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<document id="020C71E6F1F3C1D2517DE3CFE0DDB35E" ID-CLB-Dataset="300706" ID-DOI="10.11646/phytotaxa.644.1.9" ID-GBIF-Dataset="23652f99-bd8a-4230-91b4-30d925b48b89" ID-ISSN="1179-3163" ID-Zenodo-Dep="13214547" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1722597856789" checkinUser="felipe" docAuthor="Jiménez, Pedro Díaz, Croat, Thomas B., Amith, Jonathan D., Pérez-Farrera, Miguel Ángel &amp; Aguilar-Rodríguez, Pedro A." docDate="2024" docId="0328482E505E2B2FFF27FD446BCAF8A1" docLanguage="en" docName="phytotaxa.644.1.9.pdf" docOrigin="Phytotaxa 644 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.644.1.9" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Anthurium tiswatl Diaz Jim. &amp; Croat, sp. nov." docType="treatment" docVersion="4" lastPageNumber="66" masterDocId="FF113056505F2B2CFFAFFFB06E51FFE8" masterDocTitle="A new species of broad-leafed Anthurium (Araceae) from the central region of Veracruz, Mexico" masterLastPageNumber="68" masterPageNumber="63" pageNumber="64" updateTime="1724266421709" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="4A066D3810133287FDA7CEFB670B12C6">
<mods:title id="F39EA81A36D17FBC5AA28AEA7BC91D7F">A new species of broad-leafed Anthurium (Araceae) from the central region of Veracruz, Mexico</mods:title>
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<mods:name id="E1081CD7E907871D51F0F812EA59E419" type="personal">
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<mods:namePart id="81DEC0B4C60C0F26ADEF703D4B0EF394">Jiménez, Pedro Díaz</mods:namePart>
<mods:nameIdentifier id="DE329F5F6067F010EA658D0DBC361C20" type="ORCID">0000-0003-2079-674X</mods:nameIdentifier>
<mods:affiliation id="841299A78BD09C1F3ECCAADD00AF2EEC">Jardín Botánico “ Araceario Thomas B. Croat ”, s / n, Comalcalco, Tabasco, Mexico</mods:affiliation>
<mods:nameIdentifier id="F697967608805711418CCA5DE9259A11" type="email">aroid764@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="E94841B245852ADB09E6714854E33599">Croat, Thomas B.</mods:namePart>
<mods:nameIdentifier id="F711703EB2581494742038487E95CDF8" type="ORCID">0000-0001-6810-0567</mods:nameIdentifier>
<mods:affiliation id="534A6A5210D7286D3A976F0CE477F060">Missouri Botanical Garden, 4344 Shaw Blvd., St. Louis, Missouri 63110, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="25F17684ECE0D8EC314005B617B595B8" type="email">Thomas.Croat@mobot.org</mods:nameIdentifier>
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<mods:name id="E7E393D1873E07616AA4A8DBF8B79294" type="personal">
<mods:role id="47051BE2A8FA4C0AF2504AE85079AEAA">
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<mods:namePart id="F64F6FDD15EAF462B27EA21A7640365E">Amith, Jonathan D.</mods:namePart>
<mods:nameIdentifier id="D86CB80E69FD81FFC5BF6613832371AB" type="ORCID">0000-0001-7518-8456</mods:nameIdentifier>
<mods:affiliation id="608FD681C178E0E7C77DEE2A3882A149">Department of Anthropology, Gettysburg College, Box 2985, 300 N, Washington Street, Gettysburg, PA 17325, U. S. A. &amp; Research fellow, Dept. of Anthropology, Gettysburg College, 300 N. Washington Street, Gettysburg, PA 17325, U. S. A. and Research associate, Department of Botany, National Museum of Natural History, Smithsonian Institution, 10 th St. &amp; Constitution Ave. NW, Washington, DC 20004, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="9CA90CF784DA78F8D06BA5096A442207" type="email">nahuatl.biology@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="0DDB76B29B69693A6353A4A602A91D36">Pérez-Farrera, Miguel Ángel</mods:namePart>
<mods:nameIdentifier id="5FD659E4A7A13AE137E5B6F28FD3202B" type="ORCID">0000-0002-5329-1505</mods:nameIdentifier>
<mods:affiliation id="A49B9A20F0E98272F270D0EF6C212172">Herbario Eizi Matuda, Laboratorio de Ecología Evolutiva, Instituto de Ciencias Biológicas, Universidad de Ciencias y Artes de Chiapas. Libramiento Norte Poniente 1150, LaJas Maciel, Tuxtla Gutiérrez, Chiapas, 29039, Mexico</mods:affiliation>
<mods:nameIdentifier id="7B4FBE18F6BB8CBFA4E9B2C56ACA3A7E" type="email">perezfarreram@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8B5E7CAE0F22D610B18B400D3ED09406">Aguilar-Rodríguez, Pedro A.</mods:namePart>
