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<document id="D552DDD4483B97709AB8550DCF283141" ID-DOI="10.11646/zootaxa.5613.2.6" ID-ISSN="1175-5326" ID-ZooBank="57EC4EB7-22AE-4C2B-9CBC-5466D047E1CD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1744668308653" checkinUser="plazi" docAuthor="Jaroenchaiwattanachote, Chavanut, Pramual, Pairot, Wangwasit, Khanit, Bunchalee, Pasakorn &amp; Thanee, Isara" docDate="2025" docId="CF14316547104535FF5A02B61FF3EEDE" docLanguage="en" docName="zootaxa.5613.2.6.pdf" docOrigin="Zootaxa 5613 (2)" docSource="https://doi.org/10.11646/zootaxa.5613.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Triplectides buengkanensis Jaroenchaiwattanachote, Pramual, Wangwasit, Bunchalee &amp; Thanee, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="318" masterDocId="332D491D4716453EFFCD04421E7FED5F" masterDocTitle="Integrative taxonomy and DNA barcoding of Thai Caddisflies (Trichoptera), with the description of a new Species" masterLastPageNumber="322" masterPageNumber="307" pageNumber="313" updateTime="1744670767338" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="996F4BB4E33E931F455E11D4C7847D08">Integrative taxonomy and DNA barcoding of Thai Caddisflies (Trichoptera), with the description of a new Species</mods:title>
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<mods:namePart id="21B0F104B54BA615BDD0B9244EB24FC7">Jaroenchaiwattanachote, Chavanut</mods:namePart>
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<mods:affiliation id="993319D2BEFD74256A8584322D38B88B">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Mahasarakham 44150, Thailand.</mods:affiliation>
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<mods:namePart id="6C0675D5889638249D63B8FF64F33E85">Pramual, Pairot</mods:namePart>
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<mods:affiliation id="D62B8906B276692BA07AA33F00B10F8E">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Mahasarakham 44150, Thailand.</mods:affiliation>
<mods:nameIdentifier id="FB5F9F81F7FBD015DC784A286B0316DB" type="email">pairot.p@msu.ac.th</mods:nameIdentifier>
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<mods:namePart id="540C3F995F2CA97C7AFF9B1C811A5DFE">Wangwasit, Khanit</mods:namePart>
<mods:nameIdentifier id="C0CED11548C85F20EF4923449AB71B67" type="ORCID">0009-0002-0870-0549</mods:nameIdentifier>
<mods:affiliation id="4E7B78999C760E1835D4BA6ECE3499CD">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Mahasarakham 44150, Thailand.</mods:affiliation>
<mods:nameIdentifier id="7331B4172A049B94794151965AF76CB3" type="email">khanit.w@msu.ac.th</mods:nameIdentifier>
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<mods:namePart id="0C92E141AFBDB6AFAB80FDE38F2A7EDE">Bunchalee, Pasakorn</mods:namePart>
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<mods:affiliation id="A6D1C556AA1F2942A383B984EC7B9CDA">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Mahasarakham 44150, Thailand.</mods:affiliation>
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<mods:namePart id="FA2854EF5BE305B3B74C52859494F7B9">Thanee, Isara</mods:namePart>
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<mods:affiliation id="9A485690CD73E7E00712069BB586B8E4">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Mahasarakham 44150, Thailand.</mods:affiliation>
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<treatment id="CF14316547104535FF5A02B61FF3EEDE" LSID="urn:lsid:plazi:treatment:CF14316547104535FF5A02B61FF3EEDE" httpUri="http://treatment.plazi.org/id/CF14316547104535FF5A02B61FF3EEDE" lastPageId="11" lastPageNumber="318" pageId="6" pageNumber="313">
<subSubSection id="0FA7D3F847104538FF5A02B61AEAEA50" box="[151,1173,1780,1807]" pageId="6" pageNumber="313" type="nomenclature">
<paragraph id="4702807347104538FF5A02B61AEAEA50" blockId="6.[151,1173,1780,1843]" box="[151,1173,1780,1807]" pageId="6" pageNumber="313">
<heading id="1C4A371F47104538FF5A02B61AEAEA50" bold="true" box="[151,1173,1780,1807]" fontSize="11" level="1" pageId="6" pageNumber="313" reason="1">
<emphasis id="75C95C6147104538FF5A02B61AEAEA50" bold="true" box="[151,1173,1780,1807]" pageId="6" pageNumber="313">
<taxonomicName id="80BDFBF047104538FF5A02B61F9FEA50" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[151,480,1780,1807]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="313" phylum="Arthropoda" rank="species" species="buengkanensis" status="sp. nov.">
<emphasis id="75C95C6147104538FF5A02B61F9FEA50" bold="true" box="[151,480,1780,1807]" italics="true" pageId="6" pageNumber="313">Triplectides buengkanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EFAE11A47104538FE2A02B71C3AEA50" box="[487,581,1781,1807]" pageId="6" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
Jaroenchaiwatthanachote, Pramual, &amp; Thanee
</emphasis>
</heading>
</paragraph>
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<subSubSection id="0FA7D3F847104538FF5A035B1F7FEA6C" box="[151,256,1817,1843]" pageId="6" pageNumber="313" type="description">
<paragraph id="4702807347104538FF5A035B1F7FEA6C" blockId="6.[151,1173,1780,1843]" box="[151,256,1817,1843]" pageId="6" pageNumber="313">
<figureCitation id="DF869CF647104538FF5A035B1F7FEA6C" box="[151,256,1817,1843]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="7.[151,250,1943,1967]" captionStartId-1="8.[151,250,1897,1921]" captionStartId-2="9.[151,250,1397,1421]" captionStartId-3="10.[151,250,1695,1719]" captionTargetBox-0="[192,1395,182,1919]" captionTargetBox-1="[151,1435,182,1873]" captionTargetBox-2="[151,1435,181,1374]" captionTargetBox-3="[151,1435,345,1671]" captionTargetId-0="figure-20@7.[192,1395,181,1919]" captionTargetId-1="figure-19@8.[151,1435,181,1873]" captionTargetId-2="figure-20@9.[151,1435,181,1374]" captionTargetId-3="figure-92@10.[151,1435,345,1671]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 3. Triplectides buengkanensis sp. nov., adult male, holotype. 3A, habitus, dorsal; 3B, venation of right forewing, dorsal; 3C, venation of right hind wing, dorsal. Scale bars: 3A = 2 mm; 3B, 3C = 1 mm." captionText-1="FIGURE 4. Triplectides buengkanensis sp. nov., adult female, paratype. 4A, habitus, dorsal; 4B, venation of right forewing, dorsal; 4C, venation of right hind wing, dorsal. Scale bars: 4A = 2 mm; 4B, 4C = 1 mm." captionText-2="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." captionText-3="FIGURE 6. Triplectides buengkanensis sp. nov., adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm." pageId="6" pageNumber="313">Figs. 3–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0FA7D3F847104537FF5A03231F44EA45" lastPageId="9" lastPageNumber="316" pageId="6" pageNumber="313" type="diagnosis">
