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Subfamily
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<paragraph pageId="3" pageNumber="10">Type genus.</paragraph>
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<taxonomicName class="Insecta" family="Acroceridae" genus="Panops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Panops" order="Diptera" pageId="3" pageNumber="10" phylum="Arthropoda" rank="genus">Panops</taxonomicName>
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<taxonomicName id="61F68051F238143FBDF5B01114DBC53D" class="Insecta" family="Acroceridae" genus="Panops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Panops" order="Diptera" pageId="3" pageNumber="10" phylum="Arthropoda" rank="genus">Panops</taxonomicName>
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Lamarck, 1804: 263.
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<paragraph pageId="3" pageNumber="10">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="10">Usually large and densely pilose, body shape never arched; antennal flagellum elongate cylindrical to paddle-shaped, sometimes tapered but never stylate, usually lacking terminal setae; postpronotal lobes never meeting medially; wing venation complete to wing margin (rarely reduced), cells m3, d, bm and basal r4+5 typically present, closed distally; tibial spines present (rarely absent); larvae exclusively parasitoids of mygalomorph spiders.</paragraph>
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<paragraph id="86D67F1B983005C11F97110AFD3F48D9" pageId="3" pageNumber="10">Usually large and densely pilose, body shape never arched; antennal flagellum elongate cylindrical to paddle-shaped, sometimes tapered but never stylate, usually lacking terminal setae; postpronotal lobes never meeting medially; wing venation complete to wing margin (rarely reduced), cells m3, d, bm and basal r4+5 typically present, closed distally; tibial spines present (rarely absent); larvae exclusively parasitoids of mygalomorph spiders.</paragraph>
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@ -1,289 +1,289 @@
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<taxonomicNameLabel pageId="0" pageNumber="129">comb. nov.</taxonomicNameLabel>
|
||||
<treatment id="E35D008D421E5A10861EFB0A1EF660D5" ID-GBIF-Taxon="206092924" LSID="urn:lsid:plazi:treatment:E35D008D421E5A10861EFB0A1EF660D5" httpUri="http://treatment.plazi.org/id/E35D008D421E5A10861EFB0A1EF660D5" lastPageNumber="129" pageId="0" pageNumber="129" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae" scope_tribe="Quediini">
|
||||
<subSubSection id="909BAAF53B1F80F2990D621ACB70FCAC" pageId="0" pageNumber="129" type="nomenclature">
|
||||
<paragraph id="D564AB69549CC1A87501E9EDDEF18CAB" pageId="0" pageNumber="129">
|
||||
<taxonomicName id="8F39605A464D07621582DA6F070A4864" LSID="E35D008D-421E-5A10-861E-FB0A1EF660D5" authority="(Smetana, 1971)" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="comb. nov.">Iratiquedius amabilis (Smetana, 1971)</taxonomicName>
|
||||
<taxonomicNameLabel id="C463DC9BC64C58248A604C9C0E5B5AFC" pageId="0" pageNumber="129">comb. nov.</taxonomicNameLabel>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="description">
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">Figs 2A, B</figureCitation>
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">, 3C, E</figureCitation>
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">, 4A, B</figureCitation>
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">, 6A, B, E, F, H, I</figureCitation>
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">, 9A</figureCitation>
|
||||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributions of A Iratiquedius amabilis (Smetana) (circles), I. mutator (Smetana) (squares) B I. prostans (Horn) (black - specimens examined, white - literature records) C I. seriatus (Horn) (circles), I. uncifer sp. nov. (squares) D Paraquedius puncticeps (Horn) (circles), P. marginicollis sp. nov. (squares) E Quediellus debilis (Horn) (circles) (black - specimens examined, white - literature records)." figureDoi="10.3897/zookeys.1134.87853.figure11" httpUri="https://binary.pensoft.net/fig/779769" pageId="0" pageNumber="129">, 11A (map)</figureCitation>
|
||||
<subSubSection id="476783BBD69C2D83913006507CFE49E7" pageId="0" pageNumber="129" type="description">
|
||||
<paragraph id="3EF0A735EED1CB8BEFC9F1FDF3AC46A5" pageId="0" pageNumber="129">
|
||||
<figureCitation id="52358EF2AE2E7A37B0BDD737B6BC59BA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">Figs 2A, B</figureCitation>
|
||||
<figureCitation id="65973667C78003AAE99E541D8050E013" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A-D dorsal head A, B showing confluence of inner eye margin and supra-antennal carina (arrow) E, F abdominal tergites A, D Iratiquedius seriatus (Horn) B Quedius (Raphirus) probus (Casey) C, E I. amabilis (Smetana) F I. prostans (Horn). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.1 mm (A, B); 0.5 mm (C-F)." figureDoi="10.3897/zookeys.1134.87853.figure3" httpUri="https://binary.pensoft.net/fig/779761" pageId="0" pageNumber="129">, 3C, E</figureCitation>
|
||||
<figureCitation id="2C5CD613DE3B6D62E15D05686E95AEA3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">, 4A, B</figureCitation>
|
||||
<figureCitation id="0EFC60A91A325D076D1AA92BA33DD658" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">, 6A, B, E, F, H, I</figureCitation>
|
||||
<figureCitation id="663C7438E721DAAE9EA5A6F26CACC139" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">, 9A</figureCitation>
|
||||
<figureCitation id="C5028F91BFE6BFBFF3B4D3709057F962" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributions of A Iratiquedius amabilis (Smetana) (circles), I. mutator (Smetana) (squares) B I. prostans (Horn) (black - specimens examined, white - literature records) C I. seriatus (Horn) (circles), I. uncifer sp. nov. (squares) D Paraquedius puncticeps (Horn) (circles), P. marginicollis sp. nov. (squares) E Quediellus debilis (Horn) (circles) (black - specimens examined, white - literature records)." figureDoi="10.3897/zookeys.1134.87853.figure11" httpUri="https://binary.pensoft.net/fig/779769" pageId="0" pageNumber="129">, 11A (map)</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="reference_group">
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) amabilis</taxonomicName>
|
||||
<subSubSection id="FA9EF7B69F013121C4517689E62281F9" pageId="0" pageNumber="129" type="reference_group">
|
||||
<paragraph id="8036B50259A5538DF296A2557290BD09" pageId="0" pageNumber="129">
|
||||
<taxonomicName id="FD20AFBAE5ACA3282B474E335CCB778E" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) amabilis</taxonomicName>
|
||||
Smetana, 1971: 205.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) amabilis</taxonomicName>
|
||||
<paragraph id="0D302C45947FBC0FBF413FB1E19FE038" pageId="0" pageNumber="129">
|
||||
<taxonomicName id="D70B9766E6C844D9D446D83136B5804B" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) amabilis</taxonomicName>
|
||||
:
|
||||
<bibRefCitation author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="129" pagination="95 - 104" refId="B12" refString="Smetana, A, 1990. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6. Coleopterists Bulletin 44: 95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana 1990</bibRefCitation>
|
||||
