FC4AFA84FC1E" box="[1209,1312,942,966]" name="Colombia" pageId="3" pageNumber="204">Colombia</collectingCountry>
,
<collectorName id="682AF0CCFFBDFFC96756FC4BFA38FC1E" box="[1322,1436,942,966]" pageId="3" pageNumber="204">De Dalmas</collectorName>
coll., in MNHP, examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="C560951AFFBDFFC962EBFC08FDAEFBDC" blockId="3.[151,1436,911,1028]" box="[151,522,1004,1028]" pageId="3" pageNumber="204">
<treatmentCitationGroup id="E5CFB234FFBDFFC962EBFC08FDAEFBDC" box="[151,522,1004,1028]" pageId="3" pageNumber="204">
<taxonomicName id="02DFEE99FFBDFFC962EBFC08FE89FBDC" ID-CoL="6NG93" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[151,301,1004,1028]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFBDFFC962EBFC08FE89FBDC" box="[151,301,1004,1028]" italics="true" pageId="3" pageNumber="204">Josa chazaliae</emphasis>
</taxonomicName>
:
<treatmentCitation id="447EB30BFFBDFFC96346FC08FDA2FBDC" author="Ramirez, M. J." box="[314,518,1004,1028]" page="109" pageId="3" pageNumber="204" year="2003">
<bibRefCitation id="A14EE8EBFFBDFFC96346FC08FDA2FBDC" author="Ramirez, M. J." box="[314,518,1004,1028]" pageId="3" pageNumber="204" pagination="1 - 262" refId="ref15379" refString="Ramirez, M. J. (2003) The spider subfamily Amaurobioidinae (Araneae, Anyphaenidae): a phylogenetic revision at the generic level. Bulletin of the American Museum of natural History, 2003 (277), 1 - 262. https: // doi. org / 10.1206 / 0003 - 0090 (2003) 277 &lt;0001: TSSAAA&gt; 2.0. CO; 2" type="journal article" year="2003">Ramírez, 2003: 109</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFBDFFC962EBFBD6FBADFA6C" pageId="3" pageNumber="204" type="materials_examined">
<paragraph id="C560951AFFBDFFC962EBFBD6FAFEFB60" blockId="3.[151,1437,1074,2037]" pageId="3" pageNumber="204">
<emphasis id="F7AB4908FFBDFFC962EBFBD6FF72FB94" bold="true" box="[151,214,1074,1100]" pageId="3" pageNumber="204">Note.</emphasis>
The
<typeStatus id="1A642BB8FFBDFFC96369FBD6FED3FB94" box="[277,375,1074,1100]" pageId="3" pageNumber="204" type="syntype">syntypes</typeStatus>
correspond to two different species; the
<typeStatus id="1A642BB8FFBDFFC9612DFBD6FC1CFB94" box="[849,952,1074,1100]" pageId="3" pageNumber="204" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="A3CE0DDFFFBDFFC961B5FBD6FBBFFB94" box="[969,1051,1074,1100]" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="3" pageNumber="204">MNHP</collectionCode>
18396 vchLAM-243) and one of the
<typeStatus id="1A642BB8FFBDFFC962BDFBB2FEC7FBA8" box="[193,355,1110,1136]" pageId="3" pageNumber="204" type="paralectotype">paralectotypes</typeStatus>
belong to
<taxonomicName id="02DFEE99FFBDFFC963A6FBB3FDDBFBA8" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[474,639,1110,1136]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFBDFFC963A6FBB3FDDBFBA8" box="[474,639,1110,1136]" italics="true" pageId="3" pageNumber="204">Josa chazaliae</emphasis>
</taxonomicName>
(
<collectionCode id="A3CE0DDFFFBDFFC960F2FBB2FD44FBA8" box="[654,736,1110,1136]" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="3" pageNumber="204">MNHP</collectionCode>
vchLAM-242) and the other to an undescribed species, which is described below as
<taxonomicName id="02DFEE99FFBDFFC963E5FB9FFD97FB4C" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[409,563,1147,1172]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="samaria" status="sp. nov.">
<emphasis id="F7AB4908FFBDFFC963E5FB9FFD97FB4C" box="[409,563,1147,1172]" italics="true" pageId="3" pageNumber="204">Josa samaria</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFBDFFC96041FB9EFD3DFB4C" bold="true" box="[573,665,1146,1172]" pageId="3" pageNumber="204">
<taxonomicNameLabel id="EC98F473FFBDFFC96041FB9EFD3DFB4C" box="[573,665,1146,1172]" pageId="3" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectionCode id="A3CE0DDFFFBDFFC960D7FB9EFD59FB4C" box="[683,765,1146,1172]" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="3" pageNumber="204">MNHP</collectionCode>
vchLAM-244). Only records from the additional material examined were mapped, as the types label lacks a specific locality within the Sierra
<collectingRegion id="071B5BF8FFBDFFC96631FB7AFB07FB60" box="[1101,1187,1182,1208]" country="United States of America" name="Nevada" pageId="3" pageNumber="204">Nevada</collectingRegion>
de Santa Marta.
</paragraph>
<paragraph id="C560951AFFBDFFC962BBFB25FBADFA6C" blockId="3.[151,1437,1074,2037]" pageId="3" pageNumber="204">
<emphasis id="F7AB4908FFBDFFC962BBFB25FDA0FB03" bold="true" box="[199,516,1217,1243]" pageId="3" pageNumber="204">Other material examined:</emphasis>
<materialsCitation id="75B79F47FFBDFFC96072FB26FAE5FAFC" collectingDate="1975-05-09" collectorName="Loma de la Cebolleta &amp; J. Kochalka" country="Colombia" county="Cienaga" elevation="3505" location="Parque Natural Nacional Sierra Nevada de Santa Marta" municipality="San Pedro" pageId="3" pageNumber="204" specimenCount="2" specimenCount-male="2" stateProvince="Magdalena">
<collectingCountry id="BDC8D58AFFBDFFC96072FB26FD03FB04" box="[526,679,1218,1244]" name="Colombia" pageId="3" pageNumber="204">COLOMBIA</collectingCountry>
.
<collectingRegion id="071B5BF8FFBDFFC960C8FB26FC91FB04" box="[692,821,1218,1244]" country="Colombia" name="Magdalena" pageId="3" pageNumber="204">Magdalena</collectingRegion>
:
<collectingCounty id="2C01ED96FFBDFFC9613EFB26FC05FB04" box="[834,929,1218,1244]" pageId="3" pageNumber="204">Ciénaga</collectingCounty>
,
<collectingMunicipality id="25040F60FFBDFFC961D3FB26FB82FB04" box="[943,1062,1218,1244]" pageId="3" pageNumber="204">San Pedro</collectingMunicipality>
,
<location id="C000C3C1FFBDFFC96649FB26FE3BFAD8" LSID="urn:lsid:plazi:treatment:4D76240CFFBDFFC662EBFCC3FBFAFAA1:C000C3C1FFBDFFC96649FB26FE3BFAD8" country="Colombia" county="Cienaga" municipality="San Pedro" name="Parque Natural Nacional Sierra Nevada de Santa Marta" pageId="3" pageNumber="204" stateProvince="Magdalena">Parque Natural Nacional Sierra Nevada de Santa Marta</location>
,
<collectorName id="682AF0CCFFBDFFC963D5FB02FD3EFAD8" box="[425,666,1254,1280]" pageId="3" pageNumber="204">
<location id="C000C3C1FFBDFFC963D5FB02FD3EFAD8" LSID="urn:lsid:plazi:treatment:4D76240CFFBDFFC662EBFCC3FBFAFAA1:C000C3C1FFBDFFC963D5FB02FD3EFAD8" box="[425,666,1254,1280]" country="Colombia" county="Cienaga" municipality="San Pedro" name="Loma de la Cebolleta" pageId="3" pageNumber="204" stateProvince="Magdalena">Loma de la Cebolleta</location>
</collectorName>
, misc. vegetation, low-medium, manual caught,
<quantity id="022738FFFFBDFFC966CBFB02FAA9FAD8" box="[1207,1293,1254,1281]" metricMagnitude="3" metricUnit="m" metricValue="3.505" pageId="3" pageNumber="204" unit="m" value="3505.0">
<elevation id="4EF27229FFBDFFC966CBFB02FAA9FAD8" box="[1207,1293,1254,1281]" metricMagnitude="3" metricUnit="m" metricValue="3.505" pageId="3" pageNumber="204" unit="m" value="3505.0">3505 m</elevation>
</quantity>
,
<collectorName id="682AF0CCFFBDFFC96765FB02FA38FAD8" box="[1305,1436,1254,1280]" pageId="3" pageNumber="204">J. Kochalka</collectorName>
leg.,
<date id="B161B3DAFFBDFFC962A4FAEEFE9BFAFD" box="[216,319,1290,1317]" pageId="3" pageNumber="204" value="1975-05-09">
<collectingDate id="A1254A32FFBDFFC962A4FAEEFE9BFAFD" box="[216,319,1290,1317]" pageId="3" pageNumber="204" value="1975-05-09">9.V.1975</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFBDFFC96328FAEEFE23FAFC" box="[340,391,1290,1316]" count="1" pageId="3" pageNumber="204" type="male">1 ♂</specimenCount>
(MUSENUV-Ar-2671; vchLAM-245),
<specimenCount id="D3D95E93FFBDFFC9611FFAEEFC33FAFC" box="[867,919,1290,1316]" count="1" pageId="3" pageNumber="204" type="male">1 ♂</specimenCount>
(MACN-Ar 46690; vchLAM-246)
</materialsCitation>
.
<materialsCitation id="75B79F47FFBDFFC96729FAEEFBA1FA6C" collectingDate="1973-01-02" collectingDateMax="1974-01-18" collectingDateMin="1973-01-02" collectionCode="NW" country="Colombia" county="Valledupar" elevation="3974" latitude="10.797223" location="Meoyaca" longLatPrecision="1" longitude="-73.62625" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" pageId="3" pageNumber="204" specimenCount="4" specimenCount-female="4" stateProvince="Cesar">
<collectingRegion id="071B5BF8FFBDFFC96729FAEEFA3DFAFC" box="[1365,1433,1290,1316]" country="Colombia" name="Cesar" pageId="3" pageNumber="204">Cesar</collectingRegion>
:
<collectingCounty id="2C01ED96FFBDFFC962EBFACAFEB6FA90" box="[151,274,1326,1352]" pageId="3" pageNumber="204">Valledupar</collectingCounty>
,
<collectingMunicipality id="25040F60FFBDFFC96360FACAFC26FA90" box="[284,898,1326,1352]" pageId="3" pageNumber="204">Parque Nacional Natural Sierra Nevada de Santa Marta</collectingMunicipality>
(
<collectionCode id="A3CE0DDFFFBDFFC961F3FACAFC1BFA90" box="[911,959,1326,1352]" pageId="3" pageNumber="204">NW</collectionCode>
Donachuí),
<location id="C000C3C1FFBDFFC96634FACAFB16FA90" LSID="urn:lsid:plazi:treatment:4D76240CFFBDFFC662EBFCC3FBFAFAA1:C000C3C1FFBDFFC96634FACAFB16FA90" box="[1096,1202,1326,1352]" country="Colombia" county="Valledupar" latitude="10.797223" longLatPrecision="1" longitude="-73.62625" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" name="Meoyaca" pageId="3" pageNumber="204" stateProvince="Cesar">Meoyaca</location>
,
<location id="C000C3C1FFBDFFC966C0FACAFAFEFA90" LSID="urn:lsid:plazi:treatment:4D76240CFFBDFFC662EBFCC3FBFAFAA1:C000C3C1FFBDFFC966C0FACAFAFEFA90" box="[1212,1370,1326,1352]" country="Colombia" county="Valledupar" latitude="10.797223" longLatPrecision="1" longitude="-73.62625" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" name="Rio Donachui" pageId="3" pageNumber="204" stateProvince="Cesar">Río Donachuí</location>
,
<quantity id="022738FFFFBDFFC96718FACAFF0BFAB4" metricMagnitude="3" metricUnit="m" metricValue="3.9739999999999993" pageId="3" pageNumber="204" unit="m" value="3974.0">
<elevation id="4EF27229FFBDFFC96718FACAFF0BFAB4" metricMagnitude="3" metricUnit="m" metricValue="3.9739999999999993" pageId="3" pageNumber="204" unit="m" value="3974.0">3974 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFBDFFC962BEFAB7FEC6FAB6" box="[194,354,1363,1390]" degrees="10" direction="north" minutes="47" orientation="latitude" pageId="3" pageNumber="204" precision="1" seconds="39.9" value="10.794417">10°4739.9″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFBDFFC96312FAB7FDB0FAB6" box="[366,532,1363,1390]" degrees="73" direction="west" minutes="36" orientation="longitude" pageId="3" pageNumber="204" precision="1" seconds="03.6" value="-73.601">73°3603.6″W</geoCoordinate>
],
<date id="B161B3DAFFBDFFC9605AFAB6FD37FAB4" box="[550,659,1362,1388]" pageId="3" pageNumber="204" value="1974-01-18">
<collectingDate id="A1254A32FFBDFFC9605AFAB6FD37FAB4" box="[550,659,1362,1388]" pageId="3" pageNumber="204" value="1974-01-18">18.I.1974</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFBDFFC960DCFAB6FD6FFAB4" box="[672,715,1362,1388]" count="1" pageId="3" pageNumber="204" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2672; vchLAM-247);
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<elevation id="4EF27229FFBDFFC966F2FAB6FB41FAB4" box="[1166,1253,1362,1389]" metricMagnitude="3" metricUnit="m" metricValue="3.596" pageId="3" pageNumber="204" unit="m" value="3596.0">3596 m</elevation>
</quantity>
, [
<geoCoordinate id="A0EBF3DDFFBDFFC96684FAB7FA3CFAB6" box="[1272,1432,1363,1390]" degrees="10" direction="north" minutes="47" orientation="latitude" pageId="3" pageNumber="204" precision="1" seconds="04.5" value="10.784584">10°4704.5″N</geoCoordinate>
,
<geoCoordinate id="A0EBF3DDFFBDFFC962EBFA93FE99FA4A" box="[151,317,1399,1426]" degrees="73" direction="west" minutes="33" orientation="longitude" pageId="3" pageNumber="204" precision="1" seconds="40.4" value="-73.561226">73°3340.4″W</geoCoordinate>
],
<date id="B161B3DAFFBDFFC96331FA92FE08FA48" box="[333,428,1398,1424]" pageId="3" pageNumber="204" value="1973-01-02">
<collectingDate id="A1254A32FFBDFFC96331FA92FE08FA48" box="[333,428,1398,1424]" pageId="3" pageNumber="204" value="1973-01-02">2.I.1973</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFBDFFC963CAFA92FE79FA48" box="[438,477,1398,1424]" count="2" pageId="3" pageNumber="204" type="female">2 ♀</specimenCount>
(MACN-Ar 46691; vchLAM-248); Donachuí,
<location id="C000C3C1FFBDFFC9619AFA92FB27FA48" LSID="urn:lsid:plazi:treatment:4D76240CFFBDFFC662EBFCC3FBFAFAA1:C000C3C1FFBDFFC9619AFA92FB27FA48" box="[998,1155,1398,1424]" country="Colombia" county="Valledupar" latitude="10.797223" longLatPrecision="1" longitude="-73.62625" municipality="Parque Nacional Natural Sierra Nevada de Santa Marta" name="Rio Donachui" pageId="3" pageNumber="204" stateProvince="Cesar">Rio Donachuí</location>
, top of woody vegetation,
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</quantity>
, [
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,
<geoCoordinate id="A0EBF3DDFFBDFFC963D0FA7FFDF5FA6E" box="[428,593,1435,1462]" degrees="73" direction="west" minutes="37" orientation="longitude" pageId="3" pageNumber="204" precision="1" seconds="34.5" value="-73.62625">73°3734.5″W</geoCoordinate>
],
<date id="B161B3DAFFBDFFC9601FFA7EFD67FA6C" box="[611,707,1434,1460]" pageId="3" pageNumber="204" value="1973-01-06">
<collectingDate id="A1254A32FFBDFFC9601FFA7EFD67FA6C" box="[611,707,1434,1460]" pageId="3" pageNumber="204" value="1973-01-06">6.I.1973</collectingDate>
</date>
,
<specimenCount id="D3D95E93FFBDFFC960B3FA7EFD5DFA6C" box="[719,761,1434,1460]" count="1" pageId="3" pageNumber="204" type="female">1 ♀</specimenCount>
(MUSENUV-Ar-2673)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFBDFFC962BBFA59FA9CF920" pageId="3" pageNumber="204" type="diagnosis">
<paragraph id="C560951AFFBDFFC962BBFA59FA9CF920" blockId="3.[151,1437,1074,2037]" pageId="3" pageNumber="204">
<emphasis id="F7AB4908FFBDFFC962BBFA59FEE6FA0F" bold="true" box="[199,322,1469,1495]" pageId="3" pageNumber="204">Diagnosis.</emphasis>
Males of
<taxonomicName id="02DFEE99FFBDFFC963CFFA5BFDFDFA00" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[435,601,1470,1496]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFBDFFC963CFFA5BFDFDFA00" box="[435,601,1470,1496]" italics="true" pageId="3" pageNumber="204">Josa chazaliae</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="02DFEE99FFBDFFC96122FA5BFC5CFA00" authorityName="Martínez &amp; Kochalka &amp; Cabra-Garcia &amp; Ramírez" authorityYear="2025" box="[862,1016,1470,1496]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="204" phylum="Arthropoda" rank="species" species="berlandi" status="sp. nov.">
<emphasis id="F7AB4908FFBDFFC96122FA5BFC5CFA00" box="[862,1016,1470,1496]" italics="true" pageId="3" pageNumber="204">Josa berlandi</emphasis>
</taxonomicName>
<emphasis id="F7AB4908FFBDFFC96183FA5AFBFFFA00" bold="true" box="[1023,1115,1470,1496]" pageId="3" pageNumber="204">
<taxonomicNameLabel id="EC98F473FFBDFFC96183FA5AFBFFFA00" box="[1023,1115,1470,1496]" pageId="3" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the small conductor with the external region apically bifid and by the short embolus (
<figureCitation id="5DE4899FFFBDFFC9614CFA06FC00FA24" box="[816,932,1506,1532]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="3" pageNumber="204">Figs 7HI</figureCitation>
, compare with
<figureCitation id="5DE4899FFFBDFFC96633FA06FB7FFA25" box="[1103,1243,1506,1533]" captionStart="FIGURE 25" captionStartId="33.[151,250,1836,1860]" captionTargetBox="[151,1436,212,1812]" captionTargetId="figure-17@33.[151,1436,181,1812]" captionTargetPageId="33" captionText="FIGURE 25. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): E Ventral view; F Dorsal view; Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum, T, tegulum." pageId="3" pageNumber="204">Figs 25AD</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC9669AFA06FA9EFA24" box="[1254,1338,1506,1532]" captionStart="FIGURE 26" captionStartId="34.[151,250,1866,1890]" captionTargetBox="[158,1427,185,1840]" captionTargetId="figure-22@34.[151,1435,181,1842]" captionTargetPageId="34" captionText="FIGURE 26. Josa berlandi sp. nov., male (MUSENUV-Ar-2675; vchLAM-253) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female genitalia (MUSENUV-Ar-2676; vchLAM-255): D Ventral view; E Dorsal view. Abbreviations: AB, accessory bulb; C, conductor; CD, copulatory duct; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="3" pageNumber="204">26AC</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC96738FA06FA2BFA24" box="[1348,1423,1506,1532]" captionStart="FIGURE 27" captionStartId="35.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-17@35.[151,1435,181,1756]" captionTargetPageId="35" captionText="FIGURE 27. Josa berlandi sp. nov., male (MACN-Ar 46694; vchLAM-254) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus; C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Details of embolus, prolateral view; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; St, subtegulum; T, tegulum." pageId="3" pageNumber="204">27AI</figureCitation>
), but can be differentiated from this species by the conductor widening towards the apex with both apical tips of the external section of the conductor of about the same length, the posterior one curved towards the internal region and with a subapical pronounced bump (
<figureCitation id="5DE4899FFFBDFFC96050F9AAFD03F9B0" box="[556,679,1614,1640]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="3" pageNumber="204">Fig. 7AD</figureCitation>
, arrow), median apophysis bifid with its anterior branch tubular and slightly curved (
<figureCitation id="5DE4899FFFBDFFC96333F996FE09F955" box="[335,429,1650,1677]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="3" pageNumber="204">Figs 5A</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC963C6F996FE7BF954" box="[442,479,1650,1676]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="3" pageNumber="204">6A</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC96397F996FD97F954" box="[491,563,1650,1676]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="3" pageNumber="204">7AD</figureCitation>
), and the very small and rugous retrobasal tegular lobe (
<figureCitation id="5DE4899FFFBDFFC966CDF996FA8EF954" box="[1201,1322,1650,1676]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="3" pageNumber="204">Fig. 7A, F</figureCitation>
). Females are easily distinguishable by the very thin and short ducts of the accessory bulb restricted to the lateral regions of the vulva (
<figureCitation id="5DE4899FFFBDFFC96370F95EFE2CF90C" box="[268,392,1722,1748]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="3" pageNumber="204">Figs 6DE</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC963EFF95EFE7FF90C" box="[403,475,1722,1748]" captionStart="FIGURE 8" captionStartId="11.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@11.[151,1435,181,1708]" captionTargetPageId="11" captionText="FIGURE 8. Josa chazaliae (Simon, 1897), female (MACN-Ar 46691; vchLAM-248) genitalia SEM images: A Dorsal view; B Dorsal view; C Detail of dorsal view; D Accessory bulb; E Posterior-ventral view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="3" pageNumber="204">8AD</figureCitation>
), the swollen, triangular basal sclerotized projections of the epigyne (