<paragraph id="4702807347104537FF5A03231F44EA45" blockId="6.[151,1437,1889,2023]" lastBlockId="9.[151,1437,1576,2035]" lastPageId="9" lastPageNumber="316" pageId="6" pageNumber="313">
<emphasis id="75C95C6147104538FF5A03231F6DEA24" bold="true" box="[151,274,1889,1915]" pageId="6" pageNumber="313">Diagnosis.</emphasis>
<taxonomicName id="80BDFBF047104538FED103231C30EA24" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[284,591,1889,1915]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="313" phylum="Arthropoda" rank="species" species="buengkanensis" status="sp. nov.">
<emphasis id="75C95C6147104538FED103231C30EA24" box="[284,591,1889,1915]" italics="true" pageId="6" pageNumber="313">Triplectides buengkanensis</emphasis>
</taxonomicName>
<emphasis id="75C95C6147104538FD9403231CC9EA24" bold="true" box="[601,694,1889,1915]" pageId="6" pageNumber="313">
<taxonomicNameLabel id="6EFAE11A47104538FD9403231CC9EA24" box="[601,694,1889,1915]" pageId="6" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="80BDFBF047104538FC9803201AAFEA24" authority="(Matsumura 1931)" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[853,1232,1889,1915]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="313" phylum="Arthropoda" rank="species" species="misakianus">
<emphasis id="75C95C6147104538FC9803201D8DEA24" box="[853,1010,1889,1915]" italics="true" pageId="6" pageNumber="313">T. misakianus</emphasis>
(
<bibRefCitation id="232CFD8247104538FBCE03231AB7EA24" author="Matsumura, S." box="[1027,1224,1889,1915]" pageId="6" pageNumber="313" refId="ref8459" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan Empire. Tokoshoin, Tokyo, 1496 pp. [in Japanese]" type="book" year="1931">Matsumura 1931</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="232CFD8247104538FB2F03231BE8EA24" author="Kuranishi, R. B." box="[1250,1431,1889,1915]" pageId="6" pageNumber="313" pagination="43 - 44" refId="ref7841" refString="Kuranishi, R. B. (1999) A collecting record of the Triplectides misakiana (Matsumura) (Leptoceridae; Trichoptera) after 88 years from eastern Japan. The Nature and Insect, 34 (10), 43-44. [in Japanese]" type="journal article" year="1999">Kuranishi 1999</bibRefCitation>
;
<bibRefCitation id="232CFD8247104538FF5A03C71FA9EAC0" author="Katsuma, N. &amp; Kuranishi, R. B." box="[151,470,1925,1951]" pageId="6" pageNumber="313" pagination="234 - 242" refId="ref7601" refString="Katsuma, N. &amp; Kuranishi, R. B. (2016) Redescription of Triplectides misakianus (Matsumura 1931) (Trichoptera, Leptoceridae) in Japan with notes on its habitat. Zoosymposia, 10, 234-242. https://doi.org/10.11646/zoosymposia.10.1.21" type="journal article" year="2016">Katsuma &amp; Kuranishi 2016</bibRefCitation>
),
<taxonomicName id="80BDFBF047104538FE2103C41A2AEAC0" authority="Desiderio &amp; Pes (Desiderio et al. 2020)" box="[492,1109,1925,1951]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="313" phylum="Arthropoda" rank="species" species="nessimiani">
<emphasis id="75C95C6147104538FE2103C41CFEEAC0" box="[492,641,1926,1951]" italics="true" pageId="6" pageNumber="313">T. nessimiani</emphasis>
Desidério &amp; Pes (
<bibRefCitation id="232CFD8247104538FC9B03C71A32EAC0" author="Desiderio, G. R. &amp; Pes, A. M. &amp; Barcelos-Silva, P. &amp; Hamada, N." box="[854,1101,1925,1951]" pageId="6" pageNumber="313" pagination="1 - 11" refId="ref7053" refString="Desiderio, G. R., Pes, A. M., Barcelos-Silva, P. &amp; Hamada, N. (2020) Triplectides Kolenati (Trichoptera: Leptoceridae) from Brazil: A new species, new records and an identification key. European Journal of Taxonomy, 677, 1-11. https://doi.org/10.5852/ejt.2020.677" type="journal article" year="2020">
Desidério
<emphasis id="75C95C6147104538FC0303C41A7EEAC0" box="[974,1025,1925,1951]" italics="true" pageId="6" pageNumber="313">et al</emphasis>
. 2020
</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="80BDFBF047104538FB5903C41BE3EAFF" authority="Walker 1852" authorityName="Walker" authorityYear="1852" box="[1172,1436,1925,1952]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="313" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="75C95C6147104538FB5903C41B7CEAC0" box="[1172,1283,1925,1951]" italics="true" pageId="6" pageNumber="313">T. indicus</emphasis>
<bibRefCitation id="232CFD8247104538FAC603C71BE3EAFF" author="Walker, F." box="[1291,1436,1925,1952]" pageId="6" pageNumber="313" refId="ref9735" refString="Walker, F. (1852) Cataloque of the Specimens of Neuropterous Insects in the Collection of the British Museum, Part I: (Phryganides-Perlides). Order of the Trustees, London, 192 pp." type="book" year="1852">
Walker
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</taxonomicName>
in many characteristics including those of the forewings and hind wings and the shapes and lengths of the preanal appendages and of the basoventral and basomesal lobes and apically acute second articles of the inferior appendages. However,
<taxonomicName id="80BDFBF0471F4537FEC1026A1C42EB1D" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[268,573,1576,1602]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="buengkanensis" status="sp. nov.">
<emphasis id="75C95C61471F4537FEC1026A1C42EB1D" box="[268,573,1576,1602]" italics="true" pageId="9" pageNumber="316">Triplectides buengkanensis</emphasis>
</taxonomicName>
