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<document id="C8C596AB20644698EF61B0DCA39CE9A5" ID-CLB-Dataset="20614" ID-DOI="10.11646/zootaxa.5100.1.2" ID-GBIF-Dataset="9a754e9d-4435-47a0-b1d9-7a4d2b46b377" ID-ISSN="1175-5326" ID-Zenodo-Dep="6127749" ID-ZooBank="EB3B7AAB-C886-4B80-AC84-27E17BFE2EA8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645084568560" checkinUser="plazi" docAuthor="Kullander, Sven &amp; Norén, Michael" docDate="2022" docId="08571F4FFFE51B5AFF28D0B3FAAE70D3" docLanguage="en" docName="zootaxa.5100.1.2.pdf" docOrigin="Zootaxa 5100 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Devario ahlanderi Kullander &amp; Norén 2022, new species" docType="treatment" docVersion="7" lastPageNumber="64" masterDocId="F46E6737FFE21B50FFBFD530FFCA776A" masterDocTitle="The real Devario browni from the Irrawaddy River basin, and the new Devario ahlanderi from the Salween River basin in Myanmar (Teleostei: Cyprinidae: Danioninae)" masterLastPageNumber="72" masterPageNumber="54" pageNumber="61" updateTime="1699314175049" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FE9D9EA15513BCB495FE367663489BD8">The real Devario browni from the Irrawaddy River basin, and the new Devario ahlanderi from the Salween River basin in Myanmar (Teleostei: Cyprinidae: Danioninae)</mods:title>
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<mods:name id="9EFEA7AF4FA894DA76D2B450FA6AA58A" type="personal">
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<mods:namePart id="398A79200A9B382F17531E1DA1CAA2A0">Kullander, Sven</mods:namePart>
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<mods:affiliation id="331BE53D80BF8CDC344C5314A3135D04">Department of Zoology, Swedish Museum of Natural History, PO Box 50007, SE- 104 05 Stockholm, Sweden &amp; sven. kullander @ nrm. se; https: // orcid. org / 0000 - 0001 - 6075 - 0266</mods:affiliation>
<mods:nameIdentifier id="B04FFA90FA5DCC696D1C94DE82C6DCEC" type="email">sven.kullander@nrm.se</mods:nameIdentifier>
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<mods:namePart id="EEA89F56B7CD8BB659417F9CD6DC145B">Norén, Michael</mods:namePart>
<mods:nameIdentifier id="CA46049AE206B71D019B63D5DC8C0849" type="ORCID">0000-0003-2561-6760</mods:nameIdentifier>
<mods:affiliation id="80F6C4986B7F3A210A885893FD193B37">Department of Zoology, Swedish Museum of Natural History, PO Box 50007, SE- 104 05 Stockholm, Sweden &amp; michael. noren @ nrm. se; https: // orcid. org / 0000 - 0003 - 2561 - 6760</mods:affiliation>
<mods:nameIdentifier id="6C078AA220CFEF57D7B3D20DB7068A72" type="email">michael.noren@nrm.se</mods:nameIdentifier>
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<mods:date id="E3288B77243464F0D0CF32E400AC7761">2022</mods:date>
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<treatment id="08571F4FFFE51B5AFF28D0B3FAAE70D3" ID-DOI="http://doi.org/10.5281/zenodo.6312394" ID-GBIF-Taxon="193386703" ID-Zenodo-Dep="6312394" LSID="urn:lsid:plazi:treatment:08571F4FFFE51B5AFF28D0B3FAAE70D3" httpUri="http://treatment.plazi.org/id/08571F4FFFE51B5AFF28D0B3FAAE70D3" lastPageId="10" lastPageNumber="64" pageId="7" pageNumber="61">
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<heading id="DB091935FFE51B57FF28D0B3FDD972F6" bold="true" box="[151,531,1410,1437]" fontSize="11" level="1" pageId="7" pageNumber="61" reason="1">
<emphasis id="B28A724BFFE51B57FF28D0B3FDD972F6" bold="true" box="[151,531,1410,1437]" pageId="7" pageNumber="61">
<emphasis id="B28A724BFFE51B57FF28D0B3FEB172F7" bold="true" box="[151,379,1410,1437]" italics="true" pageId="7" pageNumber="61">
<taxonomicName id="47FED5DAFFE51B57FF28D0B3FEBC72F7" authority="Kullander &amp; Norén, 2022" authorityName="Kullander &amp; Norén" authorityYear="2022" box="[151,374,1410,1437]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="ahlanderi" status="sp. nov.">Devario ahlanderi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A9B9CF30FFE51B57FE3CD0B3FDD972F6" box="[387,531,1410,1437]" pageId="7" pageNumber="61" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8041AE59FFE51B57FF28D0FBFDF07147" blockId="7.[151,1437,1482,2013]" pageId="7" pageNumber="61">
<emphasis id="B28A724BFFE51B57FF28D0FBFEEE728E" bold="true" box="[151,292,1482,1509]" pageId="7" pageNumber="61">Type series.</emphasis>
<materialsCitation id="3096A404FFE51B57FE94D0FBFF197147" ID-GBIF-Occurrence="3467469301" accessionNumber="NRM 57999" collectingDate="2008-03-13" collectionCode="NRM" country="Myanmar" county="Naung Al Village" location="Thein Win" municipality="Kak-ku" pageId="7" pageNumber="61" specimenCount="1" stateProvince="Shan" typeStatus="holotype">
<typeStatus id="5F4510FBFFE51B57FE94D0FBFE5C728F" box="[299,406,1483,1509]" pageId="7" pageNumber="61" type="holotype">Holotype</typeStatus>
,
<collectionCode id="E6EF369CFFE51B57FE1CD0FBFE29728F" box="[419,483,1483,1509]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="7" pageNumber="61" type="Museum">NRM</collectionCode>
57999 (
<figureCitation id="18C5B2DCFFE51B57FDFFD0FBFD6B728F" box="[576,673,1483,1509]" captionStart="FIGURE 4" captionStartId="8.[151,250,1888,1912]" captionTargetBox="[152,1435,741,1865]" captionTargetId="figure-256@8.[152,1435,741,1865]" captionTargetPageId="8" captionText="FIGURE 4. Devario ahlanderi. All from Myanmar, Shan State, Salween River basin. A, holotype, NRM 57999, adult female, 65.2 mm SL; stream close to Naung Al Village, about 6 miles (9 km) east of Kak-ku. B, paratype, NRM 58062, adult male, 56.9 mm SL, same locality as holotype; C, paratype, NRM 58061, adult female, 55.6 mm SL; small stream running under Naung Pic bridge near Naung Pic Village, on the way from Kak-ku to Taunggyi." figureDoi="http://doi.org/10.5281/zenodo.6127757" httpUri="https://zenodo.org/record/6127757/files/figure.png" pageId="7" pageNumber="61">Fig. 4 A</figureCitation>
), adult female,
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SL;
<collectingCountry id="F8E9EEC9FFE51B57FC51D0FBFB95728F" box="[1006,1119,1483,1509]" name="Myanmar" pageId="7" pageNumber="61">Myanmar</collectingCountry>
:
<collectingRegion id="423A60BBFFE51B57FBD4D0FBFB2D728F" box="[1131,1255,1483,1509]" country="Myanmar" name="Shan" pageId="7" pageNumber="61">Shan State</collectingRegion>
: stream close to
<collectingCounty id="6920D6D5FFE51B57FF28D0DFFE947163" box="[151,350,1519,1545]" pageId="7" pageNumber="61">Naung Al Village</collectingCounty>
, about
<quantity id="470603BCFFE51B57FE12D0DFFE357163" box="[429,511,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="7" pageNumber="61" unit="mi" value="6.0">6 miles</quantity>
(
<quantity id="470603BCFFE51B57FDB1D0DFFD807160" box="[526,586,1519,1546]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="61" unit="km" value="9.0">9 km</quantity>
) east of
<collectingMunicipality id="60253423FFE51B57FD16D0DFFCCA7163" box="[681,768,1519,1545]" pageId="7" pageNumber="61">Kak-ku</collectingMunicipality>
, 20265N, 97132E;
<location id="8521F882FFE51B57FBBAD0DFFBB17163" LSID="urn:lsid:plazi:treatment:08571F4FFFE51B5AFF28D0B3FAAE70D3:8521F882FFE51B57FBBAD0DFFBB17163" box="[1029,1147,1519,1545]" country="Myanmar" county="Naung Al Village" municipality="Kak-ku" name="Thein Win" pageId="7" pageNumber="61" stateProvince="Shan">Thein Win</location>
and
<location id="8521F882FFE51B57FB0ED0DFFAEF7163" LSID="urn:lsid:plazi:treatment:08571F4FFFE51B5AFF28D0B3FAAE70D3:8521F882FFE51B57FB0ED0DFFAEF7163" box="[1201,1317,1519,1545]" country="Myanmar" county="Naung Al Village" municipality="Kak-ku" name="Thein Tan" pageId="7" pageNumber="61" stateProvince="Shan">Thein Tan</location>
,
<date id="F4408899FFE51B57FA8FD0DFFF197147" pageId="7" pageNumber="61" value="2008-03-13">
<collectingDate id="E4047171FFE51B57FA8FD0DFFF197147" pageId="7" pageNumber="61" value="2008-03-13">13 March 2008</collectingDate>
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.
