Add updates up until 2025-02-06 14:30:58

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<mods:title id="B4E1EB688554F2D9FECE351115365EE0">Revision of the genus Centrophorus (Squaliformes: Centrophoridae): Part 2 - Description of two new species of Centrophorus and clarification of the status of Centrophorus lusitanicus Barbosa du Bocage &amp; de Brito Capello, 1864</mods:title>
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<mods:title id="564346A314F4D7F80DA1A92EF2032BC4">Revision of the genus Centrophorus (Squaliformes: Centrophoridae): Part 2 - Description of two new species of Centrophorus and clarification of the status of Centrophorus lusitanicus Barbosa du Bocage &amp; de Brito Capello, 1864</mods:title>
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<subSubSection id="C3A3FBB82320FFC81ECBCF23667AF9C0" box="[151,575,1547,1572]" pageId="3" type="nomenclature">
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<heading id="D04E1F5F2320FFC81ECBCF23667AF9C0" bold="true" box="[151,575,1547,1572]" fontSize="10" level="2" pageId="3" reason="2">
<emphasis id="B9CD74212320FFC81ECBCF23667AF9C0" bold="true" box="[151,575,1547,1572]" pageId="3">
Validity of
<taxonomicName id="4CB9D3B02320FFC81F46CF23667AF9C0" box="[282,575,1547,1572]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81F46CF23667AF9C0" ID-CoL="69HN7" box="[282,575,1547,1572]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81F46CF23667AF9C0" bold="true" box="[282,575,1547,1572]" italics="true" pageId="3">Centrophorus lusitanicus</emphasis>
</taxonomicName>
</emphasis>
@ -63,11 +63,11 @@ Validity of
<subSubSection id="C3A3FBB82320FFCE1ECBCF666687FBD5" lastPageId="5" pageId="3" type="description">
<paragraph id="8B06A8332320FFC81ECBCF6667E8F936" blockId="3.[151,1436,1614,1747]" pageId="3">
The complicated nomenclatural history of
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<taxonomicName id="4CB9D3B02320FFC81C28CF67674BF982" box="[628,782,1614,1638]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81C28CF67674BF982" box="[628,782,1614,1638]" italics="true" pageId="3">C. lusitanicus</emphasis>
</taxonomicName>
, as discussed above, has hindered resolution of this group of gulper sharks. When considering Barbosa du Bocage &amp; de Brito Capellos (1864) description of
<taxonomicName id="4CB9D3B02320FFC81ABACF5B613AF96E" box="[1254,1407,1650,1674]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81ABACF5B613AF96E" box="[1254,1407,1650,1674]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81ABACF5B613AF96E" box="[1254,1407,1650,1674]" italics="true" pageId="3">C. lusitanicus</emphasis>
</taxonomicName>
in relation to the currently recognised species of
@ -88,40 +88,40 @@ in relation to the currently recognised species of
</paragraph>
<paragraph id="8B06A8332320FFC81ECBCE2B664DF8FF" blockId="3.[151,891,1794,2035]" box="[151,520,1794,1819]" pageId="3">
<taxonomicName id="4CB9D3B02320FFC81E9BCE2B6524F8FE" box="[199,353,1794,1818]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81E9BCE2B6524F8FE" box="[199,353,1794,1818]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81E9BCE2B6524F8FE" box="[199,353,1794,1818]" italics="true" pageId="3">C. lusitanicus</emphasis>
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is much larger
</paragraph>
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<taxonomicName id="4CB9D3B02320FFC81E9BCE0F6524F8DA" box="[199,353,1830,1854]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81E9BCE0F6524F8DA" box="[199,353,1830,1854]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81E9BCE0F6524F8DA" box="[199,353,1830,1854]" italics="true" pageId="3">C. lusitanicus</emphasis>
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has a blackish violet colour
</paragraph>
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• first dorsal fin base is longer in
<taxonomicName id="4CB9D3B02320FFC81C76CE636681F886" box="[554,708,1866,1890]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81C76CE636681F886" box="[554,708,1866,1890]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81C76CE636681F886" box="[554,708,1866,1890]" italics="true" pageId="3">C. lusitanicus</emphasis>
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• free rear tip of first dorsal fin more produced in
<taxonomicName id="4CB9D3B02320FFC81CBDCE47673EF862" box="[737,891,1902,1926]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81CBDCE47673EF862" box="[737,891,1902,1926]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81CBDCE47673EF862" box="[737,891,1902,1926]" italics="true" pageId="3">C. lusitanicus</emphasis>
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• dorsal spines shorter and weaker in
<taxonomicName id="4CB9D3B02320FFC81C04CEBB66B7F84E" box="[600,754,1938,1962]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81C04CEBB66B7F84E" box="[600,754,1938,1962]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81C04CEBB66B7F84E" box="[600,754,1938,1962]" italics="true" pageId="3">C. lusitanicus</emphasis>
</taxonomicName>
<taxonomicName id="4CB9D3B02320FFC81E9BCE9F6524F82A" box="[199,353,1974,1998]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81E9BCE9F6524F82A" box="[199,353,1974,1998]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81E9BCE9F6524F82A" box="[199,353,1974,1998]" italics="true" pageId="3">C. lusitanicus</emphasis>
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has a shorter snout
</paragraph>
<paragraph id="8B06A8332320FFC81ECBCEF3675DF816" blockId="3.[151,891,1794,2035]" box="[151,792,2010,2035]" pageId="3">
• pectoral-fin free tip is less produced in
<taxonomicName id="4CB9D3B02320FFC81C22CEF3675DF816" box="[638,792,2010,2034]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02320FFC81C22CEF3675DF816" box="[638,792,2010,2034]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="3" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212320FFC81C22CEF3675DF816" box="[638,792,2010,2034]" italics="true" pageId="3">C. lusitanicus</emphasis>
</taxonomicName>
</paragraph>
@ -137,9 +137,9 @@ These characteristics are perfectly consistent with the key diagnostic character
, which differs from
<emphasis id="B9CD74212327FFCF1B77C9C861D9FF13" box="[1323,1436,224,247]" italics="true" pageId="4">C. uyato</emphasis>
in having a much rougher skin. This information together with the character information provided in the description and the associated illustration (
<figureCitation id="1382B4B62327FFCF1C20C80F66F8FEA4" box="[636,701,295,320]" captionStart="FIGURE 1" captionStartId="4.[151,250,951,973]" captionTargetBox="[170,1403,643,904]" captionTargetId="figure@4.[151,1436,625,930]" captionTargetPageId="4" captionText="FIGURE 1. Lateral view of BMNH 1867.7. 23.2 (juvenile male 742 mm TL), previously considered to be a possible syntype of Centrophorus lusitanicus (now a paratype of Centrophorus lesliei n. sp.)." httpUri="https://zenodo.org/record/1042281/files/figure.png" pageId="4">Fig. 1</figureCitation>
<figureCitation id="1382B4B62327FFCF1C20C80F66F8FEA4" box="[636,701,295,320]" captionStart="FIGURE 1" captionStartId="4.[151,250,951,973]" captionTargetBox="[170,1403,643,904]" captionTargetId="figure@4.[151,1436,625,930]" captionTargetPageId="4" captionText="FIGURE 1. Lateral view of BMNH 1867.7. 23.2 (juvenile male 742 mm TL), previously considered to be a possible syntype of Centrophorus lusitanicus (now a paratype of Centrophorus lesliei n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.1042281" httpUri="https://zenodo.org/record/1042281/files/figure.png" pageId="4">Fig. 1</figureCitation>
) strongly suggest that
<taxonomicName id="4CB9D3B02327FFCF1D9FC800601BFEA4" box="[963,1118,296,320]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1D9FC800601BFEA4" box="[963,1118,296,320]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1D9FC800601BFEA4" box="[963,1118,296,320]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
is a synonym of the true
@ -153,7 +153,7 @@ is a synonym of the true
<emphasis id="B9CD74212327FFCF1CC7C8646773FE80" box="[667,822,332,356]" italics="true" pageId="4">Centrophorus</emphasis>
</taxonomicName>
species with the very long first-dorsal fin base that has been referred to as
<taxonomicName id="4CB9D3B02327FFCF1FDBC8586662FE6C" box="[391,551,368,392]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1FDBC8586662FE6C" box="[391,551,368,392]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1FDBC8586662FE6C" box="[391,551,368,392]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
in recent literature. Based on our current understanding, three species of
@ -185,11 +185,11 @@ and
[=
<emphasis id="B9CD74212327FFCF1A8BC890616DFE2B" box="[1239,1320,440,463]" italics="true" pageId="4">uyato</emphasis>
] and their new
<taxonomicName id="4CB9D3B02327FFCF1E92C8F4652CFE10" box="[206,361,476,500]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1E92C8F4652CFE10" box="[206,361,476,500]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1E92C8F4652CFE10" box="[206,361,476,500]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
. Given the scope of Barbuso du Bocage &amp; de Brito Capellos paper, it seems unlikely that the authors would have missed this common species, thus lending more support to the claim that
<taxonomicName id="4CB9D3B02327FFCF1A92CB28612EFDFC" box="[1230,1387,512,536]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1A92CB28612EFDFC" box="[1230,1387,512,536]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1A92CB28612EFDFC" box="[1230,1387,512,536]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
is a synonym of the true
@ -198,15 +198,15 @@ is a synonym of the true
</taxonomicName>
.