<emphasis id="75C95C61471F4537FD88026A1CE0EB1D" bold="true" box="[581,671,1576,1602]" pageId="9" pageNumber="316">
<taxonomicNameLabel id="6EFAE11A471F4537FD88026A1CE0EB1D" box="[581,671,1576,1602]" pageId="9" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="80BDFBF0471F4537FC19026B1A10EB1D" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[980,1135,1576,1602]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="misakianus">
<emphasis id="75C95C61471F4537FC19026B1A10EB1D" box="[980,1135,1576,1602]" italics="true" pageId="9" pageNumber="316">T. misakianus</emphasis>
</taxonomicName>
in the female by length of the fork I petiole (shorter in
<taxonomicName id="80BDFBF0471F4537FE16020F1C09EB39" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[475,630,1612,1638]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="misakianus">
<emphasis id="75C95C61471F4537FE16020F1C09EB39" box="[475,630,1612,1638]" italics="true" pageId="9" pageNumber="316">T. misakianus</emphasis>
</taxonomicName>
) and absence of the transverse vein
<emphasis id="75C95C61471F4537FBD8020F1A42EB39" box="[1045,1085,1612,1638]" italics="true" pageId="9" pageNumber="316">r -m</emphasis>
in the hind wing (present in
<taxonomicName id="80BDFBF0471F4537FA45020F1F6AEBD5" baseAuthorityName="Matsumura" baseAuthorityYear="1931" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="misakianus">
<emphasis id="75C95C61471F4537FA45020F1F6AEBD5" italics="true" pageId="9" pageNumber="316">T. misakianus</emphasis>
</taxonomicName>
). In the male,
<taxonomicName id="80BDFBF0471F4537FE7202321C8EEBD5" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[447,753,1648,1674]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="buengkanensis" status="sp. nov.">
<emphasis id="75C95C61471F4537FE7202321C8EEBD5" box="[447,753,1648,1674]" italics="true" pageId="9" pageNumber="316">Triplectides buengkanensis</emphasis>
</taxonomicName>
<emphasis id="75C95C61471F4537FD3702321D2AEBD5" bold="true" box="[762,853,1648,1674]" pageId="9" pageNumber="316">
<taxonomicNameLabel id="6EFAE11A471F4537FD3702321D2AEBD5" box="[762,853,1648,1674]" pageId="9" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from these species by tergum X being obliquely truncate in dorsal view (suboval in
<taxonomicName id="80BDFBF0471F4537FD3E02D71D1FEBF1" authorityName="Walker" authorityYear="1852" box="[755,864,1684,1710]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="75C95C61471F4537FD3E02D71D1FEBF1" box="[755,864,1684,1710]" italics="true" pageId="9" pageNumber="316">T. indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="80BDFBF0471F4537FC5B02D71A4FEBF1" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[918,1072,1684,1710]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="misakianus">
<emphasis id="75C95C61471F4537FC5B02D71A4FEBF1" box="[918,1072,1684,1710]" italics="true" pageId="9" pageNumber="316">T. misakianus</emphasis>
</taxonomicName>
) and having a shallow V-shaped apicomesal incision (deep in
<taxonomicName id="80BDFBF0471F4537FE2E02FB1C01EB8D" baseAuthorityName="Matsumura" baseAuthorityYear="1931" box="[483,638,1720,1746]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="misakianus">
<emphasis id="75C95C61471F4537FE2E02FB1C01EB8D" box="[483,638,1720,1746]" italics="true" pageId="9" pageNumber="316">T. misakianus</emphasis>
</taxonomicName>
,
<taxonomicName id="80BDFBF0471F4537FD4002FB1D5DEB8D" box="[653,802,1721,1746]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="nessimiani">
<emphasis id="75C95C61471F4537FD4002FB1D5DEB8D" box="[653,802,1721,1746]" italics="true" pageId="9" pageNumber="316">T. nessimiani</emphasis>
</taxonomicName>
, and
<taxonomicName id="80BDFBF0471F4537FCAF02FB1DAFEB8D" authorityName="Walker" authorityYear="1852" box="[866,976,1720,1746]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="75C95C61471F4537FCAF02FB1DAFEB8D" box="[866,976,1720,1746]" italics="true" pageId="9" pageNumber="316">T. indicus</emphasis>
</taxonomicName>
), the absence of striae on the basomesal lobes of the inferior appendages (present in
<taxonomicName id="80BDFBF0471F4537FD48029F1D66EBA9" box="[645,793,1757,1782]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="nessimiani">
<emphasis id="75C95C61471F4537FD48029F1D66EBA9" box="[645,793,1757,1782]" italics="true" pageId="9" pageNumber="316">T. nessimiani</emphasis>
</taxonomicName>
), and the suboval phallotremal sclerite (subpentagonal in
<taxonomicName id="80BDFBF0471F4537FF5A03431F54EA45" box="[151,299,1793,1818]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="nessimiani">
<emphasis id="75C95C61471F4537FF5A03431F54EA45" box="[151,299,1793,1818]" italics="true" pageId="9" pageNumber="316">T. nessimiani</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="13C2D0FB47114539FF5A03D51A6CEA8C" pageId="7" pageNumber="314" startId="7.[151,250,1943,1967]" targetBox="[192,1395,182,1919]" targetPageId="7" targetType="figure">
<paragraph id="4702807347114539FF5A03D51A6CEA8C" blockId="7.[151,1437,1943,2004]" pageId="7" pageNumber="314">
<emphasis id="75C95C6147114539FF5A03D51F6AEAF0" bold="true" box="[151,277,1943,1967]" pageId="7" pageNumber="314">FIGURE 3.</emphasis>
<taxonomicName id="80BDFBF047114539FED303DA1C4BEAEF" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[286,564,1944,1968]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="314" phylum="Arthropoda" rank="species" species="buengkanensis" status="sp. nov.">
<emphasis id="75C95C6147114539FED303DA1C4BEAEF" box="[286,564,1944,1968]" italics="true" pageId="7" pageNumber="314">Triplectides buengkanensis</emphasis>
</taxonomicName>
<emphasis id="75C95C6147114539FDF003DA1CEFEAEF" bold="true" box="[573,656,1944,1968]" pageId="7" pageNumber="314">
<taxonomicNameLabel id="6EFAE11A47114539FDF003DA1CEFEAEF" box="[573,656,1944,1968]" pageId="7" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male, holotype. 3A, habitus, dorsal; 3B, venation of right forewing, dorsal; 3C, venation of right hind wing, dorsal. Scale bars: 3A = 2 mm; 3B, 3C = 1 mm.
