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<taxonomicName LSID="79373CA5-B519-5C8F-A0F3-D1F0BF6F8C21" authority="(Smetana, 1971)" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Iratiquedius mutator" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="mutator" status="comb. nov.">Iratiquedius mutator (Smetana, 1971)</taxonomicName>
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<taxonomicName id="014D41BDBE009AD8812CEB91D4BBC71A" LSID="79373CA5-B519-5C8F-A0F3-D1F0BF6F8C21" authority="(Smetana, 1971)" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator" status="comb. nov.">Iratiquedius mutator (Smetana, 1971)</taxonomicName>
<taxonomicNameLabel id="1028CF398727B91BBE5CF0AE3296C4BF" pageId="0" pageNumber="129">comb. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">Figs 2C, D</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">, 4C, D</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">, 6C, D, G, J, K</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">, 9B</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributions of A Iratiquedius amabilis (Smetana) (circles), I. mutator (Smetana) (squares) B I. prostans (Horn) (black - specimens examined, white - literature records) C I. seriatus (Horn) (circles), I. uncifer sp. nov. (squares) D Paraquedius puncticeps (Horn) (circles), P. marginicollis sp. nov. (squares) E Quediellus debilis (Horn) (circles) (black - specimens examined, white - literature records)." figureDoi="10.3897/zookeys.1134.87853.figure11" httpUri="https://binary.pensoft.net/fig/779769" pageId="0" pageNumber="129">, 11A (map)</figureCitation>
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<paragraph id="4B19048C5CEDD06A0EE0DEEB8AB7191B" pageId="0" pageNumber="129">
<figureCitation id="8E3BD353FD0B436EB391EF975C080EBC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A-H dorsal habitus of A, B Iratiquedius amabilis (Smetana) A male holotype B female non-type C, D I. mutator (Smetana) C male non-type D female holotype E I. prostans (Horn) F I. seriatus (Horn) G Paraquedius puncticeps (Horn) H P. marginicollis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1134.87853.figure2" httpUri="https://binary.pensoft.net/fig/779760" pageId="0" pageNumber="129">Figs 2C, D</figureCitation>
<figureCitation id="56F19E04223F3C263DBAF99209214D49" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">, 4C, D</figureCitation>
<figureCitation id="087E152935C253D99CAC8CCBB933D316" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">, 6C, D, G, J, K</figureCitation>
<figureCitation id="0BB4B782555CA3F39A9DDD1066D93E57" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">, 9B</figureCitation>
<figureCitation id="F0A81C615DA8738D0D3C93BF48150922" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributions of A Iratiquedius amabilis (Smetana) (circles), I. mutator (Smetana) (squares) B I. prostans (Horn) (black - specimens examined, white - literature records) C I. seriatus (Horn) (circles), I. uncifer sp. nov. (squares) D Paraquedius puncticeps (Horn) (circles), P. marginicollis sp. nov. (squares) E Quediellus debilis (Horn) (circles) (black - specimens examined, white - literature records)." figureDoi="10.3897/zookeys.1134.87853.figure11" httpUri="https://binary.pensoft.net/fig/779769" pageId="0" pageNumber="129">, 11A (map)</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) mutator</taxonomicName>
<subSubSection id="06E83196B0C47234234FF8DA7902B5BD" pageId="0" pageNumber="129" type="reference_group">
<paragraph id="0C4796C0A71F1D6D8DE0CCFB0486AF87" pageId="0" pageNumber="129">
<taxonomicName id="C576541C44067A4E2FAFA173955BFE0E" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) mutator</taxonomicName>
Smetana, 1971: 206.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) mutator</taxonomicName>
<paragraph id="5298991434B1B835C8E8540EE264B377" pageId="0" pageNumber="129">
<taxonomicName id="681160DB31CDD81EE95873B86AFFD361" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator" status="comb. nov." subGenus="Raphirus">Quedius (Raphirus) mutator</taxonomicName>
:
<bibRefCitation author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="129" pagination="95 - 104" refId="B12" refString="Smetana, A, 1990. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6. Coleopterists Bulletin 44: 95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana 1990</bibRefCitation>
<bibRefCitation id="BE6529EF92340793B2A5BFB7E3828CAA" author="Smetana, A" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="129" pagination="95 - 104" refId="B12" refString="Smetana, A, 1990. Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6. Coleopterists Bulletin 44: 95 - 104" title="Revision of the tribe Quediini of America North of Mexico (Coleoptera: Staphylinidae). Supplementum 6." volume="44" year="1990">Smetana 1990</bibRefCitation>
(distributional records, misidentification of
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" family="Staphylinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Q. amabilis" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="amabilis" status="comb. nov.">Q. amabilis</taxonomicName>
<taxonomicName id="AD0F62C17FD54C2F3708BDF0507E51A4" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Brunke" baseAuthorityYear="2022" class="Insecta" family="Staphylinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Q. amabilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="amabilis" status="comb. nov.">Q. amabilis</taxonomicName>
, in part).
</paragraph>
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<subSubSection pageId="0" pageNumber="129" type="type locality">
<paragraph pageId="0" pageNumber="129">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="129">
<subSubSection id="34871253DD1DDE84D0C450E437979A81" pageId="0" pageNumber="129" type="type locality">
<paragraph id="9DE561C92BEE268D8B737A4154FCF249" pageId="0" pageNumber="129">Type locality.</paragraph>
<paragraph id="4928D1B3AF255560B9F233C4EB2102C8" pageId="0" pageNumber="129">
8 miles north of Post Pile Camp [possibly 'Valentine
<normalizedToken originalValue="Spring">Spring'</normalizedToken>
<normalizedToken id="5E5075F9234EC61622377927412CDEBA" originalValue="Spring">Spring'</normalizedToken>
, ~ 1660 m], Tehama County, California, United States.
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<paragraph pageId="0" pageNumber="129">Type material.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Holotype</emphasis>
<subSubSection id="2B34D69531E228651C354C5DD2C68D2C" pageId="0" pageNumber="129" type="type material">
<paragraph id="31467B3077C59C5396192D789FE32BD9" pageId="0" pageNumber="129">Type material.</paragraph>
<paragraph id="443001027AF34202E2C3766FF8F05EBA" pageId="0" pageNumber="129">
<emphasis id="DDC567114F063532CF80B8572B810DE2" bold="true" italics="true" pageId="0" pageNumber="129">Holotype</emphasis>
(female, CNC): 8 mi N Post Pile Camp, Tehema [Tehama] Co., Cal, VIII-30-60 [handwritten label] / R.O. Schuster Collector [printed label] / HOLOTYPE Quedius mutator Smetana 1968, CNC No. 10877 [red printed label] / CNC [handwritten label] / CNC93512 [identifier].