<figureCitation id="5DE4899FFFBDFFC966A3F95EFA9DF90C" box="[1247,1337,1722,1748]" captionStart="FIGURE 2" captionStartId="5.[151,250,1825,1849]" captionTargetBox="[158,1435,188,1794]" captionTargetId="figure-17@5.[151,1436,181,1801]" captionTargetPageId="5" captionText="FIGURE 2. Josa chazaliae (Simon, 1897), female paralectotype (MNHN; vchLAM-242): A Carapace; B Abdomen, dorsal view; C Sternum; D Abdomen, ventral view; E Epigyne, ventral view (arrow indicates the anterior excavations); F Label of type series." pageId="3" pageNumber="204">Figs 2E</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC96738F95EFAC2F90D" box="[1348,1382,1722,1749]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="3" pageNumber="204">5E</figureCitation>
), the long median septum, and median field oval with deep copulatory openings (
<figureCitation id="5DE4899FFFBDFFC96196F93AFBE0F920" box="[1002,1092,1758,1784]" captionStart="FIGURE 2" captionStartId="5.[151,250,1825,1849]" captionTargetBox="[158,1435,188,1794]" captionTargetId="figure-17@5.[151,1436,181,1801]" captionTargetPageId="5" captionText="FIGURE 2. Josa chazaliae (Simon, 1897), female paralectotype (MNHN; vchLAM-242): A Carapace; B Abdomen, dorsal view; C Sternum; D Abdomen, ventral view; E Epigyne, ventral view (arrow indicates the anterior excavations); F Label of type series." pageId="3" pageNumber="204">Figs 2E</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC9662CF93AFB29F921" box="[1104,1165,1758,1785]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="3" pageNumber="204">5EF</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC966E4F93AFB78F920" box="[1176,1244,1758,1784]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="3" pageNumber="204">6DE</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC96694F93AFA88F920" box="[1256,1324,1758,1784]" captionStart="FIGURE 8" captionStartId="11.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@11.[151,1435,181,1708]" captionTargetPageId="11" captionText="FIGURE 8. Josa chazaliae (Simon, 1897), female (MACN-Ar 46691; vchLAM-248) genitalia SEM images: A Dorsal view; B Dorsal view; C Detail of dorsal view; D Accessory bulb; E Posterior-ventral view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="3" pageNumber="204">8AF</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFBDFFC662BBF8E5FBF8FA8D" lastPageId="12" lastPageNumber="213" pageId="3" pageNumber="204" type="description">
<paragraph id="C560951AFFBDFFC662BBF8E5FEDEFDE5" blockId="3.[151,1437,1074,2037]" lastBlockId="12.[151,1437,151,1401]" lastPageId="12" lastPageNumber="213" pageId="3" pageNumber="204">
<emphasis id="F7AB4908FFBDFFC962BBF8E5FEFDF8C3" bold="true" box="[199,345,1793,1819]" pageId="3" pageNumber="204">Description.</emphasis>
<emphasis id="F7AB4908FFBDFFC96321F8E6FE15F8C4" box="[349,433,1794,1820]" italics="true" pageId="3" pageNumber="204">Female</emphasis>
(description based on the
<typeStatus id="1A642BB8FFBDFFC960B4F8E6FC96F8C4" box="[712,818,1794,1820]" pageId="3" pageNumber="204" type="lectotype">lectotype</typeStatus>
, MNHP 18396; vchLAM-243 and a female MUSENUV- Ar-2671; vchLAM-245). (
<figureCitation id="5DE4899FFFBDFFC963BCF8C2FD82F898" box="[448,550,1830,1856]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1873,1897]" captionStartId-1="5.[151,250,1825,1849]" captionTargetBox-0="[159,1424,189,1844]" captionTargetBox-1="[158,1435,188,1794]" captionTargetId-0="figure-22@4.[151,1436,181,1849]" captionTargetId-1="figure-17@5.[151,1436,181,1801]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Josa chazaliae (Simon, 1897), female lectotype (MNHN 18396; vchLAM-243):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." captionText-1="FIGURE 2. Josa chazaliae (Simon, 1897), female paralectotype (MNHN; vchLAM-242): A Carapace; B Abdomen, dorsal view; C Sternum; D Abdomen, ventral view; E Epigyne, ventral view (arrow indicates the anterior excavations); F Label of type series." pageId="3" pageNumber="204">Figs 12</figureCitation>
,
<figureCitation id="5DE4899FFFBDFFC9604FF8C2FDE0F898" box="[563,580,1830,1856]" captionStart="FIGURE 4" captionStartId="7.[151,250,1884,1908]" captionTargetBox="[159,1427,194,1858]" captionTargetId="figure-17@7.[151,1436,181,1860]" captionTargetPageId="7" captionText="FIGURE 4. Josa chazaliae (Simon, 1897), female (MUSENUV-Ar-2672; vchLAM-247): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="3" pageNumber="204">4</figureCitation>
): Total length 7.42. Carapace length 3.70, width 2.56, height 1.64. Clypeus length 0.22. Sternum length 1.88, width 1.29. Legs. Leg I: ti length 1.90/width 0.51, mt 1.53. II: ti 1.81/0.47, mt 1.49. III: ti 1.61/0.41, mt 1.57. IV: ti 2.21/0.46, mt 2.18. Spines: legs III, femur d 1-1-1, p 2ap; tibia v 2-2-2; metatarsus v 2bas. III, femur d 1-1-1, p 0-d1-d1, r 0-d1-d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-0-2, p d1-1-0-1, r d1-1-0-1. IV=III but femur r d1ap; patella d 1-1, r 1; metatarsus v 2-2-2. Abdomen: length 3.75, width 1.86, spiracleepigastrium 1.26. Body dehydrated, damaged. Color: carapace dark brown, with two wide longitudinal dark bands on sides, narrowed in cephalic region. Ocular area on dark patch with eyes bordered by black rings. Carapace with fringe of white setae on sides. Clypeus brown, low, with some white setae. Chelicerae brown with black patches and fine setae on front. Endites, labium, and sternum dark brown, labium and endites light in apical region. Legs dark brown, with long darker stripes on dorsal edges of femora. All articles covered by white setae. Abdomen dorsally with median spots on dark gray background and abundant white setae; anterior region with bunch of thick setae; venter uniformly dark gray, with light dots. Epigyne: anterior plate sclerotized with deep anterior excavations, median notch long and narrow (
<figureCitation id="5DE4899FFFB2FFC66678FE8BFBC6FE51" box="[1028,1122,367,393]" captionStart="FIGURE 2" captionStartId="5.[151,250,1825,1849]" captionTargetBox="[158,1435,188,1794]" captionTargetId="figure-17@5.[151,1436,181,1801]" captionTargetPageId="5" captionText="FIGURE 2. Josa chazaliae (Simon, 1897), female paralectotype (MNHN; vchLAM-242): A Carapace; B Abdomen, dorsal view; C Sternum; D Abdomen, ventral view; E Epigyne, ventral view (arrow indicates the anterior excavations); F Label of type series." pageId="12" pageNumber="213">Figs 2E</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC6660DFE8AFB37FE51" box="[1137,1171,366,393]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">5E</figureCitation>
, arrows). Median field transverse, large, oval in posterior view, with medial depression (
<figureCitation id="5DE4899FFFB2FFC66127FE77FC69FE75" box="[859,973,403,429]" captionStart="FIGURE 8" captionStartId="11.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@11.[151,1435,181,1708]" captionTargetPageId="11" captionText="FIGURE 8. Josa chazaliae (Simon, 1897), female (MACN-Ar 46691; vchLAM-248) genitalia SEM images: A Dorsal view; B Dorsal view; C Detail of dorsal view; D Accessory bulb; E Posterior-ventral view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="12" pageNumber="213">Fig. 8EF</figureCitation>
). Lateral lobes with sclerotized extensions very wide, forming a longitudinal concavity, posterior extensions rounded at base (
<figureCitation id="5DE4899FFFB2FFC66641FE53FB3CFE09" box="[1085,1176,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1825,1849]" captionTargetBox="[158,1435,188,1794]" captionTargetId="figure-17@5.[151,1436,181,1801]" captionTargetPageId="5" captionText="FIGURE 2. Josa chazaliae (Simon, 1897), female paralectotype (MNHN; vchLAM-242): A Carapace; B Abdomen, dorsal view; C Sternum; D Abdomen, ventral view; E Epigyne, ventral view (arrow indicates the anterior excavations); F Label of type series." pageId="12" pageNumber="213">Figs 2E</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC666D9FE52FB63FE09" box="[1189,1223,438,465]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">5E</figureCitation>
). Copulatory ducts long, touching with the accessory bulb ducts medially (
<figureCitation id="5DE4899FFFB2FFC66179FE3FFCF8FE2D" box="[773,860,475,501]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">Fig. 6E</figureCitation>
). Spermathecae rounded medially positioned in the epigyne, oriented towards median region (
<figureCitation id="5DE4899FFFB2FFC66017FE1BFD65FDC1" box="[619,705,510,537]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Figs 5F</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC660B7FE1BFD49FDC1" box="[715,749,511,537]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">6E</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC6608BFE1BFC99FDC1" box="[759,829,511,537]" captionStart="FIGURE 8" captionStartId="11.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@11.[151,1435,181,1708]" captionTargetPageId="11" captionText="FIGURE 8. Josa chazaliae (Simon, 1897), female (MACN-Ar 46691; vchLAM-248) genitalia SEM images: A Dorsal view; B Dorsal view; C Detail of dorsal view; D Accessory bulb; E Posterior-ventral view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="12" pageNumber="213">8AE</figureCitation>
). Ducts of accessory bulbs long, thin, convergent (
<figureCitation id="5DE4899FFFB2FFC66716FE1BFF1DFDE5" captionStart="FIGURE 2" captionStartId="5.[151,250,1825,1849]" captionTargetBox="[158,1435,188,1794]" captionTargetId="figure-17@5.[151,1436,181,1801]" captionTargetPageId="5" captionText="FIGURE 2. Josa chazaliae (Simon, 1897), female paralectotype (MNHN; vchLAM-242): A Carapace; B Abdomen, dorsal view; C Sternum; D Abdomen, ventral view; E Epigyne, ventral view (arrow indicates the anterior excavations); F Label of type series." pageId="12" pageNumber="213">Figs 2E</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC662B8FDC6FEA5FDE5" box="[196,257,546,573]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">5EF</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC66371FDC7FEF5FDE5" box="[269,337,547,573]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">6DE</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC66320FDC7FEC8FDE5" box="[348,364,547,573]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="12" pageNumber="213">7</figureCitation>
).
</paragraph>
<caption id="91A0C592FFBAFFCE62EBF8B5FD0BF855" pageId="4" pageNumber="205" startId="4.[151,250,1873,1897]" targetBox="[159,1424,189,1844]" targetPageId="4" targetType="figure">
<paragraph id="C560951AFFBAFFCE62EBF8B5FD0BF855" blockId="4.[151,1436,1873,1933]" pageId="4" pageNumber="205">
<emphasis id="F7AB4908FFBAFFCE62EBF8B5FEB5F8B1" bold="true" box="[151,273,1873,1897]" pageId="4" pageNumber="205">FIGURE 1.</emphasis>
<taxonomicName id="02DFEE99FFBAFFCE6369F8B6FDE4F8B1" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[277,576,1873,1897]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFBAFFCE6369F8B6FE0DF8B1" box="[277,425,1873,1897]" italics="true" pageId="4" pageNumber="205">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFBAFFCE63C8F8B5FD9CF8B1" author="Simon, E." box="[436,568,1873,1897]" pageId="4" pageNumber="205" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</taxonomicName>
, female lectotype (MNHN 18396; vchLAM-243):A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFBBFFCF62EBF8C5FF7CF859" pageId="5" pageNumber="206" startId="5.[151,250,1825,1849]" targetBox="[158,1435,188,1794]" targetPageId="5" targetType="figure">
<paragraph id="C560951AFFBBFFCF62EBF8C5FF7CF859" blockId="5.[151,1437,1825,1921]" pageId="5" pageNumber="206">
<emphasis id="F7AB4908FFBBFFCF62EBF8C5FEB1F8E1" bold="true" box="[151,277,1825,1849]" pageId="5" pageNumber="206">FIGURE 2.</emphasis>
<taxonomicName id="02DFEE99FFBBFFCF6362F8C6FDF3F8E2" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[286,599,1825,1850]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFBBFFCF6362F8C6FE13F8E1" box="[286,439,1825,1849]" italics="true" pageId="5" pageNumber="206">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFBBFFCF63BBF8C5FDF4F8E2" author="Simon, E." box="[455,592,1825,1850]" pageId="5" pageNumber="206" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</taxonomicName>
, female paralectotype (MNHN; vchLAM-242): A Carapace; B Abdomen, dorsal view; C Sternum; D Abdomen, ventral view; E Epigyne, ventral view (arrow indicates the anterior excavations); F Label of type series.
</paragraph>
</caption>
<caption id="91A0C592FFB8FFCC62EBF8ABFD3CF854" pageId="6" pageNumber="207" startId="6.[151,250,1871,1895]" targetBox="[159,1426,189,1841]" targetPageId="6" targetType="figure">
<paragraph id="C560951AFFB8FFCC62EBF8ABFD3CF854" blockId="6.[151,1436,1871,1932]" pageId="6" pageNumber="207">
<emphasis id="F7AB4908FFB8FFCC62EBF8ABFEB6F8BF" bold="true" box="[151,274,1871,1895]" pageId="6" pageNumber="207">FIGURE 3.</emphasis>
<taxonomicName id="02DFEE99FFB8FFCC6364F8B4FDECF8B0" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[280,584,1872,1896]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFB8FFCC6364F8B4FE0AF8B0" box="[280,430,1872,1896]" italics="true" pageId="6" pageNumber="207">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFB8FFCC63C7F8B4FDE4F8B0" author="Simon, E." box="[443,576,1872,1896]" pageId="6" pageNumber="207" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</taxonomicName>
, male (MUSENUV-Ar-2671; vchLAM-245): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFB9FFCD62EBF8B8FD0BF840" pageId="7" pageNumber="208" startId="7.[151,250,1884,1908]" targetBox="[159,1427,194,1858]" targetPageId="7" targetType="figure">
<paragraph id="C560951AFFB9FFCD62EBF8B8FD0BF840" blockId="7.[151,1437,1884,1944]" pageId="7" pageNumber="208">
<emphasis id="F7AB4908FFB9FFCD62EBF8B8FEB7F8AC" bold="true" box="[151,275,1884,1908]" pageId="7" pageNumber="208">FIGURE 4.</emphasis>
<taxonomicName id="02DFEE99FFB9FFCD6365F8B9FDEDF8AC" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[281,585,1884,1908]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFB9FFCD6365F8B9FE0AF8AC" box="[281,430,1884,1908]" italics="true" pageId="7" pageNumber="208">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFB9FFCD63C0F8B8FDE5F8AC" author="Simon, E." box="[444,577,1884,1908]" pageId="7" pageNumber="208" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</taxonomicName>
, female (MUSENUV-Ar-2672; vchLAM-247): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum.
</paragraph>
</caption>
<caption id="91A0C592FFB6FFC262EBF8A7FCA7F833" pageId="8" pageNumber="209" startId="8.[151,250,1859,1883]" targetBox="[159,1428,192,1829]" targetPageId="8" targetType="figure">
<paragraph id="C560951AFFB6FFC262EBF8A7FCA7F833" blockId="8.[151,1436,1859,2027]" pageId="8" pageNumber="209">
<emphasis id="F7AB4908FFB6FFC262EBF8A7FEB0F883" bold="true" box="[151,276,1859,1883]" pageId="8" pageNumber="209">FIGURE 5.</emphasis>
<taxonomicName id="02DFEE99FFB6FFC26367F8A0FDF4F883" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[283,592,1859,1883]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFB6FFC26367F8A0FE16F883" box="[283,434,1859,1883]" italics="true" pageId="8" pageNumber="209">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFB6FFC263BDF8A7FDECF883" author="Simon, E." box="[449,584,1859,1883]" pageId="8" pageNumber="209" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</taxonomicName>
, male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view.
<emphasis id="F7AB4908FFB6FFC2608BF86FFC3CF87B" bold="true" box="[759,920,1931,1955]" pageId="8" pageNumber="209">Abbreviations:</emphasis>
C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe.
</paragraph>
</caption>
<caption id="91A0C592FFB7FFC362EBF8B4FC9DF820" pageId="9" pageNumber="210" startId="9.[151,250,1872,1896]" targetBox="[151,1409,185,1846]" targetPageId="9" targetType="figure">
<paragraph id="C560951AFFB7FFC362EBF8B4FC9DF820" blockId="9.[151,1436,1872,2040]" pageId="9" pageNumber="210">
<emphasis id="F7AB4908FFB7FFC362EBF8B4FEB0F8B0" bold="true" box="[151,276,1872,1896]" pageId="9" pageNumber="210">FIGURE 6.</emphasis>
<taxonomicName id="02DFEE99FFB7FFC36360F8B5FDF6F8B0" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[284,594,1872,1896]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFB7FFC36360F8B5FE10F8B0" box="[284,436,1872,1896]" italics="true" pageId="9" pageNumber="210">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFB7FFC363BFF8B4FDEEF8B0" author="Simon, E." box="[451,586,1872,1896]" pageId="9" pageNumber="210" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</taxonomicName>
, male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology.
<emphasis id="F7AB4908FFB7FFC3630DF87CFDB6F868" bold="true" box="[369,530,1944,1968]" pageId="9" pageNumber="210">Abbreviations:</emphasis>
AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct.
</paragraph>
</caption>
<caption id="91A0C592FFB4FFC062EBF917FB57F844" pageId="10" pageNumber="211" startId="10.[151,250,1779,1803]" targetBox="[151,1435,181,1756]" targetPageId="10" targetType="figure">
<paragraph id="C560951AFFB4FFC062EBF917FB57F844" blockId="10.[151,1436,1779,1948]" pageId="10" pageNumber="211">
<emphasis id="F7AB4908FFB4FFC062EBF917FEB5F8D4" bold="true" box="[151,273,1779,1804]" pageId="10" pageNumber="211">FIGURE 7.</emphasis>
<taxonomicName id="02DFEE99FFB4FFC0636AF910FDE7F8D4" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[278,579,1780,1804]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFB4FFC0636AF910FE0FF8D4" box="[278,427,1780,1804]" italics="true" pageId="10" pageNumber="211">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFB4FFC063CBF910FD98F8D4" author="Simon, E." box="[439,572,1780,1804]" pageId="10" pageNumber="211" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</taxonomicName>
, male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images: A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE.
<emphasis id="F7AB4908FFB4FFC06094F8BBFC2DF8AF" bold="true" box="[744,905,1887,1911]" pageId="10" pageNumber="211">Abbreviations:</emphasis>
C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum.
</paragraph>
</caption>
<caption id="91A0C592FFB5FFC162EBF927FEB9F890" pageId="11" pageNumber="212" startId="11.[151,250,1731,1755]" targetBox="[151,1435,181,1708]" targetPageId="11" targetType="figure">
<paragraph id="C560951AFFB5FFC162EBF927FEB9F890" blockId="11.[151,1436,1731,1864]" pageId="11" pageNumber="212">
<emphasis id="F7AB4908FFB5FFC162EBF927FEB0F904" bold="true" box="[151,276,1731,1756]" pageId="11" pageNumber="212">FIGURE 8.</emphasis>
<taxonomicName id="02DFEE99FFB5FFC16360F920FDF5F904" authority="(Simon, 1897)" baseAuthorityName="Simon" baseAuthorityYear="1897" box="[284,593,1732,1756]" class="Arachnida" family="Anyphaenidae" genus="Josa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="chazaliae">
<emphasis id="F7AB4908FFB5FFC16360F920FE17F904" box="[284,435,1732,1756]" italics="true" pageId="11" pageNumber="212">Josa chazaliae</emphasis>
(
<bibRefCitation id="A14EE8EBFFB5FFC163BEF920FDEDF904" author="Simon, E." box="[450,585,1732,1756]" pageId="11" pageNumber="212" refId="ref15550" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, 192 pp. https: // doi. org / 10.5962 / bhl. title. 51973" type="book" year="1897">Simon, 1897</bibRefCitation>
)
</taxonomicName>
, female (MACN-Ar 46691; vchLAM-248) genitalia SEM images: A Dorsal view; B Dorsal view; C Detail of dorsal view; D Accessory bulb; E Posterior-ventral view; F Posterior view.
<emphasis id="F7AB4908FFB5FFC166B6F903FACFF927" bold="true" box="[1226,1387,1767,1791]" pageId="11" pageNumber="212">Abbreviations:</emphasis>
AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca.
