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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.666.9191" ID-PMC="PMC5534527" ID-Pensoft-Pub="1313-2970-666-1" ID-Pensoft-UUID="C43EEB41A94B930FFE439D1FAD29FF9C" ID-PubMed="28769631" ID-Zenodo-Dep="579453" ID-ZooBank="B3C377E8BBB14F328AECA2C22D1E625A" ModsDocID="1313-2970-666-1" checkinTime="1491551283800" checkinUser="pensoft" docAuthor="Moore, Matthew R., Jameson, Mary L., Garner, Beulah H., Audibert, Cedric, Smith, Andrew B. T. &amp; Seidel, Matthias" docDate="2017" docId="1E0602D4C7AB56649A1DF2AF6F2DF2C4" docLanguage="en" docName="ZooKeys 666: 1-349" docOrigin="ZooKeys 666" docPubDate="2017-04-06" docSource="http://dx.doi.org/10.3897/zookeys.666.9191" docTitle="Pelidnota unicolor subsp. unicolor unicolor (Drury 1782" docType="treatment" docVersion="3" id="C43EEB41A94B930FFE439D1FAD29FF9C" lastPageNumber="1" masterDocId="C43EEB41A94B930FFE439D1FAD29FF9C" masterDocTitle="Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies" masterLastPageNumber="349" masterPageNumber="1" pageNumber="1" updateTime="1668164364999" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moore, Matthew R.</mods:namePart>
<mods:affiliation>Department of Entomology and Nematology, University of Florida Building 1881 Natural Drive Area, Steinmetz Hall, Box 110620, Gainesville, FL 32611 - 0620, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jameson, Mary L.</mods:namePart>
<mods:affiliation>Department of Biological Sciences, Wichita State University 1845 Fairmount, Box 26, Wichita, KS 67260 - 0026, USA</mods:affiliation>
<mods:nameIdentifier type="email">maryliz.jameson@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garner, Beulah H.</mods:namePart>
<mods:affiliation>Natural History Museum, Insects Division, Department of Life Sciences, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Audibert, Cedric</mods:namePart>
<mods:affiliation>Musee des Confluences, Centre de Conservation et d'Etude des Collections, 13 A Rue Bancel, F- 69007 Lyon, France</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Andrew B. T.</mods:namePart>
<mods:affiliation>Research Division, Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-04-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>666</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>349</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.666.9191</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.666.9191</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-666-1</mods:identifier>
<mods:identifier type="ZooBank">B3C377E8BBB14F328AECA2C22D1E625A</mods:identifier>
<mods:identifier type="Pensoft-UUID">C43EEB41A94B930FFE439D1FAD29FF9C</mods:identifier>
<mods:identifier type="Zenodo-Dep">579453</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182230972" LSID="urn:lsid:plazi:treatment:1E0602D4C7AB56649A1DF2AF6F2DF2C4" httpUri="http://treatment.plazi.org/id/1E0602D4C7AB56649A1DF2AF6F2DF2C4" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="1E0602D4-C7AB-5664-9A1D-F2AF6F2DF2C4" authority="unicolor (Drury, 1782)" authorityName="unicolor (Drury" authorityYear="1782" baseAuthorityName="Drury" baseAuthorityYear="1782" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor subsp. unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="unicolor" subSpecies="unicolor">Pelidnota unicolor unicolor (Drury, 1782)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Scarabaeidae" genus="Scarabeus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Scarabeus unicolor" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="unicolor">Scarabeus unicolor</taxonomicName>
Drury, 1782: 61 [original combination].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Drury" baseAuthorityYear="1782" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="unicolor">Pelidnota unicolor</taxonomicName>
(Drury) [new combination by
<bibRefCitation author="Blanchard, CE" editor="Milne-Edwards, H" journalOrPublisher="Catalogue de la Collection Entomologique. Classe des Insectes. Volume 1. Part 1. Gide and Baudry, Paris" pageId="0" pageNumber="1" pagination="129 - 240" refId="B17" refString="Blanchard, CE, 1851. Ordre des Coleoptera. In: Milne-Edwards, H, Blanchard, CE, Lucas, H, Eds., Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Volume 1. Part 1. Gide and Baudry, Paris: 129 - 240" title="Ordre des Coleoptera." volumeTitle="Museum d'Histoire Naturelle de Paris." year="1851">Blanchard 1851</bibRefCitation>
: 211]
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Drury" baseAuthorityYear="1782" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota (Pelidnota) unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="unicolor" subGenus="Pelidnota">Pelidnota (Pelidnota) unicolor</taxonomicName>