</paragraph>
</caption>
<caption id="13C2D0FB471E4536FF5A032B1A6CEAFA" pageId="8" pageNumber="315" startId="8.[151,250,1897,1921]" targetBox="[151,1435,182,1873]" targetPageId="8" targetType="figure">
<paragraph id="47028073471E4536FF5A032B1A6CEAFA" blockId="8.[151,1437,1897,1957]" pageId="8" pageNumber="315">
<emphasis id="75C95C61471E4536FF5A032B1F6BEADE" bold="true" box="[151,276,1897,1921]" pageId="8" pageNumber="315">FIGURE 4.</emphasis>
<taxonomicName id="80BDFBF0471E4536FED1032B1C4FEADE" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[284,560,1897,1921]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="315" phylum="Arthropoda" rank="species" species="buengkanensis" status="sp. nov.">
<emphasis id="75C95C61471E4536FED1032B1C4FEADE" box="[284,560,1897,1921]" italics="true" pageId="8" pageNumber="315">Triplectides buengkanensis</emphasis>
</taxonomicName>
<emphasis id="75C95C61471E4536FDF5032B1CF5EADE" bold="true" box="[568,650,1897,1921]" pageId="8" pageNumber="315">
<taxonomicNameLabel id="6EFAE11A471E4536FDF5032B1CF5EADE" box="[568,650,1897,1921]" pageId="8" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female, paratype. 4A, habitus, dorsal; 4B, venation of right forewing, dorsal; 4C, venation of right hind wing, dorsal. Scale bars: 4A = 2 mm; 4B, 4C = 1 mm.
</paragraph>
</caption>
<caption id="13C2D0FB471F4537FF5A01371B03E88A" pageId="9" pageNumber="316" startId="9.[151,250,1397,1421]" targetBox="[151,1435,181,1374]" targetPageId="9" targetType="figure">
<paragraph id="47028073471F4537FF5A01371B03E88A" blockId="9.[151,1437,1397,1494]" pageId="9" pageNumber="316">
<emphasis id="75C95C61471F4537FF5A01371F6EE8D1" bold="true" box="[151,273,1397,1422]" pageId="9" pageNumber="316">FIGURE 5.</emphasis>
<taxonomicName id="80BDFBF0471F4537FEDA01341C57E8D1" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[279,552,1398,1422]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="316" phylum="Arthropoda" rank="species" species="buengkanensis" status="sp. nov.">
<emphasis id="75C95C61471F4537FEDA01341C57E8D1" box="[279,552,1398,1422]" italics="true" pageId="9" pageNumber="316">Triplectides buengkanensis</emphasis>
</taxonomicName>
<emphasis id="75C95C61471F4537FDE001341C03E8D1" bold="true" box="[557,636,1398,1422]" pageId="9" pageNumber="316">
<taxonomicNameLabel id="6EFAE11A471F4537FDE001341C03E8D1" box="[557,636,1398,1422]" pageId="9" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal. Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2
<superScript id="B0C82D3B471F4537FC0401FC1DA7E893" attach="left" box="[969,984,1470,1484]" fontSize="6" pageId="9" pageNumber="316">nd</superScript>
art. = second article. Scale bars: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="0FA7D3F8471F4537FF0A03661B0DEAF5" pageId="9" pageNumber="316" type="materials_examined">
<paragraph id="47028073471F4537FF0A03661AE2EAD9" blockId="9.[151,1437,1576,2035]" pageId="9" pageNumber="316">
<emphasis id="75C95C61471F4537FF0A03661FD0EA61" bold="true" box="[199,431,1828,1854]" pageId="9" pageNumber="316">Material examined.</emphasis>
<materialsCitation id="F7D58A2E471F4537FE7803661AE2EAD9" collectingDate="2023-02-04" collectorName="K. Wangwasit &amp; P. Bunchalee &amp; I. Thanee &amp; Mahasarakham University &amp; Thailand." country="Thailand" county="Bueng Khong Long District" elevation="168" latitude="18.02334" location="Khong Long" longLatPrecision="1" longitude="104.01569" municipality="Bueng" pageId="9" pageNumber="316" specimenCount="1" specimenCount-male="1" stateProvince="Bueng Kan" typeStatus="holotype">
<typeStatus id="98063ED1471F4537FE7803661C62EA61" box="[437,541,1828,1854]" pageId="9" pageNumber="316" type="holotype">Holotype</typeStatus>
.
<specimenCount id="51BB4BFA471F4537FDE703661C08EA61" box="[554,631,1828,1854]" count="1" pageId="9" pageNumber="316" type="male">1 male</specimenCount>
.
