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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.423.7536" ID-GBIF-Dataset="1f17876d-2290-4d36-aa1f-10b951701679" ID-PMC="PMC4106094" ID-Pensoft-Pub="1313-2970-423-1" ID-PubMed="25061393" ID-ZBK="55B4F9C858934F88841660FF730E8872" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-423-1" ModsDocOrigin="ZooKeys 423" ModsDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" checkinTime="1451245649604" checkinUser="pensoft" docAuthor="Brunke, Adam J. &amp; Solodovnikov, Alexey" docDate="2014" docId="F8FB13EAB6621D8B480AF3DF6BE50E30" docLanguage="en" docName="ZooKeys 423: 1-113" docOrigin="ZooKeys 423" docSource="http://dx.doi.org/10.3897/zookeys.423.7536" docTitle="Bolitogyrus viridescens Brunke, sp. n." docType="treatment" docUuid="438E36F1-20E8-4350-A8F4-E5202D9277D4" docUuidSource="ZooBank" docVersion="4" lastPageNumber="64" masterDocId="FFAB12065F4CFFED925DFFB46A5FFFFB" masterDocTitle="A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)" masterLastPageNumber="113" masterPageNumber="1" pageNumber="63" updateTime="1668158857111" updateUser="ExternalLinkService">
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<mods:title>A revision of the Neotropical species of Bolitogyrus Chevrolat, a geographically disjunct lineage of Staphylinini (Coleoptera, Staphylinidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Brunke, Adam J.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Solodovnikov, Alexey</mods:namePart>
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<subSubSection pageId="62" pageNumber="63" type="multiple">
<paragraph pageId="62" pageNumber="63">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
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<subSubSection pageId="62" pageNumber="63" type="nomenclature">
<paragraph pageId="62" pageNumber="63">
<taxonomicName LSID="http://zoobank.org/438E36F1-20E8-4350-A8F4-E5202D9277D4" authority="Brunke" class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus viridescens" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="viridescens">Bolitogyrus viridescens Brunke</taxonomicName>
<taxonomicNameLabel pageId="62" pageNumber="63">sp. n.</taxonomicNameLabel>
Figs 2D, 6G, 9F, 10F, 13F, 18
<normalizedToken originalValue="A–E">A-E</normalizedToken>
, 23L, 26I, 31C (map)
</paragraph>
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<subSubSection pageId="62" pageNumber="63" type="type locality">
<paragraph pageId="62" pageNumber="63">Type locality.</paragraph>
<paragraph pageId="62" pageNumber="63">
MEXICO, Puebla, 9.66 km east of
<normalizedToken originalValue="Teziutlán">Teziutlan</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="63" type="type material">
<paragraph pageId="62" pageNumber="63">Type material.</paragraph>
<paragraph pageId="62" pageNumber="63">
Holotype ♂ (CNC): 6mi. East, Tezuitlan, Pueb. MEX., VIII 4-6 [19]60, H.F. Howden [white printed label] / Holotype,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus viridescens" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="viridescens">Bolitogyrus viridescens</taxonomicName>
Brunke, sp. n. [red printed label].
</paragraph>
<paragraph pageId="62" pageNumber="63">Paratypes (2 ♂, 1 ♀, CNC): MEXICO: same data as holotype.</paragraph>
</subSubSection>
<subSubSection pageId="62" pageNumber="63" type="diagnosis">
<paragraph pageId="62" pageNumber="63">Diagnosis.</paragraph>
<paragraph pageId="62" pageNumber="63">Within Bullatus Lineage: elytra with strigulose patch (Fig. 9F); head without distinct posterior protuberances, without large glossy, impunctate areas (Fig. 6G); head without strigulose sculpture (Fig. 6G).</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="description">
<paragraph pageId="63" pageNumber="64">
<pageBreakToken pageId="63" pageNumber="64" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="63" pageNumber="64">Measurements ♂ (n=3): HW/HL 1.38-1.48; PW/PL 1.35-1.43; EW/EL 1.03-1.17; ESut/PL 1.00; PW/HW 1.00; forebody length 3.6-3.7 mm.</paragraph>
<paragraph pageId="63" pageNumber="64">Measurements ♀ (n=1): HW/HL 1.38; PW/PL 1.42; EW/EL 1.09; ESut/PL 0.96; PW/HW 1.03; forebody length 4.1 mm.</paragraph>
<paragraph pageId="63" pageNumber="64">
Similar to
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus strigifrons" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="strigifrons">Bolitogyrus strigifrons</taxonomicName>
and differing only in the following: head, lateral part of pronotum, scutellum and patches of strigulose sculpture on elytra with brilliant greenish-blue metallic reflection, remaining areas on pronotum and elytra with bronze metallic reflections; maxillary and labial palpi yellow; antennomeres
<normalizedToken originalValue="I–V">I-V</normalizedToken>
yellow-orange,
<normalizedToken originalValue="VI–XI">VI-XI</normalizedToken>
gradually darkening to brown apex; forecoxa yellow, meso- and metacoxa yellow (male) or yellow and dark yellow-brown (female), femur yellow with dark brown band near apex (male) or base and apex (female), tibia brown to dark brown, with or without medial face lighter, tarsus brown; forebody longer; median frontal impression indistinct, in most specimens, obliterated by punctures; base of head without distinct posterior protuberances; dorsal surface of head densely, coarsely and rugosely punctate, punctures not becoming strigulose; pronotum slightly less transverse; entire surface with many micropunctures, lateral portion with deep, rugose and asetose punctures; pronotal protuberance indistinct in lateral view; elytra slightly less transverse; elytral suture more or less as long as pronotum at middle; elytra with punctation becoming strigulose only lateromedially; tergites
<normalizedToken originalValue="III–V">III-V</normalizedToken>
with impunctate areas medioapically, approximately equal in size; abdominal sternites with dense microsculpture, interspaces about as wide as lines; median lobe in lateral view broad, narrowed at midlength, widened in apical third and abruptly narrowed to minutely produced, acute apex (Fig. 18D); in parameral view, median lobe subparallel, near apex, narrowed to obtuse, blunt apex (Fig. 18A); paramere not divided apically but with median suture, shape as in Fig. 18A; peg setae densely distributed on entire apical portion of paramere (Fig. 18E); internal sac with well-sclerotized, horseshoe-shaped sclerite with a broadly expanded base (Fig. 18
<normalizedToken originalValue="B–C">B-C</normalizedToken>
); male sternite VIII with transverse basal line broken at middle, with broadly, shallowly emarginate apex, impressed and glabrous in small triangular area near emargination; male sternite IX sparsely setose, moderately asymmetrical at base and emarginate at apex (Fig. 23L); female tergite X elongate, with broadly rounded to nearly truncate apex (Fig. 26I); basal margin of laterotergal sclerites fused and thickened across base of tergite X, female laterotergal sclerites greatly expanded and overlapping with tergite X (Fig. 26I).
