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<document id="70CCA8AC76CAF272FCB3AC5219477774" ID-CLB-Dataset="299340" ID-DOI="10.3897/zookeys.1205.120033" ID-GBIF-Dataset="15d96bea-75a1-4b35-a306-3aecdc1513a7" ID-ZooBank="1093570E-D1C7-4918-BD8F-0BC3D3F486E5" ID-publisher-id="120033" URI-arpha="541CAC61-3728-5591-9EFB-B1793F52FBE7" URI-zoobank="http://zoobank.org/1093570E-D1C7-4918-BD8F-0BC3D3F486E5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1719861896249" checkinUser="pensoft" docAuthor="Wood, Thomas J., Ismael, Halgurd R., Baiocchi, Daniele, Hamad, Mudhafar I., Bapir, Tahseen T. &amp; Selis, Marco" docDate="2024" docId="64180BEE5A8355D4A3089FA3C2E06A0F" docLanguage="en" docName="ZooKeys 1205: 267-298" docOrigin="ZooKeys 1205" docSource="https://zookeys.pensoft.net/article/120033/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Andrena (Aciandrena) duhokensis Wood &amp; Ismael &amp; Baiocchi &amp; Hamad &amp; Bapir &amp; Selis 2024, sp. nov." docType="treatment" docUuid="3C35CEE1-7063-4C23-8577-1BA21A6CCAFE" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="298" masterDocId="541CAC61372855919EFBB1793F52FBE7" masterDocTitle="A first revision of the Andrena of Iraq (Hymenoptera, Andrenidae), with the description of two new species from Iraqi Kurdistan and additional records from surrounding countries" masterLastPageNumber="298" masterPageNumber="267" pageNumber="267" updateTime="1719886393335" updateUser="ExternalLinkService">
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<mods:title id="CA72CDFA1AE4775420E8566EDF7FC235">A first revision of the Andrena of Iraq (Hymenoptera, Andrenidae), with the description of two new species from Iraqi Kurdistan and additional records from surrounding countries</mods:title>
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<mods:namePart id="0C7337BA37EF94BF1D3EDA9FDD7A7500">Wood, Thomas J.</mods:namePart>
<mods:nameIdentifier id="0EA05B0A735EDC56FB15A0E53A494237" type="ORCID">0000-0001-5653-224X</mods:nameIdentifier>
<mods:affiliation id="069CF9F7BEC4B5254C9CD49B329B9DEF">Naturalis Biodiversity Center, 2333 CR, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name id="6720427BAC4C5F0DE730238202B1FD94" type="personal">
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</mods:role>
<mods:namePart id="82E815EF445D9422736BD0F9DF6A34E2">Ismael, Halgurd R.</mods:namePart>
<mods:affiliation id="767EDE94C560D1E1438362852717A26D">Department of Plant Protection, College of Agricultural Engineering Sciences, University of Duhok, Duhok, Kurdistan Region, Iraq</mods:affiliation>
</mods:name>
<mods:name id="174387A2E60B06097BC2349E22A85F74" type="personal">
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<mods:namePart id="79E46BCCA4A32209D82845DF4D07E0AF">Baiocchi, Daniele</mods:namePart>
<mods:affiliation id="DEB175CAFE0E8252074756100A38E828">Via Matteo Babini 26, 00139, Roma, Italy</mods:affiliation>
</mods:name>
<mods:name id="560729E3ED4438415842A2178BB80927" type="personal">
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<mods:namePart id="098F4FCCF9221038D5DDE821B6FA63A8">Hamad, Mudhafar I.</mods:namePart>
<mods:affiliation id="66979B4D12FF512D8D9FB6D0373D0F82">Khabat Technical Institute, Erbil Polytechnic University, Erbil, Kurdistan Region, Iraq</mods:affiliation>
</mods:name>
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<mods:namePart id="0A1DAA23D67CB7004B2E7354330B65BC">Bapir, Tahseen T.</mods:namePart>
<mods:affiliation id="315C7D8675BE2734A52E6896EC9E52D1">Department of Plant Protection, College of Agricultural Engineering Sciences, University of Duhok, Duhok, Kurdistan Region, Iraq</mods:affiliation>
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<mods:namePart id="5BEC51190666191AE2270CCEF608152E">Selis, Marco</mods:namePart>
<mods:nameIdentifier id="797D79BB9AB56168DB8FD16762AC7390" type="ORCID">0000-0002-8200-9441</mods:nameIdentifier>
<mods:affiliation id="B445CDDC14C5BF95288181901053DF72">Via dei Tarquini 22, 01100 VT, Viterbo, Italy</mods:affiliation>
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<mods:date id="93D1DBBAFED10D27C4C5FAA38504D93C">2024</mods:date>
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<treatment id="64180BEE5A8355D4A3089FA3C2E06A0F" ID-DOI="http://doi.org/10.5281/zenodo.12615775" ID-GBIF-Taxon="234396735" ID-Zenodo-Dep="12615775" ID-arpha="64180BEE-5A83-55D4-A308-9FA3C2E06A0F" ID-zoobank="https://zoobank.org/3C35CEE1-7063-4C23-8577-1BA21A6CCAFE" LSID="urn:lsid:zoobank.org:act:3C35CEE1-7063-4C23-8577-1BA21A6CCAFE" LSID-ZBK="urn:lsid:zoobank.org:act:3C35CEE1-7063-4C23-8577-1BA21A6CCAFE" httpUri="http://treatment.plazi.org/id/64180BEE5A8355D4A3089FA3C2E06A0F">
<subSubSection id="BB6995671859C9DFA104E977F7845CF3" type="nomenclature">
<paragraph id="0AA6C8214F5028E33964942D7D2E575F">
<heading id="3F133AC0F59106E4E3F86E11EFCCB7F8" reason="label">20.</heading>
<taxonomicName id="7E5DFF77950880DA2CA653D52DC95737" authority="Wood" authorityName="Wood &amp; Ismael &amp; Baiocchi &amp; Hamad &amp; Bapir &amp; Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis" status="sp. nov." subGenus="Aciandrena">
<emphasis id="D1255133B2BF3DD8CF90599C645FBBEC" italics="true">Andrena</emphasis>
(
<emphasis id="EF7AACFDE2A85F711CBA08CE24BE2079" italics="true">Aciandrena</emphasis>
)
<emphasis id="87C597DC5258A5E709D5CCAE1AF01FC1" italics="true">duhokensis</emphasis>
Wood
</taxonomicName>
,
<taxonomicNameLabel id="BAD01DDB0652A61ADE5D6DACE7E78281" rank="species">sp. nov.</taxonomicNameLabel>
*
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVGBG" type="material">
<paragraph id="140AEC86B833FA08ABE15C0AEBDB8132">
<heading id="A8A1A87C83F94B76DB98DCA2CB17B212" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="00E8930BB27594B42AA3C946E2EE3D9D">
<materialsCitation id="7ADB7E1F159A56B62683892C1F21D014" ID-GBIF-Occurrence="4900833338" collectingDate="2023-05-11" collectionCode="RMNH" collectorName="D. Baiocchi" country="Iraq" elevation="1912" latitude="37.0158" location="Mt. Gara" longLatPrecision="6" longitude="43.3506" specimenCount="1" specimenCount-male="1" stateProvince="Duhok" typeStatus="holotype">
<emphasis id="54F8A1EA2A7308BD227EA7196BDDCEE2" bold="true">
<emphasis id="F56CD63A55F7A72C3FBF9BA6EA46FBF2" bold="true" italics="true">
<typeStatus id="65F42F68E17F57CABDB919CE8E32F67A" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="E56BDF9E67AAA2B70FCFFD776E0E9B12" name="Iraq">Iraq</collectingCountry>
</emphasis>
:
<collectingRegion id="151A315C4B0C5A836537CFB5DA25EDE0" country="Iraq" name="Dihok">Duhok</collectingRegion>
,
<location id="726FDC1CF835EF2CEB05E61863D4A729" LSID="urn:lsid:plazi:treatment:64180BEE5A8355D4A3089FA3C2E06A0F:726FDC1CF835EF2CEB05E61863D4A729" country="Iraq" latitude="37.0158" longLatPrecision="6" longitude="43.3506" name="Mt. Gara" stateProvince="Duhok">Mt. Gara</location>
[
<locationDeviation id="ED46EDC7696A5620F87D2AC8024773D9" location="Sarsing">S of Sarsing</locationDeviation>
],
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDHBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[43.350600,37.015800]}">
<geoCoordinate id="3E2D98E19051D15A86C5417144796126" degrees="37.0158" direction="north" orientation="latitude" precision="5" value="37.0158">37.0158 ° N</geoCoordinate>
,
<geoCoordinate id="AC72D885A1A115D344A2AEA5D0502537" degrees="43.3506" direction="east" orientation="longitude" precision="5" value="43.3506">43.3506 ° E</geoCoordinate>
</named-content>
,
<quantity id="D738D1FD564538A383E2B08506575980" metricMagnitude="3" metricUnit="m" metricValue="1.9119999999999997" unit="m" value="1912.0">
<elevation id="BE77B087F5CEA538596E7A413EF28646" metricMagnitude="3" metricUnit="m" metricValue="1.9119999999999997" unit="m" value="1912.0">1912 m</elevation>
</quantity>
,
<date id="A36F671E113B0E81FDBD16523B54D4F8" value="2023-05-11">
<collectingDate id="D8F10D0654934504D603E59CC0C05D64" value="2023-05-11">11. v. 2023</collectingDate>
</date>
,
<specimenCount id="D24608BB8963B5332FCFDB1C05937910" count="1" type="male">1 ♂</specimenCount>
, leg.
<collectorName id="1E79EAEB1311BC86B77B09F16E5C910E">D. Baiocchi</collectorName>
,
<named-content id="A3CE293D764A6B9EA187921617BD40AF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-natural-history-naturalis" xlink_title="Naturalis Biodiversity Center, Leiden, the Netherlands">
<collectionCode id="441111ABDDA8BCAEA9414453ED2C88B7" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</named-content>
</materialsCitation>
.
<materialsCitation id="1E4E9B958F6B49E8DB6673F47F261991" ID-GBIF-Occurrence="4900833378" collectingDate="2023-05-11" collectionCode="MSVI, RMNH, TJWC, DUMAI" collectorName="D. Baiocchi" country="Iraq" elevation="1912" latitude="37.0158" location="Iraq" longLatPrecision="6" longitude="43.3506" specimenCount="7" specimenCount-male="7" stateProvince="Duhok" typeStatus="paratype">
<emphasis id="C6BB22E9B841E69C341536FAB2CA8BE5" bold="true">
<emphasis id="8BDCB9649CC07FA390AE48599CDC7C1A" bold="true" italics="true">
<typeStatus id="F6FB95B3AFB55B2D8E4A3A89C053F06E" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="D556241DF525407DB358A7EC8B41D8A2" name="Iraq">Iraq</collectingCountry>
</emphasis>
: Same information as holotype,
<specimenCount id="6B2C7F5199C9A6C323C4DCA7166A37C7" count="7" type="male">7 ♂</specimenCount>
,
<abbrev content-type="institution" id="ABBRID0ERHBG" xlink_title="Personal collection of Marco Selis, Viterbo, Italy">
<collectionCode id="D6289B393E6A5D671719A397BA3A3FF6">MSVI</collectionCode>
</abbrev>
/
<named-content id="74B2CA9B16CC38C39D1AEB2B834E614F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-natural-history-naturalis" xlink_title="Naturalis Biodiversity Center, Leiden, the Netherlands">
<collectionCode id="3ABBFD4725242578D9FC0B4F1BDD348B" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</named-content>
/
<abbrev content-type="institution" id="ABBRID0E2HBG" xlink_title="Personal collection of Thomas J. Wood, Leiden, the Netherlands">
<collectionCode id="1969C0916FD24826CCFBA8BFD914E736">TJWC</collectionCode>
</abbrev>
/
<abbrev content-type="institution" id="ABBRID0EAIBG" xlink_title="University of Duhok, Museum of Agriculture College, Duhok, Iraq">
<collectionCode id="59BD481750119CE9AEBC6B1A2F0A89B5">DUMAI</collectionCode>
</abbrev>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFIBG" type="diagnosis">
<paragraph id="E7FC112FC5A7EA66320C13EB0AE86C53">
<heading id="84E4AC04E6EA3A8239F09D7DAEE4F2E3" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="E7E267655A78F3B27C341F2E57691719">
<taxonomicName id="FC56A65DA3030319029E8755672BFA76" authorityName="Wood &amp; Ismael &amp; Baiocchi &amp; Hamad &amp; Bapir &amp; Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis">
<emphasis id="9E10CCEFD8AF06269B9365C2AAB39183" italics="true">Andrena duhokensis</emphasis>
</taxonomicName>
can be recognised as part of the subgenus
<taxonomicName id="1D1511589782D20DC361952BE4E4BB96" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" rank="subGenus" subGenus="Aciandrena">
<emphasis id="14BC667555CA14C2E4EA771F56BD7AE3" italics="true">Aciandrena</emphasis>
Warncke, 1968
</taxonomicName>
due to the small body size, dark integument with the exception of the pale-marked clypeus (Fig.
<figureCitation id="CF59674746FE9AD5B3D8FDF35F1BC08B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 A</figureCitation>
), finely shagreened propodeal triangle (Fig.
<figureCitation id="CA414881187B2557E0E011A6ACADCDA4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 D</figureCitation>
; without lateral or basal rugae), more or less impunctate terga (Fig.
<figureCitation id="7BB791D67879B1B406FFCFC25A2E4B63" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 E</figureCitation>
), strongly antefurcal nervulus, and typical simple genital capsule (Fig.
<figureCitation id="C63F9C6870560A6446621D638C3E2CD8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 F; e</figureCitation>
. g., without completely reduced gonocoxae as in most members of the subgenus
<taxonomicName id="AB1EF55F9AA6A6DCB535B4973B95973C" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" rank="subGenus" subGenus="Graecandrena">
<emphasis id="2B986CA7DB0CD31CA37CCFDBBCE49D4B" italics="true">Graecandrena</emphasis>
Warncke, 1968
</taxonomicName>
). Identification of
<taxonomicName id="1ACB8C3B8BB06F333AD5573945AD55A1" authorityName="Warncke" authorityYear="1968" rank="subGenus" subGenus="Aciandrena">
<emphasis id="AC16AFDC9AE8D565BF26236FDEFB1401" italics="true">Aciandrena</emphasis>
</taxonomicName>
species is extremely challenging due to the large number of often cryptic species found in dry environments, their often very local distributions, and lack of good characters in the female sex. Fortunately, males are a little easier to identify through the examination of the genital capsule.
<taxonomicName id="72816D1E74B829FA7A3F29C7FCC7EB34" authorityName="Wood &amp; Ismael &amp; Baiocchi &amp; Hamad &amp; Bapir &amp; Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis">
<emphasis id="E8AE63D28EA4D3EFD7B670A6097B86FF" italics="true">Andrena duhokensis</emphasis>
</taxonomicName>
can be initially recognised due to its almost entirely yellow-marked clypeus (with the exception of two small black marks [sometimes absent] and some narrow black areas at the edges of the clypeus), placing it closest to
<taxonomicName id="6694CDECA143DFE1EE46DE59EA3D426E" authority="Morawitz, 1877" authorityName="Morawitz" authorityYear="1877" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="1AD3DDBF540D188044957CBD4335850A" italics="true">A. tenuis</emphasis>
Morawitz, 1877
</taxonomicName>
which was described from the Caucasus. However,
<taxonomicName id="5C88312CCA2267D070AEF977A78FEC50" authorityName="Wood &amp; Ismael &amp; Baiocchi &amp; Hamad &amp; Bapir &amp; Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis">
<emphasis id="C2B4944B2AA444BAFBF4E09B98C3593A" italics="true">A. duhokensis</emphasis>
</taxonomicName>
can be recognised due to the unique form of the genital capsule which has the gonocoxal teeth with their apexes strongly truncate and asymmetrical so that they diverge from each other (Fig.
<figureCitation id="7E0473549770949F39C2C997A75ADBB2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 F</figureCitation>
); in other words, they form somewhat pointed but flattened and diverging teeth. In comparative species of
<taxonomicName id="2FC47F344D0DA53F19B53D8125E5053E" authorityName="Warncke" authorityYear="1968" rank="subGenus" subGenus="Aciandrena">
<emphasis id="A6A1DF3580194954FF9B203D3782C2BA" italics="true">Aciandrena</emphasis>
</taxonomicName>
, the gonocoxal teeth are either strongly produced and truncate (e. g.,
<taxonomicName id="BD0D4C2034CA03193B521273DA49519B" authority="Wood 2022" authorityName="Wood" authorityYear="2022" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="deminuta">
<emphasis id="DE0CC1C76BBC25E5B5E311EBA5E8623D" italics="true">A. deminuta</emphasis>
<bibRefCitation id="1E7CFFC189CB164ECF3B345C3C529053" DOI="10.5852/ejt.2022.843.1947" author="Wood TJ &amp; Monfared A" journalOrPublisher="European Journal of Taxonomy" pagination="1-136" refId="B41" refString="Wood TJ, Monfared A (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843. 1947" title="A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species." volume="843" year="2022">Wood 2022</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="2BCDD8F2EAD9E1E9BA1FABB9441057F5" name="Iran">Iran</collectingCountry>
), produced into narrowly pointed or more broadly rounded teeth (e. g.,
<taxonomicName id="24AE8DE89A8FCFA65CD3BF84CE56AECC" authorityName="Morawitz" authorityYear="1877" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="E8D91B739A48D3308E93F57B746504EB" italics="true">A. tenuis</emphasis>
</taxonomicName>
from
<collectingCountry id="399405FE3D2308073185432919ABA30B" name="Turkey">Turkey</collectingCountry>
and the Caucasus,
<taxonomicName id="0DD565CDC38AF9E1A584BAC3612E3812" authority="Morawitz, 1886" authorityName="Morawitz" authorityYear="1886" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aciculata">
<emphasis id="28AFF22B15CF7261661A394CD940A938" italics="true">A. aciculata</emphasis>
Morawitz, 1886
</taxonomicName>
from Europe to the Caucasus), or reduced and truncate but never forming diverging teeth (e. g.,
<taxonomicName id="2278620F4C42CAF79DC2C604B97852C1" authority="Scheuchl &amp; Pisanty, 2016" authorityName="Scheuchl &amp; Pisanty" authorityYear="2016" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="judaea">
<emphasis id="20913C65A1DBCC9F264D47AC04E6105C" italics="true">A. judaea</emphasis>
<bibRefCitation id="4D9DFFCCA03E1A3D8FAEE16CCE304025" DOI="10.11646/zootaxa.4189.3.3" author="Pisanty G &amp; Scheuchl E &amp; Dorchin N" issue="3" journalOrPublisher="Zootaxa" pagination="485-515" refId="B21" refString="Pisanty G, Scheuchl E, Dorchin N (2016) Eight new species of Andrena Fabricius (Hymenoptera: Apoidea: Andrenidae) from Israel - a Mediterranean hotspot for wild bees. Zootaxa 4189 (3): 485 - 515. https: // doi. org / 10.11646 / zootaxa. 4189.3. 3" title="Eight new species of Andrena Fabricius (Hymenoptera: Apoidea: Andrenidae) from Israel - a Mediterranean hotspot for wild bees." volume="4189" year="2016">Scheuchl &amp; Pisanty, 2016</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="63727013C3E561D6630E47A375C913C8" name="Israel">Israel</collectingCountry>
). In this context,
<taxonomicName id="EE9B9C9EDABBBEADDDC4638C1FB368D7" authorityName="Wood &amp; Ismael &amp; Baiocchi &amp; Hamad &amp; Bapir &amp; Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis">
<emphasis id="E004B0C7AA796FD300A1BD6C29B0826C" italics="true">A. duhokensis</emphasis>
</taxonomicName>
can be separated from all similar species.
</paragraph>
<caption id="D0FA8CE8F33782DFBDEDC5D51EA331B7" ID-DOI="10.3897/zookeys.1205.120033.figure5" ID-arpha="96C67188-9C86-5C49-B2DE-977A05249501" httpUri="https://binary.pensoft.net/fig/1083013" startId="F5">
<paragraph id="834D8F7CF2A4BA8E5ED0A3C9F73EC404">
<label id="367B1D49D080EBAAE9E84A95E98281CE">Figure 5.</label>
</paragraph>
<paragraph id="FAFD3F374F1DDCAAFDC245609714D6EC">
<taxonomicName id="07D120DA8826CACA2C70C8E3673C5F4C" authorityName="Wood &amp; Ismael &amp; Baiocchi &amp; Hamad &amp; Bapir &amp; Selis" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="duhokensis" status="sp. nov." subGenus="Aciandrena">
<emphasis id="DA0248160D1579AE4DB59A28249FEFE0" italics="true">Andrena</emphasis>
(
<emphasis id="8495E7EA28C7664D5CFAB7ACFB1B4B3E" italics="true">Aciandrena</emphasis>
)
<emphasis id="D3342A1B8264AAEFB3F3EE47735078B7" italics="true">duhokensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F9AD5D2E9AC460ACAD685D22D2C8F495" rank="species">sp. nov.</taxonomicNameLabel>
male
<emphasis id="3ECDE2227EEB5677647C30FDDD9FF218" bold="true">A</emphasis>
habitus, lateral view
<emphasis id="426CC6EC2D06AD607482042C674CB0D6" bold="true">B</emphasis>
face, frontal view
<emphasis id="28CDF0125951BC041FC1F6BC48ED7BA8" bold="true">C</emphasis>
scutum, dorsal view
<emphasis id="AC2C850F4358BD6CCAF816B6AE3A2A9D" bold="true">D</emphasis>
propodeum, dorsal view
<emphasis id="BC0EE7BB6C3564BE577625EDAAC75C3F" bold="true">E</emphasis>
terga, dorsal view
<emphasis id="FC6DF7B9BF08166E32CCBB1C52099F0E" bold="true">F</emphasis>
genital capsule, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EJOBG" type="description">
<paragraph id="2CD635EB2AE1930C40255F66AFB9451B">
<heading id="E562E88700F4505C61E9CF211FA1BFE2" reason="title">Description.</heading>
</paragraph>
<paragraph id="9DC66673A52E78E99336AAA7BA160F50">
<emphasis id="31B57C859F94C5CB173FF6C81C1B7924" bold="true">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="F44579123C7D2F3786D93C231497063B">
<emphasis id="53D2D3CE4BE193277F0FA69491033C9A" bold="true">Male.</emphasis>
Body length
<quantity id="4B19AD8ACC6F12FF106F7CA5D17C9929" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
(Fig.
<figureCitation id="795C216E7784296D35264FEB3267156E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 A</figureCitation>
).
<emphasis id="2E2FCBCB35BD56F7844B045CF0249A97" bold="true" italics="true">Head</emphasis>
: Dark, 1.25 × wider than long (Fig.
<figureCitation id="069D1036022C28CB82A1F99577DCE948" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 B</figureCitation>
). Clypeus weakly domed, predominantly yellow or pale yellow marked with exception of two black dots (sometimes absent) and narrow black markings laterally; surface finely shagreened and weakly shining, irregularly punctate, punctures separated by 13 puncture diameters. Process of labrum small, rounded trapezoidal, 2 × wider than long, apical margin weakly emarginate. Gena subequal to diameter of compound eye; ocelloccipital distance equals diameter of lateral ocellus. Head covered with dirty white hairs, none equalling length of scape. Antennae basally dark, A 413 ventrally lightened by presence of grey scales; A 3 exceeding length of A 4, shorter than A 4 + 5; A 4 sub-square, shorter than long, A 5 quadrate, as long as broad.
</paragraph>
<paragraph id="8FD19EFBF074221DF2480ED42881798A">
<emphasis id="70F25E41C6098B2745B20048BB0AE4B0" bold="true" italics="true">Mesosoma</emphasis>
: Scutum and scutellum with fine granular shagreen, weakly shining, irregularly punctate, punctures separated by 14 puncture diameters (Fig.
<figureCitation id="2812B794CEF94E21DDFF5002480D5BBE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 C</figureCitation>
). Mesepisternum and dorsolateral parts of propodeum with regular but large microreticulation, dull to weakly shining; propodeal triangle large, indicated by change in surface sculpture, internal surface with fine granular microreticulation, in some parts with weakly raised network of carinae between microreticulation, surface dull (Fig.
<figureCitation id="AE26C1D8705FC8B6049744FF797D5082" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 D</figureCitation>
). Mesosoma with sparse short whitish hairs. Legs predominantly dark, apical tarsal segments slightly lightened orange, pubescence white. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma, and venation pale orange, nervulus very strongly antefurcal.
</paragraph>
<paragraph id="774B4FF9F2491536CCB9826A147BDF25">
<emphasis id="6F590F1A4BE1A0F28754936EB73B8E12" bold="true" italics="true">Metasoma</emphasis>
: Terga predominantly dark, tergal margins weakly but distinctly depressed, broadly lightened hyaline yellow-brown (Fig.
<figureCitation id="934ADD3678552A0C5C59266FC0D58122" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 E</figureCitation>
). Tergal discs finely microreticulate to shagreened, weakly shining, surface essentially impunctate, with scattered obscure punctures disappearing into underlying sculpture. Terga with sparse scattered short hairs, not forming hairbands. T 6 and T 7 with long light brown hairs overlying pseudopygidial plate of T 7. S 8 narrow, apex slightly broadened like a fish-tail, truncate, ventral surface with dense lateral fan of brown hairs. Genital capsule compact, gonocoxae produced into apical teeth, teeth strongly truncate, apexes angled and diverging from each other to form angular anteriorly projecting teeth (Fig.
<figureCitation id="652DF2F903460D81F1B4C261B03C07C8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Andrena (Aciandrena) duhokensis sp. nov. male A habitus, lateral view B face, frontal view C scutum, dorsal view D propodeum, dorsal view E terga, dorsal view F genital capsule, dorsal view." figureDoi="10.3897/zookeys.1205.120033.figure5" httpUri="https://binary.pensoft.net/fig/1083013">5 F</figureCitation>
). Gonostyli flattened and spatulate, internal margins raised, forming slight bump on inner margin basally. Penis valves moderate, occupying ½ space between gonostyli basally, strongly tapering apically.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5PBG" type="etymology">
<paragraph id="A3FE49049E56B1D5DBE3822E3DCB2759">
<heading id="142C13DCE607CEEE71E9261E3BB446F3" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="C515FE34D1BCC8B0BB7CCDDE568B3CE5">
The name is derived from the city of
<collectingRegion id="B64A90FBD251D047CE89342FA06F8E2B" country="Iraq" name="Dihok">Duhok</collectingRegion>
which gives its name to the province in which these specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDQBG" type="distribution">
<paragraph id="F88FBCC73DF2B8EFCCCE3AD4CDFBA2E0">
<heading id="C7BFEA600DD6A4AB7CE62F2D01232B1C" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="599E7DBB4E346A0691CBB0BB134C3AD3">
<collectingCountry id="BB25C866A0F8CE811118215BABB7A8FA" name="Iraq">Iraq</collectingCountry>
(Kurdistan region).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,57 +1,59 @@
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<treatment id="644DE316FF80FFA9D34FDA46FC726554" ID-DOI="http://doi.org/10.5281/zenodo.11067323" ID-Zenodo-Dep="11067323" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554" httpUri="http://treatment.plazi.org/id/644DE316FF80FFA9D34FDA46FC726554" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466">
<treatment id="644DE316FF80FFA9D34FDA46FC726554" ID-DOI="http://doi.org/10.5281/zenodo.11067323" ID-GBIF-Taxon="234412275" ID-Zenodo-Dep="11067323" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554" httpUri="http://treatment.plazi.org/id/644DE316FF80FFA9D34FDA46FC726554" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466">
<subSubSection id="A4FE018BFF80FFB7D34FDA46FD2B627F" pageId="15" pageNumber="466" type="nomenclature">
<paragraph id="EC5B5200FF80FFB7D34FDA46FE33621E" blockId="15.[123,737,545,600]" box="[123,505,545,569]" pageId="15" pageNumber="466">
<heading id="B713E56CFF80FFB7D34FDA46FE33621E" box="[123,505,545,569]" fontSize="10" level="2" pageId="15" pageNumber="466" reason="4">
@ -63,7 +65,10 @@ Andrewes
</taxonomicName>
</heading>
</paragraph>
<paragraph id="EC5B5200FF80FFB7D34FDA25FD2B627F" blockId="15.[123,737,545,600]" box="[123,737,578,600]" pageId="15" pageNumber="466">Figures 5, 6, 10, 13C, 14C, 15C, 16C, 17C, 19A-B, 21, and 23.</paragraph>
<paragraph id="EC5B5200FF80FFB7D34FDA25FD2B627F" blockId="15.[123,737,545,600]" box="[123,737,578,600]" pageId="15" pageNumber="466">
<figureCitation id="74DF4E85FF80FFB7D34FDA25FF1C627F" box="[123,214,578,600]" captionStart="FIGURE" captionStartId="5.[686,698,839,858]" captionTargetBox="[666,1158,216,826]" captionTargetId="figure-473@5.[652,1176,216,826]" captionText="FIGURE 5. Left hind tibial medial spur and tarsomere 1, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12628237" httpUri="https://zenodo.org/record/12628237/files/figure.png" pageId="15" pageNumber="466">Figures 5</figureCitation>
, 6, 10, 13C, 14C, 15C, 16C, 17C, 19A-B, 21, and 23.
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF80FFB7D34FDA0BFD84629A" pageId="15" pageNumber="466" type="reference_group">
<paragraph id="EC5B5200FF80FFB7D34FDA0BFD026287" blockId="15.[123,1174,620,701]" pageId="15" pageNumber="466">
@ -76,9 +81,7 @@ Andrewes
</treatmentCitation>
.
<typeStatus id="335FECA2FF80FFB7D178DA0BFD0762A5" box="[588,717,620,642]" pageId="15" pageNumber="466" type="holotype">HOLOTYPE</typeStatus>
, a female, in
<collectionCode id="8AF5CAC5FF80FFB7D054DA0AFC7462A5" box="[864,958,621,642]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="15" pageNumber="466" type="Museum">NHMUK</collectionCode>
. Type locality:
, a female, in NHMUK. Type locality:
<collectingCountry id="94F31290FF80FFB7D76ADA0BFB5862A5" box="[1118,1170,620,642]" name="India" pageId="15" pageNumber="466">India</collectingCountry>
,
<collectingRegion id="2E209CE2FF80FFB7D3AFDAEDFEDE6287" box="[155,276,650,672]" country="India" name="Uttarakhand" pageId="15" pageNumber="466">Uttarakhand</collectingRegion>
@ -116,16 +119,11 @@ species in the study area by the following combination of character states: Body
<quantity id="2B1CFFE5FF80FFB7D3CBDB75FE53630D" box="[255,409,786,810]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.2" metricValueMin="4.4" pageId="15" pageNumber="466" unit="mm" value="4.8" valueMax="5.2" valueMin="4.4">4.4 to 5.2 mm</quantity>
, females
<quantity id="2B1CFFE5FF80FFB7D2CBDB75FD50630D" box="[511,666,786,810]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.6" metricValueMin="4.6" pageId="15" pageNumber="466" unit="mm" value="5.1" valueMax="5.6" valueMin="4.6">4.6 to 5.6 mm</quantity>
; pronotum dark, black or piceous, with distinct greenish or bronze metallic reflection, slightly narrowed basally (ratio PWM/
<collectionCode id="8AF5CAC5FF80FFB7D083DB54FC3A636C" box="[951,1008,819,843]" pageId="15" pageNumber="466">PWB</collectionCode>
=1.13 to 1.20); elytral color pattern as in
; pronotum dark, black or piceous, with distinct greenish or bronze metallic reflection, slightly narrowed basally (ratio PWM/PWB =1.13 to 1.20); elytral color pattern as in
<figureCitation id="74DF4E85FF80FFB7D2A1DB33FE32634B" box="[405,504,852,876]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="15" pageNumber="466">Fig. 10A</figureCitation>
, with dark areas more expansive; connection between medial portion of the basal dark band and humeral portion broader, at least one-quarter length of the humeral portion; middle band with portion between anterior and posterior dark edges distinctly darker (orange-brown) than remainder of pale elytral areas; femora pale tan; pronotum widest at or anterior to middle, slightly narrowed basally (ratio PWM/
<collectionCode id="8AF5CAC5FF80FFB7D1C0DBB1FCE763C9" box="[756,813,982,1006]" pageId="15" pageNumber="466">PWB</collectionCode>
=1.13 to 1.20); elytra slightly to distinctly and obliquely truncate apically; front tarsomeres 1 to 3 without lateral expansions (
, with dark areas more expansive; connection between medial portion of the basal dark band and humeral portion broader, at least one-quarter length of the humeral portion; middle band with portion between anterior and posterior dark edges distinctly darker (orange-brown) than remainder of pale elytral areas; femora pale tan; pronotum widest at or anterior to middle, slightly narrowed basally (ratio PWM/PWB =1.13 to 1.20); elytra slightly to distinctly and obliquely truncate apically; front tarsomeres 1 to 3 without lateral expansions (
<figureCitation id="74DF4E85FF80FFB7D754DB90FF5F6417" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="15" pageNumber="466">Figs. 14</figureCitation>
<collectionCode id="8AF5CAC5FF80FFB7D3A1DC7FFF776417" box="[149,189,1048,1072]" country="USA" name="Chicago Academy of Sciences" pageId="15" pageNumber="466" type="Museum">CA</collectionCode>
, 15C); male with middle tarsomeres 1 to 4 (
CA, 15C); male with middle tarsomeres 1 to 4 (
<figureCitation id="74DF4E85FF80FFB7D191DC7FFCCD6417" box="[677,775,1048,1072]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="15" pageNumber="466">Fig. 16C</figureCitation>
) distinctly wider than in female and with pads of adhesive setae ventrally (
<figureCitation id="74DF4E85FF80FFB7D123DC5FFDB36477" box="[535,633,1080,1104]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="15" pageNumber="466">Fig. 17C</figureCitation>
@ -179,15 +177,12 @@ adults. Within the Gaoligong
range.
</paragraph>
<paragraph id="EC5B5200FF80FFA9D39FDE44FE916274" blockId="15.[123,1174,721,1726]" lastBlockId="17.[123,1174,219,1396]" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466">
<emphasis id="DE908E12FF80FFB7D39FDE44FCD6661D" bold="true" box="[171,796,1571,1594]" pageId="15" pageNumber="466">
Geographical distribution within the Gaoligong
<materialsCitation id="5C8C585DFF80FFB7D1EFDE44FC0166BB" collectingDate="2002-10-13" collectionCode="CAS" collectorName="We &amp; D. H. Kavanaugh &amp; P. E. Marek &amp; H. B. Liang &amp; D. Z. Dong" county="Fugong County" elevation="1185" location="Nu Jiang" municipality="Shangpa Township" pageId="15" pageNumber="466" specimenCount="215" specimenCount-female="66" specimenCount-male="42" stateProvince="Shan">
<collectingRegion id="2E209CE2FF80FFB7D1EFDE44FCDD661D" box="[731,791,1571,1594]" country="Myanmar" name="Shan" pageId="15" pageNumber="466">Shan</collectingRegion>
.
<emphasis id="DE908E12FF80FFB7D39FDE44FD18661D" bold="true" box="[171,722,1571,1594]" pageId="15" pageNumber="466">Geographical distribution within the Gaoligong</emphasis>
<emphasis id="DE908E12FF80FFB7D1EFDE44FCD6661D" bold="true" box="[731,796,1571,1594]" pageId="15" pageNumber="466">Shan.</emphasis>
<figureCitation id="74DF4E85FF80FFB7D011DE45FC42661D" box="[805,904,1570,1594]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="15" pageNumber="466">Fig. 10B</figureCitation>
.
<collectorName id="411137D6FF80FFB7D0A0DE44FC7D661D" box="[916,951,1571,1594]" pageId="15" pageNumber="466">We</collectorName>
examined a total of
<materialsCitation id="5C8C585DFF80FFB7D0A0DE44FCDC66BB" ID-GBIF-Occurrence="4902827319" collectingDate="2002-10-13" collectionCode="CAS" collectorName="We &amp; D. H. Kavanaugh &amp; P. E. Marek &amp; H. B. Liang &amp; D. Z. Dong" county="Fugong County" elevation="1185" location="Nu Jiang" municipality="Shangpa Township" pageId="15" pageNumber="466" specimenCount="215" specimenCount-female="66" specimenCount-male="42" stateProvince="Shan">
We examined a total of
<specimenCount id="FAE29989FF80FFB7D34FDE24FED3667C" box="[123,281,1603,1627]" count="107" pageId="15" pageNumber="466" type="generic">107 specimens</specimenCount>
(
<specimenCount id="FAE29989FF80FFB7D21DDE24FE41667C" box="[297,395,1603,1627]" count="42" pageId="15" pageNumber="466" type="male">42 males</specimenCount>
@ -218,14 +213,15 @@ and
&amp;
<collectorName id="411137D6FF80FFB7D117DEE2FD5C66BA" box="[547,662,1669,1693]" pageId="15" pageNumber="466">D.Z. Dong</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF80FFB7D023DEE3FC0166BB" ID-GBIF-Occurrence="4902827326" box="[791,971,1668,1692]" collectingDate="2002-10-13" collectionCode="CAS" collectorName="We &amp; D. H. Kavanaugh &amp; P. E. Marek &amp; H. B. Liang &amp; D. Z. Dong" county="Fugong County" elevation="1185" location="Nu Jiang" municipality="Shangpa Township" pageId="15" pageNumber="466" specimenCount="215" specimenCount-female="66" specimenCount-male="42" stateProvince="Shan">
<specimenCount id="FAE29989FF80FFB7D023DEE3FCBD66BB" box="[791,887,1668,1692]" count="1" pageId="15" pageNumber="466" type="female">1 female</specimenCount>
;
<collectionCode id="8AF5CAC5FF80FFB7D0B6DEE3FC7666BB" box="[898,956,1668,1692]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="466" type="Museum">CAS</collectionCode>
])
</materialsCitation>
</emphasis>
.
<materialsCitation id="5C8C585DFF80FFA9D0E2DEE2FB5B60D4" collectingDate="2002-10-08" collectingDateMax="2002-10-09" collectingDateMin="2002-10-08" collectionCode="CAS, IOZ" collectorName="Nu Jiang &amp; D. H. Kavanaugh &amp; P. E. Marek &amp; Liang" county="Gongshan County" elevation="1430" lastPageId="17" lastPageNumber="468" location="Dashada" municipality="Cikai Township" pageId="15" pageNumber="466" specimenCount="19" specimenCount-female="13" specimenCount-male="6" stateProvince="Shan">
<materialsCitation id="5C8C585DFF80FFA9D0E2DEE2FD0760D4" ID-GBIF-Occurrence="4902827336" collectingDate="2002-10-08" collectingDateMax="2002-10-09" collectingDateMin="2002-10-08" collectionCode="CAS, IOZ" collectorName="Nu Jiang &amp; D. H. Kavanaugh &amp; P. E. Marek &amp; Liang" county="Gongshan County" elevation="1430" lastPageId="17" lastPageNumber="468" location="Dashada" municipality="Cikai Township" pageId="15" pageNumber="466" specimenCount="19" specimenCount-female="13" specimenCount-male="6" stateProvince="Shan">
<emphasis id="DE908E12FF80FFB7D0E2DEE2FF5D6699" bold="true" italics="true" pageId="15" pageNumber="466">
<collectingCounty id="053A2A8CFF80FFB7D0E2DEE2FF5A6699" pageId="15" pageNumber="466">Gongshan County</collectingCounty>
:
@ -250,6 +246,8 @@ at
&amp; H.B.
<collectorName id="411137D6FF9EFFA9D134D8BCFDF460D4" box="[512,574,219,243]" pageId="17" pageNumber="468">Liang</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D1F9D8BCFB5B60D4" ID-GBIF-Occurrence="4902827322" box="[717,1169,219,243]" collectingDate="2002-10-08" collectingDateMax="2002-10-09" collectingDateMin="2002-10-08" collectionCode="CAS, IOZ" collectorName="Nu Jiang &amp; D. H. Kavanaugh &amp; P. E. Marek &amp; Liang" county="Gongshan County" elevation="1430" location="Dashada" municipality="Cikai Township" pageId="17" pageNumber="468" specimenCount="19" specimenCount-female="13" specimenCount-male="6" stateProvince="Shan">
<specimenCount id="FAE29989FF9EFFA9D1F9D8BCFCE060D4" box="[717,810,219,243]" count="6" pageId="17" pageNumber="468" type="male">6 males</specimenCount>
and
<specimenCount id="FAE29989FF9EFFA9D045D8BCFC3B60D4" box="[881,1009,219,243]" count="13" pageId="17" pageNumber="468" type="female">13 females</specimenCount>
@ -260,7 +258,7 @@ and
])
</materialsCitation>
;
<materialsCitation id="5C8C585DFF9EFFA9D34FD89CFB876194" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Silalong He &amp; D. H. Kavanaugh &amp; G. Tang &amp; D. Z. Dong &amp; Kongdang &amp; H. B Liang &amp; Y. Xu &amp; Pukawang &amp; N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
<materialsCitation id="5C8C585DFF9EFFA9D34FD89CFEA16114" ID-GBIF-Occurrence="4902827339" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Silalong He &amp; D. H. Kavanaugh &amp; G. Tang &amp; D. Z. Dong &amp; Kongdang &amp; H. B Liang &amp; Y. Xu &amp; Pukawang &amp; N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
<collectingCounty id="053A2A8CFF9EFFA9D34FD89CFEB96134" box="[123,371,251,275]" pageId="17" pageNumber="468">Dulongjiang Township</collectingCounty>
(
<collectingMunicipality id="0C3FC87AFF9EFFA9D2B2D89CFE086134" box="[390,450,251,275]" pageId="17" pageNumber="468">Bapo</collectingMunicipality>
@ -277,6 +275,8 @@ and
,
<collectorName id="411137D6FF9EFFA9D34FD97BFF3E6114" box="[123,244,283,307]" pageId="17" pageNumber="468">H.B. Liang</collectorName>
collector [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D25FD97CFEAD6154" ID-GBIF-Occurrence="4902827323" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Silalong He &amp; D. H. Kavanaugh &amp; G. Tang &amp; D. Z. Dong &amp; Kongdang &amp; H. B Liang &amp; Y. Xu &amp; Pukawang &amp; N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
<specimenCount id="FAE29989FF9EFFA9D25FD97CFE0A6114" box="[363,448,283,307]" count="3" pageId="17" pageNumber="468" type="male">3 males</specimenCount>
and
<specimenCount id="FAE29989FF9EFFA9D2CDD97CFDBB6114" box="[505,625,283,307]" count="11" pageId="17" pageNumber="468" type="female">11 females</specimenCount>
@ -303,6 +303,8 @@ along
&amp;
<collectorName id="411137D6FF9EFFA9D34FD93BFF266153" box="[123,236,348,372]" pageId="17" pageNumber="468">D.Z. Dong</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D253D93CFD6A61B4" ID-GBIF-Occurrence="4902827338" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Silalong He &amp; D. H. Kavanaugh &amp; G. Tang &amp; D. Z. Dong &amp; Kongdang &amp; H. B Liang &amp; Y. Xu &amp; Pukawang &amp; N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
<specimenCount id="FAE29989FF9EFFA9D253D93CFE706154" box="[359,442,347,371]" count="2" pageId="17" pageNumber="468" type="male">2 males</specimenCount>
and
<specimenCount id="FAE29989FF9EFFA9D2D8D93CFD9F6154" box="[492,597,347,371]" count="3" pageId="17" pageNumber="468" type="female">3 females</specimenCount>
@ -325,6 +327,8 @@ and
&amp;
<collectorName id="411137D6FF9EFFA9D2FDD91BFDCD61B4" box="[457,519,380,403]" pageId="17" pageNumber="468">Y. Xu</collectorName>
Y. collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D194D91CFC6E6194" ID-GBIF-Occurrence="4902827321" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Silalong He &amp; D. H. Kavanaugh &amp; G. Tang &amp; D. Z. Dong &amp; Kongdang &amp; H. B Liang &amp; Y. Xu &amp; Pukawang &amp; N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
<specimenCount id="FAE29989FF9EFFA9D194D91CFD3F61B4" box="[672,757,379,403]" count="8" pageId="17" pageNumber="468" type="male">8 males</specimenCount>
,
<specimenCount id="FAE29989FF9EFFA9D034D91CFCA061B4" box="[768,874,379,403]" count="5" pageId="17" pageNumber="468" type="female">5 females</specimenCount>
@ -344,13 +348,15 @@ Y. collectors [
&amp;
<collectorName id="411137D6FF9EFFA9D1DED9FBFCE26194" box="[746,808,412,435]" pageId="17" pageNumber="468">Y. Xu</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D090D9FCFB876194" ID-GBIF-Occurrence="4902827306" box="[932,1101,411,435]" collectingDate="2004-10-30" collectingDateMax="2021-05-30" collectingDateMin="2004-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Silalong He &amp; D. H. Kavanaugh &amp; G. Tang &amp; D. Z. Dong &amp; Kongdang &amp; H. B Liang &amp; Y. Xu &amp; Pukawang &amp; N, H. B Liang" county="Dulongjiang Township" elevation="1355" location="Mulangdang" municipality="Bapo" pageId="17" pageNumber="468" specimenCount="33" specimenCount-female="20" specimenCount-male="13" stateProvince="Shan">
<specimenCount id="FAE29989FF9EFFA9D090D9FCFBC86194" box="[932,1026,411,435]" count="1" pageId="17" pageNumber="468" type="female">1 female</specimenCount>
,
<collectionCode id="8AF5CAC5FF9EFFA9D738D9FCFBF56194" box="[1036,1087,411,435]" pageId="17" pageNumber="468">IOZ</collectionCode>
])
</materialsCitation>
.
<materialsCitation id="5C8C585DFF9EFFA9D763D9FCFD0A6234" collectingDate="2006-06-03" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="65" specimenCount-female="35" specimenCount-male="30" stateProvince="Shan">
<materialsCitation id="5C8C585DFF9EFFA9D763D9FCFF326234" ID-GBIF-Occurrence="4902827304" collectingDate="2006-06-03" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="65" specimenCount-female="35" specimenCount-male="30" stateProvince="Shan">
<emphasis id="DE908E12FF9EFFA9D763D9FCFEED61F4" bold="true" italics="true" pageId="17" pageNumber="468">
<collectingCounty id="053A2A8CFF9EFFA9D763D9FCFEEA61F4" pageId="17" pageNumber="468">Tengchong County</collectingCounty>
:
@ -379,6 +385,8 @@ just below bridge at
&amp;
<collectorName id="411137D6FF9EFFA9D717D9BBFB5F61D3" box="[1059,1173,476,500]" pageId="17" pageNumber="468">D.Z. Dong</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D3C3D99CFD0A6234" ID-GBIF-Occurrence="4902827312" box="[247,704,507,531]" collectingDate="2006-06-03" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="65" specimenCount-female="35" specimenCount-male="30" stateProvince="Shan">
<specimenCount id="FAE29989FF9EFFA9D3C3D99CFEAB6234" box="[247,353,507,531]" count="30" pageId="17" pageNumber="468" type="male">30 males</specimenCount>
and
<specimenCount id="FAE29989FF9EFFA9D290D99CFDE96234" box="[420,547,507,531]" count="35" pageId="17" pageNumber="468" type="female">35 females</specimenCount>
@ -389,14 +397,14 @@ and
])
</materialsCitation>
,
<materialsCitation id="5C8C585DFF9EFFA9D1E6D99CFEAC6214" collectorName="Longjiang Bridge" location="Longchuan Jiang" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Shan">
<materialsCitation id="5C8C585DFF9EFFA9D1E6D99CFEAC6214" ID-GBIF-Occurrence="4902827337" collectorName="Longjiang Bridge" location="Longchuan Jiang" pageId="17" pageNumber="468" specimenCount="1" stateProvince="Shan">
(
<location id="E93B04DBFF9EFFA9D1EED99CFC506234" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D1EED99CFC506234" box="[730,922,507,531]" name="Longchuan Jiang" pageId="17" pageNumber="468" stateProvince="Shan">Longchuan Jiang</location>
at
<collectorName id="411137D6FF9EFFA9D0FED99CFB5B6234" box="[970,1169,507,531]" pageId="17" pageNumber="468">Longjiang Bridge</collectorName>
, 24.89889°/98.66667°,
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D25ADA7CFE916274" collectingDate="2003-10-28" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Shi" elevation="1215" pageId="17" pageNumber="468" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Shan">
<materialsCitation id="5C8C585DFF9EFFA9D25ADA7CFBC06214" ID-GBIF-Occurrence="4902827328" box="[366,1034,539,563]" collectingDate="2003-10-28" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Shi" elevation="1215" pageId="17" pageNumber="468" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Shan">
<quantity id="2B1CFFE5FF9EFFA9D25ADA7CFE086214" box="[366,450,539,563]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="17" pageNumber="468" unit="m" value="1215.0">
<elevation id="67C9B533FF9EFFA9D25ADA7CFE086214" box="[366,450,539,563]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="17" pageNumber="468" unit="m" value="1215.0">1215 m</elevation>
</quantity>
@ -409,6 +417,8 @@ at
&amp; X.C.
<collectorName id="411137D6FF9EFFA9D053DA7CFC416214" box="[871,907,539,563]" pageId="17" pageNumber="468">Shi</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D73EDA7CFE916274" ID-GBIF-Occurrence="4902827330" collectingDate="2003-10-28" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; Shi" elevation="1215" pageId="17" pageNumber="468" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Shan">
<specimenCount id="FAE29989FF9EFFA9D73EDA7CFB996214" box="[1034,1107,539,563]" count="1" pageId="17" pageNumber="468" type="male">1 male</specimenCount>
and
<specimenCount id="FAE29989FF9EFFA9D7BCDA7CFF1B6274" count="2" pageId="17" pageNumber="468" type="female">2 females</specimenCount>
@ -440,7 +450,6 @@ Members of this species were collected from the northern to the southern parts o
(
<figureCitation id="74DF4E85FF9EFFA9D176DB7CFD5E6314" box="[578,660,795,819]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="17" pageNumber="468">Fig. 23</figureCitation>
).
<materialsCitation id="5C8C585DFF9EFFA9D19CDB7CFDD46454" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang &amp; Km &amp; Xia Zayu &amp; Y. Liu &amp; It &amp; Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
<collectingMunicipality id="0C3FC87AFF9EFFA9D19CDB7CFD1D6314" box="[680,727,795,819]" pageId="17" pageNumber="468">This</collectingMunicipality>
species has been known previously only from a few localities on the southern slope of the
<location id="E93B04DBFF9EFFA9D196DB5CFCC06374" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D196DB5CFCC06374" box="[674,778,827,851]" country="India" county="Medog County" latitude="29.3519" longLatPrecision="6" longitude="95.3417" municipality="This" name="Himalaya" pageId="17" pageNumber="468" stateProvince="Uttarakhand">Himalaya</location>
@ -448,13 +457,11 @@ in northcentral
<collectingCountry id="94F31290FF9EFFA9D08EDB5CFC3B6374" box="[954,1009,827,851]" name="India" pageId="17" pageNumber="468">India</collectingCountry>
(
<collectingRegion id="2E209CE2FF9EFFA9D734DB5CFB416374" box="[1024,1163,827,851]" country="India" name="Uttarakhand" pageId="17" pageNumber="468">Uttarakhand</collectingRegion>
).
<location id="E93B04DBFF9EFFA9D34FDB3CFF186354" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D34FDB3CFF186354" box="[123,210,859,883]" country="India" county="Medog County" latitude="29.3519" longLatPrecision="6" longitude="95.3417" municipality="This" name="Records" pageId="17" pageNumber="468" stateProvince="Uttarakhand">Records</location>
of its occurrence in the study area represent the first for
). Records of its occurrence in the study area represent the first for
<collectingCountry id="94F31290FF9EFFA9D009DB3CFCB76354" box="[829,893,859,883]" name="China" pageId="17" pageNumber="468">China</collectingCountry>
and the easternmost limit of its known distribution.
<location id="E93B04DBFF9EFFA9D2A6DB1CFE0563B4" LSID="urn:lsid:plazi:treatment:644DE316FF80FFA9D34FDA46FC726554:E93B04DBFF9EFFA9D2A6DB1CFE0563B4" box="[402,463,891,915]" country="India" county="Medog County" latitude="29.3519" longLatPrecision="6" longitude="95.3417" municipality="This" name="Other" pageId="17" pageNumber="468" stateProvince="Uttarakhand">Other</location>
new records from
<materialsCitation id="5C8C585DFF9EFFA9D2A6DB1CFC0B6394" ID-GBIF-Occurrence="4902827333" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang &amp; Km &amp; Xia Zayu &amp; Y. Liu &amp; It &amp; Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
Other new records from
<collectingCountry id="94F31290FF9EFFA9D1AFDB1CFD1163B4" box="[667,731,891,915]" name="China" pageId="17" pageNumber="468">China</collectingCountry>
are within
<collectingRegion id="2E209CE2FF9EFFA9D061DB1CFC4663B4" box="[853,908,891,915]" country="China" name="Xizang" pageId="17" pageNumber="468">Tibet</collectingRegion>
@ -481,6 +488,8 @@ bridge,
,
<collectorName id="411137D6FF9EFFA9D1E5DBFBFC806394" box="[721,842,923,947]" pageId="17" pageNumber="468">H.B. Liang</collectorName>
collector [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D0F5DBFCFF2363D4" ID-GBIF-Occurrence="4902827309" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang &amp; Km &amp; Xia Zayu &amp; Y. Liu &amp; It &amp; Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
<specimenCount id="FAE29989FF9EFFA9D0F5DBFCFBD36394" box="[961,1049,923,947]" count="5" pageId="17" pageNumber="468" type="male">5 males</specimenCount>
,
<specimenCount id="FAE29989FF9EFFA9D712DBFCFB586394" box="[1062,1170,923,947]" count="5" pageId="17" pageNumber="468" type="female">5 females</specimenCount>
@ -501,6 +510,8 @@ collector [
,
<collectorName id="411137D6FF9EFFA9D728DBDBFB5F63F4" box="[1052,1173,955,979]" pageId="17" pageNumber="468">H.B. Liang</collectorName>
collector [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D3DEDBBCFED86434" ID-GBIF-Occurrence="4902827305" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang &amp; Km &amp; Xia Zayu &amp; Y. Liu &amp; It &amp; Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
<specimenCount id="FAE29989FF9EFFA9D3DEDBBCFEFF63D4" box="[234,309,987,1011]" count="1" pageId="17" pageNumber="468" type="male">1 male</specimenCount>
,
<collectionCode id="8AF5CAC5FF9EFFA9D276DBBCFEBE63D4" box="[322,372,987,1011]" pageId="17" pageNumber="468">IOZ</collectionCode>
@ -521,12 +532,15 @@ collector [
,
<collectorName id="411137D6FF9EFFA9D74BDBBBFF6A6434" pageId="17" pageNumber="468">Y. Liu</collectorName>
collector [
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D227DB9CFDFA6434" ID-GBIF-Occurrence="4902827324" box="[275,560,1019,1043]" collectingDate="2011-07-04" collectingDateMax="2015-08-30" collectingDateMin="2011-07-04" collectionCode="IOZ" collectorName="H. B. Liang &amp; Km &amp; Xia Zayu &amp; Y. Liu &amp; It &amp; Himalayan" country="India" county="Medog County" elevation="707" latitude="29.3519" location="Himalaya" longLatPrecision="6" longitude="95.3417" municipality="This" pageId="17" pageNumber="468" specimenCount="24" specimenCount-female="9" specimenCount-male="15" stateProvince="Uttarakhand">
<specimenCount id="FAE29989FF9EFFA9D227DB9CFEA06434" box="[275,362,1019,1043]" count="9" pageId="17" pageNumber="468" type="male">9 males</specimenCount>
,
<specimenCount id="FAE29989FF9EFFA9D241DB9CFE156434" box="[373,479,1019,1043]" count="4" pageId="17" pageNumber="468" type="female">4 females</specimenCount>
,
<collectionCode id="8AF5CAC5FF9EFFA9D2DEDB9CFDD76434" box="[490,541,1019,1043]" pageId="17" pageNumber="468">IOZ</collectionCode>
]).
