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<mods:title id="430BD6B7A1E600AFD093FF846A94CA17">New Middle Triassic Bell-Shaped Nassellarian Radiolaria From Alpine And Carpathian Areas</mods:title>
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<emphasis id="C1DB8B1AC624203EFC7BFD9FFB77FDCF" bold="true" box="[982,1205,522,546]" italics="true" pageId="7" pageNumber="58">Gorispela fenestrata</emphasis>
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<figureCitation id="6B944B8DC624203EFBECFDBEFB4FFDAE" box="[1089,1165,555,579]" captionStart="Fig" captionStartId="8.[197,232,1750,1771]" captionTargetBox="[212,1390,203,1661]" captionTargetId="figure-171@8.[197,1410,178,1721]" captionTargetPageId="8" captionText="Fig. 3. a, a1. Gorispela cornubovis n. sp., holotype in lateral and basal views, respectively, Rc4; b. Gorispela fenestrata n. sp., holotype in lateral view, BV 85-70; c-e. Gorispela hungarica n. sp., FÖ 87, e. holotype. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582784" httpUri="https://zenodo.org/record/12582784/files/figure.png" pageId="7" pageNumber="58">Fig. 3b</figureCitation>
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<emphasis id="C1DB8B1AC624203EFC9DFDF8FC67FD68" bold="true" box="[816,933,621,645]" italics="true" pageId="7" pageNumber="58">Diagonisis</emphasis>
. Shell dicyrtid, campanulate. Cephalis practically imperforate except for a few small pores at the contact of the secondary lateral rays with the shell wall. Apical and ventral rays of the initial spicule forming two very robust horns diverging at about 90°, its bisectrice tilted at an angle of about 15° relative to the shell axis. Horns robust, conical on the proximal half or more and three-bladed and pyramidal on the distal portion. Apical horn tilted in dorsal direction, and apical ray prolonged along the inner side of the apical horn up to the base of the three-bladed portion. Ventral horn with proximal half conical and thick, and distal half three-bladed with pointed end and thin blades. Ventral ray of the initial spicule straight and very long inside shell, prolonged in the centre of the hollow conical half up to the base of the three-bladed portion. Collar boundary at the level of a circumferential row of small circular pores above the first circumferential ridge or, above it, at the level of MB that is marked at the surface by a constriction at the dorsal part of cephalis or a similar constriction at the base of the ventral horn. Thorax truncate-conical, with three circumferential ridges, the first one sometimes less marked. Each interval between two circumferential ridges with a row of very large oval pores forming a fenestrated structure. Distal end of shell with a thin circumferential band-like rings and, below it, a narrow girdle, both originated in the same place.
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<emphasis id="C1DB8B1AC624203EFC9DFA7EFC24F9EE" bold="true" box="[816,998,1515,1539]" italics="true" pageId="7" pageNumber="58">Studied material</emphasis>
.
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in the sample BV 85-70.
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<materialsCitation id="43C75D55C624203EFC9DF999FA82F9C9" box="[816,1344,1548,1572]" collectionCode="MGL" pageId="7" pageNumber="58" specimenCount="1" typeStatus="holotype">
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<typeStatus id="2C14E9AAC624203EFC9DF999FC51F9C9" box="[816,915,1548,1572]" pageId="7" pageNumber="58" type="holotype">Holotype</typeStatus>
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.
<figureCitation id="6B944B8DC624203EFC0CF999FC2DF9C9" box="[929,1007,1548,1572]" captionStart="Fig" captionStartId="8.[197,232,1750,1771]" captionTargetBox="[212,1390,203,1661]" captionTargetId="figure-171@8.[197,1410,178,1721]" captionTargetPageId="8" captionText="Fig. 3. a, a1. Gorispela cornubovis n. sp., holotype in lateral and basal views, respectively, Rc4; b. Gorispela fenestrata n. sp., holotype in lateral view, BV 85-70; c-e. Gorispela hungarica n. sp., FÖ 87, e. holotype. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582784" httpUri="https://zenodo.org/record/12582784/files/figure.png" pageId="7" pageNumber="58">Fig. 3b</figureCitation>
, BV 85-70, coll.
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.110273.
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<emphasis id="C1DB8B1AC624203EFC9DF9B8FB10F9A8" box="[816,1234,1581,1605]" italics="true" pageId="7" pageNumber="58">
<emphasis id="C1DB8B1AC624203EFC9DF9B8FC71F9A8" bold="true" box="[816,947,1581,1605]" italics="true" pageId="7" pageNumber="58">Dimensions</emphasis>
(measured on
<typeStatus id="2C14E9AAC624203EFBC0F9B8FB08F9A8" box="[1133,1226,1581,1605]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
)
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. Length of shell without apical horn, measured on
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, 90 µm, of cephalis 57 µm, of thorax 34 µm, diameter of cephalis at its base 64 µm, of distal ring of thorax 127 µm, length of apical horn 65 µm, its diameter at base 27 µm, length of ventral horn 60 µm, its diameter at base 34 µm.
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<subSubSection id="BBB50483C624203EFC9DF961FC5AF8C1" pageId="7" pageNumber="58" type="etymology">
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<emphasis id="C1DB8B1AC624203EFC9DF961FC67F8E1" bold="true" box="[816,933,1780,1804]" italics="true" pageId="7" pageNumber="58">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC624203EFBC3F966FB21F8E6" box="[1134,1251,1779,1803]" italics="true" pageId="7" pageNumber="58">fenestratus</emphasis>
, -
<emphasis id="C1DB8B1AC624203EFB52F961FACEF8E6" box="[1279,1292,1780,1803]" italics="true" pageId="7" pageNumber="58">a</emphasis>
, -
<emphasis id="C1DB8B1AC624203EFA8AF961FA8AF8E6" box="[1319,1352,1780,1803]" italics="true" pageId="7" pageNumber="58">um</emphasis>
with windows.
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<emphasis id="C1DB8B1AC624203EFC9DF8A3FC50F8A3" bold="true" box="[816,914,1846,1870]" italics="true" pageId="7" pageNumber="58">Remarks</emphasis>
. This species differs from the other species of the genus
<taxonomicName id="34AF2C8BC624203EFC08F8C2FBD2F882" authorityName="Dumitrică" authorityYear="2024" box="[933,1040,1879,1903]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="7" pageNumber="58" phylum="Radiozoa" rank="genus">
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by having a fenestrate thorax with three circumferential ridges and a very narrow distal cylindrical circle.
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<emphasis id="C1DB8B1AC624203EFC9DF82FFBCEF83F" bold="true" box="[816,1036,1978,2002]" italics="true" pageId="7" pageNumber="58">Stratigraphic range</emphasis>
. Lower Ladinian of sample BV 85- 70 from the Livinallongo Formation.
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<figureCitation id="6B944B8DC624203EFE3DFBA9FDC3FBB9" box="[400,513,1084,1108]" captionStart="Fig" captionStartId="8.[197,232,1750,1771]" captionTargetBox="[212,1390,203,1661]" captionTargetId="figure-171@8.[197,1410,178,1721]" captionTargetPageId="8" captionText="Fig. 3. a, a1. Gorispela cornubovis n. sp., holotype in lateral and basal views, respectively, Rc4; b. Gorispela fenestrata n. sp., holotype in lateral view, BV 85-70; c-e. Gorispela hungarica n. sp., FÖ 87, e. holotype. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582784" httpUri="https://zenodo.org/record/12582784/files/figure.png" pageId="7" pageNumber="58">Figs. 3a, a</figureCitation>
<figureCitation id="6B944B8DC624203EFDADFBA9FDC8FBB9" box="[512,522,1084,1108]" captionStart="Fig" captionStartId="1.[199,234,1402,1423]" captionTargetBox="[197,1410,160,1373]" captionTargetId="figure-425@1.[197,1410,160,1373]" captionTargetPageId="1" captionText="Fig. 1. Geological map of the locality Cristian area, Braşov Mountains, Romania, modified from the geological maps Zărneşti and Braşov sheets, scale 1:50000, Săndulescu et al. (1972)." figureDoi="http://doi.org/10.5281/zenodo.12582778" httpUri="https://zenodo.org/record/12582778/files/figure.png" pageId="7" pageNumber="58">1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C624203EFF3AFBEAFD04F8A0" pageId="7" pageNumber="58" type="diagnosis">
<paragraph id="F3105708C624203EFF3AFBEAFD6AF8C1" blockId="7.[151,772,1151,2069]" pageId="7" pageNumber="58">
<emphasis id="C1DB8B1AC624203EFF3AFBEAFEC7FB7A" bold="true" box="[151,261,1151,1175]" italics="true" pageId="7" pageNumber="58">Diagnosis</emphasis>
. Shell dicyrtid and very short with blunt apex and long, very divergent and apically curved apical and ventral horns resembling the horns of an ox. Cephalis smooth-surfaced, with blunt and imperforate apex, its distal part, at the contact with the thorax, with sparse and very small pores. Initial spicule without dorsal ray. Primary and secondary lateral rays not extend outside. Apical and ventral spines very divergent, practically symmetrical and extended outside into two long and curved horns. Both horns hollow and straight on their proximal halves, then curved in apical direction and three-bladed, with an apically directed blade and two laterally directed blades. At the base of the laterally directed blades a well visible spur or nudge is developed. Thorax very short and divided into two parts by a well-marked and thin protruding circumferential ring: a truncate cone-shaped proximal part with sparse and circular or oval pores of variable sizes and irregularly disposed, and a distal and very short cylindrical part with larger, circular or oval pores. Distal end irregularly terminated as if the skeleton was not yet finished.
</paragraph>
<paragraph id="F3105708C624203EFF3AF8A3FD04F8A0" blockId="7.[151,772,1151,2069]" box="[151,710,1845,1870]" pageId="7" pageNumber="58">
<emphasis id="C1DB8B1AC624203EFF3AF8A3FE8FF8A3" bold="true" box="[151,333,1846,1870]" italics="true" pageId="7" pageNumber="58">Studied material</emphasis>
. A single specimen in sample Rc4.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C624203EFF3AF8C2FEF3F87D" pageId="7" pageNumber="58" type="materials_examined">
<paragraph id="F3105708C624203EFF3AF8C2FEF3F87D" blockId="7.[151,772,1151,2069]" pageId="7" pageNumber="58">
<materialsCitation id="43C75D55C624203EFF3AF8C2FEF3F87D" collectionCode="MGL" pageId="7" pageNumber="58" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC624203EFF3AF8C2FF38F882" bold="true" box="[151,250,1879,1903]" italics="true" pageId="7" pageNumber="58">
<typeStatus id="2C14E9AAC624203EFF3AF8C2FF38F882" box="[151,250,1879,1903]" pageId="7" pageNumber="58" type="holotype">Holotype</typeStatus>
</emphasis>
. The illustrated specimen, Rc4, coll.
<collectionCode id="95BECFCDC624203EFF3AF8EDFF14F87D" box="[151,214,1912,1936]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="7" pageNumber="58">MGL</collectionCode>
.110272.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C624203EFF3AF80CFC0AFF5B" pageId="7" pageNumber="58" type="description">
<paragraph id="F3105708C624203EFF3AF80CFC0AFF5B" blockId="7.[151,772,1151,2069]" lastBlockId="7.[816,1436,158,446]" pageId="7" pageNumber="58">
<emphasis id="C1DB8B1AC624203EFF3AF80CFED8F85C" bold="true" box="[151,282,1945,1969]" italics="true" pageId="7" pageNumber="58">Dimensions</emphasis>
. Length of shell without spines 73 µm, of cephalothorax 56 µm, of distal part of thorax 17 µm, of A and V horns 80 µm, diameter of cephalis 35 µm, of circular ring 62 µm, of distal cylindrical part of thorax 56 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C624203EFC9DFF2AFA89FF15" pageId="7" pageNumber="58" type="etymology">
<paragraph id="F3105708C624203EFC9DFF2AFA89FF15" blockId="7.[816,1436,158,446]" pageId="7" pageNumber="58">
<emphasis id="C1DB8B1AC624203EFC9DFF2AFC67FF3A" bold="true" box="[816,933,191,215]" italics="true" pageId="7" pageNumber="58">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC624203EFBCFFF55FB51FF3A" box="[1122,1171,192,215]" italics="true" pageId="7" pageNumber="58">corn</emphasis>
horn and
<emphasis id="C1DB8B1AC624203EFAB7FF2AFAFDFF3A" box="[1306,1343,191,215]" italics="true" pageId="7" pageNumber="58">bos</emphasis>
,
<emphasis id="C1DB8B1AC624203EFAE3FF2AFA44FF3A" box="[1358,1414,191,215]" italics="true" pageId="7" pageNumber="58">bovis</emphasis>
ox, due to its resemblance with the horns of an ox.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C624203EFC9DFE94FB14FE53" pageId="7" pageNumber="58" type="discussion">
<paragraph id="F3105708C624203EFC9DFE94FCBEFE91" blockId="7.[816,1436,158,446]" pageId="7" pageNumber="58">
<emphasis id="C1DB8B1AC624203EFC9DFE94FC50FEF4" bold="true" box="[816,914,257,281]" italics="true" pageId="7" pageNumber="58">Remarks</emphasis>
. This species differs from all the other species of the genus
<taxonomicName id="34AF2C8BC624203EFC16FEB7FBE4FED7" authorityName="Dumitrică" authorityYear="2024" box="[955,1062,290,314]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="7" pageNumber="58" phylum="Radiozoa" rank="genus">
<emphasis id="C1DB8B1AC624203EFC16FEB7FBE4FED7" box="[955,1062,290,314]" italics="true" pageId="7" pageNumber="58">Gorispela</emphasis>
</taxonomicName>
by having the A and V horns very divergent and curved in apical direction on their distal halves.
</paragraph>
<paragraph id="F3105708C624203EFC9DFE13FB14FE53" blockId="7.[816,1436,158,446]" pageId="7" pageNumber="58">
<emphasis id="C1DB8B1AC624203EFC9DFE13FBDFFE73" bold="true" box="[816,1053,390,414]" italics="true" pageId="7" pageNumber="58">Stratigraphic range</emphasis>
. Lower Ladinian (Fassanian), Recoaro, Vicentinian Alps, North
<collectingCountry id="8BB81798C624203EFB33FE33FB10FE53" box="[1182,1234,422,446]" name="Italy" pageId="7" pageNumber="58">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,109 @@
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<mods:title id="FA1967EE4D87FE4215FAAFACF0D0F03D">New Middle Triassic Bell-Shaped Nassellarian Radiolaria From Alpine And Carpathian Areas</mods:title>
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<mods:namePart id="7BC0B676CEAAF736726E6DB279EA805C">Dumitrică, Paulian</mods:namePart>
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<treatment id="7B06E61EC625203FFC09FEE2FB0AFAE1" LSID="urn:lsid:plazi:treatment:7B06E61EC625203FFC09FEE2FB0AFAE1" httpUri="http://treatment.plazi.org/id/7B06E61EC625203FFC09FEE2FB0AFAE1" lastPageNumber="57" pageId="6" pageNumber="57">
<subSubSection id="BBB50483C625203FFC09FEE2FAEBFE62" box="[932,1321,375,399]" pageId="6" pageNumber="57" type="nomenclature">
<paragraph id="F3105708C625203FFC09FEE2FAEBFE62" blockId="6.[932,1321,375,399]" box="[932,1321,375,399]" pageId="6" pageNumber="57">
<heading id="A858E064C625203FFC09FEE2FAEBFE62" box="[932,1321,375,399]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="57" reason="5">
<emphasis id="C1DB8B1AC625203FFC09FEE2FAEBFE62" bold="true" box="[932,1321,375,399]" pageId="6" pageNumber="57">
<taxonomicName id="34AF2C8BC625203FFC09FEE2FB24FE62" authorityName="Dumitrică" authorityYear="2024" box="[932,1254,375,399]" class="Polycystina" family="Pseudosaturniformidae" genus="Colpotrelus" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="6" pageNumber="57" rank="species" species="campanuliformis" status="sp. nov.">
<emphasis id="C1DB8B1AC625203FFC09FEE2FB24FE62" bold="true" box="[932,1254,375,399]" italics="true" pageId="6" pageNumber="57">Colpotrelus campanuliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C625203FFB40FEE2FAEBFE62" box="[1261,1321,375,399]" pageId="6" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C625203FFB9DFE0DFB5EFE5C" box="[1072,1180,408,433]" pageId="6" pageNumber="57" type="description">
<paragraph id="F3105708C625203FFB9DFE0DFB5EFE5C" blockId="6.[1072,1180,408,433]" box="[1072,1180,408,433]" pageId="6" pageNumber="57">
<figureCitation id="6B944B8DC625203FFB9DFE0DFB5EFE5C" box="[1072,1180,408,433]" captionStart="Fig" captionStartId="5.[197,232,1764,1785]" captionTargetBox="[209,1395,175,1686]" captionTargetId="figure-163@5.[197,1411,160,1696]" captionTargetPageId="5" captionText="Fig. 2. a-d. Silicotintinnabulum formosum n. sp.: a, a1, b. sample Rc4; a1. basal view of fig. a. showing the initial spicule without D ray; c. paratype, FÖ 87; d. holotype, Rc1. 2e. Silicotintinnabulum transitum n. sp., holotype, Rc4; f, g. Colpotrelus campanuliformis n. sp., Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582780" httpUri="https://zenodo.org/record/12582780/files/figure.png" pageId="6" pageNumber="57">Figs. 2f, g</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C625203FFC9DFE4FFB2CFC92" pageId="6" pageNumber="57" type="diagnosis">
<paragraph id="F3105708C625203FFC9DFE4FFAE9FCB3" blockId="6.[816,1437,474,1293]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFC9DFE4FFC5CFE1F" bold="true" box="[816,926,474,498]" italics="true" pageId="6" pageNumber="57">Diagnosis</emphasis>
. Shell very small, dicyrtid. Cephalis with a robust and long conical apical horn with pointed distal end. Cephalis globular with smooth surface and partly poreless or with small pores on its basal part. Initial spicule without dorsal ray. Ventral spine very small and conical. Collar boundary not marked by a constriction; it could be situated at the level where the shell starts becoming conical and where the pores increase in size and frequency. Thorax conical in the proximal part, with a well-marked circumferential ridge. Beyond it, shell is very short and cylindrical or inverted conical. Its pores are circular or oval, sparse and of various sizes.
</paragraph>
<paragraph id="F3105708C625203FFC9DFCF2FB2CFC92" blockId="6.[816,1437,474,1293]" box="[816,1262,871,895]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFC9DFCF2FC24FC92" bold="true" box="[816,998,871,895]" italics="true" pageId="6" pageNumber="57">Studied material</emphasis>
.
<specimenCount id="E5A99C81C625203FFC5EFCFDFB58FC92" box="[1011,1178,871,895]" count="2" pageId="6" pageNumber="57" type="generic">Two specimens</specimenCount>
in Rc4.
</paragraph>
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<subSubSection id="BBB50483C625203FFC9DFC1CFC3DFC2C" pageId="6" pageNumber="57" type="materials_examined">
<paragraph id="F3105708C625203FFC9DFC1CFC3DFC2C" blockId="6.[816,1437,474,1293]" pageId="6" pageNumber="57">
<materialsCitation id="43C75D55C625203FFC9DFC1CFC3DFC2C" collectionCode="MGL" pageId="6" pageNumber="57" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC625203FFC9DFC1CFC51FC4C" bold="true" box="[816,915,905,929]" italics="true" pageId="6" pageNumber="57">
<typeStatus id="2C14E9AAC625203FFC9DFC1CFC51FC4C" box="[816,915,905,929]" pageId="6" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
. The specimen from the sample Rc4, fig.
<quantity id="3457FAEDC625203FFAD6FC1DFA55FC4D" box="[1403,1431,904,928]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="6" pageNumber="57" unit="g" value="2.0">2g</quantity>
, coll.
<collectionCode id="95BECFCDC625203FFCCBFC3CFC64FC2C" box="[870,934,937,961]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="6" pageNumber="57">MGL</collectionCode>
.110271.
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</paragraph>
</subSubSection>
<subSubSection id="BBB50483C625203FFC9DFC5EFC63FBAB" pageId="6" pageNumber="57" type="description">
<paragraph id="F3105708C625203FFC9DFC5EFC63FBAB" blockId="6.[816,1437,474,1293]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFC9DFC5EFC71FC0E" bold="true" box="[816,947,971,995]" italics="true" pageId="6" pageNumber="57">Dimensions</emphasis>
. Total length with apical horn 100-115 µm, without horn 53-75 µm, diameter of cephalis 38-39 µm, of the circumferential ridge 69-77 µm, of distal segment 62-66 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C625203FFC9DFBDAFAA0FB8A" box="[816,1378,1103,1127]" pageId="6" pageNumber="57" type="etymology">
<paragraph id="F3105708C625203FFC9DFBDAFAA0FB8A" blockId="6.[816,1437,474,1293]" box="[816,1378,1103,1127]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFC9DFBDAFC67FB8A" bold="true" box="[816,933,1103,1127]" italics="true" pageId="6" pageNumber="57">Etymology</emphasis>
. From the campanulate shape of its shell.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C625203FFC9DFBE5FB0AFAE1" pageId="6" pageNumber="57" type="discussion">
<paragraph id="F3105708C625203FFC9DFBE5FACAFB06" blockId="6.[816,1437,474,1293]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFC9DFBE5FC50FB65" bold="true" box="[816,914,1136,1160]" italics="true" pageId="6" pageNumber="57">Remarks</emphasis>
. This species differs from the species of the other genera herein described by the very small size of its ventral spine, the absence of a three-bladed morphology of the apical horn, and its massive structure.
</paragraph>
<paragraph id="F3105708C625203FFC9DFB60FB0AFAE1" blockId="6.[816,1437,474,1293]" box="[816,1224,1268,1293]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFC9DFB60FBCAFAE0" bold="true" box="[816,1032,1269,1293]" italics="true" pageId="6" pageNumber="57">Stratigraphic range</emphasis>
. Lower Ladinian.
</paragraph>
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<heading id="A858E064C625203FFEA8FF25FD54FF25" box="[261,662,176,200]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="57" reason="5">
<emphasis id="C1DB8B1AC625203FFEA8FF25FD54FF25" bold="true" box="[261,662,176,200]" pageId="6" pageNumber="57">
<taxonomicName id="34AF2C8BC625203FFEA8FF25FD91FF25" authorityName="Dumitrică" authorityYear="2024" box="[261,595,176,200]" class="Polycystina" family="Pseudosaturniformidae" genus="Silicotintinnabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="6" pageNumber="57" rank="species" species="transitum" status="sp. nov.">
<emphasis id="C1DB8B1AC625203FFEA8FF25FD91FF25" bold="true" box="[261,595,176,200]" italics="true" pageId="6" pageNumber="57">Silicotintinnabulum transitum</emphasis>
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<taxonomicNameLabel id="DAE83661C625203FFDF7FF25FD54FF25" box="[602,662,176,200]" pageId="6" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="BBB50483C625203FFE05FF44FE30FF04" box="[424,498,209,233]" pageId="6" pageNumber="57" type="description">
<paragraph id="F3105708C625203FFE05FF44FE30FF04" blockId="6.[424,498,209,233]" box="[424,498,209,233]" pageId="6" pageNumber="57">
<figureCitation id="6B944B8DC625203FFE05FF44FE30FF04" box="[424,498,209,233]" captionStart="Fig" captionStartId="5.[197,232,1764,1785]" captionTargetBox="[209,1395,175,1686]" captionTargetId="figure-163@5.[197,1411,160,1696]" captionTargetPageId="5" captionText="Fig. 2. a-d. Silicotintinnabulum formosum n. sp.: a, a1, b. sample Rc4; a1. basal view of fig. a. showing the initial spicule without D ray; c. paratype, FÖ 87; d. holotype, Rc1. 2e. Silicotintinnabulum transitum n. sp., holotype, Rc4; f, g. Colpotrelus campanuliformis n. sp., Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582780" httpUri="https://zenodo.org/record/12582780/files/figure.png" pageId="6" pageNumber="57">Fig. 2e</figureCitation>
</paragraph>
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<paragraph id="F3105708C625203FFF3AFE81FEE2FD37" blockId="6.[151,772,275,1458]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFF3AFE81FEC7FEC1" bold="true" box="[151,261,276,300]" italics="true" pageId="6" pageNumber="57">Diagnosis</emphasis>
. Shell bell-shaped with a relatively large spherical and poreless cephalis with a relatively long three-bladed, axially directed and pointed apical horn. Collar boundary with a well-marked depression due to the spherical shape of the cephalis and its bigger size. Thorax short, wide conical bearing a distal row of around 12 wide elongate pores or windows and an irregular row of smaller, circular or oval pores above it, below the collar boundary. A few much smaller and irregularly arranged pores may also occur above these smaller pores. Distal end with a circular horizontal platy ring and, below it, a circular band with smaller diameter. Initial spiculat system not known, but should be similar to that of the
<typeStatus id="2C14E9AAC625203FFF3AFD57FF07FD37" box="[151,197,706,730]" pageId="6" pageNumber="57">type</typeStatus>
species.
</paragraph>
<paragraph id="F3105708C625203FFF3AFD76FD04FD16" blockId="6.[151,772,275,1458]" box="[151,710,739,763]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFF3AFD76FE8FFD16" bold="true" box="[151,333,739,763]" italics="true" pageId="6" pageNumber="57">Studied material</emphasis>
. A single specimen in sample Rc4.
</paragraph>
</subSubSection>
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<paragraph id="F3105708C625203FFF3AFC91FD7AFCF1" blockId="6.[151,772,275,1458]" box="[151,696,772,796]" pageId="6" pageNumber="57">
<materialsCitation id="43C75D55C625203FFF3AFC91FD7AFCF1" box="[151,696,772,796]" collectionCode="MGL" pageId="6" pageNumber="57" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC625203FFF3AFC91FF38FCF1" bold="true" box="[151,250,772,796]" italics="true" pageId="6" pageNumber="57">
<typeStatus id="2C14E9AAC625203FFF3AFC91FF38FCF1" box="[151,250,772,796]" pageId="6" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC625203FFEA5FC91FE96FCF1" box="[264,340,772,796]" captionStart="Fig" captionStartId="5.[197,232,1764,1785]" captionTargetBox="[209,1395,175,1686]" captionTargetId="figure-163@5.[197,1411,160,1696]" captionTargetPageId="5" captionText="Fig. 2. a-d. Silicotintinnabulum formosum n. sp.: a, a1, b. sample Rc4; a1. basal view of fig. a. showing the initial spicule without D ray; c. paratype, FÖ 87; d. holotype, Rc1. 2e. Silicotintinnabulum transitum n. sp., holotype, Rc4; f, g. Colpotrelus campanuliformis n. sp., Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582780" httpUri="https://zenodo.org/record/12582780/files/figure.png" pageId="6" pageNumber="57">Fig. 2e</figureCitation>
, sample Rc4, coll.
<collectionCode id="95BECFCDC625203FFDB2FC91FD9CFCF1" box="[543,606,772,796]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="6" pageNumber="57">MGL</collectionCode>
.110270.
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</paragraph>
</subSubSection>
<subSubSection id="BBB50483C625203FFF3AFCB0FF26FC6D" pageId="6" pageNumber="57" type="description">
<paragraph id="F3105708C625203FFF3AFCB0FF26FC6D" blockId="6.[151,772,275,1458]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFF3AFCB0FED8FCD0" bold="true" box="[151,282,805,829]" italics="true" pageId="6" pageNumber="57">Dimensions</emphasis>
. Total length of shell 107 µm, of thorax 37 µm, diameter of cephalis 44 µm, of distal end of thorax 98 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C625203FFF3AFC1CFEBAFBEE" pageId="6" pageNumber="57" type="etymology">
<paragraph id="F3105708C625203FFF3AFC1CFEBAFBEE" blockId="6.[151,772,275,1458]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFF3AFC1CFECEFC4C" bold="true" box="[151,268,905,929]" italics="true" pageId="6" pageNumber="57">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC625203FFDA9FC1CFDA3FC4D" box="[516,609,905,928]" italics="true" pageId="6" pageNumber="57">transitus</emphasis>
, -
<emphasis id="C1DB8B1AC625203FFD24FC1CFD54FC4D" box="[649,662,905,928]" italics="true" pageId="6" pageNumber="57">a</emphasis>
,
<emphasis id="C1DB8B1AC625203FFD18FC1CFD1CFC4D" box="[693,734,905,928]" italics="true" pageId="6" pageNumber="57">-um</emphasis>
transitional, this species being considered transitional between the genus
<taxonomicName id="34AF2C8BC625203FFED0FC5FFD8DFC0F" authorityName="Dumitrică" authorityYear="2024" box="[381,591,970,994]" class="Polycystina" family="Pseudosaturniformidae" genus="Silicotintinnabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="6" pageNumber="57" rank="genus">
<emphasis id="C1DB8B1AC625203FFED0FC5FFD8DFC0F" box="[381,591,970,994]" italics="true" pageId="6" pageNumber="57">Silicotintinnabulum</emphasis>
</taxonomicName>
and the genus
<taxonomicName id="34AF2C8BC625203FFF3AFC7EFEB3FBEE" authorityName="Kozur &amp; Mostler" authorityYear="1979" box="[151,369,1003,1027]" genus="Pseudosaturniforma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="57" rank="genus">
<emphasis id="C1DB8B1AC625203FFF3AFC7EFEB3FBEE" box="[151,369,1003,1027]" italics="true" pageId="6" pageNumber="57">Pseudosaturniforma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C625203FFF3AFB98FDE4FA5C" pageId="6" pageNumber="57" type="discussion">
<paragraph id="F3105708C625203FFF3AFB98FED4FA9D" blockId="6.[151,772,275,1458]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFF3AFB98FF3BFBC8" bold="true" box="[151,249,1037,1061]" italics="true" pageId="6" pageNumber="57">Remarks</emphasis>
. Although this species is only based on the
<typeStatus id="2C14E9AAC625203FFF3AFBBBFF35FBAB" box="[151,247,1070,1094]" pageId="6" pageNumber="57" type="holotype">holotype</typeStatus>
, it differs essentially from the type species of the genus
<taxonomicName id="34AF2C8BC625203FFF42FBDAFE03FB8A" authorityName="Dumitrică" authorityYear="2024" box="[239,449,1103,1127]" class="Polycystina" family="Pseudosaturniformidae" genus="Silicotintinnabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="6" pageNumber="57" rank="genus">
<emphasis id="C1DB8B1AC625203FFF42FBDAFE03FB8A" box="[239,449,1103,1127]" italics="true" pageId="6" pageNumber="57">Silicotintinnabulum</emphasis>
</taxonomicName>
by its morphology and especially by the larger size and spherical shape of the cephalis and the appearance of the row of wide and logitudinally elongated distal pores that suggest the beginning of the transition to the genus
<taxonomicName id="34AF2C8BC625203FFF3AFB61FEB3FAE1" authorityName="Kozur &amp; Mostler" authorityYear="1979" box="[151,369,1268,1292]" genus="Pseudosaturniforma" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="57" rank="genus">
<emphasis id="C1DB8B1AC625203FFF3AFB61FEB3FAE1" box="[151,369,1268,1292]" italics="true" pageId="6" pageNumber="57">Pseudosaturniforma</emphasis>
</taxonomicName>
. As remains of the genus
<taxonomicName id="34AF2C8BC625203FFF3AFA80FEABFAC0" authorityName="Dumitrică" authorityYear="2024" box="[151,361,1301,1325]" class="Polycystina" family="Pseudosaturniformidae" genus="Silicotintinnabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="6" pageNumber="57" rank="genus">
<emphasis id="C1DB8B1AC625203FFF3AFA80FEABFAC0" box="[151,361,1301,1325]" italics="true" pageId="6" pageNumber="57">Silicotintinnabulum</emphasis>
</taxonomicName>
we can mention the small pores from the base of the cephalis and the long and three-bladed apical horn.
</paragraph>
<paragraph id="F3105708C625203FFF3AFAECFDE4FA5C" blockId="6.[151,772,275,1458]" pageId="6" pageNumber="57">
<emphasis id="C1DB8B1AC625203FFF3AFAECFEB0FA7C" bold="true" box="[151,370,1401,1425]" italics="true" pageId="6" pageNumber="57">Stratigraphic range</emphasis>
. Very rare in the lower Ladinian of the Buchenstein Formation, Recoaro.
</paragraph>
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<taxonomicName id="34AF2C8BC627203DFC36FD15FB2DFD75" authorityName="Dumitrică" authorityYear="2024" box="[923,1263,640,664]" class="Polycystina" family="Pseudosaturniformidae" genus="Silicotintinnabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="4" pageNumber="55" rank="species" species="formosum" status="sp. nov.">
<emphasis id="C1DB8B1AC627203DFC36FD15FB2DFD75" bold="true" box="[923,1263,640,664]" italics="true" pageId="4" pageNumber="55">Silicotintinnabulum formosum</emphasis>
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<figureCitation id="6B944B8DC627203DFB83FD35FBB4FD55" box="[1070,1142,672,696]" captionStart="Fig" captionStartId="5.[197,232,1764,1785]" captionTargetBox="[209,1395,175,1686]" captionTargetId="figure-163@5.[197,1411,160,1696]" captionTargetPageId="5" captionText="Fig. 2. a-d. Silicotintinnabulum formosum n. sp.: a, a1, b. sample Rc4; a1. basal view of fig. a. showing the initial spicule without D ray; c. paratype, FÖ 87; d. holotype, Rc1. 2e. Silicotintinnabulum transitum n. sp., holotype, Rc4; f, g. Colpotrelus campanuliformis n. sp., Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582780" httpUri="https://zenodo.org/record/12582780/files/figure.png" pageId="4" pageNumber="55">Figs. 2</figureCitation>
a-d
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<paragraph id="F3105708C627203DFC9DFD76FC32FA3E" blockId="4.[816,1437,739,2053]" pageId="4" pageNumber="55">
<emphasis id="C1DB8B1AC627203DFC9DFD76FC5CFD16" bold="true" box="[816,926,739,763]" italics="true" pageId="4" pageNumber="55">Diagnosis</emphasis>
. Shell dicyrtid, campanulate. Cephalis globular, imperforate or bearing sparse and very small circular or oval pores, especially on its distal part. Apical horn relatively long, cylindrical and hollow on its proximal third and three-bladed and pointed on its distal part. Blades thin or thick separated by wide grooves, and bearing a few short teeth or, very rarely, numerous and very short spinules on their distal end. Initial spicule with apical ray vertically positioned and prolonged into the apical horn, and ventral ray not extended outside into a spine or horn. Thorax conical with about two or three irregularly disposed ranges of large pores that increase in size very fast distally so that on the distal part only about five large pores can be seen on half the perimeter. Pores circular, oval or rounded polygonal and commonly intricately arranged (fig. 1a-d). Distal end well-marked by a thin circumferential ring disposed in horizontal plane and another circumferential ring with the blade in vertical plane, both originated in the same region of the distal end of shell. Collar boundary marked by a slight constriction and considered as being situated at the appearance of larger pores, a little below the level of the median bar of the initial spicule.
</paragraph>
<paragraph id="F3105708C627203DFC9DFA49FB26F9DA" blockId="4.[816,1437,739,2053]" pageId="4" pageNumber="55">
<emphasis id="C1DB8B1AC627203DFC9DFA49FC2EFA19" bold="true" box="[816,1004,1500,1524]" italics="true" pageId="4" pageNumber="55">Studied material</emphasis>
.
<materialsCitation id="43C75D55C627203DFC52FA49FB22F9DA" collectionCode="SMF" country="Hungary" location="Buchenstein Formation" pageId="4" pageNumber="55" specimenCode="SMF4" specimenCount="9">
<specimenCount id="E5A99C81C627203DFC52FA49FB6CFA19" box="[1023,1198,1500,1524]" count="9" pageId="4" pageNumber="55" type="generic">Nine specimens</specimenCount>
in the samples Rc1, Rc4, and
<specimenCode id="A309FF73C627203DFC0EFA68FC24F9F8" box="[931,998,1533,1557]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="55">SMF4</specimenCode>
from the
<location id="F67001D3C627203DFBCCFA68FAA7F9F8" LSID="urn:lsid:plazi:treatment:7B06E61EC627203CFC36FD15FC6AF815:F67001D3C627203DFBCCFA68FAA7F9F8" box="[1121,1381,1533,1557]" country="Hungary" name="Buchenstein Formation" pageId="4" pageNumber="55">Buchenstein Formation</location>
, and two in the sample FÖ 87, from
<collectingCountry id="8BB81798C627203DFBD2F98AFB22F9DA" box="[1151,1248,1567,1591]" name="Hungary" pageId="4" pageNumber="55">Hungary</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C627203DFC9DF9AAFA46F995" pageId="4" pageNumber="55" type="materials_examined">
<paragraph id="F3105708C627203DFC9DF9AAFAACF9BA" blockId="4.[816,1437,739,2053]" box="[816,1390,1599,1623]" pageId="4" pageNumber="55">
<materialsCitation id="43C75D55C627203DFC9DF9AAFAACF9BA" box="[816,1390,1599,1623]" collectionCode="MGL" pageId="4" pageNumber="55" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC627203DFC9DF9AAFC51F9BA" bold="true" box="[816,915,1599,1623]" italics="true" pageId="4" pageNumber="55">
<typeStatus id="2C14E9AAC627203DFC9DF9AAFC51F9BA" box="[816,915,1599,1623]" pageId="4" pageNumber="55" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC627203DFC0CF9AAFBC8F9BA" box="[929,1034,1599,1623]" captionStart="Fig" captionStartId="5.[197,232,1764,1785]" captionTargetBox="[209,1395,175,1686]" captionTargetId="figure-163@5.[197,1411,160,1696]" captionTargetPageId="5" captionText="Fig. 2. a-d. Silicotintinnabulum formosum n. sp.: a, a1, b. sample Rc4; a1. basal view of fig. a. showing the initial spicule without D ray; c. paratype, FÖ 87; d. holotype, Rc1. 2e. Silicotintinnabulum transitum n. sp., holotype, Rc4; f, g. Colpotrelus campanuliformis n. sp., Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582780" httpUri="https://zenodo.org/record/12582780/files/figure.png" pageId="4" pageNumber="55">Figure 2d</figureCitation>
, sample Rc1, coll.
<collectionCode id="95BECFCDC627203DFB78F9AAFAD7F9BA" box="[1237,1301,1599,1623]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="4" pageNumber="55">MGL</collectionCode>
.110268.
</materialsCitation>
</paragraph>
<paragraph id="F3105708C627203DFC9DF9F4FA46F995" blockId="4.[816,1437,739,2053]" box="[816,1412,1632,1657]" pageId="4" pageNumber="55">
<materialsCitation id="43C75D55C627203DFC9DF9F4FAE8F995" box="[816,1322,1632,1657]" collectionCode="MGL" pageId="4" pageNumber="55" specimenCount="1" typeStatus="paratype">
<emphasis id="C1DB8B1AC627203DFC9DF9F4FC50F994" bold="true" box="[816,914,1633,1657]" italics="true" pageId="4" pageNumber="55">
<typeStatus id="2C14E9AAC627203DFC9DF9F4FC50F994" box="[816,914,1633,1657]" pageId="4" pageNumber="55" type="paratype">Paratype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC627203DFC0DF9F5FBC5F995" box="[928,1031,1632,1656]" captionStart="Fig" captionStartId="5.[197,232,1764,1785]" captionTargetBox="[209,1395,175,1686]" captionTargetId="figure-163@5.[197,1411,160,1696]" captionTargetPageId="5" captionText="Fig. 2. a-d. Silicotintinnabulum formosum n. sp.: a, a1, b. sample Rc4; a1. basal view of fig. a. showing the initial spicule without D ray; c. paratype, FÖ 87; d. holotype, Rc1. 2e. Silicotintinnabulum transitum n. sp., holotype, Rc4; f, g. Colpotrelus campanuliformis n. sp., Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582780" httpUri="https://zenodo.org/record/12582780/files/figure.png" pageId="4" pageNumber="55">Figure 2c</figureCitation>
, sample FÖ 87, coll.
<collectionCode id="95BECFCDC627203DFB47F9F5FAE8F995" box="[1258,1322,1632,1656]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="4" pageNumber="55">MGL</collectionCode>
</materialsCitation>
.110269.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C627203DFC9DF917FB2DF88D" pageId="4" pageNumber="55" type="description">
<paragraph id="F3105708C627203DFC9DF917FB54F910" blockId="4.[816,1437,739,2053]" pageId="4" pageNumber="55">
<emphasis id="C1DB8B1AC627203DFC9DF917FC71F977" bold="true" box="[816,947,1666,1690]" italics="true" pageId="4" pageNumber="55">Dimensions</emphasis>
. Total length of shell 100-104 µm, of cephalis with apical horn 65-70 µm, of apical horn 40 µm, of thorax 30-40 µm; diameter of cephalis 35-40 µm, of distal part of thorax 83-95 µm.
</paragraph>
<paragraph id="F3105708C627203DFC9DF893FB2DF88D" blockId="4.[816,1437,739,2053]" pageId="4" pageNumber="55">
<emphasis id="C1DB8B1AC627203DFC9DF893FC67F8F3" bold="true" box="[816,933,1798,1822]" italics="true" pageId="4" pageNumber="55">Variability</emphasis>
. As the drawings show, the variability of this species is especially visible in the shape and arrangement of pores and less in the size of specimens.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C627203DFC9DF8FCFA54F86C" box="[816,1430,1897,1921]" pageId="4" pageNumber="55" type="etymology">
<paragraph id="F3105708C627203DFC9DF8FCFA54F86C" blockId="4.[816,1437,739,2053]" box="[816,1430,1897,1921]" pageId="4" pageNumber="55">
<emphasis id="C1DB8B1AC627203DFC9DF8FCFC67F86C" bold="true" box="[816,933,1897,1921]" italics="true" pageId="4" pageNumber="55">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC627203DFBF7F8FCFB7EF86C" box="[1114,1212,1897,1921]" italics="true" pageId="4" pageNumber="55">formosus</emphasis>
, -
<emphasis id="C1DB8B1AC627203DFB7FF8FFFB1DF86C" box="[1234,1247,1898,1921]" italics="true" pageId="4" pageNumber="55">a</emphasis>
, -
<emphasis id="C1DB8B1AC627203DFB58F8FFFAD4F86C" box="[1269,1302,1898,1921]" italics="true" pageId="4" pageNumber="55">um</emphasis>
beautiful.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C627203CFC9DF81EFC6AF815" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55" type="discussion">
<paragraph id="F3105708C627203CFC9DF81EFCAFF899" blockId="4.[816,1437,739,2053]" lastBlockId="5.[816,1436,1884,2040]" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
<emphasis id="C1DB8B1AC627203DFC9DF81EFC50F84E" bold="true" box="[816,914,1931,1955]" italics="true" pageId="4" pageNumber="55">Remarks</emphasis>
. The apical part of shell, including cephalis and apical horn, has a rather constant morphology, except a single specimen (
<figureCitation id="6B944B8DC627203DFC40F859FBF8F809" box="[1005,1082,1996,2020]" captionStart="Fig" captionStartId="5.[197,232,1764,1785]" captionTargetBox="[209,1395,175,1686]" captionTargetId="figure-163@5.[197,1411,160,1696]" captionTargetPageId="5" captionText="Fig. 2. a-d. Silicotintinnabulum formosum n. sp.: a, a1, b. sample Rc4; a1. basal view of fig. a. showing the initial spicule without D ray; c. paratype, FÖ 87; d. holotype, Rc1. 2e. Silicotintinnabulum transitum n. sp., holotype, Rc4; f, g. Colpotrelus campanuliformis n. sp., Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582780" httpUri="https://zenodo.org/record/12582780/files/figure.png" pageId="4" pageNumber="55">Fig. 2c</figureCitation>
) of Illyrian age from the sample FÖ 87 that bears a bunch of short spinules on its distal end. Interestingly, that a rather similar
<typeStatus id="2C14E9AAC626203CFD97F8C8FDAAF898" box="[570,616,1885,1909]" pageId="5" pageNumber="56">type</typeStatus>
of short distal spinules is present on the apical horn of the species
<taxonomicName id="34AF2C8BC626203CFF3AF80BFEA9F85B" authorityName="Dumitrică" authorityYear="2024" box="[151,363,1950,1974]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="5" pageNumber="56" phylum="Radiozoa" rank="species" species="mimetica" status="sp. nov.">
<emphasis id="C1DB8B1AC626203CFF3AF80BFEA9F85B" box="[151,363,1950,1974]" italics="true" pageId="5" pageNumber="56">Gorispela mimetica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C626203CFEDEF80AFE6FF85B" box="[371,429,1951,1974]" pageId="5" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="34AF2C8BC626203CFE4FF80BFD03F85B" authorityName="Dumitrică" authorityYear="2024" box="[482,705,1950,1974]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="5" pageNumber="56" phylum="Radiozoa" rank="species" species="hungarica" status="sp. nov.">
<emphasis id="C1DB8B1AC626203CFE4FF80BFD03F85B" box="[482,705,1950,1974]" italics="true" pageId="5" pageNumber="56">Gorispela hungarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C626203CFD64F80AFCC1F85B" box="[713,771,1951,1974]" pageId="5" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
(see below), two species occurring in the same sample FÖ 87 from the Felsőörs section. Pores of thorax are very variable in shape and arrangement but are always very large.
</paragraph>
<caption id="A7D00780C626203CFF68F971FC76F8DB" ID-DOI="http://doi.org/10.5281/zenodo.12582780" ID-Zenodo-Dep="12582780" httpUri="https://zenodo.org/record/12582780/files/figure.png" pageId="5" pageNumber="56" startId="5.[197,232,1764,1785]" targetBox="[209,1395,175,1686]" targetPageId="5" targetType="figure">
<paragraph id="F3105708C626203CFF68F971FC76F8DB" blockId="5.[197,1412,1764,1846]" pageId="5" pageNumber="56">
<emphasis id="C1DB8B1AC626203CFF68F971FF3DF914" bold="true" box="[197,255,1764,1785]" pageId="5" pageNumber="56">Fig. 2</emphasis>
.
<emphasis id="C1DB8B1AC626203CFEA6F971FEF1F914" bold="true" box="[267,307,1764,1786]" pageId="5" pageNumber="56">a -d.</emphasis>
<taxonomicName id="34AF2C8BC626203CFEE6F971FDADF917" authorityName="Dumitrică" authorityYear="2024" box="[331,623,1764,1786]" class="Polycystina" family="Pseudosaturniformidae" genus="Silicotintinnabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="5" pageNumber="56" rank="species" species="formosum" status="sp. nov.">
<emphasis id="C1DB8B1AC626203CFEE6F971FDADF917" box="[331,623,1764,1786]" italics="true" pageId="5" pageNumber="56">Silicotintinnabulum formosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C626203CFDD8F970FD6EF917" box="[629,684,1765,1786]" pageId="5" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="C1DB8B1AC626203CFD1BF971FD00F914" bold="true" box="[694,706,1764,1785]" pageId="5" pageNumber="56">a</emphasis>
,
<emphasis id="C1DB8B1AC626203CFD63F971FD24F914" bold="true" box="[718,742,1764,1785]" pageId="5" pageNumber="56">a1</emphasis>
,
<emphasis id="C1DB8B1AC626203CFD5FF971FCC4F914" bold="true" box="[754,774,1764,1785]" pageId="5" pageNumber="56">b.</emphasis>
sample Rc4;
<emphasis id="C1DB8B1AC626203CFC31F971FC78F914" bold="true" box="[924,954,1764,1785]" pageId="5" pageNumber="56">a1.</emphasis>
basal view of fig.
<emphasis id="C1DB8B1AC626203CFBC3F971FB42F914" bold="true" box="[1134,1152,1764,1785]" pageId="5" pageNumber="56">a.</emphasis>
showing the initial spicule without D ray;
<emphasis id="C1DB8B1AC626203CFEF2F897FEB2F8FA" bold="true" box="[351,368,1794,1815]" pageId="5" pageNumber="56">c.</emphasis>
paratype, FÖ 87;
<emphasis id="C1DB8B1AC626203CFD90F897FD92F8FA" bold="true" box="[573,592,1794,1815]" pageId="5" pageNumber="56">d.</emphasis>
holotype, Rc1.
<emphasis id="C1DB8B1AC626203CFCA8F897FCE0F8FA" bold="true" box="[773,802,1794,1815]" pageId="5" pageNumber="56">2e.</emphasis>
<taxonomicName id="34AF2C8BC626203CFC92F897FBA3F8F5" authorityName="Dumitrică" authorityYear="2024" box="[831,1121,1794,1816]" class="Polycystina" family="Pseudosaturniformidae" genus="Silicotintinnabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="5" pageNumber="56" rank="species" species="transitum" status="sp. nov.">
<emphasis id="C1DB8B1AC626203CFC92F897FBA3F8F5" box="[831,1121,1794,1816]" italics="true" pageId="5" pageNumber="56">Silicotintinnabulum transitum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C626203CFBC4F896FB63F8F5" box="[1129,1185,1795,1816]" pageId="5" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
, holotype, Rc4;
<emphasis id="C1DB8B1AC626203CFAEAF897FA8DF8FA" bold="true" box="[1351,1359,1794,1815]" pageId="5" pageNumber="56">f</emphasis>
,
<emphasis id="C1DB8B1AC626203CFAF0F897FAADF8FA" bold="true" box="[1373,1391,1794,1815]" pageId="5" pageNumber="56">g.</emphasis>
<taxonomicName id="34AF2C8BC626203CFF68F8B5FE22F8DB" authorityName="Dumitrică" authorityYear="2024" box="[197,480,1824,1846]" class="Polycystina" family="Pseudosaturniformidae" genus="Colpotrelus" higherTaxonomySource="GBIF" kingdom="Chromista" order="Nassellaria" pageId="5" pageNumber="56" rank="species" species="campanuliformis" status="sp. nov.">
<emphasis id="C1DB8B1AC626203CFF68F8B5FE22F8DB" box="[197,480,1824,1846]" italics="true" pageId="5" pageNumber="56">Colpotrelus campanuliformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C626203CFE48F8B5FDD9F8D8" box="[485,539,1824,1845]" pageId="5" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
, Rc4. Scale bar of 50 µm is for all figures.
</paragraph>
</caption>
<paragraph id="F3105708C626203CFC9DF8E8FC6AF815" blockId="5.[816,1436,1884,2040]" pageId="5" pageNumber="56">
<emphasis id="C1DB8B1AC626203CFC9DF8E8FBD5F878" bold="true" box="[816,1047,1917,1941]" italics="true" pageId="5" pageNumber="56">Stratigraphic range</emphasis>
. Upper Anisian (Illyrian) from
<collectingCountry id="8BB81798C626203CFC9DF80AFC4FF85A" box="[816,909,1951,1975]" name="Hungary" pageId="5" pageNumber="56">Hungary</collectingCountry>
and lower Ladinian (Fassanian) of the Buchenstein Formation from Recoaro area, Vicentinian Alps,
<collectingCountry id="8BB81798C626203CFCDDF875FC66F815" box="[880,932,2016,2040]" name="Italy" pageId="5" pageNumber="56">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:identifier id="11ED2B663E5D83360295252EB0ECFAD4" type="ISSN">1842-371x</mods:identifier>
<mods:identifier id="0FBCD8181D15439B7D23FF00B283DE6A" type="Zenodo-Dep">12582776</mods:identifier>
</mods:mods>
<treatment id="7B06E61EC6282032FC65FEB7FC18F924" LSID="urn:lsid:plazi:treatment:7B06E61EC6282032FC65FEB7FC18F924" httpUri="http://treatment.plazi.org/id/7B06E61EC6282032FC65FEB7FC18F924" lastPageNumber="62" pageId="11" pageNumber="62">
<subSubSection id="BBB50483C6282032FC65FEB7FAC7FED7" box="[968,1285,290,314]" pageId="11" pageNumber="62" type="nomenclature">
<paragraph id="F3105708C6282032FC65FEB7FAC7FED7" blockId="11.[968,1285,290,314]" box="[968,1285,290,314]" pageId="11" pageNumber="62">
<heading id="A858E064C6282032FC65FEB7FAC7FED7" box="[968,1285,290,314]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="62" reason="5">
<emphasis id="C1DB8B1AC6282032FC65FEB7FAC7FED7" bold="true" box="[968,1285,290,314]" pageId="11" pageNumber="62">
<taxonomicName id="34AF2C8BC6282032FC65FEB7FB00FED7" authorityName="Dumitrică" authorityYear="2024" box="[968,1218,290,314]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="11" pageNumber="62" phylum="Radiozoa" rank="species" species="vicentiniana" status="sp. nov.">
<emphasis id="C1DB8B1AC6282032FC65FEB7FB00FED7" bold="true" box="[968,1218,290,314]" italics="true" pageId="11" pageNumber="62">Gorispela vicentiniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6282032FB64FEB7FAC7FED7" box="[1225,1285,290,314]" pageId="11" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FB9DFED6FB5FFEB6" box="[1072,1181,323,347]" pageId="11" pageNumber="62" type="description">
<paragraph id="F3105708C6282032FB9DFED6FB5FFEB6" blockId="11.[1072,1181,323,347]" box="[1072,1181,323,347]" pageId="11" pageNumber="62">
<figureCitation id="6B944B8DC6282032FB9DFED6FBBBFEB6" box="[1072,1145,323,347]" captionStart="Fig" captionStartId="10.[197,232,1730,1751]" captionText="Fig. 4. a. Gorispela victoria n. sp., holotype, Rc4; b. Gorispela mimetica n. sp., holotype, FÖ 87; c. Gorispela haeckeli n. sp., holotype, Rc4; d-g. Gorispela vicentiniana n. sp.: g.- Rc4, d. Rc2, holotype, e. apical horn of another specimen, f. Rc1d; h. Gorispela bragini n. sp., holotype., basal view of fig. 4c; f, g.- Rc1. Scale bar of 50 µm is for all figures." pageId="11" pageNumber="62">Figs. 4</figureCitation>
d-g
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FC9DFE13FC25FC1C" pageId="11" pageNumber="62" type="diagnosis">
<paragraph id="F3105708C6282032FC9DFE13FAE3FC42" blockId="11.[816,1437,389,1737]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FC9DFE13FC5CFE73" bold="true" box="[816,926,390,414]" italics="true" pageId="11" pageNumber="62">Diagnosis</emphasis>
. Shell dicyrtid bell-shaped with initial spicules without dorsal ray. Cephalis usually poreless except its distal part that can have several small circular pores in the vicinity of the thorax. Apical horn conical, usually slightly dorsally tilted, with distal part slightly three-bladed, interbladed grooves more or less visible and probably connected with the cavity of the horn (fig. 4e). Ventral horn in subhorizontal position, shorter than the apical horn and three-bladed, its basal part with a very short conical portion. Ventral ray directed to the axis of the horn. Thorax truncate conical, bearing two circumferential ridges, the last one better marked and the interval between them having usually a ring of larger pores. Distal end represented by a relatively short cylindrical body with a row of small pores below the last circumferential ridge or with small and irregularly disposed pores. Its distal end usually irregular.
</paragraph>
<paragraph id="F3105708C6282032FC9DFC2DFC25FC1C" blockId="11.[816,1437,389,1737]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FC9DFC2DFC28FC3D" bold="true" box="[816,1002,952,976]" italics="true" pageId="11" pageNumber="62">Studied material</emphasis>
.
<specimenCount id="E5A99C81C6282032FC51FC2CFB6AFC3D" box="[1020,1192,952,976]" count="4" pageId="11" pageNumber="62" type="generic">Four specimens</specimenCount>
of which one in Rc2 and three in Rc4.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FC9DFC6FFA91FBFF" box="[816,1363,1018,1042]" pageId="11" pageNumber="62" type="materials_examined">
<paragraph id="F3105708C6282032FC9DFC6FFA91FBFF" blockId="11.[816,1437,389,1737]" box="[816,1363,1018,1042]" pageId="11" pageNumber="62">
<materialsCitation id="43C75D55C6282032FC9DFC6FFA91FBFF" box="[816,1363,1018,1042]" collectionCode="MGL" pageId="11" pageNumber="62" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC6282032FC9DFC6FFC51FBFF" bold="true" box="[816,915,1018,1042]" italics="true" pageId="11" pageNumber="62">
<typeStatus id="2C14E9AAC6282032FC9DFC6FFC51FBFF" box="[816,915,1018,1042]" pageId="11" pageNumber="62" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC6282032FC0CFC6FFC2DFBFF" box="[929,1007,1018,1042]" captionStart="Fig" captionStartId="10.[197,232,1730,1751]" captionText="Fig. 4. a. Gorispela victoria n. sp., holotype, Rc4; b. Gorispela mimetica n. sp., holotype, FÖ 87; c. Gorispela haeckeli n. sp., holotype, Rc4; d-g. Gorispela vicentiniana n. sp.: g.- Rc4, d. Rc2, holotype, e. apical horn of another specimen, f. Rc1d; h. Gorispela bragini n. sp., holotype., basal view of fig. 4c; f, g.- Rc1. Scale bar of 50 µm is for all figures." pageId="11" pageNumber="62">Fig. 4d</figureCitation>
, sample Rc2, coll.
<collectionCode id="95BECFCDC6282032FB17FC6FFB38FBFF" box="[1210,1274,1018,1042]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="11" pageNumber="62">MGL</collectionCode>
.110279.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FC9DFB89FB99FB7A" pageId="11" pageNumber="62" type="description">
<paragraph id="F3105708C6282032FC9DFB89FB99FB7A" blockId="11.[816,1437,389,1737]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FC9DFB89FC71FBD9" bold="true" box="[816,947,1052,1076]" italics="true" pageId="11" pageNumber="62">Dimensions</emphasis>
. Length of shell with apical horn 110-130 µm, without apical horn 80-83 µm, of apical horn 40-50 µm, of ventral spine 28-30 µm; diameter of cephalis 43- 45 µm, of thorax 73-82 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FC9DFB35FC6CFB34" pageId="11" pageNumber="62" type="etymology">
<paragraph id="F3105708C6282032FC9DFB35FC6CFB34" blockId="11.[816,1437,389,1737]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FC9DFB35FC67FB55" bold="true" box="[816,933,1184,1208]" italics="true" pageId="11" pageNumber="62">Etymology</emphasis>
. From its occurrence in the Vicentinian Alps, North
<collectingCountry id="8BB81798C6282032FCD8FB54FC6BFB34" box="[885,937,1217,1241]" name="Italy" pageId="11" pageNumber="62">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FC9DFB77FC18F924" pageId="11" pageNumber="62" type="discussion">
<paragraph id="F3105708C6282032FC9DFB77FC77F96A" blockId="11.[816,1437,389,1737]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FC9DFB77FC50FB17" bold="true" box="[816,914,1250,1274]" italics="true" pageId="11" pageNumber="62">Remarks</emphasis>
.
<taxonomicName id="34AF2C8BC6282032FC0EFB77FB5EFB17" authorityName="Dumitrică" authorityYear="2024" box="[931,1180,1250,1274]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="11" pageNumber="62" phylum="Radiozoa" rank="species" species="vicentiniana" status="sp. nov.">
<emphasis id="C1DB8B1AC6282032FC0EFB77FB5EFB17" box="[931,1180,1250,1274]" italics="true" pageId="11" pageNumber="62">Gorispela vicentiniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6282032FB0BFB76FB21FB17" box="[1190,1251,1251,1274]" pageId="11" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
differs from the other species above mentioned by having the unbladed proximal part of ventral horn very short or almost absent. It is very close to
<taxonomicName id="34AF2C8BC6282032FC5CFAD0FB7EFAB0" authorityName="Dumitrică" authorityYear="2024" box="[1009,1212,1349,1373]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="11" pageNumber="62" phylum="Radiozoa" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="C1DB8B1AC6282032FC5CFAD0FB7EFAB0" box="[1009,1212,1349,1373]" italics="true" pageId="11" pageNumber="62">Gorispela haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6282032FB6EFAD3FB3CFAB0" box="[1219,1278,1350,1373]" pageId="11" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
, from which it differs by having the pores in the space between the circumferential rings arranged usually in a single circumferential row and the circumferential rings narrower. Also, it is very close to
<taxonomicName id="34AF2C8BC6282032FB0EFA5CFA43FA0C" authorityName="Dumitrică" authorityYear="2024" box="[1187,1409,1481,1505]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="11" pageNumber="62" phylum="Radiozoa" rank="species" species="fenestrata" status="sp. nov.">
<emphasis id="C1DB8B1AC6282032FB0EFA5CFA43FA0C" box="[1187,1409,1481,1505]" italics="true" pageId="11" pageNumber="62">Gorispela fenestrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6282032FA25FA5FFC8DF9EF" pageId="11" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
by having the pores between circumferential rings disposed in single rows, from which it differs by not having the ventral horn with a conical proximal portion, the pores between rows smaller and the rows closer to one another.
</paragraph>
<paragraph id="F3105708C6282032FC9DF905FC18F924" blockId="11.[816,1437,389,1737]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FC9DF905FBD3F945" bold="true" box="[816,1041,1680,1704]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FC9DF905FBC8F945" bold="true" box="[816,1034,1680,1704]" italics="true" pageId="11" pageNumber="62">Stratigraphic range</emphasis>
.
</emphasis>
Buchenstein Formation of Recoaro, lower Ladinian.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="38D51A46512B36CF8714DA89E8E9A127">New Middle Triassic Bell-Shaped Nassellarian Radiolaria From Alpine And Carpathian Areas</mods:title>
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<treatment id="7B06E61EC6282032FEE6FBCBFBDAFF15" LSID="urn:lsid:plazi:treatment:7B06E61EC6282032FEE6FBCBFBDAFF15" httpUri="http://treatment.plazi.org/id/7B06E61EC6282032FEE6FBCBFBDAFF15" lastPageNumber="62" pageId="11" pageNumber="62">
<subSubSection id="BBB50483C6282032FEE6FBCBFD8DFB9B" box="[331,591,1118,1142]" pageId="11" pageNumber="62" type="nomenclature">
<paragraph id="F3105708C6282032FEE6FBCBFD8DFB9B" blockId="11.[331,591,1118,1142]" box="[331,591,1118,1142]" pageId="11" pageNumber="62">
<heading id="A858E064C6282032FEE6FBCBFD8DFB9B" box="[331,591,1118,1142]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="62" reason="5">
<emphasis id="C1DB8B1AC6282032FEE6FBCBFD8DFB9B" bold="true" box="[331,591,1118,1142]" pageId="11" pageNumber="62">
<taxonomicName id="34AF2C8BC6282032FEE6FBCBFDCFFB9B" authorityName="Dumitrică" authorityYear="2024" box="[331,525,1118,1142]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="11" pageNumber="62" phylum="Radiozoa" rank="species" species="bragini" status="sp. nov.">
<emphasis id="C1DB8B1AC6282032FEE6FBCBFDCFFB9B" bold="true" box="[331,525,1118,1142]" italics="true" pageId="11" pageNumber="62">Gorispela bragini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6282032FDBEFBCBFD8DFB9B" box="[531,591,1118,1142]" pageId="11" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FE22FBEAFDC9FB7A" box="[399,523,1151,1175]" pageId="11" pageNumber="62" type="description">
<paragraph id="F3105708C6282032FE22FBEAFDC9FB7A" blockId="11.[399,523,1151,1175]" box="[399,523,1151,1175]" pageId="11" pageNumber="62">
<figureCitation id="6B944B8DC6282032FE22FBEAFE25FB7A" box="[399,487,1151,1175]" captionStart="Fig" captionStartId="10.[197,232,1730,1751]" captionText="Fig. 4. a. Gorispela victoria n. sp., holotype, Rc4; b. Gorispela mimetica n. sp., holotype, FÖ 87; c. Gorispela haeckeli n. sp., holotype, Rc4; d-g. Gorispela vicentiniana n. sp.: g.- Rc4, d. Rc2, holotype, e. apical horn of another specimen, f. Rc1d; h. Gorispela bragini n. sp., holotype., basal view of fig. 4c; f, g.- Rc1. Scale bar of 50 µm is for all figures." pageId="11" pageNumber="62">Figs. 4h</figureCitation>
,
<figureCitation id="6B944B8DC6282032FE5FFBEAFDC9FB7A" box="[498,523,1151,1175]" captionStart="Fig" captionStartId="13.[197,232,1409,1430]" captionTargetBox="[213,1380,173,1369]" captionTargetId="figure-420@13.[197,1411,160,1381]" captionTargetPageId="13" captionText="Fig. 5. a, b. Gorispela tuba n. sp., a. Rc4, b. holotype, FÖ 87; c. Gorispela bragini n. sp., holotype, Rc4 (see also fig. 3h); d-f. Pozsvartia multicingulata n. sp., d. holotype, FÖ 87, e. CRH, f. CR25, D dorsal spine. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582786" httpUri="https://zenodo.org/record/12582786/files/figure.png" pageId="11" pageNumber="62">5c</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FF3AFB54FD77F98B" pageId="11" pageNumber="62" type="diagnosis">
<paragraph id="F3105708C6282032FF3AFB54FD86F9A8" blockId="11.[151,772,1217,2068]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FF3AFB54FEC7FB34" bold="true" box="[151,261,1217,1241]" italics="true" pageId="11" pageNumber="62">Diagnosis</emphasis>
. Shell bell-shaped with practically poreles cephalis. Apical horn slightly tilted dorsally, pointed, hollow on the proximal portion and slightly three-bladed on the distal one. Ventral horn three-bladed and pyramidal arising directly from the cephalis without a conical intermediary portion. Ventral ray of initial spicule connected with the centre of the horn. Collar boundary marked by a slight constriction. Thorax thin-walled, truncate conical and bearing 2-3 circumferential ridges bearing in the interridge areas a row of circular, oval or rectangular windows. Distal end with a very narrow and vertically directed circumferential band.
</paragraph>
<paragraph id="F3105708C6282032FF3AF9DBFD77F98B" blockId="11.[151,772,1217,2068]" box="[151,693,1614,1638]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FF3AF9DBFE8FF98B" bold="true" box="[151,333,1614,1638]" italics="true" pageId="11" pageNumber="62">Studied material</emphasis>
.
<specimenCount id="E5A99C81C6282032FEF7F9DBFDCCF98B" box="[346,526,1614,1638]" count="3" pageId="11" pageNumber="62" type="generic">Three specimens</specimenCount>
in sample Rc4.
</paragraph>
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<subSubSection id="BBB50483C6282032FF3AF9FAFEF3F945" pageId="11" pageNumber="62" type="materials_examined">
<paragraph id="F3105708C6282032FF3AF9FAFEF3F945" blockId="11.[151,772,1217,2068]" pageId="11" pageNumber="62">
<materialsCitation id="43C75D55C6282032FF3AF9FAFEF3F945" collectionCode="MGL" pageId="11" pageNumber="62" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC6282032FF3AF9FAFF38F96A" bold="true" box="[151,250,1647,1671]" italics="true" pageId="11" pageNumber="62">
<typeStatus id="2C14E9AAC6282032FF3AF9FAFF38F96A" box="[151,250,1647,1671]" pageId="11" pageNumber="62" type="holotype">Holotype</typeStatus>
</emphasis>
. The illustrated specimen, sample Rc4, coll.
<collectionCode id="95BECFCDC6282032FF3AF905FF15F945" box="[151,215,1680,1704]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="11" pageNumber="62">MGL</collectionCode>
.110278.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FF3AF927FEBFF8C0" pageId="11" pageNumber="62" type="description">
<paragraph id="F3105708C6282032FF3AF927FEBFF8C0" blockId="11.[151,772,1217,2068]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FF3AF927FED8F927" bold="true" box="[151,282,1714,1738]" italics="true" pageId="11" pageNumber="62">Dimensions</emphasis>
(measured only on
<typeStatus id="2C14E9AAC6282032FE43F924FD92F924" box="[494,592,1713,1737]" pageId="11" pageNumber="62" type="holotype">holotype</typeStatus>
). Length of shell with apical horn 130 µm, length of horn 60 µm, of cephalis 47 µm, of thorax 48 µm, diameter of cephalis 48 µm, of thorax 85 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6282032FF3AF8A3FEDFF87D" pageId="11" pageNumber="62" type="etymology">
<paragraph id="F3105708C6282032FF3AF8A3FEDFF87D" blockId="11.[151,772,1217,2068]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FF3AF8A3FECEF8A3" bold="true" box="[151,268,1846,1870]" italics="true" pageId="11" pageNumber="62">Etymology</emphasis>
. The species is dedicated to Dr. Nikita Bragin for his contribution to the knowledge of Triassic radiolarians.
</paragraph>
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<subSubSection id="BBB50483C6282032FF3AF80CFBDAFF15" pageId="11" pageNumber="62" type="discussion">
<paragraph id="F3105708C6282032FF3AF80CFB77FF5B" blockId="11.[151,772,1217,2068]" lastBlockId="11.[816,1436,158,248]" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FF3AF80CFF3BF85C" bold="true" box="[151,249,1945,1969]" italics="true" pageId="11" pageNumber="62">Remarks</emphasis>
. By its morphology, this species resembles
<taxonomicName id="34AF2C8BC6282032FF3AF82FFEB6F83F" authorityName="Dumitrică" authorityYear="2024" box="[151,372,1978,2002]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="11" pageNumber="62" phylum="Radiozoa" rank="species" species="fenestrata" status="sp. nov.">
<emphasis id="C1DB8B1AC6282032FF3AF82FFEB6F83F" box="[151,372,1978,2002]" italics="true" pageId="11" pageNumber="62">Gorispela fenestrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6282032FED7F82EFE74F83F" box="[378,438,1979,2002]" pageId="11" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
, from which it differs by being smaller and by not having the intermediary conical hollow portion between the distal three-bladed part of the ventral horn and the cephalic cavity.
</paragraph>
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<emphasis id="C1DB8B1AC6282032FC9DFF2AFBDBFF3A" bold="true" box="[816,1049,191,215]" italics="true" pageId="11" pageNumber="62">Stratigraphic range</emphasis>
. Buchenstein Formation, lower Ladinian (Fassanian).
</paragraph>
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</treatment>
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<heading id="A858E064C6282032FC59F961FB1BF8E1" box="[1012,1241,1780,1804]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="62" reason="5">
<emphasis id="C1DB8B1AC6282032FC59F961FB1BF8E1" bold="true" box="[1012,1241,1780,1804]" pageId="11" pageNumber="62">
<taxonomicName id="34AF2C8BC6282032FC59F961FB54F8E1" authorityName="Dumitrică" authorityYear="2024" box="[1012,1174,1780,1804]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="11" pageNumber="62" phylum="Radiozoa" rank="species" species="tuba" status="sp. nov.">
<emphasis id="C1DB8B1AC6282032FC59F961FB54F8E1" bold="true" box="[1012,1174,1780,1804]" italics="true" pageId="11" pageNumber="62">Gorispela tuba</emphasis>
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<taxonomicNameLabel id="DAE83661C6282032FB30F961FB1BF8E1" box="[1181,1241,1780,1804]" pageId="11" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<paragraph id="F3105708C6282032FB82F881FB5CF8C1" blockId="11.[1071,1182,1812,1836]" box="[1071,1182,1812,1836]" pageId="11" pageNumber="62">
<figureCitation id="6B944B8DC6282032FB82F881FB5CF8C1" box="[1071,1182,1812,1836]" captionStart="Fig" captionStartId="13.[197,232,1409,1430]" captionTargetBox="[213,1380,173,1369]" captionTargetId="figure-420@13.[197,1411,160,1381]" captionTargetPageId="13" captionText="Fig. 5. a, b. Gorispela tuba n. sp., a. Rc4, b. holotype, FÖ 87; c. Gorispela bragini n. sp., holotype, Rc4 (see also fig. 3h); d-f. Pozsvartia multicingulata n. sp., d. holotype, FÖ 87, e. CRH, f. CR25, D dorsal spine. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582786" httpUri="https://zenodo.org/record/12582786/files/figure.png" pageId="11" pageNumber="62">Figs. 5a, b</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="F3105708C6282035FC9DF8C2FEB1FE83" blockId="11.[816,1437,1879,2068]" lastBlockId="12.[151,771,176,895]" lastPageId="12" lastPageNumber="63" pageId="11" pageNumber="62">
<emphasis id="C1DB8B1AC6282032FC9DF8C2FC5CF882" bold="true" box="[816,926,1879,1903]" italics="true" pageId="11" pageNumber="62">Diagnosis</emphasis>
. Shell dicyrtid, conical, trumpet-shaped and smooth-surfaced with distal portion expanded. Apical horn conical, pointed and slightly dorsally tilted, its inner side partly hollow on half proximal portion. Ventral horn conical and pointed, with hollow proximal portion. Cephalis impossible to separate from thorax, most part of its wall imperforated or with very small and sparse rounded pores. Pores start increasing in size and frequency from the level of the MB, where on consider that is the boundary between the cephalis and thorax. On thorax pores are larger but of different sizes and irregularly arranged.
</paragraph>
<paragraph id="F3105708C62F2035FF3AFEE2FEAFFE5D" blockId="12.[151,771,176,895]" pageId="12" pageNumber="63">
<emphasis id="C1DB8B1AC62F2035FF3AFEE2FE93FE62" bold="true" box="[151,337,375,399]" italics="true" pageId="12" pageNumber="63">Studied material</emphasis>
.
<specimenCount id="E5A99C81C62F2035FECCFEE2FDDBFE62" box="[353,537,375,399]" count="3" pageId="12" pageNumber="63" type="generic">Three specimens</specimenCount>
of which two in FÖ 87, and one in Rc 4.
</paragraph>
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<materialsCitation id="43C75D55C62F2035FF3AFE2CFD29FE3C" box="[151,747,441,466]" collectionCode="MGL" country="Hungary" location="Hungary" pageId="12" pageNumber="63" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC62F2035FF3AFE2CFF38FE3C" bold="true" box="[151,250,441,465]" italics="true" pageId="12" pageNumber="63">
<typeStatus id="2C14E9AAC62F2035FF3AFE2CFF38FE3C" box="[151,250,441,465]" pageId="12" pageNumber="63" type="holotype">Holotype</typeStatus>
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.
<figureCitation id="6B944B8DC62F2035FEA5FE2CFE97FE3C" box="[264,341,441,465]" captionStart="Fig" captionStartId="13.[197,232,1409,1430]" captionTargetBox="[213,1380,173,1369]" captionTargetId="figure-420@13.[197,1411,160,1381]" captionTargetPageId="13" captionText="Fig. 5. a, b. Gorispela tuba n. sp., a. Rc4, b. holotype, FÖ 87; c. Gorispela bragini n. sp., holotype, Rc4 (see also fig. 3h); d-f. Pozsvartia multicingulata n. sp., d. holotype, FÖ 87, e. CRH, f. CR25, D dorsal spine. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582786" httpUri="https://zenodo.org/record/12582786/files/figure.png" pageId="12" pageNumber="63">Fig. 5b</figureCitation>
, FÖ 87,
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, coll.
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.110280.
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<emphasis id="C1DB8B1AC62F2035FF3AFE4FFED8FE1F" bold="true" box="[151,282,474,498]" italics="true" pageId="12" pageNumber="63">Dimensions</emphasis>
. Total length of shell 128-143 µm, of shell without apical horn 80-90 µm, diameter of cephalis at the level of MB 40-48 µm, of distal end 80-103 µm.
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<emphasis id="C1DB8B1AC62F2035FF3AFDABFECEFDBB" bold="true" box="[151,268,574,598]" italics="true" pageId="12" pageNumber="63">Etymology</emphasis>
. From the Latin
<taxonomicName id="34AF2C8BC62F2035FE6DFDA8FE2DFDB8" authorityName="Dumitrică" authorityYear="2024" box="[448,495,573,597]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="12" pageNumber="63" phylum="Radiozoa" rank="species" species="tuba">
<emphasis id="C1DB8B1AC62F2035FE6DFDA8FE2DFDB8" box="[448,495,573,597]" italics="true" pageId="12" pageNumber="63">tuba</emphasis>
</taxonomicName>
- trumpet.
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<subSubSection id="BBB50483C62F2035FF3AFDCAFD26FC92" pageId="12" pageNumber="63" type="discussion">
<paragraph id="F3105708C62F2035FF3AFDCAFF18FCF1" blockId="12.[151,771,176,895]" pageId="12" pageNumber="63">
<emphasis id="C1DB8B1AC62F2035FF3AFDCAFF3BFD9A" bold="true" box="[151,249,607,631]" italics="true" pageId="12" pageNumber="63">Remarks</emphasis>
.
<taxonomicName id="34AF2C8BC62F2035FEA4FDCBFE6EFD9B" authorityName="Dumitrică" authorityYear="2024" box="[265,428,606,630]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="12" pageNumber="63" phylum="Radiozoa" rank="species" species="tuba" status="sp. nov.">
<emphasis id="C1DB8B1AC62F2035FEA4FDCBFE6EFD9B" box="[265,428,606,630]" italics="true" pageId="12" pageNumber="63">Gorispela tuba</emphasis>
</taxonomicName>
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has in common with the other species of this genus the general size of the shell and the hollow structure of the apical and ventral horns. It only differs by the absence of circumferential ridges and by the absence of three-bladed distal portions of the two horns.
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<emphasis id="C1DB8B1AC62F2035FF3AFCB0FEBCFCD0" bold="true" box="[151,382,805,829]" italics="true" pageId="12" pageNumber="63">Stratigraphic range</emphasis>
. Upper Anisian (Illyrian) from
<collectingCountry id="8BB81798C62F2035FF3AFCD2FF35FCB2" box="[151,247,839,863]" name="Hungary" pageId="12" pageNumber="63">Hungary</collectingCountry>
, sample FÖ 87, to lower Ladinian (Fassanian) of the Buchenstein Formation from Recoaro, sample Rc4.
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<emphasis id="C1DB8B1AC6292033FEE8F802FDD1F842" bold="true" box="[325,531,1943,1967]" italics="true" pageId="10" pageNumber="61">Gorispela haeckeli</emphasis>
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<figureCitation id="6B944B8DC6292033FE05F822FE30F822" box="[424,498,1975,1999]" captionStart="Fig" captionStartId="10.[197,232,1730,1751]" captionText="Fig. 4. a. Gorispela victoria n. sp., holotype, Rc4; b. Gorispela mimetica n. sp., holotype, FÖ 87; c. Gorispela haeckeli n. sp., holotype, Rc4; d-g. Gorispela vicentiniana n. sp.: g.- Rc4, d. Rc2, holotype, e. apical horn of another specimen, f. Rc1d; h. Gorispela bragini n. sp., holotype., basal view of fig. 4c; f, g.- Rc1. Scale bar of 50 µm is for all figures." pageId="10" pageNumber="61">Fig. 4c</figureCitation>
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<heading id="A858E064C62A2030FEE7FA5FFD93FA0F" box="[330,593,1482,1506]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="60" reason="5">
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<emphasis id="C1DB8B1AC62A2030FEE7FA5FFDCCFA0F" bold="true" box="[330,526,1482,1506]" italics="true" pageId="9" pageNumber="60">Gorispela victoria</emphasis>
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<paragraph id="F3105708C62A2030FE05FA7EFE30F9EE" blockId="9.[424,498,1515,1539]" box="[424,498,1515,1539]" pageId="9" pageNumber="60">
<figureCitation id="6B944B8DC62A2030FE05FA7EFE30F9EE" box="[424,498,1515,1539]" captionStart="Fig" captionStartId="10.[197,232,1730,1751]" captionText="Fig. 4. a. Gorispela victoria n. sp., holotype, Rc4; b. Gorispela mimetica n. sp., holotype, FÖ 87; c. Gorispela haeckeli n. sp., holotype, Rc4; d-g. Gorispela vicentiniana n. sp.: g.- Rc4, d. Rc2, holotype, e. apical horn of another specimen, f. Rc1d; h. Gorispela bragini n. sp., holotype., basal view of fig. 4c; f, g.- Rc1. Scale bar of 50 µm is for all figures." pageId="9" pageNumber="60">Fig. 4a</figureCitation>
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<paragraph id="F3105708C62A2030FF3AF9B8FABAFED7" blockId="9.[151,772,1581,2069]" lastBlockId="9.[816,1437,158,744]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FF3AF9B8FEC7F9A8" bold="true" box="[151,261,1581,1605]" italics="true" pageId="9" pageNumber="60">Diagnosis</emphasis>
. Shell short conical with one circumferential ridge around the middle part of thorax. Initial spicule without dorsal ray. Apical and ventral rays prolonged outside into two equal, robust and straight horns situated practically symmetrical relative to the axis of shell and diverging from one another under an angle of 100°. Horns with proximal cylindrical portion short and hollow, and middle and distal portions three-bladed. Blades of apical horn bear a crown of small teeth, whereas the ventral horn bears only one tooth on the external blade. Both horns pyramidal beyond the level of the tooth. Cephalic cavity triangular in lateral view and poreless. Thorax truncate conical with about two circumferential rows of large pores on the distal end in an intercalary position. At its middle part there is a very well marked, thin and well-extending circumferential ring. Distal part of thorax below the circumferential ridge very short, inverted conical and with irregular distal border. It has large pores in the vicinity of the circumferential ridge and smaller ones and irregularly disposed on its distal part.
</paragraph>
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<emphasis id="C1DB8B1AC62A2030FC9DFED1FC2FFEB6" box="[816,1005,324,348]" italics="true" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FC9DFED1FC24FEB1" bold="true" box="[816,998,324,348]" italics="true" pageId="9" pageNumber="60">Studied material</emphasis>
.
</emphasis>
A single specimen in the sample Rc4.
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<typeStatus id="2C14E9AAC62A2030FC9DFEF0FC51FE90" box="[816,915,357,381]" pageId="9" pageNumber="60" type="holotype">Holotype</typeStatus>
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.
<figureCitation id="6B944B8DC62A2030FC0CFEF1FC2FFE91" box="[929,1005,356,380]" captionStart="Fig" captionStartId="10.[197,232,1730,1751]" captionText="Fig. 4. a. Gorispela victoria n. sp., holotype, Rc4; b. Gorispela mimetica n. sp., holotype, FÖ 87; c. Gorispela haeckeli n. sp., holotype, Rc4; d-g. Gorispela vicentiniana n. sp.: g.- Rc4, d. Rc2, holotype, e. apical horn of another specimen, f. Rc1d; h. Gorispela bragini n. sp., holotype., basal view of fig. 4c; f, g.- Rc1. Scale bar of 50 µm is for all figures." pageId="9" pageNumber="60">Fig. 4a</figureCitation>
, coll.
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.110275.
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<emphasis id="C1DB8B1AC62A2030FC9DFE13FC71FE73" bold="true" box="[816,947,390,414]" italics="true" pageId="9" pageNumber="60">Dimensions</emphasis>
. Length of shell without horns 72 µm, diameter of base of cephalis 40 µm, of circumferential ridge 87 µm.
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<subSubSection id="BBB50483C62A2030FC9DFE7CFB07FDCF" pageId="9" pageNumber="60" type="etymology">
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<emphasis id="C1DB8B1AC62A2030FC9DFE7CFC67FDEC" bold="true" box="[816,933,489,513]" italics="true" pageId="9" pageNumber="60">Etymology</emphasis>
. Due to the position of the two horns signifying victory,
<taxonomicName id="34AF2C8BC62A2030FC51FD9EFB8DFDCF" authorityName="Dumitrică" authorityYear="2024" box="[1020,1103,523,546]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="victoria">
<emphasis id="C1DB8B1AC62A2030FC51FD9EFB8DFDCF" box="[1020,1103,523,546]" italics="true" pageId="9" pageNumber="60">victoria</emphasis>
</taxonomicName>
in Latin.
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<emphasis id="C1DB8B1AC62A2030FC9DFDBEFC50FDAE" bold="true" box="[816,914,555,579]" italics="true" pageId="9" pageNumber="60">Remarks</emphasis>
. Structurally, this species is close to
<taxonomicName id="34AF2C8BC62A2030FA9FFDBEFC65FD89" authorityName="Dumitrică" authorityYear="2024" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="cornubovis" status="sp. nov.">
<emphasis id="C1DB8B1AC62A2030FA9FFDBEFC65FD89" italics="true" pageId="9" pageNumber="60">Gorispela cornubovis</emphasis>
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, from which it differs especially by having straight horns and wide conical thorax with large pores.
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<emphasis id="C1DB8B1AC62A2030FC9DFD25FBDBFD25" bold="true" box="[816,1049,688,712]" italics="true" pageId="9" pageNumber="60">Stratigraphic range</emphasis>
. Buchenstein Formation, lower Ladinian.
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<treatment id="7B06E61EC62A2033FC77FC86FDB1F881" LSID="urn:lsid:plazi:treatment:7B06E61EC62A2033FC77FC86FDB1F881" httpUri="http://treatment.plazi.org/id/7B06E61EC62A2033FC77FC86FDB1F881" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60">
<subSubSection id="BBB50483C62A2030FC77FC86FB31FCC6" box="[986,1267,787,811]" pageId="9" pageNumber="60" type="nomenclature">
<paragraph id="F3105708C62A2030FC77FC86FB31FCC6" blockId="9.[986,1267,787,811]" box="[986,1267,787,811]" pageId="9" pageNumber="60">
<heading id="A858E064C62A2030FC77FC86FB31FCC6" box="[986,1267,787,811]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="60" reason="5">
<emphasis id="C1DB8B1AC62A2030FC77FC86FB31FCC6" bold="true" box="[986,1267,787,811]" pageId="9" pageNumber="60">
<taxonomicName id="34AF2C8BC62A2030FC77FC86FB72FCC6" authorityName="Dumitrică" authorityYear="2024" box="[986,1200,787,811]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="mimetica" status="sp. nov.">
<emphasis id="C1DB8B1AC62A2030FC77FC86FB72FCC6" bold="true" box="[986,1200,787,811]" italics="true" pageId="9" pageNumber="60">Gorispela mimetica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62A2030FB1AFC86FB31FCC6" box="[1207,1267,787,811]" pageId="9" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2030FBECFCA6FB4EFCA6" box="[1089,1164,819,843]" pageId="9" pageNumber="60" type="description">
<paragraph id="F3105708C62A2030FBECFCA6FB4EFCA6" blockId="9.[1089,1164,819,843]" box="[1089,1164,819,843]" pageId="9" pageNumber="60">
<figureCitation id="6B944B8DC62A2030FBECFCA6FB4EFCA6" box="[1089,1164,819,843]" captionStart="Fig" captionStartId="10.[197,232,1730,1751]" captionText="Fig. 4. a. Gorispela victoria n. sp., holotype, Rc4; b. Gorispela mimetica n. sp., holotype, FÖ 87; c. Gorispela haeckeli n. sp., holotype, Rc4; d-g. Gorispela vicentiniana n. sp.: g.- Rc4, d. Rc2, holotype, e. apical horn of another specimen, f. Rc1d; h. Gorispela bragini n. sp., holotype., basal view of fig. 4c; f, g.- Rc1. Scale bar of 50 µm is for all figures." pageId="9" pageNumber="60">Fig. 4b</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2030FC9DFCE3FC3FFAD0" pageId="9" pageNumber="60" type="diagnosis">
<paragraph id="F3105708C62A2030FC9DFCE3FC25FB17" blockId="9.[816,1437,886,2068]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FC9DFCE3FC5CFC63" bold="true" box="[816,926,886,910]" italics="true" pageId="9" pageNumber="60">Diagnosis</emphasis>
. Shell conical, bell-shaped with robust, conical and hollow apical and ventral horns. Apical horn practically axial bearing a few distal spinules on its distal end, whereas the ventral one has no spinules and is horizontally positioned. Both horns have also one or two large pores on the distal end of their hollow portion. Cephalis poreless except its distal part that has a series of small circular and irregularly disposed pores. Thorax truncate conical with pores that increase rapidly in diameter towards the distal end. This end has a protruding distal circumferential ring and a vertically positioned and thin distal girdle.
</paragraph>
<paragraph id="F3105708C62A2030FC9DFA96FC3FFAD0" blockId="9.[816,1437,886,2068]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FC9DFA96FC29FAF6" bold="true" box="[816,1003,1283,1307]" italics="true" pageId="9" pageNumber="60">Studied material</emphasis>
. A single specimen in sample FÖ 87, Felsőörs,
<collectingCountry id="8BB81798C62A2030FC35FAB0FC3BFAD0" box="[920,1017,1317,1341]" name="Hungary" pageId="9" pageNumber="60">Hungary</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2030FC9DFAD3FC08FA93" pageId="9" pageNumber="60" type="materials_examined">
<paragraph id="F3105708C62A2030FC9DFAD3FC08FA93" blockId="9.[816,1437,886,2068]" pageId="9" pageNumber="60">
<materialsCitation id="43C75D55C62A2030FC9DFAD3FC08FA93" collectionCode="MGL" pageId="9" pageNumber="60" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC62A2030FC9DFAD3FC51FAB3" bold="true" box="[816,915,1350,1374]" italics="true" pageId="9" pageNumber="60">
<typeStatus id="2C14E9AAC62A2030FC9DFAD3FC51FAB3" box="[816,915,1350,1374]" pageId="9" pageNumber="60" type="holotype">Holotype</typeStatus>
</emphasis>
. The specimen from
<figureCitation id="6B944B8DC62A2030FB4DFAD0FA8AFAB0" box="[1248,1352,1349,1373]" captionStart="Fig" captionStartId="8.[197,232,1750,1771]" captionTargetBox="[212,1390,203,1661]" captionTargetId="figure-171@8.[197,1410,178,1721]" captionTargetPageId="8" captionText="Fig. 3. a, a1. Gorispela cornubovis n. sp., holotype in lateral and basal views, respectively, Rc4; b. Gorispela fenestrata n. sp., holotype in lateral view, BV 85-70; c-e. Gorispela hungarica n. sp., FÖ 87, e. holotype. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582784" httpUri="https://zenodo.org/record/12582784/files/figure.png" pageId="9" pageNumber="60">Fig. 3b</figureCitation>
, coll.
<collectionCode id="95BECFCDC62A2030FC9DFAF3FCADFA93" box="[816,879,1382,1406]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="9" pageNumber="60">MGL</collectionCode>
.110276.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2030FC9DFA1DFBE7FA0F" pageId="9" pageNumber="60" type="description">
<paragraph id="F3105708C62A2030FC9DFA1DFBE7FA0F" blockId="9.[816,1437,886,2068]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FC9DFA1DFC71FA4D" bold="true" box="[816,947,1416,1440]" italics="true" pageId="9" pageNumber="60">Dimensions</emphasis>
. Length of shell with apical horn 138 µm, without apical horn 92 µm, diameter of cephalis 43 µm, of distal ridge 126 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2030FC9DFA7EFC02F98B" pageId="9" pageNumber="60" type="etymology">
<paragraph id="F3105708C62A2030FC9DFA7EFC02F98B" blockId="9.[816,1437,886,2068]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FC9DFA7EFC67F9EE" bold="true" box="[816,933,1515,1539]" italics="true" pageId="9" pageNumber="60">Etymology</emphasis>
. Due to its thorax, the morphology of which resembles perfectly that of the thorax of
<taxonomicName id="34AF2C8BC62A2030FC9DF9B8FBBCF9A8" authorityName="Dumitrică" authorityYear="2024" box="[816,1150,1581,1605]" class="Polycystina" family="Oertlispongidae" genus="Silicotinntinabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="formosum" status="sp. nov.">
<emphasis id="C1DB8B1AC62A2030FC9DF9B8FBBCF9A8" box="[816,1150,1581,1605]" italics="true" pageId="9" pageNumber="60">Silicotinntinabulum formosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62A2030FB23F9B8FB13F9A9" box="[1166,1233,1581,1604]" pageId="9" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
occurring in the same sample.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2033FC9DF9FAFDB1F881" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60" type="discussion">
<paragraph id="F3105708C62A2033FC9DF9FAFED4F8A6" blockId="9.[816,1437,886,2068]" lastBlockId="10.[151,627,1843,1900]" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FC9DF9FAFC50F96A" bold="true" box="[816,914,1647,1671]" italics="true" pageId="9" pageNumber="60">Remarks</emphasis>
. This species combines the morphology of two species assigned to two coeval genera:
<taxonomicName id="34AF2C8BC62A2030FC9DF924FBBBF924" authorityName="Dumitrică" authorityYear="2024" box="[816,1145,1713,1737]" class="Polycystina" family="Oertlispongidae" genus="Silicotinntinabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="formosum" status="n. sp. and species">
<emphasis id="C1DB8B1AC62A2030FC9DF924FBBBF924" box="[816,1145,1713,1737]" italics="true" pageId="9" pageNumber="60">Silicotinntinabulum formosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62A2030FB28F927FA8CF924" box="[1157,1358,1713,1737]" pageId="9" pageNumber="60" rank="species">n. sp. and species</taxonomicNameLabel>
of the genus
<taxonomicName id="34AF2C8BC62A2030FCD7F947FC27F907" authorityName="Dumitrică" authorityYear="2024" box="[890,997,1746,1770]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="genus">
<emphasis id="C1DB8B1AC62A2030FCD7F947FC27F907" box="[890,997,1746,1770]" italics="true" pageId="9" pageNumber="60">Gorispela</emphasis>
</taxonomicName>
due the presence of a robust and long ventral horn. From the former species it has the cephalis without dorsal (D) ray, and the ventral horn and thorax completely similar; from the genus
<taxonomicName id="34AF2C8BC62A2030FB12F8A0FAE8F8A0" authorityName="Dumitrică" authorityYear="2024" box="[1215,1322,1845,1869]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="genus">
<emphasis id="C1DB8B1AC62A2030FB12F8A0FAE8F8A0" box="[1215,1322,1845,1869]" italics="true" pageId="9" pageNumber="60">Gorispela</emphasis>
</taxonomicName>
it has the same
<typeStatus id="2C14E9AAC62A2030FCDDF8C2FC5CF882" box="[880,926,1879,1903]" pageId="9" pageNumber="60">type</typeStatus>
of ventral horn. Moreover, its apical horn has short spinules at the distal end similar to the specimen of the species
<taxonomicName id="34AF2C8BC62A2030FC01F80CFB2DF85C" authorityName="Dumitrică" authorityYear="2024" box="[940,1263,1945,1969]" class="Polycystina" family="Oertlispongidae" genus="Silicotinntinabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="formosum" status="sp. nov.">
<emphasis id="C1DB8B1AC62A2030FC01F80CFB2DF85C" box="[940,1263,1945,1969]" italics="true" pageId="9" pageNumber="60">Silicotinntinabulum formosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62A2030FB5BF80FFAF2F85C" box="[1270,1328,1946,1969]" pageId="9" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
occurring in the same fauna of the sample FÖ 87 from Felsőörs (horizontal gene transfer?). What differs is the apical horn that is not three-bladed, as in
<taxonomicName id="34AF2C8BC62A2030FBD2F868FACEF7F9" authorityName="Dumitrică" authorityYear="2024" box="[1151,1292,2044,2068]" class="Polycystina" family="Oertlispongidae" genus="Silicotinntinabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="formosum">
<emphasis id="C1DB8B1AC62A2030FBD2F868FACEF7F9" box="[1151,1292,2044,2068]" italics="true" pageId="9" pageNumber="60">S. formosum</emphasis>
</taxonomicName>
, but conical and hollow.
</paragraph>
<caption id="A7D00780C6292033FF68F957FA83F8FE" pageId="10" pageNumber="61" startId="10.[197,232,1730,1751]" targetType="figure">
<paragraph id="F3105708C6292033FF68F957FA83F8FE" blockId="10.[197,1408,1730,1811]" pageId="10" pageNumber="61">
<emphasis id="C1DB8B1AC6292033FF68F957FF3DF93A" bold="true" box="[197,255,1730,1751]" pageId="10" pageNumber="61">Fig. 4</emphasis>
.
<emphasis id="C1DB8B1AC6292033FEA1F957FEDCF93A" bold="true" box="[268,286,1730,1751]" pageId="10" pageNumber="61">a.</emphasis>
<taxonomicName id="34AF2C8BC6292033FE9AF957FE2AF93A" authorityName="Dumitrică" authorityYear="2024" box="[311,488,1730,1752]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="10" pageNumber="61" phylum="Radiozoa" rank="species" species="victoria" status="sp. nov.">
<emphasis id="C1DB8B1AC6292033FE9AF957FE2AF93A" box="[311,488,1730,1752]" italics="true" pageId="10" pageNumber="61">Gorispela victoria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6292033FE43F957FDE6F93A" box="[494,548,1730,1751]" pageId="10" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
, holotype, Rc4;
<emphasis id="C1DB8B1AC6292033FD6EF957FD14F93A" bold="true" box="[707,726,1730,1751]" pageId="10" pageNumber="61">b.</emphasis>
<taxonomicName id="34AF2C8BC6292033FD42F957FC6CF93A" authorityName="Dumitrică" authorityYear="2024" box="[751,942,1730,1752]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="10" pageNumber="61" phylum="Radiozoa" rank="species" species="mimetica" status="sp. nov.">
<emphasis id="C1DB8B1AC6292033FD42F957FC6CF93A" box="[751,942,1730,1752]" italics="true" pageId="10" pageNumber="61">Gorispela mimetica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6292033FC19F957FC28F93A" box="[948,1002,1730,1751]" pageId="10" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
, holotype, FÖ 87;
<emphasis id="C1DB8B1AC6292033FB32F957FB72F93A" bold="true" box="[1183,1200,1730,1751]" pageId="10" pageNumber="61">c.</emphasis>
<taxonomicName id="34AF2C8BC6292033FB64F957FABDF935" authorityName="Dumitrică" authorityYear="2024" box="[1225,1407,1730,1752]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="10" pageNumber="61" phylum="Radiozoa" rank="species" species="haeckeli" status="sp. nov.">
<emphasis id="C1DB8B1AC6292033FB64F957FABDF935" box="[1225,1407,1730,1752]" italics="true" pageId="10" pageNumber="61">Gorispela haeckeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6292033FF68F975FF3BF918" box="[197,249,1760,1781]" pageId="10" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
, holotype, Rc4;
<emphasis id="C1DB8B1AC6292033FE36F975FE01F918" bold="true" box="[411,451,1760,1781]" pageId="10" pageNumber="61">d -g.</emphasis>
<taxonomicName id="34AF2C8BC6292033FE71F94AFD7BF918" authorityName="Dumitrică" authorityYear="2024" box="[476,697,1759,1781]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="10" pageNumber="61" phylum="Radiozoa" rank="species" species="vicentiniana" status="sp. nov.">
<emphasis id="C1DB8B1AC6292033FE71F94AFD7BF918" box="[476,697,1759,1781]" italics="true" pageId="10" pageNumber="61">Gorispela vicentiniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6292033FD6CF975FD3AF918" box="[705,760,1760,1781]" pageId="10" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="C1DB8B1AC6292033FCAFF975FCCCF918" bold="true" box="[770,782,1760,1781]" pageId="10" pageNumber="61">g</emphasis>
.- Rc4,
<emphasis id="C1DB8B1AC6292033FCFAF975FCA8F918" bold="true" box="[855,874,1760,1781]" pageId="10" pageNumber="61">d.</emphasis>
Rc2, holotype,
<emphasis id="C1DB8B1AC6292033FBB5F975FBEBF918" bold="true" box="[1048,1065,1760,1781]" pageId="10" pageNumber="61">e.</emphasis>
apical horn of another specimen,
<emphasis id="C1DB8B1AC6292033FF68F968FF11F8FF" bold="true" box="[197,211,1789,1810]" pageId="10" pageNumber="61">f.</emphasis>
Rc1d;
<emphasis id="C1DB8B1AC6292033FE86F968FEFCF8FF" bold="true" box="[299,318,1789,1810]" pageId="10" pageNumber="61">h.</emphasis>
<taxonomicName id="34AF2C8BC6292033FEFBF968FDC1F8FE" authorityName="Dumitrică" authorityYear="2024" box="[342,515,1789,1811]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="10" pageNumber="61" phylum="Radiozoa" rank="species" species="bragini" status="sp. nov.">
<emphasis id="C1DB8B1AC6292033FEFBF968FDC1F8FE" box="[342,515,1789,1811]" italics="true" pageId="10" pageNumber="61">Gorispela bragini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6292033FDA4F96BFDFCF8FE" box="[521,574,1790,1811]" pageId="10" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
, holotype., basal view of fig. 4c;
<emphasis id="C1DB8B1AC6292033FCD3F968FC44F8FF" bold="true" box="[894,902,1789,1810]" pageId="10" pageNumber="61">f</emphasis>
,
<emphasis id="C1DB8B1AC6292033FC3FF968FC5CF8FF" bold="true" box="[914,926,1789,1810]" pageId="10" pageNumber="61">g</emphasis>
.- Rc1. Scale bar of 50 µm is for all figures.
</paragraph>
</caption>
<paragraph id="F3105708C6292033FF3AF8C1FDB1F881" blockId="10.[151,627,1843,1900]" box="[151,627,1876,1900]" pageId="10" pageNumber="61">
<emphasis id="C1DB8B1AC6292033FF3AF8C1FEADF881" bold="true" box="[151,367,1876,1900]" italics="true" pageId="10" pageNumber="61">Stratigraphic range</emphasis>
. Late Anisian (Illyrian).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="C0CDFAB8CD262CF000233643695A34CB">New Middle Triassic Bell-Shaped Nassellarian Radiolaria From Alpine And Carpathian Areas</mods:title>
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<paragraph id="F3105708C62B2031FE97F8DBFDA2F88B" blockId="8.[314,608,1870,1894]" box="[314,608,1870,1894]" pageId="8" pageNumber="59">
<heading id="A858E064C62B2031FE97F8DBFDA2F88B" box="[314,608,1870,1894]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="59" reason="5">
<emphasis id="C1DB8B1AC62B2031FE97F8DBFDA2F88B" bold="true" box="[314,608,1870,1894]" pageId="8" pageNumber="59">
<taxonomicName id="34AF2C8BC62B2031FE97F8DBFDDCF88B" authorityName="Dumitrică" authorityYear="2024" box="[314,542,1870,1894]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="8" pageNumber="59" phylum="Radiozoa" rank="species" species="hungarica" status="sp. nov.">
<emphasis id="C1DB8B1AC62B2031FE97F8DBFDDCF88B" bold="true" box="[314,542,1870,1894]" italics="true" pageId="8" pageNumber="59">Gorispela hungarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62B2031FD89F8DBFDA2F88B" box="[548,608,1870,1894]" pageId="8" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62B2031FE34F8FBFDC0F86B" box="[409,514,1902,1926]" pageId="8" pageNumber="59" type="description">
<paragraph id="F3105708C62B2031FE34F8FBFDC0F86B" blockId="8.[409,514,1902,1926]" box="[409,514,1902,1926]" pageId="8" pageNumber="59">
<figureCitation id="6B944B8DC62B2031FE34F8FBFE1DF86B" box="[409,479,1902,1926]" captionStart="Fig" captionStartId="8.[197,232,1750,1771]" captionTargetBox="[212,1390,203,1661]" captionTargetId="figure-171@8.[197,1410,178,1721]" captionTargetPageId="8" captionText="Fig. 3. a, a1. Gorispela cornubovis n. sp., holotype in lateral and basal views, respectively, Rc4; b. Gorispela fenestrata n. sp., holotype in lateral view, BV 85-70; c-e. Gorispela hungarica n. sp., FÖ 87, e. holotype. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582784" httpUri="https://zenodo.org/record/12582784/files/figure.png" pageId="8" pageNumber="59">Figs. 3</figureCitation>
c-e
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62B2030FF3AF824FEF6FDEC" lastPageId="9" lastPageNumber="60" pageId="8" pageNumber="59" type="diagnosis">
<paragraph id="F3105708C62B2030FF3AF824FEBDFE53" blockId="8.[151,771,1969,2059]" lastBlockId="9.[151,772,158,1440]" lastPageId="9" lastPageNumber="60" pageId="8" pageNumber="59">
<emphasis id="C1DB8B1AC62B2031FF3AF824FEC7F824" bold="true" box="[151,261,1969,1993]" italics="true" pageId="8" pageNumber="59">Diagnosis</emphasis>
. Shell bell-shaped, conical bearing two or three circumferential ridges. Cephalis with robust and hollow conical apical and ventral horns. Its wall imperforated or with sparse rounded small pores of various sizes and irregular dispositions. Apical horn in axial position, its distal end with a few short spinules or irregular surface. Ventral horn in horizontal or subhorizontal position, its distal end three-bladed, blades thick or practically absent, and with irregular surface. Initial spicule with distal ends of apical and ventral rays attached to the inner surface of horns. Thorax truncate conical, bearing two or three circumferential ridges, its wall with irregularly distributed pores of various forms, from circular to oval or triangular. Size of pores increases from the proximal to the distal part. Distal end represented by an imperforate band. Collar boundary not marked by a change in shape but considered to be above the level of the first circumferential ridge.
</paragraph>
<paragraph id="F3105708C62A2030FF3AFE5DFEF6FDEC" blockId="9.[151,772,158,1440]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FF3AFE5DFE91FE0D" bold="true" box="[151,339,456,480]" italics="true" pageId="9" pageNumber="60">Studied material</emphasis>
. Three illustrated specimens from the sample FÖ 87.
</paragraph>
</subSubSection>
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<paragraph id="F3105708C62A2030FF3AFD9FFDECFDCF" blockId="9.[151,772,158,1440]" box="[151,558,522,546]" pageId="9" pageNumber="60">
<materialsCitation id="43C75D55C62A2030FF3AFD9FFDECFDCF" box="[151,558,522,546]" collectionCode="MGL" pageId="9" pageNumber="60" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC62A2030FF3AFD9FFF38FDCF" bold="true" box="[151,250,522,546]" italics="true" pageId="9" pageNumber="60">
<typeStatus id="2C14E9AAC62A2030FF3AFD9FFF38FDCF" box="[151,250,522,546]" pageId="9" pageNumber="60" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC62A2030FEA5FD9FFE96FDCF" box="[264,340,522,546]" captionStart="Fig" captionStartId="8.[197,232,1750,1771]" captionTargetBox="[212,1390,203,1661]" captionTargetId="figure-171@8.[197,1410,178,1721]" captionTargetPageId="8" captionText="Fig. 3. a, a1. Gorispela cornubovis n. sp., holotype in lateral and basal views, respectively, Rc4; b. Gorispela fenestrata n. sp., holotype in lateral view, BV 85-70; c-e. Gorispela hungarica n. sp., FÖ 87, e. holotype. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582784" httpUri="https://zenodo.org/record/12582784/files/figure.png" pageId="9" pageNumber="60">Fig. 3e</figureCitation>
, coll.
<collectionCode id="95BECFCDC62A2030FE39FD9FFE11FDCF" box="[404,467,522,546]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="9" pageNumber="60">MGL</collectionCode>
.110274.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2030FF3AFDBEFEFEFC81" pageId="9" pageNumber="60" type="description">
<paragraph id="F3105708C62A2030FF3AFDBEFE1FFD4A" blockId="9.[151,772,158,1440]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FF3AFDBEFED8FDAE" bold="true" box="[151,282,555,579]" italics="true" pageId="9" pageNumber="60">Dimensions</emphasis>
. Length of shell with apical horn 155-160 µm, without horn 100 µm, of apical horn 50-60 µm, of ventral horn 40-45 µm; diameter of horns at base 20-25 µm, of base of thorax 108 µm.
</paragraph>
<paragraph id="F3105708C62A2030FF3AFD25FEFEFC81" blockId="9.[151,772,158,1440]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FF3AFD25FECEFD25" bold="true" box="[151,268,688,712]" italics="true" pageId="9" pageNumber="60">Variability</emphasis>
. The three illustrated specimens, the only ones found in the Felsőörs sample, prove that the variability of the species is visible especially in the number of circumferential rings and, depending on their number, the distance between them because the size of shell remains rather constant.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2030FF3AFCE3FDB4FC62" box="[151,630,886,911]" pageId="9" pageNumber="60" type="etymology">
<paragraph id="F3105708C62A2030FF3AFCE3FDB4FC62" blockId="9.[151,772,158,1440]" box="[151,630,886,911]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FF3AFCE3FECEFC63" bold="true" box="[151,268,886,910]" italics="true" pageId="9" pageNumber="60">Etymology</emphasis>
. From its occurrence in
<collectingCountry id="8BB81798C62A2030FDBCFCE2FDB0FC62" box="[529,626,887,911]" name="Hungary" pageId="9" pageNumber="60">Hungary</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62A2030FF3AFC02FD36FA72" pageId="9" pageNumber="60" type="discussion">
<paragraph id="F3105708C62A2030FF3AFC02FEC7FA93" blockId="9.[151,772,158,1440]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FF3AFC02FF3BFC42" bold="true" box="[151,249,919,943]" italics="true" pageId="9" pageNumber="60">Remarks</emphasis>
.
<taxonomicName id="34AF2C8BC62A2030FEB1FC02FDD3FC42" authorityName="Dumitrică" authorityYear="2024" box="[284,529,919,943]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="hungarica" status="sp. nov.">
<emphasis id="C1DB8B1AC62A2030FEB1FC02FDD3FC42" box="[284,529,919,943]" italics="true" pageId="9" pageNumber="60">Gorispela hungarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62A2030FD80FC0DFDBEFC42" box="[557,636,920,943]" pageId="9" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="34AF2C8BC62A2030FF3AFC2DFEB9FC3D" authorityName="Dumitrică" authorityYear="2024" box="[151,379,952,976]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="fenestrata" status="sp. nov.">
<emphasis id="C1DB8B1AC62A2030FF3AFC2DFEB9FC3D" box="[151,379,952,976]" italics="true" pageId="9" pageNumber="60">Gorispela fenestrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62A2030FE25FC2CFE0AFC3D" box="[392,456,953,976]" pageId="9" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
by having a rather similar shape, but differs by having the apical horn in axial position, the ventral horn in horizontal position, and by not having two distal bands. It differs also by the position of the ventral ray of the initial spicule, that has not a central position in the ventral horn but it is connected to the lower side of its inner wall. It is interesting to mention that the same types of spinules present on the distal part of the apical horn of the
<specimenCount id="E5A99C81C62A2030FDB1FB54FD0AFB34" box="[540,712,1217,1241]" count="3" pageId="9" pageNumber="60" type="generic">three specimens</specimenCount>
have been remarked in the same position on the
<typeStatus id="2C14E9AAC62A2030FDD6FB77FD21FB17" box="[635,739,1250,1274]" pageId="9" pageNumber="60" type="holotype">holotypes</typeStatus>
of the species
<taxonomicName id="34AF2C8BC62A2030FEB3FA96FE35FAF6" authorityName="Dumitrică" authorityYear="2024" box="[286,503,1283,1307]" class="Polycystina" family="Oertlispongidae" genus="Gorispela" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="mimetica" status="sp. nov.">
<emphasis id="C1DB8B1AC62A2030FEB3FA96FE35FAF6" box="[286,503,1283,1307]" italics="true" pageId="9" pageNumber="60">Gorispela mimetica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62A2030FDAEFA91FD80FAF6" box="[515,578,1284,1307]" pageId="9" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
(see below) and
<taxonomicName id="34AF2C8BC62A2030FF3AFAB1FE2CFAD1" authorityName="Dumitrică" authorityYear="2024" box="[151,494,1316,1340]" class="Polycystina" family="Oertlispongidae" genus="Silicotinntinabulum" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="9" pageNumber="60" phylum="Radiozoa" rank="species" species="formosum" status="sp. nov.">
<emphasis id="C1DB8B1AC62A2030FF3AFAB1FE2CFAD1" box="[151,494,1316,1340]" italics="true" pageId="9" pageNumber="60">Silicotinntinabulum formosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62A2030FDA5FAB0FD94FAD1" box="[520,598,1317,1340]" pageId="9" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
, two species occurring in the same sample (case of horizontal gene transfer?).
</paragraph>
<paragraph id="F3105708C62A2030FF3AFA1DFD36FA72" blockId="9.[151,772,158,1440]" box="[151,756,1415,1440]" pageId="9" pageNumber="60">
<emphasis id="C1DB8B1AC62A2030FF3AFA1DFEADFA4D" bold="true" box="[151,367,1416,1440]" italics="true" pageId="9" pageNumber="60">Stratigraphic range</emphasis>
. Late Anisian (Illyrian) of
<collectingCountry id="8BB81798C62A2030FD3DFA1DFD2FFA4D" box="[656,749,1416,1440]" name="Hungary" pageId="9" pageNumber="60">Hungary</collectingCountry>
.
</paragraph>
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<taxonomicName id="34AF2C8BC62C2036FE80F872FDE9F812" authorityName="Dumitrică" authorityYear="2024" box="[301,555,2023,2047]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="15" pageNumber="66" phylum="Radiozoa" rank="species" species="deweveri" status="sp. nov.">
<emphasis id="C1DB8B1AC62C2036FE80F872FDE9F812" bold="true" box="[301,555,2023,2047]" italics="true" pageId="15" pageNumber="66">Humerocyrtis deweveri</emphasis>
</taxonomicName>
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<figureCitation id="6B944B8DC62C2036FE05F79DFE30F7CD" box="[424,498,2056,2080]" captionStart="Fig" captionStartId="15.[197,232,1741,1762]" captionTargetBox="[213,1405,175,1620]" captionTargetId="figure-137@15.[197,1418,160,1713]" captionTargetPageId="15" captionText="Fig. 6. a, b. Gorispela (?) triangulocephalis n. sp., FÖ 87: a. specimen showing internal structure, b. holotype, D dorsal ray. c. Humerocyrtis superba n. sp., holotype, BV 85-70; 6d, 6c. Humerocyrtis jekeli n. sp., Rc4, 6c. holotype, d. - paratype; e. Humerocyrtis deweveri n. sp., holotype, BW 85-70; f. Humerocyrtis problematica n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582788" httpUri="https://zenodo.org/record/12582788/files/figure.png" pageId="15" pageNumber="66">Fig. 6e</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62C2029FC9DF8F6FF1FFD37" lastPageId="16" lastPageNumber="67" pageId="15" pageNumber="66" type="diagnosis">
<paragraph id="F3105708C62C2029FC9DF8F6FDA2FD7A" blockId="15.[816,1436,1891,2080]" lastBlockId="16.[151,772,176,1325]" lastPageId="16" lastPageNumber="67" pageId="15" pageNumber="66">
<emphasis id="C1DB8B1AC62C2036FC9DF8F6FC5CF896" bold="true" box="[816,926,1891,1915]" italics="true" pageId="15" pageNumber="66">Diagnosis</emphasis>
. Shell relatively large, bell-shaped with rounded shoulder. Proximal part, consisting of cephalis and upper part of thorax, conical with concave lateral sides. Cephalis conical with two three-bladed horns: an apical horn in axial position and a smaller ventral horn, both pointed and having thin blades. Cephalic wall perforated by small circular or subcircular and irregularly arranged pores. Thorax of this part of shell wide conical. It has much larger and irregularly disposed pores. Shoulder rounded and having small pores of variable sizes and irregular arrangement, especially on the proximal side. On distal side of thorax the shoulder has two circumferential rows of rounded pores of various sizes. Below them the thorax is constricted and distally flared, bearing three circumferential rows of large pores arranged into 3 circumferential intercalary rows of
<specimenCount id="E5A99C81C6332029FD76FE4EFF11FDF9" count="2" pageId="16" pageNumber="67" type="generic" typeStatus="types">two types</specimenCount>
: the first two rows with very large rounded rectangular or oval pores with long axes longitudinally positioned, the last row with slightly smaller pores arranged with transversally directed long axes, each row containing 6-7 pores on half the perimeter.
</paragraph>
<paragraph id="F3105708C6332029FF3AFD34FF1FFD37" blockId="16.[151,772,176,1325]" pageId="16" pageNumber="67">
<emphasis id="C1DB8B1AC6332029FF3AFD34FE93FD54" bold="true" box="[151,337,673,697]" italics="true" pageId="16" pageNumber="67">Studied material</emphasis>
. A single specimen in the sample BV 85-70.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6332029FF3AFD76FD35FD16" box="[151,759,739,763]" pageId="16" pageNumber="67" type="materials_examined">
<paragraph id="F3105708C6332029FF3AFD76FD35FD16" blockId="16.[151,772,176,1325]" box="[151,759,739,763]" pageId="16" pageNumber="67">
<materialsCitation id="43C75D55C6332029FF3AFD76FD35FD16" box="[151,759,739,763]" collectionCode="MGL" pageId="16" pageNumber="67" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC6332029FF3AFD76FF38FD16" bold="true" box="[151,250,739,763]" italics="true" pageId="16" pageNumber="67">
<typeStatus id="2C14E9AAC6332029FF3AFD76FF38FD16" box="[151,250,739,763]" pageId="16" pageNumber="67" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC6332029FEA5FD76FE96FD16" box="[264,340,739,763]" captionStart="Fig" captionStartId="15.[197,232,1741,1762]" captionTargetBox="[213,1405,175,1620]" captionTargetId="figure-137@15.[197,1418,160,1713]" captionTargetPageId="15" captionText="Fig. 6. a, b. Gorispela (?) triangulocephalis n. sp., FÖ 87: a. specimen showing internal structure, b. holotype, D dorsal ray. c. Humerocyrtis superba n. sp., holotype, BV 85-70; 6d, 6c. Humerocyrtis jekeli n. sp., Rc4, 6c. holotype, d. - paratype; e. Humerocyrtis deweveri n. sp., holotype, BW 85-70; f. Humerocyrtis problematica n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582788" httpUri="https://zenodo.org/record/12582788/files/figure.png" pageId="16" pageNumber="67">Fig. 6e</figureCitation>
, sample BV 85-70, coll.
<collectionCode id="95BECFCDC6332029FDF3FD76FD5CFD16" box="[606,670,739,763]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="16" pageNumber="67">MGL</collectionCode>
.110285.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6332029FF3AFC91FE6FFC4C" pageId="16" pageNumber="67" type="description">
<paragraph id="F3105708C6332029FF3AFC91FE6FFC4C" blockId="16.[151,772,176,1325]" pageId="16" pageNumber="67">
<emphasis id="C1DB8B1AC6332029FF3AFC91FED8FCF1" bold="true" box="[151,282,772,796]" italics="true" pageId="16" pageNumber="67">Dimensions</emphasis>
. Length of shell without apical horn 114 µm, with apical horn 160 µm, length of shell up to the circumferential ring 50 µm, of distal part 65 µm, diameter of cephalis 40 µm, of shoulder or circumferential ring 74 µm, of distal end 120 µm.
