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; previousIdentifications: paratype of N. clavipes tenda; Location: islandGroup: Hawaiian Islands; island: Oahu; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Honolulu; verbatimLocality: Kaluanui Valley; verbatimElevation: 2000 ft.; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="14.iii.1946">14.iii.1946</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">BPBM</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="ii.1958" collectionCode="BPBM" collectorName="NLH Krauss" country="United States" location="United States" pageId="0" pageNumber="1110" specimenCount="5" specimenCount-adult="5" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="1110">NLH Krauss</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="1110">5</specimenCount>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Maui; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Maui; verbatimLocality: Nahiku; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="ii.1958">ii.1958</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">BPBM</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="31.vii.1958" collectionCode="BPBM" collectorName="LW Quate" country="United States" location="United States" pageId="0" pageNumber="1110" specimenCount="2" specimenCount-adult="2" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="1110">LW Quate</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="1110">2</specimenCount>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Hawaii; verbatimLocality: Kohala Mountains; verbatimElevation: 1900 ft.; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="31.vii.1958">31.vii.1958</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">BPBM</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="14.iii.1961" collectionCode="BPBM" collectorName="LW Quate" country="United States" location="United States" pageId="0" pageNumber="1110" specimenCount="3" specimenCount-adult="3" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="1110">LW Quate</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="1110">3</specimenCount>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Maui; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Maui; verbatimLocality: Waikamoi; verbatimElevation: 3936 ft.; Identification: identifiedBy: WW Wirth; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="14.iii.1961">14.iii.1961</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">BPBM</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="23.iii.1970" collectionCode="BPBM" collectorName="JL Gressitt" country="United States" location="United States" pageId="0" pageNumber="1110" specimenCount="1" specimenCount-adult="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="1110">JL Gressitt</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="1110">1</specimenCount>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Molokai; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Maui; verbatimLocality: West Halawa Valley; verbatimElevation: 164 ft.; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="23.iii.1970">23.iii.1970</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">BPBM</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="21.vii.1980" collectionCode="BPBM" collectorName="WA Steffan" country="United States" location="United States" pageId="0" pageNumber="1110" specimenCount="1" specimenCount-adult="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: recordedBy:
<collectorName pageId="0" pageNumber="1110">WA Steffan</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="1110">1</specimenCount>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Maui; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Maui; verbatimLocality: Kipahulu Valley, Pualuu Stream; verbatimElevation: 984 ft.; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="21.vii.1980">21.vii.1980</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">BPBM</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="4.i.2010" collectionCode="EMEC, 567.6" collectorName="PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja, E Owen" country="United States" latitude="22.131422" location="United States" longLatPrecision="1" longitude="-159.6217" pageId="0" pageNumber="1110" specimenCode="EMEC 7356041" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1110">EMEC7356041</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1110">PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja, E Owen</collectorName>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Kauai; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Kauai; verbatimLocality: Kawaikoi Stream at Sugi Grove; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="22.131422">22°7'53.12&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-159.6217">159°37'18.12&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="4.i.2010">4.i.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="1110">567.6</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="11.i.2010" collectionCode="EMEC, 590.3" collectorName="PM O'Grady, RT Lapoint, GM Bennett, BS Ort" country="United States" latitude="22.208723" location="United States" longLatPrecision="1" longitude="-159.59773" pageId="0" pageNumber="1110" specimenCode="EMEC 7356035" specimenCount="7" specimenCount-adult="7" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1110">EMEC7356035</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1110">PM O'Grady, RT Lapoint, GM Bennett, BS Ort</collectorName>
; individualCount:
<specimenCount pageId="0" pageNumber="1110">7</specimenCount>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Kauai; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Kauai; verbatimLocality: Hanapakai Stream, sweeping beach and stream; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="22.208723">22°12'31.40&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-159.59773">159°35'51.88&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="11.i.2010">11.i.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="1110">590.3</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="11.i.2010" collectionCode="EMEC, 592.2" collectorName="PM O'Grady, RT Lapoint, GM Bennett, BS Ort" country="United States" latitude="22.206366" location="United States" longLatPrecision="1" longitude="-159.6011" pageId="0" pageNumber="1110" specimenCode="EMEC 7356042" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1110">EMEC7356042</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1110">PM O'Grady, RT Lapoint, GM Bennett, BS Ort</collectorName>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Kauai; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Kauai; verbatimLocality: Honokoa Trail, seep #2; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="22.206366">22°12'22.91&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-159.6011">159°36'3.96&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="11.i.2010">11.i.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="1110">592.2</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="2.viii.2010" collectionCode="EMEC, 613.2" collectorName="PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja" country="United States" latitude="19.795055" location="United States" longLatPrecision="1" longitude="-155.09404" pageId="0" pageNumber="1110" specimenCode="EMEC 7356036" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1110">EMEC7356036</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1110">PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja</collectorName>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Hawaii; verbatimLocality: 0.5 miles from start of scenic drive to Onomea Bay, first major bridge, sweeping along stream; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="19.795055">19°47'42.20&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-155.09404">155°5'38.58&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="2.viii.2010">2.viii.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="1110">613.2</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="2.viii.2010" collectionCode="EMEC, 614.2" collectorName="PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja" country="United States" latitude="19.809038" location="United States" longLatPrecision="1" longitude="-155.09305" pageId="0" pageNumber="1110" specimenCode="EMEC 7356038" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1110">EMEC7356038</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1110">PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja</collectorName>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Hawaii; verbatimLocality: Alakahi Trail to Onomea Bay, adjacent to Hawaiia Botanical Garden, sweeping rocks in surf; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="19.809038">19°48'32.54&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-155.09305">155°5'34.98&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="2.viii.2010">2.viii.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="1110">614.2</collectionCode>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="2.viii.2010" collectionCode="EMEC, 615.2" collectorName="PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja" country="United States" latitude="19.811693" location="United States" longLatPrecision="1" longitude="-155.09386" pageId="0" pageNumber="1110" specimenCode="EMEC 7356039" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1110">EMEC7356039</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1110">PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja</collectorName>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Hawaii; verbatimLocality: Kawainui Stream, 6 ton bridge on Scenic Loop; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="19.811693">19°48'42.10&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-155.09386">155°5'37.88&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="2.viii.2010">2.viii.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
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</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="2.viii.2010" collectionCode="EMEC, 616.2" collectorName="PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja" country="United States" latitude="19.88276" location="United States" longLatPrecision="1" longitude="-155.11923" pageId="0" pageNumber="1110" specimenCode="EMEC 7356037" specimenCount="1" typeStatus="Other material">
Type status:
<typeStatus pageId="0" pageNumber="1110">Other material</typeStatus>
. Occurrence: catalogNumber:
<specimenCode pageId="0" pageNumber="1110">EMEC7356037</specimenCode>
; recordedBy:
<collectorName pageId="0" pageNumber="1110">PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja</collectorName>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Hawaii; verbatimLocality: Kolekole Beach Park, sweeping along ocean coast; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="19.88276">19°52'57.93&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-155.11923">155°7'9.22&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="2.viii.2010">2.viii.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
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</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="2.viii.2010" collectionCode="EMEC, 617.1" collectorName="PM O'Grady, RT Lapoint, GM Bennett, BS Ort, NA Pantoja" country="United States" latitude="20.076796" location="United States" longLatPrecision="1" longitude="-155.6191" pageId="0" pageNumber="1110" specimenCode="EMEC 7356040" specimenCount="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
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; recordedBy:
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; lifeStage:
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; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Hawaii; verbatimLocality: Waipio Stream, near the base of He'elawae Falls; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.076796">20°4'36.46&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-155.6191">155°37'8.72&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="2.viii.2010">2.viii.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
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</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="2.viii.2010" collectionCode="EMEC, BO 7.1" collectorName="BS Ort" country="United States" latitude="19.795055" location="United States" longLatPrecision="1" longitude="-155.09404" pageId="0" pageNumber="1110" specimenCode="EMEC 7356043" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
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; recordedBy:
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; individualCount:
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; sex:
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; lifeStage:
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; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
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; stateProvince: Hawaii; county: Hawaii; verbatimLocality: Ohanakupa Rd, 1st bridge at south end of road; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="19.795055">19°47'42.20&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-155.09404">155°5'38.58&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="2.viii.2010">2.viii.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
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</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
<materialsCitation collectingDate="2.viii.2010" collectionCode="EMEC, BO 10.2" collectorName="BS Ort" country="United States" latitude="20.076796" location="United States" longLatPrecision="1" longitude="-155.6191" pageId="0" pageNumber="1110" specimenCode="EMEC 7356044" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
Type status:
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. Occurrence: catalogNumber:
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; recordedBy:
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; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
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; stateProvince: Hawaii; county: Hawaii; verbatimLocality: Waipio Valley, near base of He'eilawe Falls at dammed reservoir; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.076796">20°4'36.46&quot;N</geoCoordinate>
; verbatimLongitude:
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; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="2.viii.2010">2.viii.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
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</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1110">
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Type status:
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. Occurrence: catalogNumber:
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; recordedBy:
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; individualCount:
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<specimenType pageId="0" pageNumber="1110">females</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="1110">adult</specimenType>
; Location: islandGroup: Hawaiian Islands; island: Hawaii; country:
<collectingCountry pageId="0" pageNumber="1110">United States</collectingCountry>
; stateProvince: Hawaii; county: Hawaii; verbatimLocality: Waipio Valley, near base of He'eilawe Falls at dammed reservoir; verbatimLatitude:
<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.076796">20°4'36.46&quot;N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-155.6191">155°37'8.72&quot;W</geoCoordinate>
; geodeticDatum: WGS84; Identification: identifiedBy: PM O'Grady; Event: verbatimEventDate:
<collectingDate pageId="0" pageNumber="1110" value="2.viii.2010">2.viii.2010</collectingDate>
; Record Level: institutionCode:
<collectionCode pageId="0" pageNumber="1110">EMEC</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="1110">BO10.2</collectionCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1110" type="ecological interactions">
<paragraph pageId="0" pageNumber="1110">Ecological interactions</paragraph>
<subSection pageId="0" pageNumber="1110" type="native status">
<paragraph pageId="0" pageNumber="1110">Native status</paragraph>
<paragraph pageId="0" pageNumber="1110">Endemic.</paragraph>
</subSection>
</subSubSection>
<subSubSection pageId="0" pageNumber="1110" type="distribution">
<paragraph pageId="0" pageNumber="1110">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1110">Hawaiian Islands: Kauai, Oahu, Molokai, Maui, Hawaii.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1110" type="notes">
<paragraph pageId="0" pageNumber="1110">Notes</paragraph>
<paragraph pageId="0" pageNumber="1110">52 specimens from USNM not examined.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,414 @@
<document ID-DOI="10.5281/zenodo.1038187" ID-GBIF-Dataset="5a8c5e73-63b2-445b-9ecc-96a601b1d901" ID-Zenodo-Dep="1038187" approvalRequired="18" approvalRequired_for_bibRefs="2" approvalRequired_for_illustrations="9" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="2" approvalRequired_for_treatments="2" checkinTime="1509026756856" checkinUser="jeremy" docAuthor="Leidy, J." docDate="1868" docId="BB1087B9FFBCFFC1C4828274FB1A9E19" docLanguage="en" docName="Leidy_1868_Megalosaurus(almost done).pdf.imf" docOrigin="Proceedings of the Academy of Natural Sciences of Philadelphia 20" docStyle="DocumentStyle{}" docTitle="Aublysodon mirandus Leidy 1868" docType="treatment" docVersion="8" lastPageNumber="200" masterDocId="4729FFC1FFBDFFC2C52D855AFFA69928" masterDocTitle="Remarks on a Jaw Fragment of Megalosaurus" masterLastPageNumber="200" masterPageNumber="197" pageNumber="198" updateTime="1673869449073" updateUser="jeremy">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Remarks on a Jaw Fragment of Megalosaurus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Leidy, J.</mods:namePart>
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<mods:titleInfo>
<mods:title>Proceedings of the Academy of Natural Sciences of Philadelphia</mods:title>
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<mods:part>
<mods:date>1868</mods:date>
<mods:detail type="pubDate">
<mods:number>1868-12-31</mods:number>
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<mods:detail type="volume">
<mods:number>20</mods:number>
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<mods:start>197</mods:start>
<mods:end>200</mods:end>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3508434" ID-GBIF-Taxon="165088550" ID-Zenodo-Dep="3508434" LSID="urn:lsid:plazi:treatment:BB1087B9FFBCFFC1C4828274FB1A9E19" httpUri="http://treatment.plazi.org/id/BB1087B9FFBCFFC1C4828274FB1A9E19" lastPageId="3" lastPageNumber="200" pageId="1" pageNumber="198">
<subSubSection pageId="1" pageNumber="198" type="discussion">
<paragraph blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">
The present opportunity is an appropriate one to make
<emphasis bold="true" box="[1242,1258,1838,1866]" pageId="1" pageNumber="198">a</emphasis>
few remarks on the American allies of
<emphasis box="[665,858,1871,1899]" italics="true" pageId="1" pageNumber="198">
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[665,854,1871,1899]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
.
</emphasis>
Since I have had the opportunity of inspecting the remains of the remarkable reptile from the green sand of New Jersey, described by Prof. Cope (
<treatmentCitation author="Cope" box="[724,949,1937,1965]" httpUri="http://tb.plazi.org/GgServer/html/A52387D30775FFC1FF97FAB1B85ED2A6" pageId="1" pageNumber="198" year="1866">Proc. 1866, 275</treatmentCitation>
) under the name of
<emphasis box="[1238,1508,1937,1965]" italics="true" pageId="1" pageNumber="198">
<taxonomicName authority="Cope 1866" authorityName="Cope" authorityYear="1866" baseAuthorityName="Cope" baseAuthorityYear="1866" box="[1238,1503,1937,1965]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="species" species="aquilunguis">Laelaps aquilunguis</taxonomicName>
,
</emphasis>
in observing the comparative
<emphasis box="[797,956,1971,1999]" italics="true" pageId="1" pageNumber="198">uniformity</emphasis>
of the teeth, identical in character with those of
<emphasis box="[530,722,2003,2031]" italics="true" pageId="1" pageNumber="198">
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[530,717,2003,2031]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
</emphasis>
I am more strongly impressed with the idea that the teeth of like shape forming part of those referred by me to
<emphasis box="[1330,1451,2036,2064]" italics="true" pageId="1" pageNumber="198">
<taxonomicName authorityName="Leidy" authorityYear="1856" box="[1330,1446,2036,2064]" class="Reptilia" family="Tyrannosauridae" genus="Dinodon" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Dinodon</taxonomicName>
,
</emphasis>
alone belong to this genus. The others, of which no representatives have been discovered or recognized as belonging to
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[1038,1222,2101,2129]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">
<emphasis box="[1038,1222,2101,2129]" italics="true" pageId="1" pageNumber="198">Megalosaurus</emphasis>
</taxonomicName>
or
<emphasis box="[1275,1383,2101,2129]" italics="true" pageId="1" pageNumber="198">
<taxonomicName authorityName="Cope" authorityYear="1866" box="[1275,1378,2101,2129]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Laelaps</taxonomicName>
,
</emphasis>
most probably indicate a distinct genus and species,
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="198" type="nomenclature">
<paragraph blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">
for which I propose the name of
<taxonomicName authorityName="Leidy" authorityYear="1868" box="[395,728,2167,2195]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="species" species="mirandus" status="sp. nov.">
<emphasis box="[395,577,2167,2195]" italics="true" pageId="1" pageNumber="198">Aublysodon</emphasis>
<emphasis box="[589,728,2171,2194]" italics="true" pageId="1" pageNumber="198">mirandus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="200" pageId="1" pageNumber="198" type="discussion">
<paragraph blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">
Future discovery may prove
<taxonomicName authorityName="Cope" authorityYear="1866" box="[871,971,2208,2236]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">
<emphasis box="[871,971,2208,2236]" italics="true" pageId="1" pageNumber="198">Laelaps</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Leidy" authorityYear="1856" box="[1064,1181,2208,2236]" class="Reptilia" family="Tyrannosauridae" genus="Dinodon" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">
<emphasis box="[1064,1181,2208,2236]" italics="true" pageId="1" pageNumber="198">Dinodon</emphasis>
</taxonomicName>
identical, and, judging from the comparison of corresponding parts of the jaws and the teeth, will be found to be more closely allied to
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[961,1146,2273,2301]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">
<emphasis box="[961,1146,2273,2301]" italics="true" pageId="1" pageNumber="198">Megalosaurus</emphasis>
</taxonomicName>
than was suspected, even should they not prove to be generically the same.
</paragraph>
<paragraph blockId="1.[388,1549,559,2517]" lastBlockId="2.[391,1561,555,2522]" lastPageId="2" lastPageNumber="199" pageId="1" pageNumber="198">
It is clear, from an examination of the anterior portion of the mandible of
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[393,579,2371,2399]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">
<emphasis box="[393,579,2371,2399]" italics="true" pageId="1" pageNumber="198">Megalosaurus</emphasis>
</taxonomicName>
described and figured by Buckland, Cuvier, Owen, etc., that no such teeth as those now referred to
<taxonomicName authorityName="Leidy" authorityYear="1868" box="[938,1099,2405,2433]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis box="[938,1099,2405,2433]" italics="true" pageId="1" pageNumber="198">Aublysodon</emphasis>
</taxonomicName>
occupied the forepart of the jaw. It is also probable that the upper teeth of
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[1186,1369,2437,2465]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">
<emphasis box="[1186,1369,2437,2465]" italics="true" pageId="1" pageNumber="198">Megalosaurus</emphasis>
</taxonomicName>
and of its allies differ in no important point from those below. It follows, therefore, that the teeth now referred to
<emphasis box="[851,1019,588,616]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authorityName="Leidy" authorityYear="1868" box="[851,1014,588,616]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus" status="gen. nov.">Aublysodon</taxonomicName>
,
</emphasis>
if they belong to the maxillary or mandibular series of
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[738,923,621,649]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[738,923,621,649]" italics="true" pageId="2" pageNumber="199">Megalosaurus</emphasis>
</taxonomicName>
or its allies, could only pertain to the back part. The variation in form of the teeth in question appears too great for such a position.
</paragraph>
<paragraph blockId="2.[391,1561,555,2522]" pageId="2" pageNumber="199">
The teeth now viewed as characteristic of
<taxonomicName authorityName="Leidy" authorityYear="1868" box="[1089,1252,721,749]" class="Reptilia" family="Tyrannosauridae" genus="Aublysodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis box="[1089,1252,721,749]" italics="true" pageId="2" pageNumber="199">Aublysodon</emphasis>
</taxonomicName>
are represented in
<figureCitation box="[396,661,754,782]" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="2" pageNumber="199" targetBox="[57,1772,29,2210]" targetPageId="17">figs. 36—45, pl. ix</figureCitation>
of vol. xi of the Transactions of the American Philosophical Society. The specimens consist of parts of three teeth, which differ much in size and other important points. In general the crowns are laterally compressed conical, with the anterior part thick and convex transversely as well as longitudinally, and with the sides nearly parallel. The posterior part forms a surface nearly as wide as the thickness of any part of the crown, and
<emphasis box="[396,419,953,981]" italics="true" pageId="2" pageNumber="199">is</emphasis>
defined from the lateral surfaces at right angles. In the two larger teeth these angles or borders are denticulated, like the trenchant borders of the teeth of
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[522,707,1019,1047]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[522,707,1019,1047]" italics="true" pageId="2" pageNumber="199">Megalosaurus</emphasis>
</taxonomicName>
and its American allies. In the longest tooth (
<figureCitation captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="2" pageNumber="199" targetBox="[57,1772,29,2210]" targetPageId="17">
fig. 35, 36, op.
<emphasis box="[508,552,1052,1080]" italics="true" pageId="2" pageNumber="199">cit</emphasis>
.
</figureCitation>
) the posterior surface forms an even plane; in the second sized tooth (
<figureCitation box="[498,674,1086,1114]" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="2" pageNumber="199" targetBox="[57,1772,29,2210]" targetPageId="17">figs. 37—40</figureCitation>
) the posterior surface presents a median elevation. In the smallest tooth (
<figureCitation box="[707,885,1119,1147]" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="2" pageNumber="199" targetBox="[57,1772,29,2210]" targetPageId="17">figs. 41—45</figureCitation>
), which indeed may belong to a different animal from the preceding, the borders defining the posterior surface are somewhat prominent backward, non-denticulate, and subside approaching the base of the crown so as to make a transverse section in this position oval (
<figureCitation box="[410,511,1256,1284]" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="2" pageNumber="199" targetBox="[57,1772,29,2210]" targetPageId="17">fig. 45</figureCitation>
).
</paragraph>
<paragraph blockId="2.[391,1561,555,2522]" pageId="2" pageNumber="199">
<emphasis box="[432,730,1284,1312]" italics="true" pageId="2" pageNumber="199">
<taxonomicName box="[432,725,1284,1312]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="species" species="foulkii">Hadrosaurus Foulkii</taxonomicName>
,
</emphasis>
the bulky vegetable feeder, and cotemporary of the rapacious
<emphasis box="[555,825,1318,1346]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authority="Cope 1866" authorityName="Cope" authorityYear="1866" baseAuthorityName="Cope" baseAuthorityYear="1866" box="[555,820,1318,1346]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="species" species="aquilunguis">Laelaps aquilunguis</taxonomicName>
,
</emphasis>
was at most probably only specifically distinct from
<emphasis box="[484,761,1351,1379]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authorityName="Leidy" authorityYear="1856" box="[484,756,1351,1379]" class="Reptilia" family="Hadrosauridae" genus="Trachodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="species" species="mirabilis">Trachodon mirabilis</taxonomicName>
,
</emphasis>
the teeth of which were found in association with those of
<emphasis box="[530,654,1384,1412]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authorityName="Leidy" authorityYear="1856" box="[530,649,1384,1412]" class="Reptilia" family="Tyrannosauridae" genus="Dinodon" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Dinodon</taxonomicName>
,
</emphasis>
so that, according to the laws of nomenclature, as
<taxonomicName box="[1408,1552,1384,1412]" class="Reptilia" family="Hadrosauridae" genus="Trachodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[1408,1552,1384,1412]" italics="true" pageId="2" pageNumber="199">Trachodon</emphasis>
</taxonomicName>
has priority of name, I suppose the first mentioned animal must be called
<taxonomicName box="[404,660,1451,1479]" class="Reptilia" family="Hadrosauridae" genus="Trachodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="species" species="foulkii">
<emphasis box="[404,546,1451,1479]" italics="true" pageId="2" pageNumber="199">Trachodon</emphasis>
Foulkii
</taxonomicName>
, though the names of
<taxonomicName box="[1019,1308,1451,1479]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="species" species="foulkii">
<emphasis box="[1019,1196,1451,1479]" italics="true" pageId="2" pageNumber="199">Hadrosaurus</emphasis>
Foulkii
</taxonomicName>
and
<taxonomicName baseAuthorityName="Leidy" baseAuthorityYear="1856" box="[1385,1553,1451,1479]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="species" species="mirabilis">
<emphasis box="[1385,1553,1451,1479]" italics="true" pageId="2" pageNumber="199">H. mirabilis</emphasis>
</taxonomicName>
would appear more appropriate for these powerful dinosaurs.