<collectingCountry id="3FAAC0E3471F4537FD4D03661D6AEA61" box="[640,789,1828,1854]" name="Thailand" pageId="9" pageNumber="316">THAILAND</collectingCountry>
:
<collectingRegion id="85794E91471F4537FCD303661A78EA61" box="[798,1031,1828,1854]" country="Thailand" name="Bueng Kan" pageId="9" pageNumber="316">Bueng Kan Province</collectingRegion>
,
<collectingCounty id="AE63F8FF471F4537FBDC03661B35EA61" box="[1041,1354,1828,1854]" pageId="9" pageNumber="316">Bueng Khong Long District</collectingCounty>
,
<collectingMunicipality id="A7661A09471F4537FA9E03661BE3EA61" box="[1363,1436,1828,1854]" pageId="9" pageNumber="316">Bueng</collectingMunicipality>
[= Swamp]
<location id="4262D6A8471F4537FED0030A1FD0EA3D" LSID="urn:lsid:plazi:treatment:CF14316547104535FF5A02B61FF3EEDE:4262D6A8471F4537FED0030A1FD0EA3D" box="[285,431,1864,1890]" country="Thailand" county="Bueng Khong Long District" latitude="18.02334" longLatPrecision="1" longitude="104.01569" municipality="Bueng" name="Khong Long" pageId="9" pageNumber="316" stateProvince="Bueng Kan">Khong Long</location>
;
<geoCoordinate id="2289E6B4471F4537FE76030A1C32EA3D" box="[443,589,1864,1890]" degrees="18.02334" direction="north" orientation="latitude" pageId="9" pageNumber="316" precision="1" value="18.02334">18.02334° N</geoCoordinate>
,
<geoCoordinate id="2289E6B4471F4537FD94030A1C89EA3D" box="[601,758,1864,1891]" degrees="104.01569" direction="east" orientation="longitude" pageId="9" pageNumber="316" precision="1" value="104.01569">104.01569° E</geoCoordinate>
;
<quantity id="80452D96471F4537FCCE030A1D32EA3D" box="[771,845,1864,1890]" metricMagnitude="2" metricUnit="m" metricValue="1.68" pageId="9" pageNumber="316" unit="m" value="168.0">
<elevation id="CC906740471F4537FCCE030A1D32EA3D" box="[771,845,1864,1890]" metricMagnitude="2" metricUnit="m" metricValue="1.68" pageId="9" pageNumber="316" unit="m" value="168.0">168 m</elevation>
</quantity>
;
<date id="3303A6B3471F4537FC94030A1DBFEA3D" box="[857,960,1864,1890]" pageId="9" pageNumber="316" value="2023-02-04">
<collectingDate id="23475F5B471F4537FC94030A1DBFEA3D" box="[857,960,1864,1890]" pageId="9" pageNumber="316" value="2023-02-04">4.ii.2023</collectingDate>
</date>
;
<collectingMethod id="9EFCF864471F4537FC01030A1A49EA3D" box="[972,1078,1864,1890]" pageId="9" pageNumber="316">light trap</collectingMethod>
;
<collectorName id="EA48E5A5471F4537FB8F030A1A9CEA3D" box="[1090,1251,1864,1890]" pageId="9" pageNumber="316">K. Wangwasit</collectorName>
,
<collectorName id="EA48E5A5471F4537FB23030A1B00EA3D" box="[1262,1407,1864,1890]" pageId="9" pageNumber="316">P. Bunchalee</collectorName>
&amp;
<collectorName id="EA48E5A5471F4537FF5A032E1F7BEAD9" box="[151,260,1900,1926]" pageId="9" pageNumber="316">I. Thanee</collectorName>
;
<location id="4262D6A8471F4537FEC2032E1C6BEAD9" LSID="urn:lsid:plazi:treatment:CF14316547104535FF5A02B61FF3EEDE:4262D6A8471F4537FEC2032E1C6BEAD9" box="[271,532,1900,1926]" country="Thailand" county="Bueng Khong Long District" latitude="18.02334" longLatPrecision="1" longitude="104.01569" municipality="Bueng" name="Department of Biology" pageId="9" pageNumber="316" stateProvince="Bueng Kan">Department of Biology</location>
,
<location id="4262D6A8471F4537FDD2032E1C8DEAD9" LSID="urn:lsid:plazi:treatment:CF14316547104535FF5A02B61FF3EEDE:4262D6A8471F4537FDD2032E1C8DEAD9" box="[543,754,1900,1926]" country="Thailand" county="Bueng Khong Long District" latitude="18.02334" longLatPrecision="1" longitude="104.01569" municipality="Bueng" name="Faculty of Science" pageId="9" pageNumber="316" stateProvince="Bueng Kan">Faculty of Science</location>
,
<collectorName id="EA48E5A5471F4537FD30032E1A58EAD9" box="[765,1063,1900,1926]" pageId="9" pageNumber="316">Mahasarakham University</collectorName>
,
<collectorName id="EA48E5A5471F4537FBFC032E1AE2EAD9" box="[1073,1181,1900,1926]" pageId="9" pageNumber="316">Thailand.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="47028073471F4537FF0A03D21B0DEAF5" blockId="9.[151,1437,1576,2035]" box="[199,1394,1936,1962]" pageId="9" pageNumber="316">
<materialsCitation id="F7D58A2E471F4537FF0A03D21B11EAF5" box="[199,1390,1936,1962]" collectingDate="2023-02-04" collectorName="K. Wangwasit &amp; P. Bunchalee &amp; I. Thanee &amp; Mahasarakham University &amp; Thailand." country="Thailand" county="Bueng Khong Long District" elevation="168" latitude="18.02334" location="Khong Long" longLatPrecision="1" longitude="104.01569" municipality="Bueng" pageId="9" pageNumber="316" specimenCount="15" specimenCount-female="4" specimenCount-male="11" stateProvince="Bueng Kan" typeStatus="paratype">
<typeStatus id="98063ED1471F4537FF0A03D21F4BEAF5" box="[199,308,1936,1962]" pageId="9" pageNumber="316" type="paratype">Paratypes</typeStatus>
. Same data as holotype, except
<specimenCount id="51BB4BFA471F4537FD5603D21D7EEAF5" box="[667,769,1936,1962]" count="11" pageId="9" pageNumber="316" type="male">11 males</specimenCount>
,
<specimenCount id="51BB4BFA471F4537FCC003D21D02EAF5" box="[781,893,1936,1962]" count="4" pageId="9" pageNumber="316" type="female">4 females</specimenCount>
