Add updates up until 2024-07-04 07:50:13

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<mods:title id="0EC1A37EBDC6CE72AF76EF6D705EA285">Taxonomic Assessment of Acetes indicus H. Milne Edwards, 1830 (Crustacea, Decapoda, Sergestoidea) as Revealed from Molecular and Morphological Analyses: Re-validation of A. spiniger Hansen, 1919 and Designation of a New Species</mods:title>
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<treatment id="830A4D479879FFB8C773FCF25DABFCCE" ID-DOI="http://doi.org/10.5281/zenodo.12572404" ID-GBIF-Taxon="233810457" ID-Zenodo-Dep="12572404" LSID="urn:lsid:plazi:treatment:830A4D479879FFB8C773FCF25DABFCCE" httpUri="http://treatment.plazi.org/id/830A4D479879FFB8C773FCF25DABFCCE" lastPageId="17" lastPageNumber="66" pageId="14" pageNumber="63">
<subSubSection id="43B9AFDA9879FFA7C773FCF25B56FC9E" box="[811,1131,818,842]" pageId="14" pageNumber="63" type="nomenclature">
<paragraph id="0B1CFC519879FFA7C773FCF25B56FC9E" blockId="14.[811,1131,818,842]" box="[811,1131,818,842]" pageId="14" pageNumber="63">
<heading id="50544B3D9879FFA7C773FCF25B56FC9E" box="[811,1131,818,842]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="63" reason="4">
<taxonomicName id="CCA387D29879FFA7C773FCF25B56FC9E" ID-CoL="9XFDD" authority="Hansen, 1919" authorityName="Hansen" authorityYear="1919" box="[811,1131,818,842]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="39D720439879FFA7C773FCF25CEFFC9E" bold="true" box="[811,978,818,842]" italics="true" pageId="14" pageNumber="63">Acetes spiniger</emphasis>
<bibRefCitation id="6F3281A09879FFA7C781FCF25B56FC9E" author="Hansen, H. J." box="[985,1131,818,842]" pageId="14" pageNumber="63" pagination="1 - 65" refId="ref10693" refString="Hansen, H. J. 1919. The Sergestidae of the Siboga-Expedition. Siboga Expeditie 38: 1 - 65, pls 1 - 5." type="journal article" year="1919">Hansen, 1919</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA9879FFA7C7F1FCA15CD0FCA3" box="[937,1005,865,887]" pageId="14" pageNumber="63" type="description">
<paragraph id="0B1CFC519879FFA7C7F1FCA15CD0FCA3" blockId="14.[937,1005,865,887]" box="[937,1005,865,887]" pageId="14" pageNumber="63">
(
<figureCitation id="9398E0D49879FFA7C7E8FCA15CDBFCA3" box="[944,998,865,887]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="14" pageNumber="63">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA9879FFA7C69BFC515B8AFBC8" pageId="14" pageNumber="63" type="reference_group">
<paragraph id="0B1CFC519879FFA7C69BFC515B5FFC73" blockId="14.[707,1237,913,1052]" box="[707,1122,913,935]" pageId="14" pageNumber="63">
<treatmentCitationGroup id="2BB3DB7F9879FFA7C69BFC515B5FFC73" box="[707,1122,913,935]" pageId="14" pageNumber="63">
<taxonomicName id="CCA387D29879FFA7C69BFC515CEEFC72" ID-CoL="9XFDD" authority="Hansen, 1919" authorityName="Hansen" authorityYear="1919" box="[707,979,913,935]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="39D720439879FFA7C69BFC515C73FC73" box="[707,846,913,935]" italics="true" pageId="14" pageNumber="63">Acetes spiniger</emphasis>
<bibRefCitation id="6F3281A09879FFA7C70CFC525CEEFC72" author="Hansen, H. J." box="[852,979,913,935]" pageId="14" pageNumber="63" pagination="1 - 65" refId="ref10693" refString="Hansen, H. J. 1919. The Sergestidae of the Siboga-Expedition. Siboga Expeditie 38: 1 - 65, pls 1 - 5." type="journal article" year="1919">Hansen, 1919</bibRefCitation>
</taxonomicName>
; pl. 4, fig. 5af.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0B1CFC519879FFA7C69BFC6F5B96FC2B" blockId="14.[707,1237,913,1052]" pageId="14" pageNumber="63">
<treatmentCitationGroup id="2BB3DB7F9879FFA7C69BFC6F5BE8FC10" box="[707,1237,942,964]" pageId="14" pageNumber="63">
<taxonomicName id="CCA387D29879FFA7C69BFC6F5C75FC17" ID-CoL="9XFD2" authorityName="H. Milne Edwards" authorityYear="1830" box="[707,840,942,964]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D720439879FFA7C69BFC6F5C75FC17" box="[707,840,942,964]" italics="true" pageId="14" pageNumber="63">Acetes indicus</emphasis>
</taxonomicName>
:
<treatmentCitation id="8A02DA409879FFA7C700FC6F5CCCFC10" author="Kemp, S." box="[856,1009,943,964]" page="47" pageId="14" pageNumber="63" year="1917">
<bibRefCitation id="6F3281A09879FFA7C700FC6F5CCCFC10" author="Kemp, S." box="[856,1009,943,964]" pageId="14" pageNumber="63" pagination="43 - 58" refId="ref11017" refString="Kemp, S. 1917. Notes on Crustacea Decapoda in the Indian Museum. The genus Acetes Milne-Edwards. Records of the Indian Museum 53: 43 - 58." type="journal article" year="1917">Kemp, 1917: 47</bibRefCitation>
</treatmentCitation>
(in part);
<bibRefCitation id="6F3281A09879FFA7C00EFC6E5BECFC10" author="Omori, M." box="[1110,1233,942,964]" pageId="14" pageNumber="63" pagination="1 - 89" refId="ref11166" refString="Omori, M. 1975. The systematics, biogeography, and fish- ery of epipelagic shrimps of the genus Acetes (Crustacea, Decapoda, Sergestidae). Bulletin of the Ocean Research Institute, University of Tokyo 7: 1 - 89." type="journal article" year="1975">
<collectingRegion id="C96732B39879FFA7C00EFC6E5BA9FC10" box="[1110,1172,942,964]" country="Japan" name="Aomori" pageId="14" pageNumber="63">Omori</collectingRegion>
, 1975
</bibRefCitation>
:
</treatmentCitationGroup>
(in part; at least fig. 15a, b, d, f, gi);
<bibRefCitation id="6F3281A09879FFA7C00CFC0C5BE8FC36" author="Ravindranath, K." box="[1108,1237,972,994]" pageId="14" pageNumber="63" pagination="253 - 273" refId="ref11394" refString="Ravindranath, K. 1980. Shrimps of the genus Acetes H. Milne Edwards (Crustacea, Decapoda, Sergestidae) from the estuarine system of river Krishna. Proceedings of the Indian Academy of Sciences (Animal Sciences) 89: 253 - 273." type="journal article" year="1980">Ravindranath,</bibRefCitation>
1980: 255, figs 13;
<bibRefCitation id="6F3281A09879FFA7C7C0FC295B9AFC2B" author="Pathan, D. I. &amp; D. R. Jalihal" box="[920,1191,1001,1023]" pageId="14" pageNumber="63" pagination="495 - 514" refId="ref11273" refString="Pathan, D. I. and D. R. Jalihal 1997. Proposed taxonomic revision of some important penaeid prawn genera (Crustacea: Decapoda) of Konkan coast (west coast of India). Journal of the Bombay Natural History Society 94: 495 - 514." type="journal article" year="1997">Pathan and Jalihal, 1997: 510</bibRefCitation>
.
</paragraph>
<paragraph id="0B1CFC519879FFA7C69BFBC75B8AFBC8" blockId="14.[707,1237,913,1052]" box="[707,1207,1030,1052]" pageId="14" pageNumber="63">
?
<taxonomicName id="CCA387D29879FFA7C695FBC75C72FBCF" authorityName="H. Milne Edwards" authorityYear="1830" box="[717,847,1030,1052]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D720439879FFA7C695FBC75C72FBCF" box="[717,847,1030,1052]" italics="true" pageId="14" pageNumber="63">Acetes indicus</emphasis>
</taxonomicName>
:
<bibRefCitation id="6F3281A09879FFA7C703FBC65B62FBC8" author="Tirmizi, N. &amp; N. A. Ghani" box="[859,1119,1030,1052]" pageId="14" pageNumber="63" pagination="44 - 53" refId="ref11693" refString="Tirmizi, N. and N. A. Ghani 1982. New distributional records for three species of Acetes (Decapoda, Sergestidae). Crustaceana 42: 44 - 53." type="journal article" year="1982">Tirmizi and Ghani, 1982: 44</bibRefCitation>
, figs 1, 2.
</paragraph>
</subSubSection>
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<paragraph id="0B1CFC519879FFA7C685FBFA5C0AF9A9" blockId="14.[707,1236,1082,1799]" pageId="14" pageNumber="63">
<emphasis id="39D720439879FFA7C685FBFA5C9AFB86" box="[733,935,1082,1106]" italics="true" pageId="14" pageNumber="63">Material examined</emphasis>
.
<materialsCitation id="BBCBF60C9879FFA7C791FBFA5CA8FAF6" ID-GBIF-Occurrence="4899861309" collectingDate="2012-09-23" pageId="14" pageNumber="63">
<collectingCountry id="73B4BCC19879FFA7C791FBFA5B13FB86" box="[969,1070,1082,1106]" name="Malaysia" pageId="14" pageNumber="63">Malaysia</collectingCountry>
.
<emphasis id="39D720439879FFA7C009FBFA5C1EFBA1" italics="true" pageId="14" pageNumber="63">
<location id="0E7CAA8A9879FFA7C009FBFA5BE9FB86" LSID="urn:lsid:plazi:treatment:830A4D479879FFB8C773FCF25DABFCCE:0E7CAA8A9879FFA7C009FBFA5BE9FB86" box="[1105,1236,1082,1106]" country="Malaysia" name="Off Kuala" pageId="14" pageNumber="63" stateProvince="Selangor">Off Kuala</location>
<collectingRegion id="C96732B39879FFA7C69BFB9D5C1EFBA1" box="[707,803,1117,1141]" country="Malaysia" name="Selangor" pageId="14" pageNumber="63">Selangor</collectingRegion>
</emphasis>
,
<emphasis id="39D720439879FFA7C76CFB9D5B63FBA0" box="[820,1118,1117,1141]" italics="true" pageId="14" pageNumber="63">
<collectingRegion id="C96732B39879FFA7C76CFB9D5CA9FBA1" box="[820,916,1117,1141]" country="Malaysia" name="Selangor" pageId="14" pageNumber="63">Selangor</collectingRegion>
(Malacca Straits)
</emphasis>
:
<specimenCount id="1DA537D89879FFA7C028FB9D5BE9FBA1" box="[1136,1236,1117,1141]" count="31" pageId="14" pageNumber="63" type="male">31 males</specimenCount>
(cl
<quantity id="CC5B51B49879FFA7C6B2FBBF5C56FB43" box="[746,875,1151,1175]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.3" metricValueMin="3.2" pageId="14" pageNumber="63" unit="mm" value="3.75" valueMax="4.3" valueMin="3.2">3.24.3 mm</quantity>
),
<specimenCount id="1DA537D89879FFA7C7DAFBBF5CC1FB43" box="[898,1020,1151,1175]" count="38" pageId="14" pageNumber="63" type="female">38 females</specimenCount>
(cl
<quantity id="CC5B51B49879FFA7C077FBBF5B92FB43" box="[1071,1199,1151,1175]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.8" metricValueMin="3.3" pageId="14" pageNumber="63" unit="mm" value="4.55" valueMax="5.8" valueMin="3.3">3.35.8 mm</quantity>
),
<specimenCount id="1DA537D89879FFA7C09FFBBF5C1EFB6E" count="9" pageId="14" pageNumber="63" type="juvenile">9 juveniles</specimenCount>
and damaged specimens (cl 2.84.0 mm), several body parts, off-shore commercial catch,
<date id="7F1DDA919879FFA7C752FB275CA1FB2B" box="[778,924,1255,1279]" pageId="14" pageNumber="63" value="2012-09-23">
<collectingDate id="6F5923799879FFA7C752FB275CA1FB2B" box="[778,924,1255,1279]" pageId="14" pageNumber="63" value="2012-09-23">23 Sept 2012</collectingDate>
</date>
, coll.
<collectorName id="A65699879879FFA7C786FB285B51FB2B" box="[990,1132,1256,1279]" pageId="14" pageNumber="63">Y. Hanamura</collectorName>
&amp;
<collectorName id="A65699879879FFA7C0C9FB275BE9FB2B" box="[1169,1236,1255,1279]" pageId="14" pageNumber="63">Faizul</collectorName>
(NSMT-Cr 31637);
</materialsCitation>
<materialsCitation id="BBCBF60C9879FFA7C7F9FACA5B86FA5E" ID-GBIF-Occurrence="4899861305" pageId="14" pageNumber="63">
<specimenCount id="1DA537D89879FFA7C7F9FACA5B14FAF6" box="[929,1065,1290,1314]" count="2" pageId="14" pageNumber="63" type="generic">2 specimens</specimenCount>
used for DNA analysis (sex and size undetermined), data same as previous (NSMT-Cr 31638, 31639):
<specimenCount id="1DA537D89879FFA7C025FA905BE9FAB3" box="[1149,1236,1359,1383]" count="7" pageId="14" pageNumber="63" type="male">7 males</specimenCount>
(cl
<quantity id="CC5B51B49879FFA7C6B2FAB25C56FA5E" box="[746,875,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.4" metricValueMin="3.5" pageId="14" pageNumber="63" unit="mm" value="3.95" valueMax="4.4" valueMin="3.5">3.54.4 mm</quantity>
),
<specimenCount id="1DA537D89879FFA7C7DAFAB25CC1FA5E" box="[898,1020,1394,1418]" count="10" pageId="14" pageNumber="63" type="female">10 females</specimenCount>
(cl
<quantity id="CC5B51B49879FFA7C077FAB25B92FA5E" box="[1071,1199,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.8" metricValueMin="3.5" pageId="14" pageNumber="63" unit="mm" value="4.65" valueMax="5.8" valueMin="3.5">3.55.8 mm</quantity>
),
</materialsCitation>
<materialsCitation id="BBCBF60C9879FFA7C09FFAB25C2FF9C1" ID-GBIF-Occurrence="4899861310" pageId="14" pageNumber="63">
2 abdomens, data same as previous (NSMT-Cr 31640);
<specimenCount id="1DA537D89879FFA7C779FA775C4AFA1B" box="[801,887,1463,1487]" count="3" pageId="14" pageNumber="63" type="male">3 males</specimenCount>
(cl 3.74.0 mm),
<specimenCount id="1DA537D89879FFA7C019FA775B90FA1B" box="[1089,1197,1463,1487]" count="2" pageId="14" pageNumber="63" type="female">2 females</specimenCount>
(cl 5.2,
<quantity id="CC5B51B49879FFA7C6AEFA1A5C7AFA26" box="[758,839,1498,1522]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="14" pageNumber="63" unit="mm" value="5.8">5.8 mm</quantity>
), data same as previous (NSMT-Cr 31641)
</materialsCitation>
.
<materialsCitation id="BBCBF60C9879FFA7C744FA3D5C09F9A9" ID-GBIF-Occurrence="4899861311" collectingDate="2005-08-05" collectorName="Y. Hanamura &amp; Faizul" country="Malaysia" location="Malacca" pageId="14" pageNumber="63" specimenCount="14" specimenCount-female="13" specimenCount-male="1">
<emphasis id="39D720439879FFA7C744FA3D5C44F9C1" box="[796,889,1533,1557]" italics="true" pageId="14" pageNumber="63">
<collectingCountry id="73B4BCC19879FFA7C744FA3D5C44F9C1" box="[796,889,1533,1557]" name="Malaysia" pageId="14" pageNumber="63">Malacca</collectingCountry>
</emphasis>
:
<specimenCount id="1DA537D89879FFA7C7D0FA3D5CEDF9C1" box="[904,976,1533,1557]" count="1" pageId="14" pageNumber="63" type="male">1 male</specimenCount>
(cl
<quantity id="CC5B51B49879FFA7C7A2FA3D5B71F9C1" box="[1018,1100,1533,1557]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="63" unit="mm" value="3.8">3.8 mm</quantity>
),
<specimenCount id="1DA537D89879FFA7C007FA3D5BE9F9C1" box="[1119,1236,1533,1557]" count="13" pageId="14" pageNumber="63" type="female">13 females</specimenCount>
(cl
<quantity id="CC5B51B49879FFA7C6B2F9DF5C57F9E3" box="[746,874,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="6.3" metricValueMin="3.5" pageId="14" pageNumber="63" unit="mm" value="4.9" valueMax="6.3" valueMin="3.5">3.56.3 mm</quantity>
), fish market (dried products),
<date id="7F1DDA919879FFA7C09FF9DF5C0FF98E" pageId="14" pageNumber="63" value="2005-08-05">
<collectingDate id="6F5923799879FFA7C09FF9DF5C0FF98E" pageId="14" pageNumber="63" value="2005-08-05">5 Aug 2005</collectingDate>
</date>
, coll.
