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<mods:title id="8739E930826A446B2701F905FB1137DA">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="0544E3005CED1E27D594F6FE174D80D4">Dominiak, Patrycja</mods:namePart>
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<taxonomicName id="47B2F798FFF6FFCD42AB30B803C989D4" ID-CoL="8BQQR" authority="Remm" authorityName="Remm" box="[151,478,151,177]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="similaris">
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<emphasis id="B2C65009FFF6FFCD42AB30B8039489D4" bold="true" box="[151,387,151,177]" italics="true" pageId="8" pageNumber="141">Dasyhelea similaris</emphasis>
Remm
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<taxonomicName id="47B2F798FFF6FFCD42AB30F000078991" ID-CoL="8BQQR" authority="Remm, 1972: 77" authorityName="Remm" authorityPageNumber="77" authorityYear="1972" box="[151,528,223,245]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="similaris">
<emphasis id="B2C65009FFF6FFCD42AB30F003768990" box="[151,353,223,245]" italics="true" pageId="8" pageNumber="141">Dasyhelea similaris</emphasis>
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</taxonomicName>
(
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: Buryatia).
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</paragraph>
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<emphasis id="B2C65009FFF6FFCD42AB310F03E9885C" bold="true" box="[151,510,288,313]" pageId="8" pageNumber="141">
Material examined.
<collectingCountry id="F8A5CC8BFFF6FFCD43B1310E03EF885C" box="[397,504,289,313]" name="Lebanon" pageId="8" pageNumber="141">Lebanon</collectingCountry>
.
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Aarbit Kozhayya, St. Antonios Kozhayya Monaster, N 34°
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35°56.689,
<date id="F40CAADBFFF6FFCD42AB316A03188838" box="[151,271,325,349]" pageId="8" pageNumber="141" value="2012-10-28">28.X.2012</date>
, shallow cave, net,
<specimenCount id="96B44792FFF6FFCD43D7316A00468838" box="[491,593,325,349]" pageId="8" pageNumber="141" type="male">17 males</specimenCount>
, leg. P. Dominiak (LU &amp; UG).
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<paragraph id="800D8C1BFFF6FFCD42FB3147076488C0" blockId="8.[151,1436,288,421]" pageId="8" pageNumber="141">
<emphasis id="B2C65009FFF6FFCD42FB3147037788E4" bold="true" box="[199,352,360,385]" pageId="8" pageNumber="141">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFF6FFCD4356314603AF88E4" box="[362,440,361,385]" name="Poland" pageId="8" pageNumber="141">Poland</collectingCountry>
(
<bibRefCitation id="E423F1EAFFF6FFCD43F53146012088E4" author="Dominiak, P. &amp; Szadziewski, R." box="[457,823,361,385]" firstAuthor="Dominiak" journalOrPublisher="Dipteron" pageId="8" pageNumber="141" pagination="4 - 7" part="22" refId="ref6555" refString="Dominiak, P. &amp; Szadziewski, R. (2006) Biting midges of the genus Dasyhelea Kieffer, 1911 (Diptera: Ceratopogonidae) collected from Polish peat-bogs. Dipteron, 22, 4 - 7." title="Biting midges of the genus Dasyhelea Kieffer, 1911 (Diptera: Ceratopogonidae) collected from Polish peat-bogs" type="journal article" year="2006">Dominiak &amp; Szadziewski 2006</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFF6FFCD4171314601A588E4" box="[845,946,361,385]" name="Romania" pageId="8" pageNumber="141">Romania</collectingCountry>
(
<bibRefCitation id="E423F1EAFFF6FFCD41F83146072588E4" author="Dominiak, P. &amp; Szadziewski, R." box="[964,1330,361,385]" firstAuthor="Dominiak" journalOrPublisher="Zootaxa" pageId="8" pageNumber="141" pagination="1 - 37" part="2437" refId="ref6594" refString="Dominiak, P. &amp; Szadziewski, R. (2010) Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). Zootaxa, 2437, 1 - 37." title="Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2010">Dominiak &amp; Szadziewski 2010</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFF6FFCD47743146078B88E4" box="[1352,1436,361,385]" name="Estonia" pageId="8" pageNumber="141">Estonia</collectingCountry>
(
<bibRefCitation id="E423F1EAFFF6FFCD42A231A2033C88C0" author="Remm, H." box="[158,299,397,421]" firstAuthor="Remm" journalOrPublisher="Eesti NSV Teaduste Akadeemia Tartu" pageId="8" pageNumber="141" pagination="40 - 60" refId="ref7363" refString="Remm, H. (1979) A catalogue of the Ceratopogonidae (Diptera) of the Estonian S. S. R. Eesti NSV Teaduste Akadeemia Tartu, pp. 40 - 60." title="A catalogue of the Ceratopogonidae (Diptera) of the Estonian S. S. R." type="book chapter" year="1979">Remm 1979</bibRefCitation>
) and
<collectingCountry id="F8A5CC8BFFF6FFCD435531A203A388C0" box="[361,436,397,421]" name="Russia" pageId="8" pageNumber="141">Russia</collectingCountry>
(Buryatia) (
<bibRefCitation id="E423F1EAFFF6FFCD400131A200DB88C0" author="Remm, H." box="[573,716,397,421]" firstAuthor="Remm" journalOrPublisher="Tartu Riikliku Ulikodi Toimetised" pageId="8" pageNumber="141" pagination="62 - 90" part="293" refId="ref7333" refString="Remm, H. (1972) New species of Ceratopogonidae (Diptera) from the South Siberia. Tartu Riikliku Ulikodi Toimetised, 293, 62 - 90." title="New species of Ceratopogonidae (Diptera) from the South Siberia" type="journal article" year="1972">Remm 1972</bibRefCitation>
).
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<emphasis id="B2C65009FFF6FFCD40E331A201A988C0" box="[735,958,397,421]" italics="true" pageId="8" pageNumber="141">Dasyhelea similaris</emphasis>
</taxonomicName>
is new for
<collectingCountry id="F8A5CC8BFFF6FFCD460131A2068888C0" box="[1085,1183,397,421]" name="Lebanon" pageId="8" pageNumber="141">Lebanon</collectingCountry>
and the Near East.
</paragraph>
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</treatment>
</document>

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<mods:title id="1D29202DF3425A08EF18120D02942360">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<taxonomicName id="47B2F798FFF6FFCD42AB31D603F38B77" ID-CoL="8BQR2" authority="Kieffer" authorityName="Kieffer" box="[151,484,504,531]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="turficola">
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<emphasis id="B2C65009FFF6FFCD42AB31D603938B76" bold="true" box="[151,388,505,531]" italics="true" pageId="8" pageNumber="141">Dasyhelea turficola</emphasis>
Kieffer
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</paragraph>
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<taxonomicName id="47B2F798FFF6FFCD42AB326F00868B33" ID-CoL="8BQR2" authority="Kieffer, 1925a: 152" authorityName="Kieffer" authorityPageNumber="152" authorityYear="1925" box="[151,657,576,598]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="turficola">
<emphasis id="B2C65009FFF6FFCD42AB326F03488B33" box="[151,351,576,598]" italics="true" pageId="8" pageNumber="141">Dasyhelea turficola</emphasis>
<treatmentCitation id="0113AA0AFFF6FFCD4359326F003C8B33" author="Kieffer" box="[357,555,576,598]" page="152" pageId="8" pageNumber="141" year="1925">
<bibRefCitation id="E423F1EAFFF6FFCD4359326F003C8B33" author="Kieffer, J. J." box="[357,555,576,598]" firstAuthor="Kieffer" journalOrPublisher="Beitrage zur Kunde Estlands" pageId="8" pageNumber="141" pagination="145 - 163" part="10" refId="ref7040" refString="Kieffer, J. J. (1925 a) Chironomiden der Hochmoore Nordeuropas und des ostlichen Mitteleuropas. Beitrage zur Kunde Estlands, 10, 145 - 163." title="Chironomiden der Hochmoore Nordeuropas und des ostlichen Mitteleuropas" type="journal article" year="1925" yearSuffix="a">Kieffer, 1925a: 152</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F8A5CC8BFFF6FFCD4004326F009D8B33" box="[568,650,576,598]" name="Estonia" pageId="8" pageNumber="141">Estonia</collectingCountry>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF6FFCD42AB32AD03478B87" pageId="8" pageNumber="141" type="materials_examined">
<paragraph id="800D8C1BFFF6FFCD42AB32AD03478B87" blockId="8.[151,1436,642,846]" pageId="8" pageNumber="141">
<emphasis id="B2C65009FFF6FFCD42AB32AD03E78BFE" bold="true" box="[151,496,642,667]" pageId="8" pageNumber="141">
Material examined.
<collectingCountry id="F8A5CC8BFFF6FFCD43BA32AC03E78BFE" box="[390,496,643,667]" name="Lebanon" pageId="8" pageNumber="141">Lebanon</collectingCountry>
</emphasis>
. Tyre, N 33°
<geoCoordinate id="E586EADCFFF6FFCD404132AD00F38BFF" box="[637,740,642,666]" direction="east" orientation="longitude" pageId="8" pageNumber="141" precision="55" value="15.535">15.535 E</geoCoordinate>
35°12.726,
<date id="F40CAADBFFF6FFCD415032AD01F68BFF" box="[876,993,642,666]" pageId="8" pageNumber="141" value="2012-05-06">06.V.2012</date>
, beach, net,
<specimenCount id="96B44792FFF6FFCD465732AD06D48BFF" box="[1131,1219,642,666]" pageId="8" pageNumber="141" type="male">2 males</specimenCount>
, leg. A. Alwin &amp; P. Dominiak (LU &amp; UG). Tyre, Al-Bass ruins, N 33°
<geoCoordinate id="E586EADCFFF6FFCD40E8328901298BDB" box="[724,830,678,702]" direction="east" orientation="longitude" pageId="8" pageNumber="141" precision="55" value="16.232">16.232 E</geoCoordinate>
35°12.706, 19 m asl,
<date id="F40CAADBFFF6FFCD4607328906A78BDB" box="[1083,1200,678,702]" pageId="8" pageNumber="141" value="2012-05-12">12.V.2012</date>
, net,
<specimenCount id="96B44792FFF6FFCD46D33289075D8BDB" box="[1263,1354,678,702]" pageId="8" pageNumber="141" type="male">2 males</specimenCount>
, leg. P. Dominiak (UG).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF6FFCD42FB32C103598A2B" pageId="8" pageNumber="141" type="distribution">
<paragraph id="800D8C1BFFF6FFCD42FB32C103598A2B" blockId="8.[151,1436,642,846]" pageId="8" pageNumber="141">
<emphasis id="B2C65009FFF6FFCD42FB32C103778A62" bold="true" box="[199,352,750,775]" pageId="8" pageNumber="141">Distribution.</emphasis>
<taxonomicName id="47B2F798FFF6FFCD435632C1005C8A63" box="[362,587,750,774]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="141" phylum="Arthropoda" rank="species" species="turficola">
<emphasis id="B2C65009FFF6FFCD435632C1005C8A63" box="[362,587,750,774]" italics="true" pageId="8" pageNumber="141">Dasyhelea turficola</emphasis>
</taxonomicName>
is widely distributed in the Palaearctic region (
<bibRefCitation id="E423F1EAFFF6FFCD464832C102C48A4F" author="Dominiak, P. &amp; Szadziewski, R." firstAuthor="Dominiak" journalOrPublisher="Zootaxa" pageId="8" pageNumber="141" pagination="1 - 37" part="2437" refId="ref6594" refString="Dominiak, P. &amp; Szadziewski, R. (2010) Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). Zootaxa, 2437, 1 - 37." title="Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2010">Dominiak &amp; Szadziewski 2010</bibRefCitation>
), but from the Near East previously known only from
<collectingCountry id="F8A5CC8BFFF6FFCD416E333D01878A4F" box="[850,912,786,810]" name="Israel" pageId="8" pageNumber="141">Israel</collectingCountry>
(
<bibRefCitation id="E423F1EAFFF6FFCD419D333D06B78A4F" author="Dominiak, P. &amp; Braverman, Y. &amp; Szadziewski, R." box="[929,1184,786,811]" etAl="et al." firstAuthor="Dominiak" journalOrPublisher="Polish Journal of Entomology" pageId="8" pageNumber="141" pagination="257 - 265" part="76" refId="ref6499" refString="Dominiak, P., Braverman, Y. &amp; Szadziewski, R. (2007) New records of biting midges of the genus Dasyhelea from Israel, with notes on synonymy in the genera Culicoides and Forcipomyia (Diptera: Ceratopogonidae). Polish Journal of Entomology, 76, 257 - 265." title="New records of biting midges of the genus Dasyhelea from Israel, with notes on synonymy in the genera Culicoides and Forcipomyia (Diptera: Ceratopogonidae)" type="journal article" year="2007">
Dominiak
<emphasis id="B2C65009FFF6FFCD4621333B06458A4E" box="[1053,1106,787,811]" italics="true" pageId="8" pageNumber="141">et al</emphasis>
. 2007
</bibRefCitation>
). New species for the Lebanese fauna.
</paragraph>
</subSubSection>
</treatment>
</document>

147
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<mods:title id="751342193E310C9D9532F77D9E0109BC">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<treatment id="081B3D0DFFF8FFC242AB365207598B00" ID-DOI="http://doi.org/10.5281/zenodo.4500030" ID-GBIF-Taxon="119571716" ID-Zenodo-Dep="4500030" LSID="urn:lsid:zoobank.org:act:C4645ABA-0CD2-4F4B-9B66-990525A32A30" httpUri="http://treatment.plazi.org/id/081B3D0DFFF8FFC242AB365207598B00" lastPageId="7" lastPageNumber="140" pageId="6" pageNumber="139">
<subSubSection id="C8A8DF90FFF8FFC342AB365203318FDD" pageId="6" pageNumber="139" type="nomenclature">
<paragraph id="800D8C1BFFF8FFC342AB365200A78FF2" blockId="6.[151,688,1661,1720]" box="[151,688,1661,1687]" pageId="6" pageNumber="139">
<heading id="DB453B77FFF8FFC342AB365200A78FF2" bold="true" box="[151,688,1661,1687]" fontSize="11" level="1" pageId="6" pageNumber="139" reason="1">
<emphasis id="B2C65009FFF8FFC342AB365200A78FF2" bold="true" box="[151,688,1661,1687]" pageId="6" pageNumber="139">
<taxonomicName id="47B2F798FFF8FFC342AB3652005D8FF2" ID-CoL="9HN7N" authority="Dominiak" authorityName="Dominiak" box="[151,586,1661,1687]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="sandrageorgei" status="sp. nov.">
<emphasis id="B2C65009FFF8FFC342AB365203DF8FF2" bold="true" box="[151,456,1661,1687]" italics="true" pageId="6" pageNumber="139">Dasyhelea sandrageorgei</emphasis>
Dominiak
</taxonomicName>
<taxonomicNameLabel id="A9F5ED72FFF8FFC3406E365200A78FF2" box="[594,688,1661,1687]" pageId="6" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="800D8C1BFFF8FFC342AB368F03318FDD" blockId="6.[151,688,1661,1720]" box="[151,294,1696,1720]" pageId="6" pageNumber="139">
(
<figureCitation id="1889909EFFF8FFC342A3368F03098FDD" box="[159,286,1696,1720]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Figs 13–15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342AB36C7005C8E41" pageId="6" pageNumber="139" type="materials_examined">
<paragraph id="800D8C1BFFF8FFC342AB36C7005C8E41" blockId="6.[151,1437,1767,2013]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342AB36C7035C8E65" bold="true" box="[151,331,1767,1792]" pageId="6" pageNumber="139">
<typeStatus id="5F0932B9FFF8FFC342AB36C702C58E65" box="[151,210,1768,1792]" pageId="6" pageNumber="139">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5F0932B9FFF8FFC3436436C703D28E65" box="[344,453,1768,1792]" pageId="6" pageNumber="139" type="holotype">Holotype</typeStatus>
: adult male.
<emphasis id="B2C65009FFF8FFC3405A36C700C08E65" bold="true" box="[614,727,1768,1792]" pageId="6" pageNumber="139">
<collectingCountry id="F8A5CC8BFFF8FFC3405A36C700C58E65" box="[614,722,1768,1792]" name="Lebanon" pageId="6" pageNumber="139">Lebanon</collectingCountry>
.
</emphasis>
Tyre, Al-Bass ruins, N 33°
<geoCoordinate id="E586EADCFFF8FFC3461936C706848E65" box="[1061,1171,1768,1792]" direction="east" orientation="longitude" pageId="6" pageNumber="139" precision="55" value="16.232">16.232 E</geoCoordinate>
35°12.706, 19 m asl,
<date id="F40CAADBFFF8FFC342AB3723031C8E41" box="[151,267,1804,1828]" pageId="6" pageNumber="139" value="2012-05-12">12.V.2012</date>
, net, leg. P. Dominiak (UG).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB370006D18E09" pageId="6" pageNumber="139" type="diagnosis">
<paragraph id="800D8C1BFFF8FFC342FB370006D18E09" blockId="6.[151,1437,1767,2013]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB370003568E2D" bold="true" box="[199,321,1839,1864]" pageId="6" pageNumber="139">Diagnosis.</emphasis>
This species has a pair of slightly convergent aedeagal processes, which are membranous in their lateral parts, and slender posterior process of right paramere, distinctly bent in its distal portion.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC242AB375801D5889C" lastPageId="7" lastPageNumber="140" pageId="6" pageNumber="139" type="description">
<paragraph id="800D8C1BFFF8FFC342AB3758078A8EBD" blockId="6.[151,1437,1767,2013]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342AB375803638EF5" bold="true" box="[151,372,1911,1936]" pageId="6" pageNumber="139">Description. Male.</emphasis>
Head. Flagellum length
<quantity id="474A21FEFFF8FFC340B1375700E48EF5" box="[653,755,1912,1936]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="6" pageNumber="139" unit="mm" value="0.56">0.56 mm</quantity>
, AR 0.86. Distal flagellomeres with two rows of long setae, flagellomere 13 without apical prolongation (
<figureCitation id="1889909EFFF8FFC3409137B301118ED1" box="[685,774,1948,1972]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 13</figureCitation>
). Clypeus composed of two separate sclerites, elongate, bearing 9 strong setae. Maxillary palp (
<figureCitation id="1889909EFFF8FFC3406F37EF00B28EBD" box="[595,677,1984,2008]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 14</figureCitation>
) with third palpal segment slender, 62 μm long, PR III 5.17. Thorax.
