Add updates up until 2024-07-04 07:50:13

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ggserver 2024-07-12 01:09:19 +00:00
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<paragraph pageNumber="688">RCN: 3603.</paragraph>
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<paragraph id="9E2C75A017FF97C73096DEE59B58394A" pageNumber="688">RCN: 3603.</paragraph>
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<paragraph pageNumber="688">Type not designated.</paragraph>
<subSubSection id="548464DA8411E4A35CCF17ACD4018CC7" pageNumber="688" type="materials_examined">
<paragraph id="446B10A61948238DCCBA9A0E1666528F" pageNumber="688">Type not designated.</paragraph>
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<paragraph pageNumber="688">Original material: none traced.</paragraph>
<subSubSection id="DC9B9788F7327DC0FB24211195B2B3B8" pageNumber="688" type="materials_examined">
<paragraph id="B52C61D1D8F97F2C50926E545490B51E" pageNumber="688">Original material: none traced.</paragraph>
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<subSubSection id="57A76AA3AEB50CAA3ABA3313490C0DBA" pageNumber="688" type="nomenclature">
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Current name:
<emphasis bold="true" pageNumber="688">
<taxonomicName authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Myrtaceae" genus="Myrtus" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" phylum="Tracheophyta" rank="genus">
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sp.
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(
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<taxonomicName id="B4E374E880D2EE5DD5205CB1E47F26D7" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
).
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<emphasis italics="true" pageNumber="688">Note:</emphasis>
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<emphasis id="78A017F3F203888BC48CC0EA20F67BA9" italics="true" pageNumber="688">Note:</emphasis>
The application of this name appears uncertain.
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@ -1,108 +1,108 @@
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<bibCitation author="Linnaeus" pagination="1060" title="Mesembryanthemum edule" volumeTitle="Systema Naturae, ed. 10, 2" year="1759">
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<emphasis bold="true" pageNumber="669">Mesembryanthemum edule</emphasis>
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<emphasis id="B529E33778D136E2A64598772430BBA1" bold="true" pageNumber="669">Mesembryanthemum edule</emphasis>
Linnaeus
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,
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<emphasis id="6BFBBB4857BFFA28EDA393F8089A5A6D" italics="true" pageNumber="669">Systema Naturae</emphasis>
, ed. 10, 2
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<paragraph pageNumber="669">[&quot;Habitat ad Cap. b. spei.&quot;] Sp. Pl., ed. 2, 1: 695 (1762). RCN: 3697.</paragraph>
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<paragraph id="4252CFCA64A78B602848EF4EFFF9B94C" pageNumber="669">[&quot;Habitat ad Cap. b. spei.&quot;] Sp. Pl., ed. 2, 1: 695 (1762). RCN: 3697.</paragraph>
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Replaced synonym:
<taxonomicName authority="L. (1753)" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Aizoaceae" genus="Mesembryanthemum" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="acinaciforme" variety="flavum">
<emphasis italics="true" pageNumber="669">Mesembryanthemum acinaciforme</emphasis>
<taxonomicName id="863B505E8CDFC1D40D252531CA150C48" authority="L. (1753)" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Aizoaceae" genus="Mesembryanthemum" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="variety" species="acinaciforme" variety="flavum">
<emphasis id="A3C8A137D326813E7C602A68E9CA6EFA" italics="true" pageNumber="669">Mesembryanthemum acinaciforme</emphasis>
L. var.
<emphasis italics="true" pageNumber="669">flavum</emphasis>
<emphasis id="E35BA694AA61C6CAC02EC73B7A19A6A1" italics="true" pageNumber="669">flavum</emphasis>
L. (1753)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageNumber="669" type="materials_examined">
<paragraph pageNumber="669">
<typeStatus>Lectotype</typeStatus>
<subSubSection id="71718FD8B2C2FD0E5E4ABBFF854E153F" pageNumber="669" type="materials_examined">
<paragraph id="669E253FFA18B4EB807808116F437EA4" pageNumber="669">
<typeStatus id="7AC340DE3F82DCB0E8CB93E79E888E66">Lectotype</typeStatus>
(Blake in
<emphasis italics="true" pageNumber="669">Contr. Queensland Herb</emphasis>
<emphasis id="D8F2DBA0C3356BF2F9E9C552AB7D24EF" italics="true" pageNumber="669">Contr. Queensland Herb</emphasis>
. 7: 17. 1969): [icon] &quot;
<emphasis italics="true" pageNumber="669">
<taxonomicName authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Aizoaceae" genus="Mesembryanthemum" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="falcatum">Mesembryanthemum falcatum</taxonomicName>
<emphasis id="70105E30B5C588E6669F90B56294012B" italics="true" pageNumber="669">
<taxonomicName id="B143F839FF9861B44BD11947DEB42B81" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Aizoaceae" genus="Mesembryanthemum" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="falcatum">Mesembryanthemum falcatum</taxonomicName>
majus, flore amplo luteo
</emphasis>
&quot; in Dillenius, Hort. Eltham. 2: 283, t. 212, f. 272. 1732.
</paragraph>
</subSubSection>
<subSubSection pageNumber="669" type="nomenclature">
<paragraph pageNumber="669">
<subSubSection id="9B6D48395FAAE720E7D4B3C672DCE481" pageNumber="669" type="nomenclature">
<paragraph id="9C7023A75DD75B90D836668B406D807B" pageNumber="669">
Current name:
<taxonomicName authority="(L.) N. E. Br." authorityName="N. E. Br." baseAuthorityName="Linnaeus" baseAuthorityYear="1753" class="Magnoliopsida" family="Aizoaceae" genus="Carpobrotus" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="edulis">
<emphasis italics="true" pageNumber="669">
<emphasis bold="true" italics="true" pageNumber="669">Carpobrotus edulis</emphasis>
<taxonomicName id="9F6E50CFA74D9428752CBB43617BD0DA" ID-CoL="RG7K" authority="(L.) N. E. Br." authorityName="N. E. Br." baseAuthorityName="Linnaeus" baseAuthorityYear="1753" class="Magnoliopsida" family="Aizoaceae" genus="Carpobrotus" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="species" species="edulis">
<emphasis id="5A69D05880FD5F8C9B0144816AEEEA0D" italics="true" pageNumber="669">
<emphasis id="02892F90C87DB6B8394DFA8429C0286B" bold="true" italics="true" pageNumber="669">Carpobrotus edulis</emphasis>
</emphasis>
(L.) N.E. Br.
</taxonomicName>
(
<taxonomicName class="Magnoliopsida" family="Aizoaceae" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="family">Aizoaceae</taxonomicName>
<taxonomicName id="0C7D2BCEC3E01718A9518CAD53D55B65" class="Magnoliopsida" family="Aizoaceae" kingdom="Plantae" order="Caryophyllales" phylum="Tracheophyta" rank="family">Aizoaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageNumber="669" type="discussion">
<paragraph pageNumber="669">
<emphasis italics="true" pageNumber="669">Note:</emphasis>
<subSubSection id="CA4D4E0C1C6576AED92946E2C172C5D2" pageNumber="669" type="discussion">
<paragraph id="C7144E85458150343A4C9FF43F9C25D4" pageNumber="669">
<emphasis id="B1D505B3C0BB5B214C6D38F26C8299CF" italics="true" pageNumber="669">Note:</emphasis>
See Preston &amp; Sell (in
<emphasis italics="true" pageNumber="669">Watsonia</emphasis>
<emphasis id="9A129004B14AEF4AA39FA7E1AC23C993" italics="true" pageNumber="669">Watsonia</emphasis>
17: 237-238, f. 7. 1989) and Jonkers (in
<emphasis italics="true" pageNumber="669">Brit. Cact. Succ. J.</emphasis>
<emphasis id="FF9928B063774610073A22CF5A950B70" italics="true" pageNumber="669">Brit. Cact. Succ. J.</emphasis>
19: 182, f. 6. 2001), who both reproduce the type illustration.
</paragraph>
</subSubSection>

View file

@ -1,174 +1,174 @@
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<taxonomicNameLabel id="D83E8831C7A152EF10F70E887BDC6529" pageId="0" pageNumber="107">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution records of Salsa neneba sp. nov." figureDoi="10.3897/zookeys.1102.82388.figure16" httpUri="https://binary.pensoft.net/fig/686615" pageId="0" pageNumber="107">, 16</figureCitation>
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<paragraph pageId="0" pageNumber="107">Type specimen.</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Holotype</emphasis>
<subSubSection id="7AE6CF5CCD85F23E8710241863CE7E2F" pageId="0" pageNumber="107" type="type specimen">
<paragraph id="0B2B515D81BE6FC452AA191B987AB28F" pageId="0" pageNumber="107">Type specimen.</paragraph>
<paragraph id="1A544FBA0F7FB7A9CC7647E2DCADFFE6" pageId="0" pageNumber="107">
<emphasis id="1566B750860E6268101BEE7A6B0A988E" bold="true" italics="true" pageId="0" pageNumber="107">Holotype</emphasis>
female, Neneba (
<geoCoordinate degrees="8" direction="south" minutes="45" orientation="latitude" precision="925" value="-8.75">8°45'S</geoCoordinate>
<geoCoordinate id="0D5E11ACCEEC171FF0D9AA31A0549F8D" degrees="8" direction="south" minutes="45" orientation="latitude" precision="925" value="-8.75">8°45'S</geoCoordinate>
,
<geoCoordinate degrees="147" direction="east" minutes="30" orientation="longitude" precision="925" value="147.5">147°30'E</geoCoordinate>
<geoCoordinate id="8904AED258BD7742578128521EA3BB4C" degrees="147" direction="east" minutes="30" orientation="longitude" precision="925" value="147.5">147°30'E</geoCoordinate>
, Papua New Guinea), 9 November 1896 (QM S111920).
