Add updates up until 2025-03-25 23:35:56

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<document id="2D4D3A4FCDBE0F2DA7F207728E062E21" ID-DOI="10.1016/j.jcz.2019.05.006" ID-Zenodo-Dep="15086123" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1742943580750" checkinUser="carolina" docAuthor="Zotto, Matteo Dal, Neuhaus, Birger, Yamasaki, Hiroshi &amp; Todaro, Antonio" docDate="2019" docId="03B487E6FFE7FF86FCEFFE60FA55F9A6" docLanguage="en" docName="ZoolAnz.282.206-231.pdf" docOrigin="Zoologischer Anzeiger 282" docTitle="Condyloderes multispinosus Higgins 1969" docType="treatment" docVersion="4" lastPageNumber="228" masterDocId="FF8DFF9EFFF4FF90FFD0FFCDF80DFF9F" masterDocTitle="The genus Condyloderes (Kinorhyncha: Cyclorhagida) in the Mediterranean Sea, including the description of two new species with novel characters" masterLastPageNumber="231" masterPageNumber="206" pageNumber="225" updateTime="1742945744644" updateUser="ExternalLinkService">
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<mods:title id="39502761FA8E793EED870470288CC1F9">The genus Condyloderes (Kinorhyncha: Cyclorhagida) in the Mediterranean Sea, including the description of two new species with novel characters</mods:title>
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<mods:namePart id="8D75319B3572C3CBA49247CDC7A11423">Zotto, Matteo Dal</mods:namePart>
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<mods:namePart id="B4C798B291F28AB28A264972E41E1221">Neuhaus, Birger</mods:namePart>
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<mods:namePart id="06E6525B8BE57902F8C4A9B2196E251F">Yamasaki, Hiroshi</mods:namePart>
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<mods:namePart id="524C6FD2690848380DE51CC88B7F33D4">Todaro, Antonio</mods:namePart>
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<mods:title id="C49C1323AC978F13FFB2C7110041BD10">Zoologischer Anzeiger</mods:title>
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<mods:date id="A8E8DF9D16D62816B87C812EA5010ABC">2019</mods:date>
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<mods:number id="BCDE0AF1A5264F19107E81661FFCDE1C">2019-06-05</mods:number>
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<emphasis id="B969EAE2FFE7FF83FCEFFE60FB6FFE5E" box="[831,866,429,449]" italics="true" pageId="19" pageNumber="225">3.4.</emphasis>
<taxonomicName id="4C1D4D73FFE7FF83FCBEFE63FDB3FE5E" ID-CoL="5ZRP2" authority="(McIntyre, 1962) Higgins, 1969" authorityName="Higgins" authorityYear="1969" baseAuthorityName="McIntyre" baseAuthorityYear="1962" box="[878,1470,429,449]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="19" pageNumber="225" phylum="Kinorhyncha" rank="species" species="multispinosus">
Condyloderes multispinosus
<emphasis id="B969EAE2FFE7FF83FB43FE60FDB3FE5E" box="[1171,1470,429,449]" italics="true" pageId="19" pageNumber="225">(McIntyre, 1962) Higgins, 1969</emphasis>
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<th id="76FC5DCEFFE7006FFF51FE77F8D4FE53" box="[129,217,442,460]" gridcol="0" gridrow="0" pageId="19" pageNumber="225">Specimen</th>
<th id="76FC5DCEFFE7006FFED0FE77F94AFE53" box="[256,327,442,460]" gridcol="1" gridrow="0" pageId="19" pageNumber="225"></th>
<th id="76FC5DCEFFE7006FFEBEFE77F9B8FE53" box="[366,437,442,460]" gridcol="2" gridrow="0" pageId="19" pageNumber="225"></th>
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<th id="76FC5DCEFFE7006FFD99FE77FA9DFE53" box="[585,656,442,460]" gridcol="4" gridrow="0" pageId="19" pageNumber="225"></th>
<th id="76FC5DCEFFE7006FFD66FE77FAF3FE53" box="[694,766,442,460]" gridcol="5" gridrow="0" pageId="19" pageNumber="225">♂1</th>
</tr>
<tr id="352D34B2FFE7006FFF51FE18FAF3FE7C" box="[129,766,469,483]" gridrow="1" pageId="19" pageNumber="225" rowspan-0="1" rowspan-5="1">
<td id="76FC5DCEFFE7006FFED0FE18F94AFE7C" box="[256,327,469,483]" gridcol="1" gridrow="1" pageId="19" pageNumber="225">1562541</td>
<td id="76FC5DCEFFE7006FFEBEFE18F9B8FE7C" box="[366,437,469,483]" gridcol="2" gridrow="1" pageId="19" pageNumber="225">1562542</td>
<td id="76FC5DCEFFE7006FFE0BFE18FA2FFE7C" box="[475,546,469,483]" gridcol="3" gridrow="1" pageId="19" pageNumber="225">1562543</td>
<td id="76FC5DCEFFE7006FFD99FE18FA9DFE7C" box="[585,656,469,483]" gridcol="4" gridrow="1" pageId="19" pageNumber="225">1562540</td>
</tr>
<tr id="352D34B2FFE7006FFF51FE35FAF3FD99" box="[129,766,504,518]" gridrow="2" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FE35F8D4FD99" box="[129,217,504,518]" gridcol="0" gridrow="2" pageId="19" pageNumber="225">Location</th>
<td id="76FC5DCEFFE7006FFED0FE35F94AFD99" box="[256,327,504,518]" gridcol="1" gridrow="2" pageId="19" pageNumber="225">Banyuls</td>
<td id="76FC5DCEFFE7006FFEBEFE35F9B8FD99" box="[366,437,504,518]" gridcol="2" gridrow="2" pageId="19" pageNumber="225">Banyuls</td>
<td id="76FC5DCEFFE7006FFE0BFE35FA2FFD99" box="[475,546,504,518]" gridcol="3" gridrow="2" pageId="19" pageNumber="225">Banyuls</td>
<td id="76FC5DCEFFE7006FFD99FE35FA9DFD99" box="[585,656,504,518]" gridcol="4" gridrow="2" pageId="19" pageNumber="225">Rovinj</td>
<td id="76FC5DCEFFE7006FFD66FE35FAF3FD99" box="[694,766,504,518]" gridcol="5" gridrow="2" pageId="19" pageNumber="225">Trappeto</td>
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<tr id="352D34B2FFE7006FFF51FDC2FAF3FD82" box="[129,766,527,541]" gridrow="3" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FDC2FAF3FD82" box="[129,766,527,541]" colspan="6" colspanRight="5" gridcol="0" gridrow="3" pageId="19" pageNumber="225">Character</th>
</tr>
<tr id="352D34B2FFE7006FFF51FDFFFAF3FDDF" box="[129,766,562,576]" gridrow="4" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FDFFF8D4FDDF" box="[129,217,562,576]" gridcol="0" gridrow="4" pageId="19" pageNumber="225">tl</th>
<td id="76FC5DCEFFE7006FFED0FDFFF94AFDDF" box="[256,327,562,576]" gridcol="1" gridrow="4" pageId="19" pageNumber="225">246</td>
<td id="76FC5DCEFFE7006FFEBEFDFFF9B8FDDF" box="[366,437,562,576]" gridcol="2" gridrow="4" pageId="19" pageNumber="225">272</td>
<td id="76FC5DCEFFE7006FFE0BFDFFFA2FFDDF" box="[475,546,562,576]" gridcol="3" gridrow="4" pageId="19" pageNumber="225">220</td>
<td id="76FC5DCEFFE7006FFD99FDFFFA9DFDDF" box="[585,656,562,576]" gridcol="4" gridrow="4" pageId="19" pageNumber="225">290</td>
<td id="76FC5DCEFFE7006FFD66FDFFFAF3FDDF" box="[694,766,562,576]" gridcol="5" gridrow="4" pageId="19" pageNumber="225">298</td>
</tr>
<tr id="352D34B2FFE7006FFF51FD85FAF3FDC8" box="[129,766,584,599]" gridrow="5" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FD85F8D4FDC8" box="[129,217,584,599]" gridcol="0" gridrow="5" pageId="19" pageNumber="225">s1</th>
<td id="76FC5DCEFFE7006FFED0FD85F94AFDC8" box="[256,327,584,599]" gridcol="1" gridrow="5" pageId="19" pageNumber="225">14</td>
<td id="76FC5DCEFFE7006FFEBEFD85F9B8FDC8" box="[366,437,584,599]" gridcol="2" gridrow="5" pageId="19" pageNumber="225">15</td>
<td id="76FC5DCEFFE7006FFE0BFD85FA2FFDC8" box="[475,546,584,599]" gridcol="3" gridrow="5" pageId="19" pageNumber="225">14</td>
<td id="76FC5DCEFFE7006FFD99FD85FA9DFDC8" box="[585,656,584,599]" gridcol="4" gridrow="5" pageId="19" pageNumber="225">15</td>
<td id="76FC5DCEFFE7006FFD66FD85FAF3FDC8" box="[694,766,584,599]" gridcol="5" gridrow="5" pageId="19" pageNumber="225">18</td>
</tr>
<tr id="352D34B2FFE7006FFF51FD92FAF3FDF2" box="[129,766,607,621]" gridrow="6" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FD92F8D4FDF2" box="[129,217,607,621]" gridcol="0" gridrow="6" pageId="19" pageNumber="225">s2</th>
<td id="76FC5DCEFFE7006FFED0FD92F94AFDF2" box="[256,327,607,621]" gridcol="1" gridrow="6" pageId="19" pageNumber="225">26</td>
<td id="76FC5DCEFFE7006FFEBEFD92F9B8FDF2" box="[366,437,607,621]" gridcol="2" gridrow="6" pageId="19" pageNumber="225">23</td>
<td id="76FC5DCEFFE7006FFE0BFD92FA2FFDF2" box="[475,546,607,621]" gridcol="3" gridrow="6" pageId="19" pageNumber="225">21</td>
<td id="76FC5DCEFFE7006FFD99FD92FA9DFDF2" box="[585,656,607,621]" gridcol="4" gridrow="6" pageId="19" pageNumber="225">26</td>
<td id="76FC5DCEFFE7006FFD66FD92FAF3FDF2" box="[694,766,607,621]" gridcol="5" gridrow="6" pageId="19" pageNumber="225">28</td>
</tr>
<tr id="352D34B2FFE7006FFF51FDBBFAF3FD1B" box="[129,766,630,644]" gridrow="7" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FDBBF8D4FD1B" box="[129,217,630,644]" gridcol="0" gridrow="7" pageId="19" pageNumber="225">s3</th>
<td id="76FC5DCEFFE7006FFED0FDBBF94AFD1B" box="[256,327,630,644]" gridcol="1" gridrow="7" pageId="19" pageNumber="225">28</td>
<td id="76FC5DCEFFE7006FFEBEFDBBF9B8FD1B" box="[366,437,630,644]" gridcol="2" gridrow="7" pageId="19" pageNumber="225">26</td>
<td id="76FC5DCEFFE7006FFE0BFDBBFA2FFD1B" box="[475,546,630,644]" gridcol="3" gridrow="7" pageId="19" pageNumber="225">26</td>
<td id="76FC5DCEFFE7006FFD99FDBBFA9DFD1B" box="[585,656,630,644]" gridcol="4" gridrow="7" pageId="19" pageNumber="225">28</td>
<td id="76FC5DCEFFE7006FFD66FDBBFAF3FD1B" box="[694,766,630,644]" gridcol="5" gridrow="7" pageId="19" pageNumber="225">26</td>
</tr>
<tr id="352D34B2FFE7006FFF51FD40FAF3FD04" box="[129,766,653,667]" gridrow="8" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FD40F8D4FD04" box="[129,217,653,667]" gridcol="0" gridrow="8" pageId="19" pageNumber="225">s4</th>
<td id="76FC5DCEFFE7006FFED0FD40F94AFD04" box="[256,327,653,667]" gridcol="1" gridrow="8" pageId="19" pageNumber="225">29</td>
<td id="76FC5DCEFFE7006FFEBEFD40F9B8FD04" box="[366,437,653,667]" gridcol="2" gridrow="8" pageId="19" pageNumber="225">29</td>
<td id="76FC5DCEFFE7006FFE0BFD40FA2FFD04" box="[475,546,653,667]" gridcol="3" gridrow="8" pageId="19" pageNumber="225">28</td>
<td id="76FC5DCEFFE7006FFD99FD40FA9DFD04" box="[585,656,653,667]" gridcol="4" gridrow="8" pageId="19" pageNumber="225">28</td>
<td id="76FC5DCEFFE7006FFD66FD40FAF3FD04" box="[694,766,653,667]" gridcol="5" gridrow="8" pageId="19" pageNumber="225">27</td>
</tr>
<tr id="352D34B2FFE7006FFF51FD69FAF3FD2D" box="[129,766,676,690]" gridrow="9" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FD69F8D4FD2D" box="[129,217,676,690]" gridcol="0" gridrow="9" pageId="19" pageNumber="225">s5</th>
<td id="76FC5DCEFFE7006FFED0FD69F94AFD2D" box="[256,327,676,690]" gridcol="1" gridrow="9" pageId="19" pageNumber="225">28</td>
<td id="76FC5DCEFFE7006FFEBEFD69F9B8FD2D" box="[366,437,676,690]" gridcol="2" gridrow="9" pageId="19" pageNumber="225">30</td>
<td id="76FC5DCEFFE7006FFE0BFD69FA2FFD2D" box="[475,546,676,690]" gridcol="3" gridrow="9" pageId="19" pageNumber="225">28</td>
<td id="76FC5DCEFFE7006FFD99FD69FA9DFD2D" box="[585,656,676,690]" gridcol="4" gridrow="9" pageId="19" pageNumber="225">31</td>
<td id="76FC5DCEFFE7006FFD66FD69FAF3FD2D" box="[694,766,676,690]" gridcol="5" gridrow="9" pageId="19" pageNumber="225">30</td>
</tr>
<tr id="352D34B2FFE7006FFF51FD76FAF3FD56" box="[129,766,699,713]" gridrow="10" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FD76F8D4FD56" box="[129,217,699,713]" gridcol="0" gridrow="10" pageId="19" pageNumber="225">s6</th>
<td id="76FC5DCEFFE7006FFED0FD76F94AFD56" box="[256,327,699,713]" gridcol="1" gridrow="10" pageId="19" pageNumber="225">29</td>
<td id="76FC5DCEFFE7006FFEBEFD76F9B8FD56" box="[366,437,699,713]" gridcol="2" gridrow="10" pageId="19" pageNumber="225">32</td>
<td id="76FC5DCEFFE7006FFE0BFD76FA2FFD56" box="[475,546,699,713]" gridcol="3" gridrow="10" pageId="19" pageNumber="225">27</td>
<td id="76FC5DCEFFE7006FFD99FD76FA9DFD56" box="[585,656,699,713]" gridcol="4" gridrow="10" pageId="19" pageNumber="225">31</td>
<td id="76FC5DCEFFE7006FFD66FD76FAF3FD56" box="[694,766,699,713]" gridcol="5" gridrow="10" pageId="19" pageNumber="225">33</td>
</tr>
<tr id="352D34B2FFE7006FFF51FD1FFAF3FD7F" box="[129,766,722,736]" gridrow="11" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FD1FF8D4FD7F" box="[129,217,722,736]" gridcol="0" gridrow="11" pageId="19" pageNumber="225">s7</th>
<td id="76FC5DCEFFE7006FFED0FD1FF94AFD7F" box="[256,327,722,736]" gridcol="1" gridrow="11" pageId="19" pageNumber="225">32</td>
<td id="76FC5DCEFFE7006FFEBEFD1FF9B8FD7F" box="[366,437,722,736]" gridcol="2" gridrow="11" pageId="19" pageNumber="225">33</td>
<td id="76FC5DCEFFE7006FFE0BFD1FFA2FFD7F" box="[475,546,722,736]" gridcol="3" gridrow="11" pageId="19" pageNumber="225">31</td>
<td id="76FC5DCEFFE7006FFD99FD1FFA9DFD7F" box="[585,656,722,736]" gridcol="4" gridrow="11" pageId="19" pageNumber="225">33</td>
<td id="76FC5DCEFFE7006FFD66FD1FFAF3FD7F" box="[694,766,722,736]" gridcol="5" gridrow="11" pageId="19" pageNumber="225">36</td>
</tr>
<tr id="352D34B2FFE7006FFF51FD25FAF3FD69" box="[129,766,744,758]" gridrow="12" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FD25F8D4FD69" box="[129,217,744,758]" gridcol="0" gridrow="12" pageId="19" pageNumber="225">s8</th>