</paragraph>
<paragraph id="800D8C1BFFF9FFC242AB30B7017688E8" blockId="7.[151,1437,151,613]" pageId="7" pageNumber="140">
Transverse suture absent. Wing membrane hyaline, covered with fine macrotrichia. Only one radial cell present. Wing length
<quantity id="474A21FEFFF9FFC243173093038389B1" box="[299,404,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="140" unit="mm" value="0.76">0.76 mm</quantity>
, CR 0.41. Scutellum yellow, with 6 stout setae. Legs uniformly colored,
<collectingCountry id="F8A5CC8BFFF9FFC246EB309206EC89B1" box="[1239,1275,189,212]" name="Chile" pageId="7" pageNumber="140">TR</collectingCountry>
I 2.3,
<collectingCountry id="F8A5CC8BFFF9FFC247073092074889B1" box="[1339,1375,189,212]" name="Chile" pageId="7" pageNumber="140">TR</collectingCountry>
II 2.4,
<collectingCountry id="F8A5CC8BFFF9FFC242AB30CC02AC899F" box="[151,187,227,250]" name="Chile" pageId="7" pageNumber="140">TR</collectingCountry>
III 2.0. Genitalia (
<figureCitation id="1889909EFFF9FFC2434430CD03DB899F" box="[376,460,226,250]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="7" pageNumber="140">Fig. 15</figureCitation>
). Tergite 9 tapered distally, bearing two cornet-shaped apicolateral processes. Posterior margin of sternite 9 extended, conical, fused to aedeagus by conspicuous bridge. Gonostylus evenly arched, with short, ventral, finger-like projection in basal part. Parameres asymmetrical and fused. Posterior process broadly fused to right basal arm, long, slender and distinctly bent apically. Submedian projections of aedeagus slightly convergent, with recurved tips and membranous lateral borders.
</paragraph>
<paragraph id="800D8C1BFFF9FFC242FB31B5004988D4" blockId="7.[151,1437,151,613]" box="[199,606,409,434]" pageId="7" pageNumber="140">
<emphasis id="B2C65009FFF9FFC242FB31B503F688D7" bold="true" box="[199,481,409,434]" pageId="7" pageNumber="140">Female and immatures.</emphasis>
Unknown.
</paragraph>
<paragraph id="800D8C1BFFF9FFC242FB319201D5889C" blockId="7.[151,1437,151,613]" pageId="7" pageNumber="140">
<emphasis id="B2C65009FFF9FFC242FB319203F388B3" bold="true" box="[199,484,445,470]" pageId="7" pageNumber="140">Derivation of the name.</emphasis>
The species is named after Sandra Abou Najm and George Kachacha, who deserve special thanks for their kind assistance during field research in
<collectingCountry id="F8A5CC8BFFF9FFC2416531CE01A8889C" box="[857,959,481,505]" name="Lebanon" pageId="7" pageNumber="140">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF9FFC242FB322A03C68B78" box="[199,465,517,542]" pageId="7" pageNumber="140" type="distribution">
<paragraph id="800D8C1BFFF9FFC242FB322A03C68B78" blockId="7.[151,1437,151,613]" box="[199,465,517,542]" pageId="7" pageNumber="140">
<emphasis id="B2C65009FFF9FFC242FB322A03778B7B" bold="true" box="[199,352,517,542]" pageId="7" pageNumber="140">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFF9FFC2435B322A03DA8B78" box="[359,461,517,541]" name="Lebanon" pageId="7" pageNumber="140">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF9FFC242FB320607598B00" pageId="7" pageNumber="140" type="discussion">
<paragraph id="800D8C1BFFF9FFC242FB320607598B00" blockId="7.[151,1437,151,613]" pageId="7" pageNumber="140">
<emphasis id="B2C65009FFF9FFC242FB320603588B27" bold="true" box="[199,335,553,578]" pageId="7" pageNumber="140">Comments.</emphasis>
<taxonomicName id="47B2F798FFF9FFC2436B3206006F8B24" box="[343,632,553,577]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="140" phylum="Arthropoda" rank="species" species="sandrageorgei">
<emphasis id="B2C65009FFF9FFC2436B3206006F8B24" box="[343,632,553,577]" italics="true" pageId="7" pageNumber="140">Dasyhelea sandrageorgei</emphasis>
</taxonomicName>
is close to
<taxonomicName id="47B2F798FFF9FFC240C0320506338B24" authority="Remm. The" authorityName="Remm. The" box="[764,1060,553,577]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="140" phylum="Arthropoda" rank="species" species="leptoclada">
<emphasis id="B2C65009FFF9FFC240C0320501078B24" box="[764,784,554,577]" italics="true" pageId="7" pageNumber="140">D</emphasis>
.
<emphasis id="B2C65009FFF9FFC2411C320601818B24" box="[800,918,553,577]" italics="true" pageId="7" pageNumber="140">leptoclada</emphasis>
Remm. The
</taxonomicName>
two species slightly differ in the shape of aedeagus and in the length of the bent part of paramere that is markedly shorter in the new species.
</paragraph>
</subSubSection>
</treatment>
</document>

158
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<mods:title id="F38CD818AF070850D6E36C12952E503F">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<treatment id="081B3D0DFFF8FFC342AB310800108DC1" ID-DOI="http://doi.org/10.5281/zenodo.4500016" ID-GBIF-Taxon="119571715" ID-Zenodo-Dep="4500016" LSID="urn:lsid:zoobank.org:act:2733A340-8ADE-46EF-AC4B-7504DC249E61" httpUri="http://treatment.plazi.org/id/081B3D0DFFF8FFC342AB310800108DC1" lastPageNumber="139" pageId="6" pageNumber="139">
<subSubSection id="C8A8DF90FFF8FFC342AB3108030E8807" pageId="6" pageNumber="139" type="nomenclature">
<paragraph id="800D8C1BFFF8FFC342AB3108005E8824" blockId="6.[151,585,295,354]" box="[151,585,295,321]" pageId="6" pageNumber="139">
<heading id="DB453B77FFF8FFC342AB3108005E8824" bold="true" box="[151,585,295,321]" fontSize="11" level="1" pageId="6" pageNumber="139" reason="1">
<emphasis id="B2C65009FFF8FFC342AB3108005E8824" bold="true" box="[151,585,295,321]" pageId="6" pageNumber="139">
<taxonomicName id="47B2F798FFF8FFC342AB310803F38824" ID-CoL="9HN7L" authority="Dominiak" authorityName="Dominiak" box="[151,484,295,321]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="nauta" status="sp. nov.">
<emphasis id="B2C65009FFF8FFC342AB310803778824" bold="true" box="[151,352,295,321]" italics="true" pageId="6" pageNumber="139">Dasyhelea nauta</emphasis>
Dominiak
</taxonomicName>
<taxonomicNameLabel id="A9F5ED72FFF8FFC343D73108005E8824" box="[491,585,295,321]" pageId="6" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="800D8C1BFFF8FFC342AB3165030E8807" blockId="6.[151,585,295,354]" box="[151,281,330,354]" pageId="6" pageNumber="139">
(
<figureCitation id="1889909EFFF8FFC342A3316503068807" box="[159,273,330,354]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Figs 9–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342AB31BC03B38897" pageId="6" pageNumber="139" type="materials_examined">
<paragraph id="800D8C1BFFF8FFC342AB31BC03B38897" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342AB31BC035288CE" bold="true" box="[151,325,402,427]" pageId="6" pageNumber="139">
<typeStatus id="5F0932B9FFF8FFC342AB31BC02C788CE" box="[151,208,403,427]" pageId="6" pageNumber="139">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5F0932B9FFF8FFC3437331BD03AB88CF" box="[335,444,402,426]" pageId="6" pageNumber="139" type="holotype">Holotype</typeStatus>
: adult male.
<emphasis id="B2C65009FFF8FFC3406D31BC00B188CE" bold="true" box="[593,678,403,427]" pageId="6" pageNumber="139">Ye m e n.</emphasis>
Al Mukalla, N 14˚33' E 49˚08',
<date id="F40CAADBFFF8FFC3461D31BD06DE88CF" box="[1057,1225,402,426]" pageId="6" pageNumber="139" value="2003-07" valueMax="2003-08-31" valueMin="2003-07-00">VII–VIII.2003</date>
, light trap, leg. A. van Harten (UG).
<typeStatus id="5F0932B9FFF8FFC3435D319803C588AA" box="[353,466,439,463]" pageId="6" pageNumber="139" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B2C65009FFF8FFC343E03198002688AA" bold="true" box="[476,561,439,463]" pageId="6" pageNumber="139">
<collectingCountry id="F8A5CC8BFFF8FFC343E03198003D88AA" box="[476,554,439,463]" name="Yemen" pageId="6" pageNumber="139">Yemen</collectingCountry>
.
</emphasis>
Al Mukalla, N 14˚33' E 49˚08',
<date id="F40CAADBFFF8FFC341A73199065588AB" box="[923,1090,438,462]" pageId="6" pageNumber="139" value="2003-07" valueMax="2003-08-31" valueMin="2003-07-00">VII–VIII.2003</date>
, light trap,
<specimenCount id="96B44792FFF8FFC346833199070088AB" box="[1215,1303,438,462]" pageId="6" pageNumber="139" type="male">3 males</specimenCount>
, leg. A. van Harten (
<collectionCode id="E6A314DEFFF8FFC342CD31F503508897" box="[241,327,474,498]" pageId="6" pageNumber="139">BMNH</collectionCode>
&amp; UG).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB31D101EC8B5F" pageId="6" pageNumber="139" type="diagnosis">
<paragraph id="800D8C1BFFF8FFC342FB31D101EC8B5F" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB31D103568B72" bold="true" box="[199,321,510,535]" pageId="6" pageNumber="139">Diagnosis.</emphasis>
Adult male of
<taxonomicName id="47B2F798FFF8FFC3403C31D100D48B72" box="[512,707,510,535]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="nauta">
<emphasis id="B2C65009FFF8FFC3403C31D100D48B72" box="[512,707,510,535]" italics="true" pageId="6" pageNumber="139">Dasyhelea nauta</emphasis>
</taxonomicName>
has unique eight-shaped median projections of aedeagus and symmetrical parameres. It is also distinct in having short distal flagellomeres.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB326906078D71" pageId="6" pageNumber="139" type="description">
<paragraph id="800D8C1BFFF8FFC342FB326902CF8ACD" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB326903B18B3A" bold="true" box="[199,422,582,607]" pageId="6" pageNumber="139">Description. Male.</emphasis>
Head. Flagellum length
<quantity id="474A21FEFFF8FFC340F6326901658B3B" box="[714,882,582,606]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="6" pageNumber="139" unit="mm" value="0.48" valueMax="0.51" valueMin="0.45">0.45–0.51 mm</quantity>
, AR 0.58–0.66 (n=4). Four distal flagellomeres short, with one row of long setae (
<figureCitation id="1889909EFFF8FFC3402F324500408BE7" box="[531,599,618,642]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 9</figureCitation>
). Last flagellomere without apical prolongation. Clypeus composed of two sclerites, with 5–6 strong setae. Maxillary palp (
<figureCitation id="1889909EFFF8FFC3409232A100E98BC3" box="[686,766,654,678]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 10</figureCitation>
) with third palpal segment cylindrical, 43–54 μm long, PR III 2.74–3.19 (n=4). Thorax. Transverse suture absent, number of supraalar setae vary from 6 to 7. Scutellum dark, with 6 strong setae. Wing with one nearly obliterated radial cell. Wing length
<quantity id="474A21FEFFF8FFC3463D32F606B18B94" box="[1025,1190,729,753]" metricMagnitude="-4" metricUnit="m" metricValue="8.3" metricValueMax="8.6" metricValueMin="8.0" pageId="6" pageNumber="139" unit="mm" value="0.83" valueMax="0.86" valueMin="0.8">0.80–0.86 mm</quantity>
, CR 0.48–0.50 (n=4). Legs uniform in coloration,
<collectingCountry id="F8A5CC8BFFF8FFC343D732D100198A70" box="[491,526,766,789]" name="Chile" pageId="6" pageNumber="139">TR</collectingCountry>
I 2.5–2.6 (n=4),
<collectingCountry id="F8A5CC8BFFF8FFC340E932D100EE8A70" box="[725,761,766,789]" name="Chile" pageId="6" pageNumber="139">TR</collectingCountry>
II 2.6–3.0 (n=4),
<collectingCountry id="F8A5CC8BFFF8FFC341F932D101FE8A70" box="[965,1001,766,789]" name="Chile" pageId="6" pageNumber="139">TR</collectingCountry>
III 2.3–2.4 (n=4). Genitalia (
<figureCitation id="1889909EFFF8FFC3470A32D207998A70" box="[1334,1422,765,789]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 11</figureCitation>
). Posterior margin of tergite 9 rounded, with a pair of short apicolateral processes. Sternite 9 slightly expanded. Gonostylus long and slender, gently arched. Parameres symmetrical, stout basally and very thin in apical part, fused (
<figureCitation id="1889909EFFF8FFC342DA3343032A8AE1" box="[230,317,876,900]" captionStart="FIGURES 9 – 15" captionStartId="7.[151,263,1941,1964]" captionTargetBox="[297,1288,685,1900]" captionTargetId="figure@7.[280,1307,666,1920]" captionTargetPageId="7" captionText="FIGURES 9 – 15. Dasyhelea nauta Dominiak sp. nov. (9 – 12), holotype male: 9. flagellomeres 9 – 13, 10. palpus, 11. ventral view of genitalia, 12. parameres; Dasyhelea sandrageorgei Dominiak sp. nov. (13 – 15), holotype male: 13. distal flagellomeres, 14. palpus, 15. ventral view of genitalia." httpUri="https://zenodo.org/record/219727/files/figure.png" pageId="6" pageNumber="139">Fig. 12</figureCitation>
). Median projections of aedeagus strongly sclerotized, eight-shaped, with short and sharp apical tooth.
</paragraph>
<paragraph id="800D8C1BFFF8FFC342FB339A00498AA9" blockId="6.[151,1437,402,1188]" box="[199,606,948,973]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB339A03F68AA8" bold="true" box="[199,481,948,973]" pageId="6" pageNumber="139">Female and immatures.</emphasis>
Unknown.
</paragraph>
<paragraph id="800D8C1BFFF8FFC342FB33F806078D71" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB33F803F58A95" bold="true" box="[199,482,983,1008]" pageId="6" pageNumber="139">Derivation of the name.</emphasis>
The Latin word
<taxonomicName id="47B2F798FFF8FFC3409C33F600F78A95" box="[672,736,985,1008]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="species" species="nauta">
<emphasis id="B2C65009FFF8FFC3409C33F600F78A95" box="[672,736,985,1008]" italics="true" pageId="6" pageNumber="139">nauta</emphasis>
</taxonomicName>
,-
<emphasis id="B2C65009FFF8FFC340CC33F6011C8A95" box="[752,779,985,1008]" italics="true" pageId="6" pageNumber="139">ae</emphasis>
means sailor or shipowner. The species is named after the
<typeStatus id="5F0932B9FFF8FFC342AB33D202D08D70" box="[151,199,1021,1045]" pageId="6" pageNumber="139">type</typeStatus>
locality, Al Mukalla, an important sea port in the province of Hadhramaut.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB343003AE8D5D" box="[199,441,1055,1080]" pageId="6" pageNumber="139" type="distribution">
<paragraph id="800D8C1BFFF8FFC342FB343003AE8D5D" blockId="6.[151,1437,402,1188]" box="[199,441,1055,1080]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB343003778D5D" bold="true" box="[199,352,1055,1080]" pageId="6" pageNumber="139">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFF8FFC3435B340E03A58D5D" box="[359,434,1057,1080]" name="Yemen" pageId="6" pageNumber="139">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFF8FFC342FB346B00108DC1" pageId="6" pageNumber="139" type="discussion">
<paragraph id="800D8C1BFFF8FFC342FB346B00108DC1" blockId="6.[151,1437,402,1188]" pageId="6" pageNumber="139">
<emphasis id="B2C65009FFF8FFC342FB346B03588D38" bold="true" box="[199,335,1092,1117]" pageId="6" pageNumber="139">Comments.</emphasis>
The characteristic armature of male genitalia, including the shape of both aedeagus and parameres, distinguishes this species from all other
<taxonomicName id="47B2F798FFF8FFC3413A3447016B8DE5" box="[774,892,1128,1152]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="B2C65009FFF8FFC3413A3447016B8DE5" box="[774,892,1128,1152]" italics="true" pageId="6" pageNumber="139">Dasyhelea</emphasis>
</taxonomicName>
. The built of antenna, with very short distal flagellomeres, is also distinctive.