<mods:nameIdentifier id="128A5EECFAB606D28B2F3130EE59D2C7" type="ORCID">0000-0002-9275-4322</mods:nameIdentifier>
<mods:affiliation id="B9EF72489A182877653B658C60AC4758">Instituto de Investigaciones Forestales, Universidad Veracruzana, Parque Ecológico “ El Haya ”, Carretera Antigua a Coatepec, 91070 Xalapa-Enríquez, Veracruz, Mexico</mods:affiliation>
<mods:nameIdentifier id="4837CD793EBFF8860AC3574AA8A6EAE5" type="email">pedroaguilarr@gmail.com</mods:nameIdentifier>
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<mods:title id="EAE710EF4528845DB4C70607FFC07A30">Phytotaxa</mods:title>
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<treatment id="0328482E505E2B2FFF27FD446BCAF8A1" ID-DOI="http://doi.org/10.5281/zenodo.13357000" ID-GBIF-Taxon="237647136" ID-Zenodo-Dep="13357000" LSID="urn:lsid:plazi:treatment:0328482E505E2B2FFF27FD446BCAF8A1" httpUri="http://treatment.plazi.org/id/0328482E505E2B2FFF27FD446BCAF8A1" lastPageId="3" lastPageNumber="66" pageId="1" pageNumber="64">
<subSubSection id="C39BAAB3505E2B2DFF27FD446CBFFCE6" box="[136,750,756,783]" pageId="1" pageNumber="64" type="nomenclature">
<paragraph id="8B3EF938505E2B2DFF27FD446CBFFCE6" blockId="1.[136,750,756,783]" box="[136,750,756,783]" pageId="1" pageNumber="64">
<heading id="D0764E54505E2B2DFF27FD446CBFFCE6" box="[136,750,756,783]" fontSize="11" level="2" pageId="1" pageNumber="64" reason="3">
<taxonomicName id="4C8182BB505E2B2DFF27FD446C63FCE6" authority="Diaz Jim. &amp; Croat" authorityName="Diaz Jim. &amp; Croat" box="[136,562,756,783]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="tiswatl" status="sp. nov.">
<emphasis id="B9F5252A505E2B2DFF27FD446F0AFCE7" bold="true" box="[136,347,756,783]" italics="true" pageId="1" pageNumber="64">Anthurium tiswatl</emphasis>
Díaz Jim. &amp; Croat
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,
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<emphasis id="B9F5252A505E2B2DFD92FD456CDDFCE6" box="[573,652,757,782]" italics="true" pageId="1" pageNumber="64">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="13BAE5BD505E2B2DFD0CFD446CB2FCE6" box="[675,739,756,782]" captionStart="FIGURE 1" captionStartId="2.[136,229,1877,1899]" captionTargetBox="[151,1435,187,1849]" captionTargetId="figure-23@2.[151,1435,187,1849]" captionTargetPageId="2" captionText="FIGURE 1. Anthurium tiswatl. A. Adult plant in its habit; B. Adaxial view of the leaf blade with a portion of the petiole; C. Inflorescence at anthesis (male phase) showing yellowish-green spathe, tinged with purple on the margins; D. Infructescence showing yellowish-green immature berries. Photos by P. Díaz Jiménez." figureDoi="http://doi.org/10.5281/zenodo.13214549" httpUri="https://zenodo.org/record/13214549/files/figure.png" pageId="1" pageNumber="64">Fig. 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C39BAAB3505E2B2DFF27FCF06D17FBE2" pageId="1" pageNumber="64" type="diagnosis">
<paragraph id="8B3EF938505E2B2DFF27FCF06D17FBE2" blockId="1.[136,1452,832,1106]" pageId="1" pageNumber="64">
<taxonomicName id="4C8182BB505E2B2DFF27FCF06FB9FCBD" authority="Diaz Jim. &amp; Croat" authorityName="Diaz Jim. &amp; Croat" box="[136,488,832,854]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="tiswatl">
<emphasis id="B9F5252A505E2B2DFF27FCF06F65FCBD" box="[136,308,832,853]" italics="true" pageId="1" pageNumber="64">Anthurium tiswatl</emphasis>
Diaz Jim. &amp; Croat
</taxonomicName>
is similar to
<taxonomicName id="4C8182BB505E2B2DFDCAFCF06DA4FCBD" ID-CoL="67JCP" authority="Matuda (1951: 373 - 374)" authorityName="Matuda" authorityPageNumber="373 - 374" authorityYear="1951" box="[613,1013,832,854]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="macdougallii">
<emphasis id="B9F5252A505E2B2DFDCAFCF06CAFFCBD" box="[613,766,832,853]" italics="true" pageId="1" pageNumber="64">A. macdougallii</emphasis>
<bibRefCitation id="EF1084C9505E2B2DFCACFCF06DA4FCBD" author="Matuda, E." box="[771,1013,832,854]" pageId="1" pageNumber="64" pagination="369 - 383" refId="ref3767" refString="Matuda, E. (1951) Nuevas Araceas de Mexico. Anales del instituto de Biologia de la Universidad Nacional de Mexico 22: 369 - 383." type="journal article" year="1951">Matuda (1951: 373–374)</bibRefCitation>
