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@ -311,7 +313,7 @@ locality and a closely adjacent small outcrop on a logging road in the lower mem
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@ -111,7 +113,7 @@ A second Gries Ranch glycymeridid species, described subsequently by
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<emphasis id="78E4EAD9FFE8292CFCC0F9E1EA5FBB30" bold="true" box="[863,1115,1605,1631]" pageId="16" pageNumber="15">Material examined</emphasis>
—
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from Loc.
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@ -125,28 +127,28 @@ from Loc.
information
</emphasis>
—Figured hypotypes, all from Loc.
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hypotype)
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Formation
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) (Kavanov et al. 2000, 2001).
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Although the genus is absent from the tropical Paleogene faunas of the
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, it apparently was present at much higher latitudes in cold water during the Late Cretaceous and earliest Paleogene. Marincovich (1993) described

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this species also occurs stratigraphically higher in the overlying Pittsburg Bluff Formation and Miocene Astoria Formation. In
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it occurs in the
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<emphasis id="78A9EAA4FF84FFF0B6FE60C26E28DB17" bold="true" box="[199,345,1518,1544]" pageId="8" pageNumber="120">Description.</emphasis>
Morphometric data in
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. Medium-sized loricariid with maximum standard length up to
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SL. Body robust, depressed anteriorly and compressed posteriorly (
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<emphasis id="78A9EAA4FF85FFF1B6A165B46E7EDDAF" bold="true" box="[152,271,152,176]" pageId="9" pageNumber="121">TABLE 2.</emphasis>
Morphometric data of

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<mods:affiliation id="19408977EFA9BB7C08DA69C43E0C5580">Departamento de Biología, Facultad de Ciencias, Escuela Politécnica Nacional, Quito, Ecuador</mods:affiliation>
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<mods:role id="A6F29526600CC199381790A66146CF6F">
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<mods:namePart id="5086B139F4CEBB692177F6BF286C20C4">Stewart, Donald J.</mods:namePart>
<mods:affiliation id="749AE92F320EAD833602B4942EF60DF5">Department of Environmental Biology, SUNY College of Environmental Science and Forestry, Syracuse, New York, 13210, USA</mods:affiliation>
<mods:namePart id="886929CEB5C836D459EFFC383A6DE2D4">Stewart, Donald J.</mods:namePart>
<mods:affiliation id="A97ACA0759344963660E68B09D951464">Department of Environmental Biology, SUNY College of Environmental Science and Forestry, Syracuse, New York, 13210, USA</mods:affiliation>
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<mods:date id="D39A3477596A75BF2E7CCC3E87C72AAB">2024</mods:date>
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<treatment id="C27487A0FF8EFFF0B6AE644A6DECDCEB" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB" httpUri="http://treatment.plazi.org/id/C27487A0FF8EFFF0B6AE644A6DECDCEB" lastPageId="8" lastPageNumber="120" pageId="2" pageNumber="114">
<treatment id="C27487A0FF8EFFF0B6AE644A6DECDCEB" ID-DOI="http://doi.org/10.5281/zenodo.11645869" ID-GBIF-Taxon="232750432" ID-Zenodo-Dep="11645869" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB" httpUri="http://treatment.plazi.org/id/C27487A0FF8EFFF0B6AE644A6DECDCEB" lastPageId="8" lastPageNumber="120" pageId="2" pageNumber="114">
<subSubSection id="02C7653DFF8EFFFAB6AE644A6CA6DCBA" pageId="2" pageNumber="114" type="nomenclature">
<paragraph id="4A6236B6FF8EFFFAB6AE644A6CA6DCBA" blockId="2.[151,983,358,457]" pageId="2" pageNumber="114">