</paragraph>
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<paragraph id="6AFF21724BD13C4F4A9F528E796637F4" pageId="0" pageNumber="129">
The female holotype has the shortest, most sparsely punctate elytra of all known individuals attributed here to either
<taxonomicName lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
<emphasis italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
<taxonomicName id="805AC2A573448281FA15B020964623D4" lsidName="I. mutator" pageId="0" pageNumber="129" rank="species" species="mutator">
<emphasis id="A134343C05C4746469B2E5C4ABC6D91B" italics="true" pageId="0" pageNumber="129">I. mutator</emphasis>
</taxonomicName>
or
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
<taxonomicName id="1B00B381FDE1613E4CCC417755E192CA" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis id="A9468F2A51A570FDC61EC9F69C871309" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
</taxonomicName>
. The shape of female tergite X is unique and differs from the other examined females (here attributed to
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
<taxonomicName id="5D3EBB4778974D5D2348603327F799A4" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis id="44A87E31ADAA4AA80B8BA71BC3E56FDC" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
</taxonomicName>
) by the projected, sharp apex (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">9B</figureCitation>
<figureCitation id="C563503B0D25AEFA7495189ABB438FE9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="type material">
<paragraph pageId="0" pageNumber="129">Non-type material.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">United States: California</emphasis>
<subSubSection id="531DC037127BD83CD0704EE54BA55976" pageId="0" pageNumber="129" type="type material">
<paragraph id="6B0769DAB782D4068E5DBDA526CCF1FA" pageId="0" pageNumber="129">Non-type material.</paragraph>
<paragraph id="3035725882B3E39930A196E278034C5B" pageId="0" pageNumber="129">
<emphasis id="522191885F095A5F616FC71DE0C1A4C3" bold="true" pageId="0" pageNumber="129">United States: California</emphasis>
: Butte Co.: 3 mi NE Loma [Loma Rica], 39.354, -121.411, 151 m, 3.V.1981, sifting litter along spring, D. Chandler (2 males, FMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius mutator" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="mutator">
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius mutator</emphasis>
<subSubSection id="91473048BAA8BF013226A1045CD3B00A" pageId="0" pageNumber="129" type="diagnosis">
<paragraph id="6A99423294E4ACEADEE2940FA34F7E97" pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph id="D81032E4C280E3D450603984900D786E" pageId="0" pageNumber="129">
<taxonomicName id="761F1535D7EDD9FCAA003E209C767B68" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator">
<emphasis id="C6043783E25A453F1B5E3F8548450E80" italics="true" pageId="0" pageNumber="129">Iratiquedius mutator</emphasis>
</taxonomicName>
may be recognized within the genus by a combination of the evenly punctate elytra, the lack of golden setae or impressions at the base of the abdominal tergites and the indistinct micropunctures on the anterior angles of the pronotum. The species most closely resembles
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
<taxonomicName id="FFD999901D72650DFB09E61625E0C64F" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis id="85851D91F7950A1DDD2D2AD9F983C1F6" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
</taxonomicName>
and can be distinguished from it by the indistinct micropunctures of the anterior angles of the pronotum, the longer apex of the median lobe in lateral view in males or sharp, pointed apex of female tergite X.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="redescription">
<paragraph pageId="0" pageNumber="129">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="129">
<subSubSection id="C84C52E4EF1704BCA0AE6738FB0CF802" pageId="0" pageNumber="129" type="redescription">
<paragraph id="B79A7ED6FC6621018C72B64091055DBC" pageId="0" pageNumber="129">Redescription.</paragraph>
<paragraph id="266E054E6E58F6ADBEB0837197EE56C8" pageId="0" pageNumber="129">
Measurements ♂ (
<emphasis italics="true" pageId="0" pageNumber="129">n</emphasis>
<emphasis id="267C1488DB70632453F4ABF7B967829D" italics="true" pageId="0" pageNumber="129">n</emphasis>
= 2): HW/HL 1.22-1.25; PW/PL 1.20-1.24; EW/EL 1.37-1.38; ESut/PL 0.68-0.71; PW/HW 1.21-1.23; forebody length 3.4-3.5 mm.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<paragraph id="D808B94E548DAA3F313081CFAA52A5C8" pageId="0" pageNumber="129">
Measurements ♀ (
<emphasis italics="true" pageId="0" pageNumber="129">n</emphasis>
<emphasis id="318A047E32A5CDF591542FA981CB4F6E" italics="true" pageId="0" pageNumber="129">n</emphasis>
= 1): HW/HL 1.20; PW/PL 1.22; EW/EL 1.70; ESut/PL 0.58; PW/HW 1.16; forebody length 2.8 mm.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<paragraph id="69C34EFFC8728E7EACCF9CDE2BAB7C67" pageId="0" pageNumber="129">
Extremely similar to
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
<taxonomicName id="C44B751037599A4C02FA11C5E0B63DF3" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis id="C698386B330464ABC909A0DBB60EC1A2" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
</taxonomicName>
, and differing only in the following: antennomeres 1-4 (macropterous) or 1-3 (brachypterous) clearly elongate; pronotum moderately (macropterous) to weakly (brachypterous) transverse; pronotum with micropunctures of anterior angles indistinct (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">4C, D</figureCitation>
<figureCitation id="3136F2299A6A9B57357B9CF9F1B92992" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A-D anterior angle of the pronotum A, B Iratiquedius amabilis (Smetana) C, D I. mutator (Smetana) A, C macropterous morphotype B, D brachypterous morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure4" httpUri="https://binary.pensoft.net/fig/779762" pageId="0" pageNumber="129">4C, D</figureCitation>
); elytra overall shorter, and at suture relatively shorter than pronotum when comparing morphotypes; male sternite VIII with emargination vaguely more rounded at middle but varying from moderately narrow to as wide as
<taxonomicName lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
<taxonomicName id="075FE62A3EE3453EEEAFC0FB9629125E" lsidName="I. amabilis" pageId="0" pageNumber="129" rank="species" species="amabilis">
<emphasis id="0B63ADAFEFFDE92D4FC2743465923AFB" italics="true" pageId="0" pageNumber="129">I. amabilis</emphasis>
</taxonomicName>
; male tergite X more slender with narrower apex; male sternite IX broader, more strongly convergent to apex; apex of median lobe in lateral view longer, more elongate and sharp (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6C, D</figureCitation>
<figureCitation id="B8371FC6E28822F2F82F69142D41F264" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6C, D</figureCitation>
), median lobe in ventral view with apex entire and pointed (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6G</figureCitation>
<figureCitation id="69A584A9AA03B7D6945513283CCC7366" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6G</figureCitation>
); paramere with marginal row becoming sparser distally (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6J, K</figureCitation>
<figureCitation id="3EAD1B6B296A040290586727E565B160" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A-G, L-O median lobe of aedeagus A-D, N, O in lateral view E-G, L, M in ventral view O with internal sac everted H-K, P underside of paramere showing peg setae A, B, E, F, H, I Iratiquedius amabilis (Smetana) C, D, G, J, K I. mutator (Smetana) L-P I. prostans (Horn). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure6" httpUri="https://binary.pensoft.net/fig/779764" pageId="0" pageNumber="129">6J, K</figureCitation>
); apex of female tergite X sharply projected to a point, with sparse marginal setae (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">9B</figureCitation>
<figureCitation id="62B1777C413A0C38E0818FD7259E2726" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">9B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="distribution">
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">United States</emphasis>
<subSubSection id="751CFFD6BDFA8796AA491132E52A3100" pageId="0" pageNumber="129" type="distribution">
<paragraph id="AE112DA47F03CFBDDA45168308B6AA49" pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph id="61208C75B5D9A1A4A99DF5A27FE312BF" pageId="0" pageNumber="129">
<emphasis id="74685C4A6D0BE4B5751D3ADD40EDE949" bold="true" pageId="0" pageNumber="129">United States</emphasis>
: CA.