<bibRefCitation id="FEE0010F3858F737733DCC4AAC8DC831" author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="129" pagination="95 - 104" refId="B12" refString="Smetana, A, 1990. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6. Coleopterists Bulletin 44: 95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana 1990</bibRefCitation>
|
||||
(distributional records).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="comb. nov.">Quedius amabilis</taxonomicName>
|
||||
<paragraph id="0E64F598A6528ED9688C18EFC85B8DBA" pageId="0" pageNumber="129">
|
||||
<taxonomicName id="492F2DD8A394A6B8253D2EC28DB4CAA7" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="comb. nov.">Quedius amabilis</taxonomicName>
|
||||
:
|
||||
<bibRefCitation DOI="https://doi.org/10.1111/syen.12468" author="Brunke, AJ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="396 - 421" refId="B4" refString="Brunke, AJ, Hansen, AK, Salnitska, M, Kypke, JL, Predeus, AV, Escalona, HE, Chapados, JT, Eyres, J, Richter, R, Smetana, A, Ślipinski, SA, Zwick, A, Hajek, J, Leschen, RAB, Solodovnikov, A, Dettman, JR, 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46 (2): 396 - 421, DOI: https://doi.org/10.1111/syen.12468" title="The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics." url="https://doi.org/10.1111/syen.12468" volume="46" year="2021">Brunke et al. 2021</bibRefCitation>
|
||||
<bibRefCitation id="06CA845F5A1F86983726FBE2A2758909" DOI="https://doi.org/10.1111/syen.12468" author="Brunke, AJ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="396 - 421" refId="B4" refString="Brunke, AJ, Hansen, AK, Salnitska, M, Kypke, JL, Predeus, AV, Escalona, HE, Chapados, JT, Eyres, J, Richter, R, Smetana, A, Ślipinski, SA, Zwick, A, Hajek, J, Leschen, RAB, Solodovnikov, A, Dettman, JR, 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46 (2): 396 - 421, DOI: https://doi.org/10.1111/syen.12468" title="The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics." url="https://doi.org/10.1111/syen.12468" volume="46" year="2021">Brunke et al. 2021</bibRefCitation>
|
||||
(member of 'Clade
|
||||
<normalizedToken originalValue="A’">A'</normalizedToken>
|
||||
<normalizedToken id="4D41EC24315E740DFA9ACE099419D1BB" originalValue="A’">A'</normalizedToken>
|
||||
, non-
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raphirus" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus" status="comb. nov.">Raphirus</taxonomicName>
|
||||
<taxonomicName id="9D0503A52CF23DBBEAF3F116BD4EC00A" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raphirus" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus" status="comb. nov.">Raphirus</taxonomicName>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="129">Type locality.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">Near Strawberry, El Dorado County, California, United States.</paragraph>
|
||||
<subSubSection id="C7E3DE799BE2891C0DD59B0557F8865F" pageId="0" pageNumber="129" type="type locality">
|
||||
<paragraph id="0B4AAD0C52E41CF2061E6ACFF6E7B548" pageId="0" pageNumber="129">Type locality.</paragraph>
|
||||
<paragraph id="411DBF1DE96D2D5B0509FD5BB9618D13" pageId="0" pageNumber="129">Near Strawberry, El Dorado County, California, United States.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="type material">
|
||||
<paragraph pageId="0" pageNumber="129">Type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Holotype</emphasis>
|
||||
<subSubSection id="CA2DA70905735A22BF863DE1E143D11D" pageId="0" pageNumber="129" type="type material">
|
||||
<paragraph id="861B2011E95D925F108533B8D5054ABB" pageId="0" pageNumber="129">Type material.</paragraph>
|
||||
<paragraph id="D90D63DD550034C809BDF8030F4C0DB8" pageId="0" pageNumber="129">
|
||||
<emphasis id="51ECF8659C09F2646239B81A973124E1" bold="true" italics="true" pageId="0" pageNumber="129">Holotype</emphasis>
|
||||
(male, CNC): N.r. Strawberry, Eldorado Co. Cal., 16-17.61 [handwritten label] / #67, 45 [illegible, 'mi
|
||||
<normalizedToken originalValue="camp’">camp'</normalizedToken>
|
||||
<normalizedToken id="BF8573B6BDEDD8A3FC8C840C707BBCF6" originalValue="camp’">camp'</normalizedToken>
|
||||
?] [handwritten label] /R. Schuster Collector [printed label] / HOLOTYPE
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Quedius amabilis</emphasis>
|
||||
<taxonomicName id="5EB4FF299FD9622FBEB082675C179991" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||||
<emphasis id="002D5D710194BAA850F9E14AE90D9297" italics="true" pageId="0" pageNumber="129">Quedius amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
Smetana 1968, CNC No. 10876 [red printed label] / CNC935809 [identifier label].
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">The aedeagus of the male holotype was never fully dissected by A. Smetana and was glued to the point with the specimen. For the present study, the aedeagus was relaxed, photographed and placed in glycerin within a genitalia vial.</paragraph>
|
||||
<paragraph id="95AA3975820B42459A626A5901429A02" pageId="0" pageNumber="129">The aedeagus of the male holotype was never fully dissected by A. Smetana and was glued to the point with the specimen. For the present study, the aedeagus was relaxed, photographed and placed in glycerin within a genitalia vial.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="type material">
|
||||
<paragraph pageId="0" pageNumber="129">Non-type material.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">United States: California</emphasis>
|
||||
<subSubSection id="5DF161F13A0A1456CB433733195B11FF" pageId="0" pageNumber="129" type="type material">
|
||||
<paragraph id="E2E52F8330E111C4B8746AD2F33A108D" pageId="0" pageNumber="129">Non-type material.</paragraph>
|
||||
<paragraph id="506956613D95D31B9B7968F0D484B7C0" pageId="0" pageNumber="129">
|
||||
<emphasis id="7B33B0B8729D396B57C47CC1D4974F70" bold="true" pageId="0" pageNumber="129">United States: California</emphasis>
|
||||
: Sierra Co.: 14 mi E Sierra City, Yuba Pass, 6,700' [2,042 m], 26.VI.1976, L. & N. Herman (1 male, CNC); same except 26-28.VI.1976 (2 females, CNC).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius amabilis</emphasis>
|
||||
<subSubSection id="E224D7719A8A5866059E54A12098B128" pageId="0" pageNumber="129" type="diagnosis">
|
||||
<paragraph id="38E599A38D928CF19707FE5C2B3C0FC0" pageId="0" pageNumber="129">Diagnosis.</paragraph>
|
||||
<paragraph id="C4993381C5792A3E53283FF0DA4FC4F3" pageId="0" pageNumber="129">
|
||||
<taxonomicName id="0BF21FB72ED3B5631EF82D9C6783AA2C" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||||
<emphasis id="54CEADE6186D89DE39F2F5083B4C1572" italics="true" pageId="0" pageNumber="129">Iratiquedius amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
may be recognized within the genus by a combination of the evenly punctate elytra, the lack of golden setae or impressions at the base of the abdominal tergites and the distinctly impressed micropunctures on the anterior angles of the pronotum. The species most closely resembles
|
||||
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
<taxonomicName id="B1DF2D2D3E13A11173DAEE74ABCFCA27" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis id="05442F0F85C8798E6282C6D893738D7F" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
and can be distinguished from it by the distinct micropunctures of the anterior angles of the pronotum, the shorter apex of the median lobe in lateral view in males or broader, more densely setose apex of female tergite X.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="redescription">
|
||||
