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<document ID-CLB-Dataset="297092" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="29b06347-2a16-47e8-8deb-c8084f3522bd" ID-ISBN="0-8018-8221-4" checkinTime="1670905252754" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="085E334FE45294A2F947B6FDF57FF43D" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Sciuridae.754-818.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Eupetaurus cinereus Thomas 1888" docType="treatment" docVersion="4" id="91C4B717759630DB830679AF52629BE0" lastPageNumber="767" masterDocId="91C4B717759630DB830679AF52629BE0" masterDocTitle="Order Rodentia - Family Sciuridae" masterLastPageNumber="818" masterPageNumber="754" pageNumber="767" updateTime="1716778896931" updateUser="ExternalLinkService">
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<taxonomicName id="6DB8610F5450930CC591A74013D230D2" ID-CoL="3CLXL" authority="Thomas 1888" authorityName="Thomas" authorityYear="1888" class="Mammalia" family="Sciuridae" genus="Eupetaurus" kingdom="Animalia" order="Rodentia" pageNumber="767" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="BF2FF045B7A70B654E9137A04866988B" italics="true" pageNumber="767">Eupetaurus cinereus</emphasis>
Thomas 1888
</taxonomicName>
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<treatmentCitation id="FE4EC0C1D391C3F38379C3FB55A647B9" author="Thomas" authority="Thomas 1888" authorityName="Thomas" authorityYear="1888" class="Mammalia" family="Sciuridae" genus="Eupetaurus" journal="J. Asiat. Soc. Bengal" kingdom="Animalia" order="Rodentia" page="258" pageNumber="767" phylum="Chordata" rank="species" species="cinereus" title="Eupetaurus cinereus" volume="57" year="1888">
<taxonomicName id="C694892C5A6D877DDC59F4A61ABBABB4" ID-CoL="3CLXL" authority="Thomas 1888" authorityName="Thomas" authorityYear="1888" class="Mammalia" family="Sciuridae" genus="Eupetaurus" kingdom="Animalia" order="Rodentia" pageNumber="767" phylum="Chordata" rank="species" species="cinereus">
<emphasis id="0DB36A5AD55E649384CF3C5D974A4C50" italics="true" pageNumber="767">Eupetaurus cinereus</emphasis>
Thomas 1888
</taxonomicName>
,
<bibCitation id="0B4759290DC1A691C638BDF0014A1DB5" author="Thomas" journal="J. Asiat. Soc. Bengal" pageNumber="767" pagination="258" title="Eupetaurus cinereus" volume="57" year="1888">J. Asiat. Soc. Bengal, 57: 258</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="CC1EA18FED945A6BCA4CD8D07826B4C0" pageNumber="767">
<emphasis id="5D3F9C5D35096DD436D0AADB53A2CF75" inLineHeading="true" pageNumber="767">Type Locality:</emphasis>
<materialsCitation id="184923EFCA99051D08CDE376E7E193B2" ID-GBIF-Occurrence="4867319301" country="Pakistan" elevation="1829.0" pageNumber="767">
&quot;Gilgit [Valley],...about
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&quot; [
<collectingCountry id="EEDA4DE10B83AC00DEFF46AC740B3DC6" name="Pakistan">Pakistan</collectingCountry>
,
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<elevation id="24BE34D191A758508D621F20A6F26703">1829 m</elevation>
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].
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</paragraph>
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<subSubSection id="8FC4F0BAA14B9E0E67FA643FD50AEBDC" pageNumber="767" type="vernacular_names">
<paragraph id="F5BC48B8DDED09796A4554ECB19A4A5F" pageNumber="767">
<emphasis id="C928E788A9968A2E9D99DAAFF8C45B25" inLineHeading="true" pageNumber="767">Vernacular Names:</emphasis>
<vernacularName id="BDC0B67928B0D9E2EDAC970A3269A5CB" language="eng" pageNumber="767">Woolly Flying Squirrel</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="FA850F77682EE20435552DC25E689442" pageNumber="767" type="distribution">
<paragraph id="43A3F908E4CCFBB68705B89B633D6AA3" pageNumber="767">
<emphasis id="E472A4B82A87F76B3C725EFC7E3CBE5B" inLineHeading="true" pageNumber="767">Distribution:</emphasis>
High elevations from N
<collectingCountry id="61B0F169BAA9D1A8E74F66C946A7E8BE" name="Pakistan">Pakistan</collectingCountry>
and Kashmir to
<collectingRegion id="53FE0BA2FB6689907B9A914E598319C9" country="India" name="Sikkim">Sikkim</collectingRegion>
(
<collectingCountry id="01C8196A3AE31929D9FF954A4F200BF6" name="India">India</collectingCountry>
) to
<collectingRegion id="A8AE2CA38A2369F83FA25BE3CA40848E" country="China" name="Xizang">Tibet</collectingRegion>
and possibly
<collectingRegion id="B9A6B1359316113E64FEA4D6E4DC6600" country="China" name="Yunnan">Yunnan</collectingRegion>
(
<collectingCountry id="4C78DAEA9897AE153149779F509273C0" name="China">China</collectingCountry>
;
<bibRefCitation id="48E0242301FB2A4E617400A4A8F39F6A" author="Agrawal, V. C. &amp; S. Chakraborty" refId="ref2060" refString="Agrawal, V. C., and S. Chakraborty. 1970. Occurrence of the woolly flying squirrel, Eupetaurus cinereus Thomas (Mammalia: Rodentia: Sciuridae) in North Sikkim. Journal of the Bombay Natural History Society, 66: 615 - 616." year="1970">Agrawal and Chakraborty, 1970</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="36A27CA0EC9901D629863EBE79EAE40B" pageNumber="767" type="conservation">
<paragraph id="73F737FBE0BCB937E31F72AABD2A9CA8" pageNumber="767">
<emphasis id="1817C1E89A6A6B32E6424A7A1C5203C6" inLineHeading="true" pageNumber="767">Conservation:</emphasis>
<collectionCode id="D28745479AFB208DC584C19AED65FF37">IUCN</collectionCode>
Endangered.
</paragraph>
</subSubSection>
<subSubSection id="783E38CC9D6F80B090BAB768B35C621D" pageNumber="767" type="discussion">
<paragraph id="5EF3EFF7E0FB927E48548781F6B710A6" pageNumber="767">
<emphasis id="6A06287707087EADE2254968968309EA" inLineHeading="true" pageNumber="767">Discussion:</emphasis>
Reviewed by
<bibRefCitation id="D50D6E8FD15A1B7AE83956067F39CF32" author="McKenna, M. C." refId="ref233659" refString="McKenna, M. C. 1962. Eupetaurus and the living petauristine sciurids. American Museum Novitates, 2104: 1 - 38." year="1962">McKenna (1962)</bibRefCitation>
. In 1994, the first specimen to be found in &quot;...over a century...&quot; was obtained in the Gilgit district (
<bibRefCitation id="049939D224DE3D33B3DFEFD7C146EF63" author="Zahler, P." refId="ref404008" refString="Zahler, P. 1996. Rediscovery of the woolly flying squirrel (Eupetaurus cinereus). Journal of Mammalogy, 77 (1): 54 - 57." year="1996">Zahler, 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="F7E85D2369576D1FB767F48E36B61EEC" ID-DOI="10.5281/zenodo.5412309" ID-ISSN="0253-116X" ID-Zenodo-Dep="5412309" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630354631537" checkinUser="valdenar" docAuthor="Vavrová, Ľ. &amp; Šteffek, J." docDate="2007" docId="085E9A0B6001D55FFF0E3AEAFDD01BD3" docLanguage="en" docName="LinzerbiolBeitr.2007.39.1.667-676.pdf" docOrigin="Linzer biologische Beiträge 39 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Vertigo angustior JEFFREYS 1830" docType="treatment" docVersion="4" lastPageNumber="671" masterDocId="F467E2736005D55BFF993962FFCB1F21" masterDocTitle="Favourable conservation status of European important species of genus Vertigo spp. in Slovakia - categories and criteria" masterLastPageNumber="676" masterPageNumber="667" pageNumber="671" updateTime="1699243242624" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CF61EA7116CF229C2B35C43349F73340">Favourable conservation status of European important species of genus Vertigo spp. in Slovakia - categories and criteria</mods:title>
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<mods:namePart id="2633677AE0135E16B8EA4B3C478270F7">Šteffek, J.</mods:namePart>
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<taxonomicName id="47F7509E6001D55FFF0E3AEAFE3D1CBF" ID-CoL="7FRLB" authority="JEFFREYS 1830" authorityName="JEFFREYS" authorityYear="1830" box="[151,502,904,926]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="671" phylum="Mollusca" rank="species" species="angustior">
<emphasis id="B283F70F6001D55FFF0E3AEAFE841CBC" bold="true" box="[151,335,904,925]" italics="true" pageId="4" pageNumber="671">Vertigo angustior</emphasis>
JEFFREYS 1830
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(
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)
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</heading>
</paragraph>
<paragraph id="80482B1D6001D55FFF0E3AD9FE181BAC" blockId="4.[151,1059,955,1165]" pageId="4" pageNumber="671">
Currently there is a lack of information about the population size and abundance of this species in
<collectingCountry id="F8E06B8D6001D55FFE983AB9FE961CD0" box="[257,349,987,1009]" name="Slovakia" pageId="4" pageNumber="671">Slovakia</collectingCountry>
. Despite the fact, the criterion C1 was evaluated as &quot;good&quot; because of the relatively widespread distribution of the species in
<collectingCountry id="F8E06B8D6001D55FFD733A98FC8D1B31" box="[746,838,1018,1040]" name="Slovakia" pageId="4" pageNumber="671">Slovakia</collectingCountry>
. Even though a low weight (2 or 1) was appointed for the criterion C1. The criterion C2 (Habitats) was evaluated as &quot;good&quot; from the habitat status point of view and &quot;average&quot; concerning the water regime because of the long-term decreasing of numbers of suitable wetland habitats within
<collectingCountry id="F8E06B8D6001D55FFE913D15FEA81BAC" box="[264,355,1143,1165]" name="Slovakia" pageId="4" pageNumber="671">Slovakia</collectingCountry>
, generally.
</paragraph>
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<paragraph id="80482B1D6001D55FFF0E3DFDFDD01BD3" blockId="4.[151,1059,1182,1266]" pageId="4" pageNumber="671">
<taxonomicName id="47F7509E6001D55FFF0E3DFDFEDD1B95" authorityName="JEFFREYS" authorityYear="1830" box="[151,278,1183,1204]" class="Gastropoda" family="Vertiginidae" genus="Vertigo" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="671" phylum="Mollusca" rank="species" species="angustior">
<emphasis id="B283F70F6001D55FFF0E3DFDFEDD1B95" box="[151,278,1183,1204]" italics="true" pageId="4" pageNumber="671">V. angustior</emphasis>
</taxonomicName>
is listed as vulnerable (VU) in the National Red List of Molluscs of
<collectingCountry id="F8E06B8D6001D55FFC793DFCFF7D1BF2" name="Slovakia" pageId="4" pageNumber="671">Slovakia</collectingCountry>
(ŠTEFFEK &amp; VAVROVÁ 2006), which was prepared in 2006 under the categories and criterion of IUCN, version 3.1 (2001).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:titleInfo id="470B33E2A2D6FDEDF356E3E2E4DE2A80">
<mods:title id="90BB13959399CD1F76A670170847BD6E">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="E2B8E7395387EE8EC82BC4D195968ED3">
<mods:date id="A080AA37B9B1E5872D1D529265FCD0F8">2001</mods:date>
<mods:detail id="6BE556C06CE0E248BB887A304D8CAC95" type="volume">
<mods:number id="1E6F34BBD8737A3337E98F012FFF08F2">35</mods:number>
</mods:detail>
<mods:extent id="7E16453885AA7E69D705E9A9DE7F8ACB" unit="page">
<mods:start id="8FDF3B3327E3C47C2526866F1CB30499">583</mods:start>
<mods:end id="722D08A623ABD4EC8F1D643BF8787731">616</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="57C52525B9BD5B956CBB9B9AD520EDF6">journal article</mods:classification>
<mods:identifier id="93DA5996484CE65EF7B4EF6F2807F600" type="ISSN">1464-5262</mods:identifier>
</mods:mods>
<treatment id="085EF2314D647260625BA50D41A3FC37" LSID="urn:lsid:plazi:treatment:085EF2314D647260625BA50D41A3FC37" httpUri="http://treatment.plazi.org/id/085EF2314D647260625BA50D41A3FC37" lastPageId="29" lastPageNumber="610" pageId="24" pageNumber="605">
<subSubSection id="C8ED10AC4D647265625BA50D40E0FC45" pageId="24" pageNumber="605" type="nomenclature">
<paragraph id="804843274D647265625BA50D4173FC65" blockId="24.[341,797,893,950]" box="[341,797,893,917]" pageId="24" pageNumber="605">
<heading id="DB00F44B4D647265625BA50D4173FC65" box="[341,797,893,917]" centered="true" fontSize="10" level="2" pageId="24" pageNumber="605" reason="3">
<taxonomicName id="47F738A44D647265625BA50D40BAFC65" ID-CoL="8BZ7X" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[341,724,893,917]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="605" phylum="Arthropoda" rank="species" species="samaruensis" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D647265625BA50D40BAFC65" bold="true" box="[341,724,893,917]" pageId="24" pageNumber="605">
<emphasis id="B2839F354D647265625BA50D43F4FC65" bold="true" box="[341,410,893,917]" italics="true" pageId="24" pageNumber="605">Empis</emphasis>
(
<emphasis id="B2839F354D64726562A2A50D4053FC65" bold="true" box="[428,573,893,917]" italics="true" pageId="24" pageNumber="605">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D647265615EA50D40BAFC65" bold="true" box="[592,724,893,917]" italics="true" pageId="24" pageNumber="605">samaruensis</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D64726561D0A50E4173FC65" box="[734,797,894,917]" pageId="24" pageNumber="605" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="804843274D64726562EAA5ED40E0FC45" blockId="24.[341,797,893,950]" box="[484,654,925,950]" pageId="24" pageNumber="605">(®gures 39±41)</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D6472606352A58241A3FC37" lastPageId="29" lastPageNumber="610" pageId="24" pageNumber="605" type="materials_examined">
<paragraph id="804843274D6472656352A5824296FBF9" blockId="24.[92,1047,1009,1097]" box="[92,248,1009,1033]" pageId="24" pageNumber="605">
<emphasis id="B2839F354D6472656352A5824296FBF9" box="[92,248,1009,1033]" italics="true" pageId="24" pageNumber="605">Type material</emphasis>
</paragraph>
<paragraph id="804843274D647265638AA26140A5FBB9" blockId="24.[92,1047,1009,1097]" pageId="24" pageNumber="605">
<materialsCitation id="309F497A4D647265638AA261467FFBD9" box="[132,1041,1041,1065]" collectingDate="1972-08-02" collectionCode="NHM" collectorName="J. C. Deeming" country="Nigeria" location="Zaria" municipality="Samaru" pageId="24" pageNumber="605" specimenCount="1" typeStatus="holotype">
<typeStatus id="5F4CFD854D647265638AA2614299FBD9" box="[132,247,1041,1065]" pageId="24" pageNumber="605" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F8E003B74D6472656222A26143EEFBD9" box="[300,384,1041,1065]" name="Nigeria" pageId="24" pageNumber="605">Nigeria</collectingCountry>
,
<location id="852815FC4D647265629FA26143A1FBD9" LSID="urn:lsid:plazi:treatment:085EF2314D647260625BA50D41A3FC37:852815FC4D647265629FA26143A1FBD9" box="[401,463,1041,1065]" country="Nigeria" municipality="Samaru" name="Zaria" pageId="24" pageNumber="605">Zaria</location>
,
<collectingMunicipality id="602CD95D4D64726562EEA2614059FBD9" box="[480,567,1041,1065]" pageId="24" pageNumber="605">Samaru</collectingMunicipality>
,
<date id="F44965E74D6472656147A261409EFBD9" box="[585,752,1041,1065]" pageId="24" pageNumber="605" value="1972-08-02">
<collectingDate id="E40D9C0F4D6472656147A261409EFBD9" box="[585,752,1041,1065]" pageId="24" pageNumber="605" value="1972-08-02">2 August 1972</collectingDate>
</date>
,
<collectorName id="2D0226F14D647265600FA26141C6FBD9" box="[769,936,1041,1065]" pageId="24" pageNumber="605">J. C. Deeming</collectorName>
(
<collectionCode id="E6E6DBE24D64726560B3A261466CFBD8" box="[957,1026,1041,1064]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="24" pageNumber="605" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="309F497A4D6472656352A24140A8FBB9" box="[92,710,1073,1097]" collectingDate="1971-08-19" collectionCode="MNHN" collectorName="J. C. Deeming" country="Nigeria" location="Zaria" municipality="Samaru" pageId="24" pageNumber="605" specimenCount="1" typeStatus="paratype">
<typeStatus id="5F4CFD854D6472656352A24142A7FBB9" box="[92,201,1073,1097]" pageId="24" pageNumber="605" type="paratype">PARATYPE</typeStatus>
: 1, same data but`
<date id="F44965E74D64726562B1A2414051FBB9" box="[447,575,1073,1097]" pageId="24" pageNumber="605" value="1971-08-19">
<collectingDate id="E40D9C0F4D64726562B1A2414051FBB9" box="[447,575,1073,1097]" pageId="24" pageNumber="605" value="1971-08-19">19.viii.1971</collectingDate>
</date>
(
<collectionCode id="E6E6DBE24D6472656152A24140D6FBB8" box="[604,696,1073,1096]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="605">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="804843274D6472656352A20442B4FB7B" blockId="24.[92,1047,1140,1674]" box="[92,218,1140,1163]" pageId="24" pageNumber="605">
<emphasis id="B2839F354D6472656352A20442B4FB7B" box="[92,218,1140,1163]" italics="true" pageId="24" pageNumber="605">Description</emphasis>
</paragraph>
<paragraph id="804843274D647265638AA2E343BBFB5A" blockId="24.[92,1047,1140,1674]" box="[132,469,1171,1195]" pageId="24" pageNumber="605">Male: length of wing 3.9 mm.</paragraph>
<paragraph id="804843274D647265638AA2C340C4FABA" blockId="24.[92,1047,1140,1674]" pageId="24" pageNumber="605">Head with occiput blackish. Ocellar triangle prominent with pair of short bristles. Face blackish, shining in lower part. Scape, pedicel dark brown, ¯agellum blackish. Proboscis not very long (1.5 times the head height), labrum brown to dark brown, labium with distinct annulations, strongly sclerotized, labella blackish, palpus blackish. Eyes holoptic, upper ommatidia enlarged.</paragraph>
<paragraph id="804843274D647265638AA32240F8F999" blockId="24.[92,1047,1140,1674]" pageId="24" pageNumber="605">Thorax with antepronotum dark brown with a few ®ne, short lateral bristles. Proepisternum bare. Prosternum with one ®ne, short bristle. Postpronotal lobes brown with one distinct, not very strong basal bristle. Scutum black, somewhat shining to strongly dusted greyish in prescutellar depression. Acrostichals and dorsocentrals apparently uniserial, ®ne, very short, latter with one strong, long prescutellar bristle. One distinct, not very strong postsutural supraalar, one ®ner, shorter presutural. One strong, long notopleural. One strong, long postalar. Scutellum with two strong, long apical bristles, two ®ne, shorter subapicals. Laterotergite with fan of bristles. Anterior and posterior spiracles blackish.</paragraph>
<paragraph id="804843274D647264638AA0024037FB1A" blockId="24.[92,1047,1140,1674]" lastBlockId="25.[92,1047,1043,1673]" lastPageId="25" lastPageNumber="606" pageId="24" pageNumber="605">Legs brownish. Fore femur with one row of ®ne, not very long ventral bristles. Fore tibia with one row of strong, long bristles especially on apical half. First tarsomere of fore tarsus with distinct short bristles especially dorsally. Mid femur with one row of strong, long posteroventrals, a few distinct anteroventrals at basal tip. Mid tibia with rather strong, not very long ventrals. First tarsomere of mid tarsus with distinct dorsals and ventrals. Hind femur with ®ne, rather long ventrals. Hind tibia with numerous strong, rather long dorsals. First tarsomere of hind tarsus with numerous distinct dorsals and ventrals.</paragraph>
<caption id="D48813AF4D6572646352A50C434CFC11" pageId="25" pageNumber="606" startId="25.[92,105,892,913]" targetBox="[102,1038,119,863]" targetPageId="25">
<paragraph id="804843274D6572646352A50C434CFC11" blockId="25.[92,1048,891,993]" pageId="25" pageNumber="606">
F IGS 3941.
<taxonomicName id="47F738A44D65726463E4A50C4051FC61" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[234,575,891,913]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="606" phylum="Arthropoda" rank="species" species="samaruensis" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D65726463E4A50C4349FC61" box="[234,295,892,913]" italics="true" pageId="25" pageNumber="606">Empis</emphasis>
(
<emphasis id="B2839F354D657264623AA50B43D9FC61" box="[308,439,891,913]" italics="true" pageId="25" pageNumber="606">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D65726462C6A50C4051FC61" box="[456,575,892,913]" italics="true" pageId="25" pageNumber="606">samaruensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D6572646146A50C40ECFC61" box="[584,642,892,913]" pageId="25" pageNumber="606" rank="species">sp. n.</taxonomicNameLabel>
: (39) male sternites 6, 7 and 8, ventral view (scale 0.1 mm); (40) male segment 8, lateral view (scale 0.1 mm); (41) male hypopygium, lateral view (scale 0.1 mm). st 7 5 sternite 7; st 8 5 sternite 8; tg 8 5 tergite 8.
</paragraph>
</caption>
<paragraph id="804843274D657264638AA28340AFFABA" blockId="25.[92,1047,1043,1673]" pageId="25" pageNumber="606">
Wing clear. Sc abbreviated.
<collectionCode id="E6E6DBE24D65726462C4A28343B3FAFB" box="[458,477,1267,1291]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="606">R</collectionCode>
41 5 branched at right-angle. Discal cell strongly truncate. M1, M2 abbreviated. A1 complete. Anal lobe well developed, anal angle acute. Halter with yellow base and stem, black knob.
