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<document id="21D36E0BEFCCF40D70622D204C99BEB7" ID-DOI="10.11646/zootaxa.5613.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="15216331" ID-ZooBank="2FB8BE35-114F-4A97-8610-C87ECC6F0BBA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1744667564505" checkinUser="plazi" docAuthor="Ortiz, Manuel, Capetillo, Norberto &amp; Lopeztegui, Alexander" docDate="2025" docId="0389334C5612FFB2FF7788E6FDC0029F" docLanguage="en" docName="zootaxa.5613.1.7.pdf" docOrigin="Zootaxa 5613 (1)" docSource="https://doi.org/10.11646/zootaxa.5613.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Batea floreshiguerai Ortiz, Capetillo &amp; Lopeztegui, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="159" masterDocId="FFB04B345610FFB5FFE08C15FF8D0266" masterDocTitle="The family Bateidae Stebbing, 1906 (Crustacea, Peracarida) in tropical America with the description of a new species" masterLastPageNumber="164" masterPageNumber="153" pageNumber="155" updateTime="1744670529614" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="952F61E0F3A39DF6AC14C602EB859D4A">The family Bateidae Stebbing, 1906 (Crustacea, Peracarida) in tropical America with the description of a new species</mods:title>
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<mods:namePart id="AB81F69EF228FDB479DF12F36A3C0AF1">Ortiz, Manuel</mods:namePart>
<mods:affiliation id="316438DF9E9908A6DA9C232933E27981">Laboratorio de Crustáceos, Facultad de Estudios Superiores Iztacala, Universidad Nacional Autónoma de México, Avenida de Los Barrios 1, Los Reyes Iztacala, Tlalnepantla, Estado de México, C. P. 54090, México.</mods:affiliation>
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<mods:namePart id="55E277C887C520BBACF6B93258E2A0CC">Capetillo, Norberto</mods:namePart>
<mods:affiliation id="EB1C4242194B8494C40E643F9660F1D2">Federación de Sociedades Cooperativas Pesqueras Zona Centro de Baja California Sur, Melchor Ocampo, e / Durango y Chiapas, No. 1637</mods:affiliation>
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<mods:namePart id="FC87D8C7CA9A3EB48DA1BC3DF2A3A856">Lopeztegui, Alexander</mods:namePart>
<mods:affiliation id="8704A08218196414525534DBF85050F1">Centro de Investigaciones Biológicas del Noroeste. Km. 1 Carretera a San Juan de La Costa, “ EL COMITAN ”. La Paz, Baja California Sur, México. C. P. 23205.</mods:affiliation>
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<mods:title id="B95FF63385E640F213C5869F800A20E5">Zootaxa</mods:title>
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<mods:date id="18557150C7156775E77230FCCB794877">2025</mods:date>
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<mods:number id="7F4DEAE59D20448F6BB4C97A3CA834AD">2025-03-26</mods:number>
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<treatment id="0389334C5612FFB2FF7788E6FDC0029F" ID-DOI="http://doi.org/10.5281/zenodo.15216284" ID-Zenodo-Dep="15216284" LSID="urn:lsid:plazi:treatment:0389334C5612FFB2FF7788E6FDC0029F" httpUri="http://treatment.plazi.org/id/0389334C5612FFB2FF7788E6FDC0029F" lastPageId="7" lastPageNumber="159" pageId="2" pageNumber="155">
<subSubSection id="C33AD1D15612FFB7FF7788E6FE78076B" box="[151,501,1267,1294]" pageId="2" pageNumber="155" type="nomenclature">
<paragraph id="8B9F825A5612FFB7FF7788E6FE78076B" blockId="2.[151,501,1267,1330]" box="[151,501,1267,1294]" pageId="2" pageNumber="155">
<heading id="D0D735365612FFB7FF7788E6FE78076B" bold="true" box="[151,501,1267,1294]" fontSize="11" level="1" pageId="2" pageNumber="155" reason="1">
<emphasis id="B9545E485612FFB7FF7788E6FE78076B" bold="true" box="[151,501,1267,1294]" pageId="2" pageNumber="155">
<taxonomicName id="4C20F9D95612FFB7FF7788E6FE1D0768" authorityName="Ortiz &amp; Capetillo &amp; Lopeztegui" authorityYear="2025" box="[151,400,1267,1294]" class="Malacostraca" family="Bateidae" genus="Batea" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="155" phylum="Arthropoda" rank="species" species="floreshiguerai" status="sp. nov.">
<emphasis id="B9545E485612FFB7FF7788E6FE1D0768" bold="true" box="[151,400,1267,1294]" italics="true" pageId="2" pageNumber="155">Batea floreshiguerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A267E3335612FFB7FE7788E6FE78076B" box="[407,501,1267,1293]" pageId="2" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33AD1D15612FFB7FF778902FE860754" box="[151,267,1303,1330]" pageId="2" pageNumber="155" type="description">
<paragraph id="8B9F825A5612FFB7FF778902FE860754" blockId="2.[151,501,1267,1330]" box="[151,267,1303,1330]" pageId="2" pageNumber="155">
(
<figureCitation id="131B9EDF5612FFB7FF7F8902FE8E0754" box="[159,259,1303,1330]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="3.[151,250,805,829]" captionStartId-1="4.[151,250,1976,2000]" captionStartId-2="5.[151,250,1447,1471]" captionStartId-3="6.[151,250,1552,1576]" captionTargetBox-0="[418,1168,234,779]" captionTargetBox-1="[280,1308,630,1947]" captionTargetBox-2="[301,1286,179,1421]" captionTargetBox-3="[403,1184,179,1527]" captionTargetId-0="figure-49@3.[418,1168,234,779]" captionTargetId-1="figure-223@4.[280,1308,630,1949]" captionTargetId-2="figure-18@5.[301,1286,179,1421]" captionTargetId-3="figure-27@6.[403,1184,179,1527]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 2. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). Lateral view of holotype." captionText-1="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." captionText-2="FIGURE 4. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, gnathopod 1; B, gnathopod 2." captionText-3="FIGURE 5. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, pereopod 3; B, pereopod 4; C, pereopod 5." figureDoi-0="http://doi.org/10.5281/zenodo.15216335" figureDoi-1="http://doi.org/10.5281/zenodo.15216337" figureDoi-2="http://doi.org/10.5281/zenodo.15216339" figureDoi-3="http://doi.org/10.5281/zenodo.15216341" httpUri-0="https://zenodo.org/record/15216335/files/figure.png" httpUri-1="https://zenodo.org/record/15216337/files/figure.png" httpUri-2="https://zenodo.org/record/15216339/files/figure.png" httpUri-3="https://zenodo.org/record/15216341/files/figure.png" pageId="2" pageNumber="155">Figs 2–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33AD1D15612FFB7FF77894AFA0607A7" pageId="2" pageNumber="155" type="materials_examined">
<paragraph id="8B9F825A5612FFB7FF77894AFEE507F8" blockId="2.[151,1437,1375,2014]" pageId="2" pageNumber="155">
<emphasis id="B9545E485612FFB7FF77894AFEB3071F" bold="true" box="[151,318,1375,1401]" pageId="2" pageNumber="155">Type material</emphasis>
.
<materialsCitation id="3B4888075612FFB7FEAD894AFEE507F8" collectionCode="CNCR" elevation="2" location="Deep" municipality="Mangrove" pageId="2" pageNumber="155" specimenCode="CNCR 37442" specimenCount="1" typeStatus="holotype">
<typeStatus id="549B3CF85612FFB7FEAD894AFE34071F" box="[333,441,1375,1401]" pageId="2" pageNumber="155" type="holotype">Holotype</typeStatus>
; ovigerous female;
<quantity id="4CD82FBF5612FFB7FD76894AFD60071C" box="[662,749,1375,1402]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="155" unit="mm" value="3.2">3.2 mm</quantity>
(
<specimenCode id="DB862A215612FFB7FD1C894AFC11071F" box="[764,924,1375,1401]" collectionCode="CNCR" pageId="2" pageNumber="155">CNCR 37442</specimenCode>
);
<quantity id="4CD82FBF5612FFB7FC50894AFC7C071C" box="[944,1009,1375,1402]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="155" unit="m" value="1.5">
<elevation id="000D65695612FFB7FC50894AFC7C071C" box="[944,1009,1375,1402]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="155" unit="m" value="1.5">1.5 m</elevation>
</quantity>
<location id="8EFFD4815612FFB7FC188975FBBA071C" LSID="urn:lsid:plazi:treatment:0389334C5612FFB2FF7788E6FDC0029F:8EFFD4815612FFB7FC188975FBBA071C" box="[1016,1079,1376,1402]" municipality="Mangrove" name="Deep" pageId="2" pageNumber="155">Deep</location>
; soft-bottom, mud sediments of
<collectingMunicipality id="6BFB18205612FFB7FF778991FE8707F8" box="[151,266,1412,1438]" pageId="2" pageNumber="155">Mangrove</collectingMunicipality>
swamp.
</materialsCitation>
</paragraph>
<paragraph id="8B9F825A5612FFB7FF2789B2FA0607A7" blockId="2.[151,1437,1375,2014]" box="[199,1419,1447,1474]" pageId="2" pageNumber="155">
<materialsCitation id="3B4888075612FFB7FF2789B2FA0A07A7" box="[199,1415,1447,1474]" latitude="24.250652" location="Restoration Subzone of Puerto Gato" longLatPrecision="1" longitude="-110.31397" municipality="Natural Protected Area" pageId="2" pageNumber="155" specimenCount="1" typeStatus="holotype">
<emphasis id="B9545E485612FFB7FF2789B2FEEB07A7" bold="true" box="[199,358,1447,1473]" pageId="2" pageNumber="155">
<typeStatus id="549B3CF85612FFB7FF2789B2FE8F07A7" box="[199,258,1447,1473]" pageId="2" pageNumber="155">Type</typeStatus>
locality.
</emphasis>
<location id="8EFFD4815612FFB7FE8C89B2FC8D07A7" LSID="urn:lsid:plazi:treatment:0389334C5612FFB2FF7788E6FDC0029F:8EFFD4815612FFB7FE8C89B2FC8D07A7" box="[364,768,1447,1474]" latitude="24.250652" longLatPrecision="1" longitude="-110.31397" municipality="Natural Protected Area" name="Restoration Subzone of Puerto Gato" pageId="2" pageNumber="155">Restoration Subzone of Puerto Gato</location>
a
<collectingMunicipality id="6BFB18205612FFB7FCFB89B2FB9607A7" box="[795,1051,1447,1473]" pageId="2" pageNumber="155">Natural Protected Area</collectingMunicipality>
(-
<geoCoordinate id="EE14E49D5612FFB7FBD189B2FB5007A4" box="[1073,1245,1447,1474]" degrees="110.313974" direction="west" orientation="longitude" pageId="2" pageNumber="155" precision="1" value="-110.31397">110.313974 W</geoCoordinate>
;
<geoCoordinate id="EE14E49D5612FFB7FB0889B2FAF307A7" box="[1256,1406,1447,1474]" degrees="24.250652" direction="north" orientation="latitude" pageId="2" pageNumber="155" precision="1" value="24.250652">24.250652 N</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33AD1D15612FFB7FF2789DEFE1B044B" pageId="2" pageNumber="155" type="etymology">
<paragraph id="8B9F825A5612FFB7FF2789DEFE1B044B" blockId="2.[151,1437,1375,2014]" pageId="2" pageNumber="155">
<emphasis id="B9545E485612FFB7FF2789DEFEC50783" bold="true" box="[199,328,1483,1509]" pageId="2" pageNumber="155">Etymology</emphasis>
. This new species is named in honor of José de Jesús
<collectingCountry id="F337C2CA5612FFB7FC7A89DEFC6D0783" box="[922,992,1483,1509]" name="Indonesia" pageId="2" pageNumber="155">Flores</collectingCountry>
Higuera, president of the Federación de Cooperativas Pesqueras Zona Centro de Baja California Sur, friend and activist for the conservation and protection of marine biodiversity.
</paragraph>
</subSubSection>
<subSubSection id="C33AD1D15612FFB7FF278A22FF4105F0" pageId="2" pageNumber="155" type="diagnosis">
<paragraph id="8B9F825A5612FFB7FF278A22FF4105F0" blockId="2.[151,1437,1375,2014]" pageId="2" pageNumber="155">
<emphasis id="B9545E485612FFB7FF278A22FECF0437" bold="true" box="[199,322,1591,1617]" pageId="2" pageNumber="155">Diagnosis.</emphasis>
Head with rostrum and cephalic lobe not pointed; eyes kidney shape. Body devoid of posterodorsal processes. Antenna 1 peduncular article 1 longest, as long as articles 2–3 combined, without ventral process; accessory flagellum vestigial; main flagellum article 3 posterior margin set of callynophores. Antenna 2 peduncular article 4 thicker than 5. Mandibles with 6 incisors; molar triturative with long seta on lateral margin; lacinia mobilis 2- or 4-dentate. Gnathopod 1 rudimentary; basis thickening distally with 6 long and 5 short distal simple setae, anterior marginal with single seta, posterior margin with 11 long. Gnathopod 2 propodus palmar margin oblique; palmar corner defined by 2 strong robust setae; carpus 1.3 times as long as propodus. pereopods 3–5 propodus fisures posterodistal margin. Uropod 3 rami equal in length; inner ramus inner margin with 6 simple setae basally; 8 plumose long setae distally; 8 teeth epimerum 3 posterior margin; telson longer than wide, bilobed, notched half way.
</paragraph>
</subSubSection>
<subSubSection id="C33AD1D15612FFB0FF278B8AFEFB05F5" lastPageId="5" lastPageNumber="158" pageId="2" pageNumber="155" type="description">
<paragraph id="8B9F825A5612FFB6FF278B8AFA9002B3" blockId="2.[151,1437,1375,2014]" lastBlockId="3.[151,1437,151,213]" lastPageId="3" lastPageNumber="156" pageId="2" pageNumber="155">
<emphasis id="B9545E485612FFB7FF278B8AFED405DF" bold="true" box="[199,345,1951,1977]" pageId="2" pageNumber="155">Description.</emphasis>
Based on the female
<typeStatus id="549B3CF85612FFB7FDA78B8AFD2405DF" box="[583,681,1951,1977]" pageId="2" pageNumber="155" type="holotype">holotype</typeStatus>
(CNCR 37442).
<emphasis id="B9545E485612FFB7FC888B8AFC2F05DF" box="[872,930,1951,1977]" italics="true" pageId="2" pageNumber="155">Body</emphasis>
. Head anteroventrally angular, length as long as pereonites 1–2 combined; eyes kidney-shape; rostrum flexed, length 0.7 times as long as basal article of antenna 1 peduncle; cephalic lobe small, rounded; head anteroventrally angular. Pereonites 2–6 increasing their length towards the back; pereonite 7 twice as long as pereonite 1; pleonites and urosomites dorsally smooth (
<figureCitation id="131B9EDF5613FFB6FB4D8CAEFA9D02B3" box="[1197,1296,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,805,829]" captionTargetBox="[418,1168,234,779]" captionTargetId="figure-49@3.[418,1168,234,779]" captionTargetPageId="3" captionText="FIGURE 2. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). Lateral view of holotype." figureDoi="http://doi.org/10.5281/zenodo.15216335" httpUri="https://zenodo.org/record/15216335/files/figure.png" pageId="3" pageNumber="156">Figure 2</figureCitation>
).
</paragraph>
<caption id="DF5FD2D25613FFB6FF778F30FAE10158" ID-DOI="http://doi.org/10.5281/zenodo.15216335" ID-Zenodo-Dep="15216335" box="[151,1388,805,830]" httpUri="https://zenodo.org/record/15216335/files/figure.png" pageId="3" pageNumber="156" startId="3.[151,250,805,829]" targetBox="[418,1168,234,779]" targetPageId="3" targetType="figure">
<paragraph id="8B9F825A5613FFB6FF778F30FAE10158" blockId="3.[151,1388,805,830]" box="[151,1388,805,830]" pageId="3" pageNumber="156">
<emphasis id="B9545E485613FFB6FF778F30FE9E015B" bold="true" box="[151,275,805,829]" pageId="3" pageNumber="156">FIGURE 2.</emphasis>
<taxonomicName id="4C20F9D95613FFB6FEF98F33FE670158" authorityName="Ortiz &amp; Capetillo &amp; Lopeztegui" authorityYear="2025" box="[281,490,806,830]" class="Malacostraca" family="Bateidae" genus="Batea" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="156" phylum="Arthropoda" rank="species" species="floreshiguerai" status="sp. nov.">
<emphasis id="B9545E485613FFB6FEF98F33FE670158" box="[281,490,806,830]" italics="true" pageId="3" pageNumber="156">Batea floreshiguerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A267E3335613FFB6FE108F33FDCF015B" box="[496,578,806,830]" pageId="3" pageNumber="156" rank="species">
<emphasis id="B9545E485613FFB6FE108F33FDB60158" bold="true" box="[496,571,806,830]" pageId="3" pageNumber="156">sp. nov</emphasis>
.
</taxonomicNameLabel>
Holotype, ovigerous female, 3.2 mm (CNCR 37442). Lateral view of holotype.
</paragraph>
</caption>
<paragraph id="8B9F825A5613FFB6FF278F85FB150670" blockId="3.[151,1437,912,2018]" pageId="3" pageNumber="156">
<emphasis id="B9545E485613FFB6FF278F85FECC01CC" bold="true" box="[199,321,912,938]" pageId="3" pageNumber="156">Antennae.</emphasis>
<emphasis id="B9545E485613FFB6FEA88F85FE3701CC" box="[328,442,912,938]" italics="true" pageId="3" pageNumber="156">Antenna 1</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FE298F85FDB301CC" box="[457,574,912,938]" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3A</figureCitation>
) peduncle article 1 longest, as long as articles 2–3 combined, without ventral process; accessory flagellum vestigial; main flagellum articles 1–3 as length as peduncular article 2, main flagellum article 3 posterior margin set of callynophores; rest flagellum missing.
<emphasis id="B9545E485613FFB6FC578FCDFBA40194" box="[951,1065,984,1010]" italics="true" pageId="3" pageNumber="156">Antenna 2</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FBD98FCDFB210194" box="[1081,1196,984,1010]" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3B</figureCitation>
) peduncular article 3 with ventrodistal robust seta, article 4 thicker than 5, of same length rest flagellum missing.
</paragraph>
<paragraph id="8B9F825A5613FFB6FF278835FE3607A0" blockId="3.[151,1437,912,2018]" pageId="3" pageNumber="156">
<emphasis id="B9545E485613FFB6FF278835FEDB065C" bold="true" box="[199,342,1056,1082]" pageId="3" pageNumber="156">Mouthparts</emphasis>
<emphasis id="B9545E485613FFB6FE808835FE5D065C" box="[352,464,1056,1082]" italics="true" pageId="3" pageNumber="156">Maxilla 1</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FE018835FDDA065C" box="[481,599,1056,1082]" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3C</figureCitation>
) inner plate tapering distally with 6 curved setae marginally, and 1 tiny seta distally; outer plate with 14 apical slender setae; palp 2 articulate; proximal article 0.7 as length as distal article; distal article 5 long and 4 short robust setae.
<emphasis id="B9545E485613FFB6FD3D887DFCC406E4" box="[733,841,1128,1154]" italics="true" pageId="3" pageNumber="156">Maxilla 2</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FCB7887DFC4606E4" box="[855,971,1128,1154]" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3D</figureCitation>
) inner plate narrower than outer, tapering distally, 7 anteromedial long setae; outer plate distally wide long smooth setae apically.
<emphasis id="B9545E485613FFB6FB868899FB5106C0" box="[1126,1244,1164,1190]" italics="true" pageId="3" pageNumber="156">Mandibles</emphasis>
5 incisors, molar triturative; long seta in lateral margin; palp 3 articles, article 2 thickens distally, 3 setae next to the base of article 3, distal article short, stout, 8 long curved robust setae on inner margin.
<emphasis id="B9545E485613FFB6FC4088C1FBC10688" box="[928,1100,1236,1262]" italics="true" pageId="3" pageNumber="156">Right mandible</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FBBB88C1FB410688" box="[1115,1228,1236,1262]" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3E</figureCitation>
) 2 teeth on lacinia mobilis, 4 accessory setae.
<emphasis id="B9545E485613FFB6FE2488EDFDD00774" box="[452,605,1272,1298]" italics="true" pageId="3" pageNumber="156">Left mandible</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FD8B88EDFD560774" box="[619,731,1272,1298]" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3F</figureCitation>
) 4 teeth on lacinia mobilis, 5 accessory setae.
<emphasis id="B9545E485613FFB6FB3988ECFAC90774" box="[1241,1348,1272,1298]" italics="true" pageId="3" pageNumber="156">Upper lip</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FAB188EDFF370750" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3G</figureCitation>
) longer than wide, not tapering.
<emphasis id="B9545E485613FFB6FDC38908FD020750" box="[547,655,1308,1334]" italics="true" pageId="3" pageNumber="156">Lower lip</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FD7C8909FC820750" box="[668,783,1308,1334]" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3H</figureCitation>
) as long as wide, inner plate absent, mandibular lobes well developed.
<emphasis id="B9545E485613FFB6FEFA8955FE1E073C" box="[282,403,1344,1370]" italics="true" pageId="3" pageNumber="156">Maxilliped</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FE438955FD82073C" box="[419,527,1344,1370]" captionStart="FIGURE 3" captionStartId="4.[151,250,1976,2000]" captionTargetBox="[280,1308,630,1947]" captionTargetId="figure-223@4.[280,1308,630,1949]" captionTargetPageId="4" captionText="FIGURE 3. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped." figureDoi="http://doi.org/10.5281/zenodo.15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="3" pageNumber="156">Figure 3I</figureCitation>
) inner plate 3 molariform robust setae distally, 6 simple setae, 3 marginal setae subdistally, outer plate a bit shorter than palp article 2, 8 molariform teeth, 1 simple distal seta, 5 long distal setae in palp article 2, 1 subdistally, article 3 half-length as article 2, article 3, 7 distal setae, 6 subdistally setae, dactylus 0.9 as length as propodus.
</paragraph>
<paragraph id="8B9F825A5613FFB6FF2789DAFE980480" blockId="3.[151,1437,912,2018]" pageId="3" pageNumber="156">
<emphasis id="B9545E485613FFB6FF2789DAFED2078C" bold="true" box="[199,351,1487,1514]" italics="true" pageId="3" pageNumber="156">Gnathopod 1</emphasis>
(Figura 4A) rudimentary; completely hidden behind coxa 2; basis thickening distally; sets of 6 long and 5 short distal simple setae; 1 single marginal seta; 11 long on the opposite side.
<emphasis id="B9545E485613FFB6FB9B89E6FA9F0468" bold="true" box="[1147,1298,1523,1550]" italics="true" pageId="3" pageNumber="156">Gnathopod 2</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FAC089E1FA190468" box="[1312,1428,1524,1550]" captionStart="FIGURE 4" captionStartId="5.[151,250,1447,1471]" captionTargetBox="[301,1286,179,1421]" captionTargetId="figure-18@5.[301,1286,179,1421]" captionTargetPageId="5" captionText="FIGURE 4. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, gnathopod 1; B, gnathopod 2." figureDoi="http://doi.org/10.5281/zenodo.15216339" httpUri="https://zenodo.org/record/15216339/files/figure.png" pageId="3" pageNumber="156">Figure 4B</figureCitation>
) coxa as long as it is tall; as long as carpus length; basis as long as merus and carpus combined; anterior margin one long; 6 short setae; posterior margin 3 distal setae; ischium shorter than basis; merus 4 posterior margin subdistal setae; carpus distoventral lobe 8 long setae; carpus 1.3 longer than propodus; 2 anterodorsal setae; propodus oblique; 2 setae anterior margin; 6 anterior; 12 palmar margin setae; palmar corner defined by 2 strong robust setae; posterior margin 2 short setae; dactylus fixing palm; oostegite large as long as the entire gnathopod; laminar, long marginal extensions.
</paragraph>
<paragraph id="8B9F825A5613FFB1FF278AFAFE7B005B" blockId="3.[151,1437,912,2018]" lastBlockId="4.[151,1437,151,573]" lastPageId="4" lastPageNumber="157" pageId="3" pageNumber="156">
<emphasis id="B9545E485613FFB6FF278AFAFEC6056C" bold="true" box="[199,331,1775,1802]" italics="true" pageId="3" pageNumber="156">Pereopod 3</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FEBB8AE5FE5C056D" box="[347,465,1776,1803]" captionStart="FIGURE 5" captionStartId="6.[151,250,1552,1576]" captionTargetBox="[403,1184,179,1527]" captionTargetId="figure-27@6.[403,1184,179,1527]" captionTargetPageId="6" captionText="FIGURE 5. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, pereopod 3; B, pereopod 4; C, pereopod 5." figureDoi="http://doi.org/10.5281/zenodo.15216341" httpUri="https://zenodo.org/record/15216341/files/figure.png" pageId="3" pageNumber="156">Figure 5A</figureCitation>
) coxa quadrangular, thickening ventrally; basis 2 posterodistal short setae; isquium 1 posterodistal seta; merus—carpus same length than propodus; 2 long curved setae merus anterior lobe; posterior margin 2 midway short setae; 1 distally; propodus 2 short setae anterodistally; posterior margin 2 short setae midway; posterodistal fissure present; dactylus as length as propodus; oostegite as length as the entire pereopod; long extensions anterior margin; shorter in posterior margin.
<emphasis id="B9545E485613FFB6FCB68B6AFC5705FC" bold="true" box="[854,986,1919,1946]" italics="true" pageId="3" pageNumber="156">Pereopod 4</emphasis>
(
<figureCitation id="131B9EDF5613FFB6FC0B8B95FBEC05FD" box="[1003,1121,1920,1947]" captionStart="FIGURE 5" captionStartId="6.[151,250,1552,1576]" captionTargetBox="[403,1184,179,1527]" captionTargetId="figure-27@6.[403,1184,179,1527]" captionTargetPageId="6" captionText="FIGURE 5. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, pereopod 3; B, pereopod 4; C, pereopod 5." figureDoi="http://doi.org/10.5281/zenodo.15216341" httpUri="https://zenodo.org/record/15216341/files/figure.png" pageId="3" pageNumber="156">Figure 5B</figureCitation>
) coxa the largest; posterior lobe broad; posterior margin convex; basis 1 antero and postero distal setae; ischium naked; merus and carpus as length as propodus; merus anterodistal lobe small; 1 apical seta; 1 midway anterior and posterior seta; 2 long 2 short posterodistal setae; Midway carpus 4 distal 2 midway posterior setae; propodus posterodistal fissure present; dactylus curved; 0.4 as length as propodus; oostegite not surpass carpus.
<emphasis id="B9545E485614FFB1FC3C8CAFFBED02B3" bold="true" box="[988,1120,186,213]" italics="true" pageId="4" pageNumber="157">Pereopod 5</emphasis>
(
<figureCitation id="131B9EDF5614FFB1FB908CAEFB6802B3" box="[1136,1253,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1552,1576]" captionTargetBox="[403,1184,179,1527]" captionTargetId="figure-27@6.[403,1184,179,1527]" captionTargetPageId="6" captionText="FIGURE 5. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, pereopod 3; B, pereopod 4; C, pereopod 5." figureDoi="http://doi.org/10.5281/zenodo.15216341" httpUri="https://zenodo.org/record/15216341/files/figure.png" pageId="4" pageNumber="157">Figure 5C</figureCitation>
) coxa widening ventrally; basis 2 posterodistal, 3 midway short setae; propodus as legth as isquium—carpus combined; merus anterodistal lobe 3 setae; 1 posterodistally; carpus 2 anterodistal 2 posterodistal, 2 midway short setae; propodus 2 midway, 1 anterodistal seta; posterodistal fissure present; 5 anterodistal, 2 midway setae; curved dactylus as length as merus; oostegite not surpass carpus.
<emphasis id="B9545E485614FFB1FDD08D5EFD3B0302" bold="true" box="[560,694,330,357]" pageId="4" pageNumber="157">Pereopod 6</emphasis>
(
<figureCitation id="131B9EDF5614FFB1FD248D5EFCB70303" box="[708,826,331,357]" captionStart="FIGURE 6" captionStartId="7.[151,250,1667,1691]" captionTargetBox="[396,1192,270,1638]" captionTargetId="figure-61@7.[396,1192,270,1641]" captionTargetPageId="7" captionText="FIGURE 6. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, pereopod 6; B, pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.15216343" httpUri="https://zenodo.org/record/15216343/files/figure.png" pageId="4" pageNumber="157">Figure 6A</figureCitation>
) pereopod 6 coxa curved in right angle; 4 submarginal posterior setae; basis subrectangular; wide; mid line 4 short, 2 long robust setae; anterior lobe 7 plumose robust setae; ischium triangular; 1 short subdistal anterior seta; merus as length as basis; anterior margin 3 sets 2–3 setae; discret posterodistal lobe 2 long setae; 2 single setae; 2 sets 2–3 setae; rest missing.
<emphasis id="B9545E485614FFB1FBAD8DA2FB5903B7" bold="true" box="[1101,1236,439,465]" pageId="4" pageNumber="157">Pereopod 7</emphasis>
(
<figureCitation id="131B9EDF5614FFB1FB038DA2FADA03B7" box="[1251,1367,439,465]" captionStart="FIGURE 6" captionStartId="7.[151,250,1667,1691]" captionTargetBox="[396,1192,270,1638]" captionTargetId="figure-61@7.[396,1192,270,1641]" captionTargetPageId="7" captionText="FIGURE 6. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, pereopod 6; B, pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.15216343" httpUri="https://zenodo.org/record/15216343/files/figure.png" pageId="4" pageNumber="157">Figure 6B</figureCitation>
) coxa ovoidal; anterior margin 3 setae; 2 apically; posterior margin 7 marginal setae; discreete posterior lobe; ischium triangular; anterodistal seta; merus posterior lobe 2 robust and 2 slender setae; 2 sets of 2 setae posterior margin; 2 anterodistal setae: rest missing.
</paragraph>
<caption id="DF5FD2D25614FFB1FF778BADFB1D0592" ID-DOI="http://doi.org/10.5281/zenodo.15216337" ID-Zenodo-Dep="15216337" httpUri="https://zenodo.org/record/15216337/files/figure.png" pageId="4" pageNumber="157" startId="4.[151,250,1976,2000]" targetBox="[280,1308,630,1947]" targetPageId="4" targetType="figure">
<paragraph id="8B9F825A5614FFB1FF778BADFB1D0592" blockId="4.[151,1436,1976,2036]" pageId="4" pageNumber="157">
<emphasis id="B9545E485614FFB1FF778BADFE9E05B6" bold="true" box="[151,275,1976,2000]" pageId="4" pageNumber="157">FIGURE 3.</emphasis>
<taxonomicName id="4C20F9D95614FFB1FEF98BACFE6705B6" authorityName="Ortiz &amp; Capetillo &amp; Lopeztegui" authorityYear="2025" box="[281,490,1976,2000]" class="Malacostraca" family="Bateidae" genus="Batea" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="floreshiguerai" status="sp. nov.">
<emphasis id="B9545E485614FFB1FEF98BACFE6705B6" box="[281,490,1976,2000]" italics="true" pageId="4" pageNumber="157">Batea floreshiguerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A267E3335614FFB1FE108BADFDCF05A9" box="[496,578,1976,2000]" pageId="4" pageNumber="157" rank="species">
<emphasis id="B9545E485614FFB1FE108BADFDB605B6" bold="true" box="[496,571,1976,2000]" pageId="4" pageNumber="157">sp. nov</emphasis>
.
</taxonomicNameLabel>
Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, antenna 1; B, antenna 2; C, maxilla 1; D, maxilla 2; E, right mandible; F, left mandible; G, upper lip; H, lower lip; I, maxilliped.
</paragraph>
</caption>
<caption id="DF5FD2D25615FFB0FF7789B2FF270785" ID-DOI="http://doi.org/10.5281/zenodo.15216339" ID-Zenodo-Dep="15216339" httpUri="https://zenodo.org/record/15216339/files/figure.png" pageId="5" pageNumber="158" startId="5.[151,250,1447,1471]" targetBox="[301,1286,179,1421]" targetPageId="5" targetType="figure">
<paragraph id="8B9F825A5615FFB0FF7789B2FF270785" blockId="5.[151,1436,1447,1507]" pageId="5" pageNumber="158">
<emphasis id="B9545E485615FFB0FF7789B2FE9E07D9" bold="true" box="[151,275,1447,1471]" pageId="5" pageNumber="158">FIGURE 4.</emphasis>
<taxonomicName id="4C20F9D95615FFB0FEF989BDFE6407D9" authorityName="Ortiz &amp; Capetillo &amp; Lopeztegui" authorityYear="2025" box="[281,489,1447,1471]" class="Malacostraca" family="Bateidae" genus="Batea" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="floreshiguerai" status="sp. nov.">
<emphasis id="B9545E485615FFB0FEF989BDFE6407D9" box="[281,489,1447,1471]" italics="true" pageId="5" pageNumber="158">Batea floreshiguerai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A267E3335615FFB0FE0E89B2FDCD07D9" box="[494,576,1447,1471]" pageId="5" pageNumber="158" rank="species">
<emphasis id="B9545E485615FFB0FE0E89B2FDB707D9" bold="true" box="[494,570,1447,1471]" pageId="5" pageNumber="158">sp. nov</emphasis>
.
</taxonomicNameLabel>
Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, gnathopod 1; B, gnathopod 2.
</paragraph>
</caption>
<paragraph id="8B9F825A5615FFB0FF278A20FDF40541" blockId="5.[151,1437,1589,1939]" pageId="5" pageNumber="158">
<emphasis id="B9545E485615FFB0FF278A20FEBA0429" bold="true" box="[199,311,1589,1616]" italics="true" pageId="5" pageNumber="158">Uropod 1</emphasis>
(
<figureCitation id="131B9EDF5615FFB0FEA98A20FE4F0429" box="[329,450,1589,1615]" captionStart="FIGURE 7" captionStartId="8.[151,250,1505,1529]" captionTargetBox="[346,1241,179,1473]" captionTargetId="figure-27@8.[346,1241,179,1479]" captionTargetPageId="8" captionText="FIGURE 7. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, uropod 1; B, uropod 2; C, uropod 3; D, telson; E, epimeral plates 1–3." figureDoi="http://doi.org/10.5281/zenodo.15216345" httpUri="https://zenodo.org/record/15216345/files/figure.png" pageId="5" pageNumber="158">Figure 7A</figureCitation>
) peduncle 4 setae outer margin; 1 robust 1 simple subdistal setae outer ramus; bifid distally; as length as peduncle; 1.3 as length as inner ramus; 5 setae outer margin; 5 setae inner margin; 2 joint robust distal setae.
<emphasis id="B9545E485615FFB0FEC38A68FE1D04F1" bold="true" box="[291,400,1661,1688]" italics="true" pageId="5" pageNumber="158">Uropod 2</emphasis>
(
<figureCitation id="131B9EDF5615FFB0FE7F8A68FD9E04F1" box="[415,531,1661,1687]" captionStart="FIGURE 7" captionStartId="8.[151,250,1505,1529]" captionTargetBox="[346,1241,179,1473]" captionTargetId="figure-27@8.[346,1241,179,1479]" captionTargetPageId="8" captionText="FIGURE 7. Batea floreshiguerai sp. nov. Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, uropod 1; B, uropod 2; C, uropod 3; D, telson; E, epimeral plates 1–3." figureDoi="http://doi.org/10.5281/zenodo.15216345" httpUri="https://zenodo.org/record/15216345/files/figure.png" pageId="5" pageNumber="158">Figure 7B</figureCitation>
) the shortest; peduncle 1 subdistal robust seta; 2 times longer than outer ramus; outer ramus 1 distal 1 subdistal setae; inner ramus inner margin 1 robust seta midway; distally curved, pointed.
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3
</emphasis>
(Figura 7C) peduncle 0.3 as long as rami; rami equals in length; inner ramus inner margin 6 simple setae basally; 8 plumose long setae distally; inner margin 1 small subdistal seta; outer margin 1 short seta outer margin; 1 midway long seta; 2 subdistal; not pointed.
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(
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). 3 longer than wide; bilobed; notched half way; each lobe 1 lateral, 1 subterminal short setae.
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posteroventral corner curved.
<emphasis id="B9545E485615FFB0FD278B43FCB10509" box="[711,828,1878,1903]" italics="true" pageId="5" pageNumber="158">Epimerum</emphasis>
2 almost acute; 8 strong teeth.
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posterior margin (
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).
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<emphasis id="B9545E485616FFB3FF778A05FE9E044F" bold="true" box="[151,275,1552,1577]" pageId="6" pageNumber="159">FIGURE 5.</emphasis>
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Holotype, ovigerous female, 3.2 mm (CNCR 37442). A, pereopod 3; B, pereopod 4; C, pereopod 5.
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. The 15 known species of
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can be separated in 2 groups. The former for those with body dorsally smooth, which includes
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<emphasis id="B9545E485616FFB3FD388AD6FCA204BB" bold="true" box="[728,815,1731,1757]" pageId="6" pageNumber="159">sp. nov</emphasis>
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, and the second for those with 1 or more pereonal or pleonal posterodorsal processes. Therefore, the present discussion focuses on distinguishing species with completely smooth back. In this sense the new species differs from the species in the group by having the combination of the following characters: kidney-shaped eyes; peduncular article 1 of antenna 1 without ventral process; vestigial accessory flagellum of antenna 1; main flagellum article 3 of antenna 1 posterior margin with a set of callynophore; left and right mandibles with lacinia mobilis bearing 4 and 2 teeth, respectively; mandibular lobes of the lower lip well developed; propodus of gnathopod 2 palmar margin oblique, palmar angle defined by two robust setae; pereopods 3–5 posterodistal margin of propodi with fissures (described for the first time in the genus); pereopod 6 coxa curved at right angles, with four posterior submarginal setae; subrectangular basis of pereopod 6; epimeral plate 3 posterior margin with eight strong teeth. For more details you can consult the dichotomous key that is offered to distinguish all known
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species.
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Genus
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Family
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Bateidae
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. Head rostrate; eyes large, occupying at least 1/3 of head surface; rostrum and cephalic lobe present; rostrum distally pointed or blunt. Body smooth or dentate dorsally. Antenna 1 accessory flagellum vestigial or absent. Mandible with molar and three articulated palp. Maxilla 1 inner plate conical with one small distal and seven inner marginal setae; palp two-articulate. Lower lip without inner plate. Gnathopod 1 degraded, only present coxa and feeble linear basis, both hidden by coxa 2. Gnathopod 2 subchelate; coxa 4 the largest; coxa 4 strong posterior lobe. Uropod 2 shorter than uropods 1 and 3 which are of similar length. Telson flat; cleft half way.
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Composition. Genus
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<emphasis id="B9545E485612FFB7FE548FA5FE7801AF" box="[436,501,944,969]" italics="true" pageId="2" pageNumber="155">Batea</emphasis>
<bibRefCitation id="EFB1FFAB5612FFB7FE1D8FBAFD0301AC" author="Muller F." box="[509,654,943,970]" pageId="2" pageNumber="155" pagination="276 - 277" refId="ref6425" refString="Muller F. (1865) Description of a new genus of amphipod Crustacea. Annals and Magazine of Natural History, Series 3, 15, 276-277. https://doi.org/10.1080/00222936508681804" type="journal article" year="1865">Müller, 1865</bibRefCitation>
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<emphasis id="B949EAE5FF900C1DFF4257D8FE22F89C" bold="true" box="[151,397,1770,1797]" italics="true" pageId="6" pageNumber="132">Tepperia vagabunda</emphasis>
Tsuji &amp; Yoshitake
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(
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,
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&amp;
<figureCitation id="13062A72FF900C1DFE48563DFE4BF8B0" box="[413,484,1806,1833]" captionStart-0="FIGURES 37–50" captionStart-1="FIGURES 51–54" captionStartId-0="9.[151,264,1328,1352]" captionStartId-1="10.[151,264,1485,1509]" captionTargetBox-0="[167,1419,188,1296]" captionTargetBox-1="[164,1423,561,1461]" captionTargetId-0="figure-28@9.[164,1423,181,1304]" captionTargetId-1="figure-171@10.[164,1423,561,1461]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 37–50. Terminalia of Tepperia vagabunda; 37–45, Male; 46–50, female. 37 &amp; 46, Tergites VII; 38 &amp; 39, penis; 40, apex of penis, at different angle; 41 &amp; 42, copulatory sclerite; 43, tegmen; 44, sternite VIII; 45, spiculum gastrale; 47, right stylus and gonocoxite; 48, spermatheca; 49, tergite VIII; 50, spiculum ventrale. 38–40 &amp; 43–45, Holotype; 37, 41, 42 &amp; 46–50, paratypes. 37, 38, 41, 43, 46–47 &amp; 49, Dorsal aspect; 39 &amp; 42, lateral aspect; 44, 45 &amp; 50, ventral aspect. Scale A: 0.5 mm for 37, 46, 49 &amp; 50; 0.2 mm for 41, 42, 47 &amp; 48. Scale B: 0.2 mm for 38–40 &amp; 43–45." captionText-1="FIGURES 51–54. Biology of Tepperia vagabunda on Iriomotejima Island, the Ryukyu Islands, Japan. 51, A Heritiera littoralis fruit washed ashore (indicated by an arrow); 52, inside of a H. littoralis fruit infested by the weevil; 53, an adult and a larva in a H. littoralis seed; 54, an adult on a H. littoralis twig." figureDoi-0="http://doi.org/10.5281/zenodo.15216288" figureDoi-1="http://doi.org/10.5281/zenodo.15216290" httpUri-0="https://zenodo.org/record/15216288/files/figure.png" httpUri-1="https://zenodo.org/record/15216290/files/figure.png" pageId="6" pageNumber="132">37–54</figureCitation>
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<emphasis id="B949EAE5FF900C1DFF425649FEBDF80D" bold="true" box="[151,274,1914,1940]" pageId="6" pageNumber="132">Diagnosis.</emphasis>
This species is similar to
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<emphasis id="B949EAE5FF900C1DFDEC5648FDE6F80D" box="[569,585,1915,1940]" italics="true" pageId="6" pageNumber="132">T</emphasis>
.
<emphasis id="B949EAE5FF900C1DFD8D5648FD74F80C" box="[600,731,1915,1941]" italics="true" pageId="6" pageNumber="132">palawanica</emphasis>
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in general appearance but can be clearly distinguished from it by having paired brown longitudinal scaly bands on the pronotum and a thicker rostrum whose dorsal contour is more strongly arcuate.
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<emphasis id="B949EAE5FF910C1CFF4254B7FEF9FA04" bold="true" box="[151,342,1412,1437]" pageId="7" pageNumber="133">FIGURES 19–24.</emphasis>
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<emphasis id="B949EAE5FF910C1CFE8B54B6FE18FA05" box="[350,439,1413,1436]" italics="true" pageId="7" pageNumber="133">Tepperia</emphasis>
spp.
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19–21,
<taxonomicName id="4C3D4D74FF910C1CFDEA54B6FC1CFA04" authority="(Heller, 1913)" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" baseAuthorityName="Heller" baseAuthorityYear="1913" box="[575,947,1413,1437]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="palawanica">
<emphasis id="B949EAE5FF910C1CFDEA54B6FCB8FA04" box="[575,791,1413,1437]" italics="true" pageId="7" pageNumber="133">Tepperia palawanica</emphasis>
(
<bibRefCitation id="EFAC4B06FF910C1CFCF354B6FC04FA04" author="Heller, K. M." box="[806,939,1413,1437]" pageId="7" pageNumber="133" pagination="135 - 163" refId="ref7669" refString="Heller, K. M. (1913) Neue Kafer von den Philippinen. Philippine Journal of Science, section D. General biology, ethnology, and anthropology, 8, 135-163." type="journal article" year="1913">Heller, 1913</bibRefCitation>
)
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; 22–24,
<taxonomicName id="4C3D4D74FF910C1CFBD854B6FB4FFA04" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" box="[1037,1248,1413,1437]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="vagabunda">
<emphasis id="B949EAE5FF910C1CFBD854B6FB4FFA04" box="[1037,1248,1413,1437]" italics="true" pageId="7" pageNumber="133">Tepperia vagabunda</emphasis>
</taxonomicName>
; 19, 20, 22 &amp; 23, heads, dorsal aspect; 21 &amp; 24, left mesotibiae, anterior aspect. 19 &amp; 21,
<taxonomicName id="4C3D4D74FF910C1CFCBD549AFB5EFA58" authority="Heller, 1913" authorityName="Heller" authorityYear="1913" box="[872,1265,1449,1473]" class="Insecta" family="Curculionidae" genus="Cyamobolus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="133" phylum="Arthropoda" rank="species" species="palawanicus">
<emphasis id="B949EAE5FF910C1CFCBD549AFBC2FA58" box="[872,1133,1449,1473]" italics="true" pageId="7" pageNumber="133">Cyamobolus palawanicus</emphasis>
<bibRefCitation id="EFAC4B06FF910C1CFBA7549AFB5EFA58" author="Heller, K. M." box="[1138,1265,1449,1473]" pageId="7" pageNumber="133" pagination="135 - 163" refId="ref7669" refString="Heller, K. M. (1913) Neue Kafer von den Philippinen. Philippine Journal of Science, section D. General biology, ethnology, and anthropology, 8, 135-163." type="journal article" year="1913">Heller, 1913</bibRefCitation>
</taxonomicName>
, lectotype, male; 20, ditto, paralectotype, female. 22 &amp; 24, Holotype, male; 23, paratype, female. Scale: 2.0 mm for 19, 20, 22 &amp; 23, 1.5 mm for 21 &amp; 24.
</paragraph>
</caption>
<subSubSection id="C327657CFF910C11FF125768FA9DFEFC" lastPageId="10" lastPageNumber="136" pageId="7" pageNumber="133" type="description">
<paragraph id="8B8236F7FF910C1CFF125768FC23F900" blockId="7.[151,1437,1627,2013]" pageId="7" pageNumber="133">
<emphasis id="B949EAE5FF910C1CFF125768FE33F9EC" bold="true" box="[199,412,1627,1653]" pageId="7" pageNumber="133">Description. Male</emphasis>
. Measurements (n = 10; in mm): TL 7.1–8.2 (
<typeStatus id="54868855FF910C1CFC425768FC54F9EC" box="[919,1019,1627,1653]" pageId="7" pageNumber="133" type="holotype">holotype</typeStatus>
7.9, mean 7.7), LP 2.1–2.6 (2.5, 2.4); WP 2.4–2.8 (2.7, 2.7); LE 5.0–5.6 (5.4, 5.3); WE 3.3–3.9 (3.8, 3.7).
</paragraph>
<paragraph id="8B8236F7FF910C13FF125790FB38FE10" blockId="7.[151,1437,1627,2013]" lastBlockId="8.[151,1437,151,573]" lastPageId="8" lastPageNumber="134" pageId="7" pageNumber="133">Most parts of body covered with dull-white to yellowish brown recumbent scales. Vertex with paired medial patches of dark brown scales behind eyes. Rostrum covered with brownish subovate to oblong scales, which become smaller and slenderer toward apex. Pronotum mainly covered with close light-colored subovate scales, sparsely mingled with whitish and brownish suberect scales, which become slenderer toward apex; medial part bearing paired narrow triangular patches of brownish scales along midline; apical margin with paired tufts of brownish suberect scales on middle. Lateral parts of prothorax densely covered with light-colored scales, sparsely mingled with dark brown suberect scales; prosternal canal mainly glabrous except apical part densely covered with light-colored slender scales. Scutellar shield subglabrous, except apico-lateral part fringed with minute scales. Elytra mostly covered with light-colored to brownish subovate scales, with the following two paired markings of dark suberect scales: obscure stripes on subbasal parts of 4th intervals and spots on behind middle of 4th intervals; dark spots sometimes extending to 3rd intervals; strial punctures each bearing whitish subrecumbent spathulate scale. Mesoventral receptacle densely covered with minute scales on apical margin. Metaventrite and metanepisterna also covered with light-colored subovate scales, except glabrous patch on post-median part of metaventrite. Legs with femora and tibiae irregularly mottled with light-colored to brown scales. Profemur with triangular glabrous patch on apico-ventral part; meso- and metafemur glabrous ventrally along ventral sulci. Abdominal ventrites closely covered with pale brown recumbent scales and sparsely mingled with subrecumbent scales.</paragraph>
<paragraph id="8B8236F7FF9E0C13FF1250A0FC5CFD80" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="134">
<emphasis id="B949EAE5FF9E0C13FF1250A0FEA8FE34" bold="true" box="[199,263,403,429]" pageId="8" pageNumber="134">Head</emphasis>
(
<figureCitation id="13062A72FF9E0C13FEC250A0FEC3FE34" box="[279,364,403,429]" captionStart="FIGURES 19–24" captionStartId="7.[151,264,1412,1436]" captionTargetBox="[166,1421,181,1388]" captionTargetId="figure-28@7.[166,1421,181,1389]" captionTargetPageId="7" captionText="FIGURES 19–24. Tepperia spp. 19–21, Tepperia palawanica (Heller, 1913); 22–24, Tepperia vagabunda; 19, 20, 22 &amp; 23, heads, dorsal aspect; 21 &amp; 24, left mesotibiae, anterior aspect. 19 &amp; 21, Cyamobolus palawanicus Heller, 1913, lectotype, male; 20, ditto, paralectotype, female. 22 &amp; 24, Holotype, male; 23, paratype, female. Scale: 2.0 mm for 19, 20, 22 &amp; 23, 1.5 mm for 21 &amp; 24." figureDoi="http://doi.org/10.5281/zenodo.15216282" httpUri="https://zenodo.org/record/15216282/files/figure.png" pageId="8" pageNumber="134">Fig. 22</figureCitation>
). Rostrum LR/WRM 2.6–2.9, LR/LP 0.7–0.8, thicker, in profile more strongly curved ventrad; sides more weakly attenuate from base to before middle; dorsal contour more strongly arcuate. Antennae inserted just behind middle; approximate length ratio of 1st to 7th funicular segments as follows: 2.8–3.5: 2: 1: 1: 1: 1.2: 1.2–1.4; club 2.3–2.6 times as long as wide, 0.7–0.8 times as long as funicle.
</paragraph>
<paragraph id="8B8236F7FF9E0C13FF125310FB0FFDA4" blockId="8.[151,1437,151,573]" box="[199,1184,547,573]" pageId="8" pageNumber="134">
<emphasis id="B949EAE5FF9E0C13FF125310FE8FFDA4" bold="true" box="[199,288,547,573]" pageId="8" pageNumber="134">Thorax</emphasis>
. Prothorax LP/WP 0.8–0.9; sides slightly attenuate from base to behind middle.
</paragraph>
<caption id="DF42667FFF9E0C13FF425653FABDF87D" ID-DOI="http://doi.org/10.5281/zenodo.15216286" ID-Zenodo-Dep="15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="8" pageNumber="134" startId="8.[151,264,1888,1912]" targetBox="[160,1427,607,1850]" targetPageId="8" targetType="figure">
<paragraph id="8B8236F7FF9E0C13FF425653FABDF87D" blockId="8.[151,1437,1888,2020]" pageId="8" pageNumber="134">
<emphasis id="B949EAE5FF9E0C13FF425653FEF8F8E1" bold="true" box="[151,343,1888,1912]" pageId="8" pageNumber="134">FIGURES 25–36.</emphasis>
Terminalia of
<taxonomicName id="4C3D4D74FF9E0C13FE205652FCC3F8E1" authority="(Heller, 1913)" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" baseAuthorityName="Heller" baseAuthorityYear="1913" box="[501,876,1888,1912]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="134" phylum="Arthropoda" rank="species" species="palawanica">
<emphasis id="B949EAE5FF9E0C13FE205652FD61F8E1" box="[501,718,1888,1912]" italics="true" pageId="8" pageNumber="134">Tepperia palawanica</emphasis>
(
<bibRefCitation id="EFAC4B06FF9E0C13FD0B5653FCCBF8E1" author="Heller, K. M." box="[734,868,1888,1912]" pageId="8" pageNumber="134" pagination="135 - 163" refId="ref7669" refString="Heller, K. M. (1913) Neue Kafer von den Philippinen. Philippine Journal of Science, section D. General biology, ethnology, and anthropology, 8, 135-163." type="journal article" year="1913">Heller, 1913</bibRefCitation>
)
</taxonomicName>
. 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31.
</paragraph>
</caption>
<caption id="DF42667FFF9F0C12FF425403FC4FFA41" ID-DOI="http://doi.org/10.5281/zenodo.15216288" ID-Zenodo-Dep="15216288" httpUri="https://zenodo.org/record/15216288/files/figure.png" pageId="9" pageNumber="135" startId="9.[151,264,1328,1352]" targetBox="[167,1419,188,1296]" targetPageId="9" targetType="figure">
<paragraph id="8B8236F7FF9F0C12FF425403FC4FFA41" blockId="9.[151,1437,1328,1496]" pageId="9" pageNumber="135">
<emphasis id="B949EAE5FF9F0C12FF425403FEFBFAD1" bold="true" box="[151,340,1328,1352]" pageId="9" pageNumber="135">FIGURES 37–50.</emphasis>
Terminalia of
<taxonomicName id="4C3D4D74FF9F0C12FE3F5402FD15FAD1" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" box="[490,698,1328,1352]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="135" phylum="Arthropoda" rank="species" species="vagabunda">
<emphasis id="B949EAE5FF9F0C12FE3F5402FD15FAD1" box="[490,698,1328,1352]" italics="true" pageId="9" pageNumber="135">Tepperia vagabunda</emphasis>
</taxonomicName>
; 37–45, Male; 46–50, female. 37 &amp; 46, Tergites VII; 38 &amp; 39, penis; 40, apex of penis, at different angle; 41 &amp; 42, copulatory sclerite; 43, tegmen; 44, sternite VIII; 45, spiculum gastrale; 47, right stylus and gonocoxite; 48, spermatheca; 49, tergite VIII; 50, spiculum ventrale. 38–40 &amp; 43–45, Holotype; 37, 41, 42 &amp; 46–50, paratypes. 37, 38, 41, 43, 46–47 &amp; 49, Dorsal aspect; 39 &amp; 42, lateral aspect; 44, 45 &amp; 50, ventral aspect. Scale A: 0.5 mm for 37, 46, 49 &amp; 50; 0.2 mm for 41, 42, 47 &amp; 48. Scale B: 0.2 mm for 38–40 &amp; 43–45.
</paragraph>
</caption>
<paragraph id="8B8236F7FF9F0C12FF125713FA82F91B" blockId="9.[151,1436,1567,2026]" pageId="9" pageNumber="135">Elytra LE/WE 1.4–1.5, WE/WP 1.4–1.5, with subbasal part weakly separately prominent on sutural intervals and 3rd and 4th intervals. Sutural intervals with minute irregular granules; 9th weakly granulate-costate from about middle to subapical part; 10th weakly granulate-costate. 10th striae more deeply sulcate on entire length.</paragraph>
<paragraph id="8B8236F7FF9F0C12FF1257BFFC30F88B" blockId="9.[151,1436,1567,2026]" pageId="9" pageNumber="135">
Meso- and metafemora with ventral sulci arising from base. Tibiae weakly tuberculate basally on outer margins, then subparallel-sided apically; mesotibiae (
<figureCitation id="13062A72FF9F0C12FD505783FD79F953" box="[645,726,1712,1738]" captionStart="FIGURES 19–24" captionStartId="7.[151,264,1412,1436]" captionTargetBox="[166,1421,181,1388]" captionTargetId="figure-28@7.[166,1421,181,1389]" captionTargetPageId="7" captionText="FIGURES 19–24. Tepperia spp. 19–21, Tepperia palawanica (Heller, 1913); 22–24, Tepperia vagabunda; 19, 20, 22 &amp; 23, heads, dorsal aspect; 21 &amp; 24, left mesotibiae, anterior aspect. 19 &amp; 21, Cyamobolus palawanicus Heller, 1913, lectotype, male; 20, ditto, paralectotype, female. 22 &amp; 24, Holotype, male; 23, paratype, female. Scale: 2.0 mm for 19, 20, 22 &amp; 23, 1.5 mm for 21 &amp; 24." figureDoi="http://doi.org/10.5281/zenodo.15216282" httpUri="https://zenodo.org/record/15216282/files/figure.png" pageId="9" pageNumber="135">Fig. 24</figureCitation>
) slightly curved inward. Tarsi with 1st tarsomere 1.5–2.3 times as wide as long, 1.7–1.9 times as long as and as wide as 2nd, 1.5–1.8 times as long as 3rd; 2nd subquadrate, as long as wide; 3rd 1.6–1.8 times as wide as long; 5th almost as long as 3rd.
</paragraph>
<paragraph id="8B8236F7FF9F0C12FF12562FFAA8F8AF" blockId="9.[151,1436,1567,2026]" box="[199,1287,1820,1846]" pageId="9" pageNumber="135">Metaventrite approximately 1.3 times as long along midline as maximum length of mesoventrite.</paragraph>
<paragraph id="8B8236F7FF9F0C12FF12560CFD17F8E7" blockId="9.[151,1436,1567,2026]" pageId="9" pageNumber="135">
<emphasis id="B949EAE5FF9F0C12FF12560CFE93F8C0" bold="true" box="[199,316,1855,1881]" pageId="9" pageNumber="135">Abdomen</emphasis>
. Ventrites irregularly punctate; ventrite I slightly longer, 1.6–1.9 times as long as II along midline, hardly to barely emarginate along apical margin.
</paragraph>
<paragraph id="8B8236F7FF9F0C12FF1256B4FEC7F873" blockId="9.[151,1436,1567,2026]" pageId="9" pageNumber="135">
<emphasis id="B949EAE5FF9F0C12FF1256B4FEE5F838" bold="true" box="[199,330,1927,1953]" pageId="9" pageNumber="135">Terminalia</emphasis>
(
<figureCitation id="13062A72FF9F0C12FE8D56BBFE4FF83B" box="[344,480,1927,1954]" captionStart="FIGURES 37–50" captionStartId="9.[151,264,1328,1352]" captionTargetBox="[167,1419,188,1296]" captionTargetId="figure-28@9.[164,1423,181,1304]" captionTargetPageId="9" captionText="FIGURES 37–50. Terminalia of Tepperia vagabunda; 37–45, Male; 46–50, female. 37 &amp; 46, Tergites VII; 38 &amp; 39, penis; 40, apex of penis, at different angle; 41 &amp; 42, copulatory sclerite; 43, tegmen; 44, sternite VIII; 45, spiculum gastrale; 47, right stylus and gonocoxite; 48, spermatheca; 49, tergite VIII; 50, spiculum ventrale. 38–40 &amp; 43–45, Holotype; 37, 41, 42 &amp; 46–50, paratypes. 37, 38, 41, 43, 46–47 &amp; 49, Dorsal aspect; 39 &amp; 42, lateral aspect; 44, 45 &amp; 50, ventral aspect. Scale A: 0.5 mm for 37, 46, 49 &amp; 50; 0.2 mm for 41, 42, 47 &amp; 48. Scale B: 0.2 mm for 38–40 &amp; 43–45." figureDoi="http://doi.org/10.5281/zenodo.15216288" httpUri="https://zenodo.org/record/15216288/files/figure.png" pageId="9" pageNumber="135">Figs. 37–45</figureCitation>
). Penis with body approximately 1.4 times as long as wide, subparallel-sided or slightly expanded in basal 1/3, then gently convergent apically, in profile slightly thinner; apex more weakly arcuate, faintly notched at middle.
</paragraph>
<paragraph id="8B8236F7FF9C0C11FF1251A4FB0DFE84" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="136">
<emphasis id="B949EAE5FF9C0C11FF1251A4FEB1FF28" bold="true" box="[199,286,151,177]" pageId="10" pageNumber="136">Female</emphasis>
. Measurements (n = 10; in mm): TL 5.4–8.3 (mean 7.6), LP 1.6–2.5 (2.3); WP 1.8–2.8 (2.6); LE 3.8–5.8 (5.2); WE 2.6–3.9 (3.7). Rostrum (
<figureCitation id="13062A72FF9C0C11FDC25188FDC8FF4C" box="[535,615,187,213]" captionStart="FIGURES 19–24" captionStartId="7.[151,264,1412,1436]" captionTargetBox="[166,1421,181,1388]" captionTargetId="figure-28@7.[166,1421,181,1389]" captionTargetPageId="7" captionText="FIGURES 19–24. Tepperia spp. 19–21, Tepperia palawanica (Heller, 1913); 22–24, Tepperia vagabunda; 19, 20, 22 &amp; 23, heads, dorsal aspect; 21 &amp; 24, left mesotibiae, anterior aspect. 19 &amp; 21, Cyamobolus palawanicus Heller, 1913, lectotype, male; 20, ditto, paralectotype, female. 22 &amp; 24, Holotype, male; 23, paratype, female. Scale: 2.0 mm for 19, 20, 22 &amp; 23, 1.5 mm for 21 &amp; 24." figureDoi="http://doi.org/10.5281/zenodo.15216282" httpUri="https://zenodo.org/record/15216282/files/figure.png" pageId="10" pageNumber="136">Fig. 23</figureCitation>
) LR/WRM 2.5–3.0, LR/LP 0.7–0.8; dorsal surface with punctures sparser on apical half; sides more strongly attenuate from base to middle. Prothorax LP/WP 0.9. Elytra LE/WE 1.4–1.5, WE/WP 1.4–1.5. Ventrite I slightly inflated medially excepting abdominal process flattened.
</paragraph>
<paragraph id="8B8236F7FF9C0C11FF125015FA9DFEFC" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="136">
<emphasis id="B949EAE5FF9C0C11FF125015FEE5FED9" bold="true" box="[199,330,294,320]" pageId="10" pageNumber="136">Terminalia</emphasis>
(
<figureCitation id="13062A72FF9C0C11FE895014FE48FED8" box="[348,487,294,321]" captionStart="FIGURES 37–50" captionStartId="9.[151,264,1328,1352]" captionTargetBox="[167,1419,188,1296]" captionTargetId="figure-28@9.[164,1423,181,1304]" captionTargetPageId="9" captionText="FIGURES 37–50. Terminalia of Tepperia vagabunda; 37–45, Male; 46–50, female. 37 &amp; 46, Tergites VII; 38 &amp; 39, penis; 40, apex of penis, at different angle; 41 &amp; 42, copulatory sclerite; 43, tegmen; 44, sternite VIII; 45, spiculum gastrale; 47, right stylus and gonocoxite; 48, spermatheca; 49, tergite VIII; 50, spiculum ventrale. 38–40 &amp; 43–45, Holotype; 37, 41, 42 &amp; 46–50, paratypes. 37, 38, 41, 43, 46–47 &amp; 49, Dorsal aspect; 39 &amp; 42, lateral aspect; 44, 45 &amp; 50, ventral aspect. Scale A: 0.5 mm for 37, 46, 49 &amp; 50; 0.2 mm for 41, 42, 47 &amp; 48. Scale B: 0.2 mm for 38–40 &amp; 43–45." figureDoi="http://doi.org/10.5281/zenodo.15216288" httpUri="https://zenodo.org/record/15216288/files/figure.png" pageId="10" pageNumber="136">Figs. 46–50</figureCitation>
). Tergite VIII with apical margin simple, not notched at middle. Spiculum ventrale with apical plate subparallel-sided basally. Spermatheca thicker, with cornu more strongly curved apicad.
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF9C0C11FF12505CFDE9FE34" pageId="10" pageNumber="136" type="etymology">
<paragraph id="8B8236F7FF9C0C11FF12505CFDE9FE34" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="136">
<emphasis id="B949EAE5FF9C0C11FF12505CFEE1FE10" bold="true" box="[199,334,367,393]" pageId="10" pageNumber="136">Etymology.</emphasis>
The specific name,
<taxonomicName id="4C3D4D74FF9C0C11FDE5505CFD17FE10" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" box="[560,696,367,393]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="136" phylum="Arthropoda" rank="species" species="vagabunda">
“
<emphasis id="B949EAE5FF9C0C11FDE9505CFD17FE10" box="[572,696,367,393]" italics="true" pageId="10" pageNumber="136">vagabunda</emphasis>
</taxonomicName>
”, is a Latin adjective meaning vagabond, referring to its dispersal habit with fruits of
<taxonomicName id="4C3D4D74FF9C0C11FEBA50A0FD90FE34" box="[367,575,403,429]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FEBA50A0FD90FE34" box="[367,575,403,429]" italics="true" pageId="10" pageNumber="136">Heritiera littoralis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF9C0C10FF125084FC94FEFC" lastPageId="11" lastPageNumber="137" pageId="10" pageNumber="136" type="materials_examined">
<paragraph id="8B8236F7FF9C0C11FF125084FE5EFD80" blockId="10.[151,1437,150,537]" pageId="10" pageNumber="136">
<emphasis id="B949EAE5FF9C0C11FF125084FEDBFE49" bold="true" box="[199,372,438,465]" pageId="10" pageNumber="136">Type material.</emphasis>
<materialsCitation id="3B553CAAFF9C0C11FEAF5084FE5EFD80" collectingDate="2005-04-28" collectionCode="KUM" collectorName="I. Matoba" location="Nakaragawa River" pageId="10" pageNumber="136" specimenCount="2" typeStatus="holotype">
<typeStatus id="54868855FF9C0C11FEAF5084FE4AFE48" box="[378,485,439,465]" pageId="10" pageNumber="136" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="ED2CAE32FF9C0C11FDE65084FDD4FE48" box="[563,635,439,465]" pageId="10" pageNumber="136">KUM</collectionCode>
): “(Ryukyus) /
<location id="8EE2602CFF9C0C11FCF45084FC5EFE48" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9C0C11FCF45084FC5EFE48" box="[801,1009,439,465]" name="Nakaragawa River" pageId="10" pageNumber="136">Nakaragawa River</location>
/ Iriomote I. /
<collectorName id="26C85321FF9C0C11FB5A5084FAAFFE48" box="[1167,1280,439,465]" pageId="10" pageNumber="136">I. Matoba</collectorName>
”, “
<date id="FF831037FF9C0C11FACB5085FA33FE48" box="[1310,1436,438,465]" pageId="10" pageNumber="136" value="2005-04-28">
<collectingDate id="EFC7E9DFFF9C0C11FACB5085FA33FE48" box="[1310,1436,438,465]" pageId="10" pageNumber="136" value="2005-04-28">28.IV.2005</collectingDate>
</date>
<emphasis id="B949EAE5FF9C0C11FF4250E8FDCBFE6C" bold="true" box="[151,612,475,501]" pageId="10" pageNumber="136">/ サキシマスオウノキ / DZƗよりī化</emphasis>
”, “
<typeStatus id="54868855FF9C0C11FD5250E8FCB1FE6C" box="[647,798,475,501]" pageId="10" pageNumber="136" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C3D4D74FF9C0C11FCE750E8FC3AFE6D" box="[818,917,475,500]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAE5FF9C0C11FCE750E8FC3AFE6D" box="[818,917,475,500]" italics="true" pageId="10" pageNumber="136">Tepperia</emphasis>
</taxonomicName>
/
<taxonomicName id="4C3D4D74FF9C0C11FC7D50E8FB8BFE6C" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" box="[936,1060,475,501]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="136" phylum="Arthropoda" rank="species" species="vagabunda">
<emphasis id="B949EAE5FF9C0C11FC7D50E8FB8BFE6C" box="[936,1060,475,501]" italics="true" pageId="10" pageNumber="136">vagabunda</emphasis>
</taxonomicName>
/ Tsuji et Yoshitake, 2025” (typed on a red card; newly attached).
</materialsCitation>
</paragraph>
<caption id="DF42667FFF9C0C11FF4254FEFD10F9B7" ID-DOI="http://doi.org/10.5281/zenodo.15216290" ID-Zenodo-Dep="15216290" httpUri="https://zenodo.org/record/15216290/files/figure.png" pageId="10" pageNumber="136" startId="10.[151,264,1485,1509]" targetBox="[164,1423,561,1461]" targetPageId="10" targetType="figure">
<paragraph id="8B8236F7FF9C0C11FF4254FEFD10F9B7" blockId="10.[151,1437,1485,1582]" pageId="10" pageNumber="136">
<emphasis id="B949EAE5FF9C0C11FF4254FEFEFCFA7C" bold="true" box="[151,339,1485,1509]" pageId="10" pageNumber="136">FIGURES 51–54.</emphasis>
Biology of
<taxonomicName id="4C3D4D74FF9C0C11FE1E54FDFD34FA7F" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" box="[459,667,1486,1510]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="136" phylum="Arthropoda" rank="species" species="vagabunda">
<emphasis id="B949EAE5FF9C0C11FE1E54FDFD34FA7F" box="[459,667,1486,1510]" italics="true" pageId="10" pageNumber="136">Tepperia vagabunda</emphasis>
</taxonomicName>
on Iriomotejima Island, the Ryukyu Islands, Japan. 51, A
<taxonomicName id="4C3D4D74FF9C0C11FB3454FDFA33FA7F" box="[1249,1436,1486,1510]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FB3454FDFA33FA7F" box="[1249,1436,1486,1510]" italics="true" pageId="10" pageNumber="136">Heritiera littoralis</emphasis>
</taxonomicName>
fruit washed ashore (indicated by an arrow); 52, inside of a
<taxonomicName id="4C3D4D74FF9C0C11FD2254C1FCC1F993" box="[759,878,1522,1546]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FD2254C1FCA6F990" box="[759,777,1522,1545]" italics="true" pageId="10" pageNumber="136">H</emphasis>
.
<emphasis id="B949EAE5FF9C0C11FCC254C1FCC1F993" box="[791,878,1522,1546]" italics="true" pageId="10" pageNumber="136">littoralis</emphasis>
</taxonomicName>
fruit infested by the weevil; 53, an adult and a larva in a
<taxonomicName id="4C3D4D74FF9C0C11FF7C5725FEB0F9B7" box="[169,287,1558,1582]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FF7C5725FEB0F9B7" box="[169,287,1558,1582]" italics="true" pageId="10" pageNumber="136">H. littoralis</emphasis>
</taxonomicName>
seed; 54, an adult on a
<taxonomicName id="4C3D4D74FF9C0C11FDDB5725FD2BF9B7" box="[526,644,1558,1582]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FDDB5725FD8FF9B4" box="[526,544,1558,1581]" italics="true" pageId="10" pageNumber="136">H</emphasis>
.
<emphasis id="B949EAE5FF9C0C11FDF85725FD2BF9B7" box="[557,644,1558,1582]" italics="true" pageId="10" pageNumber="136">littoralis</emphasis>
</taxonomicName>
twig.
</paragraph>
</caption>
<paragraph id="8B8236F7FF9C0C10FF1257B3FC08FE84" blockId="10.[151,1437,1664,2015]" lastBlockId="11.[151,1437,150,537]" lastPageId="11" lastPageNumber="137" pageId="10" pageNumber="136">
<materialsCitation id="3B553CAAFF9C0C11FF1257B3FC90F903" box="[199,831,1664,1690]" collectorName="Ishigakijima Is." country="Japan" location="Ryukyu Islands" pageId="10" pageNumber="136" specimenCount="1" typeStatus="paratype">
<typeStatus id="54868855FF9C0C11FF1257B3FE97F903" box="[199,312,1664,1690]" pageId="10" pageNumber="136" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F32A7667FF9C0C11FE9657B3FE28F903" box="[323,391,1664,1690]" name="Japan" pageId="10" pageNumber="136">Japan</collectingCountry>
: the
<location id="8EE2602CFF9C0C11FE6B57B3FDDDF903" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9C0C11FE6B57B3FDDDF903" box="[446,626,1664,1690]" country="Japan" name="Ryukyu Islands" pageId="10" pageNumber="136">Ryukyu Islands</location>
: [
<collectorName id="26C85321FF9C0C11FD5657B3FC98F903" box="[643,823,1664,1690]" pageId="10" pageNumber="136">Ishigakijima Is.</collectorName>
]
</materialsCitation>
<materialsCitation id="3B553CAAFF9C0C11FC9D57B3FF40F927" collectingDate="1964-03-16" collectionCode="KUM" collectorName="Y. Miyatake" country="Japan" location="Omoto-dake" pageId="10" pageNumber="136" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FC9D57B3FC02F903" box="[840,941,1664,1690]" count="1" pageId="10" pageNumber="136" type="female">1 female</specimenCount>
,
<location id="8EE2602CFF9C0C11FC6C57B3FBE7F903" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9C0C11FC6C57B3FBE7F903" box="[953,1096,1664,1690]" country="Japan" name="Omoto-dake" pageId="10" pageNumber="136">Omoto-dake</location>
,
<date id="FF831037FF9C0C11FB8057B3FB7AF903" box="[1109,1237,1664,1690]" pageId="10" pageNumber="136" value="1964-03-16">
<collectingDate id="EFC7E9DFFF9C0C11FB8057B3FB7AF903" box="[1109,1237,1664,1690]" pageId="10" pageNumber="136" value="1964-03-16">16.III.1964</collectingDate>
</date>
,
<collectorName id="26C85321FF9C0C11FB3457B3FAC4F903" box="[1249,1387,1664,1690]" pageId="10" pageNumber="136">Y. Miyatake</collectorName>
leg. (
<collectionCode id="ED2CAE32FF9C0C11FF4A5797FF49F927" box="[159,230,1700,1726]" pageId="10" pageNumber="136">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9C0C11FF2E5797FCECF927" box="[251,835,1700,1727]" collectingDate="1975-06-08" collectionCode="KUM" collectorName="T. Takahashi" country="Japan" location="Omoto" pageId="10" pageNumber="136" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FF2E5797FE9AF927" box="[251,309,1700,1726]" count="1" pageId="10" pageNumber="136" type="generic">1 ex.</specimenCount>
,
<location id="8EE2602CFF9C0C11FE965797FE3BF927" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9C0C11FE965797FE3BF927" box="[323,404,1700,1726]" country="Japan" name="Omoto" pageId="10" pageNumber="136">Omoto</location>
,
<date id="FF831037FF9C0C11FE755797FDBAF926" box="[416,533,1700,1727]" pageId="10" pageNumber="136" value="1975-06-08">
<collectingDate id="EFC7E9DFFF9C0C11FE755797FDBAF926" box="[416,533,1700,1727]" pageId="10" pageNumber="136" value="1975-06-08">8.VI.1975</collectingDate>
</date>
,
<collectorName id="26C85321FF9C0C11FDF45797FD1EF927" box="[545,689,1700,1726]" pageId="10" pageNumber="136">T. Takahashi</collectorName>
leg. (
<collectionCode id="ED2CAE32FF9C0C11FD265797FC95F927" box="[755,826,1700,1726]" pageId="10" pageNumber="136">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9C0C11FC9A5797FBB7F927" box="[847,1048,1700,1726]" country="Japan" location="Iriomotejima Is." pageId="10" pageNumber="136" specimenCount="1" typeStatus="paratype">
[
<location id="8EE2602CFF9C0C11FC805797FBBFF927" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9C0C11FC805797FBBFF927" box="[853,1040,1700,1726]" country="Japan" name="Iriomotejima Is." pageId="10" pageNumber="136">Iriomotejima Is.</location>
]
</materialsCitation>
<materialsCitation id="3B553CAAFF9C0C11FBF45797FADCF97B" collectingDate="2023-03-22" collectingDateMax="2023-05-06" collectingDateMin="2023-03-22" collectionCode="PCKF" collectorName="S. Katoh &amp; H. Hoshito &amp; K. Nishida" country="Japan" location="Haiminaka" pageId="10" pageNumber="136" specimenCount="1" stateProvince="Fukui" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FBF45797FBF4F927" box="[1057,1115,1700,1726]" count="1" pageId="10" pageNumber="136" type="generic">1 ex.</specimenCount>
,
<location id="8EE2602CFF9C0C11FBBC5797FB48F927" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9C0C11FBBC5797FB48F927" box="[1129,1255,1700,1726]" country="Japan" name="Haiminaka" pageId="10" pageNumber="136" stateProvince="Fukui">Haiminaka</location>
,
<date id="FF831037FF9C0C11FB265797FADBF927" box="[1267,1396,1700,1726]" pageId="10" pageNumber="136" value="2023-03-22">
<collectingDate id="EFC7E9DFFF9C0C11FB265797FADBF927" box="[1267,1396,1700,1726]" pageId="10" pageNumber="136" value="2023-03-22">22.III.2023</collectingDate>
</date>
, K.
<collectingRegion id="49F9F815FF9C0C11FF4257FBFF74F97B" box="[151,219,1736,1762]" country="Japan" name="Fukui" pageId="10" pageNumber="136">Fukui</collectingRegion>
,
<collectorName id="26C85321FF9C0C11FF3F57FBFEFEF97B" box="[234,337,1736,1762]" pageId="10" pageNumber="136">S. Katoh</collectorName>
,
<collectorName id="26C85321FF9C0C11FE8A57FBFE71F97B" box="[351,478,1736,1762]" pageId="10" pageNumber="136">H. Hoshito</collectorName>
&amp;
<collectorName id="26C85321FF9C0C11FDD257FBFD2CF97B" box="[519,643,1736,1762]" pageId="10" pageNumber="136">K. Nishida</collectorName>
leg., as a fruit of
<taxonomicName id="4C3D4D74FF9C0C11FC8E57FAFB82F97B" box="[859,1069,1736,1762]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FC8E57FAFB82F97B" box="[859,1069,1736,1762]" italics="true" pageId="10" pageNumber="136">Heritiera littoralis</emphasis>
</taxonomicName>
, emerged
<date id="FF831037FF9C0C11FB7D57FBFAA3F97B" box="[1192,1292,1736,1762]" pageId="10" pageNumber="136" value="2023-05-06">
<collectingDate id="EFC7E9DFFF9C0C11FB7D57FBFAA3F97B" box="[1192,1292,1736,1762]" pageId="10" pageNumber="136" value="2023-05-06">6.V.2023</collectingDate>
</date>
(
<collectionCode id="ED2CAE32FF9C0C11FACB57FBFAC4F97B" box="[1310,1387,1736,1762]" pageId="10" pageNumber="136">PCKF</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9C0C11FA5557FBFD6FF89F" collectingDate="2004-12-25" collectionCode="PCHY" collectorName="J. Kantoh" country="Japan" location="Komi" pageId="10" pageNumber="136" specimenCount="10" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FA5557FBFF69F89F" count="10" pageId="10" pageNumber="136" type="generic">10 exs.</specimenCount>
,
<location id="8EE2602CFF9C0C11FF0657DFFEB9F89F" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9C0C11FF0657DFFEB9F89F" box="[211,278,1772,1798]" country="Japan" name="Komi" pageId="10" pageNumber="136">Komi</location>
,
<date id="FF831037FF9C0C11FEF757DFFE02F89E" box="[290,429,1772,1799]" pageId="10" pageNumber="136" value="2004-12-25">
<collectingDate id="EFC7E9DFFF9C0C11FEF757DFFE02F89E" box="[290,429,1772,1799]" pageId="10" pageNumber="136" value="2004-12-25">25.XII.2004</collectingDate>
</date>
,
<collectorName id="26C85321FF9C0C11FE6F57DFFD89F89F" box="[442,550,1772,1798]" pageId="10" pageNumber="136">J. Kantoh</collectorName>
leg. (
<collectionCode id="ED2CAE32FF9C0C11FDB257DFFD17F89F" box="[615,696,1772,1798]" pageId="10" pageNumber="136">PCHY</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D3BFD7EFF9C0C11FD1857DFFC9DF89F" box="[717,818,1772,1798]" count="1" pageId="10" pageNumber="136" type="female">1 female</specimenCount>
,
<date id="FF831037FF9C0C11FCEB57DFFC1BF89E" box="[830,948,1772,1799]" pageId="10" pageNumber="136" value="2005-05-20">20.V.2005</date>
, I. Matoba leg., emerged from a fruit of
<taxonomicName id="4C3D4D74FF9C0C11FA5457DEFEF7F8B3" authority="(KUM)" baseAuthorityName="KUM" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FA5457DEFF58F8B3" italics="true" pageId="10" pageNumber="136">H. littoralis</emphasis>
(KUM)
</taxonomicName>
;
<specimenCount id="9D3BFD7EFF9C0C11FEB05623FE12F8B3" box="[357,445,1808,1834]" count="2" pageId="10" pageNumber="136" type="male">2 males</specimenCount>
&amp;
<specimenCount id="9D3BFD7EFF9C0C11FE315623FDE5F8B3" box="[484,586,1808,1834]" count="1" pageId="10" pageNumber="136" type="female">1 female</specimenCount>
,
<date id="FF831037FF9C0C11FD825623FD63F8B2" box="[599,716,1808,1835]" pageId="10" pageNumber="136" value="2005-05-21">21.V.2005</date>
, I. Matoba leg., emerged from fruits of
<taxonomicName id="4C3D4D74FF9C0C11FB405622FA2EF8B3" authority=", (KUM)" baseAuthorityName="KUM" box="[1173,1409,1808,1834]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FB405622FAB6F8B3" box="[1173,1305,1808,1834]" italics="true" pageId="10" pageNumber="136">H. littoralis</emphasis>
, (KUM)
</taxonomicName>
;
<specimenCount id="9D3BFD7EFF9C0C11FA5B5623FF7FF8D7" count="1" pageId="10" pageNumber="136" type="male">1 male</specimenCount>
,
<date id="FF831037FF9C0C11FF0B5607FEFDF8D6" box="[222,338,1844,1871]" pageId="10" pageNumber="136" value="2005-06-07">7.VI.2005</date>
, I. Matoba leg., emerged from a fruit of
<taxonomicName id="4C3D4D74FF9C0C11FCF95606FBB6F8D7" authority=", (KUM)" baseAuthorityName="KUM" box="[812,1049,1844,1870]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FCF95606FC1EF8D7" box="[812,945,1844,1870]" italics="true" pageId="10" pageNumber="136">H. littoralis</emphasis>
, (KUM)
</taxonomicName>
;
<materialsCitation id="3B553CAAFF9C0C11FBF25607FCC6F8EB" collectingDate="2020-03-29" collectingDateMax="2020-05-01" collectingDateMin="2020-03-29" collectionCode="PCHY" collectorName="H. Yoshitake" country="Japan" pageId="10" pageNumber="136" specimenCount="7" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FBF25607FBC2F8D7" box="[1063,1133,1844,1870]" count="7" pageId="10" pageNumber="136" type="generic">7 exs.</specimenCount>
,
<date id="FF831037FF9C0C11FBAE5607FB53F8D7" box="[1147,1276,1844,1870]" pageId="10" pageNumber="136" value="2020-03-29">
<collectingDate id="EFC7E9DFFF9C0C11FBAE5607FB53F8D7" box="[1147,1276,1844,1870]" pageId="10" pageNumber="136" value="2020-03-29">29.III.2020</collectingDate>
</date>
,
<collectorName id="26C85321FF9C0C11FADF5607FA33F8D7" box="[1290,1436,1844,1870]" pageId="10" pageNumber="136">H. Yoshitake</collectorName>
leg., as fruits of
<taxonomicName id="4C3D4D74FF9C0C11FE87566AFE7AF8EB" box="[338,469,1880,1906]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FE87566AFEC9F8EB" box="[338,358,1881,1906]" italics="true" pageId="10" pageNumber="136">H</emphasis>
.
<emphasis id="B949EAE5FF9C0C11FEA0566BFE7AF8EB" box="[373,469,1880,1906]" italics="true" pageId="10" pageNumber="136">littoralis</emphasis>
</taxonomicName>
, emerged
<date id="FF831037FF9C0C11FD98566BFCAFF8EA" box="[589,768,1880,1907]" pageId="10" pageNumber="136" value="2020-04-15" valueMax="2020-05-01" valueMin="2020-04-15">
<collectingDate id="EFC7E9DFFF9C0C11FD98566BFCAFF8EA" box="[589,768,1880,1907]" pageId="10" pageNumber="136" value="2020-04-15" valueMax="2020-05-01" valueMin="2020-04-15">15.IV–1.V.2020</collectingDate>
</date>
(
<collectionCode id="ED2CAE32FF9C0C11FCC5566BFCCEF8EB" box="[784,865,1880,1906]" pageId="10" pageNumber="136">PCHY</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9C0C11FCA0566BFB3DF80F" collectingDate="2015-03-05" collectingDateMax="2015-04-27" collectingDateMin="2015-03-05" collectionCode="KUM" collectorName="M. Wakabayashi" country="Japan" county="Tudwumari-hama" location="Beach" municipality="Tsukigahama" pageId="10" pageNumber="136" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FCA0566BFC6BF8EB" box="[885,964,1880,1906]" count="1" pageId="10" pageNumber="136" type="male">1 male</specimenCount>
,
<collectingCounty id="62E34E7BFF9C0C11FC05566BFB37F8EB" box="[976,1176,1880,1906]" pageId="10" pageNumber="136">Tudwumari-hama</collectingCounty>
(
<collectingMunicipality id="6BE6AC8DFF9C0C11FB72566BFAEFF8EB" box="[1191,1344,1880,1906]" pageId="10" pageNumber="136">Tsukigahama</collectingMunicipality>
)
<location id="8EE2602CFF9C0C11FA9A566BFA37F8EB" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9C0C11FA9A566BFA37F8EB" box="[1359,1432,1880,1906]" country="Japan" county="Tudwumari-hama" municipality="Tsukigahama" name="Beach" pageId="10" pageNumber="136">Beach</location>
,
<date id="FF831037FF9C0C11FF42564FFEA5F80E" box="[151,266,1916,1943]" pageId="10" pageNumber="136" value="2015-03-05">
<collectingDate id="EFC7E9DFFF9C0C11FF42564FFEA5F80E" box="[151,266,1916,1943]" pageId="10" pageNumber="136" value="2015-03-05">5.III.2015</collectingDate>
</date>
,
<collectorName id="26C85321FF9C0C11FECC564FFE77F80F" box="[281,472,1916,1942]" pageId="10" pageNumber="136">M. Wakabayashi</collectorName>
leg., as a fruit of
<taxonomicName id="4C3D4D74FF9C0C11FD67564EFC97F80F" box="[690,824,1916,1942]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FD67564EFC97F80F" box="[690,824,1916,1942]" italics="true" pageId="10" pageNumber="136">H. littoralis</emphasis>
</taxonomicName>
, emerged
<date id="FF831037FF9C0C11FC61564FFB9FF80E" box="[948,1072,1916,1943]" pageId="10" pageNumber="136" value="2015-04-27">
<collectingDate id="EFC7E9DFFF9C0C11FC61564FFB9FF80E" box="[948,1072,1916,1943]" pageId="10" pageNumber="136" value="2015-04-27">27.IV.2015</collectingDate>
</date>
(
<collectionCode id="ED2CAE32FF9C0C11FB96564FFB25F80F" box="[1091,1162,1916,1942]" pageId="10" pageNumber="136">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B553CAAFF9C0C11FB74564FFB1BF823" collectingDate="2022-03-26" collectingDateMax="2022-05-05" collectingDateMin="2022-03-26" collectionCode="KUM" collectorName="M. Maruyama &amp; T. Hashizume &amp; S. Imada" country="Japan" pageId="10" pageNumber="136" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FB74564FFB72F80F" box="[1185,1245,1916,1942]" count="1" pageId="10" pageNumber="136" type="generic">1 ex.</specimenCount>
,
<date id="FF831037FF9C0C11FB38564FFAC1F80F" box="[1261,1390,1916,1942]" pageId="10" pageNumber="136" value="2022-03-26">
<collectingDate id="EFC7E9DFFF9C0C11FB38564FFAC1F80F" box="[1261,1390,1916,1942]" pageId="10" pageNumber="136" value="2022-03-26">26.III.2022</collectingDate>
</date>
,
<collectorName id="26C85321FF9C0C11FAA9564FFEBCF823" pageId="10" pageNumber="136">M. Maruyama</collectorName>
,
<collectorName id="26C85321FF9C0C11FEC85693FE18F823" box="[285,439,1952,1978]" pageId="10" pageNumber="136">T. Hashizume</collectorName>
&amp;
<collectorName id="26C85321FF9C0C11FE0C5693FD93F823" box="[473,572,1952,1978]" pageId="10" pageNumber="136">S. Imada</collectorName>
leg., as a fruit of
<taxonomicName id="4C3D4D74FF9C0C11FD295692FCD1F823" box="[764,894,1952,1978]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="136" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9C0C11FD295692FCBFF823" box="[764,784,1953,1978]" italics="true" pageId="10" pageNumber="136">H</emphasis>
.
<emphasis id="B949EAE5FF9C0C11FCCB5693FCD1F823" box="[798,894,1952,1978]" italics="true" pageId="10" pageNumber="136">littoralis</emphasis>
</taxonomicName>
, emerged
<date id="FF831037FF9C0C11FC245693FBF9F822" box="[1009,1110,1952,1979]" pageId="10" pageNumber="136" value="2022-05-05">
<collectingDate id="EFC7E9DFFF9C0C11FC245693FBF9F822" box="[1009,1110,1952,1979]" pageId="10" pageNumber="136" value="2022-05-05">5.V.2022</collectingDate>
</date>
(
<collectionCode id="ED2CAE32FF9C0C11FBB15693FB04F823" box="[1124,1195,1952,1978]" pageId="10" pageNumber="136">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9C0C11FB685693FDB3F847" collectingDate="2005-04-28" collectionCode="KUM" collectorName="I. Matoba" country="Japan" location="Nakaragawa River" pageId="10" pageNumber="136" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FB685693FAA7F823" box="[1213,1288,1952,1978]" count="1" pageId="10" pageNumber="136" type="male">1 male</specimenCount>
&amp;
<specimenCount id="9D3BFD7EFF9C0C11FAFF5693FA37F823" box="[1322,1432,1952,1978]" count="3" pageId="10" pageNumber="136" type="female">3 females</specimenCount>
, same data as the holotype (
<collectionCode id="ED2CAE32FF9C0C11FE1956F7FDBCF847" box="[460,531,1988,2014]" pageId="10" pageNumber="136">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B553CAAFF9C0C10FDFD56F7FF40FF28" collectingDate="2005-04-25" collectionCode="KUM" collectorName="I. Matoba" country="Japan" lastPageId="11" lastPageNumber="137" location="Nakaragawa River" pageId="10" pageNumber="136" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9C0C11FDFD56F7FD2FF847" box="[552,640,1988,2014]" count="2" pageId="10" pageNumber="136" type="male">2 males</specimenCount>
&amp;
<specimenCount id="9D3BFD7EFF9C0C11FD7356F7FCA3F847" box="[678,780,1988,2014]" count="1" pageId="10" pageNumber="136" type="female">1 female</specimenCount>
, same locality as the holotype,
<date id="FF831037FF9C0C11FBA456F7FB5FF846" box="[1137,1264,1988,2015]" pageId="10" pageNumber="136" value="2005-04-25">
<collectingDate id="EFC7E9DFFF9C0C11FBA456F7FB5FF846" box="[1137,1264,1988,2015]" pageId="10" pageNumber="136" value="2005-04-25">25.IV.2005</collectingDate>
</date>
,
<collectorName id="26C85321FF9C0C11FB2856F7FAC4F847" box="[1277,1387,1988,2014]" pageId="10" pageNumber="136">I. Matoba</collectorName>
leg. (
<collectionCode id="ED2CAE32FF9D0C10FF4A51A4FF49FF28" box="[159,230,151,177]" pageId="11" pageNumber="137">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B553CAAFF9D0C10FF2C51A4FC7FFF28" box="[249,976,150,177]" collectingDate="2005-04-26" collectionCode="KUM, PCIM" collectorName="I. Matoba" country="Japan" pageId="11" pageNumber="137" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9D0C10FF2C51A4FEE1FF28" box="[249,334,151,177]" count="2" pageId="11" pageNumber="137" type="male">2 males</specimenCount>
&amp;
<specimenCount id="9D3BFD7EFF9D0C10FEA551A4FE71FF28" box="[368,478,151,177]" count="2" pageId="11" pageNumber="137" type="female">2 females</specimenCount>
,
<date id="FF831037FF9D0C10FE3D51A5FDC9FF28" box="[488,614,150,177]" pageId="11" pageNumber="137" value="2005-04-26">
<collectingDate id="EFC7E9DFFF9D0C10FE3D51A5FDC9FF28" box="[488,614,150,177]" pageId="11" pageNumber="137" value="2005-04-26">26.IV.2005</collectingDate>
</date>
,
<collectorName id="26C85321FF9D0C10FDA451A4FD72FF28" box="[625,733,151,177]" pageId="11" pageNumber="137">I. Matoba</collectorName>
leg. (
<collectionCode id="ED2CAE32FF9D0C10FCCF51A4FCF2FF28" box="[794,861,151,177]" pageId="11" pageNumber="137">KUM</collectionCode>
&amp;
<collectionCode id="ED2CAE32FF9D0C10FCAA51A4FC66FF28" box="[895,969,151,177]" pageId="11" pageNumber="137">PCIM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B553CAAFF9D0C10FC0F51A4FEB1FF4C" collectingDate="2005-05-01" collectionCode="KUM" collectorName="I. Matoba" country="Japan" pageId="11" pageNumber="137" specimenCount="8" specimenCount-female="7" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9D0C10FC0F51A4FB8BFF28" box="[986,1060,151,177]" count="1" pageId="11" pageNumber="137" type="male">1 male</specimenCount>
&amp;
<specimenCount id="9D3BFD7EFF9D0C10FB9251A4FB1AFF28" box="[1095,1205,151,177]" count="7" pageId="11" pageNumber="137" type="female">7 females</specimenCount>
,
<date id="FF831037FF9D0C10FB6A51A5FA89FF28" box="[1215,1318,150,177]" pageId="11" pageNumber="137" value="2005-05-01">
<collectingDate id="EFC7E9DFFF9D0C10FB6A51A5FA89FF28" box="[1215,1318,150,177]" pageId="11" pageNumber="137" value="2005-05-01">1.V.2005</collectingDate>
</date>
,
<collectorName id="26C85321FF9D0C10FAE551A4FA32FF28" box="[1328,1437,151,177]" pageId="11" pageNumber="137">I. Matoba</collectorName>
leg. (
<collectionCode id="ED2CAE32FF9D0C10FF1A5188FEB9FF4C" box="[207,278,187,213]" pageId="11" pageNumber="137">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9D0C10FEFD5188FBF7FF4C" box="[296,1112,187,213]" collectingDate="1977-05-30" collectionCode="KUM" collectorName="J. Okuma" country="Japan" location="Sonai" pageId="11" pageNumber="137" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9D0C10FEFD5188FEDAFF4C" box="[296,373,187,213]" count="1" pageId="11" pageNumber="137" type="male">1 male</specimenCount>
, “Sonae” [sic!] (
<location id="8EE2602CFF9D0C10FDF85188FDDDFF4C" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9D0C10FDF85188FDDDFF4C" box="[557,626,187,213]" country="Japan" name="Sonai" pageId="11" pageNumber="137">Sonai</location>
),
<date id="FF831037FF9D0C10FD505188FD54FF4C" box="[645,763,187,213]" pageId="11" pageNumber="137" value="1977-05-30">
<collectingDate id="EFC7E9DFFF9D0C10FD505188FD54FF4C" box="[645,763,187,213]" pageId="11" pageNumber="137" value="1977-05-30">30.V.1977</collectingDate>
</date>
,
<collectorName id="26C85321FF9D0C10FCD05188FCC0FF4C" box="[773,879,187,213]" pageId="11" pageNumber="137">J. Okuma</collectorName>
leg., at light (
<collectionCode id="ED2CAE32FF9D0C10FBDC5188FBFFFF4C" box="[1033,1104,187,213]" pageId="11" pageNumber="137">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9D0C10FBB75188FE26FF60" collectingDate="1988-10-29" collectionCode="KUM" collectorName="M. Iwata" country="Japan" location="Amitori" pageId="11" pageNumber="137" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9D0C10FBB75188FB1FFF4C" box="[1122,1200,187,213]" count="1" pageId="11" pageNumber="137" type="male">1 male</specimenCount>
,
<location id="8EE2602CFF9D0C10FB6C5188FABAFF4C" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9D0C10FB6C5188FABAFF4C" box="[1209,1301,187,213]" country="Japan" name="Amitori" pageId="11" pageNumber="137">Amitori</location>
,
<date id="FF831037FF9D0C10FACA5188FA38FF4C" box="[1311,1431,187,213]" pageId="11" pageNumber="137" value="1988-10-29">
<collectingDate id="EFC7E9DFFF9D0C10FACA5188FA38FF4C" box="[1311,1431,187,213]" pageId="11" pageNumber="137" value="1988-10-29">29.X.1988</collectingDate>
</date>
,
<collectorName id="26C85321FF9D0C10FF4251ECFF54FF60" box="[151,251,223,249]" pageId="11" pageNumber="137">M. Iwata</collectorName>
leg. (
<collectionCode id="ED2CAE32FF9D0C10FEEC51ECFE2FFF60" box="[313,384,223,249]" pageId="11" pageNumber="137">KUM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9D0C10FE4751ECFF57FE84" collectingDate="2011-10-16" collectingDateMax="2011-12-10" collectingDateMin="2011-10-16" collectionCode="PCHY" collectorName="S. Tetsu" country="Japan" pageId="11" pageNumber="137" specimenCount="2" typeStatus="paratype">
<specimenCount id="9D3BFD7EFF9D0C10FE4751ECFE7AFF60" box="[402,469,223,249]" count="2" pageId="11" pageNumber="137" type="generic">2 exs.</specimenCount>
, east coast,
<date id="FF831037FF9D0C10FD8C51ECFD55FF60" box="[601,762,223,249]" pageId="11" pageNumber="137" value="2011-10-16" valueMax="2011-10-19" valueMin="2011-10-16">
<collectingDate id="EFC7E9DFFF9D0C10FD8C51ECFD55FF60" box="[601,762,223,249]" pageId="11" pageNumber="137" value="2011-10-16" valueMax="2011-10-19" valueMin="2011-10-16">16–19.X.2011</collectingDate>
</date>
,
<collectorName id="26C85321FF9D0C10FCD051ECFCF1FF60" box="[773,862,223,249]" pageId="11" pageNumber="137">S. Tetsu</collectorName>
leg., as a fruit of
<taxonomicName id="4C3D4D74FF9D0C10FBCA51ECFB0EFF60" box="[1055,1185,223,249]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="137" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9D0C10FBCA51ECFB0EFF60" box="[1055,1185,223,249]" italics="true" pageId="11" pageNumber="137">H. littoralis</emphasis>
</taxonomicName>
, emerged
<date id="FF831037FF9D0C10FAC151ECFA33FF60" box="[1300,1436,223,249]" pageId="11" pageNumber="137" value="2011-12-10">
<collectingDate id="EFC7E9DFFF9D0C10FAC151ECFA33FF60" box="[1300,1436,223,249]" pageId="11" pageNumber="137" value="2011-12-10">10.XII.2011</collectingDate>
</date>
(
<collectionCode id="ED2CAE32FF9D0C10FF4A5030FF5FFE84" box="[159,240,259,285]" pageId="11" pageNumber="137">PCHY</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B553CAAFF9D0C10FED75030FC0CFE84" box="[258,931,259,285]" collectingDate="1921-08-31" collectionCode="KUM" collectorName="T. Esaki" country="Taiwan" location="Kuraru" pageId="11" pageNumber="137" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="F32A7667FF9D0C10FED75030FEF7FE84" box="[258,344,259,285]" name="Taiwan" pageId="11" pageNumber="137">Taiwan</collectingCountry>
:
<specimenCount id="9D3BFD7EFF9D0C10FEB65030FE1EFE84" box="[355,433,259,285]" count="1" pageId="11" pageNumber="137" type="male">1 male</specimenCount>
,
<location id="8EE2602CFF9D0C10FE695030FDA1FE84" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9D0C10FE695030FDA1FE84" box="[444,526,259,285]" country="Taiwan" name="Kuraru" pageId="11" pageNumber="137">Kuraru</location>
,
<date id="FF831037FF9D0C10FDCF5030FD01FE84" box="[538,686,259,285]" pageId="11" pageNumber="137" value="1921-08-31">
<collectingDate id="EFC7E9DFFF9D0C10FDCF5030FD01FE84" box="[538,686,259,285]" pageId="11" pageNumber="137" value="1921-08-31">31.VIII.1921</collectingDate>
</date>
,
<collectorName id="26C85321FF9D0C10FD6C5030FCBBFE84" box="[697,788,259,285]" pageId="11" pageNumber="137">T. Esaki</collectorName>
leg. (
<collectionCode id="ED2CAE32FF9D0C10FC815030FC34FE84" box="[852,923,259,285]" pageId="11" pageNumber="137">KUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236F7FF9D0C10FF125014FC94FEFC" blockId="11.[151,1437,150,537]" pageId="11" pageNumber="137">
<materialsCitation id="3B553CAAFF9D0C10FF125014FC98FEFC" collectingDate="2005-04-28" collectionCode="KUM" collectorName="Iriomotejima Is." country="Japan" location="Ryukyu Islands" pageId="11" pageNumber="137" specimenCount="1" specimenCount-female="1">
<emphasis id="B949EAE5FF9D0C10FF125014FDB0FED9" bold="true" box="[199,543,294,321]" pageId="11" pageNumber="137">
<typeStatus id="54868855FF9D0C10FF125014FDB5FED9" box="[199,538,294,321]" isOtherTypeStatus="true" pageId="11" pageNumber="137">Non-type material examined</typeStatus>
.
</emphasis>
<collectingCountry id="F32A7667FF9D0C10FDF15014FDC7FED8" box="[548,616,295,321]" name="Japan" pageId="11" pageNumber="137">Japan</collectingCountry>
: the
<location id="8EE2602CFF9D0C10FD425014FCE7FED8" LSID="urn:lsid:plazi:treatment:039487E1FF900C10FF4257D8FCECFD80:8EE2602CFF9D0C10FD425014FCE7FED8" box="[663,840,295,321]" country="Japan" name="Ryukyu Islands" pageId="11" pageNumber="137">Ryukyu Islands</location>
: [
<collectorName id="26C85321FF9D0C10FC835014FBA2FED8" box="[854,1037,295,321]" pageId="11" pageNumber="137">Iriomotejima Is.</collectorName>
]
<specimenCount id="9D3BFD7EFF9D0C10FBCF5014FBD3FED8" box="[1050,1148,295,321]" count="1" pageId="11" pageNumber="137" type="female">1 female</specimenCount>
, same data as the holotype (
<collectionCode id="ED2CAE32FF9D0C10FF4A5078FF4AFEFC" box="[159,229,331,357]" pageId="11" pageNumber="137">KUM</collectionCode>
; an abnormal specimen with distorted elytral apices)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF9D0C10FF12505DFCECFD80" pageId="11" pageNumber="137" type="distribution">
<paragraph id="8B8236F7FF9D0C10FF12505DFB55FE10" blockId="11.[151,1437,150,537]" box="[199,1274,366,393]" pageId="11" pageNumber="137">
<emphasis id="B949EAE5FF9D0C10FF12505DFECEFE11" bold="true" box="[199,353,366,392]" pageId="11" pageNumber="137">Distribution.</emphasis>
<collectingCountry id="F32A7667FF9D0C10FEB2505CFE04FE10" box="[359,427,367,393]" name="Japan" pageId="11" pageNumber="137">Japan</collectingCountry>
: the Ryukyu Islands (Ishigakijima Island and Iriomotejima Island);
<collectingCountry id="F32A7667FF9D0C10FB75505CFB59FE10" box="[1184,1270,367,393]" name="Taiwan" pageId="11" pageNumber="137">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="8B8236F7FF9D0C10FF1250A0FCECFD80" blockId="11.[151,1437,150,537]" pageId="11" pageNumber="137">
<emphasis id="B949EAE5FF9D0C10FF1250A0FE29FE35" bold="true" box="[199,390,402,429]" pageId="11" pageNumber="137">Natural history.</emphasis>
<taxonomicName id="4C3D4D74FF9D0C10FE5850A0FDDBFE34" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" box="[397,628,403,429]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="137" phylum="Arthropoda" rank="species" species="vagabunda">
<emphasis id="B949EAE5FF9D0C10FE5850A0FDDBFE34" box="[397,628,403,429]" italics="true" pageId="11" pageNumber="137">Tepperia vagabunda</emphasis>
</taxonomicName>
is associated with
<taxonomicName id="4C3D4D74FF9D0C10FC9F50A0FBB5FE34" box="[842,1050,403,429]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="137" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9D0C10FC9F50A0FBB5FE34" box="[842,1050,403,429]" italics="true" pageId="11" pageNumber="137">Heritiera littoralis</emphasis>
</taxonomicName>
. The
<typeStatus id="54868855FF9D0C10FB8950A0FB23FE34" box="[1116,1164,403,429]" pageId="11" pageNumber="137">type</typeStatus>
series emerged from
<taxonomicName id="4C3D4D74FF9D0C10FA5450A0FF58FE48" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="137" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9D0C10FA5450A0FF58FE48" italics="true" pageId="11" pageNumber="137">H. littoralis</emphasis>
</taxonomicName>
fruits washed ashore (
<figureCitation id="13062A72FF9D0C10FE255084FDECFE48" box="[496,579,438,465]" captionStart="FIGURES 51–54" captionStartId="10.[151,264,1485,1509]" captionTargetBox="[164,1423,561,1461]" captionTargetId="figure-171@10.[164,1423,561,1461]" captionTargetPageId="10" captionText="FIGURES 51–54. Biology of Tepperia vagabunda on Iriomotejima Island, the Ryukyu Islands, Japan. 51, A Heritiera littoralis fruit washed ashore (indicated by an arrow); 52, inside of a H. littoralis fruit infested by the weevil; 53, an adult and a larva in a H. littoralis seed; 54, an adult on a H. littoralis twig." figureDoi="http://doi.org/10.5281/zenodo.15216290" httpUri="https://zenodo.org/record/15216290/files/figure.png" pageId="11" pageNumber="137">Fig. 51</figureCitation>
). Larvae grew in and fed on
<taxonomicName id="4C3D4D74FF9D0C10FCAA5084FBAFFE48" box="[895,1024,439,465]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="137" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9D0C10FCAA5084FC3CFE49" box="[895,915,439,464]" italics="true" pageId="11" pageNumber="137">H</emphasis>
.
<emphasis id="B949EAE5FF9D0C10FC755084FBAFFE48" box="[928,1024,439,465]" italics="true" pageId="11" pageNumber="137">littoralis</emphasis>
</taxonomicName>
seeds (
<figureCitation id="13062A72FF9D0C10FB845084FB44FE48" box="[1105,1259,438,465]" captionStart="FIGURES 51–54" captionStartId="10.[151,264,1485,1509]" captionTargetBox="[164,1423,561,1461]" captionTargetId="figure-171@10.[164,1423,561,1461]" captionTargetPageId="10" captionText="FIGURES 51–54. Biology of Tepperia vagabunda on Iriomotejima Island, the Ryukyu Islands, Japan. 51, A Heritiera littoralis fruit washed ashore (indicated by an arrow); 52, inside of a H. littoralis fruit infested by the weevil; 53, an adult and a larva in a H. littoralis seed; 54, an adult on a H. littoralis twig." figureDoi="http://doi.org/10.5281/zenodo.15216290" httpUri="https://zenodo.org/record/15216290/files/figure.png" pageId="11" pageNumber="137">Figs. 52 &amp; 53</figureCitation>
) and pupated in the seeds when they fully matured. One or more individuals were found inside a seed (
<figureCitation id="13062A72FF9D0C10FB8B50E8FB1EFE6C" box="[1118,1201,474,501]" captionStart="FIGURES 51–54" captionStartId="10.[151,264,1485,1509]" captionTargetBox="[164,1423,561,1461]" captionTargetId="figure-171@10.[164,1423,561,1461]" captionTargetPageId="10" captionText="FIGURES 51–54. Biology of Tepperia vagabunda on Iriomotejima Island, the Ryukyu Islands, Japan. 51, A Heritiera littoralis fruit washed ashore (indicated by an arrow); 52, inside of a H. littoralis fruit infested by the weevil; 53, an adult and a larva in a H. littoralis seed; 54, an adult on a H. littoralis twig." figureDoi="http://doi.org/10.5281/zenodo.15216290" httpUri="https://zenodo.org/record/15216290/files/figure.png" pageId="11" pageNumber="137">Fig. 53</figureCitation>
). New adults emerge through holes made in
<taxonomicName id="4C3D4D74FF9D0C10FE4D50CCFDB5FD80" box="[408,538,511,537]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="137" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF9D0C10FE4D50CCFE03FD81" box="[408,428,511,536]" italics="true" pageId="11" pageNumber="137">H</emphasis>
.
<emphasis id="B949EAE5FF9D0C10FE6F50CCFDB5FD80" box="[442,538,511,537]" italics="true" pageId="11" pageNumber="137">littoralis</emphasis>
</taxonomicName>
pericarps (
<figureCitation id="13062A72FF9D0C10FD4250CCFC99FD80" box="[663,822,510,537]" captionStart="FIGURES 51–54" captionStartId="10.[151,264,1485,1509]" captionTargetBox="[164,1423,561,1461]" captionTargetId="figure-171@10.[164,1423,561,1461]" captionTargetPageId="10" captionText="FIGURES 51–54. Biology of Tepperia vagabunda on Iriomotejima Island, the Ryukyu Islands, Japan. 51, A Heritiera littoralis fruit washed ashore (indicated by an arrow); 52, inside of a H. littoralis fruit infested by the weevil; 53, an adult and a larva in a H. littoralis seed; 54, an adult on a H. littoralis twig." figureDoi="http://doi.org/10.5281/zenodo.15216290" httpUri="https://zenodo.org/record/15216290/files/figure.png" pageId="11" pageNumber="137">Figs. 53 &amp; 54</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

