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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.909.46838" ID-GBIF-Dataset="5cc495f4-4a41-41a0-b362-0e6d8c629b52" ID-PMC="PMC7015954" ID-Pensoft-Pub="1313-2970-909-79" ID-Pensoft-UUID="0AEF1785B8655964A36D859FBA7D8755" ID-PubMed="32089636" ID-ZooBank="22B7FF16D0A240F9B20EF7C6E0AF1842" ModsDocID="1313-2970-909-79" checkinTime="1580983798284" checkinUser="pensoft" docAuthor="Vujic, Ante, Likov, Laura, Radenkovic, Snezana, Tubic, Natasa Kocis, Djan, Mihajla, Sebic, Anja, Perez-Banon, Celeste, Barkalov, Anatolij, Hayat, Ruestem, Rojo, Santos, Andric, Andrijana &amp; Stahls, Gunilla" docDate="2020" docId="3EF7414EA1FA5C65961F5D765282F57D" docLanguage="en" docName="ZooKeys 909: 79-158" docOrigin="ZooKeys 909" docSource="http://dx.doi.org/10.3897/zookeys.909.46838" docTitle="Merodon trizonus nomen dubium" docType="treatment" docVersion="4" id="0AEF1785B8655964A36D859FBA7D8755" lastPageNumber="79" masterDocId="0AEF1785B8655964A36D859FBA7D8755" masterDocTitle="Revision of the Merodon serrulatus group (Diptera, Syrphidae)" masterLastPageNumber="158" masterPageNumber="79" pageNumber="79" updateTime="1668168397753" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the Merodon serrulatus group (Diptera, Syrphidae)</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vujic, Ante</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Likov, Laura</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Radenkovic, Snezana</mods:namePart>
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<mods:name type="personal">
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<mods:namePart>Tubic, Natasa Kocis</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Djan, Mihajla</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sebic, Anja</mods:namePart>
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<mods:namePart>Perez-Banon, Celeste</mods:namePart>
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<mods:name type="personal">
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<mods:namePart>Barkalov, Anatolij</mods:namePart>
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<mods:namePart>Hayat, Ruestem</mods:namePart>
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<mods:namePart>Andric, Andrijana</mods:namePart>
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<taxonomicName LSID="3EF7414E-A1FA-5C65-961F-5D765282F57D" authority="(Szilady, 1940) nomen dubium" authorityName="nomen dubium" baseAuthorityName="Szilady" baseAuthorityYear="1940" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon trizonus" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="trizonus">
Merodon trizonus (
<normalizedToken originalValue="Szilády">Szilady</normalizedToken>
, 1940) nomen dubium
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</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
The identity of
<taxonomicName authorityName="nomen dubium" baseAuthorityName="Szilady" baseAuthorityYear="1940" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon trizonus" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="trizonus">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon trizonus</emphasis>
</taxonomicName>
remains unclear. The species was described based on two male and two female syntypes labelled &quot;La Calle [el Kala], Algeria&quot; and &quot;Ain Draham, Tunisia&quot;, which were not examined. Originally, the syntypes were located in the Hungarian National Museum in Budapest, but the
<taxonomicName lsidName="" order="Diptera" pageId="0" pageNumber="79" rank="order">Diptera</taxonomicName>
collection was destroyed by a fire in 1956. The description of
<bibRefCitation author="Szilady, L" journalOrPublisher="Annales Musei Nationalis Hungarici" pageId="0" pageNumber="79" pagination="34 - 70" refId="B66" refString="Szilady, L, 1940. Ueber Palaearktische Syrphiden. Annales Musei Nationalis Hungarici 33: 34 - 70" title="Ueber Palaearktische Syrphiden." volume="33" year="1940">
<normalizedToken originalValue="Szilády">Szilady</normalizedToken>
(1940)
</bibRefCitation>
is incomplete and based on a few differences from the related species
<taxonomicName lsidName="M. hirsutus" pageId="0" pageNumber="79" rank="species" species="hirsutus">
<emphasis italics="true" pageId="0" pageNumber="79">M. hirsutus</emphasis>
</taxonomicName>
. The types of
<taxonomicName lsidName="M. trizonus" pageId="0" pageNumber="79" rank="species" species="trizonus">