</paragraph>
</subSubSection>
</treatment>
</document>

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(
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Material examined.
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.
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Tyre, N 33°
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35°12.726,
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, beach, net,
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, leg. A. Alwin &amp; P. Dominiak (LU &amp; UG).
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<emphasis id="B2C65009FFF8FFC342FB35E603778C87" bold="true" box="[199,352,1481,1506]" pageId="6" pageNumber="139">Distribution.</emphasis>
A distribution of this species ranges from
<collectingCountry id="F8A5CC8BFFF8FFC3417035E6019B8C84" box="[844,908,1481,1505]" name="Spain" pageId="6" pageNumber="139">Spain</collectingCountry>
to
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(
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). In the region of the Near East it was mentioned from
<collectingCountry id="F8A5CC8BFFF8FFC340E735C2010E8F60" box="[731,793,1517,1541]" name="Israel" pageId="6" pageNumber="139">Israel</collectingCountry>
(
<bibRefCitation id="E423F1EAFFF8FFC3411435C206098F60" author="Dominiak, P. &amp; Braverman, Y. &amp; Szadziewski, R." box="[808,1054,1517,1542]" etAl="et al." firstAuthor="Dominiak" journalOrPublisher="Polish Journal of Entomology" pageId="6" pageNumber="139" pagination="257 - 265" part="76" refId="ref6499" refString="Dominiak, P., Braverman, Y. &amp; Szadziewski, R. (2007) New records of biting midges of the genus Dasyhelea from Israel, with notes on synonymy in the genera Culicoides and Forcipomyia (Diptera: Ceratopogonidae). Polish Journal of Entomology, 76, 257 - 265." title="New records of biting midges of the genus Dasyhelea from Israel, with notes on synonymy in the genera Culicoides and Forcipomyia (Diptera: Ceratopogonidae)" type="journal article" year="2007">
Dominiak
<emphasis id="B2C65009FFF8FFC3419E35C001C38F63" box="[930,980,1518,1542]" italics="true" pageId="6" pageNumber="139">et al</emphasis>
. 2007
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) and
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(
<bibRefCitation id="E423F1EAFFF8FFC3468235C207878F60" author="Szadziewski, R." box="[1214,1424,1517,1541]" firstAuthor="Szadziewski" journalOrPublisher="Polish Journal of Entomology" pageId="6" pageNumber="139" pagination="79 - 98" part="55" refId="ref7434" refString="Szadziewski, R. (1985) A review of the Palaearctic Dasyhelea (Pseudoculicoides) species of the johannseni group, with a description of two new species (Diptera, Ceratopogonidae). Polish Journal of Entomology, 55, 79 - 98." title="A review of the Palaearctic Dasyhelea (Pseudoculicoides) species of the johannseni group, with a description of two new species (Diptera, Ceratopogonidae)" type="journal article" year="1985">Szadziewski 1985</bibRefCitation>
). The present record is first for
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.
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<emphasis id="B2C65009FFFAFFC142AB37B703728ECB" box="[151,357,1944,1966]" italics="true" pageId="4" pageNumber="137">Dasyhelea deemingi</emphasis>
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<bibRefCitation id="E423F1EAFFFAFFC1435737B600B68ECB" author="Boorman, J. &amp; van Harten, A." box="[363,673,1945,1966]" firstAuthor="Boorman" journalOrPublisher="Fauna of Arabia" pageId="4" pageNumber="137" pagination="427 - 462" part="19" refId="ref6017" refString="Boorman, J. &amp; van Harten, A. (2002) Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen. Fauna of Arabia, 19, 427 - 462." title="Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen" type="journal article" year="2002">Boorman &amp; Harten, 2002: 441</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F8A5CC8BFFFAFFC1409337B600E58ECA" box="[687,754,1945,1967]" name="Oman" pageId="4" pageNumber="137">Oman</collectingCountry>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="800D8C1BFFFAFFC142AB379801768EA9" blockId="4.[151,865,1944,1997]" box="[151,865,1974,1997]" pageId="4" pageNumber="137">
<taxonomicName id="47B2F798FFFAFFC142AB379800B38EA9" ID-CoL="9HSFG" authority="Mazumdar &amp; Chaudhuri, 2009" authorityName="Mazumdar &amp; Chaudhuri" authorityYear="2009" box="[151,676,1975,1997]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFAFFC142AB379803758EA8" box="[151,354,1975,1997]" italics="true" pageId="4" pageNumber="137">Dasyhelea labinoda</emphasis>
<bibRefCitation id="E423F1EAFFFAFFC14354379800B38EA9" author="Mazumdar, A. &amp; Chaudhuri, P. K." box="[360,676,1975,1997]" firstAuthor="Mazumdar" journalOrPublisher="International Journal of Dipterological Research" pageId="4" pageNumber="137" pagination="191 - 201" part="20" refId="ref7183" refString="Mazumdar, A. &amp; Chaudhuri, P. K. (2009) Indian species of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). International Journal of Dipterological Research, 20, 191 - 201." title="Indian species of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2009">Mazumdar &amp; Chaudhuri, 2009</bibRefCitation>
</taxonomicName>
: (
<collectingCountry id="F8A5CC8BFFFAFFC14089379800F98EA8" box="[693,750,1975,1997]" name="India" pageId="4" pageNumber="137">India</collectingCountry>
).
<emphasis id="B2C65009FFFAFFC140C3379901768EA9" bold="true" box="[767,865,1974,1997]" pageId="4" pageNumber="137">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042AB30B801D38879" pageId="5" pageNumber="138" type="materials_examined">
<paragraph id="800D8C1BFFFBFFC042AB30B8012E89B1" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042AB30B803C989D5" bold="true" box="[151,478,151,176]" pageId="5" pageNumber="138">
Material examined.
<collectingCountry id="F8A5CC8BFFFBFFC043BA30B703CE89D5" box="[390,473,152,176]" name="Yemen" pageId="5" pageNumber="138">Yemen</collectingCountry>
.
</emphasis>
Al Mukalla, N 14˚33' E 49˚08',
<date id="F40CAADBFFFBFFC0417830B801A789CA" box="[836,944,151,175]" pageId="5" pageNumber="138" value="2003-01" valueMax="2003-02-31" valueMin="2003-01-00">
I–
<bibRefCitation id="E423F1EAFFFBFFC0416130B801A789CA" author="II" box="[861,944,151,175]" pageId="5" pageNumber="138" refString="Szadziewski, R. &amp; Borkent, A. (2003) New synonyms, combinations and records of biting midges (Diptera: Ceratopogonidae). Polish Journal of Entomology, 72, 249 - 260." type="journal article" year="2003">II.2003</bibRefCitation>
</date>
, light trap,
<specimenCount id="96B44792FFFBFFC0461230B8066C89CA" box="[1070,1147,151,175]" pageId="5" pageNumber="138" type="male">1 male</specimenCount>
, leg. A. van Harten (UG);
<date id="F40CAADBFFFBFFC042AB3093032989B1" box="[151,318,188,212]" pageId="5" pageNumber="138" value="2003-07" valueMax="2003-08-31" valueMin="2003-07-00">VII–VIII.2003</date>
, light trap,
<specimenCount id="96B44792FFFBFFC043813093000189B1" box="[445,534,188,212]" pageId="5" pageNumber="138" type="male">5 males</specimenCount>
, leg. A. van Harten (UG).
</paragraph>
<paragraph id="800D8C1BFFFBFFC042FB30CF01D38879" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB30CF0362899D" bold="true" box="[199,373,223,248]" pageId="5" pageNumber="138">
<typeStatus id="5F0932B9FFFBFFC042FB30CF0315899D" box="[199,258,224,248]" pageId="5" pageNumber="138">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5F0932B9FFFBFFC0434130CF03C8899D" box="[381,479,224,248]" pageId="5" pageNumber="138" type="paratype">Paratype</typeStatus>
male of
<taxonomicName id="47B2F798FFFBFFC0407F30CE00DD899D" box="[579,714,224,248]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFBFFC0407F30CE0040899D" box="[579,599,225,248]" italics="true" pageId="5" pageNumber="138">D</emphasis>
.
<emphasis id="B2C65009FFFBFFC0405A30CF00DD899D" box="[614,714,224,248]" italics="true" pageId="5" pageNumber="138">labinoda</emphasis>
</taxonomicName>
, labeled as follows: Para Adult 3, Loc. Burdwan, West Bengal,
<date id="F40CAADBFFFBFFC042AB312B03068879" box="[151,273,260,284]" pageId="5" pageNumber="138" value="1989-11-02">02.xi.1989</date>
, Coll. N. Saha,
<taxonomicName id="47B2F798FFFBFFC043FD312B01D38879" authority="Mazumdar &amp; Chaudhuri." authorityName="Mazumdar &amp; Chaudhuri." box="[449,964,260,284]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFBFFC043FD312B00B58879" box="[449,674,260,284]" italics="true" pageId="5" pageNumber="138">Dasyhelea labinoda</emphasis>
Mazumdar &amp; Chaudhuri.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042FB3108037488E2" pageId="5" pageNumber="138" type="distribution">
<paragraph id="800D8C1BFFFBFFC042FB3108037488E2" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB310803778825" bold="true" box="[199,352,295,320]" pageId="5" pageNumber="138">Distribution.</emphasis>
Up to now this species was mentioned from
<collectingCountry id="F8A5CC8BFFFBFFC0414B31080607885A" box="[887,1040,295,319]" name="Saudi Arabia" pageId="5" pageNumber="138">Saudi Arabia</collectingCountry>
,
<collectingCountry id="F8A5CC8BFFFBFFC046233108066C885A" box="[1055,1147,295,319]" name="Bahrain" pageId="5" pageNumber="138">Bahrain</collectingCountry>
,
<collectingCountry id="F8A5CC8BFFFBFFC046B5310806D9885A" box="[1161,1230,295,319]" name="Oman" pageId="5" pageNumber="138">Oman</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFBFFC046E3310803318801" author="Boorman, J. &amp; van Harten, A." firstAuthor="Boorman" journalOrPublisher="Fauna of Arabia" pageId="5" pageNumber="138" pagination="427 - 462" part="19" refId="ref6017" refString="Boorman, J. &amp; van Harten, A. (2002) Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen. Fauna of Arabia, 19, 427 - 462." title="Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen" type="journal article" year="2002">Boorman &amp; van Harten 2002</bibRefCitation>
), the
<collectingCountry id="F8A5CC8BFFFBFFC0435F3163004E8801" box="[355,601,332,356]" name="United Arab Emirates" pageId="5" pageNumber="138">United Arab Emirates</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFBFFC04054316301638801" author="Szadziewski, R. &amp; Gwizdalska-Kentzer, M. &amp; Gilka, W." box="[616,884,332,356]" editor="van Harten, A." etAl="et al." firstAuthor="Szadziewski" journalOrPublisher="Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE" pageId="5" pageNumber="138" pagination="636 - 653" refId="ref7546" refString="Szadziewski, R., Gwizdalska-Kentzer, M. &amp; Gilka, W. (2011) Order Diptera, family Ceratopogonidae. In: Sheikh Tahnoon Bin Zayed Al Nahyan, H. H. (patron), van Harten, A. (Ed.), Arthropod fauna of the United Arab Emirates. Volume 4. Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE, pp. 636 - 653." title="Order Diptera, family Ceratopogonidae" type="book chapter" volumeTitle="Arthropod fauna of the United Arab Emirates. Volume 4" year="2011">
Szadziewski
<emphasis id="B2C65009FFFBFFC040C0316201398801" box="[764,814,332,356]" italics="true" pageId="5" pageNumber="138">et al</emphasis>
. 2011
</bibRefCitation>
) and
<collectingCountry id="F8A5CC8BFFFBFFC0418F316301FA8801" box="[947,1005,332,356]" name="India" pageId="5" pageNumber="138">India</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFBFFC041C7316307438801" author="Mazumdar, A. &amp; Chaudhuri, P. K." box="[1019,1364,332,356]" firstAuthor="Mazumdar" journalOrPublisher="International Journal of Dipterological Research" pageId="5" pageNumber="138" pagination="191 - 201" part="20" refId="ref7183" refString="Mazumdar, A. &amp; Chaudhuri, P. K. (2009) Indian species of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). International Journal of Dipterological Research, 20, 191 - 201." title="Indian species of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2009">Mazumdar &amp; Chaudhuri 2009</bibRefCitation>
). New record for
<collectingCountry id="F8A5CC8BFFFBFFC04331315F034888E2" box="[269,351,368,391]" name="Yemen" pageId="5" pageNumber="138">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042FB31BC01FC8B7D" pageId="5" pageNumber="138" type="discussion">
<paragraph id="800D8C1BFFFBFFC042FB31BC01FC8B7D" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB31BC035888C9" bold="true" box="[199,335,403,428]" pageId="5" pageNumber="138">Comments.</emphasis>
After examination of the
<typeStatus id="5F0932B9FFFBFFC040A431BA00EF88C8" box="[664,760,405,429]" pageId="5" pageNumber="138" type="paratype">paratype</typeStatus>
male of
<taxonomicName id="47B2F798FFFBFFC0414F31BB064B88C9" box="[883,1116,404,428]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFBFFC0414F31BB064B88C9" box="[883,1116,404,428]" italics="true" pageId="5" pageNumber="138">Dasyhelea labinoda</emphasis>
</taxonomicName>
we consider there are no differences in the morphological characters between this species and
<taxonomicName id="47B2F798FFFBFFC041823196065C88B5" box="[958,1099,440,464]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="deemingi">
<emphasis id="B2C65009FFFBFFC04182319601C588B5" box="[958,978,441,464]" italics="true" pageId="5" pageNumber="138">D</emphasis>
.
<emphasis id="B2C65009FFFBFFC041D83197065C88B5" box="[996,1099,440,464]" italics="true" pageId="5" pageNumber="138">deemingi</emphasis>
</taxonomicName>
. Szadziewski and co-authors (
<bibRefCitation id="E423F1EAFFFBFFC042A231F303A68891" author="Szadziewski, R. &amp; Gwizdalska-Kentzer, M. &amp; Gilka, W." box="[158,433,476,500]" editor="van Harten, A." etAl="et al." firstAuthor="Szadziewski" journalOrPublisher="Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE" pageId="5" pageNumber="138" pagination="636 - 653" refId="ref7546" refString="Szadziewski, R., Gwizdalska-Kentzer, M. &amp; Gilka, W. (2011) Order Diptera, family Ceratopogonidae. In: Sheikh Tahnoon Bin Zayed Al Nahyan, H. H. (patron), van Harten, A. (Ed.), Arthropod fauna of the United Arab Emirates. Volume 4. Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE, pp. 636 - 653." title="Order Diptera, family Ceratopogonidae" type="book chapter" volumeTitle="Arthropod fauna of the United Arab Emirates. Volume 4" year="2011">
Szadziewski
<emphasis id="B2C65009FFFBFFC0430931F2037E8891" box="[309,361,476,500]" italics="true" pageId="5" pageNumber="138">et al</emphasis>
. 2011
</bibRefCitation>
) suggested that these two names are probably conspecific, and currently we propose to treat
<taxonomicName id="47B2F798FFFBFFC042F3322E03438B7D" box="[207,340,512,536]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFBFFC042F3322E02F48B7D" box="[207,227,513,536]" italics="true" pageId="5" pageNumber="138">D</emphasis>
.
<emphasis id="B2C65009FFFBFFC042CD322F03438B7D" box="[241,340,512,536]" italics="true" pageId="5" pageNumber="138">labinoda</emphasis>
</taxonomicName>
as a new junior synonym of
<taxonomicName id="47B2F798FFFBFFC040A6322E01358B7D" box="[666,802,512,536]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="deemingi">
<emphasis id="B2C65009FFFBFFC040A6322E00B98B7D" box="[666,686,513,536]" italics="true" pageId="5" pageNumber="138">D</emphasis>
.
<emphasis id="B2C65009FFFBFFC04080322F01358B7D" box="[700,802,512,536]" italics="true" pageId="5" pageNumber="138">deemingi</emphasis>
</taxonomicName>
.
<emphasis id="B2C65009FFFBFFC0410C322F01FC8B7D" bold="true" box="[816,1003,512,536]" pageId="5" pageNumber="138">New synonymy.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

157
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<subSubSection id="C8A8DF90FFFAFFC142AB30B8031C89B7" pageId="4" pageNumber="137" type="nomenclature">
<paragraph id="800D8C1BFFFAFFC142AB30B8002189D4" blockId="4.[151,566,151,210]" box="[151,566,151,177]" pageId="4" pageNumber="137">
<heading id="DB453B77FFFAFFC142AB30B8002189D4" bold="true" box="[151,566,151,177]" fontSize="11" level="1" pageId="4" pageNumber="137" reason="1">
<emphasis id="B2C65009FFFAFFC142AB30B8002189D4" bold="true" box="[151,566,151,177]" pageId="4" pageNumber="137">
<taxonomicName id="47B2F798FFFAFFC142AB30B803C689D4" ID-CoL="9HN7C" authority="Dominiak" authorityName="Dominiak" box="[151,465,151,177]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="avia" status="sp. nov.">
<emphasis id="B2C65009FFFAFFC142AB30B8035989D4" bold="true" box="[151,334,151,177]" italics="true" pageId="4" pageNumber="137">Dasyhelea avia</emphasis>
Dominiak
</taxonomicName>
<taxonomicNameLabel id="A9F5ED72FFFAFFC143E430B8002189D4" box="[472,566,151,177]" pageId="4" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="800D8C1BFFFAFFC142AB3095031C89B7" blockId="4.[151,566,151,210]" box="[151,267,186,210]" pageId="4" pageNumber="137">
(
<figureCitation id="1889909EFFFAFFC142A33095031489B7" box="[159,259,186,210]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="4" pageNumber="137">Figs 4–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFAFFC142AB312C064B8807" pageId="4" pageNumber="137" type="materials_examined">
<paragraph id="800D8C1BFFFAFFC142AB312C064B8807" blockId="4.[151,1437,258,976]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142AB312C0350887E" bold="true" box="[151,327,258,283]" pageId="4" pageNumber="137">
<typeStatus id="5F0932B9FFFAFFC142AB312C02C5887E" box="[151,210,259,283]" pageId="4" pageNumber="137">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5F0932B9FFFAFFC1436C312D03AA887F" box="[336,445,258,282]" pageId="4" pageNumber="137" type="holotype">Holotype</typeStatus>
: adult male.