<collectorName id="A65699879879FFA7C72FF9825B3BF98D" box="[887,1030,1602,1625]" pageId="14" pageNumber="63">Y. Hanamura</collectorName>
&amp;
<collectorName id="A65699879879FFA7C076F9825B4CF98E" box="[1070,1137,1602,1626]" pageId="14" pageNumber="63">Faizul</collectorName>
(NSMT-Cr 31642)
</materialsCitation>
.
</paragraph>
<paragraph id="0B1CFC519879FFA7C685F9475BFCF931" blockId="14.[707,1236,1082,1799]" pageId="14" pageNumber="63">
<materialsCitation id="BBCBF60C9879FFA7C685F9475B80F931" ID-GBIF-Occurrence="4899861301" collectingDate="1972-12-01" collectionCode="T" collectorName="Achuthanktty &amp; M. Omori" country="India" location="Off Calangule" municipality="Goa" pageId="14" pageNumber="63" specimenCount="5" specimenCount-female="1" specimenCount-male="4">
<collectingCountry id="73B4BCC19879FFA7C685F9475C2AF94B" box="[733,791,1671,1695]" name="India" pageId="14" pageNumber="63">India</collectingCountry>
.
<emphasis id="39D720439879FFA7C77AF9475C86F94B" box="[802,955,1671,1695]" italics="true" pageId="14" pageNumber="63">
<location id="0E7CAA8A9879FFA7C77AF9475C86F94B" LSID="urn:lsid:plazi:treatment:830A4D479879FFB8C773FCF25DABFCCE:0E7CAA8A9879FFA7C77AF9475C86F94B" box="[802,955,1671,1695]" country="India" municipality="Goa" name="Off Calangule" pageId="14" pageNumber="63">Off Calangule</location>
</emphasis>
,
<emphasis id="39D720439879FFA7C791F9485CCAF94B" box="[969,1015,1672,1695]" italics="true" pageId="14" pageNumber="63">
<collectingMunicipality id="EB78662B9879FFA7C791F9485CCAF94B" box="[969,1015,1672,1695]" pageId="14" pageNumber="63">Goa</collectingMunicipality>
</emphasis>
;
<specimenCount id="1DA537D89879FFA7C05EF9475B64F94B" box="[1030,1113,1671,1695]" count="4" pageId="14" pageNumber="63" type="male">4 males</specimenCount>
(cl
<quantity id="CC5B51B49879FFA7C0DCF9475DCDF916" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.8" metricValueMin="3.9" pageId="14" pageNumber="63" unit="mm" value="4.35" valueMax="4.8" valueMin="3.9">3.94.8 mm</quantity>
),
<specimenCount id="1DA537D89879FFA7C75AF96A5C63F916" box="[770,862,1706,1730]" count="1" pageId="14" pageNumber="63" type="female">1 female</specimenCount>
(cl
<quantity id="CC5B51B49879FFA7C7DFF96A5CE5F916" box="[903,984,1706,1730]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="14" pageNumber="63" unit="mm" value="6.4">6.4 mm</quantity>
),
<date id="7F1DDA919879FFA7C7B3F96A5B5AF916" box="[1003,1127,1706,1730]" pageId="14" pageNumber="63" value="1972-12-01">
<collectingDate id="6F5923799879FFA7C7B3F96A5B5AF916" box="[1003,1127,1706,1730]" pageId="14" pageNumber="63" value="1972-12-01">1 Dec 1972</collectingDate>
</date>
, coll. C.
<collectionCode id="6DB264949879FFA7C0E6F96A5BEDF916" box="[1214,1232,1706,1730]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="63">T</collectionCode>
.
<collectorName id="A65699879879FFA7C69BF90D5C68F931" box="[707,853,1741,1765]" pageId="14" pageNumber="63">Achuthanktty</collectorName>
, id.
<collectorName id="A65699879879FFA7C7DAF90D5CD0F931" box="[898,1005,1741,1765]" pageId="14" pageNumber="63">M. Omori</collectorName>
(NSMT-Cr 22267)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA9879FFB9C685F9305B0EFA5E" lastPageId="16" lastPageNumber="65" pageId="14" pageNumber="63" type="description">
<paragraph id="0B1CFC519879FFA6C685F9305DA1F8D3" blockId="14.[707,1236,1082,1799]" lastBlockId="15.[140,669,1637,1799]" lastPageId="15" lastPageNumber="64" pageId="14" pageNumber="63">
<emphasis id="39D720439879FFA7C685F9305C89F8D3" box="[733,948,1775,1799]" italics="true" pageId="14" pageNumber="63">Description. Male</emphasis>
. Rostrum (
<figureCitation id="9398E0D49879FFA7C068F92F5B43F8D3" box="[1072,1150,1775,1799]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="14" pageNumber="63">Fig. 7a</figureCitation>
) subtriangular, elevated slightly from dorsal margin of carapace, apex ending into sharp tooth, and armed dorsally with 2 teeth, posterior tooth larger than anterior one. Carapace (Fig. 9a) smooth, with small post-orbital and hepatic teeth.
</paragraph>
<caption id="5FDCACD99878FFA6C4FBFA8E5D1BF9C0" ID-DOI="http://doi.org/10.5281/zenodo.12572364" ID-Zenodo-Dep="12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="15" pageNumber="64" startId="15.[163,196,1358,1380]" targetBox="[275,1103,215,1323]" targetPageId="15" targetType="figure">
<paragraph id="0B1CFC519878FFA6C4FBFA8E5D1BF9C0" blockId="15.[163,1214,1358,1556]" pageId="15" pageNumber="64">
Fig. 7.
<taxonomicName id="CCA387D29878FFA6C4A1FA8E5D36FAB7" authority="Hansen, 1919" authorityName="Hansen" authorityYear="1919" box="[249,523,1358,1379]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="64" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="39D720439878FFA6C4A1FA8E5EB8FAB7" box="[249,389,1358,1379]" italics="true" pageId="15" pageNumber="64">Acetes spiniger</emphasis>
<bibRefCitation id="6F3281A09878FFA6C5D3FA8E5D36FAB7" author="Hansen, H. J." box="[395,523,1358,1379]" pageId="15" pageNumber="64" pagination="1 - 65" refId="ref10693" refString="Hansen, H. J. 1919. The Sergestidae of the Siboga-Expedition. Siboga Expeditie 38: 1 - 65, pls 1 - 5." type="journal article" year="1919">Hansen, 1919</bibRefCitation>
</taxonomicName>
. (ae) male (cl 4.3 mm) and (fi) female (cl. 5.8 mm) from off Kuala Selangor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively.
</paragraph>
</caption>
<paragraph id="0B1CFC519878FFB9C685F9A55EDAFE83" blockId="15.[707,1236,1637,1799]" lastBlockId="16.[140,670,215,1799]" lastPageId="16" lastPageNumber="65" pageId="15" pageNumber="64">
Abdomen with first sternite possessing procurved sharp tooth at mid-part between right and left pleopods; sixth somite (
<figureCitation id="9398E0D49878FFA6C059F96A5B6EF916" box="[1025,1107,1706,1730]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="15" pageNumber="64">Fig. 7b</figureCitation>
) somewhat compressed, 2.12.3 times as long as fifth, bearing small tooth or point at postero-median part, without red spot on postero-ventral part. Telson (
<figureCitation id="9398E0D49867FFB9C4CCFF3A5FCAFEC6" box="[148,247,250,274]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7b, i</figureCitation>
) 0.60.7 times as long as sixth abdominal somite, posterior part somewhat narrowing, then forming sub-triangular end.
</paragraph>
<paragraph id="0B1CFC519867FFB9C4FFFEA25DB9FE48" blockId="16.[140,670,215,1799]" pageId="16" pageNumber="65">
Eye (
<figureCitation id="9398E0D49867FFB9C4B8FEA25E13FEAE" box="[224,302,354,378]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7a</figureCitation>
) with well pigmented cornea, latter wider than ocular peduncle in dorsal aspect.
</paragraph>
<paragraph id="0B1CFC519867FFB9C4FFFE675D49FC3E" blockId="16.[140,670,215,1799]" pageId="16" pageNumber="65">
Antennule (
<figureCitation id="9398E0D49867FFB9C57EFE675E49FE6B" box="[294,372,423,447]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7a</figureCitation>
) with elongate peduncle in adults, particularly in third segment, latter 0.92 1.10 times as long as combined length of first and second segments: lower flagellum (
<figureCitation id="9398E0D49867FFB9C61DFDCF5DA8FDF3" box="[581,661,527,551]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7c</figureCitation>
) modified into clasping organ, most commonly with 12 sub-segments, distal part of first segment of trunk part forming nearly vertical or weak conjunction with second one; major clasping setal-spine somewhat slender, extending as far as mid-length of fourth segment of main branch, with single line of serrated denticles and rudimentary clasping spine placed near base of major spine; first segment of main branch with or without setal-spine; second and third segments with single setal-spine, fourth segment 0.50.7 times as long as combined length of anterior 2 segments, bearing 46 setal-spines on distal half.
</paragraph>
<paragraph id="0B1CFC519867FFB9C4FFFC355FC7FB43" blockId="16.[140,670,215,1799]" pageId="16" pageNumber="65">
Antennal scale (
<figureCitation id="9398E0D49867FFB9C538FC355E8EFBD9" box="[352,435,1013,1037]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7a</figureCitation>
) 0.700.81 times as long as carapace, barely reaching end of second segment of antennular peduncle but not extending beyond it, outer margin with small tooth at distal end.
</paragraph>
<paragraph id="0B1CFC519867FFB9C4FFFB625E3DF98D" blockId="16.[140,670,215,1799]" pageId="16" pageNumber="65">
Third maxilliped reaching mid-length of third segment of antennular peduncle, distal segment two-thirds times as long as penultimate segment. First pereopod shortest, falling slightly short of cornea of eye; second pereopod falling short of anterior end of antennal scale; third pereopod (
<figureCitation id="9398E0D49867FFB9C4CCFAB25FDAFA5E" box="[148,231,1394,1418]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7d</figureCitation>
) longest, reaching mid-length of third segment of antennular peduncle; coxa with stout tooth or projection on distal inner margin; basis with obtuse tooth near distal inner part. Genital coxa narrowly tapering distally in lateral aspect, but widely truncated or weakly grooved in ventral aspect.
</paragraph>
<paragraph id="0B1CFC519867FFB9C4FFF9A55B96FEC6" blockId="16.[140,670,215,1799]" lastBlockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">
Petasma (
<figureCitation id="9398E0D49867FFB9C54BF9A55E59F9A9" box="[275,356,1637,1661]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7e</figureCitation>
) with capitulum robust and somewhat rounded distally, bearing numerous hooks on distal and lateral surfaces; processus ventralis narrow, reaching as far as distal end of capitulum; pars astringen occasionally noticeable as weak elevation or vestigial lobule. Appendix masculina with 2 or 3 hooks on distal margin.
</paragraph>
<paragraph id="0B1CFC519867FFB9C685FEDD5B2AFE36" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">
Uropod (
<figureCitation id="9398E0D49867FFB9C71BFEDD5CA8FEE1" box="[835,917,285,309]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7b</figureCitation>
) with exopod bearing small but distinct tooth at distal 0.640.76 on lateral margin, basal part from which is naked while setose in distal part; no confident data for its number of red spots, but at least 1 or 2 spots present in preserved specimens.
</paragraph>
<paragraph id="0B1CFC519867FFB9C685FE2D5DCCFD9D" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">
<emphasis id="39D720439867FFB9C685FE2D5C10FDD1" box="[733,813,493,517]" italics="true" pageId="16" pageNumber="65">Female</emphasis>
. Usually larger in body size than males and also differing in several sexual characters.
</paragraph>
<paragraph id="0B1CFC519867FFB9C685FD955CA7FCCE" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">
Antennule (
<figureCitation id="9398E0D49867FFB9C704FD955C9AFDB9" box="[860,935,597,621]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7f</figureCitation>
) with first segment longest, and third segment of peduncle not elongated as in male, 0.330.47 times as long as combined length of second and third segments; lower flagellum simple and not modified, composed of 19 or 20 sub-segments.
</paragraph>
<paragraph id="0B1CFC519867FFB9C685FCE55B88FC56" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">
Antennal scale (
<figureCitation id="9398E0D49867FFB9C7C0FCE55CD4FCE9" box="[920,1001,805,829]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="16" pageNumber="65">Fig. 7f</figureCitation>
) 0.640.73 times as long as carapace, reaching about basal 2/51/3 length of third segment of antennule peduncle.
</paragraph>
<paragraph id="0B1CFC519867FFB9C685FC4D5C58FBD9" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">Third maxilliped extending beyond antennular peduncle by distal and 1/21/3 of penultimate segment, distal segment 0.700.75 times as long as penultimate.</paragraph>
<paragraph id="0B1CFC519867FFB9C685FBD75CEAFB6E" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">First pereopod shortest, fully reaching cornea of eye; second pereopod falling slightly short of distal end of antennular peduncle; third pereopod extending beyond antennular peduncle by length of propodus and dactylus.</paragraph>
<paragraph id="0B1CFC519867FFB9C685FB055B3AFA91" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">
Third and fourth thoracic sternites forming longitudinal groove rather wide and shallow, not conspicuously narrowing anteriorly and not swelling markedly (
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Fig.
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, h
</figureCitation>
)
</paragraph>
<paragraph id="0B1CFC519867FFB9C685FA8F5B0EFA5E" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">
<emphasis id="39D720439867FFB9C685FA8F5C6FFAB3" box="[733,850,1359,1383]" italics="true" pageId="16" pageNumber="65">Body size</emphasis>
. Largest male cl
<quantity id="CC5B51B49867FFB9C012FA8F5BA5FAB3" box="[1098,1176,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="16" pageNumber="65" unit="mm" value="4.8">4.8 mm</quantity>
and female cl
<quantity id="CC5B51B49867FFB9C774FAB25C47FA5D" box="[812,890,1394,1418]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="16" pageNumber="65" unit="mm" value="6.4">6.4 mm</quantity>
from Goa,
<collectingCountry id="73B4BCC19867FFB9C7ADFAB25B12FA5E" box="[1013,1071,1394,1418]" name="India" pageId="16" pageNumber="65">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA9867FFB9C685FA555C5CF9E3" pageId="16" pageNumber="65" type="materials_examined">
<paragraph id="0B1CFC519867FFB9C685FA555C5CF9E3" blockId="16.[707,1236,215,1799]" pageId="16" pageNumber="65">
<emphasis id="39D720439867FFB9C685FA555C56FA79" box="[733,875,1429,1453]" italics="true" pageId="16" pageNumber="65">
<typeStatus id="D41842F39867FFB9C685FA555C32FA78" box="[733,783,1429,1452]" pageId="16" pageNumber="65">Type</typeStatus>
locality
</emphasis>
. The
<typeStatus id="D41842F39867FFB9C798FA555CD3FA79" box="[960,1006,1429,1453]" pageId="16" pageNumber="65">type</typeStatus>
locality of this species should be off
<collectingRegion id="C96732B39867FFB9C7C1FA785CC1FA04" box="[921,1020,1464,1488]" country="Myanmar" name="Yangon" pageId="16" pageNumber="65">Rangoon</collectingRegion>
, Burma (now Yangon,
<collectingCountry id="73B4BCC19867FFB9C6A1FA1A5C58FA26" box="[761,869,1498,1522]" name="Myanmar" pageId="16" pageNumber="65">Myanmar</collectingCountry>
), based on which materials
<bibRefCitation id="6F3281A09867FFB9C0D6FA1A5C37F9C0" author="Omori, M." pageId="16" pageNumber="65" pagination="1 - 89" refId="ref11166" refString="Omori, M. 1975. The systematics, biogeography, and fish- ery of epipelagic shrimps of the genus Acetes (Crustacea, Decapoda, Sergestidae). Bulletin of the Ocean Research Institute, University of Tokyo 7: 1 - 89." type="journal article" year="1975">Omori (1975)</bibRefCitation>
evidently noted the morphological traits of this species.