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="distribution">
<paragraph pageId="63" pageNumber="64">Distribution.</paragraph>
<paragraph pageId="63" pageNumber="64">
Figure 31C.
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus viridescens" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="viridescens">Bolitogyrus viridescens</taxonomicName>
is known only from
<normalizedToken originalValue="Teziutlán">Teziutlan</normalizedToken>
in Puebla, Mexico.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="bionomics">
<paragraph pageId="63" pageNumber="64">Bionomics.</paragraph>
<paragraph pageId="63" pageNumber="64">The type series was collected in early August.</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="etymology">
<paragraph pageId="63" pageNumber="64">Etymology.</paragraph>
<paragraph pageId="63" pageNumber="64">The species epithet refers to the brilliant green metallic reflection of the forebody.</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="comments">
<paragraph pageId="63" pageNumber="64">Comments.</paragraph>
<paragraph pageId="63" pageNumber="64">
Among potentially sympatric species,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus viridescens" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="viridescens">Bolitogyrus viridescens</taxonomicName>
is similar to
<taxonomicName class="Insecta" family="Staphylinidae" genus="Bolitogyrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bolitogyrus strigifrons" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="strigifrons">Bolitogyrus strigifrons</taxonomicName>
but is easily distinguished by the non-strigulose sculpture on the head.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="B696A30CD36FE2458638B37FF24DFF99" class="Insecta" family="Eulophidae" genus="Syntomosphyrum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syntomosphyrum indicum" order="Hymenoptera" pageId="0" pageNumber="8013" phylum="Arthropoda" rank="species" species="indicum">Syntomosphyrum indicum</taxonomicName>
Silvestri, 1910
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<taxonomicName id="86D3E6090814EA68BB08B9525762BBC2" genus="Chalcidoidea" lsidName="Chalcidoidea australia" pageId="0" pageNumber="8013" rank="species" species="australia">australia</taxonomicName>
Girault, 1917
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<paragraph id="C7E0D906ECB0054B25E7A25E6CC34682" pageId="0" pageNumber="8013">England</paragraph>
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<paragraph id="2A292A4226F66BAC00DC28744AE2AF9C" pageId="0" pageNumber="8013">Notes</paragraph>
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<bibRefCitation id="F1446941B606A24E32F3AF778ABD1D46" author="Graham, M. W. R. de V." journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="8013" pagination="1 - 322" title="A reclassification of the European Tetrastichinae (Hymenoptera: Eulophidae): revision of the remaining genera." volume="49" year="1991">Graham (1991a)</bibRefCitation>
records an English specimen that he supposed represented an accidental introduction.
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<taxonomicName LSID="F8FB28B5-B1DD-683B-FF28-EF0452ECA412" authority="Bourguignat, 1884" authorityName="Bourguignat" authorityYear="1884" class="Gastropoda" family="Melanopsidae" genus="Couphaniana" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Couphaniana" order="Sorbeoconcha" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Couphaniana Bourguignat, 1884</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="original source">
<paragraph pageId="5" pageNumber="6">Original source.</paragraph>
<paragraph pageId="5" pageNumber="6">
<bibRefCitation author="Bourguignat, JR" journalOrPublisher="Annales de Malacologie" pageId="303" pageNumber="304" pagination="1 - 168" publicationUrl="http://www.biodiversitylibrary.org/item/55067page/9/mode/1up" refId="B48" refString="Bourguignat, JR, 1884. Histoire des melaniens du systeme europeen. Annales de Malacologie 2: 1 - 168, http://www.biodiversitylibrary.org/item/55067page/9/mode/1up" title="Histoire des melaniens du systeme europeen." url="http://www.biodiversitylibrary.org/item/55067page/9/mode/1up" volume="2" year="1884">Bourguignat 1884</bibRefCitation>
: 74.
</paragraph>
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<paragraph pageId="5" pageNumber="6">Original classification.</paragraph>
<paragraph pageId="5" pageNumber="6">
Subgenus of
<taxonomicName authorityName="J. B. L. d'A. de Ferussac in J. B. L. d'A. de Ferussac &amp; A. E. J. P. J. F. d'A. de Ferussac" authorityYear="1807" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" lsidName="Melanopsis" order="Pteropoda" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="6">Melanopsis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<paragraph pageId="5" pageNumber="6">Type species.</paragraph>
<paragraph pageId="5" pageNumber="6">
<taxonomicName authorityName="Bourguignat" authorityYear="1884" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" lsidName="Melanopsis coupha" order="Pteropoda" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="Coupha">
<emphasis italics="true" pageId="5" pageNumber="6">Melanopsis coupha</emphasis>
</taxonomicName>
Bourguignat, 1884, by monotypy.
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</document>

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<mods:namePart>Du, Jianguo</mods:namePart>
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<mods:namePart>Ooi, Jillian Lean Sim</mods:namePart>
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<mods:namePart>Ma, Zhiyuan</mods:namePart>
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<mods:namePart>Rizman-Idid, Mohammed</mods:namePart>
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<taxonomicName LSID="F8FB66E0-AB52-568E-9C09-37ABA5748A1D" authority="Bloch, 1787" authorityName="Bloch" authorityYear="1787" class="Actinopterygii" family="Siganidae" genus="Siganus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siganus guttatus" order="Perciformes" pageId="0" pageNumber="47537" phylum="Chordata" rank="species" species="guttatus">Siganus guttatus (Bloch, 1787)</taxonomicName>
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<paragraph pageId="0" pageNumber="47537">Materials</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="47537">Type status:</emphasis>
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.
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country:
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; locality:
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;
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identifiedBy:
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<paragraph pageId="0" pageNumber="47537">Notes</paragraph>
<paragraph pageId="0" pageNumber="47537">
<bibRefCitation author="Harborne, A." journalOrPublisher="Coral Cay Conservation Ltd, London" pageId="0" pageNumber="47537" refId="B5376848" refString="Harborne, A., Fenner, D., Barnes, A., Beger, M., Harding, S., Roxburgh, T., 2000. Status report on the coral reefs of the east coast of Peninsula Malaysia. Coral Cay Conservation Ltd, London" title="Status report on the coral reefs of the east coast of Peninsula Malaysia" year="2000">Harborne et al. 2000</bibRefCitation>
;
<bibRefCitation author="Yusuf, Y. B." journalOrPublisher="Department of Fisheries, Kuala Lumpur and KUSTEM, Kuala Terengganu, Malaysia" pageId="0" pageNumber="47537" refId="B5376898" refString="Yusuf, Y. B., Mohd-Norizam, M., Ali, A. B., Zaidnuddin, I., 2001. Coral reef fish of some selected sites at Pulau Redang Marine Park, Terenggganu: A brief study. Department of Fisheries, Kuala Lumpur and KUSTEM, Kuala Terengganu, Malaysia" title="Coral reef fish of some selected sites at Pulau Redang Marine Park, Terenggganu: A brief study" year="2001">Yusuf et al. 2001</bibRefCitation>
; This study.
