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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1004.57526" ID-GBIF-Dataset="4e633bc1-16fc-4e83-b8a0-cb9289897616" ID-PMC="PMC7758311" ID-Pensoft-Pub="1313-2970-1004-27" ID-Pensoft-UUID="BEECC18115E95A798CA5250990A350E3" ID-PubMed="33384565" ID-ZooBank="320559CF19B5423CB7FB72555290241A" ModsDocID="1313-2970-1004-27" checkinTime="1608157838683" checkinUser="pensoft" docAuthor="Maddison, Wayne P., Beattie, Imara, Marathe, Kiran, Ng, Paul Y. C., Kanesharatnam, Nilani, Benjamin, Suresh P. &amp; Kunte, Krushnamegh" docDate="2020" docId="08C20831109B547A8FA6CD03F9DD2518" docLanguage="en" docName="ZooKeys 1004: 27-97" docOrigin="ZooKeys 1004" docPubDate="2020-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1004.57526" docTitle="Piranthus bakau Maddison, sp. nov." docType="treatment" docVersion="4" id="BEECC18115E95A798CA5250990A350E3" lastPageNumber="27" masterDocId="BEECC18115E95A798CA5250990A350E3" masterDocTitle="A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)" masterLastPageNumber="97" masterPageNumber="27" pageNumber="27" updateTime="1668146624181" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maddison, Wayne P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4953-4575</mods:nameIdentifier>
<mods:affiliation>Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
<mods:nameIdentifier type="email">wmaddisn@mail.ubc.ca</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beattie, Imara</mods:namePart>
<mods:affiliation>Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marathe, Kiran</mods:namePart>
<mods:affiliation>Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada &amp; National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Paul Y. C.</mods:namePart>
<mods:affiliation>205 River Valley Road, # 16 - 53, Singapore 238274, Republic of Singapore</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kanesharatnam, Nilani</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6070-8989</mods:nameIdentifier>
<mods:affiliation>National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka &amp; Department of Zoology, Faculty of Science, Eastern University, Vantharumoolai, Sri Lanka</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Benjamin, Suresh P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4666-0330</mods:nameIdentifier>
<mods:affiliation>National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kunte, Krushnamegh</mods:namePart>
<mods:affiliation>National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-12-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1004</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>97</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1004.57526</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1004.57526</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1004-27</mods:identifier>
<mods:identifier type="ZooBank">320559CF19B5423CB7FB72555290241A</mods:identifier>
<mods:identifier type="Pensoft-UUID">BEECC18115E95A798CA5250990A350E3</mods:identifier>
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<treatment ID-GBIF-Taxon="176088135" LSID="urn:lsid:zoobank.org:act:1273542D-3BE3-404B-870E-246EEAA12967" httpUri="http://treatment.plazi.org/id/08C20831109B547A8FA6CD03F9DD2518" lastPageNumber="27" pageId="0" pageNumber="27">
<subSubSection pageId="0" pageNumber="27" type="nomenclature">
<paragraph pageId="0" pageNumber="27">
<taxonomicName LSID="http://zoobank.org/1273542D-3BE3-404B-870E-246EEAA12967" authority="Maddison, sp. nov." authorityName="Maddison, sp. nov." class="Arachnida" family="Salticidae" genus="Piranthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piranthus bakau" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="bakau">Piranthus bakau Maddison, sp. nov.</taxonomicName>
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">Figs 237-254</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="type material">
<paragraph pageId="0" pageNumber="27">Type material.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Holotype</emphasis>
male (specimen SWK12-0561, also known as d424), in UBCZ, from Malaysia: Sarawak: Bako Nat. Pk., Mangroves, beach forest,
<geoCoordinate degrees="1.722" direction="north" orientation="latitude" precision="55" value="1.722">1.722°N</geoCoordinate>
,
<geoCoordinate degrees="110.446" direction="east" orientation="longitude" precision="55" value="110.446">110.446°E</geoCoordinate>
, 0 m el. 8 March 2012 Maddison/Piascik/Ang/Lee WPM#12-003.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="etymology">
<paragraph pageId="0" pageNumber="27">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="27">
Referring to the type locality and to the
<normalizedToken originalValue="holotype’s">holotype's</normalizedToken>
habitat, mangroves (Malay,
<taxonomicName class="Arachnida" family="Salticidae" genus="Piranthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piranthus bakau" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="bakau">
<emphasis italics="true" pageId="0" pageNumber="27">Piranthus bakau</emphasis>
</taxonomicName>
= mangrove).