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA9867FFB8C685F9825F83FE6B" lastPageId="17" lastPageNumber="66" pageId="16" pageNumber="65" type="discussion">
<paragraph id="0B1CFC519867FFB8C685F9825F83FE6B" blockId="16.[707,1236,215,1799]" lastBlockId="17.[140,670,215,794]" lastPageId="17" lastPageNumber="66" pageId="16" pageNumber="65">
<emphasis id="39D720439867FFB9C685F9825C07F98E" box="[733,826,1602,1626]" italics="true" pageId="16" pageNumber="65">Remarks</emphasis>
.
<bibRefCitation id="6F3281A09867FFB9C704F9825B4EF98E" author="Tirmizi, N. &amp; N. A. Ghani" box="[860,1139,1602,1626]" pageId="16" pageNumber="65" pagination="44 - 53" refId="ref11693" refString="Tirmizi, N. and N. A. Ghani 1982. New distributional records for three species of Acetes (Decapoda, Sergestidae). Crustaceana 42: 44 - 53." type="journal article" year="1982">Tirmizi and Ghani (1982)</bibRefCitation>
reported
<taxonomicName id="CCA387D29867FFB9C69BF9A55C13F9A9" authorityName="H. Milne Edwards" authorityYear="1830" box="[707,814,1637,1661]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D720439867FFB9C69BF9A55C13F9A9" box="[707,814,1637,1661]" italics="true" pageId="16" pageNumber="65">A. indicus</emphasis>
</taxonomicName>
from
<collectingCountry id="73B4BCC19867FFB9C728F9A55CF0F9A9" box="[880,973,1637,1661]" name="Pakistan" pageId="16" pageNumber="65">Pakistan</collectingCountry>
, as a species resembling Omori`s llarge formz (=
<taxonomicName id="CCA387D29867FFB9C78BF9485B05F94B" authority="Alamri &amp; Alotaiby, 2024" authorityName="Alamri &amp; Alotaiby" authorityYear="2024" box="[979,1080,1672,1695]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="39D720439867FFB9C78BF9485B05F94B" box="[979,1080,1672,1695]" italics="true" pageId="16" pageNumber="65">A. omorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22E49D389867FFB9C018F9485BA5F94B" box="[1088,1176,1671,1695]" pageId="16" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
). The male lower flagellum figured by these authors appeared to show an unusual form for the l
<taxonomicName id="CCA387D29867FFB9C0E5F90D5C2DF8D3" authorityName="H. Milne Edwards" authorityYear="1830" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="65" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D720439867FFB9C0E5F90D5C2DF8D3" italics="true" pageId="16" pageNumber="65">A. indicus</emphasis>
</taxonomicName>
z species complex in having an extremely elongated first segment of the main branch as well as a relatively short clasping tooth. However, the form of the female genital area seemed similar to
<taxonomicName id="CCA387D29866FFB8C4A7FE805E4AFE83" authorityName="Hansen" authorityYear="1919" box="[255,375,320,343]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="66" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="39D720439866FFB8C4A7FE805E4AFE83" box="[255,375,320,343]" italics="true" pageId="17" pageNumber="66">A. spiniger</emphasis>
</taxonomicName>
rather than the latter by a broad sternal groove, although future confirmation is in need for specimens occurring in
<collectingCountry id="73B4BCC19866FFB8C63DFE455F87FE6B" name="Pakistan" pageId="17" pageNumber="66">Pakistan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA9866FFB8C4FFFE0A5DABFCCE" pageId="17" pageNumber="66" type="distribution">
<paragraph id="0B1CFC519866FFB8C4FFFE0A5DABFCCE" blockId="17.[140,670,215,794]" pageId="17" pageNumber="66">
<emphasis id="39D720439866FFB8C4FFFE0A5E14FE36" box="[167,297,458,482]" italics="true" pageId="17" pageNumber="66">Distribution</emphasis>
. This species is known with certainty in the peripheral region of the north-eastern part of the Indian Ocean, from the
<collectingCountry id="73B4BCC19866FFB8C61BFDCF5DA0FDF3" box="[579,669,527,551]" name="Malaysia" pageId="17" pageNumber="66">Malacca</collectingCountry>
Straits and off Galangule, Goa,
<collectingCountry id="73B4BCC19866FFB8C5BAFDF25D24FD9E" box="[482,537,562,586]" name="India" pageId="17" pageNumber="66">India</collectingCountry>
(and probably to
<collectingCountry id="73B4BCC19866FFB8C48BFD955E0FFDB9" box="[211,306,597,621]" name="Pakistan" pageId="17" pageNumber="66">Pakistan</collectingCountry>
). In the
<collectingCountry id="73B4BCC19866FFB8C5D4FD955EDBFDB9" box="[396,486,597,621]" name="Malaysia" pageId="17" pageNumber="66">Malacca</collectingCountry>
Straits,
<taxonomicName id="CCA387D29866FFB8C619FD955F8DFD5B" authorityName="Hansen" authorityYear="1919" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="66" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="39D720439866FFB8C619FD955F8DFD5B" italics="true" pageId="17" pageNumber="66">A. spiniger</emphasis>
</taxonomicName>
is commercially fished by offshore trawls. Contrary to this,
<taxonomicName id="CCA387D29866FFB8C51BFD5A5E95FD66" authority="Alamri &amp; Alotaiby, 2024" authorityName="Alamri &amp; Alotaiby" authorityYear="2024" box="[323,424,666,690]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="66" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="39D720439866FFB8C51BFD5A5E95FD66" box="[323,424,666,690]" italics="true" pageId="17" pageNumber="66">A. omorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22E49D389866FFB8C5F7FD5A5D3EFD65" box="[431,515,666,689]" pageId="17" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
occur in great abundance in the inshore waters including mangrove estuaries, suggesting that the two closest relatives are sharing their major habitats (
<figureCitation id="9398E0D49866FFB8C61FFCC25DB5FCCE" box="[583,648,770,794]" captionStart="Fig" captionStartId="8.[163,196,811,833]" captionTargetBox="[235,1140,214,775]" captionTargetId="figure-280@8.[234,1141,213,777]" captionTargetPageId="8" captionText="Fig. 3. Map showing sampling localities and distribution ranges of Acetes indicus, A. omorii sp. nov. and A. spiniger recognised in this study with certainty: blue solid circles (nos. 12) for A. indicus, green solid squares and open square (museum vouchered specimens) for A. spiniger, and red solid triangles for A. omorii sp. nov. Numerals indicating sampling locations: 1, Chiringa; 2, Kumira-Sandwip Ship Ghat; 3, Merbok Mangrove; 4; Matang Mangrove, 5; Kuala Selangor; 6, Malacca; 7, Samut Prakan; 8, Setarap Estuary, South Kalimantan; and 9, Goa." figureDoi="http://doi.org/10.5281/zenodo.12572342" httpUri="https://zenodo.org/record/12572342/files/figure.png" pageId="17" pageNumber="66">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,518 @@
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<taxonomicName id="CCA387D2987CFFA2C4B7FB355E89FAFB" authority="Hanamura, Imai and Hardianto" authorityName="Hanamura, Imai and Hardianto" class="Malacostraca" family="Acetidae" genus="Acetes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="omorii">
<emphasis id="39D72043987CFFA2C4B7FB355EBBFAD9" bold="true" box="[239,390,1269,1293]" italics="true" pageId="11" pageNumber="60">Acetes omorii</emphasis>
Hanamura, Imai and Hardianto
</taxonomicName>
<taxonomicNameLabel id="22E49D38987CFFA2C5E3FAD85D30FAFB" box="[443,525,1304,1327]" pageId="11" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA987CFFA2C53DFA865E7DFA4A" pageId="11" pageNumber="60" type="description">
<paragraph id="0B1CFC51987CFFA2C53DFA865EFEFA88" blockId="11.[140,669,1350,1438]" box="[357,451,1350,1372]" pageId="11" pageNumber="60">
(
<figureCitation id="9398E0D4987CFFA2C534FA865E9AFA88" box="[364,423,1350,1372]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="11" pageNumber="60">Figs 5</figureCitation>
,
<figureCitation id="9398E0D4987CFFA2C5E8FA865E81FA88" box="[432,444,1350,1372]" captionStart="Fig" captionStartId="13.[163,196,1092,1114]" captionTargetBox="[273,1102,215,1048]" captionTargetId="figure-225@13.[272,1104,213,1049]" captionTargetPageId="13" captionText="Fig. 6. Acetes omorii Hanamura, Imai &amp; Hardianto sp. nov. (a) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) (NSMT-Cr 31622) from Matang Mangrove, Malaysia: a, b, right lower antennular flagellum of male and female, respectively; c, basal segments of third pereopod and third and fourth abdominal sternites, ventral; d, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572358" httpUri="https://zenodo.org/record/12572358/files/figure.png" pageId="11" pageNumber="60">6</figureCitation>
)
</paragraph>
<paragraph id="0B1CFC51987CFFA2C4D4FAAA5E7DFA4A" blockId="11.[140,669,1350,1438]" pageId="11" pageNumber="60">urn:lsid:zoobank.org:act:1199DE94-F3F4-477F-9E5B-3AD3A9C65111</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA987CFFA2C4D4FA785E34F99A" pageId="11" pageNumber="60" type="reference_group">
<paragraph id="0B1CFC51987CFFA2C4D4FA785DA6F9C1" blockId="11.[140,670,1463,1615]" pageId="11" pageNumber="60">
<treatmentCitationGroup id="2BB3DB7F987CFFA2C4D4FA785DA6F9C1" pageId="11" pageNumber="60">
<taxonomicName id="CCA387D2987CFFA2C4D4FA785E1AFA1B" ID-CoL="9XFD2" authorityName="H. Milne Edwards" authorityYear="1830" box="[140,295,1463,1487]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D72043987CFFA2C4D4FA785E1AFA1B" box="[140,295,1463,1487]" italics="true" pageId="11" pageNumber="60">Acetes indicus</emphasis>
</taxonomicName>
;
<treatmentCitation id="8A02DA40987CFFA2C56EFA775ED7FA1B" author="Omori, M." box="[310,490,1463,1487]" page="15" pageId="11" pageNumber="60" year="1975">
<bibRefCitation id="6F3281A0987CFFA2C56EFA775ED7FA1B" author="Omori, M." box="[310,490,1463,1487]" pageId="11" pageNumber="60" pagination="1 - 89" refId="ref11166" refString="Omori, M. 1975. The systematics, biogeography, and fish- ery of epipelagic shrimps of the genus Acetes (Crustacea, Decapoda, Sergestidae). Bulletin of the Ocean Research Institute, University of Tokyo 7: 1 - 89." type="journal article" year="1975">
<collectingRegion id="C96732B3987CFFA2C56EFA775E43FA1B" box="[310,382,1463,1487]" country="Japan" name="Aomori" pageId="11" pageNumber="60">Omori</collectingRegion>
, 1975: 15
</bibRefCitation>
</treatmentCitation>
(in part; at least fig. 15c, e, i);
<treatmentCitation id="8A02DA40987CFFA2C570FA1A5F8BF9C1" author="Perez Farfante, I. &amp; B. Kensley" page="187" pageId="11" pageNumber="60" year="1997">
<bibRefCitation id="6F3281A0987CFFA2C570FA1A5F8BF9C1" author="Perez Farfante, I. &amp; B. Kensley" pageId="11" pageNumber="60" pagination="1 - 233" refId="ref11352" refString="Perez Farfante, I. and B. Kensley 1997. Penaeoid and sergestoid shrimps and prawns of the world: keys and diagnoses for the families and genera. Memoires du Museum National d`Histoire Naturelle 175: 1 - 233." type="journal article" year="1997">Pérez Farfante and Kensley, 1997: 187</bibRefCitation>
</treatmentCitation>
, figs 128130 (not H.
<bibRefCitation id="6F3281A0987CFFA2C5FDFA3D5DADF9C1" author="Milne Edwards, M. H." box="[421,656,1533,1557]" pageId="11" pageNumber="60" pagination="333 - 352" refId="ref11123" refString="Milne Edwards, M. H. 1830. Description des genres Glaucothoe, Sicyonie, Sergeste et Acete, de l`ordre des Crustaces decapodes. Annales des Sciences Naturelles, Paris 19: 333 - 352, pls 8 - 11." type="journal article" year="1830">Milne Edwards, 1830</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0B1CFC51987CFFA2C4D4F9DC5E34F99A" blockId="11.[140,670,1463,1615]" pageId="11" pageNumber="60">
<treatmentCitationGroup id="2BB3DB7F987CFFA2C4D4F9DC5DA3F9E6" box="[140,670,1564,1586]" pageId="11" pageNumber="60">
<emphasis id="39D72043987CFFA2C4D4F9DC5E21F9E5" box="[140,284,1564,1585]" italics="true" pageId="11" pageNumber="60">
?
<taxonomicName id="CCA387D2987CFFA2C4CCF9DC5E21F9E5" ID-CoL="9XFD2" authorityName="H. Milne Edwards" authorityYear="1830" box="[148,284,1564,1585]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="indicus">Acetes indicus</taxonomicName>
</emphasis>
:
<treatmentCitation id="8A02DA40987CFFA2C573F9DC5E80F9E5" author="Kemp, S." box="[299,445,1564,1585]" page="47" pageId="11" pageNumber="60" year="1917">
<bibRefCitation id="6F3281A0987CFFA2C573F9DC5E80F9E5" author="Kemp, S." box="[299,445,1564,1585]" pageId="11" pageNumber="60" pagination="43 - 58" refId="ref11017" refString="Kemp, S. 1917. Notes on Crustacea Decapoda in the Indian Museum. The genus Acetes Milne-Edwards. Records of the Indian Museum 53: 43 - 58." type="journal article" year="1917">Kemp 1917: 47</bibRefCitation>
</treatmentCitation>
(in part):
<bibRefCitation id="6F3281A0987CFFA2C678F9DC5DA3F9E6" author="Boone, L." box="[544,670,1564,1586]" pageId="11" pageNumber="60" pagination="1 - 246" refId="ref10390" refString="Boone, L. 1935. Scientific results of the world cruise of the yacht lAlvaz, 1931, William K. Vanderbilt, commanding. Crustacea: Anomura, Macrura, Euphausiacea, Isopoda, Amphipoda and Echinodermata: Asteroidea and Echinoidea. Bulletin of the Vanderbilt Marine Museum 6: 1 - 246, pls 1 - 96." type="journal article" year="1935">Boone, 1935:</bibRefCitation>
</treatmentCitationGroup>
101, fig. 9.
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA987CFFA7C4FFF9475CD0FD23" lastPageId="14" lastPageNumber="63" pageId="11" pageNumber="60" type="materials_examined">
<paragraph id="0B1CFC51987CFFA2C4FFF9475C59FD5B" blockId="11.[140,670,1671,1799]" lastBlockId="11.[707,1237,215,1799]" pageId="11" pageNumber="60">
<emphasis id="39D72043987CFFA2C4FFF9475E4CF94B" box="[167,369,1671,1695]" italics="true" pageId="11" pageNumber="60">Material examined</emphasis>
.
<materialsCitation id="BBCBF60C987CFFA2C5CAF9475B64FF3B" ID-GBIF-Occurrence="4899861307" collectingDate="2012-11-25" collectionCode="R" collectorName="Y. Hanamura &amp; Faizul" country="Malaysia" location="Matang Mangrove Estuary" pageId="11" pageNumber="60" specimenCode="R 2" specimenCount="1" typeStatus="holotype">
<typeStatus id="D41842F3987CFFA2C5CAF9475EC4F94B" box="[402,505,1671,1695]" pageId="11" pageNumber="60" type="holotype">Holotype</typeStatus>
: male (cl
<quantity id="CC5B51B4987CFFA2C4D4F96A5FE3F916" box="[140,222,1706,1730]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="60" unit="mm" value="5.3">5.3 mm</quantity>
), stn
<specimenCode id="5B05542A987CFFA2C57EF96A5E6BF916" box="[294,342,1706,1730]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="60">R 2</specimenCode>
,
<location id="0E7CAA8A987CFFA2C532F96A5DA7F916" LSID="urn:lsid:plazi:treatment:830A4D47987CFFA7C4B7FB355CD0FD23:0E7CAA8A987CFFA2C532F96A5DA7F916" box="[362,666,1706,1730]" country="Malaysia" name="Matang Mangrove Estuary" pageId="11" pageNumber="60">Matang Mangrove Estuary</location>
,
<collectingCountry id="73B4BCC1987CFFA2C4D4F90D5FCDF931" box="[140,240,1741,1765]" name="Malaysia" pageId="11" pageNumber="60">Malaysia</collectingCountry>
, sledge,
<date id="7F1DDA91987CFFA2C537F90C5D18F931" box="[367,549,1740,1765]" pageId="11" pageNumber="60" value="2012-11-25">
<collectingDate id="6F592379987CFFA2C537F90C5D18F931" box="[367,549,1740,1765]" pageId="11" pageNumber="60" value="2012-11-25">25 Nov. 2012</collectingDate>
</date>
, coll.
