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<document ID-CLB-Dataset="291551" ID-GBIF-Dataset="8011f729-cae2-44a9-8412-ae9c7345c970" ID-ISBN="978-3-258-08047-5" checkinTime="1706569699811" checkinUser="plazi" docAuthor="Konrad Lauber, Gerhart Wagner &amp; Andreas Gygax" docDate="2018" docId="1E0365388F7A406D0AE2240AD161B581" docLanguage="de" docName="FloraHelv.Asteraceae.xml" docOrigin="Flora Helvetica, Bern: Haupt Verlag" docTitle="Artemisia dracunculus L." docType="treatment" docVersion="7" id="8ABA94EA572036683CE015D8D3406DA0" lastPageNumber="1116" masterDocId="43055F1E0D42AF44E8DC23E024F52E45" masterDocTitle="Flora Helvetica - Asteraceae" masterLastPageNumber="1250" masterPageNumber="1074" pageNumber="1116" updateTime="1711157813649" updateUser="ExternalLinkService">
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<mods:title id="3E486A9FD362212EB736E5D37452379C">Flora Helvetica - Asteraceae</mods:title>
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<mods:namePart id="73D3F910725BFC3C8E31DD20F7D8B4FE">Konrad Lauber</mods:namePart>
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<mods:namePart id="4ABB5F558A05D9297FED0138D96519FF">Gerhart Wagner</mods:namePart>
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<mods:namePart id="1D9F3B9BC2BA1FA854DE4006D17CE91D">Andreas Gygax</mods:namePart>
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<mods:title id="D53BF9B88DF8A466CC8036A64C6CBC00">Flora Helvetica</mods:title>
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<treatment id="1E0365388F7A406D0AE2240AD161B581" ID-DOI="http://doi.org/10.5281/zenodo.10855949" ID-GBIF-Taxon="224052688" ID-Zenodo-Dep="10855949" LSID="urn:lsid:plazi:treatment:1E0365388F7A406D0AE2240AD161B581" httpUri="http://treatment.plazi.org/id/1E0365388F7A406D0AE2240AD161B581" lastPageNumber="1116" pageNumber="1116">
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<taxonomicName id="04B07DF65D7B6E4AB99FC44515D0833C" ID-CoL="GW9Y" authority="L." authorityName="L." class="Magnoliopsida" family="Asteraceae" genus="Artemisia" kingdom="Plantae" order="Asterales" pageNumber="1138" phylum="Tracheophyta" rank="species" species="dracunculus">
<emphasis id="87FFF6979EA31AD150EBD485B94E1FD0" italics="true" pageNumber="1116">Artemisia dracunculus</emphasis>
L.
</taxonomicName>
</paragraph>
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Artbeschreibung: 60-120(-150) cm hoch,
<emphasis id="270BB24BDB748E9844CB8710B7320959" bold="true" pageNumber="1116">aromatisch</emphasis>
, kahl.
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<normalizedToken id="C8FB6D492AD28EA4ABD28AD5E61A0AD9" originalValue="Blätter" pageNumber="1138">Blaetter</normalizedToken>
ungeteilt, schmal-lanzettlich
</emphasis>
, ganzrandig,
<quantity id="B6DA73FD8099BBDCBFF21AD65400AEC9" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageNumber="1138" unit="cm" value="6.0" valueMax="10.0" valueMin="2.0">2-10 cm</quantity>
lang und
<quantity id="A0132785AD52BBBA2F54C43C959482B8" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageNumber="1138" unit="mm" value="5.0" valueMax="8.0" valueMin="2.0">2-8 mm</quantity>
breit, mit
<normalizedToken id="0F9961F1761BD1F88A3F02A66708B74F" originalValue="verschmälertem" pageNumber="1138">verschmaelertem</normalizedToken>
Grund sitzend.
<normalizedToken id="69B619BE8EBD6BC602F14DF05DCD38EC" originalValue="Blütenköpfe" pageNumber="1138">Bluetenkoepfe</normalizedToken>
kurz gestielt oder sitzend, nickend,
<emphasis id="F6DCE2FFE6F9ECF4539DF6BDE1CBBB94" bold="true" pageNumber="1116">
Durchmesser
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, mit
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<normalizedToken id="82DB03EA601391B00D4B31B53A2DA43D" originalValue="Röhrenblüten" pageNumber="1138">Roehrenblueten</normalizedToken>
</emphasis>
, ohne
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, in allseitswendiger, lockerer Rispe.
</paragraph>
</subSubSection>
<subSubSection id="D94750606C2346C82CD06AC7DB95384E" pageNumber="1116" type="biology_ecology">
<paragraph id="8C933A74ED5F4CD27FE1404A9951A0E7" pageNumber="1116">
<normalizedToken id="9A26C5D637A6859677C91238D1A27A71" originalValue="Blütezeit" pageNumber="1138">Bluetezeit</normalizedToken>
: 8-9
</paragraph>
<paragraph id="ADFE90448D53243968277C54D939C4E2" pageNumber="1116">
Standort und Verbreitung in der Schweiz: Ufer,
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, als
<normalizedToken id="2009E16456594B448B3C36F35DB062AB" originalValue="Gewürzpflanze" pageNumber="1138">Gewuerzpflanze</normalizedToken>
kultiviert und selten verwildert / kollin-montan(-subalpin) /
</paragraph>
</subSubSection>
<subSubSection id="F9C92DD176DF2F5C4D996DB27723FD59" pageNumber="1116" type="distribution">
<paragraph id="3AB3B0C38E51917DE77E103570B656FE" pageNumber="1116">Verbreitung global: Asiatisch-nordamerikanisch</paragraph>
</subSubSection>
<subSubSection id="32E26A5DA9936B771764D89B77CA4865" pageNumber="1116" type="biology_ecology">
<paragraph id="479127C22C5626D97391CF80D6609C8D" pageNumber="1116">
<normalizedToken id="FCE62B7BE42DBB00C5E307E36ECA3C74" originalValue="Ökologische">Oekologische</normalizedToken>
Zeigerwerte nach
<bibRefCitation id="AC5D72CCE92B2D40272D8C0F62CC9E1B" author="Landolt, E. &amp; Bäumler, B. &amp; Erhardt, A. &amp; Hegg, O. &amp; Klötzli, F. &amp; Lämmler, W. &amp; Nobis, M. &amp; Rudmann-Maurer, K. &amp; Theurillat, J.P. &amp; Urmi, E. &amp; Vust, M. &amp; Wohlgemuth, T." pageNumber="1116" publicationUrl="https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" refString="Landolt, E., Bäumler, B., Erhardt, A., Hegg, O., Klötzli, F., Lämmler, W., Nobis, M., Rudmann-Maurer, K., Theurillat, J.P., Urmi, E., Vust, M., Wohlgemuth, T. 2010. Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen (2nd ed.). https://www.dora.lib4ri.ch/wsl/islandora/object/wsl%3A9966" title="Flora indicativa. Ökologische Zeigerwerte und biologische Kennzeichen zur Flora der Schweiz und der Alpen" year="2010">Landolt &amp; al. (2010)</bibRefCitation>
</paragraph>
<paragraph id="60DA5F5E6FDECAD141A9BF1C6F0C01A6" pageNumber="1116">
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<tr id="4D5B7B43A0E20C7836D168752BF0AA1D" pageNumber="1116">
<th id="665086B4A8E85870B32413AF52BF919C" colspan="2" pageNumber="1116">
<emphasis id="60C39CCD0D4D7FC79B447EDF5BBF491E" bold="true" pageNumber="1116">Bodenfaktoren</emphasis>
</th>
<th id="7EAD430F87563F6C63541888BD356E1A" colspan="2" pageNumber="1116">
<emphasis id="0042B95A53413FB0CA25DDFCE7CF4502" bold="true" pageNumber="1116">Klimafaktoren</emphasis>
</th>
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<emphasis id="7C1AF23528CDDED879CA0590B49E4ECC" bold="true" pageNumber="1116">Salztoleranz</emphasis>
</th>
</tr>
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<td id="C018C79BC61835608B8C5F8C16703FD6" pageNumber="1116">Feuchtezahl F</td>
<td id="B6A64F4EAE0C8FC22FEBC27BA82D1B21" pageNumber="1116" style="text-align:left;width:100px">
frisch; Feuchtigkeit stark wechselnd (mehr als
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2 Stufen)
</td>
<td id="68CD6004D6FF63A879421AB5A9465961" pageNumber="1116">Lichtzahl L</td>
<td id="3FFC21A098A4875B72363EFFE3CD55C9" pageNumber="1116" style="text-align:left;width:60px">halbschattig</td>
<td id="D9C16F4B58CB519E5EF63F73421BC4E2" pageNumber="1116">Salzzeichen</td>
<td id="A92BCA6973A8E1054AD199EDDB4DB53F" pageNumber="1116" style="text-align:left;width:60px">--</td>
</tr>
<tr id="798E7EC290FFD38515CBCFBFF67D0E21" pageNumber="1116">
<td id="4B9EAF88222813ADDFA1AC624645D69B" pageNumber="1116">Reaktionszahl R</td>
<td id="78F9971E8A30DD4B151A033677779009" pageNumber="1116" style="text-align:left">schwach sauer bis neutral (pH 4.5-7.5)</td>
<td id="A5A0DDBD9E89DF5473ACA886EF0FB6E4" pageNumber="1116">Temperaturzahl T</td>
<td id="C9F88D56C3FD6D5D0EB9F0553CD10D51" pageNumber="1116" style="text-align:left">
sehr warm-kollin (nur an
<normalizedToken id="5F631F0A42B9407E88FCA5AF1B94D3DC" originalValue="wärmsten">waermsten</normalizedToken>
Stellen, Hauptverbreitung in
<normalizedToken id="27094D27B51E35FE0CD12D3945397A70" originalValue="Südeuropa">Suedeuropa</normalizedToken>
)
</td>
</tr>
<tr id="8B42D35DE8EC0705C0E90BBB5911AAF3" pageNumber="1116">
<td id="2BB57359030D18037B8433D03DA1EDD8" pageNumber="1116">
<normalizedToken id="D5B344A52E600C27C30B942FCC4B8BB4" originalValue="Nährstoffzahl">Naehrstoffzahl</normalizedToken>
N
</td>
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<normalizedToken id="7478C5C9F4AC0B9EB5E57348B6557D71" originalValue="nährstoffreich">naehrstoffreich</normalizedToken>
</td>
<td id="7078EDAB6E315C378D1367163335D9CE" pageNumber="1116">
<normalizedToken id="51A8DD1CC7446527FBEE35990F97F6EF" originalValue="Kontinentalitätszahl">Kontinentalitaetszahl</normalizedToken>
K
</td>
<td id="9CEB8DA34DFBE5DCC403E964590C51EE" pageNumber="1116" style="text-align:left">subkontinental (niedrige relative Luftfeuchtigkeit, grosse Temperaturschwankungen, eher kalte Winter)</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="62936EF1658D2B0F399F76F4A349AB9C" pageNumber="1116" type="vernacular_names">
<paragraph id="2C377A22C0CB2840CC4AED1ECF2F1437" pageNumber="1116">
Volksname Deutscher Name:
<vernacularName id="ED359A4EC0DA4A9C57272B5B2E9AF197" language="deu" pageNumber="1116">Estragon</vernacularName>
Nom
<normalizedToken id="D2C69A4CB0CF51B6D2B4D97B40594F0D" originalValue="français">francais</normalizedToken>
:
<vernacularName id="589CE2CDE9A8DD36B610DE47933CAA9C" language="fra" pageNumber="1116">Estragon</vernacularName>
Nome italiano:
<vernacularName id="6E404DB461BE7D58FABD714FE0C8786F" language="ita" pageNumber="1116">Assenzio dragoncello</vernacularName>
,
<vernacularName id="5C27A255A1E39F110E414635E111A678" language="ita" pageNumber="1116">Estragone</vernacularName>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="C2F3B51DB0F989C3962AAA2EA613216A" ID-CLB-Dataset="2765" ID-DOI="10.11646/zootaxa.5087.1.4" ID-GBIF-Dataset="c5ac2c46-ae52-4f48-8bc1-61f73b4d06f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5819794" ID-ZooBank="B4061200-FCA1-43A5-811C-0C558043BA46" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641371341574" checkinUser="plazi" docAuthor="Lee, Ga-Eun &amp; Li, Houhun" docDate="2022" docId="1E0387F2FFCAFFE044A8FC0B91EA725E" docLanguage="en" docName="zootaxa.5087.1.4.pdf" docOrigin="Zootaxa 5087 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tenupalpa xanthodora Lee &amp; Li 2022, comb. nov." docType="treatment" docVersion="7" lastPageNumber="107" masterDocId="E23AFF8AFFEAFFC0443FFF819035751D" masterDocTitle="Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species" masterLastPageNumber="111" masterPageNumber="75" pageNumber="107" updateTime="1699295762844" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="17639FF4E86F8D60E9D111A31C7E6FA8">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<mods:affiliation id="2FBEC0F19031BEA789737A4C9A159BA9">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China &amp; The Research Institute for Sciences, Jeju National University, Jeju 63243, Korea &amp; gaeunlee 486 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5774 - 5850</mods:affiliation>
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<treatment id="1E0387F2FFCAFFE044A8FC0B91EA725E" ID-DOI="http://doi.org/10.5281/zenodo.5819868" ID-GBIF-Taxon="191928518" ID-Zenodo-Dep="5819868" LSID="urn:lsid:plazi:treatment:1E0387F2FFCAFFE044A8FC0B91EA725E" httpUri="http://treatment.plazi.org/id/1E0387F2FFCAFFE044A8FC0B91EA725E" lastPageNumber="107" pageId="32" pageNumber="107">
<subSubSection id="DEB0656FFFCAFFE044A8FC0B933376B8" box="[151,774,906,933]" pageId="32" pageNumber="107" type="nomenclature">
<paragraph id="961536E4FFCAFFE044A8FC0B933376B8" blockId="32.[151,774,906,969]" box="[151,774,906,933]" pageId="32" pageNumber="107">
<heading id="CD5D8188FFCAFFE044A8FC0B933376B8" bold="true" box="[151,774,906,933]" fontSize="11" level="1" pageId="32" pageNumber="107" reason="1">
<emphasis id="A4DEEAF6FFCAFFE044A8FC0B933376B8" bold="true" box="[151,774,906,933]" pageId="32" pageNumber="107">
<taxonomicName id="51AA4D67FFCAFFE044A8FC0B924D76B9" ID-CoL="98YHX" authority="(Meyrick, 1923)" authorityName="Lee &amp; Li" authorityYear="2022" baseAuthorityName="Meyrick" baseAuthorityYear="1923" box="[151,632,906,933]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="species" species="xanthodora" status="comb. nov.">
<emphasis id="A4DEEAF6FFCAFFE044A8FC0B919F76B8" bold="true" box="[151,426,906,933]" italics="true" pageId="32" pageNumber="107">Tenupalpa xanthodora</emphasis>
(
<bibRefCitation id="F23B4B15FFCAFFE04585FC0B924576B9" author="Meyrick, E." box="[442,624,906,932]" pageId="32" pageNumber="107" pagination="1 - 32" refId="ref23866" refString="Meyrick, E. (1923) Exotic Microlepidoptera III. Thornhanger, Marlborough, Wilts, pp. 1 - 32." type="book chapter" year="1923">Meyrick, 1923</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BFED578DFFCAFFE046BFFC0A933376B8" box="[640,774,907,933]" pageId="32" pageNumber="107" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFCAFFE044A8FC2E910076D4" box="[151,309,943,969]" pageId="32" pageNumber="107" type="description">
<paragraph id="961536E4FFCAFFE044A8FC2E910076D4" blockId="32.[151,774,906,969]" box="[151,309,943,969]" pageId="32" pageNumber="107">
(
<figureCitation id="0E912A61FFCAFFE044A0FC2E911876D4" box="[159,301,943,969]" captionStart="FIGURES 98–102" captionStartId="33.[151,264,1913,1937]" captionTargetBox="[177,1411,181,1825]" captionTargetId="figure-18@33.[177,1411,181,1825]" captionTargetPageId="33" captionText="FIGURES 98–102. Type specimen of Tenupalpa xanthodora deposited in NHMUK. 98. Holotype, male; 99. Label data; 100, 101. Male genitalia (holotype), slide no. J.F.G.C. 8380, 100a, detail of tegumen tubercles; 102. Male sternum VIII and coremata." figureDoi="http://doi.org/10.5281/zenodo.5819842" httpUri="https://zenodo.org/record/5819842/files/figure.png" pageId="32" pageNumber="107">Figs 98–102</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFCAFFE044A8FC7292C07134" pageId="32" pageNumber="107" type="reference_group">
<paragraph id="961536E4FFCAFFE044A8FC7292C07134" blockId="32.[151,1436,1011,1066]" pageId="32" pageNumber="107">
<treatmentCitationGroup id="B6BA11CAFFCAFFE044A8FC7292C07134" pageId="32" pageNumber="107">
<treatmentCitation id="170B10F5FFCAFFE044A8FC7292767116" author="Meyrick, E." box="[151,579,1011,1035]" page="24" pageId="32" pageNumber="107" year="1923">
<taxonomicName id="51AA4D67FFCAFFE044A8FC7292767116" ID-CoL="93L49" authority="Meyrick, 1923: 24" authorityName="Meyrick" authorityPageNumber="24" authorityYear="1923" box="[151,579,1011,1035]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="species" species="xanthodora">
<emphasis id="A4DEEAF6FFCAFFE044A8FC72914B7116" box="[151,382,1011,1035]" italics="true" pageId="32" pageNumber="107">Thiotricha xanthodora</emphasis>
<bibRefCitation id="F23B4B15FFCAFFE045BBFC7292767116" author="Meyrick, E." box="[388,579,1011,1035]" pageId="32" pageNumber="107" pagination="1 - 32" refId="ref23866" refString="Meyrick, E. (1923) Exotic Microlepidoptera III. Thornhanger, Marlborough, Wilts, pp. 1 - 32." type="book chapter" year="1923">Meyrick, 1923: 24</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="170B10F5FFCAFFE04671FC72928D7116" author="Meyrick, E." box="[590,696,1011,1035]" page="103" pageId="32" pageNumber="107" year="1925">
<bibRefCitation id="F23B4B15FFCAFFE04671FC72928D7116" author="Meyrick, E." box="[590,696,1011,1035]" pageId="32" pageNumber="107" pagination="1 - 5" refId="ref23889" refString="Meyrick, E. (1925) Lepidoptera Heterocera. Family Gelechiadae [sic]. In: Wytsman, P. (Ed.), Genera Insectorum. Vol. 184. Louis Desmet-Vertneuil, Bruxelles, pp. 1 - 290, pls. 1 - 5." type="book chapter" year="1925">1925: 103</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="170B10F5FFCAFFE046FDFC72934C7116" author="Gaede, M." box="[706,889,1011,1035]" page="307" pageId="32" pageNumber="107" year="1937">
<bibRefCitation id="F23B4B15FFCAFFE046FDFC72934C7116" author="Gaede, M." box="[706,889,1011,1035]" pageId="32" pageNumber="107" pagination="1 - 630" refId="ref23337" refString="Gaede, M. (1937) Gelechiidae. Lepidopterorum Catalogus, 79, 1 - 630." type="journal article" year="1937">Gaede, 1937: 307</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="170B10F5FFCAFFE047BBFC72940B7116" author="Clarke, J. F. G." box="[900,1086,1011,1035]" page="475" pageId="32" pageNumber="107" year="1969">
<bibRefCitation id="F23B4B15FFCAFFE047BBFC72940B7116" author="Clarke, J. F. G." box="[900,1086,1011,1035]" pageId="32" pageNumber="107" refId="ref23257" refString="Clarke, J. F. G. (1969) Catalogue of the type specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. 7. Trustees of the British Museum, London, 531 pp., 265 pls." type="book" year="1969">Clarke, 1969: 475</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="170B10F5FFCAFFE04076FC7295097116" author="Ivinskis P. P. &amp; Piskunov, V. I. &amp; Emeliyanov, I. M." box="[1097,1340,1011,1035]" page="41" pageId="32" pageNumber="107" year="1984">
<bibRefCitation id="F23B4B15FFCAFFE04076FC7295097116" author="Ivinskis P. P. &amp; Piskunov, V. I. &amp; Emeliyanov, I. M." box="[1097,1340,1011,1035]" pageId="32" pageNumber="107" pagination="37 - 44" refId="ref23428" refString="Ivinskis P. P., Piskunov, V. I. &amp; Emeliyanov, I. M. (1984) [Review of the fauna of Gelechiid moths of the Thiotricha Meyrick, 1886 genus and its place in the system of Lepidoptera: Gelechiidae, Dichomerinae]. Trudy AN Lithuanian SSR, 2 (86), 37 - 44. [in Russian]" type="journal article" year="1984">
Ivinskis
<emphasis id="A4DEEAF6FFCAFFE0409FFC7594F97116" box="[1184,1228,1011,1035]" italics="true" pageId="32" pageNumber="107">et al</emphasis>
., 1984: 41
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="170B10F5FFCAFFE04178FC75917F7137" author="Kyaw, K. M. M. &amp; Yagi, S. &amp; Oku, J. &amp; Sakamaki, Y. &amp; Hirowatari, T." page="88" pageId="32" pageNumber="107" year="2019">
<bibRefCitation id="F23B4B15FFCAFFE04178FC75917F7137" author="Kyaw, K. M. M. &amp; Yagi, S. &amp; Oku, J. &amp; Sakamaki, Y. &amp; Hirowatari, T." pageId="32" pageNumber="107" pagination="67 - 99" refId="ref23620" refString="Kyaw, K. M. M., Yagi, S., Oku, J., Sakamaki, Y. &amp; Hirowatari, T. (2019) Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys, 897, 67 - 99." type="journal article" year="2019">
Kyaw
<emphasis id="A4DEEAF6FFCAFFE041B5FC7590EE7137" italics="true" pageId="32" pageNumber="107">et al</emphasis>
., 2019: 88
</bibRefCitation>
</treatmentCitation>
. TL: Pyinmana,
<collectingCountry id="EEBD7674FFCAFFE045CDFB9392637137" box="[498,598,1042,1066]" name="Myanmar" pageId="32" pageNumber="107">Myanmar</collectingCountry>
. TD: NHMUK.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFCAFFE044A8FBD9912E718B" pageId="32" pageNumber="107" type="materials_examined">
<paragraph id="961536E4FFCAFFE044A8FBD9912E718B" blockId="32.[151,1437,1111,1859]" pageId="32" pageNumber="107">
<emphasis id="A4DEEAF6FFCAFFE044A8FBD9917E716F" bold="true" box="[151,331,1112,1138]" pageId="32" pageNumber="107">Type material.</emphasis>
<materialsCitation id="26C23CB9FFCAFFE04568FBD9912E718B" ID-GBIF-Occurrence="3427696347" collectionCode="NHMUK" collectorName="J.F.G. Clarke" country="Myanmar" location="MYANMAR" pageId="32" pageNumber="107" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="49118846FFCAFFE04568FBD9918A716F" box="[343,447,1112,1138]" pageId="32" pageNumber="107" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFCAFFE045F3FBD891D6716E" box="[460,483,1113,1139]" pageId="32" pageNumber="107" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFCAFFE045CBFBD992BC716F" box="[500,649,1112,1138]" name="Myanmar" pageId="32" pageNumber="107">MYANMAR</collectingCountry>
, Pyinmana, 18.iii.[?], leg. T.B.F., genitalia slide no. J.
<collectorName id="3B5F5332FFCAFFE04132FBD9956D716F" box="[1293,1368,1112,1138]" name="J.F.G. Clarke" pageId="32" pageNumber="107">F.G.C.</collectorName>
8380 (
<collectionCode id="F0BBAE21FFCAFFE044A0FBFD9125718B" box="[159,272,1148,1174]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="32" pageNumber="107" type="Museum">NHMUK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFCAFFE044F8FB21935F7073" pageId="32" pageNumber="107" type="diagnosis">
<paragraph id="961536E4FFCAFFE044F8FB21935F7073" blockId="32.[151,1437,1111,1859]" pageId="32" pageNumber="107">
<emphasis id="A4DEEAF6FFCAFFE044F8FB21917771A7" bold="true" box="[199,322,1184,1210]" pageId="32" pageNumber="107">Diagnosis.</emphasis>
The forewing pattern of
<taxonomicName id="51AA4D67FFCAFFE0465FFB2092CB71A7" authorityName="Meyrick" authorityYear="1923" box="[608,766,1184,1210]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="species" species="xanthodora">
<emphasis id="A4DEEAF6FFCAFFE0465FFB2092CB71A7" box="[608,766,1184,1210]" italics="true" pageId="32" pageNumber="107">T. xanthodora</emphasis>
</taxonomicName>
is quite different from that of its congeners, with the basal 1/3 marked with black and yellow longitudinal streaks (
<figureCitation id="0E912A61FFCAFFE04735FB45936A71C3" box="[778,863,1220,1246]" captionStart="FIGURES 98–102" captionStartId="33.[151,264,1913,1937]" captionTargetBox="[177,1411,181,1825]" captionTargetId="figure-18@33.[177,1411,181,1825]" captionTargetPageId="33" captionText="FIGURES 98–102. Type specimen of Tenupalpa xanthodora deposited in NHMUK. 98. Holotype, male; 99. Label data; 100, 101. Male genitalia (holotype), slide no. J.F.G.C. 8380, 100a, detail of tegumen tubercles; 102. Male sternum VIII and coremata." figureDoi="http://doi.org/10.5281/zenodo.5819842" httpUri="https://zenodo.org/record/5819842/files/figure.png" pageId="32" pageNumber="107">Fig. 98</figureCitation>
). Other species of
<taxonomicName id="51AA4D67FFCAFFE0400FFB45949071C3" authorityName="Lee et Li" authorityYear="2021" box="[1072,1189,1220,1246]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFCAFFE0400FFB45949071C3" box="[1072,1189,1220,1246]" italics="true" pageId="32" pageNumber="107">Tenupalpa</emphasis>
</taxonomicName>
usually have the base uniformly colored without distinct pattern elements. The male genitalia are somewhat similar to those of
<taxonomicName id="51AA4D67FFCAFFE04102FB6890D3703B" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFCAFFE04102FB6890D3703B" italics="true" pageId="32" pageNumber="107">T. flavitermina</emphasis>
</taxonomicName>
and
<taxonomicName id="51AA4D67FFCAFFE04522FA8C91B9703B" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[285,396,1293,1318]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFCAFFE04522FA8C91B9703B" box="[285,396,1293,1318]" italics="true" pageId="32" pageNumber="107">T. acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFCAFFE045ACFA8D91D9703B" bold="true" box="[403,492,1292,1318]" pageId="32" pageNumber="107">
<taxonomicNameLabel id="BFED578DFFCAFFE045ACFA8D91D9703B" box="[403,492,1292,1318]" pageId="32" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having a sinuous distal part of the aedeagus and oval anellus lobes bearing a long apical bristle. However, the genitalia of
<taxonomicName id="51AA4D67FFCAFFE04665FAB092C27057" authorityName="Meyrick" authorityYear="1923" box="[602,759,1328,1354]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="species" species="xanthodora">
<emphasis id="A4DEEAF6FFCAFFE04665FAB092C27057" box="[602,759,1328,1354]" italics="true" pageId="32" pageNumber="107">T. xanthodora</emphasis>
</taxonomicName>
can be distinguished from those of the latter two species by the valva expanded in the apical 1/3 and the larger anellus lobes.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFCAFFE044F8FAF991EF708E" box="[199,474,1400,1427]" pageId="32" pageNumber="107" type="distribution">
<paragraph id="961536E4FFCAFFE044F8FAF991EF708E" blockId="32.[151,1437,1111,1859]" box="[199,474,1400,1427]" pageId="32" pageNumber="107">
<emphasis id="A4DEEAF6FFCAFFE044F8FAF9916C708F" bold="true" box="[199,345,1400,1426]" pageId="32" pageNumber="107">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFCAFFE04558FAF891E3708E" box="[359,470,1401,1427]" name="Myanmar" pageId="32" pageNumber="107">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFCAFFE044F8FA1D91EA725E" pageId="32" pageNumber="107" type="discussion">
<paragraph id="961536E4FFCAFFE044F8FA1D91E3735B" blockId="32.[151,1437,1111,1859]" pageId="32" pageNumber="107">
<emphasis id="A4DEEAF6FFCAFFE044F8FA1D910E70AB" bold="true" box="[199,315,1436,1462]" pageId="32" pageNumber="107">Remarks.</emphasis>
The species was originally described in
<taxonomicName id="51AA4D67FFCAFFE046C4FA1D934470AB" authorityName="Meyrick" authorityYear="1886" box="[763,881,1436,1462]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="A4DEEAF6FFCAFFE046C4FA1D934470AB" box="[763,881,1436,1462]" italics="true" pageId="32" pageNumber="107">Thiotricha</emphasis>
</taxonomicName>
based on a single specimen collected from
<collectingCountry id="EEBD7674FFCAFFE0416DFA1C90F670C6" name="Myanmar" pageId="32" pageNumber="107">Myanmar</collectingCountry>
(labeled as
<collectingCountry id="EEBD7674FFCAFFE04576FA4091AF70C6" box="[329,410,1473,1499]" name="Myanmar" pageId="32" pageNumber="107">Burma</collectingCountry>
). The male genitalia and the abdomen of the
<typeStatus id="49118846FFCAFFE047AEFA4193C670C7" box="[913,1011,1472,1498]" pageId="32" pageNumber="107" type="holotype">holotype</typeStatus>
housed in NHMUK are characteristic of
<taxonomicName id="51AA4D67FFCAFFE04486FA65911B70E3" authorityName="Lee et Li" authorityYear="2021" box="[185,302,1508,1534]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFCAFFE04486FA65911B70E3" box="[185,302,1508,1534]" italics="true" pageId="32" pageNumber="107">Tenupalpa</emphasis>
</taxonomicName>
: small uncus, tegumen bearing a bundle of tubercles (
<figureCitation id="0E912A61FFCAFFE04796FA65943B70E3" box="[937,1038,1508,1534]" captionStart="FIGURES 98–102" captionStartId="33.[151,264,1913,1937]" captionTargetBox="[177,1411,181,1825]" captionTargetId="figure-18@33.[177,1411,181,1825]" captionTargetPageId="33" captionText="FIGURES 98–102. Type specimen of Tenupalpa xanthodora deposited in NHMUK. 98. Holotype, male; 99. Label data; 100, 101. Male genitalia (holotype), slide no. J.F.G.C. 8380, 100a, detail of tegumen tubercles; 102. Male sternum VIII and coremata." figureDoi="http://doi.org/10.5281/zenodo.5819842" httpUri="https://zenodo.org/record/5819842/files/figure.png" pageId="32" pageNumber="107">Fig. 100</figureCitation>
), rounded sternum VIII, and short external coremata (
<figureCitation id="0E912A61FFCAFFE04551F98991E5733F" box="[366,464,1544,1570]" captionStart="FIGURES 98–102" captionStartId="33.[151,264,1913,1937]" captionTargetBox="[177,1411,181,1825]" captionTargetId="figure-18@33.[177,1411,181,1825]" captionTargetPageId="33" captionText="FIGURES 98–102. Type specimen of Tenupalpa xanthodora deposited in NHMUK. 98. Holotype, male; 99. Label data; 100, 101. Male genitalia (holotype), slide no. J.F.G.C. 8380, 100a, detail of tegumen tubercles; 102. Male sternum VIII and coremata." figureDoi="http://doi.org/10.5281/zenodo.5819842" httpUri="https://zenodo.org/record/5819842/files/figure.png" pageId="32" pageNumber="107">Fig. 102</figureCitation>
). Based on these morphological traits, we propose the new combination,
<taxonomicName id="51AA4D67FFCAFFE0413FF988917A735B" authority="(Meyrick, 1923)" authorityName="Lee &amp; Li" authorityYear="2022" baseAuthorityName="Meyrick" baseAuthorityYear="1923" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="species" species="xanthodora" status="comb. nov.">
<emphasis id="A4DEEAF6FFCAFFE0413FF98895A9733F" box="[1280,1436,1544,1570]" italics="true" pageId="32" pageNumber="107">T. xanthodora</emphasis>
(
<bibRefCitation id="F23B4B15FFCAFFE044A0F9AD9173735B" author="Meyrick, E." box="[159,326,1580,1606]" pageId="32" pageNumber="107" pagination="1 - 32" refId="ref23866" refString="Meyrick, E. (1923) Exotic Microlepidoptera III. Thornhanger, Marlborough, Wilts, pp. 1 - 32." type="book chapter" year="1923">Meyrick, 1923</bibRefCitation>
)
</taxonomicName>
<emphasis id="A4DEEAF6FFCAFFE04569F9AD91E3735B" bold="true" box="[342,470,1580,1606]" pageId="32" pageNumber="107">
<taxonomicNameLabel id="BFED578DFFCAFFE04569F9AD91E3735B" box="[342,470,1580,1606]" pageId="32" pageNumber="107" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="961536E4FFCAFFE044F8F9D091EA725E" blockId="32.[151,1437,1111,1859]" pageId="32" pageNumber="107">
In our previous work (
<bibRefCitation id="F23B4B15FFCAFFE045FAF9D192407377" author="Lee, G. - E. &amp; Han, T. M. &amp; Park, H. C. &amp; Qi, M. J. &amp; Li, H. H." box="[453,629,1616,1643]" pageId="32" pageNumber="107" pagination="357 - 379" refId="ref23692" refString="Lee, G. - E., Han, T. M., Park, H. C., Qi, M. J. &amp; Li, H. H. (2021) A phylogeny of the subfamily Thiotrichinae (Lepidoptera: Gel- echiidae) with a revision of the generic classification based on molecular and morphological analyses. Systematic Entomology, 46, 357 - 379. https: // doi. org / 10.1111 / syen. 12466" type="journal article" year="2021">
Lee
<emphasis id="A4DEEAF6FFCAFFE045C6F9D0921E7377" box="[505,555,1616,1642]" italics="true" pageId="32" pageNumber="107">et al</emphasis>
. 2021
</bibRefCitation>
), we incorrectly transferred
<taxonomicName id="51AA4D67FFCAFFE0478BF9D195017377" authority="Meyrick, 1908" authorityName="Meyrick" authorityYear="1908" box="[948,1332,1616,1642]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="species" species="glenias">
<emphasis id="A4DEEAF6FFCAFFE0478BF9D194B67377" box="[948,1155,1616,1642]" italics="true" pageId="32" pageNumber="107">Thiotricha glenias</emphasis>
Meyrick, 1908
</taxonomicName>
, to
<taxonomicName id="51AA4D67FFCAFFE04162F9D090E27393" authorityName="Lee et Li" authorityYear="2021" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFCAFFE04162F9D090E27393" italics="true" pageId="32" pageNumber="107">Tenupalpa</emphasis>
</taxonomicName>
.
<taxonomicName id="51AA4D67FFCAFFE044DAF9F591817393" authorityName="Meyrick" authorityYear="1908" box="[229,436,1652,1678]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="species" species="glenias">
<emphasis id="A4DEEAF6FFCAFFE044DAF9F591817393" box="[229,436,1652,1678]" italics="true" pageId="32" pageNumber="107">Thiotricha glenias</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="51AA4D67FFCAFFE046E3F9F5939F7393" box="[732,938,1652,1678]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A4DEEAF6FFCAFFE046E3F9F593647393" box="[732,849,1652,1678]" italics="true" pageId="32" pageNumber="107">Tenupalpa</emphasis>
species
</taxonomicName>
in wing pattern and coloration; however, its male possesses an expansible hair pencil dorsally on the labial palpus, setae are present on the dorsal surface of the tegumen, and tegumen tubercles are absent. These features are characteristic of many species of
<taxonomicName id="51AA4D67FFCAFFE040C8F93D955873CB" authorityName="Meyrick" authorityYear="1886" box="[1271,1389,1724,1750]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="A4DEEAF6FFCAFFE040C8F93D955873CB" box="[1271,1389,1724,1750]" italics="true" pageId="32" pageNumber="107">Thiotricha</emphasis>
</taxonomicName>
and
<emphasis id="A4DEEAF6FFCAFFE044A8F961913273E7" box="[151,263,1760,1786]" italics="true" pageId="32" pageNumber="107">Pulchrala</emphasis>
. The generic position of the species requires further study, but the combination of morphological characters is certainly not typical of
<taxonomicName id="51AA4D67FFCAFFE04638F88592497203" authorityName="Lee et Li" authorityYear="2021" box="[519,636,1796,1822]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFCAFFE04638F88592497203" box="[519,636,1796,1822]" italics="true" pageId="32" pageNumber="107">Tenupalpa</emphasis>
</taxonomicName>
. Therefore, we propose
<taxonomicName id="51AA4D67FFCAFFE047ACF88594567203" authorityName="Lee &amp; Li" authorityYear="2022" box="[915,1123,1796,1822]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="107" phylum="Arthropoda" rank="species" species="glenias" status="stat. nov.">
<emphasis id="A4DEEAF6FFCAFFE047ACF88594567203" box="[915,1123,1796,1822]" italics="true" pageId="32" pageNumber="107">Thiotricha glenias</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFCAFFE04054F88594E77203" bold="true" box="[1131,1234,1796,1822]" pageId="32" pageNumber="107">
<taxonomicNameLabel id="BFED578DFFCAFFE04054F88594E77203" box="[1131,1234,1796,1822]" pageId="32" pageNumber="107">stat. rev.</taxonomicNameLabel>
</emphasis>
, resurrected from combination with
<taxonomicName id="51AA4D67FFCAFFE0455CF8A991ED725F" authorityName="Lee et Li" authorityYear="2021" box="[355,472,1832,1858]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFCAFFE0455CF8A991ED725F" box="[355,472,1832,1858]" italics="true" pageId="32" pageNumber="107">Tenupalpa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

210
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<mods:title id="DB90263EF666455232DE2B309CEA71E3">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<mods:affiliation id="2C2E6F77EE3D47F9142F350EB8BA4F0A">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China &amp; The Research Institute for Sciences, Jeju National University, Jeju 63243, Korea &amp; gaeunlee 486 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5774 - 5850</mods:affiliation>
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<treatment id="1E0387F2FFE4FFCE44A8FDFA91FE72C2" ID-DOI="http://doi.org/10.5281/zenodo.5819834" ID-GBIF-Taxon="191928541" ID-Zenodo-Dep="5819834" LSID="urn:lsid:zoobank.org:act:0402A49C-1511-41DD-9117-3B7E8E635F99" httpUri="http://treatment.plazi.org/id/1E0387F2FFE4FFCE44A8FDFA91FE72C2" lastPageNumber="89" pageId="14" pageNumber="89">
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<paragraph id="961536E4FFE4FFCE44A8FDFA9241778B" blockId="14.[151,915,635,731]" box="[151,628,635,662]" pageId="14" pageNumber="89">
<heading id="CD5D8188FFE4FFCE44A8FDFA9241778B" bold="true" box="[151,628,635,662]" fontSize="11" level="1" pageId="14" pageNumber="89" reason="1">
<emphasis id="A4DEEAF6FFE4FFCE44A8FDFA9241778B" bold="true" box="[151,628,635,662]" pageId="14" pageNumber="89">
<taxonomicName id="51AA4D67FFE4FFCE44A8FDFA923C778B" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,521,635,662]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="89" phylum="Tracheophyta" rank="species" species="isabellina" status="sp. nov.">
<emphasis id="A4DEEAF6FFE4FFCE44A8FDFA91A6778B" bold="true" box="[151,403,635,662]" italics="true" pageId="14" pageNumber="89">Tenupalpa isabellina</emphasis>
Lee
<emphasis id="A4DEEAF6FFE4FFCE45EFFDFD91D0778B" bold="true" box="[464,485,636,662]" italics="true" pageId="14" pageNumber="89">et</emphasis>
Li
</taxonomicName>
,
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<paragraph id="961536E4FFE4FFCE44A8FD1E91A277A4" blockId="14.[151,915,635,731]" box="[151,407,670,697]" pageId="14" pageNumber="89">
(
<figureCitation id="0E912A61FFE4FFCE44A0FD1E90D277A4" box="[159,231,671,697]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="14" pageNumber="89">Figs 8</figureCitation>
,
<figureCitation id="0E912A61FFE4FFCE44CBFD1E912777A4" box="[244,274,671,697]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="14" pageNumber="89">31</figureCitation>
,
<figureCitation id="0E912A61FFE4FFCE4521FD1F910977A4" box="[286,316,670,697]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="14" pageNumber="89">51</figureCitation>
,
<figureCitation id="0E912A61FFE4FFCE4577FD1E915277A4" box="[328,359,671,697]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="14" pageNumber="89">70</figureCitation>
,
<figureCitation id="0E912A61FFE4FFCE454DFD1E91BA77A4" box="[370,399,671,697]" captionStart="FIGURES 87–90" captionStartId="19.[151,264,1943,1967]" captionTargetBox="[229,1327,198,1905]" captionTargetId="figure-18@19.[196,1391,181,1920]" captionTargetPageId="19" captionText="FIGURES 87–90. Female genitalia of Tenupalpa spp. 87. T. immaculata sp. nov., paratype, slide no. LGE18002; 88. T. isabellina sp. nov., paratype, slide no. LGE17283f; 89. T. flavitermina, slide no. LGE19021; 90. T. biformis, slide no. LLJ15251 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819832" httpUri="https://zenodo.org/record/5819832/files/figure.png" pageId="14" pageNumber="89">88</figureCitation>
)
</paragraph>
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<emphasis id="A4DEEAF6FFE4FFCE44A8FC86917D763D" bold="true" box="[151,328,774,801]" pageId="14" pageNumber="89">Type material.</emphasis>
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and
<collectorName id="3B5F5332FFE4FFCE4667FCA892C57659" box="[600,752,809,836]" pageId="14" pageNumber="89">Yinghui Mou</collectorName>
, genitalia slide no. LGE17282
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.
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<typeStatus id="49118846FFE4FFCE4072FCA894F2765E" box="[1101,1223,809,835]" pageId="14" pageNumber="89" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="546EF806FFE4FFCE40EDFCA895A9765E" box="[1234,1436,809,835]" country="China" name="Guangxi" pageId="14" pageNumber="89">Guangxi Zhuang</collectingRegion>
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:
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,
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, same data as holotype except
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, genitalia slide nos
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f,
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LGE
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<subSubSection id="DEB0656FFFE4FFCE44F8FC1092E776EC" pageId="14" pageNumber="89" type="diagnosis">
<paragraph id="961536E4FFE4FFCE44F8FC1092E776EC" blockId="14.[151,1437,774,2015]" pageId="14" pageNumber="89">
<emphasis id="A4DEEAF6FFE4FFCE44F8FC10917776B6" bold="true" box="[199,322,913,939]" pageId="14" pageNumber="89">Diagnosis.</emphasis>
The species is most similar to
<taxonomicName id="51AA4D67FFE4FFCE46ABFC13930776B6" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[660,818,913,939]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="89" phylum="Tracheophyta" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE4FFCE46ABFC13930776B6" box="[660,818,913,939]" italics="true" pageId="14" pageNumber="89">T. immaculata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE4FFCE4707FC1093A476B6" bold="true" box="[824,913,913,939]" pageId="14" pageNumber="89">
<taxonomicNameLabel id="BFED578DFFE4FFCE4707FC1093A476B6" box="[824,913,913,939]" pageId="14" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and differences between the two are presented under the latter species. In the male genitalia, the shape of the uncus is unique within the genus, with its distal part narrowly elongated compared to that of congeners.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE4FFCE44F8FC789495726B" pageId="14" pageNumber="89" type="description">
<paragraph id="961536E4FFE4FFCE44F8FC7892757183" blockId="14.[151,1437,774,2015]" pageId="14" pageNumber="89">
<emphasis id="A4DEEAF6FFE4FFCE44F8FC78916C710E" bold="true" box="[199,345,1017,1043]" pageId="14" pageNumber="89">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFE4FFCE458EFC7892307109" box="[433,517,1017,1044]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="14" pageNumber="89">Fig. 51</figureCitation>
).
<emphasis id="A4DEEAF6FFE4FFCE4625FC789262710E" box="[538,599,1017,1043]" italics="true" pageId="14" pageNumber="89">Head</emphasis>
: Vertex greyish brown or fuscous, frons creamy white. Labial palpus pale brown; apex of palpomere I fuscous on outer surface, apex of palpomere II often slightly tinged with fuscous on outer surface; palpomere III slightly longer than II, strongly mixed with grey. Antenna with scape creamy white except posterior margin fuscous; flagellum dark fuscous except basal half of dorsal surface creamy white to pale brown; male cilia as long as diameter.
</paragraph>
<paragraph id="961536E4FFE4FFCE44F8FB2691F4708C" blockId="14.[151,1437,774,2015]" pageId="14" pageNumber="89">
<emphasis id="A4DEEAF6FFE4FFCE44F8FB26912D71DC" box="[199,280,1191,1217]" italics="true" pageId="14" pageNumber="89">Thorax</emphasis>
: Dorsum and tegula pale brown except anterior 1/2 or 2/5 dark fuscous. Wingspan 9.0–10.5 mm; forewing ground color pale greyish brown, costa dark grey, base dark fuscous above fold, middle of wing often strongly suffused with grey or dark grey; two indistinct grey lines arising from outer margin of former greyish suffusion: upper line narrowly to broadly extending toward apex along anterior 1/3 of wing, broadened and smudged near apex, lower line extending along fold and broadened near tornus; fringe scales pale greyish brown. Hindwing grey; fringe scales pale greyish brown. Legs creamy white to pale brown; mid-tibia with a fuscous spot at distal 1/3; hind tibia broadly fuscous at middle.
</paragraph>
<paragraph id="961536E4FFE4FFCE44F8FA189495726B" blockId="14.[151,1437,774,2015]" pageId="14" pageNumber="89">
<emphasis id="A4DEEAF6FFE4FFCE44F8FA18910470AE" box="[199,305,1433,1459]" italics="true" pageId="14" pageNumber="89">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFE4FFCE4500FA1891B670AE" box="[319,387,1433,1459]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="14" pageNumber="89">Fig. 8</figureCitation>
): Male coremata nearly as long as sternum VIII; sternum VIII pentagonal. Male genitalia (
<figureCitation id="0E912A61FFE4FFCE4150FA18908170CB" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="14" pageNumber="89">Fig. 70</figureCitation>
) with uncus widest near base, gradually narrowing to middle, then uniformly elongate to rounded apex. Gnathos hook moderately long, sharply pointed apically. Tegumen approximately 4 times as long as uncus; with numerous ampulliform tubercles situated at middle of ventral surface, each bearing a long seta (
<figureCitation id="0E912A61FFE4FFCE4075F98094A87306" box="[1098,1181,1537,1563]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="14" pageNumber="89">Fig. 31</figureCitation>
). Valva slender, gradually narrowing from base to basal 1/3, then uniformly elongate to 2/3; distal 1/3 spatulate, rounded at apex. Anellus lobe elliptical, 1/7 length of valva, sparsely setose on inner margin; apical bristle longer than anellus lobe, curved inward. Juxta setose in distal 1/3 of inner margin, near base of valva slightly protruded. Vinculum simple, with posterior margin broadly convex. Saccus broad, sub-triangular, exceeding tegumen pedunculus. Aedeagus with basal 1/2 globular; distal 1/2 slender, straight or curved downward. Female genitalia (
<figureCitation id="0E912A61FFE4FFCE47DEF92E940773D4" box="[993,1074,1711,1737]" captionStart="FIGURES 87–90" captionStartId="19.[151,264,1943,1967]" captionTargetBox="[229,1327,198,1905]" captionTargetId="figure-18@19.[196,1391,181,1920]" captionTargetPageId="19" captionText="FIGURES 87–90. Female genitalia of Tenupalpa spp. 87. T. immaculata sp. nov., paratype, slide no. LGE18002; 88. T. isabellina sp. nov., paratype, slide no. LGE17283f; 89. T. flavitermina, slide no. LGE19021; 90. T. biformis, slide no. LLJ15251 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819832" httpUri="https://zenodo.org/record/5819832/files/figure.png" pageId="14" pageNumber="89">Fig. 88</figureCitation>
) with papillae anales simple and setose. Apophyses anteriores 1/2 length of apophyses posteriores. Tergum VIII divided into two sclerites; sternum VIII produced anteriorly, with two sub-triangular flaps covering ostium bursae. Ostium bursae near anterior margin of sternum VIII. Ductus bursae narrow at base, then gradually widening toward corpus bursae, ductus seminalis arising from posterior 1/3 of ductus bursae. Corpus bursae ovate, approximately 1.5 times as long as ductus bursae; signum small, slender, somewhat elongate fusiform, situated near entrance of ductus bursae.
</paragraph>
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<subSubSection id="DEB0656FFFE4FFCE44F8F8FF92197287" box="[199,556,1918,1946]" pageId="14" pageNumber="89" type="distribution">
<paragraph id="961536E4FFE4FFCE44F8F8FF92197287" blockId="14.[151,1437,774,2015]" box="[199,556,1918,1946]" pageId="14" pageNumber="89">
<emphasis id="A4DEEAF6FFE4FFCE44F8F8FF916C7285" bold="true" box="[199,345,1918,1944]" pageId="14" pageNumber="89">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFE4FFCE4558F8FE919F7284" box="[359,426,1919,1945]" name="China" pageId="14" pageNumber="89">China</collectingCountry>
(
<collectingRegion id="546EF806FFE4FFCE4586F8FE92157287" box="[441,544,1919,1946]" country="China" name="Guangxi" pageId="14" pageNumber="89">Guangxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE4FFCE44F8F82091FE72C2" pageId="14" pageNumber="89" type="etymology">
<paragraph id="961536E4FFE4FFCE44F8F82091FE72C2" blockId="14.[151,1437,774,2015]" pageId="14" pageNumber="89">
<emphasis id="A4DEEAF6FFE4FFCE44F8F820917D72A6" bold="true" box="[199,328,1953,1979]" pageId="14" pageNumber="89">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFE4FFCE4770F82093FD72A6" box="[847,968,1953,1979]" italics="true" pageId="14" pageNumber="89">isabellinus</emphasis>
(isabelline), referring to the pale greyish brown forewing coloration.
</paragraph>
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<taxonomicName id="51AA4D67FFE6FFCC44A8FAF69236708F" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,515,1399,1426]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="fuscalata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE6FFCC44A8FAF691B8708F" bold="true" box="[151,397,1399,1426]" italics="true" pageId="12" pageNumber="87">Tenupalpa fuscalata</emphasis>
Lee
<emphasis id="A4DEEAF6FFE6FFCC45F6FAF991EB708F" bold="true" box="[457,478,1400,1426]" italics="true" pageId="12" pageNumber="87">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFE6FFCC462FFAF9925B708F" box="[528,622,1400,1426]" pageId="12" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="DEB0656FFFE6FFCC44A8FA1D939070CE" pageId="12" pageNumber="87" type="description">
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(
<figureCitation id="0E912A61FFE6FFCC44A0FA1D90DD70AB" box="[159,232,1436,1462]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="12" pageNumber="87">Figs 7</figureCitation>
,
<figureCitation id="0E912A61FFE6FFCC44CBFA1D912770AB" box="[244,274,1436,1462]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="12" pageNumber="87">29</figureCitation>
,
<figureCitation id="0E912A61FFE6FFCC4521FA1D910870AB" box="[286,317,1436,1462]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="12" pageNumber="87">49</figureCitation>
,
<figureCitation id="0E912A61FFE6FFCC4577FA1D915370AB" box="[328,358,1436,1462]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="12" pageNumber="87">68</figureCitation>
,
<figureCitation id="0E912A61FFE6FFCC454DFA1D91BA70AA" box="[370,399,1436,1463]" captionStart="FIGURES 84–86" captionStartId="18.[151,264,1943,1967]" captionTargetBox="[351,1240,197,1904]" captionTargetId="figure-22@18.[327,1260,181,1920]" captionTargetPageId="18" captionText="FIGURES 84–86. Genitalia of Tenupalpa spp. 84. T. dissimilis sp. nov., male, holotype, slide no. LGE19007; 85. T. angustella, female, slide no. LGE17065; 86. T. fuscalata sp. nov., female, paratype, slide no. LGE18832 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819830" httpUri="https://zenodo.org/record/5819830/files/figure.png" pageId="12" pageNumber="87">86</figureCitation>
)
</paragraph>
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<subSubSection id="DEB0656FFFE6FFCC44A8F98092F37221" pageId="12" pageNumber="87" type="materials_examined">
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<emphasis id="A4DEEAF6FFE6FFCC44A8F98091707306" bold="true" box="[151,325,1537,1563]" pageId="12" pageNumber="87">Type material.</emphasis>
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<collectingDate id="F250E9CCFFE6FFCC44C4F9A491427322" box="[251,375,1573,1599]" pageId="12" pageNumber="87" value="2018-08-03">3.viii.2018</collectingDate>
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, leg.
<collectorName id="3B5F5332FFE6FFCC458BF9A492287322" box="[436,541,1573,1599]" pageId="12" pageNumber="87">Mujie Qi</collectorName>
, genitalia slide no. LGE18829
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<typeStatus id="49118846FFE6FFCC47BAF9A493CA7322" box="[901,1023,1573,1599]" pageId="12" pageNumber="87" type="paratype">Paratypes</typeStatus>
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:
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<date id="E2141024FFE6FFCC473CF9C8934A737E" box="[771,895,1609,1635]" pageId="12" pageNumber="87" value="2018-08-06">
<collectingDate id="F250E9CCFFE6FFCC473CF9C8934A737E" box="[771,895,1609,1635]" pageId="12" pageNumber="87" value="2018-08-06">6.viii.2018</collectingDate>
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, leg.
<collectorName id="3B5F5332FFE6FFCC4785F9C89414737E" box="[954,1057,1609,1635]" pageId="12" pageNumber="87">Mujie Qi</collectorName>
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f;
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<collectingCounty id="7F744E68FFE6FFCC4788F910946973B6" box="[951,1116,1681,1707]" pageId="12" pageNumber="87">Linzhi County</collectingCounty>
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<date id="E2141024FFE6FFCC4058F91094C473B6" box="[1127,1265,1681,1707]" pageId="12" pageNumber="87" value="2018-08-17">
<collectingDate id="F250E9CCFFE6FFCC4058F91094C473B6" box="[1127,1265,1681,1707]" pageId="12" pageNumber="87" value="2018-08-17">17.viii.2018</collectingDate>
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, leg.
<collectorName id="3B5F5332FFE6FFCC4111F91095A273B6" box="[1326,1431,1681,1707]" pageId="12" pageNumber="87">Mujie Qi</collectorName>
, genitalia slide no. LGE18832
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:
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<specimenCount id="80ACFD6DFFE6FFCC46F4F93492C173CD" box="[715,756,1717,1744]" pageId="12" pageNumber="87" type="female">1 ♀</specimenCount>
,
<collectingCounty id="7F744E68FFE6FFCC473DF93494B173D2" box="[770,1156,1717,1743]" pageId="12" pageNumber="87">Qinlangdang Conservation Station</collectingCounty>
(
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,
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),
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</quantity>
,
<collectingMunicipality id="7671AC9EFFE6FFCC4483F958921F73E9" box="[188,554,1753,1780]" pageId="12" pageNumber="87">Gaoligongshan Nature Reserves</collectingMunicipality>
,
<location id="9375603FFFE6FFCC4609F95892A673EE" LSID="urn:lsid:plazi:treatment:1E0387F2FFE6FFCD44A8FAF69154710E:9375603FFFE6FFCC4609F95892A673EE" box="[566,659,1753,1779]" country="China" county="Qinlangdang Conservation Station" latitude="27.69" longLatPrecision="741" longitude="98.27" municipality="Gaoligongshan Nature Reserves" name="Nujiang" pageId="12" pageNumber="87" stateProvince="Yunnan">Nujiang</location>
,
<date id="E2141024FFE6FFCC469FF958932473EE" box="[672,785,1753,1779]" pageId="12" pageNumber="87" value="2017-05-28">
<collectingDate id="F250E9CCFFE6FFCC469FF958932473EE" box="[672,785,1753,1779]" pageId="12" pageNumber="87" value="2017-05-28">28.v.2017</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFE6FFCC4770F95893D473E9" box="[847,993,1753,1780]" pageId="12" pageNumber="87">Kaijian Teng</collectorName>
<emphasis id="A4DEEAF6FFE6FFCC47D6F95B942E73EE" box="[1001,1051,1753,1779]" italics="true" pageId="12" pageNumber="87">et al</emphasis>
., genitalia slide no. LGE18837
</materialsCitation>
;
<materialsCitation id="26C23CB9FFE6FFCC41B1F95892F77221" ID-GBIF-Occurrence="3427696448" accessionNumber="LGE18835" collectingDate="2017-06" collectorName="Kaijian Teng" country="China" county="Gaoligongshan County" elevation="1335" latitude="27.84" location="Dulongjiang Town" longLatPrecision="741" longitude="98.32" municipality="Pukawang Village" pageId="12" pageNumber="87" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFE6FFCC41B1F95890F77205" pageId="12" pageNumber="87" type="female">2 ♀♀</specimenCount>
,
<collectingMunicipality id="7671AC9EFFE6FFCC44F0F97C91AE720A" box="[207,411,1789,1815]" pageId="12" pageNumber="87">Pukawang Village</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFE6FFCC4596F97C9239720A" box="[425,524,1789,1815]" degrees="27.84" direction="north" orientation="latitude" pageId="12" pageNumber="87" precision="555" value="27.84">27.84°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFE6FFCC4629F97C9243720A" box="[534,630,1789,1815]" degrees="98.32" direction="east" orientation="longitude" pageId="12" pageNumber="87" precision="555" value="98.32">98.32°E</geoCoordinate>
),
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<elevation id="1D87D1D7FFE6FFCC46B6F97C92EA7205" box="[649,735,1789,1816]" metricMagnitude="3" metricUnit="m" metricValue="1.335" pageId="12" pageNumber="87" unit="m" value="1335.0">1335 m</elevation>
</quantity>
,
<location id="9375603FFFE6FFCC46D4F97C938A7205" LSID="urn:lsid:plazi:treatment:1E0387F2FFE6FFCD44A8FAF69154710E:9375603FFFE6FFCC46D4F97C938A7205" box="[747,959,1789,1816]" country="China" county="Gaoligongshan County" latitude="27.84" longLatPrecision="741" longitude="98.32" municipality="Pukawang Village" name="Dulongjiang Town" pageId="12" pageNumber="87" stateProvince="Yunnan Province">Dulongjiang Town</location>
,
<collectingCounty id="7F744E68FFE6FFCC47F5F97C94FB720A" box="[970,1230,1789,1815]" pageId="12" pageNumber="87">Gaoligongshan County</collectingCounty>
, 7,12.
<date id="E2141024FFE6FFCC412DF97C955D720A" box="[1298,1384,1789,1815]" pageId="12" pageNumber="87" value="2017-06">
<collectingDate id="F250E9CCFFE6FFCC412DF97C955D720A" box="[1298,1384,1789,1815]" pageId="12" pageNumber="87" value="2017-06">vi.2017</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFE6FFCC44A8F8A0911D7221" box="[151,296,1825,1852]" pageId="12" pageNumber="87">Kaijian Teng</collectorName>
<emphasis id="A4DEEAF6FFE6FFCC4510F8A391557226" box="[303,352,1825,1851]" italics="true" pageId="12" pageNumber="87">et al</emphasis>
., genitalia slide no. LGE18835
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE6FFCC44F8F8C4947472F2" pageId="12" pageNumber="87" type="diagnosis">
<paragraph id="961536E4FFE6FFCC44F8F8C4947472F2" blockId="12.[151,1437,1537,2032]" pageId="12" pageNumber="87">
<emphasis id="A4DEEAF6FFE6FFCC44F8F8C491777242" bold="true" box="[199,322,1861,1887]" pageId="12" pageNumber="87">Diagnosis.</emphasis>
The species can be distinguished from congeners by the dark fuscous forewing with the distal half strongly mixed with brownish grey. The male genitalia of
<taxonomicName id="51AA4D67FFE6FFCC4713F8EB9385729E" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[812,944,1897,1923]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="fuscalata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE6FFCC4713F8EB9385729E" box="[812,944,1897,1923]" italics="true" pageId="12" pageNumber="87">T. fuscalata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE6FFCC4787F8E89426729E" bold="true" box="[952,1043,1897,1923]" pageId="12" pageNumber="87">
<taxonomicNameLabel id="BFED578DFFE6FFCC4787F8E89426729E" box="[952,1043,1897,1923]" pageId="12" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are most similar to those of
<taxonomicName id="51AA4D67FFE6FFCC4165F8EB90CA72BA" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE6FFCC4165F8EB90CA72BA" italics="true" pageId="12" pageNumber="87">T. immaculata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE6FFCC4538F80C915772BA" bold="true" box="[263,354,1933,1959]" pageId="12" pageNumber="87">
<taxonomicNameLabel id="BFED578DFFE6FFCC4538F80C915772BA" box="[263,354,1933,1959]" pageId="12" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but the uncus is broader and there are fewer tegumen tubercles than in the latter species. Also, the anellus lobes of
<taxonomicName id="51AA4D67FFE6FFCC4545F83391C872D6" authorityName="Lee &amp; Li" authorityYear="2022" box="[378,509,1969,1995]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="fuscalata">
<emphasis id="A4DEEAF6FFE6FFCC4545F83391C872D6" box="[378,509,1969,1995]" italics="true" pageId="12" pageNumber="87">T. fuscalata</emphasis>
</taxonomicName>
are fusiform, constricted basally and apically, whereas those of
<taxonomicName id="51AA4D67FFE6FFCC40ECF833954672D6" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1235,1395,1969,1995]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE6FFCC40ECF833954672D6" box="[1235,1395,1969,1995]" italics="true" pageId="12" pageNumber="87">T. immaculata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE6FFCC4144F83090FD72F2" bold="true" pageId="12" pageNumber="87">
<taxonomicNameLabel id="BFED578DFFE6FFCC4144F83090FD72F2" pageId="12" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are ovate. In the female genitalia, the boomerang-shaped signum is distinctive.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE7FFCD44F8FF17935B76B6" pageId="13" pageNumber="88" type="description">
<paragraph id="961536E4FFE7FFCD44F8FF1794EA75EB" blockId="13.[151,1437,150,1043]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8FF17916C75AD" bold="true" box="[199,345,150,176]" pageId="13" pageNumber="88">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFE7FFCD458FFF16923175AC" box="[432,516,151,177]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="13" pageNumber="88">Fig. 49</figureCitation>
).
<emphasis id="A4DEEAF6FFE7FFCD4627FF16926075AC" box="[536,597,151,177]" italics="true" pageId="13" pageNumber="88">Head</emphasis>
: Vertex pale greyish brown, frons grey to dark fuscous. Labial palpus pale greyish brown except palpomere III fuscous. Antenna with scape pale greyish brown to dark fuscous; flagellum dark fuscous, sometimes with dorsal surface slightly suffused with grey, male cilia as long as diameter.
</paragraph>
<paragraph id="961536E4FFE7FFCD44F8FF7E937874F4" blockId="13.[151,1437,150,1043]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8FF7E912D7404" box="[199,280,255,281]" italics="true" pageId="13" pageNumber="88">Thorax</emphasis>
: Dark fuscous, except posterior half pale brown. Tegula dark fuscous, sometimes tinged with pale brown posteriorly. Wingspan 7.5–10.0 mm; forewing with basal 1/2 dark fuscous to black, near base usually pale greyish brown below fold; distal 1/2 pale greyish brown, strongly mixed with fuscous toward apex, with two indistinct lines at anterior 1/3 of wing and on fold, respectively, both merged with fuscous suffusion near apex; fringe scales dark grey to dark fuscous. Hindwing and fringe scales dark fuscous. Legs pale greyish brown; fore coxa sometimes strongly mixed with grey; mid-tibia dark grey along dorsal margin, with a fuscous spot at distal 1/3 of outer surface; hind tibia broadly dark fuscous at base, at middle and at apex.