</paragraph>
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<subSubSection pageId="0" pageNumber="107" type="materials_examined">
<paragraph pageId="0" pageNumber="107">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="107">
<subSubSection id="DE2CF44BB53F6927BD2BBB30635362A6" pageId="0" pageNumber="107" type="materials_examined">
<paragraph id="CB78F2A7AC0821D106F3106CB3923DF7" pageId="0" pageNumber="107">Other material examined.</paragraph>
<paragraph id="0710CB475FB1DB20F26E458176440617" pageId="0" pageNumber="107">
Only known from
<typeStatus>holotype</typeStatus>
<typeStatus id="5024B1AF3574C325D322484E5267350D">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="etymology">
<paragraph pageId="0" pageNumber="107">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="107">The specific epithet is a noun in apposition referring to the type locality.</paragraph>
<subSubSection id="CAA2FA8553FCE31A4F04037DCA9C4E53" pageId="0" pageNumber="107" type="etymology">
<paragraph id="7AA2A3A822C7DE792682CE43D71AB8CB" pageId="0" pageNumber="107">Etymology.</paragraph>
<paragraph id="A54539080D900874590FD043F31B0C4B" pageId="0" pageNumber="107">The specific epithet is a noun in apposition referring to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="diagnosis">
<paragraph pageId="0" pageNumber="107">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="107">
<subSubSection id="BDE007EC3C22CB1D2A5498E52DDD7AC3" pageId="0" pageNumber="107" type="diagnosis">
<paragraph id="747C0646AFC446F9A1129D1424D75503" pageId="0" pageNumber="107">Diagnosis.</paragraph>
<paragraph id="068614CCD165EE7BABDE216FAE939A9E" pageId="0" pageNumber="107">
Males of
<taxonomicName genus="S." lsidName="S. neneba" order="Araneida" pageId="0" pageNumber="107" rank="species" species="neneba">
<emphasis italics="true" pageId="0" pageNumber="107">S. neneba</emphasis>
<taxonomicName id="88E20F9BF88CCAB33F1FD307A90DBF99" lsidName="S. neneba" order="Araneida" pageId="0" pageNumber="107" rank="species" species="neneba">
<emphasis id="CAE60410089E7075F6CAD5E99A720F11" italics="true" pageId="0" pageNumber="107">S. neneba</emphasis>
</taxonomicName>
sp. nov. are unknown. Genitalia of female specimens of
<taxonomicName genus="S." lsidName="S. neneba" order="Araneida" pageId="0" pageNumber="107" rank="species" species="neneba">
<emphasis italics="true" pageId="0" pageNumber="107">S. neneba</emphasis>
<taxonomicName id="2D9CA08AFB5F3F4BC9A7D18DBA2B1F31" lsidName="S. neneba" order="Araneida" pageId="0" pageNumber="107" rank="species" species="neneba">
<emphasis id="6556B5209540F8865F07119E6B67D40C" italics="true" pageId="0" pageNumber="107">S. neneba</emphasis>
</taxonomicName>
sp. nov. can be recognised by the somewhat sinuous antero-lateral edges of the epigyne plate and the transverse edges in the atrium (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15C</figureCitation>
<figureCitation id="CEE9D552EBA2322F1D4BC90735757B9B" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="description">
<paragraph pageId="0" pageNumber="107">Description.</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" pageId="0" pageNumber="107">Male.</emphasis>
<subSubSection id="3CCE3ECC0BB8CB183135D370CFEE97BE" pageId="0" pageNumber="107" type="description">
<paragraph id="58678F39B49DEBC6C1749A84B9410CBB" pageId="0" pageNumber="107">Description.</paragraph>
<paragraph id="6B4772F71A9B754EAD8CB46DE8ED4678" pageId="0" pageNumber="107">
<emphasis id="88C63667F71B990B5CDB95462F6BA21D" bold="true" pageId="0" pageNumber="107">Male.</emphasis>
Unknown.
</paragraph>
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" pageId="0" pageNumber="107">Female</emphasis>
<paragraph id="7D38C1825BAA2A9BC7DE67501A53CEB5" pageId="0" pageNumber="107">
<emphasis id="1EF4BA6088271B2FD393BC57C08A5731" bold="true" pageId="0" pageNumber="107">Female</emphasis>
(based on holotype, QM S111920): Total length 10.5. Carapace 4.5 long, 3.2 wide; dark reddish brown and covered by white setae anteriorly from fovea (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15A</figureCitation>
<figureCitation id="721C8542DE8E66DE459583E46F8428B7" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15A</figureCitation>
). Eye diameter AME 0.23, ALE 0.18, PME 0.20, PLE 0.16; row of eyes: AME 0.68, PME 0.56, PLE 2.25. Chelicerae reddish brown, four promarginal teeth (apical and second basal largest) and three retromarginal teeth (apical smallest). Legs yellowish brown, femora slightly darker (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15A, B</figureCitation>
<figureCitation id="756274BF1636530C10D1D2066E17A251" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15A, B</figureCitation>
). Pedipalp length of segments (femur + patella + tibia + tarsus = total length): 1.2 + 0.5 + 0.8 + 1.1 = 3.6. Leg formula I&gt; II&gt; IV&gt; III; length of segments (femur + patella + tibia + metatarsus + tarsus = total length): I - 3.8 + 1.7 + 3.2 + 2.8 + 1.0 = 12.5, II - 3.5 + 1.6 + 2.6 + 2.6 + 0.9 = 11.2, III - 2.1 + 1.0 + 1.2 + 1.3 + 0.7 = 6.3, IV - 3.3 + 1.4 + 2.3 + 2.3 + 0.8 = 10.1. Labium 0.58 long, 0.77 wide, reddish brown; endites reddish brown (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15B</figureCitation>
<figureCitation id="3B4B610D7F82FB251BDCEAC56C8A7D69" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15B</figureCitation>
). Sternum 2.0 long, 1.8 wide, reddish brown (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15B</figureCitation>
<figureCitation id="289D0993805DCF22EB5AB692C4B5A8B2" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15B</figureCitation>
). Abdomen 6.1 long, 5.0 wide; posterior hump distinct (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15A, B</figureCitation>
<figureCitation id="CDFE183F70FCD5A9CC083C49607D015C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15A, B</figureCitation>
); dorsum colouration poorly preserved, beige with indistinct greyish folium pattern (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15A</figureCitation>
<figureCitation id="D7FB5A43A7BC39BE3138FD91F7F3B0BA" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15A</figureCitation>
); venter olive-brown with two spindle-shaped pale lateral bands (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15B</figureCitation>
<figureCitation id="FDE7C63DD219EA58FDE81AAF5BDCD6B4" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15B</figureCitation>
). Epigyne ca. as long as wide, with sinuous antero-lateral borders and transvers ridges within the atrium (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15C</figureCitation>
<figureCitation id="AD5F33EFB01BD84057C9F322AC7E13B7" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15C</figureCitation>
); scape slightly longer than half the length of the epigyne base, slightly narrowest centrally (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15C</figureCitation>
<figureCitation id="6D5B3DFBE6AEAD1A74D1908FAFDA76EF" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Salsa neneba sp. nov., female holotype (QM S 111920) A dorsal habitus B ventral habitus C epigyne, ventral view. Scale bars: 5 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107">15C</figureCitation>
); epigyne not dissected to investigate posterior and internal morphology as only known from fragile holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="variation">
<paragraph pageId="0" pageNumber="107">Variation.</paragraph>
<paragraph pageId="0" pageNumber="107">Only known from holotype.</paragraph>
<subSubSection id="990FA25DE4BDAC888D2789DFF5C94FFF" pageId="0" pageNumber="107" type="variation">
<paragraph id="20149A89C619CEA9BC96348D1B9C6595" pageId="0" pageNumber="107">Variation.</paragraph>
<paragraph id="6E991067B669098527D6EA904C291C70" pageId="0" pageNumber="107">Only known from holotype.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="life history and habitat preferences">
<paragraph pageId="0" pageNumber="107">Life history and habitat preferences.</paragraph>
<paragraph pageId="0" pageNumber="107">Unknown.</paragraph>
<subSubSection id="7D8549A195C12D122B41E3520C752CCD" pageId="0" pageNumber="107" type="life history and habitat preferences">
<paragraph id="4FD99EEA879EABEBB1D89EAA80629E51" pageId="0" pageNumber="107">Life history and habitat preferences.</paragraph>
<paragraph id="CFBF193DA44928E214AD28CA9609E4D2" pageId="0" pageNumber="107">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="distribution">
<paragraph pageId="0" pageNumber="107">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="107">
<subSubSection id="13025B3C3C07A80A0C088E8DF1AD22A3" pageId="0" pageNumber="107" type="distribution">
<paragraph id="8BEE112BFA78CFE74CEBB1A3076EC74E" pageId="0" pageNumber="107">Distribution.</paragraph>
<paragraph id="90CA3E28C01BBDCA66C84C828B2DFBC7" pageId="0" pageNumber="107">
Currently only known from type locality, Neneba in Papua New Guinea (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution records of Salsa neneba sp. nov." figureDoi="10.3897/zookeys.1102.82388.figure16" httpUri="https://binary.pensoft.net/fig/686615" pageId="0" pageNumber="107">16</figureCitation>
<figureCitation id="F16C21AB36D78EA556825E12423D8F9A" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution records of Salsa neneba sp. nov." figureDoi="10.3897/zookeys.1102.82388.figure16" httpUri="https://binary.pensoft.net/fig/686615" pageId="0" pageNumber="107">16</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" pageId="0" pageNumber="107">Figure 15.</emphasis>