<typeStatus id="5F4510FBFFE51B57FF62D323FE807147" box="[221,330,1555,1581]" pageId="7" pageNumber="61" type="paratype">Paratypes</typeStatus>
listed in Appendix 1.
</paragraph>
<paragraph id="8041AE59FFE51B57FF78D306FCC8711F" blockId="7.[151,1437,1482,2013]" pageId="7" pageNumber="61">
<emphasis id="B28A724BFFE51B57FF78D306FE8C713A" bold="true" box="[199,326,1590,1616]" pageId="7" pageNumber="61">Definition.</emphasis>
Distinguished from all other species of
<taxonomicName id="47FED5DAFFE51B57FCBAD307FCAB713A" authorityName="Heckel" authorityYear="1843" box="[773,865,1591,1616]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="genus">
<emphasis id="B28A724BFFE51B57FCBAD307FCAB713A" box="[773,865,1591,1616]" italics="true" pageId="7" pageNumber="61">Devario</emphasis>
</taxonomicName>
by the modification in adults of the anterior half of the P stripe into a series of short irregular bars or spots.
</paragraph>
</subSubSection>
<subSubSection id="C8E4FDD2FFE51B5AFF78D34EFAAE70D3" lastPageId="10" lastPageNumber="64" pageId="7" pageNumber="61" type="description">
<paragraph id="8041AE59FFE51B57FF78D34EFB4F71F3" blockId="7.[151,1437,1482,2013]" box="[199,1157,1662,1689]" pageId="7" pageNumber="61">
<emphasis id="B28A724BFFE51B57FF78D34EFE3E71F2" bold="true" box="[199,500,1662,1689]" pageId="7" pageNumber="61">
Description of
<typeStatus id="5F4510FBFFE51B57FEC8D34FFE6071F3" box="[375,426,1663,1689]" pageId="7" pageNumber="61">type</typeStatus>
series
</emphasis>
. Morphometric and meristic data are given in Appendix 2.
</paragraph>
<paragraph id="8041AE59FFE51B58FF28D393FC897677" blockId="7.[151,1437,1482,2013]" lastBlockId="8.[151,1437,150,717]" lastPageId="8" lastPageNumber="62" pageId="7" pageNumber="61">
Body laterally compressed, elongate, sexes not significantly different in body depth, but sample small. Predorsal contour straight ascending or with minor indentation above orbit, dorsal contour sloping posterior to posterior dorsal-fin insertion. Prepelvic contour strongly curved, more so in females; chest conspicuously more compressed below pectoral fin, but not keeled. Snout short, rounded in dorsal aspect, subtriangular in lateral aspect, about as long as eye diameter. Infraorbital process broader than high, with truncate distal margin which usually slightly irregular, not projecting from skin. Danionine notch caudally margined by well-developed anteromediad projecting laminar dentary process. Skin cover absent from distal part of dentary process. Mouth terminal, directed obliquely upwards. Bony knob at dentary symphysis minor, fitting in shallow depression in upper jaw. Maxilla reaching to below anterior margin of orbit. Jaws equal anteriorly; lower jaw ending anteriorly at horizontal through middle of eye. Three specimens,
<quantity id="470603BCFFEA1B58FE2BD5A6FDFC77DB" box="[404,566,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.955" metricValueMax="5.6899999999999995" metricValueMin="4.22" pageId="8" pageNumber="62" unit="mm" value="49.55" valueMax="56.9" valueMin="42.2">42.256.9 mm</quantity>
SL identified as adult males by presence of irregular band or two distinct rows of short tubercles along orad margin of dentary and presence of dense bands of sharp tubercles on pectoral-fin rays. Tubercles absent from lower jaw in
<typeStatus id="5F4510FBFFEA1B58FD95D5EFFD467793" box="[554,652,223,249]" pageId="8" pageNumber="62" type="holotype">holotype</typeStatus>
and remaining
<typeStatus id="5F4510FBFFEA1B58FC84D5EFFC607793" box="[827,938,223,249]" pageId="8" pageNumber="62" type="paratype">paratypes</typeStatus>
. Rostral barbel short, reaching to junction of lacrimal and first infraorbital; maxillary barbel much shorter.
</paragraph>
<paragraph id="8041AE59FFEA1B58FF78D417FC097628" blockId="8.[151,1437,150,717]" box="[199,963,294,322]" pageId="8" pageNumber="62">
Pharyngeal teeth in three rows: 5,42,2,4,5 (
<specimenCount id="96F865D0FFEA1B58FD16D417FC8E762B" box="[681,836,295,321]" pageId="8" pageNumber="62" type="generic">one specimen</specimenCount>
dissected).