</paragraph>
<paragraph id="961536E4FFE7FFCD44F8FE70935B76B6" blockId="13.[151,1437,150,1043]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8FE7091047716" box="[199,305,497,523]" italics="true" pageId="13" pageNumber="88">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFE7FFCD457FFE7091B37711" box="[320,390,497,524]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="13" pageNumber="88">Fig. 7</figureCitation>
): Male coremata slightly shorter than sternum VIII; sternum III–VI well-sclerotized, sternum VIII rounded. Male genitalia (
<figureCitation id="0E912A61FFE7FFCD45CEFD9592707732" box="[497,581,532,559]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="13" pageNumber="88">Fig. 68</figureCitation>
) with uncus short, sub-pentagonal, with a medial ridge on ventral surface. Gnathos hook short, sharply pointed apically. Tegumen approximately 6 times as long as uncus; with a bundle of tubercles situated at middle of ventral surface, each tubercle ampulliform (
<figureCitation id="0E912A61FFE7FFCD47DBFDD8940F776E" box="[996,1082,601,627]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="13" pageNumber="88">Fig. 29</figureCitation>
). Valva slender, slightly curved inward, spatulate distally.Anellus lobe somewhat fusiform, approximately 1/5 length of valva; apical bristle slightly shorter than anellus lobe. Juxta with inner margin broadly convex at middle. Vinculum broad, posterior margin convex. Saccus short and broad, sub-triangular. Aedeagus with basal 2/5 dilated, slender and nearly straight distally. Female genitalia (
<figureCitation id="0E912A61FFE7FFCD455CFD65918077E2" box="[355,437,740,767]" captionStart="FIGURES 84–86" captionStartId="18.[151,264,1943,1967]" captionTargetBox="[351,1240,197,1904]" captionTargetId="figure-22@18.[327,1260,181,1920]" captionTargetPageId="18" captionText="FIGURES 84–86. Genitalia of Tenupalpa spp. 84. T. dissimilis sp. nov., male, holotype, slide no. LGE19007; 85. T. angustella, female, slide no. LGE17065; 86. T. fuscalata sp. nov., female, paratype, slide no. LGE18832 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819830" httpUri="https://zenodo.org/record/5819830/files/figure.png" pageId="13" pageNumber="88">Fig. 86</figureCitation>
) with papillae anales simple and setose. Apophyses anteriores approximately 1/2 length of apophyses posteriores. Tergum VIII divided into two sclerites; sternum VIII nearly straight posteriorly, broadly convex anteriorly, weakly sclerotized near ostium bursae and near posterior margin. Ostium bursae near anterior margin of sternum VIII. Ductus bursae slender, broadening toward corpus bursae, ductus seminalis arising from near base. Corpus bursae elongate-ovate, about as long as ductus bursae; signum long, slender, boomerang-shaped or elongate triangular, situated in posterior half of corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE7FFCD44F8FC35924A76D2" box="[199,639,948,975]" pageId="13" pageNumber="88" type="distribution">
<paragraph id="961536E4FFE7FFCD44F8FC35924A76D2" blockId="13.[151,1437,150,1043]" box="[199,639,948,975]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8FC35916C76D3" bold="true" box="[199,345,948,974]" pageId="13" pageNumber="88">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFE7FFCD4558FC35919F76D3" box="[359,426,948,974]" name="China" pageId="13" pageNumber="88">China</collectingCountry>
(
<collectingRegion id="546EF806FFE7FFCD4586FC35923B76D3" box="[441,526,948,974]" country="China" name="Xizang" pageId="13" pageNumber="88">Xizang</collectingRegion>
,
<collectingRegion id="546EF806FFE7FFCD4627FC34924676D2" box="[536,627,949,975]" country="China" name="Yunnan" pageId="13" pageNumber="88">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE7FFCD44F8FC569154710E" pageId="13" pageNumber="88" type="etymology">
<paragraph id="961536E4FFE7FFCD44F8FC569154710E" blockId="13.[151,1437,150,1043]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8FC56917D76EC" bold="true" box="[199,328,983,1009]" pageId="13" pageNumber="88">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFE7FFCD4703FC5693B576EC" box="[828,896,983,1009]" italics="true" pageId="13" pageNumber="88">fusco-</emphasis>
(dark) and
<emphasis id="A4DEEAF6FFE7FFCD403FFC56947176EC" box="[1024,1092,983,1009]" italics="true" pageId="13" pageNumber="88">alatus</emphasis>
(winged), referring to the dark fuscous forewing.
</paragraph>
</subSubSection>
</treatment>
</document>

221
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<mods:title id="5C00F56388CF4AE469514995AEA82D9E">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<mods:name id="61ABCFFE8C8221E7F48911EB494B6763" type="personal">
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<mods:namePart id="ADB3EF67F03C4F5792EDBA883294E817">Lee, Ga-Eun</mods:namePart>
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<mods:affiliation id="8C37EC68CE5C3B3E4DBD9A8C18F11A9A">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China &amp; The Research Institute for Sciences, Jeju National University, Jeju 63243, Korea &amp; gaeunlee 486 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5774 - 5850</mods:affiliation>
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<mods:role id="EA5CEB26B17592CE9A9E333E98787CDF">
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<mods:namePart id="3BE7832EDE9C247CF51F3773551818E8">Li, Houhun</mods:namePart>
<mods:affiliation id="6EE4421B22FB2F8BA789E1872BAF3CBA">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China</mods:affiliation>
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<mods:title id="BFF22DD37DB319722484F3DC1B7EF00C">Zootaxa</mods:title>
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<mods:date id="C45274582035AFE462F592AC00605178">2022</mods:date>
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<mods:number id="9FC56CD0EE4F58BC7557842CBFF48A56">2022-01-05</mods:number>
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<mods:number id="A857A31A0C24ABE5DAAB16FA2A5449DB">5087</mods:number>
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<mods:identifier id="3753FF5AACD8BAB1FF35FD95488896D2" type="CLB-Dataset">2765</mods:identifier>
<mods:identifier id="CC0926828BEED9AA1EDE3DCE834CD160" type="DOI">10.11646/zootaxa.5087.1.4</mods:identifier>
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<treatment id="1E0387F2FFE7FFCE44A8FBE091BD7733" ID-DOI="http://doi.org/10.5281/zenodo.5819828" ID-GBIF-Taxon="191928539" ID-Zenodo-Dep="5819828" LSID="urn:lsid:zoobank.org:act:9FA70607-8700-49D3-BCE8-76AAA5D28EFD" httpUri="http://treatment.plazi.org/id/1E0387F2FFE7FFCE44A8FBE091BD7733" lastPageId="14" lastPageNumber="89" pageId="13" pageNumber="88">
<subSubSection id="DEB0656FFFE7FFCD44A8FBE092BB7166" box="[151,654,1121,1148]" pageId="13" pageNumber="88" type="nomenclature">
<paragraph id="961536E4FFE7FFCD44A8FBE092BB7166" blockId="13.[151,940,1121,1216]" box="[151,654,1121,1148]" pageId="13" pageNumber="88">
<heading id="CD5D8188FFE7FFCD44A8FBE092BB7166" bold="true" box="[151,654,1121,1148]" fontSize="11" level="1" pageId="13" pageNumber="88" reason="1">
<emphasis id="A4DEEAF6FFE7FFCD44A8FBE092BB7166" bold="true" box="[151,654,1121,1148]" pageId="13" pageNumber="88">
<taxonomicName id="51AA4D67FFE7FFCD44A8FBE092167166" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,547,1121,1148]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="88" phylum="Tracheophyta" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE7FFCD44A8FBE091987161" bold="true" box="[151,429,1121,1148]" italics="true" pageId="13" pageNumber="88">Tenupalpa immaculata</emphasis>
Lee
<emphasis id="A4DEEAF6FFE7FFCD45D5FBE391CA7161" bold="true" box="[490,511,1122,1148]" italics="true" pageId="13" pageNumber="88">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFE7FFCD460EFBE092BB7166" box="[561,654,1121,1147]" pageId="13" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="DEB0656FFFE7FFCD44A8FB05939971DD" pageId="13" pageNumber="88" type="description">
<paragraph id="961536E4FFE7FFCD44A8FB0591907183" blockId="13.[151,940,1121,1216]" box="[151,421,1156,1183]" pageId="13" pageNumber="88">
(
<figureCitation id="0E912A61FFE7FFCD44A0FB0590C37183" box="[159,246,1156,1182]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="13" pageNumber="88">Figs 17</figureCitation>
,
<figureCitation id="0E912A61FFE7FFCD453DFB0591157183" box="[258,288,1156,1182]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="13" pageNumber="88">30</figureCitation>
,
<figureCitation id="0E912A61FFE7FFCD4513FB05917F7182" box="[300,330,1156,1183]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="13" pageNumber="88">50</figureCitation>
,
<figureCitation id="0E912A61FFE7FFCD4569FB0591417183" box="[342,372,1156,1182]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="13" pageNumber="88">69</figureCitation>
,
<figureCitation id="0E912A61FFE7FFCD45BFFB0591A87183" box="[384,413,1156,1182]" captionStart="FIGURES 87–90" captionStartId="19.[151,264,1943,1967]" captionTargetBox="[229,1327,198,1905]" captionTargetId="figure-18@19.[196,1391,181,1920]" captionTargetPageId="19" captionText="FIGURES 87–90. Female genitalia of Tenupalpa spp. 87. T. immaculata sp. nov., paratype, slide no. LGE18002; 88. T. isabellina sp. nov., paratype, slide no. LGE17283f; 89. T. flavitermina, slide no. LGE19021; 90. T. biformis, slide no. LLJ15251 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819832" httpUri="https://zenodo.org/record/5819832/files/figure.png" pageId="13" pageNumber="88">87</figureCitation>
)
</paragraph>
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urn:lsid:zoobank.org:act:
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<subSubSection id="DEB0656FFFE7FFCD44A8FB6D92817056" pageId="13" pageNumber="88" type="materials_examined">
<paragraph id="961536E4FFE7FFCD44A8FB6D92817056" blockId="13.[151,1437,1260,2017]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44A8FB6D9171701B" bold="true" box="[151,324,1260,1286]" pageId="13" pageNumber="88">Type material.</emphasis>
<materialsCitation id="26C23CB9FFE7FFCD4575FB6D921B7034" ID-GBIF-Occurrence="3427696330" accessionNumber="LGE18025" collectingDate="2002-06-01" collectorName="Xinpu Wang" country="China" elevation="1700" location="Huixiangping" municipality="Mt. Fanjing" pageId="13" pageNumber="88" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="49118846FFE7FFCD4575FB6D9187701B" box="[330,434,1260,1286]" pageId="13" pageNumber="88" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFE7FFCD4587FB6C91FA701A" box="[440,463,1261,1287]" pageId="13" pageNumber="88" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFE7FFCD45E5FB6D9203701B" box="[474,566,1260,1286]" name="China" pageId="13" pageNumber="88">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFE7FFCD467FFB6D9338701B" box="[576,781,1260,1286]" country="China" name="Guizhou" pageId="13" pageNumber="88">Guizhou Province</collectingRegion>
,
<collectingMunicipality id="7671AC9EFFE7FFCD4728FB6C93AB701B" box="[791,926,1260,1287]" pageId="13" pageNumber="88">Mt. Fanjing</collectingMunicipality>
,
<location id="9375603FFFE7FFCD4797FB6D9470701B" LSID="urn:lsid:plazi:treatment:1E0387F2FFE7FFCE44A8FBE091BD7733:9375603FFFE7FFCD4797FB6D9470701B" box="[936,1093,1260,1286]" country="China" municipality="Mt. Fanjing" name="Huixiangping" pageId="13" pageNumber="88" stateProvince="Guizhou">Huixiangping</location>
,
<quantity id="51529B01FFE7FFCD4070FB6D9490701A" box="[1103,1189,1260,1287]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="88" unit="m" value="1700.0">
<elevation id="1D87D1D7FFE7FFCD4070FB6D9490701A" box="[1103,1189,1260,1287]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="88" unit="m" value="1700.0">1700 m</elevation>
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,
<date id="E2141024FFE7FFCD408FFB6D9529701B" box="[1200,1308,1260,1286]" pageId="13" pageNumber="88" value="2002-06-01">
<collectingDate id="F250E9CCFFE7FFCD408FFB6D9529701B" box="[1200,1308,1260,1286]" pageId="13" pageNumber="88" value="2002-06-01">1.vi.2002</collectingDate>
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, leg.
<collectorName id="3B5F5332FFE7FFCD4169FB6D90EE7034" pageId="13" pageNumber="88">Xinpu Wang</collectorName>
, genitalia slide no. LGE18025
</materialsCitation>
.
<materialsCitation id="26C23CB9FFE7FFCD4608FA8E92857056" ID-GBIF-Occurrence="3427696405" accessionNumber="LGE18002" collectingDate="2002-06-02" collectorName="Xinpu Wang" country="China" elevation="530" location="Mt. Fanjing" municipality="Heiwanhe" pageId="13" pageNumber="88" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="A4DEEAF6FFE7FFCD4608FA8E93A57035" bold="true" box="[567,912,1294,1321]" pageId="13" pageNumber="88">
<typeStatus id="49118846FFE7FFCD4608FA8E92937034" box="[567,678,1295,1321]" pageId="13" pageNumber="88" type="paratype">Paratype</typeStatus>
.
<collectingRegion id="546EF806FFE7FFCD468EFA8F93BE7035" box="[689,907,1294,1320]" country="China" name="Guizhou" pageId="13" pageNumber="88">Guizhou Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFE7FFCD47A9FA8E938B7034" box="[918,958,1295,1321]" pageId="13" pageNumber="88" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="7671AC9EFFE7FFCD47F4FA8E940A7034" box="[971,1087,1295,1321]" pageId="13" pageNumber="88">Heiwanhe</collectingMunicipality>
,
<location id="9375603FFFE7FFCD4076FA8E94E57034" LSID="urn:lsid:plazi:treatment:1E0387F2FFE7FFCE44A8FBE091BD7733:9375603FFFE7FFCD4076FA8E94E57034" box="[1097,1232,1295,1321]" country="China" municipality="Heiwanhe" name="Mt. Fanjing" pageId="13" pageNumber="88" stateProvince="Guizhou">Mt. Fanjing</location>
,
<quantity id="51529B01FFE7FFCD40E6FA8F95147034" box="[1241,1313,1294,1321]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="13" pageNumber="88" unit="m" value="530.0">
<elevation id="1D87D1D7FFE7FFCD40E6FA8F95147034" box="[1241,1313,1294,1321]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="13" pageNumber="88" unit="m" value="530.0">530 m</elevation>
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,
<date id="E2141024FFE7FFCD4114FA8E95A27034" box="[1323,1431,1295,1321]" pageId="13" pageNumber="88" value="2002-06-02">
<collectingDate id="F250E9CCFFE7FFCD4114FA8E95A27034" box="[1323,1431,1295,1321]" pageId="13" pageNumber="88" value="2002-06-02">2.vi.2002</collectingDate>
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, leg.
<collectorName id="3B5F5332FFE7FFCD44F8FAB0916D7051" box="[199,344,1329,1356]" pageId="13" pageNumber="88">Xinpu Wang</collectorName>
, genitalia slide no. LGE18002
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.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE7FFCD44F8FAD5955C70E4" pageId="13" pageNumber="88" type="diagnosis">
<paragraph id="961536E4FFE7FFCD44F8FAD5955C70E4" blockId="13.[151,1437,1260,2017]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8FAD5910A7073" bold="true" box="[199,319,1364,1390]" pageId="13" pageNumber="88">Diagnosis.</emphasis>
The species may be confused with the similar
<taxonomicName id="51AA4D67FFE7FFCD476BFAD493E87073" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[852,989,1364,1390]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="88" phylum="Tracheophyta" rank="species" species="isabellina" status="sp. nov.">
<emphasis id="A4DEEAF6FFE7FFCD476BFAD493E87073" box="[852,989,1364,1390]" italics="true" pageId="13" pageNumber="88">T. isabellina</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE7FFCD47D9FAD594757073" bold="true" box="[998,1088,1364,1390]" pageId="13" pageNumber="88">
<taxonomicNameLabel id="BFED578DFFE7FFCD47D9FAD594757073" box="[998,1088,1364,1390]" pageId="13" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but the forewing coloration is somewhat paler than that of the latter. Diagnostic characters for
<taxonomicName id="51AA4D67FFE7FFCD475FFAF693CB708C" authorityName="Lee et Li" authorityYear="2022" box="[864,1022,1399,1425]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="88" phylum="Tracheophyta" rank="species" species="immaculata">
<emphasis id="A4DEEAF6FFE7FFCD475FFAF693CB708C" box="[864,1022,1399,1425]" italics="true" pageId="13" pageNumber="88">T. immaculata</emphasis>
</taxonomicName>
are present on the legs: the mid-tibia lacks a spot, and the hind tibia is dark fuscous apically. In
<taxonomicName id="51AA4D67FFE7FFCD4727FA1B93AA70AE" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[792,927,1433,1459]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="88" phylum="Tracheophyta" rank="species" species="isabellina" status="sp. nov.">
<emphasis id="A4DEEAF6FFE7FFCD4727FA1B93AA70AE" box="[792,927,1433,1459]" italics="true" pageId="13" pageNumber="88">T. isabellina</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE7FFCD4799FA1893CB70AE" bold="true" box="[934,1022,1433,1459]" pageId="13" pageNumber="88">
<taxonomicNameLabel id="BFED578DFFE7FFCD4799FA1893CB70AE" box="[934,1022,1433,1459]" pageId="13" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, a fuscous spot is present at the distal 1/3 of the mid-tibia, and the apex of the hind tibia is pale orange. The male genitalia of
<taxonomicName id="51AA4D67FFE7FFCD407EFA3C94E870CB" authorityName="Lee et Li" authorityYear="2022" box="[1089,1245,1468,1494]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="88" phylum="Tracheophyta" rank="species" species="immaculata">
<emphasis id="A4DEEAF6FFE7FFCD407EFA3C94E870CB" box="[1089,1245,1468,1494]" italics="true" pageId="13" pageNumber="88">T. immaculata</emphasis>
</taxonomicName>
are differentiated from those of
<taxonomicName id="51AA4D67FFE7FFCD450EFA5E918D70E4" authorityName="Lee &amp; Li" authorityYear="2022" box="[305,440,1503,1529]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="88" phylum="Tracheophyta" rank="species" species="isabellina">
<emphasis id="A4DEEAF6FFE7FFCD450EFA5E918D70E4" box="[305,440,1503,1529]" italics="true" pageId="13" pageNumber="88">T. isabellina</emphasis>
</taxonomicName>
by the shorter uncus, narrower apex of valva, and the anellus lobe distally constricted.
</paragraph>
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<subSubSection id="DEB0656FFFE7FFCE44F8F98093C274DB" lastPageId="14" lastPageNumber="89" pageId="13" pageNumber="88" type="description">
<paragraph id="961536E4FFE7FFCD44F8F9809451739E" blockId="13.[151,1437,1260,2017]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8F980916C7306" bold="true" box="[199,345,1537,1563]" pageId="13" pageNumber="88">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFE7FFCD458CF980923C7301" box="[435,521,1537,1564]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="13" pageNumber="88">Fig. 50</figureCitation>
).
<emphasis id="A4DEEAF6FFE7FFCD4621F980926E7306" box="[542,603,1537,1563]" italics="true" pageId="13" pageNumber="88">Head</emphasis>
: Vertex pale brown to greyish brown, frons creamy white. Labial palpus pale brown; palpomere I tinged with fuscous apically; palpomere III as long as II, strongly mixed with grey.Antenna with scape pale greyish brown except posterior margin tinged with fuscous distally; flagellum dark fuscous, basal half of dorsal surface slightly mixed with creamy white, male cilia as long as diameter.
</paragraph>
<paragraph id="961536E4FFE7FFCD44F8F90D94957249" blockId="13.[151,1437,1260,2017]" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8F90D912D73BB" box="[199,280,1676,1702]" italics="true" pageId="13" pageNumber="88">Thorax</emphasis>
: Pale brown, anterior margin sometimes suffused with fuscous. Tegula pale brown except anterior 3/5 dark fuscous. Wingspan 8.5–9.0 mm; forewing ground color pale greyish brown, grey along costa, base dark fuscous near costa, middle of wing broadly suffused with grey; two indistinct grey lines arising from outer margin of former greyish suffusion: upper line longitudinally running toward apex at anterior 1/3 of wing, lower line running to tornus along fold; fringe scales pale greyish brown. Hindwing grey to pale greyish brown; fringe scales pale greyish brown. Legs creamy white to pale brown; hind tibia broadly fuscous at middle and apex.
</paragraph>
<paragraph id="961536E4FFE7FFCE44F8F8DD93C274DB" blockId="13.[151,1437,1260,2017]" lastBlockId="14.[151,1437,151,559]" lastPageId="14" lastPageNumber="89" pageId="13" pageNumber="88">
<emphasis id="A4DEEAF6FFE7FFCD44F8F8DD9104726B" box="[199,305,1884,1910]" italics="true" pageId="13" pageNumber="88">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFE7FFCD457EF8DD91A1726B" box="[321,404,1884,1910]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="13" pageNumber="88">Fig. 17</figureCitation>
): Male coremata nearly as long as sternum VIII; sternum VIII sub-triangular, lateral margin broadly convex. Male genitalia (
<figureCitation id="0E912A61FFE7FFCD463AF80192627286" box="[517,599,1920,1947]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="13" pageNumber="88">Fig. 69</figureCitation>
) with uncus short, sub-pentagonal, with a medial ridge on ventral surface. Gnathos hook moderately long, sharply pointed apically. Tegumen approximately 6 times as long as uncus; with abundant tubercles situated at middle of ventral surface, each tubercle ampulliform and bearing a long seta (
<figureCitation id="0E912A61FFE7FFCD4102F84695A572FC" box="[1341,1424,1991,2017]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="13" pageNumber="88">Fig. 30</figureCitation>
). Valva slender, slightly broadened at base and in distal 1/3, rounded at apex. Anellus lobe ovate, approximately 1/6 length of valva, sparely setose near apex; apical bristle longer than anellus lobe. Juxta simple, inner margin broadly convex. Vinculum simple, broadly convex on posterior margin. Saccus sub-triangular, slightly exceeding tegumen pedunculus. Aedeagus with basal half globular, distal half slender and straight. Female genitalia (
<figureCitation id="0E912A61FFE4FFCE40E2FF7E951A7404" box="[1245,1327,255,281]" captionStart="FIGURES 87–90" captionStartId="19.[151,264,1943,1967]" captionTargetBox="[229,1327,198,1905]" captionTargetId="figure-18@19.[196,1391,181,1920]" captionTargetPageId="19" captionText="FIGURES 87–90. Female genitalia of Tenupalpa spp. 87. T. immaculata sp. nov., paratype, slide no. LGE18002; 88. T. isabellina sp. nov., paratype, slide no. LGE17283f; 89. T. flavitermina, slide no. LGE19021; 90. T. biformis, slide no. LLJ15251 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819832" httpUri="https://zenodo.org/record/5819832/files/figure.png" pageId="14" pageNumber="89">Fig. 87</figureCitation>
) with papillae anales simple and setose. Apophyses anteriores approximately 1/3 length of apophyses posteriores. Sternum VIII produced anteriorly, with two short sub-triangular flaps covering ostium bursae. Ostium bursae opening near anterior margin of sternum VIII. Ductus bursae narrow at base, gradually widening toward corpus bursae from posterior 1/3; ductus seminalis arising from posterior 1/3 of ductus bursae. Corpus bursae ovate, abruptly bulged at base; signum small, crescent-shaped, situated near entrance of ductus bursae.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE4FFCE44F8FE4F921974F7" box="[199,556,462,490]" pageId="14" pageNumber="89" type="distribution">
<paragraph id="961536E4FFE4FFCE44F8FE4F921974F7" blockId="14.[151,1437,151,559]" box="[199,556,462,490]" pageId="14" pageNumber="89">
<emphasis id="A4DEEAF6FFE4FFCE44F8FE4F916C74F5" bold="true" box="[199,345,462,488]" pageId="14" pageNumber="89">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFE4FFCE4558FE4E919F74F4" box="[359,426,463,489]" name="China" pageId="14" pageNumber="89">China</collectingCountry>
(
<collectingRegion id="546EF806FFE4FFCE4586FE4E921574F7" box="[441,544,463,490]" country="China" name="Guizhou" pageId="14" pageNumber="89">Guizhou</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE4FFCE44F8FE7091BD7733" pageId="14" pageNumber="89" type="etymology">
<paragraph id="961536E4FFE4FFCE44F8FE7091BD7733" blockId="14.[151,1437,151,559]" pageId="14" pageNumber="89">
<emphasis id="A4DEEAF6FFE4FFCE44F8FE70917D7716" bold="true" box="[199,328,497,523]" pageId="14" pageNumber="89">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFE4FFCE4774FE7093EF7716" box="[843,986,497,523]" italics="true" pageId="14" pageNumber="89">immaculatus</emphasis>
(spotless), referring to the absence of a spot on the mid-tibia.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="4B3F088FE5E47644C905DC4DCA79994D">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<paragraph id="961536E4FFE8FFC244A8FD7A93907609" blockId="2.[151,933,762,789]" box="[151,933,762,789]" pageId="2" pageNumber="77">
<heading id="CD5D8188FFE8FFC244A8FD7A93907609" bold="true" box="[151,933,762,789]" fontSize="11" level="1" pageId="2" pageNumber="77" reason="1">
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Key to species of
<taxonomicName id="51AA4D67FFE8FFC24551FD7B91DB7608" authorityName="Lee et Li" authorityYear="2021" box="[366,494,762,789]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFE8FFC24551FD7B91DB7608" bold="true" box="[366,494,762,789]" italics="true" pageId="2" pageNumber="77">Tenupalpa</emphasis>
</taxonomicName>
based on adults and male genitalia
</emphasis>
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</subSubSection>
<subSubSection id="DEB0656FFFE8FFC244A8FCC195A972F7" pageId="2" pageNumber="77" type="key">
<keyStep id="2D5E2E41FFE8FFC244A8FCC195A9766F" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FCC195A87648" blockId="2.[151,1437,832,2027]" box="[151,1437,832,854]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FCC195A87648" box="[151,1437,832,854]" pageId="2" pageNumber="77">
1. Forewing with black and yellow longitudinal streaks in basal 1/3.....................................
<taxonomicName id="51AA4D67FFE8FFC24129FCC195A87648" ID-CoL="93L49" authorityName="Meyrick" authorityYear="1923" box="[1302,1437,832,853]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="xanthodora">
<emphasis id="A4DEEAF6FFE8FFC24129FCC195A87648" box="[1302,1437,832,853]" italics="true" pageId="2" pageNumber="77">T. xanthodora</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FCDC95A9766F" blockId="2.[151,1437,832,2027]" box="[151,1436,861,883]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FCDC95A9766F" box="[151,1436,861,883]" pageId="2" pageNumber="77">- Forewing without black and yellow streaks in basal 1/3....................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FCFB95A976B0" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FCFB95A97692" blockId="2.[151,1437,832,2027]" box="[151,1436,890,912]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FCFB95A97692" box="[151,1436,890,912]" pageId="2" pageNumber="77">2. Forewing without an orange distal patch; hindwing uniformly grey to fuscous..................................... 3</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FC1995A976B0" blockId="2.[151,1437,832,2027]" box="[151,1436,920,942]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FC1995A976B0" box="[151,1436,920,942]" pageId="2" pageNumber="77">- Forewing with a distinctive orange distal patch; hindwing with apex tinged with black, subapex white.................. 9</keyLead>
</paragraph>
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<keyStep id="2D5E2E41FFE8FFC244A8FC3495A976F5" pageId="2" pageNumber="77">
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<keyLead id="2D5B95D1FFE8FFC244A8FC3495A976D7" box="[151,1436,949,971]" pageId="2" pageNumber="77">
3.
<collectingRegion id="546EF806FFE8FFC244DDFC34912C76D7" box="[226,281,949,970]" country="New Zealand" name="West Coast" pageId="2" pageNumber="77">Costa</collectingRegion>
of valva with a projection at middle.................................................................. 4
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FC5295A976F5" blockId="2.[151,1437,832,2027]" box="[151,1436,978,1000]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FC5295A976F5" box="[151,1436,978,1000]" pageId="2" pageNumber="77">
-
<collectingRegion id="546EF806FFE8FFC244DDFC53912C76FA" box="[226,281,978,999]" country="New Zealand" name="West Coast" pageId="2" pageNumber="77">Costa</collectingRegion>
of valva without a projection....................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FC7195A9713F" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FC7195A97118" blockId="2.[151,1437,832,2027]" box="[151,1436,1008,1030]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FC7195A97118" box="[151,1436,1008,1030]" pageId="2" pageNumber="77">
4. Forewing darkened apically; valva with an elongate-triangular projection at middle of costa.........
<taxonomicName id="51AA4D67FFE8FFC240F9FC71957F7118" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1222,1354,1008,1029]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="latifasciata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240F9FC71957F7118" box="[1222,1354,1008,1029]" italics="true" pageId="2" pageNumber="77">T. latifasciata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FFC7195A97118" bold="true" box="[1360,1436,1008,1029]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FFC7195A97118" box="[1360,1436,1008,1029]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FB8C95A9713F" blockId="2.[151,1437,832,2027]" box="[151,1436,1037,1059]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FB8C95A9713F" box="[151,1436,1037,1059]" pageId="2" pageNumber="77">
- Forewing not darkened apically; valva with a trapezoidal projection at middle of costa...................
<taxonomicName id="51AA4D67FFE8FFC24132FB8F95A9713F" baseAuthorityName="Meyrick" baseAuthorityYear="1918" box="[1293,1436,1037,1059]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFE8FFC24132FB8F95A9713F" box="[1293,1436,1037,1059]" italics="true" pageId="2" pageNumber="77">T. nephodesma</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FBAB95A97143" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FBAB95A97122" blockId="2.[151,1437,832,2027]" box="[151,1436,1066,1088]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FBAB95A97122" box="[151,1436,1066,1088]" pageId="2" pageNumber="77">
5. Anellus lobe with a robust apical bristle (or spine), almost reaching apex of valva.........................
<taxonomicName id="51AA4D67FFE8FFC2411EFBAA95A97122" ID-CoL="98YHR" baseAuthorityName="Omelko" baseAuthorityYear="1984" box="[1313,1436,1066,1088]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="angustella">
<emphasis id="A4DEEAF6FFE8FFC2411EFBAA95A97122" box="[1313,1436,1066,1088]" italics="true" pageId="2" pageNumber="77">T. angustella</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FBC995A97143" blockId="2.[151,1437,832,2027]" box="[151,1436,1096,1118]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FBC995A97143" box="[151,1436,1096,1118]" pageId="2" pageNumber="77">- Anellus lobe without a robust apical bristle................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FBE495A97185" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FBE495A97167" blockId="2.[151,1437,832,2027]" box="[151,1436,1125,1147]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FBE495A97167" box="[151,1436,1125,1147]" pageId="2" pageNumber="77">
6. Junction of valva and juxta strongly angled; base of valva strongly curved inward...................
<taxonomicName id="51AA4D67FFE8FFC240E5FBE7957F7167" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1242,1354,1125,1147]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="angulosa" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240E5FBE7957F7167" box="[1242,1354,1125,1147]" italics="true" pageId="2" pageNumber="77">T. angulosa</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FFBE495A97167" bold="true" box="[1360,1436,1125,1146]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FFBE495A97167" box="[1360,1436,1125,1146]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FB0295A97185" blockId="2.[151,1437,832,2027]" box="[151,1436,1154,1176]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FB0295A97185" box="[151,1436,1154,1176]" pageId="2" pageNumber="77">- Junction of valva and juxta not strongly angled; based of valva slightly or not curved inward......................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FB2195A971CF" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FB2195A971A8" blockId="2.[151,1437,832,2027]" box="[151,1436,1184,1206]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FB2195A971A8" box="[151,1436,1184,1206]" pageId="2" pageNumber="77">
7. Uncus widest at base, elongated distally...................................................
<taxonomicName id="51AA4D67FFE8FFC240ECFB21957F71A8" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1235,1354,1184,1205]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="isabellina" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240ECFB21957F71A8" box="[1235,1354,1184,1205]" italics="true" pageId="2" pageNumber="77">T. isabellina</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FFB2195A971A8" bold="true" box="[1360,1436,1184,1205]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FFB2195A971A8" box="[1360,1436,1184,1205]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FB3C95A971CF" blockId="2.[151,1437,832,2027]" box="[151,1436,1213,1235]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FB3C95A971CF" box="[151,1436,1213,1235]" pageId="2" pageNumber="77">- Uncus widest at middle, not elongated distally.............................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FB5B95A97010" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FB5B95A971F2" blockId="2.[151,1437,832,2027]" box="[151,1436,1242,1264]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FB5B95A971F2" box="[151,1436,1242,1264]" pageId="2" pageNumber="77">
8. Sternum VIII rounded; anellus lobe fusiform.................................................
<taxonomicName id="51AA4D67FFE8FFC240E5FB5A957F71F2" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1242,1354,1242,1264]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="fuscalata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240E5FB5A957F71F2" box="[1242,1354,1242,1264]" italics="true" pageId="2" pageNumber="77">T. fuscalata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FFB5B95A971F2" bold="true" box="[1360,1436,1242,1263]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FFB5B95A971F2" box="[1360,1436,1242,1263]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FB7995A97010" blockId="2.[151,1437,832,2027]" box="[151,1436,1272,1294]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FB7995A97010" box="[151,1436,1272,1294]" pageId="2" pageNumber="77">
- Sternum VIII sub-triangular; anellus lobe ovate............................................
<taxonomicName id="51AA4D67FFE8FFC240FEFB79957C7010" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1217,1353,1272,1293]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240FEFB79957C7010" box="[1217,1353,1272,1293]" italics="true" pageId="2" pageNumber="77">T. immaculata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FFB7995A97010" bold="true" box="[1360,1436,1272,1293]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FFB7995A97010" box="[1360,1436,1272,1293]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FA9495A9705A" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FA9495A97037" blockId="2.[151,1437,832,2027]" box="[151,1436,1301,1323]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FA9495A97037" box="[151,1436,1301,1323]" pageId="2" pageNumber="77">9. Forewing apical spot absent............................................................................ 10</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FAB295A9705A" blockId="2.[151,1437,832,2027]" box="[151,1436,1330,1352]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FAB295A9705A" box="[151,1436,1330,1352]" pageId="2" pageNumber="77">- Forewing apical spot present........................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FAD195A9709F" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FAD195A97078" blockId="2.[151,1437,832,2027]" box="[151,1436,1360,1382]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FAD195A97078" box="[151,1436,1360,1382]" pageId="2" pageNumber="77">
10. Valva not spatulate distally; anellus lobe oval with a long apical bristle................................
<taxonomicName id="51AA4D67FFE8FFC24130FAD195A97078" ID-CoL="98YHN" baseAuthorityName="Kyaw, Yagi &amp; Hirowatari" baseAuthorityYear="2019" box="[1295,1436,1360,1381]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFE8FFC24130FAD195A97078" box="[1295,1436,1360,1381]" italics="true" pageId="2" pageNumber="77">T. flavitermina</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FAEC95A9709F" blockId="2.[151,1437,832,2027]" box="[151,1436,1389,1411]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FAEC95A9709F" box="[151,1436,1389,1411]" pageId="2" pageNumber="77">
- Valva spatulate distally; anellus lobe falcate with a short subapical bristle............................
<taxonomicName id="51AA4D67FFE8FFC240D0FAEF957F709F" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1263,1354,1389,1411]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="falcata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240D0FAEF957F709F" box="[1263,1354,1389,1411]" italics="true" pageId="2" pageNumber="77">T. falcata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FFAEC95A9709F" bold="true" box="[1360,1436,1389,1410]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FFAEC95A9709F" box="[1360,1436,1389,1410]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FA0B95A970A0" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FA0B95A97082" blockId="2.[151,1437,832,2027]" box="[151,1436,1418,1440]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FA0B95A97082" box="[151,1436,1418,1440]" pageId="2" pageNumber="77">
11. Valva acute apically..................................................................
<taxonomicName id="51AA4D67FFE8FFC240FDFA0A957F7082" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1218,1354,1418,1440]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="uncativalva" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240FDFA0A957F7082" box="[1218,1354,1418,1440]" italics="true" pageId="2" pageNumber="77">T. uncativalva</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FFA0B95A97082" bold="true" box="[1360,1436,1418,1439]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FFA0B95A97082" box="[1360,1436,1418,1439]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FA2995A970A0" blockId="2.[151,1437,832,2027]" box="[151,1436,1448,1470]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FA2995A970A0" box="[151,1436,1448,1470]" pageId="2" pageNumber="77">- Valva rounded apically................................................................................ 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8FA4495A970EA" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8FA4495A970C7" blockId="2.[151,1437,832,2027]" box="[151,1436,1477,1499]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FA4495A970C7" box="[151,1436,1477,1499]" pageId="2" pageNumber="77">
12. Anellus lobes asymmetrical..............................................................
<taxonomicName id="51AA4D67FFE8FFC240E8FA47957F70C7" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1239,1354,1477,1499]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="dissimilis" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240E8FA47957F70C7" box="[1239,1354,1477,1499]" italics="true" pageId="2" pageNumber="77">T. dissimilis</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FFA4495A970C7" bold="true" box="[1360,1436,1477,1498]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FFA4495A970C7" box="[1360,1436,1477,1498]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8FA6295A970EA" blockId="2.[151,1437,832,2027]" box="[151,1436,1506,1528]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8FA6295A970EA" box="[151,1436,1506,1528]" pageId="2" pageNumber="77">- Anellus lobes symmetrical............................................................................. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8F98195A9732F" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8F98195A97308" blockId="2.[151,1437,832,2027]" box="[151,1436,1536,1558]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F98195A97308" box="[151,1436,1536,1558]" pageId="2" pageNumber="77">
13. Anellus lobe linear, exceeding apex of valva.......................................................
<taxonomicName id="51AA4D67FFE8FFC2411AF98195A97308" ID-CoL="98YJC" baseAuthorityName="Omelko" baseAuthorityYear="1984" box="[1317,1436,1536,1557]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="venustalis">
<emphasis id="A4DEEAF6FFE8FFC2411AF98195A97308" box="[1317,1436,1536,1557]" italics="true" pageId="2" pageNumber="77">T. venustalis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8F99C95A9732F" blockId="2.[151,1437,832,2027]" box="[151,1436,1565,1587]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F99C95A9732F" box="[151,1436,1565,1587]" pageId="2" pageNumber="77">- Anellus lobe not linear, not exceeding apex of valva......................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8F9BB95A97370" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8F9BB95A97352" blockId="2.[151,1437,832,2027]" box="[151,1436,1594,1616]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F9BB95A97352" box="[151,1436,1594,1616]" pageId="2" pageNumber="77">
14. Anellus lobe without an apical or subapical bristle.........................................
<taxonomicName id="51AA4D67FFE8FFC2408AF9BA957C7352" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1205,1353,1594,1616]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="perparviloba" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC2408AF9BA957C7352" box="[1205,1353,1594,1616]" italics="true" pageId="2" pageNumber="77">T. perparviloba</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FF9BB95A97352" bold="true" box="[1360,1436,1594,1615]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FF9BB95A97352" box="[1360,1436,1594,1615]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8F9D995A97370" blockId="2.[151,1437,832,2027]" box="[151,1436,1623,1646]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F9D995A97370" box="[151,1436,1623,1646]" pageId="2" pageNumber="77">- Anellus lobe with an apical or subapical bristle............................................................. 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8F9F495A973B5" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8F9F495A97397" blockId="2.[151,1437,832,2027]" box="[151,1436,1653,1675]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F9F495A97397" box="[151,1436,1653,1675]" pageId="2" pageNumber="77">
15. Anellus lobe digitate, curved inward..............................................................
<taxonomicName id="51AA4D67FFE8FFC2410AF9F795A97397" ID-CoL="98YHZ" baseAuthorityName="Omelko" baseAuthorityYear="1984" box="[1333,1436,1653,1675]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFE8FFC2410AF9F795A97397" box="[1333,1436,1653,1675]" italics="true" pageId="2" pageNumber="77">T. biformis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8F91295A973B5" blockId="2.[151,1437,832,2027]" box="[151,1436,1682,1704]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F91295A973B5" box="[151,1436,1682,1704]" pageId="2" pageNumber="77">- Anellus lobe round or oval, not curved................................................................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8F93195A973E2" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8F93195A973FF" blockId="2.[151,1437,832,2027]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F93195A973FF" pageId="2" pageNumber="77">
16. Distal margin of uncus deeply concave; anellus lobe with a strongly developed, horn-like apical bristle; juxta with a large digitate projection...........................................................................
<taxonomicName id="51AA4D67FFE8FFC240CCF94F957F73FF" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1267,1354,1741,1763]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="zhengi" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240CCF94F957F73FF" box="[1267,1354,1741,1763]" italics="true" pageId="2" pageNumber="77">T. zhengi</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FF94C95A973FF" bold="true" box="[1360,1436,1741,1762]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FF94C95A973FF" box="[1360,1436,1741,1762]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8F96A95A973E2" blockId="2.[151,1437,832,2027]" box="[151,1436,1770,1792]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F96A95A973E2" box="[151,1436,1770,1792]" pageId="2" pageNumber="77">- Uncus not concave distally; anellus lobe with a hair-like apical bristle; juxta without a digitate projection.............. 17</keyLead>
</paragraph>
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<paragraph id="961536E4FFE8FFC244A8F88995A97200" blockId="2.[151,1437,832,2027]" box="[151,1436,1800,1822]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F88995A97200" box="[151,1436,1800,1822]" pageId="2" pageNumber="77">
17.
<collectingRegion id="546EF806FFE8FFC244DDF889912C7200" box="[226,281,1800,1821]" country="New Zealand" name="West Coast" pageId="2" pageNumber="77">Costa</collectingRegion>
of valva acute medially...........................................................
<taxonomicName id="51AA4D67FFE8FFC240F8F889957F7200" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1223,1354,1800,1821]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="acutimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240F8F889957F7200" box="[1223,1354,1800,1821]" italics="true" pageId="2" pageNumber="77">T. acutimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FF88995A97200" bold="true" box="[1360,1436,1800,1821]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FF88995A97200" box="[1360,1436,1800,1821]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="961536E4FFE8FFC244A8F8A495A97227" blockId="2.[151,1437,832,2027]" box="[151,1436,1829,1851]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F8A495A97227" box="[151,1436,1829,1851]" pageId="2" pageNumber="77">
-
<collectingRegion id="546EF806FFE8FFC244DDF8A4912C7227" box="[226,281,1829,1850]" country="New Zealand" name="West Coast" pageId="2" pageNumber="77">Costa</collectingRegion>
of valva not acute medially....................................................................... 18
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<paragraph id="961536E4FFE8FFC244A8F8C395A9724A" blockId="2.[151,1437,832,2027]" box="[151,1436,1858,1880]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F8C395A9724A" box="[151,1436,1858,1880]" pageId="2" pageNumber="77">
18.
<collectingRegion id="546EF806FFE8FFC244DDF8C3912C724A" box="[226,281,1858,1879]" country="New Zealand" name="West Coast" pageId="2" pageNumber="77">Costa</collectingRegion>
of valva broadly convex medially.................................................
<taxonomicName id="51AA4D67FFE8FFC2408DF8C2957F7245" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1202,1354,1859,1880]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="magniguttata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC2408DF8C2957F7245" box="[1202,1354,1859,1880]" italics="true" pageId="2" pageNumber="77">T. magniguttata</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="BFED578DFFE8FFC2416FF8C395A9724A" box="[1360,1436,1858,1879]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="961536E4FFE8FFC244A8F8E195A97268" blockId="2.[151,1437,832,2027]" box="[151,1436,1888,1910]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F8E195A97268" box="[151,1436,1888,1910]" pageId="2" pageNumber="77">
-
<collectingRegion id="546EF806FFE8FFC244DDF8E1912C7268" box="[226,281,1888,1909]" country="New Zealand" name="West Coast" pageId="2" pageNumber="77">Costa</collectingRegion>
of valva not convex medially...................................................................... 19
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<paragraph id="961536E4FFE8FFC244A8F8FC95A9728F" blockId="2.[151,1437,832,2027]" box="[151,1436,1917,1939]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F8FC95A9728F" box="[151,1436,1917,1939]" pageId="2" pageNumber="77">
19. Apex of valva usually with a short projection; aedeagus acute at apex..............................
<taxonomicName id="51AA4D67FFE8FFC240D5F8FF957F728E" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1258,1354,1918,1939]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC240D5F8FF957F728E" box="[1258,1354,1918,1939]" italics="true" pageId="2" pageNumber="77">T. acutata</emphasis>
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<taxonomicNameLabel id="BFED578DFFE8FFC2416FF8FC95A9728F" box="[1360,1436,1917,1938]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="961536E4FFE8FFC244A8F81A95A972B2" blockId="2.[151,1437,832,2027]" box="[151,1436,1946,1968]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F81A95A972B2" box="[151,1436,1946,1968]" pageId="2" pageNumber="77">- Apex of valva without a projection; aedeagus not acute at apex................................................ 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="2D5E2E41FFE8FFC244A8F83995A972F7" pageId="2" pageNumber="77">
<paragraph id="961536E4FFE8FFC244A8F83995A972D0" blockId="2.[151,1437,832,2027]" box="[151,1436,1975,1998]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F83995A972D0" box="[151,1436,1975,1998]" pageId="2" pageNumber="77">
20. Gnathos thickened medially; juxta with a short projection; aedeagus with basal 2/5 dilated..........
<taxonomicName id="51AA4D67FFE8FFC24081F839957F72D0" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1214,1354,1976,1997]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="crassimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC24081F839957F72D0" box="[1214,1354,1976,1997]" italics="true" pageId="2" pageNumber="77">T. crassimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE8FFC2416FF83995A972D0" bold="true" box="[1360,1436,1976,1997]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FF83995A972D0" box="[1360,1436,1976,1997]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="961536E4FFE8FFC244A8F85495A972F7" blockId="2.[151,1437,832,2027]" box="[151,1436,2005,2027]" pageId="2" pageNumber="77">
<keyLead id="2D5B95D1FFE8FFC244A8F85495A972F7" box="[151,1436,2005,2027]" pageId="2" pageNumber="77">
- Gnathos not thickened medially; juxta without a projection; aedeagus with basal half dilated.......
<taxonomicName id="51AA4D67FFE8FFC2408FF857957F72F7" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1200,1354,2005,2027]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="species" species="breviaedeaga" status="sp. nov.">
<emphasis id="A4DEEAF6FFE8FFC2408FF857957F72F7" box="[1200,1354,2005,2027]" italics="true" pageId="2" pageNumber="77">T. breviaedeaga</emphasis>
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<emphasis id="A4DEEAF6FFE8FFC2416FF85495A972F7" bold="true" box="[1360,1436,2005,2026]" pageId="2" pageNumber="77">
<taxonomicNameLabel id="BFED578DFFE8FFC2416FF85495A972F7" box="[1360,1436,2005,2026]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
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<mods:title id="0656E3D5B661C4D9AC1C85716C5D549C">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<emphasis id="A4DEEAF6FFEBFFC144A8FA979122702C" bold="true" box="[151,279,1302,1329]" italics="true" pageId="1" pageNumber="76">Tenupalpa</emphasis>
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Lee
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Li, 2021
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Lee
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species:
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<emphasis id="A4DEEAF6FFEBFFC1464EFADA937E706E" box="[625,843,1371,1395]" italics="true" pageId="1" pageNumber="76">Tenupalpa angustella</emphasis>
(
<bibRefCitation id="F23B4B15FFEBFFC14767FADA93DA706E" author="Omelko, M. M." box="[856,1007,1371,1395]" pageId="1" pageNumber="76" pagination="28 - 34" refId="ref23940" refString="Omelko, M. M. (1984) Description of three new species of Gelechiidae from South Primorye. The Fauna and Ecology of Insects in the South of the Far East, 112 (215), 28 - 34. [in Russian]" type="journal article" year="1984">Omelko, 1984</bibRefCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="961536E4FFEBFFC144A8FA209146731E" blockId="1.[151,1437,1441,2008]" pageId="1" pageNumber="76">
<emphasis id="A4DEEAF6FFEBFFC144A8FA2091B070A6" bold="true" box="[151,389,1441,1467]" pageId="1" pageNumber="76">Generic characters.</emphasis>
<emphasis id="A4DEEAF6FFEBFFC145B0FA2091F970A6" box="[399,460,1441,1467]" italics="true" pageId="1" pageNumber="76">Head</emphasis>
: With glossy scales. Maxillary palpus 4–articled. Labial palpus slender, strongly recurved, palpomere I very short, palpomere II usually as long as III. Antenna 2/3 length of forewing, male flagellum with long fine cilia.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFEBFFC244F8F98C94D574B0" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76" type="description">
<paragraph id="961536E4FFEBFFC144F8F98C91C67271" blockId="1.[151,1437,1441,2008]" pageId="1" pageNumber="76">
<emphasis id="A4DEEAF6FFEBFFC144F8F98C912D733A" box="[199,280,1549,1575]" italics="true" pageId="1" pageNumber="76">Thorax</emphasis>
: Glossy white, cream or grey, without markings. Legs same color as head; fore femur, tibia, and tarsi dark fuscous on outer surface; hind tibia sometimes dark fuscous at middle and at apex, with long bristles along basal 2/3 of dorsal margin, whorls of bristles at apex. Forewing slender, ground color glossy white, cream, grey, or black, distal 1/3 to 1/4 often with an orange patch, a black apical spot often present; ventral forewing with a group of stiff scales near basal fork of 1A+2A; pterostigma weakly or not developed. Forewing venation (
<figureCitation id="0E912A61FFEBFFC140C9F91C956E73AA" box="[1270,1371,1693,1719]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1569,1593]" captionTargetBox="[170,1427,421,1519]" captionTargetId="figure-69@3.[151,1435,396,1546]" captionTargetPageId="3" captionText="FIGURES 1–2. Wing venation of Tenupalpa spp. 1, T. angustella, male, slide no. NKLGE013; 2, T. venustalis, female, slide no. LGE18032W (Scale bars = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819796" httpUri="https://zenodo.org/record/5819796/files/figure.png" pageId="1" pageNumber="76">Figs 1, 2</figureCitation>
) with Sc reaching basal 1/4 or 1/3 of costa, R and Rs
<subScript id="0A2E34A1FFEBFFC1469DF955929F73FF" attach="left" box="[674,682,1748,1762]" fontSize="6" pageId="1" pageNumber="76">1</subScript>
free, Rs
<subScript id="0A2E34A1FFEBFFC14737F955932573FF" attach="left" box="[776,784,1748,1762]" fontSize="6" pageId="1" pageNumber="76">2</subScript>
stalked or connate with Rs
<subScript id="0A2E34A1FFEBFFC1407FF955947D73FF" attach="left" box="[1088,1096,1748,1762]" fontSize="6" pageId="1" pageNumber="76">3</subScript>
, M
<subScript id="0A2E34A1FFEBFFC14050F955944273FF" attach="left" box="[1135,1143,1748,1762]" fontSize="6" pageId="1" pageNumber="76">1</subScript>
free from or connate with Rs
<subScript id="0A2E34A1FFEBFFC1448AF9799088721B" attach="left" box="[181,189,1784,1798]" fontSize="6" pageId="1" pageNumber="76">3</subScript>
, M
<subScript id="0A2E34A1FFEBFFC144DCF97990DE721B" attach="left" box="[227,235,1784,1798]" fontSize="6" pageId="1" pageNumber="76">2</subScript>
close to M
<subScript id="0A2E34A1FFEBFFC1455AF9799158721B" attach="left" box="[357,365,1784,1798]" fontSize="6" pageId="1" pageNumber="76">3</subScript>
, CuA
<subScript id="0A2E34A1FFEBFFC14590F9799182721B" attach="left" box="[431,439,1784,1798]" fontSize="6" pageId="1" pageNumber="76">1</subScript>
and CuA
<subScript id="0A2E34A1FFEBFFC1461EF979921C721B" attach="left" box="[545,553,1784,1798]" fontSize="6" pageId="1" pageNumber="76">2</subScript>
free, 1A+2A forked near base. Hindwing narrower than forewing, ground color grey, fuscous- or black; apex acute, often narrowly black-tipped; fringe scales sometimes with a black terminal band. Hindwing venation with R joining Sc beyond base, Rs stalked with M
<subScript id="0A2E34A1FFEBFFC147DAF8C193D87253" attach="left" box="[997,1005,1856,1870]" fontSize="6" pageId="1" pageNumber="76">1</subScript>
at distal 1/5, M
<subScript id="0A2E34A1FFEBFFC140A6F8C194947253" attach="left" box="[1177,1185,1856,1870]" fontSize="6" pageId="1" pageNumber="76">2</subScript>
often weak, CuA
<subScript id="0A2E34A1FFEBFFC1415AF8C195587253" attach="left" box="[1381,1389,1856,1870]" fontSize="6" pageId="1" pageNumber="76">1</subScript>
and CuA
<subScript id="0A2E34A1FFEBFFC144F3F8E590E1726F" attach="left" box="[204,212,1892,1906]" fontSize="6" pageId="1" pageNumber="76">2</subScript>
parallel, discal cell open.
</paragraph>
<paragraph id="961536E4FFEBFFC244F8F8F494D574B0" blockId="1.[151,1437,1441,2008]" lastBlockId="2.[151,1437,151,681]" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76">
<emphasis id="A4DEEAF6FFEBFFC144F8F8F491047292" box="[199,305,1909,1935]" italics="true" pageId="1" pageNumber="76">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFEBFFC1457DF8F491997292" box="[322,428,1909,1935]" captionStart="FIGURES 3–6" captionStartId="4.[151,264,1894,1918]" captionTargetBox="[161,1422,453,1848]" captionTargetId="figure-154@4.[151,1435,453,1871]" captionTargetPageId="4" captionText="FIGURES 3–6. Abdominal and genital structures of Tenupalpa spp. 3, distal segments of the abdomen and male genitalia of T. angustella in external view; 4, denuded male 8th abdominal segment of T. angustella (image from Lee et al., 2021: Fig. 7I), slide no. LGE18719; 5, male genitalia of T. angustella, slide no. LGE16107; 6, female genitalia of T. nephodesma, slide no. LGE19022 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819798" httpUri="https://zenodo.org/record/5819798/files/figure.png" pageId="1" pageNumber="76">Figs 3, 4</figureCitation>
). Male sternum II–VII with sclerotized plates, VIII round, sub-triangular or pentagonal; tergum VIII reduced, with mediolateral coremata, a linear sclerotization between coremata. Male genitalia (
<figureCitation id="0E912A61FFEBFFC1416FF81895A672A9" box="[1360,1427,1945,1972]" captionStart="FIGURES 3–6" captionStartId="4.[151,264,1894,1918]" captionTargetBox="[161,1422,453,1848]" captionTargetId="figure-154@4.[151,1435,453,1871]" captionTargetPageId="4" captionText="FIGURES 3–6. Abdominal and genital structures of Tenupalpa spp. 3, distal segments of the abdomen and male genitalia of T. angustella in external view; 4, denuded male 8th abdominal segment of T. angustella (image from Lee et al., 2021: Fig. 7I), slide no. LGE18719; 5, male genitalia of T. angustella, slide no. LGE16107; 6, female genitalia of T. nephodesma, slide no. LGE19022 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819798" httpUri="https://zenodo.org/record/5819798/files/figure.png" pageId="1" pageNumber="76">Fig. 5</figureCitation>
) with uncus small and triangular, often with medial ridge. Culcitula usually present. Gnathos short, hook-shaped. Tegumen sub-triangular, usually with round tubercles medially, anterior margin deeply concave. Valva slender and simple, apex rounded, rarely pointed. Anellus lobe well developed, round, oval, or digitate, rarely asymmetrical, typically bearing a long bristle at apex, bristle sometimes strongly developed as a spine. Juxta simple, divided into two lobes, sometimes with a projection. Vinculum simple, often with posterior margin broadly concave. Saccus broad. Aedeagus with bulbous base and slender distal part, often curved distally. Female genitalia (
<figureCitation id="0E912A61FFE8FFC240D3FEA69505745F" box="[1260,1328,295,322]" captionStart="FIGURES 3–6" captionStartId="4.[151,264,1894,1918]" captionTargetBox="[161,1422,453,1848]" captionTargetId="figure-154@4.[151,1435,453,1871]" captionTargetPageId="4" captionText="FIGURES 3–6. Abdominal and genital structures of Tenupalpa spp. 3, distal segments of the abdomen and male genitalia of T. angustella in external view; 4, denuded male 8th abdominal segment of T. angustella (image from Lee et al., 2021: Fig. 7I), slide no. LGE18719; 5, male genitalia of T. angustella, slide no. LGE16107; 6, female genitalia of T. nephodesma, slide no. LGE19022 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819798" httpUri="https://zenodo.org/record/5819798/files/figure.png" pageId="2" pageNumber="77">Fig. 6</figureCitation>
) with papillae anales setose. Apophyses anteriores as long as or shorter than apophyses posteriores. Sternum VIII medially emarginate or convex. Ostium often small, at anterior or posterior margin of sternum VIII. Ductus bursae slender and simple. Corpus bursae elongate oval; signum semilunar or a linear sclerite, sometimes absent.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE8FFC244F8FE37937574CC" box="[199,832,438,465]" pageId="2" pageNumber="77" type="distribution">
<paragraph id="961536E4FFE8FFC244F8FE37937574CC" blockId="2.[151,1437,151,681]" box="[199,832,438,465]" pageId="2" pageNumber="77">
<emphasis id="A4DEEAF6FFE8FFC244F8FE37915474CD" bold="true" box="[199,353,438,464]" pageId="2" pageNumber="77">Distribution.</emphasis>
Oriental and Palearctic regions (
<figureCitation id="0E912A61FFE8FFC246EEFE36930674CC" box="[721,819,439,465]" captionStart="FIGURES 103" captionStartId="34.[151,264,1128,1152]" captionTargetBox="[177,1411,201,1105]" captionTargetId="figure-22@34.[177,1411,201,1105]" captionTargetPageId="34" captionText="FIGURES 103. Global distribution map of the genus Tenupalpa." figureDoi="http://doi.org/10.5281/zenodo.5819846" httpUri="https://zenodo.org/record/5819846/files/figure.png" pageId="2" pageNumber="77">Fig. 103</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFE8FFC244F8FE5A921177B4" pageId="2" pageNumber="77" type="discussion">
<paragraph id="961536E4FFE8FFC244F8FE5A921177B4" blockId="2.[151,1437,151,681]" pageId="2" pageNumber="77">
<emphasis id="A4DEEAF6FFE8FFC244F8FE5A910E74E8" bold="true" box="[199,315,475,501]" pageId="2" pageNumber="77">Remarks.</emphasis>
Superficially, many species of
<taxonomicName id="51AA4D67FFE8FFC24694FE5A931574E8" authorityName="Lee et Li" authorityYear="2021" box="[683,800,475,501]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFE8FFC24694FE5A931574E8" box="[683,800,475,501]" italics="true" pageId="2" pageNumber="77">Tenupalpa</emphasis>
</taxonomicName>
are similar to those of
<emphasis id="A4DEEAF6FFE8FFC24009FE5A949374E8" box="[1078,1190,475,501]" italics="true" pageId="2" pageNumber="77">Pulchrala</emphasis>
in having a white to greyish forewing with an orange distal patch. However, males of
<taxonomicName id="51AA4D67FFE8FFC2474FFE7E93D07704" authorityName="Lee et Li" authorityYear="2021" box="[880,997,511,537]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFE8FFC2474FFE7E93D07704" box="[880,997,511,537]" italics="true" pageId="2" pageNumber="77">Tenupalpa</emphasis>
</taxonomicName>
typically have unmodified labial palpi, whereas those of
<emphasis id="A4DEEAF6FFE8FFC24565FDA291FF7720" box="[346,458,547,573]" italics="true" pageId="2" pageNumber="77">Pulchrala</emphasis>
have expansible hair pencils on the dorsal surface. Also, the coremata of
<taxonomicName id="51AA4D67FFE8FFC240C1FDA295467720" authorityName="Lee et Li" authorityYear="2021" box="[1278,1395,547,573]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFE8FFC240C1FDA295467720" box="[1278,1395,547,573]" italics="true" pageId="2" pageNumber="77">Tenupalpa</emphasis>
</taxonomicName>
are short and situated mediolaterally on tergum VIII, whereas they are rather long and situated laterally in
<emphasis id="A4DEEAF6FFE8FFC2411AFDC695A8777C" box="[1317,1437,583,609]" italics="true" pageId="2" pageNumber="77">Pulchrala.</emphasis>
The presence of tegumen tubercles, which is typical of
<taxonomicName id="51AA4D67FFE8FFC246C8FDEA93597798" authorityName="Lee et Li" authorityYear="2021" box="[759,876,619,645]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="77" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFE8FFC246C8FDEA93597798" box="[759,876,619,645]" italics="true" pageId="2" pageNumber="77">Tenupalpa</emphasis>
</taxonomicName>
, has been reported elsewhere only in some species of
<taxonomicName id="51AA4D67FFE8FFC24489FD0E910477B4" authorityName="Rondani" authorityYear="1876" box="[182,305,655,681]" class="Insecta" family="Gelechiidae" genus="Palumbina" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="A4DEEAF6FFE8FFC24489FD0E910477B4" box="[182,305,655,681]" italics="true" pageId="2" pageNumber="77">Palumbina</emphasis>
</taxonomicName>
within
<taxonomicName id="51AA4D67FFE8FFC245BAFD0E921577B4" authorityName="Thiotrichinae Karsholt, Mutanen, Lee &amp; Kaila" authorityYear="2013" box="[389,544,655,681]" pageId="2" pageNumber="110" rank="subFamily" subFamily="Thiotrichinae">Thiotrichinae</taxonomicName>
.
</paragraph>
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<taxonomicName id="51AA4D67FFECFFC644A8FCE7922F769D" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,538,870,897]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="species" species="latifasciata" status="sp. nov.">
<emphasis id="A4DEEAF6FFECFFC644A8FCE79191769C" bold="true" box="[151,420,870,897]" italics="true" pageId="6" pageNumber="81">Tenupalpa latifasciata</emphasis>
Lee
<emphasis id="A4DEEAF6FFECFFC645DFFCE691C0769C" bold="true" box="[480,501,871,897]" italics="true" pageId="6" pageNumber="81">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFECFFC64618FCE692B0769C" box="[551,645,871,897]" pageId="6" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644A8FC0A938476DF" pageId="6" pageNumber="81" type="description">
<paragraph id="961536E4FFECFFC644A8FC0A914E76BB" blockId="6.[151,945,870,962]" box="[151,379,907,934]" pageId="6" pageNumber="81">
(
<figureCitation id="0E912A61FFECFFC644A0FC0A90C376B8" box="[159,246,907,933]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="6" pageNumber="81">Figs 16</figureCitation>
,
<figureCitation id="0E912A61FFECFFC6453DFC0A911576B8" box="[258,288,907,933]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="6" pageNumber="81">28</figureCitation>
,
<figureCitation id="0E912A61FFECFFC64513FC0A917F76B8" box="[300,330,907,933]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="6" pageNumber="81">48</figureCitation>
,
<figureCitation id="0E912A61FFECFFC64569FC0A914676BB" box="[342,371,907,934]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="6" pageNumber="81">67</figureCitation>
)
</paragraph>
<paragraph id="961536E4FFECFFC644A8FC2B938476DF" blockId="6.[151,945,870,962]" box="[151,945,938,962]" pageId="6" pageNumber="81">
<uri id="E23B3AE6FFECFFC644A8FC2B938476DF" box="[151,945,938,962]" pageId="6" pageNumber="81">
urn:lsid:zoobank.org:act:
<uuid id="E20C0C31FFECFFC6459FFC2B938476DF" box="[416,945,938,962]" pageId="6" pageNumber="81">6D9CCEE3-3C46-45C6-AFD9-E51A2C02C6E9</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644A8FC71919A7186" pageId="6" pageNumber="81" type="materials_examined">
<paragraph id="961536E4FFECFFC644A8FC71919A7186" blockId="6.[151,1437,1008,2007]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644A8FC7191737117" bold="true" box="[151,326,1008,1034]" pageId="6" pageNumber="81">Type material.</emphasis>
<materialsCitation id="26C23CB9FFECFFC64570FC71957A7133" ID-GBIF-Occurrence="3427696442" accessionNumber="LGE16111" collectingDate="2015-07-21" collectorName="Mujie Qi &amp; Shengnan Zhao" country="China" county="Autonomous Region" elevation="1364" latitude="24.15" location="Dayaoshan Nature Reserves" longLatPrecision="751" longitude="110.21" municipality="Jinxiu" pageId="6" pageNumber="81" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus id="49118846FFECFFC64570FC7191827117" box="[335,439,1008,1034]" pageId="6" pageNumber="81" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFECFFC64580FC7091E37116" box="[447,470,1009,1035]" pageId="6" pageNumber="81" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFECFFC645DBFC7192757117" box="[484,576,1008,1034]" name="China" pageId="6" pageNumber="81">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFECFFC64673FC71933F7117" box="[588,778,1008,1034]" country="China" name="Guangxi" pageId="6" pageNumber="81">Guangxi Zhuang</collectingRegion>
<collectingCounty id="7F744E68FFECFFC6472EFC7193CB7117" box="[785,1022,1008,1034]" pageId="6" pageNumber="81">Autonomous Region</collectingCounty>
,
<collectingMunicipality id="7671AC9EFFECFFC64035FC7194647117" box="[1034,1105,1008,1034]" pageId="6" pageNumber="81">Jinxiu</collectingMunicipality>
,
<location id="9375603FFFECFFC64061FC7195A97116" LSID="urn:lsid:plazi:treatment:1E0387F2FFECFFCC44A8FCE791EE703B:9375603FFFECFFC64061FC7195A97116" box="[1118,1436,1008,1035]" country="China" county="Autonomous Region" latitude="24.15" longLatPrecision="751" longitude="110.21" municipality="Jinxiu" name="Dayaoshan Nature Reserves" pageId="6" pageNumber="81" stateProvince="Guangxi">Dayaoshan Nature Reserves</location>
(
<geoCoordinate id="F39E5023FFECFFC644A0FB9591377132" box="[159,258,1044,1071]" degrees="24.15" direction="north" orientation="latitude" pageId="6" pageNumber="81" precision="555" value="24.15">24.15°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFECFFC64532FB95914C7133" box="[269,377,1044,1070]" degrees="110.21" direction="east" orientation="longitude" pageId="6" pageNumber="81" precision="555" value="110.21">110.21°E</geoCoordinate>
),
<quantity id="51529B01FFECFFC645B3FB9591D67132" box="[396,483,1044,1071]" metricMagnitude="3" metricUnit="m" metricValue="1.364" pageId="6" pageNumber="81" unit="m" value="1364.0">
<elevation id="1D87D1D7FFECFFC645B3FB9591D67132" box="[396,483,1044,1071]" metricMagnitude="3" metricUnit="m" metricValue="1.364" pageId="6" pageNumber="81" unit="m" value="1364.0">1364 m</elevation>
</quantity>
,
<date id="E2141024FFECFFC645D0FB9592457132" box="[495,624,1044,1071]" pageId="6" pageNumber="81" value="2015-07-21">
<collectingDate id="F250E9CCFFECFFC645D0FB9592457132" box="[495,624,1044,1071]" pageId="6" pageNumber="81" value="2015-07-21">21.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFECFFC64693FB9593277133" box="[684,786,1044,1070]" pageId="6" pageNumber="81">Mujie Qi</collectorName>
and
<collectorName id="3B5F5332FFECFFC64777FB9593CC7133" box="[840,1017,1044,1070]" pageId="6" pageNumber="81">Shengnan Zhao</collectorName>
, genitalia slide no. LGE16111
</materialsCitation>
.