</paragraph>
<caption id="DFC6F8BB2327FFCF1ECBCA9F673CFC0F" httpUri="https://zenodo.org/record/1042281/files/figure.png" pageId="4" targetBox="[170,1403,643,904]" targetPageId="4">
<caption id="DFC6F8BB2327FFCF1ECBCA9F673CFC0F" ID-DOI="http://doi.org/10.5281/zenodo.1042281" httpUri="https://zenodo.org/record/1042281/files/figure.png" pageId="4" targetBox="[170,1403,643,904]" targetPageId="4">
<paragraph id="8B06A8332327FFCF1ECBCA9F673CFC0F" blockId="4.[151,1436,951,1003]" pageId="4">
<emphasis id="B9CD74212327FFCF1ECBCA9F6557FC29" bold="true" box="[151,274,951,973]" pageId="4">FIGURE 1.</emphasis>
Lateral view of BMNH 1867.7.23.2 (juvenile male 742 mm TL), previously considered to be a possible syntype of
<taxonomicName id="4CB9D3B02327FFCF1ECBCAFD65D3FC0F" box="[151,406,981,1003]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1ECBCAFD65D3FC0F" box="[151,406,981,1003]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1ECBCAFD65D3FC0F" box="[151,406,981,1003]" italics="true" pageId="4">Centrophorus lusitanicus</emphasis>
</taxonomicName>
(now a paratype of
<taxonomicName id="4CB9D3B02327FFCF1C3ECAFD6775FC0F" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[610,816,981,1003]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lesliei" status="sp. nov.">
<taxonomicName id="4CB9D3B02327FFCF1C3ECAFD6775FC0F" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[610,816,981,1003]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordara" rank="species" species="lesliei" status="sp. nov.">
<emphasis id="B9CD74212327FFCF1C3ECAFD6775FC0F" box="[610,816,981,1003]" italics="true" pageId="4">Centrophorus lesliei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2FEC95A2327FFCF1D6ACAFD672CFC0F" box="[822,873,981,1003]" pageId="4" rank="species">
@ -221,11 +221,11 @@ Since the authors refer to the fresh condition, the
locality from off
<collectingCountry id="F3AEE8A32327FFCF1A59CD3B6021FBC8" box="[1029,1124,1043,1068]" name="Portugal" pageId="4">Portugal</collectingCountry>
for
<taxonomicName id="4CB9D3B02327FFCF1ACBCD3C6171FBCF" box="[1175,1332,1043,1067]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1ACBCD3C6171FBCF" box="[1175,1332,1043,1067]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1ACBCD3C6171FBCF" box="[1175,1332,1043,1067]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
is not in question. However, the surviving apparent syntype of
<taxonomicName id="4CB9D3B02327FFCF1D79CD106786FBAB" box="[805,963,1079,1103]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1D79CD106786FBAB" box="[805,963,1079,1103]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1D79CD106786FBAB" box="[805,963,1079,1103]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
has long been considered to be BMNH 1867.7.23.2 and is clearly not conspecific with the true
@ -233,15 +233,15 @@ has long been considered to be BMNH 1867.7.23.2 and is clearly not conspecific w
<emphasis id="B9CD74212327FFCF1D57CD7467EDFB97" box="[779,936,1115,1139]" italics="true" pageId="4">C. granulosus</emphasis>
</taxonomicName>
(
<figureCitation id="1382B4B62327FFCF1DE5CD7367B9FB90" box="[953,1020,1115,1140]" captionStart="FIGURE 2" captionStartId="5.[151,250,574,596]" captionTargetBox="[351,1220,324,517]" captionTargetId="figure@5.[340,1247,301,553]" captionTargetPageId="5" captionText="FIGURE 2. Illustration of Centrophorus lusitanicus in the original description (Barbosa du Bocage &amp; Brito Capello, 1864)." httpUri="https://zenodo.org/record/1042283/files/figure.png" pageId="4">Fig. 2</figureCitation>
<figureCitation id="1382B4B62327FFCF1DE5CD7367B9FB90" box="[953,1020,1115,1140]" captionStart="FIGURE 2" captionStartId="5.[151,250,574,596]" captionTargetBox="[351,1220,324,517]" captionTargetId="figure@5.[340,1247,301,553]" captionTargetPageId="5" captionText="FIGURE 2. Illustration of Centrophorus lusitanicus in the original description (Barbosa du Bocage &amp; Brito Capello, 1864)." figureDoi="http://doi.org/10.5281/zenodo.1042283" httpUri="https://zenodo.org/record/1042283/files/figure.png" pageId="4">Fig. 2</figureCitation>
) as it has a much longer first dorsalfin base. This led to an investigation into the
<typeStatus id="540216912327FFCF1CC8CDA86681FB7C" box="[660,708,1152,1176]" pageId="4">type</typeStatus>
status of
<taxonomicName id="4CB9D3B02327FFCF1D6DCDA8678EFB73" box="[817,971,1151,1175]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1D6DCDA8678EFB73" box="[817,971,1151,1175]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1D6DCDA8678EFB73" box="[817,971,1151,1175]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
noting that no reference was made in the original description of
<taxonomicName id="4CB9D3B02327FFCF1FCACD8C6675FB5F" box="[406,560,1187,1211]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1FCACD8C6675FB5F" box="[406,560,1187,1211]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1FCACD8C6675FB5F" box="[406,560,1187,1211]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
to any
@ -249,13 +249,13 @@ to any
specimens.
<bibRefCitation id="EF28D5C22327FFCF1D60CD8B67ACFB58" author="Gunther" box="[828,1001,1187,1212]" pageId="4" refString="Gunther, A. (1870) Catalogue of the fishes in the British Museum. In: Catalogue of the Physostomi, containing the families Gymnotidae, Symbranchidae, Muraenidae, Pegasidae, and of the Lophobranchii, Plectognathi, Dipnoi, ... [thru] ... Leptocardii, in collection of the British Museum, 8, pp. 1 - 549." type="journal article" year="1870">Günther (1870)</bibRefCitation>
appears to be the first to link the name
<taxonomicName id="4CB9D3B02327FFCF1ECBCDE06575FB3B" box="[151,304,1223,1247]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1ECBCDE06575FB3B" box="[151,304,1223,1247]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1ECBCDE06575FB3B" box="[151,304,1223,1247]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
with a specimen of the long-finned species in the British Museum, referring to a male of
<quantity id="4C4105D62327FFCF1B4ACDEF613AFB04" box="[1302,1407,1223,1248]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" pageId="4" unit="in" value="29.0">29 inches</quantity>
in length. This corresponds to the size and sex of BMNH 1867.7.23.2 (
<figureCitation id="1382B4B62327FFCF1DF2CDC367B0FAE0" box="[942,1013,1259,1284]" captionStart="FIGURE 2" captionStartId="5.[151,250,574,596]" captionTargetBox="[351,1220,324,517]" captionTargetId="figure@5.[340,1247,301,553]" captionTargetPageId="5" captionText="FIGURE 2. Illustration of Centrophorus lusitanicus in the original description (Barbosa du Bocage &amp; Brito Capello, 1864)." httpUri="https://zenodo.org/record/1042283/files/figure.png" pageId="4">Fig. 2</figureCitation>
<figureCitation id="1382B4B62327FFCF1DF2CDC367B0FAE0" box="[942,1013,1259,1284]" captionStart="FIGURE 2" captionStartId="5.[151,250,574,596]" captionTargetBox="[351,1220,324,517]" captionTargetId="figure@5.[340,1247,301,553]" captionTargetPageId="5" captionText="FIGURE 2. Illustration of Centrophorus lusitanicus in the original description (Barbosa du Bocage &amp; Brito Capello, 1864)." figureDoi="http://doi.org/10.5281/zenodo.1042283" httpUri="https://zenodo.org/record/1042283/files/figure.png" pageId="4">Fig. 2</figureCitation>
).