<td id="76FC5DCEFFE7006FFED0FD25F94AFD69" box="[256,327,744,758]" gridcol="1" gridrow="12" pageId="19" pageNumber="225">33</td>
<td id="76FC5DCEFFE7006FFEBEFD25F9B8FD69" box="[366,437,744,758]" gridcol="2" gridrow="12" pageId="19" pageNumber="225">33</td>
<td id="76FC5DCEFFE7006FFE0BFD25FA2FFD69" box="[475,546,744,758]" gridcol="3" gridrow="12" pageId="19" pageNumber="225">29</td>
<td id="76FC5DCEFFE7006FFD99FD25FA9DFD69" box="[585,656,744,758]" gridcol="4" gridrow="12" pageId="19" pageNumber="225">33</td>
<td id="76FC5DCEFFE7006FFD66FD25FAF3FD69" box="[694,766,744,758]" gridcol="5" gridrow="12" pageId="19" pageNumber="225">35</td>
</tr>
<tr id="352D34B2FFE7006FFF51FD32FAF3FC92" box="[129,766,767,781]" gridrow="13" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FD32F8D4FC92" box="[129,217,767,781]" gridcol="0" gridrow="13" pageId="19" pageNumber="225">s9</th>
<td id="76FC5DCEFFE7006FFED0FD32F94AFC92" box="[256,327,767,781]" gridcol="1" gridrow="13" pageId="19" pageNumber="225">35</td>
<td id="76FC5DCEFFE7006FFEBEFD32F9B8FC92" box="[366,437,767,781]" gridcol="2" gridrow="13" pageId="19" pageNumber="225">35</td>
<td id="76FC5DCEFFE7006FFE0BFD32FA2FFC92" box="[475,546,767,781]" gridcol="3" gridrow="13" pageId="19" pageNumber="225">30</td>
<td id="76FC5DCEFFE7006FFD99FD32FA9DFC92" box="[585,656,767,781]" gridcol="4" gridrow="13" pageId="19" pageNumber="225">37</td>
<td id="76FC5DCEFFE7006FFD66FD32FAF3FC92" box="[694,766,767,781]" gridcol="5" gridrow="13" pageId="19" pageNumber="225">36</td>
</tr>
<tr id="352D34B2FFE7006FFF51FCDBFAF3FCBB" box="[129,766,790,804]" gridrow="14" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FCDBF8D4FCBB" box="[129,217,790,804]" gridcol="0" gridrow="14" pageId="19" pageNumber="225">s10</th>
<td id="76FC5DCEFFE7006FFED0FCDBF94AFCBB" box="[256,327,790,804]" gridcol="1" gridrow="14" pageId="19" pageNumber="225">28</td>
<td id="76FC5DCEFFE7006FFEBEFCDBF9B8FCBB" box="[366,437,790,804]" gridcol="2" gridrow="14" pageId="19" pageNumber="225">31</td>
<td id="76FC5DCEFFE7006FFE0BFCDBFA2FFCBB" box="[475,546,790,804]" gridcol="3" gridrow="14" pageId="19" pageNumber="225">24</td>
<td id="76FC5DCEFFE7006FFD99FCDBFA9DFCBB" box="[585,656,790,804]" gridcol="4" gridrow="14" pageId="19" pageNumber="225">31</td>
<td id="76FC5DCEFFE7006FFD66FCDBFAF3FCBB" box="[694,766,790,804]" gridcol="5" gridrow="14" pageId="19" pageNumber="225">32</td>
</tr>
<tr id="352D34B2FFE7006FFF51FCE0FAF3FCA4" box="[129,766,813,827]" gridrow="15" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FCE0F8D4FCA4" box="[129,217,813,827]" gridcol="0" gridrow="15" pageId="19" pageNumber="225">s11</th>
<td id="76FC5DCEFFE7006FFED0FCE0F94AFCA4" box="[256,327,813,827]" gridcol="1" gridrow="15" pageId="19" pageNumber="225">16</td>
<td id="76FC5DCEFFE7006FFEBEFCE0F9B8FCA4" box="[366,437,813,827]" gridcol="2" gridrow="15" pageId="19" pageNumber="225">18</td>
<td id="76FC5DCEFFE7006FFE0BFCE0FA2FFCA4" box="[475,546,813,827]" gridcol="3" gridrow="15" pageId="19" pageNumber="225">20</td>
<td id="76FC5DCEFFE7006FFD99FCE0FA9DFCA4" box="[585,656,813,827]" gridcol="4" gridrow="15" pageId="19" pageNumber="225">23</td>
<td id="76FC5DCEFFE7006FFD66FCE0FAF3FCA4" box="[694,766,813,827]" gridcol="5" gridrow="15" pageId="19" pageNumber="225">25</td>
</tr>
<tr id="352D34B2FFE7006FFF51FC9DFAF3FCC1" box="[129,766,848,862]" gridrow="16" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FC9DF8D4FCC1" box="[129,217,848,862]" gridcol="0" gridrow="16" pageId="19" pageNumber="225">sw-8</th>
<td id="76FC5DCEFFE7006FFED0FC9DF94AFCC1" box="[256,327,848,862]" gridcol="1" gridrow="16" pageId="19" pageNumber="225">68</td>
<td id="76FC5DCEFFE7006FFEBEFC9DF9B8FCC1" box="[366,437,848,862]" gridcol="2" gridrow="16" pageId="19" pageNumber="225">68</td>
<td id="76FC5DCEFFE7006FFE0BFC9DFA2FFCC1" box="[475,546,848,862]" gridcol="3" gridrow="16" pageId="19" pageNumber="225">n. a.</td>
<td id="76FC5DCEFFE7006FFD99FC9DFA9DFCC1" box="[585,656,848,862]" gridcol="4" gridrow="16" pageId="19" pageNumber="225">70</td>
<td id="76FC5DCEFFE7006FFD66FC9DFAF3FCC1" box="[694,766,848,862]" gridcol="5" gridrow="16" pageId="19" pageNumber="225">71</td>
</tr>
<tr id="352D34B2FFE7006FFF51FCAAFAF3FCEA" box="[129,766,871,885]" gridrow="17" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FCAAF8D4FCEA" box="[129,217,871,885]" gridcol="0" gridrow="17" pageId="19" pageNumber="225">sw-10</th>
<td id="76FC5DCEFFE7006FFED0FCAAF94AFCEA" box="[256,327,871,885]" gridcol="1" gridrow="17" pageId="19" pageNumber="225">55</td>
<td id="76FC5DCEFFE7006FFEBEFCAAF9B8FCEA" box="[366,437,871,885]" gridcol="2" gridrow="17" pageId="19" pageNumber="225">57</td>
<td id="76FC5DCEFFE7006FFE0BFCAAFA2FFCEA" box="[475,546,871,885]" gridcol="3" gridrow="17" pageId="19" pageNumber="225">n. a.</td>
<td id="76FC5DCEFFE7006FFD99FCAAFA9DFCEA" box="[585,656,871,885]" gridcol="4" gridrow="17" pageId="19" pageNumber="225">58</td>
<td id="76FC5DCEFFE7006FFD66FCAAFAF3FCEA" box="[694,766,871,885]" gridcol="5" gridrow="17" pageId="19" pageNumber="225">61</td>
</tr>
<tr id="352D34B2FFE7006FFF51FC47FAF3FC07" box="[129,766,906,920]" gridrow="18" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FC47F8D4FC07" box="[129,217,906,920]" gridcol="0" gridrow="18" pageId="19" pageNumber="225">md1 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FC47F94AFC07" box="[256,327,906,920]" gridcol="1" gridrow="18" pageId="19" pageNumber="225">37</td>
<td id="76FC5DCEFFE7006FFEBEFC47F9B8FC07" box="[366,437,906,920]" gridcol="2" gridrow="18" pageId="19" pageNumber="225">45</td>
<td id="76FC5DCEFFE7006FFE0BFC47FA2FFC07" box="[475,546,906,920]" gridcol="3" gridrow="18" pageId="19" pageNumber="225">38</td>
<td id="76FC5DCEFFE7006FFD99FC47FA9DFC07" box="[585,656,906,920]" gridcol="4" gridrow="18" pageId="19" pageNumber="225">41</td>
<td id="76FC5DCEFFE7006FFD66FC47FAF3FC07" box="[694,766,906,920]" gridcol="5" gridrow="18" pageId="19" pageNumber="225">33</td>
</tr>
<tr id="352D34B2FFE7006FFF51FC6CFAF3FC30" box="[129,766,929,943]" gridrow="19" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FC6CF8D4FC30" box="[129,217,929,943]" gridcol="0" gridrow="19" pageId="19" pageNumber="225">md2 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FC6CF94AFC30" box="[256,327,929,943]" gridcol="1" gridrow="19" pageId="19" pageNumber="225">40</td>
<td id="76FC5DCEFFE7006FFEBEFC6CF9B8FC30" box="[366,437,929,943]" gridcol="2" gridrow="19" pageId="19" pageNumber="225">34</td>
<td id="76FC5DCEFFE7006FFE0BFC6CFA2FFC30" box="[475,546,929,943]" gridcol="3" gridrow="19" pageId="19" pageNumber="225">45</td>
<td id="76FC5DCEFFE7006FFD99FC6CFA9DFC30" box="[585,656,929,943]" gridcol="4" gridrow="19" pageId="19" pageNumber="225">39</td>
<td id="76FC5DCEFFE7006FFD66FC6CFAF3FC30" box="[694,766,929,943]" gridcol="5" gridrow="19" pageId="19" pageNumber="225">30</td>
</tr>
<tr id="352D34B2FFE7006FFF51FC75FAF3FC59" box="[129,766,952,966]" gridrow="20" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FC75F8D4FC59" box="[129,217,952,966]" gridcol="0" gridrow="20" pageId="19" pageNumber="225">md3 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FC75F94AFC59" box="[256,327,952,966]" gridcol="1" gridrow="20" pageId="19" pageNumber="225">38</td>
<td id="76FC5DCEFFE7006FFEBEFC75F9B8FC59" box="[366,437,952,966]" gridcol="2" gridrow="20" pageId="19" pageNumber="225">39</td>
<td id="76FC5DCEFFE7006FFE0BFC75FA2FFC59" box="[475,546,952,966]" gridcol="3" gridrow="20" pageId="19" pageNumber="225">42</td>
<td id="76FC5DCEFFE7006FFD99FC75FA9DFC59" box="[585,656,952,966]" gridcol="4" gridrow="20" pageId="19" pageNumber="225">41</td>
<td id="76FC5DCEFFE7006FFD66FC75FAF3FC59" box="[694,766,952,966]" gridcol="5" gridrow="20" pageId="19" pageNumber="225">30</td>
</tr>
<tr id="352D34B2FFE7006FFF51FC03FAF3FC42" box="[129,766,974,989]" gridrow="21" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FC03F8D4FC42" box="[129,217,974,989]" gridcol="0" gridrow="21" pageId="19" pageNumber="225">md4 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FC03F94AFC42" box="[256,327,974,989]" gridcol="1" gridrow="21" pageId="19" pageNumber="225">40</td>
<td id="76FC5DCEFFE7006FFEBEFC03F9B8FC42" box="[366,437,974,989]" gridcol="2" gridrow="21" pageId="19" pageNumber="225">42</td>
<td id="76FC5DCEFFE7006FFE0BFC03FA2FFC42" box="[475,546,974,989]" gridcol="3" gridrow="21" pageId="19" pageNumber="225">45</td>
<td id="76FC5DCEFFE7006FFD99FC03FA9DFC42" box="[585,656,974,989]" gridcol="4" gridrow="21" pageId="19" pageNumber="225">38</td>
<td id="76FC5DCEFFE7006FFD66FC03FAF3FC42" box="[694,766,974,989]" gridcol="5" gridrow="21" pageId="19" pageNumber="225">32</td>
</tr>
<tr id="352D34B2FFE7006FFF51FC28FAF3FC6C" box="[129,766,997,1011]" gridrow="22" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FC28F8D4FC6C" box="[129,217,997,1011]" gridcol="0" gridrow="22" pageId="19" pageNumber="225">md5 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FC28F94AFC6C" box="[256,327,997,1011]" gridcol="1" gridrow="22" pageId="19" pageNumber="225">43</td>
<td id="76FC5DCEFFE7006FFEBEFC28F9B8FC6C" box="[366,437,997,1011]" gridcol="2" gridrow="22" pageId="19" pageNumber="225">45</td>
<td id="76FC5DCEFFE7006FFE0BFC28FA2FFC6C" box="[475,546,997,1011]" gridcol="3" gridrow="22" pageId="19" pageNumber="225">48</td>
<td id="76FC5DCEFFE7006FFD99FC28FA9DFC6C" box="[585,656,997,1011]" gridcol="4" gridrow="22" pageId="19" pageNumber="225">43</td>
<td id="76FC5DCEFFE7006FFD66FC28FAF3FC6C" box="[694,766,997,1011]" gridcol="5" gridrow="22" pageId="19" pageNumber="225">33</td>
</tr>
<tr id="352D34B2FFE7006FFF51FC36FAF3FB95" box="[129,766,1019,1034]" gridrow="23" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FC36F8D4FB95" box="[129,217,1019,1034]" gridcol="0" gridrow="23" pageId="19" pageNumber="225">md5 (cu)</th>
<td id="76FC5DCEFFE7006FFED0FC36F94AFB95" box="[256,327,1019,1034]" gridcol="1" gridrow="23" pageId="19" pageNumber="225">17</td>
<td id="76FC5DCEFFE7006FFEBEFC36F9B8FB95" box="[366,437,1019,1034]" gridcol="2" gridrow="23" pageId="19" pageNumber="225">n. a.</td>
<td id="76FC5DCEFFE7006FFE0BFC36FA2FFB95" box="[475,546,1019,1034]" gridcol="3" gridrow="23" pageId="19" pageNumber="225">18</td>
<td id="76FC5DCEFFE7006FFD99FC36FA9DFB95" box="[585,656,1019,1034]" gridcol="4" gridrow="23" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FD99FC36FA5BFB95" bold="true" box="[585,598,1019,1034]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFD66FC36FAF3FB95" box="[694,766,1019,1034]" gridcol="5" gridrow="23" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FD66FC36FACEFB95" bold="true" box="[694,707,1019,1034]" pageId="19" pageNumber="225"></emphasis>
</td>
</tr>
<tr id="352D34B2FFE7006FFF51FBDEFAF3FBBE" box="[129,766,1043,1057]" gridrow="24" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FBDEF8D4FBBE" box="[129,217,1043,1057]" gridcol="0" gridrow="24" pageId="19" pageNumber="225">md6 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FBDEF94AFBBE" box="[256,327,1043,1057]" gridcol="1" gridrow="24" pageId="19" pageNumber="225">43</td>
<td id="76FC5DCEFFE7006FFEBEFBDEF9B8FBBE" box="[366,437,1043,1057]" gridcol="2" gridrow="24" pageId="19" pageNumber="225">48</td>
<td id="76FC5DCEFFE7006FFE0BFBDEFA2FFBBE" box="[475,546,1043,1057]" gridcol="3" gridrow="24" pageId="19" pageNumber="225">50</td>
<td id="76FC5DCEFFE7006FFD99FBDEFA9DFBBE" box="[585,656,1043,1057]" gridcol="4" gridrow="24" pageId="19" pageNumber="225">46</td>
<td id="76FC5DCEFFE7006FFD66FBDEFAF3FBBE" box="[694,766,1043,1057]" gridcol="5" gridrow="24" pageId="19" pageNumber="225">n. a.</td>
</tr>
<tr id="352D34B2FFE7006FFF51FBE7FAF3FBA7" box="[129,766,1066,1080]" gridrow="25" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FBE7F8D4FBA7" box="[129,217,1066,1080]" gridcol="0" gridrow="25" pageId="19" pageNumber="225">md7 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FBE7F94AFBA7" box="[256,327,1066,1080]" gridcol="1" gridrow="25" pageId="19" pageNumber="225">43</td>
<td id="76FC5DCEFFE7006FFEBEFBE7F9B8FBA7" box="[366,437,1066,1080]" gridcol="2" gridrow="25" pageId="19" pageNumber="225">55</td>
<td id="76FC5DCEFFE7006FFE0BFBE7FA2FFBA7" box="[475,546,1066,1080]" gridcol="3" gridrow="25" pageId="19" pageNumber="225">50</td>
<td id="76FC5DCEFFE7006FFD99FBE7FA9DFBA7" box="[585,656,1066,1080]" gridcol="4" gridrow="25" pageId="19" pageNumber="225">44</td>
<td id="76FC5DCEFFE7006FFD66FBE7FAF3FBA7" box="[694,766,1066,1080]" gridcol="5" gridrow="25" pageId="19" pageNumber="225">43</td>
</tr>