<emphasis id="B2C65009FFFAFFC1406D312D0170887E" bold="true" box="[593,871,258,283]" pageId="4" pageNumber="137">
<collectingCountry id="F8A5CC8BFFFAFFC1406D312D0176887E" box="[593,865,258,283]" name="United Arab Emirates" pageId="4" pageNumber="137">United Arab Emirates</collectingCountry>
.
</emphasis>
Al-Ajban, N 24˚36' E 55˚01',
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asl,
<date id="F40CAADBFFFAFFC14706312D030E885B" pageId="4" pageNumber="137" value="2006-08-21" valueMax="2006-09-09" valueMin="2006-08-21">21.VIII– 09.IX.2006</date>
, Malaise trap, leg. A. van Harten (UG).
<typeStatus id="5F0932B9FFFAFFC140C73108017B885A" box="[763,876,295,319]" pageId="4" pageNumber="137" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B2C65009FFFAFFC1414731090683885A" bold="true" box="[891,1172,294,319]" pageId="4" pageNumber="137">
<collectingCountry id="F8A5CC8BFFFAFFC1414731090698885A" box="[891,1167,294,319]" name="United Arab Emirates" pageId="4" pageNumber="137">United Arab Emirates</collectingCountry>
.
</emphasis>
Al-Ajban, N 24˚36' E 55˚01',
<quantity id="474A21FEFFFAFFC142D7316503338807" box="[235,292,330,354]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="137" unit="m" value="60.0">60 m</quantity>
asl,
<date id="F40CAADBFFFAFFC143643165002B8807" box="[344,572,330,354]" pageId="4" pageNumber="137" value="2006-08-21" valueMax="2006-09-09" valueMin="2006-08-21">21.VIII–09.IX.2006</date>
,
<specimenCount id="96B44792FFFAFFC1407B316500B78807" box="[583,672,330,354]" pageId="4" pageNumber="137" type="male">2 males</specimenCount>
, Malaise trap, leg. A. van Harten (UG).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFAFFC142FB3141037188CF" pageId="4" pageNumber="137" type="diagnosis">
<paragraph id="800D8C1BFFFAFFC142FB3141037188CF" blockId="4.[151,1437,258,976]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142FB3141035688E2" bold="true" box="[199,321,366,391]" pageId="4" pageNumber="137">Diagnosis.</emphasis>
This species is easily diagnosed by very short, hook-like submedian projections of aedeagus and wshaped parameres.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFAFFC142FB319901FC8A5A" pageId="4" pageNumber="137" type="description">
<paragraph id="800D8C1BFFFAFFC142FB319900758B92" blockId="4.[151,1437,258,976]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142FB319903B188AA" bold="true" box="[199,422,438,463]" pageId="4" pageNumber="137">Description. Male.</emphasis>
Head. Flagellum length
<quantity id="474A21FEFFFAFFC140F53199016788AB" box="[713,880,438,462]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="4" pageNumber="137" unit="mm" value="0.54" valueMax="0.55" valueMin="0.53">0.53–0.55 mm</quantity>
, AR 0.89–0.94 (n=2). Distal flagellomeres with two rows of long setae (
<figureCitation id="1889909EFFFAFFC1439931F503FC8897" box="[421,491,474,498]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="4" pageNumber="137">Fig. 4</figureCitation>
). Frontal sclerite rather broad. Clypeus bearing 8 strong setae. Third palpal segment rather long and slender, length 47–49 μm, PR III 3.92–4.45 (n=2). Thorax. Transverse suture absent, number of supraalar setae from 7 to 8. Scutellum pale, with 6 long setae. Wing membrane covered with macrotrichia, hyaline, only one radial cell present. Wing length
<quantity id="474A21FEFFFAFFC140593266011D8B04" box="[613,778,585,609]" metricMagnitude="-4" metricUnit="m" metricValue="7.05" metricValueMax="7.1" metricValueMin="7.0" pageId="4" pageNumber="137" unit="mm" value="0.705" valueMax="0.71" valueMin="0.7">0.70–0.71 mm</quantity>
, CR 0.42–0.44 (n=3). Legs unicolored,
<collectingCountry id="F8A5CC8BFFFAFFC146F4326506FB8B04" box="[1224,1260,586,609]" name="Chile" pageId="4" pageNumber="137">TR</collectingCountry>
I 2.2 (n=2),
<collectingCountry id="F8A5CC8BFFFAFFC14752326507858B04" box="[1390,1426,586,609]" name="Chile" pageId="4" pageNumber="137">TR</collectingCountry>
II 2.3 (n=2),
<collectingCountry id="F8A5CC8BFFFAFFC14331325E03268BED" box="[269,305,625,648]" name="Chile" pageId="4" pageNumber="137">TR</collectingCountry>
III 2.2 (n=1). Genitalia (
<figureCitation id="1889909EFFFAFFC14008325F006F8BED" box="[564,632,624,648]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="4" pageNumber="137">Fig. 5</figureCitation>
). Tergite 9 rounded distally, apicolateral processes short, with sharp tip. Posterior margin of sternite 9 extended, fused to aedeagus by conspicuous bridge. Gonostylus slightly curved, slender. Parameres fused and symmetrical, w-shaped. Aedeagus with one pair of short submedian projections, both with recurved tips resemble small hooks.
</paragraph>
<paragraph id="800D8C1BFFFAFFC142FB332C00498A7E" blockId="4.[151,1437,258,976]" box="[199,606,770,795]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142FB332C03F68A7E" bold="true" box="[199,481,770,795]" pageId="4" pageNumber="137">Female and immatures.</emphasis>
Unknown.
</paragraph>
<paragraph id="800D8C1BFFFAFFC142FB330901FC8A5A" blockId="4.[151,1437,258,976]" box="[199,1003,806,831]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142FB330903F78A5A" bold="true" box="[199,480,806,831]" pageId="4" pageNumber="137">Derivation of the name.</emphasis>
The Latin noun
<taxonomicName id="47B2F798FFFAFFC140A7330700DC8A5A" box="[667,715,808,831]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="avia">
<emphasis id="B2C65009FFFAFFC140A7330700DC8A5A" box="[667,715,808,831]" italics="true" pageId="4" pageNumber="137">avia</emphasis>
</taxonomicName>
,-
<emphasis id="B2C65009FFFAFFC140E7330701018A5A" box="[731,790,808,831]" italics="true" pageId="4" pageNumber="137">orum</emphasis>
means wilderness.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFAFFC142FB336500808A06" box="[199,663,842,867]" pageId="4" pageNumber="137" type="distribution">
<paragraph id="800D8C1BFFFAFFC142FB336500808A06" blockId="4.[151,1437,258,976]" box="[199,663,842,867]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142FB336503778A06" bold="true" box="[199,352,842,867]" pageId="4" pageNumber="137">Distribution.</emphasis>
The
<collectingCountry id="F8A5CC8BFFFAFFC143A6336400848A06" box="[410,659,843,867]" name="United Arab Emirates" pageId="4" pageNumber="137">United Arab Emirates</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFAFFC142FB334106358AAA" pageId="4" pageNumber="137" type="discussion">
<paragraph id="800D8C1BFFFAFFC142FB334106358AAA" blockId="4.[151,1437,258,976]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142FB334103588AE2" bold="true" box="[199,335,878,903]" pageId="4" pageNumber="137">Comments.</emphasis>
<taxonomicName id="47B2F798FFFAFFC1436B334000118AE2" box="[343,518,879,903]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="avia">
<emphasis id="B2C65009FFFAFFC1436B334000118AE2" box="[343,518,879,903]" italics="true" pageId="4" pageNumber="137">Dasyhelea avia</emphasis>
</taxonomicName>
and the species of the
<taxonomicName id="47B2F798FFFAFFC14131334001798AE2" box="[781,878,879,903]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="turficola">
<emphasis id="B2C65009FFFAFFC14131334001798AE2" box="[781,878,879,903]" italics="true" pageId="4" pageNumber="137">turficola</emphasis>
</taxonomicName>
group are close in similar structure of the bridge joining sternite 9 and aedeagus, and the fused symmetrical parameres. It can be easily separated from the other species by the shape of aedeagal projections, which are very short and hook-like.
</paragraph>
</subSubSection>
</treatment>
</document>

119
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<mods:title id="62CDACBE290E947CE0BD958A094AE636">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<treatment id="081B3D0DFFFAFFC142AB340C01128C31" ID-DOI="http://doi.org/10.5281/zenodo.4499980" ID-GBIF-Taxon="119571711" ID-Zenodo-Dep="4499980" LSID="urn:lsid:plazi:treatment:081B3D0DFFFAFFC142AB340C01128C31" httpUri="http://treatment.plazi.org/id/081B3D0DFFFAFFC142AB340C01128C31" lastPageNumber="137" pageId="4" pageNumber="137">
<subSubSection id="C8A8DF90FFFAFFC142AB340C03F98D59" box="[151,494,1058,1085]" pageId="4" pageNumber="137" type="nomenclature">
<paragraph id="800D8C1BFFFAFFC142AB340C03F98D59" blockId="4.[151,494,1058,1085]" box="[151,494,1058,1085]" pageId="4" pageNumber="137">
<heading id="DB453B77FFFAFFC142AB340C03F98D59" bold="true" box="[151,494,1058,1085]" fontSize="11" level="1" pageId="4" pageNumber="137" reason="1">
<taxonomicName id="47B2F798FFFAFFC142AB340C03F98D59" ID-CoL="8BQP8" authority="Kieffer" authorityName="Kieffer" box="[151,494,1058,1085]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="bicrenata">
<emphasis id="B2C65009FFFAFFC142AB340C03F98D59" bold="true" box="[151,494,1058,1085]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142AB340C03998D58" bold="true" box="[151,398,1059,1085]" italics="true" pageId="4" pageNumber="137">Dasyhelea bicrenata</emphasis>
Kieffer
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFAFFC142AB344600848D1A" box="[151,659,1129,1151]" pageId="4" pageNumber="137" type="reference_group">
<paragraph id="800D8C1BFFFAFFC142AB344600848D1A" blockId="4.[151,659,1129,1151]" box="[151,659,1129,1151]" pageId="4" pageNumber="137">
<treatmentCitationGroup id="A0A2AB35FFFAFFC142AB344600848D1A" box="[151,659,1129,1151]" pageId="4" pageNumber="137">
<taxonomicName id="47B2F798FFFAFFC142AB344600988D1A" ID-CoL="8BQP8" authority="Kieffer, 1923: 668" authorityName="Kieffer" authorityPageNumber="668" authorityYear="1923" box="[151,655,1129,1151]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="bicrenata">
<emphasis id="B2C65009FFFAFFC142AB3446037F8D1A" box="[151,360,1129,1151]" italics="true" pageId="4" pageNumber="137">Dasyhelea bicrenata</emphasis>
<treatmentCitation id="0113AA0AFFFAFFC143523446003E8D1A" author="Kieffer" box="[366,553,1129,1151]" page="668" pageId="4" pageNumber="137" year="1923">
<bibRefCitation id="E423F1EAFFFAFFC143523446003E8D1A" author="Kieffer, J. J." box="[366,553,1129,1151]" firstAuthor="Kieffer" journalOrPublisher="Archives de l'Institut Pasteur Algerie" pageId="4" pageNumber="137" pagination="654 - 683" part="1" refId="ref7013" refString="Kieffer, J. J. (1923) Ceratopogonines recueillis au Sahara Constantinois. Archives de l'Institut Pasteur Algerie, 1, 654 - 683." title="Ceratopogonines recueillis au Sahara Constantinois" type="journal article" year="1923">Kieffer, 1923: 668</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F8A5CC8BFFFAFFC1400A3446009F8D1A" box="[566,648,1129,1151]" name="Algeria" pageId="4" pageNumber="137">Algeria</collectingCountry>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFAFFC142AB3483038C8D8D" pageId="4" pageNumber="137" type="materials_examined">
<paragraph id="800D8C1BFFFAFFC142AB3483038C8D8D" blockId="4.[151,1436,1196,1364]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142AB348303EC8DA0" bold="true" box="[151,507,1196,1221]" pageId="4" pageNumber="137">
Material examined.
<collectingCountry id="F8A5CC8BFFFAFFC143B6348203E18DA0" box="[394,502,1197,1221]" name="Lebanon" pageId="4" pageNumber="137">Lebanon</collectingCountry>
.
</emphasis>
Tyre, Al-Bass ruins, N 33°
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35°12.706, 19 m asl,
<date id="F40CAADBFFFAFFC146A93483071D8DA1" box="[1173,1290,1196,1220]" pageId="4" pageNumber="137" value="2012-05-12">12.V.2012</date>
, net,
<specimenCount id="96B44792FFFAFFC14774348307808DA1" box="[1352,1431,1196,1220]" pageId="4" pageNumber="137" type="male">1 male</specimenCount>
, leg. P. Dominiak (UG).
</paragraph>
</subSubSection>
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<paragraph id="800D8C1BFFFAFFC142FB34DB01128C31" blockId="4.[151,1436,1196,1364]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142FB34DB03778C68" bold="true" box="[199,352,1268,1293]" pageId="4" pageNumber="137">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFFAFFC1435134DB03BA8C69" box="[365,429,1268,1292]" name="Spain" pageId="4" pageNumber="137">Spain</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC143FE34DB01B58C69" author="Sahuquillo Herraiz, C. &amp; Gil Collado, J." box="[450,930,1268,1292]" firstAuthor="Sahuquillo Herraiz" journalOrPublisher="Revista de Sanidad e Higiene Publica" pageId="4" pageNumber="137" pagination="743 - 752" part="56" refId="ref7399" refString="Sahuquillo Herraiz, C. &amp; Gil Collado, J. (1982) Ceratopogonidae (Diptera Nematocera) de Navarra. Revista de Sanidad e Higiene Publica, 56, 743 - 752." title="Ceratopogonidae (Diptera Nematocera) de Navarra" type="journal article" year="1982">Sahuquillo Herráiz &amp; Gil Collado 1982</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC1418734DB06188C69" box="[955,1039,1268,1292]" name="Algeria" pageId="4" pageNumber="137">Algeria</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC1461834DB06A88C69" author="Kieffer, J. J." box="[1060,1215,1268,1292]" firstAuthor="Kieffer" journalOrPublisher="Archives de l'Institut Pasteur Algerie" pageId="4" pageNumber="137" pagination="654 - 683" part="1" refId="ref7013" refString="Kieffer, J. J. (1923) Ceratopogonines recueillis au Sahara Constantinois. Archives de l'Institut Pasteur Algerie, 1, 654 - 683." title="Ceratopogonines recueillis au Sahara Constantinois" type="journal article" year="1923">Kieffer 1923</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC146E534DB073D8C69" box="[1241,1322,1268,1292]" name="Poland" pageId="4" pageNumber="137">Poland</collectingCountry>
,
<collectingCountry id="F8A5CC8BFFFAFFC1470734DB078C8C69" box="[1339,1435,1268,1292]" name="Bulgaria" pageId="4" pageNumber="137">Bulgaria</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC142A2353703E38C55" author="Szadziewski, R. &amp; Borkent, A." box="[158,500,1304,1328]" firstAuthor="Szadziewski" journalOrPublisher="Polish Journal of Entomology" pageId="4" pageNumber="137" pagination="249 - 260" part="72" refId="ref7508" refString="Szadziewski, R. &amp; Borkent, A. (2003) New synonyms, combinations and records of biting midges (Diptera: Ceratopogonidae). Polish Journal of Entomology, 72, 249 - 260." title="New synonyms, combinations and records of biting midges (Diptera: Ceratopogonidae)" type="journal article" year="2003">Szadziewski &amp; Borkent 2003</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC14035353700748C55" box="[521,611,1304,1328]" name="Ukraine" pageId="4" pageNumber="137">Ukraine</collectingCountry>
(Crimea) (
<bibRefCitation id="E423F1EAFFFAFFC140DE353706598C55" author="Dominiak, P. &amp; Szadziewski, R." box="[738,1102,1304,1328]" firstAuthor="Dominiak" journalOrPublisher="Zootaxa" pageId="4" pageNumber="137" pagination="1 - 37" part="2437" refId="ref6594" refString="Dominiak, P. &amp; Szadziewski, R. (2010) Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). Zootaxa, 2437, 1 - 37." title="Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2010">Dominiak &amp; Szadziewski 2010</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC1465F353706AA8C55" box="[1123,1213,1304,1328]" name="Georgia" pageId="4" pageNumber="137">Georgia</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC146F1353707488C55" author="Remm, H." box="[1229,1375,1304,1328]" firstAuthor="Remm" journalOrPublisher="Tartu Riikliku Ulikodi Toimetised" pageId="4" pageNumber="137" pagination="3 - 37" part="194" refId="ref7303" refString="Remm, H. (1967) On the fauna of Ceratopogonidae (Diptera) in the Caucasus. Tartu Riikliku Ulikodi Toimetised, 194, 3 - 37." title="On the fauna of Ceratopogonidae (Diptera) in the Caucasus" type="journal article" year="1967">Remm 1967</bibRefCitation>
). For the first time reported from
<collectingCountry id="F8A5CC8BFFFAFFC143F3351300268C31" box="[463,561,1340,1364]" name="Lebanon" pageId="4" pageNumber="137">Lebanon</collectingCountry>
and the Near East.