</paragraph>
<paragraph id="8041AE59FFEA1B58FF78D47BFE4C7573" blockId="8.[151,1437,150,717]" pageId="8" pageNumber="62">Lateral line complete, along 3134 scales; comprising one tubed scale followed by a canal running steeply caudoventrad under unperforated scales to slightly posterior to pectoral-fin base, where curved caudad and represented by scales with indistinct or absent perforation anteriorly, becoming distinctly perforated posterior to adpressed pectoral fin; running in a curve parallel to the ventral body outline and ending low on caudal peduncle and caudal-fin base; vertical section represented by about six scales. A row of scales covering anal-fin base. About ¼ of caudal-fin length scaled basally.</paragraph>
<paragraph id="8041AE59FFEA1B58FF78D713FEB475A7" blockId="8.[151,1437,150,717]" pageId="8" pageNumber="62">Dorsal fin inserted at highest point of dorsum, distinctly posterior to middle of body. Anal fin inserted below anterior rays of dorsal fin. Pectoral-fin insertion at about vertical through posterior margin of osseous opercle; extending to pelvic-fin insertion. Pectoral axial lobe well developed. Pelvic fin inserted slightly anterior to midbody, not reaching to anterior insertion of anal fin. Pelvic axillary scale present. Caudal fin forked, lobes of about equal length, tips rounded.</paragraph>
<caption id="D481FED1FFEA1B58FF28D250FC9C708E" ID-DOI="http://doi.org/10.5281/zenodo.6127757" ID-Zenodo-Dep="6127757" httpUri="https://zenodo.org/record/6127757/files/figure.png" pageId="8" pageNumber="62" startId="8.[151,250,1888,1912]" targetBox="[152,1435,741,1865]" targetPageId="8">
<paragraph id="8041AE59FFEA1B58FF28D250FC9C708E" blockId="8.[151,1437,1888,2021]" pageId="8" pageNumber="62">
<emphasis id="B28A724BFFEA1B58FF28D250FED97012" bold="true" box="[151,275,1888,1912]" pageId="8" pageNumber="62">FIGURE 4.</emphasis>
<emphasis id="B28A724BFFEA1B58FEA5D251FE147012" box="[282,478,1888,1912]" italics="true" pageId="8" pageNumber="62">
<taxonomicName id="47FED5DAFFEA1B58FEA5D251FE107012" authorityName="Kullander &amp; Norén" authorityYear="2022" box="[282,474,1888,1912]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="62" phylum="Chordata" rank="species" species="ahlanderi">Devario ahlanderi</taxonomicName>
.
</emphasis>
All from Myanmar, Shan State, Salween River basin.
<emphasis id="B28A724BFFEA1B58FBB9D250FBD27012" bold="true" box="[1030,1048,1888,1912]" pageId="8" pageNumber="62">A</emphasis>
, holotype, NRM 57999, adult female, 65.2 mm SL; stream close to Naung Al Village, about 6 miles (9 km) east of Kak-ku.
<emphasis id="B28A724BFFEA1B58FC50D2B4FBCA70F6" bold="true" box="[1007,1024,1924,1948]" pageId="8" pageNumber="62">B</emphasis>
, paratype, NRM 58062, adult male, 56.9 mm SL, same locality as holotype;
<emphasis id="B28A724BFFEA1B58FE45D298FDD970AA" bold="true" box="[506,531,1960,1984]" pageId="8" pageNumber="62">C,</emphasis>
paratype, NRM 58061, adult female, 55.6 mm SL; small stream running under Naung Pic bridge near Naung Pic Village, on the way from Kak-ku to Taunggyi.
</paragraph>
</caption>
<caption id="D481FED1FFEB1B59FF28D260FF1F7092" ID-DOI="http://doi.org/10.5281/zenodo.6127759" ID-Zenodo-Dep="6127759" httpUri="https://zenodo.org/record/6127759/files/figure.png" pageId="9" pageNumber="63" startId="9.[151,250,1872,1896]" targetBox="[241,1347,181,1848]" targetPageId="9">
<paragraph id="8041AE59FFEB1B59FF28D260FF1F7092" blockId="9.[151,1437,1872,2040]" pageId="9" pageNumber="63">
<emphasis id="B28A724BFFEB1B59FF28D260FED97002" bold="true" box="[151,275,1872,1896]" pageId="9" pageNumber="63">FIGURE 5.</emphasis>
<taxonomicName id="47FED5DAFFEB1B59FEA5D261FE167002" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[282,476,1872,1896]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="63" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFEB1B59FEA5D261FEA77002" box="[282,365,1873,1896]" italics="true" pageId="9" pageNumber="63">Devario</emphasis>
cf.
<emphasis id="B28A724BFFEB1B59FE2AD260FE167002" box="[405,476,1872,1896]" italics="true" pageId="9" pageNumber="63">browni</emphasis>
</taxonomicName>
, all from P.R. China,Yunnan, Salween River basin.
<emphasis id="B28A724BFFEB1B59FC52D260FC357002" bold="true" box="[1005,1023,1872,1896]" pageId="9" pageNumber="63">A</emphasis>
, ZMUYU 874064, adult male, 60.5 mm SL: Canguyan County: Nangun River.
<emphasis id="B28A724BFFEB1B59FD8FD244FD8D70E6" bold="true" box="[560,583,1908,1932]" pageId="9" pageNumber="63">B.</emphasis>
ZMYU 80650, young female 42.1 mm SL; Gengma County: Qingshui River.
<emphasis id="B28A724BFFEB1B59FA3BD244FA5C70E6" bold="true" box="[1412,1430,1908,1932]" pageId="9" pageNumber="63">C</emphasis>
, ZMYU 80650, young female, 49.9 mm SL, right side, reversed: Gengma County: Qingshui River.
<emphasis id="B28A724BFFEB1B59FB2ED2A8FB6070DA" bold="true" box="[1169,1194,1944,1968]" pageId="9" pageNumber="63">D,</emphasis>
ZMYU, young female, 50.0 mm SL; Canguyan County: Nangun River;
<emphasis id="B28A724BFFEB1B59FD2AD28CFD6C70BE" bold="true" box="[661,678,1980,2004]" pageId="9" pageNumber="63">E</emphasis>
, ZMYU 80650, young female, 42.7 mm SL; Gengma County: Qingshui River.
</paragraph>
</caption>
<paragraph id="8041AE59FFE81B5AFF78D5A6FD5E7677" blockId="10.[151,1437,150,645]" pageId="10" pageNumber="64">
<emphasis id="B28A724BFFE81B5AFF78D5A6FDDB77DA" bold="true" box="[199,529,150,176]" pageId="10" pageNumber="64">Colouration in preservative</emphasis>
. Dorsum pale grey, dark grey line from occiput to base of upper caudal fin rays; sides off-white, chest and abdomen off-white or yellowish white depending on conservation. Opercle silvery, powdered with scattered melanophores; a brown patch dorsally. Dark brown vertically oriented cleithral spot covering part of first lateral-line scale and scale above.
</paragraph>
<paragraph id="8041AE59FFE81B5AFF78D417FA8B75EF" blockId="10.[151,1437,150,645]" pageId="10" pageNumber="64">
Adult flank colour pattern (
<figureCitation id="18C5B2DCFFE81B5AFE41D417FD88762B" box="[510,578,295,321]" captionStart="FIGURE 4" captionStartId="8.[151,250,1888,1912]" captionTargetBox="[152,1435,741,1865]" captionTargetId="figure-256@8.[152,1435,741,1865]" captionTargetPageId="8" captionText="FIGURE 4. Devario ahlanderi. All from Myanmar, Shan State, Salween River basin. A, holotype, NRM 57999, adult female, 65.2 mm SL; stream close to Naung Al Village, about 6 miles (9 km) east of Kak-ku. B, paratype, NRM 58062, adult male, 56.9 mm SL, same locality as holotype; C, paratype, NRM 58061, adult female, 55.6 mm SL; small stream running under Naung Pic bridge near Naung Pic Village, on the way from Kak-ku to Taunggyi." figureDoi="http://doi.org/10.5281/zenodo.6127757" httpUri="https://zenodo.org/record/6127757/files/figure.png" pageId="10" pageNumber="64">Fig. 4</figureCitation>
) varying with specimen size, and slightly on each side. Adults with black P stripe along middle of side from slight widening at end of caudal peduncle, running cephalad with irregular margins and increasing slightly in width anteroposteriorly; from about level of dorsal-fin anterior insertion replaced by black, irregularly shaped spots (rounded or as short vertical bars); anterior two spots much lighter and smaller than remainder. P-1 stripe narrower than P stripe; irregular, brown, extending from end of caudal peduncle cephalad to about vertical from anterior insertion of dorsal fin, where broken up in irregular spots paralleling or interdigitating with P stripe spots. P-2 stripe indistinct, grading into light abdomen. P+1 stripe extending from end of caudal peduncle, narrower than P stripe but wide medially and tapering in width at ends; consisting of brown blotches paralleling or interdigitating with P stripe blotches. Dorsal fin distally without pigment, basally light grey, translucent. Anal fin not pigmented. Middle rays of caudal fin light grey, fading posteriorly on fin. Pelvic fin white, translucent.