<tr id="352D34B2FFE7006FFF51FB8DFAF3FBD0" box="[129,766,1088,1103]" gridrow="26" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FB8DF8D4FBD0" box="[129,217,1088,1103]" gridcol="0" gridrow="26" pageId="19" pageNumber="225">md7 (cu)</th>
<td id="76FC5DCEFFE7006FFED0FB8DF94AFBD0" box="[256,327,1088,1103]" gridcol="1" gridrow="26" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FED0FB8DF900FBD0" bold="true" box="[256,269,1088,1103]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFEBEFB8DF9B8FBD0" box="[366,437,1088,1103]" gridcol="2" gridrow="26" pageId="19" pageNumber="225">n. a.</td>
<td id="76FC5DCEFFE7006FFE0BFB8DFA2FFBD0" box="[475,546,1088,1103]" gridcol="3" gridrow="26" pageId="19" pageNumber="225">18</td>
<td id="76FC5DCEFFE7006FFD99FB8DFA9DFBD0" box="[585,656,1088,1103]" gridcol="4" gridrow="26" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FD99FB8DFA5BFBD0" bold="true" box="[585,598,1088,1103]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFD66FB8DFAF3FBD0" box="[694,766,1088,1103]" gridcol="5" gridrow="26" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FD66FB8DFACEFBD0" bold="true" box="[694,707,1088,1103]" pageId="19" pageNumber="225"></emphasis>
</td>
</tr>
<tr id="352D34B2FFE7006FFF51FB95FAF3FBF9" box="[129,766,1112,1126]" gridrow="27" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FB95F8D4FBF9" box="[129,217,1112,1126]" gridcol="0" gridrow="27" pageId="19" pageNumber="225">md8 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FB95F94AFBF9" box="[256,327,1112,1126]" gridcol="1" gridrow="27" pageId="19" pageNumber="225">53</td>
<td id="76FC5DCEFFE7006FFEBEFB95F9B8FBF9" box="[366,437,1112,1126]" gridcol="2" gridrow="27" pageId="19" pageNumber="225">55</td>
<td id="76FC5DCEFFE7006FFE0BFB95FA2FFBF9" box="[475,546,1112,1126]" gridcol="3" gridrow="27" pageId="19" pageNumber="225">50</td>
<td id="76FC5DCEFFE7006FFD99FB95FA9DFBF9" box="[585,656,1112,1126]" gridcol="4" gridrow="27" pageId="19" pageNumber="225">52</td>
<td id="76FC5DCEFFE7006FFD66FB95FAF3FBF9" box="[694,766,1112,1126]" gridcol="5" gridrow="27" pageId="19" pageNumber="225">48</td>
</tr>
<tr id="352D34B2FFE7006FFF51FBA3FAF3FBE2" box="[129,766,1134,1149]" gridrow="28" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FBA3F8D4FBE2" box="[129,217,1134,1149]" gridcol="0" gridrow="28" pageId="19" pageNumber="225">md9 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FBA3F94AFBE2" box="[256,327,1134,1149]" gridcol="1" gridrow="28" pageId="19" pageNumber="225">57</td>
<td id="76FC5DCEFFE7006FFEBEFBA3F9B8FBE2" box="[366,437,1134,1149]" gridcol="2" gridrow="28" pageId="19" pageNumber="225">75</td>
<td id="76FC5DCEFFE7006FFE0BFBA3FA2FFBE2" box="[475,546,1134,1149]" gridcol="3" gridrow="28" pageId="19" pageNumber="225">65</td>
<td id="76FC5DCEFFE7006FFD99FBA3FA9DFBE2" box="[585,656,1134,1149]" gridcol="4" gridrow="28" pageId="19" pageNumber="225">63</td>
<td id="76FC5DCEFFE7006FFD66FBA3FAF3FBE2" box="[694,766,1134,1149]" gridcol="5" gridrow="28" pageId="19" pageNumber="225">49</td>
</tr>
<tr id="352D34B2FFE7006FFF51FB48FAF3FB0B" box="[129,766,1157,1172]" gridrow="29" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FB48F8D4FB0B" box="[129,217,1157,1172]" gridcol="0" gridrow="29" pageId="19" pageNumber="225">md10 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FB48F94AFB0B" box="[256,327,1157,1172]" gridcol="1" gridrow="29" pageId="19" pageNumber="225">40</td>
<td id="76FC5DCEFFE7006FFEBEFB48F9B8FB0B" box="[366,437,1157,1172]" gridcol="2" gridrow="29" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FEBEFB48F976FB0B" bold="true" box="[366,379,1157,1172]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFE0BFB48FA2FFB0B" box="[475,546,1157,1172]" gridcol="3" gridrow="29" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FE0BFB48F9E5FB0B" bold="true" box="[475,488,1157,1172]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFD99FB48FA9DFB0B" box="[585,656,1157,1172]" gridcol="4" gridrow="29" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FD99FB48FA5BFB0B" bold="true" box="[585,598,1157,1172]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFD66FB48FAF3FB0B" box="[694,766,1157,1172]" gridcol="5" gridrow="29" pageId="19" pageNumber="225">41</td>
</tr>
<tr id="352D34B2FFE7006FFF51FB56FAF3FB35" box="[129,766,1179,1194]" gridrow="30" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FB56F8D4FB35" box="[129,217,1179,1194]" gridcol="0" gridrow="30" pageId="19" pageNumber="225">ld10 (ac) ♂</th>
<td id="76FC5DCEFFE7006FFED0FB56F94AFB35" box="[256,327,1179,1194]" gridcol="1" gridrow="30" pageId="19" pageNumber="225">37/42</td>
<td id="76FC5DCEFFE7006FFEBEFB56F9B8FB35" box="[366,437,1179,1194]" gridcol="2" gridrow="30" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FEBEFB56F976FB35" bold="true" box="[366,379,1179,1194]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFE0BFB56FA2FFB35" box="[475,546,1179,1194]" gridcol="3" gridrow="30" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FE0BFB56F9E5FB35" bold="true" box="[475,488,1179,1194]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFD99FB56FA9DFB35" box="[585,656,1179,1194]" gridcol="4" gridrow="30" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FD99FB56FA5BFB35" bold="true" box="[585,598,1179,1194]" pageId="19" pageNumber="225"></emphasis>
</td>
<td id="76FC5DCEFFE7006FFD66FB56FAF3FB35" box="[694,766,1179,1194]" gridcol="5" gridrow="30" pageId="19" pageNumber="225">40/41</td>
</tr>
<tr id="352D34B2FFE7006FFF51FB72FAF3FB52" box="[129,766,1215,1229]" gridrow="31" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FB72F8D4FB52" box="[129,217,1215,1229]" gridcol="0" gridrow="31" pageId="19" pageNumber="225">lv1 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FB72F94AFB52" box="[256,327,1215,1229]" gridcol="1" gridrow="31" pageId="19" pageNumber="225">33/32</td>
<td id="76FC5DCEFFE7006FFEBEFB72F9B8FB52" box="[366,437,1215,1229]" gridcol="2" gridrow="31" pageId="19" pageNumber="225">37/38</td>
<td id="76FC5DCEFFE7006FFE0BFB72FA2FFB52" box="[475,546,1215,1229]" gridcol="3" gridrow="31" pageId="19" pageNumber="225">38/39</td>
<td id="76FC5DCEFFE7006FFD99FB72FA9DFB52" box="[585,656,1215,1229]" gridcol="4" gridrow="31" pageId="19" pageNumber="225">35/43</td>
<td id="76FC5DCEFFE7006FFD66FB72FAF3FB52" box="[694,766,1215,1229]" gridcol="5" gridrow="31" pageId="19" pageNumber="225">34/36</td>
</tr>
<tr id="352D34B2FFE7006FFF51FB1BFAF3FB7B" box="[129,766,1238,1252]" gridrow="32" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FB1BF8D4FB7B" box="[129,217,1238,1252]" gridcol="0" gridrow="32" pageId="19" pageNumber="225">vl2 (cu)</th>
<td id="76FC5DCEFFE7006FFED0FB1BF94AFB7B" box="[256,327,1238,1252]" gridcol="1" gridrow="32" pageId="19" pageNumber="225">13/n. a.</td>
<td id="76FC5DCEFFE7006FFEBEFB1BF9B8FB7B" box="[366,437,1238,1252]" gridcol="2" gridrow="32" pageId="19" pageNumber="225">15</td>
<td id="76FC5DCEFFE7006FFE0BFB1BFA2FFB7B" box="[475,546,1238,1252]" gridcol="3" gridrow="32" pageId="19" pageNumber="225">16/15</td>
<td id="76FC5DCEFFE7006FFD99FB1BFA9DFB7B" box="[585,656,1238,1252]" gridcol="4" gridrow="32" pageId="19" pageNumber="225">13/n. a.</td>
<td id="76FC5DCEFFE7006FFD66FB1BFAF3FB7B" box="[694,766,1238,1252]" gridcol="5" gridrow="32" pageId="19" pageNumber="225">10/n. a.</td>
</tr>
<tr id="352D34B2FFE7006FFF51FB20FAF3FB64" box="[129,766,1261,1275]" gridrow="33" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FB20F8D4FB64" box="[129,217,1261,1275]" gridcol="0" gridrow="33" pageId="19" pageNumber="225">lv2 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FB20F94AFB64" box="[256,327,1261,1275]" gridcol="1" gridrow="33" pageId="19" pageNumber="225">37/31</td>
<td id="76FC5DCEFFE7006FFEBEFB20F9B8FB64" box="[366,437,1261,1275]" gridcol="2" gridrow="33" pageId="19" pageNumber="225">41/39</td>
<td id="76FC5DCEFFE7006FFE0BFB20FA2FFB64" box="[475,546,1261,1275]" gridcol="3" gridrow="33" pageId="19" pageNumber="225">37/34</td>
<td id="76FC5DCEFFE7006FFD99FB20FA9DFB64" box="[585,656,1261,1275]" gridcol="4" gridrow="33" pageId="19" pageNumber="225">n. a./33</td>
<td id="76FC5DCEFFE7006FFD66FB20FAF3FB64" box="[694,766,1261,1275]" gridcol="5" gridrow="33" pageId="19" pageNumber="225">34/37</td>
</tr>
<tr id="352D34B2FFE7006FFF51FAC9FAF3FA8D" box="[129,766,1284,1298]" gridrow="34" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FAC9F8D4FA8D" box="[129,217,1284,1298]" gridcol="0" gridrow="34" pageId="19" pageNumber="225">lv3 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FAC9F94AFA8D" box="[256,327,1284,1298]" gridcol="1" gridrow="34" pageId="19" pageNumber="225">36/36</td>
<td id="76FC5DCEFFE7006FFEBEFAC9F9B8FA8D" box="[366,437,1284,1298]" gridcol="2" gridrow="34" pageId="19" pageNumber="225">38/41</td>
<td id="76FC5DCEFFE7006FFE0BFAC9FA2FFA8D" box="[475,546,1284,1298]" gridcol="3" gridrow="34" pageId="19" pageNumber="225">42/n. a.</td>
<td id="76FC5DCEFFE7006FFD99FAC9FA9DFA8D" box="[585,656,1284,1298]" gridcol="4" gridrow="34" pageId="19" pageNumber="225">34/36</td>
<td id="76FC5DCEFFE7006FFD66FAC9FAF3FA8D" box="[694,766,1284,1298]" gridcol="5" gridrow="34" pageId="19" pageNumber="225">41/40</td>
</tr>
<tr id="352D34B2FFE7006FFF51FAD7FAF3FAB6" box="[129,766,1306,1321]" gridrow="35" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FAD7F8D4FAB6" box="[129,217,1306,1321]" gridcol="0" gridrow="35" pageId="19" pageNumber="225">lv4 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FAD7F94AFAB6" box="[256,327,1306,1321]" gridcol="1" gridrow="35" pageId="19" pageNumber="225">36/35</td>
<td id="76FC5DCEFFE7006FFEBEFAD7F9B8FAB6" box="[366,437,1306,1321]" gridcol="2" gridrow="35" pageId="19" pageNumber="225">n. a./41</td>
<td id="76FC5DCEFFE7006FFE0BFAD7FA2FFAB6" box="[475,546,1306,1321]" gridcol="3" gridrow="35" pageId="19" pageNumber="225">39/34</td>
<td id="76FC5DCEFFE7006FFD99FAD7FA9DFAB6" box="[585,656,1306,1321]" gridcol="4" gridrow="35" pageId="19" pageNumber="225">37/40</td>
<td id="76FC5DCEFFE7006FFD66FAD7FAF3FAB6" box="[694,766,1306,1321]" gridcol="5" gridrow="35" pageId="19" pageNumber="225">42/40</td>
</tr>
<tr id="352D34B2FFE7006FFF51FAFCFAF3FADF" box="[129,766,1329,1344]" gridrow="36" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FAFCF8D4FADF" box="[129,217,1329,1344]" gridcol="0" gridrow="36" pageId="19" pageNumber="225">vl5 (cu)</th>
<td id="76FC5DCEFFE7006FFED0FAFCF94AFADF" box="[256,327,1329,1344]" gridcol="1" gridrow="36" pageId="19" pageNumber="225">16/15</td>
<td id="76FC5DCEFFE7006FFEBEFAFCF9B8FADF" box="[366,437,1329,1344]" gridcol="2" gridrow="36" pageId="19" pageNumber="225">n. a.</td>
<td id="76FC5DCEFFE7006FFE0BFAFCFA2FFADF" box="[475,546,1329,1344]" gridcol="3" gridrow="36" pageId="19" pageNumber="225">18</td>
<td id="76FC5DCEFFE7006FFD99FAFCFA9DFADF" box="[585,656,1329,1344]" gridcol="4" gridrow="36" pageId="19" pageNumber="225">16/n. a.</td>
<td id="76FC5DCEFFE7006FFD66FAFCFAF3FADF" box="[694,766,1329,1344]" gridcol="5" gridrow="36" pageId="19" pageNumber="225">15/18</td>
</tr>
<tr id="352D34B2FFE7006FFF51FA85FAF3FAC9" box="[129,766,1352,1366]" gridrow="37" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FA85F8D4FAC9" box="[129,217,1352,1366]" gridcol="0" gridrow="37" pageId="19" pageNumber="225">lv5 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FA85F94AFAC9" box="[256,327,1352,1366]" gridcol="1" gridrow="37" pageId="19" pageNumber="225">43/41</td>
<td id="76FC5DCEFFE7006FFEBEFA85F9B8FAC9" box="[366,437,1352,1366]" gridcol="2" gridrow="37" pageId="19" pageNumber="225">42/44</td>
<td id="76FC5DCEFFE7006FFE0BFA85FA2FFAC9" box="[475,546,1352,1366]" gridcol="3" gridrow="37" pageId="19" pageNumber="225">43/36</td>
<td id="76FC5DCEFFE7006FFD99FA85FA9DFAC9" box="[585,656,1352,1366]" gridcol="4" gridrow="37" pageId="19" pageNumber="225">43</td>
<td id="76FC5DCEFFE7006FFD66FA85FAF3FAC9" box="[694,766,1352,1366]" gridcol="5" gridrow="37" pageId="19" pageNumber="225">41/41</td>
</tr>
<tr id="352D34B2FFE7006FFF51FA92FAF3FAF2" box="[129,766,1375,1389]" gridrow="38" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FA92F8D4FAF2" box="[129,217,1375,1389]" gridcol="0" gridrow="38" pageId="19" pageNumber="225">lv6 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FA92F94AFAF2" box="[256,327,1375,1389]" gridcol="1" gridrow="38" pageId="19" pageNumber="225">43/44</td>
<td id="76FC5DCEFFE7006FFEBEFA92F9B8FAF2" box="[366,437,1375,1389]" gridcol="2" gridrow="38" pageId="19" pageNumber="225">46</td>
<td id="76FC5DCEFFE7006FFE0BFA92FA2FFAF2" box="[475,546,1375,1389]" gridcol="3" gridrow="38" pageId="19" pageNumber="225">43</td>
<td id="76FC5DCEFFE7006FFD99FA92FA9DFAF2" box="[585,656,1375,1389]" gridcol="4" gridrow="38" pageId="19" pageNumber="225">45/48</td>
