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<mods:title id="5EC84B8966C6D83F547854074D7A7256">A nomenclator of extant and fossil taxa of the Melanopsidae (Gastropoda, Cerithioidea)</mods:title>
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<mods:namePart id="F33720BC1317FB3DEB389A343051B93A">Neubauer, Thomas A.</mods:namePart>
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Melanopsis thomasi
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, 1877
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<paragraph id="9B47BDA13ECD303C23F4B5946924429C" pageId="277" pageNumber="278">Original source.</paragraph>
<paragraph id="80144A4CEC24757D9000BA04C528B166" pageId="277" pageNumber="278">
<bibRefCitation id="191E3C116A915AC25FF2C185CAFA76CF" author="Tournouer, R" journalOrPublisher="Journal de Conchyliologie" pageId="332" pageNumber="333" pagination="275 - 276" publicationUrl="http://www.biodiversitylibrary.org/item/54878page/289/mode/1up" refId="B518" refString="Tournouer, R, 1877. Description d'une nouvelle espece fossile de Melanopside, provenant des terrains tertiaires superieurs de la province de Constantine. Journal de Conchyliologie 25: 275 - 276, http://www.biodiversitylibrary.org/item/54878page/289/mode/1up" title="Description d'une nouvelle espece fossile de Melanopside, provenant des terrains tertiaires superieurs de la province de Constantine." url="http://www.biodiversitylibrary.org/item/54878page/289/mode/1up" volume="25" year="1877">
<normalizedToken id="06D996E72F41D59CC4AE1ACC5FC6FE66" originalValue="Tournouër">Tournouer</normalizedToken>
1877
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: 275.
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<paragraph id="7F33E5387CD711BE575EC015A66DD215" pageId="277" pageNumber="278">Type horizon.</paragraph>
<paragraph id="32420675683B0B450E095AF9CA6323A5" pageId="277" pageNumber="278">Late Pannonian, late Miocene.</paragraph>
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<paragraph id="C1A7FDBA87302C410DE2775F20D023F1" pageId="277" pageNumber="278">Type locality.</paragraph>
<paragraph id="0526FAC321558D52B2FD57593794BB4A" pageId="277" pageNumber="278">
<normalizedToken id="6C0ED5B865931B540BA5622B1CA80902" originalValue="“Smendou”">&quot;Smendou&quot;</normalizedToken>
[Zighoud Youcef], Algeria.
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<paragraph id="1061A4B1EE101BA78490C70120ABD968" pageId="3" pageNumber="16">Taxon classification Animalia Lepidoptera Tortricidae</paragraph>
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<taxonomicName id="F56247F9B39A4B482E042E6FE5CEA3F2" ID-CoL="B4VDW" LSID="http://zoobank.org/1919B149-1E92-49DA-90BB-4E1689F9C42D" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha sakartvelana" order="Lepidoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="sakartvelana">Dichrorampha sakartvelana</taxonomicName>
<taxonomicNameLabel id="8B3A3E66FB3A5449A93273345A020AE4" pageId="3" pageNumber="16">sp. n.</taxonomicNameLabel>
Figs 5-6, 11-13, 15
</paragraph>
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<paragraph id="2D0C32A6B9C2C04B3019525D01F5E14C" pageId="4" pageNumber="17">
<pageBreakToken id="44BE089DC12D5D608AED36BF3E2E1A61" pageId="4" pageNumber="17" start="start">Material</pageBreakToken>
.