303
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@ -0,0 +1,303 @@
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<mods:title id="D9D09C1DBC365E0084CE6440AC91CFD6">Presence of the genus Tepperia Lea (Coleoptera: Curculionidae) in the Oriental Region, with new combination for a Philippine species and description of a new insular species associated with the looking-glass mangrove, Heritiera littoralis Dryand. ex Aiton (Malvaceae)</mods:title>
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<mods:namePart id="5CBE65521DF5251FC540C16150365C07">Tsuji, Naomichi</mods:namePart>
<mods:affiliation id="E34D8683241F91396B88DA32F9866800">201 Corpo Passion, Komagari, Chichijima, Ogasawara, Tokyo, 100 - 2101 Japan.</mods:affiliation>
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<mods:namePart id="B62B0922D34ED5A7960CC549DF222DB7">Yoshitake, Hiraku</mods:namePart>
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<subSubSection id="C327657CFF950C18FF4257DCFD78F890" box="[151,727,1775,1802]" pageId="3" pageNumber="129" type="nomenclature">
<paragraph id="8B8236F7FF950C18FF4257DCFD78F890" blockId="3.[151,727,1775,1838]" box="[151,727,1775,1802]" pageId="3" pageNumber="129">
<heading id="D0CA819BFF950C18FF4257DCFD78F890" bold="true" box="[151,727,1775,1802]" fontSize="11" level="1" pageId="3" pageNumber="129" reason="1">
<emphasis id="B949EAE5FF950C18FF4257DCFD78F890" bold="true" box="[151,727,1775,1802]" pageId="3" pageNumber="129">
<taxonomicName id="4C3D4D74FF950C18FF4257DCFDECF890" authority="(Heller, 1913)" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" baseAuthorityName="Heller" baseAuthorityYear="1913" box="[151,579,1775,1802]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="palawanica" status="comb. nov.">
<emphasis id="B949EAE5FF950C18FF4257DCFE3DF893" bold="true" box="[151,402,1775,1802]" italics="true" pageId="3" pageNumber="129">Tepperia palawanica</emphasis>
(
<bibRefCitation id="EFAC4B06FF950C18FE7457DCFD95F890" author="Heller, K. M." box="[417,570,1775,1801]" pageId="3" pageNumber="129" pagination="135 - 163" refId="ref7669" refString="Heller, K. M. (1913) Neue Kafer von den Philippinen. Philippine Journal of Science, section D. General biology, ethnology, and anthropology, 8, 135-163." type="journal article" year="1913">Heller, 1913</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A27A579EFF950C18FD8557DCFD78F890" box="[592,727,1775,1801]" pageId="3" pageNumber="129" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF950C18FF425620FE43F8B7" box="[151,492,1811,1838]" pageId="3" pageNumber="129" type="description">
<paragraph id="8B8236F7FF950C18FF425620FE43F8B7" blockId="3.[151,727,1775,1838]" box="[151,492,1811,1838]" pageId="3" pageNumber="129">
(
<figureCitation id="13062A72FF950C18FF4A5620FE88F8B4" box="[159,295,1811,1837]" captionStart="FIGURES 12–18" captionStartId="4.[151,264,1626,1650]" captionTargetBox="[163,1424,309,1602]" captionTargetId="figure-67@4.[163,1424,309,1603]" captionTargetPageId="4" captionText="FIGURES 12–18. Tepperia spp., males. 12–14, Cyamobolus palawanicus Heller, 1913, lectotype; 15–18, Tepperia vagabunda, holotype. 12 &amp; 15, Dorsal habitus; 13 &amp; 16, lateral habitus; 14a, attached labels, surface; 14b, one of the labels, back side; 17, meso- and metaventrites; 18, venter. Scale: 2.0 mm for 12, 13, 15, 16 &amp; 18; 1.0 mm for 17." figureDoi="http://doi.org/10.5281/zenodo.15216280" httpUri="https://zenodo.org/record/15216280/files/figure.png" pageId="3" pageNumber="129">Figs. 12–14</figureCitation>
,
<figureCitation id="13062A72FF950C18FEE65620FED7F8B4" box="[307,376,1811,1837]" captionStart="FIGURES 19–24" captionStartId="7.[151,264,1412,1436]" captionTargetBox="[166,1421,181,1388]" captionTargetId="figure-28@7.[166,1421,181,1389]" captionTargetPageId="7" captionText="FIGURES 19–24. Tepperia spp. 19–21, Tepperia palawanica (Heller, 1913); 22–24, Tepperia vagabunda; 19, 20, 22 &amp; 23, heads, dorsal aspect; 21 &amp; 24, left mesotibiae, anterior aspect. 19 &amp; 21, Cyamobolus palawanicus Heller, 1913, lectotype, male; 20, ditto, paralectotype, female. 22 &amp; 24, Holotype, male; 23, paratype, female. Scale: 2.0 mm for 19, 20, 22 &amp; 23, 1.5 mm for 21 &amp; 24." figureDoi="http://doi.org/10.5281/zenodo.15216282" httpUri="https://zenodo.org/record/15216282/files/figure.png" pageId="3" pageNumber="129">19–21</figureCitation>
&amp;
<figureCitation id="13062A72FF950C18FE485620FE4BF8B7" box="[413,484,1811,1838]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="3" pageNumber="129">25–36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF950C18FF42566BFB82F817" pageId="3" pageNumber="129" type="reference_group">
<paragraph id="8B8236F7FF950C18FF42566BFB82F817" blockId="3.[151,1436,1880,1935]" pageId="3" pageNumber="129">
<treatmentCitationGroup id="AB2D11D9FF950C18FF42566BFB82F817" pageId="3" pageNumber="129">
<treatmentCitation id="0A9C10E6FF950C18FF42566BFDFAF8E9" author="Heller, K. M." box="[151,597,1880,1904]" page="140" pageId="3" pageNumber="129" year="1913">
<taxonomicName id="4C3D4D74FF950C18FF42566BFDFAF8E9" ID-CoL="6BQZM" authority="Heller, 1913: 140" authorityName="Heller" authorityPageNumber="140" authorityYear="1913" box="[151,597,1880,1904]" class="Insecta" family="Curculionidae" genus="Cyamobolus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="species" species="palawanicus">
<emphasis id="B949EAE5FF950C18FF42566BFE32F8E9" box="[151,413,1880,1904]" italics="true" pageId="3" pageNumber="129">Cyamobolus palawanicus</emphasis>
<bibRefCitation id="EFAC4B06FF950C18FE76566BFDFAF8E9" author="Heller, K. M." box="[419,597,1880,1904]" pageId="3" pageNumber="129" pagination="135 - 163" refId="ref7669" refString="Heller, K. M. (1913) Neue Kafer von den Philippinen. Philippine Journal of Science, section D. General biology, ethnology, and anthropology, 8, 135-163." type="journal article" year="1913">Heller, 1913: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54868855FF950C18FDB6566BFD3FF8E9" box="[611,656,1880,1904]" pageId="3" pageNumber="129">type</typeStatus>
locality: “
<collectingRegion id="49F9F815FF950C18FD2D566BFCD8F8E9" box="[760,887,1880,1904]" country="Philippines" name="Palawan" pageId="3" pageNumber="129">PALAWAN</collectingRegion>
”) &amp; fig 2;
<treatmentCitation id="0A9C10E6FF950C18FC30566BFB00F8E9" author="Schultze, W." box="[997,1199,1880,1904]" page="144" pageId="3" pageNumber="129" year="1916">
<bibRefCitation id="EFAC4B06FF950C18FC30566BFB00F8E9" author="Schultze, W." box="[997,1199,1880,1904]" pageId="3" pageNumber="129" pagination="95 - 194" refId="ref8351" refString="Schultze, W. (1916) A catalogue of Philippine Coleoptera. Philippine journal of science, section D. General biology, ethnology, and anthropology, 11, 95-194." type="journal article" year="1916">Schultze, 1916: 144</bibRefCitation>
</treatmentCitation>
(cataloged:
<collectingRegion id="49F9F815FF950C18FAF8566BFACDF8E9" box="[1325,1378,1880,1904]" country="Philippines" name="Cebu" pageId="3" pageNumber="129">Cebu</collectingRegion>
Is. &amp;
<collectingRegion id="49F9F815FF950C18FF125645FEB2F817" box="[199,285,1910,1934]" country="Philippines" name="Palawan" pageId="3" pageNumber="129">Palawan</collectingRegion>
Is.) &amp; 184 (biological note);
<treatmentCitation id="0A9C10E6FF950C18FD925645FCB7F816" author="Hustache, A." box="[583,792,1910,1935]" page="238" pageId="3" pageNumber="129" year="1936">
<bibRefCitation id="EFAC4B06FF950C18FD925645FCB7F816" author="Hustache, A." box="[583,792,1910,1935]" pageId="3" pageNumber="129" pagination="1 - 317" refId="ref7706" refString="Hustache, A. (1936) Curculionidae: Cryptorrhynchinae. In: Schenkling, S. (Ed.), Coleoptelorum Catalogus auspiciis et auxilio W. Junk. Vol. 151. Dr. W. Junk, s' - Gravenhage, pp. 1-317." type="book chapter" year="1936">Hustache, 1936: 238</bibRefCitation>
</treatmentCitation>
(cataloged: “
<collectingCountry id="F32A7667FF950C18FC4B5645FBB6F817" box="[926,1049,1910,1934]" name="Philippines" pageId="3" pageNumber="129">Philippines</collectingCountry>
”).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF950C1FFF42568FFF4DFF4C" lastPageId="4" lastPageNumber="130" pageId="3" pageNumber="129" type="diagnosis">
<paragraph id="8B8236F7FF950C1FFF42568FFF4DFF4C" blockId="3.[151,1437,1980,2043]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="130" pageId="3" pageNumber="129">
<emphasis id="B949EAE5FF950C18FF42568FFEBDF84F" bold="true" box="[151,274,1980,2006]" pageId="3" pageNumber="129">Diagnosis.</emphasis>
From all known
<taxonomicName id="4C3D4D74FF950C18FE19568EFD80F84F" box="[460,559,1981,2006]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAE5FF950C18FE19568EFD80F84F" box="[460,559,1981,2006]" italics="true" pageId="3" pageNumber="129">Tepperia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3D4D74FF950C18FDB7568EFC85F84E" box="[610,810,1981,2007]" pageId="3" pageNumber="129">
<emphasis id="B949EAE5FF950C18FDB7568EFD7FF84F" box="[610,720,1981,2006]" italics="true" pageId="3" pageNumber="129">Evaniocis</emphasis>
species
</taxonomicName>
, this species can be easily distinguished by the pronotum and elytra covered with light-colored scales except several brown scaly markings. In addition, this species is characterized by the rostrum whose dorsal contour is slightly curved and the elytral humeri covered by dark brown scales.
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF920C1DFF1251EDFCA6FBC0" lastPageId="6" lastPageNumber="132" pageId="4" pageNumber="130" type="description">
<paragraph id="8B8236F7FF920C1FFF1251EDFC70FE84" blockId="4.[151,1437,151,285]" pageId="4" pageNumber="130">
<emphasis id="B949EAE5FF920C1FFF1251EDFE16FF60" bold="true" box="[199,441,222,249]" pageId="4" pageNumber="130">Redescription. Male</emphasis>
. Measurements (n = 4; in mm): TL: 7.9–8.6 (
<typeStatus id="54868855FF920C1FFC6951ECFB8AFF60" box="[956,1061,223,249]" pageId="4" pageNumber="130" type="lectotype">lectotype</typeStatus>
7.9, mean 8.1); LP: 2.4–2.5 (2.4, 2.4); WP: 2.6–2.7 (2.6, 2.7); LE: 5.5–6.0 (5.5, 5.6); WE: 3.5–3.8 (3.5, 3.6).
</paragraph>
<caption id="DF42667FFF920C1FFF425769FB9CF922" ID-DOI="http://doi.org/10.5281/zenodo.15216280" ID-Zenodo-Dep="15216280" httpUri="https://zenodo.org/record/15216280/files/figure.png" pageId="4" pageNumber="130" startId="4.[151,264,1626,1650]" targetBox="[163,1424,309,1602]" targetPageId="4" targetType="figure">
<paragraph id="8B8236F7FF920C1FFF425769FB9CF922" blockId="4.[151,1437,1626,1723]" pageId="4" pageNumber="130">
<emphasis id="B949EAE5FF920C1FFF425769FEFCF9EB" bold="true" box="[151,339,1626,1650]" pageId="4" pageNumber="130">FIGURES 12–18.</emphasis>
<taxonomicName id="4C3D4D74FF920C1FFE8C5768FE4BF9EB" box="[345,484,1627,1650]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B949EAE5FF920C1FFE8C5768FE1DF9EB" box="[345,434,1627,1650]" italics="true" pageId="4" pageNumber="130">Tepperia</emphasis>
spp.
</taxonomicName>
, males. 12–14,
<taxonomicName id="4C3D4D74FF920C1FFDAA5769FBA7F9EA" authority="Heller, 1913" authorityName="Heller" authorityYear="1913" box="[639,1032,1626,1651]" class="Insecta" family="Curculionidae" genus="Cyamobolus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="palawanicus">
<emphasis id="B949EAE5FF920C1FFDAA5769FC2CF9EB" box="[639,899,1626,1650]" italics="true" pageId="4" pageNumber="130">Cyamobolus palawanicus</emphasis>
<bibRefCitation id="EFAC4B06FF920C1FFC5C5769FBA7F9EA" author="Heller, K. M." box="[905,1032,1626,1651]" pageId="4" pageNumber="130" pagination="135 - 163" refId="ref7669" refString="Heller, K. M. (1913) Neue Kafer von den Philippinen. Philippine Journal of Science, section D. General biology, ethnology, and anthropology, 8, 135-163." type="journal article" year="1913">Heller, 1913</bibRefCitation>
</taxonomicName>
, lectotype; 15–18,
<taxonomicName id="4C3D4D74FF920C1FFB125768FA39F9EB" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" box="[1223,1430,1626,1650]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="130" phylum="Arthropoda" rank="species" species="vagabunda">
<emphasis id="B949EAE5FF920C1FFB125768FA39F9EB" box="[1223,1430,1626,1650]" italics="true" pageId="4" pageNumber="130">Tepperia vagabunda</emphasis>
</taxonomicName>
, holotype. 12 &amp; 15, Dorsal habitus; 13 &amp; 16, lateral habitus; 14a, attached labels, surface; 14b, one of the labels, back side; 17, meso- and metaventrites; 18, venter. Scale: 2.0 mm for 12, 13, 15, 16 &amp; 18; 1.0 mm for 17.
</paragraph>
</caption>
<paragraph id="8B8236F7FF920C1EFF12563EFB0FFD1C" blockId="4.[151,1437,1805,2011]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="131" pageId="4" pageNumber="130">Integument reddish to dark reddish brown. Body generally covered with dull-white to yellowish brown recumbent scales. Frons and vertex covered with subovate scales; medial part of vertex with ill-defined dark brown patch. Rostrum covered with brownish subovate scales on basal half; apical margin of squamous part shallowly notched; apical half subglabrous, sparsely covered with minute scales. Pronotum covered with subovate scales, lacking suberect scales, with two pairs of dark brown scaly patches on postmedian and median parts along midline; postmedian patches connected with median ones by slightly paler scales, sometimes become obscure; median patches elongate apically with slightly paler scales; apical part covered with darker scales along margins except middle, with imbricate scales medially. Lateral parts of prothorax covered with general scales sparsely mingled with suberect scales, with irregular patch of brownish scales on middle of each side; prosternal canal mostly glabrous, but apical part densely covered with light-colored slender scales. Scutellar shield glabrous. Elytra mostly covered with subovate scales, usually with three pairs of irregular patches of dark brown scales on basal part between 2nd and 5th intervals, near humeri, and behind middle between 2nd and 4th intervals, sometimes with additional pair of patches on subapical swellings; basal patches indistinctly marginated and connected laterally with humeral patches by paler scales; sutural intervals covered with smaller scales, which become sparser on basal part; strial punctures each bearing whitish subrecumbent spathulate scale. Mesoventral receptacle glabrous on inside of receptacle canal, densely covered with minute scales on apical margin. Mesepimera, mesanepisterna, and lateral parts of mesoventrite mostly covered with subovate scales. Metaventrite and metanepisterna also covered with light-colored subovate scales; metaventrite glabrous on postmedial part; sclerolepidia absent. Femora irregularly mottled with light-colored to brown scales on anterior surfaces; each femur with triangular glabrous portion on apico-ventral part. Abdominal ventrites closely covered with pale brown scales, sparsely mingled with subrecumbent ones.</paragraph>
<paragraph id="8B8236F7FF930C1EFF1253BCFEB6FBAC" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="131">
<emphasis id="B949EAE5FF930C1EFF1253BCFEA8FD30" bold="true" box="[199,263,655,681]" pageId="5" pageNumber="131">Head</emphasis>
(
<figureCitation id="13062A72FF930C1EFECD53BCFEC4FD30" box="[280,363,655,681]" captionStart="FIGURES 19–24" captionStartId="7.[151,264,1412,1436]" captionTargetBox="[166,1421,181,1388]" captionTargetId="figure-28@7.[166,1421,181,1389]" captionTargetPageId="7" captionText="FIGURES 19–24. Tepperia spp. 19–21, Tepperia palawanica (Heller, 1913); 22–24, Tepperia vagabunda; 19, 20, 22 &amp; 23, heads, dorsal aspect; 21 &amp; 24, left mesotibiae, anterior aspect. 19 &amp; 21, Cyamobolus palawanicus Heller, 1913, lectotype, male; 20, ditto, paralectotype, female. 22 &amp; 24, Holotype, male; 23, paratype, female. Scale: 2.0 mm for 19, 20, 22 &amp; 23, 1.5 mm for 21 &amp; 24." figureDoi="http://doi.org/10.5281/zenodo.15216282" httpUri="https://zenodo.org/record/15216282/files/figure.png" pageId="5" pageNumber="131">Fig. 19</figureCitation>
). Apical part of vertex gently flattened medially; frons flattened, 0.8 times as wide as base of rostrum at narrowest point between eyes. Rostrum stout, LR/WRM 2.4–2.5, LR/LP 0.7, in profile slightly curved ventrad, very slightly attenuate toward apex, excepting apical part more strongly attenuate; dorsal surface mostly slightly convex but sometimes slightly carinate along midline, rugosely to densely punctate from base to apical part, excepting impunctate portion along midline; punctures become smaller toward apex; medial part of apex depressed, impunctate, flanked by two shallow oblique furrows; sides gradually attenuate from base to before middle, then subparallel to subapical part, and finally weakly bulged near apex; apical margin shallowly arcuate in median part; dorsal contour slightly arcuate in profile. Antennae inserted at middle; scape strongly clavate, slightly shorter than funicle; funicle with 1st and 2nd segments subconical, with 3rd to 7th barrel-shaped, successively dilated distally; approximate length ratio of 1st to 7th as follows: 3: 2: 1: 1: 1: 1: 1.3; club fusiform, 2.0–2.5 times as long as wide, 0.6–0.7 times as long as funicle, loosely segmented. Mandibles robust, each with rounded outer margin and large inner tooth.
</paragraph>
<paragraph id="8B8236F7FF930C1EFF12550CFB3FFB70" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="131">
<emphasis id="B949EAE5FF930C1EFF12550CFE8FFBC0" bold="true" box="[199,288,1087,1113]" pageId="5" pageNumber="131">Thorax</emphasis>
. Prothorax subconical, LP/WP 0.9, weakly constricted subapically; pronotum densely punctate, uniformly convex, indistinctly carinate along midline; sides parallel or slightly attenuate from base to behind middle, then roundly convergent to subapical constriction, and finally slightly attenuate to apical margin, which is strongly arched in dorsal view; lateral parts rugosely punctate, obscurely granulate on middle. Mesoscutellum with scutellar shield round to subovate, approximately as wide as elytral sutural interval, weakly convex.
</paragraph>
<paragraph id="8B8236F7FF930C1EFF1255C0FE3FF9B4" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="131">Elytra subovate, LE/WE 1.5–1.6, with weakly produced humeri, wider than prothorax, WE/WP 1.3–1.4, evenly convex excepting subbasal part weakly prominent between 2nd to 4th intervals, with weak subapical swellings; sides subparallel from humeri to apical 2/5, then gently arcuately convergent to subapical swellings, and finally rounded apically, slightly emarginate at apices. Intervals weakly convex, wider than striae, partially granulate; sutural ones faintly depressed near base; 8th and 9th irregularly granulate on humeri; 9th granulate behind humeri to just before middle, then granulate-costate to subapical part; 10th flattened basally, granulate-costate excepting base. Striae shallowly sulcate excepting middle of 9th more deeply sulcate; 10th relatively short, terminated before middle; ventral surface with submarginal ridges sublinear in ventral aspect, not arched anteriorly, with two asymmetric files on apico-internal part.</paragraph>
<paragraph id="8B8236F7FF930C1EFF125704FB46F8D4" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="131">
Femora robust, gently expanded from base to before middle, not clavate, with ventral tooth near middle, shallowly ventrally sulcate from middle to apex; dorsal contour weakly arcuate; profemora slightly longer than either meso- or metafemora. Tibiae simple along outer margins, not tuberculate; pro- and mesotibiae (
<figureCitation id="13062A72FF930C1EFB2D574CFAE7F900" box="[1272,1352,1663,1689]" captionStart="FIGURES 19–24" captionStartId="7.[151,264,1412,1436]" captionTargetBox="[166,1421,181,1388]" captionTargetId="figure-28@7.[166,1421,181,1389]" captionTargetPageId="7" captionText="FIGURES 19–24. Tepperia spp. 19–21, Tepperia palawanica (Heller, 1913); 22–24, Tepperia vagabunda; 19, 20, 22 &amp; 23, heads, dorsal aspect; 21 &amp; 24, left mesotibiae, anterior aspect. 19 &amp; 21, Cyamobolus palawanicus Heller, 1913, lectotype, male; 20, ditto, paralectotype, female. 22 &amp; 24, Holotype, male; 23, paratype, female. Scale: 2.0 mm for 19, 20, 22 &amp; 23, 1.5 mm for 21 &amp; 24." figureDoi="http://doi.org/10.5281/zenodo.15216282" httpUri="https://zenodo.org/record/15216282/files/figure.png" pageId="5" pageNumber="131">Fig. 21</figureCitation>
) gently attenuate toward apex, obtusely angulate basally on outer margins; mesotibiae slightly shorter than either pro- or metatibiae, hardly to barely curved inward; metatibiae subparallel-sided, more weakly angulate basally on outer margins than on pro- and mesotibiae. Tarsi stout; 1st tarsomere subconical, 1.6 times as wide as long, approximately twice as long as and as wide as 2nd, 1.1–1.3 times as long as 3rd; 2nd subrectangular, 1.3–1.5 times as wide as long, half as wide as 3rd; 3rd subcordate, 1.5 times as wide as long; 5th clavate, slightly longer than 3rd.
</paragraph>
<paragraph id="8B8236F7FF930C1DFF125664FC05FF4C" blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1437,151,1689]" lastPageId="6" lastPageNumber="132" pageId="5" pageNumber="131">Rostral canal reaching level between anterior margins of mesocoxae; prosternal canal approximately as wide as procoxa, gradually expanded toward apex; mesoventral receptacle subtrapezoidal, 1.3–1.4 times as wide as long, shallowly concave along anterior margin, strongly bulged ventrally, overhanging anteriorly above canal, declined posteriorly, on declivity, indistinctly costate along midline and compressed laterally. Metaventrite flattened or slightly depressed medially, 1.3–1.6 times as long as mesoventrite along midline; apical part triangularly depressed; apical depression declined medially, slightly impressed along midline.</paragraph>
<paragraph id="8B8236F7FF900C1DFF1251EDFC66FEFC" blockId="6.[151,1437,151,1689]" pageId="6" pageNumber="132">
<emphasis id="B949EAE5FF900C1DFF1251EDFE93FF61" bold="true" box="[199,316,222,248]" pageId="6" pageNumber="132">Abdomen</emphasis>
. Ventrites irregularly punctate; ventrite I with abdominal process narrowly acuminate, approximately 1.4 times as long as II along midline, flattened medially, shallowly emarginate along apical margin; II slightly longer than III and IV combined; III and IV subequal in length, each punctate in distorted row; V subtrapezoidal, approximately 3.0 times as wide as long, with apical margin subtruncate.
</paragraph>
<paragraph id="8B8236F7FF900C1DFF12505DFB0DFD30" blockId="6.[151,1437,151,1689]" pageId="6" pageNumber="132">
<emphasis id="B949EAE5FF900C1DFF12505DFEE5FE11" bold="true" box="[199,330,366,392]" pageId="6" pageNumber="132">Terminalia</emphasis>
. Tergite VII (
<figureCitation id="13062A72FF900C1DFE36505CFD9BFE10" box="[483,564,366,393]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 25</figureCitation>
) transverse, subtrapezoidal, with two longitudinal distorted rows of setiferous tubercles on medial part. Rectum with rectal ring. Penis (
<figureCitation id="13062A72FF900C1DFCC650A0FC01FE34" box="[787,942,403,429]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Figs. 26 &amp; 27</figureCitation>
) with body much shorter than its apodemes, approximately 1.5 times as long as wide, weakly sclerotized dorsally, subparallel-sided from base to subapical part, in lateral view flattened dorso-ventrally, moderately curved ventrally; apex (
<figureCitation id="13062A72FF900C1DFC2950E8FBE0FE6C" box="[1020,1103,475,501]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 28</figureCitation>
) rounded, flanked by several minute setae; endophallus spinulate basally, with tubular copulatory sclerite which is widened apically, dorsally produced in two laminae apically. Tegminal ring (
<figureCitation id="13062A72FF900C1DFD0D5310FC83FDA4" box="[728,812,547,573]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 29</figureCitation>
) with weakly sclerotized parameroid lobes, which are fused basally; diameter of tegminal ring longer than its apodeme. Sternite VIII (
<figureCitation id="13062A72FF900C1DFBC55374FBCEFDF8" box="[1040,1121,583,609]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 30</figureCitation>
) composed of two narrowly separated sublunate sclerites; spiculum relictum present, slightly sclerotized (
<figureCitation id="13062A72FF900C1DFC2A5358FBFBFD1C" box="[1023,1108,619,645]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 30</figureCitation>
). Spiculum gastrale (
<figureCitation id="13062A72FF900C1DFA975358FA3BFD1C" box="[1346,1428,619,645]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 31</figureCitation>
) shorter than penis; apical arms short, widely divergent, with narrow basal sclerotized flange.
</paragraph>
<paragraph id="8B8236F7FF900C1DFF125380FA99FCA0" blockId="6.[151,1437,151,1689]" pageId="6" pageNumber="132">
<emphasis id="B949EAE5FF900C1DFF125380FEB1FD54" bold="true" box="[199,286,691,717]" pageId="6" pageNumber="132">Female</emphasis>
. Measurements (n = 3; in mm): TL 6.7–8.7 (mean 7.8), LP 2.1–2.6 (2.4); WP 2.3–2.9 (2.6); LE 4.6–6.1 (5.4); WE 3.0–3.9 (3.5). Rostrum (
<figureCitation id="13062A72FF900C1DFDC353E4FDC8FD68" box="[534,615,727,753]" captionStart="FIGURES 19–24" captionStartId="7.[151,264,1412,1436]" captionTargetBox="[166,1421,181,1388]" captionTargetId="figure-28@7.[166,1421,181,1389]" captionTargetPageId="7" captionText="FIGURES 19–24. Tepperia spp. 19–21, Tepperia palawanica (Heller, 1913); 22–24, Tepperia vagabunda; 19, 20, 22 &amp; 23, heads, dorsal aspect; 21 &amp; 24, left mesotibiae, anterior aspect. 19 &amp; 21, Cyamobolus palawanicus Heller, 1913, lectotype, male; 20, ditto, paralectotype, female. 22 &amp; 24, Holotype, male; 23, paratype, female. Scale: 2.0 mm for 19, 20, 22 &amp; 23, 1.5 mm for 21 &amp; 24." figureDoi="http://doi.org/10.5281/zenodo.15216282" httpUri="https://zenodo.org/record/15216282/files/figure.png" pageId="6" pageNumber="132">Fig. 20</figureCitation>
) LR/WRM 2.7, LR/LP 0.6–0.7; lateral parts of dorsal surface with sparser punctures on apical half; sides more strongly attenuate from base to middle. Prothorax LP/WP 0.9–1.0. Elytra LE/ WE 1.5–1.6, WE/WP 1.3–1.4. Ventrite I slightly inflated medially excepting abdominal process flattened.
</paragraph>
<paragraph id="8B8236F7FF900C1DFF125271FCA6FBC0" blockId="6.[151,1437,151,1689]" pageId="6" pageNumber="132">
<emphasis id="B949EAE5FF900C1DFF125271FEE5FCC5" bold="true" box="[199,330,834,860]" pageId="6" pageNumber="132">Terminalia</emphasis>
. Tergite VII (
<figureCitation id="13062A72FF900C1DFE345270FD9DFCC4" box="[481,562,835,861]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 32</figureCitation>
) subtrapezoidal, slightly wider than long, with two longitudinal distorted rows of setiferous tubercles on medial part; apical margin gently arcuate; tergite VIII (
<figureCitation id="13062A72FF900C1DFBC85254FBC0FC18" box="[1053,1135,870,897]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 35</figureCitation>
) slightly longer than wide; sides rapidly tapered apicad; apical margin weakly arcuate, notched at middle. Spiculum ventrale (
<figureCitation id="13062A72FF900C1DFAD052B8FAF6FC3C" box="[1285,1369,907,933]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 36</figureCitation>
) with apical plate subovate, gradually tapered apicad, notched at middle of apical margin; apodemes short, approximately as long as apical plate. Ovipositor (
<figureCitation id="13062A72FF900C1DFDF052E0FDD8FC74" box="[549,631,979,1005]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 33</figureCitation>
) with styli slender, approximately 1/3 length of gonocoxite; vagina and bursa copulatrix simple, neither spinulate nor sclerotized. Spermatheca (
<figureCitation id="13062A72FF900C1DFC1952C4FBB1FB88" box="[972,1054,1015,1041]" captionStart="FIGURES 25–36" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[160,1427,607,1850]" captionTargetId="figure-198@8.[151,1436,597,1864]" captionTargetPageId="8" captionText="FIGURES 25–36. Terminalia of Tepperia palawanica (Heller, 1913). 25–31, Male; 32–36, female. 25 &amp; 32, Tergites VII; 26 &amp; 27, penis; 28, apex of penis, at different angle; 29, tegmen; 30, sternite VIII; 31, spiculum gastrale; 33, right stylus and gonocoxite; 34, spermatheca; 35. tergite VIII; 36, spiculum ventrale. 25, 26, 29, 32, 33 &amp; 35, Dorsal aspect; 27, lateral aspect; 30, 31 &amp; 36, ventral aspect. Scale A: 0.5 mm for 25, 32, 35 &amp; 36; 0.2 mm for 33, 34. Scale B: 0.2 mm for 26–31." figureDoi="http://doi.org/10.5281/zenodo.15216286" httpUri="https://zenodo.org/record/15216286/files/figure.png" pageId="6" pageNumber="132">Fig. 34</figureCitation>
) c-shaped; cornu short, gradually attenuate and slightly curved apically; collum obliquely convex downward; ramus vestigial; insertions of duct and gland widely separated; gland short, sclerotized basally.
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF900C1DFF125550FF50F9B4" pageId="6" pageNumber="132" type="materials_examined">
<paragraph id="8B8236F7FF900C1DFF125550FE61FA7C" blockId="6.[151,1437,151,1689]" pageId="6" pageNumber="132">
<emphasis id="B949EAE5FF900C1DFF125550FE42FBE5" bold="true" box="[199,493,1122,1149]" pageId="6" pageNumber="132">Type material examined.</emphasis>
<typeStatus id="54868855FF900C1DFE265550FDCCFBE4" box="[499,611,1123,1149]" pageId="6" pageNumber="132" type="lectotype">Lectotype</typeStatus>
(here designated;
<collectionCode id="ED2CAE32FF900C1DFCF85550FCD0FBE4" box="[813,895,1123,1149]" pageId="6" pageNumber="132">SMTD</collectionCode>
), male: “typus” (surface, typed) and “palawani- / cus Heller” (back side, handwritten) (red card), “Acc. No. 6108 / Lot / Bu.of Sci., P.I.” (typed and partly handwritten on a white card), “Collected by / E. D. Merrill.” (typed on a pale brown card); “1911 / 7” (typed and partially handwritten on a yellow card), “Staatl. Museum für / Tiekunde Dresden” (typed on a white card), and “
<typeStatus id="54868855FF900C1DFB2F55FCFA32FB70" box="[1274,1437,1231,1257]" pageId="6" pageNumber="132" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="4C3D4D74FF900C1DFF7C55C0FE9AFA94" authorityName="Schoenherr" authorityYear="1837" box="[169,309,1267,1293]" class="Insecta" family="Curculionidae" genus="Cyamobolus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAE5FF900C1DFF7C55C0FE9AFA94" box="[169,309,1267,1293]" italics="true" pageId="6" pageNumber="132">Cyamobolus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C3D4D74FF900C1DFE8455C0FE70FA94" authorityName="Heller" authorityYear="1913" box="[337,479,1267,1293]" class="Insecta" family="Curculionidae" genus="Cyamobolus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="132" phylum="Arthropoda" rank="species" species="palawanicus">
<emphasis id="B949EAE5FF900C1DFE8455C0FE70FA94" box="[337,479,1267,1293]" italics="true" pageId="6" pageNumber="132">palawanicus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFAC4B06FF900C1DFE2F55C0FD27FA94" author="Heller, K. M." box="[506,648,1267,1293]" pageId="6" pageNumber="132" pagination="135 - 163" refId="ref7669" refString="Heller, K. M. (1913) Neue Kafer von den Philippinen. Philippine Journal of Science, section D. General biology, ethnology, and anthropology, 8, 135-163." type="journal article" year="1913">Heller, 1913</bibRefCitation>
/ Des. N. Tsuji &amp; H. Yoshitake, 2025” (typed on a red card; newly attached).
<typeStatus id="54868855FF900C1DFEDE5424FE09FAA8" box="[267,422,1303,1329]" pageId="6" pageNumber="132" type="paralectotype">Paralectotype</typeStatus>
,
<specimenCount id="9D3BFD7EFF900C1DFE675424FDB8FAA8" box="[434,535,1303,1329]" count="1" pageId="6" pageNumber="132" type="female">1 female</specimenCount>
: “Acc. No. 6108 / Lot / Bu.of Sci., P.I.” (typed and partly handwritten on a white card), “Collected by / E. D. Merrill.” (typed on a pale brown card); “1911 / 7” (typed and partially handwritten on a yellow card), “
<typeStatus id="54868855FF900C1DFEAD546CFE4FFAE0" box="[376,480,1375,1401]" pageId="6" pageNumber="132" type="syntype">syntypus</typeStatus>
” (typed on a reddish orange card), “Staatl. Museum für / Tiekunde Dresden” (typed on a white card), “bei
<taxonomicName id="4C3D4D74FF900C1DFE4E54B0FDB4FA04" box="[411,539,1411,1437]" class="Insecta" family="Curculionidae" genus="Metacymia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="132" phylum="Arthropoda" rank="genus">Metacymia</taxonomicName>
? / Heller det: 1934” (partially handwritten and stamped on a white card), and “
<typeStatus id="54868855FF900C1DFF745494FE23FA58" box="[161,396,1447,1473]" pageId="6" pageNumber="132" type="paralectotype">PARALECTOTYPE</typeStatus>
/
<taxonomicName id="4C3D4D74FF900C1DFE775494FD81FA58" authorityName="Schoenherr" authorityYear="1837" box="[418,558,1447,1473]" class="Insecta" family="Curculionidae" genus="Cyamobolus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B949EAE5FF900C1DFE775494FD81FA58" box="[418,558,1447,1473]" italics="true" pageId="6" pageNumber="132">Cyamobolus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C3D4D74FF900C1DFD935494FD7BFA58" authorityName="Heller" authorityYear="1913" box="[582,724,1447,1473]" class="Insecta" family="Curculionidae" genus="Cyamobolus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="132" phylum="Arthropoda" rank="species" species="palawanicus">
<emphasis id="B949EAE5FF900C1DFD935494FD7BFA58" box="[582,724,1447,1473]" italics="true" pageId="6" pageNumber="132">palawanicus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFAC4B06FF900C1DFD3F5494FCD9FA58" author="Heller, K. M." box="[746,886,1447,1473]" pageId="6" pageNumber="132" pagination="135 - 163" refId="ref7669" refString="Heller, K. M. (1913) Neue Kafer von den Philippinen. Philippine Journal of Science, section D. General biology, ethnology, and anthropology, 8, 135-163." type="journal article" year="1913">Heller, 1913</bibRefCitation>
/ Des. N. Tsuji &amp; H. Yoshitake, 2025” (typed on a red card; newly attached).
</paragraph>
<paragraph id="8B8236F7FF900C1DFF1254DCFF50F9B4" blockId="6.[151,1437,151,1689]" pageId="6" pageNumber="132">
<emphasis id="B949EAE5FF900C1DFF1254DCFD89F991" bold="true" box="[199,550,1518,1545]" pageId="6" pageNumber="132">Non-type material examined.</emphasis>
<materialsCitation id="3B553CAAFF900C1DFDFB54DCFF54F9B4" collectionCode="SMTD" collectorName="Taytay, W &amp; W. Schultze" country="Philippines" location="The" pageId="6" pageNumber="132" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Palawan">
<location id="8EE2602CFF900C1DFDFB54DCFDF5F990" LSID="urn:lsid:plazi:treatment:039487E1FF950C1DFF4257DCFC31F900:8EE2602CFF900C1DFDFB54DCFDF5F990" box="[558,602,1519,1545]" country="Philippines" name="The" pageId="6" pageNumber="132" stateProvince="Palawan">The</location>
<collectingCountry id="F32A7667FF900C1DFDB754DCFD4AF990" box="[610,741,1519,1545]" name="Philippines" pageId="6" pageNumber="132">Philippines</collectingCountry>
: [
<collectingRegion id="49F9F815FF900C1DFD2254DCFCF6F990" box="[759,857,1519,1545]" country="Philippines" name="Palawan" pageId="6" pageNumber="132">Palawan</collectingRegion>
Is.]
<specimenCount id="9D3BFD7EFF900C1DFC5B54DCFC49F990" box="[910,998,1519,1545]" count="3" pageId="6" pageNumber="132" type="male">3 males</specimenCount>
&amp;
<specimenCount id="9D3BFD7EFF900C1DFBD854DCFBD2F990" box="[1037,1149,1519,1545]" count="2" pageId="6" pageNumber="132" type="female">2 females</specimenCount>
,
<collectorName id="26C85321FF900C1DFB5F54DCFB50F990" box="[1162,1279,1519,1545]" pageId="6" pageNumber="132">Taytay, W</collectorName>
. Schultze leg. (
<collectionCode id="ED2CAE32FF900C1DFF4A5720FF5CF9B4" box="[159,243,1555,1581]" pageId="6" pageNumber="132">SMTD</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C327657CFF900C1DFF125705FC31F900" pageId="6" pageNumber="132" type="distribution">
<paragraph id="8B8236F7FF900C1DFF125705FC0FF9C8" blockId="6.[151,1437,151,1689]" box="[199,928,1590,1617]" pageId="6" pageNumber="132">
<emphasis id="B949EAE5FF900C1DFF125705FECEF9C9" bold="true" box="[199,353,1590,1616]" pageId="6" pageNumber="132">Distribution.</emphasis>
The
<collectingCountry id="F32A7667FF900C1DFE4C5704FDB8F9C8" box="[409,535,1591,1617]" name="Philippines" pageId="6" pageNumber="132">Philippines</collectingCountry>
(
<collectingRegion id="49F9F815FF900C1DFDF35704FDCCF9C8" box="[550,611,1591,1617]" country="Philippines" name="Cebu" pageId="6" pageNumber="132">Cebu</collectingRegion>
Island and
<collectingRegion id="49F9F815FF900C1DFD305704FCEBF9C8" box="[741,836,1591,1617]" country="Philippines" name="Palawan" pageId="6" pageNumber="132">Palawan</collectingRegion>
Island).
</paragraph>
<paragraph id="8B8236F7FF900C1DFF125768FC31F900" blockId="6.[151,1437,151,1689]" pageId="6" pageNumber="132">
<emphasis id="B949EAE5FF900C1DFF125768FE29F9ED" bold="true" box="[199,390,1626,1653]" pageId="6" pageNumber="132">Natural history.</emphasis>
<bibRefCitation id="EFAC4B06FF900C1DFE5B5768FDD5F9EC" author="Schultze, W." box="[398,634,1627,1653]" pageId="6" pageNumber="132" pagination="95 - 194" refId="ref8351" refString="Schultze, W. (1916) A catalogue of Philippine Coleoptera. Philippine journal of science, section D. General biology, ethnology, and anthropology, 11, 95-194." type="journal article" year="1916">Schultze (1916: 184)</bibRefCitation>
mentioned as “Very injurious to seeds of
<taxonomicName id="4C3D4D74FF900C1DFB805768FA21F9EC" authority="Dryand." authorityName="Dryand." box="[1109,1422,1627,1653]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="6" pageNumber="132" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF900C1DFB805768FA8AF9EC" box="[1109,1317,1627,1653]" italics="true" pageId="6" pageNumber="132">Heritiera littoralis</emphasis>
Dryand.
</taxonomicName>
”, suggesting that
<taxonomicName id="4C3D4D74FF900C1DFE92574CFE4AF900" authorityName="Tsuji &amp; Yoshitake" authorityYear="2025" baseAuthorityName="Heller" baseAuthorityYear="1913" box="[327,485,1663,1689]" class="Insecta" family="Curculionidae" genus="Tepperia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="132" phylum="Arthropoda" rank="species" species="palawanica">
<emphasis id="B949EAE5FF900C1DFE92574CFE4AF900" box="[327,485,1663,1689]" italics="true" pageId="6" pageNumber="132">T. palawanica</emphasis>
</taxonomicName>
larvae develop in fruits of
<taxonomicName id="4C3D4D74FF900C1DFCC1574CFC38F900" box="[788,919,1663,1689]" class="Magnoliopsida" family="Malvaceae" genus="Heritiera" kingdom="Plantae" order="Malvales" pageId="6" pageNumber="132" phylum="Tracheophyta" rank="species" species="littoralis">
<emphasis id="B949EAE5FF900C1DFCC1574CFC38F900" box="[788,919,1663,1689]" italics="true" pageId="6" pageNumber="132">H. littoralis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

101
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@ -1,64 +1,65 @@
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: pl. 12, fig. 9).
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–B
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. On sutural ramp, fine adis, IP, adis and on siphonal canal, fine ADP, ads, MP, ms and ABP.
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FIGURE 3. A

101
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<figureCitation id="13082A0CFFD9FF962785D7F2FF15F947" box="[151,222,1685,1712]" captionStart="FIGURE 7" captionStartId="17.[151,250,1821,1845]" captionTargetBox="[151,1435,182,1797]" captionTargetId="figure-24@17.[151,1435,181,1797]" captionTargetPageId="17" captionText="FIGURE 7. Cord diagrams and apertural features of Aspellinae (Favartia)." figureDoi="http://doi.org/10.5281/zenodo.15215752" httpUri="https://zenodo.org/record/15215752/files/figure.png" pageId="21" pageNumber="22">Figs 7</figureCitation>
,
<figureCitation id="13082A0CFFD9FF9627F8D7F2FEF7F958" box="[234,316,1685,1711]" captionStart="FIGURE 11" captionStartId="22.[151,250,1083,1107]" captionTargetBox="[151,1436,181,1060]" captionTargetId="figure-32@22.[151,1436,181,1060]" captionTargetPageId="22" captionText="FIGURE 11. Favartia kovacsi sp. nov. A1–A3. Holotype, NHMW 1863/0015/0664a, Niederleis (Austria). B1–B3. paratype, NHMW 1863/0015/0664b, Niederleis (Austria). C1–C3. paratype, NHMW 1863/0015/0664c, Niederleis (Austria). D1–D3. paratype, NHMW 1863/0015/0664d, Niederleis (Austria)." pageId="21" pageNumber="22">11A–D</figureCitation>
<figureCitation id="13082A0CFFD9FF9627F8D7F2FEF7F958" box="[234,316,1685,1711]" captionStart="FIGURE 11" captionStartId="22.[151,250,1083,1107]" captionTargetBox="[151,1436,181,1060]" captionTargetId="figure-32@22.[151,1436,181,1060]" captionTargetPageId="22" captionText="FIGURE 11. Favartia kovacsi sp. nov. A1–A3. Holotype, NHMW 1863/0015/0664a, Niederleis (Austria). B1–B3. paratype, NHMW 1863/0015/0664b, Niederleis (Austria). C1–C3. paratype, NHMW 1863/0015/0664c, Niederleis (Austria). D1–D3. paratype, NHMW 1863/0015/0664d, Niederleis (Austria)." figureDoi="http://doi.org/10.5281/zenodo.15215760" httpUri="https://zenodo.org/record/15215760/files/figure.png" pageId="21" pageNumber="22">11A–D</figureCitation>
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@ -165,7 +166,7 @@ Baden Formation.
Middle Miocene, middle Badenian (Langhian).
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<emphasis id="B947EA9BFFDAFF952785D55CFEEAFBA3" bold="true" box="[151,289,1083,1108]" pageId="22" pageNumber="23">FIGURE 11.</emphasis>
<taxonomicName id="4C334D0AFFDAFF952638D55BFE13FBA3" authorityName="Harzhauser &amp; Landau &amp; Merle" authorityYear="2025" box="[298,472,1084,1108]" class="Gastropoda" family="Muricidae" genus="Favartia" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="kovacsi" status="sp. nov.">
@ -267,13 +268,13 @@ in their much broader shells and larger size.
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differs from typical specimens in the presence of weak s1 and s2 (
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<figureCitation id="13082A0CFFDBFF9423CAD1CEFAC8FF46" box="[1240,1283,151,183]" captionStart="FIGURE 1" captionStartId="7.[151,250,1747,1771]" captionTargetBox="[151,1435,181,1723]" captionTargetId="figure-24@7.[151,1435,181,1723]" captionTargetPageId="7" captionText="FIGURE 1. Cord diagrams and apertural features of Aspellinae (Aspella, Attiliosa, Dermomurex)." figureDoi="http://doi.org/10.5281/zenodo.15215740" httpUri="https://zenodo.org/record/15215740/files/figure.png" pageId="23" pageNumber="24">
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–D
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<figureCitation id="13082A0CFFDBFF942211D1CEFAC0FF40" box="[1283,1291,169,183]" captionStart="FIGURE 3" captionStartId="10.[151,250,782,806]" captionTargetBox="[151,1436,181,759]" captionTargetId="figure-32@10.[151,1436,181,759]" captionTargetPageId="10" captionText="FIGURE 3. A1–A2. Attiliosa evae (Kovács &amp; Vicián, 2024). holotype, HNHM PAL 2024.8.1., Letkés (Hungary).B1–B2. Attiliosa juhaszi Kovács &amp; Vicián, 2024. holotype, HNHM PAL 2024.1.1., Letkés (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.15215744" httpUri="https://zenodo.org/record/15215744/files/figure.png" pageId="23" pageNumber="24">3</figureCitation>
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. This might represent some variability in secondary sculpture.