<emphasis italics="true" pageId="0" pageNumber="79">M. trizonus</emphasis>
</taxonomicName>
are assumed lost and the description is insufficiently accurate to associate name to one of these species. Currently, two species from the
<taxonomicName lsidName="M. serrulatus" pageId="0" pageNumber="79" rank="species" species="serrulatus">
<emphasis italics="true" pageId="0" pageNumber="79">M. serrulatus</emphasis>
</taxonomicName>
species group, to which
<taxonomicName lsidName="M. hirsutus" pageId="0" pageNumber="79" rank="species" species="hirsutus">
<emphasis italics="true" pageId="0" pageNumber="79">M. hirsutus</emphasis>
</taxonomicName>
belongs, occur in northern Africa, namely
<taxonomicName lsidName="M. bequaerti" pageId="0" pageNumber="79" rank="species" species="bequaerti">
<emphasis italics="true" pageId="0" pageNumber="79">M. bequaerti</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="M. sophron." pageId="0" pageNumber="79" rank="species" species="sophron.">
<emphasis italics="true" pageId="0" pageNumber="79">M. sophron.</emphasis>
</taxonomicName>
Therefore, we propose to leave the name
<taxonomicName authorityName="nomen dubium" baseAuthorityName="Szilady" baseAuthorityYear="1940" class="Insecta" family="Syrphidae" genus="Merodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merodon trizonus" order="Diptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="trizonus">
<emphasis italics="true" pageId="0" pageNumber="79">Merodon trizonus</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Szilády">Szilady</normalizedToken>
, 1940) as
<emphasis italics="true" pageId="0" pageNumber="79">nomen dubium</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
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Thomson, 1878
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<paragraph id="05982531B0CA8AAB263BE43CA947D029" pageNumber="151">
<taxonomicName id="B26209C6A0CFAFCDCDF4E83C73D6267D" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="schoedli" status="sp.n.">Odontomachus schoedli</taxonomicName>
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(Figs. 4, 10, 16, 47)
</paragraph>
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<subSubSection id="D8FF5416951831FE0BB3516298B7D3A9" pageNumber="151" type="etymology">
<paragraph id="8897FCD80A3288153A072AA6227BCC32" pageNumber="151">
Etymology: This species is named for our late colleague Stefan
<normalizedToken id="C7AFE1022DD0BCABAF457D3B95793F78" originalValue="Schödl" pageNumber="151">Schoedl</normalizedToken>
, former curator for
<taxonomicName id="AC2A63FF969F921FC6D4D9ED193BFF59" class="Insecta" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
at the Natural History Museum Vienna, who collected a major part of the type series.
</paragraph>
</subSubSection>
<subSubSection id="DD4EF0951926F703D9569EFA067C8457" pageNumber="151" type="materials_examined">
<paragraph id="CAA19BF156D6A058624EB556F8CB2DD5" pageNumber="151">
Type material:
<materialsCitation id="0A41761834E7EB4CFE65F5A1A1A73D96" ID-GBIF-Occurrence="923905781" collectingDate="26. II. 1999" collectorName="S. Schoedl" location="Luzon" pageNumber="151" specimenCode="23" typeStatus="Holotype">
<typeStatus id="0551871DE8113E9224C1CC747D40CD6D" pageNumber="151">Holotype</typeStatus>
worker (UPLB):
<location id="6770A3609F682EA8E1C45B5CDA8FC10F" LSID="urn:lsid:plazi:treatment:3EF7C95A86C04E7EF9BE4ABE65763AC2:6770A3609F682EA8E1C45B5CDA8FC10F" name="Luzon" pageNumber="151">Luzon</location>
:
<location id="8D9902E5DD6245944B94BFF91A55A4BA" LSID="urn:lsid:plazi:treatment:3EF7C95A86C04E7EF9BE4ABE65763AC2:8D9902E5DD6245944B94BFF91A55A4BA" name="Mountain Province" pageNumber="151">Mountain Province</location>
: south of
<location id="EDC4DF465A6407752C85A46CF80D2CAF" LSID="urn:lsid:plazi:treatment:3EF7C95A86C04E7EF9BE4ABE65763AC2:EDC4DF465A6407752C85A46CF80D2CAF" name="Sagada" pageNumber="151">Sagada</location>
,
<location id="CBE6ED37F2581FD6050368D16913D823" LSID="urn:lsid:plazi:treatment:3EF7C95A86C04E7EF9BE4ABE65763AC2:CBE6ED37F2581FD6050368D16913D823" name="Bagnen" pageNumber="151">Bagnen</location>
, slopes of
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, 1600 m,
<collectingDate id="9E9FEA7CEB06BBAA869013AAD9331C86" pageNumber="151">26.II.1999</collectingDate>
, leg.
<collectorName id="03717718CDE20FC8E085477EB5E7B481" pageNumber="151">
S.
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(
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).