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" class="Liliopsida" family="Araneidae" genus="Salsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salsa neneba" order="Araneae" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="neneba">
<emphasis italics="true" pageId="0" pageNumber="107">Salsa neneba</emphasis>
<caption id="CC61F011C8FF5966D97CCEDC3895A8E2" doi="10.3897/zookeys.1102.82388.figure15" httpUri="https://binary.pensoft.net/fig/686614" pageId="0" pageNumber="107" start="Figure 15" startId="F15">
<paragraph id="07A21C68FDF5E3CB0D1365874DDC6493" pageId="0" pageNumber="107">
<emphasis id="0103F1ABF45A1AA93ECAB0D86BEB89C5" bold="true" pageId="0" pageNumber="107">Figure 15.</emphasis>
<taxonomicName id="65D02F15E4FED9639771D73C3A921ADE" authorityName="Framenau &amp; Castanheira" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Salsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salsa neneba" order="Araneae" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="neneba">
<emphasis id="68FC4A62E1A07DEF405085A237166B2A" italics="true" pageId="0" pageNumber="107">Salsa neneba</emphasis>
</taxonomicName>
sp. nov., female holotype (QM S111920)
<emphasis bold="true" pageId="0" pageNumber="107">A</emphasis>
<emphasis id="475A7559A77C19950D96009CFB1F096A" bold="true" pageId="0" pageNumber="107">A</emphasis>
dorsal habitus
<emphasis bold="true" pageId="0" pageNumber="107">B</emphasis>
<emphasis id="5E296A0BAA449CD35BFC2A48DDD62D5E" bold="true" pageId="0" pageNumber="107">B</emphasis>
ventral habitus
<emphasis bold="true" pageId="0" pageNumber="107">C</emphasis>
<emphasis id="9AAD548C31811806AEA810C88389FF5E" bold="true" pageId="0" pageNumber="107">C</emphasis>
epigyne, ventral view. Scale bars: 5 mm (
<emphasis bold="true" pageId="0" pageNumber="107">A, B</emphasis>
<emphasis id="2F187D2519EB231E95ACC2B76AD76EAA" bold="true" pageId="0" pageNumber="107">A, B</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="107">C</emphasis>
<emphasis id="5D3F65289C884ECD3B6C5657917C4493" bold="true" pageId="0" pageNumber="107">C</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1102.82388.figure16" httpUri="https://binary.pensoft.net/fig/686615" pageId="0" pageNumber="107" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" pageId="0" pageNumber="107">Figure 16.</emphasis>
<caption id="4DE4D2DB6188092F35CD13FF0C17B3DF" doi="10.3897/zookeys.1102.82388.figure16" httpUri="https://binary.pensoft.net/fig/686615" pageId="0" pageNumber="107" start="Figure 16" startId="F16">
<paragraph id="A88AE92888C5CD40858AB2BCAA0C0110" pageId="0" pageNumber="107">
<emphasis id="FFEE3571E6DE523272DB5E447B172740" bold="true" pageId="0" pageNumber="107">Figure 16.</emphasis>
Distribution records of
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" class="Liliopsida" family="Araneidae" genus="Salsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salsa neneba" order="Araneae" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="neneba">
<emphasis italics="true" pageId="0" pageNumber="107">Salsa neneba</emphasis>
<taxonomicName id="790CF7CA7BEA31B8C381B1ADBB1DDEF3" authorityName="Framenau &amp; Castanheira" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Salsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salsa neneba" order="Araneae" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="neneba">
<emphasis id="B2E8D335456EE3A881E23BB343D273CF" italics="true" pageId="0" pageNumber="107">Salsa neneba</emphasis>
</taxonomicName>
sp. nov.
</paragraph>

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<mods:identifier id="0C347BF05C55EFBE8DE7620113A1A36A" type="Zenodo-Dep">12586908</mods:identifier>
</mods:mods>
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<subSubSection id="92F86506FF9DFFA12D91569FFC6A2E80" box="[665,921,439,465]" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="DA5D368DFF9DFFA12D91569FFC6A2E80" blockId="1.[665,921,439,465]" box="[665,921,439,465]" pageId="1" pageNumber="46">
<heading id="811581E1FF9DFFA12D91569FFC6A2E80" box="[665,921,439,465]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="46" reason="2">
<taxonomicName id="1DE24D0EFF9DFFA12D91569FFCB12E80" ID-CoL="5422" authorityName="Papp" authorityYear="2004" box="[665,834,439,465]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E896EA9FFF9DFFA12D91569FFCB12E80" bold="true" box="[665,834,439,465]" pageId="1" pageNumber="46">Immoderatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF9DFFA12C42569FFC6A2E80" box="[842,921,439,465]" pageId="1" pageNumber="46" rank="genus">gen. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="92F86506FF9DFFA32E3556D2FE162B4F" lastPageId="3" lastPageNumber="48" pageId="1" pageNumber="46" type="materials_examined">
<paragraph id="DA5D368DFF9DFFA12E3556D2FD2B2D41" blockId="1.[317,728,506,557]" box="[317,728,506,528]" pageId="1" pageNumber="46">
<typeStatus id="0559882FFF9DFFA12E3556D2FE9D2D41" box="[317,366,506,528]" pageId="1" pageNumber="46">Type</typeStatus>
species:
<taxonomicName id="1DE24D0EFF9DFFA12ECE56D2FD6D2D41" authorityName="Papp" authorityYear="2004" box="[454,670,506,528]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF9DFFA12ECE56D2FD6D2D41" box="[454,670,506,528]" italics="true" pageId="1" pageNumber="46">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF9DFFA12DAC56D2FD2B2D41" box="[676,728,506,528]" pageId="1" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="DA5D368DFF9DFFA12E35553FFE0A2D7C" blockId="1.[317,728,506,557]" box="[317,505,535,557]" pageId="1" pageNumber="46">Gender: masculine.</paragraph>
<paragraph id="DA5D368DFF9DFFA12E765579FB2F2D36" blockId="1.[317,1271,593,1824]" box="[382,1244,593,615]" pageId="1" pageNumber="46">
Body length
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. This is far the smallest species of the subfamily
<taxonomicName id="1DE24D0EFF9DFFA12B405579FB2A2D36" box="[1096,1241,593,615]" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="subFamily" subFamily="Copromyzinae">Copromyzinae</taxonomicName>
.
</paragraph>
<paragraph id="DA5D368DFF9DFFA12E765546FC2C2C1C" blockId="1.[317,1271,593,1824]" pageId="1" pageNumber="46">
Mouth opening large, almost as long as head, palpi small,
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long. Clypeus narrow and small. Antennae in deep holes, as in
<taxonomicName id="1DE24D0EFF9DFFA12C1955A2FC982DF1" box="[785,875,650,672]" class="Insecta" family="Carnidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="family">Carnidae</taxonomicName>
. Pedicel with an extremely long dorsal sub-basal seta. First flagellomere rounded with dorsal arista and with 5 extremely long dorsal setae, which are not much shorter than arista, and with some shorter (still long) ventral setae. Face not longer than antennae. Ocelli extremely anteriorly placed: hind ocelli on border ofanterior and middle third ofpostfrons. Three pairs ofshort lateroclinate fronto-orbitals. Ofthe frontal setae only the outer vertical pair strong, postocellars indistinct. No definite ocellar setae. Two pairs of medium-long setae on fore edge of frons (over lunule). Gena narrow but postgena large.
</paragraph>
<paragraph id="DA5D368DFF9DFFA12E76547CFDC92B01" blockId="1.[317,1271,593,1824]" pageId="1" pageNumber="46">
Thorax dorsoventrally compressed, with considerably reduced sclerites. Prothorax (
<figureCitation id="42D92A08FF9DFFA12BBA547CFB1D2C3B" box="[1202,1262,852,874]" captionStart="Figs 13" captionStartId="2.[317,360,1745,1767]" captionTargetBox="[357,1262,618,1706]" captionTargetId="figure-88@2.[317,1269,614,1711]" captionTargetPageId="2" captionText="Figs 13. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 12, 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12586910" httpUri="https://zenodo.org/record/12586910/files/figure.png" pageId="1" pageNumber="46">Fig. 3</figureCitation>
) consists ofextremely reduced sclerites. Thoracic chaetotaxy: no proepisternal (but several short setu - lae), 1 notopleural, 1 supra-alar and 1 postalar, 1 rather weak dorsocentral (just anterior to scutellum), irregular rows ofsparse and fine acrostichal setulae. Scutellum very short, only
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long but
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broad. One pair oflong lateral (basal) scutellar setae, apical pair present only as minute dorsal setulae. Anepisternum reduced to an oblique sclerite. Anepimeron membranous. Katepisternum complete (i.e. sclerotization not reduced) with 1 (rather posterior) seta. Thorax actually continued caudally below abdomen, hind leg with coxa being wholly ventral to preabdomen, and even mid coxae partly so.
</paragraph>
<paragraph id="DA5D368DFF9DFFA12E76537FFC732BAC" blockId="1.[317,1271,593,1824]" pageId="1" pageNumber="46">
Legs short and thick. Mid tibia without subapical crown ofsetae, or rows ofstronger dorsal setae, but with a strong ventroapical spur and with a shorter and rather thick anterodorsal spur (
<figureCitation id="42D92A08FF9DFFA12BC7535CFEB82BF6" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="1" pageNumber="46">Fig. 5</figureCitation>
). Hind tibia extremely thick, only 3 times longer than thick (
<figureCitation id="42D92A08FF9DFFA12C9453B9FC2B2BF6" box="[924,984,1169,1191]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="1" pageNumber="46">Fig. 7</figureCitation>
), with an extremely long and thick ventroapical spur, a long dorsal seta opposite the base ofthis spur, and 3 pairs ofantero- and postero-dorsal setae. Tarsi extremely short, tarsomeres 24 shorter than broad on all the legs. Mid and hind tarsi with strong black thorns as described below.