<paragraph pageId="0" pageNumber="129">Redescription.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="32797BFA6715263A93CE066C327FEBB5" pageId="0" pageNumber="129" type="redescription">
|
||||
<paragraph id="55EC1334A9500DD18150A291CBFBC233" pageId="0" pageNumber="129">Redescription.</paragraph>
|
||||
<paragraph id="78833DB2CBF96EE65CAA7C6E11A20DD6" pageId="0" pageNumber="129">
|
||||
Measurements ♂ (
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">n</emphasis>
|
||||
<emphasis id="B5EF2211FC0686C9CDDF96CB1FB3D216" italics="true" pageId="0" pageNumber="129">n</emphasis>
|
||||
= 2): HW/HL 1.21-1.22; PW/PL 1.29-1.32; EW/EL 1.28-1.31; ESut/PL 0.83; PW/HW 1.16-1.33; forebody length 3.3-3.4 mm.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<paragraph id="FEC7ED768CBB7A14E5E9930263CB5398" pageId="0" pageNumber="129">
|
||||
Measurements ♀ (
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">n</emphasis>
|
||||
<emphasis id="F02BA5AF0895918A7F2BFB234D169A6D" italics="true" pageId="0" pageNumber="129">n</emphasis>
|
||||
= 2): HW/HL 1.18-1.21; PW/PL 1.17-1.24; EW/EL 1.45-1.59; ESut/PL 0.60-0.69; PW/HW 1.22-1.27; forebody length 2.9-3.0 mm.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">Dark brown with pronotum becoming slightly to distinctly paler toward margin, elytra variably paler at base, sides and apically, in some individuals only scutellar area darkened, pale areas varying from brownish red to yellowish red or brownish yellow; antennomeres 1-3 sometimes slightly darker than remaining segments, which are brownish yellow to dark brown; pro- and mesofemora paler, yellowish brown; abdominal tergites narrowly paler apically, sternites with broader pale area at apex.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<paragraph id="D83E452677CCD398F59509C96ED66226" pageId="0" pageNumber="129">Dark brown with pronotum becoming slightly to distinctly paler toward margin, elytra variably paler at base, sides and apically, in some individuals only scutellar area darkened, pale areas varying from brownish red to yellowish red or brownish yellow; antennomeres 1-3 sometimes slightly darker than remaining segments, which are brownish yellow to dark brown; pro- and mesofemora paler, yellowish brown; abdominal tergites narrowly paler apically, sternites with broader pale area at apex.</paragraph>
|
||||
<paragraph id="0465457535C2487606DD4F8C5B43460A" pageId="0" pageNumber="129">
|
||||
With distinct macropterous and brachypterous morphotypes (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">2A, B</figureCitation>
|
||||
<figureCitation id="7758C5ABB20F1960809BE2507425BDD7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">2A, B</figureCitation>
|
||||
). Head distinctly transverse, temples extremely short, immediately converging to neck posteriad of eyes; disc of head with microsculpture intermediate between transverse waves and transverse meshes, meshes becoming tighter on frons; posterior frontal puncture located at posterior fourth of eye; interocular punctures present or absent; labrum short, strongly transverse, forming two lobes; area between anterior frontal punctures with broad, transverse and shallow impression, encompassing interocular punctures, if present; antennomeres 1-5 (macropterous) or 1-4 (brachypterous) clearly elongate, segments becoming shorter toward apex, 8-10 weakly to moderately transverse; pronotum strongly (macropterous) to moderately (brachypterous) transverse; disc with microsculpture of transverse waves, often changing direction, becoming isodiametric meshes on anterior angles, anterior angles with distinct, shallow micropunctures, strongly (macropterous) (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">4A</figureCitation>
|
||||
<figureCitation id="0E5E4997A51CA5849F3556D58D91CDF0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">4A</figureCitation>
|
||||
) to moderately (brachypterous) impressed (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">4B</figureCitation>
|
||||
<figureCitation id="EE58A7A99366997FA5DDA13D86B323DC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">4B</figureCitation>
|
||||
); elytra moderately (macropterous), to strongly transverse (brachypterous), at suture moderately (macropterous) to markedly (brachypterous) shorter than pronotum at midline, disc without microsculpture, punctation sparse, most punctures separated by about one puncture width (brachypterous) to slightly denser, with several punctures touching each other laterally (macropterous); abdomen with dense microsculpture of transverse waves, punctures slightly (macropterous) to distinctly (brachypterous) denser on bases of segments.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129" start="Figure 4" startId="F4">
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">Figure 4.</emphasis>
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">A-D</emphasis>
|
||||
<caption id="377F996F0138C91ADA5C6CF54A58E70C" doi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129" start="Figure 4" startId="F4">
|
||||
<paragraph id="CA701B40F5E7109A450E1D7EEB9AB66E" pageId="0" pageNumber="129">
|
||||
<emphasis id="C88E768FD0747F4F716F8A7F25008FDF" bold="true" pageId="0" pageNumber="129">Figure 4.</emphasis>
|
||||
<emphasis id="220B2334ABE2953EB37C72D6514760E0" bold="true" pageId="0" pageNumber="129">A-D</emphasis>
|
||||
anterior angle of the pronotum
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">A, B</emphasis>
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius amabilis</emphasis>
|
||||
<emphasis id="8A86474F3EF3470E9FB3032125DD6C16" bold="true" pageId="0" pageNumber="129">A, B</emphasis>
|
||||
<taxonomicName id="0AA6697518B302A3ED47EAD5F0EBB963" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||||
<emphasis id="C111CA93D422CE94FC4D3BE70CB7C780" italics="true" pageId="0" pageNumber="129">Iratiquedius amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
(Smetana)
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">C, D</emphasis>
|
||||
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
<emphasis id="09C4933B325DC81A4B2F16EB53A513EC" bold="true" pageId="0" pageNumber="129">C, D</emphasis>
|
||||
<taxonomicName id="2FCE1BC0BFC2B19912CD0CDD83B0D865" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis id="B30F87F1D9964C409FD617214E96E0AB" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
(Smetana)
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">A, C</emphasis>
|
||||
<emphasis id="0647F50CAAB555D67F1904EB01C818F3" bold="true" pageId="0" pageNumber="129">A, C</emphasis>
|
||||
macropterous morphotype
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">B, D</emphasis>
|
||||
<emphasis id="458DAF9E6F6DB4AD3BFC2B6C88A3EB6C" bold="true" pageId="0" pageNumber="129">B, D</emphasis>
|
||||
brachypterous morphotype. Scale bars: 0.1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">Male.</emphasis>
|
||||
<paragraph id="93B1667DC95EC315390E4BCCB08DD51A" pageId="0" pageNumber="129">
|
||||
<emphasis id="B5CB481D31E0302525E047E41549B544" bold="true" pageId="0" pageNumber="129">Male.</emphasis>
|
||||
Sternite VIII with distinct, wide V-shaped emargination; tergite X triangular, with short, rounded apex; sternite IX overall narrow, with long asymmetrical basal part and narrow, minutely emarginate apex; median lobe in ventral view with short tooth, subparallel, with slight expansion subapically, before converging to narrow, truncate apex bearing slight to distinct emargination (Fig.