<bibRefCitation id="EF28D5C22327FFCF1A50CDC360F9FAE0" author="Gunther" box="[1036,1212,1259,1284]" pageId="4" refString="Gunther, A. (1870) Catalogue of the fishes in the British Museum. In: Catalogue of the Physostomi, containing the families Gymnotidae, Symbranchidae, Muraenidae, Pegasidae, and of the Lophobranchii, Plectognathi, Dipnoi, ... [thru] ... Leptocardii, in collection of the British Museum, 8, pp. 1 - 549." type="journal article" year="1870">Günther (1870)</bibRefCitation>
does not, however, refer to this specimen as a
@ -273,13 +273,13 @@ specimen. The original documentation of this specimen entering the Natural Histo
<emphasis id="B9CD74212327FFCF1BDECCEC6556FA1B" italics="true" pageId="4">C. granulosus</emphasis>
</taxonomicName>
, and not as
<taxonomicName id="4CB9D3B02327FFCF1FF4CCC06600FA1B" box="[424,581,1511,1535]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02327FFCF1FF4CCC06600FA1B" box="[424,581,1511,1535]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="4" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212327FFCF1FF4CCC06600FA1B" box="[424,581,1511,1535]" italics="true" pageId="4">C. lusitanicus</emphasis>
</taxonomicName>
. The final aspect which has led to much confusion is the location of the specimen as
<collectingCountry id="F3AEE8A32327FFCF1F75CF2365CEF9C0" box="[297,395,1547,1572]" name="Portugal" pageId="4">Portugal</collectingCountry>
. The original ledger (
<figureCitation id="1382B4B62327FFCF1CDFCF236682F9C0" box="[643,711,1547,1572]" captionStart="FIGURE 3" captionStartId="5.[151,250,777,799]" captionTargetBox="[340,1247,646,755]" captionTargetId="figure@5.[340,1247,646,755]" captionTargetPageId="5" captionText="FIGURE 3. Excerpt from the specimen register showing the batch of three sharks, including the possible syntype of Centrophorus lusitanicus (as C granulosus) BMNH 1867.7. 23.2, donated by Bocage. Note the lack of ditto marks beneath Portugal for Centrophorus granulosus and no mention of it being a type." httpUri="https://zenodo.org/record/1042285/files/figure.png" pageId="4">Fig. 3</figureCitation>
<figureCitation id="1382B4B62327FFCF1CDFCF236682F9C0" box="[643,711,1547,1572]" captionStart="FIGURE 3" captionStartId="5.[151,250,777,799]" captionTargetBox="[340,1247,646,755]" captionTargetId="figure@5.[340,1247,646,755]" captionTargetPageId="5" captionText="FIGURE 3. Excerpt from the specimen register showing the batch of three sharks, including the possible syntype of Centrophorus lusitanicus (as C granulosus) BMNH 1867.7. 23.2, donated by Bocage. Note the lack of ditto marks beneath Portugal for Centrophorus granulosus and no mention of it being a type." figureDoi="http://doi.org/10.5281/zenodo.1042285" httpUri="https://zenodo.org/record/1042285/files/figure.png" pageId="4">Fig. 3</figureCitation>
) includes three specimens donated from Bocage,
<taxonomicName id="4CB9D3B02327FFCF1B5DCF23655FF9A3" authorityName="Barbosa du Bocage &amp; de Brito Capello" authorityYear="1864" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="crepidalbus">
<emphasis id="B9CD74212327FFCF1B5DCF23655FF9A3" italics="true" pageId="4">Centrophorus crepidalbus</emphasis>
@ -322,7 +322,7 @@ and
(
<bibRefCitation id="EF28D5C22327FFCF1EC2CEBF6517F84B" author="Madruga" box="[158,338,1943,1968]" pageId="4" refString="Madruga, C. (2013) Jose Ficente Barbosa du Bocage (1823 - 1907). A construcao de uma persona cientifica. Dissertacao, Mestrado em Historia e Filosofia das Ciencias, Universidade de Lisboa, Lisbon, 109 pp." type="book chapter" year="2013">Madruga, 2013</bibRefCitation>
). This information, combined with the fact that the long-finned species is well known from this part of Africa, provides reasonable evidence that the BMNH (and ZMB) specimens were likely collected from The weight of the available evidence suggests that
<taxonomicName id="4CB9D3B02326FFCE1D40CAA167FCFC45" box="[796,953,905,929]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02326FFCE1D40CAA167FCFC45" box="[796,953,905,929]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="5" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212326FFCE1D40CAA167FCFC45" box="[796,953,905,929]" italics="true" pageId="5">C. lusitanicus</emphasis>
</taxonomicName>
refers to the true
@ -334,13 +334,13 @@ from off
and so must be considered a junior synonym of the latter species. The purported
<typeStatus id="540216912326FFCE1AD9CA866099FC22" box="[1157,1244,942,966]" pageId="5" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4CB9D3B02326FFCE1B5ECA8561D9FC21" box="[1282,1436,941,965]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02326FFCE1B5ECA8561D9FC21" box="[1282,1436,941,965]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="5" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212326FFCE1B5ECA8561D9FC21" box="[1282,1436,941,965]" italics="true" pageId="5">C. lusitanicus</emphasis>
</taxonomicName>
(BMNH 1867.7.23.2) is not a
<typeStatus id="540216912326FFCE1C5FCAF96676FC0D" box="[515,563,977,1001]" pageId="5">type</typeStatus>
specimen and refers to a separate long-finned species. A second Bocage specimen of the same long-finned species in the Museum für Naturkunde (ZMB 6455,
<figureCitation id="1382B4B62326FFCE1A27CADC6085FBE9" box="[1147,1216,1012,1037]" captionStart="FIGURE 4" captionStartId="6.[151,250,521,543]" captionTargetBox="[164,1403,205,485]" captionTargetId="figure@6.[151,1436,193,500]" captionTargetPageId="6" captionText="FIGURE 4. Lateral view of ZMB 6455 (female 644 mm TL), a second specimen donated from the Lisbon Museum by Bocage as Centrophorus granulosus (now a paratype of Centrophorus lesliei n. sp.)." httpUri="https://zenodo.org/record/1042287/files/figure.png" pageId="5">Fig. 4</figureCitation>
<figureCitation id="1382B4B62326FFCE1A27CADC6085FBE9" box="[1147,1216,1012,1037]" captionStart="FIGURE 4" captionStartId="6.[151,250,521,543]" captionTargetBox="[164,1403,205,485]" captionTargetId="figure@6.[151,1436,193,500]" captionTargetPageId="6" captionText="FIGURE 4. Lateral view of ZMB 6455 (female 644 mm TL), a second specimen donated from the Lisbon Museum by Bocage as Centrophorus granulosus (now a paratype of Centrophorus lesliei n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.1042287" httpUri="https://zenodo.org/record/1042287/files/figure.png" pageId="5">Fig. 4</figureCitation>
), is likely from the same collection location as the BMNH specimen.
</paragraph>
</subSubSection>

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@ -1,58 +1,58 @@
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<mods:title id="3A6BFC7FC67756810B4EA2065D0D48FA">Revision of the genus Centrophorus (Squaliformes: Centrophoridae): Part 2 - Description of two new species of Centrophorus and clarification of the status of Centrophorus lusitanicus Barbosa du Bocage &amp; de Brito Capello, 1864</mods:title>
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<mods:title id="0BEEA1E5E24D5A0DEF4AD7A1461A569F">Revision of the genus Centrophorus (Squaliformes: Centrophoridae): Part 2 - Description of two new species of Centrophorus and clarification of the status of Centrophorus lusitanicus Barbosa du Bocage &amp; de Brito Capello, 1864</mods:title>
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<figureCitation id="1382B4B6232DFFC51EC3C9F6655AFF13" box="[159,287,222,247]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStartId-0="15.[151,250,1818,1840]" captionStartId-1="16.[151,250,1054,1076]" captionStartId-2="16.[151,250,1585,1607]" captionStartId-3="18.[151,250,1956,1978]" captionStartId-4="19.[151,250,1034,1056]" captionTargetBox-0="[158,1410,262,1749]" captionTargetBox-1="[173,1436,205,1014]" captionTargetBox-2="[314,1273,1156,1563]" captionTargetBox-3="[234,1368,204,1914]" captionTargetBox-4="[333,1232,208,989]" captionTargetId-0="figure@15.[151,1436,193,1796]" captionTargetId-1="figure@16.[151,1436,193,1033]" captionTargetId-2="figure@16.[314,1273,1156,1563]" captionTargetId-3="figure@18.[192,1394,193,1935]" captionTargetId-4="figure@19.[331,1255,193,1013]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="16" captionTargetPageId-3="18" captionTargetPageId-4="19" captionText-0="FIGURE 11. Lateral view of Centrophorus longipinnis n. sp.: (A) adult male holotype (NMMB-P 15756, 720 mm TL); (B) female paratype (CSIRO H 8104 - 01, 855 mm TL); (C) female paratype (CSIRO H 7990 - 02, 825 mm TL); (D) late-term embryo paratype (CSIRO H 8103 - 02, 346 mm TL)." captionText-1="FIGURE 12. Ventral view of head and pectoral fins of Centrophorus longipinnis n. sp.: female paratype (CSIRO H 7990 - 02, 825 mm TL)." captionText-2="FIGURE 13. Upper and lower teeth of female paratype of Centrophorus longipinnis n. sp. (CSIRO H 7990 - 02, 825 mm TL)." captionText-3="FIGURE 14. Lateral trunk denticles (from below first dorsal fin) of Centrophorus longipinnis n. sp.: (A) juvenile female paratype, NMMB-P 14051, 408 mm TL; (B) female paratype, CSIRO H 8104 - 01, 855 mm TL." captionText-4="FIGURE 15. Clasper (right) of Centrophorus longipinnis (paratype CSIRO H 8104 02, adult male 679 mm TL). A, Glans not dilated; B, Glans spread. Abbreviations: AP, apopyle; CG, clasper groove; CS, clasper spine; HP, hypopyle; P 2, pelvic fin; RH, rhipidion." httpUri-0="https://zenodo.org/record/1042301/files/figure.png" httpUri-1="https://zenodo.org/record/1042303/files/figure.png" httpUri-2="https://zenodo.org/record/1042305/files/figure.png" httpUri-3="https://zenodo.org/record/1042307/files/figure.png" httpUri-4="https://zenodo.org/record/1042309/files/figure.png" pageId="14">Figs 1115</figureCitation>