</paragraph>
<paragraph pageId="0" pageNumber="129">This species is known only from two localities: one in the mountains of the northern Coast Range and one in the Central Valley region.</paragraph>
<paragraph id="13B2AE19B8B7001ECCFBCD204CCA2033" pageId="0" pageNumber="129">This species is known only from two localities: one in the mountains of the northern Coast Range and one in the Central Valley region.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="bionomics">
<paragraph pageId="0" pageNumber="129">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="129">
<subSubSection id="8075938B4E23A06691EF3E71A5CA4D20" pageId="0" pageNumber="129" type="bionomics">
<paragraph id="60C3830BC31431C0493728D29A007D38" pageId="0" pageNumber="129">Bionomics.</paragraph>
<paragraph id="588A618760CEBDA78E988C0FE2006457" pageId="0" pageNumber="129">
Both known localities are at least near springs, within areas of relatively dry, open woodland. Specimens of strongly hydrophilous species
<taxonomicName lsidName="I. uncifer" pageId="0" pageNumber="129" rank="species" species="uncifer">
<emphasis italics="true" pageId="0" pageNumber="129">I. uncifer</emphasis>
<taxonomicName id="D1489634AD4D67DE10081C36989446CE" lsidName="I. uncifer" pageId="0" pageNumber="129" rank="species" species="uncifer">
<emphasis id="09BEBB5549E64CB8B0887D27DBBEF435" italics="true" pageId="0" pageNumber="129">I. uncifer</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
<emphasis italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
<taxonomicName id="68BE40F091C17836973DEC224896B485" lsidName="I. prostans" pageId="0" pageNumber="129" rank="species" species="prostans">
<emphasis id="6305BEABDFE6EBE78409A23934CDE34F" italics="true" pageId="0" pageNumber="129">I. prostans</emphasis>
</taxonomicName>
were co-collected at the type locality. The two non-type males were collected in litter along the margins of a spring. The holotype was collected at around 1600 m, while the non-type males were collected around 150 m.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="comments">
<paragraph pageId="0" pageNumber="129">Comments.</paragraph>
<paragraph pageId="0" pageNumber="129">
<subSubSection id="A8227C6D13AE3A998FE1DFE4E5F2E626" pageId="0" pageNumber="129" type="comments">
<paragraph id="0EE0399AC6BE79DCA7F0A3C5F78FAC46" pageId="0" pageNumber="129">Comments.</paragraph>
<paragraph id="5459254F2F5C8CB578FE12F2766DAD46" pageId="0" pageNumber="129">
The concept given here for
<taxonomicName authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" family="Staphylinidae" genus="Iratiquedius" kingdom="Animalia" lsidName="Iratiquedius mutator" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="mutator">
<emphasis italics="true" pageId="0" pageNumber="129">Iratiquedius mutator</emphasis>
<taxonomicName id="3A64DDEED93E4D51353E5F44F566C0F4" authorityName="Brunke" authorityYear="2022" baseAuthorityName="Smetana" baseAuthorityYear="1971" class="Insecta" family="Staphylinidae" genus="Iratiquedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iratiquedius mutator" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="mutator">
<emphasis id="38340D62D7C667A866EE7649B3C3880C" italics="true" pageId="0" pageNumber="129">Iratiquedius mutator</emphasis>
</taxonomicName>
is considered to be a step forward but may need to be modified in the unlikely case that the population in the Central Valley is a third, undescribed species. Males from near the type locality will be needed to determine this with certainty.