</paragraph>
<paragraph id="DFAF777C9564F7CC1FD7EC81D7D6719D" pageId="4" pageNumber="17">
Holotype ♀, with three labels: &quot;Georgia, Great Caucasus | Mountains, Caucasus Range, | near Abano pass | 2280 m
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N 42°15
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E 45°30
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| 27.vii.2014, netting | leg. B. Zlatkov&quot;, &quot;Holotypus ♀ |
<taxonomicName id="2824CD27F8EC683A80CF917AF48513E2" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="genus">Dichrorampha</taxonomicName>
|
<taxonomicName id="3CD765032BEE343437D4E8E392916C2D" lsidName="sakartvelana" pageId="4" pageNumber="17" rank="species" species="sakartvelana">sakartvelana</taxonomicName>
| det. B. Zlatkov 2015 [red label]&quot;, &quot; BFUS | ♀ | Genitalia slide | No. 27.vii.2014/1&quot;, BFUS. - Paratype ♂, with three labels: the first as in Holotype; the other two: &quot;Paratypus ♂ |
<taxonomicName id="5799450E8A5805B8DD657CE6DF199801" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha" order="Lepidoptera" pageId="4" pageNumber="17" phylum="Arthropoda" rank="genus">Dichrorampha</taxonomicName>
|
<taxonomicName id="8C9B387291133D4EB9E672043FA058BB" lsidName="sakartvelana" pageId="4" pageNumber="17" rank="species" species="sakartvelana">sakartvelana</taxonomicName>
| det. B. Zlatkov 2015 [red label]&quot;, &quot;NMNHS | ♂ | Genitalia slide | No. 27.vii.2014/2&quot;, NMNHS.
</paragraph>
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<subSubSection id="A663CD0F3F98FE2D66DCEBE18CEBBAFC" lastPageId="6" lastPageNumber="19" pageId="4" pageNumber="17" type="description">
<paragraph id="C94C7B0D4292D2C0BBA15FA2312B336D" pageId="4" pageNumber="17">Description.</paragraph>
<paragraph id="86CEDE0A4E8E976B95249F0BC2D8BB16" lastPageId="5" lastPageNumber="18" pageId="4" pageNumber="17">
Adult (Figs 5-6). Sexual dimorphism subtle. Head: Frons and vertex covered with brown-grey scales. Palpus labialis with whitish basal and brown-grey distal segment; the second segment with whitish base and brown tuft at the distal end. Antennae covered with dark grey scales. Thorax: Nota, patagia and tegulae uniformly grey, thorax underside (including coxae) whitish, legs brown. Forewings comparatively wide, in male with costal fold with 1/5 of the length of the costal edge. Forewing length male 8.65, female 8.90 mm, wingspan in set specimens
<pageBreakToken id="50A4CE7CA595DE8F95DF25C1F5173C90" pageId="5" pageNumber="18" start="start">18.5</pageBreakToken>
-19 mm. The specimens are worn, but preserved areas on the forewing upperside have pale greenish-grey overlaying scales which apparently do not form an obvious pattern. No markings are visible with the exception of five pairs of faint distal costal strigulae (pairs 5-9 sensu Baixeras, 2002). Three black terminal dots are present. Underside uniformly brown. Cilia creamy with pale brown margin. Hindwings with pale brown upperside and whitish underside. Cilia whitish. Abdomen covered with pale grey scales.
