Add updates up until 2025-03-12 13:06:56

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<document id="9E0B933343085A173150064FF5CB424A" ID-DOI="10.11646/zootaxa.5604.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="15012448" ID-ZooBank="72F4414F-7769-4AE6-ACBC-14577BC32AB3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1741784003344" checkinUser="plazi" docAuthor="Gomes-Silva, José Geovane &amp; Dasilva, Marcio B." docDate="2025" docId="03A31D382841066304E1FA20C49BFD7D" docLanguage="en" docName="zootaxa.5604.1.2.pdf" docOrigin="Zootaxa 5604 (1)" docSource="https://doi.org/10.11646/zootaxa.5604.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goniosoma bragagnolo Gomes-Silva &amp; Dasilva, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="20" masterDocId="FF9A6540284006670476FFEBC76CFC3C" masterDocTitle="Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil." masterLastPageNumber="28" masterPageNumber="16" pageNumber="17" updateTime="1741784440243" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C247EF05ACF26C5D828BCBCACC764134">Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil.</mods:title>
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<mods:namePart id="01CFC2F5916338A8001545DD78D2F41B">Gomes-Silva, José Geovane</mods:namePart>
<mods:nameIdentifier id="C8C933E749FA6BA014F556B530D38903" type="ORCID">0009-0002-9811-573X</mods:nameIdentifier>
<mods:affiliation id="BE80435F549BFBAB1DCDB54FCBB69C4F">Departamento de Sistemática e Ecologia, Centro de Ciências Exatas e da Natureza, Universidade Federal da Paraíba, João Pessoa, Brasil</mods:affiliation>
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<mods:namePart id="309DE041E236C9044DC9EFE95DE8D18B">Dasilva, Marcio B.</mods:namePart>
<mods:nameIdentifier id="B43BD4FF5EC44F4EF42E321AA1A71007" type="ORCID">0000-0002-2824-1071</mods:nameIdentifier>
<mods:affiliation id="50D8311CEE4217E3F5DFFF1C6D5CD3C7">Departamento de Sistemática e Ecologia, Centro de Ciências Exatas e da Natureza, Universidade Federal da Paraíba, João Pessoa, Brasil</mods:affiliation>
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<mods:date id="F55ABFBE77BA0375B5C4160BD70E43B3">2025</mods:date>
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<treatment id="03A31D382841066304E1FA20C49BFD7D" ID-DOI="http://doi.org/10.5281/zenodo.15012473" ID-Zenodo-Dep="15012473" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D" httpUri="http://treatment.plazi.org/id/03A31D382841066304E1FA20C49BFD7D" lastPageId="4" lastPageNumber="20" pageId="1" pageNumber="17">
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<paragraph id="8BB5AC2E2841066604E1FA20C446F9D8" blockId="1.[151,810,1482,1509]" box="[151,810,1482,1509]" pageId="1" pageNumber="17">
<heading id="D0FD1B422841066604E1FA20C446F9D8" bold="true" box="[151,810,1482,1509]" fontSize="11" level="1" pageId="1" pageNumber="17" reason="1">
<emphasis id="B97E703C2841066604E1FA20C446F9D8" bold="true" box="[151,810,1482,1509]" pageId="1" pageNumber="17">
<taxonomicName id="4C0AD7AD2841066604E1FA20C6D8F9D9" authorityName="Gomes-Silva &amp; Dasilva" authorityYear="2025" box="[151,436,1482,1509]" class="Arachnida" family="Gonyleptidae" genus="Goniosoma" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="bragagnolo" status="sp. nov.">
<emphasis id="B97E703C2841066604E1FA20C6D8F9D9" bold="true" box="[151,436,1482,1509]" italics="true" pageId="1" pageNumber="17">Goniosoma bragagnolo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24DCD472841066605CDFA20C575F9D9" box="[443,537,1483,1509]" pageId="1" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1331B0AB28410666065EFA21C510F9D8" box="[552,636,1482,1508]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1930]" captionTargetBox="[179,1402,202,1856]" captionTargetId="figure-17@2.[170,1416,181,1882]" captionTargetPageId="2" captionText="FIGURE 1. AG male holotype (MZSP 31022) of Goniosoma bragagnolo sp. nov. AB habitus: A) lateral view; B) dorsal view; C) pedipalp, ventral-retrolateral view; D) femur of pedipalp, ventral view, showing prolateral seta; E) trochanter and femur IV, dorsal view; F) tibia IV, ventral-retrolateral view, showing armature of row 2 and 3 (arrows); G) metatarsus IV, ventral-retrolateral view, showing armature of row 3. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.15012450" httpUri="https://zenodo.org/record/15012450/files/figure.png" pageId="1" pageNumber="17">Figs. 1</figureCitation>
,
<figureCitation id="1331B0AB2841066606FCFA21C5F6F9D8" box="[650,666,1482,1508]" captionStart="FIGURE 3" captionStartId="6.[151,250,1601,1625]" captionTargetBox="[165,1422,187,1562]" captionTargetId="figure-17@6.[151,1436,181,1577]" captionTargetPageId="6" captionText="FIGURE 3. AB penis of male holotype (MZSP 31022) of Goniosoma bragagnolo sp. nov.: A) dorsal view; B) lateral view. CD penis of male holotype (MZSP 18974) of Serracutisoma hara sp. nov.: C) dorsal view; D) lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.15012456" httpUri="https://zenodo.org/record/15012456/files/figure.png" pageId="1" pageNumber="17">3</figureCitation>
,
<figureCitation id="1331B0AB2841066606D1FA21C5DBF9D8" box="[679,695,1482,1508]" captionStart="FIGURE 4" captionStartId="7.[151,250,662,686]" captionTargetBox="[151,1436,181,638]" captionTargetId="figure-30@7.[151,1436,181,638]" captionTargetPageId="7" captionText="FIGURE 4. Photographs of live specimens of Goniosoma bragagnolo sp. nov., female (left) and male (right)." figureDoi="http://doi.org/10.5281/zenodo.15012460" httpUri="https://zenodo.org/record/15012460/files/figure.png" pageId="1" pageNumber="17">4</figureCitation>
,
<figureCitation id="1331B0AB2841066606B3FA20C5B8F9D9" box="[709,724,1483,1509]" captionStart="FIGURE 5" captionStartId="7.[151,250,1712,1736]" captionTargetBox="[156,1433,786,1685]" captionTargetId="figure-49@7.[152,1436,782,1688]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Goniosoma bragagnolo sp. nov. and Serracutisoma hara sp. nov. along with their respective genera. Black lines: limits of areas of endemism. SEsp = Serra do Espinhaço; SC = Santa Catarina; solid lines: Congruence Core; dashed lines: Maximum Region of Endemism." figureDoi="http://doi.org/10.5281/zenodo.15012463" httpUri="https://zenodo.org/record/15012463/files/figure.png" pageId="1" pageNumber="17">5</figureCitation>
and
<figureCitation id="1331B0AB284106660767FA21C44CF9D8" box="[785,800,1482,1508]" captionStart="FIGURE 6" captionStartId="8.[151,250,1113,1137]" captionTargetBox="[155,1433,184,1087]" captionTargetId="figure-17@8.[151,1436,181,1089]" captionTargetPageId="8" captionText="FIGURE 6. Distribution of Goniosoma bragagnolo sp. nov. and Serracutisoma hara sp. nov. along with their respective genera, across the vegetation types of the Atlantic Rain Forest." figureDoi="http://doi.org/10.5281/zenodo.15012465" httpUri="https://zenodo.org/record/15012465/files/figure.png" pageId="1" pageNumber="17">6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C310FFA52841066604E1F9E7C34FFB56" pageId="1" pageNumber="17" type="materials_examined">
<paragraph id="8BB5AC2E2841066604E1F9E7C34FFB56" blockId="1.[151,1437,1548,2043]" pageId="1" pageNumber="17">
<emphasis id="B97E703C2841066604E1F9E7C62BFA1A" bold="true" box="[151,327,1548,1574]" pageId="1" pageNumber="17">Type material:</emphasis>
<materialsCitation id="3B62A673284106660538F9E7C373FB56" collectingDate="2002-05-01" collectingDateMax="2009-05-05" collectingDateMin="2002-05-01" collectionCode="MZSP" collectorName="DaSilva &amp; Bragagnolo &amp; Yamaguti &amp; Nogueira" county="Vale Verde" elevation="1820" latitude="-20.478611" location="Alto Caparao" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" pageId="1" pageNumber="17" specimenCode="MZSP 31022, MZSP 21732, MZSP 29995, MZSP 31024, MZSP 31023, MZSP 77541" specimenCount="28" specimenCount-female="8" specimenCount-immature="1" specimenCount-male="19" stateProvince="Minas Gerais" typeStatus="holotype">
male
<typeStatus id="54B1128C2841066605FAF9E7C69DFA1A" box="[396,497,1548,1574]" pageId="1" pageNumber="17" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D0C67A728410666058AF9E7C53EFA1A" box="[508,594,1548,1574]" count="6" pageId="1" pageNumber="17" type="male">6 males</specimenCount>
and
<specimenCount id="9D0C67A72841066606FFF9E7C599FA1A" box="[649,757,1548,1574]" count="4" pageId="1" pageNumber="17" type="female">4 females</specimenCount>
<typeStatus id="54B1128C28410666068AF9E6C40BFA1B" box="[764,871,1549,1575]" pageId="1" pageNumber="17" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DBAC0455284106660700F9E7C363FA1A" box="[886,1039,1548,1574]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="17" type="Museum">MZSP 31022</specimenCode>
) from
<collectingCounty id="62D4D4A228410666002DF9E7C3BBFA1A" box="[1115,1239,1548,1574]" pageId="1" pageNumber="17">Vale Verde</collectingCounty>
,
<collectingMunicipality id="6BD13654284106660094F9E6C64DFA76" pageId="1" pageNumber="17">Parque Nacional do Caparaó</collectingMunicipality>
,
<location id="8ED5FAF5284106660547F9DBC6BCFA76" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF5284106660547F9DBC6BCFA76" box="[305,464,1584,1610]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Alto Caparao" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Alto Caparaó</location>
,
<collectingRegion id="49CE62CC284106660594F9DBC5EFFA76" box="[482,643,1584,1610]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="17">Minas Gerais</collectingRegion>
.
<geoCoordinate id="EE3ECAE92841066606E5F9DBC474FA77" box="[659,792,1584,1611]" degrees="20" direction="south" minutes="25" orientation="latitude" pageId="1" pageNumber="17" precision="15" seconds="10" value="-20.419443">20°2510”S</geoCoordinate>
<geoCoordinate id="EE3ECAE9284106660753F9DBC4DCFA77" box="[805,944,1584,1611]" degrees="41" direction="west" minutes="50" orientation="longitude" pageId="1" pageNumber="17" precision="15" seconds="45" value="-41.845833">41º5045”W</geoCoordinate>
.
<date id="FFB48AEE2841066607B2F9DBC324FA76" box="[964,1096,1584,1610]" pageId="1" pageNumber="17" value="2009-04-06">
<collectingDate id="EFF073062841066607B2F9DBC324FA76" box="[964,1096,1584,1610]" pageId="1" pageNumber="17" value="2009-04-06">06/IV/2009</collectingDate>
</date>
;
<location id="8ED5FAF528410666002CF9DBC261FA76" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF528410666002CF9DBC261FA76" box="[1114,1293,1584,1610]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Pinto-da-Rocha" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Pinto-da-Rocha</location>
,
<collectorName id="26FFC9F8284106660169F9DBC215FA76" box="[1311,1401,1584,1610]" pageId="1" pageNumber="17">DaSilva</collectorName>
&amp;
<collectorName id="26FFC9F82841066604E1F9BFC671FA52" box="[151,285,1620,1646]" pageId="1" pageNumber="17">Bragagnolo</collectorName>
;
<specimenCount id="9D0C67A728410666055AF9BFC6EAFA52" box="[300,390,1620,1646]" count="2" pageId="1" pageNumber="17" type="male">2 males</specimenCount>
and
<specimenCount id="9D0C67A72841066605B2F9BFC544FA52" box="[452,552,1620,1646]" count="1" pageId="1" pageNumber="17" type="female">1 female</specimenCount>
<typeStatus id="54B1128C284106660645F9BEC5F2FA53" box="[563,670,1621,1647]" pageId="1" pageNumber="17" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DBAC04552841066606C6F9BFC421FA52" box="[688,845,1620,1646]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="17" type="Museum">MZSP 21732</specimenCode>
) from
<location id="8ED5FAF52841066607D6F9BEC382FA52" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF52841066607D6F9BEC382FA52" box="[928,1262,1620,1647]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Parque Nacional do Caparao" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Parque Nacional do Caparaó</location>
,
<location id="8ED5FAF528410666008AF9BFC2F4FA52" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF528410666008AF9BFC2F4FA52" box="[1276,1432,1620,1646]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Alto Caparao" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Alto Caparaó</location>
,
<collectingRegion id="49CE62CC2841066604E1F993C659FAAE" box="[151,309,1656,1682]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="17">Minas Gerais</collectingRegion>
.
<date id="FFB48AEE284106660534F993C68BFAAF" box="[322,487,1656,1683]" pageId="1" pageNumber="17" value="2002-05-01" valueMax="2002-05-05" valueMin="2002-05-01">
<collectingDate id="EFF07306284106660534F993C68BFAAF" box="[322,487,1656,1683]" pageId="1" pageNumber="17" value="2002-05-01" valueMax="2002-05-05" valueMin="2002-05-01">0105/V/2002</collectingDate>
</date>
;
<location id="8ED5FAF5284106660580F993C5FCFAAE" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF5284106660580F993C5FCFAAE" box="[502,656,1656,1682]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Equipe Biota" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Equipe Biota</location>
;
<specimenCount id="9D0C67A72841066606E9F993C597FAAE" box="[671,763,1656,1682]" count="4" pageId="1" pageNumber="17" type="male">4 males</specimenCount>
,
<specimenCount id="9D0C67A728410666077CF993C403FAAE" box="[778,879,1656,1682]" count="1" pageId="1" pageNumber="17" type="female">1 female</specimenCount>
and
<specimenCount id="9D0C67A72841066607DAF993C35CFAAE" box="[940,1072,1656,1682]" count="1" pageId="1" pageNumber="17" type="immature">1 immature</specimenCount>
<typeStatus id="54B1128C28410666004CF992C3C9FAAF" box="[1082,1189,1657,1683]" pageId="1" pageNumber="17" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DBAC04552841066600CEF993C238FAAF" box="[1208,1364,1656,1683]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="17" type="Museum">MZSP 29995</specimenCode>
) from
<location id="8ED5FAF52841066604E1F976C621FA8A" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF52841066604E1F976C621FA8A" box="[151,333,1692,1719]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Parque Estadual" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Parque Estadual</location>
<taxonomicName id="4C0AD7AD284106660523F977C6FCFA8A" box="[341,400,1692,1718]" genus="Serra" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="1" pageNumber="25" phylum="Mollusca" rank="genus">Serra</taxonomicName>
do
<location id="8ED5FAF52841066605CAF977C55BFA8A" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF52841066605CAF977C55BFA8A" box="[444,567,1692,1718]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Brigadeiro" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Brigadeiro</location>
,
<location id="8ED5FAF5284106660635F977C5ABFA8A" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF5284106660635F977C5ABFA8A" box="[579,711,1692,1718]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Fervedouro" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Fervedouro</location>
,
<collectingRegion id="49CE62CC2841066606A5F977C402FA8A" box="[723,878,1692,1718]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="17">Minas Gerais</collectingRegion>
.
<date id="FFB48AEE28410666070FF977C34AFA8B" box="[889,1062,1692,1719]" pageId="1" pageNumber="17" value="2005-03-21" valueMax="2005-03-24" valueMin="2005-03-21">
<collectingDate id="EFF0730628410666070FF977C34AFA8B" box="[889,1062,1692,1719]" pageId="1" pageNumber="17" value="2005-03-21" valueMax="2005-03-24" valueMin="2005-03-21">2124/III/2005</collectingDate>
</date>
;
<location id="8ED5FAF5284106660045F977C3FCFA8A" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF5284106660045F977C3FCFA8A" box="[1075,1168,1692,1718]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="DaSilva" pageId="1" pageNumber="17" stateProvince="Minas Gerais">DaSilva</location>
,
<collectorName id="26FFC9F82841066600EDF977C26BFA8A" box="[1179,1287,1692,1718]" pageId="1" pageNumber="17">Yamaguti</collectorName>
&amp;
<collectorName id="26FFC9F828410666015AF977C2F4FA8A" box="[1324,1432,1692,1718]" pageId="1" pageNumber="17">Nogueira</collectorName>
;
<specimenCount id="9D0C67A72841066604E1F92BC78EFAE6" box="[151,226,1728,1754]" count="1" pageId="1" pageNumber="17" type="male">1 male</specimenCount>
<typeStatus id="54B1128C28410666049FF92AC625FAE7" box="[233,329,1729,1755]" pageId="1" pageNumber="17" type="paratype">paratype</typeStatus>
(
<specimenCode id="DBAC045528410666052EF92BC69DFAE6" box="[344,497,1728,1754]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="17" type="Museum">MZSP 31024</specimenCode>
) from
<location id="8ED5FAF528410666064BF92BC46FFAE6" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF528410666064BF92BC46FFAE6" box="[573,771,1728,1754]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Cachoeira Bonita" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Cachoeira Bonita</location>
,
<location id="8ED5FAF5284106660778F92AC33DFAE6" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF5284106660778F92AC33DFAE6" box="[782,1105,1728,1755]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Parque Nacional do Caparao" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Parque Nacional do Caparaó</location>
,
<location id="8ED5FAF528410666002DF92BC398FAE6" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF528410666002DF92BC398FAE6" box="[1115,1268,1728,1754]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Alto Caparao" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Alto Caparaó</location>
,
<collectingRegion id="49CE62CC284106660089F92BC2F5FAE6" box="[1279,1433,1728,1754]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="17">Minas Gerais</collectingRegion>
.
<geoCoordinate id="EE3ECAE92841066604E1F90FC670FAC2" box="[151,284,1764,1790]" degrees="20" direction="south" minutes="24" orientation="latitude" pageId="1" pageNumber="17" precision="15" seconds="24" value="-20.406666">20°2424”S</geoCoordinate>
<geoCoordinate id="EE3ECAE9284106660556F90FC6C1FAC3" box="[288,429,1764,1791]" degrees="41" direction="west" minutes="50" orientation="longitude" pageId="1" pageNumber="17" precision="15" seconds="20" value="-41.838886">41°5020”W</geoCoordinate>
.
<date id="FFB48AEE2841066605CFF90FC552FAC2" box="[441,574,1764,1790]" pageId="1" pageNumber="17" value="2009-04-07">
<collectingDate id="EFF073062841066605CFF90FC552FAC2" box="[441,574,1764,1790]" pageId="1" pageNumber="17" value="2009-04-07">07/IV/2009</collectingDate>
</date>
;
<location id="8ED5FAF5284106660630F90FC595FAC2" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF5284106660630F90FC595FAC2" box="[582,761,1764,1790]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Pinto-da-Rocha" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Pinto-da-Rocha</location>
,
<collectorName id="26FFC9F8284106660777F90FC437FAC2" box="[769,859,1764,1790]" pageId="1" pageNumber="17">DaSilva</collectorName>
&amp;
<collectorName id="26FFC9F828410666070CF90FC36CFAC2" box="[890,1024,1764,1790]" pageId="1" pageNumber="17">Bragagnolo</collectorName>
;
<specimenCount id="9D0C67A728410666007FF90FC33EFAC2" box="[1033,1106,1764,1790]" count="1" pageId="1" pageNumber="17" type="male">1 male</specimenCount>
<typeStatus id="54B1128C284106660021F90EC3DBFAC3" box="[1111,1207,1765,1791]" pageId="1" pageNumber="17" type="paratype">paratype</typeStatus>
(
<specimenCode id="DBAC04552841066600B5F90FC236FAC2" box="[1219,1370,1764,1790]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="17" type="Museum">MZSP 31023</specimenCode>
) from
<location id="8ED5FAF52841066604E1F8E3C6EBFB1E" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF52841066604E1F8E3C6EBFB1E" box="[151,391,1800,1826]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Cachoeira do Aurelio" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Cachoeira do Aurélio</location>
,
<location id="8ED5FAF52841066605E7F8E2C5BDFB1E" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF52841066605E7F8E2C5BDFB1E" box="[401,721,1800,1827]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Parque Nacional do Caparao" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Parque Nacional do Caparaó</location>
,
<location id="8ED5FAF52841066606AAF8E2C405FB1E" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF52841066606AAF8E2C405FB1E" box="[732,873,1800,1827]" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Espera Feliz" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Espera Feliz</location>
,
<collectingRegion id="49CE62CC284106660704F8E3C367FB1E" box="[882,1035,1800,1826]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="17">Minas Gerais</collectingRegion>
.
<quantity id="4CF201CB284106660062F8E3C300FB1E" box="[1044,1132,1800,1826]" metricMagnitude="3" metricUnit="m" metricValue="1.8199999999999998" pageId="1" pageNumber="17" unit="m" value="1820.0">
<elevation id="00274B1D284106660062F8E3C300FB1E" box="[1044,1132,1800,1826]" metricMagnitude="3" metricUnit="m" metricValue="1.8199999999999998" pageId="1" pageNumber="17" unit="m" value="1820.0">1,820m</elevation>
</quantity>
.
<geoCoordinate id="EE3ECAE9284106660000F8E3C392FB1E" box="[1142,1278,1800,1826]" degrees="20" direction="south" minutes="28" orientation="latitude" pageId="1" pageNumber="17" precision="15" seconds="43" value="-20.478611">20°2843S</geoCoordinate>
<geoCoordinate id="EE3ECAE9284106660172F8E3C2F9FB1F" box="[1284,1429,1800,1827]" degrees="41" direction="west" minutes="49" orientation="longitude" pageId="1" pageNumber="17" precision="15" seconds="45" value="-41.829166">41°4945W</geoCoordinate>
.
<date id="FFB48AEE2841066604E1F8C7C670FB7A" box="[151,284,1836,1862]" pageId="1" pageNumber="17" value="2009-04-08">
<collectingDate id="EFF073062841066604E1F8C7C670FB7A" box="[151,284,1836,1862]" pageId="1" pageNumber="17" value="2009-04-08">08/IV/2009</collectingDate>
</date>
; Pinto-da-Rocha,
<collectorName id="26FFC9F8284106660592F8C7C552FB7A" box="[484,574,1836,1862]" pageId="1" pageNumber="17">DaSilva</collectorName>
&amp;
<collectorName id="26FFC9F8284106660614F8C7C584FB7A" box="[610,744,1836,1862]" pageId="1" pageNumber="17">Bragagnolo</collectorName>
;
<specimenCount id="9D0C67A7284106660685F8C7C424FB7A" box="[755,840,1836,1863]" count="5" pageId="1" pageNumber="17" type="male">5 males</specimenCount>
and
<specimenCount id="9D0C67A7284106660709F8C7C486FB7A" box="[895,1002,1836,1862]" count="2" pageId="1" pageNumber="17" type="female">2 females</specimenCount>
<typeStatus id="54B1128C284106660787F8C6C330FB7B" box="[1009,1116,1837,1863]" pageId="1" pageNumber="17" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DBAC045528410666001DF8C7C26FFB7B" box="[1131,1283,1836,1863]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="17" type="Museum">MZSP 77541</specimenCode>
) from
<location id="8ED5FAF5284106660138F8C6C6EAFB56" LSID="urn:lsid:plazi:treatment:03A31D382841066304E1FA20C49BFD7D:8ED5FAF5284106660138F8C6C6EAFB56" county="Vale Verde" latitude="-20.478611" longLatPrecision="21" longitude="-41.829166" municipality="Parque Nacional do Caparao" name="Parque Nacional do Caparao" pageId="1" pageNumber="17" stateProvince="Minas Gerais">Parque Nacional do Caparaó</location>
, Alto Caparaó,
<collectingRegion id="49CE62CC284106660642F8BBC5A2FB56" box="[564,718,1872,1898]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="17">Minas Gerais</collectingRegion>
.
<date id="FFB48AEE2841066606AEF8BBC411FB57" box="[728,893,1872,1899]" pageId="1" pageNumber="17" value="2009-05-01" valueMax="2009-05-05" valueMin="2009-05-01">
<collectingDate id="EFF073062841066606AEF8BBC411FB57" box="[728,893,1872,1899]" pageId="1" pageNumber="17" value="2009-05-01" valueMax="2009-05-05" valueMin="2009-05-01">0105/V/2009</collectingDate>
</date>
; Equipe Biota
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C310FFA52841066604B1F89FC459FBC6" pageId="1" pageNumber="17" type="diagnosis">
<paragraph id="8BB5AC2E2841066604B1F89FC459FBC6" blockId="1.[151,1437,1548,2043]" pageId="1" pageNumber="17">
<emphasis id="B97E703C2841066604B1F89FC628FBB2" bold="true" box="[199,324,1908,1934]" pageId="1" pageNumber="17">Diagnosis:</emphasis>
The new species is distinguished from others in the genus by the presence of a pair of small tubercles on the ocularium, and on mesotergal area III (instead of large spines, similar to
<taxonomicName id="4C0AD7AD284106660054F873C382FB8E" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1922" box="[1058,1262,1944,1971]" class="Arachnida" family="Gonyleptidae" genus="Goniosoma" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="macracanthum">
<emphasis id="B97E703C284106660054F873C35AFB8E" box="[1058,1078,1944,1970]" italics="true" pageId="1" pageNumber="17">G</emphasis>
.
<emphasis id="B97E703C284106660030F873C382FB8E" box="[1094,1262,1944,1970]" italics="true" pageId="1" pageNumber="17">macracanthum</emphasis>
</taxonomicName>
), presence of a complex dry mark resembling a mask on the dorsal scutum (similar to
<taxonomicName id="4C0AD7AD2841066607BDF857C353FBEA" authorityName="Perty" authorityYear="1833" box="[971,1087,1980,2006]" class="Arachnida" family="Gonyleptidae" genus="Goniosoma" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="varium">
<emphasis id="B97E703C2841066607BDF857C353FBEA" box="[971,1087,1980,2006]" italics="true" pageId="1" pageNumber="17">G. varium</emphasis>
</taxonomicName>
), and a ventral process with a flabellum on the penis (similar to
<taxonomicName id="4C0AD7AD284106660665F80BC5E8FBC6" authorityName="Perty" authorityYear="1833" box="[531,644,2016,2042]" class="Arachnida" family="Gonyleptidae" genus="Goniosoma" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="varium">
<emphasis id="B97E703C284106660665F80BC5E8FBC6" box="[531,644,2016,2042]" italics="true" pageId="1" pageNumber="17">G. varium</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0AD7AD2841066606CDF80BC449FBC6" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1936" box="[699,805,2016,2042]" class="Arachnida" family="Gonyleptidae" genus="Goniosoma" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="17" phylum="Arthropoda" rank="species" species="carum">
<emphasis id="B97E703C2841066606CDF80BC449FBC6" box="[699,805,2016,2042]" italics="true" pageId="1" pageNumber="17">G. carum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF75FCA62842066504E1F899C417FBCA" ID-DOI="http://doi.org/10.5281/zenodo.15012450" ID-Zenodo-Dep="15012450" httpUri="https://zenodo.org/record/15012450/files/figure.png" pageId="2" pageNumber="18" startId="2.[151,250,1906,1930]" targetBox="[179,1402,202,1856]" targetPageId="2" targetType="figure">
<paragraph id="8BB5AC2E2842066504E1F899C417FBCA" blockId="2.[151,1437,1906,2038]" pageId="2" pageNumber="18">
<emphasis id="B97E703C2842066504E1F899C679FBB6" bold="true" box="[151,277,1906,1930]" pageId="2" pageNumber="18">FIGURE 1.</emphasis>
AG male holotype (MZSP 31022) of
<taxonomicName id="4C0AD7AD2842066506D8F898C4CDFBB6" authorityName="Gomes-Silva &amp; Dasilva" authorityYear="2025" box="[686,929,1906,1930]" class="Arachnida" family="Gonyleptidae" genus="Goniosoma" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="18" phylum="Arthropoda" rank="species" species="bragagnolo" status="sp. nov.">
<emphasis id="B97E703C2842066506D8F898C4CDFBB6" box="[686,929,1906,1930]" italics="true" pageId="2" pageNumber="18">Goniosoma bragagnolo</emphasis>
</taxonomicName>
<emphasis id="B97E703C2842066507DCF899C490FBB6" bold="true" box="[938,1020,1906,1930]" pageId="2" pageNumber="18">
<taxonomicNameLabel id="A24DCD472842066507DCF899C490FBB6" box="[938,1020,1906,1930]" pageId="2" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AB habitus: A) lateral view; B) dorsal view; C) pedipalp, ventral-retrolateral view; D) femur of pedipalp, ventral view, showing prolateral seta; E) trochanter and femur IV, dorsal view; F) tibia IV, ventral-retrolateral view, showing armature of row 2 and 3 (arrows); G) metatarsus IV, ventral-retrolateral view, showing armature of row 3. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="C310FFA52843066404B1FF7CC48CFCE9" pageId="3" pageNumber="19" type="etymology">
<paragraph id="8BB5AC2E2843066404B1FF7CC48CFCE9" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FF7CC63EFC8D" bold="true" box="[199,338,151,177]" pageId="3" pageNumber="19">Etymology:</emphasis>
In honor of the opilionologist, author of many species of
<taxonomicName id="4C0AD7AD284306640792FF7CC312FC8D" authorityName="Sundevall" authorityYear="1833" box="[996,1150,151,177]" class="Arachnida" family="Gonyleptidae" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="19" phylum="Arthropoda" rank="family">Gonyleptidae</taxonomicName>
, as well as friend, Cibele Bragagnolo, who collected some of the type-specimens of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C310FFA52843066404B1FF35C210FCC5" box="[199,1404,222,249]" pageId="3" pageNumber="19" type="distribution">
<paragraph id="8BB5AC2E2843066404B1FF35C210FCC5" blockId="3.[151,1437,151,2013]" box="[199,1404,222,249]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FF35C60FFCC4" bold="true" box="[199,355,222,248]" pageId="3" pageNumber="19">Distribution:</emphasis>
slopes of the eastern end of the Mantiqueira range, in the Caparaó and Brigadeiro mountains.
</paragraph>
</subSubSection>
<subSubSection id="C310FFA52843066304B1FEE9C49BFD7D" lastPageId="4" lastPageNumber="20" pageId="3" pageNumber="19" type="description">
<paragraph id="8BB5AC2E2843066404B1FEE9C4A8FDED" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FEE9C63DFD20" bold="true" box="[199,337,258,284]" pageId="3" pageNumber="19">Description</emphasis>
(values in parentheses, for characters with variations, refer to the
<typeStatus id="54B1128C2843066400F7FEE8C38AFD21" box="[1153,1254,259,285]" pageId="3" pageNumber="19" type="holotype">holotype</typeStatus>
, figure 1AB):
<emphasis id="B97E703C2843066404E1FECDC676FD7C" bold="true" box="[151,282,294,320]" pageId="3" pageNumber="19">Ocularium</emphasis>
with a pair of small tubercles separated from each other and near to the eyes; pair of tubercles of
<emphasis id="B97E703C28430664011EFECCC7CEFD59" bold="true" pageId="3" pageNumber="19">area I</emphasis>
indistinct;
<emphasis id="B97E703C284306640552FEA0C6EEFD59" bold="true" box="[292,386,331,357]" pageId="3" pageNumber="19">area III</emphasis>
with a pair of small tubercles;
<emphasis id="B97E703C28430664069DFEA0C43AFD59" bold="true" box="[747,854,331,357]" pageId="3" pageNumber="19">carapace</emphasis>
with 1333 (33) minute granules;
<emphasis id="B97E703C284306640090FEA0C243FD59" bold="true" box="[1254,1327,331,357]" pageId="3" pageNumber="19">area I</emphasis>
with 39 (6),
<emphasis id="B97E703C2843066404BEFE84C677FDB5" bold="true" box="[200,283,367,393]" pageId="3" pageNumber="19">area II</emphasis>
with 69 (6) and
<emphasis id="B97E703C28430664059AFE84C526FDB5" bold="true" box="[492,586,367,393]" pageId="3" pageNumber="19">area III</emphasis>
with 46 (4) small granules;
<emphasis id="B97E703C2843066407E9FE84C326FDB5" bold="true" box="[927,1098,367,393]" pageId="3" pageNumber="19">dorsal scutum</emphasis>
:
<emphasis id="B97E703C28430664002DFE85C24BFDB4" bold="true" box="[1115,1319,366,392]" pageId="3" pageNumber="19">posterior margin</emphasis>
with high density of minute granules;
<emphasis id="B97E703C2843066405BFFE78C51FFD90" bold="true" box="[457,627,402,429]" pageId="3" pageNumber="19">lateral margin</emphasis>
with medium density of minute granules;
<emphasis id="B97E703C284306640035FE79C3BCFD90" bold="true" box="[1091,1232,402,428]" pageId="3" pageNumber="19">free tergites</emphasis>
,
<emphasis id="B97E703C2843066400AAFE78C2FAFD91" bold="true" box="[1244,1430,403,429]" pageId="3" pageNumber="19">anal operculum</emphasis>
,
<emphasis id="B97E703C2843066404E1FE5DC626FDED" bold="true" box="[151,330,438,465]" pageId="3" pageNumber="19">stigmatic areas</emphasis>
, and
<emphasis id="B97E703C2843066405FEFE5DC54EFDEC" bold="true" box="[392,546,438,464]" pageId="3" pageNumber="19">free sternites</emphasis>
with low density of minute granules.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1FE31C508FE25" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FE31C62AFDC8" bold="true" box="[199,326,474,500]" pageId="3" pageNumber="19">Chelicerae</emphasis>
:
<emphasis id="B97E703C284306640521FE31C6BEFDC9" bold="true" box="[343,466,474,501]" pageId="3" pageNumber="19">Segment I</emphasis>
with 3 small dorsal basal granules and 1 small apical prolateral granule;
<emphasis id="B97E703C28430664016AFE30C2F0FDC9" bold="true" box="[1308,1436,475,501]" pageId="3" pageNumber="19">segment II</emphasis>
with medium density of minute granules.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1FDC9C324FE95" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FDC9C650FE00" bold="true" box="[199,316,546,572]" pageId="3" pageNumber="19">Pedipalps</emphasis>
(figure 1CD):
<emphasis id="B97E703C284306640580FDC8C510FE01" bold="true" box="[502,636,547,573]" pageId="3" pageNumber="19">Trochanter</emphasis>
with 2 central dorsal tubercles, 1 central ventral tubercle, and 2 small apical ventral granules;
<emphasis id="B97E703C2843066405DBFDADC69AFE5C" bold="true" box="[429,502,582,608]" pageId="3" pageNumber="19">femur</emphasis>
with 6 ventral elevations 'IiIiii', and 1 subapical prolateral spine;
<emphasis id="B97E703C284306640081FDACC224FE5D" bold="true" box="[1271,1352,583,609]" pageId="3" pageNumber="19">patella</emphasis>
with 1 apical prolateral ventral tubercle;
<emphasis id="B97E703C28430664066EFD81C522FEB8" bold="true" box="[536,590,618,644]" pageId="3" pageNumber="19">tibia</emphasis>
with 1 small basal ventral tubercle, ventral armature with retrolateral 'IiIi' and prolateral 'IiIIi';
<emphasis id="B97E703C28430664050BFD64C6AAFE95" bold="true" box="[381,454,655,681]" pageId="3" pageNumber="19">tarsus</emphasis>
with ventral armature retrolateral 'IiIi' and prolateral 'IIi'.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1FD58C329FECD" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FD58C664FEF1" bold="true" box="[199,264,691,717]" pageId="3" pageNumber="19">Leg I</emphasis>
:
<emphasis id="B97E703C28430664056FFD59C63BFEF0" bold="true" box="[281,343,690,716]" pageId="3" pageNumber="19">Coxa</emphasis>
with ventral row of very large granules;
<emphasis id="B97E703C284306640746FD58C4C3FEF1" bold="true" box="[816,943,691,717]" pageId="3" pageNumber="19">trochanter</emphasis>
with low density of large ventral granules;
<emphasis id="B97E703C2843066404E1FD3DC78CFECC" bold="true" box="[151,224,726,752]" pageId="3" pageNumber="19">femur</emphasis>
and
<emphasis id="B97E703C284306640561FD3CC604FECD" bold="true" box="[279,360,727,753]" pageId="3" pageNumber="19">patella</emphasis>
with small granules;
<emphasis id="B97E703C28430664062FFD3DC5E3FECC" bold="true" box="[601,655,726,752]" pageId="3" pageNumber="19">tibia</emphasis>
and
<emphasis id="B97E703C2843066406B0FD3CC426FECD" bold="true" box="[710,842,727,753]" pageId="3" pageNumber="19">metatarsus</emphasis>
with minute granules.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1FD10C329FF05" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FD10C67CFF29" bold="true" box="[199,272,763,789]" pageId="3" pageNumber="19">Leg II</emphasis>
:
<emphasis id="B97E703C284306640568FD11C630FF28" bold="true" box="[286,348,762,788]" pageId="3" pageNumber="19">Coxa</emphasis>
with a ventral row of medium granules;
<emphasis id="B97E703C284306640768FD10C4F1FF29" bold="true" box="[798,925,763,789]" pageId="3" pageNumber="19">trochanter</emphasis>
with low density of ventral medium granules;
<emphasis id="B97E703C2843066404E1FCF5C78CFF04" bold="true" box="[151,224,798,824]" pageId="3" pageNumber="19">femur</emphasis>
and
<emphasis id="B97E703C284306640561FCF4C604FF05" bold="true" box="[279,360,799,825]" pageId="3" pageNumber="19">patella</emphasis>
with small granules;
<emphasis id="B97E703C28430664062FFCF5C5E3FF04" bold="true" box="[601,655,798,824]" pageId="3" pageNumber="19">tibia</emphasis>
and
<emphasis id="B97E703C2843066406B0FCF4C426FF05" bold="true" box="[710,842,799,825]" pageId="3" pageNumber="19">metatarsus</emphasis>
with minute granules.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1FCA8C488FFBD" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FCA8C602FF60" bold="true" box="[199,366,834,861]" pageId="3" pageNumber="19">Leg III: Coxa</emphasis>
with low density of minute granules;
<emphasis id="B97E703C284306640750FCA8C4C9FF61" bold="true" box="[806,933,835,861]" pageId="3" pageNumber="19">trochanter</emphasis>
with low density of small granules;
<emphasis id="B97E703C28430664013AFCA9C2F9FF60" bold="true" box="[1356,1429,834,860]" pageId="3" pageNumber="19">femur</emphasis>
,
<emphasis id="B97E703C2843066404E1FC8CC784FFBD" bold="true" box="[151,232,871,897]" pageId="3" pageNumber="19">patella</emphasis>
and
<emphasis id="B97E703C284306640568FC8DC638FFBC" bold="true" box="[286,340,870,896]" pageId="3" pageNumber="19">tibia</emphasis>
with medium granules;
<emphasis id="B97E703C284306640613FC8CC585FFBD" bold="true" box="[613,745,871,897]" pageId="3" pageNumber="19">metatarsus</emphasis>
with minute granules.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1FC60C502F8F9" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FC60C670FF99" bold="true" box="[199,284,907,933]" pageId="3" pageNumber="19">Leg IV</emphasis>
(figure 1B, EG):
<emphasis id="B97E703C28430664058FFC61C55BFF98" bold="true" box="[505,567,906,932]" pageId="3" pageNumber="19">Coxa</emphasis>
with low density of minute granules;
<emphasis id="B97E703C284306640786FC60C38AFF98" bold="true" box="[1008,1254,906,933]" pageId="3" pageNumber="19">prolateral apophysis</emphasis>
inserted almost dorsally, curved backwards and downwards at the apex; large
<emphasis id="B97E703C28430664070CFC45C3B5FFF4" bold="true" box="[890,1241,942,969]" pageId="3" pageNumber="19">apical retrolateral apophysis</emphasis>
pointed and the same size as the trochanter;
<emphasis id="B97E703C2843066405ADFC38C536FFD1" bold="true" box="[475,602,979,1005]" pageId="3" pageNumber="19">trochanter</emphasis>
with a small, pointed and upward-curved prolateral sub-basal apophysis, retrolateral apical apophysis pointed, curved backwards and with a granule associated;
<emphasis id="B97E703C2843066400E6FC1DC3B5F82C" bold="true" box="[1168,1241,1014,1040]" pageId="3" pageNumber="19">femur</emphasis>
short, with very strong inward curvature,
<emphasis id="B97E703C2843066405C1FBF1C47DF808" bold="true" box="[439,785,1050,1077]" pageId="3" pageNumber="19">apical retrolateral apophysis</emphasis>
very large and curved backwards,
<emphasis id="B97E703C2843066400D3FBF0C2F0F808" bold="true" box="[1189,1436,1050,1077]" pageId="3" pageNumber="19">prolateral apophysis</emphasis>
very small, row 2 with flat granules, with a high density of disorganized granules in the apical third;
<emphasis id="B97E703C284306640167FBD4C20EF865" bold="true" box="[1297,1378,1087,1113]" pageId="3" pageNumber="19">patella</emphasis>
with large prolateral granules;
<emphasis id="B97E703C2843066405C4FB89C684F840" bold="true" box="[434,488,1122,1148]" pageId="3" pageNumber="19">tibia</emphasis>
with row 2 with a group of large granules close to each other, row 3 with small spines curved backwards along the entire length, increasing from base to apex;
<emphasis id="B97E703C2843066407BCFB6CC322F89D" bold="true" box="[970,1102,1159,1185]" pageId="3" pageNumber="19">metatarsus</emphasis>
with row 3 with small spines in the first half;
<emphasis id="B97E703C28430664053AFB40C683F8F9" bold="true" box="[332,495,1195,1221]" pageId="3" pageNumber="19">tarsal process</emphasis>
very large.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1FB24C4F9F8D5" blockId="3.[151,1437,151,2013]" box="[199,917,1230,1257]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FB24C6D4F8D4" bold="true" box="[199,440,1230,1257]" pageId="3" pageNumber="19">Tarsal segmentation</emphasis>
: 810 (9); 1622 (20); 810 (9); 810 (9).
</paragraph>
<paragraph id="8BB5AC2E2843066404B1FB19C6D9FA11" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1FB19C66BF930" bold="true" box="[199,263,1266,1292]" pageId="3" pageNumber="19">Penis</emphasis>
(figure 3AB):
<emphasis id="B97E703C2843066405BDFB18C507F931" bold="true" box="[459,619,1267,1293]" pageId="3" pageNumber="19">Ventral plate</emphasis>
rectangular-shaped, longer than wide, with practically straight lateral margins, only a slight convexity in the apical region due to its elevation;
<emphasis id="B97E703C284306640785FAFDC3F0F90C" bold="true" box="[1011,1180,1302,1328]" pageId="3" pageNumber="19">apical margin</emphasis>
slightly concave with elevations above the margin at the ends. All macrosetae pointed;
<emphasis id="B97E703C2843066407D6FAD0C36AF968" bold="true" box="[928,1030,1338,1365]" pageId="3" pageNumber="19">group A</emphasis>
with 3 macrosetae aligned almost longitudinally and slightly curved downwards;
<emphasis id="B97E703C2843066406C5FAB4C43AF945" bold="true" box="[691,854,1375,1401]" pageId="3" pageNumber="19">macrosetae B</emphasis>
small, positioned more apically and ventrally than the A1;
<emphasis id="B97E703C284306640482FA68C634F9A0" bold="true" box="[244,344,1410,1437]" pageId="3" pageNumber="19">group C</emphasis>
with 3 macrosetae, C1 being slightly separated from the others and slightly curved backwards;
<emphasis id="B97E703C2843066404E1FA4CC654F9FD" bold="true" box="[151,312,1447,1473]" pageId="3" pageNumber="19">macrosetae D</emphasis>
with half size of the macrosetae of group C and positioned below C3;
<emphasis id="B97E703C284306640032FA4CC388F9FD" bold="true" box="[1092,1252,1447,1473]" pageId="3" pageNumber="19">macrosetae E</emphasis>
with half size of the macrosetae of group C and positioned more ventrally and between C1 and C2;
<emphasis id="B97E703C284306640040FA20C318F9D9" bold="true" box="[1078,1140,1483,1509]" pageId="3" pageNumber="19">glans</emphasis>
with ventral process long, with a flabellum on apex, with many projections on apical margin, a little wider than the stem;
<emphasis id="B97E703C2843066400AEFA04C270FA35" bold="true" box="[1240,1308,1519,1545]" pageId="3" pageNumber="19">stylus</emphasis>
cylindrical with a dorsal apical beak.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1F9DCC30AFA49" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1F9DCC557FA6D" bold="true" box="[199,571,1591,1617]" pageId="3" pageNumber="19">Measurements. Dorsal scutum.</emphasis>
Length: 8.910.6 (10.6); width: 8.411.1 (11.1);
<emphasis id="B97E703C28430664001DF9DDC3B2FA6C" bold="true" box="[1131,1246,1590,1616]" pageId="3" pageNumber="19">pedipalps</emphasis>
: 1314.1 (13.9);
<emphasis id="B97E703C2843066404E1F9B0C78CFA49" bold="true" box="[151,224,1627,1653]" pageId="3" pageNumber="19">legs: I</emphasis>
: 24.128.8 (28.7);
<emphasis id="B97E703C2843066405CEF9B0C6BBFA49" bold="true" box="[440,471,1627,1653]" pageId="3" pageNumber="19">II:</emphasis>
44.656.6 (56.5);
<emphasis id="B97E703C2843066406D1F9B0C5A4FA49" bold="true" box="[679,712,1627,1653]" pageId="3" pageNumber="19">III</emphasis>
: 34.943.2 (43.2);
<emphasis id="B97E703C2843066407D6F9B0C4D3FA49" bold="true" box="[928,959,1627,1653]" pageId="3" pageNumber="19">IV</emphasis>
: 4759 (58.8).
</paragraph>
<paragraph id="8BB5AC2E2843066404B1F995C6AFFB71" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1F995C624FAA4" bold="true" box="[199,328,1662,1688]" pageId="3" pageNumber="19">Coloration</emphasis>
(photos of live animals; figure 4): predominantly dark brown;
<emphasis id="B97E703C28430664006FF995C3FDFAA4" bold="true" box="[1049,1169,1662,1688]" pageId="3" pageNumber="19">chelicerae</emphasis>
dark
<emphasis id="B97E703C2843066400BDF994C3B8FAA5" bold="true" box="[1227,1236,1663,1689]" pageId="3" pageNumber="19">-</emphasis>
yellowish;
<emphasis id="B97E703C284306640124F994C782FA81" bold="true" pageId="3" pageNumber="19">dorsal scutum</emphasis>
dark brown, with yellowish
<emphasis id="B97E703C28430664065AF948C5EBFA81" bold="true" box="[556,647,1699,1725]" pageId="3" pageNumber="19">grooves</emphasis>
and
<emphasis id="B97E703C2843066406CBF948C444FA81" bold="true" box="[701,808,1699,1725]" pageId="3" pageNumber="19">carapace</emphasis>
;
<emphasis id="B97E703C284306640740F948C2F0FA80" bold="true" box="[822,1436,1698,1725]" pageId="3" pageNumber="19">apexes of trochanters of legs, tubercles of ocularium</emphasis>
and
<emphasis id="B97E703C2843066404BEF92CC50AFADD" bold="true" box="[200,614,1734,1761]" pageId="3" pageNumber="19">granules of areas of dorsal scutum</emphasis>
orange to yellow;
<emphasis id="B97E703C28430664074AF92DC355FADD" bold="true" box="[828,1081,1734,1761]" pageId="3" pageNumber="19">articular membranes</emphasis>
between coxae and trochanters pink;
<emphasis id="B97E703C2843066404A0F900C62BFB39" bold="true" box="[214,327,1771,1797]" pageId="3" pageNumber="19">dry mark</emphasis>
prominent, on the
<emphasis id="B97E703C284306640660F900C5EDFB39" bold="true" box="[534,641,1771,1797]" pageId="3" pageNumber="19">carapace</emphasis>
like complete mask, on the
<emphasis id="B97E703C2843066407B1F901C3CFFB39" bold="true" box="[967,1187,1770,1797]" pageId="3" pageNumber="19">abdominal scutum</emphasis>
following the grooves between
<emphasis id="B97E703C28430664048AF8E4C6BAFB15" bold="true" box="[252,470,1807,1833]" pageId="3" pageNumber="19">areas I, II and III,</emphasis>
between the tubercles of
<emphasis id="B97E703C284306640681F8E4C438FB15" bold="true" box="[759,852,1807,1833]" pageId="3" pageNumber="19">area III</emphasis>
as two circles and present on
<emphasis id="B97E703C2843066400DEF8E4C251FB15" bold="true" box="[1192,1341,1806,1833]" pageId="3" pageNumber="19">apex of coxa</emphasis>
, base of
<emphasis id="B97E703C2843066404E1F8D8C67AFB71" bold="true" box="[151,278,1843,1869]" pageId="3" pageNumber="19">trochanter</emphasis>
and
<emphasis id="B97E703C28430664053BF8D9C6D0FB71" bold="true" box="[333,444,1842,1869]" pageId="3" pageNumber="19">femur IV</emphasis>
.
</paragraph>
<paragraph id="8BB5AC2E2843066404B1F8BCC4D0FBE1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="19">
<emphasis id="B97E703C2843066404B1F8BCC672FB4D" bold="true" box="[199,286,1879,1905]" pageId="3" pageNumber="19">Female</emphasis>
(figure 4): legs less armed and with weaker granulation compared to males; prolateral apophysis of coxa IV small, spine-like; trochanter IV with only a small apical retrolateral apophysis; posterior margin and free tergites with larger granules compared to males, and with a pair of small tubercles at the angles; tarsal processes smaller than in the male (approximately half the size of the tarsal claw).
</paragraph>
<paragraph id="8BB5AC2E2844066304E1FF7CC49BFD7D" blockId="4.[151,1436,151,321]" pageId="4" pageNumber="20">
<taxonomicName id="4C0AD7AD2844066304E1FF7CC6BBFC8D" authority="Roewer 1930" authorityName="Roewer" authorityYear="1930" box="[151,471,151,177]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B97E703C2844066304E1FF7CC656FC8D" box="[151,314,151,177]" italics="true" pageId="4" pageNumber="20">Serracutisoma</emphasis>
<bibRefCitation id="EF9BD1DF284406630537FF7CC6BBFC8D" author="Roewer, C. F." box="[321,471,151,177]" pageId="4" pageNumber="20" pagination="341 - 452" refId="ref7982" refString="Roewer, C. F. (1930) Weitere Weberknechte IV. Erganzung der: ' Weberknechte der Erde, 1923 '. Abhandlungen. Naturwissenschaftlicher Verein zu Bremen, 27, 341-452." type="journal article" year="1930">Roewer 1930</bibRefCitation>
</taxonomicName>
now includes:
<taxonomicName id="4C0AD7AD2844066306F6FF7CC460FC8C" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1922" box="[640,780,151,177]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="proximum">
<emphasis id="B97E703C2844066306F6FF7CC460FC8C" box="[640,780,151,177]" italics="true" pageId="4" pageNumber="20">S. proximum</emphasis>
</taxonomicName>
group:
<taxonomicName id="4C0AD7AD284406630714FF7CC3C5FC8D" authority="(Simon 1879)" baseAuthorityName="Simon" baseAuthorityYear="1879" box="[866,1193,151,177]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="thalassinum">
<emphasis id="B97E703C284406630714FF7CC368FC8D" box="[866,1028,151,177]" italics="true" pageId="4" pageNumber="20">S. thalassinum</emphasis>
(
<bibRefCitation id="EF9BD1DF284406630065FF7CC3CCFC8D" author="Simon, E." box="[1043,1184,151,177]" pageId="4" pageNumber="20" pagination="183 - 241" refId="ref8165" refString="Simon, E. (1879) Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Annales de la Societe Entomologique de Belgique, 22, 183-241." type="journal article" year="1879">Simon 1879</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C0AD7AD2844066300C2FF7CC647FCE9" authority="(Mello-Leitao 1922)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1922" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="proximum">
<emphasis id="B97E703C2844066300C2FF7CC22DFC8C" box="[1204,1345,151,177]" italics="true" pageId="4" pageNumber="20">S. proximum</emphasis>
(
<bibRefCitation id="EF9BD1DF284406630138FF7CC648FCE9" author="Mello-Leitao, C. de" pageId="4" pageNumber="20" pagination="329 - 348" refId="ref7500" refString="Mello-Leitao, C. de (1922) On some new Brazilian Gonyleptidae. Annals and Magazine of Natural History, 9, 329-348. https://doi.org/10.1080/00222932208632683" type="journal article" year="1922">Mello-Leitão 1922</bibRefCitation>
)
</taxonomicName>
, type-species;
<taxonomicName id="4C0AD7AD2844066305A2FF50C44EFCE9" authority="(Mello-Leitao 1933)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1933" box="[468,802,187,213]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="molle">
<emphasis id="B97E703C2844066305A2FF50C543FCE9" box="[468,559,187,213]" italics="true" pageId="4" pageNumber="20">S. molle</emphasis>
(Mello-Leitão 1933)
</taxonomicName>
;
<taxonomicName id="4C0AD7AD284406630746FF50C38AFCE9" authority="(Soares &amp; Soares 1947)" baseAuthorityName="Soares &amp; Soares" baseAuthorityYear="1947" box="[816,1254,187,213]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="banhadoae">
<emphasis id="B97E703C284406630746FF50C452FCE9" box="[816,830,187,213]" italics="true" pageId="4" pageNumber="20">S</emphasis>
.
<emphasis id="B97E703C284406630738FF50C4A6FCE9" box="[846,970,187,213]" italics="true" pageId="4" pageNumber="20">banhadoae</emphasis>
(
<bibRefCitation id="EF9BD1DF2844066307ADFF50C3B2FCE9" author="Soares, B. A. M. &amp; Soares, H. E. M." box="[987,1246,187,213]" pageId="4" pageNumber="20" pagination="63 - 84" refId="ref8200" refString="Soares, B. A. M. &amp; Soares, H. E. M. (1947) Alotipos e formas novas de opilioes paranaenses (Opiliones-Gonyleptidae, Phalangiidae). Papeis Avulsos do Departamento de Zoologia do Estado de Sao Paulo, 8, 63-84. https://doi.org/10.11606/0031-1049.1947.8p63-84" type="journal article" year="1947">Soares &amp; Soares 1947</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C0AD7AD284406630082FF50C6ADFCC5" authority="DaSilva &amp; Gnaspini 2010" authorityName="DaSilva &amp; Gnaspini" authorityYear="2010" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="fritzmuelleri">
<emphasis id="B97E703C284406630082FF50C26EFCE9" box="[1268,1282,187,213]" italics="true" pageId="4" pageNumber="20">S</emphasis>
.
<emphasis id="B97E703C284406630164FF50C2F0FCE9" box="[1298,1436,187,213]" italics="true" pageId="4" pageNumber="20">fritzmuelleri</emphasis>
<bibRefCitation id="EF9BD1DF2844066304E1FF34C6ADFCC5" author="DaSilva, M. B. &amp; Gnaspini, P." box="[151,449,223,249]" pageId="4" pageNumber="20" pagination="530 - 624" refId="ref6500" refString="DaSilva, M. B. &amp; Gnaspini, P. (2010) A systematic revision of Goniosomatinae (Arachnida: Opiliones: Gonyleptidae), with a cladistic analysis and biogeographical notes. Invertebrate Systematics, 23, 530-624. https://doi.org/10.1071/IS09022" type="journal article" year="2010">DaSilva &amp; Gnaspini 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C0AD7AD2844066305B8FF34C4D0FCC5" authority="DaSilva &amp; Gnaspini 2010" authorityName="DaSilva &amp; Gnaspini" authorityYear="2010" box="[462,956,223,249]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="pseudovarium">
<emphasis id="B97E703C2844066305B8FF34C5E6FCC5" box="[462,650,223,249]" italics="true" pageId="4" pageNumber="20">S. pseudovarium</emphasis>
<bibRefCitation id="EF9BD1DF2844066306E4FF34C4D0FCC5" author="DaSilva, M. B. &amp; Gnaspini, P." box="[658,956,223,249]" pageId="4" pageNumber="20" pagination="530 - 624" refId="ref6500" refString="DaSilva, M. B. &amp; Gnaspini, P. (2010) A systematic revision of Goniosomatinae (Arachnida: Opiliones: Gonyleptidae), with a cladistic analysis and biogeographical notes. Invertebrate Systematics, 23, 530-624. https://doi.org/10.1071/IS09022" type="journal article" year="2010">DaSilva &amp; Gnaspini 2010</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C0AD7AD28440663078FFF34C24AFCC5" authority="DaSilva 2014" authorityName="DaSilva" authorityYear="2014" box="[1017,1318,223,249]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="gnaspinii">
<emphasis id="B97E703C28440663078FFF34C313FCC4" box="[1017,1151,223,249]" italics="true" pageId="4" pageNumber="20">S. gnaspinii</emphasis>
<bibRefCitation id="EF9BD1DF2844066300F1FF34C24AFCC5" author="DaSilva, M. B." box="[1159,1318,223,249]" pageId="4" pageNumber="20" pagination="370 - 376" refId="ref6555" refString="DaSilva, M. B. (2014) A new species of Serracutisoma (Opiliones: Gonyleptidae: Goniosomatinae) from the coastal Atlantic Rain Forest of Parana and Sao Paulo states, Brazil. Zoologia, 31 (4), 370-376. https://doi.org/10.1590/S1984-46702014000400008" type="journal article" year="2014">DaSilva 2014</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C0AD7AD284406630147FF34C2F0FCC4" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1927" box="[1329,1436,223,249]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="inerme">
<emphasis id="B97E703C284406630147FF34C2F0FCC4" box="[1329,1436,223,249]" italics="true" pageId="4" pageNumber="20">S. inerme</emphasis>
</taxonomicName>
group:
<taxonomicName id="4C0AD7AD284406630491FEE8C52DFD21" authority="(Mello-Leitao 1927)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1927" box="[231,577,259,285]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="inerme">
<emphasis id="B97E703C284406630491FEE8C63DFD20" box="[231,337,259,285]" italics="true" pageId="4" pageNumber="20">S. inerme</emphasis>
(
<bibRefCitation id="EF9BD1DF284406630516FEE8C554FD21" author="Mello-Leitao, C. de" box="[352,568,259,285]" pageId="4" pageNumber="20" pagination="395 - 418" refId="ref7537" refString="Mello-Leitao, C. de (1927) Arachnideos de Santa Catarina (Brasil). Revista do Museu Paulista, 15, 395-418." type="journal article" year="1927">Mello-Leitão 1927</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C0AD7AD28440663063BFEE8C4AEFD21" authority="(Mello-Leitao 1933)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1933" box="[589,962,259,285]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="spelaeum">
<emphasis id="B97E703C28440663063BFEE8C5BFFD21" box="[589,723,259,285]" italics="true" pageId="4" pageNumber="20">S. spelaeum</emphasis>
(Mello-Leitão 1933)
</taxonomicName>
;
<taxonomicName id="4C0AD7AD2844066307B8FEE8C325FD20" box="[974,1097,259,285]" class="Arachnida" family="Cosmetidae" genus="Santa" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="catarina">
<emphasis id="B97E703C2844066307B8FEE8C325FD20" box="[974,1097,259,285]" italics="true" pageId="4" pageNumber="20">S. catarina</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9BD1DF28440663002EFEE8C659FD7D" author="Machado, G. &amp; Pinto-da-Rocha, R. &amp; Giaretta, A. A." pageId="4" pageNumber="20" pagination="17 - 22" refId="ref7370" refString="Machado, G., Pinto-da-Rocha, R. &amp; Giaretta, A. A. (2002) Notes on the taxonomy and biology of the neotropical harvestman Goniosoma catarina sp. n. (Opiliones: Gonyleptidae). Revista Iberica de Aracnologia, 4, 17-22." type="journal article" year="2002">Machado, Pinto-da-Rocha &amp; Giaretta 2002</bibRefCitation>
);
<taxonomicName id="4C0AD7AD28440663053EFECCC469FD7D" authority="DaSilva &amp; Gnaspini 2010" authorityName="DaSilva &amp; Gnaspini" authorityYear="2010" box="[328,773,295,321]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="guaricana">
<emphasis id="B97E703C28440663053EFECCC6BBFD7C" box="[328,471,295,321]" italics="true" pageId="4" pageNumber="20">S. guaricana</emphasis>
<bibRefCitation id="EF9BD1DF2844066305A9FECCC469FD7D" author="DaSilva, M. B. &amp; Gnaspini, P." box="[479,773,295,321]" pageId="4" pageNumber="20" pagination="530 - 624" refId="ref6500" refString="DaSilva, M. B. &amp; Gnaspini, P. (2010) A systematic revision of Goniosomatinae (Arachnida: Opiliones: Gonyleptidae), with a cladistic analysis and biogeographical notes. Invertebrate Systematics, 23, 530-624. https://doi.org/10.1071/IS09022" type="journal article" year="2010">DaSilva &amp; Gnaspini 2010</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C0AD7AD284406630749FECCC4FCFD7D" authorityName="Gomes-Silva &amp; Dasilva" authorityYear="2025" box="[831,912,295,321]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="hara" status="sp. nov.">
<emphasis id="B97E703C284406630749FECCC4FCFD7D" box="[831,912,295,321]" italics="true" pageId="4" pageNumber="20">S. hara</emphasis>
</taxonomicName>
<emphasis id="B97E703C2844066307E1FECCC49CFD7D" bold="true" box="[919,1008,295,321]" pageId="4" pageNumber="20">
<taxonomicNameLabel id="A24DCD472844066307E1FECCC49CFD7D" box="[919,1008,295,321]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,358 @@
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<mods:title id="B61F5E1E543B28E5C4DFAC9B7C908BFB">Two new species of Goniosomatinae (Opiliones: Gonyleptidae) narrowly endemic to the Atlantic Forest of Brazil.</mods:title>
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<mods:namePart id="1DE13ABFCFEFC49709B4A3EB1FB3B2FC">Gomes-Silva, José Geovane</mods:namePart>
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<paragraph id="8BB5AC2E2844066304E1FE78C5B9FD90" blockId="4.[151,725,402,429]" box="[151,725,402,429]" pageId="4" pageNumber="20">
<heading id="D0FD1B422844066304E1FE78C5B9FD90" bold="true" box="[151,725,402,429]" fontSize="11" level="1" pageId="4" pageNumber="20" reason="1">
<emphasis id="B97E703C2844066304E1FE78C5B9FD90" bold="true" box="[151,725,402,429]" pageId="4" pageNumber="20">
<taxonomicName id="4C0AD7AD2844066304E1FE78C6E4FD91" authorityName="Gomes-Silva &amp; Dasilva" authorityYear="2025" box="[151,392,402,429]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="hara" status="sp. nov.">
<emphasis id="B97E703C2844066304E1FE78C6E4FD91" bold="true" box="[151,392,402,429]" italics="true" pageId="4" pageNumber="20">Serracutisoma hara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24DCD472844066305F9FE78C681FD91" box="[399,493,403,429]" pageId="4" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1331B0AB28440663058AFE79C529FD90" box="[508,581,402,428]" captionStart="FIGURE 2" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[250,1339,198,1906]" captionTargetId="figure-30@5.[237,1350,181,1919]" captionTargetPageId="5" captionText="FIGURE 2. AE male holotype (MZSP 18974) of Serracutisoma hara sp. nov. AB habitus:A) dorsal view; B) lateral view; C) pedipalp, ventral-prolateral view; D) femur III, retrolateral view, showing armature of row 3; E) trochanter and femur IV, dorsal view, showing armature of row 2 of femur. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.15012454" httpUri="https://zenodo.org/record/15012454/files/figure.png" pageId="4" pageNumber="20">Fig. 2</figureCitation>
,
<figureCitation id="1331B0AB284406630625FE79C50FFD90" box="[595,611,402,428]" captionStart="FIGURE 3" captionStartId="6.[151,250,1601,1625]" captionTargetBox="[165,1422,187,1562]" captionTargetId="figure-17@6.[151,1436,181,1577]" captionTargetPageId="6" captionText="FIGURE 3. AB penis of male holotype (MZSP 31022) of Goniosoma bragagnolo sp. nov.: A) dorsal view; B) lateral view. CD penis of male holotype (MZSP 18974) of Serracutisoma hara sp. nov.: C) dorsal view; D) lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.15012456" httpUri="https://zenodo.org/record/15012456/files/figure.png" pageId="4" pageNumber="20">3</figureCitation>
,
<figureCitation id="1331B0AB284406630606FE78C513FD91" box="[624,639,403,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1712,1736]" captionTargetBox="[156,1433,786,1685]" captionTargetId="figure-49@7.[152,1436,782,1688]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Goniosoma bragagnolo sp. nov. and Serracutisoma hara sp. nov. along with their respective genera. Black lines: limits of areas of endemism. SEsp = Serra do Espinhaço; SC = Santa Catarina; solid lines: Congruence Core; dashed lines: Maximum Region of Endemism." figureDoi="http://doi.org/10.5281/zenodo.15012463" httpUri="https://zenodo.org/record/15012463/files/figure.png" pageId="4" pageNumber="20">5</figureCitation>
and
<figureCitation id="1331B0AB2844066306CAFE79C5A7FD90" box="[700,715,402,428]" captionStart="FIGURE 6" captionStartId="8.[151,250,1113,1137]" captionTargetBox="[155,1433,184,1087]" captionTargetId="figure-17@8.[151,1436,181,1089]" captionTargetPageId="8" captionText="FIGURE 6. Distribution of Goniosoma bragagnolo sp. nov. and Serracutisoma hara sp. nov. along with their respective genera, across the vegetation types of the Atlantic Rain Forest." figureDoi="http://doi.org/10.5281/zenodo.15012465" httpUri="https://zenodo.org/record/15012465/files/figure.png" pageId="4" pageNumber="20">6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C310FFA52844066304E1FE30C623FE25" pageId="4" pageNumber="20" type="materials_examined">
<paragraph id="8BB5AC2E2844066304E1FE30C623FE25" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304E1FE30C653FDC8" bold="true" box="[151,319,474,501]" pageId="4" pageNumber="20">Type material</emphasis>
:
<materialsCitation id="3B62A673284406630539FE30C627FE25" collectingDate="1998-05-09" collectionCode="R" county="Pinto-da-Rocha" location="Gruta do Botuvera" municipality="Botuvera" pageId="4" pageNumber="20" specimenCode="MZSP 18974" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D0C67A7284406630539FE30C6F7FDC9" box="[335,411,475,501]" count="1" pageId="4" pageNumber="20" type="male">1 male</specimenCount>
<typeStatus id="54B1128C2844066305D2FE30C56AFDC9" box="[420,518,475,501]" pageId="4" pageNumber="20" type="holotype">holotype</typeStatus>
(
<specimenCode id="DBAC0455284406630663FE30C5C3FDC9" box="[533,687,475,501]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="20" type="Museum">MZSP 18974</specimenCode>
) from
<location id="8ED5FAF5284406630688FE30C4B8FDC9" LSID="urn:lsid:plazi:treatment:03A31D382844066104E1FE78C6C9FBDA:8ED5FAF5284406630688FE30C4B8FDC9" box="[766,980,475,501]" county="Pinto-da-Rocha" municipality="Botuvera" name="Gruta do Botuvera" pageId="4" pageNumber="20">Gruta do Botuverá</location>
,
<collectingMunicipality id="6BD136542844066307A9FE30C325FDC9" box="[991,1097,475,501]" pageId="4" pageNumber="20">Botuverá</collectingMunicipality>
,
<taxonomicName id="4C0AD7AD284406630023FE30C393FDC9" box="[1109,1279,475,501]" class="Arachnida" family="Cosmetidae" genus="Santa" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="catarina">Santa Catarina</taxonomicName>
.
<date id="FFB48AEE28440663017CFE30C21AFDC9" box="[1290,1398,475,501]" pageId="4" pageNumber="20" value="1998-05-09">
<collectingDate id="EFF0730628440663017CFE30C21AFDC9" box="[1290,1398,475,501]" pageId="4" pageNumber="20" value="1998-05-09">9/V/1998</collectingDate>
</date>
,
<collectionCode id="ED1B34EB2844066301F5FE30C2F5FDC9" box="[1411,1433,475,501]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="20">R</collectionCode>
.
<collectingCounty id="62D4D4A22844066304E1FE14C627FE25" box="[151,331,511,537]" pageId="4" pageNumber="20">Pinto-da-Rocha</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C310FFA52844066304B1FDC9C212FE5D" pageId="4" pageNumber="20" type="diagnosis">
<paragraph id="8BB5AC2E2844066304B1FDC9C212FE5D" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FDC9C628FE00" bold="true" box="[199,324,546,572]" pageId="4" pageNumber="20">Diagnosis:</emphasis>
Similar to other species of
<taxonomicName id="4C0AD7AD28440663060BFDC8C584FE00" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1927" box="[637,744,547,573]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="species" species="inerme">
<emphasis id="B97E703C28440663060BFDC8C584FE00" box="[637,744,547,573]" italics="true" pageId="4" pageNumber="20">S. inerme</emphasis>
</taxonomicName>
group. It differs by the combination of armature of femur III with spines close to each other and armature of row 2 of femur IV with spines only in the medial-apical portion.
</paragraph>
</subSubSection>
<subSubSection id="C310FFA52844066304B1FD80C2FBFEB9" box="[199,1431,619,645]" pageId="4" pageNumber="20" type="etymology">
<paragraph id="8BB5AC2E2844066304B1FD80C2FBFEB9" blockId="4.[151,1437,474,1941]" box="[199,1431,619,645]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FD80C63EFEB9" bold="true" box="[199,338,619,645]" pageId="4" pageNumber="20">Etymology:</emphasis>
In honor of the opilionologist, author of many species of
<taxonomicName id="4C0AD7AD2844066307AFFD80C31FFEB9" authorityName="Sundevall" authorityYear="1833" box="[985,1139,619,645]" class="Arachnida" family="Gonyleptidae" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="20" phylum="Arthropoda" rank="family">Gonyleptidae</taxonomicName>
, and friend, Marcos Hara.
</paragraph>
</subSubSection>
<subSubSection id="C310FFA52844066304B1FD65C598FE95" box="[199,756,654,681]" pageId="4" pageNumber="20" type="distribution">
<paragraph id="8BB5AC2E2844066304B1FD65C598FE95" blockId="4.[151,1437,474,1941]" box="[199,756,654,681]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FD65C60FFE94" bold="true" box="[199,355,654,680]" pageId="4" pageNumber="20">Distribution:</emphasis>
Only known from the
<typeStatus id="54B1128C284406630615FD64C5FFFE95" box="[611,659,655,681]" pageId="4" pageNumber="20">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C310FFA52844066104B1FD59C6C9FBDA" lastPageId="6" lastPageNumber="22" pageId="4" pageNumber="20" type="description">
<paragraph id="8BB5AC2E2844066304B1FD59C613FF61" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FD59C63DFEF0" bold="true" box="[199,337,690,716]" pageId="4" pageNumber="20">Description</emphasis>
(figure 2AB):
<emphasis id="B97E703C284406630664FD59C5F9FEF0" bold="true" box="[530,661,690,716]" pageId="4" pageNumber="20">Ocularium</emphasis>
with 1 pair of medium tubercles, separated and close to the eyes;
<emphasis id="B97E703C2844066304E1FD3CC66EFECD" bold="true" box="[151,258,727,753]" pageId="4" pageNumber="20">carapace</emphasis>
with 6 minute granules;
<emphasis id="B97E703C284406630661FD3CC58BFECD" bold="true" box="[535,743,727,753]" pageId="4" pageNumber="20">areas I, II and III</emphasis>
with, respectively, 10, 10 and 6 medium granules;
<emphasis id="B97E703C284406630168FD3CC208FECD" bold="true" box="[1310,1380,727,753]" pageId="4" pageNumber="20">area I</emphasis>
with 1 pair of small tubercles;
<emphasis id="B97E703C2844066305C1FD10C57FFF29" bold="true" box="[439,531,763,789]" pageId="4" pageNumber="20">area III</emphasis>
with 1 pair of medium spines;
<emphasis id="B97E703C284406630705FD10C3B5FF28" bold="true" box="[883,1241,762,789]" pageId="4" pageNumber="20">dorsal scutum: lateral margin</emphasis>
with low density of minute granules;
<emphasis id="B97E703C28440663050FFCF5C528FF04" bold="true" box="[377,580,798,824]" pageId="4" pageNumber="20">posterior margin</emphasis>
and
<emphasis id="B97E703C28440663060DFCF5C466FF04" bold="true" box="[635,778,798,824]" pageId="4" pageNumber="20">free tergites</emphasis>
practically smooth;
<emphasis id="B97E703C284406630786FCF4C271FF04" bold="true" box="[1008,1309,798,825]" pageId="4" pageNumber="20">angles of the free tergites</emphasis>
with a pair of medium granules.
</paragraph>
<paragraph id="8BB5AC2E2844066304B1FC8DC4A3FF99" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FC8DC56AFFBD" bold="true" box="[199,518,870,897]" pageId="4" pageNumber="20">Chelicerae: First segment</emphasis>
with 4 dorso-basal tubercles, 1 prolateral dorso-apical tubercle and 1 apical retrolateral tubercle;
<emphasis id="B97E703C2844066305F4FC60C550FF99" bold="true" box="[386,572,907,933]" pageId="4" pageNumber="20">second segment</emphasis>
with a medium density of granules.
</paragraph>
<paragraph id="8BB5AC2E2844066304B1FC45C4E8F865" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FC45C650FFF4" bold="true" box="[199,316,942,968]" pageId="4" pageNumber="20">Pedipalps</emphasis>
(figure 2C):
<emphasis id="B97E703C2844066305A3FC44C537FFF5" bold="true" box="[469,603,943,969]" pageId="4" pageNumber="20">Trochanter</emphasis>
with 2 medium granules and 1 apical ventral setiferous tubercle;
<emphasis id="B97E703C284406630125FC45C2F0FFF4" bold="true" box="[1363,1436,942,968]" pageId="4" pageNumber="20">femur</emphasis>
with 8 ventral tubercles IiiiIiii, and 1 prolateral spine;
<emphasis id="B97E703C284406630771FC38C434FFD1" bold="true" box="[775,856,979,1005]" pageId="4" pageNumber="20">patella</emphasis>
with 1 large prolateral granule;
<emphasis id="B97E703C2844066300B6FC39C39AFFD0" bold="true" box="[1216,1270,978,1004]" pageId="4" pageNumber="20">tibia</emphasis>
with elevation pattern on the retrolateral ventral setae IiIi, the penultimate larger, more thicker than the others, and fused with the last at the base and prolateral setae IiIi, with 4 small ventral granules and 1 small retrolateral tubercle pointed;
<emphasis id="B97E703C2844066304E1FBD4C78CF865" bold="true" box="[151,224,1087,1113]" pageId="4" pageNumber="20">tarsus</emphasis>
with prolateral ventral setae IIi and retrolateral setae IiIi.
</paragraph>
<paragraph id="8BB5AC2E2844066304B1FB88C4ADF89D" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FB88C66AF841" bold="true" box="[199,262,1123,1149]" pageId="4" pageNumber="20">Leg I</emphasis>
:
<emphasis id="B97E703C284406630562FB89C63EF840" bold="true" box="[276,338,1122,1148]" pageId="4" pageNumber="20">Coxa</emphasis>
with 1 row of medium tubercles;
<emphasis id="B97E703C2844066306BBFB88C420F841" bold="true" box="[717,844,1123,1149]" pageId="4" pageNumber="20">trochanter</emphasis>
with 1 large granule and 2 medium granules;
<emphasis id="B97E703C28440663013AFB89C2F9F840" bold="true" box="[1356,1429,1122,1148]" pageId="4" pageNumber="20">femur</emphasis>
,
<emphasis id="B97E703C2844066304E1FB6CC783F89D" bold="true" box="[151,239,1159,1185]" pageId="4" pageNumber="20">patella,</emphasis>
and
<emphasis id="B97E703C284406630553FB6DC637F89C" bold="true" box="[293,347,1158,1184]" pageId="4" pageNumber="20">tibia</emphasis>
with minute granules;
<emphasis id="B97E703C284406630628FB6CC58EF89D" bold="true" box="[606,738,1159,1185]" pageId="4" pageNumber="20">metatarsus</emphasis>
practically smooth.
</paragraph>
<paragraph id="8BB5AC2E2844066304B1FB40C602F931" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FB40C60AF8F8" bold="true" box="[199,358,1194,1221]" pageId="4" pageNumber="20">Leg II: Coxa</emphasis>
with 1 row of small tubercles;
<emphasis id="B97E703C2844066306ACFB40C435F8F9" bold="true" box="[730,857,1195,1221]" pageId="4" pageNumber="20">trochanter</emphasis>
with 1 medium granule and low density of small granules;
<emphasis id="B97E703C28440663057EFB25C63DF8D4" bold="true" box="[264,337,1230,1256]" pageId="4" pageNumber="20">femur</emphasis>
,
<emphasis id="B97E703C284406630514FB24C6D6F8D5" bold="true" box="[354,442,1231,1257]" pageId="4" pageNumber="20">patella,</emphasis>
and
<emphasis id="B97E703C284406630581FB25C541F8D4" bold="true" box="[503,557,1230,1256]" pageId="4" pageNumber="20">tibia</emphasis>
with minute granules, row 4 of
<emphasis id="B97E703C2844066307DAFB25C48EF8D4" bold="true" box="[940,994,1230,1256]" pageId="4" pageNumber="20">tibia</emphasis>
with sparse small spines;
<emphasis id="B97E703C28440663016EFB24C2F0F8D5" bold="true" box="[1304,1436,1231,1257]" pageId="4" pageNumber="20">metatarsus</emphasis>
practically smooth.
</paragraph>
<paragraph id="8BB5AC2E2844066304B1FAFCC599F945" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FAFCC677F90D" bold="true" box="[199,283,1303,1329]" pageId="4" pageNumber="20">Leg III</emphasis>
(figure 2D):
<emphasis id="B97E703C2844066305DBFAFDC687F90C" bold="true" box="[429,491,1302,1328]" pageId="4" pageNumber="20">Coxa</emphasis>
with 1 row of minute granules;
<emphasis id="B97E703C28440663073BFAFCC4A0F90D" bold="true" box="[845,972,1303,1329]" pageId="4" pageNumber="20">trochanter</emphasis>
with 1 medium granule and a low density of small granules;
<emphasis id="B97E703C28440663051EFAD1C6DDF968" bold="true" box="[360,433,1338,1364]" pageId="4" pageNumber="20">femur</emphasis>
with row 3 with spines close to each other in the apical region;
<emphasis id="B97E703C2844066300F6FAD1C3A5F968" bold="true" box="[1152,1225,1338,1364]" pageId="4" pageNumber="20">femur</emphasis>
,
<emphasis id="B97E703C2844066300A1FAD0C242F969" bold="true" box="[1239,1326,1339,1365]" pageId="4" pageNumber="20">patella,</emphasis>
and
<emphasis id="B97E703C284406630110FAD1C2F0F968" bold="true" box="[1382,1436,1338,1364]" pageId="4" pageNumber="20">tibia</emphasis>
with minute granules;
<emphasis id="B97E703C2844066305E4FAB4C57AF945" bold="true" box="[402,534,1375,1401]" pageId="4" pageNumber="20">metatarsus</emphasis>
practically smooth.
</paragraph>
<paragraph id="8BB5AC2E2844066304B1FA68C504FA49" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1FA68C677F9A1" bold="true" box="[199,283,1411,1437]" pageId="4" pageNumber="20">Leg IV</emphasis>
(figure 2E):
<emphasis id="B97E703C2844066305D9FA69C681F9A0" bold="true" box="[431,493,1410,1436]" pageId="4" pageNumber="20">Coxa</emphasis>
with low density of minute granules,
<emphasis id="B97E703C2844066307EDFA69C38CF9A0" bold="true" box="[923,1248,1410,1437]" pageId="4" pageNumber="20">apical prolateral apophysis</emphasis>
long and almost transverse with falciform apex,
<emphasis id="B97E703C284406630676FA4DC434F9FC" bold="true" box="[512,856,1446,1473]" pageId="4" pageNumber="20">apical retrolateral apophysis</emphasis>
like a small tubercle;
<emphasis id="B97E703C284406630021FA4CC3BAF9FD" bold="true" box="[1111,1238,1447,1473]" pageId="4" pageNumber="20">trochanter</emphasis>
with low density of minute granules,
<emphasis id="B97E703C28440663050AFA20C5D4F9D8" bold="true" box="[380,696,1482,1509]" pageId="4" pageNumber="20">basal prolateral apophysis</emphasis>
as a strong tubercle,
<emphasis id="B97E703C2844066307D8FA21C22BF9D8" bold="true" box="[942,1351,1482,1509]" pageId="4" pageNumber="20">apical dorsal prolateral apophysis</emphasis>
curved upwards,
<emphasis id="B97E703C284406630575FA05C53AFA34" bold="true" box="[259,598,1518,1545]" pageId="4" pageNumber="20">apical retrolateral apophysis</emphasis>
small, pointed, and with half size of the dorsal-prolateral apophysis;
<emphasis id="B97E703C284406630122FA05C2F1FA34" bold="true" box="[1364,1437,1518,1544]" pageId="4" pageNumber="20">femur</emphasis>
slightly S-shaped almost straight, with row 2 with 2 strong spines, close to each other, in the median to last third region;
<emphasis id="B97E703C284406630499F9DDC654FA6C" bold="true" box="[239,312,1590,1616]" pageId="4" pageNumber="20">femur</emphasis>
,
<emphasis id="B97E703C28440663053EF9DCC6F5FA6D" bold="true" box="[328,409,1591,1617]" pageId="4" pageNumber="20">patella</emphasis>
and
<emphasis id="B97E703C2844066305A5F9DDC565FA6C" bold="true" box="[467,521,1590,1616]" pageId="4" pageNumber="20">tibia</emphasis>
with medium spines in apical portion;
<emphasis id="B97E703C2844066307BDF9DCC323FA6D" bold="true" box="[971,1103,1591,1617]" pageId="4" pageNumber="20">metatarsus</emphasis>
with minute granules;
<emphasis id="B97E703C28440663012EF9DCC7B4FA49" bold="true" pageId="4" pageNumber="20">tarsal claws</emphasis>
smooth;
<emphasis id="B97E703C284406630537F9B0C688FA49" bold="true" box="[321,484,1627,1653]" pageId="4" pageNumber="20">tarsal process</emphasis>
very small.
</paragraph>
<paragraph id="8BB5AC2E2844066304B1F994C5B6FAA5" blockId="4.[151,1437,474,1941]" box="[199,730,1662,1689]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1F994C6ADFAA4" bold="true" box="[199,449,1662,1689]" pageId="4" pageNumber="20">Tarsal segmentation:</emphasis>
910; 1617; 11; 1112.
</paragraph>
<paragraph id="8BB5AC2E2844066304B1F949C22AFBA9" blockId="4.[151,1437,474,1941]" pageId="4" pageNumber="20">
<emphasis id="B97E703C2844066304B1F949C66BFA80" bold="true" box="[199,263,1698,1724]" pageId="4" pageNumber="20">Penis</emphasis>
(figure 3CD):
<emphasis id="B97E703C2844066305B6F948C531FA81" bold="true" box="[448,605,1699,1725]" pageId="4" pageNumber="20">Ventral plate</emphasis>
hexagonal notably broader than long,
<emphasis id="B97E703C284406630063F948C3AEFA80" bold="true" box="[1045,1218,1698,1725]" pageId="4" pageNumber="20">lateral margin</emphasis>
convex and
<emphasis id="B97E703C284406630123F949C782FADC" bold="true" pageId="4" pageNumber="20">apical margin</emphasis>
slightly concave; all macrosetae spatulate;
<emphasis id="B97E703C284406630682F92CC436FADC" bold="true" box="[756,858,1734,1761]" pageId="4" pageNumber="20">group A</emphasis>
with 3 macrosetae arranged almost transversely;
<emphasis id="B97E703C2844066304E1F900C65AFB39" bold="true" box="[151,310,1771,1797]" pageId="4" pageNumber="20">macrosetae B</emphasis>
similar and aligned with the macrosetae of group A;
<emphasis id="B97E703C28440663070FF900C4B6FB38" bold="true" box="[889,986,1770,1797]" pageId="4" pageNumber="20">group C</emphasis>
with 3 pairs of macrosetae, C1 separated of the others;
<emphasis id="B97E703C284406630558F8E4C6AEFB15" bold="true" box="[302,450,1807,1833]" pageId="4" pageNumber="20">macroseta D</emphasis>
half the size of the macrosetae of group C, aligned basally to group C; two
<emphasis id="B97E703C28440663008BF8E4C2F1FB15" bold="true" box="[1277,1437,1807,1833]" pageId="4" pageNumber="20">macrosetae E</emphasis>
half the size of the macrosetae of group C, E1 positioned between the two apical macrosetae of group C (C1 and C2) and E2 more ventral and between the two more basal macrosetae C (C2 and C3);
<emphasis id="B97E703C284406630044F8BCC31CFB4D" bold="true" box="[1074,1136,1879,1905]" pageId="4" pageNumber="20">glans</emphasis>
has a long ventral process with a wide flabellum at the apex, with many projections on ventral margin;
<emphasis id="B97E703C284406630787F890C359FBA9" bold="true" box="[1009,1077,1915,1941]" pageId="4" pageNumber="20">stylus</emphasis>
straight and cylindrical.
</paragraph>
<caption id="DF75FCA62845066204E1F87CC47FFBCB" ID-DOI="http://doi.org/10.5281/zenodo.15012454" ID-Zenodo-Dep="15012454" httpUri="https://zenodo.org/record/15012454/files/figure.png" pageId="5" pageNumber="21" startId="5.[151,250,1943,1967]" targetBox="[250,1339,198,1906]" targetPageId="5" targetType="figure">
<paragraph id="8BB5AC2E2845066204E1F87CC47FFBCB" blockId="5.[151,1437,1943,2040]" pageId="5" pageNumber="21">
<emphasis id="B97E703C2845066204E1F87CC67EFB93" bold="true" box="[151,274,1943,1967]" pageId="5" pageNumber="21">FIGURE 2.</emphasis>
AE male holotype (MZSP 18974) of
<taxonomicName id="4C0AD7AD2845066206E3F873C432FB8C" authorityName="Gomes-Silva &amp; Dasilva" authorityYear="2025" box="[661,862,1944,1968]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="hara" status="sp. nov.">
<emphasis id="B97E703C2845066206E3F873C432FB8C" box="[661,862,1944,1968]" italics="true" pageId="5" pageNumber="21">Serracutisoma hara</emphasis>
</taxonomicName>
<emphasis id="B97E703C284506620715F873C4DFFB8C" bold="true" box="[867,947,1944,1968]" pageId="5" pageNumber="21">
<taxonomicNameLabel id="A24DCD47284506620715F873C4DFFB8C" box="[867,947,1944,1968]" pageId="5" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AB habitus: A) dorsal view; B) lateral view; C) pedipalp, ventral-prolateral view; D) femur III, retrolateral view, showing armature of row 3; E) trochanter and femur IV, dorsal view, showing armature of row 2 of femur. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF75FCA62846066104E1F9AAC7AAFA9D" ID-DOI="http://doi.org/10.5281/zenodo.15012456" ID-Zenodo-Dep="15012456" httpUri="https://zenodo.org/record/15012456/files/figure.png" pageId="6" pageNumber="22" startId="6.[151,250,1601,1625]" targetBox="[165,1422,187,1562]" targetPageId="6" targetType="figure">
<paragraph id="8BB5AC2E2846066104E1F9AAC7AAFA9D" blockId="6.[151,1437,1601,1697]" pageId="6" pageNumber="22">
<emphasis id="B97E703C2846066104E1F9AAC678FA65" bold="true" box="[151,276,1601,1625]" pageId="6" pageNumber="22">FIGURE 3.</emphasis>
AB penis of male holotype (MZSP 31022) of
<taxonomicName id="4C0AD7AD284606610689F9A9C49EFA65" authorityName="Gomes-Silva &amp; Dasilva" authorityYear="2025" box="[767,1010,1601,1625]" class="Arachnida" family="Gonyleptidae" genus="Goniosoma" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="22" phylum="Arthropoda" rank="species" species="bragagnolo" status="sp. nov.">
<emphasis id="B97E703C284606610689F9A9C49EFA65" box="[767,1010,1601,1625]" italics="true" pageId="6" pageNumber="22">Goniosoma bragagnolo</emphasis>
</taxonomicName>
<emphasis id="B97E703C28460661078FF9A9C327FA66" bold="true" box="[1017,1099,1602,1626]" pageId="6" pageNumber="22">
<taxonomicNameLabel id="A24DCD4728460661078FF9A9C327FA66" box="[1017,1099,1602,1626]" pageId="6" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A) dorsal view; B) lateral view. CD penis of male holotype (MZSP 18974) of
<taxonomicName id="4C0AD7AD28460661060DF98DC42AFA41" authorityName="Gomes-Silva &amp; Dasilva" authorityYear="2025" box="[635,838,1637,1661]" class="Arachnida" family="Gonyleptidae" genus="Serracutisoma" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="22" phylum="Arthropoda" rank="species" species="hara" status="sp. nov.">
<emphasis id="B97E703C28460661060DF98DC42AFA41" box="[635,838,1637,1661]" italics="true" pageId="6" pageNumber="22">Serracutisoma hara</emphasis>
</taxonomicName>
<emphasis id="B97E703C28460661073BF98DC4F3FA42" bold="true" box="[845,927,1638,1662]" pageId="6" pageNumber="22">
<taxonomicNameLabel id="A24DCD4728460661073BF98DC4F3FA42" box="[845,927,1638,1662]" pageId="6" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: C) dorsal view; D) lateral view. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB5AC2E2846066104B1F91FC791FB0E" blockId="6.[151,1437,1779,2022]" pageId="6" pageNumber="22">
<emphasis id="B97E703C2846066104B1F91FC550FB32" bold="true" box="[199,572,1780,1806]" pageId="6" pageNumber="22">Measurements. Dorsal scutum.</emphasis>
Length: 8.6; width: 8.3;
<emphasis id="B97E703C28460661072CF918C4A1FB31" bold="true" box="[858,973,1779,1805]" pageId="6" pageNumber="22">pedipalps</emphasis>
: 13.8;
<emphasis id="B97E703C284606610068F91FC31DFB32" bold="true" box="[1054,1137,1780,1806]" pageId="6" pageNumber="22">legs: I:</emphasis>
47.2;
<emphasis id="B97E703C2846066100CCF91FC3B6FB32" bold="true" box="[1210,1242,1780,1806]" pageId="6" pageNumber="22">II:</emphasis>
102.8;
<emphasis id="B97E703C284606610147F91FC237FB32" bold="true" box="[1329,1371,1780,1806]" pageId="6" pageNumber="22">III:</emphasis>
71.9;
<emphasis id="B97E703C2846066104E1F8F3C7D1FB0E" bold="true" box="[151,189,1816,1842]" pageId="6" pageNumber="22">IV:</emphasis>
95.3.
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(in alcohol):
<emphasis id="B97E703C284606610590F8D7C439FB69" bold="true" box="[486,853,1851,1878]" pageId="6" pageNumber="22">dorsal scutum, legs IIII, tibia</emphasis>
,
<emphasis id="B97E703C284606610713F8D7C37DFB6A" bold="true" box="[869,1041,1852,1878]" pageId="6" pageNumber="22">metatarsus IV</emphasis>
,
<emphasis id="B97E703C284606610056F8D0C3FFFB69" bold="true" box="[1056,1171,1851,1877]" pageId="6" pageNumber="22">pedipalps</emphasis>
and
<emphasis id="B97E703C2846066100BBF8D0C2F1FB6A" bold="true" box="[1229,1437,1851,1878]" pageId="6" pageNumber="22">spines of area III</emphasis>
have a light-yellow coloration;
<emphasis id="B97E703C284606610588F88BC598FB46" bold="true" box="[510,756,1888,1914]" pageId="6" pageNumber="22">carapace, apophyses</emphasis>
of the
<emphasis id="B97E703C28460661073EF88BC23DFB46" bold="true" box="[840,1361,1888,1914]" pageId="6" pageNumber="22">coxa and trochanter IV, leg IV, trochanters</emphasis>
of the other legs, and the ventral side of the
<emphasis id="B97E703C28460661062CF86FC5F9FBA2" bold="true" box="[602,661,1924,1950]" pageId="6" pageNumber="22">body</emphasis>
dark brown;
<emphasis id="B97E703C284606610740F868C35AFBA2" bold="true" box="[822,1078,1923,1950]" pageId="6" pageNumber="22">articular membranes</emphasis>
light pink;
<emphasis id="B97E703C2846066100B5F86FC255FBA2" bold="true" box="[1219,1337,1924,1950]" pageId="6" pageNumber="22">dry mark</emphasis>
only on trochanter IV and the base of femur IV.
</paragraph>
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<emphasis id="B97E703C2846066104B1F827C644FBDA" bold="true" box="[199,296,1996,2022]" pageId="6" pageNumber="22">Female:</emphasis>
Unknown.
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Genus
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<bibRefCitation id="EF9F4B13FFB4FFD9FE54617AFB06F8AA" author="Kolbe, H. J." box="[425,574,1823,1849]" pageId="0" pageNumber="84" pagination="121 - 132" refId="ref1041" refString="Kolbe, H. J. (1908) Dynastidae, Cetoniidae und Scarabaeidae. In: Schultze, L. (Ed.), Zoologische und Arthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Sudafrika. Band. 1. Denkschrift Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena, 13, pp. 121-132." type="journal article" year="1908">Kolbe, 1908</bibRefCitation>
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(gender: masculinum)
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<emphasis id="B97AEAF0FFB4FFD9FF6A6125F99DF8CB" bold="true" box="[151,165,1856,1880]" pageId="0" pageNumber="84">=</emphasis>
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<emphasis id="B97AEAF0FFB4FFD9FF516125F818F8CB" box="[172,288,1856,1880]" italics="true" pageId="0" pageNumber="84">Byrrhidium</emphasis>
<bibRefCitation id="EF9F4B13FFB4FFD9FEDA6125F895F8CB" author="Harold, E. von" box="[295,429,1856,1880]" pageId="0" pageNumber="84" pagination="95 - 104" refId="ref987" refString="Harold, E. von (1869) Diagnosen neuer Coprophagen. Coleopterologische Hefte, 5, 95-104." type="journal article" year="1869">Harold, 1869</bibRefCitation>
</taxonomicName>
<emphasis id="B97AEAF0FFB4FFD9FE4E615AFB50F8CB" bold="true" box="[435,616,1855,1880]" pageId="0" pageNumber="84">junior homonym</emphasis>
(gender: neutrum)
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<emphasis id="B91D4615FFA243500EEDFF11FE3487DE" bold="true" box="[151,436,151,178]" italics="true" pageId="78" pageNumber="79">Petralycus caryapecaus</emphasis>
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(
<figureCitation id="13528682FFA243500EE5FF3DFEA887BA" box="[159,296,187,213]" captionStart="FIGURE 47" captionStartId="77.[151,250,1888,1912]" captionTargetBox="[151,1436,176,1864]" captionTargetId="figure-15@77.[151,1436,176,1864]" captionTargetPageId="77" captionText="FIGURE 47. Petralycus caryapecaus McDaniel &amp; Bolen. A—prodorsum; B—holotrichous dorsum; C—leg I; D—leg II; E—leg III; F—leg IV; G—palp; H—subcapitulum, chelicera without a seta; J—genital valves with 12 genital setae per valve; K—photograph showing parallel winding ridges and a microshield. All specimens—from Texas, San Patricio Co., copied from McDaniel &amp; Bolen (1983)." pageId="78" pageNumber="79">Fig. 47AK</figureCitation>
)
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, figs. 34;
<treatmentCitation id="0AC8BC16FFA243500D42FF79FC708678" author="Uusitalo, M." box="[824,1008,255,280]" page="76" pageId="78" pageNumber="79" year="2010">
<bibRefCitation id="EFF8E7F6FFA243500D42FF79FC708678" author="Uusitalo, M." box="[824,1008,255,280]" pageId="78" pageNumber="79" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010: 76</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA243500EEDFEC3FCED8583" pageId="78" pageNumber="79" type="description">
<paragraph id="8BD69A07FFA243500EEDFEC3FE9D86A3" blockId="78.[151,1437,325,1576]" pageId="78" pageNumber="79">
<emphasis id="B91D4615FFA243500EEDFEC3FEC58630" bold="true" box="[151,325,325,351]" pageId="78" pageNumber="79">Redescription.</emphasis>
<emphasis id="B91D4615FFA243500F34FEC1FE28860F" box="[334,424,327,352]" italics="true" pageId="78" pageNumber="79">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFA243500FA7FEC0FDC0860F" box="[477,576,326,352]" count="1" pageId="78" pageNumber="79" type="female">1 female</specimenCount>
,
<figureCitation id="13528682FFA243500C34FEC0FC80860F" box="[590,768,326,352]" captionStart="FIGURE 47" captionStartId="77.[151,250,1888,1912]" captionTargetBox="[151,1436,176,1864]" captionTargetId="figure-15@77.[151,1436,176,1864]" captionTargetPageId="77" captionText="FIGURE 47. Petralycus caryapecaus McDaniel &amp; Bolen. A—prodorsum; B—holotrichous dorsum; C—leg I; D—leg II; E—leg III; F—leg IV; G—palp; H—subcapitulum, chelicera without a seta; J—genital valves with 12 genital setae per valve; K—photograph showing parallel winding ridges and a microshield. All specimens—from Texas, San Patricio Co., copied from McDaniel &amp; Bolen (1983)." pageId="78" pageNumber="79">Fig. 47A, B, K</figureCitation>
). Length 222 µm; holotrichous number of plumose dorsal setae, slightly larger in caudal segments; naso ca. 9 µm, narrow and rounded at tip, 3 pairs of ciliate prodorsal setae (
<emphasis id="B91D4615FFA243500EDBFE09FF3586C7" box="[161,181,399,424]" italics="true" pageId="78" pageNumber="79">vi</emphasis>
,
<emphasis id="B91D4615FFA243500EB9FE09FF6786C7" box="[195,231,399,424]" italics="true" pageId="78" pageNumber="79">sce</emphasis>
,
<emphasis id="B91D4615FFA243500E8EFE09FE8A86C7" box="[244,266,399,424]" italics="true" pageId="78" pageNumber="79">in</emphasis>
), setae
<emphasis id="B91D4615FFA243500F1AFE09FE0786C7" box="[352,391,399,424]" italics="true" pageId="78" pageNumber="79">exp</emphasis>
and eyes absent, sensilla
<emphasis id="B91D4615FFA243500CDDFE09FD4686C7" box="[679,710,399,424]" italics="true" pageId="78" pageNumber="79">sci</emphasis>
filiform with short cilia, sensilla
<emphasis id="B91D4615FFA243500A41FE09FBD486C7" box="[1083,1108,399,424]" italics="true" pageId="78" pageNumber="79">ve</emphasis>
rounded, all inserted on hard integument.
</paragraph>
<paragraph id="8BD69A07FFA243500EBDFE51FD18857B" blockId="78.[151,1437,325,1576]" pageId="78" pageNumber="79">
<taxonomicName id="4C69E184FFA243500EBDFE51FE8F869F" box="[199,271,471,496]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="78" pageNumber="79" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFA243500EBDFE51FE8F869F" box="[199,271,471,496]" italics="true" pageId="78" pageNumber="79">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFA243500F5AFE50FE0C869F" box="[288,396,470,496]" captionStart="FIGURE 47" captionStartId="77.[151,250,1888,1912]" captionTargetBox="[151,1436,176,1864]" captionTargetId="figure-15@77.[151,1436,176,1864]" captionTargetPageId="77" captionText="FIGURE 47. Petralycus caryapecaus McDaniel &amp; Bolen. A—prodorsum; B—holotrichous dorsum; C—leg I; D—leg II; E—leg III; F—leg IV; G—palp; H—subcapitulum, chelicera without a seta; J—genital valves with 12 genital setae per valve; K—photograph showing parallel winding ridges and a microshield. All specimens—from Texas, San Patricio Co., copied from McDaniel &amp; Bolen (1983)." pageId="78" pageNumber="79">Figs. 47J</figureCitation>
;
<bibRefCitation id="EFF8E7F6FFA243500FE3FE50FD38869F" author="McDaniel, B. &amp; Bolen, E. G." box="[409,696,470,496]" pageId="78" pageNumber="79" pagination="811 - 817" refId="ref48059" refString="McDaniel, B. &amp; Bolen, E. G. (1983) Mites of the family Pachygnathidae (Acari: Endeostigmata) from Texas. Proceedings of the Entomological Society of Washington, 85 (4), 811-817." type="journal article" year="1983">McDaniel &amp; Bolen 1983</bibRefCitation>
: fig. 4a, c). Ventral side neotrichous with small stellate setae; 12 genital setae per valve; 4 anal setae per valve.
</paragraph>
<paragraph id="8BD69A07FFA243500EBDFD98FECB85EF" blockId="78.[151,1437,325,1576]" pageId="78" pageNumber="79">
<taxonomicName id="4C69E184FFA243500EBDFD98FED68557" authorityName="Krefft" authorityYear="1871" box="[199,342,542,568]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="78" pageNumber="79" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFA243500EBDFD98FED68557" box="[199,342,542,568]" italics="true" pageId="78" pageNumber="79">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFA243500F1EFD98FE6C8557" box="[356,492,542,568]" captionStart="FIGURE 47" captionStartId="77.[151,250,1888,1912]" captionTargetBox="[151,1436,176,1864]" captionTargetId="figure-15@77.[151,1436,176,1864]" captionTargetPageId="77" captionText="FIGURE 47. Petralycus caryapecaus McDaniel &amp; Bolen. A—prodorsum; B—holotrichous dorsum; C—leg I; D—leg II; E—leg III; F—leg IV; G—palp; H—subcapitulum, chelicera without a seta; J—genital valves with 12 genital setae per valve; K—photograph showing parallel winding ridges and a microshield. All specimens—from Texas, San Patricio Co., copied from McDaniel &amp; Bolen (1983)." pageId="78" pageNumber="79">Fig. 47G, H</figureCitation>
). 3 pairs of plumose and elongated setae on subcapitulum; chelicerae chelate-dentate, robust, and elongated without cheliceral setae; large palpal solenidion reaching over two apical eupathidia, tactile setae elongated.
</paragraph>
<paragraph id="8BD69A07FFA243500EBDFD0DFCED8583" blockId="78.[151,1437,325,1576]" pageId="78" pageNumber="79">
<emphasis id="B91D4615FFA243500EBDFD0DFF7C85CB" box="[199,252,651,676]" italics="true" pageId="78" pageNumber="79">Legs</emphasis>
(
<figureCitation id="13528682FFA243500F75FD0CFE1985CB" box="[271,409,650,676]" captionStart="FIGURE 47" captionStartId="77.[151,250,1888,1912]" captionTargetBox="[151,1436,176,1864]" captionTargetId="figure-15@77.[151,1436,176,1864]" captionTargetPageId="77" captionText="FIGURE 47. Petralycus caryapecaus McDaniel &amp; Bolen. A—prodorsum; B—holotrichous dorsum; C—leg I; D—leg II; E—leg III; F—leg IV; G—palp; H—subcapitulum, chelicera without a seta; J—genital valves with 12 genital setae per valve; K—photograph showing parallel winding ridges and a microshield. All specimens—from Texas, San Patricio Co., copied from McDaniel &amp; Bolen (1983)." pageId="78" pageNumber="79">Fig. 47CF</figureCitation>
). Pedal setae ciliate and elongated; 18 foot-shaped eupathidia ventrodistally on tarsus I; solenidial formula for tarsi, tibiae, genua and femora on legs I, II, III and IV, respectively: 1-1-0-0, 3-2-1-1, 6-2-1-2, 3-3-0-0+0; famulus I placed abaxially, near distal end of tarsus I.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA243500EBDFD73FC6E84CF" pageId="78" pageNumber="79" type="materials_examined">
<paragraph id="8BD69A07FFA243500EBDFD73FC6E84CF" blockId="78.[151,1437,325,1576]" pageId="78" pageNumber="79">
<emphasis id="B91D4615FFA243500EBDFD73FE2E8460" bold="true" box="[199,430,757,783]" pageId="78" pageNumber="79">Material examined.</emphasis>
<specimenCount id="9D6F518EFFA243500FC9FD70FD92847F" box="[435,530,758,784]" count="1" pageId="78" pageNumber="79" type="female">1 female</specimenCount>
as
<taxonomicName id="4C69E184FFA243500C49FD70FCBE847F" box="[563,830,758,784]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="caryapecaus">
<emphasis id="B91D4615FFA243500C49FD70FCBE847F" box="[563,830,758,784]" italics="true" pageId="78" pageNumber="79">Petralycus caryapecaus</emphasis>
</taxonomicName>
from sandy soil, Moorei Creek, National Battleground, Penden Co.,
<collectingRegion id="49AD54E5FFA243500F5DFC9CFE53845B" box="[295,467,794,820]" country="United States of America" name="North Carolina" pageId="78" pageNumber="79">North Carolina</collectingRegion>
,
<collectingCountry id="F37EDA97FFA243500FA4FC9CFD99845B" box="[478,537,794,820]" name="United States of America" pageId="78" pageNumber="79">USA</collectingCountry>
,
<date id="FFD7BCC7FFA243500C5FFC9CFD7D845B" box="[549,765,794,820]" pageId="78" pageNumber="79" value="1988-12-23">23 December 1988</date>
, Eric G. Bolen, slide TAMU-ENTO X1646472 at the
<collectingRegion id="49AD54E5FFA243500B21FC9CFA1C845B" box="[1371,1436,794,820]" country="United States of America" name="Texas" pageId="78" pageNumber="79">Texas</collectingRegion>
A&amp;M University Insect Collection, Department of Entomology. The examined specimen labelled as
<taxonomicName id="4C69E184FFA243500B5EFCB8FEA58413" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="caryapecaus">
<emphasis id="B91D4615FFA243500B5EFCB8FEA58413" italics="true" pageId="78" pageNumber="79">Petralycus caryapecaus</emphasis>
</taxonomicName>
by B. McDaniel was in very poor condition, shrunken and with an air bubble inside; therefore, the redescription is based on the original description (
<bibRefCitation id="EFF8E7F6FFA243500CBCFC00FC6284CF" author="McDaniel, B. &amp; Bolen, E. G." box="[710,994,902,928]" pageId="78" pageNumber="79" pagination="811 - 817" refId="ref48059" refString="McDaniel, B. &amp; Bolen, E. G. (1983) Mites of the family Pachygnathidae (Acari: Endeostigmata) from Texas. Proceedings of the Entomological Society of Washington, 85 (4), 811-817." type="journal article" year="1983">McDaniel &amp; Bolen 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA243500EBDFC2FFABC833B" pageId="78" pageNumber="79" type="diagnosis">
<paragraph id="8BD69A07FFA243500EBDFC2FFABC833B" blockId="78.[151,1437,325,1576]" pageId="78" pageNumber="79">
<emphasis id="B91D4615FFA243500EBDFC2FFE5284AC" bold="true" box="[199,466,937,963]" pageId="78" pageNumber="79">Differential diagnosis.</emphasis>
The naso of
<taxonomicName id="4C69E184FFA243500C08FC2DFCA084AB" box="[626,800,939,964]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="caryapecaus">
<emphasis id="B91D4615FFA243500C08FC2DFCA084AB" box="[626,800,939,964]" italics="true" pageId="78" pageNumber="79">P. caryapecaus</emphasis>
</taxonomicName>
is ca. half the size of the naso of
<taxonomicName id="4C69E184FFA243500AC6FC2DFADF84AB" box="[1212,1375,938,964]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA243500AC6FC2DFADF84AB" box="[1212,1375,938,964]" italics="true" pageId="78" pageNumber="79">P. celtisacinus</emphasis>
</taxonomicName>
. The solenidial pattern is unique as shown in the table above. The American species
<taxonomicName id="4C69E184FFA243500A66FC49FB3C8487" box="[1052,1212,974,1000]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA243500A66FC49FB3C8487" box="[1052,1212,974,1000]" italics="true" pageId="78" pageNumber="79">P. celtisacinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFA243500A8FFC49FA1C8487" authorityName="Uusitalo" authorityYear="2025" box="[1269,1436,974,1000]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="obtusicornis">
<emphasis id="B91D4615FFA243500A8FFC49FA1C8487" box="[1269,1436,974,1000]" italics="true" pageId="78" pageNumber="79">P. obtusicornis</emphasis>
</taxonomicName>
have chelicerae with a seta but the chelicerae of
<taxonomicName id="4C69E184FFA243500CD3FC75FCD18363" box="[681,849,1011,1036]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="caryapecaus">
<emphasis id="B91D4615FFA243500CD3FC75FCD18363" box="[681,849,1011,1036]" italics="true" pageId="78" pageNumber="79">P. caryapecaus</emphasis>
</taxonomicName>
and the European
<taxonomicName id="4C69E184FFA243500A67FC75FB218363" authorityName="Grandjean" authorityYear="1943" box="[1053,1185,1011,1036]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B91D4615FFA243500A67FC75FB218363" box="[1053,1185,1011,1036]" italics="true" pageId="78" pageNumber="79">P. unicornis</emphasis>
</taxonomicName>
are bald. All four have a holotrichous setal pattern while the African
<taxonomicName id="4C69E184FFA243500CE7FB91FCBA835F" authorityName="Theron" authorityYear="1977" box="[669,826,1046,1072]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B91D4615FFA243500CE7FB91FCBA835F" box="[669,826,1046,1072]" italics="true" pageId="78" pageNumber="79">P. longicornis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFA243500D08FB91FB8D835F" authorityName="Theron" authorityYear="1977" box="[882,1037,1046,1072]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B91D4615FFA243500D08FB91FB8D835F" box="[882,1037,1046,1072]" italics="true" pageId="78" pageNumber="79">P. brevicornis</emphasis>
</taxonomicName>
are neotrichous. The naso is blunt-tipped in the American species and
<taxonomicName id="4C69E184FFA243500C5FFBBDFD41833B" authorityName="Theron" authorityYear="1977" box="[549,705,1082,1108]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B91D4615FFA243500C5FFBBDFD41833B" box="[549,705,1082,1108]" italics="true" pageId="78" pageNumber="79">P. longicornis</emphasis>
</taxonomicName>
but with a sharp point in
<taxonomicName id="4C69E184FFA243500DA5FBBDFBE5833B" authorityName="Grandjean" authorityYear="1943" box="[991,1125,1083,1108]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B91D4615FFA243500DA5FBBDFBE5833B" box="[991,1125,1083,1108]" italics="true" pageId="78" pageNumber="79">P. unicornis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFA243500AE1FBBDFAB5833B" authorityName="Theron" authorityYear="1977" box="[1179,1333,1082,1108]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B91D4615FFA243500AE1FBBDFAB5833B" box="[1179,1333,1082,1108]" italics="true" pageId="78" pageNumber="79">P. brevicornis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA243500EBDFBD8FC8A8147" pageId="78" pageNumber="79" type="discussion">
<paragraph id="8BD69A07FFA243500EBDFBD8FB468267" blockId="78.[151,1437,325,1576]" pageId="78" pageNumber="79">
<emphasis id="B91D4615FFA243500EBDFBD8FEBB8317" bold="true" box="[199,315,1118,1144]" pageId="78" pageNumber="79">Remarks.</emphasis>
According to
<bibRefCitation id="EFF8E7F6FFA243500FAFFBD8FD7C8317" author="McDaniel, B. &amp; Bolen, E. G." box="[469,764,1118,1144]" pageId="78" pageNumber="79" pagination="811 - 817" refId="ref48059" refString="McDaniel, B. &amp; Bolen, E. G. (1983) Mites of the family Pachygnathidae (Acari: Endeostigmata) from Texas. Proceedings of the Entomological Society of Washington, 85 (4), 811-817." type="journal article" year="1983">McDaniel &amp; Bolen (1983)</bibRefCitation>
: “The
<typeStatus id="54D224A5FFA243500D38FBD8FC248317" box="[834,932,1118,1144]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
specimen of
<taxonomicName id="4C69E184FFA243500A4EFBD9FB5C8317" box="[1076,1244,1119,1144]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="caryapecaus">
<emphasis id="B91D4615FFA243500A4EFBD9FB5C8317" box="[1076,1244,1119,1144]" italics="true" pageId="78" pageNumber="79">P. caryapecaus</emphasis>
</taxonomicName>
will be deposited with National Museum of Natural History,
<collectingRegion id="49AD54E5FFA243500C0FFB04FD7D83F3" box="[629,765,1154,1180]" country="United States of America" name="Washington" pageId="78" pageNumber="79">Washington</collectingRegion>
, D.C.”—but the specimen is not in their database (Dr. Debra Creel, in litt.). The
<typeStatus id="54D224A5FFA243500F08FB20FE5483AF" box="[370,468,1190,1216]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
was collected in San Patricio Co.,
<collectingRegion id="49AD54E5FFA243500D1BFB20FC2583AF" box="[865,933,1190,1216]" country="United States of America" name="Texas" pageId="78" pageNumber="79">Texas</collectingRegion>
, a riparian woodland known as Pecan Motte with features virtually identical with those for Hackberry Motte. The mixed tree stands, among other things with hackberry (
<taxonomicName id="4C69E184FFA243500F6DFB68FED88267" box="[279,344,1262,1288]" class="Magnoliopsida" family="Cannabaceae" genus="Celtis" kingdom="Plantae" order="Rosales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFA243500F6DFB68FED88267" box="[279,344,1262,1288]" italics="true" pageId="78" pageNumber="79">Celtis</emphasis>
</taxonomicName>
) and pecan (
<taxonomicName id="4C69E184FFA243500F90FB68FDB08267" box="[490,560,1262,1288]" class="Magnoliopsida" family="Juglandaceae" genus="Carya" kingdom="Plantae" order="Fagales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFA243500F90FB68FDB08267" box="[490,560,1262,1288]" italics="true" pageId="78" pageNumber="79">Carya</emphasis>
</taxonomicName>
), obviously inspired the naming of both American species.
</paragraph>
<paragraph id="8BD69A07FFA243500EBDFA94FC8A8147" blockId="78.[151,1437,325,1576]" pageId="78" pageNumber="79">
The original figures of
<taxonomicName id="4C69E184FFA243500FAAFA95FDF18243" box="[464,625,1298,1324]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA243500FAAFA95FDF18243" box="[464,625,1298,1324]" italics="true" pageId="78" pageNumber="79">P. celtisacinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFA243500CD0FA94FCD38243" box="[682,851,1298,1324]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="82" phylum="Arthropoda" rank="species" species="caryapecaus">
P.
<emphasis id="B91D4615FFA243500CBFFA95FCD38243" box="[709,851,1299,1324]" italics="true" pageId="78" pageNumber="79">caryapecaus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF8E7F6FFA243500D19FA94FB088243" author="McDaniel, B. &amp; Bolen, E. G." box="[867,1160,1298,1324]" pageId="78" pageNumber="79" pagination="811 - 817" refId="ref48059" refString="McDaniel, B. &amp; Bolen, E. G. (1983) Mites of the family Pachygnathidae (Acari: Endeostigmata) from Texas. Proceedings of the Entomological Society of Washington, 85 (4), 811-817." type="journal article" year="1983">McDaniel &amp; Bolen, 1983</bibRefCitation>
, figs. 14) are distorted, without scale bars and the observations and measurements based on the figures are not quite reliable. The total length 222 µm given for
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<emphasis id="B91D4615FFA243500FC9FADDFDDD821B" box="[435,605,1371,1396]" italics="true" pageId="78" pageNumber="79">P. caryapecaus</emphasis>
</taxonomicName>
in the text was measured in fig. 47B from the nasal tip to caudal end and used in adjusting the scale bars for the fragments of the figure. The solenidial pattern of
<taxonomicName id="4C69E184FFA243500A24FAF9FB7C82F7" box="[1118,1276,1406,1432]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="celtisacinus">
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in the original description corresponded with the examined material, and the solenidiotaxy of
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<emphasis id="B91D4615FFA243500A4EFA25FB6182D3" box="[1076,1249,1443,1468]" italics="true" pageId="78" pageNumber="79">P. caryapecaus</emphasis>
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is also reliable. However, a sceptical attitude is advisable in regards to minor character states, such as absence of barbs on sensillus
<emphasis id="B91D4615FFA243500EEDFA6DFF30816B" box="[151,176,1515,1540]" italics="true" pageId="78" pageNumber="79">ve</emphasis>
(as mentioned in text on p. 814, but present in fig. 3f = 47A), absence of a cheliceral seta, and absence of a pointed rutellum, until fresh material is available and examined.
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<emphasis id="B91D4615FFA543570EEDF91BFE9681D7" bold="true" box="[151,278,1693,1720]" italics="true" pageId="73" pageNumber="74">Petralycus</emphasis>
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species:
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from
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,
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.
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<emphasis id="B91D4615FFA543570EEDF8AEFE1B802D" bold="true" box="[151,411,1832,1858]" pageId="73" pageNumber="74">Differential diagnosis.</emphasis>
The genus can be segregated from the other genera of
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and most of the
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by their prodorsum with less than six pairs of fundamental setae, i.e. the pair of setae
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is absent, the anterior pair of prodorsal sensilla
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being oval or globular, and the pointed or strongly tapering naso. Also, the first pair of legs is longer and thicker than legs IIIV with many foot-shaped eupathidia ventrodistally.
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<emphasis id="B91D4615FFA543570EBDF83EFEB080BD" bold="true" box="[199,304,1976,2002]" pageId="73" pageNumber="74">Remark.</emphasis>
The assessments and comments on
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from
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Petralycini
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genus:
<taxonomicName id="4C69E184FFA543570F62FAB1FDB48220" authority="Grandjean, 1943" authorityName="Grandjean" authorityYear="1943" box="[280,564,1335,1359]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFA543570F62FAB1FE048220" box="[280,388,1335,1359]" italics="true" pageId="73" pageNumber="74">Petralycus</emphasis>
<bibRefCitation id="EFF8E7F6FFA543570FF1FAB1FDB48220" author="Grandjean, F." box="[395,564,1335,1359]" pageId="73" pageNumber="74" pagination="1 - 59" refId="ref47160" refString="Grandjean, F. (1943), Quelques genres d'Acariens appartenant au groupe des Endeostigmata, 2 e Serie. Deuxieme partie. Annales des Sciences naturelles, Zoologie, 11 e Serie, 5, 1-59 (137 - 195)." type="journal article" year="1943">Grandjean, 1943</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA543570EEDFAFBFE8A8124" pageId="73" pageNumber="74" type="diagnosis">
<paragraph id="8BD69A07FFA543570EEDFAFBFE8A8124" blockId="73.[151,1437,1405,1611]" pageId="73" pageNumber="74">
<emphasis id="B91D4615FFA543570EEDFAFBFE2182F8" bold="true" box="[151,417,1405,1431]" pageId="73" pageNumber="74">Differential diagnosis.</emphasis>
This tribe (and genus
<taxonomicName id="4C69E184FFA543570CD3FAFBFCA182F8" authorityName="Grandjean" authorityYear="1943" box="[681,801,1405,1431]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFA543570CD3FAFBFCA182F8" box="[681,801,1405,1431]" italics="true" pageId="73" pageNumber="74">Petralycus</emphasis>
</taxonomicName>
) can be segregated from the rest of the
<taxonomicName id="4C69E184FFA543570A8CFAFBFA1C82F8" authorityName="Reuter" authorityYear="1909" box="[1270,1436,1405,1431]" pageId="73" pageNumber="92" rank="subOrder" subOrder="Endeostigmata">Endeostigmata</taxonomicName>
by the following combination of characters: the two pairs of prodorsal sensilla in separate bothridia, anterior pair
<emphasis id="B91D4615FFA543570EEDFA40FF3082B0" box="[151,176,1478,1503]" italics="true" pageId="73" pageNumber="74">ve</emphasis>
being globular and posterior pair
<emphasis id="B91D4615FFA543570C3FFA40FDE482B0" box="[581,612,1478,1503]" italics="true" pageId="73" pageNumber="74">sci</emphasis>
filamentous (
<figureCitation id="13528682FFA543570D7DFA43FCF3828F" box="[775,883,1477,1504]" captionStart="FIGURE 45" captionStartId="75.[151,250,1833,1857]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-15@75.[151,1436,193,1809]" captionTargetPageId="75" captionText="FIGURE 45. Petralycus celtisacinus McDaniel &amp; Bolen. A—prodorsum, setae in with elongated shaft (dorsal integument badly broken, left sensillus ve out of its socket); B—holotrichous number of microshields on dorsal integument, setae longest on caudal segments H and PS; C—gnathosoma, (subcapitulum, chelicera with a seta and palp separated); D—coxae III, IV and genital valves with 14 setae in 2 rows. All specimens—from Texas, San Patricio Co." pageId="73" pageNumber="74">Fig. 45A</figureCitation>
); strongly elongated but robust chelicerae (
<figureCitation id="13528682FFA543570B15FA43FF4A816B" captionStart="FIGURE 45" captionStartId="75.[151,250,1833,1857]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-15@75.[151,1436,193,1809]" captionTargetPageId="75" captionText="FIGURE 45. Petralycus celtisacinus McDaniel &amp; Bolen. A—prodorsum, setae in with elongated shaft (dorsal integument badly broken, left sensillus ve out of its socket); B—holotrichous number of microshields on dorsal integument, setae longest on caudal segments H and PS; C—gnathosoma, (subcapitulum, chelicera with a seta and palp separated); D—coxae III, IV and genital valves with 14 setae in 2 rows. All specimens—from Texas, San Patricio Co." pageId="73" pageNumber="74">Fig. 45C</figureCitation>
); palpal tarsi having a solenidion extending partly over a pair of distal mucronate eupathidia (
<figureCitation id="13528682FFA543570A9EFA6FFACB816B" box="[1252,1355,1513,1540]" captionStart="FIGURE 45" captionStartId="75.[151,250,1833,1857]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-15@75.[151,1436,193,1809]" captionTargetPageId="75" captionText="FIGURE 45. Petralycus celtisacinus McDaniel &amp; Bolen. A—prodorsum, setae in with elongated shaft (dorsal integument badly broken, left sensillus ve out of its socket); B—holotrichous number of microshields on dorsal integument, setae longest on caudal segments H and PS; C—gnathosoma, (subcapitulum, chelicera with a seta and palp separated); D—coxae III, IV and genital valves with 14 setae in 2 rows. All specimens—from Texas, San Patricio Co." pageId="73" pageNumber="74">Fig. 45C</figureCitation>
); pedal solenidia varies from claviform, and baculiform to uniquely thin, but not tapering (
<figureCitation id="13528682FFA543570A4AF98BFB358148" box="[1072,1205,1549,1575]" captionStart="FIGURE 46" captionStartId="76.[151,250,1778,1802]" captionTargetBox="[151,1435,176,1754]" captionTargetId="figure-24@76.[151,1436,176,1754]" captionTargetPageId="76" captionText="FIGURE 46. Petralycus celtisacinus McDaniel &amp; Bolen. A—leg I dorsal view; B—tarsus I ventral view with 33 barbed eupathidia; C—tarsus I lateral view with famulus I and foot shaped eupathidia; D—leg II; E—tibia, genu and femur III; F—tibia, genu and telofemur IV; G—anal plates with 4 insertion points for sparsely ciliated anal setae; H—prodorsum with two pairs of sensilla, median eye and massive rhagidiid-like chelicera in Proterorhagiidae. 46AG—from Texas, San Patricio Co.; 46H—from Colima, Mexico, copied from Lindquist &amp; Palacios- Vargas (1991)." pageId="73" pageNumber="74">Fig. 46AF</figureCitation>
;
<bibRefCitation id="EFF8E7F6FFA543570ABAF98BFAE18148" author="Uusitalo, M." box="[1216,1377,1549,1575]" pageId="73" pageNumber="74" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: figs. 147149).