</taxonomicName>
, but differs from that species in having longer cataphylls (11–16
<emphasis id="B9F5252A505E2B2DFEC2FCD46FD9FC91" box="[365,392,868,889]" italics="true" pageId="1" pageNumber="64">vs.</emphasis>
5.5–7.0 cm in
<taxonomicName id="4C8182BB505E2B2DFDB9FCD46CFEFC91" ID-CoL="67JCP" authorityName="Matuda" authorityYear="1951" box="[534,687,868,889]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="macdougallii">
<emphasis id="B9F5252A505E2B2DFDB9FCD46CFEFC91" box="[534,687,868,889]" italics="true" pageId="1" pageNumber="64">A. macdougallii</emphasis>
</taxonomicName>
), longer blades (44–65
<emphasis id="B9F5252A505E2B2DFC3CFCD46DFCFC91" box="[915,941,868,889]" italics="true" pageId="1" pageNumber="64">vs.</emphasis>
<quantity id="4C7954DD505E2B2DFC1DFCD46A44FC91" box="[946,1045,868,889]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="3.3" metricValueMin="1.8" pageId="1" pageNumber="64" unit="cm" value="25.5" valueMax="33.0" valueMin="18.0">18–33 cm</quantity>
), collective veins arising from first or fifth pairs of basal veins (
<emphasis id="B9F5252A505E2B2DFE2DFC386FCDFC75" box="[386,412,904,925]" italics="true" pageId="1" pageNumber="64">vs.</emphasis>
collective veins arising from the fourth or sixth pairs of basal veins), and yellowish and whitish berries when ripe (
<emphasis id="B9F5252A505E2B2DFF5DFC1C6F5DFC29" box="[242,268,940,961]" italics="true" pageId="1" pageNumber="64">vs.</emphasis>
orange). It can also be confused with
<taxonomicName id="4C8182BB505E2B2DFD2FFC1C6D97FC29" ID-CoL="5VHJL" authority="Liebmann (1849: 22)" authorityName="Liebmann" authorityPageNumber="22" authorityYear="1849" box="[640,966,940,962]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="andicola">
<emphasis id="B9F5252A505E2B2DFD2FFC1C6CA1FC29" box="[640,752,940,961]" italics="true" pageId="1" pageNumber="64">A. andicola</emphasis>
<bibRefCitation id="EF1084C9505E2B2DFD58FC1C6D97FC29" author="Liebmann, F. M." box="[759,966,940,962]" pageId="1" pageNumber="64" pagination="1 - 120" refId="ref3740" refString="Liebmann, F. M. (1849) Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i KJObenhavn (1 - 2): 1 - 120." type="book chapter" year="1849">Liebmann (1849: 22)</bibRefCitation>
</taxonomicName>
, but
<taxonomicName id="4C8182BB505E2B2DFC58FC1C6A03FC29" box="[1015,1106,940,961]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="64" phylum="Tracheophyta" rank="species" species="tiswatl">
<emphasis id="B9F5252A505E2B2DFC58FC1C6A03FC29" box="[1015,1106,940,961]" italics="true" pageId="1" pageNumber="64">A. tiswatl</emphasis>
</taxonomicName>
differs in having leaf blades with a greater number of basal lateral veins (7–9
<emphasis id="B9F5252A505E2B2DFDFBFC606C3FFC0D" box="[596,622,976,997]" italics="true" pageId="1" pageNumber="64">vs.</emphasis>
3–6), yellowish-green or light green spathe at anthesis (
<emphasis id="B9F5252A505E2B2DFB25FC606AF4FC0D" box="[1162,1189,976,997]" italics="true" pageId="1" pageNumber="64">vs.</emphasis>
dark purple to brown), and yellowish and whitish berries when ripe (
<emphasis id="B9F5252A505E2B2DFDE4FC446C34FBE1" box="[587,613,1012,1033]" italics="true" pageId="1" pageNumber="64">vs.</emphasis>
light violet and white).
</paragraph>
</subSubSection>
<subSubSection id="C39BAAB3505E2B2DFF27FBA86A51FBB9" pageId="1" pageNumber="64" type="materials_examined">
<paragraph id="8B3EF938505E2B2DFF27FBA86A51FBB9" blockId="1.[136,1452,832,1106]" pageId="1" pageNumber="64">
<materialsCitation id="3BE9F365505E2B2DFF27FBA86D49FBB9" collectingDate="2022-04-06" collectionCode="XAL" collectorName="Pedro Diaz Jimenez &amp; Gabriela Citlahua Zepahua" country="Mexico" elevation="1804" latitude="18.735834" location="Tlaxacapitzco" longLatPrecision="21" longitude="-97.06556" municipality="Municipio Tequila" pageId="1" pageNumber="64" specimenCount="1" stateProvince="Veracruz" typeStatus="holotype">
<emphasis id="B9F5252A505E2B2DFF27FBA86E93FBC5" bold="true" box="[136,194,1048,1069]" pageId="1" pageNumber="64">Type:</emphasis>
—
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.