</paragraph>
</subSubSection>
<caption id="A7D00780C6332029FCD6FC08FA8FFC5E" ID-DOI="http://doi.org/10.5281/zenodo.12582790" ID-Zenodo-Dep="12582790" box="[891,1357,925,947]" httpUri="https://zenodo.org/record/12582790/files/figure.png" pageId="16" pageNumber="67" startId="16.[891,926,925,946]" targetBox="[863,1380,302,890]" targetPageId="16" targetType="figure">
<paragraph id="F3105708C6332029FCD6FC08FA8FFC5E" blockId="16.[891,1357,925,947]" box="[891,1357,925,947]" pageId="16" pageNumber="67">
<emphasis id="C1DB8B1AC6332029FCD6FC08FC79FC5F" bold="true" box="[891,955,925,946]" pageId="16" pageNumber="67">Fig. 7.</emphasis>
-
<taxonomicName id="34AF2C8BC6332029FC62FC0BFB91FC5E" authorityName="Dumitrică" authorityYear="2024" box="[975,1107,926,947]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="16" pageNumber="67" phylum="Radiozoa" rank="genus" status="n. sp.">
<emphasis id="C1DB8B1AC6332029FC62FC0BFB91FC5E" box="[975,1107,926,947]" italics="true" pageId="16" pageNumber="67">Humerocyrtis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6332029FBF4FC0BFB4CFC5E" box="[1113,1166,926,947]" pageId="16" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
, H shoulder, Rc4.
</paragraph>
</caption>
<subSubSection id="BBB50483C6332029FF3AFC3FFE57FBEE" pageId="16" pageNumber="67" type="etymology">
<paragraph id="F3105708C6332029FF3AFC3FFE57FBEE" blockId="16.[151,772,176,1325]" pageId="16" pageNumber="67">
<emphasis id="C1DB8B1AC6332029FF3AFC3FFECEFC2F" bold="true" box="[151,268,938,962]" italics="true" pageId="16" pageNumber="67">Etymology</emphasis>
. The species is dedicated to my friend Prof. Patrick De Wever for his contribution to the knowledge of Triassic radiolarians.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C6332029FF3AFB98FF3FFAC0" pageId="16" pageNumber="67" type="discussion">
<paragraph id="F3105708C6332029FF3AFB98FDD7FB27" blockId="16.[151,772,176,1325]" pageId="16" pageNumber="67">
<emphasis id="C1DB8B1AC6332029FF3AFB98FF3BFBC8" bold="true" box="[151,249,1037,1061]" italics="true" pageId="16" pageNumber="67">Remarks</emphasis>
. This new species resembles very much the species
<taxonomicName id="34AF2C8BC6332029FF42FBBAFE21FBAB" authorityName="Dumitrică" authorityYear="2024" box="[239,483,1070,1094]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="16" pageNumber="67" phylum="Radiozoa" rank="species" species="superba" status="sp. nov.">
<emphasis id="C1DB8B1AC6332029FF42FBBAFE21FBAB" box="[239,483,1070,1094]" italics="true" pageId="16" pageNumber="67">Humerocyrtis superba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C6332029FE40FBBBFDE9FBA8" box="[493,555,1070,1093]" pageId="16" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
by having a rather similar morphology from which it differs by being smaller, by having smaller horns, shoulder with rounded outline and distal part of thorax with only three circumferential rows of large pores.
</paragraph>
<paragraph id="F3105708C6332029FF3AFB46FF3FFAC0" blockId="16.[151,772,176,1325]" pageId="16" pageNumber="67">
<emphasis id="C1DB8B1AC6332029FF3AFB46FE58FB06" bold="true" box="[151,410,1235,1259]" italics="true" pageId="16" pageNumber="67">Stratigraphic range</emphasis>
. Livinallongo Formation, Marmolada Massif, North
<collectingCountry id="8BB81798C6332029FE11FB61FE2DFAE1" box="[444,495,1268,1292]" name="Italy" pageId="16" pageNumber="67">Italy</collectingCountry>
, sample BV85-70, lower Ladinian.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,115 @@
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<treatment id="7B06E61EC62D2037FC67FC91FAAAF809" LSID="urn:lsid:plazi:treatment:7B06E61EC62D2037FC67FC91FAAAF809" httpUri="http://treatment.plazi.org/id/7B06E61EC62D2037FC67FC91FAAAF809" lastPageNumber="65" pageId="14" pageNumber="65">
<subSubSection id="BBB50483C62D2037FC67FC91FAC0FCF1" box="[970,1282,772,796]" pageId="14" pageNumber="65" type="nomenclature">
<paragraph id="F3105708C62D2037FC67FC91FAC0FCF1" blockId="14.[970,1282,772,796]" box="[970,1282,772,796]" pageId="14" pageNumber="65">
<heading id="A858E064C62D2037FC67FC91FAC0FCF1" box="[970,1282,772,796]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="65" reason="5">
<emphasis id="C1DB8B1AC62D2037FC67FC91FAC0FCF1" bold="true" box="[970,1282,772,796]" pageId="14" pageNumber="65">
<taxonomicName id="34AF2C8BC62D2037FC67FC91FB7DFCF1" authorityName="Dumitrică" authorityYear="2024" box="[970,1215,772,796]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="14" pageNumber="65" phylum="Radiozoa" rank="species" species="superba" status="sp. nov.">
<emphasis id="C1DB8B1AC62D2037FC67FC91FB7DFCF1" bold="true" box="[970,1215,772,796]" italics="true" pageId="14" pageNumber="65">Humerocyrtis superba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C62D2037FB6BFC91FAC0FCF1" box="[1222,1282,772,796]" pageId="14" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62D2037FBECFCB0FB4EFCD0" box="[1089,1164,805,829]" pageId="14" pageNumber="65" type="description">
<paragraph id="F3105708C62D2037FBECFCB0FB4EFCD0" blockId="14.[1089,1164,805,829]" box="[1089,1164,805,829]" pageId="14" pageNumber="65">
<figureCitation id="6B944B8DC62D2037FBECFCB0FB4EFCD0" box="[1089,1164,805,829]" captionStart="Fig" captionStartId="15.[197,232,1741,1762]" captionTargetBox="[213,1405,175,1620]" captionTargetId="figure-137@15.[197,1418,160,1713]" captionTargetPageId="15" captionText="Fig. 6. a, b. Gorispela (?) triangulocephalis n. sp., FÖ 87: a. specimen showing internal structure, b. holotype, D dorsal ray. c. Humerocyrtis superba n. sp., holotype, BV 85-70; 6d, 6c. Humerocyrtis jekeli n. sp., Rc4, 6c. holotype, d. - paratype; e. Humerocyrtis deweveri n. sp., holotype, BW 85-70; f. Humerocyrtis problematica n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582788" httpUri="https://zenodo.org/record/12582788/files/figure.png" pageId="14" pageNumber="65">Fig. 6c</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62D2037FC9DFCF2FA5AF974" pageId="14" pageNumber="65" type="diagnosis">
<paragraph id="F3105708C62D2037FC9DFCF2FC07F995" blockId="14.[816,1437,871,2021]" pageId="14" pageNumber="65">
<emphasis id="C1DB8B1AC62D2037FC9DFCF2FC5CFC92" bold="true" box="[816,926,871,895]" italics="true" pageId="14" pageNumber="65">Diagnosis</emphasis>
. Shell large, bell-shaped with the body divided into two parts by a median circumferential ridge corresponding to the maximum diameter of the shoulder: a proximal part and a distal part. Proximal part conical with concave lateral sides consisting of cephalis and the proximal part of the thorax. Cephalis hemisphaerical bearing two three-bladed horns: an apical horn and a ventral horn, both pointed and having thin but wide blades. Apical horn dorsally tilted. Cephalic wall perforated by very small circular or subcircular and irregularly arranged pores. Thorax of this part of shell wide conical with concave outline. It has much larger and irregularly arranged pores. Shoulder sharp and angular. Distal part of thorax much larger and longer than the proximal part, its outline concave and distal end flared. Pores of this part arranged in 6 circumferential intercalary rows of
<specimenCount id="E5A99C81C62D2037FC3CFAEDFBCBFA7C" box="[913,1033,1400,1425]" count="3" pageId="14" pageNumber="65" type="generic" typeStatus="types">three types</specimenCount>
of shapes and sizes: first two rows consisting of smaller circular, oval or elliptical pores, following two rows consisting of very large rounded rectangular or oval pores with long axes longitudinally positioned, and last two rows with elliptical pores arranged with long axes transversally directed. Each row of such thoracic pores consisting of 10-11 pores on half the perimeter.
</paragraph>
<paragraph id="F3105708C62D2037FC9DF917FA5AF974" blockId="14.[816,1437,871,2021]" box="[816,1432,1665,1690]" pageId="14" pageNumber="65">
<emphasis id="C1DB8B1AC62D2037FC9DF917FC24F977" bold="true" box="[816,998,1666,1690]" italics="true" pageId="14" pageNumber="65">Studied material</emphasis>
.
<specimenCount id="E5A99C81C62D2037FC5EF914FB49F974" box="[1011,1163,1665,1689]" count="1" pageId="14" pageNumber="65" type="generic">One specimen</specimenCount>
in the sample BV 85-70.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62D2037FC9DF936FA53F957" box="[816,1425,1698,1723]" pageId="14" pageNumber="65" type="materials_examined">
<paragraph id="F3105708C62D2037FC9DF936FA53F957" blockId="14.[816,1437,871,2021]" box="[816,1425,1698,1723]" pageId="14" pageNumber="65">
<materialsCitation id="43C75D55C62D2037FC9DF936FA53F957" box="[816,1425,1698,1723]" collectionCode="MGL" pageId="14" pageNumber="65" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC62D2037FC9DF936FC51F956" bold="true" box="[816,915,1699,1723]" italics="true" pageId="14" pageNumber="65">
<typeStatus id="2C14E9AAC62D2037FC9DF936FC51F956" box="[816,915,1699,1723]" pageId="14" pageNumber="65" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC62D2037FC0CF937FC2FF957" box="[929,1005,1698,1722]" captionStart="Fig" captionStartId="15.[197,232,1741,1762]" captionTargetBox="[213,1405,175,1620]" captionTargetId="figure-137@15.[197,1418,160,1713]" captionTargetPageId="15" captionText="Fig. 6. a, b. Gorispela (?) triangulocephalis n. sp., FÖ 87: a. specimen showing internal structure, b. holotype, D dorsal ray. c. Humerocyrtis superba n. sp., holotype, BV 85-70; 6d, 6c. Humerocyrtis jekeli n. sp., Rc4, 6c. holotype, d. - paratype; e. Humerocyrtis deweveri n. sp., holotype, BW 85-70; f. Humerocyrtis problematica n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582788" httpUri="https://zenodo.org/record/12582788/files/figure.png" pageId="14" pageNumber="65">Fig. 6c</figureCitation>
, sample BV 85-70, coll.
<collectionCode id="95BECFCDC62D2037FB5AF937FAF4F957" box="[1271,1334,1698,1722]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="14" pageNumber="65">MGL</collectionCode>
.110284.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62D2037FC9DF951FBB7F88C" pageId="14" pageNumber="65" type="description">
<paragraph id="F3105708C62D2037FC9DF951FBB7F88C" blockId="14.[816,1437,871,2021]" pageId="14" pageNumber="65">
<emphasis id="C1DB8B1AC62D2037FC9DF951FC71F931" bold="true" box="[816,947,1732,1756]" italics="true" pageId="14" pageNumber="65">Dimensions</emphasis>
. Length of shell without apical horn 155 µm, with apical horn 210 µm, length of shell up to the circumferential ring 70 µm, of distal part 85 µm, diameter of cephalis 50 µm, of shoulder or circumferential ring 110 µm, of distal end 160 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62D2037FC9DF8FCFABFF86C" box="[816,1405,1897,1921]" pageId="14" pageNumber="65" type="etymology">
<paragraph id="F3105708C62D2037FC9DF8FCFABFF86C" blockId="14.[816,1437,871,2021]" box="[816,1405,1897,1921]" pageId="14" pageNumber="65">
<emphasis id="C1DB8B1AC62D2037FC9DF8FCFC67F86C" bold="true" box="[816,933,1897,1921]" italics="true" pageId="14" pageNumber="65">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC62D2037FBF7F8FCFB78F86C" box="[1114,1210,1897,1921]" italics="true" pageId="14" pageNumber="65">superbus</emphasis>
, -
<emphasis id="C1DB8B1AC62D2037FB7DF8FFFB1FF86C" box="[1232,1245,1898,1921]" italics="true" pageId="14" pageNumber="65">a</emphasis>
, -
<emphasis id="C1DB8B1AC62D2037FB5EF8FFFAD6F86C" box="[1267,1300,1898,1921]" italics="true" pageId="14" pageNumber="65">um</emphasis>
superb.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C62D2037FC9DF81EFAAAF809" pageId="14" pageNumber="65" type="discussion">
<paragraph id="F3105708C62D2037FC9DF81EFAAAF809" blockId="14.[816,1437,871,2021]" pageId="14" pageNumber="65">
<emphasis id="C1DB8B1AC62D2037FC9DF81EFC50F84E" bold="true" box="[816,914,1931,1955]" italics="true" pageId="14" pageNumber="65">Remarks</emphasis>
. This species differs from other species of this genus by its great size and, especially, the different
<typeStatus id="2C14E9AAC62D2037FAC9F839FA5EF829" box="[1380,1436,1964,1988]" pageId="14" pageNumber="65">types</typeStatus>
and arrangement of pores on the distal part of thorax.
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<heading id="A858E064C62F2035FC15FAA2FAD7FAA2" box="[952,1301,1335,1359]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="63" reason="5">
<emphasis id="C1DB8B1AC62F2035FC15FAA2FAD7FAA2" bold="true" box="[952,1301,1335,1359]" pageId="12" pageNumber="63">
<taxonomicName id="34AF2C8BC62F2035FC15FAA2FB10FAA2" authorityName="Dumitrică" authorityYear="2024" box="[952,1234,1335,1359]" class="Polycystina" family="Oertlispongidae" genus="Pozsvartia" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="12" pageNumber="63" phylum="Radiozoa" rank="species" species="multicingulata" status="sp. nov.">
<emphasis id="C1DB8B1AC62F2035FC15FAA2FB10FAA2" bold="true" box="[952,1234,1335,1359]" italics="true" pageId="12" pageNumber="63">Pozsvartia multicingulata</emphasis>
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<taxonomicNameLabel id="DAE83661C62F2035FB74FAA2FAD7FAA2" box="[1241,1301,1335,1359]" pageId="12" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<figureCitation id="6B944B8DC62F2035FB9EFACDFBBBFA9D" box="[1075,1145,1368,1392]" captionStart="Fig" captionStartId="13.[197,232,1409,1430]" captionTargetBox="[213,1380,173,1369]" captionTargetId="figure-420@13.[197,1411,160,1381]" captionTargetPageId="13" captionText="Fig. 5. a, b. Gorispela tuba n. sp., a. Rc4, b. holotype, FÖ 87; c. Gorispela bragini n. sp., holotype, Rc4 (see also fig. 3h); d-f. Pozsvartia multicingulata n. sp., d. holotype, FÖ 87, e. CRH, f. CR25, D dorsal spine. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582786" httpUri="https://zenodo.org/record/12582786/files/figure.png" pageId="12" pageNumber="63">Figs. 5</figureCitation>
d-f
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<paragraph id="F3105708C630202AFE8CFBA8FDBBFBB8" blockId="19.[289,633,1085,1109]" box="[289,633,1085,1109]" pageId="19" pageNumber="70">
<heading id="A858E064C630202AFE8CFBA8FDBBFBB8" box="[289,633,1085,1109]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="70" reason="5">
<emphasis id="C1DB8B1AC630202AFE8CFBA8FDBBFBB8" bold="true" box="[289,633,1085,1109]" pageId="19" pageNumber="70">
<taxonomicName id="34AF2C8BC630202AFE8CFBA8FDF5FBB8" authorityName="Dumitrică" authorityYear="2024" box="[289,567,1085,1109]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="19" pageNumber="70" phylum="Radiozoa" rank="species" species="avirostrum" status="sp. nov.">
<emphasis id="C1DB8B1AC630202AFE8CFBA8FDF5FBB8" bold="true" box="[289,567,1085,1109]" italics="true" pageId="19" pageNumber="70">Humerocyrtis avirostrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C630202AFD90FBA8FDBBFBB8" box="[573,633,1085,1109]" pageId="19" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<paragraph id="F3105708C630202AFE05FBC8FE30FB98" blockId="19.[424,498,1117,1141]" box="[424,498,1117,1141]" pageId="19" pageNumber="70">
<figureCitation id="6B944B8DC630202AFE05FBC8FE30FB98" box="[424,498,1117,1141]" captionStart="Fig" captionStartId="20.[197,232,1730,1751]" captionTargetBox="[206,1394,188,1577]" captionTargetId="figure-156@20.[197,1410,178,1721]" captionTargetPageId="20" captionText="Fig. 9. a, b, f. Humerocyrtis gracilis n. sp.: a. paratype, Rc4; b. holotype, BV 85-70, f. Rc4. c. Humerocyrtis avirostrum n. sp., Rc4; d, d1, e. Humerocyrtis brevithorax n. sp., CRH: d, d1. same specimen in lateral and basal view, respectively, showing the initial spicule without D ray, e. holotype. Scale bar of 50 µm is for all figures, except fig. 7c for which the scale bar is of 75 µm." figureDoi="http://doi.org/10.5281/zenodo.12582794" httpUri="https://zenodo.org/record/12582794/files/figure.png" pageId="19" pageNumber="70">Fig. 9c</figureCitation>
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<paragraph id="F3105708C630202AFF3AFB35FEFBF8E6" blockId="19.[151,772,1184,2068]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AFB35FEC7FB55" bold="true" box="[151,261,1184,1208]" italics="true" pageId="19" pageNumber="70">Diagnosis</emphasis>
. Shell large bell-shaped, with well-developed and pointed three-bladed apical and ventral spines. Cephalis short and globular with a few very small pores. Collar boundary marked by a depression on the dorsal side and under the ventral spine that correspond to the position of MB. Proximal part of thorax tronconical and short with elliptical pores of various sizes and irregular arrangement. Shoulder rounded or slightly acute but without a circumferential ring. It corresponds to the boundary between the first two circumferential rows of pores. Distal part of thorax with concave outline and with the minimum diameter under the shoulder. Distal part of thorax conical with 4 or 5 circumferential rows of larger pores of oval, elliptical or rounded quadrangular shape. Distal end of thorax with a circular band a little broader and thicker than the bars between the circumferential rows of thoracic pores. Under this band there is a narrowing row of rounded quadrangular pores with thin intraporal bars.
</paragraph>
<paragraph id="F3105708C630202AFF3AF880FD5AF8C1" blockId="19.[151,772,1184,2068]" box="[151,664,1812,1837]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AF880FE8FF8C0" bold="true" box="[151,333,1813,1837]" italics="true" pageId="19" pageNumber="70">Studied material</emphasis>
.
<specimenCount id="E5A99C81C630202AFEF7F881FE30F8C1" box="[346,498,1812,1836]" count="1" pageId="19" pageNumber="70" type="generic">One specimen</specimenCount>
in sample Rc4.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFF3AF8A3FDB9F8A0" box="[151,635,1845,1870]" pageId="19" pageNumber="70" type="materials_examined">
<paragraph id="F3105708C630202AFF3AF8A3FDB9F8A0" blockId="19.[151,772,1184,2068]" box="[151,635,1845,1870]" pageId="19" pageNumber="70">
<materialsCitation id="43C75D55C630202AFF3AF8A3FDB9F8A0" box="[151,635,1845,1870]" collectionCode="MGL" pageId="19" pageNumber="70" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC630202AFF3AF8A3FF38F8A3" bold="true" box="[151,250,1846,1870]" italics="true" pageId="19" pageNumber="70">
<typeStatus id="2C14E9AAC630202AFF3AF8A3FF38F8A3" box="[151,250,1846,1870]" pageId="19" pageNumber="70" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="E5A99C81C630202AFEA5F8A0FE61F8A0" box="[264,419,1845,1869]" count="1" pageId="19" pageNumber="70" type="generic">One specimen</specimenCount>
, coll.
<collectionCode id="95BECFCDC630202AFE4FF8A0FDE0F8A0" box="[482,546,1845,1869]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="19" pageNumber="70">MGL</collectionCode>
.110293.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFF3AF8C2FDDAF83E" pageId="19" pageNumber="70" type="description">
<paragraph id="F3105708C630202AFF3AF8C2FDDAF83E" blockId="19.[151,772,1184,2068]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AF8C2FED8F882" bold="true" box="[151,282,1879,1903]" italics="true" pageId="19" pageNumber="70">Dimensions</emphasis>
. Length of shell with apical horn and post thoracic additional shell 270 µm, without post thoracic shell 255 µm, diameter of cephalis 55 µm, of shoulder 83 µm, of distal part of thorax 108 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFF3AF84EFBDAFF5B" pageId="19" pageNumber="70" type="etymology">
<paragraph id="F3105708C630202AFF3AF84EFBDAFF5B" blockId="19.[151,772,1184,2068]" lastBlockId="19.[816,1437,158,479]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AF84EFED1F81E" box="[151,275,2011,2035]" italics="true" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AF84EFECEF81E" bold="true" box="[151,268,2011,2035]" italics="true" pageId="19" pageNumber="70">Etymology</emphasis>
.
</emphasis>
From the Latin
<emphasis id="C1DB8B1AC630202AFE75F849FDDFF81F" box="[472,541,2012,2035]" italics="true" pageId="19" pageNumber="70">avis </emphasis>
bird and
<emphasis id="C1DB8B1AC630202AFD38F849FCC1F81F" box="[661,771,2012,2035]" italics="true" pageId="19" pageNumber="70">rostrum </emphasis>
beak, due to the resemblance between the ventral spine and the beak of birds.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFC9DFF2AFB3BFE32" pageId="19" pageNumber="70" type="discussion">
<paragraph id="F3105708C630202AFC9DFF2AFAF9FE70" blockId="19.[816,1437,158,479]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFC9DFF2AFC50FF3A" bold="true" box="[816,914,191,215]" italics="true" pageId="19" pageNumber="70">Remarks</emphasis>
.
<taxonomicName id="34AF2C8BC630202AFC01FF55FB04FF3A" authorityName="Dumitrică" authorityYear="2024" box="[940,1222,192,215]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="19" pageNumber="70" phylum="Radiozoa" rank="species" species="avirostrum" status="sp. nov.">
<emphasis id="C1DB8B1AC630202AFC01FF55FB04FF3A" box="[940,1222,192,215]" italics="true" pageId="19" pageNumber="70">Humerocyrtis avirostrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C630202AFB74FF55FADDFF3A" box="[1241,1311,192,215]" pageId="19" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
resembles generally
<taxonomicName id="34AF2C8BC630202AFC32FF74FBC3FF15" authorityName="Dumitrică" authorityYear="2024" box="[927,1025,224,248]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="19" pageNumber="70" phylum="Radiozoa" rank="species" species="lahmi" status="sp. nov.">
<emphasis id="C1DB8B1AC630202AFC32FF74FBC3FF15" box="[927,1025,224,248]" italics="true" pageId="19" pageNumber="70">H. lahmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C630202AFBA1FF74FB88FF15" box="[1036,1098,225,248]" pageId="19" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
in size and the
<typeStatus id="2C14E9AAC630202AFAA5FF74FAF4FF14" box="[1288,1334,225,249]" pageId="19" pageNumber="70">type</typeStatus>
of distal thoracic pores from which it differs by not having the shoulder with a circumferential ring and by having the distal part of thorax well constricted and the distal thoracic border with an additional post-thoracic narrowing row of pores with thin interporal bars.
</paragraph>
<paragraph id="F3105708C630202AFC9DFE32FB3BFE32" blockId="19.[816,1437,158,479]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFC9DFE32FBD3FE52" bold="true" box="[816,1041,423,447]" italics="true" pageId="19" pageNumber="70">Stratigraphic range</emphasis>
. A single specimen in the lower Ladinian from the Buchenstein Formation.
</paragraph>
</subSubSection>
</treatment>
</document>

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<treatment id="7B06E61EC630202DFC60FD9FFE40F7F8" LSID="urn:lsid:plazi:treatment:7B06E61EC630202DFC60FD9FFE40F7F8" httpUri="http://treatment.plazi.org/id/7B06E61EC630202DFC60FD9FFE40F7F8" lastPageId="20" lastPageNumber="71" pageId="19" pageNumber="70">
<subSubSection id="BBB50483C630202AFC60FD9FFB3DFDCF" box="[973,1279,522,546]" pageId="19" pageNumber="70" type="nomenclature">
<paragraph id="F3105708C630202AFC60FD9FFB3DFDCF" blockId="19.[973,1279,522,546]" box="[973,1279,522,546]" pageId="19" pageNumber="70">
<heading id="A858E064C630202AFC60FD9FFB3DFDCF" box="[973,1279,522,546]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="70" reason="5">
<emphasis id="C1DB8B1AC630202AFC60FD9FFB3DFDCF" bold="true" box="[973,1279,522,546]" pageId="19" pageNumber="70">
<taxonomicName id="34AF2C8BC630202AFC60FD9FFB7EFDCF" authorityName="Dumitrică" authorityYear="2024" box="[973,1212,522,546]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="19" pageNumber="70" phylum="Radiozoa" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="C1DB8B1AC630202AFC60FD9FFB7EFDCF" bold="true" box="[973,1212,522,546]" italics="true" pageId="19" pageNumber="70">Humerocyrtis gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C630202AFB6EFD9FFB3DFDCF" box="[1219,1279,522,546]" pageId="19" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFB89FDBEFB6BFDAE" box="[1060,1193,555,579]" pageId="19" pageNumber="70" type="description">
<paragraph id="F3105708C630202AFB89FDBEFB6BFDAE" blockId="19.[1060,1193,555,579]" box="[1060,1193,555,579]" pageId="19" pageNumber="70">
<figureCitation id="6B944B8DC630202AFB89FDBEFB6BFDAE" box="[1060,1193,555,579]" captionStart="Fig" captionStartId="20.[197,232,1730,1751]" captionTargetBox="[206,1394,188,1577]" captionTargetId="figure-156@20.[197,1410,178,1721]" captionTargetPageId="20" captionText="Fig. 9. a, b, f. Humerocyrtis gracilis n. sp.: a. paratype, Rc4; b. holotype, BV 85-70, f. Rc4. c. Humerocyrtis avirostrum n. sp., Rc4; d, d1, e. Humerocyrtis brevithorax n. sp., CRH: d, d1. same specimen in lateral and basal view, respectively, showing the initial spicule without D ray, e. holotype. Scale bar of 50 µm is for all figures, except fig. 7c for which the scale bar is of 75 µm." figureDoi="http://doi.org/10.5281/zenodo.12582794" httpUri="https://zenodo.org/record/12582794/files/figure.png" pageId="19" pageNumber="70">Figs. 9a, b, f</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFC9DFDF8FB63FB98" pageId="19" pageNumber="70" type="diagnosis">
<paragraph id="F3105708C630202AFC9DFDF8FBDEFBDE" blockId="19.[816,1437,621,2068]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFC9DFDF8FC5CFD68" bold="true" box="[816,926,621,645]" italics="true" pageId="19" pageNumber="70">Diagnosis</emphasis>
. Two-segmented bell-shaped nassellarian without dorsal ray in the initial spicular system. Cephalis ovoidal with partly imperforate wall and pyriform cavity with the acute end in the base of the apical horn. Apical horn more or less three-bladed to almost conical, subaxially disposed and usually slightly curved in ventral direction. Ventral spine very small outside cephalis and pyramidal. Thorax bell-shaped with a well-marked rounded shoulder and a rather deep concavity of its distal part. Pores circular or subcircular, relatively large and usually quincuncially arranged and increasing in diameter in distal direction. Intervening bars of pores relatively thin. Distal end flared with or without a narrow circular and imperforate band.
</paragraph>
<paragraph id="F3105708C630202AFC9DFBA8FB63FB98" blockId="19.[816,1437,621,2068]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFC9DFBA8FC2FFBB8" bold="true" box="[816,1005,1085,1109]" italics="true" pageId="19" pageNumber="70">Studied material</emphasis>
.
<specimenCount id="E5A99C81C630202AFBAFFBA9FB60FBB9" box="[1026,1186,1084,1108]" count="6" pageId="19" pageNumber="70" type="generic">Six specimens</specimenCount>
of which two in the sample Rc4 and four in BV 85-70.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFC9DFBEAFA92FB55" pageId="19" pageNumber="70" type="materials_examined">
<paragraph id="F3105708C630202AFC9DFBEAFA50FB7A" blockId="19.[816,1437,621,2068]" box="[816,1426,1151,1175]" pageId="19" pageNumber="70">
<materialsCitation id="43C75D55C630202AFC9DFBEAFA50FB7A" box="[816,1426,1151,1175]" collectionCode="MGL" pageId="19" pageNumber="70" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC630202AFC9DFBEAFC51FB7A" bold="true" box="[816,915,1151,1175]" italics="true" pageId="19" pageNumber="70">
<typeStatus id="2C14E9AAC630202AFC9DFBEAFC51FB7A" box="[816,915,1151,1175]" pageId="19" pageNumber="70" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC630202AFC0CFBEAFC2DFB7A" box="[929,1007,1151,1175]" captionStart="Fig" captionStartId="20.[197,232,1730,1751]" captionTargetBox="[206,1394,188,1577]" captionTargetId="figure-156@20.[197,1410,178,1721]" captionTargetPageId="20" captionText="Fig. 9. a, b, f. Humerocyrtis gracilis n. sp.: a. paratype, Rc4; b. holotype, BV 85-70, f. Rc4. c. Humerocyrtis avirostrum n. sp., Rc4; d, d1, e. Humerocyrtis brevithorax n. sp., CRH: d, d1. same specimen in lateral and basal view, respectively, showing the initial spicule without D ray, e. holotype. Scale bar of 50 µm is for all figures, except fig. 7c for which the scale bar is of 75 µm." figureDoi="http://doi.org/10.5281/zenodo.12582794" httpUri="https://zenodo.org/record/12582794/files/figure.png" pageId="19" pageNumber="70">Fig. 9b</figureCitation>
, sample BV 85-70, coll.
<collectionCode id="95BECFCDC630202AFB54FBEAFAFAFB7A" box="[1273,1336,1151,1175]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="19" pageNumber="70">MGL</collectionCode>
.110294.
</materialsCitation>
</paragraph>
<paragraph id="F3105708C630202AFC9DFB35FA92FB55" blockId="19.[816,1437,621,2068]" box="[816,1360,1184,1208]" pageId="19" pageNumber="70">
<materialsCitation id="43C75D55C630202AFC9DFB35FB35FB55" box="[816,1271,1184,1208]" collectionCode="MGL" pageId="19" pageNumber="70" specimenCount="1" typeStatus="paratype">
<emphasis id="C1DB8B1AC630202AFC9DFB35FC50FB55" bold="true" box="[816,914,1184,1208]" italics="true" pageId="19" pageNumber="70">
<typeStatus id="2C14E9AAC630202AFC9DFB35FC50FB55" box="[816,914,1184,1208]" pageId="19" pageNumber="70" type="paratype">Paratype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC630202AFC0DFB35FC2EFB55" box="[928,1004,1184,1208]" captionStart="Fig" captionStartId="20.[197,232,1730,1751]" captionTargetBox="[206,1394,188,1577]" captionTargetId="figure-156@20.[197,1410,178,1721]" captionTargetPageId="20" captionText="Fig. 9. a, b, f. Humerocyrtis gracilis n. sp.: a. paratype, Rc4; b. holotype, BV 85-70, f. Rc4. c. Humerocyrtis avirostrum n. sp., Rc4; d, d1, e. Humerocyrtis brevithorax n. sp., CRH: d, d1. same specimen in lateral and basal view, respectively, showing the initial spicule without D ray, e. holotype. Scale bar of 50 µm is for all figures, except fig. 7c for which the scale bar is of 75 µm." figureDoi="http://doi.org/10.5281/zenodo.12582794" httpUri="https://zenodo.org/record/12582794/files/figure.png" pageId="19" pageNumber="70">Fig. 9a</figureCitation>
, sample Rc4, coll.
<collectionCode id="95BECFCDC630202AFB1AFB35FB35FB55" box="[1207,1271,1184,1208]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="19" pageNumber="70">MGL</collectionCode>
</materialsCitation>
.110295.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFC9DFB54FAE3FAF1" pageId="19" pageNumber="70" type="description">
<paragraph id="F3105708C630202AFC9DFB54FAE3FAF1" blockId="19.[816,1437,621,2068]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFC9DFB54FC71FB34" bold="true" box="[816,947,1217,1241]" italics="true" pageId="19" pageNumber="70">Dimensions</emphasis>
. Total length of shell with apical horn 80-86 µm, of thorax 40-42 µm, diameter of cephalis 17-20 µm, of shoulder 37-38 µm, of distal end 57-59 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFC9DFAB1FAB9FAD1" box="[816,1403,1316,1340]" pageId="19" pageNumber="70" type="etymology">
<paragraph id="F3105708C630202AFC9DFAB1FAB9FAD1" blockId="19.[816,1437,621,2068]" box="[816,1403,1316,1340]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFC9DFAB1FC67FAD1" bold="true" box="[816,933,1316,1340]" italics="true" pageId="19" pageNumber="70">Etymology</emphasis>
. From the Latin adjectiv
<taxonomicName id="34AF2C8BC630202AFB18FAB1FAC4FAD1" authorityName="Dumitrică" authorityYear="2024" box="[1205,1286,1316,1340]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="19" pageNumber="70" phylum="Radiozoa" rank="species" species="gracilis">
<emphasis id="C1DB8B1AC630202AFB18FAB1FAC4FAD1" box="[1205,1286,1316,1340]" italics="true" pageId="19" pageNumber="70">gracilis</emphasis>
</taxonomicName>
delicate.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202DFC9DFAD3FE40F7F8" lastPageId="20" lastPageNumber="71" pageId="19" pageNumber="70" type="discussion">
<paragraph id="F3105708C630202AFC9DFAD3FAAFF87D" blockId="19.[816,1437,621,2068]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFC9DFAD3FC50FAB3" bold="true" box="[816,914,1350,1374]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFC9DFAD3FC45FAB3" bold="true" box="[816,903,1350,1374]" italics="true" pageId="19" pageNumber="70">Remark</emphasis>
s
</emphasis>
. Morphologically, this new species is very close to the upper Longorbardian species
<taxonomicName id="34AF2C8BC630202AFAA7FAF2FB33FA72" authority="(Tekin &amp; Mostler, 2005)" baseAuthorityName="Tekin &amp; Mostler" baseAuthorityYear="2005" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="19" pageNumber="70" phylum="Radiozoa" rank="species" species="pseudoillyrica">
<emphasis id="C1DB8B1AC630202AFAA7FAF2FC0BFA72" italics="true" pageId="19" pageNumber="70">Humerocyrtis pseudoillyrica</emphasis>
(
<bibRefCitation id="973E2AF9C630202AFC70FA12FB28FA72" author="Tekin, U. K. &amp; Mostler, H." box="[989,1258,1415,1439]" pageId="19" pageNumber="70" pagination="1 - 20" refId="ref18029" refString="Tekin, U. K. &amp; Mostler, H., 2005. Longobardian (Middle Triassic) entactinarian and nassellarian Radiolaria from the Dinarides of Bosnia and Herzegovina. Journal of Paleontology, 79 (1): 1 - 20." type="journal article" year="2005">Tekin &amp; Mostler, 2005</bibRefCitation>
)
</taxonomicName>
from which it differs by having the thoracic skeleton with larger pores, the apical horn much longer and slightly curved in ventral direction, the collar structure less marked, and the distal end of thorax without a broad imperforate zone, which is a general characteristic of many species of the genus
<taxonomicName id="34AF2C8BC630202AFC9DF9DBFC02F98B" authorityName="Kozur &amp; Mostler" authorityYear="1979" box="[816,960,1614,1638]" genus="Goestlingella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="70" rank="genus">
<emphasis id="C1DB8B1AC630202AFC9DF9DBFC02F98B" box="[816,960,1614,1638]" italics="true" pageId="19" pageNumber="70">Goestlingella</emphasis>
</taxonomicName>
. Although the ventral spine of
<taxonomicName id="34AF2C8BC630202AFA2EF9DAFC0BF96A" baseAuthorityName="Tekin &amp; Mostler" baseAuthorityYear="2005" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="19" pageNumber="70" phylum="Radiozoa" rank="species" species="pseudoillyrica">
<emphasis id="C1DB8B1AC630202AFA2EF9DAFC0BF96A" italics="true" pageId="19" pageNumber="70">H. pseudoillyrica</emphasis>
</taxonomicName>
is not mentioned in their description and not visible on their images, it must exist because it is a characteristic element of the cephalis of the genera
<taxonomicName id="34AF2C8BC630202AFC9DF947FC02F907" authorityName="Kozur &amp; Mostler" authorityYear="1979" box="[816,960,1746,1770]" genus="Goestlingella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="70" rank="genus">
<emphasis id="C1DB8B1AC630202AFC9DF947FC02F907" box="[816,960,1746,1770]" italics="true" pageId="19" pageNumber="70">Goestlingella</emphasis>
</taxonomicName>
and
<taxonomicName id="34AF2C8BC630202AFC5AF946FB4BF907" authorityName="Dumitrică" authorityYear="2024" box="[1015,1161,1747,1770]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="19" pageNumber="70" phylum="Radiozoa" rank="genus">
<emphasis id="C1DB8B1AC630202AFC5AF946FB4BF907" box="[1015,1161,1747,1770]" italics="true" pageId="19" pageNumber="70">Humerocyrtis</emphasis>
</taxonomicName>
. Its absence must be due to the dorsal position of the illustrated shells. The shape of this species is also very similar to the coeval species
<taxonomicName id="34AF2C8BC630202AFC9DF8A0FB2AF8A0" authority="Dumitrica 2017" authorityName="Dumitrica" authorityYear="2017" box="[816,1256,1845,1870]" genus="Goestlingella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="70" rank="species" species="formosa">
<emphasis id="C1DB8B1AC630202AFC9DF8A0FBE4F8A0" box="[816,1062,1845,1869]" italics="true" pageId="19" pageNumber="70">Goestlingella formosa</emphasis>
<bibRefCitation id="973E2AF9C630202AFB98F8A0FB2AF8A0" author="Dumitrica, P." box="[1077,1256,1845,1870]" pageId="19" pageNumber="70" pagination="33 - 85" refId="ref17279" refString="Dumitrica, P., 2017. On the status of the Triassic Nassellarian radiolarian family Tetraspinocyrtiidae Kozur and Mostler and description of some related taxa. Revue de micropaleontologie, 60 (2017) 33 - 85." type="journal article" year="2017">Dumitrică 2017</bibRefCitation>
</taxonomicName>
, from which it only differs by having a three-bladed apical horn, and the distal border without a rather broad imperforate zone.
</paragraph>
<paragraph id="F3105708C630202DFC9DF80CFE83F87D" blockId="19.[816,1437,621,2068]" lastBlockId="20.[151,771,1879,2069]" lastPageId="20" lastPageNumber="71" pageId="19" pageNumber="70">
The specimen illustrated in
<figureCitation id="6B944B8DC630202AFBCEF80CFB6EF85C" box="[1123,1196,1945,1969]" captionStart="Fig" captionStartId="20.[197,232,1730,1751]" captionTargetBox="[206,1394,188,1577]" captionTargetId="figure-156@20.[197,1410,178,1721]" captionTargetPageId="20" captionText="Fig. 9. a, b, f. Humerocyrtis gracilis n. sp.: a. paratype, Rc4; b. holotype, BV 85-70, f. Rc4. c. Humerocyrtis avirostrum n. sp., Rc4; d, d1, e. Humerocyrtis brevithorax n. sp., CRH: d, d1. same specimen in lateral and basal view, respectively, showing the initial spicule without D ray, e. holotype. Scale bar of 50 µm is for all figures, except fig. 7c for which the scale bar is of 75 µm." figureDoi="http://doi.org/10.5281/zenodo.12582794" httpUri="https://zenodo.org/record/12582794/files/figure.png" pageId="19" pageNumber="70">Fig. 9f</figureCitation>
from the sample Rc4 is included in this species by its shape and disposition of pores, but difers from the
<typeStatus id="2C14E9AAC630202AFBC1F84EFB0BF81E" box="[1132,1225,2011,2035]" pageId="19" pageNumber="70" type="holotype">holotype</typeStatus>
and all the
<typeStatus id="2C14E9AAC630202AFAC8F84EFC5DF7F9" isOtherTypeStatus="true" pageId="19" pageNumber="70">other specimens</typeStatus>
from the sample BV 85-70 by having wider intervening bars and smaller pores due probably to its robust skeleton.
</paragraph>
<caption id="A7D00780C637202DFF68F957FE3BF8DC" ID-DOI="http://doi.org/10.5281/zenodo.12582794" ID-Zenodo-Dep="12582794" httpUri="https://zenodo.org/record/12582794/files/figure.png" pageId="20" pageNumber="71" startId="20.[197,232,1730,1751]" targetBox="[206,1394,188,1577]" targetPageId="20" targetType="figure">
<paragraph id="F3105708C637202DFF68F957FE3BF8DC" blockId="20.[197,1412,1730,1841]" pageId="20" pageNumber="71">
<emphasis id="C1DB8B1AC637202DFF68F957FEC1F93A" bold="true" box="[197,259,1730,1751]" pageId="20" pageNumber="71">Fig. 9</emphasis>
.
<emphasis id="C1DB8B1AC637202DFEB9F957FEE2F93A" bold="true" box="[276,288,1730,1751]" pageId="20" pageNumber="71">a</emphasis>
,
<emphasis id="C1DB8B1AC637202DFE9DF957FEFFF93A" bold="true" box="[304,317,1730,1751]" pageId="20" pageNumber="71">b</emphasis>
,
<emphasis id="C1DB8B1AC637202DFEE3F957FE9EF93A" bold="true" box="[334,348,1730,1751]" pageId="20" pageNumber="71">f.</emphasis>
<taxonomicName id="34AF2C8BC637202DFED1F957FD96F935" authorityName="Dumitrică" authorityYear="2024" box="[380,596,1730,1752]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="20" pageNumber="71" phylum="Radiozoa" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="C1DB8B1AC637202DFED1F957FD96F935" box="[380,596,1730,1752]" italics="true" pageId="20" pageNumber="71">Humerocyrtis gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C637202DFDF2F957FD5BF93A" box="[607,665,1730,1751]" pageId="20" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="C1DB8B1AC637202DFD0AF957FD7BF93A" bold="true" box="[679,697,1730,1751]" pageId="20" pageNumber="71">a.</emphasis>
paratype, Rc4;
<emphasis id="C1DB8B1AC637202DFCD8F957FC4AF93A" bold="true" box="[885,904,1730,1751]" pageId="20" pageNumber="71">b.</emphasis>
holotype, BV 85-70,
<emphasis id="C1DB8B1AC637202DFB2CF957FB4BF93A" bold="true" box="[1153,1161,1730,1751]" pageId="20" pageNumber="71">f</emphasis>
. Rc4.
<emphasis id="C1DB8B1AC637202DFB70F957FB2CF93A" bold="true" box="[1245,1262,1730,1751]" pageId="20" pageNumber="71">c.</emphasis>
<taxonomicName id="34AF2C8BC637202DFAA3F956FE89F918" authorityName="Dumitrică" authorityYear="2024" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="20" pageNumber="71" phylum="Radiozoa" rank="species" species="avirostrum" status="sp. nov.">
<emphasis id="C1DB8B1AC637202DFAA3F956FE89F918" italics="true" pageId="20" pageNumber="71">Humerocyrtis avirostrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C637202DFEFCF975FE45F918" box="[337,391,1760,1781]" pageId="20" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
, Rc4;
<emphasis id="C1DB8B1AC637202DFE68F975FE10F918" bold="true" box="[453,466,1760,1781]" pageId="20" pageNumber="71">d</emphasis>
,
<emphasis id="C1DB8B1AC637202DFE73F975FE3AF918" bold="true" box="[478,504,1760,1781]" pageId="20" pageNumber="71">d1</emphasis>
,
<emphasis id="C1DB8B1AC637202DFDAEF975FDD6F918" bold="true" box="[515,532,1760,1781]" pageId="20" pageNumber="71">e.</emphasis>
<taxonomicName id="34AF2C8BC637202DFD80F975FCE4F918" authorityName="Dumitrică" authorityYear="2024" box="[557,806,1759,1781]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="20" pageNumber="71" phylum="Radiozoa" rank="species" species="brevithorax" status="sp. nov.">
<emphasis id="C1DB8B1AC637202DFD80F975FCE4F918" box="[557,806,1759,1781]" italics="true" pageId="20" pageNumber="71">Humerocyrtis brevithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C637202DFC80F975FCA0F918" box="[813,866,1760,1781]" pageId="20" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
, CRH:
<emphasis id="C1DB8B1AC637202DFC07F975FC75F918" bold="true" box="[938,951,1760,1781]" pageId="20" pageNumber="71">d</emphasis>
,
<emphasis id="C1DB8B1AC637202DFC69F975FC21F918" bold="true" box="[964,995,1760,1781]" pageId="20" pageNumber="71">d1.</emphasis>
same specimen in lateral and basal view, respectively, showing the initial spicule without D ray,
<emphasis id="C1DB8B1AC637202DFD4EF968FD36F8FF" bold="true" box="[739,756,1789,1810]" pageId="20" pageNumber="71">e.</emphasis>
holotype. Scale bar of 50 µm is for all figures, except fig. 7c for which the scale bar is of 75 µm.
</paragraph>
</caption>
<paragraph id="F3105708C637202DFF3AF80FFE40F7F8" blockId="20.[151,771,1879,2069]" pageId="20" pageNumber="71">
<emphasis id="C1DB8B1AC637202DFF3AF80FFEB9F85F" bold="true" box="[151,379,1946,1970]" pageId="20" pageNumber="71">
<emphasis id="C1DB8B1AC637202DFF3AF80FFEB6F85F" bold="true" box="[151,372,1946,1970]" italics="true" pageId="20" pageNumber="71">Stratigraphic range</emphasis>
.
</emphasis>
Frequent in the lower Ladinian of the Livinallongo Formation, Marmolada Massif, sample BV 85-70, and rather rare in the Buchenstein Formation, Recoaro, sample Rc4.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,138 @@
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<treatment id="7B06E61EC631202AFC69F8F7FE65FBFF" LSID="urn:lsid:plazi:treatment:7B06E61EC631202AFC69F8F7FE65FBFF" httpUri="http://treatment.plazi.org/id/7B06E61EC631202AFC69F8F7FE65FBFF" lastPageId="19" lastPageNumber="70" pageId="18" pageNumber="69">
<subSubSection id="BBB50483C631202BFC69F8F7FACAF897" box="[964,1288,1890,1914]" pageId="18" pageNumber="69" type="nomenclature">
<paragraph id="F3105708C631202BFC69F8F7FACAF897" blockId="18.[964,1288,1890,1914]" box="[964,1288,1890,1914]" pageId="18" pageNumber="69">
<heading id="A858E064C631202BFC69F8F7FACAF897" box="[964,1288,1890,1914]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="69" reason="5">
<emphasis id="C1DB8B1AC631202BFC69F8F7FACAF897" bold="true" box="[964,1288,1890,1914]" pageId="18" pageNumber="69">
<taxonomicName id="34AF2C8BC631202BFC69F8F7FB04F897" authorityName="Dumitrică" authorityYear="2024" box="[964,1222,1890,1914]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="18" pageNumber="69" phylum="Radiozoa" rank="species" species="undulata" status="sp. nov.">
<emphasis id="C1DB8B1AC631202BFC69F8F7FB04F897" bold="true" box="[964,1222,1890,1914]" italics="true" pageId="18" pageNumber="69">Humerocyrtis undulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C631202BFB61F8F7FACAF897" box="[1228,1288,1890,1914]" pageId="18" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C631202BFB8FF817FB69F877" box="[1058,1195,1922,1946]" pageId="18" pageNumber="69" type="description">
<paragraph id="F3105708C631202BFB8FF817FB69F877" blockId="18.[1058,1195,1922,1946]" box="[1058,1195,1922,1946]" pageId="18" pageNumber="69">
<figureCitation id="6B944B8DC631202BFB8FF817FBBBF877" box="[1058,1145,1922,1946]" captionStart="Fig" captionStartId="18.[197,232,1770,1791]" captionTargetBox="[211,1402,191,1591]" captionTargetId="figure-168@18.[197,1410,178,1721]" captionTargetPageId="18" captionText="Fig. 8. a. Humerocyrtis contraria n. sp., holotype, Rc4; b. Humerocyrtis conica n. sp., holotype, Rc4. c. Humerocyrtis jekeli n. sp., holotype, Rc2; e,?d. Humerocyrtis undulata n. sp., Rc4: e holotype. f Humerocyrtis lahmi n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582792" httpUri="https://zenodo.org/record/12582792/files/figure.png" pageId="18" pageNumber="69">Figs. 8e</figureCitation>
,?8d
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C631202AFC9DF850FD65FE32" lastPageId="19" lastPageNumber="70" pageId="18" pageNumber="69" type="diagnosis">
<paragraph id="F3105708C631202AFC9DF850FD1DFE53" blockId="18.[816,1437,1988,2079]" lastBlockId="19.[151,772,158,1042]" lastPageId="19" lastPageNumber="70" pageId="18" pageNumber="69">
<emphasis id="C1DB8B1AC631202BFC9DF850FC5CF830" bold="true" box="[816,926,1989,2013]" italics="true" pageId="18" pageNumber="69">Diagnosis</emphasis>
. Shell bell-shaped and thick-walled with a three-bladed, robust, pointed and slightly curved apical horn. One of its blades is in the plane of the ventral spine and the other two are laterally directed. Ventral spine very short and pointed. Cephalis cavity pear-shaped with pointed end in apical direction and its wall practically poreless. Collar boundary marked by a slight constriction and by the appearance of pores. Thorax short, thick-walled, with evident shoulder but without a circumferential ring. Distal end thick with undulate border. Pores of thorax rounded, separated by wide intervening bars and relatively quincuncially arranged.
</paragraph>
<paragraph id="F3105708C630202AFF3AFE5DFD65FE32" blockId="19.[151,772,158,1042]" box="[151,679,455,480]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AFE5DFE8FFE0D" bold="true" box="[151,333,456,480]" italics="true" pageId="19" pageNumber="70">Studied material</emphasis>
.
<specimenCount id="E5A99C81C630202AFEF7FE5DFDC3FE32" box="[346,513,455,479]" count="2" pageId="19" pageNumber="70" type="generic">Two specimens</specimenCount>
in sample Rc4.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFF3AFE7CFDECFDCF" pageId="19" pageNumber="70" type="materials_examined">
<paragraph id="F3105708C630202AFF3AFE7CFDECFDEC" blockId="19.[151,772,158,1042]" box="[151,558,489,513]" pageId="19" pageNumber="70">
<materialsCitation id="43C75D55C630202AFF3AFE7CFDECFDEC" box="[151,558,489,513]" collectionCode="MGL" pageId="19" pageNumber="70" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC630202AFF3AFE7CFF38FDEC" bold="true" box="[151,250,489,513]" italics="true" pageId="19" pageNumber="70">
<typeStatus id="2C14E9AAC630202AFF3AFE7CFF38FDEC" box="[151,250,489,513]" pageId="19" pageNumber="70" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC630202AFEA5FE7CFE96FDEC" box="[264,340,489,513]" captionStart="Fig" captionStartId="18.[197,232,1770,1791]" captionTargetBox="[211,1402,191,1591]" captionTargetId="figure-168@18.[197,1410,178,1721]" captionTargetPageId="18" captionText="Fig. 8. a. Humerocyrtis contraria n. sp., holotype, Rc4; b. Humerocyrtis conica n. sp., holotype, Rc4. c. Humerocyrtis jekeli n. sp., holotype, Rc2; e,?d. Humerocyrtis undulata n. sp., Rc4: e holotype. f Humerocyrtis lahmi n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582792" httpUri="https://zenodo.org/record/12582792/files/figure.png" pageId="19" pageNumber="70">Fig. 8e</figureCitation>
, coll.
<collectionCode id="95BECFCDC630202AFE39FE7CFE16FDEC" box="[404,468,489,513]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="19" pageNumber="70">MGL</collectionCode>
.110291.
</materialsCitation>
</paragraph>
<paragraph id="F3105708C630202AFF3AFD9FFDECFDCF" blockId="19.[151,772,158,1042]" box="[151,558,522,546]" pageId="19" pageNumber="70">
<materialsCitation id="43C75D55C630202AFF3AFD9FFE16FDCF" box="[151,468,522,546]" collectionCode="MGL" pageId="19" pageNumber="70" specimenCount="1" typeStatus="paratype">
<emphasis id="C1DB8B1AC630202AFF3AFD9FFF3BFDCF" bold="true" box="[151,249,522,546]" italics="true" pageId="19" pageNumber="70">
<typeStatus id="2C14E9AAC630202AFF3AFD9FFF3BFDCF" box="[151,249,522,546]" pageId="19" pageNumber="70" type="paratype">Paratype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC630202AFEABFD9FFE96FDCF" box="[262,340,522,546]" captionStart="Fig" captionStartId="18.[197,232,1770,1791]" captionTargetBox="[211,1402,191,1591]" captionTargetId="figure-168@18.[197,1410,178,1721]" captionTargetPageId="18" captionText="Fig. 8. a. Humerocyrtis contraria n. sp., holotype, Rc4; b. Humerocyrtis conica n. sp., holotype, Rc4. c. Humerocyrtis jekeli n. sp., holotype, Rc2; e,?d. Humerocyrtis undulata n. sp., Rc4: e holotype. f Humerocyrtis lahmi n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582792" httpUri="https://zenodo.org/record/12582792/files/figure.png" pageId="19" pageNumber="70">Fig. 8d</figureCitation>
, coll.
<collectionCode id="95BECFCDC630202AFE38FD9FFE16FDCF" box="[405,468,522,546]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="19" pageNumber="70">MGL</collectionCode>
</materialsCitation>
.110292.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFF3AFDBEFE5CFD4A" pageId="19" pageNumber="70" type="description">
<paragraph id="F3105708C630202AFF3AFDBEFE5CFD4A" blockId="19.[151,772,158,1042]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AFDBEFED8FDAE" bold="true" box="[151,282,555,579]" italics="true" pageId="19" pageNumber="70">Dimensions</emphasis>
. Length of shell with apical horn 120-140 µm, of apical horn 40-65 µm, length of cephalis 25-30 µm, of thorax 50-55 µm, diameter of shoulder 60-64 µm, of distal end 72-?95 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFF3AFD25FE32FD05" pageId="19" pageNumber="70" type="etymology">
<paragraph id="F3105708C630202AFF3AFD25FE32FD05" blockId="19.[151,772,158,1042]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AFD25FECEFD25" bold="true" box="[151,268,688,712]" italics="true" pageId="19" pageNumber="70">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC630202AFE6AFD3AFDF2FD2A" box="[455,560,687,711]" italics="true" pageId="19" pageNumber="70">undulatus</emphasis>
, -
<emphasis id="C1DB8B1AC630202AFDE5FD25FD97FD2A" box="[584,597,688,711]" italics="true" pageId="19" pageNumber="70">a</emphasis>
, -
<emphasis id="C1DB8B1AC630202AFDC0FD25FD4CFD2A" box="[621,654,688,711]" italics="true" pageId="19" pageNumber="70">um</emphasis>
winding due to its undulate distal border.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C630202AFF3AFD67FE65FBFF" pageId="19" pageNumber="70" type="discussion">
<paragraph id="F3105708C630202AFF3AFD67FD2AFC3D" blockId="19.[151,772,158,1042]" pageId="19" pageNumber="70">
<emphasis id="C1DB8B1AC630202AFF3AFD67FF3BFCE7" bold="true" box="[151,249,754,778]" italics="true" pageId="19" pageNumber="70">Remarks</emphasis>
. This species differs from the other species of this genus herein described by the undulate distal border. The specimen illustrated in
<figureCitation id="6B944B8DC630202AFE43FCA6FD87FCA6" box="[494,581,819,843]" captionStart="Fig" captionStartId="18.[197,232,1770,1791]" captionTargetBox="[211,1402,191,1591]" captionTargetId="figure-168@18.[197,1410,178,1721]" captionTargetPageId="18" captionText="Fig. 8. a. Humerocyrtis contraria n. sp., holotype, Rc4; b. Humerocyrtis conica n. sp., holotype, Rc4. c. Humerocyrtis jekeli n. sp., holotype, Rc2; e,?d. Humerocyrtis undulata n. sp., Rc4: e holotype. f Humerocyrtis lahmi n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582792" httpUri="https://zenodo.org/record/12582792/files/figure.png" pageId="19" pageNumber="70">Fig. 8d</figureCitation>
is questionably included in this species because its thorax is less thick, the shoulder less marked and the distal end less undulated. By its overall morphology it seems to be intermediate between
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and
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<emphasis id="C1DB8B1AC630202AFD9BFC2CFD6AFC3D" box="[566,680,952,976]" italics="true" pageId="19" pageNumber="70">H. gracilis</emphasis>
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<taxonomicNameLabel id="DAE83661C630202AFD02FC2CFD2AFC3D" box="[687,744,953,976]" pageId="19" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
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<emphasis id="C1DB8B1AC630202AFF3AFC4CFEB7FC1C" bold="true" box="[151,373,985,1009]" italics="true" pageId="19" pageNumber="70">Stratigraphic range</emphasis>
. Sparse in Buchenstein Formation, lower Ladinian, Recoaro.
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<emphasis id="C1DB8B1AC6322028FC6EFEF0FB05FE90" bold="true" box="[963,1223,357,381]" italics="true" pageId="17" pageNumber="68">Humerocyrtis contraria</emphasis>
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<figureCitation id="6B944B8DC6322028FBECFE10FB4EFE70" box="[1089,1164,389,413]" captionStart="Fig" captionStartId="18.[197,232,1770,1791]" captionTargetBox="[211,1402,191,1591]" captionTargetId="figure-168@18.[197,1410,178,1721]" captionTargetPageId="18" captionText="Fig. 8. a. Humerocyrtis contraria n. sp., holotype, Rc4; b. Humerocyrtis conica n. sp., holotype, Rc4. c. Humerocyrtis jekeli n. sp., holotype, Rc2; e,?d. Humerocyrtis undulata n. sp., Rc4: e holotype. f Humerocyrtis lahmi n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582792" httpUri="https://zenodo.org/record/12582792/files/figure.png" pageId="17" pageNumber="68">Fig. 8a</figureCitation>
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<emphasis id="C1DB8B1AC6322028FC9DFE5DFC5CFE0D" bold="true" box="[816,926,456,480]" italics="true" pageId="17" pageNumber="68">Diagnosis</emphasis>
. Test relatively short, bell-shaped with three-bladed, robust and pointed apical horn. Ventral horn pointed relatively long by comparison with other related species. Cephalis practically imperforated except a few very small pores at its lower part. Collar boundary difficult to distinguished but it could be considered to be under the level of the median bar where the pores become larger. Thorax with shoulder well marked by a rather acute angle in lateral view. Distal part of thorax very short, with a rather deep concavity under the shoulder followed by a flare-shaped outline. Distal border irregular, with a beginning of a thin additional wall developed in antapical direction.
</paragraph>
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<emphasis id="C1DB8B1AC6322028FC9DFCE3FC24FC63" bold="true" box="[816,998,886,910]" italics="true" pageId="17" pageNumber="68">Studied material</emphasis>
. A single specimen in sample Rc4.
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<typeStatus id="2C14E9AAC6322028FC9DFC02FC51FC42" box="[816,915,919,943]" pageId="17" pageNumber="68" type="holotype">Holotype</typeStatus>
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.
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, sample Rc4, coll.
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.110289.
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<emphasis id="C1DB8B1AC6322028FC9DFC2DFC71FC3D" bold="true" box="[816,947,952,976]" italics="true" pageId="17" pageNumber="68">Dimensions</emphasis>
. Total length of shell 140 µm, of thorax 60 µm, diameter of shoulder 71 µm, of distal end of thorax 90 µm.
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<subSubSection id="BBB50483C6322028FC9DFB89FC78FB7A" pageId="17" pageNumber="68" type="etymology">
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<emphasis id="C1DB8B1AC6322028FC9DFB89FC67FBD9" bold="true" box="[816,933,1052,1076]" italics="true" pageId="17" pageNumber="68">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC6322028FBD9FB89FB21FBDE" box="[1140,1251,1052,1075]" italics="true" pageId="17" pageNumber="68">contrarius</emphasis>
, -
<emphasis id="C1DB8B1AC6322028FAADFB89FAD7FBDE" box="[1280,1301,1052,1075]" italics="true" pageId="17" pageNumber="68">ia</emphasis>
contrary, because its ventral spine is longer than that of other species of the genus
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with similar morphology.
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<emphasis id="C1DB8B1AC6322028FC9DFB35FC50FB55" bold="true" box="[816,914,1184,1208]" italics="true" pageId="17" pageNumber="68">Remarks</emphasis>
. This species resembles most of the small species of the genus
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with three-bladed apical horn by its general shape, but differs from them by having a longer and more robust ventral horn, thoracic pores with very variable sizes and irregular arrangement. It differs also by its irregular distal thoracic border.
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<emphasis id="C1DB8B1AC6322028FC9DFAF3FBCEFA93" bold="true" box="[816,1036,1382,1406]" italics="true" pageId="17" pageNumber="68">Stratigraphic range</emphasis>
. Very rare in the sample Rc4 from the Buchenstein Formation from Recoaro, North
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.
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<emphasis id="C1DB8B1AC6322028FC7FFA5FFB39FA0F" bold="true" box="[978,1275,1482,1506]" pageId="17" pageNumber="68">
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<emphasis id="C1DB8B1AC6322028FC7FFA5FFB7BFA0F" bold="true" box="[978,1209,1482,1506]" italics="true" pageId="17" pageNumber="68">Humerocyrtis conica</emphasis>
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<taxonomicName id="34AF2C8BC6322028FE92FF0BFDDAFF5B" authorityName="Dumitrică" authorityYear="2024" box="[319,536,158,182]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="17" pageNumber="68" phylum="Radiozoa" rank="species" species="jekeli" status="sp. nov.">
<emphasis id="C1DB8B1AC6322028FE92FF0BFDDAFF5B" bold="true" box="[319,536,158,182]" italics="true" pageId="17" pageNumber="68">Humerocyrtis jekeli</emphasis>
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<subSubSection id="BBB50483C6322028FE22FF2AFDCEFF3A" box="[399,524,191,215]" pageId="17" pageNumber="68" type="description">
<paragraph id="F3105708C6322028FE22FF2AFDCEFF3A" blockId="17.[399,524,191,215]" box="[399,524,191,215]" pageId="17" pageNumber="68">
<figureCitation id="6B944B8DC6322028FE22FF2AFE25FF3A" box="[399,487,191,215]" captionStart="Fig" captionStartId="15.[197,232,1741,1762]" captionTargetBox="[213,1405,175,1620]" captionTargetId="figure-137@15.[197,1418,160,1713]" captionTargetPageId="15" captionText="Fig. 6. a, b. Gorispela (?) triangulocephalis n. sp., FÖ 87: a. specimen showing internal structure, b. holotype, D dorsal ray. c. Humerocyrtis superba n. sp., holotype, BV 85-70; 6d, 6c. Humerocyrtis jekeli n. sp., Rc4, 6c. holotype, d. - paratype; e. Humerocyrtis deweveri n. sp., holotype, BW 85-70; f. Humerocyrtis problematica n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582788" httpUri="https://zenodo.org/record/12582788/files/figure.png" pageId="17" pageNumber="68">Figs. 6d</figureCitation>
,
<figureCitation id="6B944B8DC6322028FE5FFF2AFDCEFF3A" box="[498,524,191,215]" captionStart="Fig" captionStartId="18.[197,232,1770,1791]" captionTargetBox="[211,1402,191,1591]" captionTargetId="figure-168@18.[197,1410,178,1721]" captionTargetPageId="18" captionText="Fig. 8. a. Humerocyrtis contraria n. sp., holotype, Rc4; b. Humerocyrtis conica n. sp., holotype, Rc4. c. Humerocyrtis jekeli n. sp., holotype, Rc2; e,?d. Humerocyrtis undulata n. sp., Rc4: e holotype. f Humerocyrtis lahmi n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582792" httpUri="https://zenodo.org/record/12582792/files/figure.png" pageId="17" pageNumber="68">8c</figureCitation>
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<subSubSection id="BBB50483C6322028FF3AFE94FD65FD4B" pageId="17" pageNumber="68" type="diagnosis">
<paragraph id="F3105708C6322028FF3AFE94FF10FD68" blockId="17.[151,772,257,1307]" pageId="17" pageNumber="68">
<emphasis id="C1DB8B1AC6322028FF3AFE94FECEFEF4" box="[151,268,257,281]" italics="true" pageId="17" pageNumber="68">
<emphasis id="C1DB8B1AC6322028FF3AFE94FEC7FEF4" bold="true" box="[151,261,257,281]" italics="true" pageId="17" pageNumber="68">Diagnosis</emphasis>
.
</emphasis>
Shell bell-shaped consisting of a small, pear-shaped cephalic cavity and practically poreles cephalis and a thorax with concave distal portion. Apical horn, long, three-bladed and pointed. Ventral spine small, three-bladed and pointed. Proximal part of thorax very short, wide-conical with rare small circular or oval pores. Shoulder relatively well remarked and rounded. Distal part of thorax subcylindrical with concave outline, its distal end simple or bearing a short and narrower thin-walled additional shell. Pores of thorax circular or oval, of variable sizes and arranged irregularly or in oblique rows.
</paragraph>
<paragraph id="F3105708C6322028FF3AFD1BFD65FD4B" blockId="17.[151,772,257,1307]" box="[151,679,654,678]" pageId="17" pageNumber="68">
<emphasis id="C1DB8B1AC6322028FF3AFD1BFE8FFD4B" bold="true" box="[151,333,654,678]" italics="true" pageId="17" pageNumber="68">Studied material</emphasis>
.
<specimenCount id="E5A99C81C6322028FEF7FD1AFDC3FD4B" box="[346,513,654,678]" count="2" pageId="17" pageNumber="68" type="generic">Two specimens</specimenCount>
in sample Rc4.
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<paragraph id="F3105708C6322028FF3AFD25FD7AFD2A" blockId="17.[151,772,257,1307]" box="[151,696,687,712]" pageId="17" pageNumber="68">
<materialsCitation id="43C75D55C6322028FF3AFD25FD7AFD2A" box="[151,696,687,712]" collectionCode="MGL" pageId="17" pageNumber="68" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC6322028FF3AFD25FF38FD25" bold="true" box="[151,250,688,712]" italics="true" pageId="17" pageNumber="68">
<typeStatus id="2C14E9AAC6322028FF3AFD25FF38FD25" box="[151,250,688,712]" pageId="17" pageNumber="68" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC6322028FEA5FD3AFE96FD2A" box="[264,340,687,711]" captionStart="Fig" captionStartId="18.[197,232,1770,1791]" captionTargetBox="[211,1402,191,1591]" captionTargetId="figure-168@18.[197,1410,178,1721]" captionTargetPageId="18" captionText="Fig. 8. a. Humerocyrtis contraria n. sp., holotype, Rc4; b. Humerocyrtis conica n. sp., holotype, Rc4. c. Humerocyrtis jekeli n. sp., holotype, Rc2; e,?d. Humerocyrtis undulata n. sp., Rc4: e holotype. f Humerocyrtis lahmi n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582792" httpUri="https://zenodo.org/record/12582792/files/figure.png" pageId="17" pageNumber="68">Fig. 8c</figureCitation>
, sample Rc2, coll.
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.110286.
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<materialsCitation id="43C75D55C6322028FF3AFD44FD9DFD05" box="[151,607,720,745]" collectionCode="MGL" pageId="17" pageNumber="68" specimenCount="1" typeStatus="paratype">
<emphasis id="C1DB8B1AC6322028FF3AFD44FF3BFD04" bold="true" box="[151,249,721,745]" italics="true" pageId="17" pageNumber="68">
<typeStatus id="2C14E9AAC6322028FF3AFD44FF3BFD04" box="[151,249,721,745]" pageId="17" pageNumber="68" type="paratype">Paratype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC6322028FEABFD45FE96FD05" box="[262,340,720,744]" captionStart="Fig" captionStartId="15.[197,232,1741,1762]" captionTargetBox="[213,1405,175,1620]" captionTargetId="figure-137@15.[197,1418,160,1713]" captionTargetPageId="15" captionText="Fig. 6. a, b. Gorispela (?) triangulocephalis n. sp., FÖ 87: a. specimen showing internal structure, b. holotype, D dorsal ray. c. Humerocyrtis superba n. sp., holotype, BV 85-70; 6d, 6c. Humerocyrtis jekeli n. sp., Rc4, 6c. holotype, d. - paratype; e. Humerocyrtis deweveri n. sp., holotype, BW 85-70; f. Humerocyrtis problematica n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582788" httpUri="https://zenodo.org/record/12582788/files/figure.png" pageId="17" pageNumber="68">Fig. 6d</figureCitation>
, sample Rc4, coll.
<collectionCode id="95BECFCDC6322028FDB2FD45FD9DFD05" box="[543,607,720,744]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="17" pageNumber="68">MGL</collectionCode>
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.110287.
</paragraph>
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<subSubSection id="BBB50483C6322028FF3AFD67FD57FC80" pageId="17" pageNumber="68" type="description">
<paragraph id="F3105708C6322028FF3AFD67FD57FC80" blockId="17.[151,772,257,1307]" pageId="17" pageNumber="68">
<emphasis id="C1DB8B1AC6322028FF3AFD67FED8FCE7" bold="true" box="[151,282,754,778]" italics="true" pageId="17" pageNumber="68">Dimensions</emphasis>
. Maximum length of shell with apical horn 126-144 µm, without horn 87-96 µm, of cephalis 30-35 µm, of thorax 52-61 µm, diameter of cephalis 37-38 µm, of shoulder 57-58 µm, of distal end 95-106 µm.
</paragraph>
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<subSubSection id="BBB50483C6322028FF3AFCE3FE9CFC1C" pageId="17" pageNumber="68" type="etymology">
<paragraph id="F3105708C6322028FF3AFCE3FE9CFC1C" blockId="17.[151,772,257,1307]" pageId="17" pageNumber="68">
<emphasis id="C1DB8B1AC6322028FF3AFCE3FECEFC63" bold="true" box="[151,268,886,910]" italics="true" pageId="17" pageNumber="68">Etymology</emphasis>
. The species is dedicated to the geologist E. Jekelius who, at the beginning of the XXth century, studied the geology and Triassic fauna from the
<collectingRegion id="316B99EAC6322028FD15FC2CFCC1FC3D" box="[696,771,953,976]" country="Romania" name="Brasov" pageId="17" pageNumber="68">Braşov</collectingRegion>
and Cristian areas.
</paragraph>
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<subSubSection id="BBB50483C6322028FF3AFC6FFE83FAF6" pageId="17" pageNumber="68" type="discussion">
<paragraph id="F3105708C6322028FF3AFC6FFDC5FB34" blockId="17.[151,772,257,1307]" pageId="17" pageNumber="68">
<emphasis id="C1DB8B1AC6322028FF3AFC6FFF3BFBFF" bold="true" box="[151,249,1018,1042]" italics="true" pageId="17" pageNumber="68">Remarks</emphasis>
. To this species I assigned provisionally two rather different specimens. The
<typeStatus id="2C14E9AAC6322028FE5EFB89FD8DFBD9" box="[499,591,1052,1076]" pageId="17" pageNumber="68" type="paratype">paratype</typeStatus>
differs from the
<typeStatus id="2C14E9AAC6322028FF3AFBA9FF36FBB9" box="[151,244,1084,1108]" pageId="17" pageNumber="68" type="holotype">holotype</typeStatus>
by having a longer and slightly pear-shaped cephalis, very irregularly arranged pores on thorax and distal border of thorax practically without the thin additional shell. There is just a beginning of such a shell in the left side of the distal border.
</paragraph>
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<emphasis id="C1DB8B1AC6322028FF3AFB77FEB5FB17" bold="true" box="[151,375,1250,1274]" italics="true" pageId="17" pageNumber="68">Stratigraphic range</emphasis>
. Buchenstein Formation, Recoaro, lower Ladinian.
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<emphasis id="C1DB8B1AC6332029FC7BFC67FB76FBE7" bold="true" box="[982,1204,1010,1034]" italics="true" pageId="16" pageNumber="67">Humerocyrtis lahmi</emphasis>
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<paragraph id="F3105708C6332029FB93FB86FB4DFBC6" blockId="16.[1086,1167,1043,1067]" box="[1086,1167,1043,1067]" pageId="16" pageNumber="67">
<figureCitation id="6B944B8DC6332029FB93FB86FB4DFBC6" box="[1086,1167,1043,1067]" captionStart="Fig" captionStartId="18.[197,232,1770,1791]" captionTargetBox="[211,1402,191,1591]" captionTargetId="figure-168@18.[197,1410,178,1721]" captionTargetPageId="18" captionText="Fig. 8. a. Humerocyrtis contraria n. sp., holotype, Rc4; b. Humerocyrtis conica n. sp., holotype, Rc4. c. Humerocyrtis jekeli n. sp., holotype, Rc2; e,?d. Humerocyrtis undulata n. sp., Rc4: e holotype. f Humerocyrtis lahmi n. sp., holotype, Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582792" httpUri="https://zenodo.org/record/12582792/files/figure.png" pageId="16" pageNumber="67">Figs. 8f</figureCitation>
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<paragraph id="F3105708C6332029FC9DFBC3FB89FA55" blockId="16.[816,1437,1109,2027]" pageId="16" pageNumber="67">
<emphasis id="C1DB8B1AC6332029FC9DFBC3FC5CFB83" bold="true" box="[816,926,1110,1134]" italics="true" pageId="16" pageNumber="67">Diagnosis</emphasis>
. Skeleton dicyrtid bell-shaped, relatively large with hemispherical cephalis and a robust axial three-bladed apical horn with well-developed thin blades. Ventral horn, broken off, but should be shorter than the apical horn. Thorax divided into two parts (proximal and distal) by a thin circumferential ridge. Its pores are rather wide and arranged intercalately in seven circumferential rows. Its proximal half wide cylindrical and distal half flared. It bears seven circumferential rows of rounded or elliptical pores of relatively equal sizes. Distal border probably straight and thin.
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<emphasis id="C1DB8B1AC6332029FC9DFA57FC24FA37" bold="true" box="[816,998,1474,1498]" italics="true" pageId="16" pageNumber="67">Studied material</emphasis>
.
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in sample Rh03.
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<emphasis id="C1DB8B1AC6332029FC9DFA76FC51FA16" bold="true" box="[816,915,1507,1531]" italics="true" pageId="16" pageNumber="67">
<typeStatus id="2C14E9AAC6332029FC9DFA76FC51FA16" box="[816,915,1507,1531]" pageId="16" pageNumber="67" type="holotype">Holotype</typeStatus>
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.
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, lost.
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<emphasis id="C1DB8B1AC6332029FC9DF991FC71F9F1" bold="true" box="[816,947,1540,1564]" italics="true" pageId="16" pageNumber="67">Dimensions</emphasis>
. Height of shell with apical horn 220 µm, of distal part of thorax 127 µm, diameter of cephalis 48 µm, of shoulder 109 µm, of distal end 156 µm.
</paragraph>
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<emphasis id="C1DB8B1AC6332029FC9DF9F2FC67F992" bold="true" box="[816,933,1639,1663]" italics="true" pageId="16" pageNumber="67">Etymology</emphasis>
. The species is dedicated to Dr. Berhard Lahm for his excellent study of the Middle and Upper Triassic radiolarians from Recoaro (
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) and Grossreifling (
<collectingCountry id="8BB81798C6332029FC95F95FFC4FF90F" box="[824,909,1738,1762]" name="Austria" pageId="16" pageNumber="67">Austria</collectingCountry>
).
</paragraph>
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<emphasis id="C1DB8B1AC6332029FC9DF979FC50F8E9" bold="true" box="[816,914,1772,1796]" italics="true" pageId="16" pageNumber="67">Remarks</emphasis>
. By its general morphology and size and especially the presence of the circumferential ridge on the shoulder and the number of circumferential rows of pores, this species resembles very much
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, from which it differs by having the thoracic pores with practically similar shapes and sizes.
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<emphasis id="C1DB8B1AC6332029FC9DF827FBCDF827" bold="true" box="[816,1039,1970,1994]" italics="true" pageId="16" pageNumber="67">Stratigraphic range</emphasis>
. Very rare in the lower Ladinian, Buchenstein Formation, Recoaro area.
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<emphasis id="C1DB8B1AC6332029FECDFACDFE35FA9D" bold="true" box="[352,503,1368,1392]" italics="true" pageId="16" pageNumber="67">Humerocyrtis</emphasis>
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<taxonomicNameLabel id="DAE83661C6332029FE53FACDFDF8FA9D" box="[510,570,1368,1392]" pageId="16" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="F3105708C6332029FE03FAEDFE2EFA7D" blockId="16.[430,492,1400,1424]" box="[430,492,1400,1424]" pageId="16" pageNumber="67">
<figureCitation id="6B944B8DC6332029FE03FAEDFE2EFA7D" box="[430,492,1400,1424]" captionStart="Fig" captionStartId="16.[891,926,925,946]" captionTargetBox="[863,1380,302,890]" captionTargetId="figure-1087@16.[831,1416,291,918]" captionTargetPageId="16" captionText="Fig. 7. - Humerocyrtis n. sp., H shoulder, Rc4." figureDoi="http://doi.org/10.5281/zenodo.12582790" httpUri="https://zenodo.org/record/12582790/files/figure.png" pageId="16" pageNumber="67">Fig. 7</figureCitation>
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<emphasis id="C1DB8B1AC6332029FF3AFA2EFF3DFA3E" bold="true" box="[151,255,1467,1491]" pageId="16" pageNumber="67">Remarks</emphasis>
. This new species, of which I have only the SEM image of a specimen from a residue of the sample Rc4 offered in 1975 to Emil Pessagno jr. during the 14 th European Micropaleontological Colloquium held in
<collectingCountry id="8BB81798C6332029FF3AF9AAFF38F9BA" box="[151,250,1599,1623]" name="Romania" pageId="16" pageNumber="67">Romania</collectingCountry>
, and illustrated by him later. This image shows very clearly the thorax with the circumferential ridge of the shoulder, the six rows of pores of the distal part of thorax, its concave outline, the proximal part of thorax and cephalis. Unfortunately, the details of the cephalic surface and its morphology are blur. What one can see is that the colar boundary is well marked and the apical horn is three-bladed but the ventral spine is invisible. One can also see that the proximal part of the thorax has pores arranged in circumferential rows as its distal part, a character that is not seen at other species of this genus, and also that the distal border of the thorax has a well-marked broad platy ring.
</paragraph>
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<emphasis id="C1DB8B1AC6332029FC9DFF25FAC3FF25" bold="true" box="[816,1281,176,200]" italics="true" pageId="16" pageNumber="67">Studied material and Stratigraphic range</emphasis>
. Sample Rc4,
</paragraph>
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<heading id="A858E064C634202EFEA4FDD9FD53FD89" box="[265,657,588,612]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="74" reason="5">
<emphasis id="C1DB8B1AC634202EFEA4FDD9FD53FD89" bold="true" box="[265,657,588,612]" pageId="23" pageNumber="74">
<taxonomicName id="34AF2C8BC634202EFEA4FDD9FD8CFD89" authorityName="Dumitrică" authorityYear="2024" box="[265,590,588,612]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="23" pageNumber="74" phylum="Radiozoa" rank="species" species="marmoladensis" status="sp. nov.">
<emphasis id="C1DB8B1AC634202EFEA4FDD9FD8CFD89" bold="true" box="[265,590,588,612]" italics="true" pageId="23" pageNumber="74">Humerocyrtis marmoladensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C634202EFDF8FDD9FD53FD89" box="[597,657,588,612]" pageId="23" pageNumber="74" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="F3105708C634202EFE0EFDF8FE3AFD68" blockId="23.[419,504,621,645]" box="[419,504,621,645]" pageId="23" pageNumber="74">
<figureCitation id="6B944B8DC634202EFE0EFDF8FE3AFD68" box="[419,504,621,645]" captionStart="Fig" captionStartId="22.[197,232,1750,1771]" captionTargetBox="[212,1397,209,1690]" captionTargetId="figure-160@22.[197,1410,178,1721]" captionTargetPageId="22" captionText="Fig. 10. a Humerocyrtis infinita n. sp, Rc4. b, d Humerocyrtis brevithorax n. sp.: b R78/1b, d poorly preserved specimen, Rc4. c Humerocyrtis asymmetrica n. sp., Rc4. f Humerocyrtis marmoladensis n. sp., BV 85-70. e, g Humerocyrtis mediotriassica n. sp.: e CR25, g Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="23" pageNumber="74">Fig. 10f</figureCitation>
</paragraph>
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<subSubSection id="BBB50483C634202EFF3AFD25FF7BFBB9" pageId="23" pageNumber="74" type="diagnosis">
<paragraph id="F3105708C634202EFF3AFD25FD35FBFF" blockId="23.[151,771,687,1572]" pageId="23" pageNumber="74">
<emphasis id="C1DB8B1AC634202EFF3AFD25FEC7FD25" bold="true" box="[151,261,688,712]" italics="true" pageId="23" pageNumber="74">Diagnosis</emphasis>
. Shell bell-shaped, wide opened distally with cephalis bearing a long, straight and thin conical apical horn in axial position and a three-bladed ventral spine with thin but very broad triangular blades. Cephalis elongate globular and practically imperforated except its distal part that contained small and irregularly disposed pores. Its distal end passes gradually to the short and wide conical proximal part of thorax. Shoulder rather angular but without a circumferential ridge. Distal part of thorax wide open, slightly concave; distal end of shell irregular and of diameter larger than that of the shoulder.
</paragraph>
<paragraph id="F3105708C634202EFF33FB89FF7BFBB9" blockId="23.[151,771,687,1572]" pageId="23" pageNumber="74">
<emphasis id="C1DB8B1AC634202EFF33FB89FE97FBD9" bold="true" box="[158,341,1052,1076]" italics="true" pageId="23" pageNumber="74">Studied material</emphasis>
. The
<typeStatus id="2C14E9AAC634202EFE38FB8EFE30FBDE" box="[405,498,1051,1075]" pageId="23" pageNumber="74" type="holotype">holotype</typeStatus>
from the sample BV 85- 70.
</paragraph>
</subSubSection>
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<paragraph id="F3105708C634202EFF3AFBCBFD96FB98" blockId="23.[151,771,687,1572]" box="[151,596,1117,1142]" pageId="23" pageNumber="74">
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<typeStatus id="2C14E9AAC634202EFF3AFBCBFF38FB9B" box="[151,250,1118,1142]" pageId="23" pageNumber="74" type="holotype">Holotype</typeStatus>
</emphasis>
.
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, coll.
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.110301.
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<paragraph id="F3105708C634202EFF3AFBEAFDE6FB34" blockId="23.[151,771,687,1572]" pageId="23" pageNumber="74">
<emphasis id="C1DB8B1AC634202EFF3AFBEAFED8FB7A" bold="true" box="[151,282,1151,1175]" italics="true" pageId="23" pageNumber="74">Dimensions</emphasis>
. Length of shell without apical horn 108 µm, of apical horn 67 µm, of ventral horn 42 µm, diameter of shoulder 75 µm, of distal end 97 µm.
</paragraph>
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<subSubSection id="BBB50483C634202EFF3AFB77FD12FB17" box="[151,720,1250,1274]" pageId="23" pageNumber="74" type="etymology">
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<emphasis id="C1DB8B1AC634202EFF3AFB77FECEFB17" bold="true" box="[151,268,1250,1274]" italics="true" pageId="23" pageNumber="74">Etymology</emphasis>
. From the Marmolada Massif, north
<collectingCountry id="8BB81798C634202EFD35FB77FD0EFB17" box="[664,716,1250,1274]" name="Italy" pageId="23" pageNumber="74">Italy</collectingCountry>
.
</paragraph>
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<subSubSection id="BBB50483C634202EFF3AFA96FF0DF9C9" pageId="23" pageNumber="74" type="discussion">
<paragraph id="F3105708C634202EFF3AFA96FDE3FA2D" blockId="23.[151,771,687,1572]" pageId="23" pageNumber="74">
<emphasis id="C1DB8B1AC634202EFF3AFA96FF3BFAF6" bold="true" box="[151,249,1283,1307]" italics="true" pageId="23" pageNumber="74">Remarks</emphasis>
. The shape of the shell of
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<emphasis id="C1DB8B1AC634202EFDDDFA91FEF8FAD1" italics="true" pageId="23" pageNumber="74">Humerocyrtis marmoladensis</emphasis>
</taxonomicName>
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resembles very well that of the species
<taxonomicName id="34AF2C8BC634202EFF46FAD3FE14FAB0" authorityName="Dumitrică" authorityYear="2024" box="[235,470,1349,1373]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="23" pageNumber="74" phylum="Radiozoa" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="C1DB8B1AC634202EFF46FAD3FE14FAB0" box="[235,470,1349,1373]" italics="true" pageId="23" pageNumber="74">Humerocyrtis gracilis</emphasis>
</taxonomicName>
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, from which it differs by having larger and irregularly arranged pores, a very long and thin conical apical horn and the ventral horn thick, three-bladed and much longer.
</paragraph>
<paragraph id="F3105708C634202EFF3AFA5FFF0DF9C9" blockId="23.[151,771,687,1572]" pageId="23" pageNumber="74">
<emphasis id="C1DB8B1AC634202EFF3AFA5FFEBBFA0F" bold="true" box="[151,377,1482,1506]" pageId="23" pageNumber="74">
<emphasis id="C1DB8B1AC634202EFF3AFA5FFEB0FA0F" bold="true" box="[151,370,1482,1506]" italics="true" pageId="23" pageNumber="74">Stratigraphic range</emphasis>
.
</emphasis>
Very rare in the lower Ladinian of the Livinallongo Formation, Marmolada Massif, North
<collectingCountry id="8BB81798C634202EFF3AF999FF09F9C9" box="[151,203,1548,1572]" name="Italy" pageId="23" pageNumber="74">Italy</collectingCountry>
.
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<mods:title id="20687702418EABF772D3536464862292">New Middle Triassic Bell-Shaped Nassellarian Radiolaria From Alpine And Carpathian Areas</mods:title>
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<heading id="A858E064C636202CFC1DFDBEFADEFDAE" box="[944,1308,555,579]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="72" reason="5">
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<taxonomicName id="34AF2C8BC636202CFC1DFDBEFB1BFDAE" authorityName="Dumitrică" authorityYear="2024" box="[944,1241,555,579]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="21" pageNumber="72" phylum="Radiozoa" rank="species" species="asymmetrica" status="sp. nov.">
<emphasis id="C1DB8B1AC636202CFC1DFDBEFB1BFDAE" bold="true" box="[944,1241,555,579]" italics="true" pageId="21" pageNumber="72">Humerocyrtis asymmetrica</emphasis>
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<taxonomicNameLabel id="DAE83661C636202CFB4DFDBEFADEFDAE" box="[1248,1308,555,579]" pageId="21" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="F3105708C636202CFB9AFDD9FB54FD89" blockId="21.[1079,1174,588,612]" box="[1079,1174,588,612]" pageId="21" pageNumber="72">
<figureCitation id="6B944B8DC636202CFB9AFDD9FB4DFD89" box="[1079,1167,588,612]" captionStart="Fig" captionStartId="22.[197,232,1750,1771]" captionTargetBox="[212,1397,209,1690]" captionTargetId="figure-160@22.[197,1410,178,1721]" captionTargetPageId="22" captionText="Fig. 10. a Humerocyrtis infinita n. sp, Rc4. b, d Humerocyrtis brevithorax n. sp.: b R78/1b, d poorly preserved specimen, Rc4. c Humerocyrtis asymmetrica n. sp., Rc4. f Humerocyrtis marmoladensis n. sp., BV 85-70. e, g Humerocyrtis mediotriassica n. sp.: e CR25, g Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="21" pageNumber="72">Fig. 10c</figureCitation>
.
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<emphasis id="C1DB8B1AC636202CFC9DFD1BFC5CFD4B" bold="true" box="[816,926,654,678]" italics="true" pageId="21" pageNumber="72">Diagnosis</emphasis>
. Shell bell-shaped with subrectangular cephalis in lateral view bearing a thin apical horn tilted dorsally and curved in ventral direction and a similar but straight ventral horn. Wall of cephalis perforated by a few circular or subcircular small and irregularly scattered pores. Collar boundary not marked externally but it seems to be below the ventral horn and at the contact of the secondary lateral rays with the shell, where an evident stricture is visible. Proximal part of thorax very short and wide conical, Shoulder asymmetrical, the ventral part more pronounced than in dorsal part. Pores of shoulder and proximal part of thorax relatively wide and divided into very small areas by a secondary network of very thin bars resembling a spiders web. This network is also present on the distal part of thorax but become less frequent in distal direction. Distal part of thorax subcylindrical with a more or less pronounced concave outline. Pores large, oval, subcircular or rounded quadrangular or triangular. Distal end with a narrowing circular and irregular row of rounded rectangular or oval pores.
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<emphasis id="C1DB8B1AC636202CFC9DFAD3FC24FAB3" bold="true" box="[816,998,1350,1374]" italics="true" pageId="21" pageNumber="72">Studied material</emphasis>
. A single specimen in sample Rc4.
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<materialsCitation id="43C75D55C636202CFC9DFAF3FB16FA93" box="[816,1236,1382,1406]" collectionCode="MGL" pageId="21" pageNumber="72" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC636202CFC9DFAF3FC51FA93" bold="true" box="[816,915,1382,1406]" italics="true" pageId="21" pageNumber="72">
<typeStatus id="2C14E9AAC636202CFC9DFAF3FC51FA93" box="[816,915,1382,1406]" pageId="21" pageNumber="72" type="holotype">Holotype</typeStatus>
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.
<figureCitation id="6B944B8DC636202CFC0CFAF3FC39FA93" box="[929,1019,1382,1406]" captionStart="Fig" captionStartId="22.[197,232,1750,1771]" captionTargetBox="[212,1397,209,1690]" captionTargetId="figure-160@22.[197,1410,178,1721]" captionTargetPageId="22" captionText="Fig. 10. a Humerocyrtis infinita n. sp, Rc4. b, d Humerocyrtis brevithorax n. sp.: b R78/1b, d poorly preserved specimen, Rc4. c Humerocyrtis asymmetrica n. sp., Rc4. f Humerocyrtis marmoladensis n. sp., BV 85-70. e, g Humerocyrtis mediotriassica n. sp.: e CR25, g Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="21" pageNumber="72">Fig. 10c</figureCitation>
, coll.
<collectionCode id="95BECFCDC636202CFB96FAF3FBB9FA93" box="[1083,1147,1382,1406]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="21" pageNumber="72">MGL</collectionCode>
.110298.
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<emphasis id="C1DB8B1AC636202CFC9DFA1DFC71FA4D" bold="true" box="[816,947,1416,1440]" italics="true" pageId="21" pageNumber="72">Dimensions</emphasis>
. Length of shell without apical horn 101 µm, of thorax 78 µm, diameter of shoulder 78 µm, of distal end 90 µm.
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<emphasis id="C1DB8B1AC636202CFC9DFA7EFC67F9EE" bold="true" box="[816,933,1515,1539]" italics="true" pageId="21" pageNumber="72">Etymology</emphasis>
. From the Latin
<taxonomicName id="34AF2C8BC636202CFBC0FA79FB37F9EE" authorityName="Dumitrică" authorityYear="2024" box="[1133,1269,1516,1539]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="21" pageNumber="72" phylum="Radiozoa" rank="species" species="asymmetrica">
<emphasis id="C1DB8B1AC636202CFBC0FA79FB37F9EE" box="[1133,1269,1516,1539]" italics="true" pageId="21" pageNumber="72">asymmetrica</emphasis>
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asymmetric, because its thorax is asymmetric in lateral view.
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<emphasis id="C1DB8B1AC636202CFC9DF9B8FC50F9A8" bold="true" box="[816,914,1581,1605]" italics="true" pageId="21" pageNumber="72">Remarks</emphasis>
. This species differs from the other species of this genus with conical spines by the narrowing distal end of thorax and the frequency of the secondary network of thin spiders web inside the pores.
</paragraph>
<paragraph id="F3105708C636202CFC9DF927FAB5F907" blockId="21.[816,1436,654,1770]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFC9DF927FBCEF927" bold="true" box="[816,1036,1714,1738]" italics="true" pageId="21" pageNumber="72">Stratigraphic range</emphasis>
. Very rare in the sample Rc4 from the Buchenstein Limestone from Recoaro, North
<collectingCountry id="8BB81798C636202CFA92F947FAB1F907" box="[1343,1395,1746,1770]" name="Italy" pageId="21" pageNumber="72">Italy</collectingCountry>
.
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<emphasis id="C1DB8B1AC636202CFE99FB54FDE6FB34" bold="true" box="[308,548,1217,1241]" italics="true" pageId="21" pageNumber="72">Humerocyrtis infinita</emphasis>
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<emphasis id="C1DB8B1AC636202CFF3AFAB1FEC7FAD1" bold="true" box="[151,261,1316,1340]" italics="true" pageId="21" pageNumber="72">Diagnosis</emphasis>
. Shell bell-shaped dicyrtid consisting of cephalis and proximal part of thorax represented by the shoulder. Cephalis smooth with some rare sparse rounded and irregularly distributed pores especially on its lower part. Apical and ventral horns conical and thin and of similar length. Apical horn dorsally tilted and slightly curved in ventral direction. Ventral horn straight and obliquely directed. Collar stricture not marked, but it could be approximately established at the level of MB where the pore start becoming larger. Thorax represented by the shoulder of thorax which is very much expanded in lateral direction and has a rounded outline. Its wall has large subcircular or rounded triangular or quadrangular pores of irregular arrangement and sizes. Distal part of shoulder is short, constricted and its end is marked by a thin linear border.
</paragraph>
<paragraph id="F3105708C636202CFF3AF8A3FDB6F8A0" blockId="21.[151,771,1316,2068]" box="[151,628,1845,1870]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFF3AF8A3FE8FF8A3" bold="true" box="[151,333,1846,1870]" italics="true" pageId="21" pageNumber="72">Studied material</emphasis>
. A single specimen in Rc4.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFF3AF8C2FDF7F882" box="[151,565,1879,1903]" pageId="21" pageNumber="72" type="materials_examined">
<paragraph id="F3105708C636202CFF3AF8C2FDF7F882" blockId="21.[151,771,1316,2068]" box="[151,565,1879,1903]" pageId="21" pageNumber="72">
<materialsCitation id="43C75D55C636202CFF3AF8C2FDF7F882" box="[151,565,1879,1903]" collectionCode="MGL" pageId="21" pageNumber="72" specimenCode="MGL110297" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC636202CFF3AF8C2FF38F882" bold="true" box="[151,250,1879,1903]" italics="true" pageId="21" pageNumber="72">
<typeStatus id="2C14E9AAC636202CFF3AF8C2FF38F882" box="[151,250,1879,1903]" pageId="21" pageNumber="72" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC636202CFEA5F8C2FEA3F882" box="[264,353,1879,1903]" captionStart="Fig" captionStartId="22.[197,232,1750,1771]" captionTargetBox="[212,1397,209,1690]" captionTargetId="figure-160@22.[197,1410,178,1721]" captionTargetPageId="22" captionText="Fig. 10. a Humerocyrtis infinita n. sp, Rc4. b, d Humerocyrtis brevithorax n. sp.: b R78/1b, d poorly preserved specimen, Rc4. c Humerocyrtis asymmetrica n. sp., Rc4. f Humerocyrtis marmoladensis n. sp., BV 85-70. e, g Humerocyrtis mediotriassica n. sp.: e CR25, g Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="21" pageNumber="72">Fig. 10a</figureCitation>
, coll.
<specimenCode id="A309FF73C636202CFE0FF8C2FDF3F882" box="[418,561,1879,1903]" collectionCode="MGL" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="21" pageNumber="72">MGL110297</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFF3AF8EDFE82F83E" pageId="21" pageNumber="72" type="description">
<paragraph id="F3105708C636202CFF3AF8EDFE82F83E" blockId="21.[151,771,1316,2068]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFF3AF8EDFED8F87D" bold="true" box="[151,282,1912,1936]" italics="true" pageId="21" pageNumber="72">Dimensions</emphasis>
. Length of shell without horns 88 µm, of thorax 55 µm, diameter of shoulder 96 µm, of its distal opening 74 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFF3AF84EFBDDFF5B" pageId="21" pageNumber="72" type="etymology">
<paragraph id="F3105708C636202CFF3AF84EFBDDFF5B" blockId="21.[151,771,1316,2068]" lastBlockId="21.[816,1437,158,513]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFF3AF84EFECEF81E" bold="true" box="[151,268,2011,2035]" italics="true" pageId="21" pageNumber="72">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC636202CFE51F84EFD91F81E" box="[508,595,2011,2035]" italics="true" pageId="21" pageNumber="72">infinitus</emphasis>
unfinished, because this species has no concave post shoulder part with flare-shaped end.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFC9DFF2AFBCCFDEC" pageId="21" pageNumber="72" type="discussion">
<paragraph id="F3105708C636202CFC9DFF2AFC5DFE70" blockId="21.[816,1437,158,513]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFC9DFF2AFC50FF3A" bold="true" box="[816,914,191,215]" italics="true" pageId="21" pageNumber="72">Remarks</emphasis>
.
<taxonomicName id="34AF2C8BC636202CFC19FF55FB6CFF3A" authorityName="Dumitrică" authorityYear="2024" box="[948,1198,191,215]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="21" pageNumber="72" phylum="Radiozoa" rank="species" species="infinita" status="sp. nov.">
<emphasis id="C1DB8B1AC636202CFC19FF55FB6CFF3A" box="[948,1198,191,215]" italics="true" pageId="21" pageNumber="72">Humerocyrtis infinita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C636202CFB67FF55FAD5FF3A" box="[1226,1303,192,215]" pageId="21" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
resembles structurally
<taxonomicName id="34AF2C8BC636202CFC15FF74FB9FFF15" authorityName="Dumitrică" authorityYear="2024" box="[952,1117,224,248]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="21" pageNumber="72" phylum="Radiozoa" rank="species" species="brevithorax" status="sp. nov.">
<emphasis id="C1DB8B1AC636202CFC15FF74FB9FFF15" box="[952,1117,224,248]" italics="true" pageId="21" pageNumber="72">H. brevithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C636202CFBC7FF74FB69FF15" box="[1130,1195,225,248]" pageId="21" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
by having the post-shoulder shell very short consisting of the narowing portion, but differs from it by missing the circumferential ring of the shoulder and by having a well-marked distal opening showing no possibility of developing a flare distal end.
</paragraph>
<paragraph id="F3105708C636202CFC9DFE32FBCCFDEC" blockId="21.[816,1437,158,513]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFC9DFE32FBD2FE52" bold="true" box="[816,1040,423,447]" italics="true" pageId="21" pageNumber="72">Stratigraphic range</emphasis>
. Very rare in the lower Ladinian sample Rc4 from the Buchenstein Formation from Recoaro, north
<collectingCountry id="8BB81798C636202CFC7BFE7CFBC8FDEC" box="[982,1034,489,513]" name="Italy" pageId="21" pageNumber="72">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,152 @@
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<treatment id="7B06E61EC636202FFC0AF880FACFF7EA" LSID="urn:lsid:plazi:treatment:7B06E61EC636202FFC0AF880FACFF7EA" httpUri="http://treatment.plazi.org/id/7B06E61EC636202FFC0AF880FACFF7EA" lastPageId="22" lastPageNumber="73" pageId="21" pageNumber="72">
<subSubSection id="BBB50483C636202CFC0AF880FAE4F8C0" box="[935,1318,1813,1837]" pageId="21" pageNumber="72" type="nomenclature">
<paragraph id="F3105708C636202CFC0AF880FAE4F8C0" blockId="21.[935,1318,1813,1837]" box="[935,1318,1813,1837]" pageId="21" pageNumber="72">
<heading id="A858E064C636202CFC0AF880FAE4F8C0" box="[935,1318,1813,1837]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="72" reason="5">
<emphasis id="C1DB8B1AC636202CFC0AF880FAE4F8C0" bold="true" box="[935,1318,1813,1837]" pageId="21" pageNumber="72">
<taxonomicName id="34AF2C8BC636202CFC0AF880FB26F8C0" authorityName="Dumitrică" authorityYear="2024" box="[935,1252,1813,1837]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="21" pageNumber="72" phylum="Radiozoa" rank="species" species="mediotriassica" status="sp. nov.">
<emphasis id="C1DB8B1AC636202CFC0AF880FB26F8C0" bold="true" box="[935,1252,1813,1837]" italics="true" pageId="21" pageNumber="72">Humerocyrtis mediotriassica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C636202CFB47F880FAE4F8C0" box="[1258,1318,1813,1837]" pageId="21" pageNumber="72" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFB85F8A0FB66F8A3" box="[1064,1188,1845,1870]" pageId="21" pageNumber="72" type="description">
<paragraph id="F3105708C636202CFB85F8A0FB66F8A3" blockId="21.[1064,1188,1845,1870]" box="[1064,1188,1845,1870]" pageId="21" pageNumber="72">
<figureCitation id="6B944B8DC636202CFB85F8A0FB66F8A3" box="[1064,1188,1845,1870]" captionStart="Fig" captionStartId="22.[197,232,1750,1771]" captionTargetBox="[212,1397,209,1690]" captionTargetId="figure-160@22.[197,1410,178,1721]" captionTargetPageId="22" captionText="Fig. 10. a Humerocyrtis infinita n. sp, Rc4. b, d Humerocyrtis brevithorax n. sp.: b R78/1b, d poorly preserved specimen, Rc4. c Humerocyrtis asymmetrica n. sp., Rc4. f Humerocyrtis marmoladensis n. sp., BV 85-70. e, g Humerocyrtis mediotriassica n. sp.: e CR25, g Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="21" pageNumber="72">Figs. 10e, g</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202FFC9DF8EDFB92F828" lastPageId="22" lastPageNumber="73" pageId="21" pageNumber="72" type="diagnosis">
<paragraph id="F3105708C636202FFC9DF8EDFB48F86E" blockId="21.[816,1436,1912,2068]" lastBlockId="22.[816,1436,1899,2056]" lastPageId="22" lastPageNumber="73" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFC9DF8EDFC5CF87D" bold="true" box="[816,926,1912,1936]" italics="true" pageId="21" pageNumber="72">Diagnosis</emphasis>
. Shell bell-shaped relatively large. Cephalis with sparse, small and irregularly disposed pores. Apical and ventral horns thin, conical, relatively long and equal in length, the former displaced toward the dorsal side of the cephalis and slightly curved towards the axis of shell. Collar boundary weakly marked outside but considered to be at the level of the median bar, which corresponds to the appearance of larger pores. Shoulder well marked by the first lateral enlargement of shell, followed by its contraction and again enlargement so that the outline of post-shoulder thorax is concave.
</paragraph>
<caption id="A7D00780C635202FFF68F943FC71F8CA" ID-DOI="http://doi.org/10.5281/zenodo.12582796" ID-Zenodo-Dep="12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="22" pageNumber="73" startId="22.[197,232,1750,1771]" targetBox="[212,1397,209,1690]" targetPageId="22" targetType="figure">
<paragraph id="F3105708C635202FFF68F943FC71F8CA" blockId="22.[197,1412,1749,1831]" pageId="22" pageNumber="73">
<emphasis id="C1DB8B1AC635202FFF68F943FECEF906" bold="true" box="[197,268,1750,1771]" pageId="22" pageNumber="73">Fig. 10</emphasis>
.
<emphasis id="C1DB8B1AC635202FFEB5F943FEE6F906" bold="true" box="[280,292,1750,1771]" pageId="22" pageNumber="73">a</emphasis>
<taxonomicName id="34AF2C8BC635202FFE93F943FDCFF906" authorityName="Dumitrică" authorityYear="2024" box="[318,525,1749,1771]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="22" pageNumber="73" phylum="Radiozoa" rank="species" species="infinita">
<emphasis id="C1DB8B1AC635202FFE93F943FDCFF906" box="[318,525,1749,1771]" italics="true" pageId="22" pageNumber="73">Humerocyrtis infinita</emphasis>
</taxonomicName>
n. sp, Rc4.
<emphasis id="C1DB8B1AC635202FFD2EF943FD52F906" bold="true" box="[643,656,1750,1771]" pageId="22" pageNumber="73">b</emphasis>
,
<emphasis id="C1DB8B1AC635202FFD30F943FD68F906" bold="true" box="[669,682,1750,1771]" pageId="22" pageNumber="73">d</emphasis>
<taxonomicName id="34AF2C8BC635202FFD69F943FC7CF906" authorityName="Dumitrică" authorityYear="2024" box="[708,958,1749,1771]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="22" pageNumber="73" phylum="Radiozoa" rank="species" species="brevithorax" status="sp. nov.">
<emphasis id="C1DB8B1AC635202FFD69F943FC7CF906" box="[708,958,1749,1771]" italics="true" pageId="22" pageNumber="73">Humerocyrtis brevithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C635202FFC68F943FC3EF906" box="[965,1020,1750,1771]" pageId="22" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="C1DB8B1AC635202FFBAAF943FBD6F906" bold="true" box="[1031,1044,1750,1771]" pageId="22" pageNumber="73">b</emphasis>
R78/1b,
<emphasis id="C1DB8B1AC635202FFB2CF943FB4CF906" bold="true" box="[1153,1166,1750,1771]" pageId="22" pageNumber="73">d</emphasis>
poorly preserved specimen, Rc4.
<emphasis id="C1DB8B1AC635202FFE98F961FE82F8E4" bold="true" box="[309,320,1780,1801]" pageId="22" pageNumber="73">c</emphasis>
<taxonomicName id="34AF2C8BC635202FFEF5F961FD9EF8E4" authorityName="Dumitrică" authorityYear="2024" box="[344,604,1780,1801]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="22" pageNumber="73" phylum="Radiozoa" rank="species" species="asymmetrica" status="sp. nov.">
<emphasis id="C1DB8B1AC635202FFEF5F961FD9EF8E4" box="[344,604,1780,1801]" italics="true" pageId="22" pageNumber="73">Humerocyrtis asymmetrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C635202FFDCEF961FD5AF8E4" box="[611,664,1780,1801]" pageId="22" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
, Rc4.
<emphasis id="C1DB8B1AC635202FFD78F961FD1FF8E4" bold="true" box="[725,733,1780,1801]" pageId="22" pageNumber="73">f</emphasis>
<taxonomicName id="34AF2C8BC635202FFD5BF961FBD1F8E4" authorityName="Dumitrică" authorityYear="2024" box="[758,1043,1779,1801]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="22" pageNumber="73" phylum="Radiozoa" rank="species" species="marmoladensis" status="sp. nov.">
<emphasis id="C1DB8B1AC635202FFD5BF961FBD1F8E4" box="[758,1043,1779,1801]" italics="true" pageId="22" pageNumber="73">Humerocyrtis marmoladensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C635202FFBB4F961FB8CF8E4" box="[1049,1102,1780,1801]" pageId="22" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
, BV 85-70.
<emphasis id="C1DB8B1AC635202FFB69F961FB0DF8E4" bold="true" box="[1220,1231,1780,1801]" pageId="22" pageNumber="73">e</emphasis>
, g
<taxonomicName id="34AF2C8BC635202FFAADF961FE90F8CA" authorityName="Dumitrică" authorityYear="2024" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="22" pageNumber="73" phylum="Radiozoa" rank="species" species="mediotriassica" status="sp. nov.">
<emphasis id="C1DB8B1AC635202FFAADF961FE90F8CA" italics="true" pageId="22" pageNumber="73">Humerocyrtis mediotriassica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C635202FFEF4F887FE4DF8CA" box="[345,399,1810,1831]" pageId="22" pageNumber="73" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="C1DB8B1AC635202FFE34F884FE66F8CB" bold="true" box="[409,420,1809,1830]" pageId="22" pageNumber="73">e</emphasis>
CR25,
<emphasis id="C1DB8B1AC635202FFE52F884FDC9F8CB" bold="true" box="[511,523,1809,1830]" pageId="22" pageNumber="73">g</emphasis>
Rc4. Scale bar of 50 µm is for all figures.
</paragraph>
</caption>
<paragraph id="F3105708C635202FFC9DF819FB92F828" blockId="22.[816,1436,1899,2056]" pageId="22" pageNumber="73">
<emphasis id="C1DB8B1AC635202FFC9DF819FC2AF849" bold="true" box="[816,1000,1932,1956]" italics="true" pageId="22" pageNumber="73">Studied material</emphasis>
.
<specimenCount id="E5A99C81C635202FFC5AF818FB61F849" box="[1015,1187,1932,1956]" count="2" pageId="22" pageNumber="73" type="generic">Two specimens</specimenCount>
, one in the sample CR 25 and another one in Rc4.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C635202FFC9DF85AFACFF7EA" pageId="22" pageNumber="73" type="materials_examined">
<paragraph id="F3105708C635202FFC9DF85AFAE5F80B" blockId="22.[816,1436,1899,2056]" box="[816,1319,1998,2023]" pageId="22" pageNumber="73">
<materialsCitation id="43C75D55C635202FFC9DF85AFAE5F80B" box="[816,1319,1998,2023]" collectionCode="MGL" pageId="22" pageNumber="73" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC635202FFC9DF85AFC51F80A" bold="true" box="[816,915,1999,2023]" italics="true" pageId="22" pageNumber="73">
<typeStatus id="2C14E9AAC635202FFC9DF85AFC51F80A" box="[816,915,1999,2023]" pageId="22" pageNumber="73" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC635202FFC0CF85BFC39F80B" box="[929,1019,1998,2022]" captionStart="Fig" captionStartId="22.[197,232,1750,1771]" captionTargetBox="[212,1397,209,1690]" captionTargetId="figure-160@22.[197,1410,178,1721]" captionTargetPageId="22" captionText="Fig. 10. a Humerocyrtis infinita n. sp, Rc4. b, d Humerocyrtis brevithorax n. sp.: b R78/1b, d poorly preserved specimen, Rc4. c Humerocyrtis asymmetrica n. sp., Rc4. f Humerocyrtis marmoladensis n. sp., BV 85-70. e, g Humerocyrtis mediotriassica n. sp.: e CR25, g Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="22" pageNumber="73">Fig. 10e</figureCitation>
, CR 25, coll.
<collectionCode id="95BECFCDC635202FFB20F85BFB0FF80B" box="[1165,1229,1998,2022]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="22" pageNumber="73">MGL</collectionCode>
.110299.
</materialsCitation>
</paragraph>
<paragraph id="F3105708C635202FFC9DF865FACFF7EA" blockId="22.[816,1436,1899,2056]" box="[816,1293,2031,2056]" pageId="22" pageNumber="73">
<materialsCitation id="43C75D55C635202FFC9DF865FB70F7EA" box="[816,1202,2031,2056]" collectionCode="MGL" pageId="22" pageNumber="73" specimenCount="1" typeStatus="paratype">
<emphasis id="C1DB8B1AC635202FFC9DF865FC50F7E5" bold="true" box="[816,914,2032,2056]" italics="true" pageId="22" pageNumber="73">
<typeStatus id="2C14E9AAC635202FFC9DF865FC50F7E5" box="[816,914,2032,2056]" pageId="22" pageNumber="73" type="paratype">Paratype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC635202FFC0DF87AFC38F7EA" box="[928,1018,2031,2055]" captionStart="Fig" captionStartId="22.[197,232,1750,1771]" captionTargetBox="[212,1397,209,1690]" captionTargetId="figure-160@22.[197,1410,178,1721]" captionTargetPageId="22" captionText="Fig. 10. a Humerocyrtis infinita n. sp, Rc4. b, d Humerocyrtis brevithorax n. sp.: b R78/1b, d poorly preserved specimen, Rc4. c Humerocyrtis asymmetrica n. sp., Rc4. f Humerocyrtis marmoladensis n. sp., BV 85-70. e, g Humerocyrtis mediotriassica n. sp.: e CR25, g Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="22" pageNumber="73">
Fig.
<quantity id="3457FAEDC635202FFC7DF87AFC38F7EA" box="[976,1018,2031,2055]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="22" pageNumber="73" unit="g" value="10.0">10g</quantity>
</figureCitation>
, Rc4, coll.
<collectionCode id="95BECFCDC635202FFBDEF87AFB70F7EA" box="[1139,1202,2031,2055]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="22" pageNumber="73">MGL</collectionCode>
</materialsCitation>
.110300.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,132 @@
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<mods:title id="6A5510D43E46DE253A5F89EA12034175">New Middle Triassic Bell-Shaped Nassellarian Radiolaria From Alpine And Carpathian Areas</mods:title>
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<mods:classification id="8B7F1B6F998CF3940F53E647762F6B94">journal article</mods:classification>
<mods:identifier id="83BFE9E264C7F4FCFA20D6D76CB5749A" type="DOI">10.35463/j.apr.2024.01.05</mods:identifier>
<mods:identifier id="4625054E622081E9866741516D3AA75B" type="ISSN">1842-371x</mods:identifier>
<mods:identifier id="66677EAA027B6A4501FE6D84209C3D66" type="Zenodo-Dep">12582776</mods:identifier>
</mods:mods>
<treatment id="7B06E61EC637202CFC1BF8CDFE6CFB98" LSID="urn:lsid:plazi:treatment:7B06E61EC637202CFC1BF8CDFE6CFB98" httpUri="http://treatment.plazi.org/id/7B06E61EC637202CFC1BF8CDFE6CFB98" lastPageId="21" lastPageNumber="72" pageId="20" pageNumber="71">
<subSubSection id="BBB50483C637202DFC1BF8CDFAD4F89D" box="[950,1302,1880,1904]" pageId="20" pageNumber="71" type="nomenclature">
<paragraph id="F3105708C637202DFC1BF8CDFAD4F89D" blockId="20.[950,1302,1880,1904]" box="[950,1302,1880,1904]" pageId="20" pageNumber="71">
<heading id="A858E064C637202DFC1BF8CDFAD4F89D" box="[950,1302,1880,1904]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="71" reason="5">
<emphasis id="C1DB8B1AC637202DFC1BF8CDFAD4F89D" bold="true" box="[950,1302,1880,1904]" pageId="20" pageNumber="71">
<taxonomicName id="34AF2C8BC637202DFC1BF8CDFB16F89D" authorityName="Dumitrică" authorityYear="2024" box="[950,1236,1880,1904]" class="Polycystina" family="Oertlispongidae" genus="Humerocyrtis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Spumellaria" pageId="20" pageNumber="71" phylum="Radiozoa" rank="species" species="brevithorax" status="sp. nov.">
<emphasis id="C1DB8B1AC637202DFC1BF8CDFB16F89D" bold="true" box="[950,1236,1880,1904]" italics="true" pageId="20" pageNumber="71">Humerocyrtis brevithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DAE83661C637202DFB77F8CDFAD4F89D" box="[1242,1302,1880,1904]" pageId="20" pageNumber="71" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C637202DFC5AF8EDFB17F87D" box="[1015,1237,1912,1936]" pageId="20" pageNumber="71" type="description">
<paragraph id="F3105708C637202DFC5AF8EDFB17F87D" blockId="20.[1015,1237,1912,1936]" box="[1015,1237,1912,1936]" pageId="20" pageNumber="71">
<figureCitation id="6B944B8DC637202DFC5AF8EDFBA3F87D" box="[1015,1121,1912,1936]" captionStart="Fig" captionStartId="20.[197,232,1730,1751]" captionTargetBox="[206,1394,188,1577]" captionTargetId="figure-156@20.[197,1410,178,1721]" captionTargetPageId="20" captionText="Fig. 9. a, b, f. Humerocyrtis gracilis n. sp.: a. paratype, Rc4; b. holotype, BV 85-70, f. Rc4. c. Humerocyrtis avirostrum n. sp., Rc4; d, d1, e. Humerocyrtis brevithorax n. sp., CRH: d, d1. same specimen in lateral and basal view, respectively, showing the initial spicule without D ray, e. holotype. Scale bar of 50 µm is for all figures, except fig. 7c for which the scale bar is of 75 µm." figureDoi="http://doi.org/10.5281/zenodo.12582794" httpUri="https://zenodo.org/record/12582794/files/figure.png" pageId="20" pageNumber="71">Fig. 9d, d</figureCitation>
<figureCitation id="6B944B8DC637202DFBCFF8EDFBAFF87D" box="[1122,1133,1912,1936]" captionStart="Fig" captionStartId="1.[199,234,1402,1423]" captionTargetBox="[197,1410,160,1373]" captionTargetId="figure-425@1.[197,1410,160,1373]" captionTargetPageId="1" captionText="Fig. 1. Geological map of the locality Cristian area, Braşov Mountains, Romania, modified from the geological maps Zărneşti and Braşov sheets, scale 1:50000, Săndulescu et al. (1972)." figureDoi="http://doi.org/10.5281/zenodo.12582778" httpUri="https://zenodo.org/record/12582778/files/figure.png" pageId="20" pageNumber="71">1</figureCitation>
, e;
<figureCitation id="6B944B8DC637202DFB3FF8EDFB17F87D" box="[1170,1237,1912,1936]" captionStart="Fig" captionStartId="22.[197,232,1750,1771]" captionTargetBox="[212,1397,209,1690]" captionTargetId="figure-160@22.[197,1410,178,1721]" captionTargetPageId="22" captionText="Fig. 10. a Humerocyrtis infinita n. sp, Rc4. b, d Humerocyrtis brevithorax n. sp.: b R78/1b, d poorly preserved specimen, Rc4. c Humerocyrtis asymmetrica n. sp., Rc4. f Humerocyrtis marmoladensis n. sp., BV 85-70. e, g Humerocyrtis mediotriassica n. sp.: e CR25, g Rc4. Scale bar of 50 µm is for all figures." figureDoi="http://doi.org/10.5281/zenodo.12582796" httpUri="https://zenodo.org/record/12582796/files/figure.png" pageId="20" pageNumber="71">10b, d</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C637202CFC9DF82EFD75FD2A" lastPageId="21" lastPageNumber="72" pageId="20" pageNumber="71" type="diagnosis">
<paragraph id="F3105708C637202CFC9DF82EFF37FD89" blockId="20.[816,1436,1978,2070]" lastBlockId="21.[151,772,158,1141]" lastPageId="21" lastPageNumber="72" pageId="20" pageNumber="71">
<emphasis id="C1DB8B1AC637202DFC9DF82EFC5CF83E" bold="true" box="[816,926,1979,2003]" italics="true" pageId="20" pageNumber="71">Diagnosis</emphasis>
. Shell dicyrtid and very short consisting of cephalis, the shoulder plus a short portion of its narrowing distal segment. Cephalis with sparse small pores of various shapes bearing a massive and slightly curved conical apical horn and a straight, three-bladed, and well-developed ventral horn. Thorax short, represented especially by its shoulder and a short narrowing distal part. Proximal part wide and short conical bearing large circular or oval pores separated by narrow and irregularly disposed intervening bars. Circumferential ring separating the two parts of the thorax is marked either by a protruding ring, an imperforate zone, or by no special structure. Distal part of thorax is thinner-walled, very short, narrowing very fast, and its pores are smaller and irregular in size, shape and arrangement. Its distal end is usually frayed, as if not finished.