</paragraph>

View file

@ -1,239 +1,239 @@
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<mods:name type="personal">
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<mods:namePart>Hernandez-Restrepo, Margarita</mods:namePart>
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<mods:namePart>Groenewald, Johannes Zacharias</mods:namePart>
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<mods:namePart>Cai, Lei</mods:namePart>
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<mods:namePart>Crous, Pedro W.</mods:namePart>
<mods:namePart id="6267ECBB84760E4978BF6B39B88F4523">Crous, Pedro W.</mods:namePart>
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<mods:date>2020</mods:date>
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<treatment ID-GBIF-Taxon="162481720" LSID="urn:lsid:plazi:treatment:79C5335520025B6CB44A5142F644BAA9" httpUri="http://treatment.plazi.org/id/79C5335520025B6CB44A5142F644BAA9" lastPageNumber="49" pageId="0" pageNumber="49">
<subSubSection pageId="0" pageNumber="49" type="nomenclature">
<paragraph pageId="0" pageNumber="49">
<taxonomicName LSID="79C53355-2002-5B6C-B44A-5142F644BAA9" authority="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum" status="sp. nov.">Ascochyta benningiorum Hern.-Restr., L. W. Hou, L. Cai &amp; Crous</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="49">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ascochyta benningiorum (CBS 144957). A, B Colony on OA (front and reverse) C, D colony on MEA (front and reverse) E, F colony on PDA (front and reverse) G pycnidia forming on OA H pycnidium I section of pycnidium J section of pycnidial wall K-M conidiogenous cells N conidia. Scale bars: 100 μm (H, I); 10 μm (J); 5 μm (K-N)." figureDoi="10.3897/mycokeys.65.47704.figure2" httpUri="https://binary.pensoft.net/fig/388205" pageId="0" pageNumber="49">Figure 2</figureCitation>
<treatment id="79C5335520025B6CB44A5142F644BAA9" ID-GBIF-Taxon="162481720" LSID="urn:lsid:plazi:treatment:79C5335520025B6CB44A5142F644BAA9" httpUri="http://treatment.plazi.org/id/79C5335520025B6CB44A5142F644BAA9" lastPageNumber="49" pageId="0" pageNumber="49" scope_class="Dothideomycetes" scope_family="Didymellaceae" scope_order="Pleosporales">
<subSubSection id="EC55327479B47D399F6AB953A997F702" pageId="0" pageNumber="49" type="nomenclature">
<paragraph id="E42F2D27DB51C4A5227EAC6C7F95985F" pageId="0" pageNumber="49">
<taxonomicName id="C0C3744B1D1D235D224494D807542E91" ID-CoL="9YNMW" LSID="79C53355-2002-5B6C-B44A-5142F644BAA9" authority="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum" status="sp. nov.">Ascochyta benningiorum Hern.-Restr., L. W. Hou, L. Cai &amp; Crous</taxonomicName>
<taxonomicNameLabel id="57DBC1909F02CCAA56B33D30B9EE7096" pageId="0" pageNumber="49">sp. nov.</taxonomicNameLabel>
<figureCitation id="A3C7EAA6FE0D12ECAC577D18D6318C54" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Ascochyta benningiorum (CBS 144957). A, B Colony on OA (front and reverse) C, D colony on MEA (front and reverse) E, F colony on PDA (front and reverse) G pycnidia forming on OA H pycnidium I section of pycnidium J section of pycnidial wall K-M conidiogenous cells N conidia. Scale bars: 100 μm (H, I); 10 μm (J); 5 μm (K-N)." figureDoi="10.3897/mycokeys.65.47704.figure2" httpUri="https://binary.pensoft.net/fig/388205" pageId="0" pageNumber="49">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="etymology">
<paragraph pageId="0" pageNumber="49">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum">
<emphasis italics="true" pageId="0" pageNumber="49">benningiorum</emphasis>
<subSubSection id="954D9288065AA0F5D5C840FCEFE54C00" pageId="0" pageNumber="49" type="etymology">
<paragraph id="F2CA652BBA31BC3AE20ED57518921114" pageId="0" pageNumber="49">Etymology.</paragraph>
<paragraph id="AA64F5BD8B322DC2E0034993D6BC6C43" pageId="0" pageNumber="49">
<taxonomicName id="EE51ACD76AECF3C30F6C3BD808C0703D" authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum">
<emphasis id="3B7CB1C5A9A19F5B386F6D6494B38E81" italics="true" pageId="0" pageNumber="49">benningiorum</emphasis>
</taxonomicName>
refers to Eva, Bas &amp; Anne Benning who collected the soil sample from which the ex-type strain was isolated.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="typus">
<paragraph pageId="0" pageNumber="49">Typus.</paragraph>
<paragraph pageId="0" pageNumber="49">
<subSubSection id="4D3AE3FD1C3734A3D4BBC7C3110ABE7A" pageId="0" pageNumber="49" type="typus">
<paragraph id="C1875C9A0350D545DBF9E986A62FC51E" pageId="0" pageNumber="49">Typus.</paragraph>
<paragraph id="C56E4DA77B686C64E6A5A88907BF610D" pageId="0" pageNumber="49">
The Netherlands. Gelderland province, Wijchen, isolated from garden soil, Mar. 2017, E. Benning, B. Benning &amp; A. Benning (
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">holotype</emphasis>
<emphasis id="49284C0241154CFAC5D0614DF6388C50" bold="true" italics="true" pageId="0" pageNumber="49">holotype</emphasis>
designated here CBS H-24104, living ex-type culture CBS 144957 = JW 196005).
</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis italics="true" pageId="0" pageNumber="49">Conidiomata</emphasis>
<paragraph id="2B4D373DD9FF4414F0ADC463B073B16F" pageId="0" pageNumber="49">
<emphasis id="A3103715601B87FC3BE186C0D7587FC0" italics="true" pageId="0" pageNumber="49">Conidiomata</emphasis>
pycnidial, mostly solitary, sometimes confluent, globose or subglobose, irregularly-shaped with age, brown to dark brown, glabrous, mostly produced on the agar surface and some immersed, 140-480(-580)
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="3CC10F1317F090B49FFD6BF11E54F939" originalValue="×">x</normalizedToken>
100-370(-440)
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="8EDBBD099951619F64634EB09975C46C" originalValue="μm">μm</normalizedToken>
; with 1-6(-10) slightly papillate ostioles; pycnidial wall pseudoparenchymatous, 4-8 layers, 14.5-65
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="96447A5056324052719ED0A512DBFE7A" originalValue="μm">μm</normalizedToken>
thick, outer layers composed of brown, flattened polygonal cells of 11-28
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="F91F01A85CF56E7376CDDD34B751243A" originalValue="μm">μm</normalizedToken>
diam.
<emphasis italics="true" pageId="0" pageNumber="49">Conidiogenous cells</emphasis>
<emphasis id="2BE53538F01F8DD2C9D2DA56FAFA05E6" italics="true" pageId="0" pageNumber="49">Conidiogenous cells</emphasis>
phialidic, hyaline, smooth, globose, ampulliform to lageniform, 5.5-9
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9E309F2CC0A00B162A3FA0E6ADB42550" originalValue="×">x</normalizedToken>
4-6.5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="26B5392C7AD9C81ED37B07776C0821FD" originalValue="μm">μm</normalizedToken>
.
<emphasis italics="true" pageId="0" pageNumber="49">Conidia</emphasis>
<emphasis id="B714B647C7B947129650C8724B7EB751" italics="true" pageId="0" pageNumber="49">Conidia</emphasis>
cylindrical, hyaline, smooth- and thin-walled, mostly straight, occasionally curved, aseptate, (3.5-)4.5-7
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="A41B9F446BF8FAF0F184C91C26127FAD" originalValue="×">x</normalizedToken>
1.5-2.5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="CBA4A0C599D5ACDA071D474396B49CBE" originalValue="μm">μm</normalizedToken>
, 2-guttulate, small.