</paragraph>
<paragraph blockId="2.[391,1561,555,2522]" pageId="2" pageNumber="199">
The best preserved tooth of those originally referred to
<emphasis box="[1300,1449,1517,1545]" italics="true" pageId="2" pageNumber="199">
<taxonomicName box="[1300,1444,1517,1545]" class="Reptilia" family="Hadrosauridae" genus="Trachodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Trachodon</taxonomicName>
,
</emphasis>
represented in
<figureCitation box="[549,686,1551,1579]" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="2" pageNumber="199" targetBox="[57,1772,29,2210]" targetPageId="17">
<emphasis box="[549,602,1551,1579]" italics="true" pageId="2" pageNumber="199">figs</emphasis>
. 1—6
</figureCitation>
of the plate above cited, is identical in form with those referred to
<emphasis box="[572,756,1584,1612]" italics="true" pageId="2" pageNumber="199">
<taxonomicName box="[572,751,1584,1612]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Hadrosaurus</taxonomicName>
,
</emphasis>
and differs only in the absence of the rugulations of the lateral borders of the crown, and in some less important points.
</paragraph>
<paragraph blockId="2.[391,1561,555,2522]" pageId="2" pageNumber="199">
The remaining specimens of teeth referred with the former to
<emphasis box="[1401,1550,1650,1678]" italics="true" pageId="2" pageNumber="199">
<taxonomicName box="[1401,1545,1650,1678]" class="Reptilia" family="Hadrosauridae" genus="Trachodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Trachodon</taxonomicName>
,
</emphasis>
are represented in
<figureCitation box="[686,839,1683,1711]" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="2" pageNumber="199" targetBox="[57,1772,29,2210]" targetPageId="17">figs. 7—20</figureCitation>
of the plate cited. Most of them are so worn and probably altered from their original
<emphasis box="[1030,1101,1716,1744]" italics="true" pageId="2" pageNumber="199">form</emphasis>
, that it is rendered uncertain whether they belong to the same animal as the preceding tooth, and one unworn (
<figureCitation box="[539,716,1783,1811]" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="2" pageNumber="199" targetBox="[57,1772,29,2210]" targetPageId="17">figs. 18—20</figureCitation>
) has a very different shape from this. Perhaps these specimens belonged to another Dinosaur, for which the name
<taxonomicName box="[1317,1456,1816,1844]" class="Reptilia" family="Hadrosauridae" genus="Trachodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[1317,1456,1816,1844]" italics="true" pageId="2" pageNumber="199">Trachodon</emphasis>
</taxonomicName>
might be reserved, while that of
<taxonomicName box="[794,971,1849,1877]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[794,971,1849,1877]" italics="true" pageId="2" pageNumber="199">Hadrosaurus</emphasis>
</taxonomicName>
might include the first mentioned and more characteristic tooth.
</paragraph>
<paragraph blockId="2.[391,1561,555,2522]" pageId="2" pageNumber="199">
As
<taxonomicName authorityName="Mantell" authorityYear="1825" box="[494,638,1916,1944]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[494,638,1916,1944]" italics="true" pageId="2" pageNumber="199">Iguanodon</emphasis>
</taxonomicName>
had its enemy in a species of
<emphasis box="[1135,1493,1916,1944]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[1135,1321,1916,1944]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
<taxonomicName box="[1342,1488,1916,1944]" class="Reptilia" family="Hadrosauridae" genus="Trachodon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Trachodon</taxonomicName>
,
</emphasis>
the representative of the former both in the western and eastern portions of the North American continent, was accompanied by an equally
<emphasis box="[1371,1554,1982,2010]" italics="true" pageId="2" pageNumber="199">bloodthirsty</emphasis>
enemy, which may, perhaps, on nearer comparison of corresponding parts, prove to be another species of the same genus, until now supposed to be
<emphasis box="[405,532,2082,2110]" italics="true" pageId="2" pageNumber="199">different</emphasis>
, under the names of
<taxonomicName authorityName="Leidy" authorityYear="1856" box="[849,965,2082,2110]" class="Reptilia" family="Tyrannosauridae" genus="Dinodon" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[849,965,2082,2110]" italics="true" pageId="2" pageNumber="199">Dinodon</emphasis>
</taxonomicName>
and
<emphasis box="[1043,1149,2082,2110]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authorityName="Cope" authorityYear="1866" box="[1043,1145,2082,2110]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Laelaps</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph blockId="2.[391,1561,555,2522]" pageId="2" pageNumber="199">
Prof. Cope remarks of
<taxonomicName authorityName="Cope" authorityYear="1866" box="[775,874,2115,2143]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[775,874,2115,2143]" italics="true" pageId="2" pageNumber="199">Laelaps</emphasis>
</taxonomicName>
(
<treatmentCitation author="Cope" box="[898,1213,2115,2143]" httpUri="http://tb.plazi.org/GgServer/html/A52387D30775FFC1FF97FAB1B85ED2A6" pageId="2" pageNumber="199" year="1866">
Pr. A.
<emphasis box="[990,1016,2115,2143]" italics="true" pageId="2" pageNumber="199">N</emphasis>
. S. 1866, 276
</treatmentCitation>
), that “in
<emphasis box="[1363,1397,2115,2143]" italics="true" pageId="2" pageNumber="199">its</emphasis>
dentition and huge prehensile claws it resembled closely
<emphasis box="[1122,1312,2147,2175]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[1122,1306,2147,2175]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
</emphasis>
but the
<emphasis box="[1458,1547,2147,2175]" italics="true" pageId="2" pageNumber="199">femur</emphasis>
, resembling in its proximal regions more nearly the
<emphasis box="[1189,1340,2181,2209]" italics="true" pageId="2" pageNumber="199">lguanodon,</emphasis>
indicated the probable existence of other equally important
<emphasis box="[1102,1262,2214,2242]" italics="true" pageId="2" pageNumber="199">differences</emphasis>
, and its pertinence to another genus.” Thus the genus is especially distinguished by the apparent peculiarity of the
<emphasis box="[755,844,2279,2307]" italics="true" pageId="2" pageNumber="199">femur</emphasis>
, but in
<emphasis box="[973,1017,2279,2307]" italics="true" pageId="2" pageNumber="199">my</emphasis>
estimation even this disappears if the bone referred to
<taxonomicName authorityName="Cope" authorityYear="1866" box="[716,814,2311,2339]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">
<emphasis box="[716,814,2311,2339]" italics="true" pageId="2" pageNumber="199">Laelaps</emphasis>
</taxonomicName>
be viewed in the corresponding position to that of
<emphasis box="[450,645,2345,2373]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authority="Buckland, 1824" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1837" box="[450,640,2345,2373]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="species" species="bucklandi">M. Bucklandi</taxonomicName>
,
</emphasis>
represented in pl. vii, pt.
<emphasis box="[1049,1083,2345,2373]" italics="true" pageId="2" pageNumber="199">iii</emphasis>
, of Prof. Owens Monograph of the Fossil Reptiles of the
<emphasis box="[799,929,2377,2405]" italics="true" pageId="2" pageNumber="199">Wealden</emphasis>
, which appears to me to be the reversed one to that in which Prof
<emphasis box="[812,883,2410,2438]" italics="true" pageId="2" pageNumber="199">Cope</emphasis>
has described it in
<treatmentCitation author="Cope" box="[1183,1511,2410,2438]" httpUri="http://tb.plazi.org/GgServer/html/A52387D30775FFC1FF97FAB1B85ED2A6" pageId="2" pageNumber="199" year="1866">
Pr. A.
<emphasis box="[1285,1312,2410,2438]" italics="true" pageId="2" pageNumber="199">N</emphasis>
.
<emphasis box="[1331,1353,2410,2438]" italics="true" pageId="2" pageNumber="199">S</emphasis>
. 1866, 276
</treatmentCitation>
.
</paragraph>
<paragraph blockId="2.[391,1561,555,2522]" lastBlockId="3.[392,1558,549,2511]" lastPageId="3" lastPageNumber="200" pageId="2" pageNumber="199">
The teeth of
<emphasis box="[635,832,2442,2470]" italics="true" pageId="2" pageNumber="199">
<taxonomicName authorityName="Leidy" authorityYear="1853" box="[635,828,2442,2470]" class="Reptilia" family="Sphenacodontidae" genus="Bathygnathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelycosauria" pageId="2" pageNumber="199" phylum="Chordata" rank="genus">Bathygnathus</taxonomicName>
,
</emphasis>
a huge carnivorous reptile, whose remains have been found in the triassic red sandstone of Prince Edwards Island, have the same form as those of
<emphasis box="[825,1147,584,612]" italics="true" pageId="3" pageNumber="200">
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[825,1013,584,612]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
<taxonomicName authorityName="Leidy" authorityYear="1856" box="[1032,1147,584,612]" class="Reptilia" family="Tyrannosauridae" genus="Dinodon" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Dinodon</taxonomicName>
</emphasis>
and
<emphasis box="[1227,1334,584,612]" italics="true" pageId="3" pageNumber="200">
<taxonomicName authorityName="Cope" authorityYear="1866" box="[1227,1330,584,612]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Laelaps</taxonomicName>
.
</emphasis>
But here, so far as we have the corresponding parts for comparison, the resemblance ceases. The remarkable depth of the dentary bone in relation with its length in
<emphasis box="[553,751,684,712]" italics="true" pageId="3" pageNumber="200">
<taxonomicName authorityName="Leidy" authorityYear="1853" box="[553,746,684,712]" class="Reptilia" family="Sphenacodontidae" genus="Bathygnathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelycosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Bathygnathus</taxonomicName>
,
</emphasis>
indicates a form of head very different from that of
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[405,589,717,745]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">
<emphasis box="[405,589,717,745]" italics="true" pageId="3" pageNumber="200">Megalosaurus</emphasis>
</taxonomicName>
and its American representatives. It was this unusual relation of depth to breadth which led me to suspect a form of head more in accordance with that of the skeleton of an upright animal, and led me to ask the question, “was this animal probably not one of the bipeds which made the so-called bird tracks of the New Red Sandstone of the valley of the Connecticut?” (See Jour. Ac. Nat. Sc. 1854, 329)
</paragraph>
<paragraph blockId="3.[392,1558,549,2511]" pageId="3" pageNumber="200">
Subsequently, in examining the remains of
<emphasis box="[1135,1322,916,944]" italics="true" pageId="3" pageNumber="200">
<taxonomicName box="[1135,1317,916,944]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Hadrosaurus</taxonomicName>
,
</emphasis>
the American representative of
<emphasis box="[668,818,950,978]" italics="true" pageId="3" pageNumber="200">
<taxonomicName authorityName="Mantell" authorityYear="1825" box="[668,813,950,978]" class="Reptilia" family="Iguanodontidae" genus="Iguanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Iguanodon</taxonomicName>
,
</emphasis>
from the great disproportion between the fore and hind parts of the body, I was led “to suspect that this great herbivorous lizard sustained itself in a semi-erect position on the huge hinder extremities and tail, while it browsed on plants growing upon the shores of the ocean.” (Cret. Rept, of the U. S. 1865, 97.)
</paragraph>
<paragraph blockId="3.[392,1558,549,2511]" pageId="3" pageNumber="200">
The remains referred to
<taxonomicName authorityName="Cope" authorityYear="1866" box="[819,916,1116,1144]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">
<emphasis box="[819,916,1116,1144]" italics="true" pageId="3" pageNumber="200">Laelaps</emphasis>
</taxonomicName>
exhibit even a far greater disproportion between the fore and hind limbs than in
<emphasis box="[1044,1228,1149,1177]" italics="true" pageId="3" pageNumber="200">
<taxonomicName box="[1044,1223,1149,1177]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Hadrosaurus</taxonomicName>
,
</emphasis>
which, together with its long bird-like claws, etc., suggested to Prof. Cope a similar position
<emphasis bold="true" box="[1526,1556,1182,1210]" pageId="3" pageNumber="200">of</emphasis>
body to that of
<emphasis box="[637,822,1215,1243]" italics="true" pageId="3" pageNumber="200">
<taxonomicName box="[637,818,1215,1243]" class="Reptilia" family="Hadrosauridae" genus="Hadrosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Hadrosaurus</taxonomicName>
,
</emphasis>
and a use of the hind limbs in attack upon the prey of the animal analagous with that in the eagle (
<treatmentCitation author="Cope" box="[1207,1538,1248,1276]" httpUri="http://tb.plazi.org/GgServer/html/A52387D30775FFC1FF97FAB1B85ED2A6" pageId="3" pageNumber="200" year="1866">Pr. A. N. S. 1866, 279</treatmentCitation>
). The extraordinary disproportion between the fore and hind limbs of
<emphasis box="[1448,1552,1282,1310]" italics="true" pageId="3" pageNumber="200">
<taxonomicName authorityName="Cope" authorityYear="1866" box="[1448,1547,1282,1310]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Laelaps</taxonomicName>
,
</emphasis>
which appears to me so closely related with
<emphasis box="[1083,1275,1315,1343]" italics="true" pageId="3" pageNumber="200">
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[1083,1271,1315,1343]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
</emphasis>
leads me to suspect that the remains described by Buckland, Cuvier, Owen and others, and attributed to the shoulder of
<taxonomicName authority="Buckland, 1824" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1837" box="[846,1037,1381,1409]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="species" species="bucklandi">
<emphasis box="[846,881,1381,1409]" italics="true" pageId="3" pageNumber="200">M.</emphasis>
Bucklandi
</taxonomicName>
, perhaps, at least in part, belong to the pelvis, if they in whole or part do not belong to other animals. Had the humerus of
<taxonomicName authorityName="Cope" authorityYear="1866" box="[640,738,1447,1475]" class="Arachnida" family="Tyrannosauridae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">
<emphasis box="[640,738,1447,1475]" italics="true" pageId="3" pageNumber="200">Laelaps</emphasis>
</taxonomicName>
been found isolated, I never would have thought of associating it in the same skeleton with the huge bones of the hinder
<emphasis bold="true" box="[1503,1548,1481,1509]" pageId="3" pageNumber="200">extremity</emphasis>
of that animal. Perhaps, when this great disproportion comes to be known, it may be discovered that there exist specimens of remains of the fore limbs of
<emphasis box="[605,795,1581,1609]" italics="true" pageId="3" pageNumber="200">
<taxonomicName authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[605,789,1581,1609]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
</emphasis>
from the Wealden, in the British or other museums of England, which heretofore have excited no suspicion as to their true relations.
</paragraph>
<paragraph blockId="3.[392,1558,549,2511]" pageId="3" pageNumber="200">
<emphasis box="[436,617,1681,1709]" italics="true" pageId="3" pageNumber="200">
<taxonomicName authorityName="von Meyer" authorityYear="1861" box="[436,612,1681,1709]" class="Reptilia" family="Megalosauridae" genus="Teratosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">Teratosaurus</taxonomicName>
,
</emphasis>
from the upper Keuper, in the vicinity of Stuttgart, described by Meyer (Palaeontographica, 1859-61, 258), approached
<taxonomicName authorityName="Leidy" authorityYear="1853" box="[1269,1458,1713,1741]" class="Reptilia" family="Sphenacodontidae" genus="Bathygnathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelycosauria" pageId="3" pageNumber="200" phylum="Chordata" rank="genus">
<emphasis box="[1269,1458,1713,1741]" italics="true" pageId="3" pageNumber="200">Bathygnathus</emphasis>
</taxonomicName>
most in the proportions of its face, as well as resembled it in the form of the teeth, but the fossil dentarybone of the latter is even still shorter and deeper than would relate to the fossil maxillary of the former.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,132 @@
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<emphasis id="01CDEABDFFBDFFC2C49F8355FE6C9F03" box="[434,458,1551,1579]" italics="true" pageId="0" pageNumber="197">A</emphasis>
fossil worthy of notice in the Museum of the Academy consists of the fragment of a jaw, apparently of the
<taxonomicName id="F4B94D2CFFBDFFC2C6E8836BFB379F65" authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[965,1169,1585,1613]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="197" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
, which, if it does not belong to a different species from
<taxonomicName id="F4B94D2CFFBDFFC2C6AF830BFBE79F45" authority="Buckland, 1824" baseAuthorityName="Eudes-Deslongchamps" baseAuthorityYear="1837" box="[898,1089,1617,1645]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="197" phylum="Chordata" rank="species" species="bucklandi">
<emphasis id="01CDEABDFFBDFFC2C6AF830BFC039F45" box="[898,933,1617,1645]" italics="true" pageId="0" pageNumber="197">M.</emphasis>
Bucklandi
</taxonomicName>
, indicates an individual larger than any one of those referred to by Buckland, Cuvier, Owen, etc. The fossil was purchased in England, and was presented to the Academy by Dr. Thomas B. Wilson. It is labelled, “ Fragment dune machoire de
<taxonomicName id="F4B94D2CFFBDFFC2C1C883EFFA0B9FF9" authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[1253,1453,1717,1745]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="197" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
trouvé dans le lias a Boué (or Boues). Lanimal est extremement rare ici. Il avait 45 pied de longeur.” In another hand it is marked “
<emphasis id="01CDEABDFFBDFFC2C19B83ADFA989E3B" box="[1206,1342,1783,1811]" italics="true" pageId="0" pageNumber="197">Jura Mts</emphasis>
.”
</paragraph>
<paragraph id="330636AFFFBDFFC2C4998243FAFC9EB0" blockId="0.[399,1561,1449,2525]" pageId="0" pageNumber="197">The fragment contains two mutilated teeth, visible throughout their length from the inner part of the jaw being broken away. The matrix adhering to the fossil consists of an oolite composed of a homogeneous clay-colored basis, with imbedded granules, of a rounded form, brown and shining.</paragraph>
</subSubSection>
<subSubSection id="7BA36524FFBDFFC2C49882C4FC969189" pageId="0" pageNumber="197" type="nomenclature">
<paragraph id="330636AFFFBDFFC2C49882C4FC969189" blockId="0.[399,1561,1449,2525]" pageId="0" pageNumber="197">
The teeth are inserted into the jaw about two-thirds their length, and more than three-fourths the depth of the bone. They have measured 5 1/2 and 6 inches in length. The breadth at the base of the enamelled crown of the best preserved tooth is 14 3/4 lines, which is nearly the fourth of an inch greater than in the largest tooth represented in any of Prof. Owens figures in his Monograph of the Fossil Reptiles of the Wealden Formation.
<emphasis id="01CDEABDFFBDFFC2C08C8D19FA1F9177" box="[1441,1465,2115,2143]" italics="true" pageId="0" pageNumber="197">A</emphasis>
tooth apparently nearly as large in an American ally, is one referred to
<emphasis id="01CDEABDFFBDFFC2C08C8D3EFDA89189" italics="true" pageId="0" pageNumber="197">
<taxonomicName id="F4B94D2CFFBDFFC2C08C8D3EFDAF9189" authorityName="Leidy" authorityYear="1856" class="Reptilia" family="Tyrannosauridae" genus="Dinodon" kingdom="Animalia" order="Dinosauria" pageId="0" pageNumber="197" phylum="Chordata" rank="species" species="horridus" status="sp. nov.">Dinodon horridus</taxonomicName>
,
</emphasis>
and represented in
</paragraph>
</subSubSection>
<subSubSection id="7BA36524FFBDFFC2C6138DDFFCFE906E" pageId="0" pageNumber="197" type="reference_group">
<paragraph id="330636AFFFBDFFC2C6138DDFFCFE906E" blockId="0.[399,1561,1449,2525]" pageId="0" pageNumber="197">
<treatmentCitation id="B21810BEFFBDFFC2C6138DDFFB9491CB" author="Leidy" httpUri="https://treatment.plazi.org/id/E8728790-2E74-CC2D-17EA-FEF0FE4CF067" pageId="0" pageNumber="197" year="1860">
<figureCitation id="AB822A2AFFBDFFC2C6138DDFFC4E9189" box="[830,1000,2181,2209]" captionText="1-20. Trachodon mirabilis 21-48. Deinodon horridus 49-52. Palaeoscincus costatus 53-55. Troodon formosus." httpUri="https://zenodo.org/record/1068922/files/figure.png" pageId="0" pageNumber="197" targetBox="[57,1772,29,2210]" targetPageId="17">fig. 21, pl. 9</figureCitation>
, of
<bibRefCitation id="57284B5EFFBDFFC2C10F8DDFFB6C9189" author="Leidy, J." box="[1058,1226,2181,2209]" journalOrPublisher="Transactions of the American Philosophical Society" pageId="0" pageNumber="197" pagination="139-154" part="11" title="Extinct vertebrates from the Judith River and Great Lignite Formations of Nebraska" type="journal article" year="1860">my memoir</bibRefCitation>
on the
<bibRef id="EA19A259FFBDFFC2C06D8DDFFB9491CB" author="Leidy, J." journalOrPublisher="Transactions of the American Philosophical Society" pageId="0" pageNumber="197" pagination="139-154" part="11" title="Extinct vertebrates from the Judith River and Great Lignite Formations of Nebraska" type="journal article" year="1860">Extinct Vertebrata of the Judith River, published in the eleventh volume of the Transactions of the American Philosophical Society</bibRef>
</treatmentCitation>
. The reconstructed outline of this figure is, however, too large, rendered so by the too distant removal of the apex of the tooth from the other fragment. The breadth of this specimen really did not exceed an inch.
</paragraph>
</subSubSection>
<subSubSection id="7BA36524FFBDFFC3C4948C10FC3A9E30" lastPageId="1" lastPageNumber="198" pageId="0" pageNumber="197" type="description">
<paragraph id="330636AFFFBDFFC3C4948C10FCD59BB9" blockId="0.[399,1561,1449,2525]" lastBlockId="1.[388,1549,559,2517]" lastPageId="1" lastPageNumber="198" pageId="0" pageNumber="197">
The longest tooth of the fossil under inspection, for the most part broken away, exhibits a mould of the large interior pulp cavity. This mould, from the bottom of the latter to its broken end in the position of the crown, is 5 1/4 inches long. The broken end is 8 lines wide and 1 3/4 lines thick;
<emphasis id="01CDEABDFFBCFFC3C04B8768FA329B66" box="[1382,1428,562,590]" italics="true" pageId="1" pageNumber="198">the</emphasis>
widest and thickest part of the mould near the middle of the length of the tooth is 11 1/2 lines wide and 5 lines thick.