</paragraph>
<paragraph id="F3105708C636202CFF3AFDF8FD75FD2A" blockId="21.[151,772,158,1141]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFF3AFDF8FE96FD68" bold="true" box="[151,340,621,645]" italics="true" pageId="21" pageNumber="72">Studied material</emphasis>
.
<materialsCitation id="43C75D55C636202CFEC4FDFBFD71FD2A" collectionCode="R" location="Pelsonian" municipality="lower Ladinian" pageId="21" pageNumber="72" specimenCode="R78" specimenCount="5">
Four or
<specimenCount id="E5A99C81C636202CFE63FDF8FDBAFD68" box="[462,632,621,645]" count="5" pageId="21" pageNumber="72" type="generic">five specimens</specimenCount>
, two in the
<location id="F67001D3C636202CFF3AFD1BFEC0FD4B" LSID="urn:lsid:plazi:treatment:7B06E61EC637202CFC1BF8CDFE6CFB98:F67001D3C636202CFF3AFD1BFEC0FD4B" box="[151,258,654,678]" municipality="lower Ladinian" name="Pelsonian" pageId="21" pageNumber="72">Pelsonian</location>
, one in the
<collectingMunicipality id="1374CD72C636202CFE2CFD1BFDE6FD4B" box="[385,548,654,678]" pageId="21" pageNumber="72">lower Ladinian</collectingMunicipality>
from sample
<specimenCode id="A309FF73C636202CFD16FD1BFD2EFD4B" box="[699,748,654,678]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="72">R78</specimenCode>
/1, and one (poorly preserved), or two, in sample Rc4
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFF3AFD44FD16FD05" box="[151,724,720,745]" pageId="21" pageNumber="72" type="materials_examined">
<paragraph id="F3105708C636202CFF3AFD44FD16FD05" blockId="21.[151,772,158,1141]" box="[151,724,720,745]" pageId="21" pageNumber="72">
<materialsCitation id="43C75D55C636202CFF3AFD44FD16FD05" box="[151,724,720,745]" collectionCode="CRH, MGL" collectorName="Cristian" pageId="21" pageNumber="72" specimenCount="1" typeStatus="holotype">
<emphasis id="C1DB8B1AC636202CFF3AFD44FF38FD04" bold="true" box="[151,250,721,745]" italics="true" pageId="21" pageNumber="72">
<typeStatus id="2C14E9AAC636202CFF3AFD44FF38FD04" box="[151,250,721,745]" pageId="21" pageNumber="72" type="holotype">Holotype</typeStatus>
</emphasis>
.
<figureCitation id="6B944B8DC636202CFEA5FD45FE96FD05" box="[264,340,720,744]" captionStart="Fig" captionStartId="20.[197,232,1730,1751]" captionTargetBox="[206,1394,188,1577]" captionTargetId="figure-156@20.[197,1410,178,1721]" captionTargetPageId="20" captionText="Fig. 9. a, b, f. Humerocyrtis gracilis n. sp.: a. paratype, Rc4; b. holotype, BV 85-70, f. Rc4. c. Humerocyrtis avirostrum n. sp., Rc4; d, d1, e. Humerocyrtis brevithorax n. sp., CRH: d, d1. same specimen in lateral and basal view, respectively, showing the initial spicule without D ray, e. holotype. Scale bar of 50 µm is for all figures, except fig. 7c for which the scale bar is of 75 µm." figureDoi="http://doi.org/10.5281/zenodo.12582794" httpUri="https://zenodo.org/record/12582794/files/figure.png" pageId="21" pageNumber="72">Fig. 9e</figureCitation>
,
<collectionCode id="95BECFCDC636202CFEF2FD45FE5AFD05" box="[351,408,720,744]" country="Zaire" httpUri="http://grbio.org/cool/m541-ph33" name="Centre de Recherche en Hydrobiologie" pageId="21" pageNumber="72">CRH</collectionCode>
,
<collectorName id="5E5A32DEC636202CFE0EFD45FE38FD05" box="[419,506,720,744]" pageId="21" pageNumber="72">Cristian</collectorName>
, coll.
<collectionCode id="95BECFCDC636202CFD97FD45FDBBFD05" box="[570,633,720,744]" country="Switzerland" httpUri="http://grbio.org/cool/jpxv-qjfr" name="Musee Geologique de Lausanne" pageId="21" pageNumber="72">MGL</collectionCode>
.110296.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFF3AFD67FD1FFCC6" pageId="21" pageNumber="72" type="description">
<paragraph id="F3105708C636202CFF3AFD67FD1FFCC6" blockId="21.[151,772,158,1141]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFF3AFD67FED8FCE7" bold="true" box="[151,282,754,778]" italics="true" pageId="21" pageNumber="72">Dimensions</emphasis>
. Maximum length of shell without apical horn 80 µm, maximum diameter of thorax 97-112 µm.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFF3AFCA1FE98FC80" pageId="21" pageNumber="72" type="etymology">
<paragraph id="F3105708C636202CFF3AFCA1FE98FC80" blockId="21.[151,772,158,1141]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFF3AFCA1FECEFCA1" bold="true" box="[151,268,820,844]" italics="true" pageId="21" pageNumber="72">Etymology</emphasis>
. From the Latin
<emphasis id="C1DB8B1AC636202CFE68FCA6FDC4FCA6" box="[453,518,819,843]" italics="true" pageId="21" pageNumber="72">brevis</emphasis>
short and
<emphasis id="C1DB8B1AC636202CFD22FCA6FD17FCA6" box="[655,725,819,843]" italics="true" pageId="21" pageNumber="72">thorax</emphasis>
due to its short thorax.
</paragraph>
</subSubSection>
<subSubSection id="BBB50483C636202CFF3AFCE3FE6CFB98" pageId="21" pageNumber="72" type="discussion">
<paragraph id="F3105708C636202CFF3AFCE3FEE2FC1C" blockId="21.[151,772,158,1141]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFF3AFCE3FF3BFC63" bold="true" box="[151,249,886,910]" italics="true" pageId="21" pageNumber="72">Remarks</emphasis>
. This species differs from the other species of this genus by having a very short and narrowing distal part of thorax, a conical apical horn and a three-bladed ventral horn.
</paragraph>
<paragraph id="F3105708C636202CFF3AFC6FFE6CFB98" blockId="21.[151,772,158,1141]" pageId="21" pageNumber="72">
<emphasis id="C1DB8B1AC636202CFF3AFC6FFEBBFBFF" bold="true" box="[151,377,1018,1042]" italics="true" pageId="21" pageNumber="72">Stratigraphic range</emphasis>
. Samples CRH, R78/1 and Rc4, middle Anisian (Pelsonian) to lower Ladinian (Fassanian) of Eastern Carpathians and Buchenstein Formation, Recoaro area, North
<collectingCountry id="8BB81798C636202CFED8FBC8FE6BFB98" box="[373,425,1117,1141]" name="Italy" pageId="21" pageNumber="72">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,104 +1,104 @@
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Millettia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Millettia vasta" order="Fabales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="vasta">Millettia vasta</taxonomicName>
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Kosterm., Reinwardtia 5: 349 (1960). Type: Indonesia, Kalimantan [Borneo], Belajan River near Muara Lempong, June 1956, [
<normalizedToken originalValue="André">Andre</normalizedToken>
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Joseph Guillaume Henri]
<emphasis italics="true" pageId="29" pageNumber="30">Kostermans 12516A</emphasis>
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, BO-1249898 (BO, holo.!); BM000997327
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BM, iso.!); L0018805 (L, iso.!); K000880991 (K, iso.!); P03081895 (P, iso.!) ≡
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(Kosterm.) Schot, Blumea 39(1-2): 36 (1994).
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<paragraph pageId="30" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="30" pageNumber="31">Borneo: Brunei; Indonesia (Kalimantan); Malaysia (Sabah, Sarawak).</paragraph>
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<paragraph id="C89435343971A7D1DCA170E65C1A40A1" pageId="30" pageNumber="31">Borneo: Brunei; Indonesia (Kalimantan); Malaysia (Sabah, Sarawak).</paragraph>
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<paragraph pageId="30" pageNumber="31">Habitat.</paragraph>
<paragraph pageId="30" pageNumber="31">Component tree in woods and forests from sea level to 250 m.</paragraph>
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<paragraph id="094D806B28E0483CF9F540BCE4FF359D" pageId="30" pageNumber="31">Habitat.</paragraph>
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<subSubSection id="BBE1B0E7240CFF8F764A62DDF18BFC6B" box="[809,1175,951,976]" pageId="14" pageNumber="231" type="nomenclature">
<paragraph id="F344E36C240CFF8F764A62DDF18BFC6B" blockId="14.[809,1175,951,1008]" box="[809,1175,951,976]" pageId="14" pageNumber="231">
<heading id="A80C5400240CFF8F764A62DDF18BFC6B" bold="true" box="[809,1175,951,976]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="231" reason="2">
<taxonomicName id="34FB98EF240CFF8F764A62DDF18BFC6B" authority="Chassain, 2005" authorityName="Chassain" authorityYear="2005" box="[809,1175,951,976]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="231" phylum="Arthropoda" rank="species" species="foucarti">
<emphasis id="C18F3F7E240CFF8F764A62DDF18BFC6B" bold="true" box="[809,1175,951,976]" pageId="14" pageNumber="231">
<emphasis id="C18F3F7E240CFF8F764A62DDF6F6FC74" bold="true" box="[809,1002,951,975]" italics="true" pageId="14" pageNumber="231">Teslasena foucarti</emphasis>
<bibRefCitation id="976A9E9D240CFF8F769262D2F18BFC6B" author="CHASSAIN, J." box="[1009,1175,952,976]" pageId="14" pageNumber="231" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7240CFF8F76CB62B2F104FC4B" box="[936,1048,984,1008]" pageId="14" pageNumber="231" type="description">
<paragraph id="F344E36C240CFF8F76CB62B2F104FC4B" blockId="14.[809,1175,951,1008]" box="[936,1048,984,1008]" pageId="14" pageNumber="231">
<emphasis id="C18F3F7E240CFF8F76CB62B2F104FC4B" bold="true" box="[936,1048,984,1008]" pageId="14" pageNumber="231">
(
<figureCitation id="6BC0FFE9240CFF8F76CC62B2F10CFC4B" box="[943,1040,984,1008]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="14" pageNumber="231">Fig. 34C</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7240CFF8F77AD6572F153FB8B" box="[718,1103,1048,1072]" pageId="14" pageNumber="231" type="reference_group">
<paragraph id="F344E36C240CFF8F77AD6572F153FB8B" blockId="14.[718,1103,1048,1072]" box="[718,1103,1048,1072]" pageId="14" pageNumber="231">
<treatmentCitationGroup id="D3EBC442240CFF8F77AD6572F153FB8B" box="[718,1103,1048,1072]" pageId="14" pageNumber="231">
<treatmentCitation id="725AC57D240CFF8F77AD6572F157FB8B" author="CHASSAIN, J." box="[718,1099,1048,1072]" page="66" pageId="14" pageNumber="231" year="2005">
<taxonomicName id="34FB98EF240CFF8F77AD6572F157FB8B" authority="Chassain, 2005: 66" authorityName="Chassain" authorityPageNumber="66" authorityYear="2005" box="[718,1099,1048,1072]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="231" phylum="Arthropoda" rank="species" species="foucarti">
<emphasis id="C18F3F7E240CFF8F77AD6572F666FB8B" box="[718,890,1048,1072]" italics="true" pageId="14" pageNumber="231">Teslasena foucarti</emphasis>
<bibRefCitation id="976A9E9D240CFF8F76E26572F157FB8B" author="CHASSAIN, J." box="[897,1099,1048,1072]" pageId="14" pageNumber="231" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005: 66</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7240CFF8E77AD6532F5C8FC4B" lastPageId="15" lastPageNumber="232" pageId="14" pageNumber="231" type="description">
<paragraph id="F344E36C240CFF8F77AD6532F6EDFA6B" blockId="14.[718,1267,1112,1488]" pageId="14" pageNumber="231">
<emphasis id="C18F3F7E240CFF8F77AD6532F64CFBCB" box="[718,848,1112,1136]" italics="true" pageId="14" pageNumber="231">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9240CFF8F76FC6532F6E3FBCB" box="[927,1023,1112,1136]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="14" pageNumber="231">Fig. 34C</figureCitation>
): Integument brown to black, legs lighter. Total length 9.0-12.0 mm; elytra 2.3-2.4 times longer than pronotum. Head concave or with a median circular depression, frontal carina straight between antennal insertions; IEP 0.25-0.35 antenna reaching or surpassing the hind angle of pronotum by one antennomere. Pronotum 1.05-1.10x wider than long, with double punctation on disc, larger punctures 0.5-1.0 diameters apart, denser on lateral borders; densely pilose with long decumbent setae. Scutellum covered with short setae. Elytra tapered apicad from humerus.
</paragraph>
<paragraph id="F344E36C240CFF8F77AD6492F1EFF92B" blockId="14.[718,1267,1528,1680]" pageId="14" pageNumber="231">
<emphasis id="C18F3F7E240CFF8F77AD6492F632F9AB" box="[718,814,1528,1552]" italics="true" pageId="14" pageNumber="231">Aedeagus:</emphasis>
Phallobase 0.31x the total length of aedeagus, 1.10x longer than wide, length ratio between lateroposterior and median parts 4.3; penis: basal triangular part with sides straight, apical part 0.71x as long as the triangular basal part; with a ventral sclerite.
</paragraph>
<paragraph id="F344E36C240CFF8F77AD67D2F1A0F8EB" blockId="14.[718,1267,1720,1872]" pageId="14" pageNumber="231">
<materialsCitation id="4393E931240CFF8F77AD67D2F1A0F8EB" collectingDate="2001-12-15" collectionCode="MNHN" collectorName="Chapada dos Parecis &amp; A. Foucart" country="Brazil" determinerName="J. Chassain" latitude="-14.2641115" location="Usine Alcomat" longLatPrecision="1" longitude="-59.233902" pageId="14" pageNumber="231" specimenCount="3" stateProvince="Mato Grosso" typeStatus="holotype">
<emphasis id="C18F3F7E240CFF8F77AD67D2F637F96B" box="[718,811,1720,1744]" italics="true" pageId="14" pageNumber="231">
<typeStatus id="2C405DCE240CFF8F77AD67D2F63AF96B" box="[718,806,1720,1744]" pageId="14" pageNumber="231" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC240CFF8F762067D2F69EF96B" box="[835,898,1720,1744]" name="Brazil" pageId="14" pageNumber="231">Brésil</collectingCountry>
, (
<collectingRegion id="313F2D8E240CFF8F76C367D2F12AF96B" box="[928,1078,1720,1744]" country="Brazil" name="Mato Grosso" pageId="14" pageNumber="231">Mato Grosso</collectingRegion>
),
<geoCoordinate id="96CF85AB240CFF8F713767D2F1F2F96B" box="[1108,1262,1720,1744]" degrees="14" direction="south" minutes="15" orientation="latitude" pageId="14" pageNumber="231" precision="1" seconds="50.80" value="-14.2641115">14°1550.80”S</geoCoordinate>
,
<geoCoordinate id="96CF85AB240CFF8F77AD67B2F66EF94B" box="[718,882,1752,1776]" degrees="59" direction="west" minutes="14" orientation="longitude" pageId="14" pageNumber="231" precision="1" seconds="02.05" value="-59.233902">59°1402.05”W</geoCoordinate>
,
<collectorName id="5E0E86BA240CFF8F76E067B2F17BF94B" box="[899,1127,1752,1776]" pageId="14" pageNumber="231">Chapada dos Parecis</collectorName>
], [
<locationDeviation id="E2F6FA4E240CFF8F71E767B2F658F8AB" location="d'Uirapuru" pageId="14" pageNumber="231">
<quantity id="34034E89240CFF8F71E767B2F1CCF94B" box="[1156,1232,1752,1776]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="231" unit="km" value="30.0">30 km</quantity>
N dUirapuru
</locationDeviation>
,
<location id="F624B5B7240CFF8F76376792F6EAF8AB" LSID="urn:lsid:plazi:treatment:7B52527A240CFF8E764A62DDF5C8FC4B:F624B5B7240CFF8F76376792F6EAF8AB" box="[852,1014,1784,1808]" country="Brazil" latitude="-14.2641115" longLatPrecision="1" longitude="-59.233902" name="Usine Alcomat" pageId="14" pageNumber="231" stateProvince="Mato Grosso">Usine Alcomat</location>
, 1 au
<date id="8745C5AC240CFF8F71276792F1D7F8AB" box="[1092,1227,1784,1808]" pageId="14" pageNumber="231" value="2001-12-15">
<collectingDate id="97013C44240CFF8F71276792F1D7F8AB" box="[1092,1227,1784,1808]" pageId="14" pageNumber="231" value="2001-12-15">15-XII-2001</collectingDate>
</date>
,
<collectorName id="5E0E86BA240CFF8F71B86792F601F894" pageId="14" pageNumber="231">A. Foucart</collectorName>
leg.], [
<typeStatus id="2C405DCE240CFF8F76066672F6D2F88B" box="[869,974,1816,1840]" pageId="14" pageNumber="231" type="holotype">Holotype</typeStatus>
], [
<taxonomicName id="34FB98EF240CFF8F768B6672F189F88B" authorityName="Rosa" authorityYear="2014" box="[1000,1173,1816,1840]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="231" phylum="Arthropoda" rank="species" species="foucarti" status="sp. nov.">
<emphasis id="C18F3F7E240CFF8F768B6672F189F88B" box="[1000,1173,1816,1840]" italics="true" pageId="14" pageNumber="231">Teslasena foucarti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205240CFF8F71FE6672F1F3F894" box="[1181,1263,1816,1839]" pageId="14" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
,
<typeStatus id="2C405DCE240CFF8F77AD6652F62EF8EB" box="[718,818,1848,1872]" pageId="14" pageNumber="231" type="holotype">Holotype</typeStatus>
,
<determinerName id="06980FA8240CFF8F765E6652F6B3F8EB" box="[829,943,1848,1872]" pageId="14" pageNumber="231">J. Chassain</determinerName>
det. 04], male (
<collectionCode id="95EA7BA9240CFF8F71376652F1ACF8EB" box="[1108,1200,1848,1872]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="231">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C240DFF8E753D61D3F41BFF4B" blockId="15.[94,642,184,240]" pageId="15" pageNumber="232">
<materialsCitation id="4393E931240DFF8E753D61D3F41FFF4B" collectingDate="2001-12-15" collectionCode="MNHN, MZUSP" collectorName="Chapada dos Parecis &amp; A. Foucart" country="Brazil" determinerName="J. Chassain" latitude="-14.2641115" location="Usine Alcomat" longLatPrecision="1" longitude="-59.233902" pageId="15" pageNumber="232" specimenCount="5" stateProvince="Mato Grosso" typeStatus="paratype">
<emphasis id="C18F3F7E240DFF8E753D61D3F5DCFF6B" box="[94,192,185,208]" italics="true" pageId="15" pageNumber="232">
<typeStatus id="2C405DCE240DFF8E753D61D3F5A6FF6B" box="[94,186,185,208]" pageId="15" pageNumber="232" type="paratype">Paratypes</typeStatus>
:
</emphasis>
same data as holotype,
<specimenCount id="E5FD28E5240DFF8E74A961D2F715FF74" box="[458,521,184,208]" count="4" pageId="15" pageNumber="232" type="generic">4 exs.</specimenCount>
(
<collectionCode id="95EA7BA9240DFF8E777861D2F76BFF6B" box="[539,631,184,208]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="232">MNHN</collectionCode>
),
<specimenCount id="E5FD28E5240DFF8E753D61B2F58CFF54" box="[94,144,216,240]" count="1" pageId="15" pageNumber="232" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9240DFF8E75FE61B2F5E0FF4B" box="[157,252,216,240]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="15" pageNumber="232">MZUSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C240DFF8E753D6072F5B0FEEB" blockId="15.[94,642,280,336]" pageId="15" pageNumber="232">
<emphasis id="C18F3F7E240DFF8E753D6072F5FDFE8B" box="[94,225,280,304]" italics="true" pageId="15" pageNumber="232">Distribution:</emphasis>
<collectingCountry id="8BECA3FC240DFF8E758D6072F451FE8B" box="[238,333,280,304]" name="Brazil" pageId="15" pageNumber="232">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E240DFF8E743D6073F4FBFE94" box="[350,487,280,304]" italics="true" pageId="15" pageNumber="232">
<collectingRegion id="313F2D8E240DFF8E743D6073F4FEFE94" box="[350,482,280,304]" country="Brazil" name="Mato Grosso" pageId="15" pageNumber="232">Mato Grosso</collectingRegion>
:
</emphasis>
Chapada dos Parecis.
</paragraph>
<paragraph id="F344E36C240DFF8E753D6012F5C8FC4B" blockId="15.[94,643,376,1008]" pageId="15" pageNumber="232">
<emphasis id="C18F3F7E240DFF8E753D6012F439FE2B" box="[94,293,376,400]" italics="true" pageId="15" pageNumber="232">
Remarks:
<taxonomicName id="34FB98EF240DFF8E75DD6013F439FE2B" authorityName="Rosa" authorityYear="2014" box="[190,293,376,400]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="foucarti">T. foucarti</taxonomicName>
</emphasis>
(
<figureCitation id="6BC0FFE9240DFF8E74506012F48CFE2B" box="[307,400,376,400]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="15" pageNumber="232">Fig. 34C</figureCitation>
) was detailed described by
<bibRefCitation id="976A9E9D240DFF8E75E160F2F42EFE0B" author="CHASSAIN, J." box="[130,306,408,432]" pageId="15" pageNumber="232" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain (2005)</bibRefCitation>
, who distinguished this species from
<taxonomicName id="34FB98EF240DFF8E75FA60D3F411FE6B" authorityName="J. Chassain" baseAuthorityName="Lucas" baseAuthorityYear="1857" box="[153,269,440,464]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="C18F3F7E240DFF8E75FA60D3F411FE6B" box="[153,269,440,464]" italics="true" pageId="15" pageNumber="232">T. femoralis</emphasis>
</taxonomicName>
in the shape of the anterior margin of frons, which is straight in
<emphasis id="C18F3F7E240DFF8E74ED60B3F4E2FE4B" box="[398,510,472,496]" italics="true" pageId="15" pageNumber="232">
<taxonomicName id="34FB98EF240DFF8E74ED60B3F4E7FE4B" authorityName="Rosa" authorityYear="2014" box="[398,507,472,496]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="foucarti">T. foucarti</taxonomicName>
,
</emphasis>
its antennomeres relatively longer, the pronotum narrower anteriorly with punctation larger on lateral and anterior borders and the metatibial inner spur with tip curved. As discussed above, the analysis of a large series of
<taxonomicName id="34FB98EF240DFF8E753D6313F5CFFD2B" authorityName="J. Chassain" baseAuthorityName="Lucas" baseAuthorityYear="1857" box="[94,211,632,656]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="C18F3F7E240DFF8E753D6313F5CFFD2B" box="[94,211,632,656]" italics="true" pageId="15" pageNumber="232">T. femoralis</emphasis>
</taxonomicName>
revealed that those features, with the exception of the curved metatibial spur, are highly variable among the specimens and do not constitute consistent diagnostic characters. On the other hand, the curved tip of the metatibial inner spur was observed only in
<taxonomicName id="34FB98EF240DFF8E75DE6273F437FC8B" authorityName="Rosa" authorityYear="2014" box="[189,299,792,816]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="foucarti">
<emphasis id="C18F3F7E240DFF8E75DE6273F437FC8B" box="[189,299,792,816]" italics="true" pageId="15" pageNumber="232">T. foucarti</emphasis>
</taxonomicName>
specimens, which also present body pilosity longer. The most evident difference in the pilosity of these species was observed in the pronotum, which is moderately long and usually dense on
<taxonomicName id="34FB98EF240DFF8E75E162F3F5F6FC0B" authorityName="Rosa" authorityYear="2014" box="[130,234,920,944]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="foucarti">
<emphasis id="C18F3F7E240DFF8E75E162F3F5F6FC0B" box="[130,234,920,944]" italics="true" pageId="15" pageNumber="232">T. foucarti</emphasis>
</taxonomicName>
and absent or minuscule and scarce in
<emphasis id="C18F3F7E240DFF8E753D62D3F5C5FC6B" box="[94,217,952,976]" italics="true" pageId="15" pageNumber="232">
<taxonomicName id="34FB98EF240DFF8E753D62D3F5C9FC6B" authorityName="J. Chassain" baseAuthorityName="Lucas" baseAuthorityYear="1857" box="[94,213,952,976]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="232" phylum="Arthropoda" rank="species" species="femoralis">T. femoralis</taxonomicName>
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Their aedeagus do not present significant differences.
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species
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(male): Frontal carina absent, mandibles long, bidentate directed anteriorly; pronotum quadrate, weakly convex, sides widened anteriorly, anterior angles right not produced, posterior angle without carina; scutellum cordiform wider than long, convex and perpendicular to mesoscutum at base; elytra subparallel to posterior third, not fused to each other, wings absent; legs narrow, tibia and tarsomeres 1-2 with spiniform setae, tibial spurs robust, longer than setae, tarsomeres and claws simple.
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<emphasis id="C18F3F7E2410FF9375A663BDF4ACFD4B" bold="true" box="[197,432,727,752]" italics="true" pageId="18" pageNumber="235">Oligostethius capensis</emphasis>
(
<bibRefCitation id="976A9E9D2410FF9374DE63B2F77CFD4B" author="SCHWARZ, O." box="[445,608,728,752]" pageId="18" pageNumber="235" pagination="357 - 376" refId="ref51087" refString="SCHWARZ, O. 1903. Neue Elateriden aus Africa und Madagaskar. Deutsche Entomologische Zeitschrift, 1903: 357 - 376." type="journal article" year="1903">Schwarz, 1903</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72410FF9374236392F4F1FCAB" box="[320,493,760,784]" pageId="18" pageNumber="235" type="description">
<paragraph id="F344E36C2410FF9374236392F4F1FCAB" blockId="18.[197,616,727,784]" box="[320,493,760,784]" pageId="18" pageNumber="235">
<emphasis id="C18F3F7E2410FF9374236392F4F1FCAB" bold="true" box="[320,493,760,784]" pageId="18" pageNumber="235">
(
<figureCitation id="6BC0FFE92410FF9374246392F48DFCAB" box="[327,401,760,784]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Figs. 5</figureCitation>
,
<figureCitation id="6BC0FFE92410FF9374FE6392F4B0FCAB" box="[413,428,760,784]" captionStart="FIGURE 6" captionStartId="19.[95,174,723,742]" captionTargetBox="[95,1225,172,698]" captionTargetId="figure-538@19.[94,1228,170,699]" captionTargetPageId="19" captionText="FIGURE 6: Oligostethius capensis. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, G, aedeagus (dorsal, ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="18" pageNumber="235">6</figureCitation>
,
<figureCitation id="6BC0FFE92410FF9374DB6392F4FAFCAB" box="[440,486,760,784]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="18" pageNumber="235">34E</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72410FF9375E76252F46CFC2B" pageId="18" pageNumber="235" type="reference_group">
<paragraph id="F344E36C2410FF9375E76252F712FCEB" blockId="18.[132,681,824,912]" box="[132,526,824,848]" pageId="18" pageNumber="235">
<treatmentCitationGroup id="D3EBC4422410FF9375E76252F712FCEB" box="[132,526,824,848]" pageId="18" pageNumber="235">
<treatmentCitation id="725AC57D2410FF9375E76252F716FCEB" author="SCHWARZ, O." box="[132,522,824,848]" page="375" pageId="18" pageNumber="235" year="1903">
<taxonomicName id="34FB98EF2410FF9375E76252F716FCEB" authority="Schwarz 1903: 375" authorityName="Schwarz" authorityPageNumber="375" authorityYear="1903" box="[132,522,824,848]" class="Insecta" family="Elateridae" genus="Idiotropia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="235" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="C18F3F7E2410FF9375E76252F426FCEB" box="[132,314,824,848]" italics="true" pageId="18" pageNumber="235">Idiotropia capensis</emphasis>
<bibRefCitation id="976A9E9D2410FF9374226252F716FCEB" author="SCHWARZ, O." box="[321,522,824,848]" pageId="18" pageNumber="235" pagination="357 - 376" refId="ref51087" refString="SCHWARZ, O. 1903. Neue Elateriden aus Africa und Madagaskar. Deutsche Entomologische Zeitschrift, 1903: 357 - 376." type="journal article" year="1903">Schwarz 1903: 375</bibRefCitation>
</taxonomicName>
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.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F344E36C2410FF9375E76232F46CFC2B" blockId="18.[132,681,824,912]" pageId="18" pageNumber="235">
<treatmentCitationGroup id="D3EBC4422410FF9375E76232F46CFC2B" pageId="18" pageNumber="235">
<emphasis id="C18F3F7E2410FF9375E76232F478FCCB" box="[132,356,856,880]" italics="true" pageId="18" pageNumber="235">
<taxonomicName id="34FB98EF2410FF9375E76232F442FCCB" baseAuthorityName="Schwarz" baseAuthorityYear="1903" box="[132,350,856,880]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="235" phylum="Arthropoda" rank="species" species="capensis">Oligostethius capensis</taxonomicName>
;
</emphasis>
<treatmentCitation id="725AC57D2410FF9374126232F753FCCB" author="SCHWARZ, O." box="[369,591,856,880]" page="315" pageId="18" pageNumber="235" year="1906">
<bibRefCitation id="976A9E9D2410FF9374126232F753FCCB" author="SCHWARZ, O." box="[369,591,856,880]" pageId="18" pageNumber="235" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 315</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2410FF9377016232F470FC2B" author="SCHENKLING, S." page="510" pageId="18" pageNumber="235" year="1927">
<bibRefCitation id="976A9E9D2410FF9377016232F470FC2B" author="SCHENKLING, S." pageId="18" pageNumber="235" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 510</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72410FF9575E762D2F436F8EB" lastPageId="20" lastPageNumber="237" pageId="18" pageNumber="235" type="description">
<paragraph id="F344E36C2410FF9375E762D2F436FBD4" blockId="18.[132,681,952,1136]" pageId="18" pageNumber="235">
<emphasis id="C18F3F7E2410FF9375E762D2F41AFC6B" box="[132,262,952,976]" italics="true" pageId="18" pageNumber="235">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE92410FF93743462D2F4A8FC6B" box="[343,436,952,976]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="18" pageNumber="235">Fig. 34E</figureCitation>
): Body convex. Integument brown or dark brown with antennae and legs light brown, covered with very short, fine, decumbent silvery setae. Total length: 9.6-10.0 mm; elytral base 0.83x as wide as prothorax, elytra 1.91x times longer than pronotum.
</paragraph>
<paragraph id="F344E36C2410FF93766461D2F65EFCD4" blockId="18.[718,1267,184,1136]" pageId="18" pageNumber="235">
Head (
<figureCitation id="6BC0FFE92410FF93763161D2F6BAFF6B" box="[850,934,184,208]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Fig. 5C</figureCitation>
) with frons (
<figureCitation id="6BC0FFE92410FF93715861D2F192FF6B" box="[1083,1166,184,208]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Fig. 5A</figureCitation>
) concave at median line, frontal carina absent medially; frontoclypeal region gradually declivous to base of labrum; punctures umbilicate, 0.5-1.0 diameter apart; antennal insertion (
<figureCitation id="6BC0FFE92410FF9376F56052F116FEEB" box="[918,1034,312,336]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Figs. 5A, B</figureCitation>
) placed within saucerlike impression. Antenna with antennomere II cylindrical 1.3x longer than wide; antennomeres III-VI convex and serrate; III 1.7x longer than II and 1.6x wider than long; IV-VI subequal 0.92x as long as III and 1.1x wider than long. IEP 0.24; mouthparts directed ventrally. Labrum (
<figureCitation id="6BC0FFE92410FF9376B56092F138FDAB" box="[982,1060,504,528]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Fig. 5A</figureCitation>
) semielliptical, convex. Mandible (
<figureCitation id="6BC0FFE92410FF9376126372F6DDFD8B" box="[881,961,536,560]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Fig. 5A</figureCitation>
) narrow and long, bidentate, apical tooth long and acute, subapical tooth smaller; laterodorsal surface at base coarsely punctate with long setae, lateral edge evenly curved apicad; mesal margin at base with a row of short setae. Maxilla with galea oboval, densely pilose, lacinia elliptical, densely pilose; medistipes trapezoidal longer than wide with several long stiff setae and a few finer and shorter setae. Labial palpigers separate; mentum trapezoidal with a lateroposterior pair of long setae and sparsely short setae.
</paragraph>
<paragraph id="F344E36C2410FF9276646212F4DCFB74" blockId="18.[718,1267,184,1136]" lastBlockId="19.[94,643,824,1232]" lastPageId="19" lastPageNumber="236" pageId="18" pageNumber="235">
Pronotum (
<figureCitation id="6BC0FFE92410FF93761F6212F6D0FC2B" box="[892,972,888,912]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Fig. 5D</figureCitation>
) dorsally convex, 1.17-1.19x longer than wide, sides rounded, lateral carina complete (
<figureCitation id="6BC0FFE92410FF93766D62D2F647FC6B" box="[782,859,952,976]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Fig. 5F</figureCitation>
), with punctures umbilicate, coalescent forming irregular striae; anterior angle produced anteriorly, wide and rounded covering half eye (
<figureCitation id="6BC0FFE92410FF9371FA6292F1F9FBAB" box="[1177,1253,1016,1040]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235">Fig. 5F</figureCitation>
); posterior angle acute, short, with tip upturned (
<figureCitation id="6BC0FFE92410FF9377B66552F63FFBEB" box="[725,803,1080,1104]" captionStart="FIGURE 3" captionStartId="16.[132,211,1800,1819]" captionTargetBox="[136,1263,685,1777]" captionTargetId="figure-249@16.[132,1266,681,1777]" captionTargetPageId="16" captionText="FIGURE 3: Idiotropia henoni (male): A, antenna; B, C, head (dorsal, anterodorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, mesocoxal cavity (lateroventral); I, mesoventrite (lateral); J, pterothorax (dorsal) and abdominal tergites I-II; K, mesoscutum and scutellar shield (lateral); L, pterothorax (ventral), M, N, O, pro-, meso- and metathoracic legs (outer surface). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12641373" httpUri="https://zenodo.org/record/12641373/files/figure.png" pageId="18" pageNumber="235">Fig. 3F</figureCitation>
), carinate; posterior margin adjacent to posterior angle with a short incision contiguous to a carina. Hypomeron (
<figureCitation id="6BC0FFE92411FF9274736252F47CFCEB" box="[272,352,824,848]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Fig. 5G</figureCitation>
) with punctures umbilicate, 0.5-1.0 diameter apart; posterior margin straight and contiguous to tip of posterior angle, without notch. Notosternal suture (
<figureCitation id="6BC0FFE92411FF92744B62F2F468FC0B" box="[296,372,920,944]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Fig. 5E</figureCitation>
) margined by a shiny band along hypomeron margin at posterior half, grooved anteriorly. Prosternum (
<figureCitation id="6BC0FFE92411FF92743262B2F480FC4B" box="[337,412,984,1008]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Fig. 5E</figureCitation>
) parallel-sided on posterior half, with divergent sides on anterior half, 2.31x longer than wide, with punctures umbilicate, 0.5-1.0 diameters apart; anterior prosternal lobe rounded and produced, covering mentum. Prosternal process (
<figureCitation id="6BC0FFE92411FF9275056512F5CAFB2B" box="[102,214,1144,1168]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Figs. 5E, F</figureCitation>
) with ventral surface laterally compressed and ascending at about 45° to apex, 2.8x longer than procoxal diameter. Procoxae open.
</paragraph>
<caption id="A784B3E42410FF9375E76662F77AF8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641377" ID-Zenodo-Dep="12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="18" pageNumber="235" startId="18.[132,211,1800,1819]" targetBox="[133,1265,1196,1773]" targetPageId="18" targetType="figure">
<paragraph id="F344E36C2410FF9375E76662F77AF8F4" blockId="18.[132,1267,1800,1871]" pageId="18" pageNumber="235">
<emphasis id="C18F3F7E2410FF9375E76662F5F6F8A7" bold="true" box="[132,234,1800,1820]" pageId="18" pageNumber="235">FIGURE 5:</emphasis>
<emphasis id="C18F3F7E2410FF93758C6663F482F8A0" box="[239,414,1801,1820]" italics="true" pageId="18" pageNumber="235">
<taxonomicName id="34FB98EF2410FF93758C6663F485F8A0" baseAuthorityName="Schwarz" baseAuthorityYear="1903" box="[239,409,1801,1820]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="235" phylum="Arthropoda" rank="species" species="capensis">Oligostethius capensis</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2410FF9374C26662F4FAF8A0" bold="true" box="[417,486,1800,1820]" pageId="18" pageNumber="235">A, B, C,</emphasis>
head (anterodorsal, lateral, dorsal);
<emphasis id="C18F3F7E2410FF93766E6663F653F8A7" bold="true" box="[781,847,1801,1820]" pageId="18" pageNumber="235">D, E, F,</emphasis>
prothorax (dorsal, ventral, lateral);
<emphasis id="C18F3F7E2410FF9371106662F195F8A7" bold="true" box="[1139,1161,1800,1820]" pageId="18" pageNumber="235">G,</emphasis>
hypomeron;
<emphasis id="C18F3F7E2410FF9375E76648F5B2F88E" bold="true" box="[132,174,1826,1845]" pageId="18" pageNumber="235">H, I,</emphasis>
pterothorax (ventral, lateroventral);
<emphasis id="C18F3F7E2410FF9374BF6648F4F6F88E" bold="true" box="[476,490,1826,1845]" pageId="18" pageNumber="235">J,</emphasis>
mesoscutum and scutellar shield;
<emphasis id="C18F3F7E2410FF9376666648F62DF88E" bold="true" box="[773,817,1826,1845]" pageId="18" pageNumber="235">K, L,</emphasis>
abdomen (ventral, lateral). Scale = 1.0 mm.
<emphasis id="C18F3F7E2410FF9371C66648F5B2F8F5" italics="true" pageId="18" pageNumber="235">Abbreviations:</emphasis>
llp: length of lateral part, lmp length of median part.
</paragraph>
</caption>
<caption id="A784B3E42411FF92753C63B9F582FCBB" ID-DOI="http://doi.org/10.5281/zenodo.12641379" ID-Zenodo-Dep="12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="19" pageNumber="236" startId="19.[95,174,723,742]" targetBox="[95,1225,172,698]" targetPageId="19" targetType="figure">
<paragraph id="F344E36C2411FF92753C63B9F582FCBB" blockId="19.[95,1229,723,768]" pageId="19" pageNumber="236">
<emphasis id="C18F3F7E2411FF92753C63B9F5D9FD5D" bold="true" box="[95,197,723,742]" pageId="19" pageNumber="236">FIGURE 6:</emphasis>
<emphasis id="C18F3F7E2411FF9275A963B9F465FD5D" box="[202,377,723,742]" italics="true" pageId="19" pageNumber="236">
<taxonomicName id="34FB98EF2411FF9275A963B9F468FD5D" baseAuthorityName="Schwarz" baseAuthorityYear="1903" box="[202,372,723,742]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="236" phylum="Arthropoda" rank="species" species="capensis">Oligostethius capensis</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2411FF92741E63B9F48CFD5D" bold="true" box="[381,400,723,742]" pageId="19" pageNumber="236">A,</emphasis>
sternite VIII;
<emphasis id="C18F3F7E2411FF92776563B9F705FD5D" bold="true" box="[518,537,723,742]" pageId="19" pageNumber="236">B,</emphasis>
tergite VIII;
<emphasis id="C18F3F7E2411FF9277E463B9F780FD5D" bold="true" box="[647,668,723,742]" pageId="19" pageNumber="236">C,</emphasis>
sternite IX;
<emphasis id="C18F3F7E2411FF92766063B9F606FD5D" bold="true" box="[771,794,723,742]" pageId="19" pageNumber="236">D,</emphasis>
tergites IX and X;
<emphasis id="C18F3F7E2411FF9276D463B9F6E5FD5D" bold="true" box="[951,1017,723,742]" pageId="19" pageNumber="236">E, F, G,</emphasis>
aedeagus (dorsal, ventral, lateral).
</paragraph>
</caption>
<paragraph id="F344E36C2411FF92753D6592F16AFC4B" blockId="19.[94,643,1272,1872]" lastBlockId="19.[680,1229,824,1872]" pageId="19" pageNumber="236">
<emphasis id="C18F3F7E2411FF92753D6592F5CAFAAB" box="[94,214,1272,1296]" italics="true" pageId="19" pageNumber="236">Pterothorax:</emphasis>
Mesoventrite gradually inclined ventrad posteriorly about 30°, anterior articulating surfaces (
<figureCitation id="6BC0FFE92411FF9275E06452F5E0FAEB" box="[131,252,1336,1360]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Figs. 5H, I</figureCitation>
) concave marginate posteriorly by a prominent carina; borders of mesoventral cavity divergent from base to anterior margin of mesocoxae then curved and convergent. Mesoventral cavity deep, floor with a smooth median band bordered by minuscule punctation (
<figureCitation id="6BC0FFE92411FF92746D64B2F47FFA4B" box="[270,355,1496,1520]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Fig. 5H</figureCitation>
). Mesocoxal cavity (
<figureCitation id="6BC0FFE92411FF92775764B2F767FA4B" box="[564,635,1496,1520]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Fig. 5I</figureCitation>
) open to mesepisternum and mesepimeron, mesepimeron long, closing the major part of cavity; mesotrocantin visible; mesepisternum with a carina on the outer angle and a prominent carina on inner anterior angle contiguous to the anterior carina of mesoventrite. Mesoventrite separated from the metaventrite at middle by a weak groove. Metaventrite (
<figureCitation id="6BC0FFE92411FF92774567D2F766F96B" box="[550,634,1720,1744]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Fig. 5H</figureCitation>
) 1.91x wider than long, 0.9x as long as mesoventrite, with punctation 0.5-1.0 diameter apart; metepisternum about 7x longer than wide. Elytra fused to each other along the median suture, sides divergent from humerus to posterior third then rounded to apex; striae with a row of punctures, interstices flat with dense, rasp-like punctation, epipleura abruptly narrowed near metacoxae. Scutellum (
<figureCitation id="6BC0FFE92411FF9276B662F2F103FC0B" box="[981,1055,920,944]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Fig. 5J</figureCitation>
) elliptical wider than long, strongly convex, abruptly elevated above the level of mesoscutum. Hind wings absent.
</paragraph>
<paragraph id="F344E36C2411FF9277826292F14DFACB" blockId="19.[680,1229,824,1872]" pageId="19" pageNumber="236">
Metacoxa (
<figureCitation id="6BC0FFE92411FF9276376292F6D4FBAB" box="[852,968,1016,1040]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Figs. 5H, I</figureCitation>
) inclined 25° in relation to transverse axis of body, median region of the dorsal surface about 15x longer than outer region; free margin of metacoxal plate very small. Legs densely pilose, setae moderately long and curved apically. Metatrochanter and metafemur compressed laterally, metafemur 3.5x longer than wide. Tibiae compressed laterally, 2x wider at apex than at base, with two subequal spurs shorter than setae, with a row of 10-13 stiff setae along apical border; tarsomeres simple, densely setose on ventral face, decreasing in length from I-IV, V as long as III and IV together; claws simple.
</paragraph>
<paragraph id="F344E36C2411FF9577826412F781FE8B" blockId="19.[680,1229,824,1872]" lastBlockId="20.[132,681,184,304]" lastPageId="20" lastPageNumber="237" pageId="19" pageNumber="236">
Abdominal ventrites (
<figureCitation id="6BC0FFE92411FF9276AB6412F159FA2B" box="[968,1093,1400,1424]" captionStart="FIGURE 5" captionStartId="18.[132,211,1800,1819]" captionTargetBox="[133,1265,1196,1773]" captionTargetId="figure-408@18.[132,1266,1191,1777]" captionTargetPageId="18" captionText="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi="http://doi.org/10.5281/zenodo.12641377" httpUri="https://zenodo.org/record/12641377/files/figure.png" pageId="19" pageNumber="236">Figs. 5K, L</figureCitation>
) with punctures umbilicate, 0.5-1.0 diameter apart, marginal plates absent; lateral part of ventrite 1 2.0x longer than its median part, ventrite 1 narrower than 3, 3-4 subequal, 5 semioval 1.12x wider than long. Sternite VIII (
<figureCitation id="6BC0FFE92411FF9277866772F629F98B" box="[741,821,1560,1584]" captionStart="FIGURE 6" captionStartId="19.[95,174,723,742]" captionTargetBox="[95,1225,172,698]" captionTargetId="figure-538@19.[94,1228,170,699]" captionTargetPageId="19" captionText="FIGURE 6: Oligostethius capensis. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, G, aedeagus (dorsal, ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="19" pageNumber="236">Fig. 6A</figureCitation>
) subrectangular 2.73x wider than long with posterior margin emarginate, predominantly membranous with a pair of dark sclerotizations along the anterior margin and a pair of lateroposterior light scletotizations; tergite VIII (
<figureCitation id="6BC0FFE92411FF9276AA67F2F10AF90B" box="[969,1046,1688,1712]" captionStart="FIGURE 6" captionStartId="19.[95,174,723,742]" captionTargetBox="[95,1225,172,698]" captionTargetId="figure-538@19.[94,1228,170,699]" captionTargetPageId="19" captionText="FIGURE 6: Oligostethius capensis. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, G, aedeagus (dorsal, ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="19" pageNumber="236">Fig. 6B</figureCitation>
) semioval, pilose, evenly sclerotized except for a membranous triangular region on anteromedian margin; sternite IX (
<figureCitation id="6BC0FFE92411FF92711567B2F1D8F94B" box="[1142,1220,1752,1776]" captionStart="FIGURE 6" captionStartId="19.[95,174,723,742]" captionTargetBox="[95,1225,172,698]" captionTargetId="figure-538@19.[94,1228,170,699]" captionTargetPageId="19" captionText="FIGURE 6: Oligostethius capensis. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, G, aedeagus (dorsal, ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="19" pageNumber="236">Fig. 6C</figureCitation>
) partly sclerotized, fused to tergite IX at half length of sternite IX, with anterior margin bilobed and produced dorsad; posterior margin rounded, apex with short and long setae; tergite IX (
<figureCitation id="6BC0FFE92416FF95749A61D2F74EFF6B" box="[505,594,184,208]" captionStart="FIGURE 6" captionStartId="19.[95,174,723,742]" captionTargetBox="[95,1225,172,698]" captionTargetId="figure-538@19.[94,1228,170,699]" captionTargetPageId="19" captionText="FIGURE 6: Oligostethius capensis. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, G, aedeagus (dorsal, ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="20" pageNumber="237">Fig. 6D</figureCitation>
) evenly sclerotized, posterior lobes densely covered with short setae and a few long setae apically; tergite X (
<figureCitation id="6BC0FFE92416FF95772C6192F7BCFEAB" box="[591,672,248,272]" captionStart="FIGURE 6" captionStartId="19.[95,174,723,742]" captionTargetBox="[95,1225,172,698]" captionTargetId="figure-538@19.[94,1228,170,699]" captionTargetPageId="19" captionText="FIGURE 6: Oligostethius capensis. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, G, aedeagus (dorsal, ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="20" pageNumber="237">Fig. 6D</figureCitation>
) semicircular with apex membranous, sparsely pilose.
</paragraph>
<paragraph id="F344E36C2416FF9575E76032F498FD6B" blockId="20.[132,681,344,720]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9575E76032F5C1FECB" box="[132,221,344,368]" italics="true" pageId="20" pageNumber="237">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92416FF95759F6032F44DFECB" box="[252,337,344,368]" captionStart="FIGURE 6" captionStartId="19.[95,174,723,742]" captionTargetBox="[95,1225,172,698]" captionTargetId="figure-538@19.[94,1228,170,699]" captionTargetPageId="19" captionText="FIGURE 6: Oligostethius capensis. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, G, aedeagus (dorsal, ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="20" pageNumber="237">Figs. 6</figureCitation>
E-G): Phallobase semicircular, strongly convex with a median dorsal apodeme, 0.3x total length of aedeagus; length ratio between lateroposterior and median parts 0.3. Parameres convex on anterior 1/3, dorsoventrally flattened on posterior 2/3 (
<figureCitation id="6BC0FFE92416FF9575D56092F414FDAB" box="[182,264,504,528]" captionStart="FIGURE 6" captionStartId="19.[95,174,723,742]" captionTargetBox="[95,1225,172,698]" captionTargetId="figure-538@19.[94,1228,170,699]" captionTargetPageId="19" captionText="FIGURE 6: Oligostethius capensis. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, G, aedeagus (dorsal, ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641379" httpUri="https://zenodo.org/record/12641379/files/figure.png" pageId="20" pageNumber="237">Fig. 6G</figureCitation>
); apex securiform short with outer angle rounded and produced, with a few setae on ventral face. Penis parallel-sided from base of basal struts to apex, basal struts 0.28x length of penis; dorsal articulation of penis with a short and narrow process between the basal struts fused to the parameres; ventral sclerite reduced to a line.
</paragraph>
<paragraph id="F344E36C2416FF9575E76393F457FC2B" blockId="20.[132,681,760,912]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9575E76393F5F9FCAB" box="[132,229,761,784]" italics="true" pageId="20" pageNumber="237">
<typeStatus id="2C405DCE2416FF9575E76393F5C3FCAB" box="[132,223,761,784]" pageId="20" pageNumber="237" type="lectotype">Lectotype</typeStatus>
:
</emphasis>
[Coll. Schwarz], [Cap Alte Sammlung], [
<taxonomicName id="34FB98EF2416FF9575EE6272F4D7FC8B" authority="Schw." authorityName="Schw." box="[141,459,792,816]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="species" species="capensins">
<emphasis id="C18F3F7E2416FF9575EE6272F46BFC8B" box="[141,375,792,816]" italics="true" pageId="20" pageNumber="237">Oligostethius capensins</emphasis>
Schw.
</taxonomicName>
], [Dtsch. Entomol. Institut Berlin], [
<typeStatus id="2C405DCE2416FF9574226252F4B2FCEB" box="[321,430,824,848]" pageId="20" pageNumber="237" type="lectotype">Lectotype</typeStatus>
], [
<taxonomicName id="34FB98EF2416FF9574AC6252F451FCCB" authority="Schwarz, C. Girard" authorityName="Schwarz, C. Girard" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="C18F3F7E2416FF9574AC6252F7B5FCEB" box="[463,681,824,848]" italics="true" pageId="20" pageNumber="237">Oligostethius capensis</emphasis>
Schwarz, C. Girard
</taxonomicName>
vid. 1979], [coll. DEI, Müncheberg], male, (DEI).
</paragraph>
<paragraph id="F344E36C2416FF9575E762D2F5ECFC4B" blockId="20.[132,681,952,1008]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9575E762D2F445FC6B" box="[132,345,952,976]" italics="true" pageId="20" pageNumber="237">Material examined:</emphasis>
<materialsCitation id="4393E9312416FF95741162D2F5F0FC4B" collectionCode="BMNH" country="South Africa" location="SOUTH AFRICA" pageId="20" pageNumber="237" specimenCount="1">
<collectingCountry id="8BECA3FC2416FF95741162D2F754FC6B" box="[370,584,952,976]" name="South Africa" pageId="20" pageNumber="237">SOUTH AFRICA</collectingCountry>
,
<specimenCount id="E5FD28E52416FF95770662D2F7B5FC74" box="[613,681,952,976]" count="1" pageId="20" pageNumber="237" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA92416FF9575EF62B2F5F8FC4B" box="[140,228,984,1008]" pageId="20" pageNumber="237">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C2416FF9575E76572F4C9FB8B" blockId="20.[132,469,1048,1072]" box="[132,469,1048,1072]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9575E76572F41AFB8B" box="[132,262,1048,1072]" italics="true" pageId="20" pageNumber="237">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2416FF95746E6572F4CDFB8B" box="[269,465,1048,1072]" name="South Africa" pageId="20" pageNumber="237">SOUTH AFRICA</collectingCountry>
.
</paragraph>
<paragraph id="F344E36C2416FF9575E76532F46EFAAB" blockId="20.[132,681,1112,1872]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9575E76532F5C2FBCB" box="[132,222,1112,1136]" italics="true" pageId="20" pageNumber="237">Remarks:</emphasis>
The specimen deposited in the MNHN collection has broken legs and antennae. The specimen of the BMNH collection has complete legs, incomplete antennae, broken elytra and its abdomen was lost. Last segment of maxillary and labial palpi was lost in both specimens.
</paragraph>
<paragraph id="F344E36C2416FF9575DE6472F436F8EB" blockId="20.[132,681,1112,1872]" pageId="20" pageNumber="237">
The data analyzed here support a close relationship of
<taxonomicName id="34FB98EF2416FF9575BF6452F4AEFAEB" box="[220,434,1336,1360]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="C18F3F7E2416FF9575BF6452F4AEFAEB" box="[220,434,1336,1360]" italics="true" pageId="20" pageNumber="237">Oligostethius capensis</emphasis>
</taxonomicName>
and
<emphasis id="C18F3F7E2416FF9574936452F449FACB" italics="true" pageId="20" pageNumber="237">
<taxonomicName id="34FB98EF2416FF9574936452F7BAFAEB" baseAuthorityName="Abeille de Perrin" baseAuthorityYear="1894" box="[496,678,1336,1360]" class="Insecta" family="Elateridae" genus="Idiotropia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="species" species="henoni">Idiotropia henoni</taxonomicName>
.
<taxonomicName id="34FB98EF2416FF9575E76432F449FACB" box="[132,341,1368,1392]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="species" species="capensis">Oligostethius capensis</taxonomicName>
</emphasis>
also shares some noticeable characters with the
<taxonomicName id="34FB98EF2416FF95745B6412F498FA2B" authorityName="Candeze" authorityYear="1863" box="[312,388,1400,1424]" class="Insecta" family="Elateridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="tribe" tribe="Dimini">Dimini</taxonomicName>
<emphasis id="C18F3F7E2416FF9574F16412F712FA2B" box="[402,526,1400,1424]" italics="true" pageId="20" pageNumber="237">
<taxonomicName id="34FB98EF2416FF9574F16412F715FA2B" authorityName="Golbach" authorityYear="1964" box="[402,521,1400,1424]" class="Insecta" family="Elateridae" genus="Apteroelater" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="genus">Apteroelater</taxonomicName>
:
</emphasis>
sutura mesometasternal grooved and elytral suture fused to each other. As discussed above,
<taxonomicName id="34FB98EF2416FF9574F764D2F70EFA6B" authorityName="Schwarz" authorityYear="1906" box="[404,530,1464,1488]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2416FF9574F764D2F70EFA6B" box="[404,530,1464,1488]" italics="true" pageId="20" pageNumber="237">Oligostethius</emphasis>
</taxonomicName>
and
<taxonomicName id="34FB98EF2416FF95772564D2F7B4FA6B" authorityName="Schwarz" authorityYear="1897" box="[582,680,1464,1488]" class="Insecta" family="Elateridae" genus="Idiotropia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2416FF95772564D2F7B4FA6B" box="[582,680,1464,1488]" italics="true" pageId="20" pageNumber="237">Idiotropia</emphasis>
</taxonomicName>
appear to be related to
<taxonomicName id="34FB98EF2416FF9574E364B2F4D0FA4B" authorityName="Candeze" authorityYear="1863" box="[384,460,1496,1520]" class="Insecta" family="Elateridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="tribe" tribe="Dimini">Dimini</taxonomicName>
species because they share the non-carinate frons, metacoxal plate reduced laterally and loss of hind wings.
<taxonomicName id="34FB98EF2416FF9574956772F768F98B" authorityName="Schwarz" authorityYear="1906" box="[502,628,1560,1584]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2416FF9574956772F768F98B" box="[502,628,1560,1584]" italics="true" pageId="20" pageNumber="237">Oligostethius</emphasis>
</taxonomicName>
also shares with species of the genera
<taxonomicName id="34FB98EF2416FF9574BD6752F438F9CB" authority="Schimmel &amp; Platia, 1992" authorityName="Schimmel &amp; Platia" authorityYear="1992" class="Insecta" family="Elateridae" genus="Neodima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2416FF9574BD6752F724F9EB" box="[478,568,1592,1616]" italics="true" pageId="20" pageNumber="237">Neodima</emphasis>
Schimmel &amp; Platia, 1992
</taxonomicName>
(Southeast Asia) and
<taxonomicName id="34FB98EF2416FF9577776732F5E3F92B" authority="Horn, 1891" authorityName="Horn" authorityYear="1891" class="Insecta" family="Elateridae" genus="Anthracopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2416FF9577776732F7B4F9CB" box="[532,680,1624,1648]" italics="true" pageId="20" pageNumber="237">Anthracopteryx</emphasis>
Horn, 1891
</taxonomicName>
(Cental and North America) the shape of scutellum (convex, wider than long) and the oblongovate elytra. It is distinguishable from those species by its longer prothorax, which is wider than elytra, pronotum with punctation dense and coalescent forming irregular striae and posterior margin with a pair of lateral incisions.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,137 @@
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<subSubSection id="BBE1B0E72415FF9675CD61DDF72EFF6B" box="[174,562,183,208]" pageId="23" pageNumber="240" type="nomenclature">
<paragraph id="F344E36C2415FF9675CD61DDF72EFF6B" blockId="23.[174,562,183,240]" box="[174,562,183,208]" pageId="23" pageNumber="240">
<heading id="A80C54002415FF9675CD61DDF72EFF6B" bold="true" box="[174,562,183,208]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="240" reason="2">
<taxonomicName id="34FB98EF2415FF9675CD61DDF72EFF6B" authority="Fleutiaux, 1940" authorityName="Fleutiaux" authorityYear="1940" box="[174,562,183,208]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="C18F3F7E2415FF9675CD61DDF72EFF6B" bold="true" box="[174,562,183,208]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF9675CD61DDF49EFF74" bold="true" box="[174,386,183,207]" italics="true" pageId="23" pageNumber="240">Toxognathus bakeri</emphasis>
Fleutiaux, 1940
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72415FF96747261B2F4CDFF4B" box="[273,465,216,240]" pageId="23" pageNumber="240" type="description">
<paragraph id="F344E36C2415FF96747261B2F4CDFF4B" blockId="23.[174,562,183,240]" box="[273,465,216,240]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF96747261B2F4CDFF4B" bold="true" box="[273,465,216,240]" pageId="23" pageNumber="240">
(
<figureCitation id="6BC0FFE92415FF96747B61B2F47CFF4B" box="[280,352,216,240]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="23" pageNumber="240">Figs. 7</figureCitation>
A-D; 34F)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72415FF96753D6072F70CFE8B" box="[94,528,280,304]" pageId="23" pageNumber="240" type="reference_group">
<paragraph id="F344E36C2415FF96753D6072F70CFE8B" blockId="23.[94,528,280,304]" box="[94,528,280,304]" pageId="23" pageNumber="240">
<treatmentCitationGroup id="D3EBC4422415FF96753D6072F70CFE8B" box="[94,528,280,304]" pageId="23" pageNumber="240">
<treatmentCitation id="725AC57D2415FF96753D6072F710FE8B" author="FLEUTIAUX, E." box="[94,524,280,304]" page="104" pageId="23" pageNumber="240" year="1940">
<taxonomicName id="34FB98EF2415FF96753D6072F710FE8B" authority="Fleutiaux, 1940 b: 104" authorityName="Fleutiaux" authorityPageNumber="104" authorityYear="1940" box="[94,524,280,304]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="C18F3F7E2415FF96753D6072F401FE8B" box="[94,285,280,304]" italics="true" pageId="23" pageNumber="240">Toxognathus bakeri</emphasis>
<bibRefCitation id="976A9E9D2415FF9674476072F710FE8B" author="FLEUTIAUX, E." box="[292,524,280,304]" pageId="23" pageNumber="240" pagination="89 - 104" refId="ref49954" refString="FLEUTIAUX, E. 1940 b. Elaterides nouveaux. Bulletin et Annales de La Societe Entomologique de Belgique, 80: 89 - 104." type="journal article" year="1940">Fleutiaux, 1940b: 104</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72415FF96753D6032F6E1FD14" pageId="23" pageNumber="240" type="description">
<paragraph id="F344E36C2415FF96753D6032F41BFCEB" blockId="23.[94,643,344,848]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF96753D6032F5FFFECB" box="[94,227,344,368]" italics="true" pageId="23" pageNumber="240">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE92415FF9674246032F4B2FECB" box="[327,430,344,368]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="23" pageNumber="240">Fig. 34F</figureCitation>
): Integument very bright, evenly light brown. Total length 6.0 mm; elytral base 1.0x as wide as pronotum, elytra 2.9x longer than pronotum. Frons flattened; frontal carina complete. Antennae surpassing the posterior angles of pronotum by one antennomere. Mandible unidentate. Pronotum (
<figureCitation id="6BC0FFE92415FF9674E66372F4C4FD8B" box="[389,472,536,560]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="23" pageNumber="240">Fig. 7A</figureCitation>
) weakly convex 1.17x wider than long with sides sinuous converging abruptly anteriad from anterior half, lateral carina prominent posteriorly, directed ventrad anteriorly, posterior angles divergent and convex; punctures 0.5-1.0 diameter apart. Elytral interstices slightly convex. Free margin of metacoxal plate (
<figureCitation id="6BC0FFE92415FF96774463B2F766FD4B" box="[551,634,728,752]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="23" pageNumber="240">Fig. 7B</figureCitation>
) short, triangular; tibiae slightly widened apicad, out- er angle of protibia truncate. Tergite IX with anterior margin straight.
</paragraph>
<paragraph id="F344E36C2415FF96753D6212F4C3FB8B" blockId="23.[94,643,888,1072]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF96753D6212F5ABFC2B" box="[94,183,888,912]" italics="true" pageId="23" pageNumber="240">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92415FF9675AB6212F454FC2B" box="[200,328,888,912]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="23" pageNumber="240">Figs. 7C, D</figureCitation>
): With phallobase 0.43x total length of aedeagus, 1.1x longer than wide, length ratio between lateroposterior and median parts 8.16; paramere with ventral anteromedian margins narrow, elongate and convergent; basal struts 0.33x total length of penis; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C2415FF9677CB61D2F604FEEB" blockId="23.[680,1229,184,336]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF9677CB61D2F619FF6B" box="[680,773,184,208]" italics="true" pageId="23" pageNumber="240">
<typeStatus id="2C405DCE2415FF9677CB61D2F61CFF6B" box="[680,768,184,208]" pageId="23" pageNumber="240" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Island of
<collectingRegion id="313F2D8E2415FF9676EA61D2F6CAFF6B" box="[905,982,184,208]" country="Malaysia" name="Pulau Pinang" pageId="23" pageNumber="240">Penang</collectingRegion>
, Baker], [
<taxonomicName id="34FB98EF2415FF96712661D2F629FF4B" authority="Fleut." authorityName="Fleutiaux" authorityYear="1940" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="bakeri">Toxognathus bakeri Fleut.</taxonomicName>
, type, Collection Fleutiaux], [Type], [Muséum Paris, Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF2415FF9671266192F678FE8B" authority="(Fleutiaux)" baseAuthorityName="Fleutiaux" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="bakeri">Toxognathus bakeri (Fleutiaux)</taxonomicName>
, J. Chassain det. 05], [
<typeStatus id="2C405DCE2415FF9671366072F1DEFE8B" box="[1109,1218,280,304]" pageId="23" pageNumber="240" type="lectotype">Lectotype</typeStatus>
], (MNHN).
</paragraph>
<paragraph id="F344E36C2415FF9677CB6012F1BBFE2B" blockId="23.[680,1191,376,400]" box="[680,1191,376,400]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF9677CB6012F636FE2B" box="[680,810,376,400]" italics="true" pageId="23" pageNumber="240">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2415FF9676526012F6AEFE2B" box="[817,946,376,400]" name="Malaysia" pageId="23" pageNumber="240">MALAYSIA</collectingCountry>
.
<emphasis id="C18F3F7E2415FF9676D86013F115FE2B" box="[955,1033,377,400]" italics="true" pageId="23" pageNumber="240">
<collectingRegion id="313F2D8E2415FF9676D86013F118FE2B" box="[955,1028,377,400]" country="Malaysia" name="Pulau Pinang" pageId="23" pageNumber="240">Penang</collectingRegion>
:
</emphasis>
<collectingRegion id="313F2D8E2415FF9671736012F146FE2B" box="[1040,1114,376,400]" country="Malaysia" name="Pulau Pinang" pageId="23" pageNumber="240">Penang</collectingRegion>
Island.
</paragraph>
<paragraph id="F344E36C2415FF9677CB60D2F6E1FD14" blockId="23.[680,1229,440,688]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF9677CB60D2F6D1FE6B" box="[680,973,440,464]" italics="true" pageId="23" pageNumber="240">
Remarks:
<taxonomicName id="34FB98EF2415FF96766860D2F6D1FE6B" box="[779,973,440,464]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="bakeri">Toxognathus bakeri</taxonomicName>
</emphasis>
is similar to
<taxonomicName id="34FB98EF2415FF96713C60D3F1D0FE6B" authorityName="Fleutiaux" authorityYear="1940" box="[1119,1228,440,464]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="dohertyi">
<emphasis id="C18F3F7E2415FF96713C60D3F1D0FE6B" box="[1119,1228,440,464]" italics="true" pageId="23" pageNumber="240">T. dohertyi</emphasis>
</taxonomicName>
mainly because of its light brown integument and the prominent lateral carina of pronotum, but differs from this species in its smaller size, lighter integument, flattened frons, pronotal punctures smaller and not coalescent and elytral interstices less convex. It shares with
<taxonomicName id="34FB98EF2415FF96767C6313F6B9FD2B" authorityName="Fleutiaux" authorityYear="1918" box="[799,933,632,656]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="beauchenei">
<emphasis id="C18F3F7E2415FF96767C6313F6B9FD2B" box="[799,933,632,656]" italics="true" pageId="23" pageNumber="240">T. beauchenei</emphasis>
</taxonomicName>
the produced ventral anteromedian margin of the parameres.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,196 @@
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<paragraph id="F344E36C2415FF9677BC639DF189FCAB" blockId="23.[735,1173,759,816]" box="[735,1173,759,784]" pageId="23" pageNumber="240">
<heading id="A80C54002415FF9677BC639DF189FCAB" bold="true" box="[735,1173,759,784]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="240" reason="2">
<taxonomicName id="34FB98EF2415FF9677BC639DF189FCAB" authority="Fleutiaux, 1918" authorityName="Fleutiaux" authorityYear="1918" box="[735,1173,759,784]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="beauchenei">
<emphasis id="C18F3F7E2415FF9677BC639DF189FCAB" bold="true" box="[735,1173,759,784]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF9677BC639DF6F8FCB4" bold="true" box="[735,996,759,783]" italics="true" pageId="23" pageNumber="240">Toxognathus beauchenei</emphasis>
Fleutiaux, 1918
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="F344E36C2415FF9676396272F107FC8B" blockId="23.[735,1173,759,816]" box="[858,1051,792,816]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF9676396272F107FC8B" bold="true" box="[858,1051,792,816]" pageId="23" pageNumber="240">
(
<figureCitation id="6BC0FFE92415FF9676026272F6B4FC8B" box="[865,936,792,816]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="23" pageNumber="240">Figs. 7</figureCitation>
E-K; 34G)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72415FF9677CB6232F68CFC0B" pageId="23" pageNumber="240" type="reference_group">
<paragraph id="F344E36C2415FF9677CB6232F68CFC0B" blockId="23.[680,1228,856,944]" pageId="23" pageNumber="240">
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<treatmentCitation id="725AC57D2415FF9677CB6232F196FCCB" author="FLEUTIAUX, E." box="[680,1162,856,880]" page="276" pageId="23" pageNumber="240" year="1918">
<taxonomicName id="34FB98EF2415FF9677CB6232F196FCCB" authority="Fleutiaux, 1918 a: 276" authorityName="Fleutiaux" authorityYear="1918" box="[680,1162,856,880]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="beauchenei">
<emphasis id="C18F3F7E2415FF9677CB6232F68BFCCB" box="[680,919,856,880]" italics="true" pageId="23" pageNumber="240">Toxognathus beauchenei</emphasis>
<bibRefCitation id="976A9E9D2415FF9676C36232F196FCCB" author="FLEUTIAUX, E." box="[928,1162,856,880]" pageId="23" pageNumber="240" pagination="175 - 278" refId="ref49758" refString="FLEUTIAUX, E. 1918 a. Nouvelles contributions a la faune de l'Indo-Chine Francaise. Annales de La Societe Entomologique de France, 87: 175 - 278." type="journal article" year="1918">Fleutiaux, 1918a: 276</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2415FF9671F46232F68CFC2B" author="FLEUTIAUX, E." page="178" pageId="23" pageNumber="240" year="1924">
<bibRefCitation id="976A9E9D2415FF9671F46232F68CFC2B" author="FLEUTIAUX, E." pageId="23" pageNumber="240" pagination="31 - 184" refId="ref49879" refString="FLEUTIAUX, E. 1924. Melasidae et Elateridae. Faune entomologique de l'Indochine francaise. 1. Coleopteres serricornes. 2. Fam. Elateridae. Opuscules de l'Institut Scientifique de l'Indochine, 2 (7): 31 - 184." type="journal article" year="1924">Fleutiaux, 1924: 178</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2415FF9676FF6212F190FC2B" author="SCHENKLING, S." box="[924,1164,888,912]" page="509" pageId="23" pageNumber="240" year="1927">
<bibRefCitation id="976A9E9D2415FF9676FF6212F190FC2B" author="SCHENKLING, S." box="[924,1164,888,912]" pageId="23" pageNumber="240" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 509</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2415FF9671F46212F691FC0B" author="FLEUTIAUX, E." page="35" pageId="23" pageNumber="240" year="1940">
<bibRefCitation id="976A9E9D2415FF9671F46212F691FC0B" author="FLEUTIAUX, E." pageId="23" pageNumber="240" pagination="19 - 40" refId="ref49922" refString="FLEUTIAUX, E. 1940 a. Les elaterides de l'Indo-chine francaise. Huitieme et derniere partie. Annales de La Societe Entomologique de France, 109: 19 - 40." type="journal article" year="1940">Fleutiaux, 1940a: 35</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72415FF9977CB62B2F1FDFD8B" lastPageId="24" lastPageNumber="241" pageId="23" pageNumber="240" type="description">
<paragraph id="F344E36C2415FF9977CB62B2F5F2FD2B" blockId="23.[680,1229,984,1072]" lastBlockId="24.[132,681,184,656]" lastPageId="24" lastPageNumber="241" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF9677CB62B2F636FC4B" box="[680,810,984,1008]" italics="true" pageId="23" pageNumber="240">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE92415FF96761A62B2F6C6FC4B" box="[889,986,984,1008]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="23" pageNumber="240">Fig. 34G</figureCitation>
): Integument dark reddish-brown, antennae evenly brown or with antennomeres I-V yellow, VI-X black and XI reddish-brown. Total length 9.0-
<quantity id="34034E89241AFF99745861D2F489FF74" box="[315,405,184,208]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="24" pageNumber="241" unit="mm" value="9.5">9.5 mm</quantity>
; elytral base 1.0-1.05x as wide as pronotum, elytra 2.60-2.66x times longer than pronotum. Frons deeply concave along the anterior margin; frontal carina complete. Antennae (
<figureCitation id="6BC0FFE9241AFF9975EF6052F5C6FEEB" box="[140,218,312,336]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="24" pageNumber="241">Fig. 7E</figureCitation>
) reaching the posterior angles of pronotum. Mandible unidentate. Pronotum weakly convex 1.19-1.22x wider than long with sides sinuous converging abruptly anteriad from anterior half, lateral carina directed ventrad anteriorly, posterior angles divergent and convex; punctures coalescent forming weak longitudinal striae. Elytral interstices costate. Free margin of metacoxal plate absent, tibia slightly widened apicad (
<figureCitation id="6BC0FFE9241AFF9974556352F461FDEB" box="[310,381,568,592]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="24" pageNumber="241">Figs. 7</figureCitation>
G-I), outer angle of protibia truncate. Tergite IX with anterior margin straight or bisinuous.
</paragraph>
<caption id="A784B3E42415FF9675036648F6AFF8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641381" ID-Zenodo-Dep="12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="23" pageNumber="240" startId="23.[96,175,1826,1845]" targetBox="[97,1227,1106,1794]" targetPageId="23" targetType="figure">
<paragraph id="F344E36C2415FF9675036648F6AFF8F4" blockId="23.[95,1229,1826,1871]" pageId="23" pageNumber="240">
<emphasis id="C18F3F7E2415FF9675036648F5DAF88E" bold="true" box="[96,198,1826,1845]" pageId="23" pageNumber="240">FIGURE 7:</emphasis>
<emphasis id="C18F3F7E2415FF9675AD6648F46DF88E" box="[206,369,1826,1845]" italics="true" pageId="23" pageNumber="240">
<taxonomicName id="34FB98EF2415FF9675AD6648F470F88E" box="[206,364,1826,1845]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="bakeri">Toxognathus bakeri</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2415FF9674146648F496F88E" bold="true" box="[375,394,1826,1845]" pageId="23" pageNumber="240">A,</emphasis>
pronotum;
<emphasis id="C18F3F7E2415FF9674906648F71AF88E" bold="true" box="[499,518,1826,1845]" pageId="23" pageNumber="240">B,</emphasis>
metaventrite;
<emphasis id="C18F3F7E2415FF9677E16648F7A9F88E" bold="true" box="[642,693,1826,1845]" pageId="23" pageNumber="240">C, D,</emphasis>
aedeagus (dorsal, ventral).
<emphasis id="C18F3F7E2415FF9676FD6648F179F88E" box="[926,1125,1826,1845]" italics="true" pageId="23" pageNumber="240">
<taxonomicName id="34FB98EF2415FF9676FD6648F17DF88E" authorityName="Fleutiaux" authorityYear="1918" box="[926,1121,1826,1845]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="240" phylum="Arthropoda" rank="species" species="beauchenei">Toxognathus beauchenei</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2415FF9671086648F161F88E" bold="true" box="[1131,1149,1826,1845]" pageId="23" pageNumber="240">E,</emphasis>
antenna;
<emphasis id="C18F3F7E2415FF96753C6656F56CF8F4" bold="true" box="[95,112,1852,1871]" pageId="23" pageNumber="240">F,</emphasis>
hind wing;
<emphasis id="C18F3F7E2415FF9675B26656F409F8F4" bold="true" box="[209,277,1852,1871]" pageId="23" pageNumber="240">G, H, I,</emphasis>
pro-, meso- and metathoracic leg (without tarsi);
<emphasis id="C18F3F7E2415FF9677D36656F7CBF8F4" bold="true" box="[688,727,1852,1871]" pageId="23" pageNumber="240">J, K,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C241AFF9975E763D2F443FCCB" blockId="24.[132,681,696,880]" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF9975E763D2F5C1FD6B" box="[132,221,696,720]" italics="true" pageId="24" pageNumber="241">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9241AFF99758963D2F431FD6B" box="[234,301,696,720]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="24" pageNumber="241">Figs. 7</figureCitation>
J-K): Phallobase 0.42x total length of aedeagus, 1.3x longer than wide, length ratio between lateroposterior and median parts 5.4; paramere with ventral anteromedian margin narrow, elongate and convergent, basal strut 0.29x total length of penis; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C241AFF9975E762F2F5E8FB8B" blockId="24.[132,681,920,1072]" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF9975E762F2F5FDFC0B" box="[132,225,920,944]" italics="true" pageId="24" pageNumber="241">
<typeStatus id="2C405DCE241AFF9975E762F2F5C0FC0B" box="[132,220,920,944]" pageId="24" pageNumber="241" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingRegion id="313F2D8E241AFF99759062F2F426FC0B" box="[243,314,920,944]" country="Vietnam" name="Ha Noi" pageId="24" pageNumber="241">Hanoi</collectingRegion>
,
<collectingCountry id="8BECA3FC241AFF99742B62F2F484FC0B" box="[328,408,920,944]" name="Vietnam" pageId="24" pageNumber="241">Tonkin</collectingCountry>
], [
<taxonomicName id="34FB98EF241AFF9974D462F2F5DDFC6B" authority="Beauchnei, Fleut." authorityName="Beauchnei, Fleut." class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E241AFF9974D462F2F7B5FC0B" box="[439,681,920,944]" italics="true" pageId="24" pageNumber="241">Toxognathus Beauchnei,</emphasis>
Fleut.
</taxonomicName>
Type, Colletion Fleutiaux], [Type], [Muséum Paris, Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF241AFF9974D562B2F43AFBAB" authority="(Fleutiaux)" baseAuthorityName="Fleutiaux" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="241" phylum="Arthropoda" rank="species" species="beauchenei">Toxognathus beauchenei (Fleutiaux)</taxonomicName>
, J. Chassain det. 05], [
<typeStatus id="2C405DCE241AFF99775B6292F7BFFBAB" box="[568,675,1016,1040]" pageId="24" pageNumber="241" type="lectotype">Lectotype</typeStatus>
] (MNHN).
</paragraph>
<paragraph id="F344E36C241AFF9975E76532F70DFB2B" blockId="24.[132,681,1112,1168]" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF9975E76532F44AFBCB" box="[132,342,1112,1136]" italics="true" pageId="24" pageNumber="241">Material examined:</emphasis>
<materialsCitation id="4393E931241AFF9974086532F711FB2B" collectionCode="MNHN" country="Vietnam" location="Tonkin" pageId="24" pageNumber="241" specimenCount="2">
<collectingCountry id="8BECA3FC241AFF9974086532F4F5FBCB" box="[363,489,1112,1136]" name="Vietnam" pageId="24" pageNumber="241">VIETNAM</collectingCountry>
.
<location id="F624B5B7241AFF9977626532F753FBCB" LSID="urn:lsid:plazi:treatment:7B52527A2415FF9977BC639DF1FDFD8B:F624B5B7241AFF9977626532F753FBCB" box="[513,591,1112,1136]" country="Vietnam" name="Tonkin" pageId="24" pageNumber="241">Tonkin</location>
,
<specimenCount id="E5FD28E5241AFF99770B6532F7B5FBD4" box="[616,681,1112,1136]" count="1" pageId="24" pageNumber="241" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9241AFF9975EF6512F5F4FB2B" box="[140,232,1144,1168]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="241">MNHN</collectionCode>
); Tam-Dao,
<specimenCount id="E5FD28E5241AFF9974096512F487FB34" box="[362,411,1144,1168]" count="1" pageId="24" pageNumber="241" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9241AFF9974CA6512F719FB2B" box="[425,517,1144,1168]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="241">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C241AFF9975E765D2F492FB6B" blockId="24.[132,398,1208,1232]" box="[132,398,1208,1232]" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF9975E765D2F41AFB6B" box="[132,262,1208,1232]" italics="true" pageId="24" pageNumber="241">Distribution:</emphasis>
<collectingCountry id="8BECA3FC241AFF99746F65D2F496FB6B" box="[268,394,1208,1232]" name="Vietnam" pageId="24" pageNumber="241">VIETNAM</collectingCountry>
.
</paragraph>
<paragraph id="F344E36C241AFF9977AD61D2F1FDFD8B" blockId="24.[718,1267,184,560]" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF9977AD61D2F634FF6B" box="[718,808,184,208]" italics="true" pageId="24" pageNumber="241">Remarks:</emphasis>
The specimen from Tam-Dao is somewhat different from the
<typeStatus id="2C405DCE241AFF9976F361B2F6F6FF4B" box="[912,1002,216,240]" pageId="24" pageNumber="241" type="holotype">holotype</typeStatus>
in its more convex body, black integument with antennae and legs reddish-brown and longer antennae (surpassing the posterior angles by one antennomere). Its phallobase differs from that of the specimen from
<collectingCountry id="8BECA3FC241AFF99714B6032F16FFECB" box="[1064,1139,344,368]" name="Vietnam" pageId="24" pageNumber="241">Tonkin</collectingCountry>
in the relative ratio between length and width (0.98) and ratio between lateroposterior and median parts (7.3).
<taxonomicName id="34FB98EF241AFF9971A560F3F684FE6B" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="241" phylum="Arthropoda" rank="species" species="beauchenei">
<emphasis id="C18F3F7E241AFF9971A560F3F684FE6B" italics="true" pageId="24" pageNumber="241">Toxognathus beauchenei</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="34FB98EF241AFF99713A60D3F1CEFE6B" authorityName="Fairmaire" authorityYear="1878" box="[1113,1234,440,464]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="241" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="C18F3F7E241AFF99713A60D3F1CEFE6B" box="[1113,1234,440,464]" italics="true" pageId="24" pageNumber="241">T. costulatus</emphasis>
</taxonomicName>
by its size, color of integument and the costate elytral interstices, but can be easily distinguished by its protibia with dorsal apical angle truncate and not produced.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,327 @@
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<heading id="A80C54002416FF95762061DDF161FF6B" bold="true" box="[835,1149,183,208]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="237" reason="2">
<taxonomicName id="34FB98EF2416FF95762061DDF161FF6B" authority="Fairmaire, 1878" authorityName="Fairmaire" authorityYear="1878" box="[835,1149,183,208]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2416FF95762061DDF161FF6B" bold="true" box="[835,1149,183,208]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF95762061DDF6D6FF74" bold="true" box="[835,970,183,207]" italics="true" pageId="20" pageNumber="237">Toxognathus</emphasis>
<bibRefCitation id="976A9E9D2416FF9576B361D2F161FF6B" author="FAIRMAIRE, L." box="[976,1149,184,208]" pageId="20" pageNumber="237" pagination="269 - 274" refId="ref49579" refString="FAIRMAIRE, L. 1878. Coleopteres de Conchinchine recueillis par M. le docteur Morice. Annales de La Societe Entomologique de France, 5 ª ser., 8: 269 - 274." type="journal article" year="1878">Fairmaire, 1878</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72416FF9577AD6192F1C2FECB" pageId="20" pageNumber="237" type="reference_group">
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<treatmentCitation id="725AC57D2416FF9577AD6192F133FEAB" author="FAIRMAIRE, L." box="[718,1071,248,272]" page="271" pageId="20" pageNumber="237" year="1878">
<taxonomicName id="34FB98EF2416FF9577AD6192F133FEAB" authority="Fairmaire, 1878: 271" authorityName="Fairmaire" authorityPageNumber="271" authorityYear="1878" box="[718,1071,248,272]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2416FF9577AD6192F659FEAB" box="[718,837,248,272]" italics="true" pageId="20" pageNumber="237">Toxognathus</emphasis>
<bibRefCitation id="976A9E9D2416FF9576336192F133FEAB" author="FAIRMAIRE, L." box="[848,1071,248,272]" pageId="20" pageNumber="237" pagination="269 - 274" refId="ref49579" refString="FAIRMAIRE, L. 1878. Coleopteres de Conchinchine recueillis par M. le docteur Morice. Annales de La Societe Entomologique de France, 5 ª ser., 8: 269 - 274." type="journal article" year="1878">Fairmaire, 1878: 271</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2416FF95715C6192F62DFE8B" author="FLEUTIAUX, E." page="276" pageId="20" pageNumber="237" year="1918">
<bibRefCitation id="976A9E9D2416FF95715C6192F62DFE8B" author="FLEUTIAUX, E." pageId="20" pageNumber="237" pagination="175 - 278" refId="ref49758" refString="FLEUTIAUX, E. 1918 a. Nouvelles contributions a la faune de l'Indo-Chine Francaise. Annales de La Societe Entomologique de France, 87: 175 - 278." type="journal article" year="1918">Fleutiaux, 1918a: 276</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2416FF9576216072F13FFE8B" author="FLEUTIAUX, E." box="[834,1059,280,304]" page="178" pageId="20" pageNumber="237" year="1924">
<bibRefCitation id="976A9E9D2416FF9576216072F13FFE8B" author="FLEUTIAUX, E." box="[834,1059,280,304]" pageId="20" pageNumber="237" pagination="31 - 184" refId="ref49879" refString="FLEUTIAUX, E. 1924. Melasidae et Elateridae. Faune entomologique de l'Indochine francaise. 1. Coleopteres serricornes. 2. Fam. Elateridae. Opuscules de l'Institut Scientifique de l'Indochine, 2 (7): 31 - 184." type="journal article" year="1924">Fleutiaux, 1924: 178</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2416FF9571566072F62DFEEB" author="SCHENKLING, S." page="509" pageId="20" pageNumber="237" year="1927">
<bibRefCitation id="976A9E9D2416FF9571566072F62DFEEB" author="SCHENKLING, S." pageId="20" pageNumber="237" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 509</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2416FF95765A6052F11BFEEB" author="FLEUTIAUX, E." box="[825,1031,312,336]" page="34" pageId="20" pageNumber="237" year="1940">
<bibRefCitation id="976A9E9D2416FF95765A6052F11BFEEB" author="FLEUTIAUX, E." box="[825,1031,312,336]" pageId="20" pageNumber="237" pagination="19 - 40" refId="ref49922" refString="FLEUTIAUX, E. 1940 a. Les elaterides de l'Indo-chine francaise. Huitieme et derniere partie. Annales de La Societe Entomologique de France, 109: 19 - 40." type="journal article" year="1940">Fleutiaux, 1940a: 34</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2416FF9571736052F1F3FEEB" author="FLEUTIAUX, E." box="[1040,1263,312,336]" page="104" pageId="20" pageNumber="237" year="1940">
<bibRefCitation id="976A9E9D2416FF9571736052F1F3FEEB" author="FLEUTIAUX, E." box="[1040,1263,312,336]" pageId="20" pageNumber="237" pagination="89 - 104" refId="ref49954" refString="FLEUTIAUX, E. 1940 b. Elaterides nouveaux. Bulletin et Annales de La Societe Entomologique de Belgique, 80: 89 - 104." type="journal article" year="1940">Fleutiaux, 1940b: 104</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2416FF9576646032F6DEFECB" author="STIBICK, J. N. L." box="[775,962,344,368]" page="168" pageId="20" pageNumber="237" year="1979">
<bibRefCitation id="976A9E9D2416FF9576646032F6DEFECB" author="STIBICK, J. N. L." box="[775,962,344,368]" pageId="20" pageNumber="237" pagination="145 - 186" refId="ref51152" refString="STIBICK, J. N. L. 1979. Classification of the Elateridae (Coleoptera). Relationships and classification of the subfamilies and tribes. Pacific Insects, 20: 145 - 186." type="journal article" year="1979">Stibick, 1979: 168</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2416FF9576AF6032F1C6FECB" author="BOUCHARD, P. &amp; BOUSQUET, I. &amp; DAVIES, A. E. &amp; ALONSO-ZARAZAGA, M. A. &amp; LAWRENCE, J. F. &amp; LYAL C. H. C. &amp; NEWTON, A. F. &amp; REID, C. A. M. &amp; SCHMITT, M. &amp; SMITH, A. B. T." box="[972,1242,344,368]" page="319" pageId="20" pageNumber="237" year="2011">
<bibRefCitation id="976A9E9D2416FF9576AF6032F1C6FECB" author="BOUCHARD, P. &amp; BOUSQUET, I. &amp; DAVIES, A. E. &amp; ALONSO-ZARAZAGA, M. A. &amp; LAWRENCE, J. F. &amp; LYAL C. H. C. &amp; NEWTON, A. F. &amp; REID, C. A. M. &amp; SCHMITT, M. &amp; SMITH, A. B. T." box="[972,1242,344,368]" pageId="20" pageNumber="237" pagination="1 - 972" refId="ref48935" refString="BOUCHARD, P.; BOUSQUET, I.; DAVIES, A. E.; ALONSO-ZARAZAGA, M. A.; LAWRENCE, J. F.; LYAL C. H. C.; NEWTON, A. F.; REID, C. A. M.; SCHMITT, M.; SLIPINSKI; S. A. &amp; SMITH, A. B. T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys, 88: 1 - 972." type="journal article" year="2011">
Bouchard
<emphasis id="C18F3F7E2416FF9571576033F175FECB" box="[1076,1129,344,368]" italics="true" pageId="20" pageNumber="237">et al.,</emphasis>
2011: 319
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72416FF9777AD60F3F1EFFCD4" lastPageId="22" lastPageNumber="239" pageId="20" pageNumber="237" type="materials_examined">
<paragraph id="F344E36C2416FF9577AD60F3F665FE6B" blockId="20.[718,1266,408,464]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9577AD60F3F65AFE0B" box="[718,838,409,432]" italics="true" pageId="20" pageNumber="237">
<typeStatus id="2C405DCE2416FF9577AD60F3F7E7FE0B" box="[718,763,409,432]" pageId="20" pageNumber="237">Type</typeStatus>
species
</emphasis>
(by monotypy):
<taxonomicName id="34FB98EF2416FF95716F60F2F66AFE6B" authority="Fairmaire, 1878" authorityName="Fairmaire" authorityYear="1878" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="237" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="C18F3F7E2416FF95716F60F2F1EEFE0B" box="[1036,1266,408,432]" italics="true" pageId="20" pageNumber="237">Toxognathus costulatus</emphasis>
<bibRefCitation id="976A9E9D2416FF9577AD60D2F66AFE6B" author="FAIRMAIRE, L." box="[718,886,440,464]" pageId="20" pageNumber="237" pagination="269 - 274" refId="ref49579" refString="FAIRMAIRE, L. 1878. Coleopteres de Conchinchine recueillis par M. le docteur Morice. Annales de La Societe Entomologique de France, 5 ª ser., 8: 269 - 274." type="journal article" year="1878">Fairmaire, 1878</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="F344E36C2416FF9577AD6093F1EFFCEB" blockId="20.[718,1267,504,848]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9577AD6093F63BFDAB" box="[718,807,505,528]" italics="true" pageId="20" pageNumber="237">Diagnosis</emphasis>
(male): Integument densely punctate with punctures rasp-like, covered with yellow setae predominantly erect, longer on lateral borders of elytra; pronotum wider than long, elytra parallel-sided on anterior 3/4 with striae deeply impressed with a row of punctures larger than those of pronotum, interstices convex or costate. Mandible short and wide with an oblique ridge on laterodorsal face, labrum subrectangular, concave and declivous anteroventrad. Posterior angles of ventrite 1 without marginal plate, of ventrites 2-4 with acute and produced marginal plates; tarsomeres simple, claws pectinate.
</paragraph>
<paragraph id="F344E36C2416FF9577AD6212F124FC2B" blockId="20.[718,1080,888,912]" box="[718,1080,888,912]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9577AD6212F64CFC2B" box="[718,848,888,912]" italics="true" pageId="20" pageNumber="237">Distribution:</emphasis>
<collectionCode id="95EA7BA92416FF9576356212F128FC2B" box="[854,1076,888,912]" pageId="20" pageNumber="237">SOUTHEAST ASIA</collectionCode>
.
</paragraph>
<paragraph id="F344E36C2416FF9577AD62D2F668FB34" blockId="20.[718,1267,952,1872]" pageId="20" pageNumber="237">
<emphasis id="C18F3F7E2416FF9577AD62D2F64CFC6B" box="[718,848,952,976]" italics="true" pageId="20" pageNumber="237">Redescription</emphasis>
(male): Integument bright, light to dark brown, reddish-brown or black; densely pilose, setae short, longer on abdominal ventrites, yellow or golden, erect on dorsal surface, predominantly decumbent ventrally. Total length: 5.0-12.0 mm; elytral base 1.0-1.1x wider than pronotum, elytra 2.5-3.1x longer than pronotum.
</paragraph>
<paragraph id="F344E36C2416FF94766465F2F777FE0B" blockId="20.[718,1267,952,1872]" lastBlockId="21.[94,643,184,1040]" lastPageId="21" lastPageNumber="238" pageId="20" pageNumber="237">
Head (
<figureCitation id="6BC0FFE92416FF95762865F2F689FB0B" box="[843,917,1176,1200]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="20" pageNumber="237">Fig. 9E</figureCitation>
) with margins over antennal insertions straight, oblique and convergent to the median straight margin. Frontal carina complete or absent between antennal insertions; frontoclypeal region gradually or steeply declivous to base of labrum about 5x wider than long; punctures coalescent 0.2 diameter apart. Antenna (
<figureCitation id="6BC0FFE92416FF9576156432F6D1FAD4" box="[886,973,1368,1392]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="20" pageNumber="237">Figs. 7E</figureCitation>
,
<figureCitation id="6BC0FFE92416FF9576BB6432F6EBFACB" box="[984,1015,1368,1392]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="20" pageNumber="237">9A</figureCitation>
) with 11 antennomeres, serrate from antennomere
<collectionCode id="95EA7BA92416FF95768A6412F11FFA34" box="[1001,1027,1400,1423]" pageId="20" pageNumber="237">IV</collectionCode>
, usually flat; antennomeres
<collectionCode id="95EA7BA92416FF95767264F2F63FFA14" box="[785,803,1432,1455]" pageId="20" pageNumber="237">II</collectionCode>
and
<collectionCode id="95EA7BA92416FF95763B64F2F66FFA14" box="[856,883,1432,1455]" pageId="20" pageNumber="237">III</collectionCode>
subequal,
<collectionCode id="95EA7BA92416FF95768164F2F6E1FA14" box="[994,1021,1432,1455]" pageId="20" pageNumber="237">III</collectionCode>
1.0-1.05x as long as
<collectionCode id="95EA7BA92416FF9571B964F2F1F2FA14" box="[1242,1262,1432,1455]" pageId="20" pageNumber="237">II</collectionCode>
;
<collectionCode id="95EA7BA92416FF9577AD64D2F7F5FA74" box="[718,745,1464,1487]" pageId="20" pageNumber="237">IV</collectionCode>
1.9-2.5x times longer than
<collectionCode id="95EA7BA92416FF95716D64D2F137FA74" box="[1038,1067,1464,1487]" pageId="20" pageNumber="237">III</collectionCode>
;
<collectionCode id="95EA7BA92416FF95715B64D2F168FA74" box="[1080,1140,1464,1487]" pageId="20" pageNumber="237">IV-VI</collectionCode>
subequal as wide as long; VII-XI increasing gradually in length to apex.
<collectionCode id="95EA7BA92416FF9576526492F644F9B4" box="[817,856,1528,1551]" pageId="20" pageNumber="237">IEP</collectionCode>
usually 0.34-0.39. Labrum (
<figureCitation id="6BC0FFE92416FF9571F76492F1F6F9AB" box="[1172,1258,1528,1552]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="20" pageNumber="237">Fig. 9D</figureCitation>
) subrectangular, anterior edge slightly emarginate or straight, concave and declivous anteroventrad, about 3.5x wider than long, coarsely punctate, with long setae. Mouthparts directed anteroventrad. Mandible (
<figureCitation id="6BC0FFE92416FF9577B667F2F639F90B" box="[725,805,1688,1712]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="20" pageNumber="237">Fig. 9D</figureCitation>
) wide and short, unidentate or bidentate, apical tooth short, subapical tooth smaller; laterodorsal face coarsely punctate with an oblique ridge steeply declivous to apical part; lateral edge abruptly curved apicad; mesal margin at base with a row of short setae. Maxilla and labium covered with light-brown setae. Maxilla (
<figureCitation id="6BC0FFE92417FF9475D461D2F41DFF6B" box="[183,257,184,208]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">Fig. 9B</figureCitation>
) with galea securiform, outer half with long and sparse setae, inner half with dense and short setae; lacinia elongate, elliptical, densely pilose; medistipes trapezoidal longer than wide with several short to long setae; Labium (
<figureCitation id="6BC0FFE92417FF94744C6052F462FEEB" box="[303,382,312,336]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">Fig. 9C</figureCitation>
) with prementum deeply emarginate on anteromedian margin with anterolateral angles rounded, not produced; maxillary and labial palpi densely pilose with apical palpomere elliptical.
</paragraph>
<paragraph id="F344E36C2417FF9475F460D2F779FBB4" blockId="21.[94,643,184,1040]" pageId="21" pageNumber="238">
Prothorax 1.10-1.25x wider than long, lateral carina complete, entirely lateroventral (
<figureCitation id="6BC0FFE92417FF94776060B2F74BFE4B" box="[515,599,472,496]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="21" pageNumber="238">Fig. 8B</figureCitation>
) or lateral on posterior region and directed ventrad anteriorly (
<figureCitation id="6BC0FFE92417FF9475DF6372F403FD8B" box="[188,287,536,560]" captionStart="FIGURE 11" captionStartId="27.[95,174,1826,1845]" captionTargetBox="[95,1228,1274,1801]" captionTargetId="figure-182@27.[94,1228,1273,1802]" captionTargetPageId="27" captionText="FIGURE 11: Toxognathus dohertyi. A, B, aedeagus (dorsal, ventral). Toxognathus fairmairei. C, hypomeron; D, pronotum and head. Toxognathus mouhoti. E, pronotum; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641389" httpUri="https://zenodo.org/record/12641389/files/figure.png" pageId="21" pageNumber="238">Fig. 11C</figureCitation>
). Pronotum flattened to convex, densely punctate, punctures rasp-like and umbilicate; posterior angles of pronotum non-carinate. Hypomeron (
<figureCitation id="6BC0FFE92417FF9475FB6312F5F2FD2B" box="[152,238,632,656]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="21" pageNumber="238">Figs. 8C</figureCitation>
,
<figureCitation id="6BC0FFE92417FF94759A6312F439FD2B" box="[249,293,632,656]" captionStart="FIGURE 11" captionStartId="27.[95,174,1826,1845]" captionTargetBox="[95,1228,1274,1801]" captionTargetId="figure-182@27.[94,1228,1273,1802]" captionTargetPageId="27" captionText="FIGURE 11: Toxognathus dohertyi. A, B, aedeagus (dorsal, ventral). Toxognathus fairmairei. C, hypomeron; D, pronotum and head. Toxognathus mouhoti. E, pronotum; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641389" httpUri="https://zenodo.org/record/12641389/files/figure.png" pageId="21" pageNumber="238">11C</figureCitation>
) with punctures smaller than those of pronotum, sparse, umbilicate, posterior 1/5 glabrous; posterior margin with an inverted U- or V-shaped short notch adjacent to posterior angle. Prosternal suture (
<figureCitation id="6BC0FFE92417FF9475F76392F5F7FCAB" box="[148,235,760,784]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="21" pageNumber="238">Figs. 8C</figureCitation>
,
<figureCitation id="6BC0FFE92417FF9475946392F404FCAB" box="[247,280,760,784]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">9G</figureCitation>
) curved, marginate by a shiny band along hypomeral margin. Prosternum (
<figureCitation id="6BC0FFE92417FF9474926272F757FC8B" box="[497,587,792,816]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="21" pageNumber="238">Figs. 8C</figureCitation>
,
<figureCitation id="6BC0FFE92417FF9477396272F760FC8B" box="[602,636,792,816]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">9G</figureCitation>
) 1.07-1.20x wider than long, with punctures larger than those of pronotum, 0.5-1.0 diameter apart; anterior lobe truncate, covering mentum. Prosternal process (
<figureCitation id="6BC0FFE92417FF94750662F2F5A6FC0B" box="[101,186,920,944]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="21" pageNumber="238">Figs. 8C</figureCitation>
,
<figureCitation id="6BC0FFE92417FF9475A762F2F5F9FC0B" box="[196,229,920,944]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">9G</figureCitation>
) 2.5-2.6x longer than diameter of procoxae, curved between procoxae then straight, ventral surface a little narrower and shorter than the ventral surface forming a subapical tooth (
<figureCitation id="6BC0FFE92417FF9474086292F4A7FBAB" box="[363,443,1016,1040]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="21" pageNumber="238">Fig. 8D</figureCitation>
). Procoxae open.
</paragraph>
<paragraph id="F344E36C2417FF94753D6552F18FFD0B" blockId="21.[94,643,1080,1872]" lastBlockId="21.[680,1229,184,1552]" pageId="21" pageNumber="238">
<emphasis id="C18F3F7E2417FF94753D6552F5C8FBEB" box="[94,212,1080,1104]" italics="true" pageId="21" pageNumber="238">Pterothorax:</emphasis>
Mesoventrite with posterior region abruptly inclined ventrad about 30° in relation to anterior region (
<figureCitation id="6BC0FFE92417FF9475B46512F401FB2B" box="[215,285,1144,1168]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">Fig. 9I</figureCitation>
), with anterior articulating surfaces (
<figureCitation id="6BC0FFE92417FF94750565F2F5A9FB0B" box="[102,181,1176,1200]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">Fig. 9H</figureCitation>
) concave bordered posteriorly by oblique carina, borders of mesoventral cavity nearly straight and divergent to mesocoxae then convergent (
<figureCitation id="6BC0FFE92417FF94748E65B2F723FB4B" box="[493,575,1240,1264]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">Fig. 9H</figureCitation>
); floor of cavity with shiny median band. Mesocoxal cavity (
<figureCitation id="6BC0FFE92417FF9475056472F5B5FA8B" box="[102,169,1304,1328]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">Fig. 9I</figureCitation>
) open to both mesepisternum and mesepimeron, trochantin visible; mesepisternum with an oblique carina anteriorly contiguous to the anterior carina of mesoventral articulating surface. Mesometaventral suture distinct. Metaventrite (
<figureCitation id="6BC0FFE92417FF9474E664F2F4C9FA0B" box="[389,469,1432,1456]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="21" pageNumber="238">Fig. 7B</figureCitation>
) 1.4-1.5x wider than long, 1.5-1.8x longer than mesoventrite, with punctation umbilicate, 0.2-1.0 diameter apart; metepisternum about 6.3-7.0x longer than wide. Scutellum abruptly elevated above the level of mesoscutum, pentagonal to subrectangular, longer than wide. Elytra gradually widened from humerus to posterior 1/3-1/4 then gradually rounded to apex; apices conjointly rounded; striae deeply impressed with a row of punctures; interstices convex, punctate, rugose; epipleura abruptly narrowed near metacoxa, slightly widened apicad from ventrite 2. Hind wings (
<figureCitation id="6BC0FFE92417FF9474B66792F73AF8B4" box="[469,550,1784,1808]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="21" pageNumber="238">Figs. 7F</figureCitation>
,
<figureCitation id="6BC0FFE92417FF9477526792F753F8AB" box="[561,591,1784,1808]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="21" pageNumber="238">8E</figureCitation>
,
<figureCitation id="6BC0FFE92417FF9477386792F767F8AB" box="[603,635,1784,1808]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">9K</figureCitation>
) with radial cell 3.75x longer than wide, CuA
<subScript id="6F7FE1292417FF947747664DF730F88E" attach="left" box="[548,556,1831,1845]" fontSize="6" pageId="21" pageNumber="238">1</subScript>
present; wedge cell absent; apex with median and posterior field sclerotizations, convergent basad and an oblique sclerotization adjacent to the radial cell. Legs, Metacoxa inclined about 24° in relation to transverse axis of body, gradually shortened laterally, median ventral part about 3.5x longer than lateral part; free margin of metacoxal plate absent or short; metatrochanter and metafemur (
<figureCitation id="6BC0FFE92417FF9477D36012F7E4FE2B" box="[688,760,376,400]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="21" pageNumber="238">Fig. 7I</figureCitation>
) weakly convex, metafemur 2.7-3.5x longer than wide. Tibiae (
<figureCitation id="6BC0FFE92417FF94761260F2F6A4FE0B" box="[881,952,408,432]" captionStart="FIGURE 7" captionStartId="23.[96,175,1826,1845]" captionTargetBox="[97,1227,1106,1794]" captionTargetId="figure-321@23.[94,1228,1103,1802]" captionTargetPageId="23" captionText="FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641381" httpUri="https://zenodo.org/record/12641381/files/figure.png" pageId="21" pageNumber="238">Figs. 7</figureCitation>
G-I; 9J) 1.7-3.5x wider at apex than at base; apices with two subequal spurs longer than setae, with a row of 8-15 spiniform setae along each outer and inner apical border and 2-3 irregular rows of spiniform setae on dorsal margin, spiniform setae shorter than the fine setae. Tarsomeres (
<figureCitation id="6BC0FFE92417FF94711D6352F1D9FDEB" box="[1150,1221,568,592]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">Fig. 9J</figureCitation>
) decreasing in length from I-IV,
<collectionCode id="95EA7BA92417FF9476946332F115FDD4" box="[1015,1033,600,623]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="21" pageNumber="238" type="Museum">V</collectionCode>
about 1.2x longer than
<collectionCode id="95EA7BA92417FF9477836312F7E7FD34" box="[736,763,632,655]" pageId="21" pageNumber="238">III</collectionCode>
and
<collectionCode id="95EA7BA92417FF9476506312F652FD34" box="[819,846,632,655]" pageId="21" pageNumber="238">IV</collectionCode>
together, simple, densely pilose ventrally; claws pectinate (with more than six teeth).
</paragraph>
<paragraph id="F344E36C2417FF94778263D2F16FF9B4" blockId="21.[680,1229,184,1552]" pageId="21" pageNumber="238">
Abdominal ventrites (
<figureCitation id="6BC0FFE92417FF9476A663D2F106FD6B" box="[965,1050,696,720]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="21" pageNumber="238">Figs. 8F</figureCitation>
,
<figureCitation id="6BC0FFE92417FF94714B63D2F159FD6B" box="[1064,1093,696,720]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">9L</figureCitation>
) I-IV evenly covered with short setae and punctures 0.5-1.0 diameters apart, apical part of ventrite 5 with punctures coalescent forming longitudinal striae; posterior angles of ventrite 1 right without marginal plate, ventrites 2-4 with triangular marginal plates widened posteriad; ventrite 1 about 3x longer on lateral part than at middle, ventrites 2-4 subequal in length; ventrite 5 subtriangular 1.55-1.95x wider than long, strongly convex posteriorly (
<figureCitation id="6BC0FFE92417FF94761962B2F6CAFC4B" box="[890,982,984,1008]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="21" pageNumber="238">Fig. 9M</figureCitation>
). Pregenitalic segments and aedeagus covered with brown setae. Sternite VIII (
<figureCitation id="6BC0FFE92417FF9477D36572F611FB8B" box="[688,781,1048,1072]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="21" pageNumber="238">Fig. 10A</figureCitation>
) subrectangular, 1.7-2.0x wider than long, bilobed posteriorly, with a pair of light sclerotizations on lateral lobes and a dark T-shaped sclerotization on anteromedian margin; posterior margin of lobes with long brown setae; tergite VIII (
<figureCitation id="6BC0FFE92417FF9476BD65F2F12BFB0B" box="[990,1079,1176,1200]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="21" pageNumber="238">Fig. 10B</figureCitation>
) evenly sclerotized with short and long brown setae on lateroposterior margin; sternite IX (
<figureCitation id="6BC0FFE92417FF9476DC65B2F101FB4B" box="[959,1053,1240,1264]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="21" pageNumber="238">Fig. 10C</figureCitation>
) with anteromedian margin rounded or acute, produced and curved dorsad, with a longitudinal dorsal apodeme, posterior part covered with brown setae longer on lateroposteri- or and apical margin; sternite and tergite IX fused near the anterior margin; tergite IX (
<figureCitation id="6BC0FFE92417FF9476906412F14FFA2B" box="[1011,1107,1400,1424]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="21" pageNumber="238">Fig. 10D</figureCitation>
) with anterior margin bisinuose or straight, posterior lobes acute with long brown setae; tergite X (
<figureCitation id="6BC0FFE92417FF94716D64D2F173FA6B" box="[1038,1135,1464,1488]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="21" pageNumber="238">Fig. 10D</figureCitation>
) suboval with long brown setae on posterior border, apical margin membranous fringed by minuscule setae.
</paragraph>
<paragraph id="F344E36C2417FF9477CB6752F18BF8F4" blockId="21.[680,1229,1592,1872]" pageId="21" pageNumber="238">
<emphasis id="C18F3F7E2417FF9477CB6752F61DF9EB" box="[680,769,1592,1616]" italics="true" pageId="21" pageNumber="238">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92417FF9476706752F680F9EB" box="[787,924,1592,1616]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="21" pageNumber="238">Figs. 10E, F</figureCitation>
): Phallobase M-shaped with lateroposterior parts parallel-sided or divergent posteriorly. Parameres usually not contiguous ventrally (fused basally in
<taxonomicName id="34FB98EF2417FF94763A67F3F6C8F90B" authorityName="Fairmaire" authorityYear="1878" box="[857,980,1688,1712]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="238" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="C18F3F7E2417FF94763A67F3F6C8F90B" box="[857,980,1688,1712]" italics="true" pageId="21" pageNumber="238">T. costulatus</emphasis>
</taxonomicName>
), with apex securiform, partly membranous, with short and long brown setae. Penis tapering to apex, dorsal articulation with a short and narrow process between the basal struts fused to the parameres; basal struts 0.28-0.34x shorter than the total length of penis; ventral sclerite present.
</paragraph>
<paragraph id="F344E36C2414FF9775E761D2F74CFCCB" blockId="22.[132,681,184,880]" pageId="22" pageNumber="239">
<emphasis id="C18F3F7E2414FF9775E761D2F478FF6B" box="[132,356,184,208]" italics="true" pageId="22" pageNumber="239">
Remarks:
<taxonomicName id="34FB98EF2414FF97758961D2F478FF6B" authorityName="Fairmaire" authorityYear="1878" box="[234,356,184,208]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">Toxognathus</taxonomicName>
</emphasis>
shares with “Physodacylinae” genera the widened tibiae and the prosternal anterior margin truncate, and can be easily recognized by its pectinate claws.
<taxonomicName id="34FB98EF2414FF9774526072F4B7FE8B" authorityName="Fairmaire" authorityYear="1878" box="[305,427,280,304]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF9774526072F4B7FE8B" box="[305,427,280,304]" italics="true" pageId="22" pageNumber="239">Toxognathus</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="34FB98EF2414FF97770F6072F5DFFEEB" authorityName="Candeze" authorityYear="1859" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="tribe" tribe="Melanotini">Melanotini</taxonomicName>
(
<taxonomicName id="34FB98EF2414FF9775B26052F427FEEB" authorityName="Leach" authorityYear="1815" box="[209,315,312,336]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="subFamily" subFamily="Elaterinae">Elaterinae</taxonomicName>
) genera by its pectinate claws, with more than six teeth.
<taxonomicName id="34FB98EF2414FF97743D6032F739FECB" box="[350,549,344,368]" pageId="22" pageNumber="239">
<emphasis id="C18F3F7E2414FF97743D6032F4C4FECB" box="[350,472,344,368]" italics="true" pageId="22" pageNumber="239">Toxognathus</emphasis>
species
</taxonomicName>
differs from the majority of the species of
<taxonomicName id="34FB98EF2414FF9774D56012F734FE2B" authorityName="Candeze" authorityYear="1859" box="[438,552,376,400]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="tribe" tribe="Melanotini">Melanotini</taxonomicName>
in (the later in parentheses): mesocoxal cavity open to mesepimeron and mesepisternum (open to mesepimeron only) and anterior prosternal margin short and truncate (rounded and produced).
<taxonomicName id="34FB98EF2414FF9774CA6092F73FFDAB" authorityName="Fairmaire" authorityYear="1878" box="[425,547,504,528]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF9774CA6092F73FFDAB" box="[425,547,504,528]" italics="true" pageId="22" pageNumber="239">Toxognathus</emphasis>
</taxonomicName>
also differs from North American and Australian species of
<taxonomicName id="34FB98EF2414FF97770E6372F5A4FDEB" authorityName="Eschscholtz" authorityYear="1829" class="Insecta" family="Elateridae" genus="Melanotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF97770E6372F5A4FDEB" italics="true" pageId="22" pageNumber="239">Melanotus</emphasis>
</taxonomicName>
in the phallobase with anterior edge notched (truncate or rounded) and by the absence of wedge cell on hind wing (present) (
<bibRefCitation id="976A9E9D2414FF9774E66312F7B8FD2B" author="QUATE, L. W. &amp; THOMPSON, S. E." box="[389,676,632,656]" pageId="22" pageNumber="239" pagination="1 - 86" refId="ref50858" refString="QUATE, L. W. &amp; THOMPSON, S. E. 1967. Revision of Click Beetles of the genus Melanotus in America North of Mexico (Coleoptera, Elateridae). Proceedings of the United States National Museum, 121: 1 - 86." type="journal article" year="1967">Quate &amp; Thompson, 1967</bibRefCitation>
;
<bibRefCitation id="976A9E9D2414FF9775E763F2F40DFD0B" author="CALDER, A. A." box="[132,273,664,688]" pageId="22" pageNumber="239" pagination="257 - 260" refId="ref49048" refString="CALDER, A. A. 1983. Melanotus Eschscholtz, a genus new to Australia (Coleoptera: Elateridae). Journal of the Australian Entomological Society, 22: 257 - 260." type="journal article" year="1983">Calder, 1983</bibRefCitation>
). Nevertheless, some Palearctic species have phallobase more similar to that of
<emphasis id="C18F3F7E2414FF97774063D2F7B5FD6B" box="[547,681,696,720]" italics="true" pageId="22" pageNumber="239">Toxoganathus</emphasis>
species (
<bibRefCitation id="976A9E9D2414FF9775B563B2F4A7FD4B" author="KABALAK, M. &amp; SERT, O." box="[214,443,728,752]" pageId="22" pageNumber="239" pagination="71 - 82" refId="ref50357" refString="KABALAK, M. &amp; SERT, O. 2011. Systematic studies on the male genital organs of Central Anatolian Elateridae (Coleoptera) Species part I: the subfamilies Elaterinae and Melanotinae. Hacettepe Journal of Biology and Chemistry, 39: 71 - 82." type="journal article" year="2011">Kabalak &amp; Sert, 2011</bibRefCitation>
). Mouthparts and hind wings have been studied only for a few species of
<emphasis id="C18F3F7E2414FF97771A6392F5D7FC8B" italics="true" pageId="22" pageNumber="239">
<taxonomicName id="34FB98EF2414FF97771A6392F5DBFC8B" authorityName="Eschscholtz" authorityYear="1829" class="Insecta" family="Elateridae" genus="Melanotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">Melanotus</taxonomicName>
,
</emphasis>
therefore the apparent synapomorphies found in these structures of
<taxonomicName id="34FB98EF2414FF9774066252F4C3FCEB" authorityName="Fairmaire" authorityYear="1878" box="[357,479,824,848]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF9774066252F4C3FCEB" box="[357,479,824,848]" italics="true" pageId="22" pageNumber="239">Toxognathus</emphasis>
</taxonomicName>
could not be compared with those of other
<taxonomicName id="34FB98EF2414FF9774EE6232F4E3FCCB" authorityName="Candeze" authorityYear="1859" box="[397,511,856,880]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="tribe" tribe="Melanotini">Melanotini</taxonomicName>
genera.
</paragraph>
<paragraph id="F344E36C2414FF97766461D2F1EFFCD4" blockId="22.[718,1267,184,880]" pageId="22" pageNumber="239">
<bibRefCitation id="976A9E9D2414FF97766461D2F680FF6B" author="HAYEK, C. M. F. VON" box="[775,924,184,208]" pageId="22" pageNumber="239" pagination="37 - 115" refId="ref50284" refString="HAYEK, C. M. F. VON. 1990. A reclassification of the Melanotus group of genera (Coleoptera: Elateridae). Bulletin of the British Museum (Natural History), 59: 37 - 115." type="journal article" year="1990">Hayek (1990)</bibRefCitation>
revised the
<taxonomicName id="34FB98EF2414FF97714761D2F196FF6B" authorityName="Eschscholtz" authorityYear="1829" box="[1060,1162,184,208]" class="Insecta" family="Elateridae" genus="Melanotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF97714761D2F196FF6B" box="[1060,1162,184,208]" italics="true" pageId="22" pageNumber="239">Melanotus</emphasis>
</taxonomicName>
group of genera and showed that the majority of species have mesocoxal cavity open to mesepimeron, although the length of the suture that separates the mesepisternum from the coxal cavity is variable and at least one species of
<taxonomicName id="34FB98EF2414FF9776796033F11EFECB" authority="Laporte, 1840" authorityName="Laporte" authorityYear="1840" box="[794,1026,344,368]" class="Insecta" family="Elateridae" genus="Priopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF9776796033F678FECB" box="[794,868,345,368]" italics="true" pageId="22" pageNumber="239">Priopus</emphasis>
Laporte, 1840
</taxonomicName>
have mesocoxal cavity open to both mesepimeron and mesepisternum.
<taxonomicName id="34FB98EF2414FF9771A56013F639FE0B" authorityName="Fairmaire" authorityYear="1878" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF9771A56013F639FE0B" italics="true" pageId="22" pageNumber="239">Toxognathus</emphasis>
</taxonomicName>
can be separated from species of this genus by its posterior angles of prothorax shorter and not strongly pointed sharply in ventral view and the tibiae widened apically.