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Paratypes(24 workers (BMNH, CZW, NHMW, UPLB): same locality data as holotype, 13 ∑∑, leg. H. Zettel (189), 6 ∑∑. B e n g u e t: west of Baguio, at km 7 of Asin Road, 17.II. 1999, leg. S.
<normalizedToken id="2610C0522F7C9703CEEC69E60B6C331A" originalValue="Schödl" pageNumber="151">Schoedl</normalizedToken>
(11), 2 ∑∑, leg. H. Zettel (179), 2 ∑∑; Baguio, leg. C.F. Baker, 1 ∑.
</paragraph>
</subSubSection>
<subSubSection id="CBFC4730D491017C8AE53CC5AFCBA5EF" pageNumber="151" type="description">
<paragraph id="FC1B29A86A8B98D20323A93296E82972" pageNumber="151">Description of worker: Measurements: holotype worker: CI 72, HL 2.95, HW 2.12, MdI 58, MdL 1.70, MsL 4.17, PnW 1.25, PtH 1.18, PtL 1.12, PtW 0.49, SI 146, SL 3.08, TL 16.25; paratype worker with smallest HW: CI 72, HL 2.98, HW 2.13, MdI 55, MdL 1.65, MsL 4.00, PnW 1.20, PtH 1.18, PtL 1.18, PtW 0.51, SI 144, SL 3.07, TL 12.75; paratype worker with largest HW: CI 73, HL 3.60, HW 2.62, MdI 59, MdL 2.13, MsL 5.93, PnW 1.52, PtH 1.48, PtL 1.53, PtW 0.56, SI 137, SL 3.58, TL 16.13.</paragraph>
<paragraph id="88E92491A6491E131A3ABCFF945E5EA3" pageNumber="151">Structures: Head striate; striation reaching occipital margin. Pronotum with fine, transverse striation. Mesopleuron with fine transverse striation. Petiole stout, with short spine, bent backwards only very slightly.</paragraph>
<paragraph id="396D25F1DEA194621F737F1E85F20BC3" pageNumber="151">Pilosity: Pubescence sparse, long.</paragraph>
<paragraph id="A7C224B65FADB2F53D1CA400A2DBEAB6" pageNumber="151">Colour: Head and mesosoma medium brown, petiole and gaster dark brown.</paragraph>
</subSubSection>
<subSubSection id="586483EE9DFBECE175A850DAD73A4E14" pageNumber="151" type="distribution">
<paragraph id="F6217C62EF252D51C45EA8EB0BB0A7F4" pageNumber="151">Distribution (Fig. 47): Endemic to the Philippines: only in the north of Luzon: Mountain Province, Benguet.</paragraph>
</subSubSection>
<subSubSection id="7487FF64041021D4B0C031D2F4DF8B4C" pageNumber="151" type="biology_ecology">
<paragraph id="8180363B5F43F9FEE2E317F07A72601B" pageNumber="151">Habitats: This species was collected at higher elevations in strongly degraded forests.</paragraph>
</subSubSection>
<subSubSection id="E27BA9A5BA76B6A6CF9CE85CF2CFD75B" pageNumber="151" type="discussion">
<paragraph id="55429CB23FB2742EC8E416D8CA7478DD" pageNumber="151">
Notes:
<taxonomicName id="CC699EE740260E61C69D60E5FDB9817D" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="schoedli" status="sp.n.">Odontomachus schoedli</taxonomicName>
<taxonomicNameLabel id="43A68AA152C36AE9841B9A472459AC25" pageNumber="151" rank="species" rankGroup="species" rankGroupNumber="3" rankNumber="3">sp.n.</taxonomicNameLabel>
is only known from the western mountain ranges of northern Luzon. In Benguet it is sympatric with
<taxonomicName id="7D8AAB11883415FE4749431B86200B69" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="infandus">O. infandus</taxonomicName>
. Main differences between these two species are the striation of the pronotum, which is transverse in
<taxonomicName id="6E5D970BFBB0EA9DC7ABE1DEBC54D820" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="schoedli" status="sp.n.">O. schoedli</taxonomicName>
<taxonomicNameLabel id="E6F6C37E6F5FE6E04657AD9434AD2045" pageNumber="151" rank="species" rankGroup="species" rankGroupNumber="3" rankNumber="3">sp.n.</taxonomicNameLabel>
but more or less longitudinal in
<taxonomicName id="5FFF9948E00765564C78CF537E74188E" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="infandus">O. infandus</taxonomicName>
, and the shape of the petiole, which is remarkably short and with a very straight anterior face of the node (including spine) in
<taxonomicName id="5A56BCFDD779B43DCD51555BC0A45674" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="schoedli" status="sp.n.">O. schoedli</taxonomicName>
<taxonomicNameLabel id="2AB5A89FD3CDB8B02C3254EC0F03DD31" pageNumber="151" rank="species" rankGroup="species" rankGroupNumber="3" rankNumber="3">sp.n.</taxonomicNameLabel>
, whereas it has a longer, curved spine in
<taxonomicName id="B156B6EDC230AE5D3242442CB9441AC2" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="infandus">O. infandus</taxonomicName>
(only weakly developed in the smallest specimens). The uniformly light chocolate brown colour of head and mesosoma of
<taxonomicName id="D8538B559EDD04B5CA31CCDEE9B4F435" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="schoedli" status="sp.n.">O. schoedli</taxonomicName>
<taxonomicNameLabel id="882869384E6202DEE1F858D3F3ECBD86" pageNumber="151" rank="species" rankGroup="species" rankGroupNumber="3" rankNumber="3">sp.n.</taxonomicNameLabel>
is striking, but we have seen a few specimens of
<taxonomicName id="465370D6645E7830E25468F4B018D2F0" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="infandus">O. infandus</taxonomicName>
from northern Luzon with the same colouration.