</paragraph>
<paragraph id="330636AFFFBCFFC3C48287CCFDBF9AB2" blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">
The fangs of the teeth do not continue of the same width to the bottom, as in the teeth of crocodiles, and, as I believe, is considered to be the case in
<emphasis id="01CDEABDFFBCFFC3C4A38782FDEB9BDC" box="[398,589,728,756]" italics="true" pageId="1" pageNumber="198">
<taxonomicName id="F4B94D2CFFBCFFC3C4A38782FDEF9BDC" authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[398,585,728,756]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
</emphasis>
but from about their middle they contract, or become narrower, as is ordinarily the case in mammals. Indeed, one of these teeth isolated might be taken for the canine of a
<emphasis id="01CDEABDFFBCFFC3C69C8640FBF89A1E" box="[945,1118,794,822]" italics="true" pageId="1" pageNumber="198">
<taxonomicName id="F4B94D2CFFBCFFC3C69C8640FBFF9A1E" authorityName="Nesti" authorityYear="1826" box="[945,1113,794,822]" class="Squamata" family="Colubridae" genus="Drepanodon" higherTaxonomySource="CoL" kingdom="Animalia" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Drepanodon</taxonomicName>
,
</emphasis>
or sabre-toothed tiger. In the fossil the bottoms of the fangs narrow antero-posteriorly, and become thinner from without inwardly, and they also curve somewhat in the latter direction.
</paragraph>
<paragraph id="330636AFFFBCFFC3C48086C5FDFE9D48" blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">
The long fangs of the teeth in the fossil, and their becoming narrowed at bottom, at first led me to suspect the specimen belonged to a different genus from
<emphasis id="01CDEABDFFBCFFC3C4CD86BBFD069AD5" box="[480,672,993,1021]" italics="true" pageId="1" pageNumber="198">
<taxonomicName id="F4B94D2CFFBCFFC3C4CD86BBFD3D9AD5" authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[480,667,993,1021]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
</emphasis>
but a view of fig. 1, plate xii, of Prof. Owens monograph above mentioned, seems to prove by the appearance of the successional teeth within the jaw, that the fangs actually become narrowed towards the bottom in that genus.
</paragraph>
<paragraph id="330636AFFFBCFFC3C482813CFC519DCD" blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">
In the best preserved tooth of the fossil, the enamelled crown exhibits the same shape, familiar as the characteristic form of that of
<emphasis id="01CDEABDFFBCFFC3C1DD81DDFA179D8B" box="[1264,1457,1159,1187]" italics="true" pageId="1" pageNumber="198">
<taxonomicName id="F4B94D2CFFBCFFC3C1DD81DDFA0B9D8B" authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[1264,1453,1159,1187]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
.
</emphasis>
The trenchant borders of the crown are denticulate, and the enamel is comparatively smooth, or only very feebly striate.
</paragraph>
<paragraph id="330636AFFFBCFFC3C49F81B0FD089C84" blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">The contracted condition of the bottom of the fangs of the teeth would leave more space than there otherwise would be for the development of successional teeth within the jaw. In the fossil the remains of one of the latter is seen at the lower part internally of one of the functional teeth, and an impression in a corresponding position of the other functional tooth indicates a similar occupant.</paragraph>
<paragraph id="330636AFFFBCFFC3C48280EBFE6E9F9F" blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">
In the progress of the successional teeth of
<emphasis id="01CDEABDFFBCFFC3C15D80EBFA949CE5" box="[1136,1330,1457,1485]" italics="true" pageId="1" pageNumber="198">
<taxonomicName id="F4B94D2CFFBCFFC3C15D80EBFA8B9CE5" authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[1136,1325,1457,1485]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">Megalosaurus</taxonomicName>
,
</emphasis>
their summit first appeared at the margin of the jaw internally to the teeth in functional position. In the course of growth and protrusion they excited absorption in the contiguous bone and fang of their predecessors, and continuing to advance from within and beneath (in the lower jaw), as it were, shouldered the latter from the jaw.
<emphasis id="01CDEABDFFBCFFC3C7F4830DFD4B9F5B" box="[729,749,1623,1651]" italics="true" pageId="1" pageNumber="198">A</emphasis>
third tooth in
<taxonomicName id="F4B94D2CFFBCFFC3C6E3830DFB219F5B" authority="Buckland, 1824" authorityName="Buckland" authorityYear="1822" box="[974,1159,1623,1651]" class="Reptilia" family="Megalosauridae" genus="Megalosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="1" pageNumber="198" phylum="Chordata" rank="genus">
<emphasis id="01CDEABDFFBCFFC3C6E3830DFB219F5B" box="[974,1159,1623,1651]" italics="true" pageId="1" pageNumber="198">Megalosaurus</emphasis>
</taxonomicName>
appears to have occupied a position internal to the second one, before the protrusion of this from the jaw.
</paragraph>
<paragraph id="330636AFFFBCFFC3C48383E1FC3A9E30" blockId="1.[388,1549,559,2517]" pageId="1" pageNumber="198">The outer portion of the jaw bone retained in the specimen has an average depth from the alveolar border of 5 inches. Its outer surface is a vertical plane, rounding only near the base.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,347 @@
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<mods:title id="E19261CB5FCE02578D062474F9FA69FE">On the taxonomy of the genus Hymenicoides Kemp, 1917 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), with resurrection of Limnopilos Chuang &amp; Ng, 1991, and descriptions of two new species</mods:title>
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<paragraph id="33066F3E6C372C644485F9B5FDC12A0E" blockId="9.[151,589,1570,1631]" box="[151,589,1570,1596]" pageId="9" pageNumber="26">
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<taxonomicName id="F4B914BD6C372C644485F9B5FE3A2A0E" ID-CoL="6QCKT" box="[151,438,1570,1596]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C372C644485F9B5FE3A2A0E" bold="true" box="[151,438,1570,1596]" italics="true" pageId="9" pageNumber="26">Limnopilos sumatranus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AFE0E576C372C6445AFF9B5FDC12A0E" box="[445,589,1570,1596]" pageId="9" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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(
<figureCitation id="AB8273BB6C372C64448DF9D2FF622A6D" box="[159,238,1605,1631]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="9" pageNumber="26">Figs. 1</figureCitation>
e, f, 7-9)
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C674485F902FE172D48" lastPageId="10" lastPageNumber="27" pageId="9" pageNumber="26" type="materials_examined">
<paragraph id="33066F3E6C372C644485F902FEDE2ACD" blockId="9.[151,1436,1685,1991]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C644485F902FE0A2A9D" bold="true" box="[151,390,1685,1711]" pageId="9" pageNumber="26">Material examined</emphasis>
.
<emphasis id="01CDB32C6C372C644585F902FD852A9D" bold="true" box="[407,521,1685,1711]" pageId="9" pageNumber="26">
<typeStatus id="EC02D19C6C372C644585F902FD852A9D" box="[407,521,1685,1711]" pageId="9" pageNumber="26" type="holotype">Holotype</typeStatus>
</emphasis>
: male, 4.7 ×
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,
<collectionCode id="55A8F7FB6C372C64470EF902FCD02A9D" box="[796,860,1685,1711]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="26">MZB</collectionCode>
Cru 1650, Jambi, Sg. Hitam, near junction into black water reserve, along Batang Hari, Sumatra,
<collectingCountry id="4BAE2FAE6C372C6446FBF92AFCD72AE5" box="[745,859,1725,1751]" name="Indonesia" pageId="9" pageNumber="26">Indonesia</collectingCountry>
(
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,
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), coll. H. H. Tan et al.,
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.
</paragraph>
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</emphasis>
:
<specimenCount id="25BFA4B76C372C64454AF89AFE222B15" box="[344,430,1805,1831]" pageId="9" pageNumber="26" type="male">1 male</specimenCount>
, 5.0 ×
<quantity id="F441C2DB6C372C64461EF89AFDE02B15" box="[524,620,1805,1831]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="9" pageNumber="26" unit="mm" value="5.9">5.9 mm</quantity>
,
<collectionCode id="55A8F7FB6C372C64466FF89AFD3A2B15" box="[637,694,1805,1831]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="26">ZRC</collectionCode>
2007.0114;
<specimenCount id="25BFA4B76C372C644740F89AFC4E2B15" box="[850,962,1805,1831]" pageId="9" pageNumber="26" type="male">21 males</specimenCount>
, 2.1 ×
<quantity id="F441C2DB6C372C644032F89AFBF22B15" box="[1056,1150,1805,1831]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="26" unit="mm" value="2.3">2.3 mm</quantity>
5.1 × 6.0 mm,
<collectionCode id="55A8F7FB6C372C644171F89AFA102B15" box="[1379,1436,1805,1831]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="26">ZRC</collectionCode>
2007.0115;
<specimenCount id="25BFA4B76C372C644531F8A2FE152B7D" box="[291,409,1845,1871]" pageId="9" pageNumber="26" type="female">7 females</specimenCount>
, 2.8 × 3.0 3.7 × 4.0 mm,
<collectionCode id="55A8F7FB6C372C6446F6F8A2FC912B7D" box="[740,797,1845,1871]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="26">ZRC</collectionCode>
2007.0116; 28 ovig., 2.8 × 3.1 4.0 ×
<quantity id="F441C2DB6C372C6440EAF8A2FADA2B7D" box="[1272,1366,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="9" pageNumber="26" unit="mm" value="4.6">4.6 mm</quantity>
,
<collectionCode id="55A8F7FB6C372C644171F8A2FA102B7D" box="[1379,1436,1845,1871]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="26">ZRC</collectionCode>
2007.0117;
<specimenCount id="25BFA4B76C372C644530F8CAFEF32B45" box="[290,383,1885,1911]" pageId="9" pageNumber="26" type="male">3 males</specimenCount>
, 3.5 × 4.0 mm 4.1 ×
<quantity id="F441C2DB6C372C64469FF8CAFD662B45" box="[653,746,1885,1911]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="26" unit="mm" value="4.8">4.8 mm</quantity>
, 1 ovig., 3.8 ×
<quantity id="F441C2DB6C372C64478EF8CAFC752B45" box="[924,1017,1885,1911]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="26" unit="mm" value="4.3">4.3 mm</quantity>
, MNHN-B30394;
<specimenCount id="25BFA4B76C372C6440C6F8CAFAAB2B45" box="[1236,1319,1885,1911]" pageId="9" pageNumber="26" type="male">1 male</specimenCount>
, 4.6 ×
<quantity id="F441C2DB6C372C64416AF8CAFF442BAD" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="9" pageNumber="26" unit="mm" value="5.1">5.1 mm</quantity>
,
<specimenCount id="25BFA4B76C372C6444C1F812FEB02BAD" box="[211,316,1925,1951]" pageId="9" pageNumber="26" type="female">1 female</specimenCount>
, 5.4 ×
<quantity id="F441C2DB6C372C64459EF812FE652BAD" box="[396,489,1925,1951]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="9" pageNumber="26" unit="mm" value="6.3">6.3 mm</quantity>
, MNHN-B30395;
<specimenCount id="25BFA4B76C372C6446D7F812FCAE2BAD" box="[709,802,1925,1951]" pageId="9" pageNumber="26" type="male">3 males</specimenCount>
, 3.3 ×
<quantity id="F441C2DB6C372C644760F812FC402BAD" box="[882,972,1925,1951]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="26" unit="mm" value="3.7">3.7 mm</quantity>
3.8 ×
<quantity id="F441C2DB6C372C64403CF812FB072BAD" box="[1070,1163,1925,1951]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="26" unit="mm" value="4.2">4.2 mm</quantity>
, 1 ovig., 3.5 × 4.0 mm,
<collectionCode id="55A8F7FB6C372C644485F83AFF5B2BF5" box="[151,215,1965,1991]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="26">MZB</collectionCode>
Cru 1651. All
<typeStatus id="EC02D19C6C372C644595F83AFE7B2BF5" box="[391,503,1965,1991]" pageId="9" pageNumber="26" type="paratype">paratypes</typeStatus>
were collected together with
<typeStatus id="EC02D19C6C372C644740F83AFC302BF5" box="[850,956,1965,1991]" pageId="9" pageNumber="26" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="33066F3E6C342C6744D4FF00FE172D48" blockId="10.[151,1437,151,2018]" pageId="10" pageNumber="27">
<emphasis id="01CDB32C6C342C6744D4FF00FE6E2C83" bold="true" box="[198,482,151,177]" pageId="10" pageNumber="27">Comparative material</emphasis>
.
<taxonomicName id="F4B914BD6C342C6745E4FF0FFBA82C80" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[502,1060,152,178]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C342C6745E4FF0FFC8C2C83" box="[502,768,152,177]" italics="true" pageId="10" pageNumber="27">Limnopilos naiyanetri</emphasis>
(
<bibRefCitation id="572812CF6C342C674707FF0FFB972C80" author="Chuang" box="[789,1051,152,178]" pageId="10" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
)
</taxonomicName>
:
<specimenCount id="25BFA4B76C342C674027FF0FFB012C80" box="[1077,1165,152,178]" pageId="10" pageNumber="27" type="male">1 male</specimenCount>
, 5.9 × 6.0 mm,
<collectionCode id="55A8F7FB6C342C674170FF0FFA172C80" box="[1378,1435,152,178]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="27">ZRC</collectionCode>
1993.6520,
<typeStatus id="EC02D19C6C342C674536FF57FE012CE8" box="[292,397,192,218]" pageId="10" pageNumber="27" type="holotype">holotype</typeStatus>
, Mae Nam Nakhon Chaisi, Amphoe Nakhon Chaisi, Changuat Nakhom Pathom,
<collectingCountry id="4BAE2FAE6C342C67414FFF57FF402D30" name="Thailand" pageId="10" pageNumber="27">Thailand</collectingCountry>
, coll. Naunsri, 1998;
<specimenCount id="25BFA4B76C342C6745C2FF7FFDB22D30" box="[464,574,232,258]" pageId="10" pageNumber="27" type="male">19 males</specimenCount>
, 3.1 × 3.6 5.4 ×
<quantity id="F441C2DB6C342C674730FF7FFC0D2D30" box="[802,897,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="10" pageNumber="27" unit="mm" value="6.4">6.4 mm</quantity>
,
<specimenCount id="25BFA4B76C342C67479CFF7FFB9F2D30" box="[910,1043,232,258]" pageId="10" pageNumber="27" type="female">13 females</specimenCount>
, 2.9 × 3.1 4.5 × 5.0 mm,
<collectionCode id="55A8F7FB6C342C674171FF7FFA102D30" box="[1379,1436,232,258]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="27">ZRC</collectionCode>
1991.65216555,
<typeStatus id="EC02D19C6C342C67457EFE87FE532D18" box="[364,479,272,298]" pageId="10" pageNumber="27" type="paratype">paratypes</typeStatus>
, same data as
<typeStatus id="EC02D19C6C342C67469BFE87FD7F2D18" box="[649,755,272,298]" pageId="10" pageNumber="27" type="holotype">holotype</typeStatus>
.
<taxonomicName id="F4B914BD6C342C6746EDFE87FB4B2D18" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[767,1223,272,298]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="27" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C342C6746EDFE87FBB52D1B" box="[767,1081,272,297]" italics="true" pageId="10" pageNumber="27">Limnopilos microrhynchus</emphasis>
(Ng, 1995)
</taxonomicName>
:
<specimenCount id="25BFA4B76C342C6740C1FE87FAAA2D18" box="[1235,1318,272,298]" pageId="10" pageNumber="27" type="male">1 male</specimenCount>
, 2.6 ×
<quantity id="F441C2DB6C342C674165FE87FF442D60" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="10" pageNumber="27" unit="mm" value="2.8">2.8 mm</quantity>
, QM W21466, Bengalon River, east Kalimantan,
<collectingCountry id="4BAE2FAE6C342C674731FEAFFC192D60" box="[803,917,312,338]" name="Indonesia" pageId="10" pageNumber="27">Indonesia</collectingCountry>
(
<geoCoordinate id="568D09F96C342C6747B5FEAFFB8A2D60" box="[935,1030,312,338]" direction="south" orientation="latitude" pageId="10" pageNumber="27" precision="925" value="-0.7">0°42´S</geoCoordinate>
,
<geoCoordinate id="568D09F96C342C674004FEAFFB1E2D60" box="[1046,1170,312,338]" direction="east" orientation="longitude" pageId="10" pageNumber="27" precision="925" value="117.63333">117°38´E</geoCoordinate>
), coll. R. Powell &amp; J. Powell,
<date id="470749FE6C342C6744E5FEF7FE1B2D48" box="[247,407,352,378]" pageId="10" pageNumber="27" value="1996-03-16">16 Mar. 1996</date>
.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C342C6144D4FE10FCAD2B74" lastPageId="12" lastPageNumber="29" pageId="10" pageNumber="27" type="description">
<paragraph id="33066F3E6C342C6744D4FE10FE432ED0" blockId="10.[151,1437,151,2018]" pageId="10" pageNumber="27">
<emphasis id="01CDB32C6C342C6744D4FE10FEDB2D93" bold="true" box="[198,343,391,417]" pageId="10" pageNumber="27">Description</emphasis>
. Carapace (
<figureCitation id="AB8273BB6C342C6745F4FE1FFDA12D90" box="[486,557,392,418]" captionStart="FIGURE 7" captionStartId="11.[151,255,1887,1911]" captionTargetBox="[278,1314,373,1886]" captionTargetId="figure@11.[266,1322,357,1905]" captionTargetPageId="11" captionText="FIGURE 7. Limnopilos sumatranus new species. a, habitus; b, third maxilliped, left; c, male right chela; d f, male right first, third, and fourth ambulatory dactyli; g, female right fourth ambulatory dactylus. a, holotype male (MZB Cru 1650), 4.7 × 5.5 mm; b f, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; g, paratype female (MNHN-B 30395), 5.4 × 6.3 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179202/files/figure.png" pageId="10" pageNumber="27">Fig. 7</figureCitation>
a) circular, CW 1.11.2 times CL (mean 1.13, n = 14); dorsal surface flat, surrounded by circular rim, regions well demarcated by grooves, H-shaped gastric groove continuous with cervical groove, cervical groove branching anteriorly, branches confluent with anterolateral rim. Rostrum vestigial, anterior median part of rim projecting slightly, tip with a few long setae. Lateral margin of carapace lacking tooth or lobe, side wall of posterolateral region slightly expanded laterally, with posterior longitudinal groove along posterolateral rim. Laterally flattened sharp tooth between antennules, completely disconnected from vestigial rostrum; orbit indiscernible, with a sharp conical tooth on outer part of eye, tooth visible from dorsal view. Epistome long, placed dorsally compared to buccal cavern, ischum of third maxilliped partially covering posterior margin.
</paragraph>
<paragraph id="33066F3E6C342C6744D4FD67FC8F2F98" blockId="10.[151,1437,151,2018]" pageId="10" pageNumber="27">
Eyes moderately developed, visible dorsally. Antennule with long coxa, about twice length of basis. Third maxilliped (
<figureCitation id="AB8273BB6C342C674536FC8FFEE52F00" box="[292,361,792,818]" captionStart="FIGURE 7" captionStartId="11.[151,255,1887,1911]" captionTargetBox="[278,1314,373,1886]" captionTargetId="figure@11.[266,1322,357,1905]" captionTargetPageId="11" captionText="FIGURE 7. Limnopilos sumatranus new species. a, habitus; b, third maxilliped, left; c, male right chela; d f, male right first, third, and fourth ambulatory dactyli; g, female right fourth ambulatory dactylus. a, holotype male (MZB Cru 1650), 4.7 × 5.5 mm; b f, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; g, paratype female (MNHN-B 30395), 5.4 × 6.3 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179202/files/figure.png" pageId="10" pageNumber="27">Fig. 7</figureCitation>
b) narrow, covering about one-quarter of buccal cavern; merus long, about one-half times of ichium, narrower than ischium; palp long, propodus slightly longer than half length of dactylus, dactylus longer than merus, tip of dactylus almost reaching proximal end of ischium; exopod short, reaching about proximal two-third of merus, with distinct flagellum.
</paragraph>
<paragraph id="33066F3E6C342C6744D4FC2FFABF2878" blockId="10.[151,1437,151,2018]" pageId="10" pageNumber="27">
Male with relatively wide abdominal cavity (
<figureCitation id="AB8273BB6C342C6746C7FC2FFC9B2FE0" box="[725,791,952,978]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="10" pageNumber="27">Fig. 1</figureCitation>
e), sternal button present on just posterior to imaginary line joining inner ends of sutures between sternites 5 and 6. Female thoracic sternite 3 separated from sternite 4 by posteriorly convex rim, sternites 48 medially fused, vulva on imaginary line joining inner ends between sternite 6 on medial fused plate of thoracic sternum, vulva with longitudinally elliptical basal mount.
</paragraph>
<paragraph id="33066F3E6C342C6744D4FBCFFF4029B8" blockId="10.[151,1437,151,2018]" pageId="10" pageNumber="27">
Cheliped symmetrical, more massive in males; cheliped of male with short merus, as long as carpus, ventral outer margin lined with long plumose setae, with subdistal tooth, ventral inner margin sparsely lined with low granules, inner surface of merus to carpus lined with long stiff setae, outer surface of carpus rounded, not strongly elevated, outer margin with small subdistal tooth; chela (
<figureCitation id="AB8273BB6C342C674780FB47FC5B28D8" box="[914,983,1232,1258]" captionStart="FIGURE 7" captionStartId="11.[151,255,1887,1911]" captionTargetBox="[278,1314,373,1886]" captionTargetId="figure@11.[266,1322,357,1905]" captionTargetPageId="11" captionText="FIGURE 7. Limnopilos sumatranus new species. a, habitus; b, third maxilliped, left; c, male right chela; d f, male right first, third, and fourth ambulatory dactyli; g, female right fourth ambulatory dactylus. a, holotype male (MZB Cru 1650), 4.7 × 5.5 mm; b f, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; g, paratype female (MNHN-B 30395), 5.4 × 6.3 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179202/files/figure.png" pageId="10" pageNumber="27">Fig. 7</figureCitation>
c) with rounded palm, covered densely with long plumose setae from distal part of upper, lower margin of palm to both fingers; fingers with ovoid gape when closed; immovable finger with 3 teeth on proximal two-thirds, with subdistal tooth connected to tip of finger by thin lobe, forming hoof; movable finger similar to immovable finger, except for tooth on proximal part.
</paragraph>
<paragraph id="33066F3E6C342C6744D4FA0FFBE02A48" blockId="10.[151,1437,151,2018]" pageId="10" pageNumber="27">
Ambulatory legs (
<figureCitation id="AB8273BB6C342C674588FA0FFE6C2980" box="[410,480,1432,1458]" captionStart="FIGURE 7" captionStartId="11.[151,255,1887,1911]" captionTargetBox="[278,1314,373,1886]" captionTargetId="figure@11.[266,1322,357,1905]" captionTargetPageId="11" captionText="FIGURE 7. Limnopilos sumatranus new species. a, habitus; b, third maxilliped, left; c, male right chela; d f, male right first, third, and fourth ambulatory dactyli; g, female right fourth ambulatory dactylus. a, holotype male (MZB Cru 1650), 4.7 × 5.5 mm; b f, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; g, paratype female (MNHN-B 30395), 5.4 × 6.3 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179202/files/figure.png" pageId="10" pageNumber="27">Fig. 7</figureCitation>
a, dg) slender, long; second, third legs subequal, longer than others, fourth shortest, inner margin of legs fringed densely with long plumose setae, outer margin lined with short curly setae; meri slightly longer than respective propodi, distal anterior angle ending in conical tooth; dactyli terminating in sharp tooth, first subdistal tooth of inner margin stronger than terminal tooth, 68 teeth (n = 7) of inner margin arranged throughout in fourth leg, 69 teeth only on distal half in second and third legs (n = 6), 24 teeth limited to distal third in first leg (n = 5), teeth of males larger than those of females.