<collectingRegion id="494537DA505E2B2DFEEAFBA86FCFFBC5" box="[325,414,1048,1069]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="1" pageNumber="64">Veracruz</collectingRegion>
:
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,
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,
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,
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,
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,
<quantity id="4C7954DD505E2B2DFBCCFBA86AFFFBC5" box="[1123,1198,1048,1069]" metricMagnitude="3" metricUnit="m" metricValue="1.8039999999999998" pageId="1" pageNumber="64" unit="m" value="1804.0">
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,
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<collectingDate id="EF7B2610505E2B2DFB17FBA86B65FBC5" box="[1208,1332,1048,1070]" pageId="1" pageNumber="64" value="2022-04-06">6 April 2022</collectingDate>
</date>
,
<emphasis id="B9F5252A505E2B2DFA91FBA86C3BFBB9" italics="true" pageId="1" pageNumber="64">
<collectorName id="26749CEE505E2B2DFA91FBA86F5BFBB9" pageId="1" pageNumber="64">Pedro Díaz Jiménez</collectorName>
&amp;
<collectorName id="26749CEE505E2B2DFE86FB8C6C65FBB9" box="[297,564,1084,1105]" pageId="1" pageNumber="64">Gabriela Citlahua Zepahua</collectorName>
1674
</emphasis>
(
<typeStatus id="543A479A505E2B2DFDD8FB8C6C81FBBA" box="[631,720,1084,1106]" pageId="1" pageNumber="64" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED9061FD505E2B2DFD76FB8C6D40FBB9" box="[729,785,1084,1105]" country="Mexico" httpUri="http://grbio.org/cool/wbe7-qwr9" name="Instituto de Ecología, A.C." pageId="1" pageNumber="64" type="Herbarium">XAL</collectionCode>
!,
</materialsCitation>
<materialsCitation id="3BE9F365505E2B2DFCB1FB8C6DADFBB9" box="[798,1020,1084,1106]" collectingDate="2022-04-06" collectionCode="HEM, MO" collectorName="Pedro Diaz Jimenez &amp; Gabriela Citlahua Zepahua" country="Mexico" elevation="1804" latitude="18.735834" location="Tlaxacapitzco" longLatPrecision="21" longitude="-97.06556" municipality="Municipio Tequila" pageId="1" pageNumber="64" specimenCount="1" stateProvince="Veracruz" typeStatus="isotype">
<typeStatus id="543A479A505E2B2DFCB1FB8C6D21FBBA" box="[798,880,1084,1106]" pageId="1" pageNumber="64">isotypes</typeStatus>
:
<collectionCode id="ED9061FD505E2B2DFCD6FB8C6DE4FBB9" box="[889,949,1084,1105]" pageId="1" pageNumber="64">HEM</collectionCode>
!,
<collectionCode id="ED9061FD505E2B2DFC6CFB8C6DA0FBB9" box="[963,1009,1084,1105]" pageId="1" pageNumber="64">MO</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C39BAAB3505E2B2DFF27FB306FD1F7F6" pageId="1" pageNumber="64" type="description">
<paragraph id="8B3EF938505E2B2DFF27FB306FD1F7F6" blockId="1.[136,1452,1152,2079]" pageId="1" pageNumber="64">
Terrestrial or epipetric;
<emphasis id="B9F5252A505E2B2DFE20FB306F96FB72" bold="true" box="[399,455,1152,1178]" pageId="1" pageNumber="64">stem</emphasis>
thick, up to
<quantity id="4C7954DD505E2B2DFDFDFB306CC7FB73" box="[594,662,1152,1179]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="64" unit="cm" value="16.0">16 cm</quantity>
long and
<quantity id="4C7954DD505E2B2DFCABFB306D06FB73" box="[772,855,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="64" unit="cm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 cm</quantity>
in diam.; leaf scars
<quantity id="4C7954DD505E2B2DFB99FB306AD9FB73" box="[1078,1160,1152,1179]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="64" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 cm</quantity>
wide; roots thick, 2.8–5.0 mm in diam.; cataphylls subcoriaceous,
<quantity id="4C7954DD505E2B2DFDE9FB146CE2FB57" box="[582,691,1188,1215]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="1" pageNumber="64" unit="cm" value="13.5" valueMax="16.0" valueMin="11.0">11–16 cm</quantity>
long, light green, acuminate at apex, drying light brown, weathering to a fibrous network at the base, persisting.
<emphasis id="B9F5252A505E2B2DFDC2FB786CEEFB0A" bold="true" box="[621,703,1224,1250]" pageId="1" pageNumber="64">Leaves</emphasis>
erect; petioles erect,
<quantity id="4C7954DD505E2B2DFC03FB786A4AFB0B" box="[940,1051,1224,1251]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" metricValueMax="7.2" metricValueMin="4.8" pageId="1" pageNumber="64" unit="cm" value="60.0" valueMax="72.0" valueMin="48.0">48–72 cm</quantity>
long, 10.0–
<quantity id="4C7954DD505E2B2DFB0EFB786B52FB0B" box="[1185,1283,1224,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="64" unit="mm" value="19.5">19.5 mm</quantity>
diam., broadly sulcate with acute margins or flattened adaxially, light green or brown-reddish, drying dark brown; geniculum 3.0–
<quantity id="4C7954DD505E2B2DFA25FB5C6EFBFAC3" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="64" unit="cm" value="3.1">3.1 cm</quantity>
long,
<quantity id="4C7954DD505E2B2DFF5DFAA06FD0FAC3" box="[242,385,1296,1323]" metricMagnitude="-2" metricUnit="m" metricValue="1.875" metricValueMax="2.0" metricValueMin="1.75" pageId="1" pageNumber="64" unit="mm" value="18.75" valueMax="20.0" valueMin="17.5">17.5–20 mm</quantity>
diam., flattened or sulcate adaxially, green or reddish; blades widely ovate to ovate-triangular, coriaceous, acute or rounded at apex, sometimes with an apiculum up to
<quantity id="4C7954DD505E2B2DFC1EFA846DA3FAA7" box="[945,1010,1332,1359]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="64" unit="mm" value="5.0">5 mm</quantity>
long, deeply and broadly lobed at base,
<quantity id="4C7954DD505E2B2DFF27FAE86EA8FA9B" box="[136,249,1368,1395]" metricMagnitude="-1" metricUnit="m" metricValue="5.45" metricValueMax="6.5" metricValueMin="4.4" pageId="1" pageNumber="64" unit="cm" value="54.5" valueMax="65.0" valueMin="44.0">44–65 cm</quantity>
long,
<quantity id="4C7954DD505E2B2DFEEAFAE86FE7FA9B" box="[325,438,1368,1395]" metricMagnitude="-1" metricUnit="m" metricValue="4.6" metricValueMax="5.6" metricValueMin="3.6" pageId="1" pageNumber="64" unit="cm" value="46.0" valueMax="56.0" valueMin="36.0">36–56 cm</quantity>