<emphasis italics="true" pageId="0" pageNumber="49">
<emphasis id="F46EAC1ADECBC8A146247010FACC96FE" italics="true" pageId="0" pageNumber="49">
Conidia
<taxonomicName class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="matrix" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="matrix">matrix</taxonomicName>
<taxonomicName id="C23814CF0748C9F0693767EA50ED9547" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="matrix" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="matrix">matrix</taxonomicName>
</emphasis>
whitish.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">Culture characteristics.</paragraph>
<paragraph pageId="0" pageNumber="49">Colonies after 7 d at 25 °C, on OA reaching 50-55 mm diam, aerial mycelium floccose, olivaceous to olivaceous black, buff towards the periphery, abundant production of pycnidia, margin irregular; reverse concolorous with the surface. On MEA reaching 40-45 mm diam, aerial mycelium floccose, concentric circles, centre pink, grey olivaceous, mouse grey, rosy buff toward periphery, moderate production of pycnidia, margin irregular; reverse orange, olivaceous black toward periphery. On PDA reaching 45-50 mm diam, aerial mycelium floccose, dark brick to olivaceous grey, buff towards periphery, abundant production of pycnidia, margin irregular; reverse concolorous with the surface. NaOH spot test negative on OA.</paragraph>
<subSubSection id="30C45B912009B520E7E9417319D77E90" pageId="0" pageNumber="49" type="description">
<paragraph id="E821DB48BEF2D1D6E6D37848FC8F285D" pageId="0" pageNumber="49">Culture characteristics.</paragraph>
<paragraph id="EB20EBA8D25932C6EC9F05D8528F9532" pageId="0" pageNumber="49">Colonies after 7 d at 25 °C, on OA reaching 50-55 mm diam, aerial mycelium floccose, olivaceous to olivaceous black, buff towards the periphery, abundant production of pycnidia, margin irregular; reverse concolorous with the surface. On MEA reaching 40-45 mm diam, aerial mycelium floccose, concentric circles, centre pink, grey olivaceous, mouse grey, rosy buff toward periphery, moderate production of pycnidia, margin irregular; reverse orange, olivaceous black toward periphery. On PDA reaching 45-50 mm diam, aerial mycelium floccose, dark brick to olivaceous grey, buff towards periphery, abundant production of pycnidia, margin irregular; reverse concolorous with the surface. NaOH spot test negative on OA.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="materials_examined">
<paragraph pageId="0" pageNumber="49">Additional specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="49">The Netherlands. Gelderland province, Wijchen, isolated from garden soil, Mar. 2017, E. Benning, B. Benning &amp; A. Benning, JW 196023 = CBS 144958; ibid. JW 196013.</paragraph>
<subSubSection id="1206B073B16F2549D506584A6FD911C0" pageId="0" pageNumber="49" type="materials_examined">
<paragraph id="B5513C94932B9BBCA2D366A76DA8839F" pageId="0" pageNumber="49">Additional specimens examined.</paragraph>
<paragraph id="9F8A42D77ABD9E51E2021DDDF8A5BCE8" pageId="0" pageNumber="49">The Netherlands. Gelderland province, Wijchen, isolated from garden soil, Mar. 2017, E. Benning, B. Benning &amp; A. Benning, JW 196023 = CBS 144958; ibid. JW 196013.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="notes">
<paragraph pageId="0" pageNumber="49">Notes.</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum">
<emphasis italics="true" pageId="0" pageNumber="49">Ascochyta benningiorum</emphasis>
<subSubSection id="A2CCBA292004D008B72B1A7A81D009B3" pageId="0" pageNumber="49" type="notes">
<paragraph id="BA8CDAEC3F4264CC34879BF185E7006C" pageId="0" pageNumber="49">Notes.</paragraph>
<paragraph id="5B913B3C365CDFB999F6C23E8E5CFC50" pageId="0" pageNumber="49">
<taxonomicName id="2FA9CA23337BE528A144F8893B88A8A2" authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum">
<emphasis id="351AF823796BF3079AC8932526138F86" italics="true" pageId="0" pageNumber="49">Ascochyta benningiorum</emphasis>
</taxonomicName>
is represented in the phylogenetic tree by three isolates (CBS 144957, CBS 144958 and JW 196013) from the same soil sample collected in Wijchen (Gelderland province).
<taxonomicName authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum">
<emphasis italics="true" pageId="0" pageNumber="49">Ascochyta benningiorum</emphasis>
<taxonomicName id="F3293343F88D2D5E83E2654030A1917B" authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum">
<emphasis id="ACE3F226DA1DCAE21C065976E585866E" italics="true" pageId="0" pageNumber="49">Ascochyta benningiorum</emphasis>
</taxonomicName>
grouped in a distinct clade close to
<taxonomicName lsidName="A. phacae" pageId="0" pageNumber="49" rank="species" species="phacae">
<emphasis italics="true" pageId="0" pageNumber="49">A. phacae</emphasis>
<taxonomicName id="98560EA224DAB68C76C533092FDFB466" lsidName="A. phacae" pageId="0" pageNumber="49" rank="species" species="phacae">
<emphasis id="91F6E4C981BB7F69F15F3F8E83D903E5" italics="true" pageId="0" pageNumber="49">A. phacae</emphasis>
</taxonomicName>
(Figure
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree generated from the maximum-likelihood analysis based on the combined ITS, LSU, tub 2 and rpb 2 sequence alignment of Didymellaceae members. The RAxML bootstrap support values (BS), Bayesian posterior probabilities (PP), and parsimony bootstrap support values (PBS) are given at the nodes (BS / PP / PBS). BS and PBS values represent parsimony bootstrap support values&gt; 50 %. Full supported branches are indicated in bold. The scale bar represents the expected number of changes per site. Ex-type strains are represented in bold. Strains obtained in the current study are printed in green; among them, whilst strains that represent new taxa are printed in red. Some of the basal branches were shortened to facilitate layout (the fraction in round parentheses refers to the presented length compared to the actual length of the branch). The tree was rooted to Coniothyrium palmarum CBS 400.71 and Leptosphaeria doliolum CBS 505.75." figureDoi="10.3897/mycokeys.65.47704.figure1" httpUri="https://binary.pensoft.net/fig/388204" pageId="0" pageNumber="49">1</figureCitation>