<bibRefCitation id="976A9E9D2414FF9776D26092F159FDAB" author="HAYEK, C. M. F. VON" box="[945,1093,504,528]" pageId="22" pageNumber="239" pagination="37 - 115" refId="ref50284" refString="HAYEK, C. M. F. VON. 1990. A reclassification of the Melanotus group of genera (Coleoptera: Elateridae). Bulletin of the British Museum (Natural History), 59: 37 - 115." type="journal article" year="1990">Hayek (1990)</bibRefCitation>
presented a key to elaterid genera with pectinate claws in which she separated
<taxonomicName id="34FB98EF2414FF9776586352F6A9FDEB" authorityName="Fairmaire" authorityYear="1878" box="[827,949,568,592]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF9776586352F6A9FDEB" box="[827,949,568,592]" italics="true" pageId="22" pageNumber="239">Toxognathus</emphasis>
</taxonomicName>
from the other genera with “procoxal cavity wide open posteriorly” and “mesepisternum separated from the metasternum by the mesepimeron” by the following combination of characters: mandible unidentate, anterior margin of prosternum truncate and pronotum without basal lateral incisions. Nevertheless, some species of
<taxonomicName id="34FB98EF2414FF97711B6392F1EEFCAB" authorityName="Fairmaire" authorityYear="1878" box="[1144,1266,760,784]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2414FF97711B6392F1EEFCAB" box="[1144,1266,760,784]" italics="true" pageId="22" pageNumber="239">Toxognathus</emphasis>
</taxonomicName>
have mandibles bidentate and a pair of short lateral incisions on pronotum, therefore they can be recognized only by the anterior prosternal margin truncate.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,183 @@
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<subSubSection id="BBE1B0E72418FF9B766A61DDF1ABFF6B" box="[777,1207,183,208]" pageId="26" pageNumber="243" type="nomenclature">
<paragraph id="F344E36C2418FF9B766A61DDF1ABFF6B" blockId="26.[777,1207,183,240]" box="[777,1207,183,208]" pageId="26" pageNumber="243">
<heading id="A80C54002418FF9B766A61DDF1ABFF6B" bold="true" box="[777,1207,183,208]" centered="true" fontSize="10" level="2" pageId="26" pageNumber="243" reason="2">
<taxonomicName id="34FB98EF2418FF9B766A61DDF1ABFF6B" authority="Fleutiaux, 1940" authorityName="Fleutiaux" authorityYear="1940" box="[777,1207,183,208]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="243" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="C18F3F7E2418FF9B766A61DDF1ABFF6B" bold="true" box="[777,1207,183,208]" pageId="26" pageNumber="243">
<emphasis id="C18F3F7E2418FF9B766A61DDF11AFF74" bold="true" box="[777,1030,183,207]" italics="true" pageId="26" pageNumber="243">Toxognathus fairmairei</emphasis>
Fleutiaux, 1940
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="BBE1B0E72418FF9B761761B2F150FF4B" box="[884,1100,216,240]" pageId="26" pageNumber="243" type="description">
<paragraph id="F344E36C2418FF9B761761B2F150FF4B" blockId="26.[777,1207,183,240]" box="[884,1100,216,240]" pageId="26" pageNumber="243">
<emphasis id="C18F3F7E2418FF9B761761B2F150FF4B" bold="true" box="[884,1100,216,240]" pageId="26" pageNumber="243">
(
<figureCitation id="6BC0FFE92418FF9B761861B2F114FF4B" box="[891,1032,216,240]" captionStart="FIGURE 11" captionStartId="27.[95,174,1826,1845]" captionTargetBox="[95,1228,1274,1801]" captionTargetId="figure-182@27.[94,1228,1273,1802]" captionTargetPageId="27" captionText="FIGURE 11: Toxognathus dohertyi. A, B, aedeagus (dorsal, ventral). Toxognathus fairmairei. C, hypomeron; D, pronotum and head. Toxognathus mouhoti. E, pronotum; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641389" httpUri="https://zenodo.org/record/12641389/files/figure.png" pageId="26" pageNumber="243">Figs. 11C, D</figureCitation>
;
<figureCitation id="6BC0FFE92418FF9B717761B2F159FF4B" box="[1044,1093,216,240]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="26" pageNumber="243">34K</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72418FF9B77AD6072F1B9FE8B" box="[718,1189,280,304]" pageId="26" pageNumber="243" type="reference_group">
<paragraph id="F344E36C2418FF9B77AD6072F1B9FE8B" blockId="26.[718,1189,280,304]" box="[718,1189,280,304]" pageId="26" pageNumber="243">
<treatmentCitationGroup id="D3EBC4422418FF9B77AD6072F1B9FE8B" box="[718,1189,280,304]" pageId="26" pageNumber="243">
<treatmentCitation id="725AC57D2418FF9B77AD6072F1BDFE8B" author="FLEUTIAUX, E." box="[718,1185,280,304]" page="104" pageId="26" pageNumber="243" year="1940">
<taxonomicName id="34FB98EF2418FF9B77AD6072F1BDFE8B" authority="Fleutiaux, 1940 b: 104" authorityName="Fleutiaux" authorityPageNumber="104" authorityYear="1940" box="[718,1185,280,304]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="243" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="C18F3F7E2418FF9B77AD6072F6AEFE8B" box="[718,946,280,304]" italics="true" pageId="26" pageNumber="243">Toxognathus fairmairei</emphasis>
<bibRefCitation id="976A9E9D2418FF9B76DA6072F1BDFE8B" author="FLEUTIAUX, E." box="[953,1185,280,304]" pageId="26" pageNumber="243" pagination="89 - 104" refId="ref49954" refString="FLEUTIAUX, E. 1940 b. Elaterides nouveaux. Bulletin et Annales de La Societe Entomologique de Belgique, 80: 89 - 104." type="journal article" year="1940">Fleutiaux, 1940b: 104</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72418FF9D77AD6032F5A1FEF4" lastPageId="28" lastPageNumber="245" pageId="26" pageNumber="243" type="description">
<paragraph id="F344E36C2418FF9A77AD6032F5A9FB6B" blockId="26.[718,1266,344,496]" lastBlockId="27.[94,643,760,1232]" lastPageId="27" lastPageNumber="244" pageId="26" pageNumber="243">
<emphasis id="C18F3F7E2418FF9B77AD6032F64FFECB" box="[718,851,344,368]" italics="true" pageId="26" pageNumber="243">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE92418FF9B76D76032F101FECB" box="[948,1053,344,368]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="26" pageNumber="243">Fig. 34K</figureCitation>
): Integument dark brown with legs and ventral abdomen brown, antenna light brown. Total length
<quantity id="34034E892418FF9B712160F2F183FE14" box="[1090,1183,408,432]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="26" pageNumber="243" unit="mm" value="7.5">7.5 mm</quantity>
; elytral base 1.09x as wide as pronotum, elytra 3.14x longer than pronotum. Frons deeply concave along the anterior margin; frontal carina complete; eyes very small (IEP 0.23) (
<figureCitation id="6BC0FFE92419FF9A747F6272F49CFC8B" box="[284,384,792,816]" captionStart="FIGURE 11" captionStartId="27.[95,174,1826,1845]" captionTargetBox="[95,1228,1274,1801]" captionTargetId="figure-182@27.[94,1228,1273,1802]" captionTargetPageId="27" captionText="FIGURE 11: Toxognathus dohertyi. A, B, aedeagus (dorsal, ventral). Toxognathus fairmairei. C, hypomeron; D, pronotum and head. Toxognathus mouhoti. E, pronotum; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641389" httpUri="https://zenodo.org/record/12641389/files/figure.png" pageId="27" pageNumber="244">Fig. 11D</figureCitation>
). Antennae reaching the base of posterior angles of pronotum; antennomeres IV-X as long as wide and convex. Mandible bidentate. Pronotum weakly convex 1.20x wider than long with sides parallel from base of posterior angles to anterior third then convergent to anterior margin, lateral carina entirely lateroventral, posterior angles divergent and weakly convex; punctures less than 0.5 diameter apart; prosternum with a median longitudinal groove on posterior half and between procoxae. Elytral interstices convex. Free margin of metacoxal plate absent, tibiae slender with out- er angle truncate. Tergite IX with anterior margin straight.
</paragraph>
<caption id="A784B3E42418FF9B75E66662F111F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641385" ID-Zenodo-Dep="12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="26" pageNumber="243" startId="26.[133,212,1800,1819]" targetBox="[136,1262,516,1767]" targetPageId="26" targetType="figure">
<paragraph id="F344E36C2418FF9B75E66662F111F8F4" blockId="26.[132,1266,1800,1871]" pageId="26" pageNumber="243">
<emphasis id="C18F3F7E2418FF9B75E66662F5F5F8A0" bold="true" box="[133,233,1800,1819]" pageId="26" pageNumber="243">FIGURE 9:</emphasis>
<emphasis id="C18F3F7E2418FF9B758E6663F4B9F8A7" box="[237,421,1801,1820]" italics="true" pageId="26" pageNumber="243">
<taxonomicName id="34FB98EF2418FF9B758E6663F4BDF8A7" box="[237,417,1801,1820]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="243" phylum="Arthropoda" rank="species" species="costulatus">Toxognathus costulatus</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2418FF9B74CB6662F4A7F8A0" bold="true" box="[424,443,1800,1819]" pageId="26" pageNumber="243">A,</emphasis>
antenna,
<emphasis id="C18F3F7E2418FF9B77686663F701F8A7" bold="true" box="[523,541,1801,1820]" pageId="26" pageNumber="243">B,</emphasis>
maxilla;
<emphasis id="C18F3F7E2418FF9B77046662F760F8A0" bold="true" box="[615,636,1800,1819]" pageId="26" pageNumber="243">C,</emphasis>
labium;
<emphasis id="C18F3F7E2418FF9B77A06663F7C5F8A7" bold="true" box="[707,729,1801,1820]" pageId="26" pageNumber="243">D,</emphasis>
labrum and mandibles;
<emphasis id="C18F3F7E2418FF9B76FD6663F6ACF8A7" bold="true" box="[926,944,1801,1820]" pageId="26" pageNumber="243">E,</emphasis>
head (dorsal);
<emphasis id="C18F3F7E2418FF9B714B6663F125F8A7" bold="true" box="[1064,1081,1801,1820]" pageId="26" pageNumber="243">F,</emphasis>
pronotum;
<emphasis id="C18F3F7E2418FF9B71FA6662F1B3F8A7" bold="true" box="[1177,1199,1800,1820]" pageId="26" pageNumber="243">G,</emphasis>
prosternum;
<emphasis id="C18F3F7E2418FF9B75D56648F5D1F88E" bold="true" box="[182,205,1826,1845]" pageId="26" pageNumber="243">H,</emphasis>
mesoventrite;
<emphasis id="C18F3F7E2418FF9B74266648F44FF88E" bold="true" box="[325,339,1826,1845]" pageId="26" pageNumber="243">I,</emphasis>
mesoventrite, pleural sclerites and mesocoxa (lateroventral);
<emphasis id="C18F3F7E2418FF9B76206648F64DF88E" bold="true" box="[835,849,1826,1845]" pageId="26" pageNumber="243">J,</emphasis>
protibia and tarsus;
<emphasis id="C18F3F7E2418FF9B769A6648F111F88E" bold="true" box="[1017,1037,1826,1845]" pageId="26" pageNumber="243">K,</emphasis>
hind wing;
<emphasis id="C18F3F7E2418FF9B71126648F1BCF88E" bold="true" box="[1137,1184,1826,1845]" pageId="26" pageNumber="243">L, M,</emphasis>
abdomen (ventral, lateral).
<emphasis id="C18F3F7E2418FF9B74726656F49EF8F4" box="[273,386,1852,1871]" italics="true" pageId="26" pageNumber="243">Abbreviations:</emphasis>
ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate.
</paragraph>
</caption>
<caption id="A784B3E42419FF9A753C63AAF1D4FD68" ID-DOI="http://doi.org/10.5281/zenodo.12641387" ID-Zenodo-Dep="12641387" box="[95,1224,704,724]" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="27" pageNumber="244" startId="27.[95,174,704,723]" targetBox="[95,1225,171,680]" targetPageId="27" targetType="figure">
<paragraph id="F344E36C2419FF9A753C63AAF1D4FD68" blockId="27.[95,1224,704,724]" box="[95,1224,704,724]" pageId="27" pageNumber="244">
<emphasis id="C18F3F7E2419FF9A753C63AAF5D3FD68" bold="true" box="[95,207,704,723]" pageId="27" pageNumber="244">FIGURE 10:</emphasis>
<emphasis id="C18F3F7E2419FF9A75B763AAF491FD68" box="[212,397,704,723]" italics="true" pageId="27" pageNumber="244">
<taxonomicName id="34FB98EF2419FF9A75B763AAF495FD68" box="[212,393,704,723]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="244" phylum="Arthropoda" rank="species" species="costulatus">Toxognathus costulatus</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2419FF9A74F263AAF4B8FD68" bold="true" box="[401,420,704,723]" pageId="27" pageNumber="244">A,</emphasis>
sternite VIII;
<emphasis id="C18F3F7E2419FF9A777A63AAF730FD68" bold="true" box="[537,556,704,723]" pageId="27" pageNumber="244">B,</emphasis>
tergite VIII;
<emphasis id="C18F3F7E2419FF9A77FA63AAF7B2FD68" bold="true" box="[665,686,704,723]" pageId="27" pageNumber="244">C,</emphasis>
sternite IX;
<emphasis id="C18F3F7E2419FF9A767763AAF637FD68" bold="true" box="[788,811,704,723]" pageId="27" pageNumber="244">D,</emphasis>
tergites IX and X;
<emphasis id="C18F3F7E2419FF9A76A463AAF6F1FD68" bold="true" box="[967,1005,704,723]" pageId="27" pageNumber="244">E, F,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C2419FF9A77CB6392F10AFCEB" blockId="27.[680,1229,760,848]" pageId="27" pageNumber="244">
<emphasis id="C18F3F7E2419FF9A77CB6392F614FCAB" box="[680,776,760,784]" italics="true" pageId="27" pageNumber="244">Aedeagus:</emphasis>
Phallobase 0.36x total length of aedeagus, 1.13x wider than long. Paramere with anteromedian margin acute, ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C2419FF9A77CB6212F68EFB8B" blockId="27.[680,1229,888,1072]" pageId="27" pageNumber="244">
<emphasis id="C18F3F7E2419FF9A77CB6212F619FC2B" box="[680,773,888,912]" italics="true" pageId="27" pageNumber="244">
<typeStatus id="2C405DCE2419FF9A77CB6212F61CFC2B" box="[680,768,888,912]" pageId="27" pageNumber="244" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Foutchien, De Latouche], [Collection Fairmaire], [Collection E. Fleutiaux], [
<taxonomicName id="34FB98EF2419FF9A712662F2F64DFC6B" authority="Fleut." authorityName="Fleutiaux" authorityYear="1940" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="244" phylum="Arthropoda" rank="species" species="fairmairei">Toxognathus fairmairei Fleut.</taxonomicName>
, type, Collection Fleuxiaux], [Type], [Muséum Paris, Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF2419FF9A712662B2F696FBAB" authority="(Fleutiaux)" baseAuthorityName="Fleutiaux" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="244" phylum="Arthropoda" rank="species" species="fairmairei">Toxognathus fairmairei (Fleutiaux)</taxonomicName>
, J. Chassain det. 05], [
<typeStatus id="2C405DCE2419FF9A71EB6292F7C5FB8B" pageId="27" pageNumber="244" type="lectotype">Lectotype</typeStatus>
], male (MNHN).
</paragraph>
<paragraph id="F344E36C2419FF9A77CB6532F6C2FBCB" blockId="27.[680,990,1112,1136]" box="[680,990,1112,1136]" pageId="27" pageNumber="244">
<emphasis id="C18F3F7E2419FF9A77CB6532F636FBCB" box="[680,810,1112,1136]" italics="true" pageId="27" pageNumber="244">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2419FF9A76526532F697FBCB" box="[817,907,1112,1136]" name="China" pageId="27" pageNumber="244">CHINA</collectingCountry>
.
<emphasis id="C18F3F7E2419FF9A76F66533F6C2FBCB" box="[917,990,1113,1136]" italics="true" pageId="27" pageNumber="244">
<collectingRegion id="313F2D8E2419FF9A76F66533F6C7FBCB" box="[917,987,1113,1136]" country="China" name="Fujian" pageId="27" pageNumber="244">Fujian</collectingRegion>
.
</emphasis>
</paragraph>
<paragraph id="F344E36C2419FF9D77CB65F2F5A1FEF4" blockId="27.[680,1228,1176,1232]" lastBlockId="28.[132,681,184,335]" lastPageId="28" lastPageNumber="245" pageId="27" pageNumber="244">
<emphasis id="C18F3F7E2419FF9A77CB65F2F61FFB0B" box="[680,771,1176,1200]" italics="true" pageId="27" pageNumber="244">Remarks:</emphasis>
The aedeagus of the
<typeStatus id="2C405DCE2419FF9A769065F2F152FB0B" box="[1011,1102,1176,1200]" pageId="27" pageNumber="244" type="holotype">holotype</typeStatus>
was poorly preserved, with dark musculature and apex broken; therefore it was not possible to obtain all measurements. It is distinguishable from all
<taxonomicName id="34FB98EF241EFF9D749F61B2F5B5FEAB" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D749F61B2F764FF4B" box="[508,632,216,240]" italics="true" pageId="28" pageNumber="245">Toxognathus</emphasis>
species
</taxonomicName>
by the shape of prothorax, the very small eyes and the median longitudinal groove on the prosternum.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,176 @@
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<subSubSection id="BBE1B0E7241AFF997670631DF1B1FD2B" box="[787,1197,631,656]" pageId="24" pageNumber="241" type="nomenclature">
<paragraph id="F344E36C241AFF997670631DF1B1FD2B" blockId="24.[787,1197,631,688]" box="[787,1197,631,656]" pageId="24" pageNumber="241">
<heading id="A80C5400241AFF997670631DF1B1FD2B" bold="true" box="[787,1197,631,656]" centered="true" fontSize="10" level="2" pageId="24" pageNumber="241" reason="2">
<taxonomicName id="34FB98EF241AFF997670631DF1B1FD2B" authority="Fleutiaux, 1940" authorityName="Fleutiaux" authorityYear="1940" box="[787,1197,631,656]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="241" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="C18F3F7E241AFF997670631DF1B1FD2B" bold="true" box="[787,1197,631,656]" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF997670631DF6E0FD2B" bold="true" box="[787,1020,631,656]" italics="true" pageId="24" pageNumber="241">Toxognathus coomani</emphasis>
Fleutiaux, 1940
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241AFF9976F763F2F130FD0B" box="[916,1068,664,688]" pageId="24" pageNumber="241" type="description">
<paragraph id="F344E36C241AFF9976F763F2F130FD0B" blockId="24.[787,1197,631,688]" box="[916,1068,664,688]" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF9976F763F2F130FD0B" bold="true" box="[916,1068,664,688]" pageId="24" pageNumber="241">
(
<figureCitation id="6BC0FFE9241AFF9976F863F2F6FAFD0B" box="[923,998,664,688]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="24" pageNumber="241">Figs. 8</figureCitation>
,
<figureCitation id="6BC0FFE9241AFF99769163F2F138FD0B" box="[1010,1060,664,688]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="24" pageNumber="241">34H</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241AFF9977AD63B2F198FD4B" box="[718,1156,728,752]" pageId="24" pageNumber="241" type="reference_group">
<paragraph id="F344E36C241AFF9977AD63B2F198FD4B" blockId="24.[718,1156,728,752]" box="[718,1156,728,752]" pageId="24" pageNumber="241">
<treatmentCitationGroup id="D3EBC442241AFF9977AD63B2F198FD4B" box="[718,1156,728,752]" pageId="24" pageNumber="241">
<treatmentCitation id="725AC57D241AFF9977AD63B2F19DFD4B" author="FLEUTIAUX, E." box="[718,1153,728,752]" page="35" pageId="24" pageNumber="241" year="1940">
<taxonomicName id="34FB98EF241AFF9977AD63B2F19DFD4B" authority="Fleutiaux, 1940 a: 35" authorityName="Fleutiaux" authorityYear="1940" box="[718,1153,728,752]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="241" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="C18F3F7E241AFF9977AD63B2F6BEFD4B" box="[718,930,728,752]" italics="true" pageId="24" pageNumber="241">Toxognathus coomani</emphasis>
<bibRefCitation id="976A9E9D241AFF9976CB63B2F19DFD4B" author="FLEUTIAUX, E." box="[936,1153,728,752]" pageId="24" pageNumber="241" pagination="19 - 40" refId="ref49922" refString="FLEUTIAUX, E. 1940 a. Les elaterides de l'Indo-chine francaise. Huitieme et derniere partie. Annales de La Societe Entomologique de France, 109: 19 - 40." type="journal article" year="1940">Fleutiaux, 1940a: 35</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241AFF9877AD6272F798FB34" lastPageId="25" lastPageNumber="242" pageId="24" pageNumber="241" type="description">
<paragraph id="F344E36C241AFF9877AD6272F755FEEB" blockId="24.[718,1267,792,1232]" lastBlockId="25.[94,643,184,336]" lastPageId="25" lastPageNumber="242" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF9977AD6272F64CFC8B" box="[718,848,792,816]" italics="true" pageId="24" pageNumber="241">Redescription</emphasis>
(male, 39H): Integument brown to dark brown, legs and antennae lighter. Total length
<quantity id="34034E89241AFF9977AD6232F650FCD4" box="[718,844,856,880]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="24" pageNumber="241" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.5-6.5 mm</quantity>
; elytral base 1.03-1.08x as wide as pronotum, elytra 2.65-2.67x longer than pronotum. Frons concave along the anterior margin; frontal carina complete. Antennae surpassing the posterior angle of pronotum by one antennomere; antennomeres IV-X as long as wide and convex. Mandible bidentate (
<figureCitation id="6BC0FFE9241AFF9977B66572F636FB8B" box="[725,810,1048,1072]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="24" pageNumber="241">Fig. 8A</figureCitation>
). Pronotum (
<figureCitation id="6BC0FFE9241AFF9976D96572F116FB8B" box="[954,1034,1048,1072]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="24" pageNumber="241">Fig. 8B</figureCitation>
) flattened 1.11-1.16x wider than long with sides slightly rounded, convergent on anterior 1/4; lateral carina entirely lateroventral (not visible from dorsal view) (
<figureCitation id="6BC0FFE9241AFF9971556512F1AEFB2B" box="[1078,1202,1144,1168]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="24" pageNumber="241">Figs. 8C, D</figureCitation>
), posterior angles subparallel and convex; posterior margin with a pair of lateral incisions adjacent to the posterior angles; punctures 0.5 diameter apart on disc, coalescent on lateral border. Elytral interstices convex. Free margin of metacoxal plate short, triangular; tibiae slightly widened apicad with dorsal apical angle truncate. Tergite IX with anterior margin bisinuous.
</paragraph>
<caption id="A784B3E4241AFF9975E06648F48DF8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641383" ID-Zenodo-Dep="12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="24" pageNumber="241" startId="24.[131,210,1826,1845]" targetBox="[135,1265,1280,1799]" targetPageId="24" targetType="figure">
<paragraph id="F344E36C241AFF9975E06648F48DF8F4" blockId="24.[131,1267,1826,1871]" pageId="24" pageNumber="241">
<emphasis id="C18F3F7E241AFF9975E06648F5F6F88E" bold="true" box="[131,234,1826,1845]" pageId="24" pageNumber="241">FIGURE 8:</emphasis>
<emphasis id="C18F3F7E241AFF99758C6648F483F88E" box="[239,415,1826,1845]" italics="true" pageId="24" pageNumber="241">
<taxonomicName id="34FB98EF241AFF99758C6648F487F88E" authorityName="Fleutiaux" authorityYear="1940" box="[239,411,1826,1845]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="241" phylum="Arthropoda" rank="species" species="coomani">Toxognathus coomani</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E241AFF9974C06648F4AAF88E" bold="true" box="[419,438,1826,1845]" pageId="24" pageNumber="241">A,</emphasis>
mandible (ventral);
<emphasis id="C18F3F7E241AFF99773D6648F7BBF88E" bold="true" box="[606,679,1826,1845]" pageId="24" pageNumber="241">B, C, D,</emphasis>
prothorax (dorsal, ventral, lateral);
<emphasis id="C18F3F7E241AFF9976A86648F6C1F88E" bold="true" box="[971,989,1826,1845]" pageId="24" pageNumber="241">E,</emphasis>
hind wing;
<emphasis id="C18F3F7E241AFF9971226648F14EF88E" bold="true" box="[1089,1106,1826,1845]" pageId="24" pageNumber="241">F,</emphasis>
abdomen (ventral);
<emphasis id="C18F3F7E241AFF9975E76656F5AAF8F4" bold="true" box="[132,182,1852,1871]" pageId="24" pageNumber="241">G, H,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C241BFF98753D6012F749FDAB" blockId="25.[94,643,376,528]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF98753D6012F5ABFE2B" box="[94,183,376,400]" italics="true" pageId="25" pageNumber="242">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9241BFF9875A56012F458FE2B" box="[198,324,376,400]" captionStart="FIGURE 8" captionStartId="24.[131,210,1826,1845]" captionTargetBox="[135,1265,1280,1799]" captionTargetId="figure-431@24.[132,1266,1273,1802]" captionTargetPageId="24" captionText="FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641383" httpUri="https://zenodo.org/record/12641383/files/figure.png" pageId="25" pageNumber="242">Figs. 8G, H</figureCitation>
): Phallobase 0.37x total length of aedeagus, 1.2x longer than wide, length ratio between lateroposterior and median parts 3; paramere with ventral anteromedian margin acute; basal struts 0.28x total length of penis; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C241BFF98753D6352F4C3FD6B" blockId="25.[94,643,568,720]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF98753D6352F5A7FDEB" box="[94,187,568,592]" italics="true" pageId="25" pageNumber="242">
<typeStatus id="2C405DCE241BFF98753D6352F5AAFDEB" box="[94,182,568,592]" pageId="25" pageNumber="242" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Ton Kin, (Lac Thô),
<collectingRegion id="313F2D8E241BFF9874A66352F728FDEB" box="[453,564,568,592]" country="Vietnam" name="Hoa Binh" pageId="25" pageNumber="242">Hoa Binh</collectingRegion>
, A. de Cooman], [
<taxonomicName id="34FB98EF241BFF9875BA6332F4D5FDCB" authorityName="Fleutiaux" authorityYear="1940" box="[217,457,600,624]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="coomani">Toxognathus coomani</taxonomicName>
, type, Collection Fleutiaux], [Type], [Muséum Paris, Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF241BFF9875D163F2F700FD0B" authority="(Fleutiaux)" baseAuthorityName="Fleutiaux" box="[178,540,664,688]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="coomani">Toxognathus coomani (Fleutiaux)</taxonomicName>
, J. Chassain det. 05], [
<typeStatus id="2C405DCE241BFF98759263D2F442FD6B" box="[241,350,696,720]" pageId="25" pageNumber="242" type="lectotype">Lectotype</typeStatus>
], (MNHN).
</paragraph>
<paragraph id="F344E36C241BFF98753D6392F5D2FC8B" blockId="25.[94,643,760,816]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF98753D6392F42CFCAB" box="[94,304,760,784]" italics="true" pageId="25" pageNumber="242">Material examined:</emphasis>
<materialsCitation id="4393E931241BFF9874256392F5D7FC8B" collectionCode="MNHN" country="Vietnam" location="Tonkin" pageId="25" pageNumber="242" specimenCount="4">
VETNAM.
<collectingCountry id="8BECA3FC241BFF9874B16392F73CFCAB" box="[466,544,760,784]" name="Vietnam" pageId="25" pageNumber="242">Tonkin</collectingCountry>
,
<specimenCount id="E5FD28E5241BFF9877596392F79FFCB4" box="[570,643,760,784]" count="4" pageId="25" pageNumber="242" type="generic">4 exs.</specimenCount>
(
<collectionCode id="95EA7BA9241BFF9875056272F5DEFC8B" box="[102,194,792,816]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="242">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C241BFF98753D6232F758FCCB" blockId="25.[94,580,856,880]" box="[94,580,856,880]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF98753D6232F5FDFCCB" box="[94,225,856,880]" italics="true" pageId="25" pageNumber="242">Distribution:</emphasis>
<collectingCountry id="8BECA3FC241BFF9875856232F478FCCB" box="[230,356,856,880]" name="Vietnam" pageId="25" pageNumber="242">VIETNAM</collectingCountry>
.
<emphasis id="C18F3F7E241BFF98740D6233F4CBFCCB" box="[366,471,856,880]" italics="true" pageId="25" pageNumber="242">
<collectingRegion id="313F2D8E241BFF98740D6233F4CDFCCB" box="[366,465,856,880]" country="Vietnam" name="Hoa Binh" pageId="25" pageNumber="242">Hoa Binh</collectingRegion>
:
</emphasis>
Lac Thuy.
</paragraph>
<paragraph id="F344E36C241BFF98753D62F2F798FB34" blockId="25.[94,644,920,1168]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF98753D62F2F48DFC0B" box="[94,401,920,944]" italics="true" pageId="25" pageNumber="242">
Remarks:
<taxonomicName id="34FB98EF241BFF9875DC62F2F48DFC0B" box="[191,401,920,944]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="coomani">Toxognathus coomani</taxonomicName>
</emphasis>
shares with
<taxonomicName id="34FB98EF241BFF98777662F3F79FFC0B" authorityName="Fleutiaux" authorityYear="1918" box="[533,643,920,944]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="C18F3F7E241BFF98777662F3F79FFC0B" box="[533,643,920,944]" italics="true" pageId="25" pageNumber="242">T. mouhoti</emphasis>
</taxonomicName>
and
<taxonomicName id="34FB98EF241BFF9875EC62D3F414FC6B" box="[143,264,952,976]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="fairmarei">
<emphasis id="C18F3F7E241BFF9875EC62D3F414FC6B" box="[143,264,952,976]" italics="true" pageId="25" pageNumber="242">T. fairmarei</emphasis>
</taxonomicName>
the bidentate mandibles. It is most similar to
<taxonomicName id="34FB98EF241BFF9875AA62B3F42BFC4B" authorityName="Fleutiaux" authorityYear="1918" box="[201,311,984,1008]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="C18F3F7E241BFF9875AA62B3F42BFC4B" box="[201,311,984,1008]" italics="true" pageId="25" pageNumber="242">T. mouhoti</emphasis>
</taxonomicName>
by its body size, eye size, shape of prothorax, free margin of metacoxal plate short and triangular and widened tibiae. It can be distinguished mainly by its frons, which is concave along the anterior border, complete frontal carina, shorter antenna and a pair of short notches on posterior margin of pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,178 @@
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<subSubSection id="BBE1B0E7241BFF9875F865BDF75AFB4B" box="[155,582,1239,1264]" pageId="25" pageNumber="242" type="nomenclature">
<paragraph id="F344E36C241BFF9875F865BDF75AFB4B" blockId="25.[155,582,1239,1296]" box="[155,582,1239,1264]" pageId="25" pageNumber="242">
<heading id="A80C5400241BFF9875F865BDF75AFB4B" bold="true" box="[155,582,1239,1264]" centered="true" fontSize="10" level="2" pageId="25" pageNumber="242" reason="2">
<taxonomicName id="34FB98EF241BFF9875F865BDF75AFB4B" authority="Fairmaire, 1878" authorityName="Fairmaire" authorityYear="1878" box="[155,582,1239,1264]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="C18F3F7E241BFF9875F865BDF75AFB4B" bold="true" box="[155,582,1239,1264]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9875F865BDF48FFB54" bold="true" box="[155,403,1239,1263]" italics="true" pageId="25" pageNumber="242">Toxognathus costulatus</emphasis>
<bibRefCitation id="976A9E9D241BFF9874FA65B2F75AFB4B" author="FAIRMAIRE, L." box="[409,582,1240,1264]" pageId="25" pageNumber="242" pagination="269 - 274" refId="ref49579" refString="FAIRMAIRE, L. 1878. Coleopteres de Conchinchine recueillis par M. le docteur Morice. Annales de La Societe Entomologique de France, 5 ª ser., 8: 269 - 274." type="journal article" year="1878">Fairmaire, 1878</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241BFF9874756592F4D7FAAB" box="[278,459,1272,1296]" pageId="25" pageNumber="242" type="description">
<paragraph id="F344E36C241BFF9874756592F4D7FAAB" blockId="25.[155,582,1239,1296]" box="[278,459,1272,1296]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9874756592F4D7FAAB" bold="true" box="[278,459,1272,1296]" pageId="25" pageNumber="242">
(
<figureCitation id="6BC0FFE9241BFF98747E6592F47BFAAB" box="[285,359,1272,1296]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="25" pageNumber="242">Figs. 9</figureCitation>
,
<figureCitation id="6BC0FFE9241BFF9874106592F48CFAAB" box="[371,400,1272,1296]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="25" pageNumber="242">10</figureCitation>
,
<figureCitation id="6BC0FFE9241BFF9874FF6592F4DFFAAB" box="[412,451,1272,1296]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="25" pageNumber="242">34I</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241BFF98753D6452F45BFA2B" pageId="25" pageNumber="242" type="reference_group">
<paragraph id="F344E36C241BFF98753D6452F45BFA2B" blockId="25.[94,643,1336,1424]" pageId="25" pageNumber="242">
<treatmentCitationGroup id="D3EBC442241BFF98753D6452F45BFA2B" pageId="25" pageNumber="242">
<treatmentCitation id="725AC57D241BFF98753D6452F731FAEB" author="FAIRMAIRE, L." box="[94,557,1336,1360]" page="271" pageId="25" pageNumber="242" year="1878">
<taxonomicName id="34FB98EF241BFF98753D6452F731FAEB" authority="Fairmaire, 1878: 271" authorityName="Fairmaire" authorityPageNumber="271" authorityYear="1878" box="[94,557,1336,1360]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="C18F3F7E241BFF98753D6452F45DFAEB" box="[94,321,1336,1360]" italics="true" pageId="25" pageNumber="242">Toxognathus costulatus</emphasis>
<bibRefCitation id="976A9E9D241BFF9874286452F731FAEB" author="FAIRMAIRE, L." box="[331,557,1336,1360]" pageId="25" pageNumber="242" pagination="269 - 274" refId="ref49579" refString="FAIRMAIRE, L. 1878. Coleopteres de Conchinchine recueillis par M. le docteur Morice. Annales de La Societe Entomologique de France, 5 ª ser., 8: 269 - 274." type="journal article" year="1878">Fairmaire, 1878: 271</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="725AC57D241BFF98775F6452F457FACB" author="SCHENKLING, S." page="509" pageId="25" pageNumber="242" year="1927">
<bibRefCitation id="976A9E9D241BFF98775F6452F457FACB" author="SCHENKLING, S." pageId="25" pageNumber="242" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 509</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D241BFF9874346432F75DFACB" author="FLEUTIAUX, E." box="[343,577,1368,1392]" page="276" pageId="25" pageNumber="242" year="1918">
<bibRefCitation id="976A9E9D241BFF9874346432F75DFACB" author="FLEUTIAUX, E." box="[343,577,1368,1392]" pageId="25" pageNumber="242" pagination="175 - 278" refId="ref49758" refString="FLEUTIAUX, E. 1918 a. Nouvelles contributions a la faune de l'Indo-Chine Francaise. Annales de La Societe Entomologique de France, 87: 175 - 278." type="journal article" year="1918">Fleutiaux, 1918a: 276</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D241BFF98772E6432F45FFA2B" author="FLEUTIAUX, E." page="35" pageId="25" pageNumber="242" year="1940">
<bibRefCitation id="976A9E9D241BFF98772E6432F45FFA2B" author="FLEUTIAUX, E." pageId="25" pageNumber="242" pagination="19 - 40" refId="ref49922" refString="FLEUTIAUX, E. 1940 a. Les elaterides de l'Indo-chine francaise. Huitieme et derniere partie. Annales de La Societe Entomologique de France, 109: 19 - 40." type="journal article" year="1940">Fleutiaux, 1940a: 35</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241BFF98753D64D2F621FC4B" pageId="25" pageNumber="242" type="description">
<paragraph id="F344E36C241BFF98753D64D2F6F3FEAB" blockId="25.[94,644,1464,1872]" lastBlockId="25.[680,1229,184,272]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF98753D64D2F5FCFA6B" box="[94,224,1464,1488]" italics="true" pageId="25" pageNumber="242">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9241BFF98744864D2F462FA6B" box="[299,382,1464,1488]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="25" pageNumber="242">Fig. 34I</figureCitation>
): Integument dark brown or dark reddish-brown. Total length
<quantity id="34034E89241BFF9874BD64B2F74DFA54" box="[478,593,1496,1520]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="25" pageNumber="242" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11-12 mm</quantity>
; elytral base as wide as pronotum, elytra 2.7-3.0x longer than pronotum. Frons (
<figureCitation id="6BC0FFE9241BFF9874376772F4D2F98B" box="[340,462,1560,1584]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="25" pageNumber="242">Figs. 9D, E</figureCitation>
) concave medially, frontal carina absent medially. Antennae (
<figureCitation id="6BC0FFE9241BFF98774F6752F767F9EB" box="[556,635,1592,1616]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="25" pageNumber="242">Fig. 9A</figureCitation>
) surpassing posterior angles of pronotum by 2 antennomeres. Mandible unidentate (
<figureCitation id="6BC0FFE9241BFF9874CC6712F719F92B" box="[431,517,1656,1680]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="25" pageNumber="242">Fig. 9D</figureCitation>
). Pronotum (
<figureCitation id="6BC0FFE9241BFF98750567F2F5A8F90B" box="[102,180,1688,1712]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="25" pageNumber="242">Fig. 9F</figureCitation>
) flattened 1.15-1.30x wider than long with sides sinuous converging abruptly anteriad on anterior half, lateral carina directed ventrad anteriorly, posterior angles divergent and convex; with punctures less than 0.5 diameter apart. Elytral interstices costate. Free margin of metacoxal plate short and triangular; tibiae widened apicad, dorso apical angle of protibiae acute and produced (
<figureCitation id="6BC0FFE9241BFF9876EA61B2F6C8FF4B" box="[905,980,216,240]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="25" pageNumber="242">Fig. 9J</figureCitation>
). Tergite IX (
<figureCitation id="6BC0FFE9241BFF98710061B2F1D8FF4B" box="[1123,1220,216,240]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="25" pageNumber="242">Fig. 10D</figureCitation>
) with anterior margin bisinuous.
</paragraph>
<paragraph id="F344E36C241BFF9877CB6052F649FE4B" blockId="25.[680,1229,312,496]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9877CB6052F61DFEEB" box="[680,769,312,336]" italics="true" pageId="25" pageNumber="242">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9241BFF98766C6052F693FEEB" box="[783,911,312,336]" captionStart="FIGURE 10" captionStartId="27.[95,174,704,723]" captionTargetBox="[95,1225,171,680]" captionTargetId="figure-216@27.[94,1228,170,680]" captionTargetPageId="27" captionText="FIGURE 10: Toxognathus costulatus. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641387" httpUri="https://zenodo.org/record/12641387/files/figure.png" pageId="25" pageNumber="242">Figs. 10E, F</figureCitation>
): Phallobase 0.44x total length of aedeagus, 1.3x longer than wide, length ratio between lateroposterior and median parts 3.1; parameres fused to each other on ventral anteromedian margin; basal strut 0.34x total length of penis; ventral sclerite elliptical.
</paragraph>
<paragraph id="F344E36C241BFF9877CB6372F1D8FD2B" blockId="25.[680,1229,536,656]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9877CB6372F619FD8B" box="[680,773,536,560]" italics="true" pageId="25" pageNumber="242">
<typeStatus id="2C405DCE241BFF9877CB6372F61CFD8B" box="[680,768,536,560]" pageId="25" pageNumber="242" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Conchinch. (
<emphasis id="C18F3F7E241BFF9876FC6372F6F7FD8B" box="[927,1003,536,560]" italics="true" pageId="25" pageNumber="242">illegible</emphasis>
) 1877] [
<taxonomicName id="34FB98EF241BFF9871266372F1D1FD8B" authorityName="Rosa" authorityYear="2014" box="[1093,1229,536,560]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="292" phylum="Arthropoda" rank="genus" status="gen. et sp. nov.">Toxognathus</taxonomicName>
<taxonomicNameLabel id="DABC8205241BFF9877CB6352F64EFDF4" box="[680,850,568,592]" pageId="25" pageNumber="242" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<taxonomicName id="34FB98EF241BFF98763E6352F1DAFDEB" authority=", Fairm., Conchonchine" authorityName="Fairm., Conchonchine" box="[861,1222,568,592]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="costulatus">costulatus, Fairm., Conchonchine</taxonomicName>
] [
<taxonomicName id="34FB98EF241BFF9877D26332F170FDCB" authority="Fairm., J. Chassain" authorityName="Fairm., J. Chassain" box="[689,1132,600,624]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="C18F3F7E241BFF9877D26332F688FDCB" box="[689,916,600,624]" italics="true" pageId="25" pageNumber="242">Toxognathus costulatus</emphasis>
Fairm., J. Chassain
</taxonomicName>
det. 05] [Muséum Paris, Coll. E. Fleutiaux] male, (MNHN).
</paragraph>
<paragraph id="F344E36C241BFF9877CB63D2F680FD4B" blockId="25.[680,1228,696,752]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9877CB63D2F677FD6B" box="[680,875,696,720]" italics="true" pageId="25" pageNumber="242">Material examined:</emphasis>
<materialsCitation id="4393E931241BFF98761163D2F7FDFD4B" collectionCode="MNHN" country="Vietnam" location="Tay Ninh" pageId="25" pageNumber="242" specimenCount="1" stateProvince="Tay Ninh">
<collectingCountry id="8BECA3FC241BFF98761163D2F6ECFD6B" box="[882,1008,696,720]" name="Vietnam" pageId="25" pageNumber="242">VIETNAM</collectingCountry>
.
<specimenCount id="E5FD28E5241BFF98769963D2F130FD74" box="[1018,1068,696,720]" count="1" pageId="25" pageNumber="242" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9241BFF98715963D2F18AFD6B" box="[1082,1174,696,720]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="242">MNHN</collectionCode>
).
<collectingRegion id="313F2D8E241BFF9871C463D2F7FDFD4B" country="Vietnam" name="Tay Ninh" pageId="25" pageNumber="242">Tay Ninh</collectingRegion>
</materialsCitation>
,
<materialsCitation id="4393E931241BFF98778F63B2F685FD4B" box="[748,921,728,752]" collectionCode="MNHN" country="Vietnam" pageId="25" pageNumber="242" specimenCount="2">
<specimenCount id="E5FD28E5241BFF98778F63B2F63BFD54" box="[748,807,728,752]" count="2" pageId="25" pageNumber="242" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA9241BFF98765763B2F68CFD4B" box="[820,912,728,752]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="242">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C241BFF9877CB6272F101FC8B" blockId="25.[680,1053,792,816]" box="[680,1053,792,816]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9877CB6272F636FC8B" box="[680,810,792,816]" italics="true" pageId="25" pageNumber="242">Distribution:</emphasis>
<collectingCountry id="8BECA3FC241BFF9876536272F6B2FC8B" box="[816,942,792,816]" name="Vietnam" pageId="25" pageNumber="242">VIETNAM</collectingCountry>
.
<emphasis id="C18F3F7E241BFF9876DB6273F101FC8B" box="[952,1053,792,816]" italics="true" pageId="25" pageNumber="242">
<collectingRegion id="313F2D8E241BFF9876DB6273F105FC8B" box="[952,1049,792,816]" country="Vietnam" name="Tay Ninh" pageId="25" pageNumber="242">Tay Ninh</collectingRegion>
.
</emphasis>
</paragraph>
<paragraph id="F344E36C241BFF9877CB6232F621FC4B" blockId="25.[680,1228,856,1008]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9877CB6232F6FAFCCB" box="[680,998,856,880]" italics="true" pageId="25" pageNumber="242">
Remarks:
<taxonomicName id="34FB98EF241BFF9876646232F6FAFCCB" box="[775,998,856,880]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="costulatus">Toxognathus costulatus</taxonomicName>
</emphasis>
is easily recognized by its protibia widened apicad with dorso apical angle acute and produced (
<figureCitation id="6BC0FFE9241BFF9876E362F2F6D4FC0B" box="[896,968,920,944]" captionStart="FIGURE 9" captionStartId="26.[133,212,1800,1819]" captionTargetBox="[136,1262,516,1767]" captionTargetId="figure-99@26.[132,1266,511,1777]" captionTargetPageId="26" captionText="FIGURE 9: Toxognathus costulatus. A, antenna, B, maxilla; C, labium; D, labrum and mandibles; E, head (dorsal); F,pronotum; G, prosternum; H, mesoventrite; I, mesoventrite, pleural sclerites and mesocoxa (lateroventral); J, protibia and tarsus; K, hind wing; L, M, abdomen (ventral, lateral). Abbreviations: ala, anterolateral angle; as, articulating surface; abp, abdominal marginal plate." figureDoi="http://doi.org/10.5281/zenodo.12641385" httpUri="https://zenodo.org/record/12641385/files/figure.png" pageId="25" pageNumber="242">Fig. 9J</figureCitation>
). The parameres with anteromedian margin fused together ventrally is unique to this species.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,118 @@
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<heading id="A80C5400241BFF98778D655DF19AFBEB" bold="true" box="[750,1158,1079,1104]" centered="true" fontSize="10" level="2" pageId="25" pageNumber="242" reason="2">
<taxonomicName id="34FB98EF241BFF98778D655DF19AFBEB" authority="Fleutiaux, 1940" authorityName="Fleutiaux" authorityYear="1940" box="[750,1158,1079,1104]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="242" phylum="Arthropoda" rank="species" species="dohertyi">
<emphasis id="C18F3F7E241BFF98778D655DF19AFBEB" bold="true" box="[750,1158,1079,1104]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF98778D655DF6C9FBF4" bold="true" box="[750,981,1079,1103]" italics="true" pageId="25" pageNumber="242">Toxognathus dohertyi</emphasis>
Fleutiaux, 1940
</emphasis>
</taxonomicName>
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<paragraph id="F344E36C241BFF9876356532F103FBCB" blockId="25.[750,1158,1079,1136]" box="[854,1055,1112,1136]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9876356532F103FBCB" bold="true" box="[854,1055,1112,1136]" pageId="25" pageNumber="242">
(
<figureCitation id="6BC0FFE9241BFF98763E6532F6F8FBCB" box="[861,996,1112,1136]" captionStart="FIGURE 11" captionStartId="27.[95,174,1826,1845]" captionTargetBox="[95,1228,1274,1801]" captionTargetId="figure-182@27.[94,1228,1273,1802]" captionTargetPageId="27" captionText="FIGURE 11: Toxognathus dohertyi. A, B, aedeagus (dorsal, ventral). Toxognathus fairmairei. C, hypomeron; D, pronotum and head. Toxognathus mouhoti. E, pronotum; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641389" httpUri="https://zenodo.org/record/12641389/files/figure.png" pageId="25" pageNumber="242">Figs. 11A, B</figureCitation>
;
<figureCitation id="6BC0FFE9241BFF9876936532F104FBCB" box="[1008,1048,1112,1136]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="25" pageNumber="242">34J</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241BFF9877CB65F2F177FB0B" box="[680,1131,1176,1200]" pageId="25" pageNumber="242" type="reference_group">
<paragraph id="F344E36C241BFF9877CB65F2F177FB0B" blockId="25.[680,1131,1176,1200]" box="[680,1131,1176,1200]" pageId="25" pageNumber="242">
<treatmentCitationGroup id="D3EBC442241BFF9877CB65F2F177FB0B" box="[680,1131,1176,1200]" pageId="25" pageNumber="242">
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<emphasis id="C18F3F7E241BFF9877CB65F2F664FB0B" box="[680,888,1176,1200]" italics="true" pageId="25" pageNumber="242">Toxognathus dohertyi</emphasis>
<bibRefCitation id="976A9E9D241BFF98761C65F2F17BFB0B" author="FLEUTIAUX, E." box="[895,1127,1176,1200]" pageId="25" pageNumber="242" pagination="89 - 104" refId="ref49954" refString="FLEUTIAUX, E. 1940 b. Elaterides nouveaux. Bulletin et Annales de La Societe Entomologique de Belgique, 80: 89 - 104." type="journal article" year="1940">Fleutiaux, 1940b: 104</bibRefCitation>
</taxonomicName>
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.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241BFF9B77CB65B2F4CBFE4B" lastPageId="26" lastPageNumber="243" pageId="25" pageNumber="242" type="description">
<paragraph id="F344E36C241BFF9877CB65B2F651F96B" blockId="25.[680,1229,1240,1744]" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9877CB65B2F636FB4B" box="[680,810,1240,1264]" italics="true" pageId="25" pageNumber="242">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9241BFF98761665B2F6D4FB4B" box="[885,968,1240,1264]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="25" pageNumber="242">Fig. 34J</figureCitation>
): Integument very bright, evenly light brown. Total length 8.0 mm; elytral base as wide as pronotum, elytra 2.62x longer than pronotum. Frons deeply concave along the anterior margin; frontal carina complete. Antennae surpassing the posterior angles by two antennomeres. Mandible unidentate. Prothorax 1.12x wider than long with sides sinuous converging abruptly anteriad on anterior half; lateral carina prominent posteriorly, directed ventrad anteriorly; posterior angles divergent and convex; pronotum weakly convex with punctures coalescent forming short and irregular longitudinal striae. Elytral interstices convex. Free margin of metacoxal plate short, triangular; tibiae slightly widened apicad, outer angle of protibiae truncate. Tergite IX with anterior margin straight.
</paragraph>
<paragraph id="F344E36C241BFF9B77CB6792F7B5FF4B" blockId="25.[680,1228,1784,1872]" lastBlockId="26.[132,681,184,240]" lastPageId="26" lastPageNumber="243" pageId="25" pageNumber="242">
<emphasis id="C18F3F7E241BFF9877CB6792F61DF8AB" box="[680,769,1784,1808]" italics="true" pageId="25" pageNumber="242">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9241BFF98766C6792F68DF8AB" box="[783,913,1784,1808]" captionStart="FIGURE 11" captionStartId="27.[95,174,1826,1845]" captionTargetBox="[95,1228,1274,1801]" captionTargetId="figure-182@27.[94,1228,1273,1802]" captionTargetPageId="27" captionText="FIGURE 11: Toxognathus dohertyi. A, B, aedeagus (dorsal, ventral). Toxognathus fairmairei. C, hypomeron; D, pronotum and head. Toxognathus mouhoti. E, pronotum; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641389" httpUri="https://zenodo.org/record/12641389/files/figure.png" pageId="25" pageNumber="242">Figs. 11A, B</figureCitation>
): Phallobase 0.42x total length of aedeagus, 1.2x longer than wide, length ratio between lateroposterior and median parts 6.1; parameres with ventral anteromedian margin acute, basal strut 0.33x total length of penis; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C2418FF9B75E76072F5E8FE0B" blockId="26.[132,681,280,432]" pageId="26" pageNumber="243">
<emphasis id="C18F3F7E2418FF9B75E76072F5FDFE8B" box="[132,225,280,304]" italics="true" pageId="26" pageNumber="243">
<typeStatus id="2C405DCE2418FF9B75E76072F5C0FE8B" box="[132,220,280,304]" pageId="26" pageNumber="243" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingRegion id="313F2D8E2418FF9B75906072F42DFE8B" box="[243,305,280,304]" country="Malaysia" name="Perak" pageId="26" pageNumber="243">Perak</collectingRegion>
,
<collectingCountry id="8BECA3FC2418FF9B74226072F489FE8B" box="[321,405,280,304]" name="Malaysia" pageId="26" pageNumber="243">Malacca</collectingCountry>
(Doherty], [
<taxonomicName id="34FB98EF2418FF9B77426072F43DFEEB" authority="Dohetyi Fleut." authorityName="Dohetyi Fleut." class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="243" phylum="Arthropoda" rank="genus">Toxognathus Dohetyi Fleut.</taxonomicName>
, Type, Collection Fleutiaux], [Type], [Muséum Paris, Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF2418FF9B77426032F5CEFE2B" authority="dohetyi" authorityName="Dohetyi" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="243" phylum="Arthropoda" rank="genus">Toxognathus dohetyi</taxonomicName>
(Fleutiaux), J. Chassain det. 05], [
<typeStatus id="2C405DCE2418FF9B77526012F782FE2B" box="[561,670,376,400]" pageId="26" pageNumber="243" type="lectotype">Lectotype</typeStatus>
], (MNHN).
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<paragraph id="F344E36C2418FF9B75E760B2F4CBFE4B" blockId="26.[132,471,472,496]" box="[132,471,472,496]" pageId="26" pageNumber="243">
<emphasis id="C18F3F7E2418FF9B75E760B2F41AFE4B" box="[132,262,472,496]" italics="true" pageId="26" pageNumber="243">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2418FF9B746E60B2F492FE4B" box="[269,398,472,496]" name="Malaysia" pageId="26" pageNumber="243">MALAYSIA</collectingCountry>
.
<collectingRegion id="313F2D8E2418FF9B74F460B2F4CFFE4B" box="[407,467,472,496]" country="Malaysia" name="Perak" pageId="26" pageNumber="243">Perak</collectingRegion>
.
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<emphasis id="C18F3F7E241EFF9D75A960FDF4ADFE14" bold="true" box="[202,433,407,431]" italics="true" pageId="28" pageNumber="245">Toxognathus mouhoti</emphasis>
Fleutiaux, 1918
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E-G; 34L)
</emphasis>
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<taxonomicName id="34FB98EF241EFF9D75E76092F77DFDAB" authority="Fleutiaux, 1918 a: 277" authorityName="Fleutiaux" authorityYear="1918" box="[132,609,504,528]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="C18F3F7E241EFF9D75E76092F443FDAB" box="[132,351,504,528]" italics="true" pageId="28" pageNumber="245">Toxognathus mouhoti</emphasis>
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</taxonomicName>
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;
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<bibRefCitation id="976A9E9D241EFF9D77106092F491FD8B" author="FLEUTIAUX, E." pageId="28" pageNumber="245" pagination="205 - 236" refId="ref49788" refString="FLEUTIAUX, E. 1918 b. Coleopteres Elaterides Indochinois de la collection du Museum d'Histoire Naturelle de Paris. Catalogue et description des especes nouvelles. Bulletin du Museum D'Histoire Naturelle, 24: 205 - 236." type="journal article" year="1918">Fleutiaux, 1918b: 236</bibRefCitation>
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,
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</treatmentCitation>
;
<treatmentCitation id="725AC57D241EFF9D774C6372F437FDEB" author="SCHENKLING, S." page="509" pageId="28" pageNumber="245" year="1927">
<bibRefCitation id="976A9E9D241EFF9D774C6372F437FDEB" author="SCHENKLING, S." pageId="28" pageNumber="245" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 509</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D241EFF9D74566352F711FDEB" author="FLEUTIAUX, E." box="[309,525,568,592]" page="35" pageId="28" pageNumber="245" year="1940">
<bibRefCitation id="976A9E9D241EFF9D74566352F711FDEB" author="FLEUTIAUX, E." box="[309,525,568,592]" pageId="28" pageNumber="245" pagination="19 - 40" refId="ref49922" refString="FLEUTIAUX, E. 1940 a. Les elaterides de l'Indo-chine francaise. Huitieme et derniere partie. Annales de La Societe Entomologique de France, 109: 19 - 40." type="journal article" year="1940">Fleutiaux, 1940a: 35</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241EFF9D75E76312F101FE8B" pageId="28" pageNumber="245" type="description">
<paragraph id="F344E36C241EFF9D75E76312F5F2FB2B" blockId="28.[132,681,632,1168]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D75E76312F41AFD2B" box="[132,262,632,656]" italics="true" pageId="28" pageNumber="245">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9241EFF9D74006312F4DBFD2B" box="[355,455,632,656]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="28" pageNumber="245">Fig. 34L</figureCitation>
): Integument evenly light reddish-brown or with antenommeres I-III yellow. Total length 5.0-6.0 mm; elytral base 1.08x as wide as pronotum, elytra 2.9-3.0x longer than pronotum. Frons flattened; frontal carina absent medially. Antennae surpassing the posterior angles of pronotum by 3.5-5.0 antennomeres. Mandible bidentate. Pronotum (
<figureCitation id="6BC0FFE9241EFF9D759E6232F44BFCCB" box="[253,343,856,880]" captionStart="FIGURE 11" captionStartId="27.[95,174,1826,1845]" captionTargetBox="[95,1228,1274,1801]" captionTargetId="figure-182@27.[94,1228,1273,1802]" captionTargetPageId="27" captionText="FIGURE 11: Toxognathus dohertyi. A, B, aedeagus (dorsal, ventral). Toxognathus fairmairei. C, hypomeron; D, pronotum and head. Toxognathus mouhoti. E, pronotum; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641389" httpUri="https://zenodo.org/record/12641389/files/figure.png" pageId="28" pageNumber="245">Fig. 11E</figureCitation>
) weakly convex 1.2x wider than long with sides slightly rounded from base of posterior angles to anterior margin, convergent on anterior 1/4, lateral carina entirely lateroventral, posterior angles subparallel and weakly convex; punctures less than 0.5 diameter apart. Elytral interstices convex. Free margin of metacoxal plate short and triangular; tibiae slightly widened apicad, dorsal apical angle of protibiae truncate. Tergite IX with anterior margin bisinuous.
</paragraph>
<paragraph id="F344E36C241EFF9D75E765D2F427FACB" blockId="28.[132,681,1208,1392]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D75E765D2F5C1FB6B" box="[132,221,1208,1232]" italics="true" pageId="28" pageNumber="245">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9241EFF9D758865D2F470FB6B" box="[235,364,1208,1232]" captionStart="FIGURE 11" captionStartId="27.[95,174,1826,1845]" captionTargetBox="[95,1228,1274,1801]" captionTargetId="figure-182@27.[94,1228,1273,1802]" captionTargetPageId="27" captionText="FIGURE 11: Toxognathus dohertyi. A, B, aedeagus (dorsal, ventral). Toxognathus fairmairei. C, hypomeron; D, pronotum and head. Toxognathus mouhoti. E, pronotum; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641389" httpUri="https://zenodo.org/record/12641389/files/figure.png" pageId="28" pageNumber="245">Figs. 11F, G</figureCitation>
): Phallobase 0.39x total length of aedeagus, 1.31x longer than wide, length ratio between lateroposterior and median parts 3.82 longer than median part; paramere with anteromedian margin acute, basal strut 0.35x total length of penis; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C241EFF9D75E764F2F4E2F98B" blockId="28.[132,681,1432,1584]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D75E764F2F5FDFA0B" box="[132,225,1432,1456]" italics="true" pageId="28" pageNumber="245">
<typeStatus id="2C405DCE241EFF9D75E764F2F5C0FA0B" box="[132,220,1432,1456]" pageId="28" pageNumber="245" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Chatebon Muhot], [Collection W.W. Saunders], [
<taxonomicName id="34FB98EF241EFF9D746464D2F73AFA6B" authority=", Flet." authorityName="Flet." box="[263,550,1464,1488]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="C18F3F7E241EFF9D746464D2F4F5FA6B" box="[263,489,1464,1488]" italics="true" pageId="28" pageNumber="245">Toxognathus mouhoti,</emphasis>
Flet.
</taxonomicName>
Type, Collection Fleutiaux], [Type], [Muséum Paris, Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF241EFF9D74676492F5B0F98B" authority="(Fleut.) J. Chassain" authorityName="J. Chassain" baseAuthorityName="Fleut." class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="species" species="mouhoti">Toxognathus mouhoti (Fleut.) J. Chassain</taxonomicName>
det. 05], [
<typeStatus id="2C405DCE241EFF9D74746772F49EF98B" box="[279,386,1560,1584]" pageId="28" pageNumber="245" type="lectotype">Lectotype</typeStatus>
] (MNHN).
</paragraph>
<paragraph id="F344E36C241EFF9D75E76733F7B4F8EB" blockId="28.[132,681,1624,1872]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D75E76733F5FAF9CB" box="[132,230,1625,1648]" italics="true" pageId="28" pageNumber="245">
<typeStatus id="2C405DCE241EFF9D75E76733F5FCF9CB" box="[132,224,1625,1648]" pageId="28" pageNumber="245" type="paratype">Paratypes</typeStatus>
:
</emphasis>
[Camboj. int. Muhot], [Collection W.W. Saunders], [Muséum Paris, Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF241EFF9D771F6713F718F90B" authority="(Fleut.) J. Chassain" authorityName="J. Chassain" baseAuthorityName="Fleut." class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="C18F3F7E241EFF9D771F6713F429F90B" italics="true" pageId="28" pageNumber="245">Toxognathus mouhoti</emphasis>
(Fleut.) J. Chassain
</taxonomicName>
det. 05], [
<typeStatus id="2C405DCE241EFF9D771267F2F5FAF96B" pageId="28" pageNumber="245" type="paralectotype">Paralectotype</typeStatus>
],
<specimenCount id="E5FD28E5241EFF9D759867D2F45AF96B" box="[251,326,1720,1744]" count="1" pageId="28" pageNumber="245" type="male">1 male</specimenCount>
, (MNHN); [Museum Paris, Cap St. Jacques, Coll. A. Bonhoure, 1909], [
<taxonomicName id="34FB98EF241EFF9D774D67B2F4DCF8AB" authority="Fleut. 1917 Fleutiaux" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="C18F3F7E241EFF9D774D67B2F5CAF8AB" italics="true" pageId="28" pageNumber="245">Toxognathus mouhoti</emphasis>
Fleut. 1917 Fleutiaux
</taxonomicName>
det.], [Muséum Paris, Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF241EFF9D74006672F5FCF8EB" authority="(Fleut.) J. Chassain" authorityName="J. Chassain" baseAuthorityName="Fleut." class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="C18F3F7E241EFF9D74006672F724F88B" box="[355,568,1816,1840]" italics="true" pageId="28" pageNumber="245">Toxognathus mouhoti</emphasis>
(Fleut.) J. Chassain
</taxonomicName>
det. 05], [
<typeStatus id="2C405DCE241EFF9D74286652F4C0F8EB" box="[331,476,1848,1872]" pageId="28" pageNumber="245" type="paralectotype">Paralectotype</typeStatus>
],
<specimenCount id="E5FD28E5241EFF9D748F6652F72DF8EB" box="[492,561,1848,1872]" count="1" pageId="28" pageNumber="245" type="male">1 male</specimenCount>
(MNHN).
</paragraph>
<paragraph id="F344E36C241EFF9D77AD61D2F1D9FF6B" blockId="28.[718,1221,184,208]" box="[718,1221,184,208]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D77AD61D2F68CFF6B" box="[718,912,184,208]" italics="true" pageId="28" pageNumber="245">Material examined:</emphasis>
<materialsCitation id="4393E931241EFF9D76F461D2F1DEFF6B" box="[919,1218,184,208]" collectionCode="MNHN" country="Vietnam" location="VIETNAM" pageId="28" pageNumber="245" specimenCount="1">
<collectingCountry id="8BECA3FC241EFF9D76F461D2F109FF6B" box="[919,1045,184,208]" name="Vietnam" pageId="28" pageNumber="245">VIETNAM</collectingCountry>
.
<specimenCount id="E5FD28E5241EFF9D717D61D2F14CFF74" box="[1054,1104,184,208]" count="1" pageId="28" pageNumber="245" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9241EFF9D713E61D2F1A5FF6B" box="[1117,1209,184,208]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="245">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C241EFF9D77AD6192F101FE8B" blockId="28.[718,1266,248,304]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D77AD6192F64CFEAB" box="[718,848,248,272]" italics="true" pageId="28" pageNumber="245">Distribution:</emphasis>
<collectingCountry id="8BECA3FC241EFF9D76396192F6F4FEAB" box="[858,1000,248,272]" name="Thailand" pageId="28" pageNumber="245">THAILAND</collectingCountry>
.
<emphasis id="C18F3F7E241EFF9D76956192F163FEAB" box="[1014,1151,248,272]" italics="true" pageId="28" pageNumber="245">
<collectingRegion id="313F2D8E241EFF9D76956192F167FEAB" box="[1014,1147,248,272]" country="Thailand" name="Chanthaburi" pageId="28" pageNumber="245">Chanthaburi</collectingRegion>
.
</emphasis>
CAMBO- DIA.
<collectingCountry id="8BECA3FC241EFF9D766A6072F69BFE8B" box="[777,903,280,304]" name="Vietnam" pageId="28" pageNumber="245">VIETNAM</collectingCountry>
.
<emphasis id="C18F3F7E241EFF9D76F26073F6C2FE8B" box="[913,990,281,304]" italics="true" pageId="28" pageNumber="245">Ba Ria-</emphasis>
Vung.
</paragraph>
</subSubSection>
</treatment>
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<heading id="A80C5400241EFF9D7659601DF199FE2B" bold="true" box="[826,1157,375,400]" centered="true" fontSize="10" level="2" pageId="28" pageNumber="245" reason="2">
<taxonomicName id="34FB98EF241EFF9D7659601DF199FE2B" authority="Fleutiaux, 1892" authorityName="Fleutiaux" authorityYear="1892" box="[826,1157,375,400]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E241EFF9D7659601DF199FE2B" bold="true" box="[826,1157,375,400]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D7659601DF6C8FE34" bold="true" box="[826,980,375,399]" italics="true" pageId="28" pageNumber="245">Dactylophysus</emphasis>
<bibRefCitation id="976A9E9D241EFF9D76B86012F199FE2B" author="FLEUTIAUX, E." box="[987,1157,376,400]" pageId="28" pageNumber="245" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="BBE1B0E7241EFF9D77AD60D2F64DFD8B" pageId="28" pageNumber="245" type="reference_group">
<paragraph id="F344E36C241EFF9D77AD60D2F164FE6B" blockId="28.[718,1267,440,560]" box="[718,1144,440,464]" pageId="28" pageNumber="245">
<treatmentCitationGroup id="D3EBC442241EFF9D77AD60D2F164FE6B" box="[718,1144,440,464]" pageId="28" pageNumber="245">
<treatmentCitation id="725AC57D241EFF9D77AD60D2F12CFE6B" author="CANDEZE, E." box="[718,1072,440,464]" page="31" pageId="28" pageNumber="245" year="1859">
<taxonomicName id="34FB98EF241EFF9D77AD60D2F12CFE6B" authority="Candeze, 1859: 31" authorityName="Candeze" authorityPageNumber="31" authorityYear="1859" box="[718,1072,440,464]" class="Insecta" family="Elateridae" genus="Heterocrepidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E241EFF9D77AD60D2F67AFE6B" box="[718,870,440,464]" italics="true" pageId="28" pageNumber="245">Heterocrepidius</emphasis>
<bibRefCitation id="976A9E9D241EFF9D760E60D2F12CFE6B" author="CANDEZE, E." box="[877,1072,440,464]" pageId="28" pageNumber="245" refId="ref49242" refString="CANDEZE, E. 1859. Monographie des Elaterides. Tome second. Liege, H. Dessain. 543 p." type="book" year="1859">Candèze, 1859: 31</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(pars).
</treatmentCitationGroup>
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<paragraph id="F344E36C241EFF9D77AD60B2F64DFD8B" blockId="28.[718,1267,440,560]" pageId="28" pageNumber="245">
<treatmentCitationGroup id="D3EBC442241EFF9D77AD60B2F64DFD8B" pageId="28" pageNumber="245">
<treatmentCitation id="725AC57D241EFF9D77AD60B2F15FFE4B" author="FLEUTIAUX, E." box="[718,1091,472,496]" page="408" pageId="28" pageNumber="245" year="1892">
<taxonomicName id="34FB98EF241EFF9D77AD60B2F15FFE4B" authority="Fleutiaux, 1892: 408" authorityName="Fleutiaux" authorityPageNumber="408" authorityYear="1892" box="[718,1091,472,496]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E241EFF9D77AD60B2F64BFE4B" box="[718,855,472,496]" italics="true" pageId="28" pageNumber="245">Dactylophysus</emphasis>
<bibRefCitation id="976A9E9D241EFF9D760260B2F15FFE4B" author="FLEUTIAUX, E." box="[865,1091,472,496]" pageId="28" pageNumber="245" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 408</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="725AC57D241EFF9D713360B2F62DFDAB" author="SCHWARZ, O." page="311" pageId="28" pageNumber="245" year="1906">
<bibRefCitation id="976A9E9D241EFF9D713360B2F62DFDAB" author="SCHWARZ, O." pageId="28" pageNumber="245" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 311</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D241EFF9D76236092F122FDAB" author="BLACKWELDER, R. E." box="[832,1086,504,528]" page="292" pageId="28" pageNumber="245" year="1944">
<bibRefCitation id="976A9E9D241EFF9D76236092F122FDAB" author="BLACKWELDER, R. E." box="[832,1086,504,528]" pageId="28" pageNumber="245" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D241EFF9D712E6092F638FD8B" author="GOLBACH, R." page="27" pageId="28" pageNumber="245" year="1994">
<bibRefCitation id="976A9E9D241EFF9D712E6092F638FD8B" author="GOLBACH, R." pageId="28" pageNumber="245" pagination="1 - 48" refId="ref50116" refString="GOLBACH, R. 1994. Elateridae (Coleoptera) de la Argentina. Historia, catalogo actualizado hasta 1991 inclusive y clave de subfamilias y de generos de Centro y Sudamerica. Opera Lilloana, 41: 1 - 48." type="journal article" year="1994">Golbach, 1994: 27</bibRefCitation>
</treatmentCitation>
, 40.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7241EFF9F77AD6333F607FC0B" lastPageId="30" lastPageNumber="247" pageId="28" pageNumber="245" type="materials_examined">
<paragraph id="F344E36C241EFF9D77AD6333F1D9FD6B" blockId="28.[718,1266,600,720]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D77AD6333F657FDCB" box="[718,843,601,624]" italics="true" pageId="28" pageNumber="245">
<typeStatus id="2C405DCE241EFF9D77AD6333F7E7FDCB" box="[718,763,601,624]" pageId="28" pageNumber="245">Type</typeStatus>
species:
</emphasis>
Now fixed (under Article 70.3 of the Code) as
<taxonomicName id="34FB98EF241EFF9D76566312F15CFD2B" authorityName="Rosa" authorityYear="2014" box="[821,1088,632,656]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="species" species="capixabensis" status="nom. nov.">
<emphasis id="C18F3F7E241EFF9D76566312F15CFD2B" box="[821,1088,632,656]" italics="true" pageId="28" pageNumber="245">Dactylophysus capixabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205241EFF9D712A6312F1A4FD34" box="[1097,1208,632,655]" pageId="28" pageNumber="245">nom. nov.</taxonomicNameLabel>
, misidentified as
<taxonomicName id="34FB98EF241EFF9D763563F2F1F2FD0B" authority="Candeze, 1859" authorityName="Candeze" authorityYear="1859" box="[854,1262,664,688]" class="Insecta" family="Elateridae" genus="Heterocrepidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="245" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="C18F3F7E241EFF9D763563F2F159FD0B" box="[854,1093,664,688]" italics="true" pageId="28" pageNumber="245">Heterocrepidius mendax</emphasis>
<bibRefCitation id="976A9E9D241EFF9D712D63F2F1F2FD0B" author="CANDEZE, E." box="[1102,1262,664,688]" pageId="28" pageNumber="245" refId="ref49242" refString="CANDEZE, E. 1859. Monographie des Elaterides. Tome second. Liege, H. Dessain. 543 p." type="book" year="1859">Candèze, 1859</bibRefCitation>
</taxonomicName>
, in the subsequent designation by
<bibRefCitation id="976A9E9D241EFF9D714563D2F1DCFD6B" author="HYSLOP, J. A." box="[1062,1216,696,720]" pageId="28" pageNumber="245" pagination="621 - 680" refId="ref50326" refString="HYSLOP, J. A. 1921. Genotypes of the Elaterid beetles of the world. Proceedings of United States of National Museum, 58: 621 - 680." type="journal article" year="1921">Hyslop (1921)</bibRefCitation>
.
</paragraph>
<paragraph id="F344E36C241EFF9D77AD6393F613FB0B" blockId="28.[718,1267,760,1200]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D77AD6393F62FFCAB" box="[718,819,761,784]" italics="true" pageId="28" pageNumber="245">Diagnosis:</emphasis>
Body weakly convex, antenna flattened reaching the posterior angles of pronotum, labrum evenly convex, mandible unidentate; pronotum trapezoidal wider than long with complete lateral carina weakly impressed anteriorly, directed anteroventrally; anterior angles acute in dorsal view. Metacoxa 30° oblique to the transverse body axis with inner 1/4 2.5-3.5x longer than the outer 3/4; free margin of metacoxal plate short and narrow, triangular; tibiae widened apicad with each lateral border of apex with a row of 12-14 spiniform setae, pro- and mesofemur laterally flat, metafemur weakly convex; last abdominal ventrite V-shaped weakly convex apically or flattened.
</paragraph>
<paragraph id="F344E36C241EFF9D77AD65B2F124FB4B" blockId="28.[718,1080,1240,1264]" box="[718,1080,1240,1264]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D77AD65B2F64CFB4B" box="[718,848,1240,1264]" italics="true" pageId="28" pageNumber="245">Distribution:</emphasis>
SOUTH AMERICA.
</paragraph>
<paragraph id="F344E36C241EFF9D77AD6472F607FA74" blockId="28.[718,1267,1304,1872]" pageId="28" pageNumber="245">
<emphasis id="C18F3F7E241EFF9D77AD6472F64CFA8B" box="[718,848,1304,1328]" italics="true" pageId="28" pageNumber="245">Redescription</emphasis>
(male): Integument bright, light brown to black. Punctures umbilicate deeply impressed, densely covered with yellow setae; pilosity decumbent or erect. Total length: 9.0-
<quantity id="34034E89241EFF9D76876412F128FA34" box="[996,1076,1400,1424]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="245" unit="mm" value="12.0">12 mm</quantity>
; elytral base 1x as wide as prothorax, elytra 2.6-3.3x longer than pronotum.
</paragraph>
<paragraph id="F344E36C241EFF9C766464B2F4E8FCAB" blockId="28.[718,1267,1304,1872]" lastBlockId="29.[94,643,184,1520]" lastPageId="29" lastPageNumber="246" pageId="28" pageNumber="245">
Head (
<figureCitation id="6BC0FFE9241EFF9D763764B2F6A6FA4B" box="[852,954,1496,1520]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="28" pageNumber="245">Figs. 12F</figureCitation>
,
<figureCitation id="6BC0FFE9241EFF9D76AF64B2F6E4FA4B" box="[972,1016,1496,1520]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="28" pageNumber="245">15B</figureCitation>
) with anterior margin of frons straight or rounded, dorsal ocular margins curved inwards; frontal carina complete, weakly produced anteriad (
<figureCitation id="6BC0FFE9241EFF9D76D36752F101F9EB" box="[944,1053,1592,1616]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="28" pageNumber="245">Fig. 12H</figureCitation>
); frontoclypeal region (
<figureCitation id="6BC0FFE9241EFF9D766E6732F672F9CB" box="[781,878,1624,1648]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="28" pageNumber="245">Fig. 12G</figureCitation>
) steeply declivous to base of labrum 5.0-6.0x wider than long; punctures as large as those of pronotum, 0.2-0.5 diameter apart. Antenna (
<figureCitation id="6BC0FFE9241EFF9D77B667D2F621F96B" box="[725,829,1720,1744]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="28" pageNumber="245">Figs. 12A</figureCitation>
,
<figureCitation id="6BC0FFE9241EFF9D762867D2F664F96B" box="[843,888,1720,1744]" captionStart="FIGURE 14" captionStartId="33.[95,174,950,969]" captionTargetBox="[96,1225,170,925]" captionTargetId="figure-410@33.[94,1228,170,926]" captionTargetPageId="33" captionText="FIGURE 14: Dactylophysus capixabensis nom. nov. A, antenna; B, pronotum; C, D, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641397" httpUri="https://zenodo.org/record/12641397/files/figure.png" pageId="28" pageNumber="245">14A</figureCitation>
,
<figureCitation id="6BC0FFE9241EFF9D76E567D2F6AEF96B" box="[902,946,1720,1744]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="28" pageNumber="245">15A</figureCitation>
) with 11 antenomeres, antennomeres
<collectionCode id="95EA7BA9241EFF9D765267B2F65FF954" box="[817,835,1752,1775]" pageId="28" pageNumber="245">II</collectionCode>
and
<collectionCode id="95EA7BA9241EFF9D76E067B2F682F954" box="[899,926,1752,1775]" pageId="28" pageNumber="245">III</collectionCode>
subequal,
<collectionCode id="95EA7BA9241EFF9D717A67B2F128F954" box="[1049,1076,1752,1775]" pageId="28" pageNumber="245">III</collectionCode>
1.0-1.16x longer than
<collectionCode id="95EA7BA9241EFF9D766B6792F600F8B4" box="[776,796,1784,1807]" pageId="28" pageNumber="245">II</collectionCode>
; IV-XI laterally flattened longer than wide; IV-X serrate, IV-VII 1.3-1.6x longer than wide, VIII 1.6-1.9x longer than wide, IX-X 1.5-2.0 longer than wide, XI elliptical abruptly narrowed at apex 2.5-2.8x longer than wide.
<collectionCode id="95EA7BA9241FFF9C744561B2F451FF54" box="[294,333,216,239]" pageId="29" pageNumber="246">IEP</collectionCode>
0.4-0.5. Labrum (
<figureCitation id="6BC0FFE9241FFF9C777961B2F767FF4B" box="[538,635,216,240]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12C</figureCitation>
) evenly convex, subrectangular, 2.5-3.0x wider than long, anterior margin slightly emarginate to straight, coarsely punctate, with long setae. Mouthparts directed anteroventrally. Mandible narrow, unidentate, laterodorsal face densely punctate; lateral edge evenly curved apicad; mesoapical margin evenly curved (
<figureCitation id="6BC0FFE9241FFF9C750560D2F5DFFE6B" box="[102,195,440,464]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="29" pageNumber="246">Fig. 15B</figureCitation>
) or sinuous (
<figureCitation id="6BC0FFE9241FFF9C743060D2F4AEFE6B" box="[339,434,440,464]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12B</figureCitation>
), mesobasal margin with a row of short setae on a prominent rectangular membranous area (
<figureCitation id="6BC0FFE9241FFF9C742A6092F4B4FDAB" box="[329,424,504,528]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="29" pageNumber="246">Fig. 15B</figureCitation>
). Maxilla (
<figureCitation id="6BC0FFE9241FFF9C77796092F767FDAB" box="[538,635,504,528]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12D</figureCitation>
) with galea securiform, anterior border covered with short fine setae, denser on inner angle; lacinia elongate, tongue-like, densely pilose; medistipes trapezoidal longer than wide with several long setae; labium (
<figureCitation id="6BC0FFE9241FFF9C750563F2F5A2FD0B" box="[102,190,664,688]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12E</figureCitation>
) with prementum shortly emarginate on anteromedian margin, anterolateral angles rounded and not produced; maxillary and labial palpi pilose with fine setae, apical palpomere securiform.
</paragraph>
<paragraph id="F344E36C241FFF9C75F46272F49CFA54" blockId="29.[94,643,184,1520]" pageId="29" pageNumber="246">
Pronotum (
<figureCitation id="6BC0FFE9241FFF9C74766272F464FC8B" box="[277,376,792,816]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Figs. 12J</figureCitation>
,
<figureCitation id="6BC0FFE9241FFF9C74E96272F4ABFC8B" box="[394,439,792,816]" captionStart="FIGURE 14" captionStartId="33.[95,174,950,969]" captionTargetBox="[96,1225,170,925]" captionTargetId="figure-410@33.[94,1228,170,926]" captionTargetPageId="33" captionText="FIGURE 14: Dactylophysus capixabensis nom. nov. A, antenna; B, pronotum; C, D, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641397" httpUri="https://zenodo.org/record/12641397/files/figure.png" pageId="29" pageNumber="246">14B</figureCitation>
,
<figureCitation id="6BC0FFE9241FFF9C74AB6272F4EAFC8B" box="[456,502,792,816]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="29" pageNumber="246">15C</figureCitation>
) trapezoidal, flattened, 1.18-1.35x wider than long, lateral carina reaching the anterior margin, lateral on posterior region and directed ventrad anteriorly (
<figureCitation id="6BC0FFE9241FFF9C74856212F758FC2B" box="[486,580,888,912]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12K</figureCitation>
) with anterior part not visible dorsally; punctures homogeneously distributed 0.2-0.5 diameters apart, a little smaller on posterior margin than at disc; anterior angles acute; posterior angles of pronotum non-carinate. Hypomeron (
<figureCitation id="6BC0FFE9241FFF9C74556572F489FB8B" box="[310,405,1048,1072]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12L</figureCitation>
) with punctures usually as large as those of pronotum, denser anteriorly; posterior 1/5-1/2 glabrous; posterior margin with a U- or]-shaped notch adjacent to the posterior angle. Prosternal suture (
<figureCitation id="6BC0FFE9241FFF9C747B65F2F468FB0B" box="[280,372,1176,1200]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Figs. 12I</figureCitation>
,
<figureCitation id="6BC0FFE9241FFF9C741C65F2F4B2FB0B" box="[383,430,1176,1200]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="29" pageNumber="246">15D</figureCitation>
) curved, marginated by a shiny band along hypomeral margin, grooved at anterior part. Prosternum (
<figureCitation id="6BC0FFE9241FFF9C741B65B2F4D0FB4B" box="[376,460,1240,1264]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12I</figureCitation>
) 1.1-1.32x wider than long, with punctures larger than those of pronotum, 0.2-1.5 diameters apart; anterior lobe edge straight or rounded, produced, covering mouth parts to labial palpiger. Prosternal process (
<figureCitation id="6BC0FFE9241FFF9C74966432F767FACB" box="[501,635,1368,1392]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Figs. 12I, K</figureCitation>
) 2.30-2.83x longer than diameter of procoxae, ventral surface narrower than the dorsal surface with a subapical tooth, about 45° declivous dorsad posteriorad of procoxae. Procoxae open.
</paragraph>
<paragraph id="F344E36C241FFF9C753D6772F665FD4B" blockId="29.[94,643,1560,1872]" lastBlockId="29.[680,1229,184,1872]" pageId="29" pageNumber="246">
<emphasis id="C18F3F7E241FFF9C753D6772F5CAF98B" box="[94,214,1560,1584]" italics="true" pageId="29" pageNumber="246">Pterothorax:</emphasis>
Mesoventrite (
<figureCitation id="6BC0FFE9241FFF9C74176772F4DAF98B" box="[372,454,1560,1584]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Figs. 12</figureCitation>
M-O) with posterior region inclined about 30° above of the anterior region (
<figureCitation id="6BC0FFE9241FFF9C75CC6732F40DF9CB" box="[175,273,1624,1648]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12N</figureCitation>
), with anterior articulating surfaces concave, smooth, bordered posteriorly by a prominent carina; borders of mesoventral cavity curved and convergent anteriorly from its half length; floor of cavity with a shiny median band. Mesocoxal cavity (
<figureCitation id="6BC0FFE9241FFF9C75056792F5D0F8AB" box="[102,204,1784,1808]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12M</figureCitation>
) open to both mesepisternum and mesepimeron, mesotrochantin visible; mesepisternum with an oblique carina anteromedially contiguous to the anterior carina of mesoventrite. Meso-metaventral suture distinct. Metaventrite (
<figureCitation id="6BC0FFE9241FFF9C76A561B2F13AFF4B" box="[966,1062,216,240]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12O</figureCitation>
) 1.3-1.4x wider than long, 1.4x longer than mesoventrite; metepisternum 6.5-8.0x longer than wide. Scutellum abruptly elevated above the level of mesoscutum, pentagonal, 1.3-1.4x longer than wide, anterior margin rounded. Metanotum (
<figureCitation id="6BC0FFE9241FFF9C76516012F693FE2B" box="[818,911,376,400]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12P</figureCitation>
): prescutum contiguous medially to the scutum by an acute process; posterior part of the scutellum with a longitudinal apodeme. Elytra flattened with apices conjointly rounded, striae with a single row of punctures increasing in size posteriorly; interstices flat, rugose with punctures 1-2 diameters apart; epipleurae (
<figureCitation id="6BC0FFE9241FFF9C76036352F6A2FDEB" box="[864,958,568,592]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12Q</figureCitation>
) narrowed near metacoxa, slightly widened apicad from ventrite 2. Hind wings with radial cell 4.2x longer than wide, CuA
<subScript id="6F7FE129241FFF9C710F63EDF168FD2E" attach="left" box="[1132,1140,647,661]" fontSize="6" pageId="29" pageNumber="246">1</subScript>
present; wedge cell present or absent; apex with anterior, median and posterior field sclerotizations convergent basad, not contiguous.
</paragraph>
<paragraph id="F344E36C241FFF9C77826392F1B7FA6B" blockId="29.[680,1229,184,1872]" pageId="29" pageNumber="246">
Metacoxa (
<figureCitation id="6BC0FFE9241FFF9C76376392F6A5FCAB" box="[852,953,760,784]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12M</figureCitation>
) inclined about 30° in relation to the transverse axis of body; inner quarter 2.5-3.5x longer than outer 3/4; ventral part with inner third about 11x longer than the outer 2/3; free margin of metacoxal plate short and narrow, triangular. Tibiae (Figs. 38A-C) with 12-14 spiniform setae along each outer and inner inner apical border (Fig. 38D), dorsal margin covered with 3-4 irregular rows of spiniform setae, outer and inner surface covered with fine setae. Protibia (
<figureCitation id="6BC0FFE9241FFF9C76146572F6C8FB8B" box="[887,980,1048,1072]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12A</figureCitation>
) 2.6-3.0x wider at apex than at base, with dorsal margin rugose, straight to sinuous with apical angle acute. Mesotibia (
<figureCitation id="6BC0FFE9241FFF9C71096532F1D8FBCB" box="[1130,1220,1112,1136]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12B</figureCitation>
) 2.8-3.2x wider at apex than at base, with dorsal apical angle acute. Metatibia (
<figureCitation id="6BC0FFE9241FFF9C76A765F2F13FFB0B" box="[964,1059,1176,1200]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12C</figureCitation>
) 2.3-2.4x wider at apex than at base with apical dorsal angle acute. Pro- and mesofemur subrectangular and laterally flattened; metafemur weakly convex, metatrochanter as convex as metafemur. Tarsomeres decreasing in length from I-IV,
<collectionCode id="95EA7BA9241FFF9C767A6452F637FAF4" box="[793,811,1336,1359]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="29" pageNumber="246" type="Museum">V</collectionCode>
longer than
<collectionCode id="95EA7BA9241FFF9C76D26452F6D0FAF4" box="[945,972,1336,1359]" pageId="29" pageNumber="246">III</collectionCode>
and
<collectionCode id="95EA7BA9241FFF9C71626452F100FAF4" box="[1025,1052,1336,1359]" pageId="29" pageNumber="246">IV</collectionCode>
together; ventral surface (
<figureCitation id="6BC0FFE9241FFF9C76636432F647FACB" box="[768,859,1368,1392]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12E</figureCitation>
) densely pilose, tarsomeres I-III or II-III lamellate, lamella on tarsomeres II-III subequal, on protarsomere I-II a little smaller or minuscule, on metatarsomere
<collectionCode id="95EA7BA9241FFF9C762564D2F67DFA74" box="[838,865,1464,1487]" pageId="29" pageNumber="246">III</collectionCode>
present or absent; claws simple.
</paragraph>
<paragraph id="F344E36C241FFF9F778264B2F431FCB4" blockId="29.[680,1229,184,1872]" lastBlockId="30.[132,681,184,784]" lastPageId="30" lastPageNumber="247" pageId="29" pageNumber="246">
Abdominal ventrites (
<figureCitation id="6BC0FFE9241FFF9C76A064B2F13CFA4B" box="[963,1056,1496,1520]" captionStart="FIGURE 12" captionStartId="31.[95,174,1775,1794]" captionTargetBox="[95,1225,565,1747]" captionTargetId="figure-138@31.[94,1228,561,1752]" captionTargetPageId="31" captionText="FIGURE 12: Dactylophysus hirtus sp. nov. (paratype). A, antenna; B, mandible; C, labrum; D, maxilla; E, labium; F, G, H, head (dorsal, anterior, lateral); I, J, K, prothorax (ventral, dorsal, lateral); L, hypomeron; M, mesoventrite and pleural sclerites (lateral); N, mesoventrite and mesocoxal cavity (lateroventral); O, pterothorax (ventral); P, dorsal pterothorax and abdominal tergite I; Q, epipleura; R, abdomen (ventral). Abbreviation: fmp, free margin of metacoxal plate." figureDoi="http://doi.org/10.5281/zenodo.12641393" httpUri="https://zenodo.org/record/12641393/files/figure.png" pageId="29" pageNumber="246">Fig. 12R</figureCitation>
) weakly convex, evenly covered with dense and decumbent pilosity, short to moderately long on ventrites 2-5, shorter on ventrite 1, ventrite 5 with apex coarsely punctate with pilosity denser and stouter; posterior angle of ventrite 1 and lateral margins of ventrites 2-4 with prominent marginal plates. Ventrite 1 with lateral part 4-6x longer than the median part, 2-4 subequal in length; ventrite 5 triangular 1.5-1.61x wider than long, flattened to weakly convex apically. Pregenitalic segments and aedeagus covered with yellow setae. Sternite VIII (
<figureCitation id="6BC0FFE9241FFF9C76156652F6FDF8EB" box="[886,993,1848,1872]" captionStart="FIGURE 13" captionStartId="32.[132,211,761,780]" captionTargetBox="[136,1262,170,734]" captionTargetId="figure-384@32.[132,1266,170,737]" captionTargetPageId="32" captionText="FIGURE 13: Dactylophysus hirtus sp. nov. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral). Abbreviation: apd, apodeme." figureDoi="http://doi.org/10.5281/zenodo.12641395" httpUri="https://zenodo.org/record/12641395/files/figure.png" pageId="29" pageNumber="246">Figs. 13A</figureCitation>
,
<figureCitation id="6BC0FFE9241FFF9C76916652F101F8EB" box="[1010,1053,1848,1872]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="29" pageNumber="246">15E</figureCitation>
) subrectangular, posterior margin straight, partly covered with short setae, lateroposterior angles with long stout setae, with two lateral and one median sclerotizations posteriorly; anterior border with or without a linear transverse sclerotization; tergite VIII (
<figureCitation id="6BC0FFE9241CFF9F77646052F770FEEB" box="[519,620,312,336]" captionStart="FIGURE 13" captionStartId="32.[132,211,761,780]" captionTargetBox="[136,1262,170,734]" captionTargetId="figure-384@32.[132,1266,170,737]" captionTargetPageId="32" captionText="FIGURE 13: Dactylophysus hirtus sp. nov. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral). Abbreviation: apd, apodeme." figureDoi="http://doi.org/10.5281/zenodo.12641395" httpUri="https://zenodo.org/record/12641395/files/figure.png" pageId="30" pageNumber="247">Figs. 13B</figureCitation>
,
<figureCitation id="6BC0FFE9241CFF9F77146052F7BCFEEB" box="[631,672,312,336]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="30" pageNumber="247">15F</figureCitation>
) evenly sclerotized U-shaped with long setae on lateroposterior and apical margin; sternite and tergite IX (
<figureCitation id="6BC0FFE9241CFF9F75D360F2F40DFE0B" box="[176,273,408,432]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="30" pageNumber="247">Fig. 15G</figureCitation>
) fused near the anterior margin; sternite IX (
<figureCitation id="6BC0FFE9241CFF9F758360D2F45CFE6B" box="[224,320,440,464]" captionStart="FIGURE 13" captionStartId="32.[132,211,761,780]" captionTargetBox="[136,1262,170,734]" captionTargetId="figure-384@32.[132,1266,170,737]" captionTargetPageId="32" captionText="FIGURE 13: Dactylophysus hirtus sp. nov. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral). Abbreviation: apd, apodeme." figureDoi="http://doi.org/10.5281/zenodo.12641395" httpUri="https://zenodo.org/record/12641395/files/figure.png" pageId="30" pageNumber="247">Fig. 13A</figureCitation>
,
<figureCitation id="6BC0FFE9241CFF9F742E60D2F462FE6B" box="[333,382,440,464]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="30" pageNumber="247">15H</figureCitation>
) with anteromedian margin rounded, produced and curved dorsad; anterolateral margins strongly sclerotized, anteromedian surface membranous, posterior surface sclerotized covered with short setae, setae long on lateroposterior and apical margin; tergite IX (
<figureCitation id="6BC0FFE9241CFF9F740C6332F4C4FDCB" box="[367,472,600,624]" captionStart="FIGURE 13" captionStartId="32.[132,211,761,780]" captionTargetBox="[136,1262,170,734]" captionTargetId="figure-384@32.[132,1266,170,737]" captionTargetPageId="32" captionText="FIGURE 13: Dactylophysus hirtus sp. nov. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral). Abbreviation: apd, apodeme." figureDoi="http://doi.org/10.5281/zenodo.12641395" httpUri="https://zenodo.org/record/12641395/files/figure.png" pageId="30" pageNumber="247">Figs. 13D</figureCitation>
,
<figureCitation id="6BC0FFE9241CFF9F74876332F70FFDCB" box="[484,531,600,624]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="30" pageNumber="247">15G</figureCitation>
) with anterior margin straight, apical lobes rounded with long setae; tergite X (
<figureCitation id="6BC0FFE9241CFF9F759263F2F440FD0B" box="[241,348,664,688]" captionStart="FIGURE 13" captionStartId="32.[132,211,761,780]" captionTargetBox="[136,1262,170,734]" captionTargetId="figure-384@32.[132,1266,170,737]" captionTargetPageId="32" captionText="FIGURE 13: Dactylophysus hirtus sp. nov. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral). Abbreviation: apd, apodeme." figureDoi="http://doi.org/10.5281/zenodo.12641395" httpUri="https://zenodo.org/record/12641395/files/figure.png" pageId="30" pageNumber="247">Figs. 13D</figureCitation>
,
<figureCitation id="6BC0FFE9241CFF9F740863F2F486FD0B" box="[363,410,664,688]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="30" pageNumber="247">15G</figureCitation>
) with apex membranous, suboval, smooth or scarcely punctate, glabrous, apical part membranous, posterior margin with a fringe of minuscule setae.
</paragraph>
<paragraph id="F344E36C241CFF9F75E76252F14AFD94" blockId="30.[132,681,824,944]" lastBlockId="30.[718,1267,184,560]" pageId="30" pageNumber="247">
<emphasis id="C18F3F7E241CFF9F75E76252F5F8FCEB" box="[132,228,824,848]" italics="true" pageId="30" pageNumber="247">Aedeagus:</emphasis>
Phallobase (
<figureCitation id="6BC0FFE9241CFF9F74016252F4DAFCEB" box="[354,454,824,848]" captionStart="FIGURE 13" captionStartId="32.[132,211,761,780]" captionTargetBox="[136,1262,170,734]" captionTargetId="figure-384@32.[132,1266,170,737]" captionTargetPageId="32" captionText="FIGURE 13: Dactylophysus hirtus sp. nov. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (dorsal, ventral). Abbreviation: apd, apodeme." figureDoi="http://doi.org/10.5281/zenodo.12641395" httpUri="https://zenodo.org/record/12641395/files/figure.png" pageId="30" pageNumber="247">Figs. 13E</figureCitation>
,
<figureCitation id="6BC0FFE9241CFF9F74AC6252F4E3FCEB" box="[463,511,824,848]" captionStart="FIGURE 14" captionStartId="33.[95,174,950,969]" captionTargetBox="[96,1225,170,925]" captionTargetId="figure-410@33.[94,1228,170,926]" captionTargetPageId="33" captionText="FIGURE 14: Dactylophysus capixabensis nom. nov. A, antenna; B, pronotum; C, D, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641397" httpUri="https://zenodo.org/record/12641397/files/figure.png" pageId="30" pageNumber="247">14C</figureCitation>
,
<figureCitation id="6BC0FFE9241CFF9F776B6252F731FCEB" box="[520,557,824,848]" captionStart="FIGURE 15" captionStartId="34.[132,211,1826,1845]" captionTargetBox="[134,1265,1352,1799]" captionTargetId="figure-458@34.[132,1266,1349,1803]" captionTargetPageId="34" captionText="FIGURE 15: Dactylophysus tibialis. A, antenna; B, head (dorsal); C, pronotum; D, hypomeron; E, sternite VIII; F, tergite VIII; G, tergites IX and X; H, sternite IX; I, J, aedeagus (dorsal, ventral). Abbreviations: lbs: length of basal strut, lp: length of penis, lph: length of phallobase." figureDoi="http://doi.org/10.5281/zenodo.12641399" httpUri="https://zenodo.org/record/12641399/files/figure.png" pageId="30" pageNumber="247">15I</figureCitation>
) M-shaped, lateral parts divergent posteriorly with a longitudinal apodeme on dorsal surface, 0.4-0.5x the total length of aedeagus, 1.2-1.3x longer than wide, length ratio between lateroposterior and median parts 4.6-22.0. Parameres tapered apicad, 3.1-3.2x wider at base than the narrowest apical part; apex securiform with outer angle acute, membranous posteriorly, with or without short setae; dorsal surface of parameres more sclerotized than the ventral one; ventral surface sclerotized laterally, membranous medially, membrane contiguous to each other and to the phallobase membrane; penis tapering to apex from base of struts to apex, basal struts 0.3-0.4x the total length of penis; dorsal articulation with a narrow and short process fused to the parameres; ventral sclerite present.
</paragraph>
<paragraph id="F344E36C241CFF9F77AD6332F607FC0B" blockId="30.[718,1267,600,944]" pageId="30" pageNumber="247">
<emphasis id="C18F3F7E241CFF9F77AD6332F634FDCB" box="[718,808,600,624]" italics="true" pageId="30" pageNumber="247">Remarks:</emphasis>
All species of
<taxonomicName id="34FB98EF241CFF9F76A96332F14FFDCB" authorityName="Fleutiaux" authorityYear="1892" box="[970,1107,600,624]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E241CFF9F76A96332F14FFDCB" box="[970,1107,600,624]" italics="true" pageId="30" pageNumber="247">Dactylophysus</emphasis>
</taxonomicName>
share the synapomorphies indicated by the present cladistic analysis: labrum with anterior edge curved, prosternal chin piece long and rounded anteriorly, loss of the hind wing sclerotization between radial cell and apical anterior field and posterior part of sternite VIII with three sclerotizations. The combination of those synapomorphies and the diagnostic characters distinguish
<taxonomicName id="34FB98EF241CFF9F76736232F6F5FCCB" box="[784,1001,856,880]" pageId="30" pageNumber="247">
<emphasis id="C18F3F7E241CFF9F76736232F685FCCB" box="[784,921,856,880]" italics="true" pageId="30" pageNumber="247">Dactylophysus</emphasis>
species
</taxonomicName>
from their most similar species belonging to
<taxonomicName id="34FB98EF241CFF9F76C16212F13AFC2B" authorityName="Fischer von Waldheim" authorityYear="1823" box="[930,1062,888,912]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E241CFF9F76C16212F13AFC2B" box="[930,1062,888,912]" italics="true" pageId="30" pageNumber="247">Physodactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="34FB98EF241CFF9F71396212F607FC0B" class="Insecta" family="Elateridae" genus="Heterocrepidius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="247" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C18F3F7E241CFF9F71396212F1EEFC2B" box="[1114,1266,888,912]" italics="true" pageId="30" pageNumber="247">Heterocrepidius</emphasis>
species.
</taxonomicName>
</paragraph>
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<taxonomicName id="34FB98EF2421FFA275E460DDF745FE6B" authority="Fischer von Waldheim, 1823" authorityName="Fischer von Waldheim" authorityYear="1823" box="[135,601,439,464]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="252" phylum="Arthropoda" rank="genus">
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<emphasis id="C18F3F7E2421FFA275E460DDF401FE74" bold="true" box="[135,285,439,463]" italics="true" pageId="35" pageNumber="252">Physodactylus</emphasis>
<bibRefCitation id="976A9E9D2421FFA2744760D2F745FE6B" author="FISCHER VON WALDHEIM, M. G." box="[292,601,440,464]" pageId="35" pageNumber="252" refId="ref49615" refString="FISCHER VON WALDHEIM, M. G. 1823. Entomographia imperii russici; genera insectorum systematica exposita et analysi iconographica instructa. Mosquae, Auctoritate Societatis Caesareae Naturae Scrutatorum Mosquensis. v. 2, 262 p." type="book" year="1823">Fischer von Waldheim, 1823</bibRefCitation>
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<paragraph id="F344E36C2421FFA2753D6092F722FD0B" blockId="35.[94,643,504,752]" pageId="35" pageNumber="252">
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<treatmentCitation id="725AC57D2421FFA2753D6092F762FDAB" author="FISCHER VON WALDHEIM, M. G." box="[94,638,504,528]" page="303" pageId="35" pageNumber="252" year="1823">
<taxonomicName id="34FB98EF2421FFA2753D6092F762FDAB" authority="Fischer von Waldheim, 1823: 303" authorityName="Fischer von Waldheim" authorityPageNumber="303" authorityYear="1823" box="[94,638,504,528]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2421FFA2753D6092F5FEFDAB" box="[94,226,504,528]" italics="true" pageId="35" pageNumber="252">Physodactylus</emphasis>
<bibRefCitation id="976A9E9D2421FFA275976092F762FDAB" author="FISCHER VON WALDHEIM, M. G." box="[244,638,504,528]" pageId="35" pageNumber="252" refId="ref49615" refString="FISCHER VON WALDHEIM, M. G. 1823. Entomographia imperii russici; genera insectorum systematica exposita et analysi iconographica instructa. Mosquae, Auctoritate Societatis Caesareae Naturae Scrutatorum Mosquensis. v. 2, 262 p." type="book" year="1823">Fischer von Waldheim, 1823: 303</bibRefCitation>
</taxonomicName>
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,
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<bibRefCitation id="976A9E9D2421FFA275F46372F419FD8B" author="FISCHER VON WALDHEIM, M. G." box="[151,261,536,560]" pageId="35" pageNumber="252" pagination="448 - 451" refId="ref49654" refString="FISCHER VON WALDHEIM, M. G. 1824. Extrait d'une lettre aadressee a M. Henning sur le Physodactyle, nouveaux genre de Coleoptere voisin des Taupin. Annales des Sciences Naturelles, 3: 448 - 451, plate 27 B." type="journal article" year="1824">1824: 450</bibRefCitation>
</treatmentCitation>
; Guérin-Méneville, 1829: 43;
<treatmentCitation id="725AC57D2421FFA277236372F45CFDEB" author="LACORDAIRE, J. T." page="237" pageId="35" pageNumber="252" year="1857">
<bibRefCitation id="976A9E9D2421FFA277236372F45CFDEB" author="LACORDAIRE, J. T." pageId="35" pageNumber="252" refId="ref50447" refString="LACORDAIRE, J. T. 1857. Histoire Naturelle des Insectes. Genera des Coleopteres. Paris, Librairie Encyclopedique de Roret. v. 4, 579 p." type="book" year="1857">Lacordaire, 1857: 237</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2421FFA2742B6352F703FDEB" author="FLEUTIAUX, E." box="[328,543,568,592]" page="406" pageId="35" pageNumber="252" year="1892">
<bibRefCitation id="976A9E9D2421FFA2742B6352F703FDEB" author="FLEUTIAUX, E." box="[328,543,568,592]" pageId="35" pageNumber="252" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2421FFA277446352F418FDCB" author="SCHWARZ, O." page="311" pageId="35" pageNumber="252" year="1906">
<bibRefCitation id="976A9E9D2421FFA277446352F418FDCB" author="SCHWARZ, O." pageId="35" pageNumber="252" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 311</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2421FFA274736332F4E9FDCB" author="FLEUTIAUX, E." box="[272,501,600,624]" page="165" pageId="35" pageNumber="252" year="1940">
<bibRefCitation id="976A9E9D2421FFA274736332F4E9FDCB" author="FLEUTIAUX, E." box="[272,501,600,624]" pageId="35" pageNumber="252" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 165</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2421FFA277626332F417FD2B" author="BLACKWELDER, R. E." page="292" pageId="35" pageNumber="252" year="1944">
<bibRefCitation id="976A9E9D2421FFA277626332F417FD2B" author="BLACKWELDER, R. E." pageId="35" pageNumber="252" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2421FFA2747F6312F4F1FD2B" author="GOLBACH, R." box="[284,493,632,656]" page="27" pageId="35" pageNumber="252" year="1994">
<bibRefCitation id="976A9E9D2421FFA2747F6312F4F1FD2B" author="GOLBACH, R." box="[284,493,632,656]" pageId="35" pageNumber="252" pagination="1 - 48" refId="ref50116" refString="GOLBACH, R. 1994. Elateridae (Coleoptera) de la Argentina. Historia, catalogo actualizado hasta 1991 inclusive y clave de subfamilias y de generos de Centro y Sudamerica. Opera Lilloana, 41: 1 - 48." type="journal article" year="1994">Golbach, 1994: 27</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2421FFA2749D6312F479FD0B" author="LAWRENCE, J. F. &amp; NEWTON, A. F." page="855" pageId="35" pageNumber="252" year="1995">
<bibRefCitation id="976A9E9D2421FFA2749D6312F479FD0B" author="LAWRENCE, J. F. &amp; NEWTON, A. F." pageId="35" pageNumber="252" pagination="779 - 1006" refId="ref50478" refString="LAWRENCE, J. F. &amp; NEWTON, A. F. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). In: Pakaluk, J. &amp; Slipinski, S. A. (Eds.). Biology, Phylogeny, and Classification of Coleoptera. Warszawa, Muzeum i Instytut Zoologii PAN. v. 2, p. 779 - 1006." type="journal article" year="1995">Lawrence &amp; Newton, 1995: 855</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2421FFA2741363F2F726FD0B" author="CHASSAIN, J." box="[368,570,664,688]" page="65" pageId="35" pageNumber="252" year="2005">
<bibRefCitation id="976A9E9D2421FFA2741363F2F726FD0B" author="CHASSAIN, J." box="[368,570,664,688]" pageId="35" pageNumber="252" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005: 65</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F344E36C2421FFA2753D63D3F5CEFD4B" blockId="35.[94,643,504,752]" pageId="35" pageNumber="252">
<treatmentCitationGroup id="D3EBC4422421FFA2753D63D3F5CEFD4B" pageId="35" pageNumber="252">
<treatmentCitation id="725AC57D2421FFA2753D63D3F463FD6B" author="PERTY, J. A. M." box="[94,383,696,720]" page="24" pageId="35" pageNumber="252" year="1830">
<taxonomicName id="34FB98EF2421FFA2753D63D3F463FD6B" authority="Perty, 1830: 24" authorityName="Perty" authorityPageNumber="24" authorityYear="1830" box="[94,383,696,720]" class="Insecta" family="Elateridae" genus="Drepanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2421FFA2753D63D3F5D8FD6B" box="[94,196,697,720]" italics="true" pageId="35" pageNumber="252">Drepanius</emphasis>
<bibRefCitation id="976A9E9D2421FFA275B163D2F463FD6B" author="PERTY, J. A. M." box="[210,383,696,720]" pageId="35" pageNumber="252" pagination="1 - 12" refId="ref50792" refString="PERTY, J. A. M. 1830. Insecta Brasiliensia. In: Spix, J. &amp; Martius, C. Delectus animalium articulatorum, quae in itinere per Brasiliam annis 1817 - 1820 jussu et auspiciis Maximilliani Josephi I Bavariae regis augustissimi peracto. Munchen, Frid. Fleischer. p. 1 - 60, pls. 1 - 12." type="book chapter" year="1830">Perty, 1830: 24</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; Lacordaire; 1857: 237 (syn).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72421FFA4753D6273F1DEFEEB" lastPageId="37" lastPageNumber="254" pageId="35" pageNumber="252" type="materials_examined">
<paragraph id="F344E36C2421FFA2753D6273F5F3FCEB" blockId="35.[94,643,792,848]" pageId="35" pageNumber="252">
<emphasis id="C18F3F7E2421FFA2753D6273F5CAFC8B" box="[94,214,793,816]" italics="true" pageId="35" pageNumber="252">
<typeStatus id="2C405DCE2421FFA2753D6273F597FC8B" box="[94,139,793,816]" pageId="35" pageNumber="252">Type</typeStatus>
species
</emphasis>
(by monotypy):
<taxonomicName id="34FB98EF2421FFA274FA6272F5F0FCEB" authority="Fischer, 1823" authorityName="Fischer von Waldheim" authorityYear="1823" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="252" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2421FFA274FA6272F79FFC8B" box="[409,643,792,816]" italics="true" pageId="35" pageNumber="252">Physodactylus henningi</emphasis>
<bibRefCitation id="976A9E9D2421FFA2753D6252F5F0FCEB" author="FISCHER VON WALDHEIM, M. G." box="[94,236,824,848]" pageId="35" pageNumber="252" refId="ref49615" refString="FISCHER VON WALDHEIM, M. G. 1823. Entomographia imperii russici; genera insectorum systematica exposita et analysi iconographica instructa. Mosquae, Auctoritate Societatis Caesareae Naturae Scrutatorum Mosquensis. v. 2, 262 p." type="book" year="1823">Fischer, 1823</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="F344E36C2421FFA2753D6213F5A0FAEB" blockId="35.[94,643,888,1360]" pageId="35" pageNumber="252">
<emphasis id="C18F3F7E2421FFA2753D6213F5D8FC2B" box="[94,196,889,912]" italics="true" pageId="35" pageNumber="252">Diagnosis:</emphasis>
Body convex. Frons carinate, antenna submoniliform not surpassing the pronotal posterior angle, labrum subrectangular strongly declivous anteroventrad, mandible falciform unidentate. Prothorax trapezoidal wider than long, lateral pronotal carina directed ventrad anteriorly; anterior angles straight or obtuse in dorsal view, not produced. Metacoxa about 17° inclined to transverse body axis with inner 1/3 about 13 times wider than the outer 2/3; free margin of metacoxal plate very short or absent. Trochanter and femur convex; tibiae widened apicad with external surface covered with spiniform seta and each inner and outer apical margins with a row of 18-27 spiniform setae. Abdominal ventrite 1 nearly divided medially.
</paragraph>
<paragraph id="F344E36C2421FFA2753D6412F455FA2B" blockId="35.[94,329,1400,1424]" box="[94,329,1400,1424]" pageId="35" pageNumber="252">
<emphasis id="C18F3F7E2421FFA2753D6412F5FDFA2B" box="[94,225,1400,1424]" italics="true" pageId="35" pageNumber="252">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2421FFA275846412F45AFA2B" box="[231,326,1400,1424]" name="Brazil" pageId="35" pageNumber="252">BRAZIL</collectingCountry>
.
</paragraph>
<paragraph id="F344E36C2421FFA2753D64D2F72CF954" blockId="35.[94,643,1464,1872]" pageId="35" pageNumber="252">
<emphasis id="C18F3F7E2421FFA2753D64D2F5FCFA6B" box="[94,224,1464,1488]" italics="true" pageId="35" pageNumber="252">Redescription</emphasis>
(male): Integument dull to very shiny with variable color patterns, with punctures umbilicate deeply or weakly impressed, covered with yellow to dark-brown setae; pilosity decumbent to semi-erect on ventral surface; setae usually absent on dorsal surface except for the erect setae on the borders of pronotum and elytra; epipleura densely covered with erect setae visible dorsally around the elytra. Total length: 5.0-
<quantity id="34034E892421FFA275EB67D2F5CAF974" box="[136,214,1720,1744]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="252" unit="mm" value="20.0">20 mm</quantity>
; elytral base 0.96-1.11x as wide as prothorax, elytra 2.70-3.48x longer than pronotum.
</paragraph>
<paragraph id="F344E36C2421FFA275F46792F66EFBCB" blockId="35.[94,643,1464,1872]" lastBlockId="35.[680,1229,184,1872]" pageId="35" pageNumber="252">
Head (
<figureCitation id="6BC0FFE92421FFA275BE6792F45EF8AB" box="[221,322,1784,1808]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="35" pageNumber="252">Figs. 18B</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA2742D6792F462F8AB" box="[334,382,1784,1808]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="35" pageNumber="252">26C</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA274EA6792F4A9F8AB" box="[393,437,1784,1808]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="35" pageNumber="252">28B</figureCitation>
) with anterior margin of frons straight or rounded, dorsal ocular margins curved inwards or subparallel. Frontal carina usually present weakly to strongly impressed, produced or not anteriorly; frontoclypeal region (
<figureCitation id="6BC0FFE92421FFA2769761B2F14EFF4B" box="[1012,1106,216,240]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="35" pageNumber="252">Fig. 26D</figureCitation>
) steeply declivous to base of labrum 5.0-6.0x wider than long; punctures larger than those of pronotum, coalescent to 0.5 diameter apart. Antenna (
<figureCitation id="6BC0FFE92421FFA2716B6052F17BFEEB" box="[1032,1127,312,336]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="35" pageNumber="252">Fig. 16A</figureCitation>
) with 11 antennomeres, antennomeres
<collectionCode id="95EA7BA92421FFA276BD6032F6ECFED4" box="[990,1008,344,367]" pageId="35" pageNumber="252">II</collectionCode>
and
<collectionCode id="95EA7BA92421FFA271456032F15DFED4" box="[1062,1089,344,367]" pageId="35" pageNumber="252">III</collectionCode>
subequal,
<collectionCode id="95EA7BA92421FFA271D26032F1D0FED4" box="[1201,1228,344,367]" pageId="35" pageNumber="252">III</collectionCode>
1.0-1.1x longer than
<collectionCode id="95EA7BA92421FFA276EB6012F680FE34" box="[904,924,376,399]" pageId="35" pageNumber="252">II</collectionCode>
; IV-XI strongly convex (submoniliform); IV-X serrate, XI subcircular to elliptical.
<collectionCode id="95EA7BA92421FFA277CB60D2F7D3FE74" box="[680,719,440,463]" pageId="35" pageNumber="252">IEP</collectionCode>
0.24-0.42. Labrum (
<figureCitation id="6BC0FFE92421FFA276DF60D2F136FE6B" box="[956,1066,440,464]" captionStart="FIGURE 30" captionStartId="61.[95,174,685,704]" captionTargetBox="[96,1225,171,657]" captionTargetId="figure-450@61.[94,1228,170,661]" captionTargetPageId="61" captionText="FIGURE 30: Physodactylus sulcatus. A, antenna; B, maxilla; C, labrum and mandible; D, E, prothorax (dorsal, lateral); F, G, pro and" figureDoi="http://doi.org/10.5281/zenodo.12641441" httpUri="https://zenodo.org/record/12641441/files/figure.png" pageId="35" pageNumber="252">Figs. 30C</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA2715860D2F17AFE6B" box="[1083,1126,440,464]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="35" pageNumber="252">33E</figureCitation>
) subrectangular, anterior margin nearly straight to medially emarginate, concave and strongly declivous ventrad, 2.86-2.92x wider than long, densely and coarsely punctate, with long setae. Mouthparts directed anteroventrally. Mandible (
<figureCitation id="6BC0FFE92421FFA276CE6332F109FDCB" box="[941,1045,600,624]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="35" pageNumber="252">Figs. 18B</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA271476332F148FDCB" box="[1060,1108,600,624]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="35" pageNumber="252">22C</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA271016332F18CFDCB" box="[1122,1168,600,624]" captionStart="FIGURE 30" captionStartId="61.[95,174,685,704]" captionTargetBox="[96,1225,171,657]" captionTargetId="figure-450@61.[94,1228,170,661]" captionTargetPageId="61" captionText="FIGURE 30: Physodactylus sulcatus. A, antenna; B, maxilla; C, labrum and mandible; D, E, prothorax (dorsal, lateral); F, G, pro and" figureDoi="http://doi.org/10.5281/zenodo.12641441" httpUri="https://zenodo.org/record/12641441/files/figure.png" pageId="35" pageNumber="252">30C</figureCitation>
) narrow and long, falciform, unidentate, laterodorsal face densely punctate; lateral edge evenly curved apicad; mesal margin at base with a row of short setae on a prominent rectangular membranous area. Maxilla and labium covered with yellow to dark-brown setae. Maxilla (
<figureCitation id="6BC0FFE92421FFA276676272F675FC8B" box="[772,873,792,816]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="35" pageNumber="252">Figs. 22B</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA276176272F6BCFC8B" box="[884,928,792,816]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="35" pageNumber="252">26B</figureCitation>
) with galea securiform, anterior part usually covered with short fine setae, denser on inner angle (some species with sparse spines and spiniform setae
<figureCitation id="6BC0FFE92421FFA276036212F6A0FC2B" box="[864,956,888,912]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="35" pageNumber="252">Fig. 20B</figureCitation>
); lacinia elongate, tongue-like, densely or sparsely pilose; medistipes trapezoidal longer than wide with several long setae; labium (
<figureCitation id="6BC0FFE92421FFA277D362B2F63EFC4B" box="[688,802,984,1008]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="35" pageNumber="252">Figs. 22D</figureCitation>
) with prementum deeply emarginate on anteromedian margin with anterolateral angles rounded or truncate, not produced; maxillary and labial palpi pilose with fine to stout setae, apical palpomere securiform.
</paragraph>
<paragraph id="F344E36C2421FFA577826512F5A3FED4" blockId="35.[680,1229,184,1872]" lastBlockId="36.[132,681,184,367]" lastPageId="36" lastPageNumber="253" pageId="35" pageNumber="252">
Prothorax subtrapezoidal 1.07-1.56x wider than long, lateral carina directed ventrad anteriorly, nearly entirely visible or with anterior part not visible dorsally, not reaching the anterior margin (incomplete) or reaching the anterior margin (complete); anterior angles straight or obtuse in dorsal view. Pronotum weakly (
<figureCitation id="6BC0FFE92421FFA276606452F67AFAEB" box="[771,870,1336,1360]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="35" pageNumber="252">Fig. 21C</figureCitation>
) to strongly (
<figureCitation id="6BC0FFE92421FFA271676452F176FAEB" box="[1028,1130,1336,1360]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="35" pageNumber="252">Fig. 22H</figureCitation>
) convex, disc, lateral and anterior borders convex with punctures umbilicate deeply impressed (shallowly impressed in some species), posterior margin with very small punctures (
<figureCitation id="6BC0FFE92421FFA2760364D2F6A3FA6B" box="[864,959,1464,1488]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="35" pageNumber="252">Fig. 26F</figureCitation>
); anterior angles right or obtuse, not produced; posterior angles directed ventromedially, usually non-carinate. Hypomeron with punctures usually as large as those of pronotum, denser laterally; posterior 1/5-1/3 glabrous; posterior margin with an inverted U-shaped notch adjacent to the posterior angle (
<figureCitation id="6BC0FFE92421FFA276F96712F6E5F92B" box="[922,1017,1656,1680]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="35" pageNumber="252">Figs. 22I</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA271646712F128F92B" box="[1031,1076,1656,1680]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="35" pageNumber="252">25E</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA271236712F17BF92B" box="[1088,1127,1656,1680]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="35" pageNumber="252">26I</figureCitation>
). Prosternal suture straight (
<figureCitation id="6BC0FFE92421FFA2761B67F2F6CBF90B" box="[888,983,1688,1712]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="35" pageNumber="252">Fig. 25C</figureCitation>
) or curved (
<figureCitation id="6BC0FFE92421FFA2713E67F2F1DCF90B" box="[1117,1216,1688,1712]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="35" pageNumber="252">Fig. 26G</figureCitation>
), margined by a shiny band along hypomeral margin grooved or nor at anterior part. Prosternum (
<figureCitation id="6BC0FFE92421FFA277D36792F606F8AB" box="[688,794,1784,1808]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="35" pageNumber="252">Figs. 18D</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA276446792F648F8AB" box="[807,852,1784,1808]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="35" pageNumber="252">20E</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA276036792F68DF8AB" box="[864,913,1784,1808]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="35" pageNumber="252">22G</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA276FD6792F6D1F8AB" box="[926,973,1784,1808]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="35" pageNumber="252">25C</figureCitation>
,
<figureCitation id="6BC0FFE92421FFA276B96792F115F8AB" box="[986,1033,1784,1808]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="35" pageNumber="252">26G</figureCitation>
) 1.09-1.87x wider than long, with punctures larger than those of pronotum, 0.5-1.5 diameters apart; anterior lobe truncate, covering mouthparts to mentum. Prosternal process (
<figureCitation id="6BC0FFE92426FFA575E861B2F409FF4B" box="[139,277,216,240]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="36" pageNumber="253">Figs. 22G, H</figureCitation>
) 2.0-2.7x longer than diameter of procoxae, ventral surface narrower than the dorsal surface with or without a subapical tooth, curved between procoxae and about 45° declivous to apex. Procoxae open.