<materialsCitation id="26C23CB9FFECFFC64166FB959123716B" ID-GBIF-Occurrence="3427696335" accessionNumber="LGE16112" collectingDate="2015-07-19" collectorName="Mujie Qi &amp; Shengnan Zhao" country="China" county="Autonomous Region" elevation="1364" latitude="24.15" location="Autonomous Region" longLatPrecision="751" longitude="110.21" municipality="Jinxiu" pageId="6" pageNumber="81" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<emphasis id="A4DEEAF6FFECFFC64166FB959281714F" bold="true" pageId="6" pageNumber="81">
<typeStatus id="49118846FFECFFC64166FB9590E2714F" pageId="6" pageNumber="81" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="546EF806FFECFFC644DBFBB99185714F" box="[228,432,1080,1106]" country="China" name="Guangxi" pageId="6" pageNumber="81">Guangxi Zhuang</collectingRegion>
<location id="9375603FFFECFFC64589FBB9929B714F" LSID="urn:lsid:plazi:treatment:1E0387F2FFECFFCC44A8FCE791EE703B:9375603FFFECFFC64589FBB9929B714F" box="[438,686,1080,1106]" name="Autonomous Region" pageId="6" pageNumber="81" stateProvince="Guangxi">Autonomous Region</location>
:
</emphasis>
<specimenCount id="80ACFD6DFFECFFC64684FBB992DD714E" box="[699,744,1080,1107]" pageId="6" pageNumber="81" type="male">1 ♂</specimenCount>
, same data as holotype except
<date id="E2141024FFECFFC64079FBB994FD714E" box="[1094,1224,1080,1107]" pageId="6" pageNumber="81" value="2015-07-19">
<collectingDate id="F250E9CCFFECFFC64079FBB994FD714E" box="[1094,1224,1080,1107]" pageId="6" pageNumber="81" value="2015-07-19">19.vii.2015</collectingDate>
</date>
, genitalia slide no. LGE16112
</materialsCitation>
.
<materialsCitation id="26C23CB9FFECFFC6451FFBDD919E7186" ID-GBIF-Occurrence="3427696431" accessionNumber="ZJL05256" collectingDate="2004-05-19" collectorName="Haili Yu" country="China" elevation="740" latitude="27.45" location="Mt. Wuyi" longLatPrecision="742" longitude="117.41" municipality="Sangang" pageId="6" pageNumber="81" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="paratype">
<emphasis id="A4DEEAF6FFECFFC6451FFBDD91DC716B" bold="true" box="[288,489,1116,1142]" pageId="6" pageNumber="81">
<collectingRegion id="546EF806FFECFFC6451FFBDD91D1716B" box="[288,484,1116,1142]" country="China" name="Fujian" pageId="6" pageNumber="81">Fujian Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFECFFC645D0FBDD922F716A" box="[495,538,1116,1143]" pageId="6" pageNumber="81" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFECFFC64619FBDD92B3716B" box="[550,646,1116,1142]" pageId="6" pageNumber="81">Sangang</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFECFFC646AAFBDD92C2716A" box="[661,759,1116,1143]" degrees="27.45" direction="north" orientation="latitude" pageId="6" pageNumber="81" precision="555" value="27.45">27.45°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFECFFC6473DFBDD935A716B" box="[770,879,1116,1142]" degrees="117.41" direction="east" orientation="longitude" pageId="6" pageNumber="81" precision="555" value="117.41">117.41°E</geoCoordinate>
),
<quantity id="51529B01FFECFFC647BEFBDD93FF716A" box="[897,970,1116,1143]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="81" unit="m" value="740.0">
<elevation id="1D87D1D7FFECFFC647BEFBDD93FF716A" box="[897,970,1116,1143]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="81" unit="m" value="740.0">740 m</elevation>
</quantity>
,
<location id="9375603FFFECFFC647EAFBDC9476716B" LSID="urn:lsid:plazi:treatment:1E0387F2FFECFFCC44A8FCE791EE703B:9375603FFFECFFC647EAFBDC9476716B" box="[981,1091,1116,1143]" country="China" latitude="27.45" longLatPrecision="742" longitude="117.41" municipality="Sangang" name="Mt. Wuyi" pageId="6" pageNumber="81" stateProvince="Fujian">Mt. Wuyi</location>
,
<date id="E2141024FFECFFC64072FBDD948B716B" box="[1101,1214,1116,1142]" pageId="6" pageNumber="81" value="2004-05-19">
<collectingDate id="F250E9CCFFECFFC64072FBDD948B716B" box="[1101,1214,1116,1142]" pageId="6" pageNumber="81" value="2004-05-19">19.v.2004</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFECFFC640C6FBDD956D716A" box="[1273,1368,1116,1143]" pageId="6" pageNumber="81">Haili Yu</collectorName>
, genitalia slide no. ZJL05256
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644F8FB25938E7036" pageId="6" pageNumber="81" type="diagnosis">
<paragraph id="961536E4FFECFFC644F8FB25938E7036" blockId="6.[151,1437,1008,2007]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FB25917771A3" bold="true" box="[199,322,1188,1214]" pageId="6" pageNumber="81">Diagnosis.</emphasis>
<taxonomicName id="51AA4D67FFECFFC64578FB25927471A3" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[327,577,1188,1214]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="species" species="latifasciata" status="sp. nov.">
<emphasis id="A4DEEAF6FFECFFC64578FB25927471A3" box="[327,577,1188,1214]" italics="true" pageId="6" pageNumber="81">Tenupalpa latifasciata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFECFFC64678FB2592AA71A3" bold="true" box="[583,671,1188,1214]" pageId="6" pageNumber="81">
<taxonomicNameLabel id="BFED578DFFECFFC64678FB2592AA71A3" box="[583,671,1188,1214]" pageId="6" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="51AA4D67FFECFFC64755FB24942471A3" box="[874,1041,1188,1214]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFECFFC64755FB24942471A3" box="[874,1041,1188,1214]" italics="true" pageId="6" pageNumber="81">T. nephodesma</emphasis>
</taxonomicName>
; the two share a broad fascia on the forewing and a short projection from the costa of valva. The new species can be distinguished from the latter by the shape of the costal projection, which is longer and slenderer than that of
<taxonomicName id="51AA4D67FFECFFC647FDFB6C945C701B" box="[962,1129,1260,1286]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFECFFC647FDFB6C945C701B" box="[962,1129,1260,1286]" italics="true" pageId="6" pageNumber="81">T. nephodesma</emphasis>
</taxonomicName>
. The curved, finger-shaped anellus lobes of the male genitalia are also diagnostic for
<taxonomicName id="51AA4D67FFECFFC64724FA9093817037" authorityName="Lee &amp; Li" authorityYear="2022" box="[795,948,1296,1322]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="species" species="latifasciata">
<emphasis id="A4DEEAF6FFECFFC64724FA9093817037" box="[795,948,1296,1322]" italics="true" pageId="6" pageNumber="81">T. latifasciata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644F8FAB591A472AF" pageId="6" pageNumber="81" type="description">
<paragraph id="961536E4FFECFFC644F8FAB593F2708B" blockId="6.[151,1437,1008,2007]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FAB5916C7053" bold="true" box="[199,345,1332,1358]" pageId="6" pageNumber="81">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFECFFC64590FAB592367053" box="[431,515,1332,1358]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="6" pageNumber="81">Fig. 48</figureCitation>
).
<emphasis id="A4DEEAF6FFECFFC64628FAB592617053" box="[535,596,1332,1358]" italics="true" pageId="6" pageNumber="81">Head</emphasis>
: Creamy white. Labial palpus creamy white; palpomere III strongly mixed with brownish grey, as long as II. Antenna with scape creamy white; flagellum dark fuscous, except basal 1/3 of dorsal surface creamy white, male cilia as long as flagellomere diameter.
</paragraph>
<paragraph id="961536E4FFECFFC644F8FA2193547372" blockId="6.[151,1437,1008,2007]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FA21912D70A7" box="[199,280,1440,1466]" italics="true" pageId="6" pageNumber="81">Thorax</emphasis>
: Creamy white. Tegula creamy white, except anterior margin slightly tinged with fuscous. Wingspan 7.0–8.0 mm; forewing basal 1/3 creamy white, anterior edge of costa black; a broad, fuscous transverse fascia between 1/3 and 2/3 of wing, its posterior margin creamy white; distal 1/3 creamy white to brownish grey, strongly mixed with fuscous at apex; fringe scales brownish grey. Hindwing grey; fringe scales brownish grey. Legs creamy white; mid-tibia sometimes with a black spot at distal 1/3; hind tibia broadly black between distal 1/2 and 1/4, black at apex; mid- and hind tarsi strongly mixed with brownish grey.
</paragraph>
<paragraph id="961536E4FFECFFC644F8F9F995317292" blockId="6.[151,1437,1008,2007]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8F9F99104738F" box="[199,305,1656,1682]" italics="true" pageId="6" pageNumber="81">Abdomen</emphasis>
: Male coremata nearly as long as sternum VIII; sternum VIII pentagonal, shortly concave distally (
<figureCitation id="0E912A61FFECFFC6449FF91D90C373AA" box="[160,246,1692,1719]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="6" pageNumber="81">Fig. 16</figureCitation>
). Male genitalia (
<figureCitation id="0E912A61FFECFFC645FAF91D922C73AA" box="[453,537,1692,1719]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="6" pageNumber="81">Fig. 67</figureCitation>
) with uncus small and triangular, protruded medially, distal 1/2 covered with minute protuberances. Gnathos hook short and thick, tapering from middle to apex, pointed apically. Tegumen with diaphragma spinous in posterior 1/2, anterior 2/5 bearing papilliform tubercles (
<figureCitation id="0E912A61FFECFFC64013F96594B773E3" box="[1068,1154,1764,1790]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="6" pageNumber="81">Fig. 28</figureCitation>
). Valva slender, strongly curved inward after base, gradually broadening toward apex; costa with an elongate-triangular projection at middle. Anellus lobe digitate, 1/5 length of valva, strongly curved inward, bifid apically; subapical bristle as long as anellus lobe. Juxta with inner margin broadly convex, with sparse setae at middle. Vinculum broad, sub-trapezoidal. Saccus short and broad, sub-triangular. Aedeagus with basal half dilated and distal 1/2 slender, blunt at apex.
</paragraph>
<paragraph id="961536E4FFECFFC644F8F81991A472AF" blockId="6.[151,1437,1008,2007]" box="[199,401,1944,1970]" pageId="6" pageNumber="81">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644F8F83D92B772CA" box="[199,642,1980,2007]" pageId="6" pageNumber="81" type="distribution">
<paragraph id="961536E4FFECFFC644F8F83D92B772CA" blockId="6.[151,1437,1008,2007]" box="[199,642,1980,2007]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8F83D916C72CB" bold="true" box="[199,345,1980,2006]" pageId="6" pageNumber="81">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFECFFC64558F83D919F72CB" box="[359,426,1980,2006]" name="China" pageId="6" pageNumber="81">China</collectingCountry>
(
<collectingRegion id="546EF806FFECFFC64586F83D923072CB" box="[441,517,1980,2006]" country="China" name="Fujian" pageId="6" pageNumber="81">Fujian</collectingRegion>
,
<collectingRegion id="546EF806FFECFFC6462FF83D924072CA" box="[528,629,1980,2007]" country="China" name="Guangxi" pageId="6" pageNumber="81">Guangxi</collectingRegion>
).
</paragraph>
</subSubSection>
<caption id="C2D5666CFFEDFFC744A8F8E5936072F4" ID-DOI="http://doi.org/10.5281/zenodo.5819800" ID-Zenodo-Dep="5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="7" pageNumber="82" startId="7.[151,264,1892,1916]" targetBox="[234,1368,181,1857]" targetPageId="7">
<paragraph id="961536E4FFEDFFC744A8F8E5936072F4" blockId="7.[151,1437,1892,2025]" pageId="7" pageNumber="82">
<emphasis id="A4DEEAF6FFEDFFC744A8F8E591727260" bold="true" box="[151,327,1892,1917]" pageId="7" pageNumber="82">FIGURES 7–14.</emphasis>
Distal segment of male abdomen in
<taxonomicName id="51AA4D67FFEDFFC74685F8E4935C7261" box="[698,873,1893,1917]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="undefined-7">
<emphasis id="A4DEEAF6FFEDFFC74685F8E493117260" box="[698,804,1893,1917]" italics="true" pageId="7" pageNumber="82">Tenupalpa</emphasis>
spp. 7
</taxonomicName>
.
<taxonomicName id="51AA4D67FFEDFFC7474CF8E493DC7260" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[883,1001,1893,1917]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="fuscalata" status="sp. nov.">
<emphasis id="A4DEEAF6FFEDFFC7474CF8E493DC7260" box="[883,1001,1893,1917]" italics="true" pageId="7" pageNumber="82">T. fuscalata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFEDFFC747D0F8E4940A7260" bold="true" box="[1007,1087,1893,1917]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="BFED578DFFEDFFC747D0F8E4940A7260" box="[1007,1087,1893,1917]" pageId="7" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18829; 8.
<taxonomicName id="51AA4D67FFEDFFC74100F8E4908B72BC" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="isabellina" status="sp. nov.">
<emphasis id="A4DEEAF6FFEDFFC74100F8E4908B72BC" italics="true" pageId="7" pageNumber="82">T. isabellina</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFEDFFC744F9F808912272BC" bold="true" box="[198,279,1929,1953]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="BFED578DFFEDFFC744F9F808912272BC" box="[198,279,1929,1953]" pageId="7" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18835; 9.
<taxonomicName id="51AA4D67FFEDFFC74624F808924E72BC" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[539,635,1929,1953]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="falcata" status="sp. nov.">
<emphasis id="A4DEEAF6FFEDFFC74624F808924E72BC" box="[539,635,1929,1953]" italics="true" pageId="7" pageNumber="82">T. falcata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFEDFFC746BDF80892E172BC" bold="true" box="[642,724,1929,1953]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="BFED578DFFEDFFC746BDF80892E172BC" box="[642,724,1929,1953]" pageId="7" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18843; 10.
<taxonomicName id="51AA4D67FFEDFFC747DAF808944F72BC" baseAuthorityName="Kyaw, Yagi &amp; Hirowatari" baseAuthorityYear="2019" box="[997,1146,1929,1953]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFEDFFC747DAF808944F72BC" box="[997,1146,1929,1953]" italics="true" pageId="7" pageNumber="82">T. flavitermina</emphasis>
</taxonomicName>
, slide no. LGE18842; 11.
<taxonomicName id="51AA4D67FFEDFFC741B5F80890C972D8" baseAuthorityName="Omelko" baseAuthorityYear="1984" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="venustalis">
<emphasis id="A4DEEAF6FFEDFFC741B5F80890C972D8" italics="true" pageId="7" pageNumber="82">T. venustalis</emphasis>
</taxonomicName>
, slide no. LGE18562; 12.
<taxonomicName id="51AA4D67FFEDFFC74632F82C92A972D8" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[525,668,1965,1989]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="uncativalva" status="sp. nov.">
<emphasis id="A4DEEAF6FFEDFFC74632F82C92A972D8" box="[525,668,1965,1989]" italics="true" pageId="7" pageNumber="82">T. uncativalva</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFEDFFC7469CF82C92C072D8" bold="true" box="[675,757,1965,1989]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="BFED578DFFEDFFC7469CF82C92C072D8" box="[675,757,1965,1989]" pageId="7" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19015; 13.
<taxonomicName id="51AA4D67FFEDFFC7403AF82C945472D8" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1029,1121,1965,1989]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="zhengi" status="sp. nov.">
<emphasis id="A4DEEAF6FFEDFFC7403AF82C945472D8" box="[1029,1121,1965,1989]" italics="true" pageId="7" pageNumber="82">T. zhengi</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFEDFFC74057F82C948C72D8" bold="true" box="[1128,1209,1965,1989]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="BFED578DFFEDFFC74057F82C948C72D8" box="[1128,1209,1965,1989]" pageId="7" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19001; 14.
<taxonomicName id="51AA4D67FFEDFFC74482F850910372F4" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[189,310,2001,2025]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="dissimilis" status="sp. nov.">
<emphasis id="A4DEEAF6FFEDFFC74482F850910372F4" box="[189,310,2001,2025]" italics="true" pageId="7" pageNumber="82">T. dissimilis</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFEDFFC74503F85091B872F4" bold="true" box="[316,397,2001,2025]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="BFED578DFFEDFFC74503F85091B872F4" box="[316,397,2001,2025]" pageId="7" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19007 (Scale bars = 0.2 mm).
</paragraph>
</caption>
<caption id="C2D5666CFFE2FFC844A8F96E9358728A" ID-DOI="http://doi.org/10.5281/zenodo.5819802" ID-Zenodo-Dep="5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="8" pageNumber="83" startId="8.[151,264,1775,1799]" targetBox="[170,1407,181,1737]" targetPageId="8">
<paragraph id="961536E4FFE2FFC844A8F96E9358728A" blockId="8.[151,1437,1775,1944]" pageId="8" pageNumber="83">
<emphasis id="A4DEEAF6FFE2FFC844A8F96E9160721A" bold="true" box="[151,341,1775,1799]" pageId="8" pageNumber="83">FIGURES 15–25.</emphasis>
Male sternum VIII of
<taxonomicName id="51AA4D67FFE2FFC8467EF97192E8721A" box="[577,733,1776,1800]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A4DEEAF6FFE2FFC8467EF971929E7215" box="[577,683,1776,1800]" italics="true" pageId="8" pageNumber="83">Tenupalpa</emphasis>
spp.
</taxonomicName>
15.
<taxonomicName id="51AA4D67FFE2FFC84733F97193907215" box="[780,933,1776,1800]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFE2FFC84733F97193907215" box="[780,933,1776,1800]" italics="true" pageId="8" pageNumber="83">T. nephodesma</emphasis>
</taxonomicName>
, slide no. ZJL05343; 16.
<taxonomicName id="51AA4D67FFE2FFC8408FF97195097215" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1200,1340,1776,1800]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="latifasciata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC8408FF97195097215" box="[1200,1340,1776,1800]" italics="true" pageId="8" pageNumber="83">T. latifasciata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC8417BF97195A07215" bold="true" box="[1348,1429,1776,1800]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC8417BF97195A07215" box="[1348,1429,1776,1800]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE16112; 17.
<taxonomicName id="51AA4D67FFE2FFC845A1F89592057231" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[414,560,1812,1836]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC845A1F89592057231" box="[414,560,1812,1836]" italics="true" pageId="8" pageNumber="83">T. immaculata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC84607F89592BE7231" bold="true" box="[568,651,1812,1836]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC84607F89592BE7231" box="[568,651,1812,1836]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18025; 18.
<taxonomicName id="51AA4D67FFE2FFC8479EF895942C7231" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[929,1049,1812,1836]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="angulosa" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC8479EF895942C7231" box="[929,1049,1812,1836]" italics="true" pageId="8" pageNumber="83">T. angulosa</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC8401EF89594467231" bold="true" box="[1057,1139,1812,1836]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC8401EF89594467231" box="[1057,1139,1812,1836]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE17270; 19.
<taxonomicName id="51AA4D67FFE2FFC841B5F89590DE724D" baseAuthorityName="Omelko" baseAuthorityYear="1984" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFE2FFC841B5F89590DE724D" italics="true" pageId="8" pageNumber="83">T. biformis</emphasis>
</taxonomicName>
, slide no. LLJ15140; 20.
<taxonomicName id="51AA4D67FFE2FFC845D2F8B992647252" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[493,593,1848,1871]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC845D2F8B992647252" box="[493,593,1848,1871]" italics="true" pageId="8" pageNumber="83">T. acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC84669F8B99293724D" bold="true" box="[598,678,1848,1872]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC84669F8B99293724D" box="[598,678,1848,1872]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE17045; 21.
<taxonomicName id="51AA4D67FFE2FFC8478FF8B99477724D" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[944,1090,1848,1872]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="crassimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC8478FF8B99477724D" box="[944,1090,1848,1872]" italics="true" pageId="8" pageNumber="83">T. crassimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC84077F8B994AD724D" bold="true" box="[1096,1176,1848,1872]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC84077F8B994AD724D" box="[1096,1176,1848,1872]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18015; 22.
<taxonomicName id="51AA4D67FFE2FFC844A8F8DD91177269" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[151,290,1884,1908]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="acutimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC844A8F8DD91177269" box="[151,290,1884,1908]" italics="true" pageId="8" pageNumber="83">T. acutimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC84516F8DD914E7269" bold="true" box="[297,379,1884,1908]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC84516F8DD914E7269" box="[297,379,1884,1908]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE16247; 23.
<taxonomicName id="51AA4D67FFE2FFC846B2F8DD93057269" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[653,816,1884,1908]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="breviaedeaga" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC846B2F8DD93057269" box="[653,816,1884,1908]" italics="true" pageId="8" pageNumber="83">T. breviaedeaga</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC84708F8DD93BC7269" bold="true" box="[823,905,1884,1908]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC84708F8DD93BC7269" box="[823,905,1884,1908]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19014; 24.
<taxonomicName id="51AA4D67FFE2FFC840A4F8DD9509726E" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1179,1340,1884,1907]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="magniguttata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC840A4F8DD9509726E" box="[1179,1340,1884,1907]" italics="true" pageId="8" pageNumber="83">T. magniguttata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC8417BF8DD95A07269" bold="true" box="[1348,1429,1884,1908]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC8417BF8DD95A07269" box="[1348,1429,1884,1908]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19013; 25.
<taxonomicName id="51AA4D67FFE2FFC845A8F80192017285" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[407,564,1920,1944]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="83" phylum="Tracheophyta" rank="species" species="perparviloba" status="sp. nov.">
<emphasis id="A4DEEAF6FFE2FFC845A8F80192017285" box="[407,564,1920,1944]" italics="true" pageId="8" pageNumber="83">T. perparviloba</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE2FFC84604F80192BE7285" bold="true" box="[571,651,1920,1944]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="BFED578DFFE2FFC84604F80192BE7285" box="[571,651,1920,1944]" pageId="8" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18036.
</paragraph>
</caption>
<caption id="C2D5666CFFE3FFC944A8F95A93DB72F2" ID-DOI="http://doi.org/10.5281/zenodo.5819804" ID-Zenodo-Dep="5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="9" pageNumber="84" startId="9.[151,264,1755,1779]" targetBox="[179,1409,181,1731]" targetPageId="9">
<paragraph id="961536E4FFE3FFC944A8F95A93DB72F2" blockId="9.[151,1437,1755,2031]" pageId="9" pageNumber="84">
<emphasis id="A4DEEAF6FFE3FFC944A8F95A916073EE" bold="true" box="[151,341,1755,1779]" pageId="9" pageNumber="84">FIGURES 26–45.</emphasis>
Tegumen tubercles of
<taxonomicName id="51AA4D67FFE3FFC9467EF95A92E973EF" box="[577,732,1755,1779]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A4DEEAF6FFE3FFC9467EF95A929E73EE" box="[577,683,1755,1779]" italics="true" pageId="9" pageNumber="84">Tenupalpa</emphasis>
spp.
</taxonomicName>
26.
<taxonomicName id="51AA4D67FFE3FFC94735F95D93BB73EE" box="[778,910,1755,1779]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="angustella">
<emphasis id="A4DEEAF6FFE3FFC94735F95D93BB73EE" box="[778,910,1755,1779]" italics="true" pageId="9" pageNumber="84">T. angustella</emphasis>
</taxonomicName>
, slide no. LGE18719; 27.
<taxonomicName id="51AA4D67FFE3FFC940A0F95D950D73EE" box="[1183,1336,1755,1779]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFE3FFC940A0F95D950D73EE" box="[1183,1336,1755,1779]" italics="true" pageId="9" pageNumber="84">T. nephodesma</emphasis>
</taxonomicName>
, slide no. ZJL05343; 28.
<taxonomicName id="51AA4D67FFE3FFC9450BF88191F5720A" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[308,448,1791,1815]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="latifasciata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC9450BF88191F5720A" box="[308,448,1791,1815]" italics="true" pageId="9" pageNumber="84">T. latifasciata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC945F7F97E922C720A" bold="true" box="[456,537,1791,1815]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC945F7F97E922C720A" box="[456,537,1791,1815]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE16112; 29.
<taxonomicName id="51AA4D67FFE3FFC94715F8819394720A" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[810,929,1791,1815]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="fuscalata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC94715F8819394720A" box="[810,929,1791,1815]" italics="true" pageId="9" pageNumber="84">T. fuscalata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94797F97E93CC720A" bold="true" box="[936,1017,1791,1815]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94797F97E93CC720A" box="[936,1017,1791,1815]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18829; 30.
<taxonomicName id="51AA4D67FFE3FFC94134F88195A8720A" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1291,1437,1791,1815]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC94134F88195A8720A" box="[1291,1437,1791,1815]" italics="true" pageId="9" pageNumber="84">T. immaculata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC944A8F8A290DC7226" bold="true" box="[151,233,1827,1851]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC944A8F8A290DC7226" box="[151,233,1827,1851]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18025; 31.
<taxonomicName id="51AA4D67FFE3FFC945C3F8A5924E7226" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[508,635,1827,1851]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="isabellina" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC945C3F8A5924E7226" box="[508,635,1827,1851]" italics="true" pageId="9" pageNumber="84">T. isabellina</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC946BDF8A292E17226" bold="true" box="[642,724,1827,1851]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC946BDF8A292E17226" box="[642,724,1827,1851]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE17282; 32.
<taxonomicName id="51AA4D67FFE3FFC947D8F8A5946B7226" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[999,1118,1827,1851]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="angulosa" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC947D8F8A5946B7226" box="[999,1118,1827,1851]" italics="true" pageId="9" pageNumber="84">T. angulosa</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94059F8A2948D7226" bold="true" box="[1126,1208,1827,1851]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94059F8A2948D7226" box="[1126,1208,1827,1851]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE17270; 33.
<taxonomicName id="51AA4D67FFE3FFC94482F8C9912E7242" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[189,283,1863,1887]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="falcata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC94482F8C9912E7242" box="[189,283,1863,1887]" italics="true" pageId="9" pageNumber="84">T. falcata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC9451EF8C691447242" bold="true" box="[289,369,1863,1887]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC9451EF8C691447242" box="[289,369,1863,1887]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18843; 34.
<taxonomicName id="51AA4D67FFE3FFC94643F8C993247242" baseAuthorityName="Kyaw, Yagi &amp; Hirowatari" baseAuthorityYear="2019" box="[636,785,1863,1887]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFE3FFC94643F8C993247242" box="[636,785,1863,1887]" italics="true" pageId="9" pageNumber="84">T. flavitermina</emphasis>
</taxonomicName>
, slide no. LGE18842; 35.
<taxonomicName id="51AA4D67FFE3FFC94024F8C994B27242" baseAuthorityName="Omelko" baseAuthorityYear="1984" box="[1051,1159,1863,1887]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFE3FFC94024F8C994B27242" box="[1051,1159,1863,1887]" italics="true" pageId="9" pageNumber="84">T. biformis</emphasis>
</taxonomicName>
, slide no. LLJ15140; 36.
<taxonomicName id="51AA4D67FFE3FFC941B5F8C990C9729E" baseAuthorityName="Omelko" baseAuthorityYear="1984" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="venustalis">
<emphasis id="A4DEEAF6FFE3FFC941B5F8C990C9729E" italics="true" pageId="9" pageNumber="84">T. venustalis</emphasis>
</taxonomicName>
, slide no. LGE18493; 37.
<taxonomicName id="51AA4D67FFE3FFC94636F8ED925B729E" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[521,622,1900,1923]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC94636F8ED925B729E" box="[521,622,1900,1923]" italics="true" pageId="9" pageNumber="84">T. acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC9464BF8EA92F0729E" bold="true" box="[628,709,1899,1923]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC9464BF8EA92F0729E" box="[628,709,1899,1923]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18030; 38.
<taxonomicName id="51AA4D67FFE3FFC947EEF8ED9451729E" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[977,1124,1899,1923]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="crassimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC947EEF8ED9451729E" box="[977,1124,1899,1923]" italics="true" pageId="9" pageNumber="84">T. crassimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94054F8EA948E729E" bold="true" box="[1131,1211,1899,1923]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94054F8EA948E729E" box="[1131,1211,1899,1923]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18015; 39.
<taxonomicName id="51AA4D67FFE3FFC94483F811917072BA" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[188,325,1935,1959]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="acutimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC94483F811917072BA" box="[188,325,1935,1959]" italics="true" pageId="9" pageNumber="84">T. acutimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94575F80E91AF72BA" bold="true" box="[330,410,1935,1959]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94575F80E91AF72BA" box="[330,410,1935,1959]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE16247; 40.
<taxonomicName id="51AA4D67FFE3FFC9469CF811937172BA" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[675,836,1935,1959]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="breviaedeaga" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC9469CF811937172BA" box="[675,836,1935,1959]" italics="true" pageId="9" pageNumber="84">T. breviaedeaga</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94776F80E93AC72BA" bold="true" box="[841,921,1935,1959]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94776F80E93AC72BA" box="[841,921,1935,1959]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19014; 41.
<taxonomicName id="51AA4D67FFE3FFC9409EF811957572BA" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1185,1344,1936,1959]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="magniguttata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC9409EF811957572BA" box="[1185,1344,1936,1959]" italics="true" pageId="9" pageNumber="84">T. magniguttata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94179F80E95A072BA" bold="true" box="[1350,1429,1935,1959]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94179F80E95A072BA" box="[1350,1429,1935,1959]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19013; 42.
<taxonomicName id="51AA4D67FFE3FFC945A8F835920172D6" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[407,564,1971,1995]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="perparviloba" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC945A8F835920172D6" box="[407,564,1971,1995]" italics="true" pageId="9" pageNumber="84">T. perparviloba</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94605F83292BE72D6" bold="true" box="[570,651,1971,1995]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94605F83292BE72D6" box="[570,651,1971,1995]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE18036; 43.
<taxonomicName id="51AA4D67FFE3FFC947A8F835941372D6" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[919,1062,1971,1995]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="uncativalva" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC947A8F835941372D6" box="[919,1062,1971,1995]" italics="true" pageId="9" pageNumber="84">T. uncativalva</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94012F832944872D6" bold="true" box="[1069,1149,1971,1995]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94012F832944872D6" box="[1069,1149,1971,1995]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19015; 44.
<taxonomicName id="51AA4D67FFE3FFC941B5F83590EC72F2" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="zhengi" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC941B5F83590EC72F2" italics="true" pageId="9" pageNumber="84">T. zhengi</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC944DFF856910572F2" bold="true" box="[224,304,2007,2031]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC944DFF856910572F2" box="[224,304,2007,2031]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19001; 45.
<taxonomicName id="51AA4D67FFE3FFC94602F859928372F2" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[573,694,2007,2031]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="84" phylum="Tracheophyta" rank="species" species="dissimilis" status="sp. nov.">
<emphasis id="A4DEEAF6FFE3FFC94602F859928372F2" box="[573,694,2007,2031]" italics="true" pageId="9" pageNumber="84">T. dissimilis</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE3FFC94683F856933872F2" bold="true" box="[700,781,2007,2031]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="BFED578DFFE3FFC94683F856933872F2" box="[700,781,2007,2031]" pageId="9" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, slide no. LGE19007.
</paragraph>
</caption>
<caption id="C2D5666CFFE0FFCA44A8F816953A72E5" ID-DOI="http://doi.org/10.5281/zenodo.5819808" ID-Zenodo-Dep="5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="10" pageNumber="85" startId="10.[151,264,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="10">
<paragraph id="961536E4FFE0FFCA44A8F816953A72E5" blockId="10.[151,1436,1943,2040]" pageId="10" pageNumber="85">
<emphasis id="A4DEEAF6FFE0FFCA44A8F816916172B2" bold="true" box="[151,340,1943,1967]" pageId="10" pageNumber="85">FIGURES 46–53.</emphasis>
Adults of
<taxonomicName id="51AA4D67FFE0FFCA4582F819926272B2" box="[445,599,1944,1968]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A4DEEAF6FFE0FFCA4582F819921272AD" box="[445,551,1944,1968]" italics="true" pageId="10" pageNumber="85">Tenupalpa</emphasis>
spp.
</taxonomicName>
46.
<taxonomicName id="51AA4D67FFE0FFCA46BEF819933672AD" box="[641,771,1944,1968]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="angustella">
<emphasis id="A4DEEAF6FFE0FFCA46BEF819933672AD" box="[641,771,1944,1968]" italics="true" pageId="10" pageNumber="85">T. angustella</emphasis>
</taxonomicName>
, male; 47.
<taxonomicName id="51AA4D67FFE0FFCA474DF819943C72AD" box="[882,1033,1944,1968]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFE0FFCA474DF819943C72AD" box="[882,1033,1944,1968]" italics="true" pageId="10" pageNumber="85">T. nephodesma</emphasis>
</taxonomicName>
, male; 48.
<taxonomicName id="51AA4D67FFE0FFCA4047F819953672AD" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1144,1283,1944,1968]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="latifasciata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE0FFCA4047F819953672AD" box="[1144,1283,1944,1968]" italics="true" pageId="10" pageNumber="85">T. latifasciata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE0FFCA4137F819956D72AD" bold="true" box="[1288,1368,1944,1968]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="BFED578DFFE0FFCA4137F819956D72AD" box="[1288,1368,1944,1968]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 49.
<taxonomicName id="51AA4D67FFE0FFCA451EF83D91A372C9" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[289,406,1980,2004]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="fuscalata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE0FFCA451EF83D91A372C9" box="[289,406,1980,2004]" italics="true" pageId="10" pageNumber="85">T. fuscalata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE0FFCA45A4F83D91DE72C9" bold="true" box="[411,491,1980,2004]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="BFED578DFFE0FFCA45A4F83D91DE72C9" box="[411,491,1980,2004]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, paratype; 50.
<taxonomicName id="51AA4D67FFE0FFCA46EFF83D935572C9" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[720,864,1980,2004]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE0FFCA46EFF83D935572C9" box="[720,864,1980,2004]" italics="true" pageId="10" pageNumber="85">T. immaculata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE0FFCA475AF83D938072C9" bold="true" box="[869,949,1980,2004]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="BFED578DFFE0FFCA475AF83D938072C9" box="[869,949,1980,2004]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 51.
<taxonomicName id="51AA4D67FFE0FFCA40B7F83D953172C9" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1160,1284,1980,2004]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="isabellina" status="sp. nov.">
<emphasis id="A4DEEAF6FFE0FFCA40B7F83D953172C9" box="[1160,1284,1980,2004]" italics="true" pageId="10" pageNumber="85">T. isabellina</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE0FFCA4136F83D956C72C9" bold="true" box="[1289,1369,1980,2004]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="BFED578DFFE0FFCA4136F83D956C72C9" box="[1289,1369,1980,2004]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 52.
<taxonomicName id="51AA4D67FFE0FFCA451CF86191AC72E5" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[291,409,2016,2040]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="angulosa" status="sp. nov.">
<emphasis id="A4DEEAF6FFE0FFCA451CF86191AC72E5" box="[291,409,2016,2040]" italics="true" pageId="10" pageNumber="85">T. angulosa</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE0FFCA45A0F86191C572E5" bold="true" box="[415,496,2016,2040]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="BFED578DFFE0FFCA45A0F86191C572E5" box="[415,496,2016,2040]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 53.
<taxonomicName id="51AA4D67FFE0FFCA46F8F861931372E5" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[711,806,2016,2040]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="85" phylum="Tracheophyta" rank="species" species="falcata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE0FFCA46F8F861931372E5" box="[711,806,2016,2040]" italics="true" pageId="10" pageNumber="85">T. falcata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE0FFCA4713F861934872E5" bold="true" box="[812,893,2016,2040]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="BFED578DFFE0FFCA4713F861934872E5" box="[812,893,2016,2040]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype (Scale bars = 0.2 mm).
</paragraph>
</caption>
<caption id="C2D5666CFFE1FFCB44A8F81694E272E5" ID-DOI="http://doi.org/10.5281/zenodo.5819814" ID-Zenodo-Dep="5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="11" pageNumber="86" startId="11.[151,264,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="11">
<paragraph id="961536E4FFE1FFCB44A8F81694E272E5" blockId="11.[151,1437,1943,2040]" pageId="11" pageNumber="86">
<emphasis id="A4DEEAF6FFE1FFCB44A8F816916372B2" bold="true" box="[151,342,1943,1967]" pageId="11" pageNumber="86">FIGURES 54–61.</emphasis>
Adults of
<taxonomicName id="51AA4D67FFE1FFCB45FAF819925772B2" box="[453,610,1944,1968]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A4DEEAF6FFE1FFCB45FAF819921A72AD" box="[453,559,1944,1968]" italics="true" pageId="11" pageNumber="86">Tenupalpa</emphasis>
spp.
</taxonomicName>
54.
<taxonomicName id="51AA4D67FFE1FFCB46AFF819931372AD" baseAuthorityName="Kyaw, Yagi &amp; Hirowatari" baseAuthorityYear="2019" box="[656,806,1944,1968]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFE1FFCB46AFF819931372AD" box="[656,806,1944,1968]" italics="true" pageId="11" pageNumber="86">T. flavitermina</emphasis>
</taxonomicName>
, male; 55.
<taxonomicName id="51AA4D67FFE1FFCB47A3F819943F72AD" baseAuthorityName="Omelko" baseAuthorityYear="1984" box="[924,1034,1944,1968]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFE1FFCB47A3F819943F72AD" box="[924,1034,1944,1968]" italics="true" pageId="11" pageNumber="86">T. biformis</emphasis>
</taxonomicName>
, male; 56.
<taxonomicName id="51AA4D67FFE1FFCB40BEF819953572AD" baseAuthorityName="Omelko" baseAuthorityYear="1984" box="[1153,1280,1944,1968]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="venustalis">
<emphasis id="A4DEEAF6FFE1FFCB40BEF819953572AD" box="[1153,1280,1944,1968]" italics="true" pageId="11" pageNumber="86">T. venustalis</emphasis>
</taxonomicName>
, female; 57.
<taxonomicName id="51AA4D67FFE1FFCB41B5F81990D672CE" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE1FFCB41B5F81990D672CE" italics="true" pageId="11" pageNumber="86">T. acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE1FFCB44D5F83D910F72C9" bold="true" box="[234,314,1980,2004]" pageId="11" pageNumber="86">
<taxonomicNameLabel id="BFED578DFFE1FFCB44D5F83D910F72C9" box="[234,314,1980,2004]" pageId="11" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 58.
<taxonomicName id="51AA4D67FFE1FFCB462EF83D929172C9" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[529,676,1980,2004]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="crassimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFE1FFCB462EF83D929172C9" box="[529,676,1980,2004]" italics="true" pageId="11" pageNumber="86">T. crassimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE1FFCB4694F83D92CE72C9" bold="true" box="[683,763,1980,2004]" pageId="11" pageNumber="86">
<taxonomicNameLabel id="BFED578DFFE1FFCB4694F83D92CE72C9" box="[683,763,1980,2004]" pageId="11" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 59.
<taxonomicName id="51AA4D67FFE1FFCB47EDF83D946972C9" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[978,1116,1980,2004]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="acutimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFE1FFCB47EDF83D946972C9" box="[978,1116,1980,2004]" italics="true" pageId="11" pageNumber="86">T. acutimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE1FFCB405DF83D948672C9" bold="true" box="[1122,1203,1980,2004]" pageId="11" pageNumber="86">
<taxonomicNameLabel id="BFED578DFFE1FFCB405DF83D948672C9" box="[1122,1203,1980,2004]" pageId="11" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 60.
<taxonomicName id="51AA4D67FFE1FFCB41B5F83D911572E5" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="breviaedeaga" status="sp. nov.">
<emphasis id="A4DEEAF6FFE1FFCB41B5F83D911572E5" italics="true" pageId="11" pageNumber="86">T. breviaedeaga</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE1FFCB4519F861914272E5" bold="true" box="[294,375,2016,2040]" pageId="11" pageNumber="86">
<taxonomicNameLabel id="BFED578DFFE1FFCB4519F861914272E5" box="[294,375,2016,2040]" pageId="11" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 61.
<taxonomicName id="51AA4D67FFE1FFCB4671F86192DB72EA" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[590,750,2016,2039]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="86" phylum="Tracheophyta" rank="species" species="magniguttata" status="sp. nov.">
<emphasis id="A4DEEAF6FFE1FFCB4671F86192DB72EA" box="[590,750,2016,2039]" italics="true" pageId="11" pageNumber="86">T. magniguttata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE1FFCB46CBF861937072E5" bold="true" box="[756,837,2016,2040]" pageId="11" pageNumber="86">
<taxonomicNameLabel id="BFED578DFFE1FFCB46CBF861937072E5" box="[756,837,2016,2040]" pageId="11" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype (Scale bars = 0.2 mm).
</paragraph>
</caption>
<caption id="C2D5666CFFE6FFCC44A8FBFC94D571A7" ID-DOI="http://doi.org/10.5281/zenodo.5819816" ID-Zenodo-Dep="5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="12" pageNumber="87" startId="12.[151,264,1149,1173]" targetBox="[151,1435,181,1125]" targetPageId="12">
<paragraph id="961536E4FFE6FFCC44A8FBFC94D571A7" blockId="12.[151,1436,1149,1210]" pageId="12" pageNumber="87">
<emphasis id="A4DEEAF6FFE6FFCC44A8FBFC91617188" bold="true" box="[151,340,1149,1173]" pageId="12" pageNumber="87">FIGURES 62–65.</emphasis>
Adults of
<taxonomicName id="51AA4D67FFE6FFCC4580FBFF926F7188" box="[447,602,1150,1174]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="A4DEEAF6FFE6FFCC4580FBFF921C718B" box="[447,553,1150,1174]" italics="true" pageId="12" pageNumber="87">Tenupalpa</emphasis>
spp.
</taxonomicName>
62.
<taxonomicName id="51AA4D67FFE6FFCC46BAFBFF933C718B" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[645,777,1150,1174]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="perparviloba" status="sp. nov.">
<emphasis id="A4DEEAF6FFE6FFCC46BAFBFF933C718B" box="[645,777,1150,1174]" italics="true" pageId="12" pageNumber="87">perparviloba</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE6FFCC4730FBFF9355718B" bold="true" box="[783,864,1150,1174]" pageId="12" pageNumber="87">
<taxonomicNameLabel id="BFED578DFFE6FFCC4730FBFF9355718B" box="[783,864,1150,1174]" pageId="12" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 63.
<taxonomicName id="51AA4D67FFE6FFCC4009FBFF94F0718B" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1078,1221,1150,1174]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="uncativalva" status="sp. nov.">
<emphasis id="A4DEEAF6FFE6FFCC4009FBFF94F0718B" box="[1078,1221,1150,1174]" italics="true" pageId="12" pageNumber="87">T. uncativalva</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE6FFCC40F3FBFF9529718B" bold="true" box="[1228,1308,1150,1174]" pageId="12" pageNumber="87">
<taxonomicNameLabel id="BFED578DFFE6FFCC40F3FBFF9529718B" box="[1228,1308,1150,1174]" pageId="12" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 64.
<taxonomicName id="51AA4D67FFE6FFCC44C9FB23916471A7" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[246,337,1186,1210]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="zhengi" status="sp. nov.">
<emphasis id="A4DEEAF6FFE6FFCC44C9FB23916471A7" box="[246,337,1186,1210]" italics="true" pageId="12" pageNumber="87">T. zhengi</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE6FFCC4567FB23919D71A7" bold="true" box="[344,424,1186,1210]" pageId="12" pageNumber="87">
<taxonomicNameLabel id="BFED578DFFE6FFCC4567FB23919D71A7" box="[344,424,1186,1210]" pageId="12" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype; 65.
<taxonomicName id="51AA4D67FFE6FFCC4640FB2392CD71A7" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[639,760,1186,1210]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="87" phylum="Tracheophyta" rank="species" species="dissimilis" status="sp. nov.">
<emphasis id="A4DEEAF6FFE6FFCC4640FB2392CD71A7" box="[639,760,1186,1210]" italics="true" pageId="12" pageNumber="87">T. dissimilis</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFE6FFCC46C1FB23937A71A7" bold="true" box="[766,847,1186,1210]" pageId="12" pageNumber="87">
<taxonomicNameLabel id="BFED578DFFE6FFCC46C1FB23937A71A7" box="[766,847,1186,1210]" pageId="12" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype (Scale bars = 0.2 mm).
</paragraph>
</caption>
<subSubSection id="DEB0656FFFE6FFCC44F8FB6991EE703B" pageId="12" pageNumber="87" type="etymology">
<paragraph id="961536E4FFE6FFCC44F8FB6991EE703B" blockId="12.[151,1436,1256,1318]" pageId="12" pageNumber="87">
<emphasis id="A4DEEAF6FFE6FFCC44F8FB69917D701F" bold="true" box="[199,328,1256,1282]" pageId="12" pageNumber="87">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFE6FFCC4777FB69934B701F" box="[840,894,1256,1282]" italics="true" pageId="12" pageNumber="87">latus</emphasis>
(broad) and
<emphasis id="A4DEEAF6FFE6FFCC402EFB699440701F" box="[1041,1141,1256,1282]" italics="true" pageId="12" pageNumber="87">fasciatus</emphasis>
(fasciate), referring to the broad fascia of the forewing.
</paragraph>
</subSubSection>
</treatment>
</document>

281
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@ -0,0 +1,281 @@
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<mods:title id="F5A81005853D7715F0F7FB28A4023D12">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<mods:namePart id="AD56B641E1A70A7E81222D4550EC41A6">Lee, Ga-Eun</mods:namePart>
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<treatment id="1E0387F2FFEFFFC644A8FABB93A07608" ID-DOI="http://doi.org/10.5281/zenodo.5819818" ID-GBIF-Taxon="191928508" ID-Zenodo-Dep="5819818" LSID="urn:lsid:plazi:treatment:1E0387F2FFEFFFC644A8FABB93A07608" httpUri="http://treatment.plazi.org/id/1E0387F2FFEFFFC644A8FABB93A07608" lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80">
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<paragraph id="961536E4FFEFFFC544A8FABB92B17049" blockId="5.[151,644,1338,1401]" box="[151,644,1338,1365]" pageId="5" pageNumber="80">
<heading id="CD5D8188FFEFFFC544A8FABB92B17049" bold="true" box="[151,644,1338,1365]" fontSize="11" level="1" pageId="5" pageNumber="80" reason="1">
<taxonomicName id="51AA4D67FFEFFFC544A8FABB92B17049" authority="(Meyrick, 1918)" baseAuthorityName="Meyrick" baseAuthorityYear="1918" box="[151,644,1338,1365]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC544A8FABB92B17049" bold="true" box="[151,644,1338,1365]" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544A8FABB91837048" bold="true" box="[151,438,1338,1365]" italics="true" pageId="5" pageNumber="80">Tenupalpa nephodesma</emphasis>
(
<bibRefCitation id="F23B4B15FFEFFFC545FAFABB924E7049" author="Meyrick, E." box="[453,635,1338,1364]" pageId="5" pageNumber="80" pagination="97 - 160" refId="ref23843" refString="Meyrick, E. (1918) Exotic Microlepidoptera II. Thornhanger, Marlborough, Wilts, pp. 97 - 160." type="book chapter" year="1918">Meyrick, 1918</bibRefCitation>
)
</emphasis>
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<subSubSection id="DEB0656FFFEFFFC544A8FADE91DE7064" box="[151,491,1374,1401]" pageId="5" pageNumber="80" type="description">
<paragraph id="961536E4FFEFFFC544A8FADE91DE7064" blockId="5.[151,644,1338,1401]" box="[151,491,1374,1401]" pageId="5" pageNumber="80">
(
<figureCitation id="0E912A61FFEFFFC544A0FADE90DD7064" box="[159,232,1375,1401]" captionStart="FIGURES 3–6" captionStartId="4.[151,264,1894,1918]" captionTargetBox="[161,1422,453,1848]" captionTargetId="figure-154@4.[151,1435,453,1871]" captionTargetPageId="4" captionText="FIGURES 3–6. Abdominal and genital structures of Tenupalpa spp. 3, distal segments of the abdomen and male genitalia of T. angustella in external view; 4, denuded male 8th abdominal segment of T. angustella (image from Lee et al., 2021: Fig. 7I), slide no. LGE18719; 5, male genitalia of T. angustella, slide no. LGE16107; 6, female genitalia of T. nephodesma, slide no. LGE19022 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819798" httpUri="https://zenodo.org/record/5819798/files/figure.png" pageId="5" pageNumber="80">Figs 6</figureCitation>
,
<figureCitation id="0E912A61FFEFFFC544CBFADF91277064" box="[244,274,1374,1401]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="5" pageNumber="80">15</figureCitation>
,
<figureCitation id="0E912A61FFEFFFC54521FADE91097064" box="[286,316,1375,1401]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="5" pageNumber="80">27</figureCitation>
,
<figureCitation id="0E912A61FFEFFFC54577FADE91537064" box="[328,358,1375,1401]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="5" pageNumber="80">47</figureCitation>
,
<figureCitation id="0E912A61FFEFFFC5454DFADE91A57064" box="[370,400,1375,1401]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="5" pageNumber="80">66</figureCitation>
,
<figureCitation id="0E912A61FFEFFFC545A3FADF91D67064" box="[412,483,1374,1401]" captionStart="FIGURES 95–97" captionStartId="31.[151,264,1868,1892]" captionTargetBox="[151,1435,344,1845]" captionTargetId="figure-90@31.[151,1435,344,1845]" captionTargetPageId="31" captionText="FIGURES 95–97. Type specimen of Tenupalpa nephodesma deposited in NHMUK. 95. Holotype, female; 96. Label data; 97. Female genitalia (holotype), slide no. J.F.G.C. 8367, 97a, detail of signum." figureDoi="http://doi.org/10.5281/zenodo.5819838" httpUri="https://zenodo.org/record/5819838/files/figure.png" pageId="5" pageNumber="80">95–97</figureCitation>
)
</paragraph>
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<subSubSection id="DEB0656FFFEFFFC544A8FA22925070E4" pageId="5" pageNumber="80" type="reference_group">
<paragraph id="961536E4FFEFFFC544A8FA22916870C4" blockId="5.[151,1437,1443,1529]" pageId="5" pageNumber="80">
<treatmentCitationGroup id="B6BA11CAFFEFFFC544A8FA22916870C4" pageId="5" pageNumber="80">
<treatmentCitation id="170B10F5FFEFFFC544A8FA22926C70A6" author="Meyrick, E." box="[151,601,1443,1467]" page="103" pageId="5" pageNumber="80" year="1918">
<taxonomicName id="51AA4D67FFEFFFC544A8FA22926C70A6" ID-CoL="56HNS" authority="Meyrick, 1918: 103" authorityName="Meyrick" authorityPageNumber="103" authorityYear="1918" box="[151,601,1443,1467]" class="Insecta" family="Gelechiidae" genus="Thiotricha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC544A8FA2291BD70A6" box="[151,392,1443,1467]" italics="true" pageId="5" pageNumber="80">Thiotricha nephodesma</emphasis>
<bibRefCitation id="F23B4B15FFEFFFC545B1FA22926C70A6" author="Meyrick, E." box="[398,601,1443,1467]" pageId="5" pageNumber="80" pagination="97 - 160" refId="ref23843" refString="Meyrick, E. (1918) Exotic Microlepidoptera II. Thornhanger, Marlborough, Wilts, pp. 97 - 160." type="book chapter" year="1918">Meyrick, 1918: 103</bibRefCitation>
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;
<treatmentCitation id="170B10F5FFEFFFC5465BFA22932E70A6" author="Gaede, M." box="[612,795,1443,1467]" page="306" pageId="5" pageNumber="80" year="1937">
<bibRefCitation id="F23B4B15FFEFFFC5465BFA22932E70A6" author="Gaede, M." box="[612,795,1443,1467]" pageId="5" pageNumber="80" pagination="1 - 630" refId="ref23337" refString="Gaede, M. (1937) Gelechiidae. Lepidopterorum Catalogus, 79, 1 - 630." type="journal article" year="1937">Gaede, 1937: 306</bibRefCitation>
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;
<treatmentCitation id="170B10F5FFEFFFC5471AFA2293EB70A6" author="Clarke, J. F. G." box="[805,990,1443,1467]" page="463" pageId="5" pageNumber="80" year="1969">
<bibRefCitation id="F23B4B15FFEFFFC5471AFA2293EB70A6" author="Clarke, J. F. G." box="[805,990,1443,1467]" pageId="5" pageNumber="80" refId="ref23257" refString="Clarke, J. F. G. (1969) Catalogue of the type specimens of Microlepidoptera in the British Museum (Natural History) described by Edward Meyrick. Vol. 7. Trustees of the British Museum, London, 531 pp., 265 pls." type="book" year="1969">Clarke, 1969: 463</bibRefCitation>
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<treatmentCitation id="170B10F5FFEFFFC547D6FA2294EE70A6" author="Ivinskis P. P. &amp; Piskunov, V. I. &amp; Emeliyanov, I. M." box="[1001,1243,1443,1467]" page="39" pageId="5" pageNumber="80" year="1984">
<bibRefCitation id="F23B4B15FFEFFFC547D6FA2294EE70A6" author="Ivinskis P. P. &amp; Piskunov, V. I. &amp; Emeliyanov, I. M." box="[1001,1243,1443,1467]" pageId="5" pageNumber="80" pagination="37 - 44" refId="ref23428" refString="Ivinskis P. P., Piskunov, V. I. &amp; Emeliyanov, I. M. (1984) [Review of the fauna of Gelechiid moths of the Thiotricha Meyrick, 1886 genus and its place in the system of Lepidoptera: Gelechiidae, Dichomerinae]. Trudy AN Lithuanian SSR, 2 (86), 37 - 44. [in Russian]" type="journal article" year="1984">
Ivinskis
<emphasis id="A4DEEAF6FFEFFFC5407FFA25945970A6" box="[1088,1132,1443,1467]" italics="true" pageId="5" pageNumber="80">et al</emphasis>
., 1984: 39
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. TL:
<collectingRegion id="546EF806FFEFFFC5412FFA22956D70A6" box="[1296,1368,1443,1467]" country="India" name="Assam" pageId="5" pageNumber="80">Assam</collectingRegion>
,
<collectingCountry id="EEBD7674FFEFFFC5415DFA2295AC70A6" box="[1378,1433,1443,1467]" name="India" pageId="5" pageNumber="80">India</collectingCountry>
. TD: NHMUK.
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</paragraph>
<paragraph id="961536E4FFEFFFC544A8FA61925070E4" blockId="5.[151,1437,1443,1529]" box="[151,613,1504,1529]" pageId="5" pageNumber="80">
<treatmentCitationGroup id="B6BA11CAFFEFFFC544A8FA61925070E4" box="[151,613,1504,1529]" pageId="5" pageNumber="80">
<treatmentCitation id="170B10F5FFEFFFC544A8FA61925470E4" author="Lee, G. - E. &amp; Han, T. M. &amp; Park, H. C. &amp; Qi, M. J. &amp; Li, H. H." box="[151,609,1504,1529]" page="371" pageId="5" pageNumber="80" year="2021">
<taxonomicName id="51AA4D67FFEFFFC544A8FA61925470E4" ID-CoL="98YJ5" authority="Lee et al., 2021: 371" authorityName="Lee" authorityPageNumber="371" authorityYear="2021" box="[151,609,1504,1529]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC544A8FA6191B270E5" box="[151,391,1504,1528]" italics="true" pageId="5" pageNumber="80">Tenupalpa nephodesma</emphasis>
<bibRefCitation id="F23B4B15FFEFFFC545B2FA60925470E4" author="Lee, G. - E. &amp; Han, T. M. &amp; Park, H. C. &amp; Qi, M. J. &amp; Li, H. H." box="[397,609,1504,1529]" pageId="5" pageNumber="80" pagination="357 - 379" refId="ref23692" refString="Lee, G. - E., Han, T. M., Park, H. C., Qi, M. J. &amp; Li, H. H. (2021) A phylogeny of the subfamily Thiotrichinae (Lepidoptera: Gel- echiidae) with a revision of the generic classification based on molecular and morphological analyses. Systematic Entomology, 46, 357 - 379. https: // doi. org / 10.1111 / syen. 12466" type="journal article" year="2021">
Lee
<emphasis id="A4DEEAF6FFEFFFC54586FA6091D970E5" box="[441,492,1504,1528]" italics="true" pageId="5" pageNumber="80">et al.</emphasis>
, 2021: 371
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<paragraph id="961536E4FFEFFFC544A8F9A6912E7378" blockId="5.[151,1437,1574,2033]" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544A8F9A6917C735D" bold="true" box="[151,329,1574,1601]" pageId="5" pageNumber="80">Type material.</emphasis>
<materialsCitation id="26C23CB9FFEFFFC5456AF9A6912E7378" ID-GBIF-Occurrence="3427696324" collectingDate="1908-09" collectionCode="NHMUK" collectorName="E. Meyrick &amp; J.F.G. Clarke" country="India" location="Khasi Hills" pageId="5" pageNumber="80" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="49118846FFEFFFC5456AF9A69188735C" box="[341,445,1575,1601]" pageId="5" pageNumber="80" type="holotype">Holotype</typeStatus>
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<collectingCountry id="EEBD7674FFEFFFC545CFF9A69277735C" box="[496,578,1575,1601]" name="India" pageId="5" pageNumber="80">INDIA</collectingCountry>
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<location id="9375603FFFEFFFC5466EF9A692E2735C" LSID="urn:lsid:plazi:treatment:1E0387F2FFEFFFC644A8FABB93A07608:9375603FFFEFFFC5466EF9A692E2735C" box="[593,727,1575,1601]" country="India" name="Khasi Hills" pageId="5" pageNumber="80">Khasi Hills</location>
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<date id="E2141024FFEFFFC546D8F9A6937F735C" box="[743,842,1575,1601]" pageId="5" pageNumber="80" value="1908-09">
<collectingDate id="F250E9CCFFEFFFC546D8F9A6937F735C" box="[743,842,1575,1601]" pageId="5" pageNumber="80" value="1908-09">ix. 1908</collectingDate>
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, leg.
<collectorName id="3B5F5332FFEFFFC547B1F9A69421735C" box="[910,1044,1575,1601]" pageId="5" pageNumber="80">E. Meyrick</collectorName>
, genitalia slide no. J.
<collectorName id="3B5F5332FFEFFFC54131F9A6956C735C" box="[1294,1369,1575,1601]" name="J.F.G. Clarke" pageId="5" pageNumber="80">F.G.C.</collectorName>
8367 (
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).
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<paragraph id="961536E4FFEFFFC544F8F9EF93A173CC" blockId="5.[151,1437,1574,2033]" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544F8F9EF92007395" bold="true" box="[199,565,1646,1672]" pageId="5" pageNumber="80">Additional material examined:</emphasis>
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<emphasis id="A4DEEAF6FFEFFFC54604F9EF93B27395" bold="true" box="[571,903,1646,1672]" pageId="5" pageNumber="80">
<collectingCountry id="EEBD7674FFEFFFC54604F9EF92A87395" box="[571,669,1646,1672]" name="China" pageId="5" pageNumber="80">CHINA</collectingCountry>
:
<collectingRegion id="546EF806FFEFFFC54698F9EF93B77395" box="[679,898,1646,1672]" country="China" name="Guizhou" pageId="5" pageNumber="80">Guizhou Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFEFFFC547B2F9EE938D7394" box="[909,952,1647,1673]" pageId="5" pageNumber="80" type="male">1 ♂</specimenCount>
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),
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,
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<collectingDate id="F250E9CCFFEFFFC544A8F912912F73B0" box="[151,282,1683,1709]" pageId="5" pageNumber="80" value="2001-07-31">31.vii.2001</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFEFFFC54569F91291E473B0" box="[342,465,1683,1709]" pageId="5" pageNumber="80">Houhun Li</collectorName>
and
<collectorName id="3B5F5332FFEFFFC54637F91292AC73B0" box="[520,665,1683,1709]" pageId="5" pageNumber="80">Xinpu Wang</collectorName>
, genitalia slide no. ZJL05343
</materialsCitation>
;
<materialsCitation id="26C23CB9FFEFFFC547CAF91293A573CC" ID-GBIF-Occurrence="3427696313" accessionNumber="LGE19022" collectingDate="2004-08-19" collectorName="Yunli Xiao" country="China" county="Daozhen County" elevation="1300" location="Chengjiashan" pageId="5" pageNumber="80" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou">
<specimenCount id="80ACFD6DFFEFFFC547CAF912941573B0" box="[1013,1056,1683,1709]" pageId="5" pageNumber="80" type="female">1 ♀</specimenCount>
,
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,
<quantity id="51529B01FFEFFFC540EAF912951973B0" box="[1237,1324,1683,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="80" unit="m" value="1300.0">
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</quantity>
,
<collectingCounty id="7F744E68FFEFFFC54106F91290DE73CC" pageId="5" pageNumber="80">Daozhen County</collectingCounty>
,
<date id="E2141024FFEFFFC544C9F93691B573CC" box="[246,384,1719,1745]" pageId="5" pageNumber="80" value="2004-08-19">
<collectingDate id="F250E9CCFFEFFFC544C9F93691B573CC" box="[246,384,1719,1745]" pageId="5" pageNumber="80" value="2004-08-19">19.viii.2004</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFEFFFC54585F936920273CC" box="[442,567,1719,1745]" pageId="5" pageNumber="80">Yunli Xiao</collectorName>
, genitalia slide no. LGE19022
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFEFFFC644F8F95B916D77D0" lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80" type="diagnosis">
<paragraph id="961536E4FFEFFFC544F8F95B93A27298" blockId="5.[151,1437,1574,2033]" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544F8F95B917773E9" bold="true" box="[199,322,1754,1780]" pageId="5" pageNumber="80">Diagnosis.</emphasis>
The species is distinguished by the following characteristics: a broad transverse fascia before middle of the forewing, a fine longitudinal line arising from the fascia toward the apex, a small trapezoidal projection on the valva, and a conical-shaped anellus lobe. The female genitalia of
<taxonomicName id="51AA4D67FFEFFFC54751F8A294207220" box="[878,1045,1827,1853]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="80" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC54751F8A294207220" box="[878,1045,1827,1853]" italics="true" pageId="5" pageNumber="80">T. nephodesma</emphasis>
</taxonomicName>
can be distinguished by a long and somewhat boomerang-shaped signum in lateral view. Externally,
<taxonomicName id="51AA4D67FFEFFFC54752F8C69421727C" box="[877,1044,1863,1889]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="80" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC54752F8C69421727C" box="[877,1044,1863,1889]" italics="true" pageId="5" pageNumber="80">T. nephodesma</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="51AA4D67FFEFFFC540E3F8C69541727C" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1244,1396,1863,1889]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="80" phylum="Tracheophyta" rank="species" species="latifasciata" status="sp. nov.">
<emphasis id="A4DEEAF6FFEFFFC540E3F8C69541727C" box="[1244,1396,1863,1889]" italics="true" pageId="5" pageNumber="80">T. latifasciata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFEFFFC54144F8C690FD7298" bold="true" pageId="5" pageNumber="80">
<taxonomicNameLabel id="BFED578DFFEFFFC54144F8C690FD7298" pageId="5" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but the apex of the forewing is not darkened in
<taxonomicName id="51AA4D67FFEFFFC546D7F8EA93A57298" box="[744,912,1899,1925]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="80" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFEFFFC546D7F8EA93A57298" box="[744,912,1899,1925]" italics="true" pageId="5" pageNumber="80">T. nephodesma</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="961536E4FFEFFFC544F8F80F956172D0" blockId="5.[151,1437,1574,2033]" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544F8F80F914B72B5" bold="true" box="[199,382,1934,1960]" pageId="5" pageNumber="80">Re-description.</emphasis>
Adult (
<figureCitation id="0E912A61FFEFFFC545EFF80E921072B4" box="[464,549,1935,1961]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="5" pageNumber="80">Figs 47</figureCitation>
,
<figureCitation id="0E912A61FFEFFFC54610F80F927A72B4" box="[559,591,1934,1961]" captionStart="FIGURES 95–97" captionStartId="31.[151,264,1868,1892]" captionTargetBox="[151,1435,344,1845]" captionTargetId="figure-90@31.[151,1435,344,1845]" captionTargetPageId="31" captionText="FIGURES 95–97. Type specimen of Tenupalpa nephodesma deposited in NHMUK. 95. Holotype, female; 96. Label data; 97. Female genitalia (holotype), slide no. J.F.G.C. 8367, 97a, detail of signum." figureDoi="http://doi.org/10.5281/zenodo.5819838" httpUri="https://zenodo.org/record/5819838/files/figure.png" pageId="5" pageNumber="80">95</figureCitation>
).
<emphasis id="A4DEEAF6FFEFFFC5465EF80E92AB72B4" box="[609,670,1935,1961]" italics="true" pageId="5" pageNumber="80">Head</emphasis>
: Silvery white. Labial palpus dirty white, except palpomere III mixed with grey. Antenna with scape dirty white; flagellum dark fuscous, except base of dorsal surface dirty white.