; GenBank no: PQ009803–PQ009817 (COI)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0FA7D3F8471F4535FF0A03F61C74EE4A" lastPageId="11" lastPageNumber="318" pageId="9" pageNumber="316" type="description">
<paragraph id="47028073471F4534FF0A03F61D47EC1E" blockId="9.[151,1437,1576,2035]" lastBlockId="10.[151,1437,151,321]" lastPageId="10" lastPageNumber="317" pageId="9" pageNumber="316">
<emphasis id="75C95C61471F4537FF0A03F61F26EA91" bold="true" box="[199,345,1972,1998]" pageId="9" pageNumber="316">Description.</emphasis>
<emphasis id="75C95C61471F4537FE9303F61FA8EA91" box="[350,471,1972,1998]" italics="true" pageId="9" pageNumber="316">Adult male</emphasis>
(
<figureCitation id="DF869CF6471F4537FE2803F61C0EEA91" box="[485,625,1972,1998]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1395,182,1919]" captionTargetId="figure-20@7.[192,1395,181,1919]" captionTargetPageId="7" captionText="FIGURE 3. Triplectides buengkanensis sp. nov., adult male, holotype. 3A, habitus, dorsal; 3B, venation of right forewing, dorsal; 3C, venation of right hind wing, dorsal. Scale bars: 3A = 2 mm; 3B, 3C = 1 mm." pageId="9" pageNumber="316">Figs 3A–3C</figureCitation>
). Forewing length 11.0 mm–
<quantity id="80452D96471F4537FC7F03F61A6DEA91" box="[946,1042,1972,1999]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="316" unit="mm" value="12.5">12.5 mm</quantity>
(
<figureCitation id="DF869CF6471F4537FBED03F61A09EA91" box="[1056,1142,1972,1998]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[192,1395,182,1919]" captionTargetId="figure-20@7.[192,1395,181,1919]" captionTargetPageId="7" captionText="FIGURE 3. Triplectides buengkanensis sp. nov., adult male, holotype. 3A, habitus, dorsal; 3B, venation of right forewing, dorsal; 3C, venation of right hind wing, dorsal. Scale bars: 3A = 2 mm; 3B, 3C = 1 mm." pageId="9" pageNumber="316">Fig. 3B</figureCitation>
), body
<quantity id="80452D96471F4537FB0A03F61B47EA91" box="[1223,1336,1972,1999]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="9.0" metricValueMin="7.5" pageId="9" pageNumber="316" unit="mm" value="8.25" valueMax="9.0" valueMin="7.5">7.5–9 mm</quantity>
(
<emphasis id="75C95C61471F4537FA8A03F71B2AEA91" box="[1351,1365,1973,1998]" italics="true" pageId="9" pageNumber="316">n</emphasis>
= 12).
<emphasis id="75C95C61471F4537FF5A039A1F58EAAD" box="[151,295,2008,2034]" italics="true" pageId="9" pageNumber="316">Adult female</emphasis>
(
<figureCitation id="DF869CF6471F4537FEFB039A1FBAEAAD" box="[310,453,2008,2034]" captionStart="FIGURE 4" captionStartId="8.[151,250,1897,1921]" captionTargetBox="[151,1435,182,1873]" captionTargetId="figure-19@8.[151,1435,181,1873]" captionTargetPageId="8" captionText="FIGURE 4. Triplectides buengkanensis sp. nov., adult female, paratype. 4A, habitus, dorsal; 4B, venation of right forewing, dorsal; 4C, venation of right hind wing, dorsal. Scale bars: 4A = 2 mm; 4B, 4C = 1 mm." pageId="9" pageNumber="316">Figs 4A–4C</figureCitation>
). Forewing length
<quantity id="80452D96471F4537FD54039A1D58EAAD" box="[665,807,2008,2035]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.5" metricValueMin="1.14" pageId="9" pageNumber="316" unit="mm" value="13.2" valueMax="15.0" valueMin="11.4">11.4–15 mm</quantity>
(
<figureCitation id="DF869CF6471F4537FCFA039A1DEFEAAD" box="[823,912,2008,2034]" captionStart="FIGURE 4" captionStartId="8.[151,250,1897,1921]" captionTargetBox="[151,1435,182,1873]" captionTargetId="figure-19@8.[151,1435,181,1873]" captionTargetPageId="8" captionText="FIGURE 4. Triplectides buengkanensis sp. nov., adult female, paratype. 4A, habitus, dorsal; 4B, venation of right forewing, dorsal; 4C, venation of right hind wing, dorsal. Scale bars: 4A = 2 mm; 4B, 4C = 1 mm." pageId="9" pageNumber="316">Fig. 4B</figureCitation>
), body
<quantity id="80452D96471F4537FC29039A1A30EAAD" box="[996,1103,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="9" pageNumber="316" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
(
<emphasis id="75C95C61471F4537FBAD039B1A11EAAD" box="[1120,1134,2009,2034]" italics="true" pageId="9" pageNumber="316">n</emphasis>
= 11). In both sexes, body generally appearing yellowish brown on head, thorax, forewing, abdomen, and legs in alcohol-preserved material. Tibial spur formula 2:2:2—forelegs, midlegs, and hind legs respectively—in both sexes. Wing venation similar in male and female except male forewing with R4+5 straight behind discoidal cell and apical fork III absent, female forewing with R4+5 angled posterad subapically at
<emphasis id="75C95C61471C4534FD3C05411D66EC43" box="[753,793,259,284]" italics="true" pageId="10" pageNumber="317">r-m</emphasis>
crossvein behind discoidal cell as in several Australian species (
<bibRefCitation id="232CFD82471C4534FF3A05651F81EC1E" author="Morse, J. C. &amp; Neboiss, N." box="[247,510,295,321]" pageId="10" pageNumber="317" pagination="61 - 98" refId="ref8696" refString="Morse, J. C., Neboiss, N. (1982) Triplectides of Australia (Insecta: Trichoptera: Leptoceridae). Memoirs of the National Museum of Victoria, 43, 61-98. https://doi.org/10.24199/j.mmv.1982.43.05" type="journal article" year="1982">Morse &amp; Neboiss 1982</bibRefCitation>
) and apical fork III present.