</paragraph>
<caption id="D481FED1FFE81B5AFF28D0B4FB6F72AA" ID-DOI="http://doi.org/10.5281/zenodo.6127761" ID-Zenodo-Dep="6127761" httpUri="https://zenodo.org/record/6127761/files/figure.png" pageId="10" pageNumber="64" startId="10.[151,250,1412,1436]" targetBox="[151,1435,669,1388]" targetPageId="10">
<paragraph id="8041AE59FFE81B5AFF28D0B4FB6F72AA" blockId="10.[151,1436,1412,1472]" pageId="10" pageNumber="64">
<emphasis id="B28A724BFFE81B5AFF28D0B4FEDE72F6" bold="true" box="[151,276,1412,1436]" pageId="10" pageNumber="64">FIGURE 6.</emphasis>
<emphasis id="B28A724BFFE81B5AFEA3D0B4FE1572F6" box="[284,479,1412,1436]" italics="true" pageId="10" pageNumber="64">
<taxonomicName id="47FED5DAFFE81B5AFEA3D0B4FE1172F6" authorityName="Kullander &amp; Norén" authorityYear="2022" box="[284,475,1412,1436]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="ahlanderi">Devario ahlanderi</taxonomicName>
,
</emphasis>
subadult paratypes.
<emphasis id="B28A724BFFE81B5AFD0DD0B4FD0E72F6" bold="true" box="[690,708,1412,1436]" pageId="10" pageNumber="64">A</emphasis>
, 42.0 mm SL;
<emphasis id="B28A724BFFE81B5AFCDED0B4FCB872F6" bold="true" box="[865,882,1412,1436]" pageId="10" pageNumber="64">B</emphasis>
, 36.9 mm SL; both from Myanmar, Shan State, small stream running under Naung Pic bridge near Naung Pic Village, on the way from Kak-ku to Taunggyi.
</paragraph>
</caption>
<paragraph id="8041AE59FFE81B5AFF78D0DEFBD071F2" blockId="10.[151,1437,1518,1977]" pageId="10" pageNumber="64">
Juveniles similar to adults, but spotted pattern absent (
<figureCitation id="18C5B2DCFFE81B5AFC8CD0DEFCB37163" box="[819,889,1518,1545]" captionStart="FIGURE 6" captionStartId="10.[151,250,1412,1436]" captionTargetBox="[151,1435,669,1388]" captionTargetId="figure-256@10.[151,1435,669,1388]" captionTargetPageId="10" captionText="FIGURE 6. Devario ahlanderi, subadult paratypes. A, 42.0 mm SL; B, 36.9 mm SL; both from Myanmar, Shan State, small stream running under Naung Pic bridge near Naung Pic Village, on the way from Kak-ku to Taunggyi." figureDoi="http://doi.org/10.5281/zenodo.6127761" httpUri="https://zenodo.org/record/6127761/files/figure.png" pageId="10" pageNumber="64">Fig. 6</figureCitation>
). Smallest juveniles available
<quantity id="470603BCFFE81B5AFB6BD0DEFAEC7163" box="[1236,1318,1518,1545]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="64" unit="mm" value="20.0">20 mm</quantity>
, with pale brown mid-dorsal stripe, black line along middle of side, and suffused brown or grey pigmentation on lower flank. At about
<quantity id="470603BCFFE81B5AFEBAD306FE8A713B" box="[261,320,1590,1617]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="64" unit="m" value="30.0">30 m</quantity>
SL, wide brown P stripe, slightly increasing in width anteriorly, with contrasting marginal interstripes. At about
<quantity id="470603BCFFE81B5AFEE8D36AFE6D711F" box="[343,423,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="64" unit="mm" value="40.0">40 mm</quantity>
SL each specimen with unique colour pattern with irregular bars or small spots anteriorly in the P stripe, and the marginal stripes continuous or not in front of the P stripe.
</paragraph>
<paragraph id="8041AE59FFE81B5AFF78D392FCD47026" blockId="10.[151,1437,1518,1977]" pageId="10" pageNumber="64">
<emphasis id="B28A724BFFE81B5AFF78D392FD5871D6" bold="true" box="[199,658,1698,1724]" pageId="10" pageNumber="64">Geographical distribution and habitat.</emphasis>
<taxonomicName id="47FED5DAFFE81B5AFD27D393FCAC71D6" authorityName="Kullander &amp; Norén" authorityYear="2022" box="[664,870,1698,1724]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="ahlanderi">
<emphasis id="B28A724BFFE81B5AFD27D393FCAC71D6" box="[664,870,1698,1724]" italics="true" pageId="10" pageNumber="64">Devario ahlanderi</emphasis>
</taxonomicName>
is known only from two small streams near Taunggyi (
<figureCitation id="18C5B2DCFFE81B5AFF73D3F6FED9718A" box="[204,275,1734,1760]" captionStart="FIGURE 1" captionStartId="1.[151,250,1935,1959]" captionTargetBox="[171,1418,1070,1902]" captionTargetId="figure-368@1.[163,1425,1062,1908]" captionTargetPageId="1" captionText="FIGURE 1. Map of sampling sites of Devario ahlanderi, D. browni, Devario cf. browni, and D. kysonensis." figureDoi="http://doi.org/10.5281/zenodo.6127751" httpUri="https://zenodo.org/record/6127751/files/figure.png" pageId="10" pageNumber="64">Figs 1</figureCitation>
). The
<typeStatus id="5F4510FBFFE81B5AFEDED3F7FE5B718B" box="[353,401,1735,1761]" pageId="10" pageNumber="64">type</typeStatus>
locality was a small shaded stream 100% shaded by bamboo trees.