<td id="76FC5DCEFFE7006FFD66FA92FAF3FAF2" box="[694,766,1375,1389]" gridcol="5" gridrow="38" pageId="19" pageNumber="225">n. a./43</td>
</tr>
<tr id="352D34B2FFE7006FFF51FABBFAF3FA1B" box="[129,766,1398,1412]" gridrow="39" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FABBF8D4FA1B" box="[129,217,1398,1412]" gridcol="0" gridrow="39" pageId="19" pageNumber="225">lv7 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FABBF94AFA1B" box="[256,327,1398,1412]" gridcol="1" gridrow="39" pageId="19" pageNumber="225">45/42</td>
<td id="76FC5DCEFFE7006FFEBEFABBF9B8FA1B" box="[366,437,1398,1412]" gridcol="2" gridrow="39" pageId="19" pageNumber="225">45/48</td>
<td id="76FC5DCEFFE7006FFE0BFABBFA2FFA1B" box="[475,546,1398,1412]" gridcol="3" gridrow="39" pageId="19" pageNumber="225">46/45</td>
<td id="76FC5DCEFFE7006FFD99FABBFA9DFA1B" box="[585,656,1398,1412]" gridcol="4" gridrow="39" pageId="19" pageNumber="225">45/47</td>
<td id="76FC5DCEFFE7006FFD66FABBFAF3FA1B" box="[694,766,1398,1412]" gridcol="5" gridrow="39" pageId="19" pageNumber="225">44</td>
</tr>
<tr id="352D34B2FFE7006FFF51FA40FAF3FA04" box="[129,766,1421,1435]" gridrow="40" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FA40F8D4FA04" box="[129,217,1421,1435]" gridcol="0" gridrow="40" pageId="19" pageNumber="225">lv8 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FA40F94AFA04" box="[256,327,1421,1435]" gridcol="1" gridrow="40" pageId="19" pageNumber="225">48/44</td>
<td id="76FC5DCEFFE7006FFEBEFA40F9B8FA04" box="[366,437,1421,1435]" gridcol="2" gridrow="40" pageId="19" pageNumber="225">52/54</td>
<td id="76FC5DCEFFE7006FFE0BFA40FA2FFA04" box="[475,546,1421,1435]" gridcol="3" gridrow="40" pageId="19" pageNumber="225">36/48</td>
<td id="76FC5DCEFFE7006FFD99FA40FA9DFA04" box="[585,656,1421,1435]" gridcol="4" gridrow="40" pageId="19" pageNumber="225">48/50</td>
<td id="76FC5DCEFFE7006FFD66FA40FAF3FA04" box="[694,766,1421,1435]" gridcol="5" gridrow="40" pageId="19" pageNumber="225">46/44</td>
</tr>
<tr id="352D34B2FFE7006FFF51FA69FAF3FA2D" box="[129,766,1444,1458]" gridrow="41" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FA69F8D4FA2D" box="[129,217,1444,1458]" gridcol="0" gridrow="41" pageId="19" pageNumber="225">la8 (cu)</th>
<td id="76FC5DCEFFE7006FFED0FA69F94AFA2D" box="[256,327,1444,1458]" gridcol="1" gridrow="41" pageId="19" pageNumber="225">21/20</td>
<td id="76FC5DCEFFE7006FFEBEFA69F9B8FA2D" box="[366,437,1444,1458]" gridcol="2" gridrow="41" pageId="19" pageNumber="225">21/23</td>
<td id="76FC5DCEFFE7006FFE0BFA69FA2FFA2D" box="[475,546,1444,1458]" gridcol="3" gridrow="41" pageId="19" pageNumber="225">25</td>
<td id="76FC5DCEFFE7006FFD99FA69FA9DFA2D" box="[585,656,1444,1458]" gridcol="4" gridrow="41" pageId="19" pageNumber="225">21/23</td>
<td id="76FC5DCEFFE7006FFD66FA69FAF3FA2D" box="[694,766,1444,1458]" gridcol="5" gridrow="41" pageId="19" pageNumber="225">23/23</td>
</tr>
<tr id="352D34B2FFE7006FFF51FA77FAF3FA56" box="[129,766,1466,1481]" gridrow="42" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FA77F8D4FA56" box="[129,217,1466,1481]" gridcol="0" gridrow="42" pageId="19" pageNumber="225">vl9 (cu)</th>
<td id="76FC5DCEFFE7006FFED0FA77F94AFA56" box="[256,327,1466,1481]" gridcol="1" gridrow="42" pageId="19" pageNumber="225">18/21</td>
<td id="76FC5DCEFFE7006FFEBEFA77F9B8FA56" box="[366,437,1466,1481]" gridcol="2" gridrow="42" pageId="19" pageNumber="225">18/19</td>
<td id="76FC5DCEFFE7006FFE0BFA77FA2FFA56" box="[475,546,1466,1481]" gridcol="3" gridrow="42" pageId="19" pageNumber="225">19/20</td>
<td id="76FC5DCEFFE7006FFD99FA77FA9DFA56" box="[585,656,1466,1481]" gridcol="4" gridrow="42" pageId="19" pageNumber="225">16/n. a.</td>
<td id="76FC5DCEFFE7006FFD66FA77FAF3FA56" box="[694,766,1466,1481]" gridcol="5" gridrow="42" pageId="19" pageNumber="225">21/n. a.</td>
</tr>
<tr id="352D34B2FFE7006FFF51FA1CFAF3FA40" box="[129,766,1489,1503]" gridrow="43" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FA1CF8D4FA40" box="[129,217,1489,1503]" gridcol="0" gridrow="43" pageId="19" pageNumber="225">lv9 (ac)</th>
<td id="76FC5DCEFFE7006FFED0FA1CF94AFA40" box="[256,327,1489,1503]" gridcol="1" gridrow="43" pageId="19" pageNumber="225">57/55</td>
<td id="76FC5DCEFFE7006FFEBEFA1CF9B8FA40" box="[366,437,1489,1503]" gridcol="2" gridrow="43" pageId="19" pageNumber="225">63/62</td>
<td id="76FC5DCEFFE7006FFE0BFA1CFA2FFA40" box="[475,546,1489,1503]" gridcol="3" gridrow="43" pageId="19" pageNumber="225">58/60</td>
<td id="76FC5DCEFFE7006FFD99FA1CFA9DFA40" box="[585,656,1489,1503]" gridcol="4" gridrow="43" pageId="19" pageNumber="225">58/60</td>
<td id="76FC5DCEFFE7006FFD66FA1CFAF3FA40" box="[694,766,1489,1503]" gridcol="5" gridrow="43" pageId="19" pageNumber="225">42/45</td>
</tr>
<tr id="352D34B2FFE7006FFF51FA25FAF3FA69" box="[129,766,1512,1526]" gridrow="44" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FA25F8D4FA69" box="[129,217,1512,1526]" gridcol="0" gridrow="44" pageId="19" pageNumber="225">ltas (ac)</th>
<td id="76FC5DCEFFE7006FFED0FA25F94AFA69" box="[256,327,1512,1526]" gridcol="1" gridrow="44" pageId="19" pageNumber="225">182/180</td>
<td id="76FC5DCEFFE7006FFEBEFA25F9B8FA69" box="[366,437,1512,1526]" gridcol="2" gridrow="44" pageId="19" pageNumber="225">166/171</td>
<td id="76FC5DCEFFE7006FFE0BFA25FA2FFA69" box="[475,546,1512,1526]" gridcol="3" gridrow="44" pageId="19" pageNumber="225">183/179</td>
<td id="76FC5DCEFFE7006FFD99FA25FA9DFA69" box="[585,656,1512,1526]" gridcol="4" gridrow="44" pageId="19" pageNumber="225">broken</td>
<td id="76FC5DCEFFE7006FFD66FA25FAF3FA69" box="[694,766,1512,1526]" gridcol="5" gridrow="44" pageId="19" pageNumber="225">180/181</td>
</tr>
<tr id="352D34B2FFE7006FFF51FA32FAF3F992" box="[129,766,1535,1549]" gridrow="45" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51FA32F8D4F992" box="[129,217,1535,1549]" gridcol="0" gridrow="45" pageId="19" pageNumber="225">mts (ac)</th>
<td id="76FC5DCEFFE7006FFED0FA32F94AF992" box="[256,327,1535,1549]" gridcol="1" gridrow="45" pageId="19" pageNumber="225">94</td>
<td id="76FC5DCEFFE7006FFEBEFA32F9B8F992" box="[366,437,1535,1549]" gridcol="2" gridrow="45" pageId="19" pageNumber="225">82</td>
<td id="76FC5DCEFFE7006FFE0BFA32FA2FF992" box="[475,546,1535,1549]" gridcol="3" gridrow="45" pageId="19" pageNumber="225">82</td>
<td id="76FC5DCEFFE7006FFD99FA32FA9DF992" box="[585,656,1535,1549]" gridcol="4" gridrow="45" pageId="19" pageNumber="225">80</td>
<td id="76FC5DCEFFE7006FFD66FA32FAF3F992" box="[694,766,1535,1549]" gridcol="5" gridrow="45" pageId="19" pageNumber="225">93</td>
</tr>
<tr id="352D34B2FFE7006FFF51F9DBFAF3F9BB" box="[129,766,1558,1572]" gridrow="46" pageId="19" pageNumber="225">
<th id="76FC5DCEFFE7006FFF51F9DBF8D4F9BB" box="[129,217,1558,1572]" gridcol="0" gridrow="46" pageId="19" pageNumber="225">mts/tl</th>
<td id="76FC5DCEFFE7006FFED0F9DBF94AF9BB" box="[256,327,1558,1572]" gridcol="1" gridrow="46" pageId="19" pageNumber="225">0.38%</td>
<td id="76FC5DCEFFE7006FFEBEF9DBF9B8F9BB" box="[366,437,1558,1572]" gridcol="2" gridrow="46" pageId="19" pageNumber="225">0.30%</td>
<td id="76FC5DCEFFE7006FFE0BF9DBFA2FF9BB" box="[475,546,1558,1572]" gridcol="3" gridrow="46" pageId="19" pageNumber="225">0.37%</td>
<td id="76FC5DCEFFE7006FFD99F9DBFA9DF9BB" box="[585,656,1558,1572]" gridcol="4" gridrow="46" pageId="19" pageNumber="225">0.28%</td>
<td id="76FC5DCEFFE7006FFD66F9DBFAF3F9BB" box="[694,766,1558,1572]" gridcol="5" gridrow="46" pageId="19" pageNumber="225">0.31%</td>
</tr>
</table>
</paragraph>
<subSubSection id="C307657BFFE7FF83FC8FFE2BFDD1FD47" pageId="19" pageNumber="225" type="reference_group">
<paragraph id="8BA236F0FFE7FF83FC8FFE2BFD45FE66" blockId="19.[831,1501,486,728]" box="[863,1352,486,505]" pageId="19" pageNumber="225">
(
<figureCitation id="13262A75FFE7FF83FCB7FE2BFBBDFE66" box="[871,944,486,505]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="19" pageNumber="225">Figs. 13</figureCitation>
and
<figureCitation id="13262A75FFE7FF83FC33FE2BFBF6FE66" box="[995,1019,486,505]" captionStart="Fig" captionStartId="21.[113,138,1401,1415]" captionTargetBox="[164,1449,178,1371]" captionTargetId="figure-287@21.[164,1449,178,1371]" captionTargetPageId="21" captionText="Fig.14. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A—C, H) and male USNM 1562541 (D, G) and female USNM 1562542 (E, F) from the Gulf of Lion, dorsal view, light micrographs. A—B, Anterior segments, left side (A), and right side (B); arrows mark tergal extensions of free flap. C, Lateral terminal accessory spine with two thin areas (arrows).D—E, Segments 6‾8 (D) and 5‾7 (E) with middorsal cuspidate spine (arrows) on segment 5 (D) and segments 5 and 7 (E). F—G, Segments 9‾11.H, Segment 9 with protonephridial openings (arrows). Arrowheads in A—B, E, and H mark type-6 sensory spots. Abbreviations: bpl, broad placid; epi, epizoic protozoan; ltas, lateral terminal accessory spine; ld, laterodorsal acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; pa, pachycyclus; s, seg ment; ssp3, type-3 sensory spot. Numbers, except for sensory-spot type, refer to the segment number. Scale bars: C = 20 Mm; G = 50 Mm, valid for A, B, and D—H." figureDoi="http://doi.org/10.5281/zenodo.15086151" httpUri="https://zenodo.org/record/15086151/files/figure.png" pageId="19" pageNumber="225">14</figureCitation>
;
<figureCitation id="13262A75FFE7FF83FBD9FE2BFC47FE66" box="[1033,1098,486,505]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="19" pageNumber="225">Fig. S3</figureCitation>
; Tables 1, 2, 5;
<tableCitation id="C69F034BFFE7FF83FB37FE2BFD36FE66" box="[1255,1339,486,505]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="19" pageNumber="225">TableS2</tableCitation>
).
</paragraph>
<paragraph id="8BA236F0FFE7FF83FC8FFDCCFD12FD8A" blockId="19.[831,1501,486,728]" box="[863,1311,513,533]" pageId="19" pageNumber="225">
<taxonomicName id="4C1D4D73FFE7FF83FC8FFDCCFC53FD8A" authorityName="McIntyre" authorityYear="1962" box="[863,1118,513,533]" class="Cyclorhagida" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Kentrorhagata" pageId="19" pageNumber="225" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE7FF83FC8FFDCCFC53FD8A" box="[863,1118,513,533]" italics="true" pageId="19" pageNumber="225">Centroderes multispinosus</emphasis>
</taxonomicName>
— McIntyre (1962).
</paragraph>
<paragraph id="8BA236F0FFE7FF83FC8FFDD0FD9FFDAE" blockId="19.[831,1501,486,728]" box="[863,1426,541,561]" pageId="19" pageNumber="225">
<taxonomicName id="4C1D4D73FFE7FF83FC8FFDD0FC61FDAE" authorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" authorityYear="2019" baseAuthorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" baseAuthorityYear="1969" box="[863,1132,541,561]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="19" pageNumber="225" phylum="Kinorhyncha" rank="species" species="multispinosus" status="comb. nov.">
<emphasis id="B969EAE2FFE7FF83FC8FFDD0FC61FDAE" box="[863,1132,541,561]" italics="true" pageId="19" pageNumber="225">Condyloderes multispinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A5799FFE7FF83FBA2FDD3FCEDFDAE" box="[1138,1248,542,561]" pageId="19" pageNumber="225" rank="species">comb. nov.</taxonomicNameLabel>
— Higgins (1969).
</paragraph>
<paragraph id="8BA236F0FFE7FF83FC8FFDF4FDB5FD22" blockId="19.[831,1501,486,728]" pageId="19" pageNumber="225">
<taxonomicName id="4C1D4D73FFE7FF83FC8FFDF4FC61FDD2" authorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" authorityYear="2019" baseAuthorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" baseAuthorityYear="1969" box="[863,1132,569,589]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="19" pageNumber="225" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE7FF83FC8FFDF4FC61FDD2" box="[863,1132,569,589]" italics="true" pageId="19" pageNumber="225">Condyloderes multispinosus</emphasis>
</taxonomicName>
— Higgins (1971, 1986, 1988), Moore (1973), Brown (1985), Huys &amp; Coomans (1989), Nebelsick (1993), Adrianov &amp; Malakhov (1994, 1999), Bamber (1997), Adrianov et al. (2002), Martorelli &amp; Higgins (2004), SØrensen &amp; Pardos (2008), SØrensen et al. (2010b), Neuhaus (2013), Neuhaus et al. (2019).
</paragraph>
<paragraph id="8BA236F0FFE7FF83FC8FFD09FDD1FD47" blockId="19.[831,1501,486,728]" box="[863,1500,708,728]" pageId="19" pageNumber="225">
<taxonomicName id="4C1D4D73FFE7FF83FC8FFD09FC1FFD47" box="[863,1042,708,728]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="19" pageNumber="225" phylum="Kinorhyncha" rank="species" species="undefined-2">
<emphasis id="B969EAE2FFE7FF83FC8FFD09FBEDFD47" box="[863,992,708,728]" italics="true" pageId="19" pageNumber="225">Condyloderes</emphasis>
sp. 2
</taxonomicName>
(Tyrrhenian Sea) — Dal Zotto &amp; Todaro (2016).