]).
</materialsCitation>
<collectorName id="411137D6FF9EFFA9D102DB9BFD8C6434" box="[566,582,1020,1043]" pageId="17" pageNumber="468">It</collectorName>
is likely that
<taxonomicName id="2BE42983FF9EFFA9D1EEDB9BFC8C6434" box="[730,838,1019,1043]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="elegans">
@ -548,9 +562,7 @@ occurs also in suitable habitats in the intervening region, including northern
</quantity>
along rivers draining the southern slopes of the
<collectorName id="411137D6FF9EFFA9D3F4DC3CFEFF6454" box="[192,309,1115,1139]" pageId="17" pageNumber="468">Himalayan</collectorName>
ranges in these areas
</materialsCitation>
.
ranges in these areas.
</paragraph>
<paragraph id="EC5B5200FF9EFFA9D39FDC1BFC726554" blockId="17.[123,1174,219,1396]" pageId="17" pageNumber="468">
<emphasis id="DE908E12FF9EFFA9D39FDC1BFCA664B4" bold="true" box="[171,876,1148,1172]" pageId="17" pageNumber="468">

View file

@ -0,0 +1,178 @@
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<mods:title id="4ECB8F48D4FA54FE567C38C84F857E31">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:namePart id="8320F01EBEC7587FD57310966313E933">Cueva-Dabkoski, Mollie</mods:namePart>
<mods:affiliation id="7738C1414EDCB4BB165AB56604274758">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:namePart id="A75B8E9311D2AC2395ACDFFEEA1214B8">Liang, Hongbin</mods:namePart>
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<mods:title id="5C34708046CE38529C8C11DE33279783">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="229BB1B8F782A7D56E430CDAA7996EF9">2023</mods:date>
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Genus
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<emphasis id="DE908E12FF83FFB4D134D95BFD746171" bold="true" box="[512,702,316,342]" italics="true" pageId="12" pageNumber="463">Tetragonoderus</emphasis>
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<emphasis id="DE908E12FF83FFB4D34FD90FFF226159" box="[123,232,360,382]" italics="true" pageId="12" pageNumber="463">Bembidium</emphasis>
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, in part [unjustified emendation for
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<emphasis id="DE908E12FF83FFB4D038D90FFCBE6159" box="[780,884,360,382]" italics="true" pageId="12" pageNumber="463">Bembidion</emphasis>
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<emphasis id="DE908E12FF83FFB4D34FD9C5FED8619F" box="[123,274,418,440]" italics="true" pageId="12" pageNumber="463">Tetragonoderus</emphasis>
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<emphasis id="DE908E12FF83FFB4D157D9C5FCD2619F" box="[611,792,418,440]" italics="true" pageId="12" pageNumber="463">Carabus quadrum</emphasis>
<bibRefCitation id="88752FF1FF83FFB4D016D9C5FC74619F" author="FABRICIUS, J. C." box="[802,958,418,440]" pageId="12" pageNumber="463" refId="ref16560" refString="FABRICIUS, J. C. 1792. Entomologia systematica emendate et aucta, secundum classes, ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Tome I. Pars I. Proft, Hafniae, xx + 330 pp." type="book" year="1792">Fabricius, 1792</bibRefCitation>
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);
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.
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</paragraph>
<paragraph id="EC5B5200FF83FFB4D34FD9BBFB5B61D5" blockId="12.[123,1173,360,556]" box="[123,1169,476,498]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="CCF4752EFF83FFB4D34FD9BBFB5B61D5" box="[123,1169,476,498]" pageId="12" pageNumber="463">
<treatmentCitation id="6D457411FF83FFB4D34FD9BBFE4961D5" author="JEANNEL, R." box="[123,387,476,498]" page="870" pageId="12" pageNumber="463" year="1949">
<taxonomicName id="2BE42983FF83FFB4D34FD9BBFE4961D5" authority="Jeannel, 1949: 870" authorityName="Jeannel" authorityPageNumber="870" authorityYear="1949" box="[123,387,476,498]" class="Insecta" family="Carabidae" genus="Cyclicus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="DE908E12FF83FFB4D34FD9BBFF0461D5" box="[123,206,476,498]" italics="true" pageId="12" pageNumber="463">Cyclicus</emphasis>
<bibRefCitation id="88752FF1FF83FFB4D3E0D9BBFE4961D5" author="JEANNEL, R." box="[212,387,476,498]" pageId="12" pageNumber="463" pagination="767 - 1146" refId="ref16832" refString="JEANNEL, R. 1949. Faune de l'empire Francais XI. Coleopteres Carabiques de la region Malagache (Troisieme partie). Librairie Larose, Paris, pp. 767 - 1146." type="book chapter" year="1949">Jeannel, 1949:870</bibRefCitation>
</taxonomicName>
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(
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species
<taxonomicName id="2BE42983FF83FFB4D13AD9BBFC5261D5" ID-CoL="55MY8" authority="Fairmaire, 1900" authorityName="Fairmaire" authorityYear="1900" box="[526,920,476,498]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="DE908E12FF83FFB4D13AD9BBFD3C61D5" box="[526,758,476,498]" italics="true" pageId="12" pageNumber="463">Tetragonoderus perrieri</emphasis>
<bibRefCitation id="88752FF1FF83FFB4D1C9D9BBFC5261D5" author="FAIRMAIRE, L." box="[765,920,476,498]" pageId="12" pageNumber="463" pagination="85 - 90" refId="ref16645" refString="FAIRMAIRE, L. 1900. Descriptions de coleopteres recueillis par M. H. Perrier de la Bathie a Madagascar. Bulletin de la Societe Entomologique de France 1900: 85 - 90." type="journal article" year="1900">Fairmaire, 1900</bibRefCitation>
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, by original designation).
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</paragraph>
<paragraph id="EC5B5200FF83FFB4D34FD99EFF1A620B" blockId="12.[123,1173,360,556]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="CCF4752EFF83FFB4D34FD99EFF1A620B" pageId="12" pageNumber="463">
<treatmentCitation id="6D457411FF83FFB4D34FD99EFE786228" author="JEANNEL, R." box="[123,434,505,527]" page="871" pageId="12" pageNumber="463" year="1949">
<taxonomicName id="2BE42983FF83FFB4D34FD99EFE786228" ID-CoL="8LS7T" authority="Jeannel, 1949: 871" authorityName="Jeannel" authorityPageNumber="871" authorityYear="1949" box="[123,434,505,527]" class="Insecta" family="Carabidae" genus="Metacyclicus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="DE908E12FF83FFB4D34FD99EFF306228" box="[123,250,505,527]" italics="true" pageId="12" pageNumber="463">Metacyclicus</emphasis>
<bibRefCitation id="88752FF1FF83FFB4D235D99EFE786228" author="JEANNEL, R." box="[257,434,505,527]" pageId="12" pageNumber="463" pagination="767 - 1146" refId="ref16832" refString="JEANNEL, R. 1949. Faune de l'empire Francais XI. Coleopteres Carabiques de la region Malagache (Troisieme partie). Librairie Larose, Paris, pp. 767 - 1146." type="book chapter" year="1949">Jeannel, 1949:871</bibRefCitation>
</taxonomicName>
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(
<typeStatus id="335FECA2FF83FFB4D2F5D99EFE266228" box="[449,492,505,527]" pageId="12" pageNumber="463">type</typeStatus>
species
<taxonomicName id="2BE42983FF83FFB4D175D99EFC0F6228" ID-CoL="55MXR" authority="Alluaud, 1897" authorityName="Alluaud" authorityYear="1897" box="[577,965,505,527]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="bastardi">
<emphasis id="DE908E12FF83FFB4D175D99EFCE56228" box="[577,815,505,527]" italics="true" pageId="12" pageNumber="463">Tetragonoderus bastardi</emphasis>
<bibRefCitation id="88752FF1FF83FFB4D00CD99EFC0F6228" author="ALLUAUD, C." box="[824,965,505,527]" pageId="12" pageNumber="463" pagination="371 - 376" refId="ref15689" refString="ALLUAUD, C. 1897. Coleopteres recueillis a Majunga (Madagascar) par M. Bastard. Bulletin du Museum d'Histoire Naturelle 1896: 371 - 376." type="journal article" year="1897">Alluaud, 1897</bibRefCitation>
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, by original designation).
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</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF83FFB4D39FDA26FDB1633E" pageId="12" pageNumber="463" type="diagnosis">
<paragraph id="EC5B5200FF83FFB4D39FDA26FBD262FE" blockId="12.[123,1174,576,793]" pageId="12" pageNumber="463">
<emphasis id="DE908E12FF83FFB4D39FDA26FED5627F" bold="true" box="[171,287,577,600]" pageId="12" pageNumber="463">Diagnosis.</emphasis>
Adults of
<taxonomicName id="2BE42983FF83FFB4D2A7DA27FDF1627F" authorityName="Dejean" authorityYear="1829" box="[403,571,576,600]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="DE908E12FF83FFB4D2A7DA27FDF1627F" box="[403,571,576,600]" italics="true" pageId="12" pageNumber="463">Tetragonoderus</emphasis>
</taxonomicName>
can be distinguished from those of
<taxonomicName id="2BE42983FF83FFB4D0F4DA27FB5F627F" box="[960,1173,576,600]" pageId="12" pageNumber="463">
<emphasis id="DE908E12FF83FFB4D0F4DA27FB8B627F" box="[960,1089,576,600]" italics="true" pageId="12" pageNumber="463">Cyclosomus</emphasis>
species
</taxonomicName>
in the study area by the following combination of character states: Body size smaller, BL = less than 6.0 mm; pronotum (
<figureCitation id="74DF4E85FF83FFB4D2BFDAE7FE2B62BF" box="[395,481,640,664]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="12" pageNumber="463">Figs. 13</figureCitation>
B-F) narrower, widest at or anterior to middle; elytra slightly to distinctly and obliquely truncate apically, elytral color pattern varied; front tarsomeres 1 to 3 without distinct lateral expansions (
<figureCitation id="74DF4E85FF83FFB4D2FEDAA6FDEA62FE" box="[458,544,705,729]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="12" pageNumber="463">Figs. 14</figureCitation>
B-F, 15B-F), broader in males than in females.
</paragraph>
<paragraph id="EC5B5200FF83FFB4D39FDA85FDB1633E" blockId="12.[123,1174,576,793]" pageId="12" pageNumber="463">
<emphasis id="DE908E12FF83FFB4D39FDA85FED162DE" bold="true" box="[171,283,738,761]" pageId="12" pageNumber="463">Diversity:</emphasis>
At present, 16 species are recognized in the Asian fauna, and here we record four from the study area, including one new species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="61055D3500BC7C591CB917D5FEF80A4B">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<taxonomicName id="2BE42983FF83FFB4D34FDB25FE2C637E" ID-CoL="55MYV" authority="Dejean" authorityName="Dejean" authorityYear="1829" box="[123,486,834,858]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">
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<emphasis id="DE908E12FF83FFB4D34FDB25FE44637D" bold="true" box="[123,398,834,858]" italics="true" pageId="12" pageNumber="463">Tetragonoderus arcuatus</emphasis>
Dejean
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<figureCitation id="74DF4E85FF83FFB4D34FDB05FF1D635F" box="[123,215,866,888]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="12" pageNumber="463">Figures 9</figureCitation>
,
<figureCitation id="74DF4E85FF83FFB4D3D6DB05FEC1635F" box="[226,267,866,888]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="12" pageNumber="463">13B</figureCitation>
,
<figureCitation id="74DF4E85FF83FFB4D222DB05FE8B635F" box="[278,321,866,888]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="12" pageNumber="463">14B</figureCitation>
,
<figureCitation id="74DF4E85FF83FFB4D27FDB05FEBF635F" box="[331,373,866,888]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="12" pageNumber="463">15B</figureCitation>
,
<figureCitation id="74DF4E85FF83FFB4D2B4DB05FE60635F" box="[384,426,866,888]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="12" pageNumber="463">16B</figureCitation>
,
<figureCitation id="74DF4E85FF83FFB4D280DB05FE14635F" box="[436,478,866,888]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="12" pageNumber="463">17B</figureCitation>
,
<figureCitation id="74DF4E85FF83FFB4D2DDDB05FDCA635F" box="[489,512,866,888]" captionStart="FIGURE" captionStartId="35.[155,167,1036,1055]" captionTargetBox="[130,1144,252,995]" captionTargetId="figure-68@35.[120,1176,216,1033]" captionTargetPageId="35" captionText="FIGURE 18. Male genitalia, Tetragonoderus spp. A, B. T. arcuatus Dejean (Nepalgunj, Banke District, Nepal); C, D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). A, C. Dorsal aspect; B, D. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067329" httpUri="https://zenodo.org/record/11067329/files/figure.png" pageId="12" pageNumber="463">18</figureCitation>
A-B, 22, and 23.
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, here designated, a female, deposited in MNHN and labeled: “
<collectingCountry id="94F31290FF83FFB4D22ADBCEFEAF6398" box="[286,357,937,959]" name="Egypt" pageId="12" pageNumber="463">Egypte</collectingCountry>
” [handwritten]/ “TYPE arcuatus” [red label]/
<taxonomicName id="2BE42983FF83FFB4D02BDBCEFC096398" ID-CoL="55MYV" authority="Dejean" authorityName="Dejean" authorityYear="1829" box="[799,963,937,959]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">“arcuatus Dejean</taxonomicName>
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Coll. Dejean” [handwritten, white label with black border]/ “
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<taxonomicName id="2BE42983FF83FFB4D027DBA1FB5F63FB" ID-CoL="55MYV" authority="Dejean 1829" authorityName="Dejean" authorityYear="1829" box="[787,1173,966,988]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF83FFB4D027DBA1FBC363FB" box="[787,1033,966,988]" italics="true" pageId="12" pageNumber="463">Tetragonoderus arcuatus</emphasis>
<bibRefCitation id="88752FF1FF83FFB4D721DBA1FB5F63FB" author="DEJEAN, P. F. M. A." box="[1045,1173,966,988]" pageId="12" pageNumber="463" refId="ref16379" refString="DEJEAN, P. F. M. A. 1829. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean 1829</bibRefCitation>
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design. by D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [red label]. Type locality:
<collectingCountry id="94F31290FF83FFB4D0D5DB83FBD063DE" box="[993,1050,996,1017]" name="Egypt" pageId="12" pageNumber="463">Egypt</collectingCountry>
[but Dejean viewed this record as uncertain].
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;
<treatmentCitation id="6D457411FF83FFB4D077DC67FBFA6431" author="ASSMANN, T. &amp; E. BOUTAUD &amp; J. BUSE &amp; V. CHIKATUNOV &amp; C. DREES &amp; A. - L. - L. FRIEDMAN &amp; W. HARDTLE &amp; K. HOMBURG &amp; T. MARCUS &amp; I. RENAN &amp; D. W. WRASE" box="[835,1072,1024,1046]" page="57" pageId="12" pageNumber="463" year="2015">
<bibRefCitation id="88752FF1FF83FFB4D077DC67FBFA6431" author="ASSMANN, T. &amp; E. BOUTAUD &amp; J. BUSE &amp; V. CHIKATUNOV &amp; C. DREES &amp; A. - L. - L. FRIEDMAN &amp; W. HARDTLE &amp; K. HOMBURG &amp; T. MARCUS &amp; I. RENAN &amp; D. W. WRASE" box="[835,1072,1024,1046]" pageId="12" pageNumber="463" pagination="49 - 69" refId="ref15907" refString="ASSMANN, T., E. BOUTAUD, J. BUSE, V. CHIKATUNOV, C. DREES, A. - L. - L. FRIEDMAN, W. HARDTLE, K. HOMBURG, T. MARCUS, I. RENAN, AND D. W. WRASE. 2015. The ground beetle tribe Cyclosomini s. l. in Israel (Coleoptera, Carabidae). Spixiana 38: 49 - 69." type="journal article" year="2015">Assmann et al., 2015:57</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6D457411FF83FFB4D70FDC66FEC36414" author="BOUSQUET, Y." page="498" pageId="12" pageNumber="463" year="2017">
<bibRefCitation id="88752FF1FF83FFB4D70FDC66FEC36414" author="BOUSQUET, Y." pageId="12" pageNumber="463" refId="ref16205" refString="BOUSQUET, Y. 2017. Tribe Cyclosomini. Pages 498 - 499 in Lobl, I., and D. Lobl (editors) Catalogue of Palaearctic Coleoptera. Volume 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden, xxxiv + 1143 pp." type="book" year="2017">Bousquet (2017:498)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EC5B5200FF83FFB4D34FDC5DFC92644A" blockId="12.[123,1174,908,1133]" pageId="12" pageNumber="463">
<treatmentCitationGroup id="CCF4752EFF83FFB4D34FDC5DFC92644A" pageId="12" pageNumber="463">
<treatmentCitation id="6D457411FF83FFB4D34FDC5DFDC06477" author="BATES, H. W." box="[123,522,1082,1104]" page="338" pageId="12" pageNumber="463" year="1891">
<taxonomicName id="2BE42983FF83FFB4D34FDC5DFDC06477" ID-CoL="55N34" authority="Bates, 1891: 338" authorityName="Bates" authorityPageNumber="338" authorityYear="1891" box="[123,522,1082,1104]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="DE908E12FF83FFB4D34FDC5DFEA96477" box="[123,355,1082,1104]" italics="true" pageId="12" pageNumber="463">Tetragonoderus cardoni</emphasis>
<bibRefCitation id="88752FF1FF83FFB4D25EDC5CFDC06477" author="BATES, H. W." box="[362,522,1082,1104]" pageId="12" pageNumber="463" pagination="324 - 340" refId="ref16102" refString="BATES, H. W. 1891. List of the Carabidae (ord. Coleoptera) obtained by Pere Cardon in Chota-Nagpore. Annales de la Societe entomologique de Belgique 35: 324 - 340." type="journal article" year="1891">Bates, 1891:338</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="335FECA2FF83FFB4D120DC5DFD5F6477" box="[532,661,1082,1104]" pageId="12" pageNumber="463" type="holotype">HOLOTYPE</typeStatus>
, a female, deposited in MNHN. Type locality:
<collectingCountry id="94F31290FF83FFB4D76ADC5DFB586477" box="[1118,1170,1082,1104]" name="India" pageId="12" pageNumber="463">India</collectingCountry>
,
<collectingRegion id="2E209CE2FF83FFB4D3AFDC30FECA644A" box="[155,256,1111,1133]" country="India" name="Jharkhand" pageId="12" pageNumber="463">Jharkhand</collectingRegion>
, Konbir.
<treatmentCitation id="6D457411FF83FFB4D26EDC3FFE3E644A" author="BATES, H. W." box="[346,500,1111,1133]" page="416" pageId="12" pageNumber="463" year="1892">
<bibRefCitation id="88752FF1FF83FFB4D26EDC3FFE3E644A" author="BATES, H. W." box="[346,500,1111,1133]" pageId="12" pageNumber="463" pagination="267 - 428" refId="ref16139" refString="BATES, H. W. 1892. Viaggio di Leonardo Fea in Birmania e regioni vicine. Annali del Museuo Civico di Storia Naturale di Genova (Serie 2 a) 12: 267 - 428." type="journal article" year="1892">Bates 1892:416</bibRefCitation>
</treatmentCitation>
. Synonymy by
<treatmentCitation id="6D457411FF83FFB4D1BDDC30FC9E644A" author="ANDREWES, H. E." box="[649,852,1111,1133]" page="150" pageId="12" pageNumber="463" year="1921">
<bibRefCitation id="88752FF1FF83FFB4D1BDDC30FC9E644A" author="ANDREWES, H. E." box="[649,852,1111,1133]" pageId="12" pageNumber="463" pagination="145 - 195" refId="ref15718" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes, 1921:150</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF83FFB6D39FDCE6FD156252" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463" type="materials_examined">
<paragraph id="EC5B5200FF83FFB4D39FDCE6FC47655D" blockId="12.[123,1174,1152,1726]" pageId="12" pageNumber="463">
<emphasis id="DE908E12FF83FFB4D39FDCE6FDFD64BF" bold="true" box="[171,567,1153,1176]" pageId="12" pageNumber="463">Notes on nomenclature and types.</emphasis>
<materialsCitation id="5C8C585DFF83FFB4D175DCE6FC43655D" ID-GBIF-Occurrence="4902827334" collectionCode="MNHN, ZMHB" country="Egypt" county="Recommendation" location="Bernd Jaeger" municipality="International Code of Zoological Nomenclature" pageId="12" pageNumber="463" specimenCount="5" specimenCount-male="1" typeStatus="lectotype">
In his original description,
<bibRefCitation id="88752FF1FF83FFB4D059DCE7FBC064BF" author="DEJEAN, P. F. M. A." box="[877,1034,1152,1176]" pageId="12" pageNumber="463" refId="ref16379" refString="DEJEAN, P. F. M. A. 1829. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean (1829)</bibRefCitation>
was unclear with regard to how many specimens he had examined (Thierry Deuve, personal communication). He noted that he had received material from Klug and that he had not seen a male specimen. In addition to a single female specimen in
<collectionCode id="8AF5CAC5FF83FFB4D11ADC85FD4864DE" box="[558,642,1250,1273]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="463">MNHN</collectionCode>
, there are
<specimenCount id="FAE29989FF83FFB4D1C1DC86FC5264DE" box="[757,920,1249,1273]" count="4" pageId="12" pageNumber="463" type="generic">four specimens</specimenCount>
from
<collectingCountry id="94F31290FF83FFB4D0E9DC85FBD764DD" box="[989,1053,1250,1274]" name="Egypt" pageId="12" pageNumber="463">Egypt</collectingCountry>
in
<collectionCode id="8AF5CAC5FF83FFB4D776DC85FB5864DE" box="[1090,1170,1250,1273]" pageId="12" pageNumber="463">ZMHB</collectionCode>
, which were probably part of Klugs original series and include
<specimenCount id="FAE29989FF83FFB4D018DD65FC45653E" box="[812,911,1281,1305]" count="1" pageId="12" pageNumber="463" type="male">one male</specimenCount>
(
<location id="E93B04DBFF83FFB4D0AADD66FBE5653D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF83FFB4D0AADD66FBE5653D" box="[926,1071,1281,1306]" country="Egypt" county="Recommendation" municipality="International Code of Zoological Nomenclature" name="Bernd Jaeger" pageId="12" pageNumber="463">Bernd Jaeger</location>
, personal communication).
<location id="E93B04DBFF83FFB4D208DD44FE86651D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF83FFB4D208DD44FE86651D" box="[316,332,1315,1338]" country="Egypt" county="Recommendation" municipality="International Code of Zoological Nomenclature" name="It" pageId="12" pageNumber="463">It</location>
seems most likely that
<location id="E93B04DBFF83FFB4D156DD45FD64651D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF83FFB4D156DD45FD64651D" box="[610,686,1314,1338]" country="Egypt" county="Recommendation" municipality="International Code of Zoological Nomenclature" name="Dejean" pageId="12" pageNumber="463">Dejean</location>
saw only the single female specimen now deposited in
<collectionCode id="8AF5CAC5FF83FFB4D232DD24FE90657D" box="[262,346,1347,1370]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="463">MNHN</collectionCode>
, but we cannot be sure.
<location id="E93B04DBFF83FFB4D156DD25FD3C657D" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF83FFB4D156DD25FD3C657D" box="[610,758,1346,1370]" country="Egypt" county="Recommendation" municipality="International Code of Zoological Nomenclature" name="Consequently" pageId="12" pageNumber="463">Consequently</location>
, we designate that female as
<typeStatus id="335FECA2FF83FFB4D700DD25FB5C657D" box="[1076,1174,1346,1370]" pageId="12" pageNumber="463" type="lectotype">lectotype</typeStatus>
(
<collectingMunicipality id="0C3FC87AFF83FFB4D3B6DD05FD4F655D" box="[130,645,1378,1402]" pageId="12" pageNumber="463">International Code of Zoological Nomenclature</collectingMunicipality>
,
<collectingCounty id="053A2A8CFF83FFB4D1A4DD05FC86655D" box="[656,844,1378,1402]" pageId="12" pageNumber="463">Recommendation</collectingCounty>
73F)
</materialsCitation>
.
</paragraph>
<paragraph id="EC5B5200FF83FFB6D39FDDE4FDFD61B3" blockId="12.[123,1174,1152,1726]" lastBlockId="14.[123,1174,219,1725]" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463">
When he proposed the synonymy of
<taxonomicName id="2BE42983FF83FFB4D10DDDE3FD2265BD" authority="Bates" authorityName="Bates" authorityYear="1891" box="[569,744,1411,1435]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="DE908E12FF83FFB4D10DDDE3FD6C65BC" box="[569,678,1411,1435]" italics="true" pageId="12" pageNumber="463">T. cardoni</emphasis>
Bates
</taxonomicName>
with
<taxonomicName id="2BE42983FF83FFB4D013DDE3FB5F65BC" authority=", Andrewes (1921: 150)" authorityName="Andrewes" authorityPageNumber="150" authorityYear="1921" box="[807,1173,1411,1435]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF83FFB4D013DDE3FC5465BC" box="[807,926,1412,1435]" italics="true" pageId="12" pageNumber="463">T. arcuatus</emphasis>
,
<bibRefCitation id="88752FF1FF83FFB4D09FDDE4FB5F65BC" author="ANDREWES, H. E." box="[939,1173,1411,1435]" pageId="12" pageNumber="463" pagination="145 - 195" refId="ref15718" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:150)</bibRefCitation>
</taxonomicName>
also noted the pronounced variation in the degree of development of the sericeous sheen seen among specimens of this species. Specimens from the study area and adjacent regions appear to have the most markedly sericeous elytra while those from areas to the west, including the
<typeStatus id="335FECA2FF83FFB4D75CDD83FB5C65DB" box="[1128,1174,1508,1532]" pageId="12" pageNumber="463">type</typeStatus>
locality (
<collectingCountry id="94F31290FF83FFB4D3EDDE63FED4663B" box="[217,286,1540,1564]" name="Egypt" pageId="12" pageNumber="463">Egypt</collectingCountry>
), have the least sericeous elytra. There is also conspicuous variation in the distinctness and extent of the subapical pale band of spots. In the single specimen from the study area, and also in those from adjacent parts of the eastern end of the range of the species (e.g., Assam,
<collectingCountry id="94F31290FF83FFB4D77ADE22FF6E665B" name="Myanmar" pageId="12" pageNumber="463">Myanmar</collectingCountry>
and northern
<collectingCountry id="94F31290FF83FFB4D20DDE03FE56665B" box="[313,412,1636,1660]" name="Thailand" pageId="12" pageNumber="463">Thailand</collectingCountry>
), the subapical pale band of spots is faint to very faint, almost invisible in some specimens, and apparent only on intervals 7 and 8 or extremely faintly visible also on more mediad intervals in some specimens. In contrast, specimens from the western part of the range of the species (i.e., North Africa, the Middle East and Central Asia) have the subapical band much more distinct and extended from interval 2 to interval 8. The female
<typeStatus id="335FECA2FF81FFB6D068D89BFC406133" box="[860,906,252,276]" pageId="14" pageNumber="465">type</typeStatus>
of
<taxonomicName id="2BE42983FF81FFB6D09AD89BFBD16134" authorityName="Bates" authorityYear="1891" box="[942,1051,251,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="DE908E12FF81FFB6D09AD89BFBD16134" box="[942,1051,251,275]" italics="true" pageId="14" pageNumber="465">T. cardoni</emphasis>
</taxonomicName>
is intermediate between specimens from the western part of the species range and those from the study area and adjacent regions in both of these features. Because of this pronounced variation, features noted in the key above and in the diagnosis presented below are applicable specifically for specimens from the study area and adjacent regions.
</paragraph>
<caption id="B89B0288FF82FFB5D3AFDE5FFE376659" ID-DOI="http://doi.org/10.5281/zenodo.11067303" ID-Zenodo-Dep="11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="13" pageNumber="464" startId="13.[155,167,1592,1611]" targetBox="[128,1175,226,1577]" targetPageId="13" targetType="figure">
<paragraph id="EC5B5200FF82FFB5D3AFDE5FFE376659" blockId="13.[123,1174,1592,1663]" pageId="13" pageNumber="464">
<smallCapsWord id="EABDC4DCFF82FFB5D3AFDE5FFF12666D" baselines="1606,1606" box="[155,216,1592,1611]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="13" pageNumber="464">FIGURE</smallCapsWord>
9.
<taxonomicName id="2BE42983FF82FFB5D3C7DE5FFDD1666C" authority="(Dejean)" baseAuthorityName="Dejean" box="[243,539,1592,1611]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF82FFB5D3C7DE5FFE03666C" box="[243,457,1592,1611]" italics="true" pageId="13" pageNumber="464">Tetragonoderus arcuatus</emphasis>
(Dejean)
</taxonomicName>
. A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<paragraph id="EC5B5200FF81FFB6D39FD9FBFD156252" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
Adults of several additional Asian taxa presently treated as distinct species are very similar to those of
<taxonomicName id="2BE42983FF81FFB6D3EAD9DAFE9D61F3" box="[222,343,445,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF81FFB6D3EAD9DAFE9D61F3" box="[222,343,445,468]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
</taxonomicName>
. These include
<taxonomicName id="2BE42983FF81FFB6D13FD9DBFC1561F3" authority="Jedlicka, 1964" authorityName="Jedlicka" authorityYear="1964" box="[523,991,444,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="DE908E12FF81FFB6D13FD9DBFCFF61F3" box="[523,821,444,468]" italics="true" pageId="14" pageNumber="465">Tetragonoderus assamensis</emphasis>
<bibRefCitation id="88752FF1FF81FFB6D00BD9DBFC1561F3" author="JEDLICKA, A." box="[831,991,444,468]" pageId="14" pageNumber="465" pagination="305 - 318" refId="ref16866" refString="JEDLICKA, A. 1964. Neue Carabiden aus Indien (Coleoptera-Carabidae). Entomologische Arbeiten aus dem Museum G. Frey 15: 305 - 318." type="journal article" year="1964">Jedlička, 1964</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2BE42983FF81FFB6D0D9D9DBFE4C61D3" authority="Jedlicka, 1964" authorityName="Jedlicka" authorityYear="1964" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="cinchona">
<emphasis id="DE908E12FF81FFB6D0D9D9DBFF1761D3" italics="true" pageId="14" pageNumber="465">Tetragonoderus cinchona</emphasis>
<bibRefCitation id="88752FF1FF81FFB6D3D2D9BBFE4C61D3" author="JEDLICKA, A." box="[230,390,476,500]" pageId="14" pageNumber="465" pagination="305 - 318" refId="ref16866" refString="JEDLICKA, A. 1964. Neue Carabiden aus Indien (Coleoptera-Carabidae). Entomologische Arbeiten aus dem Museum G. Frey 15: 305 - 318." type="journal article" year="1964">Jedlička, 1964</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2BE42983FF81FFB6D2A0D9BBFC9761D3" authority="Solsky, 1874" authorityName="Solsky" authorityYear="1874" box="[404,861,476,500]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="DE908E12FF81FFB6D2A0D9BBFD0E61D3" box="[404,708,476,500]" italics="true" pageId="14" pageNumber="465">Tetragonoderus intermedius</emphasis>
<bibRefCitation id="88752FF1FF81FFB6D1F9D9BBFC9761D3" author="SOLSKY, S. M." box="[717,861,476,500]" pageId="14" pageNumber="465" refId="ref17856" refString="SOLSKY, S. M. 1874. Zhestkokrylye (Coleoptera). In: Fedchenko, A. P. Puteshestvie v Turkestan. Izvestiya Imperatorskogo Obshchestva Lyubitelei Estestvoznaniya, Anthropologii I Etnografii (5) 11: iv + 222 pp. + 1 pl." type="book" year="1874">Solsky, 1874</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="2BE42983FF81FFB6D0AFD9BBFED06232" authority="Jedlicka, 1966" authorityName="Jedlicka" authorityYear="1966" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="nakaoi">
<emphasis id="DE908E12FF81FFB6D0AFD9BBFB5C61D3" box="[923,1174,476,500]" italics="true" pageId="14" pageNumber="465">Tetragonoderus nakaoi</emphasis>
<bibRefCitation id="88752FF1FF81FFB6D34FD99AFED06232" author="JEDLICKA, A." box="[123,282,509,533]" pageId="14" pageNumber="465" pagination="11 - 16" refId="ref16894" refString="JEDLICKA, A. 1966. 2. Family Carabidae. Neue Carabiden aus Vietnam und Thailand (Coleoptera-Carabidae). Memoirs of the Faculty of Education. Kagawa University (Part II) 140: 11 - 16" type="journal article" year="1966">Jedlička, 1966</bibRefCitation>
</taxonomicName>
. The taxonomic status of these names is currently under review, but none of them have been recorded from the study area and all are more recently described than
<taxonomicName id="2BE42983FF81FFB6D0EFDA79FB986212" box="[987,1106,542,565]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF81FFB6D0EFDA79FB986212" box="[987,1106,542,565]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
</taxonomicName>
. Consequently, none of them would challenge
<taxonomicName id="2BE42983FF81FFB6D171DA59FD776272" box="[581,701,574,597]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF81FFB6D171DA59FD776272" box="[581,701,574,597]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
</taxonomicName>
as the valid name of this species if one or more are synonymized with the latter name in the future.
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF81FFB7D39FDA19FC2A61DF" lastPageId="15" lastPageNumber="466" pageId="14" pageNumber="465" type="diagnosis">
<paragraph id="EC5B5200FF81FFB6D39FDA19FC29641F" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
<emphasis id="DE908E12FF81FFB6D39FDA19FED562B2" bold="true" box="[171,287,638,661]" pageId="14" pageNumber="465">Diagnosis.</emphasis>
Adults of
<taxonomicName id="2BE42983FF81FFB6D2BADA18FDC962B1" box="[398,515,639,662]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF81FFB6D2BADA18FDC962B1" box="[398,515,639,662]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
</taxonomicName>
can be distinguished from those of other cyclosomine species in the study area by the following combination of character states: Body size medium for genus, BL males =
<quantity id="2B1CFFE5FF81FFB6D3CADAD9FE5C62F1" box="[254,406,702,726]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.1" metricValueMin="4.9" pageId="14" pageNumber="465" unit="mm" value="5.0" valueMax="5.1" valueMin="4.9">4.9 to 5.1 mm</quantity>
, females = 5.0 to
<quantity id="2B1CFFE5FF81FFB6D160DAD9FD6062F1" box="[596,682,702,726]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="14" pageNumber="465" unit="mm" value="5.7">5.7 mm</quantity>
; forebody uniformly dark, elytra dark without pale markings in basal half, in the study area with faint pale band of spots in apical half (
<figureCitation id="74DF4E85FF81FFB6D70FDAB9FB5B62D1" box="[1083,1169,734,758]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="14" pageNumber="465">Fig. 9A</figureCitation>
, mainly visible on intervals 7 and 8, femora dark black or piceous, tibiae paler; dorsal surfaces dull, with slight to distinct bronze/copper metallic reflection, elytra sericeous (silky) and mottled, especially in posterior half, with elytral microsculpture comprised of a complex pattern of transverse, longitudinal, and oblique patches of elongate meshes, most distorted near discal and umbilicate setal pore punctures; pronotum widest at or anterior to middle; elytra slightly to distinctly and obliquely truncate apically, posterior discal setiferous puncture on interval 3 inserted distinctly anterior to level of pale subapical markings; front tarsomeres 1 to 3 without lateral expansions (
<figureCitation id="74DF4E85FF81FFB6D3B7DB87FF2563DF" box="[131,239,992,1016]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="14" pageNumber="465">Figs. 14B</figureCitation>
,
<figureCitation id="74DF4E85FF81FFB6D3CEDB87FEE363DF" box="[250,297,992,1016]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="14" pageNumber="465">15B</figureCitation>
), middle tarsi of males (
<figureCitation id="74DF4E85FF81FFB6D104DB87FD5A63DF" box="[560,656,992,1016]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="14" pageNumber="465">Fig. 16B</figureCitation>
) with tarsomeres 1 to 4 broader than in females and with pads of adhesive setae ventrally (
<figureCitation id="74DF4E85FF81FFB6D170DC67FD6C643F" box="[580,678,1024,1048]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="14" pageNumber="465">Fig. 17B</figureCitation>
); median lobe of male genitalia (
<figureCitation id="74DF4E85FF81FFB6D73DDC67FB47643F" box="[1033,1165,1024,1048]" captionStart="FIGURE" captionStartId="35.[155,167,1036,1055]" captionTargetBox="[130,1144,252,995]" captionTargetId="figure-68@35.[120,1176,216,1033]" captionTargetPageId="35" captionText="FIGURE 18. Male genitalia, Tetragonoderus spp. A, B. T. arcuatus Dejean (Nepalgunj, Banke District, Nepal); C, D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). A, C. Dorsal aspect; B, D. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067329" httpUri="https://zenodo.org/record/11067329/files/figure.png" pageId="14" pageNumber="465">Figs. 18A,B</figureCitation>
) long and slender, with apical lamella also elongate, slender and rounded apically.
</paragraph>
<paragraph id="EC5B5200FF81FFB6D39FDC26FB5C649E" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
In the study area, specimens of
<taxonomicName id="2BE42983FF81FFB6D2C9DC25FDB9647E" box="[509,627,1090,1113]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF81FFB6D2C9DC25FDB9647E" box="[509,627,1090,1113]" italics="true" pageId="14" pageNumber="465">T. arcuatus</emphasis>
</taxonomicName>
(
<figureCitation id="74DF4E85FF81FFB6D1B6DC26FD1C647E" box="[642,726,1089,1113]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="14" pageNumber="465">Fig. 9A</figureCitation>
) are similar only to those of
<taxonomicName id="2BE42983FF81FFB6D73EDC25FB5C647E" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[1034,1174,1089,1113]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="parviculus">
<emphasis id="DE908E12FF81FFB6D73EDC25FB5C647E" box="[1034,1174,1089,1113]" italics="true" pageId="14" pageNumber="465">T. parviculus</emphasis>
</taxonomicName>
(
<figureCitation id="74DF4E85FF81FFB6D3B7DC06FF2C645E" box="[131,230,1121,1145]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="14" pageNumber="465">Fig. 11A</figureCitation>
) but differ from them in having larger body size, elytra with less distinct and more restricted subapical band and distinctly sericeous sheen, and male genitalia with the median lobe more slender and with a longer and narrower apical lamella (refer to key for additional differences).
</paragraph>
<paragraph id="EC5B5200FF81FFB6D39FDCA5FD94651D" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
<emphasis id="DE908E12FF81FFB6D39FDCA5FE5064FE" bold="true" box="[171,410,1218,1241]" pageId="14" pageNumber="465">Habitat distribution.</emphasis>
The lone specimen collected in the study area was found in daytime under a stone on an open beach of the Longchuan River on a mix of sandy and rocky substrate and at an elevation of
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.
<bibRefCitation id="88752FF1FF81FFB6D295DD65FD58653D" author="ASSMANN, T. &amp; E. BOUTAUD &amp; J. BUSE &amp; V. CHIKATUNOV &amp; C. DREES &amp; A. - L. - L. FRIEDMAN &amp; W. HARDTLE &amp; K. HOMBURG &amp; T. MARCUS &amp; I. RENAN &amp; D. W. WRASE" box="[417,658,1282,1306]" pageId="14" pageNumber="465" pagination="49 - 69" refId="ref15907" refString="ASSMANN, T., E. BOUTAUD, J. BUSE, V. CHIKATUNOV, C. DREES, A. - L. - L. FRIEDMAN, W. HARDTLE, K. HOMBURG, T. MARCUS, I. RENAN, AND D. W. WRASE. 2015. The ground beetle tribe Cyclosomini s. l. in Israel (Coleoptera, Carabidae). Spixiana 38: 49 - 69." type="journal article" year="2015">Assmann et al. (2015)</bibRefCitation>
found members of this species “on heavy soils close to water in semi-desert areas” in
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.
</paragraph>
<paragraph id="EC5B5200FF81FFB6D39FDD24FF79659C" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">
<emphasis id="DE908E12FF81FFB6D39FDD24FD16657D" bold="true" box="[171,732,1347,1370]" pageId="14" pageNumber="465">Geographical distribution within the Gaoligong</emphasis>
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<emphasis id="DE908E12FF81FFB6D1DCDD24FCE3657D" bold="true" box="[744,809,1347,1370]" pageId="14" pageNumber="465">
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.
</emphasis>
<figureCitation id="74DF4E85FF81FFB6D000DD25FC46657D" box="[820,908,1346,1370]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="14" pageNumber="465">Fig. 9B</figureCitation>
.
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examined a single female specimen from the following locality:
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<collectingCounty id="053A2A8CFF81FFB6D1A3DD04FCB9655C" box="[663,883,1379,1403]" pageId="14" pageNumber="465">Tengchong County</collectingCounty>
:
</emphasis>
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(
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,
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,
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,
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,
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&amp;
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collectors)
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.
</paragraph>
<paragraph id="EC5B5200FF81FFB6D39FDDA4FEDF65DC" blockId="14.[123,1174,219,1725]" pageId="14" pageNumber="465">At present, this species has been recorded only from the southwestern part of the study area (Core Area 6).</paragraph>
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<emphasis id="DE908E12FF81FFB6D39FDE63FDE7663C" bold="true" box="[171,557,1540,1563]" pageId="14" pageNumber="465">Overall geographical distribution.</emphasis>
<figureCitation id="74DF4E85FF81FFB6D106DE63FD4A663B" box="[562,640,1540,1564]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="14" pageNumber="465">Fig. 23</figureCitation>
.
<collectingMunicipality id="0C3FC87AFF81FFB6D1BDDE63FD78663B" box="[649,690,1540,1564]" pageId="14" pageNumber="465">The</collectingMunicipality>
known range of this species as presently conceived extends from
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(
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) eastward across the
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(
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(
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and
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),
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<collectingCountry id="94F31290FF81FFB6D22CDE03FE85665B" box="[280,335,1636,1660]" name="India" pageId="14" pageNumber="465">India</collectingCountry>
(
<collectingRegion id="2E209CE2FF81FFB6D255DE03FE55665B" box="[353,415,1636,1660]" country="India" name="Bihar" pageId="14" pageNumber="465">Bihar</collectingRegion>
,
<location id="E93B04DBFF81FFB6D29ADE03FDDB665B" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D29ADE03FDDB665B" box="[430,529,1636,1660]" country="Niger, Chad, Sudan" county="Lancang County" latitude="22.57925" longLatPrecision="1" longitude="99.99849" municipality="The" name="Jharkand" pageId="14" pageNumber="465" stateProvince="Bihar">Jharkand</location>
,
<collectingRegion id="2E209CE2FF81FFB6D114DE03FDBA665B" box="[544,624,1636,1660]" country="India" name="Sikkim" pageId="14" pageNumber="465">Sikkim</collectingRegion>
, and
<collectingRegion id="2E209CE2FF81FFB6D19ADE03FD32665B" box="[686,760,1636,1660]" country="India" name="Assam" pageId="14" pageNumber="465">Assam</collectingRegion>
)
<collectingCountry id="94F31290FF81FFB6D03FDE03FC22665B" box="[779,1000,1636,1660]" isEnumeration="true" pageId="14" pageNumber="465">Bhutan, Bangladesh</collectingCountry>
,
<collectingCountry id="94F31290FF81FFB6D0C3DE02FBAA665A" box="[1015,1120,1637,1661]" name="Myanmar" pageId="14" pageNumber="465">Myanmar</collectingCountry>
, and western
<collectingCountry id="94F31290FF81FFB6D3E3DEE3FEDD66BB" box="[215,279,1668,1692]" name="China" pageId="14" pageNumber="465">China</collectingCountry>
(western
<collectingRegion id="2E209CE2FF81FFB6D2B0DEE2FDF566BB" box="[388,575,1668,1692]" country="China" name="Yunnan" pageId="14" pageNumber="465">Yunnan Province</collectingRegion>
) and south to northwestern
<collectingCountry id="94F31290FF81FFB6D047DEE3FC1F66BB" box="[883,981,1668,1692]" name="Thailand" pageId="14" pageNumber="465">Thailand</collectingCountry>
. A large series of specimens (a total of
<specimenCount id="FAE29989FF81FFB6D246DEC2FE1C669A" box="[370,470,1701,1725]" count="95" pageId="14" pageNumber="465" type="male">95 males</specimenCount>
and females, in
<collectionCode id="8AF5CAC5FF81FFB6D1ADDEC2FD0D669A" box="[665,711,1701,1725]" pageId="14" pageNumber="465">IOZ</collectionCode>
) from
<location id="E93B04DBFF81FFB6D02DDEC2FCA5669A" LSID="urn:lsid:plazi:treatment:644DE316FF83FFB7D34FDB25FC2A61DF:E93B04DBFF81FFB6D02DDEC2FCA5669A" box="[793,879,1701,1725]" country="Niger, Chad, Sudan" county="Lancang County" latitude="22.57925" longLatPrecision="1" longitude="99.99849" municipality="The" name="Mafang" pageId="14" pageNumber="465" stateProvince="Bihar">Mafang</location>
,
<collectingCounty id="053A2A8CFF81FFB6D04BDEC2FBFC669A" box="[895,1078,1701,1725]" pageId="14" pageNumber="465">Lancang County</collectingCounty>
,
<collectorName id="411137D6FF81FFB6D770DEC2FB5F669B" box="[1092,1173,1701,1724]" pageId="14" pageNumber="465">Yunnan</collectorName>
(
<geoCoordinate id="89D034C7FF80FFB7D3B6D8BCFECE60D4" box="[130,260,219,243]" degrees="22.57925" direction="north" orientation="latitude" pageId="15" pageNumber="466" precision="1" value="22.57925">N22.57925°</geoCoordinate>
/
<geoCoordinate id="89D034C7FF80FFB7D220D8BCFE6960D4" box="[276,419,219,243]" degrees="099.99849" direction="east" orientation="longitude" pageId="15" pageNumber="466" precision="1" value="99.99849">E099.99849°</geoCoordinate>
,
<quantity id="2B1CFFE5FF80FFB7D29BD8BCFE3660D4" box="[431,508,219,243]" metricMagnitude="3" metricUnit="m" metricValue="1.723" pageId="15" pageNumber="466" unit="m" value="1723.0">
<elevation id="67C9B533FF80FFB7D29BD8BCFE3660D4" box="[431,508,219,243]" metricMagnitude="3" metricUnit="m" metricValue="1.723" pageId="15" pageNumber="466" unit="m" value="1723.0">1723m</elevation>
</quantity>
,
<collectorName id="411137D6FF80FFB7D132D8BCFDAF60D4" box="[518,613,219,243]" pageId="15" pageNumber="466">W.B. Gu</collectorName>
collector,
<date id="985A74C0FF80FFB7D1E2D8BCFC4A60D4" box="[726,896,219,243]" pageId="15" pageNumber="466" value="2003-12">
<collectingDate id="881E8D28FF80FFB7D1E2D8BCFC4A60D4" box="[726,896,219,243]" pageId="15" pageNumber="466" value="2003-12">December 2003</collectingDate>
</date>
-
<date id="985A74C0FF80FFB7D0B3D8BCFBD160D4" box="[903,1051,219,243]" pageId="15" pageNumber="466" value="2005-01">
<collectingDate id="881E8D28FF80FFB7D0B3D8BCFBD160D4" box="[903,1051,219,243]" pageId="15" pageNumber="466" value="2005-01">January 2005</collectingDate>
</date>
) represents the eastern most record for this species.
</paragraph>
<paragraph id="EC5B5200FF80FFB7D39FD97AFC2A61DF" blockId="15.[123,1174,219,505]" pageId="15" pageNumber="466">
<emphasis id="DE908E12FF80FFB7D39FD97AFCBA6113" bold="true" box="[171,880,284,308]" pageId="15" pageNumber="466">
Geographical relationships with other
<taxonomicName id="2BE42983FF80FFB7D152D97BFCBA6113" box="[614,880,284,308]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="DE908E12FF80FFB7D152D97BFCD86113" bold="true" box="[614,786,284,308]" italics="true" pageId="15" pageNumber="466">Tetragonoderus</emphasis>
species.
</taxonomicName>
</emphasis>
The geographical range of
<taxonomicName id="2BE42983FF80FFB7D34FD959FF396172" box="[123,243,318,341]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF80FFB7D34FD959FF396172" box="[123,243,318,341]" italics="true" pageId="15" pageNumber="466">T. arcuatus</emphasis>
</taxonomicName>
overlaps with several other species in different parts of its range, and these relationships will be addressed in a forthcoming review of the Asian
<taxonomicName id="2BE42983FF80FFB7D015D93AFC036152" authorityName="Dejean" authorityYear="1829" box="[801,969,349,373]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="genus">
<emphasis id="DE908E12FF80FFB7D015D93AFC036152" box="[801,969,349,373]" italics="true" pageId="15" pageNumber="466">Tetragonoderus</emphasis>
</taxonomicName>
fauna (in preparation). In the study area, its range overlaps with those of
<taxonomicName id="2BE42983FF80FFB7D1FAD918FCF361B1" box="[718,825,382,406]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF80FFB7D1FAD918FCF361B1" box="[718,825,382,406]" italics="true" pageId="15" pageNumber="466">T. elegans</emphasis>
</taxonomicName>
,
<taxonomicName id="2BE42983FF80FFB7D073D918FC1861B1" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[839,978,382,406]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="parviculus">
<emphasis id="DE908E12FF80FFB7D073D918FC1861B1" box="[839,978,382,406]" italics="true" pageId="15" pageNumber="466">T. parviculus</emphasis>
</taxonomicName>
, and
<taxonomicName id="2BE42983FF80FFB7D739D918FB4561B1" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[1037,1167,383,406]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF80FFB7D739D918FB4561B1" box="[1037,1167,383,406]" italics="true" pageId="15" pageNumber="466">T. punctatus</emphasis>
</taxonomicName>
, although it has not yet been found syntopic with any of these species. As noted below,
<taxonomicName id="2BE42983FF80FFB7D727D9C7FF5561F0" authorityName="Jian &amp; Tian" authorityYear="2009" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF80FFB7D727D9C7FF5561F0" italics="true" pageId="15" pageNumber="466">T. microthorax</emphasis>
</taxonomicName>
(see treatment for
<taxonomicName id="2BE42983FF80FFB7D244D9A7FE3F61F0" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[368,501,448,471]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF80FFB7D244D9A7FE3F61F0" box="[368,501,448,471]" italics="true" pageId="15" pageNumber="466">T. punctatus</emphasis>
</taxonomicName>
) has not yet been recorded from the study area, but its range overlaps with that of
<taxonomicName id="2BE42983FF80FFB7D254D986FE1D61DF" box="[352,471,481,504]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF80FFB7D254D986FE1D61DF" box="[352,471,481,504]" italics="true" pageId="15" pageNumber="466">T. arcuatus</emphasis>
</taxonomicName>
elsewhere and may also do so in the study area.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,142 @@
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<mods:title id="237DE7089E67776536A6C1979BC90F21">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<emphasis id="DE908E12FF87FFB0D281DB2BFC916341" bold="true" box="[437,859,844,870]" pageId="8" pageNumber="459">
Genus
<taxonomicName id="2BE42983FF87FFB0D139DB2BFC916341" ID-CoL="8KWHT" authority="Latreille, 1829" authorityName="Latreille" authorityYear="1829" box="[525,859,844,870]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="DE908E12FF87FFB0D139DB2BFD546341" bold="true" box="[525,670,844,870]" italics="true" pageId="8" pageNumber="459">Cyclosomus</emphasis>
<bibRefCitation id="88752FF1FF87FFB0D192DB2BFC916341" author="LATREILLE, P. A." box="[678,859,844,870]" pageId="8" pageNumber="459" refId="ref17387" refString="LATREILLE, P. A. 1829. Les crustaces, les arachnides et les insectes, distribues en familles naturelles, ouvrage format les tomes 4 et 5 de celui de M. le Baron Cuvier sur le regne animal (deuxieme edition). Tome premier. Deterville, Paris, xxvii + 584 pp." type="book" year="1829">Latreille, 1829</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF87FFB0D34FDB1FFBE963E0" pageId="8" pageNumber="459" type="reference_group">
<paragraph id="EC5B5200FF87FFB0D34FDB1FFBE963E0" blockId="8.[123,1174,888,968]" pageId="8" pageNumber="459">
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<taxonomicName id="2BE42983FF87FFB0D34FDB1FFE5B63A9" ID-CoL="8HFVS" authority="Fabricius, 1790: 221" authorityName="Fabricius" authorityPageNumber="221" authorityYear="1790" box="[123,401,888,910]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="DE908E12FF87FFB0D34FDB1FFF0163A9" box="[123,203,888,910]" italics="true" pageId="8" pageNumber="459">Scolytus</emphasis>
<bibRefCitation id="88752FF1FF87FFB0D3E4DB1FFE5B63A9" author="FABRICIUS, J. C." box="[208,401,888,910]" pageId="8" pageNumber="459" pagination="213 - 228" refId="ref16539" refString="FABRICIUS, J. C. 1790. Nova insectorum genera. Skrivter af Naturhistorie-Selskabet 1: 213 - 228." type="journal article" year="1790">Fabricius, 1790:221</bibRefCitation>
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, in part [junior homonym of
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<emphasis id="DE908E12FF87FFB0D196DB1FFD3863A9" box="[674,754,888,910]" italics="true" pageId="8" pageNumber="459">Scolytus</emphasis>
<bibRefCitation id="88752FF1FF87FFB0D1CCDB1FFC4163A9" author="GEOFFROY, E. L." box="[760,907,888,910]" pageId="8" pageNumber="459" refId="ref16709" refString="GEOFFROY, E. L. 1762. Histoire abregee des insectes qui se trouvent aux environs de Paris; dans laquelle ces animaux sont ranges suivant un ordre methodique. Tome premier. Durand, Paris. xxviii + 523 pp. + 10 pls." type="book" year="1762">Geoffroy, 1762</bibRefCitation>
</taxonomicName>
] (
<typeStatus id="335FECA2FF87FFB0D0A9DB1FFC0263A9" box="[925,968,888,910]" pageId="8" pageNumber="459">type</typeStatus>
species
<emphasis id="DE908E12FF87FFB0D723DB1FFF05638C" italics="true" pageId="8" pageNumber="459">
<taxonomicName id="2BE42983FF87FFB0D723DB1FFBA063A9" ID-CoL="8KVGQ" authorityName="Linnaeus" authorityYear="1758" box="[1047,1130,888,910]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">Carabus</taxonomicName>
limbatus
</emphasis>
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, designated by
<bibRefCitation id="88752FF1FF87FFB0D130DBF2FD18638C" author="LATREILLE, P. A." box="[516,722,917,939]" pageId="8" pageNumber="459" refId="ref17337" refString="LATREILLE, P. A. 1810. Considerations generales sur l'ordre naturel des animaux composant les classes des crustaces, des arachnides, et des insectes; avec un tableau methodique de leurs genres, disposes. en familles. F. Schoell, Paris, 444 pp." type="book" year="1810">Latreille (1810:426))</bibRefCitation>
.