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6E, F</figureCitation>
|
||||
<figureCitation id="F77AD6BE2AA632E0E228CCB34899DB82" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6E, F</figureCitation>
|
||||
); median lobe in lateral view arcuate ventrad, with moderately long apical area, wide tooth and narrow, rounded apex (Fig.
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6A, B</figureCitation>
|
||||
<figureCitation id="7EF404E5145D67CBB7743148B39E704F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6A, B</figureCitation>
|
||||
); paramere subparallel, slightly expanded subapically, converging to moderately narrow, rounded apex, with or without small emargination, peg setae arranged in dense marginal row (Fig.
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6H, I</figureCitation>
|
||||
<figureCitation id="764FBF18252FAF648D626D05E08CA2D9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6H, I</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">Female.</emphasis>
|
||||
<paragraph id="2606360D09D7D3D49C2C576077F526A3" pageId="0" pageNumber="129">
|
||||
<emphasis id="C621CC83A72866891E75B79D9C3A99AE" bold="true" pageId="0" pageNumber="129">Female.</emphasis>
|
||||
Tergite X narrowly triangular, with apex slightly attenuate, with dense marginal setae (Fig.
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">9A</figureCitation>
|
||||
<figureCitation id="5C94E4648539F896E891473154EE0236" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">9A</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<emphasis bold="true" pageId="0" pageNumber="129">United States</emphasis>
|
||||
<subSubSection id="78B7E708A89E8AF898CCADC06B2AB13B" pageId="0" pageNumber="129" type="distribution">
|
||||
<paragraph id="FCE2CC950155D628495AA9E26F2D7D61" pageId="0" pageNumber="129">Distribution.</paragraph>
|
||||
<paragraph id="709716E92250D2914A010E279EC177DC" pageId="0" pageNumber="129">
|
||||
<emphasis id="86D1AC8FA7CEBDE26C9AF6160B3FF381" bold="true" pageId="0" pageNumber="129">United States</emphasis>
|
||||
: CA.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">This species is known only from two rather close localities in the Sierra Nevada of California.</paragraph>
|
||||
<paragraph id="9D40408D8DE8B33DDCFD9AFAAD2AF421" pageId="0" pageNumber="129">This species is known only from two rather close localities in the Sierra Nevada of California.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="bionomics">
|
||||
<paragraph pageId="0" pageNumber="129">Bionomics.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<subSubSection id="FAACFC33BBF1CEC952027CD90ABFE2C4" pageId="0" pageNumber="129" type="bionomics">
|
||||
<paragraph id="7BB8F02834B0BF2AA29A27DCB5EFB187" pageId="0" pageNumber="129">Bionomics.</paragraph>
|
||||
<paragraph id="2EBA6A48AE7D99D76F36BB032DA3700D" pageId="0" pageNumber="129">
|
||||
Nothing specific is known about this
|
||||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||||
<normalizedToken id="5517C2E867DCD431B58A41894CF27966" originalValue="species’">species'</normalizedToken>
|
||||
microhabitat preferences, though it probably lives in moss along the margins of springs and spring-fed creeks.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="129" type="comments">
|
||||
<paragraph pageId="0" pageNumber="129">Comments.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<bibRefCitation DOI="https://doi.org/10.4039/entm10379fv" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="129" pagination="1 - 303" refId="B10" refString="Smetana, A, 1971. Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 103 (79): 1 - 303, DOI: https://doi.org/10.4039/entm10379fv" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." url="https://doi.org/10.4039/entm10379fv" volume="103" year="1971">Smetana (1971)</bibRefCitation>
|
||||
<subSubSection id="186EAE36D1BEC1649CEE78F49D41AFDB" pageId="0" pageNumber="129" type="comments">
|
||||
<paragraph id="FCADB6A6FF6BAF29D538182D8CC6FE0E" pageId="0" pageNumber="129">Comments.</paragraph>
|
||||
<paragraph id="8F7495AD4C857BDC1A133C00D2FE263A" pageId="0" pageNumber="129">
|
||||
<bibRefCitation id="E3E4C2B9529E1514C3E77C1802A0E872" DOI="https://doi.org/10.4039/entm10379fv" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="129" pagination="1 - 303" refId="B10" refString="Smetana, A, 1971. Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 103 (79): 1 - 303, DOI: https://doi.org/10.4039/entm10379fv" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." url="https://doi.org/10.4039/entm10379fv" volume="103" year="1971">Smetana (1971)</bibRefCitation>
|
||||
described
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Quedius amabilis</emphasis>
|
||||
<taxonomicName id="FD7FF1385C91EA1E588C63AF7734B43D" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||||
<emphasis id="480724F790EF27C6EB99A4651A5F4AB3" italics="true" pageId="0" pageNumber="129">Quedius amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName family="Staphylinidae" kingdom="Animalia" lsidName="Q. mutator" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Q. mutator</emphasis>
|
||||
<taxonomicName id="4A6C3813952C1489B9001D90265CF0D3" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Q. mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator">
|
||||
<emphasis id="3F34AF33831128439FB874AB26FD0767" italics="true" pageId="0" pageNumber="129">Q. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
as the only members of the Amabilis group of
|
||||
<taxonomicName authority="" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus)" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="subGenus" subGenus="Raphirus">Quedius (Raphirus)</taxonomicName>
|
||||
<taxonomicName id="679360D6122CA3A9AC761EFE916128B1" authority="" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus)" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="subGenus" subGenus="Raphirus">Quedius (Raphirus)</taxonomicName>
|
||||
. The former was known from one male and one female, both macropterous and collected from the Sierra Nevada, while the latter was only known from a single, brachypterous female collected at a different locality in the northern Coast Range mountains to the west. In addition to elytral and wing size,
|
||||
<bibRefCitation DOI="https://doi.org/10.4039/entm10379fv" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="129" pagination="1 - 303" refId="B10" refString="Smetana, A, 1971. Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 103 (79): 1 - 303, DOI: https://doi.org/10.4039/entm10379fv" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." url="https://doi.org/10.4039/entm10379fv" volume="103" year="1971">Smetana (1971)</bibRefCitation>
|
||||
<bibRefCitation id="CF92D6478C91CF786A7E015569DA6E21" DOI="https://doi.org/10.4039/entm10379fv" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="129" pagination="1 - 303" refId="B10" refString="Smetana, A, 1971. Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 103 (79): 1 - 303, DOI: https://doi.org/10.4039/entm10379fv" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." url="https://doi.org/10.4039/entm10379fv" volume="103" year="1971">Smetana (1971)</bibRefCitation>
|
||||
cited differences in the punctation of the abdominal tergites, shape of the antennomeres, elytral punctation and pronotum shape. Nothing was reported for many years until
|
||||