</paragraph>
</subSubSection>
<subSubSection id="C307657BFFE7FF83FCEFFD31FC48FABD" pageId="19" pageNumber="225" type="diagnosis">
<paragraph id="8BA236F0FFE7FF83FCEFFD31FC3BFC8F" blockId="19.[831,1501,764,1314]" box="[831,1078,764,784]" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FCEFFD31FC3BFC8F" box="[831,1078,764,784]" italics="true" pageId="19" pageNumber="225">3.4.1. Emended diagnosis</emphasis>
</paragraph>
<paragraph id="8BA236F0FFE7FF83FC8FFCD4FC48FABD" blockId="19.[831,1501,764,1314]" pageId="19" pageNumber="225">
Neck placids with condyles in one lateral row in narrower placids and three lateral rows in broader placids; each narrower placid with one basal condyle; each broader placid with two condyles both in apical, intermediate, and basal lateral row; midventral placid with three apical and three basal condyles and two in intermediate lateral row; acicular spine middorsally on segments 1
<emphasis id="B969EAE2FFE7FF83FC9BFC0DFB57FC4C" bold="true" box="[843,858,960,979]" pageId="19" pageNumber="225"></emphasis>
9 (on segment 10 in male only), midterminally on segment 11, laterodorsally on segment 10 in male only, lateroventrally on segments 1
<emphasis id="B969EAE2FFE7FF83FC46FC35FBA8FB94" bold="true" box="[918,933,1016,1035]" pageId="19" pageNumber="225"></emphasis>
9, and laterodorsally on segment 10; cuspidate spine ventrolaterally on segments 2, 5, and 9 and lateral accessorily on segment 8, sometimes also middorsally on segments 5 and/or 7; type-6 sensory spot ventromedially on segments 1 (slightly more paraventrally), 2
<emphasis id="B969EAE2FFE7FF83FC36FBA5FBFBFBE4" bold="true" box="[998,1014,1128,1147]" pageId="19" pageNumber="225"></emphasis>
4 and 6
<emphasis id="B969EAE2FFE7FF83FB95FBA5FC5BFBE4" bold="true" box="[1093,1110,1128,1147]" pageId="19" pageNumber="225"></emphasis>
9, lateroventrally on segment 10, sublaterally on segments 1
<emphasis id="B969EAE2FFE7FF83FBF5FB4EFC39FB09" bold="true" box="[1061,1076,1155,1174]" pageId="19" pageNumber="225"></emphasis>
3 and 5
<emphasis id="B969EAE2FFE7FF83FBADFB4EFC80FB09" bold="true" box="[1149,1165,1155,1174]" pageId="19" pageNumber="225"></emphasis>
9, midlaterally on segments 1 and 2, laterodorsally on segments 1
<emphasis id="B969EAE2FFE7FF83FBA8FB52FC8AFB2D" bold="true" box="[1144,1159,1183,1202]" pageId="19" pageNumber="225"></emphasis>
9, subdorsally on segment 10, and paradorsally on segments 1 (almost subdorsally), 2, and 4
<emphasis id="B969EAE2FFE7FF83FAAAFB76FD86FB51" bold="true" box="[1402,1419,1211,1230]" pageId="19" pageNumber="225"></emphasis>
9; gland cell outlet ventromedially on segment 10; ventromedial appendage on segments 7 and 8 and area of micropapillae ventromedially on segment 9 in female only.
</paragraph>
</subSubSection>
<subSubSection id="C307657BFFE7FF83FCEFFA8BFBF9F9A6" pageId="19" pageNumber="225" type="materials_examined">
<paragraph id="8BA236F0FFE7FF83FCEFFA8BFC3BFAC5" blockId="19.[831,1501,1350,1593]" box="[831,1078,1350,1370]" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FCEFFA8BFC3BFAC5" box="[831,1078,1350,1370]" italics="true" pageId="19" pageNumber="225">3.4.2. Examined material</emphasis>
</paragraph>
<paragraph id="8BA236F0FFE7FF83FC8FFAAEFBF9F9A6" blockId="19.[831,1501,1350,1593]" pageId="19" pageNumber="225">
<materialsCitation id="3B753CADFFE7FF83FC8FFAAEFC80FAEA" box="[863,1165,1378,1398]" collectionCode="USNM" location="off Banyuls-sur-Mer" pageId="19" pageNumber="225" specimenCode="USNM 1562541" specimenCount="1" specimenCount-juvenile="1" specimenCount-male="1">
<specimenCount id="9D1BFD79FFE7FF83FC8FFAAEFBCBFAE9" box="[863,966,1379,1398]" count="1" pageId="19" pageNumber="225" type="male">One male</specimenCount>
(
<specimenCode id="DBBB9E8BFFE7FF83FC0AFAAEFC8CFAEA" box="[986,1153,1378,1398]" pageId="19" pageNumber="225">USNM 1562541</specimenCode>
),
</materialsCitation>
<materialsCitation id="3B753CADFFE7FF83FB4AFAAEFC2FFA0E" collectionCode="USNM" location="off Rovinj" pageId="19" pageNumber="225" specimenCode="USNM 1562542, USNM 1562543" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D1BFD79FFE7FF83FB4AFAAEFD10FAE9" box="[1178,1309,1379,1398]" count="2" pageId="19" pageNumber="225" type="female">two females</specimenCount>
(
<specimenCode id="DBBB9E8BFFE7FF83FAE0FAAEFDD4FAEA" box="[1328,1497,1378,1398]" pageId="19" pageNumber="225">USNM 1562542</specimenCode>
,
<specimenCode id="DBBB9E8BFFE7FF83FCEFFAB2FBE8FA0E" box="[831,997,1406,1426]" pageId="19" pageNumber="225">USNM 1562543</specimenCode>
), and
</materialsCitation>
<materialsCitation id="3B753CADFFE7FF83FBFFFAB2FC65FA32" collectionCode="USNM" location="off Banyuls-sur-Mer" pageId="19" pageNumber="225" specimenCode="USNM 1562544" specimenCount="1" specimenCount-juvenile="1">
<specimenCount id="9D1BFD79FFE7FF83FBFFFAB2FCBFFA0E" box="[1071,1202,1406,1426]" count="1" pageId="19" pageNumber="225" type="juvenile">one juvenile</specimenCount>
(
<specimenCode id="DBBB9E8BFFE7FF83FB14FAB2FD60FA0E" box="[1220,1389,1406,1426]" pageId="19" pageNumber="225">USNM 1562544</specimenCode>
), from
<location id="8EC2602BFFE7FF83FA12FAB3FBEAFA32" LSID="urn:lsid:plazi:treatment:03B487E6FFE7FF86FCEFFE60FA55F9A6:8EC2602BFFE7FF83FA12FAB3FBEAFA32" name="off Banyuls-sur-Mer" pageId="19" pageNumber="225">off Banyuls-sur-Mer</location>
,
<location id="8EC2602BFFE7FF83FC3FFA57FC69FA32" LSID="urn:lsid:plazi:treatment:03B487E6FFE7FF86FCEFFE60FA55F9A6:8EC2602BFFE7FF83FC3FFA57FC69FA32" box="[1007,1124,1434,1453]" name="Gulf of Lion" pageId="19" pageNumber="225">Gulf of Lion</location>
.
</materialsCitation>
<materialsCitation id="3B753CADFFE7FF83FBBCFA57FC70FA56" collectionCode="USNM" location="off Rovinj" pageId="19" pageNumber="225" specimenCode="USNM 1562540" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1BFD79FFE7FF83FBBCFA57FCD3FA32" box="[1132,1246,1434,1453]" count="1" pageId="19" pageNumber="225" type="female">One female</specimenCount>
from
<location id="8EC2602BFFE7FF83FAC9FA57FD74FA32" LSID="urn:lsid:plazi:treatment:03B487E6FFE7FF86FCEFFE60FA55F9A6:8EC2602BFFE7FF83FAC9FA57FD74FA32" box="[1305,1401,1434,1453]" name="off Rovinj" pageId="19" pageNumber="225">off Rovinj</location>
,
<location id="8EC2602BFFE7FF83FA52FA57FBB7FA56" LSID="urn:lsid:plazi:treatment:03B487E6FFE7FF86FCEFFE60FA55F9A6:8EC2602BFFE7FF83FA52FA57FBB7FA56" name="northern Adriatic Sea" pageId="19" pageNumber="225">northern Adriatic Sea</location>
(
<specimenCode id="DBBB9E8BFFE7FF83FC1BFA7BFC7FFA56" box="[971,1138,1462,1481]" pageId="19" pageNumber="225">USNM 1562540</specimenCode>
).
</materialsCitation>
<materialsCitation id="3B753CADFFE7FF83FB56FA7BFD1BFA7A" pageId="19" pageNumber="225" specimenCount="1">
All specimens mounted for light microscopy (
<figureCitation id="13262A75FFE7FF83FC12FA1FFC00FA7A" box="[962,1037,1490,1509]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="19" pageNumber="225">Figs. 13</figureCitation>
and
<figureCitation id="13262A75FFE7FF83FB94FA1FFC51FA7A" box="[1092,1116,1490,1509]" captionStart="Fig" captionStartId="21.[113,138,1401,1415]" captionTargetBox="[164,1449,178,1371]" captionTargetId="figure-287@21.[164,1449,178,1371]" captionTargetPageId="21" captionText="Fig.14. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A—C, H) and male USNM 1562541 (D, G) and female USNM 1562542 (E, F) from the Gulf of Lion, dorsal view, light micrographs. A—B, Anterior segments, left side (A), and right side (B); arrows mark tergal extensions of free flap. C, Lateral terminal accessory spine with two thin areas (arrows).D—E, Segments 6‾8 (D) and 5‾7 (E) with middorsal cuspidate spine (arrows) on segment 5 (D) and segments 5 and 7 (E). F—G, Segments 9‾11.H, Segment 9 with protonephridial openings (arrows). Arrowheads in A—B, E, and H mark type-6 sensory spots. Abbreviations: bpl, broad placid; epi, epizoic protozoan; ltas, lateral terminal accessory spine; ld, laterodorsal acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; pa, pachycyclus; s, seg ment; ssp3, type-3 sensory spot. Numbers, except for sensory-spot type, refer to the segment number. Scale bars: C = 20 Mm; G = 50 Mm, valid for A, B, and D—H." figureDoi="http://doi.org/10.5281/zenodo.15086151" httpUri="https://zenodo.org/record/15086151/files/figure.png" pageId="19" pageNumber="225">14</figureCitation>
;
<figureCitation id="13262A75FFE7FF83FBBCFA1FFCA3FA7A" box="[1132,1198,1490,1509]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="19" pageNumber="225">Fig. S3</figureCitation>
; Table 5).
</materialsCitation>
<materialsCitation id="3B753CADFFE7FF83FACEFA1FFBF9F9A6" location="off Trappeto" pageId="19" pageNumber="225" specimenCount="1" specimenCount-male="1" stateProvince="Sicily">
<specimenCount id="9D1BFD79FFE7FF83FACEFA1FFD8DFA7A" box="[1310,1408,1490,1509]" count="1" pageId="19" pageNumber="225" type="male">One male</specimenCount>
from
<location id="8EC2602BFFE7FF83FA12FA1FFB97F99E" LSID="urn:lsid:plazi:treatment:03B487E6FFE7FF86FCEFFE60FA55F9A6:8EC2602BFFE7FF83FA12FA1FFB97F99E" name="off Trappeto" pageId="19" pageNumber="225" stateProvince="Sicily">off Trappeto</location>
,
<location id="8EC2602BFFE7FF83FC76FA23FC81F99E" LSID="urn:lsid:plazi:treatment:03B487E6FFE7FF86FCEFFE60FA55F9A6:8EC2602BFFE7FF83FC76FA23FC81F99E" box="[934,1164,1518,1537]" name="Gulf of Castellammare" pageId="19" pageNumber="225" stateProvince="Sicily">Gulf of Castellammare</location>
,
<collectingRegion id="49D9F812FFE7FF83FB49FA23FCC2F99E" box="[1177,1231,1518,1537]" pageId="19" pageNumber="225">Sicily</collectingRegion>
,
<location id="8EC2602BFFE7FF83FB0BFA23FDD4F99E" LSID="urn:lsid:plazi:treatment:03B487E6FFE7FF86FCEFFE60FA55F9A6:8EC2602BFFE7FF83FB0BFA23FDD4F99E" box="[1243,1497,1518,1537]" name="southern Tyrrhenian Sea" pageId="19" pageNumber="225" stateProvince="Sicily">southern Tyrrhenian Sea</location>
, lost during observation; thus only few photographs were available for this specimen.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C307657BFFE7FF86FCEFF991FA55F9A6" lastPageId="22" lastPageNumber="228" pageId="19" pageNumber="225" type="description">
<paragraph id="8BA236F0FFE7FF83FCEFF991FC13F9EF" blockId="19.[831,1501,1628,1984]" box="[831,1054,1628,1648]" pageId="19" pageNumber="225">
<emphasis id="B969EAE2FFE7FF83FCEFF991FC13F9EF" box="[831,1054,1628,1648]" italics="true" pageId="19" pageNumber="225">3.4.3. Brief description</emphasis>
</paragraph>
<paragraph id="8BA236F0FFE7FF83FC8FF9B4FC6CF887" blockId="19.[831,1501,1628,1984]" pageId="19" pageNumber="225">
The morphology of the specimens from the Mediterranean Sea agrees widely with that of the recently re-described specimens of
<taxonomicName id="4C1D4D73FFE7FF83FCEFF97DFBE8F95B" authorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" authorityYear="2019" baseAuthorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" baseAuthorityYear="1969" box="[831,997,1712,1732]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="19" pageNumber="225" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE7FF83FCEFF97DFBE8F95B" box="[831,997,1712,1732]" italics="true" pageId="19" pageNumber="225">C. multispinosus</emphasis>
</taxonomicName>
from the North Sea, so only differences are mentioned for the Mediterranean specimens here. For illustrations, data and morphological measurements see
<figureCitation id="13262A75FFE7FF83FADEF924FD51F963" box="[1294,1372,1769,1788]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="19" pageNumber="225">Figs. 13</figureCitation>
,
<figureCitation id="13262A75FFE7FF83FABAF924FD8FF963" box="[1386,1410,1769,1788]" captionStart="Fig" captionStartId="21.[113,138,1401,1415]" captionTargetBox="[164,1449,178,1371]" captionTargetId="figure-287@21.[164,1449,178,1371]" captionTargetPageId="21" captionText="Fig.14. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A—C, H) and male USNM 1562541 (D, G) and female USNM 1562542 (E, F) from the Gulf of Lion, dorsal view, light micrographs. A—B, Anterior segments, left side (A), and right side (B); arrows mark tergal extensions of free flap. C, Lateral terminal accessory spine with two thin areas (arrows).D—E, Segments 6‾8 (D) and 5‾7 (E) with middorsal cuspidate spine (arrows) on segment 5 (D) and segments 5 and 7 (E). F—G, Segments 9‾11.H, Segment 9 with protonephridial openings (arrows). Arrowheads in A—B, E, and H mark type-6 sensory spots. Abbreviations: bpl, broad placid; epi, epizoic protozoan; ltas, lateral terminal accessory spine; ld, laterodorsal acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; pa, pachycyclus; s, seg ment; ssp3, type-3 sensory spot. Numbers, except for sensory-spot type, refer to the segment number. Scale bars: C = 20 Mm; G = 50 Mm, valid for A, B, and D—H." figureDoi="http://doi.org/10.5281/zenodo.15086151" httpUri="https://zenodo.org/record/15086151/files/figure.png" pageId="19" pageNumber="225">14</figureCitation>
,
<figureCitation id="13262A75FFE7FF83FA41F924FDDAF963" box="[1425,1495,1769,1788]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="19" pageNumber="225">Fig. S3</figureCitation>
,
<tableCitation id="C69F034BFFE7FF83FCEFF8C8FB9CF887" box="[831,913,1797,1816]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="19" pageNumber="225">Tables 2</tableCitation>
and 5, and
<tableCitation id="C69F034BFFE7FF83FBD8F8C8FC51F887" box="[1032,1116,1797,1816]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="19" pageNumber="225">Table S2</tableCitation>
.
</paragraph>
<paragraph id="8BA236F0FFE7FF84FC8FF8ECFB87F824" blockId="19.[831,1501,1628,1984]" lastBlockId="20.[805,1475,1848,1979]" lastPageId="20" lastPageNumber="226" pageId="19" pageNumber="225">
Our findings report for the first time a female adult stage for
<emphasis id="B969EAE2FFE7FF83FCEFF8F1FBEFF8CF" box="[831,994,1852,1872]" italics="true" pageId="19" pageNumber="225">
<taxonomicName id="4C1D4D73FFE7FF83FCEFF8F1FBD3F8CF" authorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" authorityYear="2019" baseAuthorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" baseAuthorityYear="1969" box="[831,990,1852,1872]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="19" pageNumber="225" phylum="Kinorhyncha" rank="species" species="multispinosus">C. multispinosus</taxonomicName>
.