</paragraph>
<paragraph id="961536E4FFEFFFC644F8F85691EC7400" blockId="5.[151,1437,1574,2033]" lastBlockId="6.[151,1437,151,789]" lastPageId="6" lastPageNumber="81" pageId="5" pageNumber="80">
<emphasis id="A4DEEAF6FFEFFFC544F8F856912D72EC" box="[199,280,2007,2033]" italics="true" pageId="5" pageNumber="80">Thorax</emphasis>
: Dorsum, tegula, and forewing dirty white mixed and pale brown; wingspan 9.0 mm; forewing with anterior edge of costa black in basal half, a very broad fuscous fascia between basal 1/3 and 2/3 of wing, a fine fuscous median line from outer margin of former fascia, running longitudinally toward apex; fringe scales grey. Hindwing and fringe scales grey. Legs dirty white and pale brown mixed, hind tibia broadly dark fuscous after middle and shortly dark fuscous at apex.
</paragraph>
<paragraph id="961536E4FFECFFC644F8FEA6916D77D0" blockId="6.[151,1437,151,789]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FEA69104745C" box="[199,305,295,321]" italics="true" pageId="6" pageNumber="81">Abdomen</emphasis>
: Male with coremata slightly shorter than sternum VIII; sternum VIII sub-pentagonal, rounded laterally (
<figureCitation id="0E912A61FFECFFC644EEFECA91107478" box="[209,293,330,357]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="6" pageNumber="81">Fig. 15</figureCitation>
). Male genitalia (
<figureCitation id="0E912A61FFECFFC645D3FECA920B747B" box="[492,574,331,358]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="6" pageNumber="81">Fig. 66</figureCitation>
) with uncus small, sub-pentagonal, protruded medially. Gnathos hook short, gradually narrowing to apex. Tegumen with a set of papilliform tubercles at middle (
<figureCitation id="0E912A61FFECFFC64061FEEE94817494" box="[1118,1204,367,393]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="6" pageNumber="81">Fig. 27</figureCitation>
). Valva with a small trapezoidal projection near middle of costa, apex rounded. Anellus lobe elongate-conical with a short subapical bristle, slightly curved, pointed apically. Juxta convex in basal 1/3 of inner margin. Vinculum simple. Saccus broad, slightly exceeding tegumen pedunculus. Aedeagus with basal half dilated, distal part slender with apex rounded. Female genitalia (
<figureCitation id="0E912A61FFECFFC6455DFE7E91997704" box="[354,428,511,537]" captionStart="FIGURES 3–6" captionStartId="4.[151,264,1894,1918]" captionTargetBox="[161,1422,453,1848]" captionTargetId="figure-154@4.[151,1435,453,1871]" captionTargetPageId="4" captionText="FIGURES 3–6. Abdominal and genital structures of Tenupalpa spp. 3, distal segments of the abdomen and male genitalia of T. angustella in external view; 4, denuded male 8th abdominal segment of T. angustella (image from Lee et al., 2021: Fig. 7I), slide no. LGE18719; 5, male genitalia of T. angustella, slide no. LGE16107; 6, female genitalia of T. nephodesma, slide no. LGE19022 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819798" httpUri="https://zenodo.org/record/5819798/files/figure.png" pageId="6" pageNumber="81">Figs 6</figureCitation>
,
<figureCitation id="0E912A61FFECFFC64586FE7E91E37704" box="[441,470,511,537]" captionStart="FIGURES 95–97" captionStartId="31.[151,264,1868,1892]" captionTargetBox="[151,1435,344,1845]" captionTargetId="figure-90@31.[151,1435,344,1845]" captionTargetPageId="31" captionText="FIGURES 95–97. Type specimen of Tenupalpa nephodesma deposited in NHMUK. 95. Holotype, female; 96. Label data; 97. Female genitalia (holotype), slide no. J.F.G.C. 8367, 97a, detail of signum." figureDoi="http://doi.org/10.5281/zenodo.5819838" httpUri="https://zenodo.org/record/5819838/files/figure.png" pageId="6" pageNumber="81">97</figureCitation>
) with papillae anales simple. Apophyses anteriores slightly shorter than 1/2 length of apophyses posteriores. Tergum VIII with posterior margin deeply concave at middle in V-shape, bearing a bundle of setae at the cleft; sternum VIII broadly convex anteriorly. Ostium bursae round, opening at middle of sternum VIII. Ductus bursae broad at base, constricted after base then gradually broadening toward corpus bursae; colliculum situated near base. Corpus bursae ovate, slightly shorter than ductus bursae; signum linear, situated in posterior half of corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644F8FD57945577EC" box="[199,1120,726,754]" pageId="6" pageNumber="81" type="distribution">
<paragraph id="961536E4FFECFFC644F8FD57945577EC" blockId="6.[151,1437,151,789]" box="[199,1120,726,754]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FD57916C77ED" bold="true" box="[199,345,726,752]" pageId="6" pageNumber="81">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFECFFC64558FD56919F77EC" box="[359,426,727,753]" name="China" pageId="6" pageNumber="81">China</collectingCountry>
(
<collectingRegion id="546EF806FFECFFC64586FD56922A77EF" box="[441,543,727,754]" country="China" name="Guizhou" pageId="6" pageNumber="81">Guizhou</collectingRegion>
),
<collectingCountry id="EEBD7674FFECFFC6460DFD56925A77EC" box="[562,623,727,753]" name="India" pageId="6" pageNumber="81">India</collectingCountry>
. This species is newly recorded from
<collectingCountry id="EEBD7674FFECFFC64029FD56946877EC" box="[1046,1117,727,753]" name="China" pageId="6" pageNumber="81">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFECFFC644F8FD7A93A07608" box="[199,917,763,789]" pageId="6" pageNumber="81" type="discussion">
<paragraph id="961536E4FFECFFC644F8FD7A93A07608" blockId="6.[151,1437,151,789]" box="[199,917,763,789]" pageId="6" pageNumber="81">
<emphasis id="A4DEEAF6FFECFFC644F8FD7A910E7608" bold="true" box="[199,315,763,789]" pageId="6" pageNumber="81">Remarks.</emphasis>
The male genitalia are newly illustrated in this paper.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="DC6ED20DA292BF3A4B07BB703D09472D">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<mods:identifier id="DCBAE277AC8A12A0E1897909485B4AD6" type="Zenodo-Dep">5819794</mods:identifier>
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<treatment id="1E0387F2FFF0FFDB44A8F91692FF76D5" ID-DOI="http://doi.org/10.5281/zenodo.5819854" ID-GBIF-Taxon="191928537" ID-Zenodo-Dep="5819854" LSID="urn:lsid:zoobank.org:act:749B998D-7A91-4521-BE2E-20E78171A850" httpUri="http://treatment.plazi.org/id/1E0387F2FFF0FFDB44A8F91692FF76D5" lastPageId="27" lastPageNumber="102" pageId="26" pageNumber="101">
<subSubSection id="DEB0656FFFF0FFDA44A8F91692B073AF" box="[151,645,1687,1714]" pageId="26" pageNumber="101" type="nomenclature">
<paragraph id="961536E4FFF0FFDA44A8F91692B073AF" blockId="26.[151,916,1687,1779]" box="[151,645,1687,1714]" pageId="26" pageNumber="101">
<heading id="CD5D8188FFF0FFDA44A8F91692B073AF" bold="true" box="[151,645,1687,1714]" fontSize="11" level="1" pageId="26" pageNumber="101" reason="1">
<emphasis id="A4DEEAF6FFF0FFDA44A8F91692B073AF" bold="true" box="[151,645,1687,1714]" pageId="26" pageNumber="101">
<taxonomicName id="51AA4D67FFF0FFDA44A8F916922F73AF" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,538,1687,1714]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="26" pageNumber="101" phylum="Tracheophyta" rank="species" species="acutimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFF0FFDA44A8F916919173AF" bold="true" box="[151,420,1687,1714]" italics="true" pageId="26" pageNumber="101">Tenupalpa acutimedia</emphasis>
Lee
<emphasis id="A4DEEAF6FFF0FFDA45DFF91991C073AF" bold="true" box="[480,501,1688,1714]" italics="true" pageId="26" pageNumber="101">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFF0FFDA4618F91992B073AF" box="[551,645,1688,1714]" pageId="26" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDA44A8F93D93A173EE" pageId="26" pageNumber="101" type="description">
<paragraph id="961536E4FFF0FFDA44A8F93D914E73CB" blockId="26.[151,916,1687,1779]" box="[151,379,1724,1751]" pageId="26" pageNumber="101">
(
<figureCitation id="0E912A61FFF0FFDA44A0F93D90C273CB" box="[159,247,1724,1750]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="26" pageNumber="101">Figs 22</figureCitation>
,
<figureCitation id="0E912A61FFF0FFDA453DF93D911573CB" box="[258,288,1724,1750]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="26" pageNumber="101">39</figureCitation>
,
<figureCitation id="0E912A61FFF0FFDA4513F93D917F73CA" box="[300,330,1724,1751]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="26" pageNumber="101">59</figureCitation>
,
<figureCitation id="0E912A61FFF0FFDA4569F93D914673CB" box="[342,371,1724,1750]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="26" pageNumber="101">78</figureCitation>
)
</paragraph>
<paragraph id="961536E4FFF0FFDA44A8F95A93A173EE" blockId="26.[151,916,1687,1779]" box="[151,916,1755,1779]" pageId="26" pageNumber="101">
<uri id="E23B3AE6FFF0FFDA44A8F95A93A173EE" box="[151,916,1755,1779]" pageId="26" pageNumber="101">
urn:lsid:zoobank.org:act:
<uuid id="E20C0C31FFF0FFDA45A3F95A93A173EE" box="[412,916,1755,1779]" pageId="26" pageNumber="101">749B998D-7A91-4521-BE2E-20E78171A850</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDA44A8F8A094327242" pageId="26" pageNumber="101" type="materials_examined">
<paragraph id="961536E4FFF0FFDA44A8F8A094327242" blockId="26.[151,1437,1825,2032]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44A8F8A0917F7226" bold="true" box="[151,330,1825,1851]" pageId="26" pageNumber="101">Type material.</emphasis>
<materialsCitation id="26C23CB9FFF0FFDA4569F8A094327242" ID-GBIF-Occurrence="3427696310" accessionNumber="LGE16247" collectingDate="2016-06-27" collectorName="Kaijian Teng &amp; Xiaofei Yang" country="China" elevation="1115" latitude="30.07" location="Bifengxia" longLatPrecision="734" longitude="102.97" municipality="Ya'an City" pageId="26" pageNumber="101" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="49118846FFF0FFDA4569F8A0918B7226" box="[342,446,1825,1851]" pageId="26" pageNumber="101" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFF0FFDA45F5F8A391D47221" box="[458,481,1826,1852]" pageId="26" pageNumber="101" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFF0FFDA45CDF8A0927B7226" box="[498,590,1825,1851]" name="China" pageId="26" pageNumber="101">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFF0FFDA4662F8A0931C7226" box="[605,809,1825,1851]" country="China" name="Sichuan" pageId="26" pageNumber="101">Sichuan Province</collectingRegion>
,
<collectingMunicipality id="7671AC9EFFF0FFDA4706F8A093867226" box="[825,947,1825,1851]" pageId="26" pageNumber="101">Ya'an City</collectingMunicipality>
,
<location id="9375603FFFF0FFDA47FCF8A094047226" LSID="urn:lsid:plazi:treatment:1E0387F2FFF0FFDB44A8F91692FF76D5:9375603FFFF0FFDA47FCF8A094047226" box="[963,1073,1825,1851]" country="China" latitude="30.07" longLatPrecision="734" longitude="102.97" municipality="Ya'an City" name="Bifengxia" pageId="26" pageNumber="101" stateProvince="Sichuan">Bifengxia</location>
(
<geoCoordinate id="F39E5023FFF0FFDA407AF8A0949D7226" box="[1093,1192,1825,1851]" degrees="30.07" direction="north" orientation="latitude" pageId="26" pageNumber="101" precision="555" value="30.07">30.07°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF0FFDA4087F8A095107226" box="[1208,1317,1825,1851]" degrees="102.97" direction="east" orientation="longitude" pageId="26" pageNumber="101" precision="555" value="102.97">102.97°E</geoCoordinate>
),
<quantity id="51529B01FFF0FFDA4101F8A095A27221" box="[1342,1431,1825,1852]" metricMagnitude="3" metricUnit="m" metricValue="1.115" pageId="26" pageNumber="101" unit="m" value="1115.0">
<elevation id="1D87D1D7FFF0FFDA4101F8A095A27221" box="[1342,1431,1825,1852]" metricMagnitude="3" metricUnit="m" metricValue="1.115" pageId="26" pageNumber="101" unit="m" value="1115.0">1115 m</elevation>
</quantity>
,
<date id="E2141024FFF0FFDA44A8F8C491247242" box="[151,273,1861,1887]" pageId="26" pageNumber="101" value="2016-06-27">
<collectingDate id="F250E9CCFFF0FFDA44A8F8C491247242" box="[151,273,1861,1887]" pageId="26" pageNumber="101" value="2016-06-27">27.vi.2016</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF0FFDA4572F8C491EB727D" box="[333,478,1861,1888]" pageId="26" pageNumber="101">Kaijian Teng</collectorName>
and
<collectorName id="3B5F5332FFF0FFDA462BF8C4929E727D" box="[532,683,1861,1888]" pageId="26" pageNumber="101">Xiaofei Yang</collectorName>
, genitalia slide no. LGE16247.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDA44F8F8E8942C72D6" pageId="26" pageNumber="101" type="diagnosis">
<paragraph id="961536E4FFF0FFDA44F8F8E8942C72D6" blockId="26.[151,1437,1825,2032]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44F8F8E89177729E" bold="true" box="[199,322,1897,1923]" pageId="26" pageNumber="101">Diagnosis.</emphasis>
The forewing ground color of
<taxonomicName id="51AA4D67FFF0FFDA46A8F8EB931B729E" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[663,814,1897,1923]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="26" pageNumber="101" phylum="Tracheophyta" rank="species" species="acutimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFF0FFDA46A8F8EB931B729E" box="[663,814,1897,1923]" italics="true" pageId="26" pageNumber="101">T. acutimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF0FFDA470BF8E893B8729E" bold="true" box="[820,909,1897,1923]" pageId="26" pageNumber="101">
<taxonomicNameLabel id="BFED578DFFF0FFDA470BF8E893B8729E" box="[820,909,1897,1923]" pageId="26" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is somewhat lighter and the orange distal patch smaller than in related species. Also, the hind tibia and mid- and hind tarsi are less mixed with fuscous. The male genitalia are unique in that the costa of valva has a pointed projection at middle.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDB44F8F85491A47641" lastPageId="27" lastPageNumber="102" pageId="26" pageNumber="101" type="description">
<paragraph id="961536E4FFF0FFDB44F8F85490CA75E5" blockId="26.[151,1437,1825,2032]" lastBlockId="27.[151,1437,150,969]" lastPageId="27" lastPageNumber="102" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44F8F854916C72F2" bold="true" box="[199,345,2005,2031]" pageId="26" pageNumber="101">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF0FFDA4590F854923672ED" box="[431,515,2005,2032]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="26" pageNumber="101">Fig. 59</figureCitation>
).
<emphasis id="A4DEEAF6FFF0FFDA4628F854926172F2" box="[535,596,2005,2031]" italics="true" pageId="26" pageNumber="101">Head</emphasis>
: Silvery white. Labial palpus silvery white, distal half of palpomere II suffused with fuscous on outer surface; palpomere III as long as II, fuscous dorsally. Antenna with scape silvery white; flagellum dark fuscous except basal half of dorsal surface mixed with silvery white, male cilia slightly longer than diameter.
</paragraph>
<paragraph id="961536E4FFF1FFDB44F8FE8390C574E9" blockId="27.[151,1437,150,969]" pageId="27" pageNumber="102">
<emphasis id="A4DEEAF6FFF1FFDB44F8FE83912D7401" box="[199,280,258,284]" italics="true" pageId="27" pageNumber="102">Thorax</emphasis>
: Dorsum and tegula silvery white. Wingspan 9.0 mm; forewing with basal 1/3 silvery white mixed with light grey, anterior edge of costa black, distal 2/3 grey; a roundish orange patch in distal 1/4 of wing, its anterior margin reaching costa, posterior margin bordered by fuscous scales above dorsum; a narrow silvery white line extending from costa to near tornus between patch and apical spot; a small, somewhat indistinct black spot on termen below apical spot; fringe scales grey. Hindwing grey, except black at apex and silvery white near apex; fringe scales grey. Legs silvery white; hind tibia dark fuscous apically; mid- and hind tarsi strongly mixed with fuscous on inner surface.
</paragraph>
<paragraph id="961536E4FFF1FFDB44F8FE7F91647625" blockId="27.[151,1437,150,969]" pageId="27" pageNumber="102">
<emphasis id="A4DEEAF6FFF1FFDB44F8FE7F91047705" box="[199,305,510,536]" italics="true" pageId="27" pageNumber="102">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFF1FFDB457FFE7F91A67705" box="[320,403,510,536]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="27" pageNumber="102">Fig. 22</figureCitation>
): Male coremata as long as sternum VIII; sternum VIII pentagonal, approximately 1/8 length of abdomen. Male genitalia (
<figureCitation id="0E912A61FFF1FFDB45E6FDA3921F7721" box="[473,554,546,572]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="27" pageNumber="102">Fig. 78</figureCitation>
) with uncus small, pentagonal, with a medial ridge. Gnathos hook short, thickened after base, pointed apically. Tegumen approximately 6 times as long as uncus; papilliform tubercles at middle, each bearing a long single hair (
<figureCitation id="0E912A61FFF1FFDB462EFDEB925D7799" box="[529,616,618,644]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="27" pageNumber="102">Fig. 39</figureCitation>
). Valva curved inward, gradually broadened from base to middle, then abruptly narrowed at costa after middle and forming a short projection, spatulate distally. Anellus lobe oval, approximately 1/4 length of valva; apical bristle twice as long as anellus lobe, strongly curved inward. Juxta projected at middle of inner margin. Vinculum simple, posterior margin rounded. Saccus sub-triangular, rounded apically, slightly exceeding tegumen pedunculus. Aedeagus with basal 2/5 globular, distal 3/5 slender and slightly sinuous, pointed apically.
</paragraph>
<paragraph id="961536E4FFF1FFDB44F8FCC391A47641" blockId="27.[151,1437,150,969]" box="[199,401,834,860]" pageId="27" pageNumber="102">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF1FFDB44F8FCE79213769C" box="[199,550,870,897]" pageId="27" pageNumber="102" type="distribution">
<paragraph id="961536E4FFF1FFDB44F8FCE79213769C" blockId="27.[151,1437,150,969]" box="[199,550,870,897]" pageId="27" pageNumber="102">
<emphasis id="A4DEEAF6FFF1FFDB44F8FCE7916C769D" bold="true" box="[199,345,870,896]" pageId="27" pageNumber="102">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFF1FFDB4558FCE7919F769D" box="[359,426,870,896]" name="China" pageId="27" pageNumber="102">China</collectingCountry>
(
<collectingRegion id="546EF806FFF1FFDB4586FCE7922F769C" box="[441,538,870,897]" country="China" name="Sichuan" pageId="27" pageNumber="102">Sichuan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF1FFDB44F8FC0B92FF76D5" pageId="27" pageNumber="102" type="etymology">
<paragraph id="961536E4FFF1FFDB44F8FC0B92FF76D5" blockId="27.[151,1437,150,969]" pageId="27" pageNumber="102">
<emphasis id="A4DEEAF6FFF1FFDB44F8FC0B917D76B9" bold="true" box="[199,328,906,932]" pageId="27" pageNumber="102">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFF1FFDB477DFC0A93BE76B9" box="[834,907,907,932]" italics="true" pageId="27" pageNumber="102">acutus</emphasis>
(pointed) and
<emphasis id="A4DEEAF6FFF1FFDB4010FC0B944B76B9" box="[1071,1150,906,932]" italics="true" pageId="27" pageNumber="102">medius</emphasis>
(middle), referring to the costa of valva with a pointed projection at middle.
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Lee
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Li
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(
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,
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,
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,
<figureCitation id="0E912A61FFF1FFDB4569FBBF91417145" box="[342,372,1086,1112]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="27" pageNumber="102">79</figureCitation>
,
<figureCitation id="0E912A61FFF1FFDB45BFFBBF91A87145" box="[384,413,1086,1112]" captionStart="FIGURES 91–94" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[240,1347,186,1895]" captionTargetId="figure-22@20.[203,1385,181,1920]" captionTargetPageId="20" captionText="FIGURES 91–94. Female genitalia of Tenupalpa spp. 91. T. venustalis, slide no. LGE18596; 92. T. acutata sp. nov., paratype, slide no. LGE18846; 93. T. crassimedia sp. nov., paratype, slide no. LGE18019; 94. T. breviaedeaga sp. nov., paratype, slide no. LGE18037 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819836" httpUri="https://zenodo.org/record/5819836/files/figure.png" pageId="27" pageNumber="102">94</figureCitation>
)
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<date id="E2141024FFF1FFDB4129FB6A95AD7018" box="[1302,1432,1259,1285]" pageId="27" pageNumber="102" value="2014-07-16">
<collectingDate id="F250E9CCFFF1FFDB4129FB6A95AD7018" box="[1302,1432,1259,1285]" pageId="27" pageNumber="102" value="2014-07-16">16.vii.2014</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF1FFDB44F7FA8E91607037" box="[200,341,1295,1322]" pageId="27" pageNumber="102">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFF1FFDB455EFA8E91E77034" box="[353,466,1295,1321]" pageId="27" pageNumber="102">Linjie Liu</collectorName>
and
<collectorName id="3B5F5332FFF1FFDB4636FA8E926A7037" box="[521,607,1295,1322]" pageId="27" pageNumber="102">Sha Hu</collectorName>
, genitalia slide no. LGE19012
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF1FFDB47FBFA91953A7053" ID-GBIF-Occurrence="3427696359" accessionNumber="LGE19017" collectingDate="2016-08-09" collectorName="Xia Bai &amp; Shuonan Qian &amp; Wanding Qi" country="China" elevation="787" latitude="18.73" location="Jianfengling" longLatPrecision="765" longitude="108.87" municipality="Tianchi" pageId="27" pageNumber="102" specimenCount="1" specimenCount-male="1" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF1FFDB47FBFA9193C57037" box="[964,1008,1296,1322]" pageId="27" pageNumber="102" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF1FFDB47C3FA8E94647034" box="[1020,1105,1295,1321]" pageId="27" pageNumber="102">Tianchi</collectingMunicipality>
(
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,
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),
<quantity id="51529B01FFF1FFDB4171FA9195A27037" box="[1358,1431,1296,1322]" metricMagnitude="2" metricUnit="m" metricValue="7.87" pageId="27" pageNumber="102" unit="m" value="787.0">
<elevation id="1D87D1D7FFF1FFDB4171FA9195A27037" box="[1358,1431,1296,1322]" metricMagnitude="2" metricUnit="m" metricValue="7.87" pageId="27" pageNumber="102" unit="m" value="787.0">787 m</elevation>
</quantity>
,
<location id="9375603FFFF1FFDB44A8FAB291117050" LSID="urn:lsid:plazi:treatment:1E0387F2FFF1FFDC44A8FB9B915B7720:9375603FFFF1FFDB44A8FAB291117050" box="[151,292,1331,1357]" country="China" latitude="18.73" longLatPrecision="765" longitude="108.87" municipality="Tianchi" name="Jianfengling" pageId="27" pageNumber="102" stateProvince="Hainan Province">Jianfengling</location>
,
<date id="E2141024FFF1FFDB450EFAB291997050" box="[305,428,1331,1357]" pageId="27" pageNumber="102" value="2016-08-09">
<collectingDate id="F250E9CCFFF1FFDB450EFAB291997050" box="[305,428,1331,1357]" pageId="27" pageNumber="102" value="2016-08-09">9.viii.2016</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF1FFDB45D5FAB292767050" box="[490,579,1331,1357]" pageId="27" pageNumber="102">Xia Bai</collectorName>
,
<collectorName id="3B5F5332FFF1FFDB466FFAB292C57050" box="[592,752,1331,1357]" pageId="27" pageNumber="102">Shuonan Qian</collectorName>
and
<collectorName id="3B5F5332FFF1FFDB4717FAB293877050" box="[808,946,1331,1357]" pageId="27" pageNumber="102">Wanding Qi</collectorName>
, genitalia slide no. LGE19017
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF1FFDB4124FAB591B37088" ID-GBIF-Occurrence="3427696354" accessionNumber="LGE18037" collectingDate="2017-08-09" collectorName="Xia Bai &amp; Ping Liu &amp; Shuai Yu" country="China" elevation="954" latitude="18.74" location="Jianfengling" longLatPrecision="765" longitude="108.84" municipality="Mingfenggu" pageId="27" pageNumber="102" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF1FFDB4124FAB595687053" box="[1307,1373,1332,1358]" pageId="27" pageNumber="102" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="80ACFD6DFFF1FFDB4155FAB595A07053" box="[1386,1429,1332,1358]" pageId="27" pageNumber="102" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF1FFDB44A8FAD69114706C" box="[151,289,1367,1393]" pageId="27" pageNumber="102">Mingfenggu</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFF1FFDB450FFAD991A7706F" box="[304,402,1368,1394]" degrees="18.74" direction="north" orientation="latitude" pageId="27" pageNumber="102" precision="555" value="18.74">18.74°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF1FFDB45A3FAD9923C706F" box="[412,521,1368,1394]" degrees="108.84" direction="east" orientation="longitude" pageId="27" pageNumber="102" precision="555" value="108.84">108.84°E</geoCoordinate>
),
<quantity id="51529B01FFF1FFDB4624FAD69256706F" box="[539,611,1367,1394]" metricMagnitude="2" metricUnit="m" metricValue="9.54" pageId="27" pageNumber="102" unit="m" value="954.0">
<elevation id="1D87D1D7FFF1FFDB4624FAD69256706F" box="[539,611,1367,1394]" metricMagnitude="2" metricUnit="m" metricValue="9.54" pageId="27" pageNumber="102" unit="m" value="954.0">954 m</elevation>
</quantity>
,
<location id="9375603FFFF1FFDB4651FAD692CE706C" LSID="urn:lsid:plazi:treatment:1E0387F2FFF1FFDC44A8FB9B915B7720:9375603FFFF1FFDB4651FAD692CE706C" box="[622,763,1367,1393]" country="China" latitude="18.74" longLatPrecision="765" longitude="108.84" municipality="Mingfenggu" name="Jianfengling" pageId="27" pageNumber="102" stateProvince="Hainan Province">Jianfengling</location>
, 8,
<date id="E2141024FFF1FFDB471FFAD693A9706C" box="[800,924,1367,1393]" pageId="27" pageNumber="102" value="2017-08-09">
<collectingDate id="F250E9CCFFF1FFDB471FFAD693A9706C" box="[800,924,1367,1393]" pageId="27" pageNumber="102" value="2017-08-09">9.viii.2017</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF1FFDB47EAFAD69419706C" box="[981,1068,1367,1393]" pageId="27" pageNumber="102">Xia Bai</collectorName>
,
<collectorName id="3B5F5332FFF1FFDB4008FAD694A2706C" box="[1079,1175,1367,1393]" pageId="27" pageNumber="102">Ping Liu</collectorName>
and
<collectorName id="3B5F5332FFF1FFDB40F4FAD69507706F" box="[1227,1330,1367,1394]" pageId="27" pageNumber="102">Shuai Yu</collectorName>
, genitalia slide nos. LGE18037
</materialsCitation>
f, LGE
<quantity id="51529B01FFF1FFDB45EAFAFD922E708B" box="[469,539,1404,1430]" metricMagnitude="4" metricUnit="m" metricValue="1.9014" pageId="27" pageNumber="102" unit="m" value="19014.0">19014</quantity>
m.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF1FFDB44F8FA1E91B47338" pageId="27" pageNumber="102" type="diagnosis">
<paragraph id="961536E4FFF1FFDB44F8FA1E91B47338" blockId="27.[151,1437,1187,2042]" pageId="27" pageNumber="102">
<emphasis id="A4DEEAF6FFF1FFDB44F8FA1E917770A4" bold="true" box="[199,322,1439,1465]" pageId="27" pageNumber="102">Diagnosis.</emphasis>
Compared to its allies, the species has a somewhat larger distal patch on the forewing. The male genitalia are similar to those of
<taxonomicName id="51AA4D67FFF1FFDB45C8FA45925370C0" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[503,614,1476,1501]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="27" pageNumber="102" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFF1FFDB45C8FA45925370C0" box="[503,614,1476,1501]" italics="true" pageId="27" pageNumber="102">T. acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF1FFDB4653FA4592F070C3" bold="true" box="[620,709,1476,1502]" pageId="27" pageNumber="102">
<taxonomicNameLabel id="BFED578DFFF1FFDB4653FA4592F070C3" box="[620,709,1476,1502]" pageId="27" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="51AA4D67FFF1FFDB46C5FA4593A970C0" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[762,924,1475,1501]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="27" pageNumber="102" phylum="Tracheophyta" rank="species" species="crassimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFF1FFDB46C5FA4593A970C0" box="[762,924,1475,1501]" italics="true" pageId="27" pageNumber="102">T. crassimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF1FFDB479CFA4593C970C3" bold="true" box="[931,1020,1476,1502]" pageId="27" pageNumber="102">
<taxonomicNameLabel id="BFED578DFFF1FFDB479CFA4593C970C3" box="[931,1020,1476,1502]" pageId="27" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but the valva is broader apically and the aedeagus is much shorter. The female genitalia are characterized by a semicircular signum situated at the middle of the corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF1FFDC44F8F9AE93C674CC" lastPageId="28" lastPageNumber="103" pageId="27" pageNumber="102" type="description">
<paragraph id="961536E4FFF1FFDB44F8F9AE9559738C" blockId="27.[151,1437,1187,2042]" pageId="27" pageNumber="102">
<emphasis id="A4DEEAF6FFF1FFDB44F8F9AE916C7354" bold="true" box="[199,345,1583,1609]" pageId="27" pageNumber="102">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF1FFDB458CF9AE923F7357" box="[435,522,1583,1610]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="27" pageNumber="102">Fig. 60</figureCitation>
).
<emphasis id="A4DEEAF6FFF1FFDB461FF9AE92687354" box="[544,605,1583,1609]" italics="true" pageId="27" pageNumber="102">Head</emphasis>
: Silvery white. Labial palpus silvery white, palpomere II slightly mixed with grey on outer surface; palpomere III as long as II, fuscous dorsally.Antenna with scape silvery white; flagellum dark fuscous except basal half of dorsal surface mixed with silvery white, male cilia twice as long as diameter.
</paragraph>
<paragraph id="961536E4FFF1FFDB44F8F91A936F7293" blockId="27.[151,1437,1187,2042]" pageId="27" pageNumber="102">
<emphasis id="A4DEEAF6FFF1FFDB44F8F91A912D73A8" box="[199,280,1691,1717]" italics="true" pageId="27" pageNumber="102">Thorax</emphasis>
: Dorsum and tegula silvery white to grey. Wingspan 6.0–7.5 mm; forewing glossy grey, anterior edge of costa black in basal half; an elliptical orange patch in distal 2/5 of wing between costa and posterior 1/4; a narrow white line extending from costa to tornus between former patch and apical spot; sometimes a small indistinct black spot on termen below apical spot; dorsum suffused with fuscous in distal 1/3, sometimes slightly tinged with orange near tornus; fringe scales grey. Hindwing grey except black at apex and white near apex; fringe scales grey. Legs silvery white; mid-tibia slightly tinged with grey near apex on outer surface; hind tibia grey to dark grey at middle and at apex; mid- and hind tarsi strongly mixed with dark grey.
</paragraph>
<paragraph id="961536E4FFF1FFDC44F8F81693C674CC" blockId="27.[151,1437,1187,2042]" lastBlockId="28.[151,1437,151,573]" lastPageId="28" lastPageNumber="103" pageId="27" pageNumber="102">
<emphasis id="A4DEEAF6FFF1FFDB44F8F816910472AC" box="[199,305,1943,1969]" italics="true" pageId="27" pageNumber="102">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFF1FFDB457EF81691A272AF" box="[321,407,1943,1970]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="27" pageNumber="102">Fig. 23</figureCitation>
): Male coremata slightly shorter than sternum VIII; sternum VIII sub-pentagonal, approximately 1/7 length of abdomen. Male genitalia (
<figureCitation id="0E912A61FFF1FFDB469BF83A92C072CB" box="[676,757,1979,2006]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="27" pageNumber="102">Fig. 79</figureCitation>
) with uncus small, pentagonal, with a medial ridge. Gnathos hook moderately long, acute apically. Tegumen approximately 6 times as long as uncus; papilliform tubercles at middle, each bearing a long single hair (
<figureCitation id="0E912A61FFF6FFDC4661FF16928675AC" box="[606,691,151,177]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="28" pageNumber="103">Fig. 40</figureCitation>
). Valva slender and nearly straight in basal half, spatulate distally. Anellus lobe oval, approximately 1/4 or 1/5 length of valva; apical bristle slightly longer than anellus lobe, curved inward. Juxta simple, setose. Vinculum simple, posterior margin broadly convex. Saccus sub-triangular, sometimes constricted at middle, slightly exceeding tegumen pedunculus. Aedeagus with basal half globular, distally sinuous or straight, truncate apically. Female genitalia (
<figureCitation id="0E912A61FFF6FFDC46E4FEA69318745C" box="[731,813,295,321]" captionStart="FIGURES 91–94" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[240,1347,186,1895]" captionTargetId="figure-22@20.[203,1385,181,1920]" captionTargetPageId="20" captionText="FIGURES 91–94. Female genitalia of Tenupalpa spp. 91. T. venustalis, slide no. LGE18596; 92. T. acutata sp. nov., paratype, slide no. LGE18846; 93. T. crassimedia sp. nov., paratype, slide no. LGE18019; 94. T. breviaedeaga sp. nov., paratype, slide no. LGE18037 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819836" httpUri="https://zenodo.org/record/5819836/files/figure.png" pageId="28" pageNumber="103">Fig. 94</figureCitation>
) with papillae anales simple. Apophyses anteriores approximately half the length of apophyses posteriores. Tergum VIII divided into two sclerites; sternum VIII broadly convex anteriorly, strongly sclerotized along anterior margin. Ostium bursae round, opening at middle of sternum VIII. Ductus bursae slightly constricted after base, then gradually broadening toward corpus bursae. Corpus bursae elongate ovate; signum semicircular, situated near entrance of ductus bursae.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF6FFDC44F8FE5B922974EB" box="[199,540,474,502]" pageId="28" pageNumber="103" type="distribution">
<paragraph id="961536E4FFF6FFDC44F8FE5B922974EB" blockId="28.[151,1437,151,573]" box="[199,540,474,502]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44F8FE5B916C74E9" bold="true" box="[199,345,474,500]" pageId="28" pageNumber="103">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFF6FFDC4558FE5A919F74E8" box="[359,426,475,501]" name="China" pageId="28" pageNumber="103">China</collectingCountry>
(
<collectingRegion id="546EF806FFF6FFDC4586FE5A922574EB" box="[441,528,475,502]" country="China" name="Hainan" pageId="28" pageNumber="103">Hainan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF6FFDC44F8FE7E915B7720" pageId="28" pageNumber="103" type="etymology">
<paragraph id="961536E4FFF6FFDC44F8FE7E915B7720" blockId="28.[151,1437,151,573]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44F8FE7E917D7704" bold="true" box="[199,328,511,537]" pageId="28" pageNumber="103">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFF6FFDC476EFE7E93BC7704" box="[849,905,511,537]" italics="true" pageId="28" pageNumber="103">brevi</emphasis>
- (short) and
<emphasis id="A4DEEAF6FFF6FFDC401FFE7E94BF7704" box="[1056,1162,511,537]" italics="true" pageId="28" pageNumber="103">aedeagus</emphasis>
(aedeagus), referring to the short aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>

825
data/1E/03/87/1E0387F2FFF2FFD944A8FC2F93277213.xml Normal file
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@ -0,0 +1,825 @@
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<mods:title id="BC2A864A71F5C0EDCFC91CE9019A8D8E">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<mods:namePart id="BADC049F80EC54FB32DDF081D1221CE0">Lee, Ga-Eun</mods:namePart>
<mods:nameIdentifier id="F333DDD16F21992FB8DA0629A3FEA8BA" type="ORCID">0000-0002-5774-5850</mods:nameIdentifier>
<mods:affiliation id="C940C312BB96D8FEDE840A25574F6257">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China &amp; The Research Institute for Sciences, Jeju National University, Jeju 63243, Korea &amp; gaeunlee 486 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5774 - 5850</mods:affiliation>
<mods:nameIdentifier id="32989A89C2BDF0BF52DF58382156F251" type="email">gaeunlee486@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B4DD1D2C50B45ECA2079C68B69B49280">Li, Houhun</mods:namePart>
<mods:affiliation id="19E5452AE1B84EA235EC364E612AC74B">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China</mods:affiliation>
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<mods:title id="BF494E30624C4E1B4BB5A1343AEA1C14">Zootaxa</mods:title>
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<mods:part id="B1A7A5BEA38AF58D872683CD50B8F5F6">
<mods:date id="A67C7B41345E7E82262EAADF3CCBB8A5">2022</mods:date>
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<mods:number id="7709C1C7F17A9321FE6A297B82765386">2022-01-05</mods:number>
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<mods:number id="1A6D6D04FFEEC9396E67CCC4134A02E2">5087</mods:number>
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<taxonomicName id="51AA4D67FFF2FFD844A8FC2F91DB76D5" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,494,942,969]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="99" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFF2FFD844A8FC2F914D76D4" bold="true" box="[151,376,942,969]" italics="true" pageId="24" pageNumber="99">Tenupalpa acutata</emphasis>
Lee
<emphasis id="A4DEEAF6FFF2FFD8458BFC2E91FC76D4" bold="true" box="[436,457,943,969]" italics="true" pageId="24" pageNumber="99">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFF2FFD845C4FC2E926C76D4" box="[507,601,943,969]" pageId="24" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="0E912A61FFF2FFD844A0FC5290C276F0" box="[159,247,979,1005]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="24" pageNumber="99">Figs 20</figureCitation>
,
<figureCitation id="0E912A61FFF2FFD8453DFC52911576F0" box="[258,288,979,1005]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="24" pageNumber="99">37</figureCitation>
,
<figureCitation id="0E912A61FFF2FFD84513FC53917F76F0" box="[300,330,978,1005]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="24" pageNumber="99">57</figureCitation>
,
<figureCitation id="0E912A61FFF2FFD84569FC52914076F0" box="[342,373,979,1005]" captionStart="FIGURES 72–77" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 72–77. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="24" pageNumber="99">76</figureCitation>
,
<figureCitation id="0E912A61FFF2FFD845BFFC5291A876F0" box="[384,413,979,1005]" captionStart="FIGURES 91–94" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[240,1347,186,1895]" captionTargetId="figure-22@20.[203,1385,181,1920]" captionTargetPageId="20" captionText="FIGURES 91–94. Female genitalia of Tenupalpa spp. 91. T. venustalis, slide no. LGE18596; 92. T. acutata sp. nov., paratype, slide no. LGE18846; 93. T. crassimedia sp. nov., paratype, slide no. LGE18019; 94. T. breviaedeaga sp. nov., paratype, slide no. LGE18037 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819836" httpUri="https://zenodo.org/record/5819836/files/figure.png" pageId="24" pageNumber="99">92</figureCitation>
)
</paragraph>
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urn:lsid:zoobank.org:act:
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<emphasis id="A4DEEAF6FFF2FFD844A8FBB9917C714F" bold="true" box="[151,329,1080,1106]" pageId="24" pageNumber="99">Type material.</emphasis>
<materialsCitation id="26C23CB9FFF2FFD8456BFBB994E7716B" ID-GBIF-Occurrence="3427696447" accessionNumber="LGE18030" collectingDate="2017-05-18" collectorName="Shuonan Qian &amp; Ga-Eun Lee" country="China" county="Kaihua County" elevation="310" latitude="29.23" location="Gutianshan" longLatPrecision="4880" longitude="118.1" municipality="Quzhou City" pageId="24" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="holotype">
<typeStatus id="49118846FFF2FFD8456BFBB99189714F" box="[340,444,1080,1106]" pageId="24" pageNumber="99" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFF2FFD845F7FBB891EA714E" box="[456,479,1081,1107]" pageId="24" pageNumber="99" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFF2FFD845D0FBB9927E714F" box="[495,587,1080,1106]" name="China" pageId="24" pageNumber="99">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFF2FFD84665FBB9931A714F" box="[602,815,1080,1106]" country="China" name="Zhejiang" pageId="24" pageNumber="99">Zhejiang Province</collectingRegion>
,
<collectingMunicipality id="7671AC9EFFF2FFD84700FBB993E1714F" box="[831,980,1080,1106]" pageId="24" pageNumber="99">Quzhou City</collectingMunicipality>
,
<collectingCounty id="7F744E68FFF2FFD847DBFBB994A1714F" box="[996,1172,1080,1106]" pageId="24" pageNumber="99">Kaihua County</collectingCounty>
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(
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<date id="E2141024FFF2FFD8455EFBDD91E7716B" box="[353,466,1116,1142]" pageId="24" pageNumber="99" value="2017-05-18">
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, leg.
<collectorName id="3B5F5332FFF2FFD8462FFBDD9285716B" box="[528,688,1116,1142]" pageId="24" pageNumber="99">Shuonan Qian</collectorName>
and
<collectorName id="3B5F5332FFF2FFD846D6FBDD9340716A" box="[745,885,1116,1143]" pageId="24" pageNumber="99">Ga-Eun Lee</collectorName>
, genitalia slide no. LGE18030
</materialsCitation>
.
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<typeStatus id="49118846FFF2FFD840E2FBDD9562716B" box="[1245,1367,1116,1142]" pageId="24" pageNumber="99" type="paratype">Paratypes</typeStatus>
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<collectingRegion id="546EF806FFF2FFD8415BFBDD917E7187" country="China" name="Zhejiang" pageId="24" pageNumber="99">Zhejiang Province</collectingRegion>
:
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<specimenCount id="80ACFD6DFFF2FFD84566FB0191A97186" box="[345,412,1152,1179]" pageId="24" pageNumber="99" type="male">2 ♂♂</specimenCount>
, same data as holotype except
<date id="E2141024FFF2FFD8473DFB0193477187" box="[770,882,1152,1178]" pageId="24" pageNumber="99" value="2017-05-20">
<collectingDate id="F250E9CCFFF2FFD8473DFB0193477187" box="[770,882,1152,1178]" pageId="24" pageNumber="99" value="2017-05-20">20.v.2017</collectingDate>
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, genitalia slide no. LGE17045
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;
<materialsCitation id="26C23CB9FFF2FFD840DEFB0191B771FE" ID-GBIF-Occurrence="3427696302" accessionNumber="LGE17259" collectingDate="2017-08-25" collectorName="Shuonan Qian &amp; Yanyan Jia &amp; Juan Li" country="China" elevation="412" latitude="29.24" location="Quzhou City" longLatPrecision="736" longitude="118.11" municipality="Gutianshan" pageId="24" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD840DEFB01953A7186" box="[1249,1295,1152,1179]" pageId="24" pageNumber="99" type="male">1 ♂</specimenCount>
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<collectingMunicipality id="7671AC9EFFF2FFD84122FB0195A87187" box="[1309,1437,1152,1178]" pageId="24" pageNumber="99">Gutianshan</collectingMunicipality>
(
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<location id="9375603FFFF2FFD845CFFB2592B071A3" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD845CFFB2592B071A3" box="[496,645,1188,1214]" country="China" latitude="29.24" longLatPrecision="736" longitude="118.11" municipality="Gutianshan" name="Quzhou City" pageId="24" pageNumber="99" stateProvince="Zhejiang Province">Quzhou City</location>
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<date id="E2141024FFF2FFD846ABFB25932B71A2" box="[660,798,1188,1215]" pageId="24" pageNumber="99" value="2017-08-25">
<collectingDate id="F250E9CCFFF2FFD846ABFB25932B71A2" box="[660,798,1188,1215]" pageId="24" pageNumber="99" value="2017-08-25">25.viii.2017</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD8475DFB25943D71A3" box="[866,1032,1188,1214]" pageId="24" pageNumber="99">Shuonan Qian</collectorName>
,
<collectorName id="3B5F5332FFF2FFD84029FB2494A171A3" box="[1046,1172,1188,1215]" pageId="24" pageNumber="99">Yanyan Jia</collectorName>
and
<collectorName id="3B5F5332FFF2FFD840ECFB24951971A3" box="[1235,1324,1188,1215]" pageId="24" pageNumber="99">Juan Li</collectorName>
, genitalia slide no. LGE17259
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD845AEFB4992FB701A" ID-GBIF-Occurrence="3427696374" accessionNumber="LGE19005" collectingDate="2018-08-11" collectorName="Shuai Yu &amp; Mengran Xing &amp; Chen Liu" country="China" elevation="465" latitude="28.21" location="Laofoyan Village" longLatPrecision="740" longitude="118.41" pageId="24" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD845AEFB4991F571FE" box="[401,448,1224,1251]" pageId="24" pageNumber="99" type="male">1 ♂</specimenCount>
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(
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,
<geoCoordinate id="F39E5023FFF2FFD84728FB4993B671FF" box="[791,899,1224,1250]" degrees="118.41" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="118.41">118.41°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD847A6FB4993D071FE" box="[921,997,1224,1251]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="24" pageNumber="99" unit="m" value="465.0">
<elevation id="1D87D1D7FFF2FFD847A6FB4993D071FE" box="[921,997,1224,1251]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="24" pageNumber="99" unit="m" value="465.0">465 m</elevation>
</quantity>
,
<date id="E2141024FFF2FFD847CBFB49944971FF" box="[1012,1148,1224,1250]" pageId="24" pageNumber="99" value="2018-08-11">
<collectingDate id="F250E9CCFFF2FFD847CBFB49944971FF" box="[1012,1148,1224,1250]" pageId="24" pageNumber="99" value="2018-08-11">11.viii.2018</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84081FB49951C71FE" box="[1214,1321,1224,1251]" pageId="24" pageNumber="99">Shuai Yu</collectorName>
,
<collectorName id="3B5F5332FFF2FFD84108FB4890FA701B" pageId="24" pageNumber="99">Mengran Xing</collectorName>
and
<collectorName id="3B5F5332FFF2FFD84538FB6D9141701B" box="[263,372,1260,1286]" pageId="24" pageNumber="99">Chen Liu</collectorName>
, genitalia slide no. LGE19005
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD846E4FB6D949C7037" ID-GBIF-Occurrence="3427696441" collectingDate="2018-08" collectorName="Shuai Yu &amp; Mengran Xing &amp; Chen Liu" country="China" county="Kaihua County" elevation="416" latitude="29.14" location="Gutianshan" longLatPrecision="737" longitude="118.07" pageId="24" pageNumber="99" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Zhejiang Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD846E4FB6D9329701A" box="[731,796,1260,1287]" pageId="24" pageNumber="99" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="80ACFD6DFFF2FFD84716FB6D935F701A" box="[809,874,1260,1287]" pageId="24" pageNumber="99" type="female">3 ♀♀</specimenCount>
,
<location id="9375603FFFF2FFD84748FB6D93C2701B" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD84748FB6D93C2701B" box="[887,1015,1260,1286]" country="China" county="Kaihua County" latitude="29.14" longLatPrecision="737" longitude="118.07" name="Gutianshan" pageId="24" pageNumber="99" stateProvince="Zhejiang Province">Gutianshan</location>
(
<geoCoordinate id="F39E5023FFF2FFD84039FB6D945C701B" box="[1030,1129,1260,1286]" degrees="29.14" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="29.14">29.14°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD8404AFB6D94D4701B" box="[1141,1249,1260,1286]" degrees="118.07" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="118.07">118.07°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD840CAFB6D950A701A" box="[1269,1343,1260,1287]" metricMagnitude="2" metricUnit="m" metricValue="4.16" pageId="24" pageNumber="99" unit="m" value="416.0">
<elevation id="1D87D1D7FFF2FFD840CAFB6D950A701A" box="[1269,1343,1260,1287]" metricMagnitude="2" metricUnit="m" metricValue="4.16" pageId="24" pageNumber="99" unit="m" value="416.0">416 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFF2FFD84174FB6D90DE7037" pageId="24" pageNumber="99">Kaihua County</collectingCounty>
, 13,15.
<date id="E2141024FFF2FFD84505FA9191977036" box="[314,418,1296,1323]" pageId="24" pageNumber="99" value="2018-08">
<collectingDate id="F250E9CCFFF2FFD84505FA9191977036" box="[314,418,1296,1323]" pageId="24" pageNumber="99" value="2018-08">viii.2018</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD845E3FA9192767036" box="[476,579,1296,1323]" pageId="24" pageNumber="99">Shuai Yu</collectorName>
,
<collectorName id="3B5F5332FFF2FFD84671FA9092C47037" box="[590,753,1296,1323]" pageId="24" pageNumber="99">Mengran Xing</collectorName>
and
<collectorName id="3B5F5332FFF2FFD84719FA9193A47037" box="[806,913,1296,1322]" pageId="24" pageNumber="99">Chen Liu</collectorName>
, genitalis slide nos. LGE
</materialsCitation>
<materialsCitation id="26C23CB9FFF2FFD84096FA9195B87037" ID-GBIF-Occurrence="3427696426" accessionNumber="LGE18846" box="[1193,1421,1296,1322]" country="China" elevation="18710" pageId="24" pageNumber="99" specimenCount="1" stateProvince="Zhejiang Province" typeStatus="paratype">
<quantity id="51529B01FFF2FFD84096FA9195307037" box="[1193,1285,1296,1322]" metricMagnitude="4" metricUnit="m" metricValue="1.871" pageId="24" pageNumber="99" unit="m" value="18710.0">
<elevation id="1D87D1D7FFF2FFD84096FA9195307037" box="[1193,1285,1296,1322]" metricMagnitude="4" metricUnit="m" metricValue="1.871" pageId="24" pageNumber="99" unit="m" value="18710.0">18710m</elevation>
</quantity>
, LGE18846
</materialsCitation>
f, LGE
<quantity id="51529B01FFF2FFD844EFFAB591237053" box="[208,278,1332,1358]" metricMagnitude="4" metricUnit="m" metricValue="1.9006" pageId="24" pageNumber="99" unit="m" value="19006.0">19006</quantity>
m, LGE
<quantity id="51529B01FFF2FFD8454FFAB591807053" box="[368,437,1332,1358]" metricMagnitude="4" metricUnit="m" metricValue="1.9007999999999998" pageId="24" pageNumber="99" unit="m" value="19008.0">19008</quantity>
m, LGE
<quantity id="51529B01FFF2FFD8462FFAB592637053" box="[528,598,1332,1358]" metricMagnitude="4" metricUnit="m" metricValue="1.9009" pageId="24" pageNumber="99" unit="m" value="19009.0">19009</quantity>
m.
<materialsCitation id="26C23CB9FFF2FFD84649FAB5948B706E" ID-GBIF-Occurrence="3427696369" accessionNumber="LGE18005" collectingDate="2008-07-27" collectorName="Weichun Li &amp; Yongling Sun &amp; Haiyan Bai" country="China" elevation="740" latitude="27.45" location="Mt. Wuyi" longLatPrecision="742" longitude="117.41" municipality="Sangang" pageId="24" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Fujian" typeStatus="paratype">
<emphasis id="A4DEEAF6FFF2FFD84649FAB5930A7053" bold="true" box="[630,831,1332,1358]" pageId="24" pageNumber="99">
<collectingRegion id="546EF806FFF2FFD84649FAB5930F7053" box="[630,826,1332,1358]" country="China" name="Fujian" pageId="24" pageNumber="99">Fujian Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFF2FFD8477AFAB593457052" box="[837,880,1332,1359]" pageId="24" pageNumber="99" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD84743FAB593E97053" box="[892,988,1332,1358]" pageId="24" pageNumber="99">Sangang</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFF2FFD847D4FAB594787052" box="[1003,1101,1332,1359]" degrees="27.45" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="27.45">27.45°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD84067FAB594F17053" box="[1112,1220,1332,1358]" degrees="117.41" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="117.41">117.41°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD840E8FAB5952A7052" box="[1239,1311,1332,1359]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="24" pageNumber="99" unit="m" value="740.0">
<elevation id="1D87D1D7FFF2FFD840E8FAB5952A7052" box="[1239,1311,1332,1359]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="24" pageNumber="99" unit="m" value="740.0">740 m</elevation>
</quantity>
,
<location id="9375603FFFF2FFD84114FAB495AD7053" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD84114FAB495AD7053" box="[1323,1432,1332,1359]" country="China" latitude="27.45" longLatPrecision="742" longitude="117.41" municipality="Sangang" name="Mt. Wuyi" pageId="24" pageNumber="99" stateProvince="Fujian">Mt. Wuyi</location>
,
<date id="E2141024FFF2FFD844A8FAD9912F706F" box="[151,282,1368,1394]" pageId="24" pageNumber="99" value="2008-07-27">
<collectingDate id="F250E9CCFFF2FFD844A8FAD9912F706F" box="[151,282,1368,1394]" pageId="24" pageNumber="99" value="2008-07-27">27.vii.2008</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84562FAD991D0706F" box="[349,485,1368,1394]" pageId="24" pageNumber="99">Weichun Li</collectorName>
,
<collectorName id="3B5F5332FFF2FFD845CBFAD992A7706F" box="[500,658,1368,1394]" pageId="24" pageNumber="99">Yongling Sun</collectorName>
and
<collectorName id="3B5F5332FFF2FFD846EFFAD99360706F" box="[720,853,1368,1394]" pageId="24" pageNumber="99">Haiyan Bai</collectorName>
, genitalia slide no. LGE18005
</materialsCitation>
.
<emphasis id="A4DEEAF6FFF2FFD840F2FAD991A0708B" bold="true" pageId="24" pageNumber="99">
<materialsCitation id="26C23CB9FFF2FFD840F2FAD991A5708B" ID-GBIF-Occurrence="3427696332" country="China" location="Autonomous Region" pageId="24" pageNumber="99" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
<collectingRegion id="546EF806FFF2FFD840F2FAD995A9706F" box="[1229,1436,1368,1394]" country="China" name="Guangxi" pageId="24" pageNumber="99">Guangxi Zhuang</collectingRegion>
<location id="9375603FFFF2FFD844A8FAFD91A5708B" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD844A8FAFD91A5708B" box="[151,400,1404,1430]" country="China" name="Autonomous Region" pageId="24" pageNumber="99" stateProvince="Guangxi">Autonomous Region</location>
</materialsCitation>
:
</emphasis>
<materialsCitation id="26C23CB9FFF2FFD845A2FAFD92A970A7" ID-GBIF-Occurrence="3427696433" accessionNumber="LGE18702" collectingDate="2008-05-06" collectorName="Hui Zhen &amp; Li Zhang" country="China" county="Yongfu County" elevation="160" latitude="24.98" location="Qinmu Village" longLatPrecision="749" longitude="109.98" pageId="24" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi Zhuang" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD845A2FAFD91FE708A" box="[413,459,1404,1431]" pageId="24" pageNumber="99" type="male">1 ♂</specimenCount>
,
<location id="9375603FFFF2FFD845E7FAFD924B708B" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD845E7FAFD924B708B" box="[472,638,1404,1430]" country="China" county="Yongfu County" latitude="24.98" longLatPrecision="749" longitude="109.98" name="Qinmu Village" pageId="24" pageNumber="99" stateProvince="Guangxi Zhuang">Qinmu Village</location>
(
<geoCoordinate id="F39E5023FFF2FFD846B1FAFD92C4708B" box="[654,753,1404,1430]" degrees="24.98" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="24.98">24.98°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD846C2FAFD935F708B" box="[765,874,1404,1430]" degrees="109.98" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="109.98">109.98°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD84740FAFD93FC708A" box="[895,969,1404,1431]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="24" pageNumber="99" unit="m" value="160.0">
<elevation id="1D87D1D7FFF2FFD84740FAFD93FC708A" box="[895,969,1404,1431]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="24" pageNumber="99" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFF2FFD847EAFAFD94B1708B" box="[981,1156,1404,1430]" pageId="24" pageNumber="99">Yongfu County</collectingCounty>
,
<date id="E2141024FFF2FFD840AFFAFD94C7708B" box="[1168,1266,1404,1430]" pageId="24" pageNumber="99" value="2008-05-06">
<collectingDate id="F250E9CCFFF2FFD840AFFAFD94C7708B" box="[1168,1266,1404,1430]" pageId="24" pageNumber="99" value="2008-05-06">6.v.2008</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD8410FFAFD95A8708B" box="[1328,1437,1404,1430]" pageId="24" pageNumber="99">Hui Zhen</collectorName>
and
<collectorName id="3B5F5332FFF2FFD844F5FA21910270A7" box="[202,311,1440,1466]" pageId="24" pageNumber="99">Li Zhang</collectorName>
, genitalia slide no. LGE18702
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD84695FA21949470C3" ID-GBIF-Occurrence="3427696406" collectingDate="2015-07-23" collectingDateMax="2015-07-26" collectingDateMin="2015-07-23" collectorName="Meiqing Yang &amp; Ga-Eun Lee" country="China" county="Huanjiang County" elevation="1180" latitude="25.29" location="Hechi" longLatPrecision="748" longitude="109.31" municipality="Yangmeiao" pageId="24" pageNumber="99" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Guangxi Zhuang" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD84695FA2192DB70A6" box="[682,750,1440,1467]" pageId="24" pageNumber="99" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="80ACFD6DFFF2FFD846C1FA21931E70A6" box="[766,811,1440,1467]" pageId="24" pageNumber="99" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD84704FA21938C70A7" box="[827,953,1440,1466]" pageId="24" pageNumber="99">Yangmeiao</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFF2FFD847F4FA21941B70A6" box="[971,1070,1440,1467]" degrees="25.29" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="25.29">25.29°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD84003FA21949C70A7" box="[1084,1193,1440,1466]" degrees="109.31" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="109.31">109.31°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD840FFFA21952C70A6" box="[1216,1305,1440,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="24" pageNumber="99" unit="m" value="1180.0">
<elevation id="1D87D1D7FFF2FFD840FFFA21952C70A6" box="[1216,1305,1440,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="24" pageNumber="99" unit="m" value="1180.0">1180 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFF2FFD84117FA2190DE70C3" pageId="24" pageNumber="99">Huanjiang County</collectingCounty>
,
<location id="9375603FFFF2FFD844C9FA45910970C3" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD844C9FA45910970C3" box="[246,316,1476,1502]" country="China" county="Huanjiang County" latitude="25.29" longLatPrecision="748" longitude="109.31" municipality="Yangmeiao" name="Hechi" pageId="24" pageNumber="99" stateProvince="Guangxi Zhuang">Hechi</location>
,
<date id="E2141024FFF2FFD84578FA4591C070C2" box="[327,501,1476,1503]" pageId="24" pageNumber="99" value="2015-07-23" valueMax="2015-07-26" valueMin="2015-07-23">
<collectingDate id="F250E9CCFFF2FFD84578FA4591C070C2" box="[327,501,1476,1503]" pageId="24" pageNumber="99" value="2015-07-23" valueMax="2015-07-26" valueMin="2015-07-23">23−26.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD8460FFA4592FA70C2" box="[560,719,1476,1503]" pageId="24" pageNumber="99">Meiqing Yang</collectorName>
and
<collectorName id="3B5F5332FFF2FFD8473AFA4593A570C2" box="[773,912,1476,1503]" pageId="24" pageNumber="99">Ga-Eun Lee</collectorName>
, genitalia slide no. LGE
</materialsCitation>
<quantity id="51529B01FFF2FFD8409DFA4594CA70C3" box="[1186,1279,1476,1502]" metricMagnitude="4" metricUnit="m" metricValue="1.7037" pageId="24" pageNumber="99" unit="m" value="17037.0">17037m</quantity>
;
<materialsCitation id="26C23CB9FFF2FFD84136FA4591E8733B" ID-GBIF-Occurrence="3427696381" collectingDate="2015-07-27" collectorName="Meiqing Yang &amp; Ga-Eun Lee" country="China" county="Rongshui County" elevation="1240" location="Liuzhou" municipality="Yangmeiao Protection Station" pageId="24" pageNumber="99" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Guangxi Zhuang" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD84136FA45957F70C2" box="[1289,1354,1476,1503]" pageId="24" pageNumber="99" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="80ACFD6DFFF2FFD84169FA4595A370C2" box="[1366,1430,1476,1503]" pageId="24" pageNumber="99" type="female">2 ♀♀</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD844A8FA6991D8731F" box="[151,493,1512,1538]" pageId="24" pageNumber="99">Yangmeiao Protection Station</collectingMunicipality>
,
<quantity id="51529B01FFF2FFD845C4FA699261731E" box="[507,596,1512,1539]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="24" pageNumber="99" unit="m" value="1240.0">
<elevation id="1D87D1D7FFF2FFD845C4FA699261731E" box="[507,596,1512,1539]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="24" pageNumber="99" unit="m" value="1240.0">1240 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFF2FFD8465CFA69931E731F" box="[611,811,1512,1538]" pageId="24" pageNumber="99">Rongshui County</collectingCounty>
,
<location id="9375603FFFF2FFD84707FA6993AD731F" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD84707FA6993AD731F" box="[824,920,1512,1538]" country="China" county="Rongshui County" municipality="Yangmeiao Protection Station" name="Liuzhou" pageId="24" pageNumber="99" stateProvince="Guangxi Zhuang">Liuzhou</location>
,
<date id="E2141024FFF2FFD84799FA69941D731E" box="[934,1064,1512,1539]" pageId="24" pageNumber="99" value="2015-07-27">
<collectingDate id="F250E9CCFFF2FFD84799FA69941D731E" box="[934,1064,1512,1539]" pageId="24" pageNumber="99" value="2015-07-27">27.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84056FA69953F731E" box="[1129,1290,1512,1539]" pageId="24" pageNumber="99">Meiqing Yang</collectorName>
and
<collectorName id="3B5F5332FFF2FFD8417AFA6990F1733A" pageId="24" pageNumber="99">Ga-Eun Lee</collectorName>
, genitalia slide nos. LGE
</materialsCitation>
<quantity id="51529B01FFF2FFD845E2F98D920D733B" box="[477,568,1548,1574]" metricMagnitude="4" metricUnit="m" metricValue="1.6011" pageId="24" pageNumber="99" unit="m" value="16011.0">16011m</quantity>
, LGE
<materialsCitation id="26C23CB9FFF2FFD84644F98D92F4733B" ID-GBIF-Occurrence="3427696378" accessionNumber="LGE16013" box="[635,705,1548,1574]" country="China" elevation="16012" pageId="24" pageNumber="99" specimenCount="1" stateProvince="Guangxi Zhuang" typeStatus="paratype">
<quantity id="51529B01FFF2FFD84644F98D92F4733B" box="[635,705,1548,1574]" metricMagnitude="4" metricUnit="m" metricValue="1.6012" pageId="24" pageNumber="99" unit="m" value="16012.0">
<elevation id="1D87D1D7FFF2FFD84644F98D92F4733B" box="[635,705,1548,1574]" metricMagnitude="4" metricUnit="m" metricValue="1.6012" pageId="24" pageNumber="99" unit="m" value="16012.0">16012</elevation>
</quantity>
</materialsCitation>
m, LGE16013
<materialsCitation id="26C23CB9FFF2FFD8475FF98D93C1733B" ID-GBIF-Occurrence="3427696382" accessionNumber="LGE19004" box="[864,1012,1548,1574]" country="China" pageId="24" pageNumber="99" specimenCount="1" stateProvince="Guangxi Zhuang" typeStatus="paratype">f, LGE19004</materialsCitation>
f.