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<taxonomicName LSID="http://zoobank.org/96870F47-6129-474B-AB13-677EDD9D12C4" authority="Pang &amp; Tian" class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong Pang &amp; Tian</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="2">sp. n.</taxonomicNameLabel>
Figs 1-19
</paragraph>
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<paragraph pageId="1" pageNumber="2">Description.</paragraph>
<paragraph pageId="1" pageNumber="2">Length: 11.5-12.5 mm; width: 3.6-3.7 mm. Habitus as in Figure 1. Body extremely slender and elongate, with very long antennae, legs and mouthpart palps.</paragraph>
<caption pageId="1" pageNumber="2">
<paragraph pageId="1" pageNumber="2">
Figure 1.
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n., habitus of male, paratype.
</paragraph>
</caption>
<paragraph pageId="1" pageNumber="2">Black, but ventral surface, femora (except basal and apical tips), clypeus, labrum and apical half of mandibles dark brown, legs including basal and apical tips of femora, antennae, palps, and basal half of mandibles yellow to yellow brown; elytra with indistinct purplish metallic sheen.</paragraph>
<paragraph pageId="1" pageNumber="2">Macrosculpture: Strongly shining, surface glabrous, polish and smooth, but base of pronotum, mesosternum, meso- and metepisterna coarsely and sparsely punctate.</paragraph>
<paragraph pageId="1" pageNumber="2">Microsculpture: Engraved meshes moderately transverse on labrum, clypeus and base of frons, and base of pronotum; strongly transverse on head; striate on disc of pronotum and elytra, but clearly isodiametric on scutellum.</paragraph>
<paragraph lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
Head very long and narrow, strongly elongated, much longer than wide, HL/HW = 2.04-2.15 (mean 2.11); widest at level of eyes, and gradually narrowed backwards to neck constriction which is short but distinct, tempora almost straight but slightly curved just before neck constriction; eyes rather flat, more or less depressed; ventral margins of eye well separated from buccal fissure; supraorbital areas with two pairs of setiferous pores, anterior closer to margin of eye than posterior; posterior pore at about middle of head from clypeal suture to neck; interspaces between anterior pores distinctly wider than that between posterior ones; distance between anterior and posterior pores slightly longer than diameter of eye; distance between eye to buccal fissure distinctly shorter than that between posterior pore to eye, but slightly longer than that between anterior pore to eye; frontal impressions rather shallow and wide, ending before anterior supraorbital pores; frons and vertex moderately convex; clypeus moderately transverse, bisetose, labrum subquadrate, almost straight at front, sexsetose; mandibles elongate,
<pageBreakToken pageId="2" pageNumber="3" start="start">gently</pageBreakToken>
and gradually narrowed towards apex, apical teeth slightly hooked; right mandible with a small but distinct anterior retinacular tooth at about median portion which is far from other teeth; both right and left mandibles with short and blunt terebral and posterior retinacular teeth near base; ligula broad, bisetose at apex; labial suture well developed; mentum bisetose, apical margin biconcave, with a long and simple median
<pageBreakToken pageId="3" pageNumber="4" start="start">tooth</pageBreakToken>
which is widened at apex, epilobes evenly rounded; submentum bearing two pairs of setae, inner ones much long the outer; palps long and slender, subcylindrical, glabrous and asetose except for labial palpomere 2 which is bisetose on inner margin; labial palpomere 2 distinctly longer than 3; maxillary palpomere 3 slightly longer than 4. Antennae long and slender, filiform, extending at about apical 1/5 of elytra in female, but 1/6 in male; antennomeres 1-3 glabrous, antennomere 1 with a long subapical seta, 2 with a short subapical seta, 3 with several apical setae; finely pubescent from antennomere 4, each of 4 to 11 with several apical setae; antennomere 2 the shortest, half as long as 1, antennomeres 3-5 longer than other, each about twice as long as 1; gradually shortened from antennomeres 6 to 11, antennomere 11 almost as long as 1.
</paragraph>
<paragraph pageId="4" pageNumber="5">
<pageBreakToken pageId="4" pageNumber="5" start="start">Pronotum</pageBreakToken>
narrow and strongly elongate, barrel-like, distinctly shorter than head, almost as wide as head including eyes; much longer than wide, PL/PW = 1.59-1.62 (mean 1.60) in male, 1.40-1.43 (mean 1.41) in female; front slightly narrower than base, PWB/PWA = 1.11-1.14 (mean 1.13) in male, 1.09-1.12 (mean 1.10) in female; widest at about middle, gently narrowed towards both fore and hind angles; front and base finely bordered, lateral margins unbordered, but with evenly and distinctly explanate-reflexed areas throughout, marginal setae absent; basal foveae wide and long; fore angle nearly rectangular, hind angle broad though somewhat rectangular; both front and base almost straight; median line deep and long; disc slightly convex, basal area rather flat, with lateral areas of basal parts evidently depressed; propleura faintly tumid, faintly visible from above, at least at the widest part; prosternal process unbordered at apex; scutellum moderately sized.
</paragraph>
<paragraph pageId="5" pageNumber="6">
<pageBreakToken pageId="5" pageNumber="6" start="start">Elytra</pageBreakToken>
very slender, elongate-ovate, much wider than head and pronotum; well bordered at base, base small, shoulders indistinct; widest at a little behind middle, EL/EW = 1.62-1.78 (mean 1.74), more contracted towards apices than towards base; disc moderately convex though rather flat in basal 1/4; striae very deep, continuous and smooth, weakly punctured; intervals strongly convex, stria 3 with three dorsal setiferous pores, basal one close to stria 3, both middle and subapical ones close to stria 2; other intervals without pore; subapical sinuation rather straight, apex broad; preapical and two apical pores present; marginal series of umbilicate pores not aggregated, which are composed of about nineteen pores, denser in subhumeral and subapical areas, sparser in middle portion; three pores (one at subhumerus, other two at subapical area) bearing much longer setae than others, which is distinctly longer than metatarsomere 4; scutellar pores present; scutellar striae deep and rather long. Hind wings reduced.