</paragraph>
<paragraph id="8D042B4FBB60CA24782E0FC5A53C6672" pageId="5" pageNumber="18">Male genitalia (Figs 11-12): Tegumen bearing a small lobe as uncus. Valva broad basally with wide basal cavity. Costal edge slightly convex. Sacculus nearly parallel to costal edge, indistinctly concave, ending with nearly straight angle. Ventral incision elongated, trapezoidal. Neck of valva slender, more than two times narrower than the basal part of valva and relatively long, 3/4 of its length. Cucullus with large dorsal lobe, densely covered with long setae, and a small rectangular ventral prominence. Phallus slender, ca. 3/5 of the length of valva, bent ventrally in the basal part, with membranous area extending at first on right, then on dorsal side. A large triangular prominence pointed dorsally at the right side of the tip is present. Circa 20 sockets of deciduous cornuti are counted.</paragraph>
<caption id="2B14586189549D036DAFF5E37E604032" pageId="5" pageNumber="18">
<paragraph id="80747E2128DB6CA25637FDD3D9B9F9FB" pageId="5" pageNumber="18">
Figures 11-13. Genitalia of
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sp. n. 11. Male genitalia, paratype; 12. Phallus, dorsal view, paratype; 13. Female genitalia, holotype. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="7AFA3E58E278220ADE9E3C2E06AA5C51" pageId="6" pageNumber="19">
<pageBreakToken id="6182775A53099AD640BCFC648A880BF6" pageId="6" pageNumber="19" start="start">Female</pageBreakToken>
genitalia (Fig. 13): Papillae anales wide. Apophyses posteriores equal in length to the apophyses anteriores, the latter look naturally deformed. Sterigma sclerotised, slightly asymmetrical, trapezoidal, with two incisions on the posterior margin of the postostial part. Ostium wide. Subgenital plate trapezoidal with distinctly sclerotised lateroposterior margins and rounded posterior angles. Antrum with the length of the membranous part of ductus bursae, well sclerotised, remarkably wide, nearly symmetrical, wineglass shaped and enveloped in a thin cuticular membrane visible after staining. The proximal sclerotisation of ductus bursae is barely discernible only under higher magnification. Ductus seminalis emerging at the middle of the membranous ductus bursae. Corpus bursae ovoid. A single moderately sized signum is present.
</paragraph>
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<subSubSection id="A31B6BC5BA7D1DF68B85759A02167A3D" pageId="6" pageNumber="19" type="diagnosis">
<paragraph id="025F79571177B3ABCB851C89539561A9" pageId="6" pageNumber="19">Diagnosis.</paragraph>
<paragraph id="737E42E23F242CFF5BEF1128BC3D466B" pageId="6" pageNumber="19">
The new species is characterized externally by uniformly coloured forewings (though the specimens are not fresh) and large size. The male genitalia resemble some species of the &quot;section petiverellae&quot; (sensu
<bibRefCitation id="B4828BC9E589DA90FAC1F11FAD1459CA" author="Danilevsky, A" editor="Byhovsky, BE" journalOrPublisher="Nauka, Leningrad" pageId="7" pageNumber="20" title="Listoviortki-Tortricidae triba plodozhorki Laspeyresiini [Tortrix Moths Tortricidae: Tribe Laspeyresiini]." volumeTitle="Fauna SSSR. Nasekomyie cheshuyekrylyie" year="1968">Danilevsky and Kuznetzov 1968</bibRefCitation>
), which apparently are closely related to
<taxonomicName id="B09A26B01620D5D776836FF0C6914A88" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha sakartvelana" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="sakartvelana">Dichrorampha sakartvelana</taxonomicName>
sp. n.
<taxonomicName id="CC81646329F65880414A3F4B6B8DF0B0" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha flavidorsana" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="flavidorsana">Dichrorampha flavidorsana</taxonomicName>
Knaggs, 1867 and
<taxonomicName id="D8D16A02CA6929CCF387C0B5FFDE05C9" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha filipjevi" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="filipjevi">Dichrorampha filipjevi</taxonomicName>
(Danilevsky, 1948) have similarly shaped valva, but the apical prominence of the phallus is pointed ventrally, and the ventral process of the cucullus is much larger in
<taxonomicName id="01C77090D1895077B919BC2E4772C96A" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha filipjevi" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="filipjevi">Dichrorampha filipjevi</taxonomicName>
.