<collectorName id="A6569987987CFFA2C6DFF90D5FC6F8D3" pageId="11" pageNumber="60">Y. Hanamura</collectorName>
&amp;
<collectorName id="A6569987987CFFA2C573F92F5E53F8D3" box="[299,366,1775,1799]" pageId="11" pageNumber="60">Faizul</collectorName>
(NSMT-Cr 31621) (transferred from formalin to 70% ethanol)
</materialsCitation>
.
<typeStatus id="D41842F3987CFFA2C03DFF185BECFF24" box="[1125,1233,216,240]" pageId="11" pageNumber="60" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="1DA537D8987CFFA2C69BFF3A5C31FEC6" box="[707,780,250,274]" count="1" pageId="11" pageNumber="60" type="male">1 male</specimenCount>
(
<quantity id="CC5B51B4987CFFA2C742FF3A5C50FEC6" box="[794,877,250,274]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="60" unit="mm" value="4.8">4.8 mm</quantity>
),
<materialsCitation id="BBCBF60C987CFFA2C7D9FF3A5B2BFE83" ID-GBIF-Occurrence="4899861308" collectingDate="2012-11-25" collectionCode="R" collectorName="Y. Hanamura &amp; Faizul" country="Malaysia" location="Matang Mangrove Estuary" pageId="11" pageNumber="60" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="1DA537D8987CFFA2C7D9FF3A5CD4FEC6" box="[897,1001,250,274]" count="3" pageId="11" pageNumber="60" type="female">3 females</specimenCount>
(cl
<quantity id="CC5B51B4987CFFA2C04CFF3A5BA9FEC6" box="[1044,1172,250,274]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.2" metricValueMin="5.9" pageId="11" pageNumber="60" unit="mm" value="6.05" valueMax="6.2" valueMin="5.9">5.96.2 mm</quantity>
), data same as holotype (NSMT-Cr 31622) (transferred from formalin to 70% ethanol)
</materialsCitation>
;
<specimenCount id="1DA537D8987CFFA2C07BFEFF5B4AFE83" box="[1059,1143,319,343]" count="5" pageId="11" pageNumber="60" type="male">5 males</specimenCount>
(cl
<quantity id="CC5B51B4987CFFA2C0FDFEFF5C29FEAE" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="5.3" metricValueMin="3.5" pageId="11" pageNumber="60" unit="mm" value="4.4" valueMax="5.3" valueMin="3.5">3.5 5.3 mm</quantity>
),
<specimenCount id="1DA537D8987CFFA2C772FEA25CA8FEAE" box="[810,917,354,378]" count="9" pageId="11" pageNumber="60" type="female">9 females</specimenCount>
(cl 5.56.0 mm), 2 individuals (sex &amp; size undetermined), sledge,
<date id="7F1DDA91987CFFA2C027FE445DC6FE6B" pageId="11" pageNumber="60" value="2008-11-25">25 Nov 2008</date>
, coll. Y. Hanamura &amp; Faizul (NSMT-Cr 31623);
<specimenCount id="1DA537D8987CFFA2C746FE0A5C9FFE36" box="[798,930,458,482]" count="2" pageId="11" pageNumber="60" type="generic">2 specimens</specimenCount>
used for DNA analysis (sex and size undetermined), data same as previous (NSMT-Cr 31624, 31625);
<specimenCount id="1DA537D8987CFFA2C054FDCF5B57FDF3" box="[1036,1130,527,551]" count="2" pageId="11" pageNumber="60" type="male">2 males</specimenCount>
(cl 4.7,
<quantity id="CC5B51B4987CFFA2C69BFDF25C29FD9E" box="[707,788,562,586]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="11" pageNumber="60" unit="mm" value="5.5">5.5 mm</quantity>
),
<materialsCitation id="BBCBF60C987CFFA2C769FDF25BEDFD9E" ID-GBIF-Occurrence="4899861306" box="[817,1232,562,586]" collectionCode="R" country="Malaysia" pageId="11" pageNumber="60" specimenCode="R2" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="1DA537D8987CFFA2C769FDF25C9EFD9E" box="[817,931,562,586]" count="3" pageId="11" pageNumber="60" type="female">3 females</specimenCount>
(cl 6.47.0 mm), stn
<specimenCode id="5B05542A987CFFA2C0F6FDF25BEDFD9E" box="[1198,1232,562,586]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="60">R2</specimenCode>
</materialsCitation>
,
<materialsCitation id="BBCBF60C987CFFA2C69BFD955C5CFD5B" ID-GBIF-Occurrence="4899861304" collectingDate="2012-08-29" collectionCode="FRI" collectorName="Y. Hanamura &amp; Faizul" country="Malaysia" pageId="11" pageNumber="60" specimenCode="CR0011" specimenCount="1" typeStatus="paratype">
sledge,
<date id="7F1DDA91987CFFA2C74DFD955C9FFDB9" box="[789,930,597,621]" pageId="11" pageNumber="60" value="2012-08-29">
<collectingDate id="6F592379987CFFA2C74DFD955C9FFDB9" box="[789,930,597,621]" pageId="11" pageNumber="60" value="2012-08-29">29 Aug 2012</collectingDate>
</date>
, coll.
<collectorName id="A6569987987CFFA2C7BAFD955B53FDB8" box="[994,1134,597,620]" pageId="11" pageNumber="60">Y. Hanamura</collectorName>
&amp;
<collectorName id="A6569987987CFFA2C0C9FD955BE9FDB9" box="[1169,1236,597,621]" pageId="11" pageNumber="60">Faizul</collectorName>
(
<collectionCode id="6DB26494987CFFA2C692FDB75DC8FD5B" box="[714,757,631,655]" country="Japan" httpUri="http://grbio.org/cool/7zx4-v798" name="Food Research Institute, Ministry of Agriculture, Forestry and Fisheries" pageId="11" pageNumber="60">FRI</collectionCode>
<specimenCode id="5B05542A987CFFA2C6A4FDB75C64FD5B" box="[764,857,631,655]" collectionCode="FRI" country="Japan" httpUri="http://grbio.org/cool/7zx4-v798" name="Food Research Institute, Ministry of Agriculture, Forestry and Fisheries" pageId="11" pageNumber="60">CR0011</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0B1CFC51987CFFA2C685FD5A5B56FBD9" blockId="11.[707,1237,215,1799]" pageId="11" pageNumber="60">
<emphasis id="39D72043987CFFA2C685FD5A5CC5FD66" box="[733,1016,666,690]" italics="true" pageId="11" pageNumber="60">Other non-type materials</emphasis>
.
<materialsCitation id="BBCBF60C987CFFA2C041FD5A5B5AFBD9" ID-GBIF-Occurrence="4899861303" collectingDate="2005-07" collectingDateMax="2008-11-25" collectingDateMin="2005-07" collectorName="Y. Hanamura &amp; Faizul" country="Malaysia" location="Mouth of Merbok Mangrove Estuary" pageId="11" pageNumber="60" specimenCount="33" specimenCount-female="9" specimenCount-juvenile="3" specimenCount-male="21">
<collectingCountry id="73B4BCC1987CFFA2C041FD5A5B43FD66" box="[1049,1150,666,690]" name="Malaysia" pageId="11" pageNumber="60">Malaysia</collectingCountry>
.
<emphasis id="39D72043987CFFA2C0D6FD5A5B36FD00" italics="true" pageId="11" pageNumber="60">
<location id="0E7CAA8A987CFFA2C0D6FD5A5B36FD00" LSID="urn:lsid:plazi:treatment:830A4D47987CFFA7C4B7FB355CD0FD23:0E7CAA8A987CFFA2C0D6FD5A5B36FD00" country="Malaysia" name="Mouth of Merbok Mangrove Estuary" pageId="11" pageNumber="60">Mouth of Merbok Mangrove Estuary</location>
</emphasis>
:
<specimenCount id="1DA537D8987CFFA2C045FD7C5B4EFD01" box="[1053,1139,700,725]" count="5" pageId="11" pageNumber="60" type="male">5 males</specimenCount>
(cl 4.5 5.0 mm), sledge,
<date id="7F1DDA91987CFFA2C723FD1F5CD5FD23" box="[891,1000,735,759]" pageId="11" pageNumber="60" value="2005-07">
<collectingDate id="6F592379987CFFA2C723FD1F5CD5FD23" box="[891,1000,735,759]" pageId="11" pageNumber="60" value="2005-07">July 2005</collectingDate>
</date>
, coll.
<collectorName id="A6569987987CFFA2C072FD205B8AFD23" box="[1066,1207,736,759]" pageId="11" pageNumber="60">Y. Hanamura</collectorName>
&amp;
<collectorName id="A6569987987CFFA2C69BFCC25C3BFCCE" box="[707,774,770,794]" pageId="11" pageNumber="60">Faizul</collectorName>
(NSMT-Cr 31626):
<emphasis id="39D72043987CFFA2C05AFCC35C2BFCE8" italics="true" pageId="11" pageNumber="60">Matang Mangrove Estuary</emphasis>
.
<specimenCount id="1DA537D8987CFFA2C77EFCE55C46FCE9" box="[806,891,805,829]" count="2" pageId="11" pageNumber="60" type="male">2 males</specimenCount>
(cl 4.8, 5.0 mm),
<specimenCount id="1DA537D8987CFFA2C01CFCE55B93FCE9" box="[1092,1198,805,829]" count="2" pageId="11" pageNumber="60" type="female">2 females</specimenCount>
(cl 5.9,
<quantity id="CC5B51B4987CFFA2C6A4FC875C73FC8B" box="[764,846,839,863]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="11" pageNumber="60" unit="mm" value="6.9">6.9 mm</quantity>
), sledge,
<date id="7F1DDA91987CFFA2C79FFC875B54FC8B" box="[967,1129,839,863]" pageId="11" pageNumber="60" value="2005-08-18">
<collectingDate id="6F592379987CFFA2C79FFC875B54FC8B" box="[967,1129,839,863]" pageId="11" pageNumber="60" value="2005-08-18">18 Aug 2005</collectingDate>
</date>
, coll.
<collectorName id="A6569987987CFFA2C0E5FC885C0FFC55" pageId="11" pageNumber="60">Y. Hanamura</collectorName>
&amp;
<collectorName id="A6569987987CFFA2C70DFCAA5CA5FC56" box="[853,920,874,898]" pageId="11" pageNumber="60">Faizul</collectorName>
(transferred from formalin to ethanol) (NSMT-Cr 31627):
<specimenCount id="1DA537D8987CFFA2C051FC4D5B52FC71" box="[1033,1135,909,933]" count="14" pageId="11" pageNumber="60" type="male">14 males</specimenCount>
(cl
<quantity id="CC5B51B4987CFFA2C0FDFC4D5C29FC13" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.6" metricValueMin="3.7" pageId="11" pageNumber="60" unit="mm" value="4.15" valueMax="4.6" valueMin="3.7">3.7 4.6 mm</quantity>
),
<specimenCount id="1DA537D8987CFFA2C773FC705CA8FC13" box="[811,917,943,967]" count="7" pageId="11" pageNumber="60" type="female">7 females</specimenCount>
(cl
<quantity id="CC5B51B4987CFFA2C79EFC6F5B7BFC13" box="[966,1094,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="6.4" metricValueMin="3.7" pageId="11" pageNumber="60" unit="mm" value="5.05" valueMax="6.4" valueMin="3.7">3.76.4 mm</quantity>
),
<specimenCount id="1DA537D8987CFFA2C004FC6F5BE9FC13" box="[1116,1236,943,967]" count="3" pageId="11" pageNumber="60" type="juvenile">3 juveniles</specimenCount>
(cl
<quantity id="CC5B51B4987CFFA2C6B3FC125C56FC3E" box="[747,875,978,1002]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.3" metricValueMin="1.5" pageId="11" pageNumber="60" unit="mm" value="1.9" valueMax="2.3" valueMin="1.5">1.52.3 mm</quantity>
), sledge,
<date id="7F1DDA91987CFFA2C783FC125B4EFC3E" box="[987,1139,978,1002]" pageId="11" pageNumber="60" value="2008-11-25">
<collectingDate id="6F592379987CFFA2C783FC125B4EFC3E" box="[987,1139,978,1002]" pageId="11" pageNumber="60" value="2008-11-25">25 Nov 2008</collectingDate>
</date>
, coll.
<collectorName id="A6569987987CFFA2C0E5FC125C0FFBD8" pageId="11" pageNumber="60">Y. Hanamura</collectorName>
&amp;
<collectorName id="A6569987987CFFA2C70CFC355CAAFBD9" box="[852,919,1013,1037]" pageId="11" pageNumber="60">Faizul</collectorName>
(NSMT-Cr 31628)
</materialsCitation>
.
</paragraph>
<paragraph id="0B1CFC51987CFFA2C685FBD75B5DFB2B" blockId="11.[707,1237,215,1799]" pageId="11" pageNumber="60">
<materialsCitation id="BBCBF60C987CFFA2C685FBD75B61FB2B" ID-GBIF-Occurrence="4899861302" collectingDate="2019-09-19" collectorName="E. Hardianto &amp; H. Imai" country="Indonesia" county="Setrap River Estuary" latitude="-30.800083" location="River Estuary" longLatPrecision="1" longitude="115.522285" municipality="Setrap River" pageId="11" pageNumber="60" specimenCount="19" specimenCount-female="13" specimenCount-male="2" stateProvince="Kalimantan Utara">
<collectingCountry id="73B4BCC1987CFFA2C685FBD75C75FBFB" box="[733,840,1047,1071]" name="Indonesia" pageId="11" pageNumber="60">Indonesia</collectingCountry>
.
<emphasis id="39D72043987CFFA2C700FBD85B7EFBFB" box="[856,1091,1048,1071]" italics="true" pageId="11" pageNumber="60">
<collectingCounty id="E27D84DD987CFFA2C700FBD85B7EFBFB" box="[856,1091,1048,1071]" pageId="11" pageNumber="60">
<collectingMunicipality id="EB78662B987CFFA2C700FBD85CD9FBFB" box="[856,996,1048,1071]" pageId="11" pageNumber="60">
Setrap
<location id="0E7CAA8A987CFFA2C7F2FBD85B7EFBFB" LSID="urn:lsid:plazi:treatment:830A4D47987CFFA7C4B7FB355CD0FD23:0E7CAA8A987CFFA2C7F2FBD85B7EFBFB" box="[938,1091,1048,1071]" country="Indonesia" county="Setrap River Estuary" latitude="-30.800083" longLatPrecision="1" longitude="115.522285" municipality="Setrap River" name="River Estuary" pageId="11" pageNumber="60" stateProvince="Kalimantan Utara">River Estuary</location>
</collectingMunicipality>
</collectingCounty>
</emphasis>
,
<emphasis id="39D72043987CFFA2C00EFBD75C3BFB86" italics="true" pageId="11" pageNumber="60">
<collectingRegion id="C96732B3987CFFA2C00EFBD75BE9FBFB" box="[1110,1236,1047,1071]" country="Indonesia" name="Kalimantan Utara" pageId="11" pageNumber="60">Kalimantan</collectingRegion>
<location id="0E7CAA8A987CFFA2C69BFBFA5C3BFB86" LSID="urn:lsid:plazi:treatment:830A4D47987CFFA7C4B7FB355CD0FD23:0E7CAA8A987CFFA2C69BFBFA5C3BFB86" box="[707,774,1082,1106]" country="Indonesia" county="Setrap River Estuary" latitude="-30.800083" longLatPrecision="1" longitude="115.522285" municipality="Setrap River" name="Island" pageId="11" pageNumber="60" stateProvince="Kalimantan Utara">Island</location>
</emphasis>
:
<specimenCount id="1DA537D8987CFFA2C74FFBFA5C51FB86" box="[791,876,1082,1106]" count="2" pageId="11" pageNumber="60" type="male">2 males</specimenCount>
(cl 6.5,
<quantity id="CC5B51B4987CFFA2C797FBFA5B1DFB86" box="[975,1056,1082,1106]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="60" unit="mm" value="7.5">7.5 mm</quantity>
),
<specimenCount id="1DA537D8987CFFA2C06EFBFA5B93FB86" box="[1078,1198,1082,1106]" count="13" pageId="11" pageNumber="60" type="female">13 females</specimenCount>
(cl 7.010.0 mm),
<geoCoordinate id="6E979A96987CFFA2C731FB9D5CD1FBA1" box="[873,1004,1115,1141]" degrees="30" direction="south" minutes="48.005" orientation="latitude" pageId="11" pageNumber="60" precision="1" value="-30.800083">30°48.005S</geoCoordinate>
,
<geoCoordinate id="6E979A96987CFFA2C7A5FB9C5BADFBA0" box="[1021,1168,1115,1141]" degrees="115" direction="east" minutes="31.337" orientation="longitude" pageId="11" pageNumber="60" precision="1" value="115.522285">115°31.337E</geoCoordinate>
,
<collectingMethod id="D2E28446987CFFA2C0F8FB9D5DDBFB43" pageId="11" pageNumber="60">hand net</collectingMethod>
,
<date id="7F1DDA91987CFFA2C6ABFBBF5CB5FB43" box="[755,904,1151,1175]" pageId="11" pageNumber="60" value="2019-09-19">
<collectingDate id="6F592379987CFFA2C6ABFBBF5CB5FB43" box="[755,904,1151,1175]" pageId="11" pageNumber="60" value="2019-09-19">19 Sept 2019</collectingDate>
</date>
, coll.