</paragraph>
</caption>
<paragraph id="C560951AFFB2FFC662BBFDA3FC6FFB31" blockId="12.[151,1437,151,1401]" pageId="12" pageNumber="213">
<emphasis id="F7AB4908FFB2FFC662BBFDA3FEA5FDB9" box="[199,257,583,609]" italics="true" pageId="12" pageNumber="213">Male</emphasis>
(MUSENUV-Ar-2671; vchLAM-245) (
<figureCitation id="5DE4899FFFB2FFC660B3FDA3FCB8FDB9" box="[719,796,583,609]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1895]" captionTargetBox="[159,1426,189,1841]" captionTargetId="figure-22@6.[151,1436,181,1848]" captionTargetPageId="6" captionText="FIGURE 3. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245): A Habitus, dorsal view; B Carapace; C Frontal view; D Habitus, ventral view; E Sternum." pageId="12" pageNumber="213">Figs 3</figureCitation>
): Total length 4.54. Carapace length 2.34, width 1.61, height 0.72. Clypeus length 0.10. Sternum length 1.20, width 0.79. Legs. Leg I: ti length 2.17/width 0.26, mt 1.88. II: ti 1.97/0.25, mt 1.72. III: ti 1.32/0.23, mt 1.20. IV: ti 1.62/0.27, mt 1.55. Spines: legs s III, femur d 1-1-1, p 2ap, r 0-d1-d1; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0; metatarsus v 2-2-0, p d1-d1-0, r d1-d1. IIIIV, femur d 1-1-1, p 0- d1-d1, r 0-d1-d1; patella r 1me; tibia v 2-2-2, p 1-d1-1-0, r 1-d1-1-0, d (r1-1) bristles; metatarsus d 0-p1-2, v 2-2-2, p d1-1-0-1, r d1-1-0-1. Abdomen: length 2.34, width 1.35, spiracleepigastrium 0.83. Color: carapace brown, with two faint dark bands on sides, anteriorly covered by white setae. Ocular area brown with eyes bordered in black rings. Clypeus brown, low, with abundant white setae. Chelicerae brown with black stripes and fine setae on front. Endites, labium, and sternum brown; labium and sternum darker. Legs coxae-tarsi brown, with some tenuous dark patches. Articles with a few white and black setae. Abdomen dehydrated, with a pattern of faint spots on a brown background. Dorsum covered with few fine white setae. Venter uniformly brown. Palp: tegulum with small basal retrolateral lobe, very sclerotized, rugous (
<figureCitation id="5DE4899FFFB2FFC66012FC37FD6EFC35" box="[622,714,978,1005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Figs 5A</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC660A9FC37FD5EFC35" box="[725,762,979,1005]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">6A</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC66179FC37FC8EFC35" box="[773,810,979,1005]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="12" pageNumber="213">7A</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC66148FC37FCDCFC35" box="[820,888,979,1005]" captionStart="FIGURE 8" captionStartId="11.[151,250,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-17@11.[151,1435,181,1708]" captionTargetPageId="11" captionText="FIGURE 8. Josa chazaliae (Simon, 1897), female (MACN-Ar 46691; vchLAM-248) genitalia SEM images: A Dorsal view; B Dorsal view; C Detail of dorsal view; D Accessory bulb; E Posterior-ventral view; F Posterior view. Abbreviations: AB, accessory bulb; CD, copulatory duct; dAB, duct of accessory bulb; FD, fertilization duct; LL, lateral lobe; MF, median field; S, spermatheca." pageId="12" pageNumber="213">8AF</figureCitation>
). Median apophysis large, bifid, anterior tip long, tubular, hook-shaped, posterior branch membranous at base with external border sub-rounded (
<figureCitation id="5DE4899FFFB2FFC666CFFC13FAABFBC9" box="[1203,1295,1014,1041]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Figs 5A</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC66766FC13FA9BFBC9" box="[1306,1343,1015,1041]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">6A</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC66735FC13FA35FBC9" box="[1353,1425,1015,1041]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="12" pageNumber="213">7AB</figureCitation>
). Conductor small, rounded, with projected base, external section wide, anteriorly swollen, bifid, with both tips equal in length, but posterior tip curved towards internal region (
<figureCitation id="5DE4899FFFB2FFC6610EFBDBFBABFB81" box="[882,1039,1087,1113]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="12" pageNumber="213">Fig. 7AB, D</figureCitation>
); internal section ending as a sharp tip (
<figureCitation id="5DE4899FFFB2FFC662BAFB87FE85FBA5" box="[198,289,1123,1149]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="12" pageNumber="213">Fig. 7D</figureCitation>
). Paramedian apophysis cup-shaped, small, heavily sclerotized, concave, apically sharp (
<figureCitation id="5DE4899FFFB2FFC66768FB87FA33FBA5" box="[1300,1431,1122,1149]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Figs 5A, D</figureCitation>
;
<figureCitation id="5DE4899FFFB2FFC662EBFB63FF79FB79" box="[151,221,1159,1185]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">6AC</figureCitation>
;
<figureCitation id="5DE4899FFFB2FFC66294FB63FE8BFB79" box="[232,303,1159,1185]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="12" pageNumber="213">7AE</figureCitation>
). Embolus short, not describing a complete loop around bulb, tape-shaped at base, coiled, with poorly developed laminar projection; basal process small, flat, retrolateral region pointed in posterior region, prolateral region oval, poorly pronounced into external keel (
<figureCitation id="5DE4899FFFB2FFC660ACFB2BFCF4FB31" box="[720,848,1230,1257]" captionStart="FIGURE 5" captionStartId="8.[151,250,1859,1883]" captionTargetBox="[159,1428,192,1829]" captionTargetId="figure-22@8.[151,1436,181,1835]" captionTargetPageId="8" captionText="FIGURE 5. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245), left copulatory bulb: A Ventral view; B Retrolateral view; C Prolateral view; D Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: E Ventral view (arrow indicates the anterior excavations); F Dorsal view. Abbreviations: C, conductor; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct; St, subtegulum; T, tegulum; TL, tegular lobe." pageId="12" pageNumber="213">Figs 5AD</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC66120FB2BFC05FB31" box="[860,929,1231,1257]" captionStart="FIGURE 6" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[151,1409,185,1846]" captionTargetId="figure-17@9.[151,1435,181,1848]" captionTargetPageId="9" captionText="FIGURE 6. Josa chazaliae (Simon, 1897), male (MUSENUV-Ar-2671; vchLAM-245) left copulatory bulb: A Ventral view; B Retrolateral view; C Apical view. Female (MUSENUV-Ar-2672; vchLAM-247), genitalia: D Ventral view; E Dorsal view, internal morphology. Abbreviations: AB, accessory bulb; C, conductor; CO, copulatory opening; dAB, duct of accessory bulb; E, embolus; FD, fertilization duct; LL, lateral lobe; MA, median apophysis; MF, median field; PBE, process on base of embolus; PMA, paramedian apophysis; S, spermatheca; SD, spermatic duct." pageId="12" pageNumber="213">6AC</figureCitation>
,
<figureCitation id="5DE4899FFFB2FFC661D1FB2BFC19FB31" box="[941,957,1231,1257]" captionStart="FIGURE 7" captionStartId="10.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1756]" captionTargetId="figure-22@10.[151,1435,181,1756]" captionTargetPageId="10" captionText="FIGURE 7. Josa chazaliae (Simon, 1897), male (MACN-Ar 46690; vchLAM-246) left copulatory bulb, SEM images:A Ventral view; B Details of apical region of conductor and embolus (arrow indicates the apical bump); C Paramedian apophysis, ventral view; D Male bulb, retrolateral view; E Paramedian apophysis, retrolateral view; F Tegular lobe; G Male bulb, prolateral view; H Male bulb, apical view; I Apical view, details of PBE. Abbreviations: C, conductor; E, embolus; MA, median apophysis; PBE, process on base of embolus; PMA, paramedian apophysis; T, tegulum; TL, tegular lobe, St, subtegulum." pageId="12" pageNumber="213">7</figureCitation>
).
</paragraph>
<paragraph id="C560951AFFB2FFC662BBFB16FBF8FA8D" blockId="12.[151,1437,151,1401]" pageId="12" pageNumber="213">
<emphasis id="F7AB4908FFB2FFC662BBFB16FEE5FAD4" bold="true" box="[199,321,1266,1292]" pageId="12" pageNumber="213">Variation:</emphasis>
Males (n=2): total length: 4.284.54; carapace length: 2.182.34, width: 1.571.61; abdomen length: 2.142.34, width: 1.191.35. Females (n=3): total length: 7.408.19; carapace length: 3.523.70, width: 2.472.56; abdomen length: 3.754.42, width: 1.862.64. Fresh specimens look more pigmented.
</paragraph>
</subSubSection>
<subSubSection id="8DC5C691FFB2FFC662BBFABAFBFAFAA1" box="[199,1118,1374,1401]" pageId="12" pageNumber="213" type="distribution">
<paragraph id="C560951AFFB2FFC662BBFABAFBFAFAA1" blockId="12.[151,1437,151,1401]" box="[199,1118,1374,1401]" pageId="12" pageNumber="213">
<emphasis id="F7AB4908FFB2FFC662BBFABAFEC5FAA0" bold="true" box="[199,353,1374,1400]" pageId="12" pageNumber="213">Distribution.</emphasis>
Only known for Sierra Nevada de Santa Marta,
<collectingCountry id="BDC8D58AFFB2FFC66103FABBFC49FAA1" box="[895,1005,1375,1401]" name="Colombia" pageId="12" pageNumber="213">Colombia</collectingCountry>
(
<figureCitation id="5DE4899FFFB2FFC66181FABBFBF4FAA1" box="[1021,1104,1375,1401]" captionStart="FIGURE 31" captionStartId="39.[151,250,1138,1162]" captionTargetBox="[159,1428,189,1106]" captionTargetId="figure-17@39.[151,1436,181,1114]" captionTargetPageId="39" captionText="FIGURE 31. Distribution of Josa species in Sierra Nevada de Santa Marta." pageId="12" pageNumber="213">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,223 @@
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<mods:title id="175B1DA75067C0FEC1CFE93030D2B257">Two new species of Thlaspidula Spaeth, 1901 (Coleoptera: Chrysomelidae: Cassidinae) from Sulawesi</mods:title>
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<mods:namePart id="4B6F175432641070070DD89A21FCC94F">Budi, Anang Setyo</mods:namePart>
<mods:affiliation id="CD530756A91560F8EFBD2D999A106E57">Museum Zoologicum Bogoriense, Research Center for Biosystematics and Evolution, National Research and Innovation Agency, Cibinong, Bogor, Indonesia</mods:affiliation>
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<mods:namePart id="A27F626BD5D50E1C991AD5C472709CEF">Świętojańska, Jolanta</mods:namePart>
<mods:affiliation id="32BC61130644888C2DCED3086ACE4CE2">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland.</mods:affiliation>
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<subSubSection id="9FA1657BFFA5FFF47CC1FE58094190E2" box="[151,770,474,504]" pageId="3" pageNumber="394" type="nomenclature">
<paragraph id="D70436F0FFA5FFF47CC1FE58094190E2" blockId="3.[151,770,474,537]" box="[151,770,474,504]" pageId="3" pageNumber="394">
<heading id="8C4C819CFFA5FFF47CC1FE58094190E2" bold="true" box="[151,770,474,504]" fontSize="11" level="1" pageId="3" pageNumber="394" reason="1">
<emphasis id="E5CFEAE2FFA5FFF47CC1FE58094190E2" bold="true" box="[151,770,474,504]" pageId="3" pageNumber="394">
<taxonomicName id="10BB4D73FFA5FFF47CC1FE5808D490EF" authority="Budi &amp; Swietojanska" authorityName="Budi &amp; Swietojanska" authorityYear="2025" box="[151,663,474,504]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="sarinoi" status="sp. nov.">
<emphasis id="E5CFEAE2FFA5FFF47CC1FE580BC690E2" bold="true" box="[151,389,474,501]" italics="true" pageId="3" pageNumber="394">Thlaspidula sarinoi</emphasis>
Budi &amp; Świętojańska
</taxonomicName>
,
<taxonomicNameLabel id="FEFC5799FFA5FFF47EF2FE59094190E2" box="[676,770,475,501]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="9FA1657BFFA5FFF47CC1FE7D0B6D930E" box="[151,302,510,537]" pageId="3" pageNumber="394" type="description">
<paragraph id="D70436F0FFA5FFF47CC1FE7D0B6D930E" blockId="3.[151,770,474,537]" box="[151,302,510,537]" pageId="3" pageNumber="394">
(
<figureCitation id="4F802A75FFA5FFF47CC9FE7D0B65930E" box="[159,294,510,537]" captionStart-0="FIGURES 45" captionStart-1="FIGURES 68" captionStartId-0="2.[151,264,1927,1951]" captionStartId-1="4.[151,264,1392,1416]" captionTargetBox-0="[179,1420,1341,1882]" captionTargetBox-1="[151,1436,181,1368]" captionTargetId-0="figure-49@2.[151,1436,1301,1903]" captionTargetId-1="figure-31@4.[151,1436,181,1368]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 45. Tarsal claw. 4. Thlaspidula gandangdewata sp. nov.; 5. Thlaspidula sarinoi sp. nov." captionText-1="FIGURES 68. Thlaspidula sarinoi sp. nov. 6. dorsal view; 7. ventral view; 8. lateral view." pageId="3" pageNumber="394">Figures 58</figureCitation>
)
</paragraph>
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<subSubSection id="9FA1657BFFA5FFF47CC1FDC508AD93E6" pageId="3" pageNumber="394" type="materials_examined">
<paragraph id="D70436F0FFA5FFF47CC1FDC50B6B9392" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">
<materialsCitation id="67D33CADFFA5FFF47CC1FDC50B6B9392" country="Indonesia" county="Sigi" elevation="1390" location="Namo" longitude="1.42575" municipality="Kulawi" pageId="3" pageNumber="394" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<emphasis id="E5CFEAE2FFA5FFF47CC1FDC50B769377" bold="true" box="[151,309,582,609]" pageId="3" pageNumber="394">
<typeStatus id="08008852FFA5FFF47CC1FDC50A919376" box="[151,210,583,609]" pageId="3" pageNumber="394">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="AFAC7660FFA5FFF47D6AFDC50BEF9376" box="[316,428,583,609]" name="Indonesia" pageId="3" pageNumber="394">Indonesia</collectingCountry>
,
<collectingRegion id="157FF812FFA5FFF47DE1FDC5085C9376" box="[439,543,583,609]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="394">Sulawesi</collectingRegion>
,
<collectingRegion id="157FF812FFA5FFF47E7CFDC508A89376" box="[554,747,583,609]" country="Indonesia" name="Sulawesi Tengah" pageId="3" pageNumber="394">Central Sulawesi</collectingRegion>
,
<collectingCounty id="3E654E7CFFA5FFF47EA0FDC509659376" box="[758,806,583,609]" pageId="3" pageNumber="394">Sigi</collectingCounty>
,
<collectingMunicipality id="3760AC8AFFA5FFF47F64FDC509C49376" box="[818,903,583,609]" pageId="3" pageNumber="394">Kulawi</collectingMunicipality>
,
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, approx. S:
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:119°5383.8”,
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a.s.l.
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</paragraph>
<paragraph id="D70436F0FFA5FFF47C91FD0D08AD93E6" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">
<emphasis id="E5CFEAE2FFA5FFF47C91FD0D085693BF" bold="true" box="[199,533,654,681]" pageId="3" pageNumber="394">
Type material (
<figureCitation id="4F802A75FFA5FFF47D2CFD0C084B93BF" box="[378,520,654,680]" captionStart-0="FIGURES 45" captionStart-1="FIGURES 68" captionStartId-0="2.[151,264,1927,1951]" captionStartId-1="4.[151,264,1392,1416]" captionTargetBox-0="[179,1420,1341,1882]" captionTargetBox-1="[151,1436,181,1368]" captionTargetId-0="figure-49@2.[151,1436,1301,1903]" captionTargetId-1="figure-31@4.[151,1436,181,1368]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 45. Tarsal claw. 4. Thlaspidula gandangdewata sp. nov.; 5. Thlaspidula sarinoi sp. nov." captionText-1="FIGURES 68. Thlaspidula sarinoi sp. nov. 6. dorsal view; 7. ventral view; 8. lateral view." pageId="3" pageNumber="394">Figures 58</figureCitation>
).
</emphasis>
<typeStatus id="08008852FFA5FFF47E4AFD0D08C493BE" box="[540,647,655,681]" pageId="3" pageNumber="394" type="holotype">Holotype</typeStatus>
, pinned: male,
<collectingRegion id="157FF812FFA5FFF47F61FD0D09B093BE" box="[823,1011,655,681]" country="Indonesia" name="Sulawesi Tengah" pageId="3" pageNumber="394">Central Sulawesi</collectingRegion>
<collectingCountry id="AFAC7660FFA5FFF47846FD0D0E3D93BE" box="[1040,1150,655,681]" name="Indonesia" pageId="3" pageNumber="394">Indonesia</collectingCountry>
| Sigi—Kulawi | Namo—
<date id="A3051030FFA5FFF47CC1FD310B4C93DA" box="[151,271,691,717]" pageId="3" pageNumber="394" value="2017-11-24">24.xi.2017</date>
| Sarino ||
<typeStatus id="08008852FFA5FFF47DDDFD31085C93DA" box="[395,543,691,717]" pageId="3" pageNumber="394" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="10BB4D73FFA5FFF47E60FD3108FF93DA" authorityName="Spaeth" authorityYear="1901" box="[566,700,691,717]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="E5CFEAE2FFA5FFF47E60FD3108FF93DA" box="[566,700,691,717]" italics="true" pageId="3" pageNumber="394">Thlaspidula</emphasis>
</taxonomicName>
|
<taxonomicName id="10BB4D73FFA5FFF47E85FD31096193DB" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[723,802,691,716]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="sarinoi" status="sp. nov.">
<emphasis id="E5CFEAE2FFA5FFF47E85FD31096193DB" box="[723,802,691,716]" italics="true" pageId="3" pageNumber="394">sarinoi</emphasis>
</taxonomicName>
<emphasis id="E5CFEAE2FFA5FFF47F7DFD3109C593DA" bold="true" box="[811,902,691,717]" pageId="3" pageNumber="394">
<taxonomicNameLabel id="FEFC5799FFA5FFF47F7DFD3109C593DA" box="[811,902,691,717]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Budi &amp; Świętojańska 2024 (deposited at the Museum Zoologicum Bogoriense, Bogor,
<collectingCountry id="AFAC7660FFA5FFF47E26FD5508A193E6" box="[624,738,727,753]" name="Indonesia" pageId="3" pageNumber="394">Indonesia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA5FFF47C91FD780F1592DE" pageId="3" pageNumber="394" type="diagnosis">
<paragraph id="D70436F0FFA5FFF47C91FD780F1592DE" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">
<emphasis id="E5CFEAE2FFA5FFF47C91FD780B019203" bold="true" box="[199,322,762,788]" pageId="3" pageNumber="394">Diagnosis.</emphasis>
<taxonomicName id="10BB4D73FFA5FFF47D1DFD790BFB9203" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[331,440,763,788]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="sarinoi" status="sp. nov.">
<emphasis id="E5CFEAE2FFA5FFF47D1DFD790BFB9203" box="[331,440,763,788]" italics="true" pageId="3" pageNumber="394">T. sarinoi</emphasis>
</taxonomicName>
<emphasis id="E5CFEAE2FFA5FFF47D94FD79085D9202" bold="true" box="[450,542,763,789]" pageId="3" pageNumber="394">
<taxonomicNameLabel id="FEFC5799FFA5FFF47D94FD79085D9202" box="[450,542,763,789]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with
<taxonomicName id="10BB4D73FFA5FFF47E35FD79093D9202" authority="(Boheman)" baseAuthorityName="Boheman" box="[611,894,763,789]" class="Insecta" family="Chrysomelidae" genus="Aspidomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="boisduvali">
<emphasis id="E5CFEAE2FFA5FFF47E35FD7908B59202" box="[611,758,763,789]" italics="true" pageId="3" pageNumber="394">T. boisduvali</emphasis>
(Boheman)
</taxonomicName>
and
<taxonomicName id="10BB4D73FFA5FFF47FEFFD790EC89202" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[953,1163,763,789]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gandangdewata" status="sp. nov.">
<emphasis id="E5CFEAE2FFA5FFF47FEFFD790EC89202" box="[953,1163,763,789]" italics="true" pageId="3" pageNumber="394">T. gandangdewata</emphasis>
</taxonomicName>
<emphasis id="E5CFEAE2FFA5FFF478C2FD790EB39202" bold="true" box="[1172,1264,763,789]" pageId="3" pageNumber="394">
<taxonomicNameLabel id="FEFC5799FFA5FFF478C2FD790EB39202" box="[1172,1264,763,789]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the group of species with presence of only posterolateral spots on the explanate margin of the elytra, without humeral spots. It differs from both congeners in elytral pattern with humeral spots merged with posterolateral band (separated in congeners) and scutellum with broadly brown to black margins (uniformly yellow in congeners).
<taxonomicName id="10BB4D73FFA5FFF478B2FCE50F0D9297" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[1252,1358,871,896]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="sarinoi">
<emphasis id="E5CFEAE2FFA5FFF478B2FCE50F0D9297" box="[1252,1358,871,896]" italics="true" pageId="3" pageNumber="394">T. sarinoi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="10BB4D73FFA5FFF47C81FC090BE492B2" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[215,423,907,933]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gandangdewata">
<emphasis id="E5CFEAE2FFA5FFF47C81FC090BE492B2" box="[215,423,907,933]" italics="true" pageId="3" pageNumber="394">T. gandangdewata</emphasis>
</taxonomicName>
also in distinctly micropectinate claws and elongate mandibles.