</paragraph>
<paragraph id="DA5D368DFF9DFFA12E76522CFCEC2951" blockId="1.[317,1271,593,1824]" pageId="1" pageNumber="46">
<materialsCitation id="6A8A3CD0FF9DFFA12E76522CFCE82951" collectionCode="R" collectorName="Wing &amp; Costal &amp; M. Radial" latitude="0.027" longLatPrecision="78" longitude="-0.03" pageId="1" pageNumber="46" specimenCode="R1, R2" specimenCount="1">
<collectorName id="7717535BFF9DFFA12E76522CFE402A4B" box="[382,435,1284,1306]" pageId="1" pageNumber="46">Wing</collectorName>
peculiar (
<figureCitation id="42D92A08FF9DFFA12D19522CFDBA2A4B" box="[529,585,1284,1306]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="1" pageNumber="46">Fig. 4</figureCitation>
) both in form and venation.
<collectorName id="7717535BFF9DFFA12C5A522CFC7C2A4B" box="[850,911,1284,1306]" pageId="1" pageNumber="46">Costal</collectorName>
vein continued to vein
<collectorName id="7717535BFF9DFFA12B61522CFB3B2A4B" box="[1129,1224,1284,1306]" pageId="1" pageNumber="46">M. Radial</collectorName>
vein
<specimenCode id="8A449EF6FF9DFFA12E355209FEA72A68" box="[317,340,1313,1337]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="46">
R
<subScript id="466634C8FF9DFFA12E455205FEA72A68" attach="left" box="[333,340,1325,1337]" fontSize="5" pageId="1" pageNumber="46">1</subScript>
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(i.e. 1st costal section) extremely short.
<specimenCode id="8A449EF6FF9DFFA12DDE5209FD1E2A68" box="[726,749,1313,1337]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="46">
R
<subScript id="466634C8FF9DFFA12DEE5205FD1E2A68" attach="left" box="[742,749,1325,1337]" fontSize="5" pageId="1" pageNumber="46">2</subScript>
</specimenCode>
+3 very short, second costal section only halfas long as third. First costal section with 2, second and third sections with 1315 very long, almost straight setae dorsally, which reclinate to the wing plane (i.e. not perpendicular but at an acute angle to it). Fourth costal section medially on wing apex with a
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mm
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long, backwardly directed seta. No basal cross-vein, discal cell rounded apically (
<figureCitation id="42D92A08FF9DFFA12DFA52BCFCD82AFB" box="[754,811,1428,1450]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="1" pageNumber="46">Fig. 4</figureCitation>
). Alula practically missing. No anal vein or anal cell, anal region present as a small microtomentose thickening ofwing membrane. Marginal hairs of wing dense and extremely long, on ventral halfofwing consist ofalternating hairs
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<geoCoordinate id="BFD6504AFF9DFFA12B4A52E6FB8B2AB5" box="[1090,1144,1486,1508]" degrees="0.060" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="55" value="0.06">0.060</geoCoordinate>
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mm
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and
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mm
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in length. Halter dark grey
</materialsCitation>
.
</paragraph>
<paragraph id="DA5D368DFF9DFFA12E76512FFB3D2906" blockId="1.[317,1271,593,1824]" pageId="1" pageNumber="46">
Abdomen seemingly robust, but only as a consequence ofcompressed status ofthorax. Ter - gites 15 normal, sternites less sclerotized; sternite 5 rather broad (
<figureCitation id="42D92A08FF9DFFA12CBA510CFBD3296B" box="[946,1056,1572,1594]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="1" pageNumber="46">Figs 1011</figureCitation>
). Abdominal setae, including marginal setae short, except for the comparatively very long pair on female tergite 7.
</paragraph>
<paragraph id="DA5D368DFF9DFFA12E765176FB3629FC" blockId="1.[317,1271,593,1824]" pageId="1" pageNumber="46">
Sternite 8 and a small part ofsternite 6, visible in male in dorsal view (
<figureCitation id="42D92A08FF9DFFA12B1B5176FBBE2925" box="[1043,1101,1630,1652]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="1" pageNumber="46">Fig. 9</figureCitation>
). Sternite 7 com - paratively small, visible only ventrally, its medial part bent into genital cavity. A small subtriangular sclerite present at right apex ofsternite 6 (? tergite 6), forming part ofthe wall ofthe genital cavity.
</paragraph>
<paragraph id="DA5D368DFF9DFFA22E76519CFCB52871" blockId="1.[317,1271,593,1824]" lastBlockId="2.[317,1269,1745,1824]" lastPageId="2" lastPageNumber="47" pageId="1" pageNumber="46">
Male genital morphology peculiar. Epandrium strongly convex (globose), without ventral lateral cleft, but wholly fused and closed ventrally as in some genera of
<taxonomicName id="1DE24D0EFF9DFFA12CC151F9FBB729B6" box="[969,1092,1745,1767]" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="46" phylum="Arthropoda" rank="subFamily" subFamily="Limosininae">Limosininae</taxonomicName>
. Epandrium with a pair ofcomparatively long subdorsal (caudal) setae. Cerci distinct but small, not fused with epand - rium (
<figureCitation id="42D92A08FF9DFFA12E7F5022FE4D2871" box="[375,446,1802,1824]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="1" pageNumber="46">Fig. 10</figureCitation>
), without setae. Subepandrial sclerite not found (i.e. caudal wall of genital chamber is membranous. Surstylus (
<figureCitation id="42D92A08FF9EFFA22D21569CFD9E2E9B" box="[553,621,436,458]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="2" pageNumber="47">Fig. 11</figureCitation>
) long thin, digitiform with a longer medial subapical seta and with 2 small apical setae. Hypandrium small. Phallus (ifmy interpretation is correct) consisting ofa rather normal distiphallus and a peculiar basiphallus with a thick structured dorsal projection, which is almost as large as distiphallus (
<figureCitation id="42D92A08FF9EFFA22D6C5522FD5A2D71" box="[612,681,522,544]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="2" pageNumber="47">Fig. 15</figureCitation>
). Phallapodeme slender, comparatively short, strongly curved in its distal third. Epiphallus seemingly absent. Postgonite long, slender, with narrowly rounded apex for
<figureCitation id="42D92A08FF9EFFA22C065022FCB52871" box="[782,838,1802,1824]" captionStart="Figs 13" captionStartId="2.[317,360,1745,1767]" captionTargetBox="[357,1262,618,1706]" captionTargetId="figure-88@2.[317,1269,614,1711]" captionTargetPageId="2" captionText="Figs 13. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 12, 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12586910" httpUri="https://zenodo.org/record/12586910/files/figure.png" pageId="2" pageNumber="47">Fig. 3</figureCitation>
</paragraph>
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<paragraph id="DA5D368DFF9EFFA22E3551F9FB192852" blockId="2.[317,1269,1745,1824]" pageId="2" pageNumber="47">
<emphasis id="E896EA9FFF9EFFA22E3551F9FE6B29B6" bold="true" box="[317,408,1745,1767]" pageId="2" pageNumber="47">Figs 13.</emphasis>
<taxonomicName id="1DE24D0EFF9EFFA22E9651F9FD8529B6" authorityName="Papp" authorityYear="2004" box="[414,630,1745,1767]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF9EFFA22E9651F9FD8529B6" box="[414,630,1745,1767]" italics="true" pageId="2" pageNumber="47">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF9EFFA22D7451F9FD4229B6" box="[636,689,1745,1767]" pageId="2" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
, paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 12, 0.2 mm
</paragraph>
</caption>
<paragraph id="DA5D368DFF9FFFA32E35569CFE562EB6" blockId="3.[317,1270,436,487]" pageId="3" pageNumber="48">
(
<figureCitation id="42D92A08FF9FFFA32E4C569CFE422E9B" box="[324,433,436,458]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="3" pageNumber="48">Figs 10, 13</figureCitation>
). Ejaculatory apodeme (
<figureCitation id="42D92A08FF9FFFA32D90569CFCF62E9B" box="[664,773,436,458]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="3" pageNumber="48">Figs 1415</figureCitation>
) mushroom-shaped, with an additional sclerite at its distal part.