</paragraph>
<paragraph id="804843274D657264638AA32241A4FA3A" blockId="25.[92,1047,1043,1673]" pageId="25" pageNumber="606">Abdomen (®gures 39, 40) with ®rst three segments yellowish with distinct lateral bristles, otherwise brown. Sternite 6 unmodi®ed. Sternite 7 distinct but membranous (®gure 39). Sternite 8 somewhat elongated posteriorly, with a ventroanterior process on each side (®gures 39, 40). Width of tergite 8 reduced laterally (®gure 40).</paragraph>
<paragraph id="804843274D657264638AA3A24091F999" blockId="25.[92,1047,1043,1673]" pageId="25" pageNumber="606">Hypopygium (®gure 41) with cercus made up of ventroinner bare lobe, visible in dorsal view, and well-bristled external lobe. Epandrial lamellae connected anteriorly, with rather broad rounded anterior tip and narrower rounded wellbristled posterior tip. Hypandrium well sclerotized, with characteristic shape in lateral view. Phallus rather short, thick, not pointed at tip.</paragraph>
<paragraph id="804843274D657264638AA001433FF979" blockId="25.[92,1047,1043,1673]" box="[132,337,1649,1673]" pageId="25" pageNumber="606">Female unknown.</paragraph>
<paragraph id="804843274D6672676352A60742BEFF7E" blockId="26.[92,1047,119,206]" box="[92,208,119,142]" pageId="26" pageNumber="607">
<emphasis id="B2839F354D6672676352A60742BEFF7E" box="[92,208,119,142]" italics="true" pageId="26" pageNumber="607">Discussion</emphasis>
</paragraph>
<paragraph id="804843274D667267638AA6E741FBFF3E" blockId="26.[92,1047,119,206]" pageId="26" pageNumber="607">
<taxonomicName id="47F738A44D667267638AA6E74346FF5E" authorityName="Esa" authorityYear="2001" box="[132,296,151,174]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="607" phylum="Arthropoda" rank="species" species="samaruensis">
<emphasis id="B2839F354D667267638AA6E74346FF5E" box="[132,296,151,174]" italics="true" pageId="26" pageNumber="607">E. samaruensis</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="47F738A44D66726762DAA6E740F6FF5E" authorityName="Collin" authorityYear="1930" box="[468,664,150,174]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="607" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D66726762DAA6E74380FF5E" box="[468,494,151,174]" italics="true" pageId="26" pageNumber="607">E.</emphasis>
(
<emphasis id="B2839F354D66726762F4A6E6407BFF5E" box="[506,533,150,174]" italics="true" pageId="26" pageNumber="607">C.</emphasis>
)
<emphasis id="B2839F354D667267612AA6E640F6FF5E" box="[548,664,150,174]" italics="true" pageId="26" pageNumber="607">chrysocera</emphasis>
</taxonomicName>
-complex. It may be distinguished from all other species of this complex by its unmodi®ed male sternite 6.
</paragraph>
<paragraph id="804843274D6672676352A6864319FEDD" blockId="26.[92,375,246,302]" box="[92,375,246,302]" pageId="26" pageNumber="607">
<emphasis id="B2839F354D6672676352A686428EFEFE" box="[92,224,246,270]" italics="true" pageId="26" pageNumber="607">Distribution</emphasis>
West Africa:
<collectingCountry id="F8E003B74D6672676212A766431EFEDE" box="[284,368,278,302]" name="Nigeria" pageId="26" pageNumber="607">Nigeria</collectingCountry>
.
</paragraph>
<paragraph id="804843274D6672676352A7264176FE7D" blockId="26.[92,792,342,398]" box="[92,792,342,398]" pageId="26" pageNumber="607">
<emphasis id="B2839F354D6672676352A72642BDFE9E" box="[92,211,342,366]" italics="true" pageId="26" pageNumber="607">Etymology</emphasis>
The name of the species is derived from the type-locality.
</paragraph>
<paragraph id="804843274D6672676268A7C540FDFE1D" blockId="26.[358,780,437,494]" pageId="26" pageNumber="607">
<taxonomicName id="47F738A44D6672676268A7C540ADFE3D" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[358,707,437,461]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="607" phylum="Arthropoda" rank="species" species="singulare" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D6672676268A7C540ADFE3D" bold="true" box="[358,707,437,461]" pageId="26" pageNumber="607">
<emphasis id="B2839F354D6672676268A7C543C5FE3D" bold="true" box="[358,427,437,461]" italics="true" pageId="26" pageNumber="607">Empis</emphasis>
(
<emphasis id="B2839F354D66726762B3A7C54020FE3D" bold="true" box="[445,590,437,461]" italics="true" pageId="26" pageNumber="607">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D667267616FA7C540ADFE3D" bold="true" box="[609,707,437,461]" italics="true" pageId="26" pageNumber="607">singulare</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D66726761C3A7C64162FE3D" box="[717,780,438,461]" pageId="26" pageNumber="607" rank="species">sp. n.</taxonomicNameLabel>
(®gures 42±46)
</paragraph>
<caption id="D48813AF4D6672676352A07A434CF97A" pageId="26" pageNumber="607" startId="26.[92,105,1546,1567]" targetBox="[189,949,584,1516]" targetPageId="26">
<paragraph id="804843274D6672676352A07A434CF97A" blockId="26.[92,1048,1545,1674]" pageId="26" pageNumber="607">
F IGS. 4246.
<taxonomicName id="47F738A44D66726763FEA07A4049F9EF" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[240,551,1545,1567]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="607" phylum="Arthropoda" rank="species" species="singulare" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D66726763FEA07A4343F9EF" box="[240,301,1546,1567]" italics="true" pageId="26" pageNumber="607">Empis</emphasis>
(
<emphasis id="B2839F354D6672676235A07943D0F9EF" box="[315,446,1545,1567]" italics="true" pageId="26" pageNumber="607">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D66726762C0A0794049F9EF" box="[462,551,1545,1567]" italics="true" pageId="26" pageNumber="607">singulare</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D6672676121A07A4006F9EF" box="[559,616,1546,1567]" pageId="26" pageNumber="607" rank="species">sp. n.</taxonomicNameLabel>
: (42) male sternites 7 and 8, ventral view (scale 0.1 mm); (43) male segment 8, dorsal view (scale 0.1 mm); (44) male sternite 6, ventral view (scale 0.1 mm); (45) male hypopygium, lateral view (scale 0.1 mm); (46) male left cercus, dorsal view (scale 0.1 mm). st 7 5 sternite 7; st 8 5 sternite 8; tg 8 5 tergite 8.
</paragraph>
</caption>
<paragraph id="804843274D6772666352A2044296FB7B" blockId="27.[92,1046,1139,1227]" box="[92,248,1139,1163]" pageId="27" pageNumber="608">
<emphasis id="B2839F354D6772666352A2044296FB7B" box="[92,248,1139,1163]" italics="true" pageId="27" pageNumber="608">Type material</emphasis>
</paragraph>
<paragraph id="804843274D677266638AA2E341A6FB3B" blockId="27.[92,1046,1139,1227]" pageId="27" pageNumber="608">
<materialsCitation id="309F497A4D677266638AA2E34326FB3B" collectingDate="1970-08-28" collectionCode="NHM" collectorName="P. H. Ward" country="Nigeria" location="Samaru" pageId="27" pageNumber="608" specimenCount="1" typeStatus="holotype">
<typeStatus id="5F4CFD854D677266638AA2E34292FB5B" box="[132,252,1171,1195]" pageId="27" pageNumber="608" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F8E003B74D677266623DA2E343E9FB5B" box="[307,391,1171,1195]" name="Nigeria" pageId="27" pageNumber="608">Nigeria</collectingCountry>
,
<location id="852815FC4D6772666292A2E3439DFB5B" LSID="urn:lsid:plazi:treatment:085EF2314D647260625BA50D41A3FC37:852815FC4D6772666292A2E3439DFB5B" box="[412,499,1171,1195]" country="Nigeria" name="Samaru" pageId="27" pageNumber="608">Samaru</location>
,
<collectingMethod id="59B63B304D6772666106A2E34151FB5A" box="[520,831,1171,1195]" pageId="27" pageNumber="608">mercury vapour light trap</collectingMethod>
,
<date id="F44965E74D677266605AA2E34660FB5B" box="[852,1038,1171,1195]" pageId="27" pageNumber="608" value="1970-08-28">
<collectingDate id="E40D9C0F4D677266605AA2E34660FB5B" box="[852,1038,1171,1195]" pageId="27" pageNumber="608" value="1970-08-28">28 August 1970</collectingDate>
</date>
,
<collectorName id="2D0226F14D6772666352A2C3428EFB3B" box="[92,224,1203,1227]" pageId="27" pageNumber="608">P. H. Ward</collectorName>
(
<collectionCode id="E6E6DBE24D67726663FAA2C34357FB3A" box="[244,313,1203,1226]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="27" pageNumber="608" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="309F497A4D6772666259A2C341ADFB3B" box="[343,963,1203,1227]" collectingDate="1970-09-08" collectionCode="MNHN" collectorName="P. H. Ward" country="Nigeria" location="Samaru" pageId="27" pageNumber="608" specimenCount="1" typeStatus="paratype">
<typeStatus id="5F4CFD854D6772666259A2C343AAFB3B" box="[343,452,1203,1227]" pageId="27" pageNumber="608" type="paratype">PARATYPE</typeStatus>
: 1, same data but`2±
<date id="F44965E74D67726661D6A2C34152FB3B" box="[728,828,1203,1227]" pageId="27" pageNumber="608" value="1970-09-08">
<collectingDate id="E40D9C0F4D67726661D6A2C34152FB3B" box="[728,828,1203,1227]" pageId="27" pageNumber="608" value="1970-09-08">8.ix.1970</collectingDate>
</date>
(
<collectionCode id="E6E6DBE24D6772666056A2C341DAFB3A" box="[856,948,1203,1226]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="27" pageNumber="608">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="804843274D6772666352A28342B4FAFA" blockId="27.[92,1047,1267,1674]" box="[92,218,1267,1290]" pageId="27" pageNumber="608">
<emphasis id="B2839F354D6772666352A28342B4FAFA" box="[92,218,1267,1290]" italics="true" pageId="27" pageNumber="608">Description</emphasis>
</paragraph>
<paragraph id="804843274D677266638AA36343BBFADA" blockId="27.[92,1047,1267,1674]" box="[132,469,1299,1323]" pageId="27" pageNumber="608">Male: length of wing 4.9 mm.</paragraph>
<paragraph id="804843274D677266638AA342466DFA3A" blockId="27.[92,1047,1267,1674]" pageId="27" pageNumber="608">Head with occiput black. Ocellar triangle prominent with pair of short bristles. Frons blackish. Face blackish, shining in lower part. Antenna blackish, pedicel a little lighter. Proboscis not very long (1.5 times the head height), labrum brown, labium blackish, slender, strongly sclerotized, with distinct annulations, palpus brown, reduced. Eyes narrowly separated on frons, ommatidia smaller at middle.</paragraph>
<paragraph id="804843274D677261638AA3A24086FABA" blockId="27.[92,1047,1267,1674]" lastBlockId="28.[92,1047,1299,1673]" lastPageId="28" lastPageNumber="609" pageId="27" pageNumber="608">Thorax dark brown, postpronotal lobe, postalar callus lighter. Antepronotum with distinct lateral bristles. Proepisternum bare. Prosternum with a few short lateral bristles. Postpronotal lobes with one distinct, not very strong basal bristle. One strong, long postsutural supraalar, one ®ne, shorter presutural. Two strong, long notopleurals. One strong, long postalar. Acrostichals and dorsocentrals irregularly biserial, ®ne, short, with one strong, long prescutellar bristle. Scutellum with two strong, long apical bristles, two ®ner, shorter subapicals. Laterotergite with fan of strong bristles. Anterior and posterior spiracles blackish.</paragraph>
<caption id="D48813AF4D6072616352A204401DFB29" pageId="28" pageNumber="609" startId="28.[92,105,1140,1161]" targetBox="[94,1045,118,1113]" targetPageId="28">
<paragraph id="804843274D6072616352A204401DFB29" blockId="28.[92,1047,1140,1242]" pageId="28" pageNumber="609">
FIG. 47. Distribution of species of the
<taxonomicName id="47F738A44D6072616109A2054139FB7A" authorityName="Collin" authorityYear="1930" box="[519,855,1140,1162]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D6072616109A205402AFB7A" box="[519,580,1141,1162]" italics="true" pageId="28" pageNumber="609">Empis</emphasis>
(
<emphasis id="B2839F354D607261615BA20440B6FB7A" box="[597,728,1140,1162]" italics="true" pageId="28" pageNumber="609">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D60726161E3A2044139FB7A" box="[749,855,1140,1162]" italics="true" pageId="28" pageNumber="609">chrysocera</emphasis>
</taxonomicName>
-group: 1 5
<taxonomicName id="47F738A44D60726160DFA205429DFB55" authorityName="Smith" authorityYear="1971" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="barbitos" subGenus="Coptophlebia">
<emphasis id="B2839F354D60726160DFA2054186FB7A" box="[977,1000,1141,1162]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D60726160F9A2044661FB7A" box="[1015,1039,1140,1162]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D60726163ADA2FF429DFB55" box="[163,243,1167,1189]" italics="true" pageId="28" pageNumber="609">barbitos</emphasis>
</taxonomicName>
; 2 5
<taxonomicName id="47F738A44D6072616222A2FF407EFB55" authorityName="Esa" authorityYear="2001" box="[300,528,1167,1189]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="brazzavillensis" subGenus="Coptophlebia">
<emphasis id="B2839F354D6072616222A2FF432DFB54" box="[300,323,1167,1188]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D607261625BA2FF4303FB55" box="[341,365,1167,1189]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D607261628CA2FF407EFB55" box="[386,528,1167,1189]" italics="true" pageId="28" pageNumber="609">brazzavillensis</emphasis>
</taxonomicName>
; 3 5
<taxonomicName id="47F738A44D6072616146A2FF4167FB55" authorityName="Collin" authorityYear="1930" box="[584,777,1167,1189]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D6072616146A2FF4031FB54" box="[584,607,1167,1188]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D607261617FA2FF40E7FB55" box="[625,649,1167,1189]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D6072616191A2FF4167FB55" box="[671,777,1167,1189]" italics="true" pageId="28" pageNumber="609">chrysocera</emphasis>
</taxonomicName>
; 4 5
<taxonomicName id="47F738A44D607261604FA2FF4661FB55" authorityName="Smith" authorityYear="1971" box="[833,1039,1167,1189]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="cuthbertsoni" subGenus="Coptophlebia">
<emphasis id="B2839F354D607261604FA2FF4136FB54" box="[833,856,1167,1188]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D6072616064A2FF41ECFB55" box="[874,898,1167,1189]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D6072616099A2FF4661FB55" box="[919,1039,1167,1189]" italics="true" pageId="28" pageNumber="609">cuthbertsoni</emphasis>
</taxonomicName>
; 5 5
<taxonomicName id="47F738A44D60726163C9A2DA431CFB4F" authorityName="Esa" authorityYear="2001" box="[199,370,1193,1215]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="juxtaripa" subGenus="Coptophlebia">
<emphasis id="B2839F354D60726163C9A2DA42B0FB4F" box="[199,222,1194,1215]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D60726163E3A2D9436BFB4F" box="[237,261,1193,1215]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D6072616219A2DA431CFB4F" box="[279,370,1194,1215]" italics="true" pageId="28" pageNumber="609">juxtaripa</emphasis>
</taxonomicName>
; 6 5
<taxonomicName id="47F738A44D60726162A8A2DA4072FB4F" authorityName="Smith" authorityYear="1967" box="[422,540,1193,1215]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D60726162A8A2DA43D3FB4F" box="[422,445,1194,1215]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D60726162C2A2D9438AFB4F" box="[460,484,1193,1215]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D60726162F8A2D94072FB4F" box="[502,540,1193,1215]" italics="true" pageId="28" pageNumber="609">lyra</emphasis>
</taxonomicName>
; 7 5
<taxonomicName id="47F738A44D607261615CA2DA4154FB4F" authorityName="Smith" authorityYear="1969" box="[594,826,1193,1215]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="machipandensis" subGenus="Coptophlebia">
<emphasis id="B2839F354D607261615CA2DA4007FB4F" box="[594,617,1194,1215]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D6072616176A2D940FEFB4F" box="[632,656,1193,1215]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D60726161AFA2D94154FB4F" box="[673,826,1193,1215]" italics="true" pageId="28" pageNumber="609">machipandensis</emphasis>
</taxonomicName>
; 8 5
<taxonomicName id="47F738A44D607261607EA2DA467EFB4F" authorityName="Esa" authorityYear="2001" box="[880,1040,1193,1215]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="plumata" subGenus="Coptophlebia">
<emphasis id="B2839F354D607261607EA2DA41E9FB4F" box="[880,903,1194,1215]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D6072616098A2D941C0FB4F" box="[918,942,1193,1215]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D60726160CEA2D9467EFB4F" box="[960,1040,1193,1215]" italics="true" pageId="28" pageNumber="609">plumata</emphasis>
</taxonomicName>
; 9 5
<taxonomicName id="47F738A44D60726163C9A2B543E7FB2A" authorityName="Esa" authorityYear="2001" box="[199,393,1220,1242]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="samaruensis" subGenus="Coptophlebia">
<emphasis id="B2839F354D60726163C9A2B542B0FB2A" box="[199,222,1221,1242]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D60726163E4A2B4436CFB2A" box="[234,258,1220,1242]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D607261621CA2B543E7FB2A" box="[274,393,1221,1242]" italics="true" pageId="28" pageNumber="609">samaruensis</emphasis>
</taxonomicName>
; 10 5
<taxonomicName id="47F738A44D60726162C6A2B54002FB2A" authorityName="Esa" authorityYear="2001" box="[456,620,1220,1242]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="609" phylum="Arthropoda" rank="species" species="singulare" subGenus="Coptophlebia">
<emphasis id="B2839F354D60726162C6A2B543B1FB2A" box="[456,479,1221,1242]" italics="true" pageId="28" pageNumber="609">E.</emphasis>
(
<emphasis id="B2839F354D60726162E2A2B4406DFB2A" box="[492,515,1220,1242]" italics="true" pageId="28" pageNumber="609">C.</emphasis>
)
<emphasis id="B2839F354D607261611DA2B44002FB2A" box="[531,620,1220,1242]" italics="true" pageId="28" pageNumber="609">singulare</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="804843274D607261638AA3234678F9B9" blockId="28.[92,1047,1299,1673]" pageId="28" pageNumber="609">Legs dark brown. Fore femur with only short bristles. Fore tibia with a few distinct short ventrals at apical tip, a few short dorsals on apical half. First tarsomere of fore tarsus short pennate dorsally. Mid femur with numerous very short dorsal bristles, a few posteroventral pennate bristles at apical tip. Mid tibia short pennate dorsally on apical two-thirds, with distinct ventral bristles. Mid tarsus with only short bristles. Hind femur short pennate dorsally except at tips, with one row of distinct, not very long ventral bristles. Hind tibia long pennate dorsally, antero- and posteroventrally on apical half. Hind tarsus with distinct ventral and dorsal bristles.</paragraph>
<paragraph id="804843274D607260638AA02143ADFF5E" blockId="28.[92,1047,1299,1673]" lastBlockId="29.[92,1048,118,525]" lastPageId="29" lastPageNumber="610" pageId="28" pageNumber="609">
Wing clear to brown at tip especially along
<collectionCode id="E6E6DBE24D6072616178A02140E4F999" box="[630,650,1617,1641]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="609">R</collectionCode>
21 3,
<collectionCode id="E6E6DBE24D60726161C1A021408CF999" box="[719,738,1617,1641]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="609">R</collectionCode>
4 and
<collectionCode id="E6E6DBE24D6072616025A0214151F999" box="[811,831,1617,1641]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="609">R</collectionCode>
5.
<collectionCode id="E6E6DBE24D6072616052A0214101F999" box="[860,879,1617,1641]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="609">R</collectionCode>
41 5 branched at right-angle. Discal cell strongly truncate. M1, M2 abbreviated. A1 complete, feebly sclerotized. Anal lobe well developed, anal angle acute. Halter with yellow base and black stem and knob.
</paragraph>
<paragraph id="804843274D617260638AA6C64360FE9D" blockId="29.[92,1048,118,525]" pageId="29" pageNumber="610">Abdomen (®gures 42±44) brown, lighter base with a few distinct bristles. Sternite 6 (®gure 44) thickened on its posterior margin, somewhat voluminous in lateral view. Tergite 7 somewhat membranous anteriorly. Sternite 7 fragmented in several membranous parts (®gure 42). Width of tergite 8 reduced laterally (®gure 43). Sternite 8 lengthened posteriorly, with a ventroanterior process on each side (®gures 42, 43).</paragraph>
<paragraph id="804843274D617260638AA70640C4FE1D" blockId="29.[92,1048,118,525]" pageId="29" pageNumber="610">Hypopygium (®gures 45, 46) with cercus very characteristic, made up of inner bare lobe and external well bristled lobe itself divided into anterior and posterior parts (®gure 46). Epandrial lamellae connected anteriorly with rounded strongly bristled tip. Phallus short, thick, not pointed at tip.</paragraph>
<paragraph id="804843274D617260638AA785433FFDFD" blockId="29.[92,1048,118,525]" box="[132,337,501,525]" pageId="29" pageNumber="610">Female unknown.</paragraph>
<paragraph id="804843274D6172606352A44F42BEFDA6" blockId="29.[92,1047,575,727]" box="[92,208,575,598]" pageId="29" pageNumber="610">
<emphasis id="B2839F354D6172606352A44F42BEFDA6" box="[92,208,575,598]" italics="true" pageId="29" pageNumber="610">Discussion</emphasis>
</paragraph>
<paragraph id="804843274D617260638AA42F415AFD26" blockId="29.[92,1047,575,727]" pageId="29" pageNumber="610">
<taxonomicName id="47F738A44D617260638AA42F4362FD86" authorityName="Esa" authorityYear="2001" box="[132,268,606,630]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="610" phylum="Arthropoda" rank="species" species="singulare">
<emphasis id="B2839F354D617260638AA42F4362FD86" box="[132,268,606,630]" italics="true" pageId="29" pageNumber="610">E. singulare</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="47F738A44D61726062C4A42F40F9FD86" authorityName="Collin" authorityYear="1930" box="[458,663,606,630]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="610" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D61726062C4A42F438AFD86" box="[458,484,607,630]" italics="true" pageId="29" pageNumber="610">E.</emphasis>
(
<emphasis id="B2839F354D61726062FAA42E4061FD86" box="[500,527,606,630]" italics="true" pageId="29" pageNumber="610">C.</emphasis>
)
<emphasis id="B2839F354D617260612DA42E40F9FD86" box="[547,663,606,630]" italics="true" pageId="29" pageNumber="610">chrysocera</emphasis>
</taxonomicName>
-complex. In general aspect, this species is very diOEerent from all others of this complex with darker wing at tip, two strong notopleurals, the presence of pennate bristles on mid and hind male femora and narrowly dichoptic eyes with ommatidia smaller at middle.