<emphasis italics="true" pageId="0" pageNumber="27">Other names</emphasis>
: In
<bibRefCitation author="Maddison, WP" journalOrPublisher="Journal of Arachnology" pageId="0" pageNumber="27" publicationUrl="http://salticidae.org/salticidImages/" refId="B23" refString="Maddison, WP, 2015b. Salticid images. http://salticidae.org/salticidImages/" title="Salticid images." url="http://salticidae.org/salticidImages/" year="2015 b">Maddison (2015b)</bibRefCitation>
and
<normalizedToken originalValue="WPM’s">WPM's</normalizedToken>
lab notebooks the informal code for this species was
<normalizedToken originalValue="“BKOMG”">&quot;BKOMG&quot;</normalizedToken>
. This species is shown in
<bibRefCitation author="Koh, JKH" journalOrPublisher="Sabah Forestry Department, city" pageId="0" pageNumber="27" refId="B18" refString="Koh, JKH, Bay, N, 2019. Borneo Spiders: A photographic field guide. Sabah Forestry Department, city" title="Borneo Spiders: A photographic field guide." year="2019">Koh and Bay 2019</bibRefCitation>
as the male of &quot;
<taxonomicName authorityName="Simon" authorityYear="1877" class="Arachnida" family="Pittosporaceae" genus="Bavia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bavia" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Bavia</emphasis>
</taxonomicName>
&quot; sp. D Strong-armed flat jumping spider.
</paragraph>
<caption doi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27" start="Figures 237–254" startId="F26">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figures 237-254.</emphasis>
<taxonomicName class="Arachnida" family="Salticidae" genus="Piranthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piranthus bakau" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="bakau">
<emphasis italics="true" pageId="0" pageNumber="27">Piranthus bakau</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="27">237</emphasis>
male left palp, prolateral view (holotype SWK12-0561)
<emphasis bold="true" pageId="0" pageNumber="27">238</emphasis>
same, ventral view
<emphasis bold="true" pageId="0" pageNumber="27">239</emphasis>
same, retrolateral view
<emphasis bold="true" pageId="0" pageNumber="27">240</emphasis>
epigyne, ventral (specimen AS19.2895)
<emphasis bold="true" pageId="0" pageNumber="27">241</emphasis>
vulva, dorsal
<emphasis bold="true" pageId="0" pageNumber="27">242</emphasis>
male JK.11.04.17.0040 (photograph Joseph K. H. Koh)
<emphasis bold="true" pageId="0" pageNumber="27">243-245</emphasis>
male holotype (SWK12-0561)
<emphasis bold="true" pageId="0" pageNumber="27">246</emphasis>
male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left)
<emphasis bold="true" pageId="0" pageNumber="27">247-254</emphasis>
female specimen AS19.2895
<emphasis bold="true" pageId="0" pageNumber="27">247</emphasis>
left fourth patella and tibia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="27">248</emphasis>
left first leg, prolateral view
<emphasis bold="true" pageId="0" pageNumber="27">249</emphasis>
adult female prosoma
<emphasis bold="true" pageId="0" pageNumber="27">250</emphasis>
adult female abdomen
<emphasis bold="true" pageId="0" pageNumber="27">251-254</emphasis>
same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="diagnosis">
<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
This and the closely similar
<taxonomicName lsidName="P. kohi" pageId="0" pageNumber="27" rank="species" species="kohi">
<emphasis italics="true" pageId="0" pageNumber="27">P. kohi</emphasis>
</taxonomicName>
differ from other known
<taxonomicName authorityName="Thorell" authorityYear="1895" class="Arachnida" family="Salticidae" genus="Piranthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piranthus" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Piranthus</emphasis>
</taxonomicName>
in having white transverse banding on the body and legs, and the posterior legs striped with black and translucent white.