<figureCitation id="1382B4B6232DFFC51EC3C9F6655AFF13" box="[159,287,222,247]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStart-3="FIGURE 14" captionStart-4="FIGURE 15" captionStartId-0="15.[151,250,1818,1840]" captionStartId-1="16.[151,250,1054,1076]" captionStartId-2="16.[151,250,1585,1607]" captionStartId-3="18.[151,250,1956,1978]" captionStartId-4="19.[151,250,1034,1056]" captionTargetBox-0="[158,1410,262,1749]" captionTargetBox-1="[173,1436,205,1014]" captionTargetBox-2="[314,1273,1156,1563]" captionTargetBox-3="[234,1368,204,1914]" captionTargetBox-4="[333,1232,208,989]" captionTargetId-0="figure@15.[151,1436,193,1796]" captionTargetId-1="figure@16.[151,1436,193,1033]" captionTargetId-2="figure@16.[314,1273,1156,1563]" captionTargetId-3="figure@18.[192,1394,193,1935]" captionTargetId-4="figure@19.[331,1255,193,1013]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="16" captionTargetPageId-3="18" captionTargetPageId-4="19" captionText-0="FIGURE 11. Lateral view of Centrophorus longipinnis n. sp.: (A) adult male holotype (NMMB-P 15756, 720 mm TL); (B) female paratype (CSIRO H 8104 - 01, 855 mm TL); (C) female paratype (CSIRO H 7990 - 02, 825 mm TL); (D) late-term embryo paratype (CSIRO H 8103 - 02, 346 mm TL)." captionText-1="FIGURE 12. Ventral view of head and pectoral fins of Centrophorus longipinnis n. sp.: female paratype (CSIRO H 7990 - 02, 825 mm TL)." captionText-2="FIGURE 13. Upper and lower teeth of female paratype of Centrophorus longipinnis n. sp. (CSIRO H 7990 - 02, 825 mm TL)." captionText-3="FIGURE 14. Lateral trunk denticles (from below first dorsal fin) of Centrophorus longipinnis n. sp.: (A) juvenile female paratype, NMMB-P 14051, 408 mm TL; (B) female paratype, CSIRO H 8104 - 01, 855 mm TL." captionText-4="FIGURE 15. Clasper (right) of Centrophorus longipinnis (paratype CSIRO H 8104 02, adult male 679 mm TL). A, Glans not dilated; B, Glans spread. Abbreviations: AP, apopyle; CG, clasper groove; CS, clasper spine; HP, hypopyle; P 2, pelvic fin; RH, rhipidion." figureDoi-0="http://doi.org/10.5281/zenodo.1042301" figureDoi-1="http://doi.org/10.5281/zenodo.1042303" figureDoi-2="http://doi.org/10.5281/zenodo.1042305" figureDoi-3="http://doi.org/10.5281/zenodo.1042307" figureDoi-4="http://doi.org/10.5281/zenodo.1042309" httpUri-0="https://zenodo.org/record/1042301/files/figure.png" httpUri-1="https://zenodo.org/record/1042303/files/figure.png" httpUri-2="https://zenodo.org/record/1042305/files/figure.png" httpUri-3="https://zenodo.org/record/1042307/files/figure.png" httpUri-4="https://zenodo.org/record/1042309/files/figure.png" pageId="14">Figs 1115</figureCitation>
;
<tableCitation id="C63B9D88232DFFC51F77C9F6653AFF13" box="[299,383,222,247]" captionStart="TABLE 2" captionStartId="20.[151,239,329,352]" captionText="TABLE 2. Proportional dimensions as percentages of total length for the holotype of Centrophorus longipinnis (NMMB-P 15756), and ranges for small paratypes (&lt;524 mm TL) and large paratypes (&gt; 700 mm TL)." pageId="14">Table 2</tableCitation>
)
@ -74,7 +74,7 @@
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<emphasis id="B9CD7421232DFFC51ECBC80E65DCFED8" box="[151,409,294,316]" italics="true" pageId="14">Centrophorus lusitanicus</emphasis>
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: 205
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<figureCitation id="1382B4B6232DFFC51C19C86D66C4FEBF" box="[581,641,325,347]" captionStart="FIGURE 2" captionStartId="5.[151,250,574,596]" captionTargetBox="[351,1220,324,517]" captionTargetId="figure@5.[340,1247,301,553]" captionTargetPageId="5" captionText="FIGURE 2. Illustration of Centrophorus lusitanicus in the original description (Barbosa du Bocage &amp; Brito Capello, 1864)." httpUri="https://zenodo.org/record/1042283/files/figure.png" pageId="14">Fig. 2</figureCitation>
<figureCitation id="1382B4B6232DFFC51C19C86D66C4FEBF" box="[581,641,325,347]" captionStart="FIGURE 2" captionStartId="5.[151,250,574,596]" captionTargetBox="[351,1220,324,517]" captionTargetId="figure@5.[340,1247,301,553]" captionTargetPageId="5" captionText="FIGURE 2. Illustration of Centrophorus lusitanicus in the original description (Barbosa du Bocage &amp; Brito Capello, 1864)." figureDoi="http://doi.org/10.5281/zenodo.1042283" httpUri="https://zenodo.org/record/1042283/files/figure.png" pageId="14">Fig. 2</figureCitation>
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</treatmentCitation>
,
<figureCitation id="1382B4B6232DFFC51A57C86D6003FEBF" box="[1035,1094,325,347]" captionStart="FIGURE 1" captionStartId="4.[151,250,951,973]" captionTargetBox="[170,1403,643,904]" captionTargetId="figure@4.[151,1436,625,930]" captionTargetPageId="4" captionText="FIGURE 1. Lateral view of BMNH 1867.7. 23.2 (juvenile male 742 mm TL), previously considered to be a possible syntype of Centrophorus lusitanicus (now a paratype of Centrophorus lesliei n. sp.)." httpUri="https://zenodo.org/record/1042281/files/figure.png" pageId="14">Fig. 1</figureCitation>
<figureCitation id="1382B4B6232DFFC51A57C86D6003FEBF" box="[1035,1094,325,347]" captionStart="FIGURE 1" captionStartId="4.[151,250,951,973]" captionTargetBox="[170,1403,643,904]" captionTargetId="figure@4.[151,1436,625,930]" captionTargetPageId="4" captionText="FIGURE 1. Lateral view of BMNH 1867.7. 23.2 (juvenile male 742 mm TL), previously considered to be a possible syntype of Centrophorus lusitanicus (now a paratype of Centrophorus lesliei n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.1042281" httpUri="https://zenodo.org/record/1042281/files/figure.png" pageId="14">Fig. 1</figureCitation>
(
<collectingRegion id="497D66D1232DFFC51A08C86D60EEFEBF" box="[1108,1195,325,347]" country="China" name="Taiwan)" pageId="14">
<collectingCountry id="F3AEE8A3232DFFC51A08C86D60E1FEBF" box="[1108,1188,325,347]" name="Taiwan" pageId="14">Taiwan</collectingCountry>
@ -196,7 +196,7 @@ White
: 1364
</treatmentCitation>
,
<figureCitation id="1382B4B6232DFFC51AE6C8E8615EFE32" box="[1210,1307,448,470]" captionStart="FIGURE 6" captionStartId="7.[151,250,1049,1071]" captionTargetBox="[179,1436,203,1013]" captionTargetId="figure@7.[151,1436,193,1028]" captionTargetPageId="7" captionText="FIGURE 6. Ventral view of head of the pregnant female holotype of Centrophorus lesliei n. sp. (SAMC-F 041921, 863 mm TL)." httpUri="https://zenodo.org/record/1042291/files/figure.png" pageId="14">Fig. 6a, b</figureCitation>
<figureCitation id="1382B4B6232DFFC51AE6C8E8615EFE32" box="[1210,1307,448,470]" captionStart="FIGURE 6" captionStartId="7.[151,250,1049,1071]" captionTargetBox="[179,1436,203,1013]" captionTargetId="figure@7.[151,1436,193,1028]" captionTargetPageId="7" captionText="FIGURE 6. Ventral view of head of the pregnant female holotype of Centrophorus lesliei n. sp. (SAMC-F 041921, 863 mm TL)." figureDoi="http://doi.org/10.5281/zenodo.1042291" httpUri="https://zenodo.org/record/1042291/files/figure.png" pageId="14">Fig. 6a, b</figureCitation>
(Indonesia); Ebert
<emphasis id="B9CD7421232DFFC51F58C8C86575FE10" box="[260,304,478,501]" italics="true" pageId="14">et al</emphasis>
., 2013: 289 (Taiwan).
@ -421,11 +421,11 @@ TL
<quantity id="4C4105D6232DFFC51C77CFF766C8F912" box="[555,653,1759,1783]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="14" unit="mm" value="700.0">700 mm</quantity>
TL) in TL; interdorsal space 1.28 (1.61; 1.031.38) in prepectoral length, 1.45 (1.69; 1.281.62) in pre-first dorsal length; pelviccaudal space 3.28 (2.60; 2.703.50) in pectoral pelvic space, 1.82 (1.87; 1.622.00) in prepectoral length; dorsalcaudal space 2.05 (1.81; 2.092.65) in interdorsal space. Caudal peduncle moderately short and deep, moderately compressed, its length 11.5 (12.4; 10.712.8)% TL, its height 1.47 (2.11; 1.612.15) times its width; tapering slightly towards caudal fin; ventral midline with a groove; dorsal midline with a weak ridge anteriorly becoming a weak groove posteriorly; no lateral keels; precaudal pits absent.
</paragraph>
<caption id="DFC6F8BB232CFFC41ECBCE3266E6F889" httpUri="https://zenodo.org/record/1042301/files/figure.png" pageId="15" targetBox="[158,1410,262,1749]" targetPageId="15">
<caption id="DFC6F8BB232CFFC41ECBCE3266E6F889" ID-DOI="http://doi.org/10.5281/zenodo.1042301" httpUri="https://zenodo.org/record/1042301/files/figure.png" pageId="15" targetBox="[158,1410,262,1749]" targetPageId="15">
<paragraph id="8B06A833232CFFC41ECBCE3266E6F889" blockId="15.[151,1436,1818,1901]" pageId="15">
<emphasis id="B9CD7421232CFFC41ECBCE326565F8D4" bold="true" box="[151,288,1818,1840]" pageId="15">FIGURE 11.</emphasis>
Lateral view of
<taxonomicName id="4CB9D3B0232CFFC41F93CE326690F8D4" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[463,725,1818,1840]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="15" phylum="Arthropoda" rank="species" species="longipinnis" status="sp. nov.">
<taxonomicName id="4CB9D3B0232CFFC41F93CE326690F8D4" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[463,725,1818,1840]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="15" phylum="Chordata" rank="species" species="longipinnis" status="sp. nov.">
<emphasis id="B9CD7421232CFFC41F93CE326690F8D4" box="[463,725,1818,1840]" italics="true" pageId="15">Centrophorus longipinnis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2FEC95A232CFFC41C81CE326755F8D4" box="[733,784,1818,1840]" pageId="15" rank="species">
@ -440,11 +440,11 @@ H 7990-02, 825 mm TL); (D) late-term embryo paratype (
H 8103-02, 346 mm TL).
