diff --git a/data/C6/0D/AE/C60DAE35861CD808D8FEFCDCBECAF946.xml b/data/C6/0D/AE/C60DAE35861CD808D8FEFCDCBECAF946.xml new file mode 100644 index 00000000000..6e1ef8cae23 --- /dev/null +++ b/data/C6/0D/AE/C60DAE35861CD808D8FEFCDCBECAF946.xml @@ -0,0 +1,375 @@ + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Author + +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com + + + +Author + +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany + + + +Author + +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + + + +Author + +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + +text + + +Zoological Journal of the Linnean Society + + +2024 + +Zool. J. Linn. Soc. + + +2023-10-28 + + +202 + + +1 + + +1 +42 + + + + +https://doi.org/10.1093/zoolinnean/zlad143 + +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding + + + + + + + +Loncovilius semiflavus +( +Fairmaire and Germain, 1862 +) + + + + + + + + +( +Figs 6C +, +7D +, +9D, G +, +11A–C +, 12G, 13G, 14G) + + + +Fairmaire and Germain 1862: 428 +(original description as + +Quedius semiflavus + +); +Gemminger and Harold 1868: 570 +(as synonym of + +Quedius lividipennis + +); +Bernhauer and Schubert 1916: 434 +(as + +Quedius semiflavus + +, catalogue); +Germain 1903: 443 +(as + +Loncovilius semiflavus + +, characters); +Coiffait and Sáiz 1966: 411 +(as + +Loncovilius +s.s. +semiflavus + +; characters); Coiffait and Sáiz, 1968: 365 (as + +Loncovilius +s.s. +semiflavus + +, checklist); +Sáiz 1971: 385 +(as + +Loncovilius +s.s. +semiflavus + +; additional material); +Herman 2001b: 3083 +(as + +Loncovilius semiflavus + +; catalog). + + + +Type material. + +Lectotype +: male, pinned, with genitalia in a separate microvial, with labels as follows: Coll. et det A. Fauvel + +Quedius semiflavus +Fairm and Germ. R.I.S. + +c.N.B. 17.479/ Concepcion + + +/ +Lectotype +[Red label]/ + +Loncovilius + +( +s +. +s +.) + +semiflavus + +1964 det. Coiff. et Sáiz/J. L. Reyes-Hernández det. 2022 [RBINS]. + + +Paralectotypes +: + +Coll. et det. A. Fauvel, + +Quedius semiflavus +Fairm and Germ., R.I.Sc.N.B. + +17.479 + +/ +paralectotype +[red paper]/ + +Loncovilius semiflavus + +, 1964 det. Coiff. et Sáiz/J. L. Reyes-Hernández det. 2022; concepcion/ + +semiflavus +Fairm. Germ. + +/R.I.Sc.N.B. 17.479, Coll. et det. A. Fauvel/ +paralectotype +[red paper]/ + +Loncovilius semiflavus + +, 1964 det. Coiff. et Saiz/J. L. Reyes-Hernández det. 2022 [RBINS]. + + +Other material eoamined: +Supporting Information, File S7. + + +Diagnosis: +Elytra translucent yellow or yellowish-brown. Male tergite X with broad U-shape emargination medially ( +Fig. 12G +); female tergite X slightly emarginate medio-apically ( +Fig. 13G +). Aedeagus as in +Figure 11A–C +. + + +Description: +Measurements +♂ +[min–max (average); +N += 10]: FBL = 3.82–4.19 (4.02); TL = 6.25–6.88 (6.54); HW = 1–1.05 (1.03); HL = 0.98–1.09 (1.02); HW/HL = 0.96–1.04 (1.01); PW = 1.28–1.44 (1.35); PL = 1.28–1.43 (1.35); PW/ PL = 0.99–1.01 (1); EW = 1.66–1.8 (1.74); EL = 1.5–1.8 (1.65); PW/HW = 1.27–1.37 (1.32). + + +Measurements +♀ +[min–max (average); +N += 10]: FBL = 4.1– 4.45 (4.29); TL = 7.2–8.6 (7.77); HW = 1.08–1.15 (1.1); HL = 1.04–1.13 (1.06); HW/HL = 1–1.06 (1.04); PW = 1.4– 1.5 (1.47); PL = 1.38–1.5 (1.46); PW/PL = 0.98–1.05 (1.01); EW = 1.85–1.9 (1.88); EL = 1.67–1.82 (1.77); PW/ HW = 1.27–1.39 (1.34). + +Head and pronotum yellow-orange or reddish-brown, usually females darker than males; elytra translucent yellow; antennae, mouthparts, and legs yellow or yellowish-brown; mesosternum, metasternum, metacoxa and abdomen dark reddish-brown; abdominal segments with pale posterior margin. + +Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and very fine microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with sparse setiferous punctures of medium and small size. Eyes medium size (EYL/HL = x = 0.5), from 1.2 to 1.4 times as long as temples (in lateral view); distance between eyes about 1.2 times as long as length of eye in males and 1.39 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 and 5 subequal in length; antennomeres 5 to 10 gradually becoming shorter apicad; a11 is from 1.88 to 2 times as long as a +10 in +males and from 1.6 to 1.8 times in females. Basal puncture single; parocular puncture usually single too, distinctly separated from eye margin; posterior frontal puncture located anterior to temporal puncture; ventral basal ridge rather straight, almost united with gular sutures; postgenal ridge reduced; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge distinct only laterally, nuchal and infraorbital ridges fused; gular sutures moderately and equally separated in both sexes. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere only weakly dilated apicad, subequal in width with more or less subconical apical palpomere. + +Pronotum about as wide as long, strongly convex, evenly curved at sides, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to three times as long as, diameter of punctures. Mesosternum with three or four macrosetae arranged in a row medially. +Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate. + +Abdominal tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area; tergite VIII distinctly emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sterniteVIIIwithstrong ( +Fig.14G +),V-shapedemarginationmedially; female sternite VIII with weak apical medial emargination; male sternite IX with arcuate emargination medially, its basal portion distinctly longer than distal portion; male tergite X with broad U-shape emargination medially ( +Fig. 12G +); female tergite X slightly emarginate medio-apically ( +Fig. 13G +); ovipositor, its second gonocoxite with one macroseta medially. + + +Aedeagus as in +Figure 11A–C +, its length 1.21; median lobe rod-like with moderately emarginate apex; paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging in a trapezoidal slightly emarginate apex, with small translucent ‘peg setae’, in lateral view almost straight and slightly concave apically. + + + +Distribution and habitat: +Loncovilius semiflavus + +is known only from +Chile +, from the +Biobío +, +Maule +, Ñuble, and +Santiago +Metropolitan regions where it occurs from +200 to 1200 m +of elevation. Its distribution is confined to the northern portions of the Valdivian temperate forest ecoregion. It has been collected by sifting leaf litter and beating the forest vegetation and flowers, e.g. from + +Drimys winteri + +( +Winteraceae +). + + + +Remarks: +Lonvcovilius semiflavus + +was described as a species of + +Quedius + +based on an unspecified number of specimens from ‘Concepcion’ (Concepción city in Central +Chile +) ( +Fairmaire and Germain 1862 +). +Gemminger and Harold (1868) +listed it as a synonym of + +Quedius lividipennis +Fairmaire and Germain, 1862 + +, but +Germain (1903) +considered both species valid when erecting a new genus + +Loncovilius + +for them. +Coiffait and Sáiz (1966) +examined +three specimens +from the collection of A. Fauvel in Brussels that they considered +syntypes +of + +L +. +semiflavus + +and designated +one male +as a +lectotype +. Morphology and labels of these +three specimens +match the original description and the type locality, respectively. According to Camousseight (1980), the type material of the species described by Philibert Germain is kept at the National Museum of +Chile +in +Santiago +. However, even though Camousseight (1980) listed type material for + +L. lividipennis + +(see below that