<taxonomicName lsidName="P. bakau" pageId="0" pageNumber="27" rank="species" species="bakau">
<emphasis italics="true" pageId="0" pageNumber="27">P. bakau</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">237-254</figureCitation>
) differs from
<taxonomicName lsidName="P. kohi" pageId="0" pageNumber="27" rank="species" species="kohi">
<emphasis italics="true" pageId="0" pageNumber="27">P. kohi</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figures 255–269" captionStartId="F27" captionText="Figures 255 - 269. Piranthus kohi sp. nov. 255 male left palp, prolateral view (holotype AS 19.1813) 256 same, ventral view 257 same, retrolateral view 258 epigyne, ventral (specimen JK. 12.04.11.0032) 259 Vulva, dorsal 260 - 265 holotype 266 male holotype left fourth patella and tibia, dorsal view 267 female left fourth patella and tibia, dorsal view (specimen JK. 19.07.19.0001) 268 female JK. 19.07.19.0001 269 Female JK. 12.04.11.0032 (photograph Joseph K. H. Koh). Scale bars: on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures255-269" httpUri="https://binary.pensoft.net/fig/487558" pageId="0" pageNumber="27">255-269</figureCitation>
) in having:
</paragraph>
<paragraph pageId="0" pageNumber="27">Second white transverse band on the dorsum of the abdomen (i.e., the first behind the basal band) complete or broken by only a slight space;</paragraph>
<paragraph pageId="0" pageNumber="27">Sides of thorax lacking the three distinct narrow vertical lines seen in P. kohi (at most only a hint of two);</paragraph>
<paragraph pageId="0" pageNumber="27">Carapace lateral to the PLE with a bare patch, lacking golden scales, extending from PME back to behind PLE, and lateral to the bare patch is a stripe of denser golden scales (Figs 245, 249);</paragraph>
<paragraph pageId="0" pageNumber="27">First and second tibia bicoloured (black with white tip, arrow in Fig. 248);</paragraph>
<paragraph pageId="0" pageNumber="27">Second femur bicoloured (white basally, black terminally, Fig. 249);</paragraph>
<paragraph pageId="0" pageNumber="27">Black dorsal band on the fourth tibia incomplete, beginning mid-segment and reaching to the end (Figs 246, 247).</paragraph>
<paragraph pageId="0" pageNumber="27">Carapace slightly flatter than in P. kohi.</paragraph>
<paragraph pageId="0" pageNumber="27">Embolus (Fig. 237) notably longer than in P. kohi (Fig. 255), closely resembling that of P. planolancis (Nafin et al. 2020);</paragraph>
<paragraph pageId="0" pageNumber="27">Epigyne with cavernous atria framed by a curved medial ridge, and relatively long copulatory ducts leading to a posterior tangle of tubes and spermathecae.</paragraph>
<paragraph pageId="0" pageNumber="27">Juveniles have markings consistent with those of adults, and thus can be distinguished by the non-genitalic features above.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="description">
<paragraph pageId="0" pageNumber="27">Description.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Male</emphasis>
(based on holotype, SWK12-0561). Carapace length 2.7; abdomen length 3.0.
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Carapace</emphasis>
with rugose surface, black on ocular area, dark brown otherwise, covered thinly with narrow golden to white scales except bare patch lateral to PLE, and on posterior slope.
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Clypeus</emphasis>
narrow, dark, with a few white setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Chelicerae</emphasis>
small and vertical, dark, with a few pale setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Palp</emphasis>
(Figs
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">237-239</figureCitation>
) black except band of white scales terminally on femur, and cymbium, which is dark basally but then fades to white distally. Embolus long, arising on prolateral basal corner of bulb, looping first dorsally, then proximally, then distally. RTA a long blade. Endite margin rounded laterally.
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Legs</emphasis>
relatively larger (compared to the body) than in many baviines, with first pair especially robust. First and second legs mostly dark and with terminal white annuli on the segments. First patella and tibia with ventral fringe of black setae. Third and fourth leg segments from patella to tarsus translucent white with black bands and stripes. Fourth tibia and patella with a pattern of short black and white bands. On the tibia, a black dorsal band begins not basally but half way to the end, and extends distally to the tip; basally, it is interrupted by white (Fig.
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">246</figureCitation>
; compare with Fig.
<figureCitation captionStart="Figures 255–269" captionStartId="F27" captionText="Figures 255 - 269. Piranthus kohi sp. nov. 255 male left palp, prolateral view (holotype AS 19.1813) 256 same, ventral view 257 same, retrolateral view 258 epigyne, ventral (specimen JK. 12.04.11.0032) 259 Vulva, dorsal 260 - 265 holotype 266 male holotype left fourth patella and tibia, dorsal view 267 female left fourth patella and tibia, dorsal view (specimen JK. 19.07.19.0001) 268 female JK. 19.07.19.0001 269 Female JK. 12.04.11.0032 (photograph Joseph K. H. Koh). Scale bars: on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures255-269" httpUri="https://binary.pensoft.net/fig/487558" pageId="0" pageNumber="27">266</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Abdomen</emphasis>
with three narrow transverse white bands, a basal one, a second one behind that, and a third one behind that. Posterior to the third band the abdomen is a shiny black, but anterior it is dusted with golden setae.