</paragraph>
</subSubSection>
</treatment>
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View file

@ -0,0 +1,352 @@
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<paragraph id="8BD69A07FFA643540EEDFF11FD6587DE" blockId="74.[151,741,151,214]" box="[151,741,151,178]" pageId="74" pageNumber="75">
<heading id="D09E2D6BFFA643540EEDFF11FD6587DE" bold="true" box="[151,741,151,178]" fontSize="11" level="2" pageId="74" pageNumber="75" reason="4">
<taxonomicName id="4C69E184FFA643540EEDFF11FD6587DE" ID-CoL="4FDK8" authority="McDaniel &amp; Bolen, 1983" authorityName="McDaniel &amp; Bolen" authorityYear="1983" box="[151,741,151,178]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA643540EEDFF11FD6587DE" bold="true" box="[151,741,151,178]" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540EEDFF11FE2C87DD" bold="true" box="[151,428,151,178]" italics="true" pageId="74" pageNumber="75">Petralycus celtisacinus</emphasis>
<bibRefCitation id="EFF8E7F6FFA643540FC9FF11FD6587DE" author="McDaniel, B. &amp; Bolen, E. G." box="[435,741,151,177]" pageId="74" pageNumber="75" pagination="811 - 817" refId="ref48059" refString="McDaniel, B. &amp; Bolen, E. G. (1983) Mites of the family Pachygnathidae (Acari: Endeostigmata) from Texas. Proceedings of the Entomological Society of Washington, 85 (4), 811-817." type="journal article" year="1983">McDaniel &amp; Bolen, 1983</bibRefCitation>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA643540EEDFF3DFE1B87BA" box="[151,411,187,214]" pageId="74" pageNumber="75" type="description">
<paragraph id="8BD69A07FFA643540EEDFF3DFE1B87BA" blockId="74.[151,741,151,214]" box="[151,411,187,214]" pageId="74" pageNumber="75">
(
<figureCitation id="13528682FFA643540EE5FF3DFEB387B9" box="[159,307,187,214]" captionStart="FIGURE 45" captionStartId="75.[151,250,1833,1857]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-15@75.[151,1436,193,1809]" captionTargetPageId="75" captionText="FIGURE 45. Petralycus celtisacinus McDaniel &amp; Bolen. A—prodorsum, setae in with elongated shaft (dorsal integument badly broken, left sensillus ve out of its socket); B—holotrichous number of microshields on dorsal integument, setae longest on caudal segments H and PS; C—gnathosoma, (subcapitulum, chelicera with a seta and palp separated); D—coxae III, IV and genital valves with 14 setae in 2 rows. All specimens—from Texas, San Patricio Co." pageId="74" pageNumber="75">Figs. 45AD</figureCitation>
,
<figureCitation id="13528682FFA643540F45FF3DFE1387BA" box="[319,403,187,213]" captionStart="FIGURE 46" captionStartId="76.[151,250,1778,1802]" captionTargetBox="[151,1435,176,1754]" captionTargetId="figure-24@76.[151,1436,176,1754]" captionTargetPageId="76" captionText="FIGURE 46. Petralycus celtisacinus McDaniel &amp; Bolen. A—leg I dorsal view; B—tarsus I ventral view with 33 barbed eupathidia; C—tarsus I lateral view with famulus I and foot shaped eupathidia; D—leg II; E—tibia, genu and femur III; F—tibia, genu and telofemur IV; G—anal plates with 4 insertion points for sparsely ciliated anal setae; H—prodorsum with two pairs of sensilla, median eye and massive rhagidiid-like chelicera in Proterorhagiidae. 46AG—from Texas, San Patricio Co.; 46H—from Colima, Mexico, copied from Lindquist &amp; Palacios- Vargas (1991)." pageId="74" pageNumber="75">46AG</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA643540EEDFE86FC6B8678" box="[151,1003,255,280]" pageId="74" pageNumber="75" type="reference_group">
<paragraph id="8BD69A07FFA643540EEDFE86FC6B8678" blockId="74.[151,1003,255,280]" box="[151,1003,255,280]" pageId="74" pageNumber="75">
<treatmentCitationGroup id="AB79BD29FFA643540EEDFE86FC6B8678" box="[151,1003,255,280]" pageId="74" pageNumber="75">
<treatmentCitation id="0AC8BC16FFA643540EEDFE86FD3E8677" author="McDaniel, B. &amp; Bolen, E. G." box="[151,702,256,280]" page="811" pageId="74" pageNumber="75" year="1983">
<taxonomicName id="4C69E184FFA643540EEDFE86FD3E8677" ID-CoL="4FDK8" authority="McDaniel &amp; Bolen, 1983: 811" authorityName="McDaniel &amp; Bolen" authorityPageNumber="811" authorityYear="1983" box="[151,702,256,280]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA643540EEDFE86FE028677" box="[151,386,256,280]" italics="true" pageId="74" pageNumber="75">Petralycus celtisacinus</emphasis>
<bibRefCitation id="EFF8E7F6FFA643540FF2FE86FD3E8677" author="McDaniel, B. &amp; Bolen, E. G." box="[392,702,256,280]" pageId="74" pageNumber="75" pagination="811 - 817" refId="ref48059" refString="McDaniel, B. &amp; Bolen, E. G. (1983) Mites of the family Pachygnathidae (Acari: Endeostigmata) from Texas. Proceedings of the Entomological Society of Washington, 85 (4), 811-817." type="journal article" year="1983">McDaniel &amp; Bolen, 1983: 811</bibRefCitation>
</taxonomicName>
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, figs. 12;
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.
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</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA643540EEDFEC3FCED847F" pageId="74" pageNumber="75" type="description">
<paragraph id="8BD69A07FFA643540EEDFEC3FC4F869F" blockId="74.[151,1437,325,1504]" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540EEDFEC3FEC58630" bold="true" box="[151,325,325,351]" pageId="74" pageNumber="75">Redescription.</emphasis>
<emphasis id="B91D4615FFA643540F33FEC1FE23860F" box="[329,419,327,352]" italics="true" pageId="74" pageNumber="75">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFA643540FB5FEC0FDBC860F" box="[463,572,326,352]" count="2" pageId="74" pageNumber="75" type="female">2 females</specimenCount>
,
<figureCitation id="13528682FFA643540C3EFEC0FD46860F" box="[580,710,326,353]" captionStart="FIGURE 45" captionStartId="75.[151,250,1833,1857]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-15@75.[151,1436,193,1809]" captionTargetPageId="75" captionText="FIGURE 45. Petralycus celtisacinus McDaniel &amp; Bolen. A—prodorsum, setae in with elongated shaft (dorsal integument badly broken, left sensillus ve out of its socket); B—holotrichous number of microshields on dorsal integument, setae longest on caudal segments H and PS; C—gnathosoma, (subcapitulum, chelicera with a seta and palp separated); D—coxae III, IV and genital valves with 14 setae in 2 rows. All specimens—from Texas, San Patricio Co." pageId="74" pageNumber="75">Fig. 45A, B</figureCitation>
). Length 246 µm; dorsum holotrichous, setae born on microplates, each seta plumose with a slightly swollen but tapering shaft, setae longest on caudal segments; lamellae inserted quite densely at regular intervals on continuous striae; naso ca. 20 µm, narrow and rounded at tip; setae
<emphasis id="B91D4615FFA643540B5FFE09FAB986C7" box="[1317,1337,399,424]" italics="true" pageId="74" pageNumber="75">vi</emphasis>
sparsely ciliated;
<emphasis id="B91D4615FFA643540E80FE35FE9E86A3" box="[250,286,435,460]" italics="true" pageId="74" pageNumber="75">sce</emphasis>
reduced in size; setae
<emphasis id="B91D4615FFA643540C5AFE35FDB686A3" box="[544,566,435,460]" italics="true" pageId="74" pageNumber="75">in</emphasis>
slightly elongated; setae
<emphasis id="B91D4615FFA643540D2DFE35FCFE86A3" box="[855,894,435,460]" italics="true" pageId="74" pageNumber="75">exp</emphasis>
and eyes absent; sensilla
<emphasis id="B91D4615FFA643540ADEFE35FB4386A3" box="[1188,1219,435,460]" italics="true" pageId="74" pageNumber="75">sci</emphasis>
filiform with short cilia; sensilla
<emphasis id="B91D4615FFA643540F4BFE51FECA869F" box="[305,330,471,496]" italics="true" pageId="74" pageNumber="75">ve</emphasis>
rounded, all prodorsal setae inserted on hard integument.
</paragraph>
<paragraph id="8BD69A07FFA643540EBDFE7DFDC48557" blockId="74.[151,1437,325,1504]" pageId="74" pageNumber="75">
<taxonomicName id="4C69E184FFA643540EBDFE7DFE8F857B" box="[199,271,507,532]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="74" pageNumber="75" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFA643540EBDFE7DFE8F857B" box="[199,271,507,532]" italics="true" pageId="74" pageNumber="75">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFA643540F64FE7CFE10857A" box="[286,400,506,533]" captionStart="FIGURE 45" captionStartId="75.[151,250,1833,1857]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-15@75.[151,1436,193,1809]" captionTargetPageId="75" captionText="FIGURE 45. Petralycus celtisacinus McDaniel &amp; Bolen. A—prodorsum, setae in with elongated shaft (dorsal integument badly broken, left sensillus ve out of its socket); B—holotrichous number of microshields on dorsal integument, setae longest on caudal segments H and PS; C—gnathosoma, (subcapitulum, chelicera with a seta and palp separated); D—coxae III, IV and genital valves with 14 setae in 2 rows. All specimens—from Texas, San Patricio Co." pageId="74" pageNumber="75">Figs. 45D</figureCitation>
,
<figureCitation id="13528682FFA643540FE6FE7CFE51857B" box="[412,465,506,532]" captionStart="FIGURE 46" captionStartId="76.[151,250,1778,1802]" captionTargetBox="[151,1435,176,1754]" captionTargetId="figure-24@76.[151,1436,176,1754]" captionTargetPageId="76" captionText="FIGURE 46. Petralycus celtisacinus McDaniel &amp; Bolen. A—leg I dorsal view; B—tarsus I ventral view with 33 barbed eupathidia; C—tarsus I lateral view with famulus I and foot shaped eupathidia; D—leg II; E—tibia, genu and femur III; F—tibia, genu and telofemur IV; G—anal plates with 4 insertion points for sparsely ciliated anal setae; H—prodorsum with two pairs of sensilla, median eye and massive rhagidiid-like chelicera in Proterorhagiidae. 46AG—from Texas, San Patricio Co.; 46H—from Colima, Mexico, copied from Lindquist &amp; Palacios- Vargas (1991)." pageId="74" pageNumber="75">46G</figureCitation>
). Ventral side neotrichous with small stellate setae; 14 genital setae per valve, 1 pair of eugenital setae; 4 anal setae per valve.
</paragraph>
<paragraph id="8BD69A07FFA643540EBDFDC4FECB85CB" blockId="74.[151,1437,325,1504]" pageId="74" pageNumber="75">
<taxonomicName id="4C69E184FFA643540EBDFDC4FED68533" authorityName="Krefft" authorityYear="1871" box="[199,342,578,604]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="74" pageNumber="75" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFA643540EBDFDC4FED68533" box="[199,342,578,604]" italics="true" pageId="74" pageNumber="75">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFA643540F19FDC4FE478532" box="[355,455,578,605]" captionStart="FIGURE 45" captionStartId="75.[151,250,1833,1857]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-15@75.[151,1436,193,1809]" captionTargetPageId="75" captionText="FIGURE 45. Petralycus celtisacinus McDaniel &amp; Bolen. A—prodorsum, setae in with elongated shaft (dorsal integument badly broken, left sensillus ve out of its socket); B—holotrichous number of microshields on dorsal integument, setae longest on caudal segments H and PS; C—gnathosoma, (subcapitulum, chelicera with a seta and palp separated); D—coxae III, IV and genital valves with 14 setae in 2 rows. All specimens—from Texas, San Patricio Co." pageId="74" pageNumber="75">Fig. 45C</figureCitation>
). 3 pairs of elongated setae on subcapitulum; chelicerae chelate-dentate,robust, elongated, with 1 pair of smooth cheliceral setae; large palpal solenidion reaching over two nude and apical eupathidia, tactile setae elongated.
</paragraph>
<paragraph id="8BD69A07FFA643540EBDFD29FCED847F" blockId="74.[151,1437,325,1504]" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540EBDFD29FF7C85A7" box="[199,252,687,712]" italics="true" pageId="74" pageNumber="75">Legs</emphasis>
(
<figureCitation id="13528682FFA643540F76FD28FE1D85A7" box="[268,413,686,712]" captionStart="FIGURE 46" captionStartId="76.[151,250,1778,1802]" captionTargetBox="[151,1435,176,1754]" captionTargetId="figure-24@76.[151,1436,176,1754]" captionTargetPageId="76" captionText="FIGURE 46. Petralycus celtisacinus McDaniel &amp; Bolen. A—leg I dorsal view; B—tarsus I ventral view with 33 barbed eupathidia; C—tarsus I lateral view with famulus I and foot shaped eupathidia; D—leg II; E—tibia, genu and femur III; F—tibia, genu and telofemur IV; G—anal plates with 4 insertion points for sparsely ciliated anal setae; H—prodorsum with two pairs of sensilla, median eye and massive rhagidiid-like chelicera in Proterorhagiidae. 46AG—from Texas, San Patricio Co.; 46H—from Colima, Mexico, copied from Lindquist &amp; Palacios- Vargas (1991)." pageId="74" pageNumber="75">Figs. 46AF</figureCitation>
). Pedal setae ciliate and elongated; ca. 30 foot-shaped eupathidia ventrodistally on tarsus I; solenidial formula for tarsi, tibiae, genua and femora on legs I, II, III and IV, respectively: 1-1-0-0, 2-2-2-2, 5-2-2-2, 5-1-1-2+0; famulus I placed abaxially, near distal end of tarsus I.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA643540EBDFC9FFC2B84AB" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="8BD69A07FFA643540EBDFC9FFC2B84AB" blockId="74.[151,1437,325,1504]" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540EBDFC9FFE33845C" bold="true" box="[199,435,793,819]" pageId="74" pageNumber="75">Material examined.</emphasis>
<specimenCount id="9D6F518EFFA643540FC7FC9CFDAC845B" box="[445,556,794,820]" count="2" pageId="74" pageNumber="75" type="female">2 females</specimenCount>
as
<taxonomicName id="4C69E184FFA643540C2DFC9CFD4F845B" authorityName="Grandjean" authorityYear="1943" box="[599,719,794,820]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFA643540C2DFC9CFD4F845B" box="[599,719,794,820]" italics="true" pageId="74" pageNumber="75">Petralycus</emphasis>
</taxonomicName>
from mesquite grass (
<taxonomicName id="4C69E184FFA643540DAFFC9CFB1C845B" box="[981,1180,794,820]" class="Liliopsida" family="Poaceae" genus="Hilaria" kingdom="Plantae" order="Poales" pageId="74" pageNumber="75" phylum="Tracheophyta" rank="species" species="belangeri">
<emphasis id="B91D4615FFA643540DAFFC9CFB1C845B" box="[981,1180,794,820]" italics="true" pageId="74" pageNumber="75">Hilaria belangeri</emphasis>
</taxonomicName>
), a riparian woodland known as Hackberry Motte, Monument Road, Walder Foundation, Sinton, San Patricio Co.,
<collectingRegion id="49AD54E5FFA643540A9BFCB8FAA58437" box="[1249,1317,830,856]" country="United States of America" name="Texas" pageId="74" pageNumber="75">Texas</collectingRegion>
,
<collectingCountry id="F37EDA97FFA643540B4FFCB8FAF08437" box="[1333,1392,830,856]" name="United States of America" pageId="74" pageNumber="75">USA</collectingCountry>
,
<date id="FFD7BCC7FFA643540BFAFCB8FED28413" pageId="74" pageNumber="75" value="1977-11-23">23 November 1977</date>
, Eric G. Bolen, slides TAMU-ENTO X1645288 (used for drawings), TAMU-ENTO X1642983 (collected
<date id="FFD7BCC7FFA643540F76FC00FE6584CF" box="[268,485,902,928]" pageId="74" pageNumber="75" value="1977-11-23">23 November 1977</date>
) and TAMU-ENTO X1642865, nymph (collected
<date id="FFD7BCC7FFA643540A66FC00FB5984CF" box="[1052,1241,902,928]" pageId="74" pageNumber="75" value="1977-02-02">2 February 1977</date>
); deposited at the
<collectingRegion id="49AD54E5FFA643540EEDFC2CFF5884AB" box="[151,216,938,964]" country="United States of America" name="Texas" pageId="74" pageNumber="75">Texas</collectingRegion>
A&amp;M University Insect Collection, Department of Entomology.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA643540EBDFC4BFBB58317" pageId="74" pageNumber="75" type="diagnosis">
<paragraph id="8BD69A07FFA643540EBDFC4BFBB58317" blockId="74.[151,1437,325,1504]" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540EBDFC4BFE4F8488" bold="true" box="[199,463,973,999]" pageId="74" pageNumber="75">Differential diagnosis.</emphasis>
The six presently known species of the genus
<taxonomicName id="4C69E184FFA643540DA5FC48FBD78487" authorityName="Grandjean" authorityYear="1943" box="[991,1111,974,1000]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFA643540DA5FC48FBD78487" box="[991,1111,974,1000]" italics="true" pageId="74" pageNumber="75">Petralycus</emphasis>
</taxonomicName>
, each have their own unique solenidiotaxy (
<tableCitation id="C6EBAFBCFFA643540F44FC74FE148363" box="[318,404,1010,1036]" captionStart="TABLE 1" captionStartId="74.[151,239,1551,1575]" captionText="TABLE 1. Solenidiotaxy of six presently known Petralycus species" pageId="74" pageNumber="75">Table 1</tableCitation>
). The American species
<taxonomicName id="4C69E184FFA643540CCBFC75FCD28363" box="[689,850,1010,1036]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA643540CCBFC75FCD28363" box="[689,850,1010,1036]" italics="true" pageId="74" pageNumber="75">P. celtisacinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFA643540DF6FC75FBB38363" authorityName="Uusitalo" authorityYear="2025" box="[908,1075,1010,1036]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="obtusicornis">
<emphasis id="B91D4615FFA643540DF6FC75FBB38363" box="[908,1075,1010,1036]" italics="true" pageId="74" pageNumber="75">P. obtusicornis</emphasis>
</taxonomicName>
have chelicerae with a seta but the chelicerae of
<taxonomicName id="4C69E184FFA643540F23FB91FD84835F" box="[345,516,1047,1072]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="caryapecaus">
<emphasis id="B91D4615FFA643540F23FB91FD84835F" box="[345,516,1047,1072]" italics="true" pageId="74" pageNumber="75">P. caryapecaus</emphasis>
</taxonomicName>
and the European
<taxonomicName id="4C69E184FFA643540CA2FB91FCDE835F" authorityName="Grandjean" authorityYear="1943" box="[728,862,1047,1072]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B91D4615FFA643540CA2FB91FCDE835F" box="[728,862,1047,1072]" italics="true" pageId="74" pageNumber="75">P. unicornis</emphasis>
</taxonomicName>
are bald. All four have a holotrichous setal pattern while the African species
<taxonomicName id="4C69E184FFA643540FCEFBBDFDCF833B" authorityName="Theron" authorityYear="1977" box="[436,591,1082,1108]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B91D4615FFA643540FCEFBBDFDCF833B" box="[436,591,1082,1108]" italics="true" pageId="74" pageNumber="75">P. longicornis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFA643540CF9FBBDFC9C833B" authorityName="Theron" authorityYear="1977" box="[643,796,1082,1108]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B91D4615FFA643540CF9FBBDFC9C833B" box="[643,796,1082,1108]" italics="true" pageId="74" pageNumber="75">P. brevicornis</emphasis>
</taxonomicName>
are neotrichous. The naso is blunt-tipped in the American species and
<taxonomicName id="4C69E184FFA643540F64FBD9FE3A8317" authorityName="Theron" authorityYear="1977" box="[286,442,1118,1144]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B91D4615FFA643540F64FBD9FE3A8317" box="[286,442,1118,1144]" italics="true" pageId="74" pageNumber="75">P. longicornis</emphasis>
</taxonomicName>
but with a sharp point in
<taxonomicName id="4C69E184FFA643540CA2FBD9FCDE8317" authorityName="Grandjean" authorityYear="1943" box="[728,862,1119,1144]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B91D4615FFA643540CA2FBD9FCDE8317" box="[728,862,1119,1144]" italics="true" pageId="74" pageNumber="75">P. unicornis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFA643540DEEFBD9FBAE8317" authorityName="Theron" authorityYear="1977" box="[916,1070,1118,1144]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis id="B91D4615FFA643540DEEFBD9FBAE8317" box="[916,1070,1118,1144]" italics="true" pageId="74" pageNumber="75">P. brevicornis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFA643540EBDFB04FB1C80AC" pageId="74" pageNumber="75" type="discussion">
<paragraph id="8BD69A07FFA643540EBDFB04FEB5838B" blockId="74.[151,1437,325,1504]" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540EBDFB04FEBB83F3" bold="true" box="[199,315,1154,1180]" pageId="74" pageNumber="75">Remarks.</emphasis>
According to
<bibRefCitation id="EFF8E7F6FFA643540FADFB04FC8083F3" author="McDaniel, B. &amp; Bolen, E. G." box="[471,768,1154,1180]" pageId="74" pageNumber="75" pagination="811 - 817" refId="ref48059" refString="McDaniel, B. &amp; Bolen, E. G. (1983) Mites of the family Pachygnathidae (Acari: Endeostigmata) from Texas. Proceedings of the Entomological Society of Washington, 85 (4), 811-817." type="journal article" year="1983">McDaniel &amp; Bolen (1983)</bibRefCitation>
: “The
<typeStatus id="54D224A5FFA643540D3DFB04FC2983F3" box="[839,937,1154,1180]" pageId="74" pageNumber="75" type="holotype">holotype</typeStatus>
specimen of
<taxonomicName id="4C69E184FFA643540A41FB05FB5983F3" box="[1083,1241,1154,1180]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA643540A41FB05FB5983F3" box="[1083,1241,1154,1180]" italics="true" pageId="74" pageNumber="75">P. celtisacinus</emphasis>
</taxonomicName>
will be deposited with National Museum of Natural History,
<collectingRegion id="49AD54E5FFA643540C0EFB20FD7C83AF" box="[628,764,1190,1216]" country="United States of America" name="Washington" pageId="74" pageNumber="75">Washington</collectingRegion>
, D.C.”—but the specimen is not in their database (Dr. Debra Creel, in litt.).
</paragraph>
<paragraph id="8BD69A07FFA643540EBDFB68FE60828F" blockId="74.[151,1437,325,1504]" pageId="74" pageNumber="75">
The
<specimenCount id="9D6F518EFFA643540E83FB68FE2F8267" box="[249,431,1262,1288]" count="3" pageId="74" pageNumber="75" type="generic">three specimens</specimenCount>
, labelled only as
<taxonomicName id="4C69E184FFA643540C17FB68FD658267" authorityName="Grandjean" authorityYear="1943" box="[621,741,1262,1288]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFA643540C17FB68FD658267" box="[621,741,1262,1288]" italics="true" pageId="74" pageNumber="75">Petralycus</emphasis>
</taxonomicName>
, but collected five months earlier from the same locality as the
<typeStatus id="54D224A5FFA643540EEDFA94FF798243" box="[151,249,1298,1324]" pageId="74" pageNumber="75" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C69E184FFA643540F64FA95FE3E8243" box="[286,446,1298,1324]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA643540F64FA95FE3E8243" box="[286,446,1298,1324]" italics="true" pageId="74" pageNumber="75">P. celtisacinus</emphasis>
</taxonomicName>
by Eric G. Bolen, are assumed here to be conspecific with
<taxonomicName id="4C69E184FFA643540A2CFA95FB758243" box="[1110,1269,1298,1324]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA643540A2CFA95FB758243" box="[1110,1269,1298,1324]" italics="true" pageId="74" pageNumber="75">P. celtisacinus</emphasis>
</taxonomicName>
because of the similar solenidiotaxy, in spite of scanty numbers of tactile setae. The adult specimen used in the redescription bears no dorsal or anal setae at all, but there is a holotrichous number of tiny microplates (33) indicating the insertion points of lost dorsal setae on the tattered integument, but held together by ventral skin (between two black lines in
<figureCitation id="13528682FFA643540EEDFA24FE8082D2" box="[151,256,1442,1469]" captionStart="FIGURE 45" captionStartId="75.[151,250,1833,1857]" captionTargetBox="[151,1436,193,1809]" captionTargetId="figure-15@75.[151,1436,193,1809]" captionTargetPageId="75" captionText="FIGURE 45. Petralycus celtisacinus McDaniel &amp; Bolen. A—prodorsum, setae in with elongated shaft (dorsal integument badly broken, left sensillus ve out of its socket); B—holotrichous number of microshields on dorsal integument, setae longest on caudal segments H and PS; C—gnathosoma, (subcapitulum, chelicera with a seta and palp separated); D—coxae III, IV and genital valves with 14 setae in 2 rows. All specimens—from Texas, San Patricio Co." pageId="74" pageNumber="75">Fig. 45A</figureCitation>
). Setae from the two other specimens were used as models in the depiction. The length of the species is from the original description.
</paragraph>
<caption id="DF16CA8FFFA643540EEDF989FCEC8147" box="[151,876,1551,1576]" pageId="74" pageNumber="75" startId="74.[151,239,1551,1575]" targetType="table">
<paragraph id="8BD69A07FFA643540EEDF989FCEC8147" blockId="74.[151,876,1551,1576]" box="[151,876,1551,1576]" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540EEDF989FE8A8148" bold="true" box="[151,266,1551,1575]" pageId="74" pageNumber="75">TABLE 1.</emphasis>
Solenidiotaxy of six presently known
<taxonomicName id="4C69E184FFA643540CDCF996FCEC8147" box="[678,876,1552,1576]" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540CDCF996FC988147" box="[678,792,1552,1576]" italics="true" pageId="74" pageNumber="75">Petralycus</emphasis>
species
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BD69A07FFA643540ED9F9BEFA93802E" pageId="74" pageNumber="75">
<table id="F96968A7FFA6BCE10ED9F9BEFA9A802E" box="[163,1306,1592,1857]" gridcols="5" gridrows="7" pageId="74" pageNumber="75">
<tr id="35599845FFA6BCE10ED9F9BEFA9A813F" box="[163,1306,1592,1616]" gridrow="0" pageId="74" pageNumber="75">
<th id="7688F139FFA6BCE10ED9F9BEFEBB813F" box="[163,315,1592,1616]" gridcol="0" gridrow="0" pageId="74" pageNumber="75">species</th>
<th id="7688F139FFA6BCE10FDEF9BEFE70813F" box="[420,496,1592,1616]" gridcol="1" gridrow="0" pageId="74" pageNumber="75">tarsi</th>
<th id="7688F139FFA6BCE10CDFF9BEFD70813F" box="[677,752,1592,1616]" gridcol="2" gridrow="0" pageId="74" pageNumber="75">tibiae</th>
<th id="7688F139FFA6BCE10DDCF9BEFB85813F" box="[934,1029,1592,1616]" gridcol="3" gridrow="0" pageId="74" pageNumber="75">genua</th>
<th id="7688F139FFA6BCE10ADDF9BEFA9A813F" box="[1191,1306,1592,1616]" gridcol="4" gridrow="0" pageId="74" pageNumber="75">femora</th>
</tr>
<tr id="35599845FFA6BCE10ED9F9E6FA9A8117" box="[163,1306,1632,1656]" gridrow="1" pageId="74" pageNumber="75">
<th id="7688F139FFA6BCE10ED9F9E6FEBB8117" box="[163,315,1632,1656]" gridcol="0" gridrow="1" pageId="74" pageNumber="75">
<taxonomicName id="4C69E184FFA643540ED9F9E6FEA08117" authorityName="Uusitalo" authorityYear="2025" box="[163,288,1632,1656]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="obtusicornis">
<emphasis id="B91D4615FFA643540ED9F9E6FEA08117" box="[163,288,1632,1656]" italics="true" pageId="74" pageNumber="75">obtusicornis</emphasis>
</taxonomicName>
</th>
<td id="7688F139FFA6BCE10FDEF9E6FE708117" box="[420,496,1632,1656]" gridcol="1" gridrow="1" pageId="74" pageNumber="75">1-1-0-0</td>
<td id="7688F139FFA6BCE10CDFF9E6FD708117" box="[677,752,1632,1656]" gridcol="2" gridrow="1" pageId="74" pageNumber="75">2-2-2-2</td>
<td id="7688F139FFA6BCE10DDCF9E6FB858117" box="[934,1029,1632,1656]" gridcol="3" gridrow="1" pageId="74" pageNumber="75">9-2-2-5</td>
<td id="7688F139FFA6BCE10ADDF9E6FA9A8117" box="[1191,1306,1632,1656]" gridcol="4" gridrow="1" pageId="74" pageNumber="75">12-3-1-3+1</td>
</tr>
<tr id="35599845FFA6BCE10ED9F90EFA9A81CF" box="[163,1306,1672,1696]" gridrow="2" pageId="74" pageNumber="75">
<th id="7688F139FFA6BCE10ED9F90EFEBB81CF" box="[163,315,1672,1696]" gridcol="0" gridrow="2" pageId="74" pageNumber="75">
<taxonomicName id="4C69E184FFA643540ED9F90EFE9B81CF" box="[163,283,1672,1696]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="celtisacinus">
<emphasis id="B91D4615FFA643540ED9F90EFE9B81CF" box="[163,283,1672,1696]" italics="true" pageId="74" pageNumber="75">celtisacinus</emphasis>
</taxonomicName>
</th>
<td id="7688F139FFA6BCE10FDEF90EFE7081CF" box="[420,496,1672,1696]" gridcol="1" gridrow="2" pageId="74" pageNumber="75">1-1-0-0</td>
<td id="7688F139FFA6BCE10CDFF90EFD7081CF" box="[677,752,1672,1696]" gridcol="2" gridrow="2" pageId="74" pageNumber="75">2-2-2-2</td>
<td id="7688F139FFA6BCE10DDCF90EFB8581CF" box="[934,1029,1672,1696]" gridcol="3" gridrow="2" pageId="74" pageNumber="75">5-2-2-2</td>
<td id="7688F139FFA6BCE10ADDF90EFA9A81CF" box="[1191,1306,1672,1696]" gridcol="4" gridrow="2" pageId="74" pageNumber="75">5-1-1-2+0</td>
</tr>
<tr id="35599845FFA6BCE10ED9F936FA9A81A6" box="[163,1306,1712,1737]" gridrow="3" pageId="74" pageNumber="75">
<th id="7688F139FFA6BCE10ED9F936FEBB81A6" box="[163,315,1712,1737]" gridcol="0" gridrow="3" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540ED9F937FEAF81A7" box="[163,303,1713,1736]" italics="true" pageId="74" pageNumber="75">
<taxonomicName id="4C69E184FFA643540ED9F937FEA681A7" box="[163,294,1713,1736]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="caryapecaus">caryapecaus</taxonomicName>
*
</emphasis>
</th>
<td id="7688F139FFA6BCE10FDEF936FE7081A6" box="[420,496,1712,1737]" gridcol="1" gridrow="3" pageId="74" pageNumber="75">1-1-0-0</td>
<td id="7688F139FFA6BCE10CDFF936FD7081A6" box="[677,752,1712,1737]" gridcol="2" gridrow="3" pageId="74" pageNumber="75">2-2-2-2</td>
<td id="7688F139FFA6BCE10DDCF936FB8581A6" box="[934,1029,1712,1737]" gridcol="3" gridrow="3" pageId="74" pageNumber="75">6-2-1-2</td>
<td id="7688F139FFA6BCE10ADDF936FA9A81A6" box="[1191,1306,1712,1737]" gridcol="4" gridrow="3" pageId="74" pageNumber="75">3-3-0-0+0</td>
</tr>
<tr id="35599845FFA6BCE10ED9F95FFA9A819E" box="[163,1306,1753,1777]" gridrow="4" pageId="74" pageNumber="75">
<th id="7688F139FFA6BCE10ED9F95FFEBB819E" box="[163,315,1753,1777]" gridcol="0" gridrow="4" pageId="74" pageNumber="75">
<taxonomicName id="4C69E184FFA643540ED9F95CFE83819E" authorityName="Grandjean" authorityYear="1943" box="[163,259,1754,1777]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B91D4615FFA643540ED9F95CFE83819E" box="[163,259,1754,1777]" italics="true" pageId="74" pageNumber="75">unicornis</emphasis>
</taxonomicName>
</th>
<td id="7688F139FFA6BCE10FDEF95FFE70819E" box="[420,496,1753,1777]" gridcol="1" gridrow="4" pageId="74" pageNumber="75">1-1-0-0</td>
<td id="7688F139FFA6BCE10CDFF95FFD70819E" box="[677,752,1753,1777]" gridcol="2" gridrow="4" pageId="74" pageNumber="75">4-2-2-2</td>
<td id="7688F139FFA6BCE10DDCF95FFB85819E" box="[934,1029,1753,1777]" gridcol="3" gridrow="4" pageId="74" pageNumber="75">5-2-2-2</td>
<td id="7688F139FFA6BCE10ADDF95FFA9A819E" box="[1191,1306,1753,1777]" gridcol="4" gridrow="4" pageId="74" pageNumber="75">3-2-0-1+0</td>
</tr>
<tr id="35599845FFA6BCE10ED9F887FA9A8076" box="[163,1306,1793,1817]" gridrow="5" pageId="74" pageNumber="75">
<th id="7688F139FFA6BCE10ED9F887FEBB8076" box="[163,315,1793,1817]" gridcol="0" gridrow="5" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540ED9F887FEAF8076" box="[163,303,1793,1817]" italics="true" pageId="74" pageNumber="75">
<taxonomicName id="4C69E184FFA643540ED9F887FE988076" authorityName="Theron" authorityYear="1977" box="[163,280,1793,1817]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="longicornis">longicornis</taxonomicName>
**
</emphasis>
</th>
<td id="7688F139FFA6BCE10FDEF887FE708076" box="[420,496,1793,1817]" gridcol="1" gridrow="5" pageId="74" pageNumber="75">1-1-0-0</td>
<td id="7688F139FFA6BCE10CDFF887FD708076" box="[677,752,1793,1817]" gridcol="2" gridrow="5" pageId="74" pageNumber="75">2-2-2-1</td>
<td id="7688F139FFA6BCE10DDCF887FB858076" box="[934,1029,1793,1817]" gridcol="3" gridrow="5" pageId="74" pageNumber="75">5-2-2-2\1</td>
<td id="7688F139FFA6BCE10ADDF887FA9A8076" box="[1191,1306,1793,1817]" gridcol="4" gridrow="5" pageId="74" pageNumber="75">2-0-0-0+1</td>
</tr>
<tr id="35599845FFA6BCE10ED9F8AFFA9A802E" box="[163,1306,1833,1857]" gridrow="6" pageId="74" pageNumber="75">
<th id="7688F139FFA6BCE10ED9F8AFFEBB802E" box="[163,315,1833,1857]" gridcol="0" gridrow="6" pageId="74" pageNumber="75">
<emphasis id="B91D4615FFA643540ED9F8AFFEBB802E" box="[163,315,1833,1857]" italics="true" pageId="74" pageNumber="75">
<taxonomicName id="4C69E184FFA643540ED9F8AFFE96802E" authorityName="Theron" authorityYear="1977" box="[163,278,1833,1857]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="brevicornis">brevicornis</taxonomicName>
***
</emphasis>
</th>
<td id="7688F139FFA6BCE10FDEF8AFFE70802E" box="[420,496,1833,1857]" gridcol="1" gridrow="6" pageId="74" pageNumber="75">1-1-0-0</td>
<td id="7688F139FFA6BCE10CDFF8AFFD70802E" box="[677,752,1833,1857]" gridcol="2" gridrow="6" pageId="74" pageNumber="75">2-2-2-2</td>
<td id="7688F139FFA6BCE10DDCF8AFFB85802E" box="[934,1029,1833,1857]" gridcol="3" gridrow="6" pageId="74" pageNumber="75">5-2-2-2</td>
<td id="7688F139FFA6BCE10ADDF8AFFA9A802E" box="[1191,1306,1833,1857]" gridcol="4" gridrow="6" pageId="74" pageNumber="75">2-0-0-0+0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD69A07FFA643540EEDF8E8FBF980CB" blockId="74.[151,1421,1901,1987]" pageId="74" pageNumber="75">
* There is only 1 solenidion on tarsus I (
<bibRefCitation id="EFF8E7F6FFA643540C55F8EBFCAF80E9" author="McDaniel, B. &amp; Bolen, E. G." box="[559,815,1901,1926]" pageId="74" pageNumber="75" pagination="811 - 817" refId="ref48059" refString="McDaniel, B. &amp; Bolen, E. G. (1983) Mites of the family Pachygnathidae (Acari: Endeostigmata) from Texas. Proceedings of the Entomological Society of Washington, 85 (4), 811-817." type="journal article" year="1983">McDaniel &amp; Bolen 1983</bibRefCitation>
: fig 3), but 4 are in the text that is apparently a typing error. ** Theron (1977: 42) reports either 3 setae and 2 solenidia or 4 setae and 1 solenidion on genu IV.
</paragraph>
<paragraph id="8BD69A07FFA643540EEDF82DFB1C80AC" blockId="74.[151,1421,1901,1987]" box="[151,1180,1963,1987]" pageId="74" pageNumber="75">
<materialsCitation id="3B01905AFFA643540EEDF82DFB1880AC" box="[151,1176,1963,1987]" collectionCode="II" location="Theron" pageId="74" pageNumber="75" specimenCount="1" typeStatus="holotype">
***
<figureCitation id="13528682FFA643540EBEF82DFEA580AC" box="[196,293,1963,1987]" captionStart="FIGURE 17" captionStartId="34.[151,250,1800,1824]" captionTargetBox="[167,1432,221,1792]" captionTargetId="figure-57@34.[151,1436,216,1792]" captionTargetPageId="34" captionText="FIGURE 17. Glabromichaelia vazquezrojasi sp. nov. A—prodorsum with elongated andciliate setae; B—reconstructed dorsal habitus, outlines andgrey dots symbolising the expected form of adult and number of missing setae, black dots represent the actually observed setae; C—neotrichous subcapitulum; D—ventral side between coxae without coxal plates. All specimens—from Veracruz, CICOLMA." pageId="74" pageNumber="75">
Figure
<quantity id="4C9137E2FFA643540F76F82DFEA580AC" box="[268,293,1963,1987]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="74" pageNumber="75" unit="in" value="17.0">17</quantity>
</figureCitation>
in
<location id="8EB6CCDCFFA643540F3FF82DFE0E80AC" LSID="urn:lsid:plazi:treatment:03C02B11FFA643540EEDFF11FB1C80AC:8EB6CCDCFFA643540F3FF82DFE0E80AC" box="[325,398,1963,1987]" name="Theron" pageId="74" pageNumber="75">Theron</location>
(1977: 43) shows no solenidia on leg
<collectionCode id="ED7802C2FFA643540D6BF82DFCA480AD" box="[785,804,1963,1986]" pageId="74" pageNumber="75">II</collectionCode>
, but they are present in the
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.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,258 @@
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<paragraph id="8BD69A07FFC743350EEDFA68FDBB8166" blockId="43.[151,571,1518,1545]" box="[151,571,1518,1545]" pageId="43" pageNumber="44">
<heading id="D09E2D6BFFC743350EEDFA68FDBB8166" bold="true" box="[151,571,1518,1545]" fontSize="11" level="2" pageId="43" pageNumber="44" reason="3">
<emphasis id="B91D4615FFC743350EEDFA68FDBB8166" bold="true" box="[151,571,1518,1545]" pageId="43" pageNumber="44">
Genus
<taxonomicName id="4C69E184FFC743350E94FA68FE458166" authorityName="Uusitalo" authorityYear="2025" box="[238,453,1518,1545]" class="Arachnida" family="Alycidae" genus="Minimamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D4615FFC743350E94FA68FE458166" bold="true" box="[238,453,1518,1545]" italics="true" pageId="43" pageNumber="44">Minimamichaelia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFC743350FB7FA69FDBB8166" box="[461,571,1519,1545]" pageId="43" pageNumber="44" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC743350EEDF9B2FBB58124" box="[151,1077,1587,1612]" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BD69A07FFC743350EEDF9B2FBB58124" blockId="43.[151,1077,1587,1612]" box="[151,1077,1587,1612]" pageId="43" pageNumber="44">
<materialsCitation id="3B01905AFFC743350EEDF9B2FBB18124" box="[151,1073,1587,1612]" country="Italy" location="Florence" municipality="Boboli" pageId="43" pageNumber="44" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D224A5FFC743350EEDF9B2FF498123" box="[151,201,1588,1612]" pageId="43" pageNumber="44">Type</typeStatus>
species:
<taxonomicName id="4C69E184FFC743350F5CF9B5FD308124" authority="Berlese, 1910: 201" authorityName="Berlese" authorityPageNumber="201" authorityYear="1910" box="[294,688,1587,1611]" class="Arachnida" family="Freyanidae" genus="Michaelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="subnuda">
<emphasis id="B91D4615FFC743350F5CF9B5FE698124" box="[294,489,1587,1611]" italics="true" pageId="43" pageNumber="44">Michaelia subnuda</emphasis>
<bibRefCitation id="EFF8E7F6FFC743350F8AF9B5FD308124" author="Berlese, A." box="[496,688,1587,1611]" pageId="43" pageNumber="44" pagination="199 - 234" refId="ref46521" refString="Berlese, A. (1910) Acari nuovi, Manipulus V, VI. Redia, 6, 199-234, pls. 18 - 21." type="journal article" year="1910">Berlese, 1910: 201</bibRefCitation>
</taxonomicName>
, fig. 20,
<collectingCountry id="F37EDA97FFC743350D70F9B5FCB98124" box="[778,825,1587,1611]" name="Italy" pageId="43" pageNumber="44">Italy</collectingCountry>
,
<location id="8EB6CCDCFFC743350D39F9B5FC1E8124" LSID="urn:lsid:plazi:treatment:03C02B11FFC743300EEDFA68FDAF84A6:8EB6CCDCFFC743350D39F9B5FC1E8124" box="[835,926,1587,1611]" country="Italy" municipality="Boboli" name="Florence" pageId="43" pageNumber="44">Florence</location>
,
<collectingMunicipality id="6BB2007DFFC743350DD2F9B5FC708124" box="[936,1008,1587,1611]" pageId="43" pageNumber="44">Boboli</collectingMunicipality>
, moss
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC743300EEDF9FFFB1B86E6" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8BD69A07FFC743350EEDF9FFFAE4804C" blockId="43.[151,1437,1657,1972]" pageId="43" pageNumber="44">
<emphasis id="B91D4615FFC743350EEDF9FFFE2081FC" bold="true" box="[151,416,1657,1683]" pageId="43" pageNumber="44">Differential diagnosis.</emphasis>
This genus can be separated from all the other bimichaeliine species by the absence of a primary pattern, i.e. the reticular pattern of roundish to polygonal loops formed by large lamellae is absent; granules and transversely inserted small to medium-sized lamellae are in undulant rows so that the middle-sized lamellae are clumped in culmination points (
<figureCitation id="13528682FFC743350F81F963FDE1806F" box="[507,609,1765,1792]" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="43" pageNumber="44">Fig. 51C</figureCitation>
). Other diagnostic features: tiny species; prodorsum holotrichous; dorsum sparsely neotrichous; setae sparsely ciliated; two baculiform solenidia on tarsi I (
<figureCitation id="13528682FFC743350A5AF88FFB32804B" box="[1056,1202,1801,1828]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="43" pageNumber="44">Figs. 52EH</figureCitation>
,
<figureCitation id="13528682FFC743350AC7F88FFB5B804B" box="[1213,1243,1801,1828]" captionStart="FIGURE 55" captionStartId="87.[151,250,1904,1928]" captionTargetBox="[151,1436,176,1881]" captionTargetId="figure-15@87.[151,1436,176,1881]" captionTargetPageId="87" captionText="FIGURE 55. Selected figures of undescribed species ber from South Africa. A—prodorsum; B—leg I; C—leg II; D—tibia and genu III; E—tibia and genus IV." pageId="43" pageNumber="44">55</figureCitation>
,
<emphasis id="B91D4615FFC743350A9DF88FFAD4804C" bold="true" box="[1255,1364,1801,1827]" pageId="43" pageNumber="44">Remarks</emphasis>
).
</paragraph>
<paragraph id="8BD69A07FFC743350EBDF8ABFCD680DB" blockId="43.[151,1437,1657,1972]" pageId="43" pageNumber="44">
The primary pattern of
<taxonomicName id="4C69E184FFC743350FB4F8ABFD128028" authority="Thor" authorityName="Thor" authorityYear="1902" box="[462,658,1837,1863]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC743350FB4F8ABFDD58028" box="[462,597,1837,1863]" italics="true" pageId="43" pageNumber="44">Bimichaelia</emphasis>
Thor
</taxonomicName>
forms roundish to subpolygonal loops of large lamellae; and the tiny ridges inside loops are granular; species are small to middle-sized (
<figureCitation id="13528682FFC743350DF9F8D7FC6B8003" box="[899,1003,1873,1900]" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="43" pageNumber="44">Fig. 51A</figureCitation>
); prodorsum holotrichous; dorsal setae have less than ten, mainly short barbs basally, which are difficult to discern against the intricate cuticular pattern; and usually three baculiform solenidia on tarsi I (
<figureCitation id="13528682FFC743350CC4F81FFCCA80DB" box="[702,842,1945,1972]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="43" pageNumber="44">Fig. 52AD</figureCitation>
).
</paragraph>
<caption id="DF16CA8FFFC043320EEDF8EFFEBF8085" pageId="44" pageNumber="45" startId="44.[151,250,1897,1921]" targetBox="[218,1370,153,1874]" targetPageId="44" targetType="figure">
<paragraph id="8BD69A07FFC043320EEDF8EFFDBE80EE" blockId="44.[151,1437,1897,2026]" box="[151,574,1897,1921]" pageId="44" pageNumber="45">
<emphasis id="B91D4615FFC043320EEDF8EFFEA080EE" bold="true" box="[151,288,1897,1921]" pageId="44" pageNumber="45">FIGURE 25.</emphasis>
<taxonomicName id="4C69E184FFC043320F5CF8EFFE6880EE" authorityName="Uusitalo" authorityYear="2025" box="[294,488,1897,1921]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="logani" status="sp. nov.">
<emphasis id="B91D4615FFC043320F5CF8EFFE6880EE" box="[294,488,1897,1921]" italics="true" pageId="44" pageNumber="45">Bimichaelia logani</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFC043320F94F8EFFDBE80EE" bold="true" box="[494,574,1897,1921]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="A22EFB6EFFC043320F94F8EFFDBE80EE" box="[494,574,1897,1921]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFC043320EEDF80DFEBF8085" blockId="44.[151,1437,1897,2026]" pageId="44" pageNumber="45">A—prodorsum, dorsal setae mostly with 56 clearly visible cilia; B—dorsum with ca. 300 setae; C—subcapitulum with 6 setae; D—genital valves of female with 14 genital setae per valve; E—anal area with 56 anal setae per valve. All specimens—from Utah, Cache Co.</paragraph>
</caption>
<caption id="DF16CA8FFFC143330EEDF953FB46807D" pageId="45" pageNumber="46" startId="45.[151,250,1749,1773]" targetBox="[152,1436,176,1726]" targetPageId="45" targetType="figure">
<paragraph id="8BD69A07FFC143330EEDF953FDBE8181" blockId="45.[151,1222,1749,1810]" box="[151,574,1749,1774]" pageId="45" pageNumber="46">
<emphasis id="B91D4615FFC143330EEDF953FEA08182" bold="true" box="[151,288,1749,1773]" pageId="45" pageNumber="46">FIGURE 26.</emphasis>
<taxonomicName id="4C69E184FFC143330F5CF953FE688182" authorityName="Uusitalo" authorityYear="2025" box="[294,488,1749,1773]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="logani" status="sp. nov.">
<emphasis id="B91D4615FFC143330F5CF953FE688182" box="[294,488,1749,1773]" italics="true" pageId="45" pageNumber="46">Bimichaelia logani</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFC143330F94F950FDBE8181" bold="true" box="[494,574,1750,1774]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="A22EFB6EFFC143330F94F950FDBE8181" box="[494,574,1750,1774]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFC143330EEDF97FFB46807D" blockId="45.[151,1222,1749,1810]" box="[151,1222,1785,1810]" pageId="45" pageNumber="46">A—leg I; B—leg II; C—tibia and genu III; D— tibia and genu IV. All specimens—from Utah, Cache Co.</paragraph>
</caption>
<paragraph id="8BD69A07FFC143330EBDF8C6FC6A8085" blockId="45.[151,1437,1856,2026]" pageId="45" pageNumber="46">
Members of
<taxonomicName id="4C69E184FFC143330F2FF8C6FD9F8035" authorityName="Uusitalo" authorityYear="2025" box="[341,543,1856,1882]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D4615FFC143330F2FF8C6FD9F8035" box="[341,543,1856,1882]" italics="true" pageId="45" pageNumber="46">Quartusmichaelia</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFC143330C5FF8C6FD0D8035" bold="true" box="[549,653,1856,1882]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="A22EFB6EFFC143330C5FF8C6FD0D8035" box="[549,653,1856,1882]" pageId="45" pageNumber="46" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
have a reticulate pattern of polygonally arranged large lamellae; small lamellae are transversely on ridges inside loops (
<figureCitation id="13528682FFC143330CC7F8E2FCA68011" box="[701,806,1891,1918]" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="45" pageNumber="46">Fig. 51D</figureCitation>
); species are small to middle-sized; ciliated dorsal setae are easily discernible because of several elongated cilia per seta at least on the shoulders and middorsum, but lanceolate setae are common; extra setae, if any, only on soft prodorsum; only one pair of setae
<emphasis id="B91D4615FFC143330AAFF82AFB6B80AA" box="[1237,1259,1964,1989]" italics="true" pageId="45" pageNumber="46">in</emphasis>
; and one of the two baculiform solenidia on tarsus I is hook-like or S-shaped (
<figureCitation id="13528682FFC143330D28F856FC5D8085" box="[850,989,1999,2026]" captionStart="FIGURE 53" captionStartId="85.[151,250,1681,1705]" captionTargetBox="[190,1397,153,1657]" captionTargetId="figure-15@85.[190,1397,153,1657]" captionTargetPageId="85" captionText="FIGURE 53. Genus-level character states of Bimichaeliini, part 2. AD—Quartusmichaelia shibai: ciliated setae on dorsum; prodorsum with six or seven pairs of setae; filamentous sensilla ve densely ciliated by cilia of various lengths from top to bottom; reticulate dorsal pattern; anterior solenidion ω1 hook-like on tarsus I, respectively; EH—Quartusmichaelia: dorsum with substellate polygons of primary pattern; setae lanceolate or ciliate; prodorsum with six pairs of setae or neotrichous rim of soft integument; lanceolate setae scaly or with tiny barbs; secondary pattern of even rows of small lamellae; tarsus I with two baculiform solenidia, anterior one being S-shaped or hook-like (and beak-like chelicera - typical of Bimichaeliini), respectively; IL—Laminamichaelia: dorsum with regular stellate pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with multiciliate setae; primary pattern polygonal; secondary pattern of even rows of small lamellae; three baculiform solenidia on tarsus I, respectively. 53AD—specimens from Veracruz, CICOLMA; 53EL—copied from Uusitalo (2010)." pageId="45" pageNumber="46">Fig. 53AH</figureCitation>
).
</paragraph>
<paragraph id="8BD69A07FFC243300EBDFF11FC018672" blockId="46.[151,1437,151,969]" pageId="46" pageNumber="47">
<taxonomicName id="4C69E184FFC243300EBDFF11FE0B87DE" authorityName="Uusitalo" authorityYear="2010" box="[199,395,151,177]" class="Arachnida" family="Alycidae" genus="Laminamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC243300EBDFF11FE0B87DE" box="[199,395,151,177]" italics="true" pageId="46" pageNumber="47">Laminamichaelia</emphasis>
</taxonomicName>
sensu str. have a reticulate pattern of polygonally arranged large lamellae; small lamellae are transversely on ridges inside loops (
<figureCitation id="13528682FFC243300C57FF3DFD1487BA" box="[557,660,186,213]" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="46" pageNumber="47">Fig. 51E</figureCitation>
); species are medium-sized to large; dorsal setae are often bush-like, easily discernible because of several elongated cilia per seta; both
<emphasis id="B91D4615FFC243300DF8FF59FC188797" box="[898,920,223,248]" italics="true" pageId="46" pageNumber="47">in</emphasis>
-area and soft prodorsum are neotrichous; and there are usually three baculiform solenidia on tarsi I (
<figureCitation id="13528682FFC243300C8DFE85FCF58672" box="[759,885,258,285]" captionStart="FIGURE 53" captionStartId="85.[151,250,1681,1705]" captionTargetBox="[190,1397,153,1657]" captionTargetId="figure-15@85.[190,1397,153,1657]" captionTargetPageId="85" captionText="FIGURE 53. Genus-level character states of Bimichaeliini, part 2. AD—Quartusmichaelia shibai: ciliated setae on dorsum; prodorsum with six or seven pairs of setae; filamentous sensilla ve densely ciliated by cilia of various lengths from top to bottom; reticulate dorsal pattern; anterior solenidion ω1 hook-like on tarsus I, respectively; EH—Quartusmichaelia: dorsum with substellate polygons of primary pattern; setae lanceolate or ciliate; prodorsum with six pairs of setae or neotrichous rim of soft integument; lanceolate setae scaly or with tiny barbs; secondary pattern of even rows of small lamellae; tarsus I with two baculiform solenidia, anterior one being S-shaped or hook-like (and beak-like chelicera - typical of Bimichaeliini), respectively; IL—Laminamichaelia: dorsum with regular stellate pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with multiciliate setae; primary pattern polygonal; secondary pattern of even rows of small lamellae; three baculiform solenidia on tarsus I, respectively. 53AD—specimens from Veracruz, CICOLMA; 53EL—copied from Uusitalo (2010)." pageId="46" pageNumber="47">Fig. 53IL</figureCitation>
).
</paragraph>
<paragraph id="8BD69A07FFC243300EBDFEA1FB1B86E6" blockId="46.[151,1437,151,969]" pageId="46" pageNumber="47">
<taxonomicName id="4C69E184FFC243300EBDFEA1FE05862E" authorityName="Uusitalo" authorityYear="2025" box="[199,389,295,321]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D4615FFC243300EBDFEA1FE05862E" box="[199,389,295,321]" italics="true" pageId="46" pageNumber="47">Glabromichaelia</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFC243300FF5FEA1FE7B862E" bold="true" box="[399,507,295,321]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A22EFB6EFFC243300FF5FEA1FE7B862E" box="[399,507,295,321]" pageId="46" pageNumber="47" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
have integumental ridges that are parallel and without granulae or laminae (
<figureCitation id="13528682FFC243300B15FEA1FF49860A" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="46" pageNumber="47">Fig. 51B</figureCitation>
); its single species is large in size; dorsum neotrichous with elongated and ciliated setae; both soft integument and
<emphasis id="B91D4615FFC243300EBDFEE9FF5D86E7" box="[199,221,367,392]" italics="true" pageId="46" pageNumber="47">in</emphasis>
-area on prodorsum neotrichous; and two baculiform solenidia on tarsi I (
<figureCitation id="13528682FFC243300A6BFEE9FB0E86E6" box="[1041,1166,366,393]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="46" pageNumber="47">Fig. 52IL</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC243300EBDFE15FA878432" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8BD69A07FFC243300EBDFE15FE23850E" blockId="46.[151,1437,151,969]" pageId="46" pageNumber="47">
<emphasis id="B91D4615FFC243300EBDFE15FEBB86C2" bold="true" box="[199,315,403,429]" pageId="46" pageNumber="47">Remarks.</emphasis>
The biggest lamellae in the lamellar clumps of
<taxonomicName id="4C69E184FFC243300D29FE15FC3286C2" authorityName="Berlese" authorityYear="1910" box="[851,946,403,429]" class="Arachnida" family="Freyanidae" genus="Michaelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="subnuda">
<emphasis id="B91D4615FFC243300D29FE15FC3286C2" box="[851,946,403,429]" italics="true" pageId="46" pageNumber="47">subnuda</emphasis>
</taxonomicName>
-group of
<emphasis id="B91D4615FFC243300A58FE15FADF86C2" box="[1058,1375,403,429]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="4C69E184FFC243300A58FE15FB6686C2" authorityName="Uusitalo" authorityYear="2010" box="[1058,1254,403,429]" class="Arachnida" family="Alycidae" genus="Laminamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus" sensu="lato">Laminamichaelia</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFC243300A94FE15FADF86C2" box="[1262,1375,403,429]" pageId="46" pageNumber="47" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
were mistaken as similar in size as the large lamellae in the lamellar cells of
<taxonomicName id="4C69E184FFC243300D9AFE31FBBA86BF" authorityName="Uusitalo" authorityYear="2025" baseAuthorityName="Berlese" baseAuthorityYear="1904" box="[992,1082,439,464]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setigera">
<emphasis id="B91D4615FFC243300D9AFE31FBBA86BF" box="[992,1082,439,464]" italics="true" pageId="46" pageNumber="47">setigera</emphasis>
</taxonomicName>
-group,
<taxonomicName id="4C69E184FFC243300AECFE31FA9F86BE" baseAuthorityName="Grandjean" baseAuthorityYear="1943" box="[1174,1311,439,465]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="arbusculosa">
<emphasis id="B91D4615FFC243300AECFE31FA9F86BE" box="[1174,1311,439,465]" italics="true" pageId="46" pageNumber="47">arbusculosa</emphasis>
</taxonomicName>
-group and
<taxonomicName id="4C69E184FFC243300EEDFE5DFE9E869A" authorityName="Thor" authorityYear="1902" box="[151,286,475,501]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC243300EEDFE5DFE9E869A" box="[151,286,475,501]" italics="true" pageId="46" pageNumber="47">Bimichaelia</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFC243300F54FE5DFE15869A" box="[302,405,474,501]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="46" pageNumber="47">Fig. 52G</figureCitation>
) in
<bibRefCitation id="EFF8E7F6FFC243300FBBFE5DFDF4869A" author="Uusitalo, M." box="[449,628,475,501]" pageId="46" pageNumber="47" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo (2010)</bibRefCitation>
, but in the context with the figures of reticulate integuments, the size of the lamellae rather corresponds with the middle-sized lamellae existing sporadically inside loops of large lamellae in
<taxonomicName id="4C69E184FFC243300EC9FDA5FEBA8552" authorityName="Thor" authorityYear="1902" box="[179,314,547,573]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC243300EC9FDA5FEBA8552" box="[179,314,547,573]" italics="true" pageId="46" pageNumber="47">Bimichaelia</emphasis>
</taxonomicName>
and the species groups of
<emphasis id="B91D4615FFC243300C24FDA5FC188552" box="[606,920,547,573]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="4C69E184FFC243300C24FDA5FCA28552" authorityName="Uusitalo" authorityYear="2010" box="[606,802,547,573]" class="Arachnida" family="Alycidae" genus="Laminamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus" sensu="lato">Laminamichaelia</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFC243300D52FDA5FC188552" box="[808,920,547,573]" pageId="46" pageNumber="47" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13528682FFC243300DDCFDA5FBB78552" box="[934,1079,546,573]" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="46" pageNumber="47">Figs. 51AE</figureCitation>
,
<figureCitation id="13528682FFC243300A3BFDA4FB358552" box="[1089,1205,546,573]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="46" pageNumber="47">52C, G, K</figureCitation>
;
<bibRefCitation id="EFF8E7F6FFC243300ABAFDA5FAE18552" author="Uusitalo, M." box="[1216,1377,547,573]" pageId="46" pageNumber="47" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: figs. 95, 105, 115, 125, 135).
</paragraph>
<paragraph id="8BD69A07FFC243300EBDFDEDFA878432" blockId="46.[151,1437,151,969]" pageId="46" pageNumber="47">
A species of the
<taxonomicName id="4C69E184FFC243300FF9FDEDFE6285EA" authorityName="Berlese" authorityYear="1910" box="[387,482,619,645]" class="Arachnida" family="Freyanidae" genus="Michaelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="subnuda">
<emphasis id="B91D4615FFC243300FF9FDEDFE6285EA" box="[387,482,619,645]" italics="true" pageId="46" pageNumber="47">subnuda</emphasis>
</taxonomicName>
-group (now
<taxonomicName id="4C69E184FFC243300C0FFDEDFCBB85EA" authorityName="Uusitalo" authorityYear="2025" box="[629,827,619,645]" class="Arachnida" family="Alycidae" genus="Minimamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC243300C0FFDEDFCBB85EA" box="[629,827,619,645]" italics="true" pageId="46" pageNumber="47">Minimamichaelia</emphasis>
</taxonomicName>
) from
<collectingCountry id="F37EDA97FFC243300DF0FDEDFB9985EA" box="[906,1049,619,645]" name="South Africa" pageId="46" pageNumber="47">South Africa</collectingCountry>
(
<figureCitation id="13528682FFC243300A52FDEDFBFA85EA" box="[1064,1146,618,645]" captionStart="FIGURE 55" captionStartId="87.[151,250,1904,1928]" captionTargetBox="[151,1436,176,1881]" captionTargetId="figure-15@87.[151,1436,176,1881]" captionTargetPageId="87" captionText="FIGURE 55. Selected figures of undescribed species ber from South Africa. A—prodorsum; B—leg I; C—leg II; D—tibia and genu III; E—tibia and genus IV." pageId="46" pageNumber="47">Fig. 55</figureCitation>
) is in the late Prof. Pieter Therons Collection and has been temporarily lost in removal of the collection, and the description is unpublished so far (Dr. Edward A. Ueckermann, in litt.). This clade was obtained from the analysis of the family
<taxonomicName id="4C69E184FFC243300B7EFD35FAE985A2" baseAuthorityName="Uusitalo" baseAuthorityYear="2010" box="[1284,1385,691,717]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="family">Alycidae</taxonomicName>
as a part of the terminals, which at that time where monotypic, and called by names
<taxonomicName id="4C69E184FFC243300A7CFD51FBE5859E" authorityName="Berlese" authorityYear="1910" box="[1030,1125,727,753]" class="Arachnida" family="Freyanidae" genus="Michaelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="subnuda">
<emphasis id="B91D4615FFC243300A7CFD51FBE5859E" box="[1030,1125,727,753]" italics="true" pageId="46" pageNumber="47">subnuda</emphasis>
</taxonomicName>
-group, along with
<taxonomicName id="4C69E184FFC243300B43FD51FA13859F" authorityName="Uusitalo" authorityYear="2025" baseAuthorityName="Berlese" baseAuthorityYear="1904" box="[1337,1427,727,752]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setigera">
<emphasis id="B91D4615FFC243300B43FD51FA13859F" box="[1337,1427,727,752]" italics="true" pageId="46" pageNumber="47">setigera</emphasis>
</taxonomicName>
- group, and
<taxonomicName id="4C69E184FFC243300F60FD7DFE23847A" baseAuthorityName="Grandjean" baseAuthorityYear="1943" box="[282,419,763,789]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="arbusculosa">
<emphasis id="B91D4615FFC243300F60FD7DFE23847A" box="[282,419,763,789]" italics="true" pageId="46" pageNumber="47">arbusculosa</emphasis>
</taxonomicName>
-group under
<emphasis id="B91D4615FFC243300C3AFD7DFC01847A" box="[576,897,763,789]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="4C69E184FFC243300C3AFD7DFC84847A" authorityName="Uusitalo" authorityYear="2010" box="[576,772,763,789]" class="Arachnida" family="Alycidae" genus="Laminamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus" sensu="lato">Laminamichaelia</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFC243300D77FD7DFC01847A" box="[781,897,763,789]" pageId="46" pageNumber="47" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="EFF8E7F6FFC243300DEBFD7DFBB8847A" author="Uusitalo, M." box="[913,1080,763,789]" pageId="46" pageNumber="47" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
). After detailed examination of several species outside Europe it became clear that certain character states are unique to the groups, the groups are no longer monotypic, and the characters are now used in creating generic diagnoses for each of them.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC243300EBDFCE1FDAF84A6" pageId="46" pageNumber="47" type="etymology">
<paragraph id="8BD69A07FFC243300EBDFCE1FDAF84A6" blockId="46.[151,1437,151,969]" pageId="46" pageNumber="47">
<emphasis id="B91D4615FFC243300EBDFCE1FECE84EE" bold="true" box="[199,334,871,897]" pageId="46" pageNumber="47">Etymology.</emphasis>
The name of the genus is a latinised combination of the Latin word ”minima”, meaning tiny, and “Michael”, referring to the British acarologist Albert Davidson Michael (18361927). The name refers to the minute size of the species of the new genus.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,194 @@
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<subSubSection id="C373C98CFFC743350EEDFD34FE6385A2" box="[151,483,690,717]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8BD69A07FFC743350EEDFD34FE6385A2" blockId="43.[151,483,690,753]" box="[151,483,690,717]" pageId="43" pageNumber="44">
<heading id="D09E2D6BFFC743350EEDFD34FE6385A2" bold="true" box="[151,483,690,717]" fontSize="11" level="2" pageId="43" pageNumber="44" reason="4">
<emphasis id="B91D4615FFC743350EEDFD34FE6385A2" bold="true" box="[151,483,690,717]" pageId="43" pageNumber="44">
<taxonomicName id="4C69E184FFC743350EEDFD34FEFE85A2" authorityName="Uusitalo" authorityYear="2025" box="[151,382,690,717]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="logani" status="sp. nov.">
<emphasis id="B91D4615FFC743350EEDFD34FEFE85A2" bold="true" box="[151,382,690,717]" italics="true" pageId="43" pageNumber="44">Bimichaelia logani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFC743350FFFFD35FE6385A2" box="[389,483,691,717]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BD69A07FFC743350EEDFD51FE18859E" blockId="43.[151,483,690,753]" box="[151,408,726,753]" pageId="43" pageNumber="44">
(
<figureCitation id="13528682FFC743350EE5FD51FEB1859E" box="[159,305,726,753]" captionStart="FIGURE 25" captionStartId="44.[151,250,1897,1921]" captionTargetBox="[218,1370,153,1874]" captionTargetId="figure-24@44.[218,1370,153,1874]" captionTargetPageId="44" captionText="FIGURE 25. Bimichaelia logani sp. nov. A—prodorsum, dorsal setae mostly with 56 clearly visible cilia; B—dorsum with ca. 300 setae; C—subcapitulum with 6 setae; D—genital valves of female with 14 genital setae per valve; E—anal area with 56 anal setae per valve. All specimens—from Utah, Cache Co." pageId="43" pageNumber="44">Figs. 25AE</figureCitation>
,
<figureCitation id="13528682FFC743350F46FD51FE10859E" box="[316,400,727,753]" captionStart="FIGURE 26" captionStartId="45.[151,250,1749,1773]" captionTargetBox="[152,1436,176,1726]" captionTargetId="figure-15@45.[151,1436,176,1726]" captionTargetPageId="45" captionText="FIGURE 26. Bimichaelia logani sp. nov. A—leg I; B—leg II; C—tibia and genu III; D— tibia and genu IV. All specimens—from Utah, Cache Co." pageId="43" pageNumber="44">26AD</figureCitation>
)
</paragraph>
<paragraph id="8BD69A07FFC743350EEDFC98FAB484CA" blockId="43.[151,1437,798,1437]" pageId="43" pageNumber="44">
<emphasis id="B91D4615FFC743350EEDFC98FEA98457" bold="true" box="[151,297,798,824]" pageId="43" pageNumber="44">Description.</emphasis>
<emphasis id="B91D4615FFC743350F4BFC99FE0B8457" box="[305,395,799,824]" italics="true" pageId="43" pageNumber="44">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFC743350FC1FC99FDAC8456" box="[443,556,799,825]" count="2" pageId="43" pageNumber="44" type="female">2 females</specimenCount>
,
<figureCitation id="13528682FFC743350C43FC99FD438456" box="[569,707,798,825]" captionStart="FIGURE 25" captionStartId="44.[151,250,1897,1921]" captionTargetBox="[218,1370,153,1874]" captionTargetId="figure-24@44.[218,1370,153,1874]" captionTargetPageId="44" captionText="FIGURE 25. Bimichaelia logani sp. nov. A—prodorsum, dorsal setae mostly with 56 clearly visible cilia; B—dorsum with ca. 300 setae; C—subcapitulum with 6 setae; D—genital valves of female with 14 genital setae per valve; E—anal area with 56 anal setae per valve. All specimens—from Utah, Cache Co." pageId="43" pageNumber="44">Fig. 25A, B</figureCitation>
). Length ca. 550 µm; dorsal setae with 56 clear cilia each and surrounded by 56 roundish loops of reticular primary pattern, caudal setae with elongated main cilium; prodorsum clearly reticulated, cilia of setae
<emphasis id="B91D4615FFC743350C7AFCE1FD9684EF" box="[512,534,871,896]" italics="true" pageId="43" pageNumber="44">in</emphasis>
,
<emphasis id="B91D4615FFC743350C59FCE1FDC784EF" box="[547,583,871,896]" italics="true" pageId="43" pageNumber="44">sce</emphasis>
and
<emphasis id="B91D4615FFC743350C06FCE1FD1084EF" box="[636,656,871,896]" italics="true" pageId="43" pageNumber="44">vi</emphasis>
along an elongated centre, anterior pair of sensilla
<emphasis id="B91D4615FFC743350AB2FCE1FB6184EF" box="[1224,1249,871,896]" italics="true" pageId="43" pageNumber="44">ve</emphasis>
densely ciliated, setae
<emphasis id="B91D4615FFC743350EACFC0DFF7D84CB" box="[214,253,907,932]" italics="true" pageId="43" pageNumber="44">exp</emphasis>
with long main cilium (like caudal setae), hard integument with large lamellae in rows or loops.
</paragraph>
<paragraph id="8BD69A07FFC743350EBDFC29FB2684A6" blockId="43.[151,1437,798,1437]" box="[199,1190,942,969]" pageId="43" pageNumber="44">
<taxonomicName id="4C69E184FFC743350EBDFC29FE8F84A7" box="[199,271,943,968]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="43" pageNumber="44" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFC743350EBDFC29FE8F84A7" box="[199,271,943,968]" italics="true" pageId="43" pageNumber="44">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFC743350F64FC29FE2684A6" box="[286,422,942,969]" captionStart="FIGURE 25" captionStartId="44.[151,250,1897,1921]" captionTargetBox="[218,1370,153,1874]" captionTargetId="figure-24@44.[218,1370,153,1874]" captionTargetPageId="44" captionText="FIGURE 25. Bimichaelia logani sp. nov. A—prodorsum, dorsal setae mostly with 56 clearly visible cilia; B—dorsum with ca. 300 setae; C—subcapitulum with 6 setae; D—genital valves of female with 14 genital setae per valve; E—anal area with 56 anal setae per valve. All specimens—from Utah, Cache Co." pageId="43" pageNumber="44">Fig. 25D, E</figureCitation>
). Genital valves each with 14 genital setae; 56 anal setae per valve.
</paragraph>
<paragraph id="8BD69A07FFC743350EBDFC55FE05837E" blockId="43.[151,1437,798,1437]" pageId="43" pageNumber="44">
<taxonomicName id="4C69E184FFC743350EBDFC55FED68482" authorityName="Krefft" authorityYear="1871" box="[199,342,979,1005]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="43" pageNumber="44" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFC743350EBDFC55FED68482" box="[199,342,979,1005]" italics="true" pageId="43" pageNumber="44">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFC743350F1FFC55FE4B8482" box="[357,459,978,1005]" captionStart="FIGURE 25" captionStartId="44.[151,250,1897,1921]" captionTargetBox="[218,1370,153,1874]" captionTargetId="figure-24@44.[218,1370,153,1874]" captionTargetPageId="44" captionText="FIGURE 25. Bimichaelia logani sp. nov. A—prodorsum, dorsal setae mostly with 56 clearly visible cilia; B—dorsum with ca. 300 setae; C—subcapitulum with 6 setae; D—genital valves of female with 14 genital setae per valve; E—anal area with 56 anal setae per valve. All specimens—from Utah, Cache Co." pageId="43" pageNumber="44">Fig. 25C</figureCitation>
). Apical palpal solenidion supported by one eupathidion; palpal joints complete; 3 pairs of subcapitular setae.
</paragraph>
<paragraph id="8BD69A07FFC743350EBDFB9DFBE18336" blockId="43.[151,1437,798,1437]" pageId="43" pageNumber="44">
<emphasis id="B91D4615FFC743350EBDFB9DFF7C835B" box="[199,252,1051,1076]" italics="true" pageId="43" pageNumber="44">Legs</emphasis>
(
<figureCitation id="13528682FFC743350F71FB9DFE17835A" box="[267,407,1051,1077]" captionStart="FIGURE 26" captionStartId="45.[151,250,1749,1773]" captionTargetBox="[152,1436,176,1726]" captionTargetId="figure-15@45.[151,1436,176,1726]" captionTargetPageId="45" captionText="FIGURE 26. Bimichaelia logani sp. nov. A—leg I; B—leg II; C—tibia and genu III; D— tibia and genu IV. All specimens—from Utah, Cache Co." pageId="43" pageNumber="44">Fig. 26AD</figureCitation>
). Solenidial formula for tarsi, tibiae, genua and femora on legs I, II, III and IV, respectively: 3B-2B-0-0, 1B-1B-1B-1B, 1C-1C-1C-1C, 1C-1B-0-0; famulus I laterodistal abaxially.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC743350EBDFBE5FD5983AA" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8BD69A07FFC743350EBDFBE5FD5983AA" blockId="43.[151,1437,798,1437]" pageId="43" pageNumber="44">
<emphasis id="B91D4615FFC743350EBDFBE5FEF48313" bold="true" box="[199,372,1122,1149]" pageId="43" pageNumber="44">Type material.</emphasis>
<materialsCitation id="3B01905AFFC743350F00FBE5FD5983AA" collectingDate="1972-04-05" collectorName="G. F. Knowlton. Deposited &amp; Acarology Laboratory &amp; University" country="United States of America" county="Cache Co." location="Maple" municipality="Green Canyon" pageId="43" pageNumber="44" specimenCount="1" specimenCount-female="1" stateProvince="Utah" typeStatus="holotype">
<typeStatus id="54D224A5FFC743350F00FBE5FE628312" box="[378,482,1123,1149]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
female and
<specimenCount id="9D6F518EFFC743350C10FBE5FCB08312" box="[618,816,1123,1149]" count="1" pageId="43" pageNumber="44" type="female" typeStatus="paratype">1 paratype female</specimenCount>
as
<taxonomicName id="4C69E184FFC743350D2EFBE5FC5B8312" authorityName="Thor" authorityYear="1902" box="[852,987,1123,1149]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC743350D2EFBE5FC5B8312" box="[852,987,1123,1149]" italics="true" pageId="43" pageNumber="44">Bimichaelia</emphasis>
</taxonomicName>
from
<location id="8EB6CCDCFFC743350A64FBE5FBE68312" LSID="urn:lsid:plazi:treatment:03C02B11FFC743350EEDFD34FA9582F2:8EB6CCDCFFC743350A64FBE5FBE68312" box="[1054,1126,1123,1149]" country="United States of America" county="Cache Co." municipality="Green Canyon" name="Maple" pageId="43" pageNumber="44" stateProvince="Utah">Maple</location>
duff,
<collectingMunicipality id="6BB2007DFFC743350ADDFBE5FACC8312" box="[1191,1356,1123,1149]" pageId="43" pageNumber="44">Green Canyon</collectingMunicipality>
,
<collectingCounty id="62B7E28BFFC743350B2CFBE5FF4183CE" pageId="43" pageNumber="44">Cache Co.</collectingCounty>
,
<collectingRegion id="49AD54E5FFC743350EABFB01FE8A83CE" box="[209,266,1159,1185]" country="United States of America" name="Utah" pageId="43" pageNumber="44">Utah</collectingRegion>
,
<collectingCountry id="F37EDA97FFC743350F63FB01FED483CE" box="[281,340,1159,1185]" name="United States of America" pageId="43" pageNumber="44">USA</collectingCountry>
,
<date id="FFD7BCC7FFC743350F19FB00FE7B83CE" box="[355,507,1158,1185]" pageId="43" pageNumber="44" value="1972-04-05">
<collectingDate id="EF93452FFFC743350F19FB00FE7B83CE" box="[355,507,1158,1185]" pageId="43" pageNumber="44" value="1972-04-05">5 April 1972</collectingDate>
</date>
,
<collectorName id="269CFFD1FFC743350C70FB01FCB983CE" box="[522,825,1159,1185]" pageId="43" pageNumber="44">G.F. Knowlton. Deposited</collectorName>
at the
<collectorName id="269CFFD1FFC743350DF4FB01FB0D83CE" box="[910,1165,1159,1185]" pageId="43" pageNumber="44">Acarology Laboratory</collectorName>
,
<collectingRegion id="49AD54E5FFC743350AE1FB01FA9683CE" box="[1179,1302,1159,1185]" country="United States of America" name="Ohio" pageId="43" pageNumber="44">Ohio State</collectingRegion>
<collectorName id="269CFFD1FFC743350B5BFB01FA1883CE" box="[1313,1432,1159,1185]" pageId="43" pageNumber="44">University</collectorName>
, collection number AL701,
<typeStatus id="54D224A5FFC743350FBFFB2DFDA783AA" box="[453,551,1195,1221]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
on slide NA32.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC743350EBDFB48FDC9823A" pageId="43" pageNumber="44" type="diagnosis">
<paragraph id="8BD69A07FFC743350EBDFB48FDC9823A" blockId="43.[151,1437,798,1437]" pageId="43" pageNumber="44">
<emphasis id="B91D4615FFC743350EBDFB48FE4E8387" bold="true" box="[199,462,1230,1256]" pageId="43" pageNumber="44">Differential diagnosis.</emphasis>
This species can be differentiated from other species of
<taxonomicName id="4C69E184FFC743350A3EFB49FB4B8386" authorityName="Thor" authorityYear="1902" box="[1092,1227,1231,1257]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC743350A3EFB49FB4B8386" box="[1092,1227,1231,1257]" italics="true" pageId="43" pageNumber="44">Bimichaelia</emphasis>
</taxonomicName>
by having sensilla
<emphasis id="B91D4615FFC743350EEDFB75FF308263" box="[151,176,1267,1292]" italics="true" pageId="43" pageNumber="44">ve</emphasis>
ciliated all along while other species have the sensilla
<emphasis id="B91D4615FFC743350D77FB75FCA68263" box="[781,806,1267,1292]" italics="true" pageId="43" pageNumber="44">ve</emphasis>
simple and only basally barbed, e.g.
<taxonomicName id="4C69E184FFC743350ABAFB75FF73825E" authority="(Ewing)" baseAuthorityName="Ewing" baseAuthorityYear="1913" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B91D4615FFC743350ABAFB75FA1C8262" box="[1216,1436,1267,1293]" italics="true" pageId="43" pageNumber="44">Bimichaelia pallida</emphasis>
(Ewing)
</taxonomicName>
from
<collectingRegion id="49AD54E5FFC743350F46FA91FE10825E" box="[316,400,1303,1329]" country="United States of America" name="Oregon" pageId="43" pageNumber="44">Oregon</collectingRegion>
(
<figureCitation id="13528682FFC743350FD8FA91FD8E825E" box="[418,526,1303,1329]" captionStart="FIGURE 23" captionStartId="41.[151,250,1863,1887]" captionTargetBox="[176,1412,153,1839]" captionTargetId="figure-15@41.[176,1412,153,1839]" captionTargetPageId="41" captionText="FIGURE 23. Bimichaelia pallida (Ewing). A—prodorsum, dorsal setae mostly with 3 clearly visible cilia; B—subcapitulum with 9 setae; C—dorsum with ca. 300 setae; D—original figure of Bimichaelia pallida (Ewing), dorsum with ca. 100 setae; E—genital valves of male with 1418 genital setae per valve; F—7 pairs of eugenital setae; G—anal area with 6 anal setae per valve. D—copied from Ewing (1913). 23AC, EG—from Oregon, Multinomah Co., Corbett; 23D—from Oregon, Benton Co., Corvallis, copied from Ewing (1913)." pageId="43" pageNumber="44">Fig. 23A</figureCitation>
).
<taxonomicName id="4C69E184FFC743350C59FA91FD7C825E" authorityName="Uusitalo" authorityYear="2025" box="[547,764,1303,1329]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="logani">
<emphasis id="B91D4615FFC743350C59FA91FD7C825E" box="[547,764,1303,1329]" italics="true" pageId="43" pageNumber="44">Bimichaelia logani</emphasis>
</taxonomicName>
also has more than 3 visible cilia on dorsal setae, and the single ceratiform solenidion on genu I.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC743350EBDFAD9FCD68216" box="[199,854,1375,1401]" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BD69A07FFC743350EBDFAD9FCD68216" blockId="43.[151,1437,798,1437]" box="[199,854,1375,1401]" pageId="43" pageNumber="44">
<emphasis id="B91D4615FFC743350EBDFAD9FEB08216" bold="true" box="[199,304,1375,1401]" pageId="43" pageNumber="44">Remark.</emphasis>
<taxonomicName id="4C69E184FFC743350F4DFAD9FE1E8216" authorityName="Uusitalo" authorityYear="2025" box="[311,414,1375,1401]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="logani">
<emphasis id="B91D4615FFC743350F4DFAD9FE1E8216" box="[311,414,1375,1401]" italics="true" pageId="43" pageNumber="44">B. logani</emphasis>
</taxonomicName>
was coded as
<emphasis id="B91D4615FFC743350C33FAD9FDF38216" box="[585,627,1375,1401]" italics="true" pageId="43" pageNumber="44">bim</emphasis>
in
<bibRefCitation id="EFF8E7F6FFC743350CE4FAD9FCD28216" author="Uusitalo, M." box="[670,850,1375,1401]" pageId="43" pageNumber="44" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo (2010)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC743350EBDFA05FA9582F2" box="[199,1301,1411,1437]" pageId="43" pageNumber="44" type="etymology">
<paragraph id="8BD69A07FFC743350EBDFA05FA9582F2" blockId="43.[151,1437,798,1437]" box="[199,1301,1411,1437]" pageId="43" pageNumber="44">
<emphasis id="B91D4615FFC743350EBDFA05FECE82F2" bold="true" box="[199,334,1411,1437]" pageId="43" pageNumber="44">Etymology.</emphasis>
The species name refers to the city of Logan,
<collectingRegion id="49AD54E5FFC743350D2EFA05FC0D82F2" box="[852,909,1411,1437]" country="United States of America" name="Utah" pageId="43" pageNumber="44">Utah</collectingRegion>
, which is close to the
<typeStatus id="54D224A5FFC743350AFEFA05FB3482F2" box="[1156,1204,1411,1437]" pageId="43" pageNumber="44">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,233 @@
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<paragraph id="8BD69A07FFCC433E0EEDF993FDE48140" blockId="32.[151,612,1557,1583]" box="[151,612,1557,1583]" pageId="32" pageNumber="33">
<heading id="D09E2D6BFFCC433E0EEDF993FDE48140" bold="true" box="[151,612,1557,1583]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="3">
<emphasis id="B91D4615FFCC433E0EEDF993FDE48140" bold="true" box="[151,612,1557,1583]" pageId="32" pageNumber="33">
Tribe
<taxonomicName id="4C69E184FFCC433E0E98F993FDE48140" authority="Womersley, 1944" authorityName="Womersley" authorityYear="1944" box="[226,612,1557,1583]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="tribe" tribe="Bimichaeliini">
Bimichaeliini
<bibRefCitation id="EFF8E7F6FFCC433E0FF5F993FDE48140" author="Womersley, H." box="[399,612,1557,1583]" pageId="32" pageNumber="33" pagination="133 - 143" refId="ref49733" refString="Womersley, H. (1944) Australian Acarina, Families Alycidae and Nanorchestidae. Transactions of the Royal Society of South Australia, 68, 133-143." type="journal article" year="1944">Womersley, 1944</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCC433E0EEDF9DCFD8F811E" box="[151,527,1625,1650]" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BD69A07FFCC433E0EEDF9DCFD8F811E" blockId="32.[151,527,1625,1650]" box="[151,527,1625,1650]" pageId="32" pageNumber="33">
<typeStatus id="54D224A5FFCC433E0EEDF9DCFF49811D" box="[151,201,1626,1650]" pageId="32" pageNumber="33">Type</typeStatus>
genus:
<taxonomicName id="4C69E184FFCC433E0F62F9DFFD8B811E" authority="Thor, 1902" authorityName="Thor" authorityYear="1902" box="[280,523,1625,1649]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCC433E0F62F9DFFE12811E" box="[280,402,1625,1649]" italics="true" pageId="32" pageNumber="33">Bimichaelia</emphasis>
<bibRefCitation id="EFF8E7F6FFCC433E0FE3F9DFFD8B811E" author="Thor, S." box="[409,523,1625,1649]" pageId="32" pageNumber="33" pagination="159 - 165" refId="ref48630" refString="Thor, S. (1902) Zur Systematik der Acaridenfauna Bdellidae Koch, 1842, Grube, 1859, Eupodidae Koch, 1842 und Cunaxidae Sig Thor, 1902. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft, Wien, 52, 159-165." type="journal article" year="1902">Thor, 1902</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCC433F0EEDF919FA05857B" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BD69A07FFCC433E0EEDF919FD6980FE" blockId="32.[151,1436,1695,1938]" pageId="32" pageNumber="33">
<emphasis id="B91D4615FFCC433E0EEDF919FE1D81D6" bold="true" box="[151,413,1695,1721]" pageId="32" pageNumber="33">Differential diagnosis.</emphasis>
The tribe differs from its monotypic sister group, the tribe
<taxonomicName id="4C69E184FFCC433E0A52F919FB7B81D6" box="[1064,1275,1695,1721]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="tribe" tribe="Proteromichaeliini">Proteromichaeliini</taxonomicName>
, by having the following features: either glabrous ridges (
<figureCitation id="13528682FFCC433E0C0CF945FD5D81B1" box="[630,733,1731,1758]" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="32" pageNumber="33">Fig. 51B</figureCitation>
), or ridges with middle-sized lamellae in clumps (
<figureCitation id="13528682FFCC433E0B6BF945FAF681B1" box="[1297,1398,1731,1758]" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="32" pageNumber="33">Fig. 51C</figureCitation>
) or large lamellae in separate polygons (
<figureCitation id="13528682FFCC433E0C56F961FD50806D" box="[556,720,1767,1794]" captionStart="FIGURE 51" captionStartId="83.[151,247,1733,1757]" captionTargetBox="[196,1391,153,1709]" captionTargetId="figure-15@83.[196,1391,153,1709]" captionTargetPageId="83" captionText="FIGURE 51. Genus-level character states of integument and dorsal setae of Bimichaeliini on the same page and to a similar scale. A—Bimichaelia: large lamellae in roundish cells; granulae and a few middle-sized lamellae on ridges inside cells; setae with short cilia and a few elongated cilia from basis; B—Glabromichaelia: parallel ridges glabrous; setal shafts with short cilia; C—Minimamichaelia: parallel ridges with granulae, small lamellae of various sizes and clumps of middle-sized lamellae; setal shafts with short cilia and a few elongated cilia; D—Quartusmichaelia: large lamellae in roundish to polygonal cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae lanceolate and/or sparsely ciliated; E—Laminamichaelia: large lamellae in mostly triangular cells; small and a few middle-sized lamellae transversely and densely packed on parallel ridges inside cells; setae with densely ciliated bunches. 51A, CE—copied from Uusitalo (2010); 51B—from Veracruz, CICOLMA." pageId="32" pageNumber="33">Fig. 51A, D, E</figureCitation>
) on soft integument (compared to the primary pattern formed by both clumps and rosettes of large lamellae [
<bibRefCitation id="EFF8E7F6FFCC433E0CFAF88DFCE68049" author="Uusitalo, M. &amp; Ueckermann, E. A. &amp; Theron, P. D." box="[640,870,1803,1830]" pageId="32" pageNumber="33" pagination="301 - 340" refId="ref48918" refString="Uusitalo, M., Ueckermann, E. A. &amp; Theron, P. D. (2020) A review of the family Alycidae (Acari, Acariformes) from South Africa. Zootaxa, 4858 (3), 301-340. https://doi.org/10.11646/zootaxa.4858.3.1" type="journal article" year="2020">
Uusitalo
<emphasis id="B91D4615FFCC433E0C90F88AFC9C804A" box="[746,796,1803,1829]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. 2020
</bibRefCitation>
: fig. 134A, B]); the absence of eyes (
<figureCitation id="13528682FFCC433E0B71F88DFF248025" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="32" pageNumber="33">Figs. 52B, F, J</figureCitation>
,
<figureCitation id="13528682FFCC433E0ECAF8A9FE938025" box="[176,275,1839,1866]" captionStart="FIGURE 53" captionStartId="85.[151,250,1681,1705]" captionTargetBox="[190,1397,153,1657]" captionTargetId="figure-15@85.[190,1397,153,1657]" captionTargetPageId="85" captionText="FIGURE 53. Genus-level character states of Bimichaeliini, part 2. AD—Quartusmichaelia shibai: ciliated setae on dorsum; prodorsum with six or seven pairs of setae; filamentous sensilla ve densely ciliated by cilia of various lengths from top to bottom; reticulate dorsal pattern; anterior solenidion ω1 hook-like on tarsus I, respectively; EH—Quartusmichaelia: dorsum with substellate polygons of primary pattern; setae lanceolate or ciliate; prodorsum with six pairs of setae or neotrichous rim of soft integument; lanceolate setae scaly or with tiny barbs; secondary pattern of even rows of small lamellae; tarsus I with two baculiform solenidia, anterior one being S-shaped or hook-like (and beak-like chelicera - typical of Bimichaeliini), respectively; IL—Laminamichaelia: dorsum with regular stellate pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with multiciliate setae; primary pattern polygonal; secondary pattern of even rows of small lamellae; three baculiform solenidia on tarsus I, respectively. 53AD—specimens from Veracruz, CICOLMA; 53EL—copied from Uusitalo (2010)." pageId="32" pageNumber="33">53B, F, J</figureCitation>
) and cheliceral setae (
<figureCitation id="13528682FFCC433E0C76F8A9FDF48025" box="[524,628,1839,1866]" captionStart="FIGURE 53" captionStartId="85.[151,250,1681,1705]" captionTargetBox="[190,1397,153,1657]" captionTargetId="figure-15@85.[190,1397,153,1657]" captionTargetPageId="85" captionText="FIGURE 53. Genus-level character states of Bimichaeliini, part 2. AD—Quartusmichaelia shibai: ciliated setae on dorsum; prodorsum with six or seven pairs of setae; filamentous sensilla ve densely ciliated by cilia of various lengths from top to bottom; reticulate dorsal pattern; anterior solenidion ω1 hook-like on tarsus I, respectively; EH—Quartusmichaelia: dorsum with substellate polygons of primary pattern; setae lanceolate or ciliate; prodorsum with six pairs of setae or neotrichous rim of soft integument; lanceolate setae scaly or with tiny barbs; secondary pattern of even rows of small lamellae; tarsus I with two baculiform solenidia, anterior one being S-shaped or hook-like (and beak-like chelicera - typical of Bimichaeliini), respectively; IL—Laminamichaelia: dorsum with regular stellate pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with multiciliate setae; primary pattern polygonal; secondary pattern of even rows of small lamellae; three baculiform solenidia on tarsus I, respectively. 53AD—specimens from Veracruz, CICOLMA; 53EL—copied from Uusitalo (2010)." pageId="32" pageNumber="33">Fig. 53H</figureCitation>
); and femora IV not divided (
<figureCitation id="13528682FFCC433E0DC4F8A9FBAF8025" box="[958,1071,1839,1866]" captionStart="FIGURE 28" captionStartId="48.[151,250,1956,1980]" captionTargetBox="[189,1398,193,1931]" captionTargetId="figure-24@48.[189,1398,193,1931]" captionTargetPageId="48" captionText="FIGURE 28. Minimamichaelia subnuda (Berlese). A—subcapitulum; B—leg I; C—leg II; D—leg III and leg IV. 2728 All specimens—from South Dakota, Campbell Co." pageId="32" pageNumber="33">Figs. 28D</figureCitation>
,
<figureCitation id="13528682FFCC433E0A41F8A9FBED8025" box="[1083,1133,1839,1866]" captionStart="FIGURE 35" captionStartId="58.[151,250,1926,1950]" captionTargetBox="[151,1435,232,1903]" captionTargetId="figure-36@58.[151,1435,232,1903]" captionTargetPageId="58" captionText="FIGURE 35. Quartusmichaelia shibai (Uusitalo, Ueckermann &amp; Theron). A—prodorsum with sensilla ve ciliated from top to bottom; B—dorsum; C—venter and gnathosoma; D—leg I; E—leg II. All specimens—from Veracruz, CICOLMA." pageId="32" pageNumber="33">35C</figureCitation>
). Otherwise, they share the typical bimichaeliine character states, such as presence of clavate sensilla
<emphasis id="B91D4615FFCC433E0D9AF8D2FC7F8002" box="[992,1023,1876,1901]" italics="true" pageId="32" pageNumber="33">sci</emphasis>
, stylet-like chelicerae, and an apical palpal solenidion in contact with TP-type eupathidia.
</paragraph>
<paragraph id="8BD69A07FFCD433F0EEDFF11FDAF87DE" blockId="33.[151,559,151,178]" box="[151,559,151,178]" pageId="33" pageNumber="34">
<emphasis id="B91D4615FFCD433F0EEDFF11FDAF87DE" bold="true" box="[151,559,151,178]" pageId="33" pageNumber="34">
Genus
<taxonomicName id="4C69E184FFCD433F0E94FF11FE3A87DD" authorityName="Uusitalo" authorityYear="2025" box="[238,442,151,178]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D4615FFCD433F0E94FF11FE3A87DD" bold="true" box="[238,442,151,178]" italics="true" pageId="33" pageNumber="34">Glabromichaelia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFCD433F0FBBFF11FDAF87DE" box="[449,559,151,177]" pageId="33" pageNumber="34" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFCD433F0EEDFF5AFD3A879B" blockId="33.[151,698,220,244]" box="[151,698,220,244]" pageId="33" pageNumber="34">
<typeStatus id="54D224A5FFCD433F0EEDFF5AFF49879B" box="[151,201,220,244]" pageId="33" pageNumber="34">Type</typeStatus>
species:
<taxonomicName id="4C69E184FFCD433F0F5CFF5AFDE3879C" authorityName="Uusitalo" authorityYear="2025" box="[294,611,220,244]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vazquezrojasi" status="sp. nov.">
<emphasis id="B91D4615FFCD433F0F5CFF5AFDE3879C" box="[294,611,220,244]" italics="true" pageId="33" pageNumber="34">Glabromichaelia vazquezrojasi</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFCD433F0C10FF5AFD3A879B" bold="true" box="[618,698,220,244]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A22EFB6EFFCD433F0C10FF5AFD3A879B" box="[618,698,220,244]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFCD433F0EEDFEA7FA05857B" blockId="33.[151,1437,289,1396]" pageId="33" pageNumber="34">
<emphasis id="B91D4615FFCD433F0EEDFEA7FE218654" bold="true" box="[151,417,289,315]" pageId="33" pageNumber="34">Differential diagnosis.</emphasis>
The genus differs from other genera of
<taxonomicName id="4C69E184FFCD433F0D00FEA4FB928653" authorityName="Womersley" authorityYear="1944" box="[890,1042,290,316]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="89" phylum="Arthropoda" rank="tribe" tribe="Bimichaeliini">Bimichaeliini</taxonomicName>
and
<taxonomicName id="4C69E184FFCD433F0A35FEA4FA9F8653" box="[1103,1311,290,316]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="33" phylum="Arthropoda" rank="tribe" tribe="Proteromichaeliini">Proteromichaeliini</taxonomicName>
by having parallel, glabrous ridges all along the dorsal and ventral integument (i.e. there is no stellate or cellular pattern of ridges), no primary pattern (i.e. large lamellae on ridges or in clumps), and no secondary pattern with smaller lamellae or granulae in between large lamellae (i.e. integuments reticulation, lamellae and granulae are absent) (
<figureCitation id="13528682FFCD433F0EE5FE34FEAB86A3" box="[159,299,434,460]" captionStart="FIGURE 19" captionStartId="36.[151,250,1874,1898]" captionTargetBox="[151,1435,176,1851]" captionTargetId="figure-24@36.[151,1435,176,1851]" captionTargetPageId="36" captionText="FIGURE 19. Glabromichaelia vazquezrojasi sp. nov. A—ventral view of female, with lowercase letters a-d referring to various areas enlarged in subsequent fragments; B—ventral area (a) with long setae; C—ventral area (b) between coxae with short setae; D—dorsal area (c) with long setae; E—dorsal area (d) demonstrating intensity of neotrichy; F—famulus ɛ I. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Fig. 19BD</figureCitation>
). Other diagnostic features are as follows: the unique presence of large (or baculiform) solenidia on tarsi, tibiae, genua and femora I, II, III and IV, respectively—2200, 1111, 0100, and 0100—as presented as the solenidial pattern in
<bibRefCitation id="EFF8E7F6FFCD433F0F07FE7CFDDB857A" author="Uusitalo, M." box="[381,603,506,533]" pageId="33" pageNumber="34" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo (2010: 25)</bibRefCitation>
. In other respects, the species is similar to the rest of the bimichaeliines.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCD433F0EBDFD9BFA108437" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD69A07FFCD433F0EBDFD9BFA108437" blockId="33.[151,1437,289,1396]" pageId="33" pageNumber="34">
<emphasis id="B91D4615FFCD433F0EBDFD9BFED98558" bold="true" box="[199,345,541,567]" pageId="33" pageNumber="34">Description.</emphasis>
Glabrous larger and finer ridges running parallel and covering dorsal and ventral sides (
<figureCitation id="13528682FFCD433F0B15FD98FF6C8533" captionStart="FIGURE 19" captionStartId="36.[151,250,1874,1898]" captionTargetBox="[151,1435,176,1851]" captionTargetId="figure-24@36.[151,1435,176,1851]" captionTargetPageId="36" captionText="FIGURE 19. Glabromichaelia vazquezrojasi sp. nov. A—ventral view of female, with lowercase letters a-d referring to various areas enlarged in subsequent fragments; B—ventral area (a) with long setae; C—ventral area (b) between coxae with short setae; D—dorsal area (c) with long setae; E—dorsal area (d) demonstrating intensity of neotrichy; F—famulus ɛ I. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Fig. 19BD</figureCitation>
); invaginate areas with softest skin, such as basis of chelicera, joints of leg segments and genital orifice are with rows of small, transverse laminae (
<figureCitation id="13528682FFCD433F0C24FDE0FD4B85EF" box="[606,715,614,640]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs. 20F</figureCitation>
,
<figureCitation id="13528682FFCD433F0CADFDE0FC8B85EF" box="[727,779,614,640]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">21G</figureCitation>
); elongated dorsal and ventral setae with 2030 cilia each (
<figureCitation id="13528682FFCD433F0EE5FD0CFEA885CB" box="[159,296,650,676]" captionStart="FIGURE 19" captionStartId="36.[151,250,1874,1898]" captionTargetBox="[151,1435,176,1851]" captionTargetId="figure-24@36.[151,1435,176,1851]" captionTargetPageId="36" captionText="FIGURE 19. Glabromichaelia vazquezrojasi sp. nov. A—ventral view of female, with lowercase letters a-d referring to various areas enlarged in subsequent fragments; B—ventral area (a) with long setae; C—ventral area (b) between coxae with short setae; D—dorsal area (c) with long setae; E—dorsal area (d) demonstrating intensity of neotrichy; F—famulus ɛ I. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Fig. 19BD</figureCitation>
); naso small with rounded tip and parallel sides, some accessory setae on prodorsal soft integument and
<emphasis id="B91D4615FFCD433F0EEDFD29FF2D85A7" box="[151,173,687,712]" italics="true" pageId="33" pageNumber="34">in</emphasis>
-area, anterior pair of sensilla
<emphasis id="B91D4615FFCD433F0F81FD29FD9485A7" box="[507,532,687,712]" italics="true" pageId="33" pageNumber="34">ve</emphasis>
densely ciliated by cilia of various lengths, longest being distal (
<figureCitation id="13528682FFCD433F0A92FD28FADA85A7" box="[1256,1370,686,712]" captionStart="FIGURE 17" captionStartId="34.[151,250,1800,1824]" captionTargetBox="[167,1432,221,1792]" captionTargetId="figure-57@34.[151,1436,216,1792]" captionTargetPageId="34" captionText="FIGURE 17. Glabromichaelia vazquezrojasi sp. nov. A—prodorsum with elongated andciliate setae; B—reconstructed dorsal habitus, outlines andgrey dots symbolising the expected form of adult and number of missing setae, black dots represent the actually observed setae; C—neotrichous subcapitulum; D—ventral side between coxae without coxal plates. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs. 17A</figureCitation>
,
<figureCitation id="13528682FFCD433F0B1FFD28FF2C8583" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">20A, B</figureCitation>
); chelicerae straight with some teeth but without setae (
<figureCitation id="13528682FFCD433F0D5BFD54FC118583" box="[801,913,722,748]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs. 20C</figureCitation>
,
<figureCitation id="13528682FFCD433F0DE4FD54FC508583" box="[926,976,722,748]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">21C</figureCitation>
); palps short and stout, palpal solenidion medially constricted, in contact with two eupathidia and with a basal tubercle (
<figureCitation id="13528682FFCD433F0D83FD70FBFE847F" box="[1017,1150,758,784]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Fig. 21A, B</figureCitation>
); two baculiform but slim solenidia on tarsi I and II, one baculiform solenidion on each of tibiae 1IV (instead of 2), one baculiform solenidion on genu II and femur II (
<figureCitation id="13528682FFCD433F0FD6FCB8FDBF8437" box="[428,575,830,856]" captionStart="FIGURE 18" captionStartId="35.[151,250,1834,1858]" captionTargetBox="[151,1436,176,1811]" captionTargetId="figure-15@35.[151,1436,176,1811]" captionTargetPageId="35" captionText="FIGURE 18. Glabromichaelia vazquezrojasi sp. nov. A—genital area of female with 1113 genital setae per valve and two pairs of eugenital setae; B—anal area with 10 anal setae per valve; C—leg I; D—leg II. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs. 18C, D</figureCitation>
,
<figureCitation id="13528682FFCD433F0C31FCB8FDF78437" box="[587,631,830,856]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">20F</figureCitation>
,
<figureCitation id="13528682FFCD433F0CF8FCB8FD538437" box="[642,723,830,856]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">21D, E</figureCitation>
,
<figureCitation id="13528682FFCD433F0CA5FCB8FCB78437" box="[735,823,830,856]" captionStart="FIGURE 22" captionStartId="39.[151,250,1384,1408]" captionTargetBox="[151,1436,176,1360]" captionTargetId="figure-15@39.[151,1436,176,1360]" captionTargetPageId="39" captionText="FIGURE 22. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">22AD</figureCitation>
); strong neotrichy already on protonymph (
<figureCitation id="13528682FFCD433F0B66FCB8FA048437" box="[1308,1412,830,856]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Fig. 20C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCD433F0EBDFCE4FC9B83AF" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BD69A07FFCD433F0EBDFCE4FC9B83AF" blockId="33.[151,1437,289,1396]" pageId="33" pageNumber="34">
<emphasis id="B91D4615FFCD433F0EBDFCE4FEBB8413" bold="true" box="[199,315,866,892]" pageId="33" pageNumber="34">Remarks.</emphasis>
The genera in the tribes
<taxonomicName id="4C69E184FFCD433F0C3EFCE4FD5C8413" authorityName="Womersley" authorityYear="1944" box="[580,732,866,892]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="89" phylum="Arthropoda" rank="tribe" tribe="Bimichaeliini">Bimichaeliini</taxonomicName>
and
<taxonomicName id="4C69E184FFCD433F0D75FCE4FC5F8413" box="[783,991,866,892]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="33" phylum="Arthropoda" rank="tribe" tribe="Proteromichaeliini">Proteromichaeliini</taxonomicName>
are based on the form of the integument, which assumes that the form of the integument is synapomorphic for each genus. The genus
<taxonomicName id="4C69E184FFCD433F0AB8FC00FAC984CF" authorityName="Thor" authorityYear="1902" box="[1218,1353,902,928]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCD433F0AB8FC00FAC984CF" box="[1218,1353,902,928]" italics="true" pageId="33" pageNumber="34">Bimichaelia</emphasis>
</taxonomicName>
can be characterised by its granular secondary pattern,
<taxonomicName id="4C69E184FFCD433F0CCEFC2CFCF884AB" authorityName="Uusitalo" authorityYear="2010" box="[692,888,938,964]" class="Arachnida" family="Alycidae" genus="Laminamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCD433F0CCEFC2CFCF884AB" box="[692,888,938,964]" italics="true" pageId="33" pageNumber="34">Laminamichaelia</emphasis>
</taxonomicName>
by the presence of transversely packed lamellae of the secondary pattern, the granulae/lamellae are absent in
<taxonomicName id="4C69E184FFCD433F0D46FC48FC7A8487" authorityName="Uusitalo" authorityYear="2025" box="[828,1018,974,1000]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCD433F0D46FC48FC7A8487" box="[828,1018,974,1000]" italics="true" pageId="33" pageNumber="34">Glabromichaelia</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C69E184FFCD433F0A4DFC48FB7A8487" authorityName="Uusitalo, Ueckerman &amp; Theron" authorityYear="2020" box="[1079,1274,974,1000]" class="Arachnida" family="Alycidae" genus="Proteromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCD433F0A4DFC48FB7A8487" box="[1079,1274,974,1000]" italics="true" pageId="33" pageNumber="34">Proteromichaelia</emphasis>
</taxonomicName>
has clumps of large lamellae inside roundish cells of large lamellae (=primary pattern) plus a secondary pattern by fine ridges. The difference between the patterns of baculiform solenidia of
<taxonomicName id="4C69E184FFCD433F0D43FB90FC77835F" authorityName="Uusitalo" authorityYear="2025" box="[825,1015,1046,1072]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCD433F0D43FB90FC77835F" box="[825,1015,1046,1072]" italics="true" pageId="33" pageNumber="34">Glabromichaelia</emphasis>
</taxonomicName>
(given above) and
<taxonomicName id="4C69E184FFCD433F0AA3FB90FA1C835F" authorityName="Uusitalo, Ueckerman &amp; Theron" authorityYear="2020" box="[1241,1436,1046,1072]" class="Arachnida" family="Alycidae" genus="Proteromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCD433F0AA3FB90FA1C835F" box="[1241,1436,1046,1072]" italics="true" pageId="33" pageNumber="34">Proteromichaelia</emphasis>
</taxonomicName>
is only one solenidion (
<bibRefCitation id="EFF8E7F6FFCD433F0FDFFBBCFDF7833A" author="Uusitalo, M." box="[421,631,1082,1109]" pageId="33" pageNumber="34" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010: 25</bibRefCitation>
, state 7 and appendix 3). The parallel ridges on the integument of the new genus
<taxonomicName id="4C69E184FFCD433F0F6CFBD8FE548317" authorityName="Uusitalo" authorityYear="2025" box="[278,468,1118,1144]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCD433F0F6CFBD8FE548317" box="[278,468,1118,1144]" italics="true" pageId="33" pageNumber="34">Glabromichaelia</emphasis>
</taxonomicName>
(tribe
<taxonomicName id="4C69E184FFCD433F0C5AFBD8FD3A8317" authorityName="Womersley" authorityYear="1944" box="[544,698,1118,1144]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="tribe" tribe="Bimichaeliini">Bimichaeliini</taxonomicName>
) are similar to the ridges in the tribes
<taxonomicName id="4C69E184FFCD433F0A1DFBD8FB3B8317" box="[1127,1211,1118,1144]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="82" phylum="Arthropoda" rank="tribe" tribe="Alycini">Alycini</taxonomicName>
and
<taxonomicName id="4C69E184FFCD433F0A89FBD8FAEC8317" authorityName="Uusitalo" authorityYear="2010" box="[1267,1388,1118,1144]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="89" phylum="Arthropoda" rank="tribe" tribe="Petralycini">Petralycini</taxonomicName>
and support the idea of monophyly in the family
<taxonomicName id="4C69E184FFCD433F0CF8FB04FD6783F3" baseAuthorityName="Uusitalo" baseAuthorityYear="2010" box="[642,743,1154,1180]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="family">Alycidae</taxonomicName>
(see
<bibRefCitation id="EFF8E7F6FFCD433F0D5AFB04FC6883F3" author="Uusitalo, M." box="[800,1000,1154,1180]" pageId="33" pageNumber="34" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010: 11</bibRefCitation>
). That the glabrousness of ridges is due to the loss of laminae/granulae remains an open question.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCD433F0EBDFB4CFD53821B" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BD69A07FFCD433F0EBDFB4CFD53821B" blockId="33.[151,1437,289,1396]" pageId="33" pageNumber="34">
<emphasis id="B91D4615FFCD433F0EBDFB4CFECE838B" bold="true" box="[199,334,1226,1252]" pageId="33" pageNumber="34">Etymology.</emphasis>
The first part of the generic name refers to the glabrous dorsal and ventral ridges of integument. This structure of integument is probably a plesiomorphic character state without primary and secondary pattern, i.e. without granulae and/or lamellae, which are common to all the other members of various species groups of the tribes
<taxonomicName id="4C69E184FFCD433F0EA6FAB0FEF4823F" authorityName="Womersley" authorityYear="1944" box="[220,372,1334,1360]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="89" phylum="Arthropoda" rank="tribe" tribe="Bimichaeliini">Bimichaeliini</taxonomicName>
and
<taxonomicName id="4C69E184FFCD433F0FD3FAB0FDFC823F" box="[425,636,1334,1360]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="33" phylum="Arthropoda" rank="tribe" tribe="Proteromichaeliini">Proteromichaeliini</taxonomicName>
, but also
<taxonomicName id="4C69E184FFCD433F0C98FAB0FCB6823F" box="[738,822,1334,1360]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="82" phylum="Arthropoda" rank="tribe" tribe="Alycini">Alycini</taxonomicName>
and
<taxonomicName id="4C69E184FFCD433F0D11FAB0FC68823F" authorityName="Uusitalo" authorityYear="2010" box="[875,1000,1334,1360]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="89" phylum="Arthropoda" rank="tribe" tribe="Petralycini">Petralycini</taxonomicName>
; and “Michael”, referring to the British acarologist Albert Davidson Michael (18361927).
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName id="4C69E184FFCD433F0EEDFA43FD8D828F" authorityName="Uusitalo" authorityYear="2025" box="[151,525,1477,1504]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="vazquezrojasi" status="sp. nov.">
<emphasis id="B91D4615FFCD433F0EEDFA43FD8D828F" bold="true" box="[151,525,1477,1504]" italics="true" pageId="33" pageNumber="34">Glabromichaelia vazquezrojasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFCD433F0C6FFA40FDF2828F" box="[533,626,1478,1504]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCD433F0EEDFA6CFC8F816B" box="[151,783,1514,1540]" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD69A07FFCD433F0EEDFA6CFC8F816B" blockId="33.[151,783,1477,1540]" box="[151,783,1514,1540]" pageId="33" pageNumber="34">
(
<figureCitation id="13528682FFCD433F0EE5FA6CFEB4816B" box="[159,308,1514,1540]" captionStart="FIGURE 17" captionStartId="34.[151,250,1800,1824]" captionTargetBox="[167,1432,221,1792]" captionTargetId="figure-57@34.[151,1436,216,1792]" captionTargetPageId="34" captionText="FIGURE 17. Glabromichaelia vazquezrojasi sp. nov. A—prodorsum with elongated andciliate setae; B—reconstructed dorsal habitus, outlines andgrey dots symbolising the expected form of adult and number of missing setae, black dots represent the actually observed setae; C—neotrichous subcapitulum; D—ventral side between coxae without coxal plates. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs. 17AD</figureCitation>
,
<figureCitation id="13528682FFCD433F0F45FA6CFE14816B" box="[319,404,1514,1540]" captionStart="FIGURE 18" captionStartId="35.[151,250,1834,1858]" captionTargetBox="[151,1436,176,1811]" captionTargetId="figure-15@35.[151,1436,176,1811]" captionTargetPageId="35" captionText="FIGURE 18. Glabromichaelia vazquezrojasi sp. nov. A—genital area of female with 1113 genital setae per valve and two pairs of eugenital setae; B—anal area with 10 anal setae per valve; C—leg I; D—leg II. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">18AD</figureCitation>
,
<figureCitation id="13528682FFCD433F0FDAFA6CFE6E816B" box="[416,494,1514,1540]" captionStart="FIGURE 19" captionStartId="36.[151,250,1874,1898]" captionTargetBox="[151,1435,176,1851]" captionTargetId="figure-24@36.[151,1435,176,1851]" captionTargetPageId="36" captionText="FIGURE 19. Glabromichaelia vazquezrojasi sp. nov. A—ventral view of female, with lowercase letters a-d referring to various areas enlarged in subsequent fragments; B—ventral area (a) with long setae; C—ventral area (b) between coxae with short setae; D—dorsal area (c) with long setae; E—dorsal area (d) demonstrating intensity of neotrichy; F—famulus ɛ I. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">19AF</figureCitation>
,
<figureCitation id="13528682FFCD433F0F83FA6CFDC8816B" box="[505,584,1514,1540]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">20AF</figureCitation>
,
<figureCitation id="13528682FFCD433F0C29FA6CFD28816B" box="[595,680,1514,1540]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">21AH</figureCitation>
,
<figureCitation id="13528682FFCD433F0CC9FA6CFC87816B" box="[691,775,1514,1540]" captionStart="FIGURE 22" captionStartId="39.[151,250,1384,1408]" captionTargetBox="[151,1436,176,1360]" captionTargetId="figure-15@39.[151,1436,176,1360]" captionTargetPageId="39" captionText="FIGURE 22. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">22AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCD433F0EEDF9A8FC358129" box="[151,949,1582,1606]" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="8BD69A07FFCD433F0EEDF9A8FC358129" blockId="33.[151,949,1582,1606]" box="[151,949,1582,1606]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB79BD29FFCD433F0EEDF9A8FC358129" box="[151,949,1582,1606]" pageId="33" pageNumber="34">
<taxonomicName id="4C69E184FFCD433F0EEDF9A8FE348129" ID-CoL="68KZ4" authorityName="Grandjean" authorityYear="1939" box="[151,436,1582,1606]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="campylognatha">
<emphasis id="B91D4615FFCD433F0EEDF9A8FE348129" box="[151,436,1582,1606]" italics="true" pageId="33" pageNumber="34">Bimichaelia campylognatha</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AC8BC16FFCD433F0FAEF9A8FD4E8129" author="Vazquez Rojas, I. M." box="[468,718,1582,1606]" page="19" pageId="33" pageNumber="34" year="2002">
<bibRefCitation id="EFF8E7F6FFCD433F0FAEF9A8FD4E8129" author="Vazquez Rojas, I. M." box="[468,718,1582,1606]" pageId="33" pageNumber="34" refId="ref49098" refString="Vazquez Rojas, I. M. (2002) Prostigmata (Acarida) edaficos de una zona litoral del Golfo di Mexico. PhD Thesis, Universidad Nacional Autonome de Mexico, Mexico City, 96 pp. Available from: https://www.fciencias.unam.mx/directorio (accessed 3 January 2022)" type="book" year="2002">Vázquez Rojas 2002: 19</bibRefCitation>
</treatmentCitation>
, in part (
<specimenCount id="9D6F518EFFCD433F0D52F9A8FC2A8129" box="[808,938,1582,1606]" count="2" pageId="33" pageNumber="34" type="generic">2 specimens</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCD43390EEDF9F2FAF18187" lastPageId="39" lastPageNumber="40" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD69A07FFCD433F0EEDF9F2FAD98071" blockId="33.[151,1436,1652,2039]" pageId="33" pageNumber="34">
<emphasis id="B91D4615FFCD433F0EEDF9F2FEA981E1" bold="true" box="[151,297,1652,1678]" pageId="33" pageNumber="34">Description.</emphasis>
<emphasis id="B91D4615FFCD433F0F4BF9F3FE0B81E1" box="[305,395,1653,1678]" italics="true" pageId="33" pageNumber="34">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFCD433F0FC4F9F3FD9F81E1" box="[446,543,1652,1679]" count="1" pageId="33" pageNumber="34" type="female">1 female</specimenCount>
,
<figureCitation id="13528682FFCD433F0C51F9F2FD3F81E0" box="[555,703,1652,1679]" captionStart="FIGURE 17" captionStartId="34.[151,250,1800,1824]" captionTargetBox="[167,1432,221,1792]" captionTargetId="figure-57@34.[151,1436,216,1792]" captionTargetPageId="34" captionText="FIGURE 17. Glabromichaelia vazquezrojasi sp. nov. A—prodorsum with elongated andciliate setae; B—reconstructed dorsal habitus, outlines andgrey dots symbolising the expected form of adult and number of missing setae, black dots represent the actually observed setae; C—neotrichous subcapitulum; D—ventral side between coxae without coxal plates. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs. 17A, B</figureCitation>
,
<figureCitation id="13528682FFCD433F0CB6F9F3FCA081E0" box="[716,800,1653,1679]" captionStart="FIGURE 19" captionStartId="36.[151,250,1874,1898]" captionTargetBox="[151,1435,176,1851]" captionTargetId="figure-24@36.[151,1435,176,1851]" captionTargetPageId="36" captionText="FIGURE 19. Glabromichaelia vazquezrojasi sp. nov. A—ventral view of female, with lowercase letters a-d referring to various areas enlarged in subsequent fragments; B—ventral area (a) with long setae; C—ventral area (b) between coxae with short setae; D—dorsal area (c) with long setae; E—dorsal area (d) demonstrating intensity of neotrichy; F—famulus ɛ I. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">19D, E</figureCitation>
). Length 1000 µm; intense neotrichy, ciliated setae with 1516 µm long shafts and with mutual distance 1720 µm from each other covering whole dorsum without clear signs of segments; integument with parallel ridges on dorsal, lateral and ventral sides and legs; naso rounded, slightly smaller basally, reticulate pattern beneath integument on crista and prodorsal sensillar area; prodorsum neotrichous, cilia of filamentous sensilla
<emphasis id="B91D4615FFCD433F0FA9F883FE6C8071" box="[467,492,1797,1822]" italics="true" pageId="33" pageNumber="34">ve</emphasis>
increasing in size towards distal end; setae from
<emphasis id="B91D4615FFCD433F0A69F883FBA98071" box="[1043,1065,1797,1822]" italics="true" pageId="33" pageNumber="34">in</emphasis>
-area penetrate far anterior.
</paragraph>
<paragraph id="8BD69A07FFCD433F0EBDF8AFFC348009" blockId="33.[151,1436,1652,2039]" pageId="33" pageNumber="34">
<taxonomicName id="4C69E184FFCD433F0EBDF8AFFE8F802D" box="[199,271,1833,1858]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFCD433F0EBDF8AFFE8F802D" box="[199,271,1833,1858]" italics="true" pageId="33" pageNumber="34">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFCD433F0F65F8AEFE34802C" box="[287,436,1832,1859]" captionStart="FIGURE 17" captionStartId="34.[151,250,1800,1824]" captionTargetBox="[167,1432,221,1792]" captionTargetId="figure-57@34.[151,1436,216,1792]" captionTargetPageId="34" captionText="FIGURE 17. Glabromichaelia vazquezrojasi sp. nov. A—prodorsum with elongated andciliate setae; B—reconstructed dorsal habitus, outlines andgrey dots symbolising the expected form of adult and number of missing setae, black dots represent the actually observed setae; C—neotrichous subcapitulum; D—ventral side between coxae without coxal plates. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs. 17C, D</figureCitation>
,
<figureCitation id="13528682FFCD433F0FBAF8AFFD95802C" box="[448,533,1833,1859]" captionStart="FIGURE 18" captionStartId="35.[151,250,1834,1858]" captionTargetBox="[151,1436,176,1811]" captionTargetId="figure-15@35.[151,1436,176,1811]" captionTargetPageId="35" captionText="FIGURE 18. Glabromichaelia vazquezrojasi sp. nov. A—genital area of female with 1113 genital setae per valve and two pairs of eugenital setae; B—anal area with 10 anal setae per valve; C—leg I; D—leg II. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">18A, B</figureCitation>
,
<figureCitation id="13528682FFCD433F0C5BF8AFFDF7802C" box="[545,631,1833,1859]" captionStart="FIGURE 19" captionStartId="36.[151,250,1874,1898]" captionTargetBox="[151,1435,176,1851]" captionTargetId="figure-24@36.[151,1435,176,1851]" captionTargetPageId="36" captionText="FIGURE 19. Glabromichaelia vazquezrojasi sp. nov. A—ventral view of female, with lowercase letters a-d referring to various areas enlarged in subsequent fragments; B—ventral area (a) with long setae; C—ventral area (b) between coxae with short setae; D—dorsal area (c) with long setae; E—dorsal area (d) demonstrating intensity of neotrichy; F—famulus ɛ I. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">19AC</figureCitation>
). Setae between coxal areas (cxII-cxIII) shorter than elsewhere; genital valves each with 1113 genital setae per valve; 10 anal setae per valve.
</paragraph>
<paragraph id="8BD69A07FFCD433F0EBDF8F6FAA380C1" blockId="33.[151,1436,1652,2039]" pageId="33" pageNumber="34">
<taxonomicName id="4C69E184FFCD433F0EBDF8F6FED680E5" authorityName="Krefft" authorityYear="1871" box="[199,342,1904,1930]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="33" pageNumber="34" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFCD433F0EBDF8F6FED680E5" box="[199,342,1904,1930]" italics="true" pageId="33" pageNumber="34">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFCD433F0F18F8F6FE4F80E4" box="[354,463,1904,1931]" captionStart="FIGURE 17" captionStartId="34.[151,250,1800,1824]" captionTargetBox="[167,1432,221,1792]" captionTargetId="figure-57@34.[151,1436,216,1792]" captionTargetPageId="34" captionText="FIGURE 17. Glabromichaelia vazquezrojasi sp. nov. A—prodorsum with elongated andciliate setae; B—reconstructed dorsal habitus, outlines andgrey dots symbolising the expected form of adult and number of missing setae, black dots represent the actually observed setae; C—neotrichous subcapitulum; D—ventral side between coxae without coxal plates. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs.17C</figureCitation>
,
<figureCitation id="13528682FFCD433F0FADF8F7FDAC80E4" box="[471,556,1905,1931]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">21AC</figureCitation>
).Subcapitulum neotrichous;chelicerae straight, thin integument for invagination with lamellae, lower digits with teeth; medially constricted palpal solenidion in contact with eupathidia.
</paragraph>
<paragraph id="8BD69A07FFCD433C0EBDF83FFB0887BA" blockId="33.[151,1436,1652,2039]" lastBlockId="34.[151,1436,151,213]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<emphasis id="B91D4615FFCD433F0EBDF83FFF7C80BD" box="[199,252,1977,2002]" italics="true" pageId="33" pageNumber="34">Legs</emphasis>
(
<figureCitation id="13528682FFCD433F0F74F83EFE2680BC" box="[270,422,1976,2003]" captionStart="FIGURE 17" captionStartId="34.[151,250,1800,1824]" captionTargetBox="[167,1432,221,1792]" captionTargetId="figure-57@34.[151,1436,216,1792]" captionTargetPageId="34" captionText="FIGURE 17. Glabromichaelia vazquezrojasi sp. nov. A—prodorsum with elongated andciliate setae; B—reconstructed dorsal habitus, outlines andgrey dots symbolising the expected form of adult and number of missing setae, black dots represent the actually observed setae; C—neotrichous subcapitulum; D—ventral side between coxae without coxal plates. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">Figs. 17C, D</figureCitation>
,
<figureCitation id="13528682FFCD433F0FCEF83FFD8A80BC" box="[436,522,1977,2003]" captionStart="FIGURE 18" captionStartId="35.[151,250,1834,1858]" captionTargetBox="[151,1436,176,1811]" captionTargetId="figure-15@35.[151,1436,176,1811]" captionTargetPageId="35" captionText="FIGURE 18. Glabromichaelia vazquezrojasi sp. nov. A—genital area of female with 1113 genital setae per valve and two pairs of eugenital setae; B—anal area with 10 anal setae per valve; C—leg I; D—leg II. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">18C, D</figureCitation>
,
<figureCitation id="13528682FFCD433F0C62F83FFD0580BC" box="[536,645,1977,2003]" captionStart="FIGURE 19" captionStartId="36.[151,250,1874,1898]" captionTargetBox="[151,1435,176,1851]" captionTargetId="figure-24@36.[151,1435,176,1851]" captionTargetPageId="36" captionText="FIGURE 19. Glabromichaelia vazquezrojasi sp. nov. A—ventral view of female, with lowercase letters a-d referring to various areas enlarged in subsequent fragments; B—ventral area (a) with long setae; C—ventral area (b) between coxae with short setae; D—dorsal area (c) with long setae; E—dorsal area (d) demonstrating intensity of neotrichy; F—famulus ɛ I. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">19A, 19F</figureCitation>
,
<figureCitation id="13528682FFCD433F0CE9F83FFD3F80BC" box="[659,703,1977,2003]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">20F</figureCitation>
,
<figureCitation id="13528682FFCD433F0CB6F83FFD7C80BC" box="[716,764,1977,2003]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="33" pageNumber="34">21F</figureCitation>
). Solenidial formula for tarsi, tibiae, genua and femora on legs I, II, III and IV, respectively: 2B-2B-0-0, 1B-1B-1B-1B, 5C-1B2C-1C-1C, 1C-1B-0-0, pestle-like famulus I placed abaxially near distal end of tarsus I; elastic skin of joints with rows of tiny lamellae (white arrows), enabling movements; claws with 90º angle midway between basis and distal tip; empodia clawless.