<heading id="112A81DAFF8EFFFAB6AE644A6D74DC9F" bold="true" box="[151,517,358,385]" fontSize="11" level="1" pageId="2" pageNumber="114" reason="1">
<emphasis id="78A9EAA4FF8EFFFAB6AE644A6D74DC9F" bold="true" box="[151,517,358,385]" pageId="2" pageNumber="114">
<taxonomicName id="8DDD4D35FF8EFFFAB6AE644A6E16DC9E" authority="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart, 2024" authorityName="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart" authorityYear="2024" box="[151,359,358,385]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="114" phylum="Chordata" rank="species" species="orcesi" status="sp. nov.">
<emphasis id="78A9EAA4FF8EFFFAB6AE644A6E16DC9E" bold="true" box="[151,359,358,385]" italics="true" pageId="2" pageNumber="114">Panaqolus orcesi</emphasis>
</taxonomicName>
<taxonomicName id="8DDD4D35FF8EFFFAB6AE644A6E16DC9E" authority="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart, 2024" authorityName="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart" authorityYear="2024" box="[151,359,358,385]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="114" phylum="Chordata" rank="species" species="orcesi" status="sp. nov.">Panaqolus orcesi</taxonomicName>
,
<taxonomicNameLabel id="639A57DFFF8EFFFAB74F644B6D74DC9F" box="[374,517,358,385]" pageId="2" pageNumber="114" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
<uri id="3E4C3AB4FF8EFFFAB6AE64A66CA6DCBA" box="[151,983,394,421]" pageId="2" pageNumber="114">
urn:lsid:zoobank.org:act:
@ -82,13 +80,13 @@ urn:lsid:zoobank.org:act:
and
<figureCitation id="D2E62A33FF8EFFFAB77564836E18DCD6" box="[332,361,431,457]" captionStart="FIGURE 3" captionStartId="5.[152,255,1968,1992]" captionTargetBox="[164,1423,964,1945]" captionTargetId="figure-85@5.[164,1423,964,1945]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of color patterns in specimens soon after capture for: a) Panaqolus orcesi, with cream-colored dots, and b) Panaqolus pantostiktos, with yellow- or gold-colored dots. Photos by R.B-S." figureDoi="http://doi.org/10.5281/zenodo.11610312" httpUri="https://zenodo.org/record/11610312/files/figure.png" pageId="2" pageNumber="114">3a</figureCitation>
;
<tableCitation id="075F030DFF8EFFFAB74D64836EC9DCD6" box="[372,440,431,457]" captionStart="TABLE 1" captionStartId="2.[152,244,828,852]" captionText="TABLE 1. Morphometric data of Panaqolus orcesi, expressed as percent of SL or percent of HL." pageId="2" pageNumber="114">Tab. 1</tableCitation>
<tableCitation id="075F030DFF8EFFFAB74D64836EC9DCD6" box="[372,440,431,457]" captionStart="TABLE 1" captionStartId="2.[152,244,828,852]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data of Panaqolus orcesi, expressed as percent of SL or percent of HL." httpUri="http://table.plazi.org/id/1EA2663EFF8EFFFAB6A166106BDBDE4B" pageId="2" pageNumber="114" tableUuid="1EA2663EFF8EFFFAB6A166106BDBDE4B">Tab. 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="02C7653DFF8EFFFAB6AE64DB6D7FDE12" pageId="2" pageNumber="114" type="materials_examined">
<paragraph id="4A6236B6FF8EFFFAB6AE64DB6EAFDF46" blockId="2.[151,1437,503,781]" pageId="2" pageNumber="114">
<materialsCitation id="FAB53CEBFF8EFFFAB6AE64DB6EAFDF46" collectingDate="2006-06-26" collectionCode="MEPN-I, R" county="Ecuadorian Amazon" latitude="-0.83416665" location="Quebrada La Cascada" longLatPrecision="21" longitude="-75.56612" municipality="River" pageId="2" pageNumber="114" specimenCount="1" stateProvince="Orellana Province, Napo" typeStatus="holotype">
<materialsCitation id="FAB53CEBFF8EFFFAB6AE64DB6EAFDF46" ID-GBIF-Occurrence="4876552304" collectingDate="2006-06-26" collectionCode="MEPN-I" collectorName="R. Barriga" county="Ecuadorian Amazon" latitude="-0.83416665" location="Napo River basin" longLatPrecision="21" longitude="-75.56612" municipality="River" pageId="2" pageNumber="114" specimenCount="1" stateProvince="Orellana Province" typeStatus="holotype">
<emphasis id="78A9EAA4FF8EFFFAB6AE64DB6E7ADF0E" bold="true" box="[151,267,503,529]" pageId="2" pageNumber="114">
<typeStatus id="95668814FF8EFFFAB6AE64DB6E77DF0E" box="[151,262,503,529]" pageId="2" pageNumber="114" type="holotype">Holotype</typeStatus>
.