(Drury) [new subgeneric combination by
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="1" pagination="1 - 241" refId="B278" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20: 1 - 241" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae." volume="20" year="1918">Ohaus 1918</bibRefCitation>
: 24]
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Drury" baseAuthorityYear="1782" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="unicolor">Pelidnota unicolor</taxonomicName>
(Drury) [removal of subgeneric combination by
<bibRefCitation author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula 2009</bibRefCitation>
: 91-93]
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Herbst" authorityYear="1790" class="Insecta" family="Scarabaeidae" genus="Melolontha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melolontha druryana" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="druryana">Melolontha druryana</taxonomicName>
Herbst, 1790
<emphasis bold="true" pageId="0" pageNumber="1">synonym.</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Herbst" authorityYear="1790" class="Insecta" family="Scarabaeidae" genus="Melolontha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melolontha druryana" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="druryana">Melolontha druryana</taxonomicName>
Herbst, 1790: 163 [original combination].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Drury" baseAuthorityYear="1782" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="unicolor">Pelidnota unicolor</taxonomicName>
(Drury) [syn. by
<bibRefCitation author="Blanchard, CE" editor="Milne-Edwards, H" journalOrPublisher="Catalogue de la Collection Entomologique. Classe des Insectes. Volume 1. Part 1. Gide and Baudry, Paris" pageId="0" pageNumber="1" pagination="129 - 240" refId="B17" refString="Blanchard, CE, 1851. Ordre des Coleoptera. In: Milne-Edwards, H, Blanchard, CE, Lucas, H, Eds., Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Volume 1. Part 1. Gide and Baudry, Paris: 129 - 240" title="Ordre des Coleoptera." volumeTitle="Museum d'Histoire Naturelle de Paris." year="1851">Blanchard 1851</bibRefCitation>
: 211].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Castelnau" authorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota testacea" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="testacea">Pelidnota testacea</taxonomicName>
Laporte, 1840
<emphasis bold="true" pageId="0" pageNumber="1">synonym.</emphasis>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Castelnau" authorityYear="1840" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota testacea" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="testacea">Pelidnota testacea</taxonomicName>
Laporte, 1840: 122 [original combination].
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota druryana" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="druryana">Pelidnota druryana</taxonomicName>
(Drury) [syn. by
<bibRefCitation author="Burmeister, HCC" journalOrPublisher="Enslin, Berlin" pageId="0" pageNumber="1" refId="B24" refString="Burmeister, HCC, 1844. Handbuch der Entomologie (ColeopteraLamellicornia Anthobia et Phyllophaga Systellochela). Volume 4, Part 1. T. C. F. Enslin, Berlin" title="Handbuch der Entomologie (ColeopteraLamellicornia Anthobia et Phyllophaga Systellochela). Volume 4, Part 1. T. C. F." year="1844">Burmeister 1844</bibRefCitation>
: 403].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
BRAZIL:
<normalizedToken originalValue="Espírito">Espirito</normalizedToken>
Santo, Minas Gerais, Pernambuco, Rio de Janeiro,
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo, Santa Catarina (
<bibRefCitation author="Herbst, JFW" journalOrPublisher="Pauli, Berlin" pageId="0" pageNumber="1" refId="B124" refString="Herbst, JFW, 1790. Natursystem aller bekannten in- und auslaendlischen Insekten, als eine Fortsetzung der von Bueffonschen Naturgeschichte. Der Kaefer dritter Theil. Pauli, Berlin" title="Natursystem aller bekannten in- und auslaendlischen Insekten, als eine Fortsetzung der von Bueffonschen Naturgeschichte. Der Kaefer dritter Theil." year="1790">Herbst 1790</bibRefCitation>
;