</paragraph>
<paragraph pageId="5" pageNumber="6">Legs very long and slender; fore leg short and stout (Figs 2-5); procoxa asetose, meso- and metacoxae bisetose, inner seta of metacoxa absent; trochanters unisetose; femora very slender, profemora unisetose ventrally, meso- and metafemora with three and two ventral setae respectively; tibiae and tarsomeres 1-3 longitudinal bisulcute dorsally; protarsomeres 1-3 slightly dilated in male, with two rows of short and sparse spongy setae ventrally (Fig. 3), while much narrower and without spongy setae in female (Fig. 5); protarsomeres 4 shortly but distinctly emarginate at apex, with lobes nearly symmetric, each with a row of three setae ventrally; meso- and metatarsomeres 4 without subapical setae (Figs 6-10); tarsomeres 5 glabrous ventrally; claws smooth.</paragraph>
<caption pageId="5" pageNumber="6">
<paragraph pageId="5" pageNumber="6">
Figures 2-10. Tarsi of
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n., 2 protarsi, dorsal view, male 3 protarsi, ventral view, male 4 protarsi, dorsal view, female 5 protarsi, ventral view, female 6 mesotarsomeres 3-5, dorsal view, male 7 mesotarsomeres 3-5, ventral view, male 8 mesotarsomeres 3-5, lateral view, male 9 metatasomeres 3-5, dorsal view, male 10 metatasomeres 3-5, ventral view, male.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="6">
Each of abdominal ventrite
<normalizedToken originalValue="IV–VI">IV-VI</normalizedToken>
with pair of paramedian setae in both sexes; ventrite VII with two pairs of marginal setae in female, but only one pair in male.
</paragraph>
<paragraph pageId="5" pageNumber="6">Male genitalia (Figs 11-13): The median lobe of aedeagus rather stout, basal bulb large, strongly arcuate in middle portion in lateral view, gently and gradually narrowed towards apex, blunt at tip; dorsal opening wide, nearly as half as whole length, reaching 1/3 from base; apical lamella rather long, nearly twice as long as wide, not parallel-sided, rounded at apex; internal sac with a long copulatory piece covered with scales, and a strongly sclerotized spine dorsally; left paramere styloid, not elongated, smaller and shorter than the right.</paragraph>
<caption pageId="5" pageNumber="6">
<paragraph pageId="5" pageNumber="6">
Figures 11-13. Male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n., 11 median lobe, lateral view 12 apex of median lobe, dorsal view 13 parameres.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="6">Female reproductive tract (Fig. 14): Ventrite X sparsely setose; gonosubcoxite bearing about a dozen fringe setae along apical area, gonocoxite strongly curved, sharp at apex, bearing three lateral and one dorsal ensiform setae; bursa copulatrix wide, with middle part evidently folded, basally narrower; spermathecal gland very large, elongate ovate; spermathecal gland duct thin and long, connected below base of spermathecal reservoir, which is shorter than spermathecal duct.</paragraph>
<caption pageId="5" pageNumber="6">
<paragraph pageId="5" pageNumber="6">
Figure 14. Female reproductive tract of
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n., bc bursa copulatrix; co common oviduct; gcx gonocoxite; gscx gonosubcoxite; sd spermathecal duct; sg spermathecal gland; sgd spermathecal gland duct; sp spermatheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="sexual dimorphisms">
<paragraph pageId="5" pageNumber="6">Sexual dimorphisms.</paragraph>
<paragraph pageId="5" pageNumber="6">Apart from protarsomeres 1-3 and ventrite VII, sexual differences are also shown on antennae and pronotum: a little longer or more elongate in male than in female.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="variability">
<paragraph pageId="5" pageNumber="6">Variability.</paragraph>
<paragraph pageId="5" pageNumber="6">In general, maxillary palpomere 3 distinctly longer than 4, but reverse in a male specimen which left maxilla with palpomere 3 shorter than 4.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="remarks">
<paragraph pageId="5" pageNumber="6">Remarks.</paragraph>
<paragraph pageId="5" pageNumber="6">
Within platynines, the chaetotaxy on pronotum and meso- and metatarsi are important characters (
<bibRefCitation author="Habu, A" journalOrPublisher="Keigaku-Sha, Ltd., Tokyo" pageId="8" pageNumber="9" title="Carabidae Platynini (Insecta: Coleoptera)." volumeTitle="Japanese Faunica" year="1978">Habu 1978</bibRefCitation>
;
<bibRefCitation author="Liebherr, JK" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="8" pageNumber="9" pagination="1 - 163" title="Phylogeny and revision of the Anchomenus clade: the genera Tetraleucus, Anchomenus, Sericoda, and Elliptoleus (Coleoptera: Carabidae: Platynini)." volume="202" year="1991">Liebherr 1991</bibRefCitation>
;
<bibRefCitation author="Schmidt, J" journalOrPublisher="Entomologische Blaetter" pageId="8" pageNumber="9" pagination="211 - 218" title="Archicolpodes n. gen. - eine neue Laufkaefergattung der Tribus Platynini aus China (Coleoptera: Carabidae)." volume="96" year="2001">Schmidt 2001</bibRefCitation>
;
<bibRefCitation author="Liebherr, JK" journalOrPublisher="Mitteilungen des Museums fuer Naturkunde Berlin, Deutsche entomologische Zeitschrift" pageId="8" pageNumber="9" pagination="151 - 206" title="Phylogeny and biogeography of the Laurasian genus Agonum Bonelli (Coleoptera: Carabidae: Platynini)." volume="51" year="2004">Liebherr and Schmidt 2004</bibRefCitation>
). We treat this peculiar species a member of the genus
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Morimotoidius</taxonomicName>
Habu, 1954 due to the fact that it shares the following combined generic characters with other species of this genus: pronotal setae absent, submentum quadrisetose, meso- and metatarsomeres 4 without subapical setae, meso- and metafemora bearing two or three setae, and tarsomeres 5 glabrous ventrally. Certainly, other remarkably modified characters such as the very elongate and rhadinoid body shape, the slender and markedly porrected mandibles, the very thin palps, reduced eyes, the markedly prolonged temples and the very small elytral base must be autapomorphies to adapt the subterranean habitat.