species) and some other +Amblyopinini +as being deposited there, he did not mention any types of + +L. semiflavus +. + +Presumably, they could have disappeared, as some other material did during a period when this collection was neglected (Camousseight, 1980), while the material from RBINS studied by +Coiffait and Sáiz (1966) +are +syntypes +that P. Germain could have shared with A. Fauvel, a prominent colleague of that time. Based on that material, +Coiffait and Sáiz (1966) +illustrated the aedeagus of + +L +. +semiflavus + +and, together with + +L +. +lividipennis + +, treated it as a member of the subgenus + +Loncovilius +s.s. + +. Apart from the types, a specimen from La Jaula in Curicó Province of +Chile +listed in +Sáiz (1971) +, was the only additional material reported for this species prior to our revision. + + + + \ No newline at end of file diff --git a/data/C6/0D/AE/C60DAE35861DD80AD90CFE10BDBCFC36.xml b/data/C6/0D/AE/C60DAE35861DD80AD90CFE10BDBCFC36.xml new file mode 100644 index 00000000000..21a750aeeea --- /dev/null +++ b/data/C6/0D/AE/C60DAE35861DD80AD90CFE10BDBCFC36.xml @@ -0,0 +1,112 @@ + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Author + +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com + + + +Author + +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany + + + +Author + +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + + + +Author + +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + +text + + +Zoological Journal of the Linnean Society + + +2024 + +Zool. J. Linn. Soc. + + +2023-10-28 + + +202 + + +1 + + +1 +42 + + + + +https://doi.org/10.1093/zoolinnean/zlad143 + +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding + + + + + + + +semiflavus + +group + + + + + + + +Included species: +Loncovilius barclayi + +sp. nov. +and + +L +. +semiflavus +( +Fairmaire and Germain, 1862 +) + +. + + + + +Diagnosis: +Head about as wide as long, without coarse non-setiferous punctures; nuchal ridge absent dorsally, nuchal and infraorbital ridges joined; gula without isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin; PMP (usually reduced); females with antennomere 11 distinctly longer than antennomere 10 (a11:a10 ratio>1.5). Pronotum outline evenly curved, with one SLSP. Protibiae not sexually dimorphic, in both sexes without laterodorsal row of thick spines. Tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area near apical margin ( +Fig. 9D +); tergite VIII with medial apical emargination in both sexes; ovipositor, its second gonocoxite with one macroseta medially ( +Fig. 9G +). + + + + \ No newline at end of file diff --git a/data/C6/0D/AE/C60DAE35861DD80BD95BFC44BE97FCB5.xml b/data/C6/0D/AE/C60DAE35861DD80BD95BFC44BE97FCB5.xml index ba47743173c..0777cbd0bb2 100644 --- a/data/C6/0D/AE/C60DAE35861DD80BD95BFC44BE97FCB5.xml +++ b/data/C6/0D/AE/C60DAE35861DD80BD95BFC44BE97FCB5.xml @@ -1,72 +1,72 @@ - - - -Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) - - -Author + + +Author -Reyes-Hernández, José L. -Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark -jl.reyeshdez@gmail.com +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com - - -Author + + +Author -Hansen, Aslak Kappel -Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany - - -Author + + +Author -Shaw, Josh Jenkins -Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark - - -Author + + +Author -Solodovnikov, Alexey -Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark -text - - -Zoological Journal of the Linnean Society +text + + +Zoological Journal of the Linnean Society - -2024 - -Zool. J. Linn. Soc. + +2024 + +Zool. J. Linn. Soc. - -2023-10-28 + +2023-10-28 - -202 + +202 - -1 + +1 - -1 -42 + +1 +42 - -https://doi.org/10.1093/zoolinnean/zlad143 + +https://doi.org/10.1093/zoolinnean/zlad143 -journal article -10.1093/zoolinnean/zlad143 -0024-4082 -14443945 -CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding - + @@ -84,7 +84,9 @@ ( Figs 6D -, 11D–F, 12H, 13H, 14D) +, +11D–F +, 12H, 13H, 14D) @@ -123,7 +125,7 @@ male, pinned, with genitalia in a separate microvial, with labels as follows: Loncovilius barclayi des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at FMNH. - + Paratypes : @@ -147,7 +149,7 @@ L. E. [ 1♀ FMNH]; - + CHILE : Valdivia @@ -237,7 +239,7 @@ NMPC]; [ 1♂ MNNC ex. FMNH]. All paratypes -with label: PAÞTYPE +with label: PARATYPE Loncovilius barclayi @@ -251,7 +253,9 @@ Elytra yellow or yellowish-brown with black marks around mesoscutellum and on su Fig. 12H ); female tergite X deeply emarginate medio-apically ( Fig. 13H -). Aedeagus as in Figure 11D–F. +). Aedeagus as in +Figure 11D–F +. Description: @@ -279,7 +283,7 @@ females. Basal and parocular punctures usually single; posterior frontal punctur Pronotum about as wide as long, strongly convex, evenly curved, with two PPDS, without APP, with one SLSP; flexible postcoxal hypomeral extension (or process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to two or four times as long as, diameter of punctures. Mesosternum with four macrosetae arranged in a row medially. Protibiae not sexually dimorphic, without laterodorsal row of thick spines in both sexes. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 shorter than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate. -Abdominal tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area; tergite VIII distinctly emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with strong, V-shaped emargination medially ( +Abdominal tergites III and IV with only a few rows of coarse punctures of moderate density, with large impunctate area; tergite VIII distinctly emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with strong, V-shaped emargination medially ( Fig. 14H ); female sternite VIII with weak apical medial emargination; male sternite IX not emarginate medio-apically, its basal portion distinctly longer than distal portion; male tergite X trilobed ( Fig. 12H @@ -289,10 +293,14 @@ Abdominal tergites III and IV with only a few rows of coarse punctures of modera Fig 9G ). -Aedeagus as in Figure 11D–F, its length ~ 1.46; median lobe rod-like, in parameral view apex deeply emarginate; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into a deeply emarginate apex, with dark peg setae, in lateral view almost straight and slightly concave apically. + +Aedeagus as in +Figure 11D–F +, its length ~ 1.46; median lobe rod-like, in parameral view apex deeply emarginate; internal sac with pair of long I-shaped copulatory sclerites. Paramere slightly produced over apex of median lobe, wider than median lobe, its sides converging into a deeply emarginate apex, with dark peg setae, in lateral view almost straight and slightly concave apically. + Etymology: -Ŋe species epithet is a patronym in recognition of Maxwell V.L.Barclay, in particular for his earlier mentoring of J.J.S.. +The species epithet is a patronym in recognition of Maxwell V.L.Barclay, in particular for his earlier mentoring of J.J.S.. diff --git a/data/C6/0D/AE/C60DAE35861ED80EDB33FE0EBAF8FDA5.xml b/data/C6/0D/AE/C60DAE35861ED80EDB33FE0EBAF8FDA5.xml new file mode 100644 index 00000000000..6e90e17a688 --- /dev/null +++ b/data/C6/0D/AE/C60DAE35861ED80EDB33FE0EBAF8FDA5.xml @@ -0,0 +1,337 @@ + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Author + +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com + + + +Author + +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany + + + +Author + +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + + + +Author + +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + +text + + +Zoological Journal of the Linnean Society + + +2024 + +Zool. J. Linn. Soc. + + +2023-10-28 + + +202 + + +1 + + +1 +42 + + + + +https://doi.org/10.1093/zoolinnean/zlad143 + +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding + + + + + + + +Loncovilius lividipennis +( +Fairmaire and Germain, 1862 +) + + + + + + + + +( +Figs 7A +, +9E +, +11G, H +, 12I, 13I, 14I) + + + +Fairmaire and Germain 1862: 430 +(as + +Philonthus lividipennis + +, original description); Fauvel, 1866: 338 (as + +Quedius lividipennis + +, additional characters); +Bernhauer and Schubert 1916: 427 +(as + +Quedius lividipennis + +, catalogue); Scheerpeltz, 1972: 23 [as + +Quedius (Loncovilius) lividipennis +; + +additional material]; +Germain 1903: 443 +(as + +Loncovilius lividipennis + +, characters); +Coiffait and Sáiz 1966: 409 +(as + +Loncovilius +s.s. +lividipennis + +; characters; additional material); Coiffait and Sáiz, 1968: 365 (as + +Loncovilius +s.s. +lividipennis + +, checklist); +Sáiz 1971: 385 +(as + +Loncovilius +s.s. +lividipennis + +; additional material); +Herman 2001b: 3083 +(as + +Loncovilius lividipennis + +, catalogue); + +Jenkins Shaw +et al +., 2020: 337 + +(as + +Loncovilius +nr. +semiflavus + +; phylogeny). + + +Material eoamined: +Supporting Information, File S7. + + +Diagnosis: +Pronotum very rarely with APP. Elytra yellow or yellowish-brown. Sternite III with basal transverse carina sharply pointed medioapically. Aedeagus as in +Figure 11G, H +. + + +Description: +Measurements +♂ +[min–max (average); +N += 10]: FBL = 3.09–3.36 (3.22); TL = 5.15–6.86 (6.20); HW = 0.79– 0.84 (0.81); HL = 0.78–0.84 (0.81); HW/HL = 0.98–1.02 (1); PW = 1.05–1.17 (1.11); PL = 0.97–1.08 (1.02); PW/ PL = 1.08–1.11 (1.09); EW = 1.38–1.84 (1.56); EL = 1.33– 1.44 (1.4); PW/HW = 1.31–1.42 (1.37). + + +Measurements +♀ +[min–max (average); +N += 10]: FBL = 3.13–3.68 (3.42); TL = 5.87–6.68 (6.29); HW = 0.78– 0.89 (0.85); HL = 0.8–0.9 (0.85); HW/HL = 0.98–1.02 (1); PW = 0.95–1.24 (1.14); PL = 0.89–1.16 (1.06); PW/ PL = 1.04–1.11 (1.08); EW = 1.54–1.82 (1.64); EL = 1.33– 1.63 (1.5); PW/HW = 1.1–1.41 (1.35). + +Most of antennae, mouthparts, pronotum, elytra, legs yellow or yellowish-brown; last three or five antennomeres and pronotum sometimes darker; head, mandibles, pro-, meso-, and metasternum, metacoxal, and abdomen reddish-brown; tergites and sternites with pale posterior margin. + +Head from slightly wider than long, to slightly longer than wide; dorsally and ventrally glossy with few micropunctures and microsculpture of transverse waves, without coarse non-setiferous punctures; posterior angles indistinct with several setiferous punctures of medium and small size. Eyes small (EYL/ HL = x = 0.47), from 1.07 to 1.44 times as long as temples (in lateral view) in males and +1.06 to 1.25 in +females; distance between eyes about 1.16 times as long as length of eye in males and 1.35 times in females. Antennomeres 2 and 3 subequal in length; antennomeres 4 to 7 subequal in length; antennomeres 8 to 10 subequal in length; a11 from 1.88 to 2.2 times long as a +10 in +males and from +1.72 to 1.94 in +females. Basal and parocular punctures usually single; posterior frontal puncture located posterior to temporal puncture; without small setose punctures between frontoclypeal punctures and anterior frontal punctures; ventral basal ridge rather straight, almost united with gular sutures; postgenal ridge present; postmandibular ridge reaching close to eye margin; PMP greatly reduced; nuchal ridge present laterally and dorsally, fused with infraorbital ridge; gula without microsculpture medially, gular sutures moderately separated. Mandibles without dorsolateral groove; labrum with trapezoidal sclerotized region and entire apical margin; mentum with seta alpha only; penultimate labial palpomere markedly dilated apicad, apical palpomere distinctly narrower than previous and more or less subconical. + +Pronotum slightly transverse, convex, evenly curved, with two PPDS, usually without APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to one or three times as long as, diameter of punctures. Mesosternum with seven macrosetae arranged in two rows medially. +Protibiae usually sexually dimorphic, with laterodorsal row of thick spines only in females; rarely in males too. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 moderately longer than 5, metatarsomere 4 ventrally with apical margin straight or slightly sinuate. + +Abdominal tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina descending medially at a sharp point; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with deep sharp V-shaped emargination medially ( +Fig. 14I +); female sternite VIII with arcuate emargination medially; male sternite IX with V-shaped emargination medially, its basal portion moderately longer than distal portion; male tergite X with deep U-shaped emargination medially ( +Fig. 12I +); female tergite X with V-shaped emargination medially ( +Fig. 13I +); ovipositor, its second gonocoxite with one macroseta medially. + + +Aedeagus as in +Figure 11G, H +; its total length 1.05–1.15. + + + +Distribution and habitat: +Loncovilius lividipennis + +is known from +Chile +and +Argentina +. In +Chile +, it occurs in the +Araucanía +, +Biobío +, +Los Lagos +, +Los Ríos +, and Ñuble regions. In +Argentina +, it is confined to the westernmost portions of the +Chubut +, +Neuquén +, and +Río Negro +provinces. The species occurs from +550 to 1600 m +elevation; it is reported in the + +Araucaria +- +Nothofagus + +forests from central and north-central portions of the Valdivian temperate forest ecoregion. It has been collected using Malaise and window traps, beating the forest vegetation, attracted to ultraviolet lights, sifted from moss and leaf litter, and directly from male cones of + +Araucaria araucana + +and flowers, e.g. from + +Desmaria mutabilis + +( +Loranthaceae +) on + +Nothofagus +sp. + + + +Remarks: +This species was described in the genus + +Philonthus +( +Fairmaire and Germain 1862: 430 +) + +based on an unspecified number of +syntypes +from the Andean forests near Chillan, a city in central +Chile +. Fauvel (1866) moved the species to the genus + +Quedius + +, while +Germain (1903) +to his genus + +Loncovilius + +. Scheerpeltz (1972), who treated + +Loncovilius + +at the rank of a subgenus of + +Quedius + +, listed significant new material for this species. +Coiffait and Sáiz (1966) +provided the species redescription and first illustration of the aedeagus based on the non-type material that comes from localities other than ‘Chillan’. +Sáiz (1971) +listed a few more specimens. Camousseight (1980) lists a ‘ +paratype +N 2267–2272’ of + +Philonthus lividipennis + +from Germain’s collection at the National Museum of +Chile +in +Santiago +, which is apparently a +syntype +. +Coiffait and Sáiz (1966) +denoted one of the specimens they examined as a +neotype +of + +L.lividipennis + +. Although we were unable to examine any +syntypes +of + +L. lividipennis + +, we assume that the currently accepted concept of this species is correct, because the only other externally similar species, + +L. carlsbergi + +sp. nov. +, is known only further south, from the Magellanic subpolar forests ecoregion. + + + + \ No newline at end of file diff --git a/data/C6/0D/AE/C60DAE35861FD808D90AF96BBA7DFE79.xml b/data/C6/0D/AE/C60DAE35861FD808D90AF96BBA7DFE79.xml new file