</paragraph>
<caption id="13C2D0FB471C4534FF5A02DD1CF7EBA1" pageId="10" pageNumber="317" startId="10.[151,250,1695,1719]" targetBox="[151,1435,345,1671]" targetPageId="10" targetType="figure">
<paragraph id="47028073471C4534FF5A02DD1CF7EBA1" blockId="10.[151,1437,1695,1791]" pageId="10" pageNumber="317">
<emphasis id="75C95C61471C4534FF5A02DD1F69EBE8" bold="true" box="[151,278,1695,1719]" pageId="10" pageNumber="317">FIGURE 6.</emphasis>
<taxonomicName id="80BDFBF0471C4534FEED02DD1C49EBE8" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[288,566,1695,1719]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="317" phylum="Arthropoda" rank="species" species="buengkanensis" status="sp. nov.">
<emphasis id="75C95C61471C4534FEED02DD1C49EBE8" box="[288,566,1695,1719]" italics="true" pageId="10" pageNumber="317">Triplectides buengkanensis</emphasis>
</taxonomicName>
<emphasis id="75C95C61471C4534FD8D02DD1CECEBE8" bold="true" box="[576,659,1695,1719]" pageId="10" pageNumber="317">
<taxonomicNameLabel id="6EFAE11A471C4534FD8D02DD1CECEBE8" box="[576,659,1695,1719]" pageId="10" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="47028073471C4535FF0A03131C71EC8E" blockId="10.[151,1437,1873,2044]" lastBlockId="11.[151,1437,150,897]" lastPageId="11" lastPageNumber="318" pageId="10" pageNumber="317">
<emphasis id="75C95C61471C4534FF0A03131F14EA34" box="[199,363,1873,1899]" italics="true" pageId="10" pageNumber="317">Male genitalia</emphasis>
(
<figureCitation id="DF869CF6471C4534FEB403131C7BEA33" box="[377,516,1873,1900]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="10" pageNumber="317">Figs 5A–5E</figureCitation>
). Segment IX (IX), in lateral view, tall, longitudinally short, subrectangular, longer dorsally than ventrally, constricted at 3/4 height with acute incisions on anterior and posterior margins, posterior margins projecting and triangular at 2/3 height. In dorsal view, tergum IX subpentagonal with posterior margin almost rounded, with tiny notch and membranous digitate median processes (
<figureCitation id="DF869CF6471C4534FC7E03FF1A73EA87" box="[947,1036,1981,2008]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="10" pageNumber="317">Fig. 5B</figureCitation>
). Preanal appendages (pr. ap.) long and slender, 3/4 as long as tergum X, blunt apically, with many setae (
<figureCitation id="DF869CF6471C4534FC6503A31A4AEAA3" box="[936,1077,2017,2044]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="10" pageNumber="317">Figs 5A, 5B</figureCitation>
). Tergum X (X), in lateral view, wide basally, with dorsal and ventral margins almost straight, apex round (
<figureCitation id="DF869CF6471D4535FC1C04D51A55EDEE" box="[977,1066,150,177]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 5A</figureCitation>
); in dorsal view, slightly tapering posterad, bearing short lateral and apical setae, apex obliquely truncate, with shallow, V-shaped incision (
<figureCitation id="DF869CF6471D4535FAFA04F91BF0ED8A" box="[1335,1423,186,213]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 5B</figureCitation>
). In lateral view (
<figureCitation id="DF869CF6471D4535FE8B049D1FE1EDA6" box="[326,414,222,249]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 5A</figureCitation>
), apicodorsal lobe of first article of each inferior appendage long, extending beyond hooked second article (2nd art.) and tergum X, bearing long setae; in ventral view (
<figureCitation id="DF869CF6471D4535FC2D05411A47EC42" box="[992,1080,258,285]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 5C</figureCitation>
), first article broad basally with blunt subtriangular basomesal lobe, second article crescent-shaped, wide at base, tapering apically, sharply curved inward, with pointed apex. In lateral and ventral views (
<figureCitation id="DF869CF6471D4535FCC405091DE9EC3A" box="[777,918,330,357]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Figs 5A, 5C</figureCitation>
), basoventral lobe of each inferior appendage, long, shorter than preanal appendages, apically round, with long setae. In lateral view (
<figureCitation id="DF869CF6471D4535FBA3052D1AB7ECD6" box="[1134,1224,366,393]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 5D</figureCitation>
), phallic apparatus long, tubular, thicker near middle, curved upward apically; in dorsal view, bulb-like, with small phallotremal sclerite and pair of wide apices (
<figureCitation id="DF869CF6471D4535FE6705F51C7EEC8E" box="[426,513,438,465]" captionStart="FIGURE 5" captionStartId="9.[151,250,1397,1421]" captionTargetBox="[151,1435,181,1374]" captionTargetId="figure-20@9.[151,1435,181,1374]" captionTargetPageId="9" captionText="FIGURE 5. Triplectides buengkanensis sp. nov., adult male, holotype. 5A–5C, genitalia: 5A, left lateral; 5B, dorsal; 5C, ventral. 5D, 5E, phallic apparatus: 5D, left lateral; 5E, dorsal.Abbreviations: ap. lo. = apicodorsal lobe; bv. lo. = basoventral lobe; me. lo. = basomesal lobe; ph. sc. = phallotremal sclerite; pr. ap. = preanal appendages; 2nd art. = second article. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 5E</figureCitation>
).