</paragraph>
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<treatment id="3EF7F004CAB582726DB747282E2F3E67" ID-DOI="http://doi.org/10.5281/zenodo.6281653" ID-GBIF-Taxon="125136876" ID-Zenodo-Dep="6281653" LSID="urn:lsid:plazi:treatment:3EF7F004CAB582726DB747282E2F3E67" httpUri="http://treatment.plazi.org/id/3EF7F004CAB582726DB747282E2F3E67" lastPageNumber="493" pageNumber="493">
<subSubSection id="9CF0993DA500A8F40A9EEB29B7D9F418" type="nomenclature">
<paragraph id="780D2DD2B03CB0D745599AC968210296" pageNumber="493">
<taxonomicName id="8CB029F7F471F9650579E9CDE165D971" ID-CoL="33FS7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28550" class="Insecta" family="Formicidae" genus="Cyphomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Cyphomyrmex vorticis Weber" order="Hymenoptera" pageNumber="493" phylum="Arthropoda" rank="species" species="vorticis">Cyphomyrmex vorticis Weber</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="24C82AD3ACD4F2EE492FC1B11750465A" type="reference_group">
<paragraph id="F48C36D823DAC6021BBDBB911E9F2898" pageNumber="493">
<taxonomicName id="AF699C29EB8E6B29DE4E9CC683866CDD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28550" class="Insecta" family="Formicidae" genus="Cyphomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Cyphomyrmex vorticis Weber" order="Hymenoptera" pageNumber="493" phylum="Arthropoda" rank="species" species="vorticis">Cyphomyrmex vorticis Weber</taxonomicName>
, 1940:409-10; [[worker]]. Kempf, 1966:188-90.
</paragraph>
</subSubSection>
<subSubSection id="876766C7AFD2072B879F36466F771E68" type="distribution">
<paragraph id="F0985B44B40BD5CFBF26909732B22476" pageNumber="493">Kempf (1966) reported this species only from Bolivia (Santa Helena, the type locality) and Brazil. We have seen a series of 9 workers, 2 females, and 1 male from Zent, costa rica, collected in March, 1924 by W. M. Mann (USNM). The workers agree well with Kempfs redescription, and with cotypes in the LACM, in all particulars except the pilosity. The appressed pilosity of the head and body is fine and hair-like in the cotypes and is very inconspicuous. The Zent specimens differ in having more conspicuous, though no more abundant, pilosity because the individual hairs are broad and scale-like.</paragraph>
</subSubSection>
<subSubSection id="B616E070B0167F3D92572212CF46F916" type="discussion">
<paragraph id="BF017FF13C0909C4EED36A5B57936B82" pageNumber="493">
Whether or not these represent a variant of
<taxonomicName id="186E0548098BB30FFCD39CB375109EF8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28550" class="Insecta" family="Formicidae" genus="Cyphomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Cyphomyrmex vorticis Weber" order="Hymenoptera" pageNumber="493" phylum="Arthropoda" rank="species" species="vorticis">C. vorticis</taxonomicName>
, or an undescribed species, is uncertain. More material of both forms, as well as samples from intervening localities, must be available before the status of the Costa Rican specimens is clear. For the present they are tentatively assigned to
<taxonomicName id="34C8ABBC4502979AEA6469ECA34DA9B0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28550" class="Insecta" family="Formicidae" genus="Cyphomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Cyphomyrmex vorticis Weber" order="Hymenoptera" pageNumber="493" phylum="Arthropoda" rank="species" species="vorticis">C. vorticis</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>