103
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<figureCitation id="13082A0CFFE9FFA62785D5E0FF27FB56" box="[151,236,1159,1185]" captionStart="FIGURE 21" captionStartId="38.[151,250,1432,1456]" captionTargetBox="[151,1435,273,1409]" captionTargetId="figure-66@38.[151,1436,273,1409]" captionTargetPageId="38" captionText="FIGURE 21. Cord diagrams and apertural features of Aspellinae (Pazinotus, Pygmaepterys, Tripterotyphis)." pageId="37" pageNumber="38">Figs 21</figureCitation>
<figureCitation id="13082A0CFFE9FFA62785D5E0FF27FB56" box="[151,236,1159,1185]" captionStart="FIGURE 21" captionStartId="38.[151,250,1432,1456]" captionTargetBox="[151,1435,273,1409]" captionTargetId="figure-66@38.[151,1436,273,1409]" captionTargetPageId="38" captionText="FIGURE 21. Cord diagrams and apertural features of Aspellinae (Pazinotus, Pygmaepterys, Tripterotyphis)." figureDoi="http://doi.org/10.5281/zenodo.15215786" httpUri="https://zenodo.org/record/15215786/files/figure.png" pageId="37" pageNumber="38">Figs 21</figureCitation>
,
<figureCitation id="13082A0CFFE9FFA627EAD5E0FE83FB56" box="[248,328,1159,1185]" captionStart="FIGURE 22" captionStartId="39.[151,250,1173,1197]" captionTargetBox="[151,1436,181,1149]" captionTargetId="figure-24@39.[151,1436,181,1149]" captionTargetPageId="39" captionText="FIGURE 22. Pazinotus pulcher sp. nov. A1–A3. Holotype, NHMW 1866/0001/0841, Niederleis (Austria). B1–B3. NHMW 1866/0001/0841a, paratype, Niederleis (Austria). C1–C3. NHMW 1866/0001/0841b, paratype, Niederleis (Austria). D1–D2. NHMW 1866/0001/0841c, paratype, Niederleis (Austria). E. NHMW 1866/0001/0841d, paratype, Niederleis (Austria)." pageId="37" pageNumber="38">22A–E</figureCitation>
<figureCitation id="13082A0CFFE9FFA627EAD5E0FE83FB56" box="[248,328,1159,1185]" captionStart="FIGURE 22" captionStartId="39.[151,250,1173,1197]" captionTargetBox="[151,1436,181,1149]" captionTargetId="figure-24@39.[151,1436,181,1149]" captionTargetPageId="39" captionText="FIGURE 22. Pazinotus pulcher sp. nov. A1–A3. Holotype, NHMW 1866/0001/0841, Niederleis (Austria). B1–B3. NHMW 1866/0001/0841a, paratype, Niederleis (Austria). C1–C3. NHMW 1866/0001/0841b, paratype, Niederleis (Austria). D1–D2. NHMW 1866/0001/0841c, paratype, Niederleis (Austria). E. NHMW 1866/0001/0841d, paratype, Niederleis (Austria)." figureDoi="http://doi.org/10.5281/zenodo.15216220" httpUri="https://zenodo.org/record/15216220/files/figure.png" pageId="37" pageNumber="38">22A–E</figureCitation>
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@ -194,7 +195,7 @@ Small, delicate, fusiform shell with slightly scalate spire and seven narrow, ra
Small, delicate, fusiform shell with slightly scalate spire; apical angle ~50°. Protoconch high conical of two smooth, moderately convex whorls. Teleoconch of up to four whorls. Suture strongly incised, linear. Early teleoconch whorls with shallow subsutural ramp, narrowly rounded shoulder, subcylindrical periphery, constricted below. Axial sculpture of seven narrow, raised, foliaceous varices. Spiral sculpture of weak P1 and P2; P1 forming short, open, adapically directed spines along shoulder. Last whorl attaining ~65% of total height; with shallow subsutural ramp, rounded shoulder subcylindrical periphery, moderately constricted below, high base. Axials sculpture of seven narrow, foliaceous varices, strictly aligned axially. Spiral sculpture weak, most prominent on varices; IP weak, P1–P4 and ADP forming low, broad cords. Subobsolete P5, P6 and secondary cords s1, s2 and s3. P1 forming short, adapically directed spines. Fasciole weakly swollen, delimiting narrow groove. Aperture narrowly ovate. Outer lip thickened with weak D1–D5. Anal canal wide, indistinct. Siphonal canal moderately long, narrow, open, slightly bent to the left, dorsally recurved at tip. Columella weakly excavated, smooth, twisted at siphonal canal. Columellar callus forming narrow, sharply delimited rim, erect at siphonal canal, adherent in parietal area.
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<emphasis id="B947EA9BFFEAFFA52785D4FFFEEBFA47" bold="true" box="[151,288,1432,1456]" pageId="38" pageNumber="39">FIGURE 21.</emphasis>
Cord diagrams and apertural features of
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<emphasis id="B947EA9BFFEBFFA42785D5F2FEE8FB5A" bold="true" box="[151,291,1173,1197]" pageId="39" pageNumber="40">FIGURE 22.</emphasis>
<taxonomicName id="4C334D0AFFEBFFA4263FD5F1FE2CFB59" authorityName="Harzhauser &amp; Landau &amp; Merle" authorityYear="2025" box="[301,487,1174,1198]" class="Gastropoda" family="Muricidae" genus="Pazinotus" kingdom="Animalia" order="Neogastropoda" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pulcher" status="sp. nov.">

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(
<figureCitation id="130C27CCA070FF9FC0A68E7BFB02C1E4" box="[159,277,619,645]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="3" pageNumber="374">Fig. 1E–H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A070FF9FC0AE8EA3FB91C03C" pageId="3" pageNumber="374" type="materials_examined">
<paragraph id="8B883B49A070FF9FC0AE8EA3F999C1AC" blockId="3.[151,1437,690,2013]" box="[151,910,690,717]" pageId="3" pageNumber="374">
<materialsCitation id="3B5F3114A070FF9FC0AE8EA3F99EC1AC" box="[151,905,690,717]" country="India" latitude="24.3404" location="Churachandpur" longLatPrecision="50" longitude="93.696" pageId="3" pageNumber="374" specimenCount="1" stateProvince="Manipur" typeStatus="holotype">
<emphasis id="B943E75BA070FF9FC0AE8EA3FB21C1AD" bold="true" box="[151,310,690,717]" pageId="3" pageNumber="374">
<typeStatus id="548C85EBA070FF9FC0AE8EA3FAC5C1AC" box="[151,210,691,717]" pageId="3" pageNumber="374">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3207BD9A070FF9FC1058EA3FB6FC1AC" box="[316,376,691,717]" name="India" pageId="3" pageNumber="374">India</collectingCountry>
,
<collectingRegion id="49F3F5ABA070FF9FC1BD8EA3FBF1C1AC" box="[388,486,691,717]" country="India" name="Manipur" pageId="3" pageNumber="374">Manipur</collectingRegion>
,
<location id="8EE86D92A070FF9FC1C88EA3F888C1AC" LSID="urn:lsid:plazi:treatment:039E8A5FA070FF98C0AE8E57FB2EC190:8EE86D92A070FF9FC1C88EA3F888C1AC" box="[497,671,691,717]" country="India" latitude="24.3404" longLatPrecision="50" longitude="93.696" name="Churachandpur" pageId="3" pageNumber="374" stateProvince="Manipur">Churachandpur</location>
,
<geoCoordinate id="EE035D8EA070FF9FC2928EA3F90BC1AC" box="[683,796,691,717]" degrees="24.3404" direction="north" orientation="latitude" pageId="3" pageNumber="374" precision="5" value="24.3404">24.3404N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA070FF9FC3118EA3F99EC1AC" box="[808,905,691,717]" degrees="93.696" direction="east" orientation="longitude" pageId="3" pageNumber="374" precision="55" value="93.696">93.696E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8B883B49A070FF9FC0FE8EC7FB91C03C" blockId="3.[151,1437,690,2013]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8EC7FB62C191" bold="true" box="[199,373,726,753]" pageId="3" pageNumber="374">Type material.</emphasis>
<materialsCitation id="3B5F3114A070FF9FC1458EC7FB91C03C" collectingDate="1993-05-19" collectionCode="NZCI" collectorName="M. Sil &amp; The" country="India" latitude="24.3404" location="Circuit House" longLatPrecision="50" longitude="93.696" municipality="Churachandpur" pageId="3" pageNumber="374" specimenCount="1" specimenCount-male="1" stateProvince="Manipur" typeStatus="holotype">
<typeStatus id="548C85EBA070FF9FC1458EC7FBF0C190" box="[380,487,727,753]" pageId="3" pageNumber="374" type="holotype">Holotype</typeStatus>
,
<emphasis id="B943E75BA070FF9FC1CA8EC8F81FC195" box="[499,520,728,756]" italics="true" pageId="3" pageNumber="374">
<specimenCount id="9D31F0C0A070FF9FC1CA8EC8F81FC195" box="[499,520,728,756]" count="1" pageId="3" pageNumber="374" type="male">♂</specimenCount>
</emphasis>
: “
<collectingCountry id="F3207BD9A070FF9FC2188EC7F848C190" box="[545,607,727,753]" name="India" pageId="3" pageNumber="374">India</collectingCountry>
,
<collectingRegion id="49F3F5ABA070FF9FC2528EC7F8DAC190" box="[619,717,727,753]" country="India" name="Manipur" pageId="3" pageNumber="374">Manipur</collectingRegion>
,
<collectingMunicipality id="6BECA133A070FF9FC2E08EC7F990C190" box="[729,903,727,753]" pageId="3" pageNumber="374">Churachandpur</collectingMunicipality>
,
<location id="8EE86D92A070FF9FC3AB8EC7FE26C190" LSID="urn:lsid:plazi:treatment:039E8A5FA070FF98C0AE8E57FB2EC190:8EE86D92A070FF9FC3AB8EC7FE26C190" box="[914,1073,727,753]" country="India" latitude="24.3404" longLatPrecision="50" longitude="93.696" municipality="Churachandpur" name="Circuit House" pageId="3" pageNumber="374" stateProvince="Manipur">Circuit House</location>
,
<geoCoordinate id="EE035D8EA070FF9FC4048EC7FEB8C190" box="[1085,1199,727,753]" degrees="24.3404" direction="north" orientation="latitude" pageId="3" pageNumber="374" precision="5" value="24.3404">24.3404N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA070FF9FC4828EC7FF0CC190" box="[1211,1307,727,753]" degrees="93.696" direction="east" orientation="longitude" pageId="3" pageNumber="374" precision="55" value="93.696">93.696E</geoCoordinate>
,
<date id="FF891D89A070FF9FC51E8EC7FF80C190" box="[1319,1431,727,753]" pageId="3" pageNumber="374" value="1993-05-19">
<collectingDate id="EFCDE461A070FF9FC51E8EC7FF80C190" box="[1319,1431,727,753]" pageId="3" pageNumber="374" value="1993-05-19">19.v.1993</collectingDate>
</date>
, leg.
<collectorName id="26C25E9FA070FF9FC0FE8EEBFB1BC074" box="[199,268,763,789]" pageId="3" pageNumber="374">M. Sil</collectorName>
and Party” (
<collectionCode id="ED26A38CA070FF9FC1AE8EEBFBC8C074" box="[407,479,763,789]" pageId="3" pageNumber="374">NZCI</collectionCode>
).
<collectorName id="26C25E9FA070FF9FC1C98EEBF80BC074" box="[496,540,763,789]" pageId="3" pageNumber="374">The</collectorName>
<typeStatus id="548C85EBA070FF9FC21B8EEBF893C074" box="[546,644,763,789]" pageId="3" pageNumber="374" type="holotype">holotype</typeStatus>
specimen bears additionally a red printed label with the following text: “
<taxonomicName id="4C3740CAA070FF9FC09D8F0FF816C058" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[164,513,799,825]" class="Insecta" family="Scarabaeidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="churachandpurensis" status="sp. nov.">
<emphasis id="B943E75BA070FF9FC09D8F0FF816C058" box="[164,513,799,825]" italics="true" pageId="3" pageNumber="374">Neoserica churachandpurensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA070FF9FC2318F0FF874C058" bold="true" box="[520,611,799,825]" pageId="3" pageNumber="374">
<taxonomicNameLabel id="A2705A20A070FF9FC2318F0FF874C058" box="[520,611,799,825]" pageId="3" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="548C85EBA070FF9FC2488F0FF91FC058" box="[625,776,799,825]" pageId="3" pageNumber="374" type="holotype">HOLOTYPE</typeStatus>
, Devanshu Gupta, Debika Bhunia, Dirk Ahrens &amp; Kailash Chandra, Det. 2024”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A070FF98C0FE8F76FB82C294" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374" type="description">
<paragraph id="8B883B49A070FF9FC0FE8F76FE61C0E0" blockId="3.[151,1437,690,2013]" box="[199,1142,870,897]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8F76FBF2C0E0" bold="true" box="[199,485,870,897]" pageId="3" pageNumber="374">
Description of
<typeStatus id="548C85EBA070FF9FC14E8F77FBF7C0E0" box="[375,480,871,897]" pageId="3" pageNumber="374" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CCF96ACA070FF9FC2738F77F8B5C0E0" box="[586,674,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="3" pageNumber="374" unit="mm" value="6.4">6.4 mm</quantity>
, length of elytra:
<quantity id="4CCF96ACA070FF9FC35E8F77F9A8C0E0" box="[871,959,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="374" unit="mm" value="4.7">4.7 mm</quantity>
, width:
<quantity id="4CCF96ACA070FF9FC4208F77FE65C0E0" box="[1049,1138,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="374" unit="mm" value="4.2">4.2 mm</quantity>
.
</paragraph>
<paragraph id="8B883B49A070FF9FC0FE8F9BFBE5C0A8" blockId="3.[151,1437,690,2013]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8F9BFB1CC0C4" bold="true" box="[199,267,907,933]" pageId="3" pageNumber="374">Body.</emphasis>
Shape oval, moderately convex; unicolored light reddish brown, entire dorsal surface shiny, glabrous except for a few setae on head.
</paragraph>
<paragraph id="8B883B49A070FF9FC0FE8FC3FAEBC650" blockId="3.[151,1437,690,2013]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8FC3FB19C08C" bold="true" box="[199,270,979,1005]" pageId="3" pageNumber="374">Head.</emphasis>
Labroclypeus transverse, widest at base, lateral margins strongly curved and narrowed anteriorly, clearly separated from ocular canthus by an obtuse angle, anterior angles strongly rounded, anterior margin shallowly emarginate medially, margins slightly reflexed; surface strongly convex, finely and densely punctate, with numerous, short setae. Frontoclypeal suture weakly incised, slightly curved, smooth area in front of eye about twice as wide as long, smooth; ocular canthus short and broad, finely and densely punctate, with a short terminal seta; frons finely and sparsely punctate, with a few erect, fine setae beside eyes and immediately behind frontoclypeal suture, punctures with setae somewhat larger than glabrous ones. Eyes moderately large, ratio ocular diameter/interocular distance: 0.76. Antenna with ten antennomeres, yellow, club in male with five antennomeres, as long as remaining antennomeres combined and straight, first joint of club a little shorter than club. Mentum anteriorly convexly elevated.
</paragraph>
<paragraph id="8B883B49A070FF9FC0FE892BF8B6C684" blockId="3.[151,1437,690,2013]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE892BFB57C634" bold="true" box="[199,320,1339,1365]" pageId="3" pageNumber="374">Pronotum</emphasis>
moderately wide, widest at base, lateral margins from base to middle straight, weakly narrowed, convexly curved in middle, and slightly curved and convergent in anterior half, anterior angles slightly produced and sharp, posterior angles rectangular, marginal line of anterior margin robust; surface finely and densely punctate, with minute setae in punctures, otherwise glabrous; anterior and lateral margins with sparse, long setae. Scutellum broadly triangular, punctation as on pronotum.
</paragraph>
<paragraph id="8B883B49A070FF9FC0FE89FFF823C514" blockId="3.[151,1437,690,2013]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE89FFFB04C568" bold="true" box="[199,275,1519,1545]" pageId="3" pageNumber="374">Elytra</emphasis>
widest at posterior third, striae finely impressed and finely punctate, intervals flat, sparsely and finely punctate, humerus near epipleural edge with a single, long seta, otherwise glabrous; apical elytral margin membranous, with wide rim of fine microtrichomes; robust epipleural edge ending at external apical angle of elytra; epipleura with long and sparse setae.
</paragraph>
<paragraph id="8B883B49A070FF9FC0FE8A6FF940C448" blockId="3.[151,1437,690,2013]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8A6FFB96C5F9" bold="true" box="[199,385,1662,1689]" pageId="3" pageNumber="374">Ventral surface</emphasis>
strongly, moderately densely punctate, with microscopic to fine setae, disc of metasternum densely and finely punctate, with long and dense, erect setae. Mesosternum between mesocoxae as wide as mesofemur. Abdominal sternites between fine, dense punctures with a transverse row of large punctures each bearing a robust seta. Ratio of metepisternum/metacoxa length: 1/1.44. Pygidium strongly convex, shiny, strong and irregularly densely punctate, apically with fine, long setae.
</paragraph>
<paragraph id="8B883B49A070FF98C0FE8B23F891C2CC" blockId="3.[151,1437,690,2013]" lastBlockId="4.[151,1437,151,753]" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8B23FAE8C42C" bold="true" box="[199,255,1843,1869]" pageId="3" pageNumber="374">Legs</emphasis>
short, meso- and metafemur moderately narrow, with two longitudinal rows of setae; metafemur wide, anterior margin sharply carinate, without adjacent crenulated line, fine, very sparsely punctate and glabrous, posterior margin straight and glabrous, neither ventrally nor dorsally serrated, posterior ventral margin widened at apex. Metatibia short and moderately wide, slightly narrowed towards base, subparallel in apical half, ratio width/ length: 1/2.85; dorsal margin sharply carinate, with two groups of spines, basal one at one third, apical one at three quarters of metatibial length, basally only with two additional, smaller bristles and with a very short, crenulated line; lateral face longitudinally convex, finely and sparsely punctate, completely smooth; ventral margin with four to five robust, equidistant spines; internal face smooth and glabrous, moderately concavely truncate interiorly near tarsal articulation. Metatarsi dorsally without punctures, ventrally with a serrated ridge and sparsely, shortly setose, metatarsomere 1 slightly shorter than following two tarsomeres combined and a quarter of its length longer than dorsal terminal spine, terminal spines of metatarsi short and strong. Protibia short and bidentate, distal tooth simple and moderately pointed, narrow and directed forward; protarsomeres narrow, ventrally normally setose, anterior claws symmetrical and normally developed.
</paragraph>
<paragraph id="8B883B49A077FF98C0FE8DA6F899C2B0" blockId="4.[151,1437,151,753]" box="[199,654,438,465]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8DA6FB2DC2B1" bold="true" box="[199,314,438,464]" pageId="4" pageNumber="375">Aedeagus</emphasis>
:
<figureCitation id="130C27CCA077FF98C1708DA7FBD6C2B0" box="[329,449,439,465]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="4" pageNumber="375">Fig. 1E–G</figureCitation>
. Habitus:
<figureCitation id="130C27CCA077FF98C20B8DA7F890C2B0" box="[562,647,439,465]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="4" pageNumber="375">Fig. 1H</figureCitation>
.
</paragraph>
<paragraph id="8B883B49A077FF98C0FE8DCBFB82C294" blockId="4.[151,1437,151,753]" box="[199,405,475,501]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8DCBFB09C294" bold="true" box="[199,286,475,501]" pageId="4" pageNumber="375">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A077FF98C0FE8DEEF95FC178" box="[199,840,510,537]" pageId="4" pageNumber="375" type="distribution">
<paragraph id="8B883B49A077FF98C0FE8DEEF95FC178" blockId="4.[151,1437,151,753]" box="[199,840,510,537]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8DEEFB76C179" bold="true" box="[199,353,510,536]" pageId="4" pageNumber="375">Distribution.</emphasis>
Species known only from the
<typeStatus id="548C85EBA077FF98C28E8DEFF8F0C178" box="[695,743,511,537]" pageId="4" pageNumber="375">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A077FF98C0FE8E32F946C1C8" pageId="4" pageNumber="375" type="diagnosis">
<paragraph id="8B883B49A077FF98C0FE8E32F946C1C8" blockId="4.[151,1437,151,753]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8E32FB55C15D" bold="true" box="[199,322,546,572]" pageId="4" pageNumber="375">Diagnosis.</emphasis>
<taxonomicName id="4C3740CAA077FF98C1778E33F8A7C15C" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[334,688,547,573]" class="Insecta" family="Scarabaeidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="churachandpurensis" status="sp. nov.">
<emphasis id="B943E75BA077FF98C1778E33F8A7C15C" box="[334,688,547,573]" italics="true" pageId="4" pageNumber="375">Neoserica churachandpurensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA077FF98C2848E33F90BC15C" bold="true" box="[701,796,547,573]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2705A20A077FF98C2848E33F90BC15C" box="[701,796,547,573]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
exhibits remarkable external and genital similarity to
<taxonomicName id="4C3740CAA077FF98C0AE8E57F83BC100" authority="(Frey, 1962)" baseAuthorityName="Frey" baseAuthorityYear="1962" box="[151,556,583,609]" class="Insecta" family="Scarabaeidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B943E75BA077FF98C0AE8E57FB86C101" box="[151,401,583,608]" italics="true" pageId="4" pageNumber="375">Neoserica assamensis</emphasis>
(Frey, 1962)
</taxonomicName>
. Both species can be distinguished by subtle differences in their parameres:
<taxonomicName id="4C3740CAA077FF98C0AE8E7BFBE3C1E4" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[151,500,619,645]" class="Insecta" family="Scarabaeidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="churachandpurensis" status="sp. nov.">
<emphasis id="B943E75BA077FF98C0AE8E7BFBE3C1E4" box="[151,500,619,645]" italics="true" pageId="4" pageNumber="375">Neoserica churachandpurensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA077FF98C1C28E7BF842C1E4" bold="true" box="[507,597,619,645]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2705A20A077FF98C1C28E7BF842C1E4" box="[507,597,619,645]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
possesses a grooved right paramere with a notch, whereas the paramere of
<taxonomicName id="4C3740CAA077FF98C0AE8E9FFB21C1C9" baseAuthorityName="Frey" baseAuthorityYear="1962" box="[151,310,655,680]" class="Insecta" family="Scarabaeidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="B943E75BA077FF98C0AE8E9FFB21C1C9" box="[151,310,655,680]" italics="true" pageId="4" pageNumber="375">N. assamensis</emphasis>
</taxonomicName>
is straight (
<bibRefCitation id="EFA646B8A077FF98C1808E9FF858C1C8" author="Ahrens, D." box="[441,591,655,681]" pageId="4" pageNumber="375" refId="ref9581" refString="Ahrens, D. (2004) Monographie der Sericini des Himalaya (Coleoptera, Scarabaeidae). Dissertation. de, Verlag im Internet GmbH, Berlin, 534 pp." type="book" year="2004">Ahrens 2004</bibRefCitation>
; figs 229–231, p. 415).
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A077FF98C0FE8EA3FB2EC190" pageId="4" pageNumber="375" type="etymology">
<paragraph id="8B883B49A077FF98C0FE8EA3FB2EC190" blockId="4.[151,1437,151,753]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8EA3FB59C1AC" bold="true" box="[199,334,691,717]" pageId="4" pageNumber="375">Etymology.</emphasis>
<taxonomicName id="4C3740CAA077FF98C16E8EA3F8A2C1AC" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[343,693,691,717]" class="Insecta" family="Scarabaeidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="churachandpurensis" status="sp. nov.">
<emphasis id="B943E75BA077FF98C16E8EA3F8A2C1AC" box="[343,693,691,717]" italics="true" pageId="4" pageNumber="375">Neoserica churachandpurensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA077FF98C2878EA3F90DC1AC" bold="true" box="[702,794,691,717]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2705A20A077FF98C2878EA3F90DC1AC" box="[702,794,691,717]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is named after its
<typeStatus id="548C85EBA077FF98C3CB8EA3FE35C1AC" box="[1010,1058,691,717]" pageId="4" pageNumber="375">type</typeStatus>
locality (adjective in nominative case singular).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="81FDB7009FA824789F30EA6C3262B60A" ID-DOI="10.11646/zootaxa.5613.2.11" ID-ISSN="1175-5326" ID-ZooBank="2972195A-6FD5-488A-8925-51D24677DFEC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1744667883477" checkinUser="plazi" docAuthor="Gupta, Devanshu, Bhunia, Debika, Ahrens, Dirk &amp; Chandra, Kailash" docDate="2025" docId="039E8A5FA072FF9FC0AE8DD6FB15C294" docLanguage="en" docName="zootaxa.5613.2.11.pdf" docOrigin="Zootaxa 5613 (2)" docSource="https://doi.org/10.11646/zootaxa.5613.2.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="374" masterDocId="FFA7F227A073FF9CC0398C10FA17C361" masterDocTitle="New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)" masterLastPageNumber="385" masterPageNumber="371" pageNumber="372" updateTime="1744670091748" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="83D5F7EC9AEAB098D76E34A78CA086E6">New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="C4B8D8FD34E8D474B07E5D8FB115EB2A">Bhunia, Debika</mods:namePart>
<mods:affiliation id="2F1F060F7543A26007CB99F586C4F40E">Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India</mods:affiliation>
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<mods:namePart id="CCE88B575F0E824414C883CE44A0F84E">Ahrens, Dirk</mods:namePart>
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<mods:namePart id="4FC199F9B41FA3E4481968AACCD0BE24">Chandra, Kailash</mods:namePart>
<mods:affiliation id="06B061490370140746D0CD74A738FB76">Flat No. 620, Tower Ferrara, Mahagun Mezzaria Sector- 78, Noida- 201301, Gautam Budha Nagar, U. P</mods:affiliation>
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<mods:date id="9A1AC9DBD5E28EB95C042512C4E9E3E5">2025</mods:date>
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<heading id="D0C08C25A072FF9DC0AE8DD6F9E1C280" bold="true" box="[151,1014,454,481]" fontSize="11" level="1" pageId="1" pageNumber="372" reason="1">
<emphasis id="B943E75BA072FF9DC0AE8DD6F9E1C280" bold="true" box="[151,1014,454,481]" pageId="1" pageNumber="372">
<taxonomicName id="4C3740CAA072FF9DC0AE8DD6F99CC281" ID-CoL="KTVW4" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[151,907,454,481]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="champhaiensis" status="sp. nov.">
<emphasis id="B943E75BA072FF9DC0AE8DD6FBDEC280" bold="true" box="[151,457,454,481]" italics="true" pageId="1" pageNumber="372">Maladera champhaiensis</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
,
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<subSubSection id="C32D68C2A072FF9DC0AE8DFBFB36C164" box="[151,289,491,517]" pageId="1" pageNumber="372" type="description">
<paragraph id="8B883B49A072FF9DC0AE8DFBFB36C164" blockId="1.[151,1014,454,517]" box="[151,289,491,517]" pageId="1" pageNumber="372">
(
<figureCitation id="130C27CCA072FF9DC0A68DFBFB0DC164" box="[159,282,491,517]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="1" pageNumber="372">Fig. 1A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A072FF9DC0AE8E3BFEE0C1CE" pageId="1" pageNumber="372" type="materials_examined">
<paragraph id="8B883B49A072FF9DC0AE8E3BF958C127" blockId="1.[151,1437,555,2025]" box="[151,847,555,582]" pageId="1" pageNumber="372">
<materialsCitation id="3B5F3114A072FF9DC0AE8E3BF95BC127" box="[151,844,555,582]" country="India" latitude="23.479" location="Champhai" longLatPrecision="74" longitude="93.325" pageId="1" pageNumber="372" specimenCount="1" stateProvince="Mizoram" typeStatus="holotype">
<emphasis id="B943E75BA072FF9DC0AE8E3BFB21C124" bold="true" box="[151,310,555,581]" pageId="1" pageNumber="372">
<typeStatus id="548C85EBA072FF9DC0AE8E3BFAC5C124" box="[151,210,555,581]" pageId="1" pageNumber="372">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3207BD9A072FF9DC1058E3BFB6DC124" box="[316,378,555,581]" name="India" pageId="1" pageNumber="372">India</collectingCountry>
:
<collectingRegion id="49F3F5ABA072FF9DC1BC8E3BFBF8C124" box="[389,495,555,581]" country="India" name="Mizoram" pageId="1" pageNumber="372">Mizoram</collectingRegion>
,
<location id="8EE86D92A072FF9DC1C08E3BF878C124" LSID="urn:lsid:plazi:treatment:039E8A5FA072FF9FC0AE8DD6FB15C294:8EE86D92A072FF9DC1C08E3BF878C124" box="[505,623,555,581]" country="India" latitude="23.479" longLatPrecision="74" longitude="93.325" name="Champhai" pageId="1" pageNumber="372" stateProvince="Mizoram">Champhai</location>
,
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,
<geoCoordinate id="EE035D8EA072FF9DC2D38E3BF95BC127" box="[746,844,555,582]" degrees="93.325" direction="east" orientation="longitude" pageId="1" pageNumber="372" precision="55" value="93.325">93.325E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8B883B49A072FF9DC0FE8E5EFEE0C1CE" blockId="1.[151,1437,555,2025]" pageId="1" pageNumber="372">
<emphasis id="B943E75BA072FF9DC0FE8E5EFB78C109" bold="true" box="[199,367,590,616]" pageId="1" pageNumber="372">Type material.</emphasis>
<materialsCitation id="3B5F3114A072FF9DC14D8E5EFEE0C1CE" collectingDate="1994-04-04" collectionCode="NZCI" collectorName="G. C. Sen &amp; The" country="India" latitude="23.479" location="Champhai" longLatPrecision="74" longitude="93.325" pageId="1" pageNumber="372" specimenCount="1" specimenCount-male="1" stateProvince="Mizoram" typeStatus="holotype">
<typeStatus id="548C85EBA072FF9DC14D8E5EFBCBC109" box="[372,476,590,616]" pageId="1" pageNumber="372" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D31F0C0A072FF9DC1DF8E5FFBE9C108" box="[486,510,591,617]" count="1" pageId="1" pageNumber="372" type="male">♂</specimenCount>
: “
<collectingCountry id="F3207BD9A072FF9DC2288E5EF85AC109" box="[529,589,590,616]" name="India" pageId="1" pageNumber="372">India</collectingCountry>
,
<collectingRegion id="49F3F5ABA072FF9DC2618E5EF8A9C109" box="[600,702,590,616]" country="India" name="Mizoram" pageId="1" pageNumber="372">Mizoram</collectingRegion>
,
<location id="8EE86D92A072FF9DC2F18E5EF92DC109" LSID="urn:lsid:plazi:treatment:039E8A5FA072FF9FC0AE8DD6FB15C294:8EE86D92A072FF9DC2F18E5EF92DC109" box="[712,826,590,616]" country="India" latitude="23.479" longLatPrecision="74" longitude="93.325" name="Champhai" pageId="1" pageNumber="372" stateProvince="Mizoram">Champhai</location>
,
<geoCoordinate id="EE035D8EA072FF9DC37D8E5EF9B2C109" box="[836,933,590,616]" degrees="23.479" direction="north" orientation="latitude" pageId="1" pageNumber="372" precision="55" value="23.479">23.479N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA072FF9DC3978E5EFE1AC108" box="[942,1037,590,617]" degrees="93.325" direction="east" orientation="longitude" pageId="1" pageNumber="372" precision="55" value="93.325">93.325E</geoCoordinate>
,
<date id="FF891D89A072FF9DC42F8E5EFE9CC109" box="[1046,1163,590,616]" pageId="1" pageNumber="372" value="1994-04-04">
<collectingDate id="EFCDE461A072FF9DC42F8E5EFE9CC109" box="[1046,1163,590,616]" pageId="1" pageNumber="372" value="1994-04-04">04.iv.1994</collectingDate>
</date>
, leg.
<collectorName id="26C25E9FA072FF9DC4FA8E5EFF32C109" box="[1219,1317,590,616]" pageId="1" pageNumber="372">G.C. Sen</collectorName>
and Party” (
<collectionCode id="ED26A38CA072FF9DC0A68E62FAF3C1ED" box="[159,228,626,652]" pageId="1" pageNumber="372">NZCI</collectionCode>
).
<collectorName id="26C25E9FA072FF9DC0CD8E61FB09C1EA" box="[244,286,625,651]" pageId="1" pageNumber="372">The</collectorName>
<typeStatus id="548C85EBA072FF9DC11D8E61FB94C1EA" box="[292,387,625,651]" pageId="1" pageNumber="372" type="holotype">holotype</typeStatus>
specimen bears additionally a red printed label with the following text: “
<taxonomicName id="4C3740CAA072FF9DC4B38E61FF8BC1EA" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[1162,1436,625,651]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="champhaiensis" status="sp. nov.">
<emphasis id="B943E75BA072FF9DC4B38E61FF8BC1EA" box="[1162,1436,625,651]" italics="true" pageId="1" pageNumber="372">Maladera champhaiensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA072FF9DC0AE8E85FAF9C1CE" bold="true" box="[151,238,661,687]" pageId="1" pageNumber="372">
<taxonomicNameLabel id="A2705A20A072FF9DC0AE8E85FAF9C1CE" box="[151,238,661,687]" pageId="1" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="548C85EBA072FF9DC0C58E85FB99C1CE" box="[252,398,661,687]" pageId="1" pageNumber="372" type="holotype">HOLOTYPE</typeStatus>
, Devanshu Gupta, Debika Bhunia, Dirk Ahrens &amp; Kailash Chandra, Det. 2024”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A072FF9FC0FE8EA8FB82C3B4" lastPageId="3" lastPageNumber="374" pageId="1" pageNumber="372" type="description">
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<emphasis id="B943E75BA072FF9DC0FE8EA8FBF2C1B3" bold="true" box="[199,485,696,722]" pageId="1" pageNumber="372">
Description of
<typeStatus id="548C85EBA072FF9DC14E8EA8FBF7C1B3" box="[375,480,696,722]" pageId="1" pageNumber="372" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CCF96ACA072FF9DC2738EA8F8A7C1B2" box="[586,688,696,723]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="1" pageNumber="372" unit="mm" value="10.1">10.1 mm</quantity>
, length of elytra:
<quantity id="4CCF96ACA072FF9DC34C8EA8F9DAC1B2" box="[885,973,696,723]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="372" unit="mm" value="7.8">7.8 mm</quantity>
, width: 6.0 mm.
</paragraph>
<paragraph id="8B883B49A072FF9DC0FE8ECBF926C079" blockId="1.[151,1437,555,2025]" pageId="1" pageNumber="372">
<emphasis id="B943E75BA072FF9DC0FE8ECBFB12C194" bold="true" box="[199,261,731,757]" pageId="1" pageNumber="372">Body</emphasis>
oval, uniformly dark brown, dorsal and ventral face dull. Head and anterior pronotum moderately shiny, except lateral setae of elytra and pronotum nearly glabrous.
</paragraph>
<paragraph id="8B883B49A072FF9DC0FE8F32F82CC734" blockId="1.[151,1437,555,2025]" pageId="1" pageNumber="372">
<emphasis id="B943E75BA072FF9DC0FE8F32FB19C05D" bold="true" box="[199,270,802,828]" pageId="1" pageNumber="372">Head.</emphasis>
Labroclypeus wide, trapezoidal, widest at base and shiny, lateral margins moderately convex and convergent anteriorly, producing an indistinct angle with ocular canthus, not incised before labrum; anterior angles strongly convex; anterior margin weakly emarginate medially; margins weakly reflexed; surface slightly convex, coarsely and densely punctate, with a few single and fine, erect setae on each side behind anterior margin. Frontoclypeal suture finely incised, weakly curved; ocular canthus short and wide, finely and moderately punctate, with a short terminal seta. Frons shiny, in posterior third dull, finely and moderately densely punctate, with a few single and short setae beside eyes. Eyes large, ratio ocular diameter/interocular distance: 0.9. Antenna with ten antennomeres, club with three antennomeres, as long as remaining antennomeres combined. Mentum convexly elevated, anteriorly slightly flattened.
</paragraph>
<paragraph id="8B883B49A072FF9DC0FE884EFB5AC664" blockId="1.[151,1437,555,2025]" pageId="1" pageNumber="372">
<emphasis id="B943E75BA072FF9DC0FE884EFB57C719" bold="true" box="[199,320,1118,1144]" pageId="1" pageNumber="372">Pronotum</emphasis>
widest at base, lateral margins evenly and convexly convergent anteriorly, anterior angles moderately sharp, distinctly produced, posterior angles blunt, moderately rounded at tip; anterior margin with complete marginal line, nearly straight; lateral and lateral anterior margin with long and fine setae; surface finely and densely punctate, with microscopic setae in punctures, otherwise glabrous. Scutellum wide, triangular, punctures fine and moderately dense, glabrous.
</paragraph>
<paragraph id="8B883B49A072FF9DC0FE891EF959C6F3" blockId="1.[151,1437,555,2025]" pageId="1" pageNumber="372">
<emphasis id="B943E75BA072FF9DC0FE891EFB04C649" bold="true" box="[199,275,1294,1320]" pageId="1" pageNumber="372">Elytra</emphasis>
wide, widest at middle, external apical angle strongly rounded, striae finely strongly impressed, finely and densely punctate, intervals weakly convex, finely and moderately densely punctate, with only microscopic setae in punctures; epipleural edge ending at external apical angle of elytra; epipleura with long and sparse setae; apical margin with a broad membranous rim of fine microtrichomes.
</paragraph>
<paragraph id="8B883B49A072FF9DC0FE898BFE26C507" blockId="1.[151,1437,555,2025]" pageId="1" pageNumber="372">
<emphasis id="B943E75BA072FF9DC0FE898BFB97C6D4" bold="true" box="[199,384,1435,1461]" pageId="1" pageNumber="372">Ventral surface</emphasis>
coarsely and densely punctate, with microscopic setae in punctures, with a few longer setae on mesosternum and metasternal plate. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.45. Metacoxa minutely punctate, punctures with minute setae. Abdominal sternites finely and moderately densely punctate, with a transverse row of coarse punctures each bearing a robust seta, ultimate sternite with dense and long setae. Pygidium dull, strongly convex, coarsely and moderately densely punctate, with long setae along the apical margin, otherwise with microscopic setae in punctures.
</paragraph>
<paragraph id="8B883B49A072FF9DC0FE8A7EFB0EC488" blockId="1.[151,1437,555,2025]" pageId="1" pageNumber="372">
<emphasis id="B943E75BA072FF9DC0FE8A7EFAE8C5E9" bold="true" box="[199,255,1646,1672]" pageId="1" pageNumber="372">Legs</emphasis>
wide, femora with two longitudinal rows of setae. Metafemur shiny, superficially punctate, anterior margin acute, with adjacent, straight and complete serrated line, anterior row of setae present but its setae short; posterior ventral margin almost straight, weakly widened in apical half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia moderately wide and short, widest at middle, ratio width/length: 1/2.3, sharply carinate dorsally; with two groups of spines, basal one shortly behind middle, apical one at four fifths of metatibial length, in basal third with a short, serrated line parallel to dorsal margin and 3–4 coarse punctures each bearing a fine seta. Tarsomeres impunctate dorsally, circular in cross section, with sparse, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate, teeth moderately large. All claws symmetrical, feebly curved and long, with normally developed basal tooth.
</paragraph>
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<paragraph id="8B883B49A071FF9EC0A18B64F86DC499" blockId="2.[151,1436,1908,2040]" pageId="2" pageNumber="373">
<emphasis id="B943E75BA071FF9EC0A18B64FB42C4ED" bold="true" box="[152,341,1908,1932]" pageId="2" pageNumber="373">FIGURE 1. A–D.</emphasis>
<taxonomicName id="4C3740CAA071FF9EC1628B64F9D7C4ED" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[347,960,1908,1932]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="champhaiensis" status="sp. nov.">
<emphasis id="B943E75BA071FF9EC1628B64F84BC4ED" box="[347,604,1908,1932]" italics="true" pageId="2" pageNumber="373">Maladera champhaiensis</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
<emphasis id="B943E75BA071FF9EC3FF8B64FE01C4ED" bold="true" box="[966,1046,1908,1932]" pageId="2" pageNumber="373">
<taxonomicNameLabel id="A2705A20A071FF9EC3FF8B64FE01C4ED" box="[966,1046,1908,1932]" pageId="2" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B943E75BA071FF9EC41D8B64FE4BC4ED" bold="true" box="[1060,1116,1908,1932]" pageId="2" pageNumber="373">E–H.</emphasis>
<taxonomicName id="4C3740CAA071FF9EC4588B65FBEEC4D1" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" class="Insecta" family="Scarabaeidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="churachandpurensis" status="sp. nov.">
<emphasis id="B943E75BA071FF9EC4588B65FF8BC4ED" box="[1121,1436,1908,1932]" italics="true" pageId="2" pageNumber="373">Neoserica churachandpurensis</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
<emphasis id="B943E75BA071FF9EC2398B88F845C4D1" bold="true" box="[512,594,1944,1968]" pageId="2" pageNumber="373">
<taxonomicNameLabel id="A2705A20A071FF9EC2398B88F845C4D1" box="[512,594,1944,1968]" pageId="2" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B943E75BA071FF9EC2598B88F899C4D1" bold="true" box="[608,654,1944,1968]" pageId="2" pageNumber="373">I–L.</emphasis>
<taxonomicName id="4C3740CAA071FF9EC2AC8B88FECAC4D1" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[661,1245,1944,1968]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="barasingha" status="sp. nov.">
<emphasis id="B943E75BA071FF9EC2AC8B88F964C4D1" box="[661,883,1944,1968]" italics="true" pageId="2" pageNumber="373">Maladera barasingha</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
<emphasis id="B943E75BA071FF9EC4DD8B88FF8BC4D1" bold="true" box="[1252,1436,1944,1968]" pageId="2" pageNumber="373">
<taxonomicNameLabel id="A2705A20A071FF9EC4DD8B88FF26C4D1" box="[1252,1329,1944,1968]" pageId="2" pageNumber="373" rank="species">sp. nov</taxonomicNameLabel>
.. (A, E, I)
</emphasis>
Aedeagus in lateral view (left);
<emphasis id="B943E75BA071FF9EC1EC8BACF830C4B5" bold="true" box="[469,551,1980,2004]" pageId="2" pageNumber="373">(B, F, J)</emphasis>
Aedeagus in dorsal view;
<emphasis id="B943E75BA071FF9EC3088BACF986C4B5" bold="true" box="[817,913,1980,2004]" pageId="2" pageNumber="373">(C, H, L)</emphasis>
Aedeagus in lateral view (right);
<emphasis id="B943E75BA071FF9EC4DB8BACFF55C4B5" bold="true" box="[1250,1346,1980,2004]" pageId="2" pageNumber="373">(D, H, L)</emphasis>
Habitus, dorsal view. Scale 0.5 mm; habitus not to scale.
</paragraph>
</caption>
<paragraph id="8B883B49A070FF9FC0FE8C86F880C3D0" blockId="3.[151,1436,150,501]" box="[199,663,150,177]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8C86FB2DC3D1" bold="true" box="[199,314,150,176]" pageId="3" pageNumber="374">Aedeagus</emphasis>
:
<figureCitation id="130C27CCA070FF9FC1708C87FBD5C3D0" box="[329,450,151,177]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="3" pageNumber="374">Fig. 1A–C</figureCitation>
.
<emphasis id="B943E75BA070FF9FC1F48C86F83BC3D1" bold="true" box="[461,556,150,176]" pageId="3" pageNumber="374">Habitus</emphasis>
:
<figureCitation id="130C27CCA070FF9FC2028C87F884C3D0" box="[571,659,151,177]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="3" pageNumber="374">Fig. 1D</figureCitation>
.
</paragraph>
<paragraph id="8B883B49A070FF9FC0FE8CABFB82C3B4" blockId="3.[151,1436,150,501]" box="[199,405,187,213]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8CABFB09C3B4" bold="true" box="[199,286,187,213]" pageId="3" pageNumber="374">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A070FF9FC0FE8CCEF998C398" box="[199,911,222,249]" pageId="3" pageNumber="374" type="distribution">
<paragraph id="8B883B49A070FF9FC0FE8CCEF998C398" blockId="3.[151,1436,150,501]" box="[199,911,222,249]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8CCEFB76C399" bold="true" box="[199,353,222,248]" pageId="3" pageNumber="374">Distribution.</emphasis>
The species is known only from the
<typeStatus id="548C85EBA070FF9FC2C78CCFF939C398" box="[766,814,223,249]" pageId="3" pageNumber="374">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A070FF9FC0FE8D12FAD3C2CC" pageId="3" pageNumber="374" type="diagnosis">
<paragraph id="8B883B49A070FF9FC0FE8D12FAD3C2CC" blockId="3.[151,1436,150,501]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8D12FB55C27D" bold="true" box="[199,322,258,284]" pageId="3" pageNumber="374">Diagnosis.</emphasis>
<taxonomicName id="4C3740CAA070FF9FC1738D13F870C27C" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[330,615,259,285]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="champhaiensis" status="sp. nov.">
<emphasis id="B943E75BA070FF9FC1738D13F870C27C" box="[330,615,259,285]" italics="true" pageId="3" pageNumber="374">Maladera champhaiensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA070FF9FC2498D13F8DBC27C" bold="true" box="[624,716,259,285]" pageId="3" pageNumber="374">
<taxonomicNameLabel id="A2705A20A070FF9FC2498D13F8DBC27C" box="[624,716,259,285]" pageId="3" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C3740CAA070FF9FC3748D13FAD8C220" authority="Sreedevi, Speer, Fabrizi &amp; Ahrens, 2018" authorityName="Sreedevi, Speer, Fabrizi &amp; Ahrens" authorityYear="2018" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="alloservitrita">
<emphasis id="B943E75BA070FF9FC3748D13FE11C27C" box="[845,1030,259,285]" italics="true" pageId="3" pageNumber="374">M. alloservitrita</emphasis>
<bibRefCitation id="EFA646B8A070FF9FC4368D13FAD8C220" author="Sreedevi, K. &amp; Speer, J. &amp; Fabrizi, S. &amp; Ahrens, D." pageId="3" pageNumber="374" pagination="97 - 128" refId="ref10602" refString="Sreedevi, K., Speer, J., Fabrizi, S. &amp; Ahrens, D. (2018) New species and records of Sericini scarab beetles from the Indian subcontinent (Coleoptera, Scarabaeidae). ZooKeys, 772, 97-128. https://doi.org/10.3897/zookeys.772.25320" type="journal article" year="2018">
Sreedevi, Speer, Fabrizi &amp; Ahrens,
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in its external and genital morphology. The new species differs from
<taxonomicName id="4C3740CAA070FF9FC3CC8D37FEB9C220" authorityName="Sreedevi, Speer, Fabrizi &amp; Ahrens" authorityYear="2018" box="[1013,1198,295,321]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="alloservitrita">
<emphasis id="B943E75BA070FF9FC3CC8D37FEB9C220" box="[1013,1198,295,321]" italics="true" pageId="3" pageNumber="374">M. alloservitrita</emphasis>
</taxonomicName>
by the angles of the dorsomedian process of phallobase and the shape of the parameres: the left paramere is not fused with the phallobase, the right one is short not being extended beyond the apex of the left paramere (
<bibRefCitation id="EFA646B8A070FF9FC4288D7FFEE0C2E8" author="Sreedevi, K. &amp; Speer, J. &amp; Fabrizi, S. &amp; Ahrens, D." box="[1041,1271,367,393]" pageId="3" pageNumber="374" pagination="97 - 128" refId="ref10602" refString="Sreedevi, K., Speer, J., Fabrizi, S. &amp; Ahrens, D. (2018) New species and records of Sericini scarab beetles from the Indian subcontinent (Coleoptera, Scarabaeidae). ZooKeys, 772, 97-128. https://doi.org/10.3897/zookeys.772.25320" type="journal article" year="2018">
Sreedevi
<emphasis id="B943E75BA070FF9FC4458D7FFEA2C2E8" box="[1148,1205,367,393]" italics="true" pageId="3" pageNumber="374">et al.</emphasis>
2018
</bibRefCitation>
; fig. 1 A–E, p. 99).
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A070FF9FC0FE8DA7FB15C294" pageId="3" pageNumber="374" type="etymology">
<paragraph id="8B883B49A070FF9FC0FE8DA7FB15C294" blockId="3.[151,1436,150,501]" pageId="3" pageNumber="374">
<emphasis id="B943E75BA070FF9FC0FE8DA7FB59C2B0" bold="true" box="[199,334,439,465]" pageId="3" pageNumber="374">Etymology.</emphasis>
<taxonomicName id="4C3740CAA070FF9FC16E8DA7F862C2B0" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[343,629,439,465]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="champhaiensis" status="sp. nov.">
<emphasis id="B943E75BA070FF9FC16E8DA7F862C2B0" box="[343,629,439,465]" italics="true" pageId="3" pageNumber="374">Maladera champhaiensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA070FF9FC2468DA7F8CCC2B0" bold="true" box="[639,731,439,465]" pageId="3" pageNumber="374">
<taxonomicNameLabel id="A2705A20A070FF9FC2468DA7F8CCC2B0" box="[639,731,439,465]" pageId="3" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is named after its
<typeStatus id="548C85EBA070FF9FC38F8DA7F9F1C2B0" box="[950,998,439,465]" pageId="3" pageNumber="374">type</typeStatus>
locality (adjective in nominative case singular).
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<taxonomicName id="4C3740CAA075FF9AC0AE880BF94DC755" ID-CoL="KTVWM" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[151,858,1050,1077]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="subansiriensis" status="sp. nov.">
<emphasis id="B943E75BA075FF9AC0AE880BFB8FC754" bold="true" box="[151,408,1050,1077]" italics="true" pageId="6" pageNumber="377">Serica subansiriensis</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
,
<taxonomicNameLabel id="A2705A20A075FF9AC35E880BF9D2C754" box="[871,965,1051,1077]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A075FF9AC0AE882FFB09C738" box="[151,286,1087,1113]" pageId="6" pageNumber="377" type="description">
<paragraph id="8B883B49A075FF9AC0AE882FFB09C738" blockId="6.[151,965,1050,1113]" box="[151,286,1087,1113]" pageId="6" pageNumber="377">
(
<figureCitation id="130C27CCA075FF9AC0A6882FFB02C738" box="[159,277,1087,1113]" captionStart="FIGURE 2" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[183,1413,188,1870]" captionTargetId="figure-22@7.[175,1413,179,1882]" captionTargetPageId="7" captionText="FIGURE 2. A–D. Maladera lumlaensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Serica subansiriensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="6" pageNumber="377">Fig. 2E–H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A075FF9AC0AE8890FB0DC64B" pageId="6" pageNumber="377" type="materials_examined">
<paragraph id="8B883B49A075FF9AC0AE8890FE05C7FA" blockId="6.[151,1437,1152,2043]" box="[151,1042,1152,1179]" pageId="6" pageNumber="377">
<materialsCitation id="3B5F3114A075FF9AC0AE8890FE19C7FA" box="[151,1038,1152,1179]" country="India" latitude="27.548" location="Pange" longLatPrecision="74" longitude="93.89" municipality="Subansiri" pageId="6" pageNumber="377" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<emphasis id="B943E75BA075FF9AC0AE8890FB21C7FB" bold="true" box="[151,310,1152,1178]" pageId="6" pageNumber="377">
<typeStatus id="548C85EBA075FF9AC0AE8890FAC5C7FB" box="[151,210,1152,1178]" pageId="6" pageNumber="377">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3207BD9A075FF9AC1058890FB6DC7FB" box="[316,378,1152,1178]" name="India" pageId="6" pageNumber="377">India</collectingCountry>
:
<collectingRegion id="49F3F5ABA075FF9AC1BA8890F84CC7FB" box="[387,603,1152,1178]" country="India" name="Arunachal Pradesh" pageId="6" pageNumber="377">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="6BECA133A075FF9AC25C8890F8C5C7FB" box="[613,722,1152,1178]" pageId="6" pageNumber="377">Subansiri</collectingMunicipality>
,
<location id="8EE86D92A075FF9AC2E48891F933C7FA" LSID="urn:lsid:plazi:treatment:039E8A5FA075FF94C0AE880BFACFC66C:8EE86D92A075FF9AC2E48891F933C7FA" box="[733,804,1153,1179]" country="India" latitude="27.548" longLatPrecision="74" longitude="93.89" municipality="Subansiri" name="Pange" pageId="6" pageNumber="377" stateProvince="Arunachal Pradesh">Pange</location>
,
<geoCoordinate id="EE035D8EA075FF9AC3098890F98CC7FA" box="[816,923,1152,1179]" degrees="27.548" direction="north" orientation="latitude" pageId="6" pageNumber="377" precision="55" value="27.548">27.548 N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA075FF9AC39F8890FE19C7FA" box="[934,1038,1152,1179]" degrees="93.890" direction="east" orientation="longitude" pageId="6" pageNumber="377" precision="55" value="93.89">93.890 E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8B883B49A075FF9AC0FE88B4FB0DC64B" blockId="6.[151,1437,1152,2043]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE88B4FB61C7DF" bold="true" box="[199,374,1188,1214]" pageId="6" pageNumber="377">Type material.</emphasis>
<materialsCitation id="3B5F3114A075FF9AC14788B4FB0DC64B" collectingDate="2017-06-17" collectionCode="R, NZCI" collectorName="K. Kushwaha" country="India" latitude="27.548" location="Pange" longLatPrecision="495" longitude="93.89" municipality="Subansiri" pageId="6" pageNumber="377" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<typeStatus id="548C85EBA075FF9AC14788B4FBFEC7DF" box="[382,489,1188,1214]" pageId="6" pageNumber="377" type="holotype">Holotype</typeStatus>
,
<emphasis id="B943E75BA075FF9AC1CC88B5F81DC7A0" box="[501,522,1189,1217]" italics="true" pageId="6" pageNumber="377">
<specimenCount id="9D31F0C0A075FF9AC1CC88B5F81DC7A0" box="[501,522,1189,1217]" count="1" pageId="6" pageNumber="377" type="male">♂</specimenCount>
</emphasis>
: “
<collectingCountry id="F3207BD9A075FF9AC21D88B4F873C7DF" box="[548,612,1188,1214]" name="India" pageId="6" pageNumber="377">India</collectingCountry>
:
<collectingRegion id="49F3F5ABA075FF9AC25488B4F951C7DF" box="[621,838,1188,1214]" country="India" name="Arunachal Pradesh" pageId="6" pageNumber="377">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="6BECA133A075FF9AC36888B4F9A9C7DF" box="[849,958,1188,1214]" pageId="6" pageNumber="377">Subansiri</collectingMunicipality>
,
<location id="8EE86D92A075FF9AC3F388B5FE06C7DE" LSID="urn:lsid:plazi:treatment:039E8A5FA075FF94C0AE880BFACFC66C:8EE86D92A075FF9AC3F388B5FE06C7DE" box="[970,1041,1189,1215]" country="India" latitude="27.548" longLatPrecision="495" longitude="93.89" municipality="Subansiri" name="Pange" pageId="6" pageNumber="377" stateProvince="Arunachal Pradesh">Pange</location>
,
<geoCoordinate id="EE035D8EA075FF9AC42488B4FE95C7DE" box="[1053,1154,1188,1215]" degrees="27.548" direction="north" orientation="latitude" pageId="6" pageNumber="377" precision="55" value="27.548">27.548N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA075FF9AC4B488B4FEF6C7DF" box="[1165,1249,1188,1214]" degrees="93.89" direction="east" orientation="longitude" pageId="6" pageNumber="377" precision="555" value="93.89">93.89E</geoCoordinate>
,
<date id="FF891D89A075FF9AC4D588B4FF71C7DF" box="[1260,1382,1188,1214]" pageId="6" pageNumber="377" value="2017-06-17">
<collectingDate id="EFCDE461A075FF9AC4D588B4FF71C7DF" box="[1260,1382,1188,1214]" pageId="6" pageNumber="377" value="2017-06-17">17.vi.2017</collectingDate>
</date>
, leg.
<collectionCode id="ED26A38CA075FF9AC0AE88D9FABAC782" box="[151,173,1225,1251]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="377">R</collectionCode>
.
<collectorName id="26C25E9FA075FF9AC08888D9FB58C783" box="[177,335,1224,1251]" pageId="6" pageNumber="377">K. Kushwaha</collectorName>
” (
<collectionCode id="ED26A38CA075FF9AC15388D8FBA5C783" box="[362,434,1224,1250]" pageId="6" pageNumber="377">NZCI</collectionCode>
). The
<typeStatus id="548C85EBA075FF9AC1C588D8F849C783" box="[508,606,1224,1250]" pageId="6" pageNumber="377" type="holotype">holotype</typeStatus>
specimen bears additionally a red printed label with the following text: “
<taxonomicName id="4C3740CAA075FF9AC09D88FCFB84C667" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[164,403,1260,1286]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="subansiriensis" status="sp. nov.">
<emphasis id="B943E75BA075FF9AC09D88FCFB84C667" box="[164,403,1260,1286]" italics="true" pageId="6" pageNumber="377">Serica subansiriensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA075FF9AC1A288FCFBE2C667" bold="true" box="[411,501,1260,1286]" pageId="6" pageNumber="377">
<taxonomicNameLabel id="A2705A20A075FF9AC1A288FCFBE2C667" box="[411,501,1260,1286]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="548C85EBA075FF9AC23D88FCF88CC667" box="[516,667,1260,1286]" pageId="6" pageNumber="377" type="holotype">HOLOTYPE</typeStatus>
, Devanshu Gupta, Debika Bhunia, Dirk Ahrens &amp; Kailash Chandra, Det. 2024”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A075FF94C0FE8924FB82C0A8" lastPageId="8" lastPageNumber="379" pageId="6" pageNumber="377" type="description">
<paragraph id="8B883B49A075FF9AC0FE8924FEB4C62E" blockId="6.[151,1437,1152,2043]" box="[199,1187,1332,1359]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8924F806C62F" bold="true" box="[199,529,1332,1358]" pageId="6" pageNumber="377">
Description of the
<typeStatus id="548C85EBA075FF9AC19A8924F81BC62F" box="[419,524,1332,1358]" pageId="6" pageNumber="377" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CCF96ACA075FF9AC24F8924F8D9C62E" box="[630,718,1332,1359]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="6" pageNumber="377" unit="mm" value="7.1">7.1 mm</quantity>
, length of elytra:
<quantity id="4CCF96ACA075FF9AC3AA8924F9FCC62E" box="[915,1003,1332,1359]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="377" unit="mm" value="5.3">5.3 mm</quantity>
, width:
<quantity id="4CCF96ACA075FF9AC47F8924FE88C62E" box="[1094,1183,1332,1359]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="377" unit="mm" value="3.7">3.7 mm</quantity>
.
</paragraph>
<paragraph id="8B883B49A075FF9AC0FE8948FB51C6F6" blockId="6.[151,1437,1152,2043]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8948FB12C613" bold="true" box="[199,261,1368,1394]" pageId="6" pageNumber="377">Body</emphasis>
oblong, dark reddish brown, antenna yellowish, legs reddish brown; except head dorsal surface dull, sparsely setose.
</paragraph>
<paragraph id="8B883B49A075FF9AC0FE89B0F9DFC443" blockId="6.[151,1437,1152,2043]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE89B0FB10C6DB" bold="true" box="[199,263,1440,1466]" pageId="6" pageNumber="377">Head</emphasis>
. Labroclypeus slightly wider than long, widest at middle, lateral margins convex and convergent anteriorly and posteriorly; anterior angles weakly rounded, anterior margin deeply emarginate medially; margins strongly reflexed; surface flat and shiny, finely and densely punctate, with shallow transverse wrinkles on basal part, with numerous long, erect setae; frontoclypeal suture distinctly elevated, weakly curved; smooth area anterior to eye large and convex, approximately 1.5 times as wide as long; ocular canthus short and narrow (1/4 of ocular diameter), smooth, with one short terminal seta. Frons completely shiny and impressed behind frontoclypeal suture, with fine and moderately dense but irregularly scattered punctures, with a few erect setae beside eyes and behind frontoclypeal suture. Eyes very large, ratio diameter/ interocular width: 0.95. Antenna yellowish, with ten antennomeres; antennomeres three to seven transverse and short; club with three antennomeres, 1.7 times as long as remaining antennomeres combined and weakly reflexed. Mentum weakly elevated, anteriorly flattened. Labrum transverse, short, moderately produced, moderately emarginate medially.
</paragraph>
<paragraph id="8B883B49A075FF9AC0FE8B3CF8B0C49B" blockId="6.[151,1437,1152,2043]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8B3CFB57C427" bold="true" box="[199,320,1836,1862]" pageId="6" pageNumber="377">Pronotum</emphasis>
transverse, widest shortly anterior to middle, lateral margins in basal half deeply concave, convex and convergent in anterior half; anterior angles weakly produced and rounded, posterior angles nearly sharp; anterior margin strongly convex and with a robust marginal line; surface moderately densely and finely punctate, a few punctures laterally with short and adpressed setae, otherwise only with minute setae in punctures; anterior and lateral borders sparsely setose; hypomeron not carinate at base. Scutellum slender and long, triangular, finely and densely punctate, at base impunctate, glabrous.
</paragraph>
<caption id="DF486BC1A074FF9BC0A18B65FB81C498" pageId="7" pageNumber="378" startId="7.[152,255,1909,1933]" targetBox="[183,1413,188,1870]" targetPageId="7" targetType="figure">
<paragraph id="8B883B49A074FF9BC0A18B65FB81C498" blockId="7.[151,1437,1909,2041]" pageId="7" pageNumber="378">
<emphasis id="B943E75BA074FF9BC0A18B65FB44C4EC" bold="true" box="[152,339,1909,1933]" pageId="7" pageNumber="378">FIGURE 2. A–D.</emphasis>
<taxonomicName id="4C3740CAA074FF9BC16E8B65F99CC4EC" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[343,907,1909,1933]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="lumlaensis" status="sp. nov.">
<emphasis id="B943E75BA074FF9BC16E8B65F83BC4EC" box="[343,556,1909,1933]" italics="true" pageId="7" pageNumber="378">Maladera lumlaensis</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
<emphasis id="B943E75BA074FF9BC3A98B65F9C8C4EC" bold="true" box="[912,991,1909,1933]" pageId="7" pageNumber="378">
<taxonomicNameLabel id="A2705A20A074FF9BC3A98B65F9C8C4EC" box="[912,991,1909,1933]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B943E75BA074FF9BC3D28B65FE34C4EC" bold="true" box="[1003,1059,1909,1933]" pageId="7" pageNumber="378">E–H.</emphasis>
<taxonomicName id="4C3740CAA074FF9BC4118B65FB43C4D0" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="subansiriensis" status="sp. nov.">
<emphasis id="B943E75BA074FF9BC4118B65FEE9C4EC" box="[1064,1278,1909,1933]" italics="true" pageId="7" pageNumber="378">Serica subansiriensis</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
<emphasis id="B943E75BA074FF9BC1608B89FBF2C4D0" bold="true" box="[345,485,1945,1969]" pageId="7" pageNumber="378">
<taxonomicNameLabel id="A2705A20A074FF9BC1608B89FBBAC4D0" box="[345,429,1945,1969]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
; I–L.
</emphasis>
<taxonomicName id="4C3740CAA074FF9BC1D38B89F9FDC4D0" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[490,1002,1945,1969]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="onam" status="sp. nov.">
<emphasis id="B943E75BA074FF9BC1D38B89F89CC4D0" box="[490,651,1945,1969]" italics="true" pageId="7" pageNumber="378">Maladera onam</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
<emphasis id="B943E75BA074FF9BC3D68B89FEB7C4D1" bold="true" box="[1007,1184,1944,1969]" pageId="7" pageNumber="378">
<taxonomicNameLabel id="A2705A20A074FF9BC3D68B89FE2CC4D0" box="[1007,1083,1945,1969]" pageId="7" pageNumber="378" rank="species">sp. nov</taxonomicNameLabel>
.. (A, E, I)
</emphasis>
Aedeagus in lateral view (left);
<emphasis id="B943E75BA074FF9BC0E18BACFB3AC4B5" bold="true" box="[216,301,1980,2005]" pageId="7" pageNumber="378">(B, F, J)</emphasis>
Aedeagus in dorsal view;
<emphasis id="B943E75BA074FF9BC2078BACF8B7C4B5" bold="true" box="[574,672,1980,2005]" pageId="7" pageNumber="378">(C, H, L)</emphasis>
Aedeagus in lateral view (right);
<emphasis id="B943E75BA074FF9BC3C18BACFE4CC4B5" bold="true" box="[1016,1115,1980,2005]" pageId="7" pageNumber="378">(D, H, L)</emphasis>
Habitus, dorsal view. Scale 0.5 mm; habitus not to scale.
</paragraph>
</caption>
<paragraph id="8B883B49A07BFF94C0FE8C87F8F8C2B0" blockId="8.[151,1437,151,1293]" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE8C87FB04C3D0" bold="true" box="[199,275,151,177]" pageId="8" pageNumber="379">Elytra</emphasis>
oblong, widest at middle, striae distinctly impressed, finely and densely punctate, intervals flat, with fine, moderately dense punctures concentrated along striae, intervals with sparsely scattered, fine, short, adpressed, white setae, with a few dark, impunctate spots; epipleural edge fine, ending at moderately curved external apical angle of elytra; epipleura densely setose, apical border chitinous, without microtrichomes (magnification 100×).
<emphasis id="B943E75BA07BFF94C57A8D13FAF9C221" bold="true" pageId="8" pageNumber="379">Ventral surface</emphasis>
dull, finely and not densely punctate, moderately densely setose, metacoxa glabrous, with a few long setae only laterally. Abdominal sternites finely and densely punctate, with a transverse row of coarse punctures, each bearing a short seta. Mesosternum between mesocoxae half as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.44. Pygidium moderately convex and dull, finely and densely punctate, with smooth darker midline, with sparsely scattered, moderately dense, long setae.
</paragraph>
<paragraph id="8B883B49A07BFF94C0FE8DCBF9CDC0E0" blockId="8.[151,1437,151,1293]" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE8DCBFAE8C294" bold="true" box="[199,255,475,501]" pageId="8" pageNumber="379">Legs</emphasis>
very slender and long; femora with two longitudinal rows of setae, between rows coarsely and not densely punctate; metafemur shiny, anterior margin acute, without a continuously serrated line behind anterior margin, ventral posterior margin serrated in apical half and not widened, dorsal posterior margin completely serrated, in basal half with a few long setae which are half as long as width of metafemur. Metatibia slender and long, widest at apex, ratio of width/length: 1/5.2, dorsally carinate, with two groups of spines, basal group well before middle, apical group at three quarters of metatibial length, basally with a few single robust setae; external face slightly longitudinally concave, nearly impunctate, with numerous longitudinal wrinkles; ventral margin serrated, with four robust, equidistant setae; medial face flat, glabrous and impunctate, apex interiorly near tarsal articulation distinctly but bluntly truncate. Tarsomeres ventrally with sparse, short setae, dorsally smooth; metatarsomeres laterally and dorsally carinate, with a strongly serrated ridge ventrally; first metatarsomere distinctly longer than following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia long, bidentate, external edge basally smooth, anterior claws asymmetrical, basal tooth of inner claw small and lobiform.
</paragraph>
<paragraph id="8B883B49A07BFF94C0FE8F9AF881C0C4" blockId="8.[151,1437,151,1293]" box="[199,662,906,933]" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE8F9AFB2DC0C5" bold="true" box="[199,314,906,932]" pageId="8" pageNumber="379">Aedeagus</emphasis>
:
<figureCitation id="130C27CCA07BFF94C1708F9BFBD6C0C4" box="[329,449,907,933]" captionStart="FIGURE 2" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[183,1413,188,1870]" captionTargetId="figure-22@7.[175,1413,179,1882]" captionTargetPageId="7" captionText="FIGURE 2. A–D. Maladera lumlaensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Serica subansiriensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="8" pageNumber="379">Fig. 2E–G</figureCitation>
.
<emphasis id="B943E75BA07BFF94C1F58F9AF83CC0C5" bold="true" box="[460,555,906,932]" pageId="8" pageNumber="379">Habitus</emphasis>
:
<figureCitation id="130C27CCA07BFF94C2008F9BF898C0C4" box="[569,655,907,933]" captionStart="FIGURE 2" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[183,1413,188,1870]" captionTargetId="figure-22@7.[175,1413,179,1882]" captionTargetPageId="7" captionText="FIGURE 2. A–D. Maladera lumlaensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Serica subansiriensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="8" pageNumber="379">Fig. 2H</figureCitation>
.
</paragraph>
<paragraph id="8B883B49A07BFF94C0FE8FBFFB82C0A8" blockId="8.[151,1437,151,1293]" box="[199,405,943,969]" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE8FBFFB09C0A8" bold="true" box="[199,286,943,969]" pageId="8" pageNumber="379">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A07BFF94C0FE8FC2FB7BC7A4" pageId="8" pageNumber="379" type="diagnosis">
<paragraph id="8B883B49A07BFF94C0FE8FC2FB7BC7A4" blockId="8.[151,1437,151,1293]" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE8FC2FB55C08D" bold="true" box="[199,322,978,1004]" pageId="8" pageNumber="379">Diagnosis.</emphasis>
<taxonomicName id="4C3740CAA07BFF94C1738FC3F82CC08C" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[330,571,979,1005]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="subansiriensis" status="sp. nov.">
<emphasis id="B943E75BA07BFF94C1738FC3F82CC08C" box="[330,571,979,1005]" italics="true" pageId="8" pageNumber="379">Serica subansiriensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA07BFF94C27D8FC3F8B7C08C" bold="true" box="[580,672,979,1005]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A2705A20A07BFF94C27D8FC3F8B7C08C" box="[580,672,979,1005]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is very similar to
<taxonomicName id="4C3740CAA07BFF94C34F8FC3FAD8C770" authority="Sreedevi, Speer, Fabrizi &amp; Ahrens, 2018" authorityName="Sreedevi, Speer, Fabrizi &amp; Ahrens" authorityYear="2018" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="zianii">
<emphasis id="B943E75BA07BFF94C34F8FC3FE13C08D" box="[886,1028,979,1005]" italics="true" pageId="8" pageNumber="379">Serica zianii</emphasis>
<bibRefCitation id="EFA646B8A07BFF94C4378FC3FAD8C770" author="Sreedevi, K. &amp; Speer, J. &amp; Fabrizi, S. &amp; Ahrens, D." pageId="8" pageNumber="379" pagination="97 - 128" refId="ref10602" refString="Sreedevi, K., Speer, J., Fabrizi, S. &amp; Ahrens, D. (2018) New species and records of Sericini scarab beetles from the Indian subcontinent (Coleoptera, Scarabaeidae). ZooKeys, 772, 97-128. https://doi.org/10.3897/zookeys.772.25320" type="journal article" year="2018">Sreedevi, Speer, Fabrizi &amp; Ahrens, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3740CAA07BFF94C1368FE7F857C770" authority="Brenske, 1896" authorityName="Brenske" authorityYear="1896" box="[271,576,1015,1041]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B943E75BA07BFF94C1368FE7FB99C770" box="[271,398,1015,1041]" italics="true" pageId="8" pageNumber="379">S. olivacea</emphasis>
<bibRefCitation id="EFA646B8A07BFF94C1A38FE7F857C770" author="Brenske, E." box="[410,576,1015,1041]" pageId="8" pageNumber="379" pagination="150 - 164" refId="ref10330" refString="Brenske, E. (1896) Insectes du Bengale. Melolonthidae. Annales de la Societe Entomologique de Belgique, 40, 150-164." type="journal article" year="1896">
Brenske,
<quantity id="4CCF96ACA07BFF94C2318FE7F857C770" box="[520,576,1015,1041]" metricMagnitude="1" metricUnit="m" metricValue="4.81584" pageId="8" pageNumber="379" unit="in" value="1896.0">1896</quantity>
</bibRefCitation>
</taxonomicName>
in external appearance and genital morphology. The male genitalia in
<taxonomicName id="4C3740CAA07BFF94C5BE8FE7FB20C754" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="subansiriensis" status="sp. nov.">
<emphasis id="B943E75BA07BFF94C5BE8FE7FB20C754" italics="true" pageId="8" pageNumber="379">S. subansiriensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA07BFF94C106880BFB8EC754" bold="true" box="[319,409,1051,1077]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A2705A20A07BFF94C106880BFB8EC754" box="[319,409,1051,1077]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is well distinct from both: the parameres in
<taxonomicName id="4C3740CAA07BFF94C3B7880BFE5CC754" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[910,1099,1051,1077]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="subansiriensis" status="sp. nov.">
<emphasis id="B943E75BA07BFF94C3B7880BFE5CC754" box="[910,1099,1051,1077]" italics="true" pageId="8" pageNumber="379">S. subansiriensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA07BFF94C46A880BFEBAC754" bold="true" box="[1107,1197,1051,1077]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A2705A20A07BFF94C46A880BFEBAC754" box="[1107,1197,1051,1077]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are distinctly shorter compared to the phallobase, the right parameres is strongly reflexed at apex (lateral view), while it is almost flat in
<taxonomicName id="4C3740CAA07BFF94C08C8873FB27C71C" authorityName="Brenske" authorityYear="1896" box="[181,304,1123,1149]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B943E75BA07BFF94C08C8873FB27C71C" box="[181,304,1123,1149]" italics="true" pageId="8" pageNumber="379">S. olivacea</emphasis>
</taxonomicName>
and weakly reflexed in
<taxonomicName id="4C3740CAA07BFF94C2788873F88CC71D" authorityName="Sreedevi, Speer, Fabrizi &amp; Ahrens" authorityYear="2018" box="[577,667,1123,1149]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="zianii">
<emphasis id="B943E75BA07BFF94C2788873F88CC71D" box="[577,667,1123,1149]" italics="true" pageId="8" pageNumber="379">S. zianii</emphasis>
</taxonomicName>
; left parameres in
<taxonomicName id="4C3740CAA07BFF94C3498873F9DCC71D" authorityName="Sreedevi, Speer, Fabrizi &amp; Ahrens" authorityYear="2018" box="[880,971,1123,1149]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="zianii">
<emphasis id="B943E75BA07BFF94C3498873F9DCC71D" box="[880,971,1123,1149]" italics="true" pageId="8" pageNumber="379">S. zianii</emphasis>
</taxonomicName>
is distinctly shorter than in the other two species being less than half as long as the right paramere (
<bibRefCitation id="EFA646B8A07BFF94C3178897F9D3C7C0" author="Ahrens, D." box="[814,964,1159,1185]" pageId="8" pageNumber="379" pagination="205 - 332" refId="ref9546" refString="Ahrens, D. (1999) Revision der Gattung Serica (s. str.) MacLeay des Himalaya-Gebiets (Coleoptera, Melolonthidae). Fragmenta Entomologica, 31, 205-332." type="journal article" year="1999">Ahrens 1999</bibRefCitation>
; figs 36, 37, p. 212;
<bibRefCitation id="EFA646B8A07BFF94C4888897FF80C7C0" author="Sreedevi, K. &amp; Speer, J. &amp; Fabrizi, S. &amp; Ahrens, D." box="[1201,1431,1159,1185]" pageId="8" pageNumber="379" pagination="97 - 128" refId="ref10602" refString="Sreedevi, K., Speer, J., Fabrizi, S. &amp; Ahrens, D. (2018) New species and records of Sericini scarab beetles from the Indian subcontinent (Coleoptera, Scarabaeidae). ZooKeys, 772, 97-128. https://doi.org/10.3897/zookeys.772.25320" type="journal article" year="2018">
Sreedevi
<emphasis id="B943E75BA07BFF94C5228897FF42C7C0" box="[1307,1365,1159,1185]" italics="true" pageId="8" pageNumber="379">et al.</emphasis>
2018
</bibRefCitation>
; figs 4E–H, p. 111).
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A07BFF94C0FE88DFFACFC66C" pageId="8" pageNumber="379" type="etymology">
<paragraph id="8B883B49A07BFF94C0FE88DFFACFC66C" blockId="8.[151,1437,151,1293]" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE88DFFB59C788" bold="true" box="[199,334,1231,1257]" pageId="8" pageNumber="379">Etymology.</emphasis>
<taxonomicName id="4C3740CAA07BFF94C16188DFF85EC788" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[344,585,1231,1257]" class="Insecta" family="Scarabaeidae" genus="Serica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="subansiriensis" status="sp. nov.">
<emphasis id="B943E75BA07BFF94C16188DFF85EC788" box="[344,585,1231,1257]" italics="true" pageId="8" pageNumber="379">Serica subansiriensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA07BFF94C26A88DFF8B8C788" bold="true" box="[595,687,1231,1257]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A2705A20A07BFF94C26A88DFF8B8C788" box="[595,687,1231,1257]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is named after its
<typeStatus id="548C85EBA07BFF94C3B288DFF9ACC788" box="[907,955,1231,1257]" pageId="8" pageNumber="379">type</typeStatus>
locality (adjective in nominative singular case).
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="EDED295910FC58F443890BA95EFA1852">New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)</mods:title>
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<heading id="D0C08C25A076FF99C0AE8FBEF9D4C0A8" bold="true" box="[151,963,942,969]" fontSize="11" level="1" pageId="5" pageNumber="376" reason="1">
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<taxonomicName id="4C3740CAA076FF99C0AE8FBEF94FC0A9" ID-CoL="KTVWV" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[151,856,942,969]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="lumlaensis" status="sp. nov.">
<emphasis id="B943E75BA076FF99C0AE8FBEFB81C0A8" bold="true" box="[151,406,942,969]" italics="true" pageId="5" pageNumber="376">Maladera lumlaensis</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
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,
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(
<figureCitation id="130C27CCA076FF99C0A68FC3FB0DC08C" box="[159,282,979,1005]" captionStart="FIGURE 2" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[183,1413,188,1870]" captionTargetId="figure-22@7.[175,1413,179,1882]" captionTargetPageId="7" captionText="FIGURE 2. A–D. Maladera lumlaensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Serica subansiriensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="5" pageNumber="376">Fig. 2A–D</figureCitation>
)
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<materialsCitation id="3B5F3114A076FF99C0AE8804FE17C74E" box="[151,1024,1044,1071]" country="India" latitude="27.53" location="Lumla" longLatPrecision="74" longitude="91.722" municipality="Tawang" pageId="5" pageNumber="376" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<emphasis id="B943E75BA076FF99C0AE8804FB21C74F" bold="true" box="[151,310,1044,1070]" pageId="5" pageNumber="376">
<typeStatus id="548C85EBA076FF99C0AE8804FAC5C74F" box="[151,210,1044,1070]" pageId="5" pageNumber="376">Type</typeStatus>
locality.
</emphasis>
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:
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,
<collectingMunicipality id="6BECA133A076FF99C25C8805F8D7C74E" box="[613,704,1045,1071]" pageId="5" pageNumber="376">Tawang</collectingMunicipality>
,
<location id="8EE86D92A076FF99C2F28804F900C74F" LSID="urn:lsid:plazi:treatment:039E8A5FA076FF9AC0AE8FBEFACFC0A8:8EE86D92A076FF99C2F28804F900C74F" box="[715,791,1044,1070]" country="India" latitude="27.53" longLatPrecision="74" longitude="91.722" municipality="Tawang" name="Lumla" pageId="5" pageNumber="376" stateProvince="Arunachal Pradesh">Lumla</location>
,
<geoCoordinate id="EE035D8EA076FF99C31B8804F99AC74E" box="[802,909,1044,1071]" degrees="27.530" direction="north" orientation="latitude" pageId="5" pageNumber="376" precision="55" value="27.53">27.530 N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA076FF99C3A18804FE17C74E" box="[920,1024,1044,1071]" degrees="91.722" direction="east" orientation="longitude" pageId="5" pageNumber="376" precision="55" value="91.722">91.722 E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8B883B49A076FF99C0FE8828FB0DC7DF" blockId="5.[151,1437,1044,2043]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8828FB6EC733" bold="true" box="[199,377,1080,1106]" pageId="5" pageNumber="376">Type material.</emphasis>
<materialsCitation id="3B5F3114A076FF99C1BC8828FB0DC7DF" collectingDate="2018-04-15" collectionCode="NZCI" collectorName="Jagdish Saini &amp; The" country="India" latitude="27.53" location="Lumla" longLatPrecision="74" longitude="91.722" municipality="Tawang" pageId="5" pageNumber="376" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
<typeStatus id="548C85EBA076FF99C1BC8828FBE7C733" box="[389,496,1080,1106]" pageId="5" pageNumber="376" type="holotype">Holotype</typeStatus>
,
<emphasis id="B943E75BA076FF99C1C68829F803C734" box="[511,532,1081,1109]" italics="true" pageId="5" pageNumber="376">
<specimenCount id="9D31F0C0A076FF99C1C68829F803C734" box="[511,532,1081,1109]" count="1" pageId="5" pageNumber="376" type="male">♂</specimenCount>
</emphasis>
: “
<collectingCountry id="F3207BD9A076FF99C20B8828F865C733" box="[562,626,1080,1106]" name="India" pageId="5" pageNumber="376">India</collectingCountry>
:
<collectingRegion id="49F3F5ABA076FF99C2468828F94CC733" box="[639,859,1080,1106]" country="India" name="Arunachal Pradesh" pageId="5" pageNumber="376">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="6BECA133A076FF99C3538829F9D2C732" box="[874,965,1081,1107]" pageId="5" pageNumber="376">Tawang</collectingMunicipality>
,
<location id="8EE86D92A076FF99C3ED8828FE37C733" LSID="urn:lsid:plazi:treatment:039E8A5FA076FF9AC0AE8FBEFACFC0A8:8EE86D92A076FF99C3ED8828FE37C733" box="[980,1056,1080,1106]" country="India" latitude="27.53" longLatPrecision="74" longitude="91.722" municipality="Tawang" name="Lumla" pageId="5" pageNumber="376" stateProvince="Arunachal Pradesh">Lumla</location>
,
<geoCoordinate id="EE035D8EA076FF99C4098828FE88C732" box="[1072,1183,1080,1107]" degrees="27.530" direction="north" orientation="latitude" pageId="5" pageNumber="376" precision="55" value="27.53">27.530 N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA076FF99C4968828FF18C733" box="[1199,1295,1080,1106]" degrees="91.722" direction="east" orientation="longitude" pageId="5" pageNumber="376" precision="55" value="91.722">91.722E</geoCoordinate>
,
<date id="FF891D89A076FF99C5268828FF8FC732" box="[1311,1432,1080,1107]" pageId="5" pageNumber="376" value="2018-04-15">
<collectingDate id="EFCDE461A076FF99C5268828FF8FC732" box="[1311,1432,1080,1107]" pageId="5" pageNumber="376" value="2018-04-15">15.iv.2018</collectingDate>
</date>
, leg.
<collectorName id="26C25E9FA076FF99C0F0884CFB76C717" box="[201,353,1116,1142]" pageId="5" pageNumber="376">Jagdish Saini</collectorName>
” (
<collectionCode id="ED26A38CA076FF99C145884CFBD3C717" box="[380,452,1116,1142]" pageId="5" pageNumber="376">NZCI</collectionCode>
).
<collectorName id="26C25E9FA076FF99C1EE884CF814C717" box="[471,515,1116,1142]" pageId="5" pageNumber="376">The</collectorName>
<typeStatus id="548C85EBA076FF99C233884CF87BC717" box="[522,620,1116,1142]" pageId="5" pageNumber="376" type="holotype">holotype</typeStatus>
specimen bears additionally a red printed label with the following text: “
<taxonomicName id="4C3740CAA076FF99C09D8890FB85C7FB" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[164,402,1152,1178]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="lumlaensis" status="sp. nov.">
<emphasis id="B943E75BA076FF99C09D8890FB85C7FB" box="[164,402,1152,1178]" italics="true" pageId="5" pageNumber="376">Maladera lumlaensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA076FF99C1A38890FBE3C7FB" bold="true" box="[410,500,1152,1178]" pageId="5" pageNumber="376">
<taxonomicNameLabel id="A2705A20A076FF99C1A38890FBE3C7FB" box="[410,500,1152,1178]" pageId="5" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="548C85EBA076FF99C23A8890F88DC7FB" box="[515,666,1152,1178]" pageId="5" pageNumber="376" type="holotype">HOLOTYPE</typeStatus>
, Devanshu Gupta, Debika Bhunia, Dirk Ahrens &amp; Kailash Chandra, Det. 2024”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A076FF9AC0FE88D8FB82C1AC" lastPageId="6" lastPageNumber="377" pageId="5" pageNumber="376" type="description">
<paragraph id="8B883B49A076FF99C0FE88D8FEA7C782" blockId="5.[151,1437,1044,2043]" box="[199,1200,1224,1251]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE88D8F806C783" bold="true" box="[199,529,1224,1250]" pageId="5" pageNumber="376">
Description of the
<typeStatus id="548C85EBA076FF99C19A88D8F81BC783" box="[419,524,1224,1250]" pageId="5" pageNumber="376" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CCF96ACA076FF99C24F88D8F8CCC782" box="[630,731,1224,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="5" pageNumber="376" unit="mm" value="11.5">11.5 mm</quantity>
, length of elytra:
<quantity id="4CCF96ACA076FF99C39988D8F9EFC782" box="[928,1016,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="5" pageNumber="376" unit="mm" value="9.4">9.4 mm</quantity>
, width:
<quantity id="4CCF96ACA076FF99C46A88D8FEBBC782" box="[1107,1196,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="5" pageNumber="376" unit="mm" value="8.4">8.4 mm</quantity>
.
</paragraph>
<paragraph id="8B883B49A076FF99C0FE88FCFAE8C64B" blockId="5.[151,1437,1044,2043]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE88FCFB12C667" bold="true" box="[199,261,1260,1286]" pageId="5" pageNumber="376">Body</emphasis>
oval, dark reddish brown, dull, labroclypeus shiny, except some single setae on head dorsal surface nearly glabrous.
</paragraph>
<paragraph id="8B883B49A076FF99C0FE8924FEA1C52B" blockId="5.[151,1437,1044,2043]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8924FB10C62F" bold="true" box="[199,263,1332,1358]" pageId="5" pageNumber="376">Head</emphasis>
. Labroclypeus moderately wide and trapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly emarginate medially, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct blunt angle; surface flat, finely and densely punctate, glabrous; frontoclypeal suture distinctly incised, evenly curved; smooth area anterior to eye weakly convex, 2.5 times as wide as long; ocular canthus short and wide (1/3 of ocular diameter), finely and densely punctate, with a terminal seta. Frons with sparse, fine punctures and a single long seta beside each eye. Eyes large, ratio diameter/ interocular width: 0.78. Antenna with ten antennomeres; club with three antennomeres and straight, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8B883B49A076FF99C0FE8A44FEB2C59E" blockId="5.[151,1437,1044,2043]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8A44FB57C50F" bold="true" box="[199,320,1620,1646]" pageId="5" pageNumber="376">Pronotum</emphasis>
moderately wide, widest shortly before base, lateral margins evenly convex and convergent anteriorly, slightly convexly narrowed towards base, anterior angles distinctly produced and sharp, posterior angles blunt, strongly rounded at tip; anterior margin convex, with marginal line widely lacking, base without marginal line; surface sparsely and finely punctate, glabrous; lateral margin finely setose, anterior margin glabrous; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, dense punctures.
</paragraph>
<paragraph id="8B883B49A076FF99C0FE8B18F8C7C4EF" blockId="5.[151,1437,1044,2043]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8B18FB04C443" bold="true" box="[199,275,1800,1826]" pageId="5" pageNumber="376">Elytra</emphasis>
widest at posterior third, striae distinctly impressed, finely and sparsely punctate, intervals nearly flat, with fine and evenly dense punctures, nearly entirely glabrous; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membranous, with a rim of microtrichomes (visible at ca 100x magnification).
</paragraph>
<paragraph id="8B883B49A076FF9AC0FE8B88FF6CC3B4" blockId="5.[151,1437,1044,2043]" lastBlockId="6.[151,1437,151,969]" lastPageId="6" lastPageNumber="377" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8B88FB96C4D3" bold="true" box="[199,385,1944,1970]" pageId="5" pageNumber="376">Ventral surface</emphasis>
dull, coarsely and densely punctate, glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a loose transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.49. Pygidium weakly convex and dull, coarsely and densely punctate, without smooth midline, glabrous except a few robust setae along apical margin.
</paragraph>
<paragraph id="8B883B49A075FF9AC0FE8CCFF949C1E4" blockId="6.[151,1437,151,969]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8CCFFAE8C398" bold="true" box="[199,255,223,249]" pageId="6" pageNumber="377">Legs</emphasis>
short and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without adjacent serrated line, anterior row of setae completely reduced; posterior margin smooth, widened and smooth at apex ventrally, not serrate dorsally, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.7, sharply carinate dorsally, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, with a few short, robust setae and a short serrated line parallel to dorsal margin basally; lateral face longitudinally convex, shiny but basal third dull, impunctate and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.
</paragraph>
<paragraph id="8B883B49A075FF9AC0FE8E9EF880C1C8" blockId="6.[151,1437,151,969]" box="[199,663,654,681]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8E9EFB2DC1C9" bold="true" box="[199,314,654,680]" pageId="6" pageNumber="377">Aedeagus</emphasis>
:
<figureCitation id="130C27CCA075FF9AC1708E9FFBD4C1C8" box="[329,451,655,681]" captionStart="FIGURE 2" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[183,1413,188,1870]" captionTargetId="figure-22@7.[175,1413,179,1882]" captionTargetPageId="7" captionText="FIGURE 2. A–D. Maladera lumlaensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Serica subansiriensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="6" pageNumber="377">Fig. 2A–C</figureCitation>
.
<emphasis id="B943E75BA075FF9AC1F48E9EF83BC1C9" bold="true" box="[461,556,654,680]" pageId="6" pageNumber="377">Habitus</emphasis>
:
<figureCitation id="130C27CCA075FF9AC2028E9FF884C1C8" box="[571,659,655,681]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="6" pageNumber="377">Fig. 1D</figureCitation>
.
</paragraph>
<paragraph id="8B883B49A075FF9AC0FE8EA3FB82C1AC" blockId="6.[151,1437,151,969]" box="[199,405,691,717]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8EA3FB09C1AC" bold="true" box="[199,286,691,717]" pageId="6" pageNumber="377">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A075FF9AC0FE8EC6F95FC190" box="[199,840,726,753]" pageId="6" pageNumber="377" type="distribution">
<paragraph id="8B883B49A075FF9AC0FE8EC6F95FC190" blockId="6.[151,1437,151,969]" box="[199,840,726,753]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8EC6FB76C191" bold="true" box="[199,353,726,752]" pageId="6" pageNumber="377">Distribution.</emphasis>
Species only known from the
<typeStatus id="548C85EBA075FF9AC28E8EC7F8F0C190" box="[695,743,727,753]" pageId="6" pageNumber="377">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A075FF9AC0FE8EEAF806C0E0" pageId="6" pageNumber="377" type="diagnosis">
<paragraph id="8B883B49A075FF9AC0FE8EEAF806C0E0" blockId="6.[151,1437,151,969]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8EEAFB2DC075" bold="true" box="[199,314,762,788]" pageId="6" pageNumber="377">Diagnosis</emphasis>
.
<taxonomicName id="4C3740CAA075FF9AC1708EEBF82FC074" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[329,568,763,789]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="lumlaensis" status="sp. nov.">
<emphasis id="B943E75BA075FF9AC1708EEBF82FC074" box="[329,568,763,789]" italics="true" pageId="6" pageNumber="377">Maladera lumlaensis</emphasis>
</taxonomicName>
<emphasis id="B943E75BA075FF9AC2068EEBF88DC074" bold="true" box="[575,666,763,789]" pageId="6" pageNumber="377">
<taxonomicNameLabel id="A2705A20A075FF9AC2068EEBF88DC074" box="[575,666,763,789]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is in the shape of genitalia and external morphology very similar to
<taxonomicName id="4C3740CAA075FF9AC0AE8F0FFBE0C058" authority="(Brenske, 1899)" baseAuthorityName="Brenske" baseAuthorityYear="1899" box="[151,503,799,825]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="sempiterna">
<emphasis id="B943E75BA075FF9AC0AE8F0FFB2EC059" box="[151,313,799,824]" italics="true" pageId="6" pageNumber="377">M. sempiterna</emphasis>
(
<bibRefCitation id="EFA646B8A075FF9AC1708F0FFBF8C058" author="Brenske, E." box="[329,495,799,825]" pageId="6" pageNumber="377" pagination="205 - 403" refId="ref10357" refString="Brenske, E. (1899) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205-403." type="journal article" year="1899">Brenske, 1899</bibRefCitation>
)
</taxonomicName>
. The new species differs from
<taxonomicName id="4C3740CAA075FF9AC3768F0FF9E7C059" baseAuthorityName="Brenske" baseAuthorityYear="1899" box="[847,1008,799,824]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="sempiterna">
<emphasis id="B943E75BA075FF9AC3768F0FF9E7C059" box="[847,1008,799,824]" italics="true" pageId="6" pageNumber="377">M. sempiterna</emphasis>
</taxonomicName>
by the longer antennal club as well as by the straight parameres which are distinctly cleft before the apex (they are not cleft in
<taxonomicName id="4C3740CAA075FF9AC44A8F53FF03C03D" baseAuthorityName="Brenske" baseAuthorityYear="1899" box="[1139,1300,835,860]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="sempiterna">
<emphasis id="B943E75BA075FF9AC44A8F53FF03C03D" box="[1139,1300,835,860]" italics="true" pageId="6" pageNumber="377">M. sempiterna</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFA646B8A075FF9AC5148F53FB3DC0E0" author="Ahrens, D. &amp; Fabrizi, S." pageId="6" pageNumber="377" pagination="1 - 355" refId="ref9865" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1-355." type="journal article" year="2016">Ahrens &amp; Fabrizi 2016</bibRefCitation>
; fig. 11P–R, p. 299).
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A075FF9AC0FE8F9BFACFC0A8" pageId="6" pageNumber="377" type="etymology">
<paragraph id="8B883B49A075FF9AC0FE8F9BFACFC0A8" blockId="6.[151,1437,151,969]" pageId="6" pageNumber="377">
<emphasis id="B943E75BA075FF9AC0FE8F9BFB59C0C4" bold="true" box="[199,334,907,933]" pageId="6" pageNumber="377">Etymology.</emphasis>
The newly described species is named after its
<typeStatus id="548C85EBA075FF9AC3BF8F9BF9A1C0C4" box="[902,950,907,933]" pageId="6" pageNumber="377">type</typeStatus>
locality (adjective in nominative singular case).
</paragraph>
</subSubSection>
</treatment>
</document>