</paragraph>
</subSubSection>
</treatment>
</document>

140
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@ -0,0 +1,140 @@
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<mods:title id="7CF061D8EAAE23E5B87F382925414340">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<emphasis id="B2C65009FFFAFFC142AB358703768CA7" bold="true" box="[151,353,1448,1474]" italics="true" pageId="4" pageNumber="137">Dasyhelea bifida</emphasis>
Zilahi-Sebess
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<taxonomicName id="47B2F798FFFAFFC142AB35C003AA8F60" ID-CoL="348JG" authority="Kieffer" authorityName="Kieffer" box="[151,445,1519,1541]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="fasciigera">
<emphasis id="B2C65009FFFAFFC142AB35C0037A8F60" box="[151,365,1519,1541]" italics="true" pageId="4" pageNumber="137">Dasyhelea fasciigera</emphasis>
Kieffer
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var.
<emphasis id="B2C65009FFFAFFC143D235C0003E8F60" box="[494,553,1519,1541]" italics="true" pageId="4" pageNumber="137">bifida</emphasis>
<treatmentCitation id="0113AA0AFFFAFFC1401335C001088F61" author="Zilahi-Sebess" box="[559,799,1519,1541]" page="44" pageId="4" pageNumber="137" year="1936">
<bibRefCitation id="E423F1EAFFFAFFC1401335C001088F61" author="Zilahi-Sebess, G." box="[559,799,1519,1541]" firstAuthor="Zilahi-Sebess" journalOrPublisher="Acta Biologica (Szeged)" pageId="4" pageNumber="137" pagination="35 - 49" part="4" refId="ref7785" refString="Zilahi-Sebess, G. (1936) Die Heleiden-Fauna von Szeged und Umgebung. Acta Biologica (Szeged), 4, 35 - 49." title="Die Heleiden-Fauna von Szeged und Umgebung" type="journal article" year="1936">Zilahi-Sebess, 1936: 44</bibRefCitation>
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(
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)
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.
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<emphasis id="B2C65009FFFAFFC142AB361E03E88F2F" bold="true" box="[151,511,1585,1610]" pageId="4" pageNumber="137">
Material examined.
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.
</emphasis>
Tyre, Tyre Coast Nature Reserve, N 33°
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35°12.794,
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, net,
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, leg. A. Alwin (UG).
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<emphasis id="B2C65009FFFAFFC142FB365603778FF7" bold="true" box="[199,352,1657,1682]" pageId="4" pageNumber="137">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFFAFFC14357365603AE8FF4" box="[363,441,1657,1681]" name="Poland" pageId="4" pageNumber="137">Poland</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC143F73656012B8FF4" author="Dominiak, P. &amp; Szadziewski, R." box="[459,828,1657,1681]" firstAuthor="Dominiak" journalOrPublisher="Zootaxa" pageId="4" pageNumber="137" pagination="1 - 37" part="2437" refId="ref6594" refString="Dominiak, P. &amp; Szadziewski, R. (2010) Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). Zootaxa, 2437, 1 - 37." title="Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2010">Dominiak &amp; Szadziewski 2010</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC1416F365501A28FF7" box="[851,949,1658,1682]" name="Hungary" pageId="4" pageNumber="137">Hungary</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC141F4365606BE8FF4" author="Zilahi-Sebess, G." box="[968,1193,1657,1681]" firstAuthor="Zilahi-Sebess" journalOrPublisher="Acta Biologica (Szeged)" pageId="4" pageNumber="137" pagination="35 - 49" part="4" refId="ref7785" refString="Zilahi-Sebess, G. (1936) Die Heleiden-Fauna von Szeged und Umgebung. Acta Biologica (Szeged), 4, 35 - 49." title="Die Heleiden-Fauna von Szeged und Umgebung" type="journal article" year="1936">Zilahi-Sebess 1936</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC146FC365607328FF4" box="[1216,1317,1657,1681]" name="Romania" pageId="4" pageNumber="137">Romania</collectingCountry>
(Damian-
<bibRefCitation id="E423F1EAFFFAFFC142AB36B203458FD0" author="Damian-Georgescu, A." box="[151,338,1693,1717]" journalOrPublisher="Studii s i Cercetari de Biologie, Seria Zoologie" pageId="4" pageNumber="137" pagination="97 - 100" part="27" refId="ref6390" refString="Damian-Georgescu, A. (1975) Ceratopogonidae (Diptera) noi pentru fauna Romaniei. Studii s i Cercetari de Biologie, Seria Zoologie, 27, 97 - 100." title="Ceratopogonidae (Diptera) noi pentru fauna Romaniei" type="journal article" year="1975">Georgescu 1975</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC1435536B203D48FD0" box="[361,451,1693,1717]" name="Ukraine" pageId="4" pageNumber="137">Ukraine</collectingCountry>
(Crimea) (
<bibRefCitation id="E423F1EAFFFAFFC1407E36B201BA8FD0" author="Dominiak, P. &amp; Szadziewski, R." box="[578,941,1693,1717]" firstAuthor="Dominiak" journalOrPublisher="Zootaxa" pageId="4" pageNumber="137" pagination="1 - 37" part="2437" refId="ref6594" refString="Dominiak, P. &amp; Szadziewski, R. (2010) Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). Zootaxa, 2437, 1 - 37." title="Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2010">Dominiak &amp; Szadziewski 2010</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC141FE36B206178FD0" box="[962,1024,1693,1717]" name="Israel" pageId="4" pageNumber="137">Israel</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC1462D36B2071B8FD0" author="Dominiak, P. &amp; Braverman, Y. &amp; Szadziewski, R." box="[1041,1292,1693,1718]" etAl="et al." firstAuthor="Dominiak" journalOrPublisher="Polish Journal of Entomology" pageId="4" pageNumber="137" pagination="257 - 265" part="76" refId="ref6499" refString="Dominiak, P., Braverman, Y. &amp; Szadziewski, R. (2007) New records of biting midges of the genus Dasyhelea from Israel, with notes on synonymy in the genera Culicoides and Forcipomyia (Diptera: Ceratopogonidae). Polish Journal of Entomology, 76, 257 - 265." title="New records of biting midges of the genus Dasyhelea from Israel, with notes on synonymy in the genera Culicoides and Forcipomyia (Diptera: Ceratopogonidae)" type="journal article" year="2007">
Dominiak
<emphasis id="B2C65009FFFAFFC146B036B006D78FD3" box="[1164,1216,1694,1718]" italics="true" pageId="4" pageNumber="137">et al</emphasis>
. 2007
</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC1471D36B2078B8FD0" box="[1313,1436,1693,1717]" name="Azerbaijan" pageId="4" pageNumber="137">Azerbaijan</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC142A236EE03248FBC" author="Remm, H." box="[158,307,1729,1753]" firstAuthor="Remm" journalOrPublisher="Tartu Riikliku Ulikodi Toimetised" pageId="4" pageNumber="137" pagination="3 - 37" part="194" refId="ref7303" refString="Remm, H. (1967) On the fauna of Ceratopogonidae (Diptera) in the Caucasus. Tartu Riikliku Ulikodi Toimetised, 194, 3 - 37." title="On the fauna of Ceratopogonidae (Diptera) in the Caucasus" type="journal article" year="1967">Remm 1967</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC1437136EE03C18FBC" box="[333,470,1729,1753]" name="Afghanistan" pageId="4" pageNumber="137">Afghanistan</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFAFFC143D736EE006F8FBC" author="Navai, S." box="[491,632,1729,1753]" firstAuthor="Navai" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="4" pageNumber="137" pagination="357 - 399" part="41" refId="ref7222" refString="Navai, S. (1994) Biting midges of the genus Dasyhelea from Afghanistan, with descriptions of new species (Diptera, Ceratopogonidae). Deutsche Entomologische Zeitschrift, 41, 357 - 399. http: // dx. doi. org / 10.1002 / mmnd. 19940410207" title="Biting midges of the genus Dasyhelea from Afghanistan, with descriptions of new species (Diptera, Ceratopogonidae)" type="journal article" year="1994">Navai 1994</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFAFFC140AE36EE00C28FBC" box="[658,725,1729,1753]" name="China" pageId="4" pageNumber="137">China</collectingCountry>
(Yu
<emphasis id="B2C65009FFFAFFC1412B36EC01598FBF" box="[791,846,1730,1754]" italics="true" pageId="4" pageNumber="137">et al</emphasis>
. 2006), Hawaii (
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(
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Material examined.
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.
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Jezzine waterfall,
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, leg. P. Dominiak (LU &amp; UG).
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The species was previously known only from Europe, as reported from
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Hagan
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(
<bibRefCitation id="E423F1EAFFFBFFC041153312063F8A30" author="Tothova, A. &amp; Knoz, J." box="[809,1064,829,853]" firstAuthor="Tothova" journalOrPublisher="Studia dipterologica" pageId="5" pageNumber="138" pagination="103 - 114" part="13" refId="ref7629" refString="Tothova, A. &amp; Knoz, J. (2006) A revised check-list of the biting midges (Diptera, Ceratopogonidae) of the Czech Republic and Slovakia. Studia dipterologica, 13, 103 - 114." title="A revised check-list of the biting midges (Diptera, Ceratopogonidae) of the Czech Republic and Slovakia" type="journal article" year="2006">Tóthová &amp; Knoz 2006</bibRefCitation>
) and
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(
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). This is the first record from
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and from the region of the Near East.
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(
<figureCitation id="1889909EFFFBFFC042A333C003148D62" box="[159,259,1007,1031]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="5" pageNumber="138">Figs 6–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042AB341701B58C42" pageId="5" pageNumber="138" type="materials_examined">
<paragraph id="800D8C1BFFFBFFC042AB341701B58C42" blockId="5.[151,1437,1079,2009]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042AB3417035E8D35" bold="true" box="[151,329,1079,1104]" pageId="5" pageNumber="138">
<typeStatus id="5F0932B9FFFBFFC042AB341702C58D35" box="[151,210,1080,1104]" pageId="5" pageNumber="138">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5F0932B9FFFBFFC04368341803D68D2A" box="[340,449,1079,1103]" pageId="5" pageNumber="138" type="holotype">Holotype</typeStatus>
: adult male.
<emphasis id="B2C65009FFFBFFC04066341700DC8D35" bold="true" box="[602,715,1080,1104]" pageId="5" pageNumber="138">
<collectingCountry id="F8A5CC8BFFFBFFC04066341700D18D35" box="[602,710,1080,1104]" name="Lebanon" pageId="5" pageNumber="138">Lebanon</collectingCountry>
.
</emphasis>
Tyre, Tyre Coast Nature Reserve, N 33°
<geoCoordinate id="E586EADCFFFBFFC04696341807018D2A" box="[1194,1302,1079,1103]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="15.206">15.206 E</geoCoordinate>
35°12.794,
<date id="F40CAADBFFFBFFC042AB3473031B8D11" box="[151,268,1116,1140]" pageId="5" pageNumber="138" value="2012-05-06">06.V.2012</date>
, net, leg. P. Dominiak (
<collectionCode id="E6A314DEFFFBFFC04015347300938D11" box="[553,644,1116,1140]" pageId="5" pageNumber="138">BMNH</collectionCode>
).
<typeStatus id="5F0932B9FFFBFFC040A73472011B8D10" box="[667,780,1117,1141]" pageId="5" pageNumber="138" type="paratype">Paratypes</typeStatus>
:
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<collectingCountry id="F8A5CC8BFFFBFFC041203473019F8D11" box="[796,904,1116,1140]" name="Lebanon" pageId="5" pageNumber="138">Lebanon</collectingCountry>
.
</emphasis>
Tyre, N 33°
<geoCoordinate id="E586EADCFFFBFFC0461F347306988D11" box="[1059,1167,1116,1140]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="15.535">15.535 E</geoCoordinate>
35°12.726,
<date id="F40CAADBFFFBFFC0471E347307808D11" box="[1314,1431,1116,1140]" pageId="5" pageNumber="138" value="2012-05-06">06.V.2012</date>
, beach, net,
<specimenCount id="96B44792FFFBFFC0431E345003978DF2" box="[290,384,1151,1175]" pageId="5" pageNumber="138" type="male">4 males</specimenCount>
, leg. A. Alwin &amp; P. Dominiak (UG). Tyre, Tyre Coast Nature Reserve, N 33°
<geoCoordinate id="E586EADCFFFBFFC047123450078B8DF2" box="[1326,1436,1151,1175]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="15.206">15.206 E</geoCoordinate>
35°12.794,
<date id="F40CAADBFFFBFFC04326348B03988DD9" box="[282,399,1188,1212]" pageId="5" pageNumber="138" value="2012-05-06">06.V.2012</date>
, net,
<specimenCount id="96B44792FFFBFFC043F1348B00318DD9" box="[461,550,1188,1212]" pageId="5" pageNumber="138" type="male">3 males</specimenCount>
, leg. P. Dominiak (LU &amp; UG). Nahr al-Kalb, N 33°
<geoCoordinate id="E586EADCFFFBFFC04652348B06C08DD9" box="[1134,1239,1188,1212]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="55.842">55.842 E</geoCoordinate>
35°35.326, 69 m asl,
<date id="F40CAADBFFFBFFC042F634E803578DBA" box="[202,320,1223,1247]" pageId="5" pageNumber="138" value="2012-05-03">03.V.2012</date>
, net,
<specimenCount id="96B44792FFFBFFC043B434E803CB8DBA" box="[392,476,1223,1247]" pageId="5" pageNumber="138" type="male">1 male</specimenCount>
, leg. P. Dominiak (UG). Aarbit Kozhayya, St. Antonios Kozhayya Monaster, N 34°
<geoCoordinate id="E586EADCFFFBFFC0428034C303308C61" box="[188,295,1260,1284]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="17.046">17.046 E</geoCoordinate>
35°56.689,
<date id="F40CAADBFFFBFFC0438534C300258C61" box="[441,562,1260,1284]" pageId="5" pageNumber="138" value="2012-10-28">28.X.2012</date>
, shallow cave, net,
<specimenCount id="96B44792FFFBFFC0412234C301678C61" box="[798,880,1260,1284]" pageId="5" pageNumber="138" type="male">1 male</specimenCount>
, leg. P. Dominiak (UG). Fanar, N 33°
<geoCoordinate id="E586EADCFFFBFFC0471334C3078B8C61" box="[1327,1436,1260,1284]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="52.68">52.680 E</geoCoordinate>
35°33.547, 138 m asl,
<date id="F40CAADBFFFBFFC043A83520001E8C42" box="[404,521,1295,1319]" pageId="5" pageNumber="138" value="2012-05-03">03.V.2012</date>
, net,
<specimenCount id="96B44792FFFBFFC04078352000858C42" box="[580,658,1295,1319]" pageId="5" pageNumber="138" type="male">1 male</specimenCount>
, leg. P. Dominiak (UG).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042FB351C03128CF1" pageId="5" pageNumber="138" type="diagnosis">
<paragraph id="800D8C1BFFFBFFC042FB351C03128CF1" blockId="5.[151,1437,1079,2009]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB351C03568C29" bold="true" box="[199,321,1331,1356]" pageId="5" pageNumber="138">Diagnosis.</emphasis>
The species belongs to the
<taxonomicName id="47B2F798FFFBFFC040AE351B01158C29" box="[658,770,1332,1356]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="fasciigera">
<emphasis id="B2C65009FFFBFFC040AE351B01158C29" box="[658,770,1332,1356]" italics="true" pageId="5" pageNumber="138">fasciigera</emphasis>
</taxonomicName>
group, and differs from all other members by stout and truncated apicolateral processes of 9th tergite in male genitalia, as well as by weakly sclerotized lateral borders of aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042FB35B001968E08" pageId="5" pageNumber="138" type="description">
<paragraph id="800D8C1BFFFBFFC042FB35B007328E40" blockId="5.[151,1437,1079,2009]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB35B003B28CDD" bold="true" box="[199,421,1439,1464]" pageId="5" pageNumber="138">Description. Male.</emphasis>
Flagellum length
<quantity id="474A21FEFFFBFFC04045358F01378CDD" box="[633,800,1440,1464]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.3" metricValueMin="5.6" pageId="5" pageNumber="138" unit="mm" value="0.595" valueMax="0.63" valueMin="0.56">0.56–0.63 mm</quantity>
, AR 0.81–0.92 (n=9). Distal flagellomeres bearing two rows of long setae (
<figureCitation id="1889909EFFFBFFC043B135EB03CF8CB9" box="[397,472,1476,1500]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="5" pageNumber="138">Fig. 6</figureCitation>
). Clypeus elongate, composed of two separate sclerites, with 7–12 strong setae. Maxillary palp as in fig. 7. Third palpal segment slender, 57–67 μm long, PR III 3.80–5.58 (n=10). Thorax. Transverse suture absent, 8–10 supraalar setae present. Scutellum entirely pale or dark, with 6–7 strong and few small setae. Wing length
<quantity id="474A21FEFFFBFFC043F8361D007A8F2F" box="[452,621,1586,1610]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.1" metricValueMin="7.9" pageId="5" pageNumber="138" unit="mm" value="0.85" valueMax="0.91" valueMin="0.79">0.79–0.91 mm</quantity>
, CR 0.43–0.46 (n=11). Only one radial cell present, wing membrane covered with macrotrichia. Legs unicolored, dark,
<collectingCountry id="F8A5CC8BFFFBFFC040E1367701168F0A" box="[733,769,1624,1647]" name="Chile" pageId="5" pageNumber="138">TR</collectingCountry>
I 1.9–2.1 (n=9),
<collectingCountry id="F8A5CC8BFFFBFFC04181367701F68F0A" box="[957,993,1624,1647]" name="Chile" pageId="5" pageNumber="138">TR</collectingCountry>
II 2.3–2.5 (n=10),
<collectingCountry id="F8A5CC8BFFFBFFC04693367706C48F0A" box="[1199,1235,1624,1647]" name="Chile" pageId="5" pageNumber="138">TR</collectingCountry>
III 2.0–2.2 (n=11). Genitalia (
<figureCitation id="1889909EFFFBFFC0432D3652034D8FF0" box="[273,346,1661,1685]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="5" pageNumber="138">Fig. 8</figureCitation>
). Posterior margin of tergite 9 with a pair of stout, apically truncated apicolateral processes. Posterior margin of sternite 9 elongate, conical. Gonostylus composed of two parts. Outer gonostylus long and straight, with varying length of finger-like prolongation near its base. Inner gonostylus strongly curved, sharp. Parameres asymmetrical. Posterior process broadly fused to right basal arm, stout basally, with divided apex. Lateral projections of aedeagus bent in apical half, with weakly sclerotized membranous lateral borders.