</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Female</emphasis>
(based on specimen AS19.2895). Carapace length 3.0; abdomen length 4.1. The specimen was collected and photographed as a small juvenile (Figs
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">251-254</figureCitation>
); as an adult its basic appearance is similar (Figs
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">249</figureCitation>
,
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">250</figureCitation>
). Structure and markings of carapace and legs as in male.
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Abdomen</emphasis>
with second transverse band oblique.
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Epigyne</emphasis>
(Fig.
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">240</figureCitation>
) with medial septum dividing deep broad atria. What serves as the ECP is not obvious; there may be two, folds posterior to the atrium on each side, midway between the epigastric furrow and the posteriormost part of the medial septum. If so, it may be intermediate between
<taxonomicName lsidName="P. decorus" pageId="0" pageNumber="27" rank="species" species="decorus">
<emphasis italics="true" pageId="0" pageNumber="27">P. decorus</emphasis>
</taxonomicName>
, with a medial ECP (
<bibRefCitation DOI="https://doi.org/10.2476/asjaa.66.25" author="Caleb, JTD" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="27" pagination="25 - 29" refId="B6" refString="Caleb, JTD, Sanap, RV, 2017. Rediscovery of Piranthus decorus Thorell 1895 (Araneae: Salticidae) after 122 years since the original description. Acta Arachnologica 66: 25 - 29, DOI: https://doi.org/10.2476/asjaa.66.25" title="Rediscovery of Piranthus decorus Thorell 1895 (Araneae: Salticidae) after 122 years since the original description." url="https://doi.org/10.2476/asjaa.66.25" volume="66" year="2017">Caleb and Sanap 2017</bibRefCitation>
), and
<taxonomicName lsidName="P. planolancis" pageId="0" pageNumber="27" rank="species" species="planolancis">
<emphasis italics="true" pageId="0" pageNumber="27">P. planolancis</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.2.10" author="Malamel, JJ" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="27" pagination="267 - 276" refId="B34" refString="Malamel, JJ, Nafin, NS, Sudhikumar, AV, Sebastian, PA, 2019. Two new species of the jumping spiders (Araneae: Salticidae) from the genera Epeus Peckham et Peckham, 1886 and Piranthus Thorell, 1895 from India. Arthropoda Selecta 28: 267 - 276, DOI: https://doi.org/10.15298/arthsel.28.2.10" title="Two new species of the jumping spiders (Araneae: Salticidae) from the genera Epeus Peckham et Peckham, 1886 and Piranthus Thorell, 1895 from India." url="https://doi.org/10.15298/arthsel.28.2.10" volume="28" year="2019">Malamel et al. 2019</bibRefCitation>
; Nafin et al. 2020), with a pair of distantly separated lateral ECPs (Nafin et al. 2020: fig. 17).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="male-female matching">
<paragraph pageId="0" pageNumber="27">Male-female matching.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName lsidName="P. bakau" pageId="0" pageNumber="27" rank="species" species="bakau">
<emphasis italics="true" pageId="0" pageNumber="27">P. bakau</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. kohi" pageId="0" pageNumber="27" rank="species" species="kohi">
<emphasis italics="true" pageId="0" pageNumber="27">P. kohi</emphasis>
</taxonomicName>
are similar in general appearance, have overlapping geographical ranges, and have been collected to date with only adult males or adult females at a locality, not both. This leads to a question of which male matches which female. Unless there are additional closely similar species in the same areas, the inferred matching is well supported by the differences in markings, carapace shape, and lengths of embolus/copulatory ducts. The male of
<taxonomicName lsidName="P. bakau" pageId="0" pageNumber="27" rank="species" species="bakau">
<emphasis italics="true" pageId="0" pageNumber="27">P. bakau</emphasis>
</taxonomicName>
and the female from Tengkorak inferred to match it share the diagnostic traits mentioned above. Doubt might arise because of one difference in their markings: the female has the second transverse abdominal band more oblique, with its two sides meeting at a central peak, while in the holotype the band is straight across. However, the second male, from Brunei, shows a peak (Fig.