</paragraph>
<paragraph id="F344E36C2426FFA575E760F2F491FAEB" blockId="36.[132,681,408,1872]" pageId="36" pageNumber="253">
<emphasis id="C18F3F7E2426FFA575E760F2F5E0FE0B" box="[132,252,408,432]" italics="true" pageId="36" pageNumber="253">Pterothorax:</emphasis>
Mesoventrite with posterior region inclined about 30° above the anterior region (
<figureCitation id="6BC0FFE92426FFA5772760D2F7BDFE6B" box="[580,673,440,464]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="36" pageNumber="253">Fig. 26K</figureCitation>
) with anterior articulating surfaces concave and smooth (
<figureCitation id="6BC0FFE92426FFA575836092F454FDAB" box="[224,328,504,528]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="36" pageNumber="253">Figs. 22K</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA574306092F466FDAB" box="[339,378,504,528]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="36" pageNumber="253">26J</figureCitation>
); borders of mesoventral cavity curved, convergent anteriorly from its half length, floor of cavity with or without shiny median band. Mesocoxal cavity (
<figureCitation id="6BC0FFE92426FFA574326332F4A3FDCB" box="[337,447,600,624]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="36" pageNumber="253">Figs. 22K</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA574B16332F4E0FDCB" box="[466,508,600,624]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="36" pageNumber="253">26L</figureCitation>
) open to both mesepisternum and mesepimeron, trochantin visible; mesepisternum with an oblique carina anteromedially contiguous to the posterior limit of the mesoventral articulating surface. Mesometaventral suture distinct or indistinct. Metaventrite (
<figureCitation id="6BC0FFE92426FFA574C76392F71DFCAB" box="[420,513,760,784]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="36" pageNumber="253">Fig. 22K</figureCitation>
) 1.4-1.6x wider than long, 1.4-1.5x longer than mesoventrite; metepisternum about 7.0 times longer than wide. Scutellum (
<figureCitation id="6BC0FFE92426FFA575E86232F5F3FCCB" box="[139,239,856,880]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="36" pageNumber="253">Figs. 22J</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5759D6232F42FFCCB" box="[254,307,856,880]" captionStart="FIGURE 30" captionStartId="61.[95,174,685,704]" captionTargetBox="[96,1225,171,657]" captionTargetId="figure-450@61.[94,1228,170,661]" captionTargetPageId="61" captionText="FIGURE 30: Physodactylus sulcatus. A, antenna; B, maxilla; C, labrum and mandible; D, E, prothorax (dorsal, lateral); F, G, pro and" figureDoi="http://doi.org/10.5281/zenodo.12641441" httpUri="https://zenodo.org/record/12641441/files/figure.png" pageId="36" pageNumber="253">30M</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA574206232F472FCCB" box="[323,366,856,880]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">33E</figureCitation>
) abruptly elevated above the level of mesoscutum, pentagonal, 1.1-1.5 longer than wide, anterior edge rounded. Metanotum (
<figureCitation id="6BC0FFE92426FFA5775D62F2F781FC0B" box="[574,669,920,944]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="36" pageNumber="253">Fig. 26R</figureCitation>
): prescutum contiguous medially to the scutum by an acute process; posterior part of the scutellum with a longitudinal apodeme. Elytra with apices conjointly rounded, striae with a single (
<figureCitation id="6BC0FFE92426FFA574AF6572F72CFB8B" box="[460,560,1048,1072]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">Fig. 33N</figureCitation>
) or several (
<figureCitation id="6BC0FFE92426FFA575EF6552F5F6FBEB" box="[140,234,1080,1104]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">Fig. 33O</figureCitation>
) rows of punctures usually increasing in size posteriorly; interstices flat to convex, impunctate or sparsely punctate; epipleura abruptly narrowed near metacoxa, slightly widened apicad from ventrite 2. Hind wings (
<figureCitation id="6BC0FFE92426FFA5747065D2F469FB6B" box="[275,373,1208,1232]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="36" pageNumber="253">Fig. 26Q</figureCitation>
) with radial cell 3.5x longer than wide, CuA
<subScript id="6F7FE1292426FFA57451658DF426FB4E" attach="left" box="[306,314,1255,1269]" fontSize="6" pageId="36" pageNumber="253">1</subScript>
present; wedge cell present; apex with anterior, median and posterior field sclerotizations convergent basad and an oblique sclerotization adjacent to the radial cell.
</paragraph>
<paragraph id="F344E36C2426FFA575DE6432F64AFDAB" blockId="36.[132,681,408,1872]" lastBlockId="36.[718,1267,184,1392]" pageId="36" pageNumber="253">
Metacoxa (
<figureCitation id="6BC0FFE92426FFA574546432F480FACB" box="[311,412,1368,1392]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="36" pageNumber="253">Fig. 22K</figureCitation>
) inclined about 17° in relation to transverse axis of body, ventral part as long as the dorsal one, abruptly shortened on out- er 2/3, inner 1/3 about 13 times longer than outer 2/3; free margin of metacoxal plate absent or very short. Tibiae with 18-27 spiniform setae along each outer and inner apical border (
<figureCitation id="6BC0FFE92426FFA574BE6772F723F98B" box="[477,575,1560,1584]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">Fig. 33L</figureCitation>
). Protibia (
<figureCitation id="6BC0FFE92426FFA575E86752F5F5F9EB" box="[139,233,1592,1616]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="36" pageNumber="253">Figs. 25F</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA575926752F43FF9EB" box="[241,291,1592,1616]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="36" pageNumber="253">26N</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA574486752F448F9EB" box="[299,340,1592,1616]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="36" pageNumber="253">28F</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5743F6752F497F9EB" box="[348,395,1592,1616]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">33G</figureCitation>
) 2.2-3.5x wider at apex than at base, with dorsal margin rugose, slightly to strongly curved with apical angle usually acute and produced; dorsal margin and outer half covered with spiniform setae. Mesotibia (
<figureCitation id="6BC0FFE92426FFA5742167D2F483F96B" box="[322,415,1720,1744]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="36" pageNumber="253">Fig. 18F</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA574CC67D2F4C2F96B" box="[431,478,1720,1744]" captionStart="FIGURE 19" captionStartId="43.[95,174,1826,1845]" captionTargetBox="[100,1222,389,1798]" captionTargetId="figure-93@43.[94,1228,385,1802]" captionTargetPageId="43" captionText="FIGURE 19: Physodactylus chassaini sp. nov. A, antenna; B, pronotum; C, mesotibia; D, E, aedeagus (dorsal, ventral). Physodactylus fischeri. F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641409" httpUri="https://zenodo.org/record/12641409/files/figure.png" pageId="36" pageNumber="253">19C</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5748C67D2F700F96B" box="[495,540,1720,1744]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="36" pageNumber="253">21E</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5774E67D2F743F96B" box="[557,607,1720,1744]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="36" pageNumber="253">26O</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5771367D2F7BCF96B" box="[624,672,1720,1744]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">33H</figureCitation>
) 2.0-3.1 wider at apex than at base, with dorsal margin curved, dorsal apical angle obtuse; covered with spiniform setae, dense on dorsal margin and outer surface, sparse on inner surface. Metatibia (
<figureCitation id="6BC0FFE92426FFA574926652F747F8EB" box="[497,603,1848,1872]" captionStart="FIGURE 17" captionStartId="40.[132,211,1852,1871]" captionTargetBox="[134,1263,1263,1821]" captionTargetId="figure-462@40.[132,1266,1261,1828]" captionTargetPageId="40" captionText="FIGURE 17: Physodactylus brasiliensis. A, antenna; B, C, prothorax (dorsal, lateral); D, metatibia; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641405" httpUri="https://zenodo.org/record/12641405/files/figure.png" pageId="36" pageNumber="253">Figs. 17D</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA577046652F793F8EB" box="[615,655,1848,1872]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="36" pageNumber="253">21F</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA577F86652F7EDFF6B" box="[667,753,184,1872]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="13.[95,174,1724,1743]" captionStartId-1="16.[132,211,1800,1819]" captionStartId-2="17.[95,174,723,742]" captionStartId-3="18.[132,211,1800,1819]" captionTargetBox-0="[100,1222,299,1700]" captionTargetBox-1="[136,1263,685,1777]" captionTargetBox-2="[97,1225,170,699]" captionTargetBox-3="[133,1265,1196,1773]" captionTargetId-0="figure-57@13.[94,1228,298,1700]" captionTargetId-1="figure-249@16.[132,1266,681,1777]" captionTargetId-2="figure-467@17.[94,1228,170,699]" captionTargetId-3="figure-408@18.[132,1266,1191,1777]" captionTargetPageId-0="13" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIGURE 2: Teslasena femoralis (male): A, antenna; B, C, head (dorsal, anterior); D, labrum, E, mandible (dorsal); F, G, H, prothorax (ventral, dorsal, lateral); I, J, K, pterothorax (ventral, lateroventral, lateral); L, pterothorax (dorsal) and abdominal tergite I; M, hind wing; N, O, abdomen (ventral, lateral); P,tergite VIII; Q, sternite VIII; R, sternite IX; S, tergites IX and X; T, U, V, aedeagus (ventral, dorsal, lateral). Abbreviations: af, mf and pf: anterior, median and posterior field sclerotizations; llpp: length of lateroposterior part of phallobase, lp: length of prosternum, lpp: lateroposterior part of phallobase, mp: median part of phallobase, ppp: posterior part of penis; tbpp: triangular basal part of penis, wp: width of prosternum." captionText-1="FIGURE 3: Idiotropia henoni (male): A, antenna; B, C, head (dorsal, anterodorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, mesocoxal cavity (lateroventral); I, mesoventrite (lateral); J, pterothorax (dorsal) and abdominal tergites I-II; K, mesoscutum and scutellar shield (lateral); L, pterothorax (ventral), M, N, O, pro-, meso- and metathoracic legs (outer surface). Scale = 1.0 mm." captionText-2="FIGURE 4: Idiotropia henoni. A, sternite IX; B, tergites IX and X; C, sternite VIII; D, tergite VIII. E, F, aedeagus (dorsal, ventral). Scale = 0.1 mm. Abbreviations: ar, dorsal articulation; vs, ventral sclerite." captionText-3="FIGURE 5: Oligostethius capensis. A, B, C, head (anterodorsal, lateral, dorsal); D, E, F, prothorax (dorsal, ventral, lateral); G, hypomeron; H, I, pterothorax (ventral, lateroventral); J, mesoscutum and scutellar shield; K, L, abdomen (ventral, lateral). Scale = 1.0 mm. Abbreviations: llp: length of lateral part, lmp length of median part." figureDoi-0="http://doi.org/10.5281/zenodo.12641371" figureDoi-1="http://doi.org/10.5281/zenodo.12641373" figureDoi-2="http://doi.org/10.5281/zenodo.12641375" figureDoi-3="http://doi.org/10.5281/zenodo.12641377" httpUri-0="https://zenodo.org/record/12641371/files/figure.png" httpUri-1="https://zenodo.org/record/12641373/files/figure.png" httpUri-2="https://zenodo.org/record/12641375/files/figure.png" httpUri-3="https://zenodo.org/record/12641377/files/figure.png" pageId="36" pageNumber="253">2 5G</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5766361D2F634FF6B" box="[768,808,184,208]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="36" pageNumber="253">26P</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5765561D2F675FF6B" box="[822,873,184,208]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="36" pageNumber="253">28H</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5761461D2F6B9FF6B" box="[887,933,184,208]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">33K</figureCitation>
) 2.4-3.5 wider at apex than at base with apical dorsal angle produced (not produced in some
<taxonomicName id="34FB98EF2426FFA5764E6193F695FEAB" authorityName="Mannerheim" authorityYear="1842" box="[813,905,248,272]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="253" phylum="Arthropoda" rank="species" species="besckei">
<emphasis id="C18F3F7E2426FFA5764E6193F695FEAB" box="[813,905,248,272]" italics="true" pageId="36" pageNumber="253">P. besckei</emphasis>
</taxonomicName>
specimens); densely covered with spiniform setae. Femur (
<figureCitation id="6BC0FFE92426FFA576B36072F173FE8B" box="[976,1135,280,304]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">Figs. 33F, H, J</figureCitation>
) rectangular with outer and inner surfaces convex; metatrochanter more convex than metafemur. Tarsomeres decreasing in length from I-IV,
<collectionCode id="95EA7BA92426FFA576A86012F6C1FE34" box="[971,989,376,399]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="36" pageNumber="253" type="Museum">V</collectionCode>
longer than
<collectionCode id="95EA7BA92426FFA571016012F161FE34" box="[1122,1149,376,399]" pageId="36" pageNumber="253">III</collectionCode>
and
<collectionCode id="95EA7BA92426FFA571D16012F1D1FE34" box="[1202,1229,376,399]" pageId="36" pageNumber="253">IV</collectionCode>
together; ventral surface densely pilose, tarsomeres I-III or II-III lamellate (
<figureCitation id="6BC0FFE92426FFA576F760D2F6E5FE6B" box="[916,1017,440,464]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="36" pageNumber="253">Fig. 33M</figureCitation>
), lamellae of tarsomeres II-III subequal, I a little smaller, minuscule or absent; claws simple.
</paragraph>
<paragraph id="F344E36C2426FFA576646372F1EFFAD4" blockId="36.[718,1267,184,1392]" pageId="36" pageNumber="253">
Abdomen (
<figureCitation id="6BC0FFE92426FFA5761F6372F114FD8B" box="[892,1032,536,560]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="36" pageNumber="253">Figs. 22L, M</figureCitation>
) with variable pilosity patterns on ventrites 1-4, ventrite 5 with apex coarsely punctate with pilosity dense and stout; posterior angle of ventrite 1 and lateral margins of 2-4 with prominent marginal plates (
<figureCitation id="6BC0FFE92426FFA5768963F2F157FD0B" box="[1002,1099,664,688]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="36" pageNumber="253">Fig. 25H</figureCitation>
); ventrite 5 subtriangular 1.1-1.5x wider than long, evenly convex. Ventrite 1 nearly divided medially, 2-4 subequal in length. Sternite VIII (
<figureCitation id="6BC0FFE92426FFA576D26392F113FCAB" box="[945,1039,760,784]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="36" pageNumber="253">Figs. 18I</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA571786392F154FCAB" box="[1051,1096,760,784]" captionStart="FIGURE 23" captionStartId="52.[132,211,1852,1871]" captionTargetBox="[133,1263,1197,1817]" captionTargetId="figure-469@52.[132,1266,1185,1828]" captionTargetPageId="52" captionText="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641421" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="36" pageNumber="253">23A</figureCitation>
) subrectangular, with posterior margin straight to emarginate medially, partly covered with short setae, lateroposterior angles usually with long stout setae; with a pair of light sclerotizations on lateroposterior angles and a transverse band-like dark sclerotization on anteromedian border. Tergite VIII (
<figureCitation id="6BC0FFE92426FFA576B462D2F128FC6B" box="[983,1076,952,976]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="36" pageNumber="253">Figs. 18J</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5715D62D2F170FC6B" box="[1086,1132,952,976]" captionStart="FIGURE 23" captionStartId="52.[132,211,1852,1871]" captionTargetBox="[133,1263,1197,1817]" captionTargetId="figure-469@52.[132,1266,1185,1828]" captionTargetPageId="52" captionText="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641421" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="36" pageNumber="253">23B</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5711562D2F1B8FC6B" box="[1142,1188,952,976]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="36" pageNumber="253">28K</figureCitation>
) evenly sclerotized U-shaped, covered with short setae, setae longer on lateroposterior and apical margins. Sternite IX (
<figureCitation id="6BC0FFE92426FFA577946572F67DFB8B" box="[759,865,1048,1072]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="36" pageNumber="253">Figs. 18G</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA5760E6572F680FB8B" box="[877,924,1048,1072]" captionStart="FIGURE 23" captionStartId="52.[132,211,1852,1871]" captionTargetBox="[133,1263,1197,1817]" captionTargetId="figure-469@52.[132,1266,1185,1828]" captionTargetPageId="52" captionText="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641421" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="36" pageNumber="253">23C</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA576CA6572F6D1FB8B" box="[937,973,1048,1072]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="36" pageNumber="253">28I</figureCitation>
) with anteromedian margin rounded, produced and curved dorsad; latero-anterior margins strongly sclerotized, anteromedian surface translucent, posterior surface sclerotized and covered with moderately long setae, longer on lateroposterior and apical margins; sternite and tergite IX fused near the anterior margin. Tergite IX (
<figureCitation id="6BC0FFE92426FFA5717565B2F163FB4B" box="[1046,1151,1240,1264]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="36" pageNumber="253">Figs. 18H</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA571EA65B2F1A7FB4B" box="[1161,1211,1240,1264]" captionStart="FIGURE 23" captionStartId="52.[132,211,1852,1871]" captionTargetBox="[133,1263,1197,1817]" captionTargetId="figure-469@52.[132,1266,1185,1828]" captionTargetPageId="52" captionText="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641421" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="36" pageNumber="253">23D</figureCitation>
,
<figureCitation id="6BC0FFE92426FFA571A565B2F1F6FB4B" box="[1222,1258,1240,1264]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="36" pageNumber="253">28J</figureCitation>
) with anterior margin straight or sinuous, apical lobes acute or rounded with long setae. Tergite X with apex membranous, suboval, scarcely punctate, usually glabrous, apical margin with a fringe of minuscule setae.
</paragraph>
<paragraph id="F344E36C2426FFA477AD64F2F4EFFF54" blockId="36.[718,1266,1432,1872]" lastBlockId="37.[94,642,184,240]" lastPageId="37" lastPageNumber="254" pageId="36" pageNumber="253">
<emphasis id="C18F3F7E2426FFA577AD64F2F632FA0B" box="[718,814,1432,1456]" italics="true" pageId="36" pageNumber="253">Aedeagus:</emphasis>
Phallobase M-shaped, lateral parts divergent posteriorly with a longitudinal apodeme on dorsal surface, 0.4-0.5x the total length of aedeagus, 0.9-1.6x longer than wide, length ratio between lateroposterior and median parts 3.3-3.4. Parameres tapered apicad, 2.1-4.8x wider at base than the narrowest apical part; with dorsal surface more sclerotized than the ventral one; ventral surface sclerotized laterally, membranous medially, membrane contiguous to each other and to the phallobase membrane; apex securiform with outer angle acute, evenly sclerotized or membranous posteriorly, with or without short setae. Penis with basal struts 0.2-0.3x the total length of penis; dorsal articulation fused to the parameres by a median sclerotized process (
<figureCitation id="6BC0FFE92427FFA475DA61D2F40BFF6B" box="[185,279,184,208]" captionStart="FIGURE 17" captionStartId="40.[132,211,1852,1871]" captionTargetBox="[134,1263,1263,1821]" captionTargetId="figure-462@40.[132,1266,1261,1828]" captionTargetPageId="40" captionText="FIGURE 17: Physodactylus brasiliensis. A, antenna; B, C, prothorax (dorsal, lateral); D, metatibia; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641405" httpUri="https://zenodo.org/record/12641405/files/figure.png" pageId="37" pageNumber="254">Fig. 17E</figureCitation>
) or by a membranous translucent area (
<figureCitation id="6BC0FFE92427FFA475F661B2F5EFFF4B" box="[149,243,216,240]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="37" pageNumber="254">Fig. 18K</figureCitation>
); ventral sclerite present.
</paragraph>
<paragraph id="F344E36C2427FFA4753D6072F1DEFEEB" blockId="37.[94,643,280,336]" lastBlockId="37.[680,1229,184,336]" pageId="37" pageNumber="254">
<emphasis id="C18F3F7E2427FFA4753D6072F5ABFE8B" box="[94,183,280,304]" italics="true" pageId="37" pageNumber="254">Remarks:</emphasis>
The combination of the most conspicuous characters of
<taxonomicName id="34FB98EF2427FFA475836052F4B6FEEB" box="[224,426,312,336]" pageId="37" pageNumber="254">
<emphasis id="C18F3F7E2427FFA475836052F47DFEEB" box="[224,353,312,336]" italics="true" pageId="37" pageNumber="254">Physodactylus</emphasis>
species
</taxonomicName>
(shape of labrum, the short and laterally convex antennae almost moniliform in some species, fossorial legs, body convex, metacoxae almost horizontal and reduced laterally and tarsomeres lamellate) easily differentiate them from other similar species as those of
<taxonomicName id="34FB98EF2427FFA476C86052F11EFEEB" authorityName="Fleutiaux" authorityYear="1892" box="[939,1026,312,336]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="C18F3F7E2427FFA476C86052F11EFEEB" box="[939,1026,312,336]" italics="true" pageId="37" pageNumber="254">Teslasena</emphasis>
</taxonomicName>
and
<emphasis id="C18F3F7E2427FFA471566052F1DEFEEB" box="[1077,1218,312,336]" italics="true" pageId="37" pageNumber="254">
<taxonomicName id="34FB98EF2427FFA471566052F1A2FEEB" authorityName="Fleutiaux" authorityYear="1892" box="[1077,1214,312,336]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="254" phylum="Arthropoda" rank="genus">Dactylophysus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,179 @@
<document id="2088A68FA64F65E1332671E19519B030" ID-DOI="10.1590/0031-1049.2014.54.18" ID-ISSN="1807-0205" ID-Zenodo-Dep="12641367" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720058375023" checkinUser="felipe" docAuthor="Rosa, Simone Policena" docDate="2014" docId="7B52527A2424FFA6759C659DF128FA8B" docLanguage="en" docName="PapAvZool.54.18.217-292.pdf" docOrigin="Papéis Avulsos de Zoologia (São Paulo) 54 (18)" docSource="http://dx.doi.org/10.1590/0031-1049.2014.54.18" docStyle="DocumentStyle:8E93E160036069519CAC16E940FF2271.1:PapAvZool.2014-2017.journal_article" docStyleId="8E93E160036069519CAC16E940FF2271" docStyleName="PapAvZool.2014-2017.journal_article" docStyleVersion="1" docTitle="Physodactylus asper Rosa, 2014, sp. nov." docType="treatment" docVersion="2" lastPageNumber="256" masterDocId="876B2A022402FF817563616AF51CFFBB" masterDocTitle="Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)" masterLastPageNumber="292" masterPageNumber="217" pageNumber="255" updateTime="1720066190901" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D16E98B42ED5705617A1A27A1171810A">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<mods:date id="23265F86F3DA4EED91ABAC8E52A47304">2014</mods:date>
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<subSubSection id="BBE1B0E72424FFA7759C659DF732FAAB" box="[255,558,1271,1296]" pageId="38" pageNumber="255" type="nomenclature">
<paragraph id="F344E36C2424FFA7759C659DF732FAAB" blockId="38.[255,558,1271,1328]" box="[255,558,1271,1296]" pageId="38" pageNumber="255">
<heading id="A80C54002424FFA7759C659DF732FAAB" bold="true" box="[255,558,1271,1296]" centered="true" fontSize="10" level="2" pageId="38" pageNumber="255" reason="2">
<emphasis id="C18F3F7E2424FFA7759C659DF732FAAB" bold="true" box="[255,558,1271,1296]" pageId="38" pageNumber="255">
<taxonomicName id="34FB98EF2424FFA7759C659DF4C8FAB4" authorityName="Rosa" authorityYear="2014" box="[255,468,1271,1295]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="255" phylum="Arthropoda" rank="species" species="asper" status="sp. nov.">
<emphasis id="C18F3F7E2424FFA7759C659DF4C8FAB4" bold="true" box="[255,468,1271,1295]" italics="true" pageId="38" pageNumber="255">Physodactylus asper</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052424FFA774B86592F732FAAB" box="[475,558,1272,1296]" pageId="38" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72424FFA6744F6472F128FA8B" lastPageId="39" lastPageNumber="256" pageId="38" pageNumber="255" type="description">
<paragraph id="F344E36C2424FFA7744F6472F71CFA8B" blockId="38.[255,558,1271,1328]" box="[300,512,1304,1328]" pageId="38" pageNumber="255">
<emphasis id="C18F3F7E2424FFA7744F6472F71CFA8B" bold="true" box="[300,512,1304,1328]" pageId="38" pageNumber="255">
(
<figureCitation id="6BC0FFE92424FFA774506472F494FA8B" box="[307,392,1304,1328]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="38" pageNumber="255">Figs. 16</figureCitation>
A-D, 34Q)
</emphasis>
</paragraph>
<paragraph id="F344E36C2424FFA775E76432F4EEFA2B" blockId="38.[132,679,1368,1424]" pageId="38" pageNumber="255">
<emphasis id="C18F3F7E2424FFA775E76432F5F3FACB" box="[132,239,1368,1392]" italics="true" pageId="38" pageNumber="255">Etymology:</emphasis>
From Latin, asper = rough; alluding to the rugose and coarsely punctate elytra.
</paragraph>
<paragraph id="F344E36C2424FFA675E764D3F4F1FB6B" blockId="38.[132,681,1464,1872]" lastBlockId="39.[94,643,1080,1232]" lastPageId="39" lastPageNumber="256" pageId="38" pageNumber="255">
<emphasis id="C18F3F7E2424FFA775E764D3F5EFFA6B" box="[132,243,1465,1488]" italics="true" pageId="38" pageNumber="255">Description</emphasis>
(male,
<figureCitation id="6BC0FFE92424FFA7743764D2F4A2FA6B" box="[340,446,1464,1488]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="38" pageNumber="255">Fig. 34Q</figureCitation>
): Integument bright; head, antennae, legs and ventral surface black, pronotum reddish-brown, elytra dark brown with epipleura lighter;
<specimenCount id="E5FD28E52424FFA775BB6772F474F98B" box="[216,360,1560,1584]" count="1" pageId="38" pageNumber="255" type="generic">one specimen</specimenCount>
with pterothorax and ventral abdomen reddish-brown on the median longitudinal line; pilosity brown. Total length
<quantity id="34034E892424FFA774926732F745F9D4" box="[497,601,1624,1648]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="38" pageNumber="255" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8-10 mm</quantity>
; elytral base 1.0x as wide as prothorax, elytra 3.0-3.2x times longer than pronotum. Frons slightly concave, frontal carina weakly impressed; antenna (
<figureCitation id="6BC0FFE92424FFA7748B67D2F758F96B" box="[488,580,1720,1744]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="38" pageNumber="255">Fig. 16A</figureCitation>
) reaching the posterior third of pronotum, antennomere IV as wide as long, V-X 1.17-1.20x wider than long; XI oval 1.5x longer than wide. Pronotum (
<figureCitation id="6BC0FFE92424FFA774BC6672F72AF88B" box="[479,566,1816,1840]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="38" pageNumber="255">Fig. 16B</figureCitation>
) 1.20-1.26 wider than long, with lateral sides nearly straight to rounded convergent from posterior angles to anterior margin; anteromedian margin weakly produced; posterior angles short, convex, parallel; lateral carina prominent nearly entirely visible dorsally, complete; disc with punctures 2-3 diameters apart, larger and denser on lateral and anterior borders; prosternal process without subapical tooth. Lamellae of pro- and mesotarsomeres I smaller than the others, absent on metatarsomere I. Elytra tapering to apex from anterior third or subparallel on anterior 2/3 then tapering to apex; striae with large and small dense punctation, striae indistinct on apical region where the punctation is mixed and coalescent; interstices convex and rugose, scarcely punctate; apical striae with punctures larger than those of pronotal lateral border. Abdomen with ventrites 2-4 strongly concave laterally; ventrite 1 densely punctate with minuscule fine setae, lateral border smooth with a few long, very fine setae; ventrites 2-4 punctate as ventrite 1 with long, decumbent and semi-erect fine setae, lateral border smooth with a few finer setae. Pregenitalic segments and aedeagus covered with stout dark-brown setae. Sternite VIII emarginate posteromedially, anterior sclerotization 0.62x times the total width of sternite; sternite IX with apex rounded.
</paragraph>
<caption id="A784B3E42425FFA6753D62B1F771FBB3" ID-DOI="http://doi.org/10.5281/zenodo.12641403" ID-Zenodo-Dep="12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="39" pageNumber="256" startId="39.[94,173,987,1006]" targetBox="[95,1224,173,961]" targetPageId="39" targetType="figure">
<paragraph id="F344E36C2425FFA6753D62B1F771FBB3" blockId="39.[94,1229,987,1032]" pageId="39" pageNumber="256">
<emphasis id="C18F3F7E2425FFA6753D62B1F5CCFC54" bold="true" box="[94,208,987,1007]" pageId="39" pageNumber="256">FIGURE 16:</emphasis>
<taxonomicName id="34FB98EF2425FFA675BB62B6F46CFC55" authorityName="Rosa" authorityYear="2014" box="[216,368,988,1007]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="256" phylum="Arthropoda" rank="species" species="asper" status="sp. nov.">
<emphasis id="C18F3F7E2425FFA675BB62B6F46CFC55" box="[216,368,988,1007]" italics="true" pageId="39" pageNumber="256">Physodactylus asper</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052425FFA6741B62B6F4A5FC54" box="[376,441,988,1007]" pageId="39" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E2425FFA674DC62B1F4CEFC55" bold="true" box="[447,466,987,1006]" pageId="39" pageNumber="256">A,</emphasis>
antenna;
<emphasis id="C18F3F7E2425FFA6774462B6F726FC54" bold="true" box="[551,570,988,1007]" pageId="39" pageNumber="256">B,</emphasis>
pronotum;
<emphasis id="C18F3F7E2425FFA677C262B1F7CFFC54" bold="true" box="[673,723,987,1007]" pageId="39" pageNumber="256">C, D,</emphasis>
aedeagus (dorsal, ventral).
<emphasis id="C18F3F7E2425FFA676DA62B6F17BFC54" box="[953,1127,988,1007]" italics="true" pageId="39" pageNumber="256">
<taxonomicName id="34FB98EF2425FFA676DA62B6F17FFC54" authorityName="Mannerheim" authorityYear="1842" box="[953,1123,988,1007]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="256" phylum="Arthropoda" rank="species" species="besckei">Physodactylus besckei</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2425FFA6710F62B6F162FC54" bold="true" box="[1132,1150,988,1007]" pageId="39" pageNumber="256">E,</emphasis>
antenna;
<emphasis id="C18F3F7E2425FFA6753D629FF594FBB3" bold="true" box="[94,136,1013,1032]" pageId="39" pageNumber="256">F, G,</emphasis>
prothorax (dorsal, lateral);
<emphasis id="C18F3F7E2425FFA6740A629FF48FFBB3" bold="true" box="[361,403,1013,1032]" pageId="39" pageNumber="256">H, I,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C2425FFA6753D6592F798FA6B" blockId="39.[94,644,1272,1488]" pageId="39" pageNumber="256">
<emphasis id="C18F3F7E2425FFA6753D6592F5ABFAAB" box="[94,183,1272,1296]" italics="true" pageId="39" pageNumber="256">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92425FFA675AB6592F444FAAB" box="[200,344,1272,1296]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="39" pageNumber="256">Figs. 16C, D</figureCitation>
): Phallobase 0.49x the total length of aedeagus, 1.26x longer than wide, length ratio between lateroposterior and median parts 34; paramere with ventral surface nearly entirely sclerotized with anteromedian margin distinct; penis with basal strut 0.19x its total length, articulated to the parameres by a sclerotized short process; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C2425FFA6753D6492F496F9EB" blockId="39.[94,643,1528,1616]" pageId="39" pageNumber="256">
<materialsCitation id="4393E9312425FFA6753D6492F496F9EB" collectingDate="1965-11-19" collectionCode="UF-PARANA, DZUP" collectorName="F. M. Oliveira" country="Brazil" location="BAHIA" pageId="39" pageNumber="256" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="C18F3F7E2425FFA6753D6492F5A7F9AB" box="[94,187,1528,1552]" italics="true" pageId="39" pageNumber="256">
<typeStatus id="2C405DCE2425FFA6753D6492F5AAF9AB" box="[94,182,1528,1552]" pageId="39" pageNumber="256" type="holotype">Holotype</typeStatus>
:
</emphasis>
[DPTº ZOOL,
<collectionCode id="95EA7BA92425FFA674F76492F736F9A8" box="[404,554,1528,1555]" pageId="39" pageNumber="256">UF-PARANÁ</collectionCode>
], [MARACÁS,
<collectingCountry id="8BECA3FC2425FFA675AC6772F431F98B" box="[207,301,1560,1584]" name="Brazil" pageId="39" pageNumber="256">BRAZIL</collectingCountry>
,
<collectingRegion id="313F2D8E2425FFA6742A6772F480F98B" box="[329,412,1560,1584]" country="Brazil" name="Bahia" pageId="39" pageNumber="256">BAHIA</collectingRegion>
,
<date id="8745C5AC2425FFA674D46772F72AF98B" box="[439,566,1560,1584]" pageId="39" pageNumber="256" value="1965-11-19">
<collectingDate id="97013C442425FFA674D46772F72AF98B" box="[439,566,1560,1584]" pageId="39" pageNumber="256" value="1965-11-19">19/11/1965</collectingDate>
</date>
,
<collectorName id="5E0E86BA2425FFA677316772F5ADF9EB" pageId="39" pageNumber="256">F.M. Oliveira</collectorName>
leg.], male (
<collectionCode id="95EA7BA92425FFA674526752F462F9EB" box="[305,382,1592,1616]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="39" pageNumber="256" type="University or college">DZUP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C2425FFA6753D6713F455F90B" blockId="39.[94,642,1656,1712]" pageId="39" pageNumber="256">
<materialsCitation id="4393E9312425FFA6753D6713F458F90B" collectionCode="DZUP, MZUSP" pageId="39" pageNumber="256" specimenCount="3" typeStatus="paratype">
<emphasis id="C18F3F7E2425FFA6753D6713F5DCF92B" box="[94,192,1657,1680]" italics="true" pageId="39" pageNumber="256">
<typeStatus id="2C405DCE2425FFA6753D6713F5A6F92B" box="[94,186,1657,1680]" pageId="39" pageNumber="256" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same labels as
<typeStatus id="2C405DCE2425FFA6740D6712F4D7F92B" box="[366,459,1656,1680]" pageId="39" pageNumber="256" type="holotype">holotype</typeStatus>
,
<specimenCount id="E5FD28E52425FFA674B96712F704F934" box="[474,536,1656,1680]" count="2" pageId="39" pageNumber="256" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA92425FFA677496712F76BF92B" box="[554,631,1656,1680]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="39" pageNumber="256" type="University or college">DZUP</collectionCode>
); idem,
<specimenCount id="E5FD28E52425FFA675C367F2F5CDF914" box="[160,209,1688,1712]" count="1" pageId="39" pageNumber="256" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA92425FFA675BC67F2F422F90B" box="[223,318,1688,1712]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="39" pageNumber="256">MZUSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C2425FFA6753D67B2F4E8F94B" blockId="39.[94,500,1752,1776]" box="[94,500,1752,1776]" pageId="39" pageNumber="256">
<emphasis id="C18F3F7E2425FFA6753D67B2F5FDF94B" box="[94,225,1752,1776]" italics="true" pageId="39" pageNumber="256">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2425FFA6758467B2F45AF94B" box="[231,326,1752,1776]" name="Brazil" pageId="39" pageNumber="256">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2425FFA6742C67B2F48CF94B" box="[335,400,1752,1776]" italics="true" pageId="39" pageNumber="256">
<collectingRegion id="313F2D8E2425FFA6742C67B2F497F94B" box="[335,395,1752,1776]" country="Brazil" name="Bahia" pageId="39" pageNumber="256">Bahia</collectingRegion>
:
</emphasis>
Maracás.
</paragraph>
<paragraph id="F344E36C2425FFA6753D6672F128FA8B" blockId="39.[94,642,1816,1872]" lastBlockId="39.[680,1229,1080,1328]" pageId="39" pageNumber="256">
<emphasis id="C18F3F7E2425FFA6753D6672F496F88B" box="[94,394,1816,1840]" italics="true" pageId="39" pageNumber="256">
Remarks:
<taxonomicName id="34FB98EF2425FFA675A46672F496F88B" authorityName="Rosa" authorityYear="2014" box="[199,394,1816,1840]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="256" phylum="Arthropoda" rank="species" species="asper" status="sp. nov.">Physodactylus asper</taxonomicName>
</emphasis>
<taxonomicNameLabel id="DABC82052425FFA674FA6672F4F3F894" box="[409,495,1816,1839]" pageId="39" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="34FB98EF2425FFA6753D6653F5A2F8EB" authorityName="Fleutiaux" authorityYear="1940" box="[94,190,1848,1872]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="256" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="C18F3F7E2425FFA6753D6653F5A2F8EB" box="[94,190,1848,1872]" italics="true" pageId="39" pageNumber="256">P. fischeri</emphasis>
</taxonomicName>
in the shape of prothorax, brown pilosity and the punctation and roughness of elytra, differing from this species in its smaller size, longer antenna, weaker impressed frontal carina, lateral pronotal carina complete, pilosity pattern of the abdominal ventrites, aedeagus with parameres more extensively sclerotized ventrally and articulated to penis through a sclerotized process, basal struts shorter and wider and median part of phallobase shorter.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,177 @@
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<taxonomicName id="34FB98EF2425FFA677B9641DF186FA2B" authority="Mannerheim, 1842" authorityName="Mannerheim" authorityYear="1842" box="[730,1178,1399,1424]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="256" phylum="Arthropoda" rank="species" species="besckei">
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<emphasis id="C18F3F7E2425FFA677B9641DF6DEFA34" bold="true" box="[730,962,1399,1423]" italics="true" pageId="39" pageNumber="256">Physodactylus besckei</emphasis>
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(
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E-I, 34R)
</emphasis>
</paragraph>
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<taxonomicName id="34FB98EF2425FFA677CB64B2F160FA4B" authority="Mannerheim, 1842: 93" authorityName="Mannerheim" authorityPageNumber="93" authorityYear="1842" box="[680,1148,1496,1520]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="256" phylum="Arthropoda" rank="species" species="besckei">
<emphasis id="C18F3F7E2425FFA677CB64B2F664FA4B" box="[680,888,1496,1520]" italics="true" pageId="39" pageNumber="256">Physodactylus besckei</emphasis>
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;
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;
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<bibRefCitation id="976A9E9D2425FFA676CA6492F192F9AB" author="FLEUTIAUX, E." box="[937,1166,1528,1552]" pageId="39" pageNumber="256" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
</treatmentCitation>
, 407;
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<bibRefCitation id="976A9E9D2425FFA677826772F6A1F98B" author="SCHWARZ, O." box="[737,957,1560,1584]" pageId="39" pageNumber="256" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 311</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2425FFA676AC6772F1D4F98B" author="SCHENKLING, S." box="[975,1224,1560,1584]" page="508" pageId="39" pageNumber="256" year="1927">
<bibRefCitation id="976A9E9D2425FFA676AC6772F1D4F98B" author="SCHENKLING, S." box="[975,1224,1560,1584]" pageId="39" pageNumber="256" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 508</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2425FFA677826752F6D9F9EB" author="FLEUTIAUX, E." box="[737,965,1592,1616]" page="168" pageId="39" pageNumber="256" year="1940">
<bibRefCitation id="976A9E9D2425FFA677826752F6D9F9EB" author="FLEUTIAUX, E." box="[737,965,1592,1616]" pageId="39" pageNumber="256" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 168</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2425FFA676B26752F1DAF9EB" author="BLACKWELDER, R. E." box="[977,1222,1592,1616]" page="292" pageId="39" pageNumber="256" year="1944">
<bibRefCitation id="976A9E9D2425FFA676B26752F1DAF9EB" author="BLACKWELDER, R. E." box="[977,1222,1592,1616]" pageId="39" pageNumber="256" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F344E36C2425FFA677CB6732F65CF92B" blockId="39.[680,1229,1496,1680]" pageId="39" pageNumber="256">
<treatmentCitationGroup id="D3EBC4422425FFA677CB6732F65CF92B" pageId="39" pageNumber="256">
<treatmentCitation id="725AC57D2425FFA677CB6732F195F9CB" author="MANNERHEIM, C. G." box="[680,1161,1624,1648]" page="93" pageId="39" pageNumber="256" year="1842">
<taxonomicName id="34FB98EF2425FFA677CB6732F195F9CB" authority="Mannerheim, 1842: 93" authorityName="Mannerheim" authorityPageNumber="93" authorityYear="1842" box="[680,1161,1624,1648]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="256" phylum="Arthropoda" rank="species" species="besckii">
<emphasis id="C18F3F7E2425FFA677CB6732F667F9CB" box="[680,891,1624,1648]" italics="true" pageId="39" pageNumber="256">Physodactylus besckii</emphasis>
<bibRefCitation id="976A9E9D2425FFA676EA6732F195F9CB" author="MANNERHEIM, C. G." box="[905,1161,1624,1648]" pageId="39" pageNumber="256" pagination="93 - 99" refId="ref50735" refString="MANNERHEIM, C. G. 1842. Descripton d'une nouvelle espece de genre Physodactylus. Acta Societatis Scientiarum Fennicae, 1: 93 - 99." type="journal article" year="1842">Mannerheim, 1842: 93</bibRefCitation>
</taxonomicName>
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(misspelling).
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</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72425FFA977CB67D2F129FC74" lastPageId="40" lastPageNumber="257" pageId="39" pageNumber="256" type="description">
<paragraph id="F344E36C2425FFA977CB67D2F7B5FB8B" blockId="39.[680,1229,1720,1872]" lastBlockId="40.[132,681,184,1072]" lastPageId="40" lastPageNumber="257" pageId="39" pageNumber="256">
<emphasis id="C18F3F7E2425FFA677CB67D2F636F96B" box="[680,810,1720,1744]" italics="true" pageId="39" pageNumber="256">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE92425FFA676EE67D2F6EAF96B" box="[909,1014,1720,1744]" captionStart="FIGURE 34" captionStartId="65.[95,174,1749,1768]" captionTargetBox="[94,1227,535,1726]" captionTargetId="figure-138@65.[94,1228,535,1726]" captionTargetPageId="65" captionText="FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="39" pageNumber="256">Fig. 34R</figureCitation>
): Integument light yellowish-brown except for head and antennae dark brown, metaventrite darker on lateral borders, abdominal ventrites 1-4 partly black; pilosity yellow. Total length 11.0-13.0 mm; elytral base 1.0-1.1x as wide as prothorax, elytra 3.0-3.1x longer than pronotum. Frons concave along the anterior border with a transverse weak protuberance between antennal insertions, frontal carina produced; antenna (
<figureCitation id="6BC0FFE9242AFFA974846072F75DFE8B" box="[487,577,280,304]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="40" pageNumber="257">Fig. 16E</figureCitation>
) reaching the base of pronotal posterior angles; antennomere IV-VIII as long as wide; IX and X 1.25 and 1.12x longer than wide, XI oval 1.75x longer than wide. Pronotum (
<figureCitation id="6BC0FFE9242AFFA975DC60F2F406FE0B" box="[191,282,408,432]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="40" pageNumber="257">Fig. 16F</figureCitation>
) convex, 1.12-1.18x wider than long, lateral sides subparallel on posterior 1/3 then abruptly convergent anteriad, posterior angles short, non-carinate; lateral carina complete, dorsally visible only on posterior third; pronotal disc with punctures umbilicate, 1-2 diameters apart, larger and denser on lateral and anterior borders; prosternal process (
<figureCitation id="6BC0FFE9242AFFA9775D6332F7BDFDCB" box="[574,673,600,624]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="40" pageNumber="257">Fig. 16G</figureCitation>
) without subapical tooth. Metatibia with dorsal apical angle not produced to weakly produced; lamellae of pro- and mesotarsomeres I a little smaller than the others, present on metatarsomere I. Elytra subparallel on anterior 2/3 then tapering to apex; punctures of apical striae as large as those of lateral pronotum; elytral interstices weakly convex, smooth, impunctate or scarcely punctate. Abdomen with ventrites 2-3 strongly concave laterally with long decumbent setae on ventrites 2-4, a little shorter on ventrite 1. Pregenitalic segments and aedeagus covered with yellow setae. Sternite VIII with posteromedian margin emarginate, anterior sclerotization inverted V-shaped 0.27x the total width of sternite; sternite IX with apex rounded.
</paragraph>
<paragraph id="F344E36C242AFFA975E76532F1FFFE94" blockId="40.[132,681,1112,1232]" lastBlockId="40.[718,1266,184,304]" pageId="40" pageNumber="257">
<emphasis id="C18F3F7E242AFFA975E76532F5C7FBCB" box="[132,219,1112,1136]" italics="true" pageId="40" pageNumber="257">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9242AFFA9758C6532F465FBCB" box="[239,377,1112,1136]" captionStart="FIGURE 16" captionStartId="39.[94,173,987,1006]" captionTargetBox="[95,1224,173,961]" captionTargetId="figure-294@39.[94,1228,170,964]" captionTargetPageId="39" captionText="FIGURE 16: Physodactylus asper sp. nov. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral). Physodactylus besckei. E, antenna; F, G, prothorax (dorsal, lateral); H, I, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641403" httpUri="https://zenodo.org/record/12641403/files/figure.png" pageId="40" pageNumber="257">Figs. 16H, I</figureCitation>
): Phallobase 0.43x the total length of aedeagus, 1.09x longer than wide, ratio between lateroposterior and median parts 6.12; paramere with ventral surface predominantly membranous except for the lateral border and an anteromedian submarginal band weakly sclerotized; penis with basal strut 0.34x its total length, articulated to the parameres by a sclerotized process; ventral sclerite tapering to apex.
</paragraph>
<paragraph id="F344E36C242AFFA977AD6032F6D0FECB" blockId="40.[718,972,344,368]" box="[718,972,344,368]" pageId="40" pageNumber="257">
<emphasis id="C18F3F7E242AFFA977AD6032F637FECB" box="[718,811,344,368]" italics="true" pageId="40" pageNumber="257">
<typeStatus id="2C405DCE242AFFA977AD6032F63AFECB" box="[718,806,344,368]" pageId="40" pageNumber="257" type="holotype">Holotype</typeStatus>
:
</emphasis>
Not examined.
</paragraph>
<paragraph id="F344E36C242AFFA977AD60F2F6C5FE6B" blockId="40.[718,1266,408,464]" pageId="40" pageNumber="257">
<emphasis id="C18F3F7E242AFFA977AD60F2F688FE0B" box="[718,916,408,432]" italics="true" pageId="40" pageNumber="257">Material examined:</emphasis>
<materialsCitation id="4393E931242AFFA976FC60F2F6C5FE6B" collectionCode="MNHN" country="Brazil" location="Sao Paulo" pageId="40" pageNumber="257" specimenCount="6" specimenCount-male="4" stateProvince="Sao Paulo" typeStatus="holotype">
<collectingCountry id="8BECA3FC242AFFA976FC60F2F6E2FE0B" box="[927,1022,408,432]" name="Brazil" pageId="40" pageNumber="257">BRAZIL</collectingCountry>
.
<specimenCount id="E5FD28E5242AFFA9716860F2F156FE14" box="[1035,1098,408,432]" count="2" pageId="40" pageNumber="257" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA9242AFFA9713860F2F1ABFE0B" box="[1115,1207,408,432]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="40" pageNumber="257">MNHN</collectionCode>
);
<collectingRegion id="313F2D8E242AFFA971AD60F2F616FE6B" country="Brazil" name="Sao Paulo" pageId="40" pageNumber="257">São Paulo</collectingRegion>
,
<specimenCount id="E5FD28E5242AFFA9767660D2F67FFE6B" box="[789,867,440,464]" count="4" pageId="40" pageNumber="257" type="male">4 males</specimenCount>
(
<collectionCode id="95EA7BA9242AFFA9761260D2F6D1FE6B" box="[881,973,440,464]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="40" pageNumber="257">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C242AFFA977AD6092F136FDAB" blockId="40.[718,1066,504,528]" box="[718,1066,504,528]" pageId="40" pageNumber="257">
<emphasis id="C18F3F7E242AFFA977AD6092F64CFDAB" box="[718,848,504,528]" italics="true" pageId="40" pageNumber="257">Distribution:</emphasis>
<collectingCountry id="8BECA3FC242AFFA976356092F6A9FDAB" box="[854,949,504,528]" name="Brazil" pageId="40" pageNumber="257">BRAZIL</collectingCountry>
.
<collectingRegion id="313F2D8E242AFFA976DC6092F13BFDAB" box="[959,1063,504,528]" country="Brazil" name="Sao Paulo" pageId="40" pageNumber="257">São Paulo</collectingRegion>
.
</paragraph>
<paragraph id="F344E36C242AFFA977AD6352F129FC74" blockId="40.[718,1267,568,976]" pageId="40" pageNumber="257">
<emphasis id="C18F3F7E242AFFA977AD6352F63BFDEB" box="[718,807,568,592]" italics="true" pageId="40" pageNumber="257">Remarks:</emphasis>
<typeStatus id="2C405DCE242AFFA976576352F689FDEB" box="[820,917,568,592]" pageId="40" pageNumber="257" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="34FB98EF242AFFA976A26352F18FFDEB" authorityName="Mannerheim" authorityYear="1842" box="[961,1171,568,592]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="257" phylum="Arthropoda" rank="species" species="besckei">
<emphasis id="C18F3F7E242AFFA976A26352F18FFDEB" box="[961,1171,568,592]" italics="true" pageId="40" pageNumber="257">Physodactylus besckei</emphasis>
</taxonomicName>
was not examined, therefore the redescription, based on specimens identified by E. Fleutiaux, requires confirmation. This species is most similar to
<taxonomicName id="34FB98EF242AFFA9713263F3F1C8FD0B" authorityName="Fleutiaux" authorityYear="1892" box="[1105,1236,664,688]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="257" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="C18F3F7E242AFFA9713263F3F1C8FD0B" box="[1105,1236,664,688]" italics="true" pageId="40" pageNumber="257">P. brasiliensis</emphasis>
</taxonomicName>
in the coloration of the dorsal surface, abdominal pilosity, shape and relative length of the antenna. It differs from this species in its frons concave along anterior border, less shiny integument, shape of prothorax, more convex pronotum, which is more produced anteriorly over head, sternite IX with rounded apex, dorsal articulation of penis with process less sclerotized, shape of ventral sclerite and the sclerotization of the anteromedian border of parameres.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,268 @@
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<mods:title id="5443A33F55677E0ED3EEB57B822E5FC3">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<mods:namePart id="63D6A6AD12C3AA653CC6CEDDF70C5737">Rosa, Simone Policena</mods:namePart>
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<subSubSection id="BBE1B0E72428FFAB7657675DF191F9EB" box="[820,1165,1591,1616]" pageId="42" pageNumber="259" type="nomenclature">
<paragraph id="F344E36C2428FFAB7657675DF191F9EB" blockId="42.[820,1165,1591,1648]" box="[820,1165,1591,1616]" pageId="42" pageNumber="259">
<heading id="A80C54002428FFAB7657675DF191F9EB" bold="true" box="[820,1165,1591,1616]" centered="true" fontSize="10" level="2" pageId="42" pageNumber="259" reason="2">
<emphasis id="C18F3F7E2428FFAB7657675DF191F9EB" bold="true" box="[820,1165,1591,1616]" pageId="42" pageNumber="259">
<taxonomicName id="34FB98EF2428FFAB7657675DF128F9F4" authorityName="Rosa" authorityYear="2014" box="[820,1076,1591,1615]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="259" phylum="Arthropoda" rank="species" species="chassaini" status="sp. nov.">
<emphasis id="C18F3F7E2428FFAB7657675DF128F9F4" bold="true" box="[820,1076,1591,1615]" italics="true" pageId="42" pageNumber="259">Physodactylus chassaini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052428FFAB71596752F191F9EB" box="[1082,1165,1592,1616]" pageId="42" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72428FFAD76196732F182FDCB" lastPageId="44" lastPageNumber="261" pageId="42" pageNumber="259" type="description">
<paragraph id="F344E36C2428FFAB76196732F15AF9CB" blockId="42.[820,1165,1591,1648]" box="[890,1094,1624,1648]" pageId="42" pageNumber="259">
<emphasis id="C18F3F7E2428FFAB76196732F15AF9CB" bold="true" box="[890,1094,1624,1648]" pageId="42" pageNumber="259">
(
<figureCitation id="6BC0FFE92428FFAB76E26732F6CAF9CB" box="[897,982,1624,1648]" captionStart="FIGURE 19" captionStartId="43.[95,174,1826,1845]" captionTargetBox="[100,1222,389,1798]" captionTargetId="figure-93@43.[94,1228,385,1802]" captionTargetPageId="43" captionText="FIGURE 19: Physodactylus chassaini sp. nov. A, antenna; B, pronotum; C, mesotibia; D, E, aedeagus (dorsal, ventral). Physodactylus fischeri. F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641409" httpUri="https://zenodo.org/record/12641409/files/figure.png" pageId="42" pageNumber="259">Figs. 19</figureCitation>
A-E, 35C)
</emphasis>
</paragraph>
<paragraph id="F344E36C2428FFAB77AD67F2F6C6F954" blockId="42.[718,1266,1688,1776]" pageId="42" pageNumber="259">
<emphasis id="C18F3F7E2428FFAB77AD67F2F625F90B" box="[718,825,1688,1712]" italics="true" pageId="42" pageNumber="259">Etymology:</emphasis>
In homage to the entomologist Mr. Jacques Chassain, for his dedication to the
<taxonomicName id="34FB98EF2428FFAB712E67D2F1EEF96B" box="[1101,1266,1720,1744]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="259" phylum="Arthropoda" rank="subFamily" subFamily="Physodactylinae">Physodactylinae</taxonomicName>
collection of the MNHN.
</paragraph>
<paragraph id="F344E36C2428FFAD77AD6673F404FC6B" blockId="42.[718,1266,1816,1872]" lastBlockId="44.[132,681,184,976]" lastPageId="44" lastPageNumber="261" pageId="42" pageNumber="259">
<emphasis id="C18F3F7E2428FFAB77AD6673F622F88B" box="[718,830,1817,1840]" italics="true" pageId="42" pageNumber="259">Description</emphasis>
(male,
<figureCitation id="6BC0FFE92428FFAB76FC6672F114F88B" box="[927,1032,1816,1840]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="42" pageNumber="259">Fig. 35C</figureCitation>
): Integument bright; dark brown to black, pronotum reddish-brown with irregular black areas along the borders, epipleurae usually lighter than elytra; pilosity brown. Total length 11.0-15.0 mm; elytral base 1.0-1.1x as wide as prothorax, elytra 3.0-3.25x longer than pronotum. Frons concave, frontal carina not produced; antenna (
<figureCitation id="6BC0FFE92429FFAA766D61D2F673FF6B" box="[782,879,184,208]" captionStart="FIGURE 19" captionStartId="43.[95,174,1826,1845]" captionTargetBox="[100,1222,389,1798]" captionTargetId="figure-93@43.[94,1228,385,1802]" captionTargetPageId="43" captionText="FIGURE 19: Physodactylus chassaini sp. nov. A, antenna; B, pronotum; C, mesotibia; D, E, aedeagus (dorsal, ventral). Physodactylus fischeri. F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641409" httpUri="https://zenodo.org/record/12641409/files/figure.png" pageId="43" pageNumber="260">Fig. 19A</figureCitation>
) reaching the posterior third of pronotum; antennomere IV 1.14x wider than long, V-X 1.28-1.50x wider than long; XI oval 1.22x longer than wide. Pronotum (
<figureCitation id="6BC0FFE92429FFAA76D66072F113FE8B" box="[949,1039,280,304]" captionStart="FIGURE 19" captionStartId="43.[95,174,1826,1845]" captionTargetBox="[100,1222,389,1798]" captionTargetId="figure-93@43.[94,1228,385,1802]" captionTargetPageId="43" captionText="FIGURE 19: Physodactylus chassaini sp. nov. A, antenna; B, pronotum; C, mesotibia; D, E, aedeagus (dorsal, ventral). Physodactylus fischeri. F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641409" httpUri="https://zenodo.org/record/12641409/files/figure.png" pageId="43" pageNumber="260">Fig. 19B</figureCitation>
) 1.16-1.31x wider than long, lateral sides nearly straight to rounded convergent from posterior angles to anterior margin; anteromedian margin weakly produced; posterior angles short, convex, parallel; lateral carina complete prominent nearly or entirely visible dorsally; disc with punctures heterogeneously distributed, 2-8 diameters apart, larger and denser on lateral (2-4 diameters apart) and anterior borders (0.5-1.0 diameters apart); prosternal process with a subapical tooth. Lamellae of pro- and mesotarsomeres I smaller than the others, present on metatarsomere I. Elytra tapering to apex from anterior third; striae with a single row of punctures; interstices convex and weakly rugose, scarcely punctate; apical striae with punctures a little larger than those of pronotal lateral border. Abdomen with ventrites 1-4 strongly concave laterally, ventrite 1 densely punctate with setae about 1/3 as long as those of ventrite 2, lateral border impuctate. Ventrites 2-4 densely punctate with long and semi-erect stout setae, lateral border sparsely punctate with a few fine setae. Pregenitalic segments and aedeagus covered with stout dark-brown setae. Sternite VIII with lateroposterior sclerotizations separate or contiguous medially; posterior margin straight, anterior sclerotization 0.4-0.6x the total width of sternite; sternite IX with apex rounded.
</paragraph>
<caption id="A784B3E42429FFAA753C6648F47FF8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641409" ID-Zenodo-Dep="12641409" httpUri="https://zenodo.org/record/12641409/files/figure.png" pageId="43" pageNumber="260" startId="43.[95,174,1826,1845]" targetBox="[100,1222,389,1798]" targetPageId="43" targetType="figure">
<paragraph id="F344E36C2429FFAA753C6648F47FF8F4" blockId="43.[94,1230,1826,1871]" pageId="43" pageNumber="260">
<emphasis id="C18F3F7E2429FFAA753C6648F5D2F88E" bold="true" box="[95,206,1826,1845]" pageId="43" pageNumber="260">FIGURE 19:</emphasis>
<taxonomicName id="34FB98EF2429FFAA75B06648F49BF88E" authorityName="Rosa" authorityYear="2014" box="[211,391,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="260" phylum="Arthropoda" rank="species" species="chassaini" status="sp. nov.">
<emphasis id="C18F3F7E2429FFAA75B06648F49BF88E" box="[211,391,1826,1845]" italics="true" pageId="43" pageNumber="260">Physodactylus chassaini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052429FFAA74EF6649F4D7F88D" box="[396,459,1827,1846]" pageId="43" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E2429FFAA74AD6648F4FDF88E" bold="true" box="[462,481,1826,1845]" pageId="43" pageNumber="260">A,</emphasis>
antenna;
<emphasis id="C18F3F7E2429FFAA77526648F758F88E" bold="true" box="[561,580,1826,1845]" pageId="43" pageNumber="260">B,</emphasis>
pronotum;
<emphasis id="C18F3F7E2429FFAA77C56648F7A7F88E" bold="true" box="[678,699,1826,1845]" pageId="43" pageNumber="260">C,</emphasis>
mesotibia;
<emphasis id="C18F3F7E2429FFAA767B6648F659F88E" bold="true" box="[792,837,1826,1845]" pageId="43" pageNumber="260">D, E,</emphasis>
aedeagus (dorsal, ventral).
<emphasis id="C18F3F7E2429FFAA71426648F1D2F88E" box="[1057,1230,1826,1845]" italics="true" pageId="43" pageNumber="260">
<taxonomicName id="34FB98EF2429FFAA71426648F1D6F88E" authorityName="Fleutiaux" authorityYear="1940" box="[1057,1226,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="260" phylum="Arthropoda" rank="species" species="fischeri">Physodactylus fischeri</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2429FFAA753D6656F594F8F4" bold="true" box="[94,136,1852,1871]" pageId="43" pageNumber="260">F, G,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C242EFFAD75E76292F78AFB54" blockId="44.[132,681,1016,1264]" pageId="44" pageNumber="261">
<emphasis id="C18F3F7E242EFFAD75E76292F5C1FBAB" box="[132,221,1016,1040]" italics="true" pageId="44" pageNumber="261">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9242EFFAD758C6292F461FBAB" box="[239,381,1016,1040]" captionStart="FIGURE 19" captionStartId="43.[95,174,1826,1845]" captionTargetBox="[100,1222,389,1798]" captionTargetId="figure-93@43.[94,1228,385,1802]" captionTargetPageId="43" captionText="FIGURE 19: Physodactylus chassaini sp. nov. A, antenna; B, pronotum; C, mesotibia; D, E, aedeagus (dorsal, ventral). Physodactylus fischeri. F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641409" httpUri="https://zenodo.org/record/12641409/files/figure.png" pageId="44" pageNumber="261">Figs. 19D, E</figureCitation>
): Phallobase 0.47x the total length of aedeagus, 1.39x longer than wide, length ratio between lateroposterior and median parts 7.4; paramere with ventral surface nearly entirely sclerotized with anteromedian margin distinct, acute and produced anteriad; penis with basal strut 0.20x its total length, articulated to the parameres by a sclerotized short process; ventral sclerite tapering to apex.
</paragraph>
<paragraph id="F344E36C242EFFAD75E76472F751FAEB" blockId="44.[132,680,1304,1360]" pageId="44" pageNumber="261">
<materialsCitation id="4393E931242EFFAD75E76472F751FAEB" collectingDate="1998-12" collectionCode="MG, MZUSP" collectorName="Bello" country="Brazil" location="Aguas Vermelhas" municipality="Vaz de Mello" pageId="44" pageNumber="261" specimenCount="1" typeStatus="holotype">
<emphasis id="C18F3F7E242EFFAD75E76472F5FDFA8B" box="[132,225,1304,1328]" italics="true" pageId="44" pageNumber="261">
<typeStatus id="2C405DCE242EFFAD75E76472F5C0FA8B" box="[132,220,1304,1328]" pageId="44" pageNumber="261" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC242EFFAD758F6472F455FA8B" box="[236,329,1304,1328]" name="Brazil" pageId="44" pageNumber="261">BRASIL</collectingCountry>
,
<collectionCode id="95EA7BA9242EFFAD74316472F49EFA8B" box="[338,386,1304,1328]" country="China" name="Museum of Zoology" pageId="44" pageNumber="261">MG</collectionCode>
:
<location id="F624B5B7242EFFAD74E96472F725FA8B" LSID="urn:lsid:plazi:treatment:7B52527A2428FFAD7657675DF182FDCB:F624B5B7242EFFAD74E96472F725FA8B" box="[394,569,1304,1331]" country="Brazil" municipality="Vaz de Mello" name="Aguas Vermelhas" pageId="44" pageNumber="261">Águas Vermelhas</location>
,
<date id="8745C5AC242EFFAD77216472F7B8FA8B" box="[578,676,1304,1328]" pageId="44" pageNumber="261" value="1998-12">
<collectingDate id="97013C44242EFFAD77216472F7B8FA8B" box="[578,676,1304,1328]" pageId="44" pageNumber="261" value="1998-12">XII.1998</collectingDate>
</date>
,
<collectingMunicipality id="13207916242EFFAD75E76452F416FAEB" box="[132,266,1336,1360]" pageId="44" pageNumber="261">Vaz de Mello</collectingMunicipality>
&amp;
<collectorName id="5E0E86BA242EFFAD744D6452F47FFAEB" box="[302,355,1336,1360]" pageId="44" pageNumber="261">Bello</collectorName>
col.], male (
<collectionCode id="95EA7BA9242EFFAD74806452F75EFAEB" box="[483,578,1336,1360]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="44" pageNumber="261">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C242EFFAD75E76413F401F9CB" blockId="44.[132,681,1400,1648]" pageId="44" pageNumber="261">
<materialsCitation id="4393E931242EFFAD75E76413F405F9CB" collectingDate="1972-11" collectingDateMax="1980-12" collectingDateMin="1972-11" collectionCode="MZUSP, UFMT, CNC, MNRJ, DZUP" collectorName="M. Alvarenga &amp; da Divisa &amp; B. Silva &amp; Colecao M. Alvarenga" country="Brazil" elevation="980" location="Same" pageId="44" pageNumber="261" specimenCount="1022" stateProvince="Bahia" typeStatus="paratype">
<emphasis id="C18F3F7E242EFFAD75E76413F5FAFA2B" box="[132,230,1401,1424]" italics="true" pageId="44" pageNumber="261">
<typeStatus id="2C405DCE242EFFAD75E76413F5FCFA2B" box="[132,224,1401,1424]" pageId="44" pageNumber="261" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<location id="F624B5B7242EFFAD758D6412F439FA2B" LSID="urn:lsid:plazi:treatment:7B52527A2428FFAD7657675DF182FDCB:F624B5B7242EFFAD758D6412F439FA2B" box="[238,293,1400,1424]" country="Brazil" name="Same" pageId="44" pageNumber="261" stateProvince="Bahia">Same</location>
label as
<typeStatus id="2C405DCE242EFFAD741C6412F4C0FA2B" box="[383,476,1400,1424]" pageId="44" pageNumber="261" type="holotype">holotype</typeStatus>
,
<specimenCount id="E5FD28E5242EFFAD748B6412F72DFA34" box="[488,561,1400,1424]" count="33" pageId="44" pageNumber="261" type="generic">33 exs.</specimenCount>
(
<collectionCode id="95EA7BA9242EFFAD775C6412F781FA2B" box="[575,669,1400,1424]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="44" pageNumber="261">MZUSP</collectionCode>
),
<specimenCount id="E5FD28E5242EFFAD75E764F2F5A0FA14" box="[132,188,1432,1456]" count="1" pageId="44" pageNumber="261" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9242EFFAD75AC64F2F43CFA0B" box="[207,288,1432,1456]" country="Brazil" lsid="urn:lsid:biocol.org:col:14332" name="Universidade Federal de Mato Grosso" pageId="44" pageNumber="261" type="Herbarium">UFMT</collectionCode>
); [
<collectingCountry id="8BECA3FC242EFFAD745D64F2F461FA0B" box="[318,381,1432,1456]" name="Brazil" pageId="44" pageNumber="261">Brasil</collectingCountry>
, Encruzilhada,
<specimenCount id="E5FD28E5242EFFAD774B64F2F765FA14" box="[552,633,1432,1456]" count="980" pageId="44" pageNumber="261" type="generic">
<quantity id="34034E89242EFFAD774B64F2F769FA14" box="[552,629,1432,1456]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="44" pageNumber="261" unit="m" value="980.0">
<elevation id="78D6045F242EFFAD774B64F2F769FA14" box="[552,629,1432,1456]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="44" pageNumber="261" unit="m" value="980.0">980 m</elevation>
</quantity>
.
</specimenCount>
<collectingRegion id="313F2D8E242EFFAD77E664F2F5BAFA6B" country="Brazil" name="Bahia" pageId="44" pageNumber="261">Bahia</collectingRegion>
,
<date id="8745C5AC242EFFAD75D664D2F412FA6B" box="[181,270,1464,1488]" pageId="44" pageNumber="261" value="1974-11">
<collectingDate id="97013C44242EFFAD75D664D2F412FA6B" box="[181,270,1464,1488]" pageId="44" pageNumber="261" value="1974-11">XI.1974</collectingDate>
</date>
,
<collectorName id="5E0E86BA242EFFAD747E64D2F4AEFA6B" box="[285,434,1464,1488]" pageId="44" pageNumber="261">M. Alvarenga</collectorName>
],
<specimenCount id="E5FD28E5242EFFAD74A564D2F718FA74" box="[454,516,1464,1488]" count="4" pageId="44" pageNumber="261" type="generic">4 exs.</specimenCount>
(
<collectionCode id="95EA7BA9242EFFAD777564D2F748FA6B" box="[534,596,1464,1488]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="44" pageNumber="261">CNC</collectionCode>
),
<specimenCount id="E5FD28E5242EFFAD770864D2F7B5FA74" box="[619,681,1464,1488]" count="2" pageId="44" pageNumber="261" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA9242EFFAD75EF64B2F5F6FA4B" box="[140,234,1496,1520]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="44" pageNumber="261">MZUSP</collectionCode>
); [Estr. Rio
<collectingRegion id="313F2D8E242EFFAD740164B2F483FA4B" box="[354,415,1496,1520]" country="Brazil" name="Bahia" pageId="44" pageNumber="261">Bahia</collectingRegion>
, km 965, Motel
<collectorName id="5E0E86BA242EFFAD772A64B2F7B4FA4B" box="[585,680,1496,1520]" pageId="44" pageNumber="261">da Divisa</collectorName>
<quantity id="34034E89242EFFAD75E76492F5D6F9B4" box="[132,202,1528,1552]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="44" pageNumber="261" unit="m" value="960.0">
<elevation id="78D6045F242EFFAD75E76492F5D6F9B4" box="[132,202,1528,1552]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="44" pageNumber="261" unit="m" value="960.0">960 m</elevation>
</quantity>
, Encruzilhada,
<collectingRegion id="313F2D8E242EFFAD740A6492F4BAF9AB" box="[361,422,1528,1552]" country="Brazil" name="Bahia" pageId="44" pageNumber="261">Bahia</collectingRegion>
,
<collectingCountry id="8BECA3FC242EFFAD74D36492F4F5F9AB" box="[432,489,1528,1552]" name="Brazil" pageId="44" pageNumber="261">Brasil</collectingCountry>
<date id="8745C5AC242EFFAD748C6492F752F9AB" box="[495,590,1528,1552]" pageId="44" pageNumber="261" value="1980-12">
<collectingDate id="97013C44242EFFAD748C6492F752F9AB" box="[495,590,1528,1552]" pageId="44" pageNumber="261" value="1980-12">XII.1980</collectingDate>
</date>
<collectorName id="5E0E86BA242EFFAD770A6492F583F994" pageId="44" pageNumber="261">B. Silva</collectorName>
],
<specimenCount id="E5FD28E5242EFFAD75D36772F5FDF994" box="[176,225,1560,1584]" count="1" pageId="44" pageNumber="261" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9242EFFAD758D6772F426F98B" box="[238,314,1560,1584]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="44" pageNumber="261">MNRJ</collectionCode>
); [
<collectorName id="5E0E86BA242EFFAD74316772F723F98B" box="[338,575,1560,1584]" pageId="44" pageNumber="261">Coleção M. Alvarenga</collectorName>
], [Encruzilhada,
<quantity id="34034E89242EFFAD75AC6752F408F9F4" box="[207,276,1592,1616]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="44" pageNumber="261" unit="m" value="980.0">
<elevation id="78D6045F242EFFAD75AC6752F408F9F4" box="[207,276,1592,1616]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="44" pageNumber="261" unit="m" value="980.0">980 m</elevation>
</quantity>
,
<collectingRegion id="313F2D8E242EFFAD747C6752F440F9EB" box="[287,348,1592,1616]" country="Brazil" name="Bahia" pageId="44" pageNumber="261">Bahia</collectingRegion>
,
<collectingCountry id="8BECA3FC242EFFAD74056752F4BEF9EB" box="[358,418,1592,1616]" name="Brazil" pageId="44" pageNumber="261">Brasil</collectingCountry>
,
<date id="8745C5AC242EFFAD74CF6752F719F9EB" box="[428,517,1592,1616]" pageId="44" pageNumber="261" value="1972-11">
<collectingDate id="97013C44242EFFAD74CF6752F719F9EB" box="[428,517,1592,1616]" pageId="44" pageNumber="261" value="1972-11">XI.1972</collectingDate>
</date>
,
<collectorName id="5E0E86BA242EFFAD776C6752F783F9EB" box="[527,671,1592,1616]" pageId="44" pageNumber="261">M. Alvarenga</collectorName>
],
<specimenCount id="E5FD28E5242EFFAD75E76732F5AAF9D4" box="[132,182,1624,1648]" count="1" pageId="44" pageNumber="261" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9242EFFAD75A06732F40CF9CB" box="[195,272,1624,1648]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="44" pageNumber="261" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C242EFFAD75E767F2F499F96B" blockId="44.[132,681,1688,1747]" pageId="44" pageNumber="261">
<emphasis id="C18F3F7E242EFFAD75E767F2F41AF90B" box="[132,262,1688,1712]" italics="true" pageId="44" pageNumber="261">Distribution:</emphasis>
<collectingCountry id="8BECA3FC242EFFAD747067F2F46EF90B" box="[275,370,1688,1712]" name="Brazil" pageId="44" pageNumber="261">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E242EFFAD74E267F2F4DEF90B" box="[385,450,1688,1712]" italics="true" pageId="44" pageNumber="261">
<collectingRegion id="313F2D8E242EFFAD74E267F2F4A1F90B" box="[385,445,1688,1712]" country="Brazil" name="Bahia" pageId="44" pageNumber="261">Bahia</collectingRegion>
:
</emphasis>
Encruzilhada;
<emphasis id="C18F3F7E242EFFAD770967F3F5D5F974" italics="true" pageId="44" pageNumber="261">
<collectingRegion id="313F2D8E242EFFAD770967F3F5DFF974" country="Brazil" name="Minas Gerais" pageId="44" pageNumber="261">Minas Gerais</collectingRegion>
:
</emphasis>
Águas Vermelhas.
</paragraph>
<paragraph id="F344E36C242EFFAD75E76792F182FDCB" blockId="44.[132,681,1784,1872]" lastBlockId="44.[718,1267,184,624]" pageId="44" pageNumber="261">
<emphasis id="C18F3F7E242EFFAD75E76792F4D0F8AB" box="[132,460,1784,1808]" italics="true" pageId="44" pageNumber="261">
Remarks:
<taxonomicName id="34FB98EF242EFFAD75846792F4D0F8AB" authorityName="Rosa" authorityYear="2014" box="[231,460,1784,1808]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="chassaini" status="sp. nov.">Physodactylus chassaini</taxonomicName>
</emphasis>
<taxonomicNameLabel id="DABC8205242EFFAD74B76792F739F8B4" box="[468,549,1784,1807]" pageId="44" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<emphasis id="C18F3F7E242EFFAD75E76673F426F88B" box="[132,314,1816,1840]" italics="true" pageId="44" pageNumber="261">
<taxonomicName id="34FB98EF242EFFAD75E76673F5F9F88B" authorityName="Fleutiaux" authorityYear="1940" box="[132,229,1816,1840]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="fischeri">P. fischeri</taxonomicName>
,
<taxonomicName id="34FB98EF242EFFAD75926673F426F88B" authorityName="Rosa" authorityYear="2014" box="[241,314,1817,1840]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="asper" status="sp. nov.">P. asper</taxonomicName>
</emphasis>
<taxonomicNameLabel id="DABC8205242EFFAD74216672F48FF894" box="[322,403,1816,1839]" pageId="44" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="34FB98EF242EFFAD74AB6673F770F88B" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[456,620,1816,1840]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="foveatostriatus">
<emphasis id="C18F3F7E242EFFAD74AB6673F770F88B" box="[456,620,1816,1840]" italics="true" pageId="44" pageNumber="261">P. foveatostriatus</emphasis>
</taxonomicName>
in its color pattern, shape of prothorax, brown pilosity and shape of antenna. It differs from the two first species mainly by its elytral striae with a single row of punctures distinct to apex and interstices smoother with a few sparse punctures. Its abdominal pilosity pattern is most similar to that of
<taxonomicName id="34FB98EF242EFFAD76A36053F115FEEB" authorityName="Rosa" authorityYear="2014" box="[960,1033,313,336]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="asper" status="sp. nov.">
<emphasis id="C18F3F7E242EFFAD76A36053F115FEEB" box="[960,1033,313,336]" italics="true" pageId="44" pageNumber="261">P. asper</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205242EFFAD71726052F17FFEF4" box="[1041,1123,312,335]" pageId="44" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
, but the setae of the later are smaller on ventrite 1 and finer on ventrites 2-4.
<taxonomicName id="34FB98EF242EFFAD76596012F101FE2B" authorityName="Rosa" authorityYear="2014" box="[826,1053,376,400]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="chassaini" status="sp. nov.">
<emphasis id="C18F3F7E242EFFAD76596012F101FE2B" box="[826,1053,376,400]" italics="true" pageId="44" pageNumber="261">Physodactylus chassaini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205242EFFAD71476012F16FFE34" box="[1060,1139,376,399]" pageId="44" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
is also similar to
<taxonomicName id="34FB98EF242EFFAD766D60F3F6ADFE0B" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[782,945,408,432]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="foveatostriatus">
<emphasis id="C18F3F7E242EFFAD766D60F3F6ADFE0B" box="[782,945,408,432]" italics="true" pageId="44" pageNumber="261">P. foveatostriatus</emphasis>
</taxonomicName>
in its elytral striae punctation, but it is distinguished by its smaller punctures of these striae, interstices more convex and by the abdominal pilosity pattern. All these species have aedeagus quite similar, with a few differences mainly on the length of the median part of the phallobase, shapes of the basal struts, ventral sclerite and dorsal articulation.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="89AA1367474719279A382BA1C125CAA2">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<taxonomicName id="34FB98EF242AFFA9779E657DF1DFFB8B" authority="Fleutiaux, 1892" authorityName="Fleutiaux" authorityYear="1892" box="[765,1219,1047,1072]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="257" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="C18F3F7E242AFFA9779E657DF1DFFB8B" bold="true" box="[765,1219,1047,1072]" pageId="40" pageNumber="257">
<emphasis id="C18F3F7E242AFFA9779E657DF10EFB94" bold="true" box="[765,1042,1047,1071]" italics="true" pageId="40" pageNumber="257">Physodactylus brasiliensis</emphasis>
<bibRefCitation id="976A9E9D242AFFA9717B6572F1DFFB8B" author="FLEUTIAUX, E." box="[1048,1219,1048,1072]" pageId="40" pageNumber="257" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892</bibRefCitation>
</emphasis>
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<paragraph id="F344E36C242AFFA976F36552F12CFBEB" blockId="40.[765,1219,1047,1104]" box="[912,1072,1080,1104]" pageId="40" pageNumber="257">
<emphasis id="C18F3F7E242AFFA976F36552F12CFBEB" bold="true" box="[912,1072,1080,1104]" pageId="40" pageNumber="257">
(
<figureCitation id="6BC0FFE9242AFFA976F46552F6F3FBEB" box="[919,1007,1080,1104]" captionStart="FIGURE 17" captionStartId="40.[132,211,1852,1871]" captionTargetBox="[134,1263,1263,1821]" captionTargetId="figure-462@40.[132,1266,1261,1828]" captionTargetPageId="40" captionText="FIGURE 17: Physodactylus brasiliensis. A, antenna; B, C, prothorax (dorsal, lateral); D, metatibia; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641405" httpUri="https://zenodo.org/record/12641405/files/figure.png" pageId="40" pageNumber="257">Figs. 17</figureCitation>
,
<figureCitation id="6BC0FFE9242AFFA976986552F135FBEB" box="[1019,1065,1080,1104]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="40" pageNumber="257">35A</figureCitation>
)
</emphasis>
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<paragraph id="F344E36C242AFFA877AD6512F72CFAD4" blockId="40.[718,1267,1144,1232]" lastBlockId="41.[94,643,184,1392]" lastPageId="41" lastPageNumber="258" pageId="40" pageNumber="257">
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<treatmentCitation id="725AC57D242AFFA977AD6512F1A9FB2B" author="FLEUTIAUX, E." box="[718,1205,1144,1168]" page="406" pageId="40" pageNumber="257" year="1892">
<taxonomicName id="34FB98EF242AFFA977AD6512F1A9FB2B" authority="Fleutiaux, 1892: 406" authorityName="Fleutiaux" authorityPageNumber="406" authorityYear="1892" box="[718,1205,1144,1168]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="257" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="C18F3F7E242AFFA977AD6512F6DBFB2B" box="[718,967,1144,1168]" italics="true" pageId="40" pageNumber="257">Physodactylus brasiliensis</emphasis>
<bibRefCitation id="976A9E9D242AFFA976B26512F1A9FB2B" author="FLEUTIAUX, E." box="[977,1205,1144,1168]" pageId="40" pageNumber="257" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
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, 408;
<treatmentCitation id="725AC57D242AFFA9766465F2F6FAFB0B" author="SCHWARZ, O." box="[775,998,1176,1200]" page="311" pageId="40" pageNumber="257" year="1906">
<bibRefCitation id="976A9E9D242AFFA9766465F2F6FAFB0B" author="SCHWARZ, O." box="[775,998,1176,1200]" pageId="40" pageNumber="257" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 311</bibRefCitation>
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;
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<bibRefCitation id="976A9E9D242AFFA9769A65F2F1F1FB0B" author="FLEUTIAUX, E." box="[1017,1261,1176,1200]" pageId="40" pageNumber="257" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 168</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D242AFFA9766465D2F6EDFB6B" author="SCHENKLING, S." box="[775,1009,1208,1232]" page="508" pageId="40" pageNumber="257" year="1927">
<bibRefCitation id="976A9E9D242AFFA9766465D2F6EDFB6B" author="SCHENKLING, S." box="[775,1009,1208,1232]" pageId="40" pageNumber="257" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 508</bibRefCitation>
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;
<treatmentCitation id="725AC57D242AFFA9769865D2F1F3FB6B" author="BLACKWELDER, R. E." box="[1019,1263,1208,1232]" page="292" pageId="40" pageNumber="257" year="1944">
<bibRefCitation id="976A9E9D242AFFA9769865D2F1F3FB6B" author="BLACKWELDER, R. E." box="[1019,1263,1208,1232]" pageId="40" pageNumber="257" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
.
<emphasis id="C18F3F7E242BFFA8753D61D2F5FCFF6B" box="[94,224,184,208]" italics="true" pageId="41" pageNumber="258">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9242BFFA8744F61D2F494FF6B" box="[300,392,184,208]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="41" pageNumber="258">Fig. 35A</figureCitation>
): Integument very shiny, head dark brown with frons lighter, antennomeres I-III reddish-brown, IV-XI dark brown; pronotum light yellowish-brown or ochre with posterior angles, anterior and posterior borders black; scutellum with borders darker, elytra light brown with surrounding area of punctures darker; ventral surface dark brown to black, except for the posterior part of prosternum, epipleura, femur and lateral border of ventrites 1-4 and ventrite 5 light brown; pilosity yellow. Total length 10.0-14.0 mm; elytral base 1.10-1.12x wider than prothorax, elytra 3.12-3.33x longer than pronotum. Frons with a triangular concave area, frontal carina produced; antenna (
<figureCitation id="6BC0FFE9242BFFA8740D6332F4D4FDCB" box="[366,456,600,624]" captionStart="FIGURE 17" captionStartId="40.[132,211,1852,1871]" captionTargetBox="[134,1263,1263,1821]" captionTargetId="figure-462@40.[132,1266,1261,1828]" captionTargetPageId="40" captionText="FIGURE 17: Physodactylus brasiliensis. A, antenna; B, C, prothorax (dorsal, lateral); D, metatibia; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641405" httpUri="https://zenodo.org/record/12641405/files/figure.png" pageId="41" pageNumber="258">Fig. 17A</figureCitation>
) reaching the base to apex of the posterior pronotal angles; antennomere IV-IX as long as wide; X 1.28x longer than wide; XI oval 1.75x longer than wide. Pronotum weakly convex (
<figureCitation id="6BC0FFE9242BFFA875F063B2F5EFFD4B" box="[147,243,728,752]" captionStart="FIGURE 17" captionStartId="40.[132,211,1852,1871]" captionTargetBox="[134,1263,1263,1821]" captionTargetId="figure-462@40.[132,1266,1261,1828]" captionTargetPageId="40" captionText="FIGURE 17: Physodactylus brasiliensis. A, antenna; B, C, prothorax (dorsal, lateral); D, metatibia; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641405" httpUri="https://zenodo.org/record/12641405/files/figure.png" pageId="41" pageNumber="258">Fig. 17B</figureCitation>
) 1.17-1.19x wider than long, lateral sides straight and roundly narrowed from posterior angle to anterior margin, posterior angles long, narrow and carinate; lateral carina nearly reaching the anterior margin, nearly entirely visible dorsally; disc with punctures umbilicate, 1-2 diameters apart, a little larger and denser on lateral and anterior borders; prosternal process (
<figureCitation id="6BC0FFE9242BFFA8743D62D2F4A7FC6B" box="[350,443,952,976]" captionStart="FIGURE 17" captionStartId="40.[132,211,1852,1871]" captionTargetBox="[134,1263,1263,1821]" captionTargetId="figure-462@40.[132,1266,1261,1828]" captionTargetPageId="40" captionText="FIGURE 17: Physodactylus brasiliensis. A, antenna; B, C, prothorax (dorsal, lateral); D, metatibia; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641405" httpUri="https://zenodo.org/record/12641405/files/figure.png" pageId="41" pageNumber="258">Fig. 17C</figureCitation>
) without subapical tooth. Lamellae of pro- and mesotarsomere I a little smaller than the others, present on metatarsomere I. Elytra tapering apicad from humerus or anterior third, punctures of apical striae larger than those of the pronotal lateral border; elytral interstices weakly convex, smooth, sparsely punctate. Abdomen with ventrites 2-3 strongly concave laterally, with long decumbent setae on ventrites 2-4, a little shorter on ventrite 1. Pregenitalic segments and aedeagus covered with yellow to light-brown setae. Sternite VIII with posterior margin straight or emarginate medially, anterior sclerotization inverted V-shaped 0.28x the total width of sternite; sternite IX with apex acute.
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</paragraph>
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<paragraph id="F344E36C242AFFA975E76656F1B3F8F4" blockId="40.[132,1199,1852,1871]" box="[132,1199,1852,1871]" pageId="40" pageNumber="257">
<emphasis id="C18F3F7E242AFFA975E76656F5E8F8F4" bold="true" box="[132,244,1852,1871]" pageId="40" pageNumber="257">FIGURE 17:</emphasis>
<emphasis id="C18F3F7E242AFFA975996656F4D8F8F4" box="[250,452,1852,1871]" italics="true" pageId="40" pageNumber="257">
<taxonomicName id="34FB98EF242AFFA975996656F4DCF8F4" authorityName="Fleutiaux" authorityYear="1892" box="[250,448,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="257" phylum="Arthropoda" rank="species" species="brasiliensis">Physodactylus brasiliensis</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E242AFFA974AB6656F4C7F8F4" bold="true" box="[456,475,1852,1871]" pageId="40" pageNumber="257">A,</emphasis>
antenna;
<emphasis id="C18F3F7E242AFFA9774F6656F745F8F4" bold="true" box="[556,601,1852,1871]" pageId="40" pageNumber="257">B, C,</emphasis>
prothorax (dorsal, lateral);
<emphasis id="C18F3F7E242AFFA976596656F64DF8F4" bold="true" box="[826,849,1852,1871]" pageId="40" pageNumber="257">D,</emphasis>
metatibia;
<emphasis id="C18F3F7E242AFFA976CD6656F6C8F8F4" bold="true" box="[942,980,1852,1871]" pageId="40" pageNumber="257">E, F,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C242BFFA8753D64F2F74EF92B" blockId="41.[94,643,1432,1680]" pageId="41" pageNumber="258">
<emphasis id="C18F3F7E242BFFA8753D64F2F5ABFA0B" box="[94,183,1432,1456]" italics="true" pageId="41" pageNumber="258">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9242BFFA875A964F2F44FFA0B" box="[202,339,1432,1456]" captionStart="FIGURE 17" captionStartId="40.[132,211,1852,1871]" captionTargetBox="[134,1263,1263,1821]" captionTargetId="figure-462@40.[132,1266,1261,1828]" captionTargetPageId="40" captionText="FIGURE 17: Physodactylus brasiliensis. A, antenna; B, C, prothorax (dorsal, lateral); D, metatibia; E, F, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641405" httpUri="https://zenodo.org/record/12641405/files/figure.png" pageId="41" pageNumber="258">Figs. 17E, F</figureCitation>
): Phallobase 0.49x the total length of aedeagus, 1.36x longer than wide, length ratio between lateroposterior and median parts 5.70; paramere with ventral surface predominantly membranous except for the lateral border and anterior margin sclerotized; penis with basal strut 0.27x its total length, articulated to the parameres by a strongly sclerotized process; ventral sclerite parallel-sided.
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242BFFA8753D67D2F690FDEB" pageId="41" pageNumber="258" type="materials_examined">
<paragraph id="F344E36C242BFFA8753D67D2F4DBF8EB" blockId="41.[94,643,1720,1872]" pageId="41" pageNumber="258">
<emphasis id="C18F3F7E242BFFA8753D67D2F5A7F96B" box="[94,187,1720,1744]" italics="true" pageId="41" pageNumber="258">
<typeStatus id="2C405DCE242BFFA8753D67D2F5AAF96B" box="[94,182,1720,1744]" pageId="41" pageNumber="258" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC242BFFA875AC67D2F412F96B" box="[207,270,1720,1744]" name="Brazil" pageId="41" pageNumber="258">Brèsil</collectingCountry>
, Caraça, P. Germain, 2º semestre 1884], [
<taxonomicName id="34FB98EF242BFFA875D267B2F4E3F94B" authority="J Fleut." authorityName="J Fleut." box="[177,511,1751,1776]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="258" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="C18F3F7E242BFFA875D267B2F4B8F94B" box="[177,420,1752,1776]" italics="true" pageId="41" pageNumber="258">Physodactylus brasiliensis</emphasis>
J Fleut.
</taxonomicName>
Type
<collectingCountry id="8BECA3FC242BFFA8775967B2F765F94B" box="[570,633,1752,1776]" name="Brazil" pageId="41" pageNumber="258">Brèsil</collectingCountry>
], [Fltx. 1891], [Museum Paris ex Coll.
<collectionCode id="95EA7BA9242BFFA874936792F718F8B4" box="[496,516,1784,1807]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="41" pageNumber="258">R</collectionCode>
. Oberthur], [Type], [
<typeStatus id="2C405DCE242BFFA875D76672F43DF88B" box="[180,289,1816,1840]" pageId="41" pageNumber="258" type="lectotype">Lectotype</typeStatus>
], [
<taxonomicName id="34FB98EF242BFFA8745A6672F5A6F8EB" authority="Fleut. J. Chassain" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="258" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="C18F3F7E242BFFA8745A6672F730F88B" box="[313,556,1816,1840]" italics="true" pageId="41" pageNumber="258">Physodactylus brasiliensis</emphasis>
Fleut. J. Chassain
</taxonomicName>
det. 05], male (
<collectionCode id="95EA7BA9242BFFA8743C6652F4A7F8EB" box="[351,443,1848,1872]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="258">MNHN</collectionCode>
).
</paragraph>
<paragraph id="F344E36C242BFFA877CB61D2F6F2FEAB" blockId="41.[680,1228,184,272]" pageId="41" pageNumber="258">
<emphasis id="C18F3F7E242BFFA877CB61D2F668FF6B" box="[680,884,184,208]" italics="true" pageId="41" pageNumber="258">Material examined.</emphasis>
<materialsCitation id="4393E931242BFFA876E261D2F6B4FF4B" collectionCode="MNHN" country="Brazil" location="Monlevade" pageId="41" pageNumber="258" specimenCount="1" stateProvince="Minas Gerais">
<collectingCountry id="8BECA3FC242BFFA876E261D2F6FCFF6B" box="[897,992,184,208]" name="Brazil" pageId="41" pageNumber="258">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E242BFFA8769261D3F19EFF74" box="[1009,1154,184,208]" italics="true" pageId="41" pageNumber="258">
<collectingRegion id="313F2D8E242BFFA8769261D3F161FF74" box="[1009,1149,184,208]" country="Brazil" name="Minas Gerais" pageId="41" pageNumber="258">Minas Gerais</collectingRegion>
:
</emphasis>
<location id="F624B5B7242BFFA871F261D2F7F6FF4B" LSID="urn:lsid:plazi:treatment:7B52527A242AFFA8779E657DF690FDEB:F624B5B7242BFFA871F261D2F7F6FF4B" country="Brazil" name="Monlevade" pageId="41" pageNumber="258" stateProvince="Minas Gerais">Monlevade</location>
,
<specimenCount id="E5FD28E5242BFFA8779961B2F62DFF54" box="[762,817,216,240]" count="1" pageId="41" pageNumber="258" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9242BFFA8762761B2F6BCFF4B" box="[836,928,216,240]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="258">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4393E931242BFFA876DB61B2F1D4FF4B" box="[952,1224,216,240]" collectionCode="MZUSP" country="Brazil" location="Caraca" pageId="41" pageNumber="258" specimenCount="2" stateProvince="Minas Gerais">
<location id="F624B5B7242BFFA876DB61B2F11DFF4B" LSID="urn:lsid:plazi:treatment:7B52527A242AFFA8779E657DF690FDEB:F624B5B7242BFFA876DB61B2F11DFF4B" box="[952,1025,216,240]" country="Brazil" name="Caraca" pageId="41" pageNumber="258" stateProvince="Minas Gerais">Caraça</location>
,
<specimenCount id="E5FD28E5242BFFA8717261B2F14CFF54" box="[1041,1104,216,240]" count="2" pageId="41" pageNumber="258" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA9242BFFA8710061B2F1DDFF4B" box="[1123,1217,216,240]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="258">MZUSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4393E931242BFFA877CB6192F6F6FEAB" box="[680,1002,248,272]" collectionCode="MZUSP" country="Brazil" location="Serra do Cipo" pageId="41" pageNumber="258" specimenCount="1" stateProvince="Minas Gerais">
<location id="F624B5B7242BFFA877CB6192F627FEAB" LSID="urn:lsid:plazi:treatment:7B52527A242AFFA8779E657DF690FDEB:F624B5B7242BFFA877CB6192F627FEAB" box="[680,827,248,272]" country="Brazil" name="Serra do Cipo" pageId="41" pageNumber="258" stateProvince="Minas Gerais">Serra do Cipó</location>
,
<specimenCount id="E5FD28E5242BFFA876266192F66BFEB4" box="[837,887,248,272]" count="1" pageId="41" pageNumber="258" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9242BFFA876E76192F6FFFEAB" box="[900,995,248,272]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="258">MZUSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C242BFFA877CB6052F128FECB" blockId="41.[680,1229,312,368]" pageId="41" pageNumber="258">
<emphasis id="C18F3F7E242BFFA877CB6052F636FEEB" box="[680,810,312,336]" italics="true" pageId="41" pageNumber="258">Distribution:</emphasis>
<collectingCountry id="8BECA3FC242BFFA876596052F685FEEB" box="[826,921,312,336]" name="Brazil" pageId="41" pageNumber="258">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E242BFFA876C86053F122FEF4" box="[939,1086,312,336]" italics="true" pageId="41" pageNumber="258">
<collectingRegion id="313F2D8E242BFFA876C86053F124FEF4" box="[939,1080,312,336]" country="Brazil" name="Minas Gerais" pageId="41" pageNumber="258">Minas Gerais</collectingRegion>
:
</emphasis>
João Monlevade; Serra do Caraça; Serra do Cipó.
</paragraph>
<paragraph id="F344E36C242BFFA877CB60F2F690FDEB" blockId="41.[680,1229,408,592]" pageId="41" pageNumber="258">
<emphasis id="C18F3F7E242BFFA877CB60F2F61EFE0B" box="[680,770,408,432]" italics="true" pageId="41" pageNumber="258">Remarks:</emphasis>
The specimen with the Fleutiauxs label has terminalia with light-brown setae and sides of penis more rounded.
<taxonomicName id="34FB98EF242BFFA8762560B2F125FE4B" box="[838,1081,472,496]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="258" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="C18F3F7E242BFFA8762560B2F125FE4B" box="[838,1081,472,496]" italics="true" pageId="41" pageNumber="258">Physodactylus brasiliensis</emphasis>
</taxonomicName>
is distinguishable from the other species in its pattern color, the very shiny integument and the long and carinate posterior pronotal angles.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,272 @@
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<mods:title id="962171552A8B99B39831BF88342C73A3">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<mods:namePart id="751A8935694FBDAC2036CB8935F6CCE5">Rosa, Simone Policena</mods:namePart>
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<treatment id="7B52527A242BFFAB766E63FDF7B4F8EB" LSID="urn:lsid:plazi:treatment:7B52527A242BFFAB766E63FDF7B4F8EB" httpUri="http://treatment.plazi.org/id/7B52527A242BFFAB766E63FDF7B4F8EB" lastPageId="42" lastPageNumber="259" pageId="41" pageNumber="258">
<subSubSection id="BBE1B0E7242BFFA8766E63FDF174FD0B" box="[781,1128,663,688]" pageId="41" pageNumber="258" type="nomenclature">
<paragraph id="F344E36C242BFFA8766E63FDF174FD0B" blockId="41.[781,1128,663,720]" box="[781,1128,663,688]" pageId="41" pageNumber="258">
<heading id="A80C5400242BFFA8766E63FDF174FD0B" bold="true" box="[781,1128,663,688]" centered="true" fontSize="10" level="2" pageId="41" pageNumber="258" reason="2">
<emphasis id="C18F3F7E242BFFA8766E63FDF174FD0B" bold="true" box="[781,1128,663,688]" pageId="41" pageNumber="258">
<taxonomicName id="34FB98EF242BFFA8766E63FDF113FD14" authorityName="Rosa" authorityYear="2014" box="[781,1039,663,687]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="258" phylum="Arthropoda" rank="species" species="brunneus" status="sp. nov.">
<emphasis id="C18F3F7E242BFFA8766E63FDF113FD14" bold="true" box="[781,1039,663,687]" italics="true" pageId="41" pageNumber="258">Physodactylus brunneus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205242BFFA8717563F2F174FD0B" box="[1046,1128,664,688]" pageId="41" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242BFFAB760963D2F7B4F8EB" lastPageId="42" lastPageNumber="259" pageId="41" pageNumber="258" type="description">
<paragraph id="F344E36C242BFFA8760963D2F117FD6B" blockId="41.[781,1128,663,720]" box="[874,1035,696,720]" pageId="41" pageNumber="258">
<emphasis id="C18F3F7E242BFFA8760963D2F117FD6B" bold="true" box="[874,1035,696,720]" pageId="41" pageNumber="258">
(
<figureCitation id="6BC0FFE9242BFFA8761263D2F6D5FD6B" box="[881,969,696,720]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="41" pageNumber="258">Figs. 18</figureCitation>
,
<figureCitation id="6BC0FFE9242BFFA876B663D2F11FFD6B" box="[981,1027,696,720]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="41" pageNumber="258">35B</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="F344E36C242BFFA877CB6392F6C5FC8B" blockId="41.[680,1228,760,816]" pageId="41" pageNumber="258">
<emphasis id="C18F3F7E242BFFA877CB6392F608FCAB" box="[680,788,760,784]" italics="true" pageId="41" pageNumber="258">Etymology:</emphasis>
From latin,
<taxonomicName id="34FB98EF242BFFA876F56392F6EDFCAB" authorityName="Rosa" authorityYear="2014" box="[918,1009,760,784]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="258" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="C18F3F7E242BFFA876F56392F6EDFCAB" box="[918,1009,760,784]" italics="true" pageId="41" pageNumber="258">brunneus</emphasis>
</taxonomicName>
= brown; alluding to the body color of this species.
</paragraph>
<paragraph id="F344E36C242BFFAB77CB6233F6AEFBCB" blockId="41.[680,1229,856,1872]" lastBlockId="42.[132,1267,1006,1552]" lastPageId="42" lastPageNumber="259" pageId="41" pageNumber="258">
<emphasis id="C18F3F7E242BFFA877CB6233F60BFCCB" box="[680,791,857,880]" italics="true" pageId="41" pageNumber="258">Description</emphasis>
(male,
<figureCitation id="6BC0FFE9242BFFA8760A6232F6DBFCCB" box="[873,967,856,880]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="41" pageNumber="258">Fig. 35B</figureCitation>
): Integument very shiny; head and antenna dark brown to black, ventral pro- and pterothorax and legs dark brown, pronotum and elytra equally brown or with pronotum darker; ventral abdomen brown with irregular areas lighter; pilosity light brown to brown. Total length 8.0-11.0 mm; elytral base 1.0-1.1x wider than prothorax, elytra 3.1-3.3x longer than pronotum. Frons (
<figureCitation id="6BC0FFE9242BFFA871216552F180FBEB" box="[1090,1180,1080,1104]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="41" pageNumber="258">Fig. 18B</figureCitation>
) flat to slightly concave, frontal carina not produced anteriorly; antenna (
<figureCitation id="6BC0FFE9242BFFA8765F6512F68AFB2B" box="[828,918,1144,1168]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="41" pageNumber="258">Fig. 18A</figureCitation>
) reaching the base of posterior angles; antennomere IV-VIII 1.1x longer than wide, IX-X 1.2-1.3x longer than wide, XI elliptical 2.1x longer than wide. Pronotum (
<figureCitation id="6BC0FFE9242BFFA876A665B2F138FB4B" box="[965,1060,1240,1264]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="41" pageNumber="258">Fig. 18C</figureCitation>
) weakly convex 1.12-1.20 wider than long, lateral sides nearly straight to rounded convergent anteriad from posterior angles or anterior third; anteromedian margin slightly produced anteriad; posterior angles short, convex, parallel; lateral carina (
<figureCitation id="6BC0FFE9242BFFA876556412F692FA2B" box="[822,910,1400,1424]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="41" pageNumber="258">Fig. 18E</figureCitation>
) prominent and visible dorsally on posterior 1/3, absent on anterior 1/3-1/2; disc with punctures homogenously or heterogeneously distributed, 2-3 diameters apart, larger and denser on lateral and anterior borders; prosternal process (
<figureCitation id="6BC0FFE9242BFFA871206492F1D9F9AB" box="[1091,1221,1528,1552]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="41" pageNumber="258">Figs. 18D, E</figureCitation>
) with a subapical tooth. Lamellae of pro- and mesotarsomeres I smaller than the others, minuscule or absent on metatarsomere I. Elytra parallel-sided on anterior 2/3 then tapering to apex, striae with a row of punctures; interstices rugose, flat, sparsely punctate; apical striae with punctures a little larger than those of pronotal lateral border. Abdomen with ventrites 2-4 strongly concave laterally, ventrite 1 impuctate on anterior and median inner 2/3 with setae a little smaller than those of ventrite 2 on lateral and posterior borders; ventrites 2-4 with long decumbent and semi-erect setae, MG, Sítio Recanto Paraíso,
<date id="8745C5AC2428FFAB76936552F12AFBEB" box="[1008,1078,1080,1104]" pageId="42" pageNumber="259" value="2008-01">I.2008</date>
Luz, M.R. Santos sparser on lateral borders. Pregenitalic segments and col.],
<specimenCount id="E5FD28E52428FFAB766A6532F626FBD4" box="[777,826,1112,1136]" count="1" pageId="42" pageNumber="259" type="generic">1 ex.</specimenCount>
(MZUSP).
</paragraph>
<caption id="A784B3E42428FFAB75E56284F13AFBA0" ID-DOI="http://doi.org/10.5281/zenodo.12641407" ID-Zenodo-Dep="12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="42" pageNumber="259" startId="42.[134,213,1006,1025]" targetBox="[134,1266,176,973]" targetPageId="42" targetType="figure">
<paragraph id="F344E36C2428FFAB75E56284F13AFBA0" blockId="42.[132,1267,1006,1552]" pageId="42" pageNumber="259">
<emphasis id="C18F3F7E2428FFAB75E56284F5EAFBBA" bold="true" box="[134,246,1006,1025]" pageId="42" pageNumber="259">FIGURE 18:</emphasis>
<taxonomicName id="34FB98EF2428FFAB759D6284F4A5FBBA" authorityName="Rosa" authorityYear="2014" box="[254,441,1006,1025]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="259" phylum="Arthropoda" rank="species" species="brunneus" status="sp. nov.">
<emphasis id="C18F3F7E2428FFAB759D6284F4A5FBBA" box="[254,441,1006,1025]" italics="true" pageId="42" pageNumber="259">Physodactylus brunneus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052428FFAB74A36285F71EFBB9" box="[448,514,1007,1026]" pageId="42" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E2428FFAB776B6284F707FBBA" bold="true" box="[520,539,1006,1025]" pageId="42" pageNumber="259">A,</emphasis>
antenna;
<emphasis id="C18F3F7E2428FFAB77136285F79FFBB9" bold="true" box="[624,643,1007,1026]" pageId="42" pageNumber="259">B,</emphasis>
head and mandible (dorsal);
<emphasis id="C18F3F7E2428FFAB76E36284F6D7FBB9" bold="true" box="[896,971,1006,1026]" pageId="42" pageNumber="259">C, D, E,</emphasis>
prothorax (dorsal, ventral, lateral);
<emphasis id="C18F3F7E2428FFAB75E76562F589FBA0" bold="true" box="[132,149,1032,1051]" pageId="42" pageNumber="259">F,</emphasis>
mesotibia;
<emphasis id="C18F3F7E2428FFAB75916562F414FBA0" bold="true" box="[242,264,1032,1051]" pageId="42" pageNumber="259">G,</emphasis>
sternite IX;
<emphasis id="C18F3F7E2428FFAB740D6562F499FBA0" bold="true" box="[366,389,1032,1051]" pageId="42" pageNumber="259">H,</emphasis>
tergites IX and X;
<emphasis id="C18F3F7E2428FFAB77426562F733FBA0" bold="true" box="[545,559,1032,1051]" pageId="42" pageNumber="259">I,</emphasis>
sternite VIII;
<emphasis id="C18F3F7E2428FFAB77C76562F7AEFBA0" bold="true" box="[676,690,1032,1051]" pageId="42" pageNumber="259">J,</emphasis>
tergite VIII;
<emphasis id="C18F3F7E2428FFAB767D6562F656FBA0" bold="true" box="[798,842,1032,1051]" pageId="42" pageNumber="259">K, L,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C2428FFAB75E76512F7B5FB2B" blockId="42.[132,1267,1006,1552]" box="[132,681,1144,1168]" pageId="42" pageNumber="259">aedeagus covered with stout light-brown setae. Sternite</paragraph>
<paragraph id="F344E36C2428FFAB75E765F2F68AFB6B" blockId="42.[132,1267,1006,1552]" pageId="42" pageNumber="259">
VIII (
<figureCitation id="6BC0FFE92428FFAB75DD65F2F413FB0B" box="[190,271,1176,1200]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="42" pageNumber="259">Fig. 18I</figureCitation>
) with posterior margin straight, anterior
<emphasis id="C18F3F7E2428FFAB77AD65F2F64CFB0B" box="[718,848,1176,1200]" italics="true" pageId="42" pageNumber="259">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2428FFAB763F65F2F6A7FB0B" box="[860,955,1176,1200]" name="Brazil" pageId="42" pageNumber="259">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2428FFAB76A865F2F110FB0B" box="[971,1036,1176,1200]" italics="true" pageId="42" pageNumber="259">
<collectingRegion id="313F2D8E2428FFAB76A865F2F11BFB0B" box="[971,1031,1176,1200]" country="Brazil" name="Bahia" pageId="42" pageNumber="259">Bahia</collectingRegion>
:
</emphasis>
Encruzilhada;
<emphasis id="C18F3F7E2428FFAB71D765F3F1EFFB0B" box="[1204,1267,1177,1200]" italics="true" pageId="42" pageNumber="259">Minas</emphasis>
sclerotization linear 0.55x the total width of sternite;
<emphasis id="C18F3F7E2428FFAB77AD65D2F60EFB74" box="[718,786,1208,1231]" italics="true" pageId="42" pageNumber="259">Gerais:</emphasis>
Frei Gaspar.
</paragraph>
<paragraph id="F344E36C2428FFAB75E765B2F70EFB54" blockId="42.[132,1267,1006,1552]" box="[132,530,1240,1264]" pageId="42" pageNumber="259">
sternite IX (
<figureCitation id="6BC0FFE92428FFAB759E65B2F446FB4B" box="[253,346,1240,1264]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="42" pageNumber="259">Fig. 18G</figureCitation>
) tapering to apex.
</paragraph>
<paragraph id="F344E36C2428FFAB77AD6592F6F3FA4B" blockId="42.[132,1267,1006,1552]" pageId="42" pageNumber="259">
<emphasis id="C18F3F7E2428FFAB77AD6592F634FAAB" box="[718,808,1272,1296]" italics="true" pageId="42" pageNumber="259">Remarks:</emphasis>
The unique combination of characters of
<emphasis id="C18F3F7E2428FFAB75E76472F5C1FA8B" box="[132,221,1304,1328]" italics="true" pageId="42" pageNumber="259">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92428FFAB758C6472F467FA8B" box="[239,379,1304,1328]" captionStart="FIGURE 18" captionStartId="42.[134,213,1006,1025]" captionTargetBox="[134,1266,176,973]" captionTargetId="figure-293@42.[132,1266,170,983]" captionTargetPageId="42" captionText="FIGURE 18: Physodactylus brunneus sp. nov. A, antenna; B, head and mandible (dorsal); C, D, E, prothorax (dorsal, ventral, lateral); F, mesotibia; G, sternite IX; H, tergites IX and X; I, sternite VIII; J, tergite VIII; K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641407" httpUri="https://zenodo.org/record/12641407/files/figure.png" pageId="42" pageNumber="259">Figs. 18K, L</figureCitation>
): Phallobase 0.45x the total
<taxonomicName id="34FB98EF2428FFAB77AD6473F65AFA8B" authorityName="Rosa" authorityYear="2014" box="[718,838,1304,1328]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="259" phylum="Arthropoda" rank="species" species="brunneus" status="sp. nov.">
<emphasis id="C18F3F7E2428FFAB77AD6473F65AFA8B" box="[718,838,1304,1328]" italics="true" pageId="42" pageNumber="259">P. brunneus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052428FFAB76326472F6B9FA94" box="[849,933,1304,1327]" pageId="42" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
includes antenna reaching the length of aedeagus, 1.32x longer than wide, length base of pronotum, lateral carina incomplete, elytral ratio between lateroposterior and median parts 6.1; striae with a row of punctures, interstices flat and venparamere with ventral surface weakly sclerotized lat- trite 1 partly impuctate. It is most similar and symerally, anterior margin sclerotized and contiguous patric to
<emphasis id="C18F3F7E2428FFAB764C64F3F695FA0B" box="[815,905,1432,1456]" italics="true" pageId="42" pageNumber="259">
<taxonomicName id="34FB98EF2428FFAB764C64F3F699FA0B" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" box="[815,901,1432,1456]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="259" phylum="Arthropoda" rank="species" species="pujoli">P. pujoli</taxonomicName>
,
</emphasis>
from which is distinguished in its medially; penis with basal strut 0.28x its total length, larger pronotal punctation, color pattern, abdominal articulated to the parameres by a translucent mem- pilosity and aedeagus shape.
</paragraph>
<paragraph id="F344E36C2428FFAB75E76492F70FF9B4" blockId="42.[132,1267,1006,1552]" box="[132,531,1528,1552]" pageId="42" pageNumber="259">brane; ventral sclerite tapering to apex.</paragraph>
<paragraph id="F344E36C2428FFAB75E76752F72EF9CB" blockId="42.[132,681,1592,1648]" pageId="42" pageNumber="259">
<materialsCitation id="4393E9312428FFAB75E76752F72EF9CB" collectingDate="1974-11" collectionCode="MZUSP" collectorName="M. Alvarenga" country="Brazil" elevation="980" location="Encruzilhada" pageId="42" pageNumber="259" specimenCount="980" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="C18F3F7E2428FFAB75E76752F5FDF9EB" box="[132,225,1592,1616]" italics="true" pageId="42" pageNumber="259">
<typeStatus id="2C405DCE2428FFAB75E76752F5C0F9EB" box="[132,220,1592,1616]" pageId="42" pageNumber="259" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC2428FFAB759B6752F444F9EB" box="[248,344,1592,1616]" name="Brazil" pageId="42" pageNumber="259">BRAZIL</collectingCountry>
,
<location id="F624B5B72428FFAB740F6752F4EFF9EB" LSID="urn:lsid:plazi:treatment:7B52527A242BFFAB766E63FDF7B4F8EB:F624B5B72428FFAB740F6752F4EFF9EB" box="[364,499,1592,1616]" country="Brazil" name="Encruzilhada" pageId="42" pageNumber="259" stateProvince="Bahia">Encruzilhada</location>
<specimenCount id="E5FD28E52428FFAB77676752F744F9F4" box="[516,600,1592,1616]" count="980" pageId="42" pageNumber="259" type="generic">
<quantity id="34034E892428FFAB77676752F748F9F4" box="[516,596,1592,1616]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="42" pageNumber="259" unit="m" value="980.0">
<elevation id="78D6045F2428FFAB77676752F748F9F4" box="[516,596,1592,1616]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="42" pageNumber="259" unit="m" value="980.0">980 m</elevation>
</quantity>
.
</specimenCount>
<collectingRegion id="313F2D8E2428FFAB770B6752F7B9F9EB" box="[616,677,1592,1616]" country="Brazil" name="Bahia" pageId="42" pageNumber="259">Bahia</collectingRegion>
,
<date id="8745C5AC2428FFAB75E76732F5C1F9CB" box="[132,221,1624,1648]" pageId="42" pageNumber="259" value="1974-11">
<collectingDate id="97013C442428FFAB75E76732F5C1F9CB" box="[132,221,1624,1648]" pageId="42" pageNumber="259" value="1974-11">XI.1974</collectingDate>
</date>
,
<collectorName id="5E0E86BA2428FFAB758B6732F464F9CB" box="[232,376,1624,1648]" pageId="42" pageNumber="259">M. Alvarenga</collectorName>
], male (
<collectionCode id="95EA7BA92428FFAB74A46732F73AF9CB" box="[455,550,1624,1648]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="259">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C2428FFAB75E767F3F7B4F8EB" blockId="42.[132,681,1688,1872]" pageId="42" pageNumber="259">
<materialsCitation id="4393E9312428FFAB75E767F3F7B8F90B" box="[132,676,1688,1712]" collectionCode="MZUSP" pageId="42" pageNumber="259" specimenCount="9" typeStatus="paratype">
<emphasis id="C18F3F7E2428FFAB75E767F3F5FAF90B" box="[132,230,1689,1712]" italics="true" pageId="42" pageNumber="259">
<typeStatus id="2C405DCE2428FFAB75E767F3F5FCF90B" box="[132,224,1689,1712]" pageId="42" pageNumber="259" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same label as
<typeStatus id="2C405DCE2428FFAB74E467F2F4F8F90B" box="[391,484,1688,1712]" pageId="42" pageNumber="259" type="holotype">holotype</typeStatus>
,
<specimenCount id="E5FD28E52428FFAB749267F2F733F914" box="[497,559,1688,1712]" count="9" pageId="42" pageNumber="259" type="generic">9 exs.</specimenCount>
(
<collectionCode id="95EA7BA92428FFAB775C67F2F781F90B" box="[575,669,1688,1712]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="259">MZUSP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="4393E9312428FFAB75E767D2F43FF96B" box="[132,291,1720,1744]" collectionCode="CNC" isTailEnumeration="true" pageId="42" pageNumber="259" specimenCount="17" typeStatus="paratype">
<specimenCount id="E5FD28E52428FFAB75E767D2F5D2F974" box="[132,206,1720,1744]" count="17" pageId="42" pageNumber="259" type="generic">17 exs.</specimenCount>
(
<collectionCode id="95EA7BA92428FFAB75BD67D2F400F96B" box="[222,284,1720,1744]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="42" pageNumber="259">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4393E9312428FFAB745367D2F497F8AB" collectionCode="MZUSP" collectorName="Alvarenga &amp; Seabra" country="Brazil" elevation="960" location="Dpto. Zool." municipality="Encruzilhada" pageId="42" pageNumber="259" specimenCount="5" stateProvince="Bahia" typeStatus="paratype">
[
<location id="F624B5B72428FFAB745467D2F4A7F96B" LSID="urn:lsid:plazi:treatment:7B52527A242BFFAB766E63FDF7B4F8EB:F624B5B72428FFAB745467D2F4A7F96B" box="[311,443,1720,1744]" country="Brazil" municipality="Encruzilhada" name="Dpto. Zool." pageId="42" pageNumber="259" stateProvince="Bahia">Dpto. Zool.</location>
, UF-Paraná], [
<collectingMunicipality id="132079162428FFAB773767D2F5D9F94B" pageId="42" pageNumber="259">Encruzilhada</collectingMunicipality>
,
<collectingRegion id="313F2D8E2428FFAB75B367B2F411F94B" box="[208,269,1752,1776]" country="Brazil" name="Bahia" pageId="42" pageNumber="259">Bahia</collectingRegion>
,
<quantity id="34034E892428FFAB747B67B2F440F954" box="[280,348,1752,1776]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="42" pageNumber="259" unit="m" value="960.0">
<elevation id="78D6045F2428FFAB747B67B2F440F954" box="[280,348,1752,1776]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="42" pageNumber="259" unit="m" value="960.0">960 m</elevation>
</quantity>
<collectingCountry id="8BECA3FC2428FFAB741B67B2F4CFF94B" box="[376,467,1752,1776]" name="Brazil" pageId="42" pageNumber="259">BRAZIL</collectingCountry>
11/72,
<collectorName id="5E0E86BA2428FFAB774567B2F797F94B" box="[550,651,1752,1776]" pageId="42" pageNumber="259">Alvarenga</collectorName>
&amp;
<collectorName id="5E0E86BA2428FFAB75E76792F5D1F8AB" box="[132,205,1784,1808]" pageId="42" pageNumber="259">Seabra</collectorName>
],
<specimenCount id="E5FD28E52428FFAB75BD6792F405F8B4" box="[222,281,1784,1808]" count="5" pageId="42" pageNumber="259" type="generic">5 exs.</specimenCount>
(
<collectionCode id="95EA7BA92428FFAB74456792F498F8AB" box="[294,388,1784,1808]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="259">MZUSP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="4393E9312428FFAB74F46792F75EF8AB" box="[407,578,1784,1808]" collectionCode="DZUP" collectorName="Alvarenga &amp; Seabra" country="Brazil" elevation="960" isTailEnumeration="true" location="Dpto. Zool." municipality="Encruzilhada" pageId="42" pageNumber="259" specimenCount="11" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="E5FD28E52428FFAB74F46792F4C3F8B4" box="[407,479,1784,1808]" count="11" pageId="42" pageNumber="259" type="generic">11 exs.</specimenCount>
(
<collectionCode id="95EA7BA92428FFAB748E6792F726F8AB" box="[493,570,1784,1808]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="42" pageNumber="259" type="University or college">DZUP</collectionCode>
)
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;
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[
<collectorName id="5E0E86BA2428FFAB77316792F40BF88B" pageId="42" pageNumber="259">Coleção M. Alvarenga</collectorName>
], [
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<collectingDate id="97013C442428FFAB75E76652F5C1F8EB" box="[132,221,1848,1872]" pageId="42" pageNumber="259" value="1972-11">XI.1972</collectingDate>
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],
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(
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)
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; [Frei Gaspar
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,185 @@
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<mods:title id="55FB7F833164F76BC34D239E343DB043">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<mods:namePart id="313E91E016AFABF0DFD91CE3E754D82F">Rosa, Simone Policena</mods:namePart>
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<mods:date id="94D32A6614B3DF07C22B6B9B57F47AD7">2014</mods:date>
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<treatment id="7B52527A242CFFAE758B61DDF706F934" ID-DOI="http://doi.org/10.5281/zenodo.12641477" ID-Zenodo-Dep="12641477" LSID="urn:lsid:plazi:treatment:7B52527A242CFFAE758B61DDF706F934" httpUri="http://treatment.plazi.org/id/7B52527A242CFFAE758B61DDF706F934" lastPageId="47" lastPageNumber="264" pageId="46" pageNumber="263">
<subSubSection id="BBE1B0E7242CFFAF758B61DDF759FF6B" box="[232,581,183,208]" pageId="46" pageNumber="263" type="nomenclature">
<paragraph id="F344E36C242CFFAF758B61DDF759FF6B" blockId="46.[232,581,183,240]" box="[232,581,183,208]" pageId="46" pageNumber="263">
<heading id="A80C5400242CFFAF758B61DDF759FF6B" bold="true" box="[232,581,183,208]" centered="true" fontSize="10" level="2" pageId="46" pageNumber="263" reason="2">
<emphasis id="C18F3F7E242CFFAF758B61DDF759FF6B" bold="true" box="[232,581,183,208]" pageId="46" pageNumber="263">
<taxonomicName id="34FB98EF242CFFAF758B61DDF4F0FF74" authorityName="Rosa" authorityYear="2014" box="[232,492,183,207]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="263" phylum="Arthropoda" rank="species" species="flavifrons" status="sp. nov.">
<emphasis id="C18F3F7E242CFFAF758B61DDF4F0FF74" bold="true" box="[232,492,183,207]" italics="true" pageId="46" pageNumber="263">Physodactylus flavifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205242CFFAF749161D2F759FF6B" box="[498,581,184,208]" pageId="46" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242CFFAE745361B2F706F934" lastPageId="47" lastPageNumber="264" pageId="46" pageNumber="263" type="description">
<paragraph id="F344E36C242CFFAF745361B2F4E1FF4B" blockId="46.[232,581,183,240]" box="[304,509,216,240]" pageId="46" pageNumber="263">
<emphasis id="C18F3F7E242CFFAF745361B2F4E1FF4B" bold="true" box="[304,509,216,240]" pageId="46" pageNumber="263">
(
<figureCitation id="6BC0FFE9242CFFAF745B61B2F490FF4B" box="[312,396,216,240]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="46" pageNumber="263">Figs. 20</figureCitation>
A-G, 35F)
</emphasis>
</paragraph>
<paragraph id="F344E36C242CFFAF75E76072F480FEEB" blockId="46.[132,680,280,336]" pageId="46" pageNumber="263">
<emphasis id="C18F3F7E242CFFAF75E76072F5ECFE8B" box="[132,240,280,304]" italics="true" pageId="46" pageNumber="263">Etymology:</emphasis>
From Latin,
<emphasis id="C18F3F7E242CFFAF74146072F4B6FE8B" box="[375,426,280,304]" italics="true" pageId="46" pageNumber="263">flavis</emphasis>
= yellow; alluding to the orange head of this species.