<bibRefCitation id="88752FF1FF87FFB0D1EFDBF2FC63638C" author="FABRICIUS, J. C." box="[731,937,917,939]" pageId="8" pageNumber="459" refId="ref16560" refString="FABRICIUS, J. C. 1792. Entomologia systematica emendate et aucta, secundum classes, ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Tome I. Pars I. Proft, Hafniae, xx + 330 pp." type="book" year="1792">Fabricius (1792:180)</bibRefCitation>
;
<bibRefCitation id="88752FF1FF87FFB0D087DBF2FBBC638C" author="BOUSQUET, Y." box="[947,1142,917,939]" pageId="8" pageNumber="459" pagination="1 - 1722" refId="ref16177" refString="BOUSQUET, Y. 2012. Catalogue of Geadephaga (Coleoptera, Adephaga) of America, North of Mexico. ZooKeys 245: 1 - 1722." type="journal article" year="2012">Bousquet (2012:89)</bibRefCitation>
.
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<emphasis id="DE908E12FF87FFB0D34FDBD5FF2563EF" box="[123,239,946,968]" italics="true" pageId="8" pageNumber="459">Cyclosomus</emphasis>
<bibRefCitation id="88752FF1FF87FFB0D3C1DBD5FE6463EF" author="LATREILLE, P. A." box="[245,430,946,968]" pageId="8" pageNumber="459" refId="ref17387" refString="LATREILLE, P. A. 1829. Les crustaces, les arachnides et les insectes, distribues en familles naturelles, ouvrage format les tomes 4 et 5 de celui de M. le Baron Cuvier sur le regne animal (deuxieme edition). Tome premier. Deterville, Paris, xxvii + 584 pp." type="book" year="1829">Latreille, 1829:394</bibRefCitation>
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(
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species
<taxonomicName id="2BE42983FF87FFB0D10CDBD5FC4363EF" ID-CoL="QWMS" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[568,905,946,968]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="DE908E12FF87FFB0D10CDBD5FD2163EF" box="[568,747,946,968]" italics="true" pageId="8" pageNumber="459">Carabus flexuosus</emphasis>
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, by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF87FFB0D39FDBBCFDE36494" pageId="8" pageNumber="459" type="diagnosis">
<paragraph id="EC5B5200FF87FFB0D39FDBBCFB996454" blockId="8.[123,1174,986,1203]" pageId="8" pageNumber="459">
<emphasis id="DE908E12FF87FFB0D39FDBBCFED563D5" bold="true" box="[171,287,987,1010]" pageId="8" pageNumber="459">Diagnosis.</emphasis>
Adults of
<taxonomicName id="2BE42983FF87FFB0D2A7DBBDFDDE63D5" authorityName="Latreille" authorityYear="1829" box="[403,532,986,1010]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="DE908E12FF87FFB0D2A7DBBDFDDE63D5" box="[403,532,986,1010]" italics="true" pageId="8" pageNumber="459">Cyclosomus</emphasis>
</taxonomicName>
can be distinguished from those of
<taxonomicName id="2BE42983FF87FFB0D0ADDBBDFB5F63D5" box="[921,1173,986,1010]" pageId="8" pageNumber="459">
<emphasis id="DE908E12FF87FFB0D0ADDBBDFB8B63D5" box="[921,1089,986,1010]" italics="true" pageId="8" pageNumber="459">Tetragonoderus</emphasis>
species
</taxonomicName>
in the study area by the following combination of character states: Body size larger, BL = 7.0 mm or more; pronotum (
<figureCitation id="74DF4E85FF87FFB0D260DC7CFE7F6414" box="[340,437,1051,1075]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="8" pageNumber="459">Fig. 13A</figureCitation>
) very broad, trapezoidal, almost as wide as elytra at humeral angles, distinctly widest at base; elytra broadly rounded apically, elytral color pattern as in
<figureCitation id="74DF4E85FF87FFB0D734DC5CFB9C6474" box="[1024,1110,1083,1107]" captionStart="FIGURE" captionStartId="10.[155,167,1587,1606]" captionTargetBox="[126,1180,216,1575]" captionTargetId="figure-67@10.[120,1180,216,1575]" captionTargetPageId="10" captionText="FIGURE 7. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="8" pageNumber="459">Fig. 7A</figureCitation>
; front tarsomeres 1 to 3 markedly expanded laterally (
<figureCitation id="74DF4E85FF87FFB0D14FDC3CFD226454" box="[635,744,1115,1139]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="8" pageNumber="459">Figs. 14A</figureCitation>
,
<figureCitation id="74DF4E85FF87FFB0D1C0DC3CFCE96454" box="[756,803,1115,1139]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="8" pageNumber="459">15A</figureCitation>
) in both males and females.
</paragraph>
<paragraph id="EC5B5200FF87FFB0D39FDC1BFDE36494" blockId="8.[123,1174,986,1203]" pageId="8" pageNumber="459">
<emphasis id="DE908E12FF87FFB0D39FDC1BFED164B4" bold="true" box="[171,283,1148,1171]" pageId="8" pageNumber="459">Diversity:</emphasis>
At present, only five species are recognized in the Asian fauna, and here we record one, a new species, from the study area.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,57 +1,59 @@
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<mods:identifier id="AE9E442C1390C6240E643441A66BC7CC" type="GBIF-Dataset">9235db45-04c4-4570-8987-cb84765763a4</mods:identifier>
<mods:identifier id="9FF6417A147979220A59A3D850B6B9E4" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="670E0E33FB3FD92DBC46162444231F41" type="Zenodo-Dep">11067293</mods:identifier>
</mods:mods>
<treatment id="644DE316FF87FFB4D34FDCBCFDF06134" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134" httpUri="http://treatment.plazi.org/id/644DE316FF87FFB4D34FDCBCFDF06134" lastPageId="12" lastPageNumber="462" pageId="8" pageNumber="459">
<treatment id="644DE316FF87FFB4D34FDCBCFDF06134" ID-DOI="http://doi.org/10.5281/zenodo.12628248" ID-GBIF-Taxon="234412263" ID-Zenodo-Dep="12628248" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134" httpUri="http://treatment.plazi.org/id/644DE316FF87FFB4D34FDCBCFDF06134" lastPageId="12" lastPageNumber="462" pageId="8" pageNumber="459">
<subSubSection id="A4FE018BFF87FFB0D34FDCBCFD556536" pageId="8" pageNumber="459" type="nomenclature">
<paragraph id="EC5B5200FF87FFB0D34FDCBCFCBF64D4" blockId="8.[123,885,1243,1297]" box="[123,885,1243,1267]" pageId="8" pageNumber="459">
<heading id="B713E56CFF87FFB0D34FDCBCFCBF64D4" box="[123,885,1243,1267]" fontSize="10" level="2" pageId="8" pageNumber="459" reason="4">
@ -65,17 +67,26 @@ Kavanaugh and Cueva-Dabkoski
</emphasis>
</heading>
</paragraph>
<paragraph id="EC5B5200FF87FFB0D34FDC9CFD556536" blockId="8.[123,885,1243,1297]" box="[123,671,1275,1297]" pageId="8" pageNumber="459">Figures 5A, 7, 8, 13A, 14A, 15A, 16A, 17A, 21, and 23.</paragraph>
<paragraph id="EC5B5200FF87FFB0D34FDC9CFD556536" blockId="8.[123,885,1243,1297]" box="[123,671,1275,1297]" pageId="8" pageNumber="459">
<figureCitation id="74DF4E85FF87FFB0D34FDC9CFF236536" box="[123,233,1275,1297]" captionStart="FIGURE" captionStartId="5.[686,698,839,858]" captionTargetBox="[666,1158,216,826]" captionTargetId="figure-473@5.[652,1176,216,826]" captionText="FIGURE 5. Left hind tibial medial spur and tarsomere 1, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.12628237" httpUri="https://zenodo.org/record/12628237/files/figure.png" pageId="8" pageNumber="459">Figures 5A</figureCitation>
, 7,
<figureCitation id="74DF4E85FF87FFB0D238DC9CFED36536" box="[268,281,1275,1297]" captionStart="FIGURE" captionStartId="11.[687,699,975,994]" captionTargetBox="[657,1156,383,944]" captionTargetId="figure-502@11.[652,1176,357,962]" captionText="FIGURE 8. Male genitalia, Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012642, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Dorsal aspect; B. Left lateral aspect; scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12628241" httpUri="https://zenodo.org/record/12628241/files/figure.png" pageId="8" pageNumber="459">8</figureCitation>
, 13A, 14A, 15A, 16A, 17A, 21, and 23.
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF87FFB1D39FDD42FE8A6174" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459" type="materials_examined">
<paragraph id="EC5B5200FF87FFB0D39FDD42FB5F669A" blockId="8.[123,1174,1316,1725]" pageId="8" pageNumber="459">
<paragraph id="EC5B5200FF87FFB1D39FDD42FCD26134" blockId="8.[123,1174,1316,1725]" lastBlockId="9.[123,1174,219,1725]" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
<emphasis id="DE908E12FF87FFB0D39FDD42FE9A651B" bold="true" box="[171,336,1317,1340]" pageId="8" pageNumber="459">Type material</emphasis>
.
<materialsCitation id="5C8C585DFF87FFB0D251DD43FB5F669A" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ, CAS, R, NHMUK, BMNH" collectorName="D. H. Kavanaugh &amp; M. Cueva-Dabkoski &amp; P. Pacholatko &amp; L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCount="56" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan" typeStatus="holotype">
<materialsCitation id="5C8C585DFF87FFB0D251DD43FE32659B" ID-GBIF-Occurrence="4902827327" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ" collectorName="Liang H. - B &amp; Shi X. - C" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1015370" specimenCount="1" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan Province" typeStatus="holotype">
<typeStatus id="335FECA2FF87FFB0D251DD43FE3F651B" box="[357,501,1316,1340]" pageId="8" pageNumber="459" type="holotype">HOLOTYPE</typeStatus>
, a male, deposited in
, a
<specimenCount id="FAE29989FF87FFB0D115DD43FD9D651B" box="[545,599,1316,1340]" pageId="8" pageNumber="459" type="male">male</specimenCount>
, deposited in
<collectionCode id="8AF5CAC5FF87FFB0D034DD43FCE5651B" box="[768,815,1316,1340]" pageId="8" pageNumber="459">IOZ</collectionCode>
, labeled: “CASENT 1015370”/ “
, labeled: “
<specimenCode id="BC42FA7BFF87FFB0D099DD43FB4E651B" box="[941,1156,1316,1340]" pageId="8" pageNumber="459">CASENT 1015370</specimenCode>
”/ “
<collectingCountry id="94F31290FF87FFB0D3B0DD23FF17657B" box="[132,221,1348,1372]" name="China" pageId="8" pageNumber="459">CHINA</collectingCountry>
,
<collectingRegion id="2E209CE2FF87FFB0D3DEDD22FE6D657B" box="[234,423,1348,1372]" country="China" name="Yunnan" pageId="8" pageNumber="459">Yunnan Province</collectingRegion>
@ -84,7 +95,7 @@ Kavanaugh and Cueva-Dabkoski
,
<collectingMunicipality id="0C3FC87AFF87FFB0D15CDD23FCD1657B" box="[616,795,1348,1372]" pageId="8" pageNumber="459">Wuhe Township</collectingMunicipality>
,
<location id="E93B04DBFF87FFB0D01CDD23FF79655B" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D01CDD23FF79655B" country="China" county="Tengchong Co." latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" name="Longjiang Bridge on Longchuanjiang" pageId="8" pageNumber="459" stateProvince="Yunnan">Longjiang Bridge on Longchuanjiang</location>
<location id="E93B04DBFF87FFB0D01CDD23FF79655B" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D01CDD23FF79655B" country="China" county="Tengchong Co." latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" name="Longjiang Bridge on Longchuanjiang" pageId="8" pageNumber="459" stateProvince="Yunnan Province">Longjiang Bridge on Longchuanjiang</location>
,
<quantity id="2B1CFFE5FF87FFB0D38ADD03FED8655B" box="[190,274,1380,1404]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="8" pageNumber="459" unit="m" value="1215.0">
<elevation id="67C9B533FF87FFB0D38ADD03FED8655B" box="[190,274,1380,1404]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="8" pageNumber="459" unit="m" value="1215.0">1215 m</elevation>
@ -96,30 +107,52 @@ Kavanaugh and Cueva-Dabkoski
<date id="985A74C0FF87FFB0D16EDD03FCDB655B" box="[602,785,1380,1404]" pageId="8" pageNumber="459" value="2003-10-30">
<collectingDate id="881E8D28FF87FFB0D16EDD03FCDB655B" box="[602,785,1380,1404]" pageId="8" pageNumber="459" value="2003-10-30">30 October 2003</collectingDate>
</date>
, on beach, Liang H.-B. &amp; Shi X.-C. collectors, LHB03-42”/ “
<typeStatus id="335FECA2FF87FFB0D2A5DDE3FDEA65BB" box="[401,544,1412,1436]" pageId="8" pageNumber="459" type="holotype">HOLOTYPE</typeStatus>
, on beach,
<collectorName id="411137D6FF87FFB0D0BFDD03FBC2655B" box="[907,1032,1380,1404]" pageId="8" pageNumber="459">Liang H.-B</collectorName>
. &amp;
<collectorName id="411137D6FF87FFB0D71BDD03FB5B655B" box="[1071,1169,1380,1404]" pageId="8" pageNumber="459">Shi X.-C</collectorName>
. collectors, LHB03-42”/ “HOLOTYPE
<taxonomicName id="2BE42983FF87FFB0D118DDE3FCF165BB" authority="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[556,827,1412,1436]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="acutangulus" status="sp. nov.">
<emphasis id="DE908E12FF87FFB0D118DDE3FCF165BB" box="[556,827,1412,1436]" italics="true" pageId="8" pageNumber="459">Cyclosomus acutangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5A33369FF87FFB0D073DDE2FC4C65BB" box="[839,902,1413,1436]" pageId="8" pageNumber="459" rank="species">sp. n.</taxonomicNameLabel>
<collectorName id="411137D6FF87FFB0D0A5DDE2FB8065BB" box="[913,1098,1412,1436]" pageId="8" pageNumber="459">D.H. Kavanaugh</collectorName>
&amp;
<collectorName id="411137D6FF87FFB0D743DDE2FEE4659B" pageId="8" pageNumber="459">M. Cueva-Dabkoski</collectorName>
2022” [red label]. A total of
<specimenCount id="FAE29989FF87FFB0D145DDC3FD34659A" box="[625,766,1444,1469]" count="28" pageId="8" pageNumber="459" type="generic" typeStatus="paratype">28 paratypes</specimenCount>
:
D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [red label].
</materialsCitation>
<materialsCitation id="5C8C585DFF87FFB0D2CBDDC3FECB65DB" ID-GBIF-Occurrence="4902827302" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="CAS" collectorName="D. H. Kavanaugh &amp; M. Cueva-Dabkoski &amp; P. Pacholatko &amp; L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1012641&quot;" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Yunnan" typeStatus="holotype">
A total of 28 paratypes:
<specimenCount id="FAE29989FF87FFB0D03FDDC3FCB1659B" box="[779,891,1444,1468]" count="5" pageId="8" pageNumber="459" type="male">five males</specimenCount>
and
<specimenCount id="FAE29989FF87FFB0D087DDC3FBFB659B" box="[947,1073,1444,1468]" count="6" pageId="8" pageNumber="459" type="female">six females</specimenCount>
, in
<collectionCode id="8AF5CAC5FF87FFB0D76FDDC3FB5B659B" box="[1115,1169,1444,1468]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="459" type="Museum">CAS</collectionCode>
, same label data as holotype, except first label [unique identifier label] different (“CASENT 1012641” to “CASENT 1012646” and “CASENT 1015371” to “CASENT 1015375”, respectively);
<collectionCode id="8AF5CAC5FF87FFB0D76FDDC3FB5B659B" box="[1115,1169,1444,1468]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="459" type="Museum">CAS</collectionCode>
, same label data as holotype, except first label [unique identifier label] different (“
<specimenCode id="BC42FA7BFF87FFB0D71EDDA3FF2E65DA" pageId="8" pageNumber="459">CASENT 1012641”</specimenCode>
to
</materialsCitation>
<materialsCitation id="5C8C585DFF87FFB0D23DDD82FDD665DA" ID-GBIF-Occurrence="4902827317" box="[265,540,1509,1533]" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ, CAS, R, NHMUK, BMNH" collectorName="D. H. Kavanaugh &amp; M. Cueva-Dabkoski &amp; P. Pacholatko &amp; L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1012646" specimenCount="1" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan" typeStatus="holotype">
<specimenCode id="BC42FA7BFF87FFB0D226DD82FE2E65DA" box="[274,484,1509,1533]" pageId="8" pageNumber="459">CASENT 1012646</specimenCode>
” and
</materialsCitation>
<materialsCitation id="5C8C585DFF87FFB0D110DD82FCEF65DB" ID-GBIF-Occurrence="4902827325" box="[548,805,1509,1533]" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ, CAS, R, NHMUK, BMNH" collectorName="D. H. Kavanaugh &amp; M. Cueva-Dabkoski &amp; P. Pacholatko &amp; L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1015371" specimenCount="1" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan" typeStatus="holotype">
<specimenCode id="BC42FA7BFF87FFB0D119DD82FD3465DA" box="[557,766,1509,1533]" pageId="8" pageNumber="459">CASENT 1015371</specimenCode>
” to
</materialsCitation>
<materialsCitation id="5C8C585DFF87FFB0D019DD82FF6A663A" ID-GBIF-Occurrence="4902827303" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ, CAS, R, NHMUK, BMNH" collectorName="D. H. Kavanaugh &amp; M. Cueva-Dabkoski &amp; P. Pacholatko &amp; L. Dembicky" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCode="CASENT 1015375" specimenCount="1" specimenCount-female="15" specimenCount-male="13" stateProvince="Yunnan" typeStatus="holotype">
<specimenCode id="BC42FA7BFF87FFB0D003DD82FBC065DA" box="[823,1034,1509,1533]" pageId="8" pageNumber="459">CASENT 1015375</specimenCode>
”, respectively);
</materialsCitation>
<materialsCitation id="5C8C585DFF87FFB0D392DE62FBA2661A" ID-GBIF-Occurrence="4902827313" collectingDate="2015-03-22" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="IOZ" collectorName="Liu Y &amp; Shi H. L" country="China" county="Tengchong Co." elevation="1215" latitude="24.89889" location="Longchuan river" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCount="13" specimenCount-female="7" specimenCount-male="6" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="FAE29989FF87FFB0D392DE62FEC0663A" box="[166,266,1541,1565]" count="6" pageId="8" pageNumber="459" type="male">six males</specimenCount>
and
<specimenCount id="FAE29989FF87FFB0D20BDE62FE12663A" box="[319,472,1541,1565]" count="7" pageId="8" pageNumber="459" type="female">seven females</specimenCount>
, in
<collectionCode id="8AF5CAC5FF87FFB0D2CADE62FDE4663A" box="[510,558,1541,1565]" pageId="8" pageNumber="459">IOZ</collectionCode>
. labeled: “China,
. labeled: “
<collectingCountry id="94F31290FF87FFB0D1A9DE62FD2B663A" box="[669,737,1541,1565]" name="China" pageId="8" pageNumber="459">China</collectingCountry>
,
<collectingRegion id="2E209CE2FF87FFB0D1DFDE62FCF5663B" box="[747,831,1541,1564]" country="China" name="Yunnan" pageId="8" pageNumber="459">Yunnan</collectingRegion>
, Tengchong, Wuhe, on beach of
<location id="E93B04DBFF87FFB0D34FDE42FEE4661A" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D34FDE42FEE4661A" box="[123,302,1573,1597]" country="China" county="Tengchong Co." latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" name="Longchuan river" pageId="8" pageNumber="459" stateProvince="Yunnan">Longchuan river</location>
@ -135,17 +168,31 @@ and
<date id="985A74C0FF87FFB0D19DDE42FCDE661A" box="[681,788,1573,1597]" pageId="8" pageNumber="459" value="2015-03-22">
<collectingDate id="881E8D28FF87FFB0D19DDE42FCDE661A" box="[681,788,1573,1597]" pageId="8" pageNumber="459" value="2015-03-22">2015.3.22</collectingDate>
</date>
N, Liu Y., Shi H.L. collectors”;
N,
<collectorName id="411137D6FF87FFB0D008DE42FCB6661A" box="[828,892,1573,1597]" pageId="8" pageNumber="459">Liu Y</collectorName>
.,
<collectorName id="411137D6FF87FFB0D0BFDE42FC28661A" box="[907,994,1573,1597]" pageId="8" pageNumber="459">Shi H.L</collectorName>
. collectors”;
</materialsCitation>
<materialsCitation id="5C8C585DFF87FFB0D75BDE41FBA2665A" ID-GBIF-Occurrence="4902827316" collectingDate="2006-06-03" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="CAS, IOZ" collectorName="Kavanaugh D &amp; Brett R." country="China" county="Wuhe" elevation="1230" latitude="24.89889" location="Longjiangqiao" longLatPrecision="1" longitude="98.66667" municipality="Tengchong" pageId="8" pageNumber="459" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan Prov." typeStatus="holotype">
<specimenCount id="FAE29989FF87FFB0D75BDE41FF65667A" count="1" pageId="8" pageNumber="459" type="male">one male</specimenCount>
and
<specimenCount id="FAE29989FF87FFB0D3C4DE21FEBA667A" box="[240,368,1605,1629]" count="1" pageId="8" pageNumber="459" type="female">one female</specimenCount>
, in
<collectionCode id="8AF5CAC5FF87FFB0D290DE22FE12667A" box="[420,472,1605,1629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="459" type="Museum">CAS</collectionCode>
<collectionCode id="8AF5CAC5FF87FFB0D290DE22FE12667A" box="[420,472,1605,1629]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="459" type="Museum">CAS</collectionCode>
and
<collectionCode id="8AF5CAC5FF87FFB0D12EDE22FD83667A" box="[538,585,1605,1629]" pageId="8" pageNumber="459">IOZ</collectionCode>
, labeled: “China,
, labeled: “
<collectingCountry id="94F31290FF87FFB0D1F3DE22FCC1667A" box="[711,779,1605,1629]" name="China" pageId="8" pageNumber="459">China</collectingCountry>
,
<collectingRegion id="2E209CE2FF87FFB0D028DE21FC7F667A" box="[796,949,1606,1629]" country="China" name="Yunnan" pageId="8" pageNumber="459">Yunnan Prov.</collectingRegion>
, Tengchong, Wuhe, Longjiangqiao, beach,
,
<collectingMunicipality id="0C3FC87AFF87FFB0D0F2DE22FBF5667A" box="[966,1087,1605,1629]" pageId="8" pageNumber="459">Tengchong</collectingMunicipality>
,
<collectingCounty id="053A2A8CFF87FFB0D764DE22FB5B667A" box="[1104,1169,1605,1629]" pageId="8" pageNumber="459">Wuhe</collectingCounty>
,
<location id="E93B04DBFF87FFB0D34FDE02FED3665A" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D34FDE02FED3665A" box="[123,281,1637,1661]" country="China" county="Wuhe" latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Tengchong" name="Longjiangqiao" pageId="8" pageNumber="459" stateProvince="Yunnan Prov.">Longjiangqiao</location>
, beach,
<geoCoordinate id="89D034C7FF87FFB0D244DE02FE38665A" box="[368,498,1637,1661]" degrees="24.89176" direction="north" orientation="latitude" pageId="8" pageNumber="459" precision="1" value="24.89176">24.89176°N</geoCoordinate>
/
<geoCoordinate id="89D034C7FF87FFB0D2CFDE02FDB2665A" box="[507,632,1637,1661]" degrees="98.67551" direction="east" orientation="longitude" pageId="8" pageNumber="459" precision="1" value="98.67551">98.67551°E</geoCoordinate>
@ -157,18 +204,26 @@ and
<date id="985A74C0FF87FFB0D1D9DE02FC87665A" box="[749,845,1637,1661]" pageId="8" pageNumber="459" value="2006-06-03">
<collectingDate id="881E8D28FF87FFB0D1D9DE02FC87665A" box="[749,845,1637,1661]" pageId="8" pageNumber="459" value="2006-06-03">2006.6.3</collectingDate>
</date>
, Kavanaugh D., Brett
<collectionCode id="8AF5CAC5FF87FFB0D708DE02FB99665A" box="[1084,1107,1637,1661]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="459">R</collectionCode>
.”;
,
<collectorName id="411137D6FF87FFB0D06CDE02FC3A665A" box="[856,1008,1637,1661]" pageId="8" pageNumber="459">Kavanaugh D</collectorName>
.,
<collectorName id="411137D6FF87FFB0D734DE01FB9D665A" box="[1024,1111,1637,1661]" pageId="8" pageNumber="459">Brett R.</collectorName>
”;
</materialsCitation>
<materialsCitation id="5C8C585DFF87FFB1D75BDE01FCD26134" ID-GBIF-Occurrence="4902827314" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="NHMUK, BMNH" collectorName="P. Pacholatko &amp; L. Dembicky" country="India" county="Tengchong Co." elevation="150" lastPageId="9" lastPageNumber="460" latitude="24.89889" location="Bhalukpong" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" pageId="8" pageNumber="459" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="ASSAM" typeStatus="holotype">
<specimenCount id="FAE29989FF87FFB0D75BDE01FF6566BA" count="1" pageId="8" pageNumber="459" type="male">one male</specimenCount>
and
<specimenCount id="FAE29989FF87FFB0D3D5DEE1FE9266BA" box="[225,344,1669,1693]" count="1" pageId="8" pageNumber="459" type="female">one female</specimenCount>
, both in
<collectionCode id="8AF5CAC5FF87FFB0D287DEE1FDD166BA" box="[435,539,1670,1693]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="459" type="Museum">NHMUK</collectionCode>
, labeled: “NE India, ASSAM, Bhalukpong,
<date id="985A74C0FF87FFB0D0D9DEE2FB5866BA" box="[1005,1170,1669,1693]" pageId="8" pageNumber="459" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">
<collectingDate id="881E8D28FF87FFB0D0D9DEE2FB5866BA" box="[1005,1170,1669,1693]" pageId="8" pageNumber="459" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">26.v.-3.vi.2006</collectingDate>
</date>
, labeled: “NE
<collectingCountry id="94F31290FF87FFB0D180DEE2FD2766BA" box="[692,749,1669,1693]" name="India" pageId="8" pageNumber="459">India</collectingCountry>
,
<collectingRegion id="2E209CE2FF87FFB0D1C2DEE2FC9F66BA" box="[758,853,1669,1693]" country="India" name="Assam" pageId="8" pageNumber="459">ASSAM</collectingRegion>
,
<location id="E93B04DBFF87FFB0D06ADEE2FC2966BA" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF87FFB0D06ADEE2FC2966BA" box="[862,995,1669,1693]" country="India" county="Tengchong Co." latitude="24.89889" longLatPrecision="1" longitude="98.66667" municipality="Wuhe Township" name="Bhalukpong" pageId="8" pageNumber="459" stateProvince="ASSAM">Bhalukpong</location>
,
<date id="985A74C0FF87FFB0D0D9DEE2FB5866BA" box="[1005,1170,1669,1693]" pageId="8" pageNumber="459" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">26.v.-3.vi.2006</date>
.
<geoCoordinate id="89D034C7FF87FFB0D34FDEC2FF14669A" box="[123,222,1701,1725]" degrees="27" direction="north" minutes="02" orientation="latitude" pageId="8" pageNumber="459" precision="925" value="27.033333">27° 02N</geoCoordinate>
<geoCoordinate id="89D034C7FF87FFB0D3D2DEC2FE82669A" box="[230,328,1701,1725]" degrees="92" direction="east" minutes="35" orientation="longitude" pageId="8" pageNumber="459" precision="925" value="92.583336">92° 35E</geoCoordinate>
@ -182,23 +237,16 @@ leg.”/ “
<collectorName id="411137D6FF87FFB0D1B1DEC2FCD9669A" box="[645,787,1701,1725]" pageId="8" pageNumber="459">L. Dembicky</collectorName>
&amp;
<collectorName id="411137D6FF87FFB0D002DEC1FC0D669A" box="[822,967,1701,1725]" pageId="8" pageNumber="459">P. Pacholátko</collectorName>
<collectionCode id="8AF5CAC5FF87FFB0D0FADEC1FBD4669A" box="[974,1054,1702,1725]" pageId="8" pageNumber="459">BMNH</collectionCode>
{E} 2006-
</materialsCitation>
</paragraph>
<paragraph id="EC5B5200FF86FFB1D34FD8BCFCD26134" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
48” Each
<typeStatus id="335FECA2FF86FFB1D3DED8BBFE8C60D3" box="[234,326,220,244]" pageId="9" pageNumber="460" type="paratype">paratype</typeStatus>
also bears the following label: “
<typeStatus id="335FECA2FF86FFB1D185D8BCFCF260D4" box="[689,824,219,243]" pageId="9" pageNumber="460" type="paratype">PARATYPE</typeStatus>
<collectionCode id="8AF5CAC5FF87FFB0D0FADEC1FBD4669A" box="[974,1054,1702,1725]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="8" pageNumber="459">BMNH</collectionCode>
{E} 2006- 48” Each paratype also bears the following label: “PARATYPE
<taxonomicName id="2BE42983FF86FFB1D076D8BCFB8560D4" authority="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[834,1103,219,243]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="acutangulus" status="sp. nov.">
<emphasis id="DE908E12FF86FFB1D076D8BCFB8560D4" box="[834,1103,219,243]" italics="true" pageId="9" pageNumber="460">Cyclosomus acutangulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5A33369FF86FFB1D76DD8BBFB5F60D4" box="[1113,1173,220,243]" pageId="9" pageNumber="460" rank="species">sp. n.</taxonomicNameLabel>
D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [yellow label].
</materialsCitation>
</paragraph>
<paragraph id="EC5B5200FF86FFB1D39FD97BFE8A6174" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
<materialsCitation id="5C8C585DFF86FFB1D39FD97BFEF66174" country="China" county="Tengchong County" location="Longchuan River" municipality="Wuhe Township" pageId="9" pageNumber="460" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="DE908E12FF86FFB1D39FD97BFE826114" bold="true" box="[171,328,284,307]" pageId="9" pageNumber="460">
<typeStatus id="335FECA2FF86FFB1D39FD97BFF296114" box="[171,227,284,307]" pageId="9" pageNumber="460">Type</typeStatus>
locality.
@ -214,7 +262,6 @@ locality.
<location id="E93B04DBFF86FFB1D081D97CFBBF6114" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF86FFB1D081D97CFBBF6114" box="[949,1141,283,307]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longchuan River" pageId="9" pageNumber="460" stateProvince="Yunnan">Longchuan River</location>
at
<location id="E93B04DBFF86FFB1D34FD95CFEF66174" LSID="urn:lsid:plazi:treatment:644DE316FF87FFB4D34FDCBCFDF06134:E93B04DBFF86FFB1D34FD95CFEF66174" box="[123,316,315,339]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longjiang Bridge" pageId="9" pageNumber="460" stateProvince="Yunnan">Longjiang Bridge</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
@ -282,9 +329,7 @@ some specimens, portion on intervals 2 to 4 located at about three-fourths of el
<paragraph id="EC5B5200FF86FFB1D39FDDA4FBA2669A" blockId="9.[123,1174,219,1725]" pageId="9" pageNumber="460">
Prothorax. Pronotum (
<figureCitation id="74DF4E85FF86FFB1D2ADDDA4FE3365FC" box="[409,505,1475,1499]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="9" pageNumber="460">Fig. 13A</figureCitation>
) broad but slightly narrow for genus (ratio PWM/
<collectionCode id="8AF5CAC5FF86FFB1D724DDA4FBE565FC" box="[1040,1071,1475,1499]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="9" pageNumber="460" type="Museum">PL</collectionCode>
= 2.12 to 2.43), trapezoidal, almost as wide as elytra at humeral angles, with anterior angles acute, narrow; apical margin markedly concave between anterior angles; lateral margins slightly to distinctly sinuate near anterior angles, otherwise smoothly curved or nearly straight; basal margin markedly bisinuate, lobate medially; anterior and lateral margination absent or extremely faintly impressed, basal margination distinct laterally, obsolete medially; midlateral setae inserted slightly anterior to mid-length, basolateral setae inserted on lateral edge of pronotum and just anterior to posterior angles. Prosternal intercoxal process moderately long, lanceolate, with complete margination.
) broad but slightly narrow for genus (ratio PWM/PL = 2.12 to 2.43), trapezoidal, almost as wide as elytra at humeral angles, with anterior angles acute, narrow; apical margin markedly concave between anterior angles; lateral margins slightly to distinctly sinuate near anterior angles, otherwise smoothly curved or nearly straight; basal margin markedly bisinuate, lobate medially; anterior and lateral margination absent or extremely faintly impressed, basal margination distinct laterally, obsolete medially; midlateral setae inserted slightly anterior to mid-length, basolateral setae inserted on lateral edge of pronotum and just anterior to posterior angles. Prosternal intercoxal process moderately long, lanceolate, with complete margination.
</paragraph>
<caption id="B89B0288FF85FFB2D3AFDE54FEA766B5" ID-DOI="http://doi.org/10.5281/zenodo.11067301" ID-Zenodo-Dep="11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="10" pageNumber="461" startId="10.[155,167,1587,1606]" targetBox="[126,1180,216,1575]" targetPageId="10" targetType="figure">
<paragraph id="EC5B5200FF85FFB2D3AFDE54FEA766B5" blockId="10.[123,1174,1587,1683]" pageId="10" pageNumber="461">
@ -299,10 +344,9 @@ Kavanaugh &amp; Cueva-Dabkoski
A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<paragraph id="EC5B5200FF84FFB3D39FD8BCFEB963F4" blockId="11.[123,1174,219,1715]" pageId="11" pageNumber="462">
Elytra. Broadly ovate, not or only slightly longer than wide (ratio EL/
<collectionCode id="8AF5CAC5FF84FFB3D080D8BCFC1760D4" box="[948,989,219,243]" pageId="11" pageNumber="462">EW</collectionCode>
= 1.00 to 1.02), broadly rounded apically, humeri obtusely angulate; elytral striae deeply impressed throughout, elytral intervals slightly convex, smooth; parascutellar setiferous pore present at base of interval 1 near junction of striae 1 and 2; two discal setiferous pores present on interval three adjacent to stria 2, one near elytral mid-length and the other near apical one-third; apical seta present, inserted near apex of interval 3; umbilicate series comprised of 11 to 13 setae. Legs. Tarsal claws, smooth, edentate. Males (
<paragraph id="EC5B5200FF84FFB3D39FD8BCFE2461F5" pageId="11" pageNumber="462">Elytra. Broadly ovate, not or only slightly longer than wide (ratio EL/EW = 1.00 to 1.02), broadly rounded apically, humeri obtusely angulate; elytral striae deeply impressed throughout, elytral intervals slightly convex, smooth; parascutellar setiferous pore present at base of interval 1 near junction of striae 1 and 2; two discal setiferous pores present on interval three adjacent to stria 2, one near elytral mid-length and the other near apical one-third; apical seta present, inserted near apex of interval 3; umbilicate series comprised of 11 to 13 setae.</paragraph>
<paragraph id="EC5B5200FF84FFB3D39FD9BCFE966334" pageId="11" pageNumber="462">
Legs. Tarsal claws, smooth, edentate. Males (
<figureCitation id="74DF4E85FF84FFB3D3F9D99CFEFB6234" box="[205,305,507,531]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="11" pageNumber="462">Fig. 14A</figureCitation>
) and females with front tarsi similar in shape and width, but males with tarsomeres 1 to 3 with two rows of adhesive squamosetae (
<bibRefCitation id="88752FF1FF84FFB3D224DA3CFE436254" author="STORK, N. E." box="[272,393,603,627]" pageId="11" pageNumber="462" pagination="173 - 306" refId="ref17965" refString="STORK, N. E. 1980. A scanning electron microscope study of tarsal adhesive setae in Coleoptera). Zoological Journal of the Linnaean Society 68: 173 - 306." type="journal article" year="1980">Stork 1980</bibRefCitation>
@ -312,32 +356,41 @@ Elytra. Broadly ovate, not or only slightly longer than wide (ratio EL/
<figureCitation id="74DF4E85FF84FFB3D3B7DADCFF2F62F4" box="[131,229,699,723]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="11" pageNumber="462">Fig. 16A</figureCitation>
) than in females and with two rows of adhesive squamosetae ventrally (
<figureCitation id="74DF4E85FF84FFB3D134DABCFDAE62D4" box="[512,612,731,755]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="11" pageNumber="462">Fig. 17A</figureCitation>
), absent from females. Male genitalia. Median lobe (Fig. 8) stout, smoothly arcuate, and with apical lamella tapered to a rounded point apically in lateral view, slightly sinusoidal and with apical lamella broadly rounded in dorsal view. Internal sac without evident spines.
), absent from females.
</paragraph>
<paragraph id="EC5B5200FF84FFB4D19BDBA8FDF06134" blockId="11.[123,1174,219,1715]" lastBlockId="12.[123,1174,219,275]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
<subScript id="70605045FF84FFB3D19BDBA8FB5C63C6" attach="right" box="[687,1174,974,994]" fontSize="8" pageId="11" pageNumber="462">
<paragraph id="EC5B5200FF84FFB3D39FDB7CFEB963F4" pageId="11" pageNumber="462">
Male genitalia. Median lobe (
<figureCitation id="74DF4E85FF84FFB3D2D3DB7CFDEC6314" box="[487,550,795,819]" captionStart="FIGURE" captionStartId="11.[687,699,975,994]" captionTargetBox="[657,1156,383,944]" captionTargetId="figure-502@11.[652,1176,357,962]" captionText="FIGURE 8. Male genitalia, Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012642, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Dorsal aspect; B. Left lateral aspect; scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12628241" httpUri="https://zenodo.org/record/12628241/files/figure.png" pageId="11" pageNumber="462">Fig. 8</figureCitation>
) stout, smoothly arcuate, and with apical lamella tapered to a rounded point apically in lateral view, slightly sinusoidal and with apical lamella broadly rounded in dorsal view. Internal sac without evident spines.
</paragraph>
<caption id="B89B0288FF84FFB3D19BDBA8FBF5646F" ID-DOI="http://doi.org/10.5281/zenodo.12628241" ID-Zenodo-Dep="12628241" httpUri="https://zenodo.org/record/12628241/files/figure.png" pageId="11" pageNumber="462" startId="11.[687,699,975,994]" targetBox="[657,1156,383,944]" targetPageId="11">
<paragraph id="EC5B5200FF84FFB3D19BDBA8FBF5646F" pageId="11" pageNumber="462">
<subScript id="70605045FF84FFB3D19BDBA8FB5F6432" attach="right" fontSize="8" pageId="11" pageNumber="462">
<smallCapsWord id="EABDC4DCFF84FFB3D19BDBA8FD2763C6" baselines="989,989" box="[687,749,975,994]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="11" pageNumber="462">FIGURE</smallCapsWord>
8. Male genitalia,
<taxonomicName id="2BE42983FF84FFB3D087DBA9FB5C63C6" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[947,1174,974,993]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="DE908E12FF84FFB3D087DBA9FB5C63C6" box="[947,1174,974,993]" italics="true" pageId="11" pageNumber="462">Cyclosomus acutangulus</emphasis>
</taxonomicName>
Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012642, Longchuan Jiang at Longjiang Bridge, Wuhe
</subScript>
Geographical variation. The two speci-
<subScript id="70605045FF84FFB3D1BBDB8FFB5C63DC" attach="right" box="[655,1174,1000,1020]" fontSize="8" pageId="11" pageNumber="462">Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT</subScript>
mens from
<collectingCountry id="94F31290FF84FFB3D236DB9CFEF36434" box="[258,313,1019,1043]" name="India" pageId="11" pageNumber="462">India</collectingCountry>
lack a preapical dark spot
<subScript id="70605045FF84FFB3D1BBDC65FB5F6432" attach="right" box="[655,1173,1026,1045]" fontSize="8" pageId="11" pageNumber="462">1012642, Longchuan Jiang at Longjiang Bridge, Wuhe</subScript>
found in specimens from the study area, but Township, Tengchong County,
Township, Tengchong County,
<collectingRegion id="2E209CE2FF84FFB3D091DC7BFC226408" box="[933,1000,1052,1071]" country="China" name="Yunnan" pageId="11" pageNumber="462">Yunnan</collectingRegion>
,
<collectingCountry id="94F31290FF84FFB3D0C2DC7CFBE76409" box="[1014,1069,1051,1070]" name="China" pageId="11" pageNumber="462">China</collectingCountry>
). A. Dorsal otherwise are similar.
). A. Dorsal
<superScript id="1B91FF48FF84FFB3D1BBDC52FBF5646F" attach="right" box="[655,1087,1077,1096]" fontSize="8" pageId="11" pageNumber="462">
aspect; B. Left lateral aspect; scale line =
<quantity id="2B1CFFE5FF84FFB3D0C3DC52FBF6646F" box="[1015,1084,1077,1096]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="462" unit="mm" value="0.5">0.5 mm</quantity>
.
</superScript>
</paragraph>
</caption>
<paragraph id="EC5B5200FF84FFB3D39FDBBCFEA86474" blockId="11.[123,626,987,1107]" pageId="11" pageNumber="462">
Geographical variation. The two specimens from
<collectingCountry id="94F31290FF84FFB3D236DB9CFEF36434" box="[258,313,1019,1043]" name="India" pageId="11" pageNumber="462">India</collectingCountry>
lack a preapical dark spot found in specimens from the study area, but otherwise are similar.
</paragraph>
<paragraph id="EC5B5200FF84FFB3D39FDC3BFB5C64D4" blockId="11.[123,1174,1115,1715]" pageId="11" pageNumber="462">
<emphasis id="DE908E12FF84FFB3D39FDC3BFE526454" bold="true" box="[171,408,1116,1139]" pageId="11" pageNumber="462">Habitat distribution.</emphasis>
Specimens of this species were collected mainly at night, with the aid of headlamps, on the upper edges of open sandy beaches along the Longchuan River (
<figureCitation id="74DF4E85FF84FFB3D717DC1CFBBB64B4" box="[1059,1137,1147,1171]" captionStart="FIGURE" captionStartId="38.[155,167,1623,1642]" captionTargetBox="[136,1161,216,1607]" captionTargetId="figure-10@38.[136,1161,216,1607]" captionTargetPageId="38" captionText="FIGURE 21. Photographs of habitat for cyclosomine species, Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., Tetragonoderus elegans Andrewes, T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., and T. punctatus Wiedemann at Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China. A. lower river bank; B. sandy upper bank with sparse vegetation. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067337" httpUri="https://zenodo.org/record/11067337/files/figure.png" pageId="11" pageNumber="462">Fig. 21</figureCitation>
@ -345,7 +398,9 @@ Specimens of this species were collected mainly at night, with the aid of headla
<quantity id="2B1CFFE5FF84FFB3D216DCFCFEBC6495" box="[290,374,1179,1203]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="11" pageNumber="462" unit="m" value="1215.0">1215 m</quantity>
, where they were found active on the surface of dry or only slightly damp sandy substrate with no or only widely scattered and low vegetation. Members of this genus are known to burrow quickly into dry sand to hide when disturbed during the daytime (
<bibRefCitation id="88752FF1FF84FFB3D0CCDCBCFB4364D4" author="NIETNER, J." box="[1016,1161,1243,1267]" pageId="11" pageNumber="462" pagination="132 - 153" refId="ref17620" refString="NIETNER, J. 1857. Entomological papers, being chiefly descriptions of new Ceylon Coleoptera with such observations on their habits etc., as appear in any way interesting. Journal of the Asiatic Society of Bengal 26: 132 - 153." type="journal article" year="1857">Nietner 1857</bibRefCitation>
).
).
</paragraph>
<paragraph id="EC5B5200FF84FFB3D39FDC9BFBAA6514" blockId="11.[123,1174,1115,1715]" pageId="11" pageNumber="462">
<emphasis id="DE908E12FF84FFB3D39FDC9BFCD96534" bold="true" box="[171,787,1276,1299]" pageId="11" pageNumber="462">Geographical distribution within the Gaoligong Shan.</emphasis>
<figureCitation id="74DF4E85FF84FFB3D02EDC9CFCA56535" box="[794,879,1275,1299]" captionStart="FIGURE" captionStartId="10.[155,167,1587,1606]" captionTargetBox="[126,1180,216,1575]" captionTargetId="figure-67@10.[120,1180,216,1575]" captionTargetPageId="10" captionText="FIGURE 7. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="11" pageNumber="462">Fig. 7B</figureCitation>
. We examined a total of
@ -356,7 +411,10 @@ and
<specimenCount id="FAE29989FF84FFB3D2BBDD7CFDCC6514" box="[399,518,1307,1331]" count="16" pageId="11" pageNumber="462" type="female">16 females</specimenCount>
) from the study area (see list of
<typeStatus id="335FECA2FF84FFB3D050DD7CFC586514" box="[868,914,1307,1331]" pageId="11" pageNumber="462">type</typeStatus>
specimens above). Members of this species were collected only in the southwestern part of the study area (Core Area 6).
specimens above).
</paragraph>
<paragraph id="EC5B5200FF84FFB3D39FDD5CFF196554" blockId="11.[123,1174,1115,1715]" pageId="11" pageNumber="462">Members of this species were collected only in the southwestern part of the study area (Core Area 6).</paragraph>
<paragraph id="EC5B5200FF84FFB3D39FDD1BFE276614" blockId="11.[123,1174,1115,1715]" pageId="11" pageNumber="462">
<emphasis id="DE908E12FF84FFB3D39FDD1BFDFA65B4" bold="true" box="[171,560,1404,1427]" pageId="11" pageNumber="462">Overall geographical distribution.</emphasis>
<figureCitation id="74DF4E85FF84FFB3D103DD1CFD4E65B4" box="[567,644,1403,1427]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="11" pageNumber="462">Fig. 23</figureCitation>
. At present, this species is known only from single areas in western
@ -377,7 +435,9 @@ apart, and it is likely that this species occurs also in suitable habitats in th
<collectingCountry id="94F31290FF84FFB3D0D7DDBCFBD665D4" box="[995,1052,1499,1523]" name="India" pageId="11" pageNumber="462">India</collectingCountry>
, at low elevations (below
<quantity id="2B1CFFE5FF84FFB3D217DD9CFEB26634" box="[291,376,1531,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="462" unit="m" value="1500.0">1500 m</quantity>
) along rivers draining the western and southern slopes of the Himalayan ranges in these areas, respectively.
) along rivers draining the western and southern slopes of the Himalayan ranges in these areas, respectively.
</paragraph>
<paragraph id="EC5B5200FF84FFB4D39FDE5BFDF06134" blockId="11.[123,1174,1115,1715]" lastBlockId="12.[123,1174,219,275]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
<emphasis id="DE908E12FF84FFB3D39FDE5BFCF76674" bold="true" box="[171,829,1595,1619]" pageId="11" pageNumber="462">
Geographical relationships with other
<taxonomicName id="2BE42983FF84FFB3D169DE5CFCF76674" box="[605,829,1595,1619]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="undetermined">
@ -399,9 +459,7 @@ were found syntopic with
</taxonomicName>
members at the locality in
<collectingRegion id="2E209CE2FF84FFB3D18DDE1CFCC866B4" box="[697,770,1659,1683]" country="India" name="Assam" pageId="11" pageNumber="462">Assam</collectingRegion>
[
<collectionCode id="8AF5CAC5FF84FFB3D03BDE1BFCB666B4" box="[783,892,1660,1683]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="462" type="Museum">NHMUK</collectionCode>
], which is the easternmost known locality for the latter species and the westernmost for
[NHMUK], which is the easternmost known locality for the latter species and the westernmost for
<taxonomicName id="2BE42983FF84FFB3D023DEFCFC706694" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[791,954,1691,1715]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="DE908E12FF84FFB3D023DEFCFC706694" box="[791,954,1691,1715]" italics="true" pageId="11" pageNumber="462">C. acutangulus</emphasis>
</taxonomicName>

View file

@ -0,0 +1,229 @@
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<mods:title id="0672CB942B4FC036B95F7F9B0520352B">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:affiliation id="15F60ED1ECACD702A2BA0890600B58B2">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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Key for identification of adult cyclosomine species of the Gaoligong Shan region of
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1 Size larger, BL =
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or more; pronotum (
<figureCitation id="74DF4E85FF88FFBFD159DB44FD0E631E" box="[621,708,803,825]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="7" pageNumber="458">Fig. 13A</figureCitation>
) very broad, trapezoidal, almost as wide as elytra at humeral angles, distinctly widest at base; elytra broadly rounded apically, elytral color pattern as in
<figureCitation id="74DF4E85FF88FFBFD39FDB3AFF366354" box="[171,252,861,883]" captionStart="FIGURE" captionStartId="10.[155,167,1587,1606]" captionTargetBox="[126,1180,216,1575]" captionTargetId="figure-67@10.[120,1180,216,1575]" captionTargetPageId="10" captionText="FIGURE 7. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT 1015370, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067301" httpUri="https://zenodo.org/record/11067301/files/figure.png" pageId="7" pageNumber="458">Fig. 7A</figureCitation>
; front tarsomeres 1 to 3 markedly expanded laterally (
<figureCitation id="74DF4E85FF88FFBFD019DB3AFC596354" box="[813,915,861,883]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="7" pageNumber="458">Figs. 14A</figureCitation>
,
<figureCitation id="74DF4E85FF88FFBFD095DB3AFC016354" box="[929,971,861,883]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="7" pageNumber="458">15A</figureCitation>
) in both males and females...................
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<taxonomicName id="2BE42983FF88FFBFD2D5DB1DFBFC63B7" authority="Kavanaugh and Cueva-Dabkoski" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[481,1078,890,912]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="acutangulus" status="sp. nov.">
<emphasis id="DE908E12FF88FFBFD2D5DB1DFD1363A8" bold="true" box="[481,729,890,911]" italics="true" pageId="7" pageNumber="458">Cyclosomus acutangulus</emphasis>
Kavanaugh and Cueva-Dabkoski
</taxonomicName>
,
<taxonomicNameLabel id="C5A33369FF88FFBFD775DB1CFB4463B7" box="[1089,1166,891,912]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
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1 Size smaller, BL = less than 6.0 mm; pronotum (
<figureCitation id="74DF4E85FF88FFBFD14ADBF0FD06638A" box="[638,716,919,941]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="7" pageNumber="458">Figs. 13</figureCitation>
B-D) narrower, widest at or anterior to middle; elytra slightly to distinctly and obliquely truncate apically, elytral color pattern varied; front tarsomeres 1 to 3 without distinct lateral expansions (
<figureCitation id="74DF4E85FF88FFBFD1BDDBB7FD2863C1" box="[649,738,976,998]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="7" pageNumber="458">Figs. 14</figureCitation>
B-F), broader in males than in females...............................................................
<taxonomicName id="2BE42983FF88FFBFD09EDB8AFB456424" box="[938,1167,1005,1027]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="undefined-2">
<emphasis id="DE908E12FF88FFBFD09EDB8AFB8B6424" box="[938,1089,1005,1027]" italics="true" pageId="7" pageNumber="458">Tetragonoderus</emphasis>
spp. 2
</taxonomicName>
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<keyStep id="57104AA5FF88FFBFD34FDC71FB456442" pageId="7" pageNumber="458">
<paragraph id="EC5B5200FF88FFBFD34FDC71FB45646E" blockId="7.[123,1174,1046,1125]" pageId="7" pageNumber="458">
<keyLead id="5715F135FF88FFBFD34FDC71FB45646E" pageId="7" pageNumber="458">
2 (1) Femora dark black or piceous; elytra dark without pale markings in basal half, with or without pale markings in apical half (
<figureCitation id="74DF4E85FF88FFBFD2A3DC54FE27646E" box="[407,493,1075,1097]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="7" pageNumber="458">Figs. 9A</figureCitation>
,
<figureCitation id="74DF4E85FF88FFBFD2CCDC54FDEA646E" box="[504,544,1075,1097]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="7" pageNumber="458">11A</figureCitation>
)................................................. 3
</keyLead>
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<paragraph id="EC5B5200FF88FFBFD34FDC28FB456442" blockId="7.[123,1174,1046,1125]" box="[123,1167,1103,1125]" pageId="7" pageNumber="458">
<keyLead id="5715F135FF88FFBFD34FDC28FB456442" box="[123,1167,1103,1125]" pageId="7" pageNumber="458">
2 Femora pale tan; elytra with pale areas in both basal and apical halves (
<figureCitation id="74DF4E85FF88FFBFD06BDC28FC0B6442" box="[863,961,1103,1125]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="7" pageNumber="458">Figs. 10A</figureCitation>
,
<figureCitation id="74DF4E85FF88FFBFD0F8DC28FC3D6442" box="[972,1015,1103,1125]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="7" pageNumber="458">12A</figureCitation>
,
<figureCitation id="74DF4E85FF88FFBFD736DC28FBD46442" box="[1026,1054,1103,1125]" captionStart="FIGURE" captionStartId="37.[155,167,1091,1110]" captionTargetBox="[384,875,236,1075]" captionTargetId="figure-14@37.[384,912,216,1079]" captionTargetPageId="37" captionText="FIGURE 20. Tetragonoderus microthorax Jian and Tian, dorsal habitus of holotype (Jinghong, Yunnan, China); scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067333" httpUri="https://zenodo.org/record/11067333/files/figure.png" pageId="7" pageNumber="458">20</figureCitation>
)........ 4
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<paragraph id="EC5B5200FF88FFBFD34FDC1FFB44651C" blockId="7.[123,1174,1144,1541]" pageId="7" pageNumber="458">
<keyLead id="5715F135FF88FFBFD34FDC1FFB44651C" pageId="7" pageNumber="458">
3 (2) Size smaller, BL =
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or less; elytra with distinct transverse subapical band of pale spots extended from interval 4 to interval 8 (
<figureCitation id="74DF4E85FF88FFBFD2D0DCF2FDF7648C" box="[484,573,1173,1195]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="7" pageNumber="458">Fig. 11A</figureCitation>
), faintly present also on interval
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some individuals, posterior discal setiferous puncture on interval 3 inserted more posteriad, at level of pale transverse subapical band; dorsal surfaces shiny, with distinct aeneous-greenish metallic reflection, of similar sheen throughout, elytral microsculpture comprised of more or less regularly transverse meshes, except their orientation slightly distorted in or near discal setal pore punctures...........................................................