<materialsCitation id="26C23CB9FFF2FFD84037F98D94307356" ID-GBIF-Occurrence="3427696351" accessionNumber="ZJL05455" collectingDate="2002-06-02" collectorName="Xinpu Wang" country="China" elevation="530" location="Mt. Fanjing" municipality="Heiwanhe" pageId="24" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="A4DEEAF6FFF2FFD84037F98D94D2733B" bold="true" box="[1032,1255,1548,1574]" pageId="24" pageNumber="99">
<collectingRegion id="546EF806FFF2FFD84037F98D94D6733B" box="[1032,1251,1548,1574]" country="China" name="Guizhou" pageId="24" pageNumber="99">Guizhou Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFF2FFD840D2F98D952C733A" box="[1261,1305,1548,1575]" pageId="24" pageNumber="99" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD8411BF98D95AD733B" box="[1316,1432,1548,1574]" pageId="24" pageNumber="99">Heiwanhe</collectingMunicipality>
,
<location id="9375603FFFF2FFD844A8F9B0912A7357" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD844A8F9B0912A7357" box="[151,287,1584,1611]" country="China" municipality="Heiwanhe" name="Mt. Fanjing" pageId="24" pageNumber="99" stateProvince="Guizhou">Mt. Fanjing</location>
,
<quantity id="51529B01FFF2FFD84516F9B191477356" box="[297,370,1584,1611]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="24" pageNumber="99" unit="m" value="530.0">
<elevation id="1D87D1D7FFF2FFD84516F9B191477356" box="[297,370,1584,1611]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="24" pageNumber="99" unit="m" value="530.0">530 m</elevation>
</quantity>
,
<date id="E2141024FFF2FFD84541F9B191DF7357" box="[382,490,1584,1610]" pageId="24" pageNumber="99" value="2002-06-02">
<collectingDate id="F250E9CCFFF2FFD84541F9B191DF7357" box="[382,490,1584,1610]" pageId="24" pageNumber="99" value="2002-06-02">2.vi.2002</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84619F9B192837356" box="[550,694,1584,1611]" pageId="24" pageNumber="99">Xinpu Wang</collectorName>
, genitalia slide no. ZJL05455
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD8402FF9B195B57373" ID-GBIF-Occurrence="3427696340" accessionNumber="LGE19002" collectingDate="2010-08-11" collectorName="Linlin Yang" country="China" county="Suiyang County" elevation="1500" latitude="28.23" location="Kuankuoshui Nature Reserves" longLatPrecision="740" longitude="107.18" pageId="24" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD8402FF9B194097356" box="[1040,1084,1584,1611]" pageId="24" pageNumber="99" type="male">1 ♂</specimenCount>
,
<location id="9375603FFFF2FFD84077F9B195A97356" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD84077F9B195A97356" box="[1096,1436,1584,1611]" country="China" county="Suiyang County" latitude="28.23" longLatPrecision="740" longitude="107.18" name="Kuankuoshui Nature Reserves" pageId="24" pageNumber="99" stateProvince="Guizhou Province">Kuankuoshui Nature Reserves</location>
(
<geoCoordinate id="F39E5023FFF2FFD844A0F9D591377373" box="[159,258,1620,1646]" degrees="28.23" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="28.23">28.23°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD8452FF9D591487373" box="[272,381,1620,1646]" degrees="107.18" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="107.18">107.18°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD845ABF9D591DB7372" box="[404,494,1620,1647]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="99" unit="m" value="1500.0">
<elevation id="1D87D1D7FFF2FFD845ABF9D591DB7372" box="[404,494,1620,1647]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="99" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFF2FFD845C2F9D592827373" box="[509,695,1620,1646]" pageId="24" pageNumber="99">Suiyang County</collectingCounty>
,
<date id="E2141024FFF2FFD846FAF9D5937B7373" box="[709,846,1620,1646]" pageId="24" pageNumber="99" value="2010-08-11">
<collectingDate id="F250E9CCFFF2FFD846FAF9D5937B7373" box="[709,846,1620,1646]" pageId="24" pageNumber="99" value="2010-08-11">11.viii.2010</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD847AFF9D5942E7372" box="[912,1051,1620,1647]" pageId="24" pageNumber="99">Linlin Yang</collectorName>
, genitalia slide no. LGE19002
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD841B1F9D595A2738E" ID-GBIF-Occurrence="3427696303" collectingDate="2015-07-19" collectorName="Meiqing Yang &amp; Ga-Eun Lee" country="China" county="Libo County" elevation="720" location="Shuizu Town" municipality="Dongdai" pageId="24" pageNumber="99" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD841B1F9D59099738E" pageId="24" pageNumber="99" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD84482F9F99115738F" box="[189,288,1656,1682]" pageId="24" pageNumber="99">Dongdai</collectingMunicipality>
,
<quantity id="51529B01FFF2FFD84511F9F9914F738E" box="[302,378,1656,1683]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="24" pageNumber="99" unit="m" value="720.0">
<elevation id="1D87D1D7FFF2FFD84511F9F9914F738E" box="[302,378,1656,1683]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="24" pageNumber="99" unit="m" value="720.0">720 m</elevation>
</quantity>
,
<location id="9375603FFFF2FFD845B7F9F99214738E" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD845B7F9F99214738E" box="[392,545,1656,1683]" country="China" county="Libo County" municipality="Dongdai" name="Shuizu Town" pageId="24" pageNumber="99" stateProvince="Guizhou Province">Shuizu Town</location>
,
<location id="9375603FFFF2FFD8460FF9F99289738F" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD8460FF9F99289738F" box="[560,700,1656,1682]" country="China" county="Libo County" municipality="Dongdai" name="Limingguan" pageId="24" pageNumber="99" stateProvince="Guizhou Province">Limingguan</location>
,
<collectingCounty id="7F744E68FFF2FFD846F6F9F99369738F" box="[713,860,1656,1682]" pageId="24" pageNumber="99">Libo County</collectingCounty>
,
<date id="E2141024FFF2FFD84756F9F993DE738E" box="[873,1003,1656,1683]" pageId="24" pageNumber="99" value="2015-07-19">
<collectingDate id="F250E9CCFFF2FFD84756F9F993DE738E" box="[873,1003,1656,1683]" pageId="24" pageNumber="99" value="2015-07-19">19.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84013F9F994FB738E" box="[1068,1230,1656,1683]" pageId="24" pageNumber="99">Meiqing Yang</collectorName>
and
<collectorName id="3B5F5332FFF2FFD84135F9F995A2738E" box="[1290,1431,1656,1683]" pageId="24" pageNumber="99">Ga-Eun Lee</collectorName>
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD844A8F91D92B773C7" ID-GBIF-Occurrence="3427696315" accessionNumber="LGE18709" collectingDate="2018-08-17" collectorName="Meiling Zheng &amp; Jiaqi Deng &amp; Xiaoju Zhu" country="China" elevation="528" latitude="27.82" location="Mt. Fanjing" longLatPrecision="741" longitude="108.78" municipality="Heiwanhe" pageId="24" pageNumber="99" specimenCount="5" specimenCount-male="5" stateProvince="Guizhou Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD844A8F91D90EF73AA" box="[151,218,1692,1719]" pageId="24" pageNumber="99" type="male">5 ♂♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD844D8F91D916C73AB" box="[231,345,1692,1718]" pageId="24" pageNumber="99">Heiwanhe</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFF2FFD84556F91D91F973AB" box="[361,460,1692,1718]" degrees="27.82" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="27.82">27.82°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD845E7F91D927173AB" box="[472,580,1692,1718]" degrees="108.78" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="108.78">108.78°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD84665F91D929173AA" box="[602,676,1692,1719]" metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" pageId="24" pageNumber="99" unit="m" value="528.0">
<elevation id="1D87D1D7FFF2FFD84665F91D929173AA" box="[602,676,1692,1719]" metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" pageId="24" pageNumber="99" unit="m" value="528.0">528 m</elevation>
</quantity>
,
<location id="9375603FFFF2FFD8468EF91C930F73AB" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD8468EF91C930F73AB" box="[689,826,1692,1719]" country="China" latitude="27.82" longLatPrecision="741" longitude="108.78" municipality="Heiwanhe" name="Mt. Fanjing" pageId="24" pageNumber="99" stateProvince="Guizhou Province">Mt. Fanjing</location>
, 16,
<date id="E2141024FFF2FFD8474DF91D93CE73AB" box="[882,1019,1692,1718]" pageId="24" pageNumber="99" value="2018-08-17">
<collectingDate id="F250E9CCFFF2FFD8474DF91D93CE73AB" box="[882,1019,1692,1718]" pageId="24" pageNumber="99" value="2018-08-17">17.viii.2018</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84005F91D94D073AB" box="[1082,1253,1692,1718]" pageId="24" pageNumber="99">Meiling Zheng</collectorName>
,
<collectorName id="3B5F5332FFF2FFD840CDF91D955973AA" box="[1266,1388,1692,1719]" pageId="24" pageNumber="99">Jiaqi Deng</collectorName>
and
<collectorName id="3B5F5332FFF2FFD844A8F941912873C7" box="[151,285,1728,1754]" pageId="24" pageNumber="99">Xiaoju Zhu</collectorName>
, genitalia slide no. LGE18709
</materialsCitation>
.
<materialsCitation id="26C23CB9FFF2FFD846B0F94193CE73E3" ID-GBIF-Occurrence="3427696329" accessionNumber="LGE17039, LGE18003" collectingDate="2008-03-30" collectorName="Bingbing Hu &amp; Haiyan Bai" country="China" elevation="810" location="Jianfengling" municipality="Tianchi" pageId="24" pageNumber="99" specimenCount="2" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<emphasis id="A4DEEAF6FFF2FFD846B0F941935173C7" bold="true" box="[655,868,1728,1754]" pageId="24" pageNumber="99">
<collectingRegion id="546EF806FFF2FFD846B0F941935573C7" box="[655,864,1728,1754]" country="China" name="Hainan" pageId="24" pageNumber="99">Hainan Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFF2FFD84751F941938773C6" box="[878,946,1728,1755]" pageId="24" pageNumber="99" type="male">2 ♂♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD847FFF941942273C7" box="[960,1047,1728,1754]" pageId="24" pageNumber="99">Tianchi</collectingMunicipality>
,
<quantity id="51529B01FFF2FFD84019F941944773C6" box="[1062,1138,1728,1755]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="24" pageNumber="99" unit="m" value="810.0">
<elevation id="1D87D1D7FFF2FFD84019F941944773C6" box="[1062,1138,1728,1755]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="24" pageNumber="99" unit="m" value="810.0">810 m</elevation>
</quantity>
,
<location id="9375603FFFF2FFD840BEF941953B73C7" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD840BEF941953B73C7" box="[1153,1294,1728,1754]" country="China" municipality="Tianchi" name="Jianfengling" pageId="24" pageNumber="99" stateProvince="Hainan">Jianfengling</location>
,
<date id="E2141024FFF2FFD84123F94195A273C7" box="[1308,1431,1728,1754]" pageId="24" pageNumber="99" value="2008-03-30">
<collectingDate id="F250E9CCFFF2FFD84123F94195A273C7" box="[1308,1431,1728,1754]" pageId="24" pageNumber="99" value="2008-03-30">30.iii.2008</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD844F8F965916D73E2" box="[199,344,1764,1791]" pageId="24" pageNumber="99">Bingbing Hu</collectorName>
and
<collectorName id="3B5F5332FFF2FFD845B1F965923B73E3" box="[398,526,1764,1790]" pageId="24" pageNumber="99">Haiyan Bai</collectorName>
, genitalia slide nos. LGE17039, LGE18003
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD84039F96591E0725B" ID-GBIF-Occurrence="3427696346" accessionNumber="LGE19010, LGE19011" collectingDate="2013-04-29" collectorName="Yinghui Sun &amp; Wei Guan &amp; Tengteng Liu" country="China" county="Ledong County" elevation="1050" latitude="18.74" location="Jianfengling" longLatPrecision="765" longitude="108.84" municipality="Tianchi" pageId="24" pageNumber="99" specimenCount="2" specimenCount-male="2" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD84039F965947273E2" box="[1030,1095,1764,1791]" pageId="24" pageNumber="99" type="male">2 ♂♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD8406DF965949273E3" box="[1106,1191,1764,1790]" pageId="24" pageNumber="99">Tianchi</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFF2FFD8408AF965952273E3" box="[1205,1303,1764,1790]" degrees="18.74" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="18.74">18.74°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD8411CF96595BA73E3" box="[1315,1423,1764,1790]" degrees="108.84" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="108.84">108.84°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD844A8F88990C5723E" box="[151,240,1800,1827]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="24" pageNumber="99" unit="m" value="1050.0">
<elevation id="1D87D1D7FFF2FFD844A8F88990C5723E" box="[151,240,1800,1827]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="24" pageNumber="99" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<location id="9375603FFFF2FFD844C1F88991BE723F" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD844C1F88991BE723F" box="[254,395,1800,1826]" country="China" county="Ledong County" latitude="18.74" longLatPrecision="765" longitude="108.84" municipality="Tianchi" name="Jianfengling" pageId="24" pageNumber="99" stateProvince="Hainan Province">Jianfengling</location>
,
<collectingCounty id="7F744E68FFF2FFD845A7F889927F723F" box="[408,586,1800,1826]" pageId="24" pageNumber="99">Ledong County</collectingCounty>
,
<date id="E2141024FFF2FFD84668F88992E5723F" box="[599,720,1800,1826]" pageId="24" pageNumber="99" value="2013-04-29">
<collectingDate id="F250E9CCFFF2FFD84668F88992E5723F" box="[599,720,1800,1826]" pageId="24" pageNumber="99" value="2013-04-29">29.iv.2013</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84732F88993AB723F" box="[781,926,1800,1826]" pageId="24" pageNumber="99">Yinghui Sun</collectorName>
,
<collectorName id="3B5F5332FFF2FFD84794F889942B723F" box="[939,1054,1800,1826]" pageId="24" pageNumber="99">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFF2FFD84068F88894C4723F" box="[1111,1265,1800,1827]" pageId="24" pageNumber="99">Tengteng Liu</collectorName>
, genitalia slide nos. LGE19010, LGE19011
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD845DFF8AD93367276" ID-GBIF-Occurrence="3427696341" collectingDate="2014-07" collectorName="Peixin Cong &amp; Linjie Liu &amp; Sha Hu" country="China" county="Ledong County" elevation="770" latitude="18.74" location="Jianfengling Nature Reserves" longLatPrecision="765" longitude="108.87" pageId="24" pageNumber="99" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD845DFF8AD923E725A" box="[480,523,1836,1863]" pageId="24" pageNumber="99" type="male">1 ♂</specimenCount>
,
<specimenCount id="80ACFD6DFFF2FFD84628F8AD9274725A" box="[535,577,1836,1863]" pageId="24" pageNumber="99" type="female">1 ♀</specimenCount>
,
<location id="9375603FFFF2FFD84670F8AD93AD725A" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD84670F8AD93AD725A" box="[591,920,1836,1863]" country="China" county="Ledong County" latitude="18.74" longLatPrecision="765" longitude="108.87" name="Jianfengling Nature Reserves" pageId="24" pageNumber="99" stateProvince="Hainan Province">Jianfengling Nature Reserves</location>
(
<geoCoordinate id="F39E5023FFF2FFD84799F8AD943C725B" box="[934,1033,1836,1862]" degrees="18.74" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="18.74">18.74°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD8402BF8AD94B5725B" box="[1044,1152,1836,1862]" degrees="108.87" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="108.87">108.87°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD840ABF8AD94E8725A" box="[1172,1245,1836,1863]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="24" pageNumber="99" unit="m" value="770.0">
<elevation id="1D87D1D7FFF2FFD840ABF8AD94E8725A" box="[1172,1245,1836,1863]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="24" pageNumber="99" unit="m" value="770.0">770 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFF2FFD840D7F8AD95AD725B" box="[1256,1432,1836,1862]" pageId="24" pageNumber="99">Ledong County</collectingCounty>
, 15,17.
<date id="E2141024FFF2FFD844E3F8D191097276" box="[220,316,1872,1899]" pageId="24" pageNumber="99" value="2014-07">
<collectingDate id="F250E9CCFFF2FFD844E3F8D191097276" box="[220,316,1872,1899]" pageId="24" pageNumber="99" value="2014-07">vii.2014</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD8454AF8D192347277" box="[373,513,1872,1898]" pageId="24" pageNumber="99">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFF2FFD84634F8D1924F7277" box="[523,634,1872,1898]" pageId="24" pageNumber="99">Linjie Liu</collectorName>
and
<collectorName id="3B5F5332FFF2FFD84692F8D193367276" box="[685,771,1872,1899]" pageId="24" pageNumber="99">Sha Hu</collectorName>
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD84733F8D194557293" ID-GBIF-Occurrence="3427696388" accessionNumber="LGE19019" collectingDate="2015-05-18" collectorName="Peixin Cong &amp; Wei Guan &amp; Sha Hu" country="China" elevation="742" latitude="18.88" location="Wuzhishan Nature Reserves" longLatPrecision="764" longitude="109.65" pageId="24" pageNumber="99" specimenCount="1" specimenCount-female="1" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD84733F8D193007276" box="[780,821,1872,1899]" pageId="24" pageNumber="99" type="female">1 ♀</specimenCount>
,
<location id="9375603FFFF2FFD8477EF8D1944F7276" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD8477EF8D1944F7276" box="[833,1146,1872,1899]" country="China" latitude="18.88" longLatPrecision="764" longitude="109.65" name="Wuzhishan Nature Reserves" pageId="24" pageNumber="99" stateProvince="Hainan Province">Wuzhishan Nature Reserves</location>
(
<geoCoordinate id="F39E5023FFF2FFD840B8F8D194DC7277" box="[1159,1257,1872,1898]" degrees="18.88" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="18.88">18.88°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD840CCF8D195547276" box="[1267,1377,1872,1899]" degrees="109.65" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="109.65">109.65°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD8414DF8D1909A7292" metricMagnitude="2" metricUnit="m" metricValue="7.42" pageId="24" pageNumber="99" unit="m" value="742.0">
<elevation id="1D87D1D7FFF2FFD8414DF8D1909A7292" metricMagnitude="2" metricUnit="m" metricValue="7.42" pageId="24" pageNumber="99" unit="m" value="742.0">742 m</elevation>
</quantity>
,
<date id="E2141024FFF2FFD84483F8F591187292" box="[188,301,1908,1935]" pageId="24" pageNumber="99" value="2015-05-18">
<collectingDate id="F250E9CCFFF2FFD84483F8F591187292" box="[188,301,1908,1935]" pageId="24" pageNumber="99" value="2015-05-18">18.v.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84555F8F591CD7293" box="[362,504,1908,1934]" pageId="24" pageNumber="99">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFF2FFD8463BF8F592437293" box="[516,630,1908,1934]" pageId="24" pageNumber="99">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFF2FFD84692F8F593317292" box="[685,772,1908,1935]" pageId="24" pageNumber="99">Sha Hu</collectorName>
, genitalia slide no. LGE19019
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD84053F8F5912D72C9" ID-GBIF-Occurrence="3427696336" accessionNumber="LGE15004" collectingDate="2015-05-24" collectingDateMax="2015-05-26" collectingDateMin="2015-05-24" collectorName="Peixin Cong &amp; Wei Guan &amp; Sha Hu" country="China" elevation="922" latitude="18.43" location="Diaoluoshan Nature Reserves" longLatPrecision="765" longitude="109.52" pageId="24" pageNumber="99" specimenCount="3" specimenCount-male="3" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD84053F8F5949B7292" box="[1132,1198,1908,1935]" pageId="24" pageNumber="99" type="male">3 ♂♂</specimenCount>
,
<location id="9375603FFFF2FFD84084F8F590C972AC" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD84084F8F590C972AC" country="China" latitude="18.43" longLatPrecision="765" longitude="109.52" name="Diaoluoshan Nature Reserves" pageId="24" pageNumber="99" stateProvince="Hainan Province">Diaoluoshan Nature Reserves</location>
(
<geoCoordinate id="F39E5023FFF2FFD84532F816915B72AC" box="[269,366,1943,1969]" degrees="18.43" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="18.43">18.43°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD84544F81791DD72AC" box="[379,488,1942,1969]" degrees="109.52" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="109.52">109.52°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD845C2F816927272AC" box="[509,583,1943,1969]" metricMagnitude="2" metricUnit="m" metricValue="9.22" pageId="24" pageNumber="99" unit="m" value="922.0">
<elevation id="1D87D1D7FFF2FFD845C2F816927272AC" box="[509,583,1943,1969]" metricMagnitude="2" metricUnit="m" metricValue="9.22" pageId="24" pageNumber="99" unit="m" value="922.0">922 m</elevation>
</quantity>
,
<date id="E2141024FFF2FFD8466BF81792DB72AC" box="[596,750,1942,1969]" pageId="24" pageNumber="99" value="2015-05-24" valueMax="2015-05-26" valueMin="2015-05-24">
<collectingDate id="F250E9CCFFF2FFD8466BF81792DB72AC" box="[596,750,1942,1969]" pageId="24" pageNumber="99" value="2015-05-24" valueMax="2015-05-26" valueMin="2015-05-24">24–26.v.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD84713F816938E72AC" box="[812,955,1943,1969]" pageId="24" pageNumber="99">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFF2FFD847F8F816940D72AC" box="[967,1080,1943,1969]" pageId="24" pageNumber="99">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFF2FFD8404FF81694FD72AC" box="[1136,1224,1943,1969]" pageId="24" pageNumber="99">Sha Hu</collectorName>
, genitalia slide no. LGE15004
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD8451DF838917972EB" ID-GBIF-Occurrence="3427696356" collectingDate="2015-07-12" collectorName="Qingyun Wang &amp; Suran Li &amp; Mengting Chen" country="China" elevation="787" latitude="18.73" location="Jianfengling" longLatPrecision="765" longitude="108.87" municipality="Tianchi" pageId="24" pageNumber="99" specimenCount="1" specimenCount-female="1" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD8451DF838917972C9" box="[290,332,1977,2004]" pageId="24" pageNumber="99" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD84566F838919B72CE" box="[345,430,1977,2003]" pageId="24" pageNumber="99">Tianchi</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFF2FFD84582F838922A72CE" box="[445,543,1977,2003]" degrees="18.73" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="18.73">18.73°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD84615F83892A272CE" box="[554,663,1977,2003]" degrees="108.87" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="108.87">108.87°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD84694F83892C172C9" box="[683,756,1977,2004]" metricMagnitude="2" metricUnit="m" metricValue="7.87" pageId="24" pageNumber="99" unit="m" value="787.0">
<elevation id="1D87D1D7FFF2FFD84694F83892C172C9" box="[683,756,1977,2004]" metricMagnitude="2" metricUnit="m" metricValue="7.87" pageId="24" pageNumber="99" unit="m" value="787.0">787 m</elevation>
</quantity>
,
<location id="9375603FFFF2FFD846C0F83893B972CE" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD846C0F83893B972CE" box="[767,908,1977,2003]" country="China" latitude="18.73" longLatPrecision="765" longitude="108.87" municipality="Tianchi" name="Jianfengling" pageId="24" pageNumber="99" stateProvince="Hainan Province">Jianfengling</location>
,
<date id="E2141024FFF2FFD847A7F838942C72C9" box="[920,1049,1977,2004]" pageId="24" pageNumber="99" value="2015-07-12">
<collectingDate id="F250E9CCFFF2FFD847A7F838942C72C9" box="[920,1049,1977,2004]" pageId="24" pageNumber="99" value="2015-07-12">12.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD8406AF838953472C9" box="[1109,1281,1977,2004]" pageId="24" pageNumber="99">Qingyun Wang</collectorName>
,
<collectorName id="3B5F5332FFF2FFD84133F838955872CE" box="[1292,1389,1977,2003]" pageId="24" pageNumber="99">Suran Li</collectorName>
and
<collectorName id="3B5F5332FFF2FFD844A8F85D917972EB" box="[151,332,2012,2038]" pageId="24" pageNumber="99">Mengting Chen</collectorName>
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF2FFD94567F85D928075AC" ID-GBIF-Occurrence="3427696317" accessionNumber="LGE16259" collectingDate="2016-08-08" collectorName="Xia Bai &amp; Shuonan Qian &amp; Wanding Qi" country="China" elevation="787" lastPageId="25" lastPageNumber="100" latitude="18.73" location="Jianfengling" longLatPrecision="765" longitude="108.87" municipality="Tianchi" pageId="24" pageNumber="99" specimenCount="1" specimenCount-male="1" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF2FFD84567F85D91B072EA" box="[344,389,2012,2039]" pageId="24" pageNumber="99" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF2FFD845ADF85D91D272EB" box="[402,487,2012,2038]" pageId="24" pageNumber="99">Tianchi</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFF2FFD845C8F85D926C72EB" box="[503,601,2012,2038]" degrees="18.73" direction="north" orientation="latitude" pageId="24" pageNumber="99" precision="555" value="18.73">18.73°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF2FFD84659F85D92E672EB" box="[614,723,2012,2038]" degrees="108.87" direction="east" orientation="longitude" pageId="24" pageNumber="99" precision="555" value="108.87">108.87°E</geoCoordinate>
),
<quantity id="51529B01FFF2FFD846D7F85D930672EA" box="[744,819,2012,2039]" metricMagnitude="2" metricUnit="m" metricValue="7.87" pageId="24" pageNumber="99" unit="m" value="787.0">
<elevation id="1D87D1D7FFF2FFD846D7F85D930672EA" box="[744,819,2012,2039]" metricMagnitude="2" metricUnit="m" metricValue="7.87" pageId="24" pageNumber="99" unit="m" value="787.0">787 m</elevation>
</quantity>
,
<location id="9375603FFFF2FFD8477FF85D93F872EB" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF2FFD8477FF85D93F872EB" box="[832,973,2012,2038]" country="China" latitude="18.73" longLatPrecision="765" longitude="108.87" municipality="Tianchi" name="Jianfengling" pageId="24" pageNumber="99" stateProvince="Hainan Province">Jianfengling</location>
,
<date id="E2141024FFF2FFD847E5F85D946372EB" box="[986,1110,2012,2038]" pageId="24" pageNumber="99" value="2016-08-08">
<collectingDate id="F250E9CCFFF2FFD847E5F85D946372EB" box="[986,1110,2012,2038]" pageId="24" pageNumber="99" value="2016-08-08">8.viii.2016</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF2FFD840ABF85D94DB72EB" box="[1172,1262,2012,2038]" pageId="24" pageNumber="99">Xia Bai</collectorName>
,
<collectorName id="3B5F5332FFF2FFD840C4F85D95A972EB" box="[1275,1436,2012,2038]" pageId="24" pageNumber="99">Shuonan Qian</collectorName>
and
<collectorName id="3B5F5332FFF3FFD944F7FF16916175AC" box="[200,340,151,177]" pageId="25" pageNumber="100">Wanding Qi</collectorName>
, genitalia slide no. LGE16259
</materialsCitation>
;
<materialsCitation id="26C23CB9FFF3FFD946FCFF16928375C9" ID-GBIF-Occurrence="3427696323" collectingDate="2018-06-12" collectorName="Ping Liu &amp; Xia Bai &amp; Shuai Yu." country="China" county="Ledong County" elevation="810" latitude="18.75" location="Jianfengling" longLatPrecision="765" longitude="108.87" pageId="25" pageNumber="100" specimenCount="1" specimenCount-female="1" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF3FFD946FCFF1692DA75AC" box="[707,751,151,177]" pageId="25" pageNumber="100" type="female">1 ♀</specimenCount>
,
<location id="9375603FFFF3FFD946C0FF1693BC75AC" LSID="urn:lsid:plazi:treatment:1E0387F2FFF2FFD944A8FC2F93277213:9375603FFFF3FFD946C0FF1693BC75AC" box="[767,905,151,177]" country="China" county="Ledong County" latitude="18.75" longLatPrecision="765" longitude="108.87" name="Jianfengling" pageId="25" pageNumber="100" stateProvince="Hainan Province">Jianfengling</location>
(
<geoCoordinate id="F39E5023FFF3FFD947A4FF1793C975AC" box="[923,1020,150,177]" degrees="18.75" direction="north" orientation="latitude" pageId="25" pageNumber="100" precision="555" value="18.75">18.75°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF3FFD94035FF16944275AC" box="[1034,1143,151,177]" degrees="108.87" direction="east" orientation="longitude" pageId="25" pageNumber="100" precision="555" value="108.87">108.87°E</geoCoordinate>
),
<quantity id="51529B01FFF3FFD940B2FF1694ED75AC" box="[1165,1240,151,177]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="25" pageNumber="100" unit="m" value="810.0">
<elevation id="1D87D1D7FFF3FFD940B2FF1694ED75AC" box="[1165,1240,151,177]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="25" pageNumber="100" unit="m" value="810.0">810 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFF3FFD940D9FF1695AC75AC" box="[1254,1433,151,177]" pageId="25" pageNumber="100">Ledong County</collectingCounty>
,
<date id="E2141024FFF3FFD944A8FF38912475CE" box="[151,273,185,211]" pageId="25" pageNumber="100" value="2018-06-12">
<collectingDate id="F250E9CCFFF3FFD944A8FF38912475CE" box="[151,273,185,211]" pageId="25" pageNumber="100" value="2018-06-12">12.vi.2018</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF3FFD94572FF38918775CE" box="[333,434,185,211]" pageId="25" pageNumber="100">Ping Liu</collectorName>
,
<collectorName id="3B5F5332FFF3FFD94583FF38922675CE" box="[444,531,185,211]" pageId="25" pageNumber="100">Xia Bai</collectorName>
and
<collectorName id="3B5F5332FFF3FFD94676FF38928375C9" box="[585,694,185,212]" pageId="25" pageNumber="100">Shuai Yu.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF3FFD944F8FF5D91CD74F4" pageId="25" pageNumber="100" type="diagnosis">
<paragraph id="961536E4FFF3FFD944F8FF5D91CD74F4" blockId="25.[151,1437,150,1807]" pageId="25" pageNumber="100">
<emphasis id="A4DEEAF6FFF3FFD944F8FF5D917775EB" bold="true" box="[199,322,220,246]" pageId="25" pageNumber="100">Diagnosis.</emphasis>
Superficially, this species closely resembles
<taxonomicName id="51AA4D67FFF3FFD94705FF5C93E875EB" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[826,989,220,246]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="crassimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD94705FF5C93E875EB" box="[826,989,220,246]" italics="true" pageId="25" pageNumber="100">T. crassimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD947DBFF5D940B75EB" bold="true" box="[996,1086,220,246]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD947DBFF5D940B75EB" box="[996,1086,220,246]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="51AA4D67FFF3FFD94073FF5C94D175EB" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1100,1252,220,246]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="acutimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD94073FF5C94D175EB" box="[1100,1252,220,246]" italics="true" pageId="25" pageNumber="100">T. acutimedia</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD940D4FF5D957075EB" bold="true" box="[1259,1349,220,246]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD940D4FF5D957075EB" box="[1259,1349,220,246]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="51AA4D67FFF3FFD9416CFF5C913F7404" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="breviaedeaga" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD9416CFF5C913F7404" italics="true" pageId="25" pageNumber="100">T. breviaedeaga</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD9452DFF7E91597404" bold="true" box="[274,364,255,281]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD9452DFF7E91597404" box="[274,364,255,281]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="51AA4D67FFF3FFD94544FF7E92187405" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[379,557,255,280]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="magniguttata" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD94544FF7E92187405" box="[379,557,255,280]" italics="true" pageId="25" pageNumber="100">T. magniguttata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD9460AFF7E92BA7404" bold="true" box="[565,655,255,281]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD9460AFF7E92BA7404" box="[565,655,255,281]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="51AA4D67FFF3FFD946A1FF7E93797404" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[670,844,255,281]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="perparviloba" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD946A1FF7E93797404" box="[670,844,255,281]" italics="true" pageId="25" pageNumber="100">T. perparviloba</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD9476BFF7E939A7404" bold="true" box="[852,943,255,281]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD9476BFF7E939A7404" box="[852,943,255,281]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="51AA4D67FFF3FFD94781FF7E94687404" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[958,1117,255,281]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="uncativalva" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD94781FF7E94687404" box="[958,1117,255,281]" italics="true" pageId="25" pageNumber="100">T. uncativalva</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD9405BFF7E948A7404" bold="true" box="[1124,1215,255,281]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD9405BFF7E948A7404" box="[1124,1215,255,281]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="51AA4D67FFF3FFD940F1FF7E95067404" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1230,1331,255,281]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="zhengi" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD940F1FF7E95067404" box="[1230,1331,255,281]" italics="true" pageId="25" pageNumber="100">T. zhengi</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD94104FF7E95A07404" bold="true" box="[1339,1429,255,281]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD94104FF7E95A07404" box="[1339,1429,255,281]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="51AA4D67FFF3FFD944F7FEA3917B7426" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[200,334,289,315]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="dissimilis" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD944F7FEA3917B7426" box="[200,334,289,315]" italics="true" pageId="25" pageNumber="100">T. dissimilis</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD94569FEA091857426" bold="true" box="[342,432,289,315]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD94569FEA091857426" box="[342,432,289,315]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and it is difficult to separate these species based only on external characters.
<taxonomicName id="51AA4D67FFF3FFD94118FEA090DE7443" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD94118FEA090DE7443" italics="true" pageId="25" pageNumber="100">Tenupalpa acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD944CDFEC5917E7443" bold="true" box="[242,331,324,350]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD944CDFEC5917E7443" box="[242,331,324,350]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is differentiated from other species by the shape of male sternum VIII, which is short and sub-rectangular compared to the round or pentagonal-shaped sternum of most other species. The male genitalia are distinguished by the slender valva, often with a minute spine near apex, and the apically pointed aedeagus. The female genitalia are similar to those of
<taxonomicName id="51AA4D67FFF3FFD945C2FE2C929774DB" box="[509,674,428,454]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFF3FFD945C2FE2C929774DB" box="[509,674,428,454]" italics="true" pageId="25" pageNumber="100">T. flavitermina</emphasis>
</taxonomicName>
, but the corpus bursae is narrower, and the signum is situated more anteriorly in
<taxonomicName id="51AA4D67FFF3FFD94517FE4E91A274F5" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[296,407,463,488]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD94517FE4E91A274F5" box="[296,407,463,488]" italics="true" pageId="25" pageNumber="100">T. acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD945A1FE4E91CD74F4" bold="true" box="[414,504,463,489]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD945A1FE4E91CD74F4" box="[414,504,463,489]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF3FFD944F8FE7091A270CB" pageId="25" pageNumber="100" type="description">
<paragraph id="961536E4FFF3FFD944F8FE709222776E" blockId="25.[151,1437,150,1807]" pageId="25" pageNumber="100">
<emphasis id="A4DEEAF6FFF3FFD944F8FE70916C7716" bold="true" box="[199,345,497,523]" pageId="25" pageNumber="100">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF3FFD9458CFE70923C7711" box="[435,521,497,524]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="25" pageNumber="100">Fig. 57</figureCitation>
).
<emphasis id="A4DEEAF6FFF3FFD94621FE70926E7716" box="[542,603,497,523]" italics="true" pageId="25" pageNumber="100">Head</emphasis>
: Silvery white, vertex sometimes mixed with grey. Labial palpus silvery white, palpomere II mixed with grey on outer surface; palpomere III as long as II, strongly mixed with fuscous. Antenna with scape silvery white; flagellum dark fuscous except basal half of dorsal surface silvery white, male cilia slightly longer than diameter.
</paragraph>
<paragraph id="961536E4FFF3FFD944F8FDFD92D976D3" blockId="25.[151,1437,150,1807]" pageId="25" pageNumber="100">
<emphasis id="A4DEEAF6FFF3FFD944F8FDFD912D778B" box="[199,280,636,662]" italics="true" pageId="25" pageNumber="100">Thorax</emphasis>
: Silvery white and light brown mixed. Tegula silvery white and light brown mixed, anterior margin usually suffused with fuscous. Wingspan 6.5–9.5 mm. Forewing silvery white to grey at base, gradually darkening toward apex; sometimes basal 1/3 silvery white and distal 2/3 dark grey; anterior edge of costa black; a large, fusiform orange patch extending along costa after middle, its inner margin nearly pointed, gradually broadened toward distal 1/6 of wing, then narrowed toward apical spot, below this an elongate orange spot on dorsum just after fold; a narrow white line between the patch and apical spot, running from costa to tornus; apical spot large, below this a small, indistinct black spot near tornus, between two spots tinged with orange along termen; fringe scales grey, mixed with cream along distal part of dorsum. Hindwing grey except apex black and near apex cream; fringe scales light grey, apical fringe cream with a black terminal band. Legs silvery white; hind tibia strongly mixed with fuscous after middle; mid- and hind tarsi mixed with fuscous.
</paragraph>
<paragraph id="961536E4FFF3FFD944F8FC5691A270CB" blockId="25.[151,1437,150,1807]" pageId="25" pageNumber="100">
<emphasis id="A4DEEAF6FFF3FFD944F8FC56910476EC" box="[199,305,983,1009]" italics="true" pageId="25" pageNumber="100">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFF3FFD9457BFC5691A876EC" box="[324,413,983,1009]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="25" pageNumber="100">Fig. 20</figureCitation>
): Male coremata as long as sternum VIII; sternum VIII short, sub-rectangular, posterior apex slightly convex medially. Male genitalia (
<figureCitation id="0E912A61FFF3FFD94694FC7892CB7109" box="[683,766,1017,1044]" captionStart="FIGURES 72–77" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 72–77. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="25" pageNumber="100">Fig. 76</figureCitation>
) with uncus small oval, with a longitudinal ridge medially. Gnathos hook short, pointed apically. Tegumen approximately 8–9 times as long as uncus; papilliform tubercles at middle, each tubercle bearing a long single hair (
<figureCitation id="0E912A61FFF3FFD946FFFBBE93237144" box="[704,790,1087,1113]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="25" pageNumber="100">Fig. 37</figureCitation>
). Valva slender, gradually narrowed from base to middle, then broadening to apex; costa with or without a minute spine before apex, rounded apically. Anellus lobe oval, approximately 1/5 length of valva; apical bristle longer than anellus lobe, sometimes sinuous, curved inward apically. Juxta with inner margin broadly convex, sometimes sinuous in distal half. Vinculum broad, posterior margin rounded. Saccus broad, sub-triangular, sometimes slightly constricted at middle, rounded apically. Aedeagus with basal 1/3 dilated, distal 2/3 slender and sinuous, acute apically. Female genitalia (
<figureCitation id="0E912A61FFF3FFD947E0FB6D9405701B" box="[991,1072,1260,1286]" captionStart="FIGURES 91–94" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[240,1347,186,1895]" captionTargetId="figure-22@20.[203,1385,181,1920]" captionTargetPageId="20" captionText="FIGURES 91–94. Female genitalia of Tenupalpa spp. 91. T. venustalis, slide no. LGE18596; 92. T. acutata sp. nov., paratype, slide no. LGE18846; 93. T. crassimedia sp. nov., paratype, slide no. LGE18019; 94. T. breviaedeaga sp. nov., paratype, slide no. LGE18037 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819836" httpUri="https://zenodo.org/record/5819836/files/figure.png" pageId="25" pageNumber="100">Fig. 92</figureCitation>
) with papillae anales simple and setose. Apophyses anteriores 1/3–1/2 length of apophyses posteriores. Tergum VIII with posterior margin deeply incised at middle, bearing a bundle of setae at the cleft; sternum VIII with posterior margin nearly straight, anterior margin convex medially. Ostium bursae round, located at middle of sternum VIII. Ductus bursae slender, slightly broadened after middle; colliculum reduced, situated at posterior 1/3, ductus seminalis arising from posterior 2/5. Corpus bursae elongate ovate, as long as ductus bursae, often constricted medially; signum small semicircular, placed at posterior 1/3.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF3FFD944F8FA5F93F470E7" box="[199,961,1502,1530]" pageId="25" pageNumber="100" type="distribution">
<paragraph id="961536E4FFF3FFD944F8FA5F93F470E7" blockId="25.[151,1437,150,1807]" box="[199,961,1502,1530]" pageId="25" pageNumber="100">
<emphasis id="A4DEEAF6FFF3FFD944F8FA5F916C70E5" bold="true" box="[199,345,1502,1528]" pageId="25" pageNumber="100">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFF3FFD94558FA5E919F70E4" box="[359,426,1503,1529]" name="China" pageId="25" pageNumber="100">China</collectingCountry>
(
<collectingRegion id="546EF806FFF3FFD94586FA5E923070E4" box="[441,517,1503,1529]" country="China" name="Fujian" pageId="25" pageNumber="100">Fujian</collectingRegion>
,
<collectingRegion id="546EF806FFF3FFD9462FFA5E924670E4" box="[528,627,1503,1529]" country="China" name="Guangxi" pageId="25" pageNumber="100">Guangxi</collectingRegion>
,
<collectingRegion id="546EF806FFF3FFD94640FA5E92D770E4" box="[639,738,1503,1529]" country="China" name="Guizhou" pageId="25" pageNumber="100">Guizhou</collectingRegion>
,
<collectingRegion id="546EF806FFF3FFD946D2FA5E937470E4" box="[749,833,1503,1529]" country="China" name="Hainan" pageId="25" pageNumber="100">Hainan</collectingRegion>
,
<collectingRegion id="546EF806FFF3FFD94773FA5E938070E7" box="[844,949,1503,1530]" country="China" name="Zhejiang" pageId="25" pageNumber="100">Zhejiang</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF3FFD944F8F98091307323" pageId="25" pageNumber="100" type="etymology">
<paragraph id="961536E4FFF3FFD944F8F98091307323" blockId="25.[151,1437,150,1807]" pageId="25" pageNumber="100">
<emphasis id="A4DEEAF6FFF3FFD944F8F980917D7306" bold="true" box="[199,328,1537,1563]" pageId="25" pageNumber="100">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFF3FFD94703F98393AE7306" box="[828,923,1538,1563]" italics="true" pageId="25" pageNumber="100">acutatus</emphasis>
(sharpened), referring to the acute apex of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF3FFD944F8F9C793277213" pageId="25" pageNumber="100" type="description">
<paragraph id="961536E4FFF3FFD944F8F9C793277213" blockId="25.[151,1437,150,1807]" pageId="25" pageNumber="100">
<emphasis id="A4DEEAF6FFF3FFD944F8F9C7910D737D" bold="true" box="[199,312,1606,1632]" pageId="25" pageNumber="100">Variation</emphasis>
.
<taxonomicName id="51AA4D67FFF3FFD9457AF9C69221737D" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[325,532,1607,1633]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD9457AF9C69221737D" box="[325,532,1607,1633]" italics="true" pageId="25" pageNumber="100">Tenupalpa acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD94626F9C69247737C" bold="true" box="[537,626,1607,1633]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD94626F9C69247737C" box="[537,626,1607,1633]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is variable in color pattern, with the forewings of some specimens gradually darkened from the base to the apex, and in others divided into white and grey areas. The termen of
<taxonomicName id="51AA4D67FFF3FFD94113F9EB95A9739E" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1324,1436,1642,1667]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD94113F9EB95A9739E" box="[1324,1436,1642,1667]" italics="true" pageId="25" pageNumber="100">T. acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF3FFD944A8F90D90C473BB" bold="true" box="[151,241,1676,1702]" pageId="25" pageNumber="100">
<taxonomicNameLabel id="BFED578DFFF3FFD944A8F90D90C473BB" box="[151,241,1676,1702]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is usually tinged with orange near the apical spot, and the dorsum is slightly suffused with orange beyond the fold, but these features can also be indistinct. At the same locality, specimens collected in May tend to be larger than those collected in August. In the male genitalia, specimens from
<collectingRegion id="546EF806FFF3FFD94790F950943A73F6" box="[943,1039,1745,1771]" country="China" name="Guangxi" pageId="25" pageNumber="100">Guangxi</collectingRegion>
often lack the small spine near the apex of valva, whereas those from
<collectingRegion id="546EF806FFF3FFD94621F97592B77213" box="[542,642,1780,1806]" country="China" name="Zhejiang" pageId="25" pageNumber="100">Zhejiang</collectingRegion>
have spines.
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="961536E4FFF3FFD944A8F8DA92BB726B" blockId="25.[151,917,1883,1979]" box="[151,654,1883,1910]" pageId="25" pageNumber="100">
<heading id="CD5D8188FFF3FFD944A8F8DA92BB726B" bold="true" box="[151,654,1883,1910]" fontSize="11" level="1" pageId="25" pageNumber="100" reason="1">
<emphasis id="A4DEEAF6FFF3FFD944A8F8DA92BB726B" bold="true" box="[151,654,1883,1910]" pageId="25" pageNumber="100">
<taxonomicName id="51AA4D67FFF3FFD944A8F8DA9216726B" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,547,1883,1910]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="100" phylum="Tracheophyta" rank="species" species="crassimedia" status="sp. nov.">
<emphasis id="A4DEEAF6FFF3FFD944A8F8DA9198726B" bold="true" box="[151,429,1883,1910]" italics="true" pageId="25" pageNumber="100">Tenupalpa crassimedia</emphasis>
Lee
<emphasis id="A4DEEAF6FFF3FFD945D5F8DD91CA726B" bold="true" box="[490,511,1884,1910]" italics="true" pageId="25" pageNumber="100">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFF3FFD9460EF8DD92BB726B" box="[561,654,1884,1910]" pageId="25" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF3FFD944A8F8FE93A072A6" pageId="25" pageNumber="100" type="description">
<paragraph id="961536E4FFF3FFD944A8F8FE91907284" blockId="25.[151,917,1883,1979]" box="[151,421,1918,1945]" pageId="25" pageNumber="100">
(
<figureCitation id="0E912A61FFF3FFD944A0F8FE90C37284" box="[159,246,1919,1945]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="25" pageNumber="100">Figs 21</figureCitation>
,
<figureCitation id="0E912A61FFF3FFD9453DF8FE91157284" box="[258,288,1919,1945]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="25" pageNumber="100">38</figureCitation>
,
<figureCitation id="0E912A61FFF3FFD94513F8FF917F7284" box="[300,330,1918,1945]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="25" pageNumber="100">58</figureCitation>
,
<figureCitation id="0E912A61FFF3FFD94569F8FE91417284" box="[342,372,1919,1945]" captionStart="FIGURES 72–77" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 72–77. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="25" pageNumber="100">77</figureCitation>
,
<figureCitation id="0E912A61FFF3FFD945BFF8FE91A87284" box="[384,413,1919,1945]" captionStart="FIGURES 91–94" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[240,1347,186,1895]" captionTargetId="figure-22@20.[203,1385,181,1920]" captionTargetPageId="20" captionText="FIGURES 91–94. Female genitalia of Tenupalpa spp. 91. T. venustalis, slide no. LGE18596; 92. T. acutata sp. nov., paratype, slide no. LGE18846; 93. T. crassimedia sp. nov., paratype, slide no. LGE18019; 94. T. breviaedeaga sp. nov., paratype, slide no. LGE18037 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819836" httpUri="https://zenodo.org/record/5819836/files/figure.png" pageId="25" pageNumber="100">93</figureCitation>
)
</paragraph>
<paragraph id="961536E4FFF3FFD944A8F82293A072A6" blockId="25.[151,917,1883,1979]" box="[151,917,1955,1979]" pageId="25" pageNumber="100">
<uri id="E23B3AE6FFF3FFD944A8F82293A072A6" box="[151,917,1955,1979]" pageId="25" pageNumber="100">
urn:lsid:zoobank.org:act:
<uuid id="E20C0C31FFF3FFD945A5F82293A072A6" box="[410,917,1955,1979]" pageId="25" pageNumber="100">2911381A-C556-4060-975A-8230DC7B6D58</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDA44A8FF16942174DA" pageId="26" pageNumber="101" type="materials_examined">
<paragraph id="961536E4FFF0FFDA44A8FF16942174DA" blockId="26.[151,1437,150,1606]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44A8FF16917375AD" bold="true" box="[151,326,150,177]" pageId="26" pageNumber="101">Type material.</emphasis>
<materialsCitation id="26C23CB9FFF0FFDA4571FF16937175C9" ID-GBIF-Occurrence="3427696417" accessionNumber="LGE18015" collectingDate="2007-05-31" collectorName="Zhiwei Zhang &amp; Weichun Li" country="China" elevation="940" latitude="18.67" location="Mt. Diaoluo" longLatPrecision="765" longitude="109.94" pageId="26" pageNumber="101" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="49118846FFF0FFDA4571FF16918375AC" box="[334,438,151,177]" pageId="26" pageNumber="101" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFF0FFDA4581FF1691E075AC" box="[446,469,151,177]" pageId="26" pageNumber="101" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFF0FFDA45DDFF16920B75AC" box="[482,574,151,177]" name="China" pageId="26" pageNumber="101">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFF0FFDA4675FF16933D75AC" box="[586,776,151,177]" country="China" name="Hainan" pageId="26" pageNumber="101">Hainan Province</collectingRegion>
,
<location id="9375603FFFF0FFDA472AFF1693AA75AC" LSID="urn:lsid:plazi:treatment:1E0387F2FFF3FFDA44A8F8DA92D2735B:9375603FFFF0FFDA472AFF1693AA75AC" box="[789,927,151,177]" country="China" latitude="18.67" longLatPrecision="765" longitude="109.94" name="Mt. Diaoluo" pageId="26" pageNumber="101" stateProvince="Hainan">Mt. Diaoluo</location>
(
<geoCoordinate id="F39E5023FFF0FFDA4790FF16942475AC" box="[943,1041,151,177]" degrees="18.67" direction="north" orientation="latitude" pageId="26" pageNumber="101" precision="555" value="18.67">18.67°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF0FFDA4021FF1694BF75AC" box="[1054,1162,151,177]" degrees="109.94" direction="east" orientation="longitude" pageId="26" pageNumber="101" precision="555" value="109.94">109.94°E</geoCoordinate>
),
<quantity id="51529B01FFF0FFDA40A0FF1694DC75AC" box="[1183,1257,151,177]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="26" pageNumber="101" unit="m" value="940.0">
<elevation id="1D87D1D7FFF0FFDA40A0FF1694DC75AC" box="[1183,1257,151,177]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="26" pageNumber="101" unit="m" value="940.0">940 m</elevation>
</quantity>
,
<date id="E2141024FFF0FFDA40C9FF16955275AC" box="[1270,1383,151,177]" pageId="26" pageNumber="101" value="2007-05-31">
<collectingDate id="F250E9CCFFF0FFDA40C9FF16955275AC" box="[1270,1383,151,177]" pageId="26" pageNumber="101" value="2007-05-31">31.v.2007</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF0FFDA44A8FF38910075CE" box="[151,309,185,211]" pageId="26" pageNumber="101">Zhiwei Zhang</collectorName>
and
<collectorName id="3B5F5332FFF0FFDA4556FF3891D875CE" box="[361,493,185,211]" pageId="26" pageNumber="101">Weichun Li</collectorName>
, genitalia slide no. LGE18015
</materialsCitation>
.
<materialsCitation id="26C23CB9FFF0FFDA4771FF38930C75EB" ID-GBIF-Occurrence="3427696435" accessionNumber="LGE18015" collectingDate="2007-05-31" collectingDateMax="2007-06-02" collectingDateMin="2007-05-31" collectorName="Zhiwei Zhang &amp; Weichun Li" country="China" elevation="940" latitude="18.67" location="Mt. Diaoluo" longLatPrecision="765" longitude="109.94" pageId="26" pageNumber="101" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<emphasis id="A4DEEAF6FFF0FFDA4771FF38949075CE" bold="true" box="[846,1189,185,211]" pageId="26" pageNumber="101">
<typeStatus id="49118846FFF0FFDA4771FF3893FD75CE" box="[846,968,185,211]" pageId="26" pageNumber="101" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="546EF806FFF0FFDA47ECFF38949575CE" box="[979,1184,185,211]" country="China" name="Hainan" pageId="26" pageNumber="101">Hainan Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFF0FFDA4094FF3894DE75C9" box="[1195,1259,185,212]" pageId="26" pageNumber="101" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="80ACFD6DFFF0FFDA40C8FF38951575C9" box="[1271,1312,185,212]" pageId="26" pageNumber="101" type="female">1 ♀</specimenCount>
, same data as holotype except
<date id="E2141024FFF0FFDA454FFF5D922E75EB" box="[368,539,220,246]" pageId="26" pageNumber="101" value="2007-05-31" valueMax="2007-06-02" valueMin="2007-05-31">
<collectingDate id="F250E9CCFFF0FFDA454FFF5D922E75EB" box="[368,539,220,246]" pageId="26" pageNumber="101" value="2007-05-31" valueMax="2007-06-02" valueMin="2007-05-31">31.v–2.vi.2007</collectingDate>
</date>
, genitalia slide nos. LGE
</materialsCitation>
<quantity id="51529B01FFF0FFDA4706FF5D93A075EB" box="[825,917,220,246]" metricMagnitude="4" metricUnit="m" metricValue="1.7038" pageId="26" pageNumber="101" unit="m" value="17038.0">17038m</quantity>
, LGE
<materialsCitation id="26C23CB9FFF0FFDA47E6FF5D942A75EB" ID-GBIF-Occurrence="3427696365" accessionNumber="LGE18017" box="[985,1055,220,246]" country="China" elevation="18016" pageId="26" pageNumber="101" specimenCount="1" stateProvince="Hainan Province" typeStatus="paratype">
<quantity id="51529B01FFF0FFDA47E6FF5D942A75EB" box="[985,1055,220,246]" metricMagnitude="4" metricUnit="m" metricValue="1.8015999999999999" pageId="26" pageNumber="101" unit="m" value="18016.0">
<elevation id="1D87D1D7FFF0FFDA47E6FF5D942A75EB" box="[985,1055,220,246]" metricMagnitude="4" metricUnit="m" metricValue="1.8015999999999999" pageId="26" pageNumber="101" unit="m" value="18016.0">18016</elevation>
</quantity>
</materialsCitation>
m, LGE18017f;
<materialsCitation id="26C23CB9FFF0FFDA40E9FF5D91747426" ID-GBIF-Occurrence="3427696395" accessionNumber="LGE19020" collectingDate="2015-05-27" collectorName="Peixin Cong &amp; Wei Guan &amp; Sha Hu" country="China" elevation="922" latitude="18.43" location="Diaoluoshan Nature Reserves" longLatPrecision="765" longitude="109.52" pageId="26" pageNumber="101" specimenCount="1" specimenCount-female="1" stateProvince="Hainan Province" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF0FFDA40E9FF5D953575EA" box="[1238,1280,220,247]" pageId="26" pageNumber="101" type="female">1 ♀</specimenCount>
,
<location id="9375603FFFF0FFDA4130FF5D91657404" LSID="urn:lsid:plazi:treatment:1E0387F2FFF3FFDA44A8F8DA92D2735B:9375603FFFF0FFDA4130FF5D91657404" country="China" latitude="18.43" longLatPrecision="765" longitude="109.52" name="Diaoluoshan Nature Reserves" pageId="26" pageNumber="101" stateProvince="Hainan Province">Diaoluoshan Nature Reserves</location>
(
<geoCoordinate id="F39E5023FFF0FFDA455EFF7E91F67404" box="[353,451,255,281]" degrees="18.43" direction="north" orientation="latitude" pageId="26" pageNumber="101" precision="555" value="18.43">18.43°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF0FFDA45F0FF7F92097404" box="[463,572,254,281]" degrees="109.52" direction="east" orientation="longitude" pageId="26" pageNumber="101" precision="555" value="109.52">109.52°E</geoCoordinate>
),
<quantity id="51529B01FFF0FFDA466EFF7E92AE7404" box="[593,667,255,281]" metricMagnitude="2" metricUnit="m" metricValue="9.22" pageId="26" pageNumber="101" unit="m" value="922.0">
<elevation id="1D87D1D7FFF0FFDA466EFF7E92AE7404" box="[593,667,255,281]" metricMagnitude="2" metricUnit="m" metricValue="9.22" pageId="26" pageNumber="101" unit="m" value="922.0">922 m</elevation>
</quantity>
,
<date id="E2141024FFF0FFDA4697FF7F932C7404" box="[680,793,254,281]" pageId="26" pageNumber="101" value="2015-05-27">
<collectingDate id="F250E9CCFFF0FFDA4697FF7F932C7404" box="[680,793,254,281]" pageId="26" pageNumber="101" value="2015-05-27">27.v.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF0FFDA4768FF7E93D07404" box="[855,997,255,281]" pageId="26" pageNumber="101">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFF0FFDA47CDFF7E94567404" box="[1010,1123,255,281]" pageId="26" pageNumber="101">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFF0FFDA40A4FF7E94C67404" box="[1179,1267,255,281]" pageId="26" pageNumber="101">Sha Hu</collectorName>
, genitalia slide no. LGE19020
</materialsCitation>
.
<emphasis id="A4DEEAF6FFF0FFDA4575FEA093207426" bold="true" box="[330,789,289,315]" pageId="26" pageNumber="101">
<materialsCitation id="26C23CB9FFF0FFDA4575FEA0933A7426" ID-GBIF-Occurrence="3427696364" box="[330,783,289,315]" country="China" location="Autonomous Region" pageId="26" pageNumber="101" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
<collectingRegion id="546EF806FFF0FFDA4575FEA092217426" box="[330,532,289,315]" country="China" name="Guangxi" pageId="26" pageNumber="101">Guangxi Zhuang</collectingRegion>
<location id="9375603FFFF0FFDA4626FEA0933A7426" LSID="urn:lsid:plazi:treatment:1E0387F2FFF3FFDA44A8F8DA92D2735B:9375603FFFF0FFDA4626FEA0933A7426" box="[537,783,289,315]" country="China" name="Autonomous Region" pageId="26" pageNumber="101" stateProvince="Guangxi">Autonomous Region</location>
</materialsCitation>
:
</emphasis>
<materialsCitation id="26C23CB9FFF0FFDA4724FEA0950C7443" ID-GBIF-Occurrence="3427696427" accessionNumber="LGE18019" collectingDate="2015-07-26" collectorName="Mujie Qi &amp; Shengnan Zhao" country="China" county="Huilongsi" elevation="1376" latitude="25.91" location="Guilin" longLatPrecision="746" longitude="110.47" municipality="Mao'ershan Nature Reserves" pageId="26" pageNumber="101" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi Zhuang" typeStatus="paratype">
<specimenCount id="80ACFD6DFFF0FFDA4724FEA093717421" box="[795,836,289,316]" pageId="26" pageNumber="101" type="female">1 ♀</specimenCount>
,
<collectingCounty id="7F744E68FFF0FFDA476EFEA0938A7426" box="[849,959,289,315]" pageId="26" pageNumber="101">Huilongsi</collectingCounty>
(
<geoCoordinate id="F39E5023FFF0FFDA47F1FEA094057421" box="[974,1072,289,316]" degrees="25.91" direction="north" orientation="latitude" pageId="26" pageNumber="101" precision="555" value="25.91">25.91°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF0FFDA4004FEA094927426" box="[1083,1191,289,315]" degrees="110.47" direction="east" orientation="longitude" pageId="26" pageNumber="101" precision="555" value="110.47">110.47°E</geoCoordinate>
),
<quantity id="51529B01FFF0FFDA4085FEA095257421" box="[1210,1296,289,316]" metricMagnitude="3" metricUnit="m" metricValue="1.376" pageId="26" pageNumber="101" unit="m" value="1376.0">
<elevation id="1D87D1D7FFF0FFDA4085FEA095257421" box="[1210,1296,289,316]" metricMagnitude="3" metricUnit="m" metricValue="1.376" pageId="26" pageNumber="101" unit="m" value="1376.0">1376 m</elevation>
</quantity>
,
<collectingMunicipality id="7671AC9EFFF0FFDA4124FEA091607442" pageId="26" pageNumber="101">Mao'ershan Nature Reserves</collectingMunicipality>
,
<location id="9375603FFFF0FFDA455DFEC591997443" LSID="urn:lsid:plazi:treatment:1E0387F2FFF3FFDA44A8F8DA92D2735B:9375603FFFF0FFDA455DFEC591997443" box="[354,428,324,350]" country="China" county="Huilongsi" latitude="25.91" longLatPrecision="746" longitude="110.47" municipality="Mao'ershan Nature Reserves" name="Guilin" pageId="26" pageNumber="101" stateProvince="Guangxi Zhuang">Guilin</location>
,
<date id="E2141024FFF0FFDA4584FEC592087442" box="[443,573,324,351]" pageId="26" pageNumber="101" value="2015-07-26">
<collectingDate id="F250E9CCFFF0FFDA4584FEC592087442" box="[443,573,324,351]" pageId="26" pageNumber="101" value="2015-07-26">26.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF0FFDA4642FEC592D37443" box="[637,742,324,350]" pageId="26" pageNumber="101">Mujie Qi</collectorName>
and
<collectorName id="3B5F5332FFF0FFDA471EFEC593E07443" box="[801,981,324,350]" pageId="26" pageNumber="101">Shengnan Zhao</collectorName>
, genitalia slide no. LGE18019
</materialsCitation>
.
<materialsCitation id="26C23CB9FFF0FFDA417AFEC59516749C" ID-GBIF-Occurrence="3427696398" accessionNumber="ZJL05252" collectingDate="1978-07-18" country="China" location="Ciping" pageId="26" pageNumber="101" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="paratype">
<emphasis id="A4DEEAF6FFF0FFDA417AFEC5913F749D" bold="true" pageId="26" pageNumber="101">
<collectingRegion id="546EF806FFF0FFDA417AFEC59130749D" country="China" name="Jiangxi" pageId="26" pageNumber="101">Jiangxi Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFF0FFDA452BFEE69176749C" box="[276,323,359,385]" pageId="26" pageNumber="101" type="male">1 ♂</specimenCount>
,
<location id="9375603FFFF0FFDA456DFEE69194749C" LSID="urn:lsid:plazi:treatment:1E0387F2FFF3FFDA44A8F8DA92D2735B:9375603FFFF0FFDA456DFEE69194749C" box="[338,417,359,385]" country="China" name="Ciping" pageId="26" pageNumber="101" stateProvince="Jiangxi">Ciping</location>
,
<date id="E2141024FFF0FFDA458FFEE69204749C" box="[432,561,359,385]" pageId="26" pageNumber="101" value="1978-07-18">
<collectingDate id="F250E9CCFFF0FFDA458FFEE69204749C" box="[432,561,359,385]" pageId="26" pageNumber="101" value="1978-07-18">18.vii.1978</collectingDate>
</date>
, leg. Chinese Academy of Science, genitalia slide no. ZJL05252
</materialsCitation>
.
<materialsCitation id="26C23CB9FFF0FFDA410EFEE7942474DA" ID-GBIF-Occurrence="3427696385" accessionNumber="LGE16256" collectingDate="2016-08-12" collectorName="Qingyun Wang &amp; Meiqing Yang &amp; Ping Liu" country="China" county="Jingning County" elevation="1301" latitude="27.67" location="Wangdongyang Wetlands Reserves" longLatPrecision="742" longitude="119.63" pageId="26" pageNumber="101" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang" typeStatus="paratype">
<emphasis id="A4DEEAF6FFF0FFDA410EFEE7913F74BE" bold="true" pageId="26" pageNumber="101">
<collectingRegion id="546EF806FFF0FFDA410EFEE7913074BE" country="China" name="Zhejiang" pageId="26" pageNumber="101">Zhejiang Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFF0FFDA452CFE08917474B9" box="[275,321,393,420]" pageId="26" pageNumber="101" type="male">1 ♂</specimenCount>
,
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, leg.
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,
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and
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, genitalia slide no. LGE16256
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDA44F8FE519224774B" pageId="26" pageNumber="101" type="diagnosis">
<paragraph id="961536E4FFF0FFDA44F8FE519224774B" blockId="26.[151,1437,150,1606]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44F8FE51917774F7" bold="true" box="[199,322,464,490]" pageId="26" pageNumber="101">Diagnosis.</emphasis>
Superficially, this species is not easily distinguished from its allies, but the orange patch of the forewing is somewhat smaller than that of congeners. The male genitalia are characterized by the elongated pentagonal uncus, the medially thickened gnathos, and a short projection of the juxta. In the female genitalia, the signum is larger than that of related species.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDA44F8FDE1933270C7" pageId="26" pageNumber="101" type="description">
<paragraph id="961536E4FFF0FFDA44F8FDE190CA77FB" blockId="26.[151,1437,150,1606]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44F8FDE1916C7767" bold="true" box="[199,345,608,634]" pageId="26" pageNumber="101">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF0FFDA458FFDE192307766" box="[432,517,608,635]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="26" pageNumber="101">Fig. 58</figureCitation>
).
<emphasis id="A4DEEAF6FFF0FFDA4626FDE192637767" box="[537,598,608,634]" italics="true" pageId="26" pageNumber="101">Head</emphasis>
: Silvery white. Labial palpus silvery white, palpomere II mixed with light brown on outer surface; palpomere III as long as II, mixed with light brown and fuscous. Antenna with scape silvery white; flagellum dark fuscous except basal 1/3 of dorsal surface silvery white, male cilia slightly longer than diameter.
</paragraph>
<paragraph id="961536E4FFF0FFDA44F8FD71911876FF" blockId="26.[151,1437,150,1606]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44F8FD71912D7617" box="[199,280,752,778]" italics="true" pageId="26" pageNumber="101">Thorax</emphasis>
: Dorsum and tegula silvery white. Wingspan 7.5–8.5 mm; forewing with basal 1/3 silvery white mixed with grey, anterior edge of costa black, distal 2/3 grey; an elliptical orange patch in distal 1/3 of wing between costa and posterior 1/3, its outer margin extending almost to apical spot; silvery white between patch and apical spot, extended along termen as a narrow line; below apical spot a smaller black spot on termen; fringe scales grey. Hindwing grey except apex black; fringe scales grey. Legs silvery white to light grey; mid-tibia sometimes slightly suffused with fuscous near apex; hind tibia strongly mixed with fuscous after middle; mid- and hind tarsi mixed with fuscous.