</emphasis>
The female exhibits a ventromedial appendage on each sternal plate of segments 7 and 8, which gives the impression of an extremely elongated sensory spot covered by numerous cuticular micropapillae (
<figureCitation id="13262A75FFE7FF83FBE5F85DFC89F83B" box="[1077,1156,1936,1956]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="19" pageNumber="225">Fig. 13E</figureCitation>
). At its base, each structure shows a sclerotized cuticular duct penetrating the trunk cuticle. The duct seems to be surrounded by a cavity. The structures occur ventromedially on segment 7 and slightly more paraventrally on segment 8 (
<figureCitation id="13262A75FFE0FF84FF1EF8BDF910F81B" box="[206,285,1904,1924]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="20" pageNumber="226">Fig. 13E</figureCitation>
;
<tableCitation id="C69F034BFFE0FF84FEF7F8BDF975F81C" box="[295,376,1904,1923]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="20" pageNumber="226">Tables 2</tableCitation>
and 5;
<tableCitation id="C69F034BFFE0FF84FE13F8BDFA1BF81C" box="[451,534,1904,1923]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="20" pageNumber="226">Table S2</tableCitation>
). Each sternal plate of segment 9 possesses a more or less rectangular area with rounded edges and prominent cuticular micropapillae (
<figureCitation id="13262A75FFE0FF84FDF4F865FA7FF824" box="[548,626,1960,1979]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="20" pageNumber="226">Fig.13E</figureCitation>
;
<tableCitation id="C69F034BFFE0FF84FDAAF865FAC6F824" box="[634,715,1960,1979]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="20" pageNumber="226">Tables 2</tableCitation>
and 5;
<tableCitation id="C69F034BFFE0FF84FC90F8F5FB98F8D3" box="[832,917,1848,1868]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="20" pageNumber="226">Table S2</tableCitation>
). Pores and ducts penetrating the trunk cuticle do not seem to exist in this area. Alarge circular gonopore with a more sclerotized edge at its anterior margin and a less sclerotized edge at the posterior margin occurs on each sternal plate of segment 11 (
<figureCitation id="13262A75FFE0FF84FCFDF865FB73F824" box="[813,894,1960,1979]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="20" pageNumber="226">Fig. 13G</figureCitation>
).
</paragraph>
<caption id="DF626678FFE0FF84FF88F9B3FC89F961" ID-DOI="http://doi.org/10.5281/zenodo.15086149" ID-Zenodo-Dep="15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="20" pageNumber="226" startId="20.[88,113,1662,1676]" targetBox="[176,1386,178,1632]" targetPageId="20" targetType="figure">
<paragraph id="8BA236F0FFE0FF84FF88F9B3FC89F961" blockId="20.[88,1476,1660,1791]" pageId="20" pageNumber="226">
<emphasis id="B969EAE2FFE0FF84FF88F9B3F89DF913" bold="true" box="[88,144,1662,1676]" pageId="20" pageNumber="226">Fig. 13.</emphasis>
<taxonomicName id="4C1D4D73FFE0FF84FF48F9B1F965F913" authorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" authorityYear="2019" baseAuthorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" baseAuthorityYear="1969" box="[152,360,1660,1676]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="20" pageNumber="226" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE0FF84FF48F9B1F965F913" box="[152,360,1660,1676]" italics="true" pageId="20" pageNumber="226">Condyloderes multispinosus</emphasis>
</taxonomicName>
, female USNM 1562540 from the northern Adriatic Sea (
<emphasis id="B969EAE2FFE0FF84FCFAF9B3FB41F913" bold="true" box="[810,844,1661,1676]" pageId="20" pageNumber="226">A‾E</emphasis>
) and females USNM 1562543 (
<emphasis id="B969EAE2FFE0FF84FB92F9B3FC46F913" bold="true" box="[1090,1099,1662,1676]" pageId="20" pageNumber="226">F</emphasis>
) and USNM 1562542 (
<emphasis id="B969EAE2FFE0FF84FAD1F9B3FD01F913" bold="true" box="[1281,1292,1662,1676]" pageId="20" pageNumber="226">G</emphasis>
) from the Gulf of Lion, ventral view, light micrographs.
<emphasis id="B969EAE2FFE0FF84FE82F959F950F93D" bold="true" box="[338,349,1684,1698]" pageId="20" pageNumber="226">A</emphasis>
, Habitus.
<emphasis id="B969EAE2FFE0FF84FE7AF958F9B8F93C" bold="true" box="[426,437,1685,1699]" pageId="20" pageNumber="226">B</emphasis>
, Neck with placids.
<emphasis id="B969EAE2FFE0FF84FD9FF959FA7DF93C" bold="true" box="[591,624,1684,1699]" pageId="20" pageNumber="226">C—E</emphasis>
, Segments 1
<emphasis id="B969EAE2FFE0FF84FD02F959FAD3F93C" bold="true" box="[722,734,1684,1699]" pageId="20" pageNumber="226"></emphasis>
3 (
<emphasis id="B969EAE2FFE0FF84FD22F959FAF1F93D" bold="true" box="[754,764,1684,1698]" pageId="20" pageNumber="226">C</emphasis>
), 4
<emphasis id="B969EAE2FFE0FF84FCC5F959FB2FF93C" bold="true" box="[789,802,1684,1699]" pageId="20" pageNumber="226"></emphasis>
6 (
<emphasis id="B969EAE2FFE0FF84FCE6F958FB4FF93C" bold="true" box="[822,834,1685,1699]" pageId="20" pageNumber="226">D</emphasis>
), and 7
<emphasis id="B969EAE2FFE0FF84FCACF959FB84F93C" bold="true" box="[892,905,1684,1699]" pageId="20" pageNumber="226"></emphasis>
9 (
<emphasis id="B969EAE2FFE0FF84FC4DF958FBAAF93C" bold="true" box="[925,935,1685,1699]" pageId="20" pageNumber="226">E</emphasis>
). Arrows in
<emphasis id="B969EAE2FFE0FF84FBD6F959FC1DF93D" bold="true" box="[1030,1040,1684,1698]" pageId="20" pageNumber="226">C</emphasis>
point to spinose extensions of free flap.
<emphasis id="B969EAE2FFE0FF84FA99F958FD5CF93C" bold="true" box="[1353,1361,1685,1699]" pageId="20" pageNumber="226">F</emphasis>
, Sternal plates of segments 2
<emphasis id="B969EAE2FFE0FF84FF13F966F8DDF925" bold="true" box="[195,208,1707,1722]" pageId="20" pageNumber="226"></emphasis>
4; arrows mark paraventral area of micropapillae.
<emphasis id="B969EAE2FFE0FF84FD80F966FA56F926" bold="true" box="[592,603,1707,1721]" pageId="20" pageNumber="226">G</emphasis>
, Segments 9
<emphasis id="B969EAE2FFE0FF84FD6DF966FAC4F925" bold="true" box="[701,713,1707,1722]" pageId="20" pageNumber="226"></emphasis>
11 with gonad and gonopores. Arrowheads in
<emphasis id="B969EAE2FFE0FF84FBFBF966FC41F926" bold="true" box="[1067,1100,1707,1722]" pageId="20" pageNumber="226">C—E</emphasis>
mark type-6 sensory spots. Abbreviations:
<emphasis id="B969EAE2FFE0FF84FA4AF966FDB3F926" bold="true" box="[1434,1470,1707,1721]" pageId="20" pageNumber="226">amp</emphasis>
, female-specific area of micropapillae;
<emphasis id="B969EAE2FFE0FF84FEAEF90FF99FF94F" bold="true" box="[382,402,1730,1744]" pageId="20" pageNumber="226">ap</emphasis>
, female-specific appendage;
<emphasis id="B969EAE2FFE0FF84FDA2F90FFA81F94F" bold="true" box="[626,652,1730,1744]" pageId="20" pageNumber="226">bpl</emphasis>
, broad placid;
<emphasis id="B969EAE2FFE0FF84FD2FF90FFB02F94F" bold="true" box="[767,783,1730,1744]" pageId="20" pageNumber="226">fa</emphasis>
, filamentous appendage;
<emphasis id="B969EAE2FFE0FF84FC06F90FFBEEF94F" bold="true" box="[982,995,1730,1744]" pageId="20" pageNumber="226">ff</emphasis>
, free flap;
<emphasis id="B969EAE2FFE0FF84FBE4F90FFC4BF94F" bold="true" box="[1076,1094,1730,1744]" pageId="20" pageNumber="226">gc</emphasis>
, gland cell;
<emphasis id="B969EAE2FFE0FF84FB73F90FFCBAF94F" bold="true" box="[1187,1207,1730,1744]" pageId="20" pageNumber="226">go</emphasis>
, female gonad;
<emphasis id="B969EAE2FFE0FF84FAE1F90FFD48F94F" bold="true" box="[1329,1349,1730,1744]" pageId="20" pageNumber="226">gp</emphasis>
, gonopore;
<emphasis id="B969EAE2FFE0FF84FA71F90FFDB3F94F" bold="true" box="[1441,1470,1730,1744]" pageId="20" pageNumber="226">ltas</emphasis>
, lateral terminal accessory spine;
<emphasis id="B969EAE2FFE0FF84FE86F914F969F978" bold="true" box="[342,356,1753,1767]" pageId="20" pageNumber="226">lv</emphasis>
, lateroventral acicular spine;
<emphasis id="B969EAE2FFE0FF84FD98F914FA6AF978" bold="true" box="[584,615,1753,1767]" pageId="20" pageNumber="226">mts</emphasis>
, midterminal spine;
<emphasis id="B969EAE2FFE0FF84FCD9F914FB3FF978" bold="true" box="[777,818,1753,1767]" pageId="20" pageNumber="226">mvpl</emphasis>
, midventral placid;
<emphasis id="B969EAE2FFE0FF84FC1DF914FBE5F978" bold="true" box="[973,1000,1753,1767]" pageId="20" pageNumber="226">npl</emphasis>
, narrow placid;
<emphasis id="B969EAE2FFE0FF84FBB6F914FC77F978" bold="true" box="[1126,1146,1753,1767]" pageId="20" pageNumber="226">pa</emphasis>
, pachycyclus;
<emphasis id="B969EAE2FFE0FF84FB39F914FCFCF978" bold="true" box="[1257,1265,1753,1767]" pageId="20" pageNumber="226">s</emphasis>
, segment;
<emphasis id="B969EAE2FFE0FF84FA96F914FD59F978" bold="true" box="[1350,1364,1753,1767]" pageId="20" pageNumber="226">st</emphasis>
, sternal plate;
<emphasis id="B969EAE2FFE0FF84FF88F93DF877F961" bold="true" box="[88,122,1776,1790]" pageId="20" pageNumber="226">vlcu</emphasis>
, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100
<emphasis id="B969EAE2FFE0FF84FC91F920FB46F960" bold="true" box="[833,843,1773,1791]" pageId="20" pageNumber="226">M</emphasis>
m; E = 50
<emphasis id="B969EAE2FFE0FF84FC4AF920FBA9F960" bold="true" box="[922,932,1773,1791]" pageId="20" pageNumber="226">M</emphasis>
m, valid for B—G; F = 30
<emphasis id="B969EAE2FFE0FF84FBB5F920FC62F960" bold="true" box="[1125,1135,1773,1791]" pageId="20" pageNumber="226">M</emphasis>
m.
</paragraph>
</caption>
<caption id="DF626678FFE1FF85FFA1FAB4FC6CFA65" ID-DOI="http://doi.org/10.5281/zenodo.15086151" ID-Zenodo-Dep="15086151" httpUri="https://zenodo.org/record/15086151/files/figure.png" pageId="21" pageNumber="227" startId="21.[113,138,1401,1415]" targetBox="[164,1449,178,1371]" targetPageId="21" targetType="figure">
<paragraph id="8BA236F0FFE1FF85FFA1FAB4FC6CFA65" blockId="21.[113,1502,1399,1531]" pageId="21" pageNumber="227">
<emphasis id="B969EAE2FFE1FF85FFA1FAB4F8A5FA18" bold="true" box="[113,168,1401,1415]" pageId="21" pageNumber="227">Fig. 14.</emphasis>
<taxonomicName id="4C1D4D73FFE1FF85FF60FABAF972FA18" authorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" authorityYear="2019" baseAuthorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" baseAuthorityYear="1969" box="[176,383,1399,1415]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="21" pageNumber="227" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE1FF85FF60FABAF972FA18" box="[176,383,1399,1415]" italics="true" pageId="21" pageNumber="227">Condyloderes multispinosus</emphasis>
</taxonomicName>
, female USNM 1562540 from the northern Adriatic Sea (
<emphasis id="B969EAE2FFE1FF85FCE5FAB4FB5AFA18" bold="true" box="[821,855,1400,1415]" pageId="21" pageNumber="227">A—C</emphasis>
,
<emphasis id="B969EAE2FFE1FF85FCB0FAB4FB60FA18" bold="true" box="[864,877,1401,1415]" pageId="21" pageNumber="227">H</emphasis>
) and male USNM 1562541 (
<emphasis id="B969EAE2FFE1FF85FB95FAB4FC5CFA18" bold="true" box="[1093,1105,1401,1415]" pageId="21" pageNumber="227">D</emphasis>
,
<emphasis id="B969EAE2FFE1FF85FB8AFAB4FC68FA18" bold="true" box="[1114,1125,1401,1415]" pageId="21" pageNumber="227">G</emphasis>
) and female USNM 1562542 (
<emphasis id="B969EAE2FFE1FF85FA9EFAB4FD55FA18" bold="true" box="[1358,1368,1401,1415]" pageId="21" pageNumber="227">E</emphasis>
,
<emphasis id="B969EAE2FFE1FF85FAB0FAB4FD64FA18" bold="true" box="[1376,1385,1401,1415]" pageId="21" pageNumber="227">F</emphasis>
) from the Gulf of Lion, dorsal view, light micrographs.
<emphasis id="B969EAE2FFE1FF85FE73FA5DF9CBFA01" bold="true" box="[419,454,1423,1438]" pageId="21" pageNumber="227">A—B</emphasis>
, Anterior segments, left side (
<emphasis id="B969EAE2FFE1FF85FD60FA5DFAB6FA01" bold="true" box="[688,699,1424,1438]" pageId="21" pageNumber="227">A</emphasis>
), and right side (
<emphasis id="B969EAE2FFE1FF85FC90FA5DFB46FA01" bold="true" box="[832,843,1424,1438]" pageId="21" pageNumber="227">B</emphasis>
); arrows mark tergal extensions of free flap.
<emphasis id="B969EAE2FFE1FF85FB79FA5DFCBEFA01" bold="true" box="[1193,1203,1424,1438]" pageId="21" pageNumber="227">C</emphasis>
, Lateral terminal accessory spine with two thin areas (arrows).
<emphasis id="B969EAE2FFE1FF85FEFEFA6AF95CFA2A" bold="true" box="[302,337,1446,1461]" pageId="21" pageNumber="227">D—E</emphasis>
, Segments 6
<emphasis id="B969EAE2FFE1FF85FE63FA6BF9CDFA2A" bold="true" box="[435,448,1446,1461]" pageId="21" pageNumber="227"></emphasis>
8 (
<emphasis id="B969EAE2FFE1FF85FE04FA6AF9EDFA2A" bold="true" box="[468,480,1447,1461]" pageId="21" pageNumber="227">D</emphasis>
) and 5
<emphasis id="B969EAE2FFE1FF85FDC6FA6BFA2FFA2A" bold="true" box="[534,546,1446,1461]" pageId="21" pageNumber="227"></emphasis>
7 (
<emphasis id="B969EAE2FFE1FF85FDE6FA6AFA4DFA2A" bold="true" box="[566,576,1447,1461]" pageId="21" pageNumber="227">E</emphasis>
) with middorsal cuspidate spine (arrows) on segment 5 (
<emphasis id="B969EAE2FFE1FF85FC2AFA6AFC0BFA2A" bold="true" box="[1018,1030,1447,1461]" pageId="21" pageNumber="227">D</emphasis>
) and segments 5 and 7 (
<emphasis id="B969EAE2FFE1FF85FB13FA6AFCC0FA2A" bold="true" box="[1219,1229,1447,1461]" pageId="21" pageNumber="227">E</emphasis>
).
<emphasis id="B969EAE2FFE1FF85FB0CFA6AFCF0FA2B" bold="true" box="[1244,1277,1446,1461]" pageId="21" pageNumber="227">F—G</emphasis>
, Segments 9
<emphasis id="B969EAE2FFE1FF85FA8FFA6BFD66FA2A" bold="true" box="[1375,1387,1446,1461]" pageId="21" pageNumber="227"></emphasis>
11.