<emphasis id="DE908E12FF88FFBFD2FBDD42FB44651C" bold="true" box="[463,1166,1317,1339]" pageId="7" pageNumber="458">
<taxonomicName id="2BE42983FF88FFBFD2FBDD42FBFC651C" authority="Kavanaugh and Cueva-Dabkoski" authorityName="Kavanaugh and Cueva-Dabkoski" authorityYear="2023" box="[463,1078,1317,1339]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="parviculus" status="sp. nov.">
<emphasis id="DE908E12FF88FFBFD2FBDD42FD13651D" bold="true" box="[463,729,1317,1338]" italics="true" pageId="7" pageNumber="458">Tetragonoderus parviculus</emphasis>
Kavanaugh and Cueva-Dabkoski
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,
<taxonomicNameLabel id="C5A33369FF88FFBFD775DD41FB44651C" box="[1089,1166,1318,1339]" pageId="7" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="EC5B5200FF88FFBFD34FDD25FB456622" blockId="7.[123,1174,1144,1541]" pageId="7" pageNumber="458">
<keyLead id="5715F135FF88FFBFD34FDD25FB456622" pageId="7" pageNumber="458">
3 Size larger, BL =
<quantity id="2B1CFFE5FF88FFBFD263DD25FE6A657F" box="[343,416,1346,1368]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="458" unit="mm" value="4.9">4.9 mm</quantity>
or more; elytra (
<figureCitation id="74DF4E85FF88FFBFD171DD25FD5B657F" box="[581,657,1346,1368]" captionStart="FIGURE" captionStartId="13.[155,167,1592,1611]" captionTargetBox="[128,1175,226,1577]" captionTargetId="figure-56@13.[120,1176,216,1581]" captionTargetPageId="13" captionText="FIGURE 9. Tetragonoderus arcuatus (Dejean). A. Dorsal aspect, scale line = 1.0 mm (CASENT1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067303" httpUri="https://zenodo.org/record/11067303/files/figure.png" pageId="7" pageNumber="458">Fig. 9A</figureCitation>
) with only faint pale markings in region of subapical band, most evident on intervals 7 and 8, posterior discal setiferous puncture on interval 3 inserted distinctly anterior to level of pale subapical markings (i.e., closer to anterior discal setiferous pore); dorsal surfaces duller, with distinct bronze/copper metallic reflection, sericeous (silky) and mottled in posterior half, with elytral microsculpture comprised of a complex pattern of transverse, longitudinal, and oblique patches of elongate meshes, most distorted near discal and umbilicate setal pore punctures..................................................................
<taxonomicName id="2BE42983FF88FFBFD067DD88FB456622" authority="Dejean" authorityName="Dejean" authorityYear="1829" box="[851,1167,1519,1541]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF88FFBFD067DD88FB8E6622" box="[851,1092,1519,1541]" italics="true" pageId="7" pageNumber="458">Tetragonoderus arcuatus</emphasis>
Dejean
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<keyStep id="57104AA5FF88FFB0D34FDE70FB4561D3" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
<paragraph id="EC5B5200FF88FFB0D34FDE70FB446129" blockId="7.[123,1174,1559,1725]" lastBlockId="8.[123,1191,219,500]" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
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4 (3) Elytral color pattern as in
<figureCitation id="74DF4E85FF88FFBFD28CDE70FDDA660A" box="[440,528,1559,1581]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="7" pageNumber="458">Fig. 10A</figureCitation>
, with dark areas more expansive; connection between medial portion of the basal dark band and humeral portion broader, at least one-quarter length of the humeral portion; middle band with portion between anterior and posterior dark edges distinctly darker (orange-brown) than remainder of pale elytral areas; pronotum dark, black or piceous, with distinct greenish or bronze metallic reflection, slightly narrowed basally (ratio PWM/PWB =1.13 to 1.20); male with middle tarsomeres 1 to 4 (
<figureCitation id="74DF4E85FF88FFBFD250DEC0FE76669A" box="[356,444,1703,1725]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="7" pageNumber="458">Fig. 16C</figureCitation>
) distinctly wider than in female and with pads of adhesive setae ventrally (
<figureCitation id="74DF4E85FF87FFB0D386D8BCFEC060D6" box="[178,266,219,241]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="8" pageNumber="459">Fig. 17C</figureCitation>
); median lobe of male genitalia (
<figureCitation id="74DF4E85FF87FFB0D17ED8BCFD0A60D6" box="[586,704,219,241]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="8" pageNumber="459">Figs. 19A,B</figureCitation>
) with conspicuous large spines on the internal sac.......................................................
<taxonomicName id="2BE42983FF87FFB0D074D89FFB446129" authority="Andrewes" authorityName="Andrewes" box="[832,1166,248,270]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF87FFB0D074D89FFBED6129" box="[832,1063,248,270]" italics="true" pageId="8" pageNumber="459">Tetragonoderus elegans</emphasis>
Andrewes
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<paragraph id="EC5B5200FF87FFB0D34FD973FB4561D3" blockId="8.[123,1191,219,500]" pageId="8" pageNumber="459">
<keyLead id="5715F135FF87FFB0D34FD973FB4561D3" pageId="8" pageNumber="459">
4 Elytral color pattern as in
<figureCitation id="74DF4E85FF87FFB0D290D973FDCE610D" box="[420,516,276,298]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="8" pageNumber="459">Figs. 12A</figureCitation>
,
<figureCitation id="74DF4E85FF87FFB0D13AD973FDE2610D" box="[526,552,276,298]" captionStart="FIGURE" captionStartId="37.[155,167,1091,1110]" captionTargetBox="[384,875,236,1075]" captionTargetId="figure-14@37.[384,912,216,1079]" captionTargetPageId="37" captionText="FIGURE 20. Tetragonoderus microthorax Jian and Tian, dorsal habitus of holotype (Jinghong, Yunnan, China); scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11067333" httpUri="https://zenodo.org/record/11067333/files/figure.png" pageId="8" pageNumber="459">20</figureCitation>
, with dark areas less expansive; connection between medial portion of the basal macula and humeral macula absent or very narrow, less than one-fifth the length of humeral macula; middle discal band with portion between anterior and posterior dark edges not or only very slightly darker than remainder of pale elytral areas; pronotum piceous or rufopiceous, with or without faint greenish metallic reflection, narrowed basally or not; male with middle mesotarsomeres (
<figureCitation id="74DF4E85FF87FFB0D751D9EFFF2C619C" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="8" pageNumber="459">Figs. 16E,F</figureCitation>
) only slightly wider than in female and with only tarsomeres 1 to 3 with pads of adhesive setae ventrally (i.e., tarsomere 4 without adhesive setae ventrally) (
<figureCitation id="74DF4E85FF87FFB0D1C1D9A5FCA361FF" box="[757,873,450,472]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="8" pageNumber="459">Figs. 17E,F</figureCitation>
); median lobe of male genitalia (
<figureCitation id="74DF4E85FF87FFB0D386D9B9FF3561D3" box="[178,255,478,500]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="8" pageNumber="459">Figs. 19</figureCitation>
C-F) without spines on the internal sac........................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="57104AA5FF87FFB0D34FDA6FFB446304" pageId="8" pageNumber="459">
<paragraph id="EC5B5200FF87FFB0D34FDA6FFB4462B6" blockId="8.[123,1174,520,803]" pageId="8" pageNumber="459">
<keyLead id="5715F135FF87FFB0D34FDA6FFB4462B6" pageId="8" pageNumber="459">
5 (4) Pronotum (
<figureCitation id="74DF4E85FF87FFB0D217DA6FFEBD6239" box="[291,375,520,542]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="8" pageNumber="459">Fig. 13E</figureCitation>
) slightly shorter and wider (ratio PWM/PL = 1.52 to 1.70), widest near mid-length and posterior to insertion of midlateral seta, not or only very slightly narrowed basally (ratio PWB/PWA = 1.15 to 1.25), lateral margins slightly and evenly convex or straight or very slightly sinuate anterior to basal angles; elytra intervals flat and striae moderately impressed; median lobe of male genitalia (
<figureCitation id="74DF4E85FF87FFB0D751DA39FF2162B6" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="8" pageNumber="459">Figs. 19C,D</figureCitation>
) with apical lamella slightly shorter and broader......
<taxonomicName id="2BE42983FF87FFB0D03EDA1CFB4462B6" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[778,1166,635,657]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF87FFB0D03EDA1CFBCF62B6" box="[778,1029,635,657]" italics="true" pageId="8" pageNumber="459">Tetragonoderus punctatus</emphasis>
(Wiedemann)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EC5B5200FF87FFB0D34FDAFFFB446304" blockId="8.[123,1174,520,803]" pageId="8" pageNumber="459">
<keyLead id="5715F135FF87FFB0D34FDAFFFB446304" pageId="8" pageNumber="459">
5 Pronotum (
<figureCitation id="74DF4E85FF87FFB0D222DAFFFEA26289" box="[278,360,664,686]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="8" pageNumber="459">Fig. 13F</figureCitation>
) slightly longer and narrower (ratio PWM/PL = 1.48 to 1.57), widest anterior to mid-length at or near insertion of midlateral seta, slightly to moderately narrowed basally (ratio PWB/PWA = 1.04 to 1.14), lateral margins slightly to moderately sinuate anterior to basal angles; elytra intervals slightly convex and striae deeply impressed; median lobe of male genitalia (
<figureCitation id="74DF4E85FF87FFB0D091DA89FBD36323" box="[933,1049,750,772]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="8" pageNumber="459">Figs. 19E,F</figureCitation>
) with apical lamella slightly longer and narrower................... [
<taxonomicName id="2BE42983FF87FFB0D1CFDB6AFB406304" authority="Jian &amp; Tian" authorityName="Jian &amp; Tian" authorityYear="2009" box="[763,1162,781,803]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF87FFB0D1CFDB6AFBC56304" box="[763,1039,781,803]" italics="true" pageId="8" pageNumber="459">Tetragonoderus microthorax</emphasis>
Jian &amp; Tian
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]
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<mods:title id="28D87BF05AE3BAF6F33773A22D3DB679">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<emphasis id="DE908E12FF9AFFADD34FD95BFE506173" bold="true" box="[123,410,316,340]" italics="true" pageId="21" pageNumber="472">Tetragonoderus punctatus</emphasis>
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<paragraph id="EC5B5200FF9AFFADD34FD93BFD636155" blockId="21.[123,681,316,370]" box="[123,681,348,370]" pageId="21" pageNumber="472">
<figureCitation id="74DF4E85FF9AFFADD34FD93BFF296155" box="[123,227,348,370]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="21" pageNumber="472">
Figures
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,
<figureCitation id="74DF4E85FF9AFFADD215D93BFE836155" box="[289,329,348,370]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="21" pageNumber="472">14E</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD260D93BFEB76155" box="[340,381,348,370]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="21" pageNumber="472">15E</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD2BCD93BFE7B6155" box="[392,433,348,370]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="21" pageNumber="472">16E</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD28FD93BFE2E6155" box="[443,484,348,370]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="21" pageNumber="472">17E</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD2DAD93BFDCF6155" box="[494,517,348,370]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="21" pageNumber="472">19</figureCitation>
C-D, 21, and 23.
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF9AFFADD34FD9E1FD3E6284" pageId="21" pageNumber="472" type="reference_group">
<paragraph id="EC5B5200FF9AFFADD34FD9E1FE4E624E" blockId="21.[123,1175,390,675]" pageId="21" pageNumber="472">
<treatmentCitationGroup id="CCF4752EFF9AFFADD34FD9E1FE4E624E" pageId="21" pageNumber="472">
<treatmentCitation id="6D457411FF9AFFADD34FD9E1FDFF61BB" author="WIEDEMANN, C. R. W." box="[123,565,390,412]" page="61" pageId="21" pageNumber="472" year="1823">
<taxonomicName id="2BE42983FF9AFFADD34FD9E1FDFF61BB" ID-CoL="LG8X" authority="Wiedemann, 1823: 61" authorityName="Wiedemann" authorityPageNumber="61" authorityYear="1823" box="[123,565,390,412]" class="Insecta" family="Carabidae" genus="Bembidium" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatum">
<emphasis id="DE908E12FF9AFFADD34FD9E1FE9D61BB" box="[123,343,390,412]" italics="true" pageId="21" pageNumber="472">Bembidium punctatum</emphasis>
<bibRefCitation id="88752FF1FF9AFFADD255D9E1FDFF61BB" author="WIEDEMANN, C. R. W." box="[353,565,390,412]" pageId="21" pageNumber="472" pagination="162 - 164" refId="ref18095" refString="WIEDEMANN, C. R. W. 1823. Zweihundert neue Kafer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2: 1 - 135, 162 - 164." type="journal article" year="1823">Wiedemann, 1823:61</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="335FECA2FF9AFFADD176D9E1FD0761BB" box="[578,717,390,412]" pageId="21" pageNumber="472" type="lectotype">LECTOTYPE</typeStatus>
, here designated, a male, in ZMUC, labeled: “TYPE” [red label]/ “Bengala Westermann
<taxonomicName id="2BE42983FF9AFFADD172D9C3FD2F619E" ID-CoL="55N24" authority="Wied." authorityName="Wied." box="[582,741,419,441]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">Punctatus Wied.</taxonomicName>
” [handwritten]/ “ZMUC 00027869”/ “
<typeStatus id="335FECA2FF9AFFADD76BD9C4FF3D61F0" pageId="21" pageNumber="472" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="2BE42983FF9AFFADD3CBD9A6FD5961F0" ID-CoL="LG8X" authority="Wiedemann, 1823" authorityName="Wiedemann" authorityYear="1823" box="[255,659,449,471]" class="Insecta" family="Carabidae" genus="Bembidium" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatum">
<emphasis id="DE908E12FF9AFFADD3CBD9A6FE1061F0" box="[255,474,449,471]" italics="true" pageId="21" pageNumber="472">Bembidium punctatum</emphasis>
<bibRefCitation id="88752FF1FF9AFFADD2D6D9A6FD5961F0" author="WIEDEMANN, C. R. W." box="[482,659,449,471]" pageId="21" pageNumber="472" pagination="162 - 164" refId="ref18095" refString="WIEDEMANN, C. R. W. 1823. Zweihundert neue Kafer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2: 1 - 135, 162 - 164." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
</taxonomicName>
design. by D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [red label].
<typeStatus id="335FECA2FF9AFFADD278D9B9FE1261D3" box="[332,472,478,500]" pageId="21" pageNumber="472" type="paralectotype">Paralectotypes</typeStatus>
examined:
<specimenCount id="FAE29989FF9AFFADD17CD9B9FD5461D3" box="[584,670,478,500]" count="1" pageId="21" pageNumber="472" type="female">1 female</specimenCount>
, in ZMUC, labeled: “Mus. Westerm.”/ “TYPE” [red label]/ “Bengal Maj. 1808.
<taxonomicName id="2BE42983FF9AFFADD286D99BFD9E6236" ID-CoL="55N24" authority="Wied." authorityName="Wied." box="[434,596,507,529]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">Punctatus Wied.</taxonomicName>
” [handwritten]/ “ZMUC 00027868” [label upside down]/ “
<typeStatus id="335FECA2FF9AFFADD390DA7FFEA76209" box="[164,365,536,558]" pageId="21" pageNumber="472" type="paralectotype">PARALECTOTYPE</typeStatus>
<taxonomicName id="2BE42983FF9AFFADD247DA7FFCCA6209" ID-CoL="LG8X" authority="Wiedemann, 1823" authorityName="Wiedemann" authorityYear="1823" box="[371,768,536,558]" class="Insecta" family="Carabidae" genus="Bembidium" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatum">
<emphasis id="DE908E12FF9AFFADD247DA7FFD816209" box="[371,587,536,558]" italics="true" pageId="21" pageNumber="472">Bembidium punctatum</emphasis>
<bibRefCitation id="88752FF1FF9AFFADD165DA7FFCCA6209" author="WIEDEMANN, C. R. W." box="[593,768,536,558]" pageId="21" pageNumber="472" pagination="162 - 164" refId="ref18095" refString="WIEDEMANN, C. R. W. 1823. Zweihundert neue Kafer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2: 1 - 135, 162 - 164." type="journal article" year="1823">Wiedemann, 1823</bibRefCitation>
</taxonomicName>
design. by D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [yellow label]. Type locality: “Bengalia” [area which now includes
<collectingRegion id="2E209CE2FF9AFFADD0FDDA51FB89626B" box="[969,1091,566,588]" country="India" name="West Bengal" pageId="21" pageNumber="472">West Bengal</collectingRegion>
(eastern
<collectingCountry id="94F31290FF9AFFADD3AFDA34FF04624E" box="[155,206,595,617]" name="India" pageId="21" pageNumber="472">India</collectingCountry>
) and
<collectingCountry id="94F31290FF9AFFADD230DA34FEB0624E" box="[260,378,595,617]" name="Bangladesh" pageId="21" pageNumber="472">Bangladesh</collectingCountry>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="EC5B5200FF9AFFADD34FDA17FD3E6284" blockId="21.[123,1175,390,675]" pageId="21" pageNumber="472">
<treatmentCitationGroup id="CCF4752EFF9AFFADD34FDA17FD3E6284" pageId="21" pageNumber="472">
<taxonomicName id="2BE42983FF9AFFADD34FDA17FDCB62A1" ID-CoL="55N24" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[123,513,624,646]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF9AFFADD34FDA17FEBF62A1" box="[123,373,624,646]" italics="true" pageId="21" pageNumber="472">Tetragonoderus punctatus</emphasis>
(Wiedemann)
</taxonomicName>
,
<treatmentCitation id="6D457411FF9AFFADD13FDA17FD0862A1" author="DEJEAN, P. F. M. A." box="[523,706,624,646]" page="505" pageId="21" pageNumber="472" year="1829">
<bibRefCitation id="88752FF1FF9AFFADD13FDA17FD0862A1" author="DEJEAN, P. F. M. A." box="[523,706,624,646]" pageId="21" pageNumber="472" refId="ref16379" refString="DEJEAN, P. F. M. A. 1829. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon-Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean (1829:505)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6D457411FF9AFFADD1F8DA17FC0C62A1" author="SCHMIDT-GOBEL, M." box="[716,966,624,646]" page="92" pageId="21" pageNumber="472" year="1846">
<bibRefCitation id="88752FF1FF9AFFADD1F8DA17FC0C62A1" author="SCHMIDT-GOBEL, M." box="[716,966,624,646]" pageId="21" pageNumber="472" pagination="1 - 3" refId="ref17793" refString="SCHMIDT-GOBEL, M. 1846. Faunula coleopterorum Birmaniae, adjectis nonnulis Bengaliae indigenis. Med. Dr. Johann Wilhelm Helfer's hinterlassene Sammlungen aus Vorder- und Hinter-Indien. Nach seinem Tode im Auftrage des bohm. National-Museums unter Mitwirkung Mehrerer bearbeitet und herausgegeben. G. Haase Sohne, Prague, viii + 94 pp. + plates 1 - 3." type="book chapter" year="1846">Schmidt-Göbel (1846:92)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6D457411FF9AFFADD0E5DA17FB5B62A1" author="CHAUDOIR, M." box="[977,1169,624,646]" page="48" pageId="21" pageNumber="472" year="1876">
<bibRefCitation id="88752FF1FF9AFFADD0E5DA17FB5B62A1" author="CHAUDOIR, M." box="[977,1169,624,646]" pageId="21" pageNumber="472" pagination="1 - 84" refId="ref16317" refString="CHAUDOIR, M. 1876. Etude monographique des Masoreides, des Tetragonoderides et du genre Nemotarsus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 51: 1 - 84." type="journal article" year="1876">Chaudoir (1876:48)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6D457411FF9AFFADD3AFDAEAFEBA6284" author="ANDREWES, H. E." box="[155,368,653,675]" page="174" pageId="21" pageNumber="472" year="1921">
<bibRefCitation id="88752FF1FF9AFFADD3AFDAEAFEBA6284" author="ANDREWES, H. E." box="[155,368,653,675]" pageId="21" pageNumber="472" pagination="145 - 195" refId="ref15718" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:174)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6D457411FF9AFFADD24FDAEAFDE46284" author="CSIKI, E." box="[379,558,653,675]" page="1298" pageId="21" pageNumber="472" year="1932">
<bibRefCitation id="88752FF1FF9AFFADD24FDAEAFDE46284" author="CSIKI, E." box="[379,558,653,675]" pageId="21" pageNumber="472" pagination="1279 - 1598" refId="ref16350" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1298)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6D457411FF9AFFADD10CDAE9FD3A6284" author="LORENZ, W." box="[568,752,653,675]" page="453" pageId="21" pageNumber="472" year="2005">
<bibRefCitation id="88752FF1FF9AFFADD10CDAE9FD3A6284" author="LORENZ, W." box="[568,752,653,675]" pageId="21" pageNumber="472" refId="ref17529" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz (2005:453)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A4FE018BFF9AFFA0D39FDADFFF6861B4" lastPageId="24" lastPageNumber="475" pageId="21" pageNumber="472" type="diagnosis">
<paragraph id="EC5B5200FF9AFFADD39FDADFFD6164B2" blockId="21.[123,1174,695,1725]" pageId="21" pageNumber="472">
<emphasis id="DE908E12FF9AFFADD39FDADFFED562E8" bold="true" box="[171,287,696,719]" pageId="21" pageNumber="472">Diagnosis.</emphasis>
Adults of
<taxonomicName id="2BE42983FF9AFFADD294DADFFDE262E8" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[416,552,696,719]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF9AFFADD294DADFFDE262E8" box="[416,552,696,719]" italics="true" pageId="21" pageNumber="472">T. punctatus</emphasis>
</taxonomicName>
can be distinguished from those of other
<emphasis id="DE908E12FF9AFFADD726DAD0FB5C62E8" box="[1042,1174,695,719]" italics="true" pageId="21" pageNumber="472">cyclosomine</emphasis>
species in the study area by the following combination of character states: Body size medium for genus, BL males =
<quantity id="2B1CFFE5FF9AFFADD273DA9FFE146337" box="[327,478,760,784]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="21" pageNumber="472" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5 to 4.8 mm</quantity>
, females =
<quantity id="2B1CFFE5FF9AFFADD163DA9FFD246337" box="[599,750,760,784]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.4" metricValueMin="4.6" pageId="21" pageNumber="472" unit="mm" value="5.0" valueMax="5.4" valueMin="4.6">4.6 to 5.4 mm</quantity>
; pronotum piceous or rufopiceous, with or without faint greenish metallic reflection; elytral color pattern as in
<figureCitation id="74DF4E85FF9AFFADD04BDB7FFC286317" box="[895,994,792,816]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="21" pageNumber="472">Fig. 12A</figureCitation>
, with dark areas less expansive, connection between medial portion of the basal dark band and humeral portion absent or very narrow, less than one-fifth the length of humeral portion, middle discal band with portion between anterior and posterior dark edges not or only slightly darker than remainder of pale elytral areas; femora pale tan; pronotum (
<figureCitation id="74DF4E85FF9AFFADD10EDBFDFD536395" box="[570,665,922,946]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="21" pageNumber="472">Fig. 13E</figureCitation>
) widest near mid-length and posterior to insertion of midlateral seta, not or only very slightly narrowed basally (ratio PWB/PWA = 1.15 to 1.25), lateral margins slightly and evenly convex or straight or very slightly sinuate anterior to basal angles; elytra slightly to distinctly truncate apically; front tarsomeres 1 to 3 without lateral expansions (
<figureCitation id="74DF4E85FF9AFFADD3F6DC7BFEE76413" box="[194,301,1052,1076]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="21" pageNumber="472">Figs. 14E</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD20FDC7BFEA36413" box="[315,361,1052,1076]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="21" pageNumber="472">15E</figureCitation>
); male with middle mesotarsomeres (
<figureCitation id="74DF4E85FF9AFFADD037DC7BFCA96413" box="[771,867,1052,1076]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="21" pageNumber="472">Fig. 16E</figureCitation>
) only slightly wider than in female and with only tarsomeres 1 to 3 with pads of adhesive setae ventrally (i.e., tarsomere 4 without adhesive setae ventrally) (
<figureCitation id="74DF4E85FF9AFFADD2F4DC3AFDEB6452" box="[448,545,1117,1141]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="21" pageNumber="472">Fig. 17E</figureCitation>
); median lobe of male genitalia (
<figureCitation id="74DF4E85FF9AFFADD0BDDC3AFBC46452" box="[905,1038,1117,1141]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="21" pageNumber="472">Figs. 19C,D</figureCitation>
) with apical lamella short and broad, internal sac without spines.
</paragraph>
<paragraph id="EC5B5200FF9AFFAFD39FDCF8FED16193" blockId="21.[123,1174,695,1725]" lastBlockId="23.[123,1174,219,1725]" lastPageId="23" lastPageNumber="474" pageId="21" pageNumber="472">
In the study area, specimens of
<taxonomicName id="2BE42983FF9AFFADD2CEDCF8FDB16491" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[506,635,1183,1206]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF9AFFADD2CEDCF8FDB16491" box="[506,635,1183,1206]" italics="true" pageId="21" pageNumber="472">T. punctatus</emphasis>
</taxonomicName>
(
<figureCitation id="74DF4E85FF9AFFADD1BDDCF9FD206491" box="[649,746,1182,1206]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="21" pageNumber="472">Fig. 12A</figureCitation>
) might be confused with those of
<taxonomicName id="2BE42983FF9AFFADD760DCF8FF6764F1" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9AFFADD760DCF8FF6764F1" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
</taxonomicName>
(
<figureCitation id="74DF4E85FF9AFFADD389DCD9FED564F1" box="[189,287,1214,1238]" captionStart="FIGURE" captionStartId="16.[155,167,1603,1622]" captionTargetBox="[120,1176,216,1581]" captionTargetId="figure-60@16.[120,1176,216,1590]" captionTargetPageId="16" captionText="FIGURE 10. Tetragonoderus elegans Andrewes. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039409, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality records (black circles) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067305" httpUri="https://zenodo.org/record/11067305/files/figure.png" pageId="21" pageNumber="472">Fig. 10A</figureCitation>
) and also with those of
<taxonomicName id="2BE42983FF9AFFADD112DCD8FD0E64F1" authorityName="Jian &amp; Tian" authorityYear="2009" box="[550,708,1214,1238]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF9AFFADD112DCD8FD0E64F1" box="[550,708,1214,1238]" italics="true" pageId="21" pageNumber="472">T. microthorax</emphasis>
</taxonomicName>
(
<figureCitation id="74DF4E85FF9AFFADD1E0DCD9FCF164F1" box="[724,827,1214,1238]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="21" pageNumber="472">Figs. 13F</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD07CDCD9FCB864F1" box="[840,882,1214,1238]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="21" pageNumber="472">14F</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD04ADCD9FC6D64F1" box="[894,935,1214,1238]" captionStart="FIGURE" captionStartId="32.[155,167,1524,1543]" captionTargetBox="[279,1026,226,1510]" captionTargetId="figure-11@32.[269,1026,216,1510]" captionTargetPageId="32" captionText="FIGURE 15. Male front tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067321" httpUri="https://zenodo.org/record/11067321/files/figure.png" pageId="21" pageNumber="472">15F</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD080DCD9FC1764F1" box="[948,989,1214,1238]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="21" pageNumber="472">16F</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD0DEDCD9FBD964F1" box="[1002,1043,1214,1238]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="21" pageNumber="472">17F</figureCitation>
,
<figureCitation id="74DF4E85FF9AFFADD714DCD9FBF364F1" box="[1056,1081,1214,1238]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="21" pageNumber="472">19</figureCitation>
E-F, and 20), although the latter has not yet been recorded from the Gaoligong
<collectingRegion id="2E209CE2FF9AFFADD055DCB8FC2E64D0" box="[865,996,1247,1271]" country="Myanmar" name="Shan" pageId="21" pageNumber="472">Shan region</collectingRegion>
.
<taxonomicName id="2BE42983FF9AFFADD0D9DCB8FF296530" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF9AFFADD0D9DCB8FF296530" italics="true" pageId="21" pageNumber="472">Tetragonoderus punctatus</emphasis>
</taxonomicName>
members differ from those of
<taxonomicName id="2BE42983FF9AFFADD11ADD67FD506530" box="[558,666,1279,1303]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9AFFADD11ADD67FD506530" box="[558,666,1279,1303]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
</taxonomicName>
in having the forebody slightly lighter in color (i.e., less heavily pigmented) and the elytral pattern of dark areas less extensive, with connection between medial portion of the basal dark band and humeral portion absent or very narrow, less than one-fifth the length of the humeral portion (at least one-fourth the length of the humeral portion in
<taxonomicName id="2BE42983FF9AFFADD34FDDE5FF2D65BE" box="[123,231,1409,1433]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9AFFADD34FDDE5FF2D65BE" box="[123,231,1409,1433]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
</taxonomicName>
), the middle discal band with the area between anterior and posterior dark edges not or only very slightly darker than remainder of pale elytral areas (distinctly darker tan-orange in
<taxonomicName id="2BE42983FF9AFFADD34FDDA4FF2265FD" box="[123,232,1474,1498]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9AFFADD34FDDA4FF2265FD" box="[123,232,1474,1498]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
</taxonomicName>
), and with a less distinct and discontinuous preapical dark band. Also the pronotum is proportionally slightly wider (ratio PWM/EW = 0.55 to 0.57; in
<taxonomicName id="2BE42983FF9AFFADD01BDD84FC5165DD" box="[815,923,1506,1530]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9AFFADD01BDD84FC5165DD" box="[815,923,1506,1530]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
</taxonomicName>
= 0.51 to 0.55), widest at or near midlength, posterior to the insertion of the midlateral seta (widest more anteriad and at or near the insertion of the midlateral seta in
<taxonomicName id="2BE42983FF9AFFADD147DE43FD2B661C" box="[627,737,1571,1595]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9AFFADD147DE43FD2B661C" box="[627,737,1571,1595]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
</taxonomicName>
), less distinctly narrowed basally (ratio PWM/PWB = 1.06 to 1.13; in
<taxonomicName id="2BE42983FF9AFFADD2FDDE22FDFC667B" box="[457,566,1604,1628]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9AFFADD2FDDE22FDFC667B" box="[457,566,1604,1628]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
</taxonomicName>
= 1.13 to 1.20), and the lateral margins are not at all or only very slightly sinuate anterior to the basal angles (distinctly sinuate in
<taxonomicName id="2BE42983FF9AFFADD0FDDE02FBF3665B" box="[969,1081,1636,1660]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF9AFFADD0FDDE02FBF3665B" box="[969,1081,1636,1660]" italics="true" pageId="21" pageNumber="472">T. elegans</emphasis>
</taxonomicName>
). Males differ in several additional features. In
<taxonomicName id="2BE42983FF9AFFADD12ADEE1FD6B66BA" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[542,673,1670,1693]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF9AFFADD12ADEE1FD6B66BA" box="[542,673,1670,1693]" italics="true" pageId="21" pageNumber="472">T. punctatus</emphasis>
</taxonomicName>
males, the middle tarsi are only slightly wider than those in females and middle tarsomere 4 lacks a pad of adhesive setae ventrally (
<figureCitation id="74DF4E85FF9AFFADD712DEC2FB5B669A" box="[1062,1169,1701,1725]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="21" pageNumber="472">Figs. 16E</figureCitation>
,
<figureCitation id="74DF4E85FF98FFAFD34FD8BCFF6360D4" box="[123,169,219,243]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="23" pageNumber="474">17E</figureCitation>
), whereas middle tarsomeres 1 to 4 are distinctly wider than in females and bear adhesive setae ventrally in
<taxonomicName id="2BE42983FF98FFAFD3CFD89BFEAC6134" box="[251,358,251,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF98FFAFD3CFD89BFEAC6134" box="[251,358,251,275]" italics="true" pageId="23" pageNumber="474">T. elegans</emphasis>
</taxonomicName>
males (
<figureCitation id="74DF4E85FF98FFAFD28ED89CFDEE6134" box="[442,548,251,275]" captionStart="FIGURE" captionStartId="33.[155,167,1521,1540]" captionTargetBox="[215,1075,216,1495]" captionTargetId="figure-133@33.[215,1081,216,1509]" captionTargetPageId="33" captionText="FIGURE 16. Male middle tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067325" httpUri="https://zenodo.org/record/11067325/files/figure.png" pageId="23" pageNumber="474">Figs. 16C</figureCitation>
,
<figureCitation id="74DF4E85FF98FFAFD11BD89CFD956134" box="[559,607,251,275]" captionStart="FIGURE" captionStartId="34.[155,167,1521,1540]" captionTargetBox="[264,1031,216,1505]" captionTargetId="figure-129@34.[244,1053,216,1509]" captionTargetPageId="34" captionText="FIGURE 17. Male middle tarsus, ventral aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT103499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067327" httpUri="https://zenodo.org/record/11067327/files/figure.png" pageId="23" pageNumber="474">17C</figureCitation>
). Males also differ in the form of their genitalia, with the median lobe (
<figureCitation id="74DF4E85FF98FFAFD200D97CFE7D6114" box="[308,439,283,307]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="23" pageNumber="474">Figs. 19C,D</figureCitation>
) slightly shorter and distinctly thicker, the apical lamella shorter and broader, and the internal sac without distinct spines in
<taxonomicName id="2BE42983FF98FFAFD1F4D95BFC886174" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[704,834,316,339]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF98FFAFD1F4D95BFC886174" box="[704,834,316,339]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
</taxonomicName>
. In contrast, the median lobe of
<taxonomicName id="2BE42983FF98FFAFD34FD93AFF206153" box="[123,234,348,372]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF98FFAFD34FD93AFF206153" box="[123,234,348,372]" italics="true" pageId="23" pageNumber="474">T. elegans</emphasis>
</taxonomicName>
males (
<figureCitation id="74DF4E85FF98FFAFD272D93BFE076153" box="[326,461,348,372]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="23" pageNumber="474">Figs. 19A,B</figureCitation>
) is longer and more slender, the apical lamella is much longer, narrower, and distinctly curved ventrad, and the internal sac is armed with five or more large and distinct spines.
</paragraph>
<caption id="B89B0288FF99FFAED3AFDE54FBC8665F" ID-DOI="http://doi.org/10.5281/zenodo.11067313" ID-Zenodo-Dep="11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="22" pageNumber="473" startId="22.[155,167,1587,1606]" targetBox="[128,1176,216,1576]" targetPageId="22" targetType="figure">
<paragraph id="EC5B5200FF99FFAED3AFDE54FBC8665F" blockId="22.[123,1174,1586,1657]" pageId="22" pageNumber="473">
<smallCapsWord id="EABDC4DCFF99FFAED3AFDE54FF126662" baselines="1601,1601" box="[155,216,1587,1606]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="22" pageNumber="473">FIGURE</smallCapsWord>
12.
<taxonomicName id="2BE42983FF99FFAED23CDE55FDA56662" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[264,623,1586,1605]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF99FFAED23CDE55FE266662" box="[264,492,1586,1605]" italics="true" pageId="22" pageNumber="473">Tetragonoderus punctatus</emphasis>
(Wiedemann)
</taxonomicName>
. A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<paragraph id="EC5B5200FF98FFAFD39FD9DBFE0663DF" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">
Members of
<taxonomicName id="2BE42983FF98FFAFD209D9DAFE0961F3" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[317,451,445,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF98FFAFD209D9DAFE0961F3" box="[317,451,445,468]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
</taxonomicName>
are extremely similar to those of
<taxonomicName id="2BE42983FF98FFAFD078D9DAFC2461F3" authorityName="Jian &amp; Tian" authorityYear="2009" box="[844,1006,444,468]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF98FFAFD078D9DAFC2461F3" box="[844,1006,444,468]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
</taxonomicName>
and these two species are no doubt confused in collections. Most specimens of
<taxonomicName id="2BE42983FF98FFAFD011D9BAFC6C61D3" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[805,934,477,500]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF98FFAFD011D9BAFC6C61D3" box="[805,934,477,500]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
</taxonomicName>
are slightly larger (BL males =
<quantity id="2B1CFFE5FF98FFAFD3E3D99AFEBA6233" box="[215,368,509,533]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="23" pageNumber="474" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.5 to 4.8 mm</quantity>
, females =
<quantity id="2B1CFFE5FF98FFAFD2D8D99AFD4F6232" box="[492,645,509,533]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.4" metricValueMin="4.6" pageId="23" pageNumber="474" unit="mm" value="5.0" valueMax="5.4" valueMin="4.6">4.6 to 5.4 mm</quantity>
) than those of
<taxonomicName id="2BE42983FF98FFAFD012D999FC0E6232" authorityName="Jian &amp; Tian" authorityYear="2009" box="[806,964,509,533]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF98FFAFD012D999FC0E6232" box="[806,964,509,533]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
</taxonomicName>
(BL males =
<quantity id="2B1CFFE5FF98FFAFD76CD99AFF1A6212" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.6" metricValueMin="3.9" pageId="23" pageNumber="474" unit="mm" value="4.25" valueMax="4.6" valueMin="3.9">3.9 to 4.6 mm</quantity>
, females =
<quantity id="2B1CFFE5FF98FFAFD279DA7AFE236212" box="[333,489,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.7" metricValueMin="4.2" pageId="23" pageNumber="474" unit="mm" value="4.45" valueMax="4.7" valueMin="4.2">4.2 to 4.7 mm</quantity>
). They have the pronotum (
<figureCitation id="74DF4E85FF98FFAFD027DA7AFCBB6212" box="[787,881,541,565]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="23" pageNumber="474">Fig. 13E</figureCitation>
) slightly shorter and wider (ratio PWM/PL = 1.52 to 1.70), widest at or near midlength and posterior to insertion of midlateral seta and not or only very slightly narrowed basally (ratio PWB/PWA = 1.15 to 1.25), and with the lateral margins slightly and evenly convex or straight or very slightly sinuate anterior to the basal angles. In
<taxonomicName id="2BE42983FF98FFAFD206DAF8FE196291" authorityName="Jian &amp; Tian" authorityYear="2009" box="[306,467,670,694]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF98FFAFD206DAF8FE196291" box="[306,467,670,694]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
</taxonomicName>
, the pronotum (
<figureCitation id="74DF4E85FF98FFAFD1BDDAF9FD236291" box="[649,745,670,694]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="23" pageNumber="474">Fig. 13F</figureCitation>
) is slightly longer and narrower (ratio PWM/PL = 1.48 to 1.57), widest anterior to mid-length at or near insertion of midlateral seta, slightly to moderately narrowed basally (ratio PWB/PWA = 1.04 to 1.14), and with the lateral margins slightly to moderately sinuate anterior to the basal angles. Unfortunately, the ranges in size and pronotal proportions (except for ratio PWB/PWA) seen among specimens of these two species overlap slightly, so measurements alone cannot distinguish all specimens of the two. The male genitalia are also very similar, except that the apical lamella of the median lobe is slightly shorter and broader in
<taxonomicName id="2BE42983FF98FFAFD215DBE7FE6F63B0" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[289,421,896,919]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF98FFAFD215DBE7FE6F63B0" box="[289,421,896,919]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
</taxonomicName>
males (
<figureCitation id="74DF4E85FF98FFAFD2C8DB18FD4B63B0" box="[508,641,895,919]" captionStart="FIGURE" captionStartId="36.[155,167,1545,1564]" captionTargetBox="[120,1164,216,1527]" captionTargetId="figure-83@36.[120,1176,216,1542]" captionTargetPageId="36" captionText="FIGURE 19. Male genitalia, Tetragonoderus spp. A, B. T. elegans Andrewes (CASENT1039498, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); C, D. T. punctatus (Wiedemann) (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E, F. T. microthorax Jian and Tian (Jamiri-Sessa, Arunachal, India). A, C, E. Dorsal aspect; B, D, F. Left lateral aspect; scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067331" httpUri="https://zenodo.org/record/11067331/files/figure.png" pageId="23" pageNumber="474">Figs. 19C,D</figureCitation>
) than in
<taxonomicName id="2BE42983FF98FFAFD1D2DBE7FC4F63B0" authorityName="Jian &amp; Tian" authorityYear="2009" box="[742,901,895,919]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF98FFAFD1D2DBE7FC4F63B0" box="[742,901,895,919]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
</taxonomicName>
(Figs. E,F) males. There are also subtle differences in the convexity of the elytral intervals (flat in
<taxonomicName id="2BE42983FF98FFAFD0BADBC7FBDB6390" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[910,1041,928,951]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF98FFAFD0BADBC7FBDB6390" box="[910,1041,928,951]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
</taxonomicName>
and slightly convex in
<taxonomicName id="2BE42983FF98FFAFD3D8DBA6FE4063FF" authorityName="Jian &amp; Tian" authorityYear="2009" box="[236,394,960,984]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF98FFAFD3D8DBA6FE4063FF" box="[236,394,960,984]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
</taxonomicName>
) and depth of impression of the striae (moderately deep in
<taxonomicName id="2BE42983FF98FFAFD738DBA6FB4563FF" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[1036,1167,961,984]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF98FFAFD738DBA6FB4563FF" box="[1036,1167,961,984]" italics="true" pageId="23" pageNumber="474">T. punctatus</emphasis>
</taxonomicName>
, even deeper in
<taxonomicName id="2BE42983FF98FFAFD22BDB86FE7763DF" authorityName="Jian &amp; Tian" authorityYear="2009" box="[287,445,992,1016]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF98FFAFD22BDB86FE7763DF" box="[287,445,992,1016]" italics="true" pageId="23" pageNumber="474">T. microthorax</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="EC5B5200FF98FFAFD39FDC66FD8864BE" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">
<emphasis id="DE908E12FF98FFAFD39FDC66FE5D643F" bold="true" box="[171,407,1025,1048]" pageId="23" pageNumber="474">Habitat distribution.</emphasis>
Members of this species were found at night, running on the surface of the open sandy shores of the Longchuan River (
<figureCitation id="74DF4E85FF98FFAFD1B2DC47FD1C641F" box="[646,726,1056,1080]" captionStart="FIGURE" captionStartId="38.[155,167,1623,1642]" captionTargetBox="[136,1161,216,1607]" captionTargetId="figure-10@38.[136,1161,216,1607]" captionTargetPageId="38" captionText="FIGURE 21. Photographs of habitat for cyclosomine species, Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., Tetragonoderus elegans Andrewes, T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., and T. punctatus Wiedemann at Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China. A. lower river bank; B. sandy upper bank with sparse vegetation. Photos by David H. Kavanaugh." figureDoi="http://doi.org/10.5281/zenodo.11067337" httpUri="https://zenodo.org/record/11067337/files/figure.png" pageId="23" pageNumber="474">Fig. 21</figureCitation>
), a medium-sized stream, at two sites (at elevations of 1215 and
<quantity id="2B1CFFE5FF98FFAFD241DC27FE02647F" box="[373,456,1088,1112]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="23" pageNumber="474" unit="m" value="1230.0">1230 m</quantity>
, respectively). They were running on the same beaches with adults of
<taxonomicName id="2BE42983FF98FFAFD3ADDC05FECC645E" box="[153,262,1121,1145]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF98FFAFD3ADDC05FECC645E" box="[153,262,1121,1145]" italics="true" pageId="23" pageNumber="474">T. elegans</emphasis>
</taxonomicName>
, which were much more abundant. They appear to prefer upper beach areas, where zones of slightly moist and dry sand meet.
</paragraph>
<paragraph id="EC5B5200FF98FFAFD39FDCC5FC0E657D" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">
<emphasis id="DE908E12FF98FFAFD39FDCC5FD18649E" bold="true" box="[171,722,1186,1209]" pageId="23" pageNumber="474">Geographical distribution within the Gaoligong</emphasis>
<emphasis id="DE908E12FF98FFAFD1EFDCC5FCD6649E" bold="true" box="[731,796,1186,1209]" pageId="23" pageNumber="474">
<collectingRegion id="2E209CE2FF98FFAFD1EFDCC5FCDD649E" box="[731,791,1186,1209]" country="Myanmar" name="Shan" pageId="23" pageNumber="474">Shan</collectingRegion>
.
</emphasis>
<figureCitation id="74DF4E85FF98FFAFD011DCC6FC42649E" box="[805,904,1185,1209]" captionStart="FIGURE" captionStartId="22.[155,167,1587,1606]" captionTargetBox="[128,1176,216,1576]" captionTargetId="figure-59@22.[120,1176,216,1576]" captionTargetPageId="22" captionText="FIGURE 12. Tetragonoderus punctatus (Wiedemann). A. Dorsal aspect, scale line = 1.0 mm (CASENT1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Map showing locality record (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067313" httpUri="https://zenodo.org/record/11067313/files/figure.png" pageId="23" pageNumber="474">Fig. 12B</figureCitation>
.
<materialsCitation id="5C8C585DFF98FFAFD0A0DCC5FBFB653D" ID-GBIF-Occurrence="4902827310" collectingDate="2003-10-28" collectingDateMax="2006-06-03" collectingDateMin="2003-10-28" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong &amp; Longchuan Jiang &amp; Longjiang Bridge &amp; X. C. Shi" county="Tengchong County" elevation="1230" location="Wuhe Township" municipality="We" pageId="23" pageNumber="474" specimenCount="15" specimenCount-female="8" specimenCount-male="2" stateProvince="Shan">
<collectingMunicipality id="0C3FC87AFF98FFAFD0A0DCC5FC7D649E" box="[916,951,1186,1209]" pageId="23" pageNumber="474">We</collectingMunicipality>
examined a total of
<specimenCount id="FAE29989FF98FFAFD34FDCA6FED764FE" box="[123,285,1217,1241]" count="5" pageId="23" pageNumber="474" type="generic">five specimens</specimenCount>
(
<specimenCount id="FAE29989FF98FFAFD21BDCA6FE5F64FE" box="[303,405,1217,1241]" count="1" pageId="23" pageNumber="474" type="male">one male</specimenCount>
and
<specimenCount id="FAE29989FF98FFAFD2E4DCA6FD9164FE" box="[464,603,1217,1241]" count="4" pageId="23" pageNumber="474" type="female">four females</specimenCount>
) from the following localities:
<emphasis id="DE908E12FF98FFAFD08CDCA5FB5F64FD" bold="true" box="[952,1173,1218,1242]" italics="true" pageId="23" pageNumber="474">
<collectingCounty id="053A2A8CFF98FFAFD08CDCA5FB4464FD" box="[952,1166,1218,1242]" pageId="23" pageNumber="474">Tengchong County</collectingCounty>
:
</emphasis>
<location id="E93B04DBFF98FFAFD34FDC86FEE164DE" LSID="urn:lsid:plazi:treatment:644DE316FF9AFFA0D34FD95BFF6861B4:E93B04DBFF98FFAFD34FDC86FEE164DE" box="[123,299,1249,1273]" county="Tengchong County" municipality="We" name="Wuhe Township" pageId="23" pageNumber="474" stateProvince="Shan">Wuhe Township</location>
(
<location id="E93B04DBFF98FFAFD20FDC86FE3C64DE" LSID="urn:lsid:plazi:treatment:644DE316FF9AFFA0D34FD95BFF6861B4:E93B04DBFF98FFAFD20FDC86FE3C64DE" box="[315,502,1249,1273]" county="Tengchong County" municipality="We" name="Longchuan Jiang" pageId="23" pageNumber="474" stateProvince="Shan">Longchuan Jiang</location>
just below bridge at
<location id="E93B04DBFF98FFAFD1D6DC86FC5764DE" LSID="urn:lsid:plazi:treatment:644DE316FF9AFFA0D34FD95BFF6861B4:E93B04DBFF98FFAFD1D6DC86FC5764DE" box="[738,925,1249,1273]" county="Tengchong County" municipality="We" name="Menglian village" pageId="23" pageNumber="474" stateProvince="Shan">Menglian village</location>
, 24.89176°/98.67551°,
<quantity id="2B1CFFE5FF98FFAFD34FDD65FF04653D" box="[123,206,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="23" pageNumber="474" unit="m" value="1230.0">
<elevation id="67C9B533FF98FFAFD34FDD65FF04653D" box="[123,206,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="23" pageNumber="474" unit="m" value="1230.0">1230 m</elevation>
</quantity>
,
<date id="985A74C0FF98FFAFD3EEDD65FE95653D" box="[218,351,1282,1306]" pageId="23" pageNumber="474" value="2006-06-03">
<collectingDate id="881E8D28FF98FFAFD3EEDD65FE95653D" box="[218,351,1282,1306]" pageId="23" pageNumber="474" value="2006-06-03">3 June 2006</collectingDate>
</date>
,
<collectorName id="411137D6FF98FFAFD25EDD64FDEB653D" box="[362,545,1282,1306]" pageId="23" pageNumber="474">D.H. Kavanaugh</collectorName>
,
<collectionCode id="8AF5CAC5FF98FFAFD118DD64FD8B653D" box="[556,577,1283,1306]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="474">R</collectionCode>
.
<collectorName id="411137D6FF98FFAFD171DD64FD51653D" box="[581,667,1283,1306]" pageId="23" pageNumber="474">L. Brett</collectorName>
,
<collectorName id="411137D6FF98FFAFD192DD64FCD4653D" box="[678,798,1282,1306]" pageId="23" pageNumber="474">H.B. Liang</collectorName>
&amp;
<collectorName id="411137D6FF98FFAFD075DD64FC7E653C" box="[833,948,1283,1307]" pageId="23" pageNumber="474">D.Z. Dong</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF98FFAFD705DD64FCDF657D" ID-GBIF-Occurrence="4902827331" collectingDate="2003-10-28" collectingDateMax="2006-06-03" collectingDateMin="2003-10-28" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong &amp; Longchuan Jiang &amp; Longjiang Bridge &amp; X. C. Shi" county="Tengchong County" elevation="1230" location="Wuhe Township" municipality="We" pageId="23" pageNumber="474" specimenCount="15" specimenCount-female="8" specimenCount-male="2" stateProvince="Shan">
<specimenCount id="FAE29989FF98FFAFD705DD64FB5C653D" box="[1073,1174,1282,1306]" count="1" pageId="23" pageNumber="474" type="male">one male</specimenCount>
and
<specimenCount id="FAE29989FF98FFAFD398DD45FE89651D" box="[172,323,1314,1338]" count="3" pageId="23" pageNumber="474" type="female">three females</specimenCount>
;
<collectionCode id="8AF5CAC5FF98FFAFD267DD45FE43651D" box="[339,393,1314,1338]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="23" pageNumber="474" type="Museum">CAS</collectionCode>
,
<collectionCode id="8AF5CAC5FF98FFAFD2ACDD45FE00651D" box="[408,458,1314,1338]" pageId="23" pageNumber="474">IOZ</collectionCode>
]), (
<collectorName id="411137D6FF98FFAFD2C4DD45FD67651D" box="[496,685,1314,1338]" pageId="23" pageNumber="474">Longchuan Jiang</collectorName>
at
<collectorName id="411137D6FF98FFAFD1E3DD45FC50651D" box="[727,922,1314,1338]" pageId="23" pageNumber="474">Longjiang Bridge</collectorName>
, 24.89889°/98.66667°,
<quantity id="2B1CFFE5FF98FFAFD34FDD25FF04657D" box="[123,206,1346,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="23" pageNumber="474" unit="m" value="1215.0">
<elevation id="67C9B533FF98FFAFD34FDD25FF04657D" box="[123,206,1346,1370]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="23" pageNumber="474" unit="m" value="1215.0">1215 m</elevation>
</quantity>
,
<date id="985A74C0FF98FFAFD3EEDD25FE5B657D" box="[218,401,1346,1370]" pageId="23" pageNumber="474" value="2003-10-28">
<collectingDate id="881E8D28FF98FFAFD3EEDD25FE5B657D" box="[218,401,1346,1370]" pageId="23" pageNumber="474" value="2003-10-28">28 October 2003</collectingDate>
</date>
,
<collectorName id="411137D6FF98FFAFD2A9DD24FDDE657D" box="[413,532,1346,1370]" pageId="23" pageNumber="474">H.B. Liang</collectorName>
&amp;
<collectorName id="411137D6FF98FFAFD10CDD25FD5C657D" box="[568,662,1346,1370]" pageId="23" pageNumber="474">X.C. Shi</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF98FFAFD021DD25FC0E657D" ID-GBIF-Occurrence="4902827315" box="[789,964,1346,1370]" collectingDate="2003-10-28" collectingDateMax="2006-06-03" collectingDateMin="2003-10-28" collectionCode="R, CAS, IOZ" collectorName="D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong &amp; Longchuan Jiang &amp; Longjiang Bridge &amp; X. C. Shi" county="Tengchong County" elevation="1230" location="Wuhe Township" municipality="We" pageId="23" pageNumber="474" specimenCount="15" specimenCount-female="8" specimenCount-male="2" stateProvince="Shan">
<specimenCount id="FAE29989FF98FFAFD021DD25FCBF657D" box="[789,885,1346,1370]" count="1" pageId="23" pageNumber="474" type="female">1 female</specimenCount>
;
<collectionCode id="8AF5CAC5FF98FFAFD0B4DD25FC73657D" box="[896,953,1346,1370]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="23" pageNumber="474" type="Museum">CAS</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="EC5B5200FF98FFAFD39FDD05FEDF65BC" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">At present, this species has been recorded only from the southwestern part of the study area (Core Area 6).</paragraph>
<paragraph id="EC5B5200FF98FFAFD39FDDC3FE7B665B" blockId="23.[123,1174,219,1725]" pageId="23" pageNumber="474">
<emphasis id="DE908E12FF98FFAFD39FDDC3FDE7659C" bold="true" box="[171,557,1444,1467]" pageId="23" pageNumber="474">Overall geographical distribution.</emphasis>
<figureCitation id="74DF4E85FF98FFAFD106DDC4FD4A659C" box="[562,640,1443,1467]" captionStart="FIGURE" captionStartId="40.[155,167,997,1016]" captionTargetBox="[120,1176,216,987]" captionTargetId="figure-10@40.[120,1176,216,987]" captionTargetPageId="40" captionText="FIGURE 23. Map showing approximate known overall geographical distributions of cyclosomine species occurring in the Gaoligong Shan. a (blue) = Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; b (yellow) = Tetragonoderus arcuatus Dejean; c (red) = T. elegans Andrewes; d (white dot) = T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov.; e (orange) = T. punctatus Wiedemann. Modified from Wikimedia Commons, World Atlas of the World, at URL: http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11067341" httpUri="https://zenodo.org/record/11067341/files/figure.png" pageId="23" pageNumber="474">Fig. 23</figureCitation>
. We have examined specimens of this species only from “Bengala” (i.e.,
<collectingRegion id="2E209CE2FF98FFAFD251DDA3FE2465FC" box="[357,494,1475,1499]" country="India" name="West Bengal" pageId="23" pageNumber="474">West Bengal</collectingRegion>
(
<collectingCountry id="94F31290FF98FFAFD2C9DDA4FDFD65FC" box="[509,567,1475,1499]" name="India" pageId="23" pageNumber="474">India</collectingCountry>
) and/or
<collectingCountry id="94F31290FF98FFAFD1A6DDA4FCDA65FC" box="[658,784,1475,1499]" name="Bangladesh" pageId="23" pageNumber="474">Bangladesh</collectingCountry>
) and the localities in the Gaoligong Shan as listed above. Its occurrence in the study area represents the eastern limit of its known geographical range and also a first record for
<collectingCountry id="94F31290FF98FFAFD14CDE64FD71663C" box="[632,699,1539,1563]" name="China" pageId="23" pageNumber="474">China</collectingCountry>
. It is likely that this species occurs also in northern
<collectingCountry id="94F31290FF98FFAFD3EFDE43FE89661B" box="[219,323,1572,1596]" name="Myanmar" pageId="23" pageNumber="474">Myanmar</collectingCountry>
and northeastern
<collectingCountry id="94F31290FF98FFAFD135DE43FDF2661B" box="[513,568,1572,1596]" name="India" pageId="23" pageNumber="474">India</collectingCountry>
along rivers draining the western slope of the Gaoligong Shan and the southern slope of the eastern part of the Himalaya Mountains, respectively, at low elevations (
<quantity id="2B1CFFE5FF98FFAFD3C2DE03FE6E665B" box="[246,420,1636,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="23" pageNumber="474" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">1000 to 1300 m</quantity>
).