</paragraph>
<paragraph id="F344E36C242CFFAE75E76013F731FCB4" blockId="46.[132,681,376,656]" lastBlockId="47.[94,643,184,784]" lastPageId="47" lastPageNumber="264" pageId="46" pageNumber="263">
<emphasis id="C18F3F7E242CFFAF75E76013F5E8FE2B" box="[132,244,377,400]" italics="true" pageId="46" pageNumber="263">Description</emphasis>
(male,
<figureCitation id="6BC0FFE9242CFFAF74226012F487FE2B" box="[321,411,376,400]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="46" pageNumber="263">Fig. 35F</figureCitation>
): Integument bright; head entirely orange except for the black tips of mandible; antennomeres I-III, prothorax, ventral pterothorax and scutellum orange; leg orange with tibia dark brown; antennomeres IV-XI and ventral abdomen dark brown to black, elytra black with an orange spot a little narrower than elytra on anterior 1/5-1/3; pilosity yellow. Total length 10.0-
<quantity id="34034E89242CFFAF74AA6332F730FDD4" box="[457,556,600,624]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="46" pageNumber="263" unit="mm" value="12.5">12.5 mm</quantity>
; elytral base 0.9-1.0 as wide as prothorax, elytra 3.2-3.3x longer than pronotum. Frons flattened, frontal carina not produced; antenna (
<figureCitation id="6BC0FFE9242CFFAF76C861B2F111FF4B" box="[939,1037,216,240]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="46" pageNumber="263">Fig. 20A</figureCitation>
) reaching the posterior third of pronotum; antennomere IV 1.22x wider than long, V-VI 1.56 and 1.62x wider than long, VII-VIII 1.44x wider than long, IX-X 1.30-1.33x wider than long, XI oval 1.54x longer than wide. Maxilla (
<figureCitation id="6BC0FFE9242CFFAF76496012F699FE2B" box="[810,901,376,400]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="46" pageNumber="263">Fig. 20B</figureCitation>
) with galea covered with a few fine and spiniform setae. Pronotum (
<figureCitation id="6BC0FFE9242CFFAF715760F2F18BFE0B" box="[1076,1175,408,432]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="46" pageNumber="263">Fig. 20C</figureCitation>
) convex 1.40-1.46x wider than long with posterior 1/3-1/2 parallel, anterior part straight convergent to anterior margin; anteromedian margin produced anteriad; posterior angles short, convex, parallel; lateral carina complete strongly prominent on posterior 2/3, sinuous in lateral view (
<figureCitation id="6BC0FFE9242CFFAF76D16332F10BFDCB" box="[946,1047,600,624]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="46" pageNumber="263">Fig. 20D</figureCitation>
), nearly entirely visible dorsally; disc with punctures heterogeneously or homogenously distributed 0.5-1.0 diameters apart, smaller and denser on anterior margin, denser on lateral border; prosternal process (
<figureCitation id="6BC0FFE9242DFFAE74A26192F73DFEAB" box="[449,545,248,272]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="47" pageNumber="264">Fig. 20D</figureCitation>
) without subapical tooth. Lamellae of pro- and mesotarsomeres I as large as those of tarsomeres II-III, small on metatarsomere I. Elytra parallel-sided on anterior 2/3 then tapering to apex; striae deeply impressed, with three rows of punctures, punctures glabrous on the median row, smaller and bearing a small seta on the lateral row; interstices convex, smooth and glabrous; apical striae with punctures smaller than those of pronotal lateral border and as large as those of elytral base. Abdomen with ventrites 2-4 concave, ventrite 1 evenly pilose with setae 1/3 as long as those of ventrite 2, ventrites 2-5 covered with very long semi-erect setae. Pregenitalic segments and aedeagus covered with stout yellow setae. Sternite VIII with posterior margin nearly straight, anterior sclerotization 0.55x the total width of sternite; sternite IX tapered to apex.
</paragraph>
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<paragraph id="F344E36C242CFFAF75E56648F134F8F4" blockId="46.[132,1267,1826,1871]" pageId="46" pageNumber="263">
<emphasis id="C18F3F7E242CFFAF75E56648F5EAF88E" bold="true" box="[134,246,1826,1845]" pageId="46" pageNumber="263">FIGURE 20:</emphasis>
<taxonomicName id="34FB98EF242CFFAF759F6648F4AAF88E" authorityName="Rosa" authorityYear="2014" box="[252,438,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="263" phylum="Arthropoda" rank="species" species="flavifrons" status="sp. nov.">
<emphasis id="C18F3F7E242CFFAF759F6648F4AAF88E" box="[252,438,1826,1845]" italics="true" pageId="46" pageNumber="263">Physodactylus flavifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205242CFFAF74DF6649F4E1F88D" box="[444,509,1827,1846]" pageId="46" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E242CFFAF77616648F708F88E" bold="true" box="[514,532,1826,1845]" pageId="46" pageNumber="263">A,</emphasis>
antenna;
<emphasis id="C18F3F7E242CFFAF770B6648F767F88E" bold="true" box="[616,635,1826,1845]" pageId="46" pageNumber="263">B,</emphasis>
maxilla;
<emphasis id="C18F3F7E242CFFAF77AB6648F60EF88E" bold="true" box="[712,786,1826,1845]" pageId="46" pageNumber="263">C, D, E,</emphasis>
prothorax (dorsal, lateral ventral);
<emphasis id="C18F3F7E242CFFAF71556648F17CF88E" bold="true" box="[1078,1120,1826,1845]" pageId="46" pageNumber="263">F, G,</emphasis>
aedeagus (dorsal, ventral).
<emphasis id="C18F3F7E242CFFAF75AD6656F4AFF8F4" box="[206,435,1852,1871]" italics="true" pageId="46" pageNumber="263">
<taxonomicName id="34FB98EF242CFFAF75AD6656F4B3F8F4" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[206,431,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="263" phylum="Arthropoda" rank="species" species="foveatostriatus">Physodactylus foveatostriatus</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E242CFFAF74D46656F4D2F8F4" bold="true" box="[439,462,1852,1871]" pageId="46" pageNumber="263">H,</emphasis>
antenna;
<emphasis id="C18F3F7E242CFFAF777C6656F75CF8F4" bold="true" box="[543,576,1852,1871]" pageId="46" pageNumber="263">I, J,</emphasis>
prothorax (dorsal, lateral);
<emphasis id="C18F3F7E242CFFAF76426656F651F8F4" bold="true" box="[801,845,1852,1871]" pageId="46" pageNumber="263">K, L,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C242DFFAE753D6252F48CFBAB" blockId="47.[94,643,824,1040]" pageId="47" pageNumber="264">
<emphasis id="C18F3F7E242DFFAE753D6252F5ABFCEB" box="[94,183,824,848]" italics="true" pageId="47" pageNumber="264">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9242DFFAE75A96252F448FCEB" box="[202,340,824,848]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="47" pageNumber="264">Figs. 20F, G</figureCitation>
): Phallobase 0.44x the total length of aedeagus, 1.2x longer than wide, length ratio between lateroposterior and median parts 4.8; paramere with ventral surface nearly entirely sclerotized; penis with basal strut 0.24x its total length, articulated to the parameres by a strongly sclerotized process; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C242DFFAE753D6552F76BFBCB" blockId="47.[94,643,1080,1136]" pageId="47" pageNumber="264">
<materialsCitation id="4393E931242DFFAE753D6552F76BFBCB" collectingDate="1968-02-01" collectingDateMax="1968-02-11" collectingDateMin="1968-02-01" collectionCode="MZUSP" collectorName="Santarem, PA" pageId="47" pageNumber="264" specimenCount="1" typeStatus="holotype">
<emphasis id="C18F3F7E242DFFAE753D6552F5A7FBEB" box="[94,187,1080,1104]" italics="true" pageId="47" pageNumber="264">
<typeStatus id="2C405DCE242DFFAE753D6552F5AAFBEB" box="[94,182,1080,1104]" pageId="47" pageNumber="264" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Fazenda Taperinha, prox.
<collectorName id="5E0E86BA242DFFAE74896552F763FBEB" box="[490,639,1080,1104]" pageId="47" pageNumber="264">Santarém, PA</collectorName>
,
<date id="8745C5AC242DFFAE753D6532F5EFFBCB" box="[94,243,1112,1136]" pageId="47" pageNumber="264" value="1968-02-01" valueMax="1968-02-11" valueMin="1968-02-01">
<collectingDate id="97013C44242DFFAE753D6532F5EFFBCB" box="[94,243,1112,1136]" pageId="47" pageNumber="264" value="1968-02-01" valueMax="1968-02-11" valueMin="1968-02-01">01-11.II.1968</collectingDate>
</date>
, Exp. Perm. Amaz.] male, (
<collectionCode id="95EA7BA9242DFFAE776E6532F770FBCB" box="[525,620,1112,1136]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="47" pageNumber="264">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C242DFFAE753D65F3F772FB0B" blockId="47.[94,622,1176,1200]" box="[94,622,1176,1200]" pageId="47" pageNumber="264">
<materialsCitation id="4393E931242DFFAE753D65F3F775FB0B" box="[94,617,1176,1200]" collectionCode="MZUSP" pageId="47" pageNumber="264" specimenCount="2" typeStatus="paratype">
<emphasis id="C18F3F7E242DFFAE753D65F3F5DCFB0B" box="[94,192,1177,1200]" italics="true" pageId="47" pageNumber="264">
<typeStatus id="2C405DCE242DFFAE753D65F3F5A6FB0B" box="[94,186,1177,1200]" pageId="47" pageNumber="264" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same label as
<typeStatus id="2C405DCE242DFFAE743665F2F4AEFB0B" box="[341,434,1176,1200]" pageId="47" pageNumber="264" type="holotype">holotype</typeStatus>
,
<specimenCount id="E5FD28E5242DFFAE74DF65F2F4EBFB14" box="[444,503,1176,1200]" count="2" pageId="47" pageNumber="264" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA9242DFFAE776765F2F77FFB0B" box="[516,611,1176,1200]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="47" pageNumber="264">MZUSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C242DFFAE753D65B2F4EEFB4B" blockId="47.[94,498,1240,1264]" box="[94,498,1240,1264]" pageId="47" pageNumber="264">
<emphasis id="C18F3F7E242DFFAE753D65B2F5FDFB4B" box="[94,225,1240,1264]" italics="true" pageId="47" pageNumber="264">Distribution:</emphasis>
<collectingCountry id="8BECA3FC242DFFAE758465B2F45AFB4B" box="[231,326,1240,1264]" name="Brazil" pageId="47" pageNumber="264">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E242DFFAE742C65B2F49FFB54" box="[335,387,1240,1263]" italics="true" pageId="47" pageNumber="264">
<collectingRegion id="313F2D8E242DFFAE742C65B2F462FB54" box="[335,382,1240,1263]" country="Brazil" name="Para" pageId="47" pageNumber="264">Pará</collectingRegion>
:
</emphasis>
Santarém.
</paragraph>
<paragraph id="F344E36C242DFFAE753D6472F706F934" blockId="47.[94,643,1304,1680]" pageId="47" pageNumber="264">
<emphasis id="C18F3F7E242DFFAE753D6472F4B0FA8B" box="[94,428,1304,1328]" italics="true" pageId="47" pageNumber="264">
Remarks:
<taxonomicName id="34FB98EF242DFFAE75A06472F4B0FA8B" authorityName="Rosa" authorityYear="2014" box="[195,428,1304,1328]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="264" phylum="Arthropoda" rank="species" species="flavifrons" status="sp. nov.">Physodactylus flavifrons</taxonomicName>
</emphasis>
<taxonomicNameLabel id="DABC8205242DFFAE74D56472F715FA94" box="[438,521,1304,1327]" pageId="47" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
is diagnosable by the unique combination of characters: pilosity yellow, antenna reaching the anterior third of pronotum, prothorax 1.40-1.46x wider than long, lamellae of pro- and mesotarsomeres I as large as the other ones, elytra with striae deeply impressed and as wide as the interstices, abdominal ventrites 2-4 concave with very long setae. This species is most similar to
<emphasis id="C18F3F7E242DFFAE753D6773F43AF98B" box="[94,294,1560,1584]" italics="true" pageId="47" pageNumber="264">
<taxonomicName id="34FB98EF242DFFAE753D6773F5DAF98B" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[94,198,1560,1584]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="264" phylum="Arthropoda" rank="species" species="sulcatus">P. sulcatus</taxonomicName>
,
<taxonomicName id="34FB98EF242DFFAE75B26773F43AF98B" authorityName="Rosa" authorityYear="2014" box="[209,294,1561,1584]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="264" phylum="Arthropoda" rank="species" species="patens" status="sp. nov.">P. patens</taxonomicName>
</emphasis>
<taxonomicNameLabel id="DABC8205242DFFAE744D6772F462F994" box="[302,382,1560,1583]" pageId="47" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="34FB98EF242DFFAE74D06773F730F98B" authorityName="Rosa" authorityYear="2014" box="[435,556,1560,1584]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="264" phylum="Arthropoda" rank="species" species="latithorax" status="sp. nov.">
<emphasis id="C18F3F7E242DFFAE74D06773F730F98B" box="[435,556,1560,1584]" italics="true" pageId="47" pageNumber="264">P. latithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205242DFFAE77576772F79FF994" box="[564,643,1560,1583]" pageId="47" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
in shape of prothorax, antenna and elytral striae; however it can be promptly distinguished by its longer abdominal pilosity and ventrites 2-4 concave.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,201 @@
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<paragraph id="F344E36C242DFFAE75F467BDF756F94B" blockId="47.[151,586,1751,1808]" box="[151,586,1751,1776]" pageId="47" pageNumber="264">
<heading id="A80C5400242DFFAE75F467BDF756F94B" bold="true" box="[151,586,1751,1776]" centered="true" fontSize="10" level="2" pageId="47" pageNumber="264" reason="2">
<taxonomicName id="34FB98EF242DFFAE75F467BDF756F94B" authority="Chassain, 2005" authorityName="Chassain" authorityYear="2005" box="[151,586,1751,1776]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="264" phylum="Arthropoda" rank="species" species="fleutiauxi">
<emphasis id="C18F3F7E242DFFAE75F467BDF756F94B" bold="true" box="[151,586,1751,1776]" pageId="47" pageNumber="264">
<emphasis id="C18F3F7E242DFFAE75F467BDF481F954" bold="true" box="[151,413,1751,1775]" italics="true" pageId="47" pageNumber="264">Physodactylus fleutiauxi</emphasis>
<bibRefCitation id="976A9E9D242DFFAE74C067B2F756F94B" author="CHASSAIN, J." box="[419,586,1752,1776]" pageId="47" pageNumber="264" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242DFFAE745B6792F4B5F8AB" box="[312,425,1784,1808]" pageId="47" pageNumber="264" type="description">
<paragraph id="F344E36C242DFFAE745B6792F4B5F8AB" blockId="47.[151,586,1751,1808]" box="[312,425,1784,1808]" pageId="47" pageNumber="264">
<emphasis id="C18F3F7E242DFFAE745B6792F4B5F8AB" bold="true" box="[312,425,1784,1808]" pageId="47" pageNumber="264">
(
<figureCitation id="6BC0FFE9242DFFAE745C6792F4BCF8AB" box="[319,416,1784,1808]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="47" pageNumber="264">Fig. 35G</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242DFFAE753D6652F187F9AB" pageId="47" pageNumber="264" type="reference_group">
<paragraph id="F344E36C242DFFAE753D6652F6DCFAB4" blockId="47.[94,540,1848,1872]" lastBlockId="47.[680,1229,184,1296]" pageId="47" pageNumber="264">
<treatmentCitationGroup id="D3EBC442242DFFAE753D6652F6DCFAB4" pageId="47" pageNumber="264">
<treatmentCitation id="725AC57D242DFFAE753D6652F705F8EB" author="CHASSAIN, J." box="[94,537,1848,1872]" page="68" pageId="47" pageNumber="264" year="2005">
<taxonomicName id="34FB98EF242DFFAE753D6652F705F8EB" authority="Chassain, 2005: 68" authorityName="Chassain" authorityPageNumber="68" authorityYear="2005" box="[94,537,1848,1872]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="264" phylum="Arthropoda" rank="species" species="fleutiauxi">
<emphasis id="C18F3F7E242DFFAE753D6652F455F8EB" box="[94,329,1848,1872]" italics="true" pageId="47" pageNumber="264">Physodactylus fleutiauxi</emphasis>
<bibRefCitation id="976A9E9D242DFFAE742C6652F705F8EB" author="CHASSAIN, J." box="[335,537,1848,1872]" pageId="47" pageNumber="264" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005: 68</bibRefCitation>
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.
<emphasis id="C18F3F7E242DFFAE77CB61D2F636FF6B" box="[680,810,184,208]" italics="true" pageId="47" pageNumber="264">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9242DFFAE76CE61D2F13BFF6B" box="[941,1063,184,208]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="47" pageNumber="264">Fig. 35G</figureCitation>
): Integument bright, evenly black; pilosity yellow. Total length
<quantity id="34034E89242DFFAE77CB6192F65FFEB4" box="[680,835,248,272]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.55" metricValueMin="1.35" pageId="47" pageNumber="264" unit="mm" value="14.5" valueMax="15.5" valueMin="13.5">13.5-15.5 mm</quantity>
; elytral base 1.0-1.1x wider than prothorax, elytra 2.7-2.8x longer than pronotum. Frons concave to convex, frontal carina not or slightly produced; IEP 0.21-0.25; antenna reaching the anterior third of pronotum; antennomere IV 1.2x wider than long, V-X 1.8x wider than long, XI oval 1.2x longer than wide. Maxilla with galea covered with fine and spiniform setae. Pronotum convex 1.3x wider than long, parallel on posterior third then straight or roundly convergent to anterior margin; anteromedian margin slightly produced anteriad; posterior angles short, convex, parallel; lateral carina evenly prominent to anterior margin; dorsally visible only on posterior third; disc with punctures medium size homogenously distributed 0.5-1.0 diameters apart, smaller and denser on anterior margin, denser on lateral border; prosternal process without subapical tooth. Lamellae of pro- and mesotarsomeres I very small, absent on metatarsomere I. Elytra parallel-sided on anterior 2/3 then tapering to apex; apex usually with a very small sutural spine; striae deeply impressed, with three rows of punctures, punctures glabrous on the median row, smaller and bearing a small seta on the lateral row; interstices convex, smooth and glabrous; apical striae with punctures smaller than those of pronotal lateral border and as large as those of elytral base. Abdomen with ventrites 1-4 strongly and evenly convex, ventrite 1 evenly pilose with short decumbent setae, 2-4 with long semi-erect setae. Pregenitalic segments and aedeagus covered with stout yellow setae. Sternite VIII subrectangular, posterior margin straight, anterior sclerotization 0.5x the total width of sternite; sternite IX tapered to apex.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F344E36C242DFFAE77CB6452F187F9AB" blockId="47.[680,1229,1336,1552]" pageId="47" pageNumber="264">
<emphasis id="C18F3F7E242DFFAE77CB6452F614FAEB" box="[680,776,1336,1360]" italics="true" pageId="47" pageNumber="264">Aedeagus:</emphasis>
Phallobase 0.5x the total length of aedeagus, 1.6x longer than wide, length ratio between lateroposterior and median parts 4.5-6.1; ventral surface of paramere with the lateral half and anterior margin sclerotized; penis with basal strut 0.2-0.3x its total length, articulated to the parameres by a strongly sclerotized process; ventral sclerite parallel-sided.
</paragraph>
</subSubSection>
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<paragraph id="F344E36C242DFFAE77CB6752F6C1F96B" blockId="47.[680,1228,1592,1744]" pageId="47" pageNumber="264">
<materialsCitation id="4393E931242DFFAE77CB6752F6C1F96B" collectingDate="2000-12-15" collectionCode="MNHN" collectorName="Chapada dos Parecis &amp; A. Foucart" country="Brazil" determinerName="J. Chassain" latitude="-14.2641115" location="d'Uirapuru" longLatPrecision="1" longitude="-59.233902" pageId="47" pageNumber="264" specimenCount="3" stateProvince="Mato Grosso" typeStatus="holotype">
<emphasis id="C18F3F7E242DFFAE77CB6752F619F9EB" box="[680,773,1592,1616]" italics="true" pageId="47" pageNumber="264">
<typeStatus id="2C405DCE242DFFAE77CB6752F61CF9EB" box="[680,768,1592,1616]" pageId="47" pageNumber="264" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC242DFFAE767D6752F641F9EB" box="[798,861,1592,1616]" name="Brazil" pageId="47" pageNumber="264">Brésil</collectingCountry>
, (
<collectingRegion id="313F2D8E242DFFAE76196752F10CF9EB" box="[890,1040,1592,1616]" country="Brazil" name="Mato Grosso" pageId="47" pageNumber="264">Mato Grosso</collectingRegion>
),
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,
<geoCoordinate id="96CF85AB242DFFAE77CB6732F650F9CB" box="[680,844,1624,1648]" degrees="59" direction="west" minutes="14" orientation="longitude" pageId="47" pageNumber="264" precision="1" seconds="02.05" value="-59.233902">59°1402.05”W</geoCoordinate>
,
<collectorName id="5E0E86BA242DFFAE76036732F154F9CB" box="[864,1096,1624,1648]" pageId="47" pageNumber="264">Chapada dos Parecis</collectorName>
,
<quantity id="34034E89242DFFAE71386732F1B4F9CB" box="[1115,1192,1624,1648]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="264" unit="km" value="30.0">30 km</quantity>
N
<location id="F624B5B7242DFFAE77CB6712F63DF92B" LSID="urn:lsid:plazi:treatment:7B52527A242DFFB175F467BDF4A1FDEB:F624B5B7242DFFAE77CB6712F63DF92B" box="[680,801,1656,1680]" country="Brazil" latitude="-14.2641115" longLatPrecision="1" longitude="-59.233902" name="d'Uirapuru" pageId="47" pageNumber="264" stateProvince="Mato Grosso">dUirapuru</location>
], [01/
<date id="8745C5AC242DFFAE76066712F6FFF92B" box="[869,995,1656,1680]" pageId="47" pageNumber="264" value="2000-12-15">
<collectingDate id="97013C44242DFFAE76066712F6FFF92B" box="[869,995,1656,1680]" pageId="47" pageNumber="264" value="2000-12-15">15-12-2000</collectingDate>
</date>
,
<collectorName id="5E0E86BA242DFFAE76966712F173F934" box="[1013,1135,1656,1680]" pageId="47" pageNumber="264">A. Foucart</collectorName>
]. [
<typeStatus id="2C405DCE242DFFAE71EF6712F7C5F90B" pageId="47" pageNumber="264" type="holotype">Holotype</typeStatus>
], [
<taxonomicName id="34FB98EF242DFFAE779067F2F6E9F90B" authorityName="Chassain" authorityYear="2005" box="[755,1013,1688,1712]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="264" phylum="Arthropoda" rank="species" species="fleutiauxi">Physodactylus fleutiauxi</taxonomicName>
,
<typeStatus id="2C405DCE242DFFAE716067F2F17BF90B" box="[1027,1127,1688,1712]" pageId="47" pageNumber="264" type="holotype">Holotype</typeStatus>
,
<determinerName id="06980FA8242DFFAE711567F2F7CCF96B" pageId="47" pageNumber="264">J. Chassain</determinerName>
det. 04], male (
<collectionCode id="95EA7BA9242DFFAE761667D2F6CDF96B" box="[885,977,1720,1744]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="47" pageNumber="264">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C242DFFAE77CB6793F1D1F8EB" blockId="47.[680,1229,1784,1872]" pageId="47" pageNumber="264">
<materialsCitation id="4393E931242DFFAE77CB6793F1D1F8EB" collectingDate="2002-12-15" collectorName="Chapada dos Parecis &amp; Uirapuru &amp; Usine" country="Brazil" latitude="-14.2641115" location="Mato grosso" longLatPrecision="1" longitude="-59.233902" pageId="47" pageNumber="264" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paratype">
<emphasis id="C18F3F7E242DFFAE77CB6793F61FF8AB" box="[680,771,1785,1808]" italics="true" pageId="47" pageNumber="264">
<typeStatus id="2C405DCE242DFFAE77CB6793F7E1F8AB" box="[680,765,1785,1808]" pageId="47" pageNumber="264" type="paratype">Paratype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC242DFFAE76416792F67DF8AB" box="[802,865,1784,1808]" name="Brazil" pageId="47" pageNumber="264">Brèsil</collectingCountry>
,
<collectingRegion id="313F2D8E242DFFAE761E6792F10EF8AB" box="[893,1042,1784,1808]" country="Brazil" name="Mato Grosso" pageId="47" pageNumber="264">Mato grosso</collectingRegion>
,
<geoCoordinate id="96CF85AB242DFFAE714D6792F1D5F8AB" box="[1070,1225,1784,1808]" degrees="14" direction="south" minutes="15" orientation="latitude" pageId="47" pageNumber="264" precision="1" seconds="50.80" value="-14.2641115">14°1550.80”S</geoCoordinate>
,
<geoCoordinate id="96CF85AB242DFFAE77CB6672F650F88B" box="[680,844,1816,1840]" degrees="59" direction="west" minutes="14" orientation="longitude" pageId="47" pageNumber="264" precision="1" seconds="02.05" value="-59.233902">59°1402.05”W</geoCoordinate>
,
<collectorName id="5E0E86BA242DFFAE763E6672F15DF88B" box="[861,1089,1816,1840]" pageId="47" pageNumber="264">Chapada dos Parecis</collectorName>
], [
<quantity id="34034E89242DFFAE713C6672F1B7F88B" box="[1119,1195,1816,1840]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="264" unit="km" value="30.0">30 km</quantity>
N
<collectorName id="5E0E86BA242DFFAE77CF6652F60DF8EB" box="[684,785,1848,1872]" pageId="47" pageNumber="264">Uirapuru</collectorName>
,
<collectorName id="5E0E86BA242DFFAE76416652F641F8EB" box="[802,861,1848,1872]" pageId="47" pageNumber="264">Usine</collectorName>
alcomat, 01 au
<date id="8745C5AC242DFFAE717E6652F1B8F8EB" box="[1053,1188,1848,1872]" pageId="47" pageNumber="264" value="2002-12-15">
<collectingDate id="97013C44242DFFAE717E6652F1B8F8EB" box="[1053,1188,1848,1872]" pageId="47" pageNumber="264" value="2002-12-15">15-XII-2002</collectingDate>
</date>
, A.
</materialsCitation>
</paragraph>
<paragraph id="F344E36C2432FFB175E761D2F74EFF4B" blockId="48.[132,680,184,240]" pageId="48" pageNumber="265">
<materialsCitation id="4393E9312432FFB175E761D2F752FF4B" collectionCode="MZUSP" collectorName="Foucart" determinerName="J. Chassain" pageId="48" pageNumber="265" specimenCount="1" typeStatus="paratype">
<collectorName id="5E0E86BA2432FFB175E761D2F5CFFF74" box="[132,211,184,207]" pageId="48" pageNumber="265">Foucart</collectorName>
leg], [
<typeStatus id="2C405DCE2432FFB1744661D2F499FF6B" box="[293,389,184,208]" pageId="48" pageNumber="265" type="paratype">Paratype</typeStatus>
], [
<taxonomicName id="34FB98EF2432FFB174C761D2F7B4FF6B" authorityName="Rosa" authorityYear="2014" box="[420,680,184,208]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="fleutiauxi" status="sp. nov.">Physodactylus fleutiauxi</taxonomicName>
<taxonomicNameLabel id="DABC82052432FFB175E761B2F5CFFF54" box="[132,211,216,239]" pageId="48" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
<determinerName id="06980FA82432FFB175BA61B2F457FF4B" box="[217,331,216,240]" pageId="48" pageNumber="265">J. Chassain</determinerName>
det 04],
<specimenCount id="E5FD28E52432FFB174CA61B2F4C7FF54" box="[425,475,216,240]" count="1" pageId="48" pageNumber="265" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA92432FFB1748B61B2F75BFF4B" box="[488,583,216,240]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="48" pageNumber="265">MZUSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C2432FFB175E76072F5CEFEEB" blockId="48.[132,680,280,336]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB175E76072F41AFE8B" box="[132,262,280,304]" italics="true" pageId="48" pageNumber="265">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2432FFB174776072F46FFE8B" box="[276,371,280,304]" name="Brazil" pageId="48" pageNumber="265">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2432FFB174E76073F711FE94" box="[388,525,280,304]" italics="true" pageId="48" pageNumber="265">
<collectingRegion id="313F2D8E2432FFB174E76073F714FE94" box="[388,520,280,304]" country="Brazil" name="Mato Grosso" pageId="48" pageNumber="265">Mato Grosso</collectingRegion>
:
</emphasis>
Chapada dos Parecis.
</paragraph>
<paragraph id="F344E36C2432FFB175E76012F4A1FDEB" blockId="48.[132,681,376,592]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB175E76012F4CDFE2B" box="[132,465,376,400]" italics="true" pageId="48" pageNumber="265">
Remarks:
<taxonomicName id="34FB98EF2432FFB175856012F4CDFE2B" authorityName="Rosa" authorityYear="2014" box="[230,465,376,400]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="fleutiauxi">Physodactylus fleutiauxi</taxonomicName>
</emphasis>
has aedeagus similar to that of
<emphasis id="C18F3F7E2432FFB1758960F3F44FFE0B" box="[234,339,408,432]" italics="true" pageId="48" pageNumber="265">
<taxonomicName id="34FB98EF2432FFB1758960F3F44CFE0B" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[234,336,408,432]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="sulcatus">P. sulcatus</taxonomicName>
,
</emphasis>
differing only in its slender phallobase.
<taxonomicName id="34FB98EF2432FFB175B060D2F4DCFE6B" authorityName="Rosa" authorityYear="2014" box="[211,448,440,464]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="fleutiauxi">
<emphasis id="C18F3F7E2432FFB175B060D2F4DCFE6B" box="[211,448,440,464]" italics="true" pageId="48" pageNumber="265">Physodactylus fleutiauxi</emphasis>
</taxonomicName>
differs also from that species in its body evenly black, pronotum weakly convex with lateral carina complete and pronotal punctation distinctly different from the finer and sparser hypomeral punctation.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,195 @@
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<heading id="A80C5400242EFFAD767863DDF1B9FD6B" bold="true" box="[795,1189,695,720]" centered="true" fontSize="10" level="2" pageId="44" pageNumber="261" reason="2">
<taxonomicName id="34FB98EF242EFFAD767863DDF1B9FD6B" authority="Chassain, 2005" authorityName="Chassain" authorityYear="2005" box="[795,1189,695,720]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="costae">
<emphasis id="C18F3F7E242EFFAD767863DDF1B9FD6B" bold="true" box="[795,1189,695,720]" pageId="44" pageNumber="261">
<emphasis id="C18F3F7E242EFFAD767863DDF6E4FD74" bold="true" box="[795,1016,695,719]" italics="true" pageId="44" pageNumber="261">Physodactylus costae</emphasis>
<bibRefCitation id="976A9E9D242EFFAD769D63D2F1B9FD6B" author="CHASSAIN, J." box="[1022,1189,696,720]" pageId="44" pageNumber="261" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242EFFAD76C463B2F105FD4B" box="[935,1049,728,752]" pageId="44" pageNumber="261" type="description">
<paragraph id="F344E36C242EFFAD76C463B2F105FD4B" blockId="44.[795,1189,695,752]" box="[935,1049,728,752]" pageId="44" pageNumber="261">
<emphasis id="C18F3F7E242EFFAD76C463B2F105FD4B" bold="true" box="[935,1049,728,752]" pageId="44" pageNumber="261">
(
<figureCitation id="6BC0FFE9242EFFAD76CD63B2F10EFD4B" box="[942,1042,728,752]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="44" pageNumber="261">Fig. 35D</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242EFFAD77AD6272F179FC8B" box="[718,1125,792,816]" pageId="44" pageNumber="261" type="reference_group">
<paragraph id="F344E36C242EFFAD77AD6272F179FC8B" blockId="44.[718,1125,792,816]" box="[718,1125,792,816]" pageId="44" pageNumber="261">
<treatmentCitationGroup id="D3EBC442242EFFAD77AD6272F179FC8B" box="[718,1125,792,816]" pageId="44" pageNumber="261">
<treatmentCitation id="725AC57D242EFFAD77AD6272F17DFC8B" author="CHASSAIN, J." box="[718,1121,792,816]" page="69" pageId="44" pageNumber="261" year="2005">
<taxonomicName id="34FB98EF242EFFAD77AD6272F17DFC8B" authority="Chassain, 2005: 69" authorityName="Chassain" authorityPageNumber="69" authorityYear="2005" box="[718,1121,792,816]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="261" phylum="Arthropoda" rank="species" species="costae">
<emphasis id="C18F3F7E242EFFAD77AD6272F68CFC8B" box="[718,912,792,816]" italics="true" pageId="44" pageNumber="261">Physodactylus costae</emphasis>
<bibRefCitation id="976A9E9D242EFFAD76F46272F17DFC8B" author="CHASSAIN, J." box="[919,1121,792,816]" pageId="44" pageNumber="261" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005: 69</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242EFFAC77AD6232F406FA34" lastPageId="45" lastPageNumber="262" pageId="44" pageNumber="261" type="description">
<paragraph id="F344E36C242EFFAC77AD6232F44BFF54" blockId="44.[718,1266,856,1872]" lastBlockId="45.[94,642,184,240]" lastPageId="45" lastPageNumber="262" pageId="44" pageNumber="261">
<emphasis id="C18F3F7E242EFFAD77AD6232F64FFCCB" box="[718,851,856,880]" italics="true" pageId="44" pageNumber="261">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9242EFFAD76D06232F101FCCB" box="[947,1053,856,880]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="44" pageNumber="261">Fig. 35D</figureCitation>
): Integument dark brown with pronotum, sutural elytral interstice and borders of elytra lighter and legs brown; pilosity yellow. Total length 12.0 mm; elytral base 1.0-1.1x wider than prothorax, elytra 3.3x longer than pronotum. Frons concave, frontal carina not or slightly produced; antenna reaching the anterior third of pronotum. IEP 0.30. Antennomere IV 1.4x wider than long, V-X 1.7x wider than long, XI 1.1x as long as wide. Maxilla with galea covered with fine and spiniform setae. Pronotum convex 1.3x wider than long, parallel on posterior third then straight convergent to anterior margin; anteromedian margin slightly produced anteriad; posterior angles short, convex, parallel; lateral carina evenly prominent to anterior margin; dorsally visible only on posterior third; disc with punctures medium size homogenously distributed 2.0-3.0 diameters apart, smaller and denser on lateral and posterior borders; prosternal process without subapical tooth. Lamellae of pro- and mesotarsomere I very small, absent on metatarsomere I. Elytra parallel-sided on anterior half then tapering to apex; apex conjointly rounded; striae deeply impressed, with three rows of punctures, punctures glabrous on the median row, smaller and bearing a small seta on the lateral row; interstices weakly convex on posterior half, flattened on anterior part, smooth and glabrous; apical striae with punctures smaller than those of pronotal lateral border and as large as those of elytral base. Ventral abdomen evenly convex, ventrites 1-5 evenly pilose with short decumbent setae. Pregenitalic segments and aedeagus covered with stout yellow setae. Sternite IX tapered to apex.
</paragraph>
<paragraph id="F344E36C242FFFAC753D6072F5F2FE4B" blockId="45.[94,643,280,496]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC753D6072F5A2FE8B" box="[94,190,280,304]" italics="true" pageId="45" pageNumber="262">Aedeagus:</emphasis>
Phallobase 0.5x total length of aedeagus, 1.5x longer than wide, length ratio between lateroposterior and median parts 5.5; ventral surface of paramere with the lateral half sclerotized; penis with basal strut 0.3x its total length, articulated to the parameres by a strongly sclerotized process; ventral sclerite parallel-sided.
</paragraph>
<paragraph id="F344E36C242FFFAC753D6372F48FFD0B" blockId="45.[94,643,536,688]" pageId="45" pageNumber="262">
<materialsCitation id="4393E931242FFFAC753D6372F48FFD0B" collectingDate="2000-12-15" collectionCode="MNHN" collectorName="Chapada dos Parecis &amp; A. Foucart" country="Brazil" determinerName="J. Chassain" latitude="-14.2641115" location="d'Uirapuru" longLatPrecision="1" longitude="-59.217236" pageId="45" pageNumber="262" specimenCount="3" stateProvince="Mato Grosso" typeStatus="holotype">
<emphasis id="C18F3F7E242FFFAC753D6372F5A7FD8B" box="[94,187,536,560]" italics="true" pageId="45" pageNumber="262">
<typeStatus id="2C405DCE242FFFAC753D6372F5AAFD8B" box="[94,182,536,560]" pageId="45" pageNumber="262" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC242FFFAC75BB6372F40BFD8B" box="[216,279,536,560]" name="Brazil" pageId="45" pageNumber="262">Brésil</collectingCountry>
,
<collectingRegion id="313F2D8E242FFFAC74526372F4D6FD8B" box="[305,458,536,560]" country="Brazil" name="Mato Grosso" pageId="45" pageNumber="262">Mato Grosso</collectingRegion>
,
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,
<geoCoordinate id="96CF85AB242FFFAC753D6352F41EFDEB" box="[94,258,568,592]" degrees="59" direction="west" minutes="13" orientation="longitude" pageId="45" pageNumber="262" precision="1" seconds="02.05" value="-59.217236">59°1302.05”W</geoCoordinate>
,
<collectorName id="5E0E86BA242FFFAC74756352F4E2FDEB" box="[278,510,568,592]" pageId="45" pageNumber="262">Chapada dos Parecis</collectorName>
,
<quantity id="34034E89242FFFAC77726352F742FDEB" box="[529,606,568,592]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="45" pageNumber="262" unit="km" value="30.0">30 km</quantity>
N
<location id="F624B5B7242FFFAC753D6332F5CBFDCB" LSID="urn:lsid:plazi:treatment:7B52527A242EFFAC767863DDF406FA34:F624B5B7242FFFAC753D6332F5CBFDCB" box="[94,215,600,624]" country="Brazil" latitude="-14.2641115" longLatPrecision="1" longitude="-59.217236" name="d'Uirapuru" pageId="45" pageNumber="262" stateProvince="Mato Grosso">dUirapuru</location>
], [01 au
<date id="8745C5AC242FFFAC74216332F4DCFDCB" box="[322,448,600,624]" pageId="45" pageNumber="262" value="2000-12-15">
<collectingDate id="97013C44242FFFAC74216332F4DCFDCB" box="[322,448,600,624]" pageId="45" pageNumber="262" value="2000-12-15">15-12-2000</collectingDate>
</date>
,
<collectorName id="5E0E86BA242FFFAC74B26332F75FFDD4" box="[465,579,600,624]" pageId="45" pageNumber="262">A. Foucart</collectorName>
leg.]. [
<typeStatus id="2C405DCE242FFFAC75066312F5D2FD2B" box="[101,206,632,656]" pageId="45" pageNumber="262" type="holotype">Holotype</typeStatus>
], [
<taxonomicName id="34FB98EF242FFFAC758A6312F4B1FD2B" authorityName="Chassain" authorityYear="2005" box="[233,429,632,656]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="costae">
<emphasis id="C18F3F7E242FFFAC758A6312F4B1FD2B" box="[233,429,632,656]" italics="true" pageId="45" pageNumber="262">Physodactylus costae</emphasis>
</taxonomicName>
,
<typeStatus id="2C405DCE242FFFAC74DE6312F73DFD2B" box="[445,545,632,656]" pageId="45" pageNumber="262" type="holotype">Holotype</typeStatus>
,
<determinerName id="06980FA8242FFFAC774D6312F59AFD0B" pageId="45" pageNumber="262">J. Chassain</determinerName>
det. 04], male (
<collectionCode id="95EA7BA9242FFFAC744863F2F49BFD0B" box="[299,391,664,688]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="45" pageNumber="262">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C242FFFAC753D63B2F5CAFCAB" blockId="45.[94,643,728,784]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC753D63B2F5FDFD4B" box="[94,225,728,752]" italics="true" pageId="45" pageNumber="262">Distribution:</emphasis>
<collectingCountry id="8BECA3FC242FFFAC758E63B2F450FD4B" box="[237,332,728,752]" name="Brazil" pageId="45" pageNumber="262">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E242FFFAC743863B3F4FFFD54" box="[347,483,728,752]" italics="true" pageId="45" pageNumber="262">
<collectingRegion id="313F2D8E242FFFAC743863B3F4C2FD54" box="[347,478,728,752]" country="Brazil" name="Mato Grosso" pageId="45" pageNumber="262">Mato Grosso</collectingRegion>
:
</emphasis>
Campo Novo dos Parecis.
</paragraph>
<paragraph id="F344E36C242FFFAC753D6252F406FA34" blockId="45.[94,643,824,1424]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC753D6252F5A4FCEB" box="[94,184,824,848]" italics="true" pageId="45" pageNumber="262">Remarks:</emphasis>
Sternite VIII was not examined.
<taxonomicName id="34FB98EF242FFFAC77416252F5D5FCCB" authorityName="Chassain" authorityYear="2005" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="costae">
<emphasis id="C18F3F7E242FFFAC77416252F5D5FCCB" italics="true" pageId="45" pageNumber="262">Physodactylus costae</emphasis>
</taxonomicName>
(
<figureCitation id="6BC0FFE9242FFFAC75B46232F42BFCCB" box="[215,311,856,880]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="45" pageNumber="262">Fig. 35D</figureCitation>
) and
<taxonomicName id="34FB98EF242FFFAC74106233F4F5FCCB" box="[371,489,856,880]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="fleutiauxi">
<emphasis id="C18F3F7E242FFFAC74106233F4F5FCCB" box="[371,489,856,880]" italics="true" pageId="45" pageNumber="262">P. fleutiauxi</emphasis>
</taxonomicName>
(
<figureCitation id="6BC0FFE9242FFFAC74946232F749FCCB" box="[503,597,856,880]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="45" pageNumber="262">Fig. 35G</figureCitation>
) are sympatric species from Chapada dos Parecis (
<collectingRegion id="313F2D8E242FFFAC772A6212F5C0FC0B" country="Brazil" name="Mato Grosso" pageId="45" pageNumber="262">Mato Grosso state</collectingRegion>
) and were described by
<bibRefCitation id="976A9E9D242FFFAC74B062F2F763FC0B" author="CHASSAIN, J." box="[467,639,920,944]" pageId="45" pageNumber="262" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain (2005)</bibRefCitation>
. They are most similar to
<taxonomicName id="34FB98EF242FFFAC741262D3F4CAFC6B" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[369,470,952,976]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="C18F3F7E242FFFAC741262D3F4CAFC6B" box="[369,470,952,976]" italics="true" pageId="45" pageNumber="262">P. sulcatus</emphasis>
</taxonomicName>
in the antennal length: this do not surpass the anterior third of pronotum. The habitus illustrations in
<bibRefCitation id="976A9E9D242FFFAC74BB6292F79EFBAB" author="CHASSAIN, J." box="[472,642,1016,1040]" pageId="45" pageNumber="262" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain (2005)</bibRefCitation>
show
<taxonomicName id="34FB98EF242FFFAC75FD6573F404FB8B" box="[158,280,1048,1072]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="fleutiauxi">
<emphasis id="C18F3F7E242FFFAC75FD6573F404FB8B" box="[158,280,1048,1072]" italics="true" pageId="45" pageNumber="262">P. fleutiauxi</emphasis>
</taxonomicName>
with lamella of meso- and metatarsomere I as large as the other ones and
<taxonomicName id="34FB98EF242FFFAC77536553F79EFBEB" authorityName="Chassain" authorityYear="2005" box="[560,642,1081,1104]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="costae">
<emphasis id="C18F3F7E242FFFAC77536553F79EFBEB" box="[560,642,1081,1104]" italics="true" pageId="45" pageNumber="262">P. costae</emphasis>
</taxonomicName>
without any lamella in all legs. Actually, as described by
<bibRefCitation id="976A9E9D242FFFAC75E36512F431FB2B" author="CHASSAIN, J." box="[128,301,1144,1168]" pageId="45" pageNumber="262" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain (2005)</bibRefCitation>
, the lamellae of the tarsomeres I are very small on the pro- and mesothoracic legs and absent on the metathoracic leg in both species.
<taxonomicName id="34FB98EF242FFFAC773665D2F5E2FB4B" authorityName="Chassain" authorityYear="2005" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="costae">
<emphasis id="C18F3F7E242FFFAC773665D2F5E2FB4B" italics="true" pageId="45" pageNumber="262">Physodactylus costae</emphasis>
</taxonomicName>
is most similar to
<emphasis id="C18F3F7E242FFFAC74AA65B2F5BCFAAB" italics="true" pageId="45" pageNumber="262">
<taxonomicName id="34FB98EF242FFFAC74AA65B2F587FAAB" authorityName="Chassain" authorityYear="2005" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="fleutiauxi">Physodactylus fleutiauxi</taxonomicName>
,
</emphasis>
from which can be separated by the features cited in the key proposed herein.
<taxonomicName id="34FB98EF242FFFAC74DC6472F79EFA8B" authorityName="Chassain" authorityYear="2005" box="[447,642,1304,1328]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="costae">
<emphasis id="C18F3F7E242FFFAC74DC6472F79EFA8B" box="[447,642,1304,1328]" italics="true" pageId="45" pageNumber="262">Physodactylus costae</emphasis>
</taxonomicName>
has aedeagus similar to that of
<emphasis id="C18F3F7E242FFFAC74C66453F739FAEB" box="[421,549,1336,1360]" italics="true" pageId="45" pageNumber="262">
<taxonomicName id="34FB98EF242FFFAC74C66453F73DFAEB" authorityName="Rosa" authorityYear="2014" box="[421,545,1336,1360]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="latithorax">P. latithorax</taxonomicName>
,
</emphasis>
however the later has penis wider at apex and narrower at base of the basal struts.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,136 @@
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<heading id="A80C5400242FFFAC75C164BDF722FA4B" bold="true" box="[162,574,1495,1520]" centered="true" fontSize="10" level="2" pageId="45" pageNumber="262" reason="2">
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<emphasis id="C18F3F7E242FFFAC75C164BDF722FA4B" bold="true" box="[162,574,1495,1520]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC75C164BDF491FA54" bold="true" box="[162,397,1495,1519]" italics="true" pageId="45" pageNumber="262">Physodactylus fischeri</emphasis>
Fleutiaux, 1940
</emphasis>
</taxonomicName>
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<paragraph id="F344E36C242FFFAC74686492F4CAF9AB" blockId="45.[162,574,1495,1552]" box="[267,470,1528,1552]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC74686492F4CAF9AB" bold="true" box="[267,470,1528,1552]" pageId="45" pageNumber="262">
(
<figureCitation id="6BC0FFE9242FFFAC74716492F47AF9AB" box="[274,358,1528,1552]" captionStart="FIGURE 19" captionStartId="43.[95,174,1826,1845]" captionTargetBox="[100,1222,389,1798]" captionTargetId="figure-93@43.[94,1228,385,1802]" captionTargetPageId="43" captionText="FIGURE 19: Physodactylus chassaini sp. nov. A, antenna; B, pronotum; C, mesotibia; D, E, aedeagus (dorsal, ventral). Physodactylus fischeri. F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641409" httpUri="https://zenodo.org/record/12641409/files/figure.png" pageId="45" pageNumber="262">Figs. 19</figureCitation>
F-G, 35E)
</emphasis>
</paragraph>
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<treatmentCitation id="725AC57D242FFFAC753D6752F72CF9EB" author="FLEUTIAUX, E." box="[94,560,1592,1616]" page="166" pageId="45" pageNumber="262" year="1940">
<taxonomicName id="34FB98EF242FFFAC753D6752F72CF9EB" authority="Fleutiaux, 1940 c: 166" authorityName="Fleutiaux" authorityPageNumber="166" authorityYear="1940" box="[94,560,1592,1616]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="C18F3F7E242FFFAC753D6752F428F9EB" box="[94,308,1592,1616]" italics="true" pageId="45" pageNumber="262">Physodactylus fischeri</emphasis>
<bibRefCitation id="976A9E9D242FFFAC74236752F72CF9EB" author="FLEUTIAUX, E." box="[320,560,1592,1616]" pageId="45" pageNumber="262" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 166</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="725AC57D242FFFAC77226752F448F9CB" author="BLACKWELDER, R. E." page="292" pageId="45" pageNumber="262" year="1944">
<bibRefCitation id="976A9E9D242FFFAC77226752F448F9CB" author="BLACKWELDER, R. E." pageId="45" pageNumber="262" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7242FFFAC753D67F2F6C9F8EB" pageId="45" pageNumber="262" type="description">
<paragraph id="F344E36C242FFFAC753D67F2F6D8FB8B" blockId="45.[94,643,1688,1872]" lastBlockId="45.[680,1229,184,1072]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC753D67F2F5FEF90B" box="[94,226,1688,1712]" italics="true" pageId="45" pageNumber="262">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9242FFFAC745F67F2F481F90B" box="[316,413,1688,1712]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="45" pageNumber="262">Fig. 35E</figureCitation>
): Integument bright, head with frons black, gradually lighter posteriorly with the same color of pronotum at base, prothorax reddish-brown, antenna, elytra and ventral abdomen black, ventral pterothorax and legs dark brown; pilosity brown. Total length 17.0 mm; elytral base 1.03x as wide as prothorax, elytra 2.98x longer than pronotum. Frons slightly concave, frontal carina not produced; antenna reaching the half length of pronotum; antennomere IV 1.05x as wide as long, V-VII 1.20x wider than long; VIII-X 1.0-1.1x as wide as long; XI oval 1.28x longer than wide. Pronotum 1.28x wider than long, lateral sides rounded convergent from posterior angles to anterior margin; anteromedian margin weakly produced; posterior angles short, convex, parallel; lateral carina dorsally visible only on posterior third, absent on anterior half; disc with punctures 2-3 diameters apart, larger and denser on lateral and anterior borders. Lamellae of pro- and mesotarsomeres I smaller than the others, absent or minuscule on metatarsomere I. Elytra subparallel on anterior 2/3 then tapering to apex; striae with large and small dense punctation, indistinct on apical region where the punctation is mixed; interstices convex and rugose, scarcely punctate; apical striae with punctures larger than those of pronotal lateral border. Abdomen with ventrites 2-4 concave laterally, 1-4 evenly covered with decumbent, moderately long pilosity, setae of ventrite 1 as long as those of ventrite 2. Pregenitalic segments and aedeagus covered with stout dark-brown setae. Sternite VIII with posteromedian margin emarginate, anterior sclerotization 0.28x the total width of sternite; sternite IX with apex rounded.
</paragraph>
<paragraph id="F344E36C242FFFAC77CB6532F16BFAEB" blockId="45.[680,1229,1112,1360]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC77CB6532F61DFBCB" box="[680,769,1112,1136]" italics="true" pageId="45" pageNumber="262">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9242FFFAC76776532F682FBCB" box="[788,926,1112,1136]" captionStart="FIGURE 19" captionStartId="43.[95,174,1826,1845]" captionTargetBox="[100,1222,389,1798]" captionTargetId="figure-93@43.[94,1228,385,1802]" captionTargetPageId="43" captionText="FIGURE 19: Physodactylus chassaini sp. nov. A, antenna; B, pronotum; C, mesotibia; D, E, aedeagus (dorsal, ventral). Physodactylus fischeri. F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641409" httpUri="https://zenodo.org/record/12641409/files/figure.png" pageId="45" pageNumber="262">Figs. 19F, G</figureCitation>
): Phallobase 0.52x the total length of aedeagus, 1.27x longer than wide, length ratio between lateroposterior and median parts 11.5; paramere with ventral surface partly sclerotized, anteromedian margin with a long median sclerotized strip convergent anteriad; penis with basal strut 0.27x its total length, articulated to the parameres by a translucent membrane; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C242FFFAC77CB6412F69CF9AB" blockId="45.[680,1229,1400,1552]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC77CB6412F619FA2B" box="[680,773,1400,1424]" italics="true" pageId="45" pageNumber="262">
<typeStatus id="2C405DCE242FFFAC77CB6412F61CFA2B" box="[680,768,1400,1424]" pageId="45" pageNumber="262" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingRegion id="313F2D8E242FFFAC76436412F643FA2B" box="[800,863,1400,1424]" country="Brazil" name="Bahia" pageId="45" pageNumber="262">Bahia</collectingRegion>
,
<collectingCountry id="8BECA3FC242FFFAC76146412F6AAFA2B" box="[887,950,1400,1424]" name="Brazil" pageId="45" pageNumber="262">Brasil</collectingCountry>
], [TYPE], [
<taxonomicName id="34FB98EF242FFFAC712B6412F629FA0B" authority="Fleut." authorityName="Fleutiaux" authorityYear="1940" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="C18F3F7E242FFFAC712B6412F7F1FA0B" italics="true" pageId="45" pageNumber="262">Physodactylus fischeri</emphasis>
Fleut.
</taxonomicName>
, COLLECTION FLEUTIAUX type], [Muséum Paris, Collection E. Fleutiaux], [
<typeStatus id="2C405DCE242FFFAC710964D2F7F1FA54" pageId="45" pageNumber="262" type="lectotype">LECTOTYPE</typeStatus>
], [
<taxonomicName id="34FB98EF242FFFAC766464B2F182FA4B" authority="Fleut., J. Chassain" authorityName="Fleut., J. Chassain" box="[775,1182,1496,1520]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="C18F3F7E242FFFAC766464B2F6C4FA4B" box="[775,984,1496,1520]" italics="true" pageId="45" pageNumber="262">Physodactylus fischeri</emphasis>
Fleut., J. Chassain
</taxonomicName>
det. 05], male (MNHN).
</paragraph>
<paragraph id="F344E36C242FFFAC77CB6752F6C0F9EB" blockId="45.[680,988,1592,1616]" box="[680,988,1592,1616]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC77CB6752F636F9EB" box="[680,810,1592,1616]" italics="true" pageId="45" pageNumber="262">Distribution:</emphasis>
<collectingCountry id="8BECA3FC242FFFAC76526752F68CF9EB" box="[817,912,1592,1616]" name="Brazil" pageId="45" pageNumber="262">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E242FFFAC76FA6752F6C0F9EB" box="[921,988,1592,1616]" italics="true" pageId="45" pageNumber="262">
<collectingRegion id="313F2D8E242FFFAC76FA6752F6C4F9EB" box="[921,984,1592,1616]" country="Brazil" name="Bahia" pageId="45" pageNumber="262">Bahia</collectingRegion>
.
</emphasis>
</paragraph>
<paragraph id="F344E36C242FFFAC77CB6712F6C9F8EB" blockId="45.[680,1229,1656,1872]" pageId="45" pageNumber="262">
<emphasis id="C18F3F7E242FFFAC77CB6712F61EF92B" box="[680,770,1656,1680]" italics="true" pageId="45" pageNumber="262">Remarks:</emphasis>
The apex of the prosternal process was not examined.
<taxonomicName id="34FB98EF242FFFAC767F67F2F6F3F90B" box="[796,1007,1688,1712]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="262" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="C18F3F7E242FFFAC767F67F2F6F3F90B" box="[796,1007,1688,1712]" italics="true" pageId="45" pageNumber="262">Physodactylus fischeri</emphasis>
</taxonomicName>
is recognized by the following combination of characters: antennae reaching the posterior half of pronotum, lateral pronotal carina absent on anterior half, elytral striae densely punctate and indistinct on apical region and pilosity of the ventrites 1-4 subequal.
</paragraph>
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<taxonomicName id="34FB98EF2430FFB3758864DDF4F5FA74" authorityName="Rosa" authorityYear="2014" box="[235,489,1463,1487]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="267" phylum="Arthropoda" rank="species" species="gounellei" status="sp. nov.">
<emphasis id="C18F3F7E2430FFB3758864DDF4F5FA74" bold="true" box="[235,489,1463,1487]" italics="true" pageId="50" pageNumber="267">Physodactylus gounellei</emphasis>
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<emphasis id="C18F3F7E2430FFB3745064B2F4E5FA4B" bold="true" box="[307,505,1496,1520]" pageId="50" pageNumber="267">
(
<figureCitation id="6BC0FFE92430FFB3745864B2F493FA4B" box="[315,399,1496,1520]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="50" pageNumber="267">Figs. 21</figureCitation>
J-M, 35J)
</emphasis>
</paragraph>
<paragraph id="F344E36C2430FFB375E76772F74CF9EB" blockId="50.[132,680,1560,1619]" pageId="50" pageNumber="267">
<emphasis id="C18F3F7E2430FFB375E76772F5ECF98B" box="[132,240,1560,1584]" italics="true" pageId="50" pageNumber="267">Etymology:</emphasis>
In honor to the French entomologist Pierre Émile Gounelle, who collected the
<typeStatus id="2C405DCE2430FFB3748D6752F750F9EB" box="[494,588,1592,1616]" pageId="50" pageNumber="267" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="F344E36C2430FFB375E76713F1BAFA34" blockId="50.[132,680,1656,1872]" lastBlockId="50.[718,1267,184,1424]" pageId="50" pageNumber="267">
<emphasis id="C18F3F7E2430FFB375E76713F5E8F92B" box="[132,244,1657,1680]" italics="true" pageId="50" pageNumber="267">Description</emphasis>
(male,
<figureCitation id="6BC0FFE92430FFB374266712F487F92B" box="[325,411,1656,1680]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="50" pageNumber="267">Fig. 35J</figureCitation>
): Integument very bright; antennae, legs, pronotum and elytra light brown to brown; head and ventral surface of thorax dark brown; abdomen with ventrites 1-3 black with lateral borders light brown, ventrite 4 brown with lateral border lighter, V evenly light brown or brown with borders lighter; pilosity light to dark brown. Total length 10.0-11.0 mm; elytral base 0.96x as wide as prothorax, elytra 2.90-3.04x longer than pronotum. Frons concave along the anterior border, with a pair of transverse protuberances between antennal insertions; frontal carina strongly produced; antenna (
<figureCitation id="6BC0FFE92430FFB371F76052F1F7FEEB" box="[1172,1259,312,336]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="50" pageNumber="267">Fig. 21J</figureCitation>
) reaching the half length of pronotum; antennomere IV 1.28x wider than long, V-IX 1.67x wider than long; X 1.58x wider than long; XI oval, 1.25x longer than wide. Pronotum (
<figureCitation id="6BC0FFE92430FFB376A460D2F134FE6B" box="[967,1064,440,464]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="50" pageNumber="267">Fig. 21K</figureCitation>
) 1.20-1.24x wider than long, lateral sides rounded from posterior angles to anterior margin, strongly narrowed on anterior angles, anteromedian margin produced anteriad over head; posterior angles elongate, convex, parallel; lateral carina complete strongly prominent on posterior 2/3, not visible dorsally on anterior third; disc and lateral margin with punctures 1-2 diameters apart, larger on lateral and anterior borders, homogenously distributed; prosternal process without subapical tooth. Lamellae of pro- and mesotarsomeres I smaller than the others, present on metatarsomere I. Elytra subparallel on anterior 2/3 then tapering to apex; apical striae with a row of punctures as large as those of pronotal lateral border, interstices convex, smooth, scarcely punctate. Abdomen with ventrites 2-3 concave laterally, pilosity of the approximately inner 3/4 (0.76-0.80) of the ventrites 1-3 separated from those of the marginal plates by a shiny impuctate longitudinal band; ventrite 1 with short setae on inner 3/4 and a little longer and denser setae on marginal plate; ventrites 2-3 with setae a little longer, semi-erect and denser than those of the inner part of ventrite 1, spars- er and decumbent on marginal plate; ventrite 4 with setae sparser and longer than those of inner part of ventrite 3, denser on lateral border; ventrite 5 with setae denser and stouter than those of ventrite 4. Pregenitalic segments and aedeagus covered with brown setae. Sternite VIII subrectangular, nearly straight posteriorly, anterior sclerotization 0.69x the total width of sternite; sternite IX tapering to apex.
</paragraph>
<paragraph id="F344E36C2430FFB377AD64D2F618F914" blockId="50.[718,1266,1464,1711]" pageId="50" pageNumber="267">
<emphasis id="C18F3F7E2430FFB377AD64D2F63BFA6B" box="[718,807,1464,1488]" italics="true" pageId="50" pageNumber="267">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92430FFB3765964D2F6D8FA6B" box="[826,964,1464,1488]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="50" pageNumber="267">Fig. 21L, M</figureCitation>
): Phallobase 0.49x the total length of aedeagus, 1.33x longer than wide, length ratio between lateroposterior and median parts 3.37; paramere with ventral surface sclerotized, with anteromedian margin distinct; penis with basal strut 0.31x its total length, articulated to the parameres by a translucent membrane; ventral sclerite tapering to apex.
</paragraph>
<paragraph id="F344E36C2430FFB277AD67B2F79FFECB" blockId="50.[718,1267,1752,1872]" lastBlockId="51.[94,644,184,368]" lastPageId="51" lastPageNumber="268" pageId="50" pageNumber="267">
<emphasis id="C18F3F7E2430FFB377AD67B2F635F94B" box="[718,809,1752,1776]" italics="true" pageId="50" pageNumber="267">
<typeStatus id="2C405DCE2430FFB377AD67B2F638F94B" box="[718,804,1752,1776]" pageId="50" pageNumber="267" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Bello Horizonte, Minas], [Museum Paris,
<collectingCountry id="8BECA3FC2430FFB377AD6792F615F8AB" box="[718,777,1784,1808]" name="Brazil" pageId="50" pageNumber="267">Brèsil</collectingCountry>
, E. Gounelle. 1911], [Comparé au type par J. Chassain = conforme], [
<taxonomicName id="34FB98EF2430FFB376DD6672F1C5F88B" authority="Fleut." authorityName="Fleut." box="[958,1241,1816,1840]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="267" phylum="Arthropoda" rank="species" species="carreti">Physodactylus carreti Fleut.</taxonomicName>
,
<taxonomicName id="34FB98EF2430FFB371816672F6E8F8EB" authority="Flx. J. Chassain" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="267" phylum="Arthropoda" rank="variety" species="carreti" variety="testaceipennis">v. testaceipennis Flx. J. Chassain</taxonomicName>
det. 05], male (MNHN).
<emphasis id="C18F3F7E2431FFB2753D61D3F5A2FF6B" box="[94,190,185,208]" italics="true" pageId="51" pageNumber="268">
<typeStatus id="2C405DCE2431FFB2753D61D3F5A4FF6B" box="[94,184,185,208]" pageId="51" pageNumber="268" type="paratype">Paratypes</typeStatus>
:
</emphasis>
[Muséum Paris, Coll. Gounelle, 1915], [Muséum Paris, Coll. E. Fleutiaux], [Comparé au type par J Chassain, conforme], [
<taxonomicName id="34FB98EF2431FFB2743E6192F5CCFE8B" authority="Fleut., J. Chassain" authorityName="Fleut., J. Chassain" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="species" species="carreti">Physodactylus carreti Fleut., J. Chassain</taxonomicName>
det 05],
<specimenCount id="E5FD28E52431FFB274516072F479FE94" box="[306,357,280,304]" count="1" pageId="51" pageNumber="268" type="generic">1 ex.</specimenCount>
(MNHN); [Muséum Paris, Coll. Gounelle, 1915], [
<taxonomicName id="34FB98EF2431FFB2742C6052F775FEEB" authority="Fleut." authorityName="Fleut." box="[335,617,312,336]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="species" species="carreti">Physodactylus carreti Fleut.</taxonomicName>
,
<taxonomicName id="34FB98EF2431FFB277106052F490FECB" authority="Flx., J. Chassain" authorityName="Flx., J. Chassain" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="variety" species="carreti" variety="testaceipennis">v. testaceipennis Flx., J. Chassain</taxonomicName>
det 05],
<specimenCount id="E5FD28E52431FFB274816032F70DFED4" box="[482,529,344,368]" count="1" pageId="51" pageNumber="268" type="generic">1 ex.</specimenCount>
(MNHN).
</paragraph>
<paragraph id="F344E36C2431FFB2753D60F2F79FFE0B" blockId="51.[94,643,408,432]" box="[94,643,408,432]" pageId="51" pageNumber="268">
<emphasis id="C18F3F7E2431FFB2753D60F2F5FDFE0B" box="[94,225,408,432]" italics="true" pageId="51" pageNumber="268">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2431FFB2758460F2F45AFE0B" box="[231,326,408,432]" name="Brazil" pageId="51" pageNumber="268">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2431FFB2742D60F3F4C4FE14" box="[334,472,408,432]" italics="true" pageId="51" pageNumber="268">
<collectingRegion id="313F2D8E2431FFB2742D60F3F4CEFE14" box="[334,466,408,432]" country="Brazil" name="Minas Gerais" pageId="51" pageNumber="268">Minas Gerais</collectingRegion>
:
</emphasis>
Belo Horizonte.
</paragraph>
<paragraph id="F344E36C2431FFB2753D60B2F1D5FE8B" blockId="51.[94,643,472,688]" lastBlockId="51.[680,1229,184,304]" pageId="51" pageNumber="268">
<emphasis id="C18F3F7E2431FFB2753D60B2F4BAFE4B" box="[94,422,472,496]" italics="true" pageId="51" pageNumber="268">
Remarks:
<taxonomicName id="34FB98EF2431FFB275A260B2F4BAFE4B" authorityName="Rosa" authorityYear="2014" box="[193,422,472,496]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="species" species="gounellei" status="sp. nov.">Physodactylus gounellei</taxonomicName>
</emphasis>
<taxonomicNameLabel id="DABC82052431FFB274CD60B2F4E3FE54" box="[430,511,472,495]" pageId="51" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="34FB98EF2431FFB2753D6093F5F2FDAB" authorityName="Rosa" authorityYear="2014" box="[94,238,504,528]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="C18F3F7E2431FFB2753D6093F5F2FDAB" box="[94,238,504,528]" italics="true" pageId="51" pageNumber="268">P. tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052431FFB275976092F45FFDB4" box="[244,323,504,527]" pageId="51" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
by its pronotal punctation and having the pilosity of the abdominal ventrites 1-3 divided laterally by a longitudinal smooth band, but differs from this species in its brown pilosity, lateral carina less prominent, anteromedian margin of pronotum less produced over head, absence of tubercle on pronotum, elytral striae with smaller punctures, the longer setae of the abdominal ventrite 1, the position of the smooth longitudinal band on the abdomen and by the stouter and denser setae on the ventrite 5.
</paragraph>
</subSubSection>
</treatment>
</document>

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<paragraph id="F344E36C2431FFB277CB601DF1D0FE2B" blockId="51.[680,1228,375,432]" box="[680,1228,375,400]" pageId="51" pageNumber="268">
<heading id="A80C54002431FFB277CB601DF1D0FE2B" bold="true" box="[680,1228,375,400]" centered="true" fontSize="10" level="2" pageId="51" pageNumber="268" reason="2">
<taxonomicName id="34FB98EF2431FFB277CB601DF1D0FE2B" authority="Fischer von Waldheim, 1823" authorityName="Fischer von Waldheim" authorityYear="1823" box="[680,1228,375,400]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2431FFB277CB601DF1D0FE2B" bold="true" box="[680,1228,375,400]" pageId="51" pageNumber="268">
<emphasis id="C18F3F7E2431FFB277CB601DF687FE34" bold="true" box="[680,923,375,399]" italics="true" pageId="51" pageNumber="268">Physodactylus henningi</emphasis>
<bibRefCitation id="976A9E9D2431FFB276C26012F1D0FE2B" author="FISCHER VON WALDHEIM, M. G." box="[929,1228,376,400]" pageId="51" pageNumber="268" refId="ref49615" refString="FISCHER VON WALDHEIM, M. G. 1823. Entomographia imperii russici; genera insectorum systematica exposita et analysi iconographica instructa. Mosquae, Auctoritate Societatis Caesareae Naturae Scrutatorum Mosquensis. v. 2, 262 p." type="book" year="1823">Fischer von Waldheim, 1823</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72431FFB2765A60F2F120FE0B" box="[825,1084,408,432]" pageId="51" pageNumber="268" type="description">
<paragraph id="F344E36C2431FFB2765A60F2F120FE0B" blockId="51.[680,1228,375,432]" box="[825,1084,408,432]" pageId="51" pageNumber="268">
<emphasis id="C18F3F7E2431FFB2765A60F2F120FE0B" bold="true" box="[825,1084,408,432]" pageId="51" pageNumber="268">
(
<figureCitation id="6BC0FFE92431FFB2765C60F2F6ABFE0B" box="[831,951,408,432]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStart-2="FIGURE 24" captionStartId-0="51.[95,174,1826,1845]" captionStartId-1="52.[132,211,1852,1871]" captionStartId-2="53.[95,174,1852,1871]" captionTargetBox-0="[95,1226,727,1799]" captionTargetBox-1="[133,1263,1197,1817]" captionTargetBox-2="[109,1212,300,1823]" captionTargetId-0="figure-197@51.[94,1228,725,1802]" captionTargetId-1="figure-469@52.[132,1266,1185,1828]" captionTargetId-2="figure-52@53.[94,1228,298,1828]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionText-0="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." captionText-1="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." captionText-2="FIGURE 24: Physodactylus henningi. A, tergite VIII; B, sternite VIII; C, ovipositor (ventral); D, ovipositor and reproductive tract (dorsal)." figureDoi-0="http://doi.org/10.5281/zenodo.12641417" figureDoi-1="http://doi.org/10.5281/zenodo.12641421" figureDoi-2="http://doi.org/10.5281/zenodo.12641423" httpUri-0="https://zenodo.org/record/12641417/files/figure.png" httpUri-1="https://zenodo.org/record/12641421/files/figure.png" httpUri-2="https://zenodo.org/record/12641423/files/figure.png" pageId="51" pageNumber="268">Figs. 22-24</figureCitation>
,
<figureCitation id="6BC0FFE92431FFB276A060F2F6C3FE0B" box="[963,991,408,432]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="51" pageNumber="268">28</figureCitation>
,
<figureCitation id="6BC0FFE92431FFB2768960F2F118FE0B" box="[1002,1028,408,432]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="51" pageNumber="268">33</figureCitation>
F-M)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72431FFB577CB60B2F431FECB" lastPageId="52" lastPageNumber="269" pageId="51" pageNumber="268" type="reference_group">
<paragraph id="F344E36C2431FFB277CB60B2F6C7FDCB" blockId="51.[680,1229,472,688]" pageId="51" pageNumber="268">
<treatmentCitationGroup id="D3EBC4422431FFB277CB60B2F6C7FDCB" pageId="51" pageNumber="268">
<treatmentCitation id="725AC57D2431FFB277CB60B2F617FDAB" author="FISCHER VON WALDHEIM, M. G." page="303" pageId="51" pageNumber="268" year="1823">
<taxonomicName id="34FB98EF2431FFB277CB60B2F617FDAB" authority="Fischer von Waldheim, 1823: 303" authorityName="Fischer von Waldheim" authorityPageNumber="303" authorityYear="1823" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2431FFB277CB60B2F691FE4B" box="[680,909,472,496]" italics="true" pageId="51" pageNumber="268">Physodactylus henningi</emphasis>
<bibRefCitation id="976A9E9D2431FFB276F560B2F617FDAB" author="FISCHER VON WALDHEIM, M. G." pageId="51" pageNumber="268" refId="ref49615" refString="FISCHER VON WALDHEIM, M. G. 1823. Entomographia imperii russici; genera insectorum systematica exposita et analysi iconographica instructa. Mosquae, Auctoritate Societatis Caesareae Naturae Scrutatorum Mosquensis. v. 2, 262 p." type="book" year="1823">Fischer von Waldheim, 1823: 303</bibRefCitation>
</taxonomicName>
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,
<treatmentCitation id="725AC57D2431FFB276786092F691FDAB" author="FISCHER VON WALDHEIM, M. G." box="[795,909,504,528]" page="450" pageId="51" pageNumber="268" year="1824">
<bibRefCitation id="976A9E9D2431FFB276786092F691FDAB" author="FISCHER VON WALDHEIM, M. G." box="[795,909,504,528]" pageId="51" pageNumber="268" pagination="448 - 451" refId="ref49654" refString="FISCHER VON WALDHEIM, M. G. 1824. Extrait d'une lettre aadressee a M. Henning sur le Physodactyle, nouveaux genre de Coleoptere voisin des Taupin. Annales des Sciences Naturelles, 3: 448 - 451, plate 27 B." type="journal article" year="1824">1824: 450</bibRefCitation>
</treatmentCitation>
; Guérin-Méneville, 1829: 43;
<treatmentCitation id="725AC57D2431FFB277826372F6CBFD8B" author="LACORDAIRE, J. T." box="[737,983,536,560]" page="237" pageId="51" pageNumber="268" year="1857">
<bibRefCitation id="976A9E9D2431FFB277826372F6CBFD8B" author="LACORDAIRE, J. T." box="[737,983,536,560]" pageId="51" pageNumber="268" refId="ref50447" refString="LACORDAIRE, J. T. 1857. Histoire Naturelle des Insectes. Genera des Coleopteres. Paris, Librairie Encyclopedique de Roret. v. 4, 579 p." type="book" year="1857">Lacordaire, 1857: 237</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2431FFB2768A6372F1D4FD8B" author="SCHWARZ, O." box="[1001,1224,536,560]" page="311" pageId="51" pageNumber="268" year="1906">
<bibRefCitation id="976A9E9D2431FFB2768A6372F1D4FD8B" author="SCHWARZ, O." box="[1001,1224,536,560]" pageId="51" pageNumber="268" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 311</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2431FFB277826352F6DBFDEB" author="FLEUTIAUX, E." box="[737,967,568,592]" page="406" pageId="51" pageNumber="268" year="1892">
<bibRefCitation id="976A9E9D2431FFB277826352F6DBFDEB" author="FLEUTIAUX, E." box="[737,967,568,592]" pageId="51" pageNumber="268" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2431FFB276BB6352F1D4FDEB" author="FLEUTIAUX, E." box="[984,1224,568,592]" page="165" pageId="51" pageNumber="268" year="1940">
<bibRefCitation id="976A9E9D2431FFB276BB6352F1D4FDEB" author="FLEUTIAUX, E." box="[984,1224,568,592]" pageId="51" pageNumber="268" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 165</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2431FFB277826332F6CBFDCB" author="BLACKWELDER, R. E." box="[737,983,600,624]" page="292" pageId="51" pageNumber="268" year="1944">
<bibRefCitation id="976A9E9D2431FFB277826332F6CBFDCB" author="BLACKWELDER, R. E." box="[737,983,600,624]" pageId="51" pageNumber="268" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="F344E36C2431FFB277CB6312F64FFD0B" blockId="51.[680,1229,472,688]" pageId="51" pageNumber="268">
<treatmentCitationGroup id="D3EBC4422431FFB277CB6312F64FFD0B" pageId="51" pageNumber="268">
<treatmentCitation id="725AC57D2431FFB277CB6312F653FD0B" author="FLEUTIAUX, E." page="406" pageId="51" pageNumber="268" year="1892">
<taxonomicName id="34FB98EF2431FFB277CB6312F653FD0B" authority="Fleutiaux, 1892: 406" authorityName="Fleutiaux" authorityPageNumber="406" authorityYear="1892" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="variety" species="henningi" variety="erythrocephalus">
<emphasis id="C18F3F7E2431FFB277CB6312F692FD2B" box="[680,910,632,656]" italics="true" pageId="51" pageNumber="268">Physodactylus henningi</emphasis>
var.
<emphasis id="C18F3F7E2431FFB276A56312F140FD2B" box="[966,1116,632,656]" italics="true" pageId="51" pageNumber="268">erythrocephalus</emphasis>
<bibRefCitation id="976A9E9D2431FFB271056312F653FD0B" author="FLEUTIAUX, E." pageId="51" pageNumber="268" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<caption id="A784B3E42431FFB2753C6648F175F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641417" ID-Zenodo-Dep="12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="51" pageNumber="268" startId="51.[95,174,1826,1845]" targetBox="[95,1226,727,1799]" targetPageId="51" targetType="figure">
<paragraph id="F344E36C2431FFB2753C6648F175F8F4" blockId="51.[94,1229,1826,1871]" pageId="51" pageNumber="268">
<emphasis id="C18F3F7E2431FFB2753C6648F5D2F88E" bold="true" box="[95,206,1826,1845]" pageId="51" pageNumber="268">FIGURE 22:</emphasis>
<emphasis id="C18F3F7E2431FFB275B06648F492F88E" box="[211,398,1826,1845]" italics="true" pageId="51" pageNumber="268">
<taxonomicName id="34FB98EF2431FFB275B06648F496F88E" authorityName="Fischer von Waldheim" authorityYear="1823" box="[211,394,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="268" phylum="Arthropoda" rank="species" species="henningi">Physodactylus henningi</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2431FFB274F16648F4B8F88E" bold="true" box="[402,420,1826,1845]" pageId="51" pageNumber="268">A,</emphasis>
antenna;
<emphasis id="C18F3F7E2431FFB274966648F714F88E" bold="true" box="[501,520,1826,1845]" pageId="51" pageNumber="268">B,</emphasis>
maxilla;
<emphasis id="C18F3F7E2431FFB277316648F77BF88E" bold="true" box="[594,615,1826,1845]" pageId="51" pageNumber="268">C,</emphasis>
mandible (dorsal);
<emphasis id="C18F3F7E2431FFB276656648F601F88E" bold="true" box="[774,797,1826,1845]" pageId="51" pageNumber="268">D,</emphasis>
labium;
<emphasis id="C18F3F7E2431FFB276066648F66BF88E" bold="true" box="[869,887,1826,1845]" pageId="51" pageNumber="268">E,</emphasis>
head (lateral);
<emphasis id="C18F3F7E2431FFB276926648F129F88E" bold="true" box="[1009,1077,1826,1845]" pageId="51" pageNumber="268">F, G, H,</emphasis>
prothorax (dorsal, ventral, lateral);
<emphasis id="C18F3F7E2431FFB275866656F5EFF8F4" bold="true" box="[229,243,1852,1871]" pageId="51" pageNumber="268">I,</emphasis>
hypomeron;
<emphasis id="C18F3F7E2431FFB274026656F473F8F4" bold="true" box="[353,367,1852,1871]" pageId="51" pageNumber="268">J,</emphasis>
mesoscutum and scutellar shield;
<emphasis id="C18F3F7E2431FFB277EB6656F781F8F4" bold="true" box="[648,669,1852,1871]" pageId="51" pageNumber="268">K,</emphasis>
pterothorax (ventral);
<emphasis id="C18F3F7E2431FFB2763B6656F694F8F4" bold="true" box="[856,904,1852,1871]" pageId="51" pageNumber="268">L, M,</emphasis>
abdomen (ventral, lateral).
</paragraph>
</caption>
<paragraph id="F344E36C2436FFB575E761D2F425FF4B" blockId="52.[132,681,184,368]" pageId="52" pageNumber="269">
<treatmentCitationGroup id="D3EBC4422436FFB575E761D2F425FF4B" pageId="52" pageNumber="269">
<treatmentCitation id="725AC57D2436FFB575E761D2F429FF4B" author="FLEUTIAUX, E." page="165" pageId="52" pageNumber="269" year="1940">
<taxonomicName id="34FB98EF2436FFB575E761D2F429FF4B" authority="Fleutiaux, 1940 c: 165" authorityName="Fleut., FLEUTIAUX" authorityPageNumber="165" authorityYear="1940" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="269" phylum="Arthropoda" rank="variety" species="henningi" variety="nigricollis">
<emphasis id="C18F3F7E2436FFB575E761D2F464FF6B" box="[132,376,184,208]" italics="true" pageId="52" pageNumber="269">Physodactylus henningi</emphasis>
var.
<emphasis id="C18F3F7E2436FFB574A861D2F736FF6B" box="[459,554,184,208]" italics="true" pageId="52" pageNumber="269">nigricollis</emphasis>
<bibRefCitation id="976A9E9D2436FFB5772161D2F429FF4B" author="FLEUTIAUX, E." pageId="52" pageNumber="269" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F344E36C2436FFB575E76192F425FE8B" blockId="52.[132,681,184,368]" pageId="52" pageNumber="269">
<treatmentCitationGroup id="D3EBC4422436FFB575E76192F425FE8B" pageId="52" pageNumber="269">
<treatmentCitation id="725AC57D2436FFB575E76192F429FE8B" author="FLEUTIAUX, E." page="165" pageId="52" pageNumber="269" year="1940">
<taxonomicName id="34FB98EF2436FFB575E76192F429FE8B" authority="Fleutiaux, 1940 c: 165" authorityName="Fleutiaux" authorityPageNumber="165" authorityYear="1940" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="269" phylum="Arthropoda" rank="variety" species="henningi" variety="puncticollis">
<emphasis id="C18F3F7E2436FFB575E76192F468FEAB" box="[132,372,248,272]" italics="true" pageId="52" pageNumber="269">Physodactylus henningi</emphasis>
var.
<emphasis id="C18F3F7E2436FFB574A36192F732FEAB" box="[448,558,248,272]" italics="true" pageId="52" pageNumber="269">puncticollis</emphasis>
<bibRefCitation id="976A9E9D2436FFB577216192F429FE8B" author="FLEUTIAUX, E." pageId="52" pageNumber="269" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F344E36C2436FFB575E76053F431FECB" blockId="52.[132,681,184,368]" pageId="52" pageNumber="269">
<treatmentCitationGroup id="D3EBC4422436FFB575E76053F431FECB" pageId="52" pageNumber="269">
<treatmentCitation id="725AC57D2436FFB575E76053F4F1FEEB" author="PERTY, J. A. M." box="[132,493,312,336]" page="25" pageId="52" pageNumber="269" year="1830">
<taxonomicName id="34FB98EF2436FFB575E76053F4F1FEEB" authority="Perty 1830: 25" authorityName="Perty" authorityPageNumber="25" authorityYear="1830" box="[132,493,312,336]" class="Insecta" family="Elateridae" genus="Drepanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="269" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="C18F3F7E2436FFB575E76053F423FEEB" box="[132,319,312,336]" italics="true" pageId="52" pageNumber="269">Drepanius clavipes</emphasis>
<bibRefCitation id="976A9E9D2436FFB574296052F4F1FEEB" author="PERTY, J. A. M." box="[330,493,312,336]" pageId="52" pageNumber="269" pagination="1 - 12" refId="ref50792" refString="PERTY, J. A. M. 1830. Insecta Brasiliensia. In: Spix, J. &amp; Martius, C. Delectus animalium articulatorum, quae in itinere per Brasiliam annis 1817 - 1820 jussu et auspiciis Maximilliani Josephi I Bavariae regis augustissimi peracto. Munchen, Frid. Fleischer. p. 1 - 60, pls. 1 - 12." type="book chapter" year="1830">Perty 1830: 25</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2436FFB574986052F5F9FECB" author="FLEUTIAUX, E." page="406" pageId="52" pageNumber="269" year="1892">
<bibRefCitation id="976A9E9D2436FFB574986052F5F9FECB" author="FLEUTIAUX, E." pageId="52" pageNumber="269" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
</treatmentCitation>
(syn.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72436FFB775E760F2F6E7F9EB" lastPageId="54" lastPageNumber="271" pageId="52" pageNumber="269" type="description">
<paragraph id="F344E36C2436FFB575E760F2F655FB34" blockId="52.[132,681,408,1168]" lastBlockId="52.[718,1267,184,1168]" pageId="52" pageNumber="269">
<emphasis id="C18F3F7E2436FFB575E760F2F41BFE0B" box="[132,263,408,432]" italics="true" pageId="52" pageNumber="269">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE92436FFB5740060F2F4D4FE0B" box="[355,456,408,432]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="52" pageNumber="269">Fig. 35K</figureCitation>
): Integument bright with brown pilosity and variable color patterns: 1) head and epipleura reddish-brown, pronotum evenly reddish-brown or with lateral carina, posterior angles and posterior margin black; antenna, ventral surface of prothorax and elytra black, pterothorax and legs dark reddish-brown, ventral abdomen black and dark reddish-brown; 2) head, antennae and elytra dark brown, prothorax reddish-brown, ventral pterothorax and abdomen dark reddish-brown; 3) head dark reddish-brown, pronotum, hypomeron, epipleura and middle of scutellum orange to reddish-brown; elytra, borders of scutellum, legs, antennae, prosternum, ventral pterothorax and abdomen dark brown; 4) evenly brown to dark reddish-brown with epipleura lighter; 5) evenly black; 6) dark brown with elytra lighter. Total length 14.0-20.0 mm; elytral base 1.0-1.08x as wide as prothorax, elytra 3.03-3.16x times longer than pronotum. Frons (
<figureCitation id="6BC0FFE92436FFB5776C62B2F771FC4B" box="[527,621,984,1008]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="52" pageNumber="269">Fig. 22E</figureCitation>
) flattened with a pair of oblique protuberances over antennal insertions, convergent posteriorly; frontal carina not produced; antenna (
<figureCitation id="6BC0FFE92436FFB574B86552F721FBEB" box="[475,573,1080,1104]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="52" pageNumber="269">Fig. 22A</figureCitation>
) reaching the posterior third of pronotum; antennomere IV 0.89-1.22x as wide as long, V-VIII 1.0-1.2x as wide as long; IX 0.89-1.07x as wide as long; X 1.14-1.43x longer than wide; XI elliptical, 1.85-2.10x longer than wide. Pronotum (
<figureCitation id="6BC0FFE92436FFB576AB6192F131FEAB" box="[968,1069,248,272]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="52" pageNumber="269">Figs. 22F</figureCitation>
,
<figureCitation id="6BC0FFE92436FFB571586192F171FEAB" box="[1083,1133,248,272]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="52" pageNumber="269">26H</figureCitation>
) 1.19-1.25x wider than long, lateral sides slightly rounded or straight convergent from posterior angles to anterior margin; anteromedian margin weakly produced; posterior angles elongate, convex, parallel; lateral carina complete, dorsally visible only on posterior third; disc with punctures 2-4 diameters apart to medium 1-2 diameters apart, larger on lateral and anterior borders, homogenously distributed; prosternal process (
<figureCitation id="6BC0FFE92436FFB576656372F675FD8B" box="[774,873,536,560]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="52" pageNumber="269">Fig. 22H</figureCitation>
) with a subapical tooth. Lamellae of pro- and mesotarsomeres I a little smaller than the others, present on metatarsomere I; pro- and mesotarsal lamellae as wide as or wider than the subsequent tarsomeres including its setae, largely visible on dorsal view. Elytra subparallel on anterior 2/3 then tapering to apex or tapering to apex from anterior third; apical striae with a row of punctures as large as those of pronotal lateral border, interstices weakly convex, rugose, sparsely punctate. Abdomen (
<figureCitation id="6BC0FFE92436FFB577B66232F679FCCB" box="[725,869,856,880]" captionStart="FIGURE 22" captionStartId="51.[95,174,1826,1845]" captionTargetBox="[95,1226,727,1799]" captionTargetId="figure-197@51.[94,1228,725,1802]" captionTargetPageId="51" captionText="FIGURE 22: Physodactylus henningi. A, antenna; B, maxilla; C, mandible (dorsal); D, labium; E, head (lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron; J, mesoscutum and scutellar shield; K, pterothorax (ventral); L, M, abdomen (ventral, lateral)." figureDoi="http://doi.org/10.5281/zenodo.12641417" httpUri="https://zenodo.org/record/12641417/files/figure.png" pageId="52" pageNumber="269">Figs. 22L, M</figureCitation>
) with ventrites 2-5 strongly concave laterally, ventrite 1 nearly entirely impunctate, except on borders which are usually scarcely pilose, rarely pilose; ventrites 2-5 densely covered with moderately long, stout and semi-erect setae. Pregenitalic segments and aedeagus covered with stout dark-brown setae. Sternite VIII (
<figureCitation id="6BC0FFE92436FFB576C16572F6E3FB8B" box="[930,1023,1048,1072]" captionStart="FIGURE 23" captionStartId="52.[132,211,1852,1871]" captionTargetBox="[133,1263,1197,1817]" captionTargetId="figure-469@52.[132,1266,1185,1828]" captionTargetPageId="52" captionText="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641421" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="52" pageNumber="269">Fig. 23A</figureCitation>
) with posterior margin nearly straight, anterior sclerotization 0.33-0.38x the total width of sternite; sternite IX (
<figureCitation id="6BC0FFE92436FFB571266532F1B9FBCB" box="[1093,1189,1112,1136]" captionStart="FIGURE 23" captionStartId="52.[132,211,1852,1871]" captionTargetBox="[133,1263,1197,1817]" captionTargetId="figure-469@52.[132,1266,1185,1828]" captionTargetPageId="52" captionText="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641421" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="52" pageNumber="269">Fig. 23C</figureCitation>
) tapering to apex.
</paragraph>
<caption id="A784B3E42436FFB575E76656F1EDF8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641421" ID-Zenodo-Dep="12641421" box="[132,1265,1852,1871]" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="52" pageNumber="269" startId="52.[132,211,1852,1871]" targetBox="[133,1263,1197,1817]" targetPageId="52" targetType="figure">
<paragraph id="F344E36C2436FFB575E76656F1EDF8F4" blockId="52.[132,1265,1852,1871]" box="[132,1265,1852,1871]" pageId="52" pageNumber="269">
<emphasis id="C18F3F7E2436FFB575E76656F5E9F8F4" bold="true" box="[132,245,1852,1871]" pageId="52" pageNumber="269">FIGURE 23:</emphasis>
<emphasis id="C18F3F7E2436FFB575996656F4AAF8F4" box="[250,438,1852,1871]" italics="true" pageId="52" pageNumber="269">
<taxonomicName id="34FB98EF2436FFB575996656F4ADF8F4" authorityName="Fischer von Waldheim" authorityYear="1823" box="[250,433,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="269" phylum="Arthropoda" rank="species" species="henningi">Physodactylus henningi</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2436FFB574D96656F4D0F8F4" bold="true" box="[442,460,1852,1871]" pageId="52" pageNumber="269">A,</emphasis>
sternite VIII;
<emphasis id="C18F3F7E2436FFB577216656F749F8F4" bold="true" box="[578,597,1852,1871]" pageId="52" pageNumber="269">B,</emphasis>
tergite VIII;
<emphasis id="C18F3F7E2436FFB577A16656F7CBF8F4" bold="true" box="[706,727,1852,1871]" pageId="52" pageNumber="269">C,</emphasis>
sternite IX;
<emphasis id="C18F3F7E2436FFB5765E6656F648F8F4" bold="true" box="[829,852,1852,1871]" pageId="52" pageNumber="269">D,</emphasis>
tergites IX and X;
<emphasis id="C18F3F7E2436FFB576936656F10AF8F4" bold="true" box="[1008,1046,1852,1871]" pageId="52" pageNumber="269">E, F,</emphasis>
aedeagus (ventral, dorsal).
</paragraph>
</caption>
<paragraph id="F344E36C2437FFB7753D61D2F42DFEAB" blockId="53.[94,643,184,272]" lastBlockId="54.[132,680,184,272]" lastPageId="54" lastPageNumber="271" pageId="53" pageNumber="270">
<emphasis id="C18F3F7E2437FFB4753D61D2F5ABFF6B" box="[94,183,184,208]" italics="true" pageId="53" pageNumber="270">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92437FFB475A761D2F45FFF6B" box="[196,323,184,208]" captionStart="FIGURE 23" captionStartId="52.[132,211,1852,1871]" captionTargetBox="[133,1263,1197,1817]" captionTargetId="figure-469@52.[132,1266,1185,1828]" captionTargetPageId="52" captionText="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641421" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="53" pageNumber="270">Figs. 23E, F</figureCitation>
): Phallobase 0.44-0.51x the total length of aedeagus, 1.17-1.51x longer than wide, length ratio between lateroposterior and median parts 4.10-10.0; paramere with ventral surface nearly entirely sclerotized with anteromedian margin distinct or partly membranous and translucent; penis with basal strut 0.23-0.32x its total length, articulated to the parameres by a translucent membrane; ventral sclerite elliptical.
</paragraph>
<caption id="A784B3E42437FFB4753C6656F1D2F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641423" ID-Zenodo-Dep="12641423" box="[95,1230,1852,1871]" httpUri="https://zenodo.org/record/12641423/files/figure.png" pageId="53" pageNumber="270" startId="53.[95,174,1852,1871]" targetBox="[109,1212,300,1823]" targetPageId="53" targetType="figure">
<paragraph id="F344E36C2437FFB4753C6656F1D2F8F4" blockId="53.[95,1230,1852,1871]" box="[95,1230,1852,1871]" pageId="53" pageNumber="270">
<emphasis id="C18F3F7E2437FFB4753C6656F5D3F8F4" bold="true" box="[95,207,1852,1871]" pageId="53" pageNumber="270">FIGURE 24:</emphasis>
<emphasis id="C18F3F7E2437FFB475B76656F493F8F4" box="[212,399,1852,1871]" italics="true" pageId="53" pageNumber="270">
<taxonomicName id="34FB98EF2437FFB475B76656F497F8F4" authorityName="Fischer von Waldheim" authorityYear="1823" box="[212,395,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="270" phylum="Arthropoda" rank="species" species="henningi">Physodactylus henningi</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2437FFB474F06656F4BAF8F4" bold="true" box="[403,422,1852,1871]" pageId="53" pageNumber="270">A,</emphasis>
tergite VIII;
<emphasis id="C18F3F7E2437FFB477716656F739F8F4" bold="true" box="[530,549,1852,1871]" pageId="53" pageNumber="270">B,</emphasis>
sternite VIII;
<emphasis id="C18F3F7E2437FFB477F96656F7B3F8F4" bold="true" box="[666,687,1852,1871]" pageId="53" pageNumber="270">C,</emphasis>
ovipositor (ventral);
<emphasis id="C18F3F7E2437FFB476386656F66EF8F4" bold="true" box="[859,882,1852,1871]" pageId="53" pageNumber="270">D,</emphasis>
ovipositor and reproductive tract (dorsal).
</paragraph>
</caption>
<paragraph id="F344E36C2434FFB775E76052F449FB4B" blockId="54.[132,681,312,1264]" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB775E76052F5CDFEEB" box="[132,209,312,336]" italics="true" pageId="54" pageNumber="271">Female:</emphasis>
Total length
<quantity id="34034E892434FFB774016052F4AFFEF4" box="[354,435,312,336]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="54" pageNumber="271" unit="mm" value="21.0">21 mm</quantity>
; elytral base as wide as prothorax, elytra 2.82x longer than pronotum. Dorsal and ventral surface strongly convex; pronotal punctation smaller and sparser, elytral punctation larger, abdominal pilosity shorter and finer than those of the male; frons concave along the anterior border, with a pair of oblique protuberances over antennal insertions convergent posteriorly; frontal carina not produced; antenna nearly reaching the half length of pronotum; antennomere IV-X 1.1-1.5 wider than long; XI oval, 1.08x longer than wide. Elytra with sides slightly divergent to posterior third then tapering to apex; interstices strongly convex. Hind wings 0.81x as long as elytra. Lamella of tarsomeres absent. Pregenitalic segments and ovipositor with stout brown setae. Sternite VIII (
<figureCitation id="6BC0FFE92434FFB774776272F473FC8B" box="[276,367,792,816]" captionStart="FIGURE 24" captionStartId="53.[95,174,1852,1871]" captionTargetBox="[109,1212,300,1823]" captionTargetId="figure-52@53.[94,1228,298,1828]" captionTargetPageId="53" captionText="FIGURE 24: Physodactylus henningi. A, tergite VIII; B, sternite VIII; C, ovipositor (ventral); D, ovipositor and reproductive tract (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641423" httpUri="https://zenodo.org/record/12641423/files/figure.png" pageId="54" pageNumber="271">Fig. 24B</figureCitation>
) with spiculum 0,7x as long as its total length, apical part with lateroanterior angles rounded, partly sclerotized with long and short setae lateroposteriorly, tergite VIII (
<figureCitation id="6BC0FFE92434FFB774BC6212F727FC2B" box="[479,571,888,912]" captionStart="FIGURE 24" captionStartId="53.[95,174,1852,1871]" captionTargetBox="[109,1212,300,1823]" captionTargetId="figure-52@53.[94,1228,298,1828]" captionTargetPageId="53" captionText="FIGURE 24: Physodactylus henningi. A, tergite VIII; B, sternite VIII; C, ovipositor (ventral); D, ovipositor and reproductive tract (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641423" httpUri="https://zenodo.org/record/12641423/files/figure.png" pageId="54" pageNumber="271">Fig. 24A</figureCitation>
) semioval, evenly sclerotized with short seta lateroposteriorly, anterior sclerotized margin emarginate. Ovipositor (
<figureCitation id="6BC0FFE92434FFB775EF62B2F400FC4B" box="[140,284,984,1008]" captionStart="FIGURE 24" captionStartId="53.[95,174,1852,1871]" captionTargetBox="[109,1212,300,1823]" captionTargetId="figure-52@53.[94,1228,298,1828]" captionTargetPageId="53" captionText="FIGURE 24: Physodactylus henningi. A, tergite VIII; B, sternite VIII; C, ovipositor (ventral); D, ovipositor and reproductive tract (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641423" httpUri="https://zenodo.org/record/12641423/files/figure.png" pageId="54" pageNumber="271">Figs. 24C, D</figureCitation>
) with coxites partly sclerotized with short setae lateroapically, without styli, baculi 3.3x longer than coxites. Reproductive tract (
<figureCitation id="6BC0FFE92434FFB777556572F780FB8B" box="[566,668,1048,1072]" captionStart="FIGURE 24" captionStartId="53.[95,174,1852,1871]" captionTargetBox="[109,1212,300,1823]" captionTargetId="figure-52@53.[94,1228,298,1828]" captionTargetPageId="53" captionText="FIGURE 24: Physodactylus henningi. A, tergite VIII; B, sternite VIII; C, ovipositor (ventral); D, ovipositor and reproductive tract (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641423" httpUri="https://zenodo.org/record/12641423/files/figure.png" pageId="54" pageNumber="271">Fig. 24D</figureCitation>
): bursa copulatrix oval with a row of spines laterally, with a slender diverticulum entering the posterior part of bursa copulatrix; a pair of sac-shaped structures posteriad of bursa copulatrix (possibly colleterial glands), lubricating glands sac-shaped near lateroposterior part of baculi.
</paragraph>
<paragraph id="F344E36C2434FFB775E76472F5EAFA2B" blockId="54.[132,681,1304,1424]" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB775E76472F457FA8B" box="[132,331,1304,1328]" italics="true" pageId="54" pageNumber="271">Material examined:</emphasis>
<materialsCitation id="4393E9312434FFB774356472F5B9FAEB" collectionCode="MNHN" country="Brazil" location="Bahia" pageId="54" pageNumber="271" specimenCount="8" stateProvince="Bahia">
<collectingCountry id="8BECA3FC2434FFB774356472F4A8FA8B" box="[342,436,1304,1328]" name="Brazil" pageId="54" pageNumber="271">BRAZIL</collectingCountry>
,
<specimenCount id="E5FD28E52434FFB774A16472F71CFA94" box="[450,512,1304,1328]" count="8" pageId="54" pageNumber="271" type="generic">8 exs.</specimenCount>
(
<collectionCode id="95EA7BA92434FFB777716472F772FA8B" box="[530,622,1304,1328]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="271">MNHN</collectionCode>
);
<collectingRegion id="313F2D8E2434FFB777E66473F5B9FAEB" country="Brazil" name="Bahia" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB777E66473F5B9FAEB" italics="true" pageId="54" pageNumber="271">Bahia</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="C18F3F7E2434FFB775C66452F5B7FAEB" box="[165,171,1336,1360]" italics="true" pageId="54" pageNumber="271">:</emphasis>
<materialsCitation id="4393E9312434FFB775D16452F441FAEB" box="[178,349,1336,1360]" collectionCode="MNHN" country="Brazil" pageId="54" pageNumber="271" specimenCount="3">
<specimenCount id="E5FD28E52434FFB775D16452F5F0FAF4" box="[178,236,1336,1360]" count="3" pageId="54" pageNumber="271" type="generic">3 exs.</specimenCount>
(
<collectionCode id="95EA7BA92434FFB775996452F44AFAEB" box="[250,342,1336,1360]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="271">MNHN</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="4393E9312434FFB7740B6452F7BFFAF4" box="[360,675,1336,1360]" collectionCode="MZUSP" country="Brazil" location="Espirito Santo" pageId="54" pageNumber="271" specimenCount="1" stateProvince="Espirito Santo">
<specimenCount id="E5FD28E52434FFB7740B6452F486FAF4" box="[360,410,1336,1360]" count="1" pageId="54" pageNumber="271" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA92434FFB774C46452F719FAEB" box="[423,517,1336,1360]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="54" pageNumber="271">MZUSP</collectionCode>
);
<collectingRegion id="313F2D8E2434FFB777746452F7BFFAF4" box="[535,675,1336,1359]" country="Brazil" name="Espirito Santo" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB777746452F7BFFAF4" box="[535,675,1336,1359]" italics="true" pageId="54" pageNumber="271">Espírito Santo</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="C18F3F7E2434FFB777C06452F7B4FAF4" box="[675,680,1336,1359]" italics="true" pageId="54" pageNumber="271">:</emphasis>
<materialsCitation id="4393E9312434FFB775E76432F439FACB" box="[132,293,1368,1392]" collectionCode="DZUP" country="Brazil" pageId="54" pageNumber="271" specimenCount="7">
<specimenCount id="E5FD28E52434FFB775E76432F5DDFAD4" box="[132,193,1368,1391]" count="7" pageId="54" pageNumber="271" type="generic">7 exs.</specimenCount>
(
<collectionCode id="95EA7BA92434FFB775B36432F401FACB" box="[208,285,1368,1392]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="54" pageNumber="271" type="University or college">DZUP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="4393E9312434FFB774516432F4C3FACB" box="[306,479,1368,1392]" collectionCode="MNRJ" country="Brazil" pageId="54" pageNumber="271" specimenCount="10">
<specimenCount id="E5FD28E52434FFB774516432F460FAD4" box="[306,380,1368,1392]" count="10" pageId="54" pageNumber="271" type="generic">10 exs.</specimenCount>
(
<collectionCode id="95EA7BA92434FFB774E86432F4CBFACB" box="[395,471,1368,1392]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="54" pageNumber="271">MNRJ</collectionCode>
)
</materialsCitation>
;
<specimenCount id="E5FD28E52434FFB7748F6432F72AFAD4" box="[492,566,1368,1392]" count="24" pageId="54" pageNumber="271" type="generic">24 exs.</specimenCount>
(
<specimenCount id="E5FD28E52434FFB7772B6432F7BEFACB" box="[584,674,1368,1392]" count="1" pageId="54" pageNumber="271" type="female">1 female</specimenCount>
) (MZUSP).
</paragraph>
<paragraph id="F344E36C2434FFB775E764D2F5C0F9AB" blockId="54.[132,681,1464,1552]" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB775E764D2F41AFA6B" box="[132,262,1464,1488]" italics="true" pageId="54" pageNumber="271">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2434FFB7747164D2F46DFA6B" box="[274,369,1464,1488]" name="Brazil" pageId="54" pageNumber="271">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2434FFB774E364D2F779FA74" box="[384,613,1464,1488]" italics="true" pageId="54" pageNumber="271">
<collectingRegion id="313F2D8E2434FFB774E364D2F4A0FA6B" box="[384,444,1464,1488]" country="Brazil" name="Bahia" pageId="54" pageNumber="271">Bahia</collectingRegion>
;
<collectingRegion id="313F2D8E2434FFB774AC64D2F77CFA74" box="[463,608,1464,1487]" country="Brazil" name="Espirito Santo" pageId="54" pageNumber="271">Espírito Santo</collectingRegion>
:
</emphasis>
Conceição da Barra, Barra do São Francisco, Linhares, Aracruz.
</paragraph>
<paragraph id="F344E36C2434FFB775E76752F601FDEB" blockId="54.[132,681,1592,1872]" lastBlockId="54.[718,1269,184,1616]" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB775E76752F5FCF9EB" box="[132,224,1592,1616]" italics="true" pageId="54" pageNumber="271">Remarks:</emphasis>
The
<typeStatus id="2C405DCE2434FFB7744A6752F499F9EB" box="[297,389,1592,1616]" pageId="54" pageNumber="271" type="holotype">holotype</typeStatus>
of
<taxonomicName id="34FB98EF2434FFB774DA6752F7B4F9EB" authorityName="Fischer von Waldheim" authorityYear="1823" box="[441,680,1592,1616]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2434FFB774DA6752F7B4F9EB" box="[441,680,1592,1616]" italics="true" pageId="54" pageNumber="271">Physodactylus henningi</emphasis>
</taxonomicName>
was supposed to be in the Zoological Museum of the Moscow State University (ZMMU), but it was neither found there nor in the German Collections of the Deutsches Entomologisches Institut (DEI, Dresden) and Zoologisches Staatssammlung (ZSM, Munich). There is no evidence that Fleutiaux compared the
<typeStatus id="2C405DCE2434FFB7758C6672F457F88B" box="[239,331,1816,1840]" pageId="54" pageNumber="271" type="holotype">holotype</typeStatus>
with the specimens he identified as
<emphasis id="C18F3F7E2434FFB775FD6653F409F8EB" box="[158,277,1848,1872]" italics="true" pageId="54" pageNumber="271">
<taxonomicName id="34FB98EF2434FFB775FD6653F40EF8EB" authorityName="Fischer von Waldheim" authorityYear="1823" box="[158,274,1848,1872]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henningi">P. henningi</taxonomicName>
,
</emphasis>
on which he based its revisions (1892, 1940c). The confirmation of the identity of these specimens and remaining material examined herein was based on the description and illustrations provided by
<bibRefCitation id="976A9E9D2434FFB776566072F167FE8B" author="FISCHER VON WALDHEIM, M. G." box="[821,1147,280,304]" pageId="54" pageNumber="271" pagination="448 - 451" refId="ref49654" refString="FISCHER VON WALDHEIM, M. G. 1824. Extrait d'une lettre aadressee a M. Henning sur le Physodactyle, nouveaux genre de Coleoptere voisin des Taupin. Annales des Sciences Naturelles, 3: 448 - 451, plate 27 B." type="journal article" year="1824">Fischer von Waldheim (1824)</bibRefCitation>
. The combination of size (
<quantity id="34034E892434FFB776E36052F6CCFEEB" box="[896,976,312,336]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="54" pageNumber="271" unit="mm" value="18.0">18 mm</quantity>
) to the general appearance, shape of antenna and tarsal lamellae of the specimen illustrated in this work is unique among the
<taxonomicName id="34FB98EF2434FFB771D16012F673FE0B" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB771D16012F601FE0B" italics="true" pageId="54" pageNumber="271">Physodactylus</emphasis>
species
</taxonomicName>
examined. The lamella of the protarsomere I of the specimens examined is usually a little narrower than those of the tarsomeres II and III, only a few specimens have it nearly as wide as the other ones as illustrated by
<bibRefCitation id="976A9E9D2434FFB771636372F605FDEB" author="FISCHER VON WALDHEIM, M. G." pageId="54" pageNumber="271" pagination="448 - 451" refId="ref49654" refString="FISCHER VON WALDHEIM, M. G. 1824. Extrait d'une lettre aadressee a M. Henning sur le Physodactyle, nouveaux genre de Coleoptere voisin des Taupin. Annales des Sciences Naturelles, 3: 448 - 451, plate 27 B." type="journal article" year="1824">Fischer von Waldheim (1824)</bibRefCitation>
.
</paragraph>
<paragraph id="F344E36C2434FFB776646332F6B6FCEB" blockId="54.[718,1269,184,1616]" pageId="54" pageNumber="271">
I have examined only photos of the dorsal habitus of
<taxonomicName id="34FB98EF2434FFB7762B6313F11AFD2B" authorityName="Perty" authorityYear="1830" box="[840,1030,632,656]" class="Insecta" family="Elateridae" genus="Drepanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="C18F3F7E2434FFB7762B6313F11AFD2B" box="[840,1030,632,656]" italics="true" pageId="54" pageNumber="271">Drepanius clavipes</emphasis>
</taxonomicName>
<typeStatus id="2C405DCE2434FFB771706312F15EFD2B" box="[1043,1090,632,656]" pageId="54" pageNumber="271">type</typeStatus>
, whose antenna illustrated by
<bibRefCitation id="976A9E9D2434FFB7760163F2F6F4FD0B" author="PERTY, J. A. M." box="[866,1000,664,688]" pageId="54" pageNumber="271" pagination="1 - 12" refId="ref50792" refString="PERTY, J. A. M. 1830. Insecta Brasiliensia. In: Spix, J. &amp; Martius, C. Delectus animalium articulatorum, quae in itinere per Brasiliam annis 1817 - 1820 jussu et auspiciis Maximilliani Josephi I Bavariae regis augustissimi peracto. Munchen, Frid. Fleischer. p. 1 - 60, pls. 1 - 12." type="book chapter" year="1830">Perty (1830)</bibRefCitation>
is different from that of
<emphasis id="C18F3F7E2434FFB777AD63D3F658FD6B" box="[718,836,696,720]" italics="true" pageId="54" pageNumber="271">
<taxonomicName id="34FB98EF2434FFB777AD63D3F65CFD6B" authorityName="Fischer von Waldheim" authorityYear="1823" box="[718,832,696,720]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henningi">P. henningi</taxonomicName>
.
</emphasis>
However, the dorsal view of the specimen deposited at ZSM (Munich) were insufficient to confirm its identity. For this reason I maintain
<taxonomicName id="34FB98EF2434FFB771E46393F1EEFCAB" authorityName="Perty" authorityYear="1830" box="[1159,1266,760,784]" class="Insecta" family="Elateridae" genus="Drepanius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="C18F3F7E2434FFB771E46393F1EEFCAB" box="[1159,1266,760,784]" italics="true" pageId="54" pageNumber="271">D. clavipes</emphasis>
</taxonomicName>
as synonym of
<emphasis id="C18F3F7E2434FFB776116273F6F6FC8B" box="[882,1002,792,816]" italics="true" pageId="54" pageNumber="271">
<taxonomicName id="34FB98EF2434FFB776116273F6FAFC8B" authorityName="Fischer von Waldheim" authorityYear="1823" box="[882,998,792,816]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henningi">P. henningi</taxonomicName>
,
</emphasis>
although this synonymy should be confirmed.