<emphasis id="B969EAE2FFE1FF85FA53FA6AFD9DFA2A" bold="true" box="[1411,1424,1447,1461]" pageId="21" pageNumber="227">H</emphasis>
, Segment 9 with protonephridial openings (arrows). Arrowheads in
<emphasis id="B969EAE2FFE1FF85FDE6FA70FA60FA53" bold="true" box="[566,621,1469,1484]" pageId="21" pageNumber="227">A—B, E</emphasis>
, and
<emphasis id="B969EAE2FFE1FF85FD48FA73FAA8FA53" bold="true" box="[664,677,1470,1484]" pageId="21" pageNumber="227">H</emphasis>
mark type-6 sensory spots. Abbreviations:
<emphasis id="B969EAE2FFE1FF85FC28FA73FC1FFA53" bold="true" box="[1016,1042,1470,1484]" pageId="21" pageNumber="227">bpl</emphasis>
, broad placid;
<emphasis id="B969EAE2FFE1FF85FB57FA70FCADFA54" bold="true" box="[1159,1184,1469,1483]" pageId="21" pageNumber="227">epi</emphasis>
, epizoic protozoan;
<emphasis id="B969EAE2FFE1FF85FAEEFA73FD56FA53" bold="true" box="[1342,1371,1470,1484]" pageId="21" pageNumber="227">ltas</emphasis>
, lateral terminal accessory spine;
<emphasis id="B969EAE2FFE1FF85FF24FA19F909FA7D" bold="true" box="[244,260,1492,1506]" pageId="21" pageNumber="227">ld</emphasis>
, laterodorsal acicular spine;
<emphasis id="B969EAE2FFE1FF85FE34FA19F9F2FA7D" bold="true" box="[484,511,1492,1506]" pageId="21" pageNumber="227">md</emphasis>
, middorsal acicular spine;
<emphasis id="B969EAE2FFE1FF85FD00FA19FAE2FA7D" bold="true" box="[720,751,1492,1506]" pageId="21" pageNumber="227">mts</emphasis>
, midterminal spine;
<emphasis id="B969EAE2FFE1FF85FC42FA19FBA0FA7D" bold="true" box="[914,941,1492,1506]" pageId="21" pageNumber="227">npl</emphasis>
, narrow placid;
<emphasis id="B969EAE2FFE1FF85FBFDFA19FC4CFA7D" bold="true" box="[1069,1089,1492,1506]" pageId="21" pageNumber="227">pa</emphasis>
, pachycyclus;
<emphasis id="B969EAE2FFE1FF85FB60FA19FCB5FA7D" bold="true" box="[1200,1208,1492,1506]" pageId="21" pageNumber="227">s</emphasis>
, seg ment;
<emphasis id="B969EAE2FFE1FF85FAC3FA19FD3BFA7D" bold="true" box="[1299,1334,1492,1506]" pageId="21" pageNumber="227">ssp3</emphasis>
, type-3 sensory spot. Numbers, except for sensory-spot type, refer to the segment number. Scale bars: C = 20
<emphasis id="B969EAE2FFE1FF85FCF6FA24FB3DFA64" bold="true" box="[806,816,1513,1531]" pageId="21" pageNumber="227">M</emphasis>
m; G = 50
<emphasis id="B969EAE2FFE1FF85FC51FA24FB86FA64" bold="true" box="[897,907,1513,1531]" pageId="21" pageNumber="227">M</emphasis>
m, valid for A, B, and D—H.
</paragraph>
</caption>
<paragraph id="8BA236F0FFE1FF85FFA1F98CF961F9CA" blockId="21.[113,783,1601,1984]" box="[113,364,1601,1621]" pageId="21" pageNumber="227">
<emphasis id="B969EAE2FFE1FF85FFA1F98CF961F9CA" box="[113,364,1601,1621]" italics="true" pageId="21" pageNumber="227">3.4.4. Sexual dimorphism</emphasis>
</paragraph>
<paragraph id="8BA236F0FFE1FF85FF41F993FDABF9EE" blockId="21.[113,783,1601,1984]" lastBlockId="21.[831,1500,1602,1649]" pageId="21" pageNumber="227">
Males can be distinguished from females by the lack of the ventromedial appendage on each sternal plate of segments 7 and 8
<emphasis id="B969EAE2FFE1FF85FFA1F959F8A3F937" box="[113,174,1684,1704]" italics="true" pageId="21" pageNumber="227">versus</emphasis>
its existence in females (
<figureCitation id="13262A75FFE1FF85FE61F958FA0CF936" box="[433,513,1685,1705]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="21" pageNumber="227">Fig. 13E</figureCitation>
;
<tableCitation id="C69F034BFFE1FF85FDDCF958FA52F937" box="[524,607,1685,1704]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="21" pageNumber="227">Tables 2</tableCitation>
and 5;
<tableCitation id="C69F034BFFE1FF85FD7EF958FB0FF937" box="[686,770,1685,1704]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="21" pageNumber="227">Table S2</tableCitation>
), the lack of a more or less rectangular area with prominent cuticular micropapillae on each sternal plate of segment 9
<emphasis id="B969EAE2FFE1FF85FDA8F901FAB8F97F" box="[632,693,1740,1760]" italics="true" pageId="21" pageNumber="227">versus</emphasis>
its existence in females (
<figureCitation id="13262A75FFE1FF85FEF0F924F963F962" box="[288,366,1769,1789]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="21" pageNumber="227">Fig.13E</figureCitation>
;
<tableCitation id="C69F034BFFE1FF85FEA6F924F9CAF963" box="[374,455,1769,1788]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="21" pageNumber="227">Tables 2</tableCitation>
and 5;
<tableCitation id="C69F034BFFE1FF85FDDFF924FA6CF963" box="[527,609,1769,1788]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="21" pageNumber="227">Table S2</tableCitation>
), the existence of a ventromedial gland cell outlet with a short sclerotized cuticular duct and an intra- and subcuticular cuticle-lined cavity on each sternal plate of segment 10
<emphasis id="B969EAE2FFE1FF85FE56F8F1F9CEF8CF" box="[390,451,1852,1872]" italics="true" pageId="21" pageNumber="227">versus</emphasis>
the lack of this outlet in females, the possession of a middorsal and a laterodorsal acicular spine on segment 10 thinner than on the remaining segments (
<figureCitation id="13262A75FFE1FF85FD64F8B8FB05F817" box="[692,776,1909,1928]" captionStart="Fig" captionStartId="21.[113,138,1401,1415]" captionTargetBox="[164,1449,178,1371]" captionTargetId="figure-287@21.[164,1449,178,1371]" captionTargetPageId="21" captionText="Fig.14. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A—C, H) and male USNM 1562541 (D, G) and female USNM 1562542 (E, F) from the Gulf of Lion, dorsal view, light micrographs. A—B, Anterior segments, left side (A), and right side (B); arrows mark tergal extensions of free flap. C, Lateral terminal accessory spine with two thin areas (arrows).D—E, Segments 6‾8 (D) and 5‾7 (E) with middorsal cuspidate spine (arrows) on segment 5 (D) and segments 5 and 7 (E). F—G, Segments 9‾11.H, Segment 9 with protonephridial openings (arrows). Arrowheads in A—B, E, and H mark type-6 sensory spots. Abbreviations: bpl, broad placid; epi, epizoic protozoan; ltas, lateral terminal accessory spine; ld, laterodorsal acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; pa, pachycyclus; s, seg ment; ssp3, type-3 sensory spot. Numbers, except for sensory-spot type, refer to the segment number. Scale bars: C = 20 Mm; G = 50 Mm, valid for A, B, and D—H." figureDoi="http://doi.org/10.5281/zenodo.15086151" httpUri="https://zenodo.org/record/15086151/files/figure.png" pageId="21" pageNumber="227">Fig. 14G</figureCitation>
)
<emphasis id="B969EAE2FFE1FF85FFA1F842F8A3F83C" box="[113,174,1935,1955]" italics="true" pageId="21" pageNumber="227">versus</emphasis>
the lack of these spines in females (
<figureCitation id="13262A75FFE1FF85FDCEF85DFA61F83B" box="[542,620,1936,1956]" captionStart="Fig" captionStartId="21.[113,138,1401,1415]" captionTargetBox="[164,1449,178,1371]" captionTargetId="figure-287@21.[164,1449,178,1371]" captionTargetPageId="21" captionText="Fig.14. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A—C, H) and male USNM 1562541 (D, G) and female USNM 1562542 (E, F) from the Gulf of Lion, dorsal view, light micrographs. A—B, Anterior segments, left side (A), and right side (B); arrows mark tergal extensions of free flap. C, Lateral terminal accessory spine with two thin areas (arrows).D—E, Segments 6‾8 (D) and 5‾7 (E) with middorsal cuspidate spine (arrows) on segment 5 (D) and segments 5 and 7 (E). F—G, Segments 9‾11.H, Segment 9 with protonephridial openings (arrows). Arrowheads in A—B, E, and H mark type-6 sensory spots. Abbreviations: bpl, broad placid; epi, epizoic protozoan; ltas, lateral terminal accessory spine; ld, laterodorsal acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; pa, pachycyclus; s, seg ment; ssp3, type-3 sensory spot. Numbers, except for sensory-spot type, refer to the segment number. Scale bars: C = 20 Mm; G = 50 Mm, valid for A, B, and D—H." figureDoi="http://doi.org/10.5281/zenodo.15086151" httpUri="https://zenodo.org/record/15086151/files/figure.png" pageId="21" pageNumber="227">Fig. 14F</figureCitation>
;
<tableCitation id="C69F034BFFE1FF85FDA6F85DFAC5F83B" box="[630,712,1936,1956]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="21" pageNumber="227">Tables 2</tableCitation>
and 5;
<tableCitation id="C69F034BFFE1FF85FFA1F861F8C7F85F" box="[113,202,1964,1984]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="21" pageNumber="227">Table S2</tableCitation>
), and by the lack of a large circular gonopore with a sclerotized edge at its anterior and posterior margin on each sternal plate of segment 11
<emphasis id="B969EAE2FFE1FF85FBDCF991FC44F9EF" box="[1036,1097,1628,1648]" italics="true" pageId="21" pageNumber="227">versus</emphasis>
its existence in females (
<figureCitation id="13262A75FFE1FF85FA99F993FD96F9EE" box="[1353,1435,1630,1649]" captionStart="Fig" captionStartId="20.[88,113,1662,1676]" captionTargetBox="[176,1386,178,1632]" captionTargetId="figure-115@20.[176,1386,178,1632]" captionTargetPageId="20" captionText="Fig. 13. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A‾E) and females USNM 1562543 (F) and USNM 1562542 (G) from the Gulf of Lion, ventral view, light micrographs. A, Habitus. B, Neck with placids. C—E, Segments 1‾3 (C), 4‾6 (D), and 7‾9 (E). Arrows in C point to spinose extensions of free flap. F, Sternal plates of segments 2‾4; arrows mark paraventral area of micropapillae.G, Segments 9‾11 with gonad and gonopores. Arrowheads in C—E mark type-6 sensory spots. Abbreviations:amp, female-specific area of micropapillae; ap, female-specific appendage; bpl, broad placid; fa, filamentous appendage; ff, free flap; gc, gland cell; go, female gonad; gp, gonopore; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; mts, midterminal spine; mvpl, midventral placid; npl, narrow placid; pa, pachycyclus; s, segment; st, sternal plate; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A = 100 Mm; E = 50 Mm, valid for B—G; F = 30 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086149" httpUri="https://zenodo.org/record/15086149/files/figure.png" pageId="21" pageNumber="227">Fig. 13G</figureCitation>
).
</paragraph>
<paragraph id="8BA236F0FFE1FF85FCEFF959FBD5F937" blockId="21.[831,1501,1684,1872]" box="[831,984,1684,1704]" pageId="21" pageNumber="227">
<emphasis id="B969EAE2FFE1FF85FCEFF959FBD5F937" box="[831,984,1684,1704]" italics="true" pageId="21" pageNumber="227">3.4.5. Variation</emphasis>
</paragraph>
<paragraph id="8BA236F0FFE1FF85FC8FF97CFC75F8CF" blockId="21.[831,1501,1684,1872]" pageId="21" pageNumber="227">
Out of the three specimens from the Gulf of Lion, one male (USNM 1562541) shows a middorsal cuspidate spine on segment 5 (
<figureCitation id="13262A75FFE1FF85FC97F924FB95F963" box="[839,920,1769,1788]" captionStart="Fig" captionStartId="21.[113,138,1401,1415]" captionTargetBox="[164,1449,178,1371]" captionTargetId="figure-287@21.[164,1449,178,1371]" captionTargetPageId="21" captionText="Fig.14. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A—C, H) and male USNM 1562541 (D, G) and female USNM 1562542 (E, F) from the Gulf of Lion, dorsal view, light micrographs. A—B, Anterior segments, left side (A), and right side (B); arrows mark tergal extensions of free flap. C, Lateral terminal accessory spine with two thin areas (arrows).D—E, Segments 6‾8 (D) and 5‾7 (E) with middorsal cuspidate spine (arrows) on segment 5 (D) and segments 5 and 7 (E). F—G, Segments 9‾11.H, Segment 9 with protonephridial openings (arrows). Arrowheads in A—B, E, and H mark type-6 sensory spots. Abbreviations: bpl, broad placid; epi, epizoic protozoan; ltas, lateral terminal accessory spine; ld, laterodorsal acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; pa, pachycyclus; s, seg ment; ssp3, type-3 sensory spot. Numbers, except for sensory-spot type, refer to the segment number. Scale bars: C = 20 Mm; G = 50 Mm, valid for A, B, and D—H." figureDoi="http://doi.org/10.5281/zenodo.15086151" httpUri="https://zenodo.org/record/15086151/files/figure.png" pageId="21" pageNumber="227">Fig. 14D</figureCitation>
), whereas two females (USNM 1562542, USNM 1562543) reveal a middorsal cuspidate spine on segments 5 and 7 (
<figureCitation id="13262A75FFE1FF85FA53F8C8FDDEF887" box="[1411,1491,1797,1816]" captionStart="Fig" captionStartId="21.[113,138,1401,1415]" captionTargetBox="[164,1449,178,1371]" captionTargetId="figure-287@21.[164,1449,178,1371]" captionTargetPageId="21" captionText="Fig.14. Condyloderes multispinosus, female USNM 1562540 from the northern Adriatic Sea (A—C, H) and male USNM 1562541 (D, G) and female USNM 1562542 (E, F) from the Gulf of Lion, dorsal view, light micrographs. A—B, Anterior segments, left side (A), and right side (B); arrows mark tergal extensions of free flap. C, Lateral terminal accessory spine with two thin areas (arrows).D—E, Segments 6‾8 (D) and 5‾7 (E) with middorsal cuspidate spine (arrows) on segment 5 (D) and segments 5 and 7 (E). F—G, Segments 9‾11.H, Segment 9 with protonephridial openings (arrows). Arrowheads in A—B, E, and H mark type-6 sensory spots. Abbreviations: bpl, broad placid; epi, epizoic protozoan; ltas, lateral terminal accessory spine; ld, laterodorsal acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; pa, pachycyclus; s, seg ment; ssp3, type-3 sensory spot. Numbers, except for sensory-spot type, refer to the segment number. Scale bars: C = 20 Mm; G = 50 Mm, valid for A, B, and D—H." figureDoi="http://doi.org/10.5281/zenodo.15086151" httpUri="https://zenodo.org/record/15086151/files/figure.png" pageId="21" pageNumber="227">Fig. 14E</figureCitation>
). The female from the northern Adriatic Sea (USNM 1562540) lacks any middorsal cuspidate spine.