</paragraph>
</caption>
<caption id="DFC6F8BB2333FFDB1ECBCD36655AFBB6" httpUri="https://zenodo.org/record/1042303/files/figure.png" pageId="16" targetBox="[173,1436,205,1014]" targetPageId="16">
<caption id="DFC6F8BB2333FFDB1ECBCD36655AFBB6" ID-DOI="http://doi.org/10.5281/zenodo.1042303" httpUri="https://zenodo.org/record/1042303/files/figure.png" pageId="16" targetBox="[173,1436,205,1014]" targetPageId="16">
<paragraph id="8B06A8332333FFDB1ECBCD36655AFBB6" blockId="16.[151,1436,1054,1106]" pageId="16">
<emphasis id="B9CD74212333FFDB1ECBCD366565FBD0" bold="true" box="[151,288,1054,1076]" pageId="16">FIGURE 12.</emphasis>
Ventral view of head and pectoral fins of
<taxonomicName id="4CB9D3B02333FFDB1C88CD36679CFBD0" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[724,985,1054,1076]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="longipinnis" status="sp. nov.">
<taxonomicName id="4CB9D3B02333FFDB1C88CD36679CFBD0" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[724,985,1054,1076]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="16" phylum="Chordata" rank="species" species="longipinnis" status="sp. nov.">
<emphasis id="B9CD74212333FFDB1C88CD36679CFBD0" box="[724,985,1054,1076]" italics="true" pageId="16">Centrophorus longipinnis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2FEC95A2333FFDB1DBDCD366056FBD0" box="[993,1043,1054,1076]" pageId="16" rank="species">
@ -455,11 +455,11 @@ Ventral view of head and pectoral fins of
H 7990-02, 825 mm TL).
</paragraph>
</caption>
<caption id="DFC6F8BB2333FFDB1ECBCF1961CEF9A3" box="[151,1419,1585,1607]" httpUri="https://zenodo.org/record/1042305/files/figure.png" pageId="16" targetBox="[314,1273,1156,1563]" targetPageId="16">
<caption id="DFC6F8BB2333FFDB1ECBCF1961CEF9A3" ID-DOI="http://doi.org/10.5281/zenodo.1042305" box="[151,1419,1585,1607]" httpUri="https://zenodo.org/record/1042305/files/figure.png" pageId="16" targetBox="[314,1273,1156,1563]" targetPageId="16">
<paragraph id="8B06A8332333FFDB1ECBCF1961CEF9A3" blockId="16.[151,1419,1585,1607]" box="[151,1419,1585,1607]" pageId="16">
<emphasis id="B9CD74212333FFDB1ECBCF19655AF9A3" bold="true" box="[151,287,1585,1607]" pageId="16">FIGURE 13.</emphasis>
Upper and lower teeth of female paratype of
<taxonomicName id="4CB9D3B02333FFDB1CB1CF1967B4F9A3" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[749,1009,1585,1607]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="longipinnis" status="sp. nov.">
<taxonomicName id="4CB9D3B02333FFDB1CB1CF1967B4F9A3" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[749,1009,1585,1607]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="16" phylum="Chordata" rank="species" species="longipinnis" status="sp. nov.">
<emphasis id="B9CD74212333FFDB1CB1CF1967B4F9A3" box="[749,1009,1585,1607]" italics="true" pageId="16">Centrophorus longipinnis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2FEC95A2333FFDB1DABCF19606FF9A3" box="[1015,1066,1585,1607]" pageId="16" rank="species">
@ -476,9 +476,9 @@ H 7990-02, 825 mm TL).
Mouth almost transverse, upper jaw slightly concave, width 1.22 (1.29; 1.061.22) in preoral length; lower labial furrows slightly longer than upper furrows; prominent postoral groove, more than twice length of upper labial furrows, extending posterolaterally from angle of jaws. Teeth strongly differentiated in upper and lower jaws, with upper teeth much smaller than lower teeth. Upper teeth of adults (based on
<taxonomicName id="4CB9D3B02332FFDA1A6AC8F460CCFE10" box="[1078,1161,476,500]" class="Insecta" family="Tenebrionidae" genus="Csiro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus">CSIRO</taxonomicName>
H 7990-02) moderately large, with erect cusps, becoming slightly oblique posteriorly towards mouth corners, bases slightly overlapping (
<figureCitation id="1382B4B62332FFDA1EC3CB0C6545FDD8" box="[159,256,548,573]" captionStart="FIGURE 13" captionStartId="16.[151,250,1585,1607]" captionTargetBox="[314,1273,1156,1563]" captionTargetId="figure@16.[314,1273,1156,1563]" captionTargetPageId="16" captionText="FIGURE 13. Upper and lower teeth of female paratype of Centrophorus longipinnis n. sp. (CSIRO H 7990 - 02, 825 mm TL)." httpUri="https://zenodo.org/record/1042305/files/figure.png" pageId="17">Fig. 13a</figureCitation>
<figureCitation id="1382B4B62332FFDA1EC3CB0C6545FDD8" box="[159,256,548,573]" captionStart="FIGURE 13" captionStartId="16.[151,250,1585,1607]" captionTargetBox="[314,1273,1156,1563]" captionTargetId="figure@16.[314,1273,1156,1563]" captionTargetPageId="16" captionText="FIGURE 13. Upper and lower teeth of female paratype of Centrophorus longipinnis n. sp. (CSIRO H 7990 - 02, 825 mm TL)." figureDoi="http://doi.org/10.5281/zenodo.1042305" httpUri="https://zenodo.org/record/1042305/files/figure.png" pageId="17">Fig. 13a</figureCitation>
). Lower teeth much larger than uppers, cusps very strongly oblique, blade-like, overlapping, edges with fine serrations (
<figureCitation id="1382B4B62332FFDA1F19CB6F65E2FD84" box="[325,423,583,608]" captionStart="FIGURE 13" captionStartId="16.[151,250,1585,1607]" captionTargetBox="[314,1273,1156,1563]" captionTargetId="figure@16.[314,1273,1156,1563]" captionTargetPageId="16" captionText="FIGURE 13. Upper and lower teeth of female paratype of Centrophorus longipinnis n. sp. (CSIRO H 7990 - 02, 825 mm TL)." httpUri="https://zenodo.org/record/1042305/files/figure.png" pageId="17">Fig. 13b</figureCitation>
<figureCitation id="1382B4B62332FFDA1F19CB6F65E2FD84" box="[325,423,583,608]" captionStart="FIGURE 13" captionStartId="16.[151,250,1585,1607]" captionTargetBox="[314,1273,1156,1563]" captionTargetId="figure@16.[314,1273,1156,1563]" captionTargetPageId="16" captionText="FIGURE 13. Upper and lower teeth of female paratype of Centrophorus longipinnis n. sp. (CSIRO H 7990 - 02, 825 mm TL)." figureDoi="http://doi.org/10.5281/zenodo.1042305" httpUri="https://zenodo.org/record/1042305/files/figure.png" pageId="17">Fig. 13b</figureCitation>
).