</paragraph>
<caption id="DF16CA8FFFCE433C0EEDF88EFBF680E2" pageId="34" pageNumber="35" startId="34.[151,250,1800,1824]" targetBox="[167,1432,221,1792]" targetPageId="34" targetType="figure">
<paragraph id="8BD69A07FFCE433C0EEDF88EFD3A804E" blockId="34.[151,1437,1800,1933]" box="[151,698,1800,1825]" pageId="34" pageNumber="35">
<emphasis id="B91D4615FFCE433C0EEDF88EFEA0804F" bold="true" box="[151,288,1800,1824]" pageId="34" pageNumber="35">FIGURE 17.</emphasis>
<taxonomicName id="4C69E184FFCE433C0F5CF88EFDE3804F" authorityName="Uusitalo" authorityYear="2025" box="[294,611,1800,1824]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="vazquezrojasi" status="sp. nov.">
<emphasis id="B91D4615FFCE433C0F5CF88EFDE3804F" box="[294,611,1800,1824]" italics="true" pageId="34" pageNumber="35">Glabromichaelia vazquezrojasi</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFCE433C0C13F88FFD3A804E" bold="true" box="[617,698,1801,1825]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="A22EFB6EFFCE433C0C13F88FFD3A804E" box="[617,698,1801,1825]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFCE433C0EEDF8AAFBF680E2" blockId="34.[151,1437,1800,1933]" pageId="34" pageNumber="35">A—prodorsum with elongated and ciliate setae; B—reconstructed dorsal habitus, outlines and grey dots symbolising the expected form of adult and number of missing setae, black dots represent the actually observed setae; C—neotrichous subcapitulum; D—ventral side between coxae without coxal plates. All specimens—from Veracruz, CICOLMA.</paragraph>
</caption>
<paragraph id="8BD69A07FFCE433C0EBDF83DFCD98096" blockId="34.[151,1436,1979,2041]" pageId="34" pageNumber="35">
<emphasis id="B91D4615FFCE433C0EBDF83DFED080BA" box="[199,336,1979,2005]" italics="true" pageId="34" pageNumber="35">Protonymph</emphasis>
(n=1,
<figureCitation id="13528682FFCE433C0FE5F83DFDAB80BA" box="[415,555,1979,2005]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStart-2="FIGURE 22" captionStartId-0="37.[151,250,1873,1897]" captionStartId-1="38.[151,250,1759,1783]" captionStartId-2="39.[151,250,1384,1408]" captionTargetBox-0="[151,1435,176,1849]" captionTargetBox-1="[151,1435,224,1735]" captionTargetBox-2="[151,1436,176,1360]" captionTargetId-0="figure-15@37.[151,1435,176,1849]" captionTargetId-1="figure-24@38.[151,1435,224,1735]" captionTargetId-2="figure-15@39.[151,1436,176,1360]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionText-0="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." captionText-1="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." captionText-2="FIGURE 22. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV. All specimens—from Veracruz, CICOLMA." pageId="34" pageNumber="35">Figs. 2022</figureCitation>
).
<emphasis id="B91D4615FFCE433C0C45F83AFD1980BA" box="[575,665,1980,2005]" italics="true" pageId="34" pageNumber="35">Dorsum</emphasis>
(
<figureCitation id="13528682FFCE433C0CD0F83DFCB880BA" box="[682,824,1979,2005]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="34" pageNumber="35">Fig. 20AD</figureCitation>
). Length 350 µm; slightly thinner neotrichy but setae similar in size with adult; a few additional setae on prodorsum.
</paragraph>
<caption id="DF16CA8FFFCF433D0EEDF8ACFC1880E5" pageId="35" pageNumber="36" startId="35.[151,250,1834,1858]" targetBox="[151,1436,176,1811]" targetPageId="35" targetType="figure">
<paragraph id="8BD69A07FFCF433D0EEDF8ACFD3A802D" blockId="35.[151,1437,1834,1930]" box="[151,698,1834,1858]" pageId="35" pageNumber="36">
<emphasis id="B91D4615FFCF433D0EEDF8ACFEA0802D" bold="true" box="[151,288,1834,1858]" pageId="35" pageNumber="36">FIGURE 18.</emphasis>
<taxonomicName id="4C69E184FFCF433D0F5CF8ACFDE3802D" authorityName="Uusitalo" authorityYear="2025" box="[294,611,1834,1858]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="vazquezrojasi" status="sp. nov.">
<emphasis id="B91D4615FFCF433D0F5CF8ACFDE3802D" box="[294,611,1834,1858]" italics="true" pageId="35" pageNumber="36">Glabromichaelia vazquezrojasi</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFCF433D0C13F8ACFD3A802D" bold="true" box="[617,698,1834,1858]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A22EFB6EFFCF433D0C13F8ACFD3A802D" box="[617,698,1834,1858]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFCF433D0EEDF8C8FC1880E5" blockId="35.[151,1437,1834,1930]" pageId="35" pageNumber="36">A—genital area of female with 1113 genital setae per valve and two pairs of eugenital setae; B—anal area with 10 anal setae per valve; C—leg I; D—leg II. All specimens—from Veracruz, CICOLMA.</paragraph>
</caption>
<caption id="DF16CA8FFFC8433A0EEDF8D4FB2180B8" pageId="36" pageNumber="37" startId="36.[151,250,1874,1898]" targetBox="[151,1435,176,1851]" targetPageId="36" targetType="figure">
<paragraph id="8BD69A07FFC8433A0EEDF8D4FD3A8004" blockId="36.[151,1436,1874,2007]" box="[151,698,1874,1899]" pageId="36" pageNumber="37">
<emphasis id="B91D4615FFC8433A0EEDF8D4FEA08005" bold="true" box="[151,288,1874,1898]" pageId="36" pageNumber="37">FIGURE 19.</emphasis>
<taxonomicName id="4C69E184FFC8433A0F5CF8D5FDE38005" authorityName="Uusitalo" authorityYear="2025" box="[294,611,1875,1899]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="vazquezrojasi" status="sp. nov.">
<emphasis id="B91D4615FFC8433A0F5CF8D5FDE38005" box="[294,611,1875,1899]" italics="true" pageId="36" pageNumber="37">Glabromichaelia vazquezrojasi</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFC8433A0C13F8D5FD3A8004" bold="true" box="[617,698,1875,1899]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A22EFB6EFFC8433A0C13F8D5FD3A8004" box="[617,698,1875,1899]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFC8433A0EEDF8F1FB2180B8" blockId="36.[151,1436,1874,2007]" pageId="36" pageNumber="37">
A—ventral view of female, with lowercase letters
<emphasis id="B91D4615FFC8433A0CE0F8F1FD3D80E0" bold="true" box="[666,701,1911,1935]" pageId="36" pageNumber="37">a -d</emphasis>
referring to various areas enlarged in subsequent fragments; B—ventral area (
<emphasis id="B91D4615FFC8433A0EAAF81DFF5D80DC" bold="true" box="[208,221,1947,1971]" pageId="36" pageNumber="37">a</emphasis>
) with long setae; C—ventral area (
<emphasis id="B91D4615FFC8433A0C44F81DFDCC80DC" bold="true" box="[574,588,1947,1971]" pageId="36" pageNumber="37">b</emphasis>
) between coxae with short setae; D—dorsal area (
<emphasis id="B91D4615FFC8433A0A3DF81DFBD280DC" bold="true" box="[1095,1106,1947,1971]" pageId="36" pageNumber="37">c</emphasis>
) with long setae; E—dorsal area (
<emphasis id="B91D4615FFC8433A0EDAF839FF2E80B8" bold="true" box="[160,174,1983,2007]" pageId="36" pageNumber="37">d</emphasis>
) demonstrating intensity of neotrichy; F—famulus ɛ I. All specimens—from Veracruz, CICOLMA.
</paragraph>
</caption>
<caption id="DF16CA8FFFC9433B0EEDF8D7FE938096" pageId="37" pageNumber="38" startId="37.[151,250,1873,1897]" targetBox="[151,1435,176,1849]" targetPageId="37" targetType="figure">
<paragraph id="8BD69A07FFC9433B0EEDF8D7FCCA8006" blockId="37.[151,1437,1873,2041]" box="[151,842,1873,1897]" pageId="37" pageNumber="38">
<emphasis id="B91D4615FFC9433B0EEDF8D7FEA08006" bold="true" box="[151,288,1873,1897]" pageId="37" pageNumber="38">FIGURE 20.</emphasis>
<taxonomicName id="4C69E184FFC9433B0F5CF8D7FDE38006" authorityName="Uusitalo" authorityYear="2025" box="[294,611,1873,1897]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vazquezrojasi" status="sp. nov.">
<emphasis id="B91D4615FFC9433B0F5CF8D7FDE38006" box="[294,611,1873,1897]" italics="true" pageId="37" pageNumber="38">Glabromichaelia vazquezrojasi</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFC9433B0C13F8D7FD3A8006" bold="true" box="[617,698,1873,1897]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="A22EFB6EFFC9433B0C13F8D7FD3A8006" box="[617,698,1873,1897]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, protonymph.
</paragraph>
<paragraph id="8BD69A07FFC9433B0EEDF8F3FE938096" blockId="37.[151,1437,1873,2041]" pageId="37" pageNumber="38">
A—prodorsum of protonyph; B—sensilla
<emphasis id="B91D4615FFC9433B0C38F8F0FDD880E2" box="[578,600,1910,1933]" italics="true" pageId="37" pageNumber="38">ve</emphasis>
and
<emphasis id="B91D4615FFC9433B0CF2F8F0FD2480E2" box="[648,676,1910,1933]" italics="true" pageId="37" pageNumber="38">sci</emphasis>
; C—lateral habitus of protonymph, with lowercase letters
<emphasis id="B91D4615FFC9433B0A89F8F3FA8080E2" bold="true" box="[1267,1280,1909,1933]" pageId="37" pageNumber="38">a</emphasis>
and
<emphasis id="B91D4615FFC9433B0B4AF8F3FABE80E2" bold="true" box="[1328,1342,1909,1933]" pageId="37" pageNumber="38">b</emphasis>
referring to integumental areas enlarged in fragments D and E; D—Integument (
<emphasis id="B91D4615FFC9433B0DFAF81FFC0D80DE" bold="true" box="[896,909,1945,1969]" pageId="37" pageNumber="38">a</emphasis>
) of dorsal side; E—integument (
<emphasis id="B91D4615FFC9433B0A9DF81FFB7580DE" bold="true" box="[1255,1269,1945,1969]" pageId="37" pageNumber="38">b</emphasis>
) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints. All specimens—from Veracruz, CICOLMA.
</paragraph>
</caption>
<caption id="DF16CA8FFFCA43380EEDF959FAB4800C" pageId="38" pageNumber="39" startId="38.[151,250,1759,1783]" targetBox="[151,1435,224,1735]" targetPageId="38" targetType="figure">
<paragraph id="8BD69A07FFCA43380EEDF959FCCA8198" blockId="38.[151,1437,1759,1891]" box="[151,842,1759,1783]" pageId="38" pageNumber="39">
<emphasis id="B91D4615FFCA43380EEDF959FEA08198" bold="true" box="[151,288,1759,1783]" pageId="38" pageNumber="39">FIGURE 21.</emphasis>
<taxonomicName id="4C69E184FFCA43380F5CF959FDE38198" authorityName="Uusitalo" authorityYear="2025" box="[294,611,1759,1783]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="vazquezrojasi" status="sp. nov.">
<emphasis id="B91D4615FFCA43380F5CF959FDE38198" box="[294,611,1759,1783]" italics="true" pageId="38" pageNumber="39">Glabromichaelia vazquezrojasi</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFCA43380C13F959FD3A8198" bold="true" box="[617,698,1759,1783]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="A22EFB6EFFCA43380C13F959FD3A8198" box="[617,698,1759,1783]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, protonymph.
</paragraph>
<paragraph id="8BD69A07FFCA43380EEDF885FAB4800C" blockId="38.[151,1437,1759,1891]" pageId="38" pageNumber="39">A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA.</paragraph>
</caption>
<caption id="DF16CA8FFFCB43390EEDFAEEFA8682CB" pageId="39" pageNumber="40" startId="39.[151,250,1384,1408]" targetBox="[151,1436,176,1360]" targetPageId="39" targetType="figure">
<paragraph id="8BD69A07FFCB43390EEDFAEEFCCA82EF" blockId="39.[151,1286,1384,1444]" box="[151,842,1384,1408]" pageId="39" pageNumber="40">
<emphasis id="B91D4615FFCB43390EEDFAEEFEA082EF" bold="true" box="[151,288,1384,1408]" pageId="39" pageNumber="40">FIGURE 22.</emphasis>
<taxonomicName id="4C69E184FFCB43390F5CFAEEFDE382EF" authorityName="Uusitalo" authorityYear="2025" box="[294,611,1384,1408]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vazquezrojasi" status="sp. nov.">
<emphasis id="B91D4615FFCB43390F5CFAEEFDE382EF" box="[294,611,1384,1408]" italics="true" pageId="39" pageNumber="40">Glabromichaelia vazquezrojasi</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFCB43390C13FAEEFD3A82EF" bold="true" box="[617,698,1384,1408]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="A22EFB6EFFCB43390C13FAEEFD3A82EF" box="[617,698,1384,1408]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, protonymph.
</paragraph>
<paragraph id="8BD69A07FFCB43390EEDFA0AFA8682CB" blockId="39.[151,1286,1384,1444]" box="[151,1286,1420,1444]" pageId="39" pageNumber="40">A—leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV. All specimens—from Veracruz, CICOLMA.</paragraph>
</caption>
<paragraph id="8BD69A07FFCB43390EBDFA55FF5B817F" blockId="39.[151,1437,1490,2021]" pageId="39" pageNumber="40">
<taxonomicName id="4C69E184FFCB43390EBDFA55FE8F8283" box="[199,271,1491,1516]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFCB43390EBDFA55FE8F8283" box="[199,271,1491,1516]" italics="true" pageId="39" pageNumber="40">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFCB43390F58FA54FE148283" box="[290,404,1490,1516]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="39" pageNumber="40">Figs. 20E</figureCitation>
,
<figureCitation id="13528682FFCB43390FDEFA54FE7E8283" box="[420,510,1490,1516]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="39" pageNumber="40">21G, H</figureCitation>
). Setae shorter than on adult; one pair of genital setae; 23 anal setae per anal valve.
</paragraph>
<paragraph id="8BD69A07FFCB43390EBDF99CFE948113" blockId="39.[151,1437,1490,2021]" pageId="39" pageNumber="40">
<taxonomicName id="4C69E184FFCB43390EBDF99CFED6815B" authorityName="Krefft" authorityYear="1871" box="[199,342,1562,1588]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="39" pageNumber="40" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFCB43390EBDF99CFED6815B" box="[199,342,1562,1588]" italics="true" pageId="39" pageNumber="40">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFCB43390F11F99CFE7B815B" box="[363,507,1562,1588]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="39" pageNumber="40">Fig. 21AC</figureCitation>
). Chelicerae straight, ca. 6 teeth in movable digits each, elastic skin enabling invagination of chelicerae with rows of tiny lamellae; palpal solenidion medially constricted, in contact with two eupathidia.
</paragraph>
<paragraph id="8BD69A07FFCB43390EBDF901FAF18187" blockId="39.[151,1437,1490,2021]" pageId="39" pageNumber="40">
<emphasis id="B91D4615FFCB43390EBDF901FF7C81CF" box="[199,252,1671,1696]" italics="true" pageId="39" pageNumber="40">Legs</emphasis>
(
<figureCitation id="13528682FFCB43390F70F900FE2F81CF" box="[266,431,1670,1696]" captionStart="FIGURE 20" captionStartId="37.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@37.[151,1435,176,1849]" captionTargetPageId="37" captionText="FIGURE 20. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—prodorsum of protonyph; B—sensilla ve and sci; C—lateral habitus of protonymph, with lowercase letters a and b referring to integumental areas enlarged in fragments D and E; D—Integument (a) of dorsal side; E—integument (b) of ventral side between coxae; F—proximal part of leg II showing granular soft integuments in movable joints.All specimens—from Veracruz, CICOLMA." pageId="39" pageNumber="40">Figs. 20C, 20F</figureCitation>
,
<figureCitation id="13528682FFCB43390FC3F900FD8781CF" box="[441,519,1670,1696]" captionStart="FIGURE 21" captionStartId="38.[151,250,1759,1783]" captionTargetBox="[151,1435,224,1735]" captionTargetId="figure-24@38.[151,1435,224,1735]" captionTargetPageId="38" captionText="FIGURE 21. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—lateral view of palps; B—palpal eupathid in contact with solenidion; C—cheliceral chelae with teeth; D—distal end of tarsus I with 2 solenidia and famulus I; E—solenidion on tarsus II; F—claws and clawless empodium; G—genital valve with 1 seta and genital papilla of protonymph; H—anal valves of protonymph. All specimens—from Veracruz, CICOLMA." pageId="39" pageNumber="40">21DF</figureCitation>
,
<figureCitation id="13528682FFCB43390C6BF900FDE981CF" box="[529,617,1670,1696]" captionStart="FIGURE 22" captionStartId="39.[151,250,1384,1408]" captionTargetBox="[151,1436,176,1360]" captionTargetId="figure-15@39.[151,1436,176,1360]" captionTargetPageId="39" captionText="FIGURE 22. Glabromichaelia vazquezrojasi sp. nov., protonymph. A—leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV. All specimens—from Veracruz, CICOLMA." pageId="39" pageNumber="40">22AD</figureCitation>
). Solenidial formula for tarsi, tibiae, genua and femora on legs I, II, III and IV, respectively: 2B-1B-0-0, 1B-1B-1B-1B, 3C-1B2C-1C-1C, 1C-1B-0-0, pestle-like famulus I placed abaxially near distal end of tarsus I; elastic skin in joints with lamellae; anterior baculiform solenidia on tarsi I (ω1) thin.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCB43390EBDF974FCAF8017" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8BD69A07FFCB43390EBDF974FCAF8017" blockId="39.[151,1437,1490,2021]" pageId="39" pageNumber="40">
<emphasis id="B91D4615FFCB43390EBDF974FEF88063" bold="true" box="[199,376,1778,1804]" pageId="39" pageNumber="40">Type material.</emphasis>
<materialsCitation id="3B01905AFFCB43390FFBF974FCAB8017" collectingDate="1992-06-12" collectingDateMax="1992-12-08" collectingDateMin="1992-06-12" collectionCode="UNAM" country="Mexico" latitude="19.5" location="Centro de Investigaciones Costeras La Mancha" longLatPrecision="1271" longitude="-96.61667" municipality="Ignacio Vazquez Rojas" pageId="39" pageNumber="40" specimenCount="2" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
Ca. 150 high-quality photos of
<specimenCount id="9D6F518EFFCB43390C96F974FCF98063" box="[748,889,1778,1804]" count="2" pageId="39" pageNumber="40" type="generic">2 specimens</specimenCount>
(female
<typeStatus id="54D224A5FFCB43390D9BF974FBC38063" box="[993,1091,1778,1804]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
and protonymph
<typeStatus id="54D224A5FFCB43390B6BF974FAF38063" box="[1297,1395,1778,1804]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
) as
<taxonomicName id="4C69E184FFCB43390EEDF890FE51805F" authorityName="Grandjean" authorityYear="1939" box="[151,465,1814,1840]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="campylognatha">
<emphasis id="B91D4615FFCB43390EEDF890FE51805F" box="[151,465,1814,1840]" italics="true" pageId="39" pageNumber="40">Bimichaelia campylognatha</emphasis>
</taxonomicName>
from litter, caducifolious rain forest (selva baja), CICOLMA (
<location id="8EB6CCDCFFCB43390A01F890FE01803B" LSID="urn:lsid:plazi:treatment:03C02B11FFCD43360EEDFA43FBD68413:8EB6CCDCFFCB43390A01F890FE01803B" country="Mexico" latitude="19.5" longLatPrecision="1271" longitude="-96.61667" municipality="Ignacio Vazquez Rojas" name="Centro de Investigaciones Costeras La Mancha" pageId="39" pageNumber="40" stateProvince="Veracruz de Ignacio de la Llave">Centro de Investigaciones Costeras La Mancha</location>
),
<geoCoordinate id="EE5DFCC0FFCB43390FE8F8BCFE73803B" box="[402,499,1850,1876]" degrees="19" direction="north" minutes="30" orientation="latitude" pageId="39" pageNumber="40" precision="925" value="19.5">19°30N</geoCoordinate>
<geoCoordinate id="EE5DFCC0FFCB43390F83F8BCFDDE803A" box="[505,606,1850,1877]" degrees="96" direction="west" minutes="37" orientation="longitude" pageId="39" pageNumber="40" precision="925" value="-96.61667">96°37W</geoCoordinate>
,
<collectingRegion id="49AD54E5FFCB43390C11F8BDFD52803A" box="[619,722,1851,1877]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="39" pageNumber="40">Veracruz</collectingRegion>
,
<collectingCountry id="F37EDA97FFCB43390CA1F8BCFCB3803B" box="[731,819,1850,1876]" name="Mexico" pageId="39" pageNumber="40">Mexico</collectingCountry>
,
<date id="FFD7BCC7FFCB43390D44F8BCFC55803B" box="[830,981,1850,1877]" pageId="39" pageNumber="40" value="1992-06-12">
<collectingDate id="EF93452FFFCB43390D44F8BCFC55803B" box="[830,981,1850,1877]" pageId="39" pageNumber="40" value="1992-06-12">12 June 1992</collectingDate>
</date>
, and
<date id="FFD7BCC7FFCB43390A74F8BCFB56803B" box="[1038,1238,1850,1876]" pageId="39" pageNumber="40" value="1992-12-08">
<collectingDate id="EF93452FFFCB43390A74F8BCFB56803B" box="[1038,1238,1850,1876]" pageId="39" pageNumber="40" value="1992-12-08">8 December 1992</collectingDate>
</date>
,
<collectingMunicipality id="6BB2007DFFCB43390A9BF8BCFF5A8017" pageId="39" pageNumber="40">Ignacio Vázquez Rojas</collectingMunicipality>
, slides ALY/VER/005 and ALY/VER/025 at
<collectionCode id="ED7802C2FFCB43390CA8F8D8FCAB8017" box="[722,811,1886,1912]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="39" pageNumber="40">UNAM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCB43390EBDF804FE0B80AF" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8BD69A07FFCB43390EBDF804FE0B80AF" blockId="39.[151,1437,1490,2021]" pageId="39" pageNumber="40">
<emphasis id="B91D4615FFCB43390EBDF804FE4B80F3" bold="true" box="[199,459,1922,1948]" pageId="39" pageNumber="40">Differential diagnosis.</emphasis>
The character states to differentiate the monotypic genus
<taxonomicName id="4C69E184FFCB43390A45F804FB7D80F3" authorityName="Uusitalo" authorityYear="2025" box="[1087,1277,1922,1948]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCB43390A45F804FB7D80F3" box="[1087,1277,1922,1948]" italics="true" pageId="39" pageNumber="40">Glabromichaelia</emphasis>
</taxonomicName>
are applicable to the species as well.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCB43360EBDF84CFBDA8796" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="discussion">
<paragraph id="8BD69A07FFCB43360EBDF84CFBDA8796" blockId="39.[151,1437,1490,2021]" lastBlockId="40.[151,1436,151,357]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="B91D4615FFCB43390EBDF84CFEBB808B" bold="true" box="[199,315,1994,2020]" pageId="39" pageNumber="40">Remarks.</emphasis>
Few solenidia are added in the process of development from the protonymph to the adult form: one baculifom solenidion to tarsi II and 2 ceratiform solenidia to genua I. Structures like the sensilla, chelicerae, palps and segments of legs were more easily observable in the protonymph and accordingly used in the description and depicted as photos to maximize the correctness of observations of this unique species.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC443360EBDFE85FA14857B" pageId="40" pageNumber="41" type="etymology">
<paragraph id="8BD69A07FFC443360EBDFE85FEF3860A" blockId="40.[151,1436,151,357]" pageId="40" pageNumber="41">
<emphasis id="B91D4615FFC443360EBDFE85FECE8672" bold="true" box="[199,334,259,285]" pageId="40" pageNumber="41">Etymology.</emphasis>
The species name (a patronymic genitive) is given in honour of Ignacio Mauro Vázquez Rojas, Professor of Zoology at the National Autonomous University of
<collectingRegion id="49AD54E5FFC443360D11FEA1FC41862E" box="[875,961,295,321]" country="Mexico" name="Mexico" pageId="40" pageNumber="41">Mexico</collectingRegion>
(UNAM), and collector and photographer of this new species.
</paragraph>
<paragraph id="8BD69A07FFC443360EEDFE30FD8E86BF" blockId="40.[151,526,438,465]" box="[151,526,438,465]" pageId="40" pageNumber="41">
<emphasis id="B91D4615FFC443360EEDFE30FD8E86BF" bold="true" box="[151,526,438,465]" pageId="40" pageNumber="41">
Genus
<taxonomicName id="4C69E184FFC443360E94FE30FD8E86BF" authority="Thor, 1902" authorityName="Thor" authorityYear="1902" box="[238,526,438,465]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC443360E94FE30FE0186BE" bold="true" box="[238,385,438,465]" italics="true" pageId="40" pageNumber="41">Bimichaelia</emphasis>
<bibRefCitation id="EFF8E7F6FFC443360FF2FE31FD8E86BF" author="Thor, S." box="[392,526,438,465]" pageId="40" pageNumber="41" pagination="159 - 165" refId="ref48630" refString="Thor, S. (1902) Zur Systematik der Acaridenfauna Bdellidae Koch, 1842, Grube, 1859, Eupodidae Koch, 1842 und Cunaxidae Sig Thor, 1902. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft, Wien, 52, 159-165." type="journal article" year="1902">Thor, 1902</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFC443360EEDFE7AFA14857B" blockId="40.[151,1428,507,532]" box="[151,1428,507,532]" pageId="40" pageNumber="41">
<materialsCitation id="3B01905AFFC443360EEDFE7AFA10857B" box="[151,1424,507,532]" country="Italy" location="Mt. St. Augusta" municipality="Treviso" pageId="40" pageNumber="41" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D224A5FFC443360EEDFE7AFF49857B" box="[151,201,508,532]" pageId="40" pageNumber="41">Type</typeStatus>
species:
<taxonomicName id="4C69E184FFC443360F5CFE7DFD0F857C" authority="Berlese, 1884" authorityName="Berlese" authorityYear="1884" box="[294,655,507,531]" class="Arachnida" family="Freyanidae" genus="Michaelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="augustana">
<emphasis id="B91D4615FFC443360F5CFE7DFE7D857C" box="[294,509,507,531]" italics="true" pageId="40" pageNumber="41">Michaelia augustana</emphasis>
<bibRefCitation id="EFF8E7F6FFC443360C79FE7DFD0F857C" author="Berlese, A." box="[515,655,507,531]" pageId="40" pageNumber="41" pagination="1882 - 1903" refId="ref46407" refString="Berlese, A. (1884) Fasciculus (Vol.) 16. In: Acari, Myriapoda et Scorpiones hucuscue in Italia reperta. 101 Vols. 1882-1903. Sumptibus Auctoris, Patavii (Padova), no pagination." type="book chapter" year="1884">Berlese, 1884</bibRefCitation>
</taxonomicName>
from
<location id="8EB6CCDCFFC443360CB6FE7AFCEE857C" LSID="urn:lsid:plazi:treatment:03C02B11FFCD43360EEDFA43FBD68413:8EB6CCDCFFC443360CB6FE7AFCEE857C" box="[716,878,507,531]" country="Italy" municipality="Treviso" name="Mt. St. Augusta" pageId="40" pageNumber="41">Mt. St. Augusta</location>
,
<collectingMunicipality id="6BB2007DFFC443360D02FE7DFC47857C" box="[888,967,507,531]" pageId="40" pageNumber="41">Treviso</collectingMunicipality>
,
<collectingCountry id="F37EDA97FFC443360DABFE7DFB83857C" box="[977,1027,507,531]" name="Italy" pageId="40" pageNumber="41">Italy</collectingCountry>
; by original designation and monotypy
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFC443360EEDFDC7FBD68413" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="8BD69A07FFC443360EEDFDC7FBD68413" blockId="40.[151,1436,577,892]" pageId="40" pageNumber="41">
<emphasis id="B91D4615FFC443360EEDFDC7FE1E8534" bold="true" box="[151,414,577,603]" pageId="40" pageNumber="41">Differential diagnosis.</emphasis>
The primary pattern of
<taxonomicName id="4C69E184FFC443360CD0FDC7FCB18534" authorityName="Thor" authorityYear="1902" box="[682,817,577,603]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC443360CD0FDC7FCB18534" box="[682,817,577,603]" italics="true" pageId="40" pageNumber="41">Bimichaelia</emphasis>
</taxonomicName>
is of roundish to subpolygonal loops of large lamellae, and granular integument inside the loops (
<figureCitation id="13528682FFC443360C13FDE3FD5A8510" box="[617,730,613,639]" captionStart="FIGURE 23" captionStartId="41.[151,250,1863,1887]" captionTargetBox="[176,1412,153,1839]" captionTargetId="figure-15@41.[176,1412,153,1839]" captionTargetPageId="41" captionText="FIGURE 23. Bimichaelia pallida (Ewing). A—prodorsum, dorsal setae mostly with 3 clearly visible cilia; B—subcapitulum with 9 setae; C—dorsum with ca. 300 setae; D—original figure of Bimichaelia pallida (Ewing), dorsum with ca. 100 setae; E—genital valves of male with 1418 genital setae per valve; F—7 pairs of eugenital setae; G—anal area with 6 anal setae per valve. D—copied from Ewing (1913). 23AC, EG—from Oregon, Multinomah Co., Corbett; 23D—from Oregon, Benton Co., Corvallis, copied from Ewing (1913)." pageId="40" pageNumber="41">Figs. 23A</figureCitation>
,
<figureCitation id="13528682FFC443360C9EFDE3FCB785EF" box="[740,823,613,640]" captionStart="FIGURE 25" captionStartId="44.[151,250,1897,1921]" captionTargetBox="[218,1370,153,1874]" captionTargetId="figure-24@44.[218,1370,153,1874]" captionTargetPageId="44" captionText="FIGURE 25. Bimichaelia logani sp. nov. A—prodorsum, dorsal setae mostly with 56 clearly visible cilia; B—dorsum with ca. 300 setae; C—subcapitulum with 6 setae; D—genital valves of female with 14 genital setae per valve; E—anal area with 56 anal setae per valve. All specimens—from Utah, Cache Co." pageId="40" pageNumber="41">25A, B</figureCitation>
,
<figureCitation id="13528682FFC443360D3BFDE3FC138510" box="[833,915,613,640]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="40" pageNumber="41">52B, C</figureCitation>
). All dorsal setae of
<taxonomicName id="4C69E184FFC443360A0EFDE3FB7B8510" authorityName="Thor" authorityYear="1902" box="[1140,1275,613,639]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC443360A0EFDE3FB7B8510" box="[1140,1275,613,639]" italics="true" pageId="40" pageNumber="41">Bimichaelia</emphasis>
</taxonomicName>
have less than 10, mainly short barbs basally, which can be difficult to discern against the intricate cuticular pattern (
<figureCitation id="13528682FFC443360B55FD0FFF2C85A8" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="40" pageNumber="41">Fig. 52A, B</figureCitation>
), whereas the setae of
<taxonomicName id="4C69E184FFC443360FD0FD2BFDE885A8" authorityName="Uusitalo" authorityYear="2025" box="[426,616,685,711]" class="Arachnida" family="Alycidae" genus="Glabromichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D4615FFC443360FD0FD2BFDE885A8" box="[426,616,685,711]" italics="true" pageId="40" pageNumber="41">Glabromichaelia</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFC443360C0AFD2BFD5985A8" bold="true" box="[624,729,685,711]" pageId="40" pageNumber="41">
<taxonomicNameLabel id="A22EFB6EFFC443360C0AFD2BFD5985A8" box="[624,729,685,711]" pageId="40" pageNumber="41" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C69E184FFC443360C9DFD2BFC2D85A8" authorityName="Uusitalo" authorityYear="2025" box="[743,941,685,711]" class="Arachnida" family="Alycidae" genus="Minimamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D4615FFC443360C9DFD2BFC2D85A8" box="[743,941,685,711]" italics="true" pageId="40" pageNumber="41">Minimamichaelia</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFC443360DC9FD2BFB9C85A8" bold="true" box="[947,1052,685,711]" pageId="40" pageNumber="41">
<taxonomicNameLabel id="A22EFB6EFFC443360DC9FD2BFB9C85A8" box="[947,1052,685,711]" pageId="40" pageNumber="41" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C69E184FFC443360A50FD2BFB7485A8" authorityName="Uusitalo" authorityYear="2025" box="[1066,1268,685,711]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B91D4615FFC443360A50FD2BFB7485A8" box="[1066,1268,685,711]" italics="true" pageId="40" pageNumber="41">Quartusmichaelia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFC443360A81FD2BFAE585A7" box="[1275,1381,685,712]" pageId="40" pageNumber="41" rank="genus">
<emphasis id="B91D4615FFC443360A81FD2BFADF85A8" bold="true" box="[1275,1375,685,711]" pageId="40" pageNumber="41">gen. nov</emphasis>
.
</taxonomicNameLabel>
, and
<taxonomicName id="4C69E184FFC443360EEDFD57FEDB8584" authorityName="Uusitalo" authorityYear="2010" box="[151,347,721,747]" class="Arachnida" family="Alycidae" genus="Laminamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC443360EEDFD57FEDB8584" box="[151,347,721,747]" italics="true" pageId="40" pageNumber="41">Laminamichaelia</emphasis>
</taxonomicName>
(
<emphasis id="B91D4615FFC443360F16FD54FE798584" box="[364,505,722,747]" italics="true" pageId="40" pageNumber="41">sensu stricto</emphasis>
)
<bibRefCitation id="EFF8E7F6FFC443360C73FD57FD2E8584" author="Uusitalo, M." box="[521,686,721,747]" pageId="40" pageNumber="41" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo, 2010</bibRefCitation>
are easily discernible because of several elongated cilia per seta at least on the shoulders and middorsum (
<figureCitation id="13528682FFC443360C33FD73FC90847F" box="[585,784,757,784]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="40" pageNumber="41">Figs. 52E, G, I, K</figureCitation>
,
<figureCitation id="13528682FFC443360D60FD73FC658460" box="[794,997,757,784]" captionStart="FIGURE 53" captionStartId="85.[151,250,1681,1705]" captionTargetBox="[190,1397,153,1657]" captionTargetId="figure-15@85.[190,1397,153,1657]" captionTargetPageId="85" captionText="FIGURE 53. Genus-level character states of Bimichaeliini, part 2. AD—Quartusmichaelia shibai: ciliated setae on dorsum; prodorsum with six or seven pairs of setae; filamentous sensilla ve densely ciliated by cilia of various lengths from top to bottom; reticulate dorsal pattern; anterior solenidion ω1 hook-like on tarsus I, respectively; EH—Quartusmichaelia: dorsum with substellate polygons of primary pattern; setae lanceolate or ciliate; prodorsum with six pairs of setae or neotrichous rim of soft integument; lanceolate setae scaly or with tiny barbs; secondary pattern of even rows of small lamellae; tarsus I with two baculiform solenidia, anterior one being S-shaped or hook-like (and beak-like chelicera - typical of Bimichaeliini), respectively; IL—Laminamichaelia: dorsum with regular stellate pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with multiciliate setae; primary pattern polygonal; secondary pattern of even rows of small lamellae; three baculiform solenidia on tarsus I, respectively. 53AD—specimens from Veracruz, CICOLMA; 53EL—copied from Uusitalo (2010)." pageId="40" pageNumber="41">53A, C, E, G, I, K</figureCitation>
). Distal famulus I is abaxial to the basis of claws (44C) and difficult to see by light microscopy. Also, the prodorsum has only 6 pairs of setae (
<figureCitation id="13528682FFC443360B5FFC9FFA18845C" box="[1317,1432,793,819]" captionStart="FIGURE 23" captionStartId="41.[151,250,1863,1887]" captionTargetBox="[176,1412,153,1839]" captionTargetId="figure-15@41.[176,1412,153,1839]" captionTargetPageId="41" captionText="FIGURE 23. Bimichaelia pallida (Ewing). A—prodorsum, dorsal setae mostly with 3 clearly visible cilia; B—subcapitulum with 9 setae; C—dorsum with ca. 300 setae; D—original figure of Bimichaelia pallida (Ewing), dorsum with ca. 100 setae; E—genital valves of male with 1418 genital setae per valve; F—7 pairs of eugenital setae; G—anal area with 6 anal setae per valve. D—copied from Ewing (1913). 23AC, EG—from Oregon, Multinomah Co., Corbett; 23D—from Oregon, Benton Co., Corvallis, copied from Ewing (1913)." pageId="40" pageNumber="41">Figs. 23A</figureCitation>
,
<figureCitation id="13528682FFC443360EEDFCBBFF4A8437" box="[151,202,829,856]" captionStart="FIGURE 25" captionStartId="44.[151,250,1897,1921]" captionTargetBox="[218,1370,153,1874]" captionTargetId="figure-24@44.[218,1370,153,1874]" captionTargetPageId="44" captionText="FIGURE 25. Bimichaelia logani sp. nov. A—prodorsum, dorsal setae mostly with 56 clearly visible cilia; B—dorsum with ca. 300 setae; C—subcapitulum with 6 setae; D—genital valves of female with 14 genital setae per valve; E—anal area with 56 anal setae per valve. All specimens—from Utah, Cache Co." pageId="40" pageNumber="41">25A</figureCitation>
,
<figureCitation id="13528682FFC443360EADFCBBFE898437" box="[215,265,829,856]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="40" pageNumber="41">52B</figureCitation>
), i.e., there are no accessory setae on the prodorsum, a character state
<taxonomicName id="4C69E184FFC443360A53FCBBFB308438" authorityName="Thor" authorityYear="1902" box="[1065,1200,829,855]" class="Arachnida" family="Alycidae" genus="Bimichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC443360A53FCBBFB308438" box="[1065,1200,829,855]" italics="true" pageId="40" pageNumber="41">Bimichaelia</emphasis>
</taxonomicName>
shares only with the
<taxonomicName id="4C69E184FFC443360EEDFCE7FEDD8414" authorityName="Uusitalo" authorityYear="2025" box="[151,349,865,891]" class="Arachnida" family="Alycidae" genus="Minimamichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFC443360EEDFCE7FEDD8414" box="[151,349,865,891]" italics="true" pageId="40" pageNumber="41">Minimamichaelia</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFC443360F16FCE7FE588413" box="[364,472,865,892]" captionStart="FIGURE 52" captionStartId="84.[151,250,1683,1707]" captionTargetBox="[190,1397,153,1659]" captionTargetId="figure-24@84.[190,1397,153,1659]" captionTargetPageId="84" captionText="FIGURE 52. Genus-level character states of Bimichaeliini, part 1. AD—Bimichaelia: habitus with roundish areoles; prodorsum with six pairs of setae; integument of C-segment with reticular primary pattern (roundish areoles); granular secondary pattern; setae with a long cilium and a few barbs of which 23 elongated; and three baculiform solenidia on tarsus I, respectively; EH—Minimamichaelia: dorsal habitus without reticulate primary pattern; prodorsum with six pairs of setae; integument of caudal segments with granules, small lamellae and clumps of middlesized lamellae on undulating ridges; elongated setae with several cilia of various length; and two baculiform solenidia on tarsus of the first pair of legs, respectively; IL—Glabromichaelia: habitus without reticulate primary pattern; on prodorsum: both soft integument and in-area neotrichous; dorsal integument with elongated, multiciliate setae and solid, parallel ridges without granulae or lamellae; two baculiform solenidia on tarsus I, respectively.Abbreviations: Na—naso, C, D, E, F, H, PS—segments of opisthosoma. 52AH—copied from Uusitalo (2010); 52IL—specimens from Veracruz, CICOLMA." pageId="40" pageNumber="41">Figs. 52F</figureCitation>
) and
<taxonomicName id="4C69E184FFC443360C6DFCE7FC518414" authority="Uusitalo et al." authorityName="Uusitalo et al." authorityYear="2025" baseAuthorityName="Uusitalo, Ueckermann &amp; Theron" baseAuthorityYear="2020" box="[535,977,865,891]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="shibai">
<emphasis id="B91D4615FFC443360C6DFCE7FCAC8414" box="[535,812,865,891]" italics="true" pageId="40" pageNumber="41">Quartusmichaelia shibai</emphasis>
Uusitalo
<emphasis id="B91D4615FFC443360DE3FCE4FC518414" box="[921,977,865,891]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFC443360D9AFCE7FBC98413" box="[992,1097,865,892]" captionStart="FIGURE 35" captionStartId="58.[151,250,1926,1950]" captionTargetBox="[151,1435,232,1903]" captionTargetId="figure-36@58.[151,1435,232,1903]" captionTargetPageId="58" captionText="FIGURE 35. Quartusmichaelia shibai (Uusitalo, Ueckermann &amp; Theron). A—prodorsum with sensilla ve ciliated from top to bottom; B—dorsum; C—venter and gnathosoma; D—leg I; E—leg II. All specimens—from Veracruz, CICOLMA." pageId="40" pageNumber="41">Fig. 35A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,201 @@
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<mods:title id="00888DF2245268FE453E20875E960BC7">A review of the family Alycidae (Acari, Acariformes) from North America</mods:title>
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<paragraph id="8BD69A07FFDB43290EEDFB48FDC18386" blockId="55.[151,577,1230,1293]" box="[151,577,1230,1257]" pageId="55" pageNumber="56">
<heading id="D09E2D6BFFDB43290EEDFB48FDC18386" bold="true" box="[151,577,1230,1257]" fontSize="11" level="2" pageId="55" pageNumber="56" reason="4">
<emphasis id="B91D4615FFDB43290EEDFB48FDC18386" bold="true" box="[151,577,1230,1257]" pageId="55" pageNumber="56">
<taxonomicName id="4C69E184FFDB43290EEDFB48FE5C8386" authorityName="Uusitalo" authorityYear="2025" box="[151,476,1230,1257]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="plumata" status="sp. nov.">
<emphasis id="B91D4615FFDB43290EEDFB48FE5C8386" bold="true" box="[151,476,1230,1257]" italics="true" pageId="55" pageNumber="56">Quartusmichaelia plumata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFDB43290F9EFB49FDC18386" box="[484,577,1231,1257]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFDB43290EEDFB75FB5A818E" pageId="55" pageNumber="56" type="description">
<paragraph id="8BD69A07FFDB43290EEDFB75FE1A8262" blockId="55.[151,577,1230,1293]" box="[151,410,1267,1293]" pageId="55" pageNumber="56">
(
<figureCitation id="13528682FFDB43290EE5FB75FEB28262" box="[159,306,1267,1293]" captionStart="FIGURE 33" captionStartId="56.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-24@56.[151,1435,176,1849]" captionTargetPageId="56" captionText="FIGURE 33. Quartusmichaelia plumata sp. nov. A—prodorsum; B—lateral view of dorsum and legs; C—claw and empodium. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">Figs. 33AC</figureCitation>
,
<figureCitation id="13528682FFDB43290F44FB75FE118262" box="[318,401,1267,1293]" captionStart="FIGURE 34" captionStartId="57.[151,250,1729,1753]" captionTargetBox="[151,1436,176,1706]" captionTargetId="figure-15@57.[151,1436,176,1706]" captionTargetPageId="57" captionText="FIGURE 34. Quartusmichaelia plumata sp. nov. A—leg I; B—leg II; C—leg III; D—leg IV; E—palp and chelicera; F—subcapitulum; G—genital area with 8 genital setae on valve and protruding genital organ with 3 pairs of eugenital setae. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">34AG</figureCitation>
)
</paragraph>
<paragraph id="8BD69A07FFDB43290EEDFABCFCE58166" blockId="55.[151,1437,1338,2013]" pageId="55" pageNumber="56">
<emphasis id="B91D4615FFDB43290EEDFABCFEA9823B" bold="true" box="[151,297,1338,1364]" pageId="55" pageNumber="56">Description.</emphasis>
<emphasis id="B91D4615FFDB43290F4AFABDFE0A823B" box="[304,394,1339,1364]" italics="true" pageId="55" pageNumber="56">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFDB43290FC5FABDFDA3823A" box="[447,547,1339,1365]" count="1" pageId="55" pageNumber="56" type="female">1 female</specimenCount>
,
<figureCitation id="13528682FFDB43290C54FABDFD35823A" box="[558,693,1339,1365]" captionStart="FIGURE 33" captionStartId="56.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-24@56.[151,1435,176,1849]" captionTargetPageId="56" captionText="FIGURE 33. Quartusmichaelia plumata sp. nov. A—prodorsum; B—lateral view of dorsum and legs; C—claw and empodium. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">Fig. 33A, B</figureCitation>
). Length 250 µm; primary pattern comprising rosettes of roundish to subpolygonal cells around a seta; ca. 50 dorsal lanceolate setae; a seta with a short petiolus, elongated shaft from which the plump setules or bristles extend to the sides and upper surface, but leaving the lower surface bare, a few ciliate setae in caudal end; on prodorsum: 1012 lanceolate setae along anterior rim; fingertip-like naso; crista eminent with sidelines and reticulation; grooved sensilla
<emphasis id="B91D4615FFDB43290D62FA4DFCB1828B" box="[792,817,1483,1508]" italics="true" pageId="55" pageNumber="56">ve</emphasis>
with a distal bunch of short barbs resembling an ear of wheat; one pair of lanceolate setae
<emphasis id="B91D4615FFDB43290C5BFA69FDB78167" box="[545,567,1519,1544]" italics="true" pageId="55" pageNumber="56">in</emphasis>
on sclerotised integument.
</paragraph>
<paragraph id="8BD69A07FFDB43290EBDF995FAC48142" blockId="55.[151,1437,1338,2013]" box="[199,1348,1555,1581]" pageId="55" pageNumber="56">
<taxonomicName id="4C69E184FFDB43290EBDF995FE8F8143" box="[199,271,1555,1580]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="55" pageNumber="56" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFDB43290EBDF995FE8F8143" box="[199,271,1555,1580]" italics="true" pageId="55" pageNumber="56">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFDB43290F64F995FE0E8142" box="[286,398,1555,1581]" captionStart="FIGURE 33" captionStartId="56.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-24@56.[151,1435,176,1849]" captionTargetPageId="56" captionText="FIGURE 33. Quartusmichaelia plumata sp. nov. A—prodorsum; B—lateral view of dorsum and legs; C—claw and empodium. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">Figs. 33B</figureCitation>
,
<figureCitation id="13528682FFDB43290FE0F995FE698142" box="[410,489,1555,1581]" captionStart="FIGURE 34" captionStartId="57.[151,250,1729,1753]" captionTargetBox="[151,1436,176,1706]" captionTargetId="figure-15@57.[151,1436,176,1706]" captionTargetPageId="57" captionText="FIGURE 34. Quartusmichaelia plumata sp. nov. A—leg I; B—leg II; C—leg III; D—leg IV; E—palp and chelicera; F—subcapitulum; G—genital area with 8 genital setae on valve and protruding genital organ with 3 pairs of eugenital setae. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">34F, G</figureCitation>
). Setae sparsely ciliated; 810 genital setae per valve; 3 anal setae per valve.
</paragraph>
<paragraph id="8BD69A07FFDB43290EBDF9B1FBD1811A" blockId="55.[151,1437,1338,2013]" pageId="55" pageNumber="56">
<taxonomicName id="4C69E184FFDB43290EBDF9B1FED6813E" authorityName="Krefft" authorityYear="1871" box="[199,342,1591,1617]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="55" pageNumber="56" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFDB43290EBDF9B1FED6813E" box="[199,342,1591,1617]" italics="true" pageId="55" pageNumber="56">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFDB43290F1EF9B1FE53813E" box="[356,467,1591,1617]" captionStart="FIGURE 33" captionStartId="56.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-24@56.[151,1435,176,1849]" captionTargetPageId="56" captionText="FIGURE 33. Quartusmichaelia plumata sp. nov. A—prodorsum; B—lateral view of dorsum and legs; C—claw and empodium. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">Figs. 33B</figureCitation>
,
<figureCitation id="13528682FFDB43290FA4F9B1FDAB813E" box="[478,555,1591,1617]" captionStart="FIGURE 34" captionStartId="57.[151,250,1729,1753]" captionTargetBox="[151,1436,176,1706]" captionTargetId="figure-15@57.[151,1436,176,1706]" captionTargetPageId="57" captionText="FIGURE 34. Quartusmichaelia plumata sp. nov. A—leg I; B—leg II; C—leg III; D—leg IV; E—palp and chelicera; F—subcapitulum; G—genital area with 8 genital setae on valve and protruding genital organ with 3 pairs of eugenital setae. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">34E, F</figureCitation>
). Chelate chelicera stylet-like with relatively short shafts; 3 pairs of ciliate setae on subcapitulum; medially constricted palpal solenidion in contact with 2 eupathidia.
</paragraph>
<paragraph id="8BD69A07FFDB43290EBDF9F9FB5A818E" blockId="55.[151,1437,1338,2013]" pageId="55" pageNumber="56">
<emphasis id="B91D4615FFDB43290EBDF9F9FF7C81F7" box="[199,252,1663,1688]" italics="true" pageId="55" pageNumber="56">Legs</emphasis>
(
<figureCitation id="13528682FFDB43290F74F9F9FE2481F6" box="[270,420,1663,1689]" captionStart="FIGURE 33" captionStartId="56.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-24@56.[151,1435,176,1849]" captionTargetPageId="56" captionText="FIGURE 33. Quartusmichaelia plumata sp. nov. A—prodorsum; B—lateral view of dorsum and legs; C—claw and empodium. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">Figs. 33B, C</figureCitation>
,
<figureCitation id="13528682FFDB43290FC8F9F9FD8981F6" box="[434,521,1663,1689]" captionStart="FIGURE 34" captionStartId="57.[151,250,1729,1753]" captionTargetBox="[151,1436,176,1706]" captionTargetId="figure-15@57.[151,1436,176,1706]" captionTargetPageId="57" captionText="FIGURE 34. Quartusmichaelia plumata sp. nov. A—leg I; B—leg II; C—leg III; D—leg IV; E—palp and chelicera; F—subcapitulum; G—genital area with 8 genital setae on valve and protruding genital organ with 3 pairs of eugenital setae. All specimens—from Quintana Roo, Sian Kaan." pageId="55" pageNumber="56">34AD</figureCitation>
). Solenidial formula for tarsi, tibiae, genua and femora on legs I, II, III and IV, respectively: 2B-2B-0-0, 2B-1B-1B-1B, 3P-1B3P-1P-1P, 4P-1B1P-0-0, ω1 on tarsus I S-shaped and on tarsus II bent adaxially, other baculiform solenidia mostly straight forward, abaxial famulus I ventrodistal.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFDB43290EBDF96DFE38801E" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="8BD69A07FFDB43290EBDF96DFE38801E" blockId="55.[151,1437,1338,2013]" pageId="55" pageNumber="56">
<emphasis id="B91D4615FFDB43290EBDF96DFEF5806B" bold="true" box="[199,373,1770,1797]" pageId="55" pageNumber="56">Type material.</emphasis>
<materialsCitation id="3B01905AFFDB43290F06F96DFC0E8046" collectingDate="1995-09-05" country="Mexico" location="Sian Ka'an" pageId="55" pageNumber="56" specimenCount="1" stateProvince="Quintana Roo" typeStatus="holotype">
<typeStatus id="54D224A5FFDB43290F06F96DFE64806A" box="[380,484,1771,1797]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
female from soil, flooded lowland tropical forest (selva baja inundable),
<location id="8EB6CCDCFFDB43290B62F96DFA18806A" LSID="urn:lsid:plazi:treatment:03C02B11FFDB43270EEDFB48FEA280DD:8EB6CCDCFFDB43290B62F96DFA18806A" box="[1304,1432,1771,1797]" country="Mexico" name="Sian Ka'an" pageId="55" pageNumber="56" stateProvince="Quintana Roo">Sian Kaan</location>
,
<collectingRegion id="49AD54E5FFDB43290EEDF889FEB88046" box="[151,312,1807,1833]" country="Mexico" name="Quintana Roo" pageId="55" pageNumber="56">Quintana Roo</collectingRegion>
,
<collectingCountry id="F37EDA97FFDB43290F3DF889FE1F8046" box="[327,415,1807,1833]" name="Mexico" pageId="55" pageNumber="56">Mexico</collectingCountry>
,
<date id="FFD7BCC7FFDB43290FD7F888FDC98046" box="[429,585,1806,1833]" pageId="55" pageNumber="56" value="1995-09-05">
<collectingDate id="EF93452FFFDB43290FD7F888FDC98046" box="[429,585,1806,1833]" pageId="55" pageNumber="56" value="1995-09-05">5. Sept. 1995</collectingDate>
</date>
, María Magdalena Vázquez
</materialsCitation>
.
<materialsCitation id="3B01905AFFDB43290DE1F889FE34801E" collectionCode="LAAH, UNAM" collectorName="de Acarologia &amp; Anita Hoffmann &amp; Universidad Nacional Autonoma" country="Mexico" location="Faculty of Sciences" municipality="Laboratorio de Acarologia" pageId="55" pageNumber="56" specimenCount="1" stateProvince="Distrito Federal" typeStatus="holotype">
<typeStatus id="54D224A5FFDB43290DE1F889FB838046" box="[923,1027,1807,1833]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
(SK-37) is deposited in the
<collectionCode id="ED7802C2FFDB43290B34F889FA1C8046" box="[1358,1436,1807,1833]" pageId="55" pageNumber="56">LAAH</collectionCode>
(
<collectingMunicipality id="6BB2007DFFDB43290EE5F8B5FE4A8022" box="[159,458,1843,1869]" pageId="55" pageNumber="56">
Laboratorio
<collectorName id="269CFFD1FFDB43290F57F8B5FE4A8022" box="[301,458,1843,1869]" pageId="55" pageNumber="56">de Acarología</collectorName>
</collectingMunicipality>
<collectorName id="269CFFD1FFDB43290FA7F8B5FD208022" box="[477,672,1843,1869]" pageId="55" pageNumber="56">Anita Hoffmann</collectorName>
”),
<location id="8EB6CCDCFFDB43290CC4F8B5FC1C8022" LSID="urn:lsid:plazi:treatment:03C02B11FFDB43270EEDFB48FEA280DD:8EB6CCDCFFDB43290CC4F8B5FC1C8022" box="[702,924,1843,1869]" country="Mexico" municipality="Laboratorio de Acarologia" name="Faculty of Sciences" pageId="55" pageNumber="56" stateProvince="Distrito Federal">Faculty of Sciences</location>
at
<collectionCode id="ED7802C2FFDB43290DBBF8B5FB978022" box="[961,1047,1843,1869]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="55" pageNumber="56">UNAM</collectionCode>
(
<collectorName id="269CFFD1FFDB43290A5DF8B5FA1C8022" box="[1063,1436,1843,1869]" pageId="55" pageNumber="56">Universidad Nacional Autonóma</collectorName>
de
<collectingCountry id="F37EDA97FFDB43290EC3F8D1FE93801E" box="[185,275,1879,1905]" name="Mexico" pageId="55" pageNumber="56">México</collectingCountry>
),
<collectingRegion id="49AD54E5FFDB43290F5FF8D1FE34801E" box="[293,436,1879,1905]" country="Mexico" name="Distrito Federal" pageId="55" pageNumber="56">Mexico City</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFDB43290EBDF8FCFCE480B2" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="8BD69A07FFDB43290EBDF8FCFCE480B2" blockId="55.[151,1437,1338,2013]" pageId="55" pageNumber="56">
<emphasis id="B91D4615FFDB43290EBDF8FCFE4E80FB" bold="true" box="[199,462,1914,1940]" pageId="55" pageNumber="56">Differential diagnosis.</emphasis>
The new species can be differentiated from other species with lanceolate setae (i.e.
<taxonomicName id="4C69E184FFDB43290BFBF8FDFF7180D7" authorityName="Uusitalo" authorityYear="2025" baseAuthorityName="Berlese" baseAuthorityYear="1904" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="setigera">
<emphasis id="B91D4615FFDB43290BFBF8FDFF7180D7" italics="true" pageId="55" pageNumber="56">Q. setigera</emphasis>
</taxonomicName>
with simple sensilla
<emphasis id="B91D4615FFDB43290F9BF819FE7A80D7" box="[481,506,1951,1976]" italics="true" pageId="55" pageNumber="56">ve</emphasis>
and
<taxonomicName id="4C69E184FFDB43290C4BF819FD4A80D6" authorityName="Uusitalo" authorityYear="2025" box="[561,714,1951,1977]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="lanceolata">
<emphasis id="B91D4615FFDB43290C4BF819FD4A80D6" box="[561,714,1951,1977]" italics="true" pageId="55" pageNumber="56">Q. lanceolata</emphasis>
</taxonomicName>
with branched sensilla
<emphasis id="B91D4615FFDB43290DAFF819FC6E80D7" box="[981,1006,1951,1976]" italics="true" pageId="55" pageNumber="56">ve</emphasis>
) by having sensilla
<emphasis id="B91D4615FFDB43290AA8F819FB6B80D7" box="[1234,1259,1951,1976]" italics="true" pageId="55" pageNumber="56">ve</emphasis>
with a grooved stalk and a distal bunch of scales, and by the solenidial formula.
</paragraph>
</subSubSection>
<caption id="DF16CA8FFFD443260EEDF8D7FF5C80DE" pageId="56" pageNumber="57" startId="56.[151,250,1873,1897]" targetBox="[151,1435,176,1849]" targetPageId="56" targetType="figure">
<paragraph id="8BD69A07FFD443260EEDF8D7FD0D8006" blockId="56.[151,1436,1873,1969]" box="[151,653,1873,1897]" pageId="56" pageNumber="57">
<emphasis id="B91D4615FFD443260EEDF8D7FEA08006" bold="true" box="[151,288,1873,1897]" pageId="56" pageNumber="57">FIGURE 33.</emphasis>
<taxonomicName id="4C69E184FFD443260F5CF8D7FDB68006" authorityName="Uusitalo" authorityYear="2025" box="[294,566,1873,1897]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="plumata" status="sp. nov.">
<emphasis id="B91D4615FFD443260F5CF8D7FDB68006" box="[294,566,1873,1897]" italics="true" pageId="56" pageNumber="57">Quartusmichaelia plumata</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFD443260C47F8D7FD0D8006" bold="true" box="[573,653,1873,1897]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="A22EFB6EFFD443260C47F8D7FD0D8006" box="[573,653,1873,1897]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFD443260EEDF8F3FF5C80DE" blockId="56.[151,1436,1873,1969]" pageId="56" pageNumber="57">A—prodorsum; B—lateral view of dorsum and legs; C—claw and empodium. All specimens—from Quintana Roo, Sian Kaan.</paragraph>
</caption>
<caption id="DF16CA8FFFD543270EEDF947FAAA804D" pageId="57" pageNumber="58" startId="57.[151,250,1729,1753]" targetBox="[151,1436,176,1706]" targetPageId="57" targetType="figure">
<paragraph id="8BD69A07FFD543270EEDF947FD0D81B5" blockId="57.[151,1436,1729,1826]" box="[151,653,1729,1754]" pageId="57" pageNumber="58">
<emphasis id="B91D4615FFD543270EEDF947FEA081B6" bold="true" box="[151,288,1729,1753]" pageId="57" pageNumber="58">FIGURE 34.</emphasis>
<taxonomicName id="4C69E184FFD543270F5CF944FDB681B5" authorityName="Uusitalo" authorityYear="2025" box="[294,566,1730,1754]" class="Arachnida" family="Alycidae" genus="Quartusmichaelia" kingdom="Animalia" order="Sarcoptiformes" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="plumata" status="sp. nov.">
<emphasis id="B91D4615FFD543270F5CF944FDB681B5" box="[294,566,1730,1754]" italics="true" pageId="57" pageNumber="58">Quartusmichaelia plumata</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFD543270C47F944FD0D81B5" bold="true" box="[573,653,1730,1754]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="A22EFB6EFFD543270C47F944FD0D81B5" box="[573,653,1730,1754]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFD543270EEDF960FAAA804D" blockId="57.[151,1436,1729,1826]" pageId="57" pageNumber="58">A—leg I; B—leg II; C—leg III; D—leg IV; E—palp and chelicera; F—subcapitulum; G—genital area with 8 genital setae on valve and protruding genital organ with 3 pairs of eugenital setae. All specimens—from Quintana Roo, Sian Kaan.</paragraph>
</caption>
<subSubSection id="C373C98CFFD543270EBDF8F2FEA280DD" pageId="57" pageNumber="58" type="etymology">
<paragraph id="8BD69A07FFD543270EBDF8F2FEA280DD" blockId="57.[151,1436,1908,1970]" pageId="57" pageNumber="58">
<emphasis id="B91D4615FFD543270EBDF8F2FECE80E1" bold="true" box="[199,334,1908,1934]" pageId="57" pageNumber="58">Etymology.</emphasis>
The species name is a feminine gender of the Latin word
<emphasis id="B91D4615FFD543270D90F8F2FBD180E1" box="[1002,1105,1908,1934]" italics="true" pageId="57" pageNumber="58">plumatus</emphasis>
and refers to the feather-like dorsal setae.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,331 @@
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<mods:title id="9E89EDA6275537E2B940AE1DBC1408E5">A review of the family Alycidae (Acari, Acariformes) from North America</mods:title>
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<treatment id="03C02B11FFE743100EEDF9B0FBF0835A" ID-DOI="http://doi.org/10.5281/zenodo.15012423" ID-Zenodo-Dep="15012423" LSID="urn:lsid:plazi:treatment:03C02B11FFE743100EEDF9B0FBF0835A" httpUri="http://treatment.plazi.org/id/03C02B11FFE743100EEDF9B0FBF0835A" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C373C98CFFE743150EEDF9B0FE49813E" box="[151,457,1590,1617]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BD69A07FFE743150EEDF9B0FE49813E" blockId="11.[151,457,1590,1653]" box="[151,457,1590,1617]" pageId="11" pageNumber="12">
<heading id="D09E2D6BFFE743150EEDF9B0FE49813E" bold="true" box="[151,457,1590,1617]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="4">
<emphasis id="B91D4615FFE743150EEDF9B0FE49813E" bold="true" box="[151,457,1590,1617]" pageId="11" pageNumber="12">
<taxonomicName id="4C69E184FFE743150EEDF9B0FEE4813E" authorityName="Uusitalo" authorityYear="2025" box="[151,356,1590,1617]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="utahensis" status="sp. nov.">
<emphasis id="B91D4615FFE743150EEDF9B0FEE4813E" bold="true" box="[151,356,1590,1617]" italics="true" pageId="11" pageNumber="12">Alycus utahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFE743150F11F9B1FE49813E" box="[363,457,1591,1617]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFE743100EEDF9DDFBB187BA" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="description">
<paragraph id="8BD69A07FFE743150EEDF9DDFEF9811A" blockId="11.[151,457,1590,1653]" box="[151,377,1627,1653]" pageId="11" pageNumber="12">
(
<figureCitation id="13528682FFE743150EE5F9DDFEA2811A" box="[159,290,1627,1653]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="11" pageNumber="12">Figs. 3AE</figureCitation>
,
<figureCitation id="13528682FFE743150F54F9DDFEF2811A" box="[302,370,1627,1653]" captionStart="FIGURE 4" captionStartId="13.[151,250,1766,1790]" captionTargetBox="[151,1435,176,1742]" captionTargetId="figure-15@13.[151,1436,176,1742]" captionTargetPageId="13" captionText="FIGURE 4. Alycus utahensis sp. nov. A—Posterior (II) coxisternal plate, neotrichy; B—Leg I; C—Leg II; D—Tibia III, genu III; E—Tibia IV, genu IV. All specimens—from Utah, Summit Co." pageId="11" pageNumber="12">4AE</figureCitation>
)
</paragraph>
<paragraph id="8BD69A07FFE743150EEDF924FD3E80FA" blockId="11.[151,1437,1698,2013]" pageId="11" pageNumber="12">
<emphasis id="B91D4615FFE743150EEDF924FEA981D3" bold="true" box="[151,297,1698,1724]" pageId="11" pageNumber="12">Description.</emphasis>
<emphasis id="B91D4615FFE743150F4BF925FE0B81D3" box="[305,395,1699,1724]" italics="true" pageId="11" pageNumber="12">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFE743150FC5F925FDA081D2" box="[447,544,1699,1725]" count="1" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
,
<figureCitation id="13528682FFE743150C57F925FD3381D2" box="[557,691,1699,1725]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="11" pageNumber="12">Figs. 3A, B</figureCitation>
). Length 570 µm; parallel ridges on soft integument meandering, mostly continuous; strong neotrichy, ciliate dorsal setae with a long central point and 57 easily distinguishable cilia, caudal setae longest with long cilia resembling prodorsal setae
<emphasis id="B91D4615FFE743150DEBF96DFC27806B" box="[913,935,1771,1796]" italics="true" pageId="11" pageNumber="12">in</emphasis>
; on prodorsum: thin strip of hard integument connecting the eye area, setae
<emphasis id="B91D4615FFE743150F94F889FD958047" box="[494,533,1807,1832]" italics="true" pageId="11" pageNumber="12">exp</emphasis>
above reduced (lensless) pair of posterior eye areas and main body of prodorsal sclerite, narrow strip of hard integument continuous from proboscis-like naso up to sensillar area and covered by longitudinal lamellae of various size classes, setae
<emphasis id="B91D4615FFE743150CACF8D1FD6A801F" box="[726,746,1879,1904]" italics="true" pageId="11" pageNumber="12">vi</emphasis>
and
<emphasis id="B91D4615FFE743150D59F8D1FCC7801F" box="[803,839,1879,1904]" italics="true" pageId="11" pageNumber="12">sce</emphasis>
on separate microplates, setae
<emphasis id="B91D4615FFE743150ADDF8D1FB3D801F" box="[1191,1213,1879,1904]" italics="true" pageId="11" pageNumber="12">in</emphasis>
longest. (One extra seta inserted proximal to the right seta
<emphasis id="B91D4615FFE743150C30F8FDFDDE80FB" box="[586,606,1915,1940]" italics="true" pageId="11" pageNumber="12">vi</emphasis>
, arrow).
</paragraph>
<paragraph id="8BD69A07FFE743150EBDF819FDB580B2" blockId="11.[151,1437,1698,2013]" pageId="11" pageNumber="12">
<taxonomicName id="4C69E184FFE743150EBDF819FE8F80D7" box="[199,271,1951,1976]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFE743150EBDF819FE8F80D7" box="[199,271,1951,1976]" italics="true" pageId="11" pageNumber="12">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFE743150F64F819FE1F80D6" box="[286,415,1951,1977]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="11" pageNumber="12">Figs. 3D, E</figureCitation>
,
<figureCitation id="13528682FFE743150FD1F819FE5280D6" box="[427,466,1951,1977]" captionStart="FIGURE 4" captionStartId="13.[151,250,1766,1790]" captionTargetBox="[151,1435,176,1742]" captionTargetId="figure-15@13.[151,1436,176,1742]" captionTargetPageId="13" captionText="FIGURE 4. Alycus utahensis sp. nov. A—Posterior (II) coxisternal plate, neotrichy; B—Leg I; C—Leg II; D—Tibia III, genu III; E—Tibia IV, genu IV. All specimens—from Utah, Summit Co." pageId="11" pageNumber="12">4A</figureCitation>
). Genital setae 2931 per valve; one pair of eugenital setae (broken lines, arrows); anal setae 7 per valve; 7 setae on coxa IV.
</paragraph>
<caption id="DF16CA8FFFE043120EEDF8D4FBC08095" pageId="12" pageNumber="13" startId="12.[151,250,1874,1898]" targetBox="[151,1436,176,1851]" targetPageId="12" targetType="figure">
<paragraph id="8BD69A07FFE043120EEDF8D4FD9A8005" blockId="12.[151,1437,1874,2042]" box="[151,538,1874,1898]" pageId="12" pageNumber="13">
<emphasis id="B91D4615FFE043120EEDF8D4FE938005" bold="true" box="[151,275,1874,1898]" pageId="12" pageNumber="13">FIGURE 3.</emphasis>
<taxonomicName id="4C69E184FFE043120F63F8D4FE438005" authorityName="Uusitalo" authorityYear="2025" box="[281,451,1874,1898]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="utahensis" status="sp. nov.">
<emphasis id="B91D4615FFE043120F63F8D4FE438005" box="[281,451,1874,1898]" italics="true" pageId="12" pageNumber="13">Alycus utahensis</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFE043120FB3F8D4FD9A8005" bold="true" box="[457,538,1874,1898]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A22EFB6EFFE043120FB3F8D4FD9A8005" box="[457,538,1874,1898]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFE043120EEDF8F0FBC08095" blockId="12.[151,1437,1874,2042]" pageId="12" pageNumber="13">
A—Prodorsum, pattern of continuous ridges on C-segment, setae
<emphasis id="B91D4615FFE043120D61F8F1FCAF80E1" box="[795,815,1911,1934]" italics="true" pageId="12" pageNumber="13">in</emphasis>
with elongated cilia, one extra seta (arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co.
</paragraph>
</caption>
<caption id="DF16CA8FFFE143130EEDF960FD8B8029" pageId="13" pageNumber="14" startId="13.[151,250,1766,1790]" targetBox="[151,1435,176,1742]" targetPageId="13" targetType="figure">
<paragraph id="8BD69A07FFE143130EEDF960FD9A8191" blockId="13.[151,1436,1766,1862]" box="[151,538,1766,1790]" pageId="13" pageNumber="14">
<emphasis id="B91D4615FFE143130EEDF960FE938191" bold="true" box="[151,275,1766,1790]" pageId="13" pageNumber="14">FIGURE 4.</emphasis>
<taxonomicName id="4C69E184FFE143130F63F960FE438191" authorityName="Uusitalo" authorityYear="2025" box="[281,451,1766,1790]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="utahensis" status="sp. nov.">
<emphasis id="B91D4615FFE143130F63F960FE438191" box="[281,451,1766,1790]" italics="true" pageId="13" pageNumber="14">Alycus utahensis</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFE143130FB3F960FD9A8191" bold="true" box="[457,538,1766,1790]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A22EFB6EFFE143130FB3F960FD9A8191" box="[457,538,1766,1790]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFE143130EEDF88CFD8B8029" blockId="13.[151,1436,1766,1862]" pageId="13" pageNumber="14">A—Posterior (II) coxisternal plate, neotrichy; B—Leg I; C—Leg II; D—Tibia III, genu III; E—Tibia IV, genu IV. All specimens—from Utah, Summit Co.</paragraph>
</caption>
<paragraph id="8BD69A07FFE143130EBDF81EFF768095" blockId="13.[151,1437,1944,2042]" pageId="13" pageNumber="14">
<taxonomicName id="4C69E184FFE143130EBDF81EFED680DD" authorityName="Krefft" authorityYear="1871" box="[199,342,1944,1970]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="13" pageNumber="14" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFE143130EBDF81EFED680DD" box="[199,342,1944,1970]" italics="true" pageId="13" pageNumber="14">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFE143130F1CF81EFE3E80DD" box="[358,446,1944,1970]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="13" pageNumber="14">Fig. 3C</figureCitation>
). Rutella with broad shaft, three ventrodistal projections, and an apical point; three pairs of adoral setae on subcapitulum; one pair of dorsal cheliceral setae. Three pairs of pseudacanthoid palpal eupathidia apically.
</paragraph>
<paragraph id="8BD69A07FFE243100EBDFF11FBB187BA" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="15">
<emphasis id="B91D4615FFE243100EBDFF11FF7C87DF" box="[199,252,151,176]" italics="true" pageId="14" pageNumber="15">Legs</emphasis>
(
<figureCitation id="13528682FFE243100F76FF11FE1187DE" box="[268,401,151,177]" captionStart="FIGURE 4" captionStartId="13.[151,250,1766,1790]" captionTargetBox="[151,1435,176,1742]" captionTargetId="figure-15@13.[151,1436,176,1742]" captionTargetPageId="13" captionText="FIGURE 4. Alycus utahensis sp. nov. A—Posterior (II) coxisternal plate, neotrichy; B—Leg I; C—Leg II; D—Tibia III, genu III; E—Tibia IV, genu IV. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Figs. 4BE</figureCitation>
). Solenidial formula for tarsi, tibiae, genua and femora of legs I, II, III and IV, respectively: 2-1-0-0, 1-2-2-0, 4-2-2-1, 2-0-0-0, tarsi I and II with laterodistal peg-like famulus.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFE243100EBDFF59FE17860A" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BD69A07FFE243100EBDFF59FE17860A" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="15">
<emphasis id="B91D4615FFE243100EBDFF59FEF68797" bold="true" box="[199,374,222,249]" pageId="14" pageNumber="15">Type material.</emphasis>
<materialsCitation id="3B01905AFFE243100F07FF59FE17860A" collectingDate="1970-08-26" collectorName="G. F. Knowlton &amp; W. J. Hanson &amp; Deposited &amp; Acarology Laboratory &amp; University. The &amp; Glyptal" country="United States of America" county="Summit Co." location="Summit Co." pageId="14" pageNumber="15" specimenCount="2" stateProvince="Utah" typeStatus="holotype">
<typeStatus id="54D224A5FFE243100F07FF59FE658796" box="[381,485,223,249]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
female from
<taxonomicName id="4C69E184FFE243100C05FF59FD3D8796" box="[639,701,223,249]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFE243100C05FF59FD3D8796" box="[639,701,223,249]" italics="true" pageId="14" pageNumber="15">Abies</emphasis>
</taxonomicName>
duff, Hayden Fork (High Uintah Mtn),
<collectingCounty id="62B7E28BFFE243100AFBFF59FA8C8796" box="[1153,1292,223,249]" pageId="14" pageNumber="15">Summit Co.</collectingCounty>
,
<collectingRegion id="49AD54E5FFE243100B62FF59FAD18796" box="[1304,1361,223,249]" country="United States of America" name="Utah" pageId="14" pageNumber="15">Utah</collectingRegion>
,
<collectingCountry id="F37EDA97FFE243100B27FF59FA188796" box="[1373,1432,223,249]" name="United States of America" pageId="14" pageNumber="15">USA</collectingCountry>
,
<date id="FFD7BCC7FFE243100EEDFE85FECD8672" box="[151,333,259,285]" pageId="14" pageNumber="15" value="1970-08-26">
<collectingDate id="EF93452FFFE243100EEDFE85FECD8672" box="[151,333,259,285]" pageId="14" pageNumber="15" value="1970-08-26">26 August 1970</collectingDate>
</date>
,
<collectorName id="269CFFD1FFE243100F23FE85FD808672" box="[345,512,259,285]" pageId="14" pageNumber="15">G.F. Knowlton</collectorName>
&amp;
<collectorName id="269CFFD1FFE243100C5EFE85FD378672" box="[548,695,259,285]" pageId="14" pageNumber="15">W.J. Hanson</collectorName>
:
<typeStatus id="54D224A5FFE243100CB8FE85FCA48672" box="[706,804,259,285]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
on slide NA1, collection number AL0709.
<collectorName id="269CFFD1FFE243100B75FE85FA018672" box="[1295,1409,259,285]" pageId="14" pageNumber="15">Deposited</collectorName>
at the
<collectorName id="269CFFD1FFE243100EBAFEA1FE3D862E" box="[192,445,295,321]" pageId="14" pageNumber="15">Acarology Laboratory</collectorName>
,
<collectingRegion id="49AD54E5FFE243100FB3FEA1FDC2862E" box="[457,578,295,321]" country="United States of America" name="Ohio" pageId="14" pageNumber="15">Ohio State</collectingRegion>
<collectorName id="269CFFD1FFE243100C30FEA1FD7A862E" box="[586,762,295,321]" pageId="14" pageNumber="15">University. The</collectorName>
specimen is dorsal side up, sealed in
<collectorName id="269CFFD1FFE243100AD0FEA1FB7E862E" box="[1194,1278,295,321]" pageId="14" pageNumber="15">Glyptal</collectorName>
and under
<quantity id="4C9137E2FFE243100BFAFEA1FF43860A" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="mm" value="20.0">20 mm</quantity>
round cover glass.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFE243100EBDFEE8FC39847A" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BD69A07FFE243100EBDFEE8FC39847A" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="15">
<emphasis id="B91D4615FFE243100EBDFEE8FE4E86E7" bold="true" box="[199,462,366,392]" pageId="14" pageNumber="15">Differential diagnosis.</emphasis>
Superficially the large (570 µm in length) American species
<taxonomicName id="4C69E184FFE243100A05FEE9FABB86E6" authorityName="Uusitalo" authorityYear="2025" box="[1151,1339,367,393]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="utahensis" status="sp. nov.">
<emphasis id="B91D4615FFE243100A05FEE9FABB86E6" box="[1151,1339,367,393]" italics="true" pageId="14" pageNumber="15">Alycus utahensis</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFE243100B38FEE9FA1C86E6" bold="true" box="[1346,1436,367,393]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A22EFB6EFFE243100B38FEE9FA1C86E6" box="[1346,1436,367,393]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles the generally smaller European
<taxonomicName id="4C69E184FFE243100C00FE15FCDB86C2" authority="Koch" authorityName="Koch" box="[634,859,403,429]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="roseus">
<emphasis id="B91D4615FFE243100C00FE15FC9586C3" box="[634,789,403,429]" italics="true" pageId="14" pageNumber="15">Alycus roseus</emphasis>
Koch
</taxonomicName>
(vs. 350560 µm), redescribed by
<bibRefCitation id="EFF8E7F6FFE243100A96FE15FF6086BE" author="Uusitalo, M." pageId="14" pageNumber="15" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo (2010: 4045</bibRefCitation>
, figs. 824), but there are qualitative and quantitative differences which justify its status of a new species. The most obvious differences are as follows: (1) neotrichy on dorsal side is ca. 30 % stronger (
<figureCitation id="13528682FFE243100AC4FE5DFA9D869A" box="[1214,1309,475,501]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Figs. 3B</figureCitation>
vs. 1B and van der
<bibRefCitation id="EFF8E7F6FFE243100E89FE79FE258576" author="Hammen, L. van der" box="[243,421,511,537]" pageId="14" pageNumber="15" pagination="127 - 139" refId="ref47262" refString="Hammen, L. van der (1989) 3.1. The morphology of Alycus roseus C. L. Koch. In: Hammen, L. van der, An introduction to Comparative Arachnology. SPB Academic Publishing bv, The Hague, pp. 127-139." type="book chapter" year="1989">Hammen, 1989</bibRefCitation>
: fig. 48); (2) the ciliated setae with elongated shafts on segments H, PS and AD have two cilia forked at the tip of the shaft and as long as the shaft (
<figureCitation id="13528682FFE243100D58FDA5FCF88552" box="[802,888,547,573]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Fig. 3B</figureCitation>
) vs. cilia on caudal setae of
<taxonomicName id="4C69E184FFE243100ACDFDA5FA9E8553" box="[1207,1310,547,573]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="roseus">
<emphasis id="B91D4615FFE243100ACDFDA5FA9E8553" box="[1207,1310,547,573]" italics="true" pageId="14" pageNumber="15">A. roseus</emphasis>
</taxonomicName>
are shorter than the shaft which is long or terminates in a single cilium in
<figureCitation id="13528682FFE243100D3EFDC1FC19850E" box="[836,921,583,609]" captionStart="FIGURE 1" captionStartId="9.[151,250,1838,1862]" captionTargetBox="[190,1397,153,1822]" captionTargetId="figure-15@9.[190,1397,153,1822]" captionTargetPageId="9" captionText="FIGURE 1. Alycus roseus C.L. Koch. A—prodorsum, pattern of fragmentary ridges on C-segment, setae in with short cilia; B—dorsal habitus, setae on caudal segments with elongated shaft and short cilia; C—subcapitulum slightly neotrichous, 3 pairs of adorals, rutella, chelicera robust, smooth cheliceral seta, palp tarsus with 3 pairs of pseudacanthoid eupathidia; D—genital valves of female with 2324 genital setae per valve, 1 pair of eugenital setae beneath valves (by broken lines, indicated by arrows,); E—anal area, 56 anal setae per valve. All specimens—from Alberta, Lamont Co." pageId="14" pageNumber="15">Fig. 1B</figureCitation>
; (3) the numbers of genital setae (
<figureCitation id="13528682FFE243100B75FDC1FAE5850E" box="[1295,1381,583,609]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Fig. 3D</figureCitation>
) and setae on coxal field IV (
<figureCitation id="13528682FFE243100FDAFDEDFE7785EA" box="[416,503,619,645]" captionStart="FIGURE 4" captionStartId="13.[151,250,1766,1790]" captionTargetBox="[151,1435,176,1742]" captionTargetId="figure-15@13.[151,1436,176,1742]" captionTargetPageId="13" captionText="FIGURE 4. Alycus utahensis sp. nov. A—Posterior (II) coxisternal plate, neotrichy; B—Leg I; C—Leg II; D—Tibia III, genu III; E—Tibia IV, genu IV. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Fig. 4A</figureCitation>
) are 2931 and 57, respectively (vs. 1824 and
<quantity id="4C9137E2FFE243100A62FDEDFBDE85EA" box="[1048,1118,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="14" pageNumber="15" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">12 in</quantity>
<figureCitation id="13528682FFE243100A1EFDEDFB4585EA" box="[1124,1221,619,645]" captionStart="FIGURE 1" captionStartId="9.[151,250,1838,1862]" captionTargetBox="[190,1397,153,1822]" captionTargetId="figure-15@9.[190,1397,153,1822]" captionTargetPageId="9" captionText="FIGURE 1. Alycus roseus C.L. Koch. A—prodorsum, pattern of fragmentary ridges on C-segment, setae in with short cilia; B—dorsal habitus, setae on caudal segments with elongated shaft and short cilia; C—subcapitulum slightly neotrichous, 3 pairs of adorals, rutella, chelicera robust, smooth cheliceral seta, palp tarsus with 3 pairs of pseudacanthoid eupathidia; D—genital valves of female with 2324 genital setae per valve, 1 pair of eugenital setae beneath valves (by broken lines, indicated by arrows,); E—anal area, 56 anal setae per valve. All specimens—from Alberta, Lamont Co." pageId="14" pageNumber="15">Figs. 1D</figureCitation>
,
<figureCitation id="13528682FFE243100AABFDEDFB7685EA" box="[1233,1270,619,645]" captionStart="FIGURE 2" captionStartId="10.[151,250,1938,1962]" captionTargetBox="[160,1427,176,1914]" captionTargetId="figure-24@10.[160,1427,176,1914]" captionTargetPageId="10" captionText="FIGURE 2. Alycus roseus C.L. Koch. A—posterior (II) coxisternal plate, slight neotrichy; B—leg I; C—leg II; D—tibia and genu III; E—tibia IV and genu IV. 1 All specimens—from Alberta, Lamont Co." pageId="14" pageNumber="15">2A</figureCitation>
, and in Europe acc.
<bibRefCitation id="EFF8E7F6FFE243100EB0FD09FE3185C6" author="Grandjean, F." box="[202,433,655,681]" pageId="14" pageNumber="15" pagination="56 - 61" refId="ref47047" refString="Grandjean, F. (1937) Le genre Pachygnathus Duges (Alycus Koch) (Acariens). Deuxieme partie. Bulletin Museum national d'Histoire naturelle, Paris, 2 e Serie, 9, 56-61." type="journal article" year="1937">Grandjean, 1937: 60</bibRefCitation>
; van der
<bibRefCitation id="EFF8E7F6FFE243100C6EFD09FD4485C6" author="Hammen, L. van der" box="[532,708,655,681]" pageId="14" pageNumber="15" pagination="127 - 139" refId="ref47262" refString="Hammen, L. van der (1989) 3.1. The morphology of Alycus roseus C. L. Koch. In: Hammen, L. van der, An introduction to Comparative Arachnology. SPB Academic Publishing bv, The Hague, pp. 127-139." type="book chapter" year="1989">Hammen, 1989</bibRefCitation>
: fig. 47); (4) the ridges on dorsal integument are more continuous (
<figureCitation id="13528682FFE243100EE5FD35FE9A85A2" box="[159,282,691,717]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Fig. 3A, D</figureCitation>
) vs. fragmentary (
<figureCitation id="13528682FFE243100F90FD35FDBF85A2" box="[490,575,691,717]" captionStart="FIGURE 1" captionStartId="9.[151,250,1838,1862]" captionTargetBox="[190,1397,153,1822]" captionTargetId="figure-15@9.[190,1397,153,1822]" captionTargetPageId="9" captionText="FIGURE 1. Alycus roseus C.L. Koch. A—prodorsum, pattern of fragmentary ridges on C-segment, setae in with short cilia; B—dorsal habitus, setae on caudal segments with elongated shaft and short cilia; C—subcapitulum slightly neotrichous, 3 pairs of adorals, rutella, chelicera robust, smooth cheliceral seta, palp tarsus with 3 pairs of pseudacanthoid eupathidia; D—genital valves of female with 2324 genital setae per valve, 1 pair of eugenital setae beneath valves (by broken lines, indicated by arrows,); E—anal area, 56 anal setae per valve. All specimens—from Alberta, Lamont Co." pageId="14" pageNumber="15">Fig. 1A</figureCitation>
and
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: figs. 810); (5) length of the prodorsal setae
<emphasis id="B91D4615FFE243100B66FD35FAB285A3" box="[1308,1330,691,716]" italics="true" pageId="14" pageNumber="15">in</emphasis>
is 40 µm (
<figureCitation id="13528682FFE243100EE5FD51FF77859E" box="[159,247,727,753]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Fig. 3A</figureCitation>
) vs. ca. 2025 µm for
<taxonomicName id="4C69E184FFE243100F89FD51FDDA859F" box="[499,602,727,753]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="roseus">
<emphasis id="B91D4615FFE243100F89FD51FDDA859F" box="[499,602,727,753]" italics="true" pageId="14" pageNumber="15">A. roseus</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFE243100C13FD51FD3D859E" box="[617,701,727,753]" captionStart="FIGURE 1" captionStartId="9.[151,250,1838,1862]" captionTargetBox="[190,1397,153,1822]" captionTargetId="figure-15@9.[190,1397,153,1822]" captionTargetPageId="9" captionText="FIGURE 1. Alycus roseus C.L. Koch. A—prodorsum, pattern of fragmentary ridges on C-segment, setae in with short cilia; B—dorsal habitus, setae on caudal segments with elongated shaft and short cilia; C—subcapitulum slightly neotrichous, 3 pairs of adorals, rutella, chelicera robust, smooth cheliceral seta, palp tarsus with 3 pairs of pseudacanthoid eupathidia; D—genital valves of female with 2324 genital setae per valve, 1 pair of eugenital setae beneath valves (by broken lines, indicated by arrows,); E—anal area, 56 anal setae per valve. All specimens—from Alberta, Lamont Co." pageId="14" pageNumber="15">Fig. 1A</figureCitation>
and
<bibRefCitation id="EFF8E7F6FFE243100C89FD51FC15859E" author="Uusitalo, M." box="[755,917,727,753]" pageId="14" pageNumber="15" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: fig 8); and (6) there is no solenidion on femur IV while
<taxonomicName id="4C69E184FFE243100F7AFD7DFEE8847B" box="[256,360,763,789]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="roseus">
<emphasis id="B91D4615FFE243100F7AFD7DFEE8847B" box="[256,360,763,789]" italics="true" pageId="14" pageNumber="15">A. roseus</emphasis>
</taxonomicName>
has one (
<figureCitation id="13528682FFE243100FA9FD7DFDB1847A" box="[467,561,763,789]" captionStart="FIGURE 4" captionStartId="13.[151,250,1766,1790]" captionTargetBox="[151,1435,176,1742]" captionTargetId="figure-15@13.[151,1436,176,1742]" captionTargetPageId="13" captionText="FIGURE 4. Alycus utahensis sp. nov. A—Posterior (II) coxisternal plate, neotrichy; B—Leg I; C—Leg II; D—Tibia III, genu III; E—Tibia IV, genu IV. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Figs. 4E</figureCitation>
vs. 2E and
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: fig. 11).
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFE243100EBDFC99FDFC837E" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BD69A07FFE243100EBDFC99FDFC837E" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="15">
<emphasis id="B91D4615FFE243100EBDFC99FEBB8456" bold="true" box="[199,315,799,825]" pageId="14" pageNumber="15">Remarks.</emphasis>
<taxonomicName id="4C69E184FFE243100F3FFC99FD378456" authority="C. L. Koch, 1842" authorityName="C. L. Koch" authorityYear="1842" box="[325,695,799,825]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="roseus">
<emphasis id="B91D4615FFE243100F3FFC99FE618457" box="[325,481,799,825]" italics="true" pageId="14" pageNumber="15">Alycus roseus</emphasis>
C. L.
<bibRefCitation id="EFF8E7F6FFE243100C55FC99FD378456" author="Koch, C. L." box="[559,695,799,825]" pageId="14" pageNumber="15" pagination="1837 - 1850" refId="ref47792" refString="Koch, C. L. (1842) 3. Heft (Vol.). In: Ubersicht des Arachniden systems. 5 Vols. 1837-1850. C. H. Zeh'schen Buchhandlung. Nurnberg, pp. 1 - 131, 13 pls." type="book chapter" year="1842">Koch, 1842</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C69E184FFE243100CBFFC99FCE48456" authorityName="Duges" authorityYear="1834" box="[709,868,799,825]" class="Arachnida" family="Alycidae" genus="Pachygnathus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFE243100CBFFC99FCE48456" box="[709,868,799,825]" italics="true" pageId="14" pageNumber="15">Pachygnathus</emphasis>
</taxonomicName>
nr.
<taxonomicName id="4C69E184FFE243100DEEFC99FC5D8456" authorityName="Grandjean" authorityYear="1942" box="[916,989,799,825]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dugesi">
<emphasis id="B91D4615FFE243100DEEFC99FC5D8456" box="[916,989,799,825]" italics="true" pageId="14" pageNumber="15">dugesi</emphasis>
</taxonomicName>
or undescribed species of
<taxonomicName id="4C69E184FFE243100B69FC99FADE8456" authorityName="C. L. Koch" authorityYear="1842" box="[1299,1374,799,825]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFE243100B69FC99FADE8456" box="[1299,1374,799,825]" italics="true" pageId="14" pageNumber="15">Alycus</emphasis>
</taxonomicName>
have been reported from
<collectingRegion id="49AD54E5FFE243100FFAFCC5FE578432" box="[384,471,835,861]" country="United States of America" name="Oregon" pageId="14" pageNumber="15">Oregon</collectingRegion>
,
<collectingRegion id="49AD54E5FFE243100F9CFCC5FDFE8432" box="[486,638,835,861]" country="United States of America" name="New Mexico" pageId="14" pageNumber="15">New Mexico</collectingRegion>
,
<collectingRegion id="49AD54E5FFE243100CF7FCC5FD7E8432" box="[653,766,835,861]" country="United States of America" name="Wyoming" pageId="14" pageNumber="15">Wyoming</collectingRegion>
,
<collectingRegion id="49AD54E5FFE243100D77FCC5FCE18432" box="[781,865,835,861]" country="United States of America" name="Florida" pageId="14" pageNumber="15">Florida</collectingRegion>
,
<collectingRegion id="49AD54E5FFE243100D0AFCC5FC428432" box="[880,962,835,861]" country="United States of America" name="Illinois" pageId="14" pageNumber="15">Illinois</collectingRegion>
,
<collectingRegion id="49AD54E5FFE243100DABFCC5FBC28432" box="[977,1090,835,861]" country="United States of America" name="Louisiana" pageId="14" pageNumber="15">Louisiana</collectingRegion>
,
<collectingRegion id="49AD54E5FFE243100A28FCC5FB3F8432" box="[1106,1215,835,861]" country="United States of America" name="Nebraska" pageId="14" pageNumber="15">Nebraska</collectingRegion>
, and
<collectingRegion id="49AD54E5FFE243100B7AFCC5FAF58432" box="[1280,1397,835,861]" country="United States of America" name="Wisconsin" pageId="14" pageNumber="15">Wisconsin</collectingRegion>
by
<bibRefCitation id="EFF8E7F6FFE243100EEDFCE1FEF484EE" author="Kamill, B. W. &amp; Steinberger, Y. &amp; Whitford, W. G." box="[151,372,870,897]" pageId="14" pageNumber="15" pagination="273 - 286" refId="ref47471" refString="Kamill, B. W., Steinberger, Y. &amp; Whitford, W. G. (1985) Soil microarthropods from the Chihuahuan Desert of New Mexico. Journal of Zoology, London, Series A, 205, 273-286. https://doi.org/10.1111/j.1469-7998.1985.tb03534.x" type="journal article" year="1985">
Kamill
<emphasis id="B91D4615FFE243100E91FCE1FEA284EE" box="[235,290,871,897]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
(1985)
</bibRefCitation>
,
<bibRefCitation id="EFF8E7F6FFE243100F04FCE1FD9284EE" author="Walter, D. E." box="[382,530,871,897]" pageId="14" pageNumber="15" pagination="271 - 290" refId="ref49287" refString="Walter, D. E. (1987) Belowground arthropods of semiarid grasslands. In: Capinera, J. L. (Ed.), Integrated Pest Management on Rangeland: A Shortgrass Prairie Perspective. Westview Press, Boulder, Colorado, pp. 271-290. https://doi.org/10.1201/9780429043673-19" type="book chapter" year="1987">Walter (1987</bibRefCitation>
,
<bibRefCitation id="EFF8E7F6FFE243100C66FCE1FDD584EE" author="Walter, D. E." box="[540,597,871,897]" pageId="14" pageNumber="15" pagination="159 - 166" refId="ref49351" refString="Walter, D. E. (1988) Predation and mycophagy by endeostigmatid mites (Acariformes: Prostigmata). Experimental and Applied Acarology, 4, 159-166. https://doi.org/10.1007/BF01193873" type="journal article" year="1988">1988</bibRefCitation>
) and
<bibRefCitation id="EFF8E7F6FFE243100CEBFCE1FCC184EE" author="Kethley, J. B." box="[657,833,871,897]" pageId="14" pageNumber="15" pagination="1058 - 1061" refId="ref47633" refString="Kethley, J. B. (1990 b) The prelarva of Alycus roseus Koch (Bimichaeliidae: Acariformes: Acari). Canadian Journal of Zoology, 68, 1058-1061. https://doi.org/10.1139/z90-154" type="journal article" year="1990">
Kethley (1990
<bibRefCitation id="EFF8E7F6FFE243100D48FCE1FCC184EE" author="Kethley, J. B." box="[818,833,871,897]" pageId="14" pageNumber="15" pagination="130 - 135" refId="ref47681" refString="Kethley, J. B. (1991) Probable tritonymphal sexual dimorphism in Alycus roseus Koch (Bimichaeliidae: Acariformes: Acari). Journal of Entomological Science, 26, 130-135. https://doi.org/10.18474/0749-8004-26.1.130" type="journal article" year="1991">b</bibRefCitation>
</bibRefCitation>
, 1991).Also unconfirmed are records of
<taxonomicName id="4C69E184FFE243100B7EFCE1FA1C84EF" box="[1284,1436,871,897]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="roseus">
<emphasis id="B91D4615FFE243100B7EFCE1FA1C84EF" box="[1284,1436,871,897]" italics="true" pageId="14" pageNumber="15">Alycus roseus</emphasis>
</taxonomicName>
from native grasslands in central
<collectingCountry id="F37EDA97FFE243100C68FC0DFDC884CA" box="[530,584,907,933]" name="United States of America" pageId="14" pageNumber="15">USA</collectingCountry>
(
<bibRefCitation id="EFF8E7F6FFE243100C20FC0DFD6884CA" author="Walter, D. E." box="[602,744,907,933]" pageId="14" pageNumber="15" pagination="271 - 290" refId="ref49287" refString="Walter, D. E. (1987) Belowground arthropods of semiarid grasslands. In: Capinera, J. L. (Ed.), Integrated Pest Management on Rangeland: A Shortgrass Prairie Perspective. Westview Press, Boulder, Colorado, pp. 271-290. https://doi.org/10.1201/9780429043673-19" type="book chapter" year="1987">Walter 1987</bibRefCitation>
). The solitary female from
<taxonomicName id="4C69E184FFE243100A5AFC0DFBDE84CA" box="[1056,1118,907,933]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFE243100A5AFC0DFBDE84CA" box="[1056,1118,907,933]" italics="true" pageId="14" pageNumber="15">Abies</emphasis>
</taxonomicName>
duff in Utah with one extra prodorsal seta (
<figureCitation id="13528682FFE243100F39FC29FE3D84A6" box="[323,445,943,969]" captionStart="FIGURE 3" captionStartId="12.[151,250,1874,1898]" captionTargetBox="[151,1436,176,1851]" captionTargetId="figure-24@12.[151,1436,176,1851]" captionTargetPageId="12" captionText="FIGURE 3. Alycus utahensis sp. nov. A—Prodorsum,patternof continuousridgeson C-segment,setaeinwithelongated cilia,oneextraseta(arrow); B—Dorsal habitus, setae with 23 elongated cilia especially in caudal segments; C—Chelicera with one seta, palp tarsus with 6 pseudacanthoid eupathidia, 3 pairs of adoral setae; D— Genital area of female, 2931 genital setae per valve, 1 pair of eugenital setae (arrows, broken lines); E—Anal area, 7 anal setae per valve. All specimens—from Utah, Summit Co." pageId="14" pageNumber="15">Fig. 3A, B</figureCitation>
, arrow) is unusual but may be anomaly of this single specimen or perhaps an expression of increased neotrichy. However, the character state combination listed above for
<taxonomicName id="4C69E184FFE243100A5EFC55FB608482" authorityName="Uusitalo" authorityYear="2025" box="[1060,1248,979,1005]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="utahensis">
<emphasis id="B91D4615FFE243100A5EFC55FB608482" box="[1060,1248,979,1005]" italics="true" pageId="14" pageNumber="15">Alycus utahensis</emphasis>
</taxonomicName>
strongly suggest that the specimen represents a new species.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFE243100EBDFB9DFBF0835A" box="[199,1136,1051,1077]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8BD69A07FFE243100EBDFB9DFBF0835A" blockId="14.[151,1437,151,1077]" box="[199,1136,1051,1077]" pageId="14" pageNumber="15">
<emphasis id="B91D4615FFE243100EBDFB9DFECE835A" bold="true" box="[199,334,1051,1077]" pageId="14" pageNumber="15">Etymology.</emphasis>
The specific name refers to the state
<collectingRegion id="49AD54E5FFE243100C8AFB9DFCA9835A" box="[752,809,1051,1077]" country="United States of America" name="Utah" pageId="14" pageNumber="15">Utah</collectingRegion>
, the state of the
<typeStatus id="54D224A5FFE243100DA5FB9DFB8F835A" box="[991,1039,1051,1077]" pageId="14" pageNumber="15">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,240 @@
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<mods:identifier id="49179D9FD5802EB60FAF1B0F42524F59" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="326E6CDBE631CED8CA383CD15362DF2B" type="ZooBank">D4E6D5D7-2723-4AAB-BAB4-A1F11E40AE37</mods:identifier>
</mods:mods>
<treatment id="03C02B11FFF143000EEDFF11FCAA835A" LSID="urn:lsid:plazi:treatment:03C02B11FFF143000EEDFF11FCAA835A" httpUri="http://treatment.plazi.org/id/03C02B11FFF143000EEDFF11FCAA835A" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="C373C98CFFF143030EEDFF11FE7587DE" box="[151,501,151,178]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BD69A07FFF143030EEDFF11FE7587DE" blockId="29.[151,501,151,214]" box="[151,501,151,178]" pageId="29" pageNumber="30">
<heading id="D09E2D6BFFF143030EEDFF11FE7587DE" bold="true" box="[151,501,151,178]" fontSize="11" level="2" pageId="29" pageNumber="30" reason="4">
<emphasis id="B91D4615FFF143030EEDFF11FE7587DE" bold="true" box="[151,501,151,178]" pageId="29" pageNumber="30">
<taxonomicName id="4C69E184FFF143030EEDFF11FE1087DD" authorityName="Uusitalo" authorityYear="2025" box="[151,400,151,178]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="B91D4615FFF143030EEDFF11FE1087DD" bold="true" box="[151,400,151,178]" italics="true" pageId="29" pageNumber="30">Amphialycus walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFF143030FEDFF11FE7587DE" box="[407,501,151,177]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF143030EEDFF3DFE1587B9" box="[151,405,187,214]" pageId="29" pageNumber="30" type="description">
<paragraph id="8BD69A07FFF143030EEDFF3DFE1587B9" blockId="29.[151,501,151,214]" box="[151,405,187,214]" pageId="29" pageNumber="30">
(
<figureCitation id="13528682FFF143030EE5FF3DFEB187BA" box="[159,305,187,213]" captionStart="FIGURE 14" captionStartId="28.[151,250,1864,1888]" captionTargetBox="[191,1397,153,1839]" captionTargetId="figure-24@28.[190,1398,153,1839]" captionTargetPageId="28" captionText="FIGURE 14. Amphialycus walteri sp. nov. A—prodorsum, naso flap-like with roundish tip and longitudinally striated; B—dorsal habitus; C—subcapitulum, rutella, 1 pair of simple cheliceral setae, palpi with 4 pairs of microtrichous eupathidia; D—genital area, 1921 genital setae per valve; E—anal area, 4 anal setae per valve. All specimens—from Alberta, Lamont Co." pageId="29" pageNumber="30">Figs. 14AE</figureCitation>
,
<figureCitation id="13528682FFF143030F46FF3DFE0D87B9" box="[316,397,187,214]" captionStart="FIGURE 15" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[151,1436,299,1919]" captionTargetId="figure-38@29.[151,1436,298,1919]" captionTargetPageId="29" captionText="FIGURE 15. Amphialycus walteri sp. nov. A—Leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV, telofemur IV. All specimens—Alberta, Lamont Co." pageId="29" pageNumber="30">15AE</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF143030EEDFE86FB8F8677" box="[151,1039,255,280]" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="8BD69A07FFF143030EEDFE86FB8F8677" blockId="29.[151,1039,255,280]" box="[151,1039,255,280]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="AB79BD29FFF143030EEDFE86FB8F8677" box="[151,1039,255,280]" pageId="29" pageNumber="30">
<taxonomicName id="4C69E184FFF143030EEDFE86FE378677" ID-CoL="CYZ8" baseAuthorityName="Grandjean" baseAuthorityYear="1937" box="[151,439,256,280]" class="Arachnida" family="Alycidae" genus="Amphialycus" isUncertain="true" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="leucogaster">
<emphasis id="B91D4615FFF143030EEDFE86FE9A8677" box="[151,282,256,280]" italics="true" pageId="29" pageNumber="30">Amphialycus</emphasis>
cf.
<emphasis id="B91D4615FFF143030F3BFE86FE378677" box="[321,439,256,280]" italics="true" pageId="29" pageNumber="30">leucogaster</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AC8BC16FFF143030FADFE86FD6C8678" author="Walter, D. E. &amp; Latonas, S." box="[471,748,255,280]" page="62" pageId="29" pageNumber="30" year="2012">
<bibRefCitation id="EFF8E7F6FFF143030FADFE86FD6C8678" author="Walter, D. E. &amp; Latonas, S." box="[471,748,255,280]" pageId="29" pageNumber="30" pagination="61 - 62" refId="ref49534" refString="Walter, D. E. &amp; Latonas, S. (2012) Almanac of Alberta Acari Part II. Ver. 2.1., The Royal Alberta Museum, Edmonton, Alberta, pp. 61-62. [http://www.royalalbertamuseum.ca/research/lifeSciences/invertebrateZoology/research.cfm]" type="book chapter" year="2012">Walter &amp; Latonas 2012: 62</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC8BC16FFF143030C8DFE86FB8B8677" author="Walter, D. E. &amp; Lumley, L. M." box="[759,1035,256,280]" page="74" pageId="29" pageNumber="30" year="2021">
<bibRefCitation id="EFF8E7F6FFF143030C8DFE86FB8B8677" author="Walter, D. E. &amp; Lumley, L. M." box="[759,1035,256,280]" pageId="29" pageNumber="30" refId="ref49595" refString="Walter, D. E. &amp; Lumley, L. M. (2021) Almanac of Alberta Acari Part II, Version 3.0. Edmonton, Alberta, 192 pp. https://doi.org/10.13140/RG.2.2.26989.72166" type="book" year="2021">Walter &amp; Lumley 2021: 74</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF16CA8FFFF143030EEDF811FAD480BB" pageId="29" pageNumber="30" startId="29.[151,250,1943,1967]" targetBox="[151,1436,299,1919]" targetPageId="29" targetType="figure">
<paragraph id="8BD69A07FFF143030EEDF811FDCE80DF" blockId="29.[151,1364,1943,2004]" box="[151,590,1943,1968]" pageId="29" pageNumber="30">
<emphasis id="B91D4615FFF143030EEDF811FEA080C0" bold="true" box="[151,288,1943,1967]" pageId="29" pageNumber="30">FIGURE 15.</emphasis>
<taxonomicName id="4C69E184FFF143030F5CF81EFE7780DF" authorityName="Uusitalo" authorityYear="2025" box="[294,503,1944,1968]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="B91D4615FFF143030F5CF81EFE7780DF" box="[294,503,1944,1968]" italics="true" pageId="29" pageNumber="30">Amphialycus walteri</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFF143030F87F81EFDCE80DF" bold="true" box="[509,590,1944,1968]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A22EFB6EFFF143030F87F81EFDCE80DF" box="[509,590,1944,1968]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFF143030EEDF83AFAD480BB" blockId="29.[151,1364,1943,2004]" box="[151,1364,1980,2004]" pageId="29" pageNumber="30">A—Leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV, telofemur IV. All specimens—Alberta, Lamont Co.</paragraph>
</caption>
<subSubSection id="C373C98CFFF243000EEDFF10FDDE850E" pageId="30" pageNumber="31" type="description">
<paragraph id="8BD69A07FFF243000EEDFF10FEBA862E" blockId="30.[151,1437,150,1077]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EEDFF10FEA987DF" bold="true" box="[151,297,150,176]" pageId="30" pageNumber="31">Description.</emphasis>
<emphasis id="B91D4615FFF243000F54FF11FE0887DF" box="[302,392,151,176]" italics="true" pageId="30" pageNumber="31">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFF243000FCFFF11FDA387DE" box="[437,547,151,177]" count="4" pageId="30" pageNumber="31" type="female">4 females</specimenCount>
,
<figureCitation id="13528682FFF243000C57FF11FD3187DE" box="[557,689,151,177]" captionStart="FIGURE 14" captionStartId="28.[151,250,1864,1888]" captionTargetBox="[191,1397,153,1839]" captionTargetId="figure-24@28.[190,1398,153,1839]" captionTargetPageId="28" captionText="FIGURE 14. Amphialycus walteri sp. nov. A—prodorsum, naso flap-like with roundish tip and longitudinally striated; B—dorsal habitus; C—subcapitulum, rutella, 1 pair of simple cheliceral setae, palpi with 4 pairs of microtrichous eupathidia; D—genital area, 1921 genital setae per valve; E—anal area, 4 anal setae per valve. All specimens—from Alberta, Lamont Co." pageId="30" pageNumber="31">Fig. 14A, B</figureCitation>
). Length 400 µm; ciliate setae with long central points (1214 µm), especially on caudal segments (1620 µm). On prodorsum: naso flap-like, longitudinally striated with roundish tip; pilose setae
<emphasis id="B91D4615FFF243000F59FF59FEB78797" box="[291,311,223,248]" italics="true" pageId="30" pageNumber="31">vi</emphasis>
set well back,
<emphasis id="B91D4615FFF243000F9EFF59FD888797" box="[484,520,223,248]" italics="true" pageId="30" pageNumber="31">sce</emphasis>
and
<emphasis id="B91D4615FFF243000C3BFF59FDE88797" box="[577,616,223,248]" italics="true" pageId="30" pageNumber="31">exp</emphasis>
plumose on hard integument,
<emphasis id="B91D4615FFF243000DB9FF59FC6A8797" box="[963,1002,223,248]" italics="true" pageId="30" pageNumber="31">exp</emphasis>
smallest and anterior to eyes, setae
<emphasis id="B91D4615FFF243000BFCFF59FA1C8797" box="[1414,1436,223,248]" italics="true" pageId="30" pageNumber="31">in</emphasis>
columnar, bushy; two pairs of filamentous sensilla
<emphasis id="B91D4615FFF243000CAFFE85FD6E8673" box="[725,750,259,284]" italics="true" pageId="30" pageNumber="31">ve</emphasis>
and
<emphasis id="B91D4615FFF243000D5FFE85FCC48673" box="[805,836,259,284]" italics="true" pageId="30" pageNumber="31">sci</emphasis>
ciliated along entire length; both pairs of lateral eyes finely striated.
</paragraph>
<paragraph id="8BD69A07FFF243000EBDFECDFBAE86E6" blockId="30.[151,1437,150,1077]" pageId="30" pageNumber="31">
<taxonomicName id="4C69E184FFF243000EBDFECDFE8F860B" box="[199,271,331,356]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFF243000EBDFECDFE8F860B" box="[199,271,331,356]" italics="true" pageId="30" pageNumber="31">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFF243000F5AFECDFE34860A" box="[288,436,331,357]" captionStart="FIGURE 14" captionStartId="28.[151,250,1864,1888]" captionTargetBox="[191,1397,153,1839]" captionTargetId="figure-24@28.[190,1398,153,1839]" captionTargetPageId="28" captionText="FIGURE 14. Amphialycus walteri sp. nov. A—prodorsum, naso flap-like with roundish tip and longitudinally striated; B—dorsal habitus; C—subcapitulum, rutella, 1 pair of simple cheliceral setae, palpi with 4 pairs of microtrichous eupathidia; D—genital area, 1921 genital setae per valve; E—anal area, 4 anal setae per valve. All specimens—from Alberta, Lamont Co." pageId="30" pageNumber="31">Figs. 14D, E</figureCitation>
,
<figureCitation id="13528682FFF243000FB8FECCFE73860A" box="[450,499,330,357]" captionStart="FIGURE 15" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[151,1436,299,1919]" captionTargetId="figure-38@29.[151,1436,298,1919]" captionTargetPageId="29" captionText="FIGURE 15. Amphialycus walteri sp. nov. A—Leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV, telofemur IV. All specimens—Alberta, Lamont Co." pageId="30" pageNumber="31">15E</figureCitation>
). Setae shorter and less ciliated than dorsal setae; genital setae of various lengths, mostly elongated closer to mesial margin, 1921 per valve; anal setae 4 per valve.
</paragraph>
<paragraph id="8BD69A07FFF243000EBDFE15FC8E869A" blockId="30.[151,1437,150,1077]" pageId="30" pageNumber="31">
<taxonomicName id="4C69E184FFF243000EBDFE15FED686C2" authorityName="Krefft" authorityYear="1871" box="[199,342,403,429]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="30" pageNumber="31" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFF243000EBDFE15FED686C2" box="[199,342,403,429]" italics="true" pageId="30" pageNumber="31">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFF243000F13FE15FE5386C2" box="[361,467,403,429]" captionStart="FIGURE 14" captionStartId="28.[151,250,1864,1888]" captionTargetBox="[191,1397,153,1839]" captionTargetId="figure-24@28.[190,1398,153,1839]" captionTargetPageId="28" captionText="FIGURE 14. Amphialycus walteri sp. nov. A—prodorsum, naso flap-like with roundish tip and longitudinally striated; B—dorsal habitus; C—subcapitulum, rutella, 1 pair of simple cheliceral setae, palpi with 4 pairs of microtrichous eupathidia; D—genital area, 1921 genital setae per valve; E—anal area, 4 anal setae per valve. All specimens—from Alberta, Lamont Co." pageId="30" pageNumber="31">Fig. 14C</figureCitation>
). One pair of smooth, abaxial cheliceral setae; rutella mitten-shaped, widening to a round hyaline membrane with two lobes apically; three pairs of well-developed adoral setae; four microtrichous eupathidia on palps close to each other but clearly apart.
</paragraph>
<paragraph id="8BD69A07FFF243000EBDFE79FDDE850E" blockId="30.[151,1437,150,1077]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EBDFE79FF7C8577" box="[199,252,511,536]" italics="true" pageId="30" pageNumber="31">Legs</emphasis>
(
<figureCitation id="13528682FFF243000F71FE79FE128576" box="[267,402,510,537]" captionStart="FIGURE 15" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[151,1436,299,1919]" captionTargetId="figure-38@29.[151,1436,298,1919]" captionTargetPageId="29" captionText="FIGURE 15. Amphialycus walteri sp. nov. A—Leg I; B—Leg II; C—Tibia III, genu III; D—Tibia IV, genu IV, telofemur IV. All specimens—Alberta, Lamont Co." pageId="30" pageNumber="31">Fig. 15AE</figureCitation>
). Pedal setae elongated; solenidial formula for tarsi, tibiae, genua and femora on legs I-II-III- IV, respectively: 2-2-0-0, 3-2-2-0, 3-2-2-1, 2-0-0-1+0, ceratiform solenidia on femora I longer than other solenidia; coxae III and IV with 3 setae,
<emphasis id="B91D4615FFF243000F90FDC1FDD7850E" bold="true" box="[490,599,583,609]" pageId="30" pageNumber="31">Remarks</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF243000EBDFDEDFC3E859E" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BD69A07FFF243000EBDFDEDFC3E859E" blockId="30.[151,1437,150,1077]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EBDFDEDFEF685EB" bold="true" box="[199,374,618,645]" pageId="30" pageNumber="31">Type material.</emphasis>
<materialsCitation id="3B01905AFFF243000F07FDEDFC3E859E" collectingDate="2008-02-17" collectionCode="TMS" collectorName="David E. Walter" country="Canada" county="Lamont Co." latitude="53.66028" location="Moose Pasture Research Site" longLatPrecision="17" longitude="-112.75584" pageId="30" pageNumber="31" specimenCode="M00016839" specimenCount="7" specimenCount-female="7" stateProvince="Alberta" typeStatus="holotype">
<typeStatus id="54D224A5FFF243000F07FDEDFE6585EA" box="[381,485,619,645]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
female and
<specimenCount id="9D6F518EFFF243000C0AFDEDFCC585EA" box="[624,837,619,645]" count="3" pageId="30" pageNumber="31" type="female" typeStatus="paratype">3 paratype females</specimenCount>
as
<taxonomicName id="4C69E184FFF243000D11FDEDFB2B85EA" baseAuthorityName="Grandjean" baseAuthorityYear="1937" box="[875,1195,619,645]" class="Arachnida" family="Alycidae" genus="Amphialycus" isUncertain="true" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="leucogaster">
<emphasis id="B91D4615FFF243000D11FDEDFC7C85EA" box="[875,1020,619,645]" italics="true" pageId="30" pageNumber="31">Amphialycus</emphasis>
cf.
<emphasis id="B91D4615FFF243000A52FDEDFB2B85EA" box="[1064,1195,619,645]" italics="true" pageId="30" pageNumber="31">leucogaster</emphasis>
</taxonomicName>
from aspen (
<taxonomicName id="4C69E184FFF243000B3AFDEDFE9A85C6" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="tremuloides">
<emphasis id="B91D4615FFF243000B3AFDEDFE9A85C6" italics="true" pageId="30" pageNumber="31">Populus tremuloides</emphasis>
</taxonomicName>
) litter under snow,
<location id="8EB6CCDCFFF243000C75FD09FCEC85C6" LSID="urn:lsid:plazi:treatment:03C02B11FFF143000EEDFF11FCAA835A:8EB6CCDCFFF243000C75FD09FCEC85C6" box="[527,876,655,681]" country="Canada" county="Lamont Co." latitude="53.66028" longLatPrecision="17" longitude="-112.75584" name="Moose Pasture Research Site" pageId="30" pageNumber="31" stateProvince="Alberta">Moose Pasture Research Site</location>
,
<locationDeviation id="9A648325FFF243000D04FD09FBDC85C6" box="[894,1116,655,681]" location="Chipman" pageId="30" pageNumber="31">
<quantity id="4C9137E2FFF243000D04FD09FC3E85C6" box="[894,958,655,681]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="31" unit="km" value="8.0">8 km</quantity>
W Chipman
</locationDeviation>
,
<collectingCounty id="62B7E28BFFF243000A14FD09FB7D85C6" box="[1134,1277,655,681]" pageId="30" pageNumber="31">Lamont Co.</collectingCounty>
,
<geoCoordinate id="EE5DFCC0FFF243000B75FD08FA1C85C6" box="[1295,1436,654,681]" degrees="53" direction="north" minutes="39" orientation="latitude" pageId="30" pageNumber="31" precision="15" seconds="37" value="53.66028">53°3937N</geoCoordinate>
<geoCoordinate id="EE5DFCC0FFF243000EEDFD34FEB585A2" box="[151,309,690,717]" degrees="112" direction="west" minutes="45" orientation="longitude" pageId="30" pageNumber="31" precision="15" seconds="21" value="-112.75584">112°4521W</geoCoordinate>
,
<collectingRegion id="49AD54E5FFF243000F3CFD35FE1D85A2" box="[326,413,691,717]" country="Canada" name="Alberta" pageId="30" pageNumber="31">Alberta</collectingRegion>
,
<collectingCountry id="F37EDA97FFF243000FD6FD35FD8385A2" box="[428,515,691,717]" name="Canada" pageId="30" pageNumber="31">Canada</collectingCountry>
,
<date id="FFD7BCC7FFF243000C68FD35FD6585A2" box="[530,741,691,717]" pageId="30" pageNumber="31" value="2008-02-17">
<collectingDate id="EF93452FFFF243000C68FD35FD6585A2" box="[530,741,691,717]" pageId="30" pageNumber="31" value="2008-02-17">17 February 2008</collectingDate>
</date>
,
<collectorName id="269CFFD1FFF243000C8EFD35FC3585A2" box="[756,949,691,717]" pageId="30" pageNumber="31">David E. Walter</collectorName>
:
<specimenCount id="9D6F518EFFF243000DB9FD35FBA885A2" box="[963,1064,691,717]" count="4" pageId="30" pageNumber="31" type="female">4 female</specimenCount>
cotypes on one slide (8)
<collectionCode id="ED7802C2FFF243000B21FD35FA1985A2" box="[1371,1433,691,717]" country="0" httpUri="http://grbio.org/cool/qe02-4904" name="Toleco Museum of Health and Natural History" pageId="30" pageNumber="31">TMS</collectionCode>
:
<specimenCode id="DBCF327CFFF243000EEDFD51FEA1859E" box="[151,289,727,753]" collectionCode="TMS" country="0" httpUri="http://grbio.org/cool/qe02-4904" name="Toleco Museum of Health and Natural History" pageId="30" pageNumber="31">M00016839</specimenCode>
at the Royal
<collectingRegion id="49AD54E5FFF243000FCEFD51FD88859E" box="[436,520,727,753]" country="Canada" name="Alberta" pageId="30" pageNumber="31">Alberta</collectingRegion>
Museum, Edmonton,
<collectingRegion id="49AD54E5FFF243000D7AFD51FCD7859E" box="[768,855,727,753]" country="Canada" name="Alberta" pageId="30" pageNumber="31">Alberta</collectingRegion>
,
<collectingCountry id="F37EDA97FFF243000D18FD51FC3A859E" box="[866,954,727,753]" name="Canada" pageId="30" pageNumber="31">Canada</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF243000EBDFD7CFA8084CA" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BD69A07FFF243000EBDFD7CFA8084CA" blockId="30.[151,1437,150,1077]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EBDFD7CFE4D847B" bold="true" box="[199,461,762,788]" pageId="30" pageNumber="31">Differential diagnosis.</emphasis>
This species can be differentiated from
<taxonomicName id="4C69E184FFF243000DF7FD7DFB21847A" box="[909,1185,763,789]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="holarcticus">
<emphasis id="B91D4615FFF243000DF7FD7DFB21847A" box="[909,1185,763,789]" italics="true" pageId="30" pageNumber="31">Amphialycus holarcticus</emphasis>
</taxonomicName>
by having a naso with a roundish tip, one cheliceral seta, fewer genital, anal and coxal setae (Uusitalo 2020: fig. 6), and two solenidia on tarsus II; and from
<taxonomicName id="4C69E184FFF243000F0EFCC5FCE78432" authority="(Grandjean, 1937)" baseAuthorityName="Grandjean" baseAuthorityYear="1937" box="[372,871,835,861]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="leucogaster">
<emphasis id="B91D4615FFF243000F0EFCC5FD108432" box="[372,656,835,861]" italics="true" pageId="30" pageNumber="31">Amphialycus leucogaster</emphasis>
(
<bibRefCitation id="EFF8E7F6FFF243000CDAFCC5FCDF8432" author="Grandjean, F." box="[672,863,835,861]" pageId="30" pageNumber="31" pagination="56 - 61" refId="ref47047" refString="Grandjean, F. (1937) Le genre Pachygnathus Duges (Alycus Koch) (Acariens). Deuxieme partie. Bulletin Museum national d'Histoire naturelle, Paris, 2 e Serie, 9, 56-61." type="journal article" year="1937">Grandjean, 1937</bibRefCitation>
)
</taxonomicName>
by having dorsal and prodorsal setae with longer shafts; crista with longitudinally (not transversely) orientated lamellae (
<bibRefCitation id="EFF8E7F6FFF243000DC5FCE1FBE484EE" author="Uusitalo, M." box="[959,1124,871,897]" pageId="30" pageNumber="31" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: figs. 7173), three pairs of well-developed adoral setae (instead of one), and two ceratiform solenidia on tibia I (instead of one).
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF243000EBDFC29FABF8482" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BD69A07FFF243000EBDFC29FABF8482" blockId="30.[151,1437,150,1077]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EBDFC29FEBB84A6" bold="true" box="[199,315,943,969]" pageId="30" pageNumber="31">Remarks.</emphasis>
Poor visibility of the distal parts of tarsi I and II prevented reliable observation of the minute famulus I and II.
<taxonomicName id="4C69E184FFF243000E82FC55FE5E8482" authorityName="Uusitalo" authorityYear="2025" box="[248,478,979,1005]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="walteri">
<emphasis id="B91D4615FFF243000E82FC55FE5E8482" box="[248,478,979,1005]" italics="true" pageId="30" pageNumber="31">Amphialycus walteri</emphasis>
</taxonomicName>
was not known when preparing the phylogenetic tree (
<bibRefCitation id="EFF8E7F6FFF243000A3FFC55FB698482" author="Uusitalo, M." box="[1093,1257,979,1005]" pageId="30" pageNumber="31" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: fig. 1).