</paragraph>
<paragraph id="800D8C1BFFFBFFC042FB371D00498E2C" blockId="5.[151,1437,1079,2009]" box="[199,606,1841,1866]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB371D03F68E2F" bold="true" box="[199,481,1841,1866]" pageId="5" pageNumber="138">Female and immatures.</emphasis>
Unknown.
</paragraph>
<paragraph id="800D8C1BFFFBFFC042FB377A01968E08" blockId="5.[151,1437,1079,2009]" box="[199,897,1877,1902]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB377A03F78E0B" bold="true" box="[199,480,1877,1902]" pageId="5" pageNumber="138">Derivation of the name.</emphasis>
The species is named after
<collectingCountry id="F8A5CC8BFFFBFFC0412B377A016A8E08" box="[791,893,1877,1901]" name="Lebanon" pageId="5" pageNumber="138">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042FB375603C68EF4" box="[199,465,1913,1938]" pageId="5" pageNumber="138" type="distribution">
<paragraph id="800D8C1BFFFBFFC042FB375603C68EF4" blockId="5.[151,1437,1079,2009]" box="[199,465,1913,1938]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB375603778EF7" bold="true" box="[199,352,1913,1938]" pageId="5" pageNumber="138">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFFBFFC0435B375603DA8EF4" box="[359,461,1913,1937]" name="Lebanon" pageId="5" pageNumber="138">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC342FB37B2001A89B1" lastPageId="6" lastPageNumber="139" pageId="5" pageNumber="138" type="discussion">
<paragraph id="800D8C1BFFFBFFC342FB37B2001A89B1" blockId="5.[151,1437,1079,2009]" lastBlockId="6.[151,1436,151,212]" lastPageId="6" lastPageNumber="139" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB37B203588ED3" bold="true" box="[199,335,1949,1974]" pageId="5" pageNumber="138">Comments.</emphasis>
This newly described species is similar to
<taxonomicName id="47B2F798FFFBFFC0410D37B106628ED0" authority="Kieffer" authorityName="Kieffer" box="[817,1141,1949,1974]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="fasciigera">
<emphasis id="B2C65009FFFBFFC0410D37B106098ED3" box="[817,1054,1950,1974]" italics="true" pageId="5" pageNumber="138">Dasyhelea fasciigera</emphasis>
Kieffer
</taxonomicName>
but differs in the shape of apicolateral processes of tergite 9, which are truncated, whereas they are rounded in the latter species. Other diagnostic characters for the new species are the shape of the lateral processes of aedeagus bearing membranous weakly chitinized lateral borders.
</paragraph>
</subSubSection>
</treatment>
</document>

140
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<heading id="DB453B77FFFDFFC642AB30B8000789D4" bold="true" box="[151,528,151,177]" fontSize="11" level="1" pageId="3" pageNumber="136" reason="1">
<taxonomicName id="47B2F798FFFDFFC642AB30B8000789D4" ID-CoL="8BQN9" authority="Winnertz" authorityName="Winnertz" box="[151,528,151,177]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="modesta">
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<emphasis id="B2C65009FFFDFFC642AB30B8036889D4" bold="true" box="[151,383,151,177]" italics="true" pageId="3" pageNumber="136">Dasyhelea modesta</emphasis>
(Winnertz)
</emphasis>
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<taxonomicName id="47B2F798FFFDFFC642AB30F000D48990" authority="Winnertz, 1852: 43" authorityName="Winnertz" authorityPageNumber="43" authorityYear="1852" box="[151,707,222,245]" class="Insecta" family="Ceratopogonidae" genus="Ceratopogon" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="B2C65009FFFDFFC642AB30F003968990" box="[151,385,223,245]" italics="true" pageId="3" pageNumber="136">Ceratopogon modestus</emphasis>
<treatmentCitation id="0113AA0AFFFDFFC643BB30F1005D8991" author="Winnertz" box="[391,586,222,244]" page="43" pageId="3" pageNumber="136" year="1852">
<bibRefCitation id="E423F1EAFFFDFFC643BB30F1005D8991" author="Winnertz, J." box="[391,586,222,244]" firstAuthor="Winnertz" journalOrPublisher="Linnaea Entomologica" pageId="3" pageNumber="136" pagination="1 - 80" part="6" refId="ref7720" refString="Winnertz, J. (1852) Beitrag zur Kenntnis der Gattung Ceratopogon Meigen. Linnaea Entomologica, 6, 1 - 80." title="Beitrag zur Kenntnis der Gattung Ceratopogon Meigen" type="journal article" year="1852">Winnertz, 1852: 43</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F8A5CC8BFFFDFFC6406430F000AB8990" box="[600,700,223,245]" name="Germany" pageId="3" pageNumber="136">Germany</collectingCountry>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFDFFC642AB310F033E88AC" pageId="3" pageNumber="136" type="materials_examined">
<paragraph id="800D8C1BFFFDFFC642AB310F033E88AC" blockId="3.[151,1436,288,565]" pageId="3" pageNumber="136">
<emphasis id="B2C65009FFFDFFC642AB310F03CF885C" bold="true" box="[151,472,288,313]" pageId="3" pageNumber="136">
Material examined.
<collectingCountry id="F8A5CC8BFFFDFFC643B1310E03C3885C" box="[397,468,289,313]" name="Israel" pageId="3" pageNumber="136">Israel</collectingCountry>
.
</emphasis>
‘En Netafim,
<date id="F40CAADBFFFDFFC640BD310E0116885C" box="[641,769,289,313]" pageId="3" pageNumber="136" value="1994-03-12">12.III.1994</date>
,
<specimenCount id="96B44792FFFDFFC6412C310E0160885C" box="[784,887,289,313]" pageId="3" pageNumber="136" type="female">1 female</specimenCount>
, leg. A. Friedberg.
<emphasis id="B2C65009FFFDFFC64666310E06DC885C" bold="true" box="[1114,1227,289,313]" pageId="3" pageNumber="136">
<collectingCountry id="F8A5CC8BFFFDFFC64666310E06D1885C" box="[1114,1222,289,313]" name="Lebanon" pageId="3" pageNumber="136">Lebanon</collectingCountry>
.
</emphasis>
Jezzine waterfall,
<date id="F40CAADBFFFDFFC642AB316A031B8838" box="[151,268,325,349]" pageId="3" pageNumber="136" value="2012-05-01">01.V.2012</date>
, net,
<specimenCount id="96B44792FFFDFFC6437B316A03BC8838" box="[327,427,325,349]" pageId="3" pageNumber="136" type="female">1 female</specimenCount>
, leg. P. Dominiak (UG). Aarbit Kozhayya, St. Antonios Kozhayya Monaster, N 34˚
<geoCoordinate id="E586EADCFFFDFFC64770316A02BF88E4" direction="east" orientation="longitude" pageId="3" pageNumber="136" precision="55" value="17.046">17.046 E</geoCoordinate>
35˚56.689,
<date id="F40CAADBFFFDFFC64306314603A788E4" box="[314,432,361,385]" pageId="3" pageNumber="136" value="2012-05-08">08.V.2012</date>
, rock church, net,
<specimenCount id="96B44792FFFDFFC640A9314600FF88E4" box="[661,744,361,385]" pageId="3" pageNumber="136" type="male">1 male</specimenCount>
, leg. P. Dominiak (UG). Ammiq Reserve, N 33˚
<geoCoordinate id="E586EADCFFFDFFC647113146078B88E4" box="[1325,1436,361,385]" direction="east" orientation="longitude" pageId="3" pageNumber="136" precision="55" value="43.996">43.996 E</geoCoordinate>
35˚47.035, 876 m asl, wetland,
<date id="F40CAADBFFFDFFC643C731A2006388C0" box="[507,628,397,421]" pageId="3" pageNumber="136" value="2012-10-24">24.X.2012</date>
, net,
<specimenCount id="96B44792FFFDFFC6408D31A2010188C0" box="[689,790,397,421]" pageId="3" pageNumber="136" type="female">1 female</specimenCount>
, leg. P. Dominiak (UG). Fghal,
<date id="F40CAADBFFFDFFC6464031A206EA88C0" box="[1148,1277,397,421]" pageId="3" pageNumber="136" value="2011-06-15">15.VI.2011</date>
, Malaise trap,
<specimenCount id="96B44792FFFDFFC642AB319E02F588AC" box="[151,226,433,457]" pageId="3" pageNumber="136" type="male">1 male</specimenCount>
(LU).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFDFFC642FB31FA00D38B50" pageId="3" pageNumber="136" type="distribution">
<paragraph id="800D8C1BFFFDFFC642FB31FA00D38B50" blockId="3.[151,1436,288,565]" pageId="3" pageNumber="136">
<emphasis id="B2C65009FFFDFFC642FB31FA0377888B" bold="true" box="[199,352,469,494]" pageId="3" pageNumber="136">Distribution.</emphasis>
<taxonomicName id="47B2F798FFFDFFC6435731FA005F8888" box="[363,584,469,493]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B2C65009FFFDFFC6435731FA005F8888" box="[363,584,469,493]" italics="true" pageId="3" pageNumber="136">Dasyhelea modesta</emphasis>
</taxonomicName>
is widespread in the Palaearctic (
<bibRefCitation id="E423F1EAFFFDFFC641E831FA07538888" author="Dominiak, P. &amp; Szadziewski, R." box="[980,1348,469,493]" firstAuthor="Dominiak" journalOrPublisher="Zootaxa" pageId="3" pageNumber="136" pagination="1 - 37" part="2437" refId="ref6594" refString="Dominiak, P. &amp; Szadziewski, R. (2010) Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). Zootaxa, 2437, 1 - 37." title="Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2010">Dominiak &amp; Szadziewski 2010</bibRefCitation>
). Up to now, in the Middle East it was known from
<collectingCountry id="F8A5CC8BFFFDFFC640B531D500DB8B77" box="[649,716,506,530]" name="Egypt" pageId="3" pageNumber="136">Egypt</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC640E731D601788B74" author="Macfie, J. W. S." box="[731,879,505,529]" firstAuthor="Macfie" journalOrPublisher="Proceedings of the Royal Entomological Society of London (B)" pageId="3" pageNumber="136" pagination="145 - 159" part="12" refId="ref7101" refString="Macfie, J. W. S. (1943) Ceratopogonidae (Diptera) from Egypt. Proceedings of the Royal Entomological Society of London (B), 12, 145 - 159." title="Ceratopogonidae (Diptera) from Egypt" type="journal article" year="1943">Macfie 1943</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFDFFC641B831D501C58B74" box="[900,978,506,529]" name="Yemen" pageId="3" pageNumber="136">Yemen</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC641DE31D607388B74" author="Boorman, J. &amp; van Harten, A." box="[994,1327,505,529]" firstAuthor="Boorman" journalOrPublisher="Fauna of Arabia" pageId="3" pageNumber="136" pagination="427 - 462" part="19" refId="ref6017" refString="Boorman, J. &amp; van Harten, A. (2002) Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen. Fauna of Arabia, 19, 427 - 462." title="Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen" type="journal article" year="2002">Boorman &amp; van Harten 2002</bibRefCitation>
) and
<collectingCountry id="F8A5CC8BFFFDFFC6475331D5078B8B74" box="[1391,1436,506,529]" name="Iran" pageId="3" pageNumber="136">Iran</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC642A2323203788B50" author="Szadziewski, R." box="[158,367,541,565]" firstAuthor="Szadziewski" journalOrPublisher="Polish Journal of Entomology" pageId="3" pageNumber="136" pagination="3 - 103" part="56" refId="ref7480" refString="Szadziewski, R. (1986) Redescriptions and notes on some Ceratopogonidae (Diptera). Polish Journal of Entomology, 56, 3 - 103." title="Redescriptions and notes on some Ceratopogonidae (Diptera)" type="journal article" year="1986">Szadziewski 1986</bibRefCitation>
). New for
<collectingCountry id="F8A5CC8BFFFDFFC643DA323200338B50" box="[486,548,541,565]" name="Israel" pageId="3" pageNumber="136">Israel</collectingCountry>
and
<collectingCountry id="F8A5CC8BFFFDFFC64066323200D78B50" box="[602,704,541,565]" name="Lebanon" pageId="3" pageNumber="136">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

155
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<paragraph id="800D8C1BFFFDFFC642AB32E1008C8B8D" blockId="3.[151,667,718,744]" box="[151,667,718,744]" pageId="3" pageNumber="136">
<heading id="DB453B77FFFDFFC642AB32E1008C8B8D" bold="true" box="[151,667,718,744]" fontSize="11" level="1" pageId="3" pageNumber="136" reason="1">
<taxonomicName id="47B2F798FFFDFFC642AB32E1008C8B8D" ID-CoL="348JP" authority="Carter, Ingram &amp; Macfie" authorityName="Carter, Ingram &amp; Macfie" box="[151,667,718,744]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="B2C65009FFFDFFC642AB32E1008C8B8D" bold="true" box="[151,667,718,744]" pageId="3" pageNumber="136">
<emphasis id="B2C65009FFFDFFC642AB32E103408B8D" bold="true" box="[151,343,718,744]" italics="true" pageId="3" pageNumber="136">Dasyhelea flava</emphasis>
Carter, Ingram &amp; Macfie
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFDFFC642AB333A01078A4E" box="[151,784,789,811]" pageId="3" pageNumber="136" type="reference_group">
<paragraph id="800D8C1BFFFDFFC642AB333A01078A4E" blockId="3.[151,784,789,811]" box="[151,784,789,811]" pageId="3" pageNumber="136">
<treatmentCitationGroup id="A0A2AB35FFFDFFC642AB333A01078A4E" box="[151,784,789,811]" pageId="3" pageNumber="136">
<taxonomicName id="47B2F798FFFDFFC642AB333A011B8A4E" ID-CoL="348JP" authority="Carter, Ingram &amp; Macfie, 1921: 196" authorityName="Carter, Ingram &amp; Macfie" authorityPageNumber="196" authorityYear="1921" box="[151,780,789,811]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="B2C65009FFFDFFC642AB333A032D8A4E" box="[151,314,789,811]" italics="true" pageId="3" pageNumber="136">Dasyhelea flava</emphasis>
<treatmentCitation id="0113AA0AFFFDFFC6437D333A00A78A4E" author="Carter" box="[321,688,789,811]" page="196" pageId="3" pageNumber="136" year="1921">
<bibRefCitation id="E423F1EAFFFDFFC6437D333A00A78A4E" author="Carter, H. F. &amp; Ingram, A. &amp; Macfie, J. W. S." box="[321,688,789,811]" firstAuthor="Carter" journalOrPublisher="Annals of Tropical Medicine and Parasitology" pageId="3" pageNumber="136" pagination="177 - 212" part="15" refId="ref6142" refString="Carter, H. F., Ingram, A. &amp; Macfie, J. W. S. (1921) Observations on the ceratopogonine midges of the Gold Coast, with descriptions of new species I - IV. Annals of Tropical Medicine and Parasitology, 15, 177 - 212." title="Observations on the ceratopogonine midges of the Gold Coast, with descriptions of new species I - IV" type="journal article" year="1921">Carter, Ingram &amp; Macfie, 1921: 196</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F8A5CC8BFFFDFFC64081333A01128A4E" box="[701,773,789,811]" name="Ghana" pageId="3" pageNumber="136">Ghana</collectingCountry>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFDFFC642AB337802C78A9A" pageId="3" pageNumber="136" type="materials_examined">
<paragraph id="800D8C1BFFFDFFC642AB337802C78A9A" blockId="3.[151,1437,855,1168]" pageId="3" pageNumber="136">
<emphasis id="B2C65009FFFDFFC642AB337803EF8A15" bold="true" box="[151,504,855,880]" pageId="3" pageNumber="136">
Material examined.
<collectingCountry id="F8A5CC8BFFFDFFC643BB337703E48A15" box="[391,499,856,880]" name="Lebanon" pageId="3" pageNumber="136">Lebanon</collectingCountry>
.