@ -98,12 +96,13 @@ and
SL,
<collectingCounty id="A3034E3AFF8EFFFAB44B64DB6C29DF0E" box="[626,856,503,529]" pageId="2" pageNumber="114">Ecuadorian Amazon</collectingCounty>
,
<collectingRegion id="8819F854FF8EFFFAB55A64DB6B09DF0E" box="[867,1144,503,529]" isEnumeration="true" pageId="2" pageNumber="114">Orellana Province, Napo</collectingRegion>
<collectingMunicipality id="AA06ACCCFF8EFFFAB24764DB6BCDDF0E" box="[1150,1212,503,529]" pageId="2" pageNumber="114">River</collectingMunicipality>
basin,
<location id="4F02606DFF8EFFFAB33264DB6F86DF2A" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB33264DB6F86DF2A" county="Ecuadorian Amazon" latitude="-0.83416665" longLatPrecision="21" longitude="-75.56612" municipality="River" name="Quebrada La Cascada" pageId="2" pageNumber="114" stateProvince="Orellana Province, Napo">Quebrada La Cascada</location>
<collectingRegion id="8819F854FF8EFFFAB55A64DB6B41DF0E" box="[867,1072,503,529]" pageId="2" pageNumber="114">Orellana Province</collectingRegion>
,
<location id="4F02606DFF8EFFFAB73A67376ED3DF2A" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB73A67376ED3DF2A" box="[259,418,539,565]" county="Ecuadorian Amazon" latitude="-0.83416665" longLatPrecision="21" longitude="-75.56612" municipality="River" name="Tiputini River" pageId="2" pageNumber="114" stateProvince="Orellana Province, Napo">Tiputini River</location>
<location id="4F02606DFF8EFFFAB20264DB6A70DF0E" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB20264DB6A70DF0E" box="[1083,1281,503,529]" county="Ecuadorian Amazon" latitude="-0.83416665" longLatPrecision="21" longitude="-75.56612" municipality="River" name="Napo River basin" pageId="2" pageNumber="114" stateProvince="Orellana Province">Napo River basin</location>
,
<location id="4F02606DFF8EFFFAB33264DB6F86DF2A" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB33264DB6F86DF2A" county="Ecuadorian Amazon" latitude="-0.83416665" longLatPrecision="21" longitude="-75.56612" municipality="River" name="Quebrada La Cascada" pageId="2" pageNumber="114" stateProvince="Orellana Province">Quebrada La Cascada</location>
,
<location id="4F02606DFF8EFFFAB73A67376ED3DF2A" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB73A67376ED3DF2A" box="[259,418,539,565]" county="Ecuadorian Amazon" latitude="-0.83416665" longLatPrecision="21" longitude="-75.56612" municipality="River" name="Tiputini River" pageId="2" pageNumber="114" stateProvince="Orellana Province">Tiputini River</location>
tributary, approx.
<geoCoordinate id="2FE95071FF8EFFFAB44C67366C72DF2A" box="[629,771,538,565]" degrees="00" direction="south" minutes="50" orientation="latitude" pageId="2" pageNumber="114" precision="15" seconds="03" value="-0.83416665">00°50’03” S</geoCoordinate>
,
@ -113,12 +112,12 @@ tributary, approx.
<collectingDate id="2E27E99EFF8EFFFAB37C67376FA3DF46" pageId="2" pageNumber="114" value="2006-06-26">26 June 2006</collectingDate>
</date>
,
<collectionCode id="2CCCAE73FF8EFFFAB6E467136F82DF46" box="[221,243,575,601]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="114">R</collectionCode>
. Barriga, RBS06-12.
<collectorName id="E7285360FF8EFFFAB6E467136E24DF46" box="[221,341,575,601]" pageId="2" pageNumber="114">R. Barriga</collectorName>
, RBS06-12.
</materialsCitation>
</paragraph>
<paragraph id="4A6236B6FF8EFFFAB6FE674F6D7FDE12" blockId="2.[151,1437,503,781]" pageId="2" pageNumber="114">
<materialsCitation id="FAB53CEBFF8EFFFAB6FE674F6D7BDE12" collectingDate="2005-04-20" collectingDateMax="2007-06-17" collectingDateMin="2005-04-20" collectionCode="MEPN-I, R, T" collectorName="J. Nenhimo &amp; Barriga &amp; J. Silva" county="Ecuadorian Amazon" elevation="100" latitude="-0.62222224" location="Rio Mengo Garita" longLatPrecision="21" longitude="-76.50694" municipality="River" pageId="2" pageNumber="114" specimenCount="2" stateProvince="Orellana Province, Napo" typeStatus="paratype">
<materialsCitation id="FAB53CEBFF8EFFFAB6FE674F6C91DFDA" ID-GBIF-Occurrence="4876552302" collectingDate="2005-04-20" collectingDateMax="2005-04-22" collectingDateMin="2005-04-20" collectionCode="MEPN-I" collectorName="R. Barriga &amp; T. Caiza &amp; J. Nenhimo" county="Ecuadorian Amazon" elevation="100" latitude="-0.62222224" location="Rio Mengo Garita" longLatPrecision="21" longitude="-76.50694" municipality="River" pageId="2" pageNumber="114" specimenCount="2" stateProvince="Orellana Province, Napo" typeStatus="paratype">
<emphasis id="78A9EAA4FF8EFFFAB6FE674F6E37DF62" bold="true" box="[199,326,611,637]" pageId="2" pageNumber="114">
<typeStatus id="95668814FF8EFFFAB6FE674F6E30DF62" box="[199,321,611,637]" pageId="2" pageNumber="114" type="paratype">Paratypes</typeStatus>
.