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF243000EBDFC71FCAA835A" pageId="30" pageNumber="31" type="etymology">
<paragraph id="8BD69A07FFF243000EBDFC71FCAA835A" blockId="30.[151,1437,150,1077]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EBDFC71FECE837E" bold="true" box="[199,334,1015,1041]" pageId="30" pageNumber="31">Etymology.</emphasis>
The specific name (a patronymic genitive) is given in honour of Dr. David Evans Walter, the collector and first recorder of this genus in North America.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,188 @@
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<subSubSection id="C373C98CFFF243000EEDFB00FD8283CF" box="[151,514,1158,1185]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BD69A07FFF243000EEDFB00FD8283CF" blockId="30.[151,514,1158,1185]" box="[151,514,1158,1185]" pageId="30" pageNumber="31">
<heading id="D09E2D6BFFF243000EEDFB00FD8283CF" bold="true" box="[151,514,1158,1185]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
<taxonomicName id="4C69E184FFF243000EEDFB00FD8283CF" ID-CoL="63HDX" authority="Zachvatkin, 1949" authorityName="Zachvatkin" authorityYear="1949" box="[151,514,1158,1185]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000EEDFB00FD8283CF" bold="true" box="[151,514,1158,1185]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EEDFB00FEA283CE" bold="true" box="[151,290,1158,1185]" italics="true" pageId="30" pageNumber="31">Orthacarus</emphasis>
<bibRefCitation id="EFF8E7F6FFF243000F53FB00FD8283CF" author="Zachvatkin, A. A." box="[297,514,1158,1184]" pageId="30" pageNumber="31" pagination="292 - 297" refId="ref49859" refString="Zachvatkin, A. A. (1949) New representatives of segmented mites (Acarina, Pachygnathidae). Entomologicheskoye Obozreniye, 30, 292-297. [in Russian]" type="journal article" year="1949">Zachvatkin, 1949</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF243000EEDFB4AFBA3838C" box="[151,1059,1227,1252]" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BD69A07FFF243000EEDFB4AFBA3838C" blockId="30.[151,1059,1227,1252]" box="[151,1059,1227,1252]" pageId="30" pageNumber="31">
<materialsCitation id="3B01905AFFF243000EEDFB4AFB9E838C" box="[151,1054,1227,1252]" country="Ukraine" location="Ukraine" pageId="30" pageNumber="31" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D224A5FFF243000EEDFB4AFF49838B" box="[151,201,1228,1252]" pageId="30" pageNumber="31">Type</typeStatus>
species:
<taxonomicName id="4C69E184FFF243000F5CFB4DFD15838C" authority="Zachvatkin, 1949" authorityName="Zachvatkin" authorityYear="1949" box="[294,661,1227,1251]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="tremli">
<emphasis id="B91D4615FFF243000F5CFB4DFE5D838C" box="[294,477,1227,1251]" italics="true" pageId="30" pageNumber="31">Orthacarus tremli</emphasis>
<bibRefCitation id="EFF8E7F6FFF243000F99FB4DFD15838C" author="Zachvatkin, A. A." box="[483,661,1227,1251]" pageId="30" pageNumber="31" pagination="292 - 297" refId="ref49859" refString="Zachvatkin, A. A. (1949) New representatives of segmented mites (Acarina, Pachygnathidae). Entomologicheskoye Obozreniye, 30, 292-297. [in Russian]" type="journal article" year="1949">Zachvatkin, 1949</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F37EDA97FFF243000CA9FB4DFCA8838C" box="[723,808,1227,1251]" name="Ukraine" pageId="30" pageNumber="31">Ukraine</collectingCountry>
; by original designation
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF243000EEDFA97FDD68148" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BD69A07FFF243000EEDFA97FDD68148" blockId="30.[151,1437,1297,2008]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EEDFA97FE218244" bold="true" box="[151,417,1297,1323]" pageId="30" pageNumber="31">Differential diagnosis.</emphasis>
This genus is similar to
<taxonomicName id="4C69E184FFF243000CB2FA97FBAB8244" authority="Zachvatkin, 1949" authorityName="Zachvatkin" authorityYear="1949" box="[712,1067,1297,1323]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000CB2FA97FCD98244" box="[712,857,1297,1323]" italics="true" pageId="30" pageNumber="31">Amphialycus</emphasis>
<bibRefCitation id="EFF8E7F6FFF243000D19FA97FBAB8244" author="Zachvatkin, A. A." box="[867,1067,1297,1323]" pageId="30" pageNumber="31" pagination="292 - 297" refId="ref49859" refString="Zachvatkin, A. A. (1949) New representatives of segmented mites (Acarina, Pachygnathidae). Entomologicheskoye Obozreniye, 30, 292-297. [in Russian]" type="journal article" year="1949">
Zachvatkin,
<quantity id="4C9137E2FFF243000D89FA97FBAB8244" box="[1011,1067,1297,1323]" metricMagnitude="1" metricUnit="m" metricValue="4.95046" pageId="30" pageNumber="31" unit="in" value="1949.0">1949</quantity>
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in having robust and elongated chelicerae, a finely striated anterior pair of eyes, and a densely setulated empodia.
<taxonomicName id="4C69E184FFF243000AF8FAB3FA838220" authorityName="Zachvatkin" authorityYear="1949" box="[1154,1283,1333,1359]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000AF8FAB3FA838220" box="[1154,1283,1333,1359]" italics="true" pageId="30" pageNumber="31">Orthacarus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C69E184FFF243000EEDFADFFEA8821C" authorityName="Zachvatkin" authorityYear="1949" box="[151,296,1369,1395]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000EEDFADFFEA8821C" box="[151,296,1369,1395]" italics="true" pageId="30" pageNumber="31">Amphialycus</emphasis>
</taxonomicName>
by having sparsely ciliated dorsal setae, a tiny and elongated body, and the prodorsal setae reduced in size. Other diagnostic features are as follows: naso is absent; prodorsal setae
<emphasis id="B91D4615FFF243000D95FAF8FB8382F8" box="[1007,1027,1406,1431]" italics="true" pageId="30" pageNumber="31">vi</emphasis>
are very close to each other; setae
<emphasis id="B91D4615FFF243000BFCFAF8FA1C82F8" box="[1414,1436,1406,1431]" italics="true" pageId="30" pageNumber="31">in</emphasis>
are pubescent or not; four palpal eupathidia are fused or basally very close to each other; legs II, III and IV are short and strongly bent; and the anus is caudal (
<figureCitation id="13528682FFF243000C09FA43FD5182B0" box="[627,721,1477,1503]" captionStart="FIGURE 16" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[151,1435,232,1851]" captionTargetId="figure-23@31.[151,1435,232,1851]" captionTargetPageId="31" captionText="FIGURE 16. Orthacarus oblongus (Halbert). A—prodorsum, setae in elongated (11,5 µm); B—prodorsum, setae in elongated (8 µm); C—leg I; D—leg II; E—tarsus and tibia III; F—tarsus, tibia and genu IV; G—palpal tarsus, lateral and ventral sides with 4 barbed and basally fused eupathidia. 16A, CG—from Quintana Roo, Sian Kaan; 16B—from Italy, Tuscany." pageId="30" pageNumber="31">Figs. 16</figureCitation>
,
<figureCitation id="13528682FFF243000CA4FA43FCC9828F" box="[734,841,1477,1504]" captionStart="FIGURE 50" captionStartId="82.[151,250,1759,1783]" captionTargetBox="[164,1423,161,1735]" captionTargetId="figure-24@82.[164,1423,161,1735]" captionTargetPageId="82" captionText="FIGURE 50. Genus-level character states of Alycini. AC—Alycus: prodorsum with two pairs of sensilla and multiciliated setae in; robust and short chelicera; and six pseudacanthoid palpal eupathidia, respectively; DF—Pachygnathus: prodorsum with two pairs of sensilla and multiciliated setae in; beak-like chelicera; andpalpal tarsus with 4 eupathidia,respectively; GI—Amphialycus: prodorsum with setae vi on both side of prominent naso and pubescent setae in; robust and elongated chelicera; and palpal tarsus with five (or four) barbed eupathidia, respectively; JL—Orthacarus: prodorsum without naso, reduced setae vi exceptionally close to each other, and setae in pubescent; robust and elongated chelicera; and palpal tarsus with four barbed and fused eupathidia, respectively. All photographs are copied from Uusitalo (2010)." pageId="30" pageNumber="31">50J, K, L</figureCitation>
,
<figureCitation id="13528682FFF243000D2CFA43FCF4828F" box="[854,884,1477,1504]" captionStart="FIGURE 54" captionStartId="86.[151,250,1908,1932]" captionTargetBox="[151,1436,176,1884]" captionTargetId="figure-24@86.[151,1436,176,1884]" captionTargetPageId="86" captionText="FIGURE 54. Selected figures of an undescribed species clu from South Africa. A—prodorsum; B—dorsum; C—dorsal seta; D—venter; E—genital valves; F—palpal eupathidia; G—Leg I; H—Empodium." pageId="30" pageNumber="31">54</figureCitation>
,
<bibRefCitation id="EFF8E7F6FFF243000DFBFA43FBA382B0" author="Uusitalo, M." box="[897,1059,1477,1503]" pageId="30" pageNumber="31" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: figs. 8286,
<bibRefCitation id="EFF8E7F6FFF243000AC6FA43FA1882B0" author="Uusitalo, M. &amp; Ueckermann, E. A. &amp; Theron, P. D." box="[1212,1432,1477,1503]" pageId="30" pageNumber="31" pagination="301 - 340" refId="ref48918" refString="Uusitalo, M., Ueckermann, E. A. &amp; Theron, P. D. (2020) A review of the family Alycidae (Acari, Acariformes) from South Africa. Zootaxa, 4858 (3), 301-340. https://doi.org/10.11646/zootaxa.4858.3.1" type="journal article" year="2020">
Uusitalo
<emphasis id="B91D4615FFF243000B59FA40FAD582B0" box="[1315,1365,1477,1503]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
2020
</bibRefCitation>
: figs 7683). The other genera of the tribe
<taxonomicName id="4C69E184FFF243000C11FA6FFD3F816C" box="[619,703,1513,1539]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="tribe" tribe="Alycini">Alycini</taxonomicName>
have either robust and non-elongated (
<taxonomicName id="4C69E184FFF243000A0CFA6FFB41816C" authorityName="C. L. Koch" authorityYear="1842" box="[1142,1217,1513,1539]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000A0CFA6FFB41816C" box="[1142,1217,1513,1539]" italics="true" pageId="30" pageNumber="31">Alycus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFF243000A8DFA6FFA13816C" authorityName="Uusitalo" authorityYear="2025" box="[1271,1427,1513,1539]" class="Arachnida" family="Alycidae" genus="Odontoalycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000A8DFA6FFA13816C" box="[1271,1427,1513,1539]" italics="true" pageId="30" pageNumber="31">Odontoalycus</emphasis>
</taxonomicName>
) or beak-like (
<taxonomicName id="4C69E184FFF243000F54F98BFE4D8148" authorityName="Duges" authorityYear="1834" box="[302,461,1549,1575]" class="Arachnida" family="Alycidae" genus="Pachygnathus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000F54F98BFE4D8148" box="[302,461,1549,1575]" italics="true" pageId="30" pageNumber="31">Pachygnathus</emphasis>
</taxonomicName>
) chelicerae.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF243000EBDF9B7FA0580B8" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BD69A07FFF243000EBDF9B7FA0580B8" blockId="30.[151,1437,1297,2008]" pageId="30" pageNumber="31">
<emphasis id="B91D4615FFF243000EBDF9B7FEB08124" bold="true" box="[199,304,1585,1611]" pageId="30" pageNumber="31">Remark.</emphasis>
The character states, suggesting a more edaphic way of life, was supposed to be unique to a single, cosmopolitan species
<taxonomicName id="4C69E184FFF243000FEEF9D0FDA08100" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[404,544,1621,1648]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="B91D4615FFF243000FEEF9D0FDA08100" box="[404,544,1621,1648]" italics="true" pageId="30" pageNumber="31">O. oblongus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C69E184FFF243000C48F9D3FD338100" authorityName="Zachvatkin" authorityYear="1949" box="[562,691,1621,1647]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000C48F9D3FD338100" box="[562,691,1621,1647]" italics="true" pageId="30" pageNumber="31">Orthacarus</emphasis>
</taxonomicName>
was not supported phylogenetically in
<bibRefCitation id="EFF8E7F6FFF243000A05F9D3FAB48100" author="Uusitalo, M." box="[1151,1332,1621,1647]" pageId="30" pageNumber="31" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo (2010)</bibRefCitation>
and was therefore tentatively considered as a subgenus of
<taxonomicName id="4C69E184FFF243000CADF9FFFCE881FC" authorityName="Zachvatkin" authorityYear="1949" box="[727,872,1657,1683]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000CADF9FFFCE881FC" box="[727,872,1657,1683]" italics="true" pageId="30" pageNumber="31">Amphialycus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF8E7F6FFF243000D00F9FFFBA081FC" author="Uusitalo, M." box="[890,1056,1657,1683]" pageId="30" pageNumber="31" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
,
<bibRefCitation id="EFF8E7F6FFF243000A54F9FFFA9A81FC" author="Uusitalo, M. &amp; Ueckermann, E. A. &amp; Theron, P. D." box="[1070,1306,1657,1683]" pageId="30" pageNumber="31" pagination="301 - 340" refId="ref48918" refString="Uusitalo, M., Ueckermann, E. A. &amp; Theron, P. D. (2020) A review of the family Alycidae (Acari, Acariformes) from South Africa. Zootaxa, 4858 (3), 301-340. https://doi.org/10.11646/zootaxa.4858.3.1" type="journal article" year="2020">
Uusitalo
<emphasis id="B91D4615FFF243000AEDF9FCFB5381FC" box="[1175,1235,1657,1683]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2020
</bibRefCitation>
). However, observations of a new species with robust and non-elongated chelicerae (e.g.
<taxonomicName id="4C69E184FFF243000D97F91BFB0981D8" authorityName="Uusitalo" authorityYear="2025" box="[1005,1161,1693,1719]" class="Arachnida" family="Alycidae" genus="Odontoalycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000D97F91BFB0981D8" box="[1005,1161,1693,1719]" italics="true" pageId="30" pageNumber="31">Odontoalycus</emphasis>
</taxonomicName>
with 20 teeth) suggested that the basic shape of the mouthparts, like robustness and elongation, may also have appeared several times in evolution (also present in
<taxonomicName id="4C69E184FFF243000FC7F963FDB58190" authorityName="Grandjean" authorityYear="1943" box="[445,565,1765,1791]" class="Arachnida" family="Alycidae" genus="Petralycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000FC7F963FDB58190" box="[445,565,1765,1791]" italics="true" pageId="30" pageNumber="31">Petralycus</emphasis>
</taxonomicName>
), and thus is not enough to justify
<taxonomicName id="4C69E184FFF243000DBAF963FBA78190" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[960,1063,1765,1791]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="B91D4615FFF243000DBAF963FBA78190" box="[960,1063,1765,1791]" italics="true" pageId="30" pageNumber="31">oblongus</emphasis>
</taxonomicName>
being in the genus
<taxonomicName id="4C69E184FFF243000B7FF963FA168190" authorityName="Zachvatkin" authorityYear="1949" box="[1285,1430,1765,1791]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000B7FF963FA168190" box="[1285,1430,1765,1791]" italics="true" pageId="30" pageNumber="31">Amphialycus</emphasis>
</taxonomicName>
. The same combination of edaphomorphic character states is present in an undescribed species
<emphasis id="B91D4615FFF243000AA6F88FFA91804C" box="[1244,1297,1801,1827]" italics="true" pageId="30" pageNumber="31">clu</emphasis>
from
<collectingCountry id="F37EDA97FFF243000B21F88FFF5F8028" name="South Africa" pageId="30" pageNumber="31">South Africa</collectingCountry>
(
<figureCitation id="13528682FFF243000E94F8ABFEC38027" box="[238,323,1837,1864]" captionStart="FIGURE 54" captionStartId="86.[151,250,1908,1932]" captionTargetBox="[151,1436,176,1884]" captionTargetId="figure-24@86.[151,1436,176,1884]" captionTargetPageId="86" captionText="FIGURE 54. Selected figures of an undescribed species clu from South Africa. A—prodorsum; B—dorsum; C—dorsal seta; D—venter; E—genital valves; F—palpal eupathidia; G—Leg I; H—Empodium." pageId="30" pageNumber="31">Fig. 54</figureCitation>
). However, the preliminary analysis missed, for example, the exceptional mutual closeness of setae
<emphasis id="B91D4615FFF243000EEDF8D4FF2B8004" box="[151,171,1874,1899]" italics="true" pageId="30" pageNumber="31">vi</emphasis>
, the shape of legs and the location of anus, which in combination is unique to the two species of
<taxonomicName id="4C69E184FFF243000A9FF8D7FAE68004" authorityName="Zachvatkin" authorityYear="1949" box="[1253,1382,1873,1899]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF243000A9FF8D7FAE68004" box="[1253,1382,1873,1899]" italics="true" pageId="30" pageNumber="31">Orthacarus</emphasis>
</taxonomicName>
, and together with other edaphomorphisms, it seems justified to reinstate the old genus name. The undescribed species
<emphasis id="B91D4615FFF243000EDBF81FFF4380DC" box="[161,195,1945,1971]" italics="true" pageId="30" pageNumber="31">clu</emphasis>
from
<collectingCountry id="F37EDA97FFF243000F75F81FFE1E80DC" box="[271,414,1945,1971]" name="South Africa" pageId="30" pageNumber="31">South Africa</collectingCountry>
(
<figureCitation id="13528682FFF243000FD7F81FFE7E80DB" box="[429,510,1945,1972]" captionStart="FIGURE 54" captionStartId="86.[151,250,1908,1932]" captionTargetBox="[151,1436,176,1884]" captionTargetId="figure-24@86.[151,1436,176,1884]" captionTargetPageId="86" captionText="FIGURE 54. Selected figures of an undescribed species clu from South Africa. A—prodorsum; B—dorsum; C—dorsal seta; D—venter; E—genital valves; F—palpal eupathidia; G—Leg I; H—Empodium." pageId="30" pageNumber="31">Fig. 54</figureCitation>
) is in the Collection of the late Prof. Pieter Therons and has been temporarily lost in removal of the collection; therefore, its description is unpublished so far (Dr. Edward A. Ueckermann, in litt.).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,332 @@
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<paragraph id="8BD69A07FFF343010EEDFF11FD6C87DE" blockId="31.[151,748,151,213]" box="[151,748,151,178]" pageId="31" pageNumber="32">
<heading id="D09E2D6BFFF343010EEDFF11FD6C87DE" bold="true" box="[151,748,151,178]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="4">
<emphasis id="B91D4615FFF343010EEDFF11FD6C87DE" bold="true" box="[151,748,151,178]" pageId="31" pageNumber="32">
<taxonomicName id="4C69E184FFF343010EEDFF11FDDE87DE" authority="(Halbert, 1920)" authorityName="Uusitalo" authorityYear="2025" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[151,606,151,178]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="oblongus" status="comb. nov.">
<emphasis id="B91D4615FFF343010EEDFF11FE1887DD" bold="true" box="[151,408,151,178]" italics="true" pageId="31" pageNumber="32">Orthacarus oblongus</emphasis>
(Halbert, 1920)
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<paragraph id="8BD69A07FFF343010EEDFF3DFEB087BA" blockId="31.[151,748,151,213]" box="[151,304,187,213]" pageId="31" pageNumber="32">
(
<figureCitation id="13528682FFF343010EE5FF3DFEA887BA" box="[159,296,187,213]" captionStart="FIGURE 16" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[151,1435,232,1851]" captionTargetId="figure-23@31.[151,1435,232,1851]" captionTargetPageId="31" captionText="FIGURE 16. Orthacarus oblongus (Halbert). A—prodorsum, setae in elongated (11,5 µm); B—prodorsum, setae in elongated (8 µm); C—leg I; D—leg II; E—tarsus and tibia III; F—tarsus, tibia and genu IV; G—palpal tarsus, lateral and ventral sides with 4 barbed and basally fused eupathidia. 16A, CG—from Quintana Roo, Sian Kaan; 16B—from Italy, Tuscany." pageId="31" pageNumber="32">Fig. 16AG</figureCitation>
)
</paragraph>
<caption id="DF16CA8FFFF343010EEDF8D4FCF880B8" pageId="31" pageNumber="32" startId="31.[151,250,1874,1898]" targetBox="[151,1435,232,1851]" targetPageId="31" targetType="figure">
<paragraph id="8BD69A07FFF343010EEDF8D4FDE98005" blockId="31.[151,1437,1874,2007]" box="[151,617,1874,1898]" pageId="31" pageNumber="32">
<emphasis id="B91D4615FFF343010EEDF8D4FEA08005" bold="true" box="[151,288,1874,1898]" pageId="31" pageNumber="32">FIGURE 16.</emphasis>
<taxonomicName id="4C69E184FFF343010F5CF8D4FDE48005" authority="(Halbert)" authorityName="Uusitalo" authorityYear="2025" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[294,612,1874,1898]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="B91D4615FFF343010F5CF8D4FE7E8005" box="[294,510,1874,1898]" italics="true" pageId="31" pageNumber="32">Orthacarus oblongus</emphasis>
(Halbert)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD69A07FFF343010EEDF8F0FCF880B8" blockId="31.[151,1437,1874,2007]" pageId="31" pageNumber="32">
A—prodorsum, setae
<emphasis id="B91D4615FFF343010F03F8F1FE0D80E1" box="[377,397,1911,1934]" italics="true" pageId="31" pageNumber="32">in</emphasis>
elongated (11,5 µm); B—prodorsum, setae
<emphasis id="B91D4615FFF343010D2FF8F1FCE980E1" box="[853,873,1911,1934]" italics="true" pageId="31" pageNumber="32">in</emphasis>
elongated (8 µm); C—leg I; D—leg II; E—tarsus and tibia III; F—tarsus, tibia and genu IV; G—palpal tarsus, lateral and ventral sides with 4 barbed and basally fused eupathidia. 16A, CG—from Quintana Roo, Sian Kaan; 16B—from Italy, Tuscany.
</paragraph>
</caption>
<paragraph id="8BD69A07FFCC433E0EEDFF11FA1D87C0" blockId="32.[151,1437,151,359]" box="[151,1437,151,175]" pageId="32" pageNumber="33">
<taxonomicName id="4C69E184FFCC433E0EEDFF11FD8487C0" authority="Halbert, 1920: 140" authorityName="Halbert" authorityPageNumber="140" authorityYear="1920" box="[151,516,151,175]" class="Insecta" family="Hesperiidae" genus="Alicus" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="B91D4615FFCC433E0EEDFF11FEBB87C0" box="[151,315,151,175]" italics="true" pageId="32" pageNumber="33">Alicus oblongus</emphasis>
Halbert, 1920: 140
</taxonomicName>
, pl. 23, figs. 23a-c, erroneous orthography of
<taxonomicName id="4C69E184FFCC433E0DA0FF11FB1387C0" authority="C. L. Koch" authorityName="C. L. Koch" authorityYear="1842" box="[986,1171,151,175]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCC433E0DA0FF11FB9E87C0" box="[986,1054,151,175]" italics="true" pageId="32" pageNumber="33">Alycus</emphasis>
C.L. Koch
</taxonomicName>
;
<typeStatus id="54D224A5FFCC433E0AE4FF11FB7787C0" box="[1182,1271,151,175]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
from Malahide,
</paragraph>
<paragraph id="8BD69A07FFCC433E0EBDFF33FA1C8664" blockId="32.[151,1437,151,359]" pageId="32" pageNumber="33">
<collectingCountry id="F37EDA97FFCC433E0EBDFF33FE9187A2" box="[199,273,181,205]" name="Ireland" pageId="32" pageNumber="33">Ireland</collectingCountry>
, from orange lichen zone at NMI (National Museum of
<collectingCountry id="F37EDA97FFCC433E0D30FF33FC1687A2" box="[842,918,181,205]" name="Ireland" pageId="32" pageNumber="33">Ireland</collectingCountry>
).
<taxonomicName id="4C69E184FFCC433E0EEDFF52FE0A8783" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[151,394,212,236]" class="Arachnida" family="Alycidae" genus="Pachygnathus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="B91D4615FFCC433E0EEDFF52FE0A8783" box="[151,394,212,236]" italics="true" pageId="32" pageNumber="33">Pachygnathus oblongus</emphasis>
</taxonomicName>
.—
<bibRefCitation id="EFF8E7F6FFCC433E0FD2FF52FD4E8783" author="Thor, S. &amp; Willmann, C." box="[424,718,212,236]" pageId="32" pageNumber="33" pagination="1 - 186" refId="ref48675" refString="Thor, S. &amp; Willmann, C. (1941) Acarina. Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae. In: Das Tierreich. Vol. 71 a. Walter de Gruyter &amp; Co., Berlin-Leipzig, pp. 1-186. https://doi.org/10.1515/9783111434568" type="book chapter" year="1941">Thor &amp; Willmann 1941: 138</bibRefCitation>
;
<bibRefCitation id="EFF8E7F6FFCC433E0CA3FF52FC708783" author="Baker, R. A. &amp; Bayliss, R. A." box="[729,1008,212,236]" pageId="32" pageNumber="33" pagination="280 - 299" refId="ref46100" refString="Baker, R. A. &amp; Bayliss, R. A. (2005) A review and update of the mesostigmatid and prostigmatid mites of the marine littoral and supralittoral in Ireland based on the work of J. N. Halbert (1872 - 1948), including a ckecklist, revised nomenclature, collecting sites and recent records of mites. Bulletin of the Irish biogeographical Society, 29, 280-299." type="journal article" year="2005">Baker &amp; Bayliss 2005: 288</bibRefCitation>
.
<taxonomicName id="4C69E184FFCC433E0EEDFF75FD878664" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[151,519,243,267]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="oblongus" subGenus="Orthacarus">
<emphasis id="B91D4615FFCC433E0EEDFF75FD878664" box="[151,519,243,267]" italics="true" pageId="32" pageNumber="33">Amphialycus (Orthacarus) oblongus</emphasis>
</taxonomicName>
.—
<bibRefCitation id="EFF8E7F6FFCC433E0C5FFF75FD5C8665" author="Uusitalo, M." box="[549,732,242,267]" pageId="32" pageNumber="33" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010: 58</bibRefCitation>
, figs. 8287;
<bibRefCitation id="EFF8E7F6FFCC433E0D19FF75FB158664" author="Vazquez-Rojas, I. M. &amp; Vazquez-Gonzalez, M. M. &amp; Lopez-Campos, M. G." box="[867,1173,243,267]" pageId="32" pageNumber="33" pagination="19 - 24" refId="ref49158" refString="Vazquez-Rojas, I. M., Vazquez-Gonzalez, M. M. &amp; Lopez-Campos, M. G. (2016) Diversidad de Endeostigmata y Prostigmata (Acari: Trombidiformes) en el area protegida &quot; Jardin Botanico Alfredo Barrera Marin &quot;, Puerto Morelos, Quintana Roo, Mexico. Entomologia mexicana, 3, 19-24." type="journal article" year="2016">
Vázquez-Rojas
<emphasis id="B91D4615FFCC433E0A79FF75FBAF8664" box="[1027,1071,243,267]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. 2016: 21
</bibRefCitation>
;
<bibRefCitation id="EFF8E7F6FFCC433E0AE5FF75FA188664" author="Uusitalo, M. &amp; Ueckermann, E. A. &amp; Theron, P. D." box="[1183,1432,243,267]" pageId="32" pageNumber="33" pagination="301 - 340" refId="ref48918" refString="Uusitalo, M., Ueckermann, E. A. &amp; Theron, P. D. (2020) A review of the family Alycidae (Acari, Acariformes) from South Africa. Zootaxa, 4858 (3), 301-340. https://doi.org/10.11646/zootaxa.4858.3.1" type="journal article" year="2020">
Uusitalo
<emphasis id="B91D4615FFCC433E0A80FF75FAA68664" box="[1274,1318,243,267]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. 2020: 321
</bibRefCitation>
,
</paragraph>
<paragraph id="8BD69A07FFCC433E0EBDFE97FD388609" blockId="32.[151,1437,151,359]" pageId="32" pageNumber="33">
figs. 7689.
<taxonomicName id="4C69E184FFCC433E0EEDFEB6FEAF8627" box="[151,303,304,328]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="roseus">
<emphasis id="B91D4615FFCC433E0EEDFEB6FEAF8627" box="[151,303,304,328]" italics="true" pageId="32" pageNumber="33">Alycus? roseus</emphasis>
</taxonomicName>
.—
<bibRefCitation id="EFF8E7F6FFCC433E0F35FEB6FDB18627" author="Womersley, H." box="[335,561,304,328]" pageId="32" pageNumber="33" pagination="133 - 143" refId="ref49733" refString="Womersley, H. (1944) Australian Acarina, Families Alycidae and Nanorchestidae. Transactions of the Royal Society of South Australia, 68, 133-143." type="journal article" year="1944">Womersley 1944: 139</bibRefCitation>
.
<taxonomicName id="4C69E184FFCC433E0EEDFEC9FECE8608" authorityName="Zachvatkin" authorityYear="1949" box="[151,334,335,359]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="tremli">
<emphasis id="B91D4615FFCC433E0EEDFEC9FECE8608" box="[151,334,335,359]" italics="true" pageId="32" pageNumber="33">Orthacarus tremli</emphasis>
</taxonomicName>
.—
<bibRefCitation id="EFF8E7F6FFCC433E0F11FEC9FDCE8608" author="Zachvatkin, A. A." box="[363,590,335,359]" pageId="32" pageNumber="33" pagination="292 - 297" refId="ref49859" refString="Zachvatkin, A. A. (1949) New representatives of segmented mites (Acarina, Pachygnathidae). Entomologicheskoye Obozreniye, 30, 292-297. [in Russian]" type="journal article" year="1949">Zachvatkin 1949: 292</bibRefCitation>
, figs. 15.
</paragraph>
<paragraph id="8BD69A07FFCC433E0EEDFE13FCCA8698" blockId="32.[151,1437,405,1475]" pageId="32" pageNumber="33">
<emphasis id="B91D4615FFCC433E0EEDFE13FEA986C0" bold="true" box="[151,297,405,431]" pageId="32" pageNumber="33">Description.</emphasis>
<emphasis id="B91D4615FFCC433E0F4AFE10FE0A86C0" box="[304,394,406,431]" italics="true" pageId="32" pageNumber="33">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFCC433E0FC3FE13FDA986C0" box="[441,553,405,431]" count="2" pageId="32" pageNumber="33" type="female">2 females</specimenCount>
,
<figureCitation id="13528682FFCC433E0C4FFE13FD4886C0" box="[565,712,405,431]" captionStart="FIGURE 16" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[151,1435,232,1851]" captionTargetId="figure-23@31.[151,1435,232,1851]" captionTargetPageId="31" captionText="FIGURE 16. Orthacarus oblongus (Halbert). A—prodorsum, setae in elongated (11,5 µm); B—prodorsum, setae in elongated (8 µm); C—leg I; D—leg II; E—tarsus and tibia III; F—tarsus, tibia and genu IV; G—palpal tarsus, lateral and ventral sides with 4 barbed and basally fused eupathidia. 16A, CG—from Quintana Roo, Sian Kaan; 16B—from Italy, Tuscany." pageId="32" pageNumber="33">Figs. 16A, B</figureCitation>
). Length 300500 µm; neotrichous setal pattern dorsally and on ventral side; naso absent, prodorsal setae
<emphasis id="B91D4615FFCC433E0C1FFE3CFDF986BC" box="[613,633,442,467]" italics="true" pageId="32" pageNumber="33">vi</emphasis>
,
<emphasis id="B91D4615FFCC433E0CFDFE3CFD2B86BC" box="[647,683,442,467]" italics="true" pageId="32" pageNumber="33">sce</emphasis>
and
<emphasis id="B91D4615FFCC433E0C9AFE3CFC8786BC" box="[736,775,442,467]" italics="true" pageId="32" pageNumber="33">exp</emphasis>
sparsely ciliated (reduced), sensilla
<emphasis id="B91D4615FFCC433E0AE7FE3CFB3686BC" box="[1181,1206,442,467]" italics="true" pageId="32" pageNumber="33">ve</emphasis>
and
<emphasis id="B91D4615FFCC433E0A91FE3CFA8A86BC" box="[1259,1290,442,467]" italics="true" pageId="32" pageNumber="33">sci</emphasis>
filamentous, distance between counterparts of setae
<emphasis id="B91D4615FFCC433E0C37FE58FDE18698" box="[589,609,478,503]" italics="true" pageId="32" pageNumber="33">vi</emphasis>
exceptionally small.
</paragraph>
<paragraph id="8BD69A07FFCC433E0EBDFD84FAC18574" blockId="32.[151,1437,405,1475]" box="[199,1345,513,539]" pageId="32" pageNumber="33">
<taxonomicName id="4C69E184FFCC433E0EBDFD84FE8F8574" box="[199,271,514,539]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFCC433E0EBDFD84FE8F8574" box="[199,271,514,539]" italics="true" pageId="32" pageNumber="33">Venter</emphasis>
</taxonomicName>
. Genital valves each with 1218 genital setae; anal valves terminally with 78 setae,
<emphasis id="B91D4615FFCC433E0AB7FD87FAC18574" bold="true" box="[1229,1345,513,539]" pageId="32" pageNumber="33">Remarks.</emphasis>
</paragraph>
<paragraph id="8BD69A07FFCC433E0EBDFDA3FAEE850C" blockId="32.[151,1437,405,1475]" pageId="32" pageNumber="33">
<taxonomicName id="4C69E184FFCC433E0EBDFDA3FED68550" authorityName="Krefft" authorityYear="1871" box="[199,342,549,575]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="32" pageNumber="33" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFCC433E0EBDFDA3FED68550" box="[199,342,549,575]" italics="true" pageId="32" pageNumber="33">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFCC433E0F1EFDA3FE4B8550" box="[356,459,549,575]" captionStart="FIGURE 16" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[151,1435,232,1851]" captionTargetId="figure-23@31.[151,1435,232,1851]" captionTargetPageId="31" captionText="FIGURE 16. Orthacarus oblongus (Halbert). A—prodorsum, setae in elongated (11,5 µm); B—prodorsum, setae in elongated (8 µm); C—leg I; D—leg II; E—tarsus and tibia III; F—tarsus, tibia and genu IV; G—palpal tarsus, lateral and ventral sides with 4 barbed and basally fused eupathidia. 16A, CG—from Quintana Roo, Sian Kaan; 16B—from Italy, Tuscany." pageId="32" pageNumber="33">Fig. 16G</figureCitation>
). Pair of smooth cheliceral setae abaxially; rutella widening to apical lobes and a pointed process, one pair of adoral setae; distally narrow palpal tarsi with four basally fused, microtrichous eupathidia.
</paragraph>
<paragraph id="8BD69A07FFCC433E0EBDFDE8FC2C85C4" blockId="32.[151,1437,405,1475]" pageId="32" pageNumber="33">
<emphasis id="B91D4615FFCC433E0EBDFDE8FF7C85E8" box="[199,252,622,647]" italics="true" pageId="32" pageNumber="33">Legs</emphasis>
(
<figureCitation id="13528682FFCC433E0F76FDEBFE1D85E8" box="[268,413,621,647]" captionStart="FIGURE 16" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[151,1435,232,1851]" captionTargetId="figure-23@31.[151,1435,232,1851]" captionTargetPageId="31" captionText="FIGURE 16. Orthacarus oblongus (Halbert). A—prodorsum, setae in elongated (11,5 µm); B—prodorsum, setae in elongated (8 µm); C—leg I; D—leg II; E—tarsus and tibia III; F—tarsus, tibia and genu IV; G—palpal tarsus, lateral and ventral sides with 4 barbed and basally fused eupathidia. 16A, CG—from Quintana Roo, Sian Kaan; 16B—from Italy, Tuscany." pageId="32" pageNumber="33">Figs. 16CF</figureCitation>
). Claw-like empodium multisetulate; solenidial formula for tarsi, tibiae, genua and femora of legs I, II, III and IV, respectively: 2-1-0-0, 2-2-2-0, 3-2-0-1, 1-0-0-0
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCC433E0EBDFD33FCB984EC" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BD69A07FFCC433E0EBDFD33FCB984EC" blockId="32.[151,1437,405,1475]" pageId="32" pageNumber="33">
<emphasis id="B91D4615FFCC433E0EBDFD33FE3085A0" bold="true" box="[199,432,693,719]" pageId="32" pageNumber="33">Material examined.</emphasis>
<materialsCitation id="3B01905AFFCC433E0FCFFD33FE46859C" location="Sian Ka'an" pageId="32" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Quintana Roo">
<specimenCount id="9D6F518EFFCC433E0FCFFD33FD9585A0" box="[437,533,693,719]" count="1" pageId="32" pageNumber="33" type="female">1 female</specimenCount>
(on slide) from leaf litter (hojarasca), floodable low forest (selva baja inundable),
<location id="8EB6CCDCFFCC433E0EEDFD5FFE99859C" LSID="urn:lsid:plazi:treatment:03C02B11FFF3433E0EEDFF11FCEF82AC:8EB6CCDCFFCC433E0EEDFD5FFE99859C" box="[151,281,729,755]" name="Sian Ka'an" pageId="32" pageNumber="33" stateProvince="Quintana Roo">Sian Kaan</location>
,
<collectingRegion id="49AD54E5FFCC433E0F5CFD5FFE46859C" box="[294,454,729,755]" country="Mexico" name="Quintana Roo" pageId="32" pageNumber="33">Quintana Roo</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B01905AFFCC433E0FAEFD5FFF2E8478" collectingDate="1996-11-15" collectorName="Maria Magdalena Vazquez" country="Mexico" location="Mexico" pageId="32" pageNumber="33" specimenCount="1">
<collectingCountry id="F37EDA97FFCC433E0FAEFD5FFDAC859C" box="[468,556,729,755]" name="Mexico" pageId="32" pageNumber="33">Mexico</collectingCountry>
,
<date id="FFD7BCC7FFCC433E0C43FD5FFC98859C" box="[569,792,729,756]" pageId="32" pageNumber="33" value="1996-11-15">
<collectingDate id="EF93452FFFCC433E0C43FD5FFC98859C" box="[569,792,729,756]" pageId="32" pageNumber="33" value="1996-11-15">15 November 1996</collectingDate>
</date>
,
<collectorName id="269CFFD1FFCC433E0D5FFD5FFBDB859C" box="[805,1115,729,755]" pageId="32" pageNumber="33">María Magdalena Vázquez</collectorName>
; several high-quality photos of
</materialsCitation>
<specimenCount id="9D6F518EFFCC433E0ECCFD7BFEAD8478" box="[182,301,765,791]" count="2" pageId="32" pageNumber="33" type="adult">two adults</specimenCount>
, 1 protonymph and
<materialsCitation id="3B01905AFFCC433E0C74FD7BFCCE8454" collectionCode="JBPM" collectorName="Puerto" country="Mexico" latitude="20.823584" location="Jardin Botanico de Puerto" longLatPrecision="1" longitude="-86.898224" pageId="32" pageNumber="33" specimenCount="1" specimenCount-larva="1" stateProvince="Morelos">
<specimenCount id="9D6F518EFFCC433E0C74FD7BFDDB8478" box="[526,603,765,791]" count="1" pageId="32" pageNumber="33" type="larva">1 larva</specimenCount>
from coastal dune, leaf litter,
<collectionCode id="ED7802C2FFCC433E0DD7FD7BFC738478" box="[941,1011,765,791]" pageId="32" pageNumber="33">JBPM</collectionCode>
(
<location id="8EB6CCDCFFCC433E0A78FD7BFAA78478" LSID="urn:lsid:plazi:treatment:03C02B11FFF3433E0EEDFF11FCEF82AC:8EB6CCDCFFCC433E0A78FD7BFAA78478" box="[1026,1319,765,791]" country="Mexico" latitude="20.823584" longLatPrecision="1" longitude="-86.898224" name="Jardin Botanico de Puerto" pageId="32" pageNumber="33" stateProvince="Morelos">Jardin Botánico de Puerto</location>
<collectingRegion id="49AD54E5FFCC433E0B55FD7BFA118478" box="[1327,1425,765,791]" country="Mexico" name="Morelos" pageId="32" pageNumber="33">Morelos</collectingRegion>
),
<geoCoordinate id="EE5DFCC0FFCC433E0EEDFCA7FEB98454" box="[151,313,801,827]" degrees="20" direction="north" minutes="49" orientation="latitude" pageId="32" pageNumber="33" precision="1" seconds="24,9" value="20.823584">20°4924,9N</geoCoordinate>
<geoCoordinate id="EE5DFCC0FFCC433E0F3AFCA7FE678454" box="[320,487,801,828]" degrees="86" direction="west" minutes="53" orientation="longitude" pageId="32" pageNumber="33" precision="1" seconds="53,6" value="-86.898224">86°5353,6W</geoCoordinate>
,
<collectorName id="269CFFD1FFCC433E0F8EFCA7FDBD8454" box="[500,573,801,827]" pageId="32" pageNumber="33">Puerto</collectorName>
<collectingRegion id="49AD54E5FFCC433E0C3EFCA7FD248454" box="[580,676,801,827]" country="Mexico" name="Morelos" pageId="32" pageNumber="33">Morelos</collectingRegion>
,
<collectingRegion id="49AD54E5FFCC433E0CCAFCA7FCCE8454" box="[688,846,801,827]" country="Mexico" name="Quintana Roo" pageId="32" pageNumber="33">Quintana Roo</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B01905AFFCC433E0D20FCA7FCB584EC" collectingDate="2015-04-30" collectionCode="LAAH, UNAM" collectorName="Maria Magdalena Vazquez. Material &amp; de Acarologia &amp; Anita Hoffmann" country="Mexico" location="Universidad Nacional Autonoma" municipality="Faculty of Sciences" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Distrito Federal">
<collectingCountry id="F37EDA97FFCC433E0D20FCA7FC328454" box="[858,946,801,827]" name="Mexico" pageId="32" pageNumber="33">Mexico</collectingCountry>
,
<date id="FFD7BCC7FFCC433E0DC7FCA7FBDC8453" box="[957,1116,801,828]" pageId="32" pageNumber="33" value="2015-04-30">
<collectingDate id="EF93452FFFCC433E0DC7FCA7FBDC8453" box="[957,1116,801,828]" pageId="32" pageNumber="33" value="2015-04-30">30 April 2015</collectingDate>
</date>
,
<collectorName id="269CFFD1FFCC433E0A1DFCA7FF768430" pageId="32" pageNumber="33">María Magdalena Vázquez. Material</collectorName>
is deposited in the
<collectionCode id="ED7802C2FFCC433E0FABFCC3FD9F8430" box="[465,543,837,863]" pageId="32" pageNumber="33">LAAH</collectionCode>
(Laboratorio
<collectorName id="269CFFD1FFCC433E0CC1FCC3FCD78430" box="[699,855,837,863]" pageId="32" pageNumber="33">de Acarología</collectorName>
<collectorName id="269CFFD1FFCC433E0D13FCC3FBAB8430" box="[873,1067,837,863]" pageId="32" pageNumber="33">Anita Hoffmann</collectorName>
”),
<collectingMunicipality id="6BB2007DFFCC433E0A32FCC3FAA48430" box="[1096,1316,837,863]" pageId="32" pageNumber="33">Faculty of Sciences</collectingMunicipality>
at
<collectionCode id="ED7802C2FFCC433E0B3DFCC3FA1D8430" box="[1351,1437,837,863]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="32" pageNumber="33">UNAM</collectionCode>
(
<location id="8EB6CCDCFFCC433E0EE5FCEFFD9184EC" LSID="urn:lsid:plazi:treatment:03C02B11FFF3433E0EEDFF11FCEF82AC:8EB6CCDCFFCC433E0EE5FCEFFD9184EC" box="[159,529,873,899]" country="Mexico" municipality="Faculty of Sciences" name="Universidad Nacional Autonoma" pageId="32" pageNumber="33" stateProvince="Distrito Federal">Universidad Nacional Autonóma</location>
de
<collectingCountry id="F37EDA97FFCC433E0C40FCEFFD1484EC" box="[570,660,873,899]" name="Mexico" pageId="32" pageNumber="33">México</collectingCountry>
),
<collectingRegion id="49AD54E5FFCC433E0CDCFCEFFCB584EC" box="[678,821,873,899]" country="Mexico" name="Distrito Federal" pageId="32" pageNumber="33">Mexico City</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCC433E0EBDFC0BFB45837C" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BD69A07FFCC433E0EBDFC0BFB45837C" blockId="32.[151,1437,405,1475]" pageId="32" pageNumber="33">
<emphasis id="B91D4615FFCC433E0EBDFC0BFE4D84C8" bold="true" box="[199,461,909,935]" pageId="32" pageNumber="33">Differential diagnosis.</emphasis>
This species differs from other neotrichous species of the genus
<taxonomicName id="4C69E184FFCC433E0AEDFC0BFAA884C8" authorityName="Zachvatkin" authorityYear="1949" box="[1175,1320,909,935]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCC433E0AEDFC0BFAA884C8" box="[1175,1320,909,935]" italics="true" pageId="32" pageNumber="33">Amphialycus</emphasis>
</taxonomicName>
by having the edaphomorphisms listed in the diagnosis of the genus. This species differs from another (still undescribed) species of the genus
<taxonomicName id="4C69E184FFCC433E0FF5FC53FD908480" authorityName="Zachvatkin" authorityYear="1949" box="[399,528,981,1007]" class="Arachnida" family="Alycidae" genus="Orthacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFCC433E0FF5FC53FD908480" box="[399,528,981,1007]" italics="true" pageId="32" pageNumber="33">Orthacarus</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFCC433E0C59FC53FDF8849F" box="[547,632,981,1008]" captionStart="FIGURE 54" captionStartId="86.[151,250,1908,1932]" captionTargetBox="[151,1436,176,1884]" captionTargetId="figure-24@86.[151,1436,176,1884]" captionTargetPageId="86" captionText="FIGURE 54. Selected figures of an undescribed species clu from South Africa. A—prodorsum; B—dorsum; C—dorsal seta; D—venter; E—genital valves; F—palpal eupathidia; G—Leg I; H—Empodium." pageId="32" pageNumber="33">Fig. 54</figureCitation>
) by having both pairs of prodorsal sensilla (
<emphasis id="B91D4615FFCC433E0AFDFC50FB208480" box="[1159,1184,982,1007]" italics="true" pageId="32" pageNumber="33">ve</emphasis>
and
<emphasis id="B91D4615FFCC433E0AA4FC50FB7D8480" box="[1246,1277,982,1007]" italics="true" pageId="32" pageNumber="33">sci</emphasis>
) filamentous, abaxially inserted cheliceral setae and the solenidial pattern as expressed in
<emphasis id="B91D4615FFCC433E0D93FC7FFBF3837C" bold="true" box="[1001,1139,1017,1043]" pageId="32" pageNumber="33">Description</emphasis>
above.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFCC433E0EBDFB9BFCEF82AC" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BD69A07FFCC433E0EBDFB9BFCEF82AC" blockId="32.[151,1437,405,1475]" pageId="32" pageNumber="33">
<emphasis id="B91D4615FFCC433E0EBDFB9BFEBB8358" bold="true" box="[199,315,1053,1079]" pageId="32" pageNumber="33">Remarks.</emphasis>
Detailed figures of the dorsum, venter, various organs, and juvenal stages of
<taxonomicName id="4C69E184FFCC433E0A9BFB9BFAF08358" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[1249,1392,1053,1079]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="oblongus">
<emphasis id="B91D4615FFCC433E0A9BFB9BFAF08358" box="[1249,1392,1053,1079]" italics="true" pageId="32" pageNumber="33">O. oblongus</emphasis>
</taxonomicName>
[as
<taxonomicName id="4C69E184FFCC433E0EEDFBC7FDB78334" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[151,567,1089,1115]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="oblongus" subGenus="Orthacarus">
<emphasis id="B91D4615FFCC433E0EEDFBC7FDB78334" box="[151,567,1089,1115]" italics="true" pageId="32" pageNumber="33">Amphialycus (Orthacarus) oblongus</emphasis>
</taxonomicName>
] has already been published twice - from Europe (
<bibRefCitation id="EFF8E7F6FFCC433E0AF3FBC7FADC8333" author="Uusitalo, M." box="[1161,1372,1089,1116]" pageId="32" pageNumber="33" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010: 58</bibRefCitation>
, figs. 8289) and from
<collectingCountry id="F37EDA97FFCC433E0F1AFBE3FE708310" box="[352,496,1125,1151]" name="South Africa" pageId="32" pageNumber="33">South Africa</collectingCountry>
(
<bibRefCitation id="EFF8E7F6FFCC433E0C7AFBE3FCA18310" author="Uusitalo, M. &amp; Ueckermann, E. A. &amp; Theron, P. D." box="[512,801,1125,1151]" pageId="32" pageNumber="33" pagination="301 - 340" refId="ref48918" refString="Uusitalo, M., Ueckermann, E. A. &amp; Theron, P. D. (2020) A review of the family Alycidae (Acari, Acariformes) from South Africa. Zootaxa, 4858 (3), 301-340. https://doi.org/10.11646/zootaxa.4858.3.1" type="journal article" year="2020">
Uusitalo
<emphasis id="B91D4615FFCC433E0C10FBE0FD1D8310" box="[618,669,1125,1151]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. 2020: 321
</bibRefCitation>
, figs. 7689). The figures in the present study focus on illustrating leg solenidia (black arrows) and variation of prodorsal setae. The reduced setae
<emphasis id="B91D4615FFCC433E0AE6FB0CFB3083CC" box="[1180,1200,1162,1187]" italics="true" pageId="32" pageNumber="33">vi</emphasis>
,
<emphasis id="B91D4615FFCC433E0AC5FB0CFB6383CC" box="[1215,1251,1162,1187]" italics="true" pageId="32" pageNumber="33">sce</emphasis>
and
<emphasis id="B91D4615FFCC433E0B60FB0CFAC183CC" box="[1306,1345,1162,1187]" italics="true" pageId="32" pageNumber="33">exp</emphasis>
are less ciliated in specimens from the European and South African populations of
<taxonomicName id="4C69E184FFCC433E0D87FB2BFA1D83A8" baseAuthorityName="Halbert" baseAuthorityYear="1920" box="[1021,1437,1197,1223]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="oblongus" subGenus="Orthacarus">
<emphasis id="B91D4615FFCC433E0D87FB2BFA1D83A8" box="[1021,1437,1197,1223]" italics="true" pageId="32" pageNumber="33">Amphialycus (Orthacarus) oblongus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF8E7F6FFCC433E0EE5FB57FEFA8384" author="Uusitalo, M." box="[159,378,1233,1259]" pageId="32" pageNumber="33" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010: 127</bibRefCitation>
, fig. 82 and
<bibRefCitation id="EFF8E7F6FFCC433E0C7CFB57FCA08384" author="Uusitalo, M. &amp; Ueckermann, E. A. &amp; Theron, P. D." box="[518,800,1233,1259]" pageId="32" pageNumber="33" pagination="301 - 340" refId="ref48918" refString="Uusitalo, M., Ueckermann, E. A. &amp; Theron, P. D. (2020) A review of the family Alycidae (Acari, Acariformes) from South Africa. Zootaxa, 4858 (3), 301-340. https://doi.org/10.11646/zootaxa.4858.3.1" type="journal article" year="2020">
Uusitalo
<emphasis id="B91D4615FFCC433E0C17FB54FD1E8384" box="[621,670,1233,1259]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. 2020: 322
</bibRefCitation>
, fig. 76), and the lengths of the columnar and bushy seta
<emphasis id="B91D4615FFCC433E0EEDFB70FF2D8260" box="[151,173,1270,1295]" italics="true" pageId="32" pageNumber="33">in</emphasis>
also varies when compared to the Mexican specimen (
<figureCitation id="13528682FFCC433E0D59FB73FC0D8260" box="[803,909,1269,1295]" captionStart="FIGURE 16" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[151,1435,232,1851]" captionTargetId="figure-23@31.[151,1435,232,1851]" captionTargetPageId="31" captionText="FIGURE 16. Orthacarus oblongus (Halbert). A—prodorsum, setae in elongated (11,5 µm); B—prodorsum, setae in elongated (8 µm); C—leg I; D—leg II; E—tarsus and tibia III; F—tarsus, tibia and genu IV; G—palpal tarsus, lateral and ventral sides with 4 barbed and basally fused eupathidia. 16A, CG—from Quintana Roo, Sian Kaan; 16B—from Italy, Tuscany." pageId="32" pageNumber="33">Fig. 16A</figureCitation>
). However, the ciliation of the prodorsal setae in a specimen from European unpublished material (
<figureCitation id="13528682FFCC433E0C88FA9FFCDA825C" box="[754,858,1305,1331]" captionStart="FIGURE 16" captionStartId="31.[151,250,1874,1898]" captionTargetBox="[151,1435,232,1851]" captionTargetId="figure-23@31.[151,1435,232,1851]" captionTargetPageId="31" captionText="FIGURE 16. Orthacarus oblongus (Halbert). A—prodorsum, setae in elongated (11,5 µm); B—prodorsum, setae in elongated (8 µm); C—leg I; D—leg II; E—tarsus and tibia III; F—tarsus, tibia and genu IV; G—palpal tarsus, lateral and ventral sides with 4 barbed and basally fused eupathidia. 16A, CG—from Quintana Roo, Sian Kaan; 16B—from Italy, Tuscany." pageId="32" pageNumber="33">Fig. 16B</figureCitation>
, from
<collectingCountry id="F37EDA97FFCC433E0DDDFA9FFC5C825C" box="[935,988,1305,1331]" name="Italy" pageId="32" pageNumber="33">Italy</collectingCountry>
) corresponds well to those form of the Mexican counterparts. Considering that the solenidial formulae are similar in specimens from the three continents - despite limited material - it is fair to assume that the observed differences are due to intraspecific variation and the specimens are conspecific. This species has also been reported from
<collectingCountry id="F37EDA97FFCC433E0DEEFA03FC7B82F0" box="[916,1019,1413,1439]" name="Australia" pageId="32" pageNumber="33">Australia</collectingCountry>
as
<taxonomicName id="4C69E184FFCC433E0A65FA03FB4782F0" box="[1055,1223,1413,1439]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="roseus">
<emphasis id="B91D4615FFCC433E0A65FA03FB4782F0" box="[1055,1223,1413,1439]" italics="true" pageId="32" pageNumber="33">Alycus? roseus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF8E7F6FFCC433E0AADFA03FF4382AC" author="Womersley, H." pageId="32" pageNumber="33" pagination="133 - 143" refId="ref49733" refString="Womersley, H. (1944) Australian Acarina, Families Alycidae and Nanorchestidae. Transactions of the Royal Society of South Australia, 68, 133-143." type="journal article" year="1944">Womersley 1944: 139</bibRefCitation>
, fig. 3FI), and it may have an anthropochorous distribution.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,255 @@
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<paragraph id="8BD69A07FFF543070EEDFC03FDFD84F0" blockId="25.[151,637,901,963]" box="[151,637,901,928]" pageId="25" pageNumber="26">
<heading id="D09E2D6BFFF543070EEDFC03FDFD84F0" bold="true" box="[151,637,901,928]" fontSize="11" level="2" pageId="25" pageNumber="26" reason="4">
<taxonomicName id="4C69E184FFF543070EEDFC03FDFD84F0" authority="Uusitalo, 2020" authorityName="Uusitalo" authorityYear="2020" box="[151,637,901,928]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="holarcticus">
<emphasis id="B91D4615FFF543070EEDFC03FDFD84F0" bold="true" box="[151,637,901,928]" pageId="25" pageNumber="26">
<emphasis id="B91D4615FFF543070EEDFC03FE4484CF" bold="true" box="[151,452,901,928]" italics="true" pageId="25" pageNumber="26">Amphialycus holarcticus</emphasis>
Uusitalo, 2020
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C373C98CFFF543070EEDFC2FFA9982B5" pageId="25" pageNumber="26" type="description">
<paragraph id="8BD69A07FFF543070EEDFC2FFE1884AC" blockId="25.[151,637,901,963]" box="[151,408,937,963]" pageId="25" pageNumber="26">
(
<figureCitation id="13528682FFF543070EE5FC2FFEB184AC" box="[159,305,937,963]" captionStart="FIGURE 12" captionStartId="26.[151,250,1863,1887]" captionTargetBox="[155,1436,153,1839]" captionTargetId="figure-24@26.[151,1436,153,1839]" captionTargetPageId="26" captionText="FIGURE 12. Amphialycus holarcticus Uusitalo. A—prodorsum, naso cone-shaped and longitudinally striated, dorsal segment C of the depicted specimen artificially folded over posterior parts of prodorsal shield; B—dorsal habitus; C—subcapitulum, rutella, 2 pairs of cheliceral setae, chb simple and cha forked (arrows), palpi with 4 pairs of microtrichous eupathidia; D—genital area with 3537 genital setae per valve, 1 pair of eugenital setae (female); E—anal area, 7 anal setae per valve. All specimens—from Utah, Cache Co." pageId="25" pageNumber="26">Figs. 12AE</figureCitation>
,
<figureCitation id="13528682FFF543070F46FC2FFE1084AC" box="[316,400,937,963]" captionStart="FIGURE 13" captionStartId="27.[151,250,1875,1899]" captionTargetBox="[151,1435,193,1851]" captionTargetId="figure-15@27.[151,1436,193,1851]" captionTargetPageId="27" captionText="FIGURE 13. Amphialycus holarcticus Uusitalo. A—leg I; B—leg II, famulus II abaxially at the level of solenidion ω1; C—tibia and genu III; D—tibia, genu, telofemur and basifemur IV. All specimens—from Utah, Cache Co." pageId="25" pageNumber="26">13AD</figureCitation>
)
</paragraph>
<paragraph id="8BD69A07FFF543070EEDFC68FB0A8369" blockId="25.[151,1162,1006,1067]" box="[151,1162,1006,1030]" pageId="25" pageNumber="26">
<taxonomicName id="4C69E184FFF543070EEDFC68FD7C8369" authority="Uusitalo, 2020: 109" authorityName="Uusitalo" authorityPageNumber="109" authorityYear="2020" box="[151,764,1006,1030]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="holarcticus" subGenus="Amphialycus">
<emphasis id="B91D4615FFF543070EEDFC68FDAB8369" box="[151,555,1006,1030]" italics="true" pageId="25" pageNumber="26">Amphialycus (Amphialycus) holarcticus</emphasis>
Uusitalo, 2020: 109
</taxonomicName>
, figs. 110;
<typeStatus id="54D224A5FFF543070D00FC68FC538369" box="[890,979,1006,1030]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
from
<taxonomicName id="4C69E184FFF543070A6AFC68FBC7836A" authorityName="Hill" authorityYear="1753" box="[1040,1095,1006,1029]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFF543070A6AFC68FBC7836A" box="[1040,1095,1006,1029]" italics="true" pageId="25" pageNumber="26">Larix</emphasis>
</taxonomicName>
stand,
</paragraph>
<paragraph id="8BD69A07FFF543070EEDFB97FE1C8345" blockId="25.[151,1162,1006,1067]" box="[151,412,1040,1067]" pageId="25" pageNumber="26">
<collectingRegion id="49AD54E5FFF543070EEDFB97FF508344" box="[151,208,1041,1067]" country="Russia" name="Tyva" pageId="25" pageNumber="26">Tuva</collectingRegion>
Republic,
<collectingCountry id="F37EDA97FFF543070F30FB96FE188345" box="[330,408,1040,1066]" name="Russia" pageId="25" pageNumber="26">Russia</collectingCountry>
.
</paragraph>
<paragraph id="8BD69A07FFF543070EEDFBDEFC3483B1" blockId="25.[151,1437,1112,2039]" pageId="25" pageNumber="26">
<emphasis id="B91D4615FFF543070EEDFBDEFEA9831D" bold="true" box="[151,297,1112,1138]" pageId="25" pageNumber="26">Description.</emphasis>
<emphasis id="B91D4615FFF543070F4BFBDFFE0B831D" box="[305,395,1113,1138]" italics="true" pageId="25" pageNumber="26">Dorsum</emphasis>
(n=
<specimenCount id="9D6F518EFFF543070FC4FBDEFD9F831D" box="[446,543,1112,1138]" count="1" pageId="25" pageNumber="26" type="female">1 female</specimenCount>
,
<figureCitation id="13528682FFF543070C51FBDEFD3E831D" box="[555,702,1112,1138]" captionStart="FIGURE 12" captionStartId="26.[151,250,1863,1887]" captionTargetBox="[155,1436,153,1839]" captionTargetId="figure-24@26.[151,1436,153,1839]" captionTargetPageId="26" captionText="FIGURE 12. Amphialycus holarcticus Uusitalo. A—prodorsum, naso cone-shaped and longitudinally striated, dorsal segment C of the depicted specimen artificially folded over posterior parts of prodorsal shield; B—dorsal habitus; C—subcapitulum, rutella, 2 pairs of cheliceral setae, chb simple and cha forked (arrows), palpi with 4 pairs of microtrichous eupathidia; D—genital area with 3537 genital setae per valve, 1 pair of eugenital setae (female); E—anal area, 7 anal setae per valve. All specimens—from Utah, Cache Co." pageId="25" pageNumber="26">Figs. 12A, B</figureCitation>
). Length 560700 µm; ciliate setae with longer central points on caudal segments. Prodorsum with: naso prominent, cone-shaped, longitudinally striated; pilose setae
<emphasis id="B91D4615FFF543070A81FBFBFA8F83F9" box="[1275,1295,1149,1174]" italics="true" pageId="25" pageNumber="26">vi</emphasis>
,
<emphasis id="B91D4615FFF543070B67FBFBFAC183F9" box="[1309,1345,1149,1174]" italics="true" pageId="25" pageNumber="26">sce</emphasis>
and
<emphasis id="B91D4615FFF543070B0FFBFBFA1C83F9" box="[1397,1436,1149,1174]" italics="true" pageId="25" pageNumber="26">exp</emphasis>
on hard integument,
<emphasis id="B91D4615FFF543070FFBFB27FE2883D5" box="[385,424,1185,1210]" italics="true" pageId="25" pageNumber="26">exp</emphasis>
smallest and anterior to eyes, setae
<emphasis id="B91D4615FFF543070D3AFB27FCD683D5" box="[832,854,1185,1210]" italics="true" pageId="25" pageNumber="26">in</emphasis>
pubescent; two pairs of filamentous sensilla
<emphasis id="B91D4615FFF543070B29FB27FAEC83D5" box="[1363,1388,1185,1210]" italics="true" pageId="25" pageNumber="26">ve</emphasis>
and
<emphasis id="B91D4615FFF543070EEDFB43FF3683B1" box="[151,182,1221,1246]" italics="true" pageId="25" pageNumber="26">sci</emphasis>
ciliated along entire length; both pairs of lateral eyes finely striated.
</paragraph>
<paragraph id="8BD69A07FFF543070EBDFB6FFB648249" blockId="25.[151,1437,1112,2039]" pageId="25" pageNumber="26">
<taxonomicName id="4C69E184FFF543070EBDFB6FFE8F826D" box="[199,271,1257,1282]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFF543070EBDFB6FFE8F826D" box="[199,271,1257,1282]" italics="true" pageId="25" pageNumber="26">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFF543070F64FB6EFE32826D" box="[286,434,1256,1282]" captionStart="FIGURE 12" captionStartId="26.[151,250,1863,1887]" captionTargetBox="[155,1436,153,1839]" captionTargetId="figure-24@26.[151,1436,153,1839]" captionTargetPageId="26" captionText="FIGURE 12. Amphialycus holarcticus Uusitalo. A—prodorsum, naso cone-shaped and longitudinally striated, dorsal segment C of the depicted specimen artificially folded over posterior parts of prodorsal shield; B—dorsal habitus; C—subcapitulum, rutella, 2 pairs of cheliceral setae, chb simple and cha forked (arrows), palpi with 4 pairs of microtrichous eupathidia; D—genital area with 3537 genital setae per valve, 1 pair of eugenital setae (female); E—anal area, 7 anal setae per valve. All specimens—from Utah, Cache Co." pageId="25" pageNumber="26">Figs. 12D, E</figureCitation>
). Setae shorter and less ciliated than dorsal setae; genital setae of various lengths, mostly elongated closer to mesial margin, 3545 per valve; anal setae 59 per valve, caudal setae longest.
</paragraph>
<paragraph id="8BD69A07FFF543070EBDFAB6FB7482FC" blockId="25.[151,1437,1112,2039]" pageId="25" pageNumber="26">
<taxonomicName id="4C69E184FFF543070EBDFAB6FED68225" authorityName="Krefft" authorityYear="1871" box="[199,342,1328,1354]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="25" pageNumber="26" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFF543070EBDFAB6FED68225" box="[199,342,1328,1354]" italics="true" pageId="25" pageNumber="26">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFF543070F1CFAB6FE4E8225" box="[358,462,1328,1354]" captionStart="FIGURE 12" captionStartId="26.[151,250,1863,1887]" captionTargetBox="[155,1436,153,1839]" captionTargetId="figure-24@26.[151,1436,153,1839]" captionTargetPageId="26" captionText="FIGURE 12. Amphialycus holarcticus Uusitalo. A—prodorsum, naso cone-shaped and longitudinally striated, dorsal segment C of the depicted specimen artificially folded over posterior parts of prodorsal shield; B—dorsal habitus; C—subcapitulum, rutella, 2 pairs of cheliceral setae, chb simple and cha forked (arrows), palpi with 4 pairs of microtrichous eupathidia; D—genital area with 3537 genital setae per valve, 1 pair of eugenital setae (female); E—anal area, 7 anal setae per valve. All specimens—from Utah, Cache Co." pageId="25" pageNumber="26">Fig. 12C</figureCitation>
). Two pairs of cheliceral setae, anterior pair abaxial and smooth, posterior pair forked, in more dorsal position; rutella widen to a round hyaline membrane with two lobes apically; three pairs of well-developed adoral setae; four microtrichous eupathidia on palps close to each other but clearly apart.
</paragraph>
<paragraph id="8BD69A07FFF543070EBDFA1BFA9982B5" blockId="25.[151,1437,1112,2039]" pageId="25" pageNumber="26">
<emphasis id="B91D4615FFF543070EBDFA1BFF7C82D9" box="[199,252,1437,1462]" italics="true" pageId="25" pageNumber="26">Legs</emphasis>
(
<figureCitation id="13528682FFF543070F76FA1AFE2382D9" box="[268,419,1436,1462]" captionStart="FIGURE 13" captionStartId="27.[151,250,1875,1899]" captionTargetBox="[151,1435,193,1851]" captionTargetId="figure-15@27.[151,1436,193,1851]" captionTargetPageId="27" captionText="FIGURE 13. Amphialycus holarcticus Uusitalo. A—leg I; B—leg II, famulus II abaxially at the level of solenidion ω1; C—tibia and genu III; D—tibia, genu, telofemur and basifemur IV. All specimens—from Utah, Cache Co." pageId="25" pageNumber="26">Figs. 13AD</figureCitation>
). Pedal setae elongated; solenidial formula for tarsi, tibiae, genua and femora on legs I-II- III-IV, respectively: 2-1-0-0, 3-2-2-0, 3-2-2-1, 2-0-0-1+0, tarsi I and II with spine-like abaxial famulus.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF543070EBDFA62FF748071" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BD69A07FFF543070EBDFA62FE6181E1" blockId="25.[151,1437,1112,2039]" pageId="25" pageNumber="26">
<emphasis id="B91D4615FFF543070EBDFA62FE328291" bold="true" box="[199,434,1508,1534]" pageId="25" pageNumber="26">Material examined.</emphasis>
<typeStatus id="54D224A5FFF543070FC0FA62FDA28291" box="[442,546,1508,1534]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
female from
<taxonomicName id="4C69E184FFF543070CC4FA63FD7B8291" authorityName="Hill" authorityYear="1753" box="[702,763,1509,1534]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFF543070CC4FA63FD7B8291" box="[702,763,1509,1534]" italics="true" pageId="25" pageNumber="26">Larix</emphasis>
</taxonomicName>
stand,
<quantity id="4C9137E2FFF543070D3FFA62FC0D8290" box="[837,909,1508,1535]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="m" value="100.0">100 m</quantity>
away from the river, on north slope, litter and soil of
<taxonomicName id="4C69E184FFF543070E91F98FFEA8814D" authorityName="Hill" authorityYear="1753" box="[235,296,1545,1570]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFF543070E91F98FFEA8814D" box="[235,296,1545,1570]" italics="true" pageId="25" pageNumber="26">Larix</emphasis>
</taxonomicName>
,
<taxonomicName id="4C69E184FFF543070F42F98EFE5D814D" box="[312,477,1544,1570]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFF543070F42F98EFE5D814D" box="[312,477,1544,1570]" italics="true" pageId="25" pageNumber="26">Rhododendron</emphasis>
</taxonomicName>
,
<taxonomicName id="4C69E184FFF543070F94F98EFDC5814D" box="[494,581,1544,1570]" class="Magnoliopsida" family="Rosaceae" genus="Spiraea" kingdom="Plantae" order="Rosales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFF543070F94F98EFDC5814D" box="[494,581,1544,1570]" italics="true" pageId="25" pageNumber="26">Spiraea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C69E184FFF543070C2CF98EFD3A814D" box="[598,698,1544,1570]" class="Magnoliopsida" family="Ranunculaceae" genus="Clematis" kingdom="Plantae" order="Ranunculales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFF543070C2CF98EFD3A814D" box="[598,698,1544,1570]" italics="true" pageId="25" pageNumber="26">Clematis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C69E184FFF543070CB1F98EFCBD814D" box="[715,829,1544,1570]" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="genus">
<emphasis id="B91D4615FFF543070CB1F98EFCBD814D" box="[715,829,1544,1570]" italics="true" pageId="25" pageNumber="26">Caragana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFF543070D02F98EFC78814D" box="[888,1016,1544,1570]" class="Bryopsida" family="Hylocomiaceae" genus="Pleurozium" kingdom="Plantae" order="Hypnales" pageId="25" pageNumber="26" phylum="Bryophyta" rank="genus">
<emphasis id="B91D4615FFF543070D02F98EFC78814D" box="[888,1016,1544,1570]" italics="true" pageId="25" pageNumber="26">Pleurozium</emphasis>
</taxonomicName>
, southern range of Tannu-Ola,
<quantity id="4C9137E2FFF543070B1EF98EFF348129" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="25" pageNumber="26" unit="m" value="1350.0">1350 m</quantity>
.a.s.l.,
<geoCoordinate id="EE5DFCC0FFF543070E83F9AAFEDA8128" box="[249,346,1580,1607]" degrees="50" direction="north" minutes="54" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="50.9">50°54N</geoCoordinate>
<geoCoordinate id="EE5DFCC0FFF543070F1EF9AAFE448129" box="[356,452,1580,1606]" degrees="94" direction="east" minutes="29" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="94.48333">94°29E</geoCoordinate>
, Aryskanny-Khem River,
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ENE of Khol-Oozha Village,
<collectingRegion id="49AD54E5FFF543070AEBF9ABFB4A8128" box="[1169,1226,1581,1607]" country="Russia" name="Tyva" pageId="25" pageNumber="26">Tuva</collectingRegion>
Republic,
<collectingCountry id="F37EDA97FFF543070B31F9AAFA198129" box="[1355,1433,1580,1606]" name="Russia" pageId="25" pageNumber="26">Russia</collectingCountry>
,
<date id="FFD7BCC7FFF543070EEDF9D6FEAE8104" box="[151,302,1616,1643]" pageId="25" pageNumber="26" value="1995-07-18">18 July 1995</date>
, Matti Uusitalo, from sample
<collectionCode id="ED7802C2FFF543070CFEF9D7FD318104" box="[644,689,1617,1643]" country="USA" name="Midwestern University" pageId="25" pageNumber="26">MU</collectionCode>
95-39. Deposited on SEM-stub U200 at the Zoological Museum, University of Turku,
<collectingCountry id="F37EDA97FFF543070FFEF9F2FE5D81E1" box="[388,477,1652,1678]" name="Finland" pageId="25" pageNumber="26">Finland</collectingCountry>
.
</paragraph>
<paragraph id="8BD69A07FFF543070EBDF91EFF748071" blockId="25.[151,1437,1112,2039]" pageId="25" pageNumber="26">
<materialsCitation id="3B01905AFFF543070EBDF91EFF5281B9" country="United States of America" county="Cache Co." location="Juniper" municipality="Blacksmith Fork Canyon" pageId="25" pageNumber="26" specimenCount="15" stateProvince="Utah">
<specimenCount id="9D6F518EFFF543070EBDF91EFEE281DD" box="[199,354,1688,1715]" count="15" pageId="25" pageNumber="26" type="generic">15 specimens</specimenCount>
from
<collectingRegion id="49AD54E5FFF543070FD7F91EFE6381DD" box="[429,483,1688,1714]" country="United States of America" name="Utah" pageId="25" pageNumber="26">Utah</collectingRegion>
<location id="8EB6CCDCFFF543070F97F91EFDBF81DD" LSID="urn:lsid:plazi:treatment:03C02B11FFF543070EEDFC03FC408099:8EB6CCDCFFF543070F97F91EFDBF81DD" box="[493,575,1688,1714]" country="United States of America" county="Cache Co." municipality="Blacksmith Fork Canyon" name="Juniper" pageId="25" pageNumber="26" stateProvince="Utah">Juniper</location>
(
<taxonomicName id="4C69E184FFF543070C29F91FFCDA81DD" box="[595,858,1689,1714]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="species" species="osteosperma">
<emphasis id="B91D4615FFF543070C29F91FFCDA81DD" box="[595,858,1689,1714]" italics="true" pageId="25" pageNumber="26">Juniperus osteosperma</emphasis>
</taxonomicName>
) duff,
<collectingMunicipality id="6BB2007DFFF543070DD6F91EFB4F81DD" box="[940,1231,1688,1714]" pageId="25" pageNumber="26">Blacksmith Fork Canyon</collectingMunicipality>
,
<collectingCounty id="62B7E28BFFF543070AA7F91EFAD581DD" box="[1245,1365,1688,1714]" pageId="25" pageNumber="26">Cache Co.</collectingCounty>
<collectingRegion id="49AD54E5FFF543070B25F91EFA1881DD" box="[1375,1432,1688,1714]" country="United States of America" name="Utah" pageId="25" pageNumber="26">Utah</collectingRegion>
,
<collectingCountry id="F37EDA97FFF543070EEDF93AFF5281B9" box="[151,210,1724,1750]" name="United States of America" pageId="25" pageNumber="26">USA</collectingCountry>
</materialsCitation>
,
<date id="FFD7BCC7FFF543070EA4F93AFEFF81B9" box="[222,383,1724,1750]" pageId="25" pageNumber="26" value="1970-04-24">24 April 1970</date>
, G.F. Knowlton &amp; W.J. Hanson; of them,
<materialsCitation id="3B01905AFFF543070D21F93AFD698195" collectorName="University" country="United States of America" location="Acarology Laboratory" pageId="25" pageNumber="26" specimenCount="14" specimenCount-female="10" specimenCount-male="4" stateProvince="Ohio">
<specimenCount id="9D6F518EFFF543070D21F93AFC5681B9" box="[859,982,1724,1750]" count="10" pageId="25" pageNumber="26" type="female">10 females</specimenCount>
and
<specimenCount id="9D6F518EFFF543070A75F93AFBE681B9" box="[1039,1126,1724,1750]" count="4" pageId="25" pageNumber="26" type="male">4 males</specimenCount>
deposited at the
<location id="8EB6CCDCFFF543070B5CF93AFE958195" LSID="urn:lsid:plazi:treatment:03C02B11FFF543070EEDFC03FC408099:8EB6CCDCFFF543070B5CF93AFE958195" country="United States of America" name="Acarology Laboratory" pageId="25" pageNumber="26" stateProvince="Ohio">Acarology Laboratory</location>
,
<collectingRegion id="49AD54E5FFF543070F5BF966FE198195" box="[289,409,1760,1786]" country="United States of America" name="Ohio" pageId="25" pageNumber="26">Ohio State</collectingRegion>
<collectorName id="269CFFD1FFF543070FDBF966FD988195" box="[417,536,1760,1786]" pageId="25" pageNumber="26">University</collectorName>
, coll. no. AL0726
</materialsCitation>
;
<materialsCitation id="3B01905AFFF543070C8FF967FF708071" country="Finland" location="Zoological Museum" municipality="University of Turku" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D6F518EFFF543070C8FF967FCF28195" box="[757,882,1760,1787]" count="1" pageId="25" pageNumber="26" type="female">one female</specimenCount>
at the
<location id="8EB6CCDCFFF543070DBBF966FB288194" LSID="urn:lsid:plazi:treatment:03C02B11FFF543070EEDFC03FC408099:8EB6CCDCFFF543070DBBF966FB288194" box="[961,1192,1760,1787]" country="Finland" municipality="University of Turku" name="Zoological Museum" pageId="25" pageNumber="26">Zoological Museum</location>
,
<collectingMunicipality id="6BB2007DFFF543070ACEF966FA188195" box="[1204,1432,1760,1786]" pageId="25" pageNumber="26">University of Turku</collectingMunicipality>
,
<collectingCountry id="F37EDA97FFF543070EEDF882FF708071" box="[151,240,1796,1822]" name="Finland" pageId="25" pageNumber="26">Finland</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF543070EBDF8AEFCC080E4" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="8BD69A07FFF543070EBDF8AEFCC080E4" blockId="25.[151,1437,1112,2039]" pageId="25" pageNumber="26">
<emphasis id="B91D4615FFF543070EBDF8AEFE51802D" bold="true" box="[199,465,1832,1858]" pageId="25" pageNumber="26">Differential diagnosis.</emphasis>
This species can be segregated from other
<taxonomicName id="4C69E184FFF543070DABF8AEFB3D802D" box="[977,1213,1832,1858]" pageId="25" pageNumber="26">
<emphasis id="B91D4615FFF543070DABF8AEFBE2802D" box="[977,1122,1832,1858]" italics="true" pageId="25" pageNumber="26">Amphialycus</emphasis>
species
</taxonomicName>
by having a broad naso with longitudinally arranged striae and tapering end, three pairs of adoral setae, two pairs of cheliceral setae (posterior being forked), and a great number of genital setae.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFF543070EBDF812FC408099" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BD69A07FFF543070EBDF812FC408099" blockId="25.[151,1437,1112,2039]" pageId="25" pageNumber="26">
<emphasis id="B91D4615FFF543070EBDF812FEBB80C1" bold="true" box="[199,315,1940,1966]" pageId="25" pageNumber="26">Remarks.</emphasis>
<taxonomicName id="4C69E184FFF543070F39F812FE6080C1" box="[323,480,1940,1966]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="holarcticus">
<emphasis id="B91D4615FFF543070F39F812FE6080C1" box="[323,480,1940,1966]" italics="true" pageId="25" pageNumber="26">A. holarcticus</emphasis>
</taxonomicName>
was coded as amp in
<bibRefCitation id="EFF8E7F6FFF543070C90F812FC1580C1" author="Uusitalo, M." box="[746,917,1940,1966]" pageId="25" pageNumber="26" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo (2010</bibRefCitation>
, fig. 1). The anterior pair of eyes is extremely finely striated (see Uusitalo 2020: SEM fig. 1, by high magnification) which is typical to the genus
<taxonomicName id="4C69E184FFF543070B7FF83EFA1680BD" authorityName="Zachvatkin" authorityYear="1949" box="[1285,1430,1976,2002]" class="Arachnida" family="Alycidae" genus="Amphialycus" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFF543070B7FF83EFA1680BD" box="[1285,1430,1976,2002]" italics="true" pageId="25" pageNumber="26">Amphialycus</emphasis>
</taxonomicName>
, but overlooked by
<bibRefCitation id="EFF8E7F6FFF543070F10F85AFD948099" author="Uusitalo, M." box="[362,532,2012,2038]" pageId="25" pageNumber="26" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo (2010</bibRefCitation>
, 2020, mentioned as glabrous in text).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,304 @@
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<paragraph id="8BD69A07FFFE430C0EEDFAB3FD888220" blockId="18.[151,520,1333,1396]" box="[151,520,1333,1360]" pageId="18" pageNumber="19">
<heading id="D09E2D6BFFFE430C0EEDFAB3FD888220" bold="true" box="[151,520,1333,1360]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="4">
<emphasis id="B91D4615FFFE430C0EEDFAB3FD888220" bold="true" box="[151,520,1333,1360]" pageId="18" pageNumber="19">
<taxonomicName id="4C69E184FFFE430C0EEDFAB3FE24823F" authorityName="Uusitalo" authorityYear="2025" box="[151,420,1333,1360]" class="Arachnida" family="Alycidae" genus="Odontoalycus" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="exoculo" status="sp. nov.">
<emphasis id="B91D4615FFFE430C0EEDFAB3FE24823F" bold="true" box="[151,420,1333,1360]" italics="true" pageId="18" pageNumber="19">Odontoalycus exoculo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22EFB6EFFFE430C0FD1FAB3FD888220" box="[427,520,1333,1359]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFFE430C0EEDFADFFABC8004" pageId="18" pageNumber="19" type="description">
<paragraph id="8BD69A07FFFE430C0EEDFADFFE46821C" blockId="18.[151,520,1333,1396]" box="[151,454,1369,1396]" pageId="18" pageNumber="19">
(
<figureCitation id="13528682FFFE430C0EE5FADFFEA6821B" box="[159,294,1369,1396]" captionStart="FIGURE 7" captionStartId="19.[151,250,1848,1872]" captionTargetBox="[151,1436,176,1824]" captionTargetId="figure-15@19.[151,1436,176,1824]" captionTargetPageId="19" captionText="FIGURE 7. Odontoalycus exoculo sp. nov., tritonymph. A—prodorsum holotrichous, eyes absent; B—dorsum holotrichous; C—rutellum and adoral setae; D—genital and anal area, 1314 genital setae per valve, 2 anal setae per valve, ventral area slighty neotrichous. All specimens—from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">Figs. 7AD</figureCitation>
,
<figureCitation id="13528682FFFE430C0F4BFADFFEF1821C" box="[305,369,1369,1395]" captionStart="FIGURE 8" captionStartId="20.[151,250,1899,1923]" captionTargetBox="[151,1436,176,1876]" captionTargetId="figure-24@20.[151,1436,176,1876]" captionTargetPageId="20" captionText="FIGURE 8. Odontoalycus exoculo sp. nov., tritonymph. A—leg I; B—leg II; C—tibia and genu III; D—tibia and genu IV; E—palp and dorsal view of chelicera; F—Movable digit of chelicera with peg-like teeth, fixed digit with a massive tooth. All specimens—from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">8AF</figureCitation>
,
<figureCitation id="13528682FFFE430C0F07FADFFE3E821C" box="[381,446,1369,1395]" captionStart="FIGURE 9" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@21.[151,1435,176,1849]" captionTargetPageId="21" captionText="FIGURE 9. Odontoalycus exoculo sp. nov., tritonymph. A—Prodorsum, soft integument by rows of transversely inserted lamellae (white arrows); B—Original dorsal view; C—Digits of left chelicera; D—Digits of right chelicera; E—Pair of claws and a clawless empodium; F—Palpal tarsus. All specimens— from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">9AF</figureCitation>
)
</paragraph>
<paragraph id="8BD69A07FFFE430C0EEDFA27FE308124" blockId="18.[151,1437,1441,2008]" pageId="18" pageNumber="19">
<emphasis id="B91D4615FFFE430C0EEDFA27FEA982D4" bold="true" box="[151,297,1441,1467]" pageId="18" pageNumber="19">Description.</emphasis>
<emphasis id="B91D4615FFFE430C0F4AFA24FE0A82D4" box="[304,394,1442,1467]" italics="true" pageId="18" pageNumber="19">Dorsum</emphasis>
(n=1 tritonymph,
<figureCitation id="13528682FFFE430C0C23FA27FD5D82D3" box="[601,733,1441,1468]" captionStart="FIGURE 7" captionStartId="19.[151,250,1848,1872]" captionTargetBox="[151,1436,176,1824]" captionTargetId="figure-15@19.[151,1436,176,1824]" captionTargetPageId="19" captionText="FIGURE 7. Odontoalycus exoculo sp. nov., tritonymph. A—prodorsum holotrichous, eyes absent; B—dorsum holotrichous; C—rutellum and adoral setae; D—genital and anal area, 1314 genital setae per valve, 2 anal setae per valve, ventral area slighty neotrichous. All specimens—from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">Figs. 7A, B</figureCitation>
,
<figureCitation id="13528682FFFE430C0C93FA27FCAF82D4" box="[745,815,1441,1467]" captionStart="FIGURE 9" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@21.[151,1435,176,1849]" captionTargetPageId="21" captionText="FIGURE 9. Odontoalycus exoculo sp. nov., tritonymph. A—Prodorsum, soft integument by rows of transversely inserted lamellae (white arrows); B—Original dorsal view; C—Digits of left chelicera; D—Digits of right chelicera; E—Pair of claws and a clawless empodium; F—Palpal tarsus. All specimens— from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">9A, B</figureCitation>
). Length 240 µm; integumental ridges densely covered by transverse lamellae (see
<emphasis id="B91D4615FFFE430C0FB2FA43FDAA82B0" bold="true" box="[456,554,1477,1503]" pageId="18" pageNumber="19">Remark</emphasis>
); holotrichous set of dorsal setae; ciliated setae with long shafts on microplates; setae slightly longer in caudal segments; prodorsum with filamentous sensilla
<emphasis id="B91D4615FFFE430C0D85FA6CFB98816C" box="[1023,1048,1514,1539]" italics="true" pageId="18" pageNumber="19">ve</emphasis>
,
<emphasis id="B91D4615FFFE430C0A5FFA6CFBC4816C" box="[1061,1092,1514,1539]" italics="true" pageId="18" pageNumber="19">sci</emphasis>
and slightly elongated setae
<emphasis id="B91D4615FFFE430C0BFCFA6CFA1C816C" box="[1414,1436,1514,1539]" italics="true" pageId="18" pageNumber="19">in</emphasis>
on prodorsal shield; ciliated setae
<emphasis id="B91D4615FFFE430C0C60F988FDAE8148" box="[538,558,1550,1575]" italics="true" pageId="18" pageNumber="19">vi</emphasis>
,
<emphasis id="B91D4615FFFE430C0C47F988FDE18148" box="[573,609,1550,1575]" italics="true" pageId="18" pageNumber="19">sce</emphasis>
and
<emphasis id="B91D4615FFFE430C0CE3F988FD408148" box="[665,704,1550,1575]" italics="true" pageId="18" pageNumber="19">exp</emphasis>
on separate microplates; mutual distances of
<emphasis id="B91D4615FFFE430C0ABCF988FB5A8148" box="[1222,1242,1550,1575]" italics="true" pageId="18" pageNumber="19">vi</emphasis>
,
<emphasis id="B91D4615FFFE430C0A93F988FA828148" box="[1257,1282,1550,1575]" italics="true" pageId="18" pageNumber="19">ve</emphasis>
and
<emphasis id="B91D4615FFFE430C0B40F988FAD08148" box="[1338,1360,1550,1575]" italics="true" pageId="18" pageNumber="19">in</emphasis>
equal; eyes absent; naso absent.
</paragraph>
<paragraph id="8BD69A07FFFE430C0EBDF9D0FAE08100" blockId="18.[151,1437,1441,2008]" box="[199,1376,1621,1648]" pageId="18" pageNumber="19">
<taxonomicName id="4C69E184FFFE430C0EBDF9D0FE8F8100" box="[199,271,1622,1647]" class="Gastropoda" family="Rissoidae" genus="Venter" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B91D4615FFFE430C0EBDF9D0FE8F8100" box="[199,271,1622,1647]" italics="true" pageId="18" pageNumber="19">Venter</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFFE430C0F64F9D3FEF88100" box="[286,376,1621,1647]" captionStart="FIGURE 7" captionStartId="19.[151,250,1848,1872]" captionTargetBox="[151,1436,176,1824]" captionTargetId="figure-15@19.[151,1436,176,1824]" captionTargetPageId="19" captionText="FIGURE 7. Odontoalycus exoculo sp. nov., tritonymph. A—prodorsum holotrichous, eyes absent; B—dorsum holotrichous; C—rutellum and adoral setae; D—genital and anal area, 1314 genital setae per valve, 2 anal setae per valve, ventral area slighty neotrichous. All specimens—from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">Fig. 7D</figureCitation>
). Setae short with less than 10 cilia; 1314 genital setae per valve; 2 anal setae per valve.
</paragraph>
<paragraph id="8BD69A07FFFE430C0EBDF9FFFE948190" blockId="18.[151,1437,1441,2008]" pageId="18" pageNumber="19">
<taxonomicName id="4C69E184FFFE430C0EBDF9FFFED681FC" authorityName="Krefft" authorityYear="1871" box="[199,342,1657,1683]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhabditida" pageId="18" pageNumber="19" phylum="Nematoda" rank="genus">
<emphasis id="B91D4615FFFE430C0EBDF9FFFED681FC" box="[199,342,1657,1683]" italics="true" pageId="18" pageNumber="19">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="13528682FFFE430C0F1DF9FFFE4A81FC" box="[359,458,1657,1683]" captionStart="FIGURE 7" captionStartId="19.[151,250,1848,1872]" captionTargetBox="[151,1436,176,1824]" captionTargetId="figure-15@19.[151,1436,176,1824]" captionTargetPageId="19" captionText="FIGURE 7. Odontoalycus exoculo sp. nov., tritonymph. A—prodorsum holotrichous, eyes absent; B—dorsum holotrichous; C—rutellum and adoral setae; D—genital and anal area, 1314 genital setae per valve, 2 anal setae per valve, ventral area slighty neotrichous. All specimens—from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">Figs. 7C</figureCitation>
,
<figureCitation id="13528682FFFE430C0FADF9FFFD9681FB" box="[471,534,1657,1684]" captionStart="FIGURE 8" captionStartId="20.[151,250,1899,1923]" captionTargetBox="[151,1436,176,1876]" captionTargetId="figure-24@20.[151,1436,176,1876]" captionTargetPageId="20" captionText="FIGURE 8. Odontoalycus exoculo sp. nov., tritonymph. A—leg I; B—leg II; C—tibia and genu III; D—tibia and genu IV; E—palp and dorsal view of chelicera; F—Movable digit of chelicera with peg-like teeth, fixed digit with a massive tooth. All specimens—from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">8E, F</figureCitation>
,
<figureCitation id="13528682FFFE430C0C58F9FFFD0981FC" box="[546,649,1657,1684]" captionStart="FIGURE 9" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@21.[151,1435,176,1849]" captionTargetPageId="21" captionText="FIGURE 9. Odontoalycus exoculo sp. nov., tritonymph. A—Prodorsum, soft integument by rows of transversely inserted lamellae (white arrows); B—Original dorsal view; C—Digits of left chelicera; D—Digits of right chelicera; E—Pair of claws and a clawless empodium; F—Palpal tarsus. All specimens— from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">9C, D, F</figureCitation>
). Chelicera robust, non-elongated, with one pair of smooth cheliceral setae in dorsal position; movable chela with 10 pairs of teeth, subequal in size; fixed digit with a pair of large teeth; subcapitulum with 3 pairs of adoral setae; rutella toothpick-like; palpal tarsus with three pairs of pseudacanthoid eupathidia.
</paragraph>
<paragraph id="8BD69A07FFFE430C0EBDF88CFABC8004" blockId="18.[151,1437,1441,2008]" pageId="18" pageNumber="19">
<emphasis id="B91D4615FFFE430C0EBDF88CFF7C804C" box="[199,252,1802,1827]" italics="true" pageId="18" pageNumber="19">Legs</emphasis>
(
<figureCitation id="13528682FFFE430C0F74F88FFE18804C" box="[270,408,1801,1827]" captionStart="FIGURE 8" captionStartId="20.[151,250,1899,1923]" captionTargetBox="[151,1436,176,1876]" captionTargetId="figure-24@20.[151,1436,176,1876]" captionTargetPageId="20" captionText="FIGURE 8. Odontoalycus exoculo sp. nov., tritonymph. A—leg I; B—leg II; C—tibia and genu III; D—tibia and genu IV; E—palp and dorsal view of chelicera; F—Movable digit of chelicera with peg-like teeth, fixed digit with a massive tooth. All specimens—from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">Figs. 8AD</figureCitation>
,
<figureCitation id="13528682FFFE430C0FDCF88FFE7B804C" box="[422,507,1801,1827]" captionStart="FIGURE 9" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@21.[151,1435,176,1849]" captionTargetPageId="21" captionText="FIGURE 9. Odontoalycus exoculo sp. nov., tritonymph. A—Prodorsum, soft integument by rows of transversely inserted lamellae (white arrows); B—Original dorsal view; C—Digits of left chelicera; D—Digits of right chelicera; E—Pair of claws and a clawless empodium; F—Palpal tarsus. All specimens— from Quintana Roo, Sian Kaan." pageId="18" pageNumber="19">9B, 9E</figureCitation>
). Solenidial formula for tarsi, tibiae, genua and femora of legs I, II, III, and IV, respectively: 2-2-0-0, 2-2-2-0, 6-2-2-1, 6-1-0-0, both solenidia on tibia I are more like baculiform, some solenidia are awl-like, short and sharp, pestle-like famulus I and famulus II abaxially near claws; empodia clawless.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFFE430C0EBDF8F3FAC280B8" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BD69A07FFFE430C0EBDF8F3FAC280B8" blockId="18.[151,1437,1441,2008]" pageId="18" pageNumber="19">
<materialsCitation id="3B01905AFFFE430C0EBDF8F3FA9280FF" box="[199,1298,1909,1936]" location="Sian Ka'an" municipality="Several" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Quintana Roo" typeStatus="holotype">
<emphasis id="B91D4615FFFE430C0EBDF8F3FEF580E0" bold="true" box="[199,373,1909,1935]" pageId="18" pageNumber="19">
<typeStatus id="54D224A5FFFE430C0EBDF8F3FE8280E0" box="[199,258,1909,1935]" pageId="18" pageNumber="19">Type</typeStatus>
material.
</emphasis>
<collectingMunicipality id="6BB2007DFFFE430C0F01F8F3FE4F80E0" box="[379,463,1909,1935]" pageId="18" pageNumber="19">Several</collectingMunicipality>
high-quality photos of 1 tritonymph from litter,
<location id="8EB6CCDCFFFE430C0D90F8F3FBE980E0" LSID="urn:lsid:plazi:treatment:03C02B11FFFE43080EEDFAB3FC628438:8EB6CCDCFFFE430C0D90F8F3FBE980E0" box="[1002,1129,1909,1935]" municipality="Several" name="Sian Ka'an" pageId="18" pageNumber="19" stateProvince="Quintana Roo">Sian Kaan</location>
,
<collectingRegion id="49AD54E5FFFE430C0A0EF8F3FA9280FF" box="[1140,1298,1909,1936]" country="Mexico" name="Quintana Roo" pageId="18" pageNumber="19">Quintana Roo</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B01905AFFFE430C0B67F8F3FABE80B8" collectingDate="2014-11-20" collectionCode="CNCA, UNAM" collectorName="Maria Magdalena Vazquez. The &amp; de Biologia &amp; Universidad Nacional Autonoma" country="Mexico" location="Instituto de Biologia" municipality="Coleccion Nacional de Acaros" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Distrito Federal" typeStatus="holotype">
<collectingCountry id="F37EDA97FFFE430C0B67F8F3FAF580E0" box="[1309,1397,1909,1935]" name="Mexico" pageId="18" pageNumber="19">Mexico</collectingCountry>
,
<date id="FFD7BCC7FFFE430C0BFAF8F3FED080DC" pageId="18" pageNumber="19" value="2014-11-20">
<collectingDate id="EF93452FFFFE430C0BFAF8F3FED080DC" pageId="18" pageNumber="19" value="2014-11-20">20 November 2014</collectingDate>
</date>
,
<collectorName id="269CFFD1FFFE430C0F21F81FFD4580DC" box="[347,709,1945,1971]" pageId="18" pageNumber="19">María Magdalena Vázquez. The</collectorName>
material will be deposited in the
<collectionCode id="ED7802C2FFFE430C0A45F81FFB0B80DC" box="[1087,1163,1945,1971]" pageId="18" pageNumber="19">CNCA</collectionCode>
(
<collectingMunicipality id="6BB2007DFFFE430C0AE0F81FFF6880B8" pageId="18" pageNumber="19">Colección Nacional de Acaros</collectingMunicipality>
) at the
<location id="8EB6CCDCFFFE430C0F41F83BFDA180B8" LSID="urn:lsid:plazi:treatment:03C02B11FFFE43080EEDFAB3FC628438:8EB6CCDCFFFE430C0F41F83BFDA180B8" box="[315,545,1981,2007]" country="Mexico" municipality="Coleccion Nacional de Acaros" name="Instituto de Biologia" pageId="18" pageNumber="19" stateProvince="Distrito Federal">
Instituto
<collectorName id="269CFFD1FFFE430C0FDAF83BFDA180B8" box="[416,545,1981,2007]" pageId="18" pageNumber="19">de Biología</collectorName>
</location>
at
<collectionCode id="ED7802C2FFFE430C0C3EF838FD1A80B7" box="[580,666,1982,2008]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="18" pageNumber="19">UNAM</collectionCode>
(
<collectorName id="269CFFD1FFFE430C0CD2F83BFB9A80B8" box="[680,1050,1981,2007]" pageId="18" pageNumber="19">Universidad Nacional Autonóma</collectorName>
de
<collectingCountry id="F37EDA97FFFE430C0A39F83BFB1D80B8" box="[1091,1181,1981,2007]" name="Mexico" pageId="18" pageNumber="19">México</collectingCountry>
),
<collectingRegion id="49AD54E5FFFE430C0AD5F83BFABE80B8" box="[1199,1342,1981,2007]" country="Mexico" name="Distrito Federal" pageId="18" pageNumber="19">Mexico City</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF16CA8FFFFF430D0EEDF8BEFF5C80D3" pageId="19" pageNumber="20" startId="19.[151,250,1848,1872]" targetBox="[151,1436,176,1824]" targetPageId="19" targetType="figure">
<paragraph id="8BD69A07FFFF430D0EEDF8BEFD57803F" blockId="19.[151,1436,1848,1980]" box="[151,727,1848,1872]" pageId="19" pageNumber="20">
<emphasis id="B91D4615FFFF430D0EEDF8BEFE93803F" bold="true" box="[151,275,1848,1872]" pageId="19" pageNumber="20">FIGURE 7.</emphasis>
<taxonomicName id="4C69E184FFFF430D0F63F8BEFE7B803F" authorityName="Uusitalo" authorityYear="2025" box="[281,507,1848,1872]" class="Arachnida" family="Alycidae" genus="Odontoalycus" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="exoculo" status="sp. nov.">
<emphasis id="B91D4615FFFF430D0F63F8BEFE7B803F" box="[281,507,1848,1872]" italics="true" pageId="19" pageNumber="20">Odontoalycus exoculo</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFFF430D0C78F8BEFDD9803F" bold="true" box="[514,601,1848,1872]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A22EFB6EFFFF430D0C78F8BEFDD4803F" box="[514,596,1848,1872]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
tritonymph.
</paragraph>
<paragraph id="8BD69A07FFFF430D0EEDF8DAFF5C80D3" blockId="19.[151,1436,1848,1980]" pageId="19" pageNumber="20">A—prodorsum holotrichous, eyes absent; B—dorsum holotrichous; C—rutellum and adoral setae; D—genital and anal area, 1314 genital setae per valve, 2 anal setae per valve, ventral area slighty neotrichous. All specimens—from Quintana Roo, Sian Kaan.</paragraph>
</caption>
<caption id="DF16CA8FFFF8430A0EEDF8EDFA8380A3" pageId="20" pageNumber="21" startId="20.[151,250,1899,1923]" targetBox="[151,1436,176,1876]" targetPageId="20" targetType="figure">
<paragraph id="8BD69A07FFF8430A0EEDF8EDFD5780EC" blockId="20.[151,1436,1899,1996]" box="[151,727,1899,1924]" pageId="20" pageNumber="21">
<emphasis id="B91D4615FFF8430A0EEDF8EDFE9380EC" bold="true" box="[151,275,1899,1923]" pageId="20" pageNumber="21">FIGURE 8.</emphasis>
<taxonomicName id="4C69E184FFF8430A0F63F8EDFE7B80EC" authorityName="Uusitalo" authorityYear="2025" box="[281,507,1899,1923]" class="Arachnida" family="Alycidae" genus="Odontoalycus" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="exoculo" status="sp. nov.">
<emphasis id="B91D4615FFF8430A0F63F8EDFE7B80EC" box="[281,507,1899,1923]" italics="true" pageId="20" pageNumber="21">Odontoalycus exoculo</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFF8430A0C78F8EAFDD980EB" bold="true" box="[514,601,1900,1924]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A22EFB6EFFF8430A0C78F8EAFDD480EB" box="[514,596,1900,1924]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
tritonymph.
</paragraph>
<paragraph id="8BD69A07FFF8430A0EEDF809FA8380A3" blockId="20.[151,1436,1899,1996]" pageId="20" pageNumber="21">A—leg I; B—leg II; C—tibia and genu III; D—tibia and genu IV; E—palp and dorsal view of chelicera; F—Movable digit of chelicera with peg-like teeth, fixed digit with a massive tooth. All specimens—from Quintana Roo, Sian Kaan.</paragraph>
</caption>
<caption id="DF16CA8FFFF9430B0EEDF8D7FE6180BA" pageId="21" pageNumber="22" startId="21.[151,250,1873,1897]" targetBox="[151,1435,176,1849]" targetPageId="21" targetType="figure">
<paragraph id="8BD69A07FFF9430B0EEDF8D7FD578006" blockId="21.[151,1436,1873,2005]" box="[151,727,1873,1897]" pageId="21" pageNumber="22">
<emphasis id="B91D4615FFF9430B0EEDF8D7FE938006" bold="true" box="[151,275,1873,1897]" pageId="21" pageNumber="22">FIGURE 9.</emphasis>
<taxonomicName id="4C69E184FFF9430B0F63F8D7FE7B8006" authorityName="Uusitalo" authorityYear="2025" box="[281,507,1873,1897]" class="Arachnida" family="Alycidae" genus="Odontoalycus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="exoculo" status="sp. nov.">
<emphasis id="B91D4615FFF9430B0F63F8D7FE7B8006" box="[281,507,1873,1897]" italics="true" pageId="21" pageNumber="22">Odontoalycus exoculo</emphasis>
</taxonomicName>
<emphasis id="B91D4615FFF9430B0C78F8D7FDD98006" bold="true" box="[514,601,1873,1897]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="A22EFB6EFFF9430B0C78F8D7FDD48006" box="[514,596,1873,1897]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
tritonymph.
</paragraph>
<paragraph id="8BD69A07FFF9430B0EEDF8F3FE6180BA" blockId="21.[151,1436,1873,2005]" pageId="21" pageNumber="22">A—Prodorsum, soft integument by rows of transversely inserted lamellae (white arrows); B—Original dorsal view; C—Digits of left chelicera; D—Digits of right chelicera; E—Pair of claws and a clawless empodium; F—Palpal tarsus. All specimens— from Quintana Roo, Sian Kaan.</paragraph>
</caption>
<subSubSection id="C373C98CFFFA43080EBDFF10FE068673" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BD69A07FFFA43080EBDFF10FE068673" blockId="22.[151,1436,150,393]" pageId="22" pageNumber="23">
<emphasis id="B91D4615FFFA43080EBDFF10FE4E87DF" bold="true" box="[199,462,150,176]" pageId="22" pageNumber="23">Differential diagnosis.</emphasis>
The species can be segregated from the other species of the tribe by having: an eyeless prodorsum; chelicera with ca. 20 tiny, beak-like teeth on the movable digit and one pair of massive teeth on the fixed digit; and a clawless empodia.
<taxonomicName id="4C69E184FFFA43080C44FF58FD098797" authorityName="C. L. Koch" authorityYear="1842" box="[574,649,222,248]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFFA43080C44FF58FD098797" box="[574,649,222,248]" italics="true" pageId="22" pageNumber="23">Alycus</emphasis>
</taxonomicName>
share the robust and non-elongated chelicera, but differ by having a holotrichous dorsum.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFFA43080EBDFEA0FEE4860B" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BD69A07FFFA43080EBDFEA0FEE4860B" blockId="22.[151,1436,150,393]" pageId="22" pageNumber="23">
<emphasis id="B91D4615FFFA43080EBDFEA0FEB0862F" bold="true" box="[199,304,294,320]" pageId="22" pageNumber="23">Remark.</emphasis>
The specimen is flattened, but transverse lamellae are visible in random places, indicated by white arrows in
<figureCitation id="13528682FFFA43080F7DFECCFEE0860B" box="[263,352,330,356]" captionStart="FIGURE 9" captionStartId="21.[151,250,1873,1897]" captionTargetBox="[151,1435,176,1849]" captionTargetId="figure-15@21.[151,1435,176,1849]" captionTargetPageId="21" captionText="FIGURE 9. Odontoalycus exoculo sp. nov., tritonymph. A—Prodorsum, soft integument by rows of transversely inserted lamellae (white arrows); B—Original dorsal view; C—Digits of left chelicera; D—Digits of right chelicera; E—Pair of claws and a clawless empodium; F—Palpal tarsus. All specimens— from Quintana Roo, Sian Kaan." pageId="22" pageNumber="23">Fig. 9A</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFFA43080EBDFEE8FA138558" pageId="22" pageNumber="23" type="etymology">
<paragraph id="8BD69A07FFFA43080EBDFEE8FB9086E6" blockId="22.[151,1436,150,393]" box="[199,1040,366,393]" pageId="22" pageNumber="23">
<emphasis id="B91D4615FFFA43080EBDFEE8FECE86E7" bold="true" box="[199,334,366,392]" pageId="22" pageNumber="23">Etymology.</emphasis>
The specific name (Lat.
<taxonomicName id="4C69E184FFFA43080C1FFEE8FD3C86E7" authorityName="Uusitalo" authorityYear="2025" box="[613,700,366,392]" class="Arachnida" family="Alycidae" genus="Odontoalycus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="exoculo">
<emphasis id="B91D4615FFFA43080C1FFEE8FD3C86E7" box="[613,700,366,392]" italics="true" pageId="22" pageNumber="23">exoculo</emphasis>
</taxonomicName>
) refers to the absence of eyes.
</paragraph>
<paragraph id="8BD69A07FFFA43080EEDFE5CFDB9869B" blockId="22.[151,569,473,501]" box="[151,569,473,501]" pageId="22" pageNumber="23">
<emphasis id="B91D4615FFFA43080EEDFE5CFDB9869B" bold="true" box="[151,569,473,501]" pageId="22" pageNumber="23">
Genus
<taxonomicName id="4C69E184FFFA43080E94FE5CFDB9869B" authority="Duges, 1834" authorityName="Duges" authorityYear="1834" box="[238,569,473,501]" class="Arachnida" family="Alycidae" genus="Pachygnathus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFFA43080E94FE5CFE1B869A" bold="true" box="[238,411,474,501]" italics="true" pageId="22" pageNumber="23">Pachygnathus</emphasis>
<bibRefCitation id="EFF8E7F6FFFA43080FD9FE5FFDB9869B" author="Duges, A." box="[419,569,473,500]" pageId="22" pageNumber="23" pagination="18 - 63" refId="ref46907" refString="Duges, A. 1834, Recherces sur l'ordre des Acariens, Troisieme Memoire, Annales des Sciences Naturelles, 2 serie, Tome 2., Zoologie, 18-63, pls. 7 - 8." type="book chapter" year="1834">Dugès, 1834</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BD69A07FFFA43080EEDFD99FA138558" blockId="22.[151,1427,543,567]" box="[151,1427,543,567]" pageId="22" pageNumber="23">
<materialsCitation id="3B01905AFFFA43080EEDFD99FA0E8558" box="[151,1422,543,567]" country="France" location="Oken" pageId="22" pageNumber="23" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D224A5FFFA43080EEDFD99FF498558" box="[151,201,543,567]" pageId="22" pageNumber="23">Type</typeStatus>
species: “
<emphasis id="B91D4615FFFA43080F48FD99FE668558" box="[306,486,543,567]" italics="true" pageId="22" pageNumber="23">Pachygnathe velu</emphasis>
<bibRefCitation id="EFF8E7F6FFFA43080F82FD99FDF88558" author="Duges, A." box="[504,632,543,567]" pageId="22" pageNumber="23" pagination="18 - 63" refId="ref46907" refString="Duges, A. 1834, Recherces sur l'ordre des Acariens, Troisieme Memoire, Annales des Sciences Naturelles, 2 serie, Tome 2., Zoologie, 18-63, pls. 7 - 8." type="book chapter" year="1834">Dugès, 1834</bibRefCitation>
(=
<taxonomicName id="4C69E184FFFA43080CEFFD99FC408558" authority="Duges" authorityName="Duges" box="[661,960,543,567]" class="Arachnida" family="Alycidae" genus="Pachygnathus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="B91D4615FFFA43080CEFFD99FCF88558" box="[661,888,543,567]" italics="true" pageId="22" pageNumber="23">Pachygnathus villosus</emphasis>
Dugès
</taxonomicName>
[in:
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], 1836 from
<collectingCountry id="F37EDA97FFFA43080AD6FD99FB768558" box="[1196,1270,543,567]" name="France" pageId="22" pageNumber="23">France</collectingCountry>
); by monotypy
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.
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFFA43080EEDFDE3FD8F85A8" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BD69A07FFFA43080EEDFDE3FD8F85A8" blockId="22.[151,1437,613,855]" pageId="22" pageNumber="23">
<emphasis id="B91D4615FFFA43080EEDFDE3FE1D8510" bold="true" box="[151,413,613,639]" pageId="22" pageNumber="23">Differential diagnosis.</emphasis>
The genus can be segregated from the other genera of the family by the beak-like chelicerae (
<figureCitation id="13528682FFFA43080EE5FD0FFE8885CC" box="[159,264,649,675]" captionStart="FIGURE 10" captionStartId="23.[151,250,1861,1885]" captionTargetBox="[164,1423,153,1837]" captionTargetId="figure-15@23.[164,1423,153,1837]" captionTargetPageId="23" captionText="FIGURE 10. Pachygnathus americanus sp. nov. A—left half of prodorsum; B—reconstruction of neotrichous dorsum with some uncertainty in setal amount and positions; C— cheliceral chelae with teeth; D—lateral lip with 2 adoral setae and rutellum; E—genital valve of female with 17 setae, one pair of eugenital setae in genital chamber; F—anal valve with 4 setae; G—holotrichous dorsum of Pachygnathus nasutus Uusitalo et al. AG—from Quintana Roo, Puerto Morelos; 10G—from South Africa, North-West Province." pageId="22" pageNumber="23">Fig. 10C</figureCitation>
), small, fork-like rutella (
<figureCitation id="13528682FFFA43080C4EFD0FFD1F85CC" box="[564,671,649,675]" captionStart="FIGURE 10" captionStartId="23.[151,250,1861,1885]" captionTargetBox="[164,1423,153,1837]" captionTargetId="figure-15@23.[164,1423,153,1837]" captionTargetPageId="23" captionText="FIGURE 10. Pachygnathus americanus sp. nov. A—left half of prodorsum; B—reconstruction of neotrichous dorsum with some uncertainty in setal amount and positions; C— cheliceral chelae with teeth; D—lateral lip with 2 adoral setae and rutellum; E—genital valve of female with 17 setae, one pair of eugenital setae in genital chamber; F—anal valve with 4 setae; G—holotrichous dorsum of Pachygnathus nasutus Uusitalo et al. AG—from Quintana Roo, Puerto Morelos; 10G—from South Africa, North-West Province." pageId="22" pageNumber="23">Fig. 10D</figureCitation>
,
<bibRefCitation id="EFF8E7F6FFFA43080CD7FD0FFCD385CC" author="Uusitalo, M." box="[685,851,649,675]" pageId="22" pageNumber="23" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
: fig. 58), and having less than six sparsely barbed eupathidia on the palpi (
<figureCitation id="13528682FFFA43080FDCFD2BFD8285A8" box="[422,514,685,711]" captionStart="FIGURE 11" captionStartId="24.[151,250,1800,1824]" captionTargetBox="[151,1435,153,1776]" captionTargetId="figure-24@24.[151,1436,153,1776]" captionTargetPageId="24" captionText="FIGURE 11. Pachygnathus americanus sp. nov. A—tarsus I; B—tibia I; C—genu and femur I; D—tarsus II; E—tibia, genu and femur II; F—tarsus III; G—tibia and genu III; H—leg IV; I—palp. All specimens—from Quintana Roo, Puerto Morelos." pageId="22" pageNumber="23">Fig. 11I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C373C98CFFFA43080EBDFD57FC628438" pageId="22" pageNumber="23" type="discussion">
<paragraph id="8BD69A07FFFA43080EBDFD57FC628438" blockId="22.[151,1437,613,855]" pageId="22" pageNumber="23">
<emphasis id="B91D4615FFFA43080EBDFD57FEBB8584" bold="true" box="[199,315,721,747]" pageId="22" pageNumber="23">Remarks.</emphasis>
The genera
<taxonomicName id="4C69E184FFFA43080FB9FD57FD8E8584" authorityName="C. L. Koch" authorityYear="1842" box="[451,526,721,747]" class="Arachnida" family="Alycidae" genus="Alycus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFFA43080FB9FD57FD8E8584" box="[451,526,721,747]" italics="true" pageId="22" pageNumber="23">Alycus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C69E184FFFA43080C38FD57FD618584" authorityName="Duges" authorityYear="1834" box="[578,737,721,747]" class="Arachnida" family="Alycidae" genus="Pachygnathus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFFA43080C38FD57FD618584" box="[578,737,721,747]" italics="true" pageId="22" pageNumber="23">Pachygnathus</emphasis>
</taxonomicName>
were sometimes considered synonymous during 19th and 20th centuries until the phylogenetic analysis of the family
<taxonomicName id="4C69E184FFFA43080D72FD73FCED8460" baseAuthorityName="Uusitalo" baseAuthorityYear="2010" box="[776,877,757,783]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="family">Alycidae</taxonomicName>
revealed their belonging to distinct evolutionary lines (
<bibRefCitation id="EFF8E7F6FFFA43080EA1FC9FFE01845C" author="Uusitalo, M." box="[219,385,793,819]" pageId="22" pageNumber="23" refId="ref48736" refString="Uusitalo, M. (2010) Revision of family Alycidae (Acariformes, Acari), with special reference to European species. PhD thesis, University of Helsinki, Helsinki University Print, Helsinki, 143 pp. Available from: http://ethesis.helsinki.fi (accessed 20 October 2021)" type="book" year="2010">Uusitalo 2010</bibRefCitation>
). Both old generic names were retained as valid, the name
<taxonomicName id="4C69E184FFFA43080A66FC9FFB3B845C" authorityName="Duges" authorityYear="1834" box="[1052,1211,793,819]" class="Arachnida" family="Alycidae" genus="Pachygnathus" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B91D4615FFFA43080A66FC9FFB3B845C" box="[1052,1211,793,819]" italics="true" pageId="22" pageNumber="23">Pachygnathus</emphasis>
</taxonomicName>
has been applied to the group of species with long and beak-like chelicerae in the tribe
<taxonomicName id="4C69E184FFFA43080DFCFCBBFC5E8438" box="[902,990,829,855]" class="Arachnida" family="Alycidae" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="tribe" tribe="Alycini">Alycini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,362 @@
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<emphasis id="B9092F22FF9895734E89FD296BBBFD56" bold="true" box="[151,234,654,681]" italics="true" pageId="4" pageNumber="33">Feltria</emphasis>
(
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<uuid id="FFDBC9E5FF9895734C8CFD156FE6FD32" box="[658,1207,690,717]" pageId="4" pageNumber="33">D7D4C3C9-B896-4AA7-A2F3-D3194D9CE537</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9895734E89FD706BAEFD0E" box="[151,255,727,753]" pageId="4" pageNumber="33" type="description">
<paragraph id="8BC2F330FF9895734E89FD706BAEFD0E" blockId="4.[151,1207,654,753]" box="[151,255,727,753]" pageId="4" pageNumber="33">
<figureCitation id="1346EFB5FF9895734E89FD706BAEFD0E" box="[151,255,727,753]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[152,255,1596,1620]" captionStartId-1="6.[152,255,1943,1967]" captionTargetBox-0="[151,1435,179,1571]" captionTargetBox-1="[187,1380,182,1917]" captionTargetId-0="figure-18@5.[151,1436,179,1571]" captionTargetId-1="figure-28@6.[187,1399,179,1917]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Feltria tienshanensis sp. nov., ♂ holotype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp; D—I-L-5 and -6 (mounted); E—I-L-6 (unmounted). Scale bars = 100 µm." captionText-1="FIGURE 4. Feltria tienshanensis sp. nov., ♀ paratype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp. Scale bars = 100 µm." figureDoi-0="http://doi.org/10.5281/zenodo.15012498" figureDoi-1="http://doi.org/10.5281/zenodo.15012500" httpUri-0="https://zenodo.org/record/15012498/files/figure.png" httpUri-1="https://zenodo.org/record/15012500/files/figure.png" pageId="4" pageNumber="33">Figs. 34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9895734E89FCB86867FC12" pageId="4" pageNumber="33" type="materials_examined">
<paragraph id="8BC2F330FF9895734E89FCB8680EFC7E" blockId="4.[151,1437,798,1977]" pageId="4" pageNumber="33">
<emphasis id="B9092F22FF9895734E89FCB86A6FFCC7" bold="true" box="[151,318,798,825]" pageId="4" pageNumber="33">Type material</emphasis>
<materialsCitation id="3B15F96DFF9895734F44FCB86FD8FCA2" collectingDate="2013-08-15" collectionCode="RMNH" collectorName="Pesic &amp; Smit." country="Kyrgyzstan" elevation="2426" latitude="41.941467" location="Son-Kul Lake" longLatPrecision="1" longitude="75.5879" pageId="4" pageNumber="33" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C64D92FF9895734F44FCB86A93FCC6" box="[346,450,799,825]" pageId="4" pageNumber="33" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7B38B9FF9895734FD7FCB86AB1FCC6" box="[457,480,799,825]" count="1" pageId="4" pageNumber="33" type="male"></specimenCount>
, dissected and slide mounted (
<collectionCode id="ED6C6BF5FF9895734D28FCB868DFFCC6" box="[822,910,799,825]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="33" type="Museum">RMNH</collectionCode>
),
<collectingCountry id="F36AB3A0FF9895734DBFFCB86F75FCC6" box="[929,1060,799,825]" name="Kyrgyzstan" pageId="4" pageNumber="33">Kyrgyzstan</collectingCountry>
, river next to the road to
<location id="8EA2A5EBFF9895734B23FCB86B9EFCA2" LSID="urn:lsid:plazi:treatment:03D44226FF9895704E89FD296BBBFD56:8EA2A5EBFF9895734B23FCB86B9EFCA2" country="Kyrgyzstan" latitude="41.941467" longLatPrecision="1" longitude="75.5879" name="Son-Kul Lake" pageId="4" pageNumber="33">Son-Kul Lake</location>
(
<figureCitation id="1346EFB5FF9895734EC1FCE46A69FCA2" box="[223,312,834,861]" captionStart="FIGURE 5" captionStartId="9.[152,255,1885,1909]" captionTargetBox="[151,1436,1009,1859]" captionTargetId="figure-233@9.[151,1436,1009,1859]" captionTargetPageId="9" captionText="FIGURE 5. Photographs of sampling sites: A—Stream next to the road to May Saz pass, 2955 m asl., 12.viii.2013 (locus typicus of Feltria kyrgyzicus sp. nov.). B—River next to the road to Son-Kul Lake, 2426 m asl., 15.viii.2013 (locus typicus of Feltria tienshanensis sp. nov.). Photos by V. Pešić." figureDoi="http://doi.org/10.5281/zenodo.15012504" httpUri="https://zenodo.org/record/15012504/files/figure.png" pageId="4" pageNumber="33">Fig. 5B</figureCitation>
),
<geoCoordinate id="EE4995F7FF9895734F53FCE56AB4FCA2" box="[333,485,834,861]" degrees="41" direction="north" minutes="56.488" orientation="latitude" pageId="4" pageNumber="33" precision="1" value="41.941467">41º56.488 N</geoCoordinate>
,
<geoCoordinate id="EE4995F7FF9895734FEFFCE569D2FCA2" box="[497,643,834,861]" degrees="75" direction="east" minutes="35.274" orientation="longitude" pageId="4" pageNumber="33" precision="1" value="75.5879">75º35.274 E</geoCoordinate>
,
<elevation id="00501403FF9895734C8CFCE46842FCA2" box="[658,787,835,861]" metricMagnitude="3" metricUnit="m" metricValue="2.4259999999999997" pageId="4" pageNumber="33" unit="m" value="2426.0">
<quantity id="4C855ED5FF9895734C8CFCE469B9FCA2" box="[658,744,835,861]" metricMagnitude="3" metricUnit="m" metricValue="2.4259999999999997" pageId="4" pageNumber="33" unit="m" value="2426.0">2426 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9895734D3AFCE568FFFCA2" box="[804,942,834,861]" pageId="4" pageNumber="33" value="2013-08-15">
<collectingDate id="EF872C18FF9895734D3AFCE568FFFCA2" box="[804,942,834,861]" pageId="4" pageNumber="33" value="2013-08-15">15.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9895734DF2FCE46F76FCA2" box="[1004,1063,835,861]" pageId="4" pageNumber="33">Pešić</collectorName>
&amp;
<collectorName id="268896E6FF9895734A52FCE46FD8FCA2" box="[1100,1161,835,861]" pageId="4" pageNumber="33">Smit.</collectorName>
</materialsCitation>
<materialsCitation id="3B15F96DFF9895734A8EFCE4680AFC7E" collectingDate="2013-08-15" collectionCode="RMNH" collectorName="Pesic &amp; Smit." country="Kyrgyzstan" elevation="2426" latitude="41.941467" location="Son-Kul Lake" longLatPrecision="1" longitude="75.5879" pageId="4" pageNumber="33" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54C64D92FF9895734A8EFCE46E50FCA2" box="[1168,1281,835,861]" pageId="4" pageNumber="33" type="paratype">Paratypes</typeStatus>
: 3/30/0, same data as the holotype,
<specimenCount id="9D7B38B9FF9895734F9AFCC06AF6FC7E" box="[388,423,871,897]" count="1" pageId="4" pageNumber="33" type="female">1♀</specimenCount>
dissected and slide mounted (
<collectionCode id="ED6C6BF5FF9895734CE4FCC06802FC7E" box="[762,851,871,897]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="33" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC2F330FF9895734ED9FC2D6867FC12" blockId="4.[151,1437,798,1977]" pageId="4" pageNumber="33">
<emphasis id="B9092F22FF9895734ED9FC2D6A2DFC5B" bold="true" box="[199,380,906,932]" pageId="4" pageNumber="33">Other material</emphasis>
<materialsCitation id="3B15F96DFF9895734F84FC2C6862FC12" collectingDate="2013-08-16" collectionCode="RMNH" collectorName="Smit &amp; Pesic" country="Kyrgyzstan" county="Son-Kul lake" elevation="3065" latitude="41.85915" location="Dzhaman-Echki" longLatPrecision="1" longitude="75.33548" municipality="Aktash river" pageId="4" pageNumber="33" specimenCount="1">
<collectingCountry id="F36AB3A0FF9895734F84FC2C694CFC5A" box="[410,541,907,933]" name="Kyrgyzstan" pageId="4" pageNumber="33">Kyrgyzstan</collectingCountry>
:
<location id="8EA2A5EBFF9895734C37FC2C698EFC5A" LSID="urn:lsid:plazi:treatment:03D44226FF9895704E89FD296BBBFD56:8EA2A5EBFF9895734C37FC2C698EFC5A" box="[553,735,907,933]" country="Kyrgyzstan" county="Son-Kul lake" latitude="41.85915" longLatPrecision="1" longitude="75.33548" municipality="Aktash river" name="Dzhaman-Echki" pageId="4" pageNumber="33">Dzhaman-Echki</location>
stream flowing into
<location id="8EA2A5EBFF9895734DD1FC2C6F23FC5A" LSID="urn:lsid:plazi:treatment:03D44226FF9895704E89FD296BBBFD56:8EA2A5EBFF9895734DD1FC2C6F23FC5A" box="[975,1138,907,933]" country="Kyrgyzstan" county="Son-Kul lake" latitude="41.85915" longLatPrecision="1" longitude="75.33548" municipality="Aktash river" name="Son-Kul Lake" pageId="4" pageNumber="33">Son-Kul Lake</location>
,
<geoCoordinate id="EE4995F7FF9895734A61FC2D6E46FC5A" box="[1151,1303,906,933]" degrees="41" direction="north" minutes="55.482" orientation="latitude" pageId="4" pageNumber="33" precision="1" value="41.9247">41º55.482 N</geoCoordinate>
,
<geoCoordinate id="EE4995F7FF9895734B3AFC2D6BFBFC36" degrees="75" direction="east" minutes="11.861" orientation="longitude" pageId="4" pageNumber="33" precision="1" value="75.197685">75º11.861 E</geoCoordinate>
,
<elevation id="00501403FF9895734EA8FC096A64FC36" box="[182,309,942,969]" metricMagnitude="3" metricUnit="m" metricValue="3.065" pageId="4" pageNumber="33" unit="m" value="3065.0">
<quantity id="4C855ED5FF9895734EA8FC096A5AFC36" box="[182,267,942,969]" metricMagnitude="3" metricUnit="m" metricValue="3.065" pageId="4" pageNumber="33" unit="m" value="3065.0">3065 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9895734F5BFC086A9EFC36" box="[325,463,943,969]" pageId="4" pageNumber="33" value="2013-08-16">
<collectingDate id="EF872C18FF9895734F5BFC086A9EFC36" box="[325,463,943,969]" pageId="4" pageNumber="33" value="2013-08-16">16.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9895734C14FC08696EFC36" box="[522,575,943,969]" pageId="4" pageNumber="33">Smit</collectorName>
1/2/0 (
<collectionCode id="ED6C6BF5FF9895734C91FC0869B9FC36" box="[655,744,943,969]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="33" type="Museum">RMNH</collectionCode>
);
<collectingMunicipality id="6BA6694AFF9895734CE7FC0868D5FC36" box="[761,900,943,969]" pageId="4" pageNumber="33">Aktash river</collectingMunicipality>
near
<collectingCounty id="62A38BBCFF9895734DDFFC086F09FC36" box="[961,1112,943,969]" pageId="4" pageNumber="33">Son-Kul lake</collectingCounty>
,
<geoCoordinate id="EE4995F7FF9895734A7DFC096FABFC36" box="[1123,1274,942,969]" degrees="41" direction="north" minutes="51.549" orientation="latitude" pageId="4" pageNumber="33" precision="1" value="41.85915">41º51.549 N</geoCoordinate>
,
<geoCoordinate id="EE4995F7FF9895734B1AFC096EC6FC36" box="[1284,1431,942,969]" degrees="75" direction="east" minutes="20.129" orientation="longitude" pageId="4" pageNumber="33" precision="1" value="75.33548">75º20.129 E</geoCoordinate>
,
<elevation id="00501403FF9895734E89FC746A46FC12" box="[151,279,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="3.048" pageId="4" pageNumber="33" unit="m" value="3048.0">
<quantity id="4C855ED5FF9895734E89FC746BBDFC12" box="[151,236,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="3.048" pageId="4" pageNumber="33" unit="m" value="3048.0">3048 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9895734F39FC746AE0FC12" box="[295,433,979,1005]" pageId="4" pageNumber="33" value="2013-08-16">
<collectingDate id="EF872C18FF9895734F39FC746AE0FC12" box="[295,433,979,1005]" pageId="4" pageNumber="33" value="2013-08-16">16.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9895734FF3FC746979FC12" box="[493,552,979,1005]" pageId="4" pageNumber="33">Pešić</collectorName>
&amp;
<collectorName id="268896E6FF9895734C52FC7469D0FC12" box="[588,641,979,1005]" pageId="4" pageNumber="33">Smit</collectorName>
0/1/0 (
<collectionCode id="ED6C6BF5FF9895734CCFFC74687BFC12" box="[721,810,979,1005]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="33" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9895734ED9FC516B84FB16" pageId="4" pageNumber="33" type="diagnosis">
<paragraph id="8BC2F330FF9895734ED9FC516B84FB16" blockId="4.[151,1437,798,1977]" pageId="4" pageNumber="33">
<emphasis id="B9092F22FF9895734ED9FC516A6BFBEF" bold="true" box="[199,314,1014,1040]" pageId="4" pageNumber="33">Diagnosis</emphasis>
—Both sexes: Coxal plates in four groups. Genital field with 4055 pairs of Ac. Male: Frontal area with a pair of rod-shaped frontal plates including pre-antenniform glandularia, pre-ocular setae and Lgl-1+2; excretory pore plate and flanking glandularia fused to the posterior margin of the genital plate. III-L-6 with a rounded ventrolateral projection in the distal third, bearing 56 setae. Female: Frontal area with pre-antenniform glandularia, pre-ocular setae and Lgl-1 fused to each other; dorsum with a dorsal shield, much longer than wider, L/W ratio 1.3, surrounded by four rounded platelets. Excretory pore in posteroventral position. Palp slender than in male.