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n. is the first member of
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Morimotoidius</taxonomicName>
found in mainland China. Other species of the genus are living either in Japan (two species) or in Taiwan Province of China (one species) (
<bibRefCitation author="Habu, A" journalOrPublisher="Bulletin of the National Institute of Agricultural Sciences (Series C (Plant Pathology and Entomology ))" pageId="8" pageNumber="9" pagination="263 - 279" title="On Trephionus otuboi Habu (Coleoptera: Carabidae)." volume="4" year="1954">Habu 1954</bibRefCitation>
,
<bibRefCitation author="Habu, A" journalOrPublisher="Keigaku-Sha, Ltd., Tokyo" pageId="8" pageNumber="9" title="Carabidae Platynini (Insecta: Coleoptera)." volumeTitle="Japanese Faunica" year="1978">1978</bibRefCitation>
;
<bibRefCitation author="Terada, K" journalOrPublisher="Miscellaneous Reports of the Hiwa Museum for Natural History" pageId="8" pageNumber="9" pagination="163 - 216" title="A checklist of the Carabidae (Coleoptera) of Taiwan." volume="45" year="2005">Terada et al. 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="etymology">
<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="6">
This new species is named after its type locality. In Chinese,
<normalizedToken originalValue="“Zhushan”">&quot;Zhushan&quot;</normalizedToken>
means a mountain or hill which is covered with bamboo forest,
<normalizedToken originalValue="“Dong”">&quot;Dong&quot;</normalizedToken>
means cave.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="materials examined">
<paragraph pageId="5" pageNumber="6">Materials examined.</paragraph>
<paragraph pageId="5" pageNumber="6">
Holotype: male, Zhushan Dong II, Dongkou Cun, Mabu Xiang, Wanzhai Xian, western Jiangxi, China, 28°02'880&quot;N, 114°22'216&quot;E, 142 m, 2-X-2012, Mingyi Tian &amp; Jingli Cheng leg. in the insect collections of South China Agricultural University, Guangzhou, China (SCAU); paratypes: 6 males and 9 females, ibid. in SCAU, except one male and one female in
<normalizedToken originalValue="Muséum">Museum</normalizedToken>
National
<normalizedToken originalValue="d’Histoire">d'Histoire</normalizedToken>
Naturelle, Paris, France, and one male and one female in Coll. J. Schmidt (Admannshagen, Germany), respectively.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="distribution">
<paragraph pageId="6" pageNumber="7">
<pageBreakToken pageId="6" pageNumber="7" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="6" pageNumber="7">China (Jiangxi) (Fig. 15).</paragraph>
<caption pageId="6" pageNumber="7">
<paragraph pageId="6" pageNumber="7">
Figure 15. Distribution map of
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="habits">
<paragraph pageId="6" pageNumber="7">Habits.</paragraph>
<paragraph pageId="6" pageNumber="7">
There is unknown about the biology and ecology of
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n. The beetles move quickly on walls and ceilings, feeding on other small arthropods, probably including eggs of the crickets which are common in the cave. The extremely modified troglomorphic characters mentioned above reveal that this beetle has ability to adapt underground environment and probably a troglobite though it has reduced eyes and pigmented body.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="about the locality cave of m. zhushandong sp. n">
<paragraph pageId="6" pageNumber="7">
About the locality cave of
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n.
</paragraph>
<paragraph pageId="6" pageNumber="7">Zhushan Dong is a touristic cave in western Jiangxi Province, located at Dongkou Cun, Mabu Xiang, Wanzai Xian, not far from the border between Wanzhai Xian and Yichun Shi (=City) (Fig. 15). Actually, there are two limestone caves in the Zhushan Dong scenic areas. Zhushan Dong I is a well-developed touristic cave, as long as 3985 m, with an underground river throughout the main passage. Zhushan Dong II is about 50 meters far from Zhushan Dong I. It is a small cave, about 30 m in length, with a small streamlet moving out at about 5 m inside of the entrance (Fig. 16, indicated by arrowhead). It is still closed for visitors. The beetles were collected by hands on walls and ceilings of the cave (Figs 17-19).</paragraph>
<caption pageId="6" pageNumber="7">
<paragraph pageId="6" pageNumber="7">
Figures 16-19. Zhushan Dong II. 16 entrance 17-18 cave walls where the type specimens were collected 19 an adult of
<taxonomicName class="Insecta" family="Carabidae" genus="Morimotoidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morimotoidius zhushandong" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="zhushandong">Morimotoidius zhushandong</taxonomicName>
sp. n. running on cave ceiling.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<subSubSection id="9A9596ECD930729CA73A74404A091EEE" pageNumber="418" type="nomenclature">
<paragraph id="43D56BBDA976EC932280B57D032DD6D9" pageNumber="418">
<taxonomicName id="9AE1DFEF2763F3F99B82BDCFF565E396" ID-CoL="7R5ZS" authority="Handley 1987" authorityName="Handley" authorityYear="1987" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageNumber="418" phylum="Chordata" rank="species" species="gnomus" subGenus="Dermanura">
<emphasis id="82D980FB7317DC2D99315F6F202D9AC3" italics="true" pageNumber="418">Artibeus (Dermanura) gnomus</emphasis>
Handley 1987
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="CF2E00000238F6EB171F91C2685ACF10" pageNumber="418" type="reference_group">
<paragraph id="B68917069BDBCB6EC631C6B496014BFF" pageNumber="418">
<treatmentCitation id="33926616D2A8BFA30318442900CA0FB4" author="Handley" authority="Handley 1987" authorityName="Handley" authorityYear="1987" class="Mammalia" family="Phyllostomidae" genus="Artibeus" journal="Fieldiana Zool." kingdom="Animalia" order="Chiroptera" page="167" pageNumber="418" phylum="Chordata" rank="species" species="gnomus" subGenus="Dermanura" title="Artibeus (Dermanura) gnomus" volume="39" year="1987">
<taxonomicName id="FC77D76CE82A450F5EBFB26DAC2F368F" ID-CoL="7R5ZS" authority="Handley 1987" authorityName="Handley" authorityYear="1987" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" pageNumber="418" phylum="Chordata" rank="species" species="gnomus" subGenus="Dermanura">
<emphasis id="DCC569CA12DA8F7A9E6D4F36C02BEA3C" italics="true" pageNumber="418">Artibeus (Dermanura) gnomus</emphasis>
Handley 1987
</taxonomicName>
,