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E765522FC252D0B" blockId="3.[317,1270,522,602]" pageId="3" pageNumber="48">Etymology. This new genus is characterised by a large set of peculiar features, for I use the word “extremely” frequently in its description. The Latin word (an adjective, used here as a noun for naming a genus) means immoderate, i.e. one who goes to extremes.</paragraph>
<paragraph id="DA5D368DFF9FFFA32E7655A8FE162B4F" blockId="3.[317,1270,640,1054]" pageId="3" pageNumber="48">
Contrary all its unique features and reductions, one can easily identify this new genus as belonging to the
<taxonomicName id="1DE24D0EFF9FFFA32DB0558CFC962DEF" box="[696,869,676,702]" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="subFamily" subFamily="Copromyzinae">Copromyzinae</taxonomicName>
through
<smallCapsWord id="DCBBA051FF9FFFA32CDC558CFBA02DEF" baselines="695,695,695" box="[980,1107,676,702]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Rohacek'" pageId="3" pageNumber="48">ROHÁČEK</smallCapsWord>
s (1998) key, which is still the most comprehensive key for the Palaearctic
<emphasis id="E896EA9FFF9FFFA32B0255EFFBC52DB0" box="[1034,1078,711,737]" italics="true" pageId="3" pageNumber="48">and</emphasis>
Oriental
<taxonomicName id="1DE24D0EFF9FFFA32BAC55EFFE5D2C55" authorityName="Macquart" authorityYear="1835" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="family">Sphaeroceridae</taxonomicName>
. However, its couplet 5 must be supplemented by an additional item to accommodate
<taxonomicName id="1DE24D0EFF9FFFA32EED5425FD732C76" authorityName="Papp" authorityYear="2004" box="[485,640,781,807]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9FFFA32EED5425FD732C76" box="[485,640,781,807]" italics="true" pageId="3" pageNumber="48">Immoderatus</emphasis>
</taxonomicName>
. This may include strong reduction in sclerotization of thoracic pleural sclerites, extremely thick and short legs, extremely strong ventroapical spur on hind tibia, the very anterior positioning ofocelli on frons, extremely long dorsal seta on pedicel and numerous very long setae on first flagellomere, absence ofanal vein and anal cell, absence ofalula, very short vein
<collectionCode id="BCF3AE48FF9FFFA32B3D54B2FBBA2CE5" box="[1077,1097,922,948]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="48">R</collectionCode>
<subScript id="466634C8FF9FFFA32B505481FB822CE6" attach="right" box="[1112,1137,937,951]" fontSize="6" pageId="3" pageNumber="48">2+3</subScript>
, extremely long perpendicular setae on costa and extremely long fringe on the hind half of wing, as well as pecularities ofthe male genitalia and the unique shape ofthe spermathecae.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,347 @@
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<taxonomicName id="1DE24D0EFF9FFFA32D605345FC762BD6" ID-CoL="3PJ93" authorityName="Papp" authorityYear="2004" box="[616,901,1133,1159]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF9FFFA32D605345FC762BD6" bold="true" box="[616,901,1133,1159]" pageId="3" pageNumber="48">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF9FFFA32C845345FC392BD6" box="[908,970,1133,1159]" pageId="3" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
</subSubSection>
<subSubSection id="92F86506FF9FFFA72DDD53B8FC682A16" lastPageId="7" lastPageNumber="52" pageId="3" pageNumber="48" type="description">
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<heading id="811581E1FF9FFFA32DDD53B8FCAE2BFB" box="[725,861,1168,1194]" centered="true" fontSize="11" level="3" pageId="3" pageNumber="48" reason="3">
(
<figureCitation id="42D92A08FF9FFFA32DD553B8FCA72BFB" box="[733,852,1168,1194]" captionStart-0="Figs 13" captionStart-1="Figs 48" captionStart-2="Figs 915" captionStart-3="Figs 1617" captionStartId-0="2.[317,360,1745,1767]" captionStartId-1="4.[317,360,1747,1769]" captionStartId-2="5.[317,360,1722,1744]" captionStartId-3="6.[317,360,1775,1797]" captionTargetBox-0="[357,1262,618,1706]" captionTargetBox-1="[342,1247,578,1717]" captionTargetBox-2="[346,1235,435,1687]" captionTargetBox-3="[321,1265,1159,1725]" captionTargetId-0="figure-88@2.[317,1269,614,1711]" captionTargetId-1="figure-74@4.[339,1247,576,1719]" captionTargetId-2="figure-11@5.[344,1243,424,1694]" captionTargetId-3="figure-362@6.[317,1269,1154,1726]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figs 13. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 12, 0.2 mm" captionText-1="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" captionText-2="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," captionText-3="Figs 1617. Immoderatus foldvarii sp. n., paratype female, genitalia. 16 = apex of abdomen, dorsal view, with cercus in lateral view, 17 = spermathecae. Scale: 0.1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.12586910" figureDoi-1="http://doi.org/10.5281/zenodo.12586912" figureDoi-2="http://doi.org/10.5281/zenodo.12586914" figureDoi-3="http://doi.org/10.5281/zenodo.12586916" httpUri-0="https://zenodo.org/record/12586910/files/figure.png" httpUri-1="https://zenodo.org/record/12586912/files/figure.png" httpUri-2="https://zenodo.org/record/12586914/files/figure.png" httpUri-3="https://zenodo.org/record/12586916/files/figure.png" pageId="3" pageNumber="48">Figs 117</figureCitation>
)
</heading>
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E7653FCFBB52A56" blockId="3.[317,1270,1236,1373]" pageId="3" pageNumber="48">
<materialsCitation id="6A8A3CD0FF9FFFA32E7653FCFBB52A56" collectingDate="2003-12-01" collectorName="Foldvari &amp; Peregovits &amp; Szappanos &amp; No" country="Thailand" location="Mae Ta Man" municipality="Mai" pageId="3" pageNumber="48" specimenCount="1" typeStatus="holotype">
<typeStatus id="0559882FFF9FFFA32E7653FCFE242BBB" box="[382,471,1236,1258]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="A2F5761DFF9FFFA32D1353FCFD692BBB" box="[539,666,1236,1258]" name="Thailand" pageId="3" pageNumber="48">THAILAND</collectingCountry>
,
<location id="DF3D6056FF9FFFA32DAC53FCFCD12BBB" LSID="urn:lsid:plazi:treatment:524B879BFF9FFFA72D605345FC682A16:DF3D6056FF9FFFA32DAC53FCFCD12BBB" box="[676,802,1236,1258]" country="Thailand" municipality="Mai" name="Mae Ta Man" pageId="3" pageNumber="48">Mae Ta Man</location>
elephant park,
<quantity id="1D1A9B68FF9FFFA32CB253FCFC0B2BBB" box="[954,1016,1236,1258]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="48" unit="km" value="45.0">45 km</quantity>
N ofChiang
<collectingMunicipality id="3A39ACF7FF9FFFA32B7F53FCFB532BBB" box="[1143,1184,1236,1258]" pageId="3" pageNumber="48">Mai</collectingMunicipality>
,
<date id="AE5C104DFF9FFFA32BA353FCFE9E2A56" pageId="3" pageNumber="48" value="2003-12-01">
<collectingDate id="BE18E9A5FF9FFFA32BA353FCFE9E2A56" pageId="3" pageNumber="48" value="2003-12-01">01. 12. 2003</collectingDate>
</date>
swept on elephant dung, leg.
<collectorName id="7717535BFF9FFFA32DA053D9FD0F2A56" box="[680,764,1265,1287]" pageId="3" pageNumber="48">Földvári</collectorName>
,
<collectorName id="7717535BFF9FFFA32C0F53D9FC9F2A56" box="[775,876,1265,1287]" pageId="3" pageNumber="48">Peregovits</collectorName>
&amp;
<collectorName id="7717535BFF9FFFA32C8653D9FC042A56" box="[910,1015,1265,1287]" pageId="3" pageNumber="48">Szappanos</collectorName>
,
<collectorName id="7717535BFF9FFFA32B0A53D9FBEC2A56" box="[1026,1055,1265,1287]" pageId="3" pageNumber="48">No</collectorName>
25.
</materialsCitation>
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E765226FE1B2A0C" blockId="3.[317,1270,1236,1373]" pageId="3" pageNumber="48">
<materialsCitation id="6A8A3CD0FF9FFFA32E765226FE172A0C" collectionCode="HNHM" pageId="3" pageNumber="48" specimenCount="37" specimenCount-female="21" specimenCount-male="16" typeStatus="paratype">
<typeStatus id="0559882FFF9FFFA32E765226FE2C2A75" box="[382,479,1294,1316]" pageId="3" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="CCE4FD04FF9FFFA32EEF5226FDCE2A75" box="[487,573,1294,1316]" count="11" pageId="3" pageNumber="48" type="male">11 males</specimenCount>
,
<specimenCount id="CCE4FD04FF9FFFA32D4E5226FD5D2A75" box="[582,686,1294,1316]" count="17" pageId="3" pageNumber="48" type="female">17 females</specimenCount>
(
<collectionCode id="BCF3AE48FF9FFFA32DB05226FCF62A75" box="[696,773,1294,1316]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="3" pageNumber="48">HNHM</collectionCode>
,
<specimenCount id="CCE4FD04FF9FFFA32C065226FCAB2A75" box="[782,856,1294,1316]" count="3" pageId="3" pageNumber="48" type="male">3 males</specimenCount>
,
<specimenCount id="CCE4FD04FF9FFFA32C695226FC4E2A75" box="[865,957,1294,1316]" count="4" pageId="3" pageNumber="48" type="female">4 females</specimenCount>
in alcohol): same as for
<typeStatus id="0559882FFF9FFFA32BAA5226FB052A75" box="[1186,1270,1294,1316]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
[genitalia of
<specimenCount id="CCE4FD04FF9FFFA32EBA5202FE082A11" box="[434,507,1322,1344]" count="2" pageId="3" pageNumber="48" type="male">2 males</specimenCount>
and females each in a plastic microvial with glycerol, wings ofa specimen pre - pared on a slide]
</materialsCitation>
.
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E7652A9FD702A81" blockId="3.[317,1270,1409,1805]" pageId="3" pageNumber="48">
Measurements in mm: body length 0.77 (
<typeStatus id="0559882FFF9FFFA32C1B52A9FC9F2AC6" box="[787,876,1409,1431]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
), 0.720.85 (
<typeStatus id="0559882FFF9FFFA32CE452A9FBB32AC6" box="[1004,1088,1409,1431]" pageId="3" pageNumber="48" type="paratype">paratype</typeStatus>
males), 0.750.95 (
<typeStatus id="0559882FFF9FFFA32E4B52B6FE642AE5" box="[323,407,1438,1460]" pageId="3" pageNumber="48" type="paratype">paratype</typeStatus>
females), wing length 0.68 (
<typeStatus id="0559882FFF9FFFA32DD152B6FCC12AE5" box="[729,818,1438,1460]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
), 0.650.75, 0.660.77, wing breadth 0.25 (
<typeStatus id="0559882FFF9FFFA32E4C5292FE6E2A81" box="[324,413,1466,1488]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
), 0.240.33, 0.260.34.