</paragraph>
<paragraph id="33066F3E6C342C6644D4F91FFC1D2D30" blockId="10.[151,1437,151,2018]" lastBlockId="11.[151,1436,152,258]" lastPageId="11" lastPageNumber="28" pageId="10" pageNumber="27">
Male abdomen-pleotelson (
<figureCitation id="AB8273BB6C342C674618F91FFDDC2A90" box="[522,592,1672,1698]" captionStart="FIGURE 8" captionStartId="12.[151,255,1467,1491]" captionTargetBox="[260,1306,186,1428]" captionTargetId="figure@12.[260,1326,172,1446]" captionTargetPageId="12" captionText="FIGURE 8. Limnopilos sumatranus new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1; d, left G 2. a, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; b, paratype female (MNHN-B 30395), 5.4 × 6.3 mm; c, d, holotype male (MZB Cru 1650), 4.7 × 5.5 mm. Scales, 1 mm." httpUri="https://zenodo.org/record/179203/files/figure.png" pageId="10" pageNumber="27">Fig. 8</figureCitation>
a) 6-segmented; first segment of male abdomen thick, hard, distal margin widely concave; second to fifth segments with distal margin concave medially, third segment widest; pleotelson trilobed, lateral lobes small, low; 2 small concavities on posterior margin, distal margin thickened, forming socket for sternal button (
<figureCitation id="AB8273BB6C342C674621F897FDFB2B28" box="[563,631,1792,1818]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="10" pageNumber="27">Fig. 1</figureCitation>
f) on inner surface. G1 (
<figureCitation id="AB8273BB6C342C67478DF897FC6A2B28" box="[927,998,1792,1818]" captionStart="FIGURE 8" captionStartId="12.[151,255,1467,1491]" captionTargetBox="[260,1306,186,1428]" captionTargetId="figure@12.[260,1326,172,1446]" captionTargetPageId="12" captionText="FIGURE 8. Limnopilos sumatranus new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1; d, left G 2. a, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; b, paratype female (MNHN-B 30395), 5.4 × 6.3 mm; c, d, holotype male (MZB Cru 1650), 4.7 × 5.5 mm. Scales, 1 mm." httpUri="https://zenodo.org/record/179203/files/figure.png" pageId="10" pageNumber="27">Fig. 8</figureCitation>
c) stout, strongly bent on distal twofifths, ventral outer surface longitudinally lined with long, stiff, distally plumose setae; distal inner end with 2 sharp processes, sub-inner process well separated from, larger than inner process, directed slightly lower than that of inner process, distal outer angle swollen, covered by tiny granule, outer ventral apex bat-like. G2 (
<figureCitation id="AB8273BB6C342C67417EF8EFFF292B88" captionStart="FIGURE 8" captionStartId="12.[151,255,1467,1491]" captionTargetBox="[260,1306,186,1428]" captionTargetId="figure@12.[260,1326,172,1446]" captionTargetPageId="12" captionText="FIGURE 8. Limnopilos sumatranus new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1; d, left G 2. a, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; b, paratype female (MNHN-B 30395), 5.4 × 6.3 mm; c, d, holotype male (MZB Cru 1650), 4.7 × 5.5 mm. Scales, 1 mm." httpUri="https://zenodo.org/record/179203/files/figure.png" pageId="10" pageNumber="27">Fig. 8</figureCitation>
d) short, base not strongly swollen. Female abdomen-pleotelson (
<figureCitation id="AB8273BB6C342C6747B6F837FC652B88" box="[932,1001,1952,1978]" captionStart="FIGURE 8" captionStartId="12.[151,255,1467,1491]" captionTargetBox="[260,1306,186,1428]" captionTargetId="figure@12.[260,1326,172,1446]" captionTargetPageId="12" captionText="FIGURE 8. Limnopilos sumatranus new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1; d, left G 2. a, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; b, paratype female (MNHN-B 30395), 5.4 × 6.3 mm; c, d, holotype male (MZB Cru 1650), 4.7 × 5.5 mm. Scales, 1 mm." httpUri="https://zenodo.org/record/179203/files/figure.png" pageId="10" pageNumber="27">Fig. 8</figureCitation>
b) demarcated to 6 segments, boundary between first and second segments movable, more or less fixed in other boundaries; second to fifth segments with corneous wide ridge medially, that of segment 6 only on proximal half, segment 6 longest, no suture separating pleotelson; pleopods (
<figureCitation id="AB8273BB6C352C664662FF57FD342CE8" box="[624,696,192,218]" captionStart="FIGURE 9" captionStartId="13.[151,255,1929,1953]" captionTargetBox="[151,1436,587,1902]" captionTargetId="figure@13.[151,1436,587,1903]" captionTargetPageId="13" captionText="FIGURE 9. Limnopilos sumatranus new species. a, b, right second and third pleopods; c, d, left fourth and fifth pleopods; e, exopods. Paratype, ovigerous female (ZRC 2007.0117), 4.0 × 4.6 mm." httpUri="https://zenodo.org/record/179204/files/figure.png" pageId="11" pageNumber="28">Fig. 9</figureCitation>
) on second to fifth segments, long, biramous from near base, developing from distal outer angle of inner side of each segment.
</paragraph>
<caption id="67C63FB66C352C664485F8C8FEDA2BE4" httpUri="https://zenodo.org/record/179202/files/figure.png" pageId="11" pageNumber="28" targetBox="[278,1314,373,1886]" targetPageId="11">
<paragraph id="33066F3E6C352C664485F8C8FEDA2BE4" blockId="11.[151,1437,1887,2006]" pageId="11" pageNumber="28">
<emphasis id="01CDB32C6C352C664485F8C8FE952B45" bold="true" box="[151,281,1887,1911]" pageId="11" pageNumber="28">FIGURE 7.</emphasis>
<taxonomicName id="F4B914BD6C352C664532F8C8FD972B44" box="[288,539,1887,1910]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="28" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C352C664532F8C8FD972B44" box="[288,539,1887,1910]" italics="true" pageId="11" pageNumber="28">Limnopilos sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C352C664630F8C8FD282B45" bold="true" box="[546,676,1887,1911]" pageId="11" pageNumber="28">
<taxonomicNameLabel id="1AFE0E576C352C664630F8C8FD282B45" box="[546,676,1887,1911]" pageId="11" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
. a, habitus; b, third maxilliped, left; c, male right chela; df, male right first, third, and fourth ambulatory dactyli; g, female right fourth ambulatory dactylus. a, holotype male (MZB Cru 1650), 4.7 × 5.5 mm; bf, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; g, paratype female (MNHN-B30395), 5.4 × 6.3 mm. Scale bars, 1 mm.
</paragraph>
</caption>
<caption id="67C63FB66C322C614485FA2CFCCA2A21" httpUri="https://zenodo.org/record/179203/files/figure.png" pageId="12" pageNumber="29" targetBox="[260,1306,186,1428]" targetPageId="12">
<paragraph id="33066F3E6C322C614485FA2CFCCA2A21" blockId="12.[151,1436,1467,1555]" pageId="12" pageNumber="29">
<emphasis id="01CDB32C6C322C614485FA2CFE9029E1" bold="true" box="[151,284,1467,1491]" pageId="12" pageNumber="29">FIGURE 8.</emphasis>
<taxonomicName id="F4B914BD6C322C614537FA2BFDAF29E1" box="[293,547,1468,1491]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C322C614537FA2BFDAF29E1" box="[293,547,1468,1491]" italics="true" pageId="12" pageNumber="29">Limnopilos sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C322C61463EFA2CFD3D29E1" bold="true" box="[556,689,1467,1491]" pageId="12" pageNumber="29">
<taxonomicNameLabel id="1AFE0E576C322C61463EFA2CFD3D29E1" box="[556,689,1467,1491]" pageId="12" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G1; d, left G2. a, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; b, paratype female (MNHN-B30395), 5.4 × 6.3 mm; c, d, holotype male (MZB Cru 1650), 4.7 × 5.5 mm. Scales, 1 mm.
</paragraph>
</caption>
<paragraph id="33066F3E6C322C6144D4F9F3FE862AFC" blockId="12.[151,1436,1636,2022]" pageId="12" pageNumber="29">
Eggs spherical, small, diameter
<quantity id="F441C2DB6C322C61462EF9F3FD792A4C" box="[572,757,1636,1662]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="12" pageNumber="29" unit="mm" value="0.33" valueMax="0.34" valueMin="0.32">0.32 0.34 mm</quantity>
[mean = 0.33, n = 10 from two females (2.8 × 3.1 and 4.0 ×
<quantity id="F441C2DB6C322C6144A3F91BFE982A94" box="[177,276,1676,1702]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="12" pageNumber="29" unit="mm" value="4.6">4.6 mm</quantity>
)], attached on endopods of pleopods, exopods growing inwards, partially covering dome-shaped abdomen.
</paragraph>
<paragraph id="33066F3E6C322C6144D4F94CFCAD2B74" blockId="12.[151,1436,1636,2022]" pageId="12" pageNumber="29">
<emphasis id="01CDB32C6C322C6144D4F94CFE772AC7" bold="true" box="[198,507,1755,1781]" pageId="12" pageNumber="29">Habitat and distribution</emphasis>
. The specimens were all collected from among the roots of a mass of floating vegetation near the centre of the stream, from which catfishes (
<taxonomicName id="F4B914BD6C322C614791F893FC712B2F" box="[899,1021,1796,1821]" class="Actinopterygii" family="Bagridae" genus="Leiocassis" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="29" phylum="Chordata" rank="genus">
<emphasis id="01CDB32C6C322C614791F893FC712B2F" box="[899,1021,1796,1821]" italics="true" pageId="12" pageNumber="29">Leiocassis</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="F4B914BD6C322C614024F893FB2E2B2C" box="[1078,1186,1796,1822]" class="Actinopterygii" family="Bagridae" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="29" phylum="Chordata" rank="family">Bagridae</taxonomicName>
) were also collected. The tannin-coloured acidic waters were gently flowing.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C322C6144D4F8C3FBC72B5C" box="[198,1099,1876,1902]" pageId="12" pageNumber="29" type="etymology">
<paragraph id="33066F3E6C322C6144D4F8C3FBC72B5C" blockId="12.[151,1436,1636,2022]" box="[198,1099,1876,1902]" pageId="12" pageNumber="29">
<emphasis id="01CDB32C6C322C6144D4F8C3FEC72B5C" bold="true" box="[198,331,1876,1902]" pageId="12" pageNumber="29">Etymology</emphasis>
. The species is named after the
<typeStatus id="EC02D19C6C322C6146D1F8C3FD7A2B5C" box="[707,758,1876,1902]" pageId="12" pageNumber="29">type</typeStatus>
locality, Sumatra,
<collectingCountry id="4BAE2FAE6C322C6147C0F8C3FBC42B5C" box="[978,1096,1876,1902]" name="Indonesia" pageId="12" pageNumber="29">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C322C6044D4F8EBFAAF2E28" lastPageId="13" lastPageNumber="30" pageId="12" pageNumber="29" type="discussion">
<paragraph id="33066F3E6C322C6044D4F8EBFD2E2DF8" blockId="12.[151,1436,1636,2022]" lastBlockId="13.[151,1436,152,538]" lastPageId="13" lastPageNumber="30" pageId="12" pageNumber="29">
<emphasis id="01CDB32C6C322C6144D4F8EBFEB42BA4" bold="true" box="[198,312,1916,1942]" pageId="12" pageNumber="29">Remarks</emphasis>
.
<taxonomicName id="F4B914BD6C322C614555F8EBFDD72BA7" box="[327,603,1916,1941]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C322C614555F8EBFDD72BA7" box="[327,603,1916,1941]" italics="true" pageId="12" pageNumber="29">Limnopilos sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C322C614670F8EBFD7E2BA4" bold="true" box="[610,754,1916,1942]" pageId="12" pageNumber="29">
<taxonomicNameLabel id="1AFE0E576C322C614670F8EBFD7E2BA4" box="[610,754,1916,1942]" pageId="12" pageNumber="29" rank="species">new species</taxonomicNameLabel>
</emphasis>
, is very close to
<taxonomicName id="F4B914BD6C322C6147ABF8EBFB702BA4" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[953,1276,1916,1942]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C322C6147ABF8EBFB092BA7" box="[953,1157,1916,1941]" italics="true" pageId="12" pageNumber="29">L. microrhynchus</emphasis>
Ng, 1995
</taxonomicName>
, especially in having a vestigial rostrum. The main difference between the two species is the location of the sub-inner process of the distal end of the G1. It is more medially placed and directed more laterally in
<taxonomicName id="F4B914BD6C322C61408CF85BFACA2BD7" box="[1182,1350,1996,2021]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="29" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="01CDB32C6C322C61408CF85BFACA2BD7" box="[1182,1350,1996,2021]" italics="true" pageId="12" pageNumber="29">L. sumatranus</emphasis>
</taxonomicName>
than in
<taxonomicName id="F4B914BD6C332C604485FF0FFEEF2C83" box="[151,355,152,177]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C332C604485FF0FFEEF2C83" box="[151,355,152,177]" italics="true" pageId="13" pageNumber="30">L. microrhynchus</emphasis>
</taxonomicName>
, where it is closer to the inner process and directed more anteriorly (
<figureCitation id="AB8273BB6C332C604093FF0FFB4A2C80" box="[1153,1222,152,178]" captionStart="FIGURE 8" captionStartId="12.[151,255,1467,1491]" captionTargetBox="[260,1306,186,1428]" captionTargetId="figure@12.[260,1326,172,1446]" captionTargetPageId="12" captionText="FIGURE 8. Limnopilos sumatranus new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1; d, left G 2. a, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; b, paratype female (MNHN-B 30395), 5.4 × 6.3 mm; c, d, holotype male (MZB Cru 1650), 4.7 × 5.5 mm. Scales, 1 mm." httpUri="https://zenodo.org/record/179203/files/figure.png" pageId="13" pageNumber="30">Fig. 8</figureCitation>
c). This difference is constant for the good series of specimens examined and evident even when similar sized specimens are compared (
<taxonomicName id="F4B914BD6C332C60450EFF7FFE482D33" box="[284,452,232,257]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="01CDB32C6C332C60450EFF7FFE482D33" box="[284,452,232,257]" italics="true" pageId="13" pageNumber="30">L. sumatranus</emphasis>
</taxonomicName>
:
<specimenCount id="25BFA4B76C332C6045C6FF7FFDBE2D30" box="[468,562,232,258]" pageId="13" pageNumber="30" type="male">3 males</specimenCount>
, ZRC 2007.0115, 2.2 × 2.32.9 ×
<quantity id="F441C2DB6C332C6047D4FF7FFBA82D30" box="[966,1060,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="30" unit="mm" value="3.2">3.2 mm</quantity>
;
<taxonomicName id="F4B914BD6C332C604022FF7FFB722D33" box="[1072,1278,232,257]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C332C604022FF7FFB722D33" box="[1072,1278,232,257]" italics="true" pageId="13" pageNumber="30">L. microrhynchus</emphasis>
</taxonomicName>
:
<specimenCount id="25BFA4B76C332C60411CFF7FFAEC2D30" box="[1294,1376,232,258]" pageId="13" pageNumber="30" type="male">1 male</specimenCount>
, QM W21466, 2.6 ×
<quantity id="F441C2DB6C332C604540FE87FE3E2D18" box="[338,434,272,298]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="30" unit="mm" value="2.8">2.8 mm</quantity>
). In addition, small specimens of
<taxonomicName id="F4B914BD6C332C604750FE87FC672D1B" box="[834,1003,272,297]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="01CDB32C6C332C604750FE87FC672D1B" box="[834,1003,272,297]" italics="true" pageId="13" pageNumber="30">L. sumatranus</emphasis>
</taxonomicName>
have a distinct sharp subdistal tooth on the ventral outer margin of the cheliped merus (indiscernible in
<taxonomicName id="F4B914BD6C332C6047D8FEAFFB162D63" box="[970,1178,312,337]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C332C6047D8FEAFFB162D63" box="[970,1178,312,337]" italics="true" pageId="13" pageNumber="30">L. microrhynchus</emphasis>
</taxonomicName>
) and an anterolateral margin which is less divergent posteriorly than in
<taxonomicName id="F4B914BD6C332C6046F2FEF7FC212D4B" box="[736,941,352,377]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C332C6046F2FEF7FC212D4B" box="[736,941,352,377]" italics="true" pageId="13" pageNumber="30">L. microrhynchus</emphasis>
</taxonomicName>
. Large individuals of
<taxonomicName id="F4B914BD6C332C6040BDFEF7FADB2D4B" box="[1199,1367,352,377]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="01CDB32C6C332C6040BDFEF7FADB2D4B" box="[1199,1367,352,377]" italics="true" pageId="13" pageNumber="30">L. sumatranus</emphasis>
</taxonomicName>
show a similar divergence of the anterolateral margin as in
<taxonomicName id="F4B914BD6C332C60470FFE1FFC602D93" box="[797,1004,392,417]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C332C60470FFE1FFC602D93" box="[797,1004,392,417]" italics="true" pageId="13" pageNumber="30">L. microrhynchus</emphasis>
</taxonomicName>
, but their difference is still obvious when similar sized specimens are compared.
</paragraph>
<paragraph id="33066F3E6C332C6044D4FE4FFAAF2E28" blockId="13.[151,1436,152,538]" pageId="13" pageNumber="30">
<taxonomicName id="F4B914BD6C332C6044D4FE4FFE732DC3" box="[198,511,472,497]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="30" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C332C6044D4FE4FFE732DC3" box="[198,511,472,497]" italics="true" pageId="13" pageNumber="30">Limnopilos microrhynchus</emphasis>
</taxonomicName>
was described from eastern Sabah (Ng, 1995), and the present specimens from eastern Kalimantan, extend its range further southwards, as well as into
<collectingCountry id="4BAE2FAE6C332C6047F0FD97FBD82E28" box="[994,1108,512,538]" name="Indonesia" pageId="13" pageNumber="30">Indonesia</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,152 @@
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<paragraph id="33066F3E6C372C644485FE68FD912E2B" blockId="9.[151,541,511,537]" box="[151,541,511,537]" pageId="9" pageNumber="26">
<heading id="684ED8526C372C644485FE68FD912E2B" bold="true" box="[151,541,511,537]" fontSize="11" level="1" pageId="9" pageNumber="26" reason="1">
<taxonomicName id="F4B914BD6C372C644485FE68FD912E2B" ID-CoL="7PMTG" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[151,541,511,537]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C644485FE68FD912E2B" bold="true" box="[151,541,511,537]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C644485FE68FEAC2E2B" bold="true" box="[151,288,511,537]" italics="true" pageId="9" pageNumber="26">Limnopilos</emphasis>
<bibRefCitation id="572812CF6C372C644534FE68FD912E2B" author="Chuang" box="[294,541,511,537]" pageId="9" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C644485FDDDFB0D2EB3" pageId="9" pageNumber="26" type="reference_group">
<paragraph id="33066F3E6C372C644485FDDDFDA72E53" blockId="9.[151,1153,586,641]" box="[151,555,586,609]" pageId="9" pageNumber="26">
<treatmentCitationGroup id="13A948106C372C644485FDDDFDA72E53" box="[151,555,586,609]" pageId="9" pageNumber="26">
<taxonomicName id="F4B914BD6C372C644485FDDDFDAB2E53" ID-CoL="7PMTG" authority="Chuang &amp; Ng, 1991: 363" authorityName="Chuang &amp; Ng" authorityPageNumber="363" authorityYear="1991" box="[151,551,586,609]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C644485FDDDFE832E53" box="[151,271,586,609]" italics="true" pageId="9" pageNumber="26">Limnopilos</emphasis>
<treatmentCitation id="B218492F6C372C644507FDDDFDAB2E53" author="Chuang" box="[277,551,586,609]" page="363" pageId="9" pageNumber="26" year="1991">
<bibRefCitation id="572812CF6C372C644507FDDDFE7D2E53" author="Chuang" box="[277,497,586,609]" pageId="9" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
: 363
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</taxonomicName>
.
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</paragraph>
<paragraph id="33066F3E6C372C644485FDFDFB0D2EB3" blockId="9.[151,1153,586,641]" box="[151,1153,618,641]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C644485FDFDFECA2EB3" box="[151,326,618,641]" italics="true" pageId="9" pageNumber="26">Hymenicoides—</emphasis>
<bibRefCitation id="572812CF6C372C64455EFDFDFDA42EB3" author="Ng" box="[332,552,618,641]" pageId="9" pageNumber="31" refString="Ng, P. K. L. &amp; Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement, 3, 1 - 82." type="journal article" year="1996">Ng &amp; Chuang, 1996</bibRefCitation>
: 50 (part); Guinot &amp; Richer de
<bibRefCitation id="572812CF6C372C644765FDFDFB892EB3" author="Forges" box="[887,1029,618,641]" pageId="9" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
: 460 (part).