<taxonomicName id="10BB4D73FFA5FFF478DDFC090EB492B3" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[1163,1271,907,932]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="sarinoi">
<emphasis id="E5CFEAE2FFA5FFF478DDFC090EB492B3" box="[1163,1271,907,932]" italics="true" pageId="3" pageNumber="394">T. sarinoi</emphasis>
</taxonomicName>
is larger from both congeners with pronotum width above
<quantity id="10439B15FFA5FFF47ED0FC2C089F92DE" box="[646,732,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="394" unit="mm" value="5.5">5.5 mm</quantity>
(below
<quantity id="10439B15FFA5FFF47F61FC2C09CE92DE" box="[823,909,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="394" unit="mm" value="5.4">5.4 mm</quantity>
in
<taxonomicName id="10BB4D73FFA5FFF47FE6FC2D0E0292DE" box="[944,1089,943,969]" class="Insecta" family="Chrysomelidae" genus="Aspidomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="boisduvali">
<emphasis id="E5CFEAE2FFA5FFF47FE6FC2D0E0292DE" box="[944,1089,943,969]" italics="true" pageId="3" pageNumber="394">T. boisduvali</emphasis>
</taxonomicName>
and
<taxonomicName id="10BB4D73FFA5FFF47821FC2D0F0692DE" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[1143,1349,943,969]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gandangdewata">
<emphasis id="E5CFEAE2FFA5FFF47821FC2D0F0692DE" box="[1143,1349,943,969]" italics="true" pageId="3" pageNumber="394">T. gandangdewata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA5FFF47C91FC5009DE9666" pageId="3" pageNumber="394" type="description">
<paragraph id="D70436F0FFA5FFF47C91FC5009109506" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">
<emphasis id="E5CFEAE2FFA5FFF47C91FC500B1A92FB" bold="true" box="[199,345,978,1004]" pageId="3" pageNumber="394">Description.</emphasis>
Length:
<quantity id="10439B15FFA5FFF47DEDFC51085192FA" box="[443,530,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="3" pageNumber="394" unit="mm" value="7.9">7.9 mm</quantity>
, width:
<quantity id="10439B15FFA5FFF47E3CFC51088292FA" box="[618,705,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="394" unit="mm" value="7.3">7.3 mm</quantity>
, length of pronotum:
<quantity id="10439B15FFA5FFF47FF9FC510E4692FA" box="[943,1029,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="394" unit="mm" value="2.7">2.7 mm</quantity>
, width of pronotum:
<quantity id="10439B15FFA5FFF478BBFC500F0792FA" box="[1261,1348,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="394" unit="mm" value="5.6">5.6 mm</quantity>
, length/ width ratio: 1.08, width/length of pronotum ratio: 2.07 (n = 2).
</paragraph>
<paragraph id="D70436F0FFA5FFF47C91FB990960941A" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">Pronotum yellow, disc at base with two small dark brown to black triangular spots, basal margin of disc darkened. Scutellum brown to black with yellow spot in the middle at the anterior margin. Elytral disc dark brown with yellow spots: two large spots in anterior half of disc, two small spots behind postscutellar elevation not connected with large anterior spots, two small spots in the middle connected with yellow lateral margins, two large posterior spots extending to the apical margin. Punctures located in area of yellow spots black. Explanate margin of elytra mostly yellow, with broad dark brown to black posterolateral spots, humeral spots absent. Clypeus, ventrites and legs yellow. Antennae yellow, segment 511 slightly infuscate.</paragraph>
<paragraph id="D70436F0FFA5FFF47C91FA950970948A" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">Pronotum broad, widest slightly behind middle, disc regularly convex, distinctly separated from explanate margin, each side is separated from the explanate margin by two distinct borders of impressions, one smaller in the area above head, one larger close to base. Explanate margin very broad, almost horizontal. Elytral disc and explanate margin surface smooth and glabrous, impunctate.</paragraph>
<paragraph id="D70436F0FFA5FFF47C91FA250B8497AA" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">Scutellum triangular. Base of elytra distinctly wider than pronotum, humeral angles strongly protruding anterad, angulate. Elytral disc strongly convex, distinctly elevated in postscutellar area without forming tubercles. Punctation of the disc is regular only in the dark part, punctures are broken in the anterior relief and sparser in the posterior relief. Punctures in rows quite dense, distance between punctures as wide as diameter. Intervals slightly convex, as wide as, to twice wider than rows, second interval thrice as wide as rows. Marginal row distinct, its punctures larger than in central rows, space between punctures slightly elevated, forming short transverse folds. Explanate margin very broad, widest part slightly narrower than width of each elytron, almost horizontal. Whole surface of disc and explanate margin glabrous.</paragraph>
<paragraph id="D70436F0FFA5FFF47C91F94509DE9666" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">
Clypeus narrow, almost as long as wide, shallowly impressed along middle, smooth and glabrous. Labrum broad, deeply impressed on sides. Mandibles elongated. Prosternal process broad, strongly expanded apically, shallowly impressed in middle, bottom of impression indistinctly granulate. Antennae in male slim and long, third segment almost twice as long as second, fourth segment 1.3x as long as third, 10
<superScript id="20CE9BB8FFA5FFF47844F8B00E5C9657" attach="left" box="[1042,1055,1842,1856]" fontSize="6" pageId="3" pageNumber="394">th</superScript>
segment 3.7x as long as wide, last segment 9.2x as long as wide. Claws large, distinctly micropectinate.
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA5FFF47C91F8F808189682" box="[199,603,1914,1941]" pageId="3" pageNumber="394" type="distribution">
<paragraph id="D70436F0FFA5FFF47C91F8F808189682" blockId="3.[151,1437,582,2013]" box="[199,603,1914,1941]" pageId="3" pageNumber="394">
<emphasis id="E5CFEAE2FFA5FFF47C91F8F80B229683" bold="true" box="[199,353,1914,1940]" pageId="3" pageNumber="394">Distribution.</emphasis>
<collectingCountry id="AFAC7660FFA5FFF47D31F8F90B979682" box="[359,468,1915,1941]" name="Indonesia" pageId="3" pageNumber="394">Indonesia</collectingCountry>
(
<collectingRegion id="157FF812FFA5FFF47DB5F8F9080D9682" box="[483,590,1915,1941]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="394">Sulawesi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA5FFF47C91F81D08F196CA" pageId="3" pageNumber="394" type="etymology">
<paragraph id="D70436F0FFA5FFF47C91F81D08F196CA" blockId="3.[151,1437,582,2013]" pageId="3" pageNumber="394">
<emphasis id="E5CFEAE2FFA5FFF47C91F81D0B0D96AE" bold="true" box="[199,334,1951,1977]" pageId="3" pageNumber="394">Etymology.</emphasis>
The epithet
<taxonomicName id="10BB4D73FFA5FFF47D8AF81D087B96AF" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[476,568,1951,1977]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="sarinoi">
<emphasis id="E5CFEAE2FFA5FFF47DBFF81D087B96AF" box="[489,568,1951,1976]" italics="true" pageId="3" pageNumber="394">sarinoi</emphasis>
</taxonomicName>
” is taken from the name Sarino, field collector of the
<typeStatus id="08008852FFA5FFF478CAF81D0EBD96AE" box="[1180,1278,1951,1977]" pageId="3" pageNumber="394" type="holotype">holotype</typeStatus>
and also staff at the Museum Zoologicum Bogoriense (MZB).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,263 @@
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<mods:title id="2F1C115D97C0EF2281D2A2A9363B59F7">Two new species of Thlaspidula Spaeth, 1901 (Coleoptera: Chrysomelidae: Cassidinae) from Sulawesi</mods:title>
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<mods:namePart id="1CABE5E31562DCF378A63A814CFB0BD6">Budi, Anang Setyo</mods:namePart>
<mods:affiliation id="7F67982F94FAB4113108EE198F66286C">Museum Zoologicum Bogoriense, Research Center for Biosystematics and Evolution, National Research and Innovation Agency, Cibinong, Bogor, Indonesia</mods:affiliation>
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<mods:namePart id="2881E1FF2901DD53B8FB1A3B4927A987">Świętojańska, Jolanta</mods:namePart>
<mods:affiliation id="CE9BD3202235DC5CFDF2378C74FEFA62">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland.</mods:affiliation>
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<treatment id="5F1287E6FFA7FFF47CC1FD0D0F3C909E" LSID="urn:lsid:plazi:treatment:5F1287E6FFA7FFF47CC1FD0D0F3C909E" httpUri="http://treatment.plazi.org/id/5F1287E6FFA7FFF47CC1FD0D0F3C909E" lastPageId="3" lastPageNumber="394" pageId="1" pageNumber="392">
<subSubSection id="9FA1657BFFA7FFF67CC1FD0D092F93BE" box="[151,876,655,685]" pageId="1" pageNumber="392" type="nomenclature">
<paragraph id="D70436F0FFA7FFF67CC1FD0D092F93BE" blockId="1.[151,876,655,717]" box="[151,876,655,685]" pageId="1" pageNumber="392">
<heading id="8C4C819CFFA7FFF67CC1FD0D092F93BE" bold="true" box="[151,876,655,685]" fontSize="11" level="1" pageId="1" pageNumber="392" reason="1">
<emphasis id="E5CFEAE2FFA7FFF67CC1FD0D092F93BE" bold="true" box="[151,876,655,685]" pageId="1" pageNumber="392">
<taxonomicName id="10BB4D73FFA7FFF67CC1FD0D094293BA" authority="Budi &amp; Swietojanska" authorityName="Budi &amp; Swietojanska" authorityYear="2025" box="[151,769,655,685]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="gandangdewata" status="sp. nov.">
<emphasis id="E5CFEAE2FFA7FFF67CC1FD0D0BAC93BD" bold="true" box="[151,495,655,682]" italics="true" pageId="1" pageNumber="392">Thlaspidula gandangdewata</emphasis>
Budi &amp; Świętojańska
</taxonomicName>
,
<taxonomicNameLabel id="FEFC5799FFA7FFF67F58FD0D092F93BE" box="[782,876,655,681]" pageId="1" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA7FFF67CC1FD310B6C93DA" box="[151,303,691,717]" pageId="1" pageNumber="392" type="description">
<paragraph id="D70436F0FFA7FFF67CC1FD310B6C93DA" blockId="1.[151,876,655,717]" box="[151,303,691,717]" pageId="1" pageNumber="392">
(
<figureCitation id="4F802A75FFA7FFF67CC9FD310B6593DA" box="[159,294,691,717]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 45" captionStartId-0="2.[151,264,1217,1241]" captionStartId-1="2.[151,264,1927,1951]" captionTargetBox-0="[151,1436,181,1193]" captionTargetBox-1="[179,1420,1341,1882]" captionTargetId-0="figure-31@2.[151,1436,181,1193]" captionTargetId-1="figure-49@2.[151,1436,1301,1903]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURES 13. Thlaspidula gandangdewata sp. nov. 1. dorsal view; 2. ventral view; 3. lateral view." captionText-1="FIGURES 45. Tarsal claw. 4. Thlaspidula gandangdewata sp. nov.; 5. Thlaspidula sarinoi sp. nov." pageId="1" pageNumber="392">Figures 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA7FFF67CC1FD790F7392B2" pageId="1" pageNumber="392" type="materials_examined">
<paragraph id="D70436F0FFA7FFF67CC1FD790815922E" blockId="1.[151,1437,763,1978]" pageId="1" pageNumber="392">
<materialsCitation id="67D33CADFFA7FFF67CC1FD790815922E" country="Indonesia" county="Mamasa" elevation="1687" location="Gandang Dewata National Park" longitude="5.0354195" municipality="Rante Pongko" pageId="1" pageNumber="392" specimenCount="1" stateProvince="Sulawesi Barat" typeStatus="holotype">
<emphasis id="E5CFEAE2FFA7FFF67CC1FD790B759202" bold="true" box="[151,310,763,789]" pageId="1" pageNumber="392">
<typeStatus id="08008852FFA7FFF67CC1FD790A919202" box="[151,210,763,789]" pageId="1" pageNumber="392">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="AFAC7660FFA7FFF67D6BFD790BEE9202" box="[317,429,763,789]" name="Indonesia" pageId="1" pageNumber="392">Indonesia</collectingCountry>
,
<collectingRegion id="157FF812FFA7FFF67DE1FD7E081D9202" box="[439,606,763,790]" country="Indonesia" name="Sulawesi Barat" pageId="1" pageNumber="392">West Sulawesi</collectingRegion>
,
<collectingCounty id="3E654E7CFFA7FFF67E3FFD7E08889201" box="[617,715,764,790]" pageId="1" pageNumber="392">Mamasa</collectingCounty>
,
<collectingMunicipality id="3760AC8AFFA7FFF67E80FD7E09359202" box="[726,886,763,790]" pageId="1" pageNumber="392">Rante Pongko</collectingMunicipality>
,
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, approx. S:02°
<geoCoordinate id="B28F5037FFA7FFF679D8FD790B4E922D" degrees="5" direction="east" minutes="2" orientation="longitude" pageId="1" pageNumber="392" precision="1" seconds="07,51" value="5.0354195">5 207,51”E</geoCoordinate>
:119°2257,03”,
<elevation id="5C96D1C3FFA7FFF67D90FC9D0815922E" box="[454,598,799,826]" metricMagnitude="3" metricUnit="m" metricValue="1.6869999999999998" pageId="1" pageNumber="392" unit="m" value="1687.0">
<quantity id="10439B15FFA7FFF67D90FC9D0858922D" box="[454,539,799,826]" metricMagnitude="3" metricUnit="m" metricValue="1.6869999999999998" pageId="1" pageNumber="392" unit="m" value="1687.0">1687 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
<paragraph id="D70436F0FFA7FFF67C91FCC10F7392B2" blockId="1.[151,1437,763,1978]" pageId="1" pageNumber="392">
<emphasis id="E5CFEAE2FFA7FFF67C91FCC10858924A" bold="true" box="[199,539,835,861]" pageId="1" pageNumber="392">
Type material (
<figureCitation id="4F802A75FFA7FFF67D2BFCC1084F924A" box="[381,524,835,861]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 45" captionStartId-0="2.[151,264,1217,1241]" captionStartId-1="2.[151,264,1927,1951]" captionTargetBox-0="[151,1436,181,1193]" captionTargetBox-1="[179,1420,1341,1882]" captionTargetId-0="figure-31@2.[151,1436,181,1193]" captionTargetId-1="figure-49@2.[151,1436,1301,1903]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURES 13. Thlaspidula gandangdewata sp. nov. 1. dorsal view; 2. ventral view; 3. lateral view." captionText-1="FIGURES 45. Tarsal claw. 4. Thlaspidula gandangdewata sp. nov.; 5. Thlaspidula sarinoi sp. nov." pageId="1" pageNumber="392">Figures 14</figureCitation>
).
</emphasis>
<typeStatus id="08008852FFA7FFF67E74FCC108CE924A" box="[546,653,835,861]" pageId="1" pageNumber="392" type="holotype">Holotype</typeStatus>
, pinned: male,
<collectingRegion id="157FF812FFA7FFF67F6BFCC109A7924A" box="[829,996,835,861]" country="Indonesia" name="Sulawesi Barat" pageId="1" pageNumber="392">West Sulawesi</collectingRegion>
<collectingCountry id="AFAC7660FFA7FFF67857FCC10E2C924A" box="[1025,1135,835,861]" name="Indonesia" pageId="1" pageNumber="392">Indonesia</collectingCountry>
| Mamasa—Rante Pongko | Gandang Dewata N.P.—
<date id="A3051030FFA7FFF67DE1FCE5086E9296" box="[439,557,871,897]" pageId="1" pageNumber="392" value="2022-06-24">24.vi.2022</date>
|
<collectionCode id="B1AAAE35FFA7FFF67E17FCEA08149295" box="[577,599,872,898]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="392">R</collectionCode>
. Pramesa N.K. &amp; Sarino ||
<typeStatus id="08008852FFA7FFF67FD4FCE50E559296" box="[898,1046,871,897]" pageId="1" pageNumber="392" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="10BB4D73FFA7FFF6787EFCE50EED9296" authorityName="Spaeth" authorityYear="1901" box="[1064,1198,871,897]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="E5CFEAE2FFA7FFF6787EFCE50EED9296" box="[1064,1198,871,897]" italics="true" pageId="1" pageNumber="392">Thlaspidula</emphasis>
</taxonomicName>
|
<taxonomicName id="10BB4D73FFA7FFF67897FCE50F379296" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[1217,1396,871,897]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="gandangdewata" status="sp. nov.">
<emphasis id="E5CFEAE2FFA7FFF67897FCE50F379296" box="[1217,1396,871,897]" italics="true" pageId="1" pageNumber="392">gandangdewata</emphasis>
</taxonomicName>
<emphasis id="E5CFEAE2FFA7FFF6792DFCE50A8B92B2" bold="true" pageId="1" pageNumber="392">
<taxonomicNameLabel id="FEFC5799FFA7FFF6792DFCE50A8B92B2" pageId="1" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
| Budi &amp; Świętojańska 2024 (deposited at the Museum Zoologicum Bogoriense, Bogor,
<collectingCountry id="AFAC7660FFA7FFF678E4FC090F6792B2" box="[1202,1316,907,933]" name="Indonesia" pageId="1" pageNumber="392">Indonesia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA7FFF67C91FC2D0F2E95FE" pageId="1" pageNumber="392" type="diagnosis">
<paragraph id="D70436F0FFA7FFF67C91FC2D0F2E95FE" blockId="1.[151,1437,763,1978]" pageId="1" pageNumber="392">
<emphasis id="E5CFEAE2FFA7FFF67C91FC2D0B0192DE" bold="true" box="[199,322,943,969]" pageId="1" pageNumber="392">Diagnosis.</emphasis>
<taxonomicName id="10BB4D73FFA7FFF67D1CFC32085A92DE" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[330,537,943,969]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="gandangdewata">
<emphasis id="E5CFEAE2FFA7FFF67D1CFC32085A92DE" box="[330,537,943,969]" italics="true" pageId="1" pageNumber="392">T. gandangdewata</emphasis>
</taxonomicName>
with
<taxonomicName id="10BB4D73FFA7FFF67E0AFC32093792DE" authority="(Boheman)" baseAuthorityName="Boheman" box="[604,884,943,969]" class="Insecta" family="Chrysomelidae" genus="Aspidomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="boisduvali">
<emphasis id="E5CFEAE2FFA7FFF67E0AFC3208AE92DE" box="[604,749,943,969]" italics="true" pageId="1" pageNumber="392">T. boisduvali</emphasis>
(Boheman)
</taxonomicName>
and
<taxonomicName id="10BB4D73FFA7FFF67FFAFC320E5B92DE" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[940,1048,944,969]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="sarinoi" status="sp. nov.">
<emphasis id="E5CFEAE2FFA7FFF67FFAFC320E5B92DE" box="[940,1048,944,969]" italics="true" pageId="1" pageNumber="392">T. sarinoi</emphasis>
</taxonomicName>
<emphasis id="E5CFEAE2FFA7FFF67877FC2D0E3F92DE" bold="true" box="[1057,1148,943,969]" pageId="1" pageNumber="392">
<taxonomicNameLabel id="FEFC5799FFA7FFF67877FC2D0E3F92DE" box="[1057,1148,943,969]" pageId="1" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
forms a group of species with presence of only posterolateral spots on the explanate margin of the elytra, without humeral spots.At first glance
<taxonomicName id="10BB4D73FFA7FFF67CC1FC7A0B259506" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[151,358,1015,1041]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="gandangdewata">
<emphasis id="E5CFEAE2FFA7FFF67CC1FC7A0B259506" box="[151,358,1015,1041]" italics="true" pageId="1" pageNumber="392">T. gandangdewata</emphasis>
</taxonomicName>
seems to be very similar to
<taxonomicName id="10BB4D73FFA7FFF67EF4FC7A09719506" box="[674,818,1015,1041]" class="Insecta" family="Chrysomelidae" genus="Aspidomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="boisduvali">
<emphasis id="E5CFEAE2FFA7FFF67EF4FC7A09719506" box="[674,818,1015,1041]" italics="true" pageId="1" pageNumber="392">T. boisduvali</emphasis>
</taxonomicName>
especially in shape of dorsal dark pattern but differs in narrowly rounded basal spots on the pronotum (subtriangular in
<taxonomicName id="10BB4D73FFA7FFF67F26FB9E0E429522" box="[880,1025,1051,1077]" class="Insecta" family="Chrysomelidae" genus="Aspidomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="boisduvali">
<emphasis id="E5CFEAE2FFA7FFF67F26FB9E0E429522" box="[880,1025,1051,1077]" italics="true" pageId="1" pageNumber="392">T. boisduvali</emphasis>
</taxonomicName>
), broad axe-shaped spot on humeral calli of elytral disc (arched spot in
<taxonomicName id="10BB4D73FFA7FFF67E78FBC20882954E" box="[558,705,1087,1113]" class="Insecta" family="Chrysomelidae" genus="Aspidomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="boisduvali">
<emphasis id="E5CFEAE2FFA7FFF67E78FBC20882954E" box="[558,705,1087,1113]" italics="true" pageId="1" pageNumber="392">T. boisduvali</emphasis>
</taxonomicName>
), stout mandibles (elongate in
<taxonomicName id="10BB4D73FFA7FFF6787EFBC20EF8954E" box="[1064,1211,1087,1113]" class="Insecta" family="Chrysomelidae" genus="Aspidomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="boisduvali">
<emphasis id="E5CFEAE2FFA7FFF6787EFBC20EF8954E" box="[1064,1211,1087,1113]" italics="true" pageId="1" pageNumber="392">T. boisduvali</emphasis>
</taxonomicName>
) and in indistinctly micropunctate tarsal claws (distinctly micropectinate in
<taxonomicName id="10BB4D73FFA7FFF67F75FBE609F4956A" box="[803,951,1123,1149]" class="Insecta" family="Chrysomelidae" genus="Aspidomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="boisduvali">
<emphasis id="E5CFEAE2FFA7FFF67F75FBE609F4956A" box="[803,951,1123,1149]" italics="true" pageId="1" pageNumber="392">T. boisduvali</emphasis>
</taxonomicName>
).