<taxonomicName id="47FED5DAFFE81B5AFB21D3F7FAA2718A" baseAuthorityName="Barman" baseAuthorityYear="1985" box="[1182,1384,1735,1760]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="jayarami">
<emphasis id="B28A724BFFE81B5AFB21D3F7FAA2718A" box="[1182,1384,1735,1760]" italics="true" pageId="10" pageNumber="64">Devario jayarami</emphasis>
</taxonomicName>
was collected together with
<taxonomicName id="47FED5DAFFE81B5AFE1CD3DBFDF9706E" authorityName="Kullander &amp; Norén" authorityYear="2022" box="[419,563,1770,1796]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="ahlanderi">
<emphasis id="B28A724BFFE81B5AFE1CD3DBFDF9706E" box="[419,563,1770,1796]" italics="true" pageId="10" pageNumber="64">D. ahlanderi</emphasis>
</taxonomicName>
, and
<taxonomicName id="47FED5DAFFE81B5AFDCCD3DBFCE3706E" baseAuthorityName="Hora and Mukerji" baseAuthorityYear="1934" box="[627,809,1770,1796]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="sondhii">
<emphasis id="B28A724BFFE81B5AFDCCD3DBFCE3706E" box="[627,809,1770,1796]" italics="true" pageId="10" pageNumber="64">Devario sondhii</emphasis>
</taxonomicName>
was available in the Taunggyi market together with
<taxonomicName id="47FED5DAFFE81B5AFA3ED3DBFECE7042" authorityName="Kullander &amp; Norén" authorityYear="2022" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="ahlanderi">
<emphasis id="B28A724BFFE81B5AFA3ED3DBFECE7042" italics="true" pageId="10" pageNumber="64">D. ahlanderi</emphasis>
</taxonomicName>
(Appendix 1). Other syntopic species were identified as
<emphasis id="B28A724BFFE81B5AFC36D23EFE707026" italics="true" pageId="10" pageNumber="64">
<taxonomicName id="47FED5DAFFE81B5AFC36D23EFB247042" baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[905,1262,1806,1832]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalichthys" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="berdmorei">Lepidocephalichthys berdmorei</taxonomicName>
,
<taxonomicName id="47FED5DAFFE81B5AFB45D23EFF1A7026" baseAuthorityName="Day" baseAuthorityYear="1871" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="stoliczkana">Pethia stoliczkana</taxonomicName>
,
<taxonomicName id="47FED5DAFFE81B5AFF63D203FE707026" baseAuthorityName="Roberts" baseAuthorityYear="2007" box="[220,442,1843,1868]" class="Actinopterygii" family="Cyprinidae" genus="Danio" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="margaritatus">Danio margaritatus</taxonomicName>
</emphasis>
, and
<emphasis id="B28A724BFFE81B5AFE48D202FCD47026" box="[503,798,1842,1868]" italics="true" pageId="10" pageNumber="64">
<taxonomicName id="47FED5DAFFE81B5AFE48D202FCD37026" baseAuthorityName="Pallas" baseAuthorityYear="1770" box="[503,793,1842,1868]" class="Actinopterygii" family="Osphronemidae" genus="Trichogaster" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="64" phylum="Chordata" rank="species" species="trichopterus">Trichogaster trichopterus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8041AE59FFE81B5AFF78D266FAAE70D3" blockId="10.[151,1437,1518,1977]" pageId="10" pageNumber="64">
<emphasis id="B28A724BFFE81B5AFF78D266FD92701A" bold="true" box="[199,600,1878,1904]" pageId="10" pageNumber="64">Explanation of the specific name:</emphasis>
The specific name is a noun in the genitive case. The species is dedicated to Erik Åhlander, long time Senior Assistant in the ichthtyology and herpetology collection of the Swedish Museum of Natural History, and a key person in the successful development and operation of ichthyology at the NRM.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,241 @@
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<mods:title id="48A19C89F824DC4EAA8E11D0FA86B3C7">The real Devario browni from the Irrawaddy River basin, and the new Devario ahlanderi from the Salween River basin in Myanmar (Teleostei: Cyprinidae: Danioninae)</mods:title>
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<mods:namePart id="89043E799A3725628C143528D0185F7A">Kullander, Sven</mods:namePart>
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<mods:affiliation id="61419F6D4130EF3441156B4B56D484BA">Department of Zoology, Swedish Museum of Natural History, PO Box 50007, SE- 104 05 Stockholm, Sweden &amp; sven. kullander @ nrm. se; https: // orcid. org / 0000 - 0001 - 6075 - 0266</mods:affiliation>
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<mods:namePart id="FC55F5B05B1AEB657CD607AA3EFB2ACE">Norén, Michael</mods:namePart>
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<mods:affiliation id="F7EC8B9C18CD70071D7CE0F22E265F24">Department of Zoology, Swedish Museum of Natural History, PO Box 50007, SE- 104 05 Stockholm, Sweden &amp; michael. noren @ nrm. se; https: // orcid. org / 0000 - 0003 - 2561 - 6760</mods:affiliation>
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<subSubSection id="C8E4FDD2FFE61B54FF28D234FAE5709C" pageId="4" pageNumber="58" type="nomenclature">
<paragraph id="8041AE59FFE61B54FF28D234FDCD7074" blockId="4.[151,519,1795,1822]" box="[151,519,1795,1822]" pageId="4" pageNumber="58">
<heading id="DB091935FFE61B54FF28D234FDCD7074" bold="true" box="[151,519,1795,1822]" fontSize="11" level="1" pageId="4" pageNumber="58" reason="1">
<taxonomicName id="47FED5DAFFE61B54FF28D234FDCD7074" ID-CoL="355SS" authority="(Regan, 1907)" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[151,519,1795,1822]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="58" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFE61B54FF28D234FDCD7074" bold="true" box="[151,519,1795,1822]" pageId="4" pageNumber="58">
<emphasis id="B28A724BFFE61B54FF28D234FE9E7074" bold="true" box="[151,340,1795,1822]" italics="true" pageId="4" pageNumber="58">Devario browni</emphasis>
(
<bibRefCitation id="E46FD3A8FFE61B54FEDCD234FE357074" author="Regan, C. T." box="[355,511,1796,1822]" pageId="4" pageNumber="58" pagination="395" refId="ref9443" refString="Regan, C. T. (1907) Description of a new cyprinid fish of the genus Danio from Upper Burma. Records of the Indian Museum, 4, 395." type="journal article" year="1907">Regan, 1907</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8041AE59FFE61B54FF28D27DFAE5709C" blockId="4.[151,1437,1868,2038]" pageId="4" pageNumber="58">
<materialsCitation id="3096A404FFE61B54FF28D27DFAE5709C" ID-GBIF-Occurrence="3467469302" collectionCode="BMNH" collectorName="Recommendations A" location="International Code of Zoological" pageId="4" pageNumber="58" specimenCode="BMNH 19071907.10" specimenCount="4" typeStatus="lectotype">
<typeStatus id="5F4510FBFFE61B54FF28D27DFECD700D" box="[151,263,1869,1895]" pageId="4" pageNumber="58" type="lectotype">Lectotype</typeStatus>
<specimenCode id="D0580622FFE61B54FEB0D27DFDCC700C" box="[271,518,1868,1895]" collectionCode="BMNH" pageId="4" pageNumber="58">BMNH 19071907.10</specimenCode>
.22.1, Adult male. 57.0 mm SL (
<figureCitation id="18C5B2DCFFE61B54FCD0D27CFC00700C" box="[879,970,1868,1894]" captionStart="FIGURE 2" captionStartId="5.[151,250,997,1021]" captionTargetBox="[151,1435,204,955]" captionTargetId="figure-24@5.[151,1435,204,955]" captionTargetPageId="5" captionText="FIGURE 2. Devario browni. BMNH 1907.10.22:14; A, Lectotype, male 57.0 mm SL; B, Paralectotype, female, 52.5 mm SL; Northern Shan States." figureDoi="http://doi.org/10.5281/zenodo.6127753" httpUri="https://zenodo.org/record/6127753/files/figure.png" pageId="4" pageNumber="58">Fig. 2A</figureCitation>
);
<typeStatus id="5F4510FBFFE61B54FC5FD27CFB48700C" box="[992,1154,1868,1894]" pageId="4" pageNumber="58" type="paralectotype">paralectotypes</typeStatus>
<collectionCode id="E6EF369CFFE61B54FB35D27DFB14700D" box="[1162,1246,1869,1895]" pageId="4" pageNumber="58">BMNH</collectionCode>
1907.10.22.24, ZSI F1872/1. The