<bibRefCitation author="Laporte, [= Castelnau] FLNCde" journalOrPublisher="Dumenil, Paris" pageId="0" pageNumber="1" refId="B163" refString="Laporte, [=Castelnau] FLNCde, 1840. Histoire naturelle des insectes Coleopteres; avec une introduction renfermant l'anatomie et la physiologie des animaux articules, par M. Brulle. Tome premier. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes Tome troisieme. P. Dumenil, Paris" title="Histoire naturelle des insectes Coleopteres; avec une introduction renfermant l'anatomie et la physiologie des animaux articules, par M. Brulle. Tome premier. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes Tome troisieme. P." year="1840">Laporte 1840</bibRefCitation>
;
<bibRefCitation author="Burmeister, HCC" journalOrPublisher="Enslin, Berlin" pageId="0" pageNumber="1" refId="B24" refString="Burmeister, HCC, 1844. Handbuch der Entomologie (ColeopteraLamellicornia Anthobia et Phyllophaga Systellochela). Volume 4, Part 1. T. C. F. Enslin, Berlin" title="Handbuch der Entomologie (ColeopteraLamellicornia Anthobia et Phyllophaga Systellochela). Volume 4, Part 1. T. C. F." year="1844">Burmeister 1844</bibRefCitation>
;
<bibRefCitation author="Blanchard, CE" editor="Milne-Edwards, H" journalOrPublisher="Catalogue de la Collection Entomologique. Classe des Insectes. Volume 1. Part 1. Gide and Baudry, Paris" pageId="0" pageNumber="1" pagination="129 - 240" refId="B17" refString="Blanchard, CE, 1851. Ordre des Coleoptera. In: Milne-Edwards, H, Blanchard, CE, Lucas, H, Eds., Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Volume 1. Part 1. Gide and Baudry, Paris: 129 - 240" title="Ordre des Coleoptera." volumeTitle="Museum d'Histoire Naturelle de Paris." year="1851">Blanchard 1851</bibRefCitation>
;
<bibRefCitation author="Harold, E" editor="Gemminger, M" journalOrPublisher="4. E. H. Gummi, Munich" pageId="0" pageNumber="1" pagination="979 - 1346" refId="B112" refString="Harold, E, 1869b. Scarabaeidae. In: Gemminger, M, Harold, E, Eds., Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus, Vol. 4. E. H. Gummi, Munich: 979 - 1346" title="Scarabaeidae." volumeTitle="Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus, Vol." year="1869 b">Harold 1869b</bibRefCitation>
;
<bibRefCitation author="Ohaus, F" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" pagination="239 - 262" refId="B267" refString="Ohaus, F, 1908a. Die Ruteliden meiner Sammelreisen in Suedamerika (Col.). Deutsche Entomologische Zeitschrift 1908: 239 - 262" title="Die Ruteliden meiner Sammelreisen in Suedamerika (Col.)." volume="1908" year="1908 a">Ohaus 1908a</bibRefCitation>
,
<bibRefCitation author="Ohaus, F" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" pagination="487 - 511" refId="B274" refString="Ohaus, F, 1913. XI. Beitrag zur Kenntnis der Ruteliden. (Col.). Deutsche Entomologische Zeitschrift 1913: 487 - 511" title="XI. Beitrag zur Kenntnis der Ruteliden. (Col.)." volume="1913" year="1913">1913</bibRefCitation>
,
<bibRefCitation author="Ohaus, F" journalOrPublisher="Coleopterorum Catalogus" pageId="0" pageNumber="1" pagination="1 - 241" refId="B278" refString="Ohaus, F, 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. Coleopterorum Catalogus 20: 1 - 241" title="Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae." volume="20" year="1918">1918</bibRefCitation>
,
<bibRefCitation author="Ohaus, F" journalOrPublisher="Mitteilungen der Deutschen Entomologischen Gesellschaft" pageId="0" pageNumber="1" refId="B286" refString="Ohaus, F, 1934b. Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T.1: Tribus Rutelini. Genera Insectorum Fasc. 199A: 1-172." title="Coleopteralamellicornia, Fam. Scarabaeidae, Subfam. Rutelinae. T. 1: Tribus Rutelini. Genera Insectorum Fasc. 199 A: 1 - 172." year="1934 b">1934b</bibRefCitation>
;
<bibRefCitation author="Guimaraes, LR" journalOrPublisher="Papeis Avulsos do Departamento de Zoologia (Sao Paulo)" pageId="0" pageNumber="1" pagination="93 - 102" refId="B101" refString="Guimaraes, LR, 1944. Rutelidae, Cetonidae, Melolonthidae e Dynastidae de Monte Alegre. Papeis Avulsos do Departamento de Zoologia (Sao Paulo) 6: 93 - 102" title="Rutelidae, Cetonidae, Melolonthidae e Dynastidae de Monte Alegre." volume="6" year="1944">
<normalizedToken originalValue="Guimarães">Guimaraes</normalizedToken>
1944
</bibRefCitation>
;
<bibRefCitation author="Machatschke, JW" journalOrPublisher="Coleopterorum Catalogus Supplementa" pageId="0" pageNumber="1" pagination="1 - 361" refId="B188" refString="Machatschke, JW, 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361" title="Scarabaeoidea: Melolonthidae, Rutelinae." volume="66" year="1972">Machatschke 1972</bibRefCitation>
;