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C644485FD25FEED2EC6" pageId="9" pageNumber="26" type="materials_examined">
<paragraph id="33066F3E6C372C644485FD25FEED2EC6" blockId="9.[151,1437,690,1476]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C644485FD25FEBF2EFE" bold="true" box="[151,307,690,716]" pageId="9" pageNumber="26">
<typeStatus id="EC02D19C6C372C644485FD25FF582EFE" box="[151,212,690,716]" pageId="9" pageNumber="26">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="F4B914BD6C372C644556FD25FCCB2EFE" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[324,839,690,716]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C372C644556FD25FDC62EF9" box="[324,586,690,715]" italics="true" pageId="9" pageNumber="26">Limnopilos naiyanetri</emphasis>
<bibRefCitation id="572812CF6C372C644640FD25FCCB2EFE" author="Chuang" box="[594,839,690,716]" pageId="9" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
</taxonomicName>
, by original designation and monotypy; gender of genus masculine.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C6444D4FC95FC6028E6" pageId="9" pageNumber="26" type="diagnosis">
<paragraph id="33066F3E6C372C6444D4FC95FC6028E6" blockId="9.[151,1437,690,1476]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C6444D4FC95FEB32F2E" bold="true" box="[198,319,770,796]" pageId="9" pageNumber="26">Diagnosis</emphasis>
. Carapace circular, pilose, dorsal surface concave; grooves distinct; rostrum absent or very weak; antenna with proximal portion of basal article posterior to inner section of proximal portion of eye; eyes, antennae, antennules visible dorsally; third maxillipeds narrow, not covering more than three-quarters of mouth field when closed, merus rectangular, dactylus styliform, approximately twice length of propodus; male chelae relatively stout, outer surface evenly convex, partially covered by dense setae, without tubercle. Female vulva placed on imaginary line joining inner ends of sutures between sternites 5, 6 on medial fused plate of thoracic sternum, vulva with basal mount. Male abdomen-pleotelson 6 segmented, pleotelson slightly trilobed, inner surface thickened distally, forming socket for sternal button (
<figureCitation id="AB8273BB6C372C64403AFB8DFBE12806" box="[1064,1133,1050,1076]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="9" pageNumber="26">Fig. 1</figureCitation>
c, d). G1 stout, bent outwards medially; distal part with distal inner processes, tuberculate distal outer angle. Female abdomen-pleotelson with six distinctly demarcated segments, boundary between first and second (
<taxonomicName id="F4B914BD6C372C6440AFFBFCFAE028B6" box="[1213,1388,1131,1156]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="sumatranus">
<emphasis id="01CDB32C6C372C6440AFFBFCFAE028B6" box="[1213,1388,1131,1156]" italics="true" pageId="9" pageNumber="26">L. sumatranus</emphasis>
</taxonomicName>
) or between second and third segments (
<taxonomicName id="F4B914BD6C372C644658FB04FD6E289E" box="[586,738,1171,1196]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C372C644658FB04FD6E289E" box="[586,738,1171,1196]" italics="true" pageId="9" pageNumber="26">L. naiyanetri</emphasis>
</taxonomicName>
) movable; long, biramous pleopods on second to fifth segments, developed from distal outer end of inner surface of each segment.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C6444D4FB75FC8B29F6" pageId="9" pageNumber="26" type="discussion">
<paragraph id="33066F3E6C372C6444D4FB75FEAE2946" blockId="9.[151,1437,690,1476]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C6444D4FB75FEB428CE" bold="true" box="[198,312,1250,1276]" pageId="9" pageNumber="26">Remarks</emphasis>
. Several characters clearly distinguish
<taxonomicName id="F4B914BD6C372C64473DFB74FC3F28CE" box="[815,947,1251,1276]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C64473DFB74FC3F28CE" box="[815,947,1251,1276]" italics="true" pageId="9" pageNumber="26">Limnopilos</emphasis>
</taxonomicName>
from
<taxonomicName id="F4B914BD6C372C64401EFB74FB3E28CE" box="[1036,1202,1251,1276]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C64401EFB74FB3E28CE" box="[1036,1202,1251,1276]" italics="true" pageId="9" pageNumber="26">Hymenicoides</emphasis>
</taxonomicName>
(see Remarks for
<taxonomicName id="F4B914BD6C372C644485FA9CFEB12916" box="[151,317,1291,1316]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C644485FA9CFEB12916" box="[151,317,1291,1316]" italics="true" pageId="9" pageNumber="26">Hymenicoides</emphasis>
</taxonomicName>
above). In addition to
<taxonomicName id="F4B914BD6C372C644672FA9CFD702916" box="[608,764,1291,1316]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C372C644672FA9CFD702916" box="[608,764,1291,1316]" italics="true" pageId="9" pageNumber="26">L. naiyanetri</emphasis>
</taxonomicName>
, the
<typeStatus id="EC02D19C6C372C64472DFA9DFCFE2916" box="[831,882,1290,1316]" pageId="9" pageNumber="26">type</typeStatus>
species,
<taxonomicName id="F4B914BD6C372C6447F4FA9CFAB12916" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[998,1341,1290,1316]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C372C6447F4FA9CFB312916" box="[998,1213,1291,1316]" italics="true" pageId="9" pageNumber="26">H. microrhynchus</emphasis>
Ng, 1995
</taxonomicName>
, and
<taxonomicName id="F4B914BD6C372C644196FA9CFEAC297E" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C372C644196FA9CFEAC297E" italics="true" pageId="9" pageNumber="26">L. sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C372C644538FAA5FE30297E" bold="true" box="[298,444,1330,1356]" pageId="9" pageNumber="26">
<taxonomicNameLabel id="1AFE0E576C372C644538FAA5FE30297E" box="[298,444,1330,1356]" pageId="9" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
, all possess the diagnostic characters of the genus and are therefore transferred to
<taxonomicName id="F4B914BD6C372C644485FACCFE972946" box="[151,283,1371,1396]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C372C644485FACCFE972946" box="[151,283,1371,1396]" italics="true" pageId="9" pageNumber="26">Limnopilos</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="33066F3E6C372C6444D4FA15FC8B29F6" blockId="9.[151,1437,690,1476]" pageId="9" pageNumber="26">
In the case of
<taxonomicName id="F4B914BD6C372C644579FA14FDB429AE" box="[363,568,1411,1436]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="microrhynchus">
<emphasis id="01CDB32C6C372C644579FA14FDB429AE" box="[363,568,1411,1436]" italics="true" pageId="9" pageNumber="26">L. microrhynchus</emphasis>
</taxonomicName>
, only male specimens are known thus far, and the female characters diagnosed above are therefore not known for this species.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,307 @@
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<treatment id="BB10DE286C3A2C644485FA50FBE12D90" ID-DOI="http://doi.org/10.5281/zenodo.6251674" ID-GBIF-Taxon="119375998" ID-Zenodo-Dep="6251674" LSID="urn:lsid:plazi:treatment:BB10DE286C3A2C644485FA50FBE12D90" httpUri="http://treatment.plazi.org/id/BB10DE286C3A2C644485FA50FBE12D90" lastPageId="9" lastPageNumber="26" pageId="4" pageNumber="21">
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<paragraph id="33066F3E6C3A2C694485FA50FDB029D3" blockId="4.[151,572,1479,1540]" box="[151,572,1479,1505]" pageId="4" pageNumber="21">
<heading id="684ED8526C3A2C694485FA50FDB029D3" bold="true" box="[151,572,1479,1505]" fontSize="11" level="1" pageId="4" pageNumber="21" reason="1">
<emphasis id="01CDB32C6C3A2C694485FA50FDB029D3" bold="true" box="[151,572,1479,1505]" pageId="4" pageNumber="21">
<taxonomicName id="F4B914BD6C3A2C694485FA50FE2929D3" ID-CoL="3NJ2J" box="[151,421,1479,1505]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="21" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C3A2C694485FA50FE2929D3" bold="true" box="[151,421,1479,1505]" italics="true" pageId="4" pageNumber="21">Hymenicoides robertsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1AFE0E576C3A2C6945BFFA50FDB029D3" box="[429,572,1479,1505]" pageId="4" pageNumber="21" rank="species">new species</taxonomicNameLabel>
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<paragraph id="33066F3E6C3A2C694485FA7DFED32A36" blockId="4.[151,572,1479,1540]" box="[151,351,1514,1540]" pageId="4" pageNumber="21">
(
<figureCitation id="AB8273BB6C3A2C69448DFA7DFF622A36" box="[159,238,1514,1540]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="4" pageNumber="21">Figs. 1</figureCitation>
a, b, 36)
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3A2C694485F9ADFC182B76" pageId="4" pageNumber="21" type="materials_examined">
<paragraph id="33066F3E6C3A2C694485F9ADFC232A4E" blockId="4.[151,1437,1594,2020]" pageId="4" pageNumber="21">
<emphasis id="01CDB32C6C3A2C694485F9ADFE0F2A66" bold="true" box="[151,387,1594,1620]" pageId="4" pageNumber="21">Material examined</emphasis>
.
<emphasis id="01CDB32C6C3A2C694581F9ADFD892A66" bold="true" box="[403,517,1594,1620]" pageId="4" pageNumber="21">
<typeStatus id="EC02D19C6C3A2C694581F9ADFD892A66" box="[403,517,1594,1620]" pageId="4" pageNumber="21" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="25BFA4B76C3A2C694606F9ADFDEB2A66" box="[532,615,1594,1620]" pageId="4" pageNumber="21" type="male">1 male</specimenCount>
, 4.9 × 6.0 mm,
<collectionCode id="55A8F7FB6C3A2C694732F9ADFCD52A66" box="[800,857,1594,1620]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="21">ZRC</collectionCode>
2007.0108, Kyaukdaw market on lower Kaladan River (tidal), Rakhine,
<collectingCountry id="4BAE2FAE6C3A2C6945B7F9F5FD952A4E" box="[421,537,1634,1660]" name="Myanmar" pageId="4" pageNumber="21">Myanmar</collectingCountry>
, coll. T. R. Roberts,
<date id="470749FE6C3A2C694718F9F5FC262A4E" box="[778,938,1634,1660]" pageId="4" pageNumber="21" value="2004-03-26">26 Mar. 2004</date>
.
</paragraph>
<paragraph id="33066F3E6C3A2C6944D4F91DFC182B76" blockId="4.[151,1437,1594,2020]" pageId="4" pageNumber="21">
<emphasis id="01CDB32C6C3A2C6944D4F91DFEC12A96" bold="true" box="[198,333,1674,1700]" pageId="4" pageNumber="21">
<typeStatus id="EC02D19C6C3A2C6944D4F91DFEC42A96" box="[198,328,1674,1700]" pageId="4" pageNumber="21" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="25BFA4B76C3A2C694546F91DFE2B2A96" box="[340,423,1674,1700]" pageId="4" pageNumber="21" type="male">1 male</specimenCount>
, 5.0 ×
<quantity id="F441C2DB6C3A2C6945E5F91DFDD82A96" box="[503,596,1674,1700]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="21" unit="mm" value="6.2">6.2 mm</quantity>
,
<collectionCode id="55A8F7FB6C3A2C69464DF91DFD142A96" box="[607,664,1674,1700]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="21">ZRC</collectionCode>
2007.0109;
<specimenCount id="25BFA4B76C3A2C69473EF91DFC142A96" box="[812,920,1674,1700]" pageId="4" pageNumber="21" type="male">42 males</specimenCount>
, 3.0 × 3.3 5.2 ×
<quantity id="F441C2DB6C3A2C69407DF91DFB402A96" box="[1135,1228,1674,1700]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="21" unit="mm" value="6.2">6.2 mm</quantity>
,
<collectionCode id="55A8F7FB6C3A2C6940C5F91DFA9C2A96" box="[1239,1296,1674,1700]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="21">ZRC</collectionCode>
2007.0110;
<specimenCount id="25BFA4B76C3A2C694485F925FE8C2AFE" box="[151,256,1714,1740]" pageId="4" pageNumber="21" type="female">1 female</specimenCount>
, 4.8 ×
<quantity id="F441C2DB6C3A2C69455DF925FE202AFE" box="[335,428,1714,1740]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="21" unit="mm" value="5.6">5.6 mm</quantity>
,
<collectionCode id="55A8F7FB6C3A2C6945AAF925FE7D2AFE" box="[440,497,1714,1740]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="21">ZRC</collectionCode>
2007.0111;
<specimenCount id="25BFA4B76C3A2C694690F925FD7A2AFE" box="[642,758,1714,1740]" pageId="4" pageNumber="21" type="female">4 females</specimenCount>
, 3.9 ×
<quantity id="F441C2DB6C3A2C694754F925FC2C2AFE" box="[838,928,1714,1740]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="21" unit="mm" value="5.5">5.5 mm</quantity>
4.8 ×
<quantity id="F441C2DB6C3A2C694013F925FBD22AFE" box="[1025,1118,1714,1740]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="21" unit="mm" value="5.6">5.6 mm</quantity>
,
<collectionCode id="55A8F7FB6C3A2C69407BF925FB2E2AFE" box="[1129,1186,1714,1740]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="21">ZRC</collectionCode>
2007.0112; 16 ovig., 4.1 ×
<quantity id="F441C2DB6C3A2C6944C8F94DFEB82AC6" box="[218,308,1754,1780]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="21" unit="mm" value="4.9">4.9 mm</quantity>
4.5 ×
<quantity id="F441C2DB6C3A2C694581F94DFE7C2AC6" box="[403,496,1754,1780]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="21" unit="mm" value="5.4">5.4 mm</quantity>
,
<collectionCode id="55A8F7FB6C3A2C6945E9F94DFDB82AC6" box="[507,564,1754,1780]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="21">ZRC</collectionCode>
2007.0113;
<specimenCount id="25BFA4B76C3A2C6946D4F94DFCAF2AC6" box="[710,803,1754,1780]" pageId="4" pageNumber="21" type="male">5 males</specimenCount>
, 4.2 × 5.0 5.0 ×
<quantity id="F441C2DB6C3A2C6947E4F94DFBDF2AC6" box="[1014,1107,1754,1780]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="4" pageNumber="21" unit="mm" value="5.8">5.8 mm</quantity>
, 2 ovig., 3.8 × 4.2, 3.8 ×
<quantity id="F441C2DB6C3A2C69416AF94DFF442B2E" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="21" unit="mm" value="4.3">4.3 mm</quantity>
, NHM 2007.600606;
<specimenCount id="25BFA4B76C3A2C6945CBF895FDBA2B2E" box="[473,566,1794,1820]" pageId="4" pageNumber="21" type="male">5 males</specimenCount>
, 4.0 × 4.8 4.9 ×
<quantity id="F441C2DB6C3A2C694702F895FCE22B2E" box="[784,878,1794,1820]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="21" unit="mm" value="5.7">5.7 mm</quantity>
, 2 ovig., 3.8 ×
<quantity id="F441C2DB6C3A2C694032F895FBF22B2E" box="[1056,1150,1794,1820]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="21" unit="mm" value="4.4">4.4 mm</quantity>
, 4.2 ×
<quantity id="F441C2DB6C3A2C6940DDF895FAA12B2E" box="[1231,1325,1794,1820]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="21" unit="mm" value="4.8">4.8 mm</quantity>
, MNHN- B30393. All
<typeStatus id="EC02D19C6C3A2C69453DF8BDFE132B76" box="[303,415,1834,1860]" pageId="4" pageNumber="21" type="paratype">paratypes</typeStatus>
were collected together with the
<typeStatus id="EC02D19C6C3A2C694734F8BDFC1D2B76" box="[806,913,1834,1860]" pageId="4" pageNumber="21" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3A2C6544D4F8C5FF782B4F" lastPageId="8" lastPageNumber="25" pageId="4" pageNumber="21" type="description">
<paragraph id="33066F3E6C3A2C6844D4F8C5FBDF2D18" blockId="4.[151,1437,1594,2020]" lastBlockId="5.[151,1436,152,298]" lastPageId="5" lastPageNumber="22" pageId="4" pageNumber="21">
<emphasis id="01CDB32C6C3A2C6944D4F8C5FEDB2B5E" bold="true" box="[198,343,1874,1900]" pageId="4" pageNumber="21">Description</emphasis>
. Carapace (
<figureCitation id="AB8273BB6C3A2C6945F1F8C5FDA52B5E" box="[483,553,1874,1900]" captionStart="FIGURE 4" captionStartId="6.[151,255,1923,1947]" captionTargetBox="[267,1330,465,1896]" captionTargetId="figure@6.[249,1427,456,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hymenicoides robertsi new species. a, habitus; b, left third maxilliped; c, male left chela, outer view; d, male left chela, dorsal view; e, right second ambulatory dactylus. a, c e, holotype male (ZRC 2007.0108), 4.9 × 6.0 mm; b, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179199/files/figure.png" pageId="4" pageNumber="21">Fig. 4</figureCitation>
a) oval, CW 1.131.24 times CL (mean 1.18, n = 9) CL; dorsal surface flat, surrounded by continuous rim, regions well demarcated by grooves, H-shaped gastric groove continuous with cervical groove, cervical groove branching anteriorly, branches confluent with anterolateral rim. Rostrum vestigial, triangular. Lateral margin of carapace lacking tooth or lobe, side wall of posterolateral region slightly expanded laterally, with longitudinal groove along posterolateral rim below. Conical tooth present between antennules, completely disconnected from vestigial rostrum; orbit indiscernible, without tooth on outer part of eye. Epistome long, placed anterior-dorsally to buccal cavern, ischium of third maxilliped partially covering posterior margin; posterior margin with trapezoidal convexity with median notch.
</paragraph>
<caption id="67C63FB66C3B2C684485FC31FB702F8F" box="[151,1276,934,958]" httpUri="https://zenodo.org/record/179198/files/figure.png" pageId="5" pageNumber="22" targetBox="[375,1210,357,907]" targetPageId="5">
<paragraph id="33066F3E6C3B2C684485FC31FB702F8F" blockId="5.[151,1276,934,958]" box="[151,1276,934,958]" pageId="5" pageNumber="22">
<emphasis id="01CDB32C6C3B2C684485FC31FE952F8C" bold="true" box="[151,281,934,958]" pageId="5" pageNumber="22">FIGURE 3.</emphasis>
Live colour of
<taxonomicName id="F4B914BD6C3B2C6845ADFC31FD3C2F8F" box="[447,688,934,957]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C3B2C6845ADFC31FD3C2F8F" box="[447,688,934,957]" italics="true" pageId="5" pageNumber="22">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3B2C6846A5FC31FCB52F8C" bold="true" box="[695,825,934,958]" pageId="5" pageNumber="22">
<taxonomicNameLabel id="1AFE0E576C3B2C6846A5FC31FCB52F8C" box="[695,825,934,958]" pageId="5" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
. Photograph was taken by Tyson Roberts.
</paragraph>
</caption>
<paragraph id="33066F3E6C3B2C6844D4FB84FB8028FF" blockId="5.[151,1437,1043,2029]" pageId="5" pageNumber="22">
Eyes moderately developed, visible dorsally. Antennule with long coxa and basis; basis more than half length of coxa. Third maxilliped (
<figureCitation id="AB8273BB6C3B2C684630FBACFDE42867" box="[546,616,1083,1109]" captionStart="FIGURE 4" captionStartId="6.[151,255,1923,1947]" captionTargetBox="[267,1330,465,1896]" captionTargetId="figure@6.[249,1427,456,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hymenicoides robertsi new species. a, habitus; b, left third maxilliped; c, male left chela, outer view; d, male left chela, dorsal view; e, right second ambulatory dactylus. a, c e, holotype male (ZRC 2007.0108), 4.9 × 6.0 mm; b, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179199/files/figure.png" pageId="5" pageNumber="22">Fig. 4</figureCitation>
b) narrow, covering about one-quarter of buccal cavern; mid-length of ischium about three-quarters of merus, distal inner angle produced; palp long, propodus about half length of dactylus, dactylus as long as merus, tip of dactylus almost reaching proximal end of ischium
<emphasis id="01CDB32C6C3B2C6840F9FB1BFAB92897" box="[1259,1333,1164,1189]" italics="true" pageId="5" pageNumber="22">in situ</emphasis>
; exopod short, reaching about proximal two-thirds of merus, with distinct flagellum.
</paragraph>
<paragraph id="33066F3E6C3B2C6844D4FB4CFEDB29A7" blockId="5.[151,1437,1043,2029]" pageId="5" pageNumber="22">
Male with relatively wide abdominal cavity (
<figureCitation id="AB8273BB6C3B2C6846CAFB4CFC9728C7" box="[728,795,1243,1269]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="5" pageNumber="22">Fig. 1</figureCitation>
a), sternal button narrow, high, just posterior to imaginary line joining inner ends of sutures between sternites 5 and 6. Female thoracic sternite 3 separated from sternite 4 by posteriorly convex rim, sterites 48 medially fused, vulva on imaginary line joining inner ends of sutures between sternites 5 and 6 on medial fused plate of thoracic sternum, vulva with longitudinally elliptical basal mount.
</paragraph>
<paragraph id="33066F3E6C3B2C6844D4FA34FE972A9F" blockId="5.[151,1437,1043,2029]" pageId="5" pageNumber="22">
Chelipeds symmetrical, relatively stouter in males; male cheliped with short merus, ventral outer margin with subdistal tooth, inner margin of merus to carpus lined with long stiff setae, outer surface of carpus rounded, not strongly elevated, outer margin with subdistal tooth; chela (
<figureCitation id="AB8273BB6C3B2C6847F7FA64FBA72A3F" box="[997,1067,1523,1549]" captionStart="FIGURE 4" captionStartId="6.[151,255,1923,1947]" captionTargetBox="[267,1330,465,1896]" captionTargetId="figure@6.[249,1427,456,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hymenicoides robertsi new species. a, habitus; b, left third maxilliped; c, male left chela, outer view; d, male left chela, dorsal view; e, right second ambulatory dactylus. a, c e, holotype male (ZRC 2007.0108), 4.9 × 6.0 mm; b, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179199/files/figure.png" pageId="5" pageNumber="22">Fig. 4</figureCitation>
c, d) with rounded palm, almost glabrous, outer surface inflated, with dorso-ventrally flattened tubercle on outer surface of proximal lower part of base of immovable finger, tubercle absent in females; fingers with ovoid gape when closed, tips slightly hoof-like; immovable finger with ca. 6 low teeth; teeth of movable finger similar to those of immovable finger.
</paragraph>
<paragraph id="33066F3E6C3B2C6844D4F92CFF572B7F" blockId="5.[151,1437,1043,2029]" pageId="5" pageNumber="22">
Ambulatory legs (
<figureCitation id="AB8273BB6C3B2C6845B4F92CFE7E2AE7" box="[422,498,1723,1749]" captionStart="FIGURE 4" captionStartId="6.[151,255,1923,1947]" captionTargetBox="[267,1330,465,1896]" captionTargetId="figure@6.[249,1427,456,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hymenicoides robertsi new species. a, habitus; b, left third maxilliped; c, male left chela, outer view; d, male left chela, dorsal view; e, right second ambulatory dactylus. a, c e, holotype male (ZRC 2007.0108), 4.9 × 6.0 mm; b, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179199/files/figure.png" pageId="5" pageNumber="22">Fig. 4</figureCitation>
a) slender, long, second longest, inner margin of propodi to dactyli fringed densely lined with long plumose setae; meri slightly longer than respective propodi, distal anterior angle not produced; dactyli (
<figureCitation id="AB8273BB6C3B2C684568F89CFE482B17" box="[378,452,1803,1829]" captionStart="FIGURE 4" captionStartId="6.[151,255,1923,1947]" captionTargetBox="[267,1330,465,1896]" captionTargetId="figure@6.[249,1427,456,1920]" captionTargetPageId="6" captionText="FIGURE 4. Hymenicoides robertsi new species. a, habitus; b, left third maxilliped; c, male left chela, outer view; d, male left chela, dorsal view; e, right second ambulatory dactylus. a, c e, holotype male (ZRC 2007.0108), 4.9 × 6.0 mm; b, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179199/files/figure.png" pageId="5" pageNumber="22">Fig. 4</figureCitation>
e) terminating in sharp tooth, with subterminal inner tooth placed far from distal tooth.