mode 100644 index 00000000000..732abfdef99 --- /dev/null +++ b/data/C6/0D/AE/C60DAE35861FD808D90AF96BBA7DFE79.xml @@ -0,0 +1,110 @@ + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Author + +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com + + + +Author + +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany + + + +Author + +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + + + +Author + +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + +text + + +Zoological Journal of the Linnean Society + + +2024 + +Zool. J. Linn. Soc. + + +2023-10-28 + + +202 + + +1 + + +1 +42 + + + + +https://doi.org/10.1093/zoolinnean/zlad143 + +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding + + + + + + + +lividipennis + +group + + + + + + + +Included species: +Loncovilius carlsbergi + +sp. nov. +and + +L +. +lividipennis +( +Fairmaire and Germain, 1862 +) + +. + + + + +Diagnosis: +Head about as wide as long, without coarse non-setiferous punctures; nuchal ridge complete dorsally, connected with infraorbital ridge; gula without isodiametric microsculpture in the middle; posterior frontal puncture located posterior to temporal puncture; postmandibular ridge located close to eye margin; PMP (usually reduced); females with antennomere 11 distinctly longer than antennomere 10 (a11:a10 ratio>1.5). Pronotum outline evenly curved. Tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area near to apical margin ( +Fig. 9E +); tergite VIII with medial apical emargination in both sexes; ovipositor, its second gonocoxite with one macroseta medially. + + + + \ No newline at end of file diff --git a/data/C6/0D/AE/C60DAE35861FD809DB8AFE10BBDFFE67.xml b/data/C6/0D/AE/C60DAE35861FD809DB8AFE10BBDFFE67.xml index c2a63f12519..5e2ad5e2489 100644 --- a/data/C6/0D/AE/C60DAE35861FD809DB8AFE10BBDFFE67.xml +++ b/data/C6/0D/AE/C60DAE35861FD809DB8AFE10BBDFFE67.xml @@ -1,70 +1,70 @@ - - - -Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) - - -Author + + +Author -Reyes-Hernández, José L. -Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark -jl.reyeshdez@gmail.com +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com - - -Author + + +Author -Hansen, Aslak Kappel -Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany - - -Author + + +Author -Shaw, Josh Jenkins -Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark - - -Author + + +Author -Solodovnikov, Alexey -Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark -text - - -Zoological Journal of the Linnean Society +text + + +Zoological Journal of the Linnean Society - -2024 - -Zool. J. Linn. Soc. + +2024 + +Zool. J. Linn. Soc. - -2023-10-28 + +2023-10-28 - -202 + +202 - -1 + +1 - -1 -42 + +1 +42 - -https://doi.org/10.1093/zoolinnean/zlad143 + +https://doi.org/10.1093/zoolinnean/zlad143 -journal article -10.1093/zoolinnean/zlad143 -0024-4082 -14443945 -CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding @@ -86,14 +86,16 @@ Figs 7B, E , 8A -, 11I, J, 12J, 13J, 14J) +, +11I, J +, 12J, 13J, 14J) Zoobank registration: urn:lsid:zoobank.org:act: -C2A7BAAE- 3D40-4ED6-9C8D-09602F86B9B3 +C2A7BAAE-3D40-4ED6-9C8D-09602F86B9B3 . @@ -127,7 +129,7 @@ and Chilean Loncovilius carlsbergi des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at NHM. - + Paratypes : @@ -137,62 +139,98 @@ des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Deposited at N : Magallanes and Chilean -Antarctica, Puerto Edén + +Antarctica, +Puerto Edén + ] -Wellington Is. Pto. Eden +Wellington Is. +Pto. Eden 4.XII.1958 / CHILE -Fr Kuschel +Fr Kuschel /NHM [blue label] [ 2♂ 6♀ -NHM; +NHM +; 1♀ -NHMD ex. NHM; +NHMD +ex. +NHM +; 1♀ -CZUG ex. NHM; +CZUG +ex. +NHM +; 1♂ -FMNH ex. NHM]; [same data as above except:] +FMNH +ex. +NHM +] + +; + +[same data as above except:] 20.XI.1958 [ 5♂ 1♀ -NHM; +NHM +; 1♀ -FMNH ex. NHM]; [ +FMNH +ex. +NHM +] + +; + +[ CHILE : Magallanes -and Chilean Antarctica] +and +Chilean Antarctica +] Nothofagus antarctica / CHILE HE -I.Puerto Eden +I.Puerto Eden , -Isla Wellington -[-49.126, -74.40] 490S 40 ss. +Isla Wellington +[-49.126, -74.40] 490S 40 ft. 29.xi.1958 /NHM [ 1♀ -NHM]; [same data as above except:] -CHILE -HE6. [ +NHM +] + +; + +[same data as above except:] CHILE HE6. [ 1♀ -NHM]; [ +NHM +] + +; + +[ CHILE : Magallanes and Chilean Antarctica] S. CHILE -Pto. Edén Wellington Is. +Pto. Edén Wellington Is. [-49.126, -74.40] Dec.7-15,’62 PJDarlinton/FIELD MUSEUM ex MCZ (retained duplic.)/ Loncovilius lividipennis @@ -201,7 +239,12 @@ and Chilean Antarctica] S. A. Newton 1993 [ 1♂ -FMNH]; [ +FMNH +] + +; + +[ CHILE : Magallanes @@ -209,8 +252,7 @@ and Chilean Antarctica] Nothofagus antarctica / CHILE -Pena.Munoz, HM -3.Gamero. 20–40 ss. [-52.33, -73.53, +Pena.Munoz, HM3.Gamero. 20–40 ft. [-52.33, -73.53, 12m @@ -221,48 +263,70 @@ and Chilean Antarctica] / NHM [ 3♂ 2♀ -NHM; +NHM +; 1♂ 1♀ -MNNC ex. NHM]; [ +MNNC +ex. +NHM +] + +; + +[ CHILE : Magallanes -and Chilean Antarctica] Munoz Gamero [-52.33, -73.53] +and Chilean Antarctica] +Munoz Gamero +[-52.33, -73.53] 27.XII.1958 -/ -CHILE +/CHILE Fr. Kuschel /Shore/NHM [ 6♂ 3♀ -NHM; +NHM +; 1♂ -NHMD ex. NHM]; [ +NHMD +ex. +NHM +] + +; + +[ CHILE : Magallanes -and Chilean Antarctica] -P.Voillians -[Puerto Williams, -54.93, -67.619] 10.1-1959/ -CHILE -Fr Kuschel/4/NHM [ +and Chilean Antarctica] P.Voillians [ +Puerto Williams +, -54.93, -67.619] 10.1-1959/CHILE +Fr Kuschel +/4/NHM [ 1♂ -CZUG ex. NHM]. All +CZUG +ex. +NHM +] + +. All paratypes -with label: PAÞTYPE +with label: PARATYPE Loncovilius carlsbergi -des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022 - -. +des. Reyes-Hernández, Hansen, Jenkins Shaw et Solodovnikov 2022. Diagnosis: -Pronotum with APP. Elytra yellowish-brown. Sternite III with basal transverse carina evenly converging at an obtuse angle. Aedeagus as in Figure 11I, J. +Pronotum with APP. Elytra yellowish-brown. Sternite III with basal transverse carina evenly converging at an obtuse angle. Aedeagus as in +Figure 11I, J +. Description: @@ -290,7 +354,7 @@ females. Basal and parocular punctures usually single; posterior frontal punctur Pronotum slightly transverse, convex, evenly curved, with two PPDS, with an APP, with one SLSP; flexible postcoxal hypomeral extension (process) not interrupted by inferior line. Elytra evenly punctate, interspaces from about as long as, to three or five times as long as, diameter of punctures. Mesosternum with five macrosetae arranged in two rows medially. Protibiae sexually dimorphic, with laterodorsal row of thick spines only in females. In males, mesotarsomeres 1–4 with pale adhesive setae, mesotarsomeres 1–3 with terminal plate. In females, mesotarsomeres 2–4 with pale adhesive setae without terminal plate. In both sexes, metatarsomeres 2–4 with pale adhesive setae, without terminal plate; metatarsomere 1 moderately longer than 5, metatarsomere 4 ventrally with apical margin bilobed. -Abdominal tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flaưened. Male sternite VIII with rectangular V-shaped emargination medially ( +Abdominal tergites III and IV with several rows of punctures of moderate size and density, without a sizeable impunctate area; tergite VIII emarginate medio-apically. Sternite III with basal transverse carina evenly converging at an obtuse angle; lateral tergal sclerites IX not dorsoventrally or laterally flattened. Male sternite VIII with rectangular V-shaped emargination medially ( Fig. 14J ); female sternite VIII with arcuate emargination medially; male sternite IX with obtuse emargination