</paragraph>
<paragraph id="DA5D368DFF9FFFA32E7652FFFB352ABC" blockId="3.[317,1270,1409,1805]" box="[382,1222,1495,1517]" pageId="3" pageNumber="48">General colour dark to black; some parts ofbody with brownish grey microtomentum.</paragraph>
<paragraph id="DA5D368DFF9FFFA42E7652DCFC4D2D71" blockId="3.[317,1270,1409,1805]" lastBlockId="4.[317,1270,436,544]" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
Head much longer than high. Head with frons broad, frons mostly dark microtomentose. Anterior part covered with adpressed small scales; ifall the large central area offrons, which includes ocelli, back to occiput, is interpreted as frontal (ocellar) triangle, this triangle is bordered by a densely hatched stripe (
<figureCitation id="42D92A08FF9FFFA32EDD5162FDFC2931" box="[469,527,1610,1632]" captionStart="Figs 13" captionStartId="2.[317,360,1745,1767]" captionTargetBox="[357,1262,618,1706]" captionTargetId="figure-88@2.[317,1269,614,1711]" captionTargetPageId="2" captionText="Figs 13. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 12, 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12586910" httpUri="https://zenodo.org/record/12586910/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
). Face (prefrons) black, with extremely deep and strongly microtomentose antennal concavities. Interantennal tubercle small. Three pairs ofshort lateroclinate fronto-orbitals; a row ofmedioclinate inner orbitals. No ocellar pair, a number ofsmall postocellar setulae, postocellar setae indistinct. A pair oflong outer verticals. Presence or absence ofinner verticals difficult to inter - pret, as longer, paired setae between outer verticals absent. Eye much reduced, slightly oval. Scape and pedicel black. Pedicel with a
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long dorsal sub-basal seta (
<figureCitation id="42D92A08FF9FFFA32B6751F2FB4229A1" box="[1135,1201,1754,1776]" captionStart="Figs 13" captionStartId="2.[317,360,1745,1767]" captionTargetBox="[357,1262,618,1706]" captionTargetId="figure-88@2.[317,1269,614,1711]" captionTargetPageId="2" captionText="Figs 13. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 12, 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12586910" httpUri="https://zenodo.org/record/12586910/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
). First flagellomere rounded, ca.
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long with dorsal arista. First flagellomere with 5 extremely long dorsal setae, not much shorter than arista (longest
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mm
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), in addition with 34
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mm
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long apical and ventral setae and with shorter cilia. Arista
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mm
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long, i.e. a little more than 2 times as long as 1st flagellomere, practically bare. Vibrissa rather long, about as long as antenna. Gena narrow. 56 peristomals. No genal seta.
</paragraph>
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<paragraph id="DA5D368DFF98FFA42E3551FBFB062857" blockId="4.[317,1270,1747,1826]" pageId="4" pageNumber="49">
<emphasis id="E896EA9FFF98FFA42E3551FBFE6429B8" bold="true" box="[317,407,1747,1769]" pageId="4" pageNumber="49">Figs 48.</emphasis>
<taxonomicName id="1DE24D0EFF98FFA42E9351FBFD8129B8" authorityName="Papp" authorityYear="2004" box="[411,626,1747,1769]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF98FFA42E9351FBFD8129B8" box="[411,626,1747,1769]" italics="true" pageId="4" pageNumber="49">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF98FFA42D7E51FBFD5A29B8" box="[630,681,1747,1769]" pageId="4" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
4 = right wing, dorsal view, 58 = paratype male, legs in anterior view: 5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-
</paragraph>
</caption>
<paragraph id="DA5D368DFF98FFA42E955024FB652873" blockId="4.[317,1270,1747,1826]" box="[413,1174,1804,1826]" pageId="4" pageNumber="49">
nification. Scales:
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for
<figureCitation id="42D92A08FF98FFA42DC85024FCDF2873" box="[704,812,1804,1826]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="4" pageNumber="49">Figs 45, 8</figureCitation>
,
<quantity id="1D1A9B68FF98FFA42C3F5024FC732873" box="[823,896,1804,1826]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="49" unit="mm" value="0.1">0.1 mm</quantity>
for
<figureCitation id="42D92A08FF98FFA42CA15024FC102873" box="[937,995,1804,1826]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="4" pageNumber="49">Fig. 6</figureCitation>
,
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for
<figureCitation id="42D92A08FF98FFA42B565024FB652873" box="[1118,1174,1804,1826]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="4" pageNumber="49">Fig. 7</figureCitation>
</paragraph>
<caption id="8E9D6605FF99FFA52E355192FB052858" ID-DOI="http://doi.org/10.5281/zenodo.12586914" ID-Zenodo-Dep="12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="5" pageNumber="50" startId="5.[317,360,1722,1744]" targetBox="[346,1235,435,1687]" targetPageId="5" targetType="figure">
<paragraph id="DA5D368DFF99FFA52E355192FB052858" blockId="5.[317,1270,1722,1830]" pageId="5" pageNumber="50">
<emphasis id="E896EA9FFF99FFA52E355192FE562981" bold="true" box="[317,421,1722,1744]" pageId="5" pageNumber="50">Figs 915.</emphasis>
<taxonomicName id="1DE24D0EFF99FFA52EA65192FD7B2981" authorityName="Papp" authorityYear="2004" box="[430,648,1722,1744]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF99FFA52EA65192FD7B2981" box="[430,648,1722,1744]" italics="true" pageId="5" pageNumber="50">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF99FFA52D985192FD342981" box="[656,711,1722,1744]" pageId="5" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme,
</paragraph>
</caption>
<paragraph id="DA5D368DFF99FFA52E4F5038FB1F2877" blockId="5.[317,1270,1722,1830]" box="[327,1260,1808,1830]" pageId="5" pageNumber="50">
sublateral view, 15 = genital complex, lateral view. Scale:
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for
<figureCitation id="42D92A08FF99FFA52CDE5038FBFD2877" box="[982,1038,1808,1830]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="5" pageNumber="50">Fig. 9</figureCitation>
,
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for
<figureCitation id="42D92A08FF99FFA52B8A5038FB1F2877" box="[1154,1260,1808,1830]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="5" pageNumber="50">Figs 1015</figureCitation>
</paragraph>
<paragraph id="DA5D368DFF9AFFA62E76569CFB452D55" blockId="6.[317,1271,436,1063]" pageId="6" pageNumber="51">
Thorax black, dark microtomentose. Prothorax (
<figureCitation id="42D92A08FF9AFFA62C4F569CFC8C2E9B" box="[839,895,436,458]" captionStart="Figs 13" captionStartId="2.[317,360,1745,1767]" captionTargetBox="[357,1262,618,1706]" captionTargetId="figure-88@2.[317,1269,614,1711]" captionTargetPageId="2" captionText="Figs 13. Immoderatus foldvarii sp. n., paratype male. 1 = head, dorsal view, 2 = first flagellomere and apex ofpedicel, lateral view, 3 = prothorax, anterior view. Scales: 0.1 mm for Figs 12, 0.2 mm" figureDoi="http://doi.org/10.5281/zenodo.12586910" httpUri="https://zenodo.org/record/12586910/files/figure.png" pageId="6" pageNumber="51">Fig. 3</figureCitation>
) consisting ofextremely reduced scler - ites. Anepisternum present as a narrow oblique sclerite, medially only
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broad, without setae. Anepimeron membranous. Katepisternum with 1 comparatively long seta at dorsal margin.
</paragraph>
<paragraph id="DA5D368DFF9AFFA62E765522FB1E2DBB" blockId="6.[317,1271,436,1063]" pageId="6" pageNumber="51">
Legs short and thick, black, but trochanters, knees, apices oftibiae, and tarsi somewhat lighter. Fore femur thickened, with longer setae on dorsal side and sub-basally on ventral side. Mid femur with 1 anterior apical seta and with a number ofmedium-long setae. Hind femur very thick with a dorsal seta at apical 1/4. Fore tibia with 3 long dorsal setae: at apical 1/4, medially and subapically. Mid tibia without the usual subapical crown ofsetae, but with a strong ventroapical spur and with a shorter, though rather thick, anterodorsal spur (
<figureCitation id="42D92A08FF9AFFA62C0855B2FCC92DE1" box="[768,826,666,688]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 5</figureCitation>
). Hind tibia extremely thick, only 3 times longer than thick (
<figureCitation id="42D92A08FF9AFFA62EC7559FFDF92D9C" box="[463,522,695,717]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 7</figureCitation>
), with an extremely long and thick ventroapical spur, with a long (
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) dorsal seta opposite the base ofventroapical spur and with 3 pairs ofantero- and postero-dorsal setae.
</paragraph>
<paragraph id="DA5D368DFF9AFFA62E3555D9FE592CC6" blockId="6.[317,1271,436,1063]" pageId="6" pageNumber="51">
Fore basitarsus large and thick, about as long as tarsomeres 2 and 3 combined. Mid basitarsus (
<figureCitation id="42D92A08FF9AFFA62BBE55D9FB1C2C56" box="[1206,1263,753,775]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 6</figureCitation>
) with an extremely thick and long ventroapical thorn and 3 other ventral thorns. Mid tarsomere 2 and 3 each with a strong apical thorn. Hind basitarsus (
<figureCitation id="42D92A08FF9AFFA62C285402FCA92C11" box="[800,858,810,832]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 8</figureCitation>
) with three anterior (anteroventral) thorn. Hind tarsomeres 2 and 3 each with an apical (anterior to ventral) thorn, which is longer and setiform on tarsomere 3. All tarsomeres with a pair ofdorsal (i.e. antero- and postero-dorsal) long thin setae (
<figureCitation id="42D92A08FF9AFFA62E4C54A9FE6F2CC6" box="[324,412,897,919]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Figs 6, 8</figureCitation>
).