</paragraph>
<paragraph id="F344E36C2434FFB776646232F6E7F9EB" blockId="54.[718,1269,184,1616]" pageId="54" pageNumber="271">
<taxonomicName id="34FB98EF2434FFB776646232F6F6FCCB" authorityName="Fischer von Waldheim" authorityYear="1823" box="[775,1002,856,880]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2434FFB776646232F6F6FCCB" box="[775,1002,856,880]" italics="true" pageId="54" pageNumber="271">Physodactylus henningi</emphasis>
</taxonomicName>
specimens from Linhares present the color pattern 1, while those from
<collectingRegion id="313F2D8E2434FFB771DB6212F1EEFC2B" box="[1208,1266,888,912]" country="Brazil" name="Bahia" pageId="54" pageNumber="271">Bahia</collectingRegion>
present the color patterns 1 and 2; specimens identified by Fleutiaux (
<emphasis id="C18F3F7E2434FFB776F662D3F6C4FC6B" box="[917,984,952,976]" italics="true" pageId="54" pageNumber="271">in litt.</emphasis>
) as
<taxonomicName id="34FB98EF2434FFB7716B62D3F166FC6B" authorityName="Fischer von Waldheim" authorityYear="1823" box="[1032,1146,952,976]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2434FFB7716B62D3F166FC6B" box="[1032,1146,952,976]" italics="true" pageId="54" pageNumber="271">P. henningi</emphasis>
</taxonomicName>
<taxonomicName id="34FB98EF2434FFB771E762D2F63AFC4B" authorityName="Fleutiaux" authorityYear="1892" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="variety" species="henningi" variety="erythrocephalus">v. erythrocephalus</taxonomicName>
without detailed locality have the pattern color 3. The specimens from Aracruz and Conceição da Barra are entirely black (color pattern 5). The specimens described by Fleutiaux (1940;
<emphasis id="C18F3F7E2434FFB771EA6553F1D0FBEB" box="[1161,1228,1080,1104]" italics="true" pageId="54" pageNumber="271">in litt.</emphasis>
) as
<taxonomicName id="34FB98EF2434FFB777AD6533F65CFBCB" authorityName="Fischer von Waldheim" authorityYear="1823" box="[718,832,1112,1136]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2434FFB777AD6533F65CFBCB" box="[718,832,1112,1136]" italics="true" pageId="54" pageNumber="271">P. henningi</emphasis>
</taxonomicName>
<taxonomicName id="34FB98EF2434FFB776286532F6C0FBCB" authorityName="Fleutiaux" authorityYear="1940" box="[843,988,1112,1136]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="variety" species="henningi" variety="puncticollis">v. puncticollis</taxonomicName>
(
<collectingRegion id="313F2D8E2434FFB7768D6532F137FBCB" box="[1006,1067,1112,1136]" country="Brazil" name="Bahia" pageId="54" pageNumber="271">Bahia</collectingRegion>
) and
<taxonomicName id="34FB98EF2434FFB7710E6533F1C4FBCB" authorityName="Fischer von Waldheim" authorityYear="1823" box="[1133,1240,1112,1136]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henning">
<emphasis id="C18F3F7E2434FFB7710E6533F1C4FBCB" box="[1133,1240,1112,1136]" italics="true" pageId="54" pageNumber="271">P. henning</emphasis>
</taxonomicName>
<taxonomicName id="34FB98EF2434FFB771816532F62FFB2B" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="variety" species="henningi" variety="nigricollis">v. nigricollis</taxonomicName>
(“
<collectingCountry id="8BECA3FC2434FFB7762F6512F697FB2B" box="[844,907,1144,1168]" name="Brazil" pageId="54" pageNumber="271">Brèsil</collectingCountry>
”) are nearly entirely black, except for the pronotum partly dark brown. The specimens from Barra do São Francisco (“Córrego Itá”) present the color patterns 4-6. There are also small variations in size and density of punctation and in the shape of phallobase, nevertheless those differences were found among individuals from same region. On the other hand, all specimens present the following combination of characters: pilosity dark brown, frons with a pair of oblique protuberances over antennal insertions, lateral carina of pronotum complete, first abdominal ventrite nearly entirely smooth and tarsal lamellae as wide as tarsomeres including its setae. These characters clearly distinguish
<taxonomicName id="34FB98EF2434FFB7769E6773F171F98B" authorityName="Fischer von Waldheim" authorityYear="1823" box="[1021,1133,1560,1584]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2434FFB7769E6773F171F98B" box="[1021,1133,1560,1584]" italics="true" pageId="54" pageNumber="271">P. henningi</emphasis>
</taxonomicName>
from the remaining species of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,222 @@
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<paragraph id="F344E36C2432FFB175C363FDF790FD0B" blockId="48.[160,652,663,720]" box="[160,652,663,688]" pageId="48" pageNumber="265">
<heading id="A80C54002432FFB175C363FDF790FD0B" bold="true" box="[160,652,663,688]" centered="true" fontSize="10" level="2" pageId="48" pageNumber="265" reason="2">
<taxonomicName id="34FB98EF2432FFB175C363FDF790FD0B" authority="Fleutiaux, 1892" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[160,652,663,688]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">
<emphasis id="C18F3F7E2432FFB175C363FDF790FD0B" bold="true" box="[160,652,663,688]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB175C363FDF4C0FD14" bold="true" box="[160,476,663,687]" italics="true" pageId="48" pageNumber="265">Physodactylus foveatostriatus</emphasis>
<bibRefCitation id="976A9E9D2432FFB1748163F2F790FD0B" author="FLEUTIAUX, E." box="[482,652,664,688]" pageId="48" pageNumber="265" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72432FFB1744F63D2F71CFD6B" box="[300,512,696,720]" pageId="48" pageNumber="265" type="description">
<paragraph id="F344E36C2432FFB1744F63D2F71CFD6B" blockId="48.[160,652,663,720]" box="[300,512,696,720]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB1744F63D2F71CFD6B" bold="true" box="[300,512,696,720]" pageId="48" pageNumber="265">
(
<figureCitation id="6BC0FFE92432FFB1745763D2F496FD6B" box="[308,394,696,720]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="48" pageNumber="265">Figs. 20</figureCitation>
H-L, 35H)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72432FFB175E76392F4F0FCEB" pageId="48" pageNumber="265" type="reference_group">
<paragraph id="F344E36C2432FFB175E76392F4F0FCEB" blockId="48.[132,680,760,848]" pageId="48" pageNumber="265">
<treatmentCitationGroup id="D3EBC4422432FFB175E76392F4F0FCEB" pageId="48" pageNumber="265">
<treatmentCitation id="725AC57D2432FFB175E76392F7B9FCAB" author="FLEUTIAUX, E." box="[132,677,760,784]" page="406" pageId="48" pageNumber="265" year="1892">
<taxonomicName id="34FB98EF2432FFB175E76392F7B9FCAB" authority="Fleutiaux, 1892: 406" authorityName="Fleutiaux, J. Chassain" authorityPageNumber="406" authorityYear="1892" box="[132,677,760,784]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">
<emphasis id="C18F3F7E2432FFB175E76392F4B9FCAB" box="[132,421,760,784]" italics="true" pageId="48" pageNumber="265">Physodactylus foveatostriatus</emphasis>
<bibRefCitation id="976A9E9D2432FFB174D66392F7B9FCAB" author="FLEUTIAUX, E." box="[437,677,760,784]" pageId="48" pageNumber="265" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2432FFB175DE6272F480FC8B" author="SCHWARZ, O." box="[189,412,792,816]" page="311" pageId="48" pageNumber="265" year="1906">
<bibRefCitation id="976A9E9D2432FFB175DE6272F480FC8B" author="SCHWARZ, O." box="[189,412,792,816]" pageId="48" pageNumber="265" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 311</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2432FFB174CC6272F7B8FC8B" author="FLEUTIAUX, E." box="[431,676,792,816]" page="164" pageId="48" pageNumber="265" year="1940">
<bibRefCitation id="976A9E9D2432FFB174CC6272F7B8FC8B" author="FLEUTIAUX, E." box="[431,676,792,816]" pageId="48" pageNumber="265" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 164</bibRefCitation>
</treatmentCitation>
, 166;
<treatmentCitation id="725AC57D2432FFB175916252F4F4FCEB" author="BLACKWELDER, R. E." box="[242,488,824,848]" page="292" pageId="48" pageNumber="265" year="1944">
<bibRefCitation id="976A9E9D2432FFB175916252F4F4FCEB" author="BLACKWELDER, R. E." box="[242,488,824,848]" pageId="48" pageNumber="265" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72432FFB175E76212F188F8EB" pageId="48" pageNumber="265" type="description">
<paragraph id="F344E36C2432FFB175E76212F6A0FE0B" blockId="48.[132,681,888,1872]" lastBlockId="48.[718,1266,184,432]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB175E76212F415FC2B" box="[132,265,888,912]" italics="true" pageId="48" pageNumber="265">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE92432FFB1743C6212F4DAFC2B" box="[351,454,888,912]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="48" pageNumber="265">Fig. 35H</figureCitation>
): Integument bright; dark brown to black with prothorax orange to reddish-brown; legs colored as elytra or with femur reddish-brown; pilosity brown. Total length 11.5-16.0 mm; elytral base 1.0-1.03x as wide as prothorax, elytra 2.7-3.0x longer than pronotum. Frons concave, disc of anterior part of head strongly convex, frontal carina weakly impressed; antenna (
<figureCitation id="6BC0FFE92432FFB175EF6512F5E2FB2B" box="[140,254,1144,1168]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="48" pageNumber="265">Figs. 20H</figureCitation>
,
<figureCitation id="6BC0FFE92432FFB174736512F422FB2B" box="[272,318,1144,1168]" captionStart="FIGURE 23" captionStartId="52.[132,211,1852,1871]" captionTargetBox="[133,1263,1197,1817]" captionTargetId="figure-469@52.[132,1266,1185,1828]" captionTargetPageId="52" captionText="FIGURE 23: Physodactylus henningi. A, sternite VIII; B, tergite VIII; C, sternite IX; D, tergites IX and X; E, F, aedeagus (ventral, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641421" httpUri="https://zenodo.org/record/12641421/files/figure.png" pageId="48" pageNumber="265">23A</figureCitation>
) reaching the posterior third of pronotum; antennomere IV 1.22x wider than long, V 1.47x wider than long, VI-X 1.25-1.37x wider than long; XI oval 1.3x longer than wide. Pronotum (
<figureCitation id="6BC0FFE92432FFB175EF6592F5F8FAAB" box="[140,228,1272,1296]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="48" pageNumber="265">Fig. 20I</figureCitation>
) 1.23-1.34x wider than long, lateral sides nearly straight to rounded convergent from posterior angles to anterior margin; anteromedian margin weakly produced; posterior angles elongate, convex, parallel; lateral carina complete weakly impressed, dorsally visible only on posterior half; disc with punctures 2-3 diameters apart, a little larger and denser on lateral and anterior borders; prosternal process (
<figureCitation id="6BC0FFE92432FFB175826492F424F9AB" box="[225,312,1528,1552]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="48" pageNumber="265">Fig. 20J</figureCitation>
) with a subapical tooth. Lamellae of pro- and mesotarsomeres I a little smaller than the others, present on metatarsomere I. Elytra parallel-sided on anterior 2/3 then tapering to apex, apex usually dehiscent; striae with a single row of punctures; interstices flattened, scarcely punctate, weakly rugose; apical striae with punctures about 2 times larger than those of pronotal lateral border. Abdomen with ventrites 2-3 slightly concave laterally, ventrite 1 punctate, covered with fine setae about 1/5 as long as those of ventrite 2, setae longer and sparser on marginal plates; ventrites 2-4 with setae moderately long, decumbent, sparser on marginal plates. Pregenitalic segments and aedeagus covered with stout dark-brown setae. Sternite VIII with lateroposterior sclerotizations separate or approximated medially; posterior margin straight, anterior sclerotization 0.3-0.4x the total width of sternite; sternite IX with apex rounded.
</paragraph>
<paragraph id="F344E36C2432FFB177AD60B2F1A5FD4B" blockId="48.[718,1267,472,752]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB177AD60B2F63BFE4B" box="[718,807,472,496]" italics="true" pageId="48" pageNumber="265">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92432FFB1765860B2F6D7FE4B" box="[827,971,472,496]" captionStart="FIGURE 20" captionStartId="46.[134,213,1826,1845]" captionTargetBox="[135,1266,706,1801]" captionTargetId="figure-235@46.[132,1266,706,1802]" captionTargetPageId="46" captionText="FIGURE 20: Physodactylus flavifrons sp. nov. A, antenna; B, maxilla; C, D, E, prothorax (dorsal, lateral ventral); F, G, aedeagus (dorsal, ventral). Physodactylus foveatostriatus. H, antenna; I, J, prothorax (dorsal, lateral); K, L, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641413" httpUri="https://zenodo.org/record/12641413/files/figure.png" pageId="48" pageNumber="265">Figs. 20K, L</figureCitation>
): Phallobase 0.4x the total length of aedeagus, 0.9-1.0 as long as wide, length ratio between lateroposterior and median parts 11.8-15.0; paramere with ventral surface weakly sclerotized, anteromedian margin with a linear sclerotization directed anteriad; penis with basal strut 0.2x times its total length, articulated to the parameres by a sclerotized short process or by a translucent membrane; ventral sclerite elongate, widened apicad.
</paragraph>
<paragraph id="F344E36C2432FFB177AD6272F1BAFC4B" blockId="48.[718,1266,792,1008]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB177AD6272F637FC8B" box="[718,811,792,816]" italics="true" pageId="48" pageNumber="265">
<typeStatus id="2C405DCE2432FFB177AD6272F63AFC8B" box="[718,806,792,816]" pageId="48" pageNumber="265" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<taxonomicName id="34FB98EF2432FFB1765B6272F1EEFC8B" authority="Chv, Brasilia" authorityName="Chv, Brasilia" box="[824,1266,792,816]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">Physodactylus foveatostriatus Chv, Brasilia</taxonomicName>
ex. mus. Castelnau, Collection Chevrolat], [Type], [Collection Chevrolat], [
<taxonomicName id="34FB98EF2432FFB176B76232F7F0FC34" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">Physodactylus foveatostriatus</taxonomicName>
J, (Chv.) Fleuti. Type,
<collectingCountry id="8BECA3FC2432FFB176906212F12EFC2B" box="[1011,1074,888,912]" name="Brazil" pageId="48" pageNumber="265">Brèsil</collectingCountry>
], [
<taxonomicName id="34FB98EF2432FFB1712E6213F16BFC0B" authority="Fleut. Ann. Soc. Ent. Fr. 1892 p. 406" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">
<emphasis id="C18F3F7E2432FFB1712E6213F1EEFC2B" box="[1101,1266,888,912]" italics="true" pageId="48" pageNumber="265">P. foveatostriatus</emphasis>
Fleut. Ann. Soc. Ent. Fr. 1892 p. 406
</taxonomicName>
, Collection FLEUTIAUX], [TYPE], [
<typeStatus id="2C405DCE2432FFB176B962D2F15BFC6B" box="[986,1095,952,976]" pageId="48" pageNumber="265" type="lectotype">Lectotype</typeStatus>
], [
<taxonomicName id="34FB98EF2432FFB1713D62D2F154FC4B" authority="Fleutiaux, J. Chassain" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">Physodactylus foveatostriatus Fleutiaux, J. Chassain</taxonomicName>
det. 05].
</paragraph>
<paragraph id="F344E36C2432FFB177AD6573F622FA2B" blockId="48.[718,1267,1048,1424]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB177AD6573F62CFB8B" box="[718,816,1049,1072]" italics="true" pageId="48" pageNumber="265">
<typeStatus id="2C405DCE2432FFB177AD6573F636FB8B" box="[718,810,1049,1072]" pageId="48" pageNumber="265" type="paratype">Paratypes</typeStatus>
:
</emphasis>
[nov. spec.], [Chevrolat], [Collection Chevrolat], [
<taxonomicName id="34FB98EF2432FFB176796553F606FBCB" authority="Fleut. Ann. Soc. Ent. Fr. 1892 p. 406" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">
<emphasis id="C18F3F7E2432FFB176796553F6A7FBEB" box="[794,955,1080,1104]" italics="true" pageId="48" pageNumber="265">P. foveatostriatus</emphasis>
Fleut. Ann. Soc. Ent. Fr. 1892 p. 406
</taxonomicName>
, Collection FLEUTIAUX], [
<typeStatus id="2C405DCE2432FFB171346532F1F4FBCB" box="[1111,1256,1112,1136]" pageId="48" pageNumber="265" type="paralectotype">Paralectotype</typeStatus>
], [
<taxonomicName id="34FB98EF2432FFB177B76512F1EEFB2B" authority="Fleutiaux, J. Chassain" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[724,1266,1144,1168]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">Physodactylus foveatostriatus Fleutiaux, J. Chassain</taxonomicName>
det. 05],
<specimenCount id="E5FD28E52432FFB1765165F2F67BFB14" box="[818,871,1176,1200]" count="1" pageId="48" pageNumber="265" type="generic">1 ex.</specimenCount>
(MNHN); idem plus [
<collectingRegion id="313F2D8E2432FFB1710265F2F1BEFB0B" box="[1121,1186,1176,1200]" country="Brazil" name="Bahia" pageId="48" pageNumber="265">Bahia</collectingRegion>
], [Coll. Fl. 335], [Muséum Paris, Collection E. Fleutiaux],
<specimenCount id="E5FD28E52432FFB177AD65B2F61DFB54" box="[718,769,1240,1264]" count="1" pageId="48" pageNumber="265" type="generic">1 ex.</specimenCount>
(MNHN); [
<collectingRegion id="313F2D8E2432FFB176E565B2F6DBFB4B" box="[902,967,1240,1264]" country="Brazil" name="Bahia" pageId="48" pageNumber="265">Bahia</collectingRegion>
], [Collection Chevrolat], [ex mus. Castelnau], [Chevrolat], [Henningi?], [
<taxonomicName id="34FB98EF2432FFB171F96593F183FA8B" authority="Fleut. Ann. Soc. Ent. Fr. 1892 p. 406" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">
<emphasis id="C18F3F7E2432FFB171F96593F63CFA8B" italics="true" pageId="48" pageNumber="265">P. foveatostriatus</emphasis>
Fleut. Ann. Soc. Ent. Fr. 1892 p. 406
</taxonomicName>
, Collection FLEUTIAUX], [
<typeStatus id="2C405DCE2432FFB176D06452F158FAEB" box="[947,1092,1336,1360]" pageId="48" pageNumber="265" type="paralectotype">Paralectotype</typeStatus>
], [
<taxonomicName id="34FB98EF2432FFB1713D6452F14DFACB" authority="Fleutiaux, J. Chassain" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">Physodactylus foveatostriatus Fleutiaux, J. Chassain</taxonomicName>
det. 05],
<specimenCount id="E5FD28E52432FFB171DD6432F1EFFAD4" box="[1214,1267,1368,1392]" count="1" pageId="48" pageNumber="265" type="generic">1 ex.</specimenCount>
(MNHN).
</paragraph>
<paragraph id="F344E36C2432FFB177AD64D2F662FA4B" blockId="48.[718,1266,1464,1520]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB177AD64D2F693FA6B" box="[718,911,1464,1488]" italics="true" pageId="48" pageNumber="265">Material examined:</emphasis>
<materialsCitation id="4393E9312432FFB176F664D2F1F0FA6B" box="[917,1260,1464,1488]" collectionCode="MNHN" country="Brazil" location="Bahia" pageId="48" pageNumber="265" specimenCount="2" stateProvince="Bahia">
<collectingCountry id="8BECA3FC2432FFB176F664D2F6EFFA6B" box="[917,1011,1464,1488]" name="Brazil" pageId="48" pageNumber="265">BRAZIL</collectingCountry>
,
<specimenCount id="E5FD28E52432FFB1769E64D2F12AFA74" box="[1021,1078,1464,1488]" count="2" pageId="48" pageNumber="265" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA92432FFB1712064D2F183FA6B" box="[1091,1183,1464,1488]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="48" pageNumber="265">MNHN</collectionCode>
);
<collectingRegion id="313F2D8E2432FFB171D364D2F1F0FA6B" box="[1200,1260,1464,1488]" country="Brazil" name="Bahia" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB171D364D2F1F0FA6B" box="[1200,1260,1464,1488]" italics="true" pageId="48" pageNumber="265">Bahia</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="C18F3F7E2432FFB1718E64D2F1EEFA6B" box="[1261,1266,1464,1488]" italics="true" pageId="48" pageNumber="265">:</emphasis>
<materialsCitation id="4393E9312432FFB177AD64B2F666FA4B" box="[718,890,1496,1520]" collectionCode="MNHN" country="Brazil" pageId="48" pageNumber="265" specimenCount="2">
<specimenCount id="E5FD28E52432FFB177AD64B2F614FA54" box="[718,776,1496,1520]" count="2" pageId="48" pageNumber="265" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA92432FFB1767564B2F66EFA4B" box="[790,882,1496,1520]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="48" pageNumber="265">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C2432FFB177AD6772F11DF98B" blockId="48.[718,1025,1560,1584]" box="[718,1025,1560,1584]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB177AD6772F64CF98B" box="[718,848,1560,1584]" italics="true" pageId="48" pageNumber="265">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2432FFB176356772F6A9F98B" box="[854,949,1560,1584]" name="Brazil" pageId="48" pageNumber="265">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2432FFB176DC6772F11DF98B" box="[959,1025,1560,1584]" italics="true" pageId="48" pageNumber="265">
<collectingRegion id="313F2D8E2432FFB176DC6772F6E2F98B" box="[959,1022,1560,1584]" country="Brazil" name="Bahia" pageId="48" pageNumber="265">Bahia</collectingRegion>
.
</emphasis>
</paragraph>
<paragraph id="F344E36C2432FFB177AD6732F188F8EB" blockId="48.[718,1266,1624,1872]" pageId="48" pageNumber="265">
<emphasis id="C18F3F7E2432FFB177AD6732F155F9CB" box="[718,1097,1624,1648]" italics="true" pageId="48" pageNumber="265">
Remarks:
<taxonomicName id="34FB98EF2432FFB176526732F155F9CB" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[817,1097,1624,1648]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="foveatostriatus">Physodactylus foveatostriatus</taxonomicName>
</emphasis>
is distinguished by its large punctures of the elytral striae, a unique character among
<taxonomicName id="34FB98EF2432FFB176E567F2F17EF90B" box="[902,1122,1688,1712]" class="Insecta" family="Elateridae" genus="Physodactylus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C18F3F7E2432FFB176E567F2F116F90B" box="[902,1034,1688,1712]" italics="true" pageId="48" pageNumber="265">Physodactylus</emphasis>
species.
</taxonomicName>
This autapomorphy combined with the shape of elytra, more tapered and dehiscent at apex, the abdominal pilosity pattern and the shorter phallobase easily separate this species from the morphologically closest species, which are
<emphasis id="C18F3F7E2432FFB1765B6653F6F6F8EB" box="[824,1002,1848,1872]" italics="true" pageId="48" pageNumber="265">
<taxonomicName id="34FB98EF2432FFB1765B6653F68BF8EB" box="[824,919,1848,1872]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="fischeri">P. fischeri</taxonomicName>
,
<taxonomicName id="34FB98EF2432FFB176C26653F6F6F8EB" authorityName="Rosa" authorityYear="2014" box="[929,1002,1849,1872]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="asper">P. asper</taxonomicName>
</emphasis>
and
<emphasis id="C18F3F7E2432FFB1717E6653F188F8EB" box="[1053,1172,1848,1872]" italics="true" pageId="48" pageNumber="265">
<taxonomicName id="34FB98EF2432FFB1717E6653F18CF8EB" authorityName="Rosa" authorityYear="2014" box="[1053,1168,1848,1872]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="265" phylum="Arthropoda" rank="species" species="chassaini">P. chassaini</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,182 @@
<document id="EEF4C89A666774FAEF1E20BDE958923B" ID-DOI="10.1590/0031-1049.2014.54.18" ID-ISSN="1807-0205" ID-Zenodo-Dep="12641367" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720058375023" checkinUser="felipe" docAuthor="Rosa, Simone Policena" docDate="2014" docId="7B52527A2433FFB375B361DDF440FACB" docLanguage="en" docName="PapAvZool.54.18.217-292.pdf" docOrigin="Papéis Avulsos de Zoologia (São Paulo) 54 (18)" docSource="http://dx.doi.org/10.1590/0031-1049.2014.54.18" docStyle="DocumentStyle:8E93E160036069519CAC16E940FF2271.1:PapAvZool.2014-2017.journal_article" docStyleId="8E93E160036069519CAC16E940FF2271" docStyleName="PapAvZool.2014-2017.journal_article" docStyleVersion="1" docTitle="Physodactylus girardi Rosa, 2014, sp. nov." docType="treatment" docVersion="2" lastPageNumber="267" masterDocId="876B2A022402FF817563616AF51CFFBB" masterDocTitle="Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)" masterLastPageNumber="292" masterPageNumber="217" pageNumber="266" updateTime="1720066190901" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A90D96EE384C3083B1E5559195A0F406">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<mods:namePart id="990C00B65B35F41954A8FDCAFD3EE783">Rosa, Simone Policena</mods:namePart>
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<mods:date id="D13AE743CA1D02D2D6555A4DA5AD9D50">2014</mods:date>
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<treatment id="7B52527A2433FFB375B361DDF440FACB" ID-DOI="http://doi.org/10.5281/zenodo.12641481" ID-Zenodo-Dep="12641481" LSID="urn:lsid:plazi:treatment:7B52527A2433FFB375B361DDF440FACB" httpUri="http://treatment.plazi.org/id/7B52527A2433FFB375B361DDF440FACB" lastPageId="50" lastPageNumber="267" pageId="49" pageNumber="266">
<subSubSection id="BBE1B0E72433FFB075B361DDF70DFF6B" box="[208,529,183,208]" pageId="49" pageNumber="266" type="nomenclature">
<paragraph id="F344E36C2433FFB075B361DDF70DFF6B" blockId="49.[208,529,183,240]" box="[208,529,183,208]" pageId="49" pageNumber="266">
<heading id="A80C54002433FFB075B361DDF70DFF6B" bold="true" box="[208,529,183,208]" centered="true" fontSize="10" level="2" pageId="49" pageNumber="266" reason="2">
<emphasis id="C18F3F7E2433FFB075B361DDF70DFF6B" bold="true" box="[208,529,183,208]" pageId="49" pageNumber="266">
<taxonomicName id="34FB98EF2433FFB075B361DDF4A4FF74" authorityName="Rosa" authorityYear="2014" box="[208,440,183,207]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="266" phylum="Arthropoda" rank="species" species="girardi" status="sp. nov.">
<emphasis id="C18F3F7E2433FFB075B361DDF4A4FF74" bold="true" box="[208,440,183,207]" italics="true" pageId="49" pageNumber="266">Physodactylus girardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052433FFB074DC61D2F70DFF6B" box="[447,529,184,208]" pageId="49" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72433FFB3747161B2F440FACB" lastPageId="50" lastPageNumber="267" pageId="49" pageNumber="266" type="description">
<paragraph id="F344E36C2433FFB0747161B2F4D3FF4B" blockId="49.[208,529,183,240]" box="[274,463,216,240]" pageId="49" pageNumber="266">
<emphasis id="C18F3F7E2433FFB0747161B2F4D3FF4B" bold="true" box="[274,463,216,240]" pageId="49" pageNumber="266">
(
<figureCitation id="6BC0FFE92433FFB0747A61B2F472FF4B" box="[281,366,216,240]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="49" pageNumber="266">Figs. 21</figureCitation>
A-I, 35I)
</emphasis>
</paragraph>
<paragraph id="F344E36C2433FFB0753D6072F750FEEB" blockId="49.[94,643,280,336]" pageId="49" pageNumber="266">
<emphasis id="C18F3F7E2433FFB0753D6072F5D6FE8B" box="[94,202,280,304]" italics="true" pageId="49" pageNumber="266">Etymology:</emphasis>
In homage to the Dr. Claude Girard, for his contribution to the knowledge of
<taxonomicName id="34FB98EF2433FFB074BC6052F755FEEB" authorityName="Leach" authorityYear="1815" box="[479,585,312,336]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="266" phylum="Arthropoda" rank="family">Elateridae</taxonomicName>
.
</paragraph>
<paragraph id="F344E36C2433FFB3753D6013F728FDEB" blockId="49.[94,643,376,592]" lastBlockId="50.[132,681,184,592]" lastPageId="50" lastPageNumber="267" pageId="49" pageNumber="266">
<emphasis id="C18F3F7E2433FFB0753D6013F5D2FE2B" box="[94,206,377,400]" italics="true" pageId="49" pageNumber="266">Description</emphasis>
(male,
<figureCitation id="6BC0FFE92433FFB0747D6012F46FFE2B" box="[286,371,376,400]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="49" pageNumber="266">Fig. 35I</figureCitation>
): Integument bright; light brown to brown, head darker, ventral abdomen black with lateral borders light brown; pilosity brown. Total length
<quantity id="34034E892433FFB075CA60B2F40DFE54" box="[169,273,472,496]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="49" pageNumber="266" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8-11 mm</quantity>
; elytral base 1.03-1.08x as wide as prothorax, elytra 2.81-2.91x longer than pronotum. Frons flat to slightly concave, frontal carina not produced anteriorly. Antenna (
<figureCitation id="6BC0FFE92433FFB074126352F4D7FDEB" box="[369,459,568,592]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="49" pageNumber="266">Fig. 21A</figureCitation>
) reaching the base of posterior angles; antennomere IV 1.05x as wide as long, V 1.14x wider than long, VI-IX 1.1-1.3x wider than long, XI oval 1.66 times longer than wide. Pronotum (
<figureCitation id="6BC0FFE92433FFB0779E6072F69FFE8B" box="[765,899,280,304]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="49" pageNumber="266">Figs. 21B, C</figureCitation>
) convex 1.07-1.12x wider than long with lateral sides gradually convergent anteriorly from posterior angle; anteromedian margin weakly produced; posterior angles short, convex, parallel and carinate; lateral carina complete, nearly entirely visible dorsally; disc with punctures 2-3 diameters apart, larger and denser on lateral and anterior borders. Protibia (
<figureCitation id="6BC0FFE92433FFB0766F6092F673FDAB" box="[780,879,504,528]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="49" pageNumber="266">Fig. 21D</figureCitation>
) with dorsal apical angle obtuse and not produced, lamellae of pro- and mesotarsomeres I minuscule, absent on metatarsomere I. Elytra parallel-sided on anterior 2/3 then tapering to apex, striae with irregular rows of punctures; interstices flat, scarcely punctate, rugose; apical striae with punctures as large as those of pronotal lateral border. Abdomen with ventrites 2-3 slightly concave laterally, ventrite 1 with setae a little smaller than those of ventrite 2, black area of ventrites 2-4 covered with moderately long setae and dense punctation, lateral brown area with punctures smaller and setae sparse. Pregenitalic segments and aedeagus covered with stout brown setae. Sternite VIII with posterior margin straight, anterior sclerotization 0.58x the total width of sternite; sternite IX (
<figureCitation id="6BC0FFE92430FFB3759C6352F441FDEB" box="[255,349,568,592]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="50" pageNumber="267">Fig. 21G</figureCitation>
) with apex rounded.
</paragraph>
<caption id="A784B3E42433FFB0753C6648F154F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641415" ID-Zenodo-Dep="12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="49" pageNumber="266" startId="49.[95,174,1826,1845]" targetBox="[97,1228,619,1795]" targetPageId="49" targetType="figure">
<paragraph id="F344E36C2433FFB0753C6648F154F8F4" blockId="49.[94,1229,1826,1871]" pageId="49" pageNumber="266">
<emphasis id="C18F3F7E2433FFB0753C6648F5D3F88E" bold="true" box="[95,207,1826,1845]" pageId="49" pageNumber="266">FIGURE 21:</emphasis>
<taxonomicName id="34FB98EF2433FFB075B66648F465F88E" authorityName="Rosa" authorityYear="2014" box="[213,377,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="266" phylum="Arthropoda" rank="species" species="girardi" status="sp. nov.">
<emphasis id="C18F3F7E2433FFB075B66648F465F88E" box="[213,377,1826,1845]" italics="true" pageId="49" pageNumber="266">Physodactylus girardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052433FFB0741D6649F4A2F88D" box="[382,446,1827,1846]" pageId="49" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E2433FFB074A16648F4C9F88E" bold="true" box="[450,469,1826,1845]" pageId="49" pageNumber="266">A,</emphasis>
antenna;
<emphasis id="C18F3F7E2433FFB077446648F748F88E" bold="true" box="[551,596,1826,1845]" pageId="49" pageNumber="266">B, C,</emphasis>
prothorax (dorsal, lateral);
<emphasis id="C18F3F7E2433FFB076566648F66BF88E" bold="true" box="[821,887,1826,1845]" pageId="49" pageNumber="266">D, E, F,</emphasis>
pro, meso- and metatibia;
<emphasis id="C18F3F7E2433FFB071356648F177F88E" bold="true" box="[1110,1131,1826,1845]" pageId="49" pageNumber="266">G,</emphasis>
sternite IX;
<emphasis id="C18F3F7E2433FFB0753D6656F594F8F4" bold="true" box="[94,136,1852,1871]" pageId="49" pageNumber="266">H, I,</emphasis>
aedeagus (ventral, dorsal).
<taxonomicName id="34FB98EF2433FFB074046656F707F8F4" authorityName="Rosa" authorityYear="2014" box="[359,539,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="266" phylum="Arthropoda" rank="species" species="gounellei" status="sp. nov.">
<emphasis id="C18F3F7E2433FFB074046656F707F8F4" box="[359,539,1852,1871]" italics="true" pageId="49" pageNumber="266">Physodactylus gounellei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052433FFB077426656F77DF8F4" box="[545,609,1852,1871]" pageId="49" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E2433FFB077076656F76EF8F4" bold="true" box="[612,626,1852,1871]" pageId="49" pageNumber="266">J,</emphasis>
antenna;
<emphasis id="C18F3F7E2433FFB077A76656F7C4F8F4" bold="true" box="[708,728,1852,1871]" pageId="49" pageNumber="266">K,</emphasis>
pronotum;
<emphasis id="C18F3F7E2433FFB0765F6656F670F8F4" bold="true" box="[828,876,1852,1871]" pageId="49" pageNumber="266">L, M,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C2430FFB375E76312F747FCEB" blockId="50.[132,681,632,848]" pageId="50" pageNumber="267">
<emphasis id="C18F3F7E2430FFB375E76312F5C1FD2B" box="[132,221,632,656]" italics="true" pageId="50" pageNumber="267">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92430FFB375936312F465FD2B" box="[240,377,632,656]" captionStart="FIGURE 21" captionStartId="49.[95,174,1826,1845]" captionTargetBox="[97,1228,619,1795]" captionTargetId="figure-201@49.[94,1228,612,1803]" captionTargetPageId="49" captionText="FIGURE 21: Physodactylus girardi sp. nov. A, antenna; B, C, prothorax (dorsal, lateral); D, E, F, pro, meso- and metatibia; G, sternite IX; H, I, aedeagus (ventral, dorsal). Physodactylus gounellei sp. nov. J, antenna; K, pronotum; L, M, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641415" httpUri="https://zenodo.org/record/12641415/files/figure.png" pageId="50" pageNumber="267">Figs. 21H, I</figureCitation>
): Phallobase 0.44x the total length of aedeagus, 1.14x longer than wide, length ratio between lateroposterior and median parts 9.5; paramere with ventral surface entirely membranous and translucent; penis with basal strut 0.31x its total length, articulated to parameres by a sclerotized process; ventral sclerite elongate, tapering apicad.
</paragraph>
<paragraph id="F344E36C2430FFB375E76212F72EFC0B" blockId="50.[132,680,888,944]" pageId="50" pageNumber="267">
<materialsCitation id="4393E9312430FFB375E76212F72EFC0B" collectingDate="1975-11" collectionCode="MZUSP" collectorName="Pirapora, M &amp; M. Alvarenga" country="Brazil" location="BRAZIL" pageId="50" pageNumber="267" specimenCount="1" typeStatus="holotype">
<emphasis id="C18F3F7E2430FFB375E76212F5FDFC2B" box="[132,225,888,912]" italics="true" pageId="50" pageNumber="267">
<typeStatus id="2C405DCE2430FFB375E76212F5C0FC2B" box="[132,220,888,912]" pageId="50" pageNumber="267" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC2430FFB375986212F440FC2B" box="[251,348,888,912]" name="Brazil" pageId="50" pageNumber="267">BRAZIL</collectingCountry>
:
<collectorName id="5E0E86BA2430FFB374106212F71CFC34" box="[371,512,888,912]" pageId="50" pageNumber="267">Pirapora, M</collectorName>
. Gervais (
<emphasis id="C18F3F7E2430FFB377E16213F785FC2B" box="[642,665,889,912]" italics="true" pageId="50" pageNumber="267">sic</emphasis>
),
<date id="8745C5AC2430FFB375E762F2F5C1FC0B" box="[132,221,920,944]" pageId="50" pageNumber="267" value="1975-11">
<collectingDate id="97013C442430FFB375E762F2F5C1FC0B" box="[132,221,920,944]" pageId="50" pageNumber="267" value="1975-11">XI.1975</collectingDate>
</date>
,
<collectorName id="5E0E86BA2430FFB3758B62F2F464FC0B" box="[232,376,920,944]" pageId="50" pageNumber="267">M. Alvarenga</collectorName>
], male (
<collectionCode id="95EA7BA92430FFB374A462F2F73AFC0B" box="[455,550,920,944]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="267">MZUSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C2430FFB375E762B3F40BFBAB" blockId="50.[132,680,984,1040]" pageId="50" pageNumber="267">
<materialsCitation id="4393E9312430FFB375E762B3F40FFBAB" collectionCode="MZUSP, CNC" pageId="50" pageNumber="267" specimenCount="4" typeStatus="paratype">
<emphasis id="C18F3F7E2430FFB375E762B3F5FAFC4B" box="[132,230,985,1008]" italics="true" pageId="50" pageNumber="267">
<typeStatus id="2C405DCE2430FFB375E762B3F5FCFC4B" box="[132,224,985,1008]" pageId="50" pageNumber="267" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same label as
<typeStatus id="2C405DCE2430FFB374EF62B2F4F5FC4B" box="[396,489,984,1008]" pageId="50" pageNumber="267" type="holotype">holotype</typeStatus>
,
<specimenCount id="E5FD28E52430FFB3749B62B2F731FC54" box="[504,557,984,1008]" count="1" pageId="50" pageNumber="267" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA92430FFB3775C62B2F781FC4B" box="[575,669,984,1008]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="50" pageNumber="267">MZUSP</collectionCode>
),
<specimenCount id="E5FD28E52430FFB375E76292F5A3FBB4" box="[132,191,1016,1040]" count="3" pageId="50" pageNumber="267" type="generic">3 exs.</specimenCount>
(
<collectionCode id="95EA7BA92430FFB375AF6292F416FBAB" box="[204,266,1016,1040]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="50" pageNumber="267">CNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C2430FFB375E76552F778FBEB" blockId="50.[132,612,1080,1104]" box="[132,612,1080,1104]" pageId="50" pageNumber="267">
<emphasis id="C18F3F7E2430FFB375E76552F41AFBEB" box="[132,262,1080,1104]" italics="true" pageId="50" pageNumber="267">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2430FFB3746E6552F470FBEB" box="[269,364,1080,1104]" name="Brazil" pageId="50" pageNumber="267">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2430FFB374166553F4E2FBF4" box="[373,510,1080,1104]" italics="true" pageId="50" pageNumber="267">
<collectingRegion id="313F2D8E2430FFB374166553F4E5FBF4" box="[373,505,1080,1104]" country="Brazil" name="Minas Gerais" pageId="50" pageNumber="267">Minas Gerais</collectingRegion>
:
</emphasis>
Pirapora.
</paragraph>
<paragraph id="F344E36C2430FFB375E76512F440FACB" blockId="50.[132,681,1144,1392]" pageId="50" pageNumber="267">
<emphasis id="C18F3F7E2430FFB375E76512F458FB2B" box="[132,324,1144,1168]" italics="true" pageId="50" pageNumber="267">
Remarks:
<taxonomicName id="34FB98EF2430FFB3758B6513F458FB2B" authorityName="Rosa" authorityYear="2014" box="[232,324,1144,1168]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="267" phylum="Arthropoda" rank="species" species="girardi">P. girardi</taxonomicName>
</emphasis>
can be recognized by the combination of the following characters: antenna reaching the base of the pronotal posterior angles, lateral carina complete and nearly entirely visible dorsally, pronotal posterior angles short and carinate, apical dorsal angle of protibia not produced, lamella of tarsomere I very small on the pro- and mesothoracic legs and absent on the metathoracic leg.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,180 @@
<document id="98F97BF86C113486735FC5E567F2B983" ID-DOI="10.1590/0031-1049.2014.54.18" ID-ISSN="1807-0205" ID-Zenodo-Dep="12641367" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720058375023" checkinUser="felipe" docAuthor="Rosa, Simone Policena" docDate="2014" docId="7B52527A2434FFB9764C67FDF4EDFDCB" docLanguage="en" docName="PapAvZool.54.18.217-292.pdf" docOrigin="Papéis Avulsos de Zoologia (São Paulo) 54 (18)" docSource="http://dx.doi.org/10.1590/0031-1049.2014.54.18" docStyle="DocumentStyle:8E93E160036069519CAC16E940FF2271.1:PapAvZool.2014-2017.journal_article" docStyleId="8E93E160036069519CAC16E940FF2271" docStyleName="PapAvZool.2014-2017.journal_article" docStyleVersion="1" docTitle="Physodactylus latithorax Rosa, 2014, sp. nov." docType="treatment" docVersion="2" lastPageNumber="272" masterDocId="876B2A022402FF817563616AF51CFFBB" masterDocTitle="Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)" masterLastPageNumber="292" masterPageNumber="217" pageNumber="271" updateTime="1720066190901" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AD78F1BBE6218F6BB5CE04558F3E8E4C">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<mods:namePart id="14B0B23CBC73273DE05723056793BC93">Rosa, Simone Policena</mods:namePart>
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<treatment id="7B52527A2434FFB9764C67FDF4EDFDCB" LSID="urn:lsid:plazi:treatment:7B52527A2434FFB9764C67FDF4EDFDCB" httpUri="http://treatment.plazi.org/id/7B52527A2434FFB9764C67FDF4EDFDCB" lastPageId="56" lastPageNumber="272" pageId="54" pageNumber="271">
<subSubSection id="BBE1B0E72434FFB7764C67FDF18DF90B" box="[815,1169,1687,1712]" pageId="54" pageNumber="271" type="nomenclature">
<paragraph id="F344E36C2434FFB7764C67FDF18DF90B" blockId="54.[815,1169,1687,1744]" box="[815,1169,1687,1712]" pageId="54" pageNumber="271">
<heading id="A80C54002434FFB7764C67FDF18DF90B" bold="true" box="[815,1169,1687,1712]" centered="true" fontSize="10" level="2" pageId="54" pageNumber="271" reason="2">
<emphasis id="C18F3F7E2434FFB7764C67FDF18DF90B" bold="true" box="[815,1169,1687,1712]" pageId="54" pageNumber="271">
<taxonomicName id="34FB98EF2434FFB7764C67FDF124F914" authorityName="Rosa" authorityYear="2014" box="[815,1080,1687,1711]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="latithorax" status="sp. nov.">
<emphasis id="C18F3F7E2434FFB7764C67FDF124F914" bold="true" box="[815,1080,1687,1711]" italics="true" pageId="54" pageNumber="271">Physodactylus latithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052434FFB7715D67F2F18DF90B" box="[1086,1169,1688,1712]" pageId="54" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72434FFB976F367D2F4EDFDCB" lastPageId="56" lastPageNumber="273" pageId="54" pageNumber="271" type="description">
<paragraph id="F344E36C2434FFB776F367D2F12CF96B" blockId="54.[815,1169,1687,1744]" box="[912,1072,1720,1744]" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB776F367D2F12CF96B" bold="true" box="[912,1072,1720,1744]" pageId="54" pageNumber="271">
(
<figureCitation id="6BC0FFE92434FFB776F467D2F6ECF96B" box="[919,1008,1720,1744]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="54" pageNumber="271">Figs. 25</figureCitation>
,
<figureCitation id="6BC0FFE92434FFB7769F67D2F135F96B" box="[1020,1065,1720,1744]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="54" pageNumber="271">35L</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="F344E36C2434FFB777AD6792F607F8EB" blockId="54.[718,1266,1784,1872]" pageId="54" pageNumber="271">
<emphasis id="C18F3F7E2434FFB777AD6792F625F8AB" box="[718,825,1784,1808]" italics="true" pageId="54" pageNumber="271">Etymology:</emphasis>
From Latin,
<emphasis id="C18F3F7E2434FFB776A96792F6E4F8AB" box="[970,1016,1784,1808]" italics="true" pageId="54" pageNumber="271">latus</emphasis>
= wide; alluding to the thorax of this species, the widest among
<taxonomicName id="34FB98EF2434FFB7710D6672F607F8EB" class="Insecta" family="Elateridae" genus="Physodactylus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="271" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="C18F3F7E2434FFB7710D6672F1EEF88B" box="[1134,1266,1816,1840]" italics="true" pageId="54" pageNumber="271">Physodactylus</emphasis>
species.
</taxonomicName>
</paragraph>
<caption id="A784B3E42435FFB6753C63B9F7DBFCBB" ID-DOI="http://doi.org/10.5281/zenodo.12641429" ID-Zenodo-Dep="12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="55" pageNumber="272" startId="55.[95,174,723,742]" targetBox="[95,1225,171,698]" targetPageId="55" targetType="figure">
<paragraph id="F344E36C2435FFB6753C63B9F7DBFCBB" blockId="55.[94,1228,723,768]" pageId="55" pageNumber="272">
<emphasis id="C18F3F7E2435FFB6753C63B9F5D2FD5D" bold="true" box="[95,206,723,742]" pageId="55" pageNumber="272">FIGURE 25:</emphasis>
<taxonomicName id="34FB98EF2435FFB675B163B9F492FD5D" authorityName="Rosa" authorityYear="2014" box="[210,398,723,742]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="272" phylum="Arthropoda" rank="species" species="latithorax" status="sp. nov.">
<emphasis id="C18F3F7E2435FFB675B163B9F492FD5D" box="[210,398,723,742]" italics="true" pageId="55" pageNumber="272">Physodactylus latithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052435FFB674F163B9F4CCFD5D" box="[402,464,723,742]" pageId="55" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E2435FFB674B063B9F4FAFD5D" bold="true" box="[467,486,723,742]" pageId="55" pageNumber="272">A,</emphasis>
antenna;
<emphasis id="C18F3F7E2435FFB6775563B9F760FD5D" bold="true" box="[566,636,723,742]" pageId="55" pageNumber="272">B, C, D,</emphasis>
prothorax (dorsal, ventral, lateral);
<emphasis id="C18F3F7E2435FFB676F863B9F6B1FD5D" bold="true" box="[923,941,723,742]" pageId="55" pageNumber="272">E,</emphasis>
hypomeron;
<emphasis id="C18F3F7E2435FFB6717863B9F15FFD5D" bold="true" box="[1051,1091,723,742]" pageId="55" pageNumber="272">F, G,</emphasis>
pro- and metatibia;
<emphasis id="C18F3F7E2435FFB675E16387F585FCBB" bold="true" box="[130,153,749,768]" pageId="55" pageNumber="272">H,</emphasis>
abdomen (ventral);
<emphasis id="C18F3F7E2435FFB674226387F452FCBB" bold="true" box="[321,334,749,768]" pageId="55" pageNumber="272">I,</emphasis>
sternite VIII;
<emphasis id="C18F3F7E2435FFB674A76387F4F7FCBB" bold="true" box="[452,491,749,768]" pageId="55" pageNumber="272">J, K,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C2435FFB6753D6253F12DFB54" blockId="55.[94,643,824,1872]" lastBlockId="55.[680,1229,824,1264]" pageId="55" pageNumber="272">
<emphasis id="C18F3F7E2435FFB6753D6253F5C8FCEB" box="[94,212,825,848]" italics="true" pageId="55" pageNumber="272">Description:</emphasis>
Integument bright ventrally, dull dorsally; head, antenna, mesoventrite, metepisternum and ventral abdomen black, pro- and mesothoracic legs dark brown with femur orange (except its apical margin dark brown), metathoracic leg as the other ones but with trochanter orange; hypomeron orange with inner margin black, prosternum orange with anterior margin and process black; pronotum and elytra brown; pilosity yellow. Total length 18.0 mm; elytral base as wide as prothorax, elytra 3.48x longer than pronotum. Frons concave, frontal carina not produced anteriorly, frontoclypeal region with a longitudinal carina medially; antenna (
<figureCitation id="6BC0FFE92435FFB6759565D2F449FB6B" box="[246,341,1208,1232]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="55" pageNumber="272">Fig. 25A</figureCitation>
) reaching the half length of pronotum; antennomere IV 1.08x longer than wide, V 1.08x wider than long, VI-VIII 1.64x wider than long, IX-X 1.5x wider than long, XI oval 1.5x longer than wide. Maxilla with galea covered with a few fine and spiniform setae. Pronotum (
<figureCitation id="6BC0FFE92435FFB674C76432F4E1FACB" box="[420,509,1368,1392]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="55" pageNumber="272">Fig. 25A</figureCitation>
) weakly convex 1.56x wider than long with posterior third subparallel then convergent to anterior margin; anteromedi- an margin weakly produced anteriad; posterior angles short, flat, parallel; lateral carina absent on anterior angle (
<figureCitation id="6BC0FFE92435FFB675C26492F41CF9AB" box="[161,256,1528,1552]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="55" pageNumber="272">Fig. 25D</figureCitation>
), visible on posterior 2/3 dorsally; disc with punctures 0.5-1.5 diameters apart, larger and denser on anterior and lateral borders; prosternal process curved 90° dorsad posteriad of procoxae, without subapical tooth (
<figureCitation id="6BC0FFE92435FFB674696712F470F92B" box="[266,364,1656,1680]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="55" pageNumber="272">Fig. 25D</figureCitation>
). Protibia with dorsal margin strongly curved and crenulate (
<figureCitation id="6BC0FFE92435FFB674A167F2F701F90B" box="[450,541,1688,1712]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="55" pageNumber="272">Fig. 25F</figureCitation>
); lamellae of pro- and mesotarsomeres I as large as those of tarsomeres II-III, small on metatarsomere I. Elytra parallel-sided on anterior 2/3 then tapering to apex; striae deeply impressed, with three rows of punctures, punctures glabrous on the median row, smaller and bearing a small seta on the lateral row; interstices flat, smooth and glabrous; apical striae with punctures smaller than those of pronotal lateral border and as large as those of elytral base. Abdomen (
<figureCitation id="6BC0FFE92435FFB676C362F2F11EFC0B" box="[928,1026,920,944]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="55" pageNumber="272">Fig. 25H</figureCitation>
) with ventrites 1-4 evenly strongly convex, ventrite 1 evenly and densely pilose with semi-erect setae as long as those of the lateral part of ventrite 2; ventrites 2-3 densely covered with short semi-erect setae laterally and long setae medially; ventrite 4 densely covered with long setae. Pregenitalic segments and aedeagus covered with stout yellow setae. Sternite VIII (
<figureCitation id="6BC0FFE92435FFB676A86512F13DFB2B" box="[971,1057,1144,1168]" captionStart="FIGURE 25" captionStartId="55.[95,174,723,742]" captionTargetBox="[95,1225,171,698]" captionTargetId="figure-525@55.[94,1228,170,699]" captionTargetPageId="55" captionText="FIGURE 25: Physodactylus latithorax sp. nov. A, antenna; B, C, D, prothorax (dorsal, ventral, lateral); E, hypomeron; F, G, pro- and metatibia; H, abdomen (ventral); I, sternite VIII; J, K, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641429" httpUri="https://zenodo.org/record/12641429/files/figure.png" pageId="55" pageNumber="272">Fig. 25I</figureCitation>
) subrectangular, posterior margin nearly straight, anterior margin with three sclerotizations occupying 0.75x the total width of sternite; sternite IX tapered to apex.
</paragraph>
<paragraph id="F344E36C2435FFB677CB6472F192F98B" blockId="55.[680,1229,1304,1584]" pageId="55" pageNumber="272">
<emphasis id="C18F3F7E2435FFB677CB6472F61AFA8B" box="[680,774,1304,1328]" italics="true" pageId="55" pageNumber="272">Aedeagus:</emphasis>
Phallobase 0.47x the total length of aedeagus, 1.27x longer than wide, length ratio between lateroposterior and median parts 8.71; paramere with ventral surface nearly entirely sclerotized, anteromedian part acute produced anteriad with tips strongly sclerotized and contiguous; penis with sides rounded posteriad of struts, abruptly narrowed near apex, basal strut 0.31x its total length, articulated to the parameres by a strongly sclerotized process; ventral sclerite parallel-sided.
</paragraph>
<paragraph id="F344E36C2435FFB677CB6732F1A6F92B" blockId="55.[680,1229,1624,1680]" pageId="55" pageNumber="272">
<materialsCitation id="4393E9312435FFB677CB6732F1A6F92B" collectingDate="1985-11-04" collectionCode="MPEG" collectorName="Serra Norte, N &amp; P. Tadeu" country="Brazil" location="Floresta" pageId="55" pageNumber="272" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<emphasis id="C18F3F7E2435FFB677CB6732F619F9CB" box="[680,773,1624,1648]" italics="true" pageId="55" pageNumber="272">
<typeStatus id="2C405DCE2435FFB677CB6732F61CF9CB" box="[680,768,1624,1648]" pageId="55" pageNumber="272" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC2435FFB6767E6732F645F9CB" box="[797,857,1624,1648]" name="Brazil" pageId="55" pageNumber="272">Brasil</collectingCountry>
<collectingRegion id="313F2D8E2435FFB676086732F686F9CB" box="[875,922,1624,1648]" country="Brazil" name="Para" pageId="55" pageNumber="272">Pará</collectingRegion>
,
<collectorName id="5E0E86BA2435FFB676D36732F17CF9CB" box="[944,1120,1624,1648]" pageId="55" pageNumber="272">Serra Norte, N</collectorName>
2-
<location id="F624B5B72435FFB671166732F1D4F9CB" LSID="urn:lsid:plazi:treatment:7B52527A2434FFB9764C67FDF4EDFDCB:F624B5B72435FFB671166732F1D4F9CB" box="[1141,1224,1624,1648]" country="Brazil" name="Floresta" pageId="55" pageNumber="272" stateProvince="Para">Floresta</location>
,
<date id="8745C5AC2435FFB677CB6712F634F92B" box="[680,808,1656,1680]" pageId="55" pageNumber="272" value="1985-11-04">
<collectingDate id="97013C442435FFB677CB6712F634F92B" box="[680,808,1656,1680]" pageId="55" pageNumber="272" value="1985-11-04">04-XI-1985</collectingDate>
</date>
], [
<collectingCountry id="8BECA3FC2435FFB6765C6712F667F92B" box="[831,891,1656,1680]" name="Brazil" pageId="55" pageNumber="272">Brasil</collectingCountry>
<collectingRegion id="313F2D8E2435FFB676E16712F6ADF92B" box="[898,945,1656,1680]" country="Brazil" name="Para" pageId="55" pageNumber="272">Pará</collectingRegion>
,
<collectorName id="5E0E86BA2435FFB676D86712F10FF92B" box="[955,1043,1656,1680]" pageId="55" pageNumber="272">P. Tadeu</collectorName>
] male (
<collectionCode id="95EA7BA92435FFB6713C6712F1B3F92B" box="[1119,1199,1656,1680]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="55" pageNumber="272">MPEG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C2435FFB677CB67D2F14EF974" blockId="55.[680,1106,1720,1744]" box="[680,1106,1720,1744]" pageId="55" pageNumber="272">
<emphasis id="C18F3F7E2435FFB677CB67D2F636F96B" box="[680,810,1720,1744]" italics="true" pageId="55" pageNumber="272">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2435FFB6765267D2F68CF96B" box="[817,912,1720,1744]" name="Brazil" pageId="55" pageNumber="272">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2435FFB676FA67D2F6D1F974" box="[921,973,1720,1743]" italics="true" pageId="55" pageNumber="272">
<collectingRegion id="313F2D8E2435FFB676FA67D2F6D4F974" box="[921,968,1720,1743]" country="Brazil" name="Para" pageId="55" pageNumber="272">Pará</collectingRegion>
:
</emphasis>
Serra Norte.
</paragraph>
<paragraph id="F344E36C2435FFB977CB6792F4EDFDCB" blockId="55.[680,1228,1784,1872]" lastBlockId="56.[132,681,184,624]" lastPageId="56" lastPageNumber="273" pageId="55" pageNumber="272">
<emphasis id="C18F3F7E2435FFB677CB6792F6E0F8AB" box="[680,1020,1784,1808]" italics="true" pageId="55" pageNumber="272">
Remarks:
<taxonomicName id="34FB98EF2435FFB6766E6792F6E0F8AB" authorityName="Rosa" authorityYear="2014" box="[781,1020,1784,1808]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="272" phylum="Arthropoda" rank="species" species="latithorax">Physodactylus latithorax</taxonomicName>
</emphasis>
is distinguished by the combination of pilosity yellow, antenna reaching the half length of pronotum, frontoclypeal region divided medially by a longitudinal carina, prothorax 1.56x wider than long, prosternal process curved 90° dorsad, lamellae of pro- and mesotarsomeres I as large as the other ones, elytra with striae deeply impressed, abdominal ventrites 2-4 strongly convex, ventrite I evenly densely covered with short and semi-erect setae; aedeagus with outer-apical angle spiniform and penis rounded posteriorly to the struts. This species is most similar to
<taxonomicName id="34FB98EF243AFFB9745C60D3F4BEFE6B" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[319,418,440,464]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="273" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="C18F3F7E243AFFB9745C60D3F4BEFE6B" box="[319,418,440,464]" italics="true" pageId="56" pageNumber="273">P. sulcatus</emphasis>
</taxonomicName>
and
<taxonomicName id="34FB98EF243AFFB974B460D3F730FE6B" authorityName="Rosa" authorityYear="2014" box="[471,556,441,464]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="273" phylum="Arthropoda" rank="species" species="patens" status="sp. nov.">
<emphasis id="C18F3F7E243AFFB974B460D3F730FE6B" box="[471,556,441,464]" italics="true" pageId="56" pageNumber="273">P. patens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243AFFB9775760D2F79AFE74" box="[564,646,440,463]" pageId="56" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
, all sharing a convex abdomen and elytral striae, from which
<taxonomicName id="34FB98EF243AFFB975A96093F45DFDAB" authorityName="Rosa" authorityYear="2014" box="[202,321,504,528]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="273" phylum="Arthropoda" rank="species" species="latithorax" status="sp. nov.">
<emphasis id="C18F3F7E243AFFB975A96093F45DFDAB" box="[202,321,504,528]" italics="true" pageId="56" pageNumber="273">P. latithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243AFFB974246092F48AFDB4" box="[327,406,504,527]" pageId="56" pageNumber="273" rank="species">sp. nov.</taxonomicNameLabel>
differs in its color pattern, larger mandible, frontoclypeal longitudinal carina, wider prothorax, longer pronotal setae, abdominal pilosity pattern and aedeagal shape.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,159 @@
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<taxonomicName id="34FB98EF2438FFBB766660FDF1A6FE0B" authority="Fleutiaux, 1892" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[773,1210,407,432]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="oberthuri">
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<emphasis id="C18F3F7E2438FFBB766660FDF115FE14" bold="true" box="[773,1033,407,431]" italics="true" pageId="58" pageNumber="275">Physodactylus oberthuri</emphasis>
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</emphasis>
</taxonomicName>
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<paragraph id="F344E36C2438FFBB761760D2F151FE6B" blockId="58.[773,1210,407,464]" box="[884,1101,440,464]" pageId="58" pageNumber="275">
<emphasis id="C18F3F7E2438FFBB761760D2F151FE6B" bold="true" box="[884,1101,440,464]" pageId="58" pageNumber="275">
(
<figureCitation id="6BC0FFE92438FFBB761860D2F11BFE6B" box="[891,1031,440,464]" captionStart="FIGURE 27" captionStartId="57.[95,174,780,799]" captionTargetBox="[98,1225,170,750]" captionTargetId="figure-548@57.[94,1228,170,756]" captionTargetPageId="57" captionText="FIGURE 27: Physodactylus niger. A, B, aedeagus (dorsal, ventral). Physodactylus oberthuri. C, D, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641433" httpUri="https://zenodo.org/record/12641433/files/figure.png" pageId="58" pageNumber="275">Figs. 27C, D</figureCitation>
;
<figureCitation id="6BC0FFE92438FFBB717760D2F158FE6B" box="[1044,1092,440,464]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="58" pageNumber="275">35N</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72438FFBB77AD6092F6C8FDCB" pageId="58" pageNumber="275" type="reference_group">
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<treatmentCitation id="725AC57D2438FFBB77AD6092F1AEFDAB" author="FLEUTIAUX, E." box="[718,1202,504,528]" page="406" pageId="58" pageNumber="275" year="1892">
<taxonomicName id="34FB98EF2438FFBB77AD6092F1AEFDAB" authority="Fleutiaux, 1892: 406" authorityName="Fleutiaux, J. Chassain" authorityPageNumber="406" authorityYear="1892" box="[718,1202,504,528]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2438FFBB77AD6092F6A0FDAB" box="[718,956,504,528]" italics="true" pageId="58" pageNumber="275">Physodactylus oberthuri</emphasis>
<bibRefCitation id="976A9E9D2438FFBB76A96092F1AEFDAB" author="FLEUTIAUX, E." box="[970,1202,504,528]" pageId="58" pageNumber="275" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
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, 407;
<treatmentCitation id="725AC57D2438FFBB76646372F6FFFD8B" author="SCHWARZ, O." box="[775,995,536,560]" page="311" pageId="58" pageNumber="275" year="1906">
<bibRefCitation id="976A9E9D2438FFBB76646372F6FFFD8B" author="SCHWARZ, O." box="[775,995,536,560]" pageId="58" pageNumber="275" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 311</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2438FFBB76976372F1F1FD8B" author="SCHENKLING, S." box="[1012,1261,536,560]" page="508" pageId="58" pageNumber="275" year="1927">
<bibRefCitation id="976A9E9D2438FFBB76976372F1F1FD8B" author="SCHENKLING, S." box="[1012,1261,536,560]" pageId="58" pageNumber="275" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 508</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2438FFBB76646352F6F0FDEB" author="FLEUTIAUX, E." box="[775,1004,568,592]" page="168" pageId="58" pageNumber="275" year="1940">
<bibRefCitation id="976A9E9D2438FFBB76646352F6F0FDEB" author="FLEUTIAUX, E." box="[775,1004,568,592]" pageId="58" pageNumber="275" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 168</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2438FFBB769B6352F1F3FDEB" author="BLACKWELDER, R. E." box="[1016,1263,568,592]" page="292" pageId="58" pageNumber="275" year="1944">
<bibRefCitation id="976A9E9D2438FFBB769B6352F1F3FDEB" author="BLACKWELDER, R. E." box="[1016,1263,568,592]" pageId="58" pageNumber="275" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D2438FFBB76646332F6CDFDCB" author="CHASSAIN, J." box="[775,977,600,624]" page="68" pageId="58" pageNumber="275" year="2005">
<bibRefCitation id="976A9E9D2438FFBB76646332F6CDFDCB" author="CHASSAIN, J." box="[775,977,600,624]" pageId="58" pageNumber="275" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005: 68</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72438FFBA77AD63F2F40EFCAB" lastPageId="59" lastPageNumber="276" pageId="58" pageNumber="275" type="description">
<paragraph id="F344E36C2438FFBB77AD63F2F1EDF9B4" blockId="58.[718,1267,664,1552]" pageId="58" pageNumber="275">
<emphasis id="C18F3F7E2438FFBB77AD63F2F651FD0B" box="[718,845,664,688]" italics="true" pageId="58" pageNumber="275">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE92438FFBB76D063F2F103FD0B" box="[947,1055,664,688]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="58" pageNumber="275">Fig. 35N</figureCitation>
): Integument very bright, dark brown with elytra lighter; pilosity brown. Total length
<quantity id="34034E892438FFBB763563B2F6A6FD54" box="[854,954,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="58" pageNumber="275" unit="mm" value="12.5">12.5 mm</quantity>
; elytral base 0.97x as wide as prothorax, elytra 2.7x longer than pronotum. Frons with a circular protuberance medially; frontal carina produced anteriorly; antenna nearly reaching the half length of pronotum; antennomere IV-X 1.2-1.3x wider than long; XI oval. Pronotum 1.18x wider than long, lateral sides rounded from posterior angles to anterior margin, strongly narrowed on anterior angle, anteromedian margin strongly produced; posterior angles elongate, convex, parallel; lateral carina complete, prominent, nearly entirely visible dorsally; disc and lateral margin with punctures very small and weakly impressed, 4-6 diameters apart, larger on anterior margin, homogenously distributed. Lamellae of pro- and mesotarsomeres I smaller than the others, present on metatarsomere I. Elytra rounded on anterior 2/3 then tapering to apex; striae with a row of punctures larger than those of lateral pronotal border, interstices slightly convex, rugose, sparsely punctate. Abdomen with ventrites 2-3 concave laterally, with setae moderately long and decumbent on ventrites 2-5, ventrite 1 with setae a little shorter than and as dense as those of ventrites 2-5. Pregenitalic segments and aedeagus covered with brown setae. Sternite VIII with posterior margin emarginate medially, anterior sclerotization curved 0.57x the total width of sternite; sternite IX with apex acute.
</paragraph>
<paragraph id="F344E36C2438FFBB77AD6752F195F8EB" blockId="58.[718,1266,1592,1872]" pageId="58" pageNumber="275">
<emphasis id="C18F3F7E2438FFBB77AD6752F63BF9EB" box="[718,807,1592,1616]" italics="true" pageId="58" pageNumber="275">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92438FFBB765B6752F6D4F9EB" box="[824,968,1592,1616]" captionStart="FIGURE 27" captionStartId="57.[95,174,780,799]" captionTargetBox="[98,1225,170,750]" captionTargetId="figure-548@57.[94,1228,170,756]" captionTargetPageId="57" captionText="FIGURE 27: Physodactylus niger. A, B, aedeagus (dorsal, ventral). Physodactylus oberthuri. C, D, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641433" httpUri="https://zenodo.org/record/12641433/files/figure.png" pageId="58" pageNumber="275">Figs. 27C, D</figureCitation>
): Phallobase 0.49x the total length of aedeagus, 1.32x longer than wide, length ratio between lateroposterior and median parts 3.7; paramere with ventral surface partly membranous, except for the lateral margin and the anterior margin sclerotized; anteromedian sclerotized margin approximate at middle; penis with basal strut 0.34x its total length, articulated to the parameres by a translucent membrane; ventral sclerite tapering apicad.
</paragraph>
<paragraph id="F344E36C2439FFBA753D61D2F74DFE0B" blockId="59.[94,643,184,432]" pageId="59" pageNumber="276">
<emphasis id="C18F3F7E2439FFBA753D61D2F5A7FF6B" box="[94,187,184,208]" italics="true" pageId="59" pageNumber="276">
<typeStatus id="2C405DCE2439FFBA753D61D2F5AAFF6B" box="[94,182,184,208]" pageId="59" pageNumber="276" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC2439FFBA75BA61D2F409FF6B" box="[217,277,184,208]" name="Brazil" pageId="59" pageNumber="276">Brèsil</collectingCountry>
mér], [ex-Musaeo Miniszech], [TYPE], [Fltx. 1891], [1940, FLEUTIAUX det.], [Fleut. Ann. Soc. Ent. Fr. 1892, p. 407, t.
<specimenCount id="E5FD28E52439FFBA77756192F725FEAB" box="[534,569,248,272]" count="8" pageId="59" pageNumber="276" type="generic">8 f.</specimenCount>
2 Collection FLEUTIAUX], [
<taxonomicName id="34FB98EF2439FFBA74026072F587FEEB" authority="J Fleut." authorityName="J Fleut." class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="oberthuri">Physodactylus oberthuri J Fleut.</taxonomicName>
type], [
<taxonomicName id="34FB98EF2439FFBA75886052F755FEEB" authority="J Fleut." authorityName="J Fleut." box="[235,585,311,336]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="oberthuri">Physodactylus oberthuri J Fleut.</taxonomicName>
type,
<collectingCountry id="8BECA3FC2439FFBA753D6032F58BFECB" box="[94,151,344,368]" name="Brazil" pageId="59" pageNumber="276">Brèsil</collectingCountry>
(Coll. Oberthur)], [Museum Paris, ex. Coll. R. Oberthur], [
<typeStatus id="2C405DCE2439FFBA759C6012F470FE2B" box="[255,364,376,400]" pageId="59" pageNumber="276" type="lectotype">Lectotype</typeStatus>
], [
<taxonomicName id="34FB98EF2439FFBA74E76012F422FE0B" authority="Fleutiaux, J. Chassain" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="oberthuri">Physodactylus oberthüri Fleutiaux, J. Chassain</taxonomicName>
det. 05], male, (MNHN).
</paragraph>
<paragraph id="F344E36C2439FFBA753D60B2F455FE4B" blockId="59.[94,329,472,496]" box="[94,329,472,496]" pageId="59" pageNumber="276">
<emphasis id="C18F3F7E2439FFBA753D60B2F5FDFE4B" box="[94,225,472,496]" italics="true" pageId="59" pageNumber="276">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2439FFBA758460B2F45AFE4B" box="[231,326,472,496]" name="Brazil" pageId="59" pageNumber="276">BRAZIL</collectingCountry>
.
</paragraph>
<paragraph id="F344E36C2439FFBA753D6372F40EFCAB" blockId="59.[94,643,536,784]" pageId="59" pageNumber="276">
<emphasis id="C18F3F7E2439FFBA753D6372F5A4FD8B" box="[94,184,536,560]" italics="true" pageId="59" pageNumber="276">Remarks:</emphasis>
The apical part of the prosternal process was not examined.
<taxonomicName id="34FB98EF2439FFBA744C6352F705FDEB" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[303,537,568,592]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2439FFBA744C6352F705FDEB" box="[303,537,568,592]" italics="true" pageId="59" pageNumber="276">Physodactylus oberthuri</emphasis>
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is similar to
<taxonomicName id="34FB98EF2439FFBA751F6333F5DAFDCB" box="[124,198,601,624]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2439FFBA751F6333F5DAFDCB" box="[124,198,601,624]" italics="true" pageId="59" pageNumber="276">P. niger</emphasis>
</taxonomicName>
and
<taxonomicName id="34FB98EF2439FFBA759D6333F493FDCB" authorityName="Rosa" authorityYear="2014" box="[254,399,600,624]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="C18F3F7E2439FFBA759D6333F493FDCB" box="[254,399,600,624]" italics="true" pageId="59" pageNumber="276">P. tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052439FFBA74F46332F4F4FDD4" box="[407,488,600,623]" pageId="59" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
in prothoracic shape. It is most similar to this latter in the shape of aedeagus, very bright integument and pronotal punctation homogenously distributed. It differs from both species manly in its brown setae and the very fine pronotal punctation.
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<emphasis id="C18F3F7E2439FFBA75B0623DF712FCCB" bold="true" box="[211,526,855,880]" pageId="59" pageNumber="276">
<taxonomicName id="34FB98EF2439FFBA75B0623DF4A9FCD4" authorityName="Rosa" authorityYear="2014" box="[211,437,855,879]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="patens" status="sp. nov.">
<emphasis id="C18F3F7E2439FFBA75B0623DF4A9FCD4" bold="true" box="[211,437,855,879]" italics="true" pageId="59" pageNumber="276">Physodactylus patens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052439FFBA74DF6232F712FCCB" box="[444,526,856,880]" pageId="59" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E72439FFBD747D6212F1A5FEAB" lastPageId="60" lastPageNumber="277" pageId="59" pageNumber="276" type="description">
<paragraph id="F344E36C2439FFBA747D6212F4DFFC2B" blockId="59.[211,526,855,912]" box="[286,451,888,912]" pageId="59" pageNumber="276">
<emphasis id="C18F3F7E2439FFBA747D6212F4DFFC2B" bold="true" box="[286,451,888,912]" pageId="59" pageNumber="276">
(
<figureCitation id="6BC0FFE92439FFBA74466212F461FC2B" box="[293,381,888,912]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">
Figs.
<geoCoordinate id="96CF85AB2439FFBA74036212F461FC2B" box="[352,381,888,912]" degrees="28, 35" direction="east" orientation="longitude" pageId="59" pageNumber="276" precision="555" value="28.35">28</geoCoordinate>
</figureCitation>
,
<figureCitation id="6BC0FFE92439FFBA74EA6212F4A7FC2B" box="[393,443,888,912]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="59" pageNumber="276">35O</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="F344E36C2439FFBA753D62D2F48CFC4B" blockId="59.[94,642,952,1008]" pageId="59" pageNumber="276">
<emphasis id="C18F3F7E2439FFBA753D62D2F5D6FC6B" box="[94,202,952,976]" italics="true" pageId="59" pageNumber="276">Etymology:</emphasis>
From latin,
<taxonomicName id="34FB98EF2439FFBA742C62D3F490FC6B" authorityName="Rosa" authorityYear="2014" box="[335,396,953,976]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="patens">
<emphasis id="C18F3F7E2439FFBA742C62D3F490FC6B" box="[335,396,953,976]" italics="true" pageId="59" pageNumber="276">patens</emphasis>
</taxonomicName>
= wide; alluding to the widened elytra of this species.
</paragraph>
<paragraph id="F344E36C2439FFBA753D6573F651FAB4" blockId="59.[94,643,1048,1296]" lastBlockId="59.[680,1229,184,1296]" pageId="59" pageNumber="276">
<emphasis id="C18F3F7E2439FFBA753D6573F5D2FB8B" box="[94,206,1049,1072]" italics="true" pageId="59" pageNumber="276">Description</emphasis>
(male,
<figureCitation id="6BC0FFE92439FFBA74526572F481FB8B" box="[305,413,1048,1072]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="59" pageNumber="276">Fig. 35O</figureCitation>
): Integument bright ventrally, dull dorsally; brown with posterior third of elytra and abdominal marginal plates black; tibiae and tarsi dark brown; pilosity yellow. Total length 17.0 mm; elytral base 1.04x as wide as prothorax, elytra 3.15x longer than pronotum. Frons (
<figureCitation id="6BC0FFE92439FFBA777C65D2F767FB6B" box="[543,635,1208,1232]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">Fig. 28B</figureCitation>
) concave; frontal carina not produced anteriorly; frontoclypeal region with a longitudinal carina medially; antenna (
<figureCitation id="6BC0FFE92439FFBA766B61D2F67FFF6B" box="[776,867,184,208]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">Fig. 28A</figureCitation>
) reaching the half length of pronotum; antennomere IV 1.27x wider than long, V 1.58x wider than long, VI-VIII 1.4-1.5x wider than long, IX-X 1.4 and 1.33x wider than long, XI suboval 1.25x longer than wide. Maxilla with galea covered with a fine and spiniform setae. Pronotum (
<figureCitation id="6BC0FFE92439FFBA717E6032F164FECB" box="[1053,1144,344,368]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">Fig. 28C</figureCitation>
) convex 1.46x wider than long with sides rounded and convergent from posterior angles to anterior margin; anteromedian margin produced anteriad; posterior angles short, convex, parallel; lateral carina nearly reaching the anterior margin, prominent, nearly entirely visible dorsally; disc with punctures small 0.5-1.0 diameters apart, denser on anterior border, larger and denser on lateral border; prosternal process without subapical tooth (
<figureCitation id="6BC0FFE92439FFBA778D6312F653FD2B" box="[750,847,632,656]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">Fig. 28D</figureCitation>
). Protibia with dorsal margin strongly curved and crenulate (
<figureCitation id="6BC0FFE92439FFBA76D163F2F113FD0B" box="[946,1039,664,688]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">Fig. 28F</figureCitation>
); lamellae of pro- and mesotarsomeres I as large as those of tarsomeres II-III, small on metatarsomere I. Elytra widened and rounded from base to posterior 1/4 then tapering to apex; striae deeply impressed, with three rows of punctures, punctures glabrous on the median row, smaller and bearing a small seta on the lateral row; interstices convex, smooth and glabrous; apical striae with punctures smaller than those of pronotal lateral border and as large as those of elytral base. Abdomen with ventrites 1-4 evenly strongly convex, ventrite 1 evenly pilose with fine setae as long as those of the lateral part of ventrite 2; ventrites 2-3 covered with short fine setae laterally and long fine setae medially, ventrite 4 with long fine setae. Pregenitalic segments (
<figureCitation id="6BC0FFE92439FFBA77D36512F61EFB2B" box="[688,770,1144,1168]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">Figs. 28</figureCitation>
I-L) and aedeagus covered with stout yellow setae. Sternite VIII (
<figureCitation id="6BC0FFE92439FFBA76E465F2F6FAFB0B" box="[903,998,1176,1200]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">Fig. 28L</figureCitation>
) subrectangular, posterior margin nearly straight, anterior sclerotization 0.72x the total width of sternite; sternite IX (
<figureCitation id="6BC0FFE92439FFBA711165B2F1D9FB4B" box="[1138,1221,1240,1264]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276">Fig. 28I</figureCitation>
) tapered to apex.
</paragraph>
<caption id="A784B3E42439FFBA753C6648F150F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641437" ID-Zenodo-Dep="12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="59" pageNumber="276" startId="59.[95,174,1826,1845]" targetBox="[95,1226,1314,1800]" targetPageId="59" targetType="figure">
<paragraph id="F344E36C2439FFBA753C6648F150F8F4" blockId="59.[94,1228,1826,1871]" pageId="59" pageNumber="276">
<emphasis id="C18F3F7E2439FFBA753C6648F5D3F88E" bold="true" box="[95,207,1826,1845]" pageId="59" pageNumber="276">FIGURE 28:</emphasis>
<taxonomicName id="34FB98EF2439FFBA75B76648F468F88E" authorityName="Rosa" authorityYear="2014" box="[212,372,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="276" phylum="Arthropoda" rank="species" species="patens" status="sp. nov.">
<emphasis id="C18F3F7E2439FFBA75B76648F468F88E" box="[212,372,1826,1845]" italics="true" pageId="59" pageNumber="276">Physodactylus patens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052439FFBA741A6649F4A5F88D" box="[377,441,1827,1846]" pageId="59" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E2439FFBA74DE6648F4CCF88E" bold="true" box="[445,464,1826,1845]" pageId="59" pageNumber="276">A,</emphasis>
antenna;
<emphasis id="C18F3F7E2439FFBA77426648F728F88E" bold="true" box="[545,564,1826,1845]" pageId="59" pageNumber="276">B,</emphasis>
head (dorsal);
<emphasis id="C18F3F7E2439FFBA77CC6648F7C3F88E" bold="true" box="[687,735,1826,1845]" pageId="59" pageNumber="276">C, D,</emphasis>
prothorax (dorsal, lateral);
<emphasis id="C18F3F7E2439FFBA76A36648F6CEF88E" bold="true" box="[960,978,1826,1845]" pageId="59" pageNumber="276">E,</emphasis>
scutellar shield;
<emphasis id="C18F3F7E2439FFBA71396648F183F88E" bold="true" box="[1114,1183,1826,1845]" pageId="59" pageNumber="276">F, G, H,</emphasis>
pro-, meso- and metatibia;
<emphasis id="C18F3F7E2439FFBA74726656F403F8F4" bold="true" box="[273,287,1852,1871]" pageId="59" pageNumber="276">I,</emphasis>
sternite IX;
<emphasis id="C18F3F7E2439FFBA74E66656F48FF8F4" bold="true" box="[389,403,1852,1871]" pageId="59" pageNumber="276">J,</emphasis>
tergites IX and X;
<emphasis id="C18F3F7E2439FFBA774C6656F758F8F4" bold="true" box="[559,580,1852,1871]" pageId="59" pageNumber="276">K,</emphasis>
tergite VIII;
<emphasis id="C18F3F7E2439FFBA77D26656F7DFF8F4" bold="true" box="[689,707,1852,1871]" pageId="59" pageNumber="276">L,</emphasis>
sternite VIII;
<emphasis id="C18F3F7E2439FFBA765B6656F670F8F4" bold="true" box="[824,876,1852,1871]" pageId="59" pageNumber="276">M, N,</emphasis>
aedeagus, (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C243EFFBD75E761D2F443FDAB" blockId="60.[132,681,184,528]" pageId="60" pageNumber="277">
<emphasis id="C18F3F7E243EFFBD75E761D2F5C1FF6B" box="[132,221,184,208]" italics="true" pageId="60" pageNumber="277">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9243EFFBD758E61D2F49EFF6B" box="[237,386,184,208]" captionStart="FIGURE 28" captionStartId="59.[95,174,1826,1845]" captionTargetBox="[95,1226,1314,1800]" captionTargetId="figure-513@59.[94,1228,1311,1802]" captionTargetPageId="59" captionText="FIGURE 28: Physodactylus patens sp. nov. A, antenna; B, head (dorsal); C, D, prothorax (dorsal, lateral); E, scutellar shield; F, G, H, pro-, meso- and metatibia; I, sternite IX; J, tergites IX and X; K, tergite VIII; L, sternite VIII; M, N, aedeagus, (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641437" httpUri="https://zenodo.org/record/12641437/files/figure.png" pageId="60" pageNumber="277">Figs. 28M, N</figureCitation>
): Phallobase 0.47x the total length of aedeagus, 1.31x longer than wide, with lateral part abruptly narrowed on posterior half, length ratio between lateroposterior and median parts 5.36; ventral surface of paramere with the lateral half sclerotized, anteromedian margin contiguous arch-shaped; penis with sides strongly widened and rounded posteriad of struts then abruptly narrowed and tapering to apex, with basal strut 0.30x its total length, articulat- ed to the parameres by a strongly sclerotized process; ventral sclerite linear.
</paragraph>
<paragraph id="F344E36C243EFFBD75E76352F761FDCB" blockId="60.[132,681,568,624]" pageId="60" pageNumber="277">
<materialsCitation id="4393E931243EFFBD75E76352F761FDCB" collectingDate="1968-12" collectionCode="DZUP" collectorName="M. Alvarenga &amp; Colecao M. Alvarenga" country="Brazil" location="Jacareacanga" pageId="60" pageNumber="277" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<emphasis id="C18F3F7E243EFFBD75E76352F5FDFDEB" box="[132,225,568,592]" italics="true" pageId="60" pageNumber="277">
<typeStatus id="2C405DCE243EFFBD75E76352F5C0FDEB" box="[132,220,568,592]" pageId="60" pageNumber="277" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<location id="F624B5B7243EFFBD758E6352F469FDEB" LSID="urn:lsid:plazi:treatment:7B52527A2439FFBD75B0623DF1A5FEAB:F624B5B7243EFFBD758E6352F469FDEB" box="[237,373,568,592]" country="Brazil" name="Jacareacanga" pageId="60" pageNumber="277" stateProvince="Para">Jacareacanga</location>
,
<collectingRegion id="313F2D8E243EFFBD741D6352F4B6FDEB" box="[382,426,568,592]" country="Brazil" name="Para" pageId="60" pageNumber="277">Pará</collectingRegion>
<collectingCountry id="8BECA3FC243EFFBD74D36352F4F0FDEB" box="[432,492,568,592]" name="Brazil" pageId="60" pageNumber="277">Brasil</collectingCountry>
,
<date id="8745C5AC243EFFBD74956352F746FDEB" box="[502,602,568,592]" pageId="60" pageNumber="277" value="1968-12">
<collectingDate id="97013C44243EFFBD74956352F746FDEB" box="[502,602,568,592]" pageId="60" pageNumber="277" value="1968-12">XII-1968</collectingDate>
</date>
,
<collectorName id="5E0E86BA243EFFBD77076352F5C4FDCB" pageId="60" pageNumber="277">M. Alvarenga</collectorName>
], [
<collectorName id="5E0E86BA243EFFBD758D6332F4C4FDCB" box="[238,472,600,624]" pageId="60" pageNumber="277">Coleção M. Alvarenga</collectorName>
] male (
<collectionCode id="95EA7BA9243EFFBD77476332F76DFDCB" box="[548,625,600,624]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="60" pageNumber="277" type="University or college">DZUP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C243EFFBD75E763F2F724FD0B" blockId="60.[132,568,664,688]" box="[132,568,664,688]" pageId="60" pageNumber="277">
<emphasis id="C18F3F7E243EFFBD75E763F2F41AFD0B" box="[132,262,664,688]" italics="true" pageId="60" pageNumber="277">Distribution:</emphasis>
<collectingCountry id="8BECA3FC243EFFBD746E63F2F470FD0B" box="[269,364,664,688]" name="Brazil" pageId="60" pageNumber="277">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E243EFFBD741663F2F4B5FD14" box="[373,425,664,687]" italics="true" pageId="60" pageNumber="277">
<collectingRegion id="313F2D8E243EFFBD741663F2F4B8FD14" box="[373,420,664,687]" country="Brazil" name="Para" pageId="60" pageNumber="277">Pará</collectingRegion>
:
</emphasis>
Jacareacanga.
</paragraph>
<paragraph id="F344E36C243EFFBD75E763B2F1A5FEAB" blockId="60.[132,682,728,1200]" lastBlockId="60.[718,1266,184,272]" pageId="60" pageNumber="277">
<emphasis id="C18F3F7E243EFFBD75E763B2F427FD4B" box="[132,315,728,752]" italics="true" pageId="60" pageNumber="277">
Remarks:
<taxonomicName id="34FB98EF243EFFBD758563B3F427FD4B" authorityName="Rosa" authorityYear="2014" box="[230,315,729,752]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="277" phylum="Arthropoda" rank="species" species="patens">P. patens</taxonomicName>
</emphasis>
is diagnosable by the combination of pilosity yellow, antenna reaching the half length of pronotum, prothorax 1.46x wider than long, lamellae of pro- and mesotarsomeres I as large as the other ones on the same leg, elytra with striae deeply impressed, abdominal ventrites 2-4 strongly convex, ventrite 1 densely covered with short, decumbent and fine setae. The aedeagus with penis strongly widened and round- ed posteriad of the struts and the phallobase with lateral parts abruptly narrowed on posterior half are autapomorphies of this species. Its strongly convex abdomen is a character shared with
<taxonomicName id="34FB98EF243EFFBD74F56553F4E5FBEB" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[406,505,1080,1104]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="277" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="C18F3F7E243EFFBD74F56553F4E5FBEB" box="[406,505,1080,1104]" italics="true" pageId="60" pageNumber="277">P. sulcatus</emphasis>
</taxonomicName>
and
<taxonomicName id="34FB98EF243EFFBD77526553F7B4FBEB" authorityName="Rosa" authorityYear="2014" box="[561,680,1080,1104]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="277" phylum="Arthropoda" rank="species" species="latithorax" status="sp. nov.">
<emphasis id="C18F3F7E243EFFBD77526553F7B4FBEB" box="[561,680,1080,1104]" italics="true" pageId="60" pageNumber="277">P. latithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243EFFBD75E76532F5C9FBD4" box="[132,213,1112,1135]" pageId="60" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
, from which it differs in its color pattern and elytral shape. It is most similar to
<taxonomicName id="34FB98EF243EFFBD74B36513F758FB2B" authorityName="Rosa" authorityYear="2014" box="[464,580,1144,1168]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="277" phylum="Arthropoda" rank="species" species="latithorax">
<emphasis id="C18F3F7E243EFFBD74B36513F758FB2B" box="[464,580,1144,1168]" italics="true" pageId="60" pageNumber="277">P. latithorax</emphasis>
</taxonomicName>
by the pilosity pattern and elytral striae, but the elytra of
<taxonomicName id="34FB98EF243EFFBD770965F3F7F7FF6B" authorityName="Rosa" authorityYear="2014" box="[618,747,185,1200]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="277" phylum="Arthropoda" rank="species" species="patens" status="sp. nov.">
<emphasis id="C18F3F7E243EFFBD770965F3F7F7FF6B" box="[618,747,185,1200]" italics="true" pageId="60" pageNumber="277">P. patens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243EFFBD779161D2F65CFF74" box="[754,832,184,207]" pageId="60" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
are wider and rounded and the abdominal setae are finer, decumbent and sparser. The carina on the frontoclypeal region is unique to this species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<heading id="A80C5400243AFFB9767E61DDF1BEFF6B" bold="true" box="[797,1186,183,208]" centered="true" fontSize="10" level="2" pageId="56" pageNumber="273" reason="2">
<taxonomicName id="34FB98EF243AFFB9767E61DDF1BEFF6B" authority="Fleutiaux, 1892" authorityName="Fleutiaux" authorityYear="1892" box="[797,1186,183,208]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="273" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E243AFFB9767E61DDF1BEFF6B" bold="true" box="[797,1186,183,208]" pageId="56" pageNumber="273">
<emphasis id="C18F3F7E243AFFB9767E61DDF6EDFF6B" bold="true" box="[797,1009,183,208]" italics="true" pageId="56" pageNumber="273">Physodactylus niger</emphasis>
<bibRefCitation id="976A9E9D243AFFB9769B61D2F1BEFF6B" author="FLEUTIAUX, E." box="[1016,1186,184,208]" pageId="56" pageNumber="273" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
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<paragraph id="F344E36C243AFFB9765661B2F190FF4B" blockId="56.[797,1186,183,240]" box="[821,1164,216,240]" pageId="56" pageNumber="273">
<emphasis id="C18F3F7E243AFFB9765661B2F190FF4B" bold="true" box="[821,1164,216,240]" pageId="56" pageNumber="273">
(
<figureCitation id="6BC0FFE9243AFFB9765F61B2F688FF4B" box="[828,916,216,240]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="56" pageNumber="273">Figs. 26</figureCitation>
;
<figureCitation id="6BC0FFE9243AFFB976C261B2F6A7FF4B" box="[929,955,216,240]" captionStart="FIGURE 27" captionStartId="57.[95,174,780,799]" captionTargetBox="[98,1225,170,750]" captionTargetId="figure-548@57.[94,1228,170,756]" captionTargetPageId="57" captionText="FIGURE 27: Physodactylus niger. A, B, aedeagus (dorsal, ventral). Physodactylus oberthuri. C, D, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641433" httpUri="https://zenodo.org/record/12641433/files/figure.png" pageId="56" pageNumber="273">27</figureCitation>
A-B; 33E, N; 35M)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7243AFFB977AD6072F1EEFEEB" pageId="56" pageNumber="273" type="reference_group">
<paragraph id="F344E36C243AFFB977AD6072F1EEFE8B" blockId="56.[718,1266,280,432]" box="[718,1266,280,304]" pageId="56" pageNumber="273">
<treatmentCitationGroup id="D3EBC442243AFFB977AD6072F1EEFE8B" box="[718,1266,280,304]" pageId="56" pageNumber="273">
<treatmentCitation id="725AC57D243AFFB977AD6072F1B6FE8B" author="FLEUTIAUX, E." box="[718,1194,280,304]" page="406" pageId="56" pageNumber="273" year="1892">
<taxonomicName id="34FB98EF243AFFB977AD6072F1B6FE8B" authority="Fleutiaux, 1892: 406" authorityName="Fleutiaux" authorityPageNumber="406" authorityYear="1892" box="[718,1194,280,304]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="273" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E243AFFB977AD6072F686FE8B" box="[718,922,280,304]" italics="true" pageId="56" pageNumber="273">Physodactylus niger</emphasis>
<bibRefCitation id="976A9E9D243AFFB976D36072F1B6FE8B" author="FLEUTIAUX, E." box="[944,1194,280,304]" pageId="56" pageNumber="273" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux, 1892: 406</bibRefCitation>
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, 407;
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<paragraph id="F344E36C243AFFB976646052F1EEFEEB" blockId="56.[718,1266,280,432]" box="[775,1266,312,336]" pageId="56" pageNumber="273">
<bibRefCitation id="976A9E9D243AFFB976646052F6FFFEEB" author="SCHWARZ, O." box="[775,995,312,336]" pageId="56" pageNumber="273" pagination="310 - 315" refId="ref51110" refString="SCHWARZ, O. 1906. Coleoptera, Fam. Elateridae Fasc. 46 A. In: Wytsman, P. (Ed.). Genera Insectorum. Bruxelles, Verteneuil &amp; Desmet, p. 310 - 315." type="book chapter" year="1906">Schwarz, 1906: 311</bibRefCitation>
;
<bibRefCitation id="976A9E9D243AFFB976976052F1F1FEEB" author="SCHENKLING, S." box="[1012,1261,312,336]" pageId="56" pageNumber="273" pagination="265 - 636" refId="ref50939" refString="SCHENKLING, S. 1927. Pars 88: Elateridae. In: Schenkling, S. (Ed.). Coleopterorum Catalogus XI. Berlin, W. Junk. p. 265 - 636." type="book chapter" year="1927">Schenkling, 1927: 508</bibRefCitation>
;
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7243AFFBB76646032F1CAFEEB" lastPageId="58" lastPageNumber="275" pageId="56" pageNumber="273" type="description">
<paragraph id="F344E36C243AFFB976646032F127FE14" blockId="56.[718,1266,280,432]" pageId="56" pageNumber="273">
<bibRefCitation id="976A9E9D243AFFB976646032F6F7FECB" author="FLEUTIAUX, E." box="[775,1003,344,368]" pageId="56" pageNumber="273" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 168</bibRefCitation>
;
<bibRefCitation id="976A9E9D243AFFB976956032F1F0FECB" author="BLACKWELDER, R. E." box="[1014,1260,344,368]" pageId="56" pageNumber="273" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
.
<taxonomicName id="34FB98EF243AFFB977AD6013F6C4FE2B" authority="Fleutiaux, 1940c" authorityName="Fleutiaux" authorityYear="1940" box="[718,984,376,400]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="273" phylum="Arthropoda" rank="species" species="carreti" status="syn. nov.">
<emphasis id="C18F3F7E243AFFB977AD6013F638FE2B" box="[718,804,377,400]" italics="true" pageId="56" pageNumber="273">P. carreti</emphasis>
<bibRefCitation id="976A9E9D243AFFB976486012F6C4FE2B" author="FLEUTIAUX, E." box="[811,984,376,400]" pageId="56" pageNumber="273" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243AFFB976BD6012F126FE34" box="[990,1082,376,400]" pageId="56" pageNumber="273">syn. nov.</taxonomicNameLabel>
<taxonomicName id="34FB98EF243AFFB977AD60F3F6C4FE0B" box="[718,984,408,432]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="273" phylum="Arthropoda" rank="variety" species="carreti" status="syn. nov." variety="testaceipennis">
<emphasis id="C18F3F7E243AFFB977AD60F3F638FE0B" box="[718,804,409,432]" italics="true" pageId="56" pageNumber="273">P. carreti</emphasis>
var.
<emphasis id="C18F3F7E243AFFB9763660F3F6C4FE0B" box="[853,984,409,432]" italics="true" pageId="56" pageNumber="273">testaceipennis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243AFFB976BC60F2F127FE14" box="[991,1083,408,432]" pageId="56" pageNumber="273">syn. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="F344E36C243AFFB877AD60B2F6A1FB6B" blockId="56.[718,1266,472,624]" lastBlockId="57.[94,1229,780,1872]" lastPageId="57" lastPageNumber="274" pageId="56" pageNumber="273">
<emphasis id="C18F3F7E243AFFB977AD60B2F652FE4B" box="[718,846,472,496]" italics="true" pageId="56" pageNumber="273">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9243AFFB976FD60B2F11FFE4B" box="[926,1027,472,496]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="56" pageNumber="273">Fig. 35M</figureCitation>
): Integument with yellow pilosity and variable color patterns: 1) brown to reddish-black with epipleura lighter; 2) black with pronotum dark reddish-brown; 3) pronotum light yellowish-brown to orange, elytra light brown to dark brown, ventral surface dark brown to black, legs evenly dark mesotarsomeres I smaller than the others, absent on brown to black or with dorsal margin of tibiae lighter; metatarsomere I. Elytra tapering apicad from base or epipleura light brown; 4) same as the former but with its half length, striae (
<figureCitation id="6BC0FFE9243BFFB876E86212F6F7FC2B" box="[907,1003,888,912]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="57" pageNumber="274">Fig. 26N</figureCitation>
) with a row of punclateral carina and pronotal anterior and posterior mar- tures, punctures of apical striae larger than those of gins black, some specimens also with a black median the lateral pronotal border; interstices slightly convex, longitudinal strip; 5) head reddish-black, pronotum, sparsely punctate. Abdominal ventrites weakly convex elytra and epipleura light yellowish-brown, ventral with setae short and decumbent on ventrites 2-4, half surface and legs darker; 6) same as the former but with shorter, very fine and sparser on ventrite 1. Pregenitaventral surface, legs and scutellum black; 7) same as lic segments and aedeagus covered with yellow setae. the former but with pronotal borders black. Antenna Sternite VIII with posterior margin straight or slightly evenly dark brown, scutellum usually with the same emarginate medially, anterior sclerotization curved or color as elytra, sometimes with borders darker. Total bisinuous 0.47-0.85x the total width of sternite; sterlength 8.0-15.0 mm; elytral base 0.97-1.03x as wide nite IX with apex rounded.
</paragraph>
<caption id="A784B3E4243AFFB975E06662F6AFF8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641431" ID-Zenodo-Dep="12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="56" pageNumber="273" startId="56.[131,210,1800,1819]" targetBox="[133,1263,663,1773]" targetPageId="56" targetType="figure">
<paragraph id="F344E36C243AFFB975E06662F6AFF8F4" blockId="56.[131,1267,1800,1871]" pageId="56" pageNumber="273">
<emphasis id="C18F3F7E243AFFB975E06662F5EBF8A0" bold="true" box="[131,247,1800,1819]" pageId="56" pageNumber="273">FIGURE 26:</emphasis>
<emphasis id="C18F3F7E243AFFB9759C6663F481F8A0" box="[255,413,1801,1820]" italics="true" pageId="56" pageNumber="273">
<taxonomicName id="34FB98EF243AFFB9759C6663F485F8A0" authorityName="Fleutiaux" authorityYear="1892" box="[255,409,1801,1820]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="273" phylum="Arthropoda" rank="species" species="niger">Physodactylus niger</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E243AFFB974C06662F4AAF8A0" bold="true" box="[419,438,1800,1819]" pageId="56" pageNumber="273">A,</emphasis>
antenna;
<emphasis id="C18F3F7E243AFFB9776F6663F703F8A7" bold="true" box="[524,543,1801,1820]" pageId="56" pageNumber="273">B,</emphasis>
maxilla;
<emphasis id="C18F3F7E243AFFB9770C6662F7A6F8A7" bold="true" box="[623,698,1800,1820]" pageId="56" pageNumber="273">C, D, E,</emphasis>
head (dorsal, anterior, lateral);
<emphasis id="C18F3F7E243AFFB976A46663F10CF8A7" bold="true" box="[967,1040,1800,1820]" pageId="56" pageNumber="273">F, G, H,</emphasis>
prothorax (dorsal, ventral, lateral);
<emphasis id="C18F3F7E243AFFB975AB6648F5CAF88E" bold="true" box="[200,214,1826,1845]" pageId="56" pageNumber="273">I,</emphasis>
hypomeron and pronotum;
<emphasis id="C18F3F7E243AFFB974A66648F4CFF88E" bold="true" box="[453,467,1826,1845]" pageId="56" pageNumber="273">J,</emphasis>
mesoventrite;
<emphasis id="C18F3F7E243AFFB9772E6648F766F88E" bold="true" box="[589,634,1826,1845]" pageId="56" pageNumber="273">K, L,</emphasis>
mesoventrite and pleural sclerites (lateral, lateroventral);
<emphasis id="C18F3F7E243AFFB971376648F171F88E" bold="true" box="[1108,1133,1826,1845]" pageId="56" pageNumber="273">M,</emphasis>
scutellar shield;
<emphasis id="C18F3F7E243AFFB975E76656F5D5F8F4" bold="true" box="[132,201,1852,1871]" pageId="56" pageNumber="273">N, O, P,</emphasis>
pro-, meso- and metatibia;
<emphasis id="C18F3F7E243AFFB974CD6656F4D9F8F4" bold="true" box="[430,453,1852,1871]" pageId="56" pageNumber="273">Q,</emphasis>
hind wing;
<emphasis id="C18F3F7E243AFFB9774B6656F720F8F4" bold="true" box="[552,572,1852,1871]" pageId="56" pageNumber="273">R,</emphasis>
pterothorax and abdominal tergite I (dorsal).
</paragraph>
</caption>
<caption id="A784B3E4243BFFB8753C6266F151FCA4" ID-DOI="http://doi.org/10.5281/zenodo.12641433" ID-Zenodo-Dep="12641433" box="[95,1101,780,799]" httpUri="https://zenodo.org/record/12641433/files/figure.png" pageId="57" pageNumber="274" startId="57.[95,174,780,799]" targetBox="[98,1225,170,750]" targetPageId="57" targetType="figure">
<paragraph id="F344E36C243BFFB8753C6266F151FCA4" blockId="57.[94,1229,780,1872]" box="[95,1101,780,799]" pageId="57" pageNumber="274">
<emphasis id="C18F3F7E243BFFB8753C6266F5D3FCA4" bold="true" box="[95,207,780,799]" pageId="57" pageNumber="274">FIGURE 27:</emphasis>
<emphasis id="C18F3F7E243BFFB875B76266F46DFCA5" box="[212,369,780,799]" italics="true" pageId="57" pageNumber="274">
<taxonomicName id="34FB98EF243BFFB875B76266F471FCA5" authorityName="Fleutiaux" authorityYear="1892" box="[212,365,780,799]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="274" phylum="Arthropoda" rank="species" species="niger">Physodactylus niger</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E243BFFB874176266F483FCA4" bold="true" box="[372,415,780,799]" pageId="57" pageNumber="274">A, B,</emphasis>
aedeagus (dorsal, ventral).
<emphasis id="C18F3F7E243BFFB8771C6266F622FCA4" box="[639,830,780,799]" italics="true" pageId="57" pageNumber="274">
<taxonomicName id="34FB98EF243BFFB8771C6266F626FCA4" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[639,826,780,799]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="274" phylum="Arthropoda" rank="species" species="oberthuri">Physodactylus oberthuri</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E243BFFB876216266F66EFCA4" bold="true" box="[834,882,780,799]" pageId="57" pageNumber="274">C, D,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<paragraph id="F344E36C243BFFB8753D65B2F798FB54" blockId="57.[94,1229,780,1872]" box="[94,644,1240,1264]" pageId="57" pageNumber="274">as prothorax, elytra 2.73-3.10x longer than pronotum.</paragraph>
<paragraph id="F344E36C243BFFB8753D6592F624F98B" blockId="57.[94,1229,780,1872]" pageId="57" pageNumber="274">
Frons (
<figureCitation id="6BC0FFE9243BFFB875C66592F5E9FAAB" box="[165,245,1272,1296]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="57" pageNumber="274">Figs. 26</figureCitation>
C-E) flattened with a weakly to strong-
<emphasis id="C18F3F7E243BFFB877CB6592F61DFAAB" box="[680,769,1272,1296]" italics="true" pageId="57" pageNumber="274">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9243BFFB8766D6592F692FAAB" box="[782,910,1272,1296]" captionStart="FIGURE 27" captionStartId="57.[95,174,780,799]" captionTargetBox="[98,1225,170,750]" captionTargetId="figure-548@57.[94,1228,170,756]" captionTargetPageId="57" captionText="FIGURE 27: Physodactylus niger. A, B, aedeagus (dorsal, ventral). Physodactylus oberthuri. C, D, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641433" httpUri="https://zenodo.org/record/12641433/files/figure.png" pageId="57" pageNumber="274">Figs. 27A, B</figureCitation>
): Phallobase 0.41-0.49x the toly produced transverse protuberance between antennal tal length of aedeagus, 1.27-1.65x longer than wide, insertions, less prominent medially; frontal carina pro- length ratio between lateroposterior and median parts duced anteriorly; antenna (
<figureCitation id="6BC0FFE9243BFFB8741A6432F4CAFACB" box="[377,470,1368,1392]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="57" pageNumber="274">Fig. 26A</figureCitation>
) nearly reaching 5.7-10.7; paramere with ventral surface partly memto surpassing by one antennomere the half length of branous, except for the lateral half and the anterior pronotum; antennomere IV 1.1-1.2x wider than long, margin sclerotized; anteromedian sclerotized mar- V-VIII 1.5-1.6x wider than long; IX-X 1.2-1.3x wider gin usually contiguous and arch-shaped; penis with than long; XI oval 1.4-1.5x longer than wide. Prono- basal strut 0.23-0.27x its total length, articulated to tum (
<figureCitation id="6BC0FFE9243BFFB875FF6492F5E5F9AB" box="[156,249,1528,1552]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="57" pageNumber="274">Fig. 26F</figureCitation>
) 1.10-1.29x wider than long, lateral the parameres by a sclerotized process; ventral sclerite sides rounded from posterior angles to anterior mar- parallel-sided.
</paragraph>
<paragraph id="F344E36C243BFFB8753D6752F11CF8EB" blockId="57.[94,1229,780,1872]" pageId="57" pageNumber="274">
gin, strongly narrowed on anterior angle, anteromedian margin strongly produced (
<figureCitation id="6BC0FFE9243BFFB874D36732F70FF9CB" box="[432,531,1624,1648]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="57" pageNumber="274">Fig. 26H</figureCitation>
); posterior
<emphasis id="C18F3F7E243BFFB877CB6732F619F9CB" box="[680,773,1624,1648]" italics="true" pageId="57" pageNumber="274">
<typeStatus id="2C405DCE243BFFB877CB6732F61CF9CB" box="[680,768,1624,1648]" pageId="57" pageNumber="274" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Saint Paul], [
<taxonomicName id="34FB98EF243BFFB876F56732F1D0F9CB" authority=", Fleut., J" authorityName="Fleut., J" box="[918,1228,1623,1648]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="274" phylum="Arthropoda" rank="species" species="niger">Physodactylus niger, Fleut., J</taxonomicName>
angles elongate, convex, parallel to convergent; lateral Type,
<collectingCountry id="8BECA3FC243BFFB8778B6712F63BF92B" box="[744,807,1656,1680]" name="Brazil" pageId="57" pageNumber="274">Brèsil</collectingCountry>
], [Fleut. Ann. Soc. Ent. Fr. 1892, p. 407, carina prominent, nearly entirely visible dorsally, reach- t. 8, f. 3, Collection FLEUTIAUX], [niger], [Physoing or nearly reaching the anterior margin (
<figureCitation id="6BC0FFE9243BFFB8774267D2F76AF96B" box="[545,630,1720,1744]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="57" pageNumber="274">Fig. 26I</figureCitation>
); dactylus
<taxonomicName id="34FB98EF243BFFB8766B67D2F6AAF96B" authority="Fleut" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[776,950,1720,1744]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="283" phylum="Arthropoda" rank="species" species="oberthuri">Oberthuri Fleut</taxonomicName>
. comparé ou type, 1940, disc with punctures umbilicate, 1-2 or 2-4 diameters Collection FLEUTIAUX] [type], [nig. J (
<emphasis id="C18F3F7E243BFFB8710767B2F1D0F94B" box="[1124,1228,1752,1776]" italics="true" pageId="57" pageNumber="274">terminalia</emphasis>
apart, usually irregularly distributed, larger and denser
<emphasis id="C18F3F7E243BFFB877CB6792F69EF8AB" box="[680,898,1784,1808]" italics="true" pageId="57" pageNumber="274">and aedeagus attached</emphasis>
)] [Museum Paris, Coll. E. Fleuon lateral and anterior borders; prosternal process with tiaux], [
<taxonomicName id="34FB98EF243BFFB877986672F187F88B" authority="Fleut., J. Chassain" authorityName="Fleut., J. Chassain" box="[763,1179,1816,1840]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="274" phylum="Arthropoda" rank="species" species="niger">Physodactylus niger Fleut., J. Chassain</taxonomicName>
det. subapical tooth (
<figureCitation id="6BC0FFE9243BFFB874696652F48BF8EB" box="[266,407,1848,1872]" captionStart="FIGURE 26" captionStartId="56.[131,210,1800,1819]" captionTargetBox="[133,1263,663,1773]" captionTargetId="figure-197@56.[132,1266,662,1777]" captionTargetPageId="56" captionText="FIGURE 26: Physodactylus niger. A, antenna; B, maxilla; C, D, E, head (dorsal, anterior, lateral); F, G, H, prothorax (dorsal, ventral, lateral); I, hypomeron and pronotum; J, mesoventrite; K, L, mesoventrite and pleural sclerites (lateral, lateroventral); M, scutellar shield; N, O, P, pro-, meso- and metatibia; Q, hind wing; R, pterothorax and abdominal tergite I (dorsal)." figureDoi="http://doi.org/10.5281/zenodo.12641431" httpUri="https://zenodo.org/record/12641431/files/figure.png" pageId="57" pageNumber="274">Figs. 26G, H</figureCitation>
). Lamellae of pro- and 05], [
<typeStatus id="2C405DCE243BFFB877BC6652F654F8EB" box="[735,840,1848,1872]" pageId="57" pageNumber="274" type="holotype">Holotype</typeStatus>
], male (MNHN).
</paragraph>
<paragraph id="F344E36C2438FFBB75E761D2F784FECB" blockId="58.[132,681,184,368]" pageId="58" pageNumber="275">
<emphasis id="C18F3F7E2438FFBB75E761D2F458FF6B" box="[132,324,184,208]" italics="true" pageId="58" pageNumber="275">Material examined:</emphasis>
<materialsCitation id="4393E9312438FFBB742A61D2F784FECB" collectionCode="MNHN, MZUSP, CNC, IBSP, UNESP-IS" country="Brazil" location="Minas Gerais" pageId="58" pageNumber="275" specimenCount="80" stateProvince="Minas Gerais" typeStatus="holotype">
Without locality:
<specimenCount id="E5FD28E52438FFBB749E61D2F729FF74" box="[509,565,184,208]" count="2" pageId="58" pageNumber="275" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB772261D2F781FF6B" box="[577,669,184,208]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="275">MNHN</collectionCode>
);
<collectingCountry id="8BECA3FC2438FFBB75E761B2F5FFFF4B" box="[132,227,216,240]" name="Brazil" pageId="58" pageNumber="275">BRAZIL</collectingCountry>
.
<typeStatus id="2C405DCE2438FFBB758D61B2F453FF4B" box="[238,335,216,240]" pageId="58" pageNumber="275" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="34FB98EF2438FFBB741661B3F4D1FF4B" authorityName="Fleutiaux" authorityYear="1940" box="[373,461,217,240]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="carreti" status="syn. nov.">
<emphasis id="C18F3F7E2438FFBB741661B3F4D1FF4B" box="[373,461,217,240]" italics="true" pageId="58" pageNumber="275">P. carreti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052438FFBB74B661B2F72EFF54" box="[469,562,216,240]" pageId="58" pageNumber="275">syn. nov.</taxonomicNameLabel>
(
<collectionCode id="95EA7BA92438FFBB772261B2F781FF4B" box="[577,669,216,240]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="275">MNHN</collectionCode>
).
<emphasis id="C18F3F7E2438FFBB75E76193F409FEB4" box="[132,277,248,272]" italics="true" pageId="58" pageNumber="275">
<collectingRegion id="313F2D8E2438FFBB75E76193F413FEB4" box="[132,271,248,272]" country="Brazil" name="Minas Gerais" pageId="58" pageNumber="275">Minas Gerais</collectingRegion>
:
</emphasis>
<specimenCount id="E5FD28E52438FFBB74416192F47FFEB4" box="[290,355,248,272]" count="2" pageId="58" pageNumber="275" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB74146192F4CFFEAB" box="[375,467,248,272]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="275">MNHN</collectionCode>
).
<emphasis id="C18F3F7E2438FFBB748F6192F747FEAB" box="[492,603,248,272]" italics="true" pageId="58" pageNumber="275">
<collectingRegion id="313F2D8E2438FFBB748F6192F749FEAB" box="[492,597,248,272]" country="Brazil" name="Sao Paulo" pageId="58" pageNumber="275">São Paulo</collectingRegion>
:
</emphasis>
<specimenCount id="E5FD28E52438FFBB770B6192F7B5FEB4" box="[616,681,248,272]" count="3" pageId="58" pageNumber="275" type="generic">3 exs.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB75EF6072F5F4FE8B" box="[140,232,280,304]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="275">MNHN</collectionCode>
),
<specimenCount id="E5FD28E52438FFBB759D6072F456FE94" box="[254,330,280,304]" count="61" pageId="58" pageNumber="275" type="generic">61 exs.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB74386072F4A5FE8B" box="[347,441,280,304]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="58" pageNumber="275">MZUSP</collectionCode>
),
<specimenCount id="E5FD28E52438FFBB74AC6072F711FE94" box="[463,525,280,304]" count="3" pageId="58" pageNumber="275" type="generic">3 exs.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB777C6072F741FE8B" box="[543,605,280,304]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="58" pageNumber="275">CNC</collectionCode>
),
<specimenCount id="E5FD28E52438FFBB77176072F7B5FE94" box="[628,681,280,304]" count="1" pageId="58" pageNumber="275" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB75EF6052F5DAFEEB" box="[140,198,312,336]" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="58" pageNumber="275">IBSP</collectionCode>
).
<emphasis id="C18F3F7E2438FFBB75B56052F40FFEF4" box="[214,275,312,335]" italics="true" pageId="58" pageNumber="275">
<collectingRegion id="313F2D8E2438FFBB75B56052F413FEF4" box="[214,271,312,335]" country="Brazil" name="Goias" pageId="58" pageNumber="275">Goiás</collectingRegion>
:
</emphasis>
<specimenCount id="E5FD28E52438FFBB747A6052F454FEF4" box="[281,328,312,336]" count="1" pageId="58" pageNumber="275" type="generic">1 ex.</specimenCount>
(
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),
<specimenCount id="E5FD28E52438FFBB74A26052F4E5FEF4" box="[449,505,312,336]" count="3" pageId="58" pageNumber="275" type="generic">3 exs.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB77666052F778FEEB" box="[517,612,312,336]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="58" pageNumber="275">MZUSP</collectionCode>
).
<emphasis id="C18F3F7E2438FFBB77106053F413FECB" italics="true" pageId="58" pageNumber="275">
<collectingRegion id="313F2D8E2438FFBB77106053F415FECB" country="Brazil" name="Mato Grosso do Sul" pageId="58" pageNumber="275">Mato Grosso do Sul</collectingRegion>
:
</emphasis>
<specimenCount id="E5FD28E52438FFBB74766032F453FED4" box="[277,335,344,368]" count="2" pageId="58" pageNumber="275" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB743E6032F4A7FECB" box="[349,443,344,368]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="58" pageNumber="275">MZUSP</collectionCode>
),
<specimenCount id="E5FD28E52438FFBB74AE6032F71BFED4" box="[461,519,344,368]" count="2" pageId="58" pageNumber="275" type="generic">2 exs.</specimenCount>
(
<collectionCode id="95EA7BA92438FFBB77776032F791FECB" box="[532,653,344,368]" pageId="58" pageNumber="275">UNESP-IS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C2438FFBB75E760F2F448FDEB" blockId="58.[132,681,408,592]" pageId="58" pageNumber="275">
<emphasis id="C18F3F7E2438FFBB75E760F2F41AFE0B" box="[132,262,408,432]" italics="true" pageId="58" pageNumber="275">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2438FFBB746C60F2F472FE0B" box="[271,366,408,432]" name="Brazil" pageId="58" pageNumber="275">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2438FFBB741A60F2F4AAFE14" box="[377,438,408,431]" italics="true" pageId="58" pageNumber="275">
<collectingRegion id="313F2D8E2438FFBB741A60F2F4AEFE14" box="[377,434,408,431]" country="Brazil" name="Goias" pageId="58" pageNumber="275">Goiás</collectingRegion>
:
</emphasis>
Jataí, Rio Verde;
<emphasis id="C18F3F7E2438FFBB771060F3F400FE6B" italics="true" pageId="58" pageNumber="275">
<collectingRegion id="313F2D8E2438FFBB771060F3F40AFE6B" country="Brazil" name="Mato Grosso do Sul" pageId="58" pageNumber="275">Mato Grosso do Sul</collectingRegion>
:
</emphasis>
Três Lagoas, Selvíria;
<emphasis id="C18F3F7E2438FFBB777B60D3F7B4FE74" box="[536,680,440,464]" italics="true" pageId="58" pageNumber="275">
<collectingRegion id="313F2D8E2438FFBB777B60D3F7BFFE74" box="[536,675,440,464]" country="Brazil" name="Minas Gerais" pageId="58" pageNumber="275">Minas Gerais</collectingRegion>
:
</emphasis>
Uberaba;
<emphasis id="C18F3F7E2438FFBB758C60B2F442FE4B" box="[239,350,472,496]" italics="true" pageId="58" pageNumber="275">
<collectingRegion id="313F2D8E2438FFBB758C60B2F444FE4B" box="[239,344,472,496]" country="Brazil" name="Sao Paulo" pageId="58" pageNumber="275">São Paulo</collectingRegion>
:
</emphasis>
Ribeirão Preto, Mogi Guaçu, Araras, Orlândia, Sud Menucci, Pradópolis, São José do Rio Preto, Pirassununga, Assis, Magda, Piracicaba, Guatapará, Batatais.
</paragraph>
<paragraph id="F344E36C2438FFBB75E76312F703FA0B" blockId="58.[132,681,632,1872]" pageId="58" pageNumber="275">
<emphasis id="C18F3F7E2438FFBB75E76312F5C2FD2B" box="[132,222,632,656]" italics="true" pageId="58" pageNumber="275">Remarks:</emphasis>
<bibRefCitation id="976A9E9D2438FFBB75866312F48EFD2B" author="FLEUTIAUX, E." box="[229,402,632,656]" pageId="58" pageNumber="275" pagination="403 - 412" refId="ref49699" refString="FLEUTIAUX, E. 1892. Note sur les Physodactylini. Annales de La Societe Entomologique de France, 61: 403 - 412." type="journal article" year="1892">Fleutiaux (1892)</bibRefCitation>
described
<taxonomicName id="34FB98EF2438FFBB77636312F5BDFD0B" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2438FFBB77636312F5BDFD0B" italics="true" pageId="58" pageNumber="275">Physodactylus niger</emphasis>
</taxonomicName>
as similar to
<taxonomicName id="34FB98EF2438FFBB744B63F3F487FD0B" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[296,411,664,688]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2438FFBB744B63F3F487FD0B" box="[296,411,664,688]" italics="true" pageId="58" pageNumber="275">P. oberthuri</emphasis>
</taxonomicName>
and distinguished it by its smaller size, the black, narrower and less convex body, the pronotum more strongly punctate with anterior margin more widely rounded. In 1940, Fleutiaux considered these features insufficient to separate those species and synonymized
<taxonomicName id="34FB98EF2438FFBB74F16253F4C1FCEB" box="[402,477,825,848]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2438FFBB74F16253F4C1FCEB" box="[402,477,825,848]" italics="true" pageId="58" pageNumber="275">P. niger</emphasis>
</taxonomicName>
under
<emphasis id="C18F3F7E2438FFBB774F6253F7B5FCEB" box="[556,681,824,848]" italics="true" pageId="58" pageNumber="275">
<taxonomicName id="34FB98EF2438FFBB774F6253F7B9FCEB" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[556,677,824,848]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="oberthuri">P. oberthuri</taxonomicName>
.
</emphasis>
The analysis of a larger series of specimens revealed that
<taxonomicName id="34FB98EF2438FFBB75D46213F41EFC2B" box="[183,258,889,912]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2438FFBB75D46213F41EFC2B" box="[183,258,889,912]" italics="true" pageId="58" pageNumber="275">P. niger</emphasis>
</taxonomicName>
is a valid species that exhibit polymorphisms in color pattern, size and punctation among specimens from same region. Nevertheless, this species can be distinguished from other
<taxonomicName id="34FB98EF2438FFBB774762B2F5D5FBAB" pageId="58" pageNumber="275">
<emphasis id="C18F3F7E2438FFBB774762B2F7B4FC4B" box="[548,680,984,1008]" italics="true" pageId="58" pageNumber="275">Physodactylus</emphasis>
species
</taxonomicName>
by the following combination of characters: general pilosity yellow, frons with a transverse protuberance, frontal carina produced anteriorly; prothorax strongly and abruptly narrowed anteriorly with lateral carina prominent, pronotal anteromedian margin produced over head, pronotal punctation heterogeneously distributed; lamella of mesotarsomere I absent; abdominal ventrites weakly convex, ventrite 1 with setae fine, half shorter and sparser them those of ventrite 2, ventrites 2-4 evenly pilose.