</paragraph>
<paragraph id="961536E4FFF0FFDA44F8FC6D933270C7" blockId="26.[151,1437,150,1606]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44F8FC6D9104711B" box="[199,305,1004,1030]" italics="true" pageId="26" pageNumber="101">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFF0FFDA457FFC6D91A1711B" box="[320,404,1004,1030]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="26" pageNumber="101">Fig. 21</figureCitation>
): Male coremata slightly shorter than sternum VIII; sternum VIII pentagonal, approximately 1/7 length of abdomen. Male genitalia (
<figureCitation id="0E912A61FFF0FFDA466AFB9192927136" box="[597,679,1040,1067]" captionStart="FIGURES 72–77" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 72–77. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="26" pageNumber="101">Fig. 77</figureCitation>
) with uncus small, elongated pentagonal, protruded medially. Gnathos hook short, thickened at middle, pointed apically. Tegumen approximately 5 times as long as uncus; papilliform tubercles at middle, each bearing a long single hair (
<figureCitation id="0E912A61FFF0FFDA46D6FBD9930B716F" box="[745,830,1112,1138]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="26" pageNumber="101">Fig. 38</figureCitation>
). Valva slender, uniformly elongate from base to near apex, curved inward at middle, slightly broadened and rounded apically. Anellus lobe ovate, approximately 1/5 length of valva; apical bristle slightly longer than anellus lobe, strongly curved inward. Juxta with a short projection at distal 1/3 of inner margin. Vinculum broad, posterior margin rounded. Saccus sub-triangular, broad basally, rounded apically. Aedeagus with basal 2/5 globular, distal 3/5 slender and slightly curved downward. Female genitalia (
<figureCitation id="0E912A61FFF0FFDA44E8FA8D911D703B" box="[215,296,1292,1318]" captionStart="FIGURES 91–94" captionStartId="20.[151,264,1943,1967]" captionTargetBox="[240,1347,186,1895]" captionTargetId="figure-22@20.[203,1385,181,1920]" captionTargetPageId="20" captionText="FIGURES 91–94. Female genitalia of Tenupalpa spp. 91. T. venustalis, slide no. LGE18596; 92. T. acutata sp. nov., paratype, slide no. LGE18846; 93. T. crassimedia sp. nov., paratype, slide no. LGE18019; 94. T. breviaedeaga sp. nov., paratype, slide no. LGE18037 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819836" httpUri="https://zenodo.org/record/5819836/files/figure.png" pageId="26" pageNumber="101">Fig. 93</figureCitation>
) with papillae anales simple and setose. Apophyses anteriores approximately 1/3 length of apophyses posteriores. Tergum VIII with posterior margin deeply incised at middle; sternum VIII shorter than tergum VIII, broadly convex anteriorly. Ostium bursae round, situated at middle of sternum VIII. Ductus bursae slender, slightly broadened near corpus bursae, colliculum in posterior 1/3, ductus seminalis arising from posterior 2/5. Corpus bursae elongate-ovate, slightly longer than ductus bursae, constricted at middle; signum elliptical or elongate diamondshaped, between posterior 1/6 and 1/2 of corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDA44F8FA65936E70E2" box="[199,859,1508,1535]" pageId="26" pageNumber="101" type="distribution">
<paragraph id="961536E4FFF0FFDA44F8FA65936E70E2" blockId="26.[151,1437,150,1606]" box="[199,859,1508,1535]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44F8FA65916C70E3" bold="true" box="[199,345,1508,1534]" pageId="26" pageNumber="101">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFF0FFDA4558FA65919F70E3" box="[359,426,1508,1534]" name="China" pageId="26" pageNumber="101">China</collectingCountry>
(
<collectingRegion id="546EF806FFF0FFDA4586FA65922B70E3" box="[441,542,1508,1534]" country="China" name="Guangxi" pageId="26" pageNumber="101">Guangxi</collectingRegion>
,
<collectingRegion id="546EF806FFF0FFDA4616FA65924870E3" box="[553,637,1508,1534]" country="China" name="Hainan" pageId="26" pageNumber="101">Hainan</collectingRegion>
,
<collectingRegion id="546EF806FFF0FFDA46B7FA6592E970E3" box="[648,732,1508,1534]" country="China" name="Jiangxi" pageId="26" pageNumber="101">Jiangxi</collectingRegion>
,
<collectingRegion id="546EF806FFF0FFDA46D8FA65936570E2" box="[743,848,1508,1535]" country="China" name="Zhejiang" pageId="26" pageNumber="101">Zhejiang</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF0FFDA44F8F98992D2735B" pageId="26" pageNumber="101" type="etymology">
<paragraph id="961536E4FFF0FFDA44F8F98992D2735B" blockId="26.[151,1437,150,1606]" pageId="26" pageNumber="101">
<emphasis id="A4DEEAF6FFF0FFDA44F8F989917D733F" bold="true" box="[199,328,1544,1570]" pageId="26" pageNumber="101">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFF0FFDA477AF98893AC733F" box="[837,921,1545,1570]" italics="true" pageId="26" pageNumber="101">crassus</emphasis>
(thickened) and
<emphasis id="A4DEEAF6FFF0FFDA4068F9899493733F" box="[1111,1190,1544,1570]" italics="true" pageId="26" pageNumber="101">medius</emphasis>
(middle), referring to the medially thickened gnathos in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="961536E4FFF4FFDE44A8FD1F927A77A4" blockId="30.[151,917,670,766]" box="[151,591,670,697]" pageId="30" pageNumber="105">
<heading id="CD5D8188FFF4FFDE44A8FD1F927A77A4" bold="true" box="[151,591,670,697]" fontSize="11" level="1" pageId="30" pageNumber="105" reason="1">
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<taxonomicName id="51AA4D67FFF4FFDE44A8FD1F91D177A5" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,484,670,697]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="30" pageNumber="105" phylum="Tracheophyta" rank="species" species="zhengi" status="sp. nov.">
<emphasis id="A4DEEAF6FFF4FFDE44A8FD1F915B77A4" bold="true" box="[151,366,670,697]" italics="true" pageId="30" pageNumber="105">Tenupalpa zhengi</emphasis>
Lee
<emphasis id="A4DEEAF6FFF4FFDE4595FD1E918A77A4" bold="true" box="[426,447,671,697]" italics="true" pageId="30" pageNumber="105">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFF4FFDE45CEFD1E927A77A4" box="[497,591,671,697]" pageId="30" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
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(
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,
<figureCitation id="0E912A61FFF4FFDE453DFD40911477C6" box="[258,289,705,731]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="30" pageNumber="105">44</figureCitation>
,
<figureCitation id="0E912A61FFF4FFDE4513FD40917F77C6" box="[300,330,705,731]" captionStart="FIGURES 62–65" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[151,1435,181,1125]" captionTargetId="figure-22@12.[151,1435,181,1125]" captionTargetPageId="12" captionText="FIGURES 62–65. Adults of Tenupalpa spp. 62. perparviloba sp. nov., male, holotype; 63. T. uncativalva sp. nov., male, holotype; 64. T. zhengi sp. nov., male, holotype; 65. T. dissimilis sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="30" pageNumber="105">64</figureCitation>
,
<figureCitation id="0E912A61FFF4FFDE4569FD40914677C6" box="[342,371,705,731]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="30" pageNumber="105">83</figureCitation>
)
</paragraph>
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<paragraph id="961536E4FFF4FFDE44A8FCA89455767B" blockId="30.[151,1437,809,1702]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44A8FCA89173765E" bold="true" box="[151,326,809,835]" pageId="30" pageNumber="105">Type material.</emphasis>
<materialsCitation id="26C23CB9FFF4FFDE4570FCA89455767B" ID-GBIF-Occurrence="3427696449" accessionNumber="LGE19001" collectingDate="2014-05-02" collectorName="Tengteng Liu &amp; Wei Guan &amp; Xuemei Hu" country="China" elevation="640" latitude="19.17" location="Limushan Nature Reserves" longLatPrecision="763" longitude="109.74" pageId="30" pageNumber="105" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="49118846FFF4FFDE4570FCA89182765E" box="[335,439,809,835]" pageId="30" pageNumber="105" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFF4FFDE45FFFCAB91E27659" box="[448,471,810,836]" pageId="30" pageNumber="105" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFF4FFDE45DBFCA89275765E" box="[484,576,809,835]" name="China" pageId="30" pageNumber="105">CHINA</collectingCountry>
,
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,
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(
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),
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<elevation id="1D87D1D7FFF4FFDE4172FCA895A27659" box="[1357,1431,809,836]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="30" pageNumber="105" unit="m" value="640.0">640 m</elevation>
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,
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<collectingDate id="F250E9CCFFF4FFDE44A8FCCD90CC767B" box="[151,249,844,870]" pageId="30" pageNumber="105" value="2014-05-02">2.v.2014</collectingDate>
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, leg.
<collectorName id="3B5F5332FFF4FFDE450AFCCC91FB767B" box="[309,462,844,871]" pageId="30" pageNumber="105">Tengteng Liu</collectorName>
,
<collectorName id="3B5F5332FFF4FFDE45E7FCCD927C767B" box="[472,585,844,870]" pageId="30" pageNumber="105">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFF4FFDE4640FCCD9336767A" box="[639,771,844,871]" pageId="30" pageNumber="105">Xuemei Hu</collectorName>
, genitalia slide no. LGE19001.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDE44F8FCEF94BC76B6" pageId="30" pageNumber="105" type="diagnosis">
<paragraph id="961536E4FFF4FFDE44F8FCEF94BC76B6" blockId="30.[151,1437,809,1702]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8FCEF91777695" bold="true" box="[199,322,878,904]" pageId="30" pageNumber="105">Diagnosis.</emphasis>
The species can be distinguished from congeners by features of the male genitalia: the anellus lobe bears a robust, horn-like apical bristle, and the juxta has a digitate projection at its middle.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDE44F8FC3591A47323" pageId="30" pageNumber="105" type="description">
<paragraph id="961536E4FFF4FFDE44F8FC35949D710E" blockId="30.[151,1437,809,1702]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8FC35916C76D3" bold="true" box="[199,345,948,974]" pageId="30" pageNumber="105">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF4FFDE4592FC35923476D2" box="[429,513,948,975]" captionStart="FIGURES 62–65" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[151,1435,181,1125]" captionTargetId="figure-22@12.[151,1435,181,1125]" captionTargetPageId="12" captionText="FIGURES 62–65. Adults of Tenupalpa spp. 62. perparviloba sp. nov., male, holotype; 63. T. uncativalva sp. nov., male, holotype; 64. T. zhengi sp. nov., male, holotype; 65. T. dissimilis sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="30" pageNumber="105">Fig. 64</figureCitation>
).
<emphasis id="A4DEEAF6FFF4FFDE462CFC35926576D3" box="[531,592,948,974]" italics="true" pageId="30" pageNumber="105">Head</emphasis>
: Silvery white. Labial palpus white and pale brown mixed; palpomere III as long as II, slightly suffused with fuscous. Antenna with scape silvery white; flagellum dark fuscous except basal 1/3 of dorsal surface mixed with silvery white, male cilia twice as long as flagellomere diameter.
</paragraph>
<paragraph id="961536E4FFF4FFDE44F8FB9D92FE71F9" blockId="30.[151,1437,809,1702]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8FB9D912D712B" box="[199,280,1052,1078]" italics="true" pageId="30" pageNumber="105">Thorax</emphasis>
: Dorsum and tegula silvery white. Wingspan 7.5 mm; forewing grey except basal 1/3 light grey, anterior edge of costa black in basal half; a roundish orange patch in distal 1/4 of wing, between costa and posterior 1/3, its outer margin almost reaching apical spot; between former patch and apical spot a narrow dirty white line, extending along termen; termen slightly tinged with black; fringe scales grey. Hindwing grey except apex black and near apex dirty white; fringe scales grey. Legs dirty white and pale brown mixed; hind tibia tinged with dark grey dorsally; mid- and hind tarsi strongly mixed with dark grey.
</paragraph>
<paragraph id="961536E4FFF4FFDE44F8FB6D95A57306" blockId="30.[151,1437,809,1702]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8FB6D9104701B" box="[199,305,1260,1286]" italics="true" pageId="30" pageNumber="105">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFF4FFDE457EFB6D91A0701B" box="[321,405,1260,1286]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="30" pageNumber="105">Fig. 13</figureCitation>
): Male coremata about half the length of sternum VIII; sternum VIII mitre-shaped: slightly constricted after base, broadly convex bilaterally, then tapered to posterior apex. Male genitalia (
<figureCitation id="0E912A61FFF4FFDE40FBFA8E95207034" box="[1220,1301,1295,1321]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="30" pageNumber="105">Fig. 83</figureCitation>
) with uncus small, distal margin deeply concave at middle, a sharp projection arising from cleft; two longitudinal ridges laterally, a row of teeth along each ridge. Gnathos hook moderately long, pointed apically. Tegumen approximately 5 times as long as uncus; a set of papilliform tubercles at anterior 1/3, each tubercle bearing a long single hair (
<figureCitation id="0E912A61FFF4FFDE4150FAF6908270AE" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="30" pageNumber="105">Fig. 44</figureCitation>
). Valva straight, ventral margin broadly convex before middle, slightly broadened apically. Anellus lobe large oval, 2/5 length of valva; apical bristle strongly developed, horn-shaped, as long as anellus lobe, slightly incurved. Juxta with a finger-shaped projection at middle of inner margin. Vinculum with posterior margin convex medially. Saccus convex. Aedeagus with basal 1/5 dilated; distal 4/5 uniformly elongate, slightly sinuous, rounded apically.
</paragraph>
<paragraph id="961536E4FFF4FFDE44F8F9A591A47323" blockId="30.[151,1437,809,1702]" box="[199,401,1572,1598]" pageId="30" pageNumber="105">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDE44F8F9C79229737F" box="[199,540,1606,1634]" pageId="30" pageNumber="105" type="distribution">
<paragraph id="961536E4FFF4FFDE44F8F9C79229737F" blockId="30.[151,1437,809,1702]" box="[199,540,1606,1634]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8F9C7916C737D" bold="true" box="[199,345,1606,1632]" pageId="30" pageNumber="105">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFF4FFDE4558F9C6919F737C" box="[359,426,1607,1633]" name="China" pageId="30" pageNumber="105">China</collectingCountry>
(
<collectingRegion id="546EF806FFF4FFDE4586F9C69225737F" box="[441,528,1607,1634]" country="China" name="Hainan" pageId="30" pageNumber="105">Hainan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDE44F8F9E8910473BB" pageId="30" pageNumber="105" type="etymology">
<paragraph id="961536E4FFF4FFDE44F8F9E8910473BB" blockId="30.[151,1437,809,1702]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8F9E8917D739E" bold="true" box="[199,328,1641,1667]" pageId="30" pageNumber="105">Etymology</emphasis>
. New species is named in memory of professor Zhemin Zheng (1932–2021), a famous Chinese entomologist.
</paragraph>
</subSubSection>
</treatment>
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196
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<mods:title id="0CE36FB0D3B2877F1A7FF73C9113A6C7">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<mods:affiliation id="E6EA087920BAAA7A4DD3A1AA5875BF84">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China &amp; The Research Institute for Sciences, Jeju National University, Jeju 63243, Korea &amp; gaeunlee 486 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5774 - 5850</mods:affiliation>
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<treatment id="1E0387F2FFF4FFE044A8F97291057624" ID-DOI="http://doi.org/10.5281/zenodo.5819866" ID-GBIF-Taxon="191928532" ID-Zenodo-Dep="5819866" LSID="urn:lsid:zoobank.org:act:4F0E736C-0061-4886-8AAD-FE22F5C612A1" httpUri="http://treatment.plazi.org/id/1E0387F2FFF4FFE044A8F97291057624" lastPageId="32" lastPageNumber="107" pageId="30" pageNumber="105">
<subSubSection id="DEB0656FFFF4FFDE44A8F972925A7213" box="[151,623,1779,1806]" pageId="30" pageNumber="105" type="nomenclature">
<paragraph id="961536E4FFF4FFDE44A8F972925A7213" blockId="30.[151,924,1779,1875]" box="[151,623,1779,1806]" pageId="30" pageNumber="105">
<heading id="CD5D8188FFF4FFDE44A8F972925A7213" bold="true" box="[151,623,1779,1806]" fontSize="11" level="1" pageId="30" pageNumber="105" reason="1">
<emphasis id="A4DEEAF6FFF4FFDE44A8F972925A7213" bold="true" box="[151,623,1779,1806]" pageId="30" pageNumber="105">
<taxonomicName id="51AA4D67FFF4FFDE44A8F97292317213" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,516,1779,1806]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="30" pageNumber="105" phylum="Tracheophyta" rank="species" species="dissimilis" status="sp. nov.">
<emphasis id="A4DEEAF6FFF4FFDE44A8F97291BB7213" bold="true" box="[151,398,1779,1806]" italics="true" pageId="30" pageNumber="105">Tenupalpa dissimilis</emphasis>
Lee
<emphasis id="A4DEEAF6FFF4FFDE45F4F97591D57213" bold="true" box="[459,480,1780,1806]" italics="true" pageId="30" pageNumber="105">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFF4FFDE462DF975925A7213" box="[530,623,1780,1806]" pageId="30" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDE44A8F89693A9724E" pageId="30" pageNumber="105" type="description">
<paragraph id="961536E4FFF4FFDE44A8F896914E722C" blockId="30.[151,924,1779,1875]" box="[151,379,1814,1841]" pageId="30" pageNumber="105">
(
<figureCitation id="0E912A61FFF4FFDE44A0F89690C3722C" box="[159,246,1815,1841]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="30" pageNumber="105">Figs 14</figureCitation>
,
<figureCitation id="0E912A61FFF4FFDE453DF8979115722C" box="[258,288,1814,1841]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="30" pageNumber="105">45</figureCitation>
,
<figureCitation id="0E912A61FFF4FFDE4513F897917F722C" box="[300,330,1814,1841]" captionStart="FIGURES 62–65" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[151,1435,181,1125]" captionTargetId="figure-22@12.[151,1435,181,1125]" captionTargetPageId="12" captionText="FIGURES 62–65. Adults of Tenupalpa spp. 62. perparviloba sp. nov., male, holotype; 63. T. uncativalva sp. nov., male, holotype; 64. T. zhengi sp. nov., male, holotype; 65. T. dissimilis sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="30" pageNumber="105">65</figureCitation>
,
<figureCitation id="0E912A61FFF4FFDE4569F8969146722C" box="[342,371,1815,1841]" captionStart="FIGURES 84–86" captionStartId="18.[151,264,1943,1967]" captionTargetBox="[351,1240,197,1904]" captionTargetId="figure-22@18.[327,1260,181,1920]" captionTargetPageId="18" captionText="FIGURES 84–86. Genitalia of Tenupalpa spp. 84. T. dissimilis sp. nov., male, holotype, slide no. LGE19007; 85. T. angustella, female, slide no. LGE17065; 86. T. fuscalata sp. nov., female, paratype, slide no. LGE18832 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819830" httpUri="https://zenodo.org/record/5819830/files/figure.png" pageId="30" pageNumber="105">84</figureCitation>
)
</paragraph>
<paragraph id="961536E4FFF4FFDE44A8F8BA93A9724E" blockId="30.[151,924,1779,1875]" box="[151,924,1851,1875]" pageId="30" pageNumber="105">
<uri id="E23B3AE6FFF4FFDE44A8F8BA93A9724E" box="[151,924,1851,1875]" pageId="30" pageNumber="105">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDE44A8F80194CD72A3" pageId="30" pageNumber="105" type="materials_examined">
<paragraph id="961536E4FFF4FFDE44A8F80194CD72A3" blockId="30.[151,1436,1920,2019]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44A8F80191717287" bold="true" box="[151,324,1920,1946]" pageId="30" pageNumber="105">Type material.</emphasis>
<materialsCitation id="26C23CB9FFF4FFDE4576F80194CD72A3" ID-GBIF-Occurrence="3427696366" accessionNumber="LGE19007" collectingDate="2013-04-30" collectorName="Yinghui Sun &amp; Wei Guan &amp; Tengteng Liu" country="China" county="Ledong County" elevation="1050" latitude="18.74" location="Tianchi" longLatPrecision="765" longitude="108.84" municipality="Jianfengling" pageId="30" pageNumber="105" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="49118846FFF4FFDE4576F80191847287" box="[329,433,1920,1946]" pageId="30" pageNumber="105" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFF4FFDE4588F80091FB7286" box="[439,462,1921,1947]" pageId="30" pageNumber="105" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFF4FFDE45E6F80192007287" box="[473,565,1920,1946]" name="China" pageId="30" pageNumber="105">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFF4FFDE4601F80192CE7287" box="[574,763,1920,1946]" country="China" name="Hainan" pageId="30" pageNumber="105">Hainan Province</collectingRegion>
,
<collectingCounty id="7F744E68FFF4FFDE473AF80193817287" box="[773,948,1920,1946]" pageId="30" pageNumber="105">Ledong County</collectingCounty>
,
<collectingMunicipality id="7671AC9EFFF4FFDE4782F801947F7287" box="[957,1098,1920,1946]" pageId="30" pageNumber="105">Jianfengling</collectingMunicipality>
,
<location id="9375603FFFF4FFDE406BF801949C7287" LSID="urn:lsid:plazi:treatment:1E0387F2FFF4FFE044A8F97291057624:9375603FFFF4FFDE406BF801949C7287" box="[1108,1193,1920,1946]" country="China" county="Ledong County" latitude="18.74" longLatPrecision="765" longitude="108.84" municipality="Jianfengling" name="Tianchi" pageId="30" pageNumber="105" stateProvince="Hainan">Tianchi</location>
(
<geoCoordinate id="F39E5023FFF4FFDE4089F801952D7287" box="[1206,1304,1920,1946]" degrees="18.74" direction="north" orientation="latitude" pageId="30" pageNumber="105" precision="555" value="18.74">18.74°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF4FFDE411CF80195BA7287" box="[1315,1423,1920,1946]" degrees="108.84" direction="east" orientation="longitude" pageId="30" pageNumber="105" precision="555" value="108.84">108.84°E</geoCoordinate>
),
<quantity id="51529B01FFF4FFDE44A8F82590DB72A2" box="[151,238,1956,1983]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="105" unit="m" value="1050.0">
<elevation id="1D87D1D7FFF4FFDE44A8F82590DB72A2" box="[151,238,1956,1983]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="30" pageNumber="105" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<date id="E2141024FFF4FFDE44C5F825914672A3" box="[250,371,1956,1982]" pageId="30" pageNumber="105" value="2013-04-30">
<collectingDate id="F250E9CCFFF4FFDE44C5F825914672A3" box="[250,371,1956,1982]" pageId="30" pageNumber="105" value="2013-04-30">30.iv.2013</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF4FFDE4592F825920972A3" box="[429,572,1956,1982]" pageId="30" pageNumber="105">Yinghui Sun</collectorName>
,
<collectorName id="3B5F5332FFF4FFDE4678F825928D72A3" box="[583,696,1956,1982]" pageId="30" pageNumber="105">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFF4FFDE46D2F82493B372A3" box="[749,902,1956,1983]" pageId="30" pageNumber="105">Tengteng Liu</collectorName>
, genitalia slide no. LGE
<quantity id="51529B01FFF4FFDE40A7F82594EB72A3" box="[1176,1246,1956,1982]" metricMagnitude="4" metricUnit="m" metricValue="1.9006999999999998" pageId="30" pageNumber="105" unit="m" value="19007.0">19007</quantity>
m.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDF44F8F849939F75C9" lastPageId="31" lastPageNumber="106" pageId="30" pageNumber="105" type="diagnosis">
<paragraph id="961536E4FFF4FFDF44F8F849939F75C9" blockId="30.[151,1436,1920,2019]" lastBlockId="31.[151,1436,150,320]" lastPageId="31" lastPageNumber="106" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8F849917772FF" bold="true" box="[199,322,1992,2018]" pageId="30" pageNumber="105">Diagnosis.</emphasis>
<taxonomicName id="51AA4D67FFF4FFDE4577F849921B72FF" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[328,558,1992,2018]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="30" pageNumber="105" phylum="Tracheophyta" rank="species" species="dissimilis" status="sp. nov.">
<emphasis id="A4DEEAF6FFF4FFDE4577F849921B72FF" box="[328,558,1992,2018]" italics="true" pageId="30" pageNumber="105">Tenupalpa dissimilis</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFF4FFDE460AF84992BA72FF" bold="true" box="[565,655,1992,2018]" pageId="30" pageNumber="105">
<taxonomicNameLabel id="BFED578DFFF4FFDE460AF84992BA72FF" box="[565,655,1992,2018]" pageId="30" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to congeneric species, as described above, and it is difficult to separate based on the external features alone. However, the male genitalia are unique in the possession of asymmetrical anellus lobes which bear a robust, hook-shaped apical bristle.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF5FFE044F8FF5F91A477D0" lastPageId="32" lastPageNumber="107" pageId="31" pageNumber="106" type="description">
<paragraph id="961536E4FFF5FFDF44F8FF5F9294745D" blockId="31.[151,1436,150,320]" pageId="31" pageNumber="106">
<emphasis id="A4DEEAF6FFF5FFDF44F8FF5F916C75E5" bold="true" box="[199,345,222,248]" pageId="31" pageNumber="106">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF5FFDF458FFF5F923075E4" box="[432,517,222,249]" captionStart="FIGURES 62–65" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[151,1435,181,1125]" captionTargetId="figure-22@12.[151,1435,181,1125]" captionTargetPageId="12" captionText="FIGURES 62–65. Adults of Tenupalpa spp. 62. perparviloba sp. nov., male, holotype; 63. T. uncativalva sp. nov., male, holotype; 64. T. zhengi sp. nov., male, holotype; 65. T. dissimilis sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="31" pageNumber="106">Fig. 65</figureCitation>
).
<emphasis id="A4DEEAF6FFF5FFDF4626FF5F926375E5" box="[537,598,222,248]" italics="true" pageId="31" pageNumber="106">Head</emphasis>
: Denuded of scales. Labial palpus dirty white except palpomere III mixed with fuscous. Antenna with scape dirty white; flagellum dark fuscous except base of dorsal surface mixed with dirty white, male cilia slightly longer than diameter.
</paragraph>
<caption id="C2D5666CFFF5FFDF44A8F8CD93B87294" ID-DOI="http://doi.org/10.5281/zenodo.5819838" ID-Zenodo-Dep="5819838" httpUri="https://zenodo.org/record/5819838/files/figure.png" pageId="31" pageNumber="106" startId="31.[151,264,1868,1892]" targetBox="[151,1435,344,1845]" targetPageId="31">
<paragraph id="961536E4FFF5FFDF44A8F8CD93B87294" blockId="31.[151,1437,1868,1929]" pageId="31" pageNumber="106">
<emphasis id="A4DEEAF6FFF5FFDF44A8F8CD91607278" bold="true" box="[151,341,1868,1893]" pageId="31" pageNumber="106">FIGURES 95–97.</emphasis>
Type specimen of
<taxonomicName id="51AA4D67FFF5FFDF462AF8CC93307278" box="[533,773,1869,1893]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="31" pageNumber="106" phylum="Tracheophyta" rank="species" species="nephodesma">
<emphasis id="A4DEEAF6FFF5FFDF462AF8CC93307278" box="[533,773,1869,1893]" italics="true" pageId="31" pageNumber="106">Tenupalpa nephodesma</emphasis>
</taxonomicName>
deposited in NHMUK. 95. Holotype, female; 96. Label data; 97. Female genitalia (holotype), slide no. J.F.G.C. 8367, 97a, detail of signum.
</paragraph>
</caption>
<paragraph id="961536E4FFCAFFE044F8FF1692FE7494" blockId="32.[151,1437,151,825]" pageId="32" pageNumber="107">
<emphasis id="A4DEEAF6FFCAFFE044F8FF16912D75AC" box="[199,280,151,177]" italics="true" pageId="32" pageNumber="107">Thorax</emphasis>
: Dorsum and tegula glossy grey. Wingspan 7.0 mm; forewing grey, anterior edge of costa black in basal half; an elliptical orange patch in distal 1/3 of wing, lying between costa and posterior 1/4, its outer margin almost reaching apical spot; a narrow dirty white line between former patch and apical spot, extending along termen; below apical spot a smaller, somewhat indistinct black spot on termen, between two spots slightly tinged with orange; dorsum suffused with orange beyond fold; fringe scales grey. Hindwing grey, except black at apex and dirty white near apex; fringe scales grey; apical fringe with a black terminal band. Legs silvery white; hind tibia dark grey dorsally; mid- and hind tarsi strongly mixed with dark grey.
</paragraph>
<paragraph id="961536E4FFCAFFE044F8FE12920077B4" blockId="32.[151,1437,151,825]" pageId="32" pageNumber="107">
<emphasis id="A4DEEAF6FFCAFFE044F8FE12910474B0" box="[199,305,403,429]" italics="true" pageId="32" pageNumber="107">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFCAFFE0457EFE1291A074B0" box="[321,405,403,429]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="32" pageNumber="107">Fig. 14</figureCitation>
): Male coremata nearly as long as sternum VIII; sternum VIII rounded. Male genitalia (
<figureCitation id="0E912A61FFCAFFE04150FE12908174CC" captionStart="FIGURES 84–86" captionStartId="18.[151,264,1943,1967]" captionTargetBox="[351,1240,197,1904]" captionTargetId="figure-22@18.[327,1260,181,1920]" captionTargetPageId="18" captionText="FIGURES 84–86. Genitalia of Tenupalpa spp. 84. T. dissimilis sp. nov., male, holotype, slide no. LGE19007; 85. T. angustella, female, slide no. LGE17065; 86. T. fuscalata sp. nov., female, paratype, slide no. LGE18832 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819830" httpUri="https://zenodo.org/record/5819830/files/figure.png" pageId="32" pageNumber="107">Fig. 84</figureCitation>
) with uncus small, obcordate; distal margin deeply incised at middle, a sharp projection arising from the cleft. Gnathos hook moderately long, with basal 1/3 broadened, acute apically. Tegumen approximately 8 times as long as uncus; a set of papilliform tubercles at anterior 1/3, each tubercle bearing a long single hair (
<figureCitation id="0E912A61FFCAFFE040FDFE7E95237704" box="[1218,1302,510,537]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="32" pageNumber="107">Fig. 45</figureCitation>
). Valva narrowed from base to middle, then broadening toward rounded apex. Anellus lobes different in size: left one much larger, rounded, approximately 1/2 length of valva, right one 1/3 length of valva; apical bristle robust, S-shaped, pointed apically. Juxta simple. Vinculum broad, simple. Saccus short, blunt. Aedeagus with basal 1/2 dilated, narrowed after middle, sinuous apically.
</paragraph>
<paragraph id="961536E4FFCAFFE044F8FD3291A477D0" blockId="32.[151,1437,151,825]" box="[199,401,691,717]" pageId="32" pageNumber="107">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFCAFFE044F8FD57922977EF" box="[199,540,726,754]" pageId="32" pageNumber="107" type="distribution">
<paragraph id="961536E4FFCAFFE044F8FD57922977EF" blockId="32.[151,1437,151,825]" box="[199,540,726,754]" pageId="32" pageNumber="107">
<emphasis id="A4DEEAF6FFCAFFE044F8FD57916C77ED" bold="true" box="[199,345,726,752]" pageId="32" pageNumber="107">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFCAFFE04558FD56919F77EC" box="[359,426,727,753]" name="China" pageId="32" pageNumber="107">China</collectingCountry>
(
<collectingRegion id="546EF806FFCAFFE04586FD56922577EF" box="[441,528,727,754]" country="China" name="Hainan" pageId="32" pageNumber="107">Hainan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFCAFFE044F8FD7A91057624" pageId="32" pageNumber="107" type="etymology">
<paragraph id="961536E4FFCAFFE044F8FD7A91057624" blockId="32.[151,1437,151,825]" pageId="32" pageNumber="107">
<emphasis id="A4DEEAF6FFCAFFE044F8FD7A917D7608" bold="true" box="[199,328,763,789]" pageId="32" pageNumber="107">Etymology</emphasis>
. The species name is derived from the Latin
<taxonomicName id="51AA4D67FFCAFFE04770FD7A938C7608" authorityName="Lee &amp; Li" authorityYear="2022" box="[847,953,763,789]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="32" pageNumber="107" phylum="Tracheophyta" rank="species" species="dissimilis">
<emphasis id="A4DEEAF6FFCAFFE04770FD7A938C7608" box="[847,953,763,789]" italics="true" pageId="32" pageNumber="107">dissimilis</emphasis>
</taxonomicName>
(dissimilar), referring to the asymmetrical anellus lobes.
</paragraph>
</subSubSection>
</treatment>
</document>

176
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<mods:title id="BCD995253E39D9A9B2C6CC6093DA7F32">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<treatment id="1E0387F2FFF6FFDC44A8FD0F929173E3" ID-DOI="http://doi.org/10.5281/zenodo.5819858" ID-GBIF-Taxon="191928531" ID-Zenodo-Dep="5819858" LSID="urn:lsid:zoobank.org:act:7DF179DD-446E-47E5-B7A7-3F9AD7DB1E27" httpUri="http://treatment.plazi.org/id/1E0387F2FFF6FFDC44A8FD0F929173E3" lastPageNumber="103" pageId="28" pageNumber="103">
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<taxonomicName id="51AA4D67FFF6FFDC44A8FD0F920077B5" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,565,654,681]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="103" phylum="Tracheophyta" rank="species" species="magniguttata" status="sp. nov.">
<emphasis id="A4DEEAF6FFF6FFDC44A8FD0F918A77B4" bold="true" box="[151,447,654,681]" italics="true" pageId="28" pageNumber="103">Tenupalpa magniguttata</emphasis>
Lee
<emphasis id="A4DEEAF6FFF6FFDC45C3FD0E922477B4" bold="true" box="[508,529,655,681]" italics="true" pageId="28" pageNumber="103">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFF6FFDC467CFD0E929577B4" box="[579,672,655,681]" pageId="28" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="DEB0656FFFF6FFDC44A8FD35939A77F7" pageId="28" pageNumber="103" type="description">
<paragraph id="961536E4FFF6FFDC44A8FD35914577D0" blockId="28.[151,943,654,746]" box="[151,368,692,717]" pageId="28" pageNumber="103">
(
<figureCitation id="0E912A61FFF6FFDC44A0FD3590C777D0" box="[159,242,692,717]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="28" pageNumber="103">Figs 24</figureCitation>
,
<figureCitation id="0E912A61FFF6FFDC44C2FD34912F77D0" box="[253,282,693,717]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="28" pageNumber="103">41</figureCitation>
,
<figureCitation id="0E912A61FFF6FFDC451AFD35917777D1" box="[293,322,692,716]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="28" pageNumber="103">61</figureCitation>
,
<figureCitation id="0E912A61FFF6FFDC4572FD34915C77D0" box="[333,361,693,717]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="28" pageNumber="103">80</figureCitation>
)
</paragraph>
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<uri id="E23B3AE6FFF6FFDC44A8FD53939A77F7" box="[151,943,722,746]" pageId="28" pageNumber="103">
urn:lsid:zoobank.org:act:
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<paragraph id="961536E4FFF6FFDC44A8FC9994FD764B" blockId="28.[151,1437,792,1791]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44A8FC999170762F" bold="true" box="[151,325,792,818]" pageId="28" pageNumber="103">Type material.</emphasis>
<materialsCitation id="26C23CB9FFF6FFDC4574FC9994FD764B" ID-GBIF-Occurrence="3427696389" accessionNumber="LGE19013" collectingDate="2015-07-07" collectorName="Qingyun Wang &amp; Suran Li &amp; Mengting Chen" country="China" elevation="766" latitude="18.88" location="Shuiman Town" longLatPrecision="764" longitude="109.67" municipality="Wuzhishan City" pageId="28" pageNumber="103" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="49118846FFF6FFDC4574FC999186762F" box="[331,435,792,818]" pageId="28" pageNumber="103" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFF6FFDC4585FC9891E4762E" box="[442,465,793,819]" pageId="28" pageNumber="103" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFF6FFDC45E2FC99920C762F" box="[477,569,792,818]" name="China" pageId="28" pageNumber="103">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFF6FFDC467BFC999334762F" box="[580,769,792,818]" country="China" name="Hainan" pageId="28" pageNumber="103">Hainan Province</collectingRegion>
,
<collectingMunicipality id="7671AC9EFFF6FFDC4733FC9993F4762F" box="[780,961,792,818]" pageId="28" pageNumber="103">Wuzhishan City</collectingMunicipality>
,
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(
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<elevation id="1D87D1D7FFF6FFDC414DFC99909A764A" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" pageId="28" pageNumber="103" unit="m" value="766.0">766 m</elevation>
</quantity>
,
<date id="E2141024FFF6FFDC4484FCBD911A764A" box="[187,303,828,855]" pageId="28" pageNumber="103" value="2015-07-07">
<collectingDate id="F250E9CCFFF6FFDC4484FCBD911A764A" box="[187,303,828,855]" pageId="28" pageNumber="103" value="2015-07-07">7.vii.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF6FFDC4554FCBD9222764A" box="[363,535,828,855]" pageId="28" pageNumber="103">Qingyun Wang</collectorName>
,
<collectorName id="3B5F5332FFF6FFDC461DFCBD92B6764B" box="[546,643,828,854]" pageId="28" pageNumber="103">Suran Li</collectorName>
and
<collectorName id="3B5F5332FFF6FFDC4686FCBD935E764B" box="[697,875,828,854]" pageId="28" pageNumber="103">Mengting Chen</collectorName>
, genitalia slide no. LGE19013.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF6FFDC44F8FCE1917876DF" pageId="28" pageNumber="103" type="diagnosis">
<paragraph id="961536E4FFF6FFDC44F8FCE1917876DF" blockId="28.[151,1437,792,1791]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44F8FCE191777667" bold="true" box="[199,322,864,890]" pageId="28" pageNumber="103">Diagnosis.</emphasis>
Superficially, this species is distinguished by the narrow forewing with a distinct black spot on the termen. The male genitalia are characterized by the medially convex costa of the valva and the straight distal part of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF6FFDC44F8FC4D91A4738F" pageId="28" pageNumber="103" type="description">
<paragraph id="961536E4FFF6FFDC44F8FC4D92947133" blockId="28.[151,1437,792,1791]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44F8FC4D916C76FB" bold="true" box="[199,345,972,998]" pageId="28" pageNumber="103">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF6FFDC458FFC4D923176FA" box="[432,516,972,999]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="28" pageNumber="103">Fig. 61</figureCitation>
).
<emphasis id="A4DEEAF6FFF6FFDC4626FC4D926376FB" box="[537,598,972,998]" italics="true" pageId="28" pageNumber="103">Head</emphasis>
: Silvery white. Labial palpus silvery white except palpomere III dark fuscous. Antenna with scape silvery white; flagellum dark fuscous except basal 1/3 of dorsal surface mixed with silvery white, male cilia slightly longer than diameter.
</paragraph>
<paragraph id="961536E4FFF6FFDC44F8FBB9928D7053" blockId="28.[151,1437,792,1791]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44F8FBB9912D714F" box="[199,280,1080,1106]" italics="true" pageId="28" pageNumber="103">Thorax</emphasis>
: Dorsum and tegula silvery white. Wingspan 5.5 mm; forewing ground color light grey, base with glossy creamy white scales, gradually darkening toward apex, anterior edge of costa black in basal half; an elliptical orange patch in distal 1/3 of wing between costa and posterior 1/4, its outer margin almost reaching apical spot; between the patch and apical spot a narrow white line extended to tornus; apical spot large, below this a smaller black spot on termen, tinged with orange between two spots; dorsum suffused with fuscous, orange after fold; fringe scales grey, mixed with cream along distal part of dorsum. Hindwing grey except apex black; fringe scales grey and cream mixed, apical fringe with a black terminal band. Legs silvery white, hind tibia suffused with fuscous in distal half, mid- and hind tarsi strongly mixed with fuscous.
</paragraph>
<paragraph id="961536E4FFF6FFDC44F8FAD990C37373" blockId="28.[151,1437,792,1791]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44F8FAD99104706F" box="[199,305,1368,1394]" italics="true" pageId="28" pageNumber="103">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFF6FFDC457DFAD991AC706F" box="[322,409,1368,1394]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="28" pageNumber="103">Fig. 24</figureCitation>
): Male coremata as long as sternum VIII; sternum VIII sub-pentagonal, rounded laterally. Male genitalia (
<figureCitation id="0E912A61FFF6FFDC457AFAFD91A0708B" box="[325,405,1404,1430]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="28" pageNumber="103">Fig. 80</figureCitation>
) with uncus small, pentagonal, with a medial ridge. Gnathos hook short, thickened after base, gradually narrowed to acute apex. Tegumen approximately 6 times as long as uncus; a set of papilliform tubercles at middle, each bearing a long hair (
<figureCitation id="0E912A61FFF6FFDC4615FA45924870C3" box="[554,637,1476,1502]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="28" pageNumber="103">Fig. 41</figureCitation>
). Valva slender, slightly curved inward, widest at middle, costa broadly convex at middle, rounded apically. Anellus lobe oval, approximately 1/5 length of valva; apical bristle longer than anellus lobe, strongly incurved. Juxta with inner margin broadly convex, a very short projection at distal 1/3. Vinculum broad and simple. Saccus knobbed. Aedeagus with basal 1/3 dilated, distal 2/3 slender and straight, acute apically.
</paragraph>
<paragraph id="961536E4FFF6FFDC44F8F9F991A4738F" blockId="28.[151,1437,792,1791]" box="[199,401,1656,1682]" pageId="28" pageNumber="103">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF6FFDC44F8F91D922973AA" box="[199,540,1692,1719]" pageId="28" pageNumber="103" type="distribution">
<paragraph id="961536E4FFF6FFDC44F8F91D922973AA" blockId="28.[151,1437,792,1791]" box="[199,540,1692,1719]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44F8F91D916C73AB" bold="true" box="[199,345,1692,1718]" pageId="28" pageNumber="103">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFF6FFDC4558F91D919F73AB" box="[359,426,1692,1718]" name="China" pageId="28" pageNumber="103">China</collectingCountry>
(
<collectingRegion id="546EF806FFF6FFDC4586F91D922573AA" box="[441,528,1692,1719]" country="China" name="Hainan" pageId="28" pageNumber="103">Hainan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF6FFDC44F8F941929173E3" pageId="28" pageNumber="103" type="etymology">
<paragraph id="961536E4FFF6FFDC44F8F941929173E3" blockId="28.[151,1437,792,1791]" pageId="28" pageNumber="103">
<emphasis id="A4DEEAF6FFF6FFDC44F8F941917D73C7" bold="true" box="[199,328,1728,1754]" pageId="28" pageNumber="103">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFF6FFDC477DF94093AC73C7" box="[834,921,1729,1754]" italics="true" pageId="28" pageNumber="103">magnus</emphasis>
(large) and
<emphasis id="A4DEEAF6FFF6FFDC401EF940944E73C7" box="[1057,1147,1729,1754]" italics="true" pageId="28" pageNumber="103">guttatus</emphasis>
(spotted), referring to the large black spot on the termen of the forewing.
</paragraph>
</subSubSection>
</treatment>
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211
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<mods:affiliation id="37869F1AA2CC3E0A2CCEC2BA79F5C26D">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China &amp; The Research Institute for Sciences, Jeju National University, Jeju 63243, Korea &amp; gaeunlee 486 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5774 - 5850</mods:affiliation>
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<paragraph id="961536E4FFF6FFDC44A8F8CE92AD7277" blockId="28.[151,922,1871,1963]" box="[151,664,1871,1898]" pageId="28" pageNumber="103">
<heading id="CD5D8188FFF6FFDC44A8F8CE92AD7277" bold="true" box="[151,664,1871,1898]" fontSize="11" level="1" pageId="28" pageNumber="103" reason="1">
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<taxonomicName id="51AA4D67FFF6FFDC44A8F8CE92187277" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,557,1871,1898]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="28" pageNumber="103" phylum="Tracheophyta" rank="species" species="perparviloba" status="sp. nov.">
<emphasis id="A4DEEAF6FFF6FFDC44A8F8CE91827277" bold="true" box="[151,439,1871,1898]" italics="true" pageId="28" pageNumber="103">Tenupalpa perparviloba</emphasis>
Lee
<emphasis id="A4DEEAF6FFF6FFDC45CBF8D1923C7277" bold="true" box="[500,521,1872,1898]" italics="true" pageId="28" pageNumber="103">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFF6FFDC4604F8D192AD7277" box="[571,664,1872,1898]" pageId="28" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="DEB0656FFFF6FFDC44A8F8F593AF72B6" pageId="28" pageNumber="103" type="description">
<paragraph id="961536E4FFF6FFDC44A8F8F5914E7293" blockId="28.[151,922,1871,1963]" box="[151,379,1908,1935]" pageId="28" pageNumber="103">
(
<figureCitation id="0E912A61FFF6FFDC44A0F8F590C37292" box="[159,246,1908,1935]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="28" pageNumber="103">Figs 25</figureCitation>
,
<figureCitation id="0E912A61FFF6FFDC453DF8F591147293" box="[258,289,1908,1934]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="28" pageNumber="103">42</figureCitation>
,
<figureCitation id="0E912A61FFF6FFDC4513F8F5917F7293" box="[300,330,1908,1934]" captionStart="FIGURES 62–65" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[151,1435,181,1125]" captionTargetId="figure-22@12.[151,1435,181,1125]" captionTargetPageId="12" captionText="FIGURES 62–65. Adults of Tenupalpa spp. 62. perparviloba sp. nov., male, holotype; 63. T. uncativalva sp. nov., male, holotype; 64. T. zhengi sp. nov., male, holotype; 65. T. dissimilis sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="28" pageNumber="103">62</figureCitation>
,
<figureCitation id="0E912A61FFF6FFDC4569F8F591467293" box="[342,371,1908,1934]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="28" pageNumber="103">81</figureCitation>
)
</paragraph>
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<emphasis id="A4DEEAF6FFF6FFDC44A8F858917D72EE" bold="true" box="[151,328,2009,2035]" pageId="28" pageNumber="103">Type material.</emphasis>
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<typeStatus id="49118846FFF6FFDC456EF858918C72EE" box="[337,441,2009,2035]" pageId="28" pageNumber="103" type="holotype">Holotype</typeStatus>
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,
<collectingCountry id="EEBD7674FFF6FFDC45D6F858927072EE" box="[489,581,2009,2035]" name="China" pageId="28" pageNumber="103">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFF6FFDC466DF858932672EE" box="[594,787,2009,2035]" country="China" name="Hainan" pageId="28" pageNumber="103">Hainan Province</collectingRegion>
,
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(
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,
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),
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<elevation id="1D87D1D7FFF6FFDC4173F85895A272E9" box="[1356,1431,2009,2036]" metricMagnitude="2" metricUnit="m" metricValue="9.54" pageId="28" pageNumber="103" unit="m" value="954.0">954 m</elevation>
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,
<date id="E2141024FFF7FFDD44A8FF16912675AC" box="[151,275,151,177]" pageId="29" pageNumber="104" value="2017-08-09">
<collectingDate id="F250E9CCFFF7FFDD44A8FF16912675AC" box="[151,275,151,177]" pageId="29" pageNumber="104" value="2017-08-09">9.viii.2017</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF7FFDD4570FF16919275AC" box="[335,423,151,177]" pageId="29" pageNumber="104">Xia Bai</collectorName>
,
<collectorName id="3B5F5332FFF7FFDD458CFF16922075AC" box="[435,533,151,177]" pageId="29" pageNumber="104">Ping Liu</collectorName>
and
<collectorName id="3B5F5332FFF7FFDD4674FF16928675AC" box="[587,691,151,177]" pageId="29" pageNumber="104">Shuai Yu</collectorName>
, genitalia slide no. LGE18036.
</materialsCitation>
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<typeStatus id="49118846FFF7FFDD4028FF1694B375AC" box="[1047,1158,151,177]" pageId="29" pageNumber="104" type="paratype">Paratype</typeStatus>
.
<collectingRegion id="546EF806FFF7FFDD40ADFF17956A75AD" box="[1170,1375,150,176]" country="China" name="Hainan" pageId="29" pageNumber="104">Hainan Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFF7FFDD4154FF1695A275AC" box="[1387,1431,151,177]" pageId="29" pageNumber="104" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="7671AC9EFFF7FFDD44A8FF3890D975CE" box="[151,236,185,211]" pageId="29" pageNumber="104">Tianchi</collectingMunicipality>
(
<geoCoordinate id="F39E5023FFF7FFDD44C3FF38916B75CE" box="[252,350,185,211]" degrees="18.74" direction="north" orientation="latitude" pageId="29" pageNumber="104" precision="555" value="18.74">18.74°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF7FFDD4554FF3891ED75CE" box="[363,472,185,211]" degrees="108.84" direction="east" orientation="longitude" pageId="29" pageNumber="104" precision="555" value="108.84">108.84°E</geoCoordinate>
),
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,
<location id="9375603FFFF7FFDD466EFF3892EB75CE" LSID="urn:lsid:plazi:treatment:1E0387F2FFF6FFDD44A8F8CE91FE71DC:9375603FFFF7FFDD466EFF3892EB75CE" box="[593,734,185,211]" county="Ledong County" latitude="18.74" longLatPrecision="765" longitude="108.84" municipality="Tianchi" name="Jianfengling" pageId="29" pageNumber="104" stateProvince="Hainan">Jianfengling</location>
,
<collectingCounty id="7F744E68FFF7FFDD46D4FF3893A875CE" box="[747,925,185,211]" pageId="29" pageNumber="104">Ledong County</collectingCounty>
,
<date id="E2141024FFF7FFDD4797FF38941475CE" box="[936,1057,185,211]" pageId="29" pageNumber="104" value="2013-04-29">
<collectingDate id="F250E9CCFFF7FFDD4797FF38941475CE" box="[936,1057,185,211]" pageId="29" pageNumber="104" value="2013-04-29">29.iv.2013</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF7FFDD4061FF3894DB75CE" box="[1118,1262,185,211]" pageId="29" pageNumber="104">Yinghui Sun</collectorName>
,
<collectorName id="3B5F5332FFF7FFDD40C5FF38955975CE" box="[1274,1388,185,211]" pageId="29" pageNumber="104">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFF7FFDD44A8FF5C910575EB" box="[151,304,220,247]" pageId="29" pageNumber="104">Tengteng Liu</collectorName>
, genitalia slide no. LGE18010
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF7FFDD44F8FF7F9179749C" pageId="29" pageNumber="104" type="diagnosis">
<paragraph id="961536E4FFF7FFDD44F8FF7F9179749C" blockId="29.[151,1437,150,1217]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8FF7F91777405" bold="true" box="[199,322,254,280]" pageId="29" pageNumber="104">Diagnosis.</emphasis>
Superficially, this species is similar to several congeners; however, it may be distinguished by the short fuscous streak on the mid-tibia, which is usually absent in related species. The male genitalia are distinguished by the distally notched uncus, the flap-like extension of the basal half of valva, and the very small anellus lobe without bristles.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF7FFDD44F8FE0891A47144" pageId="29" pageNumber="104" type="description">
<paragraph id="961536E4FFF7FFDD44F8FE08923C74F4" blockId="29.[151,1437,150,1217]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8FE08916C74BE" bold="true" box="[199,345,393,419]" pageId="29" pageNumber="104">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF7FFDD4592FE08923474B9" box="[429,513,393,420]" captionStart="FIGURES 62–65" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[151,1435,181,1125]" captionTargetId="figure-22@12.[151,1435,181,1125]" captionTargetPageId="12" captionText="FIGURES 62–65. Adults of Tenupalpa spp. 62. perparviloba sp. nov., male, holotype; 63. T. uncativalva sp. nov., male, holotype; 64. T. zhengi sp. nov., male, holotype; 65. T. dissimilis sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="29" pageNumber="104">Fig. 62</figureCitation>
).
<emphasis id="A4DEEAF6FFF7FFDD462BFE08926474BE" box="[532,593,393,419]" italics="true" pageId="29" pageNumber="104">Head</emphasis>
: Silvery white. Labial palpus creamy white, palpomere III mixed with grey. Antenna with scape silvery white; flagellum dark fuscous except basal 1/3 of dorsal surface silvery white, male cilia as long as flagellomere diameter.
</paragraph>
<paragraph id="961536E4FFF7FFDD44F8FE70955377E3" blockId="29.[151,1437,150,1217]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8FE70912D7716" box="[199,280,497,523]" italics="true" pageId="29" pageNumber="104">Thorax</emphasis>
: Dorsum and tegula silvery white to grey. Wingspan 8.0 mm; forewing grey, lighter basally, anterior edge of costa black in basal 1/3; a fusiform orange patch in distal 1/2 to 1/3 of wing between costa and posterior 1/3, its inner margin sometimes narrowed toward base; apical spot large, dirty white between apical spot and former patch, extending downward toward tornus as a narrow line; a small, indistinct black spot near tornus below apical spot, tinged with orange between two spots; dorsum with an elongate, somewhat indistinct orange spot beyond fold; fringe scales grey, dark fuscous along termen. Hindwing grey except apex black and near apex cream; fringe scales grey, apical fringe with a black terminal band. Leg silvery white; mid-tibia with a short fuscous streak on outer surface before apex; hind tibia with distal half fuscous dorsally; mid- and hind tarsi strongly mixed with fuscous.
</paragraph>
<paragraph id="961536E4FFF7FFDD44F8FC86946C712B" blockId="29.[151,1437,150,1217]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8FC869104763C" box="[199,305,775,801]" italics="true" pageId="29" pageNumber="104">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFF7FFDD457CFC8691AE763C" box="[323,411,774,801]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="29" pageNumber="104">Fig. 25</figureCitation>
): Male coremata slightly shorter than sternum VIII; sternum VIII sub-pentagonal, lateral margin rounded before middle, then gradually narrowing to posterior apex; posterior apex shortly emarginate. Male genitalia (
<figureCitation id="0E912A61FFF7FFDD4536FCCD9168767B" box="[265,349,844,870]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="29" pageNumber="104">Fig. 81</figureCitation>
) with uncus cordate: base narrow, gradually broadening toward distal margin, middle of distal margin with a deep notch. Gnathos hook long and slender, curved at middle, acute apically. Tegumen approximately 3 times as long as uncus, a set of round tubercles at middle (
<figureCitation id="0E912A61FFF7FFDD477AFC1093AF76B6" box="[837,922,913,939]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="29" pageNumber="104">Fig. 42</figureCitation>
). Valva broad basally, gradually narrowing to about 1/3, then uniformly elongate to subapex, spatulate apically; basal half of costa with a flap-like extension with a crenate costa. Anellus lobe very small, rounded, without apical or subapical bristles. Juxta with base of inner margin slightly convex. Vinculum simple, posterior margin rounded. Saccus short, sub-triangular, not exceeding tegumen pedunculus. Aedeagus with basal 2/5 dilated, distal part slender and sinuous.
</paragraph>
<paragraph id="961536E4FFF7FFDD44F8FBBE91A47144" blockId="29.[151,1437,150,1217]" box="[199,401,1087,1113]" pageId="29" pageNumber="104">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF7FFDD44F8FBE092297161" box="[199,540,1121,1148]" pageId="29" pageNumber="104" type="distribution">
<paragraph id="961536E4FFF7FFDD44F8FBE092297161" blockId="29.[151,1437,150,1217]" box="[199,540,1121,1148]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8FBE0916C7166" bold="true" box="[199,345,1121,1147]" pageId="29" pageNumber="104">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFF7FFDD4558FBE0919F7166" box="[359,426,1121,1147]" name="China" pageId="29" pageNumber="104">China</collectingCountry>
(
<collectingRegion id="546EF806FFF7FFDD4586FBE092257161" box="[441,528,1121,1148]" country="China" name="Hainan" pageId="29" pageNumber="104">Hainan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF7FFDD44F8FB0591FE71DC" pageId="29" pageNumber="104" type="etymology">
<paragraph id="961536E4FFF7FFDD44F8FB0591FE71DC" blockId="29.[151,1437,150,1217]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8FB05917D7183" bold="true" box="[199,328,1156,1182]" pageId="29" pageNumber="104">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFF7FFDD4774FB0493887183" box="[843,957,1157,1182]" italics="true" pageId="29" pageNumber="104">perparvus</emphasis>
(very small) and
<emphasis id="A4DEEAF6FFF7FFDD40B9FB0594F67183" box="[1158,1219,1156,1182]" italics="true" pageId="29" pageNumber="104">lobus</emphasis>
(lobe), referring to the very small anellus lobe.
</paragraph>
</subSubSection>
</treatment>
</document>

181
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<mods:title id="B5BE1E28B86ED60EF43F7F5A363CF5CD">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<mods:identifier id="AC65E10452D3151B442289FB72E3F901" type="Zenodo-Dep">5819794</mods:identifier>
<mods:identifier id="68B085B5E95CBF3532C4E5C012C229BF" type="ZooBank">B4061200-FCA1-43A5-811C-0C558043BA46</mods:identifier>
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<treatment id="1E0387F2FFF7FFDE44A8FA8F9195774C" ID-DOI="http://doi.org/10.5281/zenodo.5819860" ID-GBIF-Taxon="191928529" ID-Zenodo-Dep="5819860" LSID="urn:lsid:zoobank.org:act:109CB3BA-D311-4CC8-8290-37BAB19E6496" httpUri="http://treatment.plazi.org/id/1E0387F2FFF7FFDE44A8FA8F9195774C" lastPageId="30" lastPageNumber="105" pageId="29" pageNumber="104">
<subSubSection id="DEB0656FFFF7FFDD44A8FA8F92BE7034" box="[151,651,1294,1321]" pageId="29" pageNumber="104" type="nomenclature">
<paragraph id="961536E4FFF7FFDD44A8FA8F92BE7034" blockId="29.[151,933,1294,1390]" box="[151,651,1294,1321]" pageId="29" pageNumber="104">
<heading id="CD5D8188FFF7FFDD44A8FA8F92BE7034" bold="true" box="[151,651,1294,1321]" fontSize="11" level="1" pageId="29" pageNumber="104" reason="1">
<emphasis id="A4DEEAF6FFF7FFDD44A8FA8F92BE7034" bold="true" box="[151,651,1294,1321]" pageId="29" pageNumber="104">
<taxonomicName id="51AA4D67FFF7FFDD44A8FA8F92157035" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,544,1294,1321]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="104" phylum="Tracheophyta" rank="species" species="uncativalva" status="sp. nov.">
<emphasis id="A4DEEAF6FFF7FFDD44A8FA8F919F7034" bold="true" box="[151,426,1294,1321]" italics="true" pageId="29" pageNumber="104">Tenupalpa uncativalva</emphasis>
Lee
<emphasis id="A4DEEAF6FFF7FFDD45D8FA8E91C97034" bold="true" box="[487,508,1295,1321]" italics="true" pageId="29" pageNumber="104">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFF7FFDD4612FA8E92BE7034" box="[557,651,1295,1321]" pageId="29" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF7FFDD44A8FAB093907073" pageId="29" pageNumber="104" type="description">
<paragraph id="961536E4FFF7FFDD44A8FAB0914E7056" blockId="29.[151,933,1294,1390]" box="[151,379,1329,1355]" pageId="29" pageNumber="104">
(
<figureCitation id="0E912A61FFF7FFDD44A0FAB090C37056" box="[159,246,1329,1355]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="29" pageNumber="104">Figs 12</figureCitation>
,
<figureCitation id="0E912A61FFF7FFDD453DFAB091157056" box="[258,288,1329,1355]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="29" pageNumber="104">43</figureCitation>
,
<figureCitation id="0E912A61FFF7FFDD4513FAB0917F7056" box="[300,330,1329,1355]" captionStart="FIGURES 62–65" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[151,1435,181,1125]" captionTargetId="figure-22@12.[151,1435,181,1125]" captionTargetPageId="12" captionText="FIGURES 62–65. Adults of Tenupalpa spp. 62. perparviloba sp. nov., male, holotype; 63. T. uncativalva sp. nov., male, holotype; 64. T. zhengi sp. nov., male, holotype; 65. T. dissimilis sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="29" pageNumber="104">63</figureCitation>
,
<figureCitation id="0E912A61FFF7FFDD4569FAB091467056" box="[342,371,1329,1355]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="29" pageNumber="104">82</figureCitation>
)
</paragraph>
<paragraph id="961536E4FFF7FFDD44A8FAD793907073" blockId="29.[151,933,1294,1390]" box="[151,933,1366,1390]" pageId="29" pageNumber="104">
<uri id="E23B3AE6FFF7FFDD44A8FAD793907073" box="[151,933,1366,1390]" pageId="29" pageNumber="104">
urn:lsid:zoobank.org:act:
<uuid id="E20C0C31FFF7FFDD45A0FAD793907073" box="[415,933,1366,1390]" pageId="29" pageNumber="104">109CB3BA-D311-4CC8-8290-37BAB19E6496</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF7FFDD44A8FA18934270CA" pageId="29" pageNumber="104" type="materials_examined">
<paragraph id="961536E4FFF7FFDD44A8FA18934270CA" blockId="29.[151,1437,1433,2015]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44A8FA18917070AE" bold="true" box="[151,325,1433,1459]" pageId="29" pageNumber="104">Type material.</emphasis>
<materialsCitation id="26C23CB9FFF7FFDD4573FA18934270CA" ID-GBIF-Occurrence="3427696446" accessionNumber="LGE19015" collectingDate="2018-08-05" collectorName="Mujie Qi" country="China" county="Motuo County" elevation="1016" latitude="29.33" location="Chengjiao" longLatPrecision="736" longitude="95.33" municipality="Autonomous Region" pageId="29" pageNumber="104" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="49118846FFF7FFDD4573FA18918170AE" box="[332,436,1433,1459]" pageId="29" pageNumber="104" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFF7FFDD4584FA1B91E770A9" box="[443,466,1434,1460]" pageId="29" pageNumber="104" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFF7FFDD45E1FA18920F70AE" box="[478,570,1433,1459]" name="China" pageId="29" pageNumber="104">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFF7FFDD467BFA1892A070AE" box="[580,661,1433,1459]" country="China" name="Xizang" pageId="29" pageNumber="104">Xizang</collectingRegion>
<collectingMunicipality id="7671AC9EFFF7FFDD46A5FA1B93B370AE" box="[666,902,1433,1460]" pageId="29" pageNumber="104">Autonomous Region</collectingMunicipality>
,
<collectingCounty id="7F744E68FFF7FFDD47AEFA1B940370AE" box="[913,1078,1433,1460]" pageId="29" pageNumber="104">Motuo County</collectingCounty>
,
<location id="9375603FFFF7FFDD407EFA18948170AE" LSID="urn:lsid:plazi:treatment:1E0387F2FFF7FFDE44A8FA8F9195774C:9375603FFFF7FFDD407EFA18948170AE" box="[1089,1204,1433,1459]" country="China" county="Motuo County" latitude="29.33" longLatPrecision="736" longitude="95.33" municipality="Autonomous Region" name="Chengjiao" pageId="29" pageNumber="104" stateProvince="Xizang">Chengjiao</location>
(
<geoCoordinate id="F39E5023FFF7FFDD40FCFA18951370AE" box="[1219,1318,1433,1459]" degrees="29.33" direction="north" orientation="latitude" pageId="29" pageNumber="104" precision="555" value="29.33">29.33°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFF7FFDD410EFA1895BA70A9" box="[1329,1423,1433,1460]" degrees="95.33" direction="east" orientation="longitude" pageId="29" pageNumber="104" precision="555" value="95.33">95.33°E</geoCoordinate>
),
<quantity id="51529B01FFF7FFDD44A8FA3D90DB70CA" box="[151,238,1468,1495]" metricMagnitude="3" metricUnit="m" metricValue="1.016" pageId="29" pageNumber="104" unit="m" value="1016.0">
<elevation id="1D87D1D7FFF7FFDD44A8FA3D90DB70CA" box="[151,238,1468,1495]" metricMagnitude="3" metricUnit="m" metricValue="1.016" pageId="29" pageNumber="104" unit="m" value="1016.0">1016 m</elevation>
</quantity>
,
<date id="E2141024FFF7FFDD44C5FA3D914370CA" box="[250,374,1468,1495]" pageId="29" pageNumber="104" value="2018-08-05">
<collectingDate id="F250E9CCFFF7FFDD44C5FA3D914370CA" box="[250,374,1468,1495]" pageId="29" pageNumber="104" value="2018-08-05">5.viii.2018</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFF7FFDD458DFA3D922C70CB" box="[434,537,1468,1494]" pageId="29" pageNumber="104">Mujie Qi</collectorName>
, genitalia slide no. LGE19015.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF7FFDD44F8FA5F9517737C" pageId="29" pageNumber="104" type="diagnosis">
<paragraph id="961536E4FFF7FFDD44F8FA5F9517737C" blockId="29.[151,1437,1433,2015]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8FA5F917770E5" bold="true" box="[199,322,1502,1528]" pageId="29" pageNumber="104">Diagnosis.</emphasis>
The species is distinguished from other species of
<taxonomicName id="51AA4D67FFF7FFDD47BAFA5E93CF70E4" authorityName="Lee et Li" authorityYear="2021" box="[901,1018,1503,1529]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="104" phylum="Tracheophyta" rank="genus">
<emphasis id="A4DEEAF6FFF7FFDD47BAFA5E93CF70E4" box="[901,1018,1503,1529]" italics="true" pageId="29" pageNumber="104">Tenupalpa</emphasis>
</taxonomicName>
by the orange patch on the forewing that extends to an apical spot, whereas in related species the patch is separated from the apical spot by white scales. However, we examined only a single male specimen of
<taxonomicName id="51AA4D67FFF7FFDD472DF9A493877323" authorityName="Lee et Li" authorityYear="2022" box="[786,946,1572,1598]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="29" pageNumber="104" phylum="Tracheophyta" rank="species" species="uncativalva">
<emphasis id="A4DEEAF6FFF7FFDD472DF9A493877323" box="[786,946,1572,1598]" italics="true" pageId="29" pageNumber="104">T. uncativalva</emphasis>
</taxonomicName>
; hence, further examination is needed. The male genitalia are distinguished by the sharply pointed apex of the valva, which is unusual in the genus.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF7FFDE44F8F9E891A474F4" lastPageId="30" lastPageNumber="105" pageId="29" pageNumber="104" type="description">
<paragraph id="961536E4FFF7FFDD44F8F9E895A773D4" blockId="29.[151,1437,1433,2015]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8F9E8916C739E" bold="true" box="[199,345,1641,1667]" pageId="29" pageNumber="104">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFF7FFDD458CF9E8923D7399" box="[435,520,1641,1668]" captionStart="FIGURES 62–65" captionStartId="12.[151,264,1149,1173]" captionTargetBox="[151,1435,181,1125]" captionTargetId="figure-22@12.[151,1435,181,1125]" captionTargetPageId="12" captionText="FIGURES 62–65. Adults of Tenupalpa spp. 62. perparviloba sp. nov., male, holotype; 63. T. uncativalva sp. nov., male, holotype; 64. T. zhengi sp. nov., male, holotype; 65. T. dissimilis sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819816" httpUri="https://zenodo.org/record/5819816/files/figure.png" pageId="29" pageNumber="104">Fig. 63</figureCitation>
).