medially, its basal portion moderately longer than distal portion; male tergite X with U-shaped emargination medially ( Fig. 12J @@ -298,10 +362,14 @@ Abdominal tergites III and IV with several rows of punctures of moderate size an Fig. 13J ); ovipositor, its second gonocoxite with one macroseta medially. -Aedeagus as in Figure 11I, J; its total length 1.35–1.45. + +Aedeagus as in +Figure 11I, J +; its total length 1.35–1.45. + Etymology: -Ŋe species epithet is in honour of the Denmarkbased Carlsberg beer brewery, whose generous grants and beverages have facilitated much research in the Solodovnikov Lab. In addition, the male aedeagus resembles a boưle opener. +The species epithet is in honour of the Denmarkbased Carlsberg beer brewery, whose generous grants and beverages have facilitated much research in the Solodovnikov Lab. In addition, the male aedeagus resembles a bottle opener. diff --git a/data/C6/0D/AE/C60DAE358620D837D92CFB3CBED1F9A7.xml b/data/C6/0D/AE/C60DAE358620D837D92CFB3CBED1F9A7.xml new file mode 100644 index 00000000000..7c1a152b3b5 --- /dev/null +++ b/data/C6/0D/AE/C60DAE358620D837D92CFB3CBED1F9A7.xml @@ -0,0 +1,110 @@ + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Author + +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com + + + +Author + +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany + + + +Author + +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + + + +Author + +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + +text + + +Zoological Journal of the Linnean Society + + +2024 + +Zool. J. Linn. Soc. + + +2023-10-28 + + +202 + + +1 + + +1 +42 + + + + +https://doi.org/10.1093/zoolinnean/zlad143 + +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding + + + + + + + +edwardsianus + +group + + + + + + + +Included species: +Loncovilius cantharoides + +sp. nov. +, and + +L +. +edwardsianus +(Korge, 1963) + +. + + + + +Diagnosis: +Head elongate, without coarse non-setiferous punctures; nuchal and infraorbital ridges not joined; gula with isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin; PMP strongly reduced or rarely absent; females with a11 slightly longer than a10 (a11:a10 ratio <1.5). Pronotum without SLSP. Tergites III and IV with only a few rows of coarse, sparse punctures, with large impunctate area near apical margin ( +Fig. 9C +); tergite VIII without medial apical emargination; ovipositor, its second gonocoxite with three or more macrosetae medially ( +Fig. 9H +). + + + + \ No newline at end of file diff --git a/data/C6/0D/AE/C60DAE358625D832D915FD87BDEFFC26.xml b/data/C6/0D/AE/C60DAE358625D832D915FD87BDEFFC26.xml new file mode 100644 index 00000000000..d0ad0651082 --- /dev/null +++ b/data/C6/0D/AE/C60DAE358625D832D915FD87BDEFFC26.xml @@ -0,0 +1,126 @@ + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Author + +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com + + + +Author + +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany + + + +Author + +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + + + +Author + +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + +text + + +Zoological Journal of the Linnean Society + + +2024 + +Zool. J. Linn. Soc. + + +2023-10-28 + + +202 + + +1 + + +1 +42 + + + + +https://doi.org/10.1093/zoolinnean/zlad143 + +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding + + + + + + + +germaini + +group + + + + + + +Included species: + +Loncovilius germaini +( +Scheerpeltz, 1933 +) + +, + +L +. +hammondi + +sp. nov. +, and + +L +. +impunctus + +sp. nov. + + + + +Diagnosis: +Head transverse, with coarse non-setiferous punctures; nuchal ridge complete dorsally, nuchal and infraorbital ridges joined; postmandibular ridge distinctly separated from eye margin and well-developed PMP ( +Fig. 8E +; + +Brunke +et al +. 2019 + +: fig. 1); right mandible with distal tooth bifid; left mandible only with proximal, non-bifid tooth. Pronotum transverse, outline evenly curved. Tergites III and IV with two patches of punctures of moderate size and density, with a middle impunctate line ( +Fig. 9A +); ovipositor, its second gonocoxite without macroseta medially ( +Fig. 9F +). + + + + \ No newline at end of file diff --git a/data/C6/0D/AE/C60DAE358626D831D906FAD1BCD4FF3D.xml b/data/C6/0D/AE/C60DAE358626D831D906FAD1BCD4FF3D.xml new file mode 100644 index 00000000000..2ba40b9462a --- /dev/null +++ b/data/C6/0D/AE/C60DAE358626D831D906FAD1BCD4FF3D.xml @@ -0,0 +1,107 @@ + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Author + +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com + + + +Author + +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany + + + +Author + +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + + + +Author + +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + +text + + +Zoological Journal of the Linnean Society + + +2024 + +Zool. J. Linn. Soc. + + +2023-10-28 + + +202 + + +1 + + +1 +42 + + + + +https://doi.org/10.1093/zoolinnean/zlad143 + +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding + + + + + + + +variabilis + +group + + + + + + + +Included species: +Loncovilius variabilis + +sp. nov +.. + + + + +Diagnosis: +Head usually elongate, without coarse non-setiferous punctures; nuchal ridge complete dorsally, joined with infraorbital ridge; gula with isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin ( +Fig. 8F +); PMP (usually reduced); females with a11 distinctly longer than a10 (a11:a10 ratio>1.5). Pronotum slightly wider than long, its outline evenly curved. Elytra with even but sparse setiferous punctation on disc and epipleura. Protibiae not sexually dimorphic, in both sexes without laterodorsal row of thick spines. Mesotarsomeres 1–4 not sexually dimorphic, in both sexes with pale adhesive setae and mesotarsomeres +1–3 in +both sexes with terminal plate. Tergites III and IV with several rows of fine punctures of moderate density, without sizeable impunctate area near to posterior margin ( +Fig. 9B +); tergite VIII only slightly emarginate medio-apically in females; ovipositor, its second gonocoxite with one macroseta medially. Aedeagus as in +Figure 10I–K +. + + + + \ No newline at end of file diff --git a/data/C6/0D/AE/C60DAE358637D832D8B7FB26BA9CFE07.xml b/data/C6/0D/AE/C60DAE358637D832D8B7FB26BA9CFE07.xml new file mode 100644 index 00000000000..a872d1015b6 --- /dev/null +++ b/data/C6/0D/AE/C60DAE358637D832D8B7FB26BA9CFE07.xml @@ -0,0 +1,1115 @@ + + + +Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae) + + + +Author + +Reyes-Hernández, José L. +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark +jl.reyeshdez@gmail.com + + + +Author + +Hansen, Aslak Kappel +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark & Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Center for Integrative Biodiversity Discovery, Berlin, Germany + + + +Author + +Shaw, Josh Jenkins +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + + + +Author + +Solodovnikov, Alexey +Natural History Museum of Denmark, University of Copenhagen, Zoological Museum, Universitetsparken 15, 2100, Copenhagen, Denmark + +text + + +Zoological Journal of the Linnean Society + + +2024 + +Zool. J. Linn. Soc. + + +2023-10-28 + + +202 + + +1 + + +1 +42 + + + + +https://doi.org/10.1093/zoolinnean/zlad143 + +journal article +10.1093/zoolinnean/zlad143 +0024-4082 +14443945 +CE2383A-68A1-40A0-8F48-1271F96F86F1Corresponding + + + + + + +Genus + +Loncovilius +Germain, 1903 + +sensu nov. + + + + + + + +( +Figs 4–15 +) + + + + + + +Loncovilius +Germain 