<typeStatus id="5F4510FBFFE61B54FED7D240FE0570E0" box="[360,463,1904,1930]" pageId="4" pageNumber="58" type="lectotype">lectotype</typeStatus>
designation here is motivated by the uncertainty about which specimens belong in the type series, and the taxonomic uncertainty resulting from previous misidentifications. Data in this description and
<typeStatus id="5F4510FBFFE61B54FF28D288FF3470B8" box="[151,254,1976,2002]" pageId="4" pageNumber="58" type="lectotype">lectotype</typeStatus>
designation fulfil paragraph74.7 and as far as relevant
<collectorName id="2D0BCB8FFFE61B54FCDDD288FB8470B8" box="[866,1102,1976,2002]" pageId="4" pageNumber="58">Recommendations A</collectorName>
, C, D, E, and F of the
<location id="8521F882FFE61B54FAFBD289FE7D709C" LSID="urn:lsid:plazi:treatment:08571F4FFFE61B57FF28D234FCE1725A:8521F882FFE61B54FAFBD289FE7D709C" name="International Code of Zoological" pageId="4" pageNumber="58">International Code of Zoological</location>
nomenclature (International Commission for
<bibRefCitation id="E46FD3A8FFE61B54FC05D2ECFAE9709C" author="International Commission on Zoological Nomenclature" box="[954,1315,2012,2038]" pageId="4" pageNumber="58" refId="ref8715" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. Fourth Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">Zoological Nomenclature, 1999</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="D481FED1FFE71B55FF28D6D5FEBC734B" ID-DOI="http://doi.org/10.5281/zenodo.6127753" ID-Zenodo-Dep="6127753" httpUri="https://zenodo.org/record/6127753/files/figure.png" pageId="5" pageNumber="59" startId="5.[151,250,997,1021]" targetBox="[151,1435,204,955]" targetPageId="5">
<paragraph id="8041AE59FFE71B55FF28D6D5FEBC734B" blockId="5.[151,1436,997,1057]" pageId="5" pageNumber="59">
<emphasis id="B28A724BFFE71B55FF28D6D5FED97497" bold="true" box="[151,275,997,1021]" pageId="5" pageNumber="59">FIGURE 2.</emphasis>
<taxonomicName id="47FED5DAFFE71B55FEA6D6D6FE727497" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[281,440,997,1021]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="59" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFE71B55FEA6D6D6FE727497" box="[281,440,997,1021]" italics="true" pageId="5" pageNumber="59">Devario browni</emphasis>
</taxonomicName>
. BMNH 1907.10.22:14;
<emphasis id="B28A724BFFE71B55FD7DD6D5FD1E7497" bold="true" box="[706,724,997,1021]" pageId="5" pageNumber="59">A</emphasis>
, Lectotype, male 57.0 mm SL;
<emphasis id="B28A724BFFE71B55FBABD6D5FBEF7497" bold="true" box="[1044,1061,997,1021]" pageId="5" pageNumber="59">B</emphasis>
, Paralectotype, female, 52.5 mm SL; Northern Shan States.
</paragraph>
</caption>
<subSubSection id="C8E4FDD2FFE71B57FF78D17FFCE1725A" lastPageId="7" lastPageNumber="61" pageId="5" pageNumber="59" type="description">
<paragraph id="8041AE59FFE71B55FF78D17FFBF173D8" blockId="5.[151,1437,1103,1993]" pageId="5" pageNumber="59">
<emphasis id="B28A724BFFE71B55FF78D17FFE6C7303" bold="true" box="[199,422,1103,1129]" pageId="5" pageNumber="59">Revised definition.</emphasis>
Distinguished from all other species of
<taxonomicName id="47FED5DAFFE71B55FCD8D160FC097303" authorityName="Heckel" authorityYear="1843" box="[871,963,1104,1129]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B28A724BFFE71B55FCD8D160FC097303" box="[871,963,1104,1129]" italics="true" pageId="5" pageNumber="59">Devario</emphasis>
</taxonomicName>
by the shape of the P stripe: gradually expanding posteroanteriorly, either forming a uniform brown blotch anteriorly, or P stripe diverging anteriorly, forming a brown blotch enclosing a small light spot or two contiguous small light spots.
</paragraph>
<paragraph id="8041AE59FFE71B55FF78D18BFC987393" blockId="5.[151,1437,1103,1993]" pageId="5" pageNumber="59">
<emphasis id="B28A724BFFE71B55FF78D18BFCBD73BF" bold="true" box="[199,887,1211,1237]" pageId="5" pageNumber="59">
Description based on referred specimens and
<typeStatus id="5F4510FBFFE71B55FD4ED18BFCEE73BF" box="[753,804,1211,1237]" pageId="5" pageNumber="59">type</typeStatus>
series.
</emphasis>
Representative specimens are shown in
<figureCitation id="18C5B2DCFFE71B55FAF7D18BFF0E7393" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="5.[151,250,997,1021]" captionStartId-1="6.[151,250,1943,1967]" captionTargetBox-0="[151,1435,204,955]" captionTargetBox-1="[193,1395,181,1919]" captionTargetId-0="figure-24@5.[151,1435,204,955]" captionTargetId-1="figure-19@6.[193,1395,181,1919]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 2. Devario browni. BMNH 1907.10.22:14; A, Lectotype, male 57.0 mm SL; B, Paralectotype, female, 52.5 mm SL; Northern Shan States." captionText-1="FIGURE 3. Devario browni. A, NRM 36393, adult male, 62.1 mm SL, reversed; B, adult female, 58.3 mm SL, reversed; C, young male, 44.9 mm SL, reversed; D, young male, 49.5 mm SL; E, NRM 36361, young male 49.7 mm SL. All from Myanmar, Shan State, Irrawaddy River basin, Lawe Pyat stream, on Road LashioTigyaing." figureDoi-0="http://doi.org/10.5281/zenodo.6127753" figureDoi-1="http://doi.org/10.5281/zenodo.6127755" httpUri-0="https://zenodo.org/record/6127753/files/figure.png" httpUri-1="https://zenodo.org/record/6127755/files/figure.png" pageId="5" pageNumber="59">Figures 23</figureCitation>
. Morphometric and meristic data are given in Appendix 2.
</paragraph>
<paragraph id="8041AE59FFE71B55FF78D033FBF3707C" blockId="5.[151,1437,1103,1993]" pageId="5" pageNumber="59">
Body laterally compressed, elongate; sexes significantly different in body depth (ANCOVA, p&gt;0.05, but only
<specimenCount id="96F865D0FFE71B55FF28D017FEC4722B" box="[151,270,1319,1345]" pageId="5" pageNumber="59" type="adult">eight adult</specimenCount>
specimens were available. Predorsal contour ascending in straight line, sloping posteriorly from anterior dorsal-fin insertion. Prepelvic contour strongly curved; chest conspicuously compressed below pectoral fin, but not keeled. Snout short, rounded in dorsal aspect, subtriangular in lateral aspect, about as long as eye diameter. Infraorbital process broader than high, with truncate distal margin which may be slightly irregular. Danionine notch caudally margined by well-developed anteromediad projecting laminar dentary process. Skin thin or absent from distal part of infraorbital process, dentary process, and anterior margin of supraorbital. Mouth terminal, obliquely directed upwards. Bony knob at dentary symphysis minor, fitting in shallow depression in upper jaw. Blunt tubercles present in adults in 12 (males) or 1 row (females) on lateral adoral surface of dentary, and additional scattered tubercles anteriorly on dentary. Maxilla reaching to below anterior margin of orbit. Jaws equal in anterior extension; lower jaw ending anteriorly at horizontal through middle of eye. In large males, bands of strong, densely arranged sharp-tipped conical tubercles on anterior seven rays of pectoral fin; pectoral-fin tubercles absent in females, but
<specimenCount id="96F865D0FFE71B55FF28D384FEF871A7" box="[151,306,1715,1742]" pageId="5" pageNumber="59" type="generic">one specimen</specimenCount>
,
<quantity id="470603BCFFE71B55FE83D383FE5571A4" box="[316,415,1715,1742]" metricMagnitude="-2" metricUnit="m" metricValue="5.9799999999999995" pageId="5" pageNumber="59" unit="mm" value="59.8">59.8 mm</quantity>
SL potentially female with spaced tubercles on three anterior pectoral fin rays and relatively deep body. Rostral barbel short, reaching slightly beyond junction of lacrimal and first infraorbital; maxillary barbel much shorter. Pharyngeal teeth in three rows: 5,4,2,2,4,5 (
<specimenCount id="96F865D0FFE71B55FC9FD3CBFC71707F" box="[800,955,1787,1813]" pageId="5" pageNumber="59" type="generic">one specimen</specimenCount>
dissected).