</paragraph>
<paragraph id="33066F3E6C3B2C6B44D4F8CCFD2B2D90" blockId="5.[151,1437,1043,2029]" lastBlockId="6.[151,1436,152,418]" lastPageId="6" lastPageNumber="23" pageId="5" pageNumber="22">
Male abdomen-pleotelson (
<figureCitation id="AB8273BB6C3B2C684615F8CCFDC02B47" box="[519,588,1883,1909]" captionStart="FIGURE 5" captionStartId="7.[151,255,1935,1959]" captionTargetBox="[201,1409,197,1918]" captionTargetId="figure@7.[169,1454,161,1919]" captionTargetPageId="7" captionText="FIGURE 5. Hymenicoides robertsi new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1, ventral view; d, left G 1, outer view; e, left G 2. a, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm; b, paratype, female (ZRC 2007.0111), 4.8 × 5.6 mm; c e, holotype, male (ZRC 2007.0108), 4.9 × 6.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179200/files/figure.png" pageId="5" pageNumber="22">Fig. 5</figureCitation>
a) 6-segmented; first segment thick, hard, distal margin widely concave; second to fifth segments with distal margin concave medially, third segment widest; pleotelson trilobed, lateral lobes distinct, auriculate, inner surface of lateral lobes thickened externally, forming socket for sternal button on inner surface (
<figureCitation id="AB8273BB6C3B2C6845A7F844FE742BDF" box="[437,504,2003,2029]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="5" pageNumber="22">Fig. 1</figureCitation>
a, b). G1 (
<figureCitation id="AB8273BB6C3B2C684662F844FD3A2BDF" box="[624,694,2003,2029]" captionStart="FIGURE 5" captionStartId="7.[151,255,1935,1959]" captionTargetBox="[201,1409,197,1918]" captionTargetId="figure@7.[169,1454,161,1919]" captionTargetPageId="7" captionText="FIGURE 5. Hymenicoides robertsi new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1, ventral view; d, left G 1, outer view; e, left G 2. a, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm; b, paratype, female (ZRC 2007.0111), 4.8 × 5.6 mm; c e, holotype, male (ZRC 2007.0108), 4.9 × 6.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179200/files/figure.png" pageId="5" pageNumber="22">Fig. 5</figureCitation>
c, d) stout, strongly bent on distal half, with longitudinal line of long, stiff setae on distal to medial inner angle of shaft; distal inner end with beak-like chitinous projection, distal outer angle swollen, covered by tiny granule, distal outer angle dorsally connected with thumb-like projection. G2 (
<figureCitation id="AB8273BB6C382C6B4538FF7FFEE32D30" box="[298,367,232,258]" captionStart="FIGURE 5" captionStartId="7.[151,255,1935,1959]" captionTargetBox="[201,1409,197,1918]" captionTargetId="figure@7.[169,1454,161,1919]" captionTargetPageId="7" captionText="FIGURE 5. Hymenicoides robertsi new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1, ventral view; d, left G 1, outer view; e, left G 2. a, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm; b, paratype, female (ZRC 2007.0111), 4.8 × 5.6 mm; c e, holotype, male (ZRC 2007.0108), 4.9 × 6.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179200/files/figure.png" pageId="6" pageNumber="23">Fig. 5</figureCitation>
e) short, less than half length of G1. Female abdomen-pleotelson (
<figureCitation id="AB8273BB6C382C6B4068FF7FFB332D30" box="[1146,1215,232,258]" captionStart="FIGURE 5" captionStartId="7.[151,255,1935,1959]" captionTargetBox="[201,1409,197,1918]" captionTargetId="figure@7.[169,1454,161,1919]" captionTargetPageId="7" captionText="FIGURE 5. Hymenicoides robertsi new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1, ventral view; d, left G 1, outer view; e, left G 2. a, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm; b, paratype, female (ZRC 2007.0111), 4.8 × 5.6 mm; c e, holotype, male (ZRC 2007.0108), 4.9 × 6.0 mm. Scale bars, 1 mm." httpUri="https://zenodo.org/record/179200/files/figure.png" pageId="6" pageNumber="23">Fig. 5</figureCitation>
b) demarcated to 6 segments, boundary between second to fifth segments movable, more or less fixed in other boundaries; second to fifth segments with corneous wide ridge medially, that of pleotelson only on proximal half, pleotelson longest; pleopods (
<figureCitation id="AB8273BB6C382C6B4567FEF7FE372D48" box="[373,443,352,378]" captionStart="FIGURE 6" captionStartId="8.[151,255,1506,1530]" captionTargetBox="[151,1436,184,1485]" captionTargetId="figure@8.[151,1436,184,1485]" captionTargetPageId="8" captionText="FIGURE 6. Hymenicoides robertsi new species. a d, right second to fifth pleopods; e, exopods. Paratype, ovigerous female (ZRC 2007.0113), 4.2 × 5.0 mm." httpUri="https://zenodo.org/record/179201/files/figure.png" pageId="6" pageNumber="23">Fig. 6</figureCitation>
) on second to fifth segments, long, biramous from near base, developing from distal outer angle of inner surface of each segment.
</paragraph>
<caption id="67C63FB66C382C6B4485F814FCF72BED" httpUri="https://zenodo.org/record/179199/files/figure.png" pageId="6" pageNumber="23" targetBox="[267,1330,465,1896]" targetPageId="6">
<paragraph id="33066F3E6C382C6B4485F814FCF72BED" blockId="6.[151,1436,1923,2015]" pageId="6" pageNumber="23">
<emphasis id="01CDB32C6C382C6B4485F814FE972BA9" bold="true" box="[151,283,1923,1947]" pageId="6" pageNumber="23">FIGURE 4.</emphasis>
<taxonomicName id="F4B914BD6C382C6B4536F814FD942BA8" box="[292,536,1923,1946]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="23" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C382C6B4536F814FD942BA8" box="[292,536,1923,1946]" italics="true" pageId="6" pageNumber="23">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C382C6B4633F814FD2A2BA9" bold="true" box="[545,678,1923,1947]" pageId="6" pageNumber="23">
<taxonomicNameLabel id="1AFE0E576C382C6B4633F814FD2A2BA9" box="[545,678,1923,1947]" pageId="6" pageNumber="23" rank="species">new species</taxonomicNameLabel>
</emphasis>
. a, habitus; b, left third maxilliped; c, male left chela, outer view; d, male left chela, dorsal view; e, right second ambulatory dactylus. a, ce, holotype male (ZRC 2007.0108), 4.9 × 6.0 mm; b, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm. Scale bars, 1 mm.
</paragraph>
</caption>
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<paragraph id="33066F3E6C392C6A4485F818FB772BD5" blockId="7.[151,1436,1935,2023]" pageId="7" pageNumber="24">
<emphasis id="01CDB32C6C392C6A4485F818FE952B95" bold="true" box="[151,281,1935,1959]" pageId="7" pageNumber="24">FIGURE 5.</emphasis>
<taxonomicName id="F4B914BD6C392C6A4532F807FD9C2B95" box="[288,528,1936,1959]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C392C6A4532F807FD9C2B95" box="[288,528,1936,1959]" italics="true" pageId="7" pageNumber="24">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C392C6A4605F818FD162B95" bold="true" box="[535,666,1935,1959]" pageId="7" pageNumber="24">
<taxonomicNameLabel id="1AFE0E576C392C6A4605F818FD162B95" box="[535,666,1935,1959]" pageId="7" pageNumber="24" rank="species">new species</taxonomicNameLabel>
</emphasis>
. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G1, ventral view; d, left G1, outer view; e, left G2. a, paratype, male (ZRC 2007.0109), 5.0 × 6.2 mm; b, paratype, female (ZRC 2007.0111), 4.8 × 5.6 mm; ce, holotype, male (ZRC 2007.0108), 4.9 × 6.0 mm. Scale bars, 1 mm.
</paragraph>
</caption>
<caption id="67C63FB66C362C654485FA75FDCB2A28" httpUri="https://zenodo.org/record/179201/files/figure.png" pageId="8" pageNumber="25" targetBox="[151,1436,184,1485]" targetPageId="8">
<paragraph id="33066F3E6C362C654485FA75FDCB2A28" blockId="8.[151,1436,1506,1562]" pageId="8" pageNumber="25">
<emphasis id="01CDB32C6C362C654485FA75FE9729C8" bold="true" box="[151,283,1506,1530]" pageId="8" pageNumber="25">FIGURE 6.</emphasis>
<taxonomicName id="F4B914BD6C362C654537FA74FD9529C8" box="[293,537,1507,1530]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C362C654537FA74FD9529C8" box="[293,537,1507,1530]" italics="true" pageId="8" pageNumber="25">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C362C654633FA75FD2A29C8" bold="true" box="[545,678,1506,1530]" pageId="8" pageNumber="25">
<taxonomicNameLabel id="1AFE0E576C362C654633FA75FD2A29C8" box="[545,678,1506,1530]" pageId="8" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
. ad, right second to fifth pleopods; e, exopods. Paratype, ovigerous female (ZRC 2007.0113), 4.2 × 5.0 mm.
</paragraph>
</caption>
<paragraph id="33066F3E6C362C6544D4F9DCFE562A87" blockId="8.[151,1437,1611,2037]" pageId="8" pageNumber="25">
Eggs spherical, small, diameter
<quantity id="F441C2DB6C362C65462DF9DCFD772A57" box="[575,763,1611,1637]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="8" pageNumber="25" unit="mm" value="0.285" valueMax="0.3" valueMin="0.27">0.27 0.30 mm</quantity>
[mean = 0.28, n = 15 from three females (3.9 ×
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, 4.1 ×
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, 4.2 × 5.0 mm)], attached on endopods of pleopods, exopods growing inwards, partially covering dome-shaped abdomen.
</paragraph>
<paragraph id="33066F3E6C362C6544D4F955FF782B4F" blockId="8.[151,1437,1611,2037]" pageId="8" pageNumber="25">
<emphasis id="01CDB32C6C362C6544D4F955FE722AEE" bold="true" box="[198,510,1730,1756]" pageId="8" pageNumber="25">Habitat and distribution</emphasis>
. All specimens of
<taxonomicName id="F4B914BD6C362C6546F7F954FC7D2AEE" box="[741,1009,1731,1756]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="robertsi">
<emphasis id="01CDB32C6C362C6546F7F954FC7D2AEE" box="[741,1009,1731,1756]" italics="true" pageId="8" pageNumber="25">Hymenicoides robertsi</emphasis>
</taxonomicName>
were obtained from the Kyaukdaw market on the lower Kaladan River (with tidal influence), Rakhine,
<collectingCountry id="4BAE2FAE6C362C6547C6F97CFBC42B37" box="[980,1096,1771,1797]" name="Myanmar" pageId="8" pageNumber="25">Myanmar</collectingCountry>
. The specimens were found crawling over a large number of young
<taxonomicName id="F4B914BD6C362C654662F884FC9C2B1E" box="[624,784,1811,1836]" class="Actinopterygii" family="Ophichthidae" genus="Pisodonophis" kingdom="Animalia" order="Anguilliformes" pageId="8" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="01CDB32C6C362C654662F884FC9C2B1E" box="[624,784,1811,1836]" italics="true" pageId="8" pageNumber="25">Pisodonophis</emphasis>
</taxonomicName>
snake eels in a tray in the market (T. R. Roberts, pers. comm.) and had been apparently collected in the nearby area. The species is known only from
<collectingCountry id="4BAE2FAE6C362C654119F8ACFAF12B67" box="[1291,1405,1851,1877]" name="Myanmar" pageId="8" pageNumber="25">Myanmar</collectingCountry>
at present.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C362C6544D4F81DFE872BC7" pageId="8" pageNumber="25" type="etymology">
<paragraph id="33066F3E6C362C6544D4F81DFE872BC7" blockId="8.[151,1437,1611,2037]" pageId="8" pageNumber="25">
<emphasis id="01CDB32C6C362C6544D4F81DFEC72B96" bold="true" box="[198,331,1930,1956]" pageId="8" pageNumber="25">Etymology</emphasis>
. We are pleased to name this species for the well known ichthyologist, Tyson R. Roberts, who very generously provided the specimens of this new species as well as those of
<taxonomicName id="F4B914BD6C362C654023F824FB282BFE" box="[1073,1188,1971,1996]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="25" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C362C654023F824FB282BFE" box="[1073,1188,1971,1996]" italics="true" pageId="8" pageNumber="25">H. carteri</emphasis>
</taxonomicName>
from
<collectingCountry id="4BAE2FAE6C362C6540F8F824FAFF2BFF" box="[1258,1395,1971,1997]" name="Bangladesh" pageId="8" pageNumber="25">Bangladesh</collectingCountry>
for our study.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C372C6444D4FF00FBE12D90" pageId="9" pageNumber="26" type="discussion">
<paragraph id="33066F3E6C372C6444D4FF00FBE12D90" blockId="9.[151,1436,151,418]" pageId="9" pageNumber="26">
<emphasis id="01CDB32C6C372C6444D4FF00FEB42C83" bold="true" box="[198,312,151,177]" pageId="9" pageNumber="26">Remarks</emphasis>
.
<taxonomicName id="F4B914BD6C372C64455BFF0FFDD82C83" box="[329,596,152,177]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C372C64455BFF0FFDD82C83" box="[329,596,152,177]" italics="true" pageId="9" pageNumber="26">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C372C64464EFF00FD622C83" bold="true" box="[604,750,151,177]" pageId="9" pageNumber="26">
<taxonomicNameLabel id="1AFE0E576C372C64464EFF00FD622C83" box="[604,750,151,177]" pageId="9" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
, is clearly different from
<taxonomicName id="F4B914BD6C372C644036FF0FFB162C83" box="[1060,1178,152,177]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C372C644036FF0FFB162C83" box="[1060,1178,152,177]" italics="true" pageId="9" pageNumber="26">H. carteri</emphasis>
</taxonomicName>
in its proportionately more slender ambulatory legs (
<figureCitation id="AB8273BB6C372C64461AFF57FDC32CE8" box="[520,591,192,218]" captionStart="FIGURE 3" captionStartId="5.[151,255,934,958]" captionTargetBox="[375,1210,357,907]" captionTargetId="figure@5.[375,1211,357,907]" captionTargetPageId="5" captionText="FIGURE 3. Live colour of Hymenicoides robertsi new species. Photograph was taken by Tyson Roberts." httpUri="https://zenodo.org/record/179198/files/figure.png" pageId="9" pageNumber="26">Fig. 3</figureCitation>
a;
<bibRefCitation id="572812CF6C372C64467FFF57FC882CE8" author="Kemp" box="[621,772,192,218]" pageId="9" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
: Fig. 17). In addition, both species can be distinguished by the armature of the inner margin of the ambulatory dactyli. In
<taxonomicName id="F4B914BD6C372C64479FFF7FFB802D33" box="[909,1036,232,257]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="robertsi">
<emphasis id="01CDB32C6C372C64479FFF7FFB802D33" box="[909,1036,232,257]" italics="true" pageId="9" pageNumber="26">H. robertsi</emphasis>
</taxonomicName>
, all the dactyli have only one subterminal tooth, which is placed some distance from the tip, while in
<taxonomicName id="F4B914BD6C372C6447D0FE87FBBB2D1B" box="[962,1079,272,297]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C372C6447D0FE87FBBB2D1B" box="[962,1079,272,297]" italics="true" pageId="9" pageNumber="26">H. carteri</emphasis>
</taxonomicName>
, the third to fifth dactyli have 311 small teeth (posterior legs with fewer teeth) and a subdistal large tooth which is placed closer to the tip. Another major difference is the single beak-like chitinous projection of the G1 of
<taxonomicName id="F4B914BD6C372C644086FEF7FA942D4B" box="[1172,1304,352,377]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="robertsi">
<emphasis id="01CDB32C6C372C644086FEF7FA942D4B" box="[1172,1304,352,377]" italics="true" pageId="9" pageNumber="26">H. robertsi</emphasis>
</taxonomicName>
(biramous shorter projection in
<taxonomicName id="F4B914BD6C372C64459EFE1FFE732D93" box="[396,511,392,417]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C372C64459EFE1FFE732D93" box="[396,511,392,417]" italics="true" pageId="9" pageNumber="26">H. carteri</emphasis>
</taxonomicName>
) (
<figureCitation id="AB8273BB6C372C64460BFE1FFDE72D90" box="[537,619,392,418]" captionStart="FIGURE 2" captionStartId="4.[151,255,1343,1367]" captionTargetBox="[352,1266,221,1298]" captionTargetId="figure@4.[294,1293,197,1317]" captionTargetPageId="4" captionText="FIGURE 2. Male first and second gonopods of Hymenicoides carteri Kemp, 1917. Lectotype, male (NHM 1919.11.1.121), 4.0 × 4.8 mm. a, left G 1, ventral view; b, left G 1, outer view; c, left G 2. Scale bar, 1 mm." httpUri="https://zenodo.org/record/179197/files/figure.png" pageId="9" pageNumber="26">Figs. 2</figureCitation>
a, b, 4a, e, 5c, d;
<bibRefCitation id="572812CF6C372C644723FE1FFC4A2D90" author="Kemp" box="[817,966,392,418]" pageId="9" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
: Figs. 17, 20).
</paragraph>
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<subSubSection id="7BA33CB56C3D2C6E4485FCA8FF632F4E" pageId="3" pageNumber="20" type="nomenclature">
<paragraph id="33066F3E6C3D2C6E4485FCA8FDB92F6B" blockId="3.[151,565,831,892]" box="[151,565,831,857]" pageId="3" pageNumber="20">
<heading id="684ED8526C3D2C6E4485FCA8FDB92F6B" bold="true" box="[151,565,831,857]" fontSize="11" level="1" pageId="3" pageNumber="20" reason="1">
<taxonomicName id="F4B914BD6C3D2C6E4485FCA8FDB92F6B" ID-CoL="3NJ2G" authority="Kemp, 1917" authorityName="Kemp" authorityYear="1917" box="[151,565,831,857]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3D2C6E4485FCA8FDB92F6B" bold="true" box="[151,565,831,857]" pageId="3" pageNumber="20">
<emphasis id="01CDB32C6C3D2C6E4485FCA8FE152F6B" bold="true" box="[151,409,831,857]" italics="true" pageId="3" pageNumber="20">Hymenicoides carteri</emphasis>
<bibRefCitation id="572812CF6C3D2C6E45B2FCA8FDB92F6B" author="Kemp" box="[416,565,831,857]" pageId="3" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="33066F3E6C3D2C6E4485FCF5FF632F4E" blockId="3.[151,565,831,892]" box="[151,239,866,892]" pageId="3" pageNumber="20">
(
<figureCitation id="AB8273BB6C3D2C6E448DFCF5FF692F4E" box="[159,229,866,892]" captionStart="FIGURE 2" captionStartId="4.[151,255,1343,1367]" captionTargetBox="[352,1266,221,1298]" captionTargetId="figure@4.[294,1293,197,1317]" captionTargetPageId="4" captionText="FIGURE 2. Male first and second gonopods of Hymenicoides carteri Kemp, 1917. Lectotype, male (NHM 1919.11.1.121), 4.0 × 4.8 mm. a, left G 1, ventral view; b, left G 1, outer view; c, left G 2. Scale bar, 1 mm." httpUri="https://zenodo.org/record/179197/files/figure.png" pageId="3" pageNumber="20">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3D2C6E4485FC25FF7A2FDB" pageId="3" pageNumber="20" type="reference_group">
<paragraph id="33066F3E6C3D2C6E4485FC25FF7A2FDB" blockId="3.[151,1437,946,1001]" pageId="3" pageNumber="20">
<treatmentCitationGroup id="13A948106C3D2C6E4485FC25FF7A2FDB" pageId="3" pageNumber="20">
<emphasis id="01CDB32C6C3D2C6E4485FC25FE182FFB" box="[151,404,946,969]" italics="true" pageId="3" pageNumber="20">
<taxonomicName id="F4B914BD6C3D2C6E4485FC25FEA22FFB" ID-CoL="7PL39" box="[151,302,946,969]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">Hymenicoides</taxonomicName>
carteri—
</emphasis>
<treatmentCitation id="B218492F6C3D2C6E4589FC25FDDA2FFB" author="Kemp" box="[411,598,946,969]" page="268" pageId="3" pageNumber="20" year="1917">
<bibRefCitation id="572812CF6C3D2C6E4589FC25FDAD2FFB" author="Kemp" box="[411,545,946,969]" pageId="3" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
: 268
</treatmentCitation>
;
<treatmentCitation id="B218492F6C3D2C6E4673FC25FC972FFB" author="Lucas" box="[609,795,946,969]" page="197" pageId="3" pageNumber="20" year="1980">
<bibRefCitation id="572812CF6C3D2C6E4673FC25FD6A2FFB" author="Lucas" box="[609,742,946,969]" pageId="3" pageNumber="31" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas, 1980</bibRefCitation>
: 197
</treatmentCitation>
;
<treatmentCitation id="B218492F6C3D2C6E4734FC25FBA42FFB" author="Ng" box="[806,1064,946,969]" page="50" pageId="3" pageNumber="20" year="1996">
<bibRefCitation id="572812CF6C3D2C6E4734FC25FB8C2FFB" author="Ng" box="[806,1024,946,969]" pageId="3" pageNumber="31" refString="Ng, P. K. L. &amp; Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement, 3, 1 - 82." type="journal article" year="1996">Ng &amp; Chuang, 1996</bibRefCitation>
: 50
</treatmentCitation>
; Guinot &amp; Richer de
<treatmentCitation id="B218492F6C3D2C6E4118FC25FF7E2FDB" author="Forges" page="462" pageId="3" pageNumber="20" year="1997">
<bibRefCitation id="572812CF6C3D2C6E4118FC25FA142FFB" author="Forges" box="[1290,1432,946,969]" pageId="3" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
: 462
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3D2C6E4485FB8DFCF828CE" pageId="3" pageNumber="20" type="materials_examined">
<paragraph id="33066F3E6C3D2C6E4485FB8DFC3F286E" blockId="3.[151,1437,1050,1956]" pageId="3" pageNumber="20">
<emphasis id="01CDB32C6C3D2C6E4485FB8DFE082806" bold="true" box="[151,388,1050,1076]" pageId="3" pageNumber="20">Material examined</emphasis>
.
<emphasis id="01CDB32C6C3D2C6E4586FB8DFD9C2806" bold="true" box="[404,528,1050,1076]" pageId="3" pageNumber="20">
<typeStatus id="EC02D19C6C3D2C6E4586FB8DFD9C2806" box="[404,528,1050,1076]" pageId="3" pageNumber="20" type="lectotype">Lectotype</typeStatus>
</emphasis>
(present designation): male, 4.0 ×
<quantity id="F441C2DB6C3D2C6E47B9FB8DFB852806" box="[939,1033,1050,1076]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="20" unit="mm" value="4.8">4.8 mm</quantity>
, NHM 1919.11.1.121, R. Nughli, Sibpur, near Calcutta,
<collectingCountry id="4BAE2FAE6C3D2C6E458EFBD5FE57286E" box="[412,475,1090,1116]" name="India" pageId="3" pageNumber="20">India</collectingCountry>
, coll. Annandale &amp; S. Kemp,
<date id="470749FE6C3D2C6E4729FBD5FC23286E" box="[827,943,1090,1116]" pageId="3" pageNumber="20" value="1917-01">Jan. 1917</date>
.