<taxonomicName id="74AD1782A3E4A06FD4C07BE369B0EFE7" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha petiverella" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="petiverella">Dichrorampha petiverella</taxonomicName>
(Linnaeus, 1758) and
<taxonomicName id="905A4DC652A1A0DDD6362F0B15081105" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha proxima" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="proxima">Dichrorampha proxima</taxonomicName>
(Danilevsky, 1948) are also similar in general, but they have two distinct processes of the cucullus. Female genitalia do not demonstrate clear affinities to any Palaearctic
<taxonomicName id="1DA805E9D0C95A6B62C4E65768463574" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="genus">Dichrorampha</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="3FEA0A436BD7D2EC656A80D83AC36C88" pageId="6" pageNumber="19" type="preimaginal stages">
<paragraph id="FED93C4E9ACB04F2DF4890F3E0885A20" pageId="6" pageNumber="19">Preimaginal stages.</paragraph>
<paragraph id="DCD5E951EE3A8B64B0DF18768A1B1DDA" pageId="6" pageNumber="19">Unknown.</paragraph>
</subSubSection>
<subSubSection id="DD8E3BB8D71A4FF46F28E818741FEED7" pageId="6" pageNumber="19" type="biology">
<paragraph id="04F816746F2BCB6D8AAFA123CAF1005E" pageId="6" pageNumber="19">Biology.</paragraph>
<paragraph id="70902CBFCEC9754CFDC6052CBA046F0E" pageId="6" pageNumber="19">
The moths were collected at the end of July, but their condition presumes that they are on the wing earlier. The larval host plant is most probably
<taxonomicName id="8DB9CF14BD1487C52646ED874C0E1CFF" class="Magnoliopsida" family="Asteraceae" genus="Achillea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achillea" order="Asterales" pageId="6" pageNumber="19" phylum="Tracheophyta" rank="genus">Achillea</taxonomicName>
sp. above which the moths were flying (plant material was not preserved and an exact identification is thus not possible). The habitat (Fig. 15) is a subalpine meadow at an elevation of ca. 2300 m with denuded rocks where the host plant grows.
</paragraph>
<caption id="DB56B6308D30EC76F42971C6FF34B409" pageId="6" pageNumber="19">
<paragraph id="3A3DAFA60D03277E14E079D2E36AAF68" pageId="6" pageNumber="19">
Figures 14-15. Habitats. 14. Habitat of
<taxonomicName id="2FDAFCDF802044041636334DB6FC1F90" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha pentheriana" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="pentheriana">Dichrorampha pentheriana</taxonomicName>
(Rebel, 1917), Vitosha Mts, 13.vii.2012, foreground: the suspected larval host plant
<taxonomicName id="A02AE66B1F553423B724E8A74218821F" class="Magnoliopsida" family="Asteraceae" genus="Achillea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achillea lingulata" order="Asterales" pageId="6" pageNumber="19" phylum="Tracheophyta" rank="species" species="lingulata">Achillea lingulata</taxonomicName>
(photo: D. Gradinarov); 15. Habitat of
<taxonomicName id="F0E40C66E7F7D07BC0B3388E3F9DA38D" class="Insecta" family="Tortricidae" genus="Dichrorampha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichrorampha sakartvelana" order="Lepidoptera" pageId="6" pageNumber="19" phylum="Arthropoda" rank="species" species="sakartvelana">Dichrorampha sakartvelana</taxonomicName>
sp. n., Great Caucasus Mts, Tusheti Range, 27.vii.2014 (photo: M. Ilieva).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A7BA8EF7099916CE3B26DCD3F1D09870" pageId="6" pageNumber="19" type="distribution">
<paragraph id="E52628459B9FCA3F5BE5F8E94471053D" pageId="6" pageNumber="19">Distribution.</paragraph>
<paragraph id="9CFF7631A9A7EBAC2A8B281E67371EA2" pageId="6" pageNumber="19">Known from the type locality only.</paragraph>
</subSubSection>
<subSubSection id="1E5289C3FB5DECBBA934B50E2AE5F00E" pageId="6" pageNumber="19" type="etymology">
<paragraph id="99F816AF9AF76CE2F3F5681B767445B7" pageId="6" pageNumber="19">Etymology.</paragraph>
<paragraph id="735C24F9DE45849A1A0F5088B530CCEC" pageId="6" pageNumber="19">
The name of the species is an adjective, derived from the autonym for Georgia, Sakartvelo, and the specific ending for