<bibRefCitation author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="129" pagination="95 - 104" refId="B12" refString="Smetana, A, 1990. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6. Coleopterists Bulletin 44: 95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana (1990)</bibRefCitation>
|
||||
<bibRefCitation id="B759BC628F6142115F331AA388EFFDC4" author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="129" pagination="95 - 104" refId="B12" refString="Smetana, A, 1990. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6. Coleopterists Bulletin 44: 95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana (1990)</bibRefCitation>
|
||||
, reported two brachypterous female specimens as
|
||||
<taxonomicName family="Staphylinidae" kingdom="Animalia" lsidName="Q. mutator" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Q. mutator</emphasis>
|
||||
<taxonomicName id="455F8FD30A222C807E90381BE4067996" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Q. mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator">
|
||||
<emphasis id="4E39789F233C48448365B29B0243BF38" italics="true" pageId="0" pageNumber="129">Q. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
and four macropterous males (one studied here) as
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" family="Staphylinidae" kingdom="Animalia" lsidName="Q. amabilis" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Q. amabilis</emphasis>
|
||||
<taxonomicName id="CFBDF8DAF427EF43E65512FD19695A0B" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Q. amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||||
<emphasis id="033898E6DC5D69381B62F1F25067D557" italics="true" pageId="0" pageNumber="129">Q. amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
, both collected on Yuba Pass, California (Sierra Nevada) on different dates. He considered the possibility that
|
||||
<taxonomicName family="Staphylinidae" kingdom="Animalia" lsidName="Q. mutator" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Q. mutator</emphasis>
|
||||
<taxonomicName id="4B0C185396FC78D3BF09335C47042EF2" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Q. mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator">
|
||||
<emphasis id="DE44304B2F1D393071893BE3B6214B01" italics="true" pageId="0" pageNumber="129">Q. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
may simply be a brachypterous morph of
|
||||
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" family="Staphylinidae" kingdom="Animalia" lsidName="Q. amabilis" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">Q. amabilis</emphasis>
|
||||
<taxonomicName id="59C06230A50F18CC60FD1620CC6B349F" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Q. amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis">
|
||||
<emphasis id="29513226A1DAEBBA65C0B21728BE2A92" italics="true" pageId="0" pageNumber="129">Q. amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<paragraph id="1A19819CD06A873BBE3BC840A829F673" pageId="0" pageNumber="129">
|
||||
A macropterous male from Yuba Pass was dissected and its aedeagus closely resembles that of the holotype of
|
||||
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
<taxonomicName id="70D0526014CAFA0CD0BC49A8EA24A11F" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis id="9A337E2AB3C3B12F6E55C17C26FD778B" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6A, B</figureCitation>
|
||||
<figureCitation id="40430A583FA038472C47DEA4CEA75252" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6A, B</figureCitation>
|
||||
). The apical emargination of the median lobe in ventral view is less pronounced in the holotype compared to the non-type, and the paramere is slightly emarginate in the holotype, but these differences are considered to be intraspecific variation. In lateral view, the two specimens are nearly identical. The specimens are also similar externally and I agree with
|
||||
<bibRefCitation author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="129" pagination="95 - 104" refId="B12" refString="Smetana, A, 1990. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6. Coleopterists Bulletin 44: 95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana (1990)</bibRefCitation>
|
||||
<bibRefCitation id="FAF69C27343763118CD0DA9547A7F044" author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="129" pagination="95 - 104" refId="B12" refString="Smetana, A, 1990. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6. Coleopterists Bulletin 44: 95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana (1990)</bibRefCitation>
|
||||
that they are conspecific. The two brachypterous females differ from the macropterous male in all other characters (antennae, pronotum, elytra, punctation of abdominal tergites) previously used to differentiate
|
||||
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
<taxonomicName id="85DD13D55E19C7B131358F0E505EEC8E" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis id="04853885B30D13C386EEB422BE1001CB" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
<taxonomicName id="7B6BFDC0A968E14DA471EF4210E866A8" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis id="BD43A026F11E2DC1BA589F57C9E64430" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
. One male and one female from the Yuba Pass series were sequenced and their half-length (325 bp) barcodes were only 0.34% divergent (Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Neighbor-joining trees of CO 1 barcode sequences for species of Iratiquedius, Paraquedius, and Quediellus, calculated under the Kimura- 2 model for pairwise distance. White circles indicate macropterous ' debilis' morphotypes of Q. debilis, all other Q. debilis are of brachypterous ' nanulus' morphotypes. Scale bar equivalent to 2 % divergence. Sequence length, including number of ambiguous base pairs (N's), is given at the end of each sample name." figureDoi="10.3897/zookeys.1134.87853.figure10" httpUri="https://binary.pensoft.net/fig/779768" pageId="0" pageNumber="129">10</figureCitation>
|
||||
<figureCitation id="81E05D8037E05A57CEAC21170262AC4A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Neighbor-joining trees of CO 1 barcode sequences for species of Iratiquedius, Paraquedius, and Quediellus, calculated under the Kimura- 2 model for pairwise distance. White circles indicate macropterous ' debilis' morphotypes of Q. debilis, all other Q. debilis are of brachypterous ' nanulus' morphotypes. Scale bar equivalent to 2 % divergence. Sequence length, including number of ambiguous base pairs (N's), is given at the end of each sample name." figureDoi="10.3897/zookeys.1134.87853.figure10" httpUri="https://binary.pensoft.net/fig/779768" pageId="0" pageNumber="129">10</figureCitation>
|
||||
). This result suggests that the two morphotypes collected together on Yuba Pass are conspecific, correspond to
|
||||
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
<taxonomicName id="42FB3E32BDE1986D296B415FFAA3626F" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis id="658D6D3A5BBD0974EE60AE6EB9D4A954" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
and they are here treated as such. The existence of a macropterous female (the unstudied allotype), if it is indeed a female, indicates that females of
|
||||
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