wide, broadest at point of petiole attachment; anterior lobe
<quantity id="4C7954DD505E2B2DFBC5FAE86A8AFA9B" box="[1130,1243,1368,1395]" metricMagnitude="-1" metricUnit="m" metricValue="3.9" metricValueMax="4.8" metricValueMin="3.0" pageId="1" pageNumber="64" unit="cm" value="39.0" valueMax="48.0" valueMin="30.0">30–48 cm</quantity>
long, the margins broadly rounded and ± undulate; posterior lobes
<quantity id="4C7954DD505E2B2DFD10FACC6D63FA7F" box="[703,818,1404,1431]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="1" pageNumber="64" unit="cm" value="18.0" valueMax="20.0" valueMin="16.0">16–20 cm</quantity>
long,
<quantity id="4C7954DD505E2B2DFCD0FACC6DA0FA7F" box="[895,1009,1404,1431]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="1" pageNumber="64" unit="cm" value="19.0" valueMax="25.0" valueMin="13.0">13–25 cm</quantity>
wide, rounded at apex, often directed outward; sinus parabolic or ± birettiform, rounded at apex; the upper surface glossy dark green, the lower surface semiglossy light green, drying matte green or brown semiglossy on both surfaces; midrib raised above, diminished and flat at apex, raised below, dark green or yellowish-green above and light green or yellowish below; basal veins 7–9 pairs, the first or second free to base, the second to the third coalesced to
<quantity id="4C7954DD505E2B2DFC57F9BC6A19F9CF" box="[1016,1096,1548,1575]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="64" unit="cm" value="2.5">2.5 cm</quantity>
, third to the fourth of
<quantity id="4C7954DD505E2B2DFAECF9BC6BFAF9CF" box="[1347,1451,1548,1575]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="1" pageNumber="64" unit="cm" value="2.75" valueMax="4.0" valueMin="1.5">1.5–4 cm</quantity>
and fourth to remaining of
<quantity id="4C7954DD505E2B2DFE1CF9806C58F9A3" box="[435,521,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="64" unit="cm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 cm</quantity>
; posterior ribs naked; primary lateral veins 5–7 per side, departing midrib at 20–35° angle, slightly raised above, raised below, yellowish-green above and light green or yellowish below, tertiary venation visible on both sides, prominently raised below; collective veins arising from first or fifth pair of basal veins, sunken above, raised below, 4.5–24.0 mm from margin.
<emphasis id="B9F5252A505E2B2DFD1EF92C6D1EF95E" bold="true" box="[689,847,1692,1718]" pageId="1" pageNumber="64">Inflorescence</emphasis>
erect, shorter than leaves;
<emphasis id="B9F5252A505E2B2DFB2EF92C6ABDF95E" bold="true" box="[1153,1260,1692,1718]" pageId="1" pageNumber="64">peduncle</emphasis>
thick,
<quantity id="4C7954DD505E2B2DFA94F92C6BFAF95F" box="[1339,1451,1692,1719]" metricMagnitude="-1" metricUnit="m" metricValue="4.7" metricValueMax="5.4" metricValueMin="4.0" pageId="1" pageNumber="64" unit="cm" value="47.0" valueMax="54.0" valueMin="40.0">40–54 cm</quantity>
long,
<quantity id="4C7954DD505E2B2DFF65F9706F67F933" box="[202,310,1728,1755]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="1" pageNumber="64" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
diam., terete, dark green or brown-reddish;
<emphasis id="B9F5252A505E2B2DFC9EF9706D2EF932" bold="true" box="[817,895,1728,1754]" pageId="1" pageNumber="64">spathe</emphasis>
erect, lanceolate, coriaceous, yellowish-green or light green, and tinged at margins with purple, up to
<quantity id="4C7954DD505E2B2DFD4CF9546D78F917" box="[739,809,1764,1791]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="64" unit="cm" value="11.0">11 cm</quantity>
long,
<quantity id="4C7954DD505E2B2DFCDCF9546D91F917" box="[883,960,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="64" unit="cm" value="4.5">4.5 cm</quantity>
wide, acuminate at apex, subcordate at the base, inserted at 50–60° angle on peduncle, short stipe;
<emphasis id="B9F5252A505E2B2DFCABF8B86D03F8CA" bold="true" box="[772,850,1800,1826]" pageId="1" pageNumber="64">spadix</emphasis>
tapered, dark purple at anthesis, up to
<quantity id="4C7954DD505E2B2DFAA0F8B86B3BF8CB" box="[1295,1386,1800,1827]" metricMagnitude="-1" metricUnit="m" metricValue="1.48" pageId="1" pageNumber="64" unit="cm" value="14.8">14.8 cm</quantity>
long,
<quantity id="4C7954DD505E2B2DFF27F89C6E86F8AF" box="[136,215,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="64" unit="mm" value="17.0">17 mm</quantity>
diam. at the base,
<quantity id="4C7954DD505E2B2DFE06F89D6FBBF8AF" box="[425,490,1837,1863]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="64" unit="mm" value="7.0">7 mm</quantity>
diam. at the apex, with 6–7 flowers visible in the principal spiral, 8–9 flowers visible in the alternate spiral,
<emphasis id="B9F5252A505E2B2DFE25F8E06FB0F882" bold="true" box="[394,481,1872,1898]" pageId="1" pageNumber="64">flowers</emphasis>
rhombic, 2.0–
<quantity id="4C7954DD505E2B2DFD24F8E06CB0F883" box="[651,737,1872,1899]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="64" unit="mm" value="4.1">4.1 mm</quantity>
wide, 2.6–4.5 long, the sides weakly sigmoid; tepals greenish or dark, minutely papillate, the lateral tepals
<quantity id="4C7954DD505E2B2DFD2BF8C46D5DF867" box="[644,780,1908,1935]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.6" metricValueMin="1.2" pageId="1" pageNumber="64" unit="mm" value="1.9" valueMax="2.6" valueMin="1.2">1.2–2.6 mm</quantity>
wide, the inner margin ± straight;
<emphasis id="B9F5252A505E2B2DFB39F8C46A8DF866" bold="true" box="[1174,1244,1908,1934]" pageId="1" pageNumber="64">pistils</emphasis>
weakly emergent,
<quantity id="4C7954DD505E2B2DFF27F8286EABF85B" box="[136,250,1944,1971]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="64" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, dark,
<emphasis id="B9F5252A505E2B2DFED0F8286F9FF85A" bold="true" box="[383,462,1944,1970]" pageId="1" pageNumber="64">stigma</emphasis>
oblong to linear.