<taxonomicName id="10BB4D73FFA7FFF67F84FBE60E03956A" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[978,1088,1124,1149]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="sarinoi">
<emphasis id="E5CFEAE2FFA7FFF67F84FBE60E03956A" box="[978,1088,1124,1149]" italics="true" pageId="1" pageNumber="392">T. sarinoi</emphasis>
</taxonomicName>
differs in elytral pattern with humeral spots connected with posterior band thus lateral pattern looks slightly reticulate, scutellum with broadly brown to black margins (completely yellow in
<taxonomicName id="10BB4D73FFA7FFF67EF8FB2E093D95D2" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[686,894,1195,1221]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="gandangdewata">
<emphasis id="E5CFEAE2FFA7FFF67EF8FB2E093D95D2" box="[686,894,1195,1221]" italics="true" pageId="1" pageNumber="392">T. gandangdewata</emphasis>
</taxonomicName>
) and in almost uniformly yellow antennae with segments 511 only slightly infuscate (bicolored with segments 79 mostly dark brown in
<taxonomicName id="10BB4D73FFA7FFF678D9FB520F1E95FE" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[1167,1373,1231,1257]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="gandangdewata">
<emphasis id="E5CFEAE2FFA7FFF678D9FB520F1E95FE" box="[1167,1373,1231,1257]" italics="true" pageId="1" pageNumber="392">T. gandangdewata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA7FFF47C91FB7108BB9056" lastPageId="3" lastPageNumber="394" pageId="1" pageNumber="392" type="description">
<paragraph id="D70436F0FFA7FFF67C91FB7109109426" blockId="1.[151,1437,763,1978]" pageId="1" pageNumber="392">
<emphasis id="E5CFEAE2FFA7FFF67C91FB710B1A941A" bold="true" box="[199,345,1267,1293]" pageId="1" pageNumber="392">Description.</emphasis>
Length:
<quantity id="10439B15FFA7FFF67DEDFB7108519419" box="[443,530,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="392" unit="mm" value="7.9">7.9 mm</quantity>
, width:
<quantity id="10439B15FFA7FFF67E3CFB7108829419" box="[618,705,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="392" unit="mm" value="7.4">7.4 mm</quantity>
, length of pronotum:
<quantity id="10439B15FFA7FFF67FF9FB710E469419" box="[943,1029,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="392" unit="mm" value="2.7">2.7 mm</quantity>
, width of pronotum:
<quantity id="10439B15FFA7FFF678BBFB710F079419" box="[1261,1348,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="392" unit="mm" value="5.3">5.3 mm</quantity>
, length/ width ratio: 1.05, width/length of pronotum ratio: 1.96 (n = 1).
</paragraph>
<paragraph id="D70436F0FFA7FFF67C91FABE0B479746" blockId="1.[151,1437,763,1978]" pageId="1" pageNumber="392">Pronotum yellow, disc with two very small, dark brown to black, obtuse rounded triangular spots, separated by yellow at the middle of basal margin. Scutellum yellow with dark brown margins. Elytral disc yellow, with large crescent-shaped dark brown to black spot on humeral calli, triangularly protruding at mid-posterior margin, and a large square spot behind middle between suture and fifth elytral puncture row, the spot connected by a band with posterolateral spot of the explanate margin. Explanate margin of elytra mostly yellow, with broad dark brown to black posterolateral spots, humeral spots absent. Clypeus, ventrites, legs yellow. Antennae bicolored, segments 15 yellow, segment 6 yellow basally infuscate apically, segments 79 mostly dark brown and segments 1011 slightly infuscate.</paragraph>
<paragraph id="D70436F0FFA7FFF67C91F9DE0F0297AA" blockId="1.[151,1437,763,1978]" pageId="1" pageNumber="392">Pronotum broad, widest in middle, disc regularly convex, each side is separated from the explanate margin by two borders of impressions, one smaller in the area above head, one larger close to base. Explanate margin very broad, almost horizontal. Elytral disc and explanate margin with surface smooth and glabrous, impunctate.</paragraph>
<paragraph id="D70436F0FFA7FFF67C91F9450E1696AE" blockId="1.[151,1437,763,1978]" pageId="1" pageNumber="392">Scutellum triangular. Base of elytra slightly wider than pronotum, humeral angles strongly protruding anterad, angulate. Elytral disc convex, distinctly elevated in postscutellar area. Punctation of disc regular, but broken by elytral relief. Punctures in rows quite dense, distance between punctures mostly as wide as puncture diameter. Intervals slightly convex, as wide as, to twice wider than rows, second interval thrice as wide as rows. Marginal row distinct, its punctures slightly larger than in central rows, space between punctures slightly elevated, forming short transverse folds. Explanate margin very broad, widest part slightly narrower than width of each elytron, lateral borders forms a shallow gutter. Whole surface of disc and explanate margin glabrous.</paragraph>
<caption id="83C46678FFA4FFF57CC1FB430ED595CE" box="[151,1174,1217,1241]" pageId="2" pageNumber="393" startId="2.[151,264,1217,1241]" targetBox="[151,1436,181,1193]" targetPageId="2" targetType="figure">
<paragraph id="D70436F0FFA4FFF57CC1FB430ED595CE" blockId="2.[151,1174,1217,1241]" box="[151,1174,1217,1241]" pageId="2" pageNumber="393">
<emphasis id="E5CFEAE2FFA4FFF57CC1FB430B7995CE" bold="true" box="[151,314,1217,1241]" pageId="2" pageNumber="393">FIGURES 13.</emphasis>
<taxonomicName id="10BB4D73FFA4FFF57D17FB43082195CE" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[321,610,1217,1241]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="gandangdewata" status="sp. nov.">
<emphasis id="E5CFEAE2FFA4FFF57D17FB43082195CE" box="[321,610,1217,1241]" italics="true" pageId="2" pageNumber="393">Thlaspidula gandangdewata</emphasis>
</taxonomicName>
<emphasis id="E5CFEAE2FFA4FFF57E3EFB43089095CE" bold="true" box="[616,723,1217,1241]" pageId="2" pageNumber="393">
<taxonomicNameLabel id="FEFC5799FFA4FFF57E3EFB4308FA95CE" box="[616,697,1217,1241]" pageId="2" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
1.
</emphasis>
dorsal view;
<emphasis id="E5CFEAE2FFA4FFF57F0AFB43092C95CE" bold="true" box="[860,879,1217,1241]" pageId="2" pageNumber="393">2.</emphasis>
ventral view;
<emphasis id="E5CFEAE2FFA4FFF57856FB430E5095CE" bold="true" box="[1024,1043,1217,1241]" pageId="2" pageNumber="393">3.</emphasis>
lateral view.
</paragraph>
</caption>
<caption id="83C46678FFA4FFF57CC1F8050ED59688" box="[151,1174,1927,1951]" pageId="2" pageNumber="393" startId="2.[151,264,1927,1951]" targetBox="[179,1420,1341,1882]" targetPageId="2" targetType="figure">
<paragraph id="D70436F0FFA4FFF57CC1F8050ED59688" blockId="2.[151,1174,1927,1951]" box="[151,1174,1927,1951]" pageId="2" pageNumber="393">
<emphasis id="E5CFEAE2FFA4FFF57CC1F8050B789688" bold="true" box="[151,315,1927,1951]" pageId="2" pageNumber="393">FIGURES 45.</emphasis>
Tarsal claw.
<emphasis id="E5CFEAE2FFA4FFF57DE9F8050B919688" bold="true" box="[447,466,1927,1951]" pageId="2" pageNumber="393">4.</emphasis>
<taxonomicName id="10BB4D73FFA4FFF57D8EF80508B99688" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[472,762,1927,1951]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="gandangdewata" status="sp. nov.">
<emphasis id="E5CFEAE2FFA4FFF57D8EF80508B99688" box="[472,762,1927,1951]" italics="true" pageId="2" pageNumber="393">Thlaspidula gandangdewata</emphasis>
</taxonomicName>
<emphasis id="E5CFEAE2FFA4FFF57F56F80509139688" bold="true" box="[768,848,1927,1951]" pageId="2" pageNumber="393">
<taxonomicNameLabel id="FEFC5799FFA4FFF57F56F80509139688" box="[768,848,1927,1951]" pageId="2" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="E5CFEAE2FFA4FFF57F08F80509329688" bold="true" box="[862,881,1927,1951]" pageId="2" pageNumber="393">5.</emphasis>
<taxonomicName id="10BB4D73FFA4FFF57F2EF8050E7C9688" authorityName="Budi &amp; Świętojańska" authorityYear="2025" box="[888,1087,1927,1951]" class="Insecta" family="Chrysomelidae" genus="Thlaspidula" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="sarinoi" status="sp. nov.">
<emphasis id="E5CFEAE2FFA4FFF57F2EF8050E7C9688" box="[888,1087,1927,1951]" italics="true" pageId="2" pageNumber="393">Thlaspidula sarinoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEFC5799FFA4FFF57813F8050ED59688" box="[1093,1174,1927,1951]" pageId="2" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="D70436F0FFA5FFF47C91FF1508BB9056" blockId="3.[151,1436,151,393]" pageId="3" pageNumber="394">
Clypeus narrow, almost as long as wide, shallowly impressed along middle, smooth and glabrous. Labrum very broad, deeply impressed on sides. Mandibles stout. Prosternal process broad, strongly expanded apically, shallowly impressed in middle, bottom of impression indistinctly granulate. Antennae in male slim and long, third segment twice as long as second, fourth segment 1.3x as long as third, 10
<superScript id="20CE9BB8FFA5FFF47F31FE8009379007" attach="left" box="[871,884,258,272]" fontSize="6" pageId="3" pageNumber="394">th</superScript>
segment 4x as long as wide, last segment 9.4x as long as wide. Claws large, indistinctly micropectinate.
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA5FFF47C91FEC808189072" box="[199,603,330,357]" pageId="3" pageNumber="394" type="distribution">
<paragraph id="D70436F0FFA5FFF47C91FEC808189072" blockId="3.[151,1436,151,393]" box="[199,603,330,357]" pageId="3" pageNumber="394">
<emphasis id="E5CFEAE2FFA5FFF47C91FEC80B229073" bold="true" box="[199,353,330,356]" pageId="3" pageNumber="394">Distribution.</emphasis>
<collectingCountry id="AFAC7660FFA5FFF47D31FEC90B979072" box="[359,468,331,357]" name="Indonesia" pageId="3" pageNumber="394">Indonesia</collectingCountry>
(
<collectingRegion id="157FF812FFA5FFF47DB5FEC9080D9072" box="[483,590,331,357]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="394">Sulawesi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="9FA1657BFFA5FFF47C91FEED0F3C909E" box="[199,1407,367,393]" pageId="3" pageNumber="394" type="etymology">
<paragraph id="D70436F0FFA5FFF47C91FEED0F3C909E" blockId="3.[151,1436,151,393]" box="[199,1407,367,393]" pageId="3" pageNumber="394">
<emphasis id="E5CFEAE2FFA5FFF47C91FEED0B0D909E" bold="true" box="[199,334,367,393]" pageId="3" pageNumber="394">Etymology.</emphasis>
It is named after Mount Gandang Dewata in
<collectingRegion id="157FF812FFA5FFF47F1FFEED09AC909E" box="[841,1007,367,393]" country="Indonesia" name="Sulawesi Barat" pageId="3" pageNumber="394">West Sulawesi</collectingRegion>
, where the specimen was collected.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,296 @@
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<mods:title id="2C6E0C3EA586636CE06CC1CB3BC06EB6">Identification key to the species of genus Thrips (Thysanoptera, Thripidae) from India with three new species and one new record</mods:title>
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<treatment id="623FC9262A03CA2AFF553012FE9EFE65" LSID="urn:lsid:plazi:treatment:623FC9262A03CA2AFF553012FE9EFE65" httpUri="http://treatment.plazi.org/id/623FC9262A03CA2AFF553012FE9EFE65" lastPageId="22" lastPageNumber="295" pageId="19" pageNumber="292">
<subSubSection id="A28C2BBB2A03CA2FFF553012FE03F9CF" box="[151,429,1605,1632]" pageId="19" pageNumber="292" type="nomenclature">
<paragraph id="EA2978302A03CA2FFF553012FE03F9CF" blockId="19.[151,429,1605,1668]" box="[151,429,1605,1632]" pageId="19" pageNumber="292">
<heading id="B161CF5C2A03CA2FFF553012FE03F9CF" bold="true" box="[151,429,1605,1632]" fontSize="11" level="1" pageId="19" pageNumber="292" reason="1">
<emphasis id="D8E2A4222A03CA2FFF553012FE03F9CF" bold="true" box="[151,429,1605,1632]" pageId="19" pageNumber="292">
<taxonomicName id="2D9603B32A03CA2FFF553012FEE6F9CF" authorityName="Tyagi &amp; Pal &amp; Sarma &amp; Kumar" authorityYear="2025" box="[151,328,1605,1632]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="19" pageNumber="292" phylum="Arthropoda" rank="species" species="apricus" status="sp. nov.">
<emphasis id="D8E2A4222A03CA2FFF553012FEE6F9CF" bold="true" box="[151,328,1605,1632]" italics="true" pageId="19" pageNumber="292">Thrips apricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3D119592A03CA2FFE8D3012FE03F9CF" box="[335,429,1605,1631]" pageId="19" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A03CA28FF55303EFABEF84D" lastPageId="20" lastPageNumber="293" pageId="19" pageNumber="292" type="description">
<paragraph id="EA2978302A03CA2FFF55303EFEEDF914" blockId="19.[151,429,1605,1668]" box="[151,323,1641,1668]" pageId="19" pageNumber="292">
(
<figureCitation id="72AD64B52A03CA2FFF5D303EFEAEF914" box="[159,256,1641,1668]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="19" pageNumber="292">Figs 195</figureCitation>
206)
</paragraph>
<paragraph id="EA2978302A03CA28FF5530E6FF7CFE8D" blockId="19.[151,1437,1713,2028]" lastBlockId="20.[151,1437,150,285]" lastPageId="20" lastPageNumber="293" pageId="19" pageNumber="292">
<emphasis id="D8E2A4222A03CA2FFF5530E6FED2F95B" box="[151,380,1713,1739]" italics="true" pageId="19" pageNumber="292">Female macroptera.</emphasis>
Body uniformly light brown including legs (
<figureCitation id="72AD64B52A03CA2FFCB930E6FC70F95C" box="[891,990,1713,1740]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="19" pageNumber="292">Fig. 195</figureCitation>
); antennal segment III yellow, IIIVII brown (
<figureCitation id="72AD64B52A03CA2FFF2C3082FEE1F97F" box="[238,335,1749,1775]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="19" pageNumber="292">Fig. 204</figureCitation>
); forewing uniformly yellowish brown (Fig. 206). Head broader than long (
<figureCitation id="72AD64B52A03CA2FFB5B3082FB55F97F" box="[1177,1275,1749,1775]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="19" pageNumber="292">Fig. 197</figureCitation>
); eyes without pigmented facets; ocellar setae III at the tangent of ocellar triangle; six pairs of postocular setae, pair I distinctly longer than others, III and V subequal. Antennae 7-segmented, segments III and IV each with a forked sense cone (
<figureCitation id="72AD64B52A03CA2FFF1C3116FEEAF8CB" box="[222,324,1857,1883]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="19" pageNumber="292">Fig. 204</figureCitation>
). Pronotum broader than long; surface with faint transverse striations; 2 pairs of well-developed posteroangular setae present (
<figureCitation id="72AD64B52A03CA2FFE233132FDEDF8EF" box="[481,579,1893,1919]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="19" pageNumber="292">Fig. 197</figureCitation>
). Mesonotum with transverse striations; antero-median campaniform sensilla present; median pair of setae ahead of posterior margin (
<figureCitation id="72AD64B52A03CA2FFCDE31DEFCD1F833" box="[796,895,1929,1955]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="19" pageNumber="292">Fig. 198</figureCitation>
). Metanotum with longitudinal striations, a few transverse lines anteriorly; median pair of setae far behind the anterior margin; the distance between the median pair less than the median and submedian setae; campaniform sensilla present. Forewing first vein with 7 basal and 3 discal setae; second vein with 14 setae, clavus with 5 veinal and one discal seta (Fig. 206). Abdominal tergite II with 4 lateromarginal setae (
<figureCitation id="72AD64B52A04CA28FE3536ECFDF4FF45" box="[503,602,187,213]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="20" pageNumber="293">Fig. 200</figureCitation>
); abdominal tergite VIII with posteromarginal comb complete (
<figureCitation id="72AD64B52A04CA28FAEE36ECFA3EFF45" box="[1324,1424,187,213]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="20" pageNumber="293">Fig. 201</figureCitation>
). Abdominal sternites IIIVII with discal setae (
<figureCitation id="72AD64B52A04CA28FD5E3688FD50FF69" box="[668,766,223,249]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="20" pageNumber="293">Fig. 199</figureCitation>
); S1 setae on sternite VII ahead of the posterior margin (
<figureCitation id="72AD64B52A04CA28FAAD3688FF6BFE8D" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="20" pageNumber="293">Fig. 199</figureCitation>
).
</paragraph>
<caption id="BEE928B82A04CA28FF5533A6FB95F9BD" pageId="20" pageNumber="293" startId="20.[151,264,1521,1545]" targetBox="[157,1430,315,1492]" targetPageId="20" targetType="figure">
<paragraph id="EA2978302A04CA28FF5533A6FB95F9BD" blockId="20.[151,1436,1521,1581]" pageId="20" pageNumber="293">
<emphasis id="D8E2A4222A04CA28FF5533A6FEC0F999" bold="true" box="[151,366,1521,1545]" pageId="20" pageNumber="293">FIGURES 188194.</emphasis>
Sternites IIIVI, male: (188)
<taxonomicName id="2D9603B32A04CA28FD6233A5FCABF999" authorityName="Bhatti" authorityYear="1970" box="[672,773,1522,1545]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="20" pageNumber="293" phylum="Arthropoda" rank="species" species="tanicus">
<emphasis id="D8E2A4222A04CA28FD6233A5FD00F999" box="[672,686,1522,1545]" italics="true" pageId="20" pageNumber="293">T</emphasis>
.
<emphasis id="D8E2A4222A04CA28FD7E33A5FCABF999" box="[700,773,1522,1545]" italics="true" pageId="20" pageNumber="293">tanicus</emphasis>
</taxonomicName>
; (189)
<taxonomicName id="2D9603B32A04CA28FC9333A5FC66F999" authorityName="Priesner" authorityYear="1927" box="[849,968,1521,1545]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="20" pageNumber="293" phylum="Arthropoda" rank="species" species="trehernei">
<emphasis id="D8E2A4222A04CA28FC9333A5FCF1F999" box="[849,863,1522,1545]" italics="true" pageId="20" pageNumber="293">T</emphasis>
.
<emphasis id="D8E2A4222A04CA28FCAE33A6FC66F999" box="[876,968,1521,1545]" italics="true" pageId="20" pageNumber="293">trehernei</emphasis>
</taxonomicName>
. (190)
<taxonomicName id="2D9603B32A04CA28FBD633A5FBDFF999" authorityName="Bhatti" authorityYear="1980" box="[1044,1137,1522,1545]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="20" pageNumber="293" phylum="Arthropoda" rank="species" species="arorai">
<emphasis id="D8E2A4222A04CA28FBD633A5FB8CF999" box="[1044,1058,1522,1545]" italics="true" pageId="20" pageNumber="293">T</emphasis>
.
<emphasis id="D8E2A4222A04CA28FBED33A5FBDFF999" box="[1071,1137,1522,1545]" italics="true" pageId="20" pageNumber="293">arorai</emphasis>
</taxonomicName>
<emphasis id="D8E2A4222A04CA28FBB033A5FBD8F999" box="[1138,1142,1522,1545]" italics="true" pageId="20" pageNumber="293">,</emphasis>
sternites IIIVI, male; (191)
<taxonomicName id="2D9603B32A04CA28FF553041FF42F9BD" authorityName="Bhatti" authorityYear="1980" box="[151,236,1558,1581]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="20" pageNumber="293" phylum="Arthropoda" rank="species" species="xenos">
<emphasis id="D8E2A4222A04CA28FF553041FF0BF9BD" box="[151,165,1558,1581]" italics="true" pageId="20" pageNumber="293">T</emphasis>
.
<emphasis id="D8E2A4222A04CA28FF703041FF42F9BD" box="[178,236,1558,1581]" italics="true" pageId="20" pageNumber="293">xenos</emphasis>
</taxonomicName>
, sternite VII. Tergite IX, male: (192)
<taxonomicName id="2D9603B32A04CA28FDAB3041FD65F9BD" authorityName="Priesner" authorityYear="1934" box="[617,715,1557,1581]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="20" pageNumber="293" phylum="Arthropoda" rank="species" species="malloti">
<emphasis id="D8E2A4222A04CA28FDAB3041FDD9F9BD" box="[617,631,1558,1581]" italics="true" pageId="20" pageNumber="293">T</emphasis>
.