</paragraph>
<paragraph id="DA5D368DFF9AFFA62E7654B6FB582B76" blockId="6.[317,1271,436,1063]" pageId="6" pageNumber="51">
Wing peculiar (
<figureCitation id="42D92A08FF9AFFA62D1A54B6FDB92CE5" box="[530,586,926,948]" captionStart="Figs 48" captionStartId="4.[317,360,1747,1769]" captionTargetBox="[342,1247,578,1717]" captionTargetId="figure-74@4.[339,1247,576,1719]" captionTargetPageId="4" captionText="Figs 48. Immoderatus foldvarii sp. n. 4 = right wing, dorsal view, 58 = paratype male, legs in anterior view:5 = mid leg, 6 = mid tarsus, higher magnification, 7 = hind leg, 8 = hind tarsus, higher mag-" figureDoi="http://doi.org/10.5281/zenodo.12586912" httpUri="https://zenodo.org/record/12586912/files/figure.png" pageId="6" pageNumber="51">Fig. 4</figureCitation>
) both in form and venation. Wing membrane and veins unicolorous yellowish. R-M at distal part ofmid third ofdiscal cell. Costal vein with 2 long setae proximally to R
<subScript id="466634C8FF9AFFA62BE754EFFB052C82" attach="right" box="[1263,1270,967,979]" fontSize="5" pageId="6" pageNumber="51">1</subScript>
break, 13 to 15 long setae on 2nd and 3rd costal section. Wing vein ratios: C-sections 13 (
<typeStatus id="0559882FFF9AFFA62B9B54FFFB1F2CBC" box="[1171,1260,983,1005]" pageId="6" pageNumber="51" type="paratype">paratype</typeStatus>
): ca.
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,
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,
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, M vein index =
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/
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= 3.2, dM-Cu ca.
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, R-M-dM-Cu / dM-Cu ratio ca. 3.5. Halter dark grey, knob usually darker than stem.
</paragraph>
<caption id="8E9D6605FF9AFFA62E3551C7FB932873" ID-DOI="http://doi.org/10.5281/zenodo.12586916" ID-Zenodo-Dep="12586916" httpUri="https://zenodo.org/record/12586916/files/figure.png" pageId="6" pageNumber="51" startId="6.[317,360,1775,1797]" targetBox="[321,1265,1159,1725]" targetPageId="6" targetType="figure">
<paragraph id="DA5D368DFF9AFFA62E3551C7FB932873" blockId="6.[317,1270,1775,1826]" pageId="6" pageNumber="51">
<emphasis id="E896EA9FFF9AFFA62E3551C7FE5C2854" bold="true" box="[317,431,1775,1797]" pageId="6" pageNumber="51">Figs 1617.</emphasis>
<taxonomicName id="1DE24D0EFF9AFFA62EBE51C7FD7D2854" authorityName="Papp" authorityYear="2004" box="[438,654,1775,1797]" class="Insecta" family="Sphaeroceridae" genus="Immoderatus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="foldvarii" status="sp. nov.">
<emphasis id="E896EA9FFF9AFFA62EBE51C7FD7D2854" box="[438,654,1775,1797]" italics="true" pageId="6" pageNumber="51">Immoderatus foldvarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3A557E4FF9AFFA62D9B51C7FD3B2854" box="[659,712,1775,1797]" pageId="6" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
, paratype female, genitalia. 16 = apex of abdomen, dorsal view, with cercus in lateral view, 17 = spermathecae. Scale: 0.1 mm
</paragraph>
</caption>
<paragraph id="DA5D368DFF9BFFA72E76569CFC8B2D71" blockId="7.[317,1271,436,1092]" pageId="7" pageNumber="52">
Abdomen with well-sclerotized preabdominal sclerites. Terga black, broad, with fine setae also on lateral margins. Preabdominal sterna broad but less sclerotized, 5th sternum broad, unmodified. Postabdominal sclerites well-sclerotized, darker than preabdomen; 6th to 8th sterna strongly asymmetrical as in other copromyzine genera (
<figureCitation id="42D92A08FF9BFFA72C0F5522FC9F2D71" box="[775,876,522,544]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Figs 911</figureCitation>
).
</paragraph>
<paragraph id="DA5D368DFF9BFFA72E76550FFB8F2C75" blockId="7.[317,1271,436,1092]" pageId="7" pageNumber="52">
Male genital morphology peculiar, as described above. Epandrium strongly convex (globose), without ventral lateral cleft but fused and closed ventrally as in some genera of
<taxonomicName id="1DE24D0EFF9BFFA72B70556CFB002D0B" box="[1144,1267,580,602]" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="subFamily" subFamily="Limosininae">Limosininae</taxonomicName>
. Epandrium with a pair ofcomparatively long subdorsal (caudal) setae. Cerci distinct but small, not fused with epandrium (
<figureCitation id="42D92A08FF9BFFA72D115556FDAC2DC5" box="[537,607,638,660]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Fig. 10</figureCitation>
), without setae. Surstylus (
<figureCitation id="42D92A08FF9BFFA72C685556FC572DC5" box="[864,932,638,660]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Fig. 11</figureCitation>
) long thin, digitiform with a longer medial subapical seta and with 2 small apical setae. Hypandrium small. Phallic complex (
<figureCitation id="42D92A08FF9BFFA72BAF55B2FB1D2DE1" box="[1191,1262,666,688]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Fig. 15</figureCitation>
) rather compact. Phallus with rather normal distiphallus, basiphallus with a thick, intricately sclerotized dorsal projection, which is almost as large as distiphallus (
<figureCitation id="42D92A08FF9BFFA72CD755FCFBD52DBB" box="[991,1062,724,746]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Fig. 15</figureCitation>
). Phallapodeme slen- der, comparatively short, strongly curved in its distal third. Epiphallus seemingly absent. Ejaculatory apodeme (
<figureCitation id="42D92A08FF9BFFA72EAA5426FDE22C75" box="[418,529,782,804]" captionStart="Figs 915" captionStartId="5.[317,360,1722,1744]" captionTargetBox="[346,1235,435,1687]" captionTargetId="figure-11@5.[344,1243,424,1694]" captionTargetPageId="5" captionText="Figs 915. Immoderatus foldvarii sp. n., paratype male, genitalia. 9 = male postabdomen, dorsal view, 10 = same, ventral view, 11 = postabdominal sclerites, ventral view, 12 = surstylus in broadest extension, 13 = postgonite laterally, apical part in broadest extension, 14 = ejaculatory apodeme," figureDoi="http://doi.org/10.5281/zenodo.12586914" httpUri="https://zenodo.org/record/12586914/files/figure.png" pageId="7" pageNumber="52">Figs 1415</figureCitation>
) mushroom-shaped with an additional sclerite at its distal part.