<taxonomicName id="CC53490A140D48C165F138EA20FD741F" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis id="9637E3323DED6337E467281AE7313794" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
are wing dimorphic.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="129">
|
||||
<paragraph id="58408D4FA6C399458514298543EBC202" pageId="0" pageNumber="129">
|
||||
However, the female holotype of
|
||||
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
<taxonomicName id="A42563A44E85D0CDE61A2698260DC806" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis id="BF4449C4E0C27F598BEBC681183DE749" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
differs from the Yuba Pass females by the even shorter elytra, shorter antennomeres 4-10 and the distinctly different apex of tergite X (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">2D</figureCitation>
|
||||
<figureCitation id="0259528EBAC6CBBC1CADB8E92C4C399D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">2D</figureCitation>
|
||||
). This suggests that
|
||||
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
<taxonomicName id="664165F32775ACB826925807FEDE7001" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis id="11D4BDDD8F42358DB1B46AE7E68B3C29" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
is a valid species and that macropterous and brachypterous morphotypes confusingly exist in both species. All characters used previously to differentiate the two species are here considered to be associated with wing-dimorphism. Two fully winged males from the Central Valley of California were recently found in the FMNH collection (see below) and differ both externally (micropunctation of pronotum) (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">2C</figureCitation>
|
||||
<figureCitation id="D8869B717E18B2F31D3E45D63A5C6063" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">2C</figureCitation>
|
||||
) and in male genitalia from the known males of
|
||||
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
<taxonomicName id="E0D9598BD25387DEFEA3998CE76F2C38" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
|
||||
<emphasis id="50A34DD9B92A723945E6BC6CA5A1B553" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
|
||||
</taxonomicName>
|
||||
. One specimen was sequenced and its partial barcode was found to be 11.3% different from the Yuba Pass specimens (Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Neighbor-joining trees of CO 1 barcode sequences for species of Iratiquedius, Paraquedius, and Quediellus, calculated under the Kimura- 2 model for pairwise distance. White circles indicate macropterous ' debilis' morphotypes of Q. debilis, all other Q. debilis are of brachypterous ' nanulus' morphotypes. Scale bar equivalent to 2 % divergence. Sequence length, including number of ambiguous base pairs (N's), is given at the end of each sample name." figureDoi="10.3897/zookeys.1134.87853.figure10" httpUri="https://binary.pensoft.net/fig/779768" pageId="0" pageNumber="129">10</figureCitation>
|
||||
<figureCitation id="1669024C42E10D2A470F881032A9AA43" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Neighbor-joining trees of CO 1 barcode sequences for species of Iratiquedius, Paraquedius, and Quediellus, calculated under the Kimura- 2 model for pairwise distance. White circles indicate macropterous ' debilis' morphotypes of Q. debilis, all other Q. debilis are of brachypterous ' nanulus' morphotypes. Scale bar equivalent to 2 % divergence. Sequence length, including number of ambiguous base pairs (N's), is given at the end of each sample name." figureDoi="10.3897/zookeys.1134.87853.figure10" httpUri="https://binary.pensoft.net/fig/779768" pageId="0" pageNumber="129">10</figureCitation>
|
||||
). Instead, these two males more closely resemble the female holotype of
|
||||
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
<taxonomicName id="434B64B4369F9D0763E346D3E78EA883" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis id="1E09DB572B7264C9B10D6E0324904BC8" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
in external morphology, and the concept of
|
||||
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
<taxonomicName id="BCCC1F0473B49339164BD8DFBDA32486" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
|
||||
<emphasis id="B591428BBF4AF611F49B1192C93532F5" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
|
||||
</taxonomicName>
|
||||
is expanded below.
|
||||
</paragraph>
|
||||
|
|
|
|||
|
|
@ -1,154 +1,154 @@
|
|||
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<treatment ID-GBIF-Taxon="162174127" LSID="urn:lsid:zoobank.org:act:0A664B6A-03A5-4A15-BBEE-06D754270C97" httpUri="http://treatment.plazi.org/id/E38B63EB717F5ADD8DC2A322FB21927D" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName LSID="E38B63EB-717F-5ADD-8DC2-A322FB21927D" authority="Chang & Li, 2020" authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera thamsangensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thamsangensis" status="sp. nov.">Leclercera thamsangensis</taxonomicName>
|
||||
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">Figs 3</figureCitation>
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">, 4</figureCitation>
|
||||
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Cheliceral retromargin, posterior view. A Leclercera dumuzhou sp. nov. B L. ekteenensis sp. nov. C L. thamkaewensis sp. nov. D L. yamaensis sp. nov. E L. thamsangensis sp. nov. F L. hponensis sp. nov. G L. lizi sp. nov. H L. mianqiu sp. nov. I L. selasihensis sp. nov. J L. yuanzhui sp. nov. K L. paiensis sp. nov. L L. zanggaensis sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure56" httpUri="https://binary.pensoft.net/fig/380998" pageId="0" pageNumber="1">, 56E</figureCitation>
|
||||
<figureCitation captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">, 58</figureCitation>
|
||||
<treatment id="E38B63EB717F5ADD8DC2A322FB21927D" ID-GBIF-Taxon="162174127" LSID="urn:lsid:zoobank.org:act:0A664B6A-03A5-4A15-BBEE-06D754270C97" httpUri="http://treatment.plazi.org/id/E38B63EB717F5ADD8DC2A322FB21927D" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Psilodercidae" scope_order="Araneae">
|
||||
<subSubSection id="FBD3F8237CA0A8BF749D3F27B013DAAF" pageId="0" pageNumber="1" type="nomenclature">
|
||||
<paragraph id="F54114646E51039336C12181FFD56470" pageId="0" pageNumber="1">
|
||||
<taxonomicName id="BA5AA19603F765360B8D290A83974C93" ID-CoL="98ZV6" LSID="E38B63EB-717F-5ADD-8DC2-A322FB21927D" authority="Chang & Li, 2020" authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera thamsangensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thamsangensis" status="sp. nov.">Leclercera thamsangensis</taxonomicName>
|
||||
<taxonomicNameLabel id="19537E42934E2E01271624FA73352869" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
||||
<figureCitation id="7D617C2D8E706285646E1E826601727A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">Figs 3</figureCitation>
|
||||
<figureCitation id="1A356DC590D4D6476B4A4D2C875D141C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">, 4</figureCitation>
|
||||
<figureCitation id="0AC7E10FE20BD7AD89084A954888C0D2" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Cheliceral retromargin, posterior view. A Leclercera dumuzhou sp. nov. B L. ekteenensis sp. nov. C L. thamkaewensis sp. nov. D L. yamaensis sp. nov. E L. thamsangensis sp. nov. F L. hponensis sp. nov. G L. lizi sp. nov. H L. mianqiu sp. nov. I L. selasihensis sp. nov. J L. yuanzhui sp. nov. K L. paiensis sp. nov. L L. zanggaensis sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure56" httpUri="https://binary.pensoft.net/fig/380998" pageId="0" pageNumber="1">, 56E</figureCitation>