1903: 439 + +(original description; species included: + +semiflavus + +, + +lividipennis + +); +Bernhauer and Schubert 1916: 417 +(subgenus of + +Quedius + +); +Blackwelder 1952: 226 +( +type +species + +Loncovilius semiflavus + +, by subsequent designation); +Coiffait and Sáiz 1966: 401 +(characters; key to Chilean species); +Sáiz 1971: 387 +, 389 (notes on biogeography; notes on phenetic relationships). + + + + + +Type +species: + + +Loncovilius semiflavus +( +Fairmaire and Germain 1862 +) + +. + + + + + +Included species: +Loncovilius barclayi + +sp. nov. +, + +L +. +cantharoides + +sp. nov. +, + +L. carlsbergi + +sp. nov. +, + +L +. +edwardsianus +(Korge, 1963) + +, + +L +. +germaini +( +Scheerpeltz, 1933 +) + +, + +L +. +hammondi + +sp. nov. +, + +L +. +impunctus + +sp. nov. +, + +L +. +lividipennis +( +Fairmaire and Germain, 1862 +) + +, + +L +. +semiflavus +( +Fairmaire and Germain, 1862 +) + +, and + +Loncovilius variabilis + +sp. nov +.. + + + + + +Diagnosis: +Loncovilius + +can be distinguished from all other genera in +Amblyopinini +based on the combination of the following characters: frontoclypeal (epistomal) suture complete ( +Fig. 7C–E +); basisternum flat, with few punctures, without pair of macrosetae; with rounded flexible postcoxal hypomeral extension or process; two short empodial setae on each tarsomere 5; ventral side of meso- and metatarsi in males and females with pale adhesive setae ( +Fig. 7F +); abdomen without protergal glands; all tergites without posterior transverse basal carina (PTBC); lateral tergal sclerites IX at least slightly longer than tergite VIII. Free-living; females distinctly larger than males. + + + + +Description: +Frontoclypeal (epistomal) suture complete; neck constriction fully developed; neck moderately wide; nuchal ridge latero-ventrally extended anteriad beyond level of postgenal ridge, merged with, or coming very close to, infraorbital ridge; dorsal basal ridge absent; ventral basal ridge extending more or less parallel to ventral portion of postoccipital suture; postgenal ridge partially or completely missing; no extra ridge anterior to postgenal ridge; postmandibular ridge short; infraorbital ridge incomplete; frontoclypeal, anterior frontal, parocular, posterior frontal, and basal punctures present; supra-antennal and interocular punctures absent. To avoid future confusion with the terminology of the setiferous punctures of the head in ventral view we decided to follow that of figure 1 of +Smetana and Davies (2000) +except for its 'genal seta' which we rename here as postmandibular setiferous puncture (PMP) in order to not confuse it with the 'genal puncture' of +Brunke et al. (2019) +which is located in the anterodorsal genal region. PMP is found in the anteroventral genal region, in the area delimited by the margin of the eye, the base of the mandible, and the maxilla, generally associated with the anterior part of the postmandibular ridge when present. In + +Loncovilius + +, the PMP is present, in the + +germaini + +group it is well developed ( +Fig. 8E +), and in the rest of the groups, it is reduced ( +Fig. 8F +). Gula with distinct transverse basal impression; gular sutures (gs) separated from each other. Antennal insertions situated at equal distance to frontoclypeus and to eye (except + +L +. +germaini + +with antennal insertions situated closer to frontoclypeus than to eye); antennae relatively long, all antennomeres elongate, antennomere 1 shorter than or subequal to antennomeres 2 and 3 combined, antennomeres 4 to 11 with tomentose pubescence, males with antennomere 11 (a11) distinctly longer than antennomere (a10) (a11:a10 ratio>1.5), this character varies in females. Mandibles in dorsal view slightly straight with curved base and apex; right mandible with proximal tooth and distal tooth with varying shape; teeth of left mandible vary. In + +germaini + +group, on right mandible, distal tooth bifid, space between proximal and distal teeth narrow and smooth. In + +edwardsianus + +, + +lividipennis + +, + +semiflavus + +, and + +variabilis + +groups, on right mandible, distal tooth not bifid, space between proximal and distal teeth wide and rough; in germaini group left mandible only with proximal, not bifid tooth; in + +edwardsianus + +, + +lividipennis + +, + +semiflavus + +, and + +variabilis + +groups left mandible with proximal and distal teeth not bifid, the latter short and truncated, space between teeth rough, in lateral view right and left mandibles deflexed ventrally at an angle of 25–30° +c +. one-fourth to one-third distance from base to apex; ligula small and entire; labrum with transparent apical membrane; maxillary palpomere 4 (mp4, apical) more or less subconical, with evenly narrowed apex, as long as or longer than palpomere 2; preapical maxillary palpomere 3 only weakly wider than apical palpomere, about as long as mp4; labial palpomere 2 with sparse brushes of setae. + + +Ŋorao +: Chaetotaxy of pronotum varies between species groups. Paired punctures in dorsal series ( +PPDS +) may be present in different combinations; sublateral setiferous punctures ( +SLSP +) defined here as punctures found in the area between PPDS, anterior marginal row of setiferous punctures ( +AMRSP +), large lateral setiferous puncture ( +LLSP +) and large posterior setiferous puncture ( +LPSP +). + +Loncovilius carlsbergi + +sp. nov. +has the most complete chaetotaxy set where all +types +of setiferous punctures occur ( +Fig. 9A +). Pronotum and prosternum not fused, pronotosternal suture complete even in procoxal cavity; postcoxal hypomeral translucent process present, rounded or triangular; sternacostal ridge not protruding, not joining with superior line of hypomeron; basisternum (bs) flat with few punctures, triangular, its lateral arms narrowed subapically, without pair of macrosetae, bs distinctly longer than furcasternum (fs) (bs/fs ratio>1.5). Elytra without humeral spines or spine-like setae; mesoscutellum setose without posterior scutellar ridge or subbasal ridge; hind wings with veins CuA and MP4 fused in one vein, vein MP3 absent; metascutellum with mid-longitudinal suture; apex of intercoxal process in mesothorax rounded or broadly pointed, forming obtuse angle; anterior margin of mesoventrite confluent with distinctly oblique sternopleural suture; posterior border of mesocoxal cavities complete; mesometasternal suture present. Tarsal formula 5-5-5; protibiae cylindrical to slightly broadened apically, ventrally with a middle long spine directed anteriad; protarsomeres 1–3 distinctly wider than long, bilobed, and ventrally with pale adhesive setae; ventral side of meso- and metatarsomeres +1–4 in +males and females with pale adhesive setae; pale adhesive setae on meso- and metatarsi with terminal plate ( +Fig. 8B +) or without ( +Fig. 8C +); all legs with a pair of empodial setae distinctly shorter than claws; procoxae with internal and external ridges present, internal not running along external ridge, ending distinctly before or nearly joining external ridge; metacoxae with no more than four spines on posterior surface; mesotarsal segments with pale adhesive setae. + + + +Figure 3. +Ecological niche modelling of + +Loncovilius +species. A + +–D, potential suitable areas predicted with free extrapolation in South America. Binarized projection with 5% threshold for suitability values and mobility-oriented parity analysis (MOP) showing areas for which current model predictions are strictly based