</paragraph>
<paragraph id="33066F3E6C3D2C6E44D4FBFDFD1A289E" blockId="3.[151,1437,1050,1956]" pageId="3" pageNumber="20">
<emphasis id="01CDB32C6C3D2C6E44D4FBFDFEF228B6" bold="true" box="[198,382,1130,1156]" pageId="3" pageNumber="20">
<typeStatus id="EC02D19C6C3D2C6E44D4FBFDFEF228B6" box="[198,382,1130,1156]" pageId="3" pageNumber="20" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
:
<specimenCount id="25BFA4B76C3D2C6E459CFBFDFE6C28B6" box="[398,480,1130,1156]" pageId="3" pageNumber="20" type="male">1 male</specimenCount>
, 4.0 ×
<quantity id="F441C2DB6C3D2C6E4622FBFDFD0128B6" box="[560,653,1130,1156]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="20" unit="mm" value="4.8">4.8 mm</quantity>
,
<specimenCount id="25BFA4B76C3D2C6E468BFBFDFC8E28B6" box="[665,770,1130,1156]" pageId="3" pageNumber="20" type="female">1 female</specimenCount>
, 3.2 ×
<quantity id="F441C2DB6C3D2C6E4740FBFDFC2328B6" box="[850,943,1130,1156]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="20" unit="mm" value="3.7">3.7 mm</quantity>
, 1 ovigerous female, 3.0 ×
<quantity id="F441C2DB6C3D2C6E40FDFBFDFAC028B6" box="[1263,1356,1130,1156]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="20" unit="mm" value="3.7">3.7 mm</quantity>
, NHM 1919.11.1.122123, data same as
<typeStatus id="EC02D19C6C3D2C6E4631FB05FD1E289E" box="[547,658,1170,1196]" pageId="3" pageNumber="20" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="33066F3E6C3D2C6E44D4FB2DFCF828CE" blockId="3.[151,1437,1050,1956]" pageId="3" pageNumber="20">
Others.
<specimenCount id="25BFA4B76C3D2C6E4536FB2DFEFA28E6" box="[292,374,1210,1236]" pageId="3" pageNumber="20" type="male">1 male</specimenCount>
, 2.2 ×
<quantity id="F441C2DB6C3D2C6E45DAFB2DFDA928E6" box="[456,549,1210,1236]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="20" unit="mm" value="2.4">2.4 mm</quantity>
,
<specimenCount id="25BFA4B76C3D2C6E4620FB2DFD2A28E6" box="[562,678,1210,1236]" pageId="3" pageNumber="20" type="female">2 females</specimenCount>
, 2.6 ×
<quantity id="F441C2DB6C3D2C6E46EAFB2DFCD828E6" box="[760,852,1210,1236]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="20" unit="mm" value="2.9">2.9 mm</quantity>
, 2.7 ×
<quantity id="F441C2DB6C3D2C6E47B5FB2DFB8828E6" box="[935,1028,1210,1236]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="20" unit="mm" value="2.9">2.9 mm</quantity>
,
<collectionCode id="55A8F7FB6C3D2C6E4002FB2DFBC528E6" box="[1040,1097,1210,1236]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="20">ZRC</collectionCode>
2007.0107, Tangail District, North Central region, Ganges basin,
<collectingCountry id="4BAE2FAE6C3D2C6E4657FB75FD5D28CE" box="[581,721,1250,1276]" name="Bangladesh" pageId="3" pageNumber="20">Bangladesh</collectingCountry>
,
<date id="470749FE6C3D2C6E46CEFB75FCFC28CE" box="[732,880,1250,1276]" pageId="3" pageNumber="20" value="1992-11-04">4 Nov. 1992</date>
.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3D2C6E44D4FA9DFADF2B96" pageId="3" pageNumber="20" type="discussion">
<paragraph id="33066F3E6C3D2C6E44D4FA9DFAB42A56" blockId="3.[151,1437,1050,1956]" pageId="3" pageNumber="20">
<emphasis id="01CDB32C6C3D2C6E44D4FA9DFEB42916" bold="true" box="[198,312,1290,1316]" pageId="3" pageNumber="20">Remarks</emphasis>
.
<taxonomicName id="F4B914BD6C3D2C6E4558FA9CFDC62916" box="[330,586,1291,1316]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3D2C6E4558FA9CFDC62916" box="[330,586,1291,1316]" italics="true" pageId="3" pageNumber="20">Hymenicoides carteri</emphasis>
</taxonomicName>
has not been collected since
<bibRefCitation id="572812CF6C3D2C6E47A3FA9DFBDE2916" author="Kemp" box="[945,1106,1290,1316]" pageId="3" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp (1917)</bibRefCitation>
described the species from near Calcutta,
<collectingCountry id="4BAE2FAE6C3D2C6E4552FAA5FEF0297E" box="[320,380,1330,1356]" name="India" pageId="3" pageNumber="20">India</collectingCountry>
and near Khulna,
<collectingCountry id="4BAE2FAE6C3D2C6E4640FAA5FD52297E" box="[594,734,1330,1356]" name="Bangladesh" pageId="3" pageNumber="20">Bangladesh</collectingCountry>
, on the basis of
<specimenCount id="25BFA4B76C3D2C6E478AFAA5FBB6297E" box="[920,1082,1330,1356]" pageId="3" pageNumber="20" type="generic">22 specimens</specimenCount>
. In this study, we have located only four of the
<typeStatus id="EC02D19C6C3D2C6E454EFACDFE4E2946" box="[348,450,1370,1396]" pageId="3" pageNumber="20" type="syntype">syntypes</typeStatus>
in NHM (Paul Clark, pers. comm.); the rest are presumably still in the Zoological Survey of
<collectingCountry id="4BAE2FAE6C3D2C6E4502FA15FEC029AE" box="[272,332,1410,1436]" name="India" pageId="3" pageNumber="20">India</collectingCountry>
(ex Indian Museum). The G1 of
<taxonomicName id="F4B914BD6C3D2C6E46C2FA14FCCF29AE" box="[720,835,1411,1436]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3D2C6E46C2FA14FCCF29AE" box="[720,835,1411,1436]" italics="true" pageId="3" pageNumber="20">H. carteri</emphasis>
</taxonomicName>
was first described by
<bibRefCitation id="572812CF6C3D2C6E4043FA15FAA529AE" author="Lucas" box="[1105,1321,1410,1436]" pageId="3" pageNumber="31" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas (1980: 197)</bibRefCitation>
, although no figure was provided. The G1 of the
<typeStatus id="EC02D19C6C3D2C6E4649FA3DFD4A29F6" box="[603,710,1450,1476]" pageId="3" pageNumber="20" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="AB8273BB6C3D2C6E46C4FA3DFC9029F6" box="[726,796,1450,1476]" captionStart="FIGURE 2" captionStartId="4.[151,255,1343,1367]" captionTargetBox="[352,1266,221,1298]" captionTargetId="figure@4.[294,1293,197,1317]" captionTargetPageId="4" captionText="FIGURE 2. Male first and second gonopods of Hymenicoides carteri Kemp, 1917. Lectotype, male (NHM 1919.11.1.121), 4.0 × 4.8 mm. a, left G 1, ventral view; b, left G 1, outer view; c, left G 2. Scale bar, 1 mm." httpUri="https://zenodo.org/record/179197/files/figure.png" pageId="3" pageNumber="20">Fig. 2</figureCitation>
a, b, NHM, 4.0 ×
<quantity id="F441C2DB6C3D2C6E47FAFA3DFBCF29F6" box="[1000,1091,1450,1476]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="20" unit="mm" value="4.8">4.8 mm</quantity>
) is very broad, bent outwards medially and with a line of long stiff median setae along the inner part of the ventral side. The distal part of the G1 is a complicated structure; composed of a bifurcated, U-shaped projection on the distal inner angle, a denticulate lip-like margin on the distal outer angle and a long bursiform projection on the dorsal side of the base of the distal outer angle.
<emphasis id="01CDB32C6C3D2C6E45A8F9DCFD882A56" box="[442,516,1611,1636]" italics="true" pageId="3" pageNumber="20">In situ</emphasis>
, the G1 sits completely in the deep sterno-abdominal cavity (
<figureCitation id="AB8273BB6C3D2C6E40C7F9DDFA942A56" box="[1237,1304,1610,1636]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
a).
</paragraph>
<paragraph id="33066F3E6C3D2C6E44D4F9E5FADF2B96" blockId="3.[151,1437,1050,1956]" pageId="3" pageNumber="20">
Of the three recent specimens of
<taxonomicName id="F4B914BD6C3D2C6E4673F9E4FD552ABE" box="[609,729,1651,1676]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3D2C6E4673F9E4FD552ABE" box="[609,729,1651,1676]" italics="true" pageId="3" pageNumber="20">H. carteri</emphasis>
</taxonomicName>
from
<collectingCountry id="4BAE2FAE6C3D2C6E473AF9E5FC382ABE" box="[808,948,1650,1676]" name="Bangladesh" pageId="3" pageNumber="20">Bangladesh</collectingCountry>
, the only male specimen is small (ZRC 2007.0107, 2.2 ×
<quantity id="F441C2DB6C3D2C6E4575F90DFE4F2A86" box="[359,451,1690,1716]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="20" unit="mm" value="2.4">2.4 mm</quantity>
) but it is basically similar to the
<typeStatus id="EC02D19C6C3D2C6E4757F90DFCFB2A86" box="[837,887,1690,1716]" pageId="3" pageNumber="20">type</typeStatus>
series of
<taxonomicName id="F4B914BD6C3D2C6E47FBF90CFBD12A86" box="[1001,1117,1691,1716]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3D2C6E47FBF90CFBD12A86" box="[1001,1117,1691,1716]" italics="true" pageId="3" pageNumber="20">H. carteri</emphasis>
</taxonomicName>
(NHM 1919.11.1.121123 3.0 × 3.7 - 4.0 ×
<quantity id="F441C2DB6C3D2C6E454BF955FE382AEE" box="[345,436,1730,1756]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="20" unit="mm" value="4.8">4.8 mm</quantity>
) except for the G1 and the chela. The G1 is present but is still relatively undeveloped. The G1 of the
<typeStatus id="EC02D19C6C3D2C6E4553F97DFE202B36" box="[321,428,1770,1796]" pageId="3" pageNumber="20" type="lectotype">lectotype</typeStatus>
specimen (NHM 1919.11.1.121, 4.0 ×
<quantity id="F441C2DB6C3D2C6E476BF97DFC582B36" box="[889,980,1770,1796]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="20" unit="mm" value="4.8">4.8 mm</quantity>
) possesses a long bursiform projection on the dorsal side of the base of the distal outer angle (
<figureCitation id="AB8273BB6C3D2C6E4701F885FCD62B1E" box="[787,858,1810,1836]" captionStart="FIGURE 2" captionStartId="4.[151,255,1343,1367]" captionTargetBox="[352,1266,221,1298]" captionTargetId="figure@4.[294,1293,197,1317]" captionTargetPageId="4" captionText="FIGURE 2. Male first and second gonopods of Hymenicoides carteri Kemp, 1917. Lectotype, male (NHM 1919.11.1.121), 4.0 × 4.8 mm. a, left G 1, ventral view; b, left G 1, outer view; c, left G 2. Scale bar, 1 mm." httpUri="https://zenodo.org/record/179197/files/figure.png" pageId="3" pageNumber="20">Fig. 2</figureCitation>
b), which is not observed in the small male
<collectingCountry id="4BAE2FAE6C3D2C6E4146F885FF6E2B66" name="Bangladesh" pageId="3" pageNumber="20">Bangladesh</collectingCountry>
specimen. The cheliped of the
<collectingCountry id="4BAE2FAE6C3D2C6E464BF8ADFD6E2B66" box="[601,738,1850,1876]" name="Bangladesh" pageId="3" pageNumber="20">Bangladesh</collectingCountry>
male specimen is proportionally small and the tubercle on the manus is very low. These differences, however, are possibly associated with size and growth. Larger series of the specimens from
<collectingCountry id="4BAE2FAE6C3D2C6E45B1F81DFDA02B96" box="[419,556,1930,1956]" name="Bangladesh" pageId="3" pageNumber="20">Bangladesh</collectingCountry>
may help to better understand the degree of variation in this species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="E3E337F0368876700235A9B628C94614" ID-DOI="10.5281/zenodo.179195" ID-GBIF-Dataset="e0a7d367-be4e-4031-acad-42a17507fadf" ID-ISSN="1175-5326" ID-Zenodo-Dep="179195" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459941676950" checkinUser="plazi" docAuthor="Naruse, Tohru &amp; Ng, Peter K. L." docDate="2007" docId="BB10DE286C3F2C6E4485FC3DFE6E2ED0" docLanguage="en" docName="zt01621p031.pdf" docOrigin="Zootaxa 1621" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hymenicoides Kemp 1917" docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="4729A6506C3E2C6D4412FF97FF8C2C32" masterDocTitle="On the taxonomy of the genus Hymenicoides Kemp, 1917 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), with resurrection of Limnopilos Chuang &amp; Ng, 1991, and descriptions of two new species" masterLastPageNumber="31" masterPageNumber="17" pageNumber="18" updateTime="1698230522269" updateUser="plazi">
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<mods:title id="3688D8354B31A6878C964CB4CE0C4400">On the taxonomy of the genus Hymenicoides Kemp, 1917 (Crustacea: Decapoda: Brachyura: Hymenosomatidae), with resurrection of Limnopilos Chuang &amp; Ng, 1991, and descriptions of two new species</mods:title>
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<mods:namePart id="75E54257CDB3046BD3C54C1B0E5FF315">Naruse, Tohru</mods:namePart>
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<mods:namePart id="0241820A13DB152AF9A10A001A3C1DBD">Ng, Peter K. L.</mods:namePart>
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<mods:date id="B273F5E3B5640EA18D9FCCC5A208B8F3">2007</mods:date>
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<subSubSection id="7BA33CB56C3F2C6C4485FC3DFD1E281E" pageId="1" pageNumber="18" type="nomenclature">
<paragraph id="33066F3E6C3F2C6C4485FC3DFE522FF6" blockId="1.[151,478,882,964]" box="[151,478,938,964]" pageId="1" pageNumber="18">
<heading id="684ED8526C3F2C6C4485FC3DFE522FF6" bold="true" box="[151,478,938,964]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
<taxonomicName id="F4B914BD6C3F2C6C4485FC3DFE522FF6" ID-CoL="7PL39" authority="Kemp, 1917" authorityName="Kemp" authorityYear="1917" box="[151,478,938,964]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3F2C6C4485FC3DFE522FF6" bold="true" box="[151,478,938,964]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C4485FC3DFECE2FF6" bold="true" box="[151,322,938,964]" italics="true" pageId="1" pageNumber="18">Hymenicoides</emphasis>
<bibRefCitation id="572812CF6C3F2C6C455BFC3DFE522FF6" author="Kemp" box="[329,478,938,964]" pageId="1" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="33066F3E6C3F2C6C4485FC62FD1E281E" blockId="1.[151,1436,1013,1068]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C4485FC62FECA283E" box="[151,326,1013,1036]" italics="true" pageId="1" pageNumber="18">Hymenicoides—</emphasis>
<bibRefCitation id="572812CF6C3F2C6C455EFC62FE58283E" author="Kemp" box="[332,468,1013,1036]" pageId="1" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
: 267;
<bibRefCitation id="572812CF6C3F2C6C4607FC62FD17283E" author="Lucas" box="[533,667,1013,1036]" pageId="1" pageNumber="31" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas, 1980</bibRefCitation>
: 196;
<bibRefCitation id="572812CF6C3F2C6C46CEFC62FC35283E" author="Ng" box="[732,953,1013,1036]" pageId="1" pageNumber="31" refString="Ng, P. K. L. &amp; Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement, 3, 1 - 82." type="journal article" year="1996">Ng &amp; Chuang, 1996</bibRefCitation>
: 50 (part); Guinot &amp; Richer de
<bibRefCitation id="572812CF6C3F2C6C411BFC62FA14283E" author="Forges" box="[1289,1432,1013,1036]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
: 460 (part);
<bibRefCitation id="572812CF6C3F2C6C452DFB82FDD5281E" author="Guinot" box="[319,601,1045,1068]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Bouchard, J. - M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20 (4), 613 - 694." type="journal article" year="1998">Guinot &amp; Bouchard, 1998</bibRefCitation>
: 685.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3F2C6C4485FBCAFB5C2845" box="[151,1232,1117,1143]" pageId="1" pageNumber="18" type="materials_examined">
<paragraph id="33066F3E6C3F2C6C4485FBCAFB5C2845" blockId="1.[151,1437,1117,2023]" box="[151,1232,1117,1143]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C4485FBCAFEBE2845" bold="true" box="[151,306,1117,1143]" pageId="1" pageNumber="18">
<typeStatus id="EC02D19C6C3F2C6C4485FBCAFF582845" box="[151,212,1117,1143]" pageId="1" pageNumber="18">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="F4B914BD6C3F2C6C4552FBCAFD542845" authority="Kemp, 1917" authorityName="Kemp" authorityYear="1917" box="[320,728,1117,1143]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3F2C6C4552FBCAFDB12844" box="[320,573,1117,1142]" italics="true" pageId="1" pageNumber="18">Hymenicoides carteri</emphasis>
<bibRefCitation id="572812CF6C3F2C6C4657FBCAFD542845" author="Kemp" box="[581,728,1117,1143]" pageId="1" pageNumber="31" refString="Kemp, S. (1917) Notes on Crustacea Decapoda in the Indian Museum. X. Hymenosomatidae. Records of the Indian Museum, 13, 243 - 279." type="journal article" year="1917">Kemp, 1917</bibRefCitation>
</taxonomicName>
, by monotypy; gender of genus masculine.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3F2C6C44D4FB12FE702A4D" pageId="1" pageNumber="18" type="diagnosis">
<paragraph id="33066F3E6C3F2C6C44D4FB12FE702A4D" blockId="1.[151,1437,1117,2023]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C44D4FB12FECA28AD" bold="true" box="[198,326,1157,1183]" pageId="1" pageNumber="18">Diagnosis.</emphasis>
Carapace oval, dorsal surface concave; grooves distinct; rostrum absent or very weak; antenna with base of basal article placed posterior-inner part of base of eye; eyes, antennae, antennules visible dorsally; third maxillipeds narrow, not covering more than three-quarters of mouth field when closed, merus rectangular, dactylus styliform, approximately twice length of propodus; male chelae relatively stout, manus with high, prominent, dorso-ventrally compressed tubercle on distal part of outer surface, tubercle absent in females. Female vulvae placed on imaginary line joining inner ends of sutures between sternites 5 and 6 on medially fused plate of thoracic sternum, vulva with basal mount. Male abdomen-pleotelson without fused segments, male pleotelson distinctly trilobed, inner surface thickened externally, forming socket for sternal button (
<figureCitation id="AB8273BB6C3F2C6C44E1FA52FEBB29ED" box="[243,311,1477,1503]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="1" pageNumber="18">Fig. 1</figureCitation>
a). G1 stout, bent outwards medially; distal part with prominent distal inner processes, swollen distal outer angle, long, dorsal bursiform projections. Female abdomen with 6 distinctly demarcated segments, boundary between first and second segments (
<taxonomicName id="F4B914BD6C3F2C6C470BF982FC032A1C" box="[793,911,1557,1582]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3F2C6C470BF982FC032A1C" box="[793,911,1557,1582]" italics="true" pageId="1" pageNumber="18">H. carteri</emphasis>
</taxonomicName>
) or between second and third segments (
<taxonomicName id="F4B914BD6C3F2C6C416DF982FF7E2A64" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="robertsi">
<emphasis id="01CDB32C6C3F2C6C416DF982FF7E2A64" italics="true" pageId="1" pageNumber="18">H. robertsi</emphasis>
</taxonomicName>
) movable; long, biramous pleopods on second to fifth segments, developed from distal outer end of inner surface of each segment.
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3F2C6F44D4F91AFBFF2DF8" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" type="discussion">
<paragraph id="33066F3E6C3F2C6C44D4F91AFDF62BA5" blockId="1.[151,1437,1117,2023]" pageId="1" pageNumber="18">
<emphasis id="01CDB32C6C3F2C6C44D4F91AFEB42A95" bold="true" box="[198,312,1677,1703]" pageId="1" pageNumber="18">Remarks</emphasis>
. The distinctive trilobate pleotelson of
<taxonomicName id="F4B914BD6C3F2C6C4714F91AFCF62A94" box="[774,890,1677,1702]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3F2C6C4714F91AFCF62A94" box="[774,890,1677,1702]" italics="true" pageId="1" pageNumber="18">H. carteri</emphasis>
</taxonomicName>
has received some attention from taxonomists due to its unusual shape.
<bibRefCitation id="572812CF6C3F2C6C45D5F922FD2E2AFD" author="Lucas" box="[455,674,1717,1743]" pageId="1" pageNumber="31" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33 (4), 148 - 247." type="journal article" year="1980">Lucas (1980: 197)</bibRefCitation>
regarded that the lateral lobe of male pleotelson as “a cavity to receive the apice of the first pleopod”. Guinot &amp; Richer de
<bibRefCitation id="572812CF6C3F2C6C4746F94AFBBA2AC5" author="Forges" box="[852,1078,1757,1783]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges (1997: 470)</bibRefCitation>
suggested that the lateral lobe of the pleotelson may be derived from the inserted plate (see Guinot &amp; Richer de
<bibRefCitation id="572812CF6C3F2C6C4067F892FA992B2D" author="Forges" box="[1141,1301,1797,1823]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
:
<figureCitation id="AB8273BB6C3F2C6C4130F892FAE62B2D" box="[1314,1386,1797,1823]" captionStart="FIGURE 6" captionStartId="8.[151,255,1506,1530]" captionTargetBox="[151,1436,184,1485]" captionTargetId="figure@8.[151,1436,184,1485]" captionTargetPageId="8" captionText="FIGURE 6. Hymenicoides robertsi new species. a d, right second to fifth pleopods; e, exopods. Paratype, ovigerous female (ZRC 2007.0113), 4.2 × 5.0 mm." httpUri="https://zenodo.org/record/179201/files/figure.png" pageId="1" pageNumber="18">Fig. 6</figureCitation>
, for mobile intercalated platelet of
<taxonomicName id="F4B914BD6C3F2C6C461BF8BAFD682B74" box="[521,740,1837,1862]" class="Malacostraca" family="Hymenosomatidae" genus="Odiomaris" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="01CDB32C6C3F2C6C461BF8BAFD682B74" box="[521,740,1837,1862]" italics="true" pageId="1" pageNumber="18">Odiomaris pilosus</emphasis>
</taxonomicName>
).
<bibRefCitation id="572812CF6C3F2C6C4712F8BAFB0A2B75" author="Guinot" box="[768,1158,1837,1863]" pageId="1" pageNumber="31" refString="Guinot, D. &amp; Bouchard, J. - M. (1998) Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema, 20 (4), 613 - 694." type="journal article" year="1998">Guinot &amp; Bouchard (1998: 685)</bibRefCitation>
subsequently hypothesized that the abdominal socket, which is placed on the inner surface of the lateral lobe (or the intercalated platelet), is homologous with the uropod.