|
||||
<figureCitation id="1C9B37DDF7FCA8297CFDC5019E7068EC" captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">, 58</figureCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="types">
|
||||
<paragraph pageId="0" pageNumber="1">Types.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
<subSubSection id="66BCC1643624DFB98B380DFF86706559" pageId="0" pageNumber="1" type="types">
|
||||
<paragraph id="6779AEA345F0FE72F3FBEFCD6C5E2E15" pageId="0" pageNumber="1">Types.</paragraph>
|
||||
<paragraph id="367DEF07C7067D494E1C0B0322638D88" pageId="0" pageNumber="1">
|
||||
<emphasis id="A4438CEC735DE38901EAEC420C107B5F" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||||
: ♂ (IZCAS), Thailand, Loei Province, Phu Kradueng District, Phan Nok Kao Subdistrict, Tham Wat Phu Sang One,
|
||||
<geoCoordinate degrees="16" direction="north" minutes="49.0620" orientation="latitude" precision="1" value="16.8177">16°49.0620'N</geoCoordinate>
|
||||
<geoCoordinate id="42682519D2D05215A09A19BE17B38FD1" degrees="16" direction="north" minutes="49.0620" orientation="latitude" precision="1" value="16.8177">16°49.0620'N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="101" direction="east" minutes="56.4330" orientation="longitude" precision="1" value="101.94055">101°56.4330'E</geoCoordinate>
|
||||
<geoCoordinate id="5BB9637401C43107961F87483DB85384" degrees="101" direction="east" minutes="56.4330" orientation="longitude" precision="1" value="101.94055">101°56.4330'E</geoCoordinate>
|
||||
, elevation ca 385 m, 29.XI.2016, H. Zhao leg.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
|
||||
<emphasis id="7AE57D8B98BF538D90473F22889B0CD7" bold="true" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
|
||||
: 1♀ (IZCAS), same data as holotype.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">The species name is an adjective referring to the type locality.</paragraph>
|
||||
<subSubSection id="19A96D71A3896798C9B665CE19D2109F" pageId="0" pageNumber="1" type="etymology">
|
||||
<paragraph id="3637F0B4FF0712E8C34DE3F24BDA77FF" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||||
<paragraph id="08608D0B9220D60F4C61A110186ED7E7" pageId="0" pageNumber="1">The species name is an adjective referring to the type locality.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="5B9EE1D2FB6C2C16FD2F1E9F5B9362D7" pageId="0" pageNumber="1" type="diagnosis">
|
||||
<paragraph id="EDB8A891C58E345055FB13CD07244778" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||||
<paragraph id="D08ECAD2AFA5A1194A818D0852322AD4" pageId="0" pageNumber="1">
|
||||
Males of
|
||||
<taxonomicName lsidName="L. thamsangensis" pageId="0" pageNumber="1" rank="species" species="thamsangensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">L. thamsangensis</emphasis>
|
||||
<taxonomicName id="A861E84FF2FB71891B575C9F6E0A623D" lsidName="L. thamsangensis" pageId="0" pageNumber="1" rank="species" species="thamsangensis">
|
||||
<emphasis id="21CB105BD7DA4FAF91452D666FD50EC3" italics="true" pageId="0" pageNumber="1">L. thamsangensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. can be distinguished from congeners by the presence of two spines on a small retrolateral protrusion (retrolateral apophysis) of the cymbium (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">4D</figureCitation>
|
||||
<figureCitation id="A86BBB57FF5BEF03A059F9F8EA2301F1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">4D</figureCitation>
|
||||
) (vs. the absence of a retrolateral apophysis with two spines on the cymbium), the conductor and embolus are not widely separated, appearing to be similar in length and width (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">4B</figureCitation>
|
||||
<figureCitation id="14F9804F0A71091F5AB9B8A49AC0CAFF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">4B</figureCitation>
|
||||
) (vs. conductor and embolus different in congeners); females can be differentiated from congeners by rectangular, sheet-like spermathecae (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3A</figureCitation>
|
||||
<figureCitation id="C04474E70815C3CD9AD8DB4171F840C9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3A</figureCitation>
|
||||
) (vs. absence of sheet-like spermathecae in congeners).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
||||
<taxonomicName authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera thamsangensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thamsangensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Leclercera thamsangensis</emphasis>
|
||||
<caption id="E4D1CA819ACAFBCA1B1B9597B19E0CFF" doi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
|
||||
<paragraph id="2F91AC95C388CAB7CD0330275FE9FB22" pageId="0" pageNumber="1">
|
||||
<emphasis id="3D471F0DF00A1ACB816B0DF2EBDD3A03" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
||||
<taxonomicName id="084B5969C6D080B69AF2BE7BD98CCB5E" authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera thamsangensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thamsangensis">
|
||||
<emphasis id="A761CD4AF725B9B0DB0F4A8CC3A8AB45" italics="true" pageId="0" pageNumber="1">Leclercera thamsangensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., male holotype and female paratype.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="58F5D6CF5C392A778B7A1F1AF6218CE2" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
Endogyne, dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="AFEA645A3C5D2AEC1EFFE08E33CA4B5A" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
female epigastric area, ventral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="064A2F75D08822382654CFC9987F869F" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
male habitus, dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="D2E542565A63DACA5F65E741F227200D" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
female habitus, dorsal view
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
<emphasis id="CE2B8211DD7DEA2F695A2F380AD42599" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||||
female habitus, ventral view. Abbreviation: SP = spermatheca.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||||
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Male</emphasis>
|
||||
<subSubSection id="9A06CC1B2056CEFA0DCBFB57582338C0" pageId="0" pageNumber="1" type="description">
|
||||
<paragraph id="E74C4C3DB08BF8342A14C4C75B9340A7" pageId="0" pageNumber="1">Description.</paragraph>
|
||||
<paragraph id="D9CC6AA12E1F43AEDE7C188EFD6F19F7" pageId="0" pageNumber="1">
|
||||
<emphasis id="DF0E43E8E71FFCFD0FCF883A0523F74D" bold="true" pageId="0" pageNumber="1">Male</emphasis>
|
||||
(Holotype). Total length 2.44; carapace 0.82 long, 0.84 wide; abdomen 1.62 long, 0.74 wide. Carapace round and brown, with three longitudinal dark brown bands, median band two times as wide as lateral band (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3C</figureCitation>
|
||||
<figureCitation id="C7393751FA0C2B015797890A9317ED4B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3C</figureCitation>
|
||||
). Chelicerae pale brown (Fig.