<bibRefCitation author="Krajcik, M" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="0" pageNumber="1" refId="B155" refString="Krajcik, M, 2008. Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Second Edition. Plzen, 142 pp." title="Animma. X. Supplement 4. Checklist of the Scarabaeoidea of the World. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Second Edition. Plzen, 142 pp." year="2008">Krajcik 2008</bibRefCitation>
;
<bibRefCitation author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula 2009</bibRefCitation>
). PERU (
<bibRefCitation DOI="https://doi.org/10.2317/kent-88-02-186-207.1" author="Ratcliffe, BC" journalOrPublisher="The Journal of the Kansas Entomological Society" pageId="0" pageNumber="1" pagination="186 - 207" refId="B321" refString="Ratcliffe, BC, Jameson, ML, Figueroa, L, Cave, RD, Paulsen, MJ, Cano, EB, Beza-Beza, C, Jimenez-Ferbans, L, Reyes-Castillo, R, 2015. Beetles (Coleoptera) of Peru: A survey of families. Scarabaeoidea. The Journal of the Kansas Entomological Society 88: 186 - 207, DOI: https://doi.org/10.2317/kent-88-02-186-207.1" title="Beetles (Coleoptera) of Peru: A survey of families. Scarabaeoidea." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="types">
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
<paragraph pageId="0" pageNumber="1">
The following specimen is at CCECL. 1 invalid ♂ neotype: &quot;Salesopolis
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo
<normalizedToken originalValue="Brésil">Bresil</normalizedToken>
M. SOULA det 19 [obverse] II/2000//
<taxonomicName baseAuthorityName="Drury" baseAuthorityYear="1782" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis italics="true" pageId="0" pageNumber="1">Pelidnota unicolor</emphasis>
</taxonomicName>
(Dr.) M. SOULA det 2008//
<normalizedToken originalValue="Néotype">Neotype</normalizedToken>
2008
<taxonomicName family="Scarabaeidae" genus="Scarabeus" kingdom="Animalia" lsidName="Scarabeus unicolor" order="Coleoptera" pageId="0" pageNumber="1" rank="species" species="unicolor">
<emphasis italics="true" pageId="0" pageNumber="1">Scarabeus unicolor</emphasis>
</taxonomicName>
Dr Soula det.&quot; (47030861). Genitalia card-mounted underneath the invalid neotype. Box 4618683 SOULA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula (2009</bibRefCitation>
: 92) designated a neotype specimen for
<taxonomicName authority="unicolor" authorityName="unicolor (Drury" authorityYear="1782" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor subsp. unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="unicolor" subSpecies="unicolor">
<emphasis italics="true" pageId="0" pageNumber="1">Pelidnota unicolor unicolor</emphasis>
</taxonomicName>
.
<bibRefCitation author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula (2009)</bibRefCitation>
did not state where this neotype was deposited. Article 75.3.7 (ICZN 1999) requires a statement that the &quot;neotype is, or immediately upon publication has become, the property of a recognized scientific or educational institution, cited by name, that maintains a research collection, with proper facilities for preserving name-bearing types, and that makes them accessible for study&quot;. We recovered
<normalizedToken originalValue="Soula’s">Soula's</normalizedToken>
invalid neotype specimen in his formerly private collection (now at CCECL).
<bibRefCitation author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">
<normalizedToken originalValue="Soula´s">Soula's</normalizedToken>
(2009)
</bibRefCitation>
neotype is invalid because
<normalizedToken originalValue="Soula’s">Soula's</normalizedToken>
collection was private and
<bibRefCitation author="Soula, M" journalOrPublisher="AECFT, Saintry" pageId="0" pageNumber="1" refId="B360" refString="Soula, M, 2009. Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot;Pelidnotina&quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale. AECFT, Saintry" title="Les Coleopteres du Nouveau Monde. Volume 3: Rutelini 3. Revision des Pelidnotina 3. Photos de toutes les especes de Lagochile. Une revision des genres Pelidnota, Ectinoplectron, Pseudogeniates, Xenopelidnota (Coleoptera: Scarabaeidae: Rutelinae: Rutelini: &quot; Pelidnotina &quot;). Besoiro: Supplement au Bulletin de liaison de l'Association Entomologique pour la Connaissance de la Faune Tropicale." year="2009">Soula (2009)</bibRefCitation>
did not make a statement of neotype deposition.
</paragraph>
<paragraph pageId="0" pageNumber="1">
We treat
<taxonomicName authorityName="Ohaus" authorityYear="1913" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor subsp. var. var. infuscata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="variety" species="unicolor" subSpecies="var." variety="infuscata">Pelidnota unicolor var. infuscata</taxonomicName>
Ohaus (Fig.
