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<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
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<mods:title id="0677EAC888D75F6B0E7741DD680F69C3">Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
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<mods:namePart>Santos, Daubian</mods:namePart>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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<mods:namePart id="E98ECA2ACE62D4D07CB2B09B6375A0F5">Santos, Daubian</mods:namePart>
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<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
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<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
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<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
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3.3.12.
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<taxonomicName LSID="D282571B-273A-5A29-8B1A-6AA1EF780682" authority="Alexander, 1929 a, nomen dubium" authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) fuscifusus Alexander, 1929a, nomen dubium</taxonomicName>
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<taxonomicName id="539A5141D577F22DE0C3BB3A28D36E5F" ID-CoL="D436" LSID="D282571B-273A-5A29-8B1A-6AA1EF780682" authority="Alexander, 1929 a, nomen dubium" authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) fuscifusus Alexander, 1929a, nomen dubium</taxonomicName>
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<paragraph pageId="0" pageNumber="439">
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<taxonomicName authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) fuscifusus</taxonomicName>
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<subSubSection id="5585D7A64BA0B2EE7E76A1A13AC2249A" pageId="0" pageNumber="439" type="reference_group">
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<paragraph id="4CB48886978CFB8E1EC34B6CA1CD609C" pageId="0" pageNumber="439">
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<taxonomicName id="281D28D6558719D2659B4135FD482AF8" authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) fuscifusus</taxonomicName>
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Alexander, 1929a: 190 (description).
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</paragraph>
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<subSubSection pageId="0" pageNumber="439" type="remarks">
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<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="439">
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<taxonomicName authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (R.) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="R.">Amphineurus (R.) fuscifusus</taxonomicName>
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<subSubSection id="1B25FCB9C822D4FCCDF5876134D8F2A5" pageId="0" pageNumber="439" type="remarks">
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<paragraph id="455284C576DAC29CF0A5EB3949D7C3F6" pageId="0" pageNumber="439">Remarks.</paragraph>
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<paragraph id="DA63D9E6F97A737EF9F24B95FDCBBB95" pageId="0" pageNumber="439">
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<taxonomicName id="84D733A3E51954A806F271937CDD59FD" authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (R.) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="R.">Amphineurus (R.) fuscifusus</taxonomicName>
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Alexander is the only species of the genus in which the male is unknown. Additionally, the only identified specimen is the female holotype, currently housed in the Natural History Museum (BMNH). The absence of an associated male specimen and its terminalia is problematic because this is an important source of characters and taxonomic data. Lacking a well-defined diagnosis in the description, the erection of the new species was proposed mostly due to subtle differences in coloration. The only considerable difference in coloration in
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<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) fuscifusus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="fuscifusus" subGenus="R.">A. (R.) fuscifusus</taxonomicName>
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<taxonomicName id="3F30DD9D7B9C84B10A71B45271736744" class="Insecta" family="Limoniidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. (R.) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="R.">A. (R.) fuscifusus</taxonomicName>
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||||
is the pale halter, which is unlike the yellowish brown halter of
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<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) nothofagetorum" order="Diptera" pageId="0" pageNumber="439" rank="species" species="nothofagetorum" subGenus="R.">A. (R.) nothofagetorum</taxonomicName>
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||||
<taxonomicName id="A6B1F440B3420B27E375CAE0C4C2A06E" class="Insecta" family="Limoniidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. (R.) nothofagetorum" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nothofagetorum" subGenus="R.">A. (R.) nothofagetorum</taxonomicName>
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||||
Alexander.
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</paragraph>
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<paragraph pageId="0" pageNumber="439">
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<paragraph id="B0FF68F23CB11974946A76A0DD3350F1" pageId="0" pageNumber="439">
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Some morphological data described by
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<bibRefCitation author="Alexander, CP" journalOrPublisher="Diptera of Patagonia and South Chile" pageId="0" pageNumber="439" pagination="1 - 240" refId="B7" refString="Alexander, CP, 1929a. Diptera of Patagonia and South Chile. Part I. Crane-flies (Tipulidae, Trichoceridae, Tanyderidae). Diptera of Patagonia and South Chile 1: 1 - 240" title="Diptera of Patagonia and South Chile. Part I. Crane-flies (Tipulidae, Trichoceridae, Tanyderidae)." volume="1" year="1929 a">Alexander (1929a)</bibRefCitation>
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||||
<bibRefCitation id="EBA67B03B442D74CEED18B96A8A44EBF" author="Alexander, CP" journalOrPublisher="Diptera of Patagonia and South Chile" pageId="0" pageNumber="439" pagination="1 - 240" refId="B7" refString="Alexander, CP, 1929a. Diptera of Patagonia and South Chile. Part I. Crane-flies (Tipulidae, Trichoceridae, Tanyderidae). Diptera of Patagonia and South Chile 1: 1 - 240" title="Diptera of Patagonia and South Chile. Part I. Crane-flies (Tipulidae, Trichoceridae, Tanyderidae)." volume="1" year="1929 a">Alexander (1929a)</bibRefCitation>
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suggest this specimen shares similar features with clade 33 (Fig.
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<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Most parsimonious tree (part) yielded by parsimony analysis with implied weighting (K = 3), depicting recovered relationships among outgroup taxa. Numbers at tree branches designate nodes (see 4.2.). - Abbreviations: Aus, Australia; NC, New Caledonia; NG, New Guinea; NZ, New Zealand, SAm, South America; Tas, Tasmania." figureDoi="10.3897/asp.80.e83035.figure32" httpUri="https://binary.pensoft.net/fig/740585" pageId="0" pageNumber="439">32</figureCitation>
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||||
<figureCitation id="7E5E7CE67B3846902C7DA9216A262E30" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Most parsimonious tree (part) yielded by parsimony analysis with implied weighting (K = 3), depicting recovered relationships among outgroup taxa. Numbers at tree branches designate nodes (see 4.2.). - Abbreviations: Aus, Australia; NC, New Caledonia; NG, New Guinea; NZ, New Zealand, SAm, South America; Tas, Tasmania." figureDoi="10.3897/asp.80.e83035.figure32" httpUri="https://binary.pensoft.net/fig/740585" pageId="0" pageNumber="439">32</figureCitation>
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||||
). The presence of long setae on the antenna, "weak branches" in medial veins, three stripes on the scutum, and a marking near the fork of Rs are features shared between the holotype and this group. The holotype was collected in the same location as the holotype of
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||||
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) nothofagetorum" order="Diptera" pageId="0" pageNumber="439" rank="species" species="nothofagetorum" subGenus="R.">A. (R.) nothofagetorum</taxonomicName>
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||||
<taxonomicName id="34C90F496A4B57F0F00A79B08FD29404" class="Insecta" family="Limoniidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. (R.) nothofagetorum" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nothofagetorum" subGenus="R.">A. (R.) nothofagetorum</taxonomicName>
|
||||
Alexander. However, the absence of males added to the fact that there is only a single known specimen makes the synonymization or identity confirmation of the species problematic. For these reasons, we define
|
||||
<taxonomicName authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) fuscifusus</taxonomicName>
|
||||
<taxonomicName id="D909FAE0DE8AA7D3F359C54D91071D53" authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) fuscifusus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="fuscifusus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) fuscifusus</taxonomicName>
|
||||
Alexander as nomen dubium.
|
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</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
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