on extrapolation. E–H, predictions under shared socio-economic pathways (SSPs) 126, and agreement between different global climate models (GCMs), and MOP agreement among GCMs with strict extrapolative conditions under the low emissions. I–L, predictions under SSPs 585, agreement between different GCMs, and MOP agreement among GCMs with strict extrapolative conditions under the high emissions. + + + + +Figure 4. +Distribution of + +Loncovilius +species. A + +, + +semiflavus + +group. B, + +lividipennis + +group. C, + +germaini + +group. D, + +edwardsianus + +group. E, + +variabilis + +group. + + + +Abdomen +: Protergal glands absent; tergites III–VI with anterior transverse basal carina (ATBC) laterally straight continuing to paratergites, tergite VII with ATBC not continuing to paratergite, tergites VII and VIII with ATBC descending laterally without reaching tergite margin ( +Fig. 9D +); all tergites without PTBC; anterior margin of tergites and sternites III–VI entire; tergite VII with white fringe along posterior edge; tergite IX in males at least slightly longer than tergite VIII; apical margin of male sternite VIII with medial emargination; basal portion of male sternite IX asymmetrical; paramere fused to median lobe only at base and very closely appressed to median lobe along entire length, often of complex and diverse shape, with minute pale sensory peg setae or usual peg setae; internal sac often with visible sclerites; ovipositor with two paired gonocoxites and minute styli. + + + + +Distribution and habitat: +The genus is only known from +Chile +and +Argentina +; in +Chile +from the regions of +Araucanía +, +Aysén +, +Los Lagos +, +Los Ríos +, +Magallanes +and Chilean +Antarctica +, Maule, Ñuble, O’Higgins, and from Santiago Metropolitan; in +Argentina +from the provinces of +Chubut +, +Neuquén +and +Río Negro +( +Fig. 4 +). The genus occurs in the area from lowlands to +2600 m +elevation in the mountains, with an average annual rainfall of between 500 and +3000 mm +and an average annual temperature of between 1°C and 12°C. It is confined to various +types +of forests in the Chilean Matorral, Magellanic subpolar, and Valdivian temperate forest ecoregions. It has been collected using Malaise and flight intercept traps, sifting moss, dead wood, and leaf litter, beating and directly from flowers of the families +Cunoniaceae +, +Ericaceae +, +Loranthaceae +, +Myrtaceae +, and +Winteraceae +. + + + + +Figure 5. +Female habitus of + +Loncovilius +species. A + +, + +L +. +germaini + +. B, + +L +. +hammondi + +. C, + +L +. +impunctus + +. D, + +L +. +variabilis + +. + + + + +Figure 6. +Habitus of + +Loncovilius +species. A + +, male of + +L +. +edwardsianus + +. B, male of + +L +. +cantharoides + +. C, female of + +L +. +semiflavus + +. D, male of + +L +. +barclayi + +. + + + + +Figure 7. +Female habitus of + +Loncovilius +species. A + +, + +L +. +lividipennis + +. B, + +L +. +carlsbergi + +. Head. C, + +L +. +hammondi + +. D, + +L +. +semiflavus + +. E, + +L +. +carlsbergi + +. F, Male mesotarsi of + +L +. +cantharoides + +. + + + + +Figure 8. +A, pronotum of + +L +. +carlsbergi + +. AMRSP = anterior marginal row of setiferous punctures, APP = additional paired punctures adjacent to AMRSP, LLSP = large lateral setiferous puncture, LPSP = large posterior setiferous puncture, PPDS = paired punctures on dorsal serial, SLSP = sublateral setiferous puncture. B, + +L +. +cantharoides + +ventral view of mesotarsomere 3 with pale adhesive setae with terminal plate. C, + +L +. +cantharoides + +ventral view of metatarsomere 4, apical margin emarginated (bilobed), with pale adhesive setae without terminal plate. D, + +L +. + +cantharoides +tergites VI and VII. Black arrow up = anterior transverse basal carina (ATBC) laterally straight continuing to paratergites, white arrow down = ATBC descending laterally without reaching tergite margin. Head in ventral view of + +Loncovilius +species. E + +, + +L +. +hammondi + +. F, + +L +. +lividipennis + +. PMP = postmandibular puncture, PMR = postmandibular ridge, EM = eye margin. + + + + +Figure 9. +Details of the abdominal structures of + +Loncovilius +species + +; tergites III and IV (A–E); ovipositor (F–H). A, + +Loncovilius germaini + +. B, + +L +. +variabilis + +. C, + +L +. +edwardsianus + +. D, + +L +. +semiflavus + +. E, + +L +. +lividipennis + +. F, + +L +. +impunctus + +. G, + +L +. +semiflavus + +. H, + +L +. +edwardsianus + +. + + + + +Figure 10. +Aedeagus of + +Loncovilius +species. A + +–C, + +Loncovilius germaini + +. D–F, + +L +. +hammondi + +. G, H, + +L +. +impunctus + +. I–K, + +L +. +variabilis + +. L–N, + +L +. +edwardsianus + +. O–Q, + +L +. +cantharoides + +. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. + +C, F, H, K, N, Q, aedeagus laterally. + + + +Figure 11. +Aedeagus of + +Loncovilius +species. A + +–C, + +L +. +semiflavus + +. D–F, + +L +. +barclayi + +. G, H, + +L +. +lividipennis + +. I, J, + +L +. +carlsbergi + +. A, D, G, I, aedeagus dorsally (parameral side). B, E, apical portion of paramere, underside. C, F, H, J, aedeagus laterally. + + + + +Figure 12. +Male tergite X of + +Loncovilius +species. A + +, + +L +. +germaini + +. B, + +L +. +hammondi + +. C, + +L +. +impunctus + +. D, + +L +. +variabilis + +. E, + +L +. +edwardsianus + +. + + +F, + +L +. +cantharoides + +. G, + +L +. +semiflavus + +. H, + +L +. +barclayi + +. I, + +L +. +lividipennis + +. J, + +L +. +carlsbergi + +. + + + + +Figure 13. +Female tergite X of + +Loncovilius +species. A + +, + +Loncovilius germaini + +. B, + +L +. +hammondi + +. C, + +L +. +impunctus + +. D, + +L +. +variabilis + +. E, + +L +. +edwardsianus + +. F, + +L +. +cantharoides + +. G, + +L +. +semiflavus + +. H, + +L +. +barclayi + +. I, + +L +. +lividipennis + +. J, + +L +. +carlsbergi + +. + + + + +Figure 14. +Male sternite VIII of + +Loncovilius +species. A + +, + +Loncovilius germaini + +. B, + +L +. +hammondi + +. C, + +L +. +impunctus + +. D, + +L +. +variabilis + +. E, + +L +. +edwardsianus + +. F, + +L +. +cantharoides + +. G, + +L +. +semiflavus + +. H, + +L +. +barclayi + +. I, + +L +. +lividipennis + +. J, + +L +. +carlsbergi + +. + + + + +Remarks: +The monophyly of + +Loncovilius + +is supported by two unique synapomorphies: females with pale adhesive setae present at mesotarsal segments 2–4 [(character 69, state 1; reversed condition in + +variabilis + +and + +edwardsianus + +groups (character 69, state 0)]; tergite IX at least slightly longer than tergite VIII (ratio up to 1.1, character 92, state 1); and by five homoplastic synapomorphies: eyes small [(eye/head length ratio more than three-tenths but less than half, character 37, state 1; in + +semiflavus + +group and + +L +. +impunctus + +eyes of medium size (eyes/head length ratio more than half but less than three-quarters, character 37, state 2)]; metascutellar mid-longitudinal suture well developed (character 82, state 0); protergal glands absent (character 84, state 0); female sternite VIII with medial emargination at apical margin [(character 99, state 1; reversed condition in + +variabilis + +and + +edwardsianus + +groups (character 100, state 0)]; paramere(s) with black sensory peg setae [(character 105, state 1; reversed condition in + +edwardsianus + +group (character 105, state 0)]. + + + + +Key to the species of +Loncovilius + + + + +1. Head transverse, round in outline; disc with coarse non-setiferous punctures (Fig. 7C); postmandibular ridge distinctly separated from eye margin; PMP