|
||||
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Cheliceral retromargin, posterior view. A Leclercera dumuzhou sp. nov. B L. ekteenensis sp. nov. C L. thamkaewensis sp. nov. D L. yamaensis sp. nov. E L. thamsangensis sp. nov. F L. hponensis sp. nov. G L. lizi sp. nov. H L. mianqiu sp. nov. I L. selasihensis sp. nov. J L. yuanzhui sp. nov. K L. paiensis sp. nov. L L. zanggaensis sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure56" httpUri="https://binary.pensoft.net/fig/380998" pageId="0" pageNumber="1">56E</figureCitation>
|
||||
<figureCitation id="B2C6B96DCAD906353629B42E2963B59F" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Cheliceral retromargin, posterior view. A Leclercera dumuzhou sp. nov. B L. ekteenensis sp. nov. C L. thamkaewensis sp. nov. D L. yamaensis sp. nov. E L. thamsangensis sp. nov. F L. hponensis sp. nov. G L. lizi sp. nov. H L. mianqiu sp. nov. I L. selasihensis sp. nov. J L. yuanzhui sp. nov. K L. paiensis sp. nov. L L. zanggaensis sp. nov. Abbreviations: PT = promargin teeth, RT = retromargin teeth." figureDoi="10.3897/zookeys.913.48650.figure56" httpUri="https://binary.pensoft.net/fig/380998" pageId="0" pageNumber="1">56E</figureCitation>
|
||||
). Clypeus brown. Endites brown. Labium dark brown basally. Sternum purplish, delimiting a light brown band medially. Abdomen elongated, dorsum with dark brown stripes laterally, delimiting a light brown band medially, antero-ventrally dark brown with elliptical patch, posterior part with indistinct dark and light brown pattern. Legs uniformly brown; measurements: I 9.76 (2.69, 0.38, 3.13, 2.53, 1.03), II 7.48 (2.09, 0.30, 2.34. 1.94, 0.81), III 5.56 (1.56, 0.30, 1.63, 1.41, 0.66), IV 8.74 (2.50, 0.30, 2.69. 2.31, 0.94). Palp (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">4A-D</figureCitation>
|
||||
<figureCitation id="52EF43F19369C630B919E8C562090C8D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">4A-D</figureCitation>
|
||||
): femur slender, 3.5 times longer than patella; patella not swollen; tibia 1.5 times shorter than femur; cymbium 2.5 times shorter than femur, with a small retrolateral apophysis bearing two spines basally, one spine half the length of the other; bulb light brown, pyriform, with embolus and conductor arising distally; embolus straight and thin, basally connected to conductor; conductor almost as long and wide as embolus (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">4B</figureCitation>
|
||||
<figureCitation id="0DF9ED8D3DCC9114CFD866B8EA762CDB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Leclercera thamsangensis sp. nov., male holotype. A Palp, ventral view B bulb, ventral view C palp, prolateral view D palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines." figureDoi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1">4B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Female</emphasis>
|
||||
<paragraph id="D3CFEEF083B33BD23797179ABD78D622" pageId="0" pageNumber="1">
|
||||
<emphasis id="D2901C4E83DBB67D3E9BE123DDF3F9DA" bold="true" pageId="0" pageNumber="1">Female</emphasis>
|
||||
(Paratype). General features and coloration similar to that of male (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3D, E</figureCitation>
|
||||
<figureCitation id="C5FF25179B5D741B9EB00AF90605A3B7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3D, E</figureCitation>
|
||||
). Measurements: total length 1.97; carapace 0.81 long, 0.84 wide; abdomen 1.16 long, 1.00 wide. Leg measurements: I 8.97 (2.44, 0.31, 2.91, 2.34, 0.97), II 6.72 (1.94, 0.31, 1.91, 1.78, 0.78), III 5.19 (1.41, 0.31, 1.50, 1.31, 0.66), IV 7.87 (2.19, 0.31, 2.50, 2.03, 0.84). Epigastric area (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3B</figureCitation>
|
||||
<figureCitation id="4FFB3DBD5628D9A2556677D4D756F22F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3B</figureCitation>
|
||||
): purplish crescent-shaped patch with a few setae, with random purplish patterns. Endogyne (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3A</figureCitation>
|
||||
<figureCitation id="A36DB19FB721C41EB033B03B1AC9FC24" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leclercera thamsangensis sp. nov., male holotype and female paratype. A Endogyne, dorsal view B female epigastric area, ventral view C male habitus, dorsal view D female habitus, dorsal view E female habitus, ventral view. Abbreviation: SP = spermatheca." figureDoi="10.3897/zookeys.913.48650.figure3" httpUri="https://binary.pensoft.net/fig/380945" pageId="0" pageNumber="1">3A</figureCitation>
|
||||
): pair of spermathecae slightly concave toward the posterior, bearing a rectangular sheet anteriorly.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<subSubSection id="CE4DBD97ED918225B9C3835586D65121" pageId="0" pageNumber="1" type="distribution">
|
||||
<paragraph id="1ABEFDAACAF43114D4F450A0552F59A2" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||||
<paragraph id="5246AAD6F14A4AC5995FE37DA9863D53" pageId="0" pageNumber="1">
|
||||
Known only from the type locality (Fig.
|
||||
<figureCitation captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">58</figureCitation>
|
||||
<figureCitation id="29BAE01CBF8054AAF45A27E66C991D3D" captionStart="Figure 58" captionStartId="F58" captionText="Figure 58. Distribution of new Leclercera species in Southeast Asia. 1 L. selasihensis sp. nov. 2 L. mianqiu sp. nov. 3 L. thamsangensis sp. nov. 4 L. yandou sp. nov. 5 L. thamkaewensis sp. nov. 6 L. xiangbabang sp. nov. 7 L. jianzuiyu sp. nov. 8 L. yamaensis sp. nov. 9 L. banensis sp. nov. 10 L. dumuzhou sp. nov. 11 L. suwanensis sp. nov. 12 L. maochong sp. nov. 13 L. yuanzhui sp. nov. 14 L. shanzi sp. nov. 15 L. duandai sp. nov. 16 L. hponensis sp. nov. 17 L. lizi sp. nov. 18 L. xiaodai sp. nov. 19 L. paiensis sp. nov. 20 L. yanjing sp. nov. 21 L. ekteenensis sp. nov. 22 L. zanggaensis sp. nov. 23 L. zhamensis sp. nov. 24 L. sanjiao sp. nov. 25 L. aniensis sp. nov. 26 L. renqinensis sp. nov. 27 L. shergylaensis sp. nov. 28 L. pulongensis sp. nov. 29 L. tudao sp. nov. 30 L. duibaensis sp. nov. 31 L. jiazhongensis sp. nov." figureDoi="10.3897/zookeys.913.48650.figure58" httpUri="https://binary.pensoft.net/fig/381000" pageId="0" pageNumber="1">58</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
|
||||
<paragraph pageId="0" pageNumber="1">
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
|
||||
<taxonomicName authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera thamsangensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thamsangensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="1">Leclercera thamsangensis</emphasis>
|
||||
<caption id="E0245EE4EF3B2B4E879F1E94DEEA888D" doi="10.3897/zookeys.913.48650.figure4" httpUri="https://binary.pensoft.net/fig/380946" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
|
||||
<paragraph id="BF57201A76870D8C28D8E5C1C7C96045" pageId="0" pageNumber="1">
|
||||
<emphasis id="BC96E6017E35B67DA0AD9FABBF208884" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
|
||||
<taxonomicName id="01A6905C9DD265CDAD780E6F94478FB1" authorityName="Chang & Li" authorityYear="2020" class="Arachnida" family="Psilodercidae" genus="Leclercera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leclercera thamsangensis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="thamsangensis">
|
||||
<emphasis id="D00B829A500E55833DC5D8D9BAD6D6B5" italics="true" pageId="0" pageNumber="1">Leclercera thamsangensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., male holotype.
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
<emphasis id="F80CA727DFD7902C9042451751306086" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||||
Palp, ventral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
<emphasis id="A1BD2D3F2C0CDF302C326F38BE2B1EFF" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||||
bulb, ventral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
<emphasis id="72DE8CCEAEDA203AA8F0A4D377B13CB9" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||||
palp, prolateral view
|
||||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
<emphasis id="C660D2D33AE26FBA5E32DA4CBC38905F" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||||
palp, retrolateral view. Abbreviations: CO = conductor, EM = embolus, RA = retrolateral apophysis, SN = spines.
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
|
|||
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