<figureCitation id="48BA2D3965E2EA5C0533810B09A74002" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree generated from the maximum-likelihood analysis based on the combined ITS, LSU, tub 2 and rpb 2 sequence alignment of Didymellaceae members. The RAxML bootstrap support values (BS), Bayesian posterior probabilities (PP), and parsimony bootstrap support values (PBS) are given at the nodes (BS / PP / PBS). BS and PBS values represent parsimony bootstrap support values&gt; 50 %. Full supported branches are indicated in bold. The scale bar represents the expected number of changes per site. Ex-type strains are represented in bold. Strains obtained in the current study are printed in green; among them, whilst strains that represent new taxa are printed in red. Some of the basal branches were shortened to facilitate layout (the fraction in round parentheses refers to the presented length compared to the actual length of the branch). The tree was rooted to Coniothyrium palmarum CBS 400.71 and Leptosphaeria doliolum CBS 505.75." figureDoi="10.3897/mycokeys.65.47704.figure1" httpUri="https://binary.pensoft.net/fig/388204" pageId="0" pageNumber="49">1</figureCitation>
). However, it morphologically differs from
<taxonomicName lsidName="A. phacae" pageId="0" pageNumber="49" rank="species" species="phacae">
<emphasis italics="true" pageId="0" pageNumber="49">A. phacae</emphasis>
<taxonomicName id="53F6FC51FDFD0F9D0602A5F5A972F69C" lsidName="A. phacae" pageId="0" pageNumber="49" rank="species" species="phacae">
<emphasis id="9773DB5735F4D522B3B21D319CAEE232" italics="true" pageId="0" pageNumber="49">A. phacae</emphasis>
</taxonomicName>
by having smaller (3.5-7
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="5366BBC77942B9CB54008D3C84AB3A43" originalValue="×">x</normalizedToken>
1.5-2.5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="75264E6FE1547E41674025742A6108C5" originalValue="μm">μm</normalizedToken>
) and aseptate conidia. In
<taxonomicName lsidName="A. phacae" pageId="0" pageNumber="49" rank="species" species="phacae">
<emphasis italics="true" pageId="0" pageNumber="49">A. phacae</emphasis>
<taxonomicName id="E07A605E775C173085EEBD8F7A0C69BB" lsidName="A. phacae" pageId="0" pageNumber="49" rank="species" species="phacae">
<emphasis id="8E929409CC91B0FE9EAA649912E7055B" italics="true" pageId="0" pageNumber="49">A. phacae</emphasis>
</taxonomicName>
the conidia are 7-10
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="CD02144142E30F8D0459183A17EAA345" originalValue="×">x</normalizedToken>
2-4
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="EAF2618BE1F97BD8C99C55493D77AA03" originalValue="μm">μm</normalizedToken>
and 0-1-septate (
<bibRefCitation author="Corbaz, R" journalOrPublisher="Sydowia" pageId="0" pageNumber="49" pagination="229 - 230" refId="B18" refString="Corbaz, R, 1955. Sur Didymella phacae Corbaz. Sydowia 9: 229 - 230" title="Sur Didymella phacae Corbaz." volume="9" year="1955">Corbaz 1955</bibRefCitation>
<bibRefCitation id="EF2FF584E4AFEB4C7326AD6BB79E040E" author="Corbaz, R" journalOrPublisher="Sydowia" pageId="0" pageNumber="49" pagination="229 - 230" refId="B18" refString="Corbaz, R, 1955. Sur Didymella phacae Corbaz. Sydowia 9: 229 - 230" title="Sur Didymella phacae Corbaz." volume="9" year="1955">Corbaz 1955</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="49">
<paragraph id="D7BFCD662BEB83A33616CF8793B61BEF" pageId="0" pageNumber="49">
Species in
<taxonomicName class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Ascochyta</emphasis>
<taxonomicName id="8CC4D6A5E161750F2DE44F6BD0267D61" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="genus">
<emphasis id="DFC84FDCA4C2DAAF7BE00857193DAB07" italics="true" pageId="0" pageNumber="49">Ascochyta</emphasis>
</taxonomicName>
are commonly regarded as plant pathogens, especially of cereal crops and legumes (
<bibRefCitation DOI="https://doi.org/10.1007/s10658-007-9132-x" author="Davidson, JA" journalOrPublisher="European Journal of Plant Pathology" pageId="0" pageNumber="49" pagination="99 - 110" refId="B24" refString="Davidson, JA, Kimber, RBE, 2007. Integrated disease management of ascochyta blight in pulse crops. European Journal of Plant Pathology 119: 99 - 110, DOI: https://doi.org/10.1007/s10658-007-9132-x" title="Integrated disease management of ascochyta blight in pulse crops." url="https://doi.org/10.1007/s10658-007-9132-x" volume="119" year="2007">Davidson and Kimber 2007</bibRefCitation>
<bibRefCitation id="93956E73015F7E54A7AF1B9705264978" DOI="https://doi.org/10.1007/s10658-007-9132-x" author="Davidson, JA" journalOrPublisher="European Journal of Plant Pathology" pageId="0" pageNumber="49" pagination="99 - 110" refId="B24" refString="Davidson, JA, Kimber, RBE, 2007. Integrated disease management of ascochyta blight in pulse crops. European Journal of Plant Pathology 119: 99 - 110, DOI: https://doi.org/10.1007/s10658-007-9132-x" title="Integrated disease management of ascochyta blight in pulse crops." url="https://doi.org/10.1007/s10658-007-9132-x" volume="119" year="2007">Davidson and Kimber 2007</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1007/s10658-007-9117-9" author="Tivoli, B" journalOrPublisher="European Journal of Plant Pathology" pageId="0" pageNumber="49" pagination="59 - 76" refId="B61" refString="Tivoli, B, Banniza, S, 2007. Comparison of the epidemiology of ascochyta blights on grain legumes. European Journal of Plant Pathology 119: 59 - 76, DOI: https://doi.org/10.1007/s10658-007-9117-9" title="Comparison of the epidemiology of ascochyta blights on grain legumes." url="https://doi.org/10.1007/s10658-007-9117-9" volume="119" year="2007">Tivoli and Banniza 2007</bibRefCitation>
<bibRefCitation id="F914E1DA36A98D842C51D0274C658A33" DOI="https://doi.org/10.1007/s10658-007-9117-9" author="Tivoli, B" journalOrPublisher="European Journal of Plant Pathology" pageId="0" pageNumber="49" pagination="59 - 76" refId="B61" refString="Tivoli, B, Banniza, S, 2007. Comparison of the epidemiology of ascochyta blights on grain legumes. European Journal of Plant Pathology 119: 59 - 76, DOI: https://doi.org/10.1007/s10658-007-9117-9" title="Comparison of the epidemiology of ascochyta blights on grain legumes." url="https://doi.org/10.1007/s10658-007-9117-9" volume="119" year="2007">Tivoli and Banniza 2007</bibRefCitation>