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9895724ED9FB55699FF8AE" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33" type="description">
<paragraph id="8BC2F330FF9895734ED9FB5569A9FA62" blockId="4.[151,1437,798,1977]" pageId="4" pageNumber="33">
<emphasis id="B9092F22FF9895734ED9FB556A00FAF3" bold="true" box="[199,337,1266,1292]" pageId="4" pageNumber="33">Description</emphasis>
<emphasis id="B9092F22FF9895734F70FB546AF9FAF2" box="[366,424,1267,1293]" italics="true" pageId="4" pageNumber="33">Male</emphasis>
(
<typeStatus id="54C64D92FF9895734FA8FB54694EFAF2" box="[438,543,1267,1293]" pageId="4" pageNumber="33" type="holotype">holotype</typeStatus>
): Idiosoma length 359 (
<typeStatus id="54C64D92FF9895734D39FB5468C5FAF2" box="[807,916,1267,1293]" pageId="4" pageNumber="33" type="paratype">paratypes</typeStatus>
[n=3] from river on the road to Son-Kul Lake
<quantity id="4C855ED5FF9895734E89FAB16A45FACE" box="[151,276,1302,1329]" metricMagnitude="0" metricUnit="m" metricValue="9.0297" metricValueMax="9.1186" metricValueMin="8.9408" pageId="4" pageNumber="33" unit="in" value="355.5" valueMax="359.0" valueMin="352.0">352359 in</quantity>
lenght and
<quantity id="4C855ED5FF9895734F8DFAB06941FACE" box="[403,528,1303,1329]" metricMagnitude="0" metricUnit="m" metricValue="7.0104" metricValueMax="7.3152" metricValueMin="6.7056" pageId="4" pageNumber="33" unit="in" value="276.0" valueMax="288.0" valueMin="264.0">264288 in</quantity>
width, length/width ratio 1.231.36). Dorsum with a large dorsal shield (including Dl-1-3 and Dgl-1-3),
<quantity id="4C855ED5FF9895734F91FA9C6A89FAAA" box="[399,472,1339,1365]" metricMagnitude="0" metricUnit="m" metricValue="7.3914" pageId="4" pageNumber="33" unit="in" value="291.0">291 in</quantity>
length and 241 width, Dgl-4+Dl-4 on a pair of large, transverse platelets (
<figureCitation id="1346EFB5FF9895734B2DFA9C6EDEFAAA" box="[1331,1423,1339,1365]" captionStart="FIGURE 3" captionStartId="5.[152,255,1596,1620]" captionTargetBox="[151,1435,179,1571]" captionTargetId="figure-18@5.[151,1436,179,1571]" captionTargetPageId="5" captionText="FIGURE 3. Feltria tienshanensis sp. nov., ♂ holotype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp; D—I-L-5 and -6 (mounted); E—I-L-6 (unmounted). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012498" httpUri="https://zenodo.org/record/15012498/files/figure.png" pageId="4" pageNumber="33">Fig. 3A</figureCitation>
). Coxal plates in four groups, close to each other, coxal field
<quantity id="4C855ED5FF9895734D52FAF868C7FA86" box="[844,918,1375,1401]" metricMagnitude="0" metricUnit="m" metricValue="6.832599999999999" pageId="4" pageNumber="33" unit="in" value="269.0">269 in</quantity>
length, Cx-III
<quantity id="4C855ED5FF9895734A5BFAF86FDFFA86" box="[1093,1166,1375,1401]" metricMagnitude="0" metricUnit="m" metricValue="5.4864" pageId="4" pageNumber="33" unit="in" value="216.0">216 in</quantity>
width; Cx-IV posterior margin perpendicularly to longitudinal axis, (
<figureCitation id="1346EFB5FF9895734C8CFA2469BBFA62" box="[658,746,1411,1437]" captionStart="FIGURE 3" captionStartId="5.[152,255,1596,1620]" captionTargetBox="[151,1435,179,1571]" captionTargetId="figure-18@5.[151,1436,179,1571]" captionTargetPageId="5" captionText="FIGURE 3. Feltria tienshanensis sp. nov., ♂ holotype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp; D—I-L-5 and -6 (mounted); E—I-L-6 (unmounted). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012498" httpUri="https://zenodo.org/record/15012498/files/figure.png" pageId="4" pageNumber="33">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="8BC2F330FF9895734ED9FA006A54F9D2" blockId="4.[151,1437,798,1977]" pageId="4" pageNumber="33">
Genital plate separated from Cx-IV by a narrow strip of membrane bearing the Vgl and a triangular lateral platelet, transverse,
<quantity id="4C855ED5FF9895734F68FA6C6AECFA1A" box="[374,445,1483,1509]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="33" unit="in" value="100.0">100 in</quantity>
length and
<quantity id="4C855ED5FF9895734C21FA6C69D7FA1A" box="[575,646,1483,1509]" metricMagnitude="0" metricUnit="m" metricValue="6.858" pageId="4" pageNumber="33" unit="in" value="270.0">270 in</quantity>
width, with 5455 pairs of acetabula; gonopore short, length 14, in the center; platelet bearing the excretory pore broad, in the indentation of posterior margin of genital plate, fused to it (
<figureCitation id="1346EFB5FF9895734E81F9B46BA6F9D2" box="[159,247,1555,1581]" captionStart="FIGURE 3" captionStartId="5.[152,255,1596,1620]" captionTargetBox="[151,1435,179,1571]" captionTargetId="figure-18@5.[151,1436,179,1571]" captionTargetPageId="5" captionText="FIGURE 3. Feltria tienshanensis sp. nov., ♂ holotype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp; D—I-L-5 and -6 (mounted); E—I-L-6 (unmounted). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012498" httpUri="https://zenodo.org/record/15012498/files/figure.png" pageId="4" pageNumber="33">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="8BC2F330FF9895734ED9F9906F66F98A" blockId="4.[151,1437,798,1977]" pageId="4" pageNumber="33">
Palp robust and stout, P-4 equally narrowed from base to tip (
<figureCitation id="1346EFB5FF9895734D6DF990689BF9AE" box="[883,970,1591,1617]" captionStart="FIGURE 3" captionStartId="5.[152,255,1596,1620]" captionTargetBox="[151,1435,179,1571]" captionTargetId="figure-18@5.[151,1436,179,1571]" captionTargetPageId="5" captionText="FIGURE 3. Feltria tienshanensis sp. nov., ♂ holotype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp; D—I-L-5 and -6 (mounted); E—I-L-6 (unmounted). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012498" httpUri="https://zenodo.org/record/15012498/files/figure.png" pageId="4" pageNumber="33">Fig. 3C</figureCitation>
); dorsal length/height of palpal segments: P-1, 23/31; P-2, 64/52; P-3, 33/41; P-4, 80/33; P-5, 42/20; length ratio P-2/P-4 0.8.
</paragraph>
<paragraph id="8BC2F330FF9895734ED9F9D86F3AF942" blockId="4.[151,1437,798,1977]" pageId="4" pageNumber="33">
III-L-6 with a rounded ventrolateral projection in the distal third, bearing 56 setae (
<figureCitation id="1346EFB5FF9895734A74F9D86FBFF966" box="[1130,1262,1663,1689]" captionStart="FIGURE 3" captionStartId="5.[152,255,1596,1620]" captionTargetBox="[151,1435,179,1571]" captionTargetId="figure-18@5.[151,1436,179,1571]" captionTargetPageId="5" captionText="FIGURE 3. Feltria tienshanensis sp. nov., ♂ holotype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp; D—I-L-5 and -6 (mounted); E—I-L-6 (unmounted). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012498" httpUri="https://zenodo.org/record/15012498/files/figure.png" pageId="4" pageNumber="33">Figs. 3DE</figureCitation>
); dorsal lengths of III-L-1-6: 28, 44, 54, 75, 88, 98; dorsal lengths of IV-L-1-6: 61, 45, 61, 92, 103, 113.
</paragraph>
<paragraph id="8BC2F330FF9895734ED9F9606A76F846" blockId="4.[151,1437,798,1977]" pageId="4" pageNumber="33">
<emphasis id="B9092F22FF9895734ED9F9606A4AF91E" box="[199,283,1735,1761]" italics="true" pageId="4" pageNumber="33">Female</emphasis>
(from river next to the road to Son-Kul Lake; in parentheses some measurements of conserved specimens): Idiosoma 419 (422466, n=5) in length and 359 (330397, n=5) in width and
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width, length/ width ratio 1.17 (1.171.28, n=3); dorsal shield, including Dgl-
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length and
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width; Dgl-1 on platelets anterolateral to dorsal shield margins. Dl-2 elongated, Dl-3 and Dgl-3 separate from each other, Dgl-4+Dl-4 on a pair of large, transverse platelets (
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); excretory pore platelet broad, directed posteroventrally. Coxal field length 273, Cx-III width 256. Genital plates subtriangular,
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length and
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width, with about 4052 pairs of acetabula.
</paragraph>
<caption id="DF02A3B8FF9995724E86F99B68BCF986" ID-DOI="http://doi.org/10.5281/zenodo.15012498" ID-Zenodo-Dep="15012498" httpUri="https://zenodo.org/record/15012498/files/figure.png" pageId="5" pageNumber="34" startId="5.[152,255,1596,1620]" targetBox="[151,1435,179,1571]" targetPageId="5" targetType="figure">
<paragraph id="8BC2F330FF9995724E86F99B68BCF986" blockId="5.[151,1437,1596,1657]" pageId="5" pageNumber="34">
<emphasis id="B9092F22FF9995724E86F99B6A47F9AA" bold="true" box="[152,278,1596,1621]" pageId="5" pageNumber="34">FIGURE 3</emphasis>
.
<taxonomicName id="4C7D88B3FF9995724F3BF99A6AAFF9AA" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[293,510,1597,1621]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="tienshanensis" status="sp. nov.">
<emphasis id="B9092F22FF9995724F3BF99A6AAFF9AA" box="[293,510,1597,1621]" italics="true" pageId="5" pageNumber="34">Feltria tienshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A9259FF9995724C19F99A690DF9AB" box="[519,604,1597,1621]" pageId="5" pageNumber="34" rank="species">
<emphasis id="B9092F22FF9995724C19F99A6907F9AA" bold="true" box="[519,598,1597,1621]" pageId="5" pageNumber="34">sp. nov</emphasis>
.
</taxonomicNameLabel>
, ♂ holotype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp; D—I-L-5 and -6 (mounted); E—I-L-6 (unmounted). Scale bars = 100 µm.
</paragraph>
</caption>
<paragraph id="8BC2F330FF9995724ED9F96C699FF8AE" blockId="5.[151,1437,1739,1945]" pageId="5" pageNumber="34">
Chaetotaxy of palp as given in
<figureCitation id="1346EFB5FF9995724C33F96C69D4F91A" box="[557,645,1739,1765]" captionStart="FIGURE 4" captionStartId="6.[152,255,1943,1967]" captionTargetBox="[187,1380,182,1917]" captionTargetId="figure-28@6.[187,1399,179,1917]" captionTargetPageId="6" captionText="FIGURE 4. Feltria tienshanensis sp. nov., ♀ paratype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012500" httpUri="https://zenodo.org/record/15012500/files/figure.png" pageId="5" pageNumber="34">Fig. 4C</figureCitation>
; dorsal length/height of palpal segments: P-1, 21/23; P-2, 59/47; P-3, 34/35; P-4, 81/28; P-5, 45/17; length ratio P-2/P-4 0.73. Gnathosoma ventral length 108. III-L-6 without ventrolateral projection. Dorsal lengths of I-L-1-6: 41, 39, 45, 61, 67, 83; I-L-6 height 29, I-L-6 dorsal length/height ratio 2.9; dorsal lengths of IV-L-1-6: 72, 45, 63, 86, 97, 116.
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9995724ED9F8FC68FEF866" pageId="5" pageNumber="34" type="etymology">
<paragraph id="8BC2F330FF9995724ED9F8FC68FEF866" blockId="5.[151,1437,1739,1945]" pageId="5" pageNumber="34">
<emphasis id="B9092F22FF9995724ED9F8FC6A19F88A" bold="true" box="[199,328,1883,1909]" pageId="5" pageNumber="34">Etymology</emphasis>
—The new species is named after the Tien Shan Mountain range from where the new species was collected. The name is an adjective derived from a geographical name.
</paragraph>
</subSubSection>
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<paragraph id="8BC2F330FF9A95714E86F8306A82F82B" blockId="6.[151,1437,1943,2004]" pageId="6" pageNumber="35">
<emphasis id="B9092F22FF9A95714E86F8306A47F850" bold="true" box="[152,278,1943,1967]" pageId="6" pageNumber="35">FIGURE 4</emphasis>
.
<taxonomicName id="4C7D88B3FF9A95714F3BF83F6AAFF84F" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[293,510,1944,1968]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="tienshanensis" status="sp. nov.">
<emphasis id="B9092F22FF9A95714F3BF83F6AAFF84F" box="[293,510,1944,1968]" italics="true" pageId="6" pageNumber="35">Feltria tienshanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A9259FF9A95714C19F83F690CF850" box="[519,605,1944,1968]" pageId="6" pageNumber="35" rank="species">
<emphasis id="B9092F22FF9A95714C19F83F6907F84F" bold="true" box="[519,598,1944,1968]" pageId="6" pageNumber="35">sp. nov</emphasis>
.
</taxonomicNameLabel>
, ♀ paratype, river next to the road to Son-Kul Lake: A—Dorsum; B—Venter; C—Palp. Scale bars = 100 µm.
</paragraph>
</caption>
<subSubSection id="C367A0BBFF9B95704ED9FF316F0FFD9E" pageId="7" pageNumber="36" type="discussion">
<paragraph id="8BC2F330FF9B95704ED9FF3168A2FE0A" blockId="7.[151,1437,150,681]" pageId="7" pageNumber="36">
<emphasis id="B9092F22FF9B95704ED9FF316A14FF4F" bold="true" box="[199,325,150,176]" pageId="7" pageNumber="36">Discussion</emphasis>
<taxonomicName id="4C7D88B3FF9B95704F7FFF3069B8FF4E" authority="Tuzovskij, 1988" authorityName="Tuzovskij" authorityYear="1988" box="[353,745,151,177]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="kuluensis">
<emphasis id="B9092F22FF9B95704F7FFF306973FF4E" box="[353,546,151,177]" italics="true" pageId="7" pageNumber="36">Feltria kuluensis</emphasis>
<bibRefCitation id="EFEC8EC1FF9B95704C33FF3069B8FF4E" author="Tuzovskij, P. V." box="[557,745,151,177]" pageId="7" pageNumber="36" pagination="218 - 230" refId="ref6365" refString="Tuzovskij, P. V. (1988) Novyje vidy vodjanych klescej roda Feltria Koenike (Acariformes, Feltriidae). Entomologicheskoe Obozrenie, 67 (1), 218-230. [in Russian]" type="journal article" year="1988">Tuzovskij, 1988</bibRefCitation>
</taxonomicName>
, with a distribution from the Far East of
<collectingCountry id="F36AB3A0FF9B95704ACBFF306E71FF4E" box="[1237,1312,151,177]" name="Russia" pageId="7" pageNumber="36">Russia</collectingCountry>
(
<collectingRegion id="49B93DD2FF9B95704B2CFF306BB3FF2A" country="Russia" name="Magadan" pageId="7" pageNumber="36">Magadan region</collectingRegion>
;
<bibRefCitation id="EFEC8EC1FF9B95704EF1FF1C6ACEFF2A" author="Tuzovskij, P. V." box="[239,415,187,213]" pageId="7" pageNumber="36" pagination="218 - 230" refId="ref6365" refString="Tuzovskij, P. V. (1988) Novyje vidy vodjanych klescej roda Feltria Koenike (Acariformes, Feltriidae). Entomologicheskoe Obozrenie, 67 (1), 218-230. [in Russian]" type="journal article" year="1988">Tuzovskij 1988</bibRefCitation>
) to
<collectingCountry id="F36AB3A0FF9B95704FD0FF1C690DFF2A" box="[462,604,187,213]" name="South Korea" pageId="7" pageNumber="36">South Korea</collectingCountry>
(Pešić
<emphasis id="B9092F22FF9B95704CACFF1C69B4FF2A" box="[690,741,187,213]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
. 2015) is similar to the new species in the shape of the male III-L-5/6 and the arrangement of dorsal sclerites in both sexes. At present state of knowledge, males of the new species are separated by the rod-shaped frontal plates that include praeantenniform glandularia, praeocular setae and Lgl-1+2 (in
<taxonomicName id="4C7D88B3FF9B95704F88FE80694BFEBE" authorityName="Tuzovskij" authorityYear="1988" box="[406,538,295,321]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="kuluensis">
<emphasis id="B9092F22FF9B95704F88FE80694BFEBE" box="[406,538,295,321]" italics="true" pageId="7" pageNumber="36">F. kuluensis</emphasis>
</taxonomicName>
frontal plates do not reach Lgl-2, see
<bibRefCitation id="EFEC8EC1FF9B95704DDDFE806F20FEBE" author="Tuzovskij, P. V." box="[963,1137,295,321]" pageId="7" pageNumber="36" pagination="218 - 230" refId="ref6365" refString="Tuzovskij, P. V. (1988) Novyje vidy vodjanych klescej roda Feltria Koenike (Acariformes, Feltriidae). Entomologicheskoe Obozrenie, 67 (1), 218-230. [in Russian]" type="journal article" year="1988">Tuzovskij 1988</bibRefCitation>
fig. 7/1) and the excretory pore plate and the flanking glandularia fused to the posterior genital plate margin. The female of
<taxonomicName id="4C7D88B3FF9B95704AF6FEEC6E3CFE9A" authorityName="Tuzovskij" authorityYear="1988" box="[1256,1389,331,357]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="kuluensis">
<emphasis id="B9092F22FF9B95704AF6FEEC6E3CFE9A" box="[1256,1389,331,357]" italics="true" pageId="7" pageNumber="36">F. kuluensis</emphasis>
</taxonomicName>
can be distinguished by the dorsal shield only slightly longer than wide (length/width ratio
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<taxonomicName id="4C7D88B3FF9B95704AC6FEC86E0CFE76" authorityName="Tuzovskij" authorityYear="1988" box="[1240,1373,367,393]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="kuluensis">
<emphasis id="B9092F22FF9B95704AC6FEC86E0CFE76" box="[1240,1373,367,393]" italics="true" pageId="7" pageNumber="36">F. kuluensis</emphasis>
</taxonomicName>
, data taken from
<bibRefCitation id="EFEC8EC1FF9B95704F06FE346A99FE52" author="Tuzovskij, P. V." box="[280,456,403,429]" pageId="7" pageNumber="36" pagination="218 - 230" refId="ref6365" refString="Tuzovskij, P. V. (1988) Novyje vidy vodjanych klescej roda Feltria Koenike (Acariformes, Feltriidae). Entomologicheskoe Obozrenie, 67 (1), 218-230. [in Russian]" type="journal article" year="1988">Tuzovskij 1988</bibRefCitation>
;
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<taxonomicName id="4C7D88B3FF9B95704C05FE34699CFE52" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[539,717,403,429]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="tienshahensis" status="sp. nov.">
<emphasis id="B9092F22FF9B95704C05FE34699CFE52" box="[539,717,403,429]" italics="true" pageId="7" pageNumber="36">F. tienshahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A9259FF9B95704CCAFE34687CFE52" box="[724,813,403,429]" pageId="7" pageNumber="36" rank="species">
<emphasis id="B9092F22FF9B95704CCAFE346879FE52" bold="true" box="[724,808,403,429]" pageId="7" pageNumber="36">sp. nov</emphasis>
.
</taxonomicNameLabel>
) and the frontal area with praeantenniform glandularia, praeocular setae and Lgl-1 all lying on separate platelets (see
<bibRefCitation id="EFEC8EC1FF9B95704D54FE1068A8FE2E" author="Tuzovskij, P. V." box="[842,1017,439,465]" pageId="7" pageNumber="36" pagination="218 - 230" refId="ref6365" refString="Tuzovskij, P. V. (1988) Novyje vidy vodjanych klescej roda Feltria Koenike (Acariformes, Feltriidae). Entomologicheskoe Obozrenie, 67 (1), 218-230. [in Russian]" type="journal article" year="1988">Tuzovskij 1988</bibRefCitation>
fig. 7/5) (in
<taxonomicName id="4C7D88B3FF9B95704A99FE106E6BFE2E" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[1159,1338,439,465]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="tienshahensis" status="sp. nov.">
<emphasis id="B9092F22FF9B95704A99FE106E6BFE2E" box="[1159,1338,439,465]" italics="true" pageId="7" pageNumber="36">F. tienshahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A9259FF9B95704B5FFE106ECDFE2E" box="[1345,1436,439,465]" pageId="7" pageNumber="36" rank="species">
<emphasis id="B9092F22FF9B95704B5FFE106EC7FE2E" bold="true" box="[1345,1430,439,465]" pageId="7" pageNumber="36">sp. nov</emphasis>
.
</taxonomicNameLabel>
pre-antenniform glandularia, pre-ocular setae and Lgl-1 fused to each other).
</paragraph>
<paragraph id="8BC2F330FF9B95704ED9FE586F0FFD9E" blockId="7.[151,1437,150,681]" pageId="7" pageNumber="36">
<taxonomicName id="4C7D88B3FF9B95704ED9FE58694AFDE6" authority="Koenike, 1892" authorityName="Koenike" authorityYear="1892" box="[199,539,511,537]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B9092F22FF9B95704ED9FE586A3AFDE7" box="[199,363,511,537]" italics="true" pageId="7" pageNumber="36">Feltria minuta</emphasis>
Koenike, 1892
</taxonomicName>
, differs in P-4 with both ventral setae on slightly elevated, parallel longitudinal extensions, and male IV-L-6 relatively shorter with ventrolateral extension bearing one fine seta and two enlarged, transparent setae, adpressed to each other and directed ventrally (
<bibRefCitation id="EFEC8EC1FF9B95704D6EFDE06F03FD9E" author="Gerecke, R. &amp; Schatz, H. &amp; Wohltmann, A." box="[880,1106,583,609]" pageId="7" pageNumber="36" pagination="303 - 333" refId="ref5372" refString="Gerecke, R., Schatz, H. &amp; Wohltmann, A. (2009) The mites (Chelicerata: Acari) of the CRENODAT project: Faunistic records and ecological data from springs in the Autonomous Province of Trento (Italian Alps). International Journal of Acarology, 35, 303-333. https://doi.org/10.1080/01647950903059452" type="journal article" year="2009">
Gerecke
<emphasis id="B9092F22FF9B95704DC8FDE06F5EFD9E" box="[982,1039,583,609]" italics="true" pageId="7" pageNumber="36">et al.</emphasis>
2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9B95704ED9FDCD6BBBFD56" pageId="7" pageNumber="36" type="distribution">
<paragraph id="8BC2F330FF9B95704ED9FDCD6BBBFD56" blockId="7.[151,1437,150,681]" pageId="7" pageNumber="36">
<emphasis id="B9092F22FF9B95704ED9FDCD6A08FD7B" bold="true" box="[199,345,618,644]" pageId="7" pageNumber="36">Distribution</emphasis>
<collectingCountry id="F36AB3A0FF9B95704F69FDCC6AABFD7A" box="[375,506,619,645]" name="Kyrgyzstan" pageId="7" pageNumber="36">Kyrgyzstan</collectingCountry>
; know from streams in the Tien Shan mountains at an elevation of about 24003100 meters.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,182 @@
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<mods:title id="052E7272C98D9E436907AAB6F37CB528">Two new species of the genus Feltria Koenike, 1892, with notes on the water mite fauna of Kyrgyzstan (Acari, Hydrachnidia)</mods:title>
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<taxonomicName id="4C7D88B3FF9B95704E89FD5D6938FCEB" authority="Piersig, 1898" authorityName="Piersig" authorityYear="1898" box="[151,617,762,789]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="rubra" subGenus="Feltriella">
<emphasis id="B9092F22FF9B95704E89FD5D6938FCEB" bold="true" box="[151,617,762,789]" pageId="7" pageNumber="36">
<emphasis id="B9092F22FF9B95704E89FD5D6BBBFCEA" bold="true" box="[151,234,762,789]" italics="true" pageId="7" pageNumber="36">Feltria</emphasis>
(
<emphasis id="B9092F22FF9B95704EE5FD5D6A3AFCEA" bold="true" box="[251,363,762,789]" italics="true" pageId="7" pageNumber="36">Feltriella</emphasis>
)
<emphasis id="B9092F22FF9B95704F63FD5D6A90FCEA" bold="true" box="[381,449,762,789]" italics="true" pageId="7" pageNumber="36">rubra</emphasis>
Piersig, 1898
</emphasis>
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</paragraph>
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<emphasis id="B9092F22FF9B95704E89FCE56A2DFCA3" bold="true" box="[151,380,834,860]" pageId="7" pageNumber="36">Material examined</emphasis>
<materialsCitation id="3B15F96DFF9B95704F87FCE46947FC7E" collectingDate="2013-08-09" collectorName="Ala Archa NP &amp; Pesic &amp; Smit" country="Kyrgyzstan" elevation="1877" latitude="42.603382" location="Archa River" longLatPrecision="1" longitude="74.47227" pageId="7" pageNumber="36" specimenCount="1">
<collectingCountry id="F36AB3A0FF9B95704F87FCE4694DFCA2" box="[409,540,835,861]" name="Kyrgyzstan" pageId="7" pageNumber="36">Kyrgyzstan</collectingCountry>
:
<collectorName id="268896E6FF9B95704C36FCE4699BFCA2" box="[552,714,835,861]" pageId="7" pageNumber="36">Ala Archa NP</collectorName>
, tributary of
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,
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,
<geoCoordinate id="EE4995F7FF9B95704ABFFCE46E66FCA2" box="[1185,1335,835,861]" degrees="74" direction="east" minutes="28.336" orientation="longitude" pageId="7" pageNumber="36" precision="1" value="74.47227">74º28.336 E</geoCoordinate>
,
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<quantity id="4C855ED5FF9B95704B5BFCE46ECDFCA2" box="[1349,1436,835,861]" metricMagnitude="3" metricUnit="m" metricValue="1.877" pageId="7" pageNumber="36" unit="m" value="1877.0">1877 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9B95704ED5FCC06A16FC7E" box="[203,327,871,897]" pageId="7" pageNumber="36" value="2013-08-09">
<collectingDate id="EF872C18FF9B95704ED5FCC06A16FC7E" box="[203,327,871,897]" pageId="7" pageNumber="36" value="2013-08-09">9.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9B95704F9FFCC06AEDFC7E" box="[385,444,871,897]" pageId="7" pageNumber="36">Pešić</collectorName>
&amp;
<collectorName id="268896E6FF9B95704FC0FCC06947FC7E" box="[478,534,871,897]" pageId="7" pageNumber="36">Smit</collectorName>
</materialsCitation>
,
<materialsCitation id="3B15F96DFF9B95704C3FFCC06892FC5A" collectingDate="2013-08-12" collectorName="Pesic &amp; Smit" country="Kyrgyzstan" elevation="3346" latitude="42.41715" location="Karakol region" longLatPrecision="1" longitude="79.0443" municipality="May Saz" pageId="7" pageNumber="36" specimenCount="1">
1/1/0;
<location id="8EA2A5EBFF9B95704C76FCC06842FC7E" LSID="urn:lsid:plazi:treatment:03D44226FF9B95704E89FD5D68EEFB3A:8EA2A5EBFF9B95704C76FCC06842FC7E" box="[616,787,871,897]" country="Kyrgyzstan" latitude="42.41715" longLatPrecision="1" longitude="79.0443" municipality="May Saz" name="Karakol region" pageId="7" pageNumber="36">Karakol region</location>
, upper part of the stream next to the road to
<collectingMunicipality id="6BA6694AFF9B95704AE3FCC06E0FFC7E" box="[1277,1374,871,897]" pageId="7" pageNumber="36">May Saz</collectingMunicipality>
pass,
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,
<geoCoordinate id="EE4995F7FF9B95704F27FC2D6A9BFC5A" box="[313,458,906,933]" degrees="79" direction="east" minutes="02.658" orientation="longitude" pageId="7" pageNumber="36" precision="1" value="79.0443">79º02.658 E</geoCoordinate>
,
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<quantity id="4C855ED5FF9B95704FC6FC2C697CFC5A" box="[472,557,907,933]" metricMagnitude="3" metricUnit="m" metricValue="3.346" pageId="7" pageNumber="36" unit="m" value="3346.0">3346 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9B95704C79FC2C69A3FC5A" box="[615,754,907,933]" pageId="7" pageNumber="36" value="2013-08-12">
<collectingDate id="EF872C18FF9B95704C79FC2C69A3FC5A" box="[615,754,907,933]" pageId="7" pageNumber="36" value="2013-08-12">12.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9B95704D33FC2C6839FC5A" box="[813,872,907,933]" pageId="7" pageNumber="36">Pešić</collectorName>
&amp;
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</materialsCitation>
,
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0/1/0; stream next to the road to
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pass,
<geoCoordinate id="EE4995F7FF9B95704ECBFC096A3DFC36" box="[213,364,942,969]" degrees="42" direction="north" minutes="25.002" orientation="latitude" pageId="7" pageNumber="36" precision="1" value="42.4167">42º25.002 N</geoCoordinate>
,
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,
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<quantity id="4C855ED5FF9B95704C09FC09693DFC36" box="[535,620,942,969]" metricMagnitude="3" metricUnit="m" metricValue="2.955" pageId="7" pageNumber="36" unit="m" value="2955.0">2955 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9B95704CB8FC086860FC36" box="[678,817,943,969]" pageId="7" pageNumber="36" value="2013-08-12">
<collectingDate id="EF872C18FF9B95704CB8FC086860FC36" box="[678,817,943,969]" pageId="7" pageNumber="36" value="2013-08-12">12.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9B95704D72FC0868F6FC36" box="[876,935,943,969]" pageId="7" pageNumber="36">Pešić</collectorName>
&amp;
<collectorName id="268896E6FF9B95704DD5FC086F52FC36" box="[971,1027,943,969]" pageId="7" pageNumber="36">Smit</collectorName>
</materialsCitation>
, 1/0/0.
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9B95704ED9FC7468EEFB3A" pageId="7" pageNumber="36" type="discussion">
<paragraph id="8BC2F330FF9B95704ED9FC746FEDFB5E" blockId="7.[151,1437,834,1221]" pageId="7" pageNumber="36">
<emphasis id="B9092F22FF9B95704ED9FC746A65FC12" bold="true" box="[199,308,979,1005]" pageId="7" pageNumber="36">Remarks</emphasis>
—This species is most frequently recorded from many parts of Europe, but also from
<collectingCountry id="F36AB3A0FF9B95704AECFC746E64FC12" box="[1266,1333,979,1005]" name="China" pageId="7" pageNumber="36">China</collectingCountry>
(
<bibRefCitation id="EFEC8EC1FF9B95704B5DFC746B80FBEE" author="Li, H. - T. &amp; Ding, Z. - H. &amp; Gu, X. - Y. &amp; Guo, J. - J." pageId="7" pageNumber="36" pagination="410 - 441" refId="ref5583" refString="Li, H. - T., Ding, Z. - H., Gu, X. - Y. &amp; Guo, J. - J. (2022) Studies on Chinese water mites: The genus Feltria Koenike, 1892 (Acari: Feltriidae). Systematic and Applied Acarology, 27 (3), 410-441. https://doi.org/10.11158/saa.27.3.2" type="journal article" year="2022">
Li
<emphasis id="B9092F22FF9B95704B7AFC746EC4FC12" box="[1380,1429,979,1005]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
. 2022
</bibRefCitation>
) and Indian Himalayas (
<bibRefCitation id="EFEC8EC1FF9B95704FF4FC5069BBFBEE" author="Pesic, V. &amp; Panesar, A." box="[490,746,1015,1041]" pageId="7" pageNumber="36" pagination="1 - 28" refId="ref5661" refString="Pesic, V. &amp; Panesar, A. (2008) Studies on water mites (Acari, Hydrachnidia) from the Himalayas, I. The water mite genus Feltria Koenike, with descriptions of eight new species. Zootaxa, 1758 (1), 1-28. https://doi.org/10.11646/zootaxa.1758.1.1" type="journal article" year="2008">Pešić &amp; Panesar 2008</bibRefCitation>
). While males are clearly defined by the absence of a sexual dimorphism in the male III-L-
<quantity id="4C855ED5FF9B95704FF7FBBC6943FBCA" box="[489,530,1051,1077]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="36" unit="in" value="6.0">6 in</quantity>
combination with stout palps and a large dorsal shield excluding only Dgl-4+Dl- 4, the identification of the female can be difficult (
<bibRefCitation id="EFEC8EC1FF9B95704CC8FB98682AFBA6" author="Gerecke, R." box="[726,891,1087,1113]" pageId="7" pageNumber="36" pagination="13 - 47" refId="ref5337" refString="Gerecke, R. (2012) Studies on European Feltria species (Acari: Hydrachnidia: Feltriidae). Stuttgarter Beitrage zur Naturkunde A, Neue Serie, 5, 13-47." type="journal article" year="2012">Gerecke 2012</bibRefCitation>
). However, the presence of a tiny, rounded Dl-1 lying in the membrane facing the anterolateral edges of the dorsal shield, a character not found in any other species of the genus, allows easy identification of females of this species (
<bibRefCitation id="EFEC8EC1FF9B95704D63FB206F52FB5E" author="Bader, C." box="[893,1027,1159,1185]" pageId="7" pageNumber="36" pagination="1 - 127" refId="ref5272" refString="Bader, C. (1977) Die Wassermilben des Schweizerischen Nationalparks. Populationsdynamische Untersuchung zweier hochalpiner Quellen. Ergebnisse der wissenschaftlichen Untersuchungen im Schweizerischen Nationalpark, 15, 1-127." type="journal article" year="1977">Bader 1977</bibRefCitation>
;
<bibRefCitation id="EFEC8EC1FF9B95704A11FB206FE0FB5E" author="Gerecke, R." box="[1039,1201,1159,1185]" pageId="7" pageNumber="36" pagination="13 - 47" refId="ref5337" refString="Gerecke, R. (2012) Studies on European Feltria species (Acari: Hydrachnidia: Feltriidae). Stuttgarter Beitrage zur Naturkunde A, Neue Serie, 5, 13-47." type="journal article" year="2012">Gerecke 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8BC2F330FF9B95704ED9FB0C68EEFB3A" blockId="7.[151,1437,834,1221]" box="[199,959,1195,1221]" pageId="7" pageNumber="36">
<emphasis id="B9092F22FF9B95704ED9FB0C6A65FB3A" bold="true" box="[199,308,1195,1221]" pageId="7" pageNumber="36">Remarks</emphasis>
—Europe,
<collectingCountry id="F36AB3A0FF9B95704FB3FB0C6961FB3A" box="[429,560,1195,1221]" name="Turkey" pageId="7" pageNumber="36">Asia Minor</collectingCountry>
,
<collectingCountry id="F36AB3A0FF9B95704C25FB0C6926FB3A" box="[571,631,1195,1221]" name="India" pageId="7" pageNumber="36">India</collectingCountry>
,
<collectingCountry id="F36AB3A0FF9B95704C9CFB0C6998FB3A" box="[642,713,1195,1221]" name="China" pageId="7" pageNumber="36">China</collectingCountry>
. New for
<collectingCountry id="F36AB3A0FF9B95704D28FB0C68EBFB3A" box="[822,954,1195,1221]" name="Kyrgyzstan" pageId="7" pageNumber="36">Kyrgyzstan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,347 @@
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<mods:title id="0BAE6DCF4F43D77381ADCD05E668F57F">Two new species of the genus Feltria Koenike, 1892, with notes on the water mite fauna of Kyrgyzstan (Acari, Hydrachnidia)</mods:title>
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<mods:url id="F7D7C21F8E722F873FA5C4C3E8449661">https://doi.org/10.11646/zootaxa.5604.1.3</mods:url>
</mods:location>
<mods:classification id="AEC6FC72AB7ECDA28CCF135C3478704A">journal article</mods:classification>
<mods:identifier id="FDB72183EAC705E44B445B70DAAE8E95" type="DOI">10.11646/zootaxa.5604.1.3</mods:identifier>
<mods:identifier id="0B8EF290325ADF03A44117ADA7C25685" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C367A0BBFF9D95764E89FEC86FF3FE51" pageId="1" pageNumber="30" type="nomenclature">
<paragraph id="8BC2F330FF9D95764E89FEC86966FE76" blockId="1.[151,1186,367,465]" box="[151,567,367,394]" pageId="1" pageNumber="30">
<heading id="D08A445CFF9D95764E89FEC86966FE76" bold="true" box="[151,567,367,394]" fontSize="11" level="1" pageId="1" pageNumber="30" reason="1">
<emphasis id="B9092F22FF9D95764E89FEC86966FE76" bold="true" box="[151,567,367,394]" pageId="1" pageNumber="30">
<taxonomicName id="4C7D88B3FF9D95764E89FEC86A83FE75" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[151,466,367,394]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="kyrgyzica" status="sp. nov." subGenus="Feltria">
<emphasis id="B9092F22FF9D95764E89FEC86BBBFE75" bold="true" box="[151,234,367,394]" italics="true" pageId="1" pageNumber="30">Feltria</emphasis>
(
<emphasis id="B9092F22FF9D95764EE5FEC86A1FFE75" bold="true" box="[251,334,367,394]" italics="true" pageId="1" pageNumber="30">Feltria</emphasis>
)
<emphasis id="B9092F22FF9D95764F7EFEC86A83FE75" bold="true" box="[352,466,367,394]" italics="true" pageId="1" pageNumber="30">kyrgyzica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A9259FF9D95764FC7FEC86966FE76" box="[473,567,367,393]" pageId="1" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC2F330FF9D95764E89FE346FF3FE51" blockId="1.[151,1186,367,465]" box="[151,1186,403,430]" pageId="1" pageNumber="30">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C367A0BBFF9D95764E89FE106BAEFE2E" box="[151,255,439,465]" pageId="1" pageNumber="30" type="description">
<paragraph id="8BC2F330FF9D95764E89FE106BAEFE2E" blockId="1.[151,1186,367,465]" box="[151,255,439,465]" pageId="1" pageNumber="30">
<figureCitation id="1346EFB5FF9D95764E89FE106BAEFE2E" box="[151,255,439,465]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[152,255,1943,1967]" captionStartId-1="3.[152,255,1943,1967]" captionTargetBox-0="[164,1418,179,1917]" captionTargetBox-1="[172,1407,179,1917]" captionTargetId-0="figure-28@2.[164,1423,179,1917]" captionTargetId-1="figure-18@3.[172,1415,179,1917]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." captionText-1="FIGURE 2. Feltria kyrgyzica sp. nov. (A, ♂ from upper part of stream crossing the road to Son-Kul Lake; B—F, ♀ paratype, upper part of stream next to the road to May Saz pass): A, D—palp; B—Dorsum and excretory pore platelet; C—Venter (right genital plate lacking). Scale bars = 100 µm." figureDoi-0="http://doi.org/10.5281/zenodo.15012490" figureDoi-1="http://doi.org/10.5281/zenodo.15012494" httpUri-0="https://zenodo.org/record/15012490/files/figure.png" httpUri-1="https://zenodo.org/record/15012494/files/figure.png" pageId="1" pageNumber="30">Figs. 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9D95764E89FE586FDEFD32" pageId="1" pageNumber="30" type="materials_examined">
<paragraph id="8BC2F330FF9D95764E89FE586FA1FD7A" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
<emphasis id="B9092F22FF9D95764E89FE586A10FDE6" bold="true" box="[151,321,511,537]" pageId="1" pageNumber="30">Type material</emphasis>
<materialsCitation id="3B15F96DFF9D95764F43FE586BA1FD9E" collectingDate="2013-08-12" collectionCode="RMNH" collectorName="Pesic &amp; Smit." country="Kyrgyzstan" elevation="3346" latitude="42.41715" location="Karakol region" longLatPrecision="1" longitude="79.0443" municipality="May Saz" pageId="1" pageNumber="30" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C64D92FF9D95764F43FE586A94FDE6" box="[349,453,511,537]" pageId="1" pageNumber="30" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7B38B9FF9D95764FD1FDA76AB7FDE5" box="[463,486,512,538]" count="1" pageId="1" pageNumber="30" type="male"></specimenCount>
, dissected and slide mounted (
<collectionCode id="ED6C6BF5FF9D95764D55FE5868F2FDE6" box="[843,931,511,537]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="30" type="Museum">RMNH</collectionCode>
),
<collectingCountry id="F36AB3A0FF9D95764DA7FDA76F6DFDE5" box="[953,1084,512,538]" name="Kyrgyzstan" pageId="1" pageNumber="30">Kyrgyzstan</collectingCountry>
,
<location id="8EA2A5EBFF9D95764A54FE586FA9FDE6" LSID="urn:lsid:plazi:treatment:03D44226FF9D95734E89FEC86A78FDC2:8EA2A5EBFF9D95764A54FE586FA9FDE6" box="[1098,1272,511,537]" country="Kyrgyzstan" latitude="42.41715" longLatPrecision="1" longitude="79.0443" municipality="May Saz" name="Karakol region" pageId="1" pageNumber="30">Karakol region</location>
, upper part of stream next to the road to
<collectingMunicipality id="6BA6694AFF9D95764FA7FD83694AFDC2" box="[441,539,547,574]" pageId="1" pageNumber="30">May Saz</collectingMunicipality>
pass (
<figureCitation id="1346EFB5FF9D95764C7FFD8469E6FDC2" box="[609,695,547,574]" captionStart="FIGURE 5" captionStartId="9.[152,255,1885,1909]" captionTargetBox="[151,1436,1009,1859]" captionTargetId="figure-233@9.[151,1436,1009,1859]" captionTargetPageId="9" captionText="FIGURE 5. Photographs of sampling sites: A—Stream next to the road to May Saz pass, 2955 m asl., 12.viii.2013 (locus typicus of Feltria kyrgyzicus sp. nov.). B—River next to the road to Son-Kul Lake, 2426 m asl., 15.viii.2013 (locus typicus of Feltria tienshanensis sp. nov.). Photos by V. Pešić." figureDoi="http://doi.org/10.5281/zenodo.15012504" httpUri="https://zenodo.org/record/15012504/files/figure.png" pageId="1" pageNumber="30">Fig. 5A</figureCitation>
),
<geoCoordinate id="EE4995F7FF9D95764CD0FD846834FDC1" box="[718,869,547,574]" degrees="42" direction="north" minutes="25.029" orientation="latitude" pageId="1" pageNumber="30" precision="1" value="42.41715">42º25.029 N</geoCoordinate>
,
<geoCoordinate id="EE4995F7FF9D95764D71FD846F51FDC1" box="[879,1024,547,574]" degrees="79" direction="east" minutes="02.658" orientation="longitude" pageId="1" pageNumber="30" precision="1" value="79.0443">79º02.658 E</geoCoordinate>
,
<elevation id="00501403FF9D95764A10FD846FDCFDC2" box="[1038,1165,547,574]" metricMagnitude="3" metricUnit="m" metricValue="3.346" pageId="1" pageNumber="30" unit="m" value="3346.0">
<quantity id="4C855ED5FF9D95764A10FD846F33FDC1" box="[1038,1122,547,574]" metricMagnitude="3" metricUnit="m" metricValue="3.346" pageId="1" pageNumber="30" unit="m" value="3346.0">3346 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9D95764A82FD846E76FDC2" box="[1180,1319,547,573]" pageId="1" pageNumber="30" value="2013-08-12">
<collectingDate id="EF872C18FF9D95764A82FD846E76FDC2" box="[1180,1319,547,573]" pageId="1" pageNumber="30" value="2013-08-12">12.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9D95764B7FFD846ECDFDC2" box="[1377,1436,547,573]" pageId="1" pageNumber="30">Pešić</collectorName>
&amp;
<collectorName id="268896E6FF9D95764EADFDE06BA1FD9E" box="[179,240,583,609]" pageId="1" pageNumber="30">Smit.</collectorName>
</materialsCitation>
<materialsCitation id="3B15F96DFF9D95764EE8FDEF6FA1FD7A" collectingDate="2013-08-12" collectionCode="RMNH" collectorName="Pesic &amp; Smit." country="Kyrgyzstan" elevation="2955" latitude="42.4167" location="Karakol region" longLatPrecision="1" longitude="78.968285" municipality="May Saz" pageId="1" pageNumber="30" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54C64D92FF9D95764EE8FDEF6A36FD9D" box="[246,359,584,610]" pageId="1" pageNumber="30" type="paratype">Paratypes</typeStatus>
: 1/8/0 same data as the holotype,
<specimenCount id="9D7B38B9FF9D95764CC5FDE069AFFD9E" box="[731,766,583,609]" count="1" pageId="1" pageNumber="30" type="female">1♀</specimenCount>
dissected and slide mounted (
<collectionCode id="ED6C6BF5FF9D95764A53FDE06FF7FD9E" box="[1101,1190,583,609]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="30" type="Museum">RMNH</collectionCode>
); 1/1/0, stream next to the road to May Saz pass,
<geoCoordinate id="EE4995F7FF9D95764FA0FDCC6904FD79" box="[446,597,619,646]" degrees="42" direction="north" minutes="25.002" orientation="latitude" pageId="1" pageNumber="30" precision="1" value="42.4167">42º25.002 N</geoCoordinate>
,
<geoCoordinate id="EE4995F7FF9D95764C7EFDCC69A0FD79" box="[608,753,619,646]" degrees="78" direction="east" minutes="58.097" orientation="longitude" pageId="1" pageNumber="30" precision="1" value="78.968285">78º58.097 E</geoCoordinate>
,
<elevation id="00501403FF9D95764CE1FDCC682EFD7A" box="[767,895,619,646]" metricMagnitude="3" metricUnit="m" metricValue="2.955" pageId="1" pageNumber="30" unit="m" value="2955.0">
<quantity id="4C855ED5FF9D95764CE1FDCC6805FD79" box="[767,852,619,646]" metricMagnitude="3" metricUnit="m" metricValue="2.955" pageId="1" pageNumber="30" unit="m" value="2955.0">2955 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9D95764D91FDCC6F48FD7A" box="[911,1049,619,645]" pageId="1" pageNumber="30" value="2013-08-12">
<collectingDate id="EF872C18FF9D95764D91FDCC6F48FD7A" box="[911,1049,619,645]" pageId="1" pageNumber="30" value="2013-08-12">12.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9D95764A4BFDCC6FC1FD7A" box="[1109,1168,619,645]" pageId="1" pageNumber="30">Pešić</collectorName>
&amp;
<collectorName id="268896E6FF9D95764AADFDCC6FA1FD7A" box="[1203,1264,619,645]" pageId="1" pageNumber="30">Smit.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BC2F330FF9D95764ED9FD286FDEFD32" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
<emphasis id="B9092F22FF9D95764ED9FD286A29FD56" bold="true" box="[199,376,655,681]" pageId="1" pageNumber="30">Other material</emphasis>
<materialsCitation id="3B15F96DFF9D95764F88FD376FDAFD32" collectingDate="2013-08-17" collectionCode="RMNH" collectorName="Pesic &amp; Smit" country="Kyrgyzstan" elevation="3412" latitude="41.91365" location="Son-Kul Lake" longLatPrecision="1" longitude="75.42828" pageId="1" pageNumber="30" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F36AB3A0FF9D95764F88FD376948FD55" box="[406,537,656,682]" name="Kyrgyzstan" pageId="1" pageNumber="30">Kyrgyzstan</collectingCountry>
: upper part of stream crossing the road to
<location id="8EA2A5EBFF9D95764DC4FD286F28FD56" LSID="urn:lsid:plazi:treatment:03D44226FF9D95734E89FEC86A78FDC2:8EA2A5EBFF9D95764DC4FD286F28FD56" box="[986,1145,655,681]" country="Kyrgyzstan" latitude="41.91365" longLatPrecision="1" longitude="75.42828" name="Son-Kul Lake" pageId="1" pageNumber="30">Son-Kul Lake</location>
,
<geoCoordinate id="EE4995F7FF9D95764A9FFD286E4BFD55" box="[1153,1306,655,682]" degrees="41" direction="north" minutes="54.819" orientation="latitude" pageId="1" pageNumber="30" precision="1" value="41.91365">41º 54.819 N</geoCoordinate>
,
<geoCoordinate id="EE4995F7FF9D95764B3DFD286BF9FD31" degrees="75" direction="east" minutes="25.697" orientation="longitude" pageId="1" pageNumber="30" precision="1" value="75.42828">75º25.697 E</geoCoordinate>
,
<elevation id="00501403FF9D95764EA8FD146A67FD32" box="[182,310,691,718]" metricMagnitude="3" metricUnit="m" metricValue="3.412" pageId="1" pageNumber="30" unit="m" value="3412.0">
<quantity id="4C855ED5FF9D95764EA8FD146A5AFD31" box="[182,267,691,718]" metricMagnitude="3" metricUnit="m" metricValue="3.412" pageId="1" pageNumber="30" unit="m" value="3412.0">3412 m</quantity>
asl
</elevation>
.,
<date id="FFC3D5F0FF9D95764F58FD146A81FD32" box="[326,464,691,717]" pageId="1" pageNumber="30" value="2013-08-17">
<collectingDate id="EF872C18FF9D95764F58FD146A81FD32" box="[326,464,691,717]" pageId="1" pageNumber="30" value="2013-08-17">17.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="268896E6FF9D95764C12FD146916FD32" box="[524,583,691,717]" pageId="1" pageNumber="30">Pešić</collectorName>
&amp;
<collectorName id="268896E6FF9D95764C75FD1469F2FD32" box="[619,675,691,717]" pageId="1" pageNumber="30">Smit</collectorName>
,
<specimenCount id="9D7B38B9FF9D95764CB0FD146983FD32" box="[686,722,691,717]" count="1" pageId="1" pageNumber="30" type="male">1♂</specimenCount>
, dissected and slide mounted (
<collectionCode id="ED6C6BF5FF9D95764A34FD146FD2FD32" box="[1066,1155,691,717]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="30" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9D95764ED9FD706F5DFC7E" pageId="1" pageNumber="30" type="diagnosis">
<paragraph id="8BC2F330FF9D95764ED9FD706F5DFC7E" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
<emphasis id="B9092F22FF9D95764ED9FD706A6BFD0E" bold="true" box="[199,314,727,753]" pageId="1" pageNumber="30">Diagnosis</emphasis>
—Both sexes: Setae associated with Dgl-1 fine, Dgl-2 accompanied by strongly thickened setae. Coxal plates in four groups. Genital field with 70 to 90 pairs of Ac.
<emphasis id="B9092F22FF9D95764D8AFD5C689FFCEA" box="[916,974,763,789]" italics="true" pageId="1" pageNumber="30">Male</emphasis>
: Dorsum with large anterior shield and a pair of transverse posterior plates. Genital plate transverse. Palp robust and stout. III-L-6 with a long and pointed ventrolateral projection in the distal third.
<emphasis id="B9092F22FF9D95764C97FCE4698CFCA2" box="[649,733,835,861]" italics="true" pageId="1" pageNumber="30">Female</emphasis>
: Dorsum with rhomboid shield surrounded by four rounded platelets. Excretory pore in posterodorsal position. Palp slenderer than in male.
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9D95764ED9FC2C68A0F846" pageId="1" pageNumber="30" type="description">
<paragraph id="8BC2F330FF9D95764ED9FC2C69BFFB5E" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
<emphasis id="B9092F22FF9D95764ED9FC2C6A00FC5A" bold="true" box="[199,337,907,933]" pageId="1" pageNumber="30">Description</emphasis>
<emphasis id="B9092F22FF9D95764F70FC2C6AF9FC5A" box="[366,424,907,933]" italics="true" pageId="1" pageNumber="30">Male</emphasis>
(
<typeStatus id="54C64D92FF9D95764FA6FC2C694FFC5A" box="[440,542,907,933]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
, juvenile specimen; in parentheses some measurements of a mature specimen from the upper part of the stream next to the road to Son-Kul Lake): Idiosoma
<quantity id="4C855ED5FF9D95764A17FC086F01FC36" box="[1033,1104,943,970]" metricMagnitude="1" metricUnit="m" metricValue="1.143" pageId="1" pageNumber="30" unit="in" value="450.0">450 in</quantity>
length (
<typeStatus id="54C64D92FF9D95764AB5FC086E5CFC36" box="[1195,1293,943,969]" pageId="1" pageNumber="30" type="paratype">paratype</typeStatus>
from stream next to the road to May Saz pass
<quantity id="4C855ED5FF9D95764C13FC746905FC12" box="[525,596,979,1005]" metricMagnitude="0" metricUnit="m" metricValue="9.9314" pageId="1" pageNumber="30" unit="in" value="391.0">391 in</quantity>
length and
<quantity id="4C855ED5FF9D95764CC6FC746871FC12" box="[728,800,979,1005]" metricMagnitude="0" metricUnit="m" metricValue="7.3152" pageId="1" pageNumber="30" unit="in" value="288.0">288 in</quantity>
width). Dorsum with a large dorsal shield,
<quantity id="4C855ED5FF9D95764B16FC746E1EFC12" box="[1288,1359,979,1005]" metricMagnitude="0" metricUnit="m" metricValue="8.8138" pageId="1" pageNumber="30" unit="in" value="347.0">347 in</quantity>
length and
<quantity id="4C855ED5FF9D95764ED9FC506A5EFBEE" box="[199,271,1015,1042]" metricMagnitude="0" metricUnit="m" metricValue="6.4008" pageId="1" pageNumber="30" unit="in" value="252.0">252 in</quantity>
width, generally including Dl-1-3 and Dgl-1-3 (
<figureCitation id="1346EFB5FF9D95764D2EFC5068D6FBEE" box="[816,903,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="1" pageNumber="30">Fig. 1C</figureCitation>
), but occasionally (in
<typeStatus id="54C64D92FF9D95764A9DFC506FB9FBEE" box="[1155,1256,1015,1041]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
,
<figureCitation id="1346EFB5FF9D95764AEAFC506E1BFBEE" box="[1268,1354,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="1" pageNumber="30">Fig. 1A</figureCitation>
) Dgl-1 on a separate paired plate; Dgl-4+Dl-4 on a pair of large, transverse platelets. Frontal area with pre-antenniform glandularia, pre-ocular setae and Lgl-1 fused to each other. Coxal plates in four groups, close to each other, occupying more than half of the idiosoma surface (
<figureCitation id="1346EFB5FF9D95764C7FFBC469E8FB82" box="[609,697,1123,1149]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="1" pageNumber="30">Fig. 1B</figureCitation>
); anterior coxae with well-developed posterior apodemes; Cx-IV posterior margin perpendicularly to longitudinal axis.
</paragraph>
<paragraph id="8BC2F330FF9D95764ED9FB0C697BFAAA" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
Genital plate transverse, its anterior margin a straight line, separated from Cx-IV by a narrow strip of membrane bearing the Vgl and a pair of triangular lateral platelets; 118 (128) in length and 272 (300) in width, with 7280 pairs acetabula; gonopore short, slit-shaped, in the center (
<figureCitation id="1346EFB5FF9D95764CF4FB546821FAF2" box="[746,880,1267,1293]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="1" pageNumber="30">Figs. 1B, D</figureCitation>
); excretory pore on a broad platelet, laying in the posterior indentation of the genital plate margin (
<figureCitation id="1346EFB5FF9D95764CDCFAB06848FACE" box="[706,793,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="1" pageNumber="30">Fig. 1B</figureCitation>
), in a mature specimen this platelet fused to genital field (
<figureCitation id="1346EFB5FF9D95764E81FA9C6BA6FAAA" box="[159,247,1339,1365]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="1" pageNumber="30">Fig. 1D</figureCitation>
), directed posteroventrally.
</paragraph>
<paragraph id="8BC2F330FF9D95764ED9FAF86910FA1A" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
Palp robust and stout, P-2 ventral margin convex, P-4 with a weakly-developed ventral groove flanked by two setae (
<figureCitation id="1346EFB5FF9D95764F0FFA246A36FA62" box="[273,359,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="1" pageNumber="30">Fig. 1E</figureCitation>
,
<figureCitation id="1346EFB5FF9D95764F6AFA246ACBFA62" box="[372,410,1411,1437]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[172,1407,179,1917]" captionTargetId="figure-18@3.[172,1415,179,1917]" captionTargetPageId="3" captionText="FIGURE 2. Feltria kyrgyzica sp. nov. (A, ♂ from upper part of stream crossing the road to Son-Kul Lake; B—F, ♀ paratype, upper part of stream next to the road to May Saz pass): A, D—palp; B—Dorsum and excretory pore platelet; C—Venter (right genital plate lacking). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012494" httpUri="https://zenodo.org/record/15012494/files/figure.png" pageId="1" pageNumber="30">2A</figureCitation>
); dorsal length/height of palpal segments: P-1, 23/- (23/31); P-2, 72/67 (74/65); P-3, 39/45 (42/47); P-4, 90/38 (89/41); P-5, 53/24 (51/25); length P-2/P-4 ratio 0.8 (0.83). Gnathosoma ventral length 64 (91 with apodemes); chelicera length 116.
</paragraph>
<paragraph id="8BC2F330FF9D95764ED9FA486930F9AE" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
III-L-6 with a long and pointed ventrolateral projection in the distal third, bearing 45 long setae (
<figureCitation id="1346EFB5FF9D95764B29FA486EDFF9F6" box="[1335,1422,1519,1545]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="1" pageNumber="30">Fig. 1F</figureCitation>
); dorsal lengths of III-L-2-6: 52 (52), 64 (66), 92 (91), 106 (100), 114 (121); dorsal lengths of IV-L-1-6: 84, 52 (56), 72 (82), 102 (116), 118 (123), 109 (131).
</paragraph>
<paragraph id="8BC2F330FF9D95764ED9F9FC6E62F8D6" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
<emphasis id="B9092F22FF9D95764ED9F9FC6A4AF98A" box="[199,283,1627,1653]" italics="true" pageId="1" pageNumber="30">Female</emphasis>
(from upper part of stream next to the road to May Saz pass; in parentheses some measurements of conserved specimens): Idiosoma length 490 (497512, n=3) in length and (359400, n=3) in width, length/width ratio 1.281.38 (n=3); dorsal shield, including Dgl-
<quantity id="4C855ED5FF9D95764CCEF904687FF942" box="[720,814,1699,1725]" metricMagnitude="1" metricUnit="m" metricValue="5.93852" pageId="1" pageNumber="30" unit="in" value="2338.0">2, 338 in</quantity>
length and
<quantity id="4C855ED5FF9D95764DB3F90468A3F942" box="[941,1010,1699,1726]" metricMagnitude="0" metricUnit="m" metricValue="6.5024" pageId="1" pageNumber="30" unit="in" value="256.0">256 in</quantity>
width; Dgl-1 on platelets anterolateral to dorsal shield. Dl-2 elongated, Dl-3 and Dgl-3 separate from each other, Dgl-4+Dl-4 on a pair of large, transverse platelets (
<figureCitation id="1346EFB5FF9D95764F1AF94C6A0DF8FA" box="[260,348,1771,1797]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[172,1407,179,1917]" captionTargetId="figure-18@3.[172,1415,179,1917]" captionTargetPageId="3" captionText="FIGURE 2. Feltria kyrgyzica sp. nov. (A, ♂ from upper part of stream crossing the road to Son-Kul Lake; B—F, ♀ paratype, upper part of stream next to the road to May Saz pass): A, D—palp; B—Dorsum and excretory pore platelet; C—Venter (right genital plate lacking). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012494" httpUri="https://zenodo.org/record/15012494/files/figure.png" pageId="1" pageNumber="30">Fig. 2B</figureCitation>
); excretory pore platelet broad, directed posteroventrally. Coxal field
<quantity id="4C855ED5FF9D95764A79F94C6FFFF8FA" box="[1127,1198,1771,1797]" metricMagnitude="0" metricUnit="m" metricValue="7.6708" pageId="1" pageNumber="30" unit="in" value="302.0">302 in</quantity>
length, Cx-III
<quantity id="4C855ED5FF9D95764B48F94C6ECDF8FA" box="[1366,1436,1771,1797]" metricMagnitude="0" metricUnit="m" metricValue="8.026399999999999" pageId="1" pageNumber="30" unit="in" value="316.0">316 in</quantity>
width. Genital plates subtriangular,
<quantity id="4C855ED5FF9D95764C39F8A869F7F8D6" box="[551,678,1807,1834]" metricMagnitude="0" metricUnit="m" metricValue="3.1369" metricValueMax="3.175" metricValueMin="3.0987999999999998" pageId="1" pageNumber="30" unit="in" value="123.5" valueMax="125.0" valueMin="122.0">122125 in</quantity>
length and
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width, with about 90 pairs of acetabula.
</paragraph>
<paragraph id="8BC2F330FF9D95764ED9F89468A0F846" blockId="1.[151,1437,511,1978]" pageId="1" pageNumber="30">
Chaetotaxy of palp as given in
<figureCitation id="1346EFB5FF9D95764C32F89469D7F8B2" box="[556,646,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[172,1407,179,1917]" captionTargetId="figure-18@3.[172,1415,179,1917]" captionTargetPageId="3" captionText="FIGURE 2. Feltria kyrgyzica sp. nov. (A, ♂ from upper part of stream crossing the road to Son-Kul Lake; B—F, ♀ paratype, upper part of stream next to the road to May Saz pass): A, D—palp; B—Dorsum and excretory pore platelet; C—Venter (right genital plate lacking). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012494" httpUri="https://zenodo.org/record/15012494/files/figure.png" pageId="1" pageNumber="30">Fig. 2D</figureCitation>
; dorsal length/height of palpal segments: P-1, 23/28; P-2, 64/58; P-3, 39/39; P-4, 88/38; P-5, 55/20; length P-2/P-4 ratio 0.73. Gnathosoma ventral length 103; chelicera
<quantity id="4C855ED5FF9D95764AE1F8F06E16F88E" box="[1279,1351,1879,1905]" metricMagnitude="0" metricUnit="m" metricValue="3.0987999999999998" pageId="1" pageNumber="30" unit="in" value="122.0">122 in</quantity>
length. III-L-6 without ventrolateral projection. Dorsal lengths of I-L-2-6: 46, 52, 67, 80, 90; I-L-6 height 32, I-L-6 dorsal length/height ratio 2.8; dorsal lengths of IV-L-1-6: 86, 50, 78, 116, 125, 131.
</paragraph>
</subSubSection>
<caption id="DF02A3B8FF9E95754E86F830688DF807" ID-DOI="http://doi.org/10.5281/zenodo.15012490" ID-Zenodo-Dep="15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="2" pageNumber="31" startId="2.[152,255,1943,1967]" targetBox="[164,1418,179,1917]" targetPageId="2" targetType="figure">
<paragraph id="8BC2F330FF9E95754E86F830688DF807" blockId="2.[151,1436,1943,2040]" pageId="2" pageNumber="31">
<emphasis id="B9092F22FF9E95754E86F8306A42F850" bold="true" box="[152,275,1943,1967]" pageId="2" pageNumber="31">FIGURE 1</emphasis>
.
<taxonomicName id="4C7D88B3FF9E95754F3FF83F6A9FF84F" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[289,462,1944,1968]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="kyrgyzica" status="sp. nov.">
<emphasis id="B9092F22FF9E95754F3FF83F6A9FF84F" box="[289,462,1944,1968]" italics="true" pageId="2" pageNumber="31">Feltria kyrgyzica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A9259FF9E95754FC8F83F6978F850" box="[470,553,1944,1968]" pageId="2" pageNumber="31" rank="species">
<emphasis id="B9092F22FF9E95754FC8F83F6972F84F" bold="true" box="[470,547,1944,1968]" pageId="2" pageNumber="31">sp. nov</emphasis>
.
</taxonomicNameLabel>
, ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm.
</paragraph>
</caption>
<caption id="DF02A3B8FF9F95744E86F8306903F807" ID-DOI="http://doi.org/10.5281/zenodo.15012494" ID-Zenodo-Dep="15012494" httpUri="https://zenodo.org/record/15012494/files/figure.png" pageId="3" pageNumber="32" startId="3.[152,255,1943,1967]" targetBox="[172,1407,179,1917]" targetPageId="3" targetType="figure">
<paragraph id="8BC2F330FF9F95744E86F8306903F807" blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="32">
<emphasis id="B9092F22FF9F95744E86F8306A42F850" bold="true" box="[152,275,1943,1967]" pageId="3" pageNumber="32">FIGURE 2</emphasis>
.
<taxonomicName id="4C7D88B3FF9F95744F01F83F6A9DF84F" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[287,460,1944,1968]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="kyrgyzica" status="sp. nov.">
<emphasis id="B9092F22FF9F95744F01F83F6A9DF84F" box="[287,460,1944,1968]" italics="true" pageId="3" pageNumber="32">Feltria kyrgyzica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A9259FF9F95744FCCF83F6974F850" box="[466,549,1944,1968]" pageId="3" pageNumber="32" rank="species">
<emphasis id="B9092F22FF9F95744FCCF83F694FF84F" bold="true" box="[466,542,1944,1968]" pageId="3" pageNumber="32">sp. nov</emphasis>
.
</taxonomicNameLabel>
(A, ♂ from upper part of stream crossing the road to Son-Kul Lake; B—F, ♀ paratype, upper part of stream next to the road to May Saz pass): A, D—palp; B—Dorsum and excretory pore platelet; C—Venter (right genital plate lacking). Scale bars = 100 µm.
</paragraph>
</caption>
<subSubSection id="C367A0BBFF9895734ED9FF306A98FF2A" pageId="4" pageNumber="33" type="etymology">
<paragraph id="8BC2F330FF9895734ED9FF306A98FF2A" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="33">
<emphasis id="B9092F22FF9895734ED9FF306A19FF4E" bold="true" box="[199,328,151,177]" pageId="4" pageNumber="33">Etymology</emphasis>
—The new species is named after the country where it was collected. The name
<taxonomicName id="4C7D88B3FF9895734AECFF306E0DFF4E" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[1266,1372,151,177]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="kyrgyzica">
<emphasis id="B9092F22FF9895734AECFF306E0DFF4E" box="[1266,1372,151,177]" italics="true" pageId="4" pageNumber="33">kyrgyzica</emphasis>
</taxonomicName>
is an adjective, gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9895734ED9FF796F68FE0A" pageId="4" pageNumber="33" type="discussion">
<paragraph id="8BC2F330FF9895734ED9FF796F68FE0A" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="33">
<emphasis id="B9092F22FF9895734ED9FF796A14FF07" bold="true" box="[199,325,222,248]" pageId="4" pageNumber="33">Discussion</emphasis>
<taxonomicName id="4C7D88B3FF9895734F7FFF786970FF06" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[353,545,223,249]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="kyrgyzica" status="sp. nov.">
<emphasis id="B9092F22FF9895734F7FFF786970FF06" box="[353,545,223,249]" italics="true" pageId="4" pageNumber="33">Feltria kyrgyzica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23A9259FF9895734C37FF7869D5FF06" box="[553,644,223,249]" pageId="4" pageNumber="33" rank="species">
<emphasis id="B9092F22FF9895734C37FF78692FFF06" bold="true" box="[553,638,223,249]" pageId="4" pageNumber="33">sp. nov</emphasis>
.
</taxonomicNameLabel>
belongs to
<taxonomicName id="4C7D88B3FF9895734D16FF786822FF07" authorityName="Koenike" authorityYear="1892" box="[776,883,223,248]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="B9092F22FF9895734D16FF786822FF07" box="[776,883,223,248]" italics="true" pageId="4" pageNumber="33">F. minuta</emphasis>
</taxonomicName>
-complex, a group rather dishomogenous in male morphology (primarily defined by genital plate with straight anterior margin and gonopore in a central position) but in females well defined by the combination of postantenniformia included into, but Dgl-1 and Dl-2 and Dl-3 separate from, dorsal shield. The new species from
<collectingCountry id="F36AB3A0FF9895734CFCFEEC6832FE9A" box="[738,867,331,357]" name="Kyrgyzstan" pageId="4" pageNumber="33">Kyrgyzstan</collectingCountry>
is most similar to
<taxonomicName id="4C7D88B3FF9895734A27FEEC6B83FE76" authority="Pesic &amp; Panesar, 2008" authorityName="Pesic &amp; Panesar" authorityYear="2008" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B9092F22FF9895734A27FEEC6F84FE9A" box="[1081,1237,331,357]" italics="true" pageId="4" pageNumber="33">Feltria indica</emphasis>
<bibRefCitation id="EFEC8EC1FF9895734AC0FEEC6B83FE76" author="Pesic, V. &amp; Panesar, A." pageId="4" pageNumber="33" pagination="1 - 28" refId="ref5661" refString="Pesic, V. &amp; Panesar, A. (2008) Studies on water mites (Acari, Hydrachnidia) from the Himalayas, I. The water mite genus Feltria Koenike, with descriptions of eight new species. Zootaxa, 1758 (1), 1-28. https://doi.org/10.11646/zootaxa.1758.1.1" type="journal article" year="2008">Pešić &amp; Panesar, 2008</bibRefCitation>
</taxonomicName>
, known from the Indian Himalayas (
<bibRefCitation id="EFEC8EC1FF9895734C66FEC8682BFE76" author="Pesic, V. &amp; Panesar, A." box="[632,890,367,393]" pageId="4" pageNumber="33" pagination="1 - 28" refId="ref5661" refString="Pesic, V. &amp; Panesar, A. (2008) Studies on water mites (Acari, Hydrachnidia) from the Himalayas, I. The water mite genus Feltria Koenike, with descriptions of eight new species. Zootaxa, 1758 (1), 1-28. https://doi.org/10.11646/zootaxa.1758.1.1" type="journal article" year="2008">Pešić &amp; Panesar 2008</bibRefCitation>
). The male of both species with III-L-6 bearing a long and pointed ventrolateral projection in the distal third and with a similar pattern of dorsal sclerites in both sexes. The latter species can clearly be separated by having thickened setae associated with glandularia Dgl-1 (fine in
<emphasis id="B9092F22FF9895734EAAFE7C6A6EFE0A" box="[180,319,475,501]" italics="true" pageId="4" pageNumber="33">
K.
<taxonomicName id="4C7D88B3FF9895734ECBFE7C6A6EFE0A" authorityName="Pešić &amp; Smit" authorityYear="2025" box="[213,319,475,501]" class="Arachnida" family="Feltriidae" genus="Feltria" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="kyrgyzica" status="sp. nov.">kyrgyzica</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A23A9259FF9895734F58FE7C6ACEFE0A" box="[326,415,475,501]" pageId="4" pageNumber="33" rank="species">
<emphasis id="B9092F22FF9895734F58FE7C6ACBFE0A" bold="true" box="[326,410,475,501]" pageId="4" pageNumber="33">sp. nov</emphasis>
.
</taxonomicNameLabel>
) in both sexes (see
<figureCitation id="1346EFB5FF9895734C61FE7C699AFE0A" box="[639,715,475,501]" captionStart="FIGURE 1" captionStartId="2.[152,255,1943,1967]" captionTargetBox="[164,1418,179,1917]" captionTargetId="figure-28@2.[164,1423,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Feltria kyrgyzica sp. nov., ♂ (AB, EF holotype, upper part of stream next to the road to May Saz pass; CD, specimen from upper part of stream crossing the road to Son-Kul Lake): A—Dorsum; A—Venter; C—Frontal area of dorsum; D—Genital field; E—Palp (segments 25); F—I-L-5 and -6. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.15012490" httpUri="https://zenodo.org/record/15012490/files/figure.png" pageId="4" pageNumber="33">Figs. 1</figureCitation>
and
<quantity id="4C855ED5FF9895734D1FFE7C687DFE0A" box="[769,812,475,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="33" unit="in" value="7.0">7 in</quantity>
<bibRefCitation id="EFEC8EC1FF9895734D2DFE7C6F7CFE0A" author="Pesic, V. &amp; Panesar, A." box="[819,1069,475,501]" pageId="4" pageNumber="33" pagination="1 - 28" refId="ref5661" refString="Pesic, V. &amp; Panesar, A. (2008) Studies on water mites (Acari, Hydrachnidia) from the Himalayas, I. The water mite genus Feltria Koenike, with descriptions of eight new species. Zootaxa, 1758 (1), 1-28. https://doi.org/10.11646/zootaxa.1758.1.1" type="journal article" year="2008">Pešić &amp; Panesar 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C367A0BBFF9895734ED9FE596A78FDC2" pageId="4" pageNumber="33" type="distribution">
<paragraph id="8BC2F330FF9895734ED9FE596A78FDC2" blockId="4.[151,1436,151,573]" pageId="4" pageNumber="33">
<emphasis id="B9092F22FF9895734ED9FE596A08FDE7" bold="true" box="[199,345,510,536]" pageId="4" pageNumber="33">Distribution</emphasis>
<collectingCountry id="F36AB3A0FF9895734F69FE586AABFDE6" box="[375,506,511,537]" name="Kyrgyzstan" pageId="4" pageNumber="33">Kyrgyzstan</collectingCountry>
; known from streams in the Tien Shan Mountains at an elevation of about 3000 3400 meters.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,276 @@
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<mods:title id="47FF0587BC6A806D2027887A31DAE6CF">Revision of the Austro-Oriental shield-bug genus Axiagastus (Hemiptera: Heteroptera: Pentatomidae), with the description of five new species, and taxonomic notes on related genera</mods:title>
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<mods:namePart id="67E5183D646D5C17B0B996430FAB4DBA">Kment, Petr</mods:namePart>
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<mods:affiliation id="F4B3114DE2B65B6D1DA413E2EAA24221">Department of Entomology, National Museum of the Czech Republic, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
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<mods:namePart id="EEFC6C1D70CF813E0753CCF0EA46F806">Lemaître, Valérie A.</mods:namePart>
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<mods:affiliation id="E8AD73227D760E631CD497841B4E53FA">Department of Collections (Insects), The Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom; &amp; val. heureuse @ gmail. com; https: // orcid. org / 0000 - 0003 - 4802 - 2711</mods:affiliation>
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<mods:affiliation id="473E28C10B16E6207E50D23F93CC927A">Department of Collections (Insects), The Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom; &amp; mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
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<mods:namePart id="E1C43187655F8A0AFDFC64A3D6A9A934">Roca-Cusachs, Marcos</mods:namePart>
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<mods:affiliation id="85BD114DE21FAEC40C54134DC848E70C">ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, Karnataka, India shalinis. nilavu @ gmail. com; https: // orcid. org / 0000 - 0003 - 3557 - 4365 &amp; Evolution &amp; Ecology Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, Australia &amp; IRBio, Institut de Recerca de la Biodiversitat, Universitat de Barcelona, Barcelona, Spain marcosrocacusachs @ gmail. com; https: // orcid. org / 0000 - 0002 - 9174 - 6021 # Former affiliation * Corresponding author &amp; ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, Karnataka, India shalinis. nilavu @ gmail. com; https: // orcid. org / 0000 - 0003 - 3557 - 4365</mods:affiliation>
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<emphasis id="AACB9C202733FFB0FF5AF8A848A300FA" bold="true" box="[151,507,905,933]" pageId="73" pageNumber="74">
<emphasis id="AACB9C202733FFB0FF5AF8A8482000FB" bold="true" box="[151,376,906,933]" italics="true" pageId="73" pageNumber="74">Acesines breviceps</emphasis>
<bibRefCitation id="FC2E3DC32733FFB0FE4DF8AA48A300FA" author="Stal, C." box="[384,507,905,932]" pageId="73" pageNumber="74" pagination="1 - 162" refId="ref55674" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. [Checklist of Hemiptera. Contribution to a list of all hitherto known Hemiptera, along with systematic notes. 5]. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1-162. [in Latin]" type="journal article" year="1876">Stål, 1876</bibRefCitation>
</emphasis>
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</paragraph>
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<emphasis id="AACB9C202733FFB0FF5AF8EC480F00B9" box="[151,343,975,999]" italics="true" pageId="73" pageNumber="74">Acesines breviceps</emphasis>
<materialsCitation id="28D74A6F2733FFB0FE93F8EC4B4A00B9" box="[350,530,975,999]" collectionCode="NHRS" country="India" county="Typus" location="Boucard" municipality="Type" pageId="73" pageNumber="74" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<bibRefCitation id="FC2E3DC32733FFB0FE93F8EC4B4A00B9" author="Stal, C." box="[350,530,975,999]" pageId="73" pageNumber="74" pagination="1 - 162" refId="ref55674" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. [Checklist of Hemiptera. Contribution to a list of all hitherto known Hemiptera, along with systematic notes. 5]. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1-162. [in Latin]" type="journal article" year="1876">Stål, 1876: 9495</bibRefCitation>
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(original description).
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:
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, “
<collectingCountry id="E0A800A22733FFB0FC59F8EC4A9200B9" box="[916,970,975,999]" name="India" pageId="73" pageNumber="74">India</collectingCountry>
Orientalis”,
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,
<collectingMunicipality id="7864DA482733FFB0FB7CF8F34DBE00B6" box="[1201,1254,976,1000]" pageId="73" pageNumber="74">Type</collectingMunicipality>
,
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[red label, outlined black], NHRS-GULI000067472 (NHRS).
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</paragraph>
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<treatmentCitation id="191E66232733FFB0FF5AFF2E4B6F077B" author="Distant, W. L." box="[151,567,1036,1061]" page="144" pageId="73" pageNumber="74" year="1918">
<taxonomicName id="5FBF3BB12733FFB0FF5AFF2E4B6F077B" authority="Distant, 1918: 144" authorityName="Distant" authorityPageNumber="144" authorityYear="1918" box="[151,567,1036,1061]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bambusana">
<emphasis id="AACB9C202733FFB0FF5AFF2E4835077A" box="[151,365,1036,1060]" italics="true" pageId="73" pageNumber="74">Acesines bambusana</emphasis>
<bibRefCitation id="FC2E3DC32733FFB0FEBBFF2F4B6F077B" author="Distant, W. L." box="[374,567,1036,1061]" pageId="73" pageNumber="74" pagination="1 - 210" refId="ref49585" refString="Distant, W. L. (1918) Rhynchota. Vol. VII. Homoptera: Appendix. Heteroptera: Addenda. In: Shipley, A. E. &amp; Marshall, A. K. (Eds.), The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, pp. i - vii + 1-210." type="book chapter" year="1918">Distant, 1918: 144</bibRefCitation>
</taxonomicName>
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(original description).
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<typeStatus id="4704FE902733FFB0FCEAFF2E4AD0077B" box="[807,904,1037,1061]" pageId="73" pageNumber="74" type="syntype">Syntypes</typeStatus>
:
<collectingCountry id="E0A800A22733FFB0FC59FF2F4A94077A" box="[916,972,1036,1060]" name="India" pageId="73" pageNumber="74">India</collectingCountry>
:
<collectingRegion id="5A7B8ED02733FFB0FC1AFF2F4D1B077A" box="[983,1091,1036,1060]" country="India" name="Karnataka" pageId="73" pageNumber="74">Karnataka</collectingRegion>
: Chikkaballapura (
<collectorName id="354A25E42733FFB0FACAFF2E4CC5077A" box="[1287,1437,1036,1061]" pageId="73" pageNumber="74">T.V. Campbell</collectorName>
coll.,
<collectionCode id="FEAED8F72733FFB0FECDFF084808071D" box="[256,336,1067,1091]" pageId="73" pageNumber="74">BMNH</collectionCode>
), on bamboo.
<emphasis id="AACB9C202733FFB0FE12FF084A6D071D" bold="true" box="[479,821,1067,1091]" pageId="73" pageNumber="74">New junior subjective synonym</emphasis>
</materialsCitation>
<emphasis id="AACB9C202733FFB0FCF8FF084A61071D" bold="true" box="[821,825,1067,1091]" pageId="73" pageNumber="74">.</emphasis>
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</paragraph>
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<emphasis id="AACB9C202733FFB0FF5AFF5248D807D5" bold="true" box="[151,384,1137,1163]" pageId="73" pageNumber="74">Material examined.</emphasis>
<materialsCitation id="28D74A6F2733FFB0FE4AFF524DD607F1" collectingDate="2002-08-14" collectingDateMax="2002-08-22" collectingDateMin="2002-08-14" collectionCode="NMPC" collectorName="P. Sipek &amp; M. Fikacek" country="India" determinerName="P. Kment" elevation="10" latitude="15.0078335" location="Palolem" longLatPrecision="13" longitude="74.02634" municipality="Goa" pageId="73" pageNumber="74" specimenCount="1" specimenCount-female="1" stateProvince="Goa">
<emphasis id="AACB9C202733FFB0FE4AFF524B7C07D5" bold="true" box="[391,548,1137,1163]" pageId="73" pageNumber="74">
<collectingCountry id="E0A800A22733FFB0FE4AFF52488607D5" box="[391,478,1137,1163]" name="India" pageId="73" pageNumber="74">INDIA</collectingCountry>
:
<collectingRegion id="5A7B8ED02733FFB0FE24FF524B4707D5" box="[489,543,1137,1163]" country="India" name="Goa" pageId="73" pageNumber="74">Goa</collectingRegion>
:
</emphasis>
<collectingMunicipality id="7864DA482733FFB0FDE6FF524B0407D5" box="[555,604,1137,1163]" pageId="73" pageNumber="74">Goa</collectingMunicipality>
,
<locationDeviation id="89B259102733FFB0FDA5FF524A1807D2" box="[616,832,1137,1164]" location="Margao" pageId="73" pageNumber="74">
<quantity id="5F47EDD72733FFB0FDA5FF524BF607D5" box="[616,686,1137,1163]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="73" pageNumber="74" unit="km" value="30.0">30 km</quantity>
S of Margao
</locationDeviation>
(= Madgaon),
<location id="9D6016E92733FFB0FC2AFF524D1B07D5" LSID="urn:lsid:plazi:treatment:1016F1242733FFB0FF5AF8A84D340592:9D6016E92733FFB0FC2AFF524D1B07D5" box="[999,1091,1137,1163]" country="India" latitude="15.0078335" longLatPrecision="13" longitude="74.02634" municipality="Goa" name="Palolem" pageId="73" pageNumber="74" stateProvince="Goa">Palolem</location>
env. (
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),
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a.s.l.
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,
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</date>
,
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,
<collectorName id="354A25E42733FFB0FDE1FFB54BDF07EC" box="[556,647,1174,1202]" pageId="73" pageNumber="74">P. Šípek</collectorName>
&amp;
<collectorName id="354A25E42733FFB0FD67FFB54A7207F1" box="[682,810,1173,1200]" pageId="73" pageNumber="74">M. Fikáček</collectorName>
M. lgt.,
<determinerName id="6DDCACF62733FFB0FC47FFB54AA907EE" box="[906,1009,1174,1200]" pageId="73" pageNumber="74">P. Kment</determinerName>
det. (
<collectionCode id="FEAED8F72733FFB0FBFDFFB64DDE07F1" box="[1072,1158,1173,1199]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="73" pageNumber="74">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92733FFB0FF0AFF9A4B8D07A9" pageId="73" pageNumber="74" type="distribution">
<paragraph id="980040322733FFB0FF0AFF9A4B8D07A9" blockId="73.[151,1437,1137,1740]" pageId="73" pageNumber="74">
<emphasis id="AACB9C202733FFB0FF0AFF9A4839078D" bold="true" box="[199,353,1209,1235]" pageId="73" pageNumber="74">Distribution.</emphasis>
<collectingCountry id="E0A800A22733FFB0FEA4FF9A48FF078D" box="[361,423,1209,1235]" name="India" pageId="73" pageNumber="74">India</collectingCountry>
: Goa (new record),
<collectingRegion id="5A7B8ED02733FFB0FD46FF9A4BA6078D" box="[651,766,1209,1235]" country="India" name="Karnataka" pageId="73" pageNumber="74">Karnataka</collectingRegion>
(
<bibRefCitation id="FC2E3DC32733FFB0FCC3FF9A4AFC078D" author="Distant, W. L." box="[782,932,1209,1235]" pageId="73" pageNumber="74" pagination="1 - 210" refId="ref49585" refString="Distant, W. L. (1918) Rhynchota. Vol. VII. Homoptera: Appendix. Heteroptera: Addenda. In: Shipley, A. E. &amp; Marshall, A. K. (Eds.), The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, pp. i - vii + 1-210." type="book chapter" year="1918">Distant 1918</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32733FFB0FC7DFF9A4DDB078D" author="Salini, S. &amp; Gracy, R. G. &amp; Akoijam, R. &amp; Rabbani, M. K. &amp; David, K. J. &amp; Roca-Cusachs, M." box="[944,1155,1209,1235]" pageId="73" pageNumber="74" pagination="79 - 117" refId="ref54967" refString="Salini, S., Gracy, R. G., Akoijam, R., Rabbani, M. K., David, K. J. &amp; Roca-Cusachs, M. (2023 a) Revision of Acesines Stal and Dunnius Distant, resurrection of Mycterizon Breddin (Hemiptera, Heteroptera, Pentatomidae, Pentatominae), and description of a new species from India. ZooKeys, 1148, 79-117. https://doi.org/10.3897/zookeys.1148.95629" type="journal article" year="2023">
Salini
<emphasis id="AACB9C202733FFB0FC37FF994D6C078D" box="[1018,1076,1209,1235]" italics="true" pageId="73" pageNumber="74">et al.</emphasis>
2023a
</bibRefCitation>
, both as
<taxonomicName id="5FBF3BB12733FFB0FB27FF994CD4078D" authorityName="Distant" authorityYear="1918" box="[1258,1420,1209,1236]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bambusana">
<emphasis id="AACB9C202733FFB0FB27FF994CD4078D" box="[1258,1420,1209,1236]" italics="true" pageId="73" pageNumber="74">A. bambusana</emphasis>
</taxonomicName>
),
<collectingRegion id="5A7B8ED02733FFB0FF5AFFFE484407A9" box="[151,284,1245,1271]" country="India" name="Tamil Nadu" pageId="73" pageNumber="74">Tamil Nadu</collectingRegion>
(
<bibRefCitation id="FC2E3DC32733FFB0FEE7FFFE48A207A9" author="Salini, S. &amp; Gracy, R. G. &amp; Akoijam, R. &amp; Rabbani, M. K. &amp; David, K. J. &amp; Roca-Cusachs, M." box="[298,506,1245,1271]" pageId="73" pageNumber="74" pagination="79 - 117" refId="ref54967" refString="Salini, S., Gracy, R. G., Akoijam, R., Rabbani, M. K., David, K. J. &amp; Roca-Cusachs, M. (2023 a) Revision of Acesines Stal and Dunnius Distant, resurrection of Mycterizon Breddin (Hemiptera, Heteroptera, Pentatomidae, Pentatominae), and description of a new species from India. ZooKeys, 1148, 79-117. https://doi.org/10.3897/zookeys.1148.95629" type="journal article" year="2023">
Salini
<emphasis id="AACB9C202733FFB0FEB9FFFD48F507A9" box="[372,429,1245,1271]" italics="true" pageId="73" pageNumber="74">et al.</emphasis>
2023a
</bibRefCitation>
, as
<taxonomicName id="5FBF3BB12733FFB0FDE9FFFD4B9C07A9" authorityName="Distant" authorityYear="1918" box="[548,708,1245,1272]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bambusana">
<emphasis id="AACB9C202733FFB0FDE9FFFD4B9C07A9" box="[548,708,1245,1272]" italics="true" pageId="73" pageNumber="74">A. bambusana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92733FFB0FF0AFE224D340592" pageId="73" pageNumber="74" type="discussion">
<paragraph id="980040322733FFB0FF0AFE224D2906D6" blockId="73.[151,1437,1137,1740]" pageId="73" pageNumber="74">
<emphasis id="AACB9C202733FFB0FF0AFE2248630645" bold="true" box="[199,315,1281,1307]" pageId="73" pageNumber="74">Remarks.</emphasis>
<taxonomicName id="5FBF3BB12733FFB0FE8FFE214B4C0645" authorityName="Stal" authorityYear="1876" box="[322,532,1281,1308]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="AACB9C202733FFB0FE8FFE214B4C0645" box="[322,532,1281,1308]" italics="true" pageId="73" pageNumber="74">Acesines breviceps</emphasis>
</taxonomicName>
was originally described by
<bibRefCitation id="FC2E3DC32733FFB0FC9BFE224A8B0645" author="Stal, C." box="[854,979,1281,1308]" pageId="73" pageNumber="74" pagination="1 - 162" refId="ref55674" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en forteckning ofver alla hittills kanda Hemiptera, jemte systematiska meddelanden. [Checklist of Hemiptera. Contribution to a list of all hitherto known Hemiptera, along with systematic notes. 5]. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1-162. [in Latin]" type="journal article" year="1876">Stål (1876)</bibRefCitation>
based on a female specimen from “
<collectingCountry id="E0A800A22733FFB0FAADFE224CC40645" box="[1376,1436,1281,1307]" name="India" pageId="73" pageNumber="74">India</collectingCountry>
Orientalis”, which is currently housed in the Swedish Museum of Natural History,
<collectingRegion id="5A7B8ED02733FFB0FB80FE064D910661" box="[1101,1225,1317,1343]" country="Sweden" name="Stockholm" pageId="73" pageNumber="74">Stockholm</collectingRegion>
,
<collectingCountry id="E0A800A22733FFB0FB1BFE064C770661" box="[1238,1327,1317,1343]" name="Sweden" pageId="73" pageNumber="74">Sweden</collectingCountry>
(NHRS).
<taxonomicName id="5FBF3BB12733FFB0FF5AFE6948D9063D" authorityName="Distant" authorityYear="1918" box="[151,385,1353,1380]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bambusana">
<emphasis id="AACB9C202733FFB0FF5AFE6948D9063D" box="[151,385,1353,1380]" italics="true" pageId="73" pageNumber="74">Acesines bambusana</emphasis>
</taxonomicName>
was originally described by
<bibRefCitation id="FC2E3DC32733FFB0FD0EFE6A4A3D063D" author="Distant, W. L." box="[707,869,1353,1379]" pageId="73" pageNumber="74" pagination="1 - 210" refId="ref49585" refString="Distant, W. L. (1918) Rhynchota. Vol. VII. Homoptera: Appendix. Heteroptera: Addenda. In: Shipley, A. E. &amp; Marshall, A. K. (Eds.), The Fauna of British India, Including Ceylon and Burma. Taylor and Francis, London, pp. i - vii + 1-210." type="book chapter" year="1918">Distant (1918)</bibRefCitation>
based on a series of specimens from “South
<collectingCountry id="E0A800A22733FFB0FA96FE6A4CC1063D" box="[1371,1433,1353,1379]" name="India" pageId="73" pageNumber="74">India</collectingCountry>
: Chikkaballapura” collected by T. V. Campbell, which is currently housed in the BMNH.
</paragraph>
<paragraph id="980040322733FFB0FF0AFEB24D340592" blockId="73.[151,1437,1137,1740]" pageId="73" pageNumber="74">
According to
<bibRefCitation id="FC2E3DC32733FFB0FEA8FEB24B1306F5" author="Salini, S. &amp; Gracy, R. G. &amp; Akoijam, R. &amp; Rabbani, M. K. &amp; David, K. J. &amp; Roca-Cusachs, M." box="[357,587,1425,1451]" pageId="73" pageNumber="74" pagination="79 - 117" refId="ref54967" refString="Salini, S., Gracy, R. G., Akoijam, R., Rabbani, M. K., David, K. J. &amp; Roca-Cusachs, M. (2023 a) Revision of Acesines Stal and Dunnius Distant, resurrection of Mycterizon Breddin (Hemiptera, Heteroptera, Pentatomidae, Pentatominae), and description of a new species from India. ZooKeys, 1148, 79-117. https://doi.org/10.3897/zookeys.1148.95629" type="journal article" year="2023">
Salini
<emphasis id="AACB9C202733FFB0FE62FEB148B106F5" box="[431,489,1425,1451]" italics="true" pageId="73" pageNumber="74">et al.</emphasis>
(2023a)
</bibRefCitation>
, it was likely that
<taxonomicName id="5FBF3BB12733FFB0FCEDFEB14AF306F5" authorityName="Stal" authorityYear="1876" box="[800,939,1425,1452]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="AACB9C202733FFB0FCEDFEB14AF306F5" box="[800,939,1425,1452]" italics="true" pageId="73" pageNumber="74">A. breviceps</emphasis>
</taxonomicName>
was conspecific with
<taxonomicName id="5FBF3BB12733FFB0FB66FEB14C1406F5" authorityName="Distant" authorityYear="1918" box="[1195,1356,1425,1452]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bambusana">
<emphasis id="AACB9C202733FFB0FB66FEB14C1406F5" box="[1195,1356,1425,1452]" italics="true" pageId="73" pageNumber="74">A. bambusana</emphasis>
</taxonomicName>
due to the similarities in the colouration and the structural morphology of their
<typeStatus id="4704FE902733FFB0FC0DFE954AA6068E" box="[960,1022,1462,1488]" pageId="73" pageNumber="74">types</typeStatus>
, as noticed from the images received of the former species. Differences between the two species, such as the punctures on the scutellum, the colouration on the outer margins of the abdominal venter, and a small spinous projection at the caudal margins of laterotergites VIII, were considered insignificant. However, the synonymy was deferred at that time until it could be confirmed through the examination of the male genitalia structures of both species. Unfortunately, there is no way to match the male to the
<typeStatus id="4704FE902733FFB0FE87FD49482205DA" box="[330,378,1642,1668]" pageId="73" pageNumber="74">type</typeStatus>
of
<taxonomicName id="5FBF3BB12733FFB0FE6FFD494B7505DD" authorityName="Stal" authorityYear="1876" box="[418,557,1641,1668]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="AACB9C202733FFB0FE6FFD494B7505DD" box="[418,557,1641,1668]" italics="true" pageId="73" pageNumber="74">A. breviceps</emphasis>
</taxonomicName>
by locality, as no precise locality is known for its
<typeStatus id="4704FE902733FFB0FBBDFD494DFB05DA" box="[1136,1187,1642,1668]" pageId="73" pageNumber="74">type</typeStatus>
, and comparing DNA sequences may not be feasible for such an old specimen. Therefore, based on the evidence presented above, it is prudent to consider the name
<taxonomicName id="5FBF3BB12733FFB0FE2EFD914B950595" authorityName="Distant" authorityYear="1918" box="[483,717,1713,1740]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="bambusana">
<emphasis id="AACB9C202733FFB0FE2EFD914B950595" box="[483,717,1713,1740]" italics="true" pageId="73" pageNumber="74">Acesines bambusana</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="5FBF3BB12733FFB0FC11FD914D3D0595" authorityName="Stal" authorityYear="1876" box="[988,1125,1713,1740]" class="Insecta" family="Pentatomidae" genus="Acesines" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="AACB9C202733FFB0FC11FD914D3D0595" box="[988,1125,1713,1740]" italics="true" pageId="73" pageNumber="74">A. breviceps</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,826 @@
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<mods:title id="19729141F7D49E78AC97FB44A549A319">Revision of the Austro-Oriental shield-bug genus Axiagastus (Hemiptera: Heteroptera: Pentatomidae), with the description of five new species, and taxonomic notes on related genera</mods:title>
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<mods:namePart id="6BE98C258B84A70E7DCF523353A9C44D">Kment, Petr</mods:namePart>
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<mods:affiliation id="E587D077F85DFFEAEE0313761F4A66E6">Department of Entomology, National Museum of the Czech Republic, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
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<mods:affiliation id="4C1331F405A96536ADF863AA1133C68E">Department of Collections (Insects), The Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom; &amp; val. heureuse @ gmail. com; https: // orcid. org / 0000 - 0003 - 4802 - 2711</mods:affiliation>
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<mods:affiliation id="C29A5ACB065A763D3ABB314661901238">Department of Collections (Insects), The Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom; &amp; mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
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<mods:affiliation id="AAA7A88052133B014F070B60681D307A">ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, Karnataka, India shalinis. nilavu @ gmail. com; https: // orcid. org / 0000 - 0003 - 3557 - 4365 &amp; Evolution &amp; Ecology Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, Australia &amp; IRBio, Institut de Recerca de la Biodiversitat, Universitat de Barcelona, Barcelona, Spain marcosrocacusachs @ gmail. com; https: // orcid. org / 0000 - 0002 - 9174 - 6021 # Former affiliation * Corresponding author &amp; ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, Karnataka, India shalinis. nilavu @ gmail. com; https: // orcid. org / 0000 - 0003 - 3557 - 4365</mods:affiliation>
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<mods:title id="A16040684E596B1E6BEF18E674FC04F6">Zootaxa</mods:title>
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<mods:part id="97904F6F9686AF7C8F643632B50EF575">
<mods:date id="2443DDBE0E6174EA7CF34337556817F2">2025</mods:date>
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<mods:number id="328A9C5605C6F7C6A75124B134353A79">2025-03-11</mods:number>
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<mods:number id="410D1CF2393FCFA706ACA0F9AC04FDB4">5603</mods:number>
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<mods:number id="1F6ACE5154FAF324503D6EE21987809F">1</mods:number>
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<mods:url id="8D7674A4F780CF88229DDDA849A67E71">https://doi.org/10.11646/zootaxa.5603.1.1</mods:url>
</mods:location>
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<mods:identifier id="F9175EA3D000AA4A4A9A38713B3052E9" type="ISSN">1175-5334</mods:identifier>
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<treatment id="1016F1242746FFB9FF5AFBB44B4404A4" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4" httpUri="http://treatment.plazi.org/id/1016F1242746FFB9FF5AFBB44B4404A4" lastPageId="64" lastPageNumber="65" pageId="60" pageNumber="61">
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<paragraph id="980040322746FFC5FF5AFBB44BE303EF" blockId="60.[151,1174,151,250]" box="[151,699,151,178]" pageId="60" pageNumber="61">
<heading id="C348F75E2746FFC5FF5AFBB44BE303EF" bold="true" box="[151,699,151,178]" fontSize="11" level="2" pageId="60" pageNumber="61" reason="3">
<emphasis id="AACB9C202746FFC5FF5AFBB44BE303EF" bold="true" box="[151,699,151,178]" pageId="60" pageNumber="61">
<taxonomicName id="5FBF3BB12746FFC5FF5AFBB44B0D03EF" authority="Salini &amp; Kment" authorityName="Salini &amp; Kment" authorityYear="2025" box="[151,597,151,178]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="votypkai" status="sp. nov.">
<emphasis id="AACB9C202746FFC5FF5AFBB448D103EC" bold="true" box="[151,393,151,178]" italics="true" pageId="60" pageNumber="61">Axiagastus votypkai</emphasis>
Salini &amp; Kment
</taxonomicName>
<taxonomicNameLabel id="B1F8215B2746FFC5FD90FBB44BE303EF" box="[605,699,151,177]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92746FFC5FF5AFBFC481B03A4" box="[151,323,223,250]" pageId="60" pageNumber="61" type="description">
<paragraph id="980040322746FFC5FF5AFBFC481B03A4" blockId="60.[151,1174,151,250]" box="[151,323,223,250]" pageId="60" pageNumber="61">
(
<figureCitation id="00845CB72746FFC5FF52FBFC486403A4" box="[159,316,223,250]" captionStart-0="FIGURES 200206" captionStart-1="FIGURES 207213" captionStart-2="FIGURES 214219" captionStart-3="FIGURES 220225" captionStartId-0="62.[151,264,1859,1883]" captionStartId-1="63.[151,264,1909,1933]" captionStartId-2="64.[151,264,1112,1136]" captionStartId-3="65.[151,264,1107,1131]" captionTargetBox-0="[151,1436,201,1835]" captionTargetBox-1="[192,1396,181,1885]" captionTargetBox-2="[151,1436,181,1089]" captionTargetBox-3="[151,1436,181,1083]" captionTargetId-0="figure-29@62.[151,1436,201,1835]" captionTargetId-1="figure-19@63.[192,1396,181,1885]" captionTargetId-2="figure-29@64.[151,1436,181,1089]" captionTargetId-3="figure-19@65.[151,1436,181,1083]" captionTargetPageId-0="62" captionTargetPageId-1="63" captionTargetPageId-2="64" captionTargetPageId-3="65" captionText-0="FIGURES 200206. Axiagastus votypkai Salini &amp; Kment sp. nov. 200201—habitus (200—dorsal view, 201—ventral view), 202203—head (202—dorsal view, 203—lateral view); 204—external scent efferent system; 205—apex of male abdomen with genital capsule in postero-ventral view; 206—apex of female abdomen with terminalia in posteroventral view. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 1 mm (200201), 0.5 mm (202206). Orig. S. Salini." captionText-1="FIGURES 207213. Male genitalia (ESEM micrographs) of Axiagastus votypkai Salini &amp; Kment sp. nov. 207210—genital capsule (207—dorsal view, 32×; 208—ventral view, 32×; 209—caudal view, 40×; 210—lateral view, 45×); 211213—parameres (three different views, 100×). Lettering: dn—distension, te—transverse emargination. Scale: 1 mm (207210), 0.5 mm (211 213). Orig. P. Kment." captionText-2="FIGURES 214219. Axiagastus votypkai Salini &amp; Kment sp. nov., male genitalia. 214217—genital capsule (214—dorsal view, 215—ventral view, 216—caudal view, 217—lateral view); 218219—parameres (two different views). Lettering: dm— dorsal margin, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, te—transverse emargination. Scale: 0.5 mm (214217), 0.25 mm (218219). Orig. S. Salini." captionText-3="FIGURES 220225. Axiagastus votypkai Salini &amp; Kment sp. nov., male and female genitalia. 220222—phallus (220—dorsal view, 221—ventral view, 222—lateral view); 223—terminalia (dorsal view); 224—terminalia (ventral view); 225—spermatheca. Scale: 0.1 mm (220222, 225), 0.5 mm (223224). Orig. S. Salini." pageId="60" pageNumber="61">Figs 200225</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92746FFC5FF5AFA034A570208" pageId="60" pageNumber="61" type="reference_group">
<paragraph id="980040322746FFC5FF5AFA034CC50266" blockId="60.[151,1437,287,342]" box="[151,1437,287,312]" pageId="60" pageNumber="61">
<treatmentCitationGroup id="B8AF671C2746FFC5FF5AFA034CC50266" box="[151,1437,287,312]" pageId="60" pageNumber="61">
?
<taxonomicName id="5FBF3BB12746FFC5FF6FFA0348220269" authorityName="Dallas" authorityYear="1851" box="[162,378,288,311]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C202746FFC5FF6FFA0348220269" box="[162,378,288,311]" italics="true" pageId="60" pageNumber="61">Axiagastus rosmarus</emphasis>
</taxonomicName>
(misidentification, partim):
<treatmentCitation id="191E66232746FFC5FD6EFA034A280269" author="Walker, F." box="[675,880,287,312]" page="269" pageId="60" pageNumber="61" year="1867">
<bibRefCitation id="FC2E3DC32746FFC5FD6EFA034A650269" author="Walker, F." box="[675,829,287,312]" pageId="60" pageNumber="61" pagination="241 - 417" refId="ref56699" refString="Walker, F. (1867) Catalogue of the specimens of Heteropterous Hemiptera in the collection of the British Museum. Part II. Scutata. Trustees of the British Museum, London, 177 pp. [pp. 241-417]" type="book chapter" year="1867">Walker (1867)</bibRefCitation>
: 269
</treatmentCitation>
(distribution: New
<collectingCountry id="E0A800A22746FFC5FB8FFA034DC80266" box="[1090,1168,288,312]" name="Guinea-Bissau" pageId="60" pageNumber="61">Guinea</collectingCountry>
);
<bibRefCitation id="FC2E3DC32746FFC5FB69FA034CC10266" author="Cassis, G. &amp; Gross, G. F." box="[1188,1433,288,312]" pageId="60" pageNumber="61" pagination="1 - 737" refId="ref48921" refString="Cassis, G. &amp; Gross, G. F. (2002) Hemiptera: Heteroptera (Pentatomomorpha). In: Houston, W. W. K. &amp; Wells, A. (Eds.), Zoological Catalogue of Australia. Vol. 27.3 B. CSIRO Publishing, Melbourne, pp. i - xiv + 1-737." type="book chapter" year="2002">Cassis &amp; Gross (2002)</bibRefCitation>
:
</treatmentCitationGroup>
</paragraph>
<paragraph id="980040322746FFC5FF0AFA1D4A570208" blockId="60.[151,1437,287,342]" box="[199,783,318,342]" pageId="60" pageNumber="61">
458459 (catalogue, distribution:
<collectingCountry id="E0A800A22746FFC5FDD0FA1D4B260208" box="[541,638,318,342]" name="Australia" pageId="60" pageNumber="61">Australia</collectingCountry>
:
<collectingRegion id="5A7B8ED02746FFC5FD4AFA1D4A5B0208" box="[647,771,318,342]" country="Australia" name="Queensland" pageId="60" pageNumber="61">Queensland</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92746FFC4FF5AFAA348FD0242" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="materials_examined">
<paragraph id="980040322746FFC5FF5AFAA34A3802C2" blockId="60.[151,1437,384,2042]" box="[151,864,384,412]" pageId="60" pageNumber="61">
<emphasis id="AACB9C202746FFC5FF5AFAA3486E02C4" bold="true" box="[151,310,384,410]" pageId="60" pageNumber="61">Type locality.</emphasis>
<materialsCitation id="28D74A6F2746FFC5FEF1FAA34A0402C2" box="[316,860,384,412]" country="Indonesia" latitude="-3.3" location="Buare River" longLatPrecision="1307" longitude="138.71666" pageId="60" pageNumber="61" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry id="E0A800A22746FFC5FEF1FAA348F402C4" box="[316,428,384,410]" name="Indonesia" pageId="60" pageNumber="61">Indonesia</collectingCountry>
,
<collectingRegion id="5A7B8ED02746FFC5FE7AFAA348A602C4" box="[439,510,384,410]" country="Indonesia" name="Papua Barat" pageId="60" pageNumber="61">Papua</collectingRegion>
,
<location id="9D6016E92746FFC5FDC7FAA34BCB02C4" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FDC7FAA34BCB02C4" box="[522,659,384,410]" country="Indonesia" latitude="-3.3" longLatPrecision="1307" longitude="138.71666" name="Buare River" pageId="60" pageNumber="61" stateProvince="Papua Barat">Buare River</location>
,
<geoCoordinate id="FD8B26F52746FFC5FD52FAA24BB202C2" box="[671,746,385,412]" degrees="3" direction="south" minutes="18" orientation="latitude" pageId="60" pageNumber="61" precision="925" value="-3.3">3°18S</geoCoordinate>
<geoCoordinate id="FD8B26F52746FFC5FD3DFAA24A0402C2" box="[752,860,385,412]" degrees="138" direction="east" minutes="43" orientation="longitude" pageId="60" pageNumber="61" precision="925" value="138.71666">138°43E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="980040322746FFC5FF0AFA804B3601CC" blockId="60.[151,1437,384,2042]" pageId="60" pageNumber="61">
<emphasis id="AACB9C202746FFC5FF0AFA80482F02E3" bold="true" box="[199,375,419,445]" pageId="60" pageNumber="61">Type material.</emphasis>
<materialsCitation id="28D74A6F2746FFC5FE4DFA804B3601CC" collectingDate="2009-01-16" collectionCode="NIM" collectorName="A. Zamesov" country="Papua New Guinea" determinerName="Roca-Cusachs &amp; Salini, S." location="Irian" longitude="3.18" municipality="Buare River" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="4704FE902746FFC5FE4DFA8048B502E3" box="[384,493,419,445]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
:
<specimenCount id="8EB98BBB2746FFC5FE34FA874B4802E0" box="[505,528,420,446]" count="1" pageId="60" pageNumber="61" type="male"></specimenCount>
, “
<collectingCountry id="E0A800A22746FFC5FDE5FA804A1C02E3" box="[552,836,419,446]" name="Papua New Guinea" pageId="60" pageNumber="61">PAPUA NEW GUINEA</collectingCountry>
:
<location id="9D6016E92746FFC5FC9CFA804AD102E3" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FC9CFA804AD102E3" box="[849,905,419,445]" country="Papua New Guinea" longitude="3.18" municipality="Buare River" name="Irian" pageId="60" pageNumber="61">Irian</location>
,
<collectingMunicipality id="7864DA482746FFC5FC5BFA874D7802E3" box="[918,1056,419,446]" pageId="60" pageNumber="61">Buare River</collectingMunicipality>
[p] / (S
<geoCoordinate id="FD8B26F52746FFC5FBB3FA804D8B02E3" box="[1150,1235,419,445]" degrees="3, 18" direction="east" orientation="longitude" pageId="60" pageNumber="61" precision="555" value="3.18">3, 18 E</geoCoordinate>
138, 43) 5.I- [p] /
<date id="EC0166F22746FFC5FF5AFAE4485B02BF" box="[151,259,455,481]" pageId="60" pageNumber="61" value="2009-01-16">
<collectingDate id="FC459F1A2746FFC5FF5AFAE4485B02BF" box="[151,259,455,481]" pageId="60" pageNumber="61" value="2009-01-16">16.i.2009</collectingDate>
</date>
,
<collectorName id="354A25E42746FFC5FEC0FAE448CD02BF" box="[269,405,455,481]" pageId="60" pageNumber="61">A. Zamesov</collectorName>
lgt. [p] //
<taxonomicName id="5FBF3BB12746FFC5FDCEFAE44BD202BF" authorityName="Dallas" authorityYear="1851" box="[515,650,455,481]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C202746FFC5FDCEFAE44BD202BF" box="[515,650,455,481]" italics="true" pageId="60" pageNumber="61">A. rosmarus</emphasis>
</taxonomicName>
[hw] /
<determinerName id="6DDCACF62746FFC5FD14FAE44A2102BF" box="[729,889,455,481]" pageId="60" pageNumber="61">Roca-Cusachs</determinerName>
det. 2021 // dissected and illustrated by Salini. S. [p] // Roca-Cusachs Personal Collection [p] //
<typeStatus id="4704FE902746FFC5FD5AFAC94A65015A" box="[663,829,490,516]" pageId="60" pageNumber="61" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="5FBF3BB12746FFC5FC9CFAC84D23015B" authorityName="Salini &amp; Kment" authorityYear="2025" box="[849,1147,491,517]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="66" phylum="Arthropoda" rank="species" species="votypkai">
<emphasis id="AACB9C202746FFC5FC9CFAC84D23015B" box="[849,1147,491,517]" italics="true" pageId="60" pageNumber="61">AXIAGASTUS VOTYPKAI</emphasis>
</taxonomicName>
/ Salini &amp; Kment
<emphasis id="AACB9C202746FFC5FA89FAC94CC4015A" bold="true" box="[1348,1436,490,516]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="B1F8215B2746FFC5FA89FAC94CC4015A" box="[1348,1436,490,516]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
// det.
<determinerName id="6DDCACF62746FFC5FF13F92D481C0176" box="[222,324,526,552]" pageId="60" pageNumber="61">Salini, S.</determinerName>
&amp; Kment, P., 2024 [red label]” (
<collectionCode id="FEAED8F72746FFC5FD79F92D4BA80176" box="[692,752,526,552]" pageId="60" pageNumber="61">NIM</collectionCode>
). The
<typeStatus id="4704FE902746FFC5FCFBF92D4AC00176" box="[822,920,526,552]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
is pinned through the scutellum, the detached left hind leg, right antennomere IV and left antennomeres IIa, IIb and III are glued on a card and the dissected male genitalia are preserved in a glass microvial with glycerol attached to the same pin; right mesotarsomere III along with claws and right antennomere IV lost.
</materialsCitation>
</paragraph>
<paragraph id="980040322746FFC5FF0AF9BF48B704A4" blockId="60.[151,1437,384,2042]" pageId="60" pageNumber="61">
<materialsCitation id="28D74A6F2746FFC5FF0AF9BF4A4D01EB" box="[199,789,667,694]" country="Australia" location="Queensland" pageId="60" pageNumber="61" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Queensland" typeStatus="paratype">
<typeStatus id="4704FE902746FFC5FF0AF9BF486C01E8" box="[199,308,668,694]" pageId="60" pageNumber="61" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="8EB98BBB2746FFC5FE89F9BF482001E8" box="[324,376,668,694]" count="10" pageId="60" pageNumber="61" type="male">10♂</specimenCount>
,
<specimenCount id="8EB98BBB2746FFC5FE49F9BF48E001E8" box="[388,440,668,694]" count="10" pageId="60" pageNumber="61" type="female">10♀</specimenCount>
):
<emphasis id="AACB9C202746FFC5FE03F9B84A4D01EB" bold="true" box="[462,789,667,693]" pageId="60" pageNumber="61">
<collectingCountry id="E0A800A22746FFC5FE03F9B84B2F01EB" box="[462,631,667,693]" name="Australia" pageId="60" pageNumber="61">AUSTRALIA</collectingCountry>
:
<collectingRegion id="5A7B8ED02746FFC5FD4EF9B84A4D01EB" box="[643,789,667,693]" country="Australia" name="Queensland" pageId="60" pageNumber="61">Queensland</collectingRegion>
</emphasis>
</materialsCitation>
<emphasis id="AACB9C202746FFC5FCD8F9B84A4201EB" bold="true" box="[789,794,667,693]" pageId="60" pageNumber="61">:</emphasis>
<materialsCitation id="28D74A6F2746FFC5FCEFF9BF4A330187" collectingDate="1996-12-04" collectionCode="BMNH" country="Australia" location="Kuranda" municipality="Australia" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Queensland" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FCEFF9BF4A1E01E8" box="[802,838,668,694]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
, “
<collectingMunicipality id="7864DA482746FFC5FC90F9BF4A9E01E8" box="[861,966,668,694]" pageId="60" pageNumber="61">Australia</collectingMunicipality>
[hw] /
<collectingRegion id="5A7B8ED02746FFC5FBD4F9BF4DC501E8" box="[1049,1181,668,694]" country="Australia" name="Queensland" pageId="60" pageNumber="61">Queensland</collectingRegion>
[hw] /
<location id="9D6016E92746FFC5FB3DF9BF4C0801E8" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FB3DF9BF4C0801E8" box="[1264,1360,668,694]" country="Australia" municipality="Australia" name="Kuranda" pageId="60" pageNumber="61" stateProvince="Queensland">Kuranda</location>
[hw] / rain forest [hw] /
<date id="EC0166F22746FFC5FEADF99C48B80187" box="[352,480,703,729]" pageId="60" pageNumber="61" value="1996-12-04">
<collectingDate id="FC459F1A2746FFC5FEADF99C48B80187" box="[352,480,703,729]" pageId="60" pageNumber="61" value="1996-12-04">04.xii.1996</collectingDate>
</date>
[hw] / M. Shaffer [hw]” (
<collectionCode id="FEAED8F72746FFC5FCC4F99C4A3B0187" box="[777,867,703,729]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="28D74A6F2746FFC5FCBAF99C4DC50187" box="[887,1181,703,730]" country="Australia" location="Left" pageId="60" pageNumber="61" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<location id="9D6016E92746FFC5FCBAF99C4AFE0187" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FCBAF99C4AFE0187" box="[887,934,703,729]" country="Australia" name="Left" pageId="60" pageNumber="61" stateProvince="Queensland">Left</location>
antennomere IV lost
</materialsCitation>
.—
<materialsCitation id="28D74A6F2746FFC5FB09F99C4C2301A3" collectingDate="1962-01-18" collectionCode="BMNH" collectorName="E. B. Britton." country="Australia" location="Cairns" pageId="60" pageNumber="61" specimenCount="2" specimenCount-female="2" stateProvince="Queensland" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FB09F99C4D850187" box="[1220,1245,703,729]" count="1" pageId="60" pageNumber="61" type="female">1♀</specimenCount>
, “AUSTRALIA. N.Q.: /
<location id="9D6016E92746FFC5FF38F9C1486601A2" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FF38F9C1486601A2" box="[245,318,738,764]" country="Australia" name="Cairns" pageId="60" pageNumber="61" stateProvince="Queensland">Cairns</location>
/
<date id="EC0166F22746FFC5FE90F9C1489201A2" box="[349,458,738,764]" pageId="60" pageNumber="61" value="1962-01-18">
<collectingDate id="FC459F1A2746FFC5FE90F9C1489201A2" box="[349,458,738,764]" pageId="60" pageNumber="61" value="1962-01-18">18.i.1962</collectingDate>
</date>
. /
<collectorName id="354A25E42746FFC5FE20F9C14B2D01A2" box="[493,629,738,764]" pageId="60" pageNumber="61">E.B.Britton.</collectorName>
/ B.M. 1962-153. [p] // From light / trap [p] //
<specimenCount id="8EB98BBB2746FFC5FB74F9C04D9601A3" box="[1209,1230,739,765]" count="1" pageId="60" pageNumber="61" type="female"></specimenCount>
[p]” (
<collectionCode id="FEAED8F72746FFC5FAD7F9C04C2A01A3" box="[1306,1394,739,765]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="28D74A6F2746FFC5FF5AF8254B8C001A" collectionCode="BMNH" collectorName="W. Stuber" country="Indonesia" location="Humboldt Bay Dist." pageId="60" pageNumber="61" specimenCount="3" specimenCount-female="3" stateProvince="Papua Barat" typeStatus="paratype">
<emphasis id="AACB9C202746FFC5FF5AF82548C4007E" bold="true" box="[151,412,774,800]" pageId="60" pageNumber="61">
<collectingCountry id="E0A800A22746FFC5FF5AF8254862007E" box="[151,314,774,800]" name="Indonesia" pageId="60" pageNumber="61">INDONESIA</collectingCountry>
:
<collectingRegion id="5A7B8ED02746FFC5FE8BF82548CF007E" box="[326,407,774,800]" country="Indonesia" name="Papua Barat" pageId="60" pageNumber="61">Papua</collectingRegion>
:
</emphasis>
<specimenCount id="8EB98BBB2746FFC5FE6EF825489F007E" box="[419,455,774,800]" count="2" pageId="60" pageNumber="61" type="female">2♀</specimenCount>
, “DUTCH NEW GUINEA: /
<location id="9D6016E92746FFC5FCDAF8254AA2007E" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FCDAF8254AA2007E" box="[791,1018,774,801]" country="Indonesia" name="Humboldt Bay Dist." pageId="60" pageNumber="61" stateProvince="Papua Barat">Humboldt Bay Dist.</location>
/ 1937. /
<collectorName id="354A25E42746FFC5FBA9F8244D8B007E" box="[1124,1235,774,801]" pageId="60" pageNumber="61">W. Stüber</collectorName>
/ B.M. 1938-177. [p] //
<specimenCount id="8EB98BBB2746FFC5FF14F80949B6001A" box="[217,238,810,836]" count="1" pageId="60" pageNumber="61" type="female"></specimenCount>
[p] // NHMUK 013589097 [p]” (
<collectionCode id="FEAED8F72746FFC5FDBFF8094B94001A" box="[626,716,810,836]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="28D74A6F2746FFC5FD2DF80A4DB8001D" box="[736,1248,809,836]" country="Indonesia" location="Detached" pageId="60" pageNumber="61" specimenCount="1" stateProvince="Papua Barat" typeStatus="paratype">
<location id="9D6016E92746FFC5FD2DF80A4A12001D" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FD2DF80A4A12001D" box="[736,842,809,835]" country="Indonesia" name="Detached" pageId="60" pageNumber="61" stateProvince="Papua Barat">Detached</location>
abdomen glued on a piece of card
</materialsCitation>
.—
<materialsCitation id="28D74A6F2746FFC5FAC8F80A488D00D5" collectingDate="1937-06" collectionCode="BMNH" collectorName="W. Stuber." country="Indonesia" county="Humboldt Bay Dist." location="West of Tami River." municipality="Pukusam Dist." pageId="60" pageNumber="61" specimenCount="3" specimenCount-female="3" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FAC8F80A4C7A001D" box="[1285,1314,809,835]" count="2" pageId="60" pageNumber="61" type="female">2♀</specimenCount>
, “DUTCH NEW GUINEA: /
<collectingCounty id="716138BE2746FFC5FEAEF86E4B1C0039" box="[355,580,845,872]" pageId="60" pageNumber="61">Humboldt Bay Dist.</collectingCounty>
/
<collectingMunicipality id="7864DA482746FFC5FD9AF86E4BA20039" box="[599,762,845,871]" pageId="60" pageNumber="61">Pukusam Dist.</collectingMunicipality>
/
<location id="9D6016E92746FFC5FCC0F86D4AB30039" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FCC0F86D4AB30039" box="[781,1003,845,872]" country="Indonesia" county="Humboldt Bay Dist." municipality="Pukusam Dist." name="West of Tami River." pageId="60" pageNumber="61" stateProvince="Papua Barat">West of Tami River.</location>
/
<date id="EC0166F22746FFC5FC33F86E4D0A0039" box="[1022,1106,845,871]" pageId="60" pageNumber="61" value="1937-06">
<collectingDate id="FC459F1A2746FFC5FC33F86E4D0A0039" box="[1022,1106,845,871]" pageId="60" pageNumber="61" value="1937-06">vi.1937</collectingDate>
</date>
[p] //
<collectorName id="354A25E42746FFC5FB59F86D4C500039" box="[1172,1288,845,872]" pageId="60" pageNumber="61">W. Stüber.</collectorName>
/ B.M. 1938- 177. [p] //
<specimenCount id="8EB98BBB2746FFC5FEDEF852487000D5" box="[275,296,881,907]" count="1" pageId="60" pageNumber="61" type="female"></specimenCount>
[p]” (
<collectionCode id="FEAED8F72746FFC5FEA2F852489500D5" box="[367,461,881,907]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="28D74A6F2746FFC5FE34F8524AC500F0" collectingDate="1937-07" collectionCode="BMNH" collectorName="W. Stuber." country="Indonesia" location="Bewani Mts." municipality="Humboldt Bay Dist." pageId="60" pageNumber="61" specimenCount="2" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FE34F8524B4E00D5" box="[505,534,881,907]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
, “DUTCH NEW GUINEA: /
<collectingMunicipality id="7864DA482746FFC5FCA3F8534D0C00D4" box="[878,1108,880,907]" pageId="60" pageNumber="61">Humboldt Bay Dist.</collectingMunicipality>
/
<location id="9D6016E92746FFC5FBA0F8534DA700D5" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FBA0F8534DA700D5" box="[1133,1279,880,907]" country="Indonesia" municipality="Humboldt Bay Dist." name="Bewani Mts." pageId="60" pageNumber="61" stateProvince="Papua Barat">Bewani Mts.</location>
/ 400 metres. /
<date id="EC0166F22746FFC5FF6AF8B7485C00F0" box="[167,260,916,942]" pageId="60" pageNumber="61" value="1937-07">
<collectingDate id="FC459F1A2746FFC5FF6AF8B7485C00F0" box="[167,260,916,942]" pageId="60" pageNumber="61" value="1937-07">vii.1937</collectingDate>
</date>
[p] //
<collectorName id="354A25E42746FFC5FE81F8B7489B00F0" box="[332,451,916,942]" pageId="60" pageNumber="61">W. Stüber.</collectorName>
/ B.M. 1938-177. [p] //
<specimenCount id="8EB98BBB2746FFC5FD1AF8B74BB400F0" box="[727,748,916,942]" count="1" pageId="60" pageNumber="61" type="male"></specimenCount>
[p]” (
<collectionCode id="FEAED8F72746FFC5FCFEF8B74ACC00F0" box="[819,916,916,942]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="28D74A6F2746FFC5FC0CF8B749BC00AB" collectingDate="2009-01-16" collectionCode="NIM" collectorName="A. Zamesov" country="Indonesia" location="Buare River" longitude="3.18" municipality="Irian" pageId="60" pageNumber="61" specimenCount="1" specimenCount-female="1" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FC0CF8B74A8100F0" box="[961,985,916,942]" count="1" pageId="60" pageNumber="61" type="female">1♀</specimenCount>
, “
<collectingRegion id="5A7B8ED02746FFC5FC22F8B74D1200F0" box="[1007,1098,916,942]" country="Indonesia" name="Papua Barat" pageId="60" pageNumber="61">PAPUA</collectingRegion>
NEW GUINEA:
<collectingMunicipality id="7864DA482746FFC5FAD8F8B74C1500F0" box="[1301,1357,916,942]" pageId="60" pageNumber="61">Irian</collectingMunicipality>
,
<location id="9D6016E92746FFC5FA94F8B7498D008F" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FA94F8B7498D008F" country="Indonesia" longitude="3.18" municipality="Irian" name="Buare River" pageId="60" pageNumber="61" stateProvince="Papua Barat">Buare River</location>
[p] / (S
<geoCoordinate id="FD8B26F52746FFC5FEFDF89448DB008F" box="[304,387,951,977]" degrees="3, 18" direction="east" orientation="longitude" pageId="60" pageNumber="61" precision="555" value="3.18">3, 18 E</geoCoordinate>
138, 43) 5.I- [p] /
<date id="EC0166F22746FFC5FD82F8944BE3008F" box="[591,699,951,977]" pageId="60" pageNumber="61" value="2009-01-16">
<collectingDate id="FC459F1A2746FFC5FD82F8944BE3008F" box="[591,699,951,977]" pageId="60" pageNumber="61" value="2009-01-16">16.i.2009</collectingDate>
</date>
,
<collectorName id="354A25E42746FFC5FD0AF89B4A09008C" box="[711,849,952,978]" pageId="60" pageNumber="61">A. Zamesov</collectorName>
lgt. [p] // dissected and illustrated by Salini. S. [p]” (
<collectionCode id="FEAED8F72746FFC5FF52F8F8498300AB" box="[159,219,987,1013]" pageId="60" pageNumber="61">NIM</collectionCode>
)
</materialsCitation>
. Dissected female genitalia are placed in a glass microvial with glycerol attached to the same pin, terga and sterna, after extraction of terminalia and spermatheca, glued on a card, left foreleg and antennomere IV are glued to the card.