</paragraph>
<paragraph id="804843274D6172606352A5774319FCCF" blockId="29.[92,375,775,831]" box="[92,375,775,831]" pageId="29" pageNumber="610">
<emphasis id="B2839F354D6172606352A577428EFCEF" box="[92,224,775,799]" italics="true" pageId="29" pageNumber="610">Distribution</emphasis>
West Africa:
<collectingCountry id="F8E003B74D6172606212A557431EFCCF" box="[284,368,807,831]" name="Nigeria" pageId="29" pageNumber="610">Nigeria</collectingCountry>
.
</paragraph>
<paragraph id="804843274D6172606352A50042BDFC78" blockId="29.[92,1048,880,967]" box="[92,211,880,904]" pageId="29" pageNumber="610">
<emphasis id="B2839F354D6172606352A50042BDFC78" box="[92,211,880,904]" italics="true" pageId="29" pageNumber="610">Etymology</emphasis>
</paragraph>
<paragraph id="804843274D617260638AA5E141A3FC37" blockId="29.[92,1048,880,967]" pageId="29" pageNumber="610">
From the Latin word
<emphasis id="B2839F354D6172606279A5E043B1FC58" box="[375,479,912,936]" italics="true" pageId="29" pageNumber="610">singularis</emphasis>
meaning
<emphasis id="B2839F354D6172606143A5E040D6FC58" box="[589,696,912,936]" italics="true" pageId="29" pageNumber="610">particular</emphasis>
,
<emphasis id="B2839F354D61726061C8A5E04178FC58" box="[710,790,912,936]" italics="true" pageId="29" pageNumber="610">distinct</emphasis>
, in reference to several characters diOEerent of those usually found in the
<taxonomicName id="47F738A44D617260619BA5C04133FC37" authorityName="Collin" authorityYear="1930" box="[661,861,943,967]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="610" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D617260619BA5C040C1FC37" box="[661,687,944,967]" italics="true" pageId="29" pageNumber="610">E.</emphasis>
(
<emphasis id="B2839F354D61726061B3A5DF40B9FC37" box="[701,727,943,967]" italics="true" pageId="29" pageNumber="610">C.</emphasis>
)
<emphasis id="B2839F354D61726061E7A5DF4133FC37" box="[745,861,943,967]" italics="true" pageId="29" pageNumber="610">chrysocera</emphasis>
</taxonomicName>
-complex.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,165 @@
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<treatment id="085EF2314D6A72656265A2CC4663FCA3" ID-DOI="http://doi.org/10.5281/zenodo.5278970" ID-Zenodo-Dep="5278970" LSID="urn:lsid:plazi:treatment:085EF2314D6A72656265A2CC4663FCA3" httpUri="http://treatment.plazi.org/id/085EF2314D6A72656265A2CC4663FCA3" lastPageId="24" lastPageNumber="605" pageId="22" pageNumber="603">
<subSubSection id="C8ED10AC4D6A726B6265A2CC40E0FB04" pageId="22" pageNumber="603" type="nomenclature">
<paragraph id="804843274D6A726B6265A2CC4169FB23" blockId="22.[363,775,1212,1268]" box="[363,775,1212,1236]" pageId="22" pageNumber="603">
<heading id="DB00F44B4D6A726B6265A2CC4169FB23" box="[363,775,1212,1236]" centered="true" fontSize="10" level="2" pageId="22" pageNumber="603" reason="3">
<taxonomicName id="47F738A44D6A726B6265A2CC40D0FB24" ID-CoL="8BZ7K" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[363,702,1212,1236]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="603" phylum="Arthropoda" rank="species" species="plumata" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D6A726B6265A2CC40D0FB24" bold="true" box="[363,702,1212,1236]" pageId="22" pageNumber="603">
<emphasis id="B2839F354D6A726B6265A2CC43DEFB24" bold="true" box="[363,432,1212,1236]" italics="true" pageId="22" pageNumber="603">Empis</emphasis>
(
<emphasis id="B2839F354D6A726B62CCA2CC403DFB24" bold="true" box="[450,595,1212,1236]" italics="true" pageId="22" pageNumber="603">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D6A726B6168A2CC40D0FB24" bold="true" box="[614,702,1212,1236]" italics="true" pageId="22" pageNumber="603">plumata</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D6A726B61C6A2CC4169FB23" box="[712,775,1212,1235]" pageId="22" pageNumber="603" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="804843274D6A726B62EAA2AC40E0FB04" blockId="22.[363,775,1212,1268]" box="[484,654,1244,1268]" pageId="22" pageNumber="603">(®gures 36±38)</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D6A72656352A34042F4FDDD" lastPageId="24" lastPageNumber="605" pageId="22" pageNumber="603" type="materials_examined">
<paragraph id="804843274D6A726B6352A3404296FAB7" blockId="22.[92,1046,1327,1415]" box="[92,248,1327,1351]" pageId="22" pageNumber="603">
<emphasis id="B2839F354D6A726B6352A3404296FAB7" box="[92,248,1327,1351]" italics="true" pageId="22" pageNumber="603">Type material</emphasis>
</paragraph>
<paragraph id="804843274D6A726B638AA320406FFA77" blockId="22.[92,1046,1327,1415]" pageId="22" pageNumber="603">
<materialsCitation id="309F497A4D6A726B638AA3204661FA97" box="[132,1039,1359,1384]" collectingDate="1982-07-26" collectorName="de Djibelot &amp; J. Etienne" country="Senegal" location="Casamance" pageId="22" pageNumber="603" specimenCount="1" typeStatus="holotype">
<typeStatus id="5F4CFD854D6A726B638AA3204292FA97" box="[132,252,1359,1383]" pageId="22" pageNumber="603" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F8E003B74D6A726B6223A33F43ECFA97" box="[301,386,1359,1383]" name="Senegal" pageId="22" pageNumber="603">Senegal</collectingCountry>
,
<location id="852815FC4D6A726B629AA33F407DFA97" LSID="urn:lsid:plazi:treatment:085EF2314D6A72656265A2CC4663FCA3:852815FC4D6A726B629AA33F407DFA97" box="[404,531,1359,1383]" country="Senegal" name="Casamance" pageId="22" pageNumber="603">Casamance</location>
, foreÃt
<collectorName id="2D0226F14D6A726B6168A33F4086FA97" box="[614,744,1359,1383]" pageId="22" pageNumber="603">de Djibelot</collectorName>
,
<date id="F44965E74D6A726B61F7A33F41E4FA97" box="[761,906,1359,1383]" pageId="22" pageNumber="603" value="1982-07-26">
<collectingDate id="E40D9C0F4D6A726B61F7A33F41E4FA97" box="[761,906,1359,1383]" pageId="22" pageNumber="603" value="1982-07-26">26 July 1982</collectingDate>
</date>
,
<collectorName id="2D0226F14D6A726B6092A3204661FA97" box="[924,1039,1359,1383]" pageId="22" pageNumber="603">J. Etienne</collectorName>
</materialsCitation>
;
<materialsCitation id="309F497A4D6A726B6352A31F4392FA77" box="[92,508,1391,1415]" collectingDate="1982-07-26" collectionCode="MNHN" collectorName="de Djibelot &amp; J. Etienne" country="Senegal" elevation="1" location="Casamance" pageId="22" pageNumber="603" specimenCount="1" typeStatus="paratype">
<typeStatus id="5F4CFD854D6A726B6352A31F42A7FA77" box="[92,201,1391,1415]" pageId="22" pageNumber="603" type="paratype">PARATYPE</typeStatus>
:
<quantity id="470FEEC24D6A726B63DAA31F4297FA77" box="[212,249,1391,1415]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="603" unit="m" value="1.0">
<elevation id="0BDAA4144D6A726B63DAA31F4297FA77" box="[212,249,1391,1415]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="603" unit="m" value="1.0">1 m</elevation>
</quantity>
, same data (
<collectionCode id="E6E6DBE24D6A726B629FA3004383FA77" box="[401,493,1392,1415]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="603">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="804843274D6A726B6352A3C342B4FA3A" blockId="22.[92,1047,1459,1674]" box="[92,218,1459,1482]" pageId="22" pageNumber="603">
<emphasis id="B2839F354D6A726B6352A3C342B4FA3A" box="[92,218,1459,1482]" italics="true" pageId="22" pageNumber="603">Description</emphasis>
</paragraph>
<paragraph id="804843274D6A726B638AA3A243BBFA1A" blockId="22.[92,1047,1459,1674]" box="[132,469,1490,1514]" pageId="22" pageNumber="603">
<emphasis id="B2839F354D6A726B638AA3A242D0FA1A" box="[132,190,1490,1514]" italics="true" pageId="22" pageNumber="603">Male</emphasis>
: length of wing 4.6 mm.
</paragraph>
<paragraph id="804843274D6A726B638AA38140FAF979" blockId="22.[92,1047,1459,1674]" pageId="22" pageNumber="603">Head with occiput blackish to dusted greyish. Ocellar triangle blackish, prominent, without distinct bristle. Frons blackish. Face shining blackish. Antenna black. Proboscis long (three times the head height), labrum brownish, labium blackish, strongly sclerotized, with distinct annulations, labella slender, bare, palpus not visible. Eyes holoptic, upper ommatidia enlarged.</paragraph>
<caption id="D48813AF4D6B726A6352A4AC431CFCD6" pageId="23" pageNumber="604" startId="23.[92,105,732,753]" targetBox="[93,1045,119,704]" targetPageId="23">
<paragraph id="804843274D6B726A6352A4AC431CFCD6" blockId="23.[92,1047,731,806]" pageId="23" pageNumber="604">
FIGS 3638.
<taxonomicName id="47F738A44D6B726A63E4A4AC4077FD01" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[234,537,731,753]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="604" phylum="Arthropoda" rank="species" species="plumata" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D6B726A63E4A4AC4349FD01" box="[234,295,732,753]" italics="true" pageId="23" pageNumber="604">Empis</emphasis>
(
<emphasis id="B2839F354D6B726A623AA4AB43D9FD01" box="[308,439,731,753]" italics="true" pageId="23" pageNumber="604">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D6B726A62C7A4AB4077FD01" box="[457,537,731,753]" italics="true" pageId="23" pageNumber="604">plumata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D6B726A612CA4AC4035FD01" box="[546,603,732,753]" pageId="23" pageNumber="604" rank="species">sp. n.</taxonomicNameLabel>
: (36) epandrial lamella, lateral view (scale 0.1 mm); (37) male cercus, lateral view (scale 0.1 mm); (38) female tergite 7, dorsal view (scale 0.1 mm).
</paragraph>
</caption>
<paragraph id="804843274D6B726A638AA52443DFFB9B" blockId="23.[92,1048,852,1674]" pageId="23" pageNumber="604">Thorax blackish to dark brown. Antepronotu m with a few distinct lateral bristles. Postpronota l lobes shining brown, with one long, strong basal bristle. Proepisternum bare. Prosternum with a few distinct lateral bristles. Scutum without distinct stripe. Acrostichals and dorsocentrals uniserial, latter with one strong, long prescutellar bristle. One strong, long postsutural supraalar, a few ®ner, shorter others. Three strong, long notopleurals. Postalar calli brown with one strong, long bristle. Scutellum with two strong, long apical bristles, two ®ner, shorter subapicals. Laterotergite with fan of strong, long bristles. Anterior and posterior spiracles black and dark brown, respectively.</paragraph>
<paragraph id="804843274D6B726A638AA2044010FA5A" blockId="23.[92,1048,852,1674]" pageId="23" pageNumber="604">Legs dark brown, coxae lighter. Fore femur with one row of ®ne, long anteroventral bristles. Fore tibia with one row of strong, long anterodorsals except at basal tip. Mid femur with one row of strong, long anterodorsals, two rows, respectively antero- and posteroventral, of ®ner, long bristles. Mid tibia pennate anterodorsally except at basal tip, with numerous strong, long bristles antero- and posteroventrally as well as posterodorsally. First tarsomere of fore and mid tarsi with rather strong, not very long dorsals and ventrals. Hind femur with numerous distinct dorsals especially in the middle, ®ne antero- and posteroventrals. Hind tibia covered with rather strong, very long bristles especially dorsally. First tarsomere of hind tarsus with numerous distinct bristles, longer dorsally.</paragraph>
<paragraph id="804843274D6B726A638AA3C24679F9F9" blockId="23.[92,1048,852,1674]" pageId="23" pageNumber="604">
Wing brown. Sc abbreviated.
<collectionCode id="E6E6DBE24D6B726A62EAA3C24399FA3A" box="[484,503,1458,1482]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="604">R</collectionCode>
41 5 branched at right-angle,
<collectionCode id="E6E6DBE24D6B726A6052A3C24101FA3A" box="[860,879,1458,1482]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="604">R</collectionCode>
4 faint. Discal cell truncate. M1, M2 abbreviated. A1 complete, faint at tip. Anal lobe well developed, anal angle acute. Halter with brown base, yellowish stem, blackish knob.
</paragraph>
<paragraph id="804843274D6B726A638AA06143A1F999" blockId="23.[92,1048,852,1674]" pageId="23" pageNumber="604">
Abdomen with ®rst three segments yellowish with a few distinct bristles especially at base, otherwise brown, lighter than thorax. Modi®cations of male pregenital segments similar to
<emphasis id="B2839F354D6B726A624FA02243A1F999" box="[321,463,1618,1641]" italics="true" pageId="23" pageNumber="604">
<taxonomicName id="47F738A44D6B726A624FA02243A9F999" authorityName="Esa" authorityYear="2001" box="[321,455,1618,1641]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="604" phylum="Arthropoda" rank="species" species="juxtaripa">E. juxtaripa</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="804843274D6B7265638AA0014111FF5E" blockId="23.[92,1048,852,1674]" lastBlockId="24.[92,1047,118,557]" lastPageId="24" lastPageNumber="605" pageId="23" pageNumber="604">
Hypopygium (®gures 36, 37) similar to
<taxonomicName id="47F738A44D6B726A616EA0024085F979" authorityName="Esa" authorityYear="2001" box="[608,747,1650,1673]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="604" phylum="Arthropoda" rank="species" species="juxtaripa">
<emphasis id="B2839F354D6B726A616EA0024085F979" box="[608,747,1650,1673]" italics="true" pageId="23" pageNumber="604">E. juxtaripa</emphasis>
</taxonomicName>
except for the following characters: cercus with rather narrower rounded posterior lobe and truncate anterior one (®gure 37). Dorsal bristles of epandrial lamella longer (®gure 36).
</paragraph>
<paragraph id="804843274D647265638AA6C642F4FDDD" blockId="24.[92,1047,118,557]" pageId="24" pageNumber="605">Female similar to male except for the following characters: distinct pair of ocellar bristles. Eyes dichoptic, lower ommatidia enlarged. Frons shining black. Numerous distinct presutural supraalar. About 8±10 strong notopleurals. Legs a little darker. Fore femur pennate ventrally except at apical one-quarter. Fore tibia pennate ventrally on apical half. First tarsomere of fore tarsus pennate dorsally, with a few ®ne ventral pennate bristles. Mid femur pennate dorsally except at basal tip and apical one-quarter, pennate posteroventrally, and anteroventrally on apical half. Mid tibia pennate dorsally except at tips. Hind femur pennate dorsally except at tips and ventrally except at basal tip. Hind tibia densely pennate dorsally except at basal tip. Abdomen shining brown to dusted brown on tergites 7±10. Tergite 7 with U-shaped anterior margin (®gure 38). Cercus dusted brown, longer than broad, with bristly hairs.</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D6472656352A4284663FCA3" pageId="24" pageNumber="605" type="discussion">
<paragraph id="804843274D6472656352A428466FFD7F" blockId="24.[92,1025,600,655]" box="[92,1025,600,655]" pageId="24" pageNumber="605">
<emphasis id="B2839F354D6472656352A4284293FD7F" box="[92,253,600,655]" italics="true" pageId="24" pageNumber="605">
Discussion
<taxonomicName id="47F738A44D647265638AA4084293FD7F" authorityName="Esa" authorityYear="2001" box="[132,253,631,655]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="605" phylum="Arthropoda" rank="species" species="plumata">E. plumata</taxonomicName>
</emphasis>
belongs to the
<taxonomicName id="47F738A44D64726562BDA408405FFD7F" authorityName="Smith" authorityYear="1967" box="[435,561,631,655]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="605" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D64726562BDA40843A3FD7F" box="[435,461,632,655]" italics="true" pageId="24" pageNumber="605">E.</emphasis>
(
<emphasis id="B2839F354D64726562D5A407439BFD7F" box="[475,501,631,655]" italics="true" pageId="24" pageNumber="605">C.</emphasis>
)
<emphasis id="B2839F354D6472656108A407405FFD7F" box="[518,561,631,655]" italics="true" pageId="24" pageNumber="605">lyra</emphasis>
</taxonomicName>
-complex. See discussion in
<taxonomicName id="47F738A44D647265607DA4084194FD7F" authorityName="Esa" authorityYear="2001" box="[883,1018,632,655]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="605" phylum="Arthropoda" rank="species" species="juxtaripa">
<emphasis id="B2839F354D647265607DA4084194FD7F" box="[883,1018,632,655]" italics="true" pageId="24" pageNumber="605">E. juxtaripa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="804843274D6472656352A4C94317FD01" blockId="24.[92,377,697,753]" box="[92,377,697,753]" pageId="24" pageNumber="605">
<emphasis id="B2839F354D6472656352A4C9428EFD21" box="[92,224,697,721]" italics="true" pageId="24" pageNumber="605">Distribution</emphasis>
West Africa:
<collectingCountry id="F8E003B74D6472656212A4A9431FFD01" box="[284,369,729,753]" name="Senegal" pageId="24" pageNumber="605">Senegal</collectingCountry>
.