<emphasis id="A4DEEAF6FFF7FFDD4621F9E8926E739E" box="[542,603,1641,1667]" italics="true" pageId="29" pageNumber="104">Head</emphasis>
: Creamy white. Labial palpus creamy white; palpomere I slightly tinged with fuscous on outer surface; palpomere III as long as II, fuscous apically. Antenna with scape creamy white; flagellum dark fuscous except basal half of dorsal surface creamy white, male cilia as long as flagellomere diameter.
</paragraph>
<paragraph id="961536E4FFF7FFDD44F8F950942772C3" blockId="29.[151,1437,1433,2015]" pageId="29" pageNumber="104">
<emphasis id="A4DEEAF6FFF7FFDD44F8F950912D73F6" box="[199,280,1745,1771]" italics="true" pageId="29" pageNumber="104">Thorax</emphasis>
: Dorsum and tegula cream. Wingspan 8.0 mm; forewing grey except basal 2/5 strongly mixed with cream; anterior edge of costa black in basal 1/3; a semicircular orange patch in distal 1/3 of wing, its anterior margin reaching costa, gradually narrowing toward dorsum posteriorly, outer margin almost reaching apical spot; dirty white between patch and apical spot, extending along termen as a narrow line; termen slightly tinged with black below apical spot, dorsum suffused with orange beyond fold; fringe scales grey, dark fuscous on termen. Hindwing grey except apex black and near apex cream; fringe scales grey, apical fringe with a black terminal band. Legs creamy white; mid-tibia slightly tinged with fuscous after middle; hind tibia fuscous along distal half of dorsal margin, apex ringed with fuscous; mid- and hind tarsi strongly mixed with fuscous.
</paragraph>
<paragraph id="961536E4FFF4FFDE44F8FF1692EC74DB" blockId="30.[151,1437,151,593]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8FF16910475AC" box="[199,305,151,177]" italics="true" pageId="30" pageNumber="105">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFF4FFDE457FFF1691A075AC" box="[320,405,151,177]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="30" pageNumber="105">Fig. 12</figureCitation>
): Male coremata shorter than sternum VIII; sternum VIII approximately 1/7 length of abdomen, lateral margin rounded, posterior apex abruptly tapered to a shape point. Male genitalia (
<figureCitation id="0E912A61FFF4FFDE4080FF38952475CE" box="[1215,1297,185,211]" captionStart="FIGURES 78–83" captionStartId="17.[151,264,1900,1924]" captionTargetBox="[272,1326,190,1861]" captionTargetId="figure-18@17.[250,1338,181,1876]" captionTargetPageId="17" captionText="FIGURES 78–83. Male genitalia of Tenupalpa spp. 78. T. acutimedia sp. nov., holotype, slide no. LGE16247; 79. T. breviaedeaga sp. nov., paratype, slide no. LGE19014; 80. T. magniguttata sp. nov., holotype, slide no. LGE19013; 81. T. perparviloba sp. nov., holotype, slide no. LGE18036; 82. T. uncativalva sp. nov., holotype, slide no. LGE19015; 83. T. zhengi sp. nov., holotype, slide no. LGE19001 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819826" httpUri="https://zenodo.org/record/5819826/files/figure.png" pageId="30" pageNumber="105">Fig. 82</figureCitation>
) with uncus rhomboid, distal margin deeply emarginate, with a sharp projection arising from cleft, exceeding uncus. Gnathos hook moderately long, strongly curved at 1/3, acute apically. Tegumen approximately 5 times as long as uncus; a set of round tubercles at middle, each bearing a long single hair (
<figureCitation id="0E912A61FFF4FFDE477BFEA093AD7426" box="[836,920,289,315]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="30" pageNumber="105">Fig. 43</figureCitation>
). Valva curved inward, with long sparse setae, gradually narrowing from base and terminating in a sharply pointed apex, near costa with a short projection at distal 1/4. Anellus lobe oval, approximately 1/3 length of valva, with a strong apical bristle longer than anellus lobe. Juxta simple, slightly convex at middle of inner margin. Vinculum simple. Saccus short and broad, triangular. Aedeagus with basal half dilated, slender and sinuous distally.
</paragraph>
<paragraph id="961536E4FFF4FFDE44F8FE4E91A474F4" blockId="30.[151,1437,151,593]" box="[199,401,463,489]" pageId="30" pageNumber="105">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDE44F8FE7092297711" box="[199,540,497,524]" pageId="30" pageNumber="105" type="distribution">
<paragraph id="961536E4FFF4FFDE44F8FE7092297711" blockId="30.[151,1437,151,593]" box="[199,540,497,524]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8FE70916C7716" bold="true" box="[199,345,497,523]" pageId="30" pageNumber="105">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFF4FFDE4558FE70919F7716" box="[359,426,497,523]" name="China" pageId="30" pageNumber="105">China</collectingCountry>
(
<collectingRegion id="546EF806FFF4FFDE4586FE7092257711" box="[441,528,497,524]" country="China" name="Xizang" pageId="30" pageNumber="105">Xizang</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFF4FFDE44F8FD959195774C" pageId="30" pageNumber="105" type="etymology">
<paragraph id="961536E4FFF4FFDE44F8FD959195774C" blockId="30.[151,1437,151,593]" pageId="30" pageNumber="105">
<emphasis id="A4DEEAF6FFF4FFDE44F8FD95917D7733" bold="true" box="[199,328,532,558]" pageId="30" pageNumber="105">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFF4FFDE4770FD9493937733" box="[847,934,533,558]" italics="true" pageId="30" pageNumber="105">uncatus</emphasis>
(hooked) and
<emphasis id="A4DEEAF6FFF4FFDE4072FD9594BF7733" box="[1101,1162,532,558]" italics="true" pageId="30" pageNumber="105">valva</emphasis>
(valva), referring to the acute and curved valva.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5D87D4B11CD69A699141BA217DCF316B">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<emphasis id="A4DEEAF6FFFCFFD644A8FC9F91877624" bold="true" box="[151,434,798,825]" italics="true" pageId="22" pageNumber="97">Tenupalpa flavitermina</emphasis>
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<bibRefCitation id="F23B4B15FFFCFFD645FDFC9F937B7625" author="Kyaw, K. M. M. &amp; Yagi, S. &amp; Oku, J. &amp; Sakamaki, Y. &amp; Hirowatari, T." box="[450,846,798,824]" pageId="22" pageNumber="97" pagination="67 - 99" refId="ref23620" refString="Kyaw, K. M. M., Yagi, S., Oku, J., Sakamaki, Y. &amp; Hirowatari, T. (2019) Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys, 897, 67 - 99." type="journal article" year="2019">Kyaw, Yagi &amp; Hirowatari, 2019</bibRefCitation>
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<figureCitation id="0E912A61FFFCFFD644A0FCC290C37640" box="[159,246,835,861]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="22" pageNumber="97">Figs 10</figureCitation>
,
<figureCitation id="0E912A61FFFCFFD6453DFCC291157640" box="[258,288,835,861]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="22" pageNumber="97">34</figureCitation>
,
<figureCitation id="0E912A61FFFCFFD64513FCC3917F7640" box="[300,330,834,861]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="22" pageNumber="97">54</figureCitation>
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<figureCitation id="0E912A61FFFCFFD64569FCC291417640" box="[342,372,835,861]" captionStart="FIGURES 72–77" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 72–77. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="22" pageNumber="97">73</figureCitation>
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<figureCitation id="0E912A61FFFCFFD645BFFCC291A87640" box="[384,413,835,861]" captionStart="FIGURES 87–90" captionStartId="19.[151,264,1943,1967]" captionTargetBox="[229,1327,198,1905]" captionTargetId="figure-18@19.[196,1391,181,1920]" captionTargetPageId="19" captionText="FIGURES 87–90. Female genitalia of Tenupalpa spp. 87. T. immaculata sp. nov., paratype, slide no. LGE18002; 88. T. isabellina sp. nov., paratype, slide no. LGE17283f; 89. T. flavitermina, slide no. LGE19021; 90. T. biformis, slide no. LLJ15251 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819832" httpUri="https://zenodo.org/record/5819832/files/figure.png" pageId="22" pageNumber="97">89</figureCitation>
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<emphasis id="A4DEEAF6FFFCFFD644A8FC0691B17682" box="[151,388,903,927]" italics="true" pageId="22" pageNumber="97">Thiotricha flavitermina</emphasis>
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<emphasis id="A4DEEAF6FFFCFFD644A8FC2791B176A3" box="[151,388,934,958]" italics="true" pageId="22" pageNumber="97">Tenupalpa flavitermina</emphasis>
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Lee
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, 2021: 371
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Material examined.
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:
</emphasis>
<specimenCount id="80ACFD6DFFFCFFD644A8FBB590F17152" box="[151,196,1076,1103]" pageId="22" pageNumber="97" type="female">1 ♀</specimenCount>
,
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(
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),
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</quantity>
,
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,
<date id="E2141024FFFCFFD64728FBB593AF7153" box="[791,922,1076,1102]" pageId="22" pageNumber="97" value="2008-07-27">
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</date>
, leg.
<collectorName id="3B5F5332FFFCFFD647E4FBB594567153" box="[987,1123,1076,1102]" pageId="22" pageNumber="97">Weichun Li</collectorName>
,
<collectorName id="3B5F5332FFFCFFD6404DFBB5953A7153" box="[1138,1295,1076,1102]" pageId="22" pageNumber="97">Yongling Sun</collectorName>
and
<collectorName id="3B5F5332FFFCFFD64174FBB590F5716F" pageId="22" pageNumber="97">Haiyan Bai</collectorName>
, genitalia slide no. LGE19021
</materialsCitation>
.
<materialsCitation id="26C23CB9FFFCFFD64608FBD99372718B" ID-GBIF-Occurrence="3427696357" accessionNumber="LGE18844" collectingDate="2007-06-23" collectingDateMax="2007-06-29" collectingDateMin="2007-06-23" collectorName="Min Wang" country="China" latitude="23.2" location="Shaoguan City" longLatPrecision="754" longitude="115.23" municipality="Nanling" pageId="22" pageNumber="97" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong">
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:
</emphasis>
<specimenCount id="80ACFD6DFFFCFFD64773FBD9934E716E" box="[844,891,1112,1139]" pageId="22" pageNumber="97" type="male">1 ♂</specimenCount>
,
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(
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,
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),
<location id="9375603FFFFCFFD640D1FBD995AD716F" LSID="urn:lsid:plazi:treatment:1E0387F2FFFCFFD644A8FC9F92C37393:9375603FFFFCFFD640D1FBD995AD716F" box="[1262,1432,1112,1138]" country="China" latitude="23.2" longLatPrecision="754" longitude="115.23" municipality="Nanling" name="Shaoguan City" pageId="22" pageNumber="97" stateProvince="Guangdong">Shaoguan City</location>
,
<date id="E2141024FFFCFFD644A8FBFD9109718B" box="[151,316,1148,1174]" pageId="22" pageNumber="97" value="2007-06-23" valueMax="2007-06-29" valueMin="2007-06-23">
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</date>
, leg.
<collectorName id="3B5F5332FFFCFFD64548FBFD91C5718A" box="[375,496,1148,1175]" pageId="22" pageNumber="97">Min Wang</collectorName>
, genitalia slide no. LGE18844
</materialsCitation>
.
<materialsCitation id="26C23CB9FFFCFFD6476EFBFD95B671A6" ID-GBIF-Occurrence="3427696308" accessionNumber="LGE18845" collectingDate="2015-05-18" collectorName="Peixin Cong &amp; Wei Guan &amp; Sha Hu" country="China" elevation="742" latitude="18.88" location="Wuzhishan Nature Reserves" longLatPrecision="764" longitude="109.65" pageId="22" pageNumber="97" specimenCount="1" specimenCount-female="1" stateProvince="Hainan">
<emphasis id="A4DEEAF6FFFCFFD6476EFBFD9416718B" bold="true" box="[849,1059,1148,1174]" pageId="22" pageNumber="97">
<collectingRegion id="546EF806FFFCFFD6476EFBFD942B718B" box="[849,1054,1148,1174]" country="China" name="Hainan" pageId="22" pageNumber="97">Hainan Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFFCFFD64015FBFD9466718A" box="[1066,1107,1148,1175]" pageId="22" pageNumber="97" type="female">1 ♀</specimenCount>
,
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(
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,
<geoCoordinate id="F39E5023FFFCFFD64532FB21914E71A6" box="[269,379,1184,1211]" degrees="109.65" direction="east" orientation="longitude" pageId="22" pageNumber="97" precision="555" value="109.65">109.65°E</geoCoordinate>
),
<quantity id="51529B01FFFCFFD645B1FB2191ED71A6" box="[398,472,1184,1211]" metricMagnitude="2" metricUnit="m" metricValue="7.42" pageId="22" pageNumber="97" unit="m" value="742.0">
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</quantity>
,
<date id="E2141024FFFCFFD645DBFB21926071A6" box="[484,597,1184,1211]" pageId="22" pageNumber="97" value="2015-05-18">
<collectingDate id="F250E9CCFFFCFFD645DBFB21926071A6" box="[484,597,1184,1211]" pageId="22" pageNumber="97" value="2015-05-18">18.v.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFFCFFD646AEFB21932A71A7" box="[657,799,1184,1210]" pageId="22" pageNumber="97">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFFCFFD64714FB2193AE71A7" box="[811,923,1184,1210]" pageId="22" pageNumber="97">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFFCFFD647EDFB21941C71A6" box="[978,1065,1184,1211]" pageId="22" pageNumber="97">Sha Hu</collectorName>
, genitalia slide no. LGE18845
</materialsCitation>
;
<materialsCitation id="26C23CB9FFFCFFD641B1FB2191D5701F" ID-GBIF-Occurrence="3427696451" accessionNumber="LGE18027" collectingDate="2015-05-25" collectorName="Peixin Cong &amp; Wei Guan &amp; Sha Hu" country="China" elevation="922" latitude="18.43" location="Diaoluoshan Nature Reserves" longLatPrecision="765" longitude="109.52" pageId="22" pageNumber="97" specimenCount="1" specimenCount-male="1" stateProvince="Hainan">
<specimenCount id="80ACFD6DFFFCFFD641B1FB21909B71C2" pageId="22" pageNumber="97" type="male">1 ♂</specimenCount>
,
<location id="9375603FFFFCFFD64485FB45923371C2" LSID="urn:lsid:plazi:treatment:1E0387F2FFFCFFD644A8FC9F92C37393:9375603FFFFCFFD64485FB45923371C2" box="[186,518,1220,1247]" country="China" latitude="18.43" longLatPrecision="765" longitude="109.52" name="Diaoluoshan Nature Reserves" pageId="22" pageNumber="97" stateProvince="Hainan">Diaoluoshan Nature Reserves</location>
(
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,
<geoCoordinate id="F39E5023FFFCFFD646BCFB4592C571C2" box="[643,752,1220,1247]" degrees="109.52" direction="east" orientation="longitude" pageId="22" pageNumber="97" precision="555" value="109.52">109.52°E</geoCoordinate>
),
<quantity id="51529B01FFFCFFD6473CFB45937971C2" box="[771,844,1220,1247]" metricMagnitude="2" metricUnit="m" metricValue="9.22" pageId="22" pageNumber="97" unit="m" value="922.0">
<elevation id="1D87D1D7FFFCFFD6473CFB45937971C2" box="[771,844,1220,1247]" metricMagnitude="2" metricUnit="m" metricValue="9.22" pageId="22" pageNumber="97" unit="m" value="922.0">922 m</elevation>
</quantity>
,
<date id="E2141024FFFCFFD64767FB4593FC71C2" box="[856,969,1220,1247]" pageId="22" pageNumber="97" value="2015-05-25">
<collectingDate id="F250E9CCFFFCFFD64767FB4593FC71C2" box="[856,969,1220,1247]" pageId="22" pageNumber="97" value="2015-05-25">25.v.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFFCFFD6403BFB4594A471C3" box="[1028,1169,1220,1246]" pageId="22" pageNumber="97">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFFCFFD640A3FB45953871C3" box="[1180,1293,1220,1246]" pageId="22" pageNumber="97">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFFCFFD6417DFB4595AD71C2" box="[1346,1432,1220,1247]" pageId="22" pageNumber="97">Sha Hu</collectorName>
, genitalia slide no. LGE18027
</materialsCitation>
;
<materialsCitation id="26C23CB9FFFCFFD645D4FB69952D703B" ID-GBIF-Occurrence="3427696344" accessionNumber="LGE18028" collectingDate="2015-05-30" collectingDateMax="2015-06-01" collectingDateMin="2015-05-30" collectorName="Peixin Cong &amp; Wei Guan &amp; Sha Hu" country="China" county="Ledong County" elevation="770" latitude="18.74" location="Jianfengling Nature Reserves" longLatPrecision="765" longitude="108.87" pageId="22" pageNumber="97" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hainan">
<specimenCount id="80ACFD6DFFFCFFD645D4FB699223701E" box="[491,534,1256,1283]" pageId="22" pageNumber="97" type="male">1 ♂</specimenCount>
,
<specimenCount id="80ACFD6DFFFCFFD6461FFB69927C701E" box="[544,585,1256,1283]" pageId="22" pageNumber="97" type="female">1 ♀</specimenCount>
,
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(
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,
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),
<quantity id="51529B01FFFCFFD640A8FB6994EA701E" box="[1175,1247,1256,1283]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="22" pageNumber="97" unit="m" value="770.0">
<elevation id="1D87D1D7FFFCFFD640A8FB6994EA701E" box="[1175,1247,1256,1283]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="22" pageNumber="97" unit="m" value="770.0">770 m</elevation>
</quantity>
,
<collectingCounty id="7F744E68FFFCFFD640D6FB6995AC701F" box="[1257,1433,1256,1282]" pageId="22" pageNumber="97">Ledong County</collectingCounty>
,
<date id="E2141024FFFCFFD644A8FA8D9171703A" box="[151,324,1292,1319]" pageId="22" pageNumber="97" value="2015-05-30" valueMax="2015-06-01" valueMin="2015-05-30">
<collectingDate id="F250E9CCFFFCFFD644A8FA8D9171703A" box="[151,324,1292,1319]" pageId="22" pageNumber="97" value="2015-05-30" valueMax="2015-06-01" valueMin="2015-05-30">30.v−1.vi.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFFCFFD645BFFA8D923B703B" box="[384,526,1292,1318]" pageId="22" pageNumber="97">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFFCFFD64626FA8D92BC703B" box="[537,649,1292,1318]" pageId="22" pageNumber="97">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFFCFFD64680FA8D9320703A" box="[703,789,1292,1319]" pageId="22" pageNumber="97">Sha Hu</collectorName>
, genitalia slide nos. LGE
<quantity id="51529B01FFFCFFD6400EFA8D94BB703B" box="[1073,1166,1292,1318]" metricMagnitude="4" metricUnit="m" metricValue="1.8009" pageId="22" pageNumber="97" unit="m" value="18009.0">
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</quantity>
, LGE18028
</materialsCitation>
f.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFCFFD644F8FAB1938370E2" pageId="22" pageNumber="97" type="diagnosis">
<paragraph id="961536E4FFFCFFD644F8FAB1938370E2" blockId="22.[151,1437,1004,1679]" pageId="22" pageNumber="97">
<emphasis id="A4DEEAF6FFFCFFD644F8FAB191777057" bold="true" box="[199,322,1328,1354]" pageId="22" pageNumber="97">Diagnosis.</emphasis>
Wingspan 8.0–8.5 mm. This species is distinguished by the presence of an orange distal patch and the absence of a black apical spot on the forewing, although the orange patch may be absent, especially in specimens from
<collectingCountry id="EEBD7674FFFCFFD644E9FAF89123708E" box="[214,278,1401,1427]" name="Japan" pageId="22" pageNumber="97">Japan</collectingCountry>
(
<bibRefCitation id="F23B4B15FFFCFFD6451AFAF992B8708F" author="Kyaw, K. M. M. &amp; Yagi, S. &amp; Oku, J. &amp; Sakamaki, Y. &amp; Hirowatari, T." box="[293,653,1400,1426]" pageId="22" pageNumber="97" pagination="67 - 99" refId="ref23620" refString="Kyaw, K. M. M., Yagi, S., Oku, J., Sakamaki, Y. &amp; Hirowatari, T. (2019) Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys, 897, 67 - 99." type="journal article" year="2019">Kyaw, Yagi &amp; Hirowatari 2019</bibRefCitation>
). Also, there is color pattern variation in the forewing, with the distal 1/3 or 2/3 broadly grey to dark fuscous. The male genitalia are most similar to those of
<taxonomicName id="51AA4D67FFFCFFD6406AFA1C94F170AB" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[1109,1220,1437,1462]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="22" pageNumber="97" phylum="Tracheophyta" rank="species" species="acutata" status="sp. nov.">
<emphasis id="A4DEEAF6FFFCFFD6406AFA1C94F170AB" box="[1109,1220,1437,1462]" italics="true" pageId="22" pageNumber="97">T. acutata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFFCFFD640F6FA1D951770AB" bold="true" box="[1225,1314,1436,1462]" pageId="22" pageNumber="97">
<taxonomicNameLabel id="BFED578DFFFCFFD640F6FA1D951770AB" box="[1225,1314,1436,1462]" pageId="22" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Compared to the latter species,
<taxonomicName id="51AA4D67FFFCFFD64542FA40921770C7" box="[381,546,1472,1498]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="22" pageNumber="97" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFFCFFD64542FA40921770C7" box="[381,546,1472,1498]" italics="true" pageId="22" pageNumber="97">T. flavitermina</emphasis>
</taxonomicName>
has the valva without an apical projection and a shorter saccus. In addition, the posterior margin of the vinculum is narrowly convex in
<taxonomicName id="51AA4D67FFFCFFD64735FA64939A70E3" box="[778,943,1508,1534]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="22" pageNumber="97" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFFCFFD64735FA64939A70E3" box="[778,943,1508,1534]" italics="true" pageId="22" pageNumber="97">T. flavitermina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFCFFD644F8F989908C735A" pageId="22" pageNumber="97" type="distribution">
<paragraph id="961536E4FFFCFFD644F8F989908C735A" blockId="22.[151,1437,1004,1679]" pageId="22" pageNumber="97">
<emphasis id="A4DEEAF6FFFCFFD644F8F989916C733F" bold="true" box="[199,345,1544,1570]" pageId="22" pageNumber="97">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFFCFFD64557F989919E733F" box="[360,427,1544,1570]" name="China" pageId="22" pageNumber="97">China</collectingCountry>
(
<collectingRegion id="546EF806FFFCFFD64585F9899233733F" box="[442,518,1544,1570]" country="China" name="Fujian" pageId="22" pageNumber="97">Fujian</collectingRegion>
,
<collectingRegion id="546EF806FFFCFFD6462DF98992AD733F" box="[530,664,1544,1570]" country="China" name="Guangdong" pageId="22" pageNumber="97">Guangdong</collectingRegion>
,
<collectingRegion id="546EF806FFFCFFD6469BF9899332733F" box="[676,775,1544,1570]" country="China" name="Guangxi" pageId="22" pageNumber="97">Guangxi</collectingRegion>
,
<collectingRegion id="546EF806FFFCFFD6472CF989935D733E" box="[787,872,1544,1571]" country="China" name="Hainan" pageId="22" pageNumber="97">Hainan</collectingRegion>
),
<collectingCountry id="EEBD7674FFFCFFD64743F988938A733E" box="[892,959,1545,1571]" name="Japan" pageId="22" pageNumber="97">Japan</collectingCountry>
. This species is newly recorded from
<collectingCountry id="EEBD7674FFFCFFD64155F9899080735A" name="China" pageId="22" pageNumber="97">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFCFFD644F8F9D192C37393" pageId="22" pageNumber="97" type="discussion">
<paragraph id="961536E4FFFCFFD644F8F9D192C37393" blockId="22.[151,1437,1004,1679]" pageId="22" pageNumber="97">
<emphasis id="A4DEEAF6FFFCFFD644F8F9D1910E7377" bold="true" box="[199,315,1616,1642]" pageId="22" pageNumber="97">Remarks.</emphasis>
The
<typeStatus id="49118846FFFCFFD6454DF9D091977376" box="[370,418,1617,1643]" pageId="22" pageNumber="97">type</typeStatus>
materials deposited in OPU of
<collectingCountry id="EEBD7674FFFCFFD646C3F9D093097376" box="[764,828,1617,1643]" name="Japan" pageId="22" pageNumber="97">Japan</collectingCountry>
were not examined in this study. A detailed description of
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Kyaw
<emphasis id="A4DEEAF6FFFCFFD6465AF9F492A37393" box="[613,662,1652,1678]" italics="true" pageId="22" pageNumber="97">et al</emphasis>
. (2019)
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<emphasis id="A4DEEAF6FFFCFFD644A8F95E91B673E7" bold="true" box="[151,387,1759,1786]" italics="true" pageId="22" pageNumber="97">Tenupalpa biformis</emphasis>
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<bibRefCitation id="F23B4B15FFFCFFD645ACF961927773E7" author="Omelko, M. M." box="[403,578,1760,1786]" pageId="22" pageNumber="97" pagination="28 - 34" refId="ref23940" refString="Omelko, M. M. (1984) Description of three new species of Gelechiidae from South Primorye. The Fauna and Ecology of Insects in the South of the Far East, 112 (215), 28 - 34. [in Russian]" type="journal article" year="1984">Omelko, 1984</bibRefCitation>
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(
<figureCitation id="0E912A61FFFCFFD644A0F88590C37203" box="[159,246,1796,1822]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="22" pageNumber="97">Figs 19</figureCitation>
,
<figureCitation id="0E912A61FFFCFFD6453DF88591157202" box="[258,288,1796,1823]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="22" pageNumber="97">35</figureCitation>
,
<figureCitation id="0E912A61FFFCFFD64513F885917F7202" box="[300,330,1796,1823]" captionStart="FIGURES 54–61" captionStartId="11.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-18@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURES 54–61. Adults of Tenupalpa spp. 54. T. flavitermina, male; 55. T. biformis, male; 56. T. venustalis, female; 57. T. acutata sp. nov., male, holotype; 58. T. crassimedia sp. nov., male, holotype; 59. T. acutimedia sp. nov., male, holotype; 60. T. breviaedeaga sp. nov., male, holotype; 61. T. magniguttata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819814" httpUri="https://zenodo.org/record/5819814/files/figure.png" pageId="22" pageNumber="97">55</figureCitation>
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<figureCitation id="0E912A61FFFCFFD64569F88591407203" box="[342,373,1796,1822]" captionStart="FIGURES 72–77" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 72–77. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="22" pageNumber="97">74</figureCitation>
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<figureCitation id="0E912A61FFFCFFD645BFF88591A87203" box="[384,413,1796,1822]" captionStart="FIGURES 87–90" captionStartId="19.[151,264,1943,1967]" captionTargetBox="[229,1327,198,1905]" captionTargetId="figure-18@19.[196,1391,181,1920]" captionTargetPageId="19" captionText="FIGURES 87–90. Female genitalia of Tenupalpa spp. 87. T. immaculata sp. nov., paratype, slide no. LGE18002; 88. T. isabellina sp. nov., paratype, slide no. LGE17283f; 89. T. flavitermina, slide no. LGE19021; 90. T. biformis, slide no. LLJ15251 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819832" httpUri="https://zenodo.org/record/5819832/files/figure.png" pageId="22" pageNumber="97">90</figureCitation>
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<emphasis id="A4DEEAF6FFFCFFD644A8F80791697283" box="[151,348,1926,1950]" italics="true" pageId="22" pageNumber="97">Thiotricha biformis</emphasis>
<bibRefCitation id="F23B4B15FFFCFFD6455CF80792707283" author="Sohn, J. - C. &amp; Regier, J. C. &amp; Mitter, C. &amp; Adamski, D. &amp; Landry, J. - F. &amp; Heikkila, M. &amp; Park, K. - T. &amp; Harrison, T. &amp; Mitter, K. &amp; Zwick, A. &amp; Kawahara, A. Y. &amp; Cho, S. &amp; Cummings, M. P. &amp; Schmitz, P." box="[355,581,1926,1950]" pageId="22" pageNumber="97" pagination="112 - 132" refId="ref24244" refString="Sohn, J. - C., Regier, J. C., Mitter, C., Adamski, D., Landry, J. - F., Heikkila, M., Park, K. - T., Harrison, T., Mitter, K., Zwick, A., Kawahara, A. Y., Cho, S., Cummings, M. P. &amp; Schmitz, P. (2016) Phylogeny and feeding trait evolution of the mega-diverse Gelechioidea (Lepidoptera: Obtectomera): new insight from 19 nuclear genes. Systematic Entomology, 41 (1), 112 - 132. https: // doi. org / 10.1111 / syen. 12143" type="journal article" year="2016">
Sohn
<emphasis id="A4DEEAF6FFFCFFD645A1F80691FF7283" box="[414,458,1926,1950]" italics="true" pageId="22" pageNumber="97">et al</emphasis>
., 2016: 116
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;
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Kyaw
<emphasis id="A4DEEAF6FFFCFFD646ADF806928A7283" box="[658,703,1926,1950]" italics="true" pageId="22" pageNumber="97">et al</emphasis>
., 2019: 70
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</paragraph>
<paragraph id="961536E4FFFCFFD644A8F825920E72A0" blockId="22.[151,1437,1864,1981]" box="[151,571,1956,1981]" pageId="22" pageNumber="97">
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<taxonomicName id="51AA4D67FFFCFFD644A8F825920272A0" ID-CoL="98YHZ" authority="Lee et al., 2021: 371" authorityName="Lee" authorityPageNumber="371" authorityYear="2021" box="[151,567,1956,1981]" class="Insecta" family="Gelechiidae" genus="Tenupalpa" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="97" phylum="Arthropoda" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFFCFFD644A8F825916972A1" box="[151,348,1956,1980]" italics="true" pageId="22" pageNumber="97">Tenupalpa biformis</emphasis>
<bibRefCitation id="F23B4B15FFFCFFD6455DF824920272A0" author="Lee, G. - E. &amp; Han, T. M. &amp; Park, H. C. &amp; Qi, M. J. &amp; Li, H. H." box="[354,567,1956,1981]" pageId="22" pageNumber="97" pagination="357 - 379" refId="ref23692" refString="Lee, G. - E., Han, T. M., Park, H. C., Qi, M. J. &amp; Li, H. H. (2021) A phylogeny of the subfamily Thiotrichinae (Lepidoptera: Gel- echiidae) with a revision of the generic classification based on molecular and morphological analyses. Systematic Entomology, 46, 357 - 379. https: // doi. org / 10.1111 / syen. 12466" type="journal article" year="2021">
Lee
<emphasis id="A4DEEAF6FFFCFFD645B0F82491F772A1" box="[399,450,1956,1980]" italics="true" pageId="22" pageNumber="97">et al.</emphasis>
, 2021: 371
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.
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</paragraph>
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<emphasis id="A4DEEAF6FFFDFFD744A8FF17914B75AD" bold="true" box="[151,382,150,176]" pageId="23" pageNumber="98">Material examined.</emphasis>
<materialsCitation id="26C23CB9FFFDFFD745BDFF1793DC75C8" ID-GBIF-Occurrence="3427696304" accessionNumber="LLJ15140" collectingDate="2014-07-06" collectorName="Aihui Yin &amp; Xuemei Hu &amp; Qingyun Wang" country="China" elevation="555" latitude="30.33" location="Mt. Tianmu" longLatPrecision="733" longitude="119.4" municipality="Lao'an" pageId="23" pageNumber="98" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang">
<emphasis id="A4DEEAF6FFFDFFD745BDFF1792E575AD" bold="true" box="[386,720,150,176]" pageId="23" pageNumber="98">
<collectingCountry id="EEBD7674FFFDFFD745BDFF1791D175AD" box="[386,484,150,176]" name="China" pageId="23" pageNumber="98">CHINA</collectingCountry>
:
<collectingRegion id="546EF806FFFDFFD745D2FF1792FE75AD" box="[493,715,150,176]" country="China" name="Zhejiang" pageId="23" pageNumber="98">Zhejiang Province</collectingRegion>
:
</emphasis>
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,
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(
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,
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),
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<elevation id="1D87D1D7FFFDFFD74078FF1794BB75AC" box="[1095,1166,150,177]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="23" pageNumber="98" unit="m" value="555.0">555 m</elevation>
</quantity>
,
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,
<date id="E2141024FFFDFFD7411BFF1795AD75AD" box="[1316,1432,150,176]" pageId="23" pageNumber="98" value="2014-07-06">
<collectingDate id="F250E9CCFFFDFFD7411BFF1795AD75AD" box="[1316,1432,150,176]" pageId="23" pageNumber="98" value="2014-07-06">6.vii.2014</collectingDate>
</date>
, leg.
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,
<collectorName id="3B5F5332FFFDFFD74500FF3B918A75C8" box="[319,447,186,213]" pageId="23" pageNumber="98">Xuemei Hu</collectorName>
and
<collectorName id="3B5F5332FFFDFFD745CBFF3B92AA75C8" box="[500,671,186,213]" pageId="23" pageNumber="98">Qingyun Wang</collectorName>
, genitalia slide no. LLJ15140
</materialsCitation>
;
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(
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,
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,
<location id="9375603FFFFDFFD744D3FF5E914675E5" LSID="urn:lsid:plazi:treatment:1E0387F2FFFCFFD744A8F95E93067705:9375603FFFFDFFD744D3FF5E914675E5" box="[236,371,222,249]" country="China" latitude="30.37" longLatPrecision="733" longitude="119.43" municipality="Sanmuping" name="Mt. Tianmu" pageId="23" pageNumber="98" stateProvince="Zhejiang">Mt. Tianmu</location>
,
<date id="E2141024FFFDFFD74541FF5F923475E4" box="[382,513,222,249]" pageId="23" pageNumber="98" value="2015-07-17">
<collectingDate id="F250E9CCFFFDFFD74541FF5F923475E4" box="[382,513,222,249]" pageId="23" pageNumber="98" value="2015-07-17">17.vii.2015</collectingDate>
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, leg.
<collectorName id="3B5F5332FFFDFFD74605FF5F929E75E5" box="[570,683,222,248]" pageId="23" pageNumber="98">Aihui Yin</collectorName>
,
<collectorName id="3B5F5332FFFDFFD74688FF5E931275E4" box="[695,807,223,249]" pageId="23" pageNumber="98">Kang Lou</collectorName>
and
<collectorName id="3B5F5332FFFDFFD74762FF5E93E475E4" box="[861,977,223,249]" pageId="23" pageNumber="98">Tao Wang</collectorName>
, genitalia slide no. LLJ15251
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFDFFD744F8FE83957A74B0" pageId="23" pageNumber="98" type="diagnosis">
<paragraph id="961536E4FFFDFFD744F8FE83957A74B0" blockId="23.[151,1437,150,537]" pageId="23" pageNumber="98">
<emphasis id="A4DEEAF6FFFDFFD744F8FE8391777401" bold="true" box="[199,322,258,284]" pageId="23" pageNumber="98">Diagnosis.</emphasis>
Adult wingspan 8.5−10.0 mm.
<taxonomicName id="51AA4D67FFFDFFD7469FFE83934C7401" box="[672,889,258,284]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="98" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFFDFFD7469FFE83934C7401" box="[672,889,258,284]" italics="true" pageId="23" pageNumber="98">Tenupalpa biformis</emphasis>
</taxonomicName>
is diagnosed by the glossy white forewing with a broad dark grey area in distal half, an orange distal patch and a black apical spot. The male genitalia are characterized by the apically pointed, large thumb-like anellus lobes and the juxta with a short-forked projection. The female genitalia are similar to
<taxonomicName id="51AA4D67FFFDFFD745A9FEEE92147495" baseAuthorityName="Omelko" baseAuthorityYear="1984" box="[406,545,366,392]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="98" phylum="Tracheophyta" rank="species" species="venustalis">
<emphasis id="A4DEEAF6FFFDFFD745A9FEEE92147495" box="[406,545,366,392]" italics="true" pageId="23" pageNumber="98">T. venustalis</emphasis>
</taxonomicName>
as seen in the short apophyses anteriores and the corpus bursae without signum, but
<taxonomicName id="51AA4D67FFFDFFD744FDFE12910F74B1" box="[194,314,402,428]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="98" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFFDFFD744FDFE12910F74B1" box="[194,314,402,428]" italics="true" pageId="23" pageNumber="98">T. biformis</emphasis>
</taxonomicName>
can be distinguished from the latter by the medially elongate anterior margin of tergum VIII.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFDFFD744F8FE3794F774CD" box="[199,1218,438,465]" pageId="23" pageNumber="98" type="distribution">
<paragraph id="961536E4FFFDFFD744F8FE3794F774CD" blockId="23.[151,1437,150,537]" box="[199,1218,438,465]" pageId="23" pageNumber="98">
<emphasis id="A4DEEAF6FFFDFFD744F8FE37916C74CD" bold="true" box="[199,345,438,464]" pageId="23" pageNumber="98">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFFDFFD74558FE37919F74CD" box="[359,426,438,464]" name="China" pageId="23" pageNumber="98">China</collectingCountry>
(
<collectingRegion id="546EF806FFFDFFD74586FE37921674CC" box="[441,547,438,465]" country="China" name="Zhejiang" pageId="23" pageNumber="98">Zhejiang</collectingRegion>
),
<collectingCountry id="EEBD7674FFFDFFD74609FE36924C74CC" box="[566,633,439,465]" name="Japan" pageId="23" pageNumber="98">Japan</collectingCountry>
,
<collectingCountry id="EEBD7674FFFDFFD746BCFE3792E474CD" box="[643,721,438,464]" name="Russia" pageId="23" pageNumber="98">Russia</collectingCountry>
. This species is newly recorded from
<collectingCountry id="EEBD7674FFFDFFD74047FE37948A74CD" box="[1144,1215,438,464]" name="China" pageId="23" pageNumber="98">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFDFFD744F8FE5B93067705" pageId="23" pageNumber="98" type="discussion">
<paragraph id="961536E4FFFDFFD744F8FE5B93067705" blockId="23.[151,1437,150,537]" pageId="23" pageNumber="98">
<emphasis id="A4DEEAF6FFFDFFD744F8FE5B910E74E9" bold="true" box="[199,315,474,500]" pageId="23" pageNumber="98">Remarks.</emphasis>
The
<typeStatus id="49118846FFFDFFD7454AFE5A919074E8" box="[373,421,475,501]" pageId="23" pageNumber="98">type</typeStatus>
materials deposited in ZIRAS of
<collectingCountry id="EEBD7674FFFDFFD74720FE5B935F74E9" box="[799,874,474,500]" name="Russia" pageId="23" pageNumber="98">Russia</collectingCountry>
were not examined in this study. A detailed re-description of
<taxonomicName id="51AA4D67FFFDFFD74521FE7E91A27705" box="[286,407,510,536]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="98" phylum="Tracheophyta" rank="species" species="biformis">
<emphasis id="A4DEEAF6FFFDFFD74521FE7E91A27705" box="[286,407,510,536]" italics="true" pageId="23" pageNumber="98">T. biformis</emphasis>
</taxonomicName>
was provided in
<bibRefCitation id="F23B4B15FFFDFFD74667FE7E931A7705" author="Kyaw, K. M. M. &amp; Yagi, S. &amp; Oku, J. &amp; Sakamaki, Y. &amp; Hirowatari, T." box="[600,815,510,537]" pageId="23" pageNumber="98" pagination="67 - 99" refId="ref23620" refString="Kyaw, K. M. M., Yagi, S., Oku, J., Sakamaki, Y. &amp; Hirowatari, T. (2019) Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys, 897, 67 - 99." type="journal article" year="2019">
Kyaw
<emphasis id="A4DEEAF6FFFDFFD7469DFE7E92E67705" box="[674,723,510,536]" italics="true" pageId="23" pageNumber="98">et al</emphasis>
. (2019)
</bibRefCitation>
.
</paragraph>
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<mods:classification id="9512FD85BB0FFE80AE0642D53B2C032D">journal article</mods:classification>
<mods:identifier id="68ABF2BE16C2987487CF6B4C5CDB9068" type="CLB-Dataset">2765</mods:identifier>
<mods:identifier id="BDBEB5D1E4B40D303DEE2082F558D8CA" type="DOI">10.11646/zootaxa.5087.1.4</mods:identifier>
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<treatment id="1E0387F2FFFFFFD544A8FF169251708B" ID-DOI="http://doi.org/10.5281/zenodo.5819840" ID-GBIF-Taxon="191928527" ID-Zenodo-Dep="5819840" LSID="urn:lsid:zoobank.org:act:5484D760-FFEF-4F2B-8553-74848EB2B31D" httpUri="http://treatment.plazi.org/id/1E0387F2FFFFFFD544A8FF169251708B" lastPageNumber="96" pageId="21" pageNumber="96">
<subSubSection id="DEB0656FFFFFFFD544A8FF16925B75AC" box="[151,622,151,178]" pageId="21" pageNumber="96" type="nomenclature">
<paragraph id="961536E4FFFFFFD544A8FF16925B75AC" blockId="21.[151,917,151,242]" box="[151,622,151,178]" pageId="21" pageNumber="96">
<heading id="CD5D8188FFFFFFD544A8FF16925B75AC" bold="true" box="[151,622,151,178]" fontSize="11" level="1" pageId="21" pageNumber="96" reason="1">
<emphasis id="A4DEEAF6FFFFFFD544A8FF16925B75AC" bold="true" box="[151,622,151,178]" pageId="21" pageNumber="96">
<taxonomicName id="51AA4D67FFFFFFD544A8FF16923675AC" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,515,151,178]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="96" phylum="Tracheophyta" rank="species" species="angulosa" status="sp. nov.">
<emphasis id="A4DEEAF6FFFFFFD544A8FF1691B875AF" bold="true" box="[151,397,151,178]" italics="true" pageId="21" pageNumber="96">Tenupalpa angulosa</emphasis>
Lee
<emphasis id="A4DEEAF6FFFFFFD545F6FF1691EB75AC" bold="true" box="[457,478,151,177]" italics="true" pageId="21" pageNumber="96">et</emphasis>
Li
</taxonomicName>
,
<taxonomicNameLabel id="BFED578DFFFFFFD5462FFF16925B75AC" box="[528,622,151,177]" pageId="21" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFFFFD544A8FF3A93A075EF" pageId="21" pageNumber="96" type="description">
<paragraph id="961536E4FFFFFFD544A8FF3A914E75C8" blockId="21.[151,917,151,242]" box="[151,379,187,214]" pageId="21" pageNumber="96">
(
<figureCitation id="0E912A61FFFFFFD544A0FF3A90C375C8" box="[159,246,187,213]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="21" pageNumber="96">Figs 18</figureCitation>
,
<figureCitation id="0E912A61FFFFFFD5453DFF3A911575C8" box="[258,288,187,213]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="21" pageNumber="96">32</figureCitation>
,
<figureCitation id="0E912A61FFFFFFD54513FF3A917F75CB" box="[300,330,187,214]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="21" pageNumber="96">52</figureCitation>
,
<figureCitation id="0E912A61FFFFFFD54569FF3A914675C8" box="[342,371,187,213]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="21" pageNumber="96">71</figureCitation>
)
</paragraph>
<paragraph id="961536E4FFFFFFD544A8FF5B93A075EF" blockId="21.[151,917,151,242]" box="[151,917,218,242]" pageId="21" pageNumber="96">
<uri id="E23B3AE6FFFFFFD544A8FF5B93A075EF" box="[151,917,218,242]" pageId="21" pageNumber="96">
urn:lsid:zoobank.org:act:
<uuid id="E20C0C31FFFFFFD545A4FF5B93A075EF" box="[411,917,218,242]" pageId="21" pageNumber="96">5484D760-FFEF-4F2B-8553-74848EB2B31D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFFFFD544A8FEA091A274D6" pageId="21" pageNumber="96" type="materials_examined">
<paragraph id="961536E4FFFFFFD544A8FEA091A274D6" blockId="21.[151,1437,288,1430]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544A8FEA091727427" bold="true" box="[151,327,288,315]" pageId="21" pageNumber="96">Type material.</emphasis>
<materialsCitation id="26C23CB9FFFFFFD5456FFEA193837442" ID-GBIF-Occurrence="3427696416" accessionNumber="LGE17270" collectingDate="2007-04-13" collectorName="Houhun Li" country="China" elevation="210" latitude="22.44" location="Kadoorie Agricultural Research Centre" longLatPrecision="756" longitude="114.13" pageId="21" pageNumber="96" specimenCount="1" specimenCount-male="1" stateProvince="Hong Kong" typeStatus="holotype">
<typeStatus id="49118846FFFFFFD5456FFEA1918D7427" box="[336,440,288,314]" pageId="21" pageNumber="96" type="holotype">Holotype</typeStatus>
<specimenCount id="80ACFD6DFFFFFFD545FDFEA091EC7426" box="[450,473,289,315]" pageId="21" pageNumber="96" type="male">♂</specimenCount>
,
<collectingCountry id="EEBD7674FFFFFFD545D8FEA092767426" box="[487,579,289,315]" name="China" pageId="21" pageNumber="96">CHINA</collectingCountry>
,
<collectingRegion id="546EF806FFFFFFD5466FFEA092ED7426" box="[592,728,289,315]" inferredBySuffix="true" pageId="21" pageNumber="96">Hong Kong</collectingRegion>
,
<location id="9375603FFFFFFFD546D9FEA194967426" LSID="urn:lsid:plazi:treatment:1E0387F2FFFFFFD544A8FF169251708B:9375603FFFFFFFD546D9FEA194967426" box="[742,1187,288,315]" country="China" latitude="22.44" longLatPrecision="756" longitude="114.13" name="Kadoorie Agricultural Research Centre" pageId="21" pageNumber="96" stateProvince="Hong Kong">Kadoorie Agricultural Research Centre</location>
(
<geoCoordinate id="F39E5023FFFFFFD5408BFEA095227426" box="[1204,1303,289,315]" degrees="22.44" direction="north" orientation="latitude" pageId="21" pageNumber="96" precision="555" value="22.44">22.44°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFFFFFD5411BFEA095BA7426" box="[1316,1423,289,315]" degrees="114.13" direction="east" orientation="longitude" pageId="21" pageNumber="96" precision="555" value="114.13">114.13°E</geoCoordinate>
),
<quantity id="51529B01FFFFFFD544A8FEC490D57442" box="[151,224,325,351]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="21" pageNumber="96" unit="m" value="210.0">
<elevation id="1D87D1D7FFFFFFD544A8FEC490D57442" box="[151,224,325,351]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="21" pageNumber="96" unit="m" value="210.0">210 m</elevation>
</quantity>
,
<date id="E2141024FFFFFFD544D3FEC591507443" box="[236,357,324,350]" pageId="21" pageNumber="96" value="2007-04-13">
<collectingDate id="F250E9CCFFFFFFD544D3FEC591507443" box="[236,357,324,350]" pageId="21" pageNumber="96" value="2007-04-13">13.iv.2007</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFFFFFD5459FFEC5922E7443" box="[416,539,324,350]" pageId="21" pageNumber="96">Houhun Li</collectorName>
<emphasis id="A4DEEAF6FFFFFFD5461DFEC492667443" box="[546,595,324,350]" italics="true" pageId="21" pageNumber="96">et al</emphasis>
., genitalia slide no. LGE17270
</materialsCitation>
.
<materialsCitation id="26C23CB9FFFFFFD547FFFEC49438749F" ID-GBIF-Occurrence="3427696353" accessionNumber="LGE18026" collectingDate="2007-04-11" collectingDateMax="2007-04-19" collectingDateMin="2007-04-11" collectorName="Houhun Li" country="China" elevation="333" latitude="22.44" location="Kadoorie Agricultural Research Centre" longLatPrecision="756" longitude="114.13" pageId="21" pageNumber="96" specimenCount="4" specimenCount-male="4" stateProvince="Hong Kong" typeStatus="paratype">
<emphasis id="A4DEEAF6FFFFFFD547FFFEC494EC7442" bold="true" box="[960,1241,325,351]" pageId="21" pageNumber="96">
<typeStatus id="49118846FFFFFFD547FFFEC4940F7442" box="[960,1082,325,351]" pageId="21" pageNumber="96" type="paratype">Paratypes</typeStatus>
.
<collectingRegion id="546EF806FFFFFFD54079FEC494E17442" box="[1094,1236,325,351]" country="China" name="Hong Kong" pageId="21" pageNumber="96">Hong Kong</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFFFFFD540DFFEC495147442" box="[1248,1313,325,351]" pageId="21" pageNumber="96" type="male">4 ♂♂</specimenCount>
, same data as holotype except
<quantity id="51529B01FFFFFFD54549FEE991CF749E" box="[374,506,360,387]" metricMagnitude="2" metricUnit="m" metricValue="3.325" metricValueMax="4.55" metricValueMin="2.1" pageId="21" pageNumber="96" unit="m" value="332.5" valueMax="455.0" valueMin="210.0">
<elevation id="1D87D1D7FFFFFFD54549FEE991CF749E" box="[374,506,360,387]" metricMagnitude="2" metricUnit="m" metricValue="3.325" metricValueMax="4.55" metricValueMin="2.1" pageId="21" pageNumber="96" unit="m" value="332.5" valueMax="455.0" valueMin="210.0">210–455 m</elevation>
</quantity>
,
<date id="E2141024FFFFFFD54637FEE9929F749F" box="[520,682,360,386]" pageId="21" pageNumber="96" value="2007-04-11" valueMax="2007-04-19" valueMin="2007-04-11">
<collectingDate id="F250E9CCFFFFFFD54637FEE9929F749F" box="[520,682,360,386]" pageId="21" pageNumber="96" value="2007-04-11" valueMax="2007-04-19" valueMin="2007-04-11">11–19.iv.2007</collectingDate>
</date>
, genitalia slide no. LGE18026
</materialsCitation>
.
<materialsCitation id="26C23CB9FFFFFFD54025FEE9910374D6" ID-GBIF-Occurrence="3427696402" collectingDate="2016-03-15" collectorName="Yinggeling &amp; Qingyun Wang &amp; Suran Li &amp; Shengnan Zhao" country="China" elevation="540" latitude="19.08" location="Hongkan" longLatPrecision="764" longitude="109.5" pageId="21" pageNumber="96" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<emphasis id="A4DEEAF6FFFFFFD54025FEE994DA749F" bold="true" box="[1050,1263,360,386]" pageId="21" pageNumber="96">
<collectingRegion id="546EF806FFFFFFD54025FEE994DF749F" box="[1050,1258,360,386]" country="China" name="Hainan" pageId="21" pageNumber="96">Hainan Province</collectingRegion>
:
</emphasis>
<specimenCount id="80ACFD6DFFFFFFD540C7FEE89512749E" box="[1272,1319,361,387]" pageId="21" pageNumber="96" type="male">1 ♂</specimenCount>
,
<location id="9375603FFFFFFFD54109FEE995A8749F" LSID="urn:lsid:plazi:treatment:1E0387F2FFFFFFD544A8FF169251708B:9375603FFFFFFFD54109FEE995A8749F" box="[1334,1437,360,386]" country="China" latitude="19.08" longLatPrecision="764" longitude="109.5" name="Hongkan" pageId="21" pageNumber="96" stateProvince="Hainan">Hongkan</location>
(
<geoCoordinate id="F39E5023FFFFFFD544A0FE0C913474BA" box="[159,257,397,423]" degrees="19.08" direction="north" orientation="latitude" pageId="21" pageNumber="96" precision="555" value="19.08">19.08°N</geoCoordinate>
,
<geoCoordinate id="F39E5023FFFFFFD54531FE0D914E74BA" box="[270,379,396,423]" degrees="109.50" direction="east" orientation="longitude" pageId="21" pageNumber="96" precision="555" value="109.5">109.50°E</geoCoordinate>
),
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<elevation id="1D87D1D7FFFFFFD545B0FE0D91EC74BA" box="[399,473,396,423]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="21" pageNumber="96" unit="m" value="540.0">540 m</elevation>
</quantity>
,
<collectorName id="3B5F5332FFFFFFD545DAFE0D925174BB" box="[485,612,396,422]" pageId="21" pageNumber="96">Yinggeling</collectorName>
,
<date id="E2141024FFFFFFD5464FFE0D92D974BA" box="[624,748,396,423]" pageId="21" pageNumber="96" value="2016-03-15">
<collectingDate id="F250E9CCFFFFFFD5464FFE0D92D974BA" box="[624,748,396,423]" pageId="21" pageNumber="96" value="2016-03-15">15.iii.2016</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFFFFFD54715FE0D93E274BA" box="[810,983,396,423]" pageId="21" pageNumber="96">Qingyun Wang</collectorName>
,
<collectorName id="3B5F5332FFFFFFD547DCFE0C947074BB" box="[995,1093,396,423]" pageId="21" pageNumber="96">Suran Li</collectorName>
and
<collectorName id="3B5F5332FFFFFFD54042FE0D951A74BB" box="[1149,1327,396,422]" pageId="21" pageNumber="96">Shengnan Zhao</collectorName>
, genitalia slide no. LGE
</materialsCitation>
<quantity id="51529B01FFFFFFD54509FE3091A674D6" box="[310,403,433,459]" metricMagnitude="4" metricUnit="m" metricValue="1.7284" pageId="21" pageNumber="96" unit="m" value="17284.0">17284m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFFFFD544F8FE559480772B" pageId="21" pageNumber="96" type="diagnosis">
<paragraph id="961536E4FFFFFFD544F8FE559480772B" blockId="21.[151,1437,288,1430]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8FE55917774F3" bold="true" box="[199,322,468,494]" pageId="21" pageNumber="96">Diagnosis.</emphasis>
The species can be recognized by the faint fuscous median fascia and fine longitudinal lines on the forewing. The apex and tornal area are dark fuscous, but not forming definite shape as a spot. The male genitalia are characterized by a strongly angled base of the valva and the junction between valva and juxta.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFFFFD544F8FDC191A47037" pageId="21" pageNumber="96" type="description">
<paragraph id="961536E4FFFFFFD544F8FDC193E677BF" blockId="21.[151,1437,288,1430]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8FDC1916C7747" bold="true" box="[199,345,576,602]" pageId="21" pageNumber="96">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFFFFFD5458FFDC192317746" box="[432,516,576,603]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="21" pageNumber="96">Fig. 52</figureCitation>
).
<emphasis id="A4DEEAF6FFFFFFD54627FDC192607747" box="[536,597,576,602]" italics="true" pageId="21" pageNumber="96">Head</emphasis>
: Silvery white. Labial palpus yellowish white; palpomere III as long as II, tinged with dark grey. Antenna with scape silvery white to yellowish white; flagellum fuscous except basal half of dorsal surface mixed with yellowish white, male cilia as long as diameter.
</paragraph>
<paragraph id="961536E4FFFFFFD544F8FD2D943776DF" blockId="21.[151,1437,288,1430]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8FD2D912D77DB" box="[199,280,684,710]" italics="true" pageId="21" pageNumber="96">Thorax</emphasis>
: Dorsum and tegula silvery white to yellowish white. Wingspan 7.5–8.5 mm; forewing ground color creamy white, yellower in distal half; anterior edge of costa black in basal 1/3; near base with a small, fuscous inverted triangular patch extended from costa to fold; a somewhat indistinct fuscous fascia at middle, interrupted by fold; two fuscous lines arising from outer margin of former fascia: upper line running toward apex along anterior 1/3 of wing, lower line running along fold and divided into three branches in distal 2/5 of wing, first two branches reaching dorsum, the other branch running toward costa and meeting former line before apex; from apex to near tornus dark fuscous; fringe scale grey. Hindwing and fringe scale grey. Legs silvery white to creamy white; mid-tibia with a black spot at distal 1/3; hind tibia black at middle and apex on outer surface.
</paragraph>
<paragraph id="961536E4FFFFFFD544F8FC4D92BC701A" blockId="21.[151,1437,288,1430]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8FC4D910476FB" box="[199,305,972,998]" italics="true" pageId="21" pageNumber="96">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFFFFFD5457CFC4D91AC76FA" box="[323,409,972,999]" captionStart="FIGURES 15–25" captionStartId="8.[151,264,1775,1799]" captionTargetBox="[170,1407,181,1737]" captionTargetId="figure-22@8.[146,1440,181,1751]" captionTargetPageId="8" captionText="FIGURES 15–25. Male sternum VIII of Tenupalpa spp. 15. T. nephodesma, slide no. ZJL05343; 16. T. latifasciata sp. nov., slide no. LGE16112; 17. T. immaculata sp. nov., slide no. LGE18025; 18. T. angulosa sp. nov., slide no. LGE17270; 19. T. biformis, slide no. LLJ15140; 20. T. acutata sp. nov., slide no. LGE17045; 21. T. crassimedia sp. nov., slide no. LGE18015; 22. T. acutimedia sp. nov., slide no. LGE16247; 23. T. breviaedeaga sp. nov., slide no. LGE19014; 24. T. magniguttata sp. nov., slide no. LGE19013; 25. T. perparviloba sp. nov., slide no. LGE18036." figureDoi="http://doi.org/10.5281/zenodo.5819802" httpUri="https://zenodo.org/record/5819802/files/figure.png" pageId="21" pageNumber="96">Fig. 18</figureCitation>
): Male coremata nearly as long as sternum VIII; sternum VIII sub-triangular or sub-pentagonal. Male genitalia (
<figureCitation id="0E912A61FFFFFFD54599FC7191C27116" box="[422,503,1008,1035]" captionStart="FIGURES 66–71" captionStartId="15.[151,264,1879,1903]" captionTargetBox="[262,1331,187,1850]" captionTargetId="figure-18@15.[238,1348,181,1855]" captionTargetPageId="15" captionText="FIGURES 66–71. Male genitalia of Tenupalpa spp. 66. T. nephodesma, slide no. ZJL05343; 67. T. latifasciata sp. nov., paratype, slide no. LGE16112; 68. T. fuscalata sp. nov., holotype, slide no. LGE18829; 69. T. immaculata sp. nov., holotype, slide no. LGE18025; 70. T. isabellina sp. nov., holotype, slide no. LGE17282; 71. T. angulosa sp. nov., holotype, slide no. LGE17270 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819820" httpUri="https://zenodo.org/record/5819820/files/figure.png" pageId="21" pageNumber="96">Fig. 71</figureCitation>
) with uncus small, pentagonal, with a medial ridge. Gnathos hook moderately long, blunt at apex. Tegumen approximately 6 times as long as uncus, medially with a bundle of lageniform tubercles: each tubercle with a single seta (
<figureCitation id="0E912A61FFFFFFD54631FBB99251714E" box="[526,612,1080,1107]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="21" pageNumber="96">Fig. 32</figureCitation>
). Valva slender, with base strongly curved inward, gradually broadening from middle to apex, rounded at apex. Anellus lobe obovoid, 1/8 length of valva, sparsely setose; apical bristle longer than anellus lobe, curved inward. Joint between valva and juxta strongly angled. Juxta broad, with inner margin setose medially. Vinculum slender and simple. Saccus widely triangular, sinuate bilaterally, rounded at apex. Aedeagus with basal 1/2 globular, distal 1/2 slightly curved downward; a slender projection arising from middle of aedeagus interiorly, slightly exceeding apex.
</paragraph>
<paragraph id="961536E4FFFFFFD544F8FA9191A47037" blockId="21.[151,1437,288,1430]" box="[199,401,1296,1322]" pageId="21" pageNumber="96">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFFFFD544F8FAB5929B7052" box="[199,686,1332,1359]" pageId="21" pageNumber="96" type="distribution">
<paragraph id="961536E4FFFFFFD544F8FAB5929B7052" blockId="21.[151,1437,288,1430]" box="[199,686,1332,1359]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8FAB5916C7053" bold="true" box="[199,345,1332,1358]" pageId="21" pageNumber="96">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFFFFFD54558FAB5919F7053" box="[359,426,1332,1358]" name="China" pageId="21" pageNumber="96">China</collectingCountry>
(
<collectingRegion id="546EF806FFFFFFD54586FAB5923B7053" box="[441,526,1332,1358]" country="China" name="Hainan" pageId="21" pageNumber="96">Hainan</collectingRegion>
,
<collectingCountry id="EEBD7674FFFFFFD54625FAB492977052" box="[538,674,1333,1359]" name="Hong Kong" pageId="21" pageNumber="96">Hong Kong</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFFFFD544F8FAD89251708B" pageId="21" pageNumber="96" type="etymology">
<paragraph id="961536E4FFFFFFD544F8FAD89251708B" blockId="21.[151,1437,288,1430]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8FAD8917D706E" bold="true" box="[199,328,1369,1395]" pageId="21" pageNumber="96">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFFFFFD5476EFAD993F6706F" box="[849,963,1368,1394]" italics="true" pageId="21" pageNumber="96">angulosus</emphasis>
(strongly angled), referring to the angled junction between the valva and the juxta.
</paragraph>
</subSubSection>
</treatment>
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188
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<mods:title id="EACD25780699E42B9E745276E444DFE0">Taxonomic study of the genus Tenupalpa Lee et Li, 2021 (Lepidoptera: Gelechiidae Thiotrichinae) from China, with descriptions of fifteen new species</mods:title>
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<taxonomicName id="51AA4D67FFFFFFD544A8FA5C91D270EA" authority="Lee et Li" authorityName="Lee et Li" authorityYear="2022" box="[151,487,1501,1528]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="96" phylum="Tracheophyta" rank="species" species="falcata" status="sp. nov.">
<emphasis id="A4DEEAF6FFFFFFD544A8FA5C914470E5" bold="true" box="[151,369,1501,1528]" italics="true" pageId="21" pageNumber="96">Tenupalpa falcata</emphasis>
Lee
<emphasis id="A4DEEAF6FFFFFFD54591FA5F91F670E5" bold="true" box="[430,451,1502,1528]" italics="true" pageId="21" pageNumber="96">et</emphasis>
Li
</taxonomicName>
,
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<subSubSection id="DEB0656FFFFFFFD544A8F98393957324" pageId="21" pageNumber="96" type="description">
<paragraph id="961536E4FFFFFFD544A8F98391587301" blockId="21.[151,928,1501,1593]" box="[151,365,1538,1565]" pageId="21" pageNumber="96">
(
<figureCitation id="0E912A61FFFFFFD544A0F98390DD7301" box="[159,232,1538,1564]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="21" pageNumber="96">Figs 9</figureCitation>
,
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,
<figureCitation id="0E912A61FFFFFFD54521F98391097300" box="[286,316,1538,1565]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="21" pageNumber="96">53</figureCitation>
,
<figureCitation id="0E912A61FFFFFFD54577F98391507301" box="[328,357,1538,1564]" captionStart="FIGURES 72–77" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 72–77. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="21" pageNumber="96">72</figureCitation>
)
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<emphasis id="A4DEEAF6FFFFFFD544A8F9E69172739C" bold="true" box="[151,327,1639,1665]" pageId="21" pageNumber="96">Type material.</emphasis>
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,
<date id="E2141024FFFFFFD54552F90A91E873BB" box="[365,477,1675,1702]" pageId="21" pageNumber="96" value="2015-05-29">
<collectingDate id="F250E9CCFFFFFFD54552F90A91E873BB" box="[365,477,1675,1702]" pageId="21" pageNumber="96" value="2015-05-29">29.v.2015</collectingDate>
</date>
, leg.
<collectorName id="3B5F5332FFFFFFD54626F90A929273B8" box="[537,679,1675,1701]" pageId="21" pageNumber="96">Peixin Cong</collectorName>
,
<collectorName id="3B5F5332FFFFFFD5468DF90A931773B8" box="[690,802,1675,1701]" pageId="21" pageNumber="96">Wei Guan</collectorName>
and
<collectorName id="3B5F5332FFFFFFD54767F90A939B73BB" box="[856,942,1675,1702]" pageId="21" pageNumber="96">Sha Hu</collectorName>
, genitalia slide no. LGE18843.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFFFFD544F8F92E94117228" pageId="21" pageNumber="96" type="diagnosis">
<paragraph id="961536E4FFFFFFD544F8F92E94117228" blockId="21.[151,1437,1639,2026]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8F92E917773D4" bold="true" box="[199,322,1711,1737]" pageId="21" pageNumber="96">Diagnosis.</emphasis>
<taxonomicName id="51AA4D67FFFFFFD54576F92E922173D4" authority="Lee &amp; Li, 2022" authorityName="Lee &amp; Li" authorityYear="2022" box="[329,532,1711,1737]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="96" phylum="Tracheophyta" rank="species" species="falcata" status="sp. nov.">
<emphasis id="A4DEEAF6FFFFFFD54576F92E922173D4" box="[329,532,1711,1737]" italics="true" pageId="21" pageNumber="96">Tenupalpa falcata</emphasis>
</taxonomicName>
<emphasis id="A4DEEAF6FFFFFFD54623F92E924373D4" bold="true" box="[540,630,1711,1737]" pageId="21" pageNumber="96">
<taxonomicNameLabel id="BFED578DFFFFFFD54623F92E924373D4" box="[540,630,1711,1737]" pageId="21" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is superficially most similar to
<taxonomicName id="51AA4D67FFFFFFD547E2F93194B773D4" baseAuthorityName="Kyaw, Yagi &amp; Hirowatari" baseAuthorityYear="2019" box="[989,1154,1711,1737]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="96" phylum="Tracheophyta" rank="species" species="flavitermina">
<emphasis id="A4DEEAF6FFFFFFD547E2F93194B773D4" box="[989,1154,1711,1737]" italics="true" pageId="21" pageNumber="96">T. flavitermina</emphasis>
</taxonomicName>
, except the orange distal patch of the forewing is somewhat broader than in the latter.