</paragraph>
<paragraph id="47028073471D4535FF0A05991EA0EFAE" blockId="11.[151,1437,150,897]" pageId="11" pageNumber="318">
<emphasis id="75C95C61471D4535FF0A05991FF8ECAA" box="[199,391,475,501]" italics="true" pageId="11" pageNumber="318">Female genitalia</emphasis>
(
<figureCitation id="DF869CF6471D4535FE5B05991C5BECAA" box="[406,548,475,501]" captionStart="FIGURE 6" captionStartId="10.[151,250,1695,1719]" captionTargetBox="[151,1435,345,1671]" captionTargetId="figure-92@10.[151,1435,345,1671]" captionTargetPageId="10" captionText="FIGURE 6. Triplectides buengkanensis sp. nov., adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Figs 6A–6C</figureCitation>
). In lateral view (
<figureCitation id="DF869CF6471D4535FD2705991D3BECAA" box="[746,836,475,501]" captionStart="FIGURE 6" captionStartId="10.[151,250,1695,1719]" captionTargetBox="[151,1435,345,1671]" captionTargetId="figure-92@10.[151,1435,345,1671]" captionTargetPageId="10" captionText="FIGURE 6. Triplectides buengkanensis sp. nov., adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 6A</figureCitation>
), tergum IX subrectangular, with sclerotized concave area posterolaterally, sternum smaller; in dorsal view (
<figureCitation id="DF869CF6471D4535FD3E05BD1D36EF46" box="[755,841,511,537]" captionStart="FIGURE 6" captionStartId="10.[151,250,1695,1719]" captionTargetBox="[151,1435,345,1671]" captionTargetId="figure-92@10.[151,1435,345,1671]" captionTargetPageId="10" captionText="FIGURE 6. Triplectides buengkanensis sp. nov., adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 6B</figureCitation>
), subtriangular, round apically, and with pair of small papillae posteriorly. In lateral and dorsal views (
<figureCitation id="DF869CF6471D4535FD6306611D46EF62" box="[686,825,547,573]" captionStart="FIGURE 6" captionStartId="10.[151,250,1695,1719]" captionTargetBox="[151,1435,345,1671]" captionTargetId="figure-92@10.[151,1435,345,1671]" captionTargetPageId="10" captionText="FIGURE 6. Triplectides buengkanensis sp. nov., adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Figs 6A, 6C</figureCitation>
), appendages of segment X (X ap.) long and oval, with many setae; sensilla-bearing processes almost 1/2 as long as preanal appendages, blunt apically, each with 6 or 7 short, stout setae apically. In lateral view (
<figureCitation id="DF869CF6471D4535FDB506291CACEFDA" box="[632,723,619,645]" captionStart="FIGURE 6" captionStartId="10.[151,250,1695,1719]" captionTargetBox="[151,1435,345,1671]" captionTargetId="figure-92@10.[151,1435,345,1671]" captionTargetPageId="10" captionText="FIGURE 6. Triplectides buengkanensis sp. nov., adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 6A</figureCitation>
), valves (v.) semicircular, round apically, each with short setae ventrolaterally. In ventral view (
<figureCitation id="DF869CF6471D4535FDC706CD1C1AEFF6" box="[522,613,655,681]" captionStart="FIGURE 6" captionStartId="10.[151,250,1695,1719]" captionTargetBox="[151,1435,345,1671]" captionTargetId="figure-92@10.[151,1435,345,1671]" captionTargetPageId="10" captionText="FIGURE 6. Triplectides buengkanensis sp. nov., adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 6C</figureCitation>
), gonopod plate (go.pl.) sub-rectangular, with blunt anterolateral angles and round posteriorly. In ventral view (
<figureCitation id="DF869CF6471D4535FD9E06F11CD3EF92" box="[595,684,691,717]" captionStart="FIGURE 6" captionStartId="10.[151,250,1695,1719]" captionTargetBox="[151,1435,345,1671]" captionTargetId="figure-92@10.[151,1435,345,1671]" captionTargetPageId="10" captionText="FIGURE 6. Triplectides buengkanensis sp. nov., adult female genitalia, paratype. 6A, left lateral; 6B, dorsal; 6C, ventral. Abbreviations: X ap. = appendages of segment X; s-b. p. = sensilla-bearing processes; go. pl. = gonopod plate; v. = valves; v. a. = internal vaginal apparatus. Scale bars: 0.1 mm." pageId="11" pageNumber="318">Fig. 6C</figureCitation>
), internal vaginal apparatus (v. a.) subrectangular, twice as long as broad.
</paragraph>
<paragraph id="47028073471D4535FF0A06B91C74EE4A" blockId="11.[151,1437,150,897]" box="[199,523,763,789]" pageId="11" pageNumber="318">
<emphasis id="75C95C61471D4535FF0A06B91FE9EE4A" bold="true" box="[199,406,763,789]" pageId="11" pageNumber="318">Immature stages.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="0FA7D3F8471D4535FF0A075C1CB7EE66" box="[199,712,798,825]" pageId="11" pageNumber="318" type="distribution">
<paragraph id="47028073471D4535FF0A075C1CB7EE66" blockId="11.[151,1437,150,897]" box="[199,712,798,825]" pageId="11" pageNumber="318">
<emphasis id="75C95C61471D4535FF0A075C1F1EEE67" bold="true" box="[199,353,798,824]" pageId="11" pageNumber="318">Distribution.</emphasis>
<collectingCountry id="3FAAC0E3471D4535FEAA075D1FB4EE66" box="[359,459,799,825]" name="Thailand" pageId="11" pageNumber="318">Thailand</collectingCountry>
(Buengkan Province).
</paragraph>
</subSubSection>
<subSubSection id="0FA7D3F8471D4535FF0A07011FF3EEDE" pageId="11" pageNumber="318" type="etymology">
<paragraph id="47028073471D4535FF0A07011FF3EEDE" blockId="11.[151,1437,150,897]" pageId="11" pageNumber="318">
<emphasis id="75C95C61471D4535FF0A07011F31EE02" bold="true" box="[199,334,835,861]" pageId="11" pageNumber="318">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="80BDFBF0471D4535FD8D07011C9AEE02" authorityName="Jaroenchaiwattanachote &amp; Pramual &amp; Wangwasit &amp; Bunchalee &amp; Thanee" authorityYear="2025" box="[576,741,835,861]" class="Insecta" family="Leptoceridae" genus="Triplectides" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="318" phylum="Arthropoda" rank="species" species="buengkanensis">
<emphasis id="75C95C61471D4535FD8D07011C9AEE02" box="[576,741,835,861]" italics="true" pageId="11" pageNumber="318">buengkanensis</emphasis>
</taxonomicName>
, refers to the
<typeStatus id="98063ED1471D4535FC4D07011DCFEE02" box="[896,944,835,861]" pageId="11" pageNumber="318">type</typeStatus>
locality of the species, Buengkan Province, in northeast
<collectingCountry id="3FAAC0E3471D4535FEEC07251FF7EEDE" box="[289,392,871,897]" name="Thailand" pageId="11" pageNumber="318">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>