</paragraph>
<paragraph id="8B06A8332332FFDA1E9BCB44652CFCDC" blockId="17.[151,1437,151,1977]" pageId="17">
@ -487,11 +487,11 @@ Dermal denticles on flank below first dorsal fin varying greatly in shape betwee
TL) small, upright, slender, unicuspid, closely spaced (slightly overlapping), and posteriorly curved; about
<quantity id="4C4105D62332FFDA1A93CB9C6163FD28" box="[1231,1318,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="17" unit="mm" value="0.4">0.4 mm</quantity>
long (
<figureCitation id="1382B4B62332FFDA1B33CB9C6486FD14" captionStart="FIGURE 14" captionStartId="18.[151,250,1956,1978]" captionTargetBox="[234,1368,204,1914]" captionTargetId="figure@18.[192,1394,193,1935]" captionTargetPageId="18" captionText="FIGURE 14. Lateral trunk denticles (from below first dorsal fin) of Centrophorus longipinnis n. sp.: (A) juvenile female paratype, NMMB-P 14051, 408 mm TL; (B) female paratype, CSIRO H 8104 - 01, 855 mm TL." httpUri="https://zenodo.org/record/1042307/files/figure.png" pageId="17">Fig. 14a</figureCitation>
<figureCitation id="1382B4B62332FFDA1B33CB9C6486FD14" captionStart="FIGURE 14" captionStartId="18.[151,250,1956,1978]" captionTargetBox="[234,1368,204,1914]" captionTargetId="figure@18.[192,1394,193,1935]" captionTargetPageId="18" captionText="FIGURE 14. Lateral trunk denticles (from below first dorsal fin) of Centrophorus longipinnis n. sp.: (A) juvenile female paratype, NMMB-P 14051, 408 mm TL; (B) female paratype, CSIRO H 8104 - 01, 855 mm TL." figureDoi="http://doi.org/10.5281/zenodo.1042307" httpUri="https://zenodo.org/record/1042307/files/figure.png" pageId="17">Fig. 14a</figureCitation>
). Denticles of adults (based on a
<quantity id="4C4105D62332FFDA1C33CBFF6691FD0B" box="[623,724,727,752]" metricMagnitude="-1" metricUnit="m" metricValue="8.55" pageId="17" unit="mm" value="855.0">855 mm</quantity>
TL specimen) block-like, sessile (not raised on pedicels), rhomboidal, close set but not overlapping; anterior edges of crowns shallowly scalloped, posterior edge bluntly pointed (
<figureCitation id="1382B4B62332FFDA1EA6CA376519FCDC" box="[250,348,799,824]" captionStart="FIGURE 14" captionStartId="18.[151,250,1956,1978]" captionTargetBox="[234,1368,204,1914]" captionTargetId="figure@18.[192,1394,193,1935]" captionTargetPageId="18" captionText="FIGURE 14. Lateral trunk denticles (from below first dorsal fin) of Centrophorus longipinnis n. sp.: (A) juvenile female paratype, NMMB-P 14051, 408 mm TL; (B) female paratype, CSIRO H 8104 - 01, 855 mm TL." httpUri="https://zenodo.org/record/1042307/files/figure.png" pageId="17">Fig. 14b</figureCitation>
<figureCitation id="1382B4B62332FFDA1EA6CA376519FCDC" box="[250,348,799,824]" captionStart="FIGURE 14" captionStartId="18.[151,250,1956,1978]" captionTargetBox="[234,1368,204,1914]" captionTargetId="figure@18.[192,1394,193,1935]" captionTargetPageId="18" captionText="FIGURE 14. Lateral trunk denticles (from below first dorsal fin) of Centrophorus longipinnis n. sp.: (A) juvenile female paratype, NMMB-P 14051, 408 mm TL; (B) female paratype, CSIRO H 8104 - 01, 855 mm TL." figureDoi="http://doi.org/10.5281/zenodo.1042307" httpUri="https://zenodo.org/record/1042307/files/figure.png" pageId="17">Fig. 14b</figureCitation>
).
</paragraph>
<paragraph id="8B06A8332332FFDA1E9BCA6C64F8FB0C" blockId="17.[151,1437,151,1977]" pageId="17">First dorsal fin extremely long, moderately high; length 4.18 (4.18; 3.375.03) times its height, 1.76 (1.77; 1.761.98) times second dorsal-fin length; soft-fin length 2.86 (2.55; 2.483.47) times its height; height 1.14 (1.13; 0.991.25) times second dorsal-fin height; anterior margin slightly convex; apex narrowly rounded; posterior margin moderately concave, slanting well posteroventrally from top to bottom; free rear tip moderately long, relatively thick basally; inner margin nearly straight, 2.88 (2.94; 2.603.23) in soft-fin length, 1.00 (0.87; 0.82 1.20) times its height; insertion level with mid pectoralpelvic space, extremely well forward of pelvic-fin origin, pelvic-fin midpoint to first dorsal-fin insertion 15.1 (13.1; 13.217.6)% TL; base of exposed fin spine level with pectoral-fin free rear tip; spine base moderately broad, exposed anteriorly just above junction of spine and soft portion of fin; exposed fin spine relatively short, robust, tapering distally, anterior margin slightly convex; exposed portion of spine sloping strongly posterodorsally from base (of exposed portion) to apex, shorter in length to exposed portion of second dorsal-fin spine, exposed first dorsal spine length 0.39 (0.30; 0.230.39) times height of fin.</paragraph>
@ -506,16 +506,16 @@ TL) in height of fin; base of exposed fin spine level with mid pelvicfin inner m
<paragraph id="8B06A8332332FFDA1E9BCCC76550F959" blockId="17.[151,1437,151,1977]" pageId="17">Pectoral fins large; anterior margin weakly convex to nearly straight, its length 12.1 (12.3; 12.013.0)% TL; base very short, 2.68 (2.77; 2.262.99) in anterior margin length; apex somewhat angular, not falcate; posterior margin nearly straight from apex angle of free rear tip then broadly concave; inner margin slightly concave; free rear tip elongate in adults (less so in juveniles), free rear tip 1.18 (1.04; 1.011.14) in inner margin, extending past level of exposed first dorsal-fin spine; origin situated at level of mid-fifth gill slit, partially obscured by gill membrane.</paragraph>
<paragraph id="8B06A8332332FFDA1E9BCFEF67AAF85D" blockId="17.[151,1437,151,1977]" pageId="17">
Pelvic fins large, length 11.6 (11.4; 11.012.4)% TL, 1.20 (1.14; 1.181.44) times second dorsal-fin soft length; anterior margin slightly convex; apex narrowly rounded; posterior margin slightly concave; free rear tip acutely pointed, inner margin very slightly convex. Claspers of adult males relatively short, slender; tapering to a fleshy, narrowly rounded tip (
<figureCitation id="1382B4B62332FFDA1FB9CE1C667CF8A9" box="[485,569,1844,1869]" captionStart="FIGURE 15" captionStartId="19.[151,250,1034,1056]" captionTargetBox="[333,1232,208,989]" captionTargetId="figure@19.[331,1255,193,1013]" captionTargetPageId="19" captionText="FIGURE 15. Clasper (right) of Centrophorus longipinnis (paratype CSIRO H 8104 02, adult male 679 mm TL). A, Glans not dilated; B, Glans spread. Abbreviations: AP, apopyle; CG, clasper groove; CS, clasper spine; HP, hypopyle; P 2, pelvic fin; RH, rhipidion." httpUri="https://zenodo.org/record/1042309/files/figure.png" pageId="17">Fig. 15</figureCitation>
<figureCitation id="1382B4B62332FFDA1FB9CE1C667CF8A9" box="[485,569,1844,1869]" captionStart="FIGURE 15" captionStartId="19.[151,250,1034,1056]" captionTargetBox="[333,1232,208,989]" captionTargetId="figure@19.[331,1255,193,1013]" captionTargetPageId="19" captionText="FIGURE 15. Clasper (right) of Centrophorus longipinnis (paratype CSIRO H 8104 02, adult male 679 mm TL). A, Glans not dilated; B, Glans spread. Abbreviations: AP, apopyle; CG, clasper groove; CS, clasper spine; HP, hypopyle; P 2, pelvic fin; RH, rhipidion." figureDoi="http://doi.org/10.5281/zenodo.1042309" httpUri="https://zenodo.org/record/1042309/files/figure.png" pageId="17">Fig. 15</figureCitation>
); outer length 2.9 (2.53.3)% TL, 3.16 (2.983.81) times its base length (n = 8); clasper glans about
<quantity id="4C4105D62332FFDA1FF3CE7065B1F894" box="[431,500,1879,1904]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="17" unit="in" value="0.3">0.3 in</quantity>
clasper inner length; apopyle and hypopyle connected by long clasper groove; rhipidion moderately large, laterally expanded, extended from hypopyle to anterior of clasper tip; lateral edge with a slender, straight spine; dermal denticles mostly absent from dorsal surface.
</paragraph>
<caption id="DFC6F8BB2331FFD91ECBCE8C601EF83D" httpUri="https://zenodo.org/record/1042307/files/figure.png" pageId="18" targetBox="[234,1368,204,1914]" targetPageId="18">
<caption id="DFC6F8BB2331FFD91ECBCE8C601EF83D" ID-DOI="http://doi.org/10.5281/zenodo.1042307" httpUri="https://zenodo.org/record/1042307/files/figure.png" pageId="18" targetBox="[234,1368,204,1914]" targetPageId="18">
<paragraph id="8B06A8332331FFD91ECBCE8C601EF83D" blockId="18.[151,1436,1956,2009]" pageId="18">
<emphasis id="B9CD74212331FFD91ECBCE8C6566F85E" bold="true" box="[151,291,1956,1978]" pageId="18">FIGURE 14.</emphasis>
Lateral trunk denticles (from below first dorsal fin) of
<taxonomicName id="4CB9D3B02331FFD91D24CE8C60C5F85E" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[888,1152,1956,1978]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="longipinnis" status="sp. nov.">
<taxonomicName id="4CB9D3B02331FFD91D24CE8C60C5F85E" authority="White &amp; Ebert &amp; Naylor, 2017" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[888,1152,1956,1978]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="18" phylum="Chordata" rank="species" species="longipinnis" status="sp. nov.">
<emphasis id="B9CD74212331FFD91D24CE8C60C5F85E" box="[888,1152,1956,1978]" italics="true" pageId="18">Centrophorus longipinnis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2FEC95A2331FFD91AD6CE8C60F8F85E" box="[1162,1213,1956,1978]" pageId="18" rank="species">
@ -526,11 +526,11 @@ Lateral trunk denticles (from below first dorsal fin) of
H 8104-01, 855 mm TL.