<bibCitation id="D75B6FECE423CA6F9CDDD86BDA0EC663" author="Handley" journal="Fieldiana Zool." pageNumber="418" pagination="167" title="Artibeus (Dermanura) gnomus" volume="39" year="1987">Fieldiana Zool., 39: 167</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4810C9C200E5F8879BB704D7895EA811" pageNumber="418" type="materials_examined">
<paragraph id="3BDB6B5928F7EE43E6A20262401726A7" pageNumber="418">
<emphasis id="B5C395783C9D1E2DFA942A3AC2FAF481" inLineHeading="true" pageNumber="418">Type Locality:</emphasis>
<materialsCitation id="6384A0B2690A89F42D9C31B7ED47C396" ID-GBIF-Occurrence="4867283353" country="Venezuela" pageNumber="418" stateProvince="Bolivar">
<collectingCountry id="32EBF904A00643C265C349A9A742F968" name="Venezuela">Venezuela</collectingCountry>
,
<collectingRegion id="00D2E9AD00FB349C3B199B3326DB711C" country="Venezuela" name="Bolivar">Bolívar</collectingRegion>
,
<quantity id="DEB51A00215132C13852C7BC97889BAD" metricMagnitude="4" metricUnit="m" metricValue="5.9" unit="km" value="59.0">59 km</quantity>
SE El Dorado, El Manaco.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="573C9E89D5125C90876B6ADF23AAF004" pageNumber="418" type="vernacular_names">
<paragraph id="96772F9E73A53F7B0DE9381C941AE821" pageNumber="418">
<emphasis id="472B20FAF3F3022905A9FDE596B4E2DA" inLineHeading="true" pageNumber="418">Vernacular Names:</emphasis>
<vernacularName id="0374BDFA216818432A12AC6941C5F69C" language="eng" pageNumber="418">Dwarf Fruit-eating Bat</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="6522477167D4C860AD887286899D0BF6" pageNumber="418" type="distribution">
<paragraph id="2A6FABEF9A64A22FE5F14A9412CE78B8" pageNumber="418">
<emphasis id="6D6870658E5BF84CE659D8CC33F94945" inLineHeading="true" pageNumber="418">Distribution:</emphasis>
<collectingCountry id="52B6188CD2585CE333596C66B9F97984" name="Ecuador">Ecuador</collectingCountry>
,
<collectingCountry id="136DA7D24D91DD195947F06320DA73EB" name="Peru">Peru</collectingCountry>
,
<collectingCountry id="8DC6D10A21F4E4209CABD5FA5E2F222E" name="Bolivia">Bolivia</collectingCountry>
, Amazonian
<collectingCountry id="CC14A6DF54072AF04CF5F8BD51DF43F2" name="Brazil">Brazil</collectingCountry>
,
<collectingCountry id="96FDFE73B52F54C394BEA53C139DEE1C" name="Venezuela">Venezuela</collectingCountry>
, Guianas.
</paragraph>
</subSubSection>
<subSubSection id="47E0B3769494A742EA9D825E824D5F4E" pageNumber="418" type="conservation">
<paragraph id="E605593B5F4C68716651BB75AA58963F" pageNumber="418">
<emphasis id="79D7B8526BBF4CCBBC5949B744CE0EFF" inLineHeading="true" pageNumber="418">Conservation:</emphasis>
<collectionCode id="4314C4416230716EFA41EF05E1388D5F">IUCN</collectionCode>
2003 – Not listed (
<emphasis id="BA2594629982C0A820DAA00FCD5E9DDA" italics="true" pageNumber="418">lapsus</emphasis>
);
<collectionCode id="513ED6708C3AB9E3F1F9A8D26C85744A">IUCN</collectionCode>
/
<collectionCode id="B0CA77AA6EC5F5B2136789FF657725A8" country="USA" httpUri="http://grbio.org/cool/9wqd-8bvt" name="Sacramento State University" type="">SSC</collectionCode>
Action Plan (2001) – Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="97D9CD8833C0A731EB8887CADFD316AE" pageNumber="418" type="discussion">
<paragraph id="9CABEF5352BDB10CAC6B024565F6DCC1" pageNumber="418">
<emphasis id="F680B536262D0F51A9A6ADCD6F04ACC2" inLineHeading="true" pageNumber="418">Discussion:</emphasis>
Subgenus
<taxonomicName id="57C0DC5C7AADC25465DE1BF2ECCF6D1D" authorityName="Gervais" authorityYear="1856" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="subGenus" subGenus="Dermanura">
<emphasis id="8B1AAD976B6608FF08DEFF62FFB110EB" italics="true" pageNumber="418">Dermanura</emphasis>
</taxonomicName>
. Distinct from
<taxonomicName id="F546C2D21C4BC4BD083BDBC59A5EFF00" authorityName="Gervais" authorityYear="1856" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="8FDB5FF40C63C666CB41527B47F85047" italics="true" pageNumber="418">cinereus</emphasis>
</taxonomicName>
and
<taxonomicName id="7C3A3382942F24C87899A8E645F85738" authorityName="Thomas" authorityYear="1893" class="Mammalia" family="Phyllostomidae" genus="Artibeus" kingdom="Animalia" order="Chiroptera" phylum="Chordata" rank="species" species="glaucus">
<emphasis id="09B4BDB4A728DB46EF5127085245F382" italics="true" pageNumber="418">glaucus</emphasis>
</taxonomicName>
; see Handley (1987),
<bibRefCitation id="710C2F41EFDBA995F380D25C4595959C" author="Brosset, A. &amp; P. Charles-Dominique" refId="ref43246" refString="Brosset, A., and P. Charles-Dominique. 1990. The bats from French Guiana: A taxonomic, faunistic and ecological approach. Mammalia, 54: 509 - 559." year="1990">Brosset and Charles-Dominique (1990)</bibRefCitation>
, and
<bibRefCitation id="D6B6D792CF6074AD2B152BF1A6EAEB3A" author="Simmons, N. B. &amp; R. S. Voss" refId="ref328488" refString="Simmons, N. B., and R. S. Voss. 1998. The mammals of Paracou, French Guiana: A Neotropical lowland rainforest fauna. Part 1. Bats. Bulletin of the American Museum of Natural History, 237: 1 - 219." year="1998">Simmons and Voss (1998)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1120.87483" ID-GBIF-Dataset="5088bcc0-7b1c-4d9e-a05a-1ebe8ff092bc" ID-Pensoft-Pub="1313-2970-1120-67" ID-Pensoft-UUID="09E75B924CC8515D9724700E2F846EDC" ID-ZooBank="4B7BD92948A945178DF27F6780BC1AD3" ModsDocID="1313-2970-1120-67" checkinTime="1662465964255" checkinUser="pensoft" docAuthor="Han, Wei, Qiu, Lu, Zhu, Jing, Wang, Zong-Qing &amp; Che, Yan-Li" docDate="2022" docId="F8FBBA7649D5535CA37CDFDED842D126" docLanguage="en" docName="ZooKeys 1120: 67-94" docOrigin="ZooKeys 1120" docPubDate="2022-09-05" docSource="http://dx.doi.org/10.3897/zookeys.1120.87483" docTitle="Eupolyphaga reducta Qiu 2022, sp. nov." docType="treatment" docUuid="8749380B-3F77-4033-90D3-69A43F905ADD" docUuidSource="ZooBank" docVersion="2" id="09E75B924CC8515D9724700E2F846EDC" lastPageNumber="67" masterDocId="09E75B924CC8515D9724700E2F846EDC" masterDocTitle="Exploring the diversity of Eupolyphaga Chopard, 1929 (Blattodea, Corydioidea): species delimitation based on morphology and molecular analysis" masterLastPageNumber="94" masterPageNumber="67" pageNumber="67" updateTime="1662466667126" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Exploring the diversity of Eupolyphaga Chopard, 1929 (Blattodea, Corydioidea): species delimitation based on morphology and molecular analysis</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Han, Wei</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7243-1657</mods:nameIdentifier>