@ -128,62 +127,44 @@ from
<collectingCounty id="A3034E3AFF8EFFFAB47B674F6C5BDF62" box="[578,810,611,637]" pageId="2" pageNumber="114">Ecuadorian Amazon</collectingCounty>
,
<collectingRegion id="8819F854FF8EFFFAB50F674F6B3FDF62" box="[822,1102,611,637]" isEnumeration="true" pageId="2" pageNumber="114">Orellana Province, Napo</collectingRegion>
<collectingMunicipality id="AA06ACCCFF8EFFFAB26F674F6BE5DF62" box="[1110,1172,611,637]" pageId="2" pageNumber="114">River</collectingMunicipality>
basin:
River basin:
<collectionCode id="2CCCAE73FF8EFFFAB2DE674F6A36DF62" box="[1255,1351,611,637]" pageId="2" pageNumber="114">MEPN-I</collectionCode>
<quantity id="8D259B53FF8EFFFAB376674F6E79DFBE" metricMagnitude="4" metricUnit="m" metricValue="1.9493170899999999" pageId="2" pageNumber="114" unit="mm" value="1.94931709E7">19493, 170.9 mm</quantity>
SL,
19493, 170.9 mm SL,
<location id="4F02606DFF8EFFFAB70267AB6D79DFBE" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB70267AB6D79DFBE" box="[315,520,647,673]" county="Ecuadorian Amazon" latitude="-0.62222224" longLatPrecision="21" longitude="-76.50694" municipality="River" name="Rio Mengo Garita" pageId="2" pageNumber="114" stateProvince="Orellana Province, Napo">Rio Mengo Garita</location>
,
<quantity id="8D259B53FF8EFFFAB42B67AB6D29DFBE" box="[530,600,647,673]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="114" unit="m" value="100.0">
<elevation id="C1F0D185FF8EFFFAB42B67AB6D29DFBE" box="[530,600,647,673]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="114" unit="m" value="100.0">100 m</elevation>
</quantity>
upstream from bridge on
, 100 m upstream from bridge on
<location id="4F02606DFF8EFFFAB54D67AB6B76DFBE" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB54D67AB6B76DFBE" box="[884,1031,647,673]" county="Ecuadorian Amazon" latitude="-0.62222224" longLatPrecision="21" longitude="-76.50694" municipality="River" name="Yasuni River" pageId="2" pageNumber="114" stateProvince="Orellana Province, Napo">Yasuni River</location>
, approx.
<geoCoordinate id="2FE95071FF8EFFFAB25567AA6B86DFBE" box="[1132,1271,646,673]" degrees="00" direction="south" minutes="50" orientation="latitude" pageId="2" pageNumber="114" precision="15" seconds="03" value="-0.83416665">00°50’03” S</geoCoordinate>
,
<geoCoordinate id="2FE95071FF8EFFFAB33A67AB6AE9DFBE" box="[1283,1432,647,673]" degrees="76" direction="west" minutes="20" orientation="longitude" pageId="2" pageNumber="114" precision="15" seconds="42" value="-76.345">76°20’42” W</geoCoordinate>
,
<quantity id="8D259B53FF8EFFFAB6AE67866F8EDFDA" box="[151,255,682,709]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="114" unit="m" value="250.0">
<elevation id="C1F0D185FF8EFFFAB6AE67866F8EDFDA" box="[151,255,682,709]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="114" unit="m" value="250.0">250 masl</elevation>
</quantity>
,
, 250 masl,
<date id="3E631076FF8EFFFAB73067876EBEDFDA" box="[265,463,682,709]" pageId="2" pageNumber="114" value="2005-04-20" valueMax="2005-04-22" valueMin="2005-04-20">
<collectingDate id="2E27E99EFF8EFFFAB73067876EBEDFDA" box="[265,463,682,709]" pageId="2" pageNumber="114" value="2005-04-20" valueMax="2005-04-22" valueMin="2005-04-20">20–22 April 2005</collectingDate>
</date>
,
<collectionCode id="2CCCAE73FF8EFFFAB7E067876E9EDFDA" box="[473,495,683,709]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="114">R</collectionCode>
. Barriga,
<collectionCode id="2CCCAE73FF8EFFFAB46067876D1ADFDA" box="[601,619,683,709]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="114">T</collectionCode>
. Caiza &amp;
<collectorName id="E7285360FF8EFFFAB7E067876D3EDFDA" box="[473,591,683,709]" pageId="2" pageNumber="114">R. Barriga</collectorName>
,
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<emphasis id="78A9EAA4FF8EFFFAB6A166106E7EDE4B" bold="true" box="[152,271,828,852]" pageId="2" pageNumber="114">TABLE 1.</emphasis>
Morphometric data of
@ -497,7 +478,7 @@ and
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<emphasis id="78A9EAA4FF8FFFFBB6FE66146E28DE4D" bold="true" box="[199,345,824,850]" pageId="3" pageNumber="115">Description.</emphasis>
Morphometric data in
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SL. Body robust, quite depressed anteriorly and compressed posteriorly (

64
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