</paragraph>
<paragraph id="EC5B5200FF98FFA0D39FDEE2FF6861B4" blockId="23.[123,1174,219,1725]" lastBlockId="24.[123,1174,219,403]" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474">
<emphasis id="DE908E12FF98FFAFD39FDEE2FCB766BB" bold="true" box="[171,893,1669,1693]" pageId="23" pageNumber="474">
Geographical relationships with other
<taxonomicName id="2BE42983FF98FFAFD144DEE2FCB766BB" box="[624,893,1669,1693]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="DE908E12FF98FFAFD144DEE2FCD666BA" bold="true" box="[624,796,1669,1693]" italics="true" pageId="23" pageNumber="474">Tetragonoderus</emphasis>
species.
</taxonomicName>
</emphasis>
Members of this species have been found synotopic with those of
<taxonomicName id="2BE42983FF98FFAFD102DEC1FD68669B" box="[566,674,1700,1725]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DE908E12FF98FFAFD102DEC1FD68669B" box="[566,674,1700,1725]" italics="true" pageId="23" pageNumber="474">T. elegans</emphasis>
</taxonomicName>
and
<taxonomicName id="2BE42983FF98FFAFD1ECDEC1FCAE669B" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[728,868,1700,1725]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="parviculus">
<emphasis id="DE908E12FF98FFAFD1ECDEC1FCAE669B" box="[728,868,1700,1725]" italics="true" pageId="23" pageNumber="474">T. parviculus</emphasis>
</taxonomicName>
in the study area. The range of this species also overlaps that of
<taxonomicName id="2BE42983FF97FFA0D2C0D8BBFDA060D4" box="[500,618,220,243]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="arcuatus">
<emphasis id="DE908E12FF97FFA0D2C0D8BBFDA060D4" box="[500,618,220,243]" italics="true" pageId="24" pageNumber="475">T. arcuatus</emphasis>
</taxonomicName>
in the study area, but these two species have not yet been found syntopic. Also, the range of
<taxonomicName id="2BE42983FF97FFA0D11BD89BFD046134" authorityName="Jian &amp; Tian" authorityYear="2009" box="[559,718,251,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF97FFA0D11BD89BFD046134" box="[559,718,251,275]" italics="true" pageId="24" pageNumber="475">T. microthorax</emphasis>
</taxonomicName>
broadly overlaps that of
<taxonomicName id="2BE42983FF97FFA0D0D5D89BFBAF6134" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[993,1125,252,275]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="DE908E12FF97FFA0D0D5D89BFBAF6134" box="[993,1125,252,275]" italics="true" pageId="24" pageNumber="475">T. punctatus</emphasis>
</taxonomicName>
, but the former species has not yet been recorded from the study area. We note that all current records of
<taxonomicName id="2BE42983FF97FFA0D3A3D95BFEFE6174" authorityName="Jian &amp; Tian" authorityYear="2009" box="[151,308,315,339]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="microthorax">
<emphasis id="DE908E12FF97FFA0D3A3D95BFEFE6174" box="[151,308,315,339]" italics="true" pageId="24" pageNumber="475">T. microthorax</emphasis>
</taxonomicName>
are from localities well below
<quantity id="2B1CFFE5FF97FFA0D148D95CFD066174" box="[636,716,315,339]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="475" unit="m" value="1000.0">1000 m</quantity>
in elevation and such elevations are absent from all but the extreme southwesternmost part of the study area, which we were unable to sample.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,57 +1,59 @@
<document id="11854F9058697710D7FA2CFC7443FEEB" ID-DOI="10.5281/zenodo.11067293" ID-ISSN="0068-547X" ID-Zenodo-Dep="11067293" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714052207303" checkinUser="felipe" docAuthor="Kavanaugh, David H., Cueva-Dabkoski, Mollie &amp; Liang, Hongbin" docDate="2023" docId="644DE316FF9EFFADD34FDDFBFD386134" docLanguage="en" docName="ProcCalifAcadSci.67.20.451-491.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (20)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.1:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="1" docTitle="Tetragonoderus parviculus Kavanaugh and Cueva-Dabkoski 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="471" masterDocId="98749B6EFF8FFFB8D334D867FFCA6027" masterDocTitle="Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species." masterLastPageNumber="491" masterPageNumber="451" pageNumber="468" updateTime="1714052554343" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9590F57DCB5421D27FC6983B0F5E9788">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:title id="A5FE200E2338CAFA134A44488332745D">Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species.</mods:title>
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<mods:namePart id="B27B9065DD870765E7620EA1FCCB11E8">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="EFDBB55316347643A939C25167C98927">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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<mods:namePart id="6B1FD4FB801824BB3A02A3930F9EC1CC">Cueva-Dabkoski, Mollie</mods:namePart>
<mods:affiliation id="EDC3D3C964B615D3C87717219655A572">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
<mods:namePart id="0ED7FAC662408F1528C782B9C364FF41">Cueva-Dabkoski, Mollie</mods:namePart>
<mods:affiliation id="8914056A894575969A8F52DF44FF6B5D">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:namePart id="DC40050E1B827384817D4AF4151CAE40">Liang, Hongbin</mods:namePart>
<mods:affiliation id="1641921E4A37E8F033A1DC7D2FAFDC48">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
<mods:namePart id="538C3B34B354F92C1571845879915AC1">Liang, Hongbin</mods:namePart>
<mods:affiliation id="ADC66B4230D8195AA13A4B8EF491FC18">Key Laboratory of Zoological Systematics, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
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@ -85,13 +87,17 @@ C-D, 21, and 23.
</paragraph>
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<emphasis id="DE908E12FF9EFFA9D39FDD81FE9365DA" bold="true" box="[171,345,1510,1533]" pageId="17" pageNumber="468">Type material.</emphasis>
<materialsCitation id="5C8C585DFF9EFFA9D25CDD82FB5C669A" collectingDate="2006-06-03" collectionCode="IOZ, R, CAS" collectorName="L. Brett &amp; D. Z. Dong &amp; D. H. Kavanaugh &amp; M. Cueva-Dabkoski" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="7" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
<materialsCitation id="5C8C585DFF9EFFA9D25CDD82FCFD665A" ID-GBIF-Occurrence="4902827307" collectingDate="2006-06-03" collectionCode="IOZ" collectorName="D. H. Kavanaugh &amp; R. L. Brett &amp; D. Z. Dong" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCode="CASENT1039396" specimenCount="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
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, a male, deposited in
, a
<specimenCount id="FAE29989FF9EFFA9D113DD82FD9765DA" box="[551,605,1509,1533]" pageId="17" pageNumber="468" type="male">male</specimenCount>
, deposited in
<collectionCode id="8AF5CAC5FF9EFFA9D03FDD82FCF065DA" box="[779,826,1509,1533]" pageId="17" pageNumber="468">IOZ</collectionCode>
, labeled: “CASENT1039396”/ “
, labeled: “
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”/ “
<collectingCountry id="94F31290FF9EFFA9D3B0DE62FF17663A" box="[132,221,1541,1565]" name="China" pageId="17" pageNumber="468">CHINA</collectingCountry>
,
<collectingRegion id="2E209CE2FF9EFFA9D3DFDE61FEF5663A" box="[235,319,1542,1565]" country="China" name="Yunnan" pageId="17" pageNumber="468">Yunnan</collectingRegion>
@ -101,9 +107,7 @@ C-D, 21, and 23.
<collectingMunicipality id="0C3FC87AFF9EFFA9D119DE62FD28663A" box="[557,738,1541,1565]" pageId="17" pageNumber="468">Wuhe Township</collectingMunicipality>
,
<location id="E93B04DBFF9EFFA9D1C4DE62FC61663A" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9EFFA9D1C4DE62FC61663A" box="[752,939,1541,1565]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longchuan Jiang" pageId="17" pageNumber="468" stateProvince="Yunnan">Longchuan Jiang</location>
just below bridge at
<location id="E93B04DBFF9EFFA9D34FDE42FEFE661A" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9EFFA9D34FDE42FEFE661A" box="[123,308,1573,1597]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Menglian village" pageId="17" pageNumber="468" stateProvince="Yunnan">Menglian village</location>
, 24.89176°/98.67551°,”/ “
just below bridge at Menglian village, 24.89176°/98.67551°,”/ “
<quantity id="2B1CFFE5FF9EFFA9D166DE42FD62661A" box="[594,680,1573,1597]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="468" unit="m" value="1230.0">
<elevation id="67C9B533FF9EFFA9D166DE42FD62661A" box="[594,680,1573,1597]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="17" pageNumber="468" unit="m" value="1230.0">1230 m</elevation>
</quantity>
@ -111,43 +115,55 @@ just below bridge at
<date id="985A74C0FF9EFFA9D181DE42FCF1661A" box="[693,827,1573,1597]" pageId="17" pageNumber="468" value="2006-06-03">
<collectingDate id="881E8D28FF9EFFA9D181DE42FCF1661A" box="[693,827,1573,1597]" pageId="17" pageNumber="468" value="2006-06-03">3 June 2006</collectingDate>
</date>
, Stop # DHK-2006-054A, D.H. Kavanaugh,
<collectionCode id="8AF5CAC5FF9EFFA9D231DE21FED0667A" box="[261,282,1606,1629]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="468">R</collectionCode>
.
<collectorName id="411137D6FF9EFFA9D22ADE21FEBF667A" box="[286,373,1606,1629]" pageId="17" pageNumber="468">L. Brett</collectorName>
, Stop # DHK-2006-054A,
<collectorName id="411137D6FF9EFFA9D756DE41FF3D667A" pageId="17" pageNumber="468">D.H. Kavanaugh</collectorName>
,
<collectorName id="411137D6FF9EFFA9D231DE21FEBF667A" box="[261,373,1606,1629]" pageId="17" pageNumber="468">R.L. Brett</collectorName>
&amp;
<collectorName id="411137D6FF9EFFA9D2ABDE21FDDF6679" box="[415,533,1606,1630]" pageId="17" pageNumber="468">D.Z. Dong</collectorName>
collectors”/ “
<typeStatus id="335FECA2FF9EFFA9D19BDE22FCF4667A" box="[687,830,1605,1629]" pageId="17" pageNumber="468" type="holotype">HOLOTYPE</typeStatus>
collectors”/ “HOLOTYPE
<taxonomicName id="2BE42983FF9EFFA9D07DDE22FBA7667A" authority="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[841,1133,1605,1629]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="parviculus" status="sp. nov.">
<emphasis id="DE908E12FF9EFFA9D07DDE22FBA7667A" box="[841,1133,1605,1629]" italics="true" pageId="17" pageNumber="468">Tetragonoderus parviculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5A33369FF9EFFA9D743DE21FF62665A" pageId="17" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
<collectorName id="411137D6FF9EFFA9D385DE01FEAD665A" box="[177,359,1637,1661]" pageId="17" pageNumber="468">D.H. Kavanaugh</collectorName>
&amp;
<collectorName id="411137D6FF9EFFA9D2A4DE01FDA1665A" box="[400,619,1637,1661]" pageId="17" pageNumber="468">M. Cueva-Dabkoski</collectorName>
2022” [red label]. A total of
<specimenCount id="FAE29989FF9EFFA9D087DE02FB996659" box="[947,1107,1637,1662]" count="4" pageId="17" pageNumber="468" type="generic" typeStatus="paratype">four paratypes</specimenCount>
:
D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [red label].
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A total of four paratypes:
<specimenCount id="FAE29989FF9EFFA9D754DE02FF7666BA" count="3" pageId="17" pageNumber="468" type="male">three males</specimenCount>
, in
<collectionCode id="8AF5CAC5FF9EFFA9D3D5DEE2FEDF66BA" box="[225,277,1669,1693]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="468" type="Museum">CAS</collectionCode>
<collectionCode id="8AF5CAC5FF9EFFA9D3D5DEE2FEDF66BA" box="[225,277,1669,1693]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="468" type="Museum">CAS</collectionCode>
and
<collectionCode id="8AF5CAC5FF9EFFA9D27DDEE2FEB266BA" box="[329,376,1669,1693]" pageId="17" pageNumber="468">IOZ</collectionCode>
, same label data as holotype, except first label [unique identifier label] different, “CASENT1039394”, “CASENT1039395” and “CASENT1039397”, respectively;
<specimenCount id="FAE29989FF9EFFAAD75BDEC1FF0C60D4" count="1" lastPageId="18" lastPageNumber="469" pageId="17" pageNumber="468" type="female">
one
<paragraph id="EC5B5200FF9DFFAAD34FD8BCFD176153" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">
<materialsCitation id="5C8C585DFF9DFFAAD34FD8BCFD136153" collectingDate="2016-06-25" collectingDateMax="2016-06-27" collectingDateMin="2016-06-25" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Ruzicka &amp; C. - B. Wang &amp; D. H. Kavanaugh &amp; M. Cueva-Dabkoski" country="China" elevation="1290" latitude="24.613" location="Tongbiguan" longLatPrecision="12" longitude="97.75633" pageId="18" pageNumber="469" specimenCount="2" stateProvince="Yunnan" typeStatus="paratype">
, same label data as holotype, except first label [
</materialsCitation>
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unique identifier label] different, “
<specimenCode id="BC42FA7BFF9EFFA9D3EDDEC2FE6F669A" box="[217,421,1701,1725]" pageId="17" pageNumber="468">CASENT1039394</specimenCode>
”,
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D2F6DEC2FD1F669A" ID-GBIF-Occurrence="4902827308" box="[450,725,1701,1725]" collectingDate="2006-06-03" collectionCode="IOZ, R, CAS" collectorName="L. Brett &amp; D. Z. Dong &amp; D. H. Kavanaugh &amp; M. Cueva-Dabkoski" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCode="CASENT1039395" specimenCount="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
<specimenCode id="BC42FA7BFF9EFFA9D2F8DEC2FD5F669A" box="[460,661,1701,1725]" pageId="17" pageNumber="468">CASENT1039395</specimenCode>
” and
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFA9D1D0DEC2FB95669A" ID-GBIF-Occurrence="4902827329" box="[740,1119,1701,1725]" collectingDate="2006-06-03" collectionCode="IOZ, R, CAS" collectorName="L. Brett &amp; D. Z. Dong &amp; D. H. Kavanaugh &amp; M. Cueva-Dabkoski" country="China" county="Tengchong County" elevation="1230" location="Longchuan Jiang" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCode="CASENT1039397" specimenCount="1" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
<specimenCode id="BC42FA7BFF9EFFA9D1DADEC2FC70669A" box="[750,954,1701,1725]" pageId="17" pageNumber="468">CASENT1039397</specimenCode>
”, respectively;
</materialsCitation>
<materialsCitation id="5C8C585DFF9EFFAAD75BDEC1FD136153" ID-GBIF-Occurrence="4902827335" collectingDate="2006-06-03" collectingDateMax="2016-06-27" collectingDateMin="2016-06-25" collectingMethod="baited pitfall trap" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Ruzicka &amp; C. - B. Wang" country="China" county="Tengchong County" elevation="1290" lastPageId="18" lastPageNumber="469" latitude="24.613" location="YUNNAN Prov." longLatPrecision="12" longitude="97.75633" municipality="Wuhe Township" pageId="17" pageNumber="468" specimenCount="1" specimenCount-female="1" specimenCount-male="3" stateProvince="YUNNAN Prov." typeStatus="holotype">
<specimenCount id="FAE29989FF9EFFA9D75BDEC1FB5C669A" box="[1135,1174,1702,1725]" count="1" pageId="17" pageNumber="468" type="female">one</specimenCount>
female, in
<collectionCode id="8AF5CAC5FF9DFFAAD3CFD8BCFE8360D4" box="[251,329,219,243]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="18" pageNumber="469">NMPC</collectionCode>
, labeled: “
<collectingCountry id="94F31290FF9DFFAAD2F3D8BCFDE860D4" box="[455,546,219,243]" name="China" pageId="18" pageNumber="469">CHINA</collectingCountry>
:
<collectingRegion id="2E209CE2FF9DFFAAD106D8BCFD2160D5" box="[562,747,219,243]" country="China" name="Yunnan" pageId="18" pageNumber="469">YUNNAN Prov.</collectingRegion>
<locationDeviation id="FDE94B22FF9DFFAAD1CDD8BCFC3560D4" box="[761,1023,219,243]" pageId="18" pageNumber="469">
<quantity id="2B1CFFE5FF9DFFAAD1CDD8BCFC8060D4" box="[761,842,219,243]" metricMagnitude="3" metricUnit="m" metricValue="5.9" pageId="18" pageNumber="469" unit="km" value="5.9">5.9 km</quantity>
W
<location id="E93B04DBFF9DFFAAD049D8BCFC3560D4" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9DFFAAD049D8BCFC3560D4" box="[893,1023,219,243]" country="China" latitude="24.613" longLatPrecision="12" longitude="97.75633" name="Tongbiguan" pageId="18" pageNumber="469" stateProvince="Yunnan">Tongbiguan</location>
W Tongbiguan
</locationDeviation>
,
<geoCoordinate id="89D034C7FF9DFFAAD725D8BCFB5B60D4" box="[1041,1169,219,243]" degrees="24" direction="north" minutes="36.78" orientation="latitude" pageId="18" pageNumber="469" precision="9" value="24.613">24°36.78N</geoCoordinate>
,
@ -159,35 +175,23 @@ W
, &amp;
<collectorName id="411137D6FF9DFFAAD2C8D89CFDB36133" box="[508,633,251,276]" pageId="18" pageNumber="469">C.-B. Wang</collectorName>
leg.”/ “(CH12)
<date id="985A74C0FF9DFFAAD017D89CFC086134" box="[803,962,251,275]" pageId="18" pageNumber="469" value="2016-06-25" valueMax="2016-06-27" valueMin="2016-06-25">
<collectingDate id="881E8D28FF9DFFAAD017D89CFC086134" box="[803,962,251,275]" pageId="18" pageNumber="469" value="2016-06-25" valueMax="2016-06-27" valueMin="2016-06-25">25.-27.vi.2016</collectingDate>
</date>
<date id="985A74C0FF9DFFAAD017D89CFC086134" box="[803,962,251,275]" pageId="18" pageNumber="469" value="2016-06-25" valueMax="2016-06-27" valueMin="2016-06-25">25.-27.vi.2016</date>
;
<quantity id="2B1CFFE5FF9DFFAAD0F8D89CFBD56134" box="[972,1055,251,275]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="18" pageNumber="469" unit="m" value="1290.0">
<elevation id="67C9B533FF9DFFAAD0F8D89CFBD56134" box="[972,1055,251,275]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="18" pageNumber="469" unit="m" value="1290.0">1290 m</elevation>
</quantity>
,
<collectingMethod id="35A52A17FF9DFFAAD71DD89CFF046114" pageId="18" pageNumber="469">baited pitfall trap</collectingMethod>
#06 (fish meat, ripening cheese), secondary broadleaved forest, valley near stream”. Each
<typeStatus id="335FECA2FF9DFFAAD34FD95BFF1D6173" box="[123,215,316,340]" pageId="18" pageNumber="469" type="paratype">paratype</typeStatus>
also bears the following label: “
<typeStatus id="335FECA2FF9DFFAAD177D95CFD006174" box="[579,714,315,339]" pageId="18" pageNumber="469" type="paratype">PARATYPE</typeStatus>
#06 (fish meat, ripening cheese), secondary broadleaved forest, valley near stream”. Each paratype also bears the following label: “PARATYPE
<taxonomicName id="2BE42983FF9DFFAAD1E1D95CFC326174" authority="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski &amp; Liang" authorityYear="2023" box="[725,1016,315,339]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="parviculus" status="sp. nov.">
<emphasis id="DE908E12FF9DFFAAD1E1D95CFC326174" box="[725,1016,315,339]" italics="true" pageId="18" pageNumber="469">Tetragonoderus parviculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C5A33369FF9DFFAAD736D95BFB926174" box="[1026,1112,316,339]" pageId="18" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
<collectorName id="411137D6FF9DFFAAD756D95BFF3E6153" pageId="18" pageNumber="469">D.H. Kavanaugh</collectorName>
&amp;
<collectorName id="411137D6FF9DFFAAD22CD93BFE3A6153" box="[280,496,348,372]" pageId="18" pageNumber="469">M. Cueva-Dabkoski</collectorName>
2022” [yellow label]
D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [yellow label]
</materialsCitation>
.
</paragraph>
</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="EC5B5200FF9DFFAAD39FD91AFE2D6193" blockId="18.[123,1174,219,1725]" pageId="18" pageNumber="469">
<materialsCitation id="5C8C585DFF9DFFAAD39FD91AFE296193" country="China" county="Tengchong County" location="Longchuan River" municipality="Wuhe Township" pageId="18" pageNumber="469" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="DE908E12FF9DFFAAD39FD91AFE8C61B3" bold="true" box="[171,326,381,404]" pageId="18" pageNumber="469">
<typeStatus id="335FECA2FF9DFFAAD39FD91AFF2961B3" box="[171,227,381,404]" pageId="18" pageNumber="469">Type</typeStatus>
locality.
@ -203,7 +207,6 @@ locality.
<location id="E93B04DBFF9DFFAAD092D91BFBAE61B3" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9DFFAAD092D91BFBAE61B3" box="[934,1124,380,404]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longchuan River" pageId="18" pageNumber="469" stateProvince="Yunnan">Longchuan River</location>
just below bridge at
<location id="E93B04DBFF9DFFAAD21DD9FBFE296193" LSID="urn:lsid:plazi:treatment:644DE316FF9EFFADD34FDDFBFD386134:E93B04DBFF9DFFAAD21DD9FBFE296193" box="[297,483,412,436]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Menglian village" pageId="18" pageNumber="469" stateProvince="Yunnan">Menglian village</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
@ -283,13 +286,7 @@ Color. (
<paragraph id="EC5B5200FF9DFFACD39FDE03FC1A61F3" blockId="18.[123,1174,219,1725]" lastBlockId="20.[123,1174,219,1725]" lastPageId="20" lastPageNumber="471" pageId="18" pageNumber="469">
Prothorax. Pronotum (
<figureCitation id="74DF4E85FF9DFFAAD2AADE03FDCB665B" box="[414,513,1636,1660]" captionStart="FIGURE" captionStartId="30.[155,167,1512,1531]" captionTargetBox="[140,1157,216,1497]" captionTargetId="figure-11@30.[140,1157,216,1497]" captionTargetPageId="30" captionText="FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11067317" httpUri="https://zenodo.org/record/11067317/files/figure.png" pageId="18" pageNumber="469">Fig. 13D</figureCitation>
) slightly narrow (ratio PWM/
<collectionCode id="8AF5CAC5FF9DFFAAD07FDE03FCBE665B" box="[843,884,1636,1660]" pageId="18" pageNumber="469">EW</collectionCode>
= 0.61 to 0.65), distinctly narrowed basally (ratio
<collectionCode id="8AF5CAC5FF9DFFAAD24ADEE3FE7F66BB" box="[382,437,1668,1692]" pageId="18" pageNumber="469">PWB</collectionCode>
/
<collectionCode id="8AF5CAC5FF9DFFAAD288DEE3FE3C66BB" box="[444,502,1668,1692]" pageId="18" pageNumber="469">PWA</collectionCode>
= 1.04 to 1.09), widest distinctly anterior to mid-length, with anterior angles very short and broadly rounded; posterior angles slightly obtuse and slightly projected laterally; apical margin straight or very slightly concave between anterior angles; lateral margins arcuate and with a short and shallow sinuation just anterior to posterior angles; basal margin faintly bisinuate, not or only slightly lobate medially; anterior margination distinct laterally, more or less effaced medially; lateral margination distinct and complete, joined with apical and basal margination at anterior and posterior angles, respectively; basal margination distinct laterally, less distinct medially; midlateral setae inserted on lateral margination (“lateral bead”) at about one-fourth pronotal length; basolateral setae inserted on margination at posterior angles. Prosternal intercoxal process short, rounded, with complete but faint margination apically.
) slightly narrow (ratio PWM/EW = 0.61 to 0.65), distinctly narrowed basally (ratio PWB/PWA = 1.04 to 1.09), widest distinctly anterior to mid-length, with anterior angles very short and broadly rounded; posterior angles slightly obtuse and slightly projected laterally; apical margin straight or very slightly concave between anterior angles; lateral margins arcuate and with a short and shallow sinuation just anterior to posterior angles; basal margin faintly bisinuate, not or only slightly lobate medially; anterior margination distinct laterally, more or less effaced medially; lateral margination distinct and complete, joined with apical and basal margination at anterior and posterior angles, respectively; basal margination distinct laterally, less distinct medially; midlateral setae inserted on lateral margination (“lateral bead”) at about one-fourth pronotal length; basolateral setae inserted on margination at posterior angles. Prosternal intercoxal process short, rounded, with complete but faint margination apically.
</paragraph>
<caption id="B89B0288FF9CFFABD3AFDE5AFE3766BB" ID-DOI="http://doi.org/10.5281/zenodo.11067309" ID-Zenodo-Dep="11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="19" pageNumber="470" startId="19.[155,167,1597,1616]" targetBox="[123,1176,216,1583]" targetPageId="19" targetType="figure">
<paragraph id="EC5B5200FF9CFFABD3AFDE5AFE3766BB" blockId="19.[123,1174,1596,1692]" pageId="19" pageNumber="470">
@ -304,11 +301,7 @@ Kavanaugh &amp; Cueva-Dabkoski
A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km.
</paragraph>
</caption>
<paragraph id="EC5B5200FF9BFFACD39FD9BBFC0F6292" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
Elytra. Broadly ovate, longer than wide (ratio EL/
<collectionCode id="8AF5CAC5FF9BFFACD1F6D9BBFD2161D3" box="[706,747,476,500]" pageId="20" pageNumber="471">EW</collectionCode>
= 1.27 to 1.37), widest slightly posteri- or to mid-length, obliquely truncate apically, humeri obtusely rounded; elytral striae deeply impressed throughout, elytral intervals flat or slightly convex, smooth; parascutellar setiferous pore present at base of interval 1 at junction of striae 1 and 2; two discal setiferous pores present on interval three, the anterior seta inserted adjacent to stria 3 near elytral midlength, the other insert- ed adjacent to stria 2 near apical one-fourth at level of subapical pale band; apical seta present, inserted near apex of interval 3; umbilicate series comprised of 11 or 12 setae.
</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FD9BBFC0F6292" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">Elytra. Broadly ovate, longer than wide (ratio EL/EW = 1.27 to 1.37), widest slightly posterior to mid-length, obliquely truncate apically, humeri obtusely rounded; elytral striae deeply impressed throughout, elytral intervals flat or slightly convex, smooth; parascutellar setiferous pore present at base of interval 1 at junction of striae 1 and 2; two discal setiferous pores present on interval three, the anterior seta inserted adjacent to stria 3 near elytral midlength, the other inserted adjacent to stria 2 near apical one-fourth at level of subapical pale band; apical seta present, inserted near apex of interval 3; umbilicate series comprised of 11 or 12 setae.</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDAD8FBAE6311" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
Legs. Tarsal claws finely denticulate medially. Males (
<figureCitation id="74DF4E85FF9BFFACD1C0DAD9FCAB62F1" box="[756,865,702,726]" captionStart="FIGURE" captionStartId="31.[155,167,1512,1531]" captionTargetBox="[253,1040,238,1497]" captionTargetId="figure-15@31.[253,1044,216,1497]" captionTargetPageId="31" captionText="FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian &amp; Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11067319" httpUri="https://zenodo.org/record/11067319/files/figure.png" pageId="20" pageNumber="471">Figs. 14D</figureCitation>
@ -334,24 +327,25 @@ the other specimens). All of these specimens otherwise are similar.
<emphasis id="DE908E12FF9BFFACD39FDCE5FE5064BE" bold="true" box="[171,410,1154,1177]" pageId="20" pageNumber="471">Habitat distribution.</emphasis>
Members of this species were found under small stones on the open sandy shore of the Longchuan River, a medium-sized stream, at an elevation of
<quantity id="2B1CFFE5FF9BFFACD0F3DCC6FBD2649E" box="[967,1048,1185,1209]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="20" pageNumber="471" unit="m" value="1230.0">1230 m</quantity>
and togeth- er with adults of
and together with adults of
<emphasis id="DE908E12FF9BFFACD205DCA5FD9C64FE" box="[305,598,1217,1241]" italics="true" pageId="20" pageNumber="471">
<taxonomicName id="2BE42983FF9BFFACD205DCA5FE5764FE" box="[305,413,1217,1241]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="elegans">T. elegans</taxonomicName>
and
<taxonomicName id="2BE42983FF9BFFACD2E7DCA5FD9C64FE" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" box="[467,598,1218,1241]" class="Insecta" family="Carabidae" genus="Tetragonoderus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="punctatus">T. punctatus</taxonomicName>
</emphasis>
. The specimen from the Tombiguan area was collect- ed in a baited pitfall trap near a stream through secondary (disturbed) broadleaf forest at an elevation of
. The specimen from the Tombiguan area was collected in a baited pitfall trap near a stream through secondary (disturbed) broadleaf forest at an elevation of
<quantity id="2B1CFFE5FF9BFFACD3FDDD65FED4653D" box="[201,286,1282,1306]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="20" pageNumber="471" unit="m" value="1290.0">1290 m</quantity>
.
</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDD44FC3965FC" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">
<emphasis id="DE908E12FF9BFFACD39FDD44FCD6651D" bold="true" box="[171,796,1315,1338]" pageId="20" pageNumber="471">
Geographical distribution within the Gaoligong
<materialsCitation id="5C8C585DFF9BFFACD1EFDD44FC3965FC" collectingDate="2006-06-03" collectingDateMax="2016-06-27" collectingDateMin="2006-06-03" collectionCode="NMPC, R, CAS, IOZ" collectorName="J. Hajek &amp; J. Ruzicka &amp; C. B. Wang &amp; Longchuan Jiang &amp; D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong" county="Tengchong County" elevation="1290" location="Tongbiguan Township" municipality="We" pageId="20" pageNumber="471" specimenCount="15" specimenCount-female="2" specimenCount-male="8" stateProvince="Shan">
<emphasis id="DE908E12FF9BFFACD39FDD44FD18651D" bold="true" box="[171,722,1315,1338]" pageId="20" pageNumber="471">Geographical distribution within the Gaoligong</emphasis>
<emphasis id="DE908E12FF9BFFACD1EFDD44FCD6651D" bold="true" box="[731,796,1315,1338]" pageId="20" pageNumber="471">
<collectingRegion id="2E209CE2FF9BFFACD1EFDD44FCDD651D" box="[731,791,1315,1338]" country="Myanmar" name="Shan" pageId="20" pageNumber="471">Shan</collectingRegion>
.
.
</emphasis>
<figureCitation id="74DF4E85FF9BFFACD011DD45FC42651D" box="[805,904,1314,1338]" captionStart="FIGURE" captionStartId="19.[155,167,1597,1616]" captionTargetBox="[123,1176,216,1583]" captionTargetId="figure-73@19.[120,1176,216,1584]" captionTargetPageId="19" captionText="FIGURE 11. Tetragonoderus parviculus Kavanaugh &amp; Cueva-Dabkoski, sp. nov. A. Holotype, dorsal aspect, scale line = 1.0 mm (CASENT1039396, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); B. Labels associated with holotype; C. Map showing locality records (black circle) for this species in the Gaoligong Shan region, scale line = 100 km." figureDoi="http://doi.org/10.5281/zenodo.11067309" httpUri="https://zenodo.org/record/11067309/files/figure.png" pageId="20" pageNumber="471">Fig. 11B</figureCitation>
.
<materialsCitation id="5C8C585DFF9BFFACD0A0DD44FD4E65BC" ID-GBIF-Occurrence="4902827320" collectingDate="2006-06-03" collectingDateMax="2016-06-27" collectingDateMin="2006-06-03" collectionCode="NMPC, R, CAS, IOZ" collectorName="J. Hajek &amp; J. Ruzicka &amp; C. B. Wang &amp; Longchuan Jiang &amp; D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong" county="Tengchong County" elevation="1290" location="Tongbiguan Township" municipality="We" pageId="20" pageNumber="471" specimenCount="15" specimenCount-female="2" specimenCount-male="8" stateProvince="Shan">
<collectingMunicipality id="0C3FC87AFF9BFFACD0A0DD44FC7D651D" box="[916,951,1315,1338]" pageId="20" pageNumber="471">We</collectingMunicipality>
examined a total of
<specimenCount id="FAE29989FF9BFFACD34FDD25FED7657D" box="[123,285,1346,1370]" count="5" pageId="20" pageNumber="471" type="generic">five specimens</specimenCount>
@ -385,6 +379,8 @@ W of Tongbiguan
, &amp;
<collectorName id="411137D6FF9BFFACD2B3DDE4FDCA65BC" box="[391,512,1411,1435]" pageId="20" pageNumber="471">C.B. Wang</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF9BFFACD1B0DDE4FCC265FC" ID-GBIF-Occurrence="4902827301" collectingDate="2006-06-03" collectingDateMax="2016-06-27" collectingDateMin="2006-06-03" collectionCode="NMPC, R, CAS, IOZ" collectorName="J. Hajek &amp; J. Ruzicka &amp; C. B. Wang &amp; Longchuan Jiang &amp; D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong" county="Tengchong County" elevation="1290" location="Tongbiguan Township" municipality="We" pageId="20" pageNumber="471" specimenCount="15" specimenCount-female="2" specimenCount-male="8" stateProvince="Shan">
<specimenCount id="FAE29989FF9BFFACD1B0DDE4FD2D65BC" box="[644,743,1411,1435]" count="1" pageId="20" pageNumber="471" type="female">1 female</specimenCount>
;
<collectionCode id="8AF5CAC5FF9BFFACD1C1DDE4FC8D65BC" box="[757,839,1411,1435]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="471">NMPC</collectionCode>
@ -413,6 +409,8 @@ just below bridge at
&amp;
<collectorName id="411137D6FF9BFFACD12CDDA3FD4165FB" box="[536,651,1476,1500]" pageId="20" pageNumber="471">D.Z. Dong</collectorName>
collectors [
</materialsCitation>
<materialsCitation id="5C8C585DFF9BFFACD03CDDA4FC3965FC" ID-GBIF-Occurrence="4902827311" box="[776,1011,1475,1499]" collectingDate="2006-06-03" collectingDateMax="2016-06-27" collectingDateMin="2006-06-03" collectionCode="NMPC, R, CAS, IOZ" collectorName="J. Hajek &amp; J. Ruzicka &amp; C. B. Wang &amp; Longchuan Jiang &amp; D. H. Kavanaugh &amp; L. Brett &amp; H. B. Liang &amp; D. Z. Dong" county="Tengchong County" elevation="1290" location="Tongbiguan Township" municipality="We" pageId="20" pageNumber="471" specimenCount="15" specimenCount-female="2" specimenCount-male="8" stateProvince="Shan">
<specimenCount id="FAE29989FF9BFFACD03CDDA4FCAB65FC" box="[776,865,1475,1499]" count="4" pageId="20" pageNumber="471" type="male">4 males</specimenCount>
;
<collectionCode id="8AF5CAC5FF9BFFACD058DDA4FC6865FC" box="[876,930,1475,1499]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="471" type="Museum">CAS</collectionCode>
@ -420,7 +418,6 @@ collectors [
<collectionCode id="8AF5CAC5FF9BFFACD09ADDA4FC2B65FC" box="[942,993,1475,1499]" pageId="20" pageNumber="471">IOZ</collectionCode>
]).
</materialsCitation>
</emphasis>
</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDD84FF19663C" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">At present, this species is known only from the southwestern part of the study area (Core Area 6).</paragraph>
<paragraph id="EC5B5200FF9BFFACD39FDE43FE4D667B" blockId="20.[123,1174,219,1725]" pageId="20" pageNumber="471">

View file

@ -0,0 +1,199 @@
<document id="42F833723E37CC934499D3EC746A26D5" ID-DOI="10.5281/zenodo.12585046" ID-ISSN="2064-2474" ID-Zenodo-Dep="12585046" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719593661524" checkinUser="felipe" docAuthor="Papp, L." docDate="2008" docId="647F87A3FFE47373FDA2FAC6FB99FBCF" docLanguage="en" docName="ActaZoolAcadSciH.54.3.235-255.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 54 (3)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Chaetopodella (Afrochaetopodella) Papp, 2008, subgen. n." docType="treatment" docVersion="2" lastPageNumber="242" masterDocId="9846FFDBFFE17374FFC4FF8CFF98FFAF" masterDocTitle="Afrotropical Species Of Chaetopodella Duda (Diptera: Sphaeroceridae)" masterLastPageNumber="255" masterPageNumber="235" pageNumber="240" updateTime="1719595852930" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="AFE3A5DDEA8E6B4BB3545E844A197841">Afrotropical Species Of Chaetopodella Duda (Diptera: Sphaeroceridae)</mods:title>
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<mods:namePart id="50FEE14C83C1EC358844DCE6F0417C31">Papp, L.</mods:namePart>
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<mods:date id="AB548FE536370A2C0FA6E04560601FEF">2008</mods:date>
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<mods:number id="065CF909000F1F1CFAAF24645B1495FB">54</mods:number>
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<treatment id="647F87A3FFE47373FDA2FAC6FB99FBCF" LSID="urn:lsid:plazi:treatment:647F87A3FFE47373FDA2FAC6FB99FBCF" httpUri="http://treatment.plazi.org/id/647F87A3FFE47373FDA2FAC6FB99FBCF" lastPageId="7" lastPageNumber="242" pageId="5" pageNumber="240">
<subSubSection id="A4CC653EFFE47371FDA2FAC6FC54FACA" box="[614,972,1354,1381]" pageId="5" pageNumber="240" type="nomenclature">
<paragraph id="EC6936B5FFE47371FDA2FAC6FC54FACA" blockId="5.[614,972,1354,1381]" box="[614,972,1354,1381]" pageId="5" pageNumber="240">
<heading id="B72181D9FFE47371FDA2FAC6FC54FACA" box="[614,972,1354,1381]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="240" reason="2">
<taxonomicName id="2BD64D36FFE47371FDA2FAC6FCD5FACB" authorityName="Papp" authorityYear="2008" box="[614,845,1354,1380]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="240" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Afrochaetopodella">
<emphasis id="DEA2EAA7FFE47371FDA2FAC6FCD5FACB" bold="true" box="[614,845,1354,1380]" pageId="5" pageNumber="240">Afrochaetopodella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFE47371FC91FAC7FC54FACA" box="[853,972,1355,1381]" pageId="5" pageNumber="240" rank="subGenus">subgen. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4CC653EFFE47373FD0AFAE2FB99FBCF" lastPageId="7" lastPageNumber="242" pageId="5" pageNumber="240" type="description">
<paragraph id="EC6936B5FFE47371FD0AFAE2FCFCFA27" blockId="5.[718,868,1390,1416]" box="[718,868,1390,1416]" pageId="5" pageNumber="240">
<heading id="B72181D9FFE47371FD0AFAE2FCFCFA27" box="[718,868,1390,1416]" centered="true" fontSize="11" level="3" pageId="5" pageNumber="240" reason="3">
(
<figureCitation id="74ED2A30FFE47371FD12FAE2FCC4FA27" box="[726,860,1390,1416]" captionStart-0="Figs 1318" captionStart-1="Figs 1926" captionStartId-0="8.[317,360,1657,1678]" captionStartId-1="9.[317,360,1649,1670]" captionTargetBox-0="[337,1261,459,1591]" captionTargetBox-1="[320,1269,445,1600]" captionTargetId-0="figure-15@8.[317,1269,453,1601]" captionTargetId-1="figure-10@9.[317,1269,445,1603]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" captionText-1="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi-0="http://doi.org/10.5281/zenodo.12585056" figureDoi-1="http://doi.org/10.5281/zenodo.12585058" httpUri-0="https://zenodo.org/record/12585056/files/figure.png" httpUri-1="https://zenodo.org/record/12585058/files/figure.png" pageId="5" pageNumber="240">Figs 1326</figureCitation>
)
</heading>
</paragraph>
<paragraph id="EC6936B5FFE47371FEF9FA3DFC17FA69" blockId="5.[317,911,1457,1536]" box="[317,911,1457,1478]" pageId="5" pageNumber="240">
Type species:
<taxonomicName id="2BD64D36FFE47371FE02FA3DFD3BFA69" authorityName="Papp" authorityYear="2008" box="[454,675,1457,1478]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="DEA2EAA7FFE47371FE02FA3DFD3BFA69" box="[454,675,1457,1478]" italics="true" pageId="5" pageNumber="240">Chaetopodella reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFE47371FD6DFA3DFD45FA69" box="[681,733,1457,1478]" pageId="5" pageNumber="240" rank="species">sp. n.</taxonomicNameLabel>
(here designated).
</paragraph>
<paragraph id="EC6936B5FFE47371FEF9FA42FE75FA4C" blockId="5.[317,911,1457,1536]" box="[317,493,1486,1507]" pageId="5" pageNumber="240">Gender: feminine.</paragraph>
<paragraph id="EC6936B5FFE47371FEF9FA67FD49F9AF" blockId="5.[317,911,1457,1536]" box="[317,721,1515,1536]" pageId="5" pageNumber="240">
Other species included:
<taxonomicName id="2BD64D36FFE47371FDE0FA67FD0FF9AF" authorityName="Papp" authorityYear="2008" box="[548,663,1515,1536]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="keniaca" status="sp. nov.">
<emphasis id="DEA2EAA7FFE47371FDE0FA67FD0FF9AF" box="[548,663,1515,1536]" italics="true" pageId="5" pageNumber="240">Ch. keniaca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFE47371FD59FA67FD49F9AF" box="[669,721,1515,1536]" pageId="5" pageNumber="240" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="EC6936B5FFE47372FEBAF9ABFB72FE5A" blockId="5.[317,1270,1575,1813]" lastBlockId="6.[317,1270,439,571]" lastPageId="6" lastPageNumber="241" pageId="5" pageNumber="240">
In a number of characteristics it is similar to the nominate subgenus. The most important differences are as follow (most of them are synapomorphies): All inter-frontal setae short. Anterior fronto-orbital seta minute. Mid trochanter with a lateral seta, which is shorter than trochanter. Wings clear but not milky. Costal setae between veins H and R
<subScript id="705234F0FFE47371FD48F94DFD0DF97E" attach="left" box="[652,661,1729,1745]" fontSize="7" pageId="5" pageNumber="240">1</subScript>
short, much shorter than width of costal cell. Discal cell narrow and very long. Abdomen (preabdomen) very long, male sternite 5 with a pair of 3-lobed caudal processes (
<figureCitation id="74ED2A30FFE47371FD0AF977FCCCF8BA" box="[718,852,1787,1813]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="5" pageNumber="240">Figs 1415</figureCitation>
,
<figureCitation id="74ED2A30FFE47371FC9AF977FC34F8BA" box="[862,940,1787,1813]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="5" pageNumber="240">2021</figureCitation>
). Subepandrial sclerite without strong thorn-like setae (
<figureCitation id="74ED2A30FFE77372FDB1FE3BFD50FE7E" box="[629,712,439,465]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="6" pageNumber="241">Fig. 24</figureCitation>
) (that must be a reduction, i.e. a synapomorphy). Male genitalia small. Postgonites robust with several edges or tips (
<figureCitation id="74ED2A30FFE77372FB90FE57FB37FE5A" box="[1108,1199,475,501]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="6" pageNumber="241">Figs 17</figureCitation>
,
<figureCitation id="74ED2A30FFE77372FB78FE57FB45FE5A" box="[1212,1245,475,501]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="6" pageNumber="241">25</figureCitation>
).
</paragraph>
<paragraph id="EC6936B5FFE77372FEBAFE72FE32FD94" blockId="6.[317,1270,439,571]" pageId="6" pageNumber="241">The subgeneric name refers to the distribution of the two species presently included.</paragraph>
<paragraph id="EC6936B5FFE77372FE23FD06FBD3FD0A" blockId="6.[487,1099,650,677]" box="[487,1099,650,677]" pageId="6" pageNumber="241">
<taxonomicName id="2BD64D36FFE77372FE23FD06FB9DFD0B" authorityName="Papp" authorityYear="2008" box="[487,1029,650,676]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="keniaca" status="sp. nov." subGenus="Afrochaetopodella">
<emphasis id="DEA2EAA7FFE77372FE23FD06FB9DFD0B" bold="true" box="[487,1029,650,676]" pageId="6" pageNumber="241">Chaetopodella (Afrochaetopodella) keniaca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFE77372FBC9FD07FBD3FD0A" box="[1037,1099,651,677]" pageId="6" pageNumber="241" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="EC6936B5FFE77372FD0AFD22FCFDFD67" blockId="6.[718,869,686,712]" box="[718,869,686,712]" pageId="6" pageNumber="241">
<heading id="B72181D9FFE77372FD0AFD22FCFDFD67" box="[718,869,686,712]" centered="true" fontSize="11" level="3" pageId="6" pageNumber="241" reason="3">
(
<figureCitation id="74ED2A30FFE77372FD12FD22FCC4FD67" box="[726,860,686,712]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="6" pageNumber="241">Figs 1318</figureCitation>
)
</heading>
</paragraph>
<paragraph id="EC6936B5FFE77372FEBAFD7DFB6DFCA9" blockId="6.[317,1269,753,803]" box="[382,1269,753,774]" pageId="6" pageNumber="241">
<materialsCitation id="5CBE3CE8FFE77372FEBAFD7DFB6DFCA9" box="[382,1269,753,774]" collectingDate="2003-02-25" collectionCode="HNHM" country="Kenya" location="Shimba Hills Nat. Park" pageId="6" pageNumber="241" specimenCount="1" typeStatus="holotype">
<typeStatus id="336D8817FFE77372FEBAFD7DFE4FFCA9" box="[382,471,753,774]" pageId="6" pageNumber="241" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="8AC7AE70FFE77372FDD2FD7DFDFEFCA9" box="[534,614,753,774]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="241">HNHM</collectionCode>
):
<collectingCountry id="94C17625FFE77372FDB0FD7DFD54FCA9" box="[628,716,753,774]" name="Kenya" pageId="6" pageNumber="241">KENYA</collectingCountry>
:
<location id="E909606EFFE77372FD17FD7DFC29FCA9" LSID="urn:lsid:plazi:treatment:647F87A3FFE47373FDA2FAC6FB99FBCF:E909606EFFE77372FD17FD7DFC29FCA9" box="[723,945,753,774]" country="Kenya" name="Shimba Hills Nat. Park" pageId="6" pageNumber="241">Shimba Hills Nat. Park</location>
,
<date id="98681075FFE77372FC7DFD7DFBAEFCA9" box="[953,1078,753,774]" pageId="6" pageNumber="241" value="2003-02-25">
<collectingDate id="882CE99DFFE77372FC7DFD7DFBAEFCA9" box="[953,1078,753,774]" pageId="6" pageNumber="241" value="2003-02-25">2003. 02. 25.</collectingDate>
</date>
, leg.
<smallCapsWord id="EA8FA069FFE77372FBA3FD7DFB55FCAA" baselines="769,769" box="[1127,1229,753,774]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Mahunka" pageId="6" pageNumber="241">MAHUNKA</smallCapsWord>
S.
</materialsCitation>
</paragraph>
<paragraph id="EC6936B5FFE77372FEF9FC82FBA8FC8C" blockId="6.[317,1269,753,803]" box="[317,1072,782,803]" pageId="6" pageNumber="241">
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L. [Lujza] [abdomen and genitalia in a plastic microvial with glycerol].
</paragraph>
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Measurements in mm: body length 1.54, wing length 1.32, wing width 0.47. talia), lateral view. Scale:
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</paragraph>
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<emphasis id="DEA2EAA7FFE77372FEF9F95EFE37F948" bold="true" box="[317,431,1746,1767]" pageId="6" pageNumber="241">Figs 1012.</emphasis>
<taxonomicName id="2BD64D36FFE77372FE71F95FFCA8F948" authority="(DUDA)" baseAuthorityName="DUDA" baseAuthorityYear="1920" box="[437,816,1746,1768]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="denigrata" subGenus="Chaetopodella">
<emphasis id="DEA2EAA7FFE77372FE71F95FFDDAF947" box="[437,578,1747,1768]" italics="true" pageId="6" pageNumber="241">Chaetopodella</emphasis>
(
<emphasis id="DEA2EAA7FFE77372FD94F95FFDEBF947" box="[592,627,1747,1768]" italics="true" pageId="6" pageNumber="241">Ch.</emphasis>
)
<emphasis id="DEA2EAA7FFE77372FD44F95FFD45F947" box="[640,733,1747,1768]" italics="true" pageId="6" pageNumber="241">denigrata</emphasis>
(
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)
</taxonomicName>
, male postabdomen and genitalia. 1011 = surstylus: 10 = in the broadest (sublateral) view, 11 = in ventral view; 12 = phallic complex (inner geni-
</paragraph>
</caption>
<paragraph id="EC6936B5FFE67373FEBAFE38FC1AFDAC" blockId="7.[317,1271,436,1062]" pageId="7" pageNumber="242">
Head with anterior 1/3 of frons yellow, other frons dark brown. Face whitish yellow, gena yellow. Three pairs of short interfrontals. Antenna dark brown, incl. scape. Arista
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long. Vibrissa strong. Genal setae sparse but comparatively long.
</paragraph>
<paragraph id="EC6936B5FFE67373FEBAFD87FDE4FD1F" blockId="7.[317,1271,436,1062]" pageId="7" pageNumber="242">
Mesonotum microtrichose dark brown with one sagittal and two dorsocentral stripes less microtrichose, i.e. more shiny. Postpronotum, lateral half of notopleura and a small supra-alar area yellowish. Pleura lighter than mesonotum, particularly so for ventral edges of anepisternum, anepimeron and metapleural parts. Three pairs of dorsocentrals, shortening anteriorad. Scutellum
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long,
<quantity id="2B2E9B50FFE67373FE14FDF2FDBDFD3C" box="[464,549,638,659]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="242" unit="mm" value="0.31">0.31 mm</quantity>
wide, lateral scutellars
<quantity id="2B2E9B50FFE67373FCCEFDF2FCFAFD3C" box="[778,866,638,659]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="242" unit="mm" value="0.27">0.27 mm</quantity>
, apical scutellars
<quantity id="2B2E9B50FFE67373FBC8FDF2FBF9FD3C" box="[1036,1121,638,659]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="7" pageNumber="242" unit="mm" value="0.47">0.47 mm</quantity>
long. Posterior katepisternal seta
<quantity id="2B2E9B50FFE67373FE28FD17FDDBFD1F" box="[492,579,667,688]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="242" unit="mm" value="0.22">0.22 mm</quantity>
long.
</paragraph>
<paragraph id="EC6936B5FFE67373FEBAFD34FC93FC8C" blockId="7.[317,1271,436,1062]" pageId="7" pageNumber="242">
Wing 2.79 times longer than broad. Wing brownish, veins light brown, including costa. Microcilia of membrane rather strong. Strong subbasal seta on costa only
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long. Setae on first costal section only
<quantity id="2B2E9B50FFE67373FDD9FD7DFDE8FCA9" box="[541,624,753,774]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="242" unit="mm" value="0.03">0.03 mm</quantity>
long. Discal cell with a distinct small caudal vein appendage. R-M dM-Cu: dM-Cu = 19.5/6 = 3.25. Halter white.
</paragraph>
<paragraph id="EC6936B5FFE67373FEBAFCA7FB3AFCEF" blockId="7.[317,1271,436,1062]" box="[382,1186,811,832]" pageId="7" pageNumber="242">
Legs ochreous, only tarsi darker (brown). Lateral seta of mid trochanter
<quantity id="2B2E9B50FFE67373FB80FCA7FB06FCEF" box="[1092,1182,811,832]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="242" unit="mm" value="0.1">0.10 mm</quantity>
.