</paragraph>
</caption>
<caption id="DFC6F8BB2330FFD81ECBCD2264BEFBB9" httpUri="https://zenodo.org/record/1042309/files/figure.png" pageId="19" targetBox="[333,1232,208,989]" targetPageId="19">
<caption id="DFC6F8BB2330FFD81ECBCD2264BEFBB9" ID-DOI="http://doi.org/10.5281/zenodo.1042309" httpUri="https://zenodo.org/record/1042309/files/figure.png" pageId="19" targetBox="[333,1232,208,989]" targetPageId="19">
<paragraph id="8B06A8332330FFD81ECBCD2264BEFBB9" blockId="19.[151,1436,1034,1117]" pageId="19">
<emphasis id="B9CD74212330FFD81ECBCD22655AFBC4" bold="true" box="[151,287,1034,1056]" pageId="19">FIGURE 15.</emphasis>
Clasper (right) of
<taxonomicName id="4CB9D3B02330FFD81F80CD2266A5FBC4" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[476,736,1034,1056]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="longipinnis">
<taxonomicName id="4CB9D3B02330FFD81F80CD2266A5FBC4" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[476,736,1034,1056]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="19" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B9CD74212330FFD81F80CD2266A5FBC4" box="[476,736,1034,1056]" italics="true" pageId="19">Centrophorus longipinnis</emphasis>
</taxonomicName>
(paratype
@ -604,7 +604,7 @@ New Guinea
(Huon
<collectingRegion id="497D66D12330FFD81AB4CE566158F873" box="[1256,1309,1918,1943]" country="Papua New Guinea" name="Gulf" pageId="19">Gulf</collectingRegion>
) (
<figureCitation id="1382B4B62330FFD81B64CE5661CAF873" box="[1336,1423,1918,1943]" captionStart="FIGURE 10" captionStartId="13.[151,250,1838,1860]" captionTargetBox="[299,1271,1289,1807]" captionTargetId="figure@13.[299,1271,1289,1807]" captionTargetPageId="13" captionText="FIGURE 10. Map showing the collection locations, where known, of specimens of Centrophorus lesliei n. sp. (yellow and green) and Centrophorus longipinnis n. sp. (cyan and pink). Stars denote the holotypes, circles denote paratypes, and triangles denote non-type specimens (Image © NASA, TerraMetrics, Google Earth)." httpUri="https://zenodo.org/record/1042299/files/figure.png" pageId="19">Fig. 10</figureCitation>
<figureCitation id="1382B4B62330FFD81B64CE5661CAF873" box="[1336,1423,1918,1943]" captionStart="FIGURE 10" captionStartId="13.[151,250,1838,1860]" captionTargetBox="[299,1271,1289,1807]" captionTargetId="figure@13.[299,1271,1289,1807]" captionTargetPageId="13" captionText="FIGURE 10. Map showing the collection locations, where known, of specimens of Centrophorus lesliei n. sp. (yellow and green) and Centrophorus longipinnis n. sp. (cyan and pink). Stars denote the holotypes, circles denote paratypes, and triangles denote non-type specimens (Image © NASA, TerraMetrics, Google Earth)." figureDoi="http://doi.org/10.5281/zenodo.1042299" httpUri="https://zenodo.org/record/1042299/files/figure.png" pageId="19">Fig. 10</figureCitation>
).
<bibRefCitation id="EF28D5C22330FFD81ECBCE8B65F1F85F" box="[151,436,1954,1979]" pageId="19" refString="Compagno, L. J. V., Last, P. R., Stevens, J. D. &amp; Alava, M. N. R. (2005) Checklist of Philippine Chondrichthyes. CSIRO Marine Laboratories Report 243. CSIRO Marine Laboratories, Hobart, 103 pp." type="book">
Compagno
@ -638,7 +638,7 @@ Specific name a combination of the Latin
<paragraph id="8B06A8332337FFDF1ECBC861609EFE64" blockId="20.[151,1436,328,384]" pageId="20">
<emphasis id="B9CD74212337FFDF1ECBC8616554FE84" bold="true" box="[151,273,329,352]" pageId="20">TABLE 2.</emphasis>
Proportional dimensions as percentages of total length for the holotype of
<taxonomicName id="4CB9D3B02337FFDF1ADFC86061D9FE84" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[1155,1436,328,352]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="species" species="longipinnis">
<taxonomicName id="4CB9D3B02337FFDF1ADFC86061D9FE84" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[1155,1436,328,352]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="20" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B9CD74212337FFDF1ADFC86061D9FE84" box="[1155,1436,328,352]" italics="true" pageId="20">Centrophorus longipinnis</emphasis>
</taxonomicName>
(NMMB-P 15756), and ranges for small paratypes (&lt;524 mm TL) and large paratypes (&gt;700 mm TL).
@ -653,17 +653,17 @@ TL) (&gt;
TL)
<emphasis id="B9CD74212337FFDE1A01CE9C61D6F822" italics="true" lastPageId="21" pageId="20">......continued on the next page ......continued on the next page</emphasis>
The Maximum Likelihood tree generated from the ND2 sequences obtained provide further support for the separation of
<taxonomicName id="4CB9D3B02335FFDD1F6ECA9465D2FC30" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[306,407,956,980]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="lesliei">
<taxonomicName id="4CB9D3B02335FFDD1F6ECA9465D2FC30" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[306,407,956,980]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordara" rank="species" species="lesliei">
<emphasis id="B9CD74212335FFDD1F6ECA9465D2FC30" box="[306,407,956,980]" italics="true" pageId="22">C. lesliei</emphasis>
</taxonomicName>
and
<taxonomicName id="4CB9D3B02335FFDD1F8CCA946635FC30" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[464,624,956,980]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="longipinnis">
<taxonomicName id="4CB9D3B02335FFDD1F8CCA946635FC30" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[464,624,956,980]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B9CD74212335FFDD1F8CCA946635FC30" box="[464,624,956,980]" italics="true" pageId="22">C. longipinnis</emphasis>
</taxonomicName>
as valid species (
<figureCitation id="1382B4B62335FFDD1D67CA9467CAFC31" box="[827,911,956,981]" captionStart="FIGURE 16" captionStartId="23.[151,250,1604,1626]" captionTargetBox="[153,1340,303,1579]" captionTargetId="figure@23.[151,1436,301,1582]" captionTargetPageId="23" captionText="FIGURE 16. Maximum Likelihood tree estimated under the General Time Reversible model (GTR) with model terms to accommodate both Invariant site (I) and Gamma Distributed rates (G). Bootstrap support values are shown from a separate ML bootstrap analysis. Sequences used in this tree are part of the Chondrichthyan Tree of Life project (http: // sharksrays. org /)." httpUri="https://zenodo.org/record/1042311/files/figure.png" pageId="22">Fig. 16</figureCitation>
<figureCitation id="1382B4B62335FFDD1D67CA9467CAFC31" box="[827,911,956,981]" captionStart="FIGURE 16" captionStartId="23.[151,250,1604,1626]" captionTargetBox="[153,1340,303,1579]" captionTargetId="figure@23.[151,1436,301,1582]" captionTargetPageId="23" captionText="FIGURE 16. Maximum Likelihood tree estimated under the General Time Reversible model (GTR) with model terms to accommodate both Invariant site (I) and Gamma Distributed rates (G). Bootstrap support values are shown from a separate ML bootstrap analysis. Sequences used in this tree are part of the Chondrichthyan Tree of Life project (http: // sharksrays. org /)." figureDoi="http://doi.org/10.5281/zenodo.1042311" httpUri="https://zenodo.org/record/1042311/files/figure.png" pageId="22">Fig. 16</figureCitation>
).
<taxonomicName id="4CB9D3B02335FFDD1DF9CA9460CEFC30" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[933,1163,956,980]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="lesliei">
<taxonomicName id="4CB9D3B02335FFDD1DF9CA9460CEFC30" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[933,1163,956,980]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordara" rank="species" species="lesliei">
<emphasis id="B9CD74212335FFDD1DF9CA9460CEFC30" box="[933,1163,956,980]" italics="true" pageId="22">Centrophorus lesliei</emphasis>
</taxonomicName>
samples form a distinct group nested within, but separate from, the three long-snout species groups (i.e.
@ -685,19 +685,19 @@ Channel, close to the locality of the
and three
<typeStatus id="540216912335FFDD1DB9CD586015FB6C" box="[997,1104,1136,1160]" pageId="22" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4CB9D3B02335FFDD1ADFCD5860A8FB6C" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[1155,1261,1136,1160]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="lesliei">
<taxonomicName id="4CB9D3B02335FFDD1ADFCD5860A8FB6C" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[1155,1261,1136,1160]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordara" rank="species" species="lesliei">
<emphasis id="B9CD74212335FFDD1ADFCD5860A8FB6C" box="[1155,1261,1136,1160]" italics="true" pageId="22">C. lesliei</emphasis>
</taxonomicName>
.