263
data/03/9E/8A/039E8A5FA077FF99C0AE8F52FF45C03C.xml Normal file
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@ -0,0 +1,263 @@
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<mods:mods id="A932B8FBC69DEEF5FD2D2F4DE5084F30" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1696941E4581167BB4E787A336573661">New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)</mods:title>
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<mods:name id="FB24702B5ED57EE5E1C21E2C6FB3F8F1" type="personal">
<mods:role id="AD67D2467E16A6A4129CAAC1437F5788">
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<mods:namePart id="B4B28867707DFE7DD0D6EEACCB51DEC2">Gupta, Devanshu</mods:namePart>
<mods:affiliation id="1E26ED5217C7C9CF72819B11FE305FA6">Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India</mods:affiliation>
</mods:name>
<mods:name id="C18A0821CE1A2F4B2C45ECF47207B4A2" type="personal">
<mods:role id="A0CD0E74C9B2A070718B978296D424A8">
<mods:roleTerm id="94D6DFAF078647091DABC8FCB9201D2A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="859E6262E1C701CDF43A6F012D37E1FE">Bhunia, Debika</mods:namePart>
<mods:affiliation id="4A75F298FDF66B432661D745DDE9496B">Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India</mods:affiliation>
</mods:name>
<mods:name id="B57A0F215C42843237FA69E4B91A11D4" type="personal">
<mods:role id="D6B464281D17A5587C63EEBFB3BCCCC7">
<mods:roleTerm id="F45809AD85026E35A84FCBA405DD5176">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F194CA9E260811EA2FFED1ACEF826D6">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="0AA0A1F93B71FEEBD6AA981B89B35BE8" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="BB82726016CFDEEDBD0B372095A9D2F1" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="2D4F472C435E72044A485137A5021A9F" type="personal">
<mods:role id="6C967F5540289A0825B93E0F0AE18E5E">
<mods:roleTerm id="01F51D28C7D480CA5534D639BEF99D63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="14906CD10E9AE4721CF852FA69CBC626">Chandra, Kailash</mods:namePart>
<mods:affiliation id="DB883C5174DC61025B4DCD7AF45D0A56">Flat No. 620, Tower Ferrara, Mahagun Mezzaria Sector- 78, Noida- 201301, Gautam Budha Nagar, U. P</mods:affiliation>
</mods:name>
<mods:typeOfResource id="9F541DB2CBE07924946E512BE56F5D86">text</mods:typeOfResource>
<mods:relatedItem id="0D06B49ACD3682B7386D41AB4D82D716" type="host">
<mods:titleInfo id="06E5607310D705A16A04E88B22293E0F">
<mods:title id="6EEFACD7A1088DE2B29D5BA6B084DC0E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A120D20046836BF89D01910F3DD8625E">
<mods:date id="74913751CA7E60FB256161B204A712E4">2025</mods:date>
<mods:detail id="A3D85A4FB789BE204AC570267BBA4C83" type="pubDate">
<mods:number id="6E2CB49A95601169D30F655D3BF7110E">2025-03-27</mods:number>
</mods:detail>
<mods:detail id="1C413BB6372213C3800CB92CE9061903" type="volume">
<mods:number id="D4E1DDB2769B2EDD894C48CE38128F89">5613</mods:number>
</mods:detail>
<mods:detail id="D6ABEE0FA27B9B642BCA51E48F2F9F85" type="issue">
<mods:number id="BD81CE62F8BECA898586049DBF2C90B4">2</mods:number>
</mods:detail>
<mods:extent id="F3983E97C18F87EAC09DC3F108E3521F" unit="page">
<mods:start id="CF25A8DF026D0345C6965734F645F555">371</mods:start>
<mods:end id="27226D944C6DBB9651D1C4588EBBCC49">385</mods:end>
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<mods:location id="5AA3ABB7DBC4178A386B46AF159591B7">
<mods:url id="DFB542C41A01515438F4DA01D0A04271">https://doi.org/10.11646/zootaxa.5613.2.11</mods:url>
</mods:location>
<mods:classification id="6FCDD2FE97B13E6066CB1E414C2613F6">journal article</mods:classification>
<mods:identifier id="D1215185BD3D65749106EC4030E04336" type="DOI">10.11646/zootaxa.5613.2.11</mods:identifier>
<mods:identifier id="0D915DB9C643C825A42BD01E07AB4C69" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A0F6E5CD15C6BECA1436C26036007B49" type="ZooBank">2972195A-6FD5-488A-8925-51D24677DFEC</mods:identifier>
</mods:mods>
<treatment id="039E8A5FA077FF99C0AE8F52FF45C03C" LSID="urn:lsid:plazi:treatment:039E8A5FA077FF99C0AE8F52FF45C03C" httpUri="http://treatment.plazi.org/id/039E8A5FA077FF99C0AE8F52FF45C03C" lastPageId="5" lastPageNumber="376" pageId="4" pageNumber="375">
<subSubSection id="C32D68C2A077FF98C0AE8F52F9DDC03C" box="[151,970,834,861]" pageId="4" pageNumber="375" type="nomenclature">
<paragraph id="8B883B49A077FF98C0AE8F52F9DDC03C" blockId="4.[151,970,834,897]" box="[151,970,834,861]" pageId="4" pageNumber="375">
<heading id="D0C08C25A077FF98C0AE8F52F9DDC03C" bold="true" box="[151,970,834,861]" fontSize="11" level="1" pageId="4" pageNumber="375" reason="1">
<emphasis id="B943E75BA077FF98C0AE8F52F9DDC03C" bold="true" box="[151,970,834,861]" pageId="4" pageNumber="375">
<taxonomicName id="4C3740CAA077FF98C0AE8F52F948C03D" ID-CoL="KTVVZ" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[151,863,834,861]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="barasingha" status="sp. nov.">
<emphasis id="B943E75BA077FF98C0AE8F52FB8AC03C" bold="true" box="[151,413,834,861]" italics="true" pageId="4" pageNumber="375">Maladera barasingha</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
,
<taxonomicNameLabel id="A2705A20A077FF98C3558F53F9DDC03C" box="[876,970,835,861]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A077FF98C0AE8F77FB04C0E0" box="[151,275,871,897]" pageId="4" pageNumber="375" type="description">
<paragraph id="8B883B49A077FF98C0AE8F77FB04C0E0" blockId="4.[151,970,834,897]" box="[151,275,871,897]" pageId="4" pageNumber="375">
(
<figureCitation id="130C27CCA077FF98C0A68F77FB1DC0E0" box="[159,266,871,897]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="4" pageNumber="375">Fig. 1I–L</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A077FF98C0AE8FBFFAF4C738" pageId="4" pageNumber="375" type="materials_examined">
<paragraph id="8B883B49A077FF98C0AE8FBFF948C0A8" blockId="4.[151,1437,942,2013]" box="[151,863,942,969]" pageId="4" pageNumber="375">
<materialsCitation id="3B5F3114A077FF98C0AE8FBFF94BC0A8" box="[151,860,942,969]" country="India" latitude="25.0396" location="Tamenlong" longLatPrecision="50" longitude="93.468" pageId="4" pageNumber="375" specimenCount="1" stateProvince="Manipur" typeStatus="holotype">
<emphasis id="B943E75BA077FF98C0AE8FBFFB21C0A9" bold="true" box="[151,310,942,969]" pageId="4" pageNumber="375">
<typeStatus id="548C85EBA077FF98C0AE8FBFFAC5C0A8" box="[151,210,943,969]" pageId="4" pageNumber="375">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3207BD9A077FF98C1058FBFFB6FC0A8" box="[316,376,943,969]" name="India" pageId="4" pageNumber="375">India</collectingCountry>
,
<collectingRegion id="49F3F5ABA077FF98C1BD8FBFFBF1C0A8" box="[388,486,943,969]" country="India" name="Manipur" pageId="4" pageNumber="375">Manipur</collectingRegion>
,
<location id="8EE86D92A077FF98C1C88FBFF865C0A8" LSID="urn:lsid:plazi:treatment:039E8A5FA077FF99C0AE8F52FF45C03C:8EE86D92A077FF98C1C88FBFF865C0A8" box="[497,626,943,969]" country="India" latitude="25.0396" longLatPrecision="50" longitude="93.468" name="Tamenlong" pageId="4" pageNumber="375" stateProvince="Manipur">Tamenlong</location>
,
<geoCoordinate id="EE035D8EA077FF98C2458FBEF8F9C0A8" box="[636,750,942,969]" degrees="25.0396" direction="north" orientation="latitude" pageId="4" pageNumber="375" precision="5" value="25.0396">25.0396N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA077FF98C2C08FBFF94BC0A8" box="[761,860,943,969]" degrees="93.468" direction="east" orientation="longitude" pageId="4" pageNumber="375" precision="55" value="93.468">93.468E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8B883B49A077FF98C0FE8FC3FAF4C738" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8FC3FB61C08D" bold="true" box="[199,374,978,1005]" pageId="4" pageNumber="375">Type material.</emphasis>
<materialsCitation id="3B5F3114A077FF98C1448FC3FAF4C738" collectingDate="1992-11-24" collectionCode="NZCI" collectorName="A. K. Mondal &amp; The" country="India" latitude="25.0396" location="Tamenlong" longLatPrecision="50" longitude="93.468" pageId="4" pageNumber="375" specimenCount="1" specimenCount-male="1" stateProvince="Manipur" typeStatus="holotype">
<typeStatus id="548C85EBA077FF98C1448FC3FBFFC08C" box="[381,488,979,1005]" pageId="4" pageNumber="375" type="holotype">Holotype</typeStatus>
,
<emphasis id="B943E75BA077FF98C1CD8FC4F81EC091" box="[500,521,980,1008]" italics="true" pageId="4" pageNumber="375">
<specimenCount id="9D31F0C0A077FF98C1CD8FC4F81EC091" box="[500,521,980,1008]" count="1" pageId="4" pageNumber="375" type="male">♂</specimenCount>
</emphasis>
: “
<collectingCountry id="F3207BD9A077FF98C21A8FC3F876C08C" box="[547,609,979,1005]" name="India" pageId="4" pageNumber="375">India</collectingCountry>
,
<collectingRegion id="49F3F5ABA077FF98C2548FC3F8D8C08C" box="[621,719,979,1005]" country="India" name="Manipur" pageId="4" pageNumber="375">Manipur</collectingRegion>
,
<location id="8EE86D92A077FF98C2E38FC3F94CC08C" LSID="urn:lsid:plazi:treatment:039E8A5FA077FF99C0AE8F52FF45C03C:8EE86D92A077FF98C2E38FC3F94CC08C" box="[730,859,979,1005]" country="India" latitude="25.0396" longLatPrecision="50" longitude="93.468" name="Tamenlong" pageId="4" pageNumber="375" stateProvince="Manipur">Tamenlong</location>
,
<geoCoordinate id="EE035D8EA077FF98C35E8FC2F9CFC08C" box="[871,984,978,1005]" degrees="25.0396" direction="north" orientation="latitude" pageId="4" pageNumber="375" precision="5" value="25.0396">25.0396N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA077FF98C3DC8FC3FE52C08C" box="[997,1093,979,1005]" degrees="93.468" direction="east" orientation="longitude" pageId="4" pageNumber="375" precision="55" value="93.468">93.468E</geoCoordinate>
,
<date id="FF891D89A077FF98C4688FC3FEDBC08C" box="[1105,1228,979,1005]" pageId="4" pageNumber="375" value="1992-11-24">
<collectingDate id="EFCDE461A077FF98C4688FC3FEDBC08C" box="[1105,1228,979,1005]" pageId="4" pageNumber="375" value="1992-11-24">24.xi.1992</collectingDate>
</date>
, leg.
<collectorName id="26C25E9FA077FF98C53E8FC3FF8BC08C" box="[1287,1436,979,1005]" pageId="4" pageNumber="375">A.K. Mondal</collectorName>
and Party” (
<collectionCode id="ED26A38CA077FF98C1228FE7FB74C770" box="[283,355,1015,1041]" pageId="4" pageNumber="375">NZCI</collectionCode>
).
<collectorName id="26C25E9FA077FF98C14A8FE7FB88C770" box="[371,415,1015,1041]" pageId="4" pageNumber="375">The</collectorName>
<typeStatus id="548C85EBA077FF98C19C8FE7F810C770" box="[421,519,1015,1041]" pageId="4" pageNumber="375" type="holotype">holotype</typeStatus>
specimen bears additionally a red printed label with the following text: “
<taxonomicName id="4C3740CAA077FF98C5178FE7FB00C754" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="barasingha" status="sp. nov.">
<emphasis id="B943E75BA077FF98C5178FE7FB00C754" italics="true" pageId="4" pageNumber="375">Maladera barasingha</emphasis>
</taxonomicName>
<emphasis id="B943E75BA077FF98C11D880BFB94C754" bold="true" box="[292,387,1051,1077]" pageId="4" pageNumber="375">
<taxonomicNameLabel id="A2705A20A077FF98C11D880BFB94C754" box="[292,387,1051,1077]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="548C85EBA077FF98C1AE880BF839C754" box="[407,558,1051,1077]" pageId="4" pageNumber="375" type="holotype">HOLOTYPE</typeStatus>
, Devanshu Gupta, Debika Bhunia, Dirk Ahrens &amp; Kailash Chandra, Det. 2024”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A077FF99C0FE8872FB82C1E4" lastPageId="5" lastPageNumber="376" pageId="4" pageNumber="375" type="description">
<paragraph id="8B883B49A077FF98C0FE8872FEB4C71C" blockId="4.[151,1437,942,2013]" box="[199,1187,1122,1149]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8872F81DC71C" bold="true" box="[199,522,1122,1149]" pageId="4" pageNumber="375">
Description of the
<typeStatus id="548C85EBA077FF98C19A8873F81DC71C" box="[419,522,1123,1149]" pageId="4" pageNumber="375" type="holotype">holotype</typeStatus>
</emphasis>
. Length:
<quantity id="4CCF96ACA077FF98C24F8873F8D9C71C" box="[630,718,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="4" pageNumber="375" unit="mm" value="8.6">8.6 mm</quantity>
, length of elytra:
<quantity id="4CCF96ACA077FF98C3AA8873F9FCC71C" box="[915,1003,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="375" unit="mm" value="4.7">4.7 mm</quantity>
, width:
<quantity id="4CCF96ACA077FF98C47F8873FE88C71C" box="[1094,1183,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="375" unit="mm" value="4.2">4.2 mm</quantity>
.
</paragraph>
<paragraph id="8B883B49A077FF98C0FE8897FAE8C7A4" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8897FB12C7C0" bold="true" box="[199,261,1159,1185]" pageId="4" pageNumber="375">Body</emphasis>
oval, dark reddish brown, dull, labroclypeus shiny, except some single setae on head dorsal surface nearly glabrous.
</paragraph>
<paragraph id="8B883B49A077FF98C0FE88DFFEA1C684" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE88DFFB10C788" bold="true" box="[199,263,1231,1257]" pageId="4" pageNumber="375">Head</emphasis>
. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles strongly rounded, anterior margin weakly emarginate medially, margins weakly reflexed; lateral margin and ocular canthus produce an indistinct blunt angle; surface flat, finely and densely punctate, glabrous; frontoclypeal suture distinctly incised, evenly curved; smooth area anterior to eye weakly convex, 2.5 times as wide as long; ocular canthus short and wide (1/3 of ocular diameter), finely and densely punctate, with a terminal seta. Frons with sparse, fine punctures and a single, long seta beside each eye. Eyes small, ratio diameter/ interocular width: 0.58. Antenna with ten antennomeres; club with three antennomeres and reflexed, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8B883B49A077FF98C0FE89FFFEB2C5F8" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE89FFFB57C568" bold="true" box="[199,320,1519,1545]" pageId="4" pageNumber="375">Pronotum</emphasis>
moderately wide, widest shortly before base, lateral margins evenly convex and convergent anteriorly, slightly convexly narrowed towards base, anterior angles distinctly produced and sharp, posterior angles blunt, strongly rounded in the tip; anterior margin convex, with marginal line widely lacking, base without marginal line; surface sparsely and finely punctate, glabrous; lateral margin finely setose, anterior margin glabrous; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, dense punctures.
</paragraph>
<paragraph id="8B883B49A077FF98C0FE8AB3FB7AC448" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8AB3FB04C5DC" bold="true" box="[199,275,1699,1725]" pageId="4" pageNumber="375">Elytra</emphasis>
widest at posterior third, striae distinctly impressed, finely and sparsely punctate, intervals nearly flat, with fine and evenly dense punctures, nearly entirely glabrous; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a rim of microtrichomes.
</paragraph>
<paragraph id="8B883B49A077FF98C0FE8B23FF35C4BC" blockId="4.[151,1437,942,2013]" pageId="4" pageNumber="375">
<emphasis id="B943E75BA077FF98C0FE8B23FB96C42D" bold="true" box="[199,385,1842,1869]" pageId="4" pageNumber="375">Ventral surface</emphasis>
dull, coarsely and densely punctate, glabrous, metasternal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a loose transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.52. Pygidium weakly convex and dull, coarsely and densely punctate, without midline, glabrous except a few robust setae along apical margin.
</paragraph>
<paragraph id="8B883B49A076FF99C0FE8C87F949C15C" blockId="5.[151,1437,151,861]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8C87FAE8C3D0" bold="true" box="[199,255,151,177]" pageId="5" pageNumber="376">Legs</emphasis>
long and slender, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without adjacent serrated line, anterior row of setae completely reduced; posterior ventral margin smooth, widened and smooth at apex, posterior dorsal margin not serrate, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.7, sharply carinate dorsally, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, with a few short, robust setae and a short serrated line parallel to dorsal margin basally; lateral face longitudinally convex, shiny but basal third dull, impunctate and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth, subventral longitudinal carina; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.
</paragraph>
<paragraph id="8B883B49A076FF99C0FE8E56F89CC100" blockId="5.[151,1437,151,861]" box="[199,651,582,609]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8E56FB2DC101" bold="true" box="[199,314,582,608]" pageId="5" pageNumber="376">Aedeagus</emphasis>
:
<figureCitation id="130C27CCA076FF99C1708E57FBA1C100" box="[329,438,583,609]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="5" pageNumber="376">Fig. 1I–K</figureCitation>
.
<emphasis id="B943E75BA076FF99C1FD8E56F834C101" bold="true" box="[452,547,582,608]" pageId="5" pageNumber="376">Habitus</emphasis>
:
<figureCitation id="130C27CCA076FF99C20B8E57F893C100" box="[562,644,583,609]" captionStart="FIGURE 1" captionStartId="2.[152,255,1908,1932]" captionTargetBox="[207,1392,204,1854]" captionTargetId="figure-28@2.[189,1398,179,1885]" captionTargetPageId="2" captionText="FIGURE 1. A–D. Maladera champhaiensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Neoserica churachandpurensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera barasingha Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="5" pageNumber="376">
Fig.
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.
</paragraph>
<paragraph id="8B883B49A076FF99C0FE8E7BFB82C1E4" blockId="5.[151,1437,151,861]" box="[199,405,619,645]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8E7BFB09C1E4" bold="true" box="[199,286,619,645]" pageId="5" pageNumber="376">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A076FF99C0FE8E9EF973C1C8" box="[199,868,654,681]" pageId="5" pageNumber="376" type="distribution">
<paragraph id="8B883B49A076FF99C0FE8E9EF973C1C8" blockId="5.[151,1437,151,861]" box="[199,868,654,681]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8E9EFB76C1C9" bold="true" box="[199,353,654,680]" pageId="5" pageNumber="376">Distribution.</emphasis>
Species so far known only from
<typeStatus id="548C85EBA076FF99C2EA8E9FF914C1C8" box="[723,771,655,681]" pageId="5" pageNumber="376">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A076FF99C0FE8EA2FF1CC074" pageId="5" pageNumber="376" type="diagnosis">
<paragraph id="8B883B49A076FF99C0FE8EA2FF1CC074" blockId="5.[151,1437,151,861]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8EA2FB2DC1AD" bold="true" box="[199,314,690,716]" pageId="5" pageNumber="376">Diagnosis</emphasis>
.
<taxonomicName id="4C3740CAA076FF99C1708EA3F829C1AC" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[329,574,691,717]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="barasingha" status="sp. nov.">
<emphasis id="B943E75BA076FF99C1708EA3F829C1AC" box="[329,574,691,717]" italics="true" pageId="5" pageNumber="376">Maladera barasingha</emphasis>
</taxonomicName>
<emphasis id="B943E75BA076FF99C27C8EA3F888C1AC" bold="true" box="[581,671,691,717]" pageId="5" pageNumber="376">
<taxonomicNameLabel id="A2705A20A076FF99C27C8EA3F888C1AC" box="[581,671,691,717]" pageId="5" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is in shape of the male genitalia and external morphology similar to
<taxonomicName id="4C3740CAA076FF99C0AE8EC7F916C190" authority="Bhunia, Gupta, Sarkar &amp; Ahrens, 2023" authorityName="Bhunia, Gupta, Sarkar &amp; Ahrens" authorityYear="2023" box="[151,769,727,753]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="kolkataensis">
<emphasis id="B943E75BA076FF99C0AE8EC7FB51C190" box="[151,326,727,753]" italics="true" pageId="5" pageNumber="376">M. kolkataensis</emphasis>
Bhunia, Gupta, Sarkar &amp; Ahrens, 2023
</taxonomicName>
. The new species differs from
<taxonomicName id="4C3740CAA076FF99C46D8EC7FF14C190" authorityName="Bhunia, Gupta, Sarkar &amp; Ahrens" authorityYear="2023" box="[1108,1283,727,753]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="kolkataensis">
<emphasis id="B943E75BA076FF99C46D8EC7FF14C190" box="[1108,1283,727,753]" italics="true" pageId="5" pageNumber="376">M. kolkataensis</emphasis>
</taxonomicName>
by the longer antennal club as well as by the curved and protruded parameres (Bhunia
<emphasis id="B943E75BA076FF99C3FF8EEBF9E9C074" box="[966,1022,763,789]" italics="true" pageId="5" pageNumber="376">et al.</emphasis>
2023; figs 1–4, p. 353).
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A076FF99C0FE8F0FFF45C03C" pageId="5" pageNumber="376" type="etymology">
<paragraph id="8B883B49A076FF99C0FE8F0FFF45C03C" blockId="5.[151,1437,151,861]" pageId="5" pageNumber="376">
<emphasis id="B943E75BA076FF99C0FE8F0FFB5FC058" bold="true" box="[199,328,799,825]" pageId="5" pageNumber="376">Etymology</emphasis>
.
<taxonomicName id="4C3740CAA076FF99C16A8F0FF852C058" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[339,581,799,825]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="barasingha" status="sp. nov.">
<emphasis id="B943E75BA076FF99C16A8F0FF852C058" box="[339,581,799,825]" italics="true" pageId="5" pageNumber="376">Maladera barasingha</emphasis>
</taxonomicName>
<emphasis id="B943E75BA076FF99C2718F0FF889C058" bold="true" box="[584,670,799,825]" pageId="5" pageNumber="376">
<taxonomicNameLabel id="A2705A20A076FF99C2718F0FF889C058" box="[584,670,799,825]" pageId="5" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is named after the
<collectingCountry id="F3207BD9A076FF99C35B8F0FF98BC058" box="[866,924,799,825]" name="India" pageId="5" pageNumber="376">India</collectingCountry>
swamp deer(
<taxonomicName id="4C3740CAA076FF99C4168F0FFF18C058" baseAuthorityName="G. Cuvier" baseAuthorityYear="1823" box="[1071,1295,799,825]" class="Mammalia" family="Cervidae" genus="Rucervus" kingdom="Animalia" order="Artiodactyla" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="duvaucelii">
<emphasis id="B943E75BA076FF99C4168F0FFF18C058" box="[1071,1295,799,825]" italics="true" pageId="5" pageNumber="376">Rucervus duvaucelii</emphasis>
</taxonomicName>
),barasingha (noun in apposition). The aedeagus of
<taxonomicName id="4C3740CAA076FF99C27F8F53F92BC03C" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[582,828,835,861]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="barasingha" status="sp. nov.">
<emphasis id="B943E75BA076FF99C27F8F53F92BC03C" box="[582,828,835,861]" italics="true" pageId="5" pageNumber="376">Maladera barasingha</emphasis>
</taxonomicName>
<emphasis id="B943E75BA076FF99C37B8F53F98BC03C" bold="true" box="[834,924,835,861]" pageId="5" pageNumber="376">
<taxonomicNameLabel id="A2705A20A076FF99C37B8F53F98BC03C" box="[834,924,835,861]" pageId="5" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles the antler of the barasingha.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="BB1E66E83C87BBC81E5B374E33B86278">New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="AC25BD0513BD182D4E01569983A4DCEF">Bhunia, Debika</mods:namePart>
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<mods:namePart id="BFB99BF048F1FD36D57A9A5F1E812B7D">Ahrens, Dirk</mods:namePart>
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<mods:namePart id="092DEF36D0DE9A3559B73534CCD1CBF2">Chandra, Kailash</mods:namePart>
<mods:affiliation id="6AD37BF4D875A3913AA8BA3C5D4362DF">Flat No. 620, Tower Ferrara, Mahagun Mezzaria Sector- 78, Noida- 201301, Gautam Budha Nagar, U. P</mods:affiliation>
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<heading id="D0C08C25A078FF97C0AE882FF846C738" bold="true" box="[151,593,1087,1114]" fontSize="11" level="1" pageId="11" pageNumber="382" reason="1">
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<emphasis id="B943E75BA078FF97C0AE882FF846C738" bold="true" box="[151,593,1087,1114]" pageId="11" pageNumber="382">
<emphasis id="B943E75BA078FF97C0AE882FFBB7C73B" bold="true" box="[151,416,1087,1114]" italics="true" pageId="11" pageNumber="382">Maladera schenklingi</emphasis>
(Moser, 1918)
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<emphasis id="B943E75BA078FF97C0AE8897FB92C7C0" bold="true" box="[151,389,1159,1185]" pageId="11" pageNumber="382">Material examined.</emphasis>
<materialsCitation id="3B5F3114A078FF97C1A98897FE73C7A4" collectingDate="2017-07-05" collectionCode="NME" collectorName="A. Weigel" country="Nepal" elevation="180" latitude="27.580002" location="Sauraha" longLatPrecision="20" longitude="84.469444" municipality="Chitwan" pageId="11" pageNumber="382" specimenCount="9" specimenCount-female="7" specimenCount-male="2" stateProvince="Narayani">
<specimenCount id="9D31F0C0A078FF97C1A98897FBC1C7C3" box="[400,470,1159,1186]" count="2" pageId="11" pageNumber="382" type="male">2 ♂♂</specimenCount>
,
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“
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, P:
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, D:
<collectingMunicipality id="6BECA133A078FF97C3638897F9ADC7C0" box="[858,954,1159,1185]" pageId="11" pageNumber="382">Chitwan</collectingMunicipality>
~ 2kmW of
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,
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,
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<collectingDate id="EFCDE461A078FF97C5358897FF8FC7C3" box="[1292,1432,1159,1186]" pageId="11" pageNumber="382" value="2017-07-05">05.VII.2017</collectingDate>
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,
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,
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, #17-22a, leg.
<collectorName id="26C25E9FA078FF97C26988BBF8D3C7A4" box="[592,708,1195,1221]" pageId="11" pageNumber="382">A. Weigel</collectorName>
, small forest with fruit tress” (
<collectionCode id="ED26A38CA078FF97C42188BBFE4BC7A4" box="[1048,1116,1195,1221]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="11" pageNumber="382" type="">NME</collectionCode>
)
</materialsCitation>
.
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“
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, 08.1944, leg.
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8, 1950” (
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8, 1950” (
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8, 1950” (
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“
<collectingCountry id="F3207BD9A078FF97C2108FBFF870C0A8" box="[553,615,943,969]" name="India" pageId="11" pageNumber="382">India</collectingCountry>
, Bhimtal, Eing. Nr. 3/1977, leg.
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),
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“Himalaya, Chakrata,
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17.
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.1956, D.
<collectingCountry id="F3207BD9A078FF97C1838FC3F815C08C" box="[442,514,979,1005]" name="India" pageId="11" pageNumber="382">Indien</collectingCountry>
Exp, Nr 1147, Eing.Nr, 1, 1956” (
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)
</materialsCitation>
.
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110
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, D.
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1300,
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106
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<emphasis id="B943E75BA079FF96C0AE8AFBFB8FC467" bold="true" box="[151,408,1771,1798]" italics="true" pageId="10" pageNumber="381">Maladera marginella</emphasis>
(
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)
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<emphasis id="B943E75BA079FF96C0AE8B23FB97C42C" bold="true" box="[151,384,1843,1869]" pageId="10" pageNumber="381">Material examined.</emphasis>
<materialsCitation id="3B5F3114A079FF96C1BC8B23FF8FC42C" box="[389,1432,1843,1870]" collectionCode="ZMUH" collectorName="K. Benner" pageId="10" pageNumber="381" specimenCount="8" specimenCount-female="6" specimenCount-male="2">
<specimenCount id="9D31F0C0A079FF96C1BC8B23FBD2C42F" box="[389,453,1843,1870]" count="2" pageId="10" pageNumber="381" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D31F0C0A079FF96C1E98B23F819C42F" box="[464,526,1843,1870]" count="6" pageId="10" pageNumber="381" type="female">6 ♀♀</specimenCount>
“Brit. Ost-Indien, Dehradun, 8.1942, leg.
<collectorName id="26C25E9FA079FF96C3DB8B24FE41C42F" box="[994,1110,1844,1870]" pageId="10" pageNumber="381">K. Benner</collectorName>
, Eing. Nr. 8, 1950” (
<collectionCode id="ED26A38CA079FF96C5018B23FF87C42C" box="[1336,1424,1843,1869]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="10" pageNumber="381" type="">ZMUH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B5F3114A079FF96C0AE8B47F90AC410" box="[151,797,1879,1906]" collectionCode="ZMUH" collectorName="K. Benner" pageId="10" pageNumber="381" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D31F0C0A079FF96C0AE8B47FAD6C413" box="[151,193,1879,1906]" count="1" pageId="10" pageNumber="381" type="male">1 ♂</specimenCount>
“8.1945, leg.
<collectorName id="26C25E9FA079FF96C1598B48FBC2C413" box="[352,469,1880,1906]" pageId="10" pageNumber="381">K. Benner</collectorName>
, Eing. Nr. 8, 1950” (
<collectionCode id="ED26A38CA079FF96C2858B47F902C410" box="[700,789,1879,1905]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="10" pageNumber="381" type="">ZMUH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

137
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<emphasis id="B943E75BA079FF96C0AE8C87FB75C3D0" bold="true" box="[151,354,151,178]" italics="true" pageId="10" pageNumber="381">Maladera affinis</emphasis>
(
<bibRefCitation id="EFA646B8A079FF96C1488C87F856C3D0" author="Blanchard, C." box="[369,577,151,177]" pageId="10" pageNumber="381" pagination="1 - 128" refId="ref10257" refString="Blanchard, C. (1850) Ier Famille - Scarabaeidae. In: Milne-Edwards, H., Blanchard, C. E. &amp; Lucas, H. (Eds.), Catalogue de la collection entomologique du Museum d'Histoire Naturelle de Paris. Classe des insectes. Ordre des Coleopteres. Tome I. Premiere livraison. Gide et Baudry, Paris, pp. 1-128." type="book chapter" year="1850">Blanchard, 1850</bibRefCitation>
)
</emphasis>
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<emphasis id="B943E75BA079FF96C0AE8CCFFB6EC398" bold="true" box="[151,377,223,249]" pageId="10" pageNumber="381">Material examined</emphasis>
.
<materialsCitation id="3B5F3114A079FF96C1BE8CCFFF42C398" box="[391,1365,223,250]" collectingDate="1956-06-15" collectionCode="ZMUH" country="India" county="Nr" location="Timli" municipality="Exp" pageId="10" pageNumber="381" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D31F0C0A079FF96C1BE8CCFFBA6C39B" box="[391,433,223,250]" count="1" pageId="10" pageNumber="381" type="male">1 ♂</specimenCount>
“
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,
<location id="8EE86D92A079FF96C2298CCFF87FC398" LSID="urn:lsid:plazi:treatment:039E8A5FA079FF96C0AE8C87FB14C294:8EE86D92A079FF96C2298CCFF87FC398" box="[528,616,223,249]" country="India" county="Nr" municipality="Exp" name="Siwalik" pageId="10" pageNumber="381">Siwalik</location>
,
<date id="FF891D89A079FF96C24D8CCFF8E1C39B" box="[628,758,223,250]" pageId="10" pageNumber="381" value="1956-06-15">
<collectingDate id="EFCDE461A079FF96C24D8CCFF8E1C39B" box="[628,758,223,250]" pageId="10" pageNumber="381" value="1956-06-15">15.VI.1956</collectingDate>
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, D.
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,
<collectingCounty id="62E943C5A079FF96C3948CF0F9DCC39B" box="[941,971,224,250]" pageId="10" pageNumber="381">Nr</collectingCounty>
1284, Eing. Nr, 1, 1956” (
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)
</materialsCitation>
.
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<taxonomicName id="4C3740CAA079FF96C0AE8D7DF82CC2EB" authority="(Moser, 1915)" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[151,571,365,394]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="B943E75BA079FF96C0AE8D7DF82CC2EB" bold="true" box="[151,571,365,394]" pageId="10" pageNumber="381">
<emphasis id="B943E75BA079FF96C0AE8D7DFB6DC2EB" bold="true" box="[151,378,365,394]" italics="true" pageId="10" pageNumber="381">Maladera basalis</emphasis>
(Moser, 1915)
</emphasis>
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</paragraph>
<paragraph id="8B883B49A079FF96C0AE8DA7FB14C294" blockId="10.[151,1436,439,501]" pageId="10" pageNumber="381">
<emphasis id="B943E75BA079FF96C0AE8DA7FB91C2B0" bold="true" box="[151,390,439,465]" pageId="10" pageNumber="381">Material examined.</emphasis>
<materialsCitation id="3B5F3114A079FF96C1AA8DA7FAE9C294" collectingDate="1955-11-02" collectionCode="ZMUH" country="India" location="Fanvelbei" pageId="10" pageNumber="381" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D31F0C0A079FF96C1AA8DA7FBD2C2B3" box="[403,453,439,466]" count="1" pageId="10" pageNumber="381" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D31F0C0A079FF96C1EF8DA7F811C2B3" box="[470,518,439,466]" count="1" pageId="10" pageNumber="381" type="female">1 ♀</specimenCount>
“
<location id="8EE86D92A079FF96C2248DA7F886C2B0" LSID="urn:lsid:plazi:treatment:039E8A5FA079FF96C0AE8C87FB14C294:8EE86D92A079FF96C2248DA7F886C2B0" box="[541,657,439,465]" country="India" name="Fanvelbei" pageId="10" pageNumber="381">Fanvelbei</location>
,
<collectingCountry id="F3207BD9A079FF96C2988DA7F916C2B0" box="[673,769,439,465]" name="India" pageId="10" pageNumber="381">Bombay</collectingCountry>
,
<date id="FF891D89A079FF96C32B8DA7F983C2B3" box="[786,916,439,466]" pageId="10" pageNumber="381" value="1955-11-02">
<collectingDate id="EFCDE461A079FF96C32B8DA7F983C2B3" box="[786,916,439,466]" pageId="10" pageNumber="381" value="1955-11-02">02.XI.1955</collectingDate>
</date>
, D.
<collectingCountry id="F3207BD9A079FF96C3F48DA7FE02C2B0" box="[973,1045,439,465]" name="India" pageId="10" pageNumber="381">Indien</collectingCountry>
Exp.Nr 533, Eing. Nr, 1, 1956” (
<collectionCode id="ED26A38CA079FF96C0A68DCBFAE4C294" box="[159,243,475,501]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="10" pageNumber="381" type="">ZMUH</collectionCode>
)
</materialsCitation>
.
</paragraph>
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</treatment>
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127
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.1956, D.
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This is the first record of this species for Goa state of
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<materialsCitation id="3B5F3114A07AFF95C1B58AB3FE07C580" collectingDate="2022-06-20" collectionCode="NME" collectorName="A. Weigel" country="Nepal" elevation="2650" latitude="29.830278" location="Lumcha" longLatPrecision="20" longitude="81.9725" municipality="Humla Simlikot" pageId="9" pageNumber="380" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Karnali">
<specimenCount id="9D31F0C0A07AFF95C1B58AB3FBADC5DC" box="[396,442,1699,1725]" count="1" pageId="9" pageNumber="380" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D31F0C0A07AFF95C1F08AB3FBE2C5DC" box="[457,501,1699,1725]" count="1" pageId="9" pageNumber="380" type="female">1 ♀</specimenCount>
“
<collectingCountry id="F3207BD9A07AFF95C2318AB3F847C5DC" box="[520,592,1699,1725]" name="Nepal" pageId="9" pageNumber="380">Nepal</collectingCountry>
, P:
<collectingRegion id="49F3F5ABA07AFF95C2478AB3F8C2C5DC" box="[638,725,1699,1725]" country="Nepal" name="Karnali" pageId="9" pageNumber="380">Karnali</collectingRegion>
/D:
<collectingMunicipality id="6BECA133A07AFF95C2C68AB3F9A2C5DC" box="[767,949,1699,1725]" pageId="9" pageNumber="380">Humla Simlikot</collectingMunicipality>
~
<quantity id="4CCF96ACA07AFF95C3EE8AB3FE00C5DC" box="[983,1047,1699,1725]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="380" unit="km" value="21.0">21km</quantity>
SE, Rodikot, S,
<location id="8EE86D92A07AFF95C4E28AB3FF2DC5DC" LSID="urn:lsid:plazi:treatment:039E8A5FA07AFF95C0AE8A4BFB45C448:8EE86D92A07AFF95C4E28AB3FF2DC5DC" box="[1243,1338,1699,1725]" country="Nepal" latitude="29.830278" longLatPrecision="20" longitude="81.9725" municipality="Humla Simlikot" name="Lumcha" pageId="9" pageNumber="380" stateProvince="Karnali">Lumcha</location>
,
<quantity id="4CCF96ACA07AFF95C57E8AB2FF8FC5DC" box="[1351,1432,1698,1725]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="9" pageNumber="380" unit="m" value="2650.0">
<elevation id="001ADC7AA07AFF95C57E8AB2FF8FC5DC" box="[1351,1432,1698,1725]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="9" pageNumber="380" unit="m" value="2650.0">2650m</elevation>
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,
<date id="FF891D89A07AFF95C0AE8AD7FB0EC580" box="[151,281,1735,1761]" pageId="9" pageNumber="380" value="2022-06-20">
<collectingDate id="EFCDE461A07AFF95C0AE8AD7FB0EC580" box="[151,281,1735,1761]" pageId="9" pageNumber="380" value="2022-06-20">20.VI.2022</collectingDate>
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,
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,
<geoCoordinate id="EE035D8EA07AFF95C1828AD6F829C580" box="[443,574,1734,1761]" degrees="81" direction="east" minutes="58" orientation="longitude" pageId="9" pageNumber="380" precision="15" seconds="21" value="81.9725">81°58'21&quot;E</geoCoordinate>
, leg.
<collectorName id="26C25E9FA07AFF95C2448AD7F8E4C580" box="[637,755,1735,1761]" pageId="9" pageNumber="380">A. Weigel</collectorName>
, HF/KL 22-#11” (
<collectionCode id="ED26A38CA07AFF95C3FD8AD7FE1FC580" box="[964,1032,1735,1761]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="9" pageNumber="380" type="">NME</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B5F3114A07AFF95C4278AD7FB59C448" collectingDate="2022-06-30" collectionCode="NME" collectorName="A. Weigel" country="Nepal" county="Jumla Jumla" elevation="2780" latitude="29.395277" location="Bumra" longLatPrecision="20" longitude="82.148605" municipality="Buvramatichaur" pageId="9" pageNumber="380" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Karnali">
<specimenCount id="9D31F0C0A07AFF95C4278AD7FE76C580" box="[1054,1121,1735,1761]" count="2" pageId="9" pageNumber="380" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D31F0C0A07AFF95C4568AD7FE8CC580" box="[1135,1179,1735,1761]" count="1" pageId="9" pageNumber="380" type="female">1 ♀</specimenCount>
“
<collectingCountry id="F3207BD9A07AFF95C4968AD7FEE0C580" box="[1199,1271,1735,1761]" name="Nepal" pageId="9" pageNumber="380">Nepal</collectingCountry>
, P:
<collectingRegion id="49F3F5ABA07AFF95C51C8AD7FF6BC580" box="[1317,1404,1735,1761]" country="Nepal" name="Karnali" pageId="9" pageNumber="380">Karnali</collectingRegion>
/D:
<collectingCounty id="62E943C5A07AFF95C0AE8AFBFB31C464" box="[151,294,1771,1797]" pageId="9" pageNumber="380">Jumla Jumla</collectingCounty>
~
<quantity id="4CCF96ACA07AFF95C17E8AFBFB90C464" box="[327,391,1771,1797]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="380" unit="km" value="14.0">14km</quantity>
N,
<collectingMunicipality id="6BECA133A07AFF95C18D8AFBF87FC464" box="[436,616,1771,1797]" pageId="9" pageNumber="380">Buvramatichaur</collectingMunicipality>
(
<location id="8EE86D92A07AFF95C2408AFBF8DCC464" LSID="urn:lsid:plazi:treatment:039E8A5FA07AFF95C0AE8A4BFB45C448:8EE86D92A07AFF95C2408AFBF8DCC464" box="[633,715,1771,1797]" country="Nepal" county="Jumla Jumla" latitude="29.395277" longLatPrecision="20" longitude="82.148605" municipality="Buvramatichaur" name="Bumra" pageId="9" pageNumber="380" stateProvince="Karnali">Bumra</location>
),
<quantity id="4CCF96ACA07AFF95C2D98AFBF926C464" box="[736,817,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="2.78" pageId="9" pageNumber="380" unit="m" value="2780.0">
<elevation id="001ADC7AA07AFF95C2D98AFBF926C464" box="[736,817,1771,1797]" metricMagnitude="3" metricUnit="m" metricValue="2.78" pageId="9" pageNumber="380" unit="m" value="2780.0">2780m</elevation>
</quantity>
,
<geoCoordinate id="EE035D8EA07AFF95C3078AFBF9D3C464" box="[830,964,1771,1797]" degrees="29" direction="north" minutes="23" orientation="latitude" pageId="9" pageNumber="380" precision="15" seconds="43" value="29.395277">29°23'43&quot;N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA07AFF95C3EB8AFAFE42C464" box="[978,1109,1770,1797]" degrees="82" direction="east" minutes="08" orientation="longitude" pageId="9" pageNumber="380" precision="15" seconds="55" value="82.148605">82°08'55&quot;E</geoCoordinate>
,
<date id="FF891D89A07AFF95C45A8AFBFEF2C464" box="[1123,1253,1771,1797]" pageId="9" pageNumber="380" value="2022-06-30">
<collectingDate id="EFCDE461A07AFF95C45A8AFBFEF2C464" box="[1123,1253,1771,1797]" pageId="9" pageNumber="380" value="2022-06-30">30.VI.2022</collectingDate>
</date>
, leg.
<collectorName id="26C25E9FA07AFF95C51A8AFBFF8EC464" box="[1315,1433,1771,1797]" pageId="9" pageNumber="380">A. Weigel</collectorName>
, #22-21” (
<collectionCode id="ED26A38CA07AFF95C13B8B1FFB51C448" box="[258,326,1807,1833]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="9" pageNumber="380" type="">NME</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

119
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<mods:title id="AD7B5B10BA00D4E1BD85A9D1EDE12AE5">New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="5B09A98BF876A112A9D761BD6A7F0015">Ahrens, Dirk</mods:namePart>
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<heading id="D0C08C25A07AFF95C0AE8B6BF8CCC4F5" bold="true" box="[151,731,1914,1941]" fontSize="11" level="1" pageId="9" pageNumber="380" reason="1">
<taxonomicName id="4C3740CAA07AFF95C0AE8B6BF8CCC4F5" ID-CoL="7ZBZW" authority="(Brenske, 1896)" baseAuthorityName="Brenske" baseAuthorityYear="1896" box="[151,731,1914,1941]" class="Insecta" family="Scarabaeidae" genus="Lasioserica" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="subSpecies" species="maculata" subSpecies="maculata">
<emphasis id="B943E75BA07AFF95C0AE8B6BF8CCC4F5" bold="true" box="[151,731,1914,1941]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0AE8B6BF806C4F4" bold="true" box="[151,529,1914,1941]" italics="true" pageId="9" pageNumber="380">Lasioserica maculata maculata</emphasis>
(
<bibRefCitation id="EFA646B8A07AFF95C2188B6AF8C4C4F5" author="Brenske, E." box="[545,723,1914,1940]" pageId="9" pageNumber="380" pagination="150 - 164" refId="ref10330" refString="Brenske, E. (1896) Insectes du Bengale. Melolonthidae. Annales de la Societe Entomologique de Belgique, 40, 150-164." type="journal article" year="1896">Brenske, 1896</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A07AFF95C0AE8BACF93DC49B" pageId="9" pageNumber="380" type="materials_examined">
<paragraph id="8B883B49A07AFF95C0AE8BACF93DC49B" blockId="9.[151,1437,1980,2043]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0AE8BACFB96C4B7" bold="true" box="[151,385,1980,2006]" pageId="9" pageNumber="380">Material examined.</emphasis>
<materialsCitation id="3B5F3114A07AFF95C1B18BACF931C49B" collectingDate="2022-06-16" collectionCode="NME" collectorName="A. Weigel" country="Nepal" elevation="2250" latitude="29.962223" location="Humla" longLatPrecision="20" longitude="81.83917" municipality="Humla Simlikot" pageId="9" pageNumber="380" specimenCount="1" specimenCount-female="1" stateProvince="Karnali">
<specimenCount id="9D31F0C0A07AFF95C1B18BACFBA4C4B6" box="[392,435,1980,2007]" count="1" pageId="9" pageNumber="380" type="female">1 ♀</specimenCount>
“
<collectingCountry id="F3207BD9A07AFF95C1FC8BACF81AC4B7" box="[453,525,1980,2006]" name="Nepal" pageId="9" pageNumber="380">Nepal</collectingCountry>
, P:
<collectingRegion id="49F3F5ABA07AFF95C20E8BACF899C4B7" box="[567,654,1980,2006]" country="Nepal" name="Karnali" pageId="9" pageNumber="380">Karnali</collectingRegion>
, D:
<collectingMunicipality id="6BECA133A07AFF95C2848BACF966C4B7" box="[701,881,1980,2006]" pageId="9" pageNumber="380">Humla Simlikot</collectingMunicipality>
~
<quantity id="4CCF96ACA07AFF95C3B68BACF9D6C4B7" box="[911,961,1980,2006]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="380" unit="km" value="2.0">2km</quantity>
SE,
<location id="8EE86D92A07AFF95C3C18BACFE50C4B7" LSID="urn:lsid:plazi:treatment:039E8A5FA07AFF95C0AE8B6BF93DC49B:8EE86D92A07AFF95C3C18BACFE50C4B7" box="[1016,1095,1980,2006]" country="Nepal" latitude="29.962223" longLatPrecision="20" longitude="81.83917" municipality="Humla Simlikot" name="Humla" pageId="9" pageNumber="380" stateProvince="Karnali">Humla</location>
,
<collectingRegion id="49F3F5ABA07AFF95C46A8BACFEBDC4B7" box="[1107,1194,1980,2006]" country="Nepal" name="Karnali" pageId="9" pageNumber="380">Karnali</collectingRegion>
,
<quantity id="4CCF96ACA07AFF95C48C8BACFF11C4B6" box="[1205,1286,1980,2007]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="9" pageNumber="380" unit="m" value="2250.0">
<elevation id="001ADC7AA07AFF95C48C8BACFF11C4B6" box="[1205,1286,1980,2007]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="9" pageNumber="380" unit="m" value="2250.0">2250m</elevation>
</quantity>
,
<geoCoordinate id="EE035D8EA07AFF95C5288BACFF8FC4B6" box="[1297,1432,1980,2007]" degrees="29" direction="north" minutes="57" orientation="latitude" pageId="9" pageNumber="380" precision="15" seconds="44" value="29.962223">29°57'44&quot;N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA07AFF95C0AE8BF0FB0DC49A" box="[151,282,2016,2043]" degrees="81" direction="east" minutes="50" orientation="longitude" pageId="9" pageNumber="380" precision="15" seconds="21" value="81.83917">81°50'21&quot;E</geoCoordinate>
,
<date id="FF891D89A07AFF95C11F8BF0FBBFC49B" box="[294,424,2016,2042]" pageId="9" pageNumber="380" value="2022-06-16">
<collectingDate id="EFCDE461A07AFF95C11F8BF0FBBFC49B" box="[294,424,2016,2042]" pageId="9" pageNumber="380" value="2022-06-16">16.VI.2022</collectingDate>
</date>
, leg.
<collectorName id="26C25E9FA07AFF95C1DB8BF0F841C49B" box="[482,598,2016,2042]" pageId="9" pageNumber="380">A. Weigel</collectorName>
, 22-#04b” (
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