</paragraph>
<paragraph id="804843274D6472656352A56C4663FCA3" blockId="24.[92,1037,796,851]" box="[92,1037,796,851]" pageId="24" pageNumber="605">
<emphasis id="B2839F354D6472656352A56C42BDFCC4" box="[92,211,796,820]" italics="true" pageId="24" pageNumber="605">Etymology</emphasis>
The name of species is derived from the Latin word
<emphasis id="B2839F354D64726561ECA54B412AFCA3" box="[738,836,827,851]" italics="true" pageId="24" pageNumber="605">plumatus</emphasis>
meaning
<emphasis id="B2839F354D64726560B6A54B466BFCA3" box="[952,1029,827,851]" italics="true" pageId="24" pageNumber="605">feather</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,221 @@
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<mods:title id="A531CC05AB22C522F1A767E296CB3B38">Cladistics and taxonomy of the Afrotropical Empis (Coptophlebia) chrysocera-group (Diptera, Empididae)</mods:title>
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<mods:namePart id="3FA0196EF849F7CABBC76E93022B9B78">Esa, C. Daugeron</mods:namePart>
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<mods:date id="86AF02D2653AE2A8CF458CE2A37B2B4E">2001</mods:date>
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<treatment id="085EF2314D6D726E6258A5ED40FEFA3A" LSID="urn:lsid:plazi:treatment:085EF2314D6D726E6258A5ED40FEFA3A" httpUri="http://treatment.plazi.org/id/085EF2314D6D726E6258A5ED40FEFA3A" lastPageId="19" lastPageNumber="600" pageId="17" pageNumber="598">
<subSubSection id="C8ED10AC4D6D726C6258A5ED40E0FC25" pageId="17" pageNumber="598" type="nomenclature">
<paragraph id="804843274D6D726C6258A5ED4172FC45" blockId="17.[92,796,925,1013]" box="[342,796,925,949]" pageId="17" pageNumber="598">
<heading id="DB00F44B4D6D726C6258A5ED4172FC45" box="[342,796,925,949]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="598" reason="3">
<taxonomicName id="47F738A44D6D726C6258A5ED4172FC45" ID-CoL="8BZ67" authority="Smith, 1967 b" authorityName="Smith" authorityYear="1967" box="[342,796,925,949]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D6D726C6258A5ED4012FC45" bold="true" box="[342,636,925,949]" pageId="17" pageNumber="598">
<emphasis id="B2839F354D6D726C6258A5ED43F5FC45" bold="true" box="[342,411,925,949]" italics="true" pageId="17" pageNumber="598">Empis</emphasis>
(
<emphasis id="B2839F354D6D726C62A3A5ED4050FC45" bold="true" box="[429,574,925,949]" italics="true" pageId="17" pageNumber="598">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D6D726C615FA5ED4012FC45" bold="true" box="[593,636,925,949]" italics="true" pageId="17" pageNumber="598">lyra</emphasis>
</emphasis>
<bibRefCitation id="E4663ED64D6D726C6188A5ED4172FC45" author="SMITH, K. G. V." box="[646,796,925,949]" pageId="17" pageNumber="598" pagination="16" refId="ref14452" refString="SMITH, K. G. V., 1967 b, Afrikanische Empididae (Dipt.), Stuttgarter BeitraEge zur Naturkunde aus dem Staatlichen Museum fuEr Naturkunde in Stuttgart, 179, 1 ± 16." type="journal article" year="1967">Smith, 1967b</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="804843274D6D726C62EAA5CD40E0FC25" blockId="17.[92,796,925,1013]" box="[484,654,957,982]" pageId="17" pageNumber="598">(®gures 26±32)</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D6D726C6352A5904070FC05" box="[92,542,991,1013]" pageId="17" pageNumber="598" type="reference_group">
<paragraph id="804843274D6D726C6352A5904070FC05" blockId="17.[92,796,925,1013]" box="[92,542,991,1013]" pageId="17" pageNumber="598">
<treatmentCitationGroup id="A0E764094D6D726C6352A5904070FC05" box="[92,542,991,1013]" pageId="17" pageNumber="598">
<treatmentCitation id="015665364D6D726C6352A5904076FC05" author="SMITH, K. G. V." box="[92,536,991,1013]" page="11" pageId="17" pageNumber="598" year="1967">
<taxonomicName id="47F738A44D6D726C6352A5904076FC05" ID-CoL="8BZ67" authority="Smith, 1967 b: 11" authorityName="Smith" authorityPageNumber="11" authorityYear="1967" box="[92,536,991,1013]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D6D726C6352A59042F7FC05" box="[92,153,992,1013]" italics="true" pageId="17" pageNumber="598">Empis</emphasis>
(
<emphasis id="B2839F354D6D726C63A8A5AF4347FC05" box="[166,297,991,1013]" italics="true" pageId="17" pageNumber="598">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D6D726C6235A5AF430FFC05" box="[315,353,991,1013]" italics="true" pageId="17" pageNumber="598">lyra</emphasis>
<bibRefCitation id="E4663ED64D6D726C6264A5AF4076FC05" author="SMITH, K. G. V." box="[362,536,991,1013]" pageId="17" pageNumber="598" pagination="16" refId="ref14452" refString="SMITH, K. G. V., 1967 b, Afrikanische Empididae (Dipt.), Stuttgarter BeitraEge zur Naturkunde aus dem Staatlichen Museum fuEr Naturkunde in Stuttgart, 179, 1 ± 16." type="journal article" year="1967">Smith, 1967b: 11</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D6D726E6352A24243B5FAFA" lastPageId="19" lastPageNumber="600" pageId="17" pageNumber="598" type="materials_examined">
<paragraph id="804843274D6D726C6352A2424296FBB9" blockId="17.[92,1047,1073,1192]" box="[92,248,1073,1097]" pageId="17" pageNumber="598">
<emphasis id="B2839F354D6D726C6352A2424296FBB9" box="[92,248,1073,1097]" italics="true" pageId="17" pageNumber="598">Type material</emphasis>
</paragraph>
<paragraph id="804843274D6D726C638AA22141C4FB58" blockId="17.[92,1047,1073,1192]" pageId="17" pageNumber="598">
<materialsCitation id="309F497A4D6D726C638AA22142CFFB78" collectingDate="1948-06-01" collectorName="M. Steele" country="Kenya" location="Kwali Forest" pageId="17" pageNumber="598" specimenCount="1" typeStatus="holotype">
<typeStatus id="5F4CFD854D6D726C638AA2214292FB99" box="[132,252,1105,1129]" pageId="17" pageNumber="598" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F8E003B74D6D726C6224A221431AFB99" box="[298,372,1105,1129]" name="Kenya" pageId="17" pageNumber="598">Kenya</collectingCountry>
,
<location id="852815FC4D6D726C628BA2214074FB98" LSID="urn:lsid:plazi:treatment:085EF2314D6D726E6258A5ED40FEFA3A:852815FC4D6D726C628BA2214074FB98" box="[389,538,1105,1129]" country="Kenya" name="Kwali Forest" pageId="17" pageNumber="598">Kwali Forest</location>
,
<quantity id="470FEEC24D6D726C6124A22140E9FB99" box="[554,647,1105,1129]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="17" pageNumber="598" unit="mi" value="20.0">20 miles</quantity>
W. of Mombasa,
<date id="F44965E74D6D726C6053A221418BFB99" box="[861,997,1105,1129]" pageId="17" pageNumber="598" value="1948-06-01">
<collectingDate id="E40D9C0F4D6D726C6053A221418BFB99" box="[861,997,1105,1129]" pageId="17" pageNumber="598" value="1948-06-01">1 June 1948</collectingDate>
</date>
,
<collectorName id="2D0226F14D6D726C60FBA22142CFFB78" pageId="17" pageNumber="598">M. Steele</collectorName>
</materialsCitation>
;
<materialsCitation id="309F497A4D6D726C63A1A201438BFB78" box="[175,485,1136,1160]" collectingDate="1948-06-01" collectorName="M. Steele" country="Kenya" elevation="2" location="Kwali Forest" pageId="17" pageNumber="598" specimenCount="1" typeStatus="paratype">
<typeStatus id="5F4CFD854D6D726C63A1A2014345FB78" box="[175,299,1136,1160]" pageId="17" pageNumber="598" type="paratype">PARATYPES</typeStatus>
:
<quantity id="470FEEC24D6D726C6236A2004330FB77" box="[312,350,1136,1160]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="598" unit="m" value="2.0">
<elevation id="0BDAA4144D6D726C6236A2004330FB77" box="[312,350,1136,1160]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="598" unit="m" value="2.0">2 m</elevation>
</quantity>
, same data
</materialsCitation>
,
<materialsCitation id="309F497A4D6D726C62F6A20043FDFB58" collectingDate="1958-11-20" collectionCode="NHM" collectorName="Lindner" country="Zimbabwe" elevation="1" location="Livingstone" pageId="17" pageNumber="598" specimenCount="1" typeStatus="paratype">
<quantity id="470FEEC24D6D726C62F6A2004073FB77" box="[504,541,1136,1160]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="598" unit="m" value="1.0">
<elevation id="0BDAA4144D6D726C62F6A2004073FB77" box="[504,541,1136,1160]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="598" unit="m" value="1.0">1 m</elevation>
</quantity>
,
<collectingCountry id="F8E003B74D6D726C6121A20040C9FB78" box="[559,679,1136,1160]" name="Zimbabwe" pageId="17" pageNumber="598">Zimbabwe</collectingCountry>
,
<location id="852815FC4D6D726C61B4A2004150FB78" LSID="urn:lsid:plazi:treatment:085EF2314D6D726E6258A5ED40FEFA3A:852815FC4D6D726C61B4A2004150FB78" box="[698,830,1136,1160]" country="Zimbabwe" name="Livingstone" pageId="17" pageNumber="598">Livingstone</location>
, 16±
<date id="F44965E74D6D726C6074A20042FEFB58" pageId="17" pageNumber="598" value="1958-11-20">
<collectingDate id="E40D9C0F4D6D726C6074A20042FEFB58" pageId="17" pageNumber="598" value="1958-11-20">20 November 1958</collectingDate>
</date>
,
<collectorName id="2D0226F14D6D726C63AFA2E04292FB58" box="[161,252,1168,1192]" pageId="17" pageNumber="598">Lindner</collectorName>
leg. (
<collectionCode id="E6E6DBE24D6D726C6231A2E143EAFB58" box="[319,388,1169,1192]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="17" pageNumber="598" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="309F497A4D6D726C62ADA2E041CBFB58" box="[419,933,1168,1192]" collectingDate="1958-11-20" collectionCode="SMNS" collectorName="Lindner" country="Zimbabwe" elevation="2" location="Livingstone" pageId="17" pageNumber="598" specimenCount="1" typeStatus="paratype">
<quantity id="470FEEC24D6D726C62ADA2E04391FB57" box="[419,511,1168,1192]" metricMagnitude="0" metricUnit="m" metricValue="2.3" pageId="17" pageNumber="598" unit="m" value="2.3">
<elevation id="0BDAA4144D6D726C62ADA2E04391FB57" box="[419,511,1168,1192]" metricMagnitude="0" metricUnit="m" metricValue="2.3" pageId="17" pageNumber="598" unit="m" value="2.3">2, 3 m</elevation>
</quantity>
, same data than preceding (
<collectionCode id="E6E6DBE24D6D726C6049A2E041F9FB58" box="[839,919,1168,1192]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="598">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="804843274D6D726C6352A2A342B4FB1A" blockId="17.[92,1047,1235,1673]" box="[92,218,1235,1258]" pageId="17" pageNumber="598">
<emphasis id="B2839F354D6D726C6352A2A342B4FB1A" box="[92,218,1235,1258]" italics="true" pageId="17" pageNumber="598">Description</emphasis>
</paragraph>
<paragraph id="804843274D6D726C638AA28343BBFAFA" blockId="17.[92,1047,1235,1673]" box="[132,469,1267,1291]" pageId="17" pageNumber="598">Male: length of wing 4.8 mm.</paragraph>
<paragraph id="804843274D6D726C638AA36343FCFA1A" blockId="17.[92,1047,1235,1673]" pageId="17" pageNumber="598">Head with occiput dusted greyish. Ocellar triangle prominent, dusted greyish, with pair of distinct short bristles. Face broad, dusted greyish to blackish, shining in lower part. Scape and pedicel rather brownish to yellowish, ®rst ¯agellomere yellowish, second and third ¯agellomeres brown. Proboscis long (2.5 times the head height), labrum brown to yellowish, labium brownish to blackish, strongly sclerotized, with distinct annulations, labella slender, bare, palpus brown. Eyes holoptic, upper ommatidia enlarged.</paragraph>
<paragraph id="804843274D6D726F638AA38142C2FB5B" blockId="17.[92,1047,1235,1673]" lastBlockId="18.[92,1047,1011,1674]" lastPageId="18" lastPageNumber="599" pageId="17" pageNumber="598">Thorax brown to dusted greyish. Antepronotum with distinct lateral bristles. Postpronotal lobes shining dark brown, with one rather strong, long basal bristle. Proepisternum bare. Prosternum with a few lateral bristles. Scutum extensively dusted greyish, with three dark brown stripes, the ®rst under acrostichals, the two lyre-shaped others under dorsocentrals. Acrostichals uniserial, ®ne, short. Dorsocentrals uniserial, rather ®ne, short, with one strong, long prescutellar bristle. One distinct, not very long presutural supraalar. One strong, long postsutural. Three strong, more or less long notopleurals. One strong, long postalar. Scutellum with two strong, long apicals, two ®ner, shorter subapicals. Laterotergite with fan of strong, long bristles. Anterior and posterior spiracles respectively black and dark brown.</paragraph>
<caption id="D48813AF4D6E726F6206A5DC4104FC31" box="[264,874,939,961]" pageId="18" pageNumber="599" startId="18.[264,277,940,961]" targetBox="[124,1016,119,913]" targetPageId="18">
<paragraph id="804843274D6E726F6206A5DC4104FC31" blockId="18.[264,874,939,961]" box="[264,874,939,961]" pageId="18" pageNumber="599">
F IG. 26.
<taxonomicName id="47F738A44D6E726F6261A5DC40D4FC31" authority="Smith" authorityName="Smith" authorityYear="1967" box="[367,698,939,961]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="599" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D6E726F6261A5DC43C2FC31" box="[367,428,940,961]" italics="true" pageId="18" pageNumber="599">Empis</emphasis>
(
<emphasis id="B2839F354D6E726F62B4A5DB4053FC31" box="[442,573,939,961]" italics="true" pageId="18" pageNumber="599">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D6E726F6140A5DB401AFC31" box="[590,628,939,961]" italics="true" pageId="18" pageNumber="599">lyra</emphasis>
Smith
</taxonomicName>
, habitus, female.
</paragraph>
</caption>
<paragraph id="804843274D6E726F638AA2C34168FA1A" blockId="18.[92,1047,1011,1674]" pageId="18" pageNumber="599">Legs brown. Fore femur with a few distinct basoventral bristles. Fore tibia with one row of strong, rather long posterodorsal bristles, numerous ®ne ventrals. First tarsomere of fore tarsus with strong, rather short dorsal bristles. Mid femur pennate posteroventrally on apical half, with ®ne, long antero- and posteroventral bristles at basal tip. Mid tibia with one row of posteroventral and posterodorsal strong bristles, pennate dorsally and ventrally, ventrals shorter and only at basal onequarter. Hind femur with only short dorsal bristles, almost pennate apically, distinct rather ®ne ventrals, pennate antero- and posteroventrally at apical one-quarter. Hind tibia pennate like mid tibia. First tarsomere of mid and hind tarsi with numerous strong, not very long bristles, especially dorsally.</paragraph>
<paragraph id="804843274D6E726F638AA381428DF999" blockId="18.[92,1047,1011,1674]" pageId="18" pageNumber="599">
Wing (®gure 26) dark brown. Sc abbreviated.
<collectionCode id="E6E6DBE24D6E726F61A6A38140D5F9F9" box="[680,699,1521,1545]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="599">R</collectionCode>
41 5 branched at right-angle. M1, M2 abbreviated. A1 complete, faintly sclerotized towards the wing margin. Anal lobe well developed, anal angle acute. Halter with yellowish base and stem, black knob.
</paragraph>
<paragraph id="804843274D6E726E638AA00143A4FEDD" blockId="18.[92,1047,1011,1674]" lastBlockId="19.[92,1047,118,909]" lastPageId="19" lastPageNumber="600" pageId="18" pageNumber="599">Abdomen (®gures 27, 28) with ®rst three segments yellowish, otherwise dark brown. A few distinct bristles at base. Tergite 6 with rather long, strongly sclerotized, rounded medioposterior expansion (®gure 27). Tergite 7 divided into somewhat desclerotized median part and two strongly sclerotized lateral ones (®gure 27). Tergite 8 distinctly divided in a median sclerite, expanded anteroventrally, and two strongly sclerotized wing-shaped lateral ones (®gure 28). Sternite 8 developed posteriorly with rounded shape.</paragraph>
<paragraph id="804843274D6F726E638AA7464127FE3D" blockId="19.[92,1047,118,909]" pageId="19" pageNumber="600">
Hypopygium (®gures 30±32) with cercus made up of not very developed almost bare anterior lobe and well-bristled posterior lobe (®gure 30). Epandrial lamellae connected anteriorly, halter-shaped (elongated, enlarged at tip), with distinct bristles at apical tip (®gure 31), a group of strong, not very long bristles on inner face (®gure 32). Hypandrium and phallus similar to
<taxonomicName id="47F738A44D6F726E618CA7C6412CFE3D" authorityName="Esa" authorityYear="2001" box="[642,834,437,461]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="600" phylum="Arthropoda" rank="species" species="brazzavillensis">
<emphasis id="B2839F354D6F726E618CA7C6412CFE3D" box="[642,834,437,461]" italics="true" pageId="19" pageNumber="600">E. brazzavillensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="804843274D6F726E638AA7A541ADFC7C" blockId="19.[92,1047,118,909]" pageId="19" pageNumber="600">Female (®gure 26) similar to male except for the following characters: scape, pedicel distinctly yellowish. Frons as broad as face, shining blackish in upper part, dusted greyish in lower part. Eyes dichoptic, lower ommatidia a little enlarged. Fore femur short, long pennate on basal half dorsally and ventrally. Fore tibia long pennate dorsally, with short, ®ne ventral subpennate bristles. First tarsomere of fore tarsus long pennate dorsally, next two tarsomeres with a few dorsal pennate bristles. Mid femur with very short pennate dorsobasal bristles, a little longer on dorsoapical half, long pennate ventrally. Mid tibia pennate dorsally except at basal tip, ventrally on basal half. Hind femur pennate dorsally except at apical tip, long pennate anteroventrally except at basal tip, pennate posteroventrally at apical half. First three tarsomeres of mid and hind tarsi similar to these of fore tarsus. First six segments of abdomen brown to yellowish, other segments brown to blackish, without distinct bristles except at base. Tergite 7 with circular-shaped anterior and posterior margins (®gure 29). Cercus longer than broad, with ®ne, short bristly hairs.</paragraph>
<paragraph id="804843274D6F726E6352A5C542ADFC3C" blockId="19.[92,1047,949,1291]" box="[92,195,949,972]" pageId="19" pageNumber="600">
<emphasis id="B2839F354D6F726E6352A5C542ADFC3C" box="[92,195,949,972]" italics="true" pageId="19" pageNumber="600">Variation</emphasis>
</paragraph>
<paragraph id="804843274D6F726E638AA5A443B5FAFA" blockId="19.[92,1047,949,1291]" pageId="19" pageNumber="600">
<materialsCitation id="309F497A4D6F726E638AA5A443B9FAFA" country="Kenya" location="In" municipality="There" pageId="19" pageNumber="600" specimenCount="2" typeStatus="holotype">
<location id="852815FC4D6F726E638AA5A442F3FC1B" LSID="urn:lsid:plazi:treatment:085EF2314D6D726E6258A5ED40FEFA3A:852815FC4D6F726E638AA5A442F3FC1B" box="[132,157,980,1003]" country="Kenya" municipality="There" name="In" pageId="19" pageNumber="600">In</location>
the type series, two groups of specimens can be distinguished.
<collectingMunicipality id="602CD95D4D6F726E6078A5A341D6FC1B" box="[886,952,979,1003]" pageId="19" pageNumber="600">There</collectingMunicipality>
are two main variations between the ®rst group, collected in
<collectingCountry id="F8E003B74D6F726E61B2A5844168FBFC" box="[700,774,1012,1036]" name="Kenya" pageId="19" pageNumber="600">Kenya</collectingCountry>
, and the second group, collected in
<collectingCountry id="F8E003B74D6F726E63EFA2634337FBDB" box="[225,345,1043,1067]" name="Zimbabwe" pageId="19" pageNumber="600">Zimbabwe</collectingCountry>
: ®rst, the female abdomen of specimens from
<collectingCountry id="F8E003B74D6F726E6054A26441CBFBDC" box="[858,933,1044,1068]" name="Kenya" pageId="19" pageNumber="600">Kenya</collectingCountry>
is entirely brown, not shining, whereas the abdomen of specimens from
<collectingCountry id="F8E003B74D6F726E603EA24341C6FBBB" box="[816,936,1075,1099]" name="Zimbabwe" pageId="19" pageNumber="600">Zimbabwe</collectingCountry>
is lighter to subshining yellowish at base; second, the apical tip of epandrial lamella of male
<typeStatus id="5F4CFD854D6F726E6352A20342AEFB7B" box="[92,192,1139,1163]" pageId="19" pageNumber="600" type="holotype">holotype</typeStatus>
(from
<collectingCountry id="F8E003B74D6F726E6219A204430FFB7C" box="[279,353,1140,1164]" name="Kenya" pageId="19" pageNumber="600">Kenya</collectingCountry>
) is a little longer than this of specimens from
<collectingCountry id="F8E003B74D6F726E6096A203467EFB7B" box="[920,1040,1139,1163]" name="Zimbabwe" pageId="19" pageNumber="600">Zimbabwe</collectingCountry>
. Unfortunately the male genitalia of the
<typeStatus id="5F4CFD854D6F726E612DA2E340E9FB5B" box="[547,647,1171,1195]" pageId="19" pageNumber="600" type="holotype">holotype</typeStatus>
are damaged and cercus not easily visible. Nevertheless there are no diOEerences in the shape of pregenital sclerites. Thus until the discovery of new male specimens from
<collectingCountry id="F8E003B74D6F726E6193A2A34089FB1B" box="[669,743,1235,1259]" name="Kenya" pageId="19" pageNumber="600">Kenya</collectingCountry>
, I consider all males and females of type series conspeci®c
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D6F726E6352A34340FEFA3A" pageId="19" pageNumber="600" type="discussion">
<paragraph id="804843274D6F726E6352A3434663FA9A" blockId="19.[92,1037,1331,1386]" box="[92,1037,1331,1386]" pageId="19" pageNumber="600">
<emphasis id="B2839F354D6F726E6352A34342BEFA9A" box="[92,208,1331,1386]" italics="true" pageId="19" pageNumber="600">
Discussion
<taxonomicName id="47F738A44D6F726E638AA32342BEFA9A" authorityName="Smith" authorityYear="1967" box="[132,208,1362,1386]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="600" phylum="Arthropoda" rank="species" species="lyra">E. lyra</taxonomicName>
</emphasis>
belongs to the
<taxonomicName id="47F738A44D6F726E628BA323406DFA9A" authorityName="Smith" authorityYear="1967" box="[389,515,1362,1386]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="600" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D6F726E628BA32343F1FA9A" box="[389,415,1363,1386]" italics="true" pageId="19" pageNumber="600">E.</emphasis>
(
<emphasis id="B2839F354D6F726E62A2A32243A8FA9A" box="[428,454,1362,1386]" italics="true" pageId="19" pageNumber="600">C.</emphasis>
)
<emphasis id="B2839F354D6F726E62D6A322406DFA9A" box="[472,515,1362,1386]" italics="true" pageId="19" pageNumber="600">lyra</emphasis>
</taxonomicName>
-complex. See discussion in
<taxonomicName id="47F738A44D6F726E604BA3234668FA9A" authorityName="Esa" authorityYear="2001" box="[837,1030,1362,1386]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="600" phylum="Arthropoda" rank="species" species="brazzavillensis">
<emphasis id="B2839F354D6F726E604BA3234668FA9A" box="[837,1030,1362,1386]" italics="true" pageId="19" pageNumber="600">E. brazzavillensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="804843274D6F726E6352A3E240FEFA3A" blockId="19.[92,656,1426,1483]" box="[92,656,1426,1483]" pageId="19" pageNumber="600">
<emphasis id="B2839F354D6F726E6352A3E2428EFA5A" box="[92,224,1426,1450]" italics="true" pageId="19" pageNumber="600">Distribution</emphasis>
East and southern Africa:
<collectingCountry id="F8E003B74D6F726E62B8A3C3406EFA3B" box="[438,512,1459,1483]" name="Kenya" pageId="19" pageNumber="600">Kenya</collectingCountry>
,
<collectingCountry id="F8E003B74D6F726E611FA3C240E7FA3A" box="[529,649,1458,1482]" name="Zimbabwe" pageId="19" pageNumber="600">Zimbabwe</collectingCountry>
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</paragraph>
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View file

@ -0,0 +1,158 @@
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<mods:title id="30EC71A3726B567B94BC5436F633C1FE">Cladistics and taxonomy of the Afrotropical Empis (Coptophlebia) chrysocera-group (Diptera, Empididae)</mods:title>
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<heading id="DB00F44B4D707271623EA4ED412FFD45" box="[304,833,669,693]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="593" reason="3">
<taxonomicName id="47F738A44D707271623EA4ED412FFD45" ID-CoL="8BZ4D" authority="Smith, 1971" authorityName="Smith" authorityYear="1971" box="[304,833,669,693]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="593" phylum="Arthropoda" rank="species" species="cuthbertsoni" subGenus="Coptophlebia">
<emphasis id="B2839F354D707271623EA4ED40DEFD45" bold="true" box="[304,688,669,693]" pageId="12" pageNumber="593">
<emphasis id="B2839F354D707271623EA4ED431BFD45" bold="true" box="[304,373,669,693]" italics="true" pageId="12" pageNumber="593">Empis</emphasis>
(
<emphasis id="B2839F354D7072716288A4ED4079FD45" bold="true" box="[390,535,669,693]" italics="true" pageId="12" pageNumber="593">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D7072716125A4ED40DEFD45" bold="true" box="[555,688,669,693]" italics="true" pageId="12" pageNumber="593">cuthbertsoni</emphasis>
</emphasis>
<bibRefCitation id="E4663ED64D70727161B4A4ED412FFD45" author="SMITH, K. G. V." box="[698,833,669,693]" pageId="12" pageNumber="593" pagination="96" refId="ref14522" refString="SMITH, K. G. V., 1971, The sub-Saharan species of Rhamphomyia and Coptophlebia with abbreviated discal venation (Diptera: Empididae), Annals of the Natal Museum, 21, 77 ± 96." type="journal article" year="1971">Smith, 1971</bibRefCitation>
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</paragraph>
<paragraph id="804843274D70727162EAA4CD40E0FD25" blockId="12.[92,833,669,757]" box="[484,654,701,725]" pageId="12" pageNumber="593">(®gures 15±20)</paragraph>
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<emphasis id="B2839F354D7072716352A49042F7FD05" box="[92,153,736,757]" italics="true" pageId="12" pageNumber="593">Empis</emphasis>
(
<emphasis id="B2839F354D70727163A8A4AF4347FD05" box="[166,297,735,757]" italics="true" pageId="12" pageNumber="593">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D7072716235A4AF43DDFD05" box="[315,435,735,757]" italics="true" pageId="12" pageNumber="593">cuthbertsoni</emphasis>
<bibRefCitation id="E4663ED64D70727162B2A4AF4033FD05" author="SMITH, K. G. V." box="[444,605,735,757]" pageId="12" pageNumber="593" pagination="96" refId="ref14522" refString="SMITH, K. G. V., 1971, The sub-Saharan species of Rhamphomyia and Coptophlebia with abbreviated discal venation (Diptera: Empididae), Annals of the Natal Museum, 21, 77 ± 96." type="journal article" year="1971">Smith, 1971: 90</bibRefCitation>
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.