</paragraph>
<paragraph id="EC6936B5FFE67373FEBAFCC4FCCAFB89" blockId="7.[317,1271,436,1062]" pageId="7" pageNumber="242">
Male sternite 5 comparatively long but not broad (
<figureCitation id="74ED2A30FFE67373FCA7FCC4FC32FCF2" box="[867,938,840,861]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="7" pageNumber="242">Fig. 14</figureCitation>
), its caudal pair of processes large three-lobed fork, the medial one being the longest, lateral process with 3 long setae (
<figureCitation id="74ED2A30FFE67373FB99FCE8FB3BFCD6" box="[1117,1187,868,889]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="7" pageNumber="242">Fig. 15</figureCitation>
). No extremely long setae on sternite 5. Right part of the syntergosternite 68 (tergite 7 and a part of sternite 6) forms a narrow but long re-curved sclerotized process; dorsal part of the syntergosternite large, left lateral and ventral parts almost completely segregated (
<figureCitation id="74ED2A30FFE67373FC81FC37FC13FC7F" box="[837,907,955,976]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="7" pageNumber="242">Fig. 13</figureCitation>
). Surstylus (
<figureCitation id="74ED2A30FFE67373FBC4FC37FBDDFC7F" box="[1024,1093,955,976]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="7" pageNumber="242">Fig. 16</figureCitation>
) rather compact in two lobes with a number of processes but with only 1 long setae. Postgonite (
<figureCitation id="74ED2A30FFE67373FBF8FC54FB1CFC42" box="[1084,1156,984,1005]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="7" pageNumber="242">Fig. 17</figureCitation>
) thick, medium-long with blunt apex and small swellings dorsally. Apical part of the dorsal process of distiphallus (
<figureCitation id="74ED2A30FFE67373FE55FB9DFE4FFB89" box="[401,471,1041,1062]" captionStart="Figs 1318" captionStartId="8.[317,360,1657,1678]" captionTargetBox="[337,1261,459,1591]" captionTargetId="figure-15@8.[317,1269,453,1601]" captionTargetPageId="8" captionText="Figs 1318. Chaetopodella (Afrochaetopodella) keniaca sp. n., holotype male, postabdomen and genitalia.13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 =apical part of the dorsal process of distiphallus,subdorsal view. Scale: 0.1 mm for all" figureDoi="http://doi.org/10.5281/zenodo.12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="7" pageNumber="242">Fig. 18</figureCitation>
) with blunt, rather short fork apically.
</paragraph>
<paragraph id="EC6936B5FFE67373FEBAFBC7FB99FBCF" blockId="7.[382,1025,1099,1120]" box="[382,1025,1099,1120]" pageId="7" pageNumber="242">
Etymology. The specific epithet refers to its
<typeStatus id="336D8817FFE67373FCF1FBC7FCC6FBCF" box="[821,862,1099,1120]" pageId="7" pageNumber="242">type</typeStatus>
locality,
<collectingCountry id="94C17625FFE67373FC7FFBC7FC65FBCF" box="[955,1021,1099,1120]" name="Kenya" pageId="7" pageNumber="242">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,181 @@
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<mods:title id="CDC2A09D860F1716E2DB99D5C6DDB789">Afrotropical Species Of Chaetopodella Duda (Diptera: Sphaeroceridae)</mods:title>
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<subSubSection id="A4CC653EFFE67373FE23FB3DFBD3FB64" box="[487,1099,1201,1227]" pageId="7" pageNumber="242" type="nomenclature">
<paragraph id="EC6936B5FFE67373FE23FB3DFBD3FB64" blockId="7.[487,1099,1201,1227]" box="[487,1099,1201,1227]" pageId="7" pageNumber="242">
<heading id="B72181D9FFE67373FE23FB3DFBD3FB64" box="[487,1099,1201,1227]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="242" reason="2">
<taxonomicName id="2BD64D36FFE67373FE23FB3DFB9DFB64" authorityName="Papp" authorityYear="2008" box="[487,1029,1201,1227]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov." subGenus="Afrochaetopodella">
<emphasis id="DEA2EAA7FFE67373FE23FB3DFB9DFB64" bold="true" box="[487,1029,1201,1227]" pageId="7" pageNumber="242">Chaetopodella (Afrochaetopodella) reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFE67373FBC9FB3DFBD3FB64" box="[1037,1099,1201,1227]" pageId="7" pageNumber="242" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4CC653EFFE6737EFD0AFB58FB40FCC6" lastPageId="10" lastPageNumber="245" pageId="7" pageNumber="242" type="description">
<paragraph id="EC6936B5FFE67373FD0AFB58FCFCFB41" blockId="7.[718,868,1236,1262]" box="[718,868,1236,1262]" pageId="7" pageNumber="242">
<heading id="B72181D9FFE67373FD0AFB58FCFCFB41" box="[718,868,1236,1262]" centered="true" fontSize="11" level="3" pageId="7" pageNumber="242" reason="3">
(
<figureCitation id="74ED2A30FFE67373FD12FB58FCC5FB41" box="[726,861,1236,1262]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="7" pageNumber="242">Figs 1926</figureCitation>
)
</heading>
</paragraph>
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<materialsCitation id="5CBE3CE8FFE67373FEBAFA94FB6DFA82" box="[382,1269,1304,1325]" collectionCode="HNHM, EORY" country="Tanzania" location="Meru" pageId="7" pageNumber="242" specimenCount="1" typeStatus="holotype">
<typeStatus id="336D8817FFE67373FEBAFA94FE4FFA82" box="[382,471,1304,1325]" pageId="7" pageNumber="242" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="8AC7AE70FFE67373FDD3FA94FDFFFA82" box="[535,615,1304,1325]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="7" pageNumber="242">HNHM</collectionCode>
):
<collectingCountry id="94C17625FFE67373FDB2FA94FD48FA82" box="[630,720,1304,1325]" name="Tanzania" pageId="7" pageNumber="242">Tanzánia</collectingCountry>
,
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, 1979. II-III., leg. [Miklós]
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<collectionCode id="8AC7AE70FFE67373FBD2FA94FBD4FA84" box="[1046,1100,1304,1325]" pageId="7" pageNumber="242">EŐRY</collectionCode>
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[György]
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</paragraph>
<paragraph id="EC6936B5FFE67373FEF9FAB8FC17FAE6" blockId="7.[317,1269,1304,1353]" box="[317,911,1332,1353]" pageId="7" pageNumber="242">[abdomen and genitalia in a plastic microvial with glycerol].</paragraph>
<paragraph id="EC6936B5FFE67373FEBAFAE2FBF3FA2C" blockId="7.[317,1271,1390,1814]" box="[382,1131,1390,1411]" pageId="7" pageNumber="242">Measurements in mm: body length 1.70, wing length 1.52, wing width 0.55.</paragraph>
<paragraph id="EC6936B5FFE67373FEBAFA07FB6BFA59" blockId="7.[317,1271,1390,1814]" pageId="7" pageNumber="242">
Frons dark brown, anterior 1/5 reddish yellow. Ocellar triangle with frontral stripe subshiny. Face whitish yellow. One single medial interfrontal seta (not cruciate). Anterior fronto-orbital minute thin
<quantity id="2B2E9B50FFE67373FEA3FA48FE22FA76" box="[359,442,1476,1497]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="242" unit="mm" value="0.04">0.04 mm</quantity>
long, posterior pair thick,
<quantity id="2B2E9B50FFE67373FD75FA48FC9CFA76" box="[689,772,1476,1497]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="242" unit="mm" value="0.13">0.13 mm</quantity>
long. Gena and cheeks yellow, posterior part of gena yellowish grey. Genal setae sparse but comparatively long. Antennae dark brown, incl. base of scape.
</paragraph>
<paragraph id="EC6936B5FFE67373FEBAFA72FDAFF9C6" blockId="7.[317,1271,1390,1814]" pageId="7" pageNumber="242">
Mesonotum dark brown, scutellum velvety black. Pleura also dark brown, only notopleural area diffusely lighter (yellowish). Anterior katepisternal
<quantity id="2B2E9B50FFE67373FC9CF997FC37F99F" box="[856,943,1563,1584]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="242" unit="mm" value="0.09">0.09 mm</quantity>
, posterior one
<quantity id="2B2E9B50FFE67373FBFEF997FB09F99F" box="[1082,1169,1563,1584]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="242" unit="mm" value="0.2">0.20 mm</quantity>
, but rather thin. Both apical scutellars broken on the
<typeStatus id="336D8817FFE67373FD0DF9B4FC87F9E2" box="[713,799,1592,1613]" pageId="7" pageNumber="242" type="holotype">holotype</typeStatus>
, lateral scutellar
<quantity id="2B2E9B50FFE67373FC05F9B4FB80F9E2" box="[961,1048,1592,1613]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="242" unit="mm" value="0.25">0.25 mm</quantity>
; scutellum shorter than broad (0.24 vs
<quantity id="2B2E9B50FFE67373FE16F9D8FDB4F9C6" box="[466,556,1620,1641]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="242" unit="mm" value="0.3">0.30 mm</quantity>
).
</paragraph>
<paragraph id="EC6936B5FFE67373FEBAF9FDFB04F96F" blockId="7.[317,1271,1390,1814]" pageId="7" pageNumber="242">
Wing 2.76 times longer than broad. Wing yellowish, veins yellow, including costa. Medial and cubital veins rather whitish. Strong sub-basal seta on costa only
<quantity id="2B2E9B50FFE67373FC78F902FB8BF90C" box="[956,1043,1678,1699]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="242" unit="mm" value="0.09">0.09 mm</quantity>
. Discal cell with a small caudal vein appendage. R-M dM-Cu: dM-Cu = 3.23. Halter white, stalk light yellow.
</paragraph>
<paragraph id="EC6936B5FFE67373FEBAF944FC3BF972" blockId="7.[317,1271,1390,1814]" box="[382,931,1736,1757]" pageId="7" pageNumber="242">
Mid trochanter with a short lateral seta (only
<quantity id="2B2E9B50FFE67373FCF9F944FC0FF972" box="[829,919,1736,1757]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="242" unit="mm" value="0.06">0.06 mm</quantity>
).
</paragraph>
<paragraph id="EC6936B5FFE6737DFEBAF968FBFBF8B9" blockId="7.[317,1271,1390,1814]" lastBlockId="9.[317,1270,1649,1814]" lastPageId="9" lastPageNumber="244" pageId="7" pageNumber="242">
Male sternite 5 long but not broad (
<figureCitation id="74ED2A30FFE67373FD01F968FC93F956" box="[709,779,1764,1785]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="7" pageNumber="242">Fig. 20</figureCitation>
), peculiar with its very long light setae (longest ones longer than width of sternite), caudal pair of processes large, three-lobed, like in
<taxonomicName id="2BD64D36FFE67373FBF3F88DFB31F8B9" authorityName="Papp" authorityYear="2008" box="[1079,1193,1793,1814]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="keniaca">
<emphasis id="DEA2EAA7FFE67373FBF3F88DFB31F8B9" box="[1079,1193,1793,1814]" italics="true" pageId="7" pageNumber="242">Ch. keniaca</emphasis>
</taxonomicName>
, but not for
<figureCitation id="74ED2A30FFE8737DFE36F88DFDC7F8B9" box="[498,607,1793,1814]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="9" pageNumber="244">Figs 19, 22</figureCitation>
, and
<figureCitation id="74ED2A30FFE8737DFD56F88DFD40F8B9" box="[658,728,1793,1814]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="9" pageNumber="244">Fig. 20</figureCitation>
, respectively,
<quantity id="2B2E9B50FFE8737DFCA6F88DFC33F8B9" box="[866,939,1793,1814]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="244" unit="mm" value="0.1">0.1 mm</quantity>
for
<figureCitation id="74ED2A30FFE8737DFC17F88DFBFBF8B9" box="[979,1123,1793,1814]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="9" pageNumber="244">Figs 21, 2326</figureCitation>
</paragraph>
<caption id="B8A9663DFFE9737CFEF9F9F5FCE6F8AD" ID-DOI="http://doi.org/10.5281/zenodo.12585056" ID-Zenodo-Dep="12585056" httpUri="https://zenodo.org/record/12585056/files/figure.png" pageId="8" pageNumber="243" startId="8.[317,360,1657,1678]" targetBox="[337,1261,459,1591]" targetPageId="8" targetType="figure">
<paragraph id="EC6936B5FFE9737CFEF9F9F5FB6EF94A" blockId="8.[317,1270,1657,1794]" pageId="8" pageNumber="243">
<emphasis id="DEA2EAA7FFE9737CFEF9F9F5FE2AF921" bold="true" box="[317,434,1657,1678]" pageId="8" pageNumber="243">Figs 1318.</emphasis>
<taxonomicName id="2BD64D36FFE9737CFE7EF9F6FCFEF920" authorityName="Papp" authorityYear="2008" box="[442,870,1657,1679]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="243" phylum="Arthropoda" rank="species" species="keniaca" status="sp. nov." subGenus="Afrochaetopodella">
<emphasis id="DEA2EAA7FFE9737CFE7EF9F6FDDFF920" box="[442,583,1658,1679]" italics="true" pageId="8" pageNumber="243">Chaetopodella</emphasis>
(
<emphasis id="DEA2EAA7FFE9737CFD9CF9F6FC93F920" box="[600,779,1658,1679]" italics="true" pageId="8" pageNumber="243">Afrochaetopodella</emphasis>
)
<emphasis id="DEA2EAA7FFE9737CFCDFF9F6FCFEF920" box="[795,870,1658,1679]" italics="true" pageId="8" pageNumber="243">keniaca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFE9737CFCAAF9F5FC3DF921" box="[878,933,1657,1678]" pageId="8" pageNumber="243" rank="species">sp. n.</taxonomicNameLabel>
, holotype male, postabdomen and genitalia. 13 = syntergosternite 68, ventral view, 14 = 5th sternite, ventral view, 15 = caudal process of sternite 5, broadest view, 16 = surstylus at broadest (a subanteral-sublateral outer view), 17 = postgonite, broadest (= lateral) view, 18 = apical part of the dorsal process of distiphallus, subdorsal view.
</paragraph>
<paragraph id="EC6936B5FFE9737CFD70F961FCE6F8AD" blockId="8.[317,1270,1657,1794]" box="[692,894,1773,1794]" pageId="8" pageNumber="243">Scale: 0.1 mm for all</paragraph>
</caption>
<caption id="B8A9663DFFE8737DFEF9F9FDFB6DF956" ID-DOI="http://doi.org/10.5281/zenodo.12585058" ID-Zenodo-Dep="12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="9" pageNumber="244" startId="9.[317,360,1649,1670]" targetBox="[320,1269,445,1600]" targetPageId="9" targetType="figure">
<paragraph id="EC6936B5FFE8737DFEF9F9FDFB6DF956" blockId="9.[317,1270,1649,1814]" pageId="9" pageNumber="244">
<emphasis id="DEA2EAA7FFE8737DFEF9F9FDFE2AF929" bold="true" box="[317,434,1649,1670]" pageId="9" pageNumber="244">Figs 1926.</emphasis>
<taxonomicName id="2BD64D36FFE8737DFE7EF9FDFCFCF929" authorityName="Papp" authorityYear="2008" box="[442,868,1649,1670]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov." subGenus="Afrochaetopodella">
<emphasis id="DEA2EAA7FFE8737DFE7EF9FDFDDFF929" box="[442,583,1649,1670]" italics="true" pageId="9" pageNumber="244">Chaetopodella</emphasis>
(
<emphasis id="DEA2EAA7FFE8737DFD9CF9FDFC93F929" box="[600,779,1649,1670]" italics="true" pageId="9" pageNumber="244">Afrochaetopodella</emphasis>
)
<emphasis id="DEA2EAA7FFE8737DFCDFF9FDFCFCF929" box="[795,868,1649,1670]" italics="true" pageId="9" pageNumber="244">reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFE8737DFCA9F9FDFC3CF929" box="[877,932,1649,1670]" pageId="9" pageNumber="244" rank="species">sp. n.</taxonomicNameLabel>
, holotype male, postabdomen and genitalia. 19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view. Scales: 0.2 mm
</paragraph>
</caption>
<paragraph id="EC6936B5FFEB737EFEF9FE38FC4CFD62" blockId="10.[317,1270,436,717]" pageId="10" pageNumber="245">
long, the medial one bearing a large thorn, middle one with medium-long setae and lateral processes with longer setae (
<figureCitation id="74ED2A30FFEB737EFE2BFE5DFDA0FE49" box="[495,568,465,486]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="10" pageNumber="245">Fig. 21</figureCitation>
). The more sclerotized (melanized) right part of the syntergosternite 68 (tergite 7 and a part of sternite 6) forms a narrow but long re-curved sclerotized process; dorsal part of the syntergosternite large, sternite 7 and sternite 8 almost completely segregated (
<figureCitation id="74ED2A30FFEB737EFBF4FD87FBE1FD8F" box="[1072,1145,523,544]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="10" pageNumber="245">Fig. 19</figureCitation>
). Epandrium (
<figureCitation id="74ED2A30FFEB737EFE80FDA4FE12FD92" box="[324,394,552,573]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="10" pageNumber="245">Fig. 22</figureCitation>
) with a large ventral caudal process, which bears 7 very long light setae (longest
<quantity id="2B2E9B50FFEB737EFB56FDA4FB73FD92" box="[1170,1259,552,573]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="245" unit="mm" value="0.12">0.12 mm</quantity>
); caudally to surstylar base 2 similar setae (
<figureCitation id="74ED2A30FFEB737EFD00FDC8FCA9FDF6" box="[708,817,580,601]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="10" pageNumber="245">Figs 20, 22</figureCitation>
). Apical half (medial surface of epandrium with numerous long setae (
<figureCitation id="74ED2A30FFEB737EFDD7FDEDFDC5FDD9" box="[531,605,609,630]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="10" pageNumber="245">Fig. 24</figureCitation>
). Surstylus (
<figureCitation id="74ED2A30FFEB737EFD13FDEDFC86FDD9" box="[727,798,609,630]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="10" pageNumber="245">Fig. 23</figureCitation>
) rather compact with a number of processes but without long setae. Postgonite (
<figureCitation id="74ED2A30FFEB737EFDA2FDF2FD32FD3C" box="[614,682,638,659]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="10" pageNumber="245">Fig. 25</figureCitation>
) thick, rather long with sharp apex, a subapical posterior tooth and small swellings dorsally (posteriorly). Apical part of the dorsal process of distiphallus (
<figureCitation id="74ED2A30FFEB737EFB6DFD17FB77FD1F" box="[1193,1263,667,688]" captionStart="Figs 1926" captionStartId="9.[317,360,1649,1670]" captionTargetBox="[320,1269,445,1600]" captionTargetId="figure-10@9.[317,1269,445,1603]" captionTargetPageId="9" captionText="Figs 1926. Chaetopodella (Afrochaetopodella) reducta sp. n., holotype male, postabdomen and genitalia.19 = syntergosternite 68, ventral view, 20 = 5th sternite, ventral view, 21 = caudal process of sternite 5, broadest (sublateral) view; 22 = genitalia, lateral outer view, 23 = surstylus, broadest (a sublateral medial) view; 24 = apical half of epandrium, anterior view, 25 = postgonite, broadest (sublateral) view, 26 = apical part of the dorsal process of distiphallus, broadest view.Scales: 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12585058" httpUri="https://zenodo.org/record/12585058/files/figure.png" pageId="10" pageNumber="245">Fig. 26</figureCitation>
) with blunt, rather short fork apically, similar to that of
<taxonomicName id="2BD64D36FFEB737EFC9DFD34FC56FD62" authorityName="Papp" authorityYear="2008" box="[857,974,696,717]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="keniaca">
<emphasis id="DEA2EAA7FFEB737EFC9DFD34FC56FD62" box="[857,974,696,717]" italics="true" pageId="10" pageNumber="245">Ch. keniaca</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="EC6936B5FFEB737EFEBAFD78FD28FC9E" blockId="10.[317,1269,756,817]" pageId="10" pageNumber="245">Remark. There are no more data on its locality label. It seems probable that it was captured on elephant dung.</paragraph>
<paragraph id="EC6936B5FFEB737EFEBAFCD8FB40FCC6" blockId="10.[382,1240,852,873]" box="[382,1240,852,873]" pageId="10" pageNumber="245">Etymology. The specific epithet refers to the reduced frontal chaetotaxy of this species.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,215 @@
<document id="F2558F44868FCCD7920995CB032C3ED9" ID-DOI="10.5281/zenodo.12585046" ID-ISSN="2064-2474" ID-Zenodo-Dep="12585046" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719593661524" checkinUser="felipe" docAuthor="Papp, L." docDate="2008" docId="647F87A3FFEB7378FD9CFB8DFB79FB19" docLanguage="en" docName="ActaZoolAcadSciH.54.3.235-255.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 54 (3)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Chaetopodella aethiopica Papp, 2008, sp. n." docType="treatment" docVersion="2" lastPageNumber="247" masterDocId="9846FFDBFFE17374FFC4FF8CFF98FFAF" masterDocTitle="Afrotropical Species Of Chaetopodella Duda (Diptera: Sphaeroceridae)" masterLastPageNumber="255" masterPageNumber="235" pageNumber="245" updateTime="1719595852930" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="A655AEC6B10075FDE5C5412B0C4AF021">Afrotropical Species Of Chaetopodella Duda (Diptera: Sphaeroceridae)</mods:title>
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<subSubSection id="A4CC653EFFEB737EFD9CFB8DFC43FBB4" box="[600,987,1025,1051]" pageId="10" pageNumber="245" type="nomenclature">
<paragraph id="EC6936B5FFEB737EFD9CFB8DFC43FBB4" blockId="10.[600,987,1025,1051]" box="[600,987,1025,1051]" pageId="10" pageNumber="245">
<heading id="B72181D9FFEB737EFD9CFB8DFC43FBB4" box="[600,987,1025,1051]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="245" reason="2">
<taxonomicName id="2BD64D36FFEB737EFD9CFB8DFC0DFBB4" authorityName="Papp" authorityYear="2008" box="[600,917,1025,1051]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="aethiopica" status="sp. nov.">
<emphasis id="DEA2EAA7FFEB737EFD9CFB8DFC0DFBB4" bold="true" box="[600,917,1025,1051]" pageId="10" pageNumber="245">Chaetopodella aethiopica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFEB737EFC58FB8DFC43FBB4" box="[924,987,1025,1051]" pageId="10" pageNumber="245" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
</subSubSection>
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<paragraph id="EC6936B5FFEB737EFD0AFBA8FCFCFB91" blockId="10.[718,868,1060,1086]" box="[718,868,1060,1086]" pageId="10" pageNumber="245">
<heading id="B72181D9FFEB737EFD0AFBA8FCFCFB91" box="[718,868,1060,1086]" centered="true" fontSize="11" level="3" pageId="10" pageNumber="245" reason="3">
(
<figureCitation id="74ED2A30FFEB737EFD12FBA8FCC5FB91" box="[726,861,1060,1086]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="10" pageNumber="245">Figs 2733</figureCitation>
)
</heading>
</paragraph>
<paragraph id="EC6936B5FFEB737EFEBAFBE4FC8EFB19" blockId="10.[317,1270,1128,1206]" pageId="10" pageNumber="245">
<materialsCitation id="5CBE3CE8FFEB737EFEBAFBE4FB70FBD4" box="[382,1256,1128,1149]" collectingDate="1980-10-12" collectionCode="HNHM" country="Ethiopia" location="Lake Langano" pageId="10" pageNumber="245" specimenCount="1" typeStatus="holotype">
<typeStatus id="336D8817FFEB737EFEBAFBE4FE4FFBD2" box="[382,471,1128,1149]" pageId="10" pageNumber="245" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="8AC7AE70FFEB737EFDD8FBE4FDF4FBD2" box="[540,620,1128,1149]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="245">HNHM</collectionCode>
):
<collectingCountry id="94C17625FFEB737EFDB9FBE4FD49FBD2" box="[637,721,1128,1149]" name="Ethiopia" pageId="10" pageNumber="245">Ethiopia</collectingCountry>
,
<location id="E909606EFFEB737EFD1FFBE4FCFFFBD2" LSID="urn:lsid:plazi:treatment:647F87A3FFEB7378FD9CFB8DFB79FB19:E909606EFFEB737EFD1FFBE4FCFFFBD2" box="[731,871,1128,1149]" country="Ethiopia" name="Lake Langano" pageId="10" pageNumber="245">Lake Langano</location>
<date id="98681075FFEB737EFC46FBE4FC60FBD2" box="[898,1016,1128,1149]" pageId="10" pageNumber="245" value="1980-10-12">
<collectingDate id="882CE99DFFEB737EFC46FBE4FC60FBD2" box="[898,1016,1128,1149]" pageId="10" pageNumber="245" value="1980-10-12">12. X. 1980</collectingDate>
</date>
, leg. [András]
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.
<materialsCitation id="5CBE3CE8FFEB737EFEBAFB08FDA6FB36" box="[382,574,1156,1177]" collectionCode="HNHM" country="Ethiopia" pageId="10" pageNumber="245" specimenCount="1" typeStatus="paratype">
<typeStatus id="336D8817FFEB737EFEBAFB08FE43FB36" box="[382,475,1156,1177]" pageId="10" pageNumber="245" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="8AC7AE70FFEB737EFE23FB08FDAFFB36" box="[487,567,1156,1177]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="245">HNHM</collectionCode>
)
</materialsCitation>
:
<specimenCount id="FAD0FD3CFFEB737EFD82FB08FD03FB36" box="[582,667,1156,1177]" count="20" pageId="10" pageNumber="245" type="male">20 males</specimenCount>
<specimenCount id="FAD0FD3CFFEB737EFD65FB08FC98FB36" box="[673,768,1156,1177]" count="3" pageId="10" pageNumber="245" type="female">3 females</specimenCount>
: same as for
<typeStatus id="336D8817FFEB737EFCBEFB08FC56FB36" box="[890,974,1156,1177]" pageId="10" pageNumber="245" type="holotype">holotype</typeStatus>
[abdomen and genitalia of
<specimenCount id="FAD0FD3CFFEB737EFB16FB08FEEDFB19" count="2" pageId="10" pageNumber="245" type="male">two males</specimenCount>
in a plastic microvial each with glycerol].
</paragraph>
<paragraph id="EC6936B5FFEB737EFEBAFB57FB40FA86" blockId="10.[317,1270,1243,1811]" pageId="10" pageNumber="245">
Measurements in mm: body length 1.54 (
<typeStatus id="336D8817FFEB737EFCD7FB57FCF4FB5F" box="[787,876,1243,1264]" pageId="10" pageNumber="245" type="holotype">holotype</typeStatus>
), 1.311.60 (
<typeStatus id="336D8817FFEB737EFC28FB57FBD8FB5F" box="[1004,1088,1243,1264]" pageId="10" pageNumber="245" type="paratype">paratype</typeStatus>
males), 1.591.87 (
<typeStatus id="336D8817FFEB737EFE87FB74FE0FFAA2" box="[323,407,1272,1293]" pageId="10" pageNumber="245" type="paratype">paratype</typeStatus>
females), wing length 1.41 (
<typeStatus id="336D8817FFEB737EFD68FB74FC9DFAA2" box="[684,773,1272,1293]" pageId="10" pageNumber="245" type="holotype">holotype</typeStatus>
), 1.15 1.48 (
<typeStatus id="336D8817FFEB737EFC4CFB74FC44FAA2" box="[904,988,1272,1293]" pageId="10" pageNumber="245" type="paratype">paratype</typeStatus>
males), 1.48 1.67 (
<typeStatus id="336D8817FFEB737EFB65FB74FB6DFAA2" box="[1185,1269,1272,1293]" pageId="10" pageNumber="245" type="paratype">paratype</typeStatus>
females), wing width 0.54 (
<typeStatus id="336D8817FFEB737EFDFAFA98FD0FFA86" box="[574,663,1300,1321]" pageId="10" pageNumber="245" type="holotype">holotype</typeStatus>
), 0.46 0.63 (
<typeStatus id="336D8817FFEB737EFCD0FA98FCF0FA86" box="[788,872,1300,1321]" pageId="10" pageNumber="245" type="paratype">paratype</typeStatus>
males), 0.57 0.68 (
<typeStatus id="336D8817FFEB737EFBE2FA98FBE2FA86" box="[1062,1146,1300,1321]" pageId="10" pageNumber="245" type="paratype">paratype</typeStatus>
females).
</paragraph>
<paragraph id="EC6936B5FFEB737EFEBAFABDFB53FA79" blockId="10.[317,1270,1243,1811]" pageId="10" pageNumber="245">
Head finely microtrichose, subshiny; silvery spots around bases of macrochaetae. Anterior 1/32/5 of frons reddish brown; ocellar triangle with the rest of frons dark brown. Gena pale yellow in male and reddish brown in female, face pale yellow in male and brown to dark brown in female; gena with dark minute setae directed downwards, they are more stronger in male; anterior
<emphasis id="DEA2EAA7FFEB737EFB9EFA04FBE9FA32" box="[1114,1137,1416,1437]" italics="true" pageId="10" pageNumber="245">ifr</emphasis>
weak; middle and hind pairs medium-long. Posterior
<emphasis id="DEA2EAA7FFEB737EFD7DFA29FD40FA15" box="[697,728,1445,1466]" italics="true" pageId="10" pageNumber="245">ors</emphasis>
1.75 to 2.0 times longer than anterior
<emphasis id="DEA2EAA7FFEB737EFB89FA29FBF4FA15" box="[1101,1132,1445,1466]" italics="true" pageId="10" pageNumber="245">ors</emphasis>
; eye oval, antenna dark brown, arista long, somewhat less than 4 times as long as antenna, shortly ciliate.
</paragraph>
<paragraph id="EC6936B5FFEB737EFEBAFA52FDDEF90F" blockId="10.[317,1270,1243,1811]" pageId="10" pageNumber="245">
Thorax: Most parts of mesonotum silvery microtomentose, i.e. silvery spots around setal bases united (confluent) in 4 broad stripes, which leave velvety black only 3 longitudinal stripes plus a pre-scutellar area. 6 rows of acrostichal microsetae in front of suture, 3 pairs of rather strong
<emphasis id="DEA2EAA7FFEB737EFB1DF994FB68F982" box="[1241,1264,1560,1581]" italics="true" pageId="10" pageNumber="245">dc</emphasis>
. Pleura lighter grey or yellowish grey, strongly microtrichose; pale yellowish line on notopleura; 2 katepisternals, posterior one very long
<quantity id="2B2E9B50FFEB737EFD77F9DDFC93F9C9" box="[691,779,1617,1638]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="10" pageNumber="245" unit="mm" value="0.33">0.33 mm</quantity>
, anterior pair weak, only
<quantity id="2B2E9B50FFEB737EFBC4F9DDFB16F9C9" box="[1024,1166,1617,1638]" metricMagnitude="-4" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="10" pageNumber="245" unit="mm" value="0.105" valueMax="0.11" valueMin="0.1">0.100.11 mm</quantity>
; scutellum velvety black, somewhat wider than long; 2
<emphasis id="DEA2EAA7FFEB737EFD34F9E2FC9CF92C" box="[752,772,1646,1667]" italics="true" pageId="10" pageNumber="245">sc</emphasis>
long, apical one 1.81.85 and lateral one 1.11.15 times as long as scutellum.
</paragraph>
<paragraph id="EC6936B5FFEB737FFEBAF924FC70F881" blockId="10.[317,1270,1243,1811]" lastBlockId="11.[317,1271,1702,1838]" lastPageId="11" lastPageNumber="246" pageId="10" pageNumber="245">
Wing: Yellowish, veins yellowish to ochreous; costal vein somewhat darker, not extended beyond apex of R
<subScript id="705234F0FFEB737EFE16F95DFE7EF972" attach="left" box="[466,486,1745,1757]" fontSize="5" pageId="10" pageNumber="245">4+5</subScript>
, sub-basally with a very long (
<quantity id="2B2E9B50FFEB737EFCD5F948FC38F976" box="[785,928,1732,1753]" metricMagnitude="-4" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="10" pageNumber="245" unit="mm" value="0.145" valueMax="0.15" valueMin="0.14">0.140.15 mm</quantity>
) seta, first sector with much longer setae than those of second and third sectors; R
<subScript id="705234F0FFEB737EFCC8F962FCB9F955" attach="left" box="[780,801,1774,1786]" fontSize="5" pageId="10" pageNumber="245">4+5</subScript>
distinctly bent up to C; C-index (Cs2: Cs3) = 1.211.30; R-M dM-Cu: dM-Cu = about 2.5; lower corner of the discal cell angulate with a short
<quantity id="2B2E9B50FFEA737FFD8EF895FD0BF881" box="[586,659,1817,1838]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="246" unit="mm" value="0.2">0.2 mm</quantity>
for
<figureCitation id="74ED2A30FFEA737FFD78F895FC9AF881" box="[700,770,1817,1838]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="11" pageNumber="246">Fig. 27</figureCitation>
,
<quantity id="2B2E9B50FFEA737FFCC8F895FCCDF881" box="[780,853,1817,1838]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="246" unit="mm" value="0.1">0.1 mm</quantity>
for
<figureCitation id="74ED2A30FFEA737FFCB9F895FC70F881" box="[893,1000,1817,1838]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="11" pageNumber="246">Figs 2833</figureCitation>
</paragraph>
<caption id="B8A9663DFFEA737FFEF9F92AFB6FF8BD" ID-DOI="http://doi.org/10.5281/zenodo.12585060" ID-Zenodo-Dep="12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="11" pageNumber="246" startId="11.[317,360,1702,1723]" targetBox="[351,1232,427,1670]" targetPageId="11" targetType="figure">
<paragraph id="EC6936B5FFEA737FFEF9F92AFB6FF8BD" blockId="11.[317,1271,1702,1838]" pageId="11" pageNumber="246">
<emphasis id="DEA2EAA7FFEA737FFEF9F92AFE36F914" bold="true" box="[317,430,1702,1723]" pageId="11" pageNumber="246">Figs 2733.</emphasis>
<taxonomicName id="2BD64D36FFEA737FFE77F92AFD78F914" authorityName="Papp" authorityYear="2008" box="[435,736,1702,1723]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="246" phylum="Arthropoda" rank="species" species="aethiopica" status="sp. nov." subGenus="Chaetopodella">
<emphasis id="DEA2EAA7FFEA737FFE77F92AFDD8F914" box="[435,576,1702,1723]" italics="true" pageId="11" pageNumber="246">Chaetopodella</emphasis>
(
<emphasis id="DEA2EAA7FFEA737FFD88F92AFDF7F914" box="[588,623,1702,1723]" italics="true" pageId="11" pageNumber="246">Ch.</emphasis>
)
<emphasis id="DEA2EAA7FFEA737FFDBFF92AFD78F914" box="[635,736,1702,1723]" italics="true" pageId="11" pageNumber="246">aethiopica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFEA737FFD20F92AFC80F914" box="[740,792,1702,1723]" pageId="11" pageNumber="246" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, genitalia. 27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:
</paragraph>
</caption>
<paragraph id="EC6936B5FFED7378FEF9FE38FC70FE49" blockId="12.[317,1271,436,1149]" pageId="12" pageNumber="247">
vein appendage, or, in some specimens rounded without vein appendage (see below). Alula large, broad (up to
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) and rounded. Halter white to light yellowish.
</paragraph>
<paragraph id="EC6936B5FFED7378FEBAFE62FD01FD46" blockId="12.[317,1271,436,1149]" pageId="12" pageNumber="247">
Legs: Yellowish brown to dark brown (generally darker in female). Mid trochanteral seta
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long. Male fore coxa on anterior surface with 5 (6) medium-long setae (and without dense long hairs). Male fore tibia simple, fore tarsus not darkened and not flattened, tarsomeres without long dense lateral setae. Fore femur simple; male mid femur without a ventral row of setae basally. Mid tibia with 3 dorsal setae on basal 1/4, 1/3 and 3/4, 1 rather strong anterodorsal seta on 2/3, 23 weak posterodorsal setae on 1/3 1/2, rather strong posterodorsal seta on 2/3, 1 strong apicoventral seta. A ventral seta in female at distal 17/40 of tibia (this ventral seta developed also in some males). Mid metatarsal seta very strong,
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on a
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. Second hind tarsomeres swollen,
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long and
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thick.
</paragraph>
<paragraph id="EC6936B5FFED7378FEBAFD7DFBACFCF2" blockId="12.[317,1271,436,1149]" pageId="12" pageNumber="247">
Abdomen: Preabdominal sternites 24 quadratic. Tergite 5 very small, less than
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broad and less than
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long. Sternite 5 (
<figureCitation id="74ED2A30FFED7378FD33FC82FCA5FC8C" box="[759,829,782,803]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="12" pageNumber="247">Fig. 28</figureCitation>
) comparatively long, caudal pair of processes medium-long slightly medially curved and bare, symmetrical with 2 pairs of lateral thicker setae, 1 pair of blunt setae and 2 pairs of smaller medial setae. Medial platelet pilose.
</paragraph>
<paragraph id="EC6936B5FFED7378FEBAFCE8FB4BFB89" blockId="12.[317,1271,436,1149]" pageId="12" pageNumber="247">
Syntergosternite 68 (
<figureCitation id="74ED2A30FFED7378FD90FCE8FD03FCD6" box="[596,667,868,889]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="12" pageNumber="247">Fig. 27</figureCitation>
) comparatively short. S8 part of the complex short (as for the body axis), right part with intricate sclerotisation as given on the figure. Male cerci membranous, microsetose only. Subanal plate (
<figureCitation id="74ED2A30FFED7378FDB3FC12FD23FC1C" box="[631,699,926,947]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="12" pageNumber="247">Fig. 29</figureCitation>
) with one pair of a medial medium-long and one pair of longer lateral thorn-like setae. Surstylus (
<figureCitation id="74ED2A30FFED7378FD6FFC37FC81FC7F" box="[683,793,955,976]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="12" pageNumber="247">Figs 3031</figureCitation>
) comparatively simple, lateral process with some medium-long setae on the outer surface; basal lobe large. Distiphallus short and thick. Epiphallus hook-like. Postgonite (
<figureCitation id="74ED2A30FFED7378FDDEFC78FDF9FBA6" box="[538,609,1012,1033]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="12" pageNumber="247">Fig. 32</figureCitation>
) with much narrowed though not sharp apex. Phallapodeme robust. Apical part of the paired dorsal process of distiphallus with long rather thin fork apically (
<figureCitation id="74ED2A30FFED7378FB46FB9DFB50FB89" box="[1154,1224,1041,1062]" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="12" pageNumber="247">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="EC6936B5FFED7378FEBAFBA2FC98FBD2" blockId="12.[317,1271,436,1149]" pageId="12" pageNumber="247">
Also female tergites black. Lateral caudal setae on female abdominal tergites rather long, e.g.
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on tergite 6. Membrane together with sternites 47 yellow. Cerci short with a pair of
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long wavy bent hair-like setae.
</paragraph>
<paragraph id="EC6936B5FFED7378FEBAFB2DFB79FB19" blockId="12.[382,1249,1185,1206]" box="[382,1249,1185,1206]" pageId="12" pageNumber="247">
Etymology. The specific epithet refers to the Latin name of its
<typeStatus id="336D8817FFED7378FC2FFB2DFB8CFB19" box="[1003,1044,1185,1206]" pageId="12" pageNumber="247">type</typeStatus>
locality Aethiopia.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,267 @@
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<taxonomicName id="2BD64D36FFED7378FDA2FA8BFC1FFA8E" authorityName="Papp" authorityYear="2008" box="[614,903,1287,1313]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="nigeriae" status="sp. nov.">
<emphasis id="DEA2EAA7FFED7378FDA2FA8BFC1FFA8E" bold="true" box="[614,903,1287,1313]" pageId="12" pageNumber="247">Chaetopodella nigeriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFED7378FC4AFA8BFC55FA8E" box="[910,973,1287,1313]" pageId="12" pageNumber="247" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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(
<figureCitation id="74ED2A30FFED7378FD12FAA7FCC4FAEA" box="[726,860,1323,1349]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="12" pageNumber="247">Figs 3440</figureCitation>
)
</heading>
</paragraph>
<paragraph id="EC6936B5FFED7378FEBAFAE2FCB3FA0F" blockId="12.[317,1271,1390,1555]" pageId="12" pageNumber="247">
<materialsCitation id="5CBE3CE8FFED7378FEBAFAE2FCB3FA0F" collectingDate="1978-08-14" collectionCode="HNHM" country="Nigeria" location="Yanguikari Reserve" municipality="Wikki" pageId="12" pageNumber="247" specimenCount="1" typeStatus="holotype">
<typeStatus id="336D8817FFED7378FEBAFAE2FE4FFA2C" box="[382,471,1390,1411]" pageId="12" pageNumber="247" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="8AC7AE70FFED7378FDD2FAE2FDFEFA2C" box="[534,614,1390,1411]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="12" pageNumber="247">HNHM</collectionCode>
):
<collectingCountry id="94C17625FFED7378FDB0FAE2FD26FA2C" box="[628,702,1390,1411]" name="Nigeria" pageId="12" pageNumber="247">Nigeria</collectingCountry>
,
<location id="E909606EFFED7378FD03FAE2FC10FA2C" LSID="urn:lsid:plazi:treatment:647F87A3FFED737AFDA2FA8BFBCAFACC:E909606EFFED7378FD03FAE2FC10FA2C" box="[711,904,1390,1411]" country="Nigeria" municipality="Wikki" name="Yanguikari Reserve" pageId="12" pageNumber="247">Yanguikari Reserve</location>
,
<collectingMunicipality id="0C0DACCFFFED7378FC54FAE2FC54FA2C" box="[912,972,1390,1411]" pageId="12" pageNumber="247">Wikki</collectingMunicipality>
<date id="98681075FFED7378FC25FAE2FBF2FA2C" box="[993,1130,1390,1411]" pageId="12" pageNumber="247" value="1978-08-14">
<collectingDate id="882CE99DFFED7378FC25FAE2FBF2FA2C" box="[993,1130,1390,1411]" pageId="12" pageNumber="247" value="1978-08-14">Aug. 14, 1978</collectingDate>
</date>
, leg. A. DEMETER, No. 18 [hippo dung from the previous night].
</materialsCitation>
</paragraph>
<paragraph id="EC6936B5FFED7378FEBAFA24FE31F9BC" blockId="12.[317,1271,1390,1555]" pageId="12" pageNumber="247">
<materialsCitation id="5CBE3CE8FFED7378FEBAFA24FBA6FA12" box="[382,1086,1448,1469]" collectionCode="HNHM" country="Nigeria" county="Wikki" location="Yangui" municipality="Reserve" pageId="12" pageNumber="247" specimenCount="1" typeStatus="paratype">
<typeStatus id="336D8817FFED7378FEBAFA24FE43FA12" box="[382,475,1448,1469]" pageId="12" pageNumber="247" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="8AC7AE70FFED7378FE23FA24FDAFFA12" box="[487,567,1448,1469]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="12" pageNumber="247">HNHM</collectionCode>
):
<collectingCountry id="94C17625FFED7378FD81FA24FD17FA12" box="[581,655,1448,1469]" name="Nigeria" pageId="12" pageNumber="247">Nigeria</collectingCountry>
,
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[correctly: Yankari]
<collectingMunicipality id="0C0DACCFFFED7378FC62FA24FC6EFA12" box="[934,1014,1448,1469]" pageId="12" pageNumber="247">Reserve</collectingMunicipality>
,
<collectingCounty id="05084E39FFED7378FC3AFA24FBA6FA12" box="[1022,1086,1448,1469]" pageId="12" pageNumber="247">Wikki</collectingCounty>
</materialsCitation>
:
<specimenCount id="FAD0FD3CFFED7378FB82FA24FB17FA12" box="[1094,1167,1448,1469]" count="7" pageId="12" pageNumber="247" type="male">7 males</specimenCount>
<specimenCount id="FAD0FD3CFFED7378FB57FA24FB6AFA12" box="[1171,1266,1448,1469]" count="4" pageId="12" pageNumber="247" type="female">4 females</specimenCount>
: same as for
<typeStatus id="336D8817FFED7378FE68FA48FD9BFA76" box="[428,515,1476,1497]" pageId="12" pageNumber="247" type="holotype">holotype</typeStatus>
;
<materialsCitation id="5CBE3CE8FFED7378FDCFFA48FC8AFA59" collectionCode="HNHM" country="Nigeria" county="Wikki" location="Yangui" municipality="Reserve" pageId="12" pageNumber="247" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FAD0FD3CFFED7378FDCFFA48FDD3FA76" box="[523,587,1476,1497]" count="1" pageId="12" pageNumber="247" type="male">1 male</specimenCount>
<specimenCount id="FAD0FD3CFFED7378FD8BFA48FD36FA76" box="[591,686,1476,1497]" count="2" pageId="12" pageNumber="247" type="female">2 females</specimenCount>
: ibid., Aug. 11, No. 5 [2 to 3 days old buffalo dung] [abdomen and genitalia in a plastic microvial with glycerol]
</materialsCitation>
;
<materialsCitation id="5CBE3CE8FFED7378FCDEFA6DFE3DF9BC" collectionCode="HNHM" country="Nigeria" county="Wikki" location="Yangui" municipality="Reserve" pageId="12" pageNumber="247" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FAD0FD3CFFED7378FCDEFA6DFCC5FA59" box="[794,861,1505,1526]" count="1" pageId="12" pageNumber="247" type="male">1 male</specimenCount>
: ibid., Aug. 12, No. 12 [3 to 4 days old buffalo dung]
</materialsCitation>
.
</paragraph>
<paragraph id="EC6936B5FFED7378FEBAF9B4FB68F929" blockId="12.[317,1270,1592,1814]" pageId="12" pageNumber="247">
Measurements in mm: body length 1.32 (
<typeStatus id="336D8817FFED7378FCCFF9B4FCFCF9E2" box="[779,868,1592,1613]" pageId="12" pageNumber="247" type="holotype">holotype</typeStatus>
), 1.26 1.48 (
<typeStatus id="336D8817FFED7378FC2BF9B4FBDBF9E2" box="[1007,1091,1592,1613]" pageId="12" pageNumber="247" type="paratype">paratype</typeStatus>
males), 1.271.59 (
<typeStatus id="336D8817FFED7378FE87F9D8FE39F9C6" box="[323,417,1620,1641]" pageId="12" pageNumber="247" type="paratype">paratypes</typeStatus>
females), wing length 1.21 (
<typeStatus id="336D8817FFED7378FD73F9D8FC88F9C6" box="[695,784,1620,1641]" pageId="12" pageNumber="247" type="holotype">holotype</typeStatus>
), 1.181.40 (
<typeStatus id="336D8817FFED7378FC49F9D8FC79F9C6" box="[909,993,1620,1641]" pageId="12" pageNumber="247" type="paratype">paratype</typeStatus>
males), 1.211.48 (
<typeStatus id="336D8817FFED7378FB65F9D8FB6DF9C6" box="[1185,1269,1620,1641]" pageId="12" pageNumber="247" type="paratype">paratype</typeStatus>
females), wing width 0.44 (
<typeStatus id="336D8817FFED7378FD8FF9FDFD3CF929" box="[587,676,1649,1670]" pageId="12" pageNumber="247" type="holotype">holotype</typeStatus>
), 0.430.48 (
<typeStatus id="336D8817FFED7378FCE0F9FDFCE0F929" box="[804,888,1649,1670]" pageId="12" pageNumber="247" type="paratype">paratype</typeStatus>
males), 0.470.55 (
<typeStatus id="336D8817FFED7378FBFFF9FDFB17F929" box="[1083,1167,1649,1670]" pageId="12" pageNumber="247" type="paratype">paratype</typeStatus>
females).
</paragraph>
<paragraph id="EC6936B5FFED7379FEBAF902FC71F881" blockId="12.[317,1270,1592,1814]" lastBlockId="13.[317,1271,1702,1838]" lastPageId="13" lastPageNumber="248" pageId="12" pageNumber="247">
Head: Mostly blackish brown, heavily microtrichose. Frons entirely dark or reddish only on a narrow anterior part. There are silvery spots around bases of macrochaetae. Gena and face brown, in some specimens yellowish brown. Three medium-long interfrontal pairs present, upper setae slightly stronger. Anterior
<emphasis id="DEA2EAA7FFED7378FE2AF969FD95F955" box="[494,525,1765,1786]" italics="true" pageId="12" pageNumber="247">ors</emphasis>
thin, 0.065
<quantity id="2B2E9B50FFED7378FDBAF968FD4EF956" box="[638,726,1764,1785]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="12" pageNumber="247" unit="mm" value="0.07">0.07 mm</quantity>
, about 1/2 length of posterior
<emphasis id="DEA2EAA7FFED7378FC37F969FB8AF955" box="[1011,1042,1765,1786]" italics="true" pageId="12" pageNumber="247">ors</emphasis>
(
<quantity id="2B2E9B50FFED7378FBDAF968FB59F956" box="[1054,1217,1764,1785]" metricMagnitude="-5" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="12" pageNumber="247" unit="mm" value="0.014499999999999999" valueMax="0.015" valueMin="0.014">
<geoCoordinate id="89E25072FFED7378FBDAF968FBCEF956" box="[1054,1110,1764,1785]" degrees="0.014" direction="north" orientation="latitude" pageId="12" pageNumber="247" precision="55" value="0.014">0.014</geoCoordinate>
<geoCoordinate id="89E25072FFED7378FB92F968FB02F956" box="[1110,1178,1764,1785]" degrees="0.015" direction="west" orientation="longitude" pageId="12" pageNumber="247" precision="55" value="-0.015">0.015</geoCoordinate>
mm
</quantity>
). Eye large and rounded, its longest diameter (
<quantity id="2B2E9B50FFED7378FD7FF88DFC81F8B9" box="[699,793,1793,1814]" metricMagnitude="-4" metricUnit="m" metricValue="2.8499999999999996" pageId="12" pageNumber="247" unit="mm" value="0.285">0.285mm</quantity>
) about 3.5 times as long as genal width below eye
<quantity id="2B2E9B50FFEC7379FD8EF895FD0BF881" box="[586,659,1817,1838]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="248" unit="mm" value="0.2">0.2 mm</quantity>
for
<figureCitation id="74ED2A30FFEC7379FD78F895FC9AF881" box="[700,770,1817,1838]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="13" pageNumber="248">Fig. 34</figureCitation>
,
<quantity id="2B2E9B50FFEC7379FCC8F895FCCDF881" box="[780,853,1817,1838]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="248" unit="mm" value="0.1">0.1 mm</quantity>
for
<figureCitation id="74ED2A30FFEC7379FCB9F895FC71F881" box="[893,1001,1817,1838]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="13" pageNumber="248">Figs 3540</figureCitation>
</paragraph>
<caption id="B8A9663DFFEC7379FEF9F92AFB6FF8BE" ID-DOI="http://doi.org/10.5281/zenodo.12585062" ID-Zenodo-Dep="12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="13" pageNumber="248" startId="13.[317,360,1702,1723]" targetBox="[327,1265,480,1652]" targetPageId="13" targetType="figure">
<paragraph id="EC6936B5FFEC7379FEF9F92AFB6FF8BE" blockId="13.[317,1271,1702,1838]" pageId="13" pageNumber="248">
<emphasis id="DEA2EAA7FFEC7379FEF9F92AFE28F914" bold="true" box="[317,432,1702,1723]" pageId="13" pageNumber="248">Figs 3440.</emphasis>
<taxonomicName id="2BD64D36FFEC7379FE72F92AFD49F914" authorityName="Papp" authorityYear="2008" box="[438,721,1702,1723]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="nigeriae" status="sp. nov." subGenus="Chaetopodella">
<emphasis id="DEA2EAA7FFEC7379FE72F92AFDDBF914" box="[438,579,1702,1723]" italics="true" pageId="13" pageNumber="248">Chaetopodella</emphasis>
(
<emphasis id="DEA2EAA7FFEC7379FD95F92AFDECF914" box="[593,628,1702,1723]" italics="true" pageId="13" pageNumber="248">Ch.</emphasis>
)
<emphasis id="DEA2EAA7FFEC7379FD45F92AFD49F914" box="[641,721,1702,1723]" italics="true" pageId="13" pageNumber="248">nigeriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFEC7379FD13F92AFC95F914" box="[727,781,1702,1723]" pageId="13" pageNumber="248" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:
</paragraph>
</caption>
<paragraph id="EC6936B5FFEF737AFEF9FE38FDB5FE49" blockId="14.[317,1271,436,1321]" pageId="14" pageNumber="249">
(
<quantity id="2B2E9B50FFEF737AFE80FE38FE31FE66" box="[324,425,436,457]" metricMagnitude="-5" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="249" unit="mm" value="0.077">0.077 mm</quantity>
). Antenna dark brown, incl. scape, arista long, almost 4 times as long as antenna (0.59 vs.
<quantity id="2B2E9B50FFEF737AFEF9FE5DFE0FFE49" box="[317,407,465,486]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="249" unit="mm" value="0.15">0.15 mm</quantity>
, shortly ciliate.
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAFE62FC58FD3C" blockId="14.[317,1271,436,1321]" pageId="14" pageNumber="249">
Thorax: Mesonotal colour dominated by velvety black, i.e. silvery microtomentum only around setal bases and this silvery colour confluent only in 4 short stripes, which terminate at the level of posterior notopleural seta. Six rows of acrostichal microsetae in front of suture; 3
<emphasis id="DEA2EAA7FFEF737AFB4FFDA4FB3AFD92" box="[1163,1186,552,573]" italics="true" pageId="14" pageNumber="249">dc</emphasis>
; 2
<emphasis id="DEA2EAA7FFEF737AFB7AFDA4FB77FD92" box="[1214,1263,552,573]" italics="true" pageId="14" pageNumber="249">kepst</emphasis>
, posterior one very strong (
<quantity id="2B2E9B50FFEF737AFD8CFDC8FD44FDF6" box="[584,732,580,601]" metricMagnitude="-4" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="14" pageNumber="249" unit="mm" value="0.165" valueMax="0.17" valueMin="0.16">0.160.17 mm</quantity>
). Scutellum velvety black, somewhat wider than long (
<quantity id="2B2E9B50FFEF737AFE80FDEDFE01FDD9" box="[324,409,609,630]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="14" pageNumber="249" unit="mm" value="0.33">0.33 mm</quantity>
vs.
<quantity id="2B2E9B50FFEF737AFE7AFDEDFD86FDD9" box="[446,542,609,630]" metricMagnitude="-4" metricUnit="m" metricValue="2.8499999999999996" pageId="14" pageNumber="249" unit="mm" value="0.285">0.285 mm</quantity>
on
<typeStatus id="336D8817FFEF737AFD84FDEDFD00FDD9" box="[576,664,609,630]" pageId="14" pageNumber="249" type="holotype">holotype</typeStatus>
); 2
<emphasis id="DEA2EAA7FFEF737AFD7CFDEDFD54FDD9" box="[696,716,609,630]" italics="true" pageId="14" pageNumber="249">sc</emphasis>
long, apical one (
<quantity id="2B2E9B50FFEF737AFCB7FDEDFC4CFDD9" box="[883,980,609,630]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" pageId="14" pageNumber="249" unit="mm" value="0.505">0.505 mm</quantity>
on
<typeStatus id="336D8817FFEF737AFC32FDEDFBD4FDD9" box="[1014,1100,609,630]" pageId="14" pageNumber="249" type="holotype">holotype</typeStatus>
) nearly 2 times as long as lateral one (
<quantity id="2B2E9B50FFEF737AFE3BFDF2FDC1FD3C" box="[511,601,638,659]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="14" pageNumber="249" unit="mm" value="0.26">0.26 mm</quantity>
) and slightly shorter than scutellum.