<collectorName id="A6569987987CFFA2C795FB405B64FB43" box="[973,1113,1151,1175]" pageId="11" pageNumber="60">E. Hardianto</collectorName>
&amp;
<collectorName id="A6569987987CFFA2C0D8FB405BE9FB43" box="[1152,1236,1151,1175]" pageId="11" pageNumber="60">H. Imai</collectorName>
(NSMT-Cr 31631);
<specimenCount id="1DA537D8987CFFA2C7F9FB625B14FB6E" box="[929,1065,1186,1210]" count="4" pageId="11" pageNumber="60" type="generic">4 specimens</specimenCount>
used for DNA analysis (sex and size undetermined), data same as previous (NSMT-Cr 3163231635)
</materialsCitation>
.
</paragraph>
<paragraph id="0B1CFC51987CFFA2C685FACA5C2BFA79" blockId="11.[707,1237,215,1799]" pageId="11" pageNumber="60">
<materialsCitation id="BBCBF60C987CFFA2C685FACA5C2FFA79" ID-GBIF-Occurrence="4899861313" collectingDate="2016-06" collectorName="K. Nakahara" country="Thailand" location="Gulf" municipality="Bam Bo Village" pageId="11" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Samut Prakan">
<collectingCountry id="73B4BCC1987CFFA2C685FACA5C02FAF6" box="[733,831,1290,1314]" name="Thailand" pageId="11" pageNumber="60">Thailand</collectingCountry>
.
<emphasis id="39D72043987CFFA2C70EFACB5B2BFAF6" box="[854,1046,1290,1314]" italics="true" pageId="11" pageNumber="60">
<collectingMunicipality id="EB78662B987CFFA2C70EFACB5B2BFAF6" box="[854,1046,1290,1314]" pageId="11" pageNumber="60">Bam Bo Village</collectingMunicipality>
</emphasis>
,
<emphasis id="39D72043987CFFA2C068FACA5CB4FA91" italics="true" pageId="11" pageNumber="60">
<collectingRegion id="C96732B3987CFFA2C068FACA5BE9FAF6" box="[1072,1236,1290,1314]" country="Thailand" name="Samut Prakan" pageId="11" pageNumber="60">Samut Prakan</collectingRegion>
(
<location id="0E7CAA8A987CFFA2C692FAED5DC6FA91" LSID="urn:lsid:plazi:treatment:830A4D47987CFFA7C4B7FB355CD0FD23:0E7CAA8A987CFFA2C692FAED5DC6FA91" box="[714,763,1325,1349]" country="Thailand" municipality="Bam Bo Village" name="Gulf" pageId="11" pageNumber="60" stateProvince="Samut Prakan">Gulf</location>
of
<collectingCountry id="73B4BCC1987CFFA2C778FAED5CBDFA91" box="[800,896,1325,1349]" name="Thailand" pageId="11" pageNumber="60">Thailand</collectingCountry>
<emphasis id="39D72043987CFFA2C7D8FAED5CB4FA91" bold="true" box="[896,905,1325,1349]" italics="true" pageId="11" pageNumber="60">)</emphasis>
</emphasis>
:
<specimenCount id="1DA537D8987CFFA2C7C1FAED5CDFFA91" box="[921,994,1325,1349]" count="1" pageId="11" pageNumber="60" type="male">1 male</specimenCount>
(cl
<quantity id="CC5B51B4987CFFA2C056FAED5B5DFA91" box="[1038,1120,1325,1349]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="11" pageNumber="60" unit="mm" value="4.7">4.7 mm</quantity>
), commercial catch,
<date id="7F1DDA91987CFFA2C763FA905C90FAB3" box="[827,941,1359,1383]" pageId="11" pageNumber="60" value="2016-06">
<collectingDate id="6F592379987CFFA2C763FA905C90FAB3" box="[827,941,1359,1383]" pageId="11" pageNumber="60" value="2016-06">June 2016</collectingDate>
</date>
, coll.
<collectorName id="A6569987987CFFA2C7AAFA905B43FAB3" box="[1010,1150,1359,1383]" pageId="11" pageNumber="60">K. Nakahara</collectorName>
, part of appendages used for DNA analysis (NSMT-Cr 31636)
</materialsCitation>
.
</paragraph>
<paragraph id="0B1CFC51987CFFA2C685FA785CF6F94B" blockId="11.[707,1237,215,1799]" pageId="11" pageNumber="60">
<emphasis id="39D72043987CFFA2C685FA785C66FA1B" box="[733,859,1464,1487]" italics="true" pageId="11" pageNumber="60">Description</emphasis>
.
<emphasis id="39D72043987CFFA2C725FA775C89FA1B" box="[893,948,1463,1487]" italics="true" pageId="11" pageNumber="60">Male</emphasis>
. Rostrum (
<figureCitation id="9398E0D4987CFFA2C076FA775BABFA1B" box="[1070,1174,1463,1487]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="11" pageNumber="60">Fig. 5a, b</figureCitation>
) sub-triangular, elevated slightly from dorsal margin of carapace, with sharply pointed terminal tooth and small anterior tooth in addition to somewhat larger posterior tooth on its dorsal margin. Carapace entirely smooth on surface, except for post-orbital and hepatic teeth.
</paragraph>
<paragraph id="0B1CFC51987CFFA5C685F96A5FC0F930" blockId="11.[707,1237,215,1799]" lastBlockId="12.[140,669,1567,1800]" lastPageId="12" lastPageNumber="61" pageId="11" pageNumber="60">
Abdomen with first somite possessing procurved sharp tooth at median part of ventral sternite between right and left pleopods. Sixth somite (
<figureCitation id="9398E0D4987BFFA5C4BFF9DF5E04F9E3" box="[231,313,1567,1591]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="12" pageNumber="61">Fig. 5c</figureCitation>
) twice length of fifth, postero-dorsal part ending in small tooth, no red spot on postero-ventral part. Telson (
<figureCitation id="9398E0D4987BFFA5C594F9A55D07F9A9" box="[460,570,1636,1661]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="12" pageNumber="61">Fig. 5c, d</figureCitation>
) 0.70.8 times as long as sixth somite, posterior end narrowly rounded or sub-triangular with rounded distal part.
</paragraph>
<caption id="5FDCACD9987BFFA5C4FBFA905C19FA0F" ID-DOI="http://doi.org/10.5281/zenodo.12572354" ID-Zenodo-Dep="12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="12" pageNumber="61" startId="12.[163,196,1360,1382]" targetBox="[277,1104,214,1325]" targetPageId="12" targetType="figure">
<paragraph id="0B1CFC51987BFFA5C4FBFA905C19FA0F" blockId="12.[163,1213,1360,1499]" pageId="12" pageNumber="61">
Fig. 5.
<taxonomicName id="CCA387D2987BFFA5C4A1FA905D8EFAB2" authority="Hanamura, Imai and Hardianto" authorityName="Hanamura, Imai and Hardianto" authorityYear="2024" box="[249,691,1360,1382]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="61" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="39D72043987BFFA5C4A1FA905E41FAB2" box="[249,380,1360,1382]" italics="true" pageId="12" pageNumber="61">Acetes omorii</emphasis>
Hanamura, Imai and Hardianto
</taxonomicName>
<taxonomicNameLabel id="22E49D38987BFFA5C6E6FA905C36FAB1" box="[702,779,1360,1381]" pageId="12" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
(ag) holotype male (cl 5.3 mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0 mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body.
</paragraph>
</caption>
<paragraph id="0B1CFC51987BFFA5C4FFF9305CDDF9E3" blockId="12.[140,669,1567,1800]" box="[167,992,1567,1800]" lastBlockId="12.[707,1236,1567,1800]" pageId="12" pageNumber="61">
Eye (
<figureCitation id="9398E0D4987BFFA5C4BAF92F5E6DF8D3" box="[226,336,1775,1799]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="12" pageNumber="61">Fig. 5a, b</figureCitation>
) with well pigmented cornea, latter wider than eye-stalk.
</paragraph>
<paragraph id="0B1CFC51987BFFA4C685F9825C48FAF6" blockId="12.[707,1236,1567,1800]" lastBlockId="13.[707,1237,1290,1799]" lastPageId="13" lastPageNumber="62" pageId="12" pageNumber="61">
Antennule (
<figureCitation id="9398E0D4987BFFA5C707F9825CF3F98E" box="[863,974,1602,1626]" captionStart="Fig" captionStartId="15.[163,196,1358,1380]" captionTargetBox="[275,1103,215,1323]" captionTargetId="figure-81@15.[272,1104,213,1324]" captionTargetPageId="15" captionText="Fig. 7. Acetes spiniger Hansen, 1919. (ae) male (cl 4.3mm) and (fi) female (cl. 5.8mm) from off Kuala Selan- gor, Malaysia (NSMT-Cr 31637): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, right left lower antennular flagellum; d, basal segments of third pereopod and genital coxa, ventral; e, capitulum of right petasma; f, anterior part of cephalothorax, lateral; g, basal segments of third pereopod and third and fourth thoracic sternites, ventral; h, lateral profile showing channels and sternites between third and fourth thoracic somites, i, posterior part of telson, dorsal. Abbreviations: gc, genital coxa; iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572364" httpUri="https://zenodo.org/record/12572364/files/figure.png" pageId="12" pageNumber="61">Fig. 7a, b</figureCitation>
) noticeably elongate in mature males, particularly in third segment of peduncle, 0.700.86 times as long as combined length of first and second segments; lower flagellum (
<figureCitation id="9398E0D4987BFFA5C6A3F90D5C5AF931" box="[763,871,1740,1765]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="12" pageNumber="61">Figs 5a, b</figureCitation>
,
<figureCitation id="9398E0D4987BFFA5C72AF90D5CB0F931" box="[882,909,1741,1765]" captionStart="Fig" captionStartId="13.[163,196,1092,1114]" captionTargetBox="[273,1102,215,1048]" captionTargetId="figure-225@13.[272,1104,213,1049]" captionTargetPageId="13" captionText="Fig. 6. Acetes omorii Hanamura, Imai &amp; Hardianto sp. nov. (a) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) (NSMT-Cr 31622) from Matang Mangrove, Malaysia: a, b, right lower antennular flagellum of male and female, respectively; c, basal segments of third pereopod and third and fourth abdominal sternites, ventral; d, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572358" httpUri="https://zenodo.org/record/12572358/files/figure.png" pageId="12" pageNumber="61">6a</figureCitation>
) modified into clasping organ, commonly composed of 12-segmented; first segment of trunk not noticeably long, almost uniform in width through length and utmost about twice length of its width, showing weakly oblique conjunction with second one; major clasping spine moderately stout, reaching as far as mid-length of fourth segment of main branch, with 2 aligned rows of minute serrated denticles, and rudimentary clasping spine present near base of major one; first sub-segment of main branch with single basal and single distal setal-spines, second and third each with single distal and/or sub-distal setal-spine, fourth segment 1.21.5 times as long as combined length of second and third segments, and usually bearing 9 or 10 setal-spines on its entire margin, fifth segment with 13 setal-spines.
</paragraph>
<caption id="5FDCACD9987AFFA4C4FBFB845B3DFB1B" ID-DOI="http://doi.org/10.5281/zenodo.12572358" ID-Zenodo-Dep="12572358" httpUri="https://zenodo.org/record/12572358/files/figure.png" pageId="13" pageNumber="62" startId="13.[163,196,1092,1114]" targetBox="[273,1102,215,1048]" targetPageId="13" targetType="figure">
<paragraph id="0B1CFC51987AFFA4C4FBFB845B3DFB1B" blockId="13.[163,1214,1092,1231]" pageId="13" pageNumber="62">
Fig. 6.
<taxonomicName id="CCA387D2987AFFA4C4A1FB845DA9FB8E" authority="Hanamura, Imai &amp; Hardianto" authorityName="Hanamura, Imai &amp; Hardianto" authorityYear="2024" box="[249,660,1092,1114]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="39D72043987AFFA4C4A1FB845E44FB8D" box="[249,377,1092,1113]" italics="true" pageId="13" pageNumber="62">Acetes omorii</emphasis>
Hanamura, Imai &amp; Hardianto
</taxonomicName>
<taxonomicNameLabel id="22E49D38987AFFA4C6C4FB845DDBFB8D" box="[668,742,1092,1113]" pageId="13" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
(a) holotype male (cl 5.3 mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0 mm) (NSMT-Cr 31622) from Matang Mangrove, Malaysia: a, b, right lower antennular flagellum of male and female, respectively; c, basal segments of third pereopod and third and fourth abdominal sternites, ventral; d, lateral profile showing channels and sternites between third and fourth thoracic somites. Abbreviations: iiist, ivst, third and fourth thoracic sternites, respectively.
</paragraph>
</caption>
<paragraph id="0B1CFC51987AFFA4C685FAED5C9FFA79" blockId="13.[707,1237,1290,1799]" pageId="13" pageNumber="62">
Antennal scale (
<figureCitation id="9398E0D4987AFFA4C7CFFAED5CD4FA90" box="[919,1001,1324,1349]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="13" pageNumber="62">Fig. 5a</figureCitation>
) 0.700.79 times as long as carapace, slightly but distinctly extending beyond anterior margin of second segment of antennular peduncle.
</paragraph>
<paragraph id="0B1CFC51987AFFA7C685FA775E52FE48" blockId="13.[707,1237,1290,1799]" lastBlockId="14.[140,670,215,1799]" lastPageId="14" lastPageNumber="63" pageId="13" pageNumber="62">
External maxilliped and pereopods varying in length depending on body size. Third maxilliped in larger males extending as far as anterior end of antennular peduncle, distal segment about three-fourths times as long as penultimate. First pereopod in larger males extending as far as cornea. Second pereopod in larger males reaching as far as anterior end of antennal scale. Third pereopod (
<figureCitation id="9398E0D4987AFFA4C693F90D5C27F930" box="[715,794,1740,1765]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="13" pageNumber="62">Fig. 5e</figureCitation>
) in larger males extending anterior end of antennular peduncle, distinct sharp tooth present at disto-mesial part of basis; coxa with anteriorly curving on its inner margin. Genital coxa (
<figureCitation id="9398E0D49879FFA7C4CCFEDD5FD8FEE0" box="[148,229,284,309]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="14" pageNumber="63">Fig. 5e</figureCitation>
) often with slender or sub-acute distal part in lateral aspect, while sub-rectangular in ventral aspect, with widely truncated or widely sinuous anterior part.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFE685E1AFD5B" blockId="14.[140,670,215,1799]" pageId="14" pageNumber="63">
Petasma (
<figureCitation id="9398E0D49879FFA7C542FE675E52FE6B" box="[282,367,423,447]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="14" pageNumber="63">Fig. 5f</figureCitation>
) with brunt and distally rounded capitulum bearing numerous hooks along lateral surface as well as distal part; processus ventralis slender, reaching as far as distal margin of capitulum; no trace of pars astringen, Appendix masculina (
<figureCitation id="9398E0D49879FFA7C524FD955EF2FDB9" box="[380,463,596,621]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="14" pageNumber="63">
Fig.