267
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<document id="AD30B2BBFCC2B7247514DC3E4B19C026" ID-DOI="10.11646/zootaxa.5613.2.11" ID-ISSN="1175-5326" ID-ZooBank="2972195A-6FD5-488A-8925-51D24677DFEC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1744667883477" checkinUser="plazi" docAuthor="Gupta, Devanshu, Bhunia, Debika, Ahrens, Dirk &amp; Chandra, Kailash" docDate="2025" docId="039E8A5FA07BFF95C0AE894EFF20C6A0" docLanguage="en" docName="zootaxa.5613.2.11.pdf" docOrigin="Zootaxa 5613 (2)" docSource="https://doi.org/10.11646/zootaxa.5613.2.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="380" masterDocId="FFA7F227A073FF9CC0398C10FA17C361" masterDocTitle="New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)" masterLastPageNumber="385" masterPageNumber="371" pageNumber="379" updateTime="1744670091748" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="CD700B9CB6FA40DD43233F823221A9E1">New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="DFFCC81E334B38E56E686F5AABDC191B">Gupta, Devanshu</mods:namePart>
<mods:affiliation id="AE6A1A68D894822D813D602EE73C5FE9">Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India</mods:affiliation>
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<mods:namePart id="2ECAD9B29DAC1153EADE1878A7B200BD">Bhunia, Debika</mods:namePart>
<mods:affiliation id="6524A87DE1089E042FCE0DC5BF6DC7F1">Zoological Survey of India, M-Block, New Alipore, Kolkata- 700053, West Bengal, India</mods:affiliation>
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<mods:namePart id="2186E254737F3A8CE27160E86095B20A">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="AE14A8698603B8C5D4D137488BAC37A1" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="DA9CCAD3BCFD6E0468AB1C96FE507E84" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:namePart id="DEB54C5A92CE88204C80FF80788490DD">Chandra, Kailash</mods:namePart>
<mods:affiliation id="821D5337E9FE816E15F9787928686A23">Flat No. 620, Tower Ferrara, Mahagun Mezzaria Sector- 78, Noida- 201301, Gautam Budha Nagar, U. P</mods:affiliation>
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<mods:date id="80FA437D104D0AED26FD456AA9FCF83B">2025</mods:date>
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<mods:number id="7D139F6804FC6659A7FDABD08D8E866F">2025-03-27</mods:number>
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<heading id="D0C08C25A07BFF94C0AE894EF992C618" bold="true" box="[151,901,1374,1401]" fontSize="11" level="1" pageId="8" pageNumber="379" reason="1">
<emphasis id="B943E75BA07BFF94C0AE894EF992C618" bold="true" box="[151,901,1374,1401]" pageId="8" pageNumber="379">
<taxonomicName id="4C3740CAA07BFF94C0AE894EF90DC619" ID-CoL="KTVX7" authority="Gupta, Bhunia, Ahrens &amp; Chandra" authorityName="Gupta, Bhunia, Ahrens &amp; Chandra" authorityYear="2025" box="[151,794,1374,1401]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="onam" status="sp. nov.">
<emphasis id="B943E75BA07BFF94C0AE894EFB4FC618" bold="true" box="[151,344,1374,1401]" italics="true" pageId="8" pageNumber="379">Maladera onam</emphasis>
Gupta, Bhunia, Ahrens &amp; Chandra
</taxonomicName>
,
<taxonomicNameLabel id="A2705A20A07BFF94C311894FF992C618" box="[808,901,1375,1401]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C32D68C2A07BFF94C0AE8993FB04C6FC" box="[151,275,1411,1437]" pageId="8" pageNumber="379" type="description">
<paragraph id="8B883B49A07BFF94C0AE8993FB04C6FC" blockId="8.[151,901,1374,1437]" box="[151,275,1411,1437]" pageId="8" pageNumber="379">
(
<figureCitation id="130C27CCA07BFF94C0A68993FB1DC6FC" box="[159,266,1411,1437]" captionStart="FIGURE 2" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[183,1413,188,1870]" captionTargetId="figure-22@7.[175,1413,179,1882]" captionTargetPageId="7" captionText="FIGURE 2. A–D. Maladera lumlaensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Serica subansiriensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="8" pageNumber="379">Fig. 2I–L</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A07BFF94C0AE89DBFAF4C514" pageId="8" pageNumber="379" type="materials_examined">
<paragraph id="8B883B49A07BFF94C0AE89DBF9FCC684" blockId="8.[151,1437,1482,2013]" box="[151,1003,1482,1509]" pageId="8" pageNumber="379">
<materialsCitation id="3B5F3114A07BFF94C0AE89DBF9F0C684" box="[151,999,1482,1509]" country="India" latitude="12.15" location="Silent valley" longLatPrecision="558" longitude="75.142" municipality="Valiyaparamba" pageId="8" pageNumber="379" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<emphasis id="B943E75BA07BFF94C0AE89DBFB21C685" bold="true" box="[151,310,1482,1509]" pageId="8" pageNumber="379">
<typeStatus id="548C85EBA07BFF94C0AE89DBFAC5C684" box="[151,210,1483,1509]" pageId="8" pageNumber="379">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3207BD9A07BFF94C10589DBFB6FC684" box="[316,376,1483,1509]" name="India" pageId="8" pageNumber="379">India</collectingCountry>
,
<collectingRegion id="49F3F5ABA07BFF94C1BD89DBFBC6C684" box="[388,465,1483,1509]" country="India" name="Kerala" pageId="8" pageNumber="379">Kerala</collectingRegion>
,
<collectingMunicipality id="6BECA133A07BFF94C1EC89DBF897C684" box="[469,640,1483,1509]" pageId="8" pageNumber="379">Valiyaparamba</collectingMunicipality>
,
<location id="8EE86D92A07BFF94C2B289DBF90EC684" LSID="urn:lsid:plazi:treatment:039E8A5FA07BFF95C0AE894EFF20C6A0:8EE86D92A07BFF94C2B289DBF90EC684" box="[651,793,1483,1509]" country="India" latitude="12.15" longLatPrecision="558" longitude="75.142" municipality="Valiyaparamba" name="Silent valley" pageId="8" pageNumber="379" stateProvince="Kerala">Silent valley</location>
,
<geoCoordinate id="EE035D8EA07BFF94C31D89DAF96CC684" box="[804,891,1482,1509]" degrees="12.15" direction="north" orientation="latitude" pageId="8" pageNumber="379" precision="555" value="12.15">12.15N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA07BFF94C3BF89DAF9F0C684" box="[902,999,1482,1509]" degrees="75.142" direction="east" orientation="longitude" pageId="8" pageNumber="379" precision="55" value="75.142">75.142E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8B883B49A07BFF94C0FE89FFFAF4C514" blockId="8.[151,1437,1482,2013]" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE89FFFB62C569" bold="true" box="[199,373,1518,1545]" pageId="8" pageNumber="379">Type material.</emphasis>
<materialsCitation id="3B5F3114A07BFF94C14489FFFAF4C514" collectingDate="1980-01-23" collectionCode="NZCI" collectorName="S. K. Bhattacharya &amp; The" country="India" latitude="12.15" location="Silent Valley" longLatPrecision="77" longitude="75.142" municipality="Valiyaparamba" pageId="8" pageNumber="379" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="548C85EBA07BFF94C14489FFFBFFC568" box="[381,488,1519,1545]" pageId="8" pageNumber="379" type="holotype">Holotype</typeStatus>
,
<emphasis id="B943E75BA07BFF94C1CA89E0F81FC56D" box="[499,520,1520,1548]" italics="true" pageId="8" pageNumber="379">
<specimenCount id="9D31F0C0A07BFF94C1CA89E0F81FC56D" box="[499,520,1520,1548]" count="1" pageId="8" pageNumber="379" type="male">♂</specimenCount>
</emphasis>
: “
<collectingCountry id="F3207BD9A07BFF94C21B89FFF877C568" box="[546,608,1519,1545]" name="India" pageId="8" pageNumber="379">India</collectingCountry>
,
<collectingRegion id="49F3F5ABA07BFF94C25589FFF8AEC568" box="[620,697,1519,1545]" country="India" name="Kerala" pageId="8" pageNumber="379">Kerala</collectingRegion>
,
<collectingMunicipality id="6BECA133A07BFF94C2FD89FFF978C568" box="[708,879,1519,1545]" pageId="8" pageNumber="379">Valiyaparamba</collectingMunicipality>
,
<location id="8EE86D92A07BFF94C34289FFFE1BC568" LSID="urn:lsid:plazi:treatment:039E8A5FA07BFF95C0AE894EFF20C6A0:8EE86D92A07BFF94C34289FFFE1BC568" box="[891,1036,1519,1545]" country="India" latitude="12.15" longLatPrecision="77" longitude="75.142" municipality="Valiyaparamba" name="Silent Valley" pageId="8" pageNumber="379" stateProvince="Kerala">Silent Valley</location>
,
<geoCoordinate id="EE035D8EA07BFF94C42E89FEFE94C568" box="[1047,1155,1518,1545]" degrees="12.150" direction="north" orientation="latitude" pageId="8" pageNumber="379" precision="55" value="12.15">12.150 N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA07BFF94C4B789FEFEF8C568" box="[1166,1263,1518,1545]" degrees="75.142" direction="east" orientation="longitude" pageId="8" pageNumber="379" precision="55" value="75.142">75.142E</geoCoordinate>
,
<date id="FF891D89A07BFF94C4C289FFFF70C568" box="[1275,1383,1519,1545]" pageId="8" pageNumber="379" value="1980-01-23">
<collectingDate id="EFCDE461A07BFF94C4C289FFFF70C568" box="[1275,1383,1519,1545]" pageId="8" pageNumber="379" value="1980-01-23">23.i.1980</collectingDate>
</date>
, leg.
<collectorName id="26C25E9FA07BFF94C0AE8A03FB70C54C" box="[151,359,1555,1581]" pageId="8" pageNumber="379">S.K. Bhattacharya</collectorName>
” (
<collectionCode id="ED26A38CA07BFF94C1B88A03FBDEC54C" box="[385,457,1555,1581]" pageId="8" pageNumber="379">NZCI</collectionCode>
).
<collectorName id="26C25E9FA07BFF94C1E58A03F81FC54C" box="[476,520,1555,1581]" pageId="8" pageNumber="379">The</collectorName>
<typeStatus id="548C85EBA07BFF94C2368A03F866C54C" box="[527,625,1555,1581]" pageId="8" pageNumber="379" type="holotype">holotype</typeStatus>
specimen bears additionally a red printed label with the following text: “
<taxonomicName id="4C3740CAA07BFF94C09D8A27FB4FC531" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[164,344,1591,1617]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="onam" status="sp. nov.">
<emphasis id="B943E75BA07BFF94C09D8A27FB4FC531" box="[164,344,1591,1617]" italics="true" pageId="8" pageNumber="379">Maladera onam</emphasis>
</taxonomicName>
<emphasis id="B943E75BA07BFF94C1598A27FBACC530" bold="true" box="[352,443,1591,1617]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A2705A20A07BFF94C1598A27FBACC530" box="[352,443,1591,1617]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="548C85EBA07BFF94C1F38A27F876C530" box="[458,609,1591,1617]" pageId="8" pageNumber="379" type="holotype">HOLOTYPE</typeStatus>
, Devanshu Gupta, Debika Bhunia, Dirk Ahrens &amp; Kailash Chandra, Det. 2024”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A07BFF95C0FE8A6EFB82C788" lastPageId="9" lastPageNumber="380" pageId="8" pageNumber="379" type="description">
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<emphasis id="B943E75BA07BFF94C0FE8A6EF806C5F8" bold="true" box="[199,529,1662,1689]" pageId="8" pageNumber="379">
Description of the
<typeStatus id="548C85EBA07BFF94C19A8A6FF81BC5F8" box="[419,524,1663,1689]" pageId="8" pageNumber="379" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CCF96ACA07BFF94C24F8A6EF8D9C5F8" box="[630,718,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="8" pageNumber="379" unit="mm" value="9.5">9.5 mm</quantity>
, elytral length:
<quantity id="4CCF96ACA07BFF94C3448A6FF9C2C5F8" box="[893,981,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="8" pageNumber="379" unit="mm" value="7.3">7.3 mm</quantity>
, width:
<quantity id="4CCF96ACA07BFF94C4168A6FFE9FC5F8" box="[1071,1160,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="8" pageNumber="379" unit="mm" value="6.7">6.7 mm</quantity>
.
</paragraph>
<paragraph id="8B883B49A07BFF94C0FE8AB3FBD7C580" blockId="8.[151,1437,1482,2013]" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE8AB3FB12C5DC" bold="true" box="[199,261,1699,1725]" pageId="8" pageNumber="379">Body</emphasis>
oval, uniformly dark reddish brown, dorsal and ventral face dull, except lateral setae of elytra and pronotum nearly glabrous.
</paragraph>
<paragraph id="8B883B49A07BFF95C0FE8AFBFBC7C3B4" blockId="8.[151,1437,1482,2013]" lastBlockId="9.[151,1437,151,1473]" lastPageId="9" lastPageNumber="380" pageId="8" pageNumber="379">
<emphasis id="B943E75BA07BFF94C0FE8AFBFB19C464" bold="true" box="[199,270,1771,1797]" pageId="8" pageNumber="379">Head.</emphasis>
Labroclypeus wide, trapezoidal, widest at base and moderately shiny, lateral margins weakly convex and convergent anteriorly, producing a distinct blunt angle with ocular canthus, not incised before labrum, anterior angles moderately convex, anterior margin weakly emarginate medially, margins weakly reflexed; surface convex, finely and densely punctate, with a few fine erect setae behind anterior margin. Frontoclypeal suture finely incised, weakly curved. Smooth area in front of eyes 1.5 times as wide as long; ocular canthus short and wide, finely and densely punctate, with a short terminal seta. Frons finely and densely punctate, with a few single and short setae beside eyes. Eyes moderately large, ratio diameter/ interocular width: 0.67. Antenna with ten antennomeres, club with three antennomeres, slightly shorter than remaining antennomeres combined. Mentum convexly elevated, anteriorly slightly flattened.
</paragraph>
<paragraph id="8B883B49A07AFF95C0FE8CCFFB5AC2E8" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE8CCFFB57C398" bold="true" box="[199,320,223,249]" pageId="9" pageNumber="380">Pronotum</emphasis>
widest at base, lateral margins evenly and convexly convergent anteriorly, anterior angles moderately sharp, distinctly produced, posterior angles blunt, slightly rounded at tip; anterior margin with complete marginal line, straight; lateral and lateral anterior margin with long and fine setae; surface finely and densely punctate, with microscopic setae in punctures, otherwise glabrous. Scutellum wide, triangular, punctures fine and moderately dense, glabrous.
</paragraph>
<paragraph id="8B883B49A07AFF95C0FE8D83FF4FC178" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE8D83FB04C2CC" bold="true" box="[199,275,403,429]" pageId="9" pageNumber="380">Elytra</emphasis>
wide, widest at middle, external apical angle strongly rounded, striae finely impressed, finely and densely punctate, even intervals nearly flat, finely and moderately densely punctate, odd intervals slightly convex, with punctures concentrated along striae, with only microscopic setae in punctures; epipleural edge ending at external apical angle of elytra; epipleura with long and sparse setae; apical margin with a rim of fine microtrichomes.
</paragraph>
<paragraph id="8B883B49A07AFF95C0FE8E33F97AC190" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE8E33FB97C15D" bold="true" box="[199,384,546,573]" pageId="9" pageNumber="380">Ventral surface</emphasis>
coarsely and densely punctate, with microscopic setae in punctures, with a few longer setae on mesosternum and metasternal plate. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.78. Metacoxa glabrous, laterally with a few robust setae.Abdominal sternites finely and moderately densely punctate, with a transverse row of coarse punctures each bearing a robust seta, penultimate sternite with a very narrow shiny chitinous rim. Pygidium dull, weakly convex, finely and moderately densely punctate, with narrow impunctate median line, glabrous.
</paragraph>
<paragraph id="8B883B49A07AFF95C0FE8EEBF9F7C7C0" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE8EEBFAE8C074" bold="true" box="[199,255,763,789]" pageId="9" pageNumber="380">Legs</emphasis>
moderately wide; femora with two longitudinal rows of setae. Metafemur dull, superficially punctate, anterior edge acute, without adjacent serrated line, anterior and posterior row of setae absent; posterior ventral marginal most straight, strongly widened in apical half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia wide and short, widest at middle, ratio width/length: 1/2.3, sharply carinate dorsally, with two groups of spines, basal one at middle, apical one at three quarters of metatibial length, in basal third with 3–4 coarse punctures each bearing a fine seta; lateral face weakly longitudinally convex, impunctate, only ventral part with moderately dense and coarse punctures; ventral margin with four equidistant spines; medial face impunctate and glabrous, apex shallowly emarginate interiorly near tarsal articulation. Tarsomeres impunctate dorsally, circular in cross section, with sparse, fine setae ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate, teeth moderately large. All claws symmetrical, feebly curved and long, with normally developed basal tooth.
</paragraph>
<paragraph id="8B883B49A07AFF95C0FE88BAF89CC7A4" blockId="9.[151,1437,151,1473]" box="[199,651,1194,1221]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE88BAFB2DC7A5" bold="true" box="[199,314,1194,1220]" pageId="9" pageNumber="380">Aedeagus</emphasis>
:
<figureCitation id="130C27CCA07AFF95C17088BBFBA1C7A4" box="[329,438,1195,1221]" captionStart="FIGURE 2" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[183,1413,188,1870]" captionTargetId="figure-22@7.[175,1413,179,1882]" captionTargetPageId="7" captionText="FIGURE 2. A–D. Maladera lumlaensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Serica subansiriensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="9" pageNumber="380">Fig. 2I–K</figureCitation>
.
<emphasis id="B943E75BA07AFF95C1FD88BAF834C7A5" bold="true" box="[452,547,1194,1220]" pageId="9" pageNumber="380">Habitus</emphasis>
:
<figureCitation id="130C27CCA07AFF95C20B88BBF893C7A4" box="[562,644,1195,1221]" captionStart="FIGURE 2" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[183,1413,188,1870]" captionTargetId="figure-22@7.[175,1413,179,1882]" captionTargetPageId="7" captionText="FIGURE 2. A–D. Maladera lumlaensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; E–H. Serica subansiriensis Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.; I–L. Maladera onam Gupta, Bhunia, Ahrens &amp; Chandra sp. nov.. (A, E, I) Aedeagus in lateral view (left); (B, F, J) Aedeagus in dorsal view; (C, H, L) Aedeagus in lateral view (right); (D, H, L) Habitus, dorsal view. Scale 0.5 mm; habitus not to scale." pageId="9" pageNumber="380">
Fig.
<specimenCount id="9D31F0C0A07AFF95C25C88BBF893C7A4" box="[613,644,1195,1221]" count="2" pageId="9" pageNumber="380" type="generic">2L</specimenCount>
</figureCitation>
.
</paragraph>
<paragraph id="8B883B49A07AFF95C0FE88DFFB82C788" blockId="9.[151,1437,151,1473]" box="[199,405,1231,1257]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE88DFFB09C788" bold="true" box="[199,286,1231,1257]" pageId="9" pageNumber="380">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A07AFF95C0FE88E2F95FC66C" box="[199,840,1266,1293]" pageId="9" pageNumber="380" type="distribution">
<paragraph id="8B883B49A07AFF95C0FE88E2F95FC66C" blockId="9.[151,1437,151,1473]" box="[199,840,1266,1293]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE88E2FB76C66D" bold="true" box="[199,353,1266,1292]" pageId="9" pageNumber="380">Distribution.</emphasis>
Species only known from the
<typeStatus id="548C85EBA07AFF95C28E88E3F8F0C66C" box="[695,743,1267,1293]" pageId="9" pageNumber="380">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A07AFF95C0FE8906F9DAC6FC" pageId="9" pageNumber="380" type="diagnosis">
<paragraph id="8B883B49A07AFF95C0FE8906F9DAC6FC" blockId="9.[151,1437,151,1473]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE8906FB2DC651" bold="true" box="[199,314,1302,1328]" pageId="9" pageNumber="380">Diagnosis</emphasis>
.
<taxonomicName id="4C3740CAA07AFF95C1718907FBEBC651" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[328,508,1303,1329]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="onam" status="sp. nov.">
<emphasis id="B943E75BA07AFF95C1718907FBEBC651" box="[328,508,1303,1329]" italics="true" pageId="9" pageNumber="380">Maladera onam</emphasis>
</taxonomicName>
<emphasis id="B943E75BA07AFF95C23A8907F84AC650" bold="true" box="[515,605,1303,1329]" pageId="9" pageNumber="380">
<taxonomicNameLabel id="A2705A20A07AFF95C23A8907F84AC650" box="[515,605,1303,1329]" pageId="9" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is in its external characters and in the shape of the aedeagus rather similar to
<taxonomicName id="4C3740CAA07AFF95C08F892BF8C1C634" authority="Ahrens &amp; Fabrizi, 2016" authorityName="Ahrens &amp; Fabrizi" authorityYear="2016" box="[182,726,1339,1365]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="trochaloides">
<emphasis id="B943E75BA07AFF95C08F892BFBAEC634" box="[182,441,1339,1365]" italics="true" pageId="9" pageNumber="380">Maladera trochaloides</emphasis>
<bibRefCitation id="EFA646B8A07AFF95C1F8892BF8C1C634" author="Ahrens, D. &amp; Fabrizi, S." box="[449,726,1339,1365]" pageId="9" pageNumber="380" pagination="1 - 355" refId="ref9865" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1-355." type="journal article" year="2016">Ahrens &amp; Fabrizi, 2016</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C3740CAA07AFF95C2DB892BF98FC635" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[738,920,1339,1365]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="onam" status="sp. nov.">
<emphasis id="B943E75BA07AFF95C2DB892BF98FC635" box="[738,920,1339,1365]" italics="true" pageId="9" pageNumber="380">Maladera onam</emphasis>
</taxonomicName>
<emphasis id="B943E75BA07AFF95C39B892BF9E9C634" bold="true" box="[930,1022,1339,1365]" pageId="9" pageNumber="380">
<taxonomicNameLabel id="A2705A20A07AFF95C39B892BF9E9C634" box="[930,1022,1339,1365]" pageId="9" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C3740CAA07AFF95C4A0892BFF5CC634" authorityName="Ahrens &amp; Fabrizi" authorityYear="2016" box="[1177,1355,1339,1365]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="trochaloides">
<emphasis id="B943E75BA07AFF95C4A0892BFF5CC634" box="[1177,1355,1339,1365]" italics="true" pageId="9" pageNumber="380">M. trochaloides</emphasis>
</taxonomicName>
by the right paramere which is strongly curved outward having a large internal lobe; its apex is finely setose, while in
<taxonomicName id="4C3740CAA07AFF95C547894FFB35C6FC" authorityName="Ahrens &amp; Fabrizi" authorityYear="2016" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="trochaloides">
<emphasis id="B943E75BA07AFF95C547894FFB35C6FC" italics="true" pageId="9" pageNumber="380">M. trochaloides</emphasis>
</taxonomicName>
it is a glabrous (
<bibRefCitation id="EFA646B8A07AFF95C1E58993F8F2C6FC" author="Ahrens, D. &amp; Fabrizi, S." box="[476,741,1411,1437]" pageId="9" pageNumber="380" pagination="1 - 355" refId="ref9865" refString="Ahrens, D. &amp; Fabrizi, S. (2016) A monograph of the Sericini of India (Coleoptera: Scarabaeidae). Bonn Zoological Bulletin, 65, 1-355." type="journal article" year="2016">Ahrens &amp; Fabrizi 2016</bibRefCitation>
; fig. 24R–T, p. 312).
</paragraph>
</subSubSection>
<subSubSection id="C32D68C2A07AFF95C0FE89B7FF20C6A0" box="[199,1335,1447,1473]" pageId="9" pageNumber="380" type="etymology">
<paragraph id="8B883B49A07AFF95C0FE89B7FF20C6A0" blockId="9.[151,1437,151,1473]" box="[199,1335,1447,1473]" pageId="9" pageNumber="380">
<emphasis id="B943E75BA07AFF95C0FE89B7FB59C6A0" bold="true" box="[199,334,1447,1473]" pageId="9" pageNumber="380">Etymology.</emphasis>
<taxonomicName id="4C3740CAA07AFF95C16C89B7F81FC6A1" authorityName="Gupta &amp; Bhunia &amp; Ahrens &amp; Chandra" authorityYear="2025" box="[341,520,1447,1473]" class="Insecta" family="Scarabaeidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="onam" status="sp. nov.">
<emphasis id="B943E75BA07AFF95C16C89B7F81FC6A1" box="[341,520,1447,1473]" italics="true" pageId="9" pageNumber="380">Maladera onam</emphasis>
</taxonomicName>
<emphasis id="B943E75BA07AFF95C23689B7F87EC6A0" bold="true" box="[527,617,1447,1473]" pageId="9" pageNumber="380">
<taxonomicNameLabel id="A2705A20A07AFF95C23689B7F87EC6A0" box="[527,617,1447,1473]" pageId="9" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is named after a famous festival of
<collectingRegion id="49F3F5ABA07AFF95C3C389B7FE52C6A0" box="[1018,1093,1447,1473]" country="India" name="Kerala" pageId="9" pageNumber="380">Kerala</collectingRegion>
(noun in apposition).
</paragraph>
</subSubSection>
</treatment>
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<emphasis id="B943E75BA07EFF91C0AE8C87FB80C3D0" bold="true" box="[151,407,151,177]" italics="true" pageId="13" pageNumber="384">Tetraserica ferrugata</emphasis>
(
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)
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<emphasis id="B943E75BA07EFF91C0AE8CCFFB97C398" bold="true" box="[151,384,223,249]" pageId="13" pageNumber="384">Material examined.</emphasis>
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,
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“Brit. Ost-Indien, Dehradun, 9.1943, leg.
<collectorName id="26C25E9FA07EFF91C3DF8CF0FE4CC39B" box="[998,1115,224,250]" pageId="13" pageNumber="384">K. Benner</collectorName>
, Eing. Nr. 8, 1950”(
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)
</materialsCitation>
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, #17-22a, leg.
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)
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.
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225
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Brenske, 1898
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<emphasis id="B943E75BA07FFF90C0AE8873FB92C71C" bold="true" box="[151,389,1123,1149]" pageId="12" pageNumber="383">Material examined.</emphasis>
<materialsCitation id="3B5F3114A07FFF90C1A98873F89EC7C0" collectionCode="VI" collectorName="Yanchu Khola" country="Nepal" elevation="2650" latitude="29.906666" location="Ghatte" longLatPrecision="20" longitude="81.918884" municipality="Humla Simlikot" pageId="12" pageNumber="383" specimenCount="1" specimenCount-male="1" stateProvince="Karnali">
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“
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, P:
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, D:
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~
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SE,
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,
<collectorName id="26C25E9FA07FFF90C4A88873FF2FC71C" box="[1169,1336,1123,1149]" pageId="12" pageNumber="383">Yanchu Khola</collectorName>
,
<quantity id="4CCF96ACA07FFF90C57E8873FF8FC71F" box="[1351,1432,1123,1150]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="12" pageNumber="383" unit="m" value="2650.0">
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,
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<quantity id="4CCF96ACA07FFF90C1F98897F806C7C0" box="[448,529,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="2.92" pageId="12" pageNumber="383" unit="m" value="2920.0">
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,
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16–18.
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<materialsCitation id="3B5F3114A07FFF90C2B38897FEA3C7A4" collectionCode="NME, VI" collectorName="A. Weigel" country="Nepal" latitude="29.909166" location="Humla Simlikot" longLatPrecision="20" longitude="81.91861" pageId="12" pageNumber="383" specimenCount="1" specimenCount-male="1" stateProvince="Karnali">
.2022, leg.
<collectorName id="26C25E9FA07FFF90C3378897F991C7C0" box="[782,902,1159,1185]" pageId="12" pageNumber="383">A. Weigel</collectorName>
, 22-#06” (
<collectionCode id="ED26A38CA07FFF90C43A8897FE50C7C0" box="[1027,1095,1159,1185]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="12" pageNumber="383" type="">NME</collectionCode>
),
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“
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, P:
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, D:
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,
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–
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SE,
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to
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,
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16.
<collectionCode id="ED26A38CA07FFF90C4AC88BBFEA3C7A4" box="[1173,1204,1195,1221]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="383">VI</collectionCode>
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<materialsCitation id="3B5F3114A07FFF90C48C88BBF94CC66C" collectionCode="NME, VI" collectorName="A. Weigel" country="Nepal" elevation="2478" latitude="29.59" location="Mugu Jumla Simlikot" longLatPrecision="20" longitude="82.14583" pageId="12" pageNumber="383" specimenCount="1" specimenCount-male="1" stateProvince="Karnali">
.2022,
<quantity id="4CCF96ACA07FFF90C53888BBFF8FC7A7" box="[1281,1432,1195,1222]" metricMagnitude="3" metricUnit="m" metricValue="2.4775" metricValueMax="2.92" metricValueMin="2.035" pageId="12" pageNumber="383" unit="m" value="2477.5" valueMax="2920.0" valueMin="2035.0">
<elevation id="001ADC7AA07FFF90C53888BBFF8FC7A7" box="[1281,1432,1195,1222]" metricMagnitude="3" metricUnit="m" metricValue="2.4775" metricValueMax="2.92" metricValueMin="2.035" pageId="12" pageNumber="383" unit="m" value="2477.5" valueMax="2920.0" valueMin="2035.0">2035–2920m</elevation>
</quantity>
, leg.
<collectorName id="26C25E9FA07FFF90C0FF88DFFB2EC788" box="[198,313,1231,1257]" pageId="12" pageNumber="383">A. Weigel</collectorName>
, #22-05a” (
<collectionCode id="ED26A38CA07FFF90C18288DFFBE8C788" box="[443,511,1231,1257]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="12" pageNumber="383" type="">NME</collectionCode>
),
<specimenCount id="9D31F0C0A07FFF90C22B88DFF82BC78B" box="[530,572,1231,1258]" count="1" pageId="12" pageNumber="383" type="male">1 ♂</specimenCount>
“
<collectingCountry id="F3207BD9A07FFF90C27488DFF882C788" box="[589,661,1231,1257]" name="Nepal" pageId="12" pageNumber="383">Nepal</collectingCountry>
, P:
<collectingRegion id="49F3F5ABA07FFF90C28788DFF902C788" box="[702,789,1231,1257]" country="Nepal" name="Karnali" pageId="12" pageNumber="383">Karnali</collectingRegion>
, D:
<location id="8EE86D92A07FFF90C37A88C0FE2EC788" LSID="urn:lsid:plazi:treatment:039E8A5FA07FFF90C0AE880BFBC6C634:8EE86D92A07FFF90C37A88C0FE2EC788" box="[835,1081,1231,1258]" country="Nepal" latitude="29.59" longLatPrecision="20" longitude="82.14583" name="Mugu Jumla Simlikot" pageId="12" pageNumber="383" stateProvince="Karnali">Mugu Jumla Simlikot</location>
, ~
<quantity id="4CCF96ACA07FFF90C46288DFFE8CC78B" box="[1115,1179,1231,1258]" metricMagnitude="4" metricUnit="m" metricValue="4.3" pageId="12" pageNumber="383" unit="km" value="43.0">43km</quantity>
–
<quantity id="4CCF96ACA07FFF90C49088DFFEFFC78B" box="[1193,1256,1231,1258]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="383" unit="km" value="35.0">35km</quantity>
N,
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,
<geoCoordinate id="EE035D8EA07FFF90C0AE88E3FB0DC66F" box="[151,282,1267,1294]" degrees="82" direction="east" minutes="08" orientation="longitude" pageId="12" pageNumber="383" precision="15" seconds="57" value="82.14916">82°08'57&quot;E</geoCoordinate>
, to
<geoCoordinate id="EE035D8EA07FFF90C17B88E3FBDEC66F" box="[322,457,1267,1294]" degrees="29" direction="north" minutes="35" orientation="latitude" pageId="12" pageNumber="383" precision="15" seconds="24" value="29.59">29°35'24&quot;N</geoCoordinate>
,
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,
<quantity id="4CCF96ACA07FFF90C25A88E3F8A6C66F" box="[611,689,1267,1294]" metricMagnitude="3" metricUnit="m" metricValue="3.565" pageId="12" pageNumber="383" unit="m" value="3565.0">
<elevation id="001ADC7AA07FFF90C25A88E3F8A6C66F" box="[611,689,1267,1294]" metricMagnitude="3" metricUnit="m" metricValue="3.565" pageId="12" pageNumber="383" unit="m" value="3565.0">3565m</elevation>
</quantity>
–
<quantity id="4CCF96ACA07FFF90C2F988E3F918C66F" box="[704,783,1267,1294]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="383" unit="m" value="2600.0">
<elevation id="001ADC7AA07FFF90C2F988E3F918C66F" box="[704,783,1267,1294]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="383" unit="m" value="2600.0">2600m</elevation>
</quantity>
,
<date id="FF891D89A07FFF90C32388E3F94CC66C" box="[794,859,1267,1293]" pageId="12" pageNumber="383" value="2022-06-26">
26.
<collectionCode id="ED26A38CA07FFF90C30288E3F94CC66C" box="[827,859,1267,1293]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="383">VI</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B5F3114A07FFF90C36288E3FEB3C650" collectionCode="NME, VI" collectorName="A. Weigel" country="Nepal" elevation="3065" latitude="29.671389" location="Jogimara" longLatPrecision="20" longitude="82.10722" municipality="Humla Simlikot" pageId="12" pageNumber="383" specimenCount="1" specimenCount-female="1" stateProvince="Karnali">
.2022, leg.
<collectorName id="26C25E9FA07FFF90C3EF88E3FE5DC66C" box="[982,1098,1267,1293]" pageId="12" pageNumber="383">A. Weigel</collectorName>
, #22-15” (
<collectionCode id="ED26A38CA07FFF90C48788E3FF15C66C" box="[1214,1282,1267,1293]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="12" pageNumber="383" type="">NME</collectionCode>
),
<specimenCount id="9D31F0C0A07FFF90C52C88E3FF28C66F" box="[1301,1343,1267,1294]" count="1" pageId="12" pageNumber="383" type="female">1 ♀</specimenCount>
“
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, P:
<collectingRegion id="49F3F5ABA07FFF90C08D8907FB1CC650" box="[180,267,1303,1329]" country="Nepal" name="Karnali" pageId="12" pageNumber="383">Karnali</collectingRegion>
, D:
<collectingMunicipality id="6BECA133A07FFF90C10E8907FBFBC650" box="[311,492,1303,1329]" pageId="12" pageNumber="383">Humla Simlikot</collectingMunicipality>
, ~
<quantity id="4CCF96ACA07FFF90C2328907F85CC650" box="[523,587,1303,1329]" metricMagnitude="4" metricUnit="m" metricValue="4.3" pageId="12" pageNumber="383" unit="km" value="43.0">43km</quantity>
SE,
<location id="8EE86D92A07FFF90C2478907F8F2C650" LSID="urn:lsid:plazi:treatment:039E8A5FA07FFF90C0AE880BFBC6C634:8EE86D92A07FFF90C2478907F8F2C650" box="[638,741,1303,1329]" country="Nepal" latitude="29.671389" longLatPrecision="20" longitude="82.10722" municipality="Humla Simlikot" name="Jogimara" pageId="12" pageNumber="383" stateProvince="Karnali">Jogimara</location>
<geoCoordinate id="EE035D8EA07FFF90C2D38907F967C650" box="[746,880,1303,1329]" degrees="29" direction="north" minutes="40" orientation="latitude" pageId="12" pageNumber="383" precision="15" seconds="17" value="29.671389">29°40'17&quot;N</geoCoordinate>
,
<geoCoordinate id="EE035D8EA07FFF90C3428907F9E9C650" box="[891,1022,1303,1329]" degrees="82" direction="east" minutes="06" orientation="longitude" pageId="12" pageNumber="383" precision="15" seconds="26" value="82.10722">82°06'26&quot;E</geoCoordinate>
,
<quantity id="4CCF96ACA07FFF90C4318907FE4EC653" box="[1032,1113,1303,1330]" metricMagnitude="3" metricUnit="m" metricValue="3.065" pageId="12" pageNumber="383" unit="m" value="3065.0">
<elevation id="001ADC7AA07FFF90C4318907FE4EC653" box="[1032,1113,1303,1330]" metricMagnitude="3" metricUnit="m" metricValue="3.065" pageId="12" pageNumber="383" unit="m" value="3065.0">3065m</elevation>
</quantity>
,
<date id="FF891D89A07FFF90C45A8907FEB3C650" box="[1123,1188,1303,1329]" pageId="12" pageNumber="383" value="2022-06-24">
24.
<collectionCode id="ED26A38CA07FFF90C4BD8907FEB3C650" box="[1156,1188,1303,1329]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="383">VI</collectionCode>
</date>
</materialsCitation>
.2022, 2035–2920 leg.
<collectorName id="26C25E9FA07FFF90C0AE892BFB1CC634" box="[151,267,1339,1365]" pageId="12" pageNumber="383">A. Weigel</collectorName>
, #22-15” (
<collectionCode id="ED26A38CA07FFF90C1B9892BFBD3C634" box="[384,452,1339,1365]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="12" pageNumber="383" type="">NME</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>

115
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<mods:title id="F22FD22552E2FEEA14AAAC087326B7F0">New species and records of Sericinae from India and Nepal (Coleoptera: Scarabaeidae)</mods:title>
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<emphasis id="B943E75BA07FFF90C0AE89B7FB40C6A3" bold="true" box="[151,343,1447,1474]" italics="true" pageId="12" pageNumber="383">Serica khajiaris</emphasis>
Mittal, 1988
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<emphasis id="B943E75BA07FFF90C0AE89FFFB91C568" bold="true" box="[151,390,1519,1545]" pageId="12" pageNumber="383">Material examined.</emphasis>
<materialsCitation id="3B5F3114A07FFF90C1AA89FFF9E8C54C" collectingDate="2022-06-20" collectionCode="NME" collectorName="A. Weigel" country="Nepal" elevation="2650" latitude="29.830278" location="Lumcha" longLatPrecision="20" longitude="81.9725" municipality="Humla Simlikot" pageId="12" pageNumber="383" specimenCount="1" specimenCount-male="1" stateProvince="Karnali">
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“
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, P:
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, D:
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~
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SE, Rodikot, S,
<location id="8EE86D92A07FFF90C4E189FFFF20C568" LSID="urn:lsid:plazi:treatment:039E8A5FA07FFF90C0AE89B7FE14C54C:8EE86D92A07FFF90C4E189FFFF20C568" box="[1240,1335,1519,1545]" country="Nepal" latitude="29.830278" longLatPrecision="20" longitude="81.9725" municipality="Humla Simlikot" name="Lumcha" pageId="12" pageNumber="383" stateProvince="Karnali">Lumcha</location>
,
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,
<date id="FF891D89A07FFF90C0AE8A03FB0EC54C" box="[151,281,1555,1581]" pageId="12" pageNumber="383" value="2022-06-20">
<collectingDate id="EFCDE461A07FFF90C0AE8A03FB0EC54C" box="[151,281,1555,1581]" pageId="12" pageNumber="383" value="2022-06-20">20.VI.2022</collectingDate>
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,
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,
<geoCoordinate id="EE035D8EA07FFF90C18E8A03F82DC54F" box="[439,570,1555,1582]" degrees="81" direction="east" minutes="58" orientation="longitude" pageId="12" pageNumber="383" precision="15" seconds="21" value="81.9725">81°58'21&quot;E</geoCoordinate>
, leg.
<collectorName id="26C25E9FA07FFF90C24D8A03F8FFC54C" box="[628,744,1555,1581]" pageId="12" pageNumber="383">A. Weigel</collectorName>
, HF/KL 22-#11” (
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)
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.
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111
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, pasture, #22-17” (
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<emphasis id="B943E75BA07FFF90C0AE8D7FFB94C2E8" bold="true" box="[151,387,367,393]" italics="true" pageId="12" pageNumber="383">Oxyserica pruinosa</emphasis>
(
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<emphasis id="B943E75BA07FFF90C0AE8DA7FB96C2B0" bold="true" box="[151,385,439,465]" pageId="12" pageNumber="383">Material examined.</emphasis>
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<specimenCount id="9D31F0C0A07FFF90C1B08DA7FBDDC2B3" box="[393,458,439,466]" count="4" pageId="12" pageNumber="383" type="male">4 ♂♂</specimenCount>
,
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“
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,
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</quantity>
,
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20.
<collectionCode id="ED26A38CA07FFF90C3588DA7F960C2B3" box="[865,887,439,466]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="383" type="Museum">V</collectionCode>
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.1956, D. Indien Exp, Nr 961, Eing, Nr, 1, 1956” (
<collectionCode id="ED26A38CA07FFF90C0A68DCBFAE0C294" box="[159,247,475,501]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="12" pageNumber="383" type="">ZMUH</collectionCode>
),
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“
<collectingCountry id="F3207BD9A07FFF90C1768DCBFB80C294" box="[335,407,475,501]" name="Nepal" pageId="12" pageNumber="383">Nepal</collectingCountry>
, P:
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, D:
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, ~
<quantity id="4CCF96ACA07FFF90C3348DCBF95BC297" box="[781,844,475,502]" metricMagnitude="4" metricUnit="m" metricValue="4.1" pageId="12" pageNumber="383" unit="km" value="41.0">41km</quantity>
–
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N,
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<elevation id="001ADC7AA07FFF90C1BA8DEFFBC5C17B" box="[387,466,511,538]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="383" unit="m" value="2600.0">2600m</elevation>
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,
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26.
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<materialsCitation id="3B5F3114A07FFF90C2268DEFF9CBC178" box="[543,988,511,537]" collectionCode="NME" collectorName="A. Weigel" country="Nepal" pageId="12" pageNumber="383" specimenCount="1" stateProvince="Karnali">
.2022, leg.
<collectorName id="26C25E9FA07FFF90C2A38DEFF91AC178" box="[666,781,511,537]" pageId="12" pageNumber="383">A. Weigel</collectorName>
, #22-16a” (
<collectionCode id="ED26A38CA07FFF90C3B68DEFF9C4C178" box="[911,979,511,537]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="12" pageNumber="383" type="">NME</collectionCode>
)
</materialsCitation>
.
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149
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<emphasis id="B943E75BA07FFF90C0AE8F0FFBAFC058" bold="true" box="[151,440,799,826]" italics="true" pageId="12" pageNumber="383">Pachyserica marmorata</emphasis>
(
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<emphasis id="B943E75BA07FFF90C0AE8F77FB92C0E0" bold="true" box="[151,389,871,897]" pageId="12" pageNumber="383">Material examined.</emphasis>
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<specimenCount id="9D31F0C0A07FFF90C1A88F77FBA8C0E3" box="[401,447,871,898]" count="1" pageId="12" pageNumber="383" type="male">1 ♂</specimenCount>
“
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, P:
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, D:
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, ~
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–
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N,
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, to
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,
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</quantity>
–
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,
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26.
<collectionCode id="ED26A38CA07FFF90C2A08F9BF8AEC0C4" box="[665,697,907,933]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="383">VI</collectionCode>
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.2022, leg.
<collectorName id="26C25E9FA07FFF90C3038F9BF9A7C0C4" box="[826,944,907,933]" pageId="12" pageNumber="383">A. Weigel</collectorName>
, #22-16a” (
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),
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“
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,
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,
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,
<date id="FF891D89A07FFF90C0CA8FBFFB3FC0AB" box="[243,296,943,970]" pageId="12" pageNumber="383" value="1956-05-16">
16.
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<materialsCitation id="3B5F3114A07FFF90C1118FBFF9ADC0A8" box="[296,954,943,970]" collectionCode="ZMUH" country="India" location="Indien" pageId="12" pageNumber="383" specimenCount="1">
.1956, D.
<collectingCountry id="F3207BD9A07FFF90C1AF8FBFFBC9C0A8" box="[406,478,943,969]" name="India" pageId="12" pageNumber="383">Indien</collectingCountry>
Exp, Nr 961, Eing. Nr, 1, 1956” (
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)
</materialsCitation>
.
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91
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Comparison of protibia of