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</paragraph>
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<paragraph id="804843274D7072716352A5424296FCB9" blockId="12.[92,1046,817,905]" box="[92,248,817,841]" pageId="12" pageNumber="593">
<emphasis id="B2839F354D7072716352A5424296FCB9" box="[92,248,817,841]" italics="true" pageId="12" pageNumber="593">Type material</emphasis>
</paragraph>
<paragraph id="804843274D707271638AA522409CFC79" blockId="12.[92,1046,817,905]" pageId="12" pageNumber="593">
<materialsCitation id="309F497A4D707271638AA5224083FC79" collectingDate="1938-12-26" collectionCode="NHM" collectorName="A. Cuthbertson &amp; Scabiosa'" country="Zimbabwe" elevation="1" location="Wedza" pageId="12" pageNumber="593" specimenCount="1" typeStatus="paratype">
<typeStatus id="5F4CFD854D707271638AA5224291FC99" box="[132,255,849,873]" pageId="12" pageNumber="593" type="paratype">PARATYPES</typeStatus>
: 1,
<collectingCountry id="F8E003B74D7072716242A52143AAFC99" box="[332,452,849,873]" name="Zimbabwe" pageId="12" pageNumber="593">Zimbabwe</collectingCountry>
,
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,
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<collectingDate id="E40D9C0F4D707271613AA5214163FC99" box="[564,781,849,873]" pageId="12" pageNumber="593" value="1938-12-26">26 December 1938</collectingDate>
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,
<collectorName id="2D0226F14D707271602FA52141BAFC99" box="[801,980,849,873]" pageId="12" pageNumber="593">A. Cuthbertson</collectorName>
;
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, with the additional data`at ¯owers of
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(
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)
</materialsCitation>
.
</paragraph>
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<emphasis id="B2839F354D7072716352A5C542B4FC3C" box="[92,218,949,972]" italics="true" pageId="12" pageNumber="593">Description</emphasis>
</paragraph>
<paragraph id="804843274D707271638AA5A343BBFC1B" blockId="12.[92,1048,949,1674]" box="[132,469,979,1003]" pageId="12" pageNumber="593">Male: length of wing 4.5 mm.</paragraph>
<paragraph id="804843274D707271638AA58341C5FB7B" blockId="12.[92,1048,949,1674]" pageId="12" pageNumber="593">Head with occiput blackish. Ocellar triangle prominent with pair of ®ne, very short bristles. Face blackish, subshining in lower part. Scape and pedicel black, ¯agellum missing. Proboscis long (twice the head height). Labrum yellowish to brown, labium blackish, with distinct annulations, strongly sclerotized, labella slender, bare, palpus yellowish. Eyes holoptic, upper ommatidia enlarged.</paragraph>
<paragraph id="804843274D707271638AA2E342AEFA5A" blockId="12.[92,1048,949,1674]" pageId="12" pageNumber="593">Thorax subshining blackish to strongly dusted grey in prescutellar depression. Antepronotum with ®ne, short lateral bristles. Proepisternum bare. Prosternum with a few ®ne, short lateral bristles. Postpronotal lobes brownish with only a few ®ne, short bristles, apparently without distinct basal one. Acrostichals and dorsocentrals irregularly biserial, ®ne, very short, latter with one strong, long prescutellar bristle. One distinct postsutural supraalar. One strong, long notopleural. One strong, long postalar. Scutellum dark brown with two apical bristles, two ®ner, shorter subapicals. Laterotergite with fan of strong, long bristles. Anterior and posterior spiracles blackish.</paragraph>
<paragraph id="804843274D707271638AA3C34176F999" blockId="12.[92,1048,949,1674]" pageId="12" pageNumber="593">Legs dark brown to blackish. Fore femur with only a few ®ne, rather long basoventral bristles. Fore tibia with short bristles, a little longer at apical tip. Mid femur with one row of distinct rather long ventral bristles. Mid tibia with distinct short ventral and dorsal bristles. Hind femur with distinct rather long posteroventra l bristles on basal half. Hind tibia with distinct rather short basal and ventral bristles. Tarsi with distinct short bristles especially on ®rst tarsomere.</paragraph>
<paragraph id="804843274D707273638AA001404DFCA8" blockId="12.[92,1048,949,1674]" lastBlockId="14.[92,1047,801,1548]" lastPageId="14" lastPageNumber="595" pageId="12" pageNumber="593">
Wing clear. Sc abbreviated.
<collectionCode id="E6E6DBE24D70727162C4A00143B3F979" box="[458,477,1649,1673]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="593">R</collectionCode>
41 5 branched at right-angle. Discal cell strongly truncate. M1, M2 abbreviated. Anal lobe well developed, anal angle acute. Halter with yellow base and stem, black knob.
</paragraph>
<caption id="D48813AF4D7172706352A31440FDF9E9" pageId="13" pageNumber="594" startId="13.[92,105,1380,1401]" targetBox="[95,1045,119,1352]" targetPageId="13">
<paragraph id="804843274D7172706352A31440FDF9E9" blockId="13.[92,1049,1380,1561]" pageId="13" pageNumber="594">
F IGS 1520.
<taxonomicName id="47F738A44D71727063E4A31540EEFA8A" authority="Smith" authorityName="Smith" authorityYear="1971" box="[234,640,1380,1402]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="594" phylum="Arthropoda" rank="species" species="cuthbertsoni" subGenus="Coptophlebia">
<emphasis id="B2839F354D71727063E4A3154349FA8A" box="[234,295,1381,1402]" italics="true" pageId="13" pageNumber="594">Empis</emphasis>
(
<emphasis id="B2839F354D717270623DA31443D8FA8A" box="[307,438,1380,1402]" italics="true" pageId="13" pageNumber="594">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D71727062CAA3144052FA8A" box="[452,572,1380,1402]" italics="true" pageId="13" pageNumber="594">cuthbertsoni</emphasis>
Smith
</taxonomicName>
: (15) male segment 6, lateral view (scale 0.1 mm); (16) male sternite 6, ventral view (scale 0.1 mm); (17) male segments 7 and 8, lateral view (scale 0.1 mm); (18) female tergite 7, dorsal view (scale 0.1 mm); (19) male hypopygium, lateral view (scale 0.1 mm); (20) cerci, epandrial lamellae with their sclerotized anterior connection and subepandrial sclerite, anterodorsal view (scale 0.1 mm). st 7 5 sternite 7; st 8 5 sternite 8; tg 7 5 tergite 7; tg 8 5 tergite 8; cerc 5 cercus; epn 5 epandrium; scl sub 5 subepandrial sclerite.
</paragraph>
</caption>
<paragraph id="804843274D727273638AA5104668FC06" blockId="14.[92,1047,801,1548]" pageId="14" pageNumber="595">Abdomen (®gures 15±17) with ®rst three segments yellowish, otherwise dark brown to blackish, distinct bristles at base. Sternite 6 voluminous (®gures 15, 16), modi®ed in two strongly sclerotized rounded medioposterior processes ventrally. Sternite 7 membranous (®gure 17). Sternite 8 with dorso- and ventroanterior processes on each side (®gure 17). Width of tergite 8 reduced laterally (®gure 17).</paragraph>
<paragraph id="804843274D727273638AA58D40B5FB63" blockId="14.[92,1047,801,1548]" pageId="14" pageNumber="595">Hypopygium (®gures 19, 20) with cercus bent back anteriorly in dorsal view, made up of an anterodorsal bare lobe and posterior rounded well-bristled lobe (®gures 19, 20). Epandrial lamellae connected anteriorly (®gure 20), with distinct bristles only at tip. Hypandrium well sclerotized, with very short bristly hairs at tip (®gure 19). Phallus short, not pointed at tip (®gure 19).</paragraph>
<paragraph id="804843274D727273638AA2EA4105F9FB" blockId="14.[92,1047,801,1548]" pageId="14" pageNumber="595">Female similar to male except for the following characters: ¯agellum black, scape and pedicel lighter. Eyes dichoptic, lower ommatidia somewhat enlarged, frons broad, shining dark brown. Fore femur with one row of very short dorsal bristles, pennate ventrally except at basal tip. Fore tibia long pennate dorsally, with a few pennate ventral bristles at apical tip. First tarsomere of fore tarsus long pennate dorsally, short pennate ventrally. Second tarsomere of fore tarsus with a few pennate dorsal bristles. Mid femur long pennate ventrally. Mid tibia short pennate ventrally on basal half. Hind femur long pennate ventrally and dorsally except at basal tip. Hind tibia long pennate dorsally, and ventrally except at apical tip, with also a few short pennate dorsal and ventral bristles especially in the middle. Abdomen dark brown, tergite 7 with somewhat circular-shaped anterior margin, more or less desclerotized in the middle (®gure 18). Cercus longer than broad, with only bristly hairs.</paragraph>
<paragraph id="804843274D7272736352A04342BEF9BA" blockId="14.[92,1047,1587,1673]" box="[92,208,1587,1610]" pageId="14" pageNumber="595">
<emphasis id="B2839F354D7272736352A04342BEF9BA" box="[92,208,1587,1610]" italics="true" pageId="14" pageNumber="595">Discussion</emphasis>
</paragraph>
<paragraph id="804843274D727272638AA02341D6FF3E" blockId="14.[92,1047,1587,1673]" lastBlockId="15.[92,1047,118,206]" lastPageId="15" lastPageNumber="596" pageId="14" pageNumber="595">
<emphasis id="B2839F354D727273638AA0234344F99A" box="[132,298,1618,1642]" italics="true" pageId="14" pageNumber="595">
<taxonomicName id="47F738A44D727273638AA023434EF99A" box="[132,288,1618,1642]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="595" phylum="Arthropoda" rank="species" species="cuthbertson">E. cuthbertson</taxonomicName>
i
</emphasis>
belongs to the
<taxonomicName id="47F738A44D72727362DFA02340FDF99A" authorityName="Collin" authorityYear="1930" box="[465,659,1618,1642]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="595" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D72727362DFA0234385F99A" box="[465,491,1619,1642]" italics="true" pageId="14" pageNumber="595">E.</emphasis>
(
<emphasis id="B2839F354D72727362F8A022407EF99A" box="[502,528,1618,1642]" italics="true" pageId="14" pageNumber="595">C.</emphasis>
)
<emphasis id="B2839F354D7272736111A02240FDF99A" box="[543,659,1618,1642]" italics="true" pageId="14" pageNumber="595">chrysocera</emphasis>
</taxonomicName>
-complex. The following combination of characters distinguishes this species from all others belonging to the
<taxonomicName id="47F738A44D72727260C2A00242BEFF7E" authorityName="Collin" authorityYear="1930" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" lastPageId="15" lastPageNumber="596" order="Diptera" pageId="14" pageNumber="595" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D72727360C2A0024188F979" box="[972,998,1650,1673]" italics="true" pageId="14" pageNumber="595">E.</emphasis>
(
<emphasis id="B2839F354D72727360FAA0014660F979" box="[1012,1038,1649,1673]" italics="true" pageId="14" pageNumber="595">C.</emphasis>
)
<emphasis id="B2839F354D7372726352A60642BEFF7E" box="[92,208,118,142]" italics="true" pageId="15" pageNumber="596">chrysocera</emphasis>
</taxonomicName>
-group: ¯agellum black, presence of one strong notopleural, prescutellar depression strongly greyish dusted, hind tibiae with dorsal bristles shorter than tibiae width, male sternite 6 considerably voluminous, phallus not pointed at tip.
</paragraph>
<paragraph id="804843274D7372726352A68A43A5FEC1" blockId="15.[92,459,250,306]" box="[92,459,250,306]" pageId="15" pageNumber="596">
<emphasis id="B2839F354D7372726352A68A428EFEE2" box="[92,224,250,274]" italics="true" pageId="15" pageNumber="596">Distribution</emphasis>
Southern Africa:
<collectingCountry id="F8E003B74D7372726242A76A43AAFEC2" box="[332,452,282,306]" name="Zimbabwe" pageId="15" pageNumber="596">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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View file

@ -0,0 +1,169 @@
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<mods:title id="601F5EF3F243D16FF0362048958FA4E9">Cladistics and taxonomy of the Afrotropical Empis (Coptophlebia) chrysocera-group (Diptera, Empididae)</mods:title>
</mods:titleInfo>
<mods:name id="159FEFF5F612912DB76B5A685B6B1A05" type="personal">
<mods:role id="9C5DC7E9A353D5DE9E18267262786195">
<mods:roleTerm id="5C134D1ABA7A19781EE60569F45D782F">Author</mods:roleTerm>
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<mods:namePart id="BD346B07354654109E1BC61D4130321A">Esa, C. Daugeron</mods:namePart>
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<mods:titleInfo id="C3B8C84EECFF85DE84884BAD5FD478FA">
<mods:title id="04A2759D627E3C4E32839694A88D7309">Journal of Natural History</mods:title>
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<mods:part id="0FA4BA1B958C2D9082DFA6D15BA40C9F">
<mods:date id="D4236C5F0F19BD8CCDC0864E5F54660D">2001</mods:date>
<mods:detail id="2AF7F107B4FF4B50E21E08DD0887BBEC" type="volume">
<mods:number id="82BFBED8D5F3A65207EAD77C7055B26F">35</mods:number>
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<mods:identifier id="365883D803CD0607CAAB807031F63B0A" type="ISSN">1464-5262</mods:identifier>
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<treatment id="085EF2314D73726C626BA72D4291FC83" ID-DOI="http://doi.org/10.5281/zenodo.5278968" ID-Zenodo-Dep="5278968" LSID="urn:lsid:plazi:treatment:085EF2314D73726C626BA72D4291FC83" httpUri="http://treatment.plazi.org/id/085EF2314D73726C626BA72D4291FC83" lastPageId="17" lastPageNumber="598" pageId="15" pageNumber="596">
<subSubSection id="C8ED10AC4D737272626BA72D40E0FE65" pageId="15" pageNumber="596" type="nomenclature">
<paragraph id="804843274D737272626BA72D4163FE84" blockId="15.[357,781,349,405]" box="[357,781,349,373]" pageId="15" pageNumber="596">
<heading id="DB00F44B4D737272626BA72D4163FE84" box="[357,781,349,373]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="596" reason="3">
<taxonomicName id="47F738A44D737272626BA72D40AAFE85" ID-CoL="8BZ5J" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[357,708,349,373]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="596" phylum="Arthropoda" rank="species" species="juxtaripa" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D737272626BA72D40AAFE85" bold="true" box="[357,708,349,373]" pageId="15" pageNumber="596">
<emphasis id="B2839F354D737272626BA72D43C4FE85" bold="true" box="[357,426,349,373]" italics="true" pageId="15" pageNumber="596">Empis</emphasis>
(
<emphasis id="B2839F354D73727262B2A72D4023FE85" bold="true" box="[444,589,349,373]" italics="true" pageId="15" pageNumber="596">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D737272616EA72D40AAFE85" bold="true" box="[608,708,349,373]" italics="true" pageId="15" pageNumber="596">juxtaripa</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D73727261C0A72D4163FE84" box="[718,781,349,372]" pageId="15" pageNumber="596" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="804843274D73727262EAA70D40E0FE65" blockId="15.[357,781,349,405]" box="[484,654,381,405]" pageId="15" pageNumber="596">(®gures 21±25)</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D73726C6352A7A24291FC83" lastPageId="17" lastPageNumber="598" pageId="15" pageNumber="596" type="materials_examined">
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<emphasis id="B2839F354D7372726352A7A24296FE19" box="[92,248,465,489]" italics="true" pageId="15" pageNumber="596">
<typeStatus id="5F4CFD854D7372726352A7A242FAFE19" box="[92,148,466,489]" pageId="15" pageNumber="596">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="804843274D737272638AA78243FDFDD9" blockId="15.[92,1046,465,553]" pageId="15" pageNumber="596">
<materialsCitation id="309F497A4D737272638AA78243E1FDD9" collectingDate="1970-08-13" collectionCode="NHM" collectorName="NW State &amp; P. H. Ward" country="Nigeria" location="Kontagora river" pageId="15" pageNumber="596" specimenCount="1" typeStatus="holotype">
H OLOTYPE:,
<collectingCountry id="F8E003B74D737272622AA7814316FDF9" box="[292,376,497,521]" name="Nigeria" pageId="15" pageNumber="596">Nigeria</collectingCountry>
,
<collectorName id="2D0226F14D7372726288A782439AFDF9" box="[390,500,497,521]" pageId="15" pageNumber="596">NW State</collectorName>
,
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,
<quantity id="470FEEC24D73727261C8A781417AFDF9" box="[710,788,497,521]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="15" pageNumber="596" unit="mi" value="3.0">3 miles</quantity>
from
<collectingCountry id="F8E003B74D7372726059A78141F8FDF9" box="[855,918,497,521]" name="Niger" pageId="15" pageNumber="596">Niger</collectingCountry>
,
<date id="F44965E74D73727260AAA78142FEFDD9" pageId="15" pageNumber="596" value="1970-08-13">
<collectingDate id="E40D9C0F4D73727260AAA78142FEFDD9" pageId="15" pageNumber="596" value="1970-08-13">13 August 1970</collectingDate>
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,
<collectorName id="2D0226F14D73727263AFA462434BFDD9" box="[161,293,529,553]" pageId="15" pageNumber="596">P. H. Ward</collectorName>
(
<collectionCode id="E6E6DBE24D7372726234A4624311FDD9" box="[314,383,530,553]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="15" pageNumber="596" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="804843274D7372726352A42642B4FD9D" blockId="15.[92,1047,598,1673]" box="[92,218,598,621]" pageId="15" pageNumber="596">
<emphasis id="B2839F354D7372726352A42642B4FD9D" box="[92,218,598,621]" italics="true" pageId="15" pageNumber="596">Description</emphasis>
</paragraph>
<paragraph id="804843274D737272638AA40443BBFD7C" blockId="15.[92,1047,598,1673]" box="[132,469,628,652]" pageId="15" pageNumber="596">Male: length of wing 3.2 mm.</paragraph>
<paragraph id="804843274D737272638AA4E442ADFCBC" blockId="15.[92,1047,598,1673]" pageId="15" pageNumber="596">Head with occiput dark brown. Ocellar triangle prominent, without distinct bristles. Face dark brown, shining in lower part. Antenna blackish, scape and pedicel a little lighter than ¯agellum, ¯agellum rather short. Proboscis long (twice the head height), labrum brown, labium blackish, strongly sclerotized, with distinct annulations, labella slender, bare, palpus brown. Eyes holoptic, upper ommatidia enlarged.</paragraph>
<paragraph id="804843274D737272638AA52443D6FB7B" blockId="15.[92,1047,598,1673]" pageId="15" pageNumber="596">Thorax yellowish to brown. Antepronotum yellowish with a few ®ne, short lateral bristles. Postpronotal lobes yellow, with one distinct basal bristle. Proepisternum bare. Prosternum with a few ®ne, short lateral bristles. Scutum yellowish to brownish, with broad darker stripes between acrostichals and dorsocentrals, and dorsocentrals and notopleuron. Acrostichals uniserial, ®ne, short. Dorsocentrals uni- to biserial, rather ®ne, short, with one strong, long prescutellar bristle. One strong, long postsutural supraalar, a few ®ner, shorter others. Three strong, long notopleurals. One strong, long postalar. Scutellum with the insertion of pair of strong apicals. Laterotergite with fan of strong, long bristles. Anterior and posterior spiracles brown.</paragraph>
<paragraph id="804843274D737272638AA2E340FBFA3A" blockId="15.[92,1047,598,1673]" pageId="15" pageNumber="596">Legs with coxa yellowish, femur, tibia and tarsus dark brown. Fore femur with only a few ®ne bristles. Fore tibia with one row of strong, long dorsals, with one or two ®ne pennate bristles at apical tip. First tarsomere of fore tarsus somewhat dilated, with ®ve or six ®ne dorsal pennate bristles. Mid femur with ®ne, rather long dorsal and ventral bristles. Mid tibia with one row of strong, long posterodorsals, two rows, respectively antero- and posterodorsal, of pennate bristles. First tarsomere of mid tarsus with distinct bristles, especially ventrally. Hind femur with ®ne dorsals, two rows, respectively antero- and posteroventral, of ®ne bristles. Hind tibia covered with numerous strong, long dorsals, shorter ventrals. First tarsomere of hind tarsus with rather long dorsals, shorter distinct ventrals.</paragraph>
<paragraph id="804843274D737272638AA3A24306F9B9" blockId="15.[92,1047,598,1673]" pageId="15" pageNumber="596">
Wing brown. Sc abbreviated.