</paragraph>
<paragraph id="8041AE59FFE71B55FF78D22FFE3570A3" blockId="5.[151,1437,1103,1993]" pageId="5" pageNumber="59">Lateral line complete, comprising one tubed scale followed by a canal running steeply caudoventrad under five unperforated scales to slightly posterior to pectoral-fin base where canal curved caudad and represented by scales with indistinct or absent perforation anteriorly, but distinctly perforated posterior to adpressed pectoral fin; running in a curve parallel to the ventral body outline and ending low on caudal peduncle and caudal-fin base. About of caudal-fin length scaled basally.</paragraph>
<caption id="D481FED1FFE41B56FF28D2A7FC1A709D" ID-DOI="http://doi.org/10.5281/zenodo.6127755" ID-Zenodo-Dep="6127755" httpUri="https://zenodo.org/record/6127755/files/figure.png" pageId="6" pageNumber="60" startId="6.[151,250,1943,1967]" targetBox="[193,1395,181,1919]" targetPageId="6">
<paragraph id="8041AE59FFE41B56FF28D2A7FC1A709D" blockId="6.[151,1437,1943,2039]" pageId="6" pageNumber="60">
<emphasis id="B28A724BFFE41B56FF28D2A7FEC470C5" bold="true" box="[151,270,1943,1967]" pageId="6" pageNumber="60">FIGURE 3</emphasis>
.
<taxonomicName id="47FED5DAFFE41B56FEA3D2A8FE7770C5" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[284,445,1943,1967]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="60" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFE41B56FEA3D2A8FE7770C5" box="[284,445,1943,1967]" italics="true" pageId="6" pageNumber="60">Devario browni</emphasis>
</taxonomicName>
.
<emphasis id="B28A724BFFE41B56FE74D2A7FE2970C5" bold="true" box="[459,483,1943,1967]" pageId="6" pageNumber="60">A,</emphasis>
NRM 36393, adult male, 62.1 mm SL, reversed;
<emphasis id="B28A724BFFE41B56FC5AD2A7FC3670C5" bold="true" box="[997,1020,1943,1967]" pageId="6" pageNumber="60">B,</emphasis>
adult female, 58.3 mm SL, reversed;
<emphasis id="B28A724BFFE41B56FA3BD2A7FA5770C5" bold="true" box="[1412,1437,1943,1967]" pageId="6" pageNumber="60">C,</emphasis>
young male, 44.9 mm SL, reversed;
<emphasis id="B28A724BFFE41B56FDBBD28BFDDC70B9" bold="true" box="[516,534,1979,2003]" pageId="6" pageNumber="60">D</emphasis>
, young male, 49.5 mm SL;
<emphasis id="B28A724BFFE41B56FC92D28BFCF470B9" bold="true" box="[813,830,1979,2003]" pageId="6" pageNumber="60">E</emphasis>
, NRM 36361, young male 49.7 mm SL. All from Myanmar, Shan State, Irrawaddy River basin, Lawe Pyat stream, on Road LashioTigyaing.
</paragraph>
</caption>
<paragraph id="8041AE59FFE51B57FF78D5A7FA9A7677" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">Dorsal fin inserted at highest point of dorsum, slightly posterior to middle of body.Anal fin inserted below anterior rays of dorsal fin. Pectoral-fin insertion at about vertical through posterior margin of osseous opercle; extending to pelvic-fin insertion. Pectoral axial lobe present. Pelvic fin inserted slightly anterior to midbody, not reaching to anal-fin insertion. Pelvic axillary scale present. Caudal fin forked, lobes of about equal length, tips rounded.</paragraph>
<paragraph id="8041AE59FFE51B57FF78D416FD5E76C7" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">
<emphasis id="B28A724BFFE51B57FF78D416FDC5762A" bold="true" box="[199,527,294,320]" pageId="7" pageNumber="61">Colouration in preservative</emphasis>
. Dorsum pale brown; dark brown mid-dorsal stripe from occiput to base of upper caudal-fin rays; lower sides, chest and abdomen pale yellowish white. Caudal peduncle ventrally yellowish white. Opercle silvery, powdered with brown; a brown patch dorsally. Vertically oriented brown cleithral spot covering part of first lateral-line scale and scale above.
</paragraph>
<paragraph id="8041AE59FFE51B57FF78D487FAF47557" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">Flank colour pattern varying with specimen size, and slightly on each side. Adults with dark brown P stripe along middle of side, narrow at end of caudal peduncle, irregular, gradually widening cephalad to about doubled width or more at abrupt anterior termination. Light spot anteriorly in P stripe present or absent. P+1 narrower than P stripe, light brown, paralleling P stripe, running from caudal peduncle to anterior termination of P stripe.</paragraph>
<paragraph id="8041AE59FFE51B57FF78D777FF74759B" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">P+2 stripe absent. P-1 stripe narrower than P stripe; irregular, greyish brown, well demarcated against I-1 stripe, fading ventrally; extending from end of caudal peduncle cephalad to about level of anterior termination of P stripe. Two or three brown vertical bars or ovoid blotches anterior to P stripe. Interstripes I+1 and I-1 joined anterior to termination of P stripe, or light interspace of vertical bar anterior to P stripe bridging ends of interstripes I+1 and I-1.</paragraph>
<paragraph id="8041AE59FFE51B57FF78D7CBFC2C7453" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">Dorsal fin proximally pale grey, translucent; distal half diaphanous. Middle rays of caudal fin light grey, translucent, fading posteriorly on fin. Pectoral and pelvic fins white, translucent.</paragraph>
<paragraph id="8041AE59FFE51B57FF78D673FC6A74CF" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">
Juveniles about
<quantity id="470603BCFFE51B57FE3CD672FE1C7437" box="[387,470,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="61" unit="mm" value="25.0">25 mm</quantity>
similar to adults but interstripes not connected anteriorly; P stripe brown, distinct, relatively narrower than in adults. In specimens about
<quantity id="470603BCFFE51B57FD45D657FC8174EB" box="[762,843,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="61" unit="mm" value="18.0">18 mm</quantity>
, SL and shorter, horizontal stripes indistinct; P stripe represented by light brown pigmentation suffused on abdominal side.
</paragraph>
<paragraph id="8041AE59FFE51B57FF78D69FFCF7737B" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">
<emphasis id="B28A724BFFE51B57FF78D69FFDB374A3" bold="true" box="[199,633,942,969]" pageId="7" pageNumber="61">
<taxonomicName id="47FED5DAFFE51B57FF78D69FFE8674A3" box="[199,332,943,969]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B28A724BFFE51B57FF78D69FFEE974A3" bold="true" box="[199,291,943,969]" italics="true" pageId="7" pageNumber="61">Devario</emphasis>
sp.