), and only a few species were reported from soil, namely
<taxonomicName lsidName="A. fabae" pageId="0" pageNumber="49" rank="species" species="fabae">
<emphasis italics="true" pageId="0" pageNumber="49">A. fabae</emphasis>
<taxonomicName id="38816F13ADD8F0D28E40BD7D6EAE022C" lsidName="A. fabae" pageId="0" pageNumber="49" rank="species" species="fabae">
<emphasis id="4B855A5D33D151093B5358992C47D6ED" italics="true" pageId="0" pageNumber="49">A. fabae</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="A. lentis" pageId="0" pageNumber="49" rank="species" species="lentis">
<emphasis italics="true" pageId="0" pageNumber="49">A. lentis</emphasis>
<taxonomicName id="E5AAF48B2ECC8FE79E2E16B1C668E869" lsidName="A. lentis" pageId="0" pageNumber="49" rank="species" species="lentis">
<emphasis id="4E6EE5D3755D56D95F5C9ADD88AF4A68" italics="true" pageId="0" pageNumber="49">A. lentis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="A. pisi" pageId="0" pageNumber="49" rank="species" species="pisi">
<emphasis italics="true" pageId="0" pageNumber="49">A. pisi</emphasis>
<taxonomicName id="336FE1B54C692FEE07B6179D33F5DAD2" lsidName="A. pisi" pageId="0" pageNumber="49" rank="species" species="pisi">
<emphasis id="6B58E8777D0C9BEE0A4DAB6432A867BD" italics="true" pageId="0" pageNumber="49">A. pisi</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="A. rabiei" pageId="0" pageNumber="49" rank="species" species="rabiei">
<emphasis italics="true" pageId="0" pageNumber="49">A. rabiei</emphasis>
<taxonomicName id="7F6393974C97F3E2F10A353AED5C0319" lsidName="A. rabiei" pageId="0" pageNumber="49" rank="species" species="rabiei">
<emphasis id="5DE8CD3CEAFBDD63A90904584C0004FA" italics="true" pageId="0" pageNumber="49">A. rabiei</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1080/07060668609501837" author="Gossen, BD" journalOrPublisher="Canadian Journal of Plant Pathology" pageId="0" pageNumber="49" pagination="28 - 32" refId="B33" refString="Gossen, BD, Morrall, RAA, 1986. Transmission of Ascochyta lentis from infected lentil seed and plant residue. Canadian Journal of Plant Pathology 8: 28 - 32, DOI: https://doi.org/10.1080/07060668609501837" title="Transmission of Ascochyta lentis from infected lentil seed and plant residue." url="https://doi.org/10.1080/07060668609501837" volume="8" year="1986">Gossen and Morrall 1986</bibRefCitation>
<bibRefCitation id="C1EA6EE65B4E699EAAFD2C50028EE00D" DOI="https://doi.org/10.1080/07060668609501837" author="Gossen, BD" journalOrPublisher="Canadian Journal of Plant Pathology" pageId="0" pageNumber="49" pagination="28 - 32" refId="B33" refString="Gossen, BD, Morrall, RAA, 1986. Transmission of Ascochyta lentis from infected lentil seed and plant residue. Canadian Journal of Plant Pathology 8: 28 - 32, DOI: https://doi.org/10.1080/07060668609501837" title="Transmission of Ascochyta lentis from infected lentil seed and plant residue." url="https://doi.org/10.1080/07060668609501837" volume="8" year="1986">Gossen and Morrall 1986</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1007/s10658-007-9117-9" author="Tivoli, B" journalOrPublisher="European Journal of Plant Pathology" pageId="0" pageNumber="49" pagination="59 - 76" refId="B61" refString="Tivoli, B, Banniza, S, 2007. Comparison of the epidemiology of ascochyta blights on grain legumes. European Journal of Plant Pathology 119: 59 - 76, DOI: https://doi.org/10.1007/s10658-007-9117-9" title="Comparison of the epidemiology of ascochyta blights on grain legumes." url="https://doi.org/10.1007/s10658-007-9117-9" volume="119" year="2007">Tivoli and Banniza 2007</bibRefCitation>
<bibRefCitation id="0522D5C44283685CF033CEF2C79DAFA9" DOI="https://doi.org/10.1007/s10658-007-9117-9" author="Tivoli, B" journalOrPublisher="European Journal of Plant Pathology" pageId="0" pageNumber="49" pagination="59 - 76" refId="B61" refString="Tivoli, B, Banniza, S, 2007. Comparison of the epidemiology of ascochyta blights on grain legumes. European Journal of Plant Pathology 119: 59 - 76, DOI: https://doi.org/10.1007/s10658-007-9117-9" title="Comparison of the epidemiology of ascochyta blights on grain legumes." url="https://doi.org/10.1007/s10658-007-9117-9" volume="119" year="2007">Tivoli and Banniza 2007</bibRefCitation>
) and
<taxonomicName lsidName="A. syringae" pageId="0" pageNumber="49" rank="species" species="syringae">
<emphasis italics="true" pageId="0" pageNumber="49">A. syringae</emphasis>
<taxonomicName id="D3BA7A25698FA08B4238008758FFA2F1" lsidName="A. syringae" pageId="0" pageNumber="49" rank="species" species="syringae">
<emphasis id="C8320B488D2AFF7BD3DBF6DABB12C350" italics="true" pageId="0" pageNumber="49">A. syringae</emphasis>
</taxonomicName>
in the current study. Nevertheless,
<taxonomicName lsidName="A. benningiorum" pageId="0" pageNumber="49" rank="species" species="benningiorum">
<emphasis italics="true" pageId="0" pageNumber="49">A. benningiorum</emphasis>
<taxonomicName id="D9DDD682C1A5EEDC313315DA7B12D7C1" lsidName="A. benningiorum" pageId="0" pageNumber="49" rank="species" species="benningiorum">
<emphasis id="43C95FBBF7666D16AB47C5AF027EAFD1" italics="true" pageId="0" pageNumber="49">A. benningiorum</emphasis>
</taxonomicName>
is phylogenetically and morphologically distinct from these species (Figure
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree generated from the maximum-likelihood analysis based on the combined ITS, LSU, tub 2 and rpb 2 sequence alignment of Didymellaceae members. The RAxML bootstrap support values (BS), Bayesian posterior probabilities (PP), and parsimony bootstrap support values (PBS) are given at the nodes (BS / PP / PBS). BS and PBS values represent parsimony bootstrap support values&gt; 50 %. Full supported branches are indicated in bold. The scale bar represents the expected number of changes per site. Ex-type strains are represented in bold. Strains obtained in the current study are printed in green; among them, whilst strains that represent new taxa are printed in red. Some of the basal branches were shortened to facilitate layout (the fraction in round parentheses refers to the presented length compared to the actual length of the branch). The tree was rooted to Coniothyrium palmarum CBS 400.71 and Leptosphaeria doliolum CBS 505.75." figureDoi="10.3897/mycokeys.65.47704.figure1" httpUri="https://binary.pensoft.net/fig/388204" pageId="0" pageNumber="49">1</figureCitation>