<emphasis id="D8E2A4222A04CA28FD413042FD65F9BD" box="[643,715,1557,1581]" italics="true" pageId="20" pageNumber="293">malloti</emphasis>
</taxonomicName>
; (193)
<taxonomicName id="2D9603B32A04CA28FCD43041FC28F9BD" authorityName="zur Strassen" authorityYear="1978" box="[790,902,1558,1581]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="20" pageNumber="293" phylum="Arthropoda" rank="species" species="speratus">
<emphasis id="D8E2A4222A04CA28FCD43041FC8AF9BD" box="[790,804,1558,1581]" italics="true" pageId="20" pageNumber="293">T</emphasis>
.
<emphasis id="D8E2A4222A04CA28FCF23041FC28F9BD" box="[816,902,1558,1581]" italics="true" pageId="20" pageNumber="293">speratus</emphasis>
</taxonomicName>
; (194)
<taxonomicName id="2D9603B32A04CA28FC133041FB9BF9BD" authorityName="Bhatti" authorityYear="1970" box="[977,1077,1558,1581]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="20" pageNumber="293" phylum="Arthropoda" rank="species" species="tanicus">
<emphasis id="D8E2A4222A04CA28FC133041FC71F9BD" box="[977,991,1558,1581]" italics="true" pageId="20" pageNumber="293">T</emphasis>
.
<emphasis id="D8E2A4222A04CA28FC2E3041FB9BF9BD" box="[1004,1077,1558,1581]" italics="true" pageId="20" pageNumber="293">tanicus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="EA2978302A04CA28FF053028FB41F971" blockId="20.[151,1437,1663,2014]" pageId="20" pageNumber="293">
<emphasis id="D8E2A4222A04CA28FF053028FED0F909" bold="true" box="[199,382,1663,1689]" pageId="20" pageNumber="293">Measurements.</emphasis>
(
<typeStatus id="352DC6922A04CA28FE493028FE5BF909" box="[395,501,1663,1689]" pageId="20" pageNumber="293" type="holotype">Holotype</typeStatus>
female in microns). Body length 1427. Head length (width across cheeks) 87 (122); compound eye dorsal length (width) 55 (33); pronotum median length (width) 108 (135). Forewing length 846. Antenna segments IVII length (width): 21(26), 34 (24), 38 (19), 44 (17), 35 (19), 98 (37), 28 (14).
</paragraph>
<paragraph id="EA2978302A04CA28FF0530BCFAF1F8E1" blockId="20.[151,1437,1663,2014]" pageId="20" pageNumber="293">
<emphasis id="D8E2A4222A04CA28FF0530BCFE3EF895" box="[199,400,1771,1797]" italics="true" pageId="20" pageNumber="293">Male macroptera.</emphasis>
Body colour similar to female (
<figureCitation id="72AD64B52A04CA28FD3330BCFCFCF895" box="[753,850,1771,1797]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="20" pageNumber="293">Fig. 196</figureCitation>
). Abdominal tergite VIII with posteromarginal comb absent; IX with two pairs of campaniform sensilla, S1 setae on tergite IX longer than S2 and little ahead to S2, the distance between S1 setae slightly more than S1 and S2 setae, S3 strongly developed and half of the S2 length (
<figureCitation id="72AD64B52A04CA28FAAD3164FF6BF8E2" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="20" pageNumber="293">Fig. 205</figureCitation>
). Abdominal sternites IIIVIII with discal setae; sternites IIIVII each with an oval pore plate (
<figureCitation id="72AD64B52A04CA28FB323100FAFCF8E1" box="[1264,1362,1879,1905]" captionStart="FIGURES 195205" captionStartId="21.[151,264,1943,1967]" captionTargetBox="[203,1383,182,1919]" captionTargetId="figure-17@21.[196,1391,181,1919]" captionTargetPageId="21" captionText="FIGURES 195205. Thrips apricus: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female." pageId="20" pageNumber="293">Fig. 203</figureCitation>
).
</paragraph>
<paragraph id="EA2978302A04CA28FF05312CFABEF84D" blockId="20.[151,1437,1663,2014]" pageId="20" pageNumber="293">
<emphasis id="D8E2A4222A04CA28FF05312CFED0F805" bold="true" box="[199,382,1915,1941]" pageId="20" pageNumber="293">Measurements.</emphasis>
(
<typeStatus id="352DC6922A04CA28FE4F312CFE5FF805" box="[397,497,1915,1941]" pageId="20" pageNumber="293" type="paratype">Paratype</typeStatus>
male in microns). Body length 1212. Head length (width across cheeks) 86 (108); compound eye dorsal length (width) 53 (29); pronotum median length (width) 108 (135). Forewing length 747. Antenna segments IVII length (width): 14 (22), 35 (19), 41 (16), 40 (18), 34 (18), 58 (19) and 14 (7).
</paragraph>
</subSubSection>
<caption id="BEE928B82A05CA29FF5531C0FC9DF867" pageId="21" pageNumber="294" startId="21.[151,264,1943,1967]" targetBox="[203,1383,182,1919]" targetPageId="21" targetType="figure">
<paragraph id="EA2978302A05CA29FF5531C0FC9DF867" blockId="21.[151,1437,1943,2040]" pageId="21" pageNumber="294">
<emphasis id="D8E2A4222A05CA29FF5531C0FDA8F83F" bold="true" box="[151,518,1943,1967]" pageId="21" pageNumber="294">
FIGURES 195205.
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<emphasis id="D8E2A4222A05CA29FEB231C0FDA8F83F" bold="true" box="[368,518,1943,1967]" italics="true" pageId="21" pageNumber="294">Thrips apricus</emphasis>
</taxonomicName>
</emphasis>
: (195) Female; (196) Male; (197) Head and Pronotum, female; (198) Meso- and metanotum, female; (199) Sternites VVII, female; (200) Tergite II, female; (201) Tergite VIII, female; (203) Sternites IIIVII, male; (204) Antenna, female; (205) Tergite IX, male; (206) Forewing, female.
</paragraph>
</caption>
<subSubSection id="A28C2BBB2A06CA2AFF0536C1FD14FF69" pageId="22" pageNumber="295" type="materials_examined">
<paragraph id="EA2978302A06CA2AFF0536C1FD14FF69" blockId="22.[151,1436,150,501]" pageId="22" pageNumber="295">
<emphasis id="D8E2A4222A06CA2AFF0536C1FE3EFF20" bold="true" box="[199,400,150,176]" pageId="22" pageNumber="295">Material studied</emphasis>
.
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<typeStatus id="352DC6922A06CA2AFE6236C0FDA6FF21" box="[416,520,151,177]" pageId="22" pageNumber="295" type="holotype">Holotype</typeStatus>
female,
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<collectingCountry id="928138A02A06CA2AFDAC36C1FD6EFF20" box="[622,704,150,176]" name="India" pageId="22" pageNumber="295">INDIA</collectingCountry>
</emphasis>
,
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<collectingRegion id="2852B6D22A06CA2AFD1336C0FCC9FF21" box="[721,871,151,177]" country="India" name="West Bengal" pageId="22" pageNumber="295">West Bengal</collectingRegion>
</emphasis>
,
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(
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,
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,
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</quantity>
), on herb,
<date id="9E285EF02A06CA2AFF5536ECFEADFF45" box="[151,259,187,213]" pageId="22" pageNumber="295" value="1993-11-01">
<collectingDate id="8E6CA7182A06CA2AFF5536ECFEADFF45" box="[151,259,187,213]" pageId="22" pageNumber="295" value="1993-11-01">1.xi.1993</collectingDate>
</date>
, JSB, in NZC, ZSI (
<collectorName id="47631DE62A06CA2AFE2D36ECFDF8FF45" box="[495,598,187,213]" pageId="22" pageNumber="295">Reg. No.</collectorName>
26224/H17)
</materialsCitation>
.
<materialsCitation id="5AFE726D2A06CA2AFD3436ECFD19FF69" collectingDate="1993-11-01" collectorName="Reg. No." country="India" elevation="1761" latitude="31.39" location="Darjeeling" longLatPrecision="730" longitude="77.11" pageId="22" pageNumber="295" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="paratype">
<typeStatus id="352DC6922A06CA2AFD3436ECFCC9FF45" box="[758,871,187,213]" pageId="22" pageNumber="295" type="paratype">Paratypes</typeStatus>
:
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and
<specimenCount id="FC90B3B92A06CA2AFBD236ECFBF3FF45" box="[1040,1117,187,213]" count="1" pageId="22" pageNumber="295" type="male">1 male</specimenCount>
with same data as holotype (
<collectorName id="47631DE62A06CA2AFF5D3688FEABFF69" box="[159,261,223,249]" pageId="22" pageNumber="295">Reg. No.</collectorName>
26225/H17 and
<collectorName id="47631DE62A06CA2AFE023688FD8BFF69" box="[448,549,223,249]" pageId="22" pageNumber="295">Reg. No.</collectorName>
26226/H17)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A06CA2AFF053754FC78FE8D" box="[199,982,259,285]" pageId="22" pageNumber="295" type="etymology">
<paragraph id="EA2978302A06CA2AFF053754FC78FE8D" blockId="22.[151,1436,150,501]" box="[199,982,259,285]" pageId="22" pageNumber="295">
<emphasis id="D8E2A4222A06CA2AFF053754FEFCFE8D" bold="true" box="[199,338,259,285]" pageId="22" pageNumber="295">Etymology:</emphasis>
The species is named in latin after “the full of sunshine”.
</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A06CA2AFF053771FE9EFE65" pageId="22" pageNumber="295" type="discussion">
<paragraph id="EA2978302A06CA2AFF053771FE9EFE65" blockId="22.[151,1436,150,501]" pageId="22" pageNumber="295">
<emphasis id="D8E2A4222A06CA2AFF053771FEFCFED0" bold="true" box="[199,338,294,320]" pageId="22" pageNumber="295">Comments:</emphasis>
According to
<bibRefCitation id="8E0705C12A06CA2AFE3C3770FD34FED1" author="Bhatti, J. S." box="[510,666,295,321]" pageId="22" pageNumber="295" pagination="109 - 166" refId="ref18866" refString="Bhatti, J. S. (1980) Species of the genus Thrips from India. Systematic Entomology, 5, 109 - 166. https: // doi. org / 10.1111 / j. 1365 - 3113.1980. tb 00404. x" type="journal article" year="1980">Bhatti (1980)</bibRefCitation>
, this new species is similar to
<taxonomicName id="2D9603B32A06CA2AFBCB3770FAA3FED1" authority="(Bagnall)" baseAuthorityName="Bagnall" baseAuthorityYear="1916" box="[1033,1293,295,321]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="D8E2A4222A06CA2AFBCB3770FBB7FED0" box="[1033,1049,295,320]" italics="true" pageId="22" pageNumber="295">T</emphasis>
.
<emphasis id="D8E2A4222A06CA2AFBE93770FB38FED1" box="[1067,1174,295,321]" italics="true" pageId="22" pageNumber="295">longiceps</emphasis>
(Bagnall)
</taxonomicName>
, sharing the metanotal sculpture, position of ocellar setae III and body colour, comb on tergite VIII, and presence of discal setae on abdominal sternites. It can be differentiated from
<taxonomicName id="2D9603B32A06CA2AFD3D3738FC25FE19" baseAuthorityName="Bagnall" baseAuthorityYear="1916" box="[767,907,367,393]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="D8E2A4222A06CA2AFD3D3738FCA1FE18" box="[767,783,367,392]" italics="true" pageId="22" pageNumber="295">T</emphasis>
.
<emphasis id="D8E2A4222A06CA2AFCE23738FC25FE19" box="[800,907,367,393]" italics="true" pageId="22" pageNumber="295">longiceps</emphasis>
</taxonomicName>
by 7-segmented antennae (8-segmented in
<taxonomicName id="2D9603B32A06CA2AFA443738FEACFE3D" baseAuthorityName="Bagnall" baseAuthorityYear="1916" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="D8E2A4222A06CA2AFA443738FA38FE18" box="[1414,1430,367,392]" italics="true" pageId="22" pageNumber="295">T</emphasis>
.
<emphasis id="D8E2A4222A06CA2AFF5537C4FEACFE3D" box="[151,258,403,429]" italics="true" pageId="22" pageNumber="295">longiceps</emphasis>
</taxonomicName>
); antennal segments I and II yellow (brown in
<taxonomicName id="2D9603B32A06CA2AFCDE37C4FC08FE3D" baseAuthorityName="Bagnall" baseAuthorityYear="1916" box="[796,934,403,429]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="D8E2A4222A06CA2AFCDE37C4FC82FE3C" box="[796,812,403,428]" italics="true" pageId="22" pageNumber="295">T</emphasis>
.
<emphasis id="D8E2A4222A06CA2AFCF937C4FC08FE3D" box="[827,934,403,429]" italics="true" pageId="22" pageNumber="295">longiceps</emphasis>
</taxonomicName>
); postocular setae arranged in one row (two rows in
<taxonomicName id="2D9603B32A06CA2AFF3037E0FED0FE41" baseAuthorityName="Bagnall" baseAuthorityYear="1916" box="[242,382,439,465]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="D8E2A4222A06CA2AFF3037E0FEACFE40" box="[242,258,439,464]" italics="true" pageId="22" pageNumber="295">T</emphasis>
.
<emphasis id="D8E2A4222A06CA2AFED237E0FED0FE41" box="[272,382,439,465]" italics="true" pageId="22" pageNumber="295">longiceps</emphasis>
</taxonomicName>
<emphasis id="D8E2A4222A06CA2AFEBC37E0FE2CFE41" box="[382,386,439,465]" italics="true" pageId="22" pageNumber="295">,</emphasis>
poII behind poI and poIII); tergite II with 4 lateral marginal setae (3 lateral marginal setae in
<taxonomicName id="2D9603B32A06CA2AFF55378CFE8EFE65" baseAuthorityName="Bagnall" baseAuthorityYear="1916" box="[151,288,475,501]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="D8E2A4222A06CA2AFF55378CFF09FE64" box="[151,167,475,500]" italics="true" pageId="22" pageNumber="295">T</emphasis>
.
<emphasis id="D8E2A4222A06CA2AFF77378CFE8EFE65" box="[181,288,475,501]" italics="true" pageId="22" pageNumber="295">longiceps</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,236 @@
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<mods:title id="D4548DDFB7817595E74C2341B231304F">Identification key to the species of genus Thrips (Thysanoptera, Thripidae) from India with three new species and one new record</mods:title>
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<emphasis id="D8E2A4222A06CA2AFF553411FE32FDF1" bold="true" box="[151,412,582,609]" italics="true" pageId="22" pageNumber="295">Thrips hemkundensis</emphasis>
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(
<figureCitation id="72AD64B52A06CA2AFF5D343CFE92FD15" box="[159,316,619,645]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Figs 207214</figureCitation>
)
</paragraph>
<paragraph id="EA2978302A06CA2AFF5534E4FB21FB55" blockId="22.[151,1437,691,1833]" pageId="22" pageNumber="295">
<emphasis id="D8E2A4222A06CA2AFF5534E4FED1FD5C" box="[151,383,691,717]" italics="true" pageId="22" pageNumber="295">Female macroptera.</emphasis>
Body uniformly dark brown (
<figureCitation id="72AD64B52A06CA2AFD2634E4FCE4FD5D" box="[740,842,691,717]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 207</figureCitation>
); antennal segments IVIII dark brown (
<figureCitation id="72AD64B52A06CA2AFAEB34E4FA21FD5D" box="[1321,1423,691,717]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 209</figureCitation>
); forewing uniformly brown (
<figureCitation id="72AD64B52A06CA2AFE0C3480FD81FD61" box="[462,559,727,753]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 214</figureCitation>
); fore tibiae slightly pale towards apex, all tarsi yellowish brown. Head broader than long, ocellar triangle without any sculpture; ocellar setae III at the tangent of ocellar triangle; six pairs of well-developed postocular setae, pair I clearly longer than others, II placed well behind I and III (
<figureCitation id="72AD64B52A06CA2AFB783548FAB0FCA9" box="[1210,1310,799,825]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 208</figureCitation>
). Antennae 8-segmented, segment III and IV each with forked sense cone (
<figureCitation id="72AD64B52A06CA2AFC923514FC1FFCCD" box="[848,945,835,861]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 209</figureCitation>
). Pronotal surface with transverse striations, with 1822 discal setae; posteroangular setae 2 pairs; posteromarginal setae 3 pairs. Mesonotum with transverse striations; anteromedian campaniform sensilla present; median pair of setae ahead of the posterior margin. Metanotum with irregular longitudinal reticulation medially surrounded by transverse striations, few transverse lines anteriorly; median pair of setae far behind the anterior margin; the distance between the median pair less than the median and submedian setae; campaniform sensilla present (
<figureCitation id="72AD64B52A06CA2AFCB035A0FC78FB81" box="[882,982,1015,1041]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 210</figureCitation>
). Forewing first vein with 7 basal and 3 distal setae; second vein with 12 setae; clavus with 5 veinal and 1 discal seta (
<figureCitation id="72AD64B52A06CA2AFBC5324CFBC5FBA5" box="[1031,1131,1051,1077]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 214</figureCitation>
). Abdominal tergite II with 3 lateral marginal setae (
<figureCitation id="72AD64B52A06CA2AFE6E3268FDA0FBC9" box="[428,526,1087,1113]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 212</figureCitation>
); VIII with variable posteromarginal comb, microtrichia often irregular or absent medially (
<figureCitation id="72AD64B52A06CA2AFEC83234FEC0FBED" box="[266,366,1123,1149]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 213</figureCitation>
); IX with 2 pairs of campaniform sensilla. Abdominal sternites IIIVII each with discal setae, IIIVI arranged in an irregular transverse row, VII in one row (
<figureCitation id="72AD64B52A06CA2AFCA232D0FC6CFB31" box="[864,962,1159,1185]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 211</figureCitation>
); sternite VII with S1 setae far away from the posterior margin (
<figureCitation id="72AD64B52A06CA2AFE4B32FCFE44FB55" box="[393,490,1195,1221]" captionStart="FIGURES 207214" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[202,1384,182,1920]" captionTargetId="figure-17@23.[196,1390,181,1920]" captionTargetPageId="23" captionText="FIGURES 207214. Thrips hemkundensis, female: (207) Female; (208) Head and Pronotum; (209) Antenna; (210) Meso- and metanotum; (211) Sternites VVII; (212) Tergites III; (213) Tergite VIIIX; (214) Forewing." pageId="22" pageNumber="295">Fig. 211</figureCitation>
). Pleurotergites without ciliate microtrichia and discal setae.
</paragraph>
<paragraph id="EA2978302A06CA2AFF053298FA3DFAA1" blockId="22.[151,1437,691,1833]" pageId="22" pageNumber="295">
<emphasis id="D8E2A4222A06CA2AFF053298FED0FB79" bold="true" box="[199,382,1231,1257]" pageId="22" pageNumber="295">Measurements.</emphasis>
(
<typeStatus id="352DC6922A06CA2AFE523298FE54FB79" box="[400,506,1231,1257]" pageId="22" pageNumber="295" type="holotype">Holotype</typeStatus>
female in microns). Body length 1825. Head length (width across cheeks) 135 (1698); compound eye dorsal length (width) 61 (42); pronotum median length (width) 137 (224). Forewing length 842. Antenna segments IVIII length (width): 24 (31), 42 (25), 57 (21), 56 (22), 44 (21), 60 (23), 7 (9) and 11 (6).
</paragraph>
<paragraph id="EA2978302A06CA2AFF05336CFED6FAC5" blockId="22.[151,1437,691,1833]" box="[199,376,1339,1365]" pageId="22" pageNumber="295">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A06CA2AFF053309FCC2FA51" pageId="22" pageNumber="295" type="materials_examined">
<paragraph id="EA2978302A06CA2AFF053309FCC2FA51" blockId="22.[151,1437,691,1833]" pageId="22" pageNumber="295">
<emphasis id="D8E2A4222A06CA2AFF053309FE22FAE8" bold="true" box="[199,396,1374,1400]" pageId="22" pageNumber="295">Material studied</emphasis>
.