302
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<taxonomicName id="4C6138B0B26E9C3FFF07FB48FBCBFB43" authority="Perez Santa-Rita, Dombroskie, Ledezma &amp; Baixeras" authorityName="Perez Santa-Rita, Dombroskie, Ledezma &amp; Baixeras" authorityYear="2025" box="[151,1113,1164,1191]" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="interandina" status="sp. nov.">
<emphasis id="B9159F21B26E9C3FFF07FB48FE2DFB43" bold="true" box="[151,447,1164,1191]" italics="true" pageId="3" pageNumber="364">Enicotaenia interandina</emphasis>
Pérez Santa-Rita, Dombroskie, Ledezma &amp; Baixeras
</taxonomicName>
,
<emphasis id="B9159F21B26E9C3FFBF5FB48FB61FB43" bold="true" box="[1125,1267,1165,1191]" italics="true" pageId="3" pageNumber="364">
<taxonomicNameLabel id="A226225AB26E9C3FFBF5FB48FB61FB43" box="[1125,1267,1165,1191]" pageId="3" pageNumber="364" rank="species">new species</taxonomicNameLabel>
</emphasis>
</emphasis>
</heading>
<uri id="FFF04F31B26E9C3FFF07FB74FC44FB28" box="[151,982,1201,1228]" pageId="3" pageNumber="364">
urn:lsid:zoobank.org:pub:
<uuid id="FFC779E6B26E9C3FFE20FB74FC44FB28" box="[432,982,1201,1228]" pageId="3" pageNumber="364">0EB9272D-4A57-48DA-AD47-972D519B9876</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C37B10B8B26E9C3FFF07FB10FE84FB0B" box="[151,278,1237,1263]" pageId="3" pageNumber="364" type="description">
<paragraph id="8BDE4333B26E9C3FFF07FB10FE84FB0B" blockId="3.[151,1267,1164,1263]" box="[151,278,1237,1263]" pageId="3" pageNumber="364">
(
<figureCitation id="135A5FB6B26E9C3FFF0FFB10FE9CFB0B" box="[159,270,1237,1263]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[218,1378,197,1883]" captionTargetId="figure-26@4.[210,1378,179,1883]" captionTargetPageId="4" captionText="FIGURE 2. Habitus and genitalia of Enicotaenia interandina new genus and species. A, Adult male Holotype; B, Upperside forewing pattern; C, Male genitalia (GS.: 20720); D, Gnathos and transtilla of male genitalia (GS.: 20681); E, Female genitalia (GS.: 20680); F, Detail of female genitalia (GS.: 21005). Scale bars: 2 mm (A); 1 mm (B, E); 500 μm (C); 100 μm (D); 200 μm (F)." figureDoi="http://doi.org/10.5281/zenodo.15216351" httpUri="https://zenodo.org/record/15216351/files/figure.png" pageId="3" pageNumber="364">Fig 2A–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37B10B8B26E9C3FFF07FAD8FBD8FA0F" pageId="3" pageNumber="364" type="materials_examined">
<paragraph id="8BDE4333B26E9C3FFF07FAD8FBD8FA0F" blockId="3.[151,1437,1309,2020]" pageId="3" pageNumber="364">
<emphasis id="B9159F21B26E9C3FFF07FAD8FED4FAD3" bold="true" box="[151,326,1309,1335]" pageId="3" pageNumber="364">Type material.</emphasis>
<materialsCitation id="3B09496EB26E9C3FFEDDFAD8FE2BFA64" collectingDate="2011-01-28" collectionCode="MNKM" collectorName="El Millu &amp; J. Baixeras &amp; A. Valdivia &amp; G. Fernandez" country="Bolivia" county="Florida Province" elevation="1534" latitude="-17.989334" location="El Millu" longLatPrecision="1" longitude="-64.05403" municipality="Pampa Grande Municipality" pageId="3" pageNumber="364" specimenCode="GS 20681" specimenCount="1" specimenCount-male="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="54DAFD91B26E9C3FFEDDFAD8FE2BFAD3" box="[333,441,1309,1335]" pageId="3" pageNumber="364" type="holotype">Holotype</typeStatus>
. [
<collectingCountry id="F37603A3B26E9C3FFE5BFAD8FDD6FAD3" box="[459,580,1309,1335]" name="Bolivia" pageId="3" pageNumber="364">BOLIVIA</collectingCountry>
]
<specimenCount id="9D6788BAB26E9C3FFDC2FADBFDFBFADC" box="[594,617,1310,1336]" count="1" pageId="3" pageNumber="364" type="male">♂</specimenCount>
,
<collectingRegion id="49A58DD1B26E9C3FFDE6FAD8FC13FADC" box="[630,897,1309,1336]" country="Bolivia" name="Santa Cruz" pageId="3" pageNumber="364">Santa Cruz Department</collectingRegion>
,
<collectingCounty id="62BF3BBFB26E9C3FFC1DFAD8FBDEFAD3" box="[909,1100,1309,1335]" pageId="3" pageNumber="364">Florida Province</collectingCounty>
,
<collectingMunicipality id="6BBAD949B26E9C3FFBC8FADBFA0AFAD3" box="[1112,1432,1309,1336]" pageId="3" pageNumber="364">Pampa Grande Municipality</collectingMunicipality>
, locality of
<collectorName id="269426E5B26E9C3FFE80FA84FEE3FABF" box="[272,369,1345,1371]" pageId="3" pageNumber="364">
<location id="8EBE15E8B26E9C3FFE80FA84FEE3FABF" LSID="urn:lsid:plazi:treatment:03C8F225B26E9C39FF07FB48FC09FCF1:8EBE15E8B26E9C3FFE80FA84FEE3FABF" box="[272,369,1345,1371]" country="Bolivia" county="Florida Province" latitude="-17.989334" longLatPrecision="1" longitude="-64.05403" municipality="Pampa Grande Municipality" name="El Millu" pageId="3" pageNumber="364" stateProvince="Santa Cruz">El Millu</location>
</collectorName>
,
<quantity id="4C99EED6B26E9C3FFEEDFA84FE46FAB8" box="[381,468,1345,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.534" pageId="3" pageNumber="364" unit="m" value="1534.0">
<elevation id="004CA400B26E9C3FFEEDFA84FE46FAB8" box="[381,468,1345,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.534" pageId="3" pageNumber="364" unit="m" value="1534.0">1534 m</elevation>
</quantity>
,
<geoCoordinate id="EE5525F4B26E9C3FFE4FFA84FDEBFAB8" box="[479,633,1345,1372]" degrees="17" direction="south" minutes="59" orientation="latitude" pageId="3" pageNumber="364" precision="1" seconds="21.6" value="-17.989334">S17º59’21.6”</geoCoordinate>
;
<geoCoordinate id="EE5525F4B26E9C3FFD15FA84FCBBFAB8" box="[645,809,1345,1372]" degrees="64" direction="west" minutes="03" orientation="longitude" pageId="3" pageNumber="364" precision="1" seconds="14.5" value="-64.05403">W64º03’14.5”</geoCoordinate>
,
<date id="FFDF65F3B26E9C3FFCA4FA84FC2CFABF" box="[820,958,1345,1372]" pageId="3" pageNumber="364" value="2011-01-28">
<collectingDate id="EF9B9C1BB26E9C3FFCA4FA84FC2CFABF" box="[820,958,1345,1372]" pageId="3" pageNumber="364" value="2011-01-28">28 Jan 2011</collectingDate>
</date>
,
<collectorName id="269426E5B26E9C3FFC59FA87FBD5FABF" box="[969,1095,1345,1372]" pageId="3" pageNumber="364">J. Baixeras</collectorName>
,
<collectorName id="269426E5B26E9C3FFBC0FA84FB5DFABF" box="[1104,1231,1345,1371]" pageId="3" pageNumber="364">A. Valdivia</collectorName>
and
<collectorName id="269426E5B26E9C3FFA95FA84FA0FFABF" box="[1285,1437,1345,1371]" pageId="3" pageNumber="364">G. Fernández</collectorName>
(
<specimenCode id="DBC7EB48B26E9C3FFF0FFAA0FE87FA9B" box="[159,277,1381,1407]" collectionCode="GS" pageId="3" pageNumber="364">GS 20681</specimenCode>
)-coll.
<collectionCode id="ED70DBF6B26E9C3FFECBFAA3FE2BFA64" box="[347,441,1382,1408]" pageId="3" pageNumber="364">MNKM</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B09496EB26E9C3FFE52FAA3FBD4FA0F" collectingDate="2011-01-28" collectionCode="MNKM" collectorName="El Millu &amp; J. Baixeras &amp; A. Valdivia &amp; G. Fernandez" country="Bolivia" county="Florida Province" elevation="1534" latitude="-17.989334" location="El Millu" longLatPrecision="1" longitude="-64.05403" municipality="Pampa Grande Municipality" pageId="3" pageNumber="364" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Santa Cruz" typeStatus="paratype">
<typeStatus id="54DAFD91B26E9C3FFE52FAA3FDA1FA64" box="[450,563,1382,1408]" pageId="3" pageNumber="364" type="paratype">Paratypes</typeStatus>
. [BOLIVA]
<specimenCount id="9D6788BAB26E9C3FFD2CFAA0FD73FA9B" box="[700,737,1381,1407]" count="3" pageId="3" pageNumber="364" type="male">3♂</specimenCount>
,
<specimenCount id="9D6788BAB26E9C3FFD7CFAA0FC82FA9B" box="[748,784,1381,1407]" count="2" pageId="3" pageNumber="364" type="female">2♀</specimenCount>
,
<collectingRegion id="49A58DD1B26E9C3FFC8CFAA0FBB7FA64" box="[796,1061,1381,1408]" country="Bolivia" name="Santa Cruz" pageId="3" pageNumber="364">Santa Cruz Department</collectingRegion>
,
<collectingCounty id="62BF3BBFB26E9C3FFBBFFAA0FB7EFA9B" box="[1071,1260,1381,1407]" pageId="3" pageNumber="364">Florida Province</collectingCounty>
,
<collectingMunicipality id="6BBAD949B26E9C3FFB67FAA3FEBBFA47" pageId="3" pageNumber="364">Pampa Grande Municipality</collectingMunicipality>
, locality of
<collectorName id="269426E5B26E9C3FFE3DFA4CFD9CFA47" box="[429,526,1417,1443]" pageId="3" pageNumber="364">
<location id="8EBE15E8B26E9C3FFE3DFA4CFD9CFA47" LSID="urn:lsid:plazi:treatment:03C8F225B26E9C39FF07FB48FC09FCF1:8EBE15E8B26E9C3FFE3DFA4CFD9CFA47" box="[429,526,1417,1443]" country="Bolivia" county="Florida Province" latitude="-17.989334" longLatPrecision="1" longitude="-64.05403" municipality="Pampa Grande Municipality" name="El Millu" pageId="3" pageNumber="364" stateProvince="Santa Cruz">El Millu</location>
</collectorName>
,
<quantity id="4C99EED6B26E9C3FFD8AFA4CFDE3FA40" box="[538,625,1417,1444]" metricMagnitude="3" metricUnit="m" metricValue="1.534" pageId="3" pageNumber="364" unit="m" value="1534.0">
<elevation id="004CA400B26E9C3FFD8AFA4CFDE3FA40" box="[538,625,1417,1444]" metricMagnitude="3" metricUnit="m" metricValue="1.534" pageId="3" pageNumber="364" unit="m" value="1534.0">1534 m</elevation>
</quantity>
,
<geoCoordinate id="EE5525F4B26E9C3FFDEDFA4CFC85FA40" box="[637,791,1417,1444]" degrees="17" direction="south" minutes="59" orientation="latitude" pageId="3" pageNumber="364" precision="1" seconds="21.6" value="-17.989334">S17º59’21.6”</geoCoordinate>
;
<geoCoordinate id="EE5525F4B26E9C3FFCB2FA4CFC55FA40" box="[802,967,1417,1444]" degrees="64" direction="west" minutes="03" orientation="longitude" pageId="3" pageNumber="364" precision="1" seconds="14.5" value="-64.05403">W64º03’14.5”</geoCoordinate>
,
<date id="FFDF65F3B26E9C3FFC42FA4CFBC9FA47" box="[978,1115,1417,1444]" pageId="3" pageNumber="364" value="2011-01-28">
<collectingDate id="EF9B9C1BB26E9C3FFC42FA4CFBC9FA47" box="[978,1115,1417,1444]" pageId="3" pageNumber="364" value="2011-01-28">28 Jan 2011</collectingDate>
</date>
,
<collectorName id="269426E5B26E9C3FFBF6FA4FFB76FA47" box="[1126,1252,1417,1444]" pageId="3" pageNumber="364">J. Baixeras</collectorName>
,
<collectorName id="269426E5B26E9C3FFB7EFA4CFAFEFA47" box="[1262,1388,1417,1443]" pageId="3" pageNumber="364">A. Valdivia</collectorName>
and
<collectorName id="269426E5B26E9C3FFF07FA68FEBDFA23" box="[151,303,1453,1479]" pageId="3" pageNumber="364">G. Fernández</collectorName>
(GS 20680, 20717, 20720) (DNA collection codes: DNA2023006, DNA2023033, DNA2023038).
<specimenCount id="9D6788BAB26E9C3FFA1EFA68FF5FFA0F" count="1" pageId="3" pageNumber="364" type="male">1 male</specimenCount>
and
<specimenCount id="9D6788BAB26E9C3FFE94FA14FEF7FA0F" box="[260,357,1489,1515]" count="1" pageId="3" pageNumber="364" type="female">1 female</specimenCount>
deposited in
<collectionCode id="ED70DBF6B26E9C3FFE6BFA17FDCAFA08" box="[507,600,1490,1516]" pageId="3" pageNumber="364">MNKM</collectionCode>
,
<specimenCount id="9D6788BAB26E9C3FFDF4FA14FD28FA0F" box="[612,698,1489,1515]" count="2" pageId="3" pageNumber="364" type="male">2 males</specimenCount>
and
<specimenCount id="9D6788BAB26E9C3FFD60FA14FCC3FA0F" box="[752,849,1489,1515]" count="1" pageId="3" pageNumber="364" type="female">1 female</specimenCount>
deposited in ICBiBE
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B10B8B26E9C3FFF57FA30FDDFF927" pageId="3" pageNumber="364" type="diagnosis">
<paragraph id="8BDE4333B26E9C3FFF57FA30FDDFF927" blockId="3.[151,1437,1309,2020]" pageId="3" pageNumber="364">
<emphasis id="B9159F21B26E9C3FFF57FA30FED0F9EB" bold="true" box="[199,322,1525,1551]" pageId="3" pageNumber="364">Diagnosis.</emphasis>
This species may be distinguished from other
<taxonomicName id="4C6138B0B26E9C3FFCD6FA30FC23F9EB" authorityName="Pierce &amp; Metcalfe" authorityYear="1922" box="[838,945,1525,1551]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="368" phylum="Arthropoda" rank="tribe" tribe="Archipini">Archipini</taxonomicName>
by the characters discussed in the diagnosis of the genus. Only two species are known in the genus
<taxonomicName id="4C6138B0B26E9C3FFC9AF9DFFC03F9D7" authorityName="Perez Santa-Rita, Dombroskie, Ledezma &amp; Baixeras" authorityYear="2025" box="[778,913,1562,1587]" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="B9159F21B26E9C3FFC9AF9DFFC03F9D7" box="[778,913,1562,1587]" italics="true" pageId="3" pageNumber="364">Enicotaenia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6138B0B26E9C3FFC30F9DFFF4BF9B3" authority="(Razowski &amp; Becker, 2000)" authorityName="Santa-Rita &amp; Dombroskie &amp; Ledezma &amp; Baixeras" authorityYear="2025" baseAuthorityName="Razowski &amp; Becker" baseAuthorityYear="2000" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="marabana" status="comb. nov.">
<emphasis id="B9159F21B26E9C3FFC30F9DFFB30F9D7" box="[928,1186,1561,1587]" italics="true" pageId="3" pageNumber="364">Enicotaenia marabana</emphasis>
(
<bibRefCitation id="EFF03EC2B26E9C3FFB21F9DCFF43F9B3" author="Razowski, J. &amp; Becker, V. O." pageId="3" pageNumber="364" pagination="3 - 4" refId="ref6154" refString="Razowski, J. &amp; Becker, V. O. (2000) Description of nine new Neotropical genera of Archipini (Lepidoptera: Tortricidae) and their species. Acta Zoologica Cracoviensia, 43 (3-4), 199 - 206." type="journal article" year="2000">Razowski &amp; Becker, 2000</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A226225AB26E9C3FFF71F9F8FED1F9BC" box="[225,323,1597,1624]" pageId="3" pageNumber="364" rank="species">comb. n.</taxonomicNameLabel>
and
<taxonomicName id="4C6138B0B26E9C3FFEEBF9FBFD8DF9B3" authorityName="Perez Santa-Rita, Dombroskie, Ledezma &amp; Baixeras" authorityYear="2025" box="[379,543,1597,1623]" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="interandina">
<emphasis id="B9159F21B26E9C3FFEEBF9FBFD8DF9B3" box="[379,543,1597,1623]" italics="true" pageId="3" pageNumber="364">E. interandina</emphasis>
</taxonomicName>
here described, both extremely similar. The male of
<taxonomicName id="4C6138B0B26E9C3FFBE7F9FBFA98F9B3" authorityName="Santa-Rita &amp; Dombroskie &amp; Ledezma &amp; Baixeras" authorityYear="2025" baseAuthorityName="Razowski &amp; Becker" baseAuthorityYear="2000" box="[1143,1290,1597,1623]" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="marabana">
<emphasis id="B9159F21B26E9C3FFBE7F9FBFA98F9B3" box="[1143,1290,1597,1623]" italics="true" pageId="3" pageNumber="364">E. marabana</emphasis>
</taxonomicName>
is unknown, and the female is only known from the
<typeStatus id="54DAFD91B26E9C3FFDC7F9A4FD2EF99F" box="[599,700,1633,1659]" pageId="3" pageNumber="364" type="holotype">holotype</typeStatus>
, a single worn specimen from
<collectingCountry id="F37603A3B26E9C3FFB88F9A4FBF2F99F" box="[1048,1120,1633,1659]" name="Brazil" pageId="3" pageNumber="364">Brazil</collectingCountry>
. The ductus bursae is rather broad in
<taxonomicName id="4C6138B0B26E9C3FFE92F943FE3AF97B" authorityName="Perez Santa-Rita, Dombroskie, Ledezma &amp; Baixeras" authorityYear="2025" box="[258,424,1669,1695]" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="interandina">
<emphasis id="B9159F21B26E9C3FFE92F943FE3AF97B" box="[258,424,1669,1695]" italics="true" pageId="3" pageNumber="364">E. interandina</emphasis>
</taxonomicName>
and appears narrower in
<emphasis id="B9159F21B26E9C3FFD46F943FCE6F97B" box="[726,884,1669,1695]" italics="true" pageId="3" pageNumber="364">
<taxonomicName id="4C6138B0B26E9C3FFD46F943FCE2F97B" authorityName="Santa-Rita &amp; Dombroskie &amp; Ledezma &amp; Baixeras" authorityYear="2025" baseAuthorityName="Razowski &amp; Becker" baseAuthorityYear="2000" box="[726,880,1669,1695]" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="marabana">E. marabana</taxonomicName>
.
</emphasis>
The anterior hypophysis is twice as long in
<taxonomicName id="4C6138B0B26E9C3FFA14F943FE88F927" authorityName="Perez Santa-Rita, Dombroskie, Ledezma &amp; Baixeras" authorityYear="2025" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="interandina">
<emphasis id="B9159F21B26E9C3FFA14F943FE88F927" italics="true" pageId="3" pageNumber="364">E. interandina</emphasis>
</taxonomicName>
compared to
<emphasis id="B9159F21B26E9C3FFE23F96FFDDFF927" box="[435,589,1705,1731]" italics="true" pageId="3" pageNumber="364">
<taxonomicName id="4C6138B0B26E9C3FFE23F96FFDDBF927" authorityName="Santa-Rita &amp; Dombroskie &amp; Ledezma &amp; Baixeras" authorityYear="2025" baseAuthorityName="Razowski &amp; Becker" baseAuthorityYear="2000" box="[435,585,1705,1731]" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="364" phylum="Arthropoda" rank="species" species="marabana">E. marabana</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37B10B8B26E9C39FF57F908FE85FDFD" lastPageId="5" lastPageNumber="366" pageId="3" pageNumber="364" type="description">
<paragraph id="8BDE4333B26E9C3FFF57F908FEE7F8E8" blockId="3.[151,1437,1309,2020]" pageId="3" pageNumber="364">
<emphasis id="B9159F21B26E9C3FFF57F908FECBF903" bold="true" box="[199,345,1741,1767]" pageId="3" pageNumber="364">Description.</emphasis>
<emphasis id="B9159F21B26E9C3FFECFF908FE0EF903" box="[351,412,1741,1767]" italics="true" pageId="3" pageNumber="364">Head</emphasis>
: Frons, vertex, labial palpus and antenna orangish; presence of dark scales dorsally on labial palpus and antenna.
</paragraph>
<paragraph id="8BDE4333B26E9C39FF57F8D0FD61FF1D" blockId="3.[151,1437,1309,2020]" lastBlockId="5.[151,1437,151,789]" lastPageId="5" lastPageNumber="366" pageId="3" pageNumber="364">
<emphasis id="B9159F21B26E9C3FFF57F8D0FE8AF8CB" box="[199,280,1813,1839]" italics="true" pageId="3" pageNumber="364">Thorax</emphasis>
: Dorsum orangish with the presence of groups of dark scales mixed; forewing length
<quantity id="4C99EED6B26E9C3FFB62F8D0FAE9F8D4" box="[1266,1403,1813,1840]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.6" metricValueMin="5.7" pageId="3" pageNumber="364" unit="mm" value="6.15" valueMax="6.6" valueMin="5.7">5.7–6.6 mm</quantity>
(
<emphasis id="B9159F21B26E9C3FFA1CF8D1FA0EF8D4" bold="true" box="[1420,1436,1812,1840]" pageId="3" pageNumber="364">x̄</emphasis>
=6.2; n= 4) in males, 7.0–
<quantity id="4C99EED6B26E9C3FFE2BF8FCFD83F8B0" box="[443,529,1849,1876]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="364" unit="mm" value="7.4">7.4 mm</quantity>
(
<emphasis id="B9159F21B26E9C3FFDB1F8FDFDA3F8B0" bold="true" box="[545,561,1848,1876]" pageId="3" pageNumber="364">x̄</emphasis>
=7.2; n=2) in females. Forewing pattern (
<figureCitation id="135A5FB6B26E9C3FFB9AF8FCFB16F8B7" box="[1034,1156,1849,1875]" captionStart="FIGURE 2" captionStartId="4.[151,250,1909,1933]" captionTargetBox="[218,1378,197,1883]" captionTargetId="figure-26@4.[210,1378,179,1883]" captionTargetPageId="4" captionText="FIGURE 2. Habitus and genitalia of Enicotaenia interandina new genus and species. A, Adult male Holotype; B, Upperside forewing pattern; C, Male genitalia (GS.: 20720); D, Gnathos and transtilla of male genitalia (GS.: 20681); E, Female genitalia (GS.: 20680); F, Detail of female genitalia (GS.: 21005). Scale bars: 2 mm (A); 1 mm (B, E); 500 μm (C); 100 μm (D); 200 μm (F)." figureDoi="http://doi.org/10.5281/zenodo.15216351" httpUri="https://zenodo.org/record/15216351/files/figure.png" pageId="3" pageNumber="364">Fig. 2A–B</figureCitation>
) not sexually dimorphic. Forewing upperside general background colour from ochreous to orangish with scattered dark brown marks, without distinct system of strigulae; system of fasciae incomplete; basal and subbasal fasciae only expressed on dorsum as a dark brown transverse suffused area; median fascia brown, broader on costa, confluent to termen, narrowly fragmented across the wing; postmedian and preterminal fasciae fused as a preapical triangular spot; fringe dark in apical area, lighter in termen and tornus; interfascial areas light brown. Forewing underside uniformly creme, no system of strigulae. Hindwing upperside and underside fairly concolorous, uniformly light orangish brown. Female wing pattern similar to male, no obvious dimorphism.
</paragraph>
<caption id="DF1E13BBB2699C38FF07F8B0FF2FF81D" ID-DOI="http://doi.org/10.5281/zenodo.15216351" ID-Zenodo-Dep="15216351" httpUri="https://zenodo.org/record/15216351/files/figure.png" pageId="4" pageNumber="365" startId="4.[151,250,1909,1933]" targetBox="[218,1378,197,1883]" targetPageId="4" targetType="figure">
<paragraph id="8BDE4333B2699C38FF07F8B0FF2FF81D" blockId="4.[151,1436,1909,2041]" pageId="4" pageNumber="365">
<emphasis id="B9159F21B2699C38FF07F8B0FE86F869" bold="true" box="[151,276,1909,1933]" pageId="4" pageNumber="365">FIGURE 2.</emphasis>
Habitus and genitalia of
<taxonomicName id="4C6138B0B2699C38FD8AF8B3FC80F869" authorityName="Santa-Rita &amp; Dombroskie &amp; Ledezma &amp; Baixeras" authorityYear="2025" box="[538,786,1909,1933]" class="Insecta" family="Tortricidae" genus="Enicotaenia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="365" phylum="Arthropoda" rank="species" species="interandina" status="new genus and species">
<emphasis id="B9159F21B2699C38FD8AF8B3FC80F869" box="[538,786,1909,1933]" italics="true" pageId="4" pageNumber="365">Enicotaenia interandina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A226225AB2699C38FC89F8B0FB94F869" box="[793,1030,1909,1933]" pageId="4" pageNumber="365" rank="species">new genus and species</taxonomicNameLabel>
. A, Adult male Holotype; B, Upperside forewing pattern; C, Male genitalia (GS.: 20720); D, Gnathos and transtilla of male genitalia (GS.: 20681); E, Female genitalia (GS.: 20680); F, Detail of female genitalia (GS.: 21005). Scale bars: 2 mm (A); 1 mm (B, E); 500 μm (C); 100 μm (D); 200 μm (F).
</paragraph>
</caption>
<paragraph id="8BDE4333B2689C39FF57FEC6FE50FEA5" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="366">
<emphasis id="B9159F21B2689C39FF57FEC6FEA3FEF9" box="[199,305,259,285]" italics="true" pageId="5" pageNumber="366">Abdomen</emphasis>
: Dorsad orangish, brownish cephalad, paler toward the caudal segments. Male and female genitalia as described for the genus.
</paragraph>
<paragraph id="8BDE4333B2689C39FF57FE8EFC44FE35" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="366">
<emphasis id="B9159F21B2689C39FF57FE8EFD88FE80" bold="true" box="[199,538,330,357]" pageId="5" pageNumber="366">Molecular characterization.</emphasis>
We were able to obtain sequences for small fragments of COI from
<specimenCount id="9D6788BAB2689C39FAF4FE8EFE9DFE6D" count="3" pageId="5" pageNumber="366" type="generic">three specimens</specimenCount>
. Two identical small fragments of 164 bp were obtained from
<specimenCount id="9D6788BAB2689C39FC56FEAAFBB9FE6D" box="[966,1067,367,393]" count="1" pageId="5" pageNumber="366" type="male">one male</specimenCount>
and
<specimenCount id="9D6788BAB2689C39FBF2FEAAFB4FFE6D" box="[1122,1245,367,393]" count="1" pageId="5" pageNumber="366" type="female">one female</specimenCount>
specimens using the primer pair combination C_TypeF1 and C_TypeR1. Another small fragment of 307 bp was obtained from
<specimenCount id="9D6788BAB2689C39FAE4FE56FF71FE35" count="1" pageId="5" pageNumber="366" type="female">one female</specimenCount>
specimen using the pair combination LepF1 and MLepR1 primers.
</paragraph>
<paragraph id="8BDE4333B2689C39FF57FE1EFE85FDFD" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="366">The sequences are publicly available through GenBank accession numbers PV105885, PV241496 and PV241497.</paragraph>
</subSubSection>
<subSubSection id="C37B10B8B2689C39FF57FDE7FC27FD4D" pageId="5" pageNumber="366" type="biology_ecology">
<paragraph id="8BDE4333B2689C39FF57FDE7FC27FD4D" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="366">
<emphasis id="B9159F21B2689C39FF57FDE7FE6FFDD8" bold="true" box="[199,509,546,573]" pageId="5" pageNumber="366">Biology and Distribution.</emphasis>
Early stages are unknown; adults have been collected in January (n=5). Adults were collected at middle elevation (
<quantity id="4C99EED6B2689C39FDB9FD83FD13FD85" box="[553,641,582,609]" metricMagnitude="3" metricUnit="m" metricValue="1.534" pageId="5" pageNumber="366" unit="m" value="1534.0">1534 m</quantity>
) in
<collectingCountry id="F37603A3B2689C39FD3FFD82FC96FD85" box="[687,772,583,609]" name="Bolivia" pageId="5" pageNumber="366">Bolivia</collectingCountry>
,
<collectingRegion id="49A58DD1B2689C39FC80FD82FB89FD85" box="[784,1051,583,609]" country="Bolivia" name="Santa Cruz" pageId="5" pageNumber="366">Santa Cruz Department</collectingRegion>
, Florida Province, in municipality of Pampa Grande, and locality of El Millu. The collecting sites include transitions between Bolivian montane dry forest and the slopes of the northern cloud forests of Yungas ecoregion.
</paragraph>
</subSubSection>
<subSubSection id="C37B10B8B2689C39FF57FD76FC09FCF1" pageId="5" pageNumber="366" type="etymology">
<paragraph id="8BDE4333B2689C39FF57FD76FC09FCF1" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="366">
<emphasis id="B9159F21B2689C39FF57FD76FEDCFD29" bold="true" box="[199,334,691,717]" pageId="5" pageNumber="366">Etymology.</emphasis>
The specific epithet refers to the location of the collecting place, an area between inter-Andean dry valleys of the Bolivian montane dry forest and the slopes of the northern Bolivian Andes, a northern Yungas forest. This location has enormous interest from a conservation perspective.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="65DDA52592D676EE9A4DC5CB5FB11016">Three new species and phylogenetic affinity of the neotropical genus Sericobracon Shaw (Braconidae: Doryctinae)</mods:title>
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<paragraph id="8BC136B91510FFE2FF23FAB4FC61FAA5" blockId="7.[151,918,1374,1437]" box="[151,918,1374,1401]" pageId="7" pageNumber="178">
<heading id="D08981D51510FFE2FF23FAB4FC61FAA5" bold="true" box="[151,918,1374,1401]" fontSize="11" level="1" pageId="7" pageNumber="178" reason="1">
<emphasis id="B90AEAAB1510FFE2FF23FAB4FC61FAA5" bold="true" box="[151,918,1374,1401]" pageId="7" pageNumber="178">
<taxonomicName id="4C7E4D3A1510FFE2FF23FAB4FD0DFAA5" authority="Zaldivar-Riveron &amp; Shaw" authorityName="Zaldivar-Riveron &amp; Shaw" authorityYear="2025" box="[151,762,1374,1401]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="puravida" status="sp. nov.">
<emphasis id="B90AEAAB1510FFE2FF23FAB4FE5BFAA4" bold="true" box="[151,428,1374,1401]" italics="true" pageId="7" pageNumber="178">Sericobracon puravida</emphasis>
Zaldívar-Riverón &amp; Shaw
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,
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</emphasis>
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(
<figureCitation id="13452A3C1510FFE2FF2BFA69FEEBFA40" box="[159,284,1410,1437]" captionStart="FIGURE 5" captionStartId="8.[152,255,1904,1928]" captionTargetBox="[152,1435,561,1881]" captionTargetId="figure-189@8.[152,1435,561,1881]" captionTargetPageId="8" captionText="FIGURE 5. Sericobracon puravida sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216365" httpUri="https://zenodo.org/record/15216365/files/figure.png" pageId="7" pageNumber="178">Figs 5A–F</figureCitation>
).
</paragraph>
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<subSubSection id="C36465321510FFE2FF23FA20FF3DF98C" pageId="7" pageNumber="178" type="diagnosis">
<paragraph id="8BC136B91510FFE2FF23FA20FF3DF98C" blockId="7.[151,1437,1482,2013]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF23FA20FEFDFA39" bold="true" box="[151,266,1482,1508]" pageId="7" pageNumber="178">Diagnosis</emphasis>
.
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<emphasis id="B90AEAAB1510FFE2FEA8FA21FDD5FA38" box="[284,546,1483,1509]" italics="true" pageId="7" pageNumber="178">Sericobracon puravida</emphasis>
</taxonomicName>
<emphasis id="B90AEAAB1510FFE2FD98FA21FD7EFA38" bold="true" box="[556,649,1483,1509]" pageId="7" pageNumber="178">
<taxonomicNameLabel id="A23957D01510FFE2FD98FA21FD7EFA38" box="[556,649,1483,1509]" pageId="7" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C7E4D3A1510FFE2FC7EFA21FB9EFA38" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[970,1129,1483,1509]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="paulmarshi">
<emphasis id="B90AEAAB1510FFE2FC7EFA21FB9EFA38" box="[970,1129,1483,1509]" italics="true" pageId="7" pageNumber="178">S. paulmarshi</emphasis>
</taxonomicName>
by the diagnostic features mentioned above for the latter species. It can be also distinguished from
<taxonomicName id="4C7E4D3A1510FFE2FC7DFA05FB97F9D5" box="[969,1120,1519,1545]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="arimaensis">
<emphasis id="B90AEAAB1510FFE2FC7DFA05FB97F9D5" box="[969,1120,1519,1545]" italics="true" pageId="7" pageNumber="178">S. arimaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7E4D3A1510FFE2FBDBFA05FB24F9D5" box="[1135,1235,1519,1545]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="evansi">
<emphasis id="B90AEAAB1510FFE2FBDBFA05FB24F9D5" box="[1135,1235,1519,1545]" italics="true" pageId="7" pageNumber="178">S. evansi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E4D3A1510FFE2FABEFA05FA8FF9D5" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[1290,1400,1519,1545]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="zunigai">
<emphasis id="B90AEAAB1510FFE2FABEFA05FA8FF9D5" box="[1290,1400,1519,1545]" italics="true" pageId="7" pageNumber="178">S. zunigai</emphasis>
</taxonomicName>
by having a mostly smooth mesopleuron (
<figureCitation id="13452A3C1510FFE2FDE4F9F9FD30F9F0" box="[592,711,1554,1581]" captionStart="FIGURE 5" captionStartId="8.[152,255,1904,1928]" captionTargetBox="[152,1435,561,1881]" captionTargetId="figure-189@8.[152,1435,561,1881]" captionTargetPageId="8" captionText="FIGURE 5. Sericobracon puravida sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216365" httpUri="https://zenodo.org/record/15216365/files/figure.png" pageId="7" pageNumber="178">Figure 5D</figureCitation>
) (mostly coriaceous in the latter three species
<figureCitation id="13452A3C1510FFE2FB6DF9F9FAC1F9F0" box="[1241,1334,1555,1581]" captionStart="FIGURE 2" captionStartId="4.[152,255,1921,1945]" captionTargetBox="[152,1436,956,1896]" captionTargetId="figure-234@4.[152,1436,956,1896]" captionTargetPageId="4" captionText="FIGURE 2. Sericobracon arimaensis Shaw. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216359" httpUri="https://zenodo.org/record/15216359/files/figure.png" pageId="7" pageNumber="178">Figs 2D</figureCitation>
,
<figureCitation id="13452A3C1510FFE2FAF7F9F9FA91F9F0" box="[1347,1382,1555,1581]" captionStart="FIGURE 3" captionStartId="5.[152,255,1060,1084]" captionTargetBox="[151,1435,181,1036]" captionTargetId="figure-15@5.[151,1435,181,1036]" captionTargetPageId="5" captionText="FIGURE 3. Sericobracon evansi Shaw. Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216361" httpUri="https://zenodo.org/record/15216361/files/figure.png" pageId="7" pageNumber="178">3B</figureCitation>
,
<figureCitation id="13452A3C1510FFE2FAC7F9F9FA6FF9F0" box="[1395,1432,1555,1581]" captionStart="FIGURE 4" captionStartId="6.[152,255,1679,1703]" captionTargetBox="[177,1411,525,1655]" captionTargetId="figure-166@6.[177,1411,525,1655]" captionTargetPageId="6" captionText="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="7" pageNumber="178">4D</figureCitation>
,
<figureCitation id="13452A3C1510FFE2FF23F9DDFF4AF98C" box="[151,189,1591,1617]" captionStart="FIGURE 6" captionStartId="10.[152,255,1444,1468]" captionTargetBox="[152,1435,181,1420]" captionTargetId="figure-27@10.[152,1435,181,1420]" captionTargetPageId="10" captionText="FIGURE 6. Sericobracon zunigai sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216367" httpUri="https://zenodo.org/record/15216367/files/figure.png" pageId="7" pageNumber="178">6D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36465321510FFECFF73F9B0FE7FFE0C" lastPageId="9" lastPageNumber="180" pageId="7" pageNumber="178" type="description">
<paragraph id="8BC136B91510FFE2FF73F9B0FCD3F8F4" blockId="7.[151,1437,1482,2013]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73F9B0FEA6F9A9" bold="true" box="[199,337,1626,1652]" pageId="7" pageNumber="178">Description</emphasis>
.
<typeStatus id="54C5881B1510FFE2FED6F9B1FE3DF9A8" box="[354,458,1627,1653]" pageId="7" pageNumber="178" type="holotype">Holotype</typeStatus>
female. Body size
<quantity id="4C869B5C1510FFE2FD1EF9B1FCF2F9A8" box="[682,773,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="178" unit="mm" value="2.9">2.9 mm</quantity>
, fore wing
<quantity id="4C869B5C1510FFE2FC3EF9B0FC12F9A8" box="[906,997,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="178" unit="mm" value="2.5">2.5 mm</quantity>
.
<emphasis id="B90AEAAB1510FFE2FC46F9B1FBB6F9A8" box="[1010,1089,1627,1653]" italics="true" pageId="7" pageNumber="178">Colour</emphasis>
: vertex and frons brown, area surrounding eyes honey yellow; temple gena and face brown to honey yellow; mesosoma mostly brown to dark brown; first metasomal tergite brown; second tergite brown medially, remaining area pale yellow; third to fifth tergites mostly brown, lateral area pale yellow; remaining tergites mostly pale yellow with median small, brown diffused patches. Wings hyaline, pterostigma and veins honey yellow. Legs pale yellow to honey yellow. Ovipositor honey yellow, strongly sclerotised at apex; sheaths brown.
</paragraph>
<paragraph id="8BC136B91510FFEDFF73F8D9FD35FF08" blockId="7.[151,1437,1482,2013]" lastBlockId="8.[151,1437,151,537]" lastPageId="8" lastPageNumber="179" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73F8D9FEF3F890" box="[199,260,1843,1869]" italics="true" pageId="7" pageNumber="178">Head</emphasis>
: transverse in dorsal view, 2.5 × wider than its median length. Eye 1.3 × longer than wide; eye width 1.8 × longer than temple in dorsal view. Malar space 0.4 × eye height and as long as width of hypoclypeal depression; mandibles bidentate. Occipital carina complete and reaching hypostomal carina before the base of mandible; inner margins of posterior ocelli (POL) as long as the ocellar diameter (OD), 0.4 × shorter than distance between outer margin of posterior ocellus and eye (OOL). Vertex slightly striate, frons and temple smooth, frons slightly excavated, face slightly rugose, setose. Antennae broken, with 11 flagellomeres or less; first flagellomere about 4.0 × longer than wide, 1.3 × longer than second flagellomere.
</paragraph>
<paragraph id="8BC136B9151FFFEDFF73FF35FB4DFDC4" blockId="8.[151,1437,151,537]" pageId="8" pageNumber="179">
<emphasis id="B90AEAAB151FFFEDFF73FF35FEC8FF25" box="[199,319,223,248]" italics="true" pageId="8" pageNumber="179">Mesosoma</emphasis>
: 1.7 × longer than high and 2.2 × longer than wide. Pronotal collar distinct, visible in dorsal view, smooth to slightly rugose, sparsely setose; pronotal furrow wide, deep and scrobiculate. Median length of mesoscutum 1.2 × its width (dorsal view); mesoscutal lobes coriaceous, with median mesoscutal lobe slightly projected upwards from lateral lobes; notauli wide, deep, scrobiculate, not joining, obscuring before the endo of mesoscutum in a longitudinally rugose area. Scutellar disc smooth to slightly acinose, prescutellar furrow with six transverse carinae. Mesopleuron mostly smooth; precoxal sulcus wide, deep, slightly scrobiculate, 0.6 × as long as mesopleuron; mesopleural sulcus wide, relatively shallow and scrobiculate, subalar sulcus deep and scrobiculate. Metapleuron rugose-areolate. Propodeum slightly rugose-smooth on median basal half, rugose-areolate on apical half, with a median longitudinal carina running along basal third, areola distinct and complete.
</paragraph>
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<paragraph id="8BC136B9151FFFEDFF2CF89AFBBDF870" blockId="8.[151,1437,1904,1965]" pageId="8" pageNumber="179">
<emphasis id="B90AEAAB151FFFEDFF2CF89AFEEDF854" bold="true" box="[152,282,1904,1929]" pageId="8" pageNumber="179">FIGURE 5.</emphasis>
<taxonomicName id="4C7E4D3A151FFFEDFE96F89BFDFAF854" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[290,525,1905,1929]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="179" phylum="Arthropoda" rank="species" species="puravida" status="sp. nov.">
<emphasis id="B90AEAAB151FFFEDFE96F89BFDFAF854" box="[290,525,1905,1929]" italics="true" pageId="8" pageNumber="179">Sericobracon puravida</emphasis>
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<emphasis id="B90AEAAB151FFFEDFDA1F89BFD90F854" bold="true" box="[533,615,1905,1929]" pageId="8" pageNumber="179">
<taxonomicNameLabel id="A23957D0151FFFEDFDA1F89BFD90F854" box="[533,615,1905,1929]" pageId="8" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing.
</paragraph>
</caption>
<paragraph id="8BC136B9151EFFECFF73FF7DFA82FF24" blockId="9.[151,1437,150,681]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FF7DFEFCFF6D" box="[199,267,151,176]" italics="true" pageId="9" pageNumber="180">Wings</emphasis>
: fore wing 2.9 × longer than wide. Pterostigma 3.2 × longer than wide and 1.5 × as long as R. Vein r 0.9 × as long as 3RSa, 0.2 × as long as 3RSb, and as long as vein r-m. Vein m-cu antefurcal to 2RS, vein RS+Mb present. First subdiscal cell open. Hind wing vein M+CU 0.8 × as long as 1M, m-cu slightly curved towards wing apex.
</paragraph>
<paragraph id="8BC136B9151EFFECFF73FEE9FD2EFE9C" blockId="9.[151,1437,150,681]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FEE9FF0BFEC1" box="[199,252,259,284]" italics="true" pageId="9" pageNumber="180">Legs</emphasis>
: one missing, remaining fore tibia without row of spines along anterior margin. Legs mostly smooth to slightly acinose; hind coxa slightly striate ventrally.
</paragraph>
<paragraph id="8BC136B9151EFFECFF73FEA1FC61FE70" blockId="9.[151,1437,150,681]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FEA1FECBFEB9" box="[199,316,331,356]" italics="true" pageId="9" pageNumber="180">Metasoma</emphasis>
: first tergite short, 0.9 × as long as apical width, longitudinally costate. Second tergite longitudinally costate; suture between second and third tergites straight, distinct; remaining tergites mostly smooth and polished, slightly acinose apically. Ovipositor length 0.7 as long as metasoma.
</paragraph>
<paragraph id="8BC136B9151EFFECFF73FE5DFE7FFE0C" blockId="9.[151,1437,150,681]" box="[199,392,439,465]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FE5DFEFCFE0C" bold="true" box="[199,267,439,465]" pageId="9" pageNumber="180">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3646532151EFFECFF73FE30FE53FE28" box="[199,420,474,501]" pageId="9" pageNumber="180" type="biology_ecology">
<paragraph id="8BC136B9151EFFECFF73FE30FE53FE28" blockId="9.[151,1437,150,681]" box="[199,420,474,501]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FE30FED0FE29" bold="true" box="[199,295,474,500]" pageId="9" pageNumber="180">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3646532151EFFECFF73FE15FD6FFDBC" pageId="9" pageNumber="180" type="etymology">
<paragraph id="8BC136B9151EFFECFF73FE15FD6FFDBC" blockId="9.[151,1437,150,681]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FE15FEBFFDC4" bold="true" box="[199,328,511,537]" pageId="9" pageNumber="180">Etymology</emphasis>
. The name of this species is formed as an arbitrary combination of letters and refers to the popular Costa Rican expression “¡pura vida!”, meaning “this is life!”, which originally comes from the mid-twentieth century Mexican film comedian “Clavillazo”.
</paragraph>
</subSubSection>
<subSubSection id="C3646532151EFFECFF73FD80FD3CFD74" pageId="9" pageNumber="180" type="materials_examined">
<paragraph id="8BC136B9151EFFECFF73FD80FD3CFD74" blockId="9.[151,1437,150,681]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FD80FE5AFD59" bold="true" box="[199,429,618,644]" pageId="9" pageNumber="180">Material examined</emphasis>
.
<materialsCitation id="3B163CE4151EFFECFE74FD81FD3CFD74" collectingDate="1993-08" collectionCode="CNIN, IBUNAM" collectorName="P. Hanson." country="Costa Rica" location="Malaise" pageId="9" pageNumber="180" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54C5881B151EFFECFE74FD81FDDFFD58" box="[448,552,619,645]" pageId="9" pageNumber="180" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED6FAE7C151EFFECFD8FFD81FD76FD58" box="[571,641,619,645]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="9" pageNumber="180" type="">CNIN</collectionCode>
<collectionCode id="ED6FAE7C151EFFECFD38FD81FCF4FD58" box="[652,771,619,645]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="9" pageNumber="180" type="University or college">IBUNAM</collectionCode>
). Female.
<collectingCountry id="F3697629151EFFECFCC9FD81FBF7FD58" box="[893,1024,619,645]" name="Costa Rica" pageId="9" pageNumber="180">Costa Rica</collectingCountry>
:
<collectingRegion id="49BAF85B151EFFECFBBAFD81FB7BFD58" box="[1038,1164,619,645]" country="Costa Rica" name="Puntarenas" pageId="9" pageNumber="180">Puntarenas</collectingRegion>
, Pen. Osa,
<locationDeviation id="9A732F9B151EFFECFAA1FD80FF01FD74" location="Npto. Jimenez" pageId="9" pageNumber="180">
<quantity id="4C869B5C151EFFECFAA1FD80FAA5FD58" box="[1301,1362,618,645]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="180" unit="km" value="5.0">5 km</quantity>
Npto. Jiménez
</locationDeviation>
, 10, m.
<location id="8EA16062151EFFECFEFBFD65FE5DFD74" LSID="urn:lsid:plazi:treatment:03D787AF1510FFECFF23FAB4FD3CFD74:8EA16062151EFFECFEFBFD65FE5DFD74" box="[335,426,655,681]" country="Costa Rica" name="Malaise" pageId="9" pageNumber="180" stateProvince="Puntarenas">Malaise</location>
, vii,
<date id="FFC01079151EFFECFE55FD65FDBFFD74" box="[481,584,655,681]" pageId="9" pageNumber="180" value="1993-08">
<collectingDate id="EF84E991151EFFECFE55FD65FDBFFD74" box="[481,584,655,681]" pageId="9" pageNumber="180" value="1993-08">viii 1993</collectingDate>
</date>
.
<collectorName id="268B536F151EFFECFDE7FD65FD3CFD74" box="[595,715,655,681]" pageId="9" pageNumber="180">P. Hanson.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
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<treatment id="03D787AF1511FFE2FF23FF7DFB20FAD0" ID-DOI="http://doi.org/10.5281/zenodo.15216300" ID-Zenodo-Dep="15216300" LSID="urn:lsid:plazi:treatment:03D787AF1511FFE2FF23FF7DFB20FAD0" httpUri="http://treatment.plazi.org/id/03D787AF1511FFE2FF23FF7DFB20FAD0" lastPageId="7" lastPageNumber="178" pageId="6" pageNumber="177">
<subSubSection id="C36465321511FFE3FF23FF7DFC42FF6C" box="[151,949,150,178]" pageId="6" pageNumber="177" type="nomenclature">
<paragraph id="8BC136B91511FFE3FF23FF7DFC42FF6C" blockId="6.[151,949,150,213]" box="[151,949,150,178]" pageId="6" pageNumber="177">
<heading id="D08981D51511FFE3FF23FF7DFC42FF6C" bold="true" box="[151,949,150,178]" fontSize="11" level="1" pageId="6" pageNumber="177" reason="1">
<emphasis id="B90AEAAB1511FFE3FF23FF7DFC42FF6C" bold="true" box="[151,949,150,178]" pageId="6" pageNumber="177">
<taxonomicName id="4C7E4D3A1511FFE3FF23FF7DFCEEFF6C" authority="Zaldivar-Riveron &amp; Shaw" authorityName="Zaldivar-Riveron &amp; Shaw" authorityYear="2025" box="[151,793,150,178]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="paulmarshi" status="sp. nov.">
<emphasis id="B90AEAAB1511FFE3FF23FF7DFE3CFF6F" bold="true" box="[151,459,151,178]" italics="true" pageId="6" pageNumber="177">Sericobracon paulmarshi</emphasis>
Zaldívar-Riverón &amp; Shaw
</taxonomicName>
,
<taxonomicNameLabel id="A23957D01511FFE3FC91FF7DFC42FF6C" box="[805,949,151,177]" pageId="6" pageNumber="177" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36465321511FFE3FF23FF51FEE9FF08" box="[151,286,187,213]" pageId="6" pageNumber="177" type="description">
<paragraph id="8BC136B91511FFE3FF23FF51FEE9FF08" blockId="6.[151,949,150,213]" box="[151,286,187,213]" pageId="6" pageNumber="177">
(
<figureCitation id="13452A3C1511FFE3FF2BFF51FF11FF08" box="[159,230,187,213]" captionStart="FIGURE 4" captionStartId="6.[152,255,1679,1703]" captionTargetBox="[177,1411,525,1655]" captionTargetId="figure-166@6.[177,1411,525,1655]" captionTargetPageId="6" captionText="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="6" pageNumber="177">Figs 4</figureCitation>
A-E)
</paragraph>
</subSubSection>
<subSubSection id="C36465321511FFE3FF23FEE9FC5AFE28" pageId="6" pageNumber="177" type="diagnosis">
<paragraph id="8BC136B91511FFE3FF23FEE9FC5AFE28" blockId="6.[151,1437,259,502]" pageId="6" pageNumber="177">
<emphasis id="B90AEAAB1511FFE3FF23FEE9FEFDFEC0" bold="true" box="[151,266,259,285]" pageId="6" pageNumber="177">Diagnosis</emphasis>
.
<taxonomicName id="4C7E4D3A1511FFE3FEADFEE9FDCFFEC0" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[281,568,259,285]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="paulmarshi" status="sp. nov.">
<emphasis id="B90AEAAB1511FFE3FEADFEE9FDCFFEC0" box="[281,568,259,285]" italics="true" pageId="6" pageNumber="177">Sericobracon paulmarshi</emphasis>
</taxonomicName>
<emphasis id="B90AEAAB1511FFE3FD8BFEE9FD6EFEC0" bold="true" box="[575,665,259,285]" pageId="6" pageNumber="177">
<taxonomicNameLabel id="A23957D01511FFE3FD8BFEE9FD6EFEC0" box="[575,665,259,285]" pageId="6" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be morphologically distinguished from the remaining described species of its genus by having: 1) the posterior margin of the longitudinally striate of the second metasomal tergite ending in a broad “V” shape (striate area shortest medially) (
<figureCitation id="13452A3C1511FFE3FCF3FEA1FC6BFEB8" box="[839,924,331,357]" captionStart="FIGURE 4" captionStartId="6.[152,255,1679,1703]" captionTargetBox="[177,1411,525,1655]" captionTargetId="figure-166@6.[177,1411,525,1655]" captionTargetPageId="6" captionText="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="6" pageNumber="177">Fig. 4E</figureCitation>
) (striate area ending with a straight border in the remaining species) (
<figureCitation id="13452A3C1511FFE3FE2BFE85FE0CFE54" box="[415,507,367,393]" captionStart="FIGURE 3" captionStartId="5.[152,255,1060,1084]" captionTargetBox="[151,1435,181,1036]" captionTargetId="figure-15@5.[151,1435,181,1036]" captionTargetPageId="5" captionText="FIGURE 3. Sericobracon evansi Shaw. Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216361" httpUri="https://zenodo.org/record/15216361/files/figure.png" pageId="6" pageNumber="177">Figs 3D</figureCitation>
,
<figureCitation id="13452A3C1511FFE3FDB2FE85FDD0FE57" box="[518,551,367,394]" captionStart="FIGURE 5" captionStartId="8.[152,255,1904,1928]" captionTargetBox="[152,1435,561,1881]" captionTargetId="figure-189@8.[152,1435,561,1881]" captionTargetPageId="8" captionText="FIGURE 5. Sericobracon puravida sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216365" httpUri="https://zenodo.org/record/15216365/files/figure.png" pageId="6" pageNumber="177">5E</figureCitation>
,
<figureCitation id="13452A3C1511FFE3FD86FE85FDA2FE54" box="[562,597,367,393]" captionStart="FIGURE 6" captionStartId="10.[152,255,1444,1468]" captionTargetBox="[152,1435,181,1420]" captionTargetId="figure-27@10.[152,1435,181,1420]" captionTargetPageId="10" captionText="FIGURE 6. Sericobracon zunigai sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216367" httpUri="https://zenodo.org/record/15216367/files/figure.png" pageId="6" pageNumber="177">6E</figureCitation>
); 2) notauli obscuring before end of mesoscutum in a median longitudinally striate area (
<figureCitation id="13452A3C1511FFE3FEABFE79FE83FE70" box="[287,372,403,429]" captionStart="FIGURE 4" captionStartId="6.[152,255,1679,1703]" captionTargetBox="[177,1411,525,1655]" captionTargetId="figure-166@6.[177,1411,525,1655]" captionTargetPageId="6" captionText="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="6" pageNumber="177">Fig. 4B</figureCitation>
) (similar in
<taxonomicName id="4C7E4D3A1511FFE3FE48FE79FD9EFE70" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[508,617,403,429]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="zunigai" status="sp. nov.">
<emphasis id="B90AEAAB1511FFE3FE48FE79FD9EFE70" box="[508,617,403,429]" italics="true" pageId="6" pageNumber="177">S. zunigai</emphasis>
</taxonomicName>
<emphasis id="B90AEAAB1511FFE3FDDBFE79FD3FFE70" bold="true" box="[623,712,403,429]" pageId="6" pageNumber="177">
<taxonomicNameLabel id="A23957D01511FFE3FDDBFE79FD3FFE70" box="[623,712,403,429]" pageId="6" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13452A3C1511FFE3FD62FE79FCD9FE70" box="[726,814,403,429]" captionStart="FIGURE 6" captionStartId="10.[152,255,1444,1468]" captionTargetBox="[152,1435,181,1420]" captionTargetId="figure-27@10.[152,1435,181,1420]" captionTargetPageId="10" captionText="FIGURE 6. Sericobracon zunigai sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216367" httpUri="https://zenodo.org/record/15216367/files/figure.png" pageId="6" pageNumber="177">Fig. 6C</figureCitation>
); in a slightly rugose area in the remaining species (
<figureCitation id="13452A3C1511FFE3FADFFE79FF4DFE0C" captionStart="FIGURE 2" captionStartId="4.[152,255,1921,1945]" captionTargetBox="[152,1436,956,1896]" captionTargetId="figure-234@4.[152,1436,956,1896]" captionTargetPageId="4" captionText="FIGURE 2. Sericobracon arimaensis Shaw. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216359" httpUri="https://zenodo.org/record/15216359/files/figure.png" pageId="6" pageNumber="177">Figs 2C</figureCitation>
,
<figureCitation id="13452A3C1511FFE3FF71FE5DFF1FFE0C" box="[197,232,439,465]" captionStart="FIGURE 3" captionStartId="5.[152,255,1060,1084]" captionTargetBox="[151,1435,181,1036]" captionTargetId="figure-15@5.[151,1435,181,1036]" captionTargetPageId="5" captionText="FIGURE 3. Sericobracon evansi Shaw. Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216361" httpUri="https://zenodo.org/record/15216361/files/figure.png" pageId="6" pageNumber="177">3C</figureCitation>
,
<figureCitation id="13452A3C1511FFE3FF40FE5DFEE1FE0C" box="[244,278,439,465]" captionStart="FIGURE 4" captionStartId="6.[152,255,1679,1703]" captionTargetBox="[177,1411,525,1655]" captionTargetId="figure-166@6.[177,1411,525,1655]" captionTargetPageId="6" captionText="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="6" pageNumber="177">4B</figureCitation>
,
<figureCitation id="13452A3C1511FFE3FE96FE5DFEB3FE0F" box="[290,324,439,466]" captionStart="FIGURE 5" captionStartId="8.[152,255,1904,1928]" captionTargetBox="[152,1435,561,1881]" captionTargetId="figure-189@8.[152,1435,561,1881]" captionTargetPageId="8" captionText="FIGURE 5. Sericobracon puravida sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216365" httpUri="https://zenodo.org/record/15216365/files/figure.png" pageId="6" pageNumber="177">5C</figureCitation>
)); and 3) mesosoma and first metasomal tergite dark brown to black (4A, B, D, E) (brown or lighter in the remaining species;
<figureCitation id="13452A3C1511FFE3FE01FE31FDA2FE2B" box="[437,597,475,502]" captionStart="FIGURE 2" captionStartId="4.[152,255,1921,1945]" captionTargetBox="[152,1436,956,1896]" captionTargetId="figure-234@4.[152,1436,956,1896]" captionTargetPageId="4" captionText="FIGURE 2. Sericobracon arimaensis Shaw. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216359" httpUri="https://zenodo.org/record/15216359/files/figure.png" pageId="6" pageNumber="177">Figs 2A, C, D</figureCitation>
,
<figureCitation id="13452A3C1511FFE3FDD4FE31FD9BFE28" box="[608,620,475,501]" captionStart="FIGURE 3" captionStartId="5.[152,255,1060,1084]" captionTargetBox="[151,1435,181,1036]" captionTargetId="figure-15@5.[151,1435,181,1036]" captionTargetPageId="5" captionText="FIGURE 3. Sericobracon evansi Shaw. Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216361" httpUri="https://zenodo.org/record/15216361/files/figure.png" pageId="6" pageNumber="177">3</figureCitation>
A-D; 5A, C, D,E; 6A, C, E).
</paragraph>
</subSubSection>
<caption id="DF0166311511FFE3FF2CF965FBFEF916" ID-DOI="http://doi.org/10.5281/zenodo.15216363" ID-Zenodo-Dep="15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="6" pageNumber="177" startId="6.[152,255,1679,1703]" targetBox="[177,1411,525,1655]" targetPageId="6" targetType="figure">
<paragraph id="8BC136B91511FFE3FF2CF965FBFEF916" blockId="6.[151,1436,1679,1739]" pageId="6" pageNumber="177">
<emphasis id="B90AEAAB1511FFE3FF2CF965FEE0F97A" bold="true" box="[152,279,1679,1703]" pageId="6" pageNumber="177">FIGURE 4.</emphasis>
<taxonomicName id="4C7E4D3A1511FFE3FEA8F965FDEAF97A" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[284,541,1679,1703]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="paulmarshi" status="sp. nov.">
<emphasis id="B90AEAAB1511FFE3FEA8F965FDEAF97A" box="[284,541,1679,1703]" italics="true" pageId="6" pageNumber="177">Sericobracon paulmarshi</emphasis>
</taxonomicName>
<emphasis id="B90AEAAB1511FFE3FD97F965FD85F97A" bold="true" box="[547,626,1679,1703]" pageId="6" pageNumber="177">
<taxonomicNameLabel id="A23957D01511FFE3FD97F965FD85F97A" box="[547,626,1679,1703]" pageId="6" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, female. A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view.
</paragraph>
</caption>
<subSubSection id="C36465321511FFE2FF73F8F7FE7FFC30" lastPageId="7" lastPageNumber="178" pageId="6" pageNumber="177" type="description">
<paragraph id="8BC136B91511FFE3FF73F8F7FE68F87E" blockId="6.[151,1437,1821,2028]" pageId="6" pageNumber="177">
<emphasis id="B90AEAAB1511FFE3FF73F8F7FEA6F8EA" bold="true" box="[199,337,1821,1847]" pageId="6" pageNumber="177">Description</emphasis>
.
<typeStatus id="54C5881B1511FFE3FED4F8F7FE3FF8EA" box="[352,456,1821,1847]" pageId="6" pageNumber="177" type="holotype">Holotype</typeStatus>
female. Body size
<quantity id="4C869B5C1511FFE3FD2BF8F4FD00F8E5" box="[671,759,1822,1848]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="177" unit="mm" value="3.7">3.7 mm</quantity>
, fore wing
<quantity id="4C869B5C1511FFE3FCC1F8F4FC39F8E5" box="[885,974,1822,1848]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="177" unit="mm" value="3.1">3.1 mm</quantity>
.
<emphasis id="B90AEAAB1511FFE3FC6DF8F7FBDFF8EA" box="[985,1064,1821,1847]" italics="true" pageId="6" pageNumber="177">Colour</emphasis>
: vertex, frons and temple brown, face honey yellow, mesosoma and first metasomal tergites dark brown to black, remaining tergites brown to dark brown. Wings hyaline, pterostigma and veins brown. Legs pale yellow. Ovipositor honey yellow, strongly sclerotised at apex; sheaths brown.
</paragraph>
<paragraph id="8BC136B91511FFE2FF73F847FE40FEC0" blockId="6.[151,1437,1821,2028]" lastBlockId="7.[151,1437,151,1293]" lastPageId="7" lastPageNumber="178" pageId="6" pageNumber="177">
<emphasis id="B90AEAAB1511FFE3FF73F847FEF3F81A" box="[199,260,1965,1991]" italics="true" pageId="6" pageNumber="177">Head</emphasis>
: transverse in dorsal view, 1.75 × wider than its median length. Eye 1.2 × longer than wide; eye width.0 × longer than temple in dorsal view. Malar space 0.3 × eye height and 0.9 as long as width of hypoclypeal depression. Inner margins of posterior ocelli (POL) as long as the ocellar diameter (OD), 2.0 × shorter than distance between outer margin of posterior ocellus and eye (OOL). Vertex striate; temple, frons, and face smooth, frons with short sparse setae. Antennae broken, with 15 flagellomeres; first flagellomere about 3.0 × longer than wide, 1.2 × longer than second flagellomere.
</paragraph>
<paragraph id="8BC136B91510FFE2FF73FECDFA8DFDBC" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FECDFEC8FE9D" box="[199,319,295,320]" italics="true" pageId="7" pageNumber="178">Mesosoma</emphasis>
: 1.7 × longer than high and 1.6 × longer than wide. Pronotal collar short but visible in dorsal view, slightly rugose, sparsely setose; pronotal furrow wide, deep and scrobiculate. Median length of mesoscutum 1.7 × its width (dorsal view); mesoscutal lobes coriaceous, with median mesoscutal lobe slightly projected upwards from lateral lobes; notauli wide, deep, scrobiculate, not joining, obscuring before the endo of mesoscutum in a median longitudinally striate area. Scutellar disc smooth and polished, prescutellar furrow with eight transverse carinae. Mesopleuron smooth and polished; precoxal sulcus wide, deep, slightly scrobiculate, about 0.6 × as long as mesopleuron; mesopleural sulcus wide, deep and scrobiculate, subalar sulcus narrower but deep and scrobiculate. Metapleuron strongly rugose-areolate. Propodeum smooth-slightly coriaceous on median basal half, basal two thirds rugose-areolate, with a median longitudinal carina running along basal third, areola almost indistinct, complete.
</paragraph>
<paragraph id="8BC136B91510FFE2FF73FD81FF23FD2C" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FD81FEFCFD59" box="[199,267,619,644]" italics="true" pageId="7" pageNumber="178">Wings</emphasis>
: fore wing 3.6 × longer than wide. Pterostigma 3.7 × longer than wide and 0.6 × as long as R. Vein r X0.8 × as long as 3RSa, 0.2 × as long as 3RSb, and 0.7 × as long as vein r-m. Vein m-cu antefurcal to 2RS, vein RS+Mb present. First subdiscal cell open. Hind wing vein M+CU 0.8 × as long as 1M, m-cu slightly curved towards wing apex.
</paragraph>
<paragraph id="8BC136B91510FFE2FF73FD11FAF7FCC8" blockId="7.[151,1437,151,1293]" box="[199,1280,763,789]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FD11FF0BFCC9" box="[199,252,763,788]" italics="true" pageId="7" pageNumber="178">Legs</emphasis>
: fore tibia without row of spines along anterior margin. Hind femur 3.4 × longer than wide.
</paragraph>
<paragraph id="8BC136B91510FFE2FF73FCF5FE9FFC78" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FCF5FECBFCE5" box="[199,316,799,824]" italics="true" pageId="7" pageNumber="178">Metasoma</emphasis>
: first tergite short, 0.9 × as long as apical width, longitudinally striate. Second tergite with longitudinally striate area ending with a “V” shaped border, striae shortest medially; suture between second and third tergites distinct, slightly curved; remaining tergites smooth, polished to slightly acinose. Ovipositor length as long as metasoma.
</paragraph>
<paragraph id="8BC136B91510FFE2FF73FC44FB32FC14" blockId="7.[151,1437,151,1293]" box="[199,1221,942,969]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FC44FECFFC15" bold="true" box="[199,312,942,968]" pageId="7" pageNumber="178">Variation</emphasis>
. Body size
<quantity id="4C869B5C1510FFE2FE08FC45FDB1FC14" box="[444,582,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="7" pageNumber="178" unit="mm" value="3.8" valueMax="3.9" valueMin="3.7">3.7–3.9 mm</quantity>
. Thirty-one flagellomeres. Fore wing length
<quantity id="4C869B5C1510FFE2FB8FFC45FB36FC14" box="[1083,1217,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="7" pageNumber="178" unit="mm" value="3.15" valueMax="3.2" valueMin="3.1">3.1-3.2 mm</quantity>
.
</paragraph>
<paragraph id="8BC136B91510FFE2FF73FC39FE7FFC30" blockId="7.[151,1437,151,1293]" box="[199,392,979,1005]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FC39FEFCFC30" bold="true" box="[199,267,979,1005]" pageId="7" pageNumber="178">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36465321510FFE2FF73FC1CFE53FBCC" box="[199,420,1014,1041]" pageId="7" pageNumber="178" type="biology_ecology">
<paragraph id="8BC136B91510FFE2FF73FC1CFE53FBCC" blockId="7.[151,1437,151,1293]" box="[199,420,1014,1041]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FC1CFED0FBCD" bold="true" box="[199,295,1014,1040]" pageId="7" pageNumber="178">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36465321510FFE2FF73FBF1FBCCFB84" pageId="7" pageNumber="178" type="etymology">
<paragraph id="8BC136B91510FFE2FF73FBF1FBCCFB84" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FBF1FEBFFBE8" bold="true" box="[199,328,1051,1077]" pageId="7" pageNumber="178">Etymology</emphasis>
. We name this species in honour of our late colleague Dr. Paul Malcolm Marsh, who devoted his life to the taxonomic study of the braconid subfamily
<taxonomicName id="4C7E4D3A1510FFE2FD46FBD5FC99FB84" box="[754,878,1087,1113]" class="Insecta" family="Braconidae" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="subFamily" subFamily="Doryctinae">Doryctinae</taxonomicName>
for over 50 years.
</paragraph>
</subSubSection>
<subSubSection id="C36465321510FFE2FF73FB88FB20FAD0" pageId="7" pageNumber="178" type="materials_examined">
<paragraph id="8BC136B91510FFE2FF73FB88FB20FAD0" blockId="7.[151,1437,151,1293]" pageId="7" pageNumber="178">
<emphasis id="B90AEAAB1510FFE2FF73FB88FE5EFBA1" bold="true" box="[199,425,1122,1148]" pageId="7" pageNumber="178">Material examined</emphasis>
.
<materialsCitation id="3B163CE41510FFE2FE02FB89FEACFB18" collectingDate="1996-09" collectionCode="CNIN, IBUNAM, COI" collectorName="I Villegas &amp; Malaise, L. N. &amp; GenBank" country="Costa Rica" elevation="1075" location="Guanacaste" pageId="7" pageNumber="178" specimenCode="CNIN 4977" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="B90AEAAB1510FFE2FE02FB89FDDBFBA0" bold="true" box="[438,556,1123,1149]" pageId="7" pageNumber="178">
<typeStatus id="54C5881B1510FFE2FE02FB89FDD0FBA0" box="[438,551,1123,1149]" pageId="7" pageNumber="178" type="holotype">Holotype</typeStatus>
:
</emphasis>
(
<collectionCode id="ED6FAE7C1510FFE2FD8EFB89FD77FBA0" box="[570,640,1123,1149]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="7" pageNumber="178" type="">CNIN</collectionCode>
<collectionCode id="ED6FAE7C1510FFE2FD32FB89FD0EFBA0" box="[646,761,1123,1149]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="7" pageNumber="178" type="University or college">IBUNAM</collectionCode>
) Female.
<collectingCountry id="F36976291510FFE2FCD3FB89FC12FBA0" box="[871,997,1123,1149]" name="Costa Rica" pageId="7" pageNumber="178">Costa Rica</collectingCountry>
:
<collectingRegion id="49BAF85B1510FFE2FC5AFB89FB85FBA0" box="[1006,1138,1123,1149]" country="Costa Rica" name="Guanacaste" pageId="7" pageNumber="178">Guanacaste</collectingRegion>
, Est. Cacao,
<quantity id="4C869B5C1510FFE2FAB5FB88FA60FBA0" box="[1281,1431,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="7" pageNumber="178" unit="m" value="1075.0" valueMax="1150.0" valueMin="1000.0">
<elevation id="0053D18A1510FFE2FAB5FB88FA60FBA0" box="[1281,1431,1122,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="7" pageNumber="178" unit="m" value="1075.0" valueMax="1150.0" valueMin="1000.0">1000-1150 m</elevation>
</quantity>
,
<date id="FFC010791510FFE2FF23FB6DFF07FB7C" box="[151,240,1159,1185]" pageId="7" pageNumber="178" value="1996-09">
<collectingDate id="EF84E9911510FFE2FF23FB6DFF07FB7C" box="[151,240,1159,1185]" pageId="7" pageNumber="178" value="1996-09">ix 1996</collectingDate>
</date>
,
<collectorName id="268B536F1510FFE2FF48FB6DFE9CFB7C" box="[252,363,1159,1185]" pageId="7" pageNumber="178">I Villegas</collectorName>
,
<collectorName id="268B536F1510FFE2FEC3FB6DFDECFB7C" box="[375,539,1159,1185]" pageId="7" pageNumber="178">Malaise, L. N.</collectorName>
323150-375500 #4759. DNA voucher no.
<specimenCode id="DBD89EC21510FFE2FBB5FB6DFB7FFB7C" box="[1025,1160,1159,1185]" collectionCode="CNIN" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="7" pageNumber="178" type="">CNIN 4977</specimenCode>
,
<collectorName id="268B536F1510FFE2FB20FB6DFB09FB7C" box="[1172,1278,1159,1185]" pageId="7" pageNumber="178">GenBank</collectorName>
accession no. PV157332 (
<collectionCode id="ED6FAE7C1510FFE2FEAAFB41FEA3FB18" box="[286,340,1195,1221]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="7" pageNumber="178" type="Herbarium">COI</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B163CE41510FFE2FED1FB41FC9BFB18" box="[357,876,1195,1221]" collectingDate="1996-09" collectionCode="CNIN, IBUNAM, COI" collectorName="I Villegas &amp; Malaise, L. N. &amp; GenBank" country="Costa Rica" elevation="1075" location="Guanacaste" pageId="7" pageNumber="178" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="B90AEAAB1510FFE2FED1FB41FE2AFB18" bold="true" box="[357,477,1195,1221]" pageId="7" pageNumber="178">
<typeStatus id="54C5881B1510FFE2FED1FB41FE2AFB18" box="[357,477,1195,1221]" pageId="7" pageNumber="178" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D78FD301510FFE2FE5FFB41FD9EFB18" box="[491,617,1195,1221]" count="1" pageId="7" pageNumber="178" type="female">one female</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="3B163CE41510FFE2FCC2FB41FABAFB18" box="[886,1357,1195,1221]" collectingDate="1996-09" collectionCode="CNIN, IBUNAM, COI" collectorName="I Villegas &amp; Malaise, L. N. &amp; GenBank" country="Costa Rica" elevation="1075" location="Guanacaste" pageId="7" pageNumber="178" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D78FD301510FFE2FCC2FB41FC03FB18" box="[886,1012,1195,1221]" count="1" pageId="7" pageNumber="178" type="female">one female</specimenCount>
, same locality data as
<typeStatus id="54C5881B1510FFE2FB5FFB41FABAFB18" box="[1259,1357,1195,1221]" pageId="7" pageNumber="178" type="holotype">holotype</typeStatus>
</materialsCitation>
<materialsCitation id="3B163CE41510FFE2FAE7FB41FA91FB34" collectingDate="1990-06" collectorName="A. Solis &amp; Interseccion, L. N." country="Costa Rica" location="Cerro Cacao" pageId="7" pageNumber="178" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
except 1100-1200 meters,
<quantity id="4C869B5C1510FFE2FEC0FB25FE44FB34" box="[372,435,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="178" unit="km" value="2.0">2 km</quantity>
. SW de
<location id="8EA160621510FFE2FDA3FB25FD5DFB34" LSID="urn:lsid:plazi:treatment:03D787AF1511FFE2FF23FF7DFB20FAD0:8EA160621510FFE2FDA3FB25FD5DFB34" box="[535,682,1231,1257]" country="Costa Rica" name="Cerro Cacao" pageId="7" pageNumber="178" stateProvince="Guanacaste">Cerro Cacao</location>
,
<date id="FFC010791510FFE2FD03FB25FCE5FB34" box="[695,786,1231,1257]" pageId="7" pageNumber="178" value="1990-06">
<collectingDate id="EF84E9911510FFE2FD03FB25FCE5FB34" box="[695,786,1231,1257]" pageId="7" pageNumber="178" value="1990-06">vi 1990</collectingDate>
</date>
,
<collectorName id="268B536F1510FFE2FCAAFB25FC89FB34" box="[798,894,1231,1257]" pageId="7" pageNumber="178">A. Solis</collectorName>
,
<collectorName id="268B536F1510FFE2FC3FFB25FBAFFB34" box="[907,1112,1231,1257]" pageId="7" pageNumber="178">Interseccion, L.N.</collectorName>
323300-375700 #8556
</materialsCitation>
;
<materialsCitation id="3B163CE41510FFE2FAC0FB25FDB4FAD0" collectingDate="1990-06" collectorName="A. Solis &amp; Interseccion, L. N." country="Costa Rica" location="Cerro Cacao" pageId="7" pageNumber="178" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="paratype">
<specimenCount id="9D78FD301510FFE2FAC0FB25FF11FAD0" count="1" pageId="7" pageNumber="178" type="female">one female</specimenCount>
, same locality data as
<typeStatus id="54C5881B1510FFE2FE55FB19FDB4FAD0" box="[481,579,1267,1293]" pageId="7" pageNumber="178" type="holotype">holotype</typeStatus>
</materialsCitation>
except collected
<date id="FFC010791510FFE2FCB2FB19FC91FAD0" box="[774,870,1267,1293]" pageId="7" pageNumber="178" value="1996-07">vii 1996</date>
, A. Masis, Malaise trap, #47555.
</paragraph>
</subSubSection>
</treatment>
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157
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<paragraph id="8BC136B91513FFE1FF23FE14FDD3FDC5" blockId="4.[151,548,510,572]" box="[151,548,510,537]" pageId="4" pageNumber="175">
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Genus
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<emphasis id="B90AEAAB1513FFE1FF5AFE14FE78FDC4" bold="true" box="[238,399,510,537]" italics="true" pageId="4" pageNumber="175">Sericobracon</emphasis>
<bibRefCitation id="EFEF4B481513FFE1FE23FE14FDD3FDC5" author="Shaw, S. R. &amp; Edgerly, J. S." box="[407,548,510,536]" pageId="4" pageNumber="175" pagination="505 - 511" refId="ref9938" refString="Shaw, S. R. &amp; Edgerly, J. S. (1985) A new braconid genus (Hymenoptera) parasitizing webspinners (Embiidina) in Trinidad. Psyche, 92, 505-511. https://doi.org/10.1155/1985/54285" type="journal article" year="1985">Shaw, 1985</bibRefCitation>
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<paragraph id="8BC136B91513FFE1FF23FDC8FEE5FDE1" blockId="4.[151,548,510,572]" box="[151,274,546,572]" pageId="4" pageNumber="175">
(
<figureCitation id="13452A3C1513FFE1FF2BFDC8FEF2FDE1" box="[159,261,546,572]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="4.[152,255,1921,1945]" captionStartId-1="5.[152,255,1060,1084]" captionStartId-2="6.[152,255,1679,1703]" captionStartId-3="8.[152,255,1904,1928]" captionStartId-4="10.[152,255,1444,1468]" captionTargetBox-0="[152,1436,956,1896]" captionTargetBox-1="[151,1435,181,1036]" captionTargetBox-2="[177,1411,525,1655]" captionTargetBox-3="[152,1435,561,1881]" captionTargetBox-4="[152,1435,181,1420]" captionTargetId-0="figure-234@4.[152,1436,956,1896]" captionTargetId-1="figure-15@5.[151,1435,181,1036]" captionTargetId-2="figure-166@6.[177,1411,525,1655]" captionTargetId-3="figure-189@8.[152,1435,561,1881]" captionTargetId-4="figure-27@10.[152,1435,181,1420]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="10" captionText-0="FIGURE 2. Sericobracon arimaensis Shaw. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." captionText-1="FIGURE 3. Sericobracon evansi Shaw. Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." captionText-2="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." captionText-3="FIGURE 5. Sericobracon puravida sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing." captionText-4="FIGURE 6. Sericobracon zunigai sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing." figureDoi-0="http://doi.org/10.5281/zenodo.15216359" figureDoi-1="http://doi.org/10.5281/zenodo.15216361" figureDoi-2="http://doi.org/10.5281/zenodo.15216363" figureDoi-3="http://doi.org/10.5281/zenodo.15216365" figureDoi-4="http://doi.org/10.5281/zenodo.15216367" httpUri-0="https://zenodo.org/record/15216359/files/figure.png" httpUri-1="https://zenodo.org/record/15216361/files/figure.png" httpUri-2="https://zenodo.org/record/15216363/files/figure.png" httpUri-3="https://zenodo.org/record/15216365/files/figure.png" httpUri-4="https://zenodo.org/record/15216367/files/figure.png" pageId="4" pageNumber="175">Figs 2–6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36465321513FFE1FF23FD80FD6BFC78" pageId="4" pageNumber="175" type="diagnosis">
<paragraph id="8BC136B91513FFE1FF23FD80FD6BFC78" blockId="4.[151,1437,618,933]" pageId="4" pageNumber="175">
<emphasis id="B90AEAAB1513FFE1FF23FD80FEE5FD59" bold="true" box="[151,274,618,644]" pageId="4" pageNumber="175">Diagnosis.</emphasis>
Vertex striate. Clypeus concave exposing labrum. Maxillary and labial palpi 6-4 segmented. Mandibles bidentate. Occipital carinae present and complete. Occipital carina complete, reaching hypostomal carina before the base of mandible. Mesoscutal lobes mostly coriaceous, with median mesoscutal lobe slightly projected upwards from lateral lobes. Notauli complete and scrobiuculate. Prepectal carina present. Propodeum with a basal longitudinal carina followed by a distinct areola. Fore tibia without visible spines or pegs. Hind coxa with a distinct, small basoventral tubercle. Fore wing vein 2-SR complete. Hind wing vein m-cu usually slightly curved towards wing apex. First and second metasomal tergites sculptured, remaining tergites smooth and mostly polished; suture between second and third tergite distinct, straight or slightly curved; third tergite with a transverse straight furrow. Ovipositor usually long, 0.7–1.2 × as along as metasoma.
</paragraph>
</subSubSection>
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<paragraph id="8BC136B91513FFE1FF2CF86BFE4EF83C" blockId="4.[151,1437,1921,2017]" pageId="4" pageNumber="175">
<emphasis id="B90AEAAB1513FFE1FF2CF86BFEEDF844" bold="true" box="[152,282,1921,1945]" pageId="4" pageNumber="175">FIGURE 2.</emphasis>
<taxonomicName id="4C7E4D3A1513FFE1FE96F86BFD90F844" authority="Shaw." authorityName="Shaw." box="[290,615,1921,1945]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="arimaensis">
<emphasis id="B90AEAAB1513FFE1FE96F86BFDD6F845" box="[290,545,1921,1945]" italics="true" pageId="4" pageNumber="175">Sericobracon arimaensis</emphasis>
Shaw.
</taxonomicName>
Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College.
</paragraph>
</caption>
<caption id="DF0166311512FFE0FF2CFBCEFE13FB59" ID-DOI="http://doi.org/10.5281/zenodo.15216361" ID-Zenodo-Dep="15216361" httpUri="https://zenodo.org/record/15216361/files/figure.png" pageId="5" pageNumber="176" startId="5.[152,255,1060,1084]" targetBox="[151,1435,181,1036]" targetPageId="5" targetType="figure">
<paragraph id="8BC136B91512FFE0FF2CFBCEFE13FB59" blockId="5.[151,1436,1060,1156]" pageId="5" pageNumber="176">
<emphasis id="B90AEAAB1512FFE0FF2CFBCEFEEDFBE1" bold="true" box="[152,282,1060,1084]" pageId="5" pageNumber="176">FIGURE 3.</emphasis>
<taxonomicName id="4C7E4D3A1512FFE0FE96FBCEFDCDFBE1" authority="Shaw." authorityName="Shaw." box="[290,570,1060,1084]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="evansi">
<emphasis id="B90AEAAB1512FFE0FE96FBCEFE03FBE1" box="[290,500,1060,1084]" italics="true" pageId="5" pageNumber="176">Sericobracon evansi</emphasis>
Shaw.
</taxonomicName>
Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College.
</paragraph>
</caption>
<subSubSection id="C36465321512FFE0FF73FB3CFA92FAE5" pageId="5" pageNumber="176" type="distribution">
<paragraph id="8BC136B91512FFE0FF73FB3CFA92FAE5" blockId="5.[151,1437,1238,1841]" pageId="5" pageNumber="176">
<emphasis id="B90AEAAB1512FFE0FF73FB3CFE96FB2D" bold="true" box="[199,353,1238,1264]" pageId="5" pageNumber="176">Distribution.</emphasis>
The two already named species of
<taxonomicName id="4C7E4D3A1512FFE0FD41FB3CFC7BFB2D" authorityName="Shaw" authorityYear="1985" box="[757,908,1238,1264]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAAB1512FFE0FD41FB3CFC7BFB2D" box="[757,908,1238,1264]" italics="true" pageId="5" pageNumber="176">Sericobracon</emphasis>
</taxonomicName>
which were originally described for the genus occur in
<collectingCountry id="F36976291512FFE0FF4CFB10FEA0FAC9" box="[248,343,1274,1300]" name="Trinidad and Tobago" pageId="5" pageNumber="176">Trinidad</collectingCountry>
and the
<collectingCountry id="F36976291512FFE0FE02FB10FD7CFAC9" box="[438,651,1274,1300]" name="United States Virgin Islands" pageId="5" pageNumber="176">U.S. Virgin Islands</collectingCountry>
in the Caribbean Sea. The three new species of the genus described in this study occur in
<collectingCountry id="F36976291512FFE0FED8FAF4FE1EFAE5" box="[364,489,1310,1336]" name="Costa Rica" pageId="5" pageNumber="176">Costa Rica</collectingCountry>
, extending the known geographic distribution of this genus to Central America.
</paragraph>
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<subSubSection id="C36465321512FFE0FF73FAA8FCDAF8ED" pageId="5" pageNumber="176" type="discussion">
<paragraph id="8BC136B91512FFE0FF73FAA8FECBFA30" blockId="5.[151,1437,1238,1841]" pageId="5" pageNumber="176">
<emphasis id="B90AEAAB1512FFE0FF73FAA8FECCFA81" bold="true" box="[199,315,1346,1372]" pageId="5" pageNumber="176">Remarks.</emphasis>
Species of
<taxonomicName id="4C7E4D3A1512FFE0FE03FAA8FDB9FA81" authorityName="Shaw" authorityYear="1985" box="[439,590,1346,1372]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAAB1512FFE0FE03FAA8FDB9FA81" box="[439,590,1346,1372]" italics="true" pageId="5" pageNumber="176">Sericobracon</emphasis>
</taxonomicName>
can be morphologically distinguished from those belonging to the remaining New World doryctine genera by having the third and following metasomal tergites unsculptured and polished, presence of a transverse furrow on the third metasomal tergite, lacking pegs or stout spines on the foretibia, presence of a propodeal areola, lacking dorsal nodes at the ovipositor tip, and hind wing m-cu being slightly curved towards the wing apex.
</paragraph>
<paragraph id="8BC136B91512FFE0FF73FA1CFCDAF8ED" blockId="5.[151,1437,1238,1841]" pageId="5" pageNumber="176">
<taxonomicName id="4C7E4D3A1512FFE0FF73FA1CFEA9F9CD" authorityName="Shaw" authorityYear="1985" box="[199,350,1526,1552]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAAB1512FFE0FF73FA1CFEA9F9CD" box="[199,350,1526,1552]" italics="true" pageId="5" pageNumber="176">Sericobracon</emphasis>
</taxonomicName>
specimens seem to be only rarely encountered in museum collections for a couple of reasons. They are small wasps that occur only in tropical forest areas, and only (so far as is known) in close association with
<taxonomicName id="4C7E4D3A1512FFE0FF23F9D4FEEAF985" authorityName="Lameere" authorityYear="1900" box="[151,285,1598,1624]" class="Insecta" kingdom="Animalia" order="Embioptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="order">Embioptera</taxonomicName>
, which themselves are cryptic insects living hidden under sheets of spun silk. They probably spend most of their time living close to webspinners and do not disperse widely, and therefore are seldom seen, netted, or trapped. However, as it was previously noted (
<bibRefCitation id="EFEF4B481512FFE0FD7DF96CFC3BF97C" author="Shaw, S. R. &amp; Edgerly, J. S." box="[713,972,1670,1697]" pageId="5" pageNumber="176" pagination="505 - 511" refId="ref9938" refString="Shaw, S. R. &amp; Edgerly, J. S. (1985) A new braconid genus (Hymenoptera) parasitizing webspinners (Embiidina) in Trinidad. Psyche, 92, 505-511. https://doi.org/10.1155/1985/54285" type="journal article" year="1985">Shaw &amp; Edgerly 1985</bibRefCitation>
), these wasps are superficially similar in size and appearance to small doryctines such as
<taxonomicName id="4C7E4D3A1512FFE0FD0AF940FCBBF919" authorityName="Haliday" authorityYear="1836" box="[702,844,1706,1732]" class="Insecta" family="Braconidae" genus="Heterospilus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAAB1512FFE0FD0AF940FCBBF919" box="[702,844,1706,1732]" italics="true" pageId="5" pageNumber="176">Heterospilus</emphasis>
</taxonomicName>
, which are vastly more common, more diverse, and more widely dispersing species. Therefore, other specimens of
<taxonomicName id="4C7E4D3A1512FFE0FCC7F924FBFDF935" authorityName="Shaw" authorityYear="1985" box="[883,1034,1742,1768]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAAB1512FFE0FCC7F924FBFDF935" box="[883,1034,1742,1768]" italics="true" pageId="5" pageNumber="176">Sericobracon</emphasis>
</taxonomicName>
that may already exist in museum collections are likely to be still unidentified and hidden among the vast numbers of unidentified
<taxonomicName id="4C7E4D3A1512FFE0FB6AF918FA9BF8D1" authorityName="Haliday" authorityYear="1836" box="[1246,1388,1778,1804]" class="Insecta" family="Braconidae" genus="Heterospilus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAAB1512FFE0FB6AF918FA9BF8D1" box="[1246,1388,1778,1804]" italics="true" pageId="5" pageNumber="176">Heterospilus</emphasis>
</taxonomicName>
and other small doryctines that are accumulating in collections.
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="B90AEAAB151EFFECFF23FD05FE6DFCD7" bold="true" box="[151,410,751,778]" italics="true" pageId="9" pageNumber="180">Sericobracon zunigai</emphasis>
Zaldívar-Riverón &amp; Shaw
</taxonomicName>
,
<taxonomicNameLabel id="A23957D0151EFFECFD40FD1AFC73FCD7" box="[756,900,752,778]" pageId="9" pageNumber="180" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3646532151EFFECFF23FCFEFEEBFCF3" box="[151,284,788,814]" pageId="9" pageNumber="180" type="description">
<paragraph id="8BC136B9151EFFECFF23FCFEFEEBFCF3" blockId="9.[151,900,751,814]" box="[151,284,788,814]" pageId="9" pageNumber="180">
(
<figureCitation id="13452A3C151EFFECFF2BFCFEFF12FCF3" box="[159,229,788,814]" captionStart="FIGURE 6" captionStartId="10.[152,255,1444,1468]" captionTargetBox="[152,1435,181,1420]" captionTargetId="figure-27@10.[152,1435,181,1420]" captionTargetPageId="10" captionText="FIGURE 6. Sericobracon zunigai sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216367" httpUri="https://zenodo.org/record/15216367/files/figure.png" pageId="9" pageNumber="180">Figs 6</figureCitation>
A-F)
</paragraph>
</subSubSection>
<subSubSection id="C3646532151EFFECFF23FCB6FAC3FBF6" pageId="9" pageNumber="180" type="diagnosis">
<paragraph id="8BC136B9151EFFECFF23FCB6FAC3FBF6" blockId="9.[151,1437,860,2039]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF23FCB6FEFDFCAB" bold="true" box="[151,266,860,886]" pageId="9" pageNumber="180">Diagnosis</emphasis>
.
<taxonomicName id="4C7E4D3A151EFFECFEA2FCB6FDF3FCAB" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[278,516,860,886]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="zunigai" status="sp. nov.">
<emphasis id="B90AEAAB151EFFECFEA2FCB6FDF3FCAB" box="[278,516,860,886]" italics="true" pageId="9" pageNumber="180">Sericobracon zunigai</emphasis>
</taxonomicName>
<emphasis id="B90AEAAB151EFFECFDBDFCB6FD96FCAB" bold="true" box="[521,609,860,886]" pageId="9" pageNumber="180">
<taxonomicNameLabel id="A23957D0151EFFECFDBDFCB6FD96FCAB" box="[521,609,860,886]" pageId="9" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the remaining described species of
<taxonomicName id="4C7E4D3A151EFFECFAB1FCB6FA6BFCAB" authorityName="Shaw" authorityYear="1985" box="[1285,1436,860,886]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEAAB151EFFECFAB1FCB6FA6BFCAB" box="[1285,1436,860,886]" italics="true" pageId="9" pageNumber="180">Sericobracon</emphasis>
</taxonomicName>
by having: 1) propleuron, pronotum and most part of mesopleuron whitish yellow to honey yellow (
<figureCitation id="13452A3C151EFFECFAA6FC6AFA63FC47" box="[1298,1428,896,922]" captionStart="FIGURE 6" captionStartId="10.[152,255,1444,1468]" captionTargetBox="[152,1435,181,1420]" captionTargetId="figure-27@10.[152,1435,181,1420]" captionTargetPageId="10" captionText="FIGURE 6. Sericobracon zunigai sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216367" httpUri="https://zenodo.org/record/15216367/files/figure.png" pageId="9" pageNumber="180">Figs 6A, D</figureCitation>
) (mostly dark brown or black in the remaining species
<figureCitation id="13452A3C151EFFECFD58FC4EFC9EFC62" box="[748,873,932,959]" captionStart="FIGURE 2" captionStartId="4.[152,255,1921,1945]" captionTargetBox="[152,1436,956,1896]" captionTargetId="figure-234@4.[152,1436,956,1896]" captionTargetPageId="4" captionText="FIGURE 2. Sericobracon arimaensis Shaw. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216359" httpUri="https://zenodo.org/record/15216359/files/figure.png" pageId="9" pageNumber="180">Figs 2A, D</figureCitation>
,
<figureCitation id="13452A3C151EFFECFCC7FC4EFC4FFC62" box="[883,952,932,959]" captionStart="FIGURE 3" captionStartId="5.[152,255,1060,1084]" captionTargetBox="[151,1435,181,1036]" captionTargetId="figure-15@5.[151,1435,181,1036]" captionTargetPageId="5" captionText="FIGURE 3. Sericobracon evansi Shaw. Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216361" httpUri="https://zenodo.org/record/15216361/files/figure.png" pageId="9" pageNumber="180">3A, B</figureCitation>
,
<figureCitation id="13452A3C151EFFECFC77FC4EFBFEFC62" box="[963,1033,932,959]" captionStart="FIGURE 4" captionStartId="6.[152,255,1679,1703]" captionTargetBox="[177,1411,525,1655]" captionTargetId="figure-166@6.[177,1411,525,1655]" captionTargetPageId="6" captionText="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="9" pageNumber="180">4A, D</figureCitation>
,
<figureCitation id="13452A3C151EFFECFBA0FC4EFBADFC63" box="[1044,1114,932,959]" captionStart="FIGURE 5" captionStartId="8.[152,255,1904,1928]" captionTargetBox="[152,1435,561,1881]" captionTargetId="figure-189@8.[152,1435,561,1881]" captionTargetPageId="8" captionText="FIGURE 5. Sericobracon puravida sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216365" httpUri="https://zenodo.org/record/15216365/files/figure.png" pageId="9" pageNumber="180">5A, D</figureCitation>
); 3) notauli obscuring before the end of mesoscutum in a longitudinally striate- mostly smooth area (
<figureCitation id="13452A3C151EFFECFC76FC22FBEDFC3F" box="[962,1050,968,994]" captionStart="FIGURE 6" captionStartId="10.[152,255,1444,1468]" captionTargetBox="[152,1435,181,1420]" captionTargetId="figure-27@10.[152,1435,181,1420]" captionTargetPageId="10" captionText="FIGURE 6. Sericobracon zunigai sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216367" httpUri="https://zenodo.org/record/15216367/files/figure.png" pageId="9" pageNumber="180">Fig. 6C</figureCitation>
) (slightly rugose in the remaining species except
<taxonomicName id="4C7E4D3A151EFFECFEF2FC06FE12FBDB" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[326,485,1004,1030]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="paulmarshi">
<emphasis id="B90AEAAB151EFFECFEF2FC06FE12FBDB" box="[326,485,1004,1030]" italics="true" pageId="9" pageNumber="180">S. paulmarshi</emphasis>
</taxonomicName>
;
<figureCitation id="13452A3C151EFFECFE43FC06FDA3FBDB" box="[503,596,1004,1030]" captionStart="FIGURE 3" captionStartId="5.[152,255,1060,1084]" captionTargetBox="[151,1435,181,1036]" captionTargetId="figure-15@5.[151,1435,181,1036]" captionTargetPageId="5" captionText="FIGURE 3. Sericobracon evansi Shaw. Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216361" httpUri="https://zenodo.org/record/15216361/files/figure.png" pageId="9" pageNumber="180">Figs 3C</figureCitation>
,
<figureCitation id="13452A3C151EFFECFDD0FC06FD71FBDB" box="[612,646,1004,1030]" captionStart="FIGURE 4" captionStartId="6.[152,255,1679,1703]" captionTargetBox="[177,1411,525,1655]" captionTargetId="figure-166@6.[177,1411,525,1655]" captionTargetPageId="6" captionText="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="9" pageNumber="180">4B</figureCitation>
,
<figureCitation id="13452A3C151EFFECFD22FC06FD4EFBDA" box="[662,697,1004,1031]" captionStart="FIGURE 5" captionStartId="8.[152,255,1904,1928]" captionTargetBox="[152,1435,561,1881]" captionTargetId="figure-189@8.[152,1435,561,1881]" captionTargetPageId="8" captionText="FIGURE 5. Sericobracon puravida sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216365" httpUri="https://zenodo.org/record/15216365/files/figure.png" pageId="9" pageNumber="180">5C</figureCitation>
); and 4) mesopleuron mostly coriaceous (
<figureCitation id="13452A3C151EFFECFB16FC06FB04FBDB" box="[1186,1267,1004,1030]" captionStart="FIGURE 6" captionStartId="10.[152,255,1444,1468]" captionTargetBox="[152,1435,181,1420]" captionTargetId="figure-27@10.[152,1435,181,1420]" captionTargetPageId="10" captionText="FIGURE 6. Sericobracon zunigai sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216367" httpUri="https://zenodo.org/record/15216367/files/figure.png" pageId="9" pageNumber="180">Fig.6D</figureCitation>
) (similar in
<taxonomicName id="4C7E4D3A151EFFECFA33FC06FEE5FBF7" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="arimaensis">
<emphasis id="B90AEAAB151EFFECFA33FC06FEE5FBF7" italics="true" pageId="9" pageNumber="180">S. arimaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E4D3A151EFFECFEFDFBFAFE5AFBF7" box="[329,429,1040,1066]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="evansi">
<emphasis id="B90AEAAB151EFFECFEFDFBFAFE5AFBF7" box="[329,429,1040,1066]" italics="true" pageId="9" pageNumber="180">S. evansi</emphasis>
</taxonomicName>
(
<figureCitation id="13452A3C151EFFECFE0FFBFAFDEFFBF7" box="[443,536,1040,1066]" captionStart="FIGURE 2" captionStartId="4.[152,255,1921,1945]" captionTargetBox="[152,1436,956,1896]" captionTargetId="figure-234@4.[152,1436,956,1896]" captionTargetPageId="4" captionText="FIGURE 2. Sericobracon arimaensis Shaw. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216359" httpUri="https://zenodo.org/record/15216359/files/figure.png" pageId="9" pageNumber="180">Figs 2D</figureCitation>
,
<figureCitation id="13452A3C151EFFECFD90FBFAFD9EFBF6" box="[548,617,1040,1067]" captionStart="FIGURE 3" captionStartId="5.[152,255,1060,1084]" captionTargetBox="[151,1435,181,1036]" captionTargetId="figure-15@5.[151,1435,181,1036]" captionTargetPageId="5" captionText="FIGURE 3. Sericobracon evansi Shaw. Holotype, female. A) Mesosoma and metasoma, lateral view; B) mesosoma, lateral view; C) mesosoma, dorsal view; D) metasoma, dorsal view. Museum of Comparative Zoology, Harvard University; © President and Fellows of Harvard College." figureDoi="http://doi.org/10.5281/zenodo.15216361" httpUri="https://zenodo.org/record/15216361/files/figure.png" pageId="9" pageNumber="180">3A, B</figureCitation>
; mostly smooth in
<taxonomicName id="4C7E4D3A151EFFECFC8BFBFAFC2CFBF7" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[831,987,1040,1066]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="paulmarshi">
<emphasis id="B90AEAAB151EFFECFC8BFBFAFC2CFBF7" box="[831,987,1040,1066]" italics="true" pageId="9" pageNumber="180">S. paulmarshi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E4D3A151EFFECFBA5FBFAFB65FBF7" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[1041,1170,1040,1066]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="180" phylum="Arthropoda" rank="species" species="puravida">
<emphasis id="B90AEAAB151EFFECFBA5FBFAFB65FBF7" box="[1041,1170,1040,1066]" italics="true" pageId="9" pageNumber="180">S. puravida</emphasis>
</taxonomicName>
;
<figureCitation id="13452A3C151EFFECFB22FBFAFB01FBF7" box="[1174,1270,1040,1066]" captionStart="FIGURE 4" captionStartId="6.[152,255,1679,1703]" captionTargetBox="[177,1411,525,1655]" captionTargetId="figure-166@6.[177,1411,525,1655]" captionTargetPageId="6" captionText="FIGURE 4. Sericobracon paulmarshi sp. nov. Holotype, female.A) Habitus, lateral view; B) head and mesosoma, dorsal view; C) head, frontal view; D) head and mesosoma, lateral view; D) metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.15216363" httpUri="https://zenodo.org/record/15216363/files/figure.png" pageId="9" pageNumber="180">Figs 4D</figureCitation>
,
<figureCitation id="13452A3C151EFFECFAB5FBFAFAD0FBF6" box="[1281,1319,1040,1067]" captionStart="FIGURE 5" captionStartId="8.[152,255,1904,1928]" captionTargetBox="[152,1435,561,1881]" captionTargetId="figure-189@8.[152,1435,561,1881]" captionTargetPageId="8" captionText="FIGURE 5. Sericobracon puravida sp. nov. Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; metasoma, E) dorsal view; E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.15216365" httpUri="https://zenodo.org/record/15216365/files/figure.png" pageId="9" pageNumber="180">5D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3646532151EFFEFFF73FBDEFE7FF8DD" lastPageId="10" lastPageNumber="181" pageId="9" pageNumber="180" type="description">
<paragraph id="8BC136B9151EFFECFF73FBDEFD53FAFA" blockId="9.[151,1437,860,2039]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FBDEFEA6FB93" bold="true" box="[199,337,1076,1102]" pageId="9" pageNumber="180">Description</emphasis>
.
<typeStatus id="54C5881B151EFFECFED7FBDEFE3CFB93" box="[355,459,1076,1102]" pageId="9" pageNumber="180" type="holotype">Holotype</typeStatus>
female. Body size
<quantity id="4C869B5C151EFFECFD06FBDEFCF9FB92" box="[690,782,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="180" unit="mm" value="4.4">4.4 mm</quantity>
, fore wing
<quantity id="4C869B5C151EFFECFC23FBDEFC03FB92" box="[919,1012,1076,1103]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="180" unit="mm" value="3.2">3.2 mm</quantity>
.
<emphasis id="B90AEAAB151EFFECFBB6FBDEFBA6FB93" box="[1026,1105,1076,1102]" italics="true" pageId="9" pageNumber="180">Colour</emphasis>
: vertex, scapus, pedicel and flaggellomeres, mesoscutum, scutellum, propodeum, first, basal and apical median edges of second, and median areas of third to sixth metasimal tergites, and tarsi dark brown to black; frons, area surrounding eyes and temple, propleuron and pronotum, hind and middle coxae, femora and tibiae, most of second, lateral areas of third to sixth, and last three metasomal tergites, and coxae, femora, and tibiae whitish yellow; median and posterior part of mesopleuron whitish to honey yellow; wings hyaline, pterostigma and veins brown; ovipositor and sheaths honey yellow; ovipositor strongly sclerotised at apex.
</paragraph>
<paragraph id="8BC136B9151EFFECFF73FADAFB45F9FF" blockId="9.[151,1437,860,2039]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73FADAFEF3FA97" box="[199,260,1328,1354]" italics="true" pageId="9" pageNumber="180">Head</emphasis>
: transverse in dorsal view, 1.7 × wider than its median length. Eye 1.3 × longer than wide; eye width 2.4 × longer than temple in dorsal view. Malar space 0.3 × eye height and 0.8 × as long as width of hypoclypeal depression; mandibles bidentate. Occipital carina complete and reaching hypostomal carina before the base of mandible; inner margins of posterior ocelli (POL) 1.6 longer than ocellar diameter (OD), 0.6 × shorter than distance between outer margin of posterior ocellus and eye (OOL). Vertex striate, frons slightly excavated and glabrous, face transversally striate, with small, sparse setae. One antenna (left) with 32 flagellomeres, second antennae broken (right); first flagellomere about 2.6 × longer than wide, 1.2 × longer than second flagellomere.
</paragraph>
<paragraph id="8BC136B9151EFFECFF73F9C7FC2DF8BB" blockId="9.[151,1437,860,2039]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73F9C7FEC8F99B" box="[199,319,1581,1606]" italics="true" pageId="9" pageNumber="180">Mesosoma</emphasis>
: 1.4 × longer than high and 1.8 × longer than wide. Pronotal collar distinct, visible in dorsal view, acinose to slightly coriaceous, sparsely setose; pronotal furrow wide, deep and scrobiculate. Median length of mesoscutum as long as its width (dorsal view); mesoscutal lobes coriaceous, with median mesoscutal lobe slightly projected upwards from lateral lobes; notauli wide, deep, scrobiculate, not joining, obscuring before the endo of mesoscutum in a longitudinally striate-mostly smooth area. Scutellar disc smooth to slightly acinose, prescutellar furrow with six transverse carinae. Mesopleuron mostly coriaceous; precoxal sulcus wide, deep, scrobiculate and slightly coriaceous, about 0.6 × as long as mesopleuron; subalar sulcus wide, deep and scrobiculate. Metapleuron mostly coriaceous, slightly rugose anteriorly. Propodeum mostly coriaceous, rugose apically, with a median longitudinal carina running along basal third, areola distinct and complete.
</paragraph>
<paragraph id="8BC136B9151EFFECFF73F89BFF23F82A" blockId="9.[151,1437,860,2039]" pageId="9" pageNumber="180">
<emphasis id="B90AEAAB151EFFECFF73F89BFEFCF857" box="[199,267,1905,1930]" italics="true" pageId="9" pageNumber="180">Wings</emphasis>
: fore wing 4.2 × longer than wide. Pterostigma 2.0 × longer than wide and 0.6 × as long as R. Vein r as long as 3RSa, 0.2 × as long as 3RSb, and 1.1 × longer than vein r-m. Vein m-cu antefurcal to 2RS, vein RS+Mb present. First subdiscal cell open. Hind wing vein M+CU 0.6 × as long as 1M, m-cu slightly curved towards wing apex.
</paragraph>
<caption id="DF016631151DFFEFFF2CFA4EFBBEFA3D" ID-DOI="http://doi.org/10.5281/zenodo.15216367" ID-Zenodo-Dep="15216367" httpUri="https://zenodo.org/record/15216367/files/figure.png" pageId="10" pageNumber="181" startId="10.[152,255,1444,1468]" targetBox="[152,1435,181,1420]" targetPageId="10" targetType="figure">
<paragraph id="8BC136B9151DFFEFFF2CFA4EFBBEFA3D" blockId="10.[151,1437,1444,1504]" pageId="10" pageNumber="181">
<emphasis id="B90AEAAB151DFFEFFF2CFA4EFEECFA61" bold="true" box="[152,283,1444,1468]" pageId="10" pageNumber="181">FIGURE 6.</emphasis>
<taxonomicName id="4C7E4D3A151DFFEFFE90FA4EFDF7FA61" authorityName="Zaldívar-Riverón &amp; Castañeda-Osorio &amp; Shaw" authorityYear="2025" box="[292,512,1444,1468]" class="Insecta" family="Braconidae" genus="Sericobracon" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="181" phylum="Arthropoda" rank="species" species="zunigai" status="sp. nov.">
<emphasis id="B90AEAAB151DFFEFFE90FA4EFDF7FA61" box="[292,512,1444,1468]" italics="true" pageId="10" pageNumber="181">Sericobracon zunigai</emphasis>
</taxonomicName>
<emphasis id="B90AEAAB151DFFEFFDBCFA4EFDACFA61" bold="true" box="[520,603,1444,1468]" pageId="10" pageNumber="181">
<taxonomicNameLabel id="A23957D0151DFFEFFDBCFA4EFDACFA61" box="[520,603,1444,1468]" pageId="10" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, female. A) Habitus, lateral view; B) head, frontal view; C) head and mesosoma, dorsal view; D) mesosoma, lateral view; E) metasoma, dorsal view; F) fore wing.
</paragraph>
</caption>
<paragraph id="8BC136B9151DFFEFFF73F9D9FDE1F9AD" blockId="10.[151,1436,1586,1937]" pageId="10" pageNumber="181">
<emphasis id="B90AEAAB151DFFEFFF73F9D9FF0BF991" box="[199,252,1587,1612]" italics="true" pageId="10" pageNumber="181">Legs</emphasis>
: fore tibia without row of spines along anterior margin. Legs mostly smooth to slightly coriaceous. Hind coxa slightly coriaceous ventrally.
</paragraph>
<paragraph id="8BC136B9151DFFEFFF73F991FC18F900" blockId="10.[151,1436,1586,1937]" pageId="10" pageNumber="181">
<emphasis id="B90AEAAB151DFFEFFF73F991FECBF949" box="[199,316,1659,1684]" italics="true" pageId="10" pageNumber="181">Metasoma</emphasis>
: first tergite short, 0.8 × as long as apical width, longitudinally costate. Second tergite longitudinally costate; suture between second and third tergites straight, distinct; remaining tergites mostly smooth and polished, slightly acinose in some areas. Ovipositor length 0.9 × as long as metasoma.
</paragraph>
<paragraph id="8BC136B9151DFFEFFF73F90CFE7FF8DD" blockId="10.[151,1436,1586,1937]" box="[199,392,1766,1792]" pageId="10" pageNumber="181">
<emphasis id="B90AEAAB151DFFEFFF73F90CFEFCF8DD" bold="true" box="[199,267,1766,1792]" pageId="10" pageNumber="181">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3646532151DFFEFFF73F8E0FA83F8F9" box="[199,1396,1802,1829]" pageId="10" pageNumber="181" type="etymology">
<paragraph id="8BC136B9151DFFEFFF73F8E0FA83F8F9" blockId="10.[151,1436,1586,1937]" box="[199,1396,1802,1829]" pageId="10" pageNumber="181">
<emphasis id="B90AEAAB151DFFEFFF73F8E0FEBFF8F9" bold="true" box="[199,328,1802,1828]" pageId="10" pageNumber="181">Etymology</emphasis>
. The name of this species is an honorary patronym for the collector of the
<typeStatus id="54C5881B151DFFEFFB39F8E0FB05F8F9" box="[1165,1266,1802,1828]" pageId="10" pageNumber="181" type="holotype">holotype</typeStatus>
, C. Zuniga.
</paragraph>
</subSubSection>
<subSubSection id="C3646532151DFFEFFF73F8C4FE53F895" box="[199,420,1838,1864]" pageId="10" pageNumber="181" type="biology_ecology">
<paragraph id="8BC136B9151DFFEFFF73F8C4FE53F895" blockId="10.[151,1436,1586,1937]" box="[199,420,1838,1864]" pageId="10" pageNumber="181">
<emphasis id="B90AEAAB151DFFEFFF73F8C4FED0F895" bold="true" box="[199,295,1838,1864]" pageId="10" pageNumber="181">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3646532151DFFEFFF73F8B8FD17F84D" pageId="10" pageNumber="181" type="materials_examined">
<paragraph id="8BC136B9151DFFEFFF73F8B8FD17F84D" blockId="10.[151,1436,1586,1937]" pageId="10" pageNumber="181">
<emphasis id="B90AEAAB151DFFEFFF73F8B8FE5EF8B1" bold="true" box="[199,425,1874,1900]" pageId="10" pageNumber="181">Material examined</emphasis>
.
<materialsCitation id="3B163CE4151DFFEFFE03F8B8FD17F84D" collectionCode="CNIN, IBUNAM" collectorName="C. Zuniga &amp; Malaise. L. N." country="Costa Rica" elevation="600" location="Guanacaste" pageId="10" pageNumber="181" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<typeStatus id="54C5881B151DFFEFFE03F8B8FDE8F8B1" box="[439,543,1874,1900]" pageId="10" pageNumber="181" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED6FAE7C151DFFEFFD99F8B8FD84F8B1" box="[557,627,1874,1900]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="10" pageNumber="181" type="">CNIN</collectionCode>
<collectionCode id="ED6FAE7C151DFFEFFDCEF8B8FD06F8B1" box="[634,753,1874,1900]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="10" pageNumber="181" type="University or college">IBUNAM</collectionCode>
). Female.
<collectingCountry id="F3697629151DFFEFFCD7F8B8FC15F8B1" box="[867,994,1874,1900]" name="Costa Rica" pageId="10" pageNumber="181">Costa Rica</collectingCountry>
:
<collectingRegion id="49BAF85B151DFFEFFC58F8B8FB87F8B1" box="[1004,1136,1874,1900]" country="Costa Rica" name="Guanacaste" pageId="10" pageNumber="181">Guanacaste</collectingRegion>
, Est. Biol. Maritza,
<quantity id="4C869B5C151DFFEFFAFBF8B8FA60F8B0" box="[1359,1431,1874,1901]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="181" unit="m" value="600.0">
<elevation id="0053D18A151DFFEFFAFBF8B8FA60F8B0" box="[1359,1431,1874,1901]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="181" unit="m" value="600.0">600 m</elevation>
</quantity>
, xi, 1996,
<collectorName id="268B536F151DFFEFFEB5F89CFE83F84D" box="[257,372,1910,1936]" pageId="10" pageNumber="181">C. Zuniga</collectorName>
,
<collectorName id="268B536F151DFFEFFECBF89CFDD7F84C" box="[383,544,1910,1937]" pageId="10" pageNumber="181">Malaise. L. N.</collectorName>
326900-373000.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