<mods:affiliation>Institute of Entomology, College of Plant Protection, Southwest University, Beibei, Chongqing 400716, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Qiu, Lu</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0946-1634</mods:nameIdentifier>
<mods:affiliation>Institute of Entomology, College of Plant Protection, Southwest University, Beibei, Chongqing 400716, China &amp; Engineering Research Center for Forest and Grassland Disaster Prevention and Reduction, Mianyang Normal University, Mianyang 621000, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhu, Jing</mods:namePart>
<mods:affiliation>Institute of Entomology, College of Plant Protection, Southwest University, Beibei, Chongqing 400716, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Zong-Qing</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9413-1105</mods:nameIdentifier>
<mods:affiliation>Institute of Entomology, College of Plant Protection, Southwest University, Beibei, Chongqing 400716, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Che, Yan-Li</mods:namePart>
<mods:affiliation>Institute of Entomology, College of Plant Protection, Southwest University, Beibei, Chongqing 400716, China</mods:affiliation>
<mods:nameIdentifier type="email">shirleyche2000@126.com</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-05</mods:number>
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<mods:detail type="volume">
<mods:number>1120</mods:number>
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<mods:extent unit="page">
<mods:start>67</mods:start>
<mods:end>94</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1120.87483</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1120.87483</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1120-67</mods:identifier>
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<mods:identifier type="Pensoft-UUID">09E75B924CC8515D9724700E2F846EDC</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:8749380B-3F77-4033-90D3-69A43F905ADD" httpUri="http://treatment.plazi.org/id/F8FBBA7649D5535CA37CDFDED842D126" lastPageNumber="67" pageId="0" pageNumber="67">
<subSubSection pageId="0" pageNumber="67" type="nomenclature">
<paragraph pageId="0" pageNumber="67">
<taxonomicName LSID="https://zoobank.org/8749380B-3F77-4033-90D3-69A43F905ADD" authority="Qiu" authorityName="Qiu" authorityYear="2022" class="Insecta" family="Corydiidae" genus="Eupolyphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolyphaga reducta" order="Blattodea" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">Eupolyphaga reducta Qiu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="67">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="description">
<paragraph pageId="0" pageNumber="67">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="type material">
<paragraph pageId="0" pageNumber="67">Type material.</paragraph>
<paragraph pageId="0" pageNumber="67">
<emphasis bold="true" italics="true" pageId="0" pageNumber="67">Holotype</emphasis>
: China · male, Sichuan Prov., Maoxian County, Wadi Township, Shahuzhai Village; 1938 m;
<geoCoordinate degrees="31" direction="north" minutes="53.20" orientation="latitude" precision="9" value="31.886667">31°53.20'N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="28.62" orientation="longitude" precision="9" value="103.477">103°28.62'E</geoCoordinate>
; 3 October 2019; Hao Xu, Zhi-Teng Chen, Lu Qiu leg (the holotype was reared from nymph by LQ).; SWU-B-CC-010020.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="materials_examined">
<paragraph pageId="0" pageNumber="67">Other specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="67">
<specimenCount type="male">1 male</specimenCount>
(an incompletely eclosed individual, unsuitable to be designated as a type specimen); SWU-B-CC-010021 ·
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 female</specimenCount>
nymph, same collection data as holotype; SWU-B-CC-010022
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="diagnosis">
<paragraph pageId="0" pageNumber="67">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="67">
This species is easily distinguished from its congeners by the reduced and stout tegmina and hind wings in male. In comparison with other species also possessing reduced tegmina and hind wings in males (e.g.,
<taxonomicName lsidName="E. everestiana subsp. reni" pageId="0" pageNumber="67" rank="subSpecies" species="everestiana" subSpecies="reni">
<emphasis italics="true" pageId="0" pageNumber="67">E. everestiana reni</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="E. udenostyla" pageId="0" pageNumber="67" rank="species" species="udenostyla">
<emphasis italics="true" pageId="0" pageNumber="67">E. udenostyla</emphasis>
</taxonomicName>
Qiu, sp. nov.), this new species is obviously larger (24.7 mm without tegmina and wings in length), the tegmina with sparse macula only on distal half; while
<taxonomicName lsidName="E. everestiana subsp. reni" pageId="0" pageNumber="67" rank="subSpecies" species="everestiana" subSpecies="reni">
<emphasis italics="true" pageId="0" pageNumber="67">E. everestiana reni</emphasis>
</taxonomicName>
is small in size (18.0-18.5 mm without tegmina and hind wings in length) and with evenly distributed maculae on tegmina.
<taxonomicName lsidName="E. udenostyla" pageId="0" pageNumber="67" rank="species" species="udenostyla">
<emphasis italics="true" pageId="0" pageNumber="67">E. udenostyla</emphasis>
</taxonomicName>
Qiu, sp. nov. (19.7 mm without tegmina and wings in length) is an almost unicolored black species with no styli in males.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="description">
<paragraph pageId="0" pageNumber="67">Description.</paragraph>
<paragraph pageId="0" pageNumber="67">
<emphasis bold="true" pageId="0" pageNumber="67">Male holotype</emphasis>
: Body short and broad (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5A, B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="67">
<emphasis italics="true" pageId="0" pageNumber="67">Measurements</emphasis>
(mm)
</emphasis>
: Overall length: 29.5; body length: 24.7; body width (tegmina not included): 13.4; pronotum length
<normalizedToken originalValue="×">x</normalizedToken>
width: 12.5
<normalizedToken originalValue="×">x</normalizedToken>
6.9; tegmina length
<normalizedToken originalValue="×">x</normalizedToken>
width: 22.5
<normalizedToken originalValue="×">x</normalizedToken>
9.3.