<taxonomicName id="51AA4D67FFFFFFD54701F952943D73F0" authorityName="Lee &amp; Li" authorityYear="2022" box="[830,1032,1747,1773]" class="Magnoliopsida" family="Fagaceae" genus="Tenupalpa" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="96" phylum="Tracheophyta" rank="species" species="falcata">
<emphasis id="A4DEEAF6FFFFFFD54701F952943D73F0" box="[830,1032,1747,1773]" italics="true" pageId="21" pageNumber="96">Tenupalpa falcata</emphasis>
</taxonomicName>
is easily distinguished from its congeners by the features of the male genitalia, which are characterized by a falcate anellus lobe; a broad, trapezoidal vinculum; and a sinuous distal part of the aedeagus with a serrate ventral margin.
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFFFFD644F8F8BE91A4777C" lastPageId="22" lastPageNumber="97" pageId="21" pageNumber="96" type="description">
<paragraph id="961536E4FFFFFFD544F8F8BE929972BC" blockId="21.[151,1437,1639,2026]" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8F8BE916C7244" bold="true" box="[199,345,1855,1881]" pageId="21" pageNumber="96">Description.</emphasis>
Adult (
<figureCitation id="0E912A61FFFFFFD5458BF8BE923F7247" box="[436,522,1855,1882]" captionStart="FIGURES 46–53" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-22@10.[203,1385,181,1920]" captionTargetPageId="10" captionText="FIGURES 46–53. Adults of Tenupalpa spp. 46. T. angustella, male; 47. T. nephodesma, male; 48. T. latifasciata sp. nov., male, holotype; 49. T. fuscalata sp. nov., female, paratype; 50. T. immaculata sp. nov., male, holotype; 51. T. isabellina sp. nov., male, holotype; 52. T. angulosa sp. nov., male, holotype; 53. T. falcata sp. nov., male, holotype (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819808" httpUri="https://zenodo.org/record/5819808/files/figure.png" pageId="21" pageNumber="96">Fig. 53</figureCitation>
).
<emphasis id="A4DEEAF6FFFFFFD5461EF8BE926B7244" box="[545,606,1855,1881]" italics="true" pageId="21" pageNumber="96">Head</emphasis>
: Denuded [scales lost]. Labial palpus mixed light yellow and grey, palpomere III strongly mixed with grey. Antenna with scape light brown; flagellum dark fuscous, paler basally, male cilia slightly longer than flagellomere diameter.
</paragraph>
<paragraph id="961536E4FFFFFFD644F8F82A91F075E4" blockId="21.[151,1437,1639,2026]" lastBlockId="22.[151,1437,151,717]" lastPageId="22" lastPageNumber="97" pageId="21" pageNumber="96">
<emphasis id="A4DEEAF6FFFFFFD544F8F82A912D72D8" box="[199,280,1963,1989]" italics="true" pageId="21" pageNumber="96">Thorax</emphasis>
: Dorsum and tegula mixed light yellow and grey. Wingspan 8.0 mm; forewing with basal 1/3 creamy white, anterior edge of costa black, dark grey between distal 2/3 and 1/4 of wing, distal 1/4 with a light orange patch, apex dark grey; fringe scales grey. Hindwing and fringe scales grey. Legs silvery white and light brown mixed; hind tibia broadly dark fuscous at middle, tinged with dark fuscous apically; mid- and hind-tarsi strongly mixed with dark grey on inner surface.
</paragraph>
<paragraph id="961536E4FFFCFFD644F8FE8291807720" blockId="22.[151,1437,151,717]" pageId="22" pageNumber="97">
<emphasis id="A4DEEAF6FFFCFFD644F8FE8291047400" box="[199,305,259,285]" italics="true" pageId="22" pageNumber="97">Abdomen</emphasis>
(
<figureCitation id="0E912A61FFFCFFD6457CFE8291BE7400" box="[323,395,259,285]" captionStart="FIGURES 7–14" captionStartId="7.[151,264,1892,1916]" captionTargetBox="[234,1368,181,1857]" captionTargetId="figure-18@7.[220,1368,181,1869]" captionTargetPageId="7" captionText="FIGURES 7–14. Distal segment of male abdomen in Tenupalpa spp. 7. T. fuscalata sp. nov., slide no. LGE18829; 8. T. isabellina sp. nov., slide no. LGE18835; 9. T. falcata sp. nov., slide no. LGE18843; 10. T. flavitermina, slide no. LGE18842; 11. T. venustalis, slide no. LGE18562; 12. T. uncativalva sp. nov., slide no. LGE19015; 13. T. zhengi sp. nov., slide no. LGE19001; 14. T. dissimilis sp. nov., slide no. LGE19007 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819800" httpUri="https://zenodo.org/record/5819800/files/figure.png" pageId="22" pageNumber="97">Fig. 9</figureCitation>
): Male coremata slightly shorter than sternum VIII; sternum VIII sub-pentagonal, rounded laterally. Male genitalia (
<figureCitation id="0E912A61FFFCFFD64589FEA6923F745C" box="[438,522,295,321]" captionStart="FIGURES 72–77" captionStartId="16.[151,264,1934,1958]" captionTargetBox="[236,1352,194,1895]" captionTargetId="figure-22@16.[218,1370,181,1910]" captionTargetPageId="16" captionText="FIGURES 72–77. Male genitalia of Tenupalpa spp. 72. T. falcata sp. nov., holotype, slide no. LGE18843; 73. T. flavitermina, slide no. LGE18842; 74. T. biformis, slide no. LLJ15140; 75. T. venustalis, slide no. LGE18493; 76. T. acutata sp. nov., holotype, slide no. LGE18030; 77. T. crassimedia sp. nov., paratype, slide no. LGE16256 (Scale bars = 0.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.5819822" httpUri="https://zenodo.org/record/5819822/files/figure.png" pageId="22" pageNumber="97">Fig. 72</figureCitation>
) with uncus small, sub-pentagonal, with a longitudinal ridge medially. Gnathos hook short, acute apically. Tegumen approximately 9 times as long uncus; rounded tubercles at middle, each tubercle bearing a long hair (
<figureCitation id="0E912A61FFFCFFD645FDFEEE92207494" box="[450,533,367,393]" captionStart="FIGURES 26–45" captionStartId="9.[151,264,1755,1779]" captionTargetBox="[179,1409,181,1731]" captionTargetId="figure-18@9.[179,1409,181,1731]" captionTargetPageId="9" captionText="FIGURES 26–45. Tegumen tubercles of Tenupalpa spp. 26. T. angustella, slide no. LGE18719; 27. T. nephodesma, slide no. ZJL05343; 28. T. latifasciata sp. nov., slide no. LGE16112; 29. T. fuscalata sp. nov., slide no. LGE18829; 30. T. immaculata sp. nov., slide no. LGE18025; 31. T. isabellina sp. nov., slide no. LGE17282; 32. T. angulosa sp. nov., slide no. LGE17270; 33. T. falcata sp. nov., slide no. LGE18843; 34. T. flavitermina, slide no. LGE18842; 35. T. biformis, slide no. LLJ15140; 36. T. venustalis, slide no. LGE18493; 37. T. acutata sp. nov., slide no. LGE18030; 38. T. crassimedia sp. nov., slide no. LGE18015; 39. T. acutimedia sp. nov., slide no. LGE16247; 40. T. breviaedeaga sp. nov., slide no. LGE19014; 41. T. magniguttata sp. nov., slide no. LGE19013; 42. T. perparviloba sp. nov., slide no. LGE18036; 43. T. uncativalva sp. nov., slide no. LGE19015; 44. T. zhengi sp. nov., slide no. LGE19001; 45. T. dissimilis sp. nov., slide no. LGE19007." figureDoi="http://doi.org/10.5281/zenodo.5819804" httpUri="https://zenodo.org/record/5819804/files/figure.png" pageId="22" pageNumber="97">Fig. 33</figureCitation>
). Valva with basal 2/3 slender and curved inward, spatulate distally, rounded apically. Anellus lobe approximately 1/3 length of valva, falcate, gradually narrowing to pointed apex, strongly curved inward; subapical bristle arising from middle of anellus lobe, almost reaching apex of anellus lobe. Juxta simple. Vinculum very broad, somewhat trapezoidal. Saccus broad, triangular, rounded apically, not exceeding tegumen pedunculus. Aedeagus with basal 2/5 globular, distal 3/5 slender and sinuous, shortly serrate at distal 1/3 of ventral margin, rounded apically.
</paragraph>
<paragraph id="961536E4FFFCFFD644F8FDC691A4777C" blockId="22.[151,1437,151,717]" box="[199,401,583,609]" pageId="22" pageNumber="97">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFCFFD644F8FDEB9229779B" box="[199,540,618,646]" pageId="22" pageNumber="97" type="distribution">
<paragraph id="961536E4FFFCFFD644F8FDEB9229779B" blockId="22.[151,1437,151,717]" box="[199,540,618,646]" pageId="22" pageNumber="97">
<emphasis id="A4DEEAF6FFFCFFD644F8FDEB916C7799" bold="true" box="[199,345,618,644]" pageId="22" pageNumber="97">Distribution</emphasis>
.
<collectingCountry id="EEBD7674FFFCFFD64558FDEA919F7798" box="[359,426,619,645]" name="China" pageId="22" pageNumber="97">China</collectingCountry>
(
<collectingRegion id="546EF806FFFCFFD64586FDEA9225779B" box="[441,528,619,646]" country="China" name="Hainan" pageId="22" pageNumber="97">Hainan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="DEB0656FFFFCFFD644F8FD0E91B177D0" pageId="22" pageNumber="97" type="etymology">
<paragraph id="961536E4FFFCFFD644F8FD0E91B177D0" blockId="22.[151,1437,151,717]" pageId="22" pageNumber="97">
<emphasis id="A4DEEAF6FFFCFFD644F8FD0E917D77B4" bold="true" box="[199,328,655,681]" pageId="22" pageNumber="97">Etymology</emphasis>
. The species name is derived from the Latin
<emphasis id="A4DEEAF6FFFCFFD64779FD0E93AA77B4" box="[838,927,655,681]" italics="true" pageId="22" pageNumber="97">falcatus</emphasis>
(falcate), referring to the falcate anellus lobe of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>

699
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<mods:title>Eupera troglobia sp. nov.: the first troglobitic bivalve from the Americas (Mollusca, Bivalvia, Sphaeriidae)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Simone, Luiz Ricardo L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1397-9823</mods:nameIdentifier>
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<mods:nameIdentifier type="email">lrlsimone@usp.br</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ferreira, Rodrigo Lopes</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3288-4405</mods:nameIdentifier>
<mods:affiliation>Museu de Zoologia da Universidade de Sao Paulo; Cx. Postal 42494; 04218 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.42.78074</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:5B33B18C-F5F7-47BD-AF9F-EFD9305166A7" httpUri="http://treatment.plazi.org/id/1E03E206C57257A49DEB8B8CA5F2FF9D" lastPageNumber="165" pageId="0" pageNumber="165">
<subSubSection pageId="0" pageNumber="165" type="nomenclature">
<paragraph pageId="0" pageNumber="165">
<taxonomicName LSID="http://zoobank.org/5B33B18C-F5F7-47BD-AF9F-EFD9305166A7" authority="Simone &amp; Ferreira, 2022" authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia" status="sp. nov.">Eupera troglobia</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="description">
<paragraph pageId="0" pageNumber="165">
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">Figs 1-6</figureCitation>
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">, 7-13</figureCitation>
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">, 14-21</figureCitation>
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">, 22-26</figureCitation>
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">, 27-30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="materials_examined">
<paragraph pageId="0" pageNumber="165">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">
<emphasis italics="true" pageId="0" pageNumber="165">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
MZSP 155717.
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">
<typeStatus>Paratypes</typeStatus>
</emphasis>
MZSP 155716,
<specimenCount type="generic">12 specimens</specimenCount>
, MNRJ 23647,
<specimenCount type="generic">1 specimen</specimenCount>
, USNM,
<specimenCount type="generic">1 specimen</specimenCount>
, all from type locality.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="type locality">
<paragraph pageId="0" pageNumber="165">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="165">
Brazil. Tocantins; Lagoa da
<normalizedToken originalValue="Confusão">Confusao</normalizedToken>
, Casa da Pedra cave,
<geoCoordinate degrees="10" direction="south" minutes="49" orientation="latitude" precision="1" seconds="28.4" value="-10.824555">10°49'28.4&quot;S</geoCoordinate>
,
<geoCoordinate degrees="49" direction="west" minutes="37" orientation="longitude" precision="1" seconds="16.5" value="-49.621246">49°37'16.5&quot;W</geoCoordinate>
[Ferreira col., 3.viii.2021].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="diagnosis">
<paragraph pageId="0" pageNumber="165">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="165">Adult size ~4.5 mm. Lacking pigmentation in shell and soft parts. Shell very fragile, translucent, light yellow.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="description">
<paragraph pageId="0" pageNumber="165">Description.</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Shell</emphasis>
(Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">1</figureCitation>
-
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">18</figureCitation>
). Adult shell ~4.5 mm. Equivalve; height ~80% of length; width ~60% of length. Walls thin, fragile, translucent. Anterior edge rounded, smaller than posterior edge; ventral edge rounded in medium specimens (Figs
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">7-9</figureCitation>
) to slightly ascendent in larger specimens (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">1-4</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">12</figureCitation>
); posterior edge almost straight in its middle level; dorsal edge weakly convex, almost straight. Color light yellow to light greenish yellow (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">1-6</figureCitation>
). Outer surface opaque. Sculpture of uniform concentric growth lines (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">1</figureCitation>
,
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">2</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">12</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">13</figureCitation>
); ~15 per mm; each line alternating in height along its length (Fig.
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">13</figureCitation>
), but mostly 4-5 times taller than wide; interspaces ~10 times wider than each line. Growth lines usually continuous from anterior up to posterior hinge region (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">1</figureCitation>
,
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">2</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">12</figureCitation>
). Umbo (um) slightly prosogyrate, central, protruding ~10% height beyond hinge level (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">1-4</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">8-9</figureCitation>
), occupying ~20% of dorsal edge. Hinge with small, blunt cardinal tooth (ca) in LV, ~1.5 times longer than tall, tip rounded (Figs
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">14-16</figureCitation>
), shallow correspondent socket in RV (Fig.
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">16</figureCitation>
); anterior (al) and posterior (pl) lateral teeth relatively equidistant from cardinal tooth (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">4-6</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">7-11</figureCitation>
,
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">14-18</figureCitation>
), similar to each other, in both valves; located in anterior and posterior ends of hinge edge, in blunt angle preceding anterior ad posterior slopes; each lateral tooth ~4-times longer than wide, parallel to hinge edge; anterior pair of tooth usually with anterior small beak (Figs
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">14-15</figureCitation>
: al, 17-18); both lateral teeth of LV slightly more ventral than lateral teeth of RV, encasing ventrally to them; both lateral teeth of RV with narrow socket lying dorsally for counterparts of LV (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">5</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">9</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">11</figureCitation>
,
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">18</figureCitation>
). Inner surface glossy; scar of anterior adductor muscle (aa) occupying ~5% of entire inner valve surface, ~twice taller than wide, elliptic, slightly larger than scar of posterior adductor muscle (pa) (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">3-4</figureCitation>
,
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">7-9</figureCitation>
). Pallial line continuous, simple, connecting both adductor muscle scars; relatively broad; located along ventral edge ~15% of total height distant from it. Inner surface possessing minute pits, possibly of aesthetes (Figs
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">10-11</figureCitation>
: ae).
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Main muscle system</emphasis>
(Figs
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">22</figureCitation>
,
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27</figureCitation>
). Anterior adductor muscle (aa) with elliptic transverse section, dorso-ventral height of slightly twice anterior-posterior width; located close to blunt angulation between dorsal and anterior shell edges. Posterior adductor muscle (pa) slightly smaller and located slightly more ventrally than anterior muscle. Pair of anterior pedal retractor muscles (ar) originated just dorsal to anterior adductor muscle in elliptic area equivalent to ~15% of that of anterior adductor muscle; running towards posterior relatively narrow, along ~20% of shell length, attached to adjacent visceral integument; insertion splaying in antero-dorsal foot base. No detectable protractor pedal muscle. Pair of posterior pedal retractor muscles (pr) originating similarly as anterior pedal muscle, but dorsally to posterior adductor muscle; running narrow anteriorly along ~50% of shell length, as central base to local visceral mass and attached to adjacent visceral integument; insertion splaying in postero-dorsal side of pedal base.
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Foot</emphasis>
(Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
: ft). Cylindric, ~3-times longer than wide in contracted condition; ~half projected anteriorly beyond its base. Posterior region ventrally bulged, rounded. Anterior end bluntly tapering. Byssal furrow (bf) narrow, occupying ~1/4 of middle region of ventral foot surface. Byssus (by) as single, narrow, yellow thread.
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Mantle</emphasis>
(Figs
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">22</figureCitation>
,
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">26</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27</figureCitation>
). Mantle lobe thin, translucent, thickened only in edges. Colorless. Edges of both lobes fused with each other in region of anterior adductor muscle, and in region posterior to middle level of ventral edge; fusion provided by inner fold. Mantle edge in non-fused region (Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">26</figureCitation>
) with narrow, flattened outer (of) and middle (mf) folds, no papilla or special structures detectable; inner fold (if) located in inner base of middle fold, with ~half of remaining folds height, as wide as tall. Pallial musculature (pm) thin, present in base of three folds. Fusion between posterior half of mantle edges (un) simple. Incurrent siphon (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27</figureCitation>
: is) simple, cylindric, walls weakly muscular; distal edges simple, lacking papillae; length in retracted condition ~5% of shell length, ~twice longer than wide. Excurrent siphon (ex) similar to, but ~30% smaller than incurrent siphon; preserved inverted in several specimens. Siphonal musculature immersed in local mantel edges, lacking detectable bundles, neither producing pallial sinus in shell. Gill suspensory membrane (su) connected by cilia in posterior end of gill, membrane-like separating completely incurrent from excurrent chambers (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Pallial cavity</emphasis>
(Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">22</figureCitation>
). Outer demibranchs (od) with ~1/4 of shell height in its middle region; tapering gradually towards anterior, up to middle level of inner demibranch dorsal edge; tapering subtly towards posterior; lamellae very narrow, with ventral curve covering small region of inner demibranch dorsal edge (Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">23</figureCitation>
: od); dorsal connection with visceral mass via cilia (ci). Inner demibranchs (di) wide, area ~double as outer demibranchs; anterior region slightly wider than half of shell height, gradually tapering towards posterior up to certain distance from posterior adductor muscle; transversely folded; descendent lamella (Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">23</figureCitation>
: id) simple, very narrow, free from ascendent lamella; ascendent lamella with ~70% of descendent lamella length; narrow food groove (fg) in inner demibranch ventral edge; inner demibranch connections with visceral mass and its counterpart (in posterior half - Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
) via cilia (ci). Inner demibranch serving as marsupium of ~6-8 young specimens (Fig.
<figureCitation captionStart="Figures 31–32" captionStartId="F6" captionText="Figures 31 - 32. Eupera troglobia development, paratype MZSP 153866 pregnant specimen (2 - 3) 31 whole right view, shell and part of right mantle mole removed, young specimens (yo) seen in marsupium of inner demibranch by translucency 32 young specimen extracted from marsupium, right view. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.42.78074.figures31-32" httpUri="https://binary.pensoft.net/fig/671986" pageId="0" pageNumber="165">31</figureCitation>
: yo), detailed below. Pair of palps (Figs
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
,
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">25</figureCitation>
: pp) small (~half of anterior adductor muscle area), located just posterior to anterior adductor muscle; outer hemipalps (op) ~3-times longer than wide, 8-10 strong transverse folds, from edge to edge (even protruding beyond edges); tapering distally; folds ending before mouth area (mo), keeping smooth perioral area; inner hemipalp (ip) similar to outer hemipalp, but slightly smaller, usually placed close to anterior region of visceral mass.
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Visceral mass</emphasis>
(Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
). All visceral structures white. Stomach (st) occupying most of anterior half, disposed anterior-ventrally. Digestive diverticular (dg), lying along anterior region of stomach. Gonad and genital structures occupying anterior ~half of posterior half of visceral mass, covering posterior surface of stomach. Reno-pericardial structures occupying posterior half of posterior half of visceral mass, up to posterior adductor muscle. Details below.
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Circulatory and excretory systems</emphasis>
(Figs
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27</figureCitation>
). Heart occupying anterior half of reno-pericardial area. Ventricle (ve) large, as dorsal structure, totally surrounding intestine; wall thick. Anterior and posterior aortas initially running attached to adjacent intestine. Pair of auricles (au) connected to anterior region of ventricle, each one conic, running towards ventral and lateral; connecting to central region of gills. Kidneys (ki) as posterior half of reno-pericardial volume, connected anterior and dorsal surface of posterior adductor muscle; anterior region hollow, as urinary chamber; nephropore (ur) as single, small slit located in ventro-anterior surface of supra-branchial chamber; posterior region mostly filled by white, solid renal tissue.
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Digestive system</emphasis>
(Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
). Palps (pp) and mouth (mo) (Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">25</figureCitation>
) described above (pallial cavity). Esophagus (es) simple, narrow, running along ~20% of shell length from posterior region of anterior adductor muscle towards posterior and dorsal, inserting in anterior surface of stomach between its middle and dorsal thirds. Stomach (st) large, dorsally rounded, ventrally tapering towards ventral and anterior up to anterior region of foot base. No clear separation between intestine and style sac (ss). Duct to digestive diverticula (dd) located in center of both gastric lateral walls. Stomach inner surface simple, lacking chambers and large folds; gastric shield thin, located in postero-dorsal region. Intestine (in) subtly running posteriorly and dorsally after style sac end, flanking dorsal surface of foot base; short zigzag only in its middle level; intestinal length slightly larger than shell length; in pericardial region crossing directly, gradually directing ventrally and posteriorly up to posterior side of posterior adductor muscle, initially immersed in pallial edge tissue; after short distance running on supra-anal chamber (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27</figureCitation>
). Anus (an) simple, sessile, located between posterior and ventral surface of posterior adductor muscle; large anal papilla in middle of anal dorsal edge (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27</figureCitation>
: an).
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Reproductive system</emphasis>
(Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
). Gonad white, solid, small, mostly located in lateral regions of stomach. Large hollow brood pouch as posterior 2/3 of genital structures, full of large embryos (em); brood pouch tapering towards ventral and posterior, opening in both sides in small orifice (ap) located in middle level of suprabranchial chamber.
</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" italics="true" pageId="0" pageNumber="165">Central nervous system</emphasis>
(Figs
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27-29</figureCitation>
). Pair of cerebral ganglia (ce) located in region dorsal to mouth, each one ~1/20 of anterior adductor muscle size. Each ganglion (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">28</figureCitation>
: ce) elliptical, ~twice longer than wide; cerebral commissure (cc) wide, as long as each ganglion. Pair of pedal ganglia (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">29</figureCitation>
: pg) located in middle level of pedal base; both totally fused with each other, forming spheric mass; pair of statocysts (sy) very small, located in posterior side of pedal ganglia; both pedal ganglia slightly larger than one cerebral ganglion. Pair of visceral ganglia (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">27</figureCitation>
: vg) located slightly anterior to posterior adductor muscle central side; each ganglion fusiform, ~3 times longer than wide, located very close to each other, with pedal commissure very short; each visceral ganglion slightly larger than each cerebral ganglion; posteriorly single large nerve running towards posterior, ventrally to posterior adductor muscle (nv).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="development">
<paragraph pageId="0" pageNumber="165">Development.</paragraph>
<paragraph pageId="0" pageNumber="165">
Large embryos found in gonadal brood pouch located inside visceral mass (Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
: em). Embryos coming out by genital orifice (Fig.
<figureCitation captionStart="Figures 22–26" captionStartId="F4" captionText="Figures 22 - 26. Eupera troglobia anatomical drawings 22 whole right view, right valve and part of right mantle lobe removed 23 gill, transverse section in its middle level 24 whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated 25 palp region, ventral view, hemipalps slightly deflected 26 mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165">24</figureCitation>
: ap), located in middle level of inner demibranchs. Both inner demibranchs serving as external branchial brood pouches (Fig.
<figureCitation captionStart="Figures 31–32" captionStartId="F6" captionText="Figures 31 - 32. Eupera troglobia development, paratype MZSP 153866 pregnant specimen (2 - 3) 31 whole right view, shell and part of right mantle mole removed, young specimens (yo) seen in marsupium of inner demibranch by translucency 32 young specimen extracted from marsupium, right view. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.42.78074.figures31-32" httpUri="https://binary.pensoft.net/fig/671986" pageId="0" pageNumber="165">31</figureCitation>
: yo), becoming full of young specimens (yo) in their internal area between both lamellae. Young specimens with prodissoconch of ~0.2 mm (Fig.
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">20</figureCitation>
), growing up to teleoconch becoming ~3-times larger than prodissoconch (Figs
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">19</figureCitation>
,
<figureCitation captionStart="Figures 31–32" captionStartId="F6" captionText="Figures 31 - 32. Eupera troglobia development, paratype MZSP 153866 pregnant specimen (2 - 3) 31 whole right view, shell and part of right mantle mole removed, young specimens (yo) seen in marsupium of inner demibranch by translucency 32 young specimen extracted from marsupium, right view. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.42.78074.figures31-32" httpUri="https://binary.pensoft.net/fig/671986" pageId="0" pageNumber="165">32</figureCitation>
), with ~0.7 mm. Prodissoconch almost plane, circular, smooth (Figs
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">19</figureCitation>
,
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">20</figureCitation>
); teleoconch possessing only concentric undulations and growth lines (Figs
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">19</figureCitation>
,
<figureCitation captionStart="Figures 31–32" captionStartId="F6" captionText="Figures 31 - 32. Eupera troglobia development, paratype MZSP 153866 pregnant specimen (2 - 3) 31 whole right view, shell and part of right mantle mole removed, young specimens (yo) seen in marsupium of inner demibranch by translucency 32 young specimen extracted from marsupium, right view. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.42.78074.figures31-32" httpUri="https://binary.pensoft.net/fig/671986" pageId="0" pageNumber="165">32</figureCitation>
); valves translucent (Fig.
<figureCitation captionStart="Figures 31–32" captionStartId="F6" captionText="Figures 31 - 32. Eupera troglobia development, paratype MZSP 153866 pregnant specimen (2 - 3) 31 whole right view, shell and part of right mantle mole removed, young specimens (yo) seen in marsupium of inner demibranch by translucency 32 young specimen extracted from marsupium, right view. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.42.78074.figures31-32" httpUri="https://binary.pensoft.net/fig/671986" pageId="0" pageNumber="165">32</figureCitation>
). Young intra-brood pouch specimen rounded, slightly flattened, dorsal region almost straight, umbos not protruding (Figs
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">19</figureCitation>
,
<figureCitation captionStart="Figures 31–32" captionStartId="F6" captionText="Figures 31 - 32. Eupera troglobia development, paratype MZSP 153866 pregnant specimen (2 - 3) 31 whole right view, shell and part of right mantle mole removed, young specimens (yo) seen in marsupium of inner demibranch by translucency 32 young specimen extracted from marsupium, right view. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.42.78074.figures31-32" httpUri="https://binary.pensoft.net/fig/671986" pageId="0" pageNumber="165">32</figureCitation>
); shell lacking teeth in hinge and with almost no inner muscular scars (Fig.
<figureCitation captionStart="Figures 14–21" captionStartId="F3" captionText="Figures 14 - 21. Eupera troglobia shell SEM images of paratypes 153866 14 specimen 8 (part damaged), valves opened ~ 40 °, ventral view 15 same, higher magnification 16 same, higher magnification, region of cardinal tooth 17 specimen 9 (part damaged), valves opened ~ 60 °, ventral view, mainly showing hinge of left valve 18 same, right valve 19 shell of young specimen extracted from gill's marsupium, left valve, outer left view 20 same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view 21 another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500 µm (14, 17, 18), 300 µm (15), 100 µm (19), 50 µm (16, 21), 30 µm (20)." figureDoi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165">21</figureCitation>
). Gross anatomy of these young specimens (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F5" captionText="Figures 27 - 30. Eupera troglobia anatomical drawings 27 peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ 28 transition between palps and esophagus, right view, with concern to cerebral ganglia 29 pedal ganglia, dorsal view 30 gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm." figureDoi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165">30</figureCitation>
) with very small adductor muscles (aa, pa), with anterior slightly more ventral; mantle lobes edges (mb) not fused with each other, lacking siphons; gill with only inner demibranch visible (id), relatively squared, possessing 7-8 transverse folds only; foot lacking visible byssal furrow.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="materials_examined">
<paragraph pageId="0" pageNumber="165">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="165">
<typeStatus>Types</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="description">
<paragraph pageId="0" pageNumber="165">Measurements</paragraph>
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">
(in mm).
<emphasis italics="true" pageId="0" pageNumber="165">Holotype</emphasis>
</emphasis>
MZSP 155717 (Figs
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Eupera troglobia holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column 1 outer right view 2 outer left view 3 inner left view 4 inner right view 5 inner left-slightly ventral view 6 inner right-slightly ventral view." figureDoi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165">1-6</figureCitation>
): 4.3 by 3.5; Paratypes MZSP 153866: #7 (Figs
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">7-11</figureCitation>
): 2.3 by 1.9; #8 (Fig.
<figureCitation captionStart="Figures 7–13" captionStartId="F2" captionText="Figures 7 - 13. Eupera troglobia shell SEM images of paratypes 153866 7 specimen 7, both valves connected, opened ~ 120 °, ventral view 8 7, left valve, inner left view 9 7, right valve, inner left view 10 7, left valve, detail of hinge region, inner left view 11 7, right valve, detail of hinge region, inner right view 12 specimen 8, left valve, outer left view 13 same, detail of surface on middle region of ventral edge. Scale bars: 500 µm (7, 12), 300 µm (8), 200 µm (10, 13)." figureDoi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165">12</figureCitation>
): 4.5 by 3.6.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="etymology">
<paragraph pageId="0" pageNumber="165">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="165">The specific epithet refers to the troglobitic mode of life of the animal, being an adjective in the feminine nominative singular.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="165" type="habitat">
<paragraph pageId="0" pageNumber="165">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="165">
Specimens of
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
sp. nov. were only observed in the Casa de Pedra cave, and are possibly endemic to this cave (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F7" captionText="Figure 33. Location of the Casa de Pedra cave A South America, with Brazilian states highlighted B Tocantins state with Lagoa da Confusao municipality highlighted in white lines C aerial view from the Lagoa da Confusao region, where the urban area and the karst lake are visible; red star indicating the location of the Casa de Pedra cave D Casa de Pedra cave; red star indicating the location of Eupera troglobia sp. nov." figureDoi="10.3897/subtbiol.42.78074.figure33" httpUri="https://binary.pensoft.net/fig/671987" pageId="0" pageNumber="165">33A, B</figureCitation>
). The Casa de Pedra cave comprises a cave in limestone from the Couto
<normalizedToken originalValue="Magalhães">Magalhaes</normalizedToken>
Carbonatic Formation, associated with the Neoproterozoic basement of the Baixo Araguaia Supergroup, which, in addition to the limestones, presents subordinate phyllites, slates, metargilites, metarenites and quartzites (
<bibRefCitation author="Pereira, GC" journalOrPublisher="Revista GeoNorte" pageId="0" pageNumber="165" pagination="1475 - 1483" refId="B19" refString="Pereira, GC, Morais, F, 2012. Geofisica aplicada ao estudo dos fluxos subsuperficiais no entorno da Lagoa da Confusao - TO. Revista GeoNorte 3: 1475 - 1483" title="Geofisica aplicada ao estudo dos fluxos subsuperficiais no entorno da Lagoa da Confusao - TO." volume="3" year="2012">Pereira and Morais 2012</bibRefCitation>
). The climate of the region is tropical, with two distinct periods: a dry season, between May and September, and a rainy season, between October and April, with a total annual rainfall around 1750 mm (
<bibRefCitation DOI="https://doi.org/10.1590/S0100-67622002000300005" author="Martins, ICDM" journalOrPublisher="Revista Arvore" pageId="0" pageNumber="165" pagination="299 - 309" refId="B18" refString="Martins, ICDM, Soares, VP, Silva, E, Brites, RS, 2002. Diagnostico ambiental no contexto da paisagem de fragmentos florestais naturais&quot; ipucas&quot; no municipio de Lagoa da Confusao, Tocantins. Revista Arvore 26: 299 - 309, DOI: https://doi.org/10.1590/S0100-67622002000300005" title="Diagnostico ambiental no contexto da paisagem de fragmentos florestais naturais &quot; ipucas &quot; no municipio de Lagoa da Confusao, Tocantins." url="https://doi.org/10.1590/S0100-67622002000300005" volume="26" year="2002">Martins et al. 2002</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="165">
The cave has 1,038 meters of total length, with predominantly ellipsoidal conduits. There are few speleothems, in addition to thick allochthonous sediments on the cave floor. The cave is inserted in a limestone outcrop located close to the Lagoa da
<normalizedToken originalValue="Confusão">Confusao</normalizedToken>
karstic lake (Figs
<figureCitation captionStart="Figure 33" captionStartId="F7" captionText="Figure 33. Location of the Casa de Pedra cave A South America, with Brazilian states highlighted B Tocantins state with Lagoa da Confusao municipality highlighted in white lines C aerial view from the Lagoa da Confusao region, where the urban area and the karst lake are visible; red star indicating the location of the Casa de Pedra cave D Casa de Pedra cave; red star indicating the location of Eupera troglobia sp. nov." figureDoi="10.3897/subtbiol.42.78074.figure33" httpUri="https://binary.pensoft.net/fig/671987" pageId="0" pageNumber="165">33C</figureCitation>
,
<figureCitation captionStart="Figure 34" captionStartId="F8" captionText="Figure 34. Lagoa da Confusao karst area A limestone outcrops close to the Lagoa da Confusao karst lake. The dark blue indicates the floodplain (not flooded in the moment of the photograph) and the dark blue indicates those flooded areas. The arrow indicates the main entrance of the Casa de Pedra cave B outcrop where the Casa de Pedra cave is located; 1. indicates the main entrance of the cave; 2. Upper entrance." figureDoi="10.3897/subtbiol.42.78074.figure34" httpUri="https://binary.pensoft.net/fig/671988" pageId="0" pageNumber="165">34A</figureCitation>
), which overflows during the rainy season, flooding part of the flood plain surrounding it. In such periods (October to April), most cave conduits are completely filled with water. On the other hand, the cave becomes dry during the dry season, as few intermittent dams are present.
</paragraph>
<paragraph pageId="0" pageNumber="165">
A visit paid to the cave in August 2021, revealed the cave partially flooded, with most conduits inaccessible. The main entrance gallery was filled with water, which was still forming a small lake outside the cave (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F8" captionText="Figure 34. Lagoa da Confusao karst area A limestone outcrops close to the Lagoa da Confusao karst lake. The dark blue indicates the floodplain (not flooded in the moment of the photograph) and the dark blue indicates those flooded areas. The arrow indicates the main entrance of the Casa de Pedra cave B outcrop where the Casa de Pedra cave is located; 1. indicates the main entrance of the cave; 2. Upper entrance." figureDoi="10.3897/subtbiol.42.78074.figure34" httpUri="https://binary.pensoft.net/fig/671988" pageId="0" pageNumber="165">34A</figureCitation>
). Reaching the deepest areas inside the cave through the main entrance was impossible in that moment, but since there is another entrance in the middle of the cave (Figs
<figureCitation captionStart="Figure 34" captionStartId="F8" captionText="Figure 34. Lagoa da Confusao karst area A limestone outcrops close to the Lagoa da Confusao karst lake. The dark blue indicates the floodplain (not flooded in the moment of the photograph) and the dark blue indicates those flooded areas. The arrow indicates the main entrance of the Casa de Pedra cave B outcrop where the Casa de Pedra cave is located; 1. indicates the main entrance of the cave; 2. Upper entrance." figureDoi="10.3897/subtbiol.42.78074.figure34" httpUri="https://binary.pensoft.net/fig/671988" pageId="0" pageNumber="165">34B</figureCitation>
,
<figureCitation captionStart="Figure 35" captionStartId="F9" captionText="Figure 35. Casa de Pedra cave A secondary upper entrance of the cave B cave chamber that was partially flooded C area inside the cave where specimens of E. troglobia sp. nov. were found D detail of the consolidated sediment indicating the area where submerged specimens of E. troglobia sp. nov. were found E E. troglobia sp. nov. specimens in situ exposed to the air F same, detail of an air exposed specimen, with a harvestman (Eusarcus sp.) near it G in situ submerged specimens of E. troglobia sp. nov. H location of a submerged specimen of E. troglobia sp. nov." figureDoi="10.3897/subtbiol.42.78074.figure35" httpUri="https://binary.pensoft.net/fig/671989" pageId="0" pageNumber="165">35A</figureCitation>
), the inner portions were accessible (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F9" captionText="Figure 35. Casa de Pedra cave A secondary upper entrance of the cave B cave chamber that was partially flooded C area inside the cave where specimens of E. troglobia sp. nov. were found D detail of the consolidated sediment indicating the area where submerged specimens of E. troglobia sp. nov. were found E E. troglobia sp. nov. specimens in situ exposed to the air F same, detail of an air exposed specimen, with a harvestman (Eusarcus sp.) near it G in situ submerged specimens of E. troglobia sp. nov. H location of a submerged specimen of E. troglobia sp. nov." figureDoi="10.3897/subtbiol.42.78074.figure35" httpUri="https://binary.pensoft.net/fig/671989" pageId="0" pageNumber="165">35B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="165">
Individuals of
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
sp. nov. were found associated to a consolidated sediment deposit in a deeper portion of the cave (Figs
<figureCitation captionStart="Figure 33" captionStartId="F7" captionText="Figure 33. Location of the Casa de Pedra cave A South America, with Brazilian states highlighted B Tocantins state with Lagoa da Confusao municipality highlighted in white lines C aerial view from the Lagoa da Confusao region, where the urban area and the karst lake are visible; red star indicating the location of the Casa de Pedra cave D Casa de Pedra cave; red star indicating the location of Eupera troglobia sp. nov." figureDoi="10.3897/subtbiol.42.78074.figure33" httpUri="https://binary.pensoft.net/fig/671987" pageId="0" pageNumber="165">33D</figureCitation>
,
<figureCitation captionStart="Figure 35" captionStartId="F9" captionText="Figure 35. Casa de Pedra cave A secondary upper entrance of the cave B cave chamber that was partially flooded C area inside the cave where specimens of E. troglobia sp. nov. were found D detail of the consolidated sediment indicating the area where submerged specimens of E. troglobia sp. nov. were found E E. troglobia sp. nov. specimens in situ exposed to the air F same, detail of an air exposed specimen, with a harvestman (Eusarcus sp.) near it G in situ submerged specimens of E. troglobia sp. nov. H location of a submerged specimen of E. troglobia sp. nov." figureDoi="10.3897/subtbiol.42.78074.figure35" httpUri="https://binary.pensoft.net/fig/671989" pageId="0" pageNumber="165">35C</figureCitation>
). Many specimens were adhered to the sediments already exposed to the air (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F9" captionText="Figure 35. Casa de Pedra cave A secondary upper entrance of the cave B cave chamber that was partially flooded C area inside the cave where specimens of E. troglobia sp. nov. were found D detail of the consolidated sediment indicating the area where submerged specimens of E. troglobia sp. nov. were found E E. troglobia sp. nov. specimens in situ exposed to the air F same, detail of an air exposed specimen, with a harvestman (Eusarcus sp.) near it G in situ submerged specimens of E. troglobia sp. nov. H location of a submerged specimen of E. troglobia sp. nov." figureDoi="10.3897/subtbiol.42.78074.figure35" httpUri="https://binary.pensoft.net/fig/671989" pageId="0" pageNumber="165">35E, F</figureCitation>
), while others were still under water (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F9" captionText="Figure 35. Casa de Pedra cave A secondary upper entrance of the cave B cave chamber that was partially flooded C area inside the cave where specimens of E. troglobia sp. nov. were found D detail of the consolidated sediment indicating the area where submerged specimens of E. troglobia sp. nov. were found E E. troglobia sp. nov. specimens in situ exposed to the air F same, detail of an air exposed specimen, with a harvestman (Eusarcus sp.) near it G in situ submerged specimens of E. troglobia sp. nov. H location of a submerged specimen of E. troglobia sp. nov." figureDoi="10.3897/subtbiol.42.78074.figure35" httpUri="https://binary.pensoft.net/fig/671989" pageId="0" pageNumber="165">35D, H, G</figureCitation>
). However, it is important to note that the cave was still drying up, so all specimens would be exposed to the air. Considering that the cave remains out of water during at least three months a year, the individuals do survive during all this period somehow avoiding desiccation. The only known cave-restricted clam species, all belonging to the genus
<taxonomicName class="Bivalvia" family="Dreissenidae" genus="Congeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Congeria" order="Myida" pageId="0" pageNumber="165" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Congeria</emphasis>
</taxonomicName>
, from caves in the Dinaric Alps, also exhibit this behavior, presenting a notable tolerance to air exposure (
<normalizedToken originalValue="Jovanović">Jovanovic</normalizedToken>
et al. 2016). Interestingly, in the case of all three
<taxonomicName class="Bivalvia" family="Dreissenidae" genus="Congeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Congeria" order="Myida" pageId="0" pageNumber="165" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Congeria</emphasis>
</taxonomicName>
species, only part of their populations becomes exposed during dry periods, and most part of the population remains underwater in such periods. Furthermore, even considering that
<taxonomicName class="Bivalvia" family="Dreissenidae" genus="Congeria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Congeria" order="Myida" pageId="0" pageNumber="165" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Congeria</emphasis>
</taxonomicName>
specimens are able to tolerate air exposure for periods as 2 months, some individuals were observed still active, with their shells open and inhalant and exhalant syphons extruded (
<normalizedToken originalValue="Jovanović">Jovanovic</normalizedToken>
et al. 2016). In the case of
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
sp. nov. the single visit paid to the cave does not make it possible to form any hypothesis regarding the
<normalizedToken originalValue="individuals’">individuals'</normalizedToken>
behavior along the air exposure (e.g., whether they remain active or not). Accordingly, it is highly recommendable that further studies investigate the biology and life cycle of this species. It is worth mentioning that
<bibRefCitation author="Silva, FJ" journalOrPublisher="Revista GeoNorte" pageId="0" pageNumber="165" publicationUrl="https://dspace.icmbio.gov.br//jspui/handle/cecav/416" refId="B20" refString="Silva, FJ, 2006. Levantamento da fauna de invertebrados e caracterizacao bioespeleologica da caverna Casa de Pedra, municipio Lagoa da Confusao-TO. Website (11/2021). https://dspace.icmbio.gov.br//jspui/handle/cecav/416" title="Levantamento da fauna de invertebrados e caracterizacao bioespeleologica da caverna Casa de Pedra, municipio Lagoa da Confusao-TO. Website (11 / 2021)." url="https://dspace.icmbio.gov.br//jspui/handle/cecav/416" year="2006">Silva (2006)</bibRefCitation>
, in her report from the present cave, mentioned the presence of clams also associated to root masses pending from the cave ceiling during their survey (in the dry period). In that case, all specimens were also exposed to the air, and there were only few small ponds inside the cave, apparently devoid of clams.
</paragraph>
<paragraph pageId="0" pageNumber="165">
During the clam sampling in August 2021, some hydrochemical parameters were evaluated, both inside the cave and in the epigean lake (Lagoa da
<normalizedToken originalValue="Confusão">Confusao</normalizedToken>
lake), which floods to the cave during rainy periods. The parameters inside the cave were quite distinct from those from the external lake: cave waters: temperature: 23.6 °C; pH 6.17; conductivity: 0.124 mS/cm; dissolved oxygen: 0.92 mg/L; TDS (total dissolved solids): 0.08 g/L; Salinity: 0.06‰; external lake: temperature: 28.1 °C; pH 7.14; conductivity: 0.017 mS/cm; dissolved oxygen: 10.35 mg/L; TDS (total dissolved solids): 0.011 g/L; Salinity: 0.01‰. It is noticeable the differences in temperature (lower inside the cave), pH (lower inside the cave), conductivity (higher inside the cave) and dissolved oxygen (much lower inside the cave). This certainly demonstrates that the species is not only able to survive in conditions quite distinct from those observed in surface waters, but also probably tolerates high levels of variation in hydrochemical parameters along the year, considering that the cave water originates from the lake flooding.
</paragraph>
<paragraph pageId="0" pageNumber="165">
Finally, it is also worth mentioning the number of embryos found in
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
. Although in the literature, it is usual to find the term
<normalizedToken originalValue="“embryo”">&quot;embryo&quot;</normalizedToken>
referring to both the true embryos and the young, such stages are, in fact, distinct. True embryos (still in ontogenetic development) are those individuals found in the visceral marsupium. Those found in the inner demibranchs are called
<normalizedToken originalValue="“young”">&quot;young&quot;</normalizedToken>
, as they are already formed and the shell shows growth lines. In
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
, there are a maximum of 10 young in each gill (~20 in total) and another 5-6 embryos in the visceral marsupium (on each side - 10-12 in total). Hence, the species presents around 30 immatures (considering both embryos and young). In the consulted literature, only the young specimens inside gills are considered. The other already studied
<taxonomicName authorityName="Bourguignat" authorityYear="1854" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera</emphasis>
</taxonomicName>
species (all epigean), presented a considerably larger reported number. As an example,
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. platensis" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="platensis">
<emphasis italics="true" pageId="0" pageNumber="165">E. platensis</emphasis>
</taxonomicName>
had between 22 and 66 young specimens in gills (
<bibRefCitation author="Ituarte, CF" journalOrPublisher="Iheringia" pageId="0" pageNumber="165" pagination="41 - 47" refId="B12" refString="Ituarte, CF, 1988. Caracteristicas de la incubacion branquial en Eupera platensis doello-jurado, 1921 y Byssanodonta paranensis d'orbigny, 1846 (Pelecypoda: Sphaeriidae). Iheringia 68: 41 - 47" title="Caracteristicas de la incubacion branquial en Eupera platensis doello-jurado, 1921 y Byssanodonta paranensis d'orbigny, 1846 (Pelecypoda: Sphaeriidae)." volume="68" year="1988">Ituarte 1988</bibRefCitation>
);
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. cubensis" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="cubensis">
<emphasis italics="true" pageId="0" pageNumber="165">E. cubensis</emphasis>
</taxonomicName>
between 25 and 35 (
<bibRefCitation author="Heard, WH" journalOrPublisher="Nautilus" pageId="0" pageNumber="165" pagination="47 - 49" refId="B9" refString="Heard, WH, 1964. Litter size in the Sphaeriidae. Nautilus 78 (2): 47 - 49" title="Litter size in the Sphaeriidae." volume="78" year="1964">Heard 1964</bibRefCitation>
) and
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. klappenbachi" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="klappenbachi">
<emphasis italics="true" pageId="0" pageNumber="165">E. klappenbachi</emphasis>
</taxonomicName>
had between 24 and 62 (
<bibRefCitation author="Mansur, MCD" journalOrPublisher="Iheringia (Zoologia)" pageId="0" pageNumber="165" pagination="23 - 46" refId="B16" refString="Mansur, MCD, Veitenheimer, IL, 1975. Nova especie de Eupera (Bivalvia: Sphaeriidae) e primeiros estudos anatomicos dentro do genero. Iheringia (Zoologia) 47: 23 - 46" title="Nova especie de Eupera (Bivalvia: Sphaeriidae) e primeiros estudos anatomicos dentro do genero." volume="47" year="1975">Mansur and Veitenheimer 1975</bibRefCitation>
). Most cave-restricted species from several distinct groups usually have k-strategies (
<bibRefCitation DOI="https://doi.org/10.1146/annurev.en.28.010183.002053" author="Howarth, FG" journalOrPublisher="Annual Review of Entomology" pageId="0" pageNumber="165" pagination="365 - 389" refId="B10" refString="Howarth, FG, 1983. Ecology of cave arthropods. Annual Review of Entomology 28 (1): 365 - 389, DOI: https://doi.org/10.1146/annurev.en.28.010183.002053" title="Ecology of cave arthropods." url="https://doi.org/10.1146/annurev.en.28.010183.002053" volume="28" year="1983">Howarth 1983</bibRefCitation>
;
<bibRefCitation author="Belles, X" journalOrPublisher="Oecologia Aquatica" pageId="0" pageNumber="165" pagination="325 - 335" refId="B1" refString="Belles, X, 1992. Survival, opportunism and convenience in the processes of cave colonization by terrestrial faunas. Homage to Ramon Margalef, Or, why there is such pleasure in studying nature. Oecologia Aquatica 10: 325 - 335" title="Survival, opportunism and convenience in the processes of cave colonization by terrestrial faunas. Homage to Ramon Margalef, Or, why there is such pleasure in studying nature." volume="10" year="1992">
<normalizedToken originalValue="Bellés">Belles</normalizedToken>
1992
</bibRefCitation>
), due to the relatively stable environments that subterranean habitats usually present. Among the reproductive adaptations related to such strategy, there are a reduced number of offspring, increased offspring body size, parental care, among others. The reduced number of embryos compared to some epigean
<taxonomicName authorityName="Bourguignat" authorityYear="1854" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera</emphasis>
</taxonomicName>
species, associated to the proportional large size of the young observed in the visceral marsupium of
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
, probably represent another adaptation to the cave environment, confirming the cave-restricted status of this species.
</paragraph>
<caption doi="10.3897/subtbiol.42.78074.figures1-6" httpUri="https://binary.pensoft.net/fig/671981" pageId="0" pageNumber="165" start="Figures 1–6" startId="F1">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figures 1-6.</emphasis>
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
holotype MZSP 155717 shell (L 4.3 mm), right valve in left column, left valve in right column
<emphasis bold="true" pageId="0" pageNumber="165">1</emphasis>
outer right view
<emphasis bold="true" pageId="0" pageNumber="165">2</emphasis>
outer left view
<emphasis bold="true" pageId="0" pageNumber="165">3</emphasis>
inner left view
<emphasis bold="true" pageId="0" pageNumber="165">4</emphasis>
inner right view
<emphasis bold="true" pageId="0" pageNumber="165">5</emphasis>
inner left-slightly ventral view
<emphasis bold="true" pageId="0" pageNumber="165">6</emphasis>
inner right-slightly ventral view.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.42.78074.figures7-13" httpUri="https://binary.pensoft.net/fig/671982" pageId="0" pageNumber="165" start="Figures 7–13" startId="F2">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figures 7-13.</emphasis>
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
shell SEM images of paratypes 153866
<emphasis bold="true" pageId="0" pageNumber="165">7</emphasis>
specimen #7, both valves connected, opened ~120°, ventral view
<emphasis bold="true" pageId="0" pageNumber="165">8</emphasis>
#7, left valve, inner left view
<emphasis bold="true" pageId="0" pageNumber="165">9</emphasis>
#7, right valve, inner left view
<emphasis bold="true" pageId="0" pageNumber="165">10</emphasis>
#7, left valve, detail of hinge region, inner left view
<emphasis bold="true" pageId="0" pageNumber="165">11</emphasis>
#7, right valve, detail of hinge region, inner right view
<emphasis bold="true" pageId="0" pageNumber="165">12</emphasis>
specimen #8, left valve, outer left view
<emphasis bold="true" pageId="0" pageNumber="165">13</emphasis>
same, detail of surface on middle region of ventral edge. Scale bars: 500
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="165">7, 12</emphasis>
), 300
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="165">8</emphasis>
), 200
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="165">10, 13</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.42.78074.figures14-21" httpUri="https://binary.pensoft.net/fig/671983" pageId="0" pageNumber="165" start="Figures 14–21" startId="F3">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figures 14-21.</emphasis>
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
shell SEM images of paratypes 153866
<emphasis bold="true" pageId="0" pageNumber="165">14</emphasis>
specimen #8 (part damaged), valves opened ~40°, ventral view
<emphasis bold="true" pageId="0" pageNumber="165">15</emphasis>
same, higher magnification
<emphasis bold="true" pageId="0" pageNumber="165">16</emphasis>
same, higher magnification, region of cardinal tooth
<emphasis bold="true" pageId="0" pageNumber="165">17</emphasis>
specimen #9 (part damaged), valves opened ~60°, ventral view, mainly showing hinge of left valve
<emphasis bold="true" pageId="0" pageNumber="165">18</emphasis>
same, right valve
<emphasis bold="true" pageId="0" pageNumber="165">19</emphasis>
shell of young specimen extracted from
<normalizedToken originalValue="gill’s">gill's</normalizedToken>
marsupium, left valve, outer left view
<emphasis bold="true" pageId="0" pageNumber="165">20</emphasis>
same, detail of umbo showing prodissoconch in its center, left-slightly dorsal view
<emphasis bold="true" pageId="0" pageNumber="165">21</emphasis>
another intra marsupial specimen (part damaged), right valve, inner ventral-slightly left view, focus on hinge, part of left valve still attached by ligament. Scale bars: 500
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="165">14, 17, 18</emphasis>
), 300
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="165">15</emphasis>
), 100
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="165">19</emphasis>
), 50
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="165">16, 21</emphasis>
), 30
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="165">20</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.42.78074.figures22-26" httpUri="https://binary.pensoft.net/fig/671984" pageId="0" pageNumber="165" start="Figures 22–26" startId="F4">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figures 22-26.</emphasis>
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
anatomical drawings
<emphasis bold="true" pageId="0" pageNumber="165">22</emphasis>
whole right view, right valve and part of right mantle lobe removed
<emphasis bold="true" pageId="0" pageNumber="165">23</emphasis>
gill, transverse section in its middle level
<emphasis bold="true" pageId="0" pageNumber="165">24</emphasis>
whole right view, right gill removed, visceral structures seen as in situ if region was transparent, peripherical structures with only topology indicated
<emphasis bold="true" pageId="0" pageNumber="165">25</emphasis>
palp region, ventral view, hemipalps slightly deflected
<emphasis bold="true" pageId="0" pageNumber="165">26</emphasis>
mantle edge, transverse section in middle level of ventral edge. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.42.78074.figures27-30" httpUri="https://binary.pensoft.net/fig/671985" pageId="0" pageNumber="165" start="Figures 27–30" startId="F5">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figures 27-30.</emphasis>
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
anatomical drawings
<emphasis bold="true" pageId="0" pageNumber="165">27</emphasis>
peri-siphonal, posterior region, right view, right gill and mantle lobe removed, siphons longitudinally sectioned, visceral structures seen as in situ
<emphasis bold="true" pageId="0" pageNumber="165">28</emphasis>
transition between palps and esophagus, right view, with concern to cerebral ganglia
<emphasis bold="true" pageId="0" pageNumber="165">29</emphasis>
pedal ganglia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="165">30</emphasis>
gross anatomy of young specimen from gill brood pouch, right view, right mantle lobe removed. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.42.78074.figures31-32" httpUri="https://binary.pensoft.net/fig/671986" pageId="0" pageNumber="165" start="Figures 31–32" startId="F6">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figures 31-32.</emphasis>
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
development, paratype MZSP 153866 pregnant specimen (2-3)
<emphasis bold="true" pageId="0" pageNumber="165">31</emphasis>
whole right view, shell and part of right mantle mole removed, young specimens (yo) seen in marsupium of inner demibranch by translucency
<emphasis bold="true" pageId="0" pageNumber="165">32</emphasis>
young specimen extracted from marsupium, right view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.42.78074.figure33" httpUri="https://binary.pensoft.net/fig/671987" pageId="0" pageNumber="165" start="Figure 33" startId="F7">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 33.</emphasis>
Location of the Casa de Pedra cave
<emphasis bold="true" pageId="0" pageNumber="165">A</emphasis>
South America, with Brazilian states highlighted
<emphasis bold="true" pageId="0" pageNumber="165">B</emphasis>
Tocantins state with Lagoa da
<normalizedToken originalValue="Confusão">Confusao</normalizedToken>
municipality highlighted in white lines
<emphasis bold="true" pageId="0" pageNumber="165">C</emphasis>
aerial view from the Lagoa da
<normalizedToken originalValue="Confusão">Confusao</normalizedToken>
region, where the urban area and the karst lake are visible; red star indicating the location of the Casa de Pedra cave
<emphasis bold="true" pageId="0" pageNumber="165">D</emphasis>
Casa de Pedra cave; red star indicating the location of
<taxonomicName authorityName="Simone &amp; Ferreira" authorityYear="2022" class="Bivalvia" family="Sphaeriidae" genus="Eupera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupera troglobia" order="Sphaeriida" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">Eupera troglobia</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.42.78074.figure34" httpUri="https://binary.pensoft.net/fig/671988" pageId="0" pageNumber="165" start="Figure 34" startId="F8">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 34.</emphasis>
Lagoa da
<normalizedToken originalValue="Confusão">Confusao</normalizedToken>
karst area
<emphasis bold="true" pageId="0" pageNumber="165">A</emphasis>
limestone outcrops close to the Lagoa da
<normalizedToken originalValue="Confusão">Confusao</normalizedToken>
karst lake. The dark blue indicates the floodplain (not flooded in the moment of the photograph) and the dark blue indicates those flooded areas. The arrow indicates the main entrance of the Casa de Pedra cave
<emphasis bold="true" pageId="0" pageNumber="165">B</emphasis>
outcrop where the Casa de Pedra cave is located; 1. indicates the main entrance of the cave; 2. Upper entrance.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.42.78074.figure35" httpUri="https://binary.pensoft.net/fig/671989" pageId="0" pageNumber="165" start="Figure 35" startId="F9">
<paragraph pageId="0" pageNumber="165">
<emphasis bold="true" pageId="0" pageNumber="165">Figure 35.</emphasis>
Casa de Pedra cave
<emphasis bold="true" pageId="0" pageNumber="165">A</emphasis>
secondary upper entrance of the cave
<emphasis bold="true" pageId="0" pageNumber="165">B</emphasis>
cave chamber that was partially flooded
<emphasis bold="true" pageId="0" pageNumber="165">C</emphasis>
area inside the cave where specimens of
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
sp. nov. were found
<emphasis bold="true" pageId="0" pageNumber="165">D</emphasis>
detail of the consolidated sediment indicating the area where submerged specimens of
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
sp. nov. were found
<emphasis bold="true" pageId="0" pageNumber="165">E</emphasis>
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
sp. nov. specimens in situ exposed to the air
<emphasis bold="true" pageId="0" pageNumber="165">F</emphasis>
same, detail of an air exposed specimen, with a harvestman (
<taxonomicName authorityName="Perty" authorityYear="1833" class="Arachnida" family="Gonyleptidae" genus="Eusarcus" kingdom="Animalia" lsidName="Eusarcus" order="Opiliones" pageId="0" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="165">Eusarcus</emphasis>
</taxonomicName>
sp.) near it
<emphasis bold="true" pageId="0" pageNumber="165">G</emphasis>
in situ submerged specimens of
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="165">H</emphasis>
location of a submerged specimen of
<taxonomicName genus="E." kingdom="Animalia" lsidName="E. troglobia" pageId="0" pageNumber="165" phylum="Mollusca" rank="species" species="troglobia">
<emphasis italics="true" pageId="0" pageNumber="165">E. troglobia</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>

122
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<taxonomicName LSID="http://species-id.net/wiki/Aspidimerus_esakii" authority="Sasaji, 1968" authorityName="Sasaji" authorityYear="1968" class="Insecta" family="Coccinellidae" genus="Aspidimerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aspidimerus esakii" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="esakii">Aspidimerus esakii Sasaji, 1968</taxonomicName>
Figs 15-17, 43-47, 85
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<subSubSection pageId="5" pageNumber="52" type="reference_group">
<paragraph pageId="5" pageNumber="52">
<taxonomicName class="Insecta" family="Coccinellidae" genus="Aspidimerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aspidimerus esakii" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="esakii">Aspidimerus esakii</taxonomicName>
Sasaji, 1968: 16; 1986: 40;
<bibRefCitation author="Pang, XF" journalOrPublisher="Science Press, Beijing" pageId="20" pageNumber="67" title="Economic Entomology of China 14: Coleoptera Coccinellidae II." year="1979">Pang and Mao 1979</bibRefCitation>
: 54;
<bibRefCitation author="Pang, H" journalOrPublisher="Entomotaxonomia" pageId="20" pageNumber="67" pagination="185 - 193" title="The new record and new species of Aspidimerinae (Coleoptera: Coccinellidae) from China." volume="20" year="1998">Pang 1998</bibRefCitation>
: 185;
<bibRefCitation author="Yu, GY" journalOrPublisher="Spixiana" pageId="20" pageNumber="67" pagination="123 - 144" title="The Coccinellidae (Excluding Epilachinae) collected by J. Klapperich in 1977 on Taiwan." volume="18" year="1995">Yu 1995</bibRefCitation>
: 139;
<bibRefCitation author="Yu, GY" journalOrPublisher="Chemical Industry Press, Beijing" pageId="20" pageNumber="67" title="The Coccinellidae of Taiwan." year="2011">2011</bibRefCitation>
: 166;
<bibRefCitation author="Kovar, I" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="20" pageNumber="67" pagination="71 - 73" title="Coccinellidae." year="2007">
<normalizedToken originalValue="Kovář">Kovar</normalizedToken>
2007
</bibRefCitation>
: 575;
<bibRefCitation author="Ren, SX" journalOrPublisher="Science Press, Beijing" pageId="20" pageNumber="67" title="Colored pictorial handbook of ladybird beetles in China." year="2009">Ren et al. 2009</bibRefCitation>
: 106.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="52" type="diagnosis">
<paragraph pageId="5" pageNumber="52">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="52">
This species is close to
<taxonomicName class="Insecta" family="Coccinellidae" genus="Aspidimerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aspidimerus chapaensis" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="chapaensis">Aspidimerus chapaensis</taxonomicName>
in general appearance, but can be separated from it by the black elytra with four yellow spots (Fig. 15), and the antennae with 8 antennomeres. In
<taxonomicName class="Insecta" family="Coccinellidae" genus="Aspidimerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aspidimerus chapaensis" order="Coleoptera" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="chapaensis">Aspidimerus chapaensis</taxonomicName>
, elytra black with four red spots and the antennae with 9 antennomeres. The spermatheca is also diagnostic according to the illustrations given by
<bibRefCitation author="Sasaji, H" journalOrPublisher="Fukui" pageId="20" pageNumber="67" pagination="1 - 24" title="A revision of the Formosan Coccinellidae (II): Tribe Stethorini, Aspidimerini and Chilocorini (Coleoptera)." volumeTitle="Etizenia" year="1968">Sasaji (1968)</bibRefCitation>
and
<bibRefCitation author="Hoang, DN" journalOrPublisher="Nha xuat ban khoa hoc va ky thuat, Hanoi" pageId="20" pageNumber="67" title="Coccinellidae of Vietnam Part I." year="1982">
<normalizedToken originalValue="Hoàng">Hoang</normalizedToken>
(1982)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="52" type="description">
<paragraph pageId="5" pageNumber="52">Description.</paragraph>
<paragraph pageId="5" pageNumber="52">TL: 3.70mm, TW: 2.90mm, TH: 1.75mm, TL/TW: 1.28; PL/PW: 0.53; EL/EW: 1.00.</paragraph>
<paragraph pageId="5" pageNumber="52">Body oblong oval, dorsum moderately convex and densely pubescent (Figs 15-17). Head yellowish brown, clypeus reddish brown, eyes black (Fig. 17). Pronotum black except anterior corners yellowish brown. Scutellum and elytra black. Each elytron with two yellowish brown spots (Figs 15-16). Underside black, except legs and abdomen reddish brown.</paragraph>
<paragraph pageId="5" pageNumber="52">Head small, 0.45 times elytral width (HW/EW=1:2.23). Punctures on frons small, separated by 0.5-1.0 times their diameter, with dense golden pubescence. Eyes large and oval, rather finely faceted, the widest interocular distance 0.50 times head width. Pronotum 0.69 times elytral width (PW/EW=1:1.45), punctures on pronotum and scutellum fine, close, separated by 0.5-1.0 times their diameter, with thick, golden pubescence. Elytra finely punctate, with short yellow white pubescence. Surface of prosternum coarse, with sparse, long and yellowish pubescence. Punctures on metaventrite moderately large, separated by 0.5-1.0 times their diameter, with dense yellowish pubescence.</paragraph>
<paragraph pageId="5" pageNumber="52">Male genitalia: Penis relatively long, slender and strongly curved, apex of penis with membranous appendage, penis capsule with a short indistinct outer process and a long inner one (Figs 43-45). Penis guide slender, gradually tapering to apex forming a pointed tip in lateral view (Fig. 47). In ventral view, penis guide broad, widest at basal 1/3 with truncate apex (Fig. 46). Parameres slender, sparsely setose at apex, equal in length to the penis guide (Fig. 47).</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="52" type="specimens examined">
<paragraph pageId="5" pageNumber="52">Specimens examined.</paragraph>
<paragraph pageId="5" pageNumber="52">
China, Hainan: 1♂, Bawangling Natural Reserve, [
<geoCoordinate direction="north" orientation="latitude" precision="9" value="19.0915">19°05.49'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="109.10633">109°06.38'E</geoCoordinate>
], ca 260m, 5.v.2005, Wang XM leg (SCAU).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="52" type="distribution">
<paragraph pageId="5" pageNumber="52">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="52">China (Hainan, Guangxi, Taiwan).</paragraph>
</subSubSection>
</treatment>
</document>