<taxonomicName id="854FA6C211148404D9A944443CA02256" family="Tortricidae" lsidName="" pageId="6" pageNumber="19" rank="family">Tortricidae</taxonomicName>
-ana.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.139.1308" ID-GBIF-Dataset="d6a55a29-30df-409c-9879-ecff3c19fbb2" ID-PMC="PMC3260909" ID-Pensoft-Pub="1313-2970-139-45" ID-PubMed="22259309" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-139-45" ModsDocOrigin="ZooKeys 139" ModsDocTitle="A review of Biston Leach, 1815 (Lepidoptera, Geometridae, Ennominae) from China, with description of one new species" checkinTime="1451249759556" checkinUser="pensoft" docAuthor="Jiang, Nan, Xue, Dayong &amp; Han, Hongxiang" docDate="2011" docId="3EF3A28C39BF725DF6B311EAF964A27A" docLanguage="en" docName="ZooKeys 139: 45-96" docOrigin="ZooKeys 139" docSource="http://dx.doi.org/10.3897/zookeys.139.1308" docTitle="Biston thibetaria Oberthuer 1886" docType="treatment" docVersion="3" lastPageNumber="71" masterDocId="FFFD4129FFDEFF9C9A7DFF89F61DFFD3" masterDocTitle="A review of Biston Leach, 1815 (Lepidoptera, Geometridae, Ennominae) from China, with description of one new species" masterLastPageNumber="96" masterPageNumber="45" pageNumber="70" updateTime="1668152443332" updateUser="ExternalLinkService">
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<mods:title>A review of Biston Leach, 1815 (Lepidoptera, Geometridae, Ennominae) from China, with description of one new species</mods:title>
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<mods:namePart>Jiang, Nan</mods:namePart>
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<treatment ID-GBIF-Taxon="152031639" LSID="urn:lsid:plazi:treatment:3EF3A28C39BF725DF6B311EAF964A27A" httpUri="http://treatment.plazi.org/id/3EF3A28C39BF725DF6B311EAF964A27A" lastPageId="26" lastPageNumber="71" pageId="25" pageNumber="70">
<subSubSection pageId="25" pageNumber="70" type="nomenclature">
<paragraph pageId="25" pageNumber="70">
<taxonomicName LSID="http://species-id.net/wiki/Biston_thibetaria" authority="Oberthuer, 1886" authorityName="Oberthuer" authorityYear="1886" class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston thibetaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="thibetaria">
Biston thibetaria (
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
, 1886)
</taxonomicName>
Figs 55-5789114128
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="70" type="reference_group">
<paragraph pageId="25" pageNumber="70">
<taxonomicName class="Insecta" family="Geometridae" genus="Amphidasys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidasys thibetaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="thibetaria">Amphidasys thibetaria</taxonomicName>
<bibRefCitation author="Oberthuer, C" journalOrPublisher="Etudes d'Entomologie" pageId="30" pageNumber="75" pagination="13 - 38" title="Nouveaux Lepidopteres du Thibet." volume="11" year="1886">
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
, 1886
</bibRefCitation>
,
<normalizedToken originalValue="Études">Etudes</normalizedToken>
ent., 11: 32, pl. 5, fig. 30. Holotype ♀, China: Sichuan (?),
<normalizedToken originalValue="Châpa">Chapa</normalizedToken>
. (ZFMK)
</paragraph>
<paragraph pageId="25" pageNumber="70">
<taxonomicName class="Insecta" family="Geometridae" genus="Buzura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buzura thibetaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="thibetaria">Buzura thibetaria</taxonomicName>
:
<bibRefCitation pageId="25" pageNumber="70">Prout, 1915</bibRefCitation>
, in Seitz, Macrolepid. World, 4: 360, pl. 19: h.
</paragraph>
<paragraph pageId="25" pageNumber="70">
<taxonomicName class="Insecta" family="Geometridae" genus="Buzura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buzura (Blepharoctenia) thibetaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="thibetaria" subGenus="Blepharoctenia">Buzura (Blepharoctenia) thibetaria</taxonomicName>
:
<bibRefCitation pageId="25" pageNumber="70">Wehrli, 1941</bibRefCitation>
, in Seitz, Gross-Schmett. Erde, 4 (Suppl.): 436.