<quantity id="CC5B51B49879FFA7C5E9FD945EF2FDB9" box="[433,463,596,621]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="14" pageNumber="63" unit="g" value="5.0">5 g</quantity>
</figureCitation>
) with 2 or 3 short hooks distally.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFD5A5E8EFCE9" blockId="14.[140,670,215,1799]" pageId="14" pageNumber="63">
Endopod (
<figureCitation id="9398E0D49879FFA7C540FD5A5E5BFD66" box="[280,358,666,690]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="14" pageNumber="63">Fig. 5c</figureCitation>
) of uropod with 16 irregularly-shaped red-spots in fresh materials: exopod with small tooth on lateral margin demarcating non-ciliated and ciliated parts, latter about 0.7 0.8 times as long as former.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFC875D0DFC8B" blockId="14.[140,670,215,1799]" box="[167,560,839,863]" pageId="14" pageNumber="63">
<emphasis id="39D720439879FFA7C4FFFC875FCAFC8B" box="[167,247,839,863]" italics="true" pageId="14" pageNumber="63">Female</emphasis>
. Usually larger than males.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFCAA5E53FBFB" blockId="14.[140,670,215,1799]" pageId="14" pageNumber="63">
Antennule (
<figureCitation id="9398E0D49879FFA7C57DFCAA5E49FC56" box="[293,372,874,898]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="14" pageNumber="63">Fig. 5h</figureCitation>
) not elongated as in males, first segment longest, and third segments 0.39 0.46 times as long as combined length of second and third segments; lower flagellum (
<figureCitation id="9398E0D49879FFA7C67EFC125D4BFC3E" box="[550,630,978,1002]" captionStart="Fig" captionStartId="13.[163,196,1092,1114]" captionTargetBox="[273,1102,215,1048]" captionTargetId="figure-225@13.[272,1104,213,1049]" captionTargetPageId="13" captionText="Fig. 6. Acetes omorii Hanamura, Imai &amp; Hardianto sp. nov. (a) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) (NSMT-Cr 31622) from Matang Mangrove, Malaysia: a, b, right lower antennular flagellum of male and female, respectively; c, basal segments of third pereopod and third and fourth abdominal sternites, ventral; d, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572358" httpUri="https://zenodo.org/record/12572358/files/figure.png" pageId="14" pageNumber="63">Fig. 6b</figureCitation>
) as long as third segment of peduncle, constituted of 1820 sub-segments.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFBFA5FC9FB6E" blockId="14.[140,670,215,1799]" pageId="14" pageNumber="63">
Antennal scale (
<figureCitation id="9398E0D49879FFA7C538FBFA5E8EFB86" box="[352,435,1082,1106]" captionStart="Fig" captionStartId="12.[163,196,1360,1382]" captionTargetBox="[277,1104,214,1325]" captionTargetId="figure-109@12.[272,1104,213,1327]" captionTargetPageId="12" captionText="Fig. 5. Acetes omorii Hanamura, Imai and Hardianto sp. nov. (ag) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) from Matang Mangrove, Malaysia (NSMT-Cr 31622): a, cephalothorax, lateral; b, same, anterior part enlarged; c, right posterior part of abdomen, lateral; d, posterior part of telson, dorsal; e, basal segments of third pereopod and genital coxa; f, capitulum of petasma, posterior; g, right endopod of second pleopod, mesial; h, anterior part of body." figureDoi="http://doi.org/10.5281/zenodo.12572354" httpUri="https://zenodo.org/record/12572354/files/figure.png" pageId="14" pageNumber="63">Fig. 5h</figureCitation>
) 0.670.74 times as long as carapace, extending to mid-length to distal one-third of third segment of antennular peduncle.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFB055FF2FA91" blockId="14.[140,670,215,1799]" pageId="14" pageNumber="63">
Third and fourth thoracic sternites forming longitudinal groove rather deep and narrow, median swelling steep and well pronounced (
<figureCitation id="9398E0D49879FFA7C62AFACA5FFFFA91" captionStart="Fig" captionStartId="13.[163,196,1092,1114]" captionTargetBox="[273,1102,215,1048]" captionTargetId="figure-225@13.[272,1104,213,1049]" captionTargetPageId="13" captionText="Fig. 6. Acetes omorii Hanamura, Imai &amp; Hardianto sp. nov. (a) holotype male (cl 5.3mm) (NSMT-Cr 31621), and (bd) paratype female (cl 6.0mm) (NSMT-Cr 31622) from Matang Mangrove, Malaysia: a, b, right lower antennular flagellum of male and female, respectively; c, basal segments of third pereopod and third and fourth abdominal sternites, ventral; d, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: iiist, ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572358" httpUri="https://zenodo.org/record/12572358/files/figure.png" pageId="14" pageNumber="63">Fig. 6c, d</figureCitation>
).
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFA8F5EF0FA79" blockId="14.[140,670,215,1799]" pageId="14" pageNumber="63">
<emphasis id="39D720439879FFA7C4FFFA8F5E31FAB3" box="[167,268,1359,1383]" italics="true" pageId="14" pageNumber="63">Body size</emphasis>
. Largest male cl
<quantity id="CC5B51B49879FFA7C5A9FA8F5D7CFAB3" box="[497,577,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="63" unit="mm" value="7.5">7.5 mm</quantity>
, female 10.0 mm, recorded from south-eastern part of
<collectingRegion id="C96732B39879FFA7C4D4FA555E35FA79" box="[140,264,1429,1453]" country="Indonesia" name="Kalimantan Utara" pageId="14" pageNumber="63">Kalimantan</collectingRegion>
Island,
<collectingCountry id="73B4BCC19879FFA7C505FA555EF5FA79" box="[349,456,1429,1453]" name="Indonesia" pageId="14" pageNumber="63">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFA785E51FA26" blockId="14.[140,670,215,1799]" pageId="14" pageNumber="63">
<materialsCitation id="BBCBF60C9879FFA7C4FFFA785E55FA26" ID-GBIF-Occurrence="4899861312" country="Malaysia" location="Matang Mangrove Estuary" municipality="Peninsular" pageId="14" pageNumber="63" specimenCount="1" typeStatus="holotype">
<emphasis id="39D720439879FFA7C4FFFA785E12FA1B" box="[167,303,1463,1487]" italics="true" pageId="14" pageNumber="63">
<typeStatus id="D41842F39879FFA7C4FFFA785FE4FA1B" box="[167,217,1464,1487]" pageId="14" pageNumber="63">Type</typeStatus>
locality
</emphasis>
.
<location id="0E7CAA8A9879FFA7C508FA785DA7FA04" LSID="urn:lsid:plazi:treatment:830A4D47987CFFA7C4B7FB355CD0FD23:0E7CAA8A9879FFA7C508FA785DA7FA04" box="[336,666,1464,1488]" country="Malaysia" municipality="Peninsular" name="Matang Mangrove Estuary" pageId="14" pageNumber="63">Matang Mangrove Estuary</location>
,
<collectingMunicipality id="EB78662B9879FFA7C4D4FA1A5FC0FA26" box="[140,253,1498,1522]" pageId="14" pageNumber="63">Peninsular</collectingMunicipality>
<collectingCountry id="73B4BCC19879FFA7C55BFA1A5E55FA26" box="[259,360,1498,1522]" name="Malaysia" pageId="14" pageNumber="63">Malaysia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFFA3D5E11F9A8" blockId="14.[140,670,215,1799]" pageId="14" pageNumber="63">
<emphasis id="39D720439879FFA7C4FFFA3D5E24F9C1" box="[167,281,1533,1557]" italics="true" pageId="14" pageNumber="63">Etymology</emphasis>
. This species is named in honour of the late Dr. Makoto
<collectingRegion id="C96732B39879FFA7C5D3F9DF5EECF9E3" box="[395,465,1567,1591]" country="Japan" name="Aomori" pageId="14" pageNumber="63">Omori</collectingRegion>
(1937-2022), who first discussed the two morphological forms in
<taxonomicName id="CCA387D29879FFA7C4D4F9A55E18F9A9" authorityName="H. Milne Edwards" authorityYear="1830" box="[140,293,1637,1661]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D720439879FFA7C4D4F9A55E18F9A9" box="[140,293,1637,1661]" italics="true" pageId="14" pageNumber="63">Acetes indicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0B1CFC519879FFA7C4FFF9475C8BFEC5" blockId="14.[140,670,215,1799]" lastBlockId="14.[707,1236,215,759]" pageId="14" pageNumber="63">
<emphasis id="39D720439879FFA7C4FFF9475E39F94B" box="[167,260,1671,1695]" italics="true" pageId="14" pageNumber="63">Remarks</emphasis>
.
<bibRefCitation id="6F3281A09879FFA7C57DF9475D6FF94B" author="Tirmizi, N. &amp; N. A. Ghani" box="[293,594,1671,1695]" pageId="14" pageNumber="63" pagination="44 - 53" refId="ref11693" refString="Tirmizi, N. and N. A. Ghani 1982. New distributional records for three species of Acetes (Decapoda, Sergestidae). Crustaceana 42: 44 - 53." type="journal article" year="1982">Tirmizi and Ghani (1982)</bibRefCitation>
noted that specimens of
<taxonomicName id="CCA387D29879FFA7C517F96A5E86F916" authorityName="H. Milne Edwards" authorityYear="1830" box="[335,443,1706,1730]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D720439879FFA7C517F96A5E86F916" box="[335,443,1706,1730]" italics="true" pageId="14" pageNumber="63">A. indicus</emphasis>
</taxonomicName>
collected from
<collectingCountry id="73B4BCC19879FFA7C63DF96A5F8AF930" name="Pakistan" pageId="14" pageNumber="63">Pakistan</collectingCountry>
resemble Omori`s large form. However, we have the impression that their morphological features appeared to be similar to
<taxonomicName id="CCA387D29879FFA7C04EFF185BB0FF3B" authorityName="Hansen" authorityYear="1919" box="[1046,1165,216,239]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="39D720439879FFA7C04EFF185BB0FF3B" box="[1046,1165,216,239]" italics="true" pageId="14" pageNumber="63">A. spiniger</emphasis>
</taxonomicName>
rather than
<taxonomicName id="CCA387D29879FFA7C6AFFF3A5C61FEC6" authority="Alamri &amp; Alotaiby, 2024" authorityName="Alamri &amp; Alotaiby" authorityYear="2024" box="[759,860,250,274]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="39D720439879FFA7C6AFFF3A5C61FEC6" box="[759,860,250,274]" italics="true" pageId="14" pageNumber="63">A. omorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22E49D389879FFA7C73BFF3A5C8BFEC5" box="[867,950,250,273]" pageId="14" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="0B1CFC519879FFA7C685FEDD5C5CFE6B" blockId="14.[707,1236,215,759]" pageId="14" pageNumber="63">
The specimens reported from
<collectingCountry id="73B4BCC19879FFA7C011FEDD5B95FEE1" box="[1097,1192,285,309]" name="Thailand" pageId="14" pageNumber="63">Thailand</collectingCountry>
by
<bibRefCitation id="6F3281A09879FFA7C69BFEFF5B09FE83" author="Perez Farfante, I. &amp; B. Kensley" box="[707,1076,319,343]" pageId="14" pageNumber="63" pagination="1 - 233" refId="ref11352" refString="Perez Farfante, I. and B. Kensley 1997. Penaeoid and sergestoid shrimps and prawns of the world: keys and diagnoses for the families and genera. Memoires du Museum National d`Histoire Naturelle 175: 1 - 233." type="journal article" year="1997">Pérez Farfante and Kensley (1997</bibRefCitation>
; figs 128130) are regarded to be
<taxonomicName id="CCA387D29879FFA7C7C2FEA25B3EFEAE" authorityName="Alamri &amp; Alotaiby" authorityYear="2024" box="[922,1027,354,378]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="omorii">
<emphasis id="39D720439879FFA7C7C2FEA25B3EFEAE" box="[922,1027,354,378]" italics="true" pageId="14" pageNumber="63">A. omorii</emphasis>
</taxonomicName>
, because the male lower antennular flagellum agrees well with that of this species.
</paragraph>
<paragraph id="0B1CFC519879FFA7C685FE0A5CD0FD23" blockId="14.[707,1236,215,759]" pageId="14" pageNumber="63">
<emphasis id="39D720439879FFA7C685FE0A5C62FE36" box="[733,863,458,482]" italics="true" pageId="14" pageNumber="63">Distribution</emphasis>
.
<taxonomicName id="CCA387D29879FFA7C7D9FE0A5B28FE36" authority="Alamri &amp; Alotaiby, 2024" authorityName="Alamri &amp; Alotaiby" authorityYear="2024" box="[897,1045,458,482]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="39D720439879FFA7C7D9FE0A5B28FE36" box="[897,1045,458,482]" italics="true" pageId="14" pageNumber="63">Acetes omorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22E49D389879FFA7C045FE0A5B4DFE35" box="[1053,1136,458,481]" pageId="14" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
has been collected with certainty in the coastal waters of South-East Asia, from
<collectingRegion id="C96732B39879FFA7C799FDCF5B00FDF3" box="[961,1085,527,551]" country="Indonesia" name="Kalimantan Utara" pageId="14" pageNumber="63">Kalimantan</collectingRegion>
Island,
<collectingCountry id="73B4BCC19879FFA7C0C2FDCF5DC7FD9E" name="Indonesia" pageId="14" pageNumber="63">Indonesia</collectingCountry>
to the
<collectingCountry id="73B4BCC19879FFA7C71CFDF25CA3FD9E" box="[836,926,562,586]" name="Malaysia" pageId="14" pageNumber="63">Malacca</collectingCountry>
Straits, via the Gulf of
<collectingCountry id="73B4BCC19879FFA7C0C2FDF25DCCFDB9" name="Thailand" pageId="14" pageNumber="63">Thailand</collectingCountry>
(
<figureCitation id="9398E0D49879FFA7C75CFD955C74FDB9" box="[772,841,597,621]" captionStart="Fig" captionStartId="8.[163,196,811,833]" captionTargetBox="[235,1140,214,775]" captionTargetId="figure-280@8.[234,1141,213,777]" captionTargetPageId="8" captionText="Fig. 3. Map showing sampling localities and distribution ranges of Acetes indicus, A. omorii sp. nov. and A. spiniger recognised in this study with certainty: blue solid circles (nos. 12) for A. indicus, green solid squares and open square (museum vouchered specimens) for A. spiniger, and red solid triangles for A. omorii sp. nov. Numerals indicating sampling locations: 1, Chiringa; 2, Kumira-Sandwip Ship Ghat; 3, Merbok Mangrove; 4; Matang Mangrove, 5; Kuala Selangor; 6, Malacca; 7, Samut Prakan; 8, Setarap Estuary, South Kalimantan; and 9, Goa." figureDoi="http://doi.org/10.5281/zenodo.12572342" httpUri="https://zenodo.org/record/12572342/files/figure.png" pageId="14" pageNumber="63">Fig. 3</figureCitation>
). On the west coasts of Peninsular
<collectingCountry id="73B4BCC19879FFA7C69BFDB75C1AFD5B" box="[707,807,631,655]" name="Malaysia" pageId="14" pageNumber="63">Malaysia</collectingCountry>
, specimens of
<taxonomicName id="CCA387D29879FFA7C797FDB85B0AFD5B" authority="Alamri &amp; Alotaiby, 2024" authorityName="Alamri &amp; Alotaiby" authorityYear="2024" box="[975,1079,632,655]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="omorii" status="sp. nov.">
<emphasis id="39D720439879FFA7C797FDB85B0AFD5B" box="[975,1079,632,655]" italics="true" pageId="14" pageNumber="63">A. omorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22E49D389879FFA7C019FDB85BAAFD5B" box="[1089,1175,632,655]" pageId="14" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
were commonly captured in the inshore waters including mangrove estuaries (see also
<bibRefCitation id="6F3281A09879FFA7C01CFD7D5C18FD23" author="Hanamura, Y. &amp; R. Siow &amp; P. - E. Chee" pageId="14" pageNumber="63" pagination="16 - 25" refId="ref10642" refString="Hanamura, Y., R. Siow and P. - E. Chee 2007. Abundance and spatio-temporal distribution of Acetes shrimp (Crustacea, Decapoda, Sergestidae) in the Merbok and Matang mangrove estuaries, Peninsular Malaysia. Malaysian Fisheries Journal 6: 16 - 25." type="journal article" year="2007">
Hanamura
<emphasis id="39D720439879FFA7C099FD7D5DE2FD23" italics="true" pageId="14" pageNumber="63">et al.</emphasis>
, 2007
</bibRefCitation>
; under
<taxonomicName id="CCA387D29879FFA7C72AFD205CE0FD23" authorityName="H. Milne Edwards" authorityYear="1830" box="[882,989,735,759]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D720439879FFA7C72AFD205CE0FD23" box="[882,989,735,759]" italics="true" pageId="14" pageNumber="63">A. indicus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,308 @@
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<subSubSection id="43B9AFDA987FFFA1C6AEFB4B5B9CFB77" box="[758,1185,1163,1187]" pageId="8" pageNumber="57" type="nomenclature">
<paragraph id="0B1CFC51987FFFA1C6AEFB4B5B9CFB77" blockId="8.[758,1185,1163,1224]" box="[758,1185,1163,1187]" pageId="8" pageNumber="57">
<heading id="50544B3D987FFFA1C6AEFB4B5B9CFB77" box="[758,1185,1163,1187]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="57" reason="4">
<taxonomicName id="CCA387D2987FFFA1C6AEFB4B5B9CFB77" ID-CoL="9XFD2" authority="H. Milne Edwards, 1830" authorityName="H. Milne Edwards" authorityYear="1830" box="[758,1185,1163,1187]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D72043987FFFA1C6AEFB4B5CAEFB77" bold="true" box="[758,915,1163,1187]" italics="true" pageId="8" pageNumber="57">Acetes indicus</emphasis>
H.