</emphasis>
Damour River, N 33°
<geoCoordinate id="E586EADCFFFDFFC640D7337801438A0A" box="[747,852,855,879]" direction="east" orientation="longitude" pageId="3" pageNumber="136" precision="55" value="42.369">42.369 E</geoCoordinate>
35°27.637, 48 m asl,
<date id="F40CAADBFFFDFFC64676337806D48A0A" box="[1098,1219,855,879]" pageId="3" pageNumber="136" value="2012-10-26">26.X.2012</date>
, net,
<specimenCount id="96B44792FFFDFFC646C23378075B8A0A" box="[1278,1356,855,879]" pageId="3" pageNumber="136" type="male">1 male</specimenCount>
, leg. P. Dominiak (UG). Near Ras el-Ain, Tyre, N 33˚
<geoCoordinate id="E586EADCFFFDFFC6409B335301068AF1" box="[679,785,892,916]" direction="east" orientation="longitude" pageId="3" pageNumber="136" precision="55" value="12.927">12.927 E</geoCoordinate>
35˚13.056, 5 m asl,
<date id="F40CAADBFFFDFFC6463D335306618AF1" box="[1025,1142,892,916]" pageId="3" pageNumber="136" value="2012-05-06">06.V.2012</date>
, banana plantation, net,
<specimenCount id="96B44792FFFDFFC647B2335302C78AD2" pageId="3" pageNumber="136" type="male">1 male</specimenCount>
, leg. A. Alwin (UG). Ras el-Ain, Tyre, avocado plantation,
<date id="F40CAADBFFFDFFC6415433B006278AD2" box="[872,1072,927,951]" pageId="3" pageNumber="136" value="2012-09-15" valueMax="2012-10-20" valueMin="2012-09-15">15.IX–20.X.2012</date>
, yellow sticky trap,
<specimenCount id="96B44792FFFDFFC6472C33B0077F8AD2" box="[1296,1384,927,951]" pageId="3" pageNumber="136" type="male">2 males</specimenCount>
, leg. H. Abdul-Nour (UG); citrus plantation,
<date id="F40CAADBFFFDFFC6406933EB010B8AB9" box="[597,796,964,988]" pageId="3" pageNumber="136" value="2012-09-15" valueMax="2012-10-20" valueMin="2012-09-15">15.IX–20.X.2012</date>
, yellow sticky trap,
<specimenCount id="96B44792FFFDFFC641C133EB06738AB9" box="[1021,1124,964,988]" pageId="3" pageNumber="136" type="male">20 males</specimenCount>
, leg. H. Abdul-Nour (LU &amp; UG).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFDFFC642FB342400FC8DEA" pageId="3" pageNumber="136" type="distribution">
<paragraph id="800D8C1BFFFDFFC642FB342400FC8DEA" blockId="3.[151,1437,855,1168]" pageId="3" pageNumber="136">
<emphasis id="B2C65009FFFDFFC642FB342403778D41" bold="true" box="[199,352,1035,1060]" pageId="3" pageNumber="136">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFFDFFC6435B342303D78D41" box="[359,448,1036,1060]" name="Gambia" pageId="3" pageNumber="136">Gambia</collectingCountry>
(
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),
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<bibRefCitation id="E423F1EAFFFDFFC6417D3423061D8D41" author="Carter, H. F. &amp; Ingram, A. &amp; Macfie, J. W. S." box="[833,1034,1036,1060]" etAl="et al." firstAuthor="Carter" journalOrPublisher="Annals of Tropical Medicine and Parasitology" pageId="3" pageNumber="136" pagination="177 - 212" part="15" refId="ref6142" refString="Carter, H. F., Ingram, A. &amp; Macfie, J. W. S. (1921) Observations on the ceratopogonine midges of the Gold Coast, with descriptions of new species I - IV. Annals of Tropical Medicine and Parasitology, 15, 177 - 212." title="Observations on the ceratopogonine midges of the Gold Coast, with descriptions of new species I - IV" type="journal article" year="1921">
Carter
<emphasis id="B2C65009FFFDFFC641AC342201D68D41" box="[912,961,1036,1060]" italics="true" pageId="3" pageNumber="136">et al</emphasis>
. 1921
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),
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(
<bibRefCitation id="E423F1EAFFFDFFC64681342302C58D22" author="de Meillon, B. &amp; Wirth, W. W." firstAuthor="de Meillon" journalOrPublisher="Annals of the Natal Museum" pageId="3" pageNumber="136" pagination="525 - 561" part="24" refId="ref6339" refString="de Meillon, B. &amp; Wirth, W. W. (1981) Subsaharan Ceratopogonidae (Diptera) VI. New species and records of South African biting midges collected by A. L. Dyce. Annals of the Natal Museum, 24, 525 - 561." title="Subsaharan Ceratopogonidae (Diptera) VI. New species and records of South African biting midges collected by A. L. Dyce" type="journal article" year="1981">de Meillon &amp; Wirth 1981</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFDFFC642D5340003308D22" box="[233,295,1071,1095]" name="Israel" pageId="3" pageNumber="136">Israel</collectingCountry>
(
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Dominiak
<emphasis id="B2C65009FFFDFFC64389341E03FD8D2D" box="[437,490,1072,1096]" italics="true" pageId="3" pageNumber="136">et al</emphasis>
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),
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,
<collectingCountry id="F8A5CC8BFFFDFFC64091340000E58D22" box="[685,754,1071,1095]" name="Oman" pageId="3" pageNumber="136">Oman</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC6413F3400064A8D22" author="Boorman, J. &amp; van Harten, A." box="[771,1117,1071,1096]" firstAuthor="Boorman" journalOrPublisher="Fauna of Arabia" pageId="3" pageNumber="136" pagination="427 - 462" part="19" refId="ref6017" refString="Boorman, J. &amp; van Harten, A. (2002) Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen. Fauna of Arabia, 19, 427 - 462." title="Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen" type="journal article" year="2002">Boorman &amp; van Harten 2002</bibRefCitation>
), the
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(
<bibRefCitation id="E423F1EAFFFDFFC642A2347B03A68D09" author="Szadziewski, R. &amp; Gwizdalska-Kentzer, M. &amp; Gilka, W." box="[158,433,1108,1132]" editor="van Harten, A." etAl="et al." firstAuthor="Szadziewski" journalOrPublisher="Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE" pageId="3" pageNumber="136" pagination="636 - 653" refId="ref7546" refString="Szadziewski, R., Gwizdalska-Kentzer, M. &amp; Gilka, W. (2011) Order Diptera, family Ceratopogonidae. In: Sheikh Tahnoon Bin Zayed Al Nahyan, H. H. (patron), van Harten, A. (Ed.), Arthropod fauna of the United Arab Emirates. Volume 4. Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE, pp. 636 - 653." title="Order Diptera, family Ceratopogonidae" type="book chapter" volumeTitle="Arthropod fauna of the United Arab Emirates. Volume 4" year="2011">
Szadziewski
<emphasis id="B2C65009FFFDFFC64308347A03708D09" box="[308,359,1108,1132]" italics="true" pageId="3" pageNumber="136">et al</emphasis>
. 2011
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), and
<collectingCountry id="F8A5CC8BFFFDFFC643CB347B00418D09" box="[503,598,1108,1132]" name="Réunion" pageId="3" pageNumber="136">Réunion</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC6405A347B011B8D09" author="Clastrier, J." box="[614,780,1108,1132]" firstAuthor="Clastrier" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="3" pageNumber="136" pagination="412 - 446" part="37" refId="ref6197" refString="Clastrier, J. (1959) Notes sur les Ceratopogonides. 8. Ceratopogonides de l'Ile de la Reunion. Archives de l'Institut Pasteur d'Algerie, 37, 412 - 446." title="Notes sur les Ceratopogonides. 8. Ceratopogonides de l'Ile de la Reunion" type="journal article" year="1959">Clastrier 1959</bibRefCitation>
). The record from
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(
<bibRefCitation id="E423F1EAFFFDFFC6466F347B076F8D09" author="de Meillon, B. &amp; Wirth, W. W." box="[1107,1400,1108,1132]" firstAuthor="de Meillon" journalOrPublisher="Annals of the Natal Museum" pageId="3" pageNumber="136" pagination="525 - 561" part="24" refId="ref6339" refString="de Meillon, B. &amp; Wirth, W. W. (1981) Subsaharan Ceratopogonidae (Diptera) VI. New species and records of South African biting midges collected by A. L. Dyce. Annals of the Natal Museum, 24, 525 - 561." title="Subsaharan Ceratopogonidae (Diptera) VI. New species and records of South African biting midges collected by A. L. Dyce" type="journal article" year="1981">de Meillon &amp; Wirth 1981</bibRefCitation>
) is doubtful. This species is new for the Lebanese fauna.
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122
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<emphasis id="B2C65009FFFDFFC642AB364903D08FE5" bold="true" box="[151,455,1638,1664]" italics="true" pageId="3" pageNumber="136">Dasyhelea alboverrucosa</emphasis>
Remm
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<emphasis id="B2C65009FFFDFFC642AB3682038D8FA6" box="[151,410,1709,1731]" italics="true" pageId="3" pageNumber="136">Dasyhelea alboverrucosa</emphasis>
<treatmentCitation id="0113AA0AFFFDFFC6439D3681005D8FA6" author="Remm" box="[417,586,1709,1731]" page="17" pageId="3" pageNumber="136" year="1967">
<bibRefCitation id="E423F1EAFFFDFFC6439D3681005D8FA6" author="Remm, H." box="[417,586,1709,1731]" firstAuthor="Remm" journalOrPublisher="Tartu Riikliku Ulikodi Toimetised" pageId="3" pageNumber="136" pagination="3 - 37" part="194" refId="ref7303" refString="Remm, H. (1967) On the fauna of Ceratopogonidae (Diptera) in the Caucasus. Tartu Riikliku Ulikodi Toimetised, 194, 3 - 37." title="On the fauna of Ceratopogonidae (Diptera) in the Caucasus" type="journal article" year="1967">Remm, 1967: 17</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F8A5CC8BFFFDFFC6406B368200DA8FA6" box="[599,717,1709,1731]" name="Azerbaijan" pageId="3" pageNumber="136">Azerbaijan</collectingCountry>
)
</taxonomicName>
.
</treatmentCitationGroup>
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<emphasis id="B2C65009FFFDFFC642AB36C003ED8E6D" bold="true" box="[151,506,1775,1800]" pageId="3" pageNumber="136">
Material examined.
<collectingCountry id="F8A5CC8BFFFDFFC643B536DF03E28E6D" box="[393,501,1776,1800]" name="Lebanon" pageId="3" pageNumber="136">Lebanon</collectingCountry>
.
</emphasis>
Tyre, N 33°
<geoCoordinate id="E586EADCFFFDFFC640B836C000FB8E62" box="[644,748,1775,1799]" direction="east" orientation="longitude" pageId="3" pageNumber="136" precision="55" value="15.535">15.535 E</geoCoordinate>
35°12.726,
<date id="F40CAADBFFFDFFC6414B36C001FB8E62" box="[887,1004,1775,1799]" pageId="3" pageNumber="136" value="2012-05-06">06.V.2012</date>
, beach, net,
<specimenCount id="96B44792FFFDFFC6464636C006C48E62" box="[1146,1235,1775,1799]" pageId="3" pageNumber="136" type="male">2 males</specimenCount>
, leg. P. Dominiak (LU &amp; UG).
</paragraph>
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<emphasis id="B2C65009FFFDFFC642FB371803778E35" bold="true" box="[199,352,1847,1872]" pageId="3" pageNumber="136">Distribution.</emphasis>
The species is known from
<collectingCountry id="F8A5CC8BFFFDFFC640A3371801108E2A" box="[671,775,1847,1871]" name="Romania" pageId="3" pageNumber="136">Romania</collectingCountry>
,
<collectingCountry id="F8A5CC8BFFFDFFC6412E3718017B8E2A" box="[786,876,1847,1871]" name="Ukraine" pageId="3" pageNumber="136">Ukraine</collectingCountry>
(Crimea),
<collectingCountry id="F8A5CC8BFFFDFFC641D43718067D8E2A" box="[1000,1130,1847,1871]" name="Kazakhstan" pageId="3" pageNumber="136">Kazakhstan</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC64645371802C48E11" author="Dominiak, P. &amp; Szadziewski, R." firstAuthor="Dominiak" journalOrPublisher="Zootaxa" pageId="3" pageNumber="136" pagination="1 - 37" part="2437" refId="ref6594" refString="Dominiak, P. &amp; Szadziewski, R. (2010) Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). Zootaxa, 2437, 1 - 37." title="Distribution and new synonymy in European biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2010">Dominiak &amp; Szadziewski 2010</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFDFFC642D5377303308E11" box="[233,295,1884,1908]" name="Israel" pageId="3" pageNumber="136">Israel</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC64304377300248E11" author="Dominiak, P. &amp; Braverman, Y. &amp; Szadziewski, R." box="[312,563,1884,1908]" etAl="et al." firstAuthor="Dominiak" journalOrPublisher="Polish Journal of Entomology" pageId="3" pageNumber="136" pagination="257 - 265" part="76" refId="ref6499" refString="Dominiak, P., Braverman, Y. &amp; Szadziewski, R. (2007) New records of biting midges of the genus Dasyhelea from Israel, with notes on synonymy in the genera Culicoides and Forcipomyia (Diptera: Ceratopogonidae). Polish Journal of Entomology, 76, 257 - 265." title="New records of biting midges of the genus Dasyhelea from Israel, with notes on synonymy in the genera Culicoides and Forcipomyia (Diptera: Ceratopogonidae)" type="journal article" year="2007">
Dominiak
<emphasis id="B2C65009FFFDFFC6438F377203F18E11" box="[435,486,1884,1908]" italics="true" pageId="3" pageNumber="136">et al</emphasis>
. 2007
</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFDFFC64075377300D38E11" box="[585,708,1884,1908]" name="Azerbaijan" pageId="3" pageNumber="136">Azerbaijan</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC640E9377301738E11" author="Remm, H." box="[725,868,1884,1908]" firstAuthor="Remm" journalOrPublisher="Tartu Riikliku Ulikodi Toimetised" pageId="3" pageNumber="136" pagination="3 - 37" part="194" refId="ref7303" refString="Remm, H. (1967) On the fauna of Ceratopogonidae (Diptera) in the Caucasus. Tartu Riikliku Ulikodi Toimetised, 194, 3 - 37." title="On the fauna of Ceratopogonidae (Diptera) in the Caucasus" type="journal article" year="1967">Remm 1967</bibRefCitation>
) and
<collectingCountry id="F8A5CC8BFFFDFFC64194377306268E11" box="[936,1073,1884,1908]" name="Afghanistan" pageId="3" pageNumber="136">Afghanistan</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFDFFC6467E377306DB8E11" author="Navai, S." box="[1090,1228,1884,1908]" firstAuthor="Navai" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="3" pageNumber="136" pagination="357 - 399" part="41" refId="ref7222" refString="Navai, S. (1994) Biting midges of the genus Dasyhelea from Afghanistan, with descriptions of new species (Diptera, Ceratopogonidae). Deutsche Entomologische Zeitschrift, 41, 357 - 399. http: // dx. doi. org / 10.1002 / mmnd. 19940410207" title="Biting midges of the genus Dasyhelea from Afghanistan, with descriptions of new species (Diptera, Ceratopogonidae)" type="journal article" year="1994">Navai 1994</bibRefCitation>
). New species for
<collectingCountry id="F8A5CC8BFFFDFFC642AB375002EA8EF2" box="[151,253,1919,1943]" name="Lebanon" pageId="3" pageNumber="136">Lebanon</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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123
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Clastrier, Rioux &amp; Descous
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<emphasis id="B2C65009FFFFFFC442AB3185036E88A5" box="[151,377,426,448]" italics="true" pageId="1" pageNumber="134">Dasyhelea tibestiensis</emphasis>
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(
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)
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.
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<emphasis id="B2C65009FFFFFFC442AB31C203CF8B60" bold="true" box="[151,472,493,518]" pageId="1" pageNumber="134">
Material examined.
<collectingCountry id="F8A5CC8BFFFFFFC443B131C203C48B60" box="[397,467,493,517]" name="Israel" pageId="1" pageNumber="134">Israel</collectingCountry>
.
</emphasis>
Shezaf N.R., Nahal Shahaq, N 30˚
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35˚15.32,
<date id="F40CAADBFFFFFFC4467731C206F78B60" box="[1099,1248,493,517]" pageId="1" pageNumber="134" value="1999-08-13">13.VIII.1999</date>
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, leg. I. Yarom;
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, Malaise trap,
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, leg. I. Yarom.