<figureCitation id="E8AB1775E2F88AAD6F7F9489C7B388B9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic tree generated from the maximum-likelihood analysis based on the combined ITS, LSU, tub 2 and rpb 2 sequence alignment of Didymellaceae members. The RAxML bootstrap support values (BS), Bayesian posterior probabilities (PP), and parsimony bootstrap support values (PBS) are given at the nodes (BS / PP / PBS). BS and PBS values represent parsimony bootstrap support values&gt; 50 %. Full supported branches are indicated in bold. The scale bar represents the expected number of changes per site. Ex-type strains are represented in bold. Strains obtained in the current study are printed in green; among them, whilst strains that represent new taxa are printed in red. Some of the basal branches were shortened to facilitate layout (the fraction in round parentheses refers to the presented length compared to the actual length of the branch). The tree was rooted to Coniothyrium palmarum CBS 400.71 and Leptosphaeria doliolum CBS 505.75." figureDoi="10.3897/mycokeys.65.47704.figure1" httpUri="https://binary.pensoft.net/fig/388204" pageId="0" pageNumber="49">1</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.1016/j.simyco.2015.10.003" author="Chen, Q" journalOrPublisher="Studies in Mycology" pageId="0" pageNumber="49" pagination="137 - 217" refId="B15" refString="Chen, Q, Jiang, JR, Zhang, GZ, Cai, L, Crous, PW, 2015. Resolving the Phoma enigma. Studies in Mycology 82: 137 - 217, DOI: https://doi.org/10.1016/j.simyco.2015.10.003" title="Resolving the Phoma enigma." url="https://doi.org/10.1016/j.simyco.2015.10.003" volume="82" year="2015">Chen et al. 2015</bibRefCitation>
<bibRefCitation id="C4C779EE639FD94A665B37D06886AF9A" DOI="https://doi.org/10.1016/j.simyco.2015.10.003" author="Chen, Q" journalOrPublisher="Studies in Mycology" pageId="0" pageNumber="49" pagination="137 - 217" refId="B15" refString="Chen, Q, Jiang, JR, Zhang, GZ, Cai, L, Crous, PW, 2015. Resolving the Phoma enigma. Studies in Mycology 82: 137 - 217, DOI: https://doi.org/10.1016/j.simyco.2015.10.003" title="Resolving the Phoma enigma." url="https://doi.org/10.1016/j.simyco.2015.10.003" volume="82" year="2015">Chen et al. 2015</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/mycokeys.65.47704.figure2" httpUri="https://binary.pensoft.net/fig/388205" pageId="0" pageNumber="49" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 2.</emphasis>
<taxonomicName authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum">
<emphasis italics="true" pageId="0" pageNumber="49">Ascochyta benningiorum</emphasis>
<caption id="27213020BB16BC8DD20CC542F058BA56" doi="10.3897/mycokeys.65.47704.figure2" httpUri="https://binary.pensoft.net/fig/388205" pageId="0" pageNumber="49" start="Figure 2" startId="F2">
<paragraph id="245140F47156C8A88E4653134EE626C2" pageId="0" pageNumber="49">
<emphasis id="6AE5C10E96B2F8FCAF203B920AC08653" bold="true" pageId="0" pageNumber="49">Figure 2.</emphasis>
<taxonomicName id="B981BD7902D25305939F8E40A57797AA" authorityName="Hern. - Restr., L. W. Hou, L. Cai &amp; Crous" authorityYear="2020" class="Dothideomycetes" family="Didymellaceae" genus="Ascochyta" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ascochyta benningiorum" order="Pleosporales" pageId="0" pageNumber="49" phylum="Ascomycota" rank="species" species="benningiorum">
<emphasis id="5AB6505ED35D3995D4B228F035DD7172" italics="true" pageId="0" pageNumber="49">Ascochyta benningiorum</emphasis>
</taxonomicName>
(CBS 144957).
<emphasis bold="true" pageId="0" pageNumber="49">A, B</emphasis>
<emphasis id="C801834304A19DF1B374C336A5F90864" bold="true" pageId="0" pageNumber="49">A, B</emphasis>
Colony on OA (front and reverse)
<emphasis bold="true" pageId="0" pageNumber="49">C, D</emphasis>
<emphasis id="A5C1573C0B923ABB7ECA5E7458F825D3" bold="true" pageId="0" pageNumber="49">C, D</emphasis>
colony on MEA (front and reverse)
<emphasis bold="true" pageId="0" pageNumber="49">E, F</emphasis>
<emphasis id="10494701C9651B4D2AAD9FCC1EE68561" bold="true" pageId="0" pageNumber="49">E, F</emphasis>
colony on PDA (front and reverse)
<emphasis bold="true" pageId="0" pageNumber="49">G</emphasis>
<emphasis id="0E78F241905553E31D20F76363D397EC" bold="true" pageId="0" pageNumber="49">G</emphasis>
pycnidia forming on OA
<emphasis bold="true" pageId="0" pageNumber="49">H</emphasis>
<emphasis id="4E3563A430B7FE68C4D0314170696D31" bold="true" pageId="0" pageNumber="49">H</emphasis>
pycnidium
<emphasis bold="true" pageId="0" pageNumber="49">I</emphasis>
<emphasis id="E72FCAA79500D8406402093845D53B62" bold="true" pageId="0" pageNumber="49">I</emphasis>
section of pycnidium
<emphasis bold="true" pageId="0" pageNumber="49">J</emphasis>
<emphasis id="8793790FD08F6AC964244CACF0377037" bold="true" pageId="0" pageNumber="49">J</emphasis>
section of pycnidial wall
<emphasis bold="true" pageId="0" pageNumber="49">K-M</emphasis>
<emphasis id="6D5F7B85168C5D8FE43988D6AD53B663" bold="true" pageId="0" pageNumber="49">K-M</emphasis>
conidiogenous cells
<emphasis bold="true" pageId="0" pageNumber="49">N</emphasis>
<emphasis id="882EC529C928CB3E819FA88874B1F8C0" bold="true" pageId="0" pageNumber="49">N</emphasis>
conidia. Scale bars: 100
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="F782FF3A1A0E197763FBFD738BC79CC5" originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="49">H, I</emphasis>
<emphasis id="00844E34B4F0B4C10FE9909F454B92A9" bold="true" pageId="0" pageNumber="49">H, I</emphasis>
); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="BD5EC1B829347F89E16E7E36FF558AAA" originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="49">J</emphasis>
<emphasis id="2B13B5EE972DF82F11C5BE8C1C748B51" bold="true" pageId="0" pageNumber="49">J</emphasis>
); 5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="4289F58C99BA5BEAE05F3520FA64ABBE" originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="49">K-N</emphasis>
<emphasis id="9F70F09D415F835F4A2228B4A0B2F001" bold="true" pageId="0" pageNumber="49">K-N</emphasis>
).
</paragraph>
</caption>