<collectionCode id="E6E6DBE24D73727262D1A3A2439CFA1A" box="[479,498,1490,1514]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="596">R</collectionCode>
41 5 branched at right-angle. Discal cell rather short, truncate. M1, M2 abbreviated. A1 complete, faintly sclerotized towards the wing margin. Anal lobe well developed, anal angle acute. Halter with yellow base and stem, brown knob.
</paragraph>
<paragraph id="804843274D73726C638AA021414EFF1E" blockId="15.[92,1047,598,1673]" lastBlockId="17.[92,1047,118,430]" lastPageId="17" lastPageNumber="598" pageId="15" pageNumber="596">Abdomen (®gures 21±23) dark brown, lighter at base, with a few distinct brownish bristles at base. Tergite 6 unmodi®ed. Tergite 7 made up of two lateral well-sclerotized parts bearing a small dorsal process, and a central part, less sclerotized anteriorly (®gure 22). Tergite 8 divided into median part expanded anteroventrally (®gure 23) and wing-shaped lateral parts. Sternite 8 developed posteriorly, with triangular shape and distinct anterodorsal process (®gure 21).</paragraph>
<caption id="D48813AF4D6C726D6352A3BF4375F9BF" pageId="16" pageNumber="597" startId="16.[92,105,1487,1508]" targetBox="[95,1044,119,1458]" targetPageId="16">
<paragraph id="804843274D6C726D6352A3BF4375F9BF" blockId="16.[92,1047,1486,1615]" pageId="16" pageNumber="597">
FIGS 2125.
<taxonomicName id="47F738A44D6C726D63E6A3BF4048FA14" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[232,550,1486,1508]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="597" phylum="Arthropoda" rank="species" species="juxtaripa" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D6C726D63E6A3BF434BFA14" box="[232,293,1487,1508]" italics="true" pageId="16" pageNumber="597">Empis</emphasis>
(
<emphasis id="B2839F354D6C726D623BA3BE43D6FA14" box="[309,440,1486,1508]" italics="true" pageId="16" pageNumber="597">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D6C726D62C5A3BF4048FA14" box="[459,550,1487,1508]" italics="true" pageId="16" pageNumber="597">juxtaripa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D6C726D6121A3BF4005FA14" box="[559,619,1487,1508]" pageId="16" pageNumber="597" rank="species">sp. n.</taxonomicNameLabel>
: (21) male sternite 8, lateral view (scale 0.1 mm); (22) male tergite 7, dorsal view (scale 0.1 mm); (23) male tergite 8, lateral view (scale 0.1 mm); (24) male cercus and epandrial lamella, lateral view (scale 0.1 mm); (25) hypandrium and phallus, lateral view (scale 0.1 mm). cerc 5 cercus; epn 5 epandrium.
</paragraph>
</caption>
<paragraph id="804843274D6D726C638AA6864072FE7E" blockId="17.[92,1047,118,430]" pageId="17" pageNumber="598">Hypopygium (®gures 24, 25) with cercus made up of two rounded lobes: a broad, almost bare anterior one, a narrower bristled posterior one (®gure 24). Epandrial lamellae connected anteriorly, developed dorsally in rounded lobe with at least six strong inner bristles (®gure 24). Hypandrium desclerotized in its median part. Phallus rather short, not very thick (®gure 25).</paragraph>
<paragraph id="804843274D6D726C638AA7E6433FFE5D" blockId="17.[92,1047,118,430]" box="[132,337,406,430]" pageId="17" pageNumber="598">Female unknown.</paragraph>
<paragraph id="804843274D6D726C6352A7A942BEFE00" blockId="17.[92,1047,473,655]" box="[92,208,473,496]" pageId="17" pageNumber="598">
<emphasis id="B2839F354D6D726C6352A7A942BEFE00" box="[92,208,473,496]" italics="true" pageId="17" pageNumber="598">Discussion</emphasis>
</paragraph>
<paragraph id="804843274D6D726C638AA78941BFFD7F" blockId="17.[92,1047,473,655]" pageId="17" pageNumber="598">
<taxonomicName id="47F738A44D6D726C638AA7894365FDE0" authorityName="Esa" authorityYear="2001" box="[132,267,505,528]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="juxtaripa">
<emphasis id="B2839F354D6D726C638AA7894365FDE0" box="[132,267,505,528]" italics="true" pageId="17" pageNumber="598">E. juxtaripa</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="47F738A44D6D726C62CDA789402CFDE0" authorityName="Smith" authorityYear="1967" box="[451,578,504,528]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D6D726C62CDA78943B2FDE0" box="[451,476,505,528]" italics="true" pageId="17" pageNumber="598">E.</emphasis>
(
<emphasis id="B2839F354D6D726C62E4A788406BFDE0" box="[490,517,504,528]" italics="true" pageId="17" pageNumber="598">C.</emphasis>
)
<emphasis id="B2839F354D6D726C6119A788402CFDE0" box="[535,578,504,528]" italics="true" pageId="17" pageNumber="598">lyra</emphasis>
</taxonomicName>
-complex. It resembles
<taxonomicName id="47F738A44D6D726C6041A78941A7FDE0" authorityName="Esa" authorityYear="2001" box="[847,969,504,528]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="plumata">
<emphasis id="B2839F354D6D726C6041A78941A7FDE0" box="[847,969,504,528]" italics="true" pageId="17" pageNumber="598">E. plumata</emphasis>
</taxonomicName>
in the triangular shape of sternite 8, the shape and the pilosity of epandrial lamella, and the presence of numerous long dorsal bristles on hind tibia. They can be distinguished by their size (
<taxonomicName id="47F738A44D6D726C63F8A4284312FD9F" authorityName="Esa" authorityYear="2001" box="[246,380,600,623]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="juxtaripa">
<emphasis id="B2839F354D6D726C63F8A4284312FD9F" box="[246,380,600,623]" italics="true" pageId="17" pageNumber="598">E. juxtaripa</emphasis>
</taxonomicName>
is smaller), the length of proboscis (longer in
<taxonomicName id="47F738A44D6D726C609DA4284662FD9F" authorityName="Esa" authorityYear="2001" box="[915,1036,599,623]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="plumata">
<emphasis id="B2839F354D6D726C609DA4284662FD9F" box="[915,1036,599,623]" italics="true" pageId="17" pageNumber="598">E. plumata</emphasis>
</taxonomicName>
) and the colour of thorax (yellowish in
<taxonomicName id="47F738A44D6D726C6113A40840CAFD7F" authorityName="Esa" authorityYear="2001" box="[541,676,632,655]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="juxtaripa">
<emphasis id="B2839F354D6D726C6113A40840CAFD7F" box="[541,676,632,655]" italics="true" pageId="17" pageNumber="598">E. juxtaripa</emphasis>
</taxonomicName>
, brownish in
<taxonomicName id="47F738A44D6D726C604AA40841D0FD7F" authorityName="Esa" authorityYear="2001" box="[836,958,631,655]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="598" phylum="Arthropoda" rank="species" species="plumata">
<emphasis id="B2839F354D6D726C604AA40841D0FD7F" box="[836,958,631,655]" italics="true" pageId="17" pageNumber="598">E. plumata</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="804843274D6D726C6352A4C94319FD01" blockId="17.[92,375,697,753]" box="[92,375,697,753]" pageId="17" pageNumber="598">
<emphasis id="B2839F354D6D726C6352A4C9428EFD21" box="[92,224,697,721]" italics="true" pageId="17" pageNumber="598">Distribution</emphasis>
West Africa:
<collectingCountry id="F8E003B74D6D726C6212A4A9431EFD01" box="[284,368,729,753]" name="Nigeria" pageId="17" pageNumber="598">Nigeria</collectingCountry>
.
</paragraph>
<paragraph id="804843274D6D726C6352A56C42BDFCC4" blockId="17.[92,1047,796,883]" box="[92,211,796,820]" pageId="17" pageNumber="598">
<emphasis id="B2839F354D6D726C6352A56C42BDFCC4" box="[92,211,796,820]" italics="true" pageId="17" pageNumber="598">Etymology</emphasis>
</paragraph>
<paragraph id="804843274D6D726C638AA54C4291FC83" blockId="17.[92,1047,796,883]" pageId="17" pageNumber="598">
<emphasis id="B2839F354D6D726C638AA54C4283FCA3" box="[132,237,828,851]" italics="true" pageId="17" pageNumber="598">Juxtaripa</emphasis>
is a combination of the Latin words
<emphasis id="B2839F354D6D726C61BAA54C4083FCA3" box="[692,749,828,851]" italics="true" pageId="17" pageNumber="598">juxta</emphasis>
, meaning
<emphasis id="B2839F354D6D726C6065A54C41F5FCA3" box="[875,923,828,851]" italics="true" pageId="17" pageNumber="598">near</emphasis>
, and
<emphasis id="B2839F354D6D726C60EAA54C4661FCA3" box="[996,1039,828,851]" italics="true" pageId="17" pageNumber="598">ripa</emphasis>
, meaning
<emphasis id="B2839F354D6D726C63CBA52C4296FC83" box="[197,248,860,883]" italics="true" pageId="17" pageNumber="598">river</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,280 @@
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<mods:date id="C0FDBFAE72D4CB242B73F96BFD9CC6C2">2001</mods:date>
<mods:detail id="92E9A031603BBEC4D7B5BB7B50A09585" type="volume">
<mods:number id="478B06BB616E8EFE51765A940CEEC60F">35</mods:number>
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<mods:start id="2B9D9C665EFA79374FAC5FE89E303216">583</mods:start>
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<mods:classification id="7B6E9A50E3D050182F2303C5BFFD5D49">journal article</mods:classification>
<mods:identifier id="0827DEDB805789B534F446DDCF21E55A" type="ISSN">1464-5262</mods:identifier>
</mods:mods>
<treatment id="085EF2314D7872786254A5F943C7FCF4" ID-DOI="http://doi.org/10.5281/zenodo.5278962" ID-Zenodo-Dep="5278962" LSID="urn:lsid:plazi:treatment:085EF2314D7872786254A5F943C7FCF4" httpUri="http://treatment.plazi.org/id/085EF2314D7872786254A5F943C7FCF4" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585">
<subSubSection id="C8ED10AC4D7872796254A5F94179FC52" box="[346,791,905,930]" pageId="4" pageNumber="585" type="nomenclature">
<paragraph id="804843274D7872796254A5F94179FC52" blockId="4.[346,791,905,930]" box="[346,791,905,930]" pageId="4" pageNumber="585">
<heading id="DB00F44B4D7872796254A5F94179FC52" box="[346,791,905,930]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="585" reason="3">
<taxonomicName id="47F738A44D7872796254A5F940A4FC51" ID-CoL="8BZ42" authorityName="Collin" authorityYear="1930" box="[346,714,905,929]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D7872796254A5F940A4FC51" bold="true" box="[346,714,905,929]" pageId="4" pageNumber="585">
<emphasis id="B2839F354D7872796254A5F943F1FC51" bold="true" box="[346,415,905,929]" italics="true" pageId="4" pageNumber="585">Empis</emphasis>
(
<emphasis id="B2839F354D78727962BFA5F9402CFC51" bold="true" box="[433,578,905,929]" italics="true" pageId="4" pageNumber="585">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D787279615BA5F940A4FC51" bold="true" box="[597,714,905,929]" italics="true" pageId="4" pageNumber="585">chrysocera</emphasis>
</emphasis>
</taxonomicName>
-group
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D7872786352A59A4361FD75" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585" type="diagnosis">
<paragraph id="804843274D7872796352A59A42A9FBF1" blockId="4.[92,1047,1002,1248]" box="[92,199,1002,1025]" pageId="4" pageNumber="585">
<emphasis id="B2839F354D7872796352A59A42A9FBF1" box="[92,199,1002,1025]" italics="true" pageId="4" pageNumber="585">Diagnosis</emphasis>
</paragraph>
<paragraph id="804843274D787279638AA27840FFFB10" blockId="4.[92,1047,1002,1248]" pageId="4" pageNumber="585">Species brown to blackish in general colour; ¯agellum styliform; proboscis about twice the head height, labium slender, strongly sclerotized, with distinct annulations; proepisternum bare; laterotergite with fan of strong black bristles; R41 5 branched at right-angle, veins M1 and M2 abbreviated; presence of pennate bristles on legs of females; male pregenital segments always strongly modi®ed. Male cerci made up of anterior almost bare lobe and posterior bristled lobe; epandrial lamellae connected anteriorly. Spermatheca unsclerotized (®gure 14).</paragraph>
<paragraph id="804843274D7872796352A36342BEFADA" blockId="4.[92,1047,1299,1673]" box="[92,208,1299,1322]" pageId="4" pageNumber="585">
<emphasis id="B2839F354D7872796352A36342BEFADA" box="[92,208,1299,1322]" italics="true" pageId="4" pageNumber="585">Discussion</emphasis>
</paragraph>
<paragraph id="804843274D787279638AA3424299F9D9" blockId="4.[92,1047,1299,1673]" pageId="4" pageNumber="585">
With the relatively short proboscis, rather short scutal bristles, the entirely brown legs, the transformed male pregenital segments and the characteristic structure of male genitalia, species of the
<taxonomicName id="47F738A44D78727962BDA3034012FA7A" authorityName="Collin" authorityYear="1930" box="[435,636,1394,1418]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D78727962BDA30343A2FA7A" box="[435,460,1395,1418]" italics="true" pageId="4" pageNumber="585">E.</emphasis>
(
<emphasis id="B2839F354D78727962D2A3024398FA7A" box="[476,502,1394,1418]" italics="true" pageId="4" pageNumber="585">C.</emphasis>
)
<emphasis id="B2839F354D7872796106A3024012FA7A" box="[520,636,1394,1418]" italics="true" pageId="4" pageNumber="585">chrysocera</emphasis>
</taxonomicName>
-group are very distinct from those belonging to the subgenus
<taxonomicName id="47F738A44D7872796294A3E3404DFA5A" baseAuthorityName="Daugeron" baseAuthorityYear="2000" box="[410,547,1427,1450]" class="Insecta" family="Empididae" genus="Disneyempis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="B2839F354D7872796294A3E3404DFA5A" box="[410,547,1427,1450]" italics="true" pageId="4" pageNumber="585">Disneyempis</emphasis>
</taxonomicName>
and the
<taxonomicName id="47F738A44D787279619DA3E34108FA5A" authorityName="Smith" authorityYear="1971" box="[659,870,1426,1450]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="namwamba" subGenus="Coptophlebia">
<emphasis id="B2839F354D787279619DA3E340C2FA5A" box="[659,684,1427,1450]" italics="true" pageId="4" pageNumber="585">E.</emphasis>
(
<emphasis id="B2839F354D78727961B2A3E240B8FA5A" box="[700,726,1426,1450]" italics="true" pageId="4" pageNumber="585">C.</emphasis>
)
<emphasis id="B2839F354D78727961E4A3E24108FA5A" box="[746,870,1426,1450]" italics="true" pageId="4" pageNumber="585">namwamba</emphasis>
</taxonomicName>
-group. On the other hand, they can be distinguished from the remaining Afrotropical
<taxonomicName id="47F738A44D7872796084A3C2419DFA3A" baseAuthorityName="Daugeron" baseAuthorityYear="2000" box="[906,1011,1458,1482]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="tribe" tribe="Empidini">Empidini</taxonomicName>
by the following combination of characters: proepisternum bare, labella as long as prementum, presence of distinct annulations on labium and spermatheca unsclerotized.
</paragraph>
<paragraph id="804843274D787278638AA04143B7FF5E" blockId="4.[92,1047,1299,1673]" lastBlockId="5.[92,1047,118,398]" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585">
Within the
<taxonomicName id="47F738A44D7872796205A04243B9F9B9" authorityName="Collin" authorityYear="1930" box="[267,471,1585,1609]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D7872796205A042434BF9B9" box="[267,293,1586,1609]" italics="true" pageId="4" pageNumber="585">E.</emphasis>
(
<emphasis id="B2839F354D787279623AA0414321F9B9" box="[308,335,1585,1609]" italics="true" pageId="4" pageNumber="585">C.</emphasis>
)
<emphasis id="B2839F354D787279626DA04143B9F9B9" box="[355,471,1585,1609]" italics="true" pageId="4" pageNumber="585">chrysocera</emphasis>
</taxonomicName>
-group, two complexes of species can be de®ned: the
<taxonomicName id="47F738A44D7872796387A0224367F999" authorityName="Smith" authorityYear="1967" box="[137,265,1617,1641]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D7872796387A02242CDF999" box="[137,163,1618,1641]" italics="true" pageId="4" pageNumber="585">E.</emphasis>
(
<emphasis id="B2839F354D78727963BFA02142A2F999" box="[177,204,1617,1641]" italics="true" pageId="4" pageNumber="585">C.</emphasis>
)
<emphasis id="B2839F354D78727963D0A0214367F999" box="[222,265,1617,1641]" italics="true" pageId="4" pageNumber="585">lyra</emphasis>
</taxonomicName>
- and
<taxonomicName id="47F738A44D787279625EA0224074F999" authorityName="Collin" authorityYear="1930" box="[336,538,1617,1641]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D787279625EA0224304F999" box="[336,362,1618,1641]" italics="true" pageId="4" pageNumber="585">E.</emphasis>
(
<emphasis id="B2839F354D7872796277A02143FDF999" box="[377,403,1617,1641]" italics="true" pageId="4" pageNumber="585">C.</emphasis>
)
<emphasis id="B2839F354D78727962A8A0214074F999" box="[422,538,1617,1641]" italics="true" pageId="4" pageNumber="585">chrysocera</emphasis>
</taxonomicName>
-complexes (see the cladistic analysis). They can be distinguished by the colour of the male wing, the number of notopleurals, the tint of prescutellar depression, the shape and structure of male pregenital segments (see the key to species).
</paragraph>
<paragraph id="804843274D797278638AA6C6401AFE7D" blockId="5.[92,1047,118,398]" pageId="5" pageNumber="586">
Nothing is known about mating and feeding behaviour of species of the
<taxonomicName id="47F738A44D79727860C2A6C742BEFF1E" authorityName="Collin" authorityYear="1930" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D79727860C2A6C74188FF3E" box="[972,998,183,206]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727860FAA6C64660FF3E" box="[1012,1038,182,206]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D7972786352A6A642BEFF1E" box="[92,208,214,238]" italics="true" pageId="5" pageNumber="586">chrysocera</emphasis>
</taxonomicName>
-group. However, according to their label, the
<typeStatus id="5F4CFD854D79727861EFA6A6412BFF1E" box="[737,837,214,238]" pageId="5" pageNumber="586" type="holotype">holotype</typeStatus>
of
<taxonomicName id="47F738A44D7972786065A6A7418AFF1E" authorityName="Smith" authorityYear="1971" box="[875,996,214,238]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="barbitos">
<emphasis id="B2839F354D7972786065A6A7418AFF1E" box="[875,996,214,238]" italics="true" pageId="5" pageNumber="586">E. barbitos</emphasis>
</taxonomicName>
and
<specimenCount id="96F188AE4D7972786352A68642B7FEFE" box="[92,217,246,270]" pageId="5" pageNumber="586" type="female">one female</specimenCount>
<typeStatus id="5F4CFD854D79727863EAA6864326FEFE" box="[228,328,246,270]" pageId="5" pageNumber="586" type="paratype">paratype</typeStatus>
of
<taxonomicName id="47F738A44D797278627BA687404EFEFE" authorityName="Smith" authorityYear="1971" box="[373,544,246,270]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="cuthbertsoni">
<emphasis id="B2839F354D797278627BA687404EFEFE" box="[373,544,246,270]" italics="true" pageId="5" pageNumber="586">E. cuthbertsoni</emphasis>
</taxonomicName>
have been collected at ¯owers of
<taxonomicName id="47F738A44D79727860BBA6864310FEDE" class="Magnoliopsida" family="Ranunculaceae" genus="Clematis" kingdom="Plantae" order="Ranunculales" pageId="5" pageNumber="586" phylum="Tracheophyta" rank="species" species="stanleyi">
<emphasis id="B2839F354D79727860BBA68642DEFEDE" italics="true" pageId="5" pageNumber="586">Clematis stanleyi</emphasis>
(Ranunculaceae)
</taxonomicName>
and
<taxonomicName id="47F738A44D79727862CFA76640A0FEDE" box="[449,718,278,302]" class="Magnoliopsida" family="Caprifoliaceae" genus="Scabiosa" kingdom="Plantae" order="Dipsacales" pageId="5" pageNumber="586" phylum="Tracheophyta" rank="genus">
<emphasis id="B2839F354D79727862CFA766404CFEDE" box="[449,546,278,302]" italics="true" pageId="5" pageNumber="586">Scabiosa</emphasis>
(Dipsacaceae)
</taxonomicName>
, respectively. On the other hand males have holoptic eyes with upper ommatidia enlarged. Consequently, as most of
<taxonomicName id="47F738A44D79727863B6A726434FFE9E" baseAuthorityName="Daugeron" baseAuthorityYear="2000" box="[184,289,342,366]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="tribe" tribe="Empidini">Empidini</taxonomicName>
, species of the
<taxonomicName id="47F738A44D79727862C4A72740E3FE9E" authorityName="Collin" authorityYear="1930" box="[458,653,342,366]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D79727862C4A727438DFE9E" box="[458,483,343,366]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727862E1A7264064FE9E" box="[495,522,342,366]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D7972786117A72640E3FE9E" box="[537,653,342,366]" italics="true" pageId="5" pageNumber="586">chrysocera</emphasis>
</taxonomicName>
-group are probably ¯ower visitors and form swarms in which mating takes place.