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<paragraph id="800D8C1BFFFFFFC442FB321A01CE8B14" blockId="1.[151,1436,493,625]" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC442FB321A03778B2B" bold="true" box="[199,352,565,590]" pageId="1" pageNumber="134">Distribution.</emphasis>
<collectingCountry id="F8A5CC8BFFFFFFC44356321A03B28B28" box="[362,421,565,589]" name="Chad" pageId="1" pageNumber="134">Chad</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFFFFC4438B321A00B38B28" author="Clastrier, J. &amp; Rioux, J. A. &amp; Descous, S." box="[439,676,565,589]" etAl="et al." firstAuthor="Clastrier" journalOrPublisher="Archives de l'Institut Pasteur Algerie" pageId="1" pageNumber="134" pagination="49 - 98" part="39" refId="ref6230" refString="Clastrier, J., Rioux, J. A. &amp; Descous, S. (1961) Notes sur les Ceratopogonides. XII. Ceratopogonides de Nord-Tchad. Archives de l'Institut Pasteur Algerie, 39, 49 - 98." title="Notes sur les Ceratopogonides. XII. Ceratopogonides de Nord-Tchad" type="journal article" year="1961">
Clastrier
<emphasis id="B2C65009FFFFFFC440183219004F8B28" box="[548,600,565,589]" italics="true" pageId="1" pageNumber="134">et al</emphasis>
. 1961
</bibRefCitation>
),
<collectingCountry id="F8A5CC8BFFFFFFC440873219011B8B28" box="[699,780,566,589]" name="Yemen" pageId="1" pageNumber="134">Yemen</collectingCountry>
,
<collectingCountry id="F8A5CC8BFFFFFFC44127321A01778B28" box="[795,864,565,589]" name="Oman" pageId="1" pageNumber="134">Oman</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFFFFC4414D321A06DD8B28" author="Boorman, J. &amp; van Harten, A." box="[881,1226,565,589]" firstAuthor="Boorman" journalOrPublisher="Fauna of Arabia" pageId="1" pageNumber="134" pagination="427 - 462" part="19" refId="ref6017" refString="Boorman, J. &amp; van Harten, A. (2002) Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen. Fauna of Arabia, 19, 427 - 462." title="Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen" type="journal article" year="2002">Boorman &amp; van Harten 2002</bibRefCitation>
), the
<collectingCountry id="F8A5CC8BFFFFFFC44731321A02EC8B14" name="United Arab Emirates" pageId="1" pageNumber="134">United Arab Emirates</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFFFFC443353276000F8B14" author="Szadziewski, R. &amp; Gwizdalska-Kentzer, M. &amp; Gilka, W." box="[265,536,601,625]" editor="van Harten, A." etAl="et al." firstAuthor="Szadziewski" journalOrPublisher="Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE" pageId="1" pageNumber="134" pagination="636 - 653" refId="ref7546" refString="Szadziewski, R., Gwizdalska-Kentzer, M. &amp; Gilka, W. (2011) Order Diptera, family Ceratopogonidae. In: Sheikh Tahnoon Bin Zayed Al Nahyan, H. H. (patron), van Harten, A. (Ed.), Arthropod fauna of the United Arab Emirates. Volume 4. Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE, pp. 636 - 653." title="Order Diptera, family Ceratopogonidae" type="book chapter" volumeTitle="Arthropod fauna of the United Arab Emirates. Volume 4" year="2011">
Szadziewski
<emphasis id="B2C65009FFFFFFC443A1327503D88B14" box="[413,463,601,625]" italics="true" pageId="1" pageNumber="134">et al</emphasis>
. 2011
</bibRefCitation>
). Reported for the first time from
<collectingCountry id="F8A5CC8BFFFFFFC441AF327601C28B14" box="[915,981,601,625]" name="Israel" pageId="1" pageNumber="134">Israel</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

191
data/08/1B/3D/081B3D0DFFFFFFC442AB332503348E58.xml Normal file
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@ -0,0 +1,191 @@
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<mods:title id="B2EA9EEB6CD8B460FA5025CAFB9540FA">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<treatment id="081B3D0DFFFFFFC442AB332503348E58" ID-DOI="http://doi.org/10.5281/zenodo.4499930" ID-GBIF-Taxon="119571703" ID-Zenodo-Dep="4499930" LSID="urn:lsid:zoobank.org:act:3BABE4D9-78F6-481A-B9BD-C3137E990676" httpUri="http://treatment.plazi.org/id/081B3D0DFFFFFFC442AB332503348E58" lastPageNumber="134" pageId="1" pageNumber="134">
<subSubSection id="C8A8DF90FFFFFFC442AB3325031D8A20" pageId="1" pageNumber="134" type="nomenclature">
<paragraph id="800D8C1BFFFFFFC442AB3325004D8A41" blockId="1.[151,602,778,837]" box="[151,602,778,804]" pageId="1" pageNumber="134">
<heading id="DB453B77FFFFFFC442AB3325004D8A41" bold="true" box="[151,602,778,804]" fontSize="11" level="1" pageId="1" pageNumber="134" reason="1">
<emphasis id="B2C65009FFFFFFC442AB3325004D8A41" bold="true" box="[151,602,778,804]" pageId="1" pageNumber="134">
<taxonomicName id="47B2F798FFFFFFC442AB332503E28A41" ID-CoL="9HN78" authority="Dominiak" authorityName="Dominiak" box="[151,501,778,804]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="antonii" status="sp. nov.">
<emphasis id="B2C65009FFFFFFC442AB332503658A41" bold="true" box="[151,370,778,804]" italics="true" pageId="1" pageNumber="134">Dasyhelea antonii</emphasis>
Dominiak
</taxonomicName>
<taxonomicNameLabel id="A9F5ED72FFFFFFC443C03325004D8A41" box="[508,602,778,804]" pageId="1" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="800D8C1BFFFFFFC442AB3302031D8A20" blockId="1.[151,602,778,837]" box="[151,266,813,837]" pageId="1" pageNumber="134">
(
<figureCitation id="1889909EFFFFFFC442A3330203158A20" box="[159,258,813,837]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="1" pageNumber="134">Figs 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFFFFC442AB335A00B08D78" pageId="1" pageNumber="134" type="materials_examined">
<paragraph id="800D8C1BFFFFFFC442AB335A00B08D78" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC442AB335A03518AEB" bold="true" box="[151,326,885,910]" pageId="1" pageNumber="134">
<typeStatus id="5F0932B9FFFFFFC442AB335A02C58AE8" box="[151,210,885,909]" pageId="1" pageNumber="134">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5F0932B9FFFFFFC44372335A03AC8AE8" box="[334,443,885,909]" pageId="1" pageNumber="134" type="holotype">Holotype</typeStatus>
: adult male.
<collectingCountry id="F8A5CC8BFFFFFFC44070335A014E8AEB" box="[588,857,885,910]" name="United Arab Emirates" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC44070335A014E8AEB" bold="true" box="[588,857,885,910]" pageId="1" pageNumber="134">United Arab Emirates</emphasis>
</collectingCountry>
. Sharjah Desert Park, N 25˚17' E 55˚42',
<quantity id="474A21FEFFFFFFC4470F335A07798AE8" box="[1331,1390,885,909]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="134" unit="m" value="70.0">70 m</quantity>
asl,
<date id="F40CAADBFFFFFFC442AB33B603378AD4" box="[151,288,921,945]" pageId="1" pageNumber="134" value="2008-12-15">15.XII.2008</date>
–
<date id="F40CAADBFFFFFFC4430D33B6038C8AD4" box="[305,411,921,945]" pageId="1" pageNumber="134" value="2009-01-12">12.I.2009</date>
, light trap, leg. A. van Harten (UG).
<typeStatus id="5F0932B9FFFFFFC4415E33B501C48AD7" box="[866,979,922,946]" pageId="1" pageNumber="134" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B2C65009FFFFFFC441DF33B606E88AD7" bold="true" box="[995,1279,921,946]" pageId="1" pageNumber="134">
<collectingCountry id="F8A5CC8BFFFFFFC441DF33B606ED8AD7" box="[995,1274,921,946]" name="United Arab Emirates" pageId="1" pageNumber="134">United Arab Emirates</collectingCountry>
.
</emphasis>
Al-Ajban, N 24˚36' E 55˚01',
<quantity id="474A21FEFFFFFFC4436D3392039D8AB0" box="[337,394,957,981]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="134" unit="m" value="60.0">60 m</quantity>
asl,
<date id="F40CAADBFFFFFFC44383339200678AB0" box="[447,624,957,981]" pageId="1" pageNumber="134" value="2006-01-30" valueMax="2006-02-26" valueMin="2006-01-30">
30.I–26.
<bibRefCitation id="E423F1EAFFFFFFC44027339200678AB0" author="II" box="[539,624,957,981]" pageId="1" pageNumber="134" refString="Dominiak, P. &amp; Szadziewski, R. (2006) Biting midges of the genus Dasyhelea Kieffer, 1911 (Diptera: Ceratopogonidae) collected from Polish peat-bogs. Dipteron, 22, 4 - 7." type="journal article" year="2006">II.2006</bibRefCitation>
</date>
, Malaise trap,
<specimenCount id="96B44792FFFFFFC44129339201798AB0" box="[789,878,957,981]" pageId="1" pageNumber="134" type="male">2 males</specimenCount>
, leg. A. van Harten (UG). Sharjah Desert Park, N 25˚17' E 55˚42',
<quantity id="474A21FEFFFFFFC4436533CE03828A9C" box="[345,405,993,1017]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="134" unit="m" value="70.0">70 m</quantity>
asl,
<date id="F40CAADBFFFFFFC443F233CE00418A9C" box="[462,598,993,1017]" pageId="1" pageNumber="134" value="2008-12-11">11.XII.2008</date>
–
<date id="F40CAADBFFFFFFC4405A33CE00C58A9C" box="[614,722,993,1017]" pageId="1" pageNumber="134" value="2009-01-06">06.I.2009</date>
, light trap,
<specimenCount id="96B44792FFFFFFC4416633CE01BD8A9C" box="[858,938,993,1017]" pageId="1" pageNumber="134" type="male">1 male</specimenCount>
, leg. A. van Harten (UG);
<date id="F40CAADBFFFFFFC446DA33CE07808A9C" box="[1254,1431,993,1017]" pageId="1" pageNumber="134" value="2009-01-12" valueMax="2009-02-02" valueMin="2009-01-12">12.I–02.II.2009</date>
, light trap,
<specimenCount id="96B44792FFFFFFC44336342A034F8D78" box="[266,344,1029,1053]" pageId="1" pageNumber="134" type="male">1 male</specimenCount>
, leg. A. van Harten (
<collectionCode id="E6A314DEFFFFFFC4407D342A008B8D78" box="[577,668,1029,1053]" pageId="1" pageNumber="134">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFFFFC442FB340607078D00" pageId="1" pageNumber="134" type="diagnosis">
<paragraph id="800D8C1BFFFFFFC442FB340607078D00" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC442FB340603568D27" bold="true" box="[199,321,1065,1090]" pageId="1" pageNumber="134">Diagnosis.</emphasis>
<taxonomicName id="47B2F798FFFFFFC443703406000B8D24" box="[332,540,1065,1089]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="antonii">
<emphasis id="B2C65009FFFFFFC443703406000B8D24" box="[332,540,1065,1089]" italics="true" pageId="1" pageNumber="134">Dasyhelea antonii</emphasis>
</taxonomicName>
is distinct in having a fifth palpal segment nearly two times longer than the fourth one, as well as a concave and well sclerotized posterior margin of 9th sternite of male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFFFFC442FB345E06DC8F00" pageId="1" pageNumber="134" type="description">
<paragraph id="800D8C1BFFFFFFC442FB345E00E68C9C" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC442FB345E03B48DEF" bold="true" box="[199,419,1137,1162]" pageId="1" pageNumber="134">Description. Male.</emphasis>
Head. Distal flagellomeres with two rows of setae (
<figureCitation id="1889909EFFFFFFC441DB345E063F8DEC" box="[999,1064,1137,1161]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="1" pageNumber="134">Fig. 1</figureCitation>
). Total length of flagellum
<quantity id="474A21FEFFFFFFC44761345E02E88DC8" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="5.8" metricValueMin="5.6" pageId="1" pageNumber="134" unit="mm" value="0.57" valueMax="0.58" valueMin="0.56">0.56– 0.58 mm</quantity>
, AR 0.89–0.94 (n=2). Clypeus entire with lateral lobes, bearing from 8 to 10 strong setae. Third palpal segment slender, length 60–72 μm, PR III 4.28–5.64 (n=4). Fifth palpal segment elongate, 1.7–2 times longer than fourth segment, with long setae located also in middle and at base (
<figureCitation id="1889909EFFFFFFC4419334CF01E08D9D" box="[943,1015,1248,1272]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="1" pageNumber="134">Fig. 2</figureCitation>
). Thorax. Transverse suture present, number of supraalar setae varing from 6 to 8. Scutellum yellow, with 7–8 strong and few smaller setae. Wing length
<quantity id="474A21FEFFFFFFC442DB350703878C25" box="[231,400,1320,1344]" metricMagnitude="-4" metricUnit="m" metricValue="8.85" metricValueMax="9.9" metricValueMin="7.8" pageId="1" pageNumber="134" unit="mm" value="0.885" valueMax="0.99" valueMin="0.78">0.78–0.99 mm</quantity>
, CR 0.44–0.45 (n=4), two radial cells present. Legs unicolor,
<collectingCountry id="F8A5CC8BFFFFFFC4465D350606938C25" box="[1121,1156,1321,1344]" name="Chile" pageId="1" pageNumber="134">TR</collectingCountry>
I 1.9–2.1 (n=4),
<collectingCountry id="F8A5CC8BFFFFFFC4477D350607728C25" box="[1345,1381,1321,1344]" name="Chile" pageId="1" pageNumber="134">TR</collectingCountry>
II 2.0 (n=2),
<collectingCountry id="F8A5CC8BFFFFFFC442DF357F03108C02" box="[227,263,1360,1383]" name="Chile" pageId="1" pageNumber="134">TR</collectingCountry>
III 1.8–2.0 (n=4). Genitalia (
<figureCitation id="1889909EFFFFFFC44000356000978C02" box="[572,640,1359,1383]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="1" pageNumber="134">Fig. 3</figureCitation>
). Tergite 9 rounded distally, with a pair of short apicolateral processes. Posterior margin of sternite 9 concave and well sclerotized, with very small median projections. Gonostylus long, slightly arched. Parameres asymmetrical. Posterior process broadly fused to right basal arm, long and slender, gradually tapering distally, markedly curved on apical portion. Arch of aedeagus low. Submedian projections stout, triangular, highly sclerotized only on ventral margins.
</paragraph>
<paragraph id="800D8C1BFFFFFFC442FB362900498F78" blockId="1.[151,1437,885,1853]" box="[199,606,1541,1566]" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC442FB362903F68F7B" bold="true" box="[199,481,1541,1566]" pageId="1" pageNumber="134">Female and immatures.</emphasis>
Unknown.
</paragraph>
<paragraph id="800D8C1BFFFFFFC442FB360606DC8F00" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC442FB360603FF8F27" bold="true" box="[199,488,1577,1602]" pageId="1" pageNumber="134">Derivation of the name.</emphasis>
This species is named after Antonius van Harten, the leader of the entomological projects in the Middle East, who collected and kindly sent us a lot of materials of biting midges.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFFFFC442FB365E00808FEC" box="[199,663,1649,1674]" pageId="1" pageNumber="134" type="distribution">
<paragraph id="800D8C1BFFFFFFC442FB365E00808FEC" blockId="1.[151,1437,885,1853]" box="[199,663,1649,1674]" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC442FB365E03778FEF" bold="true" box="[199,352,1649,1674]" pageId="1" pageNumber="134">Distribution.</emphasis>
The
<collectingCountry id="F8A5CC8BFFFFFFC443A6365E00848FEC" box="[410,659,1649,1673]" name="United Arab Emirates" pageId="1" pageNumber="134">United Arab Emirates</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFFFFC442FB36BA03348E58" pageId="1" pageNumber="134" type="discussion">
<paragraph id="800D8C1BFFFFFFC442FB36BA03348E58" blockId="1.[151,1437,885,1853]" pageId="1" pageNumber="134">
<emphasis id="B2C65009FFFFFFC442FB36BA03588FCB" bold="true" box="[199,335,1685,1710]" pageId="1" pageNumber="134">Comments.</emphasis>
Among all other species of this subgenus,
<taxonomicName id="47B2F798FFFFFFC4416236B906278FCB" box="[862,1072,1686,1710]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="antonii">
<emphasis id="B2C65009FFFFFFC4416236B906278FCB" box="[862,1072,1686,1710]" italics="true" pageId="1" pageNumber="134">Dasyhelea antonii</emphasis>
</taxonomicName>
is characterized in having the posterior margin of sternite 9 concave and strongly chitinized, and the aedeagus with prominent submedian projections. The species resembles
<taxonomicName id="47B2F798FFFFFFC4401236F001248F90" authority="Winnertz" authorityName="Winnertz" box="[558,819,1757,1782]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="B2C65009FFFFFFC4401236F000558F93" box="[558,578,1759,1782]" italics="true" pageId="1" pageNumber="134">D</emphasis>
.
<emphasis id="B2C65009FFFFFFC4406F36F100A78F93" box="[595,688,1758,1782]" italics="true" pageId="1" pageNumber="134">modesta</emphasis>
(Winnertz)
</taxonomicName>
and
<taxonomicName id="47B2F798FFFFFFC4414C36F006558F90" authority="Clastrier" authorityName="Clastrier" box="[880,1090,1757,1782]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="wirthi">
<emphasis id="B2C65009FFFFFFC4414C36F001938F93" box="[880,900,1759,1782]" italics="true" pageId="1" pageNumber="134">D</emphasis>
.
<emphasis id="B2C65009FFFFFFC441A836F101C08F93" box="[916,983,1758,1782]" italics="true" pageId="1" pageNumber="134">wirthi</emphasis>
Clastrier
</taxonomicName>
(known only from female) in having a similar shape of the fifth palpal segment that is elongate, but in
<taxonomicName id="47B2F798FFFFFFC441EA372C065E8E7F" box="[982,1097,1794,1818]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="antonii">
<emphasis id="B2C65009FFFFFFC441EA372C01FD8E7F" box="[982,1002,1795,1818]" italics="true" pageId="1" pageNumber="134">D</emphasis>
.
<emphasis id="B2C65009FFFFFFC441C5372C065E8E7F" box="[1017,1097,1795,1818]" italics="true" pageId="1" pageNumber="134">antonii</emphasis>
</taxonomicName>
bears additional long setae in median part.
</paragraph>
</subSubSection>
</treatment>
</document>