</taxonomicName>
<taxonomicName id="47FED5DAFFE51B57FEEAD69EFE0174A3" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[341,459,942,969]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="browni">
cf.
<emphasis id="B28A724BFFE51B57FEC5D69EFE0174A3" bold="true" box="[378,459,942,969]" italics="true" pageId="7" pageNumber="61">browni</emphasis>
</taxonomicName>
from
<collectingRegion id="423A60BBFFE51B57FDABD69FFDBE74A3" box="[532,628,943,969]" country="China" name="Yunnan" pageId="7" pageNumber="61">Yunnan</collectingRegion>
.
</emphasis>
<collectingRegion id="423A60BBFFE51B57FD3ED69FFD1D74A3" box="[641,727,943,969]" country="China" name="Yunnan" pageId="7" pageNumber="61">Yunnan</collectingRegion>
specimens inadequately preserved, overall brownish and most specimens with indistinct colour pattern (
<figureCitation id="18C5B2DCFFE51B57FDD2D6E3FD797487" box="[621,691,978,1005]" captionStart="FIGURE 5" captionStartId="9.[151,250,1872,1896]" captionTargetBox="[241,1347,181,1848]" captionTargetId="figure-24@9.[241,1347,181,1848]" captionTargetPageId="9" captionText="FIGURE 5. Devario cf. browni, all from P.R. China,Yunnan, Salween River basin. A, ZMUYU 874064, adult male, 60.5 mm SL: Canguyan County: Nangun River. B. ZMYU 80650, young female 42.1 mm SL; Gengma County: Qingshui River. C, ZMYU 80650, young female, 49.9 mm SL, right side, reversed: Gengma County: Qingshui River. D, ZMYU, young female, 50.0 mm SL; Canguyan County: Nangun River; E, ZMYU 80650, young female, 42.7 mm SL; Gengma County: Qingshui River." figureDoi="http://doi.org/10.5281/zenodo.6127759" httpUri="https://zenodo.org/record/6127759/files/figure.png" pageId="7" pageNumber="61">Fig. 5</figureCitation>
). Body proportions and meristic data similar to
<taxonomicName id="47FED5DAFFE51B57FB69D6E3FA827487" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[1238,1352,979,1005]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFE51B57FB69D6E3FA827487" box="[1238,1352,979,1005]" italics="true" pageId="7" pageNumber="61">D. browni</emphasis>
</taxonomicName>
and
<taxonomicName id="47FED5DAFFE51B57FA3ED6E3FECE737B" authorityName="Kullander &amp; Norén" authorityYear="2022" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="ahlanderi">
<emphasis id="B28A724BFFE51B57FA3ED6E3FECE737B" italics="true" pageId="7" pageNumber="61">D. ahlanderi</emphasis>
</taxonomicName>
, but specimens not in a good state of preservation.
</paragraph>
<paragraph id="8041AE59FFE51B57FF78D12BFC5C7317" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">Most specimens from the Nangun River with relatively narrow P stripe, only slightly widened anteriorly and terminating in a short vertical bar. Specimens from the Qingshui River (in the Nangun River basin) with large or small dark blotch terminating P stripe at level of pelvic fin insertion.</paragraph>
<paragraph id="8041AE59FFE51B57FF78D1B6FCE1725A" blockId="7.[151,1437,151,1329]" pageId="7" pageNumber="61">
<emphasis id="B28A724BFFE51B57FF78D1B6FD5D73CA" bold="true" box="[199,663,1158,1185]" pageId="7" pageNumber="61">Geographical distribution and habitat.</emphasis>
The
<typeStatus id="5F4510FBFFE51B57FD6FD1B7FCCA73CB" box="[720,768,1159,1185]" pageId="7" pageNumber="61">type</typeStatus>
locality of
<taxonomicName id="47FED5DAFFE51B57FC3DD1B7FBF973CB" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[898,1075,1159,1185]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFE51B57FC3DD1B7FBF973CB" box="[898,1075,1159,1185]" italics="true" pageId="7" pageNumber="61">Devario browni</emphasis>
</taxonomicName>
is not known, but was probably near Lashio in the upper
<collectingRegion id="423A60BBFFE51B57FE10D19BFDE873AF" box="[431,546,1195,1221]" country="Myanmar" name="Ayeyawady" pageId="7" pageNumber="61">Irrawaddy</collectingRegion>
River basin. Specimens from streams near Lashio were identified as
<taxonomicName id="47FED5DAFFE51B57FA93D19BFA5673AF" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[1324,1436,1195,1221]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFE51B57FA93D19BFA5673AF" box="[1324,1436,1195,1221]" italics="true" pageId="7" pageNumber="61">D. browni</emphasis>
</taxonomicName>
(Appendix 1). Presence of
<taxonomicName id="47FED5DAFFE51B57FE6FD1FFFD8E7383" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[464,580,1231,1257]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFE51B57FE6FD1FFFD8E7383" box="[464,580,1231,1257]" italics="true" pageId="7" pageNumber="61">D. browni</emphasis>
</taxonomicName>
in the upper Salween River basin in
<collectingRegion id="423A60BBFFE51B57FC46D1FFFB857383" box="[1017,1103,1231,1257]" country="China" name="Yunnan" pageId="7" pageNumber="61">Yunnan</collectingRegion>
was not confirmed.
<taxonomicName id="47FED5DAFFE51B57FAFFD1FFFECE7267" baseAuthorityName="Hora" baseAuthorityYear="1928" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="shanensis">
<emphasis id="B28A724BFFE51B57FAFFD1FFFECE7267" italics="true" pageId="7" pageNumber="61">Devario shanensis</emphasis>
</taxonomicName>
was syntopic with
<taxonomicName id="47FED5DAFFE51B57FE59D1C3FD937267" baseAuthorityName="Regan" baseAuthorityYear="1907" box="[486,601,1267,1293]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="browni">
<emphasis id="B28A724BFFE51B57FE59D1C3FD937267" box="[486,601,1267,1293]" italics="true" pageId="7" pageNumber="61">D. browni</emphasis>
</taxonomicName>
near Lashio (Appendix 1).Other syntopic fish species were identified as
<emphasis id="B28A724BFFE51B57FF28D027FD92725B" box="[151,600,1303,1329]" italics="true" pageId="7" pageNumber="61">
<taxonomicName id="47FED5DAFFE51B57FF28D027FEB2725B" baseAuthorityName="Vinciguerra" baseAuthorityYear="1890" box="[151,376,1303,1329]" class="Actinopterygii" family="Cyprinidae" genus="Opsarius" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="barnoides">Opsarius barnoides</taxonomicName>
,
<taxonomicName id="47FED5DAFFE51B57FE3CD027FD9E725B" baseAuthorityName="Day" baseAuthorityYear="1871" box="[387,596,1303,1329]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="stoliczkana">Pethia stoliczkana</taxonomicName>
,
</emphasis>
and
<emphasis id="B28A724BFFE51B57FD31D027FCE1725A" box="[654,811,1303,1328]" italics="true" pageId="7" pageNumber="61">
<taxonomicName id="47FED5DAFFE51B57FD31D027FCED725A" authorityName="Fang &amp; Kottelat" authorityYear="2000" box="[654,807,1303,1328]" class="Actinopterygii" family="Cyprinidae" genus="Danio" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="61" phylum="Chordata" rank="species" species="roseus">Danio roseus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>