</paragraph>
<paragraph pageId="25" pageNumber="70">
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston thibetaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="thibetaria">Biston thibetaria</taxonomicName>
:
<bibRefCitation pageId="25" pageNumber="70">Parsons et al., 1999</bibRefCitation>
, Geometrid Moths of the World, a Catalogue, 1: 88.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="70" type="diagnosis">
<paragraph pageId="25" pageNumber="70">Diagnosis.</paragraph>
<paragraph pageId="25" pageNumber="70">
This species is very distinct and is easily recognizable by the thick black lines and yellowish green bands placed basally of the antemedial line of the forewing and distally of the postmedial lines of both wings, the large, black ringed and pale-centred discal spots on both wings, as well as the black-belted abdomen and the fresh yellow anal tuft. The male genitalia of
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston thibetaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="thibetaria">Biston thibetaria</taxonomicName>
are close to those of
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston panterinaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="panterinaria">Biston panterinaria</taxonomicName>
: the apex of the uncus is bifurcated and about four-fifths as long as the basal width; the median process of the gnathos is short and round apically; the valva is broad basally and narrow apically; the ventral margin of the valva is slightly sinuous; the juxta has a deep incision at the middle on the posterior margin; the cornutus is stick-like; a narrow sclerotized band is present on lateral side of the aedeagus. But it can be distinguished from that species by the strongly rounded basal half of the valva. The female genitalia of the species are close to those of
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston panterinaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="panterinaria">Biston panterinaria</taxonomicName>
as follows: the ostium bursae is weakly sclerotized; the ductus bursae is very short; the corpus bursae is curved medially, striated in the posterior half and enlarged at tip; the signum is oval and with marginal spines. It differs in having an oval lamella postvaginalis, which is absent in
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston panterinaria" order="Lepidoptera" pageId="25" pageNumber="70" phylum="Arthropoda" rank="species" species="panterinaria">Biston panterinaria</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="26" lastPageNumber="71" pageId="25" pageNumber="70" type="material examined">
<paragraph pageId="25" pageNumber="70">Material examineds.</paragraph>
<paragraph lastPageId="26" lastPageNumber="71" pageId="25" pageNumber="70">
CHINA, Sichuan (ZFMK): Sichuan (?),
<normalizedToken originalValue="Châpa">Chapa</normalizedToken>
, 1♂(Syntype). Hubei (IZCAS): Shennongjia, 600-700 m, 17-18.VII., 2.VIII.1998, coll. Ye Chanjuan, 3♂; Xingshan, Longmenhe, 730-1350 m,
<normalizedToken originalValue="VIVII">VI-VII</normalizedToken>
.1993, coll. Song Shimei et al., 11♂1♀; Zigui, Jiulingtou, 220-250 m, 25.VII.1993, coll. Song Shimei, 2♂;
<pageBreakToken pageId="26" pageNumber="71" start="start">Badong</pageBreakToken>
, 19.V.1989, 1♂; Hefeng, 650 m, 29.V.1989, coll. Li Wei, 1♂; Lichuan, Xingdoushan, 860 m, 6.VII.1989, coll. Li Wei, 1♂; Xianfeng, 800 m, 2.VI.1989, coll. Li Wei, 1♀. Hunan (IZCAS): Tianpingshan, 12.VIII.1981, 1♂. Fujian (IZCAS): Mt. Wuyi, Sangang, 7.VII.1982, coll. Wang Linyao, 1♂. Guangxi (IZCAS): Longsheng, 10-13.VI.1980, coll. Song Shimei and Wang Linyao, 4♂. Sichuan (IZCAS): Barkam, 2600 m, 21.VIII.1983, coll. Chai Huaicheng, 1♂; Luding, Moxi, 600-1900 m, 11-17.VI.1983, coll. Chai Huaicheng and Wang Shuyong, 3♂; Luding, Guzanjiangju