</paragraph>
<caption doi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="67">
<emphasis bold="true" pageId="0" pageNumber="67">Figure 5.</emphasis>
<taxonomicName authorityName="Qiu" authorityYear="2022" class="Insecta" family="Corydiidae" genus="Eupolyphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupolyphaga reducta" order="Blattodea" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="reducta">
<emphasis italics="true" pageId="0" pageNumber="67">Eupolyphaga reducta</emphasis>
</taxonomicName>
Qiu, sp. nov., male holotype
<emphasis bold="true" pageId="0" pageNumber="67">A</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="67">B</emphasis>
habitus, ventral view
<emphasis bold="true" pageId="0" pageNumber="67">C</emphasis>
pronotum, male holotype, dorsal view
<emphasis bold="true" pageId="0" pageNumber="67">D</emphasis>
head, male holotype, ventral view
<emphasis bold="true" pageId="0" pageNumber="67">E</emphasis>
supra-anal plate, ventral view
<emphasis bold="true" pageId="0" pageNumber="67">F</emphasis>
subgenital plate, ventral view
<emphasis bold="true" pageId="0" pageNumber="67">G</emphasis>
genitalia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="67">H</emphasis>
right phallomere, right-ventral view
<emphasis bold="true" pageId="0" pageNumber="67">I-J</emphasis>
the living holotype after moulting
<emphasis bold="true" pageId="0" pageNumber="67">K</emphasis>
habitat of
<taxonomicName lsidName="E. reducta" pageId="0" pageNumber="67" rank="species" species="reducta">
<emphasis italics="true" pageId="0" pageNumber="67">E. reducta</emphasis>
</taxonomicName>
Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province
<emphasis bold="true" pageId="0" pageNumber="67">L</emphasis>
Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (
<emphasis bold="true" pageId="0" pageNumber="67">A, B</emphasis>
); 0.2 cm (
<emphasis bold="true" pageId="0" pageNumber="67">C, D</emphasis>
); 0.1 cm (
<emphasis bold="true" pageId="0" pageNumber="67">E-H</emphasis>
). Photographs
<emphasis bold="true" pageId="0" pageNumber="67">I-L</emphasis>
by LQ.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="67">
<emphasis bold="true" pageId="0" pageNumber="67">
<emphasis italics="true" pageId="0" pageNumber="67">Coloration</emphasis>
.
</emphasis>
Body yellowish brown. Pronotum brown, anterior margin yellow (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5C</figureCitation>
). Space between vertex to the ocelli black, face fulvous with blackish brown spots. Antennae yellowish brown, ante-clypeus yellow, hind-clypeus brown. Labial palpi and maxillary palpi yellowish brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5D</figureCitation>
). Tegmina and hind wings yellowish brown, maculae brown. Legs and abdomen brown.
</paragraph>
<paragraph pageId="0" pageNumber="67">
<emphasis bold="true" italics="true" pageId="0" pageNumber="67">Head</emphasis>
: Interocular space narrow, obviously narrower than the distance between antennal sockets, and slightly narrower than the distance between ocelli. Ocelli large, ocellar ridge short, straight, covered with yellowish brown setae. Clypeus small (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5D</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="67">Pronotum</emphasis>
: Widely transverse, the widest point near the middle, anterior yellowish margin wide, the boundary between the yellowish and the brownish portion fused. Surface with short pubescence (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="67">Tegmina and hind wings</emphasis>
: Reduced and robust, exceeding the end of abdomen 4.4 mm. The apices of hind wings exceed tegmina obviously when at resting position (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5I</figureCitation>
). Tegmina translucent, yellowish brown, irregularly with several maculae apically, shoulders of tegmina wide, margins rounded. Wings translucent, light yellowish brown, almost no maculae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="67">Legs</emphasis>
: Slender, front femur type C1, arolia large, tarsal claws simple and symmetrical.
<emphasis bold="true" italics="true" pageId="0" pageNumber="67">Abdomen</emphasis>
: Supra-anal plate transversely broad, convex in the middle, cerci short (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5E</figureCitation>
). Subgenital plate slightly asymmetrical, right portion larger than left, styli short (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5F</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="67">Genitalia</emphasis>
: Well sclerotized (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5G, H</figureCitation>
). L1 slender, anteriorly prominently protruded, with a long protuberance on the left side and two well-developed protrusions on the posterior margin, L2 more developed on the right side, L3 curved, with a more rounded curvature, processa distale (pda) and processa apicale (paa) well developed, long and sturdy. Right phallomere relatively larger, R2 divided into two chunks, outer chunk larger than the inner one.
</paragraph>
<paragraph pageId="0" pageNumber="67">
<emphasis bold="true" pageId="0" pageNumber="67">Female and ootheca</emphasis>
: Unknown.
</paragraph>
<paragraph pageId="0" pageNumber="67">
<emphasis bold="true" pageId="0" pageNumber="67">Nymph</emphasis>
: Similar to the male in color, yellowish brown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="natural history">
<paragraph pageId="0" pageNumber="67">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="67">
This new species is distributed in the Heishui River Basin (Maoxian County to Heishui County). Nymphs of this new species live beneath the withered grass at the edge of a soil slope (Hao Xu, Lu Qiu, pers. obs.; Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eupolyphaga reducta Qiu, sp. nov., male holotype A habitus, dorsal view B habitus, ventral view C pronotum, male holotype, dorsal view D head, male holotype, ventral view E supra-anal plate, ventral view F subgenital plate, ventral view G genitalia, dorsal view H right phallomere, right-ventral view I-J the living holotype after moulting K habitat of E. reducta Qiu, sp. nov., in Shahuzhai Village, Maoxian County, Sichuan Province L Dr. Zhi-Teng Chen and Dr. Hao Xu collecting this species. Scale bars: 1.0 cm (A, B); 0.2 cm (C, D); 0.1 cm (E-H). Photographs I-L by LQ." figureDoi="10.3897/zookeys.1120.87483.figure5" httpUri="https://binary.pensoft.net/fig/738835" pageId="0" pageNumber="67">5K, L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="67" type="etymology">
<paragraph pageId="0" pageNumber="67">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="67">
The species epithet is from the Latin
<emphasis italics="true" pageId="0" pageNumber="67">reductus</emphasis>
indicating its reduced tegmina and wings.
</paragraph>
</subSubSection>
</treatment>
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<paragraph id="D78C0763C8273D0FE40F687C3CDE69E2" pageId="0" pageNumber="113952">China to Temperate East Asia</paragraph>
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