</paragraph>
<paragraph id="8BA236F0FFE1FF85FCEFF8B9FC09F817" blockId="21.[831,1500,1908,1984]" box="[831,1028,1908,1928]" pageId="21" pageNumber="227">
<emphasis id="B969EAE2FFE1FF85FCEFF8B9FC09F817" box="[831,1028,1908,1928]" italics="true" pageId="21" pageNumber="227">3.4.6. Juvenile stage</emphasis>
</paragraph>
<paragraph id="8BA236F0FFE1FF86FC8FF85CF940FD6B" blockId="21.[831,1500,1908,1984]" lastBlockId="22.[88,757,179,756]" lastPageId="22" lastPageNumber="228" pageId="21" pageNumber="227">
One small specimen (trunk length = 149
<emphasis id="B969EAE2FFE1FF85FAD2F843FD03F83B" bold="true" box="[1282,1294,1934,1956]" pageId="21" pageNumber="227">M</emphasis>
m) possesses a very thin cuticle, has segments 10 and 11 still fused, lacks a free flap but shows a series of postmarginal spicula with a stronger base, and reveals papilla-like sensory spots elevated above the surface of the cuticle (
<figureCitation id="13262A75FFE2FF86FF78FF26F915FF61" box="[168,280,235,254]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="22" pageNumber="228">Fig. S3A—D</figureCitation>
). Based on these observations, this specimen is regarded as a juvenile stage. Placids with condyles exist but are difficult to separate from each other. The trunk is composed of segments 1—9 separated from each other and from segments 10 + 11 which are still fused (
<figureCitation id="13262A75FFE2FF86FE56FE97F9D8FEF1" box="[390,469,346,366]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="22" pageNumber="228">Fig. S3B</figureCitation>
). Segment 1 reveals a single cuticular plate, whereas segments 2
<emphasis id="B969EAE2FFE2FF86FE0DFEBBF9E0FE16" bold="true" box="[477,493,374,393]" pageId="22" pageNumber="228"></emphasis>
11 probably possess one tergal and one sternal plate (
<figureCitation id="13262A75FFE2FF86FEADFE5FF9C1FE3A" box="[381,460,402,421]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="22" pageNumber="228">Fig. S3A</figureCitation>
). Short cuticular hairs with a stronger base cover all segments except ventromedially and midlaterally, where the dorsoventral muscles originate (
<figureCitation id="13262A75FFE2FF86FDBEFE07F868FE66" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="22" pageNumber="228">Figs. S3A and B</figureCitation>
). A type-3 sensory spot occurs ventrolaterally and subdorsally on segment 11 (
<figureCitation id="13262A75FFE2FF86FED6FDCFF954FD8A" box="[262,345,514,533]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="22" pageNumber="228">Fig. S3B</figureCitation>
). Type-6 sensory spots appear at least ventromedially on segments 2, 5, and 9, sublaterally on segments 1, 3, 6, 7, and 9, and paradorsally on segments 1 (almost subdorsally), 4, and 6
<emphasis id="B969EAE2FFE2FF86FF79FD98F8B7FDF7" bold="true" box="[169,186,597,616]" pageId="22" pageNumber="228"></emphasis>
9 (
<figureCitation id="13262A75FFE2FF86FF06FD98F94AFDF6" box="[214,327,597,617]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="22" pageNumber="228">Fig. S3A—D</figureCitation>
). Acicular spines are found lateroventrally and middorsally on segments 1
<emphasis id="B969EAE2FFE2FF86FE42FDBCF9ADFD1B" bold="true" box="[402,416,625,644]" pageId="22" pageNumber="228"></emphasis>
10 as well as in a lateral accessory position and midterminally on segment 11 (
<figureCitation id="13262A75FFE2FF86FDC0FD40FA72FD3E" box="[528,639,653,673]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="22" pageNumber="228">Fig. S3A—D</figureCitation>
). Cuspidate spines are located ventrolaterally on segments 2, 5, and 9, in a lateralaccessory position on segments 4 and 8, and middorsally on segment 5 (
<figureCitation id="13262A75FFE2FF86FF00FD2CF94DFD6B" box="[208,320,737,756]" captionStart="Fig" captionStartId="5.[113,138,1828,1842]" captionTargetBox="[240,1374,178,1799]" captionTargetId="figure-1@5.[240,1374,178,1799]" captionTargetPageId="5" captionText="Fig. 3. Condyloderes agnetis sp. nov., female holotype, ZMB 11763. Light micrographs. A, Habitus, ventral view. B, Habitus, dorsal view. C, Neck with placids and segment 1, dorsal view.D, Segments 1‾3, dorsal view. E, Segments 7‾10, dorsal view.F, Segments 8‾9, ventral view.G, Segments 9‾11 with gonopores.H, Close-up of the trunk (segments 5‾6) with an oocyte. Arrowheads in C—D mark type-6 sensory spots. Abbreviations: bpl, broad placid; gp, gonopore; lacu, lateral accessory cuspidate spine; ltas, lateral terminal accessory spine; lv, lateroventral acicular spine; md, middorsal acicular spine; mts, midterminal spine; npl, narrow placid; oo, oocyte; pdcu, paradorsal cuspidate spine; s, segment; rch, longitudinal rows of regularly arranged cuticular hairs; sdcu, subdorsal cuspidate spine; vlcu, ventrolateral cuspidate spine. Numbers refer to the segment number. Scale bars: A, B = 100 Mm; C, G, F = 20 Mm; D, E = 50 Mm." figureDoi="http://doi.org/10.5281/zenodo.15086129" httpUri="https://zenodo.org/record/15086129/files/figure.png" pageId="22" pageNumber="228">Fig. S3A—D</figureCitation>
).
</paragraph>
<paragraph id="8BA236F0FFE2FF86FF88FCD5FA1EFCB3" blockId="22.[88,758,792,1398]" box="[88,531,792,812]" pageId="22" pageNumber="228">
<emphasis id="B969EAE2FFE2FF86FF88FCD5FA1EFCB3" box="[88,531,792,812]" italics="true" pageId="22" pageNumber="228">3.4.7. Comparison with previous descriptions</emphasis>
</paragraph>
<paragraph id="8BA236F0FFE2FF86FFA7FCF8FA7FFAE9" blockId="22.[88,758,792,1398]" pageId="22" pageNumber="228">
The species was originally described as
<taxonomicName id="4C1D4D73FFE2FF86FDE8FCF9F8A7FCFC" authorityName="McIntyre" authorityYear="1962" class="Cyclorhagida" family="Centroderidae" genus="Centroderes" kingdom="Animalia" order="Kentrorhagata" pageId="22" pageNumber="228" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE2FF86FDE8FCF9F8A7FCFC" italics="true" pageId="22" pageNumber="228">Centroderes multispinosus</emphasis>
</taxonomicName>
by McIntyre (1962) and later placed in a new genus and combined as
<taxonomicName id="4C1D4D73FFE2FF86FF22FCA6FAC1FC1F" authority="(McIntyre, 1962)" authorityName="Higgins" authorityYear="1969" baseAuthorityName="McIntyre" baseAuthorityYear="1962" box="[242,716,875,896]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="22" pageNumber="228" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE2FF86FF22FCA6FA07FCE0" box="[242,522,875,895]" italics="true" pageId="22" pageNumber="228">Condyloderes multispinosus</emphasis>
(McIntyre, 1962)
</taxonomicName>
by Higgins (1969). Most recently, Neuhaus et al. (2019) re-described the male holotype and three additional males from the North Sea. The specimens from the Mediterranean Sea differ from the specimens from the North Sea in the possession of a type-6 sensory spot ventromedially on segments 5 and 11
<emphasis id="B969EAE2FFE2FF86FE05FC3AFA1FFB94" box="[469,530,1015,1035]" italics="true" pageId="22" pageNumber="228">versus</emphasis>
the lack of these spots in the specimens from the North Sea and in the existence of a middorsal cuspidate spine on segments 5 and/or 7 at least in some specimens
<emphasis id="B969EAE2FFE2FF86FF16FB86F90EFBC0" box="[198,259,1099,1119]" italics="true" pageId="22" pageNumber="228">versus</emphasis>
their lack in the northern population. The juvenile specimen from the Mediterranean Sea possesses a cuspidate spine in a lateral accessory position on segment 4, which is missing in any adult specimen from the North Sea and from the Mediterranean Sea. We regard the above mentioned differences as variation within one and the same species. It should be kept in mind that few specimens of this species are known both from the North Sea and from the Mediterranean Sea (
<tableCitation id="C69F034BFFE2FF86FE55FAC2F9D4FABD" box="[389,473,1295,1314]" captionStart="Table 2" captionStartId="10.[88,132,180,194]" captionTargetPageId="10" captionText="Table 2 Characters of Condyloderes agnetis sp.nov.,C.clarae sp.nov., and C.multispinosus present in the majority of specimens of a given species.Number of studied specimens mounted for light microscopy provided in () after species name; for C.multispinosus including specimens from Neuhaus et al. (2019). Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Characters found in one gender only are indicated.Characters occupying an intermediate position to two reference positions are arranged right or left in a column (on segment 7: paradorsally to subdorsally; on segments 5, 8 and 9: ventrolaterally to ventromedially). Positions of more frequent variation of organ (n ± 5) are highlighted by oblique hatching.Information for C. multispinosus also taken from Neuhaus et al.(2019). Abbreviations:[], indicates variation in minority of specimens;ac, acicular spine; amp, area of micropapillae; ap, appendage; gc, gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lv, lateroventral; md, middorsal; ml, midlateral; mts, midterminal spine; ne, nephridial openings; pd, paradorsal; pv, paraventral; sd, subdorsal; sh, short; sl, sublateral; ssp3, type-3 sensory spot; vl, ventrolateral; vm, ventromedial." pageId="22" pageNumber="228">Table S2</tableCitation>
). In any case, this study expands the distribution record of
<taxonomicName id="4C1D4D73FFE2FF86FE4DFAE7FA37FAA1" authorityName="Higgins" authorityYear="1969" baseAuthorityName="McIntyre" baseAuthorityYear="1962" box="[413,570,1322,1342]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="22" pageNumber="228" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE2FF86FE4DFAE7FA37FAA1" box="[413,570,1322,1342]" italics="true" pageId="22" pageNumber="228">C. multispinosus</emphasis>
</taxonomicName>
considerably from the North Sea to the Mediterranean Sea suggesting that the species should also occur in the northeastern Atlantic Ocean.
</paragraph>
<paragraph id="8BA236F0FFE2FF86FF88FA54FADDFA32" blockId="22.[88,758,1433,1593]" box="[88,720,1433,1454]" pageId="22" pageNumber="228">
<emphasis id="B969EAE2FFE2FF86FF88FA54F931FA32" box="[88,316,1433,1453]" italics="true" pageId="22" pageNumber="228">3.4.8. Co-occurrence of</emphasis>
<taxonomicName id="4C1D4D73FFE2FF86FE93FA57F9E6FA32" authorityName="Higgins" authorityYear="1969" baseAuthorityName="McIntyre" baseAuthorityYear="1962" box="[323,491,1434,1453]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="22" pageNumber="228" phylum="Kinorhyncha" rank="species" species="multispinosus">C. multispinosus</taxonomicName>
<emphasis id="B969EAE2FFE2FF86FE22FA54FA1AFA32" box="[498,535,1433,1453]" italics="true" pageId="22" pageNumber="228">and</emphasis>
<taxonomicName id="4C1D4D73FFE2FF86FDCEFA56FA72FA32" authorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" authorityYear="2019" box="[542,639,1434,1454]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="22" pageNumber="228" phylum="Kinorhyncha" rank="species" species="agnetis" status="sp. nov.">C. agnetis</taxonomicName>
<emphasis id="B969EAE2FFE2FF86FD57FA54FADDFA32" box="[647,720,1433,1453]" italics="true" pageId="22" pageNumber="228">
<taxonomicNameLabel id="A25A5799FFE2FF86FD57FA54FADDFA32" box="[647,720,1433,1453]" pageId="22" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BA236F0FFE2FF86FFA7FA7BFA55F9A6" blockId="22.[88,758,1433,1593]" pageId="22" pageNumber="228">
Asingle male of
<taxonomicName id="4C1D4D73FFE2FF86FEFBFA78F9C6FA56" authorityName="Higgins" authorityYear="1969" baseAuthorityName="McIntyre" baseAuthorityYear="1962" box="[299,459,1461,1481]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="22" pageNumber="228" phylum="Kinorhyncha" rank="species" species="multispinosus">
<emphasis id="B969EAE2FFE2FF86FEFBFA78F9C6FA56" box="[299,459,1461,1481]" italics="true" pageId="22" pageNumber="228">C. multispinosus</emphasis>
</taxonomicName>
was collected together with
<taxonomicName id="4C1D4D73FFE2FF86FF88FA1CF8B8FA7A" authorityName="Zotto &amp; Neuhaus &amp; Yamasaki &amp; Todaro" authorityYear="2019" box="[88,181,1489,1509]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="22" pageNumber="228" phylum="Kinorhyncha" rank="species" species="agnetis" status="sp. nov.">
<emphasis id="B969EAE2FFE2FF86FF88FA1CF8B8FA7A" box="[88,181,1489,1509]" italics="true" pageId="22" pageNumber="228">C. agnetis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A5799FFE2FF86FF6CFA1EF904FA79" box="[188,265,1491,1510]" pageId="22" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
at a station off Trappeto, Gulf of Castellammare, Sicily, southern Tyrrhenian Sea (Table 1). The two species also occurred in almost the same location off Rovinj, northern Adriatic Sea, but were collected in different years (Table 1).
</paragraph>
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<mods:title id="28BCAAC6A7D1444C08458A7BF9C4F998">The genus Condyloderes (Kinorhyncha: Cyclorhagida) in the Mediterranean Sea, including the description of two new species with novel characters</mods:title>
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<mods:namePart id="2118B5E209CAB5B551A24AF26DF83619">Zotto, Matteo Dal</mods:namePart>
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<paragraph id="8BA236F0FFF5FF91FC8FF9B7FCB5F912" blockId="1.[863,1290,1574,1677]" box="[863,1208,1656,1677]" pageId="1" pageNumber="207">
Genus
<taxonomicName id="4C1D4D73FFF5FF91FC74F9B5FCB5F912" ID-CoL="7NTDT" authority="Higgins, 1969" authorityName="Higgins" authorityYear="1969" box="[932,1208,1656,1677]" class="Cyclorhagida" family="Centroderidae" genus="Condyloderes" kingdom="Animalia" order="Kentrorhagata" pageId="1" pageNumber="207" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EAE2FFF5FF91FC74F9B5FC28F913" box="[932,1061,1656,1676]" italics="true" pageId="1" pageNumber="207">Condyloderes</emphasis>
Higgins, 1969.
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<heading id="D0EA819CFFF5FF91FC8FF97CFC21F95B" bold="true" box="[863,1068,1713,1732]" fontSize="8" level="1" pageId="1" pageNumber="207" reason="6">
<emphasis id="B969EAE2FFF5FF91FC8FF97CFC21F95B" bold="true" box="[863,1068,1713,1732]" pageId="1" pageNumber="207">Emended diagnosis</emphasis>
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<paragraph id="8BA236F0FFF5FF92FC8FF900F8AEFABF" blockId="1.[831,1501,1713,1984]" lastBlockId="2.[88,757,875,1312]" lastPageId="2" lastPageNumber="208" pageId="1" pageNumber="207">
Mouth cone with 9 individual outer oral styles with broader base and thin and flexible anterior part; 14 trichoscalids each originating from a trichoscalid plate and accompanied anteriorly by two pointed filamentous appendages; 16 placids with broader midventral placid, neighboured by two narrower placids and alternatingly a broader and a narrower placid; all placids with knobby projections (= condyles) in one or two lateral rows in narrower placids and two or three lateral rows in broader placids; midventral placid with two or three apical condyles, zero or two in intermediate lateral row, and 3
<emphasis id="B969EAE2FFF6FF92FDE8FCA6FA44FCE1" bold="true" box="[568,585,875,894]" pageId="2" pageNumber="208"></emphasis>
6 in basal lateral row; ventromedial to midventral free flap and primary pectinate fringe of segment 1 at least partly reduced; acicular spine middorsally on segments 1
<emphasis id="B969EAE2FFF6FF92FE8BFC73F967FC4E" bold="true" box="[347,362,958,977]" pageId="2" pageNumber="208"></emphasis>
9, midterminally on segment 11 (= midterminal spine), lateroventrally on segments 2
<emphasis id="B969EAE2FFF6FF92FD5BFC17FA91FC72" bold="true" box="[651,668,986,1005]" pageId="2" pageNumber="208"></emphasis>
9, lateroventrally or in lateral accessory position on segment 1, laterodorsally on segment 10 in male and in stage-1 female (in species with more than one adult life history stage), and lateral accessorily on segment 11 (= lateral terminal accessory spine); cuspidate spines present on two or more segments from 1 to 9; type-3 sensory spot ventrolaterally next to lateral terminal accessory spine and subdorsally or laterodorsally on segment 11; type-6 sensory spot elevated above trunk surface giving the impression to be half-drowned in the trunk cuticle; ventromedial appendage at least on two segments of segments 5
<emphasis id="B969EAE2FFF6FF92FD69FB3CFAC7FA9B" bold="true" box="[697,714,1265,1284]" pageId="2" pageNumber="208"></emphasis>
8 in female.
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