<figureCitation captionStart="Figure 94" captionStartId="F94" captionText="Figure 94. Pelidnota unicolor var. infuscata Ohaus (unavailable name) (valid name Pelidnota unicolor [Drury]) invalid type female from ZMHB. A Dorsal habitus B Lateral habitus C Specimen labels." figureDoi="10.3897/zookeys.666.9191.figure94" httpUri="https://binary.pensoft.net/fig/130522" pageId="0" pageNumber="1">94</figureCitation>
) as unambiguously infrasubspecific and thus as an
<emphasis bold="true" pageId="0" pageNumber="1">unavailable name</emphasis>
.
<bibRefCitation author="Ohaus, F" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" pagination="487 - 511" refId="B274" refString="Ohaus, F, 1913. XI. Beitrag zur Kenntnis der Ruteliden. (Col.). Deutsche Entomologische Zeitschrift 1913: 487 - 511" title="XI. Beitrag zur Kenntnis der Ruteliden. (Col.)." volume="1913" year="1913">Ohaus (1913)</bibRefCitation>
clearly described this taxon as a variety. For the purpose of Art. 45.6.4. ICZN,
<bibRefCitation author="Ohaus, F" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" pagination="487 - 511" refId="B274" refString="Ohaus, F, 1913. XI. Beitrag zur Kenntnis der Ruteliden. (Col.). Deutsche Entomologische Zeitschrift 1913: 487 - 511" title="XI. Beitrag zur Kenntnis der Ruteliden. (Col.)." volume="1913" year="1913">
<normalizedToken originalValue="Ohaus’s">Ohaus's</normalizedToken>
(1913)
</bibRefCitation>
publication described both subspecies and varieties (sometimes both for the same species, e.g.,
<taxonomicName authorityName="Blanchard" authorityYear="1850" class="Insecta" family="Scarabaeidae" genus="Homonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homonyx chalceus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chalceus">
<emphasis italics="true" pageId="0" pageNumber="1">Homonyx chalceus</emphasis>
</taxonomicName>
), thus unambiguously allowing us to treat this name in an infrasubspecific manner.
<bibRefCitation author="Ohaus, F" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="1" pagination="487 - 511" refId="B274" refString="Ohaus, F, 1913. XI. Beitrag zur Kenntnis der Ruteliden. (Col.). Deutsche Entomologische Zeitschrift 1913: 487 - 511" title="XI. Beitrag zur Kenntnis der Ruteliden. (Col.)." volume="1913" year="1913">Ohaus (1913)</bibRefCitation>
clearly and unambiguously described &quot;var.
<taxonomicName class="Insecta" family="Scarabaeidae" genus="Homonyx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="var. infuscata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="variety" species="chalceus" variety="infuscata">infuscata</taxonomicName>
&quot; as an infrasubspecific name under
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis italics="true" pageId="0" pageNumber="1">P. unicolor</emphasis>
</taxonomicName>
. As such, this name is not treated as a valid species group name. Ohaus (1908) made reference to this dark variety and subsequently named it as a variety (
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="infuscata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis italics="true" pageId="0" pageNumber="1">infuscata</emphasis>
</taxonomicName>
) in 1913. The specimen in ZMHB labeled as a type is, in fact, a dark color morph of
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis italics="true" pageId="0" pageNumber="1">P. unicolor</emphasis>
</taxonomicName>
based on comparisons with the nominotypical form. This specimen was labeled as an invalid type in ZMHB.
</paragraph>
<caption ID-Zenodo-Dep="982497" doi="10.3897/zookeys.666.9191.figure94" httpUri="https://binary.pensoft.net/fig/130522" pageId="0" pageNumber="1" start="Figure 94" startId="F94">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 94.</emphasis>
<taxonomicName authorityName="Ohaus" authorityYear="1913" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor subsp. var. var. infuscata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="variety" species="unicolor" subSpecies="var." variety="infuscata">Pelidnota unicolor var. infuscata</taxonomicName>
Ohaus (unavailable name) (valid name
<taxonomicName baseAuthorityName="Drury" baseAuthorityYear="1782" class="Insecta" family="Scarabaeidae" genus="Pelidnota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelidnota unicolor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis italics="true" pageId="0" pageNumber="1">Pelidnota unicolor</emphasis>
</taxonomicName>
[Drury]) invalid type female from ZMHB.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Dorsal habitus
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Lateral habitus
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
Specimen labels.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>