<figureCitation captionStart="Figures 237–254" captionStartId="F26" captionText="Figures 237 - 254. Piranthus bakau sp. nov. 237 male left palp, prolateral view (holotype SWK 12 - 0561) 238 same, ventral view 239 same, retrolateral view 240 epigyne, ventral (specimen AS 19.2895) 241 vulva, dorsal 242 male JK. 11.04.17.0040 (photograph Joseph K. H. Koh) 243 - 245 male holotype (SWK 12 - 0561) 246 male holotype left fourth patella and tibia, dorsal view (right leg digitally flipped to appear as left) 247 - 254 female specimen AS 19.2895 247 left fourth patella and tibia, dorsal view 248 left first leg, prolateral view 249 adult female prosoma 250 adult female abdomen 251 - 254 same specimen while immature. Scale bars on genitalia 0.1 mm; on bodies and legs 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures237-254" httpUri="https://binary.pensoft.net/fig/487557" pageId="0" pageNumber="27">242</figureCitation>
). A juvenile co-collected with the male holotype has the band peaked, and looks very much like the female when immature. The female is not designated as a paratype, however, because of the possibility it is a different but very closely related species, given the geographical distance between it and the holotype and their different habitats.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="natural history">
<paragraph pageId="0" pageNumber="27">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="27">Holotype male collected from mangroves; female from Tengkorak collected by shaking vines and understory trees near waterfall.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="materials_examined">
<paragraph pageId="0" pageNumber="27">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="27">
Two juveniles with same data as holotype. Also, one male (specimen JK.11.04.17.0040), in LKCNHM, from Brunei: Tutong, Tasek Merimbun, Zone C2, Palau Luba, Sungai Melunchur,
<geoCoordinate degrees="4.5817" direction="north" orientation="latitude" precision="5" value="4.5817">4.5817°N</geoCoordinate>
,
<geoCoordinate degrees="114.6872" direction="east" orientation="longitude" precision="5" value="114.6872">114.6872°E</geoCoordinate>
, J. K. H. Koh 17 Apr. 2011. One female (specimen AS19.2895), in UBCZ, from Malaysia: Johor: Gunung Belemut Forest, Lata Tengkorak,
<geoCoordinate degrees="2.055" direction="north" orientation="latitude" precision="55" value="2.055">2.055°N</geoCoordinate>
,
<geoCoordinate degrees="103.543" direction="east" orientation="longitude" precision="55" value="103.543">103.543°E</geoCoordinate>
, 250 m elev. 16 June 2019 Maddison/Morehouse/et al. WPM#19-057.
</paragraph>
</subSubSection>
</treatment>
</document>

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*
<taxonomicName id="54831FC44925A7A58960E98EE03A1B9B" ID-CoL="9HXQK" authority="Deeming &amp; Al-Dhafer, 2012" authorityName="Deeming &amp; Al-Dhafer" authorityYear="2012" class="Insecta" family="Chloropidae" genus="Trachysiphonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachysiphonella recurva" order="Diptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="recurva">
Trachysiphonella recurva Deeming &amp;
<normalizedToken id="95A4E855A91EDC8C01F74B17184D81FF" originalValue="Al–Dhafer">Al-Dhafer</normalizedToken>
, 2012
</taxonomicName>
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<paragraph id="4E2EF510451E33BD6273D76FC9C2E044" pageId="7" pageNumber="138">Material examined.</paragraph>
<paragraph id="C231FEC3D07AEC06DB0930118AD8C2CD" pageId="7" pageNumber="138">
Turkey: 13km NE of
<normalizedToken id="C1D476DFA3F3B2A099D7D2DF7D1EC8AF" originalValue="Muğla">Mugla</normalizedToken>
, pasture/pine wood, 1200m,
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,
<geoCoordinate id="96071AEB2D840E7FC04F2EFBC0EF0BF6" direction="east" orientation="longitude" precision="925" value="28.5">28°30'E</geoCoordinate>
, 23.-27.vi.2015, 1M; Turkey: 8 km S of
<normalizedToken id="CD72D4EED47D08F5D197AC5C7BE0986D" originalValue="Çine">Cine</normalizedToken>
, river bank, 68 m, SW + YPWT,
<geoCoordinate id="19F4710EC7934CEE3C45ECD25016747A" direction="north" orientation="latitude" precision="15" value="37.542778">37°32'34&quot;N</geoCoordinate>
,
<geoCoordinate id="A0E4416559AF041E208AB52F01C573D9" direction="east" orientation="longitude" precision="15" value="28.062777">28°03'46&quot;E</geoCoordinate>
, 28.-30.vi.2015, 1M and 1F.
</paragraph>
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<paragraph id="B475C8457A017F432970BD38AD845F1D" pageId="7" pageNumber="138">Distribution.</paragraph>
<paragraph id="08654EB2E30EF234902F1FE7B69A3303" pageId="7" pageNumber="138">this species was described from Yemen and further recorded from Oman and Saudi Arabia.</paragraph>
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