<materialsCitation id="5AFE726D2A06CA2AFE5B3308FB07FA0D" collectingDate="1984-08-17" collectorName="B. K. Negi &amp; A. K. Srivastava &amp; Reg. No." country="India" elevation="1761" latitude="31.39" location="Hemkund" longLatPrecision="730" longitude="77.11" pageId="22" pageNumber="295" specimenCount="1" stateProvince="Uttarakhand" typeStatus="holotype">
<typeStatus id="352DC6922A06CA2AFE5B3308FDAFFAE9" box="[409,513,1375,1401]" pageId="22" pageNumber="295" type="holotype">Holotype</typeStatus>
female,
<emphasis id="D8E2A4222A06CA2AFD9D3309FD1FFAE8" bold="true" box="[607,689,1374,1400]" pageId="22" pageNumber="295">
<collectingCountry id="928138A02A06CA2AFD9D3309FD1FFAE8" box="[607,689,1374,1400]" name="India" pageId="22" pageNumber="295">INDIA</collectingCountry>
</emphasis>
,
<emphasis id="D8E2A4222A06CA2AFD7C3308FCF4FAE9" bold="true" box="[702,858,1375,1401]" pageId="22" pageNumber="295">
<collectingRegion id="2852B6D22A06CA2AFD7C3308FCF4FAE9" box="[702,858,1375,1401]" country="India" name="Uttarakhand" pageId="22" pageNumber="295">Uttarakhand</collectingRegion>
</emphasis>
,
<location id="EF492EEB2A06CA2AFCA43308FC7AFAE9" LSID="urn:lsid:plazi:treatment:623FC9262A06CA2AFF553411FE1AF8B9:EF492EEB2A06CA2AFCA43308FC7AFAE9" box="[870,980,1375,1401]" country="India" latitude="31.39" longLatPrecision="730" longitude="77.11" name="Hemkund" pageId="22" pageNumber="295" stateProvince="Uttarakhand">Hemkund</location>
(
<geoCoordinate id="8FA21EF72A06CA2AFC203308FB91FAE9" box="[994,1087,1375,1401]" degrees="31.39" direction="north" orientation="latitude" pageId="22" pageNumber="295" precision="555" value="31.39">31.39 N</geoCoordinate>
,
<geoCoordinate id="8FA21EF72A06CA2AFB8B3308FB35FAE9" box="[1097,1179,1375,1401]" degrees="77.11" direction="east" orientation="longitude" pageId="22" pageNumber="295" precision="555" value="77.11">77.11E</geoCoordinate>
,
<quantity id="2D6ED5D52A06CA2AFB673308FB55FAE9" box="[1189,1275,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.761" pageId="22" pageNumber="295" unit="m" value="1761.0">
<elevation id="61BB9F032A06CA2AFB673308FB55FAE9" box="[1189,1275,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.761" pageId="22" pageNumber="295" unit="m" value="1761.0">1761 m</elevation>
</quantity>
),
<collectedFrom id="598A3BCD2A06CA2AFACC3308FEB9FA0D" pageId="22" pageNumber="295">
on
<taxonomicName id="2D9603B32A06CA2AFAED3308FEB9FA0D" class="Magnoliopsida" family="Saxifragaceae" genus="Saxifraga" kingdom="Plantae" order="Saxifragales" pageId="22" pageNumber="295" phylum="Tracheophyta" rank="species" species="diversifolia">
<emphasis id="D8E2A4222A06CA2AFAED3308FEB9FA0D" italics="true" pageId="22" pageNumber="295">Saxifraga diversifolia</emphasis>
</taxonomicName>
</collectedFrom>
,
<date id="9E285EF02A06CA2AFEE733D4FE01FA0D" box="[293,431,1411,1437]" pageId="22" pageNumber="295" value="1984-08-17">
<collectingDate id="8E6CA7182A06CA2AFEE733D4FE01FA0D" box="[293,431,1411,1437]" pageId="22" pageNumber="295" value="1984-08-17">17.viii.1984</collectingDate>
</date>
,
<collectorName id="47631DE62A06CA2AFE7933D4FD80FA0D" box="[443,558,1411,1437]" pageId="22" pageNumber="295">B.K. Negi</collectorName>
&amp;
<collectorName id="47631DE62A06CA2AFD9333D4FCA8FA0D" box="[593,774,1411,1437]" pageId="22" pageNumber="295">A.K. Srivastava</collectorName>
, in NZC, ZSI (
<collectorName id="47631DE62A06CA2AFC7233D4FBB8FA0D" box="[944,1046,1411,1437]" pageId="22" pageNumber="295">Reg. No.</collectorName>
26227/H17)
</materialsCitation>
.
<materialsCitation id="5AFE726D2A06CA2AFB7133D4FCC6FA51" collectingDate="1984-08-17" collectorName="Reg. No." country="India" elevation="1761" latitude="31.39" location="Hemkund" longLatPrecision="730" longitude="77.11" pageId="22" pageNumber="295" specimenCount="2" specimenCount-female="2" stateProvince="Uttarakhand" typeStatus="paratype">
<typeStatus id="352DC6922A06CA2AFB7133D4FA8AFA0D" box="[1203,1316,1411,1437]" pageId="22" pageNumber="295" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="FC90B3B92A06CA2AFAF233D4FA32FA0D" box="[1328,1436,1411,1437]" count="2" pageId="22" pageNumber="295" type="female">2 females</specimenCount>
with same data as holotype (
<collectorName id="47631DE62A06CA2AFE1433F0FD92FA51" box="[470,572,1447,1473]" pageId="22" pageNumber="295">Reg. No.</collectorName>
26228/H17 to26229/H17)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A06CA2AFF05339CFCEDFA75" box="[199,835,1483,1509]" pageId="22" pageNumber="295" type="etymology">
<paragraph id="EA2978302A06CA2AFF05339CFCEDFA75" blockId="22.[151,1437,691,1833]" box="[199,835,1483,1509]" pageId="22" pageNumber="295">
<emphasis id="D8E2A4222A06CA2AFF05339CFEFCFA75" bold="true" box="[199,338,1483,1509]" pageId="22" pageNumber="295">Etymology:</emphasis>
The species is named after the
<typeStatus id="352DC6922A06CA2AFD70339CFD4CFA75" box="[690,738,1483,1509]" pageId="22" pageNumber="295">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A06CA2AFF0533B9FE1AF8B9" pageId="22" pageNumber="295" type="discussion">
<paragraph id="EA2978302A06CA2AFF0533B9FE1AF8B9" blockId="22.[151,1437,691,1833]" pageId="22" pageNumber="295">
<emphasis id="D8E2A4222A06CA2AFF0533B9FEFCF998" bold="true" box="[199,338,1518,1544]" pageId="22" pageNumber="295">Comments:</emphasis>
<taxonomicName id="2D9603B32A06CA2AFE9B33B8FDE8F999" authorityName="Tyagi &amp; Pal &amp; Sarma &amp; Kumar" authorityYear="2025" box="[345,582,1519,1545]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="hemkundensis" status="sp. nov.">
<emphasis id="D8E2A4222A06CA2AFE9B33B8FDE8F999" box="[345,582,1519,1545]" italics="true" pageId="22" pageNumber="295">Thrips hemkundensis</emphasis>
</taxonomicName>
<emphasis id="D8E2A4222A06CA2AFD8F33B8FD09F999" bold="true" box="[589,679,1519,1545]" pageId="22" pageNumber="295">
<taxonomicNameLabel id="C3D119592A06CA2AFD8F33B8FD09F999" box="[589,679,1519,1545]" pageId="22" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="2D9603B32A06CA2AFCFB33B8FC5FF999" authority="Uzel" authorityName="Uzel" box="[825,1009,1519,1545]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFCFB33B8FC18F998" box="[825,950,1519,1545]" italics="true" pageId="22" pageNumber="295">Thrips pini</emphasis>
Uzel
</taxonomicName>
but it can be differentiated by smooth ocellar triangle (weakly sculptured in
<taxonomicName id="2D9603B32A06CA2AFD8A3044FD3FF9BC" box="[584,657,1555,1580]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFD8A3044FD3FF9BC" box="[584,657,1555,1580]" italics="true" pageId="22" pageNumber="295">T. pini</emphasis>
</taxonomicName>
); cheeks not rounded (rounded in
<taxonomicName id="2D9603B32A06CA2AFBD93044FBCAF9BC" box="[1051,1124,1555,1580]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFBD93044FBCAF9BC" box="[1051,1124,1555,1580]" italics="true" pageId="22" pageNumber="295">T. pini</emphasis>
</taxonomicName>
); head without tubercles in front of eyes (head usually with small tubercles arranged longitudinally in front of eyes in
<taxonomicName id="2D9603B32A06CA2AFB483060FB7FF9C0" box="[1162,1233,1591,1616]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFB483060FB7FF9C0" box="[1162,1233,1591,1616]" italics="true" pageId="22" pageNumber="295">T. pini</emphasis>
</taxonomicName>
); postocular setae biserial, poII and poIV well developed (postocular setae uniserial, po II and po IV minute in
<taxonomicName id="2D9603B32A06CA2AFB09300CFABBF9E4" box="[1227,1301,1627,1652]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFB09300CFABBF9E4" box="[1227,1301,1627,1652]" italics="true" pageId="22" pageNumber="295">T. pini</emphasis>
</taxonomicName>
); pronotum with 1822 discal setae (
<quantity id="2D6ED5D52A06CA2AFE703029FDB6F909" box="[434,536,1662,1689]" metricMagnitude="-1" metricUnit="m" metricValue="7.493" metricValueMax="8.89" metricValueMin="6.096" pageId="22" pageNumber="295" unit="in" value="29.5" valueMax="35.0" valueMin="24.0">2435 in</quantity>
<taxonomicName id="2D9603B32A06CA2AFDE23028FDC7F908" box="[544,617,1663,1688]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFDE23028FDC7F908" box="[544,617,1663,1688]" italics="true" pageId="22" pageNumber="295">T. pini</emphasis>
</taxonomicName>
); antennae brown (antennal segment I to II pale brown, III brown often with basal third yellowish, IV brown or dark brown with extreme base slightly pale, V dark brown with extreme base slightly pale in
<taxonomicName id="2D9603B32A06CA2AFE473090FE61F970" box="[389,463,1735,1760]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFE473090FE61F970" box="[389,463,1735,1760]" italics="true" pageId="22" pageNumber="295">T. pini</emphasis>
</taxonomicName>
); metanotum with longitudinal reticulations in middle (with irregular reticulations or transverse anastomosing striae at middle in in
<taxonomicName id="2D9603B32A06CA2AFD6730BCFD40F894" box="[677,750,1771,1796]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFD6730BCFD40F894" box="[677,750,1771,1796]" italics="true" pageId="22" pageNumber="295">T. pini</emphasis>
</taxonomicName>
); abdominal tergite II with 3 lateral marginal setae (4 lateral marginal setae in
<taxonomicName id="2D9603B32A06CA2AFE9F3158FE0AF8B8" box="[349,420,1807,1832]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="22" pageNumber="295" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="D8E2A4222A06CA2AFE9F3158FE0AF8B8" box="[349,420,1807,1832]" italics="true" pageId="22" pageNumber="295">T. pini</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="E4F54877F1B0BFCFB2AD84489D5EAC5A">Identification key to the species of genus Thrips (Thysanoptera, Thripidae) from India with three new species and one new record</mods:title>
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<mods:namePart id="F09257C27087A1B0E7607CDED1BE419D">Pal, Shash</mods:namePart>
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<heading id="B161CF5C2A08CA24FF5533D4FDE7FA0D" bold="true" box="[151,585,1411,1438]" fontSize="11" level="1" pageId="24" pageNumber="297" reason="1">
<taxonomicName id="2D9603B32A08CA24FF5533D4FDE7FA0D" ID-CoL="56ML3" authority="Mound &amp; Masumoto" authorityName="Mound &amp; Masumoto" authorityYear="2005" box="[151,585,1411,1438]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="24" pageNumber="297" phylum="Arthropoda" rank="species" species="safrus">
<emphasis id="D8E2A4222A08CA24FF5533D4FDE7FA0D" bold="true" box="[151,585,1411,1438]" pageId="24" pageNumber="297">
<emphasis id="D8E2A4222A08CA24FF5533D4FE97FA0D" bold="true" box="[151,313,1411,1438]" italics="true" pageId="24" pageNumber="297">Thrips safrus</emphasis>
Mound &amp; Masumoto
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<taxonomicName id="2D9603B32A08CA24FF55339FFDCEFA4F" ID-CoL="56ML3" authority="Mound &amp; Masumoto, 2005: 45" authorityName="Mound &amp; Masumoto" authorityPageNumber="45" authorityYear="2005" box="[151,608,1479,1504]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="24" pageNumber="297" phylum="Arthropoda" rank="species" species="safrus">
<emphasis id="D8E2A4222A08CA24FF55339FFEB0FA70" box="[151,286,1480,1504]" italics="true" pageId="24" pageNumber="297">Thrips safrus</emphasis>
<bibRefCitation id="8E0705C12A08CA24FEE6339FFDCEFA4F" author="Mound, L. A. &amp; Masumoto, M." box="[292,608,1479,1504]" pageId="24" pageNumber="297" pagination="1 - 64" refId="ref19226" refString="Mound, L. A. &amp; Masumoto, M. (2005) The genus Thrips (Thysanoptera, Thripidae) in Australia, New Caledonia and New Zealand. Zootaxa, 1020 (1), 1 - 64. https: // doi. org / 10.11646 / zootaxa. 1020.1.1" type="journal article" year="2005">Mound &amp; Masumoto, 2005: 45</bibRefCitation>
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<paragraph id="EA2978302A08CA24FF553059FBE6F8B4" blockId="24.[151,1437,1550,1900]" pageId="24" pageNumber="297">Diagnosis: Body and legs white, last antennal segments light brown, remaining segments pale; forewings pale. Antennae 7-segmented; ocellar setae III arise just within anterolateral margins of ocellar triangle close to first ocellus, postocular setae arranged in straight row, subequal in size. Pronotum with transverse striae, 45 posteromarginal setae, outer postero-angular seta shorter than inner seta. Metanotum irregularly reticulate medially, median setae well behind anterior margin, campaniform sensilla present. Forewing first vein usually with 3 setae on distal half; clavus with terminal seta longer than subterminal seta. Abdominal tergite II with 3 lateral setae; abdominal tergite VIII with posteromarginal comb represented by a few teeth laterally; sternite II with 1 or 2 discal setae, IIIVII with 1525 discal setae in an irregular transverse row; pleurotergites with no discal setae.</paragraph>
<paragraph id="EA2978302A08CA24FF05317AFE88F8FC" blockId="24.[151,1437,1550,1900]" pageId="24" pageNumber="297">
<emphasis id="D8E2A4222A08CA24FF05317AFE32F8D7" bold="true" box="[199,412,1837,1863]" pageId="24" pageNumber="297">Specimen studied</emphasis>
:
<materialsCitation id="5AFE726D2A08CA24FE6D3179FE8CF8FC" collectingDate="1967-04-19" collectorName="Reg. No." country="India" location="Punjab" municipality="Ludihana" pageId="24" pageNumber="297" specimenCount="1" specimenCount-female="1">
<collectingCountry id="928138A02A08CA24FE6D3179FDADF8D8" box="[431,515,1838,1864]" name="India" pageId="24" pageNumber="297">INDIA</collectingCountry>
:
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,
<collectingMunicipality id="0A4DE24A2A08CA24FDB23179FD73F8D8" box="[624,733,1838,1864]" pageId="24" pageNumber="297">Ludihana</collectingMunicipality>
,
<specimenCount id="FC90B3B92A08CA24FD293179FCFDF8D8" box="[747,851,1838,1864]" count="1" pageId="24" pageNumber="297" type="female">1 female</specimenCount>
,
<date id="9E285EF02A08CA24FCA03179FC74F8D8" box="[866,986,1838,1864]" pageId="24" pageNumber="297" value="1967-04-19">
<collectingDate id="8E6CA7182A08CA24FCA03179FC74F8D8" box="[866,986,1838,1864]" pageId="24" pageNumber="297" value="1967-04-19">19.iv.1967</collectingDate>
</date>
,
<taxonomicName id="2D9603B32A08CA24FC283178FBE9F8D8" box="[1002,1095,1839,1864]" class="Magnoliopsida" family="Verbenaceae" genus="Lantana" kingdom="Plantae" order="Lamiales" pageId="24" pageNumber="297" phylum="Tracheophyta" rank="genus">
<emphasis id="D8E2A4222A08CA24FC283178FBE9F8D8" box="[1002,1095,1839,1864]" italics="true" pageId="24" pageNumber="297">Lantana</emphasis>
</taxonomicName>
flowers, coll. JSB (
<collectorName id="47631DE62A08CA24FAF13179FA33F8D8" box="[1331,1437,1838,1864]" pageId="24" pageNumber="297">Reg. No.</collectorName>
18791/H17)
</materialsCitation>
.
</paragraph>
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<paragraph id="EA2978302A09CA25FF5531C0FCBDF843" blockId="25.[151,1436,1943,2004]" pageId="25" pageNumber="298">
<emphasis id="D8E2A4222A09CA25FF5531C0FDF6F83F" bold="true" box="[151,600,1943,1968]" pageId="25" pageNumber="298">
FIGURES 215221.
<taxonomicName id="2D9603B32A09CA25FEB631C0FDADF83F" authorityName="Tyagi &amp; Pal &amp; Sarma &amp; Kumar" authorityYear="2025" box="[372,515,1943,1967]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="25" pageNumber="298" phylum="Arthropoda" rank="species" species="roseae">
<emphasis id="D8E2A4222A09CA25FEB631C0FDADF83F" bold="true" box="[372,515,1943,1967]" italics="true" pageId="25" pageNumber="298">Thrips roseae</emphasis>
</taxonomicName>
, female
</emphasis>
: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing.
</paragraph>
</caption>
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<paragraph id="EA2978302A0ACA26FF5531C0FEE9F844" blockId="26.[151,1436,1943,2004]" pageId="26" pageNumber="299">
<emphasis id="D8E2A4222A0ACA26FF5531C0FD0BF83F" bold="true" box="[151,677,1943,1968]" pageId="26" pageNumber="299">FIGURE 222. Slides of Type specimens of studied</emphasis>
(
<taxonomicName id="2D9603B32A0ACA26FD7031CFFD5EF820" authorityName="Bhatti" authorityYear="1980" box="[690,752,1944,1968]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="alatus">
<emphasis id="D8E2A4222A0ACA26FD7031CFFD5EF820" box="[690,752,1944,1968]" italics="true" pageId="26" pageNumber="299">alatus</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FD3E31CFFC92F83F" authorityName="Bhatti" authorityYear="1980" box="[764,828,1944,1967]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="arorai">
<emphasis id="D8E2A4222A0ACA26FD3E31CFFC92F83F" box="[764,828,1944,1967]" italics="true" pageId="26" pageNumber="299">arorai</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FC8531CFFC3EF83F" authorityName="Bhatti" authorityYear="1967" box="[839,912,1944,1967]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="atactus">
<emphasis id="D8E2A4222A0ACA26FC8531CFFC3EF83F" box="[839,912,1944,1967]" italics="true" pageId="26" pageNumber="299">atactus</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FC5E31CFFC7EF820" authorityName="Bhatti" authorityYear="1980" box="[924,976,1944,1968]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="cedri">
<emphasis id="D8E2A4222A0ACA26FC5E31CFFC7EF820" box="[924,976,1944,1968]" italics="true" pageId="26" pageNumber="299">cedri</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FC1931CFFBB8F820" authorityName="Bhatti" authorityYear="1980" box="[987,1046,1944,1968]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="dorax">
<emphasis id="D8E2A4222A0ACA26FC1931CFFBB8F820" box="[987,1046,1944,1968]" italics="true" pageId="26" pageNumber="299">dorax</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FBE031CFFB6BF820" authorityName="Ananthakrishnan &amp; Jagadish" authorityYear="1966" box="[1058,1221,1944,1968]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="kodaikanalensis">
<emphasis id="D8E2A4222A0ACA26FBE031CFFB6BF820" box="[1058,1221,1944,1968]" italics="true" pageId="26" pageNumber="299">kodaikanalensis</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FB1331CFFA8DF820" authorityName="Bhatti" authorityYear="1999" box="[1233,1315,1944,1968]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="chandni">
<emphasis id="D8E2A4222A0ACA26FB1331CFFA8DF820" box="[1233,1315,1944,1968]" italics="true" pageId="26" pageNumber="299">chandni</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FAEC31CFFA38F83F" authorityName="Vijay Veer &amp; Srivastava" authorityYear="1985" box="[1326,1430,1944,1967]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="sensarmai">
<emphasis id="D8E2A4222A0ACA26FAEC31CFFA38F83F" box="[1326,1430,1944,1967]" italics="true" pageId="26" pageNumber="299">sensarmai</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FF5531EBFF4EF844" authorityName="Bhatti" authorityYear="1980" box="[151,224,1980,2004]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="garuda">
<emphasis id="D8E2A4222A0ACA26FF5531EBFF4EF844" box="[151,224,1980,2004]" italics="true" pageId="26" pageNumber="299">garuda</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0ACA26FF2F31EBFE97F844" authorityName="Tyagi &amp; Kumar" authorityYear="2015" box="[237,313,1980,2004]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="26" pageNumber="299" phylum="Arthropoda" rank="species" species="moundi">
<emphasis id="D8E2A4222A0ACA26FF2F31EBFE97F844" box="[237,313,1980,2004]" italics="true" pageId="26" pageNumber="299">moundi</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
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<paragraph id="EA2978302A0BCA27FF5530B2FBF1F96E" blockId="27.[151,1119,1765,1790]" box="[151,1119,1765,1790]" pageId="27" pageNumber="300">
<emphasis id="D8E2A4222A0BCA27FF5530B2FD3CF96D" bold="true" box="[151,658,1765,1790]" pageId="27" pageNumber="300">FIGURE 223. Slides of Type specimens studied</emphasis>
(
<taxonomicName id="2D9603B32A0BCA27FD6230B1FD46F96E" authorityName="Bhatti" authorityYear="1980" box="[672,744,1766,1790]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="species" species="levatus">
<emphasis id="D8E2A4222A0BCA27FD6230B1FD46F96E" box="[672,744,1766,1790]" italics="true" pageId="27" pageNumber="300">levatus</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0BCA27FD3730B1FC81F96D" authorityName="Bhatti" authorityYear="1967" box="[757,815,1766,1789]" class="Insecta" family="Thripidae" genus="Kemothrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="species" species="mirus">
<emphasis id="D8E2A4222A0BCA27FD3730B1FC81F96D" box="[757,815,1766,1789]" italics="true" pageId="27" pageNumber="300">mirus</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0BCA27FCF930B1FC2AF96D" authorityName="Bhatti" authorityYear="1970" box="[827,900,1766,1789]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="species" species="tanicus">
<emphasis id="D8E2A4222A0BCA27FCF930B1FC2AF96D" box="[827,900,1766,1789]" italics="true" pageId="27" pageNumber="300">tanicus</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0BCA27FC5330B1FC13F96E" authorityName="Bhatti" authorityYear="1967" box="[913,957,1766,1790]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="species" species="latis">
<emphasis id="D8E2A4222A0BCA27FC5330B1FC13F96E" box="[913,957,1766,1790]" italics="true" pageId="27" pageNumber="300">latis</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0BCA27FC0B30B1FBA4F96D" authorityName="Bhatti" authorityYear="1980" box="[969,1034,1766,1789]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="species" species="taurus">
<emphasis id="D8E2A4222A0BCA27FC0B30B1FBA4F96D" box="[969,1034,1766,1789]" italics="true" pageId="27" pageNumber="300">taurus</emphasis>
</taxonomicName>
,
<taxonomicName id="2D9603B32A0BCA27FBD530B1FBFFF96D" authorityName="Bhatti" authorityYear="1980" box="[1047,1105,1766,1789]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="species" species="xenos">
<emphasis id="D8E2A4222A0BCA27FBD530B1FBFFF96D" box="[1047,1105,1766,1789]" italics="true" pageId="27" pageNumber="300">xenos</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<subSubSection id="A28C2BBB2A0BCA20FF053107FE9FFF45" lastPageId="28" lastPageNumber="301" pageId="27" pageNumber="300" type="discussion">
<paragraph id="EA2978302A0BCA20FF053107FE9FFF45" blockId="27.[151,1437,1872,2042]" lastBlockId="28.[151,1437,151,213]" lastPageId="28" lastPageNumber="301" pageId="27" pageNumber="300">
<emphasis id="D8E2A4222A0BCA27FF053107FEE6F8FA" bold="true" box="[199,328,1872,1898]" pageId="27" pageNumber="300">Comments</emphasis>
: Described originally from various sites in northern
<collectingCountry id="928138A02A0BCA27FC473107FC41F8FA" box="[901,1007,1872,1898]" name="Australia" pageId="27" pageNumber="300">Australia</collectingCountry>
, with
<specimenCount id="FC90B3B92A0BCA27FBED3107FB07F8FA" box="[1071,1193,1872,1898]" count="1" pageId="27" pageNumber="300" type="female">one female</specimenCount>
from
<collectingCountry id="928138A02A0BCA27FB283107FA36F8FA" box="[1258,1432,1872,1898]" name="New Caledonia" pageId="27" pageNumber="300">New Caledonia</collectingCountry>
, this species has otherwise been found established only on the Hawaiian Islands (
<bibRefCitation id="8E0705C12A0BCA27FBF73123FABFF81E" author="Mound, L. A. &amp; Matsunaga, J. &amp; Bushe, B. &amp; Hoddle, M. S. &amp; Wells, A." box="[1077,1297,1908,1934]" pageId="27" pageNumber="300" pagination="17 - 28" refId="ref19280" refString="Mound, L. A., Matsunaga, J., Bushe, B., Hoddle, M. S. &amp; Wells, A. (2017) Adventive Thysanoptera Species on the Hawaiian Islands: New Records and Putative Host Associations. Proceedings of the Hawaiian Entomological Society, 49, 17 - 28." type="journal article" year="2017">
Mound
<emphasis id="D8E2A4222A0BCA27FB533122FB6AF81E" box="[1169,1220,1908,1934]" italics="true" pageId="27" pageNumber="300">et al</emphasis>
. 2017
</bibRefCitation>
). The single female listed above remained unidentified in the collections of J.S. Bhatti, but using the key to species of
<taxonomicName id="2D9603B32A0BCA27FA9131CFFA32F822" box="[1363,1436,1944,1970]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="D8E2A4222A0BCA27FA9131CFFA32F822" box="[1363,1436,1944,1970]" italics="true" pageId="27" pageNumber="300">Thrips</emphasis>
</taxonomicName>
from Indian subregion (
<bibRefCitation id="8E0705C12A0BCA27FE6131EBFD84F846" author="Bhatti, J. S." box="[419,554,1980,2006]" pageId="27" pageNumber="300" pagination="109 - 166" refId="ref18866" refString="Bhatti, J. S. (1980) Species of the genus Thrips from India. Systematic Entomology, 5, 109 - 166. https: // doi. org / 10.1111 / j. 1365 - 3113.1980. tb 00404. x" type="journal article" year="1980">Bhatti 1980</bibRefCitation>
) it reached to
<taxonomicName id="2D9603B32A0BCA27FD0D31EAFCF6F846" baseAuthorityName="Bagnall" baseAuthorityYear="1916" box="[719,856,1980,2006]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="D8E2A4222A0BCA27FD0D31EAFD71F846" box="[719,735,1981,2006]" italics="true" pageId="27" pageNumber="300">T</emphasis>
.