conspicuous (Fig. 8E); mandibles with a shallow dorso-lateral groove; right mandible with distal tooth bifid; left mandible only with proximal, non-bifid tooth. Abdominal tergites III–V with two patches of setiferous punctures separated by an impunctate area (Fig. 9A); second gonocoxite without macrosetae (Fig. 9F)..................................2 + + +- Head about as wide as long or elongate, its outline mostly longitudinally elliptical; disc without coarse non-setiferous punctures (Fig. 7D, E); postmandibular ridge situated very close to eye margin; PMP reduced and hard to see (Fig. 8F); mandibles without dorso-lateral groove; right mandible with distal tooth not bifid; left mandible with proximal and distal teeth, the latter not bifid, short and truncated, space between proximal and distal teeth rough. Abdominal tergites III–V not as described above; second gonocoxite with at least one macroseta (Fig. 9G, H) ................................................................................4 + + + + + +2. Disc of head without microsculpture; antennal insertions situated closer to frontoclypeus than to eye. Aedeagus as in Figure 10A–C........................................................................................................................................ +Loncovilius germaini +( +Scheerpeltz, 1933 +) + + + +- Disc of head with microsculpture; antennal insertions situated at equal distance to frontoclypeus and to the eye, or closer to the eye. Aedeagus not as above ....................................................................................................................................................................3 + + + + + +3. Microsculpture on dorsal surface of head meshed to isodiametric. Pronotum and elytral disc with few and scattered coarse non-setiferous punctures. Aedeagus as in Figure 10G, H ................................................................. +Loncovilius impunctus +sp. nov. + + + + +- Microsculpture on dorsal surface of head as transverse waves. Pronotum and elytral disc with conspicuous coarse non-setiferous punctures. Aedeagus as in Figure 10D–F........................................................................... +Loncovilius hammondi +sp. nov. + + + + + +4. Head: nuchal ridge present dorsally and laterally. Abdomen: tergites III and IV with more than three rows of punctures, anterior half punctate (Fig. 9B, E) ...................................................................................................................................................................5 + + +- Head: nuchal ridge missing dorsally, present laterally. Abdomen: tergites III and IV with less than three rows of punctures, anterior half impunctate (Fig. 9C, D).........................................................................................................................................................7 + + + + + +5. Protibia in both sexes without a row of thick laterodorsal spines; first metatarsomere shorter than fifth or at most both tarsomeres subequal. Abdominal tergites III and IV with several rows of sparse and fine punctures of moderate density (Fig. 9B); sternite III with basal transverse carina strongly arcuate; aedeagus as in Figure 10I–K ........................................................... .......................................................................................................................................................................... +Loncovilius variabilis +sp. nov. + + + +- Protibia with a row of thick laterodorsal spines present at least in females; first metatarsomere moderately longer than fifth. Abdominal tergites III and IV with several rows of punctures of moderate size and density (Fig. 9E); sternite III with basal transverse carina projecting medially in a sharp or obtuse angle; aedeagus not as above ...............................................................6 + + + + + +6. Pronotum rarely with additional paired punctures adjacent to anterior marginal row of setiferous punctures (Fig. 8A). Abdominal sternite III with basal transverse carina sharply pointed medioapically; aedeagus as in +Figure 11G, H +........................ ........................................................................................................................... +Loncovilius lividipennis +( +Fairmaire and Germain, 1862 +) + + + + +- Pronotum always with additional paired punctures adjacent to anterior marginal row of setiferous punctures (Fig. 8A). Abdominal sternite III with basal transverse carina evenly converging at an obtuse angle; aedeagus as in +Figure 11I, J +.......................................................................................................................................................................... +Loncovilius carlsbergi +sp. nov. + + + + + +7. Head and pronotum microsculpture consisting of isodiametric meshes. Head elongate. Pronotum without sublateral setiferous punctures. Males: aedeagus, paramere strongly produced over apex of median lobe. Females: second gonocoxite with three or more macrosetae (Fig. 9G) ...........................................................................................................................................................8 + + +- Head and pronotum with very fine microsculpture of long transverse and diagonal waves. Head about as wide as long. Pronotum with sublateral setiferous punctures. Males: aedeagus, paramere slightly shorter or produced over apex of median lobe. Females: second gonocoxite with one or two macrosetae (Fig. 9H).........................................................................................9 + + + + + +8. Head with small setiferous punctures between frontoclypeal and anterior frontal punctures. Pronotum outline evenly curved. Protibia with a row of thick laterodorsal spines only in females. Elytra rarely with weak bright metallic iridescence .................................................................................................................................................................... +Loncovilius cantharoides +sp. nov. + + + + +- Head without small setiferous punctures between frontoclypeal and anterior frontal punctures. Pronotum outline hexagonal. Protibia in both sexes without a row of thick laterodorsal spines. Elytra with bright greenish, bluish, or purplish metallic iridescence (Figs 1A, 6A)......................................................................................... +Loncovilius edwardsianus +(Korge, 1963) + + + + + + +9 Elytra with brown-black marking around mesoscutellum and on suture (Fig. 6D). All femora reddish-brown. Male sternite IX without median emargination at apex; male tergite X trilobed (Fig.12H); aedeagus as in +Figure 11D–F +.............................. ............................................................................................................................................................................ +Loncovilius barclayi +sp. nov. + + + + +- Elytra without dark mark around mesoscutellum and on suture. All femora yellow or yellowish-brown; male sternite IX with median emargination at apex; male tergite X emarginate medio-apically (Fig. 12G); aedeagus as in +Figure 11A–C +............................................................................................................................. +Loncovilius semiflavus +( +Fairmaire and Germain, 1862 +) + + + + + + + \ No newline at end of file