<bibRefCitation id="6F3281A0987FFFA1C7E3FB4B5B9CFB77" author="Milne Edwards, M. H." box="[955,1185,1163,1187]" pageId="8" pageNumber="57" pagination="333 - 352" refId="ref11123" refString="Milne Edwards, M. H. 1830. Description des genres Glaucothoe, Sicyonie, Sergeste et Acete, de l`ordre des Crustaces decapodes. Annales des Sciences Naturelles, Paris 19: 333 - 352, pls 8 - 11." type="journal article" year="1830">Milne Edwards, 1830</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA987FFFA1C7F1FB725CD3FB1C" box="[937,1006,1202,1224]" pageId="8" pageNumber="57" type="description">
<paragraph id="0B1CFC51987FFFA1C7F1FB725CD3FB1C" blockId="8.[758,1185,1163,1224]" box="[937,1006,1202,1224]" pageId="8" pageNumber="57">
(
<figureCitation id="9398E0D4987FFFA1C7E8FB725CDBFB1C" box="[944,998,1202,1224]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="8" pageNumber="57">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA987FFFA1C69BFB235BBCFAE7" pageId="8" pageNumber="57" type="reference_group">
<paragraph id="0B1CFC51987FFFA1C69BFB235C3BFAC1" blockId="8.[707,1236,1250,1331]" pageId="8" pageNumber="57">
<treatmentCitationGroup id="2BB3DB7F987FFFA1C69BFB235C3BFAC1" pageId="8" pageNumber="57">
<treatmentCitation id="8A02DA40987FFFA1C69BFB235B5BFB2C" author="Milne Edwards, M. H." box="[707,1126,1250,1272]" page="351" pageId="8" pageNumber="57" year="1830">
<taxonomicName id="CCA387D2987FFFA1C69BFB235B5BFB2C" ID-CoL="9XFD2" authority="H. Milne Edwards, 1830: 351" authorityName="H. Milne Edwards" authorityPageNumber="351" authorityYear="1830" box="[707,1126,1250,1272]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D72043987FFFA1C69BFB235C7AFB23" box="[707,839,1250,1272]" italics="true" pageId="8" pageNumber="57">Acetes indicus</emphasis>
H.
<bibRefCitation id="6F3281A0987FFFA1C734FB225B5BFB2C" author="Milne Edwards, M. H." box="[876,1126,1250,1272]" pageId="8" pageNumber="57" pagination="333 - 352" refId="ref11123" refString="Milne Edwards, M. H. 1830. Description des genres Glaucothoe, Sicyonie, Sergeste et Acete, de l`ordre des Crustaces decapodes. Annales des Sciences Naturelles, Paris 19: 333 - 352, pls 8 - 11." type="journal article" year="1830">Milne Edwards, 1830: 351</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 11, figs 19.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0B1CFC51987FFFA1C69BFADD5BBCFAE7" blockId="8.[707,1236,1250,1331]" box="[707,1153,1309,1331]" pageId="8" pageNumber="57">
<treatmentCitationGroup id="2BB3DB7F987FFFA1C69BFADD5BBCFAE7" box="[707,1153,1309,1331]" pageId="8" pageNumber="57">
?
<taxonomicName id="CCA387D2987FFFA1C695FADD5C72FAE6" ID-CoL="9XFD2" authorityName="H. Milne Edwards" authorityYear="1830" box="[717,847,1309,1330]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D72043987FFFA1C695FADD5C72FAE6" box="[717,847,1309,1330]" italics="true" pageId="8" pageNumber="57">Acetes indicus</emphasis>
</taxonomicName>
:
<treatmentCitation id="8A02DA40987FFFA1C703FADE5CCDFAE6" author="Kemp, S." box="[859,1008,1309,1331]" page="47" pageId="8" pageNumber="57" year="1917">
<bibRefCitation id="6F3281A0987FFFA1C703FADE5CCDFAE6" author="Kemp, S." box="[859,1008,1309,1331]" pageId="8" pageNumber="57" pagination="43 - 58" refId="ref11017" refString="Kemp, S. 1917. Notes on Crustacea Decapoda in the Indian Museum. The genus Acetes Milne-Edwards. Records of the Indian Museum 53: 43 - 58." type="journal article" year="1917">Kemp, 1917: 47</bibRefCitation>
</treatmentCitation>
, fig. 2a (in part)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="43B9AFDA987FFFA2C685FAB25EF2FB6E" lastPageId="11" lastPageNumber="60" pageId="8" pageNumber="57" type="materials_examined">
<paragraph id="0B1CFC51987FFFA3C685FAB25DA3FF3B" blockId="8.[707,1236,1394,1799]" lastBlockId="10.[140,670,215,1800]" lastPageId="10" lastPageNumber="59" pageId="8" pageNumber="57">
<emphasis id="39D72043987FFFA1C685FAB25CFDFA5E" box="[733,960,1394,1418]" italics="true" pageId="8" pageNumber="57">Material examined.</emphasis>
<materialsCitation id="BBCBF60C987FFFA1C781FAB25B0DFA78" ID-GBIF-Occurrence="4899861314" collectorName="Market Bazar" country="Bangladesh" location="Chiringa" pageId="8" pageNumber="57" specimenCount="1">
<collectingCountry id="73B4BCC1987FFFA1C781FAB25B64FA5E" box="[985,1113,1394,1418]" name="Bangladesh" pageId="8" pageNumber="57">Bangladesh</collectingCountry>
.
<emphasis id="39D72043987FFFA1C02CFAB25DF1FA78" italics="true" pageId="8" pageNumber="57">
<location id="0E7CAA8A987FFFA1C02CFAB25BE9FA5E" LSID="urn:lsid:plazi:treatment:830A4D47987FFFA2C6AEFB4B5EF2FB6E:0E7CAA8A987FFFA1C02CFAB25BE9FA5E" box="[1140,1236,1394,1418]" country="Bangladesh" name="Chiringa" pageId="8" pageNumber="57">Chiringa</location>
(
</emphasis>
=
<emphasis id="39D72043987FFFA1C6B2FA555C77FA79" box="[746,842,1429,1453]" italics="true" pageId="8" pageNumber="57">Chakari)</emphasis>
:
<emphasis id="39D72043987FFFA1C701FA555B0DFA78" box="[857,1072,1427,1453]" italics="true" pageId="8" pageNumber="57">
Cox`s
<collectorName id="A6569987987FFFA1C7C3FA555B0DFA78" box="[923,1072,1429,1453]" pageId="8" pageNumber="57">Market Bazar</collectorName>
</emphasis>
</materialsCitation>
,
<specimenCount id="1DA537D8987FFFA1C066FA555BACFA79" box="[1086,1169,1429,1453]" count="8" pageId="8" pageNumber="57" type="male">8 males</specimenCount>
(cl ca 3.55.0 mm),
<specimenCount id="1DA537D8987FFFA1C701FA775CECFA1B" box="[857,977,1463,1487]" count="19" pageId="8" pageNumber="57" type="female">19 females</specimenCount>
(cl ca 4.58.0 mm),
<date id="7F1DDA91987FFFA1C0E1FA775C0DFA26" pageId="8" pageNumber="57" value="2018-08-19">19 Aug 2018</date>
, coll. Kanak (formalin to 70% ethanol) (NSMT-Cr 31611);
<specimenCount id="1DA537D8987FFFA1C7C6FA3C5CC9F9C1" box="[926,1012,1532,1557]" count="5" pageId="8" pageNumber="57" type="male">5 males</specimenCount>
(cl ca 4.0
<quantity id="CC5B51B4987FFFA1C02FFA3C5BF5F9C1" box="[1143,1224,1532,1557]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="57" unit="mm" value="4.5">4.5 mm</quantity>
),
<specimenCount id="1DA537D8987FFFA1C69BF9DF5C7DF9E3" box="[707,832,1567,1591]" count="29" pageId="8" pageNumber="57" type="female">29 females</specimenCount>
(cl ca 4.58.0 mm), data same as precedent (ethanol) (NSMT-Cr 31612);
<specimenCount id="1DA537D8987FFFA1C024F9825DC6F9A8" count="2" pageId="8" pageNumber="57" type="generic">2 specimens</specimenCount>
used for DNA analysis, data same as previous (NSMT-Cr 31613, 31614):
<emphasis id="39D72043987FFFA1C075F9475C2DF916" italics="true" pageId="8" pageNumber="57">Sitakunda Fish Market</emphasis>
,
<emphasis id="39D72043987FFFA1C77AF96A5B69F916" box="[802,1108,1706,1730]" italics="true" pageId="8" pageNumber="57">Kumira-Sandwip Ship Ghat</emphasis>
:
<specimenCount id="1DA537D8987FFFA1C030F96A5BE9F916" box="[1128,1236,1706,1730]" count="9" pageId="8" pageNumber="57" type="female">9 females</specimenCount>
(cl 6.0
<quantity id="CC5B51B4987FFFA1C740F90C5C54F931" box="[792,873,1740,1765]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="57" unit="mm" value="6.5">6.5 mm</quantity>
),
<geoCoordinate id="6E979A96987FFFA1C727F90D5B35F930" box="[895,1032,1739,1765]" degrees="22" direction="north" minutes="30.157" orientation="latitude" pageId="8" pageNumber="57" precision="1" value="22.502617">22°30.157N</geoCoordinate>
,
<geoCoordinate id="6E979A96987FFFA1C04DF90D5BA6F930" box="[1045,1179,1739,1765]" degrees="91" direction="east" minutes="41.324" orientation="longitude" pageId="8" pageNumber="57" precision="1" value="91.688736">91°41.324E</geoCoordinate>
,
<date id="7F1DDA91987FFFA1C0F1F90D5DC6F8D3" pageId="8" pageNumber="57" value="2018-12">Dec 2018</date>
, coll. Md Rakel-Ul Islam (NSMT-Cr 31615);
<specimenCount id="1DA537D8987EFFA0C4BEF96A5E7BF916" box="[230,326,1706,1730]" count="35" pageId="9" pageNumber="58" type="male">35 males</specimenCount>
(cl
<quantity id="CC5B51B4987EFFA0C529F96A5ECCF916" box="[369,497,1706,1730]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.6" metricValueMin="4.3" pageId="9" pageNumber="58" unit="mm" value="4.949999999999999" valueMax="5.6" valueMin="4.3">4.35.6 mm</quantity>
),
<specimenCount id="1DA537D8987EFFA0C65DF96A5D47F916" box="[517,634,1706,1730]" count="37" pageId="9" pageNumber="58" type="female">37 females</specimenCount>
(cl 4.07.0 mm),
<date id="7F1DDA91987EFFA0C578F90D5EB4F931" box="[288,393,1741,1765]" pageId="9" pageNumber="58" value="2019-02">Feb 2019</date>
, other data same as precedent (NSMT-Cr 31616);
<specimenCount id="1DA537D8987EFFA0C5F7F92F5D0BF8D3" box="[431,566,1775,1799]" count="3" pageId="9" pageNumber="58" type="generic">3 specimens</specimenCount>
used for DNA analysis, data same as precedent (NSMT-Cr 3161731619):
<specimenCount id="1DA537D8987EFFA0C7CFF90C5CC6F931" box="[919,1019,1740,1765]" count="25" pageId="9" pageNumber="58" type="male">25 males</specimenCount>
(cl 4.66.0 mm),
<specimenCount id="1DA537D8987EFFA0C09FF90D5C2BF8D3" count="9" pageId="9" pageNumber="58" type="female">9 females</specimenCount>
(cl 6.0
<quantity id="CC5B51B4987EFFA0C72CF92F5CFBF8D3" box="[884,966,1775,1799]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="58" unit="mm" value="7.6">7.6 mm</quantity>
), commercial catch,
<date id="7F1DDA91987EFFA3C0F4F92F5FFEFF3B" lastPageId="10" lastPageNumber="59" pageId="9" pageNumber="58" value="2019-02">Feb 2019</date>
, coll. Md Rakel-UI Islam (NSMT-Cr 31620).
</paragraph>
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<paragraph id="0B1CFC51987EFFA0C4FBFA765B50F9A8" blockId="9.[163,1214,1462,1660]" pageId="9" pageNumber="58">
Fig. 4.
<taxonomicName id="CCA387D2987EFFA0C4A1FA765D5BFA1F" authority="H. Milne Edwards, 1830" authorityName="H. Milne Edwards" authorityYear="1830" box="[249,614,1462,1484]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="58" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D72043987EFFA0C4A1FA765E41FA1F" box="[249,380,1462,1483]" italics="true" pageId="9" pageNumber="58">Acetes indicus</emphasis>
H.
<bibRefCitation id="6F3281A0987EFFA0C5C7FA765D5BFA1F" author="Milne Edwards, M. H." box="[415,614,1462,1484]" pageId="9" pageNumber="58" pagination="333 - 352" refId="ref11123" refString="Milne Edwards, M. H. 1830. Description des genres Glaucothoe, Sicyonie, Sergeste et Acete, de l`ordre des Crustaces decapodes. Annales des Sciences Naturelles, Paris 19: 333 - 352, pls 8 - 11." type="journal article" year="1830">Milne Edwards, 1830</bibRefCitation>
</taxonomicName>
. (ag) male (cl 5.6 mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3 mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagellum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites. Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively.
</paragraph>
</caption>
<paragraph id="0B1CFC51987DFFA3C4FFFF3B5EF7FDD1" blockId="10.[140,670,215,1800]" pageId="10" pageNumber="59">
<emphasis id="39D72043987DFFA3C4FFFF3B5E18FEC6" box="[167,293,251,274]" italics="true" pageId="10" pageNumber="59">Description</emphasis>
.