<emphasis id="D8E2A4222A0BCA27FD2F31EBFCF6F846" box="[749,856,1980,2006]" italics="true" pageId="27" pageNumber="300">longiceps</emphasis>
</taxonomicName>
. However, from that species it differs in the colour of the body, position of ocellar setae III, and number of antennal segments. Identified here as
<taxonomicName id="2D9603B32A0BCA27FB0F31B6FA80F86A" authorityName="Mound &amp; Masumoto" authorityYear="2005" box="[1229,1326,2016,2042]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="27" pageNumber="300" phylum="Arthropoda" rank="species" species="safrus">
<emphasis id="D8E2A4222A0BCA27FB0F31B6FA80F86A" box="[1229,1326,2016,2042]" italics="true" pageId="27" pageNumber="300">T. safrus</emphasis>
</taxonomicName>
based on the Mound and Masumoto key, this female represents a particularly unusual record of a species that is presumably non-endemic.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,211 @@
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<emphasis id="D8E2A4222A08CA24FF5536C0FE92FF21" bold="true" box="[151,316,151,178]" italics="true" pageId="24" pageNumber="297">Thrips roseae</emphasis>
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<taxonomicNameLabel id="C3D119592A08CA24FE8636C0FE0CFF21" box="[324,418,151,177]" pageId="24" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="EA2978302A08CA24FF5536ECFEEDFF46" blockId="24.[151,418,151,214]" box="[151,323,187,214]" pageId="24" pageNumber="297">
(
<figureCitation id="72AD64B52A08CA24FF5D36ECFE95FF46" box="[159,315,187,214]" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Figs 215221</figureCitation>
)
</paragraph>
<paragraph id="EA2978302A08CA24FF553754FD19FD61" blockId="24.[151,1437,259,1330]" pageId="24" pageNumber="297">
<emphasis id="D8E2A4222A08CA24FF553754FED1FE8D" box="[151,383,259,285]" italics="true" pageId="24" pageNumber="297">Female macroptera.</emphasis>
Body uniformly brown including legs, fore tibiae pale towards apex, all tarsi yellow (
<figureCitation id="72AD64B52A08CA24FAAD3754FF6BFED2" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 215</figureCitation>
); antennal segments III brown, IIIIV and basal ¼ of V yellow, proximal ¾ of V and VIVIII brown (
<figureCitation id="72AD64B52A08CA24FAAD3770FF6BFEF5" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 216</figureCitation>
); forewing uniformly brown with light shade towards basal end (
<figureCitation id="72AD64B52A08CA24FC07371CFB85FEF5" box="[965,1067,331,357]" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 221</figureCitation>
). Head little broader than long; eyes without pigmented facets; ocellar setae III placed on the outer side of the tangent between fore and hind ocelli; six pairs of postocular setae, poI, and poIV distinctly longer than others, poIV longer than poI (
<figureCitation id="72AD64B52A08CA24FAEE37C4FA3EFE3D" box="[1324,1424,403,429]" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 219</figureCitation>
). Antennae 8-segmented, segment III and IV each with a forked sense cone. Pronotum broader than long; surface with transverse striations; with 2 pairs of well-developed posteroangular setae. Mesonotum with widely spaced transverse striations; antero-median campaniform sensilla present; median pair of setae ahead of posterior margin. Metanotum with longitudinal striations, few transverse lines annntomedially; median pair of setae slightly close to the anterior margin; the distance between the median pair less than the median and submedian setae; campaniform sensilla absent (
<figureCitation id="72AD64B52A08CA24FE89343CFE00FD15" box="[331,430,619,645]" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 217</figureCitation>
). Forewing first vein 7 basal and 3 discal setae; second vein with 14 setae, clavus with 5 veinal and one discal seta (
<figureCitation id="72AD64B52A08CA24FE0434D8FD87FD39" box="[454,553,655,681]" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 221</figureCitation>
). Abdominal tergite II with 4 lateromarginal setae; abdominal tergite VIII with posteromarginal comb complete (
<figureCitation id="72AD64B52A08CA24FDCE34E4FDC0FD5D" box="[524,622,691,717]" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 220</figureCitation>
). Abdominal sternites without discal setae (
<figureCitation id="72AD64B52A08CA24FB8C34E4FB01FD5D" box="[1102,1199,691,717]" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 218</figureCitation>
); sternite VII with S1 setae away from the posterior margin (
<figureCitation id="72AD64B52A08CA24FD8A3480FD04FD61" box="[584,682,727,753]" captionStart="FIGURES 215221" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[202,1383,189,1919]" captionTargetId="figure-17@25.[196,1392,181,1919]" captionTargetPageId="25" captionText="FIGURES 215221. Thrips roseae, female: (215) Female; (216) Antenna; (217) Meso- and metanotum; (218) Tergite IVVI; (219) Head and pronotum; (220) Tergite VIII; (221) Forewing." pageId="24" pageNumber="297">Fig. 218</figureCitation>
).
</paragraph>
<paragraph id="EA2978302A08CA24FF0534ACFA3DFCCD" blockId="24.[151,1437,259,1330]" pageId="24" pageNumber="297">
<emphasis id="D8E2A4222A08CA24FF0534ACFED0FC85" bold="true" box="[199,382,763,789]" pageId="24" pageNumber="297">Measurements.</emphasis>
(
<typeStatus id="352DC6922A08CA24FE5234ACFE54FC85" box="[400,506,763,789]" pageId="24" pageNumber="297" type="holotype">Holotype</typeStatus>
female in microns). Body length 1589. Head length (width across cheeks) 133 (168); compound eye dorsal length (width) 77 (47); pronotum median length (width) 142 (208). Forewing length 715. Antenna segments IVIII length (width): 28 (28), 65 (27), 45 (21), 42 (22), 33 (18), 48 (19), 6 (7) and 11 (4).
</paragraph>
<paragraph id="EA2978302A08CA24FF053530FED6FC11" blockId="24.[151,1437,259,1330]" box="[199,376,871,897]" pageId="24" pageNumber="297">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A08CA24FF0535DCFDAAFC7E" pageId="24" pageNumber="297" type="materials_examined">
<paragraph id="EA2978302A08CA24FF0535DCFDAAFC7E" blockId="24.[151,1437,259,1330]" pageId="24" pageNumber="297">
<emphasis id="D8E2A4222A08CA24FF0535DCFE22FC35" bold="true" box="[199,396,907,933]" pageId="24" pageNumber="297">Material studied</emphasis>
.
<materialsCitation id="5AFE726D2A08CA24FE5835DCFC47FC5A" collectingDate="2000-05-07" collectorName="Vikas &amp; Reg. No." country="India" elevation="1761" latitude="31.39" location="Solan" longLatPrecision="730" longitude="77.11" pageId="24" pageNumber="297" specimenCount="1" stateProvince="Himachal Pradesh" typeStatus="holotype">
<typeStatus id="352DC6922A08CA24FE5835DCFDACFC35" box="[410,514,907,933]" pageId="24" pageNumber="297" type="holotype">Holotype</typeStatus>
female,
<emphasis id="D8E2A4222A08CA24FDA335DCFD1DFC35" bold="true" box="[609,691,907,933]" pageId="24" pageNumber="297">
<collectingCountry id="928138A02A08CA24FDA335DCFD1DFC35" box="[609,691,907,933]" name="India" pageId="24" pageNumber="297">INDIA</collectingCountry>
</emphasis>
,
<emphasis id="D8E2A4222A08CA24FD0335DCFC33FC35" bold="true" box="[705,925,907,933]" pageId="24" pageNumber="297">
<collectingRegion id="2852B6D22A08CA24FD0335DCFC33FC35" box="[705,925,907,933]" country="India" name="Himachal Pradesh" pageId="24" pageNumber="297">Himachal Pradesh</collectingRegion>
</emphasis>
,
<location id="EF492EEB2A08CA24FC6835DCFC44FC35" LSID="urn:lsid:plazi:treatment:623FC9262A08CA24FF5536C0FAC4FAA2:EF492EEB2A08CA24FC6835DCFC44FC35" box="[938,1002,907,933]" country="India" latitude="31.39" longLatPrecision="730" longitude="77.11" name="Solan" pageId="24" pageNumber="297" stateProvince="Himachal Pradesh">Solan</location>
(
<geoCoordinate id="8FA21EF72A08CA24FC3A35DCFBF8FC36" box="[1016,1110,907,934]" degrees="31.39" direction="north" orientation="latitude" pageId="24" pageNumber="297" precision="555" value="31.39">31.39 N</geoCoordinate>
,
<geoCoordinate id="8FA21EF72A08CA24FBA235DCFB1CFC35" box="[1120,1202,907,933]" degrees="77.11" direction="east" orientation="longitude" pageId="24" pageNumber="297" precision="555" value="77.11">77.11E</geoCoordinate>
,
<quantity id="2D6ED5D52A08CA24FB7E35DCFABDFC35" box="[1212,1299,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.761" pageId="24" pageNumber="297" unit="m" value="1761.0">
<elevation id="61BB9F032A08CA24FB7E35DCFABDFC35" box="[1212,1299,907,933]" metricMagnitude="3" metricUnit="m" metricValue="1.761" pageId="24" pageNumber="297" unit="m" value="1761.0">1761 m</elevation>
</quantity>
), on flowers of rose (
<taxonomicName id="2D9603B32A08CA24FF3735E7FED5FC59" box="[245,379,943,969]" class="Magnoliopsida" family="Rosaceae" genus="Rosa" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="297" phylum="Tracheophyta" rank="species" species="indica">
<emphasis id="D8E2A4222A08CA24FF3735E7FED5FC59" box="[245,379,943,969]" italics="true" pageId="24" pageNumber="297">Rosa indica</emphasis>
</taxonomicName>
),
<date id="9E285EF02A08CA24FE5135F8FE58FC59" box="[403,502,943,969]" pageId="24" pageNumber="297" value="2000-05-07">
<collectingDate id="8E6CA7182A08CA24FE5135F8FE58FC59" box="[403,502,943,969]" pageId="24" pageNumber="297" value="2000-05-07">7.v.2000</collectingDate>
</date>
,
<collectorName id="47631DE62A08CA24FDC335F8FDEAFC59" box="[513,580,943,969]" pageId="24" pageNumber="297">Vikas</collectorName>
, in NZC, ZSI, (
<collectorName id="47631DE62A08CA24FD3135F8FCF4FC5A" box="[755,858,943,970]" pageId="24" pageNumber="297">Reg. No.</collectorName>
15596/H17)
</materialsCitation>
.
<materialsCitation id="5AFE726D2A08CA24FC2E35F8FDAEFC7E" collectingDate="2000-05-07" collectorName="Reg. No." country="India" elevation="1761" latitude="31.39" location="Solan" longLatPrecision="730" longitude="77.11" pageId="24" pageNumber="297" specimenCount="1" specimenCount-female="1" stateProvince="Himachal Pradesh" typeStatus="paratype">
<typeStatus id="352DC6922A08CA24FC2E35F8FBCFFC5A" box="[1004,1121,943,970]" pageId="24" pageNumber="297" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="FC90B3B92A08CA24FBA935F8FB63FC59" box="[1131,1229,943,969]" count="1" pageId="24" pageNumber="297" type="female">1 female</specimenCount>
with same data as holotype (
<collectorName id="47631DE62A08CA24FECA3584FEC0FC7E" box="[264,366,979,1006]" pageId="24" pageNumber="297">Reg. No.</collectorName>
15597/H17)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A08CA24FF0535A0FC89FB81" box="[199,807,1015,1041]" pageId="24" pageNumber="297" type="etymology">
<paragraph id="EA2978302A08CA24FF0535A0FC89FB81" blockId="24.[151,1437,259,1330]" box="[199,807,1015,1041]" pageId="24" pageNumber="297">
<emphasis id="D8E2A4222A08CA24FF0535A0FEFCFB81" bold="true" box="[199,338,1015,1041]" pageId="24" pageNumber="297">Etymology:</emphasis>
The species is named after the host plant.
</paragraph>
</subSubSection>
<subSubSection id="A28C2BBB2A08CA24FF05324CFAC4FAA2" pageId="24" pageNumber="297" type="discussion">
<paragraph id="EA2978302A08CA24FF05324CFAC4FAA2" blockId="24.[151,1437,259,1330]" pageId="24" pageNumber="297">
<emphasis id="D8E2A4222A08CA24FF05324CFEFCFBA5" bold="true" box="[199,338,1051,1077]" pageId="24" pageNumber="297">Comments:</emphasis>
<taxonomicName id="2D9603B32A08CA24FE9A324CFE4BFBA5" authorityName="Tyagi &amp; Pal &amp; Sarma &amp; Kumar" authorityYear="2025" box="[344,485,1051,1077]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="24" pageNumber="297" phylum="Arthropoda" rank="species" species="rosea" status="sp. nov.">
<emphasis id="D8E2A4222A08CA24FE9A324CFE4BFBA5" box="[344,485,1051,1077]" italics="true" pageId="24" pageNumber="297">Thrips rosea</emphasis>
</taxonomicName>
<emphasis id="D8E2A4222A08CA24FE2E324CFDE8FBA5" bold="true" box="[492,582,1051,1077]" pageId="24" pageNumber="297">
<taxonomicNameLabel id="C3D119592A08CA24FE2E324CFDE8FBA5" box="[492,582,1051,1077]" pageId="24" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="2D9603B32A08CA24FD15324CFB8AFBA5" authority="(Pelikan)" baseAuthorityName="Pelikan" baseAuthorityYear="1970" box="[727,1060,1051,1077]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="24" pageNumber="297" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis id="D8E2A4222A08CA24FD15324CFC18FBA5" box="[727,950,1051,1077]" italics="true" pageId="24" pageNumber="297">Thrips himalayanus</emphasis>
(Pelikan)
</taxonomicName>
but it can be differentiated by the presence of transverse striations on pronotal surface, forewings weakly shaded at base, antennal segments IV and basal ¼ of V yellow. In contrast, the pronotal surface smooth, forewing completely shaded, antennae dark except segment III in
<taxonomicName id="2D9603B32A08CA24FEF532DFFE4CFB31" baseAuthorityName="Pelikan" baseAuthorityYear="1970" box="[311,482,1159,1186]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="24" pageNumber="297" phylum="Arthropoda" rank="species" species="himalayanus">
<emphasis id="D8E2A4222A08CA24FEF532DFFEE9FB31" box="[311,327,1160,1185]" italics="true" pageId="24" pageNumber="297">T</emphasis>
.
<emphasis id="D8E2A4222A08CA24FE9132D0FE4CFB31" box="[339,482,1159,1185]" italics="true" pageId="24" pageNumber="297">himalayanus</emphasis>
</taxonomicName>
. This new species is very similar to
<taxonomicName id="2D9603B32A08CA24FCAD32DFFAC0FB31" authority="Masumoto &amp; Okajima (2013)" authorityName="Masumoto &amp; Okajima" authorityYear="2013" box="[879,1390,1159,1186]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="24" pageNumber="297" phylum="Arthropoda" rank="species" species="shiranesanus">
<emphasis id="D8E2A4222A08CA24FCAD32DFFCD1FB31" box="[879,895,1160,1185]" italics="true" pageId="24" pageNumber="297">T</emphasis>
.
<emphasis id="D8E2A4222A08CA24FC4932D0FBB1FB31" box="[907,1055,1159,1185]" italics="true" pageId="24" pageNumber="297">shiranesanus</emphasis>
<bibRefCitation id="8E0705C12A08CA24FBE732DFFAC0FB31" author="Masumoto, M. &amp; Okajima, S." box="[1061,1390,1159,1186]" pageId="24" pageNumber="297" pagination="1 - 65" refId="ref18954" refString="Masumoto, M. &amp; Okajima, S. (2013) Review of the genus Thrips and related genera (Thysanoptera, Thripidae) from Japan. Zootaxa, 3678 (1), 1 - 65. https: // doi. org / 10.11646 / zootaxa. 3678.1.1" type="journal article" year="2013">Masumoto &amp; Okajima (2013)</bibRefCitation>
</taxonomicName>
and can be differentiated by the presence of 8 segmented antennae, metanotum without campaniform sensilla, abdominal sternite VII with 3 pairs of setae and tergites III to VII without small scallops along posterior margin. In contrast, antennae 7-segmented, metanotum with campaniform sensilla, abdominal sternite VII with 2 pairs of setae on one side and 3 pairs on other side, tergites III to VII with small scallops along posterior margin in
<taxonomicName id="2D9603B32A08CA24FB70334FFACDFAA1" authorityName="Masumoto &amp; Okajima" authorityYear="2013" box="[1202,1379,1303,1330]" class="Insecta" family="Thripidae" genus="Thrips" kingdom="Animalia" order="Thysanoptera" pageId="24" pageNumber="297" phylum="Arthropoda" rank="species" species="shiranesanus">
<emphasis id="D8E2A4222A08CA24FB70334FFB6CFAA1" box="[1202,1218,1304,1329]" italics="true" pageId="24" pageNumber="297">T</emphasis>
.
<emphasis id="D8E2A4222A08CA24FB0D3340FACDFAA1" box="[1231,1379,1303,1329]" italics="true" pageId="24" pageNumber="297">shiranesanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>