<emphasis id="B9F5252A505E2B2DFD3DF8286D11F85A" bold="true" box="[658,832,1944,1970]" pageId="1" pageNumber="64">Infructescence</emphasis>
pendent; spadix up to
<quantity id="4C7954DD505E2B2DFB92F8286AD3F85B" box="[1085,1154,1944,1971]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="64" unit="cm" value="17.0">17 cm</quantity>
long,
<quantity id="4C7954DD505E2B2DFB67F8286B46F85B" box="[1224,1303,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="64" unit="mm" value="28.0">28 mm</quantity>
diam., tepals brown or light brown; immature
<emphasis id="B9F5252A505E2B2DFE54F80C6C1EF83E" bold="true" box="[507,591,1980,2006]" pageId="1" pageNumber="64">berries</emphasis>
green or yellowish-green when immature, yellowish and whitish when ripe, obovoid, ± round at apex, 7.2–12.0 mm long,
<quantity id="4C7954DD505E2B2DFD3EF8506CBFF813" box="[657,750,2016,2043]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="1" pageNumber="64" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">6–9 mm</quantity>
wide;
<emphasis id="B9F5252A505E2B2DFC93F8506D2BF812" bold="true" box="[828,890,2016,2042]" pageId="1" pageNumber="64">seeds</emphasis>
1 or 2, oblong, greenish to whitish, 5.5–10.0 mm long,
<quantity id="4C7954DD505E2B2DFF67F7B46F6BF7F7" box="[200,314,2052,2079]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" pageId="1" pageNumber="64" unit="mm" value="4.75" valueMax="6.0" valueMin="3.5">3.5–6 mm</quantity>
diam.
</paragraph>
</subSubSection>
<caption id="DFFEA9B0505D2B2EFF27F8E56C6DF85B" ID-DOI="http://doi.org/10.5281/zenodo.13214549" ID-Zenodo-Dep="13214549" httpUri="https://zenodo.org/record/13214549/files/figure.png" pageId="2" pageNumber="65" startId="2.[136,229,1877,1899]" targetBox="[151,1435,187,1849]" targetPageId="2" targetType="figure">
<paragraph id="8B3EF938505D2B2EFF27F8E56C6DF85B" blockId="2.[136,1452,1877,1971]" pageId="2" pageNumber="65">
<emphasis id="B9F5252A505D2B2EFF27F8E56EACF882" bold="true" box="[136,253,1877,1899]" pageId="2" pageNumber="65">FIGURE 1.</emphasis>
<taxonomicName id="4C8182BB505D2B2EFEACF8E56FFEF882" box="[259,431,1877,1898]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="65" phylum="Tracheophyta" rank="species" species="tiswatl">
<emphasis id="B9F5252A505D2B2EFEACF8E56FFEF882" box="[259,431,1877,1898]" italics="true" pageId="2" pageNumber="65">Anthurium tiswatl</emphasis>
</taxonomicName>
.
<emphasis id="B9F5252A505D2B2EFE15F8E56F83F882" bold="true" box="[442,466,1877,1898]" pageId="2" pageNumber="65">A.</emphasis>
Adult plant in its habit;
<emphasis id="B9F5252A505D2B2EFD17F8E56C9EF882" bold="true" box="[696,719,1877,1898]" pageId="2" pageNumber="65">B.</emphasis>
Adaxial view of the leaf blade with a portion of the petiole;
<emphasis id="B9F5252A505D2B2EFAA2F8E56B74F883" bold="true" box="[1293,1317,1877,1899]" pageId="2" pageNumber="65">C.</emphasis>
Inflorescence at anthesis (male phase) showing yellowish-green spathe, tinged with purple on the margins;
<emphasis id="B9F5252A505D2B2EFBA6F8C96A70F866" bold="true" box="[1033,1057,1913,1934]" pageId="2" pageNumber="65">D.</emphasis>
Infructescence showing yellowish-green immature berries. Photos by P. Díaz Jiménez.