178
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@ -0,0 +1,178 @@
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<mods:title id="BBEDB073E1608B9A5A0B33938104E447">Three new species of the genus Rachicerus Walker (Diptera, Xylophagidae) from Brazil</mods:title>
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<mods:namePart id="2EB5C97E0477D1E8921830F275C4C86B">Pujol-Luz, José Roberto</mods:namePart>
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<subSubSection id="C3698790FFB4FFFFFF7177BDFD71533A" box="[151,663,1525,1552]" pageId="8" pageNumber="302" type="nomenclature">
<paragraph id="8BCCD41BFFB4FFFFFF7177BDFD71533A" blockId="8.[151,663,1525,1588]" box="[151,663,1525,1552]" pageId="8" pageNumber="302">
<heading id="D0846377FFB4FFFFFF7177BDFD71533A" bold="true" box="[151,663,1525,1552]" fontSize="11" level="1" pageId="8" pageNumber="302" reason="1">
<emphasis id="B9070809FFB4FFFFFF7177BDFD71533A" bold="true" box="[151,663,1525,1552]" pageId="8" pageNumber="302">
<taxonomicName id="4C73AF98FFB4FFFFFF7177BDFE1C533A" authority="Pujol-Luz" authorityName="Pujol-Luz &amp; Godoi" authorityYear="2025" box="[151,506,1525,1552]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="302" phylum="Arthropoda" rank="species" species="victori" status="sp. nov.">
<emphasis id="B9070809FFB4FFFFFF7177BDFE92533A" bold="true" box="[151,372,1525,1552]" italics="true" pageId="8" pageNumber="302">Rachicerus victori</emphasis>
Pujol-Luz
</taxonomicName>
,
<taxonomicNameLabel id="A234B572FFB4FFFFFDE177BEFD71533A" box="[519,663,1525,1552]" pageId="8" pageNumber="302" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB4FFFFFF717452FEC15306" box="[151,295,1561,1588]" pageId="8" pageNumber="302" type="description">
<paragraph id="8BCCD41BFFB4FFFFFF717452FEC15306" blockId="8.[151,663,1525,1588]" box="[151,295,1561,1588]" pageId="8" pageNumber="302">
(
<figureCitation id="1348C89EFFB4FFFFFF797452FEF95306" box="[159,287,1561,1588]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[350,1238,630,1897]" captionTargetId="figure-182@9.[350,1238,630,1899]" captionTargetPageId="9" captionText="FIGURES 16–18. Rachicerus victori Pujol-Luz sp. nov. 16, Habitus in lateral view; 17, Frontal view of the head, showing the orbital triangle; 18, Left antenna. (Holotype housed in MZUSP)." pageId="8" pageNumber="302">Figs 16–18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB4FFFFFF717429FC775395" pageId="8" pageNumber="302" type="diagnosis">
<paragraph id="8BCCD41BFFB4FFFFFF717429FC775395" blockId="8.[151,1437,1633,2020]" pageId="8" pageNumber="302">
<emphasis id="B9070809FFB4FFFFFF717429FEEC534E" bold="true" box="[151,266,1633,1659]" pageId="8" pageNumber="302">Diagnosis</emphasis>
. Brownish body with blackish terminalia; Femora, tibiae and first tarsomeres with light and dark brown stripes; antenna with slender, spoon-shaped pectinate flagellomeres.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB4FFFEFF2174E2FE445450" lastPageId="9" lastPageNumber="303" pageId="8" pageNumber="302" type="description">
<paragraph id="8BCCD41BFFB4FFFEFF2174E2FE375474" blockId="8.[151,1437,1633,2020]" lastBlockId="9.[151,1436,150,573]" lastPageId="9" lastPageNumber="303" pageId="8" pageNumber="302">
<emphasis id="B9070809FFB4FFFFFF2174E2FE6653F1" bold="true" box="[199,384,1706,1732]" pageId="8" pageNumber="302">Measurements:</emphasis>
body length:
<quantity id="4C8B79FEFFB4FFFFFDC274E1FD6653F1" box="[548,640,1705,1732]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="8" pageNumber="302" unit="mm" value="6.6">6.6 mm</quantity>
; antenna:
<quantity id="4C8B79FEFFB4FFFFFD1174E2FCB253F1" box="[759,852,1706,1732]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="302" unit="mm" value="1.3">1.3 mm</quantity>
; number of antennal flagellomeres: 17; wings:
<quantity id="4C8B79FEFFB4FFFFFA9F74E1FF2053DD" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="302" unit="mm" value="6.5">6.5 mm</quantity>
.
<emphasis id="B9070809FFB4FFFFFF377486FEF753DD" bold="true" box="[209,273,1742,1768]" pageId="8" pageNumber="302">Head</emphasis>
: hemispherical, wider than thorax; front flat, shining brown, with two longitudinal ridges running from ocellar plate and converging on a suture that divides antennal plate in half; striated integument between ridges; frontal plate large with small whitish setae (pruinosity like) in inferior margin around antennae; fronto-orbital plate depressed and bare (
<figureCitation id="1348C89EFFB4FFFFFE9D7572FE295261" box="[379,463,1850,1876]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[350,1238,630,1897]" captionTargetId="figure-182@9.[350,1238,630,1899]" captionTargetPageId="9" captionText="FIGURES 16–18. Rachicerus victori Pujol-Luz sp. nov. 16, Habitus in lateral view; 17, Frontal view of the head, showing the orbital triangle; 18, Left antenna. (Holotype housed in MZUSP)." pageId="8" pageNumber="302">Fig. 17</figureCitation>
). Compound eyes dichoptic, reniform and bare; vertex with ocellar triangle developed, black; ocellus brownish (
<figureCitation id="1348C89EFFB4FFFFFE597516FDF1524D" box="[447,535,1886,1912]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[350,1238,630,1897]" captionTargetId="figure-182@9.[350,1238,630,1899]" captionTargetPageId="9" captionText="FIGURES 16–18. Rachicerus victori Pujol-Luz sp. nov. 16, Habitus in lateral view; 17, Frontal view of the head, showing the orbital triangle; 18, Left antenna. (Holotype housed in MZUSP)." pageId="8" pageNumber="302">Fig. 17</figureCitation>
). Occiput well developed and very pilose. Palps brownish and 2-segmented. Labellum brownish, membranous with elongate setae. Antennae elongate more than two times greater than width of head, with 17 pectinate spoon-shaped flagellomeres (
<figureCitation id="1348C89EFFB4FFFFFCC375EEFC5452F5" box="[805,946,1957,1984]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[350,1238,630,1897]" captionTargetId="figure-182@9.[350,1238,630,1899]" captionTargetPageId="9" captionText="FIGURES 16–18. Rachicerus victori Pujol-Luz sp. nov. 16, Habitus in lateral view; 17, Frontal view of the head, showing the orbital triangle; 18, Left antenna. (Holotype housed in MZUSP)." pageId="8" pageNumber="302">
Figs. 16
<emphasis id="B9070809FFB4FFFFFC6375EEFC7552F5" bold="true" box="[901,915,1958,1984]" pageId="8" pageNumber="302">–</emphasis>
18
</figureCitation>
).
<emphasis id="B9070809FFB4FFFFFC2C75EEFBC552F5" bold="true" box="[970,1059,1958,1984]" pageId="8" pageNumber="302">Thorax</emphasis>
: predominantly dark brown and pilose; postpronotal lobe brownish; disc of thorax with dense pilosity; anepimerom and laterotergites entirely bare; scutellum brownish, with sparse pilosity (
<figureCitation id="1348C89EFFB5FFFEFD9372DFFD2C5584" box="[629,714,151,177]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[350,1238,630,1897]" captionTargetId="figure-182@9.[350,1238,630,1899]" captionTargetPageId="9" captionText="FIGURES 16–18. Rachicerus victori Pujol-Luz sp. nov. 16, Habitus in lateral view; 17, Frontal view of the head, showing the orbital triangle; 18, Left antenna. (Holotype housed in MZUSP)." pageId="9" pageNumber="303">Fig. 16</figureCitation>
).
<emphasis id="B9070809FFB5FFFEFD0672DEFCCA5585" bold="true" box="[736,812,150,176]" pageId="9" pageNumber="303">Wings</emphasis>
: hyaline, inconspicuous pterostigma between veins Sc and R
<subScript id="17F7D65EFFB5FFFEFF3D7285FF0555EE" attach="left" box="[219,227,205,219]" fontSize="6" pageId="9" pageNumber="303">1</subScript>
; R
<subScript id="17F7D65EFFB5FFFEFEE17285FEF655EE" attach="left" box="[263,272,205,219]" fontSize="6" pageId="9" pageNumber="303">S</subScript>
stem curved upwards before bifurcation of radial sectors; halters brownish.
<emphasis id="B9070809FFB5FFFEFB9872F3FB5055E0" bold="true" box="[1150,1206,187,213]" pageId="9" pageNumber="303">Legs</emphasis>
: coxae, trochanters, femurs and tibiae brownish, posterior
<superScript id="7C067953FFB5FFFEFDAF7296FDB755D9" attach="right" box="[585,593,222,236]" fontSize="6" pageId="9" pageNumber="303">1</superScript>
⁄
<subScript id="17F7D65EFFB5FFFEFDB072B9FDB855CA" attach="left" box="[598,606,241,255]" fontSize="6" pageId="9" pageNumber="303">2</subScript>
of femurs and tibiae brownish yellow; first tarsomeres whitish; empodia pulvilliform (
<figureCitation id="1348C89EFFB5FFFEFED6734BFE635428" box="[304,389,259,285]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[350,1238,630,1897]" captionTargetId="figure-182@9.[350,1238,630,1899]" captionTargetPageId="9" captionText="FIGURES 16–18. Rachicerus victori Pujol-Luz sp. nov. 16, Habitus in lateral view; 17, Frontal view of the head, showing the orbital triangle; 18, Left antenna. (Holotype housed in MZUSP)." pageId="9" pageNumber="303">Fig. 16</figureCitation>
).
<emphasis id="B9070809FFB5FFFEFE7D734AFDF65429" bold="true" box="[411,528,258,284]" pageId="9" pageNumber="303">Abdomen</emphasis>
: entirely brownish with dark brownish to black terminalia, with dense pilosity; terminalia conical (
<figureCitation id="1348C89EFFB5FFFEFE96736FFE255474" box="[368,451,295,321]" captionStart="FIGURES 16–18" captionStartId="9.[151,264,1926,1950]" captionTargetBox="[350,1238,630,1897]" captionTargetId="figure-182@9.[350,1238,630,1899]" captionTargetPageId="9" captionText="FIGURES 16–18. Rachicerus victori Pujol-Luz sp. nov. 16, Habitus in lateral view; 17, Frontal view of the head, showing the orbital triangle; 18, Left antenna. (Holotype housed in MZUSP)." pageId="9" pageNumber="303">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BCCD41BFFB5FFFEFF217303FE445450" blockId="9.[151,1436,150,573]" box="[199,418,331,357]" pageId="9" pageNumber="303">
<emphasis id="B9070809FFB5FFFEFF217303FEC35450" bold="true" box="[199,293,331,357]" pageId="9" pageNumber="303">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB5FFFEFF217327FCB25498" pageId="9" pageNumber="303" type="etymology">
<paragraph id="8BCCD41BFFB5FFFEFF217327FC7D54BC" blockId="9.[151,1436,150,573]" box="[199,923,367,393]" pageId="9" pageNumber="303">
<emphasis id="B9070809FFB5FFFEFF217327FEAE54BC" bold="true" box="[199,328,367,393]" pageId="9" pageNumber="303">Etymology</emphasis>
: This name is given to honor my beloved son Victor.
</paragraph>
<paragraph id="8BCCD41BFFB5FFFEFF2173DAFCB25498" blockId="9.[151,1436,150,573]" box="[199,852,402,429]" pageId="9" pageNumber="303">
<emphasis id="B9070809FFB5FFFEFF2173DAFE185499" bold="true" box="[199,510,402,428]" pageId="9" pageNumber="303">Geographical distribution</emphasis>
.
<collectingCountry id="F364948BFFB5FFFEFDEA73DBFD925498" box="[524,628,403,429]" name="Brazil" pageId="9" pageNumber="303">BRAZIL</collectingCountry>
, State of
<collectingRegion id="49B71AF9FFB5FFFEFD3A73DBFCB65498" box="[732,848,403,429]" country="Brazil" name="Sao Paulo" pageId="9" pageNumber="303">São Paulo</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB5FFFEFF2173FFFDE15708" pageId="9" pageNumber="303" type="materials_examined">
<paragraph id="8BCCD41BFFB5FFFEFF2173FFFDE15708" blockId="9.[151,1436,150,573]" pageId="9" pageNumber="303">
<emphasis id="B9070809FFB5FFFEFF2173FFFE9754E5" bold="true" box="[199,369,438,465]" pageId="9" pageNumber="303">Type material</emphasis>
.
<materialsCitation id="3B1BDE46FFB5FFFEFE6773FFFDE55708" collectionCode="MZUSP" collectorName="Paulo, S. &amp; Roque, J &amp; J. Lane" country="Brazil" location="The" municipality="Museu de Zoologia da Universidade" pageId="9" pageNumber="303" specimenCode="MZ053525" specimenCount="3" stateProvince="Sao Paulo" typeStatus="holotype">
<location id="8EAC82C0FFB5FFFEFE6773FFFE4B54E4" LSID="urn:lsid:plazi:treatment:03DA650DFFB4FFFEFF7177BDFDE15708:8EAC82C0FFB5FFFEFE6773FFFE4B54E4" box="[385,429,439,465]" country="Brazil" municipality="Museu de Zoologia da Universidade" name="The" pageId="9" pageNumber="303" stateProvince="Sao Paulo">The</location>
male
<typeStatus id="54C86AB9FFB5FFFEFE1173FFFD6D54E4" box="[503,651,439,465]" pageId="9" pageNumber="303" type="holotype">HOLOTYPE</typeStatus>
is housed in the
<collectingMunicipality id="6BA84E61FFB5FFFEFCB173FFFB1C54E4" box="[855,1274,439,465]" pageId="9" pageNumber="303">Museu de Zoologia da Universidade</collectingMunicipality>
de
<collectingRegion id="49B71AF9FFB5FFFEFACE73FFFA7A54E4" box="[1320,1436,439,465]" country="Brazil" name="Sao Paulo" pageId="9" pageNumber="303">São Paulo</collectingRegion>
(
<collectionCode id="ED624CDEFFB5FFFEFF797393FEE454C0" box="[159,258,475,501]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="303" type="">MZUSP</collectionCode>
),
<collectingRegion id="49B71AF9FFB5FFFEFEF37393FE6E54C0" box="[277,392,475,501]" country="Brazil" name="Sao Paulo" pageId="9" pageNumber="303">São Paulo</collectingRegion>
,
<collectingRegion id="49B71AF9FFB5FFFEFE757393FDE054C0" box="[403,518,475,501]" country="Brazil" name="Sao Paulo" pageId="9" pageNumber="303">São Paulo</collectingRegion>
,
<collectingCountry id="F364948BFFB5FFFEFDF67393FDB154C0" box="[528,599,475,501]" name="Brazil" pageId="9" pageNumber="303">Brazil</collectingCountry>
, labeled: S.[ão]
<collectorName id="2686B1CDFFB5FFFEFCEA7393FC8B54C0" box="[780,877,475,501]" pageId="9" pageNumber="303">Paulo, S.</collectorName>
[ão]
<collectorName id="2686B1CDFFB5FFFEFC457393FBEE54C0" box="[931,1032,475,501]" pageId="9" pageNumber="303">Roque, J</collectorName>
. Lane col., XII.[19]46 [white label]/
<typeStatus id="54C86AB9FFB5FFFEFF7173B7FF1D572C" box="[151,251,511,537]" pageId="9" pageNumber="303" type="holotype">Holótipo</typeStatus>
[red label]/
<specimenCode id="DBD57C60FFB5FFFEFE6F73B6FDEE572C" box="[393,520,510,537]" collectionCode="MZ" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="9" pageNumber="303">MZ053525</specimenCode>
[white label]. The
<typeStatus id="54C86AB9FFB5FFFEFD0273B7FCA0572C" box="[740,838,511,537]" pageId="9" pageNumber="303" type="holotype">holotype</typeStatus>
is in good condition, only missing the right antenna and the lack of the right fore leg
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

255
data/03/DA/65/03DA650DFFB6FFFDFF717609FA7B53BB.xml Normal file
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@ -0,0 +1,255 @@
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<mods:title id="6BEF7AED09E9045A2DDCA940CF890BB6">Three new species of the genus Rachicerus Walker (Diptera, Xylophagidae) from Brazil</mods:title>
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<mods:affiliation id="F9E00A6CF876CE324595859C9370D0C4">Universidade de Brasília (UNB), Instituto de Ciências Biológicas, Departamento de Zoologia, Brasília, DF, Brazil.</mods:affiliation>
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<mods:affiliation id="797FF8CAD56640340C719FBDB9FCD2FC">Universidade de Brasília (UNB), Instituto de Ciências Biológicas, Departamento de Zoologia, Brasília, DF, Brazil. &amp; Universidade Federal do Amazonas (UFAM), Instituto de Ciências Biológicas, Departamento de Biologia, Manaus, AM, Brazil.</mods:affiliation>
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<treatment id="03DA650DFFB6FFFDFF717609FA7B53BB" LSID="urn:lsid:plazi:treatment:03DA650DFFB6FFFDFF717609FA7B53BB" httpUri="http://treatment.plazi.org/id/03DA650DFFB6FFFDFF717609FA7B53BB" lastPageNumber="304" pageId="10" pageNumber="304">
<subSubSection id="C3698790FFB6FFFDFF717609FCB2516E" box="[151,852,1088,1115]" pageId="10" pageNumber="304" type="nomenclature">
<paragraph id="8BCCD41BFFB6FFFDFF717609FCB2516E" blockId="10.[151,852,1088,1115]" box="[151,852,1088,1115]" pageId="10" pageNumber="304">
<heading id="D0846377FFB6FFFDFF717609FCB2516E" bold="true" box="[151,852,1088,1115]" fontSize="11" level="1" pageId="10" pageNumber="304" reason="1">
<emphasis id="B9070809FFB6FFFDFF717609FCB2516E" bold="true" box="[151,852,1088,1115]" pageId="10" pageNumber="304">
Key to Brazilian species of the genus
<taxonomicName id="4C73AF98FFB6FFFDFD817608FCB2516E" ID-CoL="63QHH" authority="Walker" authorityName="Walker" box="[615,852,1088,1115]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="B9070809FFB6FFFDFD817608FD16516E" bold="true" box="[615,752,1088,1115]" italics="true" pageId="10" pageNumber="304">Rachicerus</emphasis>
Walker
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB6FFFDFF7176CEFA7B53BB" pageId="10" pageNumber="304" type="key">
<keyStep id="3087CCBEFFB6FFFDFF7176CEFA7A518D" pageId="10" pageNumber="304">
<paragraph id="8BCCD41BFFB6FFFDFF7176CEFA7A51AE" blockId="10.[151,1437,1158,1678]" box="[151,1436,1158,1180]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF7176CEFA7A51AE" box="[151,1436,1158,1180]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF7176CEFF4551AE" bold="true" box="[151,163,1158,1179]" pageId="10" pageNumber="304">1</emphasis>
Antenna with less than 19 flagellomeres...................................................................
<emphasis id="B9070809FFB6FFFDFA7676CEFA7A51AE" bold="true" box="[1424,1436,1158,1179]" pageId="10" pageNumber="304">2</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCD41BFFB6FFFDFF7176EBFA7A518D" blockId="10.[151,1437,1158,1678]" box="[151,1436,1187,1209]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF7176EBFA7A518D" box="[151,1436,1187,1209]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF7176EBFF79518D" bold="true" box="[151,159,1187,1208]" pageId="10" pageNumber="304">-</emphasis>
Antenna with 20 or more flagellomeres...................................................................
<emphasis id="B9070809FFB6FFFDFA7676EBFA7A518D" bold="true" box="[1424,1436,1187,1208]" pageId="10" pageNumber="304">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087CCBEFFB6FFFDFF717688FA7A5025" pageId="10" pageNumber="304">
<paragraph id="8BCCD41BFFB6FFFDFF717688FA7B51E3" blockId="10.[151,1437,1158,1678]" box="[151,1437,1216,1238]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF717688FA7B51E3" box="[151,1437,1216,1238]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF717688FF4551E0" bold="true" box="[151,163,1216,1237]" pageId="10" pageNumber="304">2</emphasis>
Antenna bipectinate with 19 flagellomeres; scutum orange; abdomen black, with bluish reflections...
<taxonomicName id="4C73AF98FFB6FFFDFB227689FA7B51E3" ID-CoL="8GT9Y" authority="Pujol-Luz" authorityName="Pujol-Luz" authorityYear="2019" box="[1220,1437,1216,1238]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="carrerai">
<emphasis id="B9070809FFB6FFFDFB227689FA7B51E3" bold="true" box="[1220,1437,1216,1238]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFB227689FAD651E0" bold="true" box="[1220,1328,1216,1238]" italics="true" pageId="10" pageNumber="304">R. carrerai</emphasis>
Pujol-Luz
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCD41BFFB6FFFDFF717696FA7A5025" blockId="10.[151,1437,1158,1678]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF717696FA7A5025" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF717696FF7951C6" bold="true" box="[151,159,1246,1267]" pageId="10" pageNumber="304">-</emphasis>
Antenna single pectinate with 16 or 17 flagellomeres; scutum entirely or partially blackish to brownish; abdomen blackish to yellowish without, with bluish reflections..................................................................
<emphasis id="B9070809FFB6FFFDFA7676B3FA7A5025" bold="true" box="[1424,1436,1275,1296]" pageId="10" pageNumber="304">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087CCBEFFB6FFFDFF717750FA7A507E" pageId="10" pageNumber="304">
<paragraph id="8BCCD41BFFB6FFFDFF717750FA7B501B" blockId="10.[151,1437,1158,1678]" box="[151,1437,1304,1326]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF717750FA7B501B" box="[151,1437,1304,1326]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF717750FF455018" bold="true" box="[151,163,1304,1325]" pageId="10" pageNumber="304">3</emphasis>
Antenna with 18 flagellomeres; 1
<superScript id="7C067953FFB6FFFDFDC57750FDCA5011" attach="none" box="[547,556,1304,1316]" fontSize="5" pageId="10" pageNumber="304">st</superScript>
, 2
<superScript id="7C067953FFB6FFFDFDA27750FDB45011" attach="left" box="[580,594,1304,1316]" fontSize="5" pageId="10" pageNumber="304">nd</superScript>
and 3
<superScript id="7C067953FFB6FFFDFD6B7750FD7F5011" attach="left" box="[653,665,1304,1316]" fontSize="5" pageId="10" pageNumber="304">rd</superScript>
abdominal segments yellowish.....................
<taxonomicName id="4C73AF98FFB6FFFDFB337751FA7B501B" ID-CoL="78RS9" authority="Carrera" authorityName="Carrera" authorityYear="1940" box="[1237,1437,1304,1326]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="oliverioi">
<emphasis id="B9070809FFB6FFFDFB337751FA7B501B" bold="true" box="[1237,1437,1304,1326]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFB337751FAA4501B" bold="true" box="[1237,1346,1304,1326]" italics="true" pageId="10" pageNumber="304">R. oliverioi</emphasis>
Carrera
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCD41BFFB6FFFDFF71777EFA7A507E" blockId="10.[151,1437,1158,1678]" box="[151,1436,1334,1356]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF71777EFA7A507E" box="[151,1436,1334,1356]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF71777EFF79507E" bold="true" box="[151,159,1334,1355]" pageId="10" pageNumber="304">-</emphasis>
Antenna with 16–17 flagellomeres; abdominal segments blackish to brownish....................................
<emphasis id="B9070809FFB6FFFDFA76777EFA7A507E" bold="true" box="[1424,1436,1334,1355]" pageId="10" pageNumber="304">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087CCBEFFB6FFFDFF71771BFA7A50B3" pageId="10" pageNumber="304">
<paragraph id="8BCCD41BFFB6FFFDFF71771BFA7A505D" blockId="10.[151,1437,1158,1678]" box="[151,1436,1363,1385]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF71771BFA7A505D" box="[151,1436,1363,1385]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF71771BFF45505D" bold="true" box="[151,163,1363,1384]" pageId="10" pageNumber="304">4</emphasis>
Antenna with 16 flagellomeres..................................................
<emphasis id="B9070809FFB6FFFDFB97771BFA7A505D" bold="true" box="[1137,1436,1363,1385]" pageId="10" pageNumber="304">
<taxonomicName id="4C73AF98FFB6FFFDFB97771BFAAC505C" authority="Pujol-Luz" authorityName="Pujol-Luz &amp; Godoi" authorityYear="2025" box="[1137,1354,1363,1385]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="B9070809FFB6FFFDFB97771BFB3B505C" bold="true" box="[1137,1245,1363,1385]" italics="true" pageId="10" pageNumber="304">R. isabelae</emphasis>
Pujol-Luz
</taxonomicName>
<taxonomicNameLabel id="A234B572FFB6FFFDFAB6771BFA7A505D" box="[1360,1436,1363,1384]" pageId="10" pageNumber="304" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCD41BFFB6FFFDFF717739FA7A50B3" blockId="10.[151,1437,1158,1678]" box="[151,1436,1392,1414]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF717739FA7A50B3" box="[151,1436,1392,1414]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF717739FF7950B3" bold="true" box="[151,159,1393,1414]" pageId="10" pageNumber="304">-</emphasis>
Antenna with 17 flagellomeres...................................................
<emphasis id="B9070809FFB6FFFDFB647739FA7A50B3" bold="true" box="[1154,1436,1392,1414]" pageId="10" pageNumber="304">
<taxonomicName id="4C73AF98FFB6FFFDFB647739FAAC50B3" authority="Pujol-Luz" authorityName="Pujol-Luz &amp; Godoi" authorityYear="2025" box="[1154,1354,1392,1414]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="victori" status="sp. nov.">
<emphasis id="B9070809FFB6FFFDFB647739FB3B50B0" bold="true" box="[1154,1245,1392,1414]" italics="true" pageId="10" pageNumber="304">R. victori</emphasis>
Pujol-Luz
</taxonomicName>
<taxonomicNameLabel id="A234B572FFB6FFFDFAB67739FA7A50B3" box="[1360,1436,1393,1414]" pageId="10" pageNumber="304" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087CCBEFFB6FFFDFF7177C6FA7A50E8" pageId="10" pageNumber="304">
<paragraph id="8BCCD41BFFB6FFFDFF7177C6FA7A50F5" blockId="10.[151,1437,1158,1678]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF7177C6FA7A50F5" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF7177C6FF455096" bold="true" box="[151,163,1422,1443]" pageId="10" pageNumber="304">5</emphasis>
Antenna with 20 flagellomeres; abdominal segments black with intersegmentar membrane whitish.........................................................................................
<emphasis id="B9070809FFB6FFFDFBC777E3FA7A50F5" bold="true" box="[1057,1436,1451,1473]" pageId="10" pageNumber="304">
<taxonomicName id="4C73AF98FFB6FFFDFBC777E3FAAC50F4" authority="Pujol-Luz &amp; Godoi" authorityName="Pujol-Luz &amp; Godoi" authorityYear="2025" box="[1057,1354,1451,1473]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="rafaeli" status="sp. nov.">
<emphasis id="B9070809FFB6FFFDFBC777E3FB9850F4" bold="true" box="[1057,1150,1451,1473]" italics="true" pageId="10" pageNumber="304">R. rafaeli</emphasis>
Pujol-Luz &amp; Godoi
</taxonomicName>
<taxonomicNameLabel id="A234B572FFB6FFFDFAB677E3FA7A50F5" box="[1360,1436,1451,1472]" pageId="10" pageNumber="304" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCD41BFFB6FFFDFF717781FA7A50E8" blockId="10.[151,1437,1158,1678]" box="[151,1436,1480,1502]" pageId="10" pageNumber="304">
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<emphasis id="B9070809FFB6FFFDFF717781FF7950EB" bold="true" box="[151,159,1481,1502]" pageId="10" pageNumber="304">-</emphasis>
Antenna with 21 to 28 flagellomeres; abdominal segments predominantly blackish or brownish.......................
<emphasis id="B9070809FFB6FFFDFA767780FA7A50E8" bold="true" box="[1424,1436,1480,1501]" pageId="10" pageNumber="304">6</emphasis>
</keyLead>
</paragraph>
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<keyStep id="3087CCBEFFB6FFFDFF7177AEFA7A532D" pageId="10" pageNumber="304">
<paragraph id="8BCCD41BFFB6FFFDFF7177AEFA7A50CE" blockId="10.[151,1437,1158,1678]" box="[151,1436,1510,1532]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF7177AEFA7A50CE" box="[151,1436,1510,1532]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF7177AEFF4550CE" bold="true" box="[151,163,1510,1531]" pageId="10" pageNumber="304">6</emphasis>
Antenna with 22 or 25 flagellomeres.....................................................................
<emphasis id="B9070809FFB6FFFDFA7677AEFA7A50CE" bold="true" box="[1424,1436,1510,1531]" pageId="10" pageNumber="304">7</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCD41BFFB6FFFDFF71744BFA7A532D" blockId="10.[151,1437,1158,1678]" box="[151,1436,1539,1561]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF71744BFA7A532D" box="[151,1436,1539,1561]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF71744BFF79532D" bold="true" box="[151,159,1539,1560]" pageId="10" pageNumber="304">-</emphasis>
Antenna with 28 flagellomeres..........................................................................
<emphasis id="B9070809FFB6FFFDFA76744BFA7A532D" bold="true" box="[1424,1436,1539,1560]" pageId="10" pageNumber="304">8</emphasis>
</keyLead>
</paragraph>
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<keyStep id="3087CCBEFFB6FFFDFF717469FA7B5361" pageId="10" pageNumber="304">
<paragraph id="8BCCD41BFFB6FFFDFF717469FA7B5303" blockId="10.[151,1437,1158,1678]" box="[151,1437,1568,1590]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF717469FA7B5303" box="[151,1437,1568,1590]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF717469FF455303" bold="true" box="[151,163,1569,1590]" pageId="10" pageNumber="304">7</emphasis>
Antenna with 22 flagellomeres, 2
<superScript id="7C067953FFB6FFFDFDC47468FDD65319" attach="none" box="[546,560,1568,1580]" fontSize="5" pageId="10" pageNumber="304">nd</superScript>
, sometimes 3
<superScript id="7C067953FFB6FFFDFD537468FD275319" attach="none" box="[693,705,1568,1580]" fontSize="5" pageId="10" pageNumber="304">rd</superScript>
; abdominal segments whitish......................
<taxonomicName id="4C73AF98FFB6FFFDFB137469FA7B5303" ID-CoL="4R89W" authority="Carrera" authorityName="Carrera" authorityYear="1940" box="[1269,1437,1568,1590]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B9070809FFB6FFFDFB137469FA7B5303" bold="true" box="[1269,1437,1568,1590]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFB137469FAA45303" bold="true" box="[1269,1346,1568,1590]" italics="true" pageId="10" pageNumber="304">R. lanei</emphasis>
Carrera
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCD41BFFB6FFFDFF717476FA7B5361" blockId="10.[151,1437,1158,1678]" box="[151,1437,1597,1620]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF717476FA7B5361" box="[151,1437,1597,1620]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF717476FF795366" bold="true" box="[151,159,1598,1619]" pageId="10" pageNumber="304">-</emphasis>
Antenna with 25 flagellomeres; abdominal segments bright black.................................
<taxonomicName id="4C73AF98FFB6FFFDFB0B7476FA7B5361" ID-CoL="4R89Y" authority="Carrera" authorityName="Carrera" authorityYear="1940" box="[1261,1437,1597,1620]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="lopesi">
<emphasis id="B9070809FFB6FFFDFB0B7476FA7B5361" bold="true" box="[1261,1437,1597,1620]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFB0B7476FAA45366" bold="true" box="[1261,1346,1597,1619]" italics="true" pageId="10" pageNumber="304">R. lopesi</emphasis>
Carrera
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087CCBEFFB6FFFDFF717413FA7B53BB" pageId="10" pageNumber="304">
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<keyLead id="3082772EFFB6FFFDFF717413FA7B5344" box="[151,1437,1627,1649]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF717413FF455345" bold="true" box="[151,163,1627,1648]" pageId="10" pageNumber="304">8</emphasis>
Longitudinal veins strongly darkened; abdomen entirely black.................................
<taxonomicName id="4C73AF98FFB6FFFDFB317413FA7B5344" ID-CoL="78S46" authority="Carrera" authorityName="Carrera" authorityYear="1940" box="[1239,1437,1627,1649]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="marcusi">
<emphasis id="B9070809FFB6FFFDFB317413FA7B5344" bold="true" box="[1239,1437,1627,1649]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFB317413FAA55345" bold="true" box="[1239,1347,1627,1648]" italics="true" pageId="10" pageNumber="304">R. marcusi</emphasis>
Carrera
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCD41BFFB6FFFDFF717431FA7B53BB" blockId="10.[151,1437,1158,1678]" box="[151,1437,1656,1678]" pageId="10" pageNumber="304">
<keyLead id="3082772EFFB6FFFDFF717431FA7B53BB" box="[151,1437,1656,1678]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFF717431FF7953BB" bold="true" box="[151,159,1657,1678]" pageId="10" pageNumber="304">-</emphasis>
Longitudinal veins darkened only in costal region of the wings; 2
<superScript id="7C067953FFB6FFFDFCB27430FC8453B1" attach="left" box="[852,866,1656,1668]" fontSize="5" pageId="10" pageNumber="304">nd</superScript>
abdominal segment bright yellow...
<taxonomicName id="4C73AF98FFB6FFFDFB2F7431FA7B53BB" ID-CoL="4R8B5" authority="Carrera" authorityName="Carrera" authorityYear="1945" box="[1225,1437,1656,1678]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="304" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="B9070809FFB6FFFDFB2F7431FA7B53BB" bold="true" box="[1225,1437,1656,1678]" pageId="10" pageNumber="304">
<emphasis id="B9070809FFB6FFFDFB2F7431FAA453BB" bold="true" box="[1225,1346,1656,1678]" italics="true" pageId="10" pageNumber="304">R. shannoni</emphasis>
Carrera
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

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<mods:title id="F28C921D4E00D4506D06475F546A1018">Three new species of the genus Rachicerus Walker (Diptera, Xylophagidae) from Brazil</mods:title>
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<mods:affiliation id="C1F480B1AB516EEE830F6626B1A623C8">Universidade de Brasília (UNB), Instituto de Ciências Biológicas, Departamento de Zoologia, Brasília, DF, Brazil. &amp; Universidade Federal do Amazonas (UFAM), Instituto de Ciências Biológicas, Departamento de Biologia, Manaus, AM, Brazil.</mods:affiliation>
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<subSubSection id="C3698790FFB9FFF2FF717412FCE85341" box="[151,782,1626,1653]" pageId="5" pageNumber="299" type="nomenclature">
<paragraph id="8BCCD41BFFB9FFF2FF717412FCE85341" blockId="5.[151,782,1626,1689]" box="[151,782,1626,1653]" pageId="5" pageNumber="299">
<heading id="D0846377FFB9FFF2FF717412FCE85341" bold="true" box="[151,782,1626,1653]" fontSize="11" level="1" pageId="5" pageNumber="299" reason="1">
<emphasis id="B9070809FFB9FFF2FF717412FCE85341" bold="true" box="[151,782,1626,1653]" pageId="5" pageNumber="299">
<taxonomicName id="4C73AF98FFB9FFF2FF717412FD975341" authority="Pujol-Luz &amp; Godoi" authorityName="Pujol-Luz &amp; Godoi" authorityYear="2025" box="[151,625,1626,1653]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="299" phylum="Arthropoda" rank="species" species="rafaeli" status="sp. nov.">
<emphasis id="B9070809FFB9FFF2FF717412FE915340" bold="true" box="[151,375,1626,1653]" italics="true" pageId="5" pageNumber="299">Rachicerus rafaeli</emphasis>
Pujol-Luz &amp; Godoi
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,
<taxonomicNameLabel id="A234B572FFB9FFF2FD987413FCE85341" box="[638,782,1626,1653]" pageId="5" pageNumber="299" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C3698790FFB9FFF2FF717437FEFF53AC" box="[151,281,1662,1689]" pageId="5" pageNumber="299" type="description">
<paragraph id="8BCCD41BFFB9FFF2FF717437FEFF53AC" blockId="5.[151,782,1626,1689]" box="[151,281,1662,1689]" pageId="5" pageNumber="299">
(
<figureCitation id="1348C89EFFB9FFF2FF797437FEF753AC" box="[159,273,1662,1689]" captionStart-0="FIGURES 6–8" captionStart-1="FIGURES 9–15" captionStartId-0="7.[151,264,1445,1469]" captionStartId-1="8.[151,264,1347,1371]" captionTargetBox-0="[360,1227,179,1417]" captionTargetBox-1="[349,1239,182,1321]" captionTargetId-0="figure-26@7.[360,1227,179,1419]" captionTargetId-1="figure-17@8.[349,1239,179,1321]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 6–8. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov. 6, Habitus in lateral view; 7, Frontal view of the diaphanized head (Paratype), showing the orbital triangle, 8, Left antenna (Paratype). (Holotype housed in CZMA, Paratype housed in INPA)." captionText-1="FIGURES 9–15. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov., genital capsule. 9, Hypandrium, gonostylus and gonocoxal apodeme; 10, Hypandrium, gonocoxal apodeme, gonocoxite and gonostylus; 11, Hypandrium and gonostylus; 12, Hypandrium and gonostylus; 13, Cercus; 14, Phallus; 15, Phallus with glans pore, distal view." pageId="5" pageNumber="299">Figs 6–15</figureCitation>
)
</paragraph>
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<subSubSection id="C3698790FFB9FFF2FF71748EFE085230" pageId="5" pageNumber="299" type="diagnosis">
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<emphasis id="B9070809FFB9FFF2FF71748EFEEC53D5" bold="true" box="[151,266,1734,1760]" pageId="5" pageNumber="299">Diagnosis</emphasis>
. Body blackish; femora, tibiae and first tarsomeres striped in black and white; antenna with flagellomeres pectinate slims and lanceolate.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB9FFF0FF217547FE44535C" lastPageId="7" lastPageNumber="301" pageId="5" pageNumber="299" type="description">
<paragraph id="8BCCD41BFFB9FFF1FF217547FE2457B0" blockId="5.[151,1437,1734,2013]" lastBlockId="6.[151,1437,151,645]" lastPageId="6" lastPageNumber="300" pageId="5" pageNumber="299">
<emphasis id="B9070809FFB9FFF2FF217547FE66521C" bold="true" box="[199,384,1807,1833]" pageId="5" pageNumber="299">Measurements:</emphasis>
body length:
<quantity id="4C8B79FEFFB9FFF2FDFE7547FD47521C" box="[536,673,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="8.8" metricValueMin="6.8" pageId="5" pageNumber="299" unit="mm" value="7.8" valueMax="8.8" valueMin="6.8">6.8–8.8 mm</quantity>
; antenna:
<quantity id="4C8B79FEFFB9FFF2FCF67547FC7F521C" box="[784,921,1807,1833]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="5" pageNumber="299" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.8–2.3 mm</quantity>
; number of antennal flagellomeres: 20; wings:
<quantity id="4C8B79FEFFB9FFF2FF71757AFEC65278" box="[151,288,1842,1869]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.6" metricValueMin="5.6" pageId="5" pageNumber="299" unit="mm" value="6.1" valueMax="6.6" valueMin="5.6">5.6–6.6 mm</quantity>
.
<emphasis id="B9070809FFB9FFF2FECC757BFE8C5278" bold="true" box="[298,362,1843,1869]" pageId="5" pageNumber="299">Head</emphasis>
: hemispherical, wider than thorax; front flat, shining black, with two longitudinal ridges running from ocellar plate and converging on suture that divides antennal plate in half; frontal plate large with small whitish setae (pruinosity like) around the antennae in inferior margin; fronto-orbital plate depressed and bare. Compound eyes dichoptic and bare; vertex with ocellar triangle developed, black; ocellus brownish yellow. Occiput shinning black well developed and very pilose (
<figureCitation id="1348C89EFFB9FFF2FDAE758BFD5552E8" box="[584,691,1987,2013]" captionStart="FIGURES 6–8" captionStartId="7.[151,264,1445,1469]" captionTargetBox="[360,1227,179,1417]" captionTargetId="figure-26@7.[360,1227,179,1419]" captionTargetPageId="7" captionText="FIGURES 6–8. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov. 6, Habitus in lateral view; 7, Frontal view of the diaphanized head (Paratype), showing the orbital triangle, 8, Left antenna (Paratype). (Holotype housed in CZMA, Paratype housed in INPA)." pageId="5" pageNumber="299">
Figs. 6
<emphasis id="B9070809FFB9FFF2FD73758BFD4552E8" bold="true" box="[661,675,1987,2013]" pageId="5" pageNumber="299">–</emphasis>
7
</figureCitation>
). Palps brownish, club-like and 2-segmented. Labellum brownish, membranous with elongate setae. Antennae elongate more than two times as long as width of head, with 20 spatulate flagellomeres, last flagellomeres truncate (
<figureCitation id="1348C89EFFBAFFF1FD9872F3FD0955E0" box="[638,751,187,213]" captionStart="FIGURES 6–8" captionStartId="7.[151,264,1445,1469]" captionTargetBox="[360,1227,179,1417]" captionTargetId="figure-26@7.[360,1227,179,1419]" captionTargetPageId="7" captionText="FIGURES 6–8. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov. 6, Habitus in lateral view; 7, Frontal view of the diaphanized head (Paratype), showing the orbital triangle, 8, Left antenna (Paratype). (Holotype housed in CZMA, Paratype housed in INPA)." pageId="6" pageNumber="300">Figs. 6, 8</figureCitation>
).
<emphasis id="B9070809FFBAFFF1FCE072F3FCB955E0" bold="true" box="[774,863,187,213]" pageId="6" pageNumber="300">Thorax</emphasis>
: predominantly black and bare; postpronotal lobe blackish; scutum with small and sparse pilosity; anepimeron whitish; laterotergites shinning black and entirely bare; scutellum blackish, with dense pilosity (
<figureCitation id="1348C89EFFBAFFF1FD40734BFD0A5428" box="[678,748,259,285]" captionStart="FIGURES 6–8" captionStartId="7.[151,264,1445,1469]" captionTargetBox="[360,1227,179,1417]" captionTargetId="figure-26@7.[360,1227,179,1419]" captionTargetPageId="7" captionText="FIGURES 6–8. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov. 6, Habitus in lateral view; 7, Frontal view of the diaphanized head (Paratype), showing the orbital triangle, 8, Left antenna (Paratype). (Holotype housed in CZMA, Paratype housed in INPA)." pageId="6" pageNumber="300">Fig. 6</figureCitation>
).
<emphasis id="B9070809FFBAFFF1FCE2734AFCB65429" bold="true" box="[772,848,258,284]" pageId="6" pageNumber="300">Wings</emphasis>
: hyaline, inconspicuous pterostigma between veins Sc and R
<subScript id="17F7D65EFFBAFFF1FF197371FEE15472" attach="left" box="[255,263,313,327]" fontSize="6" pageId="6" pageNumber="300">1</subScript>
; R
<subScript id="17F7D65EFFBAFFF1FECC7371FED55472" attach="left" box="[298,307,313,327]" fontSize="6" pageId="6" pageNumber="300">S</subScript>
stem curved upwards before bifurcation of radial sectors; halters whitish (
<figureCitation id="1348C89EFFBAFFF1FB63736FFB2D5474" box="[1157,1227,295,321]" captionStart="FIGURES 6–8" captionStartId="7.[151,264,1445,1469]" captionTargetBox="[360,1227,179,1417]" captionTargetId="figure-26@7.[360,1227,179,1419]" captionTargetPageId="7" captionText="FIGURES 6–8. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov. 6, Habitus in lateral view; 7, Frontal view of the diaphanized head (Paratype), showing the orbital triangle, 8, Left antenna (Paratype). (Holotype housed in CZMA, Paratype housed in INPA)." pageId="6" pageNumber="300">Fig. 6</figureCitation>
).
<emphasis id="B9070809FFBAFFF1FB04736FFAFC5474" bold="true" box="[1250,1306,295,321]" pageId="6" pageNumber="300">Legs</emphasis>
: coxae and trochanters black; fore leg with femora black, tibia and first tarsomeres white; mid and hind legs with femur and tibia with proximal half whitish and distal half black; first tarsomere of both legs whitish; empodia pulvilliform (
<figureCitation id="1348C89EFFBAFFF1FA897327FF415498" captionStart="FIGURES 6–8" captionStartId="7.[151,264,1445,1469]" captionTargetBox="[360,1227,179,1417]" captionTargetId="figure-26@7.[360,1227,179,1419]" captionTargetPageId="7" captionText="FIGURES 6–8. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov. 6, Habitus in lateral view; 7, Frontal view of the diaphanized head (Paratype), showing the orbital triangle, 8, Left antenna (Paratype). (Holotype housed in CZMA, Paratype housed in INPA)." pageId="6" pageNumber="300">Fig. 6</figureCitation>
).
<emphasis id="B9070809FFBAFFF1FF5873DAFED55499" bold="true" box="[190,307,402,428]" pageId="6" pageNumber="300">Abdomen</emphasis>
: entirely shinning dark black, with dense small pilosity; intersegmental membranes whitish, giving black and white striped appearance to abdomen; terminalia conical with dense pilosity (
<figureCitation id="1348C89EFFBAFFF1FB6B73FFFB3254E4" box="[1165,1236,439,465]" captionStart="FIGURES 6–8" captionStartId="7.[151,264,1445,1469]" captionTargetBox="[360,1227,179,1417]" captionTargetId="figure-26@7.[360,1227,179,1419]" captionTargetPageId="7" captionText="FIGURES 6–8. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov. 6, Habitus in lateral view; 7, Frontal view of the diaphanized head (Paratype), showing the orbital triangle, 8, Left antenna (Paratype). (Holotype housed in CZMA, Paratype housed in INPA)." pageId="6" pageNumber="300">Fig. 6</figureCitation>
). Genital capsule (epandrium and hypandrium) elliptical in general form; gonocoxal apodemes conspicuously long, flat and curved inwards, arising near base of gonostylus (
<figureCitation id="1348C89EFFBAFFF1FD8173B7FD06572C" box="[615,736,511,537]" captionStart="FIGURES 9–15" captionStartId="8.[151,264,1347,1371]" captionTargetBox="[349,1239,182,1321]" captionTargetId="figure-17@8.[349,1239,179,1321]" captionTargetPageId="8" captionText="FIGURES 9–15. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov., genital capsule. 9, Hypandrium, gonostylus and gonocoxal apodeme; 10, Hypandrium, gonocoxal apodeme, gonocoxite and gonostylus; 11, Hypandrium and gonostylus; 12, Hypandrium and gonostylus; 13, Cercus; 14, Phallus; 15, Phallus with glans pore, distal view." pageId="6" pageNumber="300">
Figs. 9
<emphasis id="B9070809FFBAFFF1FD5373B7FD25572C" bold="true" box="[693,707,511,537]" pageId="6" pageNumber="300">–</emphasis>
11
</figureCitation>
); phallus tubular, long and curved, moderately depressed from median surface to glans pore (
<figureCitation id="1348C89EFFBAFFF1FE17706BFD9D5708" box="[497,635,546,573]" captionStart="FIGURES 9–15" captionStartId="8.[151,264,1347,1371]" captionTargetBox="[349,1239,182,1321]" captionTargetId="figure-17@8.[349,1239,179,1321]" captionTargetPageId="8" captionText="FIGURES 9–15. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov., genital capsule. 9, Hypandrium, gonostylus and gonocoxal apodeme; 10, Hypandrium, gonocoxal apodeme, gonocoxite and gonostylus; 11, Hypandrium and gonostylus; 12, Hypandrium and gonostylus; 13, Cercus; 14, Phallus; 15, Phallus with glans pore, distal view." pageId="6" pageNumber="300">
Figs. 14
<emphasis id="B9070809FFBAFFF1FDA8706BFDBA5708" bold="true" box="[590,604,547,573]" pageId="6" pageNumber="300">–</emphasis>
15
</figureCitation>
); gonostylus long and curved, with upper tip slightly larger than basal tip; interior margin deeply concave, covered by small and sparse pilosity (
<figureCitation id="1348C89EFFBAFFF1FC34700FFBBC5754" box="[978,1114,583,609]" captionStart="FIGURES 9–15" captionStartId="8.[151,264,1347,1371]" captionTargetBox="[349,1239,182,1321]" captionTargetId="figure-17@8.[349,1239,179,1321]" captionTargetPageId="8" captionText="FIGURES 9–15. Rachicerus rafaeli Pujol-Luz &amp; Godoi sp. nov., genital capsule. 9, Hypandrium, gonostylus and gonocoxal apodeme; 10, Hypandrium, gonocoxal apodeme, gonocoxite and gonostylus; 11, Hypandrium and gonostylus; 12, Hypandrium and gonostylus; 13, Cercus; 14, Phallus; 15, Phallus with glans pore, distal view." pageId="6" pageNumber="300">
Figs. 12
<emphasis id="B9070809FFBAFFF1FBCB700FFBDD5754" bold="true" box="[1069,1083,583,609]" pageId="6" pageNumber="300">–</emphasis>
13
</figureCitation>
). Cercus long, spoon-shaped with small, sparse pilosity.
</paragraph>
<caption id="DF0C8493FFBAFFF1FF7175F4FCD852CC" pageId="6" pageNumber="300" startId="6.[151,264,1980,2004]" targetBox="[356,1232,666,1945]" targetPageId="6" targetType="figure">
<paragraph id="8BCCD41BFFBAFFF1FF7175F4FCD852CC" blockId="6.[151,1436,1980,2041]" pageId="6" pageNumber="300">
<emphasis id="B9070809FFBAFFF1FF7175F4FED252E0" bold="true" box="[151,308,1980,2005]" pageId="6" pageNumber="300">FIGURES 3–5</emphasis>
.
<taxonomicName id="4C73AF98FFBAFFF1FEA775F5FD9C52E0" authority="Pujol-Luz" authorityName="Pujol-Luz &amp; Godoi" authorityYear="2025" box="[321,634,1981,2005]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="300" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="B9070809FFBAFFF1FEA775F5FDE852E0" box="[321,526,1981,2005]" italics="true" pageId="6" pageNumber="300">Rachicerus isabelae</emphasis>
Pujol-Luz
</taxonomicName>
<emphasis id="B9070809FFBAFFF1FD6675F5FD3752E0" bold="true" box="[640,721,1981,2005]" pageId="6" pageNumber="300">
<taxonomicNameLabel id="A234B572FFBAFFF1FD6675F5FD3752E0" box="[640,721,1981,2005]" pageId="6" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
3, Habitus in lateral view; 4, Fronto-lateral view of the head, showing the orbital triangle; 5, Left antenna. (Holotype housed in MZUSP).
</paragraph>
</caption>
<caption id="DF0C8493FFBBFFF0FF7177EDFF065330" pageId="7" pageNumber="301" startId="7.[151,264,1445,1469]" targetBox="[360,1227,179,1417]" targetPageId="7" targetType="figure">
<paragraph id="8BCCD41BFFBBFFF0FF7177EDFF065330" blockId="7.[151,1436,1445,1541]" pageId="7" pageNumber="301">
<emphasis id="B9070809FFBBFFF0FF7177EDFED35088" bold="true" box="[151,309,1445,1469]" pageId="7" pageNumber="301">FIGURES 6–8</emphasis>
.
<taxonomicName id="4C73AF98FFBBFFF0FEA477EDFD2A5088" authority="Pujol-Luz &amp; Godoi" authorityName="Pujol-Luz &amp; Godoi" authorityYear="2025" box="[322,716,1445,1469]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="301" phylum="Arthropoda" rank="species" species="rafaeli" status="sp. nov.">
<emphasis id="B9070809FFBBFFF0FEA477EDFE195088" box="[322,511,1445,1469]" italics="true" pageId="7" pageNumber="301">Rachicerus rafaeli</emphasis>
Pujol-Luz &amp; Godoi
</taxonomicName>
<emphasis id="B9070809FFBBFFF0FD3277EDFCC35088" bold="true" box="[724,805,1445,1469]" pageId="7" pageNumber="301">
<taxonomicNameLabel id="A234B572FFBBFFF0FD3277EDFCC35088" box="[724,805,1445,1469]" pageId="7" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
6, Habitus in lateral view; 7, Frontal view of the diaphanized head (Paratype), showing the orbital triangle, 8, Left antenna (Paratype). (Holotype housed in CZMA, Paratype housed in INPA).
</paragraph>
</caption>
<paragraph id="8BCCD41BFFBBFFF0FF217407FE44535C" blockId="7.[151,1437,1615,2038]" box="[199,418,1615,1641]" pageId="7" pageNumber="301">
<emphasis id="B9070809FFBBFFF0FF217407FEC3535C" bold="true" box="[199,293,1615,1641]" pageId="7" pageNumber="301">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFBBFFF0FF21743BFBE053E0" pageId="7" pageNumber="301" type="etymology">
<paragraph id="8BCCD41BFFBBFFF0FF21743BFF1D5384" blockId="7.[151,1437,1615,2038]" pageId="7" pageNumber="301">
<emphasis id="B9070809FFBBFFF0FF21743BFEAE53B8" bold="true" box="[199,328,1651,1677]" pageId="7" pageNumber="301">Etymology</emphasis>
: This name is given to honor Professor José Albertino Rafael, our friend and illustrious Brazilian dipterist.
</paragraph>
<paragraph id="8BCCD41BFFBBFFF0FF2174F3FBE053E0" blockId="7.[151,1437,1615,2038]" box="[199,1030,1723,1750]" pageId="7" pageNumber="301">
<emphasis id="B9070809FFBBFFF0FF2174F3FE1853E0" bold="true" box="[199,510,1723,1749]" pageId="7" pageNumber="301">Geographical distribution</emphasis>
.
<collectingCountry id="F364948BFFBBFFF0FDEA74F4FD9253E3" box="[524,628,1724,1750]" name="Brazil" pageId="7" pageNumber="301">BRAZIL</collectingCountry>
, states of Amazonas and
<collectingRegion id="49B71AF9FFBBFFF0FC6B74F3FBE453E0" box="[909,1026,1723,1749]" country="Brazil" name="Maranhao" pageId="7" pageNumber="301">Maranhão</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFBBFFF0FF217497FC9652C0" pageId="7" pageNumber="301" type="materials_examined">
<paragraph id="8BCCD41BFFBBFFF0FF217497FC9652C0" blockId="7.[151,1437,1615,2038]" pageId="7" pageNumber="301">
<emphasis id="B9070809FFBBFFF0FF217497FE9653CC" bold="true" box="[199,368,1759,1785]" pageId="7" pageNumber="301">Type material</emphasis>
.
<materialsCitation id="3B1BDE46FFBBFFF0FE677497FC8A52C0" collectingDate="1988-08-23" collectingDateMax="2018-12-31" collectingDateMin="1988-08-23" collectionCode="CZMA, INPA" collectorName="F. Limeira-de-Oliveira &amp; D. Limeira &amp; J. S. Brito &amp; The &amp; Manaus &amp; Amazonas &amp; Km NE Manaus &amp; Reserva Ducke &amp; J. A. Rafael &amp; Socorro Rosa" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" location="The" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" pageId="7" pageNumber="301" specimenCode="CZMA600" specimenCount="5" stateProvince="Maranhao" typeStatus="holotype">
<location id="8EAC82C0FFBBFFF0FE677497FE4B53CC" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FE677497FE4B53CC" box="[385,429,1759,1785]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="The" pageId="7" pageNumber="301" stateProvince="Maranhao">The</location>
male
<typeStatus id="54C86AB9FFBBFFF0FE107497FD6C53CC" box="[502,650,1759,1785]" pageId="7" pageNumber="301" type="holotype">HOLOTYPE</typeStatus>
is housed in the
<location id="8EAC82C0FFBBFFF0FCB27497FBCD53CC" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FCB27497FBCD53CC" box="[852,1067,1759,1785]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="Colecao Zoologica" pageId="7" pageNumber="301" stateProvince="Maranhao">Coleção Zoológica</location>
do
<collectingRegion id="49B71AF9FFBBFFF0FBBC7497FB2A53CC" box="[1114,1228,1759,1785]" country="Brazil" name="Maranhao" pageId="7" pageNumber="301">Maranhão</collectingRegion>
(
<collectionCode id="ED624CDEFFBBFFF0FB3B7497FAD253CC" box="[1245,1332,1759,1785]" pageId="7" pageNumber="301">CZMA</collectionCode>
),
<location id="8EAC82C0FFBBFFF0FAAF7497FA7E53CC" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FAAF7497FA7E53CC" box="[1353,1432,1759,1785]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="Caxias" pageId="7" pageNumber="301" stateProvince="Maranhao">Caxias</location>
,
<collectingRegion id="49B71AF9FFBBFFF0FF71754BFEED5228" box="[151,267,1795,1821]" country="Brazil" name="Maranhao" pageId="7" pageNumber="301">Maranhão</collectingRegion>
,
<collectingCountry id="F364948BFFBBFFF0FEF0754BFEB85228" box="[278,350,1795,1821]" name="Brazil" pageId="7" pageNumber="301">Brazil</collectingCountry>
.
<location id="8EAC82C0FFBBFFF0FE8E754BFE215228" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FE8E754BFE215228" box="[360,455,1795,1821]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="Labeled" pageId="7" pageNumber="301" stateProvince="Maranhao">Labeled</location>
: Brasil (MA), C[ândido]
<location id="8EAC82C0FFBBFFF0FD39754BFCDD5228" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FD39754BFCDD5228" box="[735,827,1795,1821]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="Mendes" pageId="7" pageNumber="301" stateProvince="Maranhao">Mendes</location>
,
<location id="8EAC82C0FFBBFFF0FCA3754BFC445228" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FCA3754BFC445228" box="[837,930,1795,1821]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="Fazenda" pageId="7" pageNumber="301" stateProvince="Maranhao">Fazenda</location>
7
<location id="8EAC82C0FFBBFFF0FC5B754CFBE8522B" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FC5B754CFBE8522B" box="[957,1038,1796,1822]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="Irmaos" pageId="7" pageNumber="301" stateProvince="Maranhao">Irmãos</location>
,
<geoCoordinate id="EE47B2DCFFBBFFF0FBFE754BFB7A522B" box="[1048,1180,1795,1822]" degrees="01" direction="south" minutes="52" orientation="latitude" pageId="7" pageNumber="301" precision="15" seconds="06" value="-1.8683333">
01
<superScript id="7C067953FFBBFFF0FBD2754BFBDA5224" attach="none" box="[1076,1084,1795,1809]" fontSize="6" pageId="7" pageNumber="301">o</superScript>
52’06”S
</geoCoordinate>
,
<geoCoordinate id="EE47B2DCFFBBFFF0FB41754BFAAE522B" box="[1191,1352,1795,1822]" degrees="45" direction="west" minutes="45" orientation="longitude" pageId="7" pageNumber="301" precision="1" seconds="59.7" value="-45.766582">
45
<superScript id="7C067953FFBBFFF0FB25754BFB2D5224" attach="none" box="[1219,1227,1795,1809]" fontSize="6" pageId="7" pageNumber="301">o</superScript>
45’59.7”W
</geoCoordinate>
[White label]/
<location id="8EAC82C0FFBBFFF0FF00756FFEFB5274" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FF00756FFEFB5274" box="[230,285,1831,1857]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="Arm" pageId="7" pageNumber="301" stateProvince="Maranhao">Arm</location>
[adilha]
<location id="8EAC82C0FFBBFFF0FE9F756FFE325274" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF0FF717412FC9652C0:8EAC82C0FFBBFFF0FE9F756FFE325274" box="[377,468,1831,1857]" country="Brazil" county="Invertebrados do Instituto Nacional de Pesquisas da Amazonia" latitude="-1.8683333" longLatPrecision="15" longitude="-45.766582" municipality="Colecao de Invertebrados do Instituto Nacional de Pesquisas" name="Malaise" pageId="7" pageNumber="301" stateProvince="Maranhao">Malaise</location>
,
<date id="FFCDF2DBFFBBFFF0FE07756FFD6E5277" box="[481,648,1831,1858]" pageId="7" pageNumber="301" value="2018-12-15" valueMax="2018-12-31" valueMin="2018-12-15">
<collectingDate id="EF890B33FFBBFFF0FE07756FFD6E5277" box="[481,648,1831,1858]" pageId="7" pageNumber="301" value="2018-12-15" valueMax="2018-12-31" valueMin="2018-12-15">15-31.xii.2018</collectingDate>
</date>
,
<collectorName id="2686B1CDFFBBFFF0FD707560FC715274" box="[662,919,1831,1858]" pageId="7" pageNumber="301">F. Limeira-de-Oliveira</collectorName>
,
<collectorName id="2686B1CDFFBBFFF0FC427560FBC75274" box="[932,1057,1831,1858]" pageId="7" pageNumber="301">D. Limeira</collectorName>
&amp;
<collectorName id="2686B1CDFFBBFFF0FBA1756FFB525274" box="[1095,1204,1831,1857]" pageId="7" pageNumber="301">J.S. Brito</collectorName>
, cols. [White label]/
<typeStatus id="54C86AB9FFBBFFF0FF717503FF1D5250" box="[151,251,1867,1893]" pageId="7" pageNumber="301" type="holotype">Holótipo</typeStatus>
[Red label]/
<specimenCode id="DBD57C60FFBBFFF0FE6D7503FDE35250" box="[395,517,1867,1893]" collectionCode="CZMA" pageId="7" pageNumber="301">CZMA600</specimenCode>
[White label].
<collectorName id="2686B1CDFFBBFFF0FD4A7503FD3E5250" box="[684,728,1867,1893]" pageId="7" pageNumber="301">The</collectorName>
<typeStatus id="54C86AB9FFBBFFF0FD387503FCA65250" box="[734,832,1867,1893]" pageId="7" pageNumber="301" type="holotype">holotype</typeStatus>
is in good condition, only missing the lack of the right fore leg, the mid legs and the tips of the antennae.
<typeStatus id="54C86AB9FFBBFFF0FD077538FC8A52BF" box="[737,876,1904,1930]" pageId="7" pageNumber="301" type="paratype">PARATYPE</typeStatus>
male housed in the
<collectingMunicipality id="6BA84E61FFBBFFF0FBB37527FE125298" pageId="7" pageNumber="301">
Coleção de
<collectingCounty id="62ADAC97FFBBFFF0FB3B7527FE125298" pageId="7" pageNumber="301">Invertebrados do Instituto Nacional de Pesquisas</collectingCounty>
</collectingMunicipality>
da Amazônia (
<collectionCode id="ED624CDEFFBBFFF0FD7C75DBFD395298" box="[666,735,1939,1965]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="7" pageNumber="301" type="">INPA</collectionCode>
),
<collectorName id="2686B1CDFFBBFFF0FD1775DCFCAA529B" box="[753,844,1940,1966]" pageId="7" pageNumber="301">Manaus</collectorName>
,
<collectorName id="2686B1CDFFBBFFF0FCB375DCFC28529B" box="[853,974,1940,1966]" pageId="7" pageNumber="301">Amazonas</collectorName>
,
<collectingCountry id="F364948BFFBBFFF0FC3F75DBFBC75298" box="[985,1057,1939,1965]" name="Brazil" pageId="7" pageNumber="301">Brazil</collectingCountry>
.
<collectorName id="2686B1CDFFBBFFF0FBCC75DBFBB05298" box="[1066,1110,1939,1965]" pageId="7" pageNumber="301">The</collectorName>
specimen is fixed in glycerol in a microvial, labeled: Brasil,
<collectorName id="2686B1CDFFBBFFF0FE1A75F0FD9352E7" box="[508,629,1976,2002]" pageId="7" pageNumber="301">Amazonas</collectorName>
, 26
<collectorName id="2686B1CDFFBBFFF0FD4E75F0FC8052E7" box="[680,870,1976,2002]" pageId="7" pageNumber="301">Km NE Manaus</collectorName>
,
<collectorName id="2686B1CDFFBBFFF0FC9275F0FBC252E4" box="[884,1060,1975,2002]" pageId="7" pageNumber="301">Reserva Ducke</collectorName>
,
<date id="FFCDF2DBFFBBFFF0FBD775FFFB2552E4" box="[1073,1219,1975,2001]" pageId="7" pageNumber="301" value="1988-08-23">
<collectingDate id="EF890B33FFBBFFF0FBD775FFFB2552E4" box="[1073,1219,1975,2001]" pageId="7" pageNumber="301" value="1988-08-23">23.VIII.1988</collectingDate>
</date>
[white label]/
<collectorName id="2686B1CDFFBBFFF0FA8975F0FF0652C0" pageId="7" pageNumber="301">J.A. Rafael</collectorName>
&amp;
<collectorName id="2686B1CDFFBBFFF0FEE27593FE7A52C3" box="[260,412,2011,2038]" pageId="7" pageNumber="301">Socorro Rosa</collectorName>
col. [white label]/
<typeStatus id="54C86AB9FFBBFFF0FD967593FD2B52C0" box="[624,717,2011,2037]" pageId="7" pageNumber="301" type="paratype">Parátipo</typeStatus>
[Green label]
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