<taxonomicName id="34FB98EF2438FFBB77476472F5A9FAEB" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2438FFBB77476472F5A9FAEB" italics="true" pageId="58" pageNumber="275">Physodactylus niger</emphasis>
</taxonomicName>
can be promptly distinguished from
<taxonomicName id="34FB98EF2438FFBB77566453F7B5FAEB" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[565,681,1336,1360]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2438FFBB77566453F7B5FAEB" box="[565,681,1336,1360]" italics="true" pageId="58" pageNumber="275">P. oberthuri</emphasis>
</taxonomicName>
by its larger and denser pronotal punctation, yellow pilosity and convex abdomen (brown pilosity and laterally concave abdomen in
<taxonomicName id="34FB98EF2438FFBB74FF64F3F713FA0B" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[412,527,1432,1456]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2438FFBB74FF64F3F713FA0B" box="[412,527,1432,1456]" italics="true" pageId="58" pageNumber="275">P. oberthuri</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="F344E36C2438FFBB75DE64D2F1CAFEEB" blockId="58.[132,681,632,1872]" lastBlockId="58.[718,1267,184,336]" pageId="58" pageNumber="275">
The
<typeStatus id="2C405DCE2438FFBB759264D2F457FA6B" box="[241,331,1464,1488]" pageId="58" pageNumber="275" type="holotype">holotype</typeStatus>
of
<taxonomicName id="34FB98EF2438FFBB741764D2F75DFA6B" authorityName="Fleutiaux" authorityYear="1940" box="[372,577,1464,1488]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="carreti" status="syn. nov.">
<emphasis id="C18F3F7E2438FFBB741764D2F75DFA6B" box="[372,577,1464,1488]" italics="true" pageId="58" pageNumber="275">Physodactylus carreti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052438FFBB772864D2F7B5FA74" box="[587,681,1464,1488]" pageId="58" pageNumber="275">syn. nov.</taxonomicNameLabel>
has color pattern 3, punctation 1-2 diameters apart on pronotal disc and antenna nearly reaching the base of pronotum; its aedeagus has the apex broken but does not present any consistent difference from those of other
<taxonomicName id="34FB98EF2438FFBB75BD6733F43BF9CB" box="[222,295,1625,1648]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="275" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2438FFBB75BD6733F43BF9CB" box="[222,295,1625,1648]" italics="true" pageId="58" pageNumber="275">P. niger</emphasis>
</taxonomicName>
specimens. The type of
<taxonomicName id="34FB98EF2438FFBB77456733F40EF92B" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="273" phylum="Arthropoda" rank="variety" species="carreti" status="syn. nov." variety="testaceipennis">
<emphasis id="C18F3F7E2438FFBB77456733F761F9CB" box="[550,637,1625,1648]" italics="true" pageId="58" pageNumber="275">P. carreti</emphasis>
var. testaceipennis
</taxonomicName>
<taxonomicNameLabel id="DABC82052438FFBB74786712F464F934" box="[283,376,1656,1680]" pageId="58" pageNumber="275">syn. nov.</taxonomicNameLabel>
has the color pattern 6, pronotal punctation more sparsely distributed (2-4 diameters apart on pronotal disc). Specimens collected at Ribeirão Preto and Mogi Guaçu exhibit all color patterns. The specimens from Sud Menucci and São José do Rio Preto are the smallest, while those from Rio Verde, Jataí and Três Lagoas are the largest, representing the extreme sizes of a morphocline in body length. The specimens from Três Lagoas have the anteromedian sclerotized margin of parameres more widely separate. Some specimens from other localities also present this margin separate in variable width.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,516 @@
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<mods:title id="CFC6AED4E9E36EE1E2A0F1F587DFB9D4">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<paragraph id="F344E36C243CFFBF764165BDF183FB4B" blockId="62.[802,1183,1239,1296]" box="[802,1183,1239,1264]" pageId="62" pageNumber="279">
<heading id="A80C5400243CFFBF764165BDF183FB4B" bold="true" box="[802,1183,1239,1264]" centered="true" fontSize="10" level="2" pageId="62" pageNumber="279" reason="2">
<emphasis id="C18F3F7E243CFFBF764165BDF183FB4B" bold="true" box="[802,1183,1239,1264]" pageId="62" pageNumber="279">
<taxonomicName id="34FB98EF243CFFBF764165BDF15AFB54" authorityName="Rosa" authorityYear="2014" box="[802,1094,1239,1263]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="C18F3F7E243CFFBF764165BDF15AFB54" bold="true" box="[802,1094,1239,1263]" italics="true" pageId="62" pageNumber="279">Physodactylus tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243CFFBF712F65B2F183FB4B" box="[1100,1183,1240,1264]" pageId="62" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="BBE1B0E7243CFFC576EC6592F4DFFD54" lastPageId="68" lastPageNumber="285" pageId="62" pageNumber="279" type="description">
<paragraph id="F344E36C243CFFBF76EC6592F12DFAAB" blockId="62.[802,1183,1239,1296]" box="[911,1073,1272,1296]" pageId="62" pageNumber="279">
<emphasis id="C18F3F7E243CFFBF76EC6592F12DFAAB" bold="true" box="[911,1073,1272,1296]" pageId="62" pageNumber="279">
(
<figureCitation id="6BC0FFE9243CFFBF76F56592F6F3FAAB" box="[918,1007,1272,1296]" captionStart="FIGURE 31" captionStartId="63.[95,174,647,666]" captionTargetBox="[95,1224,173,620]" captionTargetId="figure-117@63.[94,1228,170,624]" captionTargetPageId="63" captionText="FIGURE 31: Physodactylus tuberculatus sp. nov. A, B, prothorax (dorsal, lateral); C, metatibia; D, abdomen (ventral); E, sternite VIII; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641443" httpUri="https://zenodo.org/record/12641443/files/figure.png" pageId="62" pageNumber="279">Figs. 31</figureCitation>
,
<figureCitation id="6BC0FFE9243CFFBF76986592F136FAAB" box="[1019,1066,1272,1296]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="62" pageNumber="279">35R</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="F344E36C243CFFBF77AD6452F7E6FA2B" blockId="62.[718,1266,1336,1424]" pageId="62" pageNumber="279">
<emphasis id="C18F3F7E243CFFBF77AD6452F625FAEB" box="[718,825,1336,1360]" italics="true" pageId="62" pageNumber="279">Etymology:</emphasis>
From Latin,
<emphasis id="C18F3F7E243CFFBF76A46452F124FAEB" box="[967,1080,1336,1360]" italics="true" pageId="62" pageNumber="279">tuberculum</emphasis>
= tubercle; alluding to the conspicuous pronotal tubercle of this species.
</paragraph>
<paragraph id="F344E36C243CFFC177AD64D3F6A4FE2B" blockId="62.[718,1267,1464,1872]" lastBlockId="64.[718,1267,184,400]" lastPageId="64" lastPageNumber="281" pageId="62" pageNumber="279">
<emphasis id="C18F3F7E243CFFBF77AD64D3F622FA6B" box="[718,830,1465,1488]" italics="true" pageId="62" pageNumber="279">Description</emphasis>
(male
<figureCitation id="6BC0FFE9243CFFBF76EB64D2F6FAFA6B" box="[904,998,1464,1488]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="62" pageNumber="279">Fig. 35R</figureCitation>
): Integument very bright, dark brown with elytra lighter or black with elytra dark reddish-brown and epipleura lighter; pilosity yellow. Total length 11.0-
<quantity id="34034E89243CFFBF71646772F172F994" box="[1031,1134,1560,1584]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="62" pageNumber="279" unit="mm" value="13.5">13.5 mm</quantity>
; elytral base 0.93-1.05x as wide as prothorax, elytra 2.73-3.03x times longer than pronotum. Frons with a transverse or circular protuberance medially; frontal carina strongly produced anteriorly; antenna nearly reaching the half length of pronotum; antennomere IV and V 1.4-1.5x wider than long, VI-VIII 1.9-2.0x wider than long; IX-X 1.6x wider than long; XI subcircular as wide as long. Pronotum (
<figureCitation id="6BC0FFE9243CFFBF71766672F16DF88B" box="[1045,1137,1816,1840]" captionStart="FIGURE 31" captionStartId="63.[95,174,647,666]" captionTargetBox="[95,1224,173,620]" captionTargetId="figure-117@63.[94,1228,170,624]" captionTargetPageId="63" captionText="FIGURE 31: Physodactylus tuberculatus sp. nov. A, B, prothorax (dorsal, lateral); C, metatibia; D, abdomen (ventral); E, sternite VIII; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641443" httpUri="https://zenodo.org/record/12641443/files/figure.png" pageId="62" pageNumber="279">Fig. 31A</figureCitation>
) 1.18-1.21x wider than long, lateral sides rounded from posterior angles to anterior margin, strongly narrowed on anterior angles, anteromedian margin strongly produced; posterior third at middle with a small spiniform tubercle (
<figureCitation id="6BC0FFE9243DFFBE75CF6252F417FCEB" box="[172,267,824,848]" captionStart="FIGURE 31" captionStartId="63.[95,174,647,666]" captionTargetBox="[95,1224,173,620]" captionTargetId="figure-117@63.[94,1228,170,624]" captionTargetPageId="63" captionText="FIGURE 31: Physodactylus tuberculatus sp. nov. A, B, prothorax (dorsal, lateral); C, metatibia; D, abdomen (ventral); E, sternite VIII; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641443" httpUri="https://zenodo.org/record/12641443/files/figure.png" pageId="63" pageNumber="280">Fig. 31B</figureCitation>
); posterior angles elongate, convex, parallel; lateral carina complete, strongly prominent on posterior half, not visible dorsally on anterior half; disc and lateral margin with punctures 1-2 diameters apart, larger on anterior angle, homogenously distributed; prosternal process without subapical tooth. Lamellae of pro- and mesotarsomeres I smaller than the others, present on metatarsomere I. Elytra tapering from base to apex; striae apical striae with a row of punctures about 3 times larger than those of pronotal disc; interstices flat, slightly rugose, sparsely punctate. Abdomen (
<figureCitation id="6BC0FFE92442FFC1740F61D2F4CCFF6B" box="[364,464,184,208]" captionStart="FIGURE 31" captionStartId="63.[95,174,647,666]" captionTargetBox="[95,1224,173,620]" captionTargetId="figure-117@63.[94,1228,170,624]" captionTargetPageId="63" captionText="FIGURE 31: Physodactylus tuberculatus sp. nov. A, B, prothorax (dorsal, lateral); C, metatibia; D, abdomen (ventral); E, sternite VIII; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641443" httpUri="https://zenodo.org/record/12641443/files/figure.png" pageId="64" pageNumber="281">Fig. 31D</figureCitation>
) with ventrites 2-3 concave laterally, pilosity of the inner 2/3 (0.6-0.7) of the ventrites 1-3 separated from those of lateral bor- der by a glabrous and impunctate longitudinal band; ventrite 1 with short setae on inner third and twice longer, denser and stouter setae on lateral border; ventrites 2-3 with setae semi-erect twice longer, denser and stouter than those of the inner part of ventrite 1, sparser on lateral border; ventrite 4 with setae spars- er, longer and stouter than those of the inner part of ventrite 3, denser on lateral border; ventrite 5 with median setae sparse, denser on borders. Pregenitalic segments and aedeagus covered with yellow setae. Sternite VIII subrectangular with anterior margin and anterior sclerotization strongly curved, posterior margin slightly to strongly emarginate (
<figureCitation id="6BC0FFE92442FFC171566052F193FEEB" box="[1077,1167,312,336]" captionStart="FIGURE 31" captionStartId="63.[95,174,647,666]" captionTargetBox="[95,1224,173,620]" captionTargetId="figure-117@63.[94,1228,170,624]" captionTargetPageId="63" captionText="FIGURE 31: Physodactylus tuberculatus sp. nov. A, B, prothorax (dorsal, lateral); C, metatibia; D, abdomen (ventral); E, sternite VIII; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641443" httpUri="https://zenodo.org/record/12641443/files/figure.png" pageId="64" pageNumber="281">Fig. 31E</figureCitation>
), anterior sclerotization 0.4x the total width of sternite; sternite IX with apex rounded.
</paragraph>
<caption id="A784B3E4243DFFBE753C63EDF47FFD0F" ID-DOI="http://doi.org/10.5281/zenodo.12641443" ID-Zenodo-Dep="12641443" httpUri="https://zenodo.org/record/12641443/files/figure.png" pageId="63" pageNumber="280" startId="63.[95,174,647,666]" targetBox="[95,1224,173,620]" targetPageId="63" targetType="figure">
<paragraph id="F344E36C243DFFBE753C63EDF47FFD0F" blockId="63.[94,1229,647,692]" pageId="63" pageNumber="280">
<emphasis id="C18F3F7E243DFFBE753C63EDF5CDFD21" bold="true" box="[95,209,647,666]" pageId="63" pageNumber="280">FIGURE 31:</emphasis>
<taxonomicName id="34FB98EF243DFFBE75BB63EDF4B5FD21" authorityName="Rosa" authorityYear="2014" box="[216,425,647,666]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="280" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="C18F3F7E243DFFBE75BB63EDF4B5FD21" box="[216,425,647,666]" italics="true" pageId="63" pageNumber="280">Physodactylus tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243DFFBE74D363E2F4EEFD20" box="[432,498,648,667]" pageId="63" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E243DFFBE749463EDF738FD20" bold="true" box="[503,548,647,667]" pageId="63" pageNumber="280">A, B,</emphasis>
prothorax (dorsal, lateral);
<emphasis id="C18F3F7E243DFFBE766E63EDF63EFD21" bold="true" box="[781,802,647,666]" pageId="63" pageNumber="280">C,</emphasis>
metatibia;
<emphasis id="C18F3F7E243DFFBE76E163E2F685FD20" bold="true" box="[898,921,648,667]" pageId="63" pageNumber="280">D,</emphasis>
abdomen (ventral);
<emphasis id="C18F3F7E243DFFBE712563E2F144FD20" bold="true" box="[1094,1112,648,667]" pageId="63" pageNumber="280">E,</emphasis>
sternite VIII;
<emphasis id="C18F3F7E243DFFBE753D63CBF594FD0F" bold="true" box="[94,136,673,692]" pageId="63" pageNumber="280">F, G,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<caption id="A784B3E4243DFFBE753C6648F6A2F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641449" ID-Zenodo-Dep="12641449" httpUri="https://zenodo.org/record/12641449/files/figure.png" pageId="63" pageNumber="280" startId="63.[95,174,1826,1845]" targetBox="[94,1228,971,1802]" targetPageId="63" targetType="figure">
<paragraph id="F344E36C243DFFBE753C6648F6A2F8F4" blockId="63.[94,1229,1826,1871]" pageId="63" pageNumber="280">
<emphasis id="C18F3F7E243DFFBE753C6648F5CCF88E" bold="true" box="[95,208,1826,1845]" pageId="63" pageNumber="280">FIGURE 32:</emphasis>
<emphasis id="C18F3F7E243DFFBE75B46648F46FF88E" box="[215,371,1826,1845]" italics="true" pageId="63" pageNumber="280">
<taxonomicName id="34FB98EF243DFFBE75B46648F473F88E" authorityName="J. Chassain" baseAuthorityName="Lucas" baseAuthorityYear="1857" box="[215,367,1826,1845]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="280" phylum="Arthropoda" rank="species" species="femoralis">Teslasena femoralis</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E243DFFBE741B6648F497F88E" bold="true" box="[376,395,1826,1845]" pageId="63" pageNumber="280">A,</emphasis>
maxilla;
<emphasis id="C18F3F7E243DFFBE74BA6648F4F7F88E" bold="true" box="[473,491,1826,1845]" pageId="63" pageNumber="280">B,</emphasis>
detail of maxillar base;
<emphasis id="C18F3F7E243DFFBE77D66648F7D6F88E" bold="true" box="[693,714,1826,1845]" pageId="63" pageNumber="280">C,</emphasis>
labium;
<emphasis id="C18F3F7E243DFFBE76756648F631F88E" bold="true" box="[790,813,1826,1845]" pageId="63" pageNumber="280">D,</emphasis>
pronotum (laterodorsal);
<emphasis id="C18F3F7E243DFFBE71646648F105F88E" bold="true" box="[1031,1049,1826,1845]" pageId="63" pageNumber="280">E,</emphasis>
pronotal punctation;
<emphasis id="C18F3F7E243DFFBE753D6656F573F8F4" bold="true" box="[94,111,1852,1871]" pageId="63" pageNumber="280">F,</emphasis>
elytral striae;
<emphasis id="C18F3F7E243DFFBE75836656F5EAF8F4" bold="true" box="[224,246,1852,1871]" pageId="63" pageNumber="280">G,</emphasis>
elytron (dorsolateral);
<emphasis id="C18F3F7E243DFFBE74D26656F4F2F8F4" bold="true" box="[433,494,1852,1871]" pageId="63" pageNumber="280">H, I, J,</emphasis>
pro-, meso and metathoracic legs;
<emphasis id="C18F3F7E243DFFBE766E6656F63EF8F4" bold="true" box="[781,802,1852,1871]" pageId="63" pageNumber="280">K,</emphasis>
protarsus;
<emphasis id="C18F3F7E243DFFBE761E6656F692F8F4" bold="true" box="[893,910,1852,1871]" pageId="63" pageNumber="280">L,</emphasis>
claw.
</paragraph>
</caption>
<paragraph id="F344E36C2442FFC077AD60D2F718FECB" blockId="64.[718,1266,440,528]" lastBlockId="65.[94,643,184,368]" lastPageId="65" lastPageNumber="282" pageId="64" pageNumber="281">
<emphasis id="C18F3F7E2442FFC177AD60D2F63BFE6B" box="[718,807,440,464]" italics="true" pageId="64" pageNumber="281">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE92442FFC1765A60D2F6DFFE6B" box="[825,963,440,464]" captionStart="FIGURE 31" captionStartId="63.[95,174,647,666]" captionTargetBox="[95,1224,173,620]" captionTargetId="figure-117@63.[94,1228,170,624]" captionTargetPageId="63" captionText="FIGURE 31: Physodactylus tuberculatus sp. nov. A, B, prothorax (dorsal, lateral); C, metatibia; D, abdomen (ventral); E, sternite VIII; F, G, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641443" httpUri="https://zenodo.org/record/12641443/files/figure.png" pageId="64" pageNumber="281">Figs. 31F, G</figureCitation>
): Phallobase 0.47x the total length of aedeagus, 1.36x longer than wide, length ratio between lateroposterior and median parts 3.3-4.8; paramere with ventral surface predominantly membranous, except for the lateral and the anterior margin sclerotized; anteromedian sclerotized margin approximate at middle; penis with basal strut 0.32x its total length, articulated to the parameres by a translucent membrane; ventral sclerite parallel-sided.
</paragraph>
<caption id="A784B3E42442FFC175E76662F193F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641451" ID-Zenodo-Dep="12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="64" pageNumber="281" startId="64.[132,211,1800,1819]" targetBox="[132,1266,567,1776]" targetPageId="64" targetType="figure">
<paragraph id="F344E36C2442FFC175E76662F193F8F4" blockId="64.[132,1268,1800,1871]" pageId="64" pageNumber="281">
<emphasis id="C18F3F7E2442FFC175E76662F5E9F8A0" bold="true" box="[132,245,1800,1819]" pageId="64" pageNumber="281">FIGURE 33:</emphasis>
<taxonomicName id="34FB98EF2442FFC175986663F487F8A7" authorityName="Rosa" authorityYear="2014" box="[251,411,1801,1820]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="281" phylum="Arthropoda" rank="species" species="hirtus" status="sp. nov.">
<emphasis id="C18F3F7E2442FFC175986663F487F8A7" box="[251,411,1801,1820]" italics="true" pageId="64" pageNumber="281">Dactylophysus hirtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052442FFC174C26663F4FDF8A7" box="[417,481,1801,1820]" pageId="64" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C18F3F7E2442FFC174866662F736F8A0" bold="true" box="[485,554,1800,1820]" pageId="64" pageNumber="281">A, B, C,</emphasis>
pro-, meso- and metathoracic legs;
<emphasis id="C18F3F7E2442FFC176326663F674F8A7" bold="true" box="[849,872,1801,1820]" pageId="64" pageNumber="281">D,</emphasis>
apex of protibia.
<emphasis id="C18F3F7E2442FFC1769A6663F18AF8A0" box="[1017,1174,1801,1820]" italics="true" pageId="64" pageNumber="281">
<taxonomicName id="34FB98EF2442FFC1769A6663F18EF8A0" box="[1017,1170,1801,1820]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="281" phylum="Arthropoda" rank="species" species="niger">Physodactylus niger</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2442FFC171F96663F1B0F8A7" bold="true" box="[1178,1196,1801,1820]" pageId="64" pageNumber="281">E,</emphasis>
labrum. (lateroanterior).
<emphasis id="C18F3F7E2442FFC174686648F4DBF88E" box="[267,455,1826,1845]" italics="true" pageId="64" pageNumber="281">
<taxonomicName id="34FB98EF2442FFC174686648F4DEF88E" authorityName="Fischer von Waldheim" authorityYear="1823" box="[267,450,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="281" phylum="Arthropoda" rank="species" species="henningi">Physodactylus henningi</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2442FFC174A96648F4C7F88E" bold="true" box="[458,475,1826,1845]" pageId="64" pageNumber="281">F,</emphasis>
prothoracic leg;
<emphasis id="C18F3F7E2442FFC177016648F764F88E" bold="true" box="[610,632,1826,1845]" pageId="64" pageNumber="281">G,</emphasis>
protibia;
<emphasis id="C18F3F7E2442FFC177AB6648F7C3F88E" bold="true" box="[712,735,1826,1845]" pageId="64" pageNumber="281">H,</emphasis>
mesothoracic leg;
<emphasis id="C18F3F7E2442FFC176156648F698F88E" bold="true" box="[886,900,1826,1845]" pageId="64" pageNumber="281">I,</emphasis>
mesotibia;
<emphasis id="C18F3F7E2442FFC176816648F6ECF88E" bold="true" box="[994,1008,1826,1845]" pageId="64" pageNumber="281">J,</emphasis>
metathoracic leg;
<emphasis id="C18F3F7E2442FFC171E56648F187F88E" bold="true" box="[1158,1179,1826,1845]" pageId="64" pageNumber="281">K,</emphasis>
metatibia;
<emphasis id="C18F3F7E2442FFC175E76656F58AF8F4" bold="true" box="[132,150,1852,1871]" pageId="64" pageNumber="281">L,</emphasis>
apex of metatibia;
<emphasis id="C18F3F7E2442FFC174576656F450F8F4" bold="true" box="[308,332,1852,1871]" pageId="64" pageNumber="281">M,</emphasis>
lamella of protarsomere I.
<emphasis id="C18F3F7E2442FFC177486656F7DBF8F5" box="[555,711,1852,1871]" italics="true" pageId="64" pageNumber="281">
<taxonomicName id="34FB98EF2442FFC177486656F7DFF8F5" box="[555,707,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="281" phylum="Arthropoda" rank="species" species="niger">Physodactylus niger</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2442FFC177A86656F7FDF8F4" bold="true" box="[715,737,1852,1871]" pageId="64" pageNumber="281">N,</emphasis>
elytral striae.
<emphasis id="C18F3F7E2442FFC176306656F11AF8F4" box="[851,1030,1852,1871]" italics="true" pageId="64" pageNumber="281">
<taxonomicName id="34FB98EF2442FFC176306656F11DF8F4" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[851,1025,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="281" phylum="Arthropoda" rank="species" species="sulcatus">Physodactylus sulcatus</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E2442FFC1716A6656F13CF8F4" bold="true" box="[1033,1056,1852,1871]" pageId="64" pageNumber="281">O,</emphasis>
elytral striae.
</paragraph>
</caption>
<paragraph id="F344E36C2443FFC3753D60F2F5EDFF4B" blockId="65.[94,643,408,496]" lastBlockId="66.[132,681,184,240]" lastPageId="66" lastPageNumber="283" pageId="65" pageNumber="282">
<emphasis id="C18F3F7E2443FFC0753D60F2F5A7FE0B" box="[94,187,408,432]" italics="true" pageId="65" pageNumber="282">
<typeStatus id="2C405DCE2443FFC0753D60F2F5AAFE0B" box="[94,182,408,432]" pageId="65" pageNumber="282" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Gorham Collection acc69966], [
<taxonomicName id="34FB98EF2443FFC0777460F2F58DFE6B" authorityName="Fischer von Waldheim" authorityYear="1823" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="genus">Physodactylus</taxonomicName>
, nov (
<emphasis id="C18F3F7E2443FFC075AD60D2F406FE6B" box="[206,282,440,464]" italics="true" pageId="65" pageNumber="282">illegible</emphasis>
),
<collectingCountry id="8BECA3FC2443FFC0744E60D2F471FE6B" box="[301,365,440,464]" name="Brazil" pageId="65" pageNumber="282">Brazil</collectingCountry>
], [
<taxonomicName id="34FB98EF2443FFC074E160D2F763FE6B" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[386,639,440,464]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="oberthuri">Physodactylus oberthuri</taxonomicName>
, comparado ao tipo, C. Costa 1995] male, (MZUSP).
<emphasis id="C18F3F7E2443FFC077CB61D3F610FF6B" box="[680,780,185,208]" italics="true" pageId="65" pageNumber="282">
<typeStatus id="2C405DCE2443FFC077CB61D3F61BFF6B" box="[680,775,185,208]" pageId="65" pageNumber="282" type="paratype">Paratypes</typeStatus>
:
</emphasis>
[
<collectingCountry id="8BECA3FC2443FFC0767961D2F641FF6B" box="[794,861,184,208]" name="Brazil" pageId="65" pageNumber="282">Brésil</collectingCountry>
], [Collection Bonvouloir], [Fleutx. 1891], [
<taxonomicName id="34FB98EF2443FFC0766B61B2F7E5FEAB" authority=", Fleut, FLEUTIAUX" authorityName="Fleut, FLEUTIAUX" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="oberthuri">Physodactylus oberthuri, Fleut, FLEUTIAUX</taxonomicName>
det. 1944], [Museum Paris, ex Coll. R. Oberthur], [Museum Paris Coll. E. Fleutiaux], [
<taxonomicName id="34FB98EF2443FFC071FB6072F684FEEB" authority="Fleut." authorityName="Fleutiaux" authorityYear="1892" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="niger">Physodactylus niger Fleut.</taxonomicName>
?, J. Chassain det. 05]
<specimenCount id="E5FD28E52443FFC071F16052F1D4FEF4" box="[1170,1224,312,336]" count="1" pageId="65" pageNumber="282" type="generic">1 ex.</specimenCount>
, (MNHN); [Lavras,
<collectingRegion id="313F2D8E2443FFC076F76032F133FECB" box="[916,1071,344,368]" country="Brazil" name="Minas Gerais" pageId="65" pageNumber="282">Minas Gerais</collectingRegion>
,
<date id="8745C5AC2443FFC071246032F1D4FECB" box="[1095,1224,344,368]" pageId="65" pageNumber="282" value="1988-12-13">13/12/1988</date>
, R.C. Garcia], [
<taxonomicName id="34FB98EF2443FFC0763E6012F7F6FE0B" authority="(Fleut., 1892)" baseAuthorityName="Fleut." baseAuthorityYear="1892" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="oberthuri">Physodactylus oberthuri (Fleut., 1892)</taxonomicName>
, S.A.C. Chen det. 1989],
<specimenCount id="E5FD28E52443FFC0717460F2F152FE14" box="[1047,1102,408,432]" count="1" pageId="65" pageNumber="282" type="generic">1 ex.</specimenCount>
(MZUSP); [Minas Geraes], [Fry Coll., 1905.100.], [486ª],
<specimenCount id="E5FD28E52443FFC077CB60B2F7FEFE54" box="[680,738,472,496]" count="1" pageId="65" pageNumber="282" type="generic">1 ex.</specimenCount>
(BMNH), idem plus [J], [486ª], [3681], [
<taxonomicName id="34FB98EF2440FFC375EE61D3F454FF6B" authority="Fleut" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[141,328,184,208]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2440FFC375EE61D3F41AFF6B" box="[141,262,184,208]" italics="true" pageId="66" pageNumber="283">P. oberthuri</emphasis>
Fleut
</taxonomicName>
, ex desc. Det. K.G. Blair],
<specimenCount id="E5FD28E52440FFC3771761D2F7B5FF74" box="[628,681,184,208]" count="1" pageId="66" pageNumber="283" type="generic">1 ex.</specimenCount>
(BMNH).
</paragraph>
<caption id="A784B3E42443FFC0753C67BFF664F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641453" ID-Zenodo-Dep="12641453" httpUri="https://zenodo.org/record/12641453/files/figure.png" pageId="65" pageNumber="282" startId="65.[95,174,1749,1768]" targetBox="[94,1227,535,1726]" targetPageId="65" targetType="figure">
<paragraph id="F344E36C2443FFC0753C67BFF664F8F4" blockId="65.[94,1230,1749,1871]" pageId="65" pageNumber="282">
<emphasis id="C18F3F7E2443FFC0753C67BFF5CDF953" bold="true" box="[95,209,1749,1768]" pageId="65" pageNumber="282">FIGURE 34:</emphasis>
Habitus.
<emphasis id="C18F3F7E2443FFC0744467BFF426F953" bold="true" box="[295,314,1749,1768]" pageId="65" pageNumber="282">A,</emphasis>
<taxonomicName id="34FB98EF2443FFC0742267BCF71AF953" authorityName="Schwarz, C. Girard" authorityYear="1903" box="[321,518,1749,1768]" class="Insecta" family="Elateridae" genus="Margogastrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="schneideri">
<emphasis id="C18F3F7E2443FFC0742267BCF71AF953" box="[321,518,1749,1768]" italics="true" pageId="65" pageNumber="282">Margogastrius schneideri</emphasis>
</taxonomicName>
(13.0 mm);
<emphasis id="C18F3F7E2443FFC0771567BCF794F952" bold="true" box="[630,648,1750,1769]" pageId="65" pageNumber="282">B,</emphasis>
<taxonomicName id="34FB98EF2443FFC077F367BFF639F953" authorityName="J. Chassain" baseAuthorityName="Lucas" baseAuthorityYear="1857" box="[656,805,1749,1768]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="C18F3F7E2443FFC077F367BFF639F953" box="[656,805,1749,1768]" italics="true" pageId="65" pageNumber="282">Teslasena femoralis</emphasis>
</taxonomicName>
(9.0 mm);
<emphasis id="C18F3F7E2443FFC076E967BFF683F953" bold="true" box="[906,927,1749,1768]" pageId="65" pageNumber="282">C,</emphasis>
<taxonomicName id="34FB98EF2443FFC076C567BCF6E7F953" authorityName="Rosa" authorityYear="2014" box="[934,1019,1749,1768]" class="Insecta" family="Elateridae" genus="Teslasena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="foucarti">
<emphasis id="C18F3F7E2443FFC076C567BCF6E7F953" box="[934,1019,1749,1768]" italics="true" pageId="65" pageNumber="282">T. foucarti</emphasis>
</taxonomicName>
(9.5 mm);
<emphasis id="C18F3F7E2443FFC0710367BCF16BF952" bold="true" box="[1120,1143,1750,1769]" pageId="65" pageNumber="282">D,</emphasis>
<taxonomicName id="34FB98EF2443FFC0711D67BFF58FF8B9" baseAuthorityName="Abeille de Perrin" baseAuthorityYear="1894" class="Insecta" family="Elateridae" genus="Idiotropia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="henoni">
<emphasis id="C18F3F7E2443FFC0711D67BFF58FF8B9" italics="true" pageId="65" pageNumber="282">Idiotropia henoni</emphasis>
</taxonomicName>
(4.5 mm);
<emphasis id="C18F3F7E2443FFC075976785F41AF8B9" bold="true" box="[244,262,1775,1794]" pageId="65" pageNumber="282">E,</emphasis>
<taxonomicName id="34FB98EF2443FFC074686785F4AEF8B9" box="[267,434,1775,1794]" class="Insecta" family="Elateridae" genus="Oligostethius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="C18F3F7E2443FFC074686785F4AEF8B9" box="[267,434,1775,1794]" italics="true" pageId="65" pageNumber="282">Oligostethius capensis</emphasis>
</taxonomicName>
(10 mm);
<emphasis id="C18F3F7E2443FFC0776E6785F702F8B9" bold="true" box="[525,542,1775,1794]" pageId="65" pageNumber="282">F,</emphasis>
<taxonomicName id="34FB98EF2443FFC077436785F7A6F8B9" box="[544,698,1775,1794]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="C18F3F7E2443FFC077436785F7A6F8B9" box="[544,698,1775,1794]" italics="true" pageId="65" pageNumber="282">Toxognathus bakeri</emphasis>
</taxonomicName>
(6.0 mm);
<emphasis id="C18F3F7E2443FFC0767A6785F632F8B9" bold="true" box="[793,814,1775,1794]" pageId="65" pageNumber="282">G,</emphasis>
<taxonomicName id="34FB98EF2443FFC07650679AF683F8B9" box="[819,927,1775,1794]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="beauchenei">
<emphasis id="C18F3F7E2443FFC07650679AF683F8B9" box="[819,927,1775,1794]" italics="true" pageId="65" pageNumber="282">T. beauchenei</emphasis>
</taxonomicName>
(9.0 mm);
<emphasis id="C18F3F7E2443FFC0769D6785F10AF8B9" bold="true" box="[1022,1046,1775,1794]" pageId="65" pageNumber="282">H,</emphasis>
<taxonomicName id="34FB98EF2443FFC07179679AF16EF8B9" box="[1050,1138,1776,1794]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="C18F3F7E2443FFC07179679AF16EF8B9" box="[1050,1138,1776,1794]" italics="true" pageId="65" pageNumber="282">T. coomani</emphasis>
</taxonomicName>
(6.5 mm);
<emphasis id="C18F3F7E2443FFC0753D6663F570F8A7" bold="true" box="[94,108,1801,1820]" pageId="65" pageNumber="282">I,</emphasis>
<taxonomicName id="34FB98EF2443FFC075116663F5C8F8A7" box="[114,212,1801,1820]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="costulatus">
<emphasis id="C18F3F7E2443FFC075116663F5C8F8A7" box="[114,212,1801,1820]" italics="true" pageId="65" pageNumber="282">T. costulatus</emphasis>
</taxonomicName>
(11.0 mm);
<emphasis id="C18F3F7E2443FFC074206663F44CF8A7" bold="true" box="[323,336,1801,1820]" pageId="65" pageNumber="282">J,</emphasis>
<taxonomicName id="34FB98EF2443FFC074346663F4B4F8A7" box="[343,424,1801,1820]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="doherty">
<emphasis id="C18F3F7E2443FFC074346663F4B4F8A7" box="[343,424,1801,1820]" italics="true" pageId="65" pageNumber="282">T. doherty</emphasis>
</taxonomicName>
(8.0 mm);
<emphasis id="C18F3F7E2443FFC077686663F73CF8A7" bold="true" box="[523,544,1801,1820]" pageId="65" pageNumber="282">K,</emphasis>
<taxonomicName id="34FB98EF2443FFC077456663F791F8A7" box="[550,653,1801,1820]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="C18F3F7E2443FFC077456663F791F8A7" box="[550,653,1801,1820]" italics="true" pageId="65" pageNumber="282">T. fairmairei</emphasis>
</taxonomicName>
(7.5 mm);
<emphasis id="C18F3F7E2443FFC077936663F61EF8A7" bold="true" box="[752,770,1801,1820]" pageId="65" pageNumber="282">L,</emphasis>
<taxonomicName id="34FB98EF2443FFC0766A6663F67EF8A7" box="[777,866,1801,1820]" class="Insecta" family="Elateridae" genus="Toxognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="mouhoti">
<emphasis id="C18F3F7E2443FFC0766A6663F67EF8A7" box="[777,866,1801,1820]" italics="true" pageId="65" pageNumber="282">T. mouhoti</emphasis>
</taxonomicName>
(5.0 mm);
<emphasis id="C18F3F7E2443FFC076A66663F6C1F8A7" bold="true" box="[965,989,1801,1820]" pageId="65" pageNumber="282">M,</emphasis>
<taxonomicName id="34FB98EF2443FFC076876663F19AF8A7" authorityName="Rosa" authorityYear="2014" box="[996,1158,1801,1820]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="hirtus" status="sp. nov.">
<emphasis id="C18F3F7E2443FFC076876663F19AF8A7" box="[996,1158,1801,1820]" italics="true" pageId="65" pageNumber="282">Dactylophysus hirtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052443FFC071EE6663F1D2F8A7" box="[1165,1230,1801,1820]" pageId="65" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
(10.5 mm);
<emphasis id="C18F3F7E2443FFC075A06648F5C5F88E" bold="true" box="[195,217,1826,1845]" pageId="65" pageNumber="282">N,</emphasis>
<taxonomicName id="34FB98EF2443FFC075BD6649F445F88E" authorityName="Rosa" authorityYear="2014" box="[222,345,1826,1845]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="capixabensis" status="nom. nov.">
<emphasis id="C18F3F7E2443FFC075BD6649F445F88E" box="[222,345,1826,1845]" italics="true" pageId="65" pageNumber="282">D. capixabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052443FFC0743D6649F4AFF88D" box="[350,435,1827,1846]" pageId="65" pageNumber="282">nom. nov.</taxonomicNameLabel>
(9.0 mm);
<emphasis id="C18F3F7E2443FFC077716648F734F88E" bold="true" box="[530,552,1826,1845]" pageId="65" pageNumber="282">O,</emphasis>
<taxonomicName id="34FB98EF2443FFC0774E6649F762F88E" baseAuthorityName="Candeze" baseAuthorityYear="1859" box="[557,638,1826,1845]" class="Insecta" family="Elateridae" genus="Dactylophysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="C18F3F7E2443FFC0774E6649F762F88E" box="[557,638,1826,1845]" italics="true" pageId="65" pageNumber="282">D. tibialis</emphasis>
</taxonomicName>
(12.0 mm);
<emphasis id="C18F3F7E2443FFC0778A6648F7E7F88E" bold="true" box="[745,763,1826,1845]" pageId="65" pageNumber="282">P,</emphasis>
<taxonomicName id="34FB98EF2443FFC0779E6648F6A6F88E" box="[765,954,1826,1845]" class="Insecta" family="Elateridae" genus="Heterocrepidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="C18F3F7E2443FFC0779E6648F6A6F88E" box="[765,954,1826,1845]" italics="true" pageId="65" pageNumber="282">Heterocrepidius mendax</emphasis>
</taxonomicName>
(9.0 mm);
<emphasis id="C18F3F7E2443FFC071796648F12DF88E" bold="true" box="[1050,1073,1826,1845]" pageId="65" pageNumber="282">Q,</emphasis>
<taxonomicName id="34FB98EF2443FFC071556648F1D0F88E" authorityName="Rosa" authorityYear="2014" box="[1078,1228,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="asper" status="sp. nov.">
<emphasis id="C18F3F7E2443FFC071556648F1D0F88E" box="[1078,1228,1826,1845]" italics="true" pageId="65" pageNumber="282">Physodactylus asper</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052443FFC0753D6656F582F8F4" box="[94,158,1852,1871]" pageId="65" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
(9.5 mm);
<emphasis id="C18F3F7E2443FFC0759F6656F413F8F4" bold="true" box="[252,271,1852,1871]" pageId="65" pageNumber="282">R,</emphasis>
<taxonomicName id="34FB98EF2443FFC074766656F440F8F4" authorityName="Mannerheim" authorityYear="1842" box="[277,348,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="282" phylum="Arthropoda" rank="species" species="besckei">
<emphasis id="C18F3F7E2443FFC074766656F440F8F4" box="[277,348,1852,1871]" italics="true" pageId="65" pageNumber="282">P. besckei</emphasis>
</taxonomicName>
(11.0 mm).
<emphasis id="C18F3F7E2443FFC074A56656F4F3F8F4" bold="true" box="[454,495,1852,1871]" pageId="65" pageNumber="282">A, D</emphasis>
= lectotypes;
<emphasis id="C18F3F7E2443FFC077036656F611F8F4" bold="true" box="[608,781,1852,1871]" pageId="65" pageNumber="282">E-G, I, J, L, M, P, Q</emphasis>
= holotypes.
</paragraph>
</caption>
<paragraph id="F344E36C2440FFC375E76072F752FE94" blockId="66.[132,590,280,304]" box="[132,590,280,304]" pageId="66" pageNumber="283">
<emphasis id="C18F3F7E2440FFC375E76072F41AFE8B" box="[132,262,280,304]" italics="true" pageId="66" pageNumber="283">Distribution:</emphasis>
<collectingCountry id="8BECA3FC2440FFC3746E6072F470FE8B" box="[269,364,280,304]" name="Brazil" pageId="66" pageNumber="283">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E2440FFC374166073F4E2FE94" box="[373,510,280,304]" italics="true" pageId="66" pageNumber="283">
<collectingRegion id="313F2D8E2440FFC374166073F4E5FE94" box="[373,505,280,304]" country="Brazil" name="Minas Gerais" pageId="66" pageNumber="283">Minas Gerais</collectingRegion>
:
</emphasis>
Lavras.
</paragraph>
<paragraph id="F344E36C2440FFC375E76032F601FE74" blockId="66.[132,680,344,464]" lastBlockId="66.[718,1267,184,463]" pageId="66" pageNumber="283">
<emphasis id="C18F3F7E2440FFC375E76032F5C2FECB" box="[132,222,344,368]" italics="true" pageId="66" pageNumber="283">Remarks:</emphasis>
The specimen from Lavras differs from the other
<specimenCount id="E5FD28E52440FFC375A16012F445FE2B" box="[194,345,376,400]" count="2" pageId="66" pageNumber="283" type="generic">two specimens</specimenCount>
in the coloration pattern (elytra black with lighter epipleura) and relatively longer elytra. Its genitalia is also somewhat different from that of the
<typeStatus id="2C405DCE2440FFC3767261D2F677FF6B" box="[785,875,184,208]" pageId="66" pageNumber="283" type="holotype">holotype</typeStatus>
in the shape of the sternite VIII, with posterior margin slightly emarginate, and the larger ratio between lateroposterior and median parts of the phallobase (4.75). The genitalia of the MNHN and BMNH
<typeStatus id="2C405DCE2440FFC3764A6052F695FEEB" box="[809,905,312,336]" pageId="66" pageNumber="283" type="paratype">paratypes</typeStatus>
were not examined. All specimens share the conical tubercle on posterior region of pronotum and the pilosity pattern of abdomen, which are unique characters for
<taxonomicName id="34FB98EF2440FFC3768960F2F1EEFE0B" authorityName="Rosa" authorityYear="2014" box="[1002,1266,408,432]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC3768960F2F1EEFE0B" box="[1002,1266,408,432]" italics="true" pageId="66" pageNumber="283">Physodactylus tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC377AD60D2F601FE74" box="[718,797,440,463]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<caption id="A784B3E42440FFC375E767BFF77FF8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641457" ID-Zenodo-Dep="12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="66" pageNumber="283" startId="66.[132,211,1749,1768]" targetBox="[134,1266,517,1725]" targetPageId="66" targetType="figure">
<paragraph id="F344E36C2440FFC375E767BFF77FF8F4" blockId="66.[132,1267,1749,1871]" pageId="66" pageNumber="283">
<emphasis id="C18F3F7E2440FFC375E767BFF5EBF953" bold="true" box="[132,247,1749,1768]" pageId="66" pageNumber="283">FIGURE 35:</emphasis>
Habitus.
<emphasis id="C18F3F7E2440FFC3742C67BFF47DF953" bold="true" box="[335,353,1749,1768]" pageId="66" pageNumber="283">A,</emphasis>
<taxonomicName id="34FB98EF2440FFC3740A67BFF733F953" box="[361,559,1749,1768]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="C18F3F7E2440FFC3740A67BFF733F953" box="[361,559,1749,1768]" italics="true" pageId="66" pageNumber="283">Physodactylus brasiliensis</emphasis>
</taxonomicName>
(12.8 mm);
<emphasis id="C18F3F7E2440FFC377C367BCF7AFF952" bold="true" box="[672,691,1750,1769]" pageId="66" pageNumber="283">B,</emphasis>
<taxonomicName id="34FB98EF2440FFC377D867BCF605F953" authorityName="Rosa" authorityYear="2014" box="[699,793,1749,1768]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="brunneus" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC377D867BCF605F953" box="[699,793,1749,1768]" italics="true" pageId="66" pageNumber="283">P. brunneus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC3764267BCF67FF952" box="[801,867,1750,1769]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(9.0 mm);
<emphasis id="C18F3F7E2440FFC376AA67BFF6C1F953" bold="true" box="[969,989,1749,1768]" pageId="66" pageNumber="283">C,</emphasis>
<taxonomicName id="34FB98EF2440FFC3768667BCF15CF953" authorityName="Rosa" authorityYear="2014" box="[997,1088,1749,1768]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="chassaini" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC3768667BCF15CF953" box="[997,1088,1749,1768]" italics="true" pageId="66" pageNumber="283">P. chassaini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC3712B67BCF196F952" box="[1096,1162,1750,1769]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(11.5 mm);
<emphasis id="C18F3F7E2440FFC375E76785F587F8B9" bold="true" box="[132,155,1775,1794]" pageId="66" pageNumber="283">D,</emphasis>
<taxonomicName id="34FB98EF2440FFC375C0679AF5F8F8B9" authorityName="Chassain" authorityYear="2005" box="[163,228,1776,1794]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="costae">
<emphasis id="C18F3F7E2440FFC375C0679AF5F8F8B9" box="[163,228,1776,1794]" italics="true" pageId="66" pageNumber="283">P. costae</emphasis>
</taxonomicName>
(12.0 mm);
<emphasis id="C18F3F7E2440FFC374356785F474F8B9" bold="true" box="[342,360,1775,1794]" pageId="66" pageNumber="283">E,</emphasis>
<taxonomicName id="34FB98EF2440FFC37413679AF4A1F8B9" box="[368,445,1775,1794]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="C18F3F7E2440FFC37413679AF4A1F8B9" box="[368,445,1775,1794]" italics="true" pageId="66" pageNumber="283">P. fischeri</emphasis>
</taxonomicName>
(17.0 mm);
<emphasis id="C18F3F7E2440FFC3774C6785F75CF8B9" bold="true" box="[559,576,1775,1794]" pageId="66" pageNumber="283">F,</emphasis>
<taxonomicName id="34FB98EF2440FFC37726679AF7B8F8B9" authorityName="Rosa" authorityYear="2014" box="[581,676,1775,1794]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="flavifrons" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC37726679AF7B8F8B9" box="[581,676,1775,1794]" italics="true" pageId="66" pageNumber="283">P. flavifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC377CF6785F7F2F8B9" box="[684,750,1775,1794]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(10.0 mm);
<emphasis id="C18F3F7E2440FFC3763C6785F669F8B9" bold="true" box="[863,885,1775,1794]" pageId="66" pageNumber="283">G,</emphasis>
<taxonomicName id="34FB98EF2440FFC3761F679AF6C1F8B9" authorityName="Rosa" authorityYear="2014" box="[892,989,1775,1794]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="fleutiauxi">
<emphasis id="C18F3F7E2440FFC3761F679AF6C1F8B9" box="[892,989,1775,1794]" italics="true" pageId="66" pageNumber="283">P. fleutiauxi</emphasis>
</taxonomicName>
(15.0 mm);
<emphasis id="C18F3F7E2440FFC3712C6785F17AF8B9" bold="true" box="[1103,1126,1775,1794]" pageId="66" pageNumber="283">H,</emphasis>
<taxonomicName id="34FB98EF2440FFC3710D679AF1EEF8B9" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[1134,1266,1775,1794]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="foveatostriatus">
<emphasis id="C18F3F7E2440FFC3710D679AF1EEF8B9" box="[1134,1266,1775,1794]" italics="true" pageId="66" pageNumber="283">P. foveatostriatus</emphasis>
</taxonomicName>
(12.0 mm);
<emphasis id="C18F3F7E2440FFC3758B6663F5EAF8A7" bold="true" box="[232,246,1801,1820]" pageId="66" pageNumber="283">I,</emphasis>
<taxonomicName id="34FB98EF2440FFC375986663F45EF8A7" authorityName="Rosa" authorityYear="2014" box="[251,322,1801,1820]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="girardi" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC375986663F45EF8A7" box="[251,322,1801,1820]" italics="true" pageId="66" pageNumber="283">P. girardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC374246663F49BF8A7" box="[327,391,1801,1820]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(9.0 mm);
<emphasis id="C18F3F7E2440FFC374876663F4EEF8A7" bold="true" box="[484,498,1801,1820]" pageId="66" pageNumber="283">J,</emphasis>
<taxonomicName id="34FB98EF2440FFC374956663F752F8A7" authorityName="Rosa" authorityYear="2014" box="[502,590,1801,1820]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="gounellei" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC374956663F752F8A7" box="[502,590,1801,1820]" italics="true" pageId="66" pageNumber="283">P. gounellei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC377306663F78FF8A7" box="[595,659,1801,1820]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(10.5 mm);
<emphasis id="C18F3F7E2440FFC377996663F613F8A7" bold="true" box="[762,783,1801,1820]" pageId="66" pageNumber="283">K,</emphasis>
<taxonomicName id="34FB98EF2440FFC376776663F670F8A7" authorityName="Fischer von Waldheim" authorityYear="1823" box="[788,876,1801,1820]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="henningi">
<emphasis id="C18F3F7E2440FFC376776663F670F8A7" box="[788,876,1801,1820]" italics="true" pageId="66" pageNumber="283">P. henningi</emphasis>
</taxonomicName>
(15.0 mm);
<emphasis id="C18F3F7E2440FFC376B66663F6FBF8A7" bold="true" box="[981,999,1801,1820]" pageId="66" pageNumber="283">L,</emphasis>
<taxonomicName id="34FB98EF2440FFC3768F6663F157F8A7" authorityName="Rosa" authorityYear="2014" box="[1004,1099,1801,1820]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="latithorax" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC3768F6663F157F8A7" box="[1004,1099,1801,1820]" italics="true" pageId="66" pageNumber="283">P. latithorax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC371336663F18CF8A7" box="[1104,1168,1801,1820]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(18.0 mm);
<emphasis id="C18F3F7E2440FFC375E76648F581F88E" bold="true" box="[132,157,1826,1845]" pageId="66" pageNumber="283">M,</emphasis>
<taxonomicName id="34FB98EF2440FFC375C16649F5C7F88E" authorityName="Fleutiaux" authorityYear="1892" box="[162,219,1827,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2440FFC375C16649F5C7F88E" box="[162,219,1827,1845]" italics="true" pageId="66" pageNumber="283">P. niger</emphasis>
</taxonomicName>
(11.0 mm);
<emphasis id="C18F3F7E2440FFC374276648F446F88E" bold="true" box="[324,346,1826,1845]" pageId="66" pageNumber="283">N,</emphasis>
<taxonomicName id="34FB98EF2440FFC3743C6649F4A6F88E" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[351,442,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2440FFC3743C6649F4A6F88E" box="[351,442,1826,1845]" italics="true" pageId="66" pageNumber="283">P. oberthuri</emphasis>
</taxonomicName>
(12.5 mm);
<emphasis id="C18F3F7E2440FFC377406648F726F88E" bold="true" box="[547,570,1826,1845]" pageId="66" pageNumber="283">O,</emphasis>
<taxonomicName id="34FB98EF2440FFC3775D6649F79DF88E" authorityName="Rosa" authorityYear="2014" box="[574,641,1827,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="patens" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC3775D6649F79DF88E" box="[574,641,1827,1845]" italics="true" pageId="66" pageNumber="283">P. patens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC377E56649F7D9F88D" box="[646,709,1827,1846]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(17.0 mm);
<emphasis id="C18F3F7E2440FFC3764E6648F623F88E" bold="true" box="[813,831,1826,1845]" pageId="66" pageNumber="283">P,</emphasis>
<taxonomicName id="34FB98EF2440FFC376236649F69CF88E" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" box="[832,896,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="pujoli">
<emphasis id="C18F3F7E2440FFC376236649F69CF88E" box="[832,896,1826,1845]" italics="true" pageId="66" pageNumber="283">P. pujoli</emphasis>
</taxonomicName>
(12.0 mm);
<emphasis id="C18F3F7E2440FFC3768B6648F6E3F88E" bold="true" box="[1000,1023,1826,1845]" pageId="66" pageNumber="283">Q,</emphasis>
<taxonomicName id="34FB98EF2440FFC371606649F14EF88E" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[1027,1106,1826,1845]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="C18F3F7E2440FFC371606649F14EF88E" box="[1027,1106,1826,1845]" italics="true" pageId="66" pageNumber="283">P. sulcatus</emphasis>
</taxonomicName>
(9.0 mm);
<emphasis id="C18F3F7E2440FFC371D36648F1D8F88E" bold="true" box="[1200,1220,1826,1845]" pageId="66" pageNumber="283">R,</emphasis>
<taxonomicName id="34FB98EF2440FFC371AB6649F5CFF8F4" authorityName="Rosa" authorityYear="2014" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="283" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="C18F3F7E2440FFC371AB6649F5CFF8F4" italics="true" pageId="66" pageNumber="283">P. tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052440FFC375BA6656F404F8F4" box="[217,280,1852,1871]" pageId="66" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
(12.0 mm).
<emphasis id="C18F3F7E2440FFC374E26656F4E4F8F4" bold="true" box="[385,504,1852,1871]" pageId="66" pageNumber="283">A-D, F-J, L-R</emphasis>
= holotypes.
</paragraph>
</caption>
<caption id="A784B3E42441FFC2753C6662F695F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641461" ID-Zenodo-Dep="12641461" httpUri="https://zenodo.org/record/12641461/files/figure.png" pageId="67" pageNumber="284" startId="67.[95,174,1800,1819]" targetBox="[213,1108,171,1777]" targetPageId="67" targetType="figure">
<paragraph id="F344E36C2441FFC2753C6662F695F8F4" blockId="67.[94,1229,1800,1871]" pageId="67" pageNumber="284">
<emphasis id="C18F3F7E2441FFC2753C6662F5CDF8A0" bold="true" box="[95,209,1800,1819]" pageId="67" pageNumber="284">FIGURE 36:</emphasis>
Strict consensus of four most parsimonious trees from the phylogenetic analysis of
<taxonomicName id="34FB98EF2441FFC276FB6663F100F8A7" box="[920,1052,1801,1820]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="284" phylum="Arthropoda" rank="subFamily" subFamily="Physodactylinae">Physodactylinae</taxonomicName>
(length = 281 steps, CI = 0.35, RI = 0.68). Character and state numbers are respectively above and below internodes. Black circles indicate exclusive synapomorphies and white circles homoplastic synapomorphies. Absolute Bremer support in parentheses.
</paragraph>
</caption>
<paragraph id="F344E36C2446FFC575DE61D2F4DFFD54" blockId="68.[132,681,184,752]" pageId="68" pageNumber="285">
<taxonomicName id="34FB98EF2446FFC575DE61D2F4DAFF6B" authorityName="Rosa" authorityYear="2014" box="[189,454,184,208]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="285" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="C18F3F7E2446FFC575DE61D2F4DAFF6B" box="[189,454,184,208]" italics="true" pageId="68" pageNumber="285">Physodactylus tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052446FFC574B261D2F73AFF74" box="[465,550,184,207]" pageId="68" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
shares with
<taxonomicName id="34FB98EF2446FFC575E761B3F5D1FF4B" authorityName="Fleutiaux" authorityYear="1892" box="[132,205,217,240]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="285" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2446FFC575E761B3F5D1FF4B" box="[132,205,217,240]" italics="true" pageId="68" pageNumber="285">P. niger</emphasis>
</taxonomicName>
and
<taxonomicName id="34FB98EF2446FFC5746361B3F468FF4B" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[256,372,216,240]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="285" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2446FFC5746361B3F468FF4B" box="[256,372,216,240]" italics="true" pageId="68" pageNumber="285">P. oberthuri</emphasis>
</taxonomicName>
the shape of prothorax, but is easily distinguishable by the above mentioned unique characters. It shares with
<taxonomicName id="34FB98EF2446FFC574F06073F4C2FE8B" authorityName="Fleutiaux" authorityYear="1892" box="[403,478,281,304]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="285" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2446FFC574F06073F4C2FE8B" box="[403,478,281,304]" italics="true" pageId="68" pageNumber="285">P. niger</emphasis>
</taxonomicName>
the yellow pilosity and differs from this species also in its smaller and homogenously distributed pronotal punctation, anteromedian margin of pronotum more produced, elytral striae with larger punctures, flat elytral interstices, antennomeres shorter and the lateral carina more prominent posteriorly and less prominent anteriorly. It is most similar to
<taxonomicName id="34FB98EF2446FFC574326093F4D8FDAB" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[337,452,504,528]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="285" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2446FFC574326093F4D8FDAB" box="[337,452,504,528]" italics="true" pageId="68" pageNumber="285">P. oberthuri</emphasis>
</taxonomicName>
in the aedeagal shape, very bright integument, pronotal punctation homogenously distributed and the color pattern, usually with elytra lighter, a pattern not found among
<taxonomicName id="34FB98EF2446FFC577036333F7B4FDCB" authorityName="Fleutiaux" authorityYear="1892" box="[608,680,601,624]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="285" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="C18F3F7E2446FFC577036333F7B4FDCB" box="[608,680,601,624]" italics="true" pageId="68" pageNumber="285">P. niger</emphasis>
</taxonomicName>
specimens.
<taxonomicName id="34FB98EF2446FFC5759F6312F71EFD2B" authorityName="Rosa" authorityYear="2014" box="[252,514,632,656]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="285" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="C18F3F7E2446FFC5759F6312F71EFD2B" box="[252,514,632,656]" italics="true" pageId="68" pageNumber="285">Physodactylus tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC82052446FFC577686312F740FD34" box="[523,604,632,655]" pageId="68" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="34FB98EF2446FFC5743363F3F4D9FD0B" authorityName="Fleutiaux, J. Chassain" authorityYear="1892" box="[336,453,664,688]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="285" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="C18F3F7E2446FFC5743363F3F4D9FD0B" box="[336,453,664,688]" italics="true" pageId="68" pageNumber="285">P. oberthuri</emphasis>
</taxonomicName>
by its yellow pilosity, larger pronotal punctation, the pronotal tubercle and the abdominal pilosity pattern.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,243 @@
<document id="9675B27CAC76F35D8B575C0E19D2B648" ID-DOI="10.1590/0031-1049.2014.54.18" ID-ISSN="1807-0205" ID-Zenodo-Dep="12641367" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720058375023" checkinUser="felipe" docAuthor="Rosa, Simone Policena" docDate="2014" docId="7B52527A243EFFBC767A603DF4CEF8EB" docLanguage="en" docName="PapAvZool.54.18.217-292.pdf" docOrigin="Papéis Avulsos de Zoologia (São Paulo) 54 (18)" docSource="http://dx.doi.org/10.1590/0031-1049.2014.54.18" docStyle="DocumentStyle:8E93E160036069519CAC16E940FF2271.1:PapAvZool.2014-2017.journal_article" docStyleId="8E93E160036069519CAC16E940FF2271" docStyleName="PapAvZool.2014-2017.journal_article" docStyleVersion="1" docTitle="Physodactylus pujoli Fleut., FLEUTIAUX 1940" docType="treatment" docVersion="2" lastPageNumber="278" masterDocId="876B2A022402FF817563616AF51CFFBB" masterDocTitle="Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)" masterLastPageNumber="292" masterPageNumber="217" pageNumber="277" updateTime="1720066190901" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2251338744D25ED02D482D1F69ADF366">Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae)</mods:title>
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<mods:titleInfo id="23EF55D7903652D7D05FA6B8F4E3BEAB">
<mods:title id="544147C5267557E83E08A0FCF067DC3C">Papéis Avulsos de Zoologia</mods:title>
</mods:titleInfo>
<mods:part id="91AA63F8054C1E37F5D5A60A16DFD26A">
<mods:date id="1182656EE6C8FFDC68693685041C17E6">2014</mods:date>
<mods:detail id="AA39ECA36BAB4E5D5D696967D5AC3E5B" type="series">
<mods:title id="E9760F2A8247B5DF3F56ABE3F42099B3">São Paulo</mods:title>
</mods:detail>
<mods:detail id="1864B35B64DFD6F4FE13BECC477FDF86" type="volume">
<mods:number id="EDB50E5CCFE705E57DB11392EA57722E">54</mods:number>
</mods:detail>
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<mods:number id="1A0C02BE0D6A4FFB1CECAD312FD78109">18</mods:number>
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<mods:extent id="5ABB917CCF41AF00785CBBA25C269390" unit="page">
<mods:start id="3CA2F9D3A86C38D95DA2D30AAE515999">217</mods:start>
<mods:end id="88840C255739C7D567EC5A7EA3D4FD8D">292</mods:end>
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<mods:location id="7845FFA7A838A5ED6893A237DA7F7601">
<mods:url id="37FDAC22A17FF52876AF7DB88F55B69E">http://dx.doi.org/10.1590/0031-1049.2014.54.18</mods:url>
</mods:location>
<mods:classification id="31DCA5E5715B09D069092A535090CE0E">journal article</mods:classification>
<mods:identifier id="24846D829AFF129BB44CF33E3493A5A2" type="DOI">10.1590/0031-1049.2014.54.18</mods:identifier>
<mods:identifier id="79F8BE45DC04063C419631C2AB9E26D2" type="ISSN">1807-0205</mods:identifier>
<mods:identifier id="5348C7620DCC910173CBDD2E0E9141D8" type="Zenodo-Dep">12641367</mods:identifier>
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<subSubSection id="BBE1B0E7243EFFBD767A603DF1BBFECB" box="[793,1191,343,368]" pageId="60" pageNumber="277" type="nomenclature">
<paragraph id="F344E36C243EFFBD767A603DF1BBFECB" blockId="60.[793,1191,343,400]" box="[793,1191,343,368]" pageId="60" pageNumber="277">
<heading id="A80C5400243EFFBD767A603DF1BBFECB" bold="true" box="[793,1191,343,368]" centered="true" fontSize="10" level="2" pageId="60" pageNumber="277" reason="2">
<taxonomicName id="34FB98EF243EFFBD767A603DF1BBFECB" authority="Fleutiaux, 1940" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" box="[793,1191,343,368]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="277" phylum="Arthropoda" rank="species" species="pujoli">
<emphasis id="C18F3F7E243EFFBD767A603DF1BBFECB" bold="true" box="[793,1191,343,368]" pageId="60" pageNumber="277">
<emphasis id="C18F3F7E243EFFBD767A603DF6EAFED4" bold="true" box="[793,1014,343,367]" italics="true" pageId="60" pageNumber="277">Physodactylus pujoli</emphasis>
Fleutiaux, 1940
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7243EFFBD76F36012F12DFE2B" box="[912,1073,376,400]" pageId="60" pageNumber="277" type="description">
<paragraph id="F344E36C243EFFBD76F36012F12DFE2B" blockId="60.[793,1191,343,400]" box="[912,1073,376,400]" pageId="60" pageNumber="277">
<emphasis id="C18F3F7E243EFFBD76F36012F12DFE2B" bold="true" box="[912,1073,376,400]" pageId="60" pageNumber="277">
(
<figureCitation id="6BC0FFE9243EFFBD76F46012F6F3FE2B" box="[919,1007,376,400]" captionStart="FIGURE 29" captionStartId="60.[132,211,1852,1871]" captionTargetBox="[135,1264,1226,1822]" captionTargetId="figure-485@60.[132,1266,1223,1828]" captionTargetPageId="60" captionText="FIGURE 29: Physodactylus pujoli. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641439" httpUri="https://zenodo.org/record/12641439/files/figure.png" pageId="60" pageNumber="277">Figs. 29</figureCitation>
,
<figureCitation id="6BC0FFE9243EFFBD76986012F134FE2B" box="[1019,1064,376,400]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="60" pageNumber="277">35P</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7243EFFBD77AD60D2F6D4FE4B" pageId="60" pageNumber="277" type="reference_group">
<paragraph id="F344E36C243EFFBD77AD60D2F6D4FE4B" blockId="60.[718,1266,440,496]" pageId="60" pageNumber="277">
<treatmentCitationGroup id="D3EBC442243EFFBD77AD60D2F6D4FE4B" pageId="60" pageNumber="277">
<treatmentCitation id="725AC57D243EFFBD77AD60D2F1B0FE6B" author="FLEUTIAUX, E." box="[718,1196,440,464]" page="164" pageId="60" pageNumber="277" year="1940">
<taxonomicName id="34FB98EF243EFFBD77AD60D2F1B0FE6B" authority="Fleutiaux, 1940 c: 164" authorityName="Fleut., FLEUTIAUX" authorityPageNumber="164" authorityYear="1940" box="[718,1196,440,464]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="277" phylum="Arthropoda" rank="species" species="pujoli">
<emphasis id="C18F3F7E243EFFBD77AD60D2F682FE6B" box="[718,926,440,464]" italics="true" pageId="60" pageNumber="277">Physodactylus pujoli</emphasis>
<bibRefCitation id="976A9E9D243EFFBD76D360D2F1B0FE6B" author="FLEUTIAUX, E." box="[944,1196,440,464]" pageId="60" pageNumber="277" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 164</bibRefCitation>
</taxonomicName>
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, 166;
<treatmentCitation id="725AC57D243EFFBD77AD60B2F6D8FE4B" author="BLACKWELDER, R. E." box="[718,964,472,496]" page="292" pageId="60" pageNumber="277" year="1944">
<bibRefCitation id="976A9E9D243EFFBD77AD60B2F6D8FE4B" author="BLACKWELDER, R. E." box="[718,964,472,496]" pageId="60" pageNumber="277" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7243EFFBC77AD6372F4CEF8EB" lastPageId="61" lastPageNumber="278" pageId="60" pageNumber="277" type="description">
<paragraph id="F344E36C243EFFBD77AD6372F1EFFB0B" blockId="60.[718,1267,536,1200]" pageId="60" pageNumber="277">
<emphasis id="C18F3F7E243EFFBD77AD6372F64CFD8B" box="[718,848,536,560]" italics="true" pageId="60" pageNumber="277">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9243EFFBD76D56372F101FD8B" box="[950,1053,536,560]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="60" pageNumber="277">Fig. 35P</figureCitation>
): Integument very shiny; head, antenna, ventral pterothorax and legs dark brown, pronotum light brown with borders (or borders and disc) darker, elytra evenly brown or gradually darker apicad; ventral abdomen black with lateral borders brown; pilosity brown. Total length 9.5-12.0 mm; elytral base as wide as prothorax, elytra 2.8-3.0x longer than pronotum. Frons concave, frontal carina produced; antenna (
<figureCitation id="6BC0FFE9243EFFBD71646272F178FC8B" box="[1031,1124,792,816]" captionStart="FIGURE 29" captionStartId="60.[132,211,1852,1871]" captionTargetBox="[135,1264,1226,1822]" captionTargetId="figure-485@60.[132,1266,1223,1828]" captionTargetPageId="60" captionText="FIGURE 29: Physodactylus pujoli. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641439" httpUri="https://zenodo.org/record/12641439/files/figure.png" pageId="60" pageNumber="277">Fig. 29A</figureCitation>
) reaching the base of posterior angles; antennomere IV as wide as long, V-VI 1.1-1.2x wider than long, VII-VIII as wide as long, IX-X 1.1-1.2x longer than wide, XI elliptical 2.0x longer than wide, apex constricted. Pronotum (
<figureCitation id="6BC0FFE9243EFFBD766462D2F67EFC6B" box="[775,866,952,976]" captionStart="FIGURE 29" captionStartId="60.[132,211,1852,1871]" captionTargetBox="[135,1264,1226,1822]" captionTargetId="figure-485@60.[132,1266,1223,1828]" captionTargetPageId="60" captionText="FIGURE 29: Physodactylus pujoli. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641439" httpUri="https://zenodo.org/record/12641439/files/figure.png" pageId="60" pageNumber="277">Fig. 29B</figureCitation>
) weakly convex 1.22-1.27x wider than long, with lateral sides nearly straight from posterior angles or posterior third to anterior margin; anteromedian margin weakly produced anteriad; posterior angles short, convex; lateral carina complete weakly impressed anteriorly, dorsally visible only on posterior half; disc with punctures 2-4 diameters apart, a little larger and denser on lateral and anterior borders.
</paragraph>
<caption id="A784B3E4243EFFBD75E76656F694F8F4" ID-DOI="http://doi.org/10.5281/zenodo.12641439" ID-Zenodo-Dep="12641439" box="[132,904,1852,1871]" httpUri="https://zenodo.org/record/12641439/files/figure.png" pageId="60" pageNumber="277" startId="60.[132,211,1852,1871]" targetBox="[135,1264,1226,1822]" targetPageId="60" targetType="figure">
<paragraph id="F344E36C243EFFBD75E76656F694F8F4" blockId="60.[132,904,1852,1871]" box="[132,904,1852,1871]" pageId="60" pageNumber="277">
<emphasis id="C18F3F7E243EFFBD75E76656F5E9F8F4" bold="true" box="[132,245,1852,1871]" pageId="60" pageNumber="277">FIGURE 29:</emphasis>
<emphasis id="C18F3F7E243EFFBD75996656F482F8F4" box="[250,414,1852,1871]" italics="true" pageId="60" pageNumber="277">
<taxonomicName id="34FB98EF243EFFBD75996656F485F8F4" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" box="[250,409,1852,1871]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="277" phylum="Arthropoda" rank="species" species="pujoli">Physodactylus pujoli</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E243EFFBD74C16656F4A8F8F4" bold="true" box="[418,436,1852,1871]" pageId="60" pageNumber="277">A,</emphasis>
antenna;
<emphasis id="C18F3F7E243EFFBD77656656F705F8F4" bold="true" box="[518,537,1852,1871]" pageId="60" pageNumber="277">B,</emphasis>
pronotum;
<emphasis id="C18F3F7E243EFFBD771E6656F7B1F8F4" bold="true" box="[637,685,1852,1871]" pageId="60" pageNumber="277">C, D,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
</caption>
<caption id="A784B3E4243FFFBC753C63C7F1D1FD7B" ID-DOI="http://doi.org/10.5281/zenodo.12641441" ID-Zenodo-Dep="12641441" box="[95,1229,685,705]" httpUri="https://zenodo.org/record/12641441/files/figure.png" pageId="61" pageNumber="278" startId="61.[95,174,685,704]" targetBox="[96,1225,171,657]" targetPageId="61" targetType="figure">
<paragraph id="F344E36C243FFFBC753C63C7F1D1FD7B" blockId="61.[94,1229,685,1264]" box="[95,1229,685,705]" pageId="61" pageNumber="278">
<emphasis id="C18F3F7E243FFFBC753C63C7F5CDFD7B" bold="true" box="[95,209,685,704]" pageId="61" pageNumber="278">FIGURE 30:</emphasis>
<emphasis id="C18F3F7E243FFFBC75BA63C7F491FD7B" box="[217,397,685,704]" italics="true" pageId="61" pageNumber="278">
<taxonomicName id="34FB98EF243FFFBC75BA63C7F495FD7B" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[217,393,685,704]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="sulcatus">Physodactylus sulcatus</taxonomicName>
.
</emphasis>
<emphasis id="C18F3F7E243FFFBC74F063C7F4BAFD7B" bold="true" box="[403,422,685,704]" pageId="61" pageNumber="278">A,</emphasis>
antenna;
<emphasis id="C18F3F7E243FFFBC749F63C7F713FD7B" bold="true" box="[508,527,685,704]" pageId="61" pageNumber="278">B,</emphasis>
maxilla;
<emphasis id="C18F3F7E243FFFBC773C63C7F76FFD7B" bold="true" box="[607,627,685,704]" pageId="61" pageNumber="278">C,</emphasis>
labrum and mandible;
<emphasis id="C18F3F7E243FFFBC765D63C7F671FD7B" bold="true" box="[830,877,685,704]" pageId="61" pageNumber="278">D, E,</emphasis>
prothorax (dorsal, lateral);
<emphasis id="C18F3F7E243FFFBC713B63C7F19FFD7B" bold="true" box="[1112,1155,685,704]" pageId="61" pageNumber="278">F, G,</emphasis>
pro and
</paragraph>
</caption>
<paragraph id="F344E36C243FFFBC753C63ADF4A7FD61" blockId="61.[94,1229,685,1264]" box="[95,443,711,730]" pageId="61" pageNumber="278">
metatibia;
<emphasis id="C18F3F7E243FFFBC75D563ADF5FCFD61" bold="true" box="[182,224,711,730]" pageId="61" pageNumber="278">H, I,</emphasis>
aedeagus (dorsal, ventral).
</paragraph>
<paragraph id="F344E36C243FFFBC753D6392F176FB4B" blockId="61.[94,1229,685,1264]" pageId="61" pageNumber="278">
Lamellae of pro- and mesotarsomeres I smaller than
<emphasis id="C18F3F7E243FFFBC77CB6392F676FCAB" box="[680,874,760,784]" italics="true" pageId="61" pageNumber="278">Material examined:</emphasis>
<collectingCountry id="8BECA3FC243FFFBC76136392F6D3FCAB" box="[880,975,760,784]" name="Brazil" pageId="61" pageNumber="278">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E243FFFBC76B46392F104FCAB" box="[983,1048,760,784]" italics="true" pageId="61" pageNumber="278">
<collectingRegion id="313F2D8E243FFFBC76B46392F10FFCAB" box="[983,1043,760,784]" country="Brazil" name="Bahia" pageId="61" pageNumber="278">Bahia</collectingRegion>
:
</emphasis>
<specimenCount id="E5FD28E5243FFFBC717C6392F144FCB4" box="[1055,1112,760,784]" count="2" pageId="61" pageNumber="278" type="generic">2 exs.</specimenCount>
(MNHN). the others, minuscule or absent on metatarsomere I. Elytra parallel-sided on anterior 2/3 then tapering to
<emphasis id="C18F3F7E243FFFBC77CB6252F636FCEB" box="[680,810,824,848]" italics="true" pageId="61" pageNumber="278">Distribution:</emphasis>
<collectingCountry id="8BECA3FC243FFFBC76526252F68CFCEB" box="[817,912,824,848]" name="Brazil" pageId="61" pageNumber="278">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E243FFFBC76FA6252F6C6FCEB" box="[921,986,824,848]" italics="true" pageId="61" pageNumber="278">
<collectingRegion id="313F2D8E243FFFBC76FA6252F6C9FCEB" box="[921,981,824,848]" country="Brazil" name="Bahia" pageId="61" pageNumber="278">Bahia</collectingRegion>
:
</emphasis>
Ribeirão do Largo. apex, striae with rows of large and small punctation; interstices convex, scarcely punctate, rugose; apical
<emphasis id="C18F3F7E243FFFBC77CB6212F61EFC2B" box="[680,770,888,912]" italics="true" pageId="61" pageNumber="278">Remarks:</emphasis>
The prosternal process was not examined. striae with punctures about 2.0 larger than those of
<taxonomicName id="34FB98EF243FFFBC77CB62F2F66DFC0B" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" box="[680,881,920,944]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="pujoli">
<emphasis id="C18F3F7E243FFFBC77CB62F2F66DFC0B" box="[680,881,920,944]" italics="true" pageId="61" pageNumber="278">Physodactylus pujoli</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="34FB98EF243FFFBC716562F3F196FC0B" box="[1030,1162,920,944]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="C18F3F7E243FFFBC716562F3F196FC0B" box="[1030,1162,920,944]" italics="true" pageId="61" pageNumber="278">P. brasiliensis</emphasis>
</taxonomicName>
in its pronotal lateral border. Abdomen with ventrites 2-4 very shiny integument, color pattern and weakly constrongly concave laterally, ventrite 1 with setae a little vex pronotum, but can be promptly distinguished smaller than those of ventrite 2, black area of ven- from this species by its brown pilosity. It shares with trites 1-4 covered with long decumbent and semi-
<taxonomicName id="34FB98EF243FFFBC77CB6573F7ECFB8B" authorityName="Rosa" authorityYear="2014" box="[680,752,1049,1072]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="C18F3F7E243FFFBC77CB6573F7ECFB8B" box="[680,752,1049,1072]" italics="true" pageId="61" pageNumber="278">P. asper</emphasis>
</taxonomicName>
and
<taxonomicName id="34FB98EF243FFFBC76406573F662FB8B" box="[803,894,1048,1072]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="C18F3F7E243FFFBC76406573F662FB8B" box="[803,894,1048,1072]" italics="true" pageId="61" pageNumber="278">P. fischeri</emphasis>
</taxonomicName>
the elytral striae punctation, but erect setae and dense punctation, lateral brown area its striae are more deeply impressed and distinct at with punctures finer and setae sparser. Pregenitalic apex.
<emphasis id="C18F3F7E243FFFBC77846532F6B2FBCB" box="[743,942,1112,1136]" italics="true" pageId="61" pageNumber="278">
Physodactulys
<taxonomicName id="34FB98EF243FFFBC76166532F6B2FBCB" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" box="[885,942,1112,1136]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="pujoli">pujoli</taxonomicName>
</emphasis>
shares the antennal length segments and aedeagus covered with stout dark- with
<taxonomicName id="34FB98EF243FFFBC77BD6513F63BFB2B" authorityName="Rosa" authorityYear="2014" box="[734,807,1145,1168]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="asper" status="sp. nov.">
<emphasis id="C18F3F7E243FFFBC77BD6513F63BFB2B" box="[734,807,1145,1168]" italics="true" pageId="61" pageNumber="278">P. asper</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243FFFBC764C6512F663FB34" box="[815,895,1144,1167]" pageId="61" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="34FB98EF243FFFBC76D06513F111FB2B" authorityName="Rosa" authorityYear="2014" box="[947,1037,1144,1168]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="girardi" status="sp. nov.">
<emphasis id="C18F3F7E243FFFBC76D06513F111FB2B" box="[947,1037,1144,1168]" italics="true" pageId="61" pageNumber="278">P. girardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243FFFBC71766512F179FB34" box="[1045,1125,1144,1167]" pageId="61" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
It is symbrown setae. Sternite VIII subrectangular, posterior patric with the former, from which it differs in the margin nearly straight, anterior sclerotization 0.44x color pattern, pilosity of the ventrite 1, pronotal the total width of sternite; sternite IX tapered to apex. punctation and elytral striae and interstices.
</paragraph>
<paragraph id="F344E36C243FFFBC753D6472F4ACF9AB" blockId="61.[94,643,1304,1552]" pageId="61" pageNumber="278">
<emphasis id="C18F3F7E243FFFBC753D6472F5ABFA8B" box="[94,183,1304,1328]" italics="true" pageId="61" pageNumber="278">Aedeagus</emphasis>
(
<figureCitation id="6BC0FFE9243FFFBC75AA6472F444FA8B" box="[201,344,1304,1328]" captionStart="FIGURE 29" captionStartId="60.[132,211,1852,1871]" captionTargetBox="[135,1264,1226,1822]" captionTargetId="figure-485@60.[132,1266,1223,1828]" captionTargetPageId="60" captionText="FIGURE 29: Physodactylus pujoli. A, antenna; B, pronotum; C, D, aedeagus (dorsal, ventral)." figureDoi="http://doi.org/10.5281/zenodo.12641439" httpUri="https://zenodo.org/record/12641439/files/figure.png" pageId="61" pageNumber="278">Figs. 29C, D</figureCitation>
): Phallobase 0.44x the total length of aedeagus, 1.12x longer than wide, length ratio between lateroposterior and median parts 13.7; paramere with ventral surface weakly sclerotized laterally and along the anteromedian margin; penis with basal strut 0.26x its total length, articulated to the parameres by a translucent membrane; ventral sclerite linear weakly sclerotized apically.
</paragraph>
<paragraph id="F344E36C243FFFBC753D6752F441F90B" blockId="61.[94,642,1592,1712]" pageId="61" pageNumber="278">
<materialsCitation id="4393E931243FFFBC753D6752F441F90B" collectionCode="TYPE, MNHN" collectorName="Ch. Pujol &amp; Coll. E. Fleutiaux" location="Cachimbo" pageId="61" pageNumber="278" specimenCount="2" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="C18F3F7E243FFFBC753D6752F5A7F9EB" box="[94,187,1592,1616]" italics="true" pageId="61" pageNumber="278">
<typeStatus id="2C405DCE243FFFBC753D6752F5AAF9EB" box="[94,182,1592,1616]" pageId="61" pageNumber="278" type="holotype">Holotype</typeStatus>
:
</emphasis>
[
<location id="F624B5B7243FFFBC75A86752F420F9EB" LSID="urn:lsid:plazi:treatment:7B52527A243EFFBC767A603DF4CEF8EB:F624B5B7243FFFBC75A86752F420F9EB" box="[203,316,1592,1616]" name="Cachimbo" pageId="61" pageNumber="278" stateProvince="Bahia">Cachimbo</location>
, Prov.
<collectingRegion id="313F2D8E243FFFBC74E66752F4DEF9EB" box="[389,450,1592,1616]" country="Brazil" name="Bahia" pageId="61" pageNumber="278">Bahia</collectingRegion>
,
<collectorName id="5E0E86BA243FFFBC74AC6752F729F9EB" box="[463,565,1592,1616]" pageId="61" pageNumber="278">Ch. Pujol</collectorName>
1890], [
<collectionCode id="95EA7BA9243FFFBC75066732F5B0F9D4" box="[101,172,1624,1647]" pageId="61" pageNumber="278">TYPE</collectionCode>
], [
<taxonomicName id="34FB98EF243FFFBC75A76732F79EF9CB" authority="Fleut., FLEUTIAUX" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" box="[196,642,1624,1648]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="pujoli">Physodactylus pujoli Fleut., FLEUTIAUX</taxonomicName>
det. type], [Muséum Paris,
<collectorName id="5E0E86BA243FFFBC741F6712F727F92B" box="[380,571,1656,1680]" pageId="61" pageNumber="278">Coll. E. Fleutiaux</collectorName>
], [
<typeStatus id="2C405DCE243FFFBC77306712F5B8F90B" pageId="61" pageNumber="278" type="lectotype">Lectotype</typeStatus>
], male (
<collectionCode id="95EA7BA9243FFFBC759667F2F44DF90B" box="[245,337,1688,1712]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="61" pageNumber="278">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="F344E36C243FFFBC753D67B3F4CEF8EB" blockId="61.[94,642,1752,1872]" pageId="61" pageNumber="278">
<materialsCitation id="4393E931243FFFBC753D67B3F4D2F8EB" collectionCode="MNHN" collectorName="Ch. Pujol &amp; Coll. E. Fleutiaux" location="Bahia" pageId="61" pageNumber="278" specimenCount="1" stateProvince="Bahia" typeStatus="paratype">
<emphasis id="C18F3F7E243FFFBC753D67B3F5DCF94B" box="[94,192,1753,1776]" italics="true" pageId="61" pageNumber="278">
<typeStatus id="2C405DCE243FFFBC753D67B3F5A6F94B" box="[94,186,1753,1776]" pageId="61" pageNumber="278" type="paratype">Paratypes</typeStatus>
:
</emphasis>
[Cachimbo, Prov.
<collectingRegion id="313F2D8E243FFFBC74E467B2F4D8F94B" box="[391,452,1752,1776]" country="Brazil" name="Bahia" pageId="61" pageNumber="278">Bahia</collectingRegion>
,
<collectorName id="5E0E86BA243FFFBC74B367B2F72AF94B" box="[464,566,1752,1776]" pageId="61" pageNumber="278">Ch. Pujol</collectorName>
1890], [COTYPE], [
<taxonomicName id="34FB98EF243FFFBC75906792F5B2F88B" authority="Fleut., FLEUTIAUX" authorityName="Fleut., FLEUTIAUX" authorityYear="1940" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="pujoli">Physodactylus pujoli Fleut., FLEUTIAUX</taxonomicName>
det. cotype], [Muséum Paris,
<collectorName id="5E0E86BA243FFFBC74926672F58AF8EB" pageId="61" pageNumber="278">Coll. E. Fleutiaux</collectorName>
], [
<typeStatus id="2C405DCE243FFFBC75CE6652F406F8EB" box="[173,282,1848,1872]" pageId="61" pageNumber="278" type="lectotype">Lectotype</typeStatus>
],
<specimenCount id="E5FD28E5243FFFBC74486652F440F8F4" box="[299,348,1848,1872]" count="1" pageId="61" pageNumber="278" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9243FFFBC74096652F4DAF8EB" box="[362,454,1848,1872]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="61" pageNumber="278">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,271 @@
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<taxonomicName id="34FB98EF243FFFBC778B645DF190FAEB" authority="Fleutiaux, 1940" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[744,1164,1335,1360]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="C18F3F7E243FFFBC778B645DF190FAEB" bold="true" box="[744,1164,1335,1360]" pageId="61" pageNumber="278">
<emphasis id="C18F3F7E243FFFBC778B645DF6C7FAF4" bold="true" box="[744,987,1335,1359]" italics="true" pageId="61" pageNumber="278">Physodactylus sulcatus</emphasis>
Fleutiaux, 1940
</emphasis>
</taxonomicName>
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<paragraph id="F344E36C243FFFBC762A6432F130FACB" blockId="61.[744,1164,1335,1392]" box="[841,1068,1368,1392]" pageId="61" pageNumber="278">
<emphasis id="C18F3F7E243FFFBC762A6432F130FACB" bold="true" box="[841,1068,1368,1392]" pageId="61" pageNumber="278">
(
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Figs.
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,
<figureCitation id="6BC0FFE9243FFFBC76D76432F6FAFACB" box="[948,998,1368,1392]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="61" pageNumber="278">33O</figureCitation>
,
<figureCitation id="6BC0FFE9243FFFBC76916432F138FACB" box="[1010,1060,1368,1392]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="61" pageNumber="278">35Q</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7243FFFBC77CB64F2F678F934" pageId="61" pageNumber="278" type="reference_group">
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<treatmentCitationGroup id="D3EBC442243FFFBC77CB64F2F1B3FA6B" pageId="61" pageNumber="278">
<treatmentCitation id="725AC57D243FFFBC77CB64F2F196FA0B" author="FLEUTIAUX, E." box="[680,1162,1432,1456]" page="165" pageId="61" pageNumber="278" year="1940">
<taxonomicName id="34FB98EF243FFFBC77CB64F2F196FA0B" authority="Fleutiaux, 1940 c: 165" authorityName="Fleutiaux, J. Chassain" authorityPageNumber="165" authorityYear="1940" box="[680,1162,1432,1456]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="C18F3F7E243FFFBC77CB64F2F69BFA0B" box="[680,903,1432,1456]" italics="true" pageId="61" pageNumber="278">Physodactylus sulcatus</emphasis>
<bibRefCitation id="976A9E9D243FFFBC76F564F2F196FA0B" author="FLEUTIAUX, E." box="[918,1162,1432,1456]" pageId="61" pageNumber="278" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 165</bibRefCitation>
</taxonomicName>
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, 167;
<treatmentCitation id="725AC57D243FFFBC778264D2F6CBFA6B" author="BLACKWELDER, R. E." box="[737,983,1464,1488]" page="292" pageId="61" pageNumber="278" year="1944">
<bibRefCitation id="976A9E9D243FFFBC778264D2F6CBFA6B" author="BLACKWELDER, R. E." box="[737,983,1464,1488]" pageId="61" pageNumber="278" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D243FFFBC768264D2F1B7FA6B" author="CHASSAIN, J." box="[993,1195,1464,1488]" page="70" pageId="61" pageNumber="278" year="2005">
<bibRefCitation id="976A9E9D243FFFBC768264D2F1B7FA6B" author="CHASSAIN, J." box="[993,1195,1464,1488]" pageId="61" pageNumber="278" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005: 70</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="F344E36C243FFFBC77CB64B2F620F994" blockId="61.[680,1228,1432,1680]" pageId="61" pageNumber="278">
<treatmentCitationGroup id="D3EBC442243FFFBC77CB64B2F620F994" pageId="61" pageNumber="278">
<treatmentCitation id="725AC57D243FFFBC77CB64B2F19BFA4B" author="FLEUTIAUX, E." box="[680,1159,1496,1520]" page="165" pageId="61" pageNumber="278" year="1940">
<taxonomicName id="34FB98EF243FFFBC77CB64B2F19BFA4B" authority="Fleutiaux, 1940 c: 165" authorityName="Fleutiaux, J. Chassain" authorityPageNumber="165" authorityYear="1940" box="[680,1159,1496,1520]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="C18F3F7E243FFFBC77CB64B2F667FA4B" box="[680,891,1496,1520]" italics="true" pageId="61" pageNumber="278">Physodactylus obesus</emphasis>
<bibRefCitation id="976A9E9D243FFFBC76EF64B2F19BFA4B" author="FLEUTIAUX, E." box="[908,1159,1496,1520]" pageId="61" pageNumber="278" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 167;
<treatmentCitation id="725AC57D243FFFBC77826492F6FFF9AB" author="BLACKWELDER, R. E." box="[737,995,1528,1552]" page="292" pageId="61" pageNumber="278" year="1944">
<bibRefCitation id="976A9E9D243FFFBC77826492F6FFF9AB" author="BLACKWELDER, R. E." box="[737,995,1528,1552]" pageId="61" pageNumber="278" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D243FFFBC76906492F1D4F9AB" author="CHASSAIN, J." box="[1011,1224,1528,1552]" page="70" pageId="61" pageNumber="278" year="2005">
<bibRefCitation id="976A9E9D243FFFBC76906492F1D4F9AB" author="CHASSAIN, J." box="[1011,1224,1528,1552]" pageId="61" pageNumber="278" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005: 70</bibRefCitation>
</treatmentCitation>
; syn. nov.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F344E36C243FFFBC77CB6752F678F934" blockId="61.[680,1228,1432,1680]" pageId="61" pageNumber="278">
<treatmentCitationGroup id="D3EBC442243FFFBC77CB6752F678F934" pageId="61" pageNumber="278">
<treatmentCitation id="725AC57D243FFFBC77CB6752F197F9EB" author="FLEUTIAUX, E." box="[680,1163,1592,1616]" page="165" pageId="61" pageNumber="278" year="1940">
<taxonomicName id="34FB98EF243FFFBC77CB6752F197F9EB" authority="Fleutiaux, 1940 c: 165" authorityName="Fleutiaux" authorityPageNumber="165" authorityYear="1940" box="[680,1163,1592,1616]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="278" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="C18F3F7E243FFFBC77CB6752F690F9EB" box="[680,908,1592,1616]" italics="true" pageId="61" pageNumber="278">Physodactylus testaceus</emphasis>
<bibRefCitation id="976A9E9D243FFFBC76FA6752F197F9EB" author="FLEUTIAUX, E." box="[921,1163,1592,1616]" pageId="61" pageNumber="278" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux, 1940c: 165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 167;
<treatmentCitation id="725AC57D243FFFBC77826732F6FFF9CB" author="BLACKWELDER, R. E." box="[737,995,1624,1648]" page="292" pageId="61" pageNumber="278" year="1944">
<bibRefCitation id="976A9E9D243FFFBC77826732F6FFF9CB" author="BLACKWELDER, R. E." box="[737,995,1624,1648]" pageId="61" pageNumber="278" pagination="280 - 303" refId="ref48842" refString="BLACKWELDER, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. Bulletin of the United States National Museum, 185 (2): 280 - 303." type="journal article" year="1944">Blackwelder, 1944: 292</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="725AC57D243FFFBC76906732F1D4F9CB" author="CHASSAIN, J." box="[1011,1224,1624,1648]" page="68" pageId="61" pageNumber="278" year="2005">
<bibRefCitation id="976A9E9D243FFFBC76906732F1D4F9CB" author="CHASSAIN, J." box="[1011,1224,1624,1648]" pageId="61" pageNumber="278" pagination="65 - 71" refId="ref49320" refString="CHASSAIN, J. 2005. Description de trois nouvelles especes de Physodactilinae du Mato Grosso (Bresil) (Coleoptera, Elateridae). Boletin Sociedad Entomologica Aragonesa, 36: 65 - 71." type="journal article" year="2005">Chassain, 2005: 68</bibRefCitation>
</treatmentCitation>
, 70; syn. nov.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BBE1B0E7243FFFBF77CB67D2F1F8FB2B" lastPageId="62" lastPageNumber="279" pageId="61" pageNumber="278" type="description">
<paragraph id="F344E36C243FFFBF77CB67D2F4E9F954" blockId="61.[680,1229,1720,1872]" lastBlockId="62.[132,681,184,1776]" lastPageId="62" lastPageNumber="279" pageId="61" pageNumber="278">
<emphasis id="C18F3F7E243FFFBC77CB67D2F636F96B" box="[680,810,1720,1744]" italics="true" pageId="61" pageNumber="278">Redescription</emphasis>
(male,
<figureCitation id="6BC0FFE9243FFFBC76E367D2F6F9F96B" box="[896,997,1720,1744]" captionStart="FIGURE 35" captionStartId="66.[132,211,1749,1768]" captionTargetBox="[134,1266,517,1725]" captionTargetId="figure-119@66.[132,1266,516,1725]" captionTargetPageId="66" captionText="FIGURE 35: Habitus. A, Physodactylus brasiliensis (12.8 mm); B, P. brunneus sp. nov. (9.0 mm); C, P. chassaini sp. nov. (11.5 mm); D, P. costae (12.0 mm); E, P. fischeri (17.0 mm); F, P. flavifrons sp. nov. (10.0 mm); G, P. fleutiauxi (15.0 mm); H, P. foveatostriatus (12.0 mm); I, P. girardi sp. nov. (9.0 mm); J, P. gounellei sp. nov. (10.5 mm); K, P. henningi (15.0 mm); L, P. latithorax sp. nov. (18.0 mm); M, P. niger (11.0 mm); N, P. oberthuri (12.5 mm); O, P. patens sp. nov. (17.0 mm); P,P. pujoli (12.0 mm); Q, P. sulcatus (9.0 mm); R, P. tuberculatus sp. nov. (12.0 mm). A-D, F-J, L-R = holotypes." figureDoi="http://doi.org/10.5281/zenodo.12641457" httpUri="https://zenodo.org/record/12641457/files/figure.png" pageId="61" pageNumber="278">Fig. 35Q</figureCitation>
): Integument entirely bright or with elytral interstices dull; pilosity yellow; color patterns: 1) dark brown with prothorax orange; 2) head and elytra black, antenna, ventral pterothorax, legs and ventral abdomen dark brown, prothorax and epipleura orange, scutellum dark orange; 3) head, elytra, prosternum apically, ventral pterothorax and abdomen black, prothorax (except prosternum apically), epipleura and apical elytral suture light to dark orange, with or without an orange spot on base and apex of elytra, ventral abdomen evenly colored or with lateral borders lighter; 4) as the former but antenna orange and femur entirely or partly orange; 5) as pattern 3 but pronotal disc dark brown; 6) antenna, prothorax (except prosternum apically), elytra and epipleura brown, head, prosternum apically, ventral pterothorax and abdomen dark brown to black, legs dark brown with femur lighter. Total length 8.0-15.0 mm; elytral base 1.0-1.1x wider than prothorax, elytra 3.0-3.4x longer than pronotum. Frons concave to convex, frontal carina not or slightly produced; antenna (
<figureCitation id="6BC0FFE9243CFFBF75EF63D2F5F6FD6B" box="[140,234,696,720]" captionStart="FIGURE 30" captionStartId="61.[95,174,685,704]" captionTargetBox="[96,1225,171,657]" captionTargetId="figure-450@61.[94,1228,170,661]" captionTargetPageId="61" captionText="FIGURE 30: Physodactylus sulcatus. A, antenna; B, maxilla; C, labrum and mandible; D, E, prothorax (dorsal, lateral); F, G, pro and" figureDoi="http://doi.org/10.5281/zenodo.12641441" httpUri="https://zenodo.org/record/12641441/files/figure.png" pageId="62" pageNumber="279">Fig. 30A</figureCitation>
) reaching or surpassing the anterior third of pronotum but not reaching its half length; antennomere IV 1.3-1.4x wider than long, V-X 1.6-1.86x wider than long, XI circular to oval 1.0-1.1x times as long as wide. Maxilla (
<figureCitation id="6BC0FFE9243CFFBF74146252F4C9FCEB" box="[375,469,824,848]" captionStart="FIGURE 30" captionStartId="61.[95,174,685,704]" captionTargetBox="[96,1225,171,657]" captionTargetId="figure-450@61.[94,1228,170,661]" captionTargetPageId="61" captionText="FIGURE 30: Physodactylus sulcatus. A, antenna; B, maxilla; C, labrum and mandible; D, E, prothorax (dorsal, lateral); F, G, pro and" figureDoi="http://doi.org/10.5281/zenodo.12641441" httpUri="https://zenodo.org/record/12641441/files/figure.png" pageId="62" pageNumber="279">Fig. 30B</figureCitation>
) with galea covered with fine and spiniform setae. Pronotum (
<figureCitation id="6BC0FFE9243CFFBF77236232F7BCFCCB" box="[576,672,856,880]" captionStart="FIGURE 30" captionStartId="61.[95,174,685,704]" captionTargetBox="[96,1225,171,657]" captionTargetId="figure-450@61.[94,1228,170,661]" captionTargetPageId="61" captionText="FIGURE 30: Physodactylus sulcatus. A, antenna; B, maxilla; C, labrum and mandible; D, E, prothorax (dorsal, lateral); F, G, pro and" figureDoi="http://doi.org/10.5281/zenodo.12641441" httpUri="https://zenodo.org/record/12641441/files/figure.png" pageId="62" pageNumber="279">Fig. 30D</figureCitation>
) convex 1.3-1.4x wider than long, parallel on posterior third then straight or roundly convergent to anterior margin; anteromedian margin slightly produced anteriad; posterior angles short, convex, parallel; lateral carina evenly prominent to anterior margin or weaker impressed to absent on anterior third; dorsally visible only on posterior third; disc with punctures medium homogenously distributed 0.5-1.0 diameters apart, smaller and denser on anterior margin, denser on lateral border; prosternal process without subapical tooth (
<figureCitation id="6BC0FFE9243CFFBF75A965D2F43AFB6B" box="[202,294,1208,1232]" captionStart="FIGURE 30" captionStartId="61.[95,174,685,704]" captionTargetBox="[96,1225,171,657]" captionTargetId="figure-450@61.[94,1228,170,661]" captionTargetPageId="61" captionText="FIGURE 30: Physodactylus sulcatus. A, antenna; B, maxilla; C, labrum and mandible; D, E, prothorax (dorsal, lateral); F, G, pro and" figureDoi="http://doi.org/10.5281/zenodo.12641441" httpUri="https://zenodo.org/record/12641441/files/figure.png" pageId="62" pageNumber="279">Fig. 30E</figureCitation>
). Lamellae of pro- and mesotarsomeree I very small, minuscule or absent on metatarsomere I. Elytra parallel-sided on anterior 2/3 then tapering to apex; apex usually with a very small sutural spine; striae (
<figureCitation id="6BC0FFE9243CFFBF75A86452F432FAEB" box="[203,302,1336,1360]" captionStart="FIGURE 33" captionStartId="64.[132,211,1800,1819]" captionTargetBox="[132,1266,567,1776]" captionTargetId="figure-187@64.[132,1266,567,1776]" captionTargetPageId="64" captionText="FIGURE 33: Dactylophysus hirtus sp. nov. A, B, C, pro-, meso- and metathoracic legs; D, apex of protibia. Physodactylus niger. E, labrum. (lateroanterior). Physodactylus henningi. F, prothoracic leg; G, protibia; H, mesothoracic leg; I, mesotibia; J, metathoracic leg; K, metatibia; L, apex of metatibia; M, lamella of protarsomere I. Physodactylus niger. N, elytral striae. Physodactylus sulcatus. O, elytral striae." figureDoi="http://doi.org/10.5281/zenodo.12641451" httpUri="https://zenodo.org/record/12641451/files/figure.png" pageId="62" pageNumber="279">Fig. 33O</figureCitation>
) deeply impressed, with three rows of punctures, punctures glabrous on the median row, smaller and bearing a small seta on the lateral row; interstices convex, smooth and glabrous; apical striae with punctures smaller than those of pronotal lateral border and as large as those of elytral base. Abdomen with ventrites 1-4 strongly and evenly convex, ventrite 2 evenly pilose with short decumbent setae, II-IV with mixed short decumbent and long semi-erect setae, longer setae denser on median region. Pregenitalic segments and aedeagus covered with stout yellow setae. Sternite VIII subrectangular, posterior margin straight, anterior sclerotization 0.6x the total width of sternite; sternite IX tapered to apex.
</paragraph>
<paragraph id="F344E36C243CFFBF75E76672F1DDFEEB" blockId="62.[132,681,1816,1872]" lastBlockId="62.[718,1267,184,336]" pageId="62" pageNumber="279">
<emphasis id="C18F3F7E243CFFBF75E76672F5F8F88B" box="[132,228,1816,1840]" italics="true" pageId="62" pageNumber="279">Aedeagus:</emphasis>
Phallobase 0.5x the total length of aedeagus, 1.2x longer than wide, length ratio between lateroposterior and median parts 3.5-5.2; ventral surface of paramere with the lateral half and anterior margin sclerotized; penis with basal strut 0.2-0.3x its total length, articulated to the parameres by a strongly sclerotized process; ventral sclerite parallel-sided.
</paragraph>
<paragraph id="F344E36C243CFFBF77AD6012F6BAFDAB" blockId="62.[718,1267,376,528]" pageId="62" pageNumber="279">
<emphasis id="C18F3F7E243CFFBF77AD6012F637FE2B" box="[718,811,376,400]" italics="true" pageId="62" pageNumber="279">
<typeStatus id="2C405DCE243CFFBF77AD6012F63AFE2B" box="[718,806,376,400]" pageId="62" pageNumber="279" type="holotype">Holotype</typeStatus>
:
</emphasis>
[Jatahy,
<collectingRegion id="313F2D8E243CFFBF76E76012F6DFFE2B" box="[900,963,376,400]" country="Brazil" name="Goias" pageId="62" pageNumber="279">Goyas</collectingRegion>
<collectingCountry id="8BECA3FC243CFFBF76A96012F115FE2B" box="[970,1033,376,400]" name="Brazil" pageId="62" pageNumber="279">Brèsil</collectingCountry>
], [
<taxonomicName id="34FB98EF243CFFBF717E6012F62FFE0B" authority="Fleut." authorityName="Fleutiaux, J. Chassain" authorityYear="1940" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="sulcatus">Physodactylus sulcatus Fleut.</taxonomicName>
COLLECTION FLEUTIAUX], [TYPE], [Muséum Paris, Coll. E. Fleutiaux], [
<typeStatus id="2C405DCE243CFFBF711860D2F1F4FE6B" box="[1147,1256,440,464]" pageId="62" pageNumber="279" type="lectotype">Lectotype</typeStatus>
], [
<taxonomicName id="34FB98EF243CFFBF77B760B2F1A2FE4B" authority="Fleutiaux, J. Chassain" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[724,1214,472,496]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="sulcatus">Physodactylus sulcatus Fleutiaux, J. Chassain</taxonomicName>
det. 05], male (MNHN).
</paragraph>
<paragraph id="F344E36C243CFFBF77AD6352F1F7FD0B" blockId="62.[718,1267,568,688]" pageId="62" pageNumber="279">
<emphasis id="C18F3F7E243CFFBF77AD6352F68AFDEB" box="[718,918,568,592]" italics="true" pageId="62" pageNumber="279">Material examined:</emphasis>
<materialsCitation id="4393E931243CFFBF76C06352F626FDCB" collectionCode="MZUSP" country="Brazil" location="Mato Grosso" pageId="62" pageNumber="279" specimenCount="10" stateProvince="Mato Grosso">
<collectingCountry id="8BECA3FC243CFFBF76C06352F11EFDEB" box="[931,1026,568,592]" name="Brazil" pageId="62" pageNumber="279">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E243CFFBF71726353F185FDF4" box="[1041,1177,568,592]" italics="true" pageId="62" pageNumber="279">
<collectingRegion id="313F2D8E243CFFBF71726353F18FFDF4" box="[1041,1171,568,592]" country="Brazil" name="Mato Grosso" pageId="62" pageNumber="279">Mato Grosso</collectingRegion>
:
</emphasis>
<specimenCount id="E5FD28E5243CFFBF71C66352F1EFFDF4" box="[1189,1267,568,592]" count="10" pageId="62" pageNumber="279" type="generic">10 exs.</specimenCount>
(
<collectionCode id="95EA7BA9243CFFBF77B66332F62FFDCB" box="[725,819,600,624]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="62" pageNumber="279">MZUSP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="4393E931243CFFBF76336332F6C8FD2B" collectionCode="CNC, MNHN" country="Brazil" pageId="62" pageNumber="279" specimenCount="8" stateProvince="Mato Grosso" typeStatus="holotype">
<specimenCount id="E5FD28E5243CFFBF76336332F68AFDD4" box="[848,918,600,624]" count="8" pageId="62" pageNumber="279" type="generic">8 exs.</specimenCount>
(
<collectionCode id="95EA7BA9243CFFBF76CD6332F6F0FDCB" box="[942,1004,600,624]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="62" pageNumber="279">CNC</collectionCode>
),
<typeStatus id="2C405DCE243CFFBF71696332F178FDCB" box="[1034,1124,600,624]" pageId="62" pageNumber="279" type="holotype">holotype</typeStatus>
of
<taxonomicName id="34FB98EF243CFFBF71FF6333F7E8FD2B" authorityName="Fleutiaux" authorityYear="1940" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="testaceus" status="syn. nov.">
<emphasis id="C18F3F7E243CFFBF71FF6333F7E8FD2B" italics="true" pageId="62" pageNumber="279">P. testaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243CFFBF779C6312F642FD34" box="[767,862,632,656]" pageId="62" pageNumber="279">syn. nov.</taxonomicNameLabel>
(
<collectionCode id="95EA7BA9243CFFBF76136312F6D0FD2B" box="[880,972,632,656]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="62" pageNumber="279">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4393E931243CFFBF76826313F1FBFD0B" collectionCode="MNHN" country="Brazil" location="Mato Grosso do Sul" pageId="62" pageNumber="279" specimenCount="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
<emphasis id="C18F3F7E243CFFBF76826313F1AFFD2B" box="[993,1203,632,656]" italics="true" pageId="62" pageNumber="279">
<collectingRegion id="313F2D8E243CFFBF76826313F1B2FD2B" box="[993,1198,632,656]" country="Brazil" name="Mato Grosso do Sul" pageId="62" pageNumber="279">Mato Grosso do Sul</collectingRegion>
:
</emphasis>
<specimenCount id="E5FD28E5243CFFBF71DD6312F1EFFD34" box="[1214,1267,632,656]" count="1" pageId="62" pageNumber="279" type="generic">1 ex.</specimenCount>
(
<collectionCode id="95EA7BA9243CFFBF77B563F2F62EFD0B" box="[726,818,664,688]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="62" pageNumber="279">MNHN</collectionCode>
);
<typeStatus id="2C405DCE243CFFBF762763F2F682FD0B" box="[836,926,664,688]" pageId="62" pageNumber="279" type="holotype">holotype</typeStatus>
of
<taxonomicName id="34FB98EF243CFFBF76A363F3F108FD0B" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[960,1044,664,688]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="obesus" status="syn. nov.">
<emphasis id="C18F3F7E243CFFBF76A363F3F108FD0B" box="[960,1044,664,688]" italics="true" pageId="62" pageNumber="279">P. obesus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DABC8205243CFFBF717963F2F169FD14" box="[1050,1141,664,688]" pageId="62" pageNumber="279">syn. nov.</taxonomicNameLabel>
(
<collectionCode id="95EA7BA9243CFFBF71E063F2F1C3FD0B" box="[1155,1247,664,688]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="62" pageNumber="279">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F344E36C243CFFBF77AD63B2F1CEFCAB" blockId="62.[718,1266,728,784]" pageId="62" pageNumber="279">
<emphasis id="C18F3F7E243CFFBF77AD63B2F64CFD4B" box="[718,848,728,752]" italics="true" pageId="62" pageNumber="279">Distribution:</emphasis>
<collectingCountry id="8BECA3FC243CFFBF763B63B2F6ABFD4B" box="[856,951,728,752]" name="Brazil" pageId="62" pageNumber="279">BRAZIL</collectingCountry>
.
<emphasis id="C18F3F7E243CFFBF76A263B3F158FD54" box="[961,1092,728,752]" italics="true" pageId="62" pageNumber="279">
<collectingRegion id="313F2D8E243CFFBF76A263B3F123FD54" box="[961,1087,728,752]" country="Brazil" name="Mato Grosso" pageId="62" pageNumber="279">Mato Grosso</collectingRegion>
:
</emphasis>
Cuiabá, Rosário Oeste, Sinop, Vera;
<emphasis id="C18F3F7E243CFFBF76F96393F17CFCAB" box="[922,1120,760,784]" italics="true" pageId="62" pageNumber="279">
<collectingRegion id="313F2D8E243CFFBF76F96393F147FCAB" box="[922,1115,760,784]" country="Brazil" name="Mato Grosso do Sul" pageId="62" pageNumber="279">Mato Grosso do Sul</collectingRegion>
:
</emphasis>
Corumbá.
</paragraph>
<paragraph id="F344E36C243CFFBF77AD6252F1F8FB2B" blockId="62.[718,1267,824,1168]" pageId="62" pageNumber="279">
<emphasis id="C18F3F7E243CFFBF77AD6252F63BFCEB" box="[718,807,824,848]" italics="true" pageId="62" pageNumber="279">Remarks:</emphasis>
<bibRefCitation id="976A9E9D243CFFBF764F6252F6C3FCEB" author="FLEUTIAUX, E." box="[812,991,824,848]" pageId="62" pageNumber="279" pagination="164 - 168" refId="ref49980" refString="FLEUTIAUX, E. 1940 c. Revision des Physodactylus Fischer (Col. Elateridae). Revue Francaise d'Entomologie, 7: 164 - 168." type="journal article" year="1940">Fleutiaux (1940c)</bibRefCitation>
described
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<taxonomicName id="34FB98EF243CFFBF712B6252F61DFCCB" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="sulcatus">Physodactylus sulcatus</taxonomicName>
,
<taxonomicName id="34FB98EF243CFFBF76696233F647FCCB" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[778,859,856,880]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="obesus">P. obesus</taxonomicName>
</emphasis>
and
<emphasis id="C18F3F7E243CFFBF76EE6233F6E6FCCB" box="[909,1018,857,880]" italics="true" pageId="62" pageNumber="279">
<taxonomicName id="34FB98EF243CFFBF76EE6233F6EAFCCB" authorityName="Fleutiaux" authorityYear="1940" box="[909,1014,857,880]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="testaceus">P. testaceus</taxonomicName>
,
</emphasis>
each one based on a single specimen. He distinguished them by color pattern, integument brightness and size. However, the analysis of a larger series showed that the differences in those features are variations. Specimens from Vera (MT) exhibit all color patterns, including specimens with the same color (pattern 6) and size of
<taxonomicName id="34FB98EF243CFFBF71466573F190FB8B" authorityName="Fleutiaux" authorityYear="1940" box="[1061,1164,1049,1072]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="C18F3F7E243CFFBF71466573F190FB8B" box="[1061,1164,1049,1072]" italics="true" pageId="62" pageNumber="279">P. testaceus</emphasis>
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. Material examined from Sinop (MT) includes specimens as small as the
<taxonomicName id="34FB98EF243CFFBF76CB6533F117FBCB" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[936,1035,1112,1136]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="sulcatus">
<emphasis id="C18F3F7E243CFFBF76CB6533F117FBCB" box="[936,1035,1112,1136]" italics="true" pageId="62" pageNumber="279">P. sulcatus</emphasis>
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<typeStatus id="2C405DCE243CFFBF71706532F170FBCB" box="[1043,1132,1112,1136]" pageId="62" pageNumber="279" type="holotype">holotype</typeStatus>
(
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) and specimens as large as the
<taxonomicName id="34FB98EF243CFFBF76A86513F101FB2B" authorityName="Fleutiaux, J. Chassain" authorityYear="1940" box="[971,1053,1144,1168]" class="Insecta" family="Elateridae" genus="Physodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="279" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="C18F3F7E243CFFBF76A86513F101FB2B" box="[971,1053,1144,1168]" italics="true" pageId="62" pageNumber="279">P. obesus</emphasis>
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<typeStatus id="2C405DCE243CFFBF71406512F160FB2B" box="[1059,1148,1144,1168]" pageId="62" pageNumber="279" type="holotype">holotype</typeStatus>
(
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).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,143 +1,142 @@
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<taxonomicName id="7DDB932AAFBAA099EF2542E39B5F12A7" authority="sp." class="Insecta" family="Braconidae" genus="Micro" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micro sofron" order="Hymenoptera" pageId="13" pageNumber="62" phylum="Arthropoda" rank="species" species="sofron">Micro plitis sp. nr. sofron</taxonomicName>
Fig. 30
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<paragraph pageId="14" pageNumber="63">
<pageBreakToken pageId="14" pageNumber="63" start="start">Distribution</pageBreakToken>
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<paragraph id="7031EA315EAC8E6626CE874AF4CDDDB4" pageId="14" pageNumber="63">
<pageBreakToken id="B68068D40461C3236BC4084ABD6D1AF4" pageId="14" pageNumber="63" start="start">Distribution</pageBreakToken>
.
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<paragraph pageId="14" pageNumber="63">NEA. Probably a High Arctic endemic.</paragraph>
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<paragraph pageId="14" pageNumber="63">
<paragraph id="D71042D3B16F6CF216E97486428925C8" pageId="14" pageNumber="63">NEA. Probably a High Arctic endemic.</paragraph>
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<paragraph id="CC39617E22618FD295DCC81D9C545F6E" pageId="14" pageNumber="63">
Figure 30.
<taxonomicName class="Insecta" family="Braconidae" genus="Microplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microplitis sofron" order="Hymenoptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="sofron">Microplitis sp. near sofron</taxonomicName>
<taxonomicName id="8E746B5AD77F04B0F92CDE5BA62F9106" class="Insecta" family="Braconidae" genus="Microplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microplitis sofron" order="Hymenoptera" pageId="14" pageNumber="63" phylum="Arthropoda" rank="species" species="sofron">Microplitis sp. near sofron</taxonomicName>
. A Habitus, lateral B Fore wing C Mesosoma, dorsal D Head, frontal E Metasoma, dorsal.
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<paragraph pageId="14" pageNumber="63">Notes.</paragraph>
<paragraph pageId="14" pageNumber="63">
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<paragraph id="3AE0BCB4199444685D0B5796BA083A10" pageId="14" pageNumber="63">Notes.</paragraph>
<paragraph id="205BC8CDE10781B60646C37A0871ED88" pageId="14" pageNumber="63">
A total of 12 specimens from Banks and Victoria Islands. This species will run to
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<taxonomicName id="494049B00F3634EE042B2D0D9766EA58" lsidName="M. sofron" pageId="14" pageNumber="63" rank="species" species="sofron">M. sofron</taxonomicName>
in the keys provided by
<bibRefCitation author="Nixon, GEJ" journalOrPublisher="Bulletin of the British Museum (Natural History), Entomology series" pageId="16" pageNumber="65" pagination="1 - 30" title="A revision of the n. w. European species of Microplitis Foerster (Hymenoptera: Braconidae)." volume="25" year="1970">Nixon (1970)</bibRefCitation>
<bibRefCitation id="E1B592AE6FE4CAB6C5003A2BD5FFCB96" author="Nixon, GEJ" journalOrPublisher="Bulletin of the British Museum (Natural History), Entomology series" pageId="16" pageNumber="65" pagination="1 - 30" title="A revision of the n. w. European species of Microplitis Foerster (Hymenoptera: Braconidae)." volume="25" year="1970">Nixon (1970)</bibRefCitation>
,
<bibRefCitation author="Papp, J" journalOrPublisher="Entomologische Abhandlungen," pageId="16" pageNumber="65" pagination="95 - 140" title="Palaearctic species of Microgaster Latreille (= Microplitis Foerster) with description of seven new species (Hymenoptera, Braconidae, Microgastrinae)." volume="47" year="1984">Papp (1984)</bibRefCitation>
<bibRefCitation id="B2F7B2624A1BBFF7D1D71B6EF8C6DB80" author="Papp, J" journalOrPublisher="Entomologische Abhandlungen," pageId="16" pageNumber="65" pagination="95 - 140" title="Palaearctic species of Microgaster Latreille (= Microplitis Foerster) with description of seven new species (Hymenoptera, Braconidae, Microgastrinae)." volume="47" year="1984">Papp (1984)</bibRefCitation>
, and
<bibRefCitation author="van Achterberg, C" journalOrPublisher="Zoologische Mededelingen" pageId="15" pageNumber="64" pagination="13 - 62" title="The Braconidae (Hymenoptera) of Greenland." volume="80" year="2006">van Achterberg (2006)</bibRefCitation>
<bibRefCitation id="0C56A99713E2D833DC989ED6CEBD3875" author="van Achterberg, C" journalOrPublisher="Zoologische Mededelingen" pageId="15" pageNumber="64" pagination="13 - 62" title="The Braconidae (Hymenoptera) of Greenland." volume="80" year="2006">van Achterberg (2006)</bibRefCitation>
. However, its metatibia is not bright yellow, and the shape of mediotergite 1 does not resemble the illustration of
<bibRefCitation author="Papp, J" journalOrPublisher="Entomologische Abhandlungen," pageId="16" pageNumber="65" pagination="95 - 140" title="Palaearctic species of Microgaster Latreille (= Microplitis Foerster) with description of seven new species (Hymenoptera, Braconidae, Microgastrinae)." volume="47" year="1984">Papp (1984</bibRefCitation>
<bibRefCitation id="3D15BF54878AF857A92008BA9AA4D996" author="Papp, J" journalOrPublisher="Entomologische Abhandlungen," pageId="16" pageNumber="65" pagination="95 - 140" title="Palaearctic species of Microgaster Latreille (= Microplitis Foerster) with description of seven new species (Hymenoptera, Braconidae, Microgastrinae)." volume="47" year="1984">Papp (1984</bibRefCitation>
: figure 83). The color of metatibia and shape of mediotergite 1 are actually closer to
<taxonomicName lsidName="M. lugubroides" pageId="14" pageNumber="63" rank="species" species="lugubroides">M. lugubroides</taxonomicName>
<taxonomicName id="C635D574BF4DAB12A63DDE3558C589AB" lsidName="M. lugubroides" pageId="14" pageNumber="63" rank="species" species="lugubroides">M. lugubroides</taxonomicName>
, but from that species it differs in the length of the last flagellomere, the main feature that van Achterberg used to separate
<taxonomicName lsidName="M. sofron" pageId="14" pageNumber="63" rank="species" species="sofron">M. sofron</taxonomicName>
<taxonomicName id="5CC9AFADF658625369A6C99F62D32230" lsidName="M. sofron" pageId="14" pageNumber="63" rank="species" species="sofron">M. sofron</taxonomicName>
from
<taxonomicName lsidName="M. lugubroides" pageId="14" pageNumber="63" rank="species" species="lugubroides">M. lugubroides</taxonomicName>
<taxonomicName id="A5C70E5C5E2D9017ED4541012A8FF589" lsidName="M. lugubroides" pageId="14" pageNumber="63" rank="species" species="lugubroides">M. lugubroides</taxonomicName>
. The Canadian specimens probably represent a new species, but without examining the types of
<taxonomicName lsidName="sofron" pageId="14" pageNumber="63" rank="species" species="sofron">sofron</taxonomicName>
<taxonomicName id="9C8640CA46DC4A1B3F8530FA1C0EEB0B" lsidName="sofron" pageId="14" pageNumber="63" rank="species" species="sofron">sofron</taxonomicName>
and
<taxonomicName lsidName="lugubroides" pageId="14" pageNumber="63" rank="species" species="lugubroides">lugubroides</taxonomicName>
<taxonomicName id="9385F032ACE9D86215E12994F5BEE7BC" lsidName="lugubroides" pageId="14" pageNumber="63" rank="species" species="lugubroides">lugubroides</taxonomicName>
we cannot be certain. No DNA sequences are available for this species.
</paragraph>
</subSubSection>