<materialsCitation id="28D74A6F2746FFC5FECCFF024AFD0625" collectingDate="2009-01-25" collectingDateMax="2011-05-09" collectingDateMin="2009-01-25" collectionCode="NMPC, MRCC" collectorName="A. Zamesov" country="Papua New Guinea" county="Binatang Research Center" elevation="20" latitude="-5.1563888" location="Charles University in Prague" longLatPrecision="21" longitude="145.79472" municipality="Department of Parasitology" pageId="60" pageNumber="61" specimenCount="9" specimenCount-female="3" specimenCount-male="2" stateProvince="Madang" typeStatus="paratype">
<emphasis id="AACB9C202746FFC5FECCFF024BC50762" bold="true" box="[257,669,1057,1084]" pageId="60" pageNumber="61">
<collectingCountry id="E0A800A22746FFC5FECCFF024B7B0765" box="[257,547,1057,1084]" name="Papua New Guinea" pageId="60" pageNumber="61">PAPUA NEW GUINEA</collectingCountry>
:
<collectingRegion id="5A7B8ED02746FFC5FDFDFF014BC00762" box="[560,664,1058,1084]" country="Papua New Guinea" name="Madang" pageId="60" pageNumber="61">Madang</collectingRegion>
:
</emphasis>
<specimenCount id="8EB98BBB2746FFC5FD68FF014B920762" box="[677,714,1058,1084]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
, “PAPUA NG,
<collectingRegion id="5A7B8ED02746FFC5FCB6FF014A820762" box="[891,986,1058,1084]" country="Papua New Guinea" name="Madang" pageId="60" pageNumber="61">Madang</collectingRegion>
, NAGADA /
<collectingCounty id="716138BE2746FFC5FBBBFF014CC50762" box="[1142,1437,1058,1084]" pageId="60" pageNumber="61">Binatang Research Center</collectingCounty>
/
<geoCoordinate id="FD8B26F52746FFC5FF6BFF644853073C" box="[166,267,1095,1122]" degrees="5" direction="south" minutes="9" orientation="latitude" pageId="60" pageNumber="61" precision="15" seconds="23" value="-5.1563888">5°923″S</geoCoordinate>
<geoCoordinate id="FD8B26F52746FFC5FEDEFF6448FE073C" box="[275,422,1095,1122]" degrees="145" direction="east" minutes="47" orientation="longitude" pageId="60" pageNumber="61" precision="15" seconds="41" value="145.79472">145°4741″E</geoCoordinate>
,
<quantity id="5F47EDD72746FFC5FE7CFF6648B3073E" box="[433,491,1093,1120]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="60" pageNumber="61" unit="m" value="20.0">
<elevation id="1392A7012746FFC5FE7CFF6648B3073E" box="[433,491,1093,1120]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="60" pageNumber="61" unit="m" value="20.0">20 m</elevation>
</quantity>
/ gardens and ruderal vegetation /
<date id="EC0166F22746FFC5FCBFFF664A8B0701" box="[882,979,1093,1119]" pageId="60" pageNumber="61" value="2011-05-06">
<collectingDate id="FC459F1A2746FFC5FCBFFF664A8B0701" box="[882,979,1093,1119]" pageId="60" pageNumber="61" value="2011-05-06">6.v.2011</collectingDate>
</date>
, Votýpka &amp; Lukeš lgt. [p] //
<collectingMunicipality id="7864DA482746FFC5FAD5FF65481B07DD" pageId="60" pageNumber="61">Department of Parasitology</collectingMunicipality>
/
<location id="9D6016E92746FFC5FE91FF4A4BFD07DD" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FE91FF4A4BFD07DD" box="[348,677,1129,1155]" country="Papua New Guinea" county="Binatang Research Center" latitude="-5.1563888" longLatPrecision="21" longitude="145.79472" municipality="Department of Parasitology" name="Charles University in Prague" pageId="60" pageNumber="61" stateProvince="Madang">Charles University in Prague</location>
/ DISSECTION
<location id="9D6016E92746FFC5FCA7FF4A4ACB07DD" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FCA7FF4A4ACB07DD" box="[874,915,1129,1155]" country="Papua New Guinea" county="Binatang Research Center" latitude="-5.1563888" longLatPrecision="21" longitude="145.79472" municipality="Department of Parasitology" name="No." pageId="60" pageNumber="61" stateProvince="Madang">No.</location>
141 / NEGATIVE [p in green] (
<collectionCode id="FEAED8F72746FFC5FAC5FF4A4C0607DD" box="[1288,1374,1129,1155]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="60" pageNumber="61">NMPC</collectionCode>
);
<specimenCount id="8EB98BBB2746FFC5FABFFF4A4CCF07DD" box="[1394,1431,1129,1155]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
,
<specimenCount id="8EB98BBB2746FFC5FF5AFFAF49E407F8" box="[151,188,1164,1190]" count="2" pageId="60" pageNumber="61" type="female">2♀</specimenCount>
, PAPUA NG,
<collectingRegion id="5A7B8ED02746FFC5FEABFFAF489D07F8" box="[358,453,1164,1190]" country="Papua New Guinea" name="Madang" pageId="60" pageNumber="61">Madang</collectingRegion>
, NAGADA /
<location id="9D6016E92746FFC5FDABFFAF4AC907F8" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FDABFFAF4AC907F8" box="[614,913,1164,1190]" country="Papua New Guinea" county="Binatang Research Center" latitude="-5.1563888" longLatPrecision="21" longitude="145.79472" municipality="Department of Parasitology" name="Binatang Research Center" pageId="60" pageNumber="61" stateProvince="Madang">Binatang Research Center</location>
/
<geoCoordinate id="FD8B26F52746FFC5FC61FFAD4D4907F7" box="[940,1041,1166,1193]" degrees="5" direction="south" minutes="9" orientation="latitude" pageId="60" pageNumber="61" precision="15" seconds="23" value="-5.1563888">5°923″S</geoCoordinate>
<geoCoordinate id="FD8B26F52746FFC5FBD7FFAD4DF607F7" box="[1050,1198,1166,1193]" degrees="145" direction="east" minutes="47" orientation="longitude" pageId="60" pageNumber="61" precision="15" seconds="41" value="145.79472">145°4741″E</geoCoordinate>
,
<quantity id="5F47EDD72746FFC5FB76FFAF4DAF07F9" box="[1211,1271,1164,1191]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="60" pageNumber="61" unit="m" value="20.0">
<elevation id="1392A7012746FFC5FB76FFAF4DAF07F9" box="[1211,1271,1164,1191]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="60" pageNumber="61" unit="m" value="20.0">20 m</elevation>
</quantity>
/ gardens and ruderal vegetation /
<date id="EC0166F22746FFC5FEB8FF93488F0794" box="[373,471,1200,1226]" pageId="60" pageNumber="61" value="2011-05-09">
<collectingDate id="FC459F1A2746FFC5FEB8FF93488F0794" box="[373,471,1200,1226]" pageId="60" pageNumber="61" value="2011-05-09">9.v.2011</collectingDate>
</date>
, Votýpka &amp; Lukeš lgt. [p] //
<location id="9D6016E92746FFC5FCDFFF934D1E0794" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FCDFFF934D1E0794" box="[786,1094,1200,1226]" country="Papua New Guinea" county="Binatang Research Center" latitude="-5.1563888" longLatPrecision="21" longitude="145.79472" municipality="Department of Parasitology" name="Department of Parasitology" pageId="60" pageNumber="61" stateProvince="Madang">Department of Parasitology</location>
/
<location id="9D6016E92746FFC5FB97FF934CC40794" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FB97FF934CC40794" box="[1114,1436,1200,1226]" country="Papua New Guinea" county="Binatang Research Center" latitude="-5.1563888" longLatPrecision="21" longitude="145.79472" municipality="Department of Parasitology" name="Charles University in Prague" pageId="60" pageNumber="61" stateProvince="Madang">Charles University in Prague</location>
/ DISSECTION No. 442, 443, 448 [respectively] / NEGATIVE [p in green] (
<collectionCode id="FEAED8F72746FFC5FBCFFFF04D0007B3" box="[1026,1112,1235,1261]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="60" pageNumber="61">NMPC</collectionCode>
). All
<specimenCount id="8EB98BBB2746FFC5FB5BFFF04C1907B3" box="[1174,1345,1235,1261]" count="4" pageId="60" pageNumber="61" type="generic">four specimens</specimenCount>
pinned, more or less damaged due to the dissection of the abdomen, detached body parts glued on separate piece of card pinned under the particular specimen.—
<specimenCount id="8EB98BBB2746FFC5FDBDFE394BDF066A" box="[624,647,1306,1332]" count="1" pageId="60" pageNumber="61" type="female">1♀</specimenCount>
, “PAPUA NEW GUINEA. / Imino (S
<geoCoordinate id="FD8B26F52746FFC5FBF6FE394DD2066A" box="[1083,1162,1306,1332]" degrees="2" direction="east" minutes="37" orientation="longitude" pageId="60" pageNumber="61" precision="925" value="2.6166668">2 37 E</geoCoordinate>
140 10)
<date id="EC0166F22746FFC5FB27FE394CD3066B" box="[1258,1419,1306,1333]" pageId="60" pageNumber="61" value="2009-01-25" valueMax="2009-01-31" valueMin="2009-01-25">
<collectingDate id="FC459F1A2746FFC5FB27FE394CD3066B" box="[1258,1419,1306,1333]" pageId="60" pageNumber="61" value="2009-01-25" valueMax="2009-01-31" valueMin="2009-01-25">25.I-31.I.2009</collectingDate>
</date>
/
<collectorName id="354A25E42746FFC5FF5AFE1D487A0606" box="[151,290,1342,1368]" pageId="60" pageNumber="61">A. Zamesov</collectorName>
lgt. // Roca-Cusachs Personal Collection [p] //
<typeStatus id="4704FE902746FFC5FC8FFE1D4AB80606" box="[834,992,1342,1368]" pageId="60" pageNumber="61" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="5FBF3BB12746FFC5FC3AFE1D4C190606" authorityName="Salini &amp; Kment" authorityYear="2025" box="[1015,1345,1342,1368]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="66" phylum="Arthropoda" rank="species" species="votypkai">AXIAGASTUS VOTYPKAI</taxonomicName>
/ Salini &amp; Kment
<taxonomicNameLabel id="B1F8215B2746FFC5FEC3FE4248300625" box="[270,360,1377,1403]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
// det.
<bibRefCitation id="FC2E3DC32746FFC5FE72FE424BF90625" author="Roca-Cusachs, M. &amp; Rider, D. A. &amp; Kment, P. &amp; Jung, S. - H." box="[447,673,1377,1403]" pageId="60" pageNumber="61" pagination="419 - 434" refId="ref54561" refString="Roca-Cusachs, M., Rider, D. A., Kment, P. &amp; Jung, S. - H. (2024) Revision of Vitruvius (Hemiptera: Heteroptera: Pentatomidae) with the description of a new species from Cambodia and Thailand. Zootaxa, 5486 (3), 419-434. https://doi.org/10.11646/zootaxa.5486.3.5" type="journal article" year="2024">Roca-Cusachs 2024</bibRefCitation>
[red label]” (
<collectionCode id="FEAED8F72746FFC5FC8FFE424AC50625" box="[834,925,1377,1403]" pageId="60" pageNumber="61">MRCC</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="28D74A6F2746FFC5FC04FE424863069C" collectionCode="ISNB" collectorName="Coll. I. R. Sc." country="Papua New Guinea" location="Madang" pageId="60" pageNumber="61" specimenCount="2" specimenCount-male="2" stateProvince="Madang" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FC04FE424ABE0625" box="[969,998,1377,1403]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
, “
<collectorName id="354A25E42746FFC5FC32FE424DC80625" box="[1023,1168,1377,1403]" pageId="60" pageNumber="61">Coll. I.R.Sc.</collectorName>
N.B. / Canopy Mission Papua / New Guinea (
<collectingRegion id="5A7B8ED02746FFC5FE5DFEA748B606C0" box="[400,494,1412,1438]" country="Papua New Guinea" name="Madang" pageId="60" pageNumber="61">Madang</collectingRegion>
/ prov.): Baiteta
<date id="EC0166F22746FFC5FD60FEA74A6F06C1" box="[685,823,1412,1439]" pageId="60" pageNumber="61" value="1995-05-25">25.V. / 1995</date>
Fogging AR8 / Leg. Olivier Missa [p, orange label] //
<specimenCount id="8EB98BBB2746FFC5FF5AFE8B49F4069C" box="[151,172,1448,1474]" count="1" pageId="60" pageNumber="61" type="male"></specimenCount>
[p]” (
<collectionCode id="FEAED8F72746FFC5FF3DFE8B486B069C" box="[240,307,1448,1474]" pageId="60" pageNumber="61">ISNB</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="28D74A6F2746FFC5FE93FE8B4A320557" collectingDate="1968-04-30" collectionCode="BMNH" collectorName="B. Gray &amp; M. S. K. Ghauri" country="Papua New Guinea" location="Bulolo" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Morobe" typeStatus="paratype">
<emphasis id="AACB9C202746FFC5FE93FE8B489D069C" bold="true" box="[350,453,1448,1474]" pageId="60" pageNumber="61">
<collectingRegion id="5A7B8ED02746FFC5FE93FE8B4899069C" box="[350,449,1448,1474]" country="Papua New Guinea" name="Morobe" pageId="60" pageNumber="61">Morobe</collectingRegion>
:
</emphasis>
<specimenCount id="8EB98BBB2746FFC5FE00FE8B48A9069C" box="[461,497,1448,1474]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
, “attracted to house [hw] / light 9.00 PM,
<location id="9D6016E92746FFC5FC03FE8B4D42069C" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FC03FE8B4D42069C" box="[974,1050,1448,1474]" country="Papua New Guinea" name="Bulolo" pageId="60" pageNumber="61" stateProvince="Morobe">Bulolo</location>
[hw] /
<collectorName id="354A25E42746FFC5FBA0FE8B4D9F069C" box="[1133,1223,1448,1474]" pageId="60" pageNumber="61">B. Gray</collectorName>
,
<date id="EC0166F22746FFC5FB1EFE8B4C11069C" box="[1235,1353,1448,1474]" pageId="60" pageNumber="61" value="1968-04-30">
<collectingDate id="FC459F1A2746FFC5FB1EFE8B4C11069C" box="[1235,1353,1448,1474]" pageId="60" pageNumber="61" value="1968-04-30">30.iv.1968</collectingDate>
</date>
[hw] // 279 [hw] // 141 [hw] // C. I.E Coll. [p] / A. [p] 5556 [hw] //
<taxonomicName id="5FBF3BB12746FFC5FC84FEEF4A9C06B8" authorityName="Dallas" authorityYear="1851" box="[841,964,1484,1510]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="AACB9C202746FFC5FC84FEEF4A9C06B8" box="[841,964,1484,1510]" italics="true" pageId="60" pageNumber="61">Axiagastus</emphasis>
</taxonomicName>
[hw] / sp. [hw] /
<collectorName id="354A25E42746FFC5FB46FEE84C6E06BB" box="[1163,1334,1483,1509]" pageId="60" pageNumber="61">M.S.K. Ghauri</collectorName>
, det. 197 [p] 3 [hw] // dissected and illustrated by Salini. S. [p]” (
<collectionCode id="FEAED8F72746FFC5FCC5FECC4A3A0557" box="[776,866,1519,1545]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
. The dissected male genitalia are placed in a glass microvial with glycerol attached to the same pin, right antennomeres III and IV and left protarsomeres II and III lost, both hemelytra along with hindwings dislocated and spread laterally, exposing the dorsum of the abdomen, beyond the scutellum.—
<materialsCitation id="28D74A6F2746FFC5FE78FD7A486B05E4" collectionCode="BMNH" collectorName="Coll. J H Ardley" country="Papua New Guinea" determinerName="M. S. K. Ghauri" location="By Light Trapping." municipality="Bubia." pageId="60" pageNumber="61" specimenCount="2" specimenCount-male="2" stateProvince="Morobe" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FE78FD7A4888052D" box="[437,464,1625,1651]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
, “
<collectingMunicipality id="7864DA482746FFC5FE2AFD7A4B6B052D" box="[487,563,1625,1651]" pageId="60" pageNumber="61">Bubia.</collectingMunicipality>
LAE.NG. [p] / 23/5/61. /
<location id="9D6016E92746FFC5FC99FD794D76052D" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FC99FD794D76052D" box="[852,1070,1625,1652]" country="Papua New Guinea" municipality="Bubia." name="By Light Trapping." pageId="60" pageNumber="61" stateProvince="Morobe">By Light Trapping.</location>
[hw] /
<collectorName id="354A25E42746FFC5FB4CFD7A4C63052D" box="[1153,1339,1625,1652]" pageId="60" pageNumber="61">Coll. J H Ardley</collectorName>
[p] // B. 1315 [hw, pink label] // C.I.E. COLL. / NO. [p] 17721 [hw] //
<taxonomicName id="5FBF3BB12746FFC5FC90FD5E4A8105C9" authorityName="Dallas" authorityYear="1851" box="[861,985,1661,1687]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">Axiagastus</taxonomicName>
/ sp. [hw] /
<determinerName id="6DDCACF62746FFC5FBABFD5E4C5605C9" box="[1126,1294,1661,1687]" pageId="60" pageNumber="61">M.S.K. Ghauri</determinerName>
det. [p] //
<specimenCount id="8EB98BBB2746FFC5FA4AFD5E4CC405C9" box="[1415,1436,1661,1687]" count="1" pageId="60" pageNumber="61" type="male"></specimenCount>
[p]” (
<collectionCode id="FEAED8F72746FFC5FF1EFD83487305E4" box="[211,299,1696,1722]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
.—
<emphasis id="AACB9C202746FFC5FE98FD8348DF05E4" bold="true" box="[341,391,1696,1722]" pageId="60" pageNumber="61">
<materialsCitation id="28D74A6F2746FFC5FE98FD8348DB05E4" box="[341,387,1696,1722]" country="Papua New Guinea" location="Ori" pageId="60" pageNumber="61" specimenCount="1" stateProvince="Morobe" typeStatus="paratype">
<location id="9D6016E92746FFC5FE98FD8348DB05E4" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FE98FD8348DB05E4" box="[341,387,1696,1722]" country="Papua New Guinea" name="Ori" pageId="60" pageNumber="61" stateProvince="Morobe">Ori</location>
</materialsCitation>
:
</emphasis>
<materialsCitation id="28D74A6F2746FFC5FE43FD8348A70583" collectingDate="1933-09" country="Papua New Guinea" elevation="366" location="Kokoda." pageId="60" pageNumber="61" specimenCount="2" specimenCount-male="2" stateProvince="West New Britain" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FE43FD8348EC05E4" box="[398,436,1696,1722]" count="2" pageId="60" pageNumber="61" type="male">2♂</specimenCount>
, “PAPUA:
<location id="9D6016E92746FFC5FDFEFD834BCB05E4" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FDFEFD834BCB05E4" box="[563,659,1696,1722]" country="Papua New Guinea" name="Kokoda." pageId="60" pageNumber="61" stateProvince="West New Britain">Kokoda.</location>
/
<quantity id="5F47EDD72746FFC5FD65FD834A5E05E4" box="[680,774,1696,1722]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="60" pageNumber="61" unit="ft" value="1200.0">
<elevation id="1392A7012746FFC5FD65FD834A5E05E4" box="[680,774,1696,1722]" metricMagnitude="2" metricUnit="m" metricValue="3.6576" pageId="60" pageNumber="61" unit="ft" value="1200.0">1.200 ft.</elevation>
</quantity>
<date id="EC0166F22746FFC5FCC1FD834A3C05E4" box="[780,868,1696,1722]" pageId="60" pageNumber="61" value="1933-09">
<collectingDate id="FC459F1A2746FFC5FCC1FD834A3C05E4" box="[780,868,1696,1722]" pageId="60" pageNumber="61" value="1933-09">ix.1933</collectingDate>
</date>
. / L. E. Cheesman. / B.M. 1934-321 [p] / NHMUK 013589094 [p]”—
<collectingRegion id="5A7B8ED02746FFC5FEAAFDE748A70583" box="[359,511,1731,1758]" country="Papua New Guinea" name="West New Britain" pageId="60" pageNumber="61">
<emphasis id="AACB9C202746FFC5FEAAFDE748A70583" bold="true" box="[359,511,1731,1758]" pageId="60" pageNumber="61">New Britain</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="AACB9C202746FFC5FE32FDE04B5C0583" bold="true" box="[511,516,1731,1757]" pageId="60" pageNumber="61">:</emphasis>
<materialsCitation id="28D74A6F2746FFC5FDC0FDE748590416" collectionCode="BMNH" collectorName="Coll. Dept. &amp; G. M. Baloch &amp; Ex. Cluster &amp; M. S. K. Ghauri" country="Papua New Guinea" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="West New Britain" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FDC0FDE74B6A0580" box="[525,562,1732,1758]" count="1" pageId="60" pageNumber="61" type="male">1♂</specimenCount>
, “L.A.E.S. [hw] / KERAVAT [hw] / 5.iii.70 [hw] // K.790 [hw] //. Ex.
<collectorName id="354A25E42746FFC5FAA8FDE7498D045C" pageId="60" pageNumber="61">Coll. Dept.</collectorName>
[p] / Agr. P. M. [p] / No.: [p] 13326 [hw] //
<collectorName id="354A25E42746FFC5FD08FDC44A06045F" box="[709,862,1767,1794]" pageId="60" pageNumber="61">G. M. Baloch</collectorName>
[hw] // C. I. E COLL. [p] / A. [p] 3841 [hw] //
<collectorName id="354A25E42746FFC5FABBFDCB49B0047A" pageId="60" pageNumber="61">Ex. Cluster</collectorName>
palm [hw] /?
<taxonomicName id="5FBF3BB12746FFC5FE40FC294B0E047B" box="[397,598,1802,1829]" class="Liliopsida" family="Arecaceae" genus="Ptychosperma" kingdom="Plantae" order="Arecales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="AACB9C202746FFC5FE40FC294B75047A" box="[397,557,1802,1828]" italics="true" pageId="60" pageNumber="61">Ptychosperma</emphasis>
sp.
</taxonomicName>
[hw] //
<taxonomicName id="5FBF3BB12746FFC5FD79FC284A77047B" authorityName="Dallas" authorityYear="1851" box="[692,815,1803,1829]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="AACB9C202746FFC5FD79FC284A77047B" box="[692,815,1803,1829]" italics="true" pageId="60" pageNumber="61">Axiagastus</emphasis>
</taxonomicName>
[hw]
<emphasis id="AACB9C202746FFC5FCBBFC284A26047B" box="[886,894,1803,1829]" italics="true" pageId="60" pageNumber="61">/</emphasis>
sp. [hw] /
<collectorName id="354A25E42746FFC5FC33FC284DF1047A" box="[1022,1193,1802,1829]" pageId="60" pageNumber="61">M.S.K. Ghauri</collectorName>
, det. 197 [p] 0 [hw]” (
<collectionCode id="FEAED8F72746FFC5FF52FC0D49A10416" box="[159,249,1838,1864]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
. Both right pro- and mesotarsus lost, both antennae except antennomere I, all left legs except coxae and trochanters, and right hind leg except coxa lost, left hemelytron glued to card.—
<materialsCitation id="28D74A6F2746FFC5FBFAFC724A5C04EC" collectingDate="1940-09-03" collectionCode="BMNH" country="Papua New Guinea" determinerName="W. E. China" location="Bainings" pageId="60" pageNumber="61" specimenCount="2" specimenCount-female="2" stateProvince="West New Britain" typeStatus="paratype">
<specimenCount id="8EB98BBB2746FFC5FBFAFC724D080435" box="[1079,1104,1873,1899]" count="1" pageId="60" pageNumber="61" type="female">1♀</specimenCount>
, “
<collectingRegion id="5A7B8ED02746FFC5FBABFC714C780432" box="[1126,1312,1874,1900]" country="Papua New Guinea" name="West New Britain" pageId="60" pageNumber="61">NEW BRITAIN</collectingRegion>
/
<location id="9D6016E92746FFC5FAF4FC724CC50435" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FAF4FC724CC50435" box="[1337,1437,1873,1899]" country="Papua New Guinea" name="Bainings" pageId="60" pageNumber="61" stateProvince="West New Britain">Bainings</location>
/
<date id="EC0166F22746FFC5FF6AFC56487804D1" box="[167,288,1909,1935]" pageId="60" pageNumber="61" value="1940-09-03">
<collectingDate id="FC459F1A2746FFC5FF6AFC56487804D1" box="[167,288,1909,1935]" pageId="60" pageNumber="61" value="1940-09-03">3. ix. 1940</collectingDate>
</date>
/ J. L. Froggatt. / with “borers” / in cocoa stem [sic, hw] // H.545 [hw] //
<taxonomicName id="5FBF3BB12746FFC5FBAEFC564D8704D1" authorityName="Dallas" authorityYear="1851" box="[1123,1247,1909,1935]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">Axiagastus</taxonomicName>
/
<taxonomicName id="5FBF3BB12746FFC5FB3BFC564CC504D1" authority="Dall." authorityName="Dallas" authorityYear="1851" box="[1270,1437,1909,1935]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="rosmarus">rosmarus Dall.</taxonomicName>
[hw] / det.
<determinerName id="6DDCACF62746FFC5FEDAFCBB48C904EC" box="[279,401,1944,1970]" pageId="60" pageNumber="61">W.E.China</determinerName>
194[p]1[hw] //
<specimenCount id="8EB98BBB2746FFC5FD84FCBA4B0604ED" box="[585,606,1945,1971]" count="1" pageId="60" pageNumber="61" type="female"></specimenCount>
[p]” (
<collectionCode id="FEAED8F72746FFC5FD68FCBB4BA404EC" box="[677,764,1944,1970]" pageId="60" pageNumber="61">BMNH</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="28D74A6F2746FFC5FCE8FCBB4CC104EC" box="[805,1433,1944,1970]" country="Papua New Guinea" location="Each" pageId="60" pageNumber="61" specimenCount="1" stateProvince="West New Britain" typeStatus="paratype">
<location id="9D6016E92746FFC5FCE8FCBB4A0704EC" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92746FFC5FCE8FCBB4A0704EC" box="[805,863,1944,1970]" country="Papua New Guinea" name="Each" pageId="60" pageNumber="61" stateProvince="West New Britain">Each</location>
specimen bears the following identification label
</materialsCitation>
: “
<typeStatus id="4704FE902746FFC5FF6CFC9E481A0489" box="[161,322,1981,2007]" pageId="60" pageNumber="61" type="paratype">PARATYPUS</typeStatus>
/
<taxonomicName id="5FBF3BB12746FFC5FE9BFC9E4BD90489" authorityName="Salini &amp; Kment" authorityYear="2025" box="[342,641,1981,2007]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="60" pageNumber="66" phylum="Arthropoda" rank="species" species="votypkai">
<emphasis id="AACB9C202746FFC5FE9BFC9E4BD90489" box="[342,641,1981,2007]" italics="true" pageId="60" pageNumber="61">AXIAGASTUS VOTYPKAI</emphasis>
</taxonomicName>
/ Salini &amp; Kment
<taxonomicNameLabel id="B1F8215B2746FFC5FC80FC9E4AFC0489" box="[845,932,1981,2007]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
/ det. Salini, S. &amp; Kment, P., 2024 [p, yellow label], unless stated otherwise.
</paragraph>
<paragraph id="980040322747FFC4FF0AFBB548FD0242" blockId="61.[151,1437,150,2043]" pageId="61" pageNumber="62">
<emphasis id="AACB9C202747FFC4FF0AFBB54B0003EE" bold="true" box="[199,600,150,176]" pageId="61" pageNumber="62">Additional material examined.—</emphasis>
<materialsCitation id="28D74A6F2747FFC4FD95FBB548F90242" collectingDate="1969-01-16" collectionCode="BMNH" collectorName="N. Gough" country="Papua New Guinea" location="On" pageId="61" pageNumber="62" specimenCount="2" specimenCount-female="2" stateProvince="East Sepik">
<emphasis id="AACB9C202747FFC4FD95FBB54D7003EE" bold="true" box="[600,1064,150,177]" pageId="61" pageNumber="62">
<collectingCountry id="E0A800A22747FFC4FD95FBB54ADF03EE" box="[600,903,150,177]" name="Papua New Guinea" pageId="61" pageNumber="62">PAPUA NEW GUINEA</collectingCountry>
:
<collectingRegion id="5A7B8ED02747FFC4FC57FBB44D7B03EE" box="[922,1059,150,177]" country="Papua New Guinea" name="East Sepik" pageId="61" pageNumber="62">East Sepik</collectingRegion>
:
</emphasis>
<specimenCount id="8EB98BBB2747FFC4FBF8FBB44D0003EF" box="[1077,1112,151,177]" count="1" pageId="61" pageNumber="62" type="female">1♀</specimenCount>
, “
<location id="9D6016E92747FFC4FBBBFBB44DC203EF" LSID="urn:lsid:plazi:treatment:1016F1242746FFB9FF5AFBB44B4404A4:9D6016E92747FFC4FBBBFBB44DC203EF" box="[1142,1178,151,177]" country="Papua New Guinea" name="On" pageId="61" pageNumber="62" stateProvince="East Sepik">On</location>
leaf
<taxonomicName id="5FBF3BB12747FFC4FB13FBB44CD003EF" box="[1246,1416,151,177]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="61" pageNumber="62" phylum="Tracheophyta" rank="species" species="undetermined">Eucalyptus sp.</taxonomicName>
/ Kinjingini, E. S. D. /
<date id="EC0166F22747FFC4FE49FB9948A0038A" box="[388,504,186,212]" pageId="61" pageNumber="62" value="1969-01-16">
<collectingDate id="FC459F1A2747FFC4FE49FB9948A0038A" box="[388,504,186,212]" pageId="61" pageNumber="62" value="1969-01-16">16.1.1969</collectingDate>
</date>
.
<collectorName id="354A25E42747FFC4FDCFFB984B28038A" box="[514,624,186,213]" pageId="61" pageNumber="62">N. Gough</collectorName>
[hw] // 603 [hw] // 171 [hw] // C.I.E. COLL. / A. [p] 5556 [hw] //
<specimenCount id="8EB98BBB2747FFC4FA9EFB984C30038B" box="[1363,1384,187,213]" count="1" pageId="61" pageNumber="62" type="female"></specimenCount>
[p]” (
<collectionCode id="FEAED8F72747FFC4FF52FBFD49A103A6" box="[159,249,222,248]" pageId="61" pageNumber="62">BMNH</collectionCode>
). Due to its small size, slightly aberrant colouration and lack of associated males the identification of this female is only tentative
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92747FFB9FF0AFA054C350667" lastPageId="64" lastPageNumber="65" pageId="61" pageNumber="62" type="description">
<paragraph id="980040322747FFC4FF0AFA054DA60656" blockId="61.[151,1437,150,2043]" pageId="61" pageNumber="62">
<emphasis id="AACB9C202747FFC4FF0AFA054B8B021E" bold="true" box="[199,723,294,321]" pageId="61" pageNumber="62">
Description.
<emphasis id="AACB9C202747FFC4FE93FA054B8B021E" bold="true" box="[350,723,294,321]" italics="true" pageId="61" pageNumber="62">Colour, integument and vestiture</emphasis>
</emphasis>
(
<figureCitation id="00845CB72747FFC4FD2DFA054A26021E" box="[736,894,294,320]" captionStart="FIGURES 200206" captionStartId="62.[151,264,1859,1883]" captionTargetBox="[151,1436,201,1835]" captionTargetId="figure-29@62.[151,1436,201,1835]" captionTargetPageId="62" captionText="FIGURES 200206. Axiagastus votypkai Salini &amp; Kment sp. nov. 200201—habitus (200—dorsal view, 201—ventral view), 202203—head (202—dorsal view, 203—lateral view); 204—external scent efferent system; 205—apex of male abdomen with genital capsule in postero-ventral view; 206—apex of female abdomen with terminalia in posteroventral view. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 1 mm (200201), 0.5 mm (202206). Orig. S. Salini." pageId="61" pageNumber="62">Figs 200204</figureCitation>
). Ground colour stramineous, but body appearing black due to dense, coarse black punctation confluent on various parts of body (especially on scutellum and coria), forming extensive dark area intermixed with impunctate bright yellow areas; disc of head bright yellow, bordered laterally with black outline, basal half of disc with coarse black punctures, gradually becoming finer towards apex, more or less appearing as a longitudinal row of punctures on middle of mandibular plates, lateral margins of clypeus bordered by black lines; ventral surface of head, including denticles on bucculae (except black apices) bright yellow; each scape (I) and both pedicellites (IIa, IIb) stramineous (IIb sometimes darker on apical half), flagellomeres (III and IV) brown to dark brown, basally paler; labium concolourous with ventral body surface except for black apex of labiomere IV; disc of pronotum bright yellow except for black anterolateral margins, and narrow to thick, black transverse band along posterior margins sometimes extending to humeri, sometimes interrupted with isolated spot on humeri; coarse, black punctation scattered uniformly over rest of yellowish disc of pronotum; ground colour of scutellar disc bright yellow with distinct black markings as follows: 1+1 small to moderately sized, roughly round spots submedially on basal margin of scutellum, and 1+1 large, roughly quadrate spots at frenal regions of scutellar disc, and a large, transverse, roughly crescent-shaped anteapical spot (somewhat nebulous in
<specimenCount id="8EB98BBB2747FFC4FB64F9DA4C1F004C" box="[1193,1351,760,787]" count="1" pageId="61" pageNumber="62" type="generic">one specimen</specimenCount>
); bright yellow regions of scutellar disc scattered with black punctation; apex of scutellum with pale impunctate spot, with anterior margin medially incised, appearing similar to letter M; endocorium with dense, coarse, black punctation, confluent into two poorly delimited dark spots (anteriorly and posterocentrally) alternating with two paler spots (anterocentrally and posteriorly), a pale callose spot more or less apparent laterally in middle of each endocorium; connexiva pale yellow except black anterior and posterior third of each segment; fine, dense, concolourous punctures on connexiva. Ventral surface of body including legs and labium pale yellow, scattered with coarse, brown punctures confluent, mostly forming transverse broken rows especially on thorax and lateral region of abdomen; punctures more concentrated towards lateral regions of thorax and ventrites; legs with punctures round, coarse, brown, sparse on tibiae and femora; head on ventral surface with two narrow, longitudinal stripes on either side of bucculae, from their apices to base of head, black; one roughly round, moderately-sized spot lateral to base of pro-, meso- and metacoxae, one spot each on middle of lateral margins of metapleura, apex of ostiolar peritremes and antero- and posterolateral angles of ventrites IIIVII, and one longitudinal, median stripe on ventrites VVII, black; apices of buccular denticles, labium and tarsal claws, a sublateral row of small spots mesad to each spiracle coalescing with spiracular outline, black; ventrites with intersegmental sutures marked with brown, transverse stripe.
</paragraph>
<paragraph id="980040322747FFC4FF0AFE314A70060E" blockId="61.[151,1437,150,2043]" pageId="61" pageNumber="62">
<emphasis id="AACB9C202747FFC4FF0AFE3148630672" bold="true" box="[199,315,1298,1324]" italics="true" pageId="61" pageNumber="62">Structure.</emphasis>
Labium reaching or surpassing posterior margin of ventrite III, sometimes reaching posterior margin of ventrite IV. Other characters as in generic redescription.
</paragraph>
<paragraph id="980040322747FFC4FF0AFE794B8D0418" blockId="61.[151,1437,150,2043]" pageId="61" pageNumber="62">
<emphasis id="AACB9C202747FFC4FF0AFE794828062B" bold="true" box="[199,368,1370,1397]" italics="true" pageId="61" pageNumber="62">Male genitalia</emphasis>
(
<figureCitation id="00845CB72747FFC4FE4DFE7948B0062B" box="[384,488,1370,1397]" captionStart="FIGURES 200206" captionStartId="62.[151,264,1859,1883]" captionTargetBox="[151,1436,201,1835]" captionTargetId="figure-29@62.[151,1436,201,1835]" captionTargetPageId="62" captionText="FIGURES 200206. Axiagastus votypkai Salini &amp; Kment sp. nov. 200201—habitus (200—dorsal view, 201—ventral view), 202203—head (202—dorsal view, 203—lateral view); 204—external scent efferent system; 205—apex of male abdomen with genital capsule in postero-ventral view; 206—apex of female abdomen with terminalia in posteroventral view. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 1 mm (200201), 0.5 mm (202206). Orig. S. Salini." pageId="61" pageNumber="62">Figs 205</figureCitation>
,
<figureCitation id="00845CB72747FFC4FE39FE794B03062A" box="[500,603,1370,1396]" captionStart-0="FIGURES 207213" captionStart-1="FIGURES 214219" captionStart-2="FIGURES 220225" captionStartId-0="63.[151,264,1909,1933]" captionStartId-1="64.[151,264,1112,1136]" captionStartId-2="65.[151,264,1107,1131]" captionTargetBox-0="[192,1396,181,1885]" captionTargetBox-1="[151,1436,181,1089]" captionTargetBox-2="[151,1436,181,1083]" captionTargetId-0="figure-19@63.[192,1396,181,1885]" captionTargetId-1="figure-29@64.[151,1436,181,1089]" captionTargetId-2="figure-19@65.[151,1436,181,1083]" captionTargetPageId-0="63" captionTargetPageId-1="64" captionTargetPageId-2="65" captionText-0="FIGURES 207213. Male genitalia (ESEM micrographs) of Axiagastus votypkai Salini &amp; Kment sp. nov. 207210—genital capsule (207—dorsal view, 32×; 208—ventral view, 32×; 209—caudal view, 40×; 210—lateral view, 45×); 211213—parameres (three different views, 100×). Lettering: dn—distension, te—transverse emargination. Scale: 1 mm (207210), 0.5 mm (211 213). Orig. P. Kment." captionText-1="FIGURES 214219. Axiagastus votypkai Salini &amp; Kment sp. nov., male genitalia. 214217—genital capsule (214—dorsal view, 215—ventral view, 216—caudal view, 217—lateral view); 218219—parameres (two different views). Lettering: dm— dorsal margin, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, te—transverse emargination. Scale: 0.5 mm (214217), 0.25 mm (218219). Orig. S. Salini." captionText-2="FIGURES 220225. Axiagastus votypkai Salini &amp; Kment sp. nov., male and female genitalia. 220222—phallus (220—dorsal view, 221—ventral view, 222—lateral view); 223—terminalia (dorsal view); 224—terminalia (ventral view); 225—spermatheca. Scale: 0.1 mm (220222, 225), 0.5 mm (223224). Orig. S. Salini." pageId="61" pageNumber="62">207222</figureCitation>
). Genital capsule (
<figureCitation id="00845CB72747FFC4FCE2FE794A95062A" box="[815,973,1370,1396]" captionStart="FIGURES 207213" captionStartId="63.[151,264,1909,1933]" captionTargetBox="[192,1396,181,1885]" captionTargetId="figure-19@63.[192,1396,181,1885]" captionTargetPageId="63" captionText="FIGURES 207213. Male genitalia (ESEM micrographs) of Axiagastus votypkai Salini &amp; Kment sp. nov. 207210—genital capsule (207—dorsal view, 32×; 208—ventral view, 32×; 209—caudal view, 40×; 210—lateral view, 45×); 211213—parameres (three different views, 100×). Lettering: dn—distension, te—transverse emargination. Scale: 1 mm (207210), 0.5 mm (211 213). Orig. P. Kment." pageId="61" pageNumber="62">Figs 207210</figureCitation>
,
<figureCitation id="00845CB72747FFC4FC16FE794D66062A" box="[987,1086,1370,1396]" captionStart="FIGURES 214219" captionStartId="64.[151,264,1112,1136]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-29@64.[151,1436,181,1089]" captionTargetPageId="64" captionText="FIGURES 214219. Axiagastus votypkai Salini &amp; Kment sp. nov., male genitalia. 214217—genital capsule (214—dorsal view, 215—ventral view, 216—caudal view, 217—lateral view); 218219—parameres (two different views). Lettering: dm— dorsal margin, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, te—transverse emargination. Scale: 0.5 mm (214217), 0.25 mm (218219). Orig. S. Salini." pageId="61" pageNumber="62">214217</figureCitation>
) subquadrate; dorsal rim more deeply incised than ventral rim; lateral wall of dorsal rim slightly concave, smooth, sclerotized black, ending in moderately developed denticle contiguous with moderately deep, transverse emargination (te) at middle of dorsal rim; dorsal sinus of posterior aperture broadly U-shaped (ds), ventral margin of posterior aperture semicircular; ventral rim including posterolateral angles bordered with a row of moderately elongate, golden setae; posterolateral lobes in lateral view as shown in
<figureCitation id="00845CB72747FFC4FDDFFD2E4B2F0576" box="[530,631,1549,1576]" captionStart="FIGURES 207213" captionStartId="63.[151,264,1909,1933]" captionTargetBox="[192,1396,181,1885]" captionTargetId="figure-19@63.[192,1396,181,1885]" captionTargetPageId="63" captionText="FIGURES 207213. Male genitalia (ESEM micrographs) of Axiagastus votypkai Salini &amp; Kment sp. nov. 207210—genital capsule (207—dorsal view, 32×; 208—ventral view, 32×; 209—caudal view, 40×; 210—lateral view, 45×); 211213—parameres (three different views, 100×). Lettering: dn—distension, te—transverse emargination. Scale: 1 mm (207210), 0.5 mm (211 213). Orig. P. Kment." pageId="61" pageNumber="62">Figs 210</figureCitation>
,
<figureCitation id="00845CB72747FFC4FD4EFD2D4BE80576" box="[643,688,1550,1576]" captionStart="FIGURES 214219" captionStartId="64.[151,264,1112,1136]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-29@64.[151,1436,181,1089]" captionTargetPageId="64" captionText="FIGURES 214219. Axiagastus votypkai Salini &amp; Kment sp. nov., male genitalia. 214217—genital capsule (214—dorsal view, 215—ventral view, 216—caudal view, 217—lateral view); 218219—parameres (two different views). Lettering: dm— dorsal margin, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, te—transverse emargination. Scale: 0.5 mm (214217), 0.25 mm (218219). Orig. S. Salini." pageId="61" pageNumber="62">217</figureCitation>
; ventral rim broadly V-shaped at middle, infoldings of ventral rim deeply impressed on either side of moderately developed distension (dn) at middle, distension (dn) emarginated with short V-shaped incision at middle, visible on ventral view of genital capsule; infoldings of ventral rim laterally (inner to posterolateral lobes) and lateral to base of median distension with a prominent, apically acuminate, sclerotized, black denticle (dt).
<emphasis id="AACB9C202747FFC4FEB9FDBD48BB05E9" box="[372,483,1694,1719]" italics="true" pageId="61" pageNumber="62">Paramere</emphasis>
(
<figureCitation id="00845CB72747FFC4FE3FFDBE4BC805E9" box="[498,656,1693,1719]" captionStart="FIGURES 207213" captionStartId="63.[151,264,1909,1933]" captionTargetBox="[192,1396,181,1885]" captionTargetId="figure-19@63.[192,1396,181,1885]" captionTargetPageId="63" captionText="FIGURES 207213. Male genitalia (ESEM micrographs) of Axiagastus votypkai Salini &amp; Kment sp. nov. 207210—genital capsule (207—dorsal view, 32×; 208—ventral view, 32×; 209—caudal view, 40×; 210—lateral view, 45×); 211213—parameres (three different views, 100×). Lettering: dn—distension, te—transverse emargination. Scale: 1 mm (207210), 0.5 mm (211 213). Orig. P. Kment." pageId="61" pageNumber="62">Figs 211213</figureCitation>
,
<figureCitation id="00845CB72747FFC4FD51FDBE4A5905E9" box="[668,769,1693,1719]" captionStart="FIGURES 214219" captionStartId="64.[151,264,1112,1136]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-29@64.[151,1436,181,1089]" captionTargetPageId="64" captionText="FIGURES 214219. Axiagastus votypkai Salini &amp; Kment sp. nov., male genitalia. 214217—genital capsule (214—dorsal view, 215—ventral view, 216—caudal view, 217—lateral view); 218219—parameres (two different views). Lettering: dm— dorsal margin, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, te—transverse emargination. Scale: 0.5 mm (214217), 0.25 mm (218219). Orig. S. Salini." pageId="61" pageNumber="62">218, 219</figureCitation>
) simple, nearly as in
<taxonomicName id="5FBF3BB12747FFC4FC38FDBE4D2605E9" authorityName="Dallas" authorityYear="1851" box="[1013,1150,1693,1719]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C202747FFC4FC38FDBE4D2605E9" box="[1013,1150,1693,1719]" italics="true" pageId="61" pageNumber="62">A. rosmarus</emphasis>
</taxonomicName>
, numerous fine, elongate setae along periphery of laminate disc (ld); crown narrowed abruptly towards apex, acuminate in lateral view (
<figureCitation id="00845CB72747FFC4FAA7FDE249A305A1" captionStart="FIGURES 207213" captionStartId="63.[151,264,1909,1933]" captionTargetBox="[192,1396,181,1885]" captionTargetId="figure-19@63.[192,1396,181,1885]" captionTargetPageId="63" captionText="FIGURES 207213. Male genitalia (ESEM micrographs) of Axiagastus votypkai Salini &amp; Kment sp. nov. 207210—genital capsule (207—dorsal view, 32×; 208—ventral view, 32×; 209—caudal view, 40×; 210—lateral view, 45×); 211213—parameres (three different views, 100×). Lettering: dn—distension, te—transverse emargination. Scale: 1 mm (207210), 0.5 mm (211 213). Orig. P. Kment." pageId="61" pageNumber="62">Figs 211, 213</figureCitation>
,
<figureCitation id="00845CB72747FFC4FECBFDC6486C05A1" box="[262,308,1765,1791]" captionStart="FIGURES 214219" captionStartId="64.[151,264,1112,1136]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-29@64.[151,1436,181,1089]" captionTargetPageId="64" captionText="FIGURES 214219. Axiagastus votypkai Salini &amp; Kment sp. nov., male genitalia. 214217—genital capsule (214—dorsal view, 215—ventral view, 216—caudal view, 217—lateral view); 218219—parameres (two different views). Lettering: dm— dorsal margin, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, te—transverse emargination. Scale: 0.5 mm (214217), 0.25 mm (218219). Orig. S. Salini." pageId="61" pageNumber="62">218</figureCitation>
), dorsal margin (dm) of upright finger-like crown convex in lateral view (
<figureCitation id="00845CB72747FFC4FBA1FDC64D9605A1" box="[1132,1230,1765,1791]" captionStart="FIGURES 214219" captionStartId="64.[151,264,1112,1136]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-29@64.[151,1436,181,1089]" captionTargetPageId="64" captionText="FIGURES 214219. Axiagastus votypkai Salini &amp; Kment sp. nov., male genitalia. 214217—genital capsule (214—dorsal view, 215—ventral view, 216—caudal view, 217—lateral view); 218219—parameres (two different views). Lettering: dm— dorsal margin, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, te—transverse emargination. Scale: 0.5 mm (214217), 0.25 mm (218219). Orig. S. Salini." pageId="61" pageNumber="62">Fig. 218</figureCitation>
); stem moderately elongate, apodeme disc-like.
<emphasis id="AACB9C202747FFC4FE2EFC2B4B61047C" box="[483,569,1800,1826]" italics="true" pageId="61" pageNumber="62">Phallus</emphasis>
(
<figureCitation id="00845CB72747FFC4FD84FC2B4BB3047D" box="[585,747,1800,1827]" captionStart="FIGURES 220225" captionStartId="65.[151,264,1107,1131]" captionTargetBox="[151,1436,181,1083]" captionTargetId="figure-19@65.[151,1436,181,1083]" captionTargetPageId="65" captionText="FIGURES 220225. Axiagastus votypkai Salini &amp; Kment sp. nov., male and female genitalia. 220222—phallus (220—dorsal view, 221—ventral view, 222—lateral view); 223—terminalia (dorsal view); 224—terminalia (ventral view); 225—spermatheca. Scale: 0.1 mm (220222, 225), 0.5 mm (223224). Orig. S. Salini." pageId="61" pageNumber="62">Figs 220222</figureCitation>
). Articulatory apparatus as in
<figureCitation id="00845CB72747FFC4FB8CFC2B4D87047D" box="[1089,1247,1800,1827]" captionStart="FIGURES 220225" captionStartId="65.[151,264,1107,1131]" captionTargetBox="[151,1436,181,1083]" captionTargetId="figure-19@65.[151,1436,181,1083]" captionTargetPageId="65" captionText="FIGURES 220225. Axiagastus votypkai Salini &amp; Kment sp. nov., male and female genitalia. 220222—phallus (220—dorsal view, 221—ventral view, 222—lateral view); 223—terminalia (dorsal view); 224—terminalia (ventral view); 225—spermatheca. Scale: 0.1 mm (220222, 225), 0.5 mm (223224). Orig. S. Salini." pageId="61" pageNumber="62">Figs 220222</figureCitation>
; phallus as in
<taxonomicName id="5FBF3BB12747FFC4FA49FC2A48580418" authorityName="Dallas" authorityYear="1851" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C202747FFC4FA49FC2A48580418" italics="true" pageId="61" pageNumber="62">A. rosmarus</emphasis>
</taxonomicName>
and processes of aedeagus as in
<figureCitation id="00845CB72747FFC4FDBDFC0F4B890418" box="[624,721,1836,1862]" captionStart="FIGURES 220225" captionStartId="65.[151,264,1107,1131]" captionTargetBox="[151,1436,181,1083]" captionTargetId="figure-19@65.[151,1436,181,1083]" captionTargetPageId="65" captionText="FIGURES 220225. Axiagastus votypkai Salini &amp; Kment sp. nov., male and female genitalia. 220222—phallus (220—dorsal view, 221—ventral view, 222—lateral view); 223—terminalia (dorsal view); 224—terminalia (ventral view); 225—spermatheca. Scale: 0.1 mm (220222, 225), 0.5 mm (223224). Orig. S. Salini." pageId="61" pageNumber="62">Fig. 222</figureCitation>
.
</paragraph>
<paragraph id="980040322747FFC4FF0AFC73488804A4" blockId="61.[151,1437,150,2043]" pageId="61" pageNumber="62">
<emphasis id="AACB9C202747FFC4FF0AFC7348D30435" bold="true" box="[199,395,1872,1899]" italics="true" pageId="61" pageNumber="62">Female genitalia</emphasis>
(
<figureCitation id="00845CB72747FFC4FE57FC734B590434" box="[410,513,1872,1898]" captionStart="FIGURES 200206" captionStartId="62.[151,264,1859,1883]" captionTargetBox="[151,1436,201,1835]" captionTargetId="figure-29@62.[151,1436,201,1835]" captionTargetPageId="62" captionText="FIGURES 200206. Axiagastus votypkai Salini &amp; Kment sp. nov. 200201—habitus (200—dorsal view, 201—ventral view), 202203—head (202—dorsal view, 203—lateral view); 204—external scent efferent system; 205—apex of male abdomen with genital capsule in postero-ventral view; 206—apex of female abdomen with terminalia in posteroventral view. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 1 mm (200201), 0.5 mm (202206). Orig. S. Salini." pageId="61" pageNumber="62">Figs 206</figureCitation>
,
<figureCitation id="00845CB72747FFC4FDC0FC734B2C0435" box="[525,628,1872,1899]" captionStart="FIGURES 220225" captionStartId="65.[151,264,1107,1131]" captionTargetBox="[151,1436,181,1083]" captionTargetId="figure-19@65.[151,1436,181,1083]" captionTargetPageId="65" captionText="FIGURES 220225. Axiagastus votypkai Salini &amp; Kment sp. nov., male and female genitalia. 220222—phallus (220—dorsal view, 221—ventral view, 222—lateral view); 223—terminalia (dorsal view); 224—terminalia (ventral view); 225—spermatheca. Scale: 0.1 mm (220222, 225), 0.5 mm (223224). Orig. S. Salini." pageId="61" pageNumber="62">223225</figureCitation>
).
<emphasis id="AACB9C202747FFC4FD4BFC734A580434" box="[646,768,1872,1898]" italics="true" pageId="61" pageNumber="62">Terminalia</emphasis>
(
<figureCitation id="00845CB72747FFC4FCDDFC734AF60434" box="[784,942,1872,1898]" captionStart="FIGURES 220225" captionStartId="65.[151,264,1107,1131]" captionTargetBox="[151,1436,181,1083]" captionTargetId="figure-19@65.[151,1436,181,1083]" captionTargetPageId="65" captionText="FIGURES 220225. Axiagastus votypkai Salini &amp; Kment sp. nov., male and female genitalia. 220222—phallus (220—dorsal view, 221—ventral view, 222—lateral view); 223—terminalia (dorsal view); 224—terminalia (ventral view); 225—spermatheca. Scale: 0.1 mm (220222, 225), 0.5 mm (223224). Orig. S. Salini." pageId="61" pageNumber="62">Figs 223, 224</figureCitation>
) as in
<taxonomicName id="5FBF3BB12747FFC4FC34FC724DDA0434" authorityName="Dallas" authorityYear="1851" box="[1017,1154,1873,1899]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C202747FFC4FC34FC724DDA0434" box="[1017,1154,1873,1899]" italics="true" pageId="61" pageNumber="62">A. rosmarus</emphasis>
</taxonomicName>
with inner posterolateral angles of valvifers VIII obtusangulate; valvifers IX short, narrow, transverse plate with anterior margin biconcave, posterior margin more or less straight; laterotergites VIII and IX as in
<taxonomicName id="5FBF3BB12747FFC4FC12FCBB4D3304EC" authorityName="Dallas" authorityYear="1851" box="[991,1131,1944,1970]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C202747FFC4FC12FCBB4D3304EC" box="[991,1131,1944,1970]" italics="true" pageId="61" pageNumber="62">A. rosmarus</emphasis>
</taxonomicName>
except caudal margins of laterotergites VIII with 1+1 indistinct, minute sclerotized, black denticle.
<emphasis id="AACB9C202747FFC4FC0BFC9F4D670488" box="[966,1087,1980,2006]" italics="true" pageId="61" pageNumber="62">Gynatrium</emphasis>
as in
<taxonomicName id="5FBF3BB12747FFC4FBB3FC9F4DA50488" authorityName="Distant" authorityYear="1911" box="[1150,1277,1980,2006]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="cambelli">
<emphasis id="AACB9C202747FFC4FBB3FC9F4DA50488" box="[1150,1277,1980,2006]" italics="true" pageId="61" pageNumber="62">A. cambelli</emphasis>
</taxonomicName>
.
<emphasis id="AACB9C202747FFC4FAC7FC9F4CC40488" box="[1290,1436,1980,2006]" italics="true" pageId="61" pageNumber="62">Spermatheca</emphasis>
(
<figureCitation id="00845CB72747FFC4FF52FCC349A704A5" box="[159,255,2016,2043]" captionStart="FIGURES 220225" captionStartId="65.[151,264,1107,1131]" captionTargetBox="[151,1436,181,1083]" captionTargetId="figure-19@65.[151,1436,181,1083]" captionTargetPageId="65" captionText="FIGURES 220225. Axiagastus votypkai Salini &amp; Kment sp. nov., male and female genitalia. 220222—phallus (220—dorsal view, 221—ventral view, 222—lateral view); 223—terminalia (dorsal view); 224—terminalia (ventral view); 225—spermatheca. Scale: 0.1 mm (220222, 225), 0.5 mm (223224). Orig. S. Salini." pageId="61" pageNumber="62">Fig. 225</figureCitation>
) as in
<emphasis id="AACB9C202747FFC4FE84FCC3488804A4" box="[329,464,2016,2042]" italics="true" pageId="61" pageNumber="62">
<taxonomicName id="5FBF3BB12747FFC4FE84FCC3489404A4" authorityName="Distant" authorityYear="1911" box="[329,460,2016,2042]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="cambelli">A. cambelli</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="CCC010BA2744FFC7FF5AFC604C0B0499" pageId="62" pageNumber="63" startId="62.[151,264,1859,1883]" targetBox="[151,1436,201,1835]" targetPageId="62" targetType="figure">
<paragraph id="980040322744FFC7FF5AFC604C0B0499" blockId="62.[151,1437,1859,1991]" pageId="62" pageNumber="63">
<emphasis id="AACB9C202744FFC7FF5AFC6048350405" bold="true" box="[151,365,1859,1883]" pageId="62" pageNumber="63">FIGURES 200206.</emphasis>
<taxonomicName id="5FBF3BB12744FFC7FEB9FC674BBF0405" authority="Salini &amp; Kment" authorityName="Salini &amp; Kment" authorityYear="2025" box="[372,743,1859,1883]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="votypkai" status="sp. nov.">
<emphasis id="AACB9C202744FFC7FEB9FC674B180405" box="[372,576,1859,1883]" italics="true" pageId="62" pageNumber="63">Axiagastus votypkai</emphasis>
Salini &amp; Kment
</taxonomicName>
<emphasis id="AACB9C202744FFC7FD23FC604A660405" bold="true" box="[750,830,1859,1883]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="B1F8215B2744FFC7FD23FC604A660405" box="[750,830,1859,1883]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
200201—habitus (200—dorsal view, 201—ventral view), 202203—head (202—dorsal view, 203—lateral view); 204—external scent efferent system; 205—apex of male abdomen with genital capsule in postero-ventral view; 206—apex of female abdomen with terminalia in posteroventral view. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 1 mm (200201), 0.5 mm (202206). Orig. S. Salini.
</paragraph>
</caption>
<caption id="CCC010BA2745FFC6FF5AFC56482A04A6" pageId="63" pageNumber="64" startId="63.[151,264,1909,1933]" targetBox="[192,1396,181,1885]" targetPageId="63" targetType="figure">
<paragraph id="980040322745FFC6FF5AFC56482A04A6" blockId="63.[151,1436,1909,2041]" pageId="63" pageNumber="64">
<emphasis id="AACB9C202745FFC6FF5AFC56483604D3" bold="true" box="[151,366,1909,1933]" pageId="63" pageNumber="64">FIGURES 207213.</emphasis>
Male genitalia (ESEM micrographs) of
<taxonomicName id="5FBF3BB12745FFC6FCC3FC564DDC04D2" authority="Salini &amp; Kment" authorityName="Salini &amp; Kment" authorityYear="2025" box="[782,1156,1909,1933]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="votypkai" status="sp. nov.">
<emphasis id="AACB9C202745FFC6FCC3FC564A8304D3" box="[782,987,1909,1933]" italics="true" pageId="63" pageNumber="64">Axiagastus votypkai</emphasis>
Salini &amp; Kment
</taxonomicName>
<emphasis id="AACB9C202745FFC6FB41FC564D8504D3" bold="true" box="[1164,1245,1909,1933]" pageId="63" pageNumber="64">
<taxonomicNameLabel id="B1F8215B2745FFC6FB41FC564D8504D3" box="[1164,1245,1909,1933]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
207210—genital capsule (207—dorsal view, 32×; 208—ventral view, 32×; 209—caudal view, 40×; 210—lateral view, 45×); 211213—parameres (three different views, 100×). Lettering: dn—distension, te—transverse emargination. Scale: 1 mm (207210), 0.5 mm (211 213). Orig. P. Kment.
</paragraph>
</caption>
<caption id="CCC010BA273AFFB9FF5AFF7B4AFE0785" pageId="64" pageNumber="65" startId="64.[151,264,1112,1136]" targetBox="[151,1436,181,1089]" targetPageId="64" targetType="figure">
<paragraph id="98004032273AFFB9FF5AFF7B4AFE0785" blockId="64.[151,1436,1112,1243]" pageId="64" pageNumber="65">
<emphasis id="AACB9C20273AFFB9FF5AFF7B4837072F" bold="true" box="[151,367,1112,1137]" pageId="64" pageNumber="65">FIGURES 214219.</emphasis>
<taxonomicName id="5FBF3BB1273AFFB9FEB4FF7A4BAD072E" authority="Salini &amp; Kment" authorityName="Salini &amp; Kment" authorityYear="2025" box="[377,757,1113,1137]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="votypkai" status="sp. nov.">
<emphasis id="AACB9C20273AFFB9FEB4FF7A4B1F072F" box="[377,583,1113,1137]" italics="true" pageId="64" pageNumber="65">Axiagastus votypkai</emphasis>
Salini &amp; Kment
</taxonomicName>
<emphasis id="AACB9C20273AFFB9FD33FF7A4A09072F" bold="true" box="[766,849,1113,1137]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="B1F8215B273AFFB9FD33FF7A4A09072F" box="[766,849,1113,1137]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male genitalia. 214217—genital capsule (214—dorsal view, 215—ventral view, 216—caudal view, 217—lateral view); 218219—parameres (two different views). Lettering: dm— dorsal margin, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, te—transverse emargination. Scale: 0.5 mm (214217), 0.25 mm (218219). Orig. S. Salini.
</paragraph>
</caption>
<paragraph id="98004032273AFFB9FF0AFE3C4C350667" blockId="64.[151,1437,1311,2042]" box="[199,1389,1311,1337]" pageId="64" pageNumber="65">
<emphasis id="AACB9C20273AFFB9FF0AFE3C48670667" bold="true" box="[199,319,1311,1337]" pageId="64" pageNumber="65">Variation.</emphasis>
The specimen from Kinjingini is considerably smaller and paler than the remaining specimens.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B9273AFFB9FF0AFE614D510552" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="98004032273AFFB9FF0AFE614D510552" blockId="64.[151,1437,1311,2042]" pageId="64" pageNumber="65">
<emphasis id="AACB9C20273AFFB9FF0AFE6148940602" bold="true" box="[199,460,1346,1372]" pageId="64" pageNumber="65">Differential diagnosis.</emphasis>
This species is very similar to
<taxonomicName id="5FBF3BB1273AFFB9FCD3FE604AFC0602" authorityName="Dallas" authorityYear="1851" box="[798,932,1347,1373]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C20273AFFB9FCD3FE604AFC0602" box="[798,932,1347,1373]" italics="true" pageId="64" pageNumber="65">A. rosmarus</emphasis>
</taxonomicName>
from the
<collectingCountry id="E0A800A2273AFFB9FBC3FE614DD40602" box="[1038,1164,1346,1372]" name="Philippines" pageId="64" pageNumber="65">Philippines</collectingCountry>
but can be distinguished from that species by the darker or black body colour with large, black quadrate spots anteriorly on the scutellar disc. On the other hand, this scutellar pattern resembles that of
<taxonomicName id="5FBF3BB1273AFFB9FCD0FEAA4D3706FD" authority="Salini &amp; Kment" authorityName="Salini &amp; Kment" authorityYear="2025" box="[797,1135,1417,1443]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="prathapani" status="sp. nov.">
<emphasis id="AACB9C20273AFFB9FCD0FEAA4AEF06FD" box="[797,951,1417,1443]" italics="true" pageId="64" pageNumber="65">A. prathapani</emphasis>
Salini &amp; Kment
</taxonomicName>
<emphasis id="AACB9C20273AFFB9FBBBFEAA4D8806FD" bold="true" box="[1142,1232,1417,1443]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="B1F8215B273AFFB9FBBBFEAA4D8806FD" box="[1142,1232,1417,1443]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but differs by the lack of two rows of four spots anteriorly on the pronotal disc. It differs from both the species by the prevailingly pale antennae and by the male genital capsule with a distinct shape of the ventral rim, and the presence of a prominent, apically acuminate, sclerotized, black denticle within the posterolateral angles.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B9273AFFB9FF0AFD354A170528" pageId="64" pageNumber="65" type="etymology">
<paragraph id="98004032273AFFB9FF0AFD354A170528" blockId="64.[151,1437,1311,2042]" pageId="64" pageNumber="65">
<emphasis id="AACB9C20273AFFB9FF0AFD354816056E" bold="true" box="[199,334,1558,1584]" pageId="64" pageNumber="65">Etymology.</emphasis>
This species is named in honor of the eminent Czech parasitologist Jan Votýpka (Department of Parasitology, Charles University,
<collectingRegion id="5A7B8ED0273AFFB9FDDCFD1A4B3A050D" box="[529,610,1593,1619]" country="Czech Republic" name="Praha" pageId="64" pageNumber="65">Prague</collectingRegion>
,
<collectingCountry id="E0A800A2273AFFB9FDA0FD1A4A7D050D" box="[621,805,1593,1619]" name="Czech Republic" pageId="64" pageNumber="65">Czech Republic</collectingCountry>
), who collected a part of the
<typeStatus id="4704FE90273AFFB9FBA0FD1A4DC5050D" box="[1133,1181,1593,1619]" pageId="64" pageNumber="65">type</typeStatus>
series and contributed other noteworthy heteropteran samples donated to the NMPC.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B9273AFFB9FF0AFD5C4CDB045D" pageId="64" pageNumber="65" type="biology_ecology">
<paragraph id="98004032273AFFB9FF0AFD5C4CDB045D" blockId="64.[151,1437,1311,2042]" pageId="64" pageNumber="65">
<emphasis id="AACB9C20273AFFB9FF0AFD5C4B1805C7" bold="true" box="[199,576,1663,1689]" pageId="64" pageNumber="65">Biology and plant association.</emphasis>
In
<collectingRegion id="5A7B8ED0273AFFB9FDBEFDA34A5E05C7" box="[627,774,1663,1690]" country="Papua New Guinea" name="West New Britain" pageId="64" pageNumber="65">New Britain</collectingRegion>
, one of the
<typeStatus id="4704FE90273AFFB9FC6EFDA34D5605C4" box="[931,1038,1664,1690]" pageId="64" pageNumber="65" type="paratype">paratypes</typeStatus>
was collected on a cluster palm (?
<taxonomicName id="5FBF3BB1273AFFB9FF60FD81482D05E2" box="[173,373,1698,1724]" class="Liliopsida" family="Arecaceae" genus="Ptychosperma" kingdom="Plantae" order="Arecales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="AACB9C20273AFFB9FF60FD81481505E2" box="[173,333,1698,1724]" italics="true" pageId="64" pageNumber="65">Ptychosperma</emphasis>
sp.
</taxonomicName>
) (
<taxonomicName id="5FBF3BB1273AFFB9FE43FD814B5105E2" box="[398,521,1698,1724]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
), and another one “with borers in cocoa stem which probably refers to coconut palm instead (see the same mistake above regarding
<taxonomicName id="5FBF3BB1273AFFB9FD39FDE54A2D0581" authorityName="Distant" authorityYear="1911" box="[756,885,1733,1760]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="cambelli">
<emphasis id="AACB9C20273AFFB9FD39FDE54A2D0581" box="[756,885,1733,1760]" italics="true" pageId="64" pageNumber="65">A. cambelli</emphasis>
</taxonomicName>
). Another specimen from Papuan mainland was collected on leaf of
<taxonomicName id="5FBF3BB1273AFFB9FEBBFDCA4B41045D" box="[374,537,1769,1795]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="AACB9C20273AFFB9FEBBFDCA48AA045D" box="[374,498,1769,1795]" italics="true" pageId="64" pageNumber="65">Eucalyptus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="5FBF3BB1273AFFB9FDE5FDCA4BFD045D" box="[552,677,1769,1795]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="64" pageNumber="65" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
). Some of the
<typeStatus id="4704FE90273AFFB9FC88FDCA4AE8045D" box="[837,944,1769,1795]" pageId="64" pageNumber="65" type="paratype">paratypes</typeStatus>
were collected by light traps (this paper).
</paragraph>
</subSubSection>
<subSubSection id="D0A513B9273AFFB9FF0AFC2F48E10417" pageId="64" pageNumber="65" type="distribution">
<paragraph id="98004032273AFFB9FF0AFC2F48E10417" blockId="64.[151,1437,1311,2042]" pageId="64" pageNumber="65">
<emphasis id="AACB9C20273AFFB9FF0AFC2F48390478" bold="true" box="[199,353,1804,1830]" pageId="64" pageNumber="65">Distribution.</emphasis>
<collectingCountry id="E0A800A2273AFFB9FEA8FC2F48940478" box="[357,460,1804,1830]" name="Australia" pageId="64" pageNumber="65">Australia</collectingCountry>
(
<collectingRegion id="5A7B8ED0273AFFB9FE15FC2F4B380478" box="[472,608,1804,1830]" country="Australia" name="Queensland" pageId="64" pageNumber="65">Queensland</collectingRegion>
) (this paper),
<collectingCountry id="E0A800A2273AFFB9FD38FC2F4A3A0478" box="[757,866,1804,1830]" name="Indonesia" pageId="64" pageNumber="65">Indonesia</collectingCountry>
(
<collectingRegion id="5A7B8ED0273AFFB9FCA3FC2F4AEE0478" box="[878,950,1804,1830]" country="Indonesia" name="Papua Barat" pageId="64" pageNumber="65">Papua</collectingRegion>
) (this paper),
<collectingCountry id="E0A800A2273AFFB9FB81FC2F4C470478" box="[1100,1311,1804,1830]" name="Papua New Guinea" pageId="64" pageNumber="65">Papua New Guinea</collectingCountry>
(mainland,
<collectingRegion id="5A7B8ED0273AFFB9FF5AFC13487B0417" box="[151,291,1839,1866]" country="Papua New Guinea" name="West New Britain" pageId="64" pageNumber="65">New Britain</collectingRegion>
) (this paper).
</paragraph>
</subSubSection>
<subSubSection id="D0A513B9273AFFB9FF0AFC714B4404A4" pageId="64" pageNumber="65" type="discussion">
<paragraph id="98004032273AFFB9FF0AFC714B4404A4" blockId="64.[151,1437,1311,2042]" pageId="64" pageNumber="65">
<emphasis id="AACB9C20273AFFB9FF0AFC7148630432" bold="true" box="[199,315,1874,1900]" pageId="64" pageNumber="65">Remarks.</emphasis>
The
<typeStatus id="4704FE90273AFFB9FEB4FC7148830432" box="[377,475,1874,1900]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
locality label is slightly ambiguous as the GPS co-ordinates mentioned in the label indicates the collection locality as
<collectingRegion id="5A7B8ED0273AFFB9FDD9FC554B0004CE" box="[532,600,1910,1936]" country="Indonesia" name="Papua Barat" pageId="64" pageNumber="65">Papua</collectingRegion>
in
<collectingCountry id="E0A800A2273AFFB9FDB7FC564BBF04D1" box="[634,743,1909,1935]" name="Indonesia" pageId="64" pageNumber="65">Indonesia</collectingCountry>
instead of
<collectingCountry id="E0A800A2273AFFB9FC92FC554D6D04D1" box="[863,1077,1909,1936]" name="Papua New Guinea" pageId="64" pageNumber="65">Papua New Guinea</collectingCountry>
as stated on the label. Besides, a part of the type series was unambiguously collected from
<collectingRegion id="5A7B8ED0273AFFB9FCDDFCBA4A0F04ED" box="[784,855,1945,1971]" country="Indonesia" name="Papua Barat" pageId="64" pageNumber="65">Papua</collectingRegion>
,
<collectingCountry id="E0A800A2273AFFB9FCACFCBA4A8A04ED" box="[865,978,1945,1971]" name="Indonesia" pageId="64" pageNumber="65">Indonesia</collectingCountry>
. Therefore, the type locality is considered as Buare River,
<collectingRegion id="5A7B8ED0273AFFB9FE81FC9F48CB0488" box="[332,403,1980,2006]" country="Indonesia" name="Papua Barat" pageId="64" pageNumber="65">Papua</collectingRegion>
,
<collectingCountry id="E0A800A2273AFFB9FE52FC9F4B480488" box="[415,528,1980,2006]" name="Indonesia" pageId="64" pageNumber="65">Indonesia</collectingCountry>
. This is also applicable to a female
<typeStatus id="4704FE90273AFFB9FC6FFC9F4D5D0488" box="[930,1029,1980,2006]" pageId="64" pageNumber="65" type="paratype">paratype</typeStatus>
, which bears the same locality label as the
<typeStatus id="4704FE90273AFFB9FF12FCC3481C04A4" box="[223,324,2016,2042]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
, deposited in NIM.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,287 @@
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<mods:title id="7D3D9598BBF0E1C5B818C0C7F3C54784">Revision of the Austro-Oriental shield-bug genus Axiagastus (Hemiptera: Heteroptera: Pentatomidae), with the description of five new species, and taxonomic notes on related genera</mods:title>
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<mods:namePart id="E5F798CABD528B6CD970F2D4186B8A16">Kment, Petr</mods:namePart>
<mods:nameIdentifier id="13BA757D92083E8A342B1E3E678411B7" type="ORCID">0000-0002-7026-5691</mods:nameIdentifier>
<mods:affiliation id="189275355AC0A8D9B8E644B22F6BB911">Department of Entomology, National Museum of the Czech Republic, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
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<mods:namePart id="329FD80B2DC58BF7DA1FC98AB012ECB1">Lemaître, Valérie A.</mods:namePart>
<mods:nameIdentifier id="317E3EC59BECC616970CA671C1E05C9E" type="ORCID">0000-0003-4802-2711</mods:nameIdentifier>
<mods:affiliation id="2DFABAFDB737D93DFC13123B1A977F4F">Department of Collections (Insects), The Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom; &amp; val. heureuse @ gmail. com; https: // orcid. org / 0000 - 0003 - 4802 - 2711</mods:affiliation>
<mods:nameIdentifier id="D1FDC20E97030D7EDB6A29865D9D2749" type="email">val.heureuse@gmail.com</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="5EB1538EE738EF2F396108B749EFDF03">Webb, Michael D.</mods:namePart>
<mods:nameIdentifier id="ABA44BF63BD2396EDA7AA2FB34B4452F" type="ORCID">0000-0002-1312-6142</mods:nameIdentifier>
<mods:affiliation id="EB0878BA459EEDB72CA5BD71465EE14C">Department of Collections (Insects), The Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom; &amp; mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
<mods:nameIdentifier id="56FFE70BF55175E1167651AF4D79F2B5" type="email">mdwebb04@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:role id="2838DF0E0BDA0D886CBC575CE8C37B56">
<mods:roleTerm id="05726454597ABD17464A348323F3EBF1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A15D49A8E53003971FD2143183A95D6F">Roca-Cusachs, Marcos</mods:namePart>
<mods:nameIdentifier id="4485AECB3FA7BD72EC9B1901269C550D" type="ORCID">0000-0003-3557-4365</mods:nameIdentifier>
<mods:affiliation id="FF888517C6EDC93E077F5E345E2716B5">ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, Karnataka, India shalinis. nilavu @ gmail. com; https: // orcid. org / 0000 - 0003 - 3557 - 4365 &amp; Evolution &amp; Ecology Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, Australia &amp; IRBio, Institut de Recerca de la Biodiversitat, Universitat de Barcelona, Barcelona, Spain marcosrocacusachs @ gmail. com; https: // orcid. org / 0000 - 0002 - 9174 - 6021 # Former affiliation * Corresponding author &amp; ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, Karnataka, India shalinis. nilavu @ gmail. com; https: // orcid. org / 0000 - 0003 - 3557 - 4365</mods:affiliation>
<mods:nameIdentifier id="38E61FC471711AC7A3A6A9030AAADED2" type="email">shalinis.nilavu@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="D74E20F7189CEEAA1E0502E4A4D7998E">Zootaxa</mods:title>
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<mods:date id="CD755EE16AA2F530D21CD14EAEE0B9C2">2025</mods:date>
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<mods:number id="18CA22CA0E4B813D94923FAD38E1B8C3">5603</mods:number>
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<taxonomicName id="5FBF3BB12751FFD2FF5AFBB44BF103EF" authority="Salini &amp; Roca-Cusachs" authorityName="Salini &amp; Roca-Cusachs" authorityYear="2025" box="[151,681,151,178]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="luteipes" status="sp. nov.">
<emphasis id="AACB9C202751FFD2FF5AFBB448D903EC" bold="true" box="[151,385,151,178]" italics="true" pageId="43" pageNumber="44">Axiagastus luteipes</emphasis>
Salini &amp; Roca-Cusachs
</taxonomicName>
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</emphasis>
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<paragraph id="980040322751FFD2FF5AFBFC481A03A7" blockId="43.[151,1173,151,249]" box="[151,322,223,249]" pageId="43" pageNumber="44">
(
<figureCitation id="00845CB72751FFD2FF52FBFC486203A7" box="[159,314,223,249]" captionStart-0="FIGURES 117118" captionStart-1="FIGURES 119122" captionStart-2="FIGURES 123126" captionStart-3="FIGURES 127133" captionStartId-0="43.[151,264,1943,1967]" captionStartId-1="44.[151,264,1206,1230]" captionStartId-2="45.[151,264,1861,1885]" captionStartId-3="46.[151,264,1203,1227]" captionTargetBox-0="[151,1436,921,1920]" captionTargetBox-1="[151,1436,181,1182]" captionTargetBox-2="[151,1436,777,1837]" captionTargetBox-3="[151,1436,181,1179]" captionTargetId-0="figure-297@43.[151,1436,921,1920]" captionTargetId-1="figure-29@44.[151,1436,181,1182]" captionTargetId-2="figure-273@45.[151,1436,777,1837]" captionTargetId-3="figure-29@46.[151,1436,181,1179]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionTargetPageId-2="45" captionTargetPageId-3="46" captionText-0="FIGURES 117118. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., habitus. 117—dorsal view; 118—ventral view. Scale: 1 mm (117118). Orig. S. Salini." captionText-1="FIGURES 119122. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov. 119—head (dorsal view); 120—head (lateral view); 121—apex of male abdomen with genital capsule in postero-ventral view; 122—external scent efferent system. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (119122). Orig. S. Salini." captionText-2="FIGURES 123126. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 123126—genital capsule (123— dorsal view, 124—ventral view, 125—caudal view, 126—lateral view). Lettering: dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, te—transverse emargination. Scale: 0.5 mm (123126). Orig. S. Salini." captionText-3="FIGURES 127133. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 127129—paramere (three different views); 130—articulatory apparatus; 131133—phallus (131—dorsal view, 132—ventral view, 133—lateral view). Lettering: dm—dorsal margin, pa—processes of aedeagus. Scale: 0.1 mm (127133). Orig. S. Salini." pageId="43" pageNumber="44">Figs 117133</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92751FFD2FF5AFA04486E0147" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="980040322751FFD2FF5AFA044D02021F" blockId="43.[151,1437,295,898]" box="[151,1114,295,322]" pageId="43" pageNumber="44">
<materialsCitation id="28D74A6F2751FFD2FF5AFA044D02021F" box="[151,1114,295,322]" country="Myanmar" elevation="450" location="Hills" pageId="43" pageNumber="44" specimenCount="1" stateProvince="Chin" typeStatus="holotype">
<emphasis id="AACB9C202751FFD2FF5AFA04486E021F" bold="true" box="[151,310,295,321]" pageId="43" pageNumber="44">
<typeStatus id="4704FE902751FFD2FF5AFA04498A021F" box="[151,210,295,321]" pageId="43" pageNumber="44">Type</typeStatus>
locality.
</emphasis>
NW
<collectingCountry id="E0A800A22751FFD2FEBCFA0B48B8021C" box="[369,480,296,322]" name="Myanmar" pageId="43" pageNumber="44">Myanmar</collectingCountry>
,
<collectingRegion id="5A7B8ED02751FFD2FE26FA044B3C021F" box="[491,612,295,321]" country="Myanmar" name="Chin" pageId="43" pageNumber="44">Chin State</collectingRegion>
,
<collectingRegion id="5A7B8ED02751FFD2FDA2FA044BFD021F" box="[623,677,295,321]" country="Myanmar" name="Chin" pageId="43" pageNumber="44">Chin</collectingRegion>
<location id="9D6016E92751FFD2FD61FA044BBD021F" LSID="urn:lsid:plazi:treatment:1016F1242751FFD4FF5AFBB44A300193:9D6016E92751FFD2FD61FA044BBD021F" box="[684,741,295,321]" country="Myanmar" name="Hills" pageId="43" pageNumber="44" stateProvince="Chin">Hills</location>
,
<elevation id="1392A7012751FFD2FD3CFA044AF4021F" box="[753,940,295,322]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="43" pageNumber="44" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<quantity id="5F47EDD72751FFD2FD3CFA044A28021C" box="[753,880,295,322]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="43" pageNumber="44" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400500 m</quantity>
a.s.l.
</elevation>
,
<locationDeviation id="89B259102751FFD2FC75FA0B4D02021F" box="[952,1114,295,322]" location="Falam." pageId="43" pageNumber="44">NW of Falam.</locationDeviation>
</materialsCitation>
</paragraph>
<paragraph id="980040322751FFD2FF0AFA68486E0147" blockId="43.[151,1437,295,898]" pageId="43" pageNumber="44">
<emphasis id="AACB9C202751FFD2FF0AFA68482E023B" bold="true" box="[199,374,331,357]" pageId="43" pageNumber="44">Type material.</emphasis>
<materialsCitation id="28D74A6F2751FFD2FEB2FA68486E0147" collectingDate="2016-06-05" collectionCode="NIM" collectorName="The" country="Myanmar" determinerName="Salini, S." elevation="450" location="Chin State" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Chin" typeStatus="holotype">
<typeStatus id="4704FE902751FFD2FEB2FA6848B4023B" box="[383,492,331,357]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
:
<specimenCount id="8EB98BBB2751FFD2FE3AFA6F4B540238" box="[503,524,332,358]" count="1" pageId="43" pageNumber="44" type="male"></specimenCount>
(
<figureCitation id="00845CB72751FFD2FDD0FA684BE3023B" box="[541,699,331,357]" captionStart-0="FIGURES 117118" captionStart-1="FIGURES 119122" captionStart-2="FIGURES 123126" captionStart-3="FIGURES 127133" captionStartId-0="43.[151,264,1943,1967]" captionStartId-1="44.[151,264,1206,1230]" captionStartId-2="45.[151,264,1861,1885]" captionStartId-3="46.[151,264,1203,1227]" captionTargetBox-0="[151,1436,921,1920]" captionTargetBox-1="[151,1436,181,1182]" captionTargetBox-2="[151,1436,777,1837]" captionTargetBox-3="[151,1436,181,1179]" captionTargetId-0="figure-297@43.[151,1436,921,1920]" captionTargetId-1="figure-29@44.[151,1436,181,1182]" captionTargetId-2="figure-273@45.[151,1436,777,1837]" captionTargetId-3="figure-29@46.[151,1436,181,1179]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionTargetPageId-2="45" captionTargetPageId-3="46" captionText-0="FIGURES 117118. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., habitus. 117—dorsal view; 118—ventral view. Scale: 1 mm (117118). Orig. S. Salini." captionText-1="FIGURES 119122. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov. 119—head (dorsal view); 120—head (lateral view); 121—apex of male abdomen with genital capsule in postero-ventral view; 122—external scent efferent system. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (119122). Orig. S. Salini." captionText-2="FIGURES 123126. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 123126—genital capsule (123— dorsal view, 124—ventral view, 125—caudal view, 126—lateral view). Lettering: dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, te—transverse emargination. Scale: 0.5 mm (123126). Orig. S. Salini." captionText-3="FIGURES 127133. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 127129—paramere (three different views); 130—articulatory apparatus; 131133—phallus (131—dorsal view, 132—ventral view, 133—lateral view). Lettering: dm—dorsal margin, pa—processes of aedeagus. Scale: 0.1 mm (127133). Orig. S. Salini." pageId="43" pageNumber="44">Figs 117133</figureCitation>
), “NW
<collectingCountry id="E0A800A22751FFD2FCDEFA684AF0023B" box="[787,936,331,357]" name="Myanmar" pageId="43" pageNumber="44">MYANMAR</collectingCountry>
,
<collectingRegion id="5A7B8ED02751FFD2FC78FA684D74023B" box="[949,1068,331,357]" country="Myanmar" name="Chin" pageId="43" pageNumber="44">Chin State</collectingRegion>
/
<collectingRegion id="5A7B8ED02751FFD2FB88FA684DD1023B" box="[1093,1161,331,357]" country="Myanmar" name="Chin" pageId="43" pageNumber="44">CHIN</collectingRegion>
HILLS, alt.
<quantity id="5F47EDD72751FFD2FAD1FA684CC40238" box="[1308,1436,331,358]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="43" pageNumber="44" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<elevation id="1392A7012751FFD2FAD1FA684CC40238" box="[1308,1436,331,358]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="43" pageNumber="44" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400500 m</elevation>
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/ NW of FALAM /
<date id="EC0166F22751FFD2FEB5FA4C48BD02D4" box="[376,485,367,394]" pageId="43" pageNumber="44" value="2016-06-05">
<collectingDate id="FC459F1A2751FFD2FEB5FA4C48BD02D4" box="[376,485,367,394]" pageId="43" pageNumber="44" value="2016-06-05">5.vi.2016</collectingDate>
</date>
, local collector lgt. [p] // dissected and illustrated by Salini. S. [p] // Roca-Cusachs Personal Collection [p] //
<typeStatus id="4704FE902751FFD2FE78FAB04B0302F3" box="[437,603,403,429]" pageId="43" pageNumber="44" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="5FBF3BB12751FFD2FDBDFAB04ACE02F3" authorityName="Salini &amp; Roca-Cusachs" authorityYear="2025" box="[624,918,403,429]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="43" pageNumber="72" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="AACB9C202751FFD2FDBDFAB04ACE02F3" box="[624,918,403,429]" italics="true" pageId="43" pageNumber="44">AXIAGASTUS LUTEIPES</emphasis>
</taxonomicName>
/ Salini &amp; Roca-Cusachs sp. nov. // det.
<determinerName id="6DDCACF62751FFD2FA99FAB049F7028F" pageId="43" pageNumber="44">Salini, S.</determinerName>
, 2024 [p, red label]” (
<collectionCode id="FEAED8F72751FFD2FE6CFA944885028F" box="[417,477,439,465]" pageId="43" pageNumber="44">NIM</collectionCode>
).
<collectorName id="354A25E42751FFD2FE22FA944B43028F" box="[495,539,439,465]" pageId="43" pageNumber="44">The</collectorName>
<typeStatus id="4704FE902751FFD2FDECFA944BDB028F" box="[545,643,439,465]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
is pinned through the scutellum, the dissected male genitalia are placed in a glass microvial with glycerol attached to the same pin; antennomeres III and IV of both antennae and one paramere lost.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92751FFD4FF0AF9004B57023B" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44" type="description">
<paragraph id="980040322751FFD5FF0AF9004C4E05FC" blockId="43.[151,1437,295,898]" lastBlockId="44.[151,1437,1384,2023]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
<emphasis id="AACB9C202751FFD2FF0AF9004B800163" bold="true" box="[199,728,547,574]" pageId="43" pageNumber="44">
Description.
<emphasis id="AACB9C202751FFD2FE92F9004B800163" bold="true" box="[351,728,547,574]" italics="true" pageId="43" pageNumber="44">Colour, integument and vestiture</emphasis>
</emphasis>
(
<figureCitation id="00845CB72751FFD2FD2BF9004AE30163" box="[742,955,547,573]" captionStart-0="FIGURES 117118" captionStart-1="FIGURES 119122" captionStartId-0="43.[151,264,1943,1967]" captionStartId-1="44.[151,264,1206,1230]" captionTargetBox-0="[151,1436,921,1920]" captionTargetBox-1="[151,1436,181,1182]" captionTargetId-0="figure-297@43.[151,1436,921,1920]" captionTargetId-1="figure-29@44.[151,1436,181,1182]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURES 117118. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., habitus. 117—dorsal view; 118—ventral view. Scale: 1 mm (117118). Orig. S. Salini." captionText-1="FIGURES 119122. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov. 119—head (dorsal view); 120—head (lateral view); 121—apex of male abdomen with genital capsule in postero-ventral view; 122—external scent efferent system. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (119122). Orig. S. Salini." pageId="43" pageNumber="44">Figs 117120, 122</figureCitation>
). Dorsum of head, pronotum and scutellum bright yellow; coria appear black due to presence of dense, coarse punctures; coarse, black punctures scattered over disc of pronotum and scutellum (except impunctate scutellar apex), punctures outlining cicatrices on anterior disc of pronotum; disc of head with punctures fine, black, arranged in longitudinal rows; lateral margins of head black; antennae impunctate, with each scape (I) pale yellow, basi- (IIa) and distipedicellite (IIb) black (basi- and distiflagellum (IIIIV) missing); pronotum with anterolateral margins including humeri black, disc posterior to imaginary transverse line across humeri, dividing pronotal disc into anterior and posterior parts, black (appears as thick, black transverse band on basal pronotal disc), prefrenal disc of scutellum with moderately large, transverse, oval, undivided black spot; another, thick, transverse inverted U-shaped spot just before scutellar apex, black; scutellar apex appears as yellowish, more or less round spot, narrowly truncate anteriorly, devoid of punctures; thick, coarse, dense punctures on coria except costal margins and major veins of hemelytra; membranes smoky brown; connexiva bright yellow with extreme lateral margins of abdomen, including denticle at posterolateral angle of each abdominal ventrite, black; anterior and posterior fourths of each connexival segment black with black punctures, middle half of each connexival segment bright yellow, with coarse, brown punctures. Ventral surface of body including legs and labium luteous, scattered with black punctures; punctures arranged in a similar pattern as that of previous species; punctures large, scattered on legs; apices of buccular denticles, labium, and tarsal claws, a single spot at apex of each ostiolar peritreme, small round spot laterad to muscle scars on each ventrite from IIIVII, moderately large spot mesad to each spiracle coalescing with spiracular outline, anterior and posterolateral angles of ventrites IIIVII, black; ventrites with intersegmental sutures reinforced with brown transverse stripes.
</paragraph>
<caption id="CCC010BA2751FFD2FF5AFCB448BC048A" pageId="43" pageNumber="44" startId="43.[151,264,1943,1967]" targetBox="[151,1436,921,1920]" targetPageId="43" targetType="figure">
<paragraph id="980040322751FFD2FF5AFCB448BC048A" blockId="43.[151,1437,1943,2004]" pageId="43" pageNumber="44">
<emphasis id="AACB9C202751FFD2FF5AFCB4483104F1" bold="true" box="[151,361,1943,1967]" pageId="43" pageNumber="44">FIGURES 117118.</emphasis>
<taxonomicName id="5FBF3BB12751FFD2FEA2FCBB4A7B04EE" authority="Salini &amp; Roca-Cusachs" authorityName="Salini &amp; Roca-Cusachs" authorityYear="2025" box="[367,803,1944,1968]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="luteipes" status="sp. nov.">
<emphasis id="AACB9C202751FFD2FEA2FCBB4B6B04EE" box="[367,563,1944,1968]" italics="true" pageId="43" pageNumber="44">Axiagastus luteipes</emphasis>
Salini &amp; Roca-Cusachs
</taxonomicName>
<emphasis id="AACB9C202751FFD2FCE5FCBB4A2004EE" bold="true" box="[808,888,1944,1968]" pageId="43" pageNumber="44">
<taxonomicNameLabel id="B1F8215B2751FFD2FCE5FCBB4A2004EE" box="[808,888,1944,1968]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus. 117—dorsal view; 118—ventral view. Scale: 1 mm (117
<emphasis id="AACB9C202751FFD2FEC8FC9F484A048A" bold="true" box="[261,274,1980,2004]" pageId="43" pageNumber="44"></emphasis>
118). Orig. S. Salini.
</paragraph>
</caption>
<caption id="CCC010BA2756FFD5FF5AFF954DC30648" pageId="44" pageNumber="45" startId="44.[151,264,1206,1230]" targetBox="[151,1436,181,1182]" targetPageId="44" targetType="figure">
<paragraph id="980040322756FFD5FF5AFF954DC30648" blockId="44.[151,1437,1206,1302]" pageId="44" pageNumber="45">
<emphasis id="AACB9C202756FFD5FF5AFF9548350790" bold="true" box="[151,365,1206,1230]" pageId="44" pageNumber="45">FIGURES 119122.</emphasis>
<taxonomicName id="5FBF3BB12756FFD5FEBEFF954A750790" authority="Salini &amp; Roca-Cusachs" authorityName="Salini &amp; Roca-Cusachs" authorityYear="2025" box="[371,813,1206,1230]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="luteipes" status="sp. nov.">
<emphasis id="AACB9C202756FFD5FEBEFF954B600790" box="[371,568,1206,1230]" italics="true" pageId="44" pageNumber="45">Axiagastus luteipes</emphasis>
Salini &amp; Roca-Cusachs
</taxonomicName>
<emphasis id="AACB9C202756FFD5FCF9FF954ADD0790" bold="true" box="[820,901,1206,1230]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="B1F8215B2756FFD5FCF9FF954ADD0790" box="[820,901,1206,1230]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
119—head (dorsal view); 120—head (lateral view); 121—apex of male abdomen with genital capsule in postero-ventral view; 122—external scent efferent system. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (119122). Orig. S. Salini.
</paragraph>
</caption>
<paragraph id="980040322756FFD5FF0AFD8F4C280598" blockId="44.[151,1437,1384,2023]" box="[199,1392,1708,1735]" pageId="44" pageNumber="45">
<emphasis id="AACB9C202756FFD5FF0AFD8F48630598" bold="true" box="[199,315,1708,1734]" italics="true" pageId="44" pageNumber="45">Structure.</emphasis>
Labium surpassing posterior margin of ventrite III. Other characters as in generic redescription.
</paragraph>
<paragraph id="980040322756FFD4FF0AFDF34D88021F" blockId="44.[151,1437,1384,2023]" lastBlockId="45.[151,1437,151,717]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<emphasis id="AACB9C202756FFD5FF0AFDF3483405B5" bold="true" box="[199,364,1744,1771]" italics="true" pageId="44" pageNumber="45">Male genitalia</emphasis>
(
<figureCitation id="00845CB72756FFD5FEB5FDF3488305B4" box="[376,475,1744,1770]" captionStart="FIGURES 119122" captionStartId="44.[151,264,1206,1230]" captionTargetBox="[151,1436,181,1182]" captionTargetId="figure-29@44.[151,1436,181,1182]" captionTargetPageId="44" captionText="FIGURES 119122. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov. 119—head (dorsal view); 120—head (lateral view); 121—apex of male abdomen with genital capsule in postero-ventral view; 122—external scent efferent system. Lettering: d—“tusk-like” denticles, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (119122). Orig. S. Salini." pageId="44" pageNumber="45">Figs 121</figureCitation>
,
<figureCitation id="00845CB72756FFD5FE29FDF34B1205B4" box="[484,586,1744,1770]" captionStart-0="FIGURES 123126" captionStart-1="FIGURES 127133" captionStartId-0="45.[151,264,1861,1885]" captionStartId-1="46.[151,264,1203,1227]" captionTargetBox-0="[151,1436,777,1837]" captionTargetBox-1="[151,1436,181,1179]" captionTargetId-0="figure-273@45.[151,1436,777,1837]" captionTargetId-1="figure-29@46.[151,1436,181,1179]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionText-0="FIGURES 123126. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 123126—genital capsule (123— dorsal view, 124—ventral view, 125—caudal view, 126—lateral view). Lettering: dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, te—transverse emargination. Scale: 0.5 mm (123126). Orig. S. Salini." captionText-1="FIGURES 127133. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 127129—paramere (three different views); 130—articulatory apparatus; 131133—phallus (131—dorsal view, 132—ventral view, 133—lateral view). Lettering: dm—dorsal margin, pa—processes of aedeagus. Scale: 0.1 mm (127133). Orig. S. Salini." pageId="44" pageNumber="45">123133</figureCitation>
). Genital capsule (
<figureCitation id="00845CB72756FFD5FCDFFDF34AF405B4" box="[786,940,1744,1770]" captionStart="FIGURES 123126" captionStartId="45.[151,264,1861,1885]" captionTargetBox="[151,1436,777,1837]" captionTargetId="figure-273@45.[151,1436,777,1837]" captionTargetPageId="45" captionText="FIGURES 123126. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 123126—genital capsule (123— dorsal view, 124—ventral view, 125—caudal view, 126—lateral view). Lettering: dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, te—transverse emargination. Scale: 0.5 mm (123126). Orig. S. Salini." pageId="44" pageNumber="45">Figs 123126</figureCitation>
) subquadrate, dorsal rim more deeply incised than ventral rim, lateral wall of dorsal rim slightly concave, ending in moderately angular projection contiguous with moderately deep, transverse emargination (te) at middle of dorsal rim, dorsal sinus of posterior aperture broadly U-shaped (ds), ventral margin of posterior aperture semiovate, posterolateral lobes narrowly rounded in lateral view, ventral rim broadly V-shaped at middle, infoldings of ventral rim deeply impressed on either side of well-developed distension (dn) at middle, distension (dn) emarginate at middle, visible as tongue-like structure on ventral side of genital capsule, and as globose structure in lateral view, infoldings of ventral rim laterally (inner to posterolateral lobes) with a narrow sclerotized, black, more or less blunt denticle (dt).
<emphasis id="AACB9C202756FFD5FBBBFCEE4DBD04B8" box="[1142,1253,1997,2022]" italics="true" pageId="44" pageNumber="45">Paramere</emphasis>
(
<figureCitation id="00845CB72756FFD5FB3BFCEF4CCC04B8" box="[1270,1428,1996,2022]" captionStart="FIGURES 127133" captionStartId="46.[151,264,1203,1227]" captionTargetBox="[151,1436,181,1179]" captionTargetId="figure-29@46.[151,1436,181,1179]" captionTargetPageId="46" captionText="FIGURES 127133. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 127129—paramere (three different views); 130—articulatory apparatus; 131133—phallus (131—dorsal view, 132—ventral view, 133—lateral view). Lettering: dm—dorsal margin, pa—processes of aedeagus. Scale: 0.1 mm (127133). Orig. S. Salini." pageId="44" pageNumber="45">Figs 127129</figureCitation>
) simple, crown with an upright finger-like process forming an acute angle with a laminate disc, dorsal margin (dm) convex, abruptly narrowed towards apex, apex appearing acute in lateral view (
<figureCitation id="00845CB72757FFD4FBDBFB984D21038B" box="[1046,1145,187,213]" captionStart="FIGURES 127133" captionStartId="46.[151,264,1203,1227]" captionTargetBox="[151,1436,181,1179]" captionTargetId="figure-29@46.[151,1436,181,1179]" captionTargetPageId="46" captionText="FIGURES 127133. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 127129—paramere (three different views); 130—articulatory apparatus; 131133—phallus (131—dorsal view, 132—ventral view, 133—lateral view). Lettering: dm—dorsal margin, pa—processes of aedeagus. Scale: 0.1 mm (127133). Orig. S. Salini." pageId="45" pageNumber="46">Fig. 127</figureCitation>
); numerous fine, elongate setae along periphery of laminate disc (ld); stem moderately elongate, apodeme disc-like.
<emphasis id="AACB9C202757FFD4FB41FBFC4DBA03A7" box="[1164,1250,223,249]" italics="true" pageId="45" pageNumber="46">Phallus</emphasis>
(
<figureCitation id="00845CB72757FFD4FB3CFBFC4CC903A7" box="[1265,1425,223,249]" captionStart="FIGURES 127133" captionStartId="46.[151,264,1203,1227]" captionTargetBox="[151,1436,181,1179]" captionTargetId="figure-29@46.[151,1436,181,1179]" captionTargetPageId="46" captionText="FIGURES 127133. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 127129—paramere (three different views); 130—articulatory apparatus; 131133—phallus (131—dorsal view, 132—ventral view, 133—lateral view). Lettering: dm—dorsal margin, pa—processes of aedeagus. Scale: 0.1 mm (127133). Orig. S. Salini." pageId="45" pageNumber="46">Figs 131133</figureCitation>
). Articulatory apparatus as in
<figureCitation id="00845CB72757FFD4FE2CFA204B1C0243" box="[481,580,259,285]" captionStart="FIGURES 127133" captionStartId="46.[151,264,1203,1227]" captionTargetBox="[151,1436,181,1179]" captionTargetId="figure-29@46.[151,1436,181,1179]" captionTargetPageId="46" captionText="FIGURES 127133. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 127129—paramere (three different views); 130—articulatory apparatus; 131133—phallus (131—dorsal view, 132—ventral view, 133—lateral view). Lettering: dm—dorsal margin, pa—processes of aedeagus. Scale: 0.1 mm (127133). Orig. S. Salini." pageId="45" pageNumber="46">Fig. 130</figureCitation>
; phallotheca, conjunctival process and aedeagus as in
<taxonomicName id="5FBF3BB12757FFD4FB71FA2048AF021F" authority="Salini, Kment &amp; Roca-Cusachs" authorityName="Salini, Kment &amp; Roca-Cusachs" authorityYear="2025" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="chandrashekarai" status="sp. nov.">
<emphasis id="AACB9C202757FFD4FB71FA204CC40243" box="[1212,1436,259,285]" italics="true" pageId="45" pageNumber="46">A. chandrashekarai</emphasis>
Salini, Kment &amp; Roca-Cusachs
</taxonomicName>
<taxonomicNameLabel id="B1F8215B2757FFD4FE33FA044B01021F" box="[510,601,295,321]" pageId="45" pageNumber="46" rank="species">
<emphasis id="AACB9C202757FFD4FE33FA044B0A021F" bold="true" box="[510,594,295,321]" pageId="45" pageNumber="46">sp. nov</emphasis>
.
</taxonomicNameLabel>
; a pair of elongate processes of aedeagus as in
<figureCitation id="00845CB72757FFD4FBA6FA044D93021F" box="[1131,1227,295,321]" captionStart="FIGURES 127133" captionStartId="46.[151,264,1203,1227]" captionTargetBox="[151,1436,181,1179]" captionTargetId="figure-29@46.[151,1436,181,1179]" captionTargetPageId="46" captionText="FIGURES 127133. Axiagastus luteipes Salini &amp; Roca-Cusachs sp. nov., male genitalia. 127129—paramere (three different views); 130—articulatory apparatus; 131133—phallus (131—dorsal view, 132—ventral view, 133—lateral view). Lettering: dm—dorsal margin, pa—processes of aedeagus. Scale: 0.1 mm (127133). Orig. S. Salini." pageId="45" pageNumber="46">Fig. 133</figureCitation>
.
</paragraph>
<paragraph id="980040322757FFD4FF0AFA694B57023B" blockId="45.[151,1437,151,717]" box="[199,527,330,357]" pageId="45" pageNumber="46">
<emphasis id="AACB9C202757FFD4FF0AFA6948C9023B" bold="true" box="[199,401,330,357]" italics="true" pageId="45" pageNumber="46">Female genitalia.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92757FFD4FF0AFA4D4CD90163" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="980040322757FFD4FF0AFA4D4C7F02AB" blockId="45.[151,1437,151,717]" pageId="45" pageNumber="46">
<emphasis id="AACB9C202757FFD4FF0AFA4D489402D6" bold="true" box="[199,460,366,392]" pageId="45" pageNumber="46">Differential diagnosis.</emphasis>
Compared to other congeners, this species has a moderately large body size and a shinier luteous body colour with the dorsal surface with shiny black markings on a luteous background. It is very distinct in the shape of the median distension of the male genital capsule, which is emarginate at middle and is visible as a tongue-like structure on the ventral side of the genital capsule, and as a globose structure in lateral view.
</paragraph>
<paragraph id="980040322757FFD4FF0AFADC4CD90163" blockId="45.[151,1437,151,717]" pageId="45" pageNumber="46">
In colouration, this species resembles a dark specimen of
<taxonomicName id="5FBF3BB12757FFD4FC8CFADC4A850147" authorityName="Kment &amp; Lemaître &amp; Webb &amp; Roca-Cusachs" authorityYear="2025" box="[833,989,511,537]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="yeshwanthi" status="sp. nov.">
<emphasis id="AACB9C202757FFD4FC8CFADC4A850147" box="[833,989,511,537]" italics="true" pageId="45" pageNumber="46">A. yeshwanthi</emphasis>
</taxonomicName>
<emphasis id="AACB9C202757FFD4FC2FFADC4D630147" bold="true" box="[994,1083,511,537]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="B1F8215B2757FFD4FC2FFADC4D630147" box="[994,1083,511,537]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="00845CB72757FFD4FB84FADC4DF20147" box="[1097,1194,511,537]" captionStart="FIGURES 232235" captionStartId="68.[151,264,1909,1933]" captionTargetBox="[192,1396,181,1885]" captionTargetId="figure-29@68.[192,1396,181,1885]" captionTargetPageId="68" captionText="FIGURES 232235. Axiagastus yeshwanthi Salini, Kment &amp; Webb sp. nov., paratype male from India (body length 13.27 mm). 232233—habitus (232—dorsal view, 233—ventral view); 234—female from Cambodia (body length 12.00 mm), habitus, dorsal view; 235—male from Nantou, Taiwan (body length 14.00 mm), habitus, dorsal view. Scale: 1 mm (232233). Orig. S. Salini (232233), P. Kment (234235)." pageId="45" pageNumber="46">Fig. 234</figureCitation>
), but they differ in the shape of the whitish spot on the apex of the scutellum, which is shorter and anteriorly trilobate in
<taxonomicName id="5FBF3BB12757FFD4FB10F9004C220163" authorityName="Kment &amp; Lemaître &amp; Webb &amp; Roca-Cusachs" authorityYear="2025" box="[1245,1402,547,573]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="yeshwanthi">
<emphasis id="AACB9C202757FFD4FB10F9004C220163" box="[1245,1402,547,573]" italics="true" pageId="45" pageNumber="46">A. yeshwanthi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92757FFD4FF0AF964480601F7" pageId="45" pageNumber="46" type="etymology">
<paragraph id="980040322757FFD4FF0AF964480601F7" blockId="45.[151,1437,151,717]" pageId="45" pageNumber="46">
<emphasis id="AACB9C202757FFD4FF0AF9644816013F" bold="true" box="[199,334,583,609]" pageId="45" pageNumber="46">Etymology.</emphasis>
The species-group name is a Latin word combining the adjective
<emphasis id="AACB9C202757FFD4FBDAF9644D02013F" box="[1047,1114,583,609]" italics="true" pageId="45" pageNumber="46">luteus</emphasis>
(-
<emphasis id="AACB9C202757FFD4FBBCF9644D27013E" box="[1137,1151,583,608]" italics="true" pageId="45" pageNumber="46">a</emphasis>
, -
<emphasis id="AACB9C202757FFD4FB59F9644DEE013E" box="[1172,1206,583,608]" italics="true" pageId="45" pageNumber="46">um</emphasis>
), meaning “yellow”, and the noun
<emphasis id="AACB9C202757FFD4FEE2F948480C01DA" box="[303,340,619,644]" italics="true" pageId="45" pageNumber="46">pes</emphasis>
, meaning “foot”; noun in apposition. The name refers to the yellowish colouration of the species, including its legs.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92757FFD4FF0AF9914A300193" box="[199,872,690,717]" pageId="45" pageNumber="46" type="distribution">
<paragraph id="980040322757FFD4FF0AF9914A300193" blockId="45.[151,1437,151,717]" box="[199,872,690,717]" pageId="45" pageNumber="46">
<emphasis id="AACB9C202757FFD4FF0AF99148390192" bold="true" box="[199,353,690,716]" pageId="45" pageNumber="46">Distribution.</emphasis>
Known only from NW
<collectingCountry id="E0A800A22757FFD4FDA1F9904B810193" box="[620,729,691,717]" name="Myanmar" pageId="45" pageNumber="46">Myanmar</collectingCountry>
(this paper).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="59BB06437EDBE9219F8585EDA78F2149">Revision of the Austro-Oriental shield-bug genus Axiagastus (Hemiptera: Heteroptera: Pentatomidae), with the description of five new species, and taxonomic notes on related genera</mods:title>
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(checklist);
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<bibRefCitation id="FC2E3DC32754FFD7FDDCFD6A4A59053F" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[529,769,1609,1633]" pageId="46" pageNumber="47" pagination="297 - 300" refId="ref51680" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297-300, pls. 5 - 25. [book pagination: iii + 330 pp + 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng (1977)</bibRefCitation>
: 90
</treatmentCitation>
, figs 416417 (morphological illustrations), 132 (diagnosis), Pl. 17: fig. 234 (habitus photograph);
<treatmentCitation id="191E66232754FFD7FD8BFD444A7D0521" author="Chen, S. - L. &amp; Li, Y. - G. &amp; Lin, S. - M. &amp; Lin, Q. - Y." box="[582,805,1639,1663]" page="47" pageId="46" pageNumber="47" year="1985">
<bibRefCitation id="FC2E3DC32754FFD7FD8BFD444A5A0521" author="Chen, S. - L. &amp; Li, Y. - G. &amp; Lin, S. - M. &amp; Lin, Q. - Y." box="[582,770,1639,1663]" pageId="46" pageNumber="47" pagination="45 - 51" refId="ref49031" refString="Chen, S. - L., Li, Y. - G., Lin, S. - M. &amp; Lin, Q. - Y. (1985) A list of Hemiptera Pentatomidae family of Fujian Province. Journal of Fujian College of Forestry, 5 (2), 45-51. [in Chinese, English summary]" type="journal article" year="1985">
Chen
<emphasis id="AACB9C202754FFD7FD4FFD4B4BED0521" box="[642,693,1639,1663]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
(1985)
</bibRefCitation>
: 47
</treatmentCitation>
48 (distribution);
<treatmentCitation id="191E66232754FFD7FC29FD4B4D270521" author="Hua, L. - Z." box="[996,1151,1639,1663]" page="43" pageId="46" pageNumber="47" year="1989">
<bibRefCitation id="FC2E3DC32754FFD7FC29FD4B4D040521" author="Hua, L. - Z." box="[996,1116,1639,1663]" pageId="46" pageNumber="47" refId="ref51800" refString="Hua, L. - Z. (1989) List of insect specimens of Zhongshan (Sun Yatsen) University. Taxonomy Division, Institute of Entomology, Zhongshan (Sun Yatsen) University, Guangzhou, ii + 158 pp. [in Chinese, English summary]" type="book" year="1989">Hua (1989)</bibRefCitation>
: 43
</treatmentCitation>
(list);
<treatmentCitation id="191E66232754FFD7FB0EFD444C2F0521" author="Zhang, S. - M." box="[1219,1399,1639,1663]" page="42" pageId="46" pageNumber="47" year="1995">
<bibRefCitation id="FC2E3DC32754FFD7FB0EFD444C090521" author="Zhang, S. - M." box="[1219,1361,1639,1663]" pageId="46" pageNumber="47" pagination="1 - 169" refId="ref57418" refString="Zhang, S. - M. (Ed.) (1995) Hemiptera (2). In: Economic insect fauna of China. Vol. 50. Science Press, Beijing, pp. i - xiii + 1-169, XXIV pls. [in Chinese]" type="book chapter" year="1995">Zhang (1995)</bibRefCitation>
: 42
</treatmentCitation>
, Pl. XII: fig. 114 (redescription, distribution, habitus illustration);
<treatmentCitation id="191E66232754FFD7FC8DFDA54AB105C0" author="Hua, L. - Z." box="[832,1001,1670,1694]" page="171" pageId="46" pageNumber="47" year="2000">
<bibRefCitation id="FC2E3DC32754FFD7FC8DFDA54AE005C0" author="Hua, L. - Z." box="[832,952,1670,1694]" pageId="46" pageNumber="47" pagination="162 - 216" refId="ref51850" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Huayu Nature Book Trade Co. Ltd., Guangdong, pp. 162-216. [Complete pagination: 448 pp.]" type="book chapter" year="2000">Hua (2000)</bibRefCitation>
: 171
</treatmentCitation>
(checklist, plant association, distribution);
<treatmentCitation id="191E66232754FFD7FF0AFD8648E005E3" author="Rider, D. A. &amp; Zheng, L. - Y. &amp; Kerzhner, I. M." box="[199,440,1701,1725]" page="137" pageId="46" pageNumber="47" year="2002">
<bibRefCitation id="FC2E3DC32754FFD7FF0AFD8648DF05E3" author="Rider, D. A. &amp; Zheng, L. - Y. &amp; Kerzhner, I. M." box="[199,391,1701,1725]" pageId="46" pageNumber="47" pagination="135 - 153" refId="ref54497" refString="Rider, D. A., Zheng, L. - Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135-153. https://doi.org/10.31610/zsr/2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="AACB9C202754FFD7FECBFD86486205E3" box="[262,314,1701,1725]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
(2002)
</bibRefCitation>
: 137
</treatmentCitation>
(checklist, distribution);
<treatmentCitation id="191E66232754FFD7FD77FD864A2805E2" author="Rider, D. A." box="[698,880,1700,1725]" page="260" pageId="46" pageNumber="47" year="2006">
<bibRefCitation id="FC2E3DC32754FFD7FD77FD864A1805E2" author="Rider, D. A." box="[698,832,1700,1725]" pageId="46" pageNumber="47" pagination="233 - 402" refId="ref54249" refString="Rider, D. A. (2006) Family Pentatomidae Leach, 1815. In: Aukema, B. &amp; Rieger, Ch. (Eds.), Catalogue of the Heteroptera of the Palaearctic Region. Vol. 5. Pentatomomorpha II. The Netherlands Entomological Society, Amsterdam, pp. 233-402. [book pagination: xiii + 550 pp.]" type="book chapter" year="2006">Rider (2006)</bibRefCitation>
: 260
</treatmentCitation>
(catalogue, distribution);
<treatmentCitation id="191E66232754FFD7FBB4FD864C4805E3" author="Fan, Z. - H." box="[1145,1296,1701,1725]" page="71" pageId="46" pageNumber="47" year="2011">
<bibRefCitation id="FC2E3DC32754FFD7FBB4FD864DB505E3" author="Fan, Z. - H." box="[1145,1261,1701,1725]" pageId="46" pageNumber="47" refId="ref50076" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Ph. D. thesis, Nankai University, Tianjin, 614 pp. [in Chinese, English summary]" type="book" year="2011">Fan (2011)</bibRefCitation>
: 71
</treatmentCitation>
73, figs 2.29 ag (key to species, redescription, drawings of male and female genitalia, distribution), 559560, figs 6263 (habitus photographs in dorsal and ventral view).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92754FFC9FF5AFC0B4B2B06BF" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="980040322754FFD7FF5AFC0B4B69041C" blockId="46.[151,1436,1832,2039]" box="[151,561,1832,1858]" pageId="46" pageNumber="47">
<materialsCitation id="28D74A6F2754FFD7FF5AFC0B4B75041C" box="[151,557,1832,1858]" country="China" location="Island" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<emphasis id="AACB9C202754FFD7FF5AFC0B486E041C" bold="true" box="[151,310,1832,1858]" pageId="46" pageNumber="47">
<typeStatus id="4704FE902754FFD7FF5AFC0B498A041C" box="[151,210,1832,1858]" pageId="46" pageNumber="47">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E0A800A22754FFD7FEF1FC0B48DA041C" box="[316,386,1832,1858]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
,
<collectingRegion id="5A7B8ED02754FFD7FE40FC0B4886041C" box="[397,478,1832,1858]" country="China" name="Hainan" pageId="46" pageNumber="47">Hainan</collectingRegion>
<location id="9D6016E92754FFD7FE28FC0B4B75041C" LSID="urn:lsid:plazi:treatment:1016F1242754FFCAFF5AFE464CD103A6:9D6016E92754FFD7FE28FC0B4B75041C" box="[485,557,1832,1858]" country="China" name="Island" pageId="46" pageNumber="47" stateProvince="Hainan">Island</location>
</materialsCitation>
.