<taxonomicName id="4CB9D3B02335FFDD1B5DCD586550FB48" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="longipinnis">
<taxonomicName id="4CB9D3B02335FFDD1B5DCD586550FB48" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B9CD74212335FFDD1B5DCD586550FB48" italics="true" pageId="22">Centrophorus longipinnis</emphasis>
</taxonomicName>
forms a distinct group well separated from the long-snout and
<taxonomicName id="4CB9D3B02335FFDD1A43CDBC60CFFB48" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[1055,1162,1172,1196]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="lesliei">
<taxonomicName id="4CB9D3B02335FFDD1A43CDBC60CFFB48" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[1055,1162,1172,1196]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordara" rank="species" species="lesliei">
<emphasis id="B9CD74212335FFDD1A43CDBC60CFFB48" box="[1055,1162,1172,1196]" italics="true" pageId="22">C. lesliei</emphasis>
</taxonomicName>
groups. Within the
<taxonomicName id="4CB9D3B02335FFDD1BDECDBC6550FB34" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="longipinnis">
<taxonomicName id="4CB9D3B02335FFDD1BDECDBC6550FB34" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B9CD74212335FFDD1BDECDBC6550FB34" italics="true" pageId="22">C. longipinnis</emphasis>
</taxonomicName>
group, two samples (GN 11175 from
@ -705,9 +705,9 @@ group, two samples (GN 11175 from
and GN 4348 from the
<collectingCountry id="F3AEE8A32335FFDD1AD8CD9F6141FB34" box="[1156,1284,1207,1232]" name="Philippines" pageId="22">Philippines</collectingCountry>
) show some differences to the main group (
<figureCitation id="1382B4B62335FFDD1FA9CDF4660CFB11" box="[501,585,1244,1269]" captionStart="FIGURE 16" captionStartId="23.[151,250,1604,1626]" captionTargetBox="[153,1340,303,1579]" captionTargetId="figure@23.[151,1436,301,1582]" captionTargetPageId="23" captionText="FIGURE 16. Maximum Likelihood tree estimated under the General Time Reversible model (GTR) with model terms to accommodate both Invariant site (I) and Gamma Distributed rates (G). Bootstrap support values are shown from a separate ML bootstrap analysis. Sequences used in this tree are part of the Chondrichthyan Tree of Life project (http: // sharksrays. org /)." httpUri="https://zenodo.org/record/1042311/files/figure.png" pageId="22">Fig. 16</figureCitation>
<figureCitation id="1382B4B62335FFDD1FA9CDF4660CFB11" box="[501,585,1244,1269]" captionStart="FIGURE 16" captionStartId="23.[151,250,1604,1626]" captionTargetBox="[153,1340,303,1579]" captionTargetId="figure@23.[151,1436,301,1582]" captionTargetPageId="23" captionText="FIGURE 16. Maximum Likelihood tree estimated under the General Time Reversible model (GTR) with model terms to accommodate both Invariant site (I) and Gamma Distributed rates (G). Bootstrap support values are shown from a separate ML bootstrap analysis. Sequences used in this tree are part of the Chondrichthyan Tree of Life project (http: // sharksrays. org /)." figureDoi="http://doi.org/10.5281/zenodo.1042311" httpUri="https://zenodo.org/record/1042311/files/figure.png" pageId="22">Fig. 16</figureCitation>
). But morphologically, the Indonesian specimen is identical to the other
<taxonomicName id="4CB9D3B02335FFDD1BDECDF46550FAFC" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="longipinnis">
<taxonomicName id="4CB9D3B02335FFDD1BDECDF46550FAFC" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B9CD74212335FFDD1BDECDF46550FAFC" italics="true" pageId="22">C. longipinnis</emphasis>
</taxonomicName>
specimens and is considered to be this species.
@ -998,7 +998,7 @@ concluded that the BMNH specimen was
<emphasis id="B9CD74212335FFDD1DDBCC0C67A2FAD8" box="[903,999,1316,1340]" italics="true" pageId="22">C. uyato</emphasis>
</taxonomicName>
based on DNA sequence comparisons for CO1. This result is inconsistent with the findings we present herein. Morphological data suggest that the BMNH specimen is
<taxonomicName id="4CB9D3B02335FFDD1FD8CC4465AEFA60" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[388,491,1388,1412]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="lesliei">
<taxonomicName id="4CB9D3B02335FFDD1FD8CC4465AEFA60" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[388,491,1388,1412]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordara" rank="species" species="lesliei">
<emphasis id="B9CD74212335FFDD1FD8CC4465AEFA60" box="[388,491,1388,1412]" italics="true" pageId="22">C. lesliei</emphasis>
</taxonomicName>
and furthermore, is very distinct from
@ -1008,11 +1008,11 @@ and furthermore, is very distinct from
in possessing a much longer first dorsal-fin base.
<bibRefCitation id="EF28D5C22335FFDD1F12CCA7668CFA4C" author="Munoz-Chapuli" box="[334,713,1423,1448]" pageId="22" refString="Munoz-Chapuli, R. &amp; Ramos, F. (1989) Review of the Centrophorus sharks (Elasmobranchii, Squalidae) of the eastern Atlantic. Cybium, 13 (1), 65 - 81." type="journal article" year="1989">Munoz-Chapuli &amp; Ramos (1989)</bibRefCitation>
recorded vastly different first dorsal-fin ray counts between
<taxonomicName id="4CB9D3B02335FFDD1BDECCB86555FA28" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="lusitanicus">
<taxonomicName id="4CB9D3B02335FFDD1BDECCB86555FA28" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordata" rank="species" species="lusitanicus">
<emphasis id="B9CD74212335FFDD1BDECCB86555FA28" italics="true" pageId="22">C. lusitanicus</emphasis>
</taxonomicName>
(=
<taxonomicName id="4CB9D3B02335FFDD1F6DCC9C65D0FA28" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[305,405,1460,1484]" class="Insecta" family="Brentidae" genus="Centrophorus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="lesliei">
<taxonomicName id="4CB9D3B02335FFDD1F6DCC9C65D0FA28" authorityName="White &amp; Ebert &amp; Naylor" authorityYear="2017" box="[305,405,1460,1484]" class="Elasmobranchii" family="Centrophoridae" genus="Centrophorus" higherTaxonomySource="CoL" kingdom="Animalia" order="Squaliformes" pageId="22" phylum="Chordara" rank="species" species="lesliei">
<emphasis id="B9CD74212335FFDD1F6DCC9C65D0FA28" box="[305,405,1460,1484]" italics="true" pageId="22">C. lesliei</emphasis>
</taxonomicName>
) and

View file

@ -1,57 +1,59 @@
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The Balkan Green Lizard was reported on Lefkada by Werner (1894: 230; 1929: 474) and Schmid (2019: 74). Also, citizen science records are scarce: Kariotes (
The Balkan Green Lizard was reported on Lefkada by
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<paragraph id="F44E3698FFCC9D647751FAB3FDED6D59" blockId="4.[113,756,1250,1488]" box="[113,603,1280,1307]" pageId="4" pageNumber="61">
<taxonomicName id="33F14D1BFFCC9D647751FAB3FDED6D59" ID-CoL="5CDM2" authority="(Merrem, 1820)" baseAuthorityName="Merrem" baseAuthorityYear="1820" box="[113,603,1280,1307]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="61" phylum="Chordata" rank="species" species="vermicularis">
<taxonomicName id="33F14D1BFFCC9D647751FAB3FDED6D59" ID-CoL="5CDM2" authority="(Merrem, 1820)" baseAuthorityName="Merrem" baseAuthorityYear="1820" box="[113,603,1280,1307]" class="Reptilia" family="Typhlopidae" genus="Xerotyphlops" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="61" rank="species" species="vermicularis">
<emphasis id="C685EA8AFFCC9D647751FAB3FE146D59" bold="true" box="[113,418,1280,1307]" italics="true" pageId="4" pageNumber="61">
<author id="2C4CDE7DFFCC9D647751FAB3FEBF6D59" box="[113,265,1280,1307]" fns="*" inm="true" npo="*" pageId="4" pageNumber="61">Xerotyphlops</author>
<title id="86F5DDAEFFCC9D647631FAB3FE146D59" box="[273,418,1280,1307]" pageId="4" pageNumber="61">vermicularis</title>
@ -68,14 +70,22 @@
Until recently the Worm Snake on Lefkada was known only for a single specimen collected in 1929 by
<author id="2C4CDE7DFFCC9D6475B0FA8EFD426D15" box="[656,756,1341,1367]" case="TC" fns="I" in="M." ins="D" ln="Beier" lnc="TC" npo="InLn" pageId="4" pageNumber="61">M. Beier</author>
(
<bibRefCitation id="90604B69FFCC9D647759FAE8FEE76D37" author="Werner F." box="[121,337,1371,1397]" pageId="4" pageNumber="61" pagination="471 - 765" refId="ref7143" refString="Werner F., 1929 - Teil V. Reptilia, Amphibia, Orthoptera, Embidaria und Scorpiones. In: Zoologische Forschungsreise nach den Jonischen Inseln und dem Peloponnes. Beier M. (ed.). Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien - mathematisch-naturwissenschaftliche Classe, 138 (9 - 10): 471 - 765." type="journal article" year="1929">
<title id="86F5DDAEFFCC9D647759FAE8FF796D37" box="[121,207,1371,1397]" jopScore="0.037037037" pageId="4" pageNumber="61" type="_" typeScore="12">Werner</title>
,
<year id="805977CBFFCC9D6477FBFAE8FEA16D37" box="[219,279,1371,1397]" pageId="4" pageNumber="61">1929</year>
: 478).
<journalOrPublisher id="2E303C54FFCC9D647646FAE8FD9E6D37" box="[358,552,1371,1397]" jopScore="0.00397147" pageId="4" pageNumber="61">Recently Schmid</journalOrPublisher>
(
: 478
</bibRefCitation>
).
<journalOrPublisher id="2E303C54FFCC9D647646FAE8FD9E6D37" box="[358,552,1371,1397]" jopScore="0.00397147" pageId="4" pageNumber="61">
Recently
<bibRefCitation id="90604B69FFCC9D6476F2FAE8FD116D34" author="Schmid A." box="[466,679,1371,1398]" pageId="4" pageNumber="61" pagination="72 - 79" refId="ref6898" refString="Schmid A., 2019 - Ein Hornchen zum Fruhstuck: Reptiliensuche auf Lefkada. Reptilia, Munster, 24 (136): 72 - 79." type="journal article" year="2019">
Schmid (
<part id="80559C05FFCC9D647519FAE8FDC06D37" box="[569,630,1371,1397]" pageId="4" pageNumber="61" type="volume">2019</part>
: 75) reported on a second individual after about a century, unfortunately not providing information on the locality and date of the sighting, and not providing a picture.
: 75)
</bibRefCitation>
</journalOrPublisher>
reported on a second individual after about a century, unfortunately not providing information on the locality and date of the sighting, and not providing a picture.
</paragraph>
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