</paragraph>
<paragraph id="DA5D368DFF9BFFA72E765402FC022CBC" blockId="7.[317,1271,436,1092]" pageId="7" pageNumber="52">
Female tergite 7 with a pair ofextremely long setae. Cercus with a long apical and 2 thick dor - sal setae (
<figureCitation id="42D92A08FF9BFFA72E91546FFE132C0C" box="[409,480,839,861]" captionStart="Figs 1617" captionStartId="6.[317,360,1775,1797]" captionTargetBox="[321,1265,1159,1725]" captionTargetId="figure-362@6.[317,1269,1154,1726]" captionTargetPageId="6" captionText="Figs 1617. Immoderatus foldvarii sp. n., paratype female, genitalia. 16 = apex of abdomen, dorsal view, with cercus in lateral view, 17 = spermathecae. Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.12586916" httpUri="https://zenodo.org/record/12586916/files/figure.png" pageId="7" pageNumber="52">Fig. 16</figureCitation>
). Two spermathecae (
<figureCitation id="42D92A08FF9BFFA72DA7546FFD062C0C" box="[687,757,839,861]" captionStart="Figs 1617" captionStartId="6.[317,360,1775,1797]" captionTargetBox="[321,1265,1159,1725]" captionTargetId="figure-362@6.[317,1269,1154,1726]" captionTargetPageId="6" captionText="Figs 1617. Immoderatus foldvarii sp. n., paratype female, genitalia. 16 = apex of abdomen, dorsal view, with cercus in lateral view, 17 = spermathecae. Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.12586916" httpUri="https://zenodo.org/record/12586916/files/figure.png" pageId="7" pageNumber="52">Fig. 17</figureCitation>
), similar in shape but uneven in size (
<specimenCount id="CCE4FD04FF9BFFA72B50546FFB3E2C0C" box="[1112,1229,839,861]" count="2" pageId="7" pageNumber="52" type="female">two females</specimenCount>
dissected), spermathecae not globular as in several genera ofCopromyzinae (
<taxonomicName id="1DE24D0EFF9BFFA72B1C544CFB822C2B" authorityName="Lioy" authorityYear="1864" box="[1044,1137,868,890]" class="Insecta" family="Sphaeroceridae" genus="Lotophila" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72B1C544CFB822C2B" box="[1044,1137,868,890]" italics="true" pageId="7" pageNumber="52">Lotophila</emphasis>
</taxonomicName>
,
<taxonomicName id="1DE24D0EFF9BFFA72B76544CFB1C2C2B" authorityName="Macquart" authorityYear="1835" box="[1150,1263,868,890]" class="Insecta" family="Sphaeroceridae" genus="Crumomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72B76544CFB1C2C2B" box="[1150,1263,868,890]" italics="true" pageId="7" pageNumber="52">Crumomyia</emphasis>
</taxonomicName>
,
<taxonomicName id="1DE24D0EFF9BFFA72E3554A9FE392CC6" box="[317,458,897,919]" class="Insecta" family="Sphaeroceridae" genus="Achaetothorax" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72E3554A9FE392CC6" box="[317,458,897,919]" italics="true" pageId="7" pageNumber="52">Achaetothorax</emphasis>
</taxonomicName>
,
<taxonomicName id="1DE24D0EFF9BFFA72ED454A9FD882CC6" authorityName="Hedicke" authorityYear="1923" box="[476,635,897,919]" class="Insecta" family="Sphaeroceridae" genus="Gymnometopina" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72ED454A9FD882CC6" box="[476,635,897,919]" italics="true" pageId="7" pageNumber="52">Gymnometopina</emphasis>
</taxonomicName>
,
<taxonomicName id="1DE24D0EFF9BFFA72D8554A9FCE52CC6" authorityName="Vanschuytbroeck" authorityYear="1948" box="[653,790,897,919]" class="Insecta" family="Sphaeroceridae" genus="Metaborborus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72D8554A9FCE52CC6" box="[653,790,897,919]" italics="true" pageId="7" pageNumber="52">Metaborborus</emphasis>
</taxonomicName>
, etc., see
<bibRefCitation id="BE734B7CFF9BFFA72C8F54A9FB772CC6" author="NORRBOM, A. L. &amp; KIM, K. C." box="[903,1156,897,919]" pageId="7" pageNumber="52" pagination="167 - 225" refId="ref3210" refString="NORRBOM, A. L. &amp; KIM, K. C. (1985 a) Systematics ofCrumomyia Macquart and Alloborborus (Diptera: Sphaeroceridae). Systematic Entomology 10: 167 - 225." type="journal article" year="1985">
<smallCapsWord id="DCBBA051FF9BFFA72C8F54A9FC1A2CC4" baselines="913,913" box="[903,1001,897,919]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Norrbom" pageId="7" pageNumber="52">NORRBOM</smallCapsWord>
&amp;
<smallCapsWord id="DCBBA051FF9BFFA72B1B54A9FBCF2CC4" baselines="913,913" box="[1043,1084,897,919]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Kim" pageId="7" pageNumber="52">KIM</smallCapsWord>
1985
<emphasis id="E896EA9FFF9BFFA72B7F54A9FB772CC6" box="[1143,1156,897,919]" italics="true" pageId="7" pageNumber="52">a</emphasis>
</bibRefCitation>
,b,
<bibRefCitation id="BE734B7CFF9BFFA72BAF54A9FE282CE5" author="PAPP, L. &amp; NORRBOM, A. L." pageId="7" pageNumber="52" pagination="321 - 344" refId="ref3314" refString="PAPP, L. &amp; NORRBOM, A. L. (1992) Taxonomy ofthe Palaeotropical genus Achaetothorax Hedicke (Diptera, Sphaeroceridae). Acta zool. hung. 38: 321 - 344." type="journal article" year="1992">
<smallCapsWord id="DCBBA051FF9BFFA72BAF54A9FB242CC4" baselines="913,913" box="[1191,1239,897,919]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Papp" pageId="7" pageNumber="52">PAPP</smallCapsWord>
&amp;
<smallCapsWord id="DCBBA051FF9BFFA72E3554B6FE6C2CE3" baselines="942,942" box="[317,415,926,948]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Norrbom" pageId="7" pageNumber="52">NORRBOM</smallCapsWord>
1992
</bibRefCitation>
). Ducts laterally directed, which again is peculiar (similar lateroclinate ducts are known in
<taxonomicName id="1DE24D0EFF9BFFA72EAD5492FC952C81" authority="ROHACEK &amp; MARSHALL, 1998" authorityName="ROHACEK &amp; MARSHALL" authorityYear="1998" box="[421,870,954,976]" class="Insecta" family="Sphaeroceridae" genus="Sphaeromitra" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="E896EA9FFF9BFFA72EAD5492FDDA2C81" box="[421,553,954,976]" italics="true" pageId="7" pageNumber="52">Sphaeromitra</emphasis>
<bibRefCitation id="BE734B7CFF9BFFA72D3B5492FC952C81" author="ROHACEK, J. &amp; MARSHALL, S. A." box="[563,870,954,976]" pageId="7" pageNumber="52" pagination="455 - 491" refId="ref3406" refString="ROHACEK, J. &amp; MARSHALL, S. A. (1998) Revision ofHomalomitrinae subfam. n. (Diptera: Sphaero - ceridae), with description ofa new genus and three new species. European Journal ofEntomol - ogy 95: 455 - 491." type="journal article" year="1998">
<smallCapsWord id="DCBBA051FF9BFFA72D3B5492FD622C9E" baselines="970,971" box="[563,657,954,976]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Rohacek" pageId="7" pageNumber="52">ROHÁČEK</smallCapsWord>
&amp;
<smallCapsWord id="DCBBA051FF9BFFA72DB05492FCD02C9E" baselines="970,971" box="[696,803,954,976]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Marshall" pageId="7" pageNumber="52">MARSHALL</smallCapsWord>
, 1998
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="1DE24D0EFF9BFFA72C7C5492FBE62C81" box="[884,1045,954,976]" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="subFamily" subFamily="Homalomitrinae">Homalomitrinae</taxonomicName>
), but the shape ofthe spermatheca ofthis species is otherwise unique amongst sphaerocerids.
</paragraph>
<paragraph id="DA5D368DFF9BFFA72E7654DCFCA62B15" blockId="7.[317,1271,436,1092]" pageId="7" pageNumber="52">This new species is so peculiar that I think a new genus deserves to be erected for it. However, I have no objective criteria to separate generic or specific characters. For this reason, some characters are given in both the generic and specific descriptions.</paragraph>
<paragraph id="DA5D368DFF9BFFA72E76534FFCCC2BE6" blockId="7.[317,1265,1127,1207]" pageId="7" pageNumber="52">
<materialsCitation id="6A8A3CD0FF9BFFA72E76534FFCC92BE6" country="Vietnam" location="Viet Nam" pageId="7" pageNumber="52" specimenCount="1">
I name this new species to honour my young colleague, Dr
<smallCapsWord id="DCBBA051FF9BFFA72CA5534FFC0F2B2D" baselines="1143,1144" box="[941,1020,1127,1149]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Mihaly" pageId="7" pageNumber="52">MIHÁLY</smallCapsWord>
<smallCapsWord id="DCBBA051FF9BFFA72B09534FFB902B2D" baselines="1143,1144" box="[1025,1123,1127,1149]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Foldvari" pageId="7" pageNumber="52">FÖLDVÁRI</smallCapsWord>
, one ofthe collectors ofthe
<typeStatus id="0559882FFF9BFFA72EB053ACFE122BCB" box="[440,481,1156,1178]" pageId="7" pageNumber="52">type</typeStatus>
series, and who also collected much other invaluable dipterous material during his joint 2003 collection trip to
<collectingCountry id="A2F5761DFF9BFFA72D595389FD422BE6" box="[593,689,1185,1207]" name="Vietnam" pageId="7" pageNumber="52">Viet Nam</collectingCountry>
and
<collectingCountry id="A2F5761DFF9BFFA72DEA5389FCC92BE6" box="[738,826,1185,1207]" name="Thailand" pageId="7" pageNumber="52">Thailand</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="DA5D368DFF9BFFA72C1B53C1FCEC2BAE" blockId="7.[787,799,1257,1279]" box="[787,799,1257,1279]" pageId="7" pageNumber="52">*</paragraph>
<paragraph id="DA5D368DFF9BFFA72E76523CFB052A7B" blockId="7.[317,1270,1300,1351]" box="[382,1270,1300,1322]" pageId="7" pageNumber="52">
<emphasis id="E896EA9FFF9BFFA72E76523CFDDC2A7B" box="[382,559,1300,1322]" italics="true" pageId="7" pageNumber="52">Acknowledgements</emphasis>
This study was supported by the National Scientific Research Fund (OTKA,
</paragraph>
<paragraph id="DA5D368DFF9BFFA72E355219FC682A16" blockId="7.[317,1270,1300,1351]" box="[317,923,1329,1351]" pageId="7" pageNumber="52">No. T 042540). I am grateful to the two reviewers for advice.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,74 +1,74 @@
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9 L2, 14 L3,
<date value="2018-05-17">17.05.2018</date>
, KO, neig. v. Abay, PF Akzhol, rice field edge, IT; 8 L1, 11 L2, 5 L3, 4 L4, 2 L5, 24-
<date value="2018-05-26">26.05.2018</date>
<date id="197D759F99DAF7636D8B561E7EBE492B" value="2018-05-17">17.05.2018</date>
, KO, neig. v. Abay, PF Akzhol, rice field edge, IT; 8 L1, 11 L2, 5 L3, 4 L4, 2 L5, 24-
<date id="979A9FFAA66C0EF68298B8471F46B9EE" value="2018-05-26">26.05.2018</date>
, KO, neig. v. Abay, PF Akzhol, rice field edge, IT;
<specimenCount type="male">12 males</specimenCount>
<specimenCount id="2BD9C1D4D370088E5DADF4443EB90687" type="male">12 males</specimenCount>
,
<specimenCount type="female">11 females</specimenCount>
<specimenCount id="671804FFA3FAD9C36779549640BFCFB5" type="female">11 females</specimenCount>
,
<date value="2019-06-24">24.06.2019</date>
<date id="B70BBF50822FE9781E890B472B78FEB6" value="2019-06-24">24.06.2019</date>
, KO, Shieli d., PF Akmaya, rice field edge and alfalfa fields, IT,
<normalizedToken originalValue="АМ">AM</normalizedToken>
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.
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