</paragraph>
<paragraph id="980040322754FFD7FF0AFC6F4CC404A9" blockId="46.[151,1436,1832,2039]" pageId="46" pageNumber="47">
<emphasis id="AACB9C202754FFD7FF0AFC6F482C0438" bold="true" box="[199,372,1868,1894]" pageId="46" pageNumber="47">Type material.</emphasis>
<materialsCitation id="28D74A6F2754FFD7FEB7FC6F4A6A0438" box="[378,818,1868,1894]" country="China" location="Hainan" pageId="46" pageNumber="47" specimenCount="17" specimenCount-female="14" specimenCount-male="3" stateProvince="Hainan" typeStatus="syntype">
<typeStatus id="4704FE902754FFD7FEB7FC6F48B90438" box="[378,481,1868,1894]" pageId="46" pageNumber="47" type="syntype">Syntypes</typeStatus>
(
<specimenCount id="8EB98BBB2754FFD7FE3DFC6F4B4D0438" box="[496,533,1868,1894]" count="3" pageId="46" pageNumber="47" type="male">3♂</specimenCount>
,
<specimenCount id="8EB98BBB2754FFD7FDEDFC6F4B0B0438" box="[544,595,1868,1894]" count="14" pageId="46" pageNumber="47" type="female">14♀</specimenCount>
):
<emphasis id="AACB9C202754FFD7FDA5FC6F4A6A0438" bold="true" box="[616,818,1868,1894]" pageId="46" pageNumber="47">
<collectingCountry id="E0A800A22754FFD7FDA5FC6F4B920438" box="[616,714,1868,1894]" name="China" pageId="46" pageNumber="47">CHINA</collectingCountry>
:
<collectingRegion id="5A7B8ED02754FFD7FD18FC6F4A6A0438" box="[725,818,1868,1894]" country="China" name="Hainan" pageId="46" pageNumber="47">Hainan</collectingRegion>
</emphasis>
</materialsCitation>
<emphasis id="AACB9C202754FFD7FCFFFC6F4A6F0438" bold="true" box="[818,823,1868,1894]" pageId="46" pageNumber="47">:</emphasis>
<materialsCitation id="28D74A6F2754FFD7FCF0FC6F4CC004D4" collectionCode="BMNH" country="China" county="Type" location="Distant Coll." municipality="Dist." pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="2" stateProvince="Hainan" typeStatus="syntype">
<specimenCount id="8EB98BBB2754FFD7FCF0FC6F4A380438" box="[829,864,1868,1894]" count="1" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
, “
<collectingRegion id="5A7B8ED02754FFD7FCBAFC6F4AB00438" box="[887,1000,1868,1894]" country="China" name="Hainan" pageId="46" pageNumber="47">HAINAN</collectingRegion>
/ ISLAND [p] //
<collectingCounty id="716138BE2754FFD7FB69FC6E4D840439" box="[1188,1244,1869,1895]" pageId="46" pageNumber="47">Type</collectingCounty>
[p, red-margined disc // mitescens /
<collectingMunicipality id="7864DA482754FFD7FE93FC5348CD04D4" box="[350,405,1904,1930]" pageId="46" pageNumber="47">Dist.</collectingMunicipality>
[hw] //
<location id="9D6016E92754FFD7FE25FC534B2D04D4" LSID="urn:lsid:plazi:treatment:1016F1242754FFCAFF5AFE464CD103A6:9D6016E92754FFD7FE25FC534B2D04D4" box="[488,629,1904,1930]" country="China" county="Type" municipality="Dist." name="Distant Coll." pageId="46" pageNumber="47" stateProvince="Hainan">Distant Coll.</location>
/ 1911383 [p] //
<specimenCount id="8EB98BBB2754FFD7FCF8FC524A1204D5" box="[821,842,1905,1931]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
[p] // [QR code] / NHMUK010582892 [p]” (
<collectionCode id="FEAED8F72754FFD7FAFBFC534CC804D4" box="[1334,1424,1904,1930]" pageId="46" pageNumber="47">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="28D74A6F2754FFD7FF5AFCB64BBF04A8" collectionCode="BMNH" country="China" location="Distant Coll." municipality="Distant Coll." pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="2" stateProvince="Hainan" typeStatus="syntype">
<specimenCount id="8EB98BBB2754FFD7FF5AFCB649F404F1" box="[151,172,1941,1967]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
, “
<collectingRegion id="5A7B8ED02754FFD7FF0BFCB7486F04F0" box="[198,311,1940,1966]" country="China" name="Hainan" pageId="46" pageNumber="47">HAINAN</collectingRegion>
/ ISLAND [p] //
<typeStatus id="4704FE902754FFD7FDC9FCB74BDA04F0" box="[516,642,1940,1966]" pageId="46" pageNumber="47" type="syntype">SYNTYPE</typeStatus>
[p, blue-margined disc] //
<collectingMunicipality id="7864DA482754FFD7FC75FCB74D1004F0" box="[952,1096,1940,1966]" pageId="46" pageNumber="47">Distant Coll.</collectingMunicipality>
/ 1911383 [p]” (
<collectionCode id="FEAED8F72754FFD7FADDFCB74C3204F0" box="[1296,1386,1940,1966]" pageId="46" pageNumber="47">BMNH</collectionCode>
);
<specimenCount id="8EB98BBB2754FFD7FA4DFCB64CCD04F1" box="[1408,1429,1941,1967]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
,
<collectingRegion id="5A7B8ED02754FFD7FF5AFC9B485D048C" box="[151,261,1976,2002]" country="China" name="Hainan" pageId="46" pageNumber="47">HAINAN</collectingRegion>
/ ISLAND [p] //
<typeStatus id="4704FE902754FFD7FE18FC9B4B0B048C" box="[469,595,1976,2002]" pageId="46" pageNumber="47" type="syntype">SYNTYPE</typeStatus>
[p, blue-margined disc] //
<taxonomicName id="5FBF3BB12754FFD7FC47FC9B4D8E048C" authority="Distant" authorityName="Distant" authorityYear="1901" box="[906,1238,1976,2002]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="mitescens">
<emphasis id="AACB9C202754FFD7FC47FC9B4D24048C" box="[906,1148,1976,2002]" italics="true" pageId="46" pageNumber="47">Axiagastus mitescens</emphasis>
Distant
</taxonomicName>
//
<location id="9D6016E92754FFD7FB34FC9B4CD3048C" LSID="urn:lsid:plazi:treatment:1016F1242754FFCAFF5AFE464CD103A6:9D6016E92754FFD7FB34FC9B4CD3048C" box="[1273,1419,1976,2002]" country="China" municipality="Distant Coll." name="Distant Coll." pageId="46" pageNumber="47" stateProvince="Hainan">Distant Coll.</location>
/ 1911383 [p] // NHMUK 013589098 [p]” (
<collectionCode id="FEAED8F72754FFD7FD48FCFF4B8704A8" box="[645,735,2012,2038]" pageId="46" pageNumber="47">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="28D74A6F2754FFD7FD39FCFF4C2704A8" box="[756,1407,2012,2039]" country="China" location="Distant Coll." pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="syntype">
<specimenCount id="8EB98BBB2754FFD7FD39FCFF4A4F04A8" box="[756,791,2012,2038]" count="1" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
, “
<collectingRegion id="5A7B8ED02754FFD7FCFDFCFF4ADB04A8" box="[816,899,2012,2038]" country="China" name="Hainan" pageId="46" pageNumber="47">Hainan</collectingRegion>
[hw] //
<location id="9D6016E92754FFD7FC2CFCFF4D2A04A8" LSID="urn:lsid:plazi:treatment:1016F1242754FFCAFF5AFE464CD103A6:9D6016E92754FFD7FC2CFCFF4D2A04A8" box="[993,1138,2012,2038]" country="China" name="Distant Coll." pageId="46" pageNumber="47" stateProvince="Hainan">Distant Coll.</location>
/ B. M. 1911-383 [p] //
</materialsCitation>
<specimenCount id="8EB98BBB2754FFD7FA4AFCFE4CC404A9" box="[1415,1436,2013,2039]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
</paragraph>
<caption id="CCC010BA2755FFD6FF5AFC1B4AEC04C6" pageId="47" pageNumber="48" startId="47.[151,264,1848,1872]" targetBox="[151,1436,181,1824]" targetPageId="47" targetType="figure">
<paragraph id="980040322755FFD6FF5AFC1B4AEC04C6" blockId="47.[151,1437,1848,1944]" pageId="47" pageNumber="48">
<emphasis id="AACB9C202755FFD6FF5AFC1B4837040E" bold="true" box="[151,367,1848,1872]" pageId="47" pageNumber="48">FIGURES 134137.</emphasis>
<taxonomicName id="5FBF3BB12755FFD6FEBAFC1B4BBF040E" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[375,743,1848,1872]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="mitescens">
<emphasis id="AACB9C202755FFD6FEBAFC1B4B08040E" box="[375,592,1848,1872]" italics="true" pageId="47" pageNumber="48">Axiagastus mitescens</emphasis>
<bibRefCitation id="FC2E3DC32755FFD6FD94FC1B4BBF040E" author="Distant, W. L." box="[601,743,1848,1872]" pageId="47" pageNumber="48" pagination="581 - 592" refId="ref49383" refString="Distant, W. L. (1901) Contributions to a knowledge of the Rhynchota. Transactions of the Entomological Society of London, 1901 (4), 581-592, pl. 16. https://doi.org/10.1111/j.1365-2311.1901.tb01374.x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
. 134135—female habitus (134—dorsal view, 135—ventral view). Orig. J.-F. Tsai (NMNS), © The Trustees of the Natural History Museum, London; 136137—male habitus (136—dorsal view, 137—ventral view). Scale: 2 mm (134135), 1 mm (136137). Orig. S. Salini.
</paragraph>
</caption>
<caption id="CCC010BA274AFFC9FF5AFF8A4C260657" pageId="48" pageNumber="49" startId="48.[151,264,1193,1217]" targetBox="[151,1436,181,1169]" targetPageId="48" targetType="figure">
<paragraph id="98004032274AFFC9FF5AFF8A4C260657" blockId="48.[151,1436,1193,1289]" pageId="48" pageNumber="49">
<emphasis id="AACB9C20274AFFC9FF5AFF8A482A079F" bold="true" box="[151,370,1193,1217]" pageId="48" pageNumber="49">FIGURES 138141.</emphasis>
<taxonomicName id="5FBF3BB1274AFFC9FEB0FF8A4BAD079F" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[381,757,1193,1217]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="mitescens">
<emphasis id="AACB9C20274AFFC9FEB0FF8A4B01079F" box="[381,601,1193,1217]" italics="true" pageId="48" pageNumber="49">Axiagastus mitescens</emphasis>
<bibRefCitation id="FC2E3DC3274AFFC9FDA8FF8A4BAD079F" author="Distant, W. L." box="[613,757,1193,1217]" pageId="48" pageNumber="49" pagination="581 - 592" refId="ref49383" refString="Distant, W. L. (1901) Contributions to a knowledge of the Rhynchota. Transactions of the Entomological Society of London, 1901 (4), 581-592, pl. 16. https://doi.org/10.1111/j.1365-2311.1901.tb01374.x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
. 138—head (dorsal view); 139—external scent efferent system; 140—apex of male abdomen with genital capsule in postero-ventral view; 141—apex of female abdomen with terminalia in posteroventral view. Lettering: dn—distension, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (138141). Orig. S. Salini.
</paragraph>
</caption>
<paragraph id="98004032274AFFC9FF5AFE784B2B06BF" blockId="48.[151,1437,1371,2010]" pageId="48" pageNumber="49">
[p]” (
<collectionCode id="FEAED8F7274AFFC9FF18FE784877062B" box="[213,303,1371,1397]" pageId="48" pageNumber="49">BMNH</collectionCode>
);
<materialsCitation id="28D74A6F274AFFC9FE88FE784DA6062B" box="[325,1278,1371,1398]" collectionCode="BMNH" country="Iceland" location="Hainan" municipality="Distant Coll." pageId="48" pageNumber="49" specimenCount="2" specimenCount-female="2">
<specimenCount id="8EB98BBB274AFFC9FE88FE784830062B" box="[325,360,1371,1397]" count="1" pageId="48" pageNumber="49" type="female">1♀</specimenCount>
, “
<location id="9D6016E9274AFFC9FE4FFE78488D062B" LSID="urn:lsid:plazi:treatment:1016F1242754FFCAFF5AFE464CD103A6:9D6016E9274AFFC9FE4FFE78488D062B" box="[386,469,1371,1397]" country="Iceland" municipality="Distant Coll." name="Hainan" pageId="48" pageNumber="49">Hainan</location>
/
<collectingCountry id="E0A800A2274AFFC9FE22FE784B6B062B" box="[495,563,1371,1397]" name="Iceland" pageId="48" pageNumber="49">Island</collectingCountry>
[hw] //
<collectingMunicipality id="7864DA48274AFFC9FD5EFE784A7D062B" box="[659,805,1371,1397]" pageId="48" pageNumber="49">Distant Coll.</collectingMunicipality>
/ B. M. 1911-383 [p] //
<specimenCount id="8EB98BBB274AFFC9FB8DFE7F4D0D0628" box="[1088,1109,1372,1398]" count="1" pageId="48" pageNumber="49" type="female"></specimenCount>
[p]” (
<collectionCode id="FEAED8F7274AFFC9FB51FE784DAE062B" box="[1180,1270,1371,1397]" pageId="48" pageNumber="49">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="28D74A6F274AFFC9FAC1FE784BC506C7" collectionCode="BMNH" country="Iceland" location="Distant Coll." pageId="48" pageNumber="49" specimenCount="10" specimenCount-female="10">
<specimenCount id="8EB98BBB274AFFC9FAC1FE784C68062B" box="[1292,1328,1371,1397]" count="9" pageId="48" pageNumber="49" type="female">9♀</specimenCount>
, “
<location id="9D6016E9274AFFC9FA84FE78499706C7" LSID="urn:lsid:plazi:treatment:1016F1242754FFCAFF5AFE464CD103A6:9D6016E9274AFFC9FA84FE78499706C7" country="Iceland" name="Distant Coll." pageId="48" pageNumber="49">Distant Coll.</location>
/ B. M. 1911-383 [p] //
<specimenCount id="8EB98BBB274AFFC9FE2CFEA348AE06C4" box="[481,502,1408,1434]" count="1" pageId="48" pageNumber="49" type="female"></specimenCount>
[p]” (
<collectionCode id="FEAED8F7274AFFC9FDF7FE5C4BCC06C7" box="[570,660,1407,1433]" pageId="48" pageNumber="49">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="28D74A6F274AFFC9FD64FE5C4C1906C7" box="[681,1345,1407,1434]" collectionCode="BMNH" country="Iceland" location="Distant Coll." pageId="48" pageNumber="49" specimenCount="4" specimenCount-male="4">
<specimenCount id="8EB98BBB274AFFC9FD64FE5C4B9606C7" box="[681,718,1407,1433]" count="3" pageId="48" pageNumber="49" type="male">3♂</specimenCount>
, “
<location id="9D6016E9274AFFC9FD29FE5C4A2F06C7" LSID="urn:lsid:plazi:treatment:1016F1242754FFCAFF5AFE464CD103A6:9D6016E9274AFFC9FD29FE5C4A2F06C7" box="[740,887,1407,1433]" country="Iceland" name="Distant Coll." pageId="48" pageNumber="49">Distant Coll.</location>
/ B. M. 1911-383 [p] //
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[p]” (
<collectionCode id="FEAED8F7274AFFC9FB2FFE5C4C6106C7" box="[1250,1337,1407,1433]" pageId="48" pageNumber="49">BMNH</collectionCode>
)
</materialsCitation>
.—
<materialsCitation id="28D74A6F274AFFC9FAAFFE5C4B3706BF" collectorName="Salini, S. &amp; Kment, P." country="Iceland" location="Each" pageId="48" pageNumber="49" specimenCount="1" typeStatus="syntype">
<location id="9D6016E9274AFFC9FAAFFE5C4CC406C7" LSID="urn:lsid:plazi:treatment:1016F1242754FFCAFF5AFE464CD103A6:9D6016E9274AFFC9FAAFFE5C4CC406C7" box="[1378,1436,1407,1433]" country="Iceland" name="Each" pageId="48" pageNumber="49">Each</location>
specimen bears the following identification label: “
<typeStatus id="4704FE90274AFFC9FD03FE804A3906E3" box="[718,865,1443,1469]" pageId="48" pageNumber="49" type="syntype">SYNTYPUS</typeStatus>
/
<taxonomicName id="5FBF3BB1274AFFC9FCB8FE874DE906E0" authorityName="Distant" authorityYear="1901" box="[885,1201,1444,1470]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="3" phylum="Arthropoda" rank="species" species="mitescens">
<emphasis id="AACB9C20274AFFC9FCB8FE874DE906E0" box="[885,1201,1444,1470]" italics="true" pageId="48" pageNumber="49">AXIAGASTUS MITESCENS</emphasis>
</taxonomicName>
/
<bibRefCitation id="FC2E3DC3274AFFC9FB08FE804C0406E3" author="Distant, W. L." box="[1221,1372,1443,1469]" pageId="48" pageNumber="49" pagination="581 - 592" refId="ref49383" refString="Distant, W. L. (1901) Contributions to a knowledge of the Rhynchota. Transactions of the Entomological Society of London, 1901 (4), 581-592, pl. 16. https://doi.org/10.1111/j.1365-2311.1901.tb01374.x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
/ des.
<collectorName id="354A25E4274AFFC9FF5AFEE449A506BF" box="[151,253,1479,1505]" pageId="48" pageNumber="49">Salini, S.</collectorName>
&amp;
<collectorName id="354A25E4274AFFC9FEECFEEB48C906BC" box="[289,401,1480,1506]" pageId="48" pageNumber="49">Kment, P.</collectorName>
, 2024 [p, red label]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B9274AFFCBFF0AFEC84B1E0578" lastPageId="50" lastPageNumber="51" pageId="48" pageNumber="49" type="description">
<paragraph id="98004032274AFFC9FF0AFEC84C620487" blockId="48.[151,1437,1371,2010]" pageId="48" pageNumber="49">
<emphasis id="AACB9C20274AFFC9FF0AFEC84A51055B" bold="true" box="[199,777,1515,1542]" pageId="48" pageNumber="49">
Redescription.
<emphasis id="AACB9C20274AFFC9FE4DFEC84A51055B" bold="true" box="[384,777,1515,1542]" italics="true" pageId="48" pageNumber="49">Colour, integument and vestiture</emphasis>
</emphasis>
(
<figureCitation id="00845CB7274AFFC9FCD1FEC84A98055B" box="[796,960,1515,1541]" captionStart-0="FIGURES 134137" captionStart-1="FIGURES 138141" captionStartId-0="47.[151,264,1848,1872]" captionStartId-1="48.[151,264,1193,1217]" captionTargetBox-0="[151,1436,181,1824]" captionTargetBox-1="[151,1436,181,1169]" captionTargetId-0="figure-19@47.[151,1436,181,1824]" captionTargetId-1="figure-29@48.[151,1436,181,1169]" captionTargetPageId-0="47" captionTargetPageId-1="48" captionText-0="FIGURES 134137. Axiagastus mitescens Distant, 1901. 134135—female habitus (134—dorsal view, 135—ventral view). Orig. J.-F. Tsai (NMNS), © The Trustees of the Natural History Museum, London; 136137—male habitus (136—dorsal view, 137—ventral view). Scale: 2 mm (134135), 1 mm (136137). Orig. S. Salini." captionText-1="FIGURES 138141. Axiagastus mitescens Distant, 1901. 138—head (dorsal view); 139—external scent efferent system; 140—apex of male abdomen with genital capsule in postero-ventral view; 141—apex of female abdomen with terminalia in posteroventral view. Lettering: dn—distension, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (138141). Orig. S. Salini." pageId="48" pageNumber="49">Figs 134139</figureCitation>
). Body above (except hemelytra) orange yellow, with following well-expressed black markings: longitudinal stripes on disc of head along lateral margins of clypeus, extending to posterior margin of head, anterolateral margins of pronotum, transverse (sometimes broken) stripe posteriorly on pronotal disc, 1+1 moderately large, submedian spots at one-third from base of scutellar disc, broad lunulate median spot at one-third from apex of scutellar disc; disc of head dorsally with fine, black punctures arranged in one or two longitudinal rows, disc of pronotum with coarse, black punctures sparsely distributed, sometimes coalescent to form small, black spots, especially on anterior pronotal disc; ground colour of each clavus and corium yellowish, with dense, coarse, black punctures arranged on each corium, punctures finer on exocorium; scape pale, concolourous with body, remaining antennomeres black; connexiva concolourous with body; membranes smoky brown; ventral surface of body concolourous with dorsum, head on ventral surface impunctate; black, coarse punctures on thoracic sternites; ventrites impunctate; apices of ostiolar peritremes, labium, and tarsal claws, a moderately large spot coalescent with each spiracle anteriorly, a row of small spots mesad to lateral margins of abdomen, small denticle on posterolateral angle of each abdominal segment, black; legs concolourous with ventral surface of body, impunctate, sometimes with one or two small, black, more or less round spots on femora.
</paragraph>
<caption id="CCC010BA274BFFC8FF5AFF7A4D1C0783" pageId="49" pageNumber="50" startId="49.[151,264,1113,1137]" targetBox="[151,1436,181,1089]" targetPageId="49" targetType="figure">
<paragraph id="98004032274BFFC8FF5AFF7A4D1C0783" blockId="49.[151,1437,1113,1246]" pageId="49" pageNumber="50">
<emphasis id="AACB9C20274BFFC8FF5AFF7A4836072F" bold="true" box="[151,366,1113,1137]" pageId="49" pageNumber="50">FIGURES 142148.</emphasis>
<taxonomicName id="5FBF3BB1274BFFC8FEB9FF794B86072C" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[372,734,1113,1138]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="mitescens">
<emphasis id="AACB9C20274BFFC8FEB9FF794B14072F" box="[372,588,1114,1137]" italics="true" pageId="49" pageNumber="50">Axiagastus mitescens</emphasis>
<bibRefCitation id="FC2E3DC3274BFFC8FD9EFF7A4B86072C" author="Distant, W. L." box="[595,734,1113,1138]" pageId="49" pageNumber="50" pagination="581 - 592" refId="ref49383" refString="Distant, W. L. (1901) Contributions to a knowledge of the Rhynchota. Transactions of the Entomological Society of London, 1901 (4), 581-592, pl. 16. https://doi.org/10.1111/j.1365-2311.1901.tb01374.x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
, male genitalia. 142145—genital capsule (142—dorsal view, 143— ventral view, 144—caudal view, 145—lateral view); 146148—parameres (three different views). Lettering: a—angulation, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, sr—serrations, te—transverse emargination, vr—ventral rim. Scale: 0.5 mm (142145), 0.1 mm (146148). Orig. S. Salini.
</paragraph>
</caption>
<paragraph id="98004032274BFFC8FF0AFE134D990614" blockId="49.[151,1437,1328,2038]" box="[199,1217,1328,1354]" pageId="49" pageNumber="50">
<emphasis id="AACB9C20274BFFC8FF0AFE1348630614" bold="true" box="[199,315,1328,1354]" italics="true" pageId="49" pageNumber="50">Structure.</emphasis>
Labium short, reaching metacoxae. Other characters as in generic redescription.
</paragraph>
<paragraph id="98004032274BFFC8FF0AFE704B3904A8" blockId="49.[151,1437,1328,2038]" pageId="49" pageNumber="50">
<emphasis id="AACB9C20274BFFC8FF0AFE70482A0630" bold="true" box="[199,370,1363,1390]" italics="true" pageId="49" pageNumber="50">Male genitalia</emphasis>
(
<figureCitation id="00845CB7274BFFC8FE48FE7748B60630" box="[389,494,1364,1390]" captionStart="FIGURES 138141" captionStartId="48.[151,264,1193,1217]" captionTargetBox="[151,1436,181,1169]" captionTargetId="figure-29@48.[151,1436,181,1169]" captionTargetPageId="48" captionText="FIGURES 138141. Axiagastus mitescens Distant, 1901. 138—head (dorsal view); 139—external scent efferent system; 140—apex of male abdomen with genital capsule in postero-ventral view; 141—apex of female abdomen with terminalia in posteroventral view. Lettering: dn—distension, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (138141). Orig. S. Salini." pageId="49" pageNumber="50">Figs 140</figureCitation>
,
<figureCitation id="00845CB7274BFFC8FE33FE704B3D0630" box="[510,613,1363,1390]" captionStart-0="FIGURES 142148" captionStart-1="FIGURES 149154" captionStartId-0="49.[151,264,1113,1137]" captionStartId-1="50.[151,264,1168,1192]" captionTargetBox-0="[151,1436,181,1089]" captionTargetBox-1="[151,1436,181,1145]" captionTargetId-0="figure-19@49.[151,1436,181,1089]" captionTargetId-1="figure-29@50.[151,1436,181,1145]" captionTargetPageId-0="49" captionTargetPageId-1="50" captionText-0="FIGURES 142148. Axiagastus mitescens Distant, 1901, male genitalia. 142145—genital capsule (142—dorsal view, 143— ventral view, 144—caudal view, 145—lateral view); 146148—parameres (three different views). Lettering: a—angulation, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, sr—serrations, te—transverse emargination, vr—ventral rim. Scale: 0.5 mm (142145), 0.1 mm (146148). Orig. S. Salini." captionText-1="FIGURES 149154. Axiagastus mitescens Distant, 1901, male and female genitalia. 149151—phallus (149—dorsal view, 150—ventral view, 151—lateral view); 152—terminalia (dorsal); 153—terminalia (ventral); 154—spermatheca. Lettering: ad—aedeagus, cp—conjunctival processes, pa—processes of aedeagus, vcp—ventral convex projection. Scale: 0.5 mm (149 154). Orig. S. Salini." pageId="49" pageNumber="50">142151</figureCitation>
). Genital capsule (
<figureCitation id="00845CB7274BFFC8FCF2FE774A870630" box="[831,991,1363,1390]" captionStart="FIGURES 142148" captionStartId="49.[151,264,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-19@49.[151,1436,181,1089]" captionTargetPageId="49" captionText="FIGURES 142148. Axiagastus mitescens Distant, 1901, male genitalia. 142145—genital capsule (142—dorsal view, 143— ventral view, 144—caudal view, 145—lateral view); 146148—parameres (three different views). Lettering: a—angulation, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, sr—serrations, te—transverse emargination, vr—ventral rim. Scale: 0.5 mm (142145), 0.1 mm (146148). Orig. S. Salini." pageId="49" pageNumber="50">Figs 142145</figureCitation>
) subquadrate, dorsal rim more deeply incised than ventral rim; lateral wall of dorsal rim slightly concave, few minute, indistinct serrations (sr), appearing sclerotized black, ending in moderately developed angulation contiguous, with deep, transverse emargination (te) on middle of dorsal rim; dorsal sinus of posterior aperture broadly U-shaped (ds), ventral margin of posterior aperture semispherical; setae bordering ventral rim absent, unlike in other species, moderately developed setae on posterolateral lobes, and short fine setae mesad to median distension; ventral rim moderately concave at middle (vr), slightly concave sublateral to posterolateral lobes; posterolateral lobes with minute angulations, in lateral view; infoldings of ventral rim deeply impressed on either side of moderately developed distension at middle, distension (dn) sunken, with short emarginate V-shaped incision at middle, visible on dorsal view of genital capsule; infoldings of ventral rim laterally (inner to posterolateral lobes) with a prominent, obtuse or blunt, sclerotized, black denticle (dt).
<emphasis id="AACB9C20274BFFC8FF03FD9F4865058B" box="[206,317,1724,1749]" italics="true" pageId="49" pageNumber="50">Paramere</emphasis>
(
<figureCitation id="00845CB7274BFFC8FE80FD9F48B20588" box="[333,490,1724,1750]" captionStart="FIGURES 142148" captionStartId="49.[151,264,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-19@49.[151,1436,181,1089]" captionTargetPageId="49" captionText="FIGURES 142148. Axiagastus mitescens Distant, 1901, male genitalia. 142145—genital capsule (142—dorsal view, 143— ventral view, 144—caudal view, 145—lateral view); 146148—parameres (three different views). Lettering: a—angulation, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, sr—serrations, te—transverse emargination, vr—ventral rim. Scale: 0.5 mm (142145), 0.1 mm (146148). Orig. S. Salini." pageId="49" pageNumber="50">Figs 146148</figureCitation>
) simple, crown finger-like, as in
<taxonomicName id="5FBF3BB1274BFFC8FCACFD9F4AA70588" authorityName="Kment &amp; Lemaître &amp; Webb &amp; Roca-Cusachs" authorityYear="2025" box="[865,1023,1724,1750]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="yeshwanthi" status="sp. nov.">
<emphasis id="AACB9C20274BFFC8FCACFD9F4AA70588" box="[865,1023,1724,1750]" italics="true" pageId="49" pageNumber="50">A. yeshwanthi</emphasis>
</taxonomicName>
<emphasis id="AACB9C20274BFFC8FBCBFD9F4D300588" bold="true" box="[1030,1128,1724,1750]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="B1F8215B274BFFC8FBCBFD9F4D3A0588" box="[1030,1122,1724,1750]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
with dorsal margin convex at one-third in lateral view (
<figureCitation id="00845CB7274BFFC8FE1FFDC34B2905A4" box="[466,625,1760,1786]" captionStart="FIGURES 142148" captionStartId="49.[151,264,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-19@49.[151,1436,181,1089]" captionTargetPageId="49" captionText="FIGURES 142148. Axiagastus mitescens Distant, 1901, male genitalia. 142145—genital capsule (142—dorsal view, 143— ventral view, 144—caudal view, 145—lateral view); 146148—parameres (three different views). Lettering: a—angulation, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, sr—serrations, te—transverse emargination, vr—ventral rim. Scale: 0.5 mm (142145), 0.1 mm (146148). Orig. S. Salini." pageId="49" pageNumber="50">Figs 146, 147</figureCitation>
); finger-like crown on inner side with numerous fine, short setae, single moderately elongate setae on dorsal side; elongate setae along periphery of laminate disc (ld); crown gradually narrowed towards apex; apex rounded, not acuminate in lateral view (
<figureCitation id="00845CB7274BFFC8FC74FC0B4D03041C" box="[953,1115,1832,1858]" captionStart="FIGURES 142148" captionStartId="49.[151,264,1113,1137]" captionTargetBox="[151,1436,181,1089]" captionTargetId="figure-19@49.[151,1436,181,1089]" captionTargetPageId="49" captionText="FIGURES 142148. Axiagastus mitescens Distant, 1901, male genitalia. 142145—genital capsule (142—dorsal view, 143— ventral view, 144—caudal view, 145—lateral view); 146148—parameres (three different views). Lettering: a—angulation, dn—distension, ds—U-shaped posterior aperture of dorsal sinus, dt—denticle, ld—laminate disc, sr—serrations, te—transverse emargination, vr—ventral rim. Scale: 0.5 mm (142145), 0.1 mm (146148). Orig. S. Salini." pageId="49" pageNumber="50">Figs 146, 147</figureCitation>
); stem moderately elongate, apodeme disc-like.
<emphasis id="AACB9C20274BFFC8FEBDFC6F489E0438" box="[368,454,1868,1894]" italics="true" pageId="49" pageNumber="50">Phallus</emphasis>
(
<figureCitation id="00845CB7274BFFC8FE19FC6F4B2A0438" box="[468,626,1867,1894]" captionStart="FIGURES 149154" captionStartId="50.[151,264,1168,1192]" captionTargetBox="[151,1436,181,1145]" captionTargetId="figure-29@50.[151,1436,181,1145]" captionTargetPageId="50" captionText="FIGURES 149154. Axiagastus mitescens Distant, 1901, male and female genitalia. 149151—phallus (149—dorsal view, 150—ventral view, 151—lateral view); 152—terminalia (dorsal); 153—terminalia (ventral); 154—spermatheca. Lettering: ad—aedeagus, cp—conjunctival processes, pa—processes of aedeagus, vcp—ventral convex projection. Scale: 0.5 mm (149 154). Orig. S. Salini." pageId="49" pageNumber="50">Figs 149151</figureCitation>
). Articulatory apparatus as in
<figureCitation id="00845CB7274BFFC8FC73FC6F4D010438" box="[958,1113,1867,1894]" captionStart="FIGURES 149154" captionStartId="50.[151,264,1168,1192]" captionTargetBox="[151,1436,181,1145]" captionTargetId="figure-29@50.[151,1436,181,1145]" captionTargetPageId="50" captionText="FIGURES 149154. Axiagastus mitescens Distant, 1901, male and female genitalia. 149151—phallus (149—dorsal view, 150—ventral view, 151—lateral view); 152—terminalia (dorsal); 153—terminalia (ventral); 154—spermatheca. Lettering: ad—aedeagus, cp—conjunctival processes, pa—processes of aedeagus, vcp—ventral convex projection. Scale: 0.5 mm (149 154). Orig. S. Salini." pageId="49" pageNumber="50">Figs 149151</figureCitation>
; phallotheca less sclerotized, distal part transparent, with ventral convex projection (vcp) in close proximity with processes of aedeagus (pa); a pair of dorsal conjunctival processes (cp), partly sclerotized throughout length, apically rounded; aedeagus (ad) short, deflected dorsad, apex swollen, drop-like, transparent, with embedded phallotreme, processes of aedeagus much elongate and as shown in
<figureCitation id="00845CB7274BFFC8FE30FCFF4B0504A8" box="[509,605,2011,2038]" captionStart="FIGURES 149154" captionStartId="50.[151,264,1168,1192]" captionTargetBox="[151,1436,181,1145]" captionTargetId="figure-29@50.[151,1436,181,1145]" captionTargetPageId="50" captionText="FIGURES 149154. Axiagastus mitescens Distant, 1901, male and female genitalia. 149151—phallus (149—dorsal view, 150—ventral view, 151—lateral view); 152—terminalia (dorsal); 153—terminalia (ventral); 154—spermatheca. Lettering: ad—aedeagus, cp—conjunctival processes, pa—processes of aedeagus, vcp—ventral convex projection. Scale: 0.5 mm (149 154). Orig. S. Salini." pageId="49" pageNumber="50">Fig. 151</figureCitation>
.
</paragraph>
<caption id="CCC010BA2748FFCBFF5AFFB34832064A" pageId="50" pageNumber="51" startId="50.[151,264,1168,1192]" targetBox="[151,1436,181,1145]" targetPageId="50" targetType="figure">
<paragraph id="980040322748FFCBFF5AFFB34832064A" blockId="50.[151,1437,1168,1300]" pageId="50" pageNumber="51">
<emphasis id="AACB9C202748FFCBFF5AFFB3482807F7" bold="true" box="[151,368,1168,1193]" pageId="50" pageNumber="51">FIGURES 149154.</emphasis>
<taxonomicName id="5FBF3BB12748FFCBFEB4FFB24BB207F7" authority="Distant, 1901" authorityName="Distant" authorityYear="1901" box="[377,746,1169,1193]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mitescens">
<emphasis id="AACB9C202748FFCBFEB4FFB24B0B07F6" box="[377,595,1169,1192]" italics="true" pageId="50" pageNumber="51">Axiagastus mitescens</emphasis>
<bibRefCitation id="FC2E3DC32748FFCBFD90FFB24BB207F7" author="Distant, W. L." box="[605,746,1169,1193]" pageId="50" pageNumber="51" pagination="581 - 592" refId="ref49383" refString="Distant, W. L. (1901) Contributions to a knowledge of the Rhynchota. Transactions of the Entomological Society of London, 1901 (4), 581-592, pl. 16. https://doi.org/10.1111/j.1365-2311.1901.tb01374.x" type="journal article" year="1901">Distant, 1901</bibRefCitation>
</taxonomicName>
, male and female genitalia. 149151—phallus (149—dorsal view, 150—ventral view, 151—lateral view); 152—terminalia (dorsal); 153—terminalia (ventral); 154—spermatheca. Lettering: ad—aedeagus, cp—conjunctival processes, pa—processes of aedeagus, vcp—ventral convex projection. Scale: 0.5 mm (149 154). Orig. S. Salini.
</paragraph>
</caption>
<paragraph id="980040322748FFCBFF0AFE7B4B1E0578" blockId="50.[151,1437,1368,2043]" pageId="50" pageNumber="51">
<emphasis id="AACB9C202748FFCBFF0AFE7B48D7062D" bold="true" box="[199,399,1368,1395]" italics="true" pageId="50" pageNumber="51">Female genitalia</emphasis>
(
<figureCitation id="00845CB72748FFCBFE6FFE7B4B53062D" box="[418,523,1368,1395]" captionStart="FIGURES 138141" captionStartId="48.[151,264,1193,1217]" captionTargetBox="[151,1436,181,1169]" captionTargetId="figure-29@48.[151,1436,181,1169]" captionTargetPageId="48" captionText="FIGURES 138141. Axiagastus mitescens Distant, 1901. 138—head (dorsal view); 139—external scent efferent system; 140—apex of male abdomen with genital capsule in postero-ventral view; 141—apex of female abdomen with terminalia in posteroventral view. Lettering: dn—distension, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (138141). Orig. S. Salini." pageId="50" pageNumber="51">Figs 141</figureCitation>
,
<figureCitation id="00845CB72748FFCBFDD6FE7B4BDA062D" box="[539,642,1368,1395]" captionStart="FIGURES 149154" captionStartId="50.[151,264,1168,1192]" captionTargetBox="[151,1436,181,1145]" captionTargetId="figure-29@50.[151,1436,181,1145]" captionTargetPageId="50" captionText="FIGURES 149154. Axiagastus mitescens Distant, 1901, male and female genitalia. 149151—phallus (149—dorsal view, 150—ventral view, 151—lateral view); 152—terminalia (dorsal); 153—terminalia (ventral); 154—spermatheca. Lettering: ad—aedeagus, cp—conjunctival processes, pa—processes of aedeagus, vcp—ventral convex projection. Scale: 0.5 mm (149 154). Orig. S. Salini." pageId="50" pageNumber="51">152154</figureCitation>
).
<emphasis id="AACB9C202748FFCBFD54FE7B4A4B062C" box="[665,787,1368,1394]" italics="true" pageId="50" pageNumber="51">Terminalia</emphasis>
(
<figureCitation id="00845CB72748FFCBFCEBFE7B4AD7062D" box="[806,911,1368,1395]" captionStart="FIGURES 138141" captionStartId="48.[151,264,1193,1217]" captionTargetBox="[151,1436,181,1169]" captionTargetId="figure-29@48.[151,1436,181,1169]" captionTargetPageId="48" captionText="FIGURES 138141. Axiagastus mitescens Distant, 1901. 138—head (dorsal view); 139—external scent efferent system; 140—apex of male abdomen with genital capsule in postero-ventral view; 141—apex of female abdomen with terminalia in posteroventral view. Lettering: dn—distension, o—ostiole, p—ostiolar peritreme. Scale: 0.5 mm (138141). Orig. S. Salini." pageId="50" pageNumber="51">Figs 141</figureCitation>
,
<figureCitation id="00845CB72748FFCBFC6DFE7B4D52062D" box="[928,1034,1368,1395]" captionStart="FIGURES 149154" captionStartId="50.[151,264,1168,1192]" captionTargetBox="[151,1436,181,1145]" captionTargetId="figure-29@50.[151,1436,181,1145]" captionTargetPageId="50" captionText="FIGURES 149154. Axiagastus mitescens Distant, 1901, male and female genitalia. 149151—phallus (149—dorsal view, 150—ventral view, 151—lateral view); 152—terminalia (dorsal); 153—terminalia (ventral); 154—spermatheca. Lettering: ad—aedeagus, cp—conjunctival processes, pa—processes of aedeagus, vcp—ventral convex projection. Scale: 0.5 mm (149 154). Orig. S. Salini." pageId="50" pageNumber="51">152, 153</figureCitation>
). Valvifers VIII as in
<taxonomicName id="5FBF3BB12748FFCBFADCFE7A4CCC062C" authorityName="Distant" authorityYear="1911" box="[1297,1428,1368,1395]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="cambelli">
<emphasis id="AACB9C202748FFCBFADCFE7A4CCC062C" box="[1297,1428,1368,1395]" italics="true" pageId="50" pageNumber="51">A. cambelli</emphasis>
</taxonomicName>
; valvifers IX short, narrow, transverse plate, anterior margin of each angulate at middle, posterior margins slightly concave; laterotergites VIII and IX as in
<emphasis id="AACB9C202748FFCBFDAAFE824A2F06E5" box="[615,887,1441,1467]" italics="true" pageId="50" pageNumber="51">
<taxonomicName id="5FBF3BB12748FFCBFDAAFE824BAB06E4" authorityName="Dallas" authorityYear="1851" box="[615,755,1441,1467]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rosmarus">A. rosmarus</taxonomicName>
. Gynatrium
</emphasis>
as in
<emphasis id="AACB9C202748FFCBFC70FE824DBE06E4" box="[957,1254,1440,1467]" italics="true" pageId="50" pageNumber="51">
<taxonomicName id="5FBF3BB12748FFCBFC70FE824D1106E4" authorityName="Dallas" authorityYear="1851" box="[957,1097,1441,1467]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rosmarus">A. rosmarus</taxonomicName>
. Spermatheca
</emphasis>
(
<figureCitation id="00845CB72748FFCBFB3AFE834C0F06E5" box="[1271,1367,1440,1467]" captionStart="FIGURES 149154" captionStartId="50.[151,264,1168,1192]" captionTargetBox="[151,1436,181,1145]" captionTargetId="figure-29@50.[151,1436,181,1145]" captionTargetPageId="50" captionText="FIGURES 149154. Axiagastus mitescens Distant, 1901, male and female genitalia. 149151—phallus (149—dorsal view, 150—ventral view, 151—lateral view); 152—terminalia (dorsal); 153—terminalia (ventral); 154—spermatheca. Lettering: ad—aedeagus, cp—conjunctival processes, pa—processes of aedeagus, vcp—ventral convex projection. Scale: 0.5 mm (149 154). Orig. S. Salini." pageId="50" pageNumber="51">Fig. 154</figureCitation>
) as in
<taxonomicName id="5FBF3BB12748FFCBFF5AFEE648470680" authorityName="Dallas" authorityYear="1851" box="[151,287,1477,1503]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C202748FFCBFF5AFEE648470680" box="[151,287,1477,1503]" italics="true" pageId="50" pageNumber="51">A. rosmarus</emphasis>
</taxonomicName>
, distal spermathecal duct gradually widened towards proximal flange, slightly wider than intermediate part of spermatheca; apical receptacle small, with three ductules; a pair of short, subequal (one with short branch before apex), third one slightly longer.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92748FFCBFF0AFD134A720585" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="980040322748FFCBFF0AFD134DD605E8" blockId="50.[151,1437,1368,2043]" pageId="50" pageNumber="51">
<emphasis id="AACB9C202748FFCBFF0AFD1348890514" bold="true" box="[199,465,1584,1610]" pageId="50" pageNumber="51">Differential diagnosis.</emphasis>
This robust species is the only known species of
<taxonomicName id="5FBF3BB12748FFCBFBD4FD124DCC0515" authorityName="Dallas" authorityYear="1851" box="[1049,1172,1585,1611]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="AACB9C202748FFCBFBD4FD124DCC0515" box="[1049,1172,1585,1611]" italics="true" pageId="50" pageNumber="51">Axiagastus</emphasis>
</taxonomicName>
with a unique orange-yellow dorsal body colour with well-expressed black markings, especially on the scutellar disc, and the lack of a yellowish, impunctate spot on the apex of the scutellum. This species can also be distinguished by differences in the male genitalia structures, such as the shape of the paramere and processes of the aedeagus.
</paragraph>
<paragraph id="980040322748FFCBFF0AFDE24A720585" blockId="50.[151,1437,1368,2043]" box="[199,810,1728,1755]" pageId="50" pageNumber="51">
<emphasis id="AACB9C202748FFCBFF0AFDE248C00584" bold="true" box="[199,408,1728,1755]" pageId="50" pageNumber="51">Plant association.</emphasis>
<taxonomicName id="5FBF3BB12748FFCBFE52FDE24B5F0585" box="[415,519,1729,1755]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="50" pageNumber="51" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
(
<bibRefCitation id="FC2E3DC32748FFCBFDD8FDE34BF90585" author="Zhang, S. - M." box="[533,673,1728,1755]" pageId="50" pageNumber="51" pagination="1 - 169" refId="ref57418" refString="Zhang, S. - M. (Ed.) (1995) Hemiptera (2). In: Economic insect fauna of China. Vol. 50. Science Press, Beijing, pp. i - xiii + 1-169, XXIV pls. [in Chinese]" type="book chapter" year="1995">Zhang 1995</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32748FFCBFD61FDE24A460585" author="Hua, L. - Z." box="[684,798,1729,1755]" pageId="50" pageNumber="51" pagination="162 - 216" refId="ref51850" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Huayu Nature Book Trade Co. Ltd., Guangdong, pp. 162-216. [Complete pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92748FFCBFF0AFDC648F70418" pageId="50" pageNumber="51" type="etymology">
<paragraph id="980040322748FFCBFF0AFDC648F70418" blockId="50.[151,1437,1368,2043]" pageId="50" pageNumber="51">
<emphasis id="AACB9C202748FFCBFF0AFDC6481605A1" bold="true" box="[199,334,1765,1791]" pageId="50" pageNumber="51">Etymology.</emphasis>
Although the original publication did not specify the gender or the etymology, the specific name
<taxonomicName id="5FBF3BB12748FFCBFF5AFC2A4849047C" authorityName="Distant" authorityYear="1901" box="[151,273,1801,1827]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mitescens">
<emphasis id="AACB9C202748FFCBFF69FC2A4849047C" box="[164,273,1801,1826]" italics="true" pageId="50" pageNumber="51">mitescens</emphasis>
</taxonomicName>
” is the present participle in the nominative singular of the verb “
<emphasis id="AACB9C202748FFCBFC26FC2A4D19047C" box="[1003,1089,1801,1826]" italics="true" pageId="50" pageNumber="51">mitesco</emphasis>
”, meaning “to become mild or mellow”. It is invariable.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92748FFCBFF0AFC734DEB04A4" pageId="50" pageNumber="51" type="distribution">
<paragraph id="980040322748FFCBFF0AFC734C4004D1" blockId="50.[151,1437,1368,2043]" pageId="50" pageNumber="51">
<emphasis id="AACB9C202748FFCBFF0AFC7348390434" bold="true" box="[199,353,1872,1898]" pageId="50" pageNumber="51">Distribution.</emphasis>
<collectingCountry id="E0A800A22748FFCBFEAAFC7348F60434" box="[359,430,1872,1898]" name="China" pageId="50" pageNumber="51">China</collectingCountry>
:
<collectingRegion id="5A7B8ED02748FFCBFE74FC734B590434" box="[441,513,1872,1898]" country="China" name="Fujian" pageId="50" pageNumber="51">Fujian</collectingRegion>
(
<bibRefCitation id="FC2E3DC32748FFCBFDC2FC734B950435" author="Chen, S. - L. &amp; Li, Y. - G. &amp; Lin, S. - M. &amp; Lin, Q. - Y." box="[527,717,1872,1899]" pageId="50" pageNumber="51" pagination="45 - 51" refId="ref49031" refString="Chen, S. - L., Li, Y. - G., Lin, S. - M. &amp; Lin, Q. - Y. (1985) A list of Hemiptera Pentatomidae family of Fujian Province. Journal of Fujian College of Forestry, 5 (2), 45-51. [in Chinese, English summary]" type="journal article" year="1985">
Chen
<emphasis id="AACB9C202748FFCBFD9EFC724BD30434" box="[595,651,1872,1898]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
1985
</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32748FFCBFD15FC724A100435" author="Hua, L. - Z." box="[728,840,1873,1899]" pageId="50" pageNumber="51" pagination="162 - 216" refId="ref51850" refString="Hua, L. - Z. (2000) XVIII. Order Hemiptera. In: Hua, L. - Z., List of Chinese Insects. Vol. I. Huayu Nature Book Trade Co. Ltd., Guangdong, pp. 162-216. [Complete pagination: 448 pp.]" type="book chapter" year="2000">Hua 2000</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32748FFCBFC9EFC734D4A0435" author="Rider, D. A. &amp; Zheng, L. - Y. &amp; Kerzhner, I. M." box="[851,1042,1872,1899]" pageId="50" pageNumber="51" pagination="135 - 153" refId="ref54497" refString="Rider, D. A., Zheng, L. - Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135-153. https://doi.org/10.31610/zsr/2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="AACB9C202748FFCBFC55FC724A880434" box="[920,976,1872,1898]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
2002
</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32748FFCBFBD0FC724DD10435" author="Fan, Z. - H." box="[1053,1161,1873,1899]" pageId="50" pageNumber="51" refId="ref50076" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Ph. D. thesis, Nankai University, Tianjin, 614 pp. [in Chinese, English summary]" type="book" year="2011">Fan 2011</bibRefCitation>
),
<collectingRegion id="5A7B8ED02748FFCBFB51FC734C470434" box="[1180,1311,1872,1898]" country="China" name="Guangdong" pageId="50" pageNumber="51">Guangdong</collectingRegion>
(
<bibRefCitation id="FC2E3DC32748FFCBFAE0FC724CC00435" author="Wu, C. F." box="[1325,1432,1873,1899]" pageId="50" pageNumber="51" pagination="203 - 231" refId="ref57141" refString="Wu, C. F. (1933) A preliminary check list of Hemiptera Heteroptera recorded from Kwangtung Province, South China. Lingnan Science Journal Supplement, 12, 203-231." type="journal article" year="1933">Wu 1933</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32748FFCBFF5AFC57480E04D1" author="Rider, D. A. &amp; Zheng, L. - Y. &amp; Kerzhner, I. M." box="[151,342,1908,1935]" pageId="50" pageNumber="51" pagination="135 - 153" refId="ref54497" refString="Rider, D. A., Zheng, L. - Y. &amp; Kerzhner, I. M. (2002) Checklist and nomenclatural notes on the Chinese Pentatomidae (Heteroptera). II. Pentatominae. Zoosystematica Rossica, 11 (1), 135-153. https://doi.org/10.31610/zsr/2002.11.1.135" type="journal article" year="2002">
Rider
<emphasis id="AACB9C202748FFCBFF11FC56484D04D0" box="[220,277,1908,1934]" italics="true" pageId="50" pageNumber="51">et al.</emphasis>
2002
</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32748FFCBFEACFC56489404D1" author="Fan, Z. - H." box="[353,460,1909,1935]" pageId="50" pageNumber="51" refId="ref50076" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Ph. D. thesis, Nankai University, Tianjin, 614 pp. [in Chinese, English summary]" type="book" year="2011">Fan 2011</bibRefCitation>
, no exact records),
<collectingRegion id="5A7B8ED02748FFCBFD6BFC574BAF04D0" box="[678,759,1908,1934]" country="China" name="Hainan" pageId="50" pageNumber="51">Hainan</collectingRegion>
(
<bibRefCitation id="FC2E3DC32748FFCBFCC8FC574AC104D1" author="Distant, W. L." box="[773,921,1908,1935]" pageId="50" pageNumber="51" pagination="581 - 592" refId="ref49383" refString="Distant, W. L. (1901) Contributions to a knowledge of the Rhynchota. Transactions of the Entomological Society of London, 1901 (4), 581-592, pl. 16. https://doi.org/10.1111/j.1365-2311.1901.tb01374.x" type="journal article" year="1901">Distant 1901</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32748FFCBFC68FC574DCC04D1" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[933,1172,1908,1935]" pageId="50" pageNumber="51" pagination="297 - 300" refId="ref51680" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297-300, pls. 5 - 25. [book pagination: iii + 330 pp + 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng 1977</bibRefCitation>
,
<bibRefCitation id="FC2E3DC32748FFCBFB52FC564C5404D1" author="Fan, Z. - H." box="[1183,1292,1909,1935]" pageId="50" pageNumber="51" refId="ref50076" refString="Fan, Z. - H. (2011) Study on the taxonomy of Pentatominae from China (Hemiptera: Heteroptera: Pentatomidae). Ph. D. thesis, Nankai University, Tianjin, 614 pp. [in Chinese, English summary]" type="book" year="2011">Fan 2011</bibRefCitation>
).
</paragraph>
<paragraph id="980040322748FFCBFF0AFCBB4DEB04A4" blockId="50.[151,1437,1368,2043]" pageId="50" pageNumber="51">
The record from Guangdong (
<bibRefCitation id="FC2E3DC32748FFCBFDDAFCBA4BD804ED" author="Wu, C. F." box="[535,640,1945,1971]" pageId="50" pageNumber="51" pagination="203 - 231" refId="ref57141" refString="Wu, C. F. (1933) A preliminary check list of Hemiptera Heteroptera recorded from Kwangtung Province, South China. Lingnan Science Journal Supplement, 12, 203-231." type="journal article" year="1933">Wu 1933</bibRefCitation>
) may refer to Hainan, as that island was previously part of Guangdong (cf.
<bibRefCitation id="FC2E3DC32748FFCBFF09FC9F48EC0489" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." box="[196,436,1980,2007]" pageId="50" pageNumber="51" pagination="297 - 300" refId="ref51680" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297-300, pls. 5 - 25. [book pagination: iii + 330 pp + 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng 1977</bibRefCitation>
). It has also been reported to be distributed in
<collectingCountry id="E0A800A22748FFCBFC74FC9E4D700489" box="[953,1064,1981,2007]" name="Myanmar" pageId="50" pageNumber="51">Myanmar</collectingCountry>
,
<collectingCountry id="E0A800A22748FFCBFBFEFC9F4DE90488" box="[1075,1201,1980,2006]" name="Philippines" pageId="50" pageNumber="51">Philippines</collectingCountry>
and
<collectingCountry id="E0A800A22748FFCBFB2AFC9F4C130488" box="[1255,1355,1980,2006]" name="Thailand" pageId="50" pageNumber="51">Thailand</collectingCountry>
(
<bibRefCitation id="FC2E3DC32748FFCBFA94FC9F486704A5" author="Hsiao, T. - Y. &amp; Zheng, L. - Y." pageId="50" pageNumber="51" pagination="297 - 300" refId="ref51680" refString="Hsiao, T. - Y. &amp; Zheng, L. - Y. (1977) Family Pentatomidae. In: Hsiao, T. - Y., Ren, S. - Z., Zheng, L. - Y., Jing, H. - L. &amp; Liu, S. - L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. I. Science Press, Beijing, pp. 53 - 158 + 297-300, pls. 5 - 25. [book pagination: iii + 330 pp + 52 pls; in Chinese with English summary]" type="book chapter" year="1977">Hsiao &amp; Zheng 1977</bibRefCitation>
), however, we did not find any specimens or published records to confirm this.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92749FFCAFF0AFBB54CD103A6" pageId="51" pageNumber="52" type="discussion">
<paragraph id="980040322749FFCAFF0AFBB54CD103A6" blockId="51.[151,1437,150,249]" pageId="51" pageNumber="52">
<emphasis id="AACB9C202749FFCAFF0AFBB5486303EE" bold="true" box="[199,315,150,176]" pageId="51" pageNumber="52">Remarks.</emphasis>
This species was originally described from an unspecified number of specimens (syntypic) with data: “
<collectingRegion id="5A7B8ED02749FFCAFF13FB994869038A" box="[222,305,186,212]" country="China" name="Hainan" pageId="51" pageNumber="52">Hainan</collectingRegion>
Island”. The range of measurements, from “
<quantity id="5F47EDD72749FFCAFCD0FB994AF6038A" box="[797,942,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="51" pageNumber="52" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13 to 14 mm</quantity>
”, indicates that more than
<specimenCount id="8EB98BBB2749FFCAFB1FFB984C33038A" box="[1234,1387,186,213]" count="1" pageId="51" pageNumber="52" type="generic">one specimen</specimenCount>
was examined, and although the fifth joint was noted as “mutilated”, this condition is consistent across all specimens.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,233 @@
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<mods:title id="3C5AEE796225A2BBB7D3C9398975D27A">Revision of the Austro-Oriental shield-bug genus Axiagastus (Hemiptera: Heteroptera: Pentatomidae), with the description of five new species, and taxonomic notes on related genera</mods:title>
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<mods:namePart id="D30169D7E823E7B62AC529CF8B9E13E5">Kment, Petr</mods:namePart>
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<mods:affiliation id="450E9FBB5527BB93B4F5D9CCEB3A8F87">Department of Entomology, National Museum of the Czech Republic, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
<mods:nameIdentifier id="14AE4BC0ED94B1445F0E8ACA884682B1" type="email">sigara@post.cz</mods:nameIdentifier>
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<mods:namePart id="20E16D5688C6FE6D3DC49AF3634F1F7C">Lemaître, Valérie A.</mods:namePart>
<mods:nameIdentifier id="47633D50482F9539973162CEA453ECEC" type="ORCID">0000-0003-4802-2711</mods:nameIdentifier>
<mods:affiliation id="ED6096DBA95E51BB179FEAF8F49B95B1">Department of Collections (Insects), The Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom; &amp; val. heureuse @ gmail. com; https: // orcid. org / 0000 - 0003 - 4802 - 2711</mods:affiliation>
<mods:nameIdentifier id="F4E8D401AE34E8136F230C3998A8EEB9" type="email">val.heureuse@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0C1596643059CD36EF9C7503C0CA3BE4">Webb, Michael D.</mods:namePart>
<mods:nameIdentifier id="AC2920DA92A416C5979DD548FEF60D8D" type="ORCID">0000-0002-1312-6142</mods:nameIdentifier>
<mods:affiliation id="46E07B0A11AC7D6EE1AC5D0CB85FD7CF">Department of Collections (Insects), The Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom; &amp; mdwebb 04 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1312 - 6142</mods:affiliation>
<mods:nameIdentifier id="A0B9C85341E89152BC3CCBC88434904B" type="email">mdwebb04@gmail.com</mods:nameIdentifier>
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<mods:namePart id="58A249CFF3EFAFD2065654D80289C1E7">Roca-Cusachs, Marcos</mods:namePart>
<mods:nameIdentifier id="BBF712C95586C1EFC47BF5D55AE02D72" type="ORCID">0000-0003-3557-4365</mods:nameIdentifier>
<mods:affiliation id="46AF9DB2CD0A4D3FDC664CC09F566ACF">ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, Karnataka, India shalinis. nilavu @ gmail. com; https: // orcid. org / 0000 - 0003 - 3557 - 4365 &amp; Evolution &amp; Ecology Research Centre, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, Australia &amp; IRBio, Institut de Recerca de la Biodiversitat, Universitat de Barcelona, Barcelona, Spain marcosrocacusachs @ gmail. com; https: // orcid. org / 0000 - 0002 - 9174 - 6021 # Former affiliation * Corresponding author &amp; ICAR-National Bureau of Agricultural Insect Resources, Bangalore, 560024, Karnataka, India shalinis. nilavu @ gmail. com; https: // orcid. org / 0000 - 0003 - 3557 - 4365</mods:affiliation>
<mods:nameIdentifier id="C9423447ACA240F1948FC7D751AF3BB2" type="email">shalinis.nilavu@gmail.com</mods:nameIdentifier>
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<mods:title id="1C6785CD5EFA2ACA3E1AAC1A2423BAF5">Zootaxa</mods:title>
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<mods:date id="D08000064B5CBE4F3B665E8994E2A9F3">2025</mods:date>
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<mods:number id="F81E23F02820E59C9D0FEFF65ED676FA">5603</mods:number>
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<paragraph id="98004032275DFFDEFF5AFE5D4BDF06C6" blockId="39.[151,647,1405,1468]" box="[151,647,1405,1432]" pageId="39" pageNumber="40">
<heading id="C348F75E275DFFDEFF5AFE5D4BDF06C6" bold="true" box="[151,647,1405,1432]" fontSize="11" level="2" pageId="39" pageNumber="40" reason="4">
<taxonomicName id="5FBF3BB1275DFFDEFF5AFE5D4BDF06C6" authority="Jensen-Haarup, 1937" authorityName="Jensen-Haarup" authorityYear="1937" box="[151,647,1405,1432]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="AACB9C20275DFFDEFF5AFE5D4BDF06C6" bold="true" box="[151,647,1405,1432]" pageId="39" pageNumber="40">
<emphasis id="AACB9C20275DFFDEFF5AFE5D482E06C6" bold="true" box="[151,374,1405,1432]" italics="true" pageId="39" pageNumber="40">Axiagastus dubius</emphasis>
<bibRefCitation id="FC2E3DC3275DFFDEFEB0FE5D4BDF06C6" author="Jensen-Haarup, A. C." box="[381,647,1406,1432]" pageId="39" pageNumber="40" pagination="321 - 324" refId="ref51987" refString="Jensen-Haarup, A. C. (1937) Einige neue Pentatomidenarten aus der Sammlung des Zoologischen Museums in Hamburg (Hem. het.). Entomologiske Rundschau, 54 (26), 321-324." type="journal article" year="1937">Jensen-Haarup, 1937</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="D0A513B9275DFFDEFF5AFE81481906E2" box="[151,321,1442,1468]" pageId="39" pageNumber="40" type="description">
<paragraph id="98004032275DFFDEFF5AFE81481906E2" blockId="39.[151,647,1405,1468]" box="[151,321,1442,1468]" pageId="39" pageNumber="40">
(
<figureCitation id="00845CB7275DFFDEFF52FE81486106E2" box="[159,313,1442,1468]" captionStart-0="FIGURES 110111" captionStart-1="FIGURES 112113" captionStartId-0="40.[151,264,1943,1967]" captionStartId-1="41.[151,264,1131,1155]" captionTargetBox-0="[183,1404,181,1919]" captionTargetBox-1="[151,1436,199,1107]" captionTargetId-0="figure-29@40.[183,1404,181,1919]" captionTargetId-1="figure-19@41.[151,1436,199,1107]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 110111. Axiagastus dubius Jensen-Haarup, 1937—habitus. 110—dorsal view; 111—ventral view. Orig. Eileen Nguyen, © ZMUH. Scale: 1 mm (110111)." captionText-1="FIGURES 112113. Axiagastus dubius Jensen-Haarup, 1937. 112—abdomen (ventral view); 113—type labels. Orig. Eileen Nguyen, © ZMUH. Scale: 1 mm (112)." pageId="39" pageNumber="40">Figs 110113</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D0A513B9275DFFDEFF5AFEC44BAF0562" pageId="39" pageNumber="40" type="reference_group">
<paragraph id="98004032275DFFDEFF5AFEC448720543" blockId="39.[151,1436,1510,1596]" pageId="39" pageNumber="40">
<treatmentCitationGroup id="B8AF671C275DFFDEFF5AFEC448720543" pageId="39" pageNumber="40">
<treatmentCitation id="191E6623275DFFDEFF5AFEC44B0006A0" author="Jensen-Haarup, A. C." box="[151,600,1510,1534]" page="322" pageId="39" pageNumber="40" year="1937">
<taxonomicName id="5FBF3BB1275DFFDEFF5AFEC44B0006A0" authority="Jensen-Haarup, 1937: 322" authorityName="Jensen-Haarup" authorityPageNumber="322" authorityYear="1937" box="[151,600,1510,1534]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="AACB9C20275DFFDEFF5AFEC4481706A0" box="[151,335,1510,1534]" italics="true" pageId="39" pageNumber="40">Axiagastus dubius</emphasis>
<materialsCitation id="28D74A6F275DFFDEFE9EFEC44B0006A0" box="[339,600,1510,1534]" collectionCode="ZMUH" country="Singapore" location="Alte Sammlung" pageId="39" pageNumber="40" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<bibRefCitation id="FC2E3DC3275DFFDEFE9EFEC44B0006A0" author="Jensen-Haarup, A. C." box="[339,600,1510,1534]" pageId="39" pageNumber="40" pagination="321 - 324" refId="ref51987" refString="Jensen-Haarup, A. C. (1937) Einige neue Pentatomidenarten aus der Sammlung des Zoologischen Museums in Hamburg (Hem. het.). Entomologiske Rundschau, 54 (26), 321-324." type="journal article" year="1937">Jensen-Haarup, 1937: 322</bibRefCitation>
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(original description).
<typeStatus id="4704FE90275DFFDEFCF6FEC54AC506A0" box="[827,925,1510,1534]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
:
<specimenCount id="8EB98BBB275DFFDEFC68FEC44AE206A0" box="[933,954,1511,1534]" count="1" pageId="39" pageNumber="40" type="male"></specimenCount>
, “
<collectingCountry id="E0A800A2275DFFDEFC01FEC54D6F06A0" box="[972,1079,1510,1534]" name="Singapore" pageId="39" pageNumber="40">Singapore</collectingCountry>
?
<location id="9D6016E9275DFFDEFB8EFEC54DB906A0" LSID="urn:lsid:plazi:treatment:1016F124275DFFD0FF5AFE5D4C7906B5:9D6016E9275DFFDEFB8EFEC54DB906A0" box="[1091,1249,1510,1534]" country="Singapore" name="Alte Sammlung" pageId="39" pageNumber="40">Alte Sammlung</location>
[= old collection]” (ZMUH).
</treatmentCitationGroup>
</paragraph>
<paragraph id="98004032275DFFDEFF5AFD074BAF0562" blockId="39.[151,1436,1510,1596]" box="[151,759,1572,1596]" pageId="39" pageNumber="40">
<treatmentCitationGroup id="B8AF671C275DFFDEFF5AFD074BAF0562" box="[151,759,1572,1596]" pageId="39" pageNumber="40">
<taxonomicName id="5FBF3BB1275DFFDEFF5AFD0748090562" authorityName="Jensen-Haarup" authorityYear="1937" box="[151,337,1572,1596]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="AACB9C20275DFFDEFF5AFD0748090562" box="[151,337,1572,1596]" italics="true" pageId="39" pageNumber="40">Axiagastus dubius</emphasis>
</taxonomicName>
:
<treatmentCitation id="191E6623275DFFDEFE90FD074B770562" author="Weidner, H." box="[349,559,1572,1596]" page="114" pageId="39" pageNumber="40" year="1972">
<bibRefCitation id="FC2E3DC3275DFFDEFE90FD074B580562" author="Weidner, H." box="[349,512,1572,1596]" pageId="39" pageNumber="40" pagination="107 - 134" refId="ref57015" refString="Weidner, H. (1972) Die Entomologischen Sammlungen des Zoologischen Instituts und Zoologischen Museums der Universitat Hamburg. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 68, 107-134." type="journal article" year="1972">Weidner (1972)</bibRefCitation>
: 114
</treatmentCitation>
(
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information).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D0A513B9275DFFD0FF5AFD494B7D0669" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="98004032275DFFDEFF5AFD49489A05DA" blockId="39.[151,1437,1642,2028]" box="[151,450,1642,1668]" pageId="39" pageNumber="40">
<materialsCitation id="28D74A6F275DFFDEFF5AFD4948E605DA" box="[151,446,1642,1668]" country="Singapore" location="Singapore" pageId="39" pageNumber="40" specimenCount="1" typeStatus="holotype">
<emphasis id="AACB9C20275DFFDEFF5AFD49486E05DA" bold="true" box="[151,310,1642,1668]" pageId="39" pageNumber="40">
<typeStatus id="4704FE90275DFFDEFF5AFD49498A05DA" box="[151,210,1642,1668]" pageId="39" pageNumber="40">Type</typeStatus>
locality.
</emphasis>
?
<collectingCountry id="E0A800A2275DFFDEFE8AFD4948E605DA" box="[327,446,1642,1668]" name="Singapore" pageId="39" pageNumber="40">Singapore</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="98004032275DFFDEFF0AFDAD4C620402" blockId="39.[151,1437,1642,2028]" pageId="39" pageNumber="40">
<emphasis id="AACB9C20275DFFDEFF0AFDAD482B05F6" bold="true" box="[199,371,1678,1704]" pageId="39" pageNumber="40">Type material.</emphasis>
<materialsCitation id="28D74A6F275DFFDEFEB5FDAD4C620402" collectionCode="ZMUH" country="Singapore" location="Martin Husemann" municipality="University of Hamburg" pageId="39" pageNumber="40" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="4704FE90275DFFDEFEB5FDAD48BD05F6" box="[376,485,1678,1704]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
:
<specimenCount id="8EB98BBB275DFFDEFE20FDAD4B5A05F6" box="[493,514,1678,1704]" count="1" pageId="39" pageNumber="40" type="male"></specimenCount>
(
<figureCitation id="00845CB7275DFFDEFDC2FDAD4BF105F6" box="[527,681,1678,1704]" captionStart-0="FIGURES 110111" captionStart-1="FIGURES 112113" captionStartId-0="40.[151,264,1943,1967]" captionStartId-1="41.[151,264,1131,1155]" captionTargetBox-0="[183,1404,181,1919]" captionTargetBox-1="[151,1436,199,1107]" captionTargetId-0="figure-29@40.[183,1404,181,1919]" captionTargetId-1="figure-19@41.[151,1436,199,1107]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionText-0="FIGURES 110111. Axiagastus dubius Jensen-Haarup, 1937—habitus. 110—dorsal view; 111—ventral view. Orig. Eileen Nguyen, © ZMUH. Scale: 1 mm (110111)." captionText-1="FIGURES 112113. Axiagastus dubius Jensen-Haarup, 1937. 112—abdomen (ventral view); 113—type labels. Orig. Eileen Nguyen, © ZMUH. Scale: 1 mm (112)." pageId="39" pageNumber="40">Figs 110113</figureCitation>
), “
<collectingCountry id="E0A800A2275DFFDEFD08FDAD4A6405F6" box="[709,828,1678,1704]" name="Singapore" pageId="39" pageNumber="40">Singapore</collectingCountry>
. [p]? [hw] /Alte Sammlung. [p] //
<taxonomicName id="5FBF3BB1275DFFDEFB7BFDAD4C6A05F6" authorityName="Dallas" authorityYear="1851" box="[1206,1330,1678,1704]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Axiagastus</taxonomicName>
/ n. spec? / nicht Rosmarus! [Breddins hw] // A. C. Jensen-/ Haarup / determ. 1931-32 [p] // Type [p, red label] // Type Coll. J-Hrp [p, label with black submarginal line] //
<taxonomicName id="5FBF3BB1275DFFDEFD50FDF54A4105AE" authorityName="Dallas" authorityYear="1851" box="[669,793,1750,1776]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Axiagastus</taxonomicName>
/ dubius J-Hrps / Det. [hw] Jensen-Haarup. [p, label with black submarginal line] // ZMH 81525 [p]” (
<collectionCode id="FEAED8F7275DFFDEFD40FDD94BBE044A" box="[653,742,1786,1812]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="39" pageNumber="40">ZMUH</collectionCode>
). The specimen is pinned through the scutellum, considerably damaged, both hemelytra, left and right antennomeres IIbIV, and right metatarsomeres IIIII lost. We examined images provided by courtesy of Eileen Nguyen (
<collectionCode id="FEAED8F7275DFFDEFD78FC614A530402" box="[693,779,1858,1884]" country="Viet Nam" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoological Museum, University of Hanoi" pageId="39" pageNumber="40">ZMUH</collectionCode>
) and
<location id="9D6016E9275DFFDEFC84FC614D4C0402" LSID="urn:lsid:plazi:treatment:1016F124275DFFD0FF5AFE5D4C7906B5:9D6016E9275DFFDEFC84FC614D4C0402" box="[841,1044,1858,1884]" country="Singapore" municipality="University of Hamburg" name="Martin Husemann" pageId="39" pageNumber="40">Martin Husemann</location>
(
<collectingMunicipality id="7864DA48275DFFDEFBEFFC614C770402" box="[1058,1327,1858,1884]" pageId="39" pageNumber="40">University of Hamburg</collectingMunicipality>
).
</materialsCitation>
</paragraph>
<paragraph id="98004032275DFFD0FF0AFC454B7D0669" blockId="39.[151,1437,1642,2028]" lastBlockId="41.[151,1436,1273,1516]" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40">
<emphasis id="AACB9C20275DFFDEFF0AFC4548E204DE" bold="true" box="[199,442,1894,1920]" pageId="39" pageNumber="40">Original description</emphasis>
(
<bibRefCitation id="FC2E3DC3275DFFDEFE04FC454BED04DE" author="Jensen-Haarup, A. C." box="[457,693,1894,1920]" pageId="39" pageNumber="40" pagination="321 - 324" refId="ref51987" refString="Jensen-Haarup, A. C. (1937) Einige neue Pentatomidenarten aus der Sammlung des Zoologischen Museums in Hamburg (Hem. het.). Entomologiske Rundschau, 54 (26), 321-324." type="journal article" year="1937">Jensen-Haarup 1937</bibRefCitation>
, translated from the German). “Very similar to
<taxonomicName id="5FBF3BB1275DFFDEFB00FC454CC004DE" authority="Dall." authorityName="Dallas" authorityYear="1851" box="[1229,1432,1894,1920]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C20275DFFDEFB00FC454C0E04DE" box="[1229,1366,1894,1920]" italics="true" pageId="39" pageNumber="40">A. rosmarus</emphasis>
Dall.
</taxonomicName>
, but, as Breddin noted on a label placed underneath the animal, not this species. Colouring brownish and yellow. The scutellum is elongate, much longer than wide at the base. It is missing the two large black spots on the anterior part of the scutellum as well as the curved dark spot in front of the tip, which are found in
<taxonomicName id="5FBF3BB1275DFFDEFB80FCF14D8D04B2" authorityName="Dallas" authorityYear="1851" box="[1101,1237,2002,2028]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="rosmarus">
<emphasis id="AACB9C20275DFFDEFB80FCF14D8D04B2" box="[1101,1237,2002,2028]" italics="true" pageId="39" pageNumber="40">A. rosmarus</emphasis>
</taxonomicName>
; the tip is bearing an almost circular, pale spot. The hemelytra bearing, past its middle, a pale spot next to the main vein. Length of the body (without membrane)
<quantity id="5F47EDD72753FFD0FE0EFE3E4B4F0669" box="[451,535,1309,1335]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="41" pageNumber="42" unit="mm" value="14.0">14 mm</quantity>
.”
</paragraph>
</subSubSection>
<caption id="CCC010BA2752FFD1FF5AFCB44B00048A" pageId="40" pageNumber="41" startId="40.[151,264,1943,1967]" targetBox="[183,1404,181,1919]" targetPageId="40" targetType="figure">
<paragraph id="980040322752FFD1FF5AFCB44B00048A" blockId="40.[151,1437,1943,2006]" pageId="40" pageNumber="41">
<emphasis id="AACB9C202752FFD1FF5AFCB4483404F1" bold="true" box="[151,364,1943,1967]" pageId="40" pageNumber="41">FIGURES 110111.</emphasis>
<taxonomicName id="5FBF3BB12752FFD1FEBBFCBB4A4D04EE" authority="Jensen-Haarup, 1937" authorityName="Jensen-Haarup" authorityYear="1937" box="[374,789,1944,1968]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="AACB9C202752FFD1FEBBFCBB4B6B04EE" box="[374,563,1944,1968]" italics="true" pageId="40" pageNumber="41">Axiagastus dubius</emphasis>
<bibRefCitation id="FC2E3DC32752FFD1FDF1FCBB4A4D04EE" author="Jensen-Haarup, A. C." box="[572,789,1944,1968]" pageId="40" pageNumber="41" pagination="321 - 324" refId="ref51987" refString="Jensen-Haarup, A. C. (1937) Einige neue Pentatomidenarten aus der Sammlung des Zoologischen Museums in Hamburg (Hem. het.). Entomologiske Rundschau, 54 (26), 321-324." type="journal article" year="1937">Jensen-Haarup, 1937</bibRefCitation>
</taxonomicName>
—habitus. 110—dorsal view; 111—ventral view. Orig. Eileen Nguyen, © ZMUH. Scale: 1 mm (110111).
</paragraph>
</caption>
<caption id="CCC010BA2753FFD0FF5AFF484B7F07F9" pageId="41" pageNumber="42" startId="41.[151,264,1131,1155]" targetBox="[151,1436,199,1107]" targetPageId="41" targetType="figure">
<paragraph id="980040322753FFD0FF5AFF484B7F07F9" blockId="41.[151,1437,1131,1193]" pageId="41" pageNumber="42">
<emphasis id="AACB9C202753FFD0FF5AFF48483407DD" bold="true" box="[151,364,1131,1155]" pageId="41" pageNumber="42">FIGURES 112113.</emphasis>
<taxonomicName id="5FBF3BB12753FFD0FEB8FF484A4B07DD" authority="Jensen-Haarup, 1937" authorityName="Jensen-Haarup" authorityYear="1937" box="[373,787,1131,1155]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="AACB9C202753FFD0FEB8FF484B6807DD" box="[373,560,1131,1155]" italics="true" pageId="41" pageNumber="42">Axiagastus dubius</emphasis>
<bibRefCitation id="FC2E3DC32753FFD0FDF5FF4F4A4B07DD" author="Jensen-Haarup, A. C." box="[568,787,1131,1155]" pageId="41" pageNumber="42" pagination="321 - 324" refId="ref51987" refString="Jensen-Haarup, A. C. (1937) Einige neue Pentatomidenarten aus der Sammlung des Zoologischen Museums in Hamburg (Hem. het.). Entomologiske Rundschau, 54 (26), 321-324." type="journal article" year="1937">Jensen-Haarup, 1937</bibRefCitation>
</taxonomicName>
. 112—abdomen (ventral view); 113—type labels. Orig. Eileen Nguyen, © ZMUH. Scale: 1 mm (112).
</paragraph>
</caption>
<subSubSection id="D0A513B92753FFD0FF0AFE624D8C0605" box="[199,1236,1345,1372]" pageId="41" pageNumber="42" type="etymology">
<paragraph id="980040322753FFD0FF0AFE624D8C0605" blockId="41.[151,1436,1273,1516]" box="[199,1236,1345,1372]" pageId="41" pageNumber="42">
<emphasis id="AACB9C202753FFD0FF0AFE6248160605" bold="true" box="[199,334,1345,1371]" pageId="41" pageNumber="42">Etymology.</emphasis>
The specific epithet is the Latin adjective
<taxonomicName id="5FBF3BB12753FFD0FCE7FE624A2D0605" authorityName="Jensen-Haarup" authorityYear="1937" box="[810,885,1345,1371]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="AACB9C202753FFD0FCE7FE624A2D0605" box="[810,885,1345,1371]" italics="true" pageId="41" pageNumber="42">dubius</emphasis>
</taxonomicName>
(-
<emphasis id="AACB9C202753FFD0FC43FE614AC40605" box="[910,924,1346,1371]" italics="true" pageId="41" pageNumber="42">a</emphasis>
, -
<emphasis id="AACB9C202753FFD0FC7EFE614A8D0605" box="[947,981,1346,1371]" italics="true" pageId="41" pageNumber="42">um</emphasis>
), meaning “doubtful”.
</paragraph>
</subSubSection>
<subSubSection id="D0A513B92753FFD0FF0AFE464C7906B5" pageId="41" pageNumber="42" type="discussion">
<paragraph id="980040322753FFD0FF0AFE464C7906B5" blockId="41.[151,1436,1273,1516]" pageId="41" pageNumber="42">
<emphasis id="AACB9C202753FFD0FF0AFE4648630621" bold="true" box="[199,315,1381,1407]" pageId="41" pageNumber="42">Remarks.</emphasis>
<taxonomicName id="5FBF3BB12753FFD0FE88FE454B4D0621" authorityName="Jensen-Haarup" authorityYear="1937" box="[325,533,1381,1408]" class="Insecta" family="Pentatomidae" genus="Axiagastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="AACB9C202753FFD0FE88FE454B4D0621" box="[325,533,1381,1408]" italics="true" pageId="41" pageNumber="42">Axiagastus dubius</emphasis>
</taxonomicName>
was originally described by
<bibRefCitation id="FC2E3DC32753FFD0FCAAFE454D3A0621" author="Jensen-Haarup, A. C." box="[871,1122,1381,1408]" pageId="41" pageNumber="42" pagination="321 - 324" refId="ref51987" refString="Jensen-Haarup, A. C. (1937) Einige neue Pentatomidenarten aus der Sammlung des Zoologischen Museums in Hamburg (Hem. het.). Entomologiske Rundschau, 54 (26), 321-324." type="journal article" year="1937">Jensen-Haarup (1937)</bibRefCitation>
based on a male specimen collected from
<collectingCountry id="E0A800A22753FFD0FE88FEAA48EF06FD" box="[325,439,1417,1443]" name="Singapore" pageId="41" pageNumber="42">Singapore</collectingCountry>
(as a doubtful locality). The specimen is currently deposited in the Museum of Nature Hamburg—Zoology, Leibniz Institute for the Analysis of Biodiversity Change (ZMUH).This species is characterized by the chocolate brown colouration, with a more or less pale, circular spot on the apex of the scutellum.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,257 @@
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<mods:title id="01B28D2571E21E504C09A19A2621F1E9">Juliaca Melichar, 1926 (Hemiptera: Cicadellidae: Cicadellinae) from Ecuador: description of a new species and identification key</mods:title>
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<taxonomicName id="5B964D3AFF90FFFF0994FAE8FE3F9B2A" authorityName="Felix &amp; Silva &amp; Quintas &amp; Pecly &amp; Froza &amp; Mejdalani" authorityYear="2025" box="[151,401,1302,1329]" class="Insecta" family="Cicadellidae" genus="Juliaca" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="nigrosignata" status="sp. nov.">
<emphasis id="AEE2EAABFF90FFFF0994FAE8FE3F9B2A" bold="true" box="[151,401,1302,1329]" italics="true" pageId="1" pageNumber="78">Juliaca nigrosignata</emphasis>
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(
<figureCitation id="04AD2A3CFF90FFFF099CFAC5FEB69B4E" box="[159,280,1339,1365]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="1" pageNumber="78">Figs. 112</figureCitation>
)
</paragraph>
<paragraph id="9C2936B9FF90FFFF0994FA7DFC0E9B86" blockId="1.[151,1437,1411,2013]" box="[151,928,1411,1437]" pageId="1" pageNumber="78">Length. Males 6.196.38 mm (n = 2); females 6.036.24 mm (n = 2).</paragraph>
<paragraph id="9C2936B9FF90FFFF09C4FA59FE9998FA" blockId="1.[151,1437,1411,2013]" pageId="1" pageNumber="78">
Head (
<figureCitation id="04AD2A3CFF90FFFF0811FA59FEFB9BDA" box="[274,341,1447,1473]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="1" pageNumber="78">Fig. 1</figureCitation>
). Crown, in dorsal view, moderately produced anteriorly, its median length approximately 1/2 of interocular width and 3/10 of transocular width; surface smooth, without transverse concavity before ocelli; anterior margin broadly rounded; without carina at transition from crown to face. Ocelli located slightly behind imaginary transverse line between anterior angles of compound eyes, each ocellus approximately equidistant from median line of crown and adjacent anterior eye angle. Frontogenal suture extending onto crown and attaining ocellus. Antennal ledge, in dorsal view, not protuberant; in lateral view, with anterior margin approximately rectilinear and vertical. Face with frons convex medially; muscle impressions distinct. Epistomal suture obsolete. Clypeus, in lateral view, not protuberant, its contour continuing profile of frons at upper portion and more nearly horizontal at lower portion.
</paragraph>
<paragraph id="9C2936B9FF90FFFF09C4F915FA3F99C6" blockId="1.[151,1437,1411,2013]" pageId="1" pageNumber="78">
Thorax (
<figureCitation id="04AD2A3CFF90FFFF082AF915FEC5991E" box="[297,363,1771,1797]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="1" pageNumber="78">Fig. 1</figureCitation>
). Pronotum, in dorsal view, with width greater than transocular width of head; lateral margins distinctly convergent anteriorly; posterior margin slightly concave; dorsolateral carina distinct, complete, rectilinear, declivous anterad; disk smooth, without transverse rugae.Mesonotum, in dorsal view, with scutellum not transversely striate. Forewing not coriaceous, translucid, without distinct apical membranous area; with four apical cells, base of fourth more proximal than base of third, and three anteapical cells, their bases obscure; without anteapical plexus of veins. Hind wing with vein R
<subScript id="001234FCFF90FFFF08DEF84FFE5999A4" attach="left" box="[477,503,1969,1983]" fontSize="6" pageId="1" pageNumber="78">2+3</subScript>
incomplete. Hind leg with femoral apical setal formula 2:1:1; first tarsomere longer than combined length of two more distal tarsomeres, its plantar surface with two longitudinal rows of small setae.
</paragraph>
<caption id="C8E96631FF93FFFC0994F97BFA2A9936" pageId="2" pageNumber="79" startId="2.[151,264,1669,1693]" targetBox="[162,1428,181,1631]" targetPageId="2" targetType="figure">
<paragraph id="9C2936B9FF93FFFC0994F97BFA2A9936" blockId="2.[151,1437,1669,1837]" pageId="2" pageNumber="79">
<emphasis id="AEE2EAABFF93FFFC0994F97BFEE79886" bold="true" box="[151,329,1669,1693]" pageId="2" pageNumber="79">FIGURES 112.</emphasis>
<taxonomicName id="5B964D3AFF93FFFC0853F97BFD8D9886" authorityName="Felix &amp; Silva &amp; Quintas &amp; Pecly &amp; Froza &amp; Mejdalani" authorityYear="2025" box="[336,547,1669,1693]" class="Insecta" family="Cicadellidae" genus="Juliaca" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="nigrosignata" status="sp. nov.">
<emphasis id="AEE2EAABFF93FFFC0853F97BFD8D9886" box="[336,547,1669,1693]" italics="true" pageId="2" pageNumber="79">Juliaca nigrosignata</emphasis>
</taxonomicName>
<emphasis id="AEE2EAABFF93FFFC0B28F97BFDD29886" bold="true" box="[555,636,1669,1693]" pageId="2" pageNumber="79">
<taxonomicNameLabel id="B5D157D0FF93FFFC0B28F97BFDD29886" box="[555,636,1669,1693]" pageId="2" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm.
</paragraph>
</caption>
<caption id="C8E96631FF92FFFD0994F869FADD99CF" pageId="3" pageNumber="80" startId="3.[151,264,1943,1967]" targetBox="[251,1342,181,1920]" targetPageId="3" targetType="figure">
<paragraph id="9C2936B9FF92FFFD0994F869FADD99CF" blockId="3.[151,1436,1943,2004]" pageId="3" pageNumber="80">
<emphasis id="AEE2EAABFF92FFFD0994F869FEFC99B4" bold="true" box="[151,338,1943,1967]" pageId="3" pageNumber="80">FIGURES 1314.</emphasis>
13, map showing the location of the type locality of
<taxonomicName id="5B964D3AFF92FFFD0A4CF866FBB099B4" authorityName="Felix &amp; Silva &amp; Quintas &amp; Pecly &amp; Froza &amp; Mejdalani" authorityYear="2025" box="[847,1054,1944,1968]" class="Insecta" family="Cicadellidae" genus="Juliaca" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="80" phylum="Arthropoda" rank="species" species="nigrosignata" status="sp. nov.">
<emphasis id="AEE2EAABFF92FFFD0A4CF866FBB099B4" box="[847,1054,1944,1968]" italics="true" pageId="3" pageNumber="80">Juliaca nigrosignata</emphasis>
</taxonomicName>
<emphasis id="AEE2EAABFF92FFFD0D22F866FBDE99AB" bold="true" box="[1057,1136,1944,1968]" pageId="3" pageNumber="80">
<taxonomicNameLabel id="B5D157D0FF92FFFD0D22F866FBDE99AB" box="[1057,1136,1944,1968]" pageId="3" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the Province of Sucumbíos, Ecuador. 14, sub-Andean montane forest at the type locality (about 2,000 m a.s.l.). Photograph by Adriano Brilhante Kury.
</paragraph>
</caption>
<paragraph id="9C2936B9FF95FFFA09C4FF69FB8C9FEE" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="81">
Male terminalia. Pygofer (
<figureCitation id="04AD2A3CFF95FFFA08EFFF69FD819EAA" box="[492,559,151,177]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 2</figureCitation>
), in lateral view, moderately produced posteriorly; without processes; posterior margin broadly rounded; surface with macrosetae distributed mostly on posterior half of disk, some of them very elongate. Valve (
<figureCitation id="04AD2A3CFF95FFFA0851FF21FE3B9EE2" box="[338,405,223,249]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 2</figureCitation>
), in ventral view, subrectangular. Subgenital plate (
<figureCitation id="04AD2A3CFF95FFFA0AD2FF21FB949EE2" box="[977,1082,223,249]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Figs. 2, 3</figureCitation>
), in ventral view, subtriangular, distinctly narrowed along apical half; with elongate uniseriate macrosetae distributed along outer margin; microsetae also present; plates not fused to each other basally; in lateral view, extending as far posteriorly as pygofer apex. Connective (
<figureCitation id="04AD2A3CFF95FFFA0825FEB5FEC59F7E" box="[294,363,331,357]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 4</figureCitation>
), in dorsal view, Y-shaped; arms widely divergent; stalk elongate, with median keel. Style (
<figureCitation id="04AD2A3CFF95FFFA0C6CFEB5FF099F92" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 4</figureCitation>
), in dorsal view, with apophysis elongate; without preapical lobe; apex acute, directed outward. Aedeagus (
<figureCitation id="04AD2A3CFF95FFFA0C4AFE91FF089FB6" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Figs. 5, 6</figureCitation>
) symmetrical; shaft, in lateral view, short, with apical half directed ventrally; apex moderately expanded; without processes; gonopore located ventroapically. Paraphyses (
<figureCitation id="04AD2A3CFF95FFFA0A1FFE49FCCF9FCA" box="[796,865,439,465]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 7</figureCitation>
) symmetrical; stalk poorly sclerotized, articulated with apex of connective; with pair of slender spiniform rami directed posteriorly.
</paragraph>
<paragraph id="9C2936B9FF95FFFA09C4FE01FD569A42" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="81">
Female terminalia. Sternite VII (
<figureCitation id="04AD2A3CFF95FFFA0B2DFE01FDC19C02" box="[558,623,511,537]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 8</figureCitation>
), in ventral view, moderately produced and slightly narrowed posteriorly; posterior margin convex, with distinct median triangular projection. “Internal” sternite VIII, in dorsal view, without distinct sclerites. Pygofer (
<figureCitation id="04AD2A3CFF95FFFA08CDFDB9FDB89C7A" box="[462,534,583,609]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 9</figureCitation>
), in lateral view, moderately produced posteriorly; posterior margin narrowly rounded; macrosetae distributed mostly on posterior third and extending anteriorly along ventral margin. Valvifer I, in lateral view, guttiform, ventral margin distinctly rounded. Valvula I, in ventral view, with basal portion not modified, just slightly expanded; blade, in lateral view, with apical portion gradually narrowed; apex acute; ventral interlocking device located on basiventral half of valvula; dorsal sculptured area extending from basal portion to apex of blade, formed mostly by scale-like processes arranged in oblique lines; ventral sculptured area restricted to apical portion, formed by scale-like processes. Valvifer II, in lateral view, pentagonal, narrowed inferiorly; with distinct sensory field of small setae adjacent to articulation point. Valvula II (
<figureCitation id="04AD2A3CFF95FFFA0AFFFCBDFB2B9D46" box="[1020,1157,835,861]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Figs. 1012</figureCitation>
), in lateral view, slightly expanded beyond basal curvature; dorsal margin convex; preapical prominence distinct; apical portion spear-shaped; apex subacute; blade with about twelve continuous, subtriangular sclerotized teeth, most teeth at basal and median thirds elongate, those at apical third smaller; denticles distributed on teeth and on dorsal and ventral apical portions of blade, except on apex (dorsal and ventral dentate apical portions with about same size); blade with ducts extending toward teeth and apex. Gonoplac extending as far posteriorly as pygofer apex; in lateral view, with basal half narrow and apical half distinctly expanded; apex obtuse; surface with denticuli and few small setae on apical portion and extending anteriorly along ventral margin.
</paragraph>
<paragraph id="9C2936B9FF95FFFA09C4FB9DFD089AF2" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="81">
Color (
<figureCitation id="04AD2A3CFF95FFFA081BFB9DFEF29A66" box="[280,348,1123,1149]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 1</figureCitation>
). Body mostly white to brownish-white. Crown with single, conspicuous black spot at median portion, pedunculate at base, broadened at middle where it attains ocelli, and emarginate at apex; compound eye, in lateral view, mostly dark brown to black on superior portion and mostly yellow on inferior portion. Tarsal claws brown. Subgenital plate with apex dark brown.
</paragraph>
</subSubSection>
<subSubSection id="D48C6532FF95FFFA09C4FB0DFAB79B4E" pageId="4" pageNumber="81" type="etymology">
<paragraph id="9C2936B9FF95FFFA09C4FB0DFE919B2A" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="81">
<emphasis id="AEE2EAABFF95FFFA09C4FB0DFEE09B16" bold="true" box="[199,334,1267,1293]" pageId="4" pageNumber="81">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="5B964D3AFF95FFFA0B44FB0DFD7B9B17" authorityName="Felix &amp; Silva &amp; Quintas &amp; Pecly &amp; Froza &amp; Mejdalani" authorityYear="2025" box="[583,725,1267,1292]" class="Insecta" family="Cicadellidae" genus="Juliaca" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="species" species="nigrosignata">
<emphasis id="AEE2EAABFF95FFFA0B44FB0DFD7B9B17" box="[583,725,1267,1292]" italics="true" pageId="4" pageNumber="81">nigrosignata</emphasis>
</taxonomicName>
, refers to the conspicuous black spot at the median portion of crown (
<figureCitation id="04AD2A3CFF95FFFA09EFFAE9FE809B2A" box="[236,302,1303,1329]" captionStart="FIGURES 112" captionStartId="2.[151,264,1669,1693]" captionTargetBox="[162,1428,181,1631]" captionTargetId="figure-37@2.[151,1436,181,1645]" captionTargetPageId="2" captionText="FIGURES 112. Juliaca nigrosignata sp. nov. Male (17): 1, dorsal habitus (length 6.38 mm); 2, pygofer, valve, subgenital plate, and anal tube, lateral view; 3, left subgenital plate, ventral view; 4, left style and connective, dorsal view; 5, ejaculatory bulb and shaft of aedeagus, lateral view; 6, ejaculatory bulb and shaft of aedeagus, ventral view; 7, paraphyses, dorsal view. Female (812): 8, sternite VII, ventral view; 9, pygofer, lateral view; 10, ovipositor valvulae II, lateral view; 11, apical portion of valvula II, lateral view; 12, teeth at median portion of valvula II, lateral view. Scale bars of figures 2, 9, and 10 = 0.5 mm." pageId="4" pageNumber="81">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="9C2936B9FF95FFFA09C4FAC5FAB79B4E" blockId="4.[151,1437,151,1509]" box="[199,1305,1338,1365]" pageId="4" pageNumber="81">
<emphasis id="AEE2EAABFF95FFFA09C4FAC5FE149B4F" bold="true" box="[199,442,1338,1365]" pageId="4" pageNumber="81">Known distribution.</emphasis>
<collectingCountry id="E4817629FF95FFFA08C2FAC5FD8E9B4E" box="[449,544,1339,1365]" name="Ecuador" pageId="4" pageNumber="81">Ecuador</collectingCountry>
, Province of
<collectingRegion id="5E52F85BFF95FFFA0BB7FAC5FC9D9B4E" box="[692,819,1339,1365]" country="Ecuador" name="Sucumbios" pageId="4" pageNumber="81">Sucumbíos</collectingRegion>
, sub-Andean montane forest (
<figureCitation id="04AD2A3CFF95FFFA0D80FAC5FAA59B4E" box="[1155,1291,1339,1365]" captionStart="FIGURES 1314" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[251,1342,181,1920]" captionTargetId="figure-21@3.[244,1342,181,1920]" captionTargetPageId="3" captionText="FIGURES 1314.13, mapshowing the locationof the type locality of Juliacanigrosignatasp. nov. in theProvince of Sucumbíos, Ecuador. 14, sub-Andean montane forest at the type locality (about 2,000 m a.s.l.). Photograph by Adriano Brilhante Kury." pageId="4" pageNumber="81">Figs. 13, 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D48C6532FF95FFFA09C4FAA0FC389BFE" pageId="4" pageNumber="81" type="materials_examined">
<paragraph id="9C2936B9FF95FFFA09C4FAA0FC389BFE" blockId="4.[151,1437,151,1509]" pageId="4" pageNumber="81">
<emphasis id="AEE2EAABFF95FFFA09C4FAA0FE1F9B63" bold="true" box="[199,433,1374,1400]" pageId="4" pageNumber="81">Material examined.</emphasis>
<materialsCitation id="2CFE3CE4FF95FFFA08BBFAA1FEAD9BDA" collectingDate="2014-02-12" collectionCode="QCAZI" collectorName="A Giupponi" country="Ecuador" elevation="2006" latitude="0.51625" location="Male" longLatPrecision="1" longitude="-77.53477" municipality="Province" pageId="4" pageNumber="81" specimenCount="1" stateProvince="Sucumbios" typeStatus="holotype">
<collectingCountry id="E4817629FF95FFFA08BBFAA1FDB99B62" box="[440,535,1375,1401]" name="Ecuador" pageId="4" pageNumber="81">Ecuador</collectingCountry>
,
<collectingMunicipality id="7C4DACC3FF95FFFA0B20FAA1FD299B62" box="[547,647,1375,1401]" pageId="4" pageNumber="81">Province</collectingMunicipality>
of
<collectingRegion id="5E52F85BFF95FFFA0BAEFAA1FC839B62" box="[685,813,1375,1401]" country="Ecuador" name="Sucumbios" pageId="4" pageNumber="81">Sucumbíos</collectingRegion>
.
<location id="99496062FF95FFFA0A3BFAA1FCDC9B62" LSID="urn:lsid:plazi:treatment:143F87AFFF90FFFA0994FAE8FC389BFE:99496062FF95FFFA0A3BFAA1FCDC9B62" box="[824,882,1375,1401]" country="Ecuador" latitude="0.51625" longLatPrecision="1" longitude="-77.53477" municipality="Province" name="Male" pageId="4" pageNumber="81" stateProvince="Sucumbios">Male</location>
<typeStatus id="432D881BFF95FFFA0A7AFAA1FC719B62" box="[889,991,1375,1401]" pageId="4" pageNumber="81" type="holotype">holotype</typeStatus>
: “Ponto 7.
<collectingCountry id="E4817629FF95FFFA0D5FFAA1FB159B62" box="[1116,1211,1375,1401]" name="Ecuador" pageId="4" pageNumber="81">Ecuador</collectingCountry>
, \
<collectingRegion id="5E52F85BFF95FFFA0DD5FAA1FAFB9B62" box="[1238,1365,1375,1401]" country="Ecuador" name="Sucumbios" pageId="4" pageNumber="81">Sucumbíos</collectingRegion>
, rio,
<quantity id="5B6E9B5CFF95FFFA0C8DFAA0FF159B86" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="81" unit="km" value="5.0">5 km</quantity>
ao \ norte de La Bonita.”; “
<geoCoordinate id="F9A2507EFF95FFFA08EEFA7DFD2C9B86" box="[493,642,1410,1437]" degrees="00.51625" direction="north" orientation="latitude" pageId="4" pageNumber="81" precision="1" value="0.51625">N 00.51625°</geoCoordinate>
;
<geoCoordinate id="F9A2507EFF95FFFA0B8FFA7DFC9D9B86" box="[652,819,1410,1437]" degrees="077.53477" direction="west" orientation="longitude" pageId="4" pageNumber="81" precision="1" value="-77.53477">W 077.53477°</geoCoordinate>
. \ Alt.
<quantity id="5B6E9B5CFF95FFFA0A78FA7DFC7C9B86" box="[891,978,1411,1437]" metricMagnitude="3" metricUnit="m" metricValue="2.006" pageId="4" pageNumber="81" unit="m" value="2006.0">
<elevation id="17BBD18AFF95FFFA0A78FA7DFC7C9B86" box="[891,978,1411,1437]" metricMagnitude="3" metricUnit="m" metricValue="2.006" pageId="4" pageNumber="81" unit="m" value="2006.0">2006 m</elevation>
</quantity>
.
<date id="E8281079FF95FFFA0ADEFA7DFBFF9B86" box="[989,1105,1411,1437]" pageId="4" pageNumber="81" value="2014-02-12">
<collectingDate id="F86CE991FF95FFFA0ADEFA7DFBFF9B86" box="[989,1105,1411,1437]" pageId="4" pageNumber="81" value="2014-02-12">12.ii.2014</collectingDate>
</date>
, A \ Kury &amp;
<collectorName id="3163536FFF95FFFA0DDEFA7DFACE9B86" box="[1245,1376,1411,1437]" pageId="4" pageNumber="81">A Giupponi</collectorName>
leg.” (
<collectionCode id="FA87AE7CFF95FFFA099CFA59FF559BDA" box="[159,251,1447,1473]" pageId="4" pageNumber="81">QCAZI</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="2CFE3CE4FF95FFFA080CFA59FC3C9BFE" collectingDate="2014-02-12" collectionCode="QCAZI, CEIOC, MNRJ, MELQ" collectorName="A Giupponi" country="Ecuador" elevation="2006" latitude="0.51625" location="Male" longLatPrecision="1" longitude="-77.53477" municipality="Province" pageId="4" pageNumber="81" specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="Sucumbios" typeStatus="paratype">
<typeStatus id="432D881BFF95FFFA080CFA59FE2E9BDA" box="[271,384,1447,1473]" pageId="4" pageNumber="81" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="8A90FD30FF95FFFA088FFA59FDAF9BDA" box="[396,513,1447,1473]" count="5" pageId="4" pageNumber="81" type="male">five males</specimenCount>
and
<specimenCount id="8A90FD30FF95FFFA0B38FA59FD789BDA" box="[571,726,1447,1473]" count="3" pageId="4" pageNumber="81" type="female">three females</specimenCount>
, same data as the holotype (one couple at
<collectionCode id="FA87AE7CFF95FFFA0DB8FA59FABA9BDA" box="[1211,1300,1447,1473]" pageId="4" pageNumber="81">QCAZI</collectionCode>
; one couple plus
<specimenCount id="8A90FD30FF95FFFA09CEFA35FE9C9BFE" box="[205,306,1483,1509]" count="1" pageId="4" pageNumber="81" type="male">one male</specimenCount>
at
<collectionCode id="FA87AE7CFF95FFFA0856FA35FE039BFE" box="[341,429,1483,1509]" pageId="4" pageNumber="81">CEIOC</collectionCode>
; one couple at
<collectionCode id="FA87AE7CFF95FFFA0B57FA35FD0C9BFE" box="[596,674,1483,1509]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="4" pageNumber="81">MNRJ</collectionCode>
;
<specimenCount id="8A90FD30FF95FFFA0BAEFA35FCBD9BFE" box="[685,787,1483,1509]" count="1" pageId="4" pageNumber="81" type="male">one male</specimenCount>
at
<collectionCode id="FA87AE7CFF95FFFA0A35FA35FC259BFE" box="[822,907,1483,1509]" pageId="4" pageNumber="81">MELQ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,391 @@
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<mods:mods id="75B62035CF8B32840FFDFA08DD9FDFDD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3F6EF05447BC8E24AA84FB7DA80A17E6">New Ecuadorian and Colombian Lamiinae (Coleoptera, Cerambycidae)</mods:title>
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<mods:namePart id="390730A379339B5B6CF7D37F7ABA2CB7">Vlasak, Josef</mods:namePart>
<mods:affiliation id="7C1EC9CD9C3284C0E050645698EF52C7">207 Silverbrook Drive, Schwenksville, PA 19473, U. S. A.</mods:affiliation>
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<mods:namePart id="3CC37CE8CFF3163FA589A38109BD9E8C">Nascimento, Francisco Eriberto Lima</mods:namePart>
<mods:affiliation id="B14CD15B4ACD529DAFE99BF3889EB725">Laboratório de Invertebrados - LABIN, Instituto de Ciências Exatas e Naturais - ICEN, Universidade Federal de Rondonopolis, Rondonópolis, Mato Grosso, Brazil</mods:affiliation>
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</mods:role>
<mods:namePart id="BE9D19D3CB036A6EA380815BCACCCCFE">Santos-Silva, Antonio</mods:namePart>
<mods:affiliation id="B4B5745D76B382C7EC92E62CB3C89376">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="A1E01BFE11CDDC9654ECF74B53DA0CC0">Zootaxa</mods:title>
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<mods:date id="E3CDBED3FE09A9FD7AC62871F083215A">2025</mods:date>
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<treatment id="892487AC7602FFE54FD1D88EF1CAFE07" LSID="urn:lsid:plazi:treatment:892487AC7602FFE54FD1D88EF1CAFE07" httpUri="http://treatment.plazi.org/id/892487AC7602FFE54FD1D88EF1CAFE07" lastPageId="6" lastPageNumber="7" pageId="1" pageNumber="2">
<subSubSection id="499765317602FFE24FD1D88EF17BFD67" box="[151,571,718,745]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="013236BA7602FFE24FD1D88EF17BFD67" blockId="1.[151,571,718,781]" box="[151,571,718,745]" pageId="1" pageNumber="2">
<heading id="5A7A81D67602FFE24FD1D88EF17BFD67" bold="true" box="[151,571,718,745]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="33F9EAA87602FFE24FD1D88EF17BFD67" bold="true" box="[151,571,718,745]" pageId="1" pageNumber="2">
<taxonomicName id="C68D4D397602FFE24FD1D88EF296FD67" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[151,470,718,745]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="chloroxanthon" status="sp. nov.">
<emphasis id="33F9EAA87602FFE24FD1D88EF296FD67" bold="true" box="[151,470,718,745]" italics="true" pageId="1" pageNumber="2">Nealcidion chloroxanthon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28CA57D37602FFE24E9BD88FF17BFD67" box="[477,571,719,745]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="499765317602FFE74FD1D8B3F09FFB67" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="description">
<paragraph id="013236BA7602FFE24FD1D8B3F259FC83" blockId="1.[151,571,718,781]" box="[151,281,755,781]" pageId="1" pageNumber="2">
(
<figureCitation id="99B62A3F7602FFE24FD9D8B3F251FC83" box="[159,273,755,781]" captionStart-0="FIGURES 15" captionStart-1="FIGURES 610" captionStartId-0="2.[152,270,1943,1967]" captionStartId-1="5.[152,270,1891,1915]" captionTargetBox-0="[166,1421,181,1919]" captionTargetBox-1="[162,1423,462,1863]" captionTargetId-0="figure-23@2.[166,1421,181,1919]" captionTargetId-1="figure-151@5.[151,1435,453,1868]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionText-0="FIGURES 15. Nealcidion chloroxanthon sp. nov., holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Humeral area." captionText-1="FIGURES 610. Nealcidion chloroxanthon sp. nov., paratypes from Ecuador (Napo), dorsal habitus: 6) Male from Pacto Sumaco; 7) Female from Papallacta; 8) Male from Papallacta; 9) Female from Papallacta; 10) Female from Papallacta." figureDoi-0="http://doi.org/10.5281/zenodo.15012340" figureDoi-1="http://doi.org/10.5281/zenodo.15012346" httpUri-0="https://zenodo.org/record/15012340/files/figure.png" httpUri-1="https://zenodo.org/record/15012346/files/figure.png" pageId="1" pageNumber="2">Figs 110</figureCitation>
)
</paragraph>
<paragraph id="013236BA7602FFE24FD1D97AF203F9AB" blockId="1.[151,1437,826,2041]" pageId="1" pageNumber="2">
<emphasis id="33F9EAA87602FFE24FD1D97AF2A6FCDB" bold="true" box="[151,486,826,853]" pageId="1" pageNumber="2">
Description.
<typeStatus id="DE3688187602FFE24E72D97BF2E1FCDB" box="[308,417,827,853]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="99B62A3F7602FFE24EBFD97BF122FCDB" box="[505,610,826,853]" captionStart="FIGURES 15" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[166,1421,181,1919]" captionTargetId="figure-23@2.[166,1421,181,1919]" captionTargetPageId="2" captionText="FIGURES 15. Nealcidion chloroxanthon sp. nov., holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Humeral area." figureDoi="http://doi.org/10.5281/zenodo.15012340" httpUri="https://zenodo.org/record/15012340/files/figure.png" pageId="1" pageNumber="2">Figs 15</figureCitation>
). Head capsule dark brown; ventral mouthparts reddish brown, except palpomeres blackish with the apex of maxillary palpomeres IV and labial palpomeres III reddish brown; anteclypeus mostly brown posteriorly, with area close to labrum dark brown; labrum dark brown on posterior 3/4, brown on anterior quarter; mandibles dark reddish brown on basal2/3,blackish on apical third; scape irregularly greenish yellow on basal third and mostly dark brown on remaining dorsal surface, greenish yellow laterally, gradually darker from middle and becoming reddish brown close to apex, and mostly orangish brown on ventral surface; pedicel mostly dark brown; antennomere III dark brown on apex, irregularly, mostly dark brown dorsally, and irregularly orangish brown on remaining surface, this area slightly darker near dark-brown apex; antennomere IV mostly dull yellowish brown, except dark-brown apex; antennomeres VX light reddish brown except dark-brown apex. Prothorax dark brown. Ventral surface of meso- and metathorax black except dark-brown central area of mesoventrite and base of mesoventral process and orangish-brown area close to metacoxal cavities. Scutellum dark brown. Elytra dark brown basally, gradually dark green on remaining surface, this area more brownish green depending on light intensity and lighter posteriorly. Pro- and mesocoxae orange with irregular dark areas; metacoxae dark brown laterally, orange on remaining surface. Trochanters orangish. Femora orangish basally, irregularly, both orangish and greenish on remaining peduncle, light green on femoral clubs. Basal 2/3 of tibiae mostly light green, apical third of protibiae both dark brown and dark orangish brown, apical third of mesotibiae dark brown on dorsal sulcus, dark orangish brown on remaining surface except blackish dorsal apex and dark-brown ventral apex, and apical third of metatibiae brown dorsally, dark brown on apex of ventral surface, and orangish brown on remaining surface. Protarsi dark brown basally, gradually blackish toward apex; meso- and metatarsomeres I orange except dark brown apical third; meso- and metatarsomeres IIIII dark brown and IVV and claws black. Abdominal ventrites dark brown laterally, black centrally.
</paragraph>
<paragraph id="013236BA7602FFE04F81DC6FF04AFD0B" blockId="1.[151,1437,826,2041]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<emphasis id="33F9EAA87602FFE24F81DC6FF24EF9C7" bold="true" box="[199,270,1583,1609]" pageId="1" pageNumber="2">Head.</emphasis>
Frons, vertex, antennal tubercles, genae, and area behind eyes with dense greenish-yellow pubescence, except glabrous median groove, this area diamond-shaped between antennal tubercles and upper eye lobes, and apex of genae, pubescence slightly sparser on antennal tubercles and slightly darker on superior region of frons; frons with one long, erect brown seta close to eyes; frons, vertex, antennal tubercles, and area behind eyes abundantly, finely, shallowly punctate. Wide central area of postclypeus with dense greenish-yellow pubescence close to frons, almost glabrous centrally, and with moderately abundant yellowish-white setae directed forward close to anteclypeus; with a few long, erect brown setae interspersed posteriorly. Sides of postclypeus glabrous. Labrum with moderately sparse yellowish-white pubescence posteriorly, long, erect setae directed forward about middle, setae yellowish white centrally, yellowish brown laterally, and glabrous on anterior region, except fringe of yellowish-brown setae on anterior margin. Gulamentum glabrous, except yellowish-white pubescence on intermaxillary process. Maximum diameter of upper eye lobes slightly shorter than basal diameter of scape; distance between upper eye lobes 0.36 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.64 times distance between outer margins of eyes. Antennae 1.9 times elytral length, reaching elytral apex at basal third of antennomere VII. Scape with moderately abundant yellowish pubescence not obscuring integument, except dark-brown pubescence not obscuring integument on dark dorsal area. Pedicel with yellowish-white pubescence not obscuring integument on part of basal half and dark-brown pubescence not obscuring integument on remaining surface; with a few moderately long, erect dark-brown setae ventrally. Antennomere III mostly with dark-brown pubescence not obscuring integument dorsally and on apical dark area and yellowish pubescence not obscuring integument on remaining surface; with moderately long, erect, thick dark-brown setae interspersed ventrally and short dark-brown setae directed backward apically on dorsal and lateral surfaces. Antennomeres IVXI with abundant yellowish pubescence not obscuring integument on light integumental area, pubescence appearing to be darker depending on light intensity and source, and abundant dark-brown pubescence not obscuring integument on dark integumental area; IV with a few short, erect, thick dark-brown setae interspersed ventrally, distinctly more abundant apically; VIVII with a few short, erect, thick setae on ventral apex; IVVII with short, thick dark-brown setae directed backward on apex of dorsal and lateral surfaces; apex of XI with short, bristly yellowish-white setae. Antennal formula (ratio) based on antennomere III: scape = 0.92; pedicel = 0.12; IV = 0.76; V = 0.53; VI = 0.46; VII = 0.43; VIII = 0.39; IX = 0.37; X = 0.35; XI = 0.35.
</paragraph>
<caption id="55F266327601FFE14FDEDDD7F14EF85D" ID-DOI="http://doi.org/10.5281/zenodo.15012340" ID-Zenodo-Dep="15012340" httpUri="https://zenodo.org/record/15012340/files/figure.png" pageId="2" pageNumber="3" startId="2.[152,270,1943,1967]" targetBox="[166,1421,181,1919]" targetPageId="2" targetType="figure">
<paragraph id="013236BA7601FFE14FDEDDD7F14EF85D" blockId="2.[151,1437,1943,2004]" pageId="2" pageNumber="3">
<emphasis id="33F9EAA87601FFE14FDEDDD7F200F821" bold="true" box="[152,320,1943,1967]" pageId="2" pageNumber="3">FIGURES 15.</emphasis>
<taxonomicName id="C68D4D397601FFE14E01DDD8F10FF83E" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[327,591,1944,1968]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="chloroxanthon" status="sp. nov.">
<emphasis id="33F9EAA87601FFE14E01DDD8F10FF83E" box="[327,591,1944,1968]" italics="true" pageId="2" pageNumber="3">Nealcidion chloroxanthon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28CA57D37601FFE14D13DDD8F1E6F821" box="[597,678,1944,1967]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male:
<emphasis id="33F9EAA87601FFE14C0ADDD7F022F821" bold="true" box="[844,866,1943,1967]" pageId="2" pageNumber="3">1)</emphasis>
Dorsal habitus;
<emphasis id="33F9EAA87601FFE14B41DDD7F75DF821" bold="true" box="[1031,1053,1943,1967]" pageId="2" pageNumber="3">2)</emphasis>
Ventral habitus;
<emphasis id="33F9EAA87601FFE14B8EDDD7F79DF821" bold="true" box="[1224,1245,1943,1967]" pageId="2" pageNumber="3">3)</emphasis>
Lateral habitus;
<emphasis id="33F9EAA87601FFE14AC1DDD7F6DDF821" bold="true" box="[1415,1437,1943,1967]" pageId="2" pageNumber="3">4)</emphasis>
Head, frontal view;
<emphasis id="33F9EAA87601FFE14E24DDFBF237F85D" bold="true" box="[354,375,1979,2003]" pageId="2" pageNumber="3">5)</emphasis>
Humeral area.
</paragraph>
</caption>
<paragraph id="013236BA7600FFE74F81D8CFF3B9FC47" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="33F9EAA87600FFE04F81D8CFF267FD27" bold="true" box="[199,295,655,681]" pageId="3" pageNumber="4">Thorax.</emphasis>
Prothorax wider than long; anterior constriction well marked; sides divergent from anterior constriction to lateral tubercle, convergent from lateral tubercle to posterior sixth, then parallel-sided toward posterolateral angle; lateral tubercles large, subconical. Pronotum densely, minutely punctate, except smooth area close to anterior margin and posterior sulcus, and transverse row with coarse punctures near posterior margin, punctures not follow toward sides of prothorax; posterior sulcus well marked laterally, almost absent centrally; widely, longitudinally depressed centrally; with dense, wide, longitudinal greenish-yellow pubescent band centrally, from anterior to posterior margins, abundant greenish-yellow pubescence laterally, pubescence denser close to lateral tubercles of prothorax and lighter than on central pubescent band, and abundant brownish-yellow pubescence not obscuring integument on remaining surface, except glabrous anterior constriction and posterior sulcus; with a few long, erect dark-brown setae on sides of posterior third. Sides of prothorax with abundant greenish-yellow pubescence, denser on some areas, except glabrous anterior and posterior sulcus. Prosternum with dense greenish-yellow pubescence laterally and abundant greenish-yellow pubescence centrally partially obscuring integument, except glabrous anterior sulcus, narrow white pubescent area close to sides of procoxal cavities, and glabrous remaining area close to procoxal cavities. Prosternal process with abundant greenish-yellow pubescence partially obscuring integument, except glabrous sides and narrow transverse white pubescent area on apex; narrowest area 0.38 times procoxal width. Mesoventrite with abundant white pubescence not obscuring integument, except dense greenish-yellow pubescence close to superior region of mesanepisterna. Mesanepisterna and mesepimera with dense greenish-yellow pubescence. Mesoventral process with somewhat abundant whitish pubescence laterally, abundant whitish pubescence centrally, with greenish-yellow pubescence interspersed anteriorly, and sparse, both whitish and greenish-yellow pubescence on remaining surface; sides convergent from base to near apex, then slightly widened; narrowest area 0.44 times mesocoxal width. Metanepisterna with abundant whitish pubescence not obscuring integument on anterior third and dense greenish-yellow pubescence on remaining surface. Metaventrite with dense greenish-yellow pubescence laterally, except white pubescence close to mesocoxal cavities, whitish pubescence denser toward metanepisterna, and abundant, both whitish and greenish-yellow pubescence close to metacoxal cavities; remaining surface with abundant whitish pubescence not obscuring integument, with greenish-white pubescence interspersed on some areas, except very sparse whitish pubescence on large posterocentral region, absent on metathoracic discrimen and posterior area close to abdominal process. Scutellum glabrous anterocentrally, dense greenish-yellow pubescence on remaining surface, with whitish pubescence interspersed on sides of basal area.
<emphasis id="33F9EAA87600FFE04C74DC1BF0C5F9FB" bold="true" box="[818,901,1627,1653]" pageId="3" pageNumber="4">Elytra.</emphasis>
Abundantly, coarsely punctate on anterior sixth, punctures gradually sparser, finer from anterior sixth to posterior fifth, absent on posterior fifth; centrobasal crest large, elevated, rounded and without erect setae dorsally; with well-marked, longitudinal carina from centrobasal crest to posterolateral angle, carina arched inward on its anterior quarter and slightly obliquely inclined toward posterolateral angle on its posterior seventh; area between dorsal carina and suture depressed; with abundant greenish-yellow pubescence, partially obscuring integument on some areas, dense around centrobasal crest, except: four irregular, large dark-green pubescence areas, from epipleural margin to dorsal surface, pubescence lighter on posterior area, only the third reaching suture; dashed, slightly oblique white pubescent band on basal half of dorsal surface; dashed white pubescent band on epipleural margin, this area partially greenish-yellow anteriorly; dashed white pubescent band along suture, except area close to scutellum with white pubescent band interspersed among greenish-yellow pubescence; somewhat small, irregular white pubescent spots on posterior half; area between white pubescent maculae on dashed bands with somewhat abundant dark-green pubescence not obscuring integument. Sides of basal area with minute, erect, thick dark-brown setae (
<figureCitation id="99B62A3F7607FFE74C14DAFBF0D6FF5B" box="[850,918,186,213]" captionStart="FIGURES 15" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[166,1421,181,1919]" captionTargetId="figure-23@2.[166,1421,181,1919]" captionTargetPageId="2" captionText="FIGURES 15. Nealcidion chloroxanthon sp. nov., holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Humeral area." figureDoi="http://doi.org/10.5281/zenodo.15012340" httpUri="https://zenodo.org/record/15012340/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
). Apex obliquely concave, making outer angle acutely triangularly elongated and sutural angle slightly projected.
<emphasis id="33F9EAA87607FFE74CCCDA9FF089FF77" bold="true" box="[906,969,223,249]" pageId="4" pageNumber="5">Legs.</emphasis>
Coxae with abundant yellow pubescence not obscuring integument, pubescence more greenish yellow depending on light intensity, except abundant white pubescence not obscuring integument on part of area close to margins. Trochanters with moderately abundant white pubescence not obscuring integument. Femora with abundant greenish-yellow pubescence not obscuring integument, except profemoral peduncle with moderately abundant white pubescence not obscuring integument, and base of dorsal surface and entire ventral surface of meso- and metafemoral peduncles with abundant white pubescence not obscuring integument. Tibiae with abundant greenish-yellow pubescence partially obscuring integument, except; base of ventral surface glabrous; dorsal apex of protibiae glabrous; protibiae with short, erect dark-brown setae ventrally about middle; protibiae with dense, bristly yellowish-brown pubescence ventrally on apical half; pubescence sparse on posterior third of meso- and metatibiae; posterior third of meso- and metatibiae with short, erect, thick dark-brown setae, setae denser on mesotibiae; area close to apex of ventral surface of meso- and metatibiae with dense, bristly yellowish-brown pubescence; and posterior third of ventral surface of metatibiae with somewhat sparse, short, erect dark-brown setae. Protibiae arched, widened inward apically. Dorsal surface of tarsomeres I with abundant light greenish-yellow pubescence not obscuring integument, pubescence lighter on meso- and metatarsi; pro- and mesotarsomeres IIIII with somewhat abundant greenish-yellow pubescence not obscuring integument; metatarsomeres IIIII with abundant greenish-yellow pubescence laterally and sparser, somewhat greenish pubescence centrally; pro- and mesotarsomeres IVV with somewhat abundant dark-brown pubescence not obscuring integument; metatarsomeres IVV with abundant both light greenish-yellow and whitish pubescence on basal half, and somewhat abundant dark-brown pubescence not obscuring integument on posterior half; tarsomeres with a few long, erect dark-brown setae except on IV; metatarsomere I 1.25 times longer than IIIII together.
</paragraph>
<paragraph id="013236BA7607FFE74F81D992F0E8FB9F" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="5">
<emphasis id="33F9EAA87607FFE74F81D992F203FC62" bold="true" box="[199,323,978,1004]" pageId="4" pageNumber="5">Abdomen.</emphasis>
Ventrites with dense greenish-yellow pubescence laterally and sparse whitish pubescence centrally; apex of ventrite 5 widely concave, making the outer angles spiniform.
</paragraph>
<paragraph id="013236BA7607FFE74F81DE5BF028FBD7" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="5">
<emphasis id="33F9EAA87607FFE74F81DE5BF25EFBBB" bold="true" box="[199,286,1051,1077]" pageId="4" pageNumber="5">Female</emphasis>
(
<figureCitation id="99B62A3F7607FFE74E6DDE5BF2F6FBBB" box="[299,438,1051,1077]" captionStart="FIGURES 610" captionStartId="5.[152,270,1891,1915]" captionTargetBox="[162,1423,462,1863]" captionTargetId="figure-151@5.[151,1435,453,1868]" captionTargetPageId="5" captionText="FIGURES 610. Nealcidion chloroxanthon sp. nov., paratypes from Ecuador (Napo), dorsal habitus: 6) Male from Pacto Sumaco; 7) Female from Papallacta; 8) Male from Papallacta; 9) Female from Papallacta; 10) Female from Papallacta." figureDoi="http://doi.org/10.5281/zenodo.15012346" httpUri="https://zenodo.org/record/15012346/files/figure.png" pageId="4" pageNumber="5">Figs 7, 9, 10</figureCitation>
). Similar to male, differing mainly by femoral clubs stouter; additionally, antennae slightly shorter, reaching elytral apex at basal third of antennomere VIII.
</paragraph>
<paragraph id="013236BA7607FFE74F81DE22F09FFB67" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="5">
<emphasis id="33F9EAA87607FFE74F81DE22F2DBFBF2" bold="true" box="[199,411,1122,1148]" pageId="4" pageNumber="5">Dimensions (mm)</emphasis>
(
<typeStatus id="DE3688187607FFE74EECDE23F14EFBF3" box="[426,526,1123,1149]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
male/
<typeStatus id="DE3688187607FFE74D12DE23F1F4FBF3" box="[596,692,1123,1149]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
males/
<typeStatus id="DE3688187607FFE74C43DE23F026FBF3" box="[773,870,1123,1149]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
females). Total length, 8.80/7.958.50/6.709.00; prothoracic length, 1.50/1.351.50/1.051.50; anterior prothoracic width, 1.55/1.551.60/1.301.70; posterior prothoracic width, 1.85/1.751.80/1.501.95; maximum prothoracic width, 2.40/2.202.35/1.852.35; humeral width, 3.15/2.953.05/2.353.35; elytral length, 6.45/5.906.30/4.856.75.
</paragraph>
</subSubSection>
<subSubSection id="499765317607FFE74F81DEB3F01CFADB" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="013236BA7607FFE74F81DEB3F01CFADB" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="5">
<emphasis id="33F9EAA87607FFE74F81DEB3F234FA82" bold="true" box="[199,372,1266,1293]" pageId="4" pageNumber="5">Type material.</emphasis>
<materialsCitation id="B1E53CE77607FFE74E3FDEB3F6D9FA83" box="[377,1433,1266,1293]" collectingDate="2022-11-25" collectionCode="MZSP" collectorName="J. Vlasak" country="Ecuador" elevation="2100" location="Napo" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
<typeStatus id="DE3688187607FFE74E3FDEB3F2A1FA83" box="[377,481,1267,1293]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
male from
<collectingCountry id="799A762A7607FFE74D19DEB3F1A9FA83" box="[607,745,1267,1293]" name="Ecuador" pageId="4" pageNumber="5">ECUADOR</collectingCountry>
,
<emphasis id="33F9EAA87607FFE74DB5DEB3F070FA82" box="[755,816,1267,1292]" italics="true" pageId="4" pageNumber="5">
<collectingRegion id="C349F8587607FFE74DB5DEB3F070FA82" box="[755,816,1267,1292]" country="Ecuador" name="Napo" pageId="4" pageNumber="5">Napo</collectingRegion>
</emphasis>
:
<quantity id="C6759B5F7607FFE74C7BDEB3F0CEFA83" box="[829,910,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="5" unit="m" value="2100.0">
<elevation id="8AA0D1897607FFE74C7BDEB3F0CEFA83" box="[829,910,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="5" unit="m" value="2100.0">2100m</elevation>
</quantity>
, Cosanga,
<date id="7533107A7607FFE74B42DEB2F7D7FA83" box="[1028,1175,1266,1293]" pageId="4" pageNumber="5" value="2022-11-25">
<collectingDate id="6577E9927607FFE74B42DEB2F7D7FA83" box="[1028,1175,1266,1293]" pageId="4" pageNumber="5" value="2022-11-25">25 Nov 2022</collectingDate>
</date>
,
<collectorName id="AC78536C7607FFE74BE6DEB3F645FA83" box="[1184,1285,1267,1293]" pageId="4" pageNumber="5">J. Vlasak</collectorName>
leg. (
<collectionCode id="679CAE7F7607FFE74A07DEB3F6D0FA83" box="[1345,1424,1267,1293]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="5" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B1E53CE77607FFE74FD1DF57F661FABF" box="[151,1313,1302,1329]" collectingDate="2024-09-05" collectingDateMax="2024-09-07" collectingDateMin="2024-09-05" collectionCode="JVCO" collectorName="Pacto Sumaco &amp; J. Vlasak" country="Ecuador" elevation="1500" location="Napo" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<typeStatus id="DE3688187607FFE74FD1DF57F244FABF" box="[151,260,1303,1329]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
<collectingCountry id="799A762A7607FFE74E67DF57F2EBFABF" box="[289,427,1303,1329]" name="Ecuador" pageId="4" pageNumber="5">ECUADOR</collectingCountry>
,
<emphasis id="33F9EAA87607FFE74EFEDF57F2B5FABE" box="[440,501,1303,1328]" italics="true" pageId="4" pageNumber="5">
<collectingRegion id="C349F8587607FFE74EFEDF57F2B5FABE" box="[440,501,1303,1328]" country="Ecuador" name="Napo" pageId="4" pageNumber="5">Napo</collectingRegion>
</emphasis>
:
<specimenCount id="178BFD337607FFE74D40DF57F114FABF" box="[518,596,1303,1329]" count="1" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
<quantity id="C6759B5F7607FFE74D18DF56F1EEFABF" box="[606,686,1302,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="m" value="1500.0">
<elevation id="8AA0D1897607FFE74D18DF56F1EEFABF" box="[606,686,1302,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<collectorName id="AC78536C7607FFE74DFBDF57F022FABF" box="[701,866,1303,1329]" pageId="4" pageNumber="5">Pacto Sumaco</collectorName>
,
<date id="7533107A7607FFE74C36DF56F752FABF" box="[880,1042,1302,1329]" pageId="4" pageNumber="5" value="2024-09-05" valueMax="2024-09-07" valueMin="2024-09-05">
<collectingDate id="6577E9927607FFE74C36DF56F752FABF" box="[880,1042,1302,1329]" pageId="4" pageNumber="5" value="2024-09-05" valueMax="2024-09-07" valueMin="2024-09-05">57 Sep 2024</collectingDate>
</date>
,
<collectorName id="AC78536C7607FFE74B66DF57F7C9FABF" box="[1056,1161,1303,1329]" pageId="4" pageNumber="5">J. Vlasak</collectorName>
leg. (
<collectionCode id="679CAE7F7607FFE74B8BDF57F659FABF" box="[1229,1305,1303,1329]" pageId="4" pageNumber="5">JVCO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B1E53CE77607FFE74A69DF57F018FADB" collectingDate="2024-09-01" collectionCode="JVCO" collectorName="Papallacta &amp; J. Vlasak" country="Ecuador" elevation="3400" pageId="4" pageNumber="5" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<specimenCount id="178BFD337607FFE74A69DF57F6C0FABF" box="[1327,1408,1303,1329]" count="1" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
,
<specimenCount id="178BFD337607FFE74AC8DF57F3AEFADB" count="3" pageId="4" pageNumber="5" type="female">3 females</specimenCount>
<quantity id="C6759B5F7607FFE74FB3DF7BF206FADB" box="[245,326,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="5" unit="m" value="3400.0">
<elevation id="8AA0D1897607FFE74FB3DF7BF206FADB" box="[245,326,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="4" pageNumber="5" unit="m" value="3400.0">3400m</elevation>
</quantity>
,
<collectorName id="AC78536C7607FFE74E17DF7BF287FADB" box="[337,455,1339,1365]" pageId="4" pageNumber="5">Papallacta</collectorName>
,
<date id="7533107A7607FFE74E94DF7BF113FADB" box="[466,595,1339,1365]" pageId="4" pageNumber="5" value="2024-09-01">
<collectingDate id="6577E9927607FFE74E94DF7BF113FADB" box="[466,595,1339,1365]" pageId="4" pageNumber="5" value="2024-09-01">1 Sep 2024</collectingDate>
</date>
,
<collectorName id="AC78536C7607FFE74D18DF7BF184FADB" box="[606,708,1339,1365]" pageId="4" pageNumber="5">J. Vlasak</collectorName>
leg. (
<collectionCode id="679CAE7F7607FFE74C45DF7BF00FFADB" box="[771,847,1339,1365]" pageId="4" pageNumber="5">JVCO</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="499765317607FFE74F81DF1FF098FA13" pageId="4" pageNumber="5" type="etymology">
<paragraph id="013236BA7607FFE74F81DF1FF098FA13" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="5">
<emphasis id="33F9EAA87607FFE74F81DF1FF20EFAF7" bold="true" box="[199,334,1375,1401]" pageId="4" pageNumber="5">Etymology.</emphasis>
The specific epithet
<taxonomicName id="C68D4D397607FFE74D07DF1FF1B4FAF7" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[577,756,1375,1401]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chloroxanthon">“chloroxanthon</taxonomicName>
”, from Greek “χλωρός” (chloros)-pale green, and “ξανθός” (xanthos)-yellow, refers to the overall greenish-yellow color of this beetle.