</paragraph>
<paragraph id="804843274D7972786352A7B2428EFE2A" blockId="5.[92,1047,450,569]" box="[92,224,450,474]" pageId="5" pageNumber="586">
<emphasis id="B2839F354D7972786352A7B2428EFE2A" box="[92,224,450,474]" italics="true" pageId="5" pageNumber="586">Distribution</emphasis>
</paragraph>
<paragraph id="804843274D797278638AA79240AAFDC9" blockId="5.[92,1047,450,569]" pageId="5" pageNumber="586">
The
<taxonomicName id="47F738A44D79727863B4A79343EDFE0A" authorityName="Collin" authorityYear="1930" box="[186,387,482,506]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D79727863B4A79342BDFE0A" box="[186,211,483,506]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727863EDA7924293FE0A" box="[227,253,482,506]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D7972786201A79243EDFE0A" box="[271,387,482,506]" italics="true" pageId="5" pageNumber="586">chrysocera</emphasis>
</taxonomicName>
-group appears widespread in Africa, from West to East and Southern Africa, but seems absent from
<collectingCountry id="F8E003B74D797278615FA471408FFDE9" box="[593,737,513,537]" name="South Africa" pageId="5" pageNumber="586">South Africa</collectingCountry>
(®gure 47):
<collectingCountry id="F8E003B74D7972786062A47141D6FDE9" box="[876,952,513,537]" name="Democratic Republic of the Congo" pageId="5" pageNumber="586">Congo</collectingCountry>
,
<collectingCountry id="F8E003B74D79727860CBA4724661FDEA" box="[965,1039,514,538]" name="Kenya" pageId="5" pageNumber="586">Kenya</collectingCountry>
,
<collectingCountry id="F8E003B74D7972786352A451429FFDC9" box="[92,241,545,569]" name="Mozambique" pageId="5" pageNumber="586">Mozambique</collectingCountry>
,
<collectingCountry id="F8E003B74D797278620CA4514338FDC9" box="[258,342,545,569]" name="Nigeria" pageId="5" pageNumber="586">Nigeria</collectingCountry>
,
<collectingCountry id="F8E003B74D7972786269A45143D2FDC9" box="[359,444,545,569]" name="Senegal" pageId="5" pageNumber="586">Senegal</collectingCountry>
,
<collectingCountry id="F8E003B74D79727862C3A451405DFDC9" box="[461,563,545,569]" name="Tanzania" pageId="5" pageNumber="586">Tanzania</collectingCountry>
,
<collectingCountry id="F8E003B74D797278614BA45140D3FDC9" box="[581,701,545,569]" name="Zimbabwe" pageId="5" pageNumber="586">Zimbabwe</collectingCountry>
.
</paragraph>
<paragraph id="804843274D7972786352A41D4361FD75" blockId="5.[92,1048,621,773]" box="[92,271,621,645]" pageId="5" pageNumber="586">
<emphasis id="B2839F354D7972786352A41D4361FD75" box="[92,271,621,645]" italics="true" pageId="5" pageNumber="586">Included species</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D797278638AA4FE43C7FCF4" pageId="5" pageNumber="586" type="reference_group">
<paragraph id="804843274D797278638AA4FE43C7FCF4" blockId="5.[92,1048,621,773]" pageId="5" pageNumber="586">
<taxonomicName id="47F738A44D797278638AA4FE431CFD55" ID-CoL="8BZ3S" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[132,370,653,677]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="brazzavillensis" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D797278638AA4FE42F3FD55" box="[132,157,654,677]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727863A4A4FD42ABFD55" box="[170,197,653,677]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D79727863DBA4FD431CFD55" box="[213,370,653,677]" italics="true" pageId="5" pageNumber="586">brazzavillensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D7972786272A4FE43D5FD55" box="[380,443,654,677]" pageId="5" pageNumber="586" rank="species">sp. n.</taxonomicNameLabel>
;
<taxonomicName id="47F738A44D79727862C7A4FE416DFD55" ID-CoL="8BZ3M" authority="Smith, 1971" authorityName="Smith" authorityYear="1971" box="[457,771,653,677]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="barbitos" subGenus="Coptophlebia">
<emphasis id="B2839F354D79727862C7A4FE438DFD55" box="[457,483,654,677]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727862E1A4FD4064FD55" box="[495,522,653,677]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D7972786115A4FD401DFD55" box="[539,627,653,677]" italics="true" pageId="5" pageNumber="586">barbitos</emphasis>
<bibRefCitation id="E4663ED64D7972786172A4FD416DFD55" author="SMITH, K. G. V." box="[636,771,653,677]" pageId="5" pageNumber="586" pagination="96" refId="ref14522" refString="SMITH, K. G. V., 1971, The sub-Saharan species of Rhamphomyia and Coptophlebia with abbreviated discal venation (Diptera: Empididae), Annals of the Natal Museum, 21, 77 ± 96." type="journal article" year="1971">Smith, 1971</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="47F738A44D797278601DA4FE41A6FD55" ID-CoL="8BZ5J" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[787,968,653,677]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="juxtaripa" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D797278601DA4FE4142FD55" box="[787,812,654,677]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D7972786034A4FD413AFD55" box="[826,852,653,677]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D797278606AA4FE41A6FD55" box="[868,968,654,677]" italics="true" pageId="5" pageNumber="586">juxtaripa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D79727860DCA4FE467EFD55" box="[978,1040,654,677]" pageId="5" pageNumber="586" rank="species">sp. n.</taxonomicNameLabel>
;
<taxonomicName id="47F738A44D7972786352A4DE4317FD35" ID-CoL="8BZ67" authority="Smith, 1967 b" authorityName="Smith" authorityYear="1967" box="[92,377,685,709]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D7972786352A4DE421BFD35" box="[92,117,686,709]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D797278638DA4DD42F3FD35" box="[131,157,685,709]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D79727863A1A4DD42B4FD35" box="[175,218,685,709]" italics="true" pageId="5" pageNumber="586">lyra</emphasis>
<bibRefCitation id="E4663ED64D79727863EAA4DD4317FD35" author="SMITH, K. G. V." box="[228,377,685,709]" pageId="5" pageNumber="586" pagination="16" refId="ref14452" refString="SMITH, K. G. V., 1967 b, Afrikanische Empididae (Dipt.), Stuttgarter BeitraEge zur Naturkunde aus dem Staatlichen Museum fuEr Naturkunde in Stuttgart, 179, 1 ± 16." type="journal article" year="1967">Smith, 1967b</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="47F738A44D7972786285A4DE4059FD35" ID-CoL="8BZ7K" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[395,567,685,709]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="plumata" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D7972786285A4DE43CBFD35" box="[395,421,686,709]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727862BDA4DD43A3FD35" box="[435,461,685,709]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D79727862D1A4DD4059FD35" box="[479,567,685,709]" italics="true" pageId="5" pageNumber="586">plumata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D797278614FA4DE40EFFD35" box="[577,641,686,709]" pageId="5" pageNumber="586" rank="species">sp. n.</taxonomicNameLabel>
;
<taxonomicName id="47F738A44D797278619EA4DE4182FD35" ID-CoL="8BZ42" authority="Collin, 1929" authorityName="Collin" authorityYear="1930" box="[656,1004,685,709]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D797278619EA4DE40C4FD35" box="[656,682,686,709]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727861B6A4DD40BCFD35" box="[696,722,685,709]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D79727861EAA4DD4136FD35" box="[740,856,685,709]" italics="true" pageId="5" pageNumber="586">chrysocera</emphasis>
Collin, 1929
</taxonomicName>
;
<taxonomicName id="47F738A44D79727860F0A4DE43F0FD15" ID-CoL="8BZ4D" authority="Smith, 1971" authorityName="Smith" authorityYear="1971" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="cuthbertsoni" subGenus="Coptophlebia">
<emphasis id="B2839F354D79727860F0A4DE4676FD35" box="[1022,1048,686,709]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D797278636FA4BD4212FD15" box="[97,124,717,741]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D7972786382A4BD437FFD15" box="[140,273,717,741]" italics="true" pageId="5" pageNumber="586">cuthbertsoni</emphasis>
<bibRefCitation id="E4663ED64D7972786217A4BD43F0FD15" author="SMITH, K. G. V." box="[281,414,717,741]" pageId="5" pageNumber="586" pagination="96" refId="ref14522" refString="SMITH, K. G. V., 1971, The sub-Saharan species of Rhamphomyia and Coptophlebia with abbreviated discal venation (Diptera: Empididae), Annals of the Natal Museum, 21, 77 ± 96." type="journal article" year="1971">Smith, 1971</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="47F738A44D79727862A3A4BE415DFD15" ID-CoL="8BZ68" authority="Smith, 1969" authorityName="Smith" authorityYear="1969" box="[429,819,717,741]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="machipandensis" subGenus="Coptophlebia">
<emphasis id="B2839F354D79727862A3A4BE43A8FD15" box="[429,454,718,741]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727862DDA4BD4383FD15" box="[467,493,717,741]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D79727862F2A4BD40CBFD15" box="[508,677,717,741]" italics="true" pageId="5" pageNumber="586">machipandensis</emphasis>
<bibRefCitation id="E4663ED64D79727861A0A4BD415DFD15" author="SMITH, K. G. V." box="[686,819,717,741]" pageId="5" pageNumber="586" pagination="342" refId="ref14490" refString="SMITH, K. G. V., 1969, The Empididae of Southern Africa (Diptera), Annals of the Natal Museum, 19, 1 ± 342." type="journal article" year="1969">Smith, 1969</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="47F738A44D797278604DA4BE4678FD15" ID-CoL="8BZ7X" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[835,1046,717,741]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="samaruensis" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D797278604DA4BE4132FD15" box="[835,860,718,741]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D7972786066A4BD41ECFD15" box="[872,898,717,741]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D797278609CA4BE4678FD15" box="[914,1046,718,741]" italics="true" pageId="5" pageNumber="586">samaruensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D7972786352A49D42F5FCF4" box="[92,155,749,772]" pageId="5" pageNumber="586" rank="species">sp. n.</taxonomicNameLabel>
;
<taxonomicName id="47F738A44D79727863A5A49E430EFCF5" ID-CoL="8BZ82" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[171,352,749,773]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="singulare" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D79727863A5A49E42ABFCF5" box="[171,197,750,773]" italics="true" pageId="5" pageNumber="586">E.</emphasis>
(
<emphasis id="B2839F354D79727863DCA49D4282FCF5" box="[210,236,749,773]" italics="true" pageId="5" pageNumber="586">C.</emphasis>
)
<emphasis id="B2839F354D79727863F0A49D430EFCF5" box="[254,352,749,773]" italics="true" pageId="5" pageNumber="586">singulare</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D7972786264A49D43C7FCF4" box="[362,425,749,772]" pageId="5" pageNumber="586" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,165 @@
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<mods:title id="45675C93D2B2B7A2ADF56CD9E5E7B9EF">Cladistics and taxonomy of the Afrotropical Empis (Coptophlebia) chrysocera-group (Diptera, Empididae)</mods:title>
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<mods:date id="1DC06CE88F1E566643294C5CA998AFEE">2001</mods:date>
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<heading id="DB00F44B4D7972786248A5494145FCA1" box="[326,811,825,849]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="586" reason="3">
<taxonomicName id="47F738A44D7972786248A5494145FCA1" ID-CoL="8BZ3M" authority="Smith, 1971" authorityName="Smith" authorityYear="1971" box="[326,811,825,849]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="barbitos" subGenus="Coptophlebia">
<emphasis id="B2839F354D7972786248A54940F7FCA1" bold="true" box="[326,665,825,849]" pageId="5" pageNumber="586">
<emphasis id="B2839F354D7972786248A54943E5FCA1" bold="true" box="[326,395,825,849]" italics="true" pageId="5" pageNumber="586">Empis</emphasis>
(
<emphasis id="B2839F354D7972786293A5494040FCA1" bold="true" box="[413,558,825,849]" italics="true" pageId="5" pageNumber="586">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D797278614FA54940F7FCA1" bold="true" box="[577,665,825,849]" italics="true" pageId="5" pageNumber="586">barbitos</emphasis>
</emphasis>
<bibRefCitation id="E4663ED64D79727861AAA5494145FCA1" author="SMITH, K. G. V." box="[676,811,825,849]" pageId="5" pageNumber="586" pagination="96" refId="ref14522" refString="SMITH, K. G. V., 1971, The sub-Saharan species of Rhamphomyia and Coptophlebia with abbreviated discal venation (Diptera: Empididae), Annals of the Natal Museum, 21, 77 ± 96." type="journal article" year="1971">Smith, 1971</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="804843274D79727862FCA52940EFFC81" blockId="5.[92,811,825,913]" box="[498,641,857,882]" pageId="5" pageNumber="586">(®gures 1±3)</paragraph>
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<treatmentCitationGroup id="A0E764094D7972786352A50C4054FC61" box="[92,570,891,913]" pageId="5" pageNumber="586">
<treatmentCitation id="015665364D7972786352A50C405AFC61" author="SMITH, K. G. V." box="[92,564,891,913]" page="90" pageId="5" pageNumber="586" year="1971">
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<emphasis id="B2839F354D7972786352A50C42F7FC61" box="[92,153,892,913]" italics="true" pageId="5" pageNumber="586">Empis</emphasis>
(
<emphasis id="B2839F354D79727863A8A50B4347FC61" box="[166,297,891,913]" italics="true" pageId="5" pageNumber="586">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D7972786235A50B43E5FC61" box="[315,395,891,913]" italics="true" pageId="5" pageNumber="586">barbitos</emphasis>
<bibRefCitation id="E4663ED64D797278629DA50B405AFC61" author="SMITH, K. G. V." box="[403,564,891,913]" pageId="5" pageNumber="586" pagination="96" refId="ref14522" refString="SMITH, K. G. V., 1971, The sub-Saharan species of Rhamphomyia and Coptophlebia with abbreviated discal venation (Diptera: Empididae), Annals of the Natal Museum, 21, 77 ± 96." type="journal article" year="1971">Smith, 1971: 90</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<paragraph id="804843274D7972786352A04F4054F979" blockId="5.[92,1047,1599,1674]" pageId="5" pageNumber="586">
FIGS 13.
<taxonomicName id="47F738A44D79727863DCA04F403EF9A5" ID-CoL="8BZ3M" authority="Smith" authorityName="Smith" authorityYear="1971" box="[210,592,1599,1621]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="barbitos" subGenus="Coptophlebia">
<emphasis id="B2839F354D79727863DCA04F4361F9A4" box="[210,271,1599,1620]" italics="true" pageId="5" pageNumber="586">Empis</emphasis>
(
<emphasis id="B2839F354D797278622EA04F43CDF9A5" box="[288,419,1599,1621]" italics="true" pageId="5" pageNumber="586">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D79727862B9A04F4069F9A5" box="[439,519,1599,1621]" italics="true" pageId="5" pageNumber="586">barbitos</emphasis>
Smith
</taxonomicName>
: (1) male posterior lobe of cercus, lateral view (scale 0.1 mm); (2) male anterior lobe of cercus, lateral view (scale 0.1 mm); (3) phallus, lateral view (scale 0.1 mm).
</paragraph>
</caption>
<paragraph id="804843274D7A727B621AA6464296FF9E" blockId="6.[92,1046,54,110]" pageId="6" pageNumber="587">
Afrotropical
<taxonomicName id="47F738A44D7A727B62A9A647417FFFBE" ID-CoL="8BZ42" authorityName="Collin" authorityYear="1930" box="[423,785,54,78]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="chrysocera" subGenus="Coptophlebia">
<emphasis id="B2839F354D7A727B62A9A6474385FFBE" box="[423,491,55,78]" italics="true" pageId="6" pageNumber="587">Empis</emphasis>
(
<emphasis id="B2839F354D7A727B62F7A64640E4FFBE" box="[505,650,54,78]" italics="true" pageId="6" pageNumber="587">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D7A727B6192A646417FFFBE" box="[668,785,54,78]" italics="true" pageId="6" pageNumber="587">chrysocera</emphasis>
</taxonomicName>
group
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587
<emphasis id="B2839F354D7A727B6352A62742FAFF9E" box="[92,148,87,110]" italics="true" pageId="6" pageNumber="587">Type</emphasis>
</specimenCount>
material
</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D7A727A638AA60843A5FF5E" lastPageId="7" lastPageNumber="588" pageId="6" pageNumber="587" type="materials_examined">
<paragraph id="804843274D7A727B638AA60840C3FF40" blockId="6.[92,1047,119,176]" pageId="6" pageNumber="587">
<materialsCitation id="309F497A4D7A727B638AA60840C3FF40" collectingDate="1939-01-15" collectionCode="NHM" collectorName="A. Cuthbertson" country="Zimbabwe" location="Cleveland Dam" municipality="Salisbury" pageId="6" pageNumber="587" specimenCount="1" typeStatus="holotype">
<typeStatus id="5F4CFD854D7A727B638AA6084292FF7F" box="[132,252,119,143]" pageId="6" pageNumber="587" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F8E003B74D7A727B623AA60743C2FF7F" box="[308,428,119,143]" name="Zimbabwe" pageId="6" pageNumber="587">Zimbabwe</collectingCountry>
,
<location id="852815FC4D7A727B62CCA6074017FF7F" LSID="urn:lsid:plazi:treatment:085EF2314D79727A6248A54943A5FF5E:852815FC4D7A727B62CCA6074017FF7F" box="[450,633,119,143]" country="Zimbabwe" municipality="Salisbury" name="Cleveland Dam" pageId="6" pageNumber="587">Cleveland Dam</location>
,
<collectingMunicipality id="602CD95D4D7A727B6181A6074099FF7F" box="[655,759,119,143]" pageId="6" pageNumber="587">Salisbury</collectingMunicipality>
, at ¯owers of
<taxonomicName id="47F738A44D7A727B60BBA60742DEFF40" class="Magnoliopsida" family="Ranunculaceae" genus="Clematis" kingdom="Plantae" order="Ranunculales" pageId="6" pageNumber="587" phylum="Tracheophyta" rank="species" species="stanleyi">
<emphasis id="B2839F354D7A727B60BBA60742DEFF40" italics="true" pageId="6" pageNumber="587">Clematis stanleyi</emphasis>
</taxonomicName>
,
<date id="F44965E74D7A727B63CFA6E84312FF40" box="[193,380,152,176]" pageId="6" pageNumber="587" value="1939-01-15">
<collectingDate id="E40D9C0F4D7A727B63CFA6E84312FF40" box="[193,380,152,176]" pageId="6" pageNumber="587" value="1939-01-15">15 January 1939</collectingDate>
</date>
,
<collectorName id="2D0226F14D7A727B6283A6E84051FF40" box="[397,575,152,176]" pageId="6" pageNumber="587">A. Cuthbertson</collectorName>
(
<collectionCode id="E6E6DBE24D7A727B615DA6E840F6FF5F" box="[595,664,152,175]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="6" pageNumber="587" type="Herbarium">NHM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="804843274D7A727B6352A68C42B4FEE3" blockId="6.[92,1048,252,1476]" box="[92,218,252,275]" pageId="6" pageNumber="587">
<emphasis id="B2839F354D7A727B6352A68C42B4FEE3" box="[92,218,252,275]" italics="true" pageId="6" pageNumber="587">Description</emphasis>
</paragraph>
<paragraph id="804843274D7A727B638AA76C43BBFEC3" blockId="6.[92,1048,252,1476]" box="[132,469,284,308]" pageId="6" pageNumber="587">Male: length of wing 4.8 mm.</paragraph>
<paragraph id="804843274D7A727B638AA74C40B1FE25" blockId="6.[92,1048,252,1476]" pageId="6" pageNumber="587">Head with occiput blackish to dusted greyish. Ocellar triangle with pair of very short bristles. Face dusted greyish in upper part, shining black in lower part. Antenna blackish. Palpus yellowish. Proboscis long (2.5 times the head height), labrum brownish, labium blackish, strongly sclerotized, with distinct annulations, labella slender, bare. Eyes holoptic, upper ommatidia enlarged.</paragraph>
<paragraph id="804843274D7A727B638AA7AD41EBFD28" blockId="6.[92,1048,252,1476]" pageId="6" pageNumber="587">Thorax blackish to dusted greyish. Antepronotum with a few distinct lateral bristles. Postpronotal lobes subshining dark brown with one distinct basal bristle. Proepisternum bare. Prosternum with a few lateral short bristles. Acrostichals uniserial, ®ne, short. Dorsocentrals uni- to biserial, rather ®ne, short, with one strong, long prescutellar bristle. One strong, long postsutural supraalar. Three strong, long notopleurals. Postalar calli brownish with one strong, long bristle. Scutellum with two strong, long apical bristles, two ®ner, shorter subapicals. Laterotergite with fan of black bristles. Anterior spiracle black, posterior spiracle not visible.</paragraph>
<paragraph id="804843274D7A727B638AA49243B2FBED" blockId="6.[92,1048,252,1476]" pageId="6" pageNumber="587">Legs brownish, coxa dusted greyish. Fore femur with short dorsal bristles, ®ne, longer ventrals. Fore tibia with one row of distinct dorsal bristles. First tarsomere of fore tarsus with strong, short ventral bristles. Mid femur with one row of strong dorsal bristles, pennate posteroventrally on apical half. Mid tibia pennate dorsally except at tips, with distinct ventral bristles. First tarsomere of mid tarsus with strong, not very long bristles, especially ventrally. Hind femur with numerous distinct dorsal and ventral bristles, stronger dorsally, pennate posteroventrally at apical tip. Hind tibia with numerous distinct dorsal and ventral bristles, stronger dorsally, pennate dorsally in the middle. Hind tarsus similar to mid tarsus. Each tarsomere of all tarsi with distinct bristles at apical tip.</paragraph>
<paragraph id="804843274D7A727B638AA2554067FB6F" blockId="6.[92,1048,252,1476]" pageId="6" pageNumber="587">
Wing dark brown. Sc abbreviated.