<emphasis id="39D72043987DFFA3C51EFF3A5EB9FEC6" box="[326,388,250,274]" italics="true" pageId="10" pageNumber="59">Male.</emphasis>
Rostrum (
<figureCitation id="9398E0D4987DFFA3C5ADFF3A5D61FEC6" box="[501,604,250,274]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4a, h</figureCitation>
) typical of Asian members of genus in having sub-triangular plate raised slightly from dorsal margin of carapace, having sharply pointed terminal tooth and small anterior tooth in addition to somewhat larger posterior tooth on its dorsal margin. Carapace smooth on surface, except for post-orbital and hepatic teeth.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C4FFFDCF5FC7FCE9" blockId="10.[140,670,215,1800]" pageId="10" pageNumber="59">
Abdomen with first somite possessing procurved sharp tooth present at median part of ventral sternite between right and left pleopods. Sixth somite (
<figureCitation id="9398E0D4987DFFA3C57DFDB75E48FD5B" box="[293,373,631,655]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4b</figureCitation>
) twice length of fifth, postero-dorsal part ending in small postero-median tooth, no red spot on postero-ventral part. Telson (
<figureCitation id="9398E0D4987DFFA3C4CCFD1F5FC0FD23" box="[148,253,735,759]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4b, c</figureCitation>
) 0.7 times as long as sixth abdominal somite, with narrowly rounded or sub-triangular distal end.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C4FFFC885EB3FC56" blockId="10.[140,670,215,1800]" pageId="10" pageNumber="59">
Eye (
<figureCitation id="9398E0D4987DFFA3C4B8FC875E10FC8B" box="[224,301,839,863]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4a</figureCitation>
) with well pigmented cornea, latter wider than eye-stalk.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C4FFFC4D5D38F94B" blockId="10.[140,670,215,1800]" pageId="10" pageNumber="59">
Antennule noticeably elongate in mature males, particularly in third segment of peduncle, 0.810.97 times as long as combined length of first and second segments; lower flagellum (
<figureCitation id="9398E0D4987DFFA3C62AFC355FF8FBFB" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4a, d</figureCitation>
) modified into clasping organ, commonly composed of 12 segments, infrequently 13 or 14 segments; first segment of trunk rather long, noticeably narrowing at mid-length, three times or more as long as its narrowest part, forming distinct oblique conjunction with second one; major clasping spine moderately stout, reaching mid-length of fourth segment of main branch, bearing 2 incompletely aligned rows of minute serrated denticles, and also microscopically small clasping spine near base of major one; first segment of main branch with single basal setal-spine, and with or without single distal setal-spine, second and third segments each with single distal and/or sub-distal setal-spine, fourth segment 1.01.5 times as long as combined length of second and third segments, bearing usually 9 or 10 setal-spines on its entire margin, fifth segment with 13 setal spines.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C4FFF96A5C9FFF3B" blockId="10.[140,670,215,1800]" lastBlockId="10.[707,1237,215,1799]" pageId="10" pageNumber="59">
Antennal scale (
<figureCitation id="9398E0D4987DFFA3C538F96A5E8FF916" box="[352,434,1706,1730]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4a</figureCitation>
) 0.750.80 times as long as carapace, slightly but distinctly extending beyond anterior margin of second segment of antennular peduncle.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685FF3A5B7EFCCD" blockId="10.[707,1237,215,1799]" pageId="10" pageNumber="59">
External maxilliped and pereopods varying in length depending on body size. Third maxilliped in larger males extending as far as anterior end of antennular peduncle, distal segment three-fourths times as long as penultimate. First pereopod in larger males extending as far as cornea. Second pereopod in larger males reaching as far as anterior end of antennal scale. Third pereopod (
<figureCitation id="9398E0D4987DFFA3C0F1FE2D5DE0FDF3" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4e</figureCitation>
) in larger males extending anterior end of antennular peduncle, distinct sharp tooth present at disto-mesial part of basis and anteriorly curving projection on inner margin of coxa. Genital coxa (
<figureCitation id="9398E0D4987DFFA3C752FD5A5C61FD66" box="[778,860,666,690]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4e</figureCitation>
) often with slender or sub-acute distal part in lateral aspect, while sub-rectangular or obtusely rounded in ventral aspect, with truncate or widely sinuous anterior part.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685FCE55DC5FBD9" blockId="10.[707,1237,215,1799]" pageId="10" pageNumber="59">
Petasma (
<figureCitation id="9398E0D4987DFFA3C713FCE55C81FCE9" box="[843,956,805,829]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4f, g</figureCitation>
) with blunt and distally rounded capitulum bearing numerous hooks along lateral surfaces as well as distal part; processus ventralis slender, extending as far as distal margin of capitulum; no trace of pars astringen. Appendix masculina with 2 or 3 short hooks distally.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685FBD75CFFFB6E" blockId="10.[707,1237,215,1799]" pageId="10" pageNumber="59">
Endopod (
<figureCitation id="9398E0D4987DFFA3C717FBD75CA3FBFB" box="[847,926,1047,1071]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4b</figureCitation>
) of uropod with 16 irregularly-shaped red-spots in fresh materials: exopod with small tooth on lateral margin, demarcating non-ciliated and ciliated parts, latter about 0.6 times as long as former.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685FB055B5AFB09" blockId="10.[707,1237,215,1799]" box="[733,1127,1221,1245]" pageId="10" pageNumber="59">
<emphasis id="39D72043987DFFA3C685FB055C10FB09" box="[733,813,1221,1245]" italics="true" pageId="10" pageNumber="59">Female</emphasis>
. Usually larger than males.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685FB275C89FA5E" blockId="10.[707,1237,215,1799]" pageId="10" pageNumber="59">
Antennule (
<figureCitation id="9398E0D4987DFFA3C704FB275C96FB2B" box="[860,939,1255,1279]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4h</figureCitation>
) not elongated as in males, first segment longest, and third segments 0.50 0.55 times as long as combined length of first and second segments; no examined females with intact lower flagellum.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685FA555C16F9C1" blockId="10.[707,1237,215,1799]" pageId="10" pageNumber="59">
Antennal scale (
<figureCitation id="9398E0D4987DFFA3C7CEFA555CD7FA79" box="[918,1002,1429,1453]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4h</figureCitation>
) 0.710.74 times as long as carapace, extending to mid-length to distal one-third of third segment of antennular peduncle.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685F9DF5B9DF94B" blockId="10.[707,1237,215,1799]" pageId="10" pageNumber="59">
Third and fourth thoracic sternite having a somewhat shallow and wide longitudinal groove, not conspicuously narrowing anteriorly, median swelling low and not pronounced (
<figureCitation id="9398E0D4987DFFA3C06DF9475BAEF94B" box="[1077,1171,1671,1695]" captionStart="Fig" captionStartId="9.[163,196,1462,1484]" captionTargetBox="[272,1104,213,1429]" captionTargetId="figure-54@9.[272,1104,213,1429]" captionTargetPageId="9" captionText="Fig. 4. Acetes indicus H. Milne Edwards, 1830. (ag) male (cl 5.6mm) from Kumia-Sandwip Ship Ghat Bangladesh, and (hj) female (cl 7.3mm) from same locality (NSMT-Cr 31616): a, anterior part of cephalothorax, lateral; b, posterior part of abdomen, lateral; c, posterior part of telson, dorsal; d, left lower antennular flagel- lum; e, basal segments of third pereopod and genital coxa, ventral; f, right petasma, posterior; g, same, capitulum; h, anterior part of cephalothorax, lateral; i, basal segments of third pereopod and third and fourth abdominal sternites, ventral; j, lateral profile showing channels and sternites between third and fourth thoracic somites.Abbreviations: gc, genital coxa; iiist and ivst, third and fourth thoracic sternites, respectively." figureDoi="http://doi.org/10.5281/zenodo.12572348" httpUri="https://zenodo.org/record/12572348/files/figure.png" pageId="10" pageNumber="59">Fig. 4i, j</figureCitation>
).
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685F96A5B08F930" blockId="10.[707,1237,215,1799]" pageId="10" pageNumber="59">
<emphasis id="39D72043987DFFA3C685F96A5C78F916" box="[733,837,1706,1730]" italics="true" pageId="10" pageNumber="59">Body size</emphasis>
. Largest male recorded from
<collectingCountry id="73B4BCC1987DFFA3C0F8F96A5C29F931" name="Bangladesh" pageId="10" pageNumber="59">Bangladesh</collectingCountry>
cl
<quantity id="CC5B51B4987DFFA3C76DF90C5CB8F930" box="[821,901,1740,1764]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="10" pageNumber="59" unit="mm" value="5.6">5.6 mm</quantity>
, female 8.0 mm.
</paragraph>
<paragraph id="0B1CFC51987DFFA3C685F9305B22F8D3" blockId="10.[707,1237,215,1799]" box="[733,1055,1775,1799]" pageId="10" pageNumber="59">
<emphasis id="39D72043987DFFA3C685F9305C5BF8D3" box="[733,870,1775,1799]" italics="true" pageId="10" pageNumber="59">
<typeStatus id="D41842F3987DFFA3C685F9305C32F8D3" box="[733,783,1776,1799]" pageId="10" pageNumber="59">Type</typeStatus>
locality
</emphasis>
. Ganges Delta.
</paragraph>
<paragraph id="0B1CFC51987CFFA2C4FFFF175EE3FDD1" blockId="11.[140,669,215,1210]" pageId="11" pageNumber="60">
<emphasis id="39D72043987CFFA2C4FFFF175E39FF3B" box="[167,260,215,239]" italics="true" pageId="11" pageNumber="60">Remarks</emphasis>
. As mentioned above, our study evidently showed that the current concept of
<taxonomicName id="CCA387D2987CFFA2C4D4FEDD5E15FEE1" authorityName="H. Milne Edwards" authorityYear="1830" box="[140,296,285,309]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D72043987CFFA2C4D4FEDD5E15FEE1" box="[140,296,285,309]" italics="true" pageId="11" pageNumber="60">Acetes indicus</emphasis>
</taxonomicName>
is a species complex, which may have derived a consequence of cryptic speciation from a common ancestor, leading to the existence of three extant species (see
<figureCitation id="9398E0D4987CFFA2C59CFE455D34FE49" box="[452,521,389,413]" captionStart="Fig" captionStartId="5.[163,196,1203,1225]" captionTargetBox="[234,1140,213,1168]" captionTargetId="figure-137@5.[234,1141,213,1169]" captionTargetPageId="5" captionText="Fig. 1. Bayesian inference trees of Acetes indicus, A. omorii sp. nov., and A. spiniger with the outgroups of other congeneric species based on the mitochondrial 16S (a) and mitochondrial COI (b) markers. Values at the nodes are Bayesian posterior probabilities/Maximum Likelihood/Neighbor Joining support values. For the haplotype abbreviations, see Table 1." figureDoi="http://doi.org/10.5281/zenodo.12572332" httpUri="https://zenodo.org/record/12572332/files/figure.png" pageId="11" pageNumber="60">Figs 1</figureCitation>
,
<figureCitation id="9398E0D4987CFFA2C643FE455D16FE49" box="[539,555,389,413]" captionStart="Fig" captionStartId="6.[163,196,1040,1062]" captionTargetBox="[236,1139,214,1005]" captionTargetId="figure-281@6.[234,1141,213,1007]" captionTargetPageId="6" captionText="Fig. 2. Haplotype network (implemented in PopART) of the Acetes indicus, A. omorii sp. nov., and A. spiniger based on the mitochondrial 16S (a) and mitochondrial COI (b) markers using minimum spanning network (MSN) analysis with outgroups of congeneric species. Circles with different colours represent different species." figureDoi="http://doi.org/10.5281/zenodo.12572338" httpUri="https://zenodo.org/record/12572338/files/figure.png" pageId="11" pageNumber="60">2</figureCitation>
;
<tableCitation id="4621C9EA987CFFA2C661FE455DA5FE49" box="[569,664,389,413]" captionStart="Table 2" captionStartId="4.[163,213,208,230]" captionTargetBox="[145,1225,278,518]" captionText="Table 2. Base composition (%) of 16S rRNA gene amplified for species of Acetes indicus species group and two congeneric species." httpUri="http://table.plazi.org/id/5FDCACD99873FFADC4FBFF105EBCFF2B" pageId="11" pageNumber="60" tableUuid="5FDCACD99873FFADC4FBFF105EBCFF2B">Tables 2</tableCitation>
,
<tableCitation id="4621C9EA987CFFA2C4D4FE675FA6FE6B" box="[140,155,423,447]" captionStart="Table 3" captionStartId="4.[163,213,575,597]" captionTargetBox="[145,1228,645,885]" captionText="Table 3. Base composition (%) of COI gene amplified for species of Acetes indicus species group and two conge- neric species." httpUri="http://table.plazi.org/id/5FDCACD99873FFADC4FBFDFF5E76FDBA" pageId="11" pageNumber="60" tableUuid="5FDCACD99873FFADC4FBFDFF5E76FDBA">3</tableCitation>
). Among them, the population collected from the coastal waters of
<collectingCountry id="73B4BCC1987CFFA2C5DAFE0A5EC3FE36" box="[386,510,458,482]" name="Bangladesh" pageId="11" pageNumber="60">Bangladesh</collectingCountry>
was regarded the representative of
<emphasis id="39D72043987CFFA2C534FE2D5EE3FDD1" box="[364,478,493,517]" italics="true" pageId="11" pageNumber="60">
<taxonomicName id="CCA387D2987CFFA2C534FE2D5EE7FDD1" authorityName="H. Milne Edwards" authorityYear="1830" box="[364,474,493,517]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="indicus">A. indicus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="0B1CFC51987CFFA2C4FFFDCF5D41FCCE" blockId="11.[140,669,215,1210]" pageId="11" pageNumber="60">
Besides,
<bibRefCitation id="6F3281A0987CFFA2C552FDCF5D1EFDF3" author="Pathan, D. I. &amp; D. R. Jalihal" box="[266,547,527,551]" pageId="11" pageNumber="60" pagination="495 - 514" refId="ref11273" refString="Pathan, D. I. and D. R. Jalihal 1997. Proposed taxonomic revision of some important penaeid prawn genera (Crustacea: Decapoda) of Konkan coast (west coast of India). Journal of the Bombay Natural History Society 94: 495 - 514." type="journal article" year="1997">Pathan and Jalihal (1997)</bibRefCitation>
mentioned that the materials reported by
<bibRefCitation id="6F3281A0987CFFA2C58CFDF25D58FD9E" author="Kemp, S." box="[468,613,562,586]" pageId="11" pageNumber="60" pagination="43 - 58" refId="ref11017" refString="Kemp, S. 1917. Notes on Crustacea Decapoda in the Indian Museum. The genus Acetes Milne-Edwards. Records of the Indian Museum 53: 43 - 58." type="journal article" year="1917">Kemp (1917)</bibRefCitation>
contains both the large and small forms. We suspect that his materials may have included some specimens of the typical
<taxonomicName id="CCA387D2987CFFA2C530FD5A5EF1FD66" authorityName="H.Milne Edwards" authorityYear="1830" box="[360,460,666,690]" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="indicu">
<emphasis id="39D72043987CFFA2C530FD5A5EF1FD66" box="[360,460,666,690]" italics="true" pageId="11" pageNumber="60">A. indicu</emphasis>
</taxonomicName>
s because the illustration of the male lower antennular flagellum (
<bibRefCitation id="6F3281A0987CFFA2C4CBFD1F5E25FD23" author="Kemp, S." box="[147,280,735,759]" pageId="11" pageNumber="60" pagination="43 - 58" refId="ref11017" refString="Kemp, S. 1917. Notes on Crustacea Decapoda in the Indian Museum. The genus Acetes Milne-Edwards. Records of the Indian Museum 53: 43 - 58." type="journal article" year="1917">Kemp 1917</bibRefCitation>
, fig. 2a) seems similar to this species rather than its South-East Asian congener.
</paragraph>
<paragraph id="0B1CFC51987CFFA2C4FFFCE55D25FB86" blockId="11.[140,669,215,1210]" pageId="11" pageNumber="60">
Unfortunately, the sampling sites in this study were restricted to the eastern extremity of the Ganges Delta. Thus, a broad area of the northern coasts of the Gulf of Bengal was left un-surveyed, and future studies are strongly in need to clarify the nature of this species group in that region. The designation of a
<typeStatus id="D41842F3987CFFA2C589FC355D15FBD9" box="[465,552,1013,1037]" pageId="11" pageNumber="60" type="neotype">neotype</typeStatus>
, therefore, was not recommended until more detailed information are available from the region.
</paragraph>
<paragraph id="0B1CFC51987CFFA2C4FFFB9D5EF2FB6E" blockId="11.[140,669,215,1210]" pageId="11" pageNumber="60">
<emphasis id="39D72043987CFFA2C4FFFB9D5E14FBA1" box="[167,297,1117,1141]" italics="true" pageId="11" pageNumber="60">Distribution</emphasis>
. Specimens attributable to
<taxonomicName id="CCA387D2987CFFA2C634FB9D5E38FB43" authorityName="H. Milne Edwards" authorityYear="1830" class="Malacostraca" family="Acetidae" genus="Acetes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="60" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="39D72043987CFFA2C634FB9D5E38FB43" italics="true" pageId="11" pageNumber="60">Acetes indicus</emphasis>
</taxonomicName>
were found only from the coastal waters of
<collectingCountry id="73B4BCC1987CFFA2C4ADFB625E4CFB6E" box="[245,369,1186,1210]" name="Bangladesh" pageId="11" pageNumber="60">Bangladesh</collectingCountry>
(
<figureCitation id="9398E0D4987CFFA2C5D8FB625EFCFB6E" box="[384,449,1186,1210]" captionStart="Fig" captionStartId="8.[163,196,811,833]" captionTargetBox="[235,1140,214,775]" captionTargetId="figure-280@8.[234,1141,213,777]" captionTargetPageId="8" captionText="Fig. 3. Map showing sampling localities and distribution ranges of Acetes indicus, A. omorii sp. nov. and A. spiniger recognised in this study with certainty: blue solid circles (nos. 12) for A. indicus, green solid squares and open square (museum vouchered specimens) for A. spiniger, and red solid triangles for A. omorii sp. nov. Numerals indicating sampling locations: 1, Chiringa; 2, Kumira-Sandwip Ship Ghat; 3, Merbok Mangrove; 4; Matang Mangrove, 5; Kuala Selangor; 6, Malacca; 7, Samut Prakan; 8, Setarap Estuary, South Kalimantan; and 9, Goa." figureDoi="http://doi.org/10.5281/zenodo.12572342" httpUri="https://zenodo.org/record/12572342/files/figure.png" pageId="11" pageNumber="60">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>