</paragraph>
</caption>
<caption id="DFFEA9B0505C2B2FFF27FBD26AB5FB91" ID-DOI="http://doi.org/10.5281/zenodo.13214551" ID-Zenodo-Dep="13214551" box="[136,1252,1122,1145]" httpUri="https://zenodo.org/record/13214551/files/figure.png" pageId="3" pageNumber="66" startId="3.[136,229,1122,1144]" targetBox="[151,1435,190,1098]" targetPageId="3" targetType="figure">
<paragraph id="8B3EF938505C2B2FFF27FBD26AB5FB91" blockId="3.[136,1252,1122,1145]" box="[136,1252,1122,1145]" pageId="3" pageNumber="66">
<emphasis id="B9F5252A505C2B2FFF27FBD26EAFFB9F" bold="true" box="[136,254,1122,1144]" pageId="3" pageNumber="66">FIGURE 2.</emphasis>
Map showing the collection site (black dot) of
<taxonomicName id="4C8182BB505C2B2FFD6AFBD36D20FB90" box="[709,881,1123,1144]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="tiswatl" status="sp. nov.">
<emphasis id="B9F5252A505C2B2FFD6AFBD36D20FB90" box="[709,881,1123,1144]" italics="true" pageId="3" pageNumber="66">Anthurium tiswatl</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2C69851505C2B2FFCD8FBD36D90FB90" box="[887,961,1123,1144]" pageId="3" pageNumber="66" rank="species">
<emphasis id="B9F5252A505C2B2FFCD8FBD36DEDFB90" box="[887,956,1123,1144]" italics="true" pageId="3" pageNumber="66">sp. nov</emphasis>
.
</taxonomicNameLabel>
in Tequila, Veracruz, Mexico.
</paragraph>
</caption>
<subSubSection id="C39BAAB3505C2B2FFF12FB176D92FAB9" pageId="3" pageNumber="66" type="distribution">
<paragraph id="8B3EF938505C2B2FFF12FB176D92FAB9" blockId="3.[136,1452,1191,1866]" pageId="3" pageNumber="66">
<emphasis id="B9F5252A505C2B2FFF12FB176C2BFB29" bold="true" box="[189,634,1191,1217]" pageId="3" pageNumber="66">Distribution, habitat and phenology:</emphasis>
—
<taxonomicName id="4C8182BB505C2B2FFD39FB176D33FB29" box="[662,866,1191,1217]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="tiswatl">
<emphasis id="B9F5252A505C2B2FFD39FB176D33FB29" box="[662,866,1191,1217]" italics="true" pageId="3" pageNumber="66">Anthurium tiswatl</emphasis>
</taxonomicName>
is endemic to the municipality of Tequila (
<figureCitation id="13BAE5BD505C2B2FFAF9FB176BCCFB29" box="[1366,1437,1191,1217]" captionStart="FIGURE 2" captionStartId="3.[136,229,1122,1144]" captionTargetBox="[151,1435,190,1098]" captionTargetId="figure-19@3.[151,1435,190,1098]" captionTargetPageId="3" captionText="FIGURE 2. Map showing the collection site (black dot) of Anthurium tiswatl sp. nov. in Tequila, Veracruz, Mexico." figureDoi="http://doi.org/10.5281/zenodo.13214551" httpUri="https://zenodo.org/record/13214551/files/figure.png" pageId="3" pageNumber="66">Fig. 2</figureCitation>
),
<collectingRegion id="494537DA505C2B2FFF27FB7B6EBEFB0D" box="[136,239,1227,1253]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="66">Veracruz</collectingRegion>
,
<collectingCountry id="F396B9A8505C2B2FFF56FB7B6F00FB0D" box="[249,337,1227,1253]" name="Mexico" pageId="3" pageNumber="66">Mexico</collectingCountry>
, in the central region of the state, at
<quantity id="4C7954DD505C2B2FFD48FB7B6DD5FB0D" box="[743,900,1227,1253]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="3" pageNumber="66" unit="m" value="1850.0" valueMax="1900.0" valueMin="1800.0">1800–1900 m</quantity>
, in pine-oak forest. It grows in the understory, on the edges of the forest, or in forest fragments. In the vicinity,
<taxonomicName id="4C8182BB505C2B2FFCE5FB5F6A44FAE1" box="[842,1045,1263,1289]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="tiswatl">
<emphasis id="B9F5252A505C2B2FFCE5FB5F6A44FAE1" box="[842,1045,1263,1289]" italics="true" pageId="3" pageNumber="66">Anthurium tiswatl</emphasis>
</taxonomicName>
may be found in the municipalities of Atlahuilco, Ixtaczoquitlán, Los Reyes, Magdalena, Omealca, Orizaba, San Andrés Tenejapan and Zongolica. All specimens with inflorescences and infructescence were collected in April.