</paragraph>
</subSubSection>
<subSubSection id="499765317607FFE54F81DFE7F1CAFE07" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="discussion">
<paragraph id="013236BA7607FFE64F81DFE7F24CFE23" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,429]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="33F9EAA87607FFE74F81DFE7F27BFA4F" bold="true" box="[199,315,1447,1473]" pageId="4" pageNumber="5">Remarks.</emphasis>
We believe that
<taxonomicName id="C68D4D397607FFE74EA9DFE7F12AFA4F" authorityName="Monne" authorityYear="1977" box="[495,618,1447,1473]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74EA9DFE7F12AFA4F" box="[495,618,1447,1473]" italics="true" pageId="4" pageNumber="5">Nealcidion</emphasis>
</taxonomicName>
is not a natural group because it brings together species with very different features (
<bibRefCitation id="651C4B4B7607FFE74E46DF8BF2C3FA6B" author="Bates, H. W." box="[256,387,1483,1509]" pageId="4" pageNumber="5" pagination="100 - 109" refId="ref9078" refString="Bates, H. W. (1863) Contributions to an insect fauna of the Amazon Valley. Coleoptera: Longicornes The Annals and Magazine of Natural History, Series 3, 12 (68), 100-109. https://doi.org/10.1080/00222936308681491" type="journal article" year="1863">Bates 1863</bibRefCitation>
;
<bibRefCitation id="651C4B4B7607FFE74ED6DF8BF163FA6B" author="Monne, M. A." box="[400,547,1483,1509]" pageId="4" pageNumber="5" pagination="693 - 711" refId="ref9831" refString="Monne, M. A. (1977) Contribuicao ao conhecimento dosAcanthocinini. VI (Coleoptera, Cerambycidae, Lamiinae, Acanthocinini). Revista Brasileira de Biologia, 37 (4), 693-711." type="journal article" year="1977">Monné 1977</bibRefCitation>
). However, as these differences are mixed among species, it is not possible, at the moment, to know which of them could actually represent generic and not specific differences.
<bibRefCitation id="651C4B4B7607FFE74BBCDFAFF6DCF987" author="Monne, M. A." box="[1274,1436,1519,1545]" pageId="4" pageNumber="5" pagination="321 - 326" refId="ref9870" refString="Monne, M. A. (2009) Novos taxons neotropicais em Acanthocinini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 53 (3), 321-326. https://doi.org/10.1590/S0085-56262009000300001" type="journal article" year="2009">Monné (2009)</bibRefCitation>
described
<taxonomicName id="C68D4D397607FFE74E4DDC53F2EAF9A3" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[267,426,1555,1581]" class="Insecta" family="Cerambycidae" genus="Tropidocoleus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74E4DDC53F2EAF9A3" box="[267,426,1555,1581]" italics="true" pageId="4" pageNumber="5">Tropidocoleus</emphasis>
</taxonomicName>
to include
<taxonomicName id="C68D4D397607FFE74D6DDC53F026F9A3" authority="Monne, 2009" authorityName="Monne" authorityYear="2009" box="[555,870,1555,1581]" class="Insecta" family="Cerambycidae" genus="Tropidocoleus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bicarinatus">
<emphasis id="33F9EAA87607FFE74D6DDC53F187F9A3" box="[555,711,1555,1581]" italics="true" pageId="4" pageNumber="5">T. bicarinatus</emphasis>
<bibRefCitation id="651C4B4B7607FFE74D89DC53F026F9A3" author="Monne, M. A." box="[719,870,1555,1581]" pageId="4" pageNumber="5" pagination="321 - 326" refId="ref9870" refString="Monne, M. A. (2009) Novos taxons neotropicais em Acanthocinini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 53 (3), 321-326. https://doi.org/10.1590/S0085-56262009000300001" type="journal article" year="2009">Monné, 2009</bibRefCitation>
</taxonomicName>
and commented (translated): “
<taxonomicName id="C68D4D397607FFE74B83DC53F624F9A3" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[1221,1380,1555,1581]" class="Insecta" family="Cerambycidae" genus="Tropidocoleus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="33F9EAA87607FFE74B83DC53F624F9A3" box="[1221,1380,1555,1581]" italics="true" pageId="4" pageNumber="5">Tropidocoleus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28CA57D37607FFE74A2BDC53F387F9DF" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
belongs to the group of genera with elytra without erect setae, with centrobasal crest, without lateral [humeral] carinae, and with dorsal carinae. This group includes
<taxonomicName id="C68D4D397607FFE74DB2DC1BF750F9FB" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" box="[756,1040,1627,1653]" class="Insecta" family="Cerambycidae" genus="Cobelura" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74DB2DC1BF01EF9FB" box="[756,862,1627,1653]" italics="true" pageId="4" pageNumber="5">Cobelura</emphasis>
Erichson, 1847
</taxonomicName>
and the Group I (
<bibRefCitation id="651C4B4B7607FFE74B9CDC1BF6D4F9FB" author="Bates, H. W." box="[1242,1428,1626,1653]" pageId="4" pageNumber="5" pagination="100 - 109" refId="ref9078" refString="Bates, H. W. (1863) Contributions to an insect fauna of the Amazon Valley. Coleoptera: Longicornes The Annals and Magazine of Natural History, Series 3, 12 (68), 100-109. https://doi.org/10.1080/00222936308681491" type="journal article" year="1863">Bates 1863: 105</bibRefCitation>
) of
<taxonomicName id="C68D4D397607FFE74FF1DC3FF295F917" authority="Monne, 1977" authorityName="Monne" authorityYear="1977" box="[183,469,1663,1689]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74FF1DC3FF272F917" box="[183,306,1663,1689]" italics="true" pageId="4" pageNumber="5">Nealcidion</emphasis>
<bibRefCitation id="651C4B4B7607FFE74E7DDC3FF295F917" author="Monne, M. A." box="[315,469,1663,1689]" pageId="4" pageNumber="5" pagination="693 - 711" refId="ref9831" refString="Monne, M. A. (1977) Contribuicao ao conhecimento dosAcanthocinini. VI (Coleoptera, Cerambycidae, Lamiinae, Acanthocinini). Revista Brasileira de Biologia, 37 (4), 693-711." type="journal article" year="1977">Monné, 1977</bibRefCitation>
</taxonomicName>
, with dorsal carinae on elytra, from the centrobasal crest to the outer apical angle of elytra.” Additionally,
<bibRefCitation id="651C4B4B7607FFE74ECEDCE3F168F933" author="Monne, M. A." box="[392,552,1699,1725]" pageId="4" pageNumber="5" pagination="321 - 326" refId="ref9870" refString="Monne, M. A. (2009) Novos taxons neotropicais em Acanthocinini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 53 (3), 321-326. https://doi.org/10.1590/S0085-56262009000300001" type="journal article" year="2009">Monné (2009)</bibRefCitation>
commented that in
<taxonomicName id="C68D4D397607FFE74C45DCE3F0E2F933" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[771,930,1699,1725]" class="Insecta" family="Cerambycidae" genus="Tropidocoleus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74C45DCE3F0E2F933" box="[771,930,1699,1725]" italics="true" pageId="4" pageNumber="5">Tropidocoleus</emphasis>
</taxonomicName>
,
<taxonomicName id="C68D4D397607FFE74CF6DCE3F75AF933" authorityName="Erichson" authorityYear="1847" box="[944,1050,1699,1725]" class="Insecta" family="Cerambycidae" genus="Cobelura" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74CF6DCE3F75AF933" box="[944,1050,1699,1725]" italics="true" pageId="4" pageNumber="5">Cobelura</emphasis>
</taxonomicName>
, and the Group I of
<taxonomicName id="C68D4D397607FFE74BBFDCE3F634F933" authorityName="Monne" authorityYear="1977" box="[1273,1396,1699,1725]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74BBFDCE3F634F933" box="[1273,1396,1699,1725]" italics="true" pageId="4" pageNumber="5">Nealcidion</emphasis>
</taxonomicName>
the sides of the prothorax are either rounded or with a poorly developed swelling, and have one dorsal carina on each elytron. However, there are species in
<taxonomicName id="C68D4D397607FFE74D05DCABF1FEF88B" authorityName="Monne" authorityYear="1977" box="[579,702,1771,1797]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74D05DCABF1FEF88B" box="[579,702,1771,1797]" italics="true" pageId="4" pageNumber="5">Nealcidion</emphasis>
</taxonomicName>
with humeral carinae well marked, at least keel-shaped on part of the elytra (
<emphasis id="33F9EAA87607FFE74E4BDD4FF26FF8A6" box="[269,303,1807,1832]" italics="true" pageId="4" pageNumber="5">e.g</emphasis>
.
<taxonomicName id="C68D4D397607FFE74E7DDD4FF0C3F8A7" authority="Nascimento, Santos-Silva &amp; McClarin, 2020" authorityName="Nascimento, Santos-Silva &amp; McClarin" authorityYear="2020" box="[315,899,1807,1833]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kayi">
<emphasis id="33F9EAA87607FFE74E7DDD4FF2C9F8A7" box="[315,393,1807,1833]" italics="true" pageId="4" pageNumber="5">N. kayi</emphasis>
Nascimento, Santos-Silva &amp; McClarin, 2020
</taxonomicName>
;
<taxonomicName id="C68D4D397607FFE74CCADD4FF7FFF8A7" authority="(Bates, 1863)" baseAuthorityName="Bates" baseAuthorityYear="1863" box="[908,1215,1807,1833]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="trivittatum">
<emphasis id="33F9EAA87607FFE74CCADD4FF761F8A6" box="[908,1057,1807,1832]" italics="true" pageId="4" pageNumber="5">N. trivittatum</emphasis>
(
<bibRefCitation id="651C4B4B7607FFE74B69DD4FF7F7F8A7" author="Bates, H. W." box="[1071,1207,1807,1833]" pageId="4" pageNumber="5" pagination="100 - 109" refId="ref9078" refString="Bates, H. W. (1863) Contributions to an insect fauna of the Amazon Valley. Coleoptera: Longicornes The Annals and Magazine of Natural History, Series 3, 12 (68), 100-109. https://doi.org/10.1080/00222936308681491" type="journal article" year="1863">Bates, 1863</bibRefCitation>
)
</taxonomicName>
; and
<taxonomicName id="C68D4D397607FFE74BB1DD4FF2A5F8C3" authority="Vlasak &amp; Santos-Silva, 2024" authorityName="Vlasak &amp; Santos-Silva" authorityYear="2024" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="11" phylum="Arthropoda" rank="species" species="apunctatum">
N.
<emphasis id="33F9EAA87607FFE74A5EDD4FF6DCF8A6" box="[1304,1436,1807,1832]" italics="true" pageId="4" pageNumber="5">apunctatum</emphasis>
<bibRefCitation id="651C4B4B7607FFE74FD1DD73F2A5F8C3" author="Vlasak, J. &amp; Santos-Silva, A." box="[151,485,1843,1869]" pageId="4" pageNumber="5" pagination="80 - 90" refId="ref10258" refString="Vlasak, J. &amp; Santos-Silva, A. (2024) Ecuadorian Acanthocinini: description of two new species and transference and notes on Sympagus cooperi Monne &amp; Monne. Zootaxa, 5397 (1), 80-90. https://doi.org/10.11646/zootaxa.5397.1.4" type="journal article" year="2024">Vlasak &amp; Santos-Silva, 2024</bibRefCitation>
</taxonomicName>
). Furthermore, there are species in
<taxonomicName id="C68D4D397607FFE74C33DD73F0B0F8C3" authorityName="Monne" authorityYear="1977" box="[885,1008,1843,1869]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74C33DD73F0B0F8C3" box="[885,1008,1843,1869]" italics="true" pageId="4" pageNumber="5">Nealcidion</emphasis>
</taxonomicName>
with two or more dorsal carinae (
<emphasis id="33F9EAA87607FFE74A32DD73F6DCF8C2" box="[1396,1436,1843,1868]" italics="true" pageId="4" pageNumber="5">e.g.</emphasis>
<taxonomicName id="C68D4D397607FFE74FD1DD17F2B0F8FF" authority="(Aurivillius, 1900)" baseAuthorityName="Aurivillius" baseAuthorityYear="1900" box="[151,496,1879,1905]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="griseum">
<emphasis id="33F9EAA87607FFE74FD1DD17F252F8FE" box="[151,274,1879,1904]" italics="true" pageId="4" pageNumber="5">N. griseum</emphasis>
(Aurivillius, 1900)
</taxonomicName>
;
<taxonomicName id="C68D4D397607FFE74EBADD17F06EF8FF" authority="(Bates, 1881)" baseAuthorityName="Bates" baseAuthorityYear="1881" box="[508,814,1879,1905]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="eulophum">
<emphasis id="33F9EAA87607FFE74EBADD17F1CBF8FF" box="[508,651,1879,1905]" italics="true" pageId="4" pageNumber="5">N. eulophum</emphasis>
(Bates, 1881)
</taxonomicName>
; and the shape of the sides of the prothorax is variable, uniformly rounded from base to near apex, with rounded central protuberance, or somewhat distinctly projected and subconical.
<bibRefCitation id="651C4B4B7607FFE74E5BDDDFF2FDF837" author="Monne, M. A." box="[285,445,1951,1977]" pageId="4" pageNumber="5" pagination="321 - 326" refId="ref9870" refString="Monne, M. A. (2009) Novos taxons neotropicais em Acanthocinini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 53 (3), 321-326. https://doi.org/10.1590/S0085-56262009000300001" type="journal article" year="2009">Monné (2009)</bibRefCitation>
also defined
<taxonomicName id="C68D4D397607FFE74D14DDDFF1B1F837" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[594,753,1951,1977]" class="Insecta" family="Cerambycidae" genus="Tropidocoleus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87607FFE74D14DDDFF1B1F837" box="[594,753,1951,1977]" italics="true" pageId="4" pageNumber="5">Tropidocoleus</emphasis>
</taxonomicName>
as having narrow upper eye lobes, with the distance between them about five times the diameter of one lobe, and the lower eye lobes about as long as the genae. Although the length of the lower eye lobes in
<taxonomicName id="C68D4D397606FFE64EBFDAD7F134FF3F" authorityName="Monne" authorityYear="1977" box="[505,628,151,177]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87606FFE64EBFDAD7F134FF3F" box="[505,628,151,177]" italics="true" pageId="5" pageNumber="6">Nealcidion</emphasis>
</taxonomicName>
is often much longer than the genae, there are species with the lower eye lobes as long, shorter, or distinctly shorter than the genae. Finally,
<bibRefCitation id="651C4B4B7606FFE64C3DDAFBF75BFF5B" author="Monne, M. A." box="[891,1051,187,213]" pageId="5" pageNumber="6" pagination="321 - 326" refId="ref9870" refString="Monne, M. A. (2009) Novos taxons neotropicais em Acanthocinini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 53 (3), 321-326. https://doi.org/10.1590/S0085-56262009000300001" type="journal article" year="2009">Monné (2009)</bibRefCitation>
affirmed that the centrobasal crest of the elytra in
<taxonomicName id="C68D4D397606FFE64E04DA9FF2A1FF77" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[322,481,223,249]" class="Insecta" family="Cerambycidae" genus="Tropidocoleus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87606FFE64E04DA9FF2A1FF77" box="[322,481,223,249]" italics="true" pageId="5" pageNumber="6">Tropidocoleus</emphasis>
</taxonomicName>
is prominent and without erect setae [apically]. However, the centrobasal crest in the species of
<taxonomicName id="C68D4D397606FFE64E49DB43F2CAFE93" authorityName="Monne" authorityYear="1977" box="[271,394,259,285]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87606FFE64E49DB43F2CAFE93" box="[271,394,259,285]" italics="true" pageId="5" pageNumber="6">Nealcidion</emphasis>
</taxonomicName>
is extremely variable, from slightly elevated to distinctly elevated, from short to moderately long, and with or without erect setae on top. Therefore, the differences pointed out separating
<taxonomicName id="C68D4D397606FFE64BF9DB67F61EFECF" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[1215,1374,295,321]" class="Insecta" family="Cerambycidae" genus="Tropidocoleus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87606FFE64BF9DB67F61EFECF" box="[1215,1374,295,321]" italics="true" pageId="5" pageNumber="6">Tropidocoleus</emphasis>
</taxonomicName>
from
<taxonomicName id="C68D4D397606FFE64FD1DB0BF252FEEB" authorityName="Monne" authorityYear="1977" box="[151,274,331,357]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87606FFE64FD1DB0BF252FEEB" box="[151,274,331,357]" italics="true" pageId="5" pageNumber="6">Nealcidion</emphasis>
</taxonomicName>
are problematic. We are pointing out the problems regarding
<taxonomicName id="C68D4D397606FFE64CF0DB0BF715FEEB" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[950,1109,331,357]" class="Insecta" family="Cerambycidae" genus="Tropidocoleus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA87606FFE64CF0DB0BF715FEEB" box="[950,1109,331,357]" italics="true" pageId="5" pageNumber="6">Tropidocoleus</emphasis>
</taxonomicName>
here because the new species presents some features that allow it to be included in this genus, especially the shape of the lateral tubercles of the prothorax.
</paragraph>
<caption id="55F266327606FFE64FDEDD23F60CF811" ID-DOI="http://doi.org/10.5281/zenodo.15012346" ID-Zenodo-Dep="15012346" httpUri="https://zenodo.org/record/15012346/files/figure.png" pageId="5" pageNumber="6" startId="5.[152,270,1891,1915]" targetBox="[162,1423,462,1863]" targetPageId="5" targetType="figure">
<paragraph id="013236BA7606FFE64FDEDD23F60CF811" blockId="5.[151,1436,1891,1951]" pageId="5" pageNumber="6">
<emphasis id="33F9EAA87606FFE64FDEDD23F211F8F5" bold="true" box="[152,337,1891,1915]" pageId="5" pageNumber="6">FIGURES 610.</emphasis>
<taxonomicName id="C68D4D397606FFE64E1CDD23F126F8F5" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[346,614,1891,1915]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chloroxanthon" status="sp. nov.">
<emphasis id="33F9EAA87606FFE64E1CDD23F126F8F5" box="[346,614,1891,1915]" italics="true" pageId="5" pageNumber="6">Nealcidion chloroxanthon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28CA57D37606FFE64D29DD24F184F8F5" box="[623,708,1892,1915]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, paratypes from Ecuador (Napo), dorsal habitus:
<emphasis id="33F9EAA87606FFE64B8BDD23F7A2F8F5" bold="true" box="[1229,1250,1891,1915]" pageId="5" pageNumber="6">6)</emphasis>
Male from Pacto Sumaco;
<emphasis id="33F9EAA87606FFE64FB0DDC7F24CF811" bold="true" box="[246,268,1927,1951]" pageId="5" pageNumber="6">7)</emphasis>
Female from Papallacta;
<emphasis id="33F9EAA87606FFE64D56DDC7F165F811" bold="true" box="[528,549,1927,1951]" pageId="5" pageNumber="6">8)</emphasis>
Male from Papallacta;
<emphasis id="33F9EAA87606FFE64C54DDC7F068F811" bold="true" box="[786,808,1927,1951]" pageId="5" pageNumber="6">9)</emphasis>
Female from Papallacta;
<emphasis id="33F9EAA87606FFE64B6ADDC7F70EF811" bold="true" box="[1068,1102,1927,1951]" pageId="5" pageNumber="6">10)</emphasis>
Female from Papallacta.
</paragraph>
</caption>
<paragraph id="013236BA7605FFE54F81DAD7F1CAFE07" blockId="6.[151,1436,151,393]" pageId="6" pageNumber="7">
<taxonomicName id="C68D4D397605FFE54F81DAD7F2ACFF3F" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[199,492,151,177]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="chloroxanthon">
<emphasis id="33F9EAA87605FFE54F81DAD7F2ACFF3F" box="[199,492,151,177]" italics="true" pageId="6" pageNumber="7">Nealcidion chloroxanthon</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="C68D4D397605FFE54DC7DAD7F71BFF3F" authority="Vlasak &amp; Santos-Silva, 2022" authorityName="Vlasak &amp; Santos-Silva" authorityYear="2022" box="[641,1115,151,177]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obliquum">
<emphasis id="33F9EAA87605FFE54DC7DAD7F04CFF3F" box="[641,780,151,177]" italics="true" pageId="6" pageNumber="7">N. obliquum</emphasis>
<bibRefCitation id="651C4B4B7605FFE54C55DAD7F71BFF3F" author="Vlasak, J. &amp; Santos-Silva, A." box="[787,1115,151,177]" pageId="6" pageNumber="7" pagination="535 - 554" refId="ref10164" refString="Vlasak, J. &amp; Santos-Silva, A. (2022) Five new species of Nealcidion Monne, 1997 from Ecuador (Coleoptera, Cerambycidae, Lamiinae). Zootaxa, 5209 (5), 535-554. https://doi.org/10.11646/zootaxa.5209.5.2" type="journal article" year="2022">Vlasak &amp; Santos-Silva, 2022</bibRefCitation>
</taxonomicName>
(see photographs on
<bibRefCitation id="651C4B4B7605FFE54A0BDAD7F3A2FF5B" author="Bezark, L. G." pageId="6" pageNumber="7" refId="ref9130" refString="Bezark, L. G. (2025) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http://bezbycids.com/byciddb/wdefault.asp?w=n/ (accessed 6 January 2025)" type="url" year="2025">Bezark (2025)</bibRefCitation>
and
<bibRefCitation id="651C4B4B7605FFE54E5CDAFBF139FF5B" author="Vlasak, J. &amp; Santos-Silva, A." box="[282,633,187,213]" pageId="6" pageNumber="7" pagination="535 - 554" refId="ref10164" refString="Vlasak, J. &amp; Santos-Silva, A. (2022) Five new species of Nealcidion Monne, 1997 from Ecuador (Coleoptera, Cerambycidae, Lamiinae). Zootaxa, 5209 (5), 535-554. https://doi.org/10.11646/zootaxa.5209.5.2" type="journal article" year="2022">Vlasak &amp; Santos-Silva (2022))</bibRefCitation>
, but differs especially as follows: general pubescence mostly greenish-yellow; pronotum with wide, longitudinal greenish-yellow pubescence centrally and without longitudinal dark pubescent bands on sides of central region; and different pubescent pattern. In
<taxonomicName id="C68D4D397605FFE54B44DB43F7CCFE93" authorityName="Vlasak &amp; Santos-Silva" authorityYear="2022" box="[1026,1164,259,285]" class="Insecta" family="Cerambycidae" genus="Nealcidion" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obliquum">
<emphasis id="33F9EAA87605FFE54B44DB43F7CCFE93" box="[1026,1164,259,285]" italics="true" pageId="6" pageNumber="7">N. obliquum</emphasis>
</taxonomicName>
, the general pubescence is not greenish-yellow, central area of pronotum longitudinally glabrous centrally, with moderately narrow, longitudinal light pubescent band on sides of glabrous area, and large, longitudinal dark pubescent area close to central area, and different pubescent pattern.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="CDB4C0136E3A5BE48BE1930266867545">New Ecuadorian and Colombian Lamiinae (Coleoptera, Cerambycidae)</mods:title>
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<mods:name id="77648FBAD41C26423D0A954289906C6E" type="personal">
<mods:role id="021FE87CE62C115A51C9EB46224C150C">
<mods:roleTerm id="BA942987BB7827461BD693662BB18582">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C7C45248B181F8DD5D84FB51730854B7">Vlasak, Josef</mods:namePart>
<mods:affiliation id="A21FAD193BD0654C383ED29A2602B22E">207 Silverbrook Drive, Schwenksville, PA 19473, U. S. A.</mods:affiliation>
</mods:name>
<mods:name id="39AE31E5B99C6525BA5FADC422196EEF" type="personal">
<mods:role id="BFECBC8CA29A22DE5A3848212473C4BE">
<mods:roleTerm id="2E56220CBA254BC8D446D63AFB9B1E49">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0EC9C6C1DE4E0DDBD2FFA5A7E6B0005D">Nascimento, Francisco Eriberto Lima</mods:namePart>
<mods:affiliation id="BF389740768EA7FA7EBC3A7D5FFD7949">Laboratório de Invertebrados - LABIN, Instituto de Ciências Exatas e Naturais - ICEN, Universidade Federal de Rondonopolis, Rondonópolis, Mato Grosso, Brazil</mods:affiliation>
</mods:name>
<mods:name id="665C2C997C1ECB7E948E35C8AB0428F8" type="personal">
<mods:role id="135E7408943A4F24EB1E93ACA0760971">
<mods:roleTerm id="0E986964BACBDCC05CDD3B1C5CA49CBD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="35FD52D28A435447E71E6C5CA7610A6A">Santos-Silva, Antonio</mods:namePart>
<mods:affiliation id="5BB9738243A198096DE556A7265E0428">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="02135A9057ED573FFC8800ABAD7CFE3A">text</mods:typeOfResource>
<mods:relatedItem id="7BBECA395A9B35388B65BAC94AA465AA" type="host">
<mods:titleInfo id="6BFFB30AF2B35CD04B4D853EFD0A0393">
<mods:title id="DE6DE5C096A128FC376F2F9B6206A9DD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B22AEDE49D030C8D36CF7A4203DB9B39">
<mods:date id="E0D9FE7E72F54C8466B22DC4610DDD53">2025</mods:date>
<mods:detail id="3DAFF9519693718F3F6FEF553538C970" type="pubDate">
<mods:number id="A56A607C4ABE8E85259494BD0671BCF4">2025-03-12</mods:number>
</mods:detail>
<mods:detail id="3C037183D7609B04A164DD697CAFC08C" type="volume">
<mods:number id="416BC0098FB42A88E701C19292C059F3">5604</mods:number>
</mods:detail>
<mods:detail id="3AA765CBC651B98E91D4FB9942CA818E" type="issue">
<mods:number id="A91B44994CFCA3A176090B0BEA60E256">1</mods:number>
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<mods:start id="03453ECE4718FF61C340E85482FADB0F">1</mods:start>
<mods:end id="8B54F027661D81BD99CB1D2CBF6094DA">15</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4D56BEBE4A08FE3306DC4711BD4658B0">
<mods:url id="E1FD8A70D20213EC79D81E69B0935124">https://doi.org/10.11646/zootaxa.5604.1.1</mods:url>
</mods:location>
<mods:classification id="78FE7F6976894DFFF0A1D5EE8C3D03AC">journal article</mods:classification>
<mods:identifier id="97AF2D6840E460D0577621C92C7CF719" type="DOI">10.11646/zootaxa.5604.1.1</mods:identifier>
<mods:identifier id="6C50DC619F77F421DE8848978AAAF52E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F5FAC477C52677FA522DFB0F0B31ED10" type="Zenodo-Dep">15012338</mods:identifier>
<mods:identifier id="8D64C6AA08596BC0532EDA32E02CB989" type="ZooBank">2C4691AC-E712-4335-8869-6B9547DF5D7A</mods:identifier>
</mods:mods>
<treatment id="892487AC7609FFEF4FD1DE5AF06BF837" LSID="urn:lsid:plazi:treatment:892487AC7609FFEF4FD1DE5AF06BF837" httpUri="http://treatment.plazi.org/id/892487AC7609FFEF4FD1DE5AF06BF837" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="499765317609FFE94FD1DE5AF28CFBBB" box="[151,460,1050,1077]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="013236BA7609FFE94FD1DE5AF28CFBBB" blockId="10.[151,460,1050,1113]" box="[151,460,1050,1077]" pageId="10" pageNumber="11">
<heading id="5A7A81D67609FFE94FD1DE5AF28CFBBB" bold="true" box="[151,460,1050,1077]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="33F9EAA87609FFE94FD1DE5AF28CFBBB" bold="true" box="[151,460,1050,1077]" pageId="10" pageNumber="11">
<taxonomicName id="C68D4D397609FFE94FD1DE5AF227FBBB" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[151,359,1050,1077]" class="Insecta" family="Cerambycidae" genus="Antodice" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="apicalis" status="sp. nov.">
<emphasis id="33F9EAA87609FFE94FD1DE5AF227FBBB" bold="true" box="[151,359,1050,1077]" italics="true" pageId="10" pageNumber="11">Antodice apicalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28CA57D37609FFE94E28DE5BF28CFBBB" box="[366,460,1051,1077]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="499765317609FFEF4FD1DE7FF643FA83" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
<paragraph id="013236BA7609FFE94FD1DE7FF267FBD7" blockId="10.[151,460,1050,1113]" box="[151,295,1087,1113]" pageId="10" pageNumber="11">
(
<figureCitation id="99B62A3F7609FFE94FD9DE7FF25FFBD7" box="[159,287,1087,1113]" captionStart="FIGURES 1721" captionStartId="13.[152,270,1731,1755]" captionTargetBox="[151,1435,181,1708]" captionTargetId="figure-19@13.[151,1435,181,1708]" captionTargetPageId="13" captionText="FIGURES 1721. Antodice apicalis sp. nov., holotype female: 17) Dorsal habitus; 18) Ventral habitus; 19) Lateral habitus; 20) Head, frontal view; 21) Tarsal claw." figureDoi="http://doi.org/10.5281/zenodo.15012350" httpUri="https://zenodo.org/record/15012350/files/figure.png" pageId="10" pageNumber="11">Figs 1721</figureCitation>
)
</paragraph>
<paragraph id="013236BA7609FFE94FD1DEC6F7D3F9A3" blockId="10.[151,1437,1158,2013]" pageId="10" pageNumber="11">
<emphasis id="33F9EAA87609FFE94FD1DEC6F2B5FB2E" bold="true" box="[151,501,1158,1185]" pageId="10" pageNumber="11">
Description.
<typeStatus id="DE3688187609FFE94E6BDEC7F2DAFB2F" box="[301,410,1159,1185]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
female.
</emphasis>
Head capsule blackish; ventral mouthparts mostly orangish, except some palpomeres partially brown or reddish brown and with yellowish-brown apex; anteclypeus mostly dark brown centrally, irregularly dull yellowish-brown laterally; labrum reddish brown with irregular dark-brown areas; antennae blackish, except scape, pedicel and base of antennomeres IIIVII dark reddish brown; mandibles black. Prothorax blackish on pronotum and sides, dark brown ventrally, except posterior half of prosternal process orangish brown with irregular dark-brown areas interspersed. Ventral surface of meso- and metathorax black. Scutellum black. Elytra orangish brown on anterior third, gradually more brownish toward posterior region, except black inverted psi-shaped macula on anterior 2/3 and brown posterior eight. Coxae mostly dark orangish brown, except brown sides of metacoxae. Protrochanters dark brown; meso- and metatrochanters dark reddish brown. Femora black except dark reddish-brown basal area of dorsal and lateral surfaces of profemora and light reddish-brown basal area of dorsal and lateral surfaces of meso- and metafemora. Tibiae and tarsi black, except reddish-brown apex of tarsomeres V and tarsal claws. Ventral surface of abdomen dark brown, except brown central apex of ventrites 14.
</paragraph>
<paragraph id="013236BA7609FFE84F81DC77F23DFB9F" blockId="10.[151,1437,1158,2013]" lastBlockId="11.[151,1437,151,2013]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="33F9EAA87609FFE94F81DC77F24EF9DF" bold="true" box="[199,270,1591,1617]" pageId="10" pageNumber="11">Head.</emphasis>
Frons abundantly, finely punctate, with moderately coarse punctures interspersed; with abundant yellowish-brown pubescence not obscuring integument laterally and superiorly, and a few yellowish-brown setae on remaining surface; with a few long, erect dark-brown setae near eyes. Area between eyes and base of antennae with dense yellowish-brown pubescence dorsally and laterally, both dark-brown and yellowish-brown frontally. Area between antennal tubercles and anterior margin of upper eye lobes, abundantly, finely punctate, with coarse punctures interspersed, except smooth median groove, smooth area widened on its central area; with abundant yellowish-brown pubescence partially obscuring integument close to glabrous median groove and abundant dark-brown pubescence not obscuring integument laterally. Remaining surface of vertex abundantly, finely punctate, with coarse punctures interspersed; with abundant dark-brown pubescence partially obscuring integument, with short, decumbent dark yellowish-brown setae interspersed, except glabrous median groove; with a few long, erect dark-brown setae interspersed. Area behind upper eye lobes with sculpturing as on posterior region of vertex; with abundant dark-brown pubescence close to vertex and eye, and dense yellow pubescence on remaining surface; with a few long, erect dark-brown setae close to eye. Area behind lower eye lobes with sculpturing as on area behind upper eye lobes; with abundant, both yellowish-brown and yellowish-white pubescence not obscuring integument, pubescence distinctly sparser close to eye; with a few long, erect dark-brown setae interspersed. Genae 0.7 times longer than inner margin of lower eye lobe; abundantly, finely striate-punctate laterally; frontal area smooth close to eyes and on apex, sparsely, finely punctate on remaining surface; punctate frontal area with moderately abundant pale-yellow pubescence not obscuring integument, with long, erect, decumbent pale setae interspersed, striate-punctate area with moderately sparse yellowish-white pubescence and long, erect yellowish and brown setae interspersed. Antennal tubercles abundantly, finely punctate, with coarse punctures interspersed; with abundant dark-brown pubescence not obscuring integument, and both long and short, erect dark-brown setae interspersed, especially close to apex. Wide central area of postclypeus microsculptured, with fine, shallow punctures interspersed, except a few coarser punctures on its sides; almost glabrous, with long, erect yellowish-brown setae laterally. Sides of postclypeus smooth, glabrous. Labrum sparsely, coarsely punctate close to anteclypeus, smooth on remaining surface; with long, erect yellowish-brown setae posteriorly and laterally, absent on remaining surface. Gulamentum smooth, glabrous posteriorly; anterior region abundantly, finely rugose-punctate, with sparse, both short and long, erect, both brown and yellowish-white setae. Maximum diameter of upper eye lobes 3.5 times shorter than maximum diameter of scape; distance between upper eye lobes 0.14 times distance between outer margins of eyes; in frontal view, distance between lower eye lobes 0.59 times distance between outer margins of eyes. Antennae 1.15 times elytral length, reaching about posterior seventh of elytra. Scape and pedicel with abundant, short, bristly dark-brown setae, slightly denser on scape, and long, erect dark-brown setae interspersed throughout. Antennomeres IIIXI with abundant, short, bristly dark-brown setae, sparser on base of IIIVII; ventral surface with long, erect dark-brown setae, denser on IIIIV, gradually sparser from V; with sparse, long, erect dark-brown setae dorsally and laterally; with long dark-brown setae directed backward on entire apex; antennomere III wide, gradually widened from base to apex; remaining flagellomeres subcylindrical, gradually narrowed from IV. Antennal formula (ratio) based on antennomere III: scape = 1.08; pedicel = 0.21; IV = 1.17; V = 1.30; VI = 1.17; VII = 1.00; VIII = 0.76; IX = 0.69; X = 0.65; XI = 0.76.
</paragraph>
<paragraph id="013236BA7608FFEF4F81DE5BF227FBD7" blockId="11.[151,1437,151,2013]" lastBlockId="12.[151,1437,151,1977]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="33F9EAA87608FFE84F81DE5BF267FBBB" bold="true" box="[199,295,1051,1077]" pageId="11" pageNumber="12">Thorax.</emphasis>
Prothorax about as long as wide; anterior constriction distinct; sides sinuous. Pronotum densely, very finely punctate, with abundant, coarse punctures interspersed; sides with dense, wide, longitudinal orangish-yellow pubescent band, from anterior to posterior margin, almost reaching sides of prothorax; remaining surface with abundant dark-brown pubescence not obscuring integument and long, erect yellowish-white setae and a few very long, erect setae interspersed, these latter brown with pale apices. Sides of prothorax with abundant brown pubescence not obscuring integument and long, erect, dark-brown setae interspersed. Prosternum sparsely, coarsely punctate, punctures coarser and denser on anterior sulcus; with abundant brown pubescence partially obscuring integument laterally, and long, erect whitish setae interspersed; remaining surface with sparse, both, short, decumbent and long, erect whitish setae. Prosternal process with moderately abundant yellowish-white pubescence, slightly sparser on posterior half, and long, erect whitish setae interspersed on posterior half; narrowest area 0.14 times procoxal width. Mesoventrite moderately abundantly, coarsely punctate, punctures denser laterally; central area with sparse dull yellowish-brown pubescence, with long, erect yellowish-white setae interspersed; remaining surface with dense dull yellowish-brown pubescence. Mesanepisterna with abundant brown pubescence, with short, decumbent whitish setae interspersed, except dense yellowish-brown pubescence close to mesepimeron. Mesepimera with moderately abundant dark-brown pubescence not obscuring integument and long, erect setae of same color interspersed. Mesoventral process densely, moderately finely punctate; posterior half almost drop-shaped; with abundant dull yellowish-brown pubescence not obscuring integument, and long, erect yellowish setae interspersed; narrowest area 0.19 times mesocoxal width; widest posterior area 0.38 times mesocoxal width. Metanepisterna with abundant dark-brown pubescence not obscuring integument, and short, decumbent whitish setae interspersed. Metaventrite with abundant brownish pubescence not obscuring integument close to metanepisterna, with long, erect dull yellowish-brown setae interspersed; lateral area close to brownish pubescence with abundant yellowish-brown pubescence on anterior third, and long, erect setae of same color interspersed; lateral area close to brownish pubescence and metacoxal cavities with abundant yellowish pubescence not obscuring integument; posterior area with abundant yellowish pubescence close to metathoracic discrimen, and long, erect setae of same color interspersed; remaining surface with sparse yellowish pubescence, slightly more abundant anteriorly, and long, erect setae of same color interspersed, except glabrous area close to metathoracic discrimen and central area close to metacoxal cavities. Scutellum with abundant dark-brown pubescence not obscuring integument, except dense dark yellowish-brown pubescence laterally, and abundant pale pubescence posterocentrally.
<emphasis id="33F9EAA8760FFFEF4C92DAD7F767FF3F" bold="true" box="[980,1063,151,177]" pageId="12" pageNumber="13">Elytra.</emphasis>
Abundantly, coarsely punctate, punctures slightly sparser and finer on posterior quarter; dorsal surface widely, longitudinally, moderately shallowly sulcate; psi-shaped area with moderately abundant dark-brown pubescence not obscuring integument, with long, erect dark-brown setae interspersed, and a few moderately long, suberect yellowish-white setae interspersed; light integumental area on anterior half with dense orangish-yellow pubescence and long, erect yellowish setae interspersed; light integumental area on posterior half with dense orangish-yellow pubescence, except somewhat indistinct, U-shaped pale-yellow pubescent macula dorsally; area close to apex with abundant, bristly dark-brown pubescence not obscuring integument, and long, erect setae of same color interspersed; apex rounded.
<emphasis id="33F9EAA8760FFFEF4A63DBD3F624FE23" bold="true" box="[1317,1380,403,429]" pageId="12" pageNumber="13">Legs.</emphasis>
Pro- and mesocoxae subglabrous anteriorly; remaining surface with sparse pale-yellow pubescence and long, erect setae of same color interspersed. Metacoxae with abundant dull yellowish-brown pubescence antero- and posterolaterally, moderately sparse whitish pubescence on remaining lateral area, and moderately abundant pale-yellow pubescence, with long, erect setae of same color interspersed close to trochanter. Trochanters with abundant pale-yellow pubescence, pubescence almost absent anteriorly, and long, erect pale-yellow setae interspersed. Femora with moderately abundant, bristly dark-brown pubescence dorsally, with long, erect setae of same color interspersed on posterior half; with abundant yellowish pubescence on remaining surface, pubescence denser on inferior half of sides and ventrally, and long, erect yellowish setae interspersed, erect setae distinctly denser basally. Protibiae abundantly, moderately coarsely rugose-punctate, except smooth dorsal apex; dorsal and lateral surfaces with abundant dark-brown pubescence not obscuring integument and long, erect setae of same color interspersed, except glabrous smooth area; ventral surface with very sparse dark-brown pubescence on basal quarter, dense, bristly yellowish-brown pubescence from basal quarter to middle, with a few moderately long, erect dark-brown setae interspersed, and dense, bristly, mostly dark-brown pubescence on remaining surface. Meso- and metatibiae transversely striate-punctate dorsally and laterally, abundantly, finely punctate ventrally; with dark-brown pubescence dorsally and laterally, denser and bristly on some areas, except dense, bristly yellowish-brown pubescence centrally on dorsal surface, and long, erect dark-brown setae interspersed; ventral surface with abundant, bristly, mostly yellowish-brown pubescence on basal half, and bristly, gradually longer, dense dark-brown pubescence on apical half. Dorsal surface of tarsomeres with sparse dark-brown pubescence, distinctly sparser on IVV, and long, erect setae of same color interspersed.
</paragraph>
<paragraph id="013236BA760FFFEF4F81DE22F04AFB4B" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="13">
<emphasis id="33F9EAA8760FFFEF4F81DE22F203FBF2" bold="true" box="[199,323,1122,1148]" pageId="12" pageNumber="13">Abdomen.</emphasis>
Ventrites with abundant yellowish-brown pubescence not obscuring integument, pubescence appearing darker laterally depending on light source, except glabrous apex of 24, and long, erect setae of same color interspersed. Apex of ventrite 5 concave centrally.
</paragraph>
<paragraph id="013236BA760FFFEF4F81DE8EF643FA83" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="13">
<emphasis id="33F9EAA8760FFFEF4F81DE8EF2E4FB66" bold="true" box="[199,420,1230,1256]" pageId="12" pageNumber="13">Dimensions (mm).</emphasis>
Total length, 10.90; prothoracic length, 1.35; anterior prothoracic width, 1.35; posterior prothoracic width, 1.30; maximum prothoracic width, 1.35; humeral width, 2.00; elytral length, 8.55.
</paragraph>
</subSubSection>
<subSubSection id="49976531760FFFEF4F81DF57F29EFADB" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="013236BA760FFFEF4F81DF57F29EFADB" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="13">
<emphasis id="33F9EAA8760FFFEF4F81DF57F237FABE" bold="true" box="[199,375,1302,1329]" pageId="12" pageNumber="13">Type material.</emphasis>
<materialsCitation id="B1E53CE7760FFFEF4E39DF57F29EFADB" collectingDate="2023-06-18" collectionCode="MZSP" collectorName="J. Vlasak" country="Ecuador" elevation="1900" location="Cabanas Intag Colibri" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Imbabura" typeStatus="holotype">
<typeStatus id="DE368818760FFFEF4E39DF57F2A7FABF" box="[383,487,1303,1329]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
female from
<collectingCountry id="799A762A760FFFEF4DC5DF57F04DFABF" box="[643,781,1303,1329]" name="Ecuador" pageId="12" pageNumber="13">ECUADOR</collectingCountry>
,
<emphasis id="33F9EAA8760FFFEF4C5CDF57F0C8FABF" box="[794,904,1303,1329]" italics="true" pageId="12" pageNumber="13">
<collectingRegion id="C349F858760FFFEF4C5CDF57F0C8FABF" box="[794,904,1303,1329]" country="Ecuador" name="Imbabura" pageId="12" pageNumber="13">Imbabura</collectingRegion>
</emphasis>
:
<location id="04526061760FFFEF4CDFDF57F7D5FABF" LSID="urn:lsid:plazi:treatment:892487AC7609FFEF4FD1DE5AF06BF837:04526061760FFFEF4CDFDF57F7D5FABF" box="[921,1173,1303,1329]" country="Ecuador" name="Cabanas Intag Colibri" pageId="12" pageNumber="13" stateProvince="Imbabura">Cabañas Intag Colibri</location>
,
<quantity id="C6759B5F760FFFEF4BE4DF57F7BAFABF" box="[1186,1274,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="13" unit="m" value="1900.0">
<elevation id="8AA0D189760FFFEF4BE4DF57F7BAFABF" box="[1186,1274,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="13" unit="m" value="1900.0">1900 m</elevation>
</quantity>
, Intag,
<date id="7533107A760FFFEF4A17DF57F392FADB" pageId="12" pageNumber="13" value="2023-06-18">
<collectingDate id="6577E992760FFFEF4A17DF57F392FADB" pageId="12" pageNumber="13" value="2023-06-18">18 Jun 2023</collectingDate>
</date>
,
<collectorName id="AC78536C760FFFEF4F9BDF7BF204FADB" box="[221,324,1339,1365]" pageId="12" pageNumber="13">J. Vlasak</collectorName>
leg. (
<collectionCode id="679CAE7F760FFFEF4EC5DF7BF292FADB" box="[387,466,1339,1365]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13" type="Museum">MZSP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="49976531760FFFEF4F81DF1FF394FA13" pageId="12" pageNumber="13" type="etymology">
<paragraph id="013236BA760FFFEF4F81DF1FF394FA13" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="13">
<emphasis id="33F9EAA8760FFFEF4F81DF1FF20EFAF7" bold="true" box="[199,334,1375,1401]" pageId="12" pageNumber="13">Etymology.</emphasis>
The specific epithet
<taxonomicName id="C68D4D39760FFFEF4D0EDF1FF1ECFAF7" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[584,684,1375,1401]" class="Insecta" family="Cerambycidae" genus="Antodice" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="apicalis">“apicalis</taxonomicName>
” (Latin, “apex,” meaning tip or end) refers to the brown elytral apex.
</paragraph>
</subSubSection>
<subSubSection id="49976531760FFFEF4F81DFE7F06BF837" pageId="12" pageNumber="13" type="discussion">
<paragraph id="013236BA760FFFEF4F81DFE7F0F2F8C3" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="13">
<emphasis id="33F9EAA8760FFFEF4F81DFE7F27BFA4F" bold="true" box="[199,315,1447,1473]" pageId="12" pageNumber="13">Remarks.</emphasis>
<taxonomicName id="C68D4D39760FFFEF4E03DFE7F14BFA4F" authorityName="Vlasak &amp; Nascimento &amp; Santos-Silva" authorityYear="2025" box="[325,523,1447,1473]" class="Insecta" family="Cerambycidae" genus="Antodice" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="apicalis" status="sp. nov.">
<emphasis id="33F9EAA8760FFFEF4E03DFE7F14BFA4F" box="[325,523,1447,1473]" italics="true" pageId="12" pageNumber="13">Antodice apicalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28CA57D3760FFFEF4D52DFE7F12DFA4F" box="[532,621,1447,1473]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="C68D4D39760FFFEF4C4EDFE7F626FA4F" authority="Bezark, Botero &amp; Santos-Silva, 2022" authorityName="Bezark, Botero &amp; Santos-Silva" authorityYear="2022" box="[776,1382,1447,1473]" class="Insecta" family="Cerambycidae" genus="Antodice" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nascimentoi">
<emphasis id="33F9EAA8760FFFEF4C4EDFE7F0F1FA4E" box="[776,945,1447,1473]" italics="true" pageId="12" pageNumber="13">A. nascimentoi</emphasis>
<bibRefCitation id="651C4B4B760FFFEF4CFDDFE7F626FA4F" author="Bezark, L. G. &amp; Botero, J. P. &amp; Santos-Silva, A." box="[955,1382,1447,1473]" pageId="12" pageNumber="13" pagination="1 - 20" refId="ref9180" refString="Bezark, L. G., Botero, J. P. &amp; Santos-Silva, A. (2022) A new genus and seven new species of Neotropical Lamiinae (Coleoptera, Cerambycidae) with taxonomic notes. Faunitaxys, 10 (1), 1-20." type="journal article" year="2022">Bezark, Botero &amp; Santos-Silva, 2022</bibRefCitation>
</taxonomicName>
(see photographs on
<bibRefCitation id="651C4B4B760FFFEF4E0FDF8BF2A9FA6B" author="Bezark, L. G." box="[329,489,1482,1509]" pageId="12" pageNumber="13" refId="ref9130" refString="Bezark, L. G. (2025) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http://bezbycids.com/byciddb/wdefault.asp?w=n/ (accessed 6 January 2025)" type="url" year="2025">Bezark (2025)</bibRefCitation>
and in
<bibRefCitation id="651C4B4B760FFFEF4D7EDF8BF060FA6B" author="Bezark, L. G. &amp; Botero, J. P. &amp; Santos-Silva, A." box="[568,800,1483,1509]" pageId="12" pageNumber="13" pagination="1 - 20" refId="ref9180" refString="Bezark, L. G., Botero, J. P. &amp; Santos-Silva, A. (2022) A new genus and seven new species of Neotropical Lamiinae (Coleoptera, Cerambycidae) with taxonomic notes. Faunitaxys, 10 (1), 1-20." type="journal article" year="2022">
Bezark
<emphasis id="33F9EAA8760FFFEF4DCBDF8BF184FA6B" box="[653,708,1483,1509]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2022))
</bibRefCitation>
, the only other species of the genus with the antennomere III distinctly not cylindrical, but differs as follows: frons abundantly, finely punctate with coarse punctures interspersed; maximum diameter of upper eye lobes 3.5 times shorter than maximum diameter of scape; lower eye lobes 1.4 times as long as genal length; in frontal view, distance between lower eye lobes 3.5 times genal length; elytra with large, inverted letter psi-shaped black macula on anterior 2/3; posterior elytra brown on posterior eight. In
<taxonomicName id="C68D4D39760FFFEF4FFFDC3FF222F916" authorityName="Bezark, Botero &amp; Santos-Silva" authorityYear="2022" box="[185,354,1663,1689]" class="Insecta" family="Cerambycidae" genus="Antodice" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nascimentoi">
<emphasis id="33F9EAA8760FFFEF4FFFDC3FF222F916" box="[185,354,1663,1689]" italics="true" pageId="12" pageNumber="13">A. nascimentoi</emphasis>
</taxonomicName>
, the frons is very sparsely finely punctate laterally, these punctures are absent centrally, with coarse punctures interspersed, the maximum diameter of the upper eye lobes is 2.5 times shorter than the maximum diameter of the scape (although usually the width is narrower than in males in other species, the difference is always very small), the lower eye lobes are twice longer than the genal length, in frontal view, the distance between the lower eye lobes is 3.0 times the genal length, the elytra have longitudinal reddish-brown band close to suture on the anterior basal third, and the posterior half of elytra is entirely orangish.
</paragraph>
<paragraph id="013236BA760FFFEF4F81DD17F06BF837" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="13">
The non-cylindrical basal flagellomeres may suggest that this species, like
<taxonomicName id="C68D4D39760FFFEF4B50DD17F7FDF8FE" authorityName="Bezark, Botero &amp; Santos-Silva" authorityYear="2022" box="[1046,1213,1879,1905]" class="Insecta" family="Cerambycidae" genus="Antodice" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nascimentoi">
<emphasis id="33F9EAA8760FFFEF4B50DD17F7FDF8FE" box="[1046,1213,1879,1905]" italics="true" pageId="12" pageNumber="13">A. nascimentoi</emphasis>
</taxonomicName>
, does not belong to
<taxonomicName id="C68D4D39760FFFEF4FD1DD3BF3BBF81B" authorityName="Thomson" authorityYear="1864" box="[151,251,1915,1941]" class="Insecta" family="Cerambycidae" genus="Antodice" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="33F9EAA8760FFFEF4FD1DD3BF3BBF81B" box="[151,251,1915,1941]" italics="true" pageId="12" pageNumber="13">Antodice</emphasis>
</taxonomicName>
. However, the existence of species with intermediate basal flagellomeres, as for example in
<taxonomicName id="C68D4D39760FFFEF4BB3DD3BF392F837" authority="Lane, 1940" authorityName="Lane" authorityYear="1940" class="Insecta" family="Cerambycidae" genus="Antodice" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="neivai">
<emphasis id="33F9EAA8760FFFEF4BB3DD3BF617F81A" box="[1269,1367,1915,1941]" italics="true" pageId="12" pageNumber="13">A. neivai</emphasis>
Lane, 1940
</taxonomicName>
, does not allow them to be separated from this genus.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,238 @@
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<mods:title id="E78FE47C34A2C3F26FE54AFC8C50DF0F">Dr. Wolfram Meys Stonefly (Insecta: Plecoptera) Collection from Luzon, Philippines: Part II. Leuctridae</mods:title>
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<treatment id="B44A87E57C12FFA5FF069CC7FE2EFCDF" ID-DOI="http://doi.org/10.5281/zenodo.15012569" ID-Zenodo-Dep="15012569" LSID="urn:lsid:plazi:treatment:B44A87E57C12FFA5FF069CC7FE2EFCDF" httpUri="http://treatment.plazi.org/id/B44A87E57C12FFA5FF069CC7FE2EFCDF" lastPageId="6" lastPageNumber="58" pageId="5" pageNumber="57">
<subSubSection id="74F965787C12FFA6FF069CC7FD70F954" box="[151,656,1724,1751]" pageId="5" pageNumber="57" type="nomenclature">
<paragraph id="3C5C36F37C12FFA6FF069CC7FD70F954" blockId="5.[151,656,1724,1787]" box="[151,656,1724,1751]" pageId="5" pageNumber="57">
<heading id="6714819F7C12FFA6FF069CC7FD70F954" bold="true" box="[151,656,1724,1751]" fontSize="11" level="1" pageId="5" pageNumber="57" reason="1">
<emphasis id="0E97EAE17C12FFA6FF069CC7FD70F954" bold="true" box="[151,656,1724,1751]" pageId="5" pageNumber="57">
<taxonomicName id="FBE34D707C12FFA6FF069CC7FDC5F954" authority="Pelingen" authorityName="Pelingen" authorityYear="2025" box="[151,549,1724,1751]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="57" phylum="Arthropoda" rank="species" species="rotundus" status="sp. nov.">
<emphasis id="0E97EAE17C12FFA6FF069CC7FE50F954" bold="true" box="[151,432,1724,1751]" italics="true" pageId="5" pageNumber="57">Rhopalopsole rotundus</emphasis>
Pelingen
</taxonomicName>
,
<taxonomicNameLabel id="15A4579A7C12FFA6FDA39CC6FD70F954" box="[562,656,1725,1751]" pageId="5" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="74F965787C12FFA6FF069C9AFE6DF978" box="[151,397,1761,1787]" pageId="5" pageNumber="57" type="description">
<paragraph id="3C5C36F37C12FFA6FF069C9AFE6DF978" blockId="5.[151,656,1724,1787]" box="[151,397,1761,1787]" pageId="5" pageNumber="57">
(
<figureCitation id="A4D82A767C12FFA6FF0E9C9AFF10F978" box="[159,240,1761,1787]" captionStart="FIGURE 2" captionStartId="2.[151,250,1099,1123]" captionTargetBox="[314,1274,179,1068]" captionTargetId="figure-15@2.[314,1274,179,1073]" captionTargetPageId="2" captionText="FIGURE 2. Male adult dorsal habitus. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012547" httpUri="https://zenodo.org/record/15012547/files/figure.png" pageId="5" pageNumber="57">Fig. 2c</figureCitation>
,
<figureCitation id="A4D82A767C12FFA6FF6D9C9AFEF9F978" box="[252,281,1761,1787]" captionStart="FIGURE 3" captionStartId="3.[151,250,1346,1370]" captionTargetBox="[314,1274,234,1319]" captionTargetId="figure-59@3.[314,1274,234,1319]" captionTargetPageId="3" captionText="FIGURE 3. Male adult lateral habitus. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012549" httpUri="https://zenodo.org/record/15012549/files/figure.png" pageId="5" pageNumber="57">3c</figureCitation>
,
<figureCitation id="A4D82A767C12FFA6FEB49C9AFED5F978" box="[293,309,1761,1787]" captionStart="FIGURE 6" captionStartId="5.[151,250,812,836]" captionTargetBox="[159,1436,179,772]" captionTargetId="figure-25@5.[151,1436,179,786]" captionTargetPageId="5" captionText="FIGURE 6. Rhopalopsole rotundus sp. nov. male terminalia. a dorsal; b lateral; c ventral." figureDoi="http://doi.org/10.5281/zenodo.15012557" httpUri="https://zenodo.org/record/15012557/files/figure.png" pageId="5" pageNumber="57">6</figureCitation>
,
<figureCitation id="A4D82A767C12FFA6FED09C9AFEBEF978" box="[321,350,1761,1787]" captionStart="FIGURE 7" captionStartId="5.[151,250,1546,1570]" captionTargetBox="[157,1426,900,1509]" captionTargetId="figure-42@5.[151,1436,894,1521]" captionTargetPageId="5" captionText="FIGURE 7. Male terminalia. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012559" httpUri="https://zenodo.org/record/15012559/files/figure.png" pageId="5" pageNumber="57">7c</figureCitation>
,
<figureCitation id="A4D82A767C12FFA6FEF89C9AFE64F978" box="[361,388,1761,1787]" captionStart="FIGURE 8" captionStartId="6.[151,250,1315,1339]" captionTargetBox="[159,1434,919,1283]" captionTargetId="figure-320@6.[151,1436,918,1289]" captionTargetPageId="6" captionText="FIGURE 8. Epiproct and tergum 10 lateral process. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012561" httpUri="https://zenodo.org/record/15012561/files/figure.png" pageId="5" pageNumber="57">8c</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="74F965787C12FFA6FF069D52FB76F8E4" pageId="5" pageNumber="57" type="materials_examined">
<paragraph id="3C5C36F37C12FFA6FF069D52FB76F8E4" blockId="5.[151,1436,1833,1967]" pageId="5" pageNumber="57">
<materialsCitation id="8C8B3CAE7C12FFA6FF069D52FBD8F8C0" box="[151,1080,1833,1859]" collectingDate="1997-04-09" collectionCode="PNM" collectorName="Mey" country="Philippines" location="Infanta" municipality="Luzon" pageId="5" pageNumber="57" specimenCount="1" specimenCount-male="1" stateProvince="Quezon" typeStatus="holotype">
<emphasis id="0E97EAE17C12FFA6FF069D52FEEDF8C0" bold="true" box="[151,269,1833,1859]" pageId="5" pageNumber="57">
<typeStatus id="E35888517C12FFA6FF069D52FEE8F8C0" box="[151,264,1833,1859]" pageId="5" pageNumber="57" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="2AE5FD7A7C12FFA6FE879D52FECDF8C0" box="[278,301,1833,1859]" count="1" pageId="5" pageNumber="57" type="male"></specimenCount>
,
<collectingCountry id="44F476637C12FFA6FEAA9D52FE5CF8C0" box="[315,444,1833,1859]" name="Philippines" pageId="5" pageNumber="57">Philippines</collectingCountry>
,
<collectingMunicipality id="DC38AC897C12FFA6FE589D52FDF3F8C0" box="[457,531,1833,1859]" pageId="5" pageNumber="57">Luzon</collectingMunicipality>
,
<collectingRegion id="FE27F8117C12FFA6FDB19D52FD97F8C0" box="[544,631,1833,1859]" country="Philippines" name="Quezon" pageId="5" pageNumber="57">Quezon</collectingRegion>
\
<location id="393C60287C12FFA6FD019D52FD02F8C0" LSID="urn:lsid:plazi:treatment:B44A87E57C12FFA5FF069CC7FE2EFCDF:393C60287C12FFA6FD019D52FD02F8C0" box="[656,738,1833,1859]" country="Philippines" municipality="Luzon" name="Infanta" pageId="5" pageNumber="57" stateProvince="Quezon">Infanta</location>
,
<date id="485D10337C12FFA6FD7E9D52FCBDF8C0" box="[751,861,1833,1859]" pageId="5" pageNumber="57" value="1997-04-09">
<collectingDate id="5819E9DB7C12FFA6FD7E9D52FCBDF8C0" box="[751,861,1833,1859]" pageId="5" pageNumber="57" value="1997-04-09">9.IV.1997</collectingDate>
</date>
\ leg.
<collectorName id="911653257C12FFA6FC399D52FC3BF8C0" box="[936,987,1833,1859]" pageId="5" pageNumber="57">Mey</collectorName>
(
<emphasis id="0E97EAE17C12FFA6FC7C9D52FBCDF8C0" bold="true" box="[1005,1069,1833,1859]" pageId="5" pageNumber="57">
<collectionCode id="5AF2AE367C12FFA6FC7C9D52FBCDF8C0" box="[1005,1069,1833,1859]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="5" pageNumber="57">PNM</collectionCode>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="8C8B3CAE7C12FFA6FBD79D52FF1DF8E4" collectingDate="1997-04-09" collectionCode="PNM" collectorName="Mey" country="Philippines" location="Infanta" municipality="Luzon" pageId="5" pageNumber="57" specimenCount="1" specimenCount-male="1" stateProvince="Quezon" typeStatus="paratype">
<emphasis id="0E97EAE17C12FFA6FBD79D52FB27F8C0" bold="true" box="[1094,1223,1833,1859]" pageId="5" pageNumber="57">
<typeStatus id="E35888517C12FFA6FBD79D52FB22F8C0" box="[1094,1218,1833,1859]" pageId="5" pageNumber="57" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="2AE5FD7A7C12FFA6FB419D52FB1EF8C0" box="[1232,1278,1833,1859]" count="1" pageId="5" pageNumber="57" type="male">1 ♂</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="8C8B3CAE7C12FFA6FE999D36FB76F8E4" box="[264,1174,1869,1895]" collectingDate="2000-03-17" collectingDateMax="2000-03-19" collectingDateMin="2000-03-17" collectorName="Mey &amp; Richter" country="Philippines" location="Mt. Banahaw" municipality="Kinabuhayan" pageId="5" pageNumber="57" specimenCount="3" specimenCount-male="3" stateProvince="Quezon" typeStatus="paratype">
<specimenCount id="2AE5FD7A7C12FFA6FE999D36FED4F8E4" box="[264,308,1869,1895]" count="3" pageId="5" pageNumber="57" type="male">3 ♂</specimenCount>
,
<location id="393C60287C12FFA6FED19D36FE37F8E4" LSID="urn:lsid:plazi:treatment:B44A87E57C12FFA5FF069CC7FE2EFCDF:393C60287C12FFA6FED19D36FE37F8E4" box="[320,471,1869,1895]" country="Philippines" municipality="Kinabuhayan" name="Mt. Banahaw" pageId="5" pageNumber="57" stateProvince="Quezon">Mt. Banahaw</location>
\
<collectingMunicipality id="DC38AC897C12FFA6FE7C9D36FD62F8E4" box="[493,642,1869,1895]" pageId="5" pageNumber="57">Kinabuhayan</collectingMunicipality>
\
<date id="485D10337C12FFA6FD099D36FCDBF8E4" box="[664,827,1869,1895]" pageId="5" pageNumber="57" value="2000-03-17" valueMax="2000-03-19" valueMin="2000-03-17">
<collectingDate id="5819E9DB7C12FFA6FD099D36FCDBF8E4" box="[664,827,1869,1895]" pageId="5" pageNumber="57" value="2000-03-17" valueMax="2000-03-19" valueMin="2000-03-17">17-19.III.2000</collectingDate>
</date>
\ leg.
<collectorName id="911653257C12FFA6FC109D36FC54F8E4" box="[897,948,1869,1895]" pageId="5" pageNumber="57">Mey</collectorName>
&amp;
<collectorName id="911653257C12FFA6FC499D36FBCAF8E4" box="[984,1066,1869,1895]" pageId="5" pageNumber="57">Richter</collectorName>
(
<emphasis id="0E97EAE17C12FFA6FBAA9D36FB6DF8E4" bold="true" box="[1083,1165,1869,1895]" pageId="5" pageNumber="57">AdMU</emphasis>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="74F965787C12FFA6FF569D0AFAE2F82C" pageId="5" pageNumber="57" type="diagnosis">
<paragraph id="3C5C36F37C12FFA6FF569D0AFAE2F82C" blockId="5.[151,1436,1833,1967]" pageId="5" pageNumber="57">
<emphasis id="0E97EAE17C12FFA6FF569D0AFEA2F808" bold="true" box="[199,322,1905,1931]" pageId="5" pageNumber="57">Diagnosis.</emphasis>
Male adults of this species are characterized by the lateral projections of the tergum 10 composed of a giant dorsal spine and a tiny ventral tubercle. The epiproct is small and elliptical in the dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C11FFA5FF569AEDFD1FFD9A" pageId="6" pageNumber="58" type="description">
<paragraph id="3C5C36F37C11FFA5FF569AEDFB3BFF7B" blockId="6.[151,1437,150,860]" pageId="6" pageNumber="58">
<emphasis id="0E97EAE17C11FFA5FF569AEDFDA0FF33" bold="true" box="[199,576,150,176]" pageId="6" pageNumber="58">Description. Male adult habitus.</emphasis>
Forewing length is 4.0
<quantity id="FB1B9B167C11FFA5FCD79AEDFC79FF32" box="[838,921,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="58" unit="mm" value="4.2">4.2 mm</quantity>
(n=5). Head dark brown, wider than pronotum; compound eyes black; antennae light brown; mouthparts brownish to brown. Pronotum light brown without dark rugosities; wings subhyaline with darker veins; legs light brown. Abdomen brown to light brown.
</paragraph>
<paragraph id="3C5C36F37C11FFA5FF569B79FD1FFD9A" blockId="6.[151,1437,150,860]" pageId="6" pageNumber="58">
<emphasis id="0E97EAE17C11FFA5FF569B79FE70FE9F" bold="true" box="[199,400,258,284]" pageId="6" pageNumber="58">Male terminalia.</emphasis>
Tergum 9 well sclerotized anteriorly, less so posteriorly, this margin terminating in a wide, produced sclerite with tiny granules. Sternum 9 longer than wide, distal portion with narrow trapezoidal subgenital plate, the plate wider than long; vesicle in ventral view dark brown, ovoid, covered with long, dense hairs. Tergum 10 with well sclerotized lateral projections terminating into a giant dorsal spine and a tiny ventral tubercle. Central plate sclerotized and trilobed, lateral lobes subtriangular, together with a circular; medial lobe darkly pigmented, with a small knob-like projection.
<taxonomicName id="FBE34D707C11FFA5FDB29BCCFD91FE52" box="[547,625,439,465]" class="Insecta" family="Kateretidae" genus="Cercus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="0E97EAE17C11FFA5FDB29BCCFD91FE52" box="[547,625,439,465]" italics="true" pageId="6" pageNumber="58">Cercus</emphasis>
</taxonomicName>
twice as long as wide, curved dorsally, with setation, without terminal spine. Epiproct darkly sclerotized, and strongly curved cephalic in lateral view. Subanal lobe with sclerotized lateral margins and apex membranous, ventral furrows vague.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C11FFA5FF569859FD43FDE2" pageId="6" pageNumber="58" type="etymology">
<paragraph id="3C5C36F37C11FFA5FF569859FD43FDE2" blockId="6.[151,1437,150,860]" pageId="6" pageNumber="58">
<emphasis id="0E97EAE17C11FFA5FF569859FEAEFDBF" bold="true" box="[199,334,546,572]" pageId="6" pageNumber="58">Etymology.</emphasis>
The specific epithet refers to the cephalically curved epiproct in lateral view as the distinguishing characteristic from
<taxonomicName id="FBE34D707C11FFA5FEE0983CFE0BFDE3" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[369,491,582,608]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="0E97EAE17C11FFA5FEE0983CFE0BFDE3" box="[369,491,582,608]" italics="true" pageId="6" pageNumber="58">R. andreae</emphasis>
</taxonomicName>
and
<taxonomicName id="FBE34D707C11FFA5FDB0983CFD7CFDE3" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[545,668,583,608]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="cayasan">
<emphasis id="0E97EAE17C11FFA5FDB0983CFD7CFDE3" box="[545,668,583,608]" italics="true" pageId="6" pageNumber="58">R. cayasan</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C11FFA5FF569811FB97FD07" box="[199,1143,618,645]" pageId="6" pageNumber="58" type="distribution">
<paragraph id="3C5C36F37C11FFA5FF569811FB97FD07" blockId="6.[151,1437,150,860]" box="[199,1143,618,645]" pageId="6" pageNumber="58">
<emphasis id="0E97EAE17C11FFA5FF569811FE81FD07" bold="true" box="[199,353,618,644]" pageId="6" pageNumber="58">Distribution.</emphasis>
This species is known so far only from
<collectingRegion id="FE27F8117C11FFA5FCB09811FC05FD07" box="[801,997,618,644]" country="Philippines" name="Quezon" pageId="6" pageNumber="58">Quezon Province</collectingRegion>
,
<collectingCountry id="44F476637C11FFA5FC609811FB93FD07" box="[1009,1139,618,644]" name="Philippines" pageId="6" pageNumber="58">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C11FFA5FF5698F5FE2EFCDF" pageId="6" pageNumber="58" type="discussion">
<paragraph id="3C5C36F37C11FFA5FF5698F5FE2EFCDF" blockId="6.[151,1437,150,860]" pageId="6" pageNumber="58">
<emphasis id="0E97EAE17C11FFA5FF5698F5FEDBFD2B" bold="true" box="[199,315,654,680]" pageId="6" pageNumber="58">Remarks.</emphasis>
The novel species is a typical of
<taxonomicName id="FBE34D707C11FFA5FD2F98F4FCADFD2B" authorityName="Luzon, Negros, Mindanao" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[702,845,654,680]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="malayana">
<emphasis id="0E97EAE17C11FFA5FD2F98F4FCADFD2B" box="[702,845,654,680]" italics="true" pageId="6" pageNumber="58">R. malayana</emphasis>
</taxonomicName>
group,
<emphasis id="0E97EAE17C11FFA5FC3698F4FC05FD2B" box="[935,997,655,680]" italics="true" pageId="6" pageNumber="58">sensu</emphasis>
<bibRefCitation id="58724B027C11FFA5FC7F98F5FB26FD2B" author="Sivec, I. &amp; Harper, P. P. &amp; Shimizu, T." box="[1006,1222,654,680]" pageId="6" pageNumber="58" pagination="1 - 122" refId="ref4452" refString="Sivec, I., Harper, P. P. &amp; Shimizu, T. (2008) Contribution to the study of the Oriental genus Rhopalopsole (Plecoptera: Leuctridae). Scopolia, 64, 1-122. [https://www.zobodat.at/pdf/Scopolia_64_0001-0122.pdf]" type="journal article" year="2008">
Sivec
<emphasis id="0E97EAE17C11FFA5FBA498F4FB8FFD2B" box="[1077,1135,654,680]" italics="true" pageId="6" pageNumber="58">et al.</emphasis>
(2008)
</bibRefCitation>
. It can be possibly confused with
<taxonomicName id="FBE34D707C11FFA5FEAC98C8FE57FD4F" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[317,439,690,716]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="0E97EAE17C11FFA5FEAC98C8FE57FD4F" box="[317,439,690,716]" italics="true" pageId="6" pageNumber="58">R. andreae</emphasis>
</taxonomicName>
and
<taxonomicName id="FBE34D707C11FFA5FE7F98C8FD89FD4F" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[494,617,691,716]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="cayasan">
<emphasis id="0E97EAE17C11FFA5FE7F98C8FD89FD4F" box="[494,617,691,716]" italics="true" pageId="6" pageNumber="58">R. cayasan</emphasis>
</taxonomicName>
having tergum 10 lateral projections with a giant dorsal spine. However,
<taxonomicName id="FBE34D707C11FFA5FF0698ACFEFCFD73" authorityName="Pelingen" authorityYear="2025" box="[151,284,726,752]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="rotundus" status="sp. nov.">
<emphasis id="0E97EAE17C11FFA5FF0698ACFEFCFD73" box="[151,284,726,752]" italics="true" pageId="6" pageNumber="58">R. rotundus</emphasis>
</taxonomicName>
<emphasis id="0E97EAE17C11FFA5FEB798ADFE63FD73" bold="true" box="[294,387,726,752]" pageId="6" pageNumber="58">
<taxonomicNameLabel id="15A4579A7C11FFA5FEB798ADFE63FD73" box="[294,387,726,752]" pageId="6" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from
<taxonomicName id="FBE34D707C11FFA5FD5598ACFCA1FD73" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[708,833,726,752]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="0E97EAE17C11FFA5FD5598ACFCA1FD73" box="[708,833,726,752]" italics="true" pageId="6" pageNumber="58">R. andreae</emphasis>
</taxonomicName>
with its epiproct being more cephalically curved as compared to the
<taxonomicName id="FBE34D707C11FFA5FEC59880FE2EFC97" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[340,462,762,788]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="andreae">
<emphasis id="0E97EAE17C11FFA5FEC59880FE2EFC97" box="[340,462,762,788]" italics="true" pageId="6" pageNumber="58">R. andreae</emphasis>
</taxonomicName>
epiproct which is more straightly projected caudally in lateral view (
<bibRefCitation id="58724B027C11FFA5FB479881FF56FCBB" author="Sivec, I. &amp; Harper, P. P. &amp; Shimizu, T." pageId="6" pageNumber="58" pagination="1 - 122" refId="ref4452" refString="Sivec, I., Harper, P. P. &amp; Shimizu, T. (2008) Contribution to the study of the Oriental genus Rhopalopsole (Plecoptera: Leuctridae). Scopolia, 64, 1-122. [https://www.zobodat.at/pdf/Scopolia_64_0001-0122.pdf]" type="journal article" year="2008">
Sivec
<emphasis id="0E97EAE17C11FFA5FA8C9880FAB5FC97" box="[1309,1365,762,788]" italics="true" pageId="6" pageNumber="58">et al.</emphasis>
2008: 31
</bibRefCitation>
). Interestingly,
<taxonomicName id="FBE34D707C11FFA5FEF39964FE3BFCBB" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[354,475,799,824]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="cayasan">
<emphasis id="0E97EAE17C11FFA5FEF39964FE3BFCBB" box="[354,475,799,824]" italics="true" pageId="6" pageNumber="58">R. cayasan</emphasis>
</taxonomicName>
can also be differentiated from
<taxonomicName id="FBE34D707C11FFA5FCA99964FC59FCBB" authorityName="Pelingen" authorityYear="2025" box="[824,953,798,824]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="58" phylum="Arthropoda" rank="species" species="rotundus">
<emphasis id="0E97EAE17C11FFA5FCA99964FC59FCBB" box="[824,953,798,824]" italics="true" pageId="6" pageNumber="58">R. rotundus</emphasis>
</taxonomicName>
due to its shorter upright epiproct, in lateral view (
<bibRefCitation id="58724B027C11FFA5FF4D9939FE26FCDF" author="Sivec, I. &amp; Harper, P. P. &amp; Shimizu, T." box="[220,454,834,860]" pageId="6" pageNumber="58" pagination="1 - 122" refId="ref4452" refString="Sivec, I., Harper, P. P. &amp; Shimizu, T. (2008) Contribution to the study of the Oriental genus Rhopalopsole (Plecoptera: Leuctridae). Scopolia, 64, 1-122. [https://www.zobodat.at/pdf/Scopolia_64_0001-0122.pdf]" type="journal article" year="2008">
Sivec
<emphasis id="0E97EAE17C11FFA5FEB29938FEBBFCDF" box="[291,347,834,860]" italics="true" pageId="6" pageNumber="58">et al.</emphasis>
2008: 33
</bibRefCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,202 @@
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<treatment id="B44A87E57C14FFA7FF069FABFD54FE52" ID-DOI="http://doi.org/10.5281/zenodo.15012565" ID-Zenodo-Dep="15012565" LSID="urn:lsid:plazi:treatment:B44A87E57C14FFA7FF069FABFD54FE52" httpUri="http://treatment.plazi.org/id/B44A87E57C14FFA7FF069FABFD54FE52" lastPageId="4" lastPageNumber="56" pageId="3" pageNumber="55">
<subSubSection id="74F965787C14FFA0FF069FABFD67FA69" box="[151,647,1488,1515]" pageId="3" pageNumber="55" type="nomenclature">
<paragraph id="3C5C36F37C14FFA0FF069FABFD67FA69" blockId="3.[151,647,1488,1551]" box="[151,647,1488,1515]" pageId="3" pageNumber="55">
<heading id="6714819F7C14FFA0FF069FABFD67FA69" bold="true" box="[151,647,1488,1515]" fontSize="11" level="1" pageId="3" pageNumber="55" reason="1">
<emphasis id="0E97EAE17C14FFA0FF069FABFD67FA69" bold="true" box="[151,647,1488,1515]" pageId="3" pageNumber="55">
<taxonomicName id="FBE34D707C14FFA0FF069FABFDFDFA69" authority="Pelingen" authorityName="Pelingen" authorityYear="2025" box="[151,541,1488,1515]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="absentia" status="sp. nov.">
<emphasis id="0E97EAE17C14FFA0FF069FABFE48FA68" bold="true" box="[151,424,1488,1515]" italics="true" pageId="3" pageNumber="55">Rhopalopsole absentia</emphasis>
Pelingen
</taxonomicName>
,
<taxonomicNameLabel id="15A4579A7C14FFA0FDBB9FABFD67FA69" box="[554,647,1488,1514]" pageId="3" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="74F965787C14FFA0FF069F8FFE74F98D" box="[151,404,1524,1551]" pageId="3" pageNumber="55" type="description">
<paragraph id="3C5C36F37C14FFA0FF069F8FFE74F98D" blockId="3.[151,647,1488,1551]" box="[151,404,1524,1551]" pageId="3" pageNumber="55">
(
<figureCitation id="A4D82A767C14FFA0FF0E9F8FFF13F98D" box="[159,243,1524,1550]" captionStart="FIGURE 2" captionStartId="2.[151,250,1099,1123]" captionTargetBox="[314,1274,179,1068]" captionTargetId="figure-15@2.[314,1274,179,1073]" captionTargetPageId="2" captionText="FIGURE 2. Male adult dorsal habitus. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012547" httpUri="https://zenodo.org/record/15012547/files/figure.png" pageId="3" pageNumber="55">Fig. 2b</figureCitation>
,
<figureCitation id="A4D82A767C14FFA0FF6F9F8FFEFCF98D" box="[254,284,1524,1550]" captionStart="FIGURE 3" captionStartId="3.[151,250,1346,1370]" captionTargetBox="[314,1274,234,1319]" captionTargetId="figure-59@3.[314,1274,234,1319]" captionTargetPageId="3" captionText="FIGURE 3. Male adult lateral habitus. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012549" httpUri="https://zenodo.org/record/15012549/files/figure.png" pageId="3" pageNumber="55">3b</figureCitation>
,
<figureCitation id="A4D82A767C14FFA0FEB99F8FFED8F98C" box="[296,312,1524,1551]" captionStart="FIGURE 5" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[169,1418,1251,1879]" captionTargetId="figure-166@4.[151,1436,1248,1909]" captionTargetPageId="4" captionText="FIGURE 5. Rhopalopsole absentia sp. nov. male terminalia. a dorsal; b lateral; c ventral." figureDoi="http://doi.org/10.5281/zenodo.15012555" httpUri="https://zenodo.org/record/15012555/files/figure.png" pageId="3" pageNumber="55">5</figureCitation>
,
<figureCitation id="A4D82A767C14FFA0FED59F8FFE82F98D" box="[324,354,1524,1550]" captionStart="FIGURE 7" captionStartId="5.[151,250,1546,1570]" captionTargetBox="[157,1426,900,1509]" captionTargetId="figure-42@5.[151,1436,894,1521]" captionTargetPageId="5" captionText="FIGURE 7. Male terminalia. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012559" httpUri="https://zenodo.org/record/15012559/files/figure.png" pageId="3" pageNumber="55">7b</figureCitation>
,
<figureCitation id="A4D82A767C14FFA0FEFF9F8FFE6BF98D" box="[366,395,1524,1550]" captionStart="FIGURE 8" captionStartId="6.[151,250,1315,1339]" captionTargetBox="[159,1434,919,1283]" captionTargetId="figure-320@6.[151,1436,918,1289]" captionTargetPageId="6" captionText="FIGURE 8. Epiproct and tergum 10 lateral process. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012561" httpUri="https://zenodo.org/record/15012561/files/figure.png" pageId="3" pageNumber="55">8b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="74F965787C14FFA0FF069C47FD4FF9F9" pageId="3" pageNumber="55" type="materials_examined">
<paragraph id="3C5C36F37C14FFA0FF069C47FD4FF9F9" blockId="3.[151,1437,1596,2019]" pageId="3" pageNumber="55">
<materialsCitation id="8C8B3CAE7C14FFA0FF069C47FA77F9D5" box="[151,1431,1596,1623]" collectingDate="1998-11-09" collectingDateMax="1998-11-10" collectingDateMin="1998-11-09" collectionCode="PNM" collectorName="Mey &amp; Speidel" country="Philippines" county="Luzon" location="Coto" municipality="Mts" pageId="3" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Zambales" typeStatus="holotype">
<emphasis id="0E97EAE17C14FFA0FF069C47FEEDF9D5" bold="true" box="[151,269,1596,1622]" pageId="3" pageNumber="55">
<typeStatus id="E35888517C14FFA0FF069C47FEE8F9D5" box="[151,264,1596,1622]" pageId="3" pageNumber="55" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="2AE5FD7A7C14FFA0FE899C46FECFF9D4" box="[280,303,1597,1623]" count="1" pageId="3" pageNumber="55" type="male"></specimenCount>
,
<collectingCountry id="44F476637C14FFA0FEAE9C47FE20F9D5" box="[319,448,1596,1622]" name="Philippines" pageId="3" pageNumber="55">Philippines</collectingCountry>
,
<collectingCounty id="D53D4E7F7C14FFA0FE419C46FDF8F9D4" box="[464,536,1597,1623]" pageId="3" pageNumber="55">Luzon</collectingCounty>
<emphasis id="0E97EAE17C14FFA0FDB39C47FDCAF9D5" bold="true" box="[546,554,1596,1622]" pageId="3" pageNumber="55">\</emphasis>
<collectingRegion id="FE27F8117C14FFA0FDA49C47FD42F9D5" box="[565,674,1596,1622]" country="Philippines" name="Zambales" pageId="3" pageNumber="55">Zambales</collectingRegion>
<collectingMunicipality id="DC38AC897C14FFA0FD3C9C46FD3BF9D4" box="[685,731,1597,1623]" pageId="3" pageNumber="55">Mts</collectingMunicipality>
,
<location id="393C60287C14FFA0FD7B9C47FCC0F9D5" LSID="urn:lsid:plazi:treatment:B44A87E57C14FFA7FF069FABFD54FE52:393C60287C14FFA0FD7B9C47FCC0F9D5" box="[746,800,1596,1622]" country="Philippines" county="Luzon" municipality="Mts" name="Coto" pageId="3" pageNumber="55" stateProvince="Zambales">Coto</location>
\
<date id="485D10337C14FFA0FCAE9C47FC38F9D5" box="[831,984,1596,1622]" pageId="3" pageNumber="55" value="1998-11-09" valueMax="1998-11-10" valueMin="1998-11-09">
<collectingDate id="5819E9DB7C14FFA0FCAE9C47FC38F9D5" box="[831,984,1596,1622]" pageId="3" pageNumber="55" value="1998-11-09" valueMax="1998-11-10" valueMin="1998-11-09">
9-10.XI.
<quantity id="FB1B9B167C14FFA0FC0A9C47FC38F9D5" box="[923,984,1596,1622]" metricMagnitude="6" metricUnit="m" metricValue="1.99815" pageId="3" pageNumber="55" unit="m" value="1998150.0">1998</quantity>
</collectingDate>
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, 150 m \ leg.
<collectorName id="911653257C14FFA0FB149C46FB58F9D4" box="[1157,1208,1597,1623]" pageId="3" pageNumber="55">Mey</collectorName>
&amp;
<collectorName id="911653257C14FFA0FB759C47FAD8F9D5" box="[1252,1336,1596,1622]" pageId="3" pageNumber="55">Speidel</collectorName>
(
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<collectionCode id="5AF2AE367C14FFA0FADD9C47FA6CF9D5" box="[1356,1420,1596,1622]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="3" pageNumber="55">PNM</collectionCode>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="8C8B3CAE7C14FFA0FF069C1BFD4FF9F9" box="[151,687,1632,1662]" collectingDate="1998-11-09" collectingDateMax="1998-11-10" collectingDateMin="1998-11-09" collectionCode="PNM" collectorName="Mey &amp; Speidel" country="Philippines" county="Luzon" location="Coto" municipality="Mts" pageId="3" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="Zambales" typeStatus="paratype">
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<typeStatus id="E35888517C14FFA0FF069C1BFEE8F9F9" box="[151,264,1632,1658]" pageId="3" pageNumber="55" type="paratype">Paratype</typeStatus>
:
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1
<emphasis id="0E97EAE17C14FFA0FEB89C19FEDEF9FD" box="[297,318,1634,1662]" italics="true" pageId="3" pageNumber="55"></emphasis>
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, same data as holotype (
<emphasis id="0E97EAE17C14FFA0FDC29C1BFD45F9F9" bold="true" box="[595,677,1632,1658]" pageId="3" pageNumber="55">AdMU</emphasis>
)
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</paragraph>
</subSubSection>
<subSubSection id="74F965787C14FFA0FF569CFFFAEEF941" pageId="3" pageNumber="55" type="diagnosis">
<paragraph id="3C5C36F37C14FFA0FF569CFFFAEEF941" blockId="3.[151,1437,1596,2019]" pageId="3" pageNumber="55">
<emphasis id="0E97EAE17C14FFA0FF569CFFFEA2F91D" bold="true" box="[199,322,1668,1694]" pageId="3" pageNumber="55">Diagnosis.</emphasis>
Male adult of this species is characterized by the bifurcated lateral projections of the tergum and the epiproct, which is curved rather shallowly. So far, this is the only Philippine species without a vesicle.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C14FFA7FF569CB7FCACFF56" lastPageId="4" lastPageNumber="56" pageId="3" pageNumber="55" type="description">
<paragraph id="3C5C36F37C14FFA0FF569CB7FB6FF8AD" blockId="3.[151,1437,1596,2019]" pageId="3" pageNumber="55">
<emphasis id="0E97EAE17C14FFA0FF569CB7FDB1F965" bold="true" box="[199,593,1740,1766]" pageId="3" pageNumber="55">Description. Male adult habitus.</emphasis>
Forewing length is 4.55.0 mm. Head brown to dark brown, wider than pronotum; compound eyes black; antennae brown to light brown; mouthparts brown. Pronotum light brown with slightly dark rugosities; wings subhyaline with darker veins; legs brown. Abdomen brown.
</paragraph>
<paragraph id="3C5C36F37C14FFA7FF569D43FCACFF56" blockId="3.[151,1437,1596,2019]" lastBlockId="4.[151,1436,151,465]" lastPageId="4" lastPageNumber="56" pageId="3" pageNumber="55">
<emphasis id="0E97EAE17C14FFA0FF569D43FE6FF8D1" bold="true" box="[199,399,1848,1874]" pageId="3" pageNumber="55">Male terminalia.</emphasis>
Tergum 9 well sclerotized at anterior margin, less so posteriorly which terminates in a wide, produced sclerite with tiny granules. Sternum 9 longer than wide, distal portion with narrow trapezoidal subgenital plate, the plate wider than longer; vesicle. Tergum 10 with well sclerotized, bifurcating equilateral projections. Central plate sclerotized and trilobed, lateral lobes subtriangular, together with a circular; medial lobe darkly pigmented, with a small knob-like projection.
<taxonomicName id="FBE34D707C14FFA0FD089DB3FD07F861" box="[665,743,1992,2018]" class="Insecta" family="Kateretidae" genus="Cercus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">Cercus</taxonomicName>
twice as long as wide, curved dorsally, with setations, without terminal spine. Epiproct sclerotized, and rather shallowly curved in lateral view. Subanal lobe with sclerotized lateral margins and apex membranous, ventral furrows vague.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C13FFA7FF569AA4FC55FE9E" pageId="4" pageNumber="56" type="etymology">
<paragraph id="3C5C36F37C13FFA7FF569AA4FC55FE9E" blockId="4.[151,1436,151,465]" pageId="4" pageNumber="56">
<emphasis id="0E97EAE17C13FFA7FF569AA4FEAEFF7A" bold="true" box="[199,334,223,249]" pageId="4" pageNumber="56">Etymology.</emphasis>
The specific epithet refers to the absence of a vesicle on the 9th sternum and a terminal spine on the cerci which distinguishes it from all other Philippine
<taxonomicName id="FBE34D707C13FFA7FC859B78FC4EFE9E" authorityName="Klapalek" authorityYear="1912" box="[788,942,259,285]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="0E97EAE17C13FFA7FC859B78FC4EFE9E" box="[788,942,259,285]" italics="true" pageId="4" pageNumber="56">Rhopalopsole</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C13FFA7FF569B5DFADBFEC2" box="[199,1339,294,321]" pageId="4" pageNumber="56" type="distribution">
<paragraph id="3C5C36F37C13FFA7FF569B5DFADBFEC2" blockId="4.[151,1436,151,465]" box="[199,1339,294,321]" pageId="4" pageNumber="56">
<emphasis id="0E97EAE17C13FFA7FF569B5DFE81FEC3" bold="true" box="[199,353,294,320]" pageId="4" pageNumber="56">Distribution.</emphasis>
This species is known so far only from
<collectingRegion id="FE27F8117C13FFA7FCB09B5CFC1BFEC2" box="[801,1019,295,321]" country="Philippines" name="Zambales" pageId="4" pageNumber="56">Zambales Province</collectingRegion>
, Central Luzon,
<collectingCountry id="44F476637C13FFA7FB279B5CFAD8FEC2" box="[1206,1336,295,321]" name="Philippines" pageId="4" pageNumber="56">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C13FFA7FF569B30FD54FE52" pageId="4" pageNumber="56" type="discussion">
<paragraph id="3C5C36F37C13FFA7FF569B30FD54FE52" blockId="4.[151,1436,151,465]" pageId="4" pageNumber="56">
<emphasis id="0E97EAE17C13FFA7FF569B30FEDBFEE6" bold="true" box="[199,315,331,357]" pageId="4" pageNumber="56">Remarks.</emphasis>
The novel species is a typical of
<taxonomicName id="FBE34D707C13FFA7FD239B30FCDFFEE6" authorityName="Luzon, Negros, Mindanao" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[690,831,331,357]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="malayana">
<emphasis id="0E97EAE17C13FFA7FD239B30FCDFFEE6" box="[690,831,331,357]" italics="true" pageId="4" pageNumber="56">R. malayana</emphasis>
</taxonomicName>
group,
<emphasis id="0E97EAE17C13FFA7FC069B30FC35FEE7" box="[919,981,331,356]" italics="true" pageId="4" pageNumber="56">sensu</emphasis>
<bibRefCitation id="58724B027C13FFA7FC4D9B30FB4FFEE6" author="Sivec, I. &amp; Harper, P. P. &amp; Shimizu, T." box="[988,1199,331,357]" pageId="4" pageNumber="56" pagination="1 - 122" refId="ref4452" refString="Sivec, I., Harper, P. P. &amp; Shimizu, T. (2008) Contribution to the study of the Oriental genus Rhopalopsole (Plecoptera: Leuctridae). Scopolia, 64, 1-122. [https://www.zobodat.at/pdf/Scopolia_64_0001-0122.pdf]" type="journal article" year="2008">
Sivec
<emphasis id="0E97EAE17C13FFA7FBB39B30FBBAFEE6" box="[1058,1114,331,357]" italics="true" pageId="4" pageNumber="56">et al.</emphasis>
(2008)
</bibRefCitation>
. The male appears to be closely related to
<taxonomicName id="FBE34D707C13FFA7FE119B14FDEDFE0A" authorityName="Luzon, Negros, Mindanao" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[384,525,367,393]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="malayana">
<emphasis id="0E97EAE17C13FFA7FE119B14FDEDFE0A" box="[384,525,367,393]" italics="true" pageId="4" pageNumber="56">R. malayana</emphasis>
</taxonomicName>
and
<taxonomicName id="FBE34D707C13FFA7FDD59B14FD78FE0B" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[580,664,367,392]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="meyi">
<emphasis id="0E97EAE17C13FFA7FDD59B14FD78FE0B" box="[580,664,367,392]" italics="true" pageId="4" pageNumber="56">R. meyi</emphasis>
</taxonomicName>
having the same bifurcating tergum 10 lateral projections. However,
<taxonomicName id="FBE34D707C13FFA7FF069BE8FEF6FE2E" authorityName="Pelingen" authorityYear="2025" box="[151,278,403,429]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="absentia" status="sp. nov.">
<emphasis id="0E97EAE17C13FFA7FF069BE8FEF6FE2E" box="[151,278,403,429]" italics="true" pageId="4" pageNumber="56">R. absentia</emphasis>
</taxonomicName>
<emphasis id="0E97EAE17C13FFA7FE8C9BE8FE97FE2E" bold="true" box="[285,375,403,429]" pageId="4" pageNumber="56">
<taxonomicNameLabel id="15A4579A7C13FFA7FE8C9BE8FE97FE2E" box="[285,375,403,429]" pageId="4" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from both
<taxonomicName id="FBE34D707C13FFA7FD759BE8FC91FE2E" authorityName="Luzon, Negros, Mindanao" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[740,881,403,429]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="malayana">
<emphasis id="0E97EAE17C13FFA7FD759BE8FC91FE2E" box="[740,881,403,429]" italics="true" pageId="4" pageNumber="56">R. malayana</emphasis>
</taxonomicName>
and
<taxonomicName id="FBE34D707C13FFA7FC389BE8FC1EFE2F" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[937,1022,403,428]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="56" phylum="Arthropoda" rank="species" species="meyi">
<emphasis id="0E97EAE17C13FFA7FC389BE8FC1EFE2F" box="[937,1022,403,428]" italics="true" pageId="4" pageNumber="56">R. meyi</emphasis>
</taxonomicName>
with its epiproct that is only slightly curved as compared to regularly rounded hooks.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,286 @@
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<emphasis id="0E97EAE17C16FFA2FF069D5CFD41F8C1" bold="true" box="[151,673,1831,1858]" pageId="1" pageNumber="53">
<taxonomicName id="FBE34D707C16FFA2FF069D5CFDD7F8C1" authority="Pelingen" authorityName="Pelingen" authorityYear="2025" box="[151,567,1831,1858]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="53" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0E97EAE17C16FFA2FF069D5CFE22F8C1" bold="true" box="[151,450,1831,1858]" italics="true" pageId="1" pageNumber="53">Rhopalopsole luzonensis</emphasis>
Pelingen
</taxonomicName>
,
<taxonomicNameLabel id="15A4579A7C16FFA2FDD59D53FD41F8C1" box="[580,673,1832,1858]" pageId="1" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="74F965787C16FFA2FF069D37FE6DF8E5" box="[151,397,1868,1895]" pageId="1" pageNumber="53" type="description">
<paragraph id="3C5C36F37C16FFA2FF069D37FE6DF8E5" blockId="1.[151,673,1831,1895]" box="[151,397,1868,1895]" pageId="1" pageNumber="53">
(
<figureCitation id="A4D82A767C16FFA2FF0E9D37FF11F8E5" box="[159,241,1868,1894]" captionStart="FIGURE 2" captionStartId="2.[151,250,1099,1123]" captionTargetBox="[314,1274,179,1068]" captionTargetId="figure-15@2.[314,1274,179,1073]" captionTargetPageId="2" captionText="FIGURE 2. Male adult dorsal habitus. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012547" httpUri="https://zenodo.org/record/15012547/files/figure.png" pageId="1" pageNumber="53">Fig. 2a</figureCitation>
,
<figureCitation id="A4D82A767C16FFA2FF6D9D37FEF9F8E5" box="[252,281,1868,1894]" captionStart="FIGURE 3" captionStartId="3.[151,250,1346,1370]" captionTargetBox="[314,1274,234,1319]" captionTargetId="figure-59@3.[314,1274,234,1319]" captionTargetPageId="3" captionText="FIGURE 3. Male adult lateral habitus. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012549" httpUri="https://zenodo.org/record/15012549/files/figure.png" pageId="1" pageNumber="53">3a</figureCitation>
,
<figureCitation id="A4D82A767C16FFA2FEB49D37FED5F8E5" box="[293,309,1868,1894]" captionStart="FIGURE 4" captionStartId="4.[151,250,1165,1189]" captionTargetBox="[159,1402,486,1132]" captionTargetId="figure-149@4.[151,1436,486,1139]" captionTargetPageId="4" captionText="FIGURE 4. Rhopalopsole luzonensis sp. nov. male terminalia. a dorsal; b lateral; c ventral." figureDoi="http://doi.org/10.5281/zenodo.15012551" httpUri="https://zenodo.org/record/15012551/files/figure.png" pageId="1" pageNumber="53">4</figureCitation>
,
<figureCitation id="A4D82A767C16FFA2FED09D36FEBDF8E4" box="[321,349,1869,1895]" captionStart="FIGURE 7" captionStartId="5.[151,250,1546,1570]" captionTargetBox="[157,1426,900,1509]" captionTargetId="figure-42@5.[151,1436,894,1521]" captionTargetPageId="5" captionText="FIGURE 7. Male terminalia. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012559" httpUri="https://zenodo.org/record/15012559/files/figure.png" pageId="1" pageNumber="53">7a</figureCitation>
,
<figureCitation id="A4D82A767C16FFA2FEF89D37FE64F8E5" box="[361,388,1868,1894]" captionStart="FIGURE 8" captionStartId="6.[151,250,1315,1339]" captionTargetBox="[159,1434,919,1283]" captionTargetId="figure-320@6.[151,1436,918,1289]" captionTargetPageId="6" captionText="FIGURE 8. Epiproct and tergum 10 lateral process. a Rhopalopsole luzonensis sp. nov.; b Rhopalopsole absentia sp. nov.; c Rhopalopsole rotundus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.15012561" httpUri="https://zenodo.org/record/15012561/files/figure.png" pageId="1" pageNumber="53">8a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="74F965787C16FFA2FF069DEFFE0BF875" pageId="1" pageNumber="53" type="materials_examined">
<paragraph id="3C5C36F37C16FFA2FF069DEFFE0BF875" blockId="1.[151,1436,1940,2039]" pageId="1" pageNumber="53">
<materialsCitation id="8C8B3CAE7C16FFA2FF069DEFFA77F82D" box="[151,1431,1940,1967]" collectingDate="1997-11-25" collectionCode="PNM" collectorName="Adunot-river Untenauf &amp; Mey &amp; Ebert &amp; Nuss" country="Philippines" location="Luzon" pageId="1" pageNumber="53" specimenCount="1" specimenCount-male="1" stateProvince="Benguet" typeStatus="holotype">
<emphasis id="0E97EAE17C16FFA2FF069DEFFEEDF82D" bold="true" box="[151,269,1940,1966]" pageId="1" pageNumber="53">
<typeStatus id="E35888517C16FFA2FF069DEFFEE8F82D" box="[151,264,1940,1966]" pageId="1" pageNumber="53" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="2AE5FD7A7C16FFA2FE879DEEFECDF82C" box="[278,301,1941,1967]" count="1" pageId="1" pageNumber="53" type="male"></specimenCount>
,
<collectingCountry id="44F476637C16FFA2FEAA9DEFFE5CF82D" box="[315,444,1940,1966]" name="Philippines" pageId="1" pageNumber="53">Philippines</collectingCountry>
,
<location id="393C60287C16FFA2FE589DEEFDF1F82C" LSID="urn:lsid:plazi:treatment:B44A87E57C16FFA0FF069D5CFC99FF56:393C60287C16FFA2FE589DEEFDF1F82C" box="[457,529,1941,1967]" country="Philippines" name="Luzon" pageId="1" pageNumber="53" stateProvince="Benguet">Luzon</location>
\
<collectingRegion id="FE27F8117C16FFA2FDB89DEEFD69F82C" box="[553,649,1941,1967]" country="Philippines" name="Benguet" pageId="1" pageNumber="53">Benguet</collectingRegion>
,
<date id="485D10337C16FFA2FD069DEFFCF6F82C" box="[663,790,1940,1967]" pageId="1" pageNumber="53" value="1997-11-25">
<collectingDate id="5819E9DB7C16FFA2FD069DEFFCF6F82C" box="[663,790,1940,1967]" pageId="1" pageNumber="53" value="1997-11-25">25.XI.1997</collectingDate>
</date>
\
<collectorName id="911653257C16FFA2FCBF9DEFFBD1F82D" box="[814,1073,1940,1966]" pageId="1" pageNumber="53">Adunot-river Untenauf</collectorName>
\ leg.
<collectorName id="911653257C16FFA2FBED9DEEFB50F82C" box="[1148,1200,1941,1967]" pageId="1" pageNumber="53">Mey</collectorName>
,
<collectorName id="911653257C16FFA2FB2C9DEFFB1DF82D" box="[1213,1277,1940,1966]" pageId="1" pageNumber="53">Ebert</collectorName>
,
<collectorName id="911653257C16FFA2FA9B9DEEFADAF82C" box="[1290,1338,1941,1967]" pageId="1" pageNumber="53">Nuss</collectorName>
(
<emphasis id="0E97EAE17C16FFA2FADD9DEFFA6CF82D" bold="true" box="[1356,1420,1940,1966]" pageId="1" pageNumber="53">
<collectionCode id="5AF2AE367C16FFA2FADD9DEFFA6CF82D" box="[1356,1420,1940,1966]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="1" pageNumber="53">PNM</collectionCode>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="8C8B3CAE7C16FFA2FF069DC3FC80F850" box="[151,864,1976,2003]" collectingDate="1997-04-09" collectorName="Mey" country="Philippines" location="Infanta" municipality="Luzon" pageId="1" pageNumber="53" specimenCount="2" specimenCount-male="2" stateProvince="Quezon" typeStatus="paratype">
<emphasis id="0E97EAE17C16FFA2FF069DC3FEF8F851" bold="true" box="[151,280,1976,2002]" pageId="1" pageNumber="53">
<typeStatus id="E35888517C16FFA2FF069DC3FEF3F851" box="[151,275,1976,2002]" pageId="1" pageNumber="53" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="2AE5FD7A7C16FFA2FE8E9DC3FEABF850" box="[287,331,1976,2003]" count="2" pageId="1" pageNumber="53" type="male">2 ♂</specimenCount>
,
<collectingMunicipality id="DC38AC897C16FFA2FEC69DC2FE41F850" box="[343,417,1977,2003]" pageId="1" pageNumber="53">Luzon</collectingMunicipality>
,
<collectingRegion id="FE27F8117C16FFA2FE3D9DC3FDE3F851" box="[428,515,1976,2002]" country="Philippines" name="Quezon" pageId="1" pageNumber="53">Quezon</collectingRegion>
\
<location id="393C60287C16FFA2FD899DC3FD8AF851" LSID="urn:lsid:plazi:treatment:B44A87E57C16FFA0FF069D5CFC99FF56:393C60287C16FFA2FD899DC3FD8AF851" box="[536,618,1976,2002]" country="Philippines" municipality="Luzon" name="Infanta" pageId="1" pageNumber="53" stateProvince="Quezon">Infanta</location>
,
<date id="485D10337C16FFA2FDE49DC3FD04F851" box="[629,740,1976,2002]" pageId="1" pageNumber="53" value="1997-04-09">
<collectingDate id="5819E9DB7C16FFA2FDE49DC3FD04F851" box="[629,740,1976,2002]" pageId="1" pageNumber="53" value="1997-04-09">9.IV.1997</collectingDate>
</date>
\ leg.
<collectorName id="911653257C16FFA2FCB89DC2FC80F850" box="[809,864,1977,2003]" pageId="1" pageNumber="53">Mey</collectorName>
</materialsCitation>
;
<materialsCitation id="8C8B3CAE7C16FFA2FCFA9DC3FE9FF875" collectingDate="2000-03-17" collectingDateMax="2000-03-19" collectingDateMin="2000-03-17" collectorName="Mey &amp; Richter" country="Philippines" location="Kinabuhayan" municipality="Mt. Banahaw" pageId="1" pageNumber="53" specimenCount="1" specimenCount-male="1" stateProvince="Quezon" typeStatus="paratype">
<specimenCount id="2AE5FD7A7C16FFA2FCFA9DC3FC76F850" box="[875,918,1976,2003]" count="1" pageId="1" pageNumber="53" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="DC38AC897C16FFA2FC339DC2FBD9F851" box="[930,1081,1976,2003]" pageId="1" pageNumber="53">Mt. Banahaw</collectingMunicipality>
\
<location id="393C60287C16FFA2FBDF9DC3FB03F851" LSID="urn:lsid:plazi:treatment:B44A87E57C16FFA0FF069D5CFC99FF56:393C60287C16FFA2FBDF9DC3FB03F851" box="[1102,1251,1976,2002]" country="Philippines" municipality="Mt. Banahaw" name="Kinabuhayan" pageId="1" pageNumber="53" stateProvince="Quezon">Kinabuhayan</location>
\
<date id="485D10337C16FFA2FB689DC3FA7CF851" box="[1273,1436,1976,2002]" pageId="1" pageNumber="53" value="2000-03-17" valueMax="2000-03-19" valueMin="2000-03-17">
<collectingDate id="5819E9DB7C16FFA2FB689DC3FA7CF851" box="[1273,1436,1976,2002]" pageId="1" pageNumber="53" value="2000-03-17" valueMax="2000-03-19" valueMin="2000-03-17">17-19.III.2000</collectingDate>
</date>
\ leg.
<collectorName id="911653257C16FFA2FF479DA6FEE9F874" box="[214,265,2013,2039]" pageId="1" pageNumber="53">Mey</collectorName>
&amp;
<collectorName id="911653257C16FFA2FEBC9DA7FE9FF875" box="[301,383,2012,2038]" pageId="1" pageNumber="53">Richter</collectorName>
</materialsCitation>
(
<emphasis id="0E97EAE17C16FFA2FE019DA7FE02F875" bold="true" box="[400,482,2012,2038]" pageId="1" pageNumber="53">AdMU</emphasis>
)
</paragraph>
</subSubSection>
<caption id="689C667B7C15FFA1FF069E30FEA7FB04" ID-DOI="http://doi.org/10.5281/zenodo.15012547" ID-Zenodo-Dep="15012547" httpUri="https://zenodo.org/record/15012547/files/figure.png" pageId="2" pageNumber="54" startId="2.[151,250,1099,1123]" targetBox="[314,1274,179,1068]" targetPageId="2" targetType="figure">
<paragraph id="3C5C36F37C15FFA1FF069E30FEA7FB04" blockId="2.[151,1436,1099,1159]" pageId="2" pageNumber="54">
<emphasis id="0E97EAE17C15FFA1FF069E30FDA3FBE0" bold="true" box="[151,579,1099,1123]" pageId="2" pageNumber="54">FIGURE 2. Male adult dorsal habitus. a</emphasis>
<taxonomicName id="FBE34D707C15FFA1FDD99E30FCA3FBE0" authorityName="Pelingen" authorityYear="2025" box="[584,835,1099,1123]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0E97EAE17C15FFA1FDD99E30FCA3FBE0" box="[584,835,1099,1123]" italics="true" pageId="2" pageNumber="54">Rhopalopsole luzonensis</emphasis>
</taxonomicName>
<emphasis id="0E97EAE17C15FFA1FCD99E30FC78FBE0" bold="true" box="[840,920,1099,1123]" pageId="2" pageNumber="54">
<taxonomicNameLabel id="15A4579A7C15FFA1FCD99E30FC78FBE0" box="[840,920,1099,1123]" pageId="2" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="0E97EAE17C15FFA1FC359E30FC52FBE0" bold="true" box="[932,946,1099,1123]" pageId="2" pageNumber="54">b</emphasis>
<taxonomicName id="FBE34D707C15FFA1FC299E30FB7FFBE0" authorityName="Pelingen" authorityYear="2025" box="[952,1183,1099,1123]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="absentia" status="sp. nov.">
<emphasis id="0E97EAE17C15FFA1FC299E30FB7FFBE0" box="[952,1183,1099,1123]" italics="true" pageId="2" pageNumber="54">Rhopalopsole absentia</emphasis>
</taxonomicName>
<emphasis id="0E97EAE17C15FFA1FB359E30FB13FBE0" bold="true" box="[1188,1267,1099,1123]" pageId="2" pageNumber="54">
<taxonomicNameLabel id="15A4579A7C15FFA1FB359E30FB13FBE0" box="[1188,1267,1099,1123]" pageId="2" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="0E97EAE17C15FFA1FA919E30FAEBFBE0" bold="true" box="[1280,1291,1099,1123]" pageId="2" pageNumber="54">c</emphasis>
<taxonomicName id="FBE34D707C15FFA1FA809E30FF10FB04" authorityName="Pelingen" authorityYear="2025" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundus" status="sp. nov.">
<emphasis id="0E97EAE17C15FFA1FA809E30FF10FB04" italics="true" pageId="2" pageNumber="54">Rhopalopsole rotundus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="15A4579A7C15FFA1FF669E14FEA7FB04" box="[247,327,1135,1159]" pageId="2" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="74F965787C15FFA1FF569ECEFE60FA94" pageId="2" pageNumber="54" type="diagnosis">
<paragraph id="3C5C36F37C15FFA1FF569ECEFE60FA94" blockId="2.[151,1437,1205,2024]" pageId="2" pageNumber="54">
<emphasis id="0E97EAE17C15FFA1FF569ECEFEA2FB4C" bold="true" box="[199,322,1205,1231]" pageId="2" pageNumber="54">Diagnosis.</emphasis>
The lateral projections of the tergum 10 of males bifurcate, with the top spine appearing longer than the lower one in lateral view. The cercus bears long setations, and in dorsal aspect, the epiproct looks like an elongated oval form.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C15FFA1FF569F5AFEB1F94B" pageId="2" pageNumber="54" type="description">
<paragraph id="3C5C36F37C15FFA1FF569F5AFBD5FA00" blockId="2.[151,1437,1205,2024]" pageId="2" pageNumber="54">
<emphasis id="0E97EAE17C15FFA1FF569F5AFDB2FAB8" bold="true" box="[199,594,1313,1339]" pageId="2" pageNumber="54">Description. Male adult habitus.</emphasis>
Forewing length:
<quantity id="FB1B9B167C15FFA1FCBB9F5AFC55FABF" box="[810,949,1313,1340]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="2" pageNumber="54" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.54.8 mm</quantity>
(n=3). Head light brown to brown, wider than pronotum; compound eyes black; antennae light brown; mouthparts brown. Pronotum light brown with dark rugosities; wings subhyaline with darker veins; legs brown. Abdomen light brown.
</paragraph>
<paragraph id="3C5C36F37C15FFA1FF569FF6FEB1F94B" blockId="2.[151,1437,1205,2024]" pageId="2" pageNumber="54">
<emphasis id="0E97EAE17C15FFA1FF569FF6FE6EFA24" bold="true" box="[199,398,1421,1447]" pageId="2" pageNumber="54">Male terminalia.</emphasis>
Tergum 9 strongly sclerotized at anterior margin, weakly sclerotized posteriorly, ornamented throughout. Sternum 9 longer than wide, distal portion with narrow trapezoidal subgenital plate, the plate wider than longer; vesicle in ventral view dark brown, ovoid, covered with long, dense hairs. Tergum 10 from lateral view the upper spine appears longer than the lower spine, with well-sclerotized, bifurcate lateral projections that terminate in two sharp, parallel-sided points in the dorsal aspect. Central plate sclerotized and trilobed, lateral lobes an elongated subtrapezoid, medial lobe darkly pigmented.
<taxonomicName id="FBE34D707C15FFA1FCAB9C3AFC68F9D8" box="[826,904,1601,1627]" class="Insecta" family="Kateretidae" genus="Cercus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">Cercus</taxonomicName>
twice longer than wide, curved dorsally, with setation, without terminal spine. Epiproct strongly sclerotized and darkly sclerotized laterally, oval in dorsal view, longer than wide. Subanal lobe with prominent basal sclerotization, ambiguous ventral furrows, membranous lateral edges, and apex.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C15FFA1FF569CAAFAA5F968" box="[199,1349,1745,1772]" pageId="2" pageNumber="54" type="etymology">
<paragraph id="3C5C36F37C15FFA1FF569CAAFAA5F968" blockId="2.[151,1437,1205,2024]" box="[199,1349,1745,1772]" pageId="2" pageNumber="54">
<emphasis id="0E97EAE17C15FFA1FF569CAAFEAEF968" bold="true" box="[199,334,1745,1771]" pageId="2" pageNumber="54">Etymology.</emphasis>
The species is named after its current known distribution in the Luzon Island,
<collectingCountry id="44F476637C15FFA1FB2E9CAAFAA1F968" box="[1215,1345,1745,1771]" name="Philippines" pageId="2" pageNumber="54">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C15FFA1FF569C8EFAB3F88C" box="[199,1363,1781,1808]" pageId="2" pageNumber="54" type="distribution">
<paragraph id="3C5C36F37C15FFA1FF569C8EFAB3F88C" blockId="2.[151,1437,1205,2024]" box="[199,1363,1781,1808]" pageId="2" pageNumber="54">
<emphasis id="0E97EAE17C15FFA1FF569C8EFE81F88C" bold="true" box="[199,353,1781,1807]" pageId="2" pageNumber="54">Distribution.</emphasis>
This species is known so far only from
<collectingRegion id="FE27F8117C15FFA1FCB09C8DFC9EF893" box="[801,894,1782,1808]" country="Philippines" name="Benguet" pageId="2" pageNumber="54">Benguet</collectingRegion>
and
<collectingRegion id="FE27F8117C15FFA1FC249C8EFBECF88C" box="[949,1036,1781,1807]" country="Philippines" name="Quezon" pageId="2" pageNumber="54">Quezon</collectingRegion>
provinces in the
<collectingCountry id="44F476637C15FFA1FB5C9C8EFAAFF88C" box="[1229,1359,1781,1807]" name="Philippines" pageId="2" pageNumber="54">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="74F965787C15FFA0FF569D62FC99FF56" lastPageId="3" lastPageNumber="55" pageId="2" pageNumber="54" type="discussion">
<paragraph id="3C5C36F37C15FFA0FF569D62FC99FF56" blockId="2.[151,1437,1205,2024]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="55" pageId="2" pageNumber="54">
<emphasis id="0E97EAE17C15FFA1FF569D62FEDBF8B0" bold="true" box="[199,315,1817,1843]" pageId="2" pageNumber="54">Remarks.</emphasis>
The novel species is a typical of
<taxonomicName id="FBE34D707C15FFA1FD2F9D61FCAFF8B0" authorityName="Luzon, Negros, Mindanao" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[702,847,1817,1844]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="malayana">
R.
<emphasis id="0E97EAE17C15FFA1FD709D62FCAFF8B0" box="[737,847,1817,1843]" italics="true" pageId="2" pageNumber="54">malayana</emphasis>
</taxonomicName>
group,
<emphasis id="0E97EAE17C15FFA1FC389D61FC07F8B0" box="[937,999,1818,1843]" italics="true" pageId="2" pageNumber="54">sensu</emphasis>
<bibRefCitation id="58724B027C15FFA1FC619D62FB27F8B0" author="Sivec, I. &amp; Harper, P. P. &amp; Shimizu, T." box="[1008,1223,1817,1843]" pageId="2" pageNumber="54" pagination="1 - 122" refId="ref4452" refString="Sivec, I., Harper, P. P. &amp; Shimizu, T. (2008) Contribution to the study of the Oriental genus Rhopalopsole (Plecoptera: Leuctridae). Scopolia, 64, 1-122. [https://www.zobodat.at/pdf/Scopolia_64_0001-0122.pdf]" type="journal article" year="2008">
Sivec
<emphasis id="0E97EAE17C15FFA1FBA69D61FB91F8B0" box="[1079,1137,1817,1843]" italics="true" pageId="2" pageNumber="54">et al.</emphasis>
(2008)
</bibRefCitation>
. The male appears to be closely connected to
<taxonomicName id="FBE34D707C15FFA1FE5D9D45FDBBF8D4" authorityName="Luzon, Negros, Mindanao" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[460,603,1853,1879]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="malayana">
<emphasis id="0E97EAE17C15FFA1FE5D9D45FDBBF8D4" box="[460,603,1853,1879]" italics="true" pageId="2" pageNumber="54">R. malayana</emphasis>
</taxonomicName>
and more similar with
<taxonomicName id="FBE34D707C15FFA1FCFB9D45FC21F8D4" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[874,961,1854,1879]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="meyi">
<emphasis id="0E97EAE17C15FFA1FCFB9D45FC21F8D4" box="[874,961,1854,1879]" italics="true" pageId="2" pageNumber="54">R. meyi</emphasis>
</taxonomicName>
having alike bifurcated tergum 10 lateral projections and strongly hooked epiprocts. However,
<taxonomicName id="FBE34D707C15FFA1FD7D9D19FC60F8F8" authorityName="Pelingen" authorityYear="2025" box="[748,896,1889,1915]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0E97EAE17C15FFA1FD7D9D19FC60F8F8" box="[748,896,1889,1915]" italics="true" pageId="2" pageNumber="54">R. luzonensis</emphasis>
</taxonomicName>
<emphasis id="0E97EAE17C15FFA1FC169D1AFC00F8F8" bold="true" box="[903,992,1889,1915]" pageId="2" pageNumber="54">
<taxonomicNameLabel id="15A4579A7C15FFA1FC169D1AFC00F8F8" box="[903,992,1889,1915]" pageId="2" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from
<taxonomicName id="FBE34D707C15FFA1FA9E9D19FA7CF8F8" authorityName="Luzon, Negros, Mindanao" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[1295,1436,1889,1915]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="malayana">
<emphasis id="0E97EAE17C15FFA1FA9E9D19FA7CF8F8" box="[1295,1436,1889,1915]" italics="true" pageId="2" pageNumber="54">R. malayana</emphasis>
</taxonomicName>
with the upper spine of bifurcate lateral projections on tergum 10 being longer than the lower one in lateral view; while
<taxonomicName id="FBE34D707C15FFA1FF4F9DD1FE94F840" authorityName="Pelingen" authorityYear="2025" box="[222,372,1961,1987]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0E97EAE17C15FFA1FF4F9DD1FE94F840" box="[222,372,1961,1987]" italics="true" pageId="2" pageNumber="54">R. luzonensis</emphasis>
</taxonomicName>
<emphasis id="0E97EAE17C15FFA1FEED9DD2FE38F840" bold="true" box="[380,472,1961,1987]" pageId="2" pageNumber="54">
<taxonomicNameLabel id="15A4579A7C15FFA1FEED9DD2FE38F840" box="[380,472,1961,1987]" pageId="2" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differentiated from
<taxonomicName id="FBE34D707C15FFA1FD2F9DD1FCF4F840" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[702,788,1962,1987]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="meyi">
<emphasis id="0E97EAE17C15FFA1FD2F9DD1FCF4F840" box="[702,788,1962,1987]" italics="true" pageId="2" pageNumber="54">R. meyi</emphasis>
</taxonomicName>
based on a bigger and more ovoid epiproct on its dorsal view. In addition, the posterior margin of sternum 9 is curved smoothly in
<taxonomicName id="FBE34D707C15FFA1FC689DB5FB70F864" authorityName="Pelingen" authorityYear="2025" box="[1017,1168,1997,2023]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0E97EAE17C15FFA1FC689DB5FB70F864" box="[1017,1168,1997,2023]" italics="true" pageId="2" pageNumber="54">R. luzonensis</emphasis>
</taxonomicName>
<emphasis id="0E97EAE17C15FFA1FB089DB6FB15F864" bold="true" box="[1177,1269,1997,2023]" pageId="2" pageNumber="54">
<taxonomicNameLabel id="15A4579A7C15FFA1FB089DB6FB15F864" box="[1177,1269,1997,2023]" pageId="2" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
while
<taxonomicName id="FBE34D707C15FFA1FAD79DB5FA7CF864" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[1350,1436,1998,2023]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="54" phylum="Arthropoda" rank="species" species="meyi">
<emphasis id="0E97EAE17C15FFA1FAD79DB5FA7CF864" box="[1350,1436,1998,2023]" italics="true" pageId="2" pageNumber="54">R. meyi</emphasis>
</taxonomicName>
projects curving posteriorly. Its subanal lobes are also more elongated than that of
<taxonomicName id="FBE34D707C14FFA0FBBF9AECFB61FF33" authorityName="Sivec &amp; Harper" authorityYear="2008" box="[1070,1153,151,176]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="meyi">
<emphasis id="0E97EAE17C14FFA0FBBF9AECFB61FF33" box="[1070,1153,151,176]" italics="true" pageId="3" pageNumber="55">R. meyi</emphasis>
</taxonomicName>
. Lastly,
<taxonomicName id="FBE34D707C14FFA0FB709AECFA94FF32" authorityName="Pelingen" authorityYear="2025" box="[1249,1396,151,177]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
<emphasis id="0E97EAE17C14FFA0FB709AECFA94FF32" box="[1249,1396,151,177]" italics="true" pageId="3" pageNumber="55">R. luzonensis</emphasis>
</taxonomicName>
<emphasis id="0E97EAE17C14FFA0FAEA9AECFF28FF56" bold="true" pageId="3" pageNumber="55">
<taxonomicNameLabel id="15A4579A7C14FFA0FAEA9AECFF28FF56" pageId="3" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
lacks a terminal spine on its cerci in contrast to
<taxonomicName id="FBE34D707C14FFA0FD749AC0FC92FF56" authorityName="Luzon, Negros, Mindanao" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[741,882,187,213]" class="Insecta" family="Leuctridae" genus="Rhopalopsole" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="55" phylum="Arthropoda" rank="species" species="malayana">
<emphasis id="0E97EAE17C14FFA0FD749AC0FC92FF56" box="[741,882,187,213]" italics="true" pageId="3" pageNumber="55">R. malayana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>