<collectionCode id="E6E6DBE24D7A727B6128A2554057FBCD" box="[550,569,1061,1085]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="587">R</collectionCode>
41 5 branched at right-angle. Discal cell truncate. M1 abbreviated. M2 abbreviated in the right wing, complete in the left wing. A1 complete. Anal lobe well developed, anal angle rather acute. Halter with brownish base and stem, black knob.
</paragraph>
<paragraph id="804843274D7A727B638AA2D84328FAB1" blockId="6.[92,1048,252,1476]" pageId="6" pageNumber="587">Abdomen very damaged. Brownish at base with several distinct bristles on the ®rst two segments, otherwise silvery dusted (according to the original description). Tergite 7 apparently divided in two lateral parts and a less sclerotized central one. According to the original drawing of postabdomen, sternite 8 developed posteriorly with rounded shape.</paragraph>
<paragraph id="804843274D7A727B638AA33A408DFA54" blockId="6.[92,1048,252,1476]" pageId="6" pageNumber="587">Hypopygium (®gures 1±3) very damaged. Cerci broken, apparently made up of anterior reduced rounded lobe (®gure 2) and well-developed posterior lobe bearing distinct bristles (®gure 1). Phallus short, thick (®gure 3).</paragraph>
<paragraph id="804843274D7A727B638AA3DC4336FA34" blockId="6.[92,1048,252,1476]" box="[132,344,1452,1476]" pageId="6" pageNumber="587">Female: unknown.</paragraph>
<paragraph id="804843274D7A727B6352A06042BEF9D7" blockId="6.[92,1047,1552,1673]" box="[92,208,1552,1575]" pageId="6" pageNumber="587">
<emphasis id="B2839F354D7A727B6352A06042BEF9D7" box="[92,208,1552,1575]" italics="true" pageId="6" pageNumber="587">Discussion</emphasis>
</paragraph>
<paragraph id="804843274D7A727B638AA0414134F979" blockId="6.[92,1047,1552,1673]" pageId="6" pageNumber="587">
<taxonomicName id="47F738A44D7A727B638AA0414293F9B8" authorityName="Smith" authorityYear="1971" box="[132,253,1584,1608]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="barbitos">
<emphasis id="B2839F354D7A727B638AA0414293F9B8" box="[132,253,1584,1608]" italics="true" pageId="6" pageNumber="587">E. barbitos</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="47F738A44D7A727B62BAA041405CF9B8" authorityName="Smith" authorityYear="1967" box="[436,562,1584,1608]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D7A727B62BAA04143A3F9B8" box="[436,461,1585,1608]" italics="true" pageId="6" pageNumber="587">E.</emphasis>
(
<emphasis id="B2839F354D7A727B62D2A0404398F9B8" box="[476,502,1584,1608]" italics="true" pageId="6" pageNumber="587">C.</emphasis>
)
<emphasis id="B2839F354D7A727B6109A040405CF9B8" box="[519,562,1584,1608]" italics="true" pageId="6" pageNumber="587">lyra</emphasis>
</taxonomicName>
-complex. It resembles
<taxonomicName id="47F738A44D7A727B6033A04141E4F9B8" authorityName="Smith" authorityYear="1967" box="[829,906,1584,1608]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="lyra">
<emphasis id="B2839F354D7A727B6033A04141E4F9B8" box="[829,906,1584,1608]" italics="true" pageId="6" pageNumber="587">E. lyra</emphasis>
</taxonomicName>
in the basic pattern of dusting on the scutum, but the lyre-shaped stripe under the dorsocentrals is not present and the antennae are blackish, not brown-yellowish.
</paragraph>
<paragraph id="804843274D7B727A6352A60643A5FF5E" blockId="7.[92,459,118,174]" box="[92,459,118,174]" pageId="7" pageNumber="588">
<emphasis id="B2839F354D7B727A6352A606428EFF7E" box="[92,224,118,142]" italics="true" pageId="7" pageNumber="588">Distribution</emphasis>
Southern Africa:
<collectingCountry id="F8E003B74D7B727A6242A6E643AAFF5E" box="[332,452,150,174]" name="Zimbabwe" pageId="7" pageNumber="588">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,146 @@
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<mods:title id="295B24C4B2F27F3A7239936A5A0229A6">Cladistics and taxonomy of the Afrotropical Empis (Coptophlebia) chrysocera-group (Diptera, Empididae)</mods:title>
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<mods:namePart id="B01B6C5EFBBF45C61F98C812E4FB60A5">Esa, C. Daugeron</mods:namePart>
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<mods:part id="BE63F543A18A2BB9F599D25F0E5EC24C">
<mods:date id="C8764892AAE0CF3AE8DBABFB9666E55E">2001</mods:date>
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<treatment id="085EF2314D7B72746246A6944176FBDB" ID-DOI="http://doi.org/10.5281/zenodo.5278964" ID-Zenodo-Dep="5278964" LSID="urn:lsid:plazi:treatment:085EF2314D7B72746246A6944176FBDB" httpUri="http://treatment.plazi.org/id/085EF2314D7B72746246A6944176FBDB" lastPageId="9" lastPageNumber="590" pageId="7" pageNumber="588">
<subSubSection id="C8ED10AC4D7B727A6246A69440EFFEEC" pageId="7" pageNumber="588" type="nomenclature">
<paragraph id="804843274D7B727A6246A6944147FF0C" blockId="7.[328,809,228,285]" box="[328,809,228,252]" pageId="7" pageNumber="588">
<heading id="DB00F44B4D7B727A6246A6944147FF0C" box="[328,809,228,252]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="588" reason="3">
<taxonomicName id="47F738A44D7B727A6246A694408EFF0C" ID-CoL="8BZ3S" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[328,736,228,252]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="species" species="brazzavillensis" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D7B727A6246A694408EFF0C" bold="true" box="[328,736,228,252]" pageId="7" pageNumber="588">
<emphasis id="B2839F354D7B727A6246A69443E3FF0C" bold="true" box="[328,397,228,252]" italics="true" pageId="7" pageNumber="588">Empis</emphasis>
(
<emphasis id="B2839F354D7B727A6291A694405EFF0C" bold="true" box="[415,560,228,252]" italics="true" pageId="7" pageNumber="588">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D7B727A614DA694408EFF0C" bold="true" box="[579,736,228,252]" italics="true" pageId="7" pageNumber="588">brazzavillensis</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D7B727A61E5A6954147FF0C" box="[747,809,229,252]" pageId="7" pageNumber="588" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="804843274D7B727A62FCA77440EFFEEC" blockId="7.[328,809,228,285]" box="[498,641,260,285]" pageId="7" pageNumber="588">(®gures 4±9)</paragraph>
</subSubSection>
<subSubSection id="C8ED10AC4D7B72746352A7194176FBDB" lastPageId="9" lastPageNumber="590" pageId="7" pageNumber="588" type="materials_examined">
<paragraph id="804843274D7B727A6352A7194296FE70" blockId="7.[92,1047,360,448]" box="[92,248,360,384]" pageId="7" pageNumber="588">
<emphasis id="B2839F354D7B727A6352A7194296FE70" box="[92,248,360,384]" italics="true" pageId="7" pageNumber="588">Type material</emphasis>
</paragraph>
<paragraph id="804843274D7B727A638AA7F8406FFE30" blockId="7.[92,1047,360,448]" pageId="7" pageNumber="588">
<materialsCitation id="309F497A4D7B727A638AA7F8467EFE50" box="[132,1040,392,416]" collectorName="de Brazzaville &amp; E. Roubaud &amp; A. Weiss" country="Democratic Republic of the Congo" location="Congo" pageId="7" pageNumber="588" specimenCount="1" typeStatus="holotype">
<typeStatus id="5F4CFD854D7B727A638AA7F84292FE50" box="[132,252,392,416]" pageId="7" pageNumber="588" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F8E003B74D7B727A623EA7F84312FE50" box="[304,380,392,416]" name="Democratic Republic of the Congo" pageId="7" pageNumber="588">Congo</collectingCountry>
, environ
<collectorName id="2D0226F14D7B727A62FFA7F840FDFE50" box="[497,659,392,416]" pageId="7" pageNumber="588">de Brazzaville</collectorName>
, 1907,
<collectorName id="2D0226F14D7B727A61E3A7F84113FE50" box="[749,893,392,416]" pageId="7" pageNumber="588">E. Roubaud</collectorName>
&amp;
<collectorName id="2D0226F14D7B727A60A6A7F8467EFE50" box="[936,1040,392,416]" pageId="7" pageNumber="588">A. Weiss</collectorName>
</materialsCitation>
;
<materialsCitation id="309F497A4D7B727A6352A7D84392FE30" box="[92,508,424,448]" collectionCode="MNHN" collectorName="de Brazzaville &amp; E. Roubaud &amp; A. Weiss" country="Democratic Republic of the Congo" elevation="1" location="Congo" pageId="7" pageNumber="588" specimenCount="1" typeStatus="paratype">
<typeStatus id="5F4CFD854D7B727A6352A7D842A7FE30" box="[92,201,424,448]" pageId="7" pageNumber="588" type="paratype">PARATYPE</typeStatus>
:
<quantity id="470FEEC24D7B727A63DAA7D84297FE4F" box="[212,249,424,448]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="588" unit="m" value="1.0">
<elevation id="0BDAA4144D7B727A63DAA7D84297FE4F" box="[212,249,424,448]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="588" unit="m" value="1.0">1 m</elevation>
</quantity>
, same data (
<collectionCode id="E6E6DBE24D7B727A629FA7D84383FE4F" box="[401,493,424,447]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="588">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="804843274D7B727A6352A78642B4FDFD" blockId="7.[92,1047,502,1673]" box="[92,218,502,525]" pageId="7" pageNumber="588">
<emphasis id="B2839F354D7B727A6352A78642B4FDFD" box="[92,218,502,525]" italics="true" pageId="7" pageNumber="588">Description</emphasis>
</paragraph>
<paragraph id="804843274D7B727A638AA46543BBFDDD" blockId="7.[92,1047,502,1673]" box="[132,469,533,557]" pageId="7" pageNumber="588">Male: length of wing 4.7 mm.</paragraph>
<paragraph id="804843274D7B727A638AA44540D8FD1C" blockId="7.[92,1047,502,1673]" pageId="7" pageNumber="588">Head with occiput brown to dusted greyish. Ocellar triangle with pair of short bristles. Face dusted greyish in upper part, shining black in lower part. Scape and pedicel brown, ®rst ¯agellomere yellowish to brown at base and at tip, second and third ¯agellomeres dark brown. Proboscis long (twice the head height), labium blackish, strongly sclerotized, with distinct annulations, labella slender, bare, palpus brownish. Eyes holoptic, upper ommatidia enlarged.</paragraph>
<paragraph id="804843274D7B727A638AA4844181FBDB" blockId="7.[92,1047,502,1673]" pageId="7" pageNumber="588">
Thorax dark brown. Antepronotu m with short lateral bristles. Postpronota l lobes with one strong, long basal bristle. Proepisternum bare. Prosternum with a few short lateral bristles. Scutum with a more or less distinct dusted grey stripe between acrostichals and dorsocentrals, and lyre-shaped markings on dorsocentrals. Acrostichals apparently uniserial, very ®ne, short. Dorsocentrals apparently uniserial, ®ne, short although more distinct than acrostichals, not visible in the prescutellar depression. The
<typeStatus id="5F4CFD854D7B727A6252A5C443AEFC3C" box="[348,448,948,972]" pageId="7" pageNumber="588" type="holotype">holotype</typeStatus>
with distinct insertion of one postsutural supraalar. Three strong, long notopleurals. One strong, long postalar. Scutellum with two strong, long apical bristles, the
<typeStatus id="5F4CFD854D7B727A62CCA5834048FBFB" box="[450,550,1011,1035]" pageId="7" pageNumber="588" type="holotype">holotype</typeStatus>
with, between them, one ®ner, shorter one. Laterotergite with fan of black bristles. Anterior and posterior spiracles brown.
</paragraph>
<paragraph id="804843274D7B727A638AA244466DFABA" blockId="7.[92,1047,502,1673]" pageId="7" pageNumber="588">Legs brown. Fore femur with short dorsal bristles, ®ner, longer ventrals. Fore tibia with one row of strong, long dorsals. First tarsomere of fore tarsus with a few strong, short ventrals. Mid femur pennate ventrally on apical half, with one row of dorsal distinct bristles. Mid tibia pennate dorsally except at basal tip, with a few pennate ventral bristles at base. First tarsomere of mid tarsus with numerous strong, rather long bristles. Hind femur pennate antero- and posteroventrally at apical tip, with one row of ®ne, rather long ventral bristles. Hind tibia pennate dorsally except at basal tip. First tarsomere of hind tarsus with numerous distinct, rather long dorsal and ventral bristles. Each tarsomere of all tarsi with distinct bristles at apical tip.</paragraph>
<paragraph id="804843274D7B727A638AA3224144FA5A" blockId="7.[92,1047,502,1673]" pageId="7" pageNumber="588">
Wing dark brown. Sc abbreviated.
<collectionCode id="E6E6DBE24D7B727A6128A3224057FA9A" box="[550,569,1362,1386]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="588">R</collectionCode>
41 5 branched at right-angle. Discal cell truncate. M1, M2 abbreviated. A1 complete. Anal lobe well developed, anal angle acute. Halter with yellowish base and stem, dark brown knob.
</paragraph>
<paragraph id="804843274D7B727A638AA3C24662F9B9" blockId="7.[92,1047,502,1673]" pageId="7" pageNumber="588">Abdomen (®gures 4±6) dusted brownish. First three segments with rather strong, long, brownish lateral bristles. Tergite 6 with rounded medioposterior expansion (®gure 6). Tergite 7 indistinctly divided in three parts (®gure 5). Tergite 8 divided in three sclerites, a central one expanded anteroventrally, and two lateral wing-shaped ones (®gures 4, 5). Sternite 8 developed posteriorly with rounded shape (®gure 4).</paragraph>
<paragraph id="804843274D7B7274638AA02141E0FF5E" blockId="7.[92,1047,502,1673]" lastBlockId="9.[92,1047,118,589]" lastPageId="9" lastPageNumber="590" pageId="7" pageNumber="588">Hypopygium (®gures 7, 8) with cercus made up of almost bare anterior lobe, rounded dorsally, and posterior pointed lobe with numerous distinct bristles (®gure 7). Epandrial lamellae connected anteriorly, with posterodorsal rounded expansion bearing a few distinct bristles at tip (®gure 8). Phallus short.</paragraph>
<caption id="D48813AF4D7472756352A3EF438EF9C9" pageId="8" pageNumber="589" startId="8.[92,105,1439,1460]" targetBox="[104,1035,119,1410]" targetPageId="8">
<paragraph id="804843274D7472756352A3EF438EF9C9" blockId="8.[92,1047,1439,1593]" pageId="8" pageNumber="589">
F IGS 49.
<taxonomicName id="47F738A44D74727563DFA3EF4053FA45" authority="Esa, 2001" authorityName="Esa" authorityYear="2001" box="[209,573,1439,1461]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="589" phylum="Arthropoda" rank="species" species="brazzavillensis" status="sp. nov." subGenus="Coptophlebia">
<emphasis id="B2839F354D74727563DFA3EF4360FA44" box="[209,270,1439,1460]" italics="true" pageId="8" pageNumber="589">Empis</emphasis>
(
<emphasis id="B2839F354D7472756212A3EF43F1FA45" box="[284,415,1439,1461]" italics="true" pageId="8" pageNumber="589">Coptophlebia</emphasis>
)
<emphasis id="B2839F354D74727562A1A3EF4053FA45" box="[431,573,1439,1461]" italics="true" pageId="8" pageNumber="589">brazzavillensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9B0224E4D747275614BA3EF4013FA44" box="[581,637,1439,1460]" pageId="8" pageNumber="589" rank="species">sp. n.</taxonomicNameLabel>
: (4) male abdominal segments 7 and 8, lateral view (scale 0.1 mm); (5) male abdominal tergites 7 and 8, dorsal view (scale 0.1 mm); (6) male abdominal tergite 6, dorsal view (scale 0.1 mm); (7) male cercus, lateral view (scale 0.1 mm); (8) epandrial lamella, lateral view (scale 0.1 mm); (9) female tergite 7, dorsal view (scale 0.1 mm). st 7 5 sternite 7; st 8 5 sternite 8; tg 7 5 tergite 7; tg 8 5 tergite 8.
</paragraph>
</caption>
<paragraph id="804843274D757274638AA6C642F4FDBD" blockId="9.[92,1047,118,589]" pageId="9" pageNumber="590">Female similar to male except for the following characters: pedicel lighter, ¯agellum entirely yellowish. Proboscis a little shorter. Eyes dichoptic, ommatidia all of equal size. Frons shining blackish in upper part. Dorsocentrals with one strong, long prescutellar bristle. Fore femur short pennate dorsally on basal half, with two or three ventral pennate bristles at base. Fore tibia pennate dorsally except at basal tip, with ®ne bristles at apical one-quarter. Mid femur pennate dorsally (only for short distance apically) and posteroventrally. Mid tibia pennate dorsally except at basal tip, posteroventrally at basal third. Hind femur pennate dorsally except at apical tip, with ®ne, short pennate ventral bristles. Hind tibia pennate dorsally except at basal tip, ventrally especially at basal third. First tarsomere of all tarsi pennate dorsally. Abdomen brown to yellowish, almost bare. Tergite 7 with circular-shaped anterior margin (®gure 9). Cercus longer than broad, with ®ne, very short bristly hairs.</paragraph>
<paragraph id="804843274D7572746352A40542BEFD7C" blockId="9.[92,1047,629,876]" box="[92,208,629,652]" pageId="9" pageNumber="590">
<emphasis id="B2839F354D7572746352A40542BEFD7C" box="[92,208,629,652]" italics="true" pageId="9" pageNumber="590">Discussion</emphasis>
</paragraph>
<paragraph id="804843274D757274638AA4E54056FC9C" blockId="9.[92,1047,629,876]" pageId="9" pageNumber="590">
<taxonomicName id="47F738A44D757274638AA4E5432AFD5C" authorityName="Esa" authorityYear="2001" box="[132,324,660,684]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="590" phylum="Arthropoda" rank="species" species="brazzavillensis">
<emphasis id="B2839F354D757274638AA4E5432AFD5C" box="[132,324,660,684]" italics="true" pageId="9" pageNumber="590">E. brazzavillensis</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="47F738A44D75727462F6A4E5401BFD5C" authorityName="Smith" authorityYear="1967" box="[504,629,660,684]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="590" phylum="Arthropoda" rank="species" species="lyra" subGenus="Coptophlebia">
<emphasis id="B2839F354D75727462F6A4E5407FFD5C" box="[504,529,661,684]" italics="true" pageId="9" pageNumber="590">E.</emphasis>
(
<emphasis id="B2839F354D7572746111A4E44057FD5C" box="[543,569,660,684]" italics="true" pageId="9" pageNumber="590">C.</emphasis>
)
<emphasis id="B2839F354D7572746144A4E4401BFD5C" box="[586,629,660,684]" italics="true" pageId="9" pageNumber="590">lyra</emphasis>
</taxonomicName>
-complex. It resembles
<taxonomicName id="47F738A44D7572746071A4E541A2FD5C" authorityName="Smith" authorityYear="1967" box="[895,972,660,684]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="590" phylum="Arthropoda" rank="species" species="lyra">
<emphasis id="B2839F354D7572746071A4E541A2FD5C" box="[895,972,660,684]" italics="true" pageId="9" pageNumber="590">E. lyra</emphasis>
</taxonomicName>
in the male legs ornamentation, the yellowish antennae and the presence, although less distinct, of lyre-shaped markings on the scutum. Both species can be distinguished by the general colour of the body, extensively brownish for the ®rst, brown to dusted grey for the second, the shape of the male genital sclerite, especially the epandrial lamella, and the female legs ornamentation: fore tibia almost bare ventrally for the ®rst, with pennate bristles for the second.
</paragraph>
<paragraph id="804843274D7572746352A5E443E2FC3C" blockId="9.[92,396,916,972]" box="[92,396,916,972]" pageId="9" pageNumber="590">
<emphasis id="B2839F354D7572746352A5E4428EFC5C" box="[92,224,916,940]" italics="true" pageId="9" pageNumber="590">Distribution</emphasis>
Central Africa:
<collectingCountry id="F8E003B74D7572746237A5C443EBFC3C" box="[313,389,948,972]" name="Democratic Republic of the Congo" pageId="9" pageNumber="590">Congo</collectingCountry>
.
</paragraph>
<paragraph id="804843274D7572746352A5834176FBDB" blockId="9.[92,792,1011,1067]" box="[92,792,1011,1067]" pageId="9" pageNumber="590">
<emphasis id="B2839F354D7572746352A58342BDFBFB" box="[92,211,1011,1035]" italics="true" pageId="9" pageNumber="590">Etymology</emphasis>
The name of the species is derived from the type-locality.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="7D2513270DE728CD0B51CBBD1D74E33F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:230101" class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster gibbosa var. mauritania" order="Hymenoptera" pageNumber="284" phylum="Arthropoda" rank="variety" species="gibbosa" variety="mauritania">A. gibbosa Latr. v. mauritania Em.</taxonomicName>
</paragraph>
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<paragraph id="D9C09B9E0175FE7200D6B0FDF71100B2" pageNumber="284">- [[male]] (non decrit) long. 4,5-5 mill, - Noir, mandibules, antennes et pattes brunes, lisse et luisant, tete mate et ponctuee. Epinotum un peu plus, etrangle en avant que chez le type, du reste semblable.</paragraph>
</subSubSection>
<subSubSection id="28359EC6D738DDA2B2E11161EC4EB417" type="materials_examined">
<paragraph id="E68F4B924C50E9C9AB9C8DB5119631B3" pageNumber="284">Tunisie: Djbel Ouselet. Trouve dans le nid avec les [[worker]][[worker]], 1907.</paragraph>
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