</paragraph>
</subSubSection>
<subSubSection id="C39BAAB3505C2B2FFF12FAEB6A80F9A5" pageId="3" pageNumber="66" type="etymology">
<paragraph id="8B3EF938505C2B2FFF12FAEB6A80F9A5" blockId="3.[136,1452,1191,1866]" pageId="3" pageNumber="66">
<emphasis id="B9F5252A505C2B2FFF12FAEB6F6BFA9D" bold="true" box="[189,314,1371,1397]" pageId="3" pageNumber="66">Eponymy:</emphasis>
—This species is named for the local name as it is known in its
<typeStatus id="543A479A505C2B2FFBA1FAEB6A6FFA9D" box="[1038,1086,1371,1397]" pageId="3" pageNumber="66">type</typeStatus>
locality, named as such because of its morphological traits and habitat. The term
<taxonomicName id="4C8182BB505C2B2FFD65FACF6D75FA71" box="[714,804,1407,1433]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="tiswatl">“tiswatl</taxonomicName>
” is the local, central
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variant pronunciation of “teswatl”, a plant name documented in sources from the mid-16th century (e.g.,
<bibRefCitation id="EF1084C9505C2B2FFBA0FA136A99FA56" author="Hernandez, F." box="[1039,1224,1443,1470]" pageId="3" pageNumber="66" refId="ref3629" refString="Hernandez, F. (1959) Obras completas. Tomo II: Historia natural de Nueva Espana, vols. 1 y 2: Historia de las plantas de Nueva Espana. Universidad Nacional Autonoma de Mexico, Mexico City." type="journal volume" year="1959">Hernández 1959</bibRefCitation>
) and widely used in Nahuatl-speaking communities in the northeastern Sierra de
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for several
<taxonomicName id="4C8182BB505C2B2FFBA1FA776A3BFA09" box="[1038,1130,1479,1505]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
and
<taxonomicName id="4C8182BB505C2B2FFB0CFA776B34FA09" box="[1187,1381,1479,1505]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
alike. An explanation for the local use of the term
<taxonomicName id="4C8182BB505C2B2FFDDCFA5B6C9CF9ED" box="[627,717,1515,1541]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="tiswatl">“tiswatl</taxonomicName>
” for an
<taxonomicName id="4C8182BB505C2B2FFC89FA5B6DCFF9ED" authorityName="Schott" authorityYear="1829" box="[806,926,1515,1541]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B9F5252A505C2B2FFC89FA5B6DCFF9ED" box="[806,926,1515,1541]" italics="true" pageId="3" pageNumber="66">Anthurium</emphasis>
</taxonomicName>
might be related to the etymology of this term: ti- ‘rock’ or ‘stone’ and iswatl ‘broad leaf’ such as that found in maize plants and many other monocots. The
<taxonomicName id="4C8182BB505C2B2FFA9BF9BF6BFDF9C1" authorityName="Schott" authorityYear="1829" box="[1332,1452,1551,1577]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B9F5252A505C2B2FFA9BF9BF6BFDF9C1" box="[1332,1452,1551,1577]" italics="true" pageId="3" pageNumber="66">Anthurium</emphasis>
</taxonomicName>
here described is broad-leafed and is often found in rocky soil, hence the incorporated element ti-.
</paragraph>
</subSubSection>
<subSubSection id="C39BAAB3505C2B2FFF12F9E76EA2F8E9" pageId="3" pageNumber="66" type="conservation">
<paragraph id="8B3EF938505C2B2FFF12F9E76EA2F8E9" blockId="3.[136,1452,1191,1866]" pageId="3" pageNumber="66">
<emphasis id="B9F5252A505C2B2FFF12F9E76C19F999" bold="true" box="[189,584,1623,1649]" pageId="3" pageNumber="66">Preliminary conservation status:</emphasis>
—Applying the IUCN Red List Categories and Criteria and considering the area where the two populations of
<taxonomicName id="4C8182BB505C2B2FFDBCF9CB6C8FF97D" box="[531,734,1659,1685]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="tiswatl">
<emphasis id="B9F5252A505C2B2FFDBCF9CB6C8FF97D" box="[531,734,1659,1685]" italics="true" pageId="3" pageNumber="66">Anthurium tiswatl</emphasis>
</taxonomicName>
were found, in an area of about
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and on private property, with fragments of cleared forest, this species could be considered as Critically Endangered ([CR B2ab(ii,iii,iv,v)]) (
<bibRefCitation id="EF1084C9505C2B2FFF3FF9736F48F935" author="IUCN &amp; Cambridge, UK" box="[144,281,1731,1757]" pageId="3" pageNumber="66" refId="ref3671" refString="IUCN (2012) The IUCN Red List categories and criteria: Version 3.1, 2 nd ed. IUCN Species Survival Commission. Gland, Switzerland and Cambridge, UK. Available from: https: // portals. iucn. org / library / sites / library / files / documents / L- 2001 - 001 - 2 nd-Es. pdf (accessed 21 January 2024)" type="book" year="2012">IUCN 2012</bibRefCitation>
), but further studies are needed, since the species could be found in a larger area in the central region of
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.
</paragraph>
</subSubSection>
<subSubSection id="C39BAAB3505C2B2FFF12F8BB6BCAF8A1" pageId="3" pageNumber="66" type="materials_examined">
<paragraph id="8B3EF938505C2B2FFF12F8BB6BCAF8A1" blockId="3.[136,1452,1191,1866]" pageId="3" pageNumber="66">
<emphasis id="B9F5252A505C2B2FFF12F8BB6C90F8CD" bold="true" box="[189,705,1803,1829]" pageId="3" pageNumber="66">
Additional specimen examined (
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)
</emphasis>
:—
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.
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: Mun. Tequila, Tlaxacapitzco, Tecuanca,
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,
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,
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,
<date id="FF3FDFF8505C2B2FFDB2F89F6CFCF8A1" box="[541,685,1839,1865]" pageId="3" pageNumber="66" value="2022-04-06">6 April 2022</date>
,
<emphasis id="B9F5252A505C2B2FFD17F89F6B62F8A1" box="[696,1331,1839,1865]" italics="true" pageId="3" pageNumber="66">Pedro Díaz Jiménez &amp; Gabriela Citlahua Zepahua 1673</emphasis>
(HEM!).
</paragraph>
</subSubSection>
</treatment>
</document>