</paragraph>
<paragraph id="33066F3E6C3F2C6F44D4F832FBFF2DF8" blockId="1.[151,1437,1117,2023]" lastBlockId="2.[151,1436,152,458]" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
Our examination of the
<typeStatus id="EC02D19C6C3F2C6C45F3F832FD9F2B8D" box="[481,531,1957,1983]" pageId="1" pageNumber="18">type</typeStatus>
material of
<taxonomicName id="F4B914BD6C3F2C6C46B1F832FC942B8C" box="[675,792,1957,1982]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3F2C6C46B1F832FC942B8C" box="[675,792,1957,1982]" italics="true" pageId="1" pageNumber="18">H. carteri</emphasis>
</taxonomicName>
and
<taxonomicName id="F4B914BD6C3F2C6C4740F832FC5E2B8C" box="[850,978,1957,1982]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C3F2C6C4740F832FC5E2B8C" box="[850,978,1957,1982]" italics="true" pageId="1" pageNumber="18">H. robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3F2C6C47C8F832FBE62B8D" bold="true" box="[986,1130,1957,1983]" pageId="1" pageNumber="18">
<taxonomicNameLabel id="1AFE0E576C3F2C6C47C8F832FBE62B8D" box="[986,1130,1957,1983]" pageId="1" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
, shows that the inner surface of the lateral lobe of the male pleotelson is domed and that it is externally surrounded by a thickened rim, forming a semicircular cavity (
<figureCitation id="AB8273BB6C3C2C6F461AFF0FFDC12C80" box="[520,589,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="2" pageNumber="19">Fig. 1</figureCitation>
b). When the abdomen is closed, the cavity overlays the sternal button (
<figureCitation id="AB8273BB6C3C2C6F448DFF57FF6A2CE8" box="[159,230,192,218]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="2" pageNumber="19">Fig. 1</figureCitation>
a). This does not support the premise that the lateral lobe of the pleotelson is to accommodate the unusual tip of the G1 (
<figureCitation id="AB8273BB6C3C2C6F458EFF7FFE622D30" box="[412,494,232,258]" captionStart="FIGURE 2" captionStartId="4.[151,255,1343,1367]" captionTargetBox="[352,1266,221,1298]" captionTargetId="figure@4.[294,1293,197,1317]" captionTargetPageId="4" captionText="FIGURE 2. Male first and second gonopods of Hymenicoides carteri Kemp, 1917. Lectotype, male (NHM 1919.11.1.121), 4.0 × 4.8 mm. a, left G 1, ventral view; b, left G 1, outer view; c, left G 2. Scale bar, 1 mm." httpUri="https://zenodo.org/record/179197/files/figure.png" pageId="2" pageNumber="19">Figs. 2</figureCitation>
a, b, 5c, d). In fact, even when the abdomen is open, the G1 can easily be accommodated within the sterno-abdominal cavity (
<figureCitation id="AB8273BB6C3C2C6F46BDFE87FD7E2D18" box="[687,754,272,298]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="2" pageNumber="19">Fig. 1</figureCitation>
a). The sternal buttons and the lateral cavities of the pleotelson function instead to “lock” the abdomen with the thorax. It should be noted that the G1 of
<taxonomicName id="F4B914BD6C3C2C6F4134FEAFFA102D63" box="[1318,1436,312,337]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3C2C6F4134FEAFFA102D63" box="[1318,1436,312,337]" italics="true" pageId="2" pageNumber="19">H. carteri</emphasis>
</taxonomicName>
closely resembles that of
<taxonomicName id="F4B914BD6C3C2C6F45DAFEF7FD732D4B" box="[456,767,352,377]" class="Malacostraca" family="Hymenosomatidae" genus="Cancrocaeca" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="xenomorpha">
<emphasis id="01CDB32C6C3C2C6F45DAFEF7FD732D4B" box="[456,767,352,377]" italics="true" pageId="2" pageNumber="19">Cancrocaeca xenomorpha</emphasis>
</taxonomicName>
(see also Guinot &amp; Richer de
<bibRefCitation id="572812CF6C3C2C6F4062FEF7FA822D48" author="Forges" box="[1136,1294,352,378]" pageId="2" pageNumber="31" refString="Guinot, D. &amp; Richer de Forges, B. (1997) Affinites entre les Hymenosomatidae MacLeay ' 1838 et les Inachoididae Dana ' 1851 (Crustacea ' Decapoda ' Brachyura). Zoosystema, 19 (2 / 3), 453 - 502." type="journal article" year="1997">Forges, 1997</bibRefCitation>
), and in the latter species, the male abdomen is more typical in shape, lacking the trilobite pleotelson, as well as sternal buttons, seen in
<taxonomicName id="F4B914BD6C3C2C6F4547FE27FE772DFB" box="[341,507,432,457]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3C2C6F4547FE27FE772DFB" box="[341,507,432,457]" italics="true" pageId="2" pageNumber="19">Hymenicoides</emphasis>
</taxonomicName>
(Naruse, T., Ng, P.K.L. &amp; Guinot, D., in manuscript).
</paragraph>
</subSubSection>
<caption id="67C63FB66C3C2C6F4485F8EAFB212BC7" httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="2" pageNumber="19" targetBox="[268,1318,501,1901]" targetPageId="2">
<paragraph id="33066F3E6C3C2C6F4485F8EAFB212BC7" blockId="2.[151,1436,1917,2037]" pageId="2" pageNumber="19">
<emphasis id="01CDB32C6C3C2C6F4485F8EAFE952BA7" bold="true" box="[151,281,1917,1941]" pageId="2" pageNumber="19">FIGURE 1.</emphasis>
Male abdominal cavity and pleotelson of
<taxonomicName id="F4B914BD6C3C2C6F46CFF8E9FC5C2BA7" box="[733,976,1918,1941]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C3C2C6F46CFF8E9FC5C2BA7" box="[733,976,1918,1941]" italics="true" pageId="2" pageNumber="19">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3C2C6F47C4F8EAFBD52BA7" bold="true" box="[982,1113,1917,1941]" pageId="2" pageNumber="19">
<taxonomicNameLabel id="1AFE0E576C3C2C6F47C4F8EAFBD52BA7" box="[982,1113,1917,1941]" pageId="2" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="F4B914BD6C3C2C6F4086F8E9FA802BA7" box="[1172,1292,1918,1941]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3C2C6F4086F8E9FA802BA7" box="[1172,1292,1918,1941]" italics="true" pageId="2" pageNumber="19">Limnopilos</emphasis>
</taxonomicName>
species. a, b,
<taxonomicName id="F4B914BD6C3C2C6F4485F809FE042B87" box="[151,392,1950,1973]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="robertsi" status="sp. nov.">
<emphasis id="01CDB32C6C3C2C6F4485F809FE042B87" box="[151,392,1950,1973]" italics="true" pageId="2" pageNumber="19">Hymenicoides robertsi</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3C2C6F4583F80AFD992B87" bold="true" box="[401,533,1949,1973]" pageId="2" pageNumber="19">
<taxonomicNameLabel id="1AFE0E576C3C2C6F4583F80AFD992B87" box="[401,533,1949,1973]" pageId="2" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
(paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d,
<taxonomicName id="F4B914BD6C3C2C6F403FF809FA102B87" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[1069,1436,1950,1973]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C3C2C6F403FF809FB352B87" box="[1069,1209,1950,1973]" italics="true" pageId="2" pageNumber="19">L. naiyanetri</emphasis>
Chuang &amp; Ng, 1991
</taxonomicName>
(ZRC 1991.65216555, 4.9 × 5.3 mm); e, f,
<taxonomicName id="F4B914BD6C3C2C6F4666F829FC802BE7" box="[628,780,1982,2005]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C3C2C6F4666F829FC802BE7" box="[628,780,1982,2005]" italics="true" pageId="2" pageNumber="19">L. sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3C2C6F4706F82AFC1B2BE7" bold="true" box="[788,919,1981,2005]" pageId="2" pageNumber="19">
<taxonomicNameLabel id="1AFE0E576C3C2C6F4706F82AFC1B2BE7" box="[788,919,1981,2005]" pageId="2" pageNumber="19" rank="species">new species</taxonomicNameLabel>
</emphasis>
(paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G1; b, d, f, inner surface of male pleotelson.
</paragraph>
</caption>
<subSubSection id="7BA33CB56C3D2C6E44D4FF0FFBCB2EA0" pageId="3" pageNumber="20" type="reference_group">
<paragraph id="33066F3E6C3D2C6E44D4FF0FFC552DC0" blockId="3.[151,1437,152,738]" pageId="3" pageNumber="20">
<taxonomicName id="F4B914BD6C3D2C6E44D4FF0FFD442C80" ID-CoL="72CL4" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[198,712,152,178]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="naiyanetri">
<emphasis id="01CDB32C6C3D2C6E44D4FF0FFE402C83" box="[198,460,152,177]" italics="true" pageId="3" pageNumber="20">Limnopilos naiyanetri</emphasis>
<bibRefCitation id="572812CF6C3D2C6E45C6FF0FFD442C80" author="Chuang" box="[468,712,152,178]" pageId="3" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
</taxonomicName>
(
<typeStatus id="EC02D19C6C3D2C6E46CAFF0FFC802C80" box="[728,780,152,178]" pageId="3" pageNumber="20">type</typeStatus>
species of
<taxonomicName id="F4B914BD6C3D2C6E4786FF0FFA942C80" ID-CoL="7PMTG" authority="Chuang &amp; Ng, 1991" authorityName="Chuang &amp; Ng" authorityYear="1991" box="[916,1304,152,178]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E4786FF0FFB942C83" box="[916,1048,152,177]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
<bibRefCitation id="572812CF6C3D2C6E4033FF0FFA942C80" author="Chuang" box="[1057,1304,152,178]" pageId="3" pageNumber="31" refString="Chuang, C. T. N. &amp; Ng, P. K. L. (1991) Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39 (2), 363 - 368." type="journal article" year="1991">Chuang &amp; Ng, 1991</bibRefCitation>
</taxonomicName>
),
<taxonomicName id="F4B914BD6C3D2C6E413CFF0FFE062CE8" ID-CoL="8WB9Z" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="microrhynchus" status="comb. nov.">
<emphasis id="01CDB32C6C3D2C6E413CFF0FFE8E2CEB" italics="true" pageId="3" pageNumber="20">L. microrhynchus</emphasis>
(Ng, 1995)
</taxonomicName>
<taxonomicNameLabel id="1AFE0E576C3D2C6E4580FF57FDED2CE8" box="[402,609,192,218]" pageId="3" pageNumber="20">new combination</taxonomicNameLabel>
, and
<taxonomicName id="F4B914BD6C3D2C6E468CFF57FCCB2CEB" ID-CoL="6QCKT" box="[670,839,192,217]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="sumatranus" status="sp. nov.">
<emphasis id="01CDB32C6C3D2C6E468CFF57FCCB2CEB" box="[670,839,192,217]" italics="true" pageId="3" pageNumber="20">L. sumatranus</emphasis>
</taxonomicName>
<emphasis id="01CDB32C6C3D2C6E475CFF28FC522CEB" bold="true" box="[846,990,191,217]" pageId="3" pageNumber="20">
<taxonomicNameLabel id="1AFE0E576C3D2C6E475CFF28FC522CEB" box="[846,990,191,217]" pageId="3" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</emphasis>
, all possess the sternal button and the pleotelson lateral cavity (
<figureCitation id="AB8273BB6C3D2C6E45ADFF7FFD8C2D30" box="[447,512,232,258]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
cf). The lateral cavities of these three species, however, are relatively smaller and the rim is more distal in position on the cavity when compared with those of
<taxonomicName id="F4B914BD6C3D2C6E4071FE87FB542D1B" ID-CoL="3NJ2G" box="[1123,1240,272,297]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="carteri">
<emphasis id="01CDB32C6C3D2C6E4071FE87FB542D1B" box="[1123,1240,272,297]" italics="true" pageId="3" pageNumber="20">H. carteri</emphasis>
</taxonomicName>
and
<taxonomicName id="F4B914BD6C3D2C6E4106FE87FA182D1B" ID-CoL="3NJ2J" box="[1300,1428,272,297]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="robertsi">
<emphasis id="01CDB32C6C3D2C6E4106FE87FA182D1B" box="[1300,1428,272,297]" italics="true" pageId="3" pageNumber="20">H. robertsi</emphasis>
</taxonomicName>
. This may be due to the different shape of the G1 of
<taxonomicName id="F4B914BD6C3D2C6E46ECFEAFFC282D63" ID-CoL="7PL39" box="[766,932,312,337]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E46ECFEAFFC282D63" box="[766,932,312,337]" italics="true" pageId="3" pageNumber="20">Hymenicoides</emphasis>
</taxonomicName>
. The lateral cavity of
<taxonomicName id="F4B914BD6C3D2C6E40B9FEAFFADD2D63" ID-CoL="7PL39" box="[1195,1361,312,337]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E40B9FEAFFADD2D63" box="[1195,1361,312,337]" italics="true" pageId="3" pageNumber="20">Hymenicoides</emphasis>
</taxonomicName>
needs to be thickened along the external margins to effectively engage the sternal button, three sides (anterior, inner and posterior) which are surrounded by the G
<quantity id="F441C2DB6C3D2C6E46A9FE1FFD6A2D93" box="[699,742,392,418]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="20" unit="in" value="1.0">
1
<emphasis id="01CDB32C6C3D2C6E46DDFE1FFD6A2D93" box="[719,742,392,417]" italics="true" pageId="3" pageNumber="20">in</emphasis>
</quantity>
situ (
<figureCitation id="AB8273BB6C3D2C6E4739FE1FFCE32D90" box="[811,879,392,418]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
a). However, the sternal buttons of
<taxonomicName id="F4B914BD6C3D2C6E410AFE1FFA102D93" ID-CoL="7PMTG" box="[1304,1436,392,417]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E410AFE1FFA102D93" box="[1304,1436,392,417]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
are placed more distantly from the distal outer angle of the G1 (
<figureCitation id="AB8273BB6C3D2C6E4769FE27FC312DF8" box="[891,957,432,458]" captionStart="FIGURE 1" captionStartId="2.[151,255,1917,1941]" captionTargetBox="[268,1318,501,1901]" captionTargetId="figure@2.[268,1319,499,1902]" captionTargetPageId="2" captionText="FIGURE 1. Male abdominal cavity and pleotelson of Hymenicoides robertsi new species, and Limnopilos species. a, b, Hymenicoides robertsi new species (paratype, ZRC 2007.0109, 5.0 × 6.2 mm); c, d, L. naiyanetri Chuang &amp; Ng, 1991 (ZRC 1991.6521 6555, 4.9 × 5.3 mm); e, f, L. sumatranus new species (paratype, ZRC 2007.0114, 5.0 × 5.9 mm). a, c, e, male abdominal cavity with sternal button (sb) and G 1; b, d, f, inner surface of male pleotelson." httpUri="https://zenodo.org/record/179196/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
c, e), and allow the lateral cavity to more easily lock onto the sternal button by way of the distally thickened rim.
</paragraph>
<paragraph id="33066F3E6C3D2C6E44D4FD97FBCB2EA0" blockId="3.[151,1437,152,738]" pageId="3" pageNumber="20">
The G1s of the
<taxonomicName id="F4B914BD6C3D2C6E4594FD97FDA02E2B" ID-CoL="7PL39" box="[390,556,512,537]" class="Malacostraca" family="Hymenosomatidae" genus="Hymenicoides" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E4594FD97FDA02E2B" box="[390,556,512,537]" italics="true" pageId="3" pageNumber="20">Hymenicoides</emphasis>
</taxonomicName>
species (
<figureCitation id="AB8273BB6C3D2C6E468EFD97FD692E28" box="[668,741,512,538]" captionStart="FIGURE 2" captionStartId="4.[151,255,1343,1367]" captionTargetBox="[352,1266,221,1298]" captionTargetId="figure@4.[294,1293,197,1317]" captionTargetPageId="4" captionText="FIGURE 2. Male first and second gonopods of Hymenicoides carteri Kemp, 1917. Lectotype, male (NHM 1919.11.1.121), 4.0 × 4.8 mm. a, left G 1, ventral view; b, left G 1, outer view; c, left G 2. Scale bar, 1 mm." httpUri="https://zenodo.org/record/179197/files/figure.png" pageId="3" pageNumber="20">Fig. 2</figureCitation>
a, b, 5c, d) differ from those of
<taxonomicName id="F4B914BD6C3D2C6E4075FD97FB672E2B" ID-CoL="7PMTG" box="[1127,1259,512,537]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E4075FD97FB672E2B" box="[1127,1259,512,537]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
in several key aspects: they have a proportionately stouter shaft (vs. moderately stout in
<taxonomicName id="F4B914BD6C3D2C6E47EEFDBFFB0C2E73" ID-CoL="7PMTG" box="[1020,1152,552,577]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E47EEFDBFFB0C2E73" box="[1020,1152,552,577]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
), more prominent distal inner processes (vs. moderately stout in
<taxonomicName id="F4B914BD6C3D2C6E467EFDC7FD7C2E5B" ID-CoL="7PMTG" box="[620,752,592,617]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E467EFDC7FD7C2E5B" box="[620,752,592,617]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
) and with the distal outer angle more swollen than in
<taxonomicName id="F4B914BD6C3D2C6E4176FDC7FF612EA3" ID-CoL="7PMTG" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E4176FDC7FF612EA3" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
species (
<figureCitation id="AB8273BB6C3D2C6E454BFDEFFE122EA0" box="[345,414,632,658]" captionStart="FIGURE 8" captionStartId="12.[151,255,1467,1491]" captionTargetBox="[260,1306,186,1428]" captionTargetId="figure@12.[260,1326,172,1446]" captionTargetPageId="12" captionText="FIGURE 8. Limnopilos sumatranus new species. a, male abdomen-pleotelson; b, female abdomen-pleotelson; c, left G 1; d, left G 2. a, paratype male (ZRC 2007.0114), 5.0 × 5.9 mm; b, paratype female (MNHN-B 30395), 5.4 × 6.3 mm; c, d, holotype male (MZB Cru 1650), 4.7 × 5.5 mm. Scales, 1 mm." httpUri="https://zenodo.org/record/179203/files/figure.png" pageId="3" pageNumber="20">Fig. 8</figureCitation>
c;
<bibRefCitation id="572812CF6C3D2C6E45A8FDEFFD212EA0" author="Ng" box="[442,685,632,658]" pageId="3" pageNumber="31" refString="Ng, P. K. L. &amp; Chuang, C. T. N. (1996) The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement, 3, 1 - 82." type="journal article" year="1996">Ng &amp; Chuang, 1996</bibRefCitation>
: Fig. 21H; Ng, 1995: Fig. 14A, B).
</paragraph>
</subSubSection>
<subSubSection id="7BA33CB56C3D2C6E44D4FD37FE6E2ED0" pageId="3" pageNumber="20" type="description">
<paragraph id="33066F3E6C3D2C6E44D4FD37FE6E2ED0" blockId="3.[151,1437,152,738]" pageId="3" pageNumber="20">
The marked differences in the chela, abdomen and G1 lead us to now conclude that
<taxonomicName id="F4B914BD6C3D2C6E408DFD37FAAF2E8B" box="[1183,1315,672,697]" class="Malacostraca" family="Hymenosomatidae" genus="Limnopilos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="01CDB32C6C3D2C6E408DFD37FAAF2E8B" box="[1183,1315,672,697]" italics="true" pageId="3" pageNumber="20">Limnopilos</emphasis>
</taxonomicName>
should be resurrected as a valid genus.
</paragraph>
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</treatment>
</document>

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<paragraph pageId="45" pageNumber="46">Taxon classification Animalia Dactylogyridea Dactylogyridae</paragraph>
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<taxonomicName authority="Moreira, Scholz &amp; Luque, 2015" authorityName="Moreira, Scholz &amp; Luque" authorityYear="2015" class="Monogenea" family="Dactylogyridae" genus="Urocleidoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Urocleidoides xinguensis" order="Dactylogyridea" pageId="45" pageNumber="46" phylum="Platyhelminthes" rank="species" species="xinguensis">Urocleidoides xinguensis Moreira, Scholz &amp; Luque, 2015</taxonomicName>
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<paragraph pageId="45" pageNumber="46">Type host.</paragraph>
<paragraph pageId="45" pageNumber="46">
<taxonomicName class="Actinopterygii" family="Erythrinidae" genus="Hoplias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoplias aimara" order="Characiformes" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="aimara">Hoplias aimara</taxonomicName>
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<paragraph pageId="45" pageNumber="46">Infection site.</paragraph>
<paragraph pageId="45" pageNumber="46">Gills.</paragraph>
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<paragraph pageId="45" pageNumber="46">Type locality.</paragraph>
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Brazil,
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State, Xingu River around Altamira (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-3.2">3°12'S</geoCoordinate>
,
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).
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<paragraph pageId="45" pageNumber="46">Holotype.</paragraph>
<paragraph pageId="45" pageNumber="46">CHIOC 38002 a.</paragraph>
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<paragraph pageId="45" pageNumber="46">Paratypes.</paragraph>
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CHIOC 38002
<normalizedToken originalValue="bd">b-d</normalizedToken>
.
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<subSubSection pageId="45" pageNumber="46" type="remarks">
<paragraph pageId="45" pageNumber="46">Remarks.</paragraph>
<paragraph pageId="45" pageNumber="46">Two paratypes deposited in the IPCAS collection.</paragraph>
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<paragraph pageId="45" pageNumber="46">Reference.</paragraph>
<paragraph pageId="45" pageNumber="46">
<bibRefCitation author="Moreira, J" journalOrPublisher="Acta Parasitologica" pageId="69" pageNumber="70" pagination="254 - 260" title="First data on the parasites of Hopliasaimara (Characiformes): description of two new species of gill monogeneans (Dactylogyridae)." url="10.1515/ap-2015-0036" volume="60" year="2015">Moreira et al. (2015)</bibRefCitation>
.
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<taxonomicName id="06971828441AA2D6B57071809E57C831" ID-CoL="492YH" ID-ENA="497257" LSID="BB10FE82-2FF6-55FE-AD30-C72B27034023" authority="L. f." class="Magnoliopsida" family="Rubiaceae" genus="Oldenlandia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oldenlandia capensis" order="Gentianales" pageId="0" pageNumber="54205" phylum="Tracheophyta" rank="species" species="capensis">Oldenlandia capensis L.f.</taxonomicName>
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<paragraph id="E2862C7FC811CE9DCEFAE83EC3537146" pageId="0" pageNumber="54205">Distribution</paragraph>
<paragraph id="11ED9E5F9CBE7269C8E56636DF7EBE77" pageId="0" pageNumber="54205">Cosmopolitan</paragraph>
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<paragraph id="6FB4C8928C31BFD39A14C329C0AA18AD" pageId="0" pageNumber="54205">Life Form: therophyte</paragraph>
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