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAFD17FB48FCEF" blockId="14.[317,1271,436,1321]" pageId="14" pageNumber="249">
Wing: Milky whitish, veins whitish except for light brown C; C not extended beyond apex of R
<subScript id="705234F0FFEF737AFE89FD48FEFAFD7F" attach="left" box="[333,354,708,720]" fontSize="5" pageId="14" pageNumber="249">4+5</subScript>
. First costal section setae
<quantity id="2B2E9B50FFEF737AFDA5FD34FD2FFD62" box="[609,695,696,717]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="249" unit="mm" value="0.09">0.09 mm</quantity>
long, i.e. distinctly stronger setae than those of second and third sectors and longer than cell diameter. Apical part of R
<subScript id="705234F0FFEF737AFC4CFD6DFC05FD42" attach="left" box="[904,925,737,749]" fontSize="5" pageId="14" pageNumber="249">4+5</subScript>
rather strongly bent up to C. C-index = 0.40/
<quantity id="2B2E9B50FFEF737AFE63FD7DFD9EFCA9" box="[423,518,753,774]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" pageId="14" pageNumber="249" unit="mm" value="0.225">0.225 mm</quantity>
1.77 (
<typeStatus id="336D8817FFEF737AFDFBFD7DFD0EFCA9" box="[575,662,753,774]" pageId="14" pageNumber="249" type="holotype">holotype</typeStatus>
) 1.691.78 (
<typeStatus id="336D8817FFEF737AFCC8FD7DFCF6FCA9" box="[780,878,753,774]" pageId="14" pageNumber="249" type="paratype">paratypes</typeStatus>
); R-M dM-Cu: dM-Cu = 3.4 (
<typeStatus id="336D8817FFEF737AFB56FD7DFB73FCA9" box="[1170,1259,753,774]" pageId="14" pageNumber="249" type="holotype">holotype</typeStatus>
), 3.33.5 (
<typeStatus id="336D8817FFEF737AFE55FC82FE6BFC8C" box="[401,499,782,803]" pageId="14" pageNumber="249" type="paratype">paratypes</typeStatus>
); lower corner of the discal cell angulate with a short vein appendage, or, in some specimens rounded without vein appendage; alula broad and rounded; halter yellowish white.
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAFCC4FD70FC7F" blockId="14.[317,1271,436,1321]" pageId="14" pageNumber="249">
Legs brown; male fore tibia normal but tarsus slightly thickened. Male fore coxa on anterior surface with dense long (up to
<quantity id="2B2E9B50FFEF737AFD9AFCE8FD2CFCD6" box="[606,692,868,889]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="14" pageNumber="249" unit="mm" value="0.18">0.18 mm</quantity>
) light curly hairs. Posterior apex of fore coxa with 2 (3) similar but thicker and even longer (
<quantity id="2B2E9B50FFEF737AFDAEFC0DFD59FC39" box="[618,705,897,918]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="249" unit="mm" value="0.2">0.20 mm</quantity>
) hair-like setae. Female fore coxa more medially than anteriorly with similar light but shorter dense hairs. Fore femur with 23 dorsal setae. Mid femur without a ventral row of setae on basal 1/
<quantity id="2B2E9B50FFEF737AFD47FC37FD33FC7F" box="[643,683,955,976]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="14" pageNumber="249" unit="in" value="5.0">5 in</quantity>
male.
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAFC54FC22FBD2" blockId="14.[317,1271,436,1321]" pageId="14" pageNumber="249">
Abdomen: Male tergite 5 short, not divided sagittally, sclerotisation not weaker than laterally. Sternite 5 (
<figureCitation id="74ED2A30FFEF737AFE60FC78FE70FBA6" box="[420,488,1012,1033]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="14" pageNumber="249">Fig. 35</figureCitation>
) with robust paired caudal processes not far from each other, medially and apically with 3+2+1 setae. Medial platelet angulate and hairy (pilose). Syntergosternite 68 (
<figureCitation id="74ED2A30FFEF737AFBB5FB9DFB20FB89" box="[1137,1208,1041,1062]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="14" pageNumber="249">Fig. 34</figureCitation>
) short dorsally (as regards body axis), right lateral (T7) part complex. Ventralmost part of the complex comparatively weakly sclerotized. Ventral plate of subepandrial sclerite (
<figureCitation id="74ED2A30FFEF737AFC1AFBC7FBBDFBCF" box="[990,1061,1099,1120]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="14" pageNumber="249">Fig. 36</figureCitation>
) with 2 pairs of long thorn-like seta, the lateral one much thicker than the medial pair.
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAFB08FB43FB5F" blockId="14.[317,1271,436,1321]" pageId="14" pageNumber="249">
Surstylus (
<figureCitation id="74ED2A30FFEF737AFE26FB08FDD6FB36" box="[482,590,1156,1177]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="14" pageNumber="249">Figs 3738</figureCitation>
) with large lateral process, which bears some medium-long setae; basal process with 1 long and 2 short setae. Distiphallus robust, comparatively short, epiphallus triangular in profile. Postgonite (
<figureCitation id="74ED2A30FFEF737AFE33FB32FDA4FB7C" box="[503,572,1214,1235]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="14" pageNumber="249">Fig. 39</figureCitation>
) less narrowed in its apical part than that of
<taxonomicName id="2BD64D36FFEF737AFC19FB32FBFFFB7C" authorityName="Papp" authorityYear="2008" box="[989,1127,1214,1235]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="aethiopica">
<emphasis id="DEA2EAA7FFEF737AFC19FB32FBFFFB7C" box="[989,1127,1214,1235]" italics="true" pageId="14" pageNumber="249">Ch. aethiopica</emphasis>
</taxonomicName>
. Apical part of the paired dorsal process of distiphallus with a longer medial and a shorter lateral processes (
<figureCitation id="74ED2A30FFEF737AFB4EFB57FB57FB5F" box="[1162,1231,1243,1264]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="14" pageNumber="249">Fig. 40</figureCitation>
).
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAFB74FB7BFA86" blockId="14.[317,1271,436,1321]" pageId="14" pageNumber="249">
Female abdominal tergites with long setae on caudal lateral edges, longest on tergite 6,
<quantity id="2B2E9B50FFEF737AFB08FB74FEFDFA86" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="249" unit="mm" value="0.15">0.15 mm</quantity>
; cerci very short, only
<quantity id="2B2E9B50FFEF737AFD8DFA98FD07FA86" box="[585,671,1300,1321]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="249" unit="mm" value="0.08">0.08 mm</quantity>
long,
<quantity id="2B2E9B50FFEF737AFD1BFA98FCDAFA86" box="[735,834,1300,1321]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="249" unit="mm" value="0.035">0.035 mm</quantity>
broad, longest cercal setae
<quantity id="2B2E9B50FFEF737AFB97FA98FB31FA86" box="[1107,1193,1300,1321]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="249" unit="mm" value="0.08">0.08 mm</quantity>
long.
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAFAC2FBCAFACC" blockId="14.[382,1106,1358,1379]" box="[382,1106,1358,1379]" pageId="14" pageNumber="249">
Etymology. The specific epithet (noun) refers to its
<typeStatus id="336D8817FFEF737AFCB8FAC2FC3DFACC" box="[892,933,1358,1379]" pageId="14" pageNumber="249">type</typeStatus>
locality,
<collectingCountry id="94C17625FFEF737AFBC6FAC2FBD5FACC" box="[1026,1101,1358,1379]" name="Nigeria" pageId="14" pageNumber="249">Nigeria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,259 @@
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<mods:title id="3A8A309D52071AE6175F037DBD9556C8">Afrotropical Species Of Chaetopodella Duda (Diptera: Sphaeroceridae)</mods:title>
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<paragraph id="EC6936B5FFEF737AFDA5FA38FC4AFA61" blockId="14.[609,978,1460,1486]" box="[609,978,1460,1486]" pageId="14" pageNumber="249">
<heading id="B72181D9FFEF737AFDA5FA38FC4AFA61" box="[609,978,1460,1486]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="249" reason="2">
<taxonomicName id="2BD64D36FFEF737AFDA5FA38FC14FA61" authorityName="Papp" authorityYear="2008" box="[609,908,1460,1486]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="demeteri" status="sp. nov.">
<emphasis id="DEA2EAA7FFEF737AFDA5FA38FC14FA61" bold="true" box="[609,908,1460,1486]" pageId="14" pageNumber="249">Chaetopodella demeteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFEF737AFC57FA38FC4AFA61" box="[915,978,1460,1486]" pageId="14" pageNumber="249" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4CC653EFFEF737BFD0AFA5BFC87F8AE" lastPageId="15" lastPageNumber="250" pageId="14" pageNumber="249" type="description">
<paragraph id="EC6936B5FFEF737AFD0AFA5BFCFCFA5E" blockId="14.[718,868,1495,1521]" box="[718,868,1495,1521]" pageId="14" pageNumber="249">
<heading id="B72181D9FFEF737AFD0AFA5BFCFCFA5E" box="[718,868,1495,1521]" centered="true" fontSize="11" level="3" pageId="14" pageNumber="249" reason="3">
(
<figureCitation id="74ED2A30FFEF737AFD12FA5BFCC5FA5E" box="[726,861,1495,1521]" captionStart="Figs 4146" captionStartId="16.[317,360,1720,1741]" captionTargetBox="[317,1266,922,1691]" captionTargetId="figure-121@16.[317,1269,922,1691]" captionTargetPageId="16" captionText="Figs 4146. Chaetopodella (Ch.) demeteri sp. n., paratype male, genitalia. 41 = caudal process of sternite 5; 42 = subanal plate, caudal view; 4344 = surstylus: 43 = in the broadest (sublateral) view, 44 = in ventral view; 45= phallic complex (inner genitalia), lateral view, 46 = apical part of the dorsal" figureDoi="http://doi.org/10.5281/zenodo.12585064" httpUri="https://zenodo.org/record/12585064/files/figure.png" pageId="14" pageNumber="249">Figs 4146</figureCitation>
)
</heading>
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAF997FCB3F9E2" blockId="14.[317,1270,1563,1785]" pageId="14" pageNumber="249">
<materialsCitation id="5CBE3CE8FFEF737AFEBAF997FCB3F9E2" collectingDate="1978-08-14" collectionCode="HNHM" country="Nigeria" location="Yanguikari Reserve" municipality="Wikki" pageId="14" pageNumber="249" specimenCount="1" typeStatus="holotype">
<typeStatus id="336D8817FFEF737AFEBAF997FE4FF99F" box="[382,471,1563,1584]" pageId="14" pageNumber="249" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="8AC7AE70FFEF737AFDD2F997FDFEF99F" box="[534,614,1563,1584]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14" pageNumber="249">HNHM</collectionCode>
):
<collectingCountry id="94C17625FFEF737AFDB0F997FD26F99F" box="[628,702,1563,1584]" name="Nigeria" pageId="14" pageNumber="249">Nigeria</collectingCountry>
,
<location id="E909606EFFEF737AFD03F997FC10F99F" LSID="urn:lsid:plazi:treatment:647F87A3FFEF737BFDA5FA38FC87F8AE:E909606EFFEF737AFD03F997FC10F99F" box="[711,904,1563,1584]" country="Nigeria" municipality="Wikki" name="Yanguikari Reserve" pageId="14" pageNumber="249">Yanguikari Reserve</location>
,
<collectingMunicipality id="0C0DACCFFFEF737AFC54F997FC54F99F" box="[912,972,1563,1584]" pageId="14" pageNumber="249">Wikki</collectingMunicipality>
<date id="98681075FFEF737AFC25F997FBF2F99F" box="[993,1130,1563,1584]" pageId="14" pageNumber="249" value="1978-08-14">
<collectingDate id="882CE99DFFEF737AFC25F997FBF2F99F" box="[993,1130,1563,1584]" pageId="14" pageNumber="249" value="1978-08-14">Aug. 14, 1978</collectingDate>
</date>
, leg. A. DEMETER, No. 18 [hippo dung from the previous night].
</materialsCitation>
</paragraph>
<paragraph id="EC6936B5FFEF737AFEBAF9D8FC97F956" blockId="14.[317,1270,1563,1785]" pageId="14" pageNumber="249">
<materialsCitation id="5CBE3CE8FFEF737AFEBAF9D8FC70F9C6" box="[382,1000,1620,1641]" country="Nigeria" county="Wikki" location="Yangui" municipality="Reserve" pageId="14" pageNumber="249" specimenCount="1" typeStatus="paratype">
<typeStatus id="336D8817FFEF737AFEBAF9D8FE47F9C6" box="[382,479,1620,1641]" pageId="14" pageNumber="249" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="94C17625FFEF737AFE2CF9D8FDAAF9C6" box="[488,562,1620,1641]" name="Nigeria" pageId="14" pageNumber="249">Nigeria</collectingCountry>
,
<location id="E909606EFFEF737AFDFFF9D8FD1AF9C6" LSID="urn:lsid:plazi:treatment:647F87A3FFEF737BFDA5FA38FC87F8AE:E909606EFFEF737AFDFFF9D8FD1AF9C6" box="[571,642,1620,1641]" country="Nigeria" county="Wikki" municipality="Reserve" name="Yangui" pageId="14" pageNumber="249">Yangui</location>
[correctly: Yankari]
<collectingMunicipality id="0C0DACCFFFEF737AFC88F9D8FC04F9C6" box="[844,924,1620,1641]" pageId="14" pageNumber="249">Reserve</collectingMunicipality>
,
<collectingCounty id="05084E39FFEF737AFC60F9D8FC70F9C6" box="[932,1000,1620,1641]" pageId="14" pageNumber="249">Wikki</collectingCounty>
</materialsCitation>
:
<materialsCitation id="5CBE3CE8FFEF737AFC2FF9D8FB0DF9C6" box="[1003,1173,1620,1641]" country="Nigeria" county="Wikki" location="Yangui" municipality="Reserve" pageId="14" pageNumber="249" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FAD0FD3CFFEF737AFC2FF9D8FBB3F9C6" box="[1003,1067,1620,1641]" count="1" pageId="14" pageNumber="249" type="male">1 male</specimenCount>
: same data
</materialsCitation>
.
<materialsCitation id="5CBE3CE8FFEF737AFB5AF9D8FE2DF929" country="Nigeria" county="Wikki" location="Yangui" municipality="Reserve" pageId="14" pageNumber="249" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="FAD0FD3CFFEF737AFB5AF9D8FB78F9C6" box="[1182,1248,1620,1641]" count="1" pageId="14" pageNumber="249" type="male">1 male</specimenCount>
,
<specimenCount id="FAD0FD3CFFEF737AFB2EF9D8FE1AF929" count="1" pageId="14" pageNumber="249" type="female">1 female</specimenCount>
: ibid
</materialsCitation>
., Aug. 14, No. 16 [fresh buffalo dung];
<materialsCitation id="5CBE3CE8FFEF737AFCF0F9FDFC5DF929" box="[820,965,1649,1670]" country="Nigeria" county="Wikki" location="Yangui" municipality="Reserve" pageId="14" pageNumber="249" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="FAD0FD3CFFEF737AFCF0F9FDFC1FF929" box="[820,903,1649,1670]" count="1" pageId="14" pageNumber="249" type="female">1 female</specimenCount>
: ibid
</materialsCitation>
., No. 20 [fresh buffalo dung];
<materialsCitation id="5CBE3CE8FFEF737AFB2EF9FDFD9EF90C" country="Nigeria" county="Wikki" location="Yangui" municipality="Reserve" pageId="14" pageNumber="249" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="FAD0FD3CFFEF737AFB2EF9FDFEEDF90C" count="2" pageId="14" pageNumber="249" type="male">2 males</specimenCount>
<specimenCount id="FAD0FD3CFFEF737AFEBFF902FE4BF90C" box="[379,467,1678,1699]" count="1" pageId="14" pageNumber="249" type="female">1 female</specimenCount>
: ibid
</materialsCitation>
., Aug. 15, No. 22 [buffalo dung from the previous night].
<specimenCount id="FAD0FD3CFFEF737AFBF2F902FB0CF90C" box="[1078,1172,1678,1699]" count="1" pageId="14" pageNumber="249" type="male">One male</specimenCount>
of No. 22 was originally severely damaged by the pin; now wings are preserved between two small pieces of cover glass, abdomen and genitalia (dissected) in a plastic microvial with glycerol, and the rest of the body is also in another microvial with glycerol.
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAFE38FB4CFDAC" blockId="15.[317,1271,436,1437]" pageId="15" pageNumber="250">
Measurements in mm: body length 1.54 (
<typeStatus id="336D8817FFEE737BFCD7FE38FCF4FE66" box="[787,876,436,457]" pageId="15" pageNumber="250" type="holotype">holotype</typeStatus>
), 1.321.38 (
<typeStatus id="336D8817FFEE737BFC28FE38FBD8FE66" box="[1004,1088,436,457]" pageId="15" pageNumber="250" type="paratype">paratype</typeStatus>
males), 1.811.87 (
<typeStatus id="336D8817FFEE737BFE87FE5DFE39FE49" box="[323,417,465,486]" pageId="15" pageNumber="250" type="paratype">paratypes</typeStatus>
females), wing length 1.37 (
<typeStatus id="336D8817FFEE737BFD73FE5DFC88FE49" box="[695,784,465,486]" pageId="15" pageNumber="250" type="holotype">holotype</typeStatus>
), 1.151.25 (
<typeStatus id="336D8817FFEE737BFC49FE5DFC79FE49" box="[909,993,465,486]" pageId="15" pageNumber="250" type="paratype">paratype</typeStatus>
males), 1.451.59 (
<typeStatus id="336D8817FFEE737BFB65FE5DFB6DFE49" box="[1185,1269,465,486]" pageId="15" pageNumber="250" type="paratype">paratype</typeStatus>
females), wing width 0.47 (
<typeStatus id="336D8817FFEE737BFDFAFE62FD0FFDAC" box="[574,663,494,515]" pageId="15" pageNumber="250" type="holotype">holotype</typeStatus>
), 0.440.46 (
<typeStatus id="336D8817FFEE737BFCD4FE62FCFCFDAC" box="[784,868,494,515]" pageId="15" pageNumber="250" type="paratype">paratype</typeStatus>
males), 0.54 0.61 (
<typeStatus id="336D8817FFEE737BFBE5FE62FBEDFDAC" box="[1057,1141,494,515]" pageId="15" pageNumber="250" type="paratype">paratype</typeStatus>
females).
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAFD87FD5EFD3C" blockId="15.[317,1271,436,1437]" pageId="15" pageNumber="250">
Head: Silvery spots or stripes around bases of macrochaetae. Frons dark brown, at most anterior 1/5 reddish. Face dirty yellow, gena greyish yellow; 3
<emphasis id="DEA2EAA7FFEE737BFCA5FDA4FCE0FD92" box="[865,888,552,573]" italics="true" pageId="15" pageNumber="250">ifr</emphasis>
; upper setae slightly longer; anterior
<emphasis id="DEA2EAA7FFEE737BFB13FDA4FB6EFD92" box="[1239,1270,552,573]" italics="true" pageId="15" pageNumber="250">ors</emphasis>
thin
<quantity id="2B2E9B50FFEE737BFEA3FDC8FE25FDF6" box="[359,445,580,601]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="250" unit="mm" value="0.09">0.09 mm</quantity>
, posterior
<emphasis id="DEA2EAA7FFEE737BFDDAFDC9FDA5FDF5" box="[542,573,581,602]" italics="true" pageId="15" pageNumber="250">ors</emphasis>
thick
<quantity id="2B2E9B50FFEE737BFDB1FDC8FC92FDF6" box="[629,778,580,601]" metricMagnitude="-4" metricUnit="m" metricValue="1.525" metricValueMax="1.55" metricValueMin="1.5" pageId="15" pageNumber="250" unit="mm" value="0.1525" valueMax="0.155" valueMin="0.15">0.150.155 mm</quantity>
long. Eye short oval, its longest diameter (
<quantity id="2B2E9B50FFEE737BFB5DFDC8FB77FDF6" box="[1177,1263,580,601]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="15" pageNumber="250" unit="mm" value="0.27">0.27 mm</quantity>
) more than 3.0 times as long as gena below eye (
<quantity id="2B2E9B50FFEE737BFCC4FDEDFCC2FDD9" box="[768,858,609,630]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="250" unit="mm" value="0.08">0.08 mm</quantity>
). Antenna dark brown. Arista long, 4.04.4 times as long as antenna, shortly ciliate.
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAFD17FB0AFCEF" blockId="15.[317,1271,436,1437]" pageId="15" pageNumber="250">
Thorax: Mesonotal colour dominated by velvety black, i.e. silvery microtomentum only around setal bases and this silvery colour confluent only in 4 short stripes, which terminate at the level of posterior notopleural seta. Six rows of
<emphasis id="DEA2EAA7FFEE737BFCD1FD59FCB4FD45" box="[789,812,725,746]" italics="true" pageId="15" pageNumber="250">ac</emphasis>
microsetae in front of suture, 3
<emphasis id="DEA2EAA7FFEE737BFBB5FD59FB10FD45" box="[1137,1160,725,746]" italics="true" pageId="15" pageNumber="250">dc</emphasis>
shortening anteriorad. Two
<emphasis id="DEA2EAA7FFEE737BFE1FFD7DFD94FCA9" box="[475,524,753,774]" italics="true" pageId="15" pageNumber="250">kepst</emphasis>
, posterior one very long:
<quantity id="2B2E9B50FFEE737BFCC4FD7DFCF8FCA9" box="[768,864,753,774]" metricMagnitude="-4" metricUnit="m" metricValue="2.8499999999999996" pageId="15" pageNumber="250" unit="mm" value="0.285">0.285 mm</quantity>
on
<typeStatus id="336D8817FFEE737BFC47FD7DFC41FCA9" box="[899,985,753,774]" pageId="15" pageNumber="250" type="holotype">holotype</typeStatus>
,
<quantity id="2B2E9B50FFEE737BFC26FD7DFBAEFCA9" box="[994,1078,753,774]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="250" unit="mm" value="0.26">0.26 mm</quantity>
on a male
<typeStatus id="336D8817FFEE737BFB59FD7DFB6BFCA9" box="[1181,1267,753,774]" pageId="15" pageNumber="250" type="paratype">paratype</typeStatus>
. Scutellum velvety black, wider than long,
<quantity id="2B2E9B50FFEE737BFD1EFC82FCA4FC8C" box="[730,828,782,803]" metricMagnitude="-4" metricUnit="m" metricValue="2.8499999999999996" pageId="15" pageNumber="250" unit="mm" value="0.285">0.285 mm</quantity>
broad and
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long; 2 pairs of scutellar setae rather long, apical one
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, lateral one
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, scutellum (
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male).
</paragraph>
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Wing: Milky, veins white; C not extended beyond apex of R
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, first section with much longer setae than those of second and third sections; R
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distinctly bent up to C; C-index = 0.40/
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= 1.6 (1.55 on another male), R-M dM-Cu: dM-Cu = 3.27; lower corner of the discal cell angulate with a short vein appendage, or, in some specimens rounded without vein appendage; alula broad and rounded; halter with reddish yellow knob and yellow stem.
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAFC54FE2BFBCF" blockId="15.[317,1271,436,1437]" pageId="15" pageNumber="250">
Legs: Yellowish brown, fore coxa ochreous; male tarsi darker brown. Fore femur with 2 dorsal setae on apical half. Male fore tibia not thickened, fore tarsus slightly thickened and tarsomeres 25 even flattened. Fore coxa with 56 distinct setae and without long light (whitish) hairs. Mid trochanter with a thick
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long seta laterally. Mid femur with a ventral row of 3 setae on basal 1/
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male.
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAFBE4FDCAFB7C" blockId="15.[317,1271,436,1437]" pageId="15" pageNumber="250">
Abdomen: Male sternite 5 rather similar to that of
<taxonomicName id="2BD64D36FFEE737BFCADFBE4FC6FFBD2" authorityName="Papp" authorityYear="2008" box="[873,1015,1128,1149]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopica">
<emphasis id="DEA2EAA7FFEE737BFCADFBE4FC6FFBD2" box="[873,1015,1128,1149]" italics="true" pageId="15" pageNumber="250">Ch. aethiopica</emphasis>
</taxonomicName>
. However, caudal process (
<figureCitation id="74ED2A30FFEE737BFE80FB08FE16FB36" box="[324,398,1156,1177]" captionStart="Figs 4146" captionStartId="16.[317,360,1720,1741]" captionTargetBox="[317,1266,922,1691]" captionTargetId="figure-121@16.[317,1269,922,1691]" captionTargetPageId="16" captionText="Figs 4146. Chaetopodella (Ch.) demeteri sp. n., paratype male, genitalia. 41 = caudal process of sternite 5; 42 = subanal plate, caudal view; 4344 = surstylus: 43 = in the broadest (sublateral) view, 44 = in ventral view; 45= phallic complex (inner genitalia), lateral view, 46 = apical part of the dorsal" figureDoi="http://doi.org/10.5281/zenodo.12585064" httpUri="https://zenodo.org/record/12585064/files/figure.png" pageId="15" pageNumber="250">Fig. 41</figureCitation>
) short curved with shorter thorns. Medial platelet with short pilosity. Ventral plate of subepandrial sclerite (
<figureCitation id="74ED2A30FFEE737BFDCAFB2DFDCBFB19" box="[526,595,1185,1206]" captionStart="Figs 4146" captionStartId="16.[317,360,1720,1741]" captionTargetBox="[317,1266,922,1691]" captionTargetId="figure-121@16.[317,1269,922,1691]" captionTargetPageId="16" captionText="Figs 4146. Chaetopodella (Ch.) demeteri sp. n., paratype male, genitalia. 41 = caudal process of sternite 5; 42 = subanal plate, caudal view; 4344 = surstylus: 43 = in the broadest (sublateral) view, 44 = in ventral view; 45= phallic complex (inner genitalia), lateral view, 46 = apical part of the dorsal" figureDoi="http://doi.org/10.5281/zenodo.12585064" httpUri="https://zenodo.org/record/12585064/files/figure.png" pageId="15" pageNumber="250">Fig. 42</figureCitation>
) with 2 pairs of long laterally curved thorn-like seta, the lateral one not thicker than the medial pair.
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAFB57FC39FACC" blockId="15.[317,1271,436,1437]" pageId="15" pageNumber="250">
Surstylus (
<figureCitation id="74ED2A30FFEE737BFE2CFB57FDC3FB5F" box="[488,603,1243,1264]" captionStart="Figs 4146" captionStartId="16.[317,360,1720,1741]" captionTargetBox="[317,1266,922,1691]" captionTargetId="figure-121@16.[317,1269,922,1691]" captionTargetPageId="16" captionText="Figs 4146. Chaetopodella (Ch.) demeteri sp. n., paratype male, genitalia. 41 = caudal process of sternite 5; 42 = subanal plate, caudal view; 4344 = surstylus: 43 = in the broadest (sublateral) view, 44 = in ventral view; 45= phallic complex (inner genitalia), lateral view, 46 = apical part of the dorsal" figureDoi="http://doi.org/10.5281/zenodo.12585064" httpUri="https://zenodo.org/record/12585064/files/figure.png" pageId="15" pageNumber="250">Figs 4344</figureCitation>
) laterally on outer surface with several setae incl. on inner edge. Epiphallus (
<figureCitation id="74ED2A30FFEE737BFE70FB74FE65FAA2" box="[436,509,1272,1293]" captionStart="Figs 4146" captionStartId="16.[317,360,1720,1741]" captionTargetBox="[317,1266,922,1691]" captionTargetId="figure-121@16.[317,1269,922,1691]" captionTargetPageId="16" captionText="Figs 4146. Chaetopodella (Ch.) demeteri sp. n., paratype male, genitalia. 41 = caudal process of sternite 5; 42 = subanal plate, caudal view; 4344 = surstylus: 43 = in the broadest (sublateral) view, 44 = in ventral view; 45= phallic complex (inner genitalia), lateral view, 46 = apical part of the dorsal" figureDoi="http://doi.org/10.5281/zenodo.12585064" httpUri="https://zenodo.org/record/12585064/files/figure.png" pageId="15" pageNumber="250">Fig. 45</figureCitation>
) smallest in the species group, subtriangular in profile, postgonite (
<figureCitation id="74ED2A30FFEE737BFB61FB74FB77FAA2" box="[1189,1263,1272,1293]" captionStart="Figs 4146" captionStartId="16.[317,360,1720,1741]" captionTargetBox="[317,1266,922,1691]" captionTargetId="figure-121@16.[317,1269,922,1691]" captionTargetPageId="16" captionText="Figs 4146. Chaetopodella (Ch.) demeteri sp. n., paratype male, genitalia. 41 = caudal process of sternite 5; 42 = subanal plate, caudal view; 4344 = surstylus: 43 = in the broadest (sublateral) view, 44 = in ventral view; 45= phallic complex (inner genitalia), lateral view, 46 = apical part of the dorsal" figureDoi="http://doi.org/10.5281/zenodo.12585064" httpUri="https://zenodo.org/record/12585064/files/figure.png" pageId="15" pageNumber="250">Fig. 45</figureCitation>
) strongly narrowed subapically, apex almost sharp. Phallapodeme somewhat smaller than in the related species. Apical part of the paired dorsal process of distiphallus with short and thick fork (
<figureCitation id="74ED2A30FFEE737BFB0FFABDFEE5FACC" captionStart="Figs 4146" captionStartId="16.[317,360,1720,1741]" captionTargetBox="[317,1266,922,1691]" captionTargetId="figure-121@16.[317,1269,922,1691]" captionTargetPageId="16" captionText="Figs 4146. Chaetopodella (Ch.) demeteri sp. n., paratype male, genitalia. 41 = caudal process of sternite 5; 42 = subanal plate, caudal view; 4344 = surstylus: 43 = in the broadest (sublateral) view, 44 = in ventral view; 45= phallic complex (inner genitalia), lateral view, 46 = apical part of the dorsal" figureDoi="http://doi.org/10.5281/zenodo.12585064" httpUri="https://zenodo.org/record/12585064/files/figure.png" pageId="15" pageNumber="250">Figs 4546</figureCitation>
), much shorter than those of
<taxonomicName id="2BD64D36FFEE737BFD58FAC2FC8EFACC" authorityName="Papp" authorityYear="2008" box="[668,790,1358,1379]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="250" phylum="Arthropoda" rank="species" species="nigeriae">
<emphasis id="DEA2EAA7FFEE737BFD58FAC2FC8EFACC" box="[668,790,1358,1379]" italics="true" pageId="15" pageNumber="250">Ch. nigeriae</emphasis>
</taxonomicName>
(
<figureCitation id="74ED2A30FFEE737BFCE0FAC2FC0EFACC" box="[804,918,1358,1379]" captionStart="Figs 3440" captionStartId="13.[317,360,1702,1723]" captionTargetBox="[327,1265,480,1652]" captionTargetId="figure-10@13.[317,1269,474,1652]" captionTargetPageId="13" captionText="Figs 3440. Chaetopodella (Ch.) nigeriae sp. n., paratype male, genitalia. 34 = pregenital sclerites, subventral-subcaudal view, 35 = 5th sternite, ventral view, 36 = subanal plate, caudal view; 3738 = surstylus: 36 = in the broadest (sublateral) view, 38 = in ventral view; 39 = phallic complex (inner genitalia), lateral view, 40 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585062" httpUri="https://zenodo.org/record/12585062/files/figure.png" pageId="15" pageNumber="250">Figs 3940</figureCitation>
).
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAFAE7FB18FA32" blockId="15.[317,1271,436,1437]" pageId="15" pageNumber="250">
Female abdominal terga velvety black, sternites 35 (6) pale (yellowish), lateral seta on tergite 6
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long. Cerci brown, ca.
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long, longest cercal setae
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.
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAFA4DFE32F9BF" blockId="15.[317,1270,1473,1552]" pageId="15" pageNumber="250">
Etymology. The species was named to the honour of Dr.
<smallCapsWord id="EA8FA069FFEE737BFC58FA4DFC74FA7A" baselines="1489,1489" box="[924,1004,1473,1494]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Andras" pageId="15" pageNumber="250">ANDRÁS</smallCapsWord>
<smallCapsWord id="EA8FA069FFEE737BFC35FA4DFBD6FA7A" baselines="1489,1489" box="[1009,1102,1473,1494]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Demeter" pageId="15" pageNumber="250">DEMETER</smallCapsWord>
(formerly the curator of the collection of mammals in the HNHM), who collected the
<typeStatus id="336D8817FFEE737BFC0CFA52FC69FA5C" box="[968,1009,1502,1523]" pageId="15" pageNumber="250">type</typeStatus>
series of the three new species above.
</paragraph>
<paragraph id="EC6936B5FFEE737BFEBAF9BBFC87F8AE" blockId="15.[317,1270,1591,1793]" pageId="15" pageNumber="250">
Specimens with rounded discal cell. There are specimens in all the three
<taxonomicName id="2BD64D36FFEE737BFEF9F9D7FE72F9DA" box="[317,490,1627,1653]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="250" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="DEA2EAA7FFEE737BFEF9F9D7FE72F9DA" box="[317,490,1627,1653]" italics="true" pageId="15" pageNumber="250">Chaetopodella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C59157DCFFEE737BFE35F9D7FDB0F9DA" box="[497,552,1627,1653]" pageId="15" pageNumber="250" sensu="stricto">s.str.</taxonomicNameLabel>
species described here with rounded discal cell. I made some figures on such a male of
<taxonomicName id="2BD64D36FFEE737BFDA3F9F2FC8AF937" authorityName="Papp" authorityYear="2008" box="[615,786,1662,1688]" class="Insecta" family="Sphaeroceridae" genus="Chaetopodella" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="250" phylum="Arthropoda" rank="species" species="aethiopica">
<emphasis id="DEA2EAA7FFEE737BFDA3F9F2FC8AF937" box="[615,786,1662,1688]" italics="true" pageId="15" pageNumber="250">Ch. aethiopica</emphasis>
</taxonomicName>
(
<figureCitation id="74ED2A30FFEE737BFCE5F9F2FC3EF937" box="[801,934,1662,1688]" captionStart="Figs 4751" captionStartId="17.[317,360,1722,1743]" captionTargetBox="[326,1265,831,1689]" captionTargetId="figure-120@17.[320,1265,826,1693]" captionTargetPageId="17" captionText="Figs 4751. Chaetopodella (Ch.) aethiopica sp. n., genitalia of a paratype male with rounded discal cell. 47 = pregenital sclerites, subventral-subcaudal view, 48 = subanal plate, caudal view; 49 = postgonite and epiphallus, lateral view, 50 = surstylus, broadest (sublateral) view, 51 = apical part of" figureDoi="http://doi.org/10.5281/zenodo.12585068" httpUri="https://zenodo.org/record/12585068/files/figure.png" pageId="15" pageNumber="250">Figs 4751</figureCitation>
) in order to demonstrate that these are not different species. The differences one can detect if compared to
<figureCitation id="74ED2A30FFEE737BFB07F92DFE1EF971" captionStart="Figs 2733" captionStartId="11.[317,360,1702,1723]" captionTargetBox="[351,1232,427,1670]" captionTargetId="figure-10@11.[351,1235,422,1673]" captionTargetPageId="11" captionText="Figs 2733. Chaetopodella (Ch.) aethiopica sp.n., paratype male, genitalia.27 = pregenital sclerites, subventral-subcaudal view, 28 = 5th sternite, ventral view, 29 = subanal plate, caudal view; 3031 = surstylus: 30 = in the broadest (sublateral) view, 31 = in ventral view; 32 = phallic complex (inner genitalia), lateral view, 33 = apical part of the dorsal process of distiphallus, broadest view. Scales:" figureDoi="http://doi.org/10.5281/zenodo.12585060" httpUri="https://zenodo.org/record/12585060/files/figure.png" pageId="15" pageNumber="250">Figs 2733</figureCitation>
are individual variations, or, slight differences caused by the slightly different positioning of the given genital parts.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,104 +1,128 @@
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<treatment id="648ECEFA97164D471F57062AB5820F2B" ID-GBIF-Taxon="127858146" LSID="urn:lsid:plazi:treatment:648ECEFA97164D471F57062AB5820F2B" httpUri="http://treatment.plazi.org/id/648ECEFA97164D471F57062AB5820F2B" lastPageId="41" lastPageNumber="120" pageId="40" pageNumber="119">
<subSubSection id="EDCF44C83BEEB350B1C9E80A65E64C48" pageId="40" pageNumber="119" type="nomenclature">
<paragraph id="FFD3269E2E5FE154433871D60D17B7FD" pageId="40" pageNumber="119">
<taxonomicName id="8BC23BADB8158E01EBC045837DFB59ED" authority="Rayment" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amegilla (Zonamegilla) murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis" subGenus="Zonamegilla">Amegilla (Zonamegilla) murrayensis (Rayment)</taxonomicName>
Figs 11, 32, 45
<taxonomicName id="8BC23BADB8158E01EBC045837DFB59ED" ID-CoL="66B6N" LSID="BC2E1325-67B8-5FA0-A9CE-8CF6A4D2028A" authority="(Rayment)" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla (Zonamegilla) murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis" subGenus="Zonamegilla">Amegilla (Zonamegilla) murrayensis (Rayment)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="068C691E5993EA8844215594C279FCAA" pageId="40" pageNumber="119" type="description">
<paragraph id="0CCD27AAEB05287E3D64149D10998D8C" pageId="40" pageNumber="119">
<figureCitation id="018A1882823D9789F72054E4D1593FB3" captionStart="Figure 314" captionStartId="F3" captionText="Figure 3 - 14. Distribution maps ● indicate museum record, ʘ indicates DNA specimen." figureDoi="10.3897/zookeys.653.11177.figure3-14" httpUri="https://binary.pensoft.net/fig/121856" pageId="40" pageNumber="119">Figs 11</figureCitation>
<figureCitation id="36A72D5D48B3C492816FF181D8E7AD65" captionStart="Figure 2437" captionStartId="F9" captionText="Figure 24 - 37. Apical tergites of females, especially T 5. 24 Amegilla aeruginosa 25 Amegilla chlorocyanea 26 Amegilla adelaidae 27 Amegilla alpha 28 Amegilla asserta 29 Amegilla cingulata 30 Amegilla indistincta 31 Amegilla karlba 32 Amegilla murrayensis 33 Amegilla paeninsulae 34 Amegilla pulchra 35 Amegilla thorogoodi 36 Amegilla viridicingulata 37 Amegilla walkeri." figureDoi="10.3897/zookeys.653.11177.figure24-37" httpUri="https://binary.pensoft.net/fig/121864" pageId="40" pageNumber="119">, 32</figureCitation>
<figureCitation id="1ABBFBEFB70809F3555567E61E4ADB56" captionStart="Figures 3850" captionStartId="F10" captionText="Figures 38 - 50. S 7 of males. 38 Amegilla aeruginosa 39 Amegilla chlorocyanea 40 Amegilla adelaidae 41 Amegilla asserta 42 Amegilla cingulata 43 Amegilla indistincta 44 Amegilla karlba 45 Amegilla murrayensis 46 Amegilla paeninsulae 47 Amegilla pulchra 48 Amegilla thorogoodi 49 Amegilla viridicingulata 50 Amegilla walkeri." figureDoi="10.3897/zookeys.653.11177.figures38-50" httpUri="https://binary.pensoft.net/fig/121865" pageId="40" pageNumber="119">, 45</figureCitation>
</paragraph>
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<subSubSection id="C723E9CDDDDE76B66AB07461E5EA7553" pageId="40" pageNumber="119" type="reference_group">
<paragraph id="14800BC2AE141815A8A3ADF367ACBF72" pageId="40" pageNumber="119">
<taxonomicName id="117C2EF4ACFCC9D064D430CD1490DD53" class="Insecta" family="Apidae" genus="Anthophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anthophora murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">Anthophora murrayensis</taxonomicName>
<taxonomicName id="117C2EF4ACFCC9D064D430CD1490DD53" authorityName="Rayment" authorityYear="1935" class="Insecta" family="Apidae" genus="Anthophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthophora murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">Anthophora murrayensis</taxonomicName>
Rayment, 1939, p. 288.
</paragraph>
<paragraph id="9FCD67E7607810C2B4EC6CDCDCFD67EC" pageId="40" pageNumber="119">
<taxonomicName id="CC644F8E3B118966D790155D3427D4E5" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">Amegilla murrayensis</taxonomicName>
<taxonomicName id="CC644F8E3B118966D790155D3427D4E5" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">Amegilla murrayensis</taxonomicName>
(Rayment) Michener, 1965, p. 217.
</paragraph>
<paragraph id="55D8BECC41A00F8974A8BBF5BB134E7D" pageId="40" pageNumber="119">
<taxonomicName id="12849424315AE99F8013D8388552D933" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amegilla (Zonamegilla) murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis" subGenus="Zonamegilla">Amegilla (Zonamegilla) murrayensis</taxonomicName>
<taxonomicName id="12849424315AE99F8013D8388552D933" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla (Zonamegilla) murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis" subGenus="Zonamegilla">Amegilla (Zonamegilla) murrayensis</taxonomicName>
(Rayment) Brooks, 1988, p. 511.
</paragraph>
<paragraph id="C6D18F6C9652C6C2C2A23A1F5B7AA991" pageId="40" pageNumber="119">
<taxonomicName id="7E7FC7AA5C6B7BEC18FAED51FF0FDD37" class="Insecta" family="Apidae" genus="Anthophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anthophora longula" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="longula">Anthophora longula</taxonomicName>
Rayment, 1947, p. 59. n. syn.
<taxonomicName id="7E7FC7AA5C6B7BEC18FAED51FF0FDD37" authorityName="Rayment" authorityYear="1947" class="Insecta" family="Apidae" genus="Anthophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthophora longula" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="longula">Anthophora longula</taxonomicName>
Rayment, 1947, p. 59.
<emphasis id="6F5DFBE098DC6E51C0E69DED5DFA325C" bold="true" pageId="40" pageNumber="119">n. syn.</emphasis>
</paragraph>
<paragraph id="03CE365E49B5BA2E65B96CFC1071D7D3" pageId="40" pageNumber="119">
<taxonomicName id="2E034306C158DFDA3784F3A0718EF795" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amegilla longula" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="longula">Amegilla longula</taxonomicName>
<taxonomicName id="2E034306C158DFDA3784F3A0718EF795" baseAuthorityName="Rayment" baseAuthorityYear="1947" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla longula" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="longula">Amegilla longula</taxonomicName>
(Rayment) Michener, 1965, p. 217.
</paragraph>
<paragraph id="5CCB237319C35195BB497F2481B6A788" pageId="40" pageNumber="119">
<taxonomicName id="F01A1CD9F2F99D4F3FDCD8C734B4D8B1" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amegilla (Zonamegilla) longula" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="longula" subGenus="Zonamegilla">Amegilla (Zonamegilla) longula</taxonomicName>
<taxonomicName id="F01A1CD9F2F99D4F3FDCD8C734B4D8B1" baseAuthorityName="Rayment" baseAuthorityYear="1947" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla (Zonamegilla) longula" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="longula" subGenus="Zonamegilla">Amegilla (Zonamegilla) longula</taxonomicName>
(Rayment) Brooks, 1988, p. 511.
</paragraph>
<paragraph id="EE5D44901B16089917CCB4F286376BBB" pageId="40" pageNumber="119">
<taxonomicName id="63EED546D8309467C6591B91643CB7CB" class="Insecta" family="Apidae" genus="Anthophora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Anthophora subsalteri" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="subsalteri">Anthophora subsalteri</taxonomicName>
Rayment, 1947, p. 69. n. syn.
<taxonomicName id="63EED546D8309467C6591B91643CB7CB" authorityName="Rayment" authorityYear="1947" class="Insecta" family="Apidae" genus="Anthophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthophora subsalteri" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="subsalteri">Anthophora subsalteri</taxonomicName>
Rayment, 1947, p. 69.
<emphasis id="B07507CA35D68A92741B67B0BA3853D8" bold="true" pageId="40" pageNumber="119">n. syn.</emphasis>
</paragraph>
<paragraph id="8001986883C5F8AEAF65A001C34955ED" pageId="40" pageNumber="119">
<taxonomicName id="AECF443862FC2B6BDBD520974552F55C" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amegilla subsalteri" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="subsalteri">Amegilla subsalteri</taxonomicName>
<taxonomicName id="AECF443862FC2B6BDBD520974552F55C" baseAuthorityName="Rayment" baseAuthorityYear="1947" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla subsalteri" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="subsalteri">Amegilla subsalteri</taxonomicName>
(Rayment) Michener, 1965, p. 217.
</paragraph>
<paragraph id="FB1D38B13CB50E67BA3F2CD175D7E5D0" pageId="40" pageNumber="119">
<taxonomicName id="EE3C23BA3CE8CBA74FF396B5BEA3E336" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amegilla (Zonamegilla) subsalteri" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="subsalteri" subGenus="Zonamegilla">Amegilla (Zonamegilla) subsalteri</taxonomicName>
<taxonomicName id="EE3C23BA3CE8CBA74FF396B5BEA3E336" baseAuthorityName="Rayment" baseAuthorityYear="1947" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla (Zonamegilla) subsalteri" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="subsalteri" subGenus="Zonamegilla">Amegilla (Zonamegilla) subsalteri</taxonomicName>
(Rayment) Brooks, 1988, p. 511.
</paragraph>
</subSubSection>
<subSubSection id="40A63B4490799E064AB5FF5090287375" pageId="40" pageNumber="119" type="materials_examined">
<paragraph id="CC71D981385B6098581606487B8E32DB" pageId="40" pageNumber="119">Material examined.</paragraph>
<paragraph id="883DDA4F3823C28F008172FD2F4D63C2" pageId="40" pageNumber="119">229 females and 132males.</paragraph>
<paragraph id="883DDA4F3823C28F008172FD2F4D63C2" pageId="40" pageNumber="119">
<specimenCount id="5C691EDA95DA55D1A8A07FCDFC56E1BC" count="229" type="female">229 females</specimenCount>
and
<specimenCount id="EE5A200FA970947CBC08EF20B587FE3D" count="132" type="male">132males</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="A7EDDDA759658B28E13C81805244DAD3" pageId="40" pageNumber="119" type="type data">
<paragraph id="3C2941AD839AC7A67A785E17C6656DD2" pageId="40" pageNumber="119">Type data.</paragraph>
<paragraph id="CA5792C26FF2F7C1C74C56E2221D3038" pageId="40" pageNumber="119">
Syntypes of
<taxonomicName id="29D765EA41A175A80D67EE46A8C249DF" lsidName="murrayensis" pageId="40" pageNumber="119" rank="species" species="murrayensis">murrayensis</taxonomicName>
<taxonomicName id="D5CD9A96D74AEDC356D3147D2F4FF298" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">
<emphasis id="64BE5B93E6B7CD054967EB69DA36C1B4" italics="true" pageId="40" pageNumber="119">Amegilla murrayensis</emphasis>
</taxonomicName>
, male, female, Gunbower, VIC, 16 Mar. 1940, No. G500,
<normalizedToken id="735F5EE25BCF8C54C019B7A2BACC8A79" originalValue="“Type”">&quot;Type&quot;</normalizedToken>
&amp;
@ -107,51 +131,75 @@ Syntypes of
</paragraph>
<paragraph id="4FD4406F6CD8C11E5D2BC44257A96EE3" pageId="40" pageNumber="119">
Syntypes of
<taxonomicName id="D2AA9F3D1BD835A2242D0295B7F56D13" lsidName="longula" pageId="40" pageNumber="119" rank="species" species="longula">longula</taxonomicName>
<taxonomicName id="F7643432CDFFD7634A8ABF6FA590ACAF" baseAuthorityName="Rayment" baseAuthorityYear="1947" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla longula" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="longula">
<emphasis id="0AB6185683376C09E25E75C1C23012CC" italics="true" pageId="40" pageNumber="119">Amegilla longula</emphasis>
</taxonomicName>
, male, female, Orroroo, SA, 3 &amp; 10 Feb. 1940, ANIC 32-034558-9.
</paragraph>
</subSubSection>
<subSubSection id="5E6D7AC5A58A9F13AF94F0D6A10B3C92" pageId="40" pageNumber="119" type="decisions for synonymy">
<paragraph id="64E236E348A1EDB6C9981BD20151275B" pageId="40" pageNumber="119">Decisions for synonymy.</paragraph>
<paragraph id="97E5F2C645C7DD54AAA9FED638464688" pageId="40" pageNumber="119">The results of DNA analyses of specimens from across the complete geographical range showed no geographical pattern with respect to sequence variation. The uncorrected sequence divergence was found to be 0-1.3% (Table 1), which is well below the usual limits for conspecific individuals.</paragraph>
<paragraph id="97E5F2C645C7DD54AAA9FED638464688" pageId="40" pageNumber="119">
The results of DNA analyses of specimens from across the complete geographical range showed no geographical pattern with respect to sequence variation. The uncorrected sequence divergence was found to be 0-1.3% (Table
<tableCitation id="2FD0873C714A7C0C52161E1328E00D9B" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Maximum percentage uncorrected pairwise intraspecific sequence divergence of Australian Amegilla and distances to nearest other species clade A) based on 822 bp fragment of CO 1 using primers M 202 / M 70, B) based on 648 bp amplified using primers M 414 / M 423. ' - ': no data available. ' * ': inflated pairwise divergence due two multiple clades possible resulting from nuclear paralogues. (See also supplementary Figs S 1 and S 2)" httpUri="http://table.plazi.org/id/043B31A44079B965106F15E4A1CB9FE7" pageId="40" pageNumber="119" tableUuid="043B31A44079B965106F15E4A1CB9FE7">1</tableCitation>
), which is well below the usual limits for conspecific individuals.
</paragraph>
<paragraph id="BC9A386583456BCD50551BBB930650D6" pageId="40" pageNumber="119">
The syntypes of
<taxonomicName id="0429C9BF8D73739D639C44DDF106B405" lsidName="murrayensis" pageId="40" pageNumber="119" rank="species" species="murrayensis">murrayensis</taxonomicName>
<taxonomicName id="7E4B3E8615F599DC2028AB0BBE5B278F" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">
<emphasis id="E4C2166F80B5E9CA4E814B1312D135A5" italics="true" pageId="40" pageNumber="119">Amegilla murrayensis</emphasis>
</taxonomicName>
and
<taxonomicName id="E242E52361EEDBB76D1939F42BDA3EE1" lsidName="longula" pageId="40" pageNumber="119" rank="species" species="longula">longula</taxonomicName>
<taxonomicName id="37A0ED8116E19651C738269B4051C689" baseAuthorityName="Rayment" baseAuthorityYear="1947" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla longula" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="longula">
<emphasis id="7EAEB3B7F8FEE0217214975493F2A4FC" italics="true" pageId="40" pageNumber="119">Amegilla longula</emphasis>
</taxonomicName>
were examined and considered to be conspecific. Type material for
<taxonomicName id="C62F486FFEBBD8B5D08E383E036961B5" lsidName="subsalteri" pageId="40" pageNumber="119" rank="species" species="subsalteri">subsalteri</taxonomicName>
<taxonomicName id="CA06BB64106724B0F89B36CC9C1513C2" baseAuthorityName="Rayment" baseAuthorityYear="1947" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla subsalteri" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="subsalteri">
<emphasis id="CEC94CBEAD14654412356DAE31176D48" italics="true" pageId="40" pageNumber="119">Amegilla subsalteri</emphasis>
</taxonomicName>
, presumed to be the holotype by monotypy, was not found, but from
<normalizedToken id="DF503F82CD1434CB777D5A64E3AFCD92" originalValue="Rayments">Rayment's</normalizedToken>
description and drawings (
<bibRefCitation id="80F8A54B142EAEAE2D7B3656D6590CD4" pageId="40" pageNumber="119">Rayment 1947</bibRefCitation>
<bibRefCitation id="80F8A54B142EAEAE2D7B3656D6590CD4" author="Rayment, T" pageId="57" pageNumber="136" refId="B48" refString="Rayment, T, 1947. A critical revision of species in the zonata group of Anthophora by new characters (Part II), Treubia 19: 46-73." title="A critical revision of species in the zonata group of Anthophora by new characters (Part II), Treubia 19: 46 - 73." year="1947">Rayment 1947</bibRefCitation>
) together with the collection locality,
<taxonomicName id="66F9A25438D7F8C73418120939E71226" lsidName="subsalteri" pageId="40" pageNumber="119" rank="species" species="subsalteri">subsalteri</taxonomicName>
<taxonomicName id="76866336A60262B5B2145C06184CAAF3" baseAuthorityName="Rayment" baseAuthorityYear="1947" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla subsalteri" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="subsalteri">
<emphasis id="D06EAD59FE27C41C9ACBBF38A836D5F0" italics="true" pageId="40" pageNumber="119">Amegilla subsalteri</emphasis>
</taxonomicName>
is believed to be conspecific with
<taxonomicName id="2BC7AE6CBE3CB00C23F44EF4C60FE225" lsidName="murrayensis" pageId="40" pageNumber="119" rank="species" species="murrayensis">murrayensis</taxonomicName>
<taxonomicName id="49277E97AFDBA250DB867A5CAEEE7506" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">
<emphasis id="482CD3F6B46875396E7418C93B97AE65" italics="true" pageId="40" pageNumber="119">Amegilla murrayensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CA0001C6BE8CE48C2FE7C8CC75C37F99" pageId="40" pageNumber="119" type="diagnosis">
<paragraph id="ECBBD82193C31CEC9FE5FCB8DE51EDB2" pageId="40" pageNumber="119">Diagnosis.</paragraph>
<paragraph id="FA34048D3E1A8DF8CFB5499B03729743" pageId="40" pageNumber="119">
<taxonomicName id="82D5F251187FCA83F5DAC487D478260B" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">Amegilla murrayensis</taxonomicName>
<taxonomicName id="82D5F251187FCA83F5DAC487D478260B" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="40" pageNumber="119" phylum="Arthropoda" rank="species" species="murrayensis">
<emphasis id="D23F8CFFFCA77C26BE56A0A598DC9A15" italics="true" pageId="40" pageNumber="119">Amegilla murrayensis</emphasis>
</taxonomicName>
is a relatively small species with pale yellow face marks and narrow (about 0.3x the width of the disc on T2) apical hair bands which are usually pale blue, occasionally with an orange tint, but never bright orange; female hind tibia with a dark streak at least 0.5
<normalizedToken id="5415BFB8EA5FB49211D94C7BFD4DC3E1" originalValue="×">x</normalizedToken>
as long as the tibia; T5 with a patch of scattered white hair that narrows laterally and with a longitudinal line of denser white hair that does not intrude significantly into the prepygidial fimbria. Both sexes can be distinguished from other species by hair bands on T1-4 that appear broader laterally below the gradulus because of numerous scattered pale hairs on the disc (Figs 16, 17).
as long as the tibia; T5 with a patch of scattered white hair that narrows laterally and with a longitudinal line of denser white hair that does not intrude significantly into the prepygidial fimbria. Both sexes can be distinguished from other species by hair bands on T1-4 that appear broader laterally below the gradulus because of numerous scattered pale hairs on the disc (Figs
<figureCitation id="5B3CA0F641AE5029955F8DC8D6AAF442" captionStart="Figures 1619" captionStartId="F5" captionText="Figures 16 - 19. Abdomen, lateral view. 16 A. murrayensis female 17 male 18 A. pulchra female 19 male." figureDoi="10.3897/zookeys.653.11177.figures16-19" httpUri="https://binary.pensoft.net/fig/121858" pageId="40" pageNumber="119">16</figureCitation>
,
<figureCitation id="505F491A8421E4D5C9DADEFCADAABDCE" captionStart="Figures 1619" captionStartId="F5" captionText="Figures 16 - 19. Abdomen, lateral view. 16 A. murrayensis female 17 male 18 A. pulchra female 19 male." figureDoi="10.3897/zookeys.653.11177.figures16-19" httpUri="https://binary.pensoft.net/fig/121858" pageId="40" pageNumber="119">17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0CB88918DEC67DD6AF7AC5387567CAC0" pageId="40" pageNumber="119" type="description">
<paragraph id="A2A89E5310140582135A34E36CEDEE1F" pageId="40" pageNumber="119">Description.</paragraph>
<paragraph id="C5A8DB07C28FB1DEC5D0AC967AB0A8FE" pageId="40" pageNumber="119">
Female: Sunnyside, N of Murray Bridge,
<geoCoordinate id="2E1DE6E7AC6501311622D4B6EC047B98" direction="south" orientation="latitude" precision="5" value="-35.05">35.0500S</geoCoordinate>
<geoCoordinate id="298FF46CBC2E12FF40EB4C7B794F6886" direction="east" orientation="longitude" precision="5" value="139.36">139.3600E</geoCoordinate>
<emphasis id="C0D3ACD8F3E1789FA7264A1DA732A0DE" italics="true" pageId="40" pageNumber="119">Female</emphasis>
: Sunnyside, N of Murray Bridge,
<geoCoordinate id="2E1DE6E7AC6501311622D4B6EC047B98" degrees="35.0500" direction="south" orientation="latitude" precision="5" value="-35.05">35.0500S</geoCoordinate>
<geoCoordinate id="298FF46CBC2E12FF40EB4C7B794F6886" degrees="139.3600" direction="east" orientation="longitude" precision="5" value="139.36">139.3600E</geoCoordinate>
, 27 Feb 2003, R.Leijs &amp; K. Hogendoorn, SAMA 32-002635.