186
data/03/DA/65/03DA650DFFB9FFF2FF7170C6FE20533C.xml Normal file
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@ -0,0 +1,186 @@
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<mods:title id="8D51ED536F59B5381E80F88A57D888DE">Three new species of the genus Rachicerus Walker (Diptera, Xylophagidae) from Brazil</mods:title>
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<mods:namePart id="DA7BF8B4FBB688AB3DD88984B4DBE42F">Pujol-Luz, José Roberto</mods:namePart>
<mods:affiliation id="3B7C8C7E0C532434377069C7CD0D8B73">Universidade de Brasília (UNB), Instituto de Ciências Biológicas, Departamento de Zoologia, Brasília, DF, Brazil.</mods:affiliation>
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<mods:namePart id="3329889F95E32B1C24DD297565BDF457">Godoi, Fabio Siqueira Pitaluga De</mods:namePart>
<mods:affiliation id="AEAECF0BA15188061BD3B36271A5E312">Universidade de Brasília (UNB), Instituto de Ciências Biológicas, Departamento de Zoologia, Brasília, DF, Brazil. &amp; Universidade Federal do Amazonas (UFAM), Instituto de Ciências Biológicas, Departamento de Biologia, Manaus, AM, Brazil.</mods:affiliation>
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<treatment id="03DA650DFFB9FFF2FF7170C6FE20533C" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF2FF7170C6FE20533C" httpUri="http://treatment.plazi.org/id/03DA650DFFB9FFF2FF7170C6FE20533C" lastPageNumber="299" pageId="5" pageNumber="299">
<subSubSection id="C3698790FFB9FFF2FF7170C6FD4B579D" box="[151,685,654,681]" pageId="5" pageNumber="299" type="nomenclature">
<paragraph id="8BCCD41BFFB9FFF2FF7170C6FD4B579D" blockId="5.[151,685,654,717]" box="[151,685,654,681]" pageId="5" pageNumber="299">
<heading id="D0846377FFB9FFF2FF7170C6FD4B579D" bold="true" box="[151,685,654,681]" fontSize="11" level="1" pageId="5" pageNumber="299" reason="1">
<emphasis id="B9070809FFB9FFF2FF7170C6FD4B579D" bold="true" box="[151,685,654,681]" pageId="5" pageNumber="299">
<taxonomicName id="4C73AF98FFB9FFF2FF7170C6FDF6579D" authority="Pujol-Luz" authorityName="Pujol-Luz &amp; Godoi" authorityYear="2025" box="[151,528,654,681]" class="Insecta" family="Xylophagidae" genus="Rachicerus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="299" phylum="Arthropoda" rank="species" species="isabelae" status="sp. nov.">
<emphasis id="B9070809FFB9FFF2FF7170C6FE6F579C" bold="true" box="[151,393,654,681]" italics="true" pageId="5" pageNumber="299">Rachicerus isabelae</emphasis>
Pujol-Luz
</taxonomicName>
,
<taxonomicNameLabel id="A234B572FFB9FFF2FDFB70C7FD4B579D" box="[541,685,654,681]" pageId="5" pageNumber="299" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB9FFF2FF7170FBFEED57F8" box="[151,267,690,717]" pageId="5" pageNumber="299" type="description">
<paragraph id="8BCCD41BFFB9FFF2FF7170FBFEED57F8" blockId="5.[151,685,654,717]" box="[151,267,690,717]" pageId="5" pageNumber="299">
(
<figureCitation id="1348C89EFFB9FFF2FF7970FBFEE457F8" box="[159,258,690,717]" captionStart="FIGURES 3–5" captionStartId="6.[151,264,1980,2004]" captionTargetBox="[356,1232,666,1945]" captionTargetId="figure-246@6.[356,1232,666,1954]" captionTargetPageId="6" captionText="FIGURES 3–5. Rachicerus isabelae Pujol-Luz sp. nov. 3, Habitus in lateral view; 4, Fronto-lateral view of the head, showing the orbital triangle; 5, Left antenna. (Holotype housed in MZUSP)." pageId="5" pageNumber="299">Figs 3–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB9FFF2FF7170B2FAE85620" box="[151,1294,762,789]" pageId="5" pageNumber="299" type="diagnosis">
<paragraph id="8BCCD41BFFB9FFF2FF7170B2FAE85620" blockId="5.[151,1437,762,1545]" box="[151,1294,762,789]" pageId="5" pageNumber="299">
<emphasis id="B9070809FFB9FFF2FF7170B2FEEC5621" bold="true" box="[151,266,762,788]" pageId="5" pageNumber="299">Diagnosis</emphasis>
. Body dark brownish; yellow cerci; antenna with pectinate falciform-shaped flagellomeres.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB9FFF2FF217157FE6E5038" pageId="5" pageNumber="299" type="description">
<paragraph id="8BCCD41BFFB9FFF2FF217157FA7B560C" blockId="5.[151,1437,762,1545]" box="[199,1437,799,825]" pageId="5" pageNumber="299">
<emphasis id="B9070809FFB9FFF2FF217157FE66560C" bold="true" box="[199,384,799,825]" pageId="5" pageNumber="299">Measurements:</emphasis>
body length:
<quantity id="4C8B79FEFFB9FFF2FDFA7157FD93560C" box="[540,629,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="5" pageNumber="299" unit="mm" value="8.7">8.7 mm</quantity>
; antenna:
<quantity id="4C8B79FEFFB9FFF2FD0E7157FCA7560C" box="[744,833,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="299" unit="mm" value="1.3">1.3 mm</quantity>
; number of antennal flagellomeres: 16; wings:
<quantity id="4C8B79FEFFB9FFF2FAB27157FA7F560C" box="[1364,1433,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="299" unit="mm" value="7.0">7 mm</quantity>
.
</paragraph>
<paragraph id="8BCCD41BFFB9FFF2FF71710BFCE751DC" blockId="5.[151,1437,762,1545]" pageId="5" pageNumber="299">
<emphasis id="B9070809FFB9FFF2FF71710BFF315668" bold="true" box="[151,215,835,861]" pageId="5" pageNumber="299">Head</emphasis>
: hemispherical, wider than thorax; front flat, shining brown, with two longitudinal ridges running from ocellar plate and converging on a suture that divides antennal plate in half; frontal plate large with small whitish setae (pruinosity like) ventrally, around antennae; fronto-orbital plate depressed and bare. Compound eyes dichoptic and bare; vertex brownish with ocellar triangle developed with tuffs of small bristles; ocellus brownish yellow. Occiput brownish, well developed, and pilose. Palpus yellowish, club-like and bi-segmented. Labellum yellowish and membranous with elongate setae. Antennae elongate, more than two times longer than width of head, with 15 pectinate falciform-shaped flagellomeres, and last one club-like (
<figureCitation id="1348C89EFFB9FFF2FC997653FC0B5100" box="[895,1005,1050,1077]" captionStart="FIGURES 3–5" captionStartId="6.[151,264,1980,2004]" captionTargetBox="[356,1232,666,1945]" captionTargetId="figure-246@6.[356,1232,666,1954]" captionTargetPageId="6" captionText="FIGURES 3–5. Rachicerus isabelae Pujol-Luz sp. nov. 3, Habitus in lateral view; 4, Fronto-lateral view of the head, showing the orbital triangle; 5, Left antenna. (Holotype housed in MZUSP)." pageId="5" pageNumber="299">Figs. 3–5</figureCitation>
).
<emphasis id="B9070809FFB9FFF2FBE27653FBBB5100" bold="true" box="[1028,1117,1051,1077]" pageId="5" pageNumber="299">Thorax</emphasis>
: predominantly dark brown and bare; postpronotal lobe yellowish white; scutum with sparse pilosity; pleuron entirely bare; scutellum brownish, with dense pilosity.
<emphasis id="B9070809FFB9FFF2FE9E762AFE225149" bold="true" box="[376,452,1122,1148]" pageId="5" pageNumber="299">Wings</emphasis>
: hyaline, inconspicuous pterostigma between veins Sc and R
<subScript id="17F7D65EFFB9FFF2FB95763DFB9D51B6" attach="left" box="[1139,1147,1141,1155]" fontSize="6" pageId="5" pageNumber="299">1</subScript>
; R
<subScript id="17F7D65EFFB9FFF2FB7B763DFB4051B6" attach="left" box="[1181,1190,1141,1155]" fontSize="6" pageId="5" pageNumber="299">S</subScript>
stem curved upwards before bifurcation of radial sectors; halters brownish yellow.
<emphasis id="B9070809FFB9FFF2FCA476CFFC9C5194" bold="true" box="[834,890,1159,1185]" pageId="5" pageNumber="299">Legs</emphasis>
: coxae, trochanters, femurs and tibiae brownish; first tarsomeres whitish; empodia pulvilliform.
<emphasis id="B9070809FFB9FFF2FD4076E2FCFD51F1" bold="true" box="[678,795,1194,1220]" pageId="5" pageNumber="299">Abdomen</emphasis>
: entirely dark brownish, with dense small pilosity, except in second tergite; cercus yellowish, with dense pilosity.
</paragraph>
<paragraph id="8BCCD41BFFB9FFF2FF2176BBFE6E5038" blockId="5.[151,1437,762,1545]" box="[199,392,1267,1293]" pageId="5" pageNumber="299">
<emphasis id="B9070809FFB9FFF2FF2176BBFEED5038" bold="true" box="[199,267,1267,1293]" pageId="5" pageNumber="299">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB9FFF2FF21775FFC655060" pageId="5" pageNumber="299" type="etymology">
<paragraph id="8BCCD41BFFB9FFF2FF21775FFC075004" blockId="5.[151,1437,762,1545]" box="[199,993,1303,1329]" pageId="5" pageNumber="299">
<emphasis id="B9070809FFB9FFF2FF21775FFEAE5004" bold="true" box="[199,328,1303,1329]" pageId="5" pageNumber="299">Etymology</emphasis>
: This name is given to honor my beloved daughter Isabela.
</paragraph>
<paragraph id="8BCCD41BFFB9FFF2FF217772FC655060" blockId="5.[151,1437,762,1545]" box="[199,899,1338,1365]" pageId="5" pageNumber="299">
<emphasis id="B9070809FFB9FFF2FF217772FE185061" bold="true" box="[199,510,1338,1364]" pageId="5" pageNumber="299">Geographical distribution</emphasis>
.
<collectingCountry id="F364948BFFB9FFF2FDEA7773FD925060" box="[524,628,1339,1365]" name="Brazil" pageId="5" pageNumber="299">BRAZIL</collectingCountry>
, state of
<collectingRegion id="49B71AF9FFB9FFF2FD317773FC665060" box="[727,896,1339,1365]" country="Brazil" name="Santa Catarina" pageId="5" pageNumber="299">Santa Catarina</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3698790FFB9FFF2FF217717FE20533C" pageId="5" pageNumber="299" type="materials_examined">
<paragraph id="8BCCD41BFFB9FFF2FF217717FE20533C" blockId="5.[151,1437,762,1545]" pageId="5" pageNumber="299">
<emphasis id="B9070809FFB9FFF2FF217717FE89504D" bold="true" box="[199,367,1374,1401]" pageId="5" pageNumber="299">Type material</emphasis>
.
<materialsCitation id="3B1BDE46FFB9FFF2FE997717FE24533C" collectingDate="1962-11" collectionCode="MZUSP" collectorName="Fritz Plaumann &amp; White &amp; Red" country="Brazil" county="Nova Teutonia" latitude="-27.183332" location="The" longLatPrecision="1238" longitude="-52.383335" municipality="Labeled" pageId="5" pageNumber="299" specimenCode="MZ053524" specimenCount="3" stateProvince="Sao Paulo" typeStatus="holotype">
<location id="8EAC82C0FFB9FFF2FE997717FE4D504C" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF2FF7170C6FE20533C:8EAC82C0FFB9FFF2FE997717FE4D504C" box="[383,427,1375,1401]" country="Brazil" county="Nova Teutonia" latitude="-27.183332" longLatPrecision="1238" longitude="-52.383335" municipality="Labeled" name="The" pageId="5" pageNumber="299" stateProvince="Sao Paulo">The</location>
female
<typeStatus id="54C86AB9FFB9FFF2FDE17717FD7D504C" box="[519,667,1375,1401]" pageId="5" pageNumber="299" type="holotype">HOLOTYPE</typeStatus>
is housed in the
<location id="8EAC82C0FFB9FFF2FC867717FB18504C" LSID="urn:lsid:plazi:treatment:03DA650DFFB9FFF2FF7170C6FE20533C:8EAC82C0FFB9FFF2FC867717FB18504C" box="[864,1278,1375,1401]" country="Brazil" county="Nova Teutonia" latitude="-27.183332" longLatPrecision="1238" longitude="-52.383335" municipality="Labeled" name="Museu de Zoologia da Universidade" pageId="5" pageNumber="299" stateProvince="Sao Paulo">Museu de Zoologia da Universidade</location>
de
<collectingRegion id="49B71AF9FFB9FFF2FACC7717FA7A504C" box="[1322,1436,1375,1401]" country="Brazil" name="Sao Paulo" pageId="5" pageNumber="299">São Paulo</collectingRegion>
(
<collectionCode id="ED624CDEFFB9FFF2FF7977CBFEE450A8" box="[159,258,1411,1437]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="299" type="">MZUSP</collectionCode>
),
<collectingRegion id="49B71AF9FFB9FFF2FEF177CBFE6850A8" box="[279,398,1411,1437]" country="Brazil" name="Sao Paulo" pageId="5" pageNumber="299">São Paulo</collectingRegion>
,
<collectingRegion id="49B71AF9FFB9FFF2FE7C77CBFDF750A8" box="[410,529,1411,1437]" country="Brazil" name="Sao Paulo" pageId="5" pageNumber="299">São Paulo</collectingRegion>
,
<collectingCountry id="F364948BFFB9FFF2FDF877CBFD8050A8" box="[542,614,1411,1437]" name="Brazil" pageId="5" pageNumber="299">Brazil</collectingCountry>
.
<collectingMunicipality id="6BA84E61FFB9FFF2FD9477CBFD3750A8" box="[626,721,1411,1437]" pageId="5" pageNumber="299">Labeled</collectingMunicipality>
: Brasilien,
<collectingCounty id="62ADAC97FFB9FFF2FCB577CBFC1C50A8" box="[851,1018,1411,1437]" pageId="5" pageNumber="299">Nova Teutonia</collectingCounty>
[
<collectingRegion id="49B71AF9FFB9FFF2FBEF77CBFB5E50A8" box="[1033,1208,1411,1437]" country="Brazil" name="Santa Catarina" pageId="5" pageNumber="299">Santa Catarina</collectingRegion>
],
<geoCoordinate id="EE47B2DCFFB9FFF2FB2A77CBFAC350A8" box="[1228,1317,1411,1437]" degrees="27" direction="south" minutes="11" orientation="latitude" pageId="5" pageNumber="299" precision="925" value="-27.183332">
27
<superScript id="7C067953FFB9FFF2FB0E77CBFB1650A4" attach="none" box="[1256,1264,1411,1425]" fontSize="6" pageId="5" pageNumber="299">o</superScript>
11’S
</geoCoordinate>
,
<geoCoordinate id="EE47B2DCFFB9FFF2FAD277CAFA7E50A8" box="[1332,1432,1410,1437]" degrees="52" direction="west" minutes="23" orientation="longitude" pageId="5" pageNumber="299" precision="925" value="-52.383335">
52
<superScript id="7C067953FFB9FFF2FAB677CBFABE50A4" attach="none" box="[1360,1368,1411,1425]" fontSize="6" pageId="5" pageNumber="299">o</superScript>
23’W
</geoCoordinate>
,
<collectorName id="2686B1CDFFB9FFF2FF7177EFFEA550F4" box="[151,323,1447,1473]" pageId="5" pageNumber="299">Fritz Plaumann</collectorName>
col.,
<date id="FFCDF2DBFFB9FFF2FE6777EFFE3850F4" box="[385,478,1447,1473]" pageId="5" pageNumber="299" value="1962-11">
<collectingDate id="EF890B33FFB9FFF2FE6777EFFE3850F4" box="[385,478,1447,1473]" pageId="5" pageNumber="299" value="1962-11">XI.1962</collectingDate>
</date>
(300–500 meters) [
<collectorName id="2686B1CDFFB9FFF2FD5177EFFD1950F4" box="[695,767,1447,1473]" pageId="5" pageNumber="299">White</collectorName>
label]/
<typeStatus id="54C86AB9FFB9FFF2FCB277EFFC5E50F4" box="[852,952,1447,1473]" pageId="5" pageNumber="299" type="holotype">Holótipo</typeStatus>
[
<collectorName id="2686B1CDFFB9FFF2FC2377EFFC1350F4" box="[965,1013,1447,1473]" pageId="5" pageNumber="299">Red</collectorName>
label]/
<specimenCode id="DBD57C60FFB9FFF2FBAC77EEFB2E50F4" box="[1098,1224,1446,1473]" collectionCode="MZ" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="5" pageNumber="299">MZ053524</specimenCode>
[White label]. The
<typeStatus id="54C86AB9FFB9FFF2FF717783FF1F50D0" box="[151,249,1483,1509]" pageId="5" pageNumber="299" type="holotype">holotype</typeStatus>
is in good condition, except for the damaged left wing and the right wing, which has been separated and dried stored in a microvial
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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218
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<emphasis id="B93FB041FFD1FFADFF7CF996E377F9EE" bold="true" box="[151,536,1540,1567]" italics="true" pageId="10" pageNumber="396">Euthalia thawgawa wuzhenjuni</emphasis>
H.
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<figureCitation id="137070D6FFD1FFADFF7CF9BAE049F9B3" box="[151,294,1576,1602]" captionStart="FIGURES 46–49" captionStartId="8.[151,264,1481,1505]" captionTargetBox="[151,1435,273,1458]" captionTargetId="figure-52@8.[151,1435,273,1458]" captionTargetPageId="8" captionText="FIGURES 46–49. Females of Euthalia ssp., Depositary of the specimens: 46 in CHH, 47 &amp; 49 in CHSY, 48 in IPBX. a=upperside, b=underside" pageId="10" pageNumber="396">Fig. 47 &amp; 48</figureCitation>
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<emphasis id="B93FB041FFD1FFADFF7CF9FFE0B9F974" box="[151,470,1645,1669]" italics="true" pageId="10" pageNumber="396">Euthalia thawgawa wuzhenjuni</emphasis>
H.
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, figs. 240 &amp; 255.
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<emphasis id="B93FB041FFD1FFADFF7CF920E010F93D" bold="true" box="[151,383,1714,1740]" pageId="10" pageNumber="396">Material examined.</emphasis>
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,
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, leg.
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, leg.
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<emphasis id="B93FB041FFD1FFADFF2CF969E054F8E4" bold="true" box="[199,315,1787,1813]" pageId="10" pageNumber="396">Remarks.</emphasis>
This is the first record of the female of this recently described taxon, which inhabits only in the Nujiang Valley. Compared with the females of the nominate subspecies from Dulongjiang Valley (
<figureCitation id="137070D6FFD1FFADFB2EF88DE479F8C8" box="[1221,1302,1823,1849]" captionStart="FIGURES 46–49" captionStartId="8.[151,264,1481,1505]" captionTargetBox="[151,1435,273,1458]" captionTargetId="figure-52@8.[151,1435,273,1458]" captionTargetPageId="8" captionText="FIGURES 46–49. Females of Euthalia ssp., Depositary of the specimens: 46 in CHH, 47 &amp; 49 in CHSY, 48 in IPBX. a=upperside, b=underside" pageId="10" pageNumber="396">Fig. 46</figureCitation>
), Northwest Yunnan and
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, Northeast
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(
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), the females of
<taxonomicName id="4C4B17D0FFD1FFADFBF7F8D6E457F8AC" authorityName="H. Huang" authorityYear="2021" box="[1052,1336,1859,1885]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="396" phylum="Arthropoda" rank="subSpecies" species="thawgawa" subSpecies="wuzhenjuni">
<emphasis id="B93FB041FFD1FFADFBF7F8D6E457F8AC" box="[1052,1336,1859,1885]" italics="true" pageId="10" pageNumber="396">E. thawgawa wuzhenjuni</emphasis>
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differ in the absence of the white spots in spaces 6 and 7 on hindwing upperside and the medial band on forewing upperside slightly broader near the costa.
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<paragraph id="8BF46C53FFD0FFACFF7CF805E58FF822" blockId="11.[151,1436,1943,2003]" pageId="11" pageNumber="397">
<emphasis id="B93FB041FFD0FFACFF7CF805E03BF85E" bold="true" box="[151,340,1943,1967]" pageId="11" pageNumber="397">FIGURES 60–65.</emphasis>
Male genitalia of
<taxonomicName id="4C4B17D0FFD0FFACFDE6F805E3F1F85E" authorityName="Staudinger" authorityYear="1886" box="[525,670,1943,1967]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="397" phylum="Arthropoda" rank="species" species="duda">
<emphasis id="B93FB041FFD0FFACFDE6F805E3F1F85E" box="[525,670,1943,1967]" italics="true" pageId="11" pageNumber="397">Euthalia duda</emphasis>
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sspp. Depositary of the specimens: 60 in TAAHU, 61 in CLSY, 62 in MfN, 63 &amp; 65 in ZFMK, 64 in NMB. a=genitalia capsule, b=phallus in lateral view, c=tip of valva in its full view.
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<emphasis id="B93FB041FFD7FFABFF7CF957E039F92C" bold="true" box="[151,342,1733,1757]" pageId="12" pageNumber="398">FIGURES 66–68.</emphasis>
Distribution map of
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spp.
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in the current study, living adult and habitat. 66, distribution map, 67, living female of
<taxonomicName id="4C4B17D0FFD7FFABFED4F978E08FF8F0" authorityName="Yokochi" authorityYear="2009" box="[319,480,1769,1793]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="398" phylum="Arthropoda" rank="subSpecies" species="bellula" subSpecies="uedai">
<emphasis id="B93FB041FFD7FFABFED4F978E08FF8F0" box="[319,480,1769,1793]" italics="true" pageId="12" pageNumber="398">E. bellula uedai</emphasis>
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<emphasis id="B93FB041FFD7FFABFE0CF97BE32DF8F0" bold="true" box="[487,578,1769,1793]" pageId="12" pageNumber="398">stat. rev.</emphasis>
from Gongshan County, 68, habitat in Gongshan County, Northwest Yunnan.
</paragraph>
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<emphasis id="B93FB041FFD7FFABFF2CF8C1E00EF89C" bold="true" box="[199,353,1875,1901]" pageId="12" pageNumber="398">Distribution.</emphasis>
Southwest
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<emphasis id="B93FB041FFD7FFABFDC7F8C1E35AF89C" bold="true" box="[556,565,1875,1901]" pageId="12" pageNumber="398">(</emphasis>
Along Nujiang valley from Northwest
<collectingRegion id="498FA2B1FFD7FFABFC0EF8C6E554F89F" box="[997,1083,1876,1902]" country="China" name="Yunnan" pageId="12" pageNumber="398">Yunnan</collectingRegion>
(Gongshan County) to extreme
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<collectingMunicipality id="6B90F629FFD7FFABFF7CF8E5E06BF860" box="[151,260,1911,1937]" pageId="12" pageNumber="398">Southeast</collectingMunicipality>
<collectingRegion id="498FA2B1FFD7FFABFEE0F8E5E033F860" box="[267,348,1911,1937]" country="China" name="Xizang" pageId="12" pageNumber="398">Xizang</collectingRegion>
(
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,
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.
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<emphasis id="B93FB041FFD6FFAAFF7CFF05E018FF40" bold="true" box="[151,375,151,178]" italics="true" pageId="13" pageNumber="399">Euthalia mingyiae</emphasis>
H.
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<emphasis id="B93FB041FFD6FFAAFF7CFE92E03DFEE6" box="[151,338,256,280]" italics="true" pageId="13" pageNumber="399">Euthalia mingyiae</emphasis>
H.
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, fig. 10, pl. XV, figs. 6 &amp; 8;
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, figs. 68, 280.
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<emphasis id="B93FB041FFD6FFAAFF7CFED7E0EFFEAE" bold="true" box="[151,384,325,351]" pageId="13" pageNumber="399">Material examined.</emphasis>
<materialsCitation id="3B23660EFFD6FFAAFE6DFED4E19AFE75" collectingDate="2021-08-01" collectionCode="CHSY" collectorName="Si-yao Huang" county="Gongshan County" location="Bingzhongluo" pageId="13" pageNumber="399" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D4DA7DAFFD6FFAAFE6DFED4E086FE91" box="[390,489,326,352]" count="1" pageId="13" pageNumber="399" type="female">1 female</specimenCount>
,
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<collectingDate id="EFB1B37BFFD6FFAAFE1FFED4E315FE91" box="[500,634,326,352]" pageId="13" pageNumber="399" value="2021-08-01">1.VIII.2021</collectingDate>
</date>
,
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,
<collectingCounty id="629514DFFFD6FFAAFCDAFED4E292FE91" box="[817,1021,326,352]" pageId="13" pageNumber="399">Gongshan County</collectingCounty>
,
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, leg.
<collectorName id="26BE0985FFD6FFAAFAE9FED4E4F2FE91" box="[1282,1437,326,352]" pageId="13" pageNumber="399">Si-yao Huang</collectorName>
(
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<emphasis id="B93FB041FFD6FFAAFF2CFE1CE054FE59" bold="true" box="[199,315,398,424]" pageId="13" pageNumber="399">Remarks.</emphasis>
This is the first record of the female of this little-known species which is endemic to the area around Bingzhongluo, Gongshan County. Compared with the male, the size of the female is much larger (length of forewing
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vs
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in HT male), but the wing pattern and shape do not show significant difference.
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<emphasis id="B93FB041FFD6FFAAFF2CFE6BE00EFDE2" bold="true" box="[199,353,505,531]" pageId="13" pageNumber="399">Distribution.</emphasis>
Southwest
<collectingCountry id="F35C2CC3FFD6FFAAFE08FE68E349FDE5" box="[483,550,506,532]" name="China" pageId="13" pageNumber="399">China</collectingCountry>
(West
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, Gongshan County, area around Bingzhongluo).
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<mods:title id="2C35EAAB3F4841D7A8CDF965945E7013">Taxonomic notes on certain species in the subgenus Limbusa Moore of the genus Euthalia Hübner (Lepidoptera, Papilionoidea, Nymphalidae, Limenitinae) from China and adjacent countries</mods:title>
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<taxonomicName id="4C4B17D0FFD9FFA5FF7CFF4DE334FF08" authority="Yokochi, 2009" authorityName="Yokochi" authorityYear="2009" box="[151,603,223,250]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="subSpecies" species="bellula" status="stat. nov." subSpecies="uedai">
<emphasis id="B93FB041FFD9FFA5FF7CFF4DE0C9FF0B" bold="true" box="[151,422,223,250]" italics="true" pageId="2" pageNumber="388">Euthalia bellula uedai</emphasis>
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<figureCitation id="137070D6FFD9FFA5FF7CFE91E06EFEEF" box="[151,257,259,286]" captionStart="FIGURES 1–7" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[190,1398,181,1919]" captionTargetId="figure-28@3.[190,1398,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–7. Adults of Euthalia spp. Depositary of the specimens: 1 in KMNH, 2–5 in CHSY, 6 &amp; 7 in NHMUK (©The Trustees of NHMUK). a=upperside, b=underside, c=labels." pageId="2" pageNumber="388">Figs. 1–5</figureCitation>
,
<figureCitation id="137070D6FFD9FFA5FEE6FE91E039FEEF" box="[269,342,259,286]" captionStart="FIGURES 50–55" captionStartId="9.[151,264,1954,1978]" captionTargetBox="[170,1407,355,1930]" captionTargetId="figure-115@9.[164,1423,345,1930]" captionTargetPageId="9" captionText="FIGURES 50–55. Male genitalia of Euthalia spp. Depositary of the specimens: 50–52, 54 &amp; 55 in CHSY, 53 in CHH.a=genitalia capsule, b=phallus in lateral view, c=tip of valva in its full view." pageId="2" pageNumber="388">50–52</figureCitation>
,
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<emphasis id="B93FB041FFD9FFA5FF7CFEDAE181FE91" box="[151,238,328,352]" italics="true" pageId="2" pageNumber="388">Euthalia</emphasis>
(
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, figs. 1–4, 8.
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<emphasis id="B93FB041FFD9FFA5FF7CFE1FE0EBFE56" bold="true" box="[151,388,397,423]" pageId="2" pageNumber="388">Material examined.</emphasis>
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of
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(
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)
<emphasis id="B93FB041FFD9FFA5FCE6FE1CE2CDFE59" box="[781,930,398,424]" italics="true" pageId="2" pageNumber="388">bellula uedai</emphasis>
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: male, printed label “
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<collectingDate id="EFB1B37BFFD9FFA5FB4AFE1CE457FE59" box="[1185,1336,398,424]" pageId="2" pageNumber="388" value="2006-07-20">20. Jul. 2006</collectingDate>
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| Chudu Razi | (Mt. Range) |
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west of | Kawanglangpu |
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| MYANMAR | Coll.
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. Yokochi.” / printed red label “
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” (
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:
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<collectingDate id="EFB1B37BFFD9FFA5FC30FE44E50EFE01" box="[987,1121,470,496]" pageId="2" pageNumber="388" value="2021-08-06">6.VIII.2021</collectingDate>
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,
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,
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, leg.
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, genitalia dissection No.
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(
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, same data, but
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<collectingDate id="EFB1B37BFFD9FFA5FAFAFE68E4F7FDE5" box="[1297,1432,506,532]" pageId="2" pageNumber="388" value="2021-08-04">4.VIII.2021</collectingDate>
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, genitalia dissection No.
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(male) (
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, same data, but
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, genitalia dissection No.
<specimenCode id="DBEDC428FFD9FFA5FAA1FD8CE4F3FDC9" box="[1354,1436,542,568]" collectionCode="NJEU" pageId="2" pageNumber="388">NJEU3</specimenCode>
(
<collectionCode id="ED5AF496FFD9FFA5FF74FDD0E19FFDAD" box="[159,240,578,604]" pageId="2" pageNumber="388">CHSY</collectionCode>
);
<specimenCount id="9D4DA7DAFFD9FFA5FEE8FDD0E008FDAD" box="[259,359,578,604]" count="1" pageId="2" pageNumber="388" type="female">1 female</specimenCount>
,
<date id="FFF54A93FFD9FFA5FE99FDD0E096FDAD" box="[370,505,578,604]" pageId="2" pageNumber="388" value="2021-08-02">
<collectingDate id="EFB1B37BFFD9FFA5FE99FDD0E096FDAD" box="[370,505,578,604]" pageId="2" pageNumber="388" value="2021-08-02">2.VIII.2021</collectingDate>
</date>
, Bingzhongluo,
<location id="8E943A88FFD9FFA5FD5AFDD0E211FDAD" LSID="urn:lsid:plazi:treatment:03E2DD45FFD9FFA5FF7CFF4DE378FA69:8E943A88FFD9FFA5FD5AFDD0E211FDAD" box="[689,894,578,604]" country="China" county="Gongshan County" name="Gongshan County" pageId="2" pageNumber="388" stateProvince="Yunnan">Gongshan County</location>
,
<collectingRegion id="498FA2B1FFD9FFA5FC6CFDD0E525FDAD" box="[903,1098,578,604]" country="China" name="Yunnan" pageId="2" pageNumber="388">Yunnan Province</collectingRegion>
, leg.
<collectorName id="26BE0985FFD9FFA5FB6DFDD0E44EFDAD" box="[1158,1313,578,604]" pageId="2" pageNumber="388">Si-yao Huang</collectorName>
(
<collectionCode id="ED5AF496FFD9FFA5FADBFDD0E4EEFDAD" box="[1328,1409,578,604]" pageId="2" pageNumber="388">CHSY</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3513FD8FFD9FFA5FF2CFDF7E423FAF9" pageId="2" pageNumber="388" type="discussion">
<paragraph id="8BF46C53FFD9FFA5FF2CFDF7E239FC19" blockId="2.[151,1437,397,1432]" pageId="2" pageNumber="388">
<emphasis id="B93FB041FFD9FFA5FF2CFDF7E023FD8E" bold="true" box="[199,332,613,639]" pageId="2" pageNumber="388">Discussion.</emphasis>
Taxon
<emphasis id="B93FB041FFD9FFA5FE4EFDF4E08CFD71" box="[421,483,614,640]" italics="true" pageId="2" pageNumber="388">uedai</emphasis>
was originally described as a subspecies of
<taxonomicName id="4C4B17D0FFD9FFA5FC0FFDF5E46DFD70" authority="Yokochi, 2005" authorityName="Yokochi" authorityYear="2005" box="[996,1282,614,641]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B93FB041FFD9FFA5FC0FFDF5E53BFD71" box="[996,1108,614,640]" italics="true" pageId="2" pageNumber="388">E. bellula</emphasis>
<bibRefCitation id="EFDA11A2FFD9FFA5FBB7FDF4E46DFD70" author="Yokochi, T." box="[1116,1282,614,641]" pageId="2" pageNumber="388" pagination="6 - 18" refId="ref5054" refString="Yokochi, T. (2005) Description of a new species and six new subspecies of the genus Euthalia (subgenus Limbusa), with designation of lectotype of Euthalia pratti occidentalis (Lepidoptera, Nymphalidae). Transaction of the Lepidopterological Society of Japan, 56 (1), 6-18." type="journal article" year="2005">Yokochi, 2005</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="498FA2B1FFD9FFA5FAA0FDF4E1BDFD55" country="Myanmar" name="Kachin" pageId="2" pageNumber="388">Kachin State</collectingRegion>
, NE
<collectingCountry id="F35C2CC3FFD9FFA5FEE4FD18E013FD55" box="[271,380,650,676]" name="Myanmar" pageId="2" pageNumber="388">Myanmar</collectingCountry>
(
<bibRefCitation id="EFDA11A2FFD9FFA5FE66FD18E35CFD55" author="Yokochi, T." box="[397,563,650,676]" pageId="2" pageNumber="388" pagination="20 - 28" refId="ref5110" refString="Yokochi, T. (2009) Two new subspecies of the Genus Euthalia (Subgenus Limbusa) recorded from Kachin, Myanmar (Lepidoptera, Nymphalidae). Butterflies, 53, 20-28." type="journal article" year="2009">Yokochi 2009</bibRefCitation>
). Later,
<bibRefCitation id="EFDA11A2FFD9FFA5FD79FD18E22BFD55" author="Yokochi, T." box="[658,836,650,676]" pageId="2" pageNumber="388" pagination="9 - 100" refId="ref5259" refString="Yokochi, T. (2012) Revision of the Subgenus Limbusa Moore, [1897] (Lepidoptera, Nymphalidae, Adoliadini) Part 3. Descriptions of species (2). Bulletin of the Kitakyushu Museum of Natural History and Human History, Series A, 10, 9-100." type="journal article" year="2012">Yokochi (2012)</bibRefCitation>
synonymized this taxon with
<taxonomicName id="4C4B17D0FFD9FFA5FB49FD19E444FD55" authorityName="Oberthur" authorityYear="1907" box="[1186,1323,650,676]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="monbeigi">
<emphasis id="B93FB041FFD9FFA5FB49FD19E444FD55" box="[1186,1323,650,676]" italics="true" pageId="2" pageNumber="388">E. monbeigi</emphasis>
</taxonomicName>
. In 2021, a series of specimens of both sexes of this taxon were collected by the first author in his field survey in Nujiang valley, and the exmination of the wing pattern as well as the male genitalia reveals that
<emphasis id="B93FB041FFD9FFA5FB7CFD40E5BAFD1D" box="[1175,1237,722,748]" italics="true" pageId="2" pageNumber="388">uedai</emphasis>
is in fact a bona taxon, differing from its related congeners, e.g.
<taxonomicName id="4C4B17D0FFD9FFA5FD40FD65E277FCE1" authorityName="Yokochi" authorityYear="2005" box="[683,792,758,784]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B93FB041FFD9FFA5FD40FD65E277FCE1" box="[683,792,758,784]" italics="true" pageId="2" pageNumber="388">E. bellula</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4B17D0FFD9FFA5FCCDFD65E2C2FCE1" authorityName="Oberthur" authorityYear="1907" box="[806,941,758,784]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="monbeigi">
<emphasis id="B93FB041FFD9FFA5FCCDFD65E2C2FCE1" box="[806,941,758,784]" italics="true" pageId="2" pageNumber="388">E. monbeigi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4B17D0FFD9FFA5FC08FD65E518FCE1" authorityName="Koiwaya" authorityYear="1996" box="[995,1143,759,784]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="yunnanica">
<emphasis id="B93FB041FFD9FFA5FC08FD65E518FCE1" box="[995,1143,759,784]" italics="true" pageId="2" pageNumber="388">E. yunnanica</emphasis>
</taxonomicName>
by the combination of the following characters: 1) the patches in spaces
<quantity id="4CB3C1B6FFD9FFA5FD49FC88E385FCC5" box="[674,746,794,820]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="2" pageNumber="388" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
the medial band on forewing upperside are distinctly longer, especially in male; 2) the patch in space
<quantity id="4CB3C1B6FFD9FFA5FD8BFCACE3E4FCA9" box="[608,651,830,856]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="388" unit="in" value="6.0">6 in</quantity>
the medial band on forewing upperside is more oval-shaped, while in its congeners such patch is more rectangular; 3) the discal band on hindwing is much broader than
<taxonomicName id="4C4B17D0FFD9FFA5FB15FCF1E403FC8D" authorityName="Yokochi" authorityYear="2005" box="[1278,1388,866,892]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B93FB041FFD9FFA5FB15FCF1E403FC8D" box="[1278,1388,866,892]" italics="true" pageId="2" pageNumber="388">E. bellula</emphasis>
</taxonomicName>
and 4) in male genitalia, the valva is distinctly longer (the male genitalia of the true
<taxonomicName id="4C4B17D0FFD9FFA5FBF3FC15E5EAFC51" authorityName="Yokochi" authorityYear="2005" box="[1048,1157,902,928]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B93FB041FFD9FFA5FBF3FC15E5EAFC51" box="[1048,1157,902,928]" italics="true" pageId="2" pageNumber="388">E. bellula</emphasis>
</taxonomicName>
is unknown, see below). Since the male genitalia of the true
<taxonomicName id="4C4B17D0FFD9FFA5FDC7FC39E3F5FC35" authorityName="Yokochi" authorityYear="2005" box="[556,666,938,964]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B93FB041FFD9FFA5FDC7FC39E3F5FC35" box="[556,666,938,964]" italics="true" pageId="2" pageNumber="388">E. bellula</emphasis>
</taxonomicName>
from North
<collectingCountry id="F35C2CC3FFD9FFA5FCC1FC38E20FFC35" box="[810,864,938,964]" name="Laos" pageId="2" pageNumber="388">Laos</collectingCountry>
is unknown, we temporarily still treated the taxon
<emphasis id="B93FB041FFD9FFA5FF7CFC5CE1BAFC19" box="[151,213,974,1000]" italics="true" pageId="2" pageNumber="388">uedai</emphasis>
as a valid subspecies of it, viz.
<taxonomicName id="4C4B17D0FFD9FFA5FDD3FC5DE385FC19" authorityName="Yokochi" authorityYear="2009" box="[568,746,974,1000]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="subSpecies" species="bellula" subSpecies="uedai">
<emphasis id="B93FB041FFD9FFA5FDD3FC5DE385FC19" box="[568,746,974,1000]" italics="true" pageId="2" pageNumber="388">E. bellula uedai</emphasis>
</taxonomicName>
<emphasis id="B93FB041FFD9FFA5FD1AFC5CE239FC19" bold="true" box="[753,854,974,1000]" pageId="2" pageNumber="388">stat. rev.</emphasis>
</paragraph>
<paragraph id="8BF46C53FFD9FFA5FF2CFC60E423FAF9" blockId="2.[151,1437,397,1432]" pageId="2" pageNumber="388">
<emphasis id="B93FB041FFD9FFA5FF2CFC60E054FBFD" bold="true" box="[199,315,1010,1036]" pageId="2" pageNumber="388">Remarks.</emphasis>
It is worth noting that the status of
<taxonomicName id="4C4B17D0FFD9FFA5FD30FC61E224FBFD" authorityName="Yokochi" authorityYear="2005" box="[731,843,1010,1036]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B93FB041FFD9FFA5FD30FC61E224FBFD" box="[731,843,1010,1036]" italics="true" pageId="2" pageNumber="388">E. bellula</emphasis>
</taxonomicName>
is unclear. This taxon was originally described as a subspecies of
<taxonomicName id="4C4B17D0FFD9FFA5FEDCFB85E0FFFBC1" authorityName="Staudinger" authorityYear="1886" box="[311,400,1046,1072]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="duda">
<emphasis id="B93FB041FFD9FFA5FEDCFB85E0FFFBC1" box="[311,400,1046,1072]" italics="true" pageId="2" pageNumber="388">E. duda</emphasis>
</taxonomicName>
based upon a series of specimens from
<collectingCountry id="F35C2CC3FFD9FFA5FCB2FB84E2E0FBC1" box="[857,911,1046,1072]" name="Laos" pageId="2" pageNumber="388">Laos</collectingCountry>
and
<collectingCountry id="F35C2CC3FFD9FFA5FC23FB84E545FBC1" box="[968,1066,1046,1072]" name="Vietnam" pageId="2" pageNumber="388">Vietnam</collectingCountry>
, and later raised to specific rank by
<bibRefCitation id="EFDA11A2FFD9FFA5FF51FBA8E001FBA5" author="Yokochi, T." box="[186,366,1082,1108]" pageId="2" pageNumber="388" pagination="20 - 28" refId="ref5110" refString="Yokochi, T. (2009) Two new subspecies of the Genus Euthalia (Subgenus Limbusa) recorded from Kachin, Myanmar (Lepidoptera, Nymphalidae). Butterflies, 53, 20-28." type="journal article" year="2009">Yokochi (2009)</bibRefCitation>
. The
<typeStatus id="54F0D2F1FFD9FFA5FE46FBA8E0B2FBA5" box="[429,477,1082,1108]" pageId="2" pageNumber="388">type</typeStatus>
locality of
<taxonomicName id="4C4B17D0FFD9FFA5FD8AFBA9E3BFFBA5" authorityName="Yokochi" authorityYear="2005" box="[609,720,1082,1108]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B93FB041FFD9FFA5FD8AFBA9E3BFFBA5" box="[609,720,1082,1108]" italics="true" pageId="2" pageNumber="388">E. bellula</emphasis>
</taxonomicName>
is Xam Neua, North
<collectingCountry id="F35C2CC3FFD9FFA5FC2EFBA8E291FBA5" box="[965,1022,1082,1108]" name="Laos" pageId="2" pageNumber="388">Laos</collectingCountry>
, but the male genitalia illustrated by
<bibRefCitation id="EFDA11A2FFD9FFA5FF7CFBCCE02EFB89" author="Yokochi, T." box="[151,321,1118,1144]" pageId="2" pageNumber="388" pagination="20 - 28" refId="ref5110" refString="Yokochi, T. (2009) Two new subspecies of the Genus Euthalia (Subgenus Limbusa) recorded from Kachin, Myanmar (Lepidoptera, Nymphalidae). Butterflies, 53, 20-28." type="journal article" year="2009">Yokochi (2009</bibRefCitation>
,
<bibRefCitation id="EFDA11A2FFD9FFA5FEA6FBCCE0E8FB89" author="Yokochi, T." box="[333,391,1118,1144]" pageId="2" pageNumber="388" pagination="9 - 100" refId="ref5259" refString="Yokochi, T. (2012) Revision of the Subgenus Limbusa Moore, [1897] (Lepidoptera, Nymphalidae, Adoliadini) Part 3. Descriptions of species (2). Bulletin of the Kitakyushu Museum of Natural History and Human History, Series A, 10, 9-100." type="journal article" year="2012">2012</bibRefCitation>
) were from Vietnamese population, while those of the Laotian population are still unknown. Superficially, the Vietnamese population differs from Laotian population by the broader hindwing discal band, suggesting the possiblity that Vietnamese population represents a different species. For exactly the same reason, it is possible that the taxon
<emphasis id="B93FB041FFD9FFA5FE79FB58E0BFFB15" box="[402,464,1226,1252]" italics="true" pageId="2" pageNumber="388">uedai</emphasis>
, which also bears a broader hindwing discal band, should be treated as a distinct species. To clarify both issues, the examination of the male genitalia of
<taxonomicName id="4C4B17D0FFD9FFA5FCB1FB7DE2A8FAF9" authorityName="Yokochi" authorityYear="2005" box="[858,967,1262,1288]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="B93FB041FFD9FFA5FCB1FB7DE2A8FAF9" box="[858,967,1262,1288]" italics="true" pageId="2" pageNumber="388">E. bellula</emphasis>
</taxonomicName>
from its
<typeStatus id="54F0D2F1FFD9FFA5FBC6FB7CE532FAF9" box="[1069,1117,1262,1288]" pageId="2" pageNumber="388">type</typeStatus>
locality is necessary.
</paragraph>
</subSubSection>
<subSubSection id="C3513FD8FFD9FFA5FF2CFA83E28AFAA1" pageId="2" pageNumber="388" type="biology_ecology">
<paragraph id="8BF46C53FFD9FFA5FF2CFA83E28AFAA1" blockId="2.[151,1437,397,1432]" pageId="2" pageNumber="388">
<emphasis id="B93FB041FFD9FFA5FF2CFA83E048FADA" bold="true" box="[199,295,1297,1323]" pageId="2" pageNumber="388">Biology.</emphasis>
In
<collectingCountry id="F35C2CC3FFD9FFA5FEA7FA80E0FDFADD" box="[332,402,1298,1324]" name="China" pageId="2" pageNumber="388">China</collectingCountry>
,
<taxonomicName id="4C4B17D0FFD9FFA5FE76FA80E3F8FADD" authorityName="Yokochi" authorityYear="2009" box="[413,663,1298,1324]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="388" phylum="Arthropoda" rank="subSpecies" species="bellula" subSpecies="uedai">
<emphasis id="B93FB041FFD9FFA5FE76FA80E3F8FADD" box="[413,663,1298,1324]" italics="true" pageId="2" pageNumber="388">Euthalia bellula uedai</emphasis>
</taxonomicName>
is observed on wing from July to August, and can be found puddling on wet soil (
<figureCitation id="137070D6FFD9FFA5FECAFAA4E01CFAA1" box="[289,371,1334,1360]" captionStart="FIGURES 66–68" captionStartId="12.[151,264,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-18@12.[151,1436,181,1709]" captionTargetPageId="12" captionText="FIGURES 66–68. Distribution map of Euthalia spp. in the current study, living adult and habitat. 66, distribution map, 67, living female of E. bellula uedai stat. rev. from Gongshan County, 68, habitat in Gongshan County, Northwest Yunnan." pageId="2" pageNumber="388">Fig. 67</figureCitation>
) in dense forest at the altitude around
<quantity id="4CB3C1B6FFD9FFA5FCCAFAA4E219FAA1" box="[801,886,1334,1360]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="388" unit="m" value="2700.0">2700 m</quantity>
(
<figureCitation id="137070D6FFD9FFA5FC6EFAA4E2B6FAA1" box="[901,985,1334,1360]" captionStart="FIGURES 66–68" captionStartId="12.[151,264,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-18@12.[151,1436,181,1709]" captionTargetPageId="12" captionText="FIGURES 66–68. Distribution map of Euthalia spp. in the current study, living adult and habitat. 66, distribution map, 67, living female of E. bellula uedai stat. rev. from Gongshan County, 68, habitat in Gongshan County, Northwest Yunnan." pageId="2" pageNumber="388">Fig. 68</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3513FD8FFD9FFA5FF2CFACBE378FA69" pageId="2" pageNumber="388" type="distribution">
<paragraph id="8BF46C53FFD9FFA5FF2CFACBE378FA69" blockId="2.[151,1437,397,1432]" pageId="2" pageNumber="388">
<emphasis id="B93FB041FFD9FFA5FF2CFACBE00EFA82" bold="true" box="[199,353,1369,1395]" pageId="2" pageNumber="388">Distribution.</emphasis>
Southwest
<collectingCountry id="F35C2CC3FFD9FFA5FE09FAC8E34AFA85" box="[482,549,1370,1396]" name="China" pageId="2" pageNumber="388">China</collectingCountry>
(West
<collectingRegion id="498FA2B1FFD9FFA5FD9AFAC8E3A5FA85" box="[625,714,1370,1396]" country="China" name="Yunnan" pageId="2" pageNumber="388">Yunnan</collectingRegion>
, along Nujiang valley from Gongshan to Fugong), and Northeast
<collectingCountry id="F35C2CC3FFD9FFA5FF7CFAECE06BFA69" box="[151,260,1406,1432]" name="Myanmar" pageId="2" pageNumber="388">Myanmar</collectingCountry>
(
<collectingRegion id="498FA2B1FFD9FFA5FEF8FAECE0C9FA69" box="[275,422,1406,1432]" country="Myanmar" name="Kachin" pageId="2" pageNumber="388">Kachin State</collectingRegion>
) (
<figureCitation id="137070D6FFD9FFA5FE56FAECE361FA69" box="[445,526,1406,1432]" captionStart="FIGURES 66–68" captionStartId="12.[151,264,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-18@12.[151,1436,181,1709]" captionTargetPageId="12" captionText="FIGURES 66–68. Distribution map of Euthalia spp. in the current study, living adult and habitat. 66, distribution map, 67, living female of E. bellula uedai stat. rev. from Gongshan County, 68, habitat in Gongshan County, Northwest Yunnan." pageId="2" pageNumber="388">Fig. 66</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="435D76CCFFE509E0D32E90E4DCB1E861">Taxonomic notes on certain species in the subgenus Limbusa Moore of the genus Euthalia Hübner (Lepidoptera, Papilionoidea, Nymphalidae, Limenitinae) from China and adjacent countries</mods:title>
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<mods:namePart id="070A3AD7FFD304FC162D8C74F2899A9C">Huang, Si-Yao</mods:namePart>
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<mods:affiliation id="86B57FE050EEA2AD53BEA463C4BE4A57">Leibniz Institute for the Analysis of Biodiversity Change, Museum Koenig, Adenauer Allee 127, Bonn 53113, Germany.</mods:affiliation>
<mods:nameIdentifier id="734121DA230BA134A19F1D50BA0471A9" type="email">uang@leibniz-lib.de</mods:nameIdentifier>
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<mods:namePart id="B497F10AB5BEC0D4122B37B0A3595EC8">Xu, Yong-Qiang</mods:namePart>
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<mods:affiliation id="80A5FD962617E12CCF31045510DEDF57">Institute of Plateau Biology of Xizang Aotonomous Region, Lhasa, Xizang, China.</mods:affiliation>
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<mods:namePart id="DD0A159152A4733363BB9224A4382AB7">Lu, Di</mods:namePart>
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<mods:namePart id="711DCB7C37B718331B4C915BB229848C">Lu, Fei</mods:namePart>
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<mods:affiliation id="21E5F5BD4FF74EB268D458A5264421E4">College of Forestry, Southwest Forestry University, Kunming 650224, Yunnan, China.</mods:affiliation>
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<taxonomicName id="4C4B17D0FFDCFFA0FF7CF948E311F905" authority="H. Huang, 1999" authorityName="H. Huang" authorityYear="1999" box="[151,638,1754,1781]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="393" phylum="Arthropoda" rank="subSpecies" species="duda" status="stat. nov." subSpecies="tsangpoi">
<emphasis id="B93FB041FFDCFFA0FF7CF948E0DCF905" bold="true" box="[151,435,1754,1781]" italics="true" pageId="7" pageNumber="393">Euthalia duda tsangpoi</emphasis>
H.
<bibRefCitation id="EFDA11A2FFDCFFA0FE0BF949E311F905" author="Huang, H." box="[480,638,1754,1781]" pageId="7" pageNumber="393" pagination="642 - 676" refId="ref4575" refString="Huang, H. (1999) Some new butterflies from China. I. (Rhopalocera). Lambillionea, XCIX (4), 642-676." type="journal article" year="1999">Huang, 1999</bibRefCitation>
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<taxonomicNameLabel id="A20C0D3AFFDCFFA0FD6EF949E380F904" box="[645,751,1755,1781]" pageId="7" pageNumber="393">stat. rev.</taxonomicNameLabel>
</emphasis>
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<figureCitation id="137070D6FFDCFFA0FF7CF96DE071F8E8" box="[151,286,1791,1817]" captionStart="FIGURES 15–29" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-28@5.[203,1384,181,1919]" captionTargetPageId="5" captionText="FIGURES 15–29. Adults of Euthalia duda sspp., upperside. Depositary of the specimens: 15 &amp; 16 in MfN, 17 in NMB, 18–21 in ZFMK, 22 &amp; 23 in NHMUK (©The Trustees of NHMUK), 24 in CQEC, 25 in TAAHU, 26, 28 &amp; 29 in CLSY, 27 in CLF, scale bar for all except fig. 27." pageId="7" pageNumber="393">Figs. 24–29</figureCitation>
,
<figureCitation id="137070D6FFDCFFA0FEC2F96DE01DF8E8" box="[297,370,1791,1817]" captionStart="FIGURES 30–44" captionStartId="6.[151,264,1923,1947]" captionTargetBox="[151,1436,181,1899]" captionTargetId="figure-18@6.[151,1436,181,1899]" captionTargetPageId="6" captionText="FIGURES 30–44. Underside of figures 15–29, scale bar for all except fig. 27." pageId="7" pageNumber="393">39–44</figureCitation>
,
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h-k, 56, 57, 60, 61
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<treatmentCitation id="0AEA4A42FFDCFFA0FF7CF8D1E345F8AA" author="Huang, H." box="[151,554,1859,1884]" page="643" pageId="7" pageNumber="393" year="1999">
<taxonomicName id="4C4B17D0FFDCFFA0FF7CF8D1E345F8AA" authority="H. Huang, 1999: 643" authorityName="H. Huang" authorityPageNumber="643" authorityYear="1999" box="[151,554,1859,1884]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="393" phylum="Arthropoda" rank="species" species="tsangpoi">
<emphasis id="B93FB041FFDCFFA0FF7CF8D1E023F8AA" box="[151,332,1859,1883]" italics="true" pageId="7" pageNumber="393">Euthalia tsangpoi</emphasis>
H.
<bibRefCitation id="EFDA11A2FFDCFFA0FE9AF8D6E345F8AA" author="Huang, H." box="[369,554,1859,1884]" pageId="7" pageNumber="393" pagination="642 - 676" refId="ref4575" refString="Huang, H. (1999) Some new butterflies from China. I. (Rhopalocera). Lambillionea, XCIX (4), 642-676." type="journal article" year="1999">Huang, 1999: 643</bibRefCitation>
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, figs. 6, 7, 79;
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<bibRefCitation id="EFDA11A2FFDCFFA0FD54F8D1E21AF8AA" author="Yokochi, T." box="[703,885,1859,1883]" pageId="7" pageNumber="393" pagination="9 - 100" refId="ref5259" refString="Yokochi, T. (2012) Revision of the Subgenus Limbusa Moore, [1897] (Lepidoptera, Nymphalidae, Adoliadini) Part 3. Descriptions of species (2). Bulletin of the Kitakyushu Museum of Natural History and Human History, Series A, 10, 9-100." type="journal article" year="2012">Yokochi 2012: 12</bibRefCitation>
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, fig. 103.
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<emphasis id="B93FB041FFDCFFA0FF7CF81BE0EBF852" bold="true" box="[151,388,1929,1955]" pageId="7" pageNumber="393">Material examined.</emphasis>
<materialsCitation id="3B23660EFFDCFFA0FE65F81BE197F836" collectingDate="1996-07" collectionCode="CQEC" collectorName="Motuo, SE" location="Hanmi" pageId="7" pageNumber="393" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="B93FB041FFDCFFA0FE65F81BE094F852" bold="true" box="[398,507,1929,1955]" pageId="7" pageNumber="393">
<typeStatus id="54F0D2F1FFDCFFA0FE65F81BE094F852" box="[398,507,1929,1955]" pageId="7" pageNumber="393" type="holotype">Holotype</typeStatus>
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: male, late
<date id="FFF54A93FFDCFFA0FD6CF81BE395F852" box="[647,762,1929,1955]" pageId="7" pageNumber="393" value="1996-07">
<collectingDate id="EFB1B37BFFDCFFA0FD6CF81BE395F852" box="[647,762,1929,1955]" pageId="7" pageNumber="393" value="1996-07">July 1996</collectingDate>
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, on road between
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and Aniqiao,
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(
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)
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.
<emphasis id="B93FB041FFDCFFA0FEEDF83FE31DF836" bold="true" box="[262,626,1965,1991]" pageId="7" pageNumber="393">Additional material examined</emphasis>
:
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, altitude
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,
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,
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,
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,
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<collectingMunicipality id="6B90F629FFDCFFA0FF19F843E08CF81A" box="[242,483,2001,2027]" pageId="7" pageNumber="393">Autonomous Region</collectingMunicipality>
,
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, leg.
<collectorName id="26BE0985FFDCFFA0FD95F843E25CF81D" box="[638,819,2001,2028]" pageId="7" pageNumber="393">Song-yun Lang</collectorName>
, dissection
<collectorName id="26BE0985FFDCFFA0FC54F840E286F81D" box="[959,1001,2002,2028]" pageId="7" pageNumber="393">No.</collectorName>
NYM0017 &amp; NYM0060 (
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</paragraph>
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female,
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<date id="FFF54A93FFD3FFAFFF19FF05E0F9FF40" box="[242,406,151,177]" pageId="8" pageNumber="394" value="2012-08-14">
<collectingDate id="EFB1B37BFFD3FFAFFF19FF05E0F9FF40" box="[242,406,151,177]" pageId="8" pageNumber="394" value="2012-08-14">14. VIII. 2012</collectingDate>
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,
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,
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,
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<collectingMunicipality id="6B90F629FFD3FFAFFCE1FF05E299FF40" box="[778,1014,151,177]" pageId="8" pageNumber="394">Autonomous Region</collectingMunicipality>
,
<collectingCountry id="F35C2CC3FFD3FFAFFBE9FF05E527FF40" box="[1026,1096,151,177]" name="China" pageId="8" pageNumber="394">China</collectingCountry>
, leg.
<collectorName id="26BE0985FFD3FFAFFB6FFF05E45CFF40" box="[1156,1331,151,177]" pageId="8" pageNumber="394">Song-yun Lang</collectorName>
(
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);
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,
<date id="FFF54A93FFD3FFAFFF18FF29E01AFF24" box="[243,373,187,213]" pageId="8" pageNumber="394" value="2016-06-27">
27.
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<materialsCitation id="3B23660EFFD3FFAFFED8FF29E01AFF24" box="[307,373,187,213]" collectionCode="TAAHU" collectorName="Zhao-hui Pan &amp; No." country="China" county="Bomi County" location="Yigong" municipality="Autonomous Region" pageId="8" pageNumber="394" specimenCount="1" stateProvince="Xizang">.2016</materialsCitation>
</date>
,
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,
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,
<collectingRegion id="498FA2B1FFD3FFAFFD6CFF29E3B7FF24" box="[647,728,187,213]" country="China" name="Xizang" pageId="8" pageNumber="394">Xizang</collectingRegion>
<collectingMunicipality id="6B90F629FFD3FFAFFD34FF29E2A4FF24" box="[735,971,187,213]" pageId="8" pageNumber="394">Autonomous Region</collectingMunicipality>
,
<collectingCountry id="F35C2CC3FFD3FFAFFC3CFF29E575FF24" box="[983,1050,187,213]" name="China" pageId="8" pageNumber="394">China</collectingCountry>
leg.
<collectorName id="26BE0985FFD3FFAFFBB8FF29E580FF24" box="[1107,1263,187,213]" pageId="8" pageNumber="394">Zhao-hui Pan</collectorName>
, dissection
<collectorName id="26BE0985FFD3FFAFFA98FF29E4F2FF24" box="[1395,1437,187,213]" pageId="8" pageNumber="394">No.</collectorName>
STS-34888 (
<collectionCode id="ED5AF496FFD3FFAFFECEFF4DE0E3FF08" box="[293,396,223,249]" pageId="8" pageNumber="394">TAAHU</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF343CDBFFD3FFAFFF7CFA5BE0F2F9F4" pageId="8" pageNumber="394" startId="8.[151,264,1481,1505]" targetBox="[151,1435,273,1458]" targetPageId="8" targetType="figure">
<paragraph id="8BF46C53FFD3FFAFFF7CFA5BE0F2F9F4" blockId="8.[151,1437,1481,1541]" pageId="8" pageNumber="394">
<emphasis id="B93FB041FFD3FFAFFF7CFA5BE036FA10" bold="true" box="[151,345,1481,1505]" pageId="8" pageNumber="394">FIGURES 46–49.</emphasis>
Females of
<taxonomicName id="4C4B17D0FFD3FFAFFE09FA5BE356FA10" authorityName="Hubner" authorityYear="1819" box="[482,569,1481,1505]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="B93FB041FFD3FFAFFE09FA5BE356FA10" box="[482,569,1481,1505]" italics="true" pageId="8" pageNumber="394">Euthalia</emphasis>
</taxonomicName>
ssp., Depositary of the specimens: 46 in CHH, 47 &amp; 49 in CHSY, 48 in IPBX. a=upperside, b=underside
</paragraph>
</caption>
<subSubSection id="C3513FD8FFD3FFAEFF2CF9DEE5A8FEEC" lastPageId="9" lastPageNumber="395" pageId="8" pageNumber="394" type="discussion">
<paragraph id="8BF46C53FFD3FFAEFF2CF9DEE5A8FEEC" blockId="8.[151,1437,1612,2035]" lastBlockId="9.[151,1437,151,321]" lastPageId="9" lastPageNumber="395" pageId="8" pageNumber="394">
<emphasis id="B93FB041FFD3FFAFFF2CF9DEE023F997" bold="true" box="[199,332,1612,1638]" pageId="8" pageNumber="394">Discussion.</emphasis>
Taxon
<taxonomicName id="4C4B17D0FFD3FFAFFE76F9DFE092F997" authorityName="H. Huang" authorityYear="1999" box="[413,509,1613,1638]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="394" phylum="Arthropoda" rank="species" species="tsangpoi">
<emphasis id="B93FB041FFD3FFAFFE76F9DFE092F997" box="[413,509,1613,1638]" italics="true" pageId="8" pageNumber="394">tsangpoi</emphasis>
</taxonomicName>
was originally described as a bona species based on
<specimenCount id="9D4DA7DAFFD3FFAFFBD5F9DFE5C0F996" box="[1086,1199,1613,1639]" count="2" pageId="8" pageNumber="394" type="male">two males</specimenCount>
from Metok (Motuo) County, Southeast
<collectingRegion id="498FA2B1FFD3FFAFFE86F9E3E0AEF97A" box="[365,449,1649,1675]" country="China" name="Xizang" pageId="8" pageNumber="394">Xizang</collectingRegion>
,
<collectingCountry id="F35C2CC3FFD3FFAFFE25F9E3E37EF97A" box="[462,529,1649,1675]" name="China" pageId="8" pageNumber="394">China</collectingCountry>
(H.
<bibRefCitation id="EFDA11A2FFD3FFAFFDACF9E3E3B8F97A" author="Huang, H." box="[583,727,1649,1675]" pageId="8" pageNumber="394" pagination="642 - 676" refId="ref4575" refString="Huang, H. (1999) Some new butterflies from China. I. (Rhopalocera). Lambillionea, XCIX (4), 642-676." type="journal article" year="1999">Huang 1999</bibRefCitation>
). Later, H.
<bibRefCitation id="EFDA11A2FFD3FFAFFCB1F9E3E297F97A" author="Huang, H." box="[858,1016,1649,1675]" pageId="8" pageNumber="394" pagination="339 - 360" refId="ref4605" refString="Huang, H. (2002) Some new nymphalids from the valleys of Nujiang and Dulongjiang, China. Atalanta, 33 (3 / 4), 339-360" type="journal article" year="2002">Huang (2002)</bibRefCitation>
synonymized
<taxonomicName id="4C4B17D0FFD3FFAFFB74F9E3E590F97B" authorityName="H. Huang" authorityYear="1999" box="[1183,1279,1649,1674]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="394" phylum="Arthropoda" rank="species" species="tsangpoi">
<emphasis id="B93FB041FFD3FFAFFB74F9E3E590F97B" box="[1183,1279,1649,1674]" italics="true" pageId="8" pageNumber="394">tsangpoi</emphasis>
</taxonomicName>
with
<taxonomicName id="4C4B17D0FFD3FFAFFAA8F9E3E4F3F97A" authorityName="Staudinger" authorityYear="1886" box="[1347,1436,1649,1675]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="394" phylum="Arthropoda" rank="species" species="duda">
<emphasis id="B93FB041FFD3FFAFFAA8F9E3E4F3F97A" box="[1347,1436,1649,1675]" italics="true" pageId="8" pageNumber="394">E. duda</emphasis>
</taxonomicName>
(
<figureCitation id="137070D6FFD3FFAFFF74F907E046F95E" box="[159,297,1684,1711]" captionStart="FIGURES 15–29" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-28@5.[203,1384,181,1919]" captionTargetPageId="5" captionText="FIGURES 15–29. Adults of Euthalia duda sspp., upperside. Depositary of the specimens: 15 &amp; 16 in MfN, 17 in NMB, 18–21 in ZFMK, 22 &amp; 23 in NHMUK (©The Trustees of NHMUK), 24 in CQEC, 25 in TAAHU, 26, 28 &amp; 29 in CLSY, 27 in CLF, scale bar for all except fig. 27." pageId="8" pageNumber="394">Figs. 15–23</figureCitation>
,
<figureCitation id="137070D6FFD3FFAFFED3F907E0EFF95E" box="[312,384,1685,1711]" captionStart="FIGURES 30–44" captionStartId="6.[151,264,1923,1947]" captionTargetBox="[151,1436,181,1899]" captionTargetId="figure-18@6.[151,1436,181,1899]" captionTargetPageId="6" captionText="FIGURES 30–44. Underside of figures 15–29, scale bar for all except fig. 27." pageId="8" pageNumber="394">30–38</figureCitation>
,
<figureCitation id="137070D6FFD3FFAFFE65F906E0C4F95E" box="[398,427,1684,1711]" captionStart="FIGURES 45" captionStartId="7.[151,264,1648,1672]" captionTargetBox="[151,1436,596,1624]" captionTargetId="figure-241@7.[151,1436,596,1624]" captionTargetPageId="7" captionText="FIGURES 45. Comparison of details of the wing patterns of Euthalia duda sspp." pageId="8" pageNumber="394">45</figureCitation>
a-g, 58, 59, 62–65), stating that both taxa cannot be separated soundly from each other. However,
<bibRefCitation id="EFDA11A2FFD3FFAFFEE0F92BE0D2F922" author="Yokochi, T." box="[267,445,1721,1747]" pageId="8" pageNumber="394" pagination="9 - 100" refId="ref5259" refString="Yokochi, T. (2012) Revision of the Subgenus Limbusa Moore, [1897] (Lepidoptera, Nymphalidae, Adoliadini) Part 3. Descriptions of species (2). Bulletin of the Kitakyushu Museum of Natural History and Human History, Series A, 10, 9-100." type="journal article" year="2012">Yokochi (2012)</bibRefCitation>
resurrected the status of
<taxonomicName id="4C4B17D0FFD3FFAFFD35F92BE251F923" authorityName="H. Huang" authorityYear="1999" box="[734,830,1721,1746]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="394" phylum="Arthropoda" rank="species" species="tsangpoi">
<emphasis id="B93FB041FFD3FFAFFD35F92BE251F923" box="[734,830,1721,1746]" italics="true" pageId="8" pageNumber="394">tsangpoi</emphasis>
</taxonomicName>
as a distinct species mainly based on the differences found in the male genitalia, e.g. the longer valva, broader uncus, shorter and undivided cornuti and more dents at the distal end of valva. To tackle the issue, we examined a series of specimens of both sexes of
<taxonomicName id="4C4B17D0FFD3FFAFFAEFF893E432F8EA" authorityName="Staudinger" authorityYear="1886" box="[1284,1373,1793,1819]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="394" phylum="Arthropoda" rank="species" species="duda">
<emphasis id="B93FB041FFD3FFAFFAEFF893E432F8EA" box="[1284,1373,1793,1819]" italics="true" pageId="8" pageNumber="394">E. duda</emphasis>
</taxonomicName>
from various localities across its distribution including the
<typeStatus id="54F0D2F1FFD3FFAFFD01F8B7E23BF8CE" box="[746,852,1829,1855]" pageId="8" pageNumber="394" type="lectotype">lectotype</typeStatus>
, and several individuals of
<taxonomicName id="4C4B17D0FFD3FFAFFB6FF8B7E46CF8CF" authorityName="H. Huang" authorityYear="1999" box="[1156,1283,1829,1854]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="394" phylum="Arthropoda" rank="species" species="tsangpoi">
<emphasis id="B93FB041FFD3FFAFFB6FF8B7E46CF8CF" box="[1156,1283,1829,1854]" italics="true" pageId="8" pageNumber="394">E. tsangpoi</emphasis>
</taxonomicName>
including the
<typeStatus id="54F0D2F1FFD3FFAFFF7CF8DBE192F892" box="[151,253,1865,1891]" pageId="8" pageNumber="394" type="holotype">holotype</typeStatus>
. The examination of the male genitalia (
<figureCitation id="137070D6FFD3FFAFFD25F8DBE239F892" box="[718,854,1864,1891]" captionStart-0="FIGURES 56–59" captionStart-1="FIGURES 60–65" captionStartId-0="10.[151,264,1398,1422]" captionStartId-1="11.[151,264,1943,1967]" captionTargetBox-0="[151,1436,182,1373]" captionTargetBox-1="[199,1385,189,1918]" captionTargetId-0="figure-18@10.[151,1436,181,1374]" captionTargetId-1="figure-28@11.[190,1397,181,1919]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 56–59. Male genitalia of the type specimens of Euthalia duda sspp. Depositary of the specimens: 56 &amp; 57 in CHH, 58 &amp; 59 in MfN, scale bar for figs. 58 &amp; 59. a=genitalia capsule, b=phallus in lateral view, c=tip of valva in its full view." captionText-1="FIGURES 60–65. Male genitalia of Euthalia duda sspp. Depositary of the specimens: 60 in TAAHU, 61 in CLSY, 62 in MfN, 63 &amp; 65 in ZFMK, 64 in NMB. a=genitalia capsule, b=phallus in lateral view, c=tip of valva in its full view." pageId="8" pageNumber="394">Figs. 57–65</figureCitation>
) revealed that no constant difference can be found between both taxa, and the differences summarized by
<bibRefCitation id="EFDA11A2FFD3FFAFFD1DF8FFE2CAF876" author="Yokochi, T." box="[758,933,1901,1927]" pageId="8" pageNumber="394" pagination="9 - 100" refId="ref5259" refString="Yokochi, T. (2012) Revision of the Subgenus Limbusa Moore, [1897] (Lepidoptera, Nymphalidae, Adoliadini) Part 3. Descriptions of species (2). Bulletin of the Kitakyushu Museum of Natural History and Human History, Series A, 10, 9-100." type="journal article" year="2012">Yokochi (2012)</bibRefCitation>
shall be attributed to individual variation and the inaccuracy in the handdrawing (
<figureCitation id="137070D6FFD3FFAFFDDBF803E3ECF85A" box="[560,643,1936,1963]" captionStart="FIGURES 56–59" captionStartId="10.[151,264,1398,1422]" captionTargetBox="[151,1436,182,1373]" captionTargetId="figure-18@10.[151,1436,181,1374]" captionTargetPageId="10" captionText="FIGURES 56–59. Male genitalia of the type specimens of Euthalia duda sspp. Depositary of the specimens: 56 &amp; 57 in CHH, 58 &amp; 59 in MfN, scale bar for figs. 58 &amp; 59. a=genitalia capsule, b=phallus in lateral view, c=tip of valva in its full view." pageId="8" pageNumber="394">Fig. 56</figureCitation>
) of the male genitalia of the
<typeStatus id="54F0D2F1FFD3FFAFFC39F803E55BF85A" box="[978,1076,1937,1963]" pageId="8" pageNumber="394" type="holotype">holotype</typeStatus>
illustrated in H.
<bibRefCitation id="EFDA11A2FFD3FFAFFB1CF803E4F8F85A" author="Huang, H." box="[1271,1431,1937,1963]" pageId="8" pageNumber="394" pagination="642 - 676" refId="ref4575" refString="Huang, H. (1999) Some new butterflies from China. I. (Rhopalocera). Lambillionea, XCIX (4), 642-676." type="journal article" year="1999">Huang (1999)</bibRefCitation>
, compared with the image of the genitalia slide (
<figureCitation id="137070D6FFD3FFAFFD48F827E399F83E" box="[675,758,1972,1999]" captionStart="FIGURES 56–59" captionStartId="10.[151,264,1398,1422]" captionTargetBox="[151,1436,182,1373]" captionTargetId="figure-18@10.[151,1436,181,1374]" captionTargetPageId="10" captionText="FIGURES 56–59. Male genitalia of the type specimens of Euthalia duda sspp. Depositary of the specimens: 56 &amp; 57 in CHH, 58 &amp; 59 in MfN, scale bar for figs. 58 &amp; 59. a=genitalia capsule, b=phallus in lateral view, c=tip of valva in its full view." pageId="8" pageNumber="394">Fig. 57</figureCitation>
). Although
<bibRefCitation id="EFDA11A2FFD3FFAFFC9EF827E54CF83E" author="Yokochi, T." box="[885,1059,1973,1999]" pageId="8" pageNumber="394" pagination="9 - 100" refId="ref5259" refString="Yokochi, T. (2012) Revision of the Subgenus Limbusa Moore, [1897] (Lepidoptera, Nymphalidae, Adoliadini) Part 3. Descriptions of species (2). Bulletin of the Kitakyushu Museum of Natural History and Human History, Series A, 10, 9-100." type="journal article" year="2012">Yokochi (2012)</bibRefCitation>
stated that limited differences can be found in wing patterns between the type specimens of both taxa, after examining a long series of newly available material, we found that
<taxonomicName id="4C4B17D0FFD2FFAEFE4EFF05E34AFF41" authorityName="H. Huang" authorityYear="1999" box="[421,549,151,176]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="395" phylum="Arthropoda" rank="species" species="tsangpoi">
<emphasis id="B93FB041FFD2FFAEFE4EFF05E34AFF41" box="[421,549,151,176]" italics="true" pageId="9" pageNumber="395">E. tsangpoi</emphasis>
</taxonomicName>
generally differs from
<taxonomicName id="4C4B17D0FFD2FFAEFCC7FF05E2EBFF40" authorityName="Staudinger" authorityYear="1886" box="[812,900,151,177]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="395" phylum="Arthropoda" rank="species" species="duda">
<emphasis id="B93FB041FFD2FFAEFCC7FF05E2EBFF40" box="[812,900,151,177]" italics="true" pageId="9" pageNumber="395">E. duda</emphasis>
</taxonomicName>
by the basally narrower discal patch in space 3 on forewing upperside and the more obliquely placed patches in spaces 5 and 6 of forewing upperside in male (
<figureCitation id="137070D6FFD2FFAEFA84FF29E1DBFF08" captionStart="FIGURES 45" captionStartId="7.[151,264,1648,1672]" captionTargetBox="[151,1436,596,1624]" captionTargetId="figure-241@7.[151,1436,596,1624]" captionTargetPageId="7" captionText="FIGURES 45. Comparison of details of the wing patterns of Euthalia duda sspp." pageId="9" pageNumber="395">Fig. 45</figureCitation>
) and the broader discal band on hindwing in the female. Therefore, considering the evidences listed above, it is more reasonable to treat
<taxonomicName id="4C4B17D0FFD2FFAEFE47FE91E344FEED" authorityName="H. Huang" authorityYear="1999" box="[428,555,259,284]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="395" phylum="Arthropoda" rank="species" species="tsangpoi">
<emphasis id="B93FB041FFD2FFAEFE47FE91E344FEED" box="[428,555,259,284]" italics="true" pageId="9" pageNumber="395">E. tsangpoi</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="4C4B17D0FFD2FFAEFCEAFE91E237FEEC" authorityName="Staudinger" authorityYear="1886" box="[769,856,259,285]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="395" phylum="Arthropoda" rank="species" species="duda">
<emphasis id="B93FB041FFD2FFAEFCEAFE91E237FEEC" box="[769,856,259,285]" italics="true" pageId="9" pageNumber="395">E. duda</emphasis>
</taxonomicName>
, viz.
<taxonomicName id="4C4B17D0FFD2FFAEFC7CFE91E53AFEED" authorityName="Huang &amp; Xu &amp; Lu &amp; Lu" authorityYear="2025" baseAuthorityName="H. Huang" baseAuthorityYear="1999" box="[919,1109,259,285]" class="Insecta" family="Nymphalidae" genus="Euthalia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="395" phylum="Arthropoda" rank="subSpecies" species="duda" status="stat. nov." subSpecies="tsangpoi">
<emphasis id="B93FB041FFD2FFAEFC7CFE91E53AFEED" box="[919,1109,259,285]" italics="true" pageId="9" pageNumber="395">E. duda tsangpoi</emphasis>
</taxonomicName>
<emphasis id="B93FB041FFD2FFAEFBB7FE91E5A8FEEC" bold="true" box="[1116,1223,259,285]" pageId="9" pageNumber="395">
<taxonomicNameLabel id="A20C0D3AFFD2FFAEFBB7FE91E5A8FEEC" box="[1116,1223,259,285]" pageId="9" pageNumber="395">stat. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3513FD8FFD2FFAEFF2CFEB4E52BFEB0" box="[199,1092,294,321]" pageId="9" pageNumber="395" type="distribution">
<paragraph id="8BF46C53FFD2FFAEFF2CFEB4E52BFEB0" blockId="9.[151,1437,151,321]" box="[199,1092,294,321]" pageId="9" pageNumber="395">
<emphasis id="B93FB041FFD2FFAEFF2CFEB4E00EFEB1" bold="true" box="[199,353,294,320]" pageId="9" pageNumber="395">Distribution.</emphasis>
Southeast
<collectingRegion id="498FA2B1FFD2FFAEFE30FEB5E343FEB0" box="[475,556,295,321]" country="China" name="Xizang" pageId="9" pageNumber="395">Xizang</collectingRegion>
(Motuo and Bomi Counties), Southwest
<collectingCountry id="F35C2CC3FFD2FFAEFC12FEB5E52FFEB0" box="[1017,1088,295,321]" name="China" pageId="9" pageNumber="395">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

79
data/18/03/87/180387F64B01FFEE06DD33126A48F189.xml
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@ -1,54 +1,55 @@
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<figureCitation id="08912A654B27FFCB06D5334C6FC6F631" box="[159,242,1874,1900]" captionStart="FIGURE 34" captionStartId="44.[152,255,1942,1966]" captionTargetBox="[183,1404,178,1917]" captionTargetId="figure-20@44.[183,1404,178,1917]" captionTargetPageId="44" captionText="FIGURE 34. Esenbeckia yepocapa, paratype male. A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, frontal view. D. Labels." figureDoi="http://doi.org/10.5281/zenodo.15215993" httpUri="https://zenodo.org/record/15215993/files/figure.png" pageId="43" pageNumber="44">Fig. 34</figureCitation>
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83
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