</paragraph>
<paragraph id="63EB62EE1E43CB71F1576CC2A9CED63E" pageId="40" pageNumber="119">Length 12 mm; forewing length 8 mm.</paragraph>
<paragraph id="32B2FCEA334787DBCC511A21607663EF" pageId="40" pageNumber="119">
Structure. Head: clypeus protuberant, in profile 0.5 width of eye; galea in repose reaching half-way between fore and mid coxae; length of f1 2.8
<emphasis id="9D51D1A278FC8D70F940DA29B3091E1D" bold="true" italics="true" pageId="40" pageNumber="119">Structure.</emphasis>
Head: clypeus protuberant, in profile 0.5 width of eye; galea in repose reaching half-way between fore and mid coxae; length of f1 2.8
<normalizedToken id="D09F47990F589675549B9C36BBBE5BCC" originalValue="×">x</normalizedToken>
length of f2, 0.9
<normalizedToken id="608B399DAFAAD931FC4B73A86F3B36E1" originalValue="×">x</normalizedToken>
@ -163,17 +211,36 @@ width; IOD 1.4
<normalizedToken id="AD4C8AFEB82B75593ABF371795A881CA" originalValue="×">x</normalizedToken>
OOD; OS 0.8
<normalizedToken id="C552F95728EFA600803811E8DEB93B12" originalValue="×">x</normalizedToken>
OOD. Coloration. Yellow marks on labrum, mandibles, scape, clypeus, paraocular and supraclypeal areas; inverted T-shape on clypeus. Pubescence. Head: labrum white, remaining areas predominantly pale yellow with scattered black robust hairs on clypeus, paraocular areas, between antennae, near ocelli and on vertex; gena white. Thorax: scutum ginger intermixed with black hairs; pleura ginger with scattered black hairs under wing base, white ventrally; thoracic sterna white; propodeum laterally light ginger with scattered black hairs. Legs: forefemur posteriorly with long white hairs, outer surface of fore tibia and tarsus white, inner surface of tarsus dark; mid legs dark, except white hairs on apex of the femur, posteriorly on proximal one third of femur and on outer surface of tibia and basitarsus; hind legs black, except white hairs on apex of femur and outer surface of tibia; basitibial streak black, 0.6-0.9 length of femur. Metasoma: apical hair bands on T1-T4 white with iridescence varying from light blue to greenish orange; T5 laterally with long white hairs and few dispersed short hairs (Fig. 32), fimbria black, medial patch around the fimbria, narrowing laterally; S3, S4 dark, posterolateral patches of white hairs; S5 black, laterally with small white patches. Punctation. Head: clypeus with close, medium sized, deep punctures, 0.3-1.0 puncture widths apart; labrum shiny, with close, small punctures of intermediate depth, 0.2-0.9 puncture widths apart, interspaces almost smooth. Thorax: scutum somewhat shiny, with close punctures, 0.2-0.8 puncture widths apart. Metasoma: T1-T5 with close to open punctures, 0.5-2.0 puncture widths apart.
OOD.
<emphasis id="58D6D68E566784F7307999AF32DC5E20" bold="true" pageId="40" pageNumber="119">
<emphasis id="FCD4008A0E3C8A3399730076878330BF" italics="true" pageId="40" pageNumber="119">Coloration</emphasis>
.
</emphasis>
Yellow marks on labrum, mandibles, scape, clypeus, paraocular and supraclypeal areas; inverted T-shape on clypeus.
<emphasis id="CD36DCA025123E5B11CC3D97B8A456A3" bold="true" pageId="40" pageNumber="119">
<emphasis id="020D630F3DA86E9182EDD4113539A2B7" italics="true" pageId="40" pageNumber="119">Pubescence</emphasis>
.
</emphasis>
Head: labrum white, remaining areas predominantly pale yellow with scattered black robust hairs on clypeus, paraocular areas, between antennae, near ocelli and on vertex; gena white. Thorax: scutum ginger intermixed with black hairs; pleura ginger with scattered black hairs under wing base, white ventrally; thoracic sterna white; propodeum laterally light ginger with scattered black hairs. Legs: forefemur posteriorly with long white hairs, outer surface of fore tibia and tarsus white, inner surface of tarsus dark; mid legs dark, except white hairs on apex of the femur, posteriorly on proximal one third of femur and on outer surface of tibia and basitarsus; hind legs black, except white hairs on apex of femur and outer surface of tibia; basitibial streak black, 0.6-0.9 length of femur. Metasoma: apical hair bands on T1-T4 white with iridescence varying from light blue to greenish orange; T5 laterally with long white hairs and few dispersed short hairs (Fig.
<figureCitation id="D84BD4ECFC8E53F1BBC923C71E1D4A83" captionStart="Figure 2437" captionStartId="F9" captionText="Figure 24 - 37. Apical tergites of females, especially T 5. 24 Amegilla aeruginosa 25 Amegilla chlorocyanea 26 Amegilla adelaidae 27 Amegilla alpha 28 Amegilla asserta 29 Amegilla cingulata 30 Amegilla indistincta 31 Amegilla karlba 32 Amegilla murrayensis 33 Amegilla paeninsulae 34 Amegilla pulchra 35 Amegilla thorogoodi 36 Amegilla viridicingulata 37 Amegilla walkeri." figureDoi="10.3897/zookeys.653.11177.figure24-37" httpUri="https://binary.pensoft.net/fig/121864" pageId="40" pageNumber="119">32</figureCitation>
), fimbria black, medial patch around the fimbria, narrowing laterally; S3, S4 dark, posterolateral patches of white hairs; S5 black, laterally with small white patches.
<emphasis id="D93D357BCD8E95C83F9AD06C328BFB62" bold="true" pageId="40" pageNumber="119">
<emphasis id="E7786267A98C172147377E0DF3543A33" italics="true" pageId="40" pageNumber="119">Punctation</emphasis>
.
</emphasis>
Head: clypeus with close, medium sized, deep punctures, 0.3-1.0 puncture widths apart; labrum shiny, with close, small punctures of intermediate depth, 0.2-0.9 puncture widths apart, interspaces almost smooth. Thorax: scutum somewhat shiny, with close punctures, 0.2-0.8 puncture widths apart. Metasoma: T1-T5 with close to open punctures, 0.5-2.0 puncture widths apart.
</paragraph>
<paragraph id="F7CB3B6A0A3580CF0E63125F86CE5806" pageId="40" pageNumber="119">
Male: Coen,
<geoCoordinate id="147C3D7A76131A4A3BC698CD7BD7C23B" direction="south" orientation="latitude" precision="1" value="-13.94415">13.94415S</geoCoordinate>
<geoCoordinate id="077AA51997E8267E97CF678F0C85C9A1" direction="east" orientation="longitude" precision="1" value="143.20023">143.20022E</geoCoordinate>
<emphasis id="FDC34A2A3B0688A0E20A32893C6AD036" italics="true" pageId="40" pageNumber="119">Male</emphasis>
: Coen,
<geoCoordinate id="147C3D7A76131A4A3BC698CD7BD7C23B" degrees="13.94415" direction="south" orientation="latitude" precision="1" value="-13.94415">13.94415S</geoCoordinate>
<geoCoordinate id="077AA51997E8267E97CF678F0C85C9A1" degrees="143.20022" direction="east" orientation="longitude" precision="1" value="143.20023">143.20022E</geoCoordinate>
, QLD, 27 June 2007, SAMA 32-002584, R. Leijs &amp; M. Batley, DNA voucher RB280 (RL783).
</paragraph>
<paragraph id="E35525935539D006DCF752BC0250E23D" pageId="40" pageNumber="119">Length 11 mm; forewing length 8 mm.</paragraph>
<paragraph id="07E8F1C4633D7149CBC5B3E5FFC2E914" pageId="40" pageNumber="119">
Structure. Head: shortest distance between eyes 0.7 length of eye; clypeus protuberant, in profile 0.5 width of eye; galea in repose reaching halfway between fore and mid coxae; length of f1 1.5
<emphasis id="FFF058E1D3895BDC1574F3A35AAA2ACB" bold="true" italics="true" pageId="40" pageNumber="119">Structure.</emphasis>
Head: shortest distance between eyes 0.7 length of eye; clypeus protuberant, in profile 0.5 width of eye; galea in repose reaching halfway between fore and mid coxae; length of f1 1.5
<normalizedToken id="1F86B78FFEE6EDE701959304EF71CA99" originalValue="×">x</normalizedToken>
length of f2, 0.5
<normalizedToken id="5747F882A330B23B156978ABDD3802EE" originalValue="×">x</normalizedToken>
@ -189,9 +256,21 @@ OOD. Wings: length of marginal cell 0.8
<normalizedToken id="568A0BACFE40FDBD50B851B09D1B6F90" originalValue="×">x</normalizedToken>
distance from apex of marginal cell to wing tip; length of vein M of hind wing 2.6 times as long as second abscissa of M+Cu; length of jugal lobe about 0.5
<normalizedToken id="FB2C0CB731BAF07E978D621C050458B5" originalValue="×">x</normalizedToken>
length of vannal lobe. Metasoma: apicomedial emargination of S5 wide and deep; S7 windows small, median hair brush 3
<normalizedToken id="9BBE8C884D0E3FE82EF740911CD053E0" originalValue="4×">-4x</normalizedToken>
width, lateral wings of hair brush well developed making an angle of 60° with long axis of brush (Fig. 45); S8 apical emargination deep. Pubescence. Head: labrum white, remaining areas predominantly pale brown with scattered black robust hairs on clypeus, paraocular areas, between antennae, near ocelli and on vertex; gena white. Thorax: scutum light brown intermixed with black hairs; pleura brown with scattered black hairs under wing base, white ventrally; propodeum laterally light brown with scattered black hairs. Legs: foreleg: femur posteriorly with long white hairs, outer surface of tibia and tarsus white, inner surface of and tarsus dark; mid legs dark, except white hairs on the apex of femur, posteriorly on proximal one third of femur and on outer surface of tibia and basitarsus; hind legs black, except white hairs on apex of femur and outer surface of tibia, small white patch on outer base of basitarsus. Metasoma: apical hair bands on T1-T5 white; T6, T7 black when viewed from behind, light brown when viewed from the side; S2-S5: S3, S4 medially dark, lateral thirds white, S5 medially dark, lateral quarters white. Punctation. Head: clypeus with punctures 1.0-3.0 puncture widths apart; labrum with medium, shallow punctures 0.5-1.5 puncture widths apart. Thorax: scutum shiny, with close, small, shallow punctures 0.3-1.0 puncture widths apart, interspaces smooth. Metasoma: T1-T5 with shiny, open, fine, shallow punctures, 1.0-3.0 puncture widths apart.
length of vannal lobe. Metasoma: apicomedial emargination of S5 wide and deep; S7 windows small, median hair brush 3-4
<normalizedToken id="79AB6F966B48F18C5D5E05FC5CAC7A56" originalValue="×">x</normalizedToken>
width, lateral wings of hair brush well developed making an angle of 60° with long axis of brush (Fig.
<figureCitation id="8F9E6A2D724CF2D3F2A86BC78E3C53CB" captionStart="Figures 3850" captionStartId="F10" captionText="Figures 38 - 50. S 7 of males. 38 Amegilla aeruginosa 39 Amegilla chlorocyanea 40 Amegilla adelaidae 41 Amegilla asserta 42 Amegilla cingulata 43 Amegilla indistincta 44 Amegilla karlba 45 Amegilla murrayensis 46 Amegilla paeninsulae 47 Amegilla pulchra 48 Amegilla thorogoodi 49 Amegilla viridicingulata 50 Amegilla walkeri." figureDoi="10.3897/zookeys.653.11177.figures38-50" httpUri="https://binary.pensoft.net/fig/121865" pageId="40" pageNumber="119">45</figureCitation>
); S8 apical emargination deep.
<emphasis id="16C5C9BCF6F12103B1DD3B9776BC8690" bold="true" pageId="40" pageNumber="119">
<emphasis id="4583626499687FB852794711BEF54162" italics="true" pageId="40" pageNumber="119">Pubescence</emphasis>
.
</emphasis>
Head: labrum white, remaining areas predominantly pale brown with scattered black robust hairs on clypeus, paraocular areas, between antennae, near ocelli and on vertex; gena white. Thorax: scutum light brown intermixed with black hairs; pleura brown with scattered black hairs under wing base, white ventrally; propodeum laterally light brown with scattered black hairs. Legs: foreleg: femur posteriorly with long white hairs, outer surface of tibia and tarsus white, inner surface of and tarsus dark; mid legs dark, except white hairs on the apex of femur, posteriorly on proximal one third of femur and on outer surface of tibia and basitarsus; hind legs black, except white hairs on apex of femur and outer surface of tibia, small white patch on outer base of basitarsus. Metasoma: apical hair bands on T1-T5 white; T6, T7 black when viewed from behind, light brown when viewed from the side; S2-S5: S3, S4 medially dark, lateral thirds white, S5 medially dark, lateral quarters white.
<emphasis id="B92C2CFCBF09313D42EBCB86BAA778D5" bold="true" pageId="40" pageNumber="119">
<emphasis id="02E8C09678DD50836381F28F967B4FB4" italics="true" pageId="40" pageNumber="119">Punctation</emphasis>
.
</emphasis>
Head: clypeus with punctures 1.0-3.0 puncture widths apart; labrum with medium, shallow punctures 0.5-1.5 puncture widths apart. Thorax: scutum shiny, with close, small, shallow punctures 0.3-1.0 puncture widths apart, interspaces smooth. Metasoma: T1-T5 with shiny, open, fine, shallow punctures, 1.0-3.0 puncture widths apart.
</paragraph>
</subSubSection>
<subSubSection id="DAE5259DC3C98B2AC88D61AFB3ECF0A6" lastPageId="41" lastPageNumber="120" pageId="40" pageNumber="119" type="variation">
@ -200,7 +279,9 @@ width, lateral wings of hair brush well developed making an angle of 60° with l
Males from southern Western Australia often have more black hairs on the clypeus and paraclypeal areas than specimens from northern Queensland or specimens from the Lofty Ranges in South Australia. There is also some variation in
<pageBreakToken id="E5BD2B3E90481E12C3DFB96FA5DBAF14" pageId="41" pageNumber="120" start="start">the</pageBreakToken>
width of the pale patch on T5 in fresh females. Some specimens from the NW Pilbarra and Barrow Island are seemingly larger, have wider tergal bands and a more intense patch of pale hairs on female T5 and have almost ivory face marks and white apical bands on T1-4. These specimens are sufficiently different from
<taxonomicName id="393E462FEFBCFDF6E7B4696FA43D6C42" lsidName="murrayensis" pageId="41" pageNumber="120" rank="species" species="murrayensis">murrayensis</taxonomicName>
<taxonomicName id="EB2370237760072E919BB681C8D768E3" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="41" pageNumber="120" phylum="Arthropoda" rank="species" species="murrayensis">
<emphasis id="50417275EE95975A3C69232FAB55E468" italics="true" pageId="41" pageNumber="120">Amegilla murrayensis</emphasis>
</taxonomicName>
and may belong to an undescribed species. Future collections and molecular work may shed light on their identity.
</paragraph>
</subSubSection>
@ -208,19 +289,31 @@ and may belong to an undescribed species. Future collections and molecular work
<paragraph id="2F6555AA3F314E3E0732B4D9A56586A5" pageId="41" pageNumber="120">Remarks.</paragraph>
<paragraph id="EB2AA66D2648E5D0E7F534300EDF893C" pageId="41" pageNumber="120">
There were some problems with the molecular delineation of
<taxonomicName id="A1A6825017EA30BC5822C4357350E5F2" lsidName="murrayensis" pageId="41" pageNumber="120" rank="species" species="murrayensis">murrayensis</taxonomicName>
<taxonomicName id="5CF254A8609CC955FAD7693B5E856FBF" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="41" pageNumber="120" phylum="Arthropoda" rank="species" species="murrayensis">
<emphasis id="795B2137759E0A993009B609C6C82495" italics="true" pageId="41" pageNumber="120">Amegilla murrayensis</emphasis>
</taxonomicName>
and
<taxonomicName id="D9148E884E748CDB61F79E5375E2EEE6" lsidName="pulchra" pageId="41" pageNumber="120" rank="species" species="pulchra">pulchra</taxonomicName>
<taxonomicName id="A2D578E9721E638C0B88D06D027FA541" baseAuthorityName="Smith" baseAuthorityYear="1854" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla pulchra" order="Hymenoptera" pageId="41" pageNumber="120" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="BED58E21C5DDE66D071C2FE933A13B7E" italics="true" pageId="41" pageNumber="120">Amegilla pulchra</emphasis>
</taxonomicName>
, as mentioned in the Results and Discussion section. The sequences obtained with the M202/M70 primers resulted in two morphologically similar clades of
<taxonomicName id="56F2591BED80D32403618166E1BA79A6" lsidName="murrayensis" pageId="41" pageNumber="120" rank="species" species="murrayensis">murrayensis</taxonomicName>
<taxonomicName id="17023DBD214C4F4123ED905DFAAADB20" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="41" pageNumber="120" phylum="Arthropoda" rank="species" species="murrayensis">
<emphasis id="C28F1C2C6037F3F037F5F45013A5AE1E" italics="true" pageId="41" pageNumber="120">Amegilla murrayensis</emphasis>
</taxonomicName>
specimens, often with specimens from the same localities in different clades (Suppl. material 1: Fig. S1). It also resulted in a number of specimens now believed to be
<taxonomicName id="993B17CBFB80D73B334FB76920EC64A7" lsidName="pulchra" pageId="41" pageNumber="120" rank="species" species="pulchra">pulchra</taxonomicName>
<taxonomicName id="36377EBB7CC03747FDB32AB12AD312CE" baseAuthorityName="Smith" baseAuthorityYear="1854" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla pulchra" order="Hymenoptera" pageId="41" pageNumber="120" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="420FF069D9A54B958D4857E48E5DB87B" italics="true" pageId="41" pageNumber="120">Amegilla pulchra</emphasis>
</taxonomicName>
to appear in one of the
<taxonomicName id="B9ACF672C0540697B73E3E7E63ED28A5" lsidName="murrayensis" pageId="41" pageNumber="120" rank="species" species="murrayensis">murrayensis</taxonomicName>
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<emphasis id="3F7CAA07AD92E24F37D48C71AC133A59" italics="true" pageId="41" pageNumber="120">Amegilla murrayensis</emphasis>
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clades. Using the CO1 barcoding primers (M414/M423,
<bibRefCitation id="59CC4B1727F58438C681A640EF7BF1A2" author="Hebert, PDN" pageId="56" pageNumber="135" url="https://doi.org/10.1073/pnas.0406166101" year="2004">Hebert et al. 2004</bibRefCitation>
<bibRefCitation id="59CC4B1727F58438C681A640EF7BF1A2" DOI="https://doi.org/10.1073/pnas.0406166101" author="Hebert, PDN" pageId="56" pageNumber="135" refId="B26" refString="Hebert, PDN, Penton, EH, Burns, JM, Janzen, DH, Hallwachs, W, 2004. Ten species in one: DNA barcoding reveals cryptic species in the neo-tropical skipper butterfly Astraptes fulgerator. Proceedings National Academy of Sciences, USA 101: 14812-14817. https://doi.org/10.1073/pnas.0406166101" title="Ten species in one: DNA barcoding reveals cryptic species in the neo-tropical skipper butterfly Astraptes fulgerator. Proceedings National Academy of Sciences, USA 101: 14812 - 14817." url="https://doi.org/10.1073/pnas.0406166101" year="2004">Hebert et al. 2004</bibRefCitation>
) the two
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clades collapsed into one (Suppl. material 1: Fig. S2). These problems may have been caused by amplification of a mitochondrial copy in the nuclear genome.
</paragraph>
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@ -243,24 +336,60 @@ clades collapsed into one (Suppl. material 1: Fig. S2). These problems may have
<th id="711E376CD22557DA4D51F91B1F3FEE02" colspan="1" pageId="41" pageNumber="120" rowspan="1">Nov</th>
<th id="A105986A1178A80399FFA06CE292FC26" colspan="1" pageId="41" pageNumber="120" rowspan="1">Dec</th>
</tr>
<tr id="1AFB7D315A06BD166590D524E31F5B2A" pageId="41" pageNumber="120">
<td id="0F6856D41757FEAD255D49B1FE11D96A" colspan="1" pageId="41" pageNumber="120" rowspan="1">No of records N of 30°S</td>
<tr id="D54B77D4C76A928A7EAA6BE517A663F0" pageId="41" pageNumber="120">
<td id="0F6856D41757FEAD255D49B1FE11D96A" colspan="1" pageId="41" pageNumber="120" rowspan="1">
<emphasis id="F97B8793EA880AE176EB40C190CCABD4" bold="true" pageId="41" pageNumber="120">No of records N of 30°S</emphasis>
:
</td>
<td id="89108F25590D6FCBC2DCAF1CA3E69C31" colspan="1" pageId="41" pageNumber="120" rowspan="1">10</td>
<td id="F91CB4A81AE9B2742710379C3BCDFE42" colspan="1" pageId="41" pageNumber="120" rowspan="1">6</td>
<td id="67D56CC4704D74D30D84FB41C4A573B9" colspan="1" pageId="41" pageNumber="120" rowspan="1">24</td>
<td id="7F1DB9AB36356CFF3C46E7A182F97244" colspan="1" pageId="41" pageNumber="120" rowspan="1">10</td>
<td id="B221F67EDC523AE4C48F53605B67F8A7" colspan="1" pageId="41" pageNumber="120" rowspan="1">24</td>
<td id="7C9C355E3E6A7CCFEC2B49A9EB63A35F" colspan="1" pageId="41" pageNumber="120" rowspan="1">4</td>
<td id="5A4A4BCA4A5E25C9575C874004F12FD5" colspan="1" pageId="41" pageNumber="120" rowspan="1">7</td>
<td id="A67BC9129736D2D386B5103B4E579B61" colspan="1" pageId="41" pageNumber="120" rowspan="1">25</td>
<td id="F4C4FAF774E99E6CF1A671801E12B19D" colspan="1" pageId="41" pageNumber="120" rowspan="1">4</td>
<td id="BD47D4C987C0BC08A3B83585AA910BD4" colspan="1" pageId="41" pageNumber="120" rowspan="1">11</td>
<td id="680E0594B586F75247AECFFD4C94397B" colspan="1" pageId="41" pageNumber="120" rowspan="1">17</td>
<td id="DC6FCA2905E80D56F9014868E21D0812" colspan="1" pageId="41" pageNumber="120" rowspan="1">5</td>
</tr>
<tr id="664CA9A07E17DAE92E9AF678E09F2827" pageId="41" pageNumber="120">
<td id="04BB29C8D09D54B3B64D7684F4AE6075" colspan="1" pageId="41" pageNumber="120" rowspan="1">No of records S of 30°S</td>
<tr id="BA7A10211ABAB80B098037B65758A6D2" pageId="41" pageNumber="120">
<td id="04BB29C8D09D54B3B64D7684F4AE6075" colspan="1" pageId="41" pageNumber="120" rowspan="1">
<emphasis id="1AC215FAE6CFD075EA490D3382A7FFDD" bold="true" pageId="41" pageNumber="120">No of records S of 30°S</emphasis>
:
</td>
<td id="9EBEA96619D9A70B01FDE904B0358370" colspan="1" pageId="41" pageNumber="120" rowspan="1">161</td>
<td id="807A3A8F9C9174F168BEAA75E5BB47BD" colspan="1" pageId="41" pageNumber="120" rowspan="1">16</td>
<td id="F2E62B9C529308B23A06173B7E4F3A1A" colspan="1" pageId="41" pageNumber="120" rowspan="1">6</td>
<td id="814D90DAFF02A471C59C1B6A6E7538AC" colspan="1" pageId="41" pageNumber="120" rowspan="1">0</td>
<td id="D3319D5DC77661594B3D57235B87744C" colspan="1" pageId="41" pageNumber="120" rowspan="1">0</td>
<td id="3812F21EA9099975D53819DC88513356" colspan="1" pageId="41" pageNumber="120" rowspan="1">0</td>
<td id="F474D6F2A9928C8C26DFB5C5D57D922C" colspan="1" pageId="41" pageNumber="120" rowspan="1">1</td>
<td id="5AB83CE7220AA6DE992731F19B84F456" colspan="1" pageId="41" pageNumber="120" rowspan="1">0</td>
<td id="B3B009D73AB10AF00CD3C63E533E77AB" colspan="1" pageId="41" pageNumber="120" rowspan="1">0</td>
<td id="DBFD954C947A41B7CEF83824D954E0D5" colspan="1" pageId="41" pageNumber="120" rowspan="1">0</td>
<td id="44946EFF26C03ED769CD23E598182718" colspan="1" pageId="41" pageNumber="120" rowspan="1">7</td>
<td id="B6057F7E8B6238CFE0FE79CA0AA0FB4C" colspan="1" pageId="41" pageNumber="120" rowspan="1">16</td>
</tr>
</table>
</paragraph>
<paragraph id="9C828F611CEDEA96EDF13211016ADF3D" pageId="41" pageNumber="120">
<taxonomicName id="FB0B9EABB9292329D1CC73DC64D36552" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="41" pageNumber="120" phylum="Arthropoda" rank="species" species="murrayensis">Amegilla murrayensis</taxonomicName>
<taxonomicName id="FB0B9EABB9292329D1CC73DC64D36552" baseAuthorityName="Rayment" baseAuthorityYear="1935" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amegilla murrayensis" order="Hymenoptera" pageId="41" pageNumber="120" phylum="Arthropoda" rank="species" species="murrayensis">
<emphasis id="BABBED8B5BDB5B44E637511655A5D171" italics="true" pageId="41" pageNumber="120">Amegilla murrayensis</emphasis>
</taxonomicName>
is the most widespread and common species in the subgenus
<taxonomicName id="792A63A302CB0CCCD455537000DE2E60" lsidName="(Zonamegilla)" pageId="41" pageNumber="120" rank="subGenus" subGenus="Zonamegilla">Zonamegilla</taxonomicName>
<taxonomicName id="792A63A302CB0CCCD455537000DE2E60" authorityName="Popov" authorityYear="1950" class="Insecta" family="Apidae" genus="Amegilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Zonamegilla)" order="Hymenoptera" pageId="41" pageNumber="120" phylum="Arthropoda" rank="subGenus" subGenus="Zonamegilla">Zonamegilla</taxonomicName>
. Below 30°S the species is active from November until March, with a peak in January. In the north of the continent they can be found year round possibly with a peak in May.
</paragraph>
</subSubSection>
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<paragraph id="EF36E863B7153A3CD3531F2089B4E88C" pageId="41" pageNumber="120">Distribution.</paragraph>
<paragraph id="D1147E3670EC303752CA86B4DBF11BE7" pageId="41" pageNumber="120">Widespread throughout Australia, but not found in Tasmania (Fig. 11).</paragraph>
<paragraph id="D1147E3670EC303752CA86B4DBF11BE7" pageId="41" pageNumber="120">
Widespread throughout Australia, but not found in Tasmania (Fig.
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).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,255 +1,255 @@
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4.
<taxonomicName LSID="83E7F5CE-7191-561E-B4F6-175E5455F878" authority="J. Compton &amp; Schrire" authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus" status="gen. nov.">Sigmoidala J.Compton &amp; Schrire</taxonomicName>
<taxonomicNameLabel pageId="34" pageNumber="35">gen. nov.</taxonomicNameLabel>
<taxonomicName id="E36B210A99C606FE4F9D2ADD179AC62E" ID-CoL="9DNDW" LSID="83E7F5CE-7191-561E-B4F6-175E5455F878" authority="J. Compton &amp; Schrire" authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus" status="gen. nov.">Sigmoidala J.Compton &amp; Schrire</taxonomicName>
<taxonomicNameLabel id="55F22DC20084DC647B6DCFFB269E771B" pageId="34" pageNumber="35">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="diagnosis">
<paragraph pageId="34" pageNumber="35">Diagnosis.</paragraph>
<paragraph pageId="34" pageNumber="35">
<subSubSection id="6381D242F3EB6DA8A6D7C1ACB9EC65C9" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="7FB22BDF2EF0E4CFF3F8A9CB28CE2200" pageId="34" pageNumber="35">Diagnosis.</paragraph>
<paragraph id="F9C5501C312C1986B27C6F969C51EC47" pageId="34" pageNumber="35">
The monospecific
<taxonomicName authorityName="J. Compton &amp; Schrire" authorityYear="2019" baseAuthorityName="J. Compton &amp; Schrire" baseAuthorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala kityana" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="kityana">
<emphasis italics="true" pageId="34" pageNumber="35">Sigmoidala kityana</emphasis>
<taxonomicName id="A495D6011BE2035EF08A8E3FDF10E004" authorityName="J. Compton &amp; Schrire" authorityYear="2019" baseAuthorityName="J. Compton &amp; Schrire" baseAuthorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala kityana" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="kityana">
<emphasis id="1F27F6BB39AA93B9276A43B1305171C9" italics="true" pageId="34" pageNumber="35">Sigmoidala kityana</emphasis>
</taxonomicName>
has several affinities with
<taxonomicName authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" baseAuthorityName="Roxb." class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara racemosa" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="racemosa">
<emphasis italics="true" pageId="34" pageNumber="35">Endosamara racemosa</emphasis>
<taxonomicName id="FB441F3C737E465337EE632D6F48B1AA" authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" baseAuthorityName="Roxb." class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara racemosa" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="racemosa">
<emphasis id="3599BA41C08670AF4907B6CE392DB342" italics="true" pageId="34" pageNumber="35">Endosamara racemosa</emphasis>
</taxonomicName>
including the absence of bracteoles and glabrous ovaries, characters which also separate it from
<taxonomicName authorityName="Endl., Gen. Pl. Suppl. 3: 104" authorityYear="1843" class="Magnoliopsida" family="Leguminosae" genus="Callerya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Callerya" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Callerya</emphasis>
<taxonomicName id="A52A12B354B9D72D6D1C8144FD8FD50B" authorityName="Endl., Gen. Pl. Suppl. 3: 104" authorityYear="1843" class="Magnoliopsida" family="Leguminosae" genus="Callerya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Callerya" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="F045E8709ECCEF5954BBFF4D4DE0FB3E" italics="true" pageId="34" pageNumber="35">Callerya</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="34" pageNumber="35">s.str.</emphasis>
<emphasis id="54C450792FB35D9A596BCD227D0450B9" italics="true" pageId="34" pageNumber="35">s.str.</emphasis>
which has bracteoles and sericeous ovaries.
<taxonomicName authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Sigmoidala</emphasis>
<taxonomicName id="F3D5D0D9332581BA503AE39B9069B8DD" authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="07F2C5AB79A619C7775645D52EC80C9A" italics="true" pageId="34" pageNumber="35">Sigmoidala</emphasis>
</taxonomicName>
also shares with
<taxonomicName authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
<taxonomicName id="12711EE7824A5BE1054089C7815EA8F9" authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="85449205CFBBC1A0436F3FFA4A3A834A" italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
</taxonomicName>
the pubescent floral bracts and broadly campanulate, slightly oblique, subtruncate calyx as noted by
<bibRefCitation author="Schot, AM" journalOrPublisher="Blumea" pageId="76" pageNumber="77" pagination="1 - 40" refId="B91" refString="Schot, AM, 1994. A Revision of Callerya Endl. (including Padbruggea and Whitfordiodendron) (Papilionaceae: Millettieae). Blumea 39: 1 - 40" title="A Revision of Callerya Endl. (including Padbruggea and Whitfordiodendron) (Papilionaceae: Millettieae)." volume="39" year="1994">Schot (1994</bibRefCitation>
<bibRefCitation id="83BD98F3B01C3CF7EE4BBD5DC5A5F9C9" author="Schot, AM" journalOrPublisher="Blumea" pageId="76" pageNumber="77" pagination="1 - 40" refId="B91" refString="Schot, AM, 1994. A Revision of Callerya Endl. (including Padbruggea and Whitfordiodendron) (Papilionaceae: Millettieae). Blumea 39: 1 - 40" title="A Revision of Callerya Endl. (including Padbruggea and Whitfordiodendron) (Papilionaceae: Millettieae)." volume="39" year="1994">Schot (1994</bibRefCitation>
: 25) but it was placed in
<taxonomicName authorityName="Endl., Gen. Pl. Suppl. 3: 104" authorityYear="1843" class="Magnoliopsida" family="Leguminosae" genus="Callerya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Callerya" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Callerya</emphasis>
<taxonomicName id="BB1100A0C11654CDDD3544D5C80B2810" authorityName="Endl., Gen. Pl. Suppl. 3: 104" authorityYear="1843" class="Magnoliopsida" family="Leguminosae" genus="Callerya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Callerya" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="654D55B8C38E5FF9B79A79CA6B1A3B23" italics="true" pageId="34" pageNumber="35">Callerya</emphasis>
</taxonomicName>
on account of the fruits that lacked the lomented endocarp. The stipules in
<taxonomicName authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Sigmoidala</emphasis>
<taxonomicName id="43CFD7492E30A7450EDAB5BF43A69ECB" authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="C3161B9667600324A030E6308CD95A9E" italics="true" pageId="34" pageNumber="35">Sigmoidala</emphasis>
</taxonomicName>
are shorter, 3-7 mm long (vs. 9-12 mm in
<taxonomicName authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
<taxonomicName id="4792CECE8EA8344F834DA2B7CC3E15D9" authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="B2BE1B940123F618FBC1C638C38FE424" italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
</taxonomicName>
); pedicels shorter, 3-4 mm long (vs. 4-12 mm in
<taxonomicName authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
<taxonomicName id="AA64B4EC822AC2518CBEE4692448BF3A" authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="F3A08212AFEAE9D554F4CD4D9CF7003C" italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Lour., Fl. Cochinch. 2: 462" authorityYear="1790" class="Magnoliopsida" family="Leguminosae" genus="Sarcodum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sarcodum" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Sarcodum</emphasis>
<taxonomicName id="240CFFD8E60E962FEBBDD66F9C4F84F3" authorityName="Lour., Fl. Cochinch. 2: 462" authorityYear="1790" class="Magnoliopsida" family="Leguminosae" genus="Sarcodum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sarcodum" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="F4B87C0ED74D2F9BC791FA0CD53B479C" italics="true" pageId="34" pageNumber="35">Sarcodum</emphasis>
</taxonomicName>
), floral bracts linear, 6-8 mm long (vs. linear-lanceolate, 8-12 mm long in
<taxonomicName authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
<taxonomicName id="B2FC677F45691EB3EE6D16D6B349FEAB" authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="8BF951513C6C68504DE5045C7D161DD0" italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
</taxonomicName>
, 6-20 mm in
<taxonomicName authorityName="Lour., Fl. Cochinch. 2: 462" authorityYear="1790" class="Magnoliopsida" family="Leguminosae" genus="Sarcodum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sarcodum" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Sarcodum</emphasis>
<taxonomicName id="704747C9BA21F8EB10B52E9E47E9E04E" authorityName="Lour., Fl. Cochinch. 2: 462" authorityYear="1790" class="Magnoliopsida" family="Leguminosae" genus="Sarcodum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sarcodum" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="5E37A72943807F5EBBCAFA54B675486A" italics="true" pageId="34" pageNumber="35">Sarcodum</emphasis>
</taxonomicName>
); the back of the standard densely, appressed rufous pubescent (vs. glabrous in
<taxonomicName authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
<taxonomicName id="44904BDDBEFDCD8D9D6FA33ADAED61B0" authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="8AF2EB51183FF4EB1A04A769704B756B" italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Lour., Fl. Cochinch. 2: 462" authorityYear="1790" class="Magnoliopsida" family="Leguminosae" genus="Sarcodum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sarcodum" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Sarcodum</emphasis>
<taxonomicName id="FDFFE25D72B68757D0A0F11762751F31" authorityName="Lour., Fl. Cochinch. 2: 462" authorityYear="1790" class="Magnoliopsida" family="Leguminosae" genus="Sarcodum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sarcodum" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="35AB0FEA9B8FE6743EEDDEA75E6142D0" italics="true" pageId="34" pageNumber="35">Sarcodum</emphasis>
</taxonomicName>
) and the wing petals of
<taxonomicName authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Sigmoidala</emphasis>
<taxonomicName id="4730FAB1CC0C6273FAA5EF1C2E54F35A" authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="247FA29B594F2812C3AB75F8166B3076" italics="true" pageId="34" pageNumber="35">Sigmoidala</emphasis>
</taxonomicName>
are unique within Tribe
<taxonomicName authorityName="Zhu, Cathaya 6: 115 - 124" authorityYear="1994" family="Leguminosae" genus="Wisterieae" kingdom="Plantae" lsidName="Wisterieae" order="Fabales" pageId="34" pageNumber="35" rank="genus">Wisterieae</taxonomicName>
<taxonomicName id="FF268B3E3ED754D724671B285D604981" authorityName="Zhu, Cathaya 6: 115 - 124" authorityYear="1994" class="Magnoliopsida" family="Leguminosae" genus="Wisterieae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Wisterieae" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">Wisterieae</taxonomicName>
being a sigmoid shape, reflexed at the midpoint and extending outwards towards the apex (see Fig.
<figureCitation captionStart="Figure 3" captionStartId="F6" captionText="Figure 3. Sigmoidala kityana (Craib) J. Compton &amp; Schrire. A Habit B young leaf C lower surface of leaf D leaflet detail of hairs E inflorescence F flower bud with bracteole and pedicel G calyx external surface H standard petal inner surface I wing petal J keel petal K staminal column lateral view L staminal column ventral view M stamen dorsal and ventral views N ovary lateral view O style and stigma P pod Q pod detail of surface R seed ventral view S seed lateral view (all from Clark 245). Drawn by Margaret Tebbs." figureDoi="10.3897/phytokeys.125.34877.figure3" httpUri="https://binary.pensoft.net/fig/313256" pageId="34" pageNumber="35">3I</figureCitation>
<figureCitation id="05BB806F9DFDBA60D0622E733C9E7D6A" captionStart="Figure 3" captionStartId="F6" captionText="Figure 3. Sigmoidala kityana (Craib) J. Compton &amp; Schrire. A Habit B young leaf C lower surface of leaf D leaflet detail of hairs E inflorescence F flower bud with bracteole and pedicel G calyx external surface H standard petal inner surface I wing petal J keel petal K staminal column lateral view L staminal column ventral view M stamen dorsal and ventral views N ovary lateral view O style and stigma P pod Q pod detail of surface R seed ventral view S seed lateral view (all from Clark 245). Drawn by Margaret Tebbs." figureDoi="10.3897/phytokeys.125.34877.figure3" httpUri="https://binary.pensoft.net/fig/313256" pageId="34" pageNumber="35">3I</figureCitation>
). The pods of
<taxonomicName authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Sigmoidala</emphasis>
<taxonomicName id="723E60B352164FA9B48D4F9B63C3696F" authorityName="J. Compton &amp; Schrire" authorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="A01CC7A236A3C928A3F2E4B21ED673B1" italics="true" pageId="34" pageNumber="35">Sigmoidala</emphasis>
</taxonomicName>
are flattened, linear to obovate, 7-11
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="A6FED604B4604D33FA4408467C1135DF" originalValue="×">x</normalizedToken>
1-2 cm (vs. septate, flattened, linear, 10-25
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="525452C994B12C5657906121F6C152AA" originalValue="×">x</normalizedToken>
1-2 cm in
<taxonomicName authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
<taxonomicName id="F420B644A707D9A195AD61D222533BFB" authorityName="R. Geesink, Leiden Bot. Ser. 8: 93" authorityYear="1984" class="Magnoliopsida" family="Fabaceae" genus="Endosamara" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Endosamara" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="7A02FF7E18DEFCD862C1D78BFC225754" italics="true" pageId="34" pageNumber="35">Endosamara</emphasis>
</taxonomicName>
and botuliform in
<taxonomicName authorityName="Lour., Fl. Cochinch. 2: 462" authorityYear="1790" class="Magnoliopsida" family="Leguminosae" genus="Sarcodum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sarcodum" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="34" pageNumber="35">Sarcodum</emphasis>
<taxonomicName id="5AB116ECF13382685D8029AC0EDC08B2" authorityName="Lour., Fl. Cochinch. 2: 462" authorityYear="1790" class="Magnoliopsida" family="Leguminosae" genus="Sarcodum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sarcodum" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">
<emphasis id="BDB2944F731106C220D93C73ED9EFFC9" italics="true" pageId="34" pageNumber="35">Sarcodum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="35" type="type species">
<paragraph pageId="34" pageNumber="35">Type species.</paragraph>
<paragraph pageId="34" pageNumber="35">
<taxonomicName authorityName="J. Compton &amp; Schrire" authorityYear="2019" baseAuthorityName="J. Compton &amp; Schrire" baseAuthorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala kityana" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="kityana">
<emphasis italics="true" pageId="34" pageNumber="35">Sigmoidala kityana</emphasis>
<subSubSection id="DA0707B35F83AEFEF513720DED7AAC77" pageId="34" pageNumber="35" type="type species">
<paragraph id="76549BE204066C8CBE6EB299AC6245A7" pageId="34" pageNumber="35">Type species.</paragraph>
<paragraph id="C156FAEEF6523F1A41F6525C9AE3C921" pageId="34" pageNumber="35">
<taxonomicName id="80B8B8BAA23A8B28AF5D26C0D409D732" authorityName="J. Compton &amp; Schrire" authorityYear="2019" baseAuthorityName="J. Compton &amp; Schrire" baseAuthorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Sigmoidala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sigmoidala kityana" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="kityana">
<emphasis id="5C0394C8CF161128C13655D616218580" italics="true" pageId="34" pageNumber="35">Sigmoidala kityana</emphasis>
</taxonomicName>
(Craib) J.Compton &amp; Schrire ≡
<taxonomicName authorityName="Craib" authorityYear="1927" class="Magnoliopsida" family="Fabaceae" genus="Millettia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Millettia kityana" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="kityana">
<emphasis italics="true" pageId="34" pageNumber="35">Millettia kityana</emphasis>
<taxonomicName id="210A071F8D7B3D9B7EC1AF0B84FBBA33" authorityName="Craib" authorityYear="1927" class="Magnoliopsida" family="Fabaceae" genus="Millettia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Millettia kityana" order="Fabales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="kityana">
<emphasis id="364C3148CF76203AC35B0BDC22B3DF8D" italics="true" pageId="34" pageNumber="35">Millettia kityana</emphasis>
</taxonomicName>
Craib.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="description">
<paragraph pageId="35" pageNumber="36">
<pageBreakToken pageId="35" pageNumber="36" start="start">Genus</pageBreakToken>
<subSubSection id="4C43AD6274D3DEBF5ECAE8A3432A8606" pageId="35" pageNumber="36" type="description">
<paragraph id="7D046AEF340F057A575015B1D668A718" pageId="35" pageNumber="36">
<pageBreakToken id="4581BC5166C0199088AADC8D5BB76A1B" pageId="35" pageNumber="36" start="start">Genus</pageBreakToken>
description..
</paragraph>
<paragraph pageId="35" pageNumber="36">
<paragraph id="2E1300EF441A70B8EA4CEA878A9A1E2A" pageId="35" pageNumber="36">
robust, twining woody vine.
<emphasis italics="true" pageId="35" pageNumber="36">Stems</emphasis>
<emphasis id="F94515DA911D9D9593CD6323BB504B2B" italics="true" pageId="35" pageNumber="36">Stems</emphasis>
very dark green when young, terete.
<emphasis italics="true" pageId="35" pageNumber="36">Leaves</emphasis>
<emphasis id="6F41F604E2BFA3E060944B06671613AA" italics="true" pageId="35" pageNumber="36">Leaves</emphasis>
evergreen, coriaceous and nitid when mature, imparipinnate with 7-9 (- 11) leaflets, rachis 12-30 cm long.
<emphasis italics="true" pageId="35" pageNumber="36">Stipules</emphasis>
<emphasis id="8118A8BADC899BB8D5CB72647B6CABDC" italics="true" pageId="35" pageNumber="36">Stipules</emphasis>
3-7 mm long, narrowly deltoid, persistent.
<emphasis italics="true" pageId="35" pageNumber="36">Stipels</emphasis>
<emphasis id="0C7F091F6DDFA18EEFC7A525B09484A1" italics="true" pageId="35" pageNumber="36">Stipels</emphasis>
3-6 mm long, linear, persistent.
<emphasis italics="true" pageId="35" pageNumber="36">Leaflets</emphasis>
<emphasis id="E329BE6587E0FD3234E19F59902D11C3" italics="true" pageId="35" pageNumber="36">Leaflets</emphasis>
7-18
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9145DCC8BD5D541B33B93754862FF06B" originalValue="×">x</normalizedToken>
2-5 cm, elliptic to narrowly obovate, glabrous above and below, apex cuspidate, margins entire, base cordate.
<emphasis italics="true" pageId="35" pageNumber="36">Inflorescence</emphasis>
<emphasis id="6773287088BA141576460292BDEB457E" italics="true" pageId="35" pageNumber="36">Inflorescence</emphasis>
a robust many-flowered terminal panicle 20-50 cm long, peduncle sparsely hairy.
<emphasis italics="true" pageId="35" pageNumber="36">Flowers</emphasis>
<emphasis id="1AF7FD9D8CDE6428F7FAE93DC1B43084" italics="true" pageId="35" pageNumber="36">Flowers</emphasis>
16-20 mm long, emerging from August - November.
<emphasis italics="true" pageId="35" pageNumber="36">Floral bracts</emphasis>
<emphasis id="66413CCA74337104397992CDB05A60DB" italics="true" pageId="35" pageNumber="36">Floral bracts</emphasis>
6-8 mm long, caducous, linear.
<emphasis italics="true" pageId="35" pageNumber="36">Bracteoles</emphasis>
<emphasis id="C71E9FF25996583DA28B8092C466D917" italics="true" pageId="35" pageNumber="36">Bracteoles</emphasis>
absent.
<emphasis italics="true" pageId="35" pageNumber="36">Pedicels</emphasis>
<emphasis id="01562CFDF339B5106A43A093F49D60B8" italics="true" pageId="35" pageNumber="36">Pedicels</emphasis>
3-4 mm long, glabrous.
<emphasis italics="true" pageId="35" pageNumber="36">Calyx</emphasis>
<emphasis id="E3CCDB95D2AE96DC2831D0D0DA3D7DBB" italics="true" pageId="35" pageNumber="36">Calyx</emphasis>
4
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9E948219C11B2676ABDBB2D831F1A7E0" originalValue="×">x</normalizedToken>
6 mm, campanulate, green, densely puberulent externally, five lobed, lobes
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="A899FEE14F930120951D1FB01F41B017" originalValue="±">+/-</normalizedToken>
equal 1-6 mm long, rounded, obtuse or subtruncate becoming subentire after anthesis.
<emphasis italics="true" pageId="35" pageNumber="36">Standard</emphasis>
<emphasis id="B246882BC3C290FBA23C1AB0C51C9EA9" italics="true" pageId="35" pageNumber="36">Standard</emphasis>
10-12
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="E1CA0F1A15210435565092EE01464282" originalValue="×">x</normalizedToken>
12-13 mm, suborbicular, inner surface white with a pink flush, nectar guide broad, deep golden-yellow, back of standard densely appressed, ferrugineous or rufous pubescent, apex acute or emarginate. Callosities of boss type.
<emphasis italics="true" pageId="35" pageNumber="36">Wing petals</emphasis>
<emphasis id="4A5C9570995C560193BF839BB01EDBA1" italics="true" pageId="35" pageNumber="36">Wing petals</emphasis>
glabrous, longer than keel in length but notably sigmoid towards apex and thereby shortened, each narrowly semi-pandurate 10-14
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="22CA7C06650B23E53CE2DFBFE3B07136" originalValue="×">x</normalizedToken>
3 mm with basal claws 1-3 mm long.
<emphasis italics="true" pageId="35" pageNumber="36">Keel petals</emphasis>
<emphasis id="A85BAEF09010E3735CCA34E3E4AE3C3C" italics="true" pageId="35" pageNumber="36">Keel petals</emphasis>
10-12
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="1ACE03B4384751057D65F152E89D9048" originalValue="×">x</normalizedToken>
4-6 mm, glabrous, united into a long navicular cup, apex acute.
<emphasis italics="true" pageId="35" pageNumber="36">Stamens</emphasis>
<emphasis id="870F189B0E29FE6EC2AB0A9780399F43" italics="true" pageId="35" pageNumber="36">Stamens</emphasis>
diadelphous, nine fused together, the vexillary one free, all curved upwards at apex.
<emphasis italics="true" pageId="35" pageNumber="36">Ovary</emphasis>
<emphasis id="73C2D85FF134A4E42D214DD8BACF388A" italics="true" pageId="35" pageNumber="36">Ovary</emphasis>
glabrous, style glabrous, 3 mm long, curved upwards at apex,
<emphasis italics="true" pageId="35" pageNumber="36">stigma</emphasis>
<emphasis id="D5EDD026C979225715DC7CC9C2661878" italics="true" pageId="35" pageNumber="36">stigma</emphasis>
punctate.
<emphasis italics="true" pageId="35" pageNumber="36">Pods</emphasis>
<emphasis id="185B8FD65BD2AA3DB22587393A675806" italics="true" pageId="35" pageNumber="36">Pods</emphasis>
7-11
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="1933D3B6407A9A6C0C0EADDF75F330CC" originalValue="×">x</normalizedToken>
1-2 cm, flattened, linear to narrowly obovate, dehiscent, exocarp surface glabrous, speckled with small pustules, subseptate.
<emphasis italics="true" pageId="35" pageNumber="36">Seeds</emphasis>
<emphasis id="57C9403082D3B57D38E1D493F387533A" italics="true" pageId="35" pageNumber="36">Seeds</emphasis>
1
<normalizedToken originalValue="5(">-5(-</normalizedToken>
<normalizedToken id="6177E19EB9F98360FD41C5DF3357D66B" originalValue="5(">-5(-</normalizedToken>
8), ellipsoid or orbicular 12-14
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="6656A9B5C4F4B0C134A23BB87ADA1119" originalValue="×">x</normalizedToken>
12-13
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="8002D9B78D538C790F7E5BFF6257179C" originalValue="×">x</normalizedToken>
13 mm, hilum 1.6-2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="2833B2ECBA496659B3F14EEA0201E7BE" originalValue="×">x</normalizedToken>
2 mm, elliptic. Fig.
<figureCitation captionStart="Figure 3" captionStartId="F6" captionText="Figure 3. Sigmoidala kityana (Craib) J. Compton &amp; Schrire. A Habit B young leaf C lower surface of leaf D leaflet detail of hairs E inflorescence F flower bud with bracteole and pedicel G calyx external surface H standard petal inner surface I wing petal J keel petal K staminal column lateral view L staminal column ventral view M stamen dorsal and ventral views N ovary lateral view O style and stigma P pod Q pod detail of surface R seed ventral view S seed lateral view (all from Clark 245). Drawn by Margaret Tebbs." figureDoi="10.3897/phytokeys.125.34877.figure3" httpUri="https://binary.pensoft.net/fig/313256" pageId="35" pageNumber="36">
<figureCitation id="B934889D1035E3A68934BCCA3C0AD377" captionStart="Figure 3" captionStartId="F6" captionText="Figure 3. Sigmoidala kityana (Craib) J. Compton &amp; Schrire. A Habit B young leaf C lower surface of leaf D leaflet detail of hairs E inflorescence F flower bud with bracteole and pedicel G calyx external surface H standard petal inner surface I wing petal J keel petal K staminal column lateral view L staminal column ventral view M stamen dorsal and ventral views N ovary lateral view O style and stigma P pod Q pod detail of surface R seed ventral view S seed lateral view (all from Clark 245). Drawn by Margaret Tebbs." figureDoi="10.3897/phytokeys.125.34877.figure3" httpUri="https://binary.pensoft.net/fig/313256" pageId="35" pageNumber="36">
3
<normalizedToken originalValue="CD">C-D</normalizedToken>
<normalizedToken id="00F5D2817202B18957C1C739DED11E18" originalValue="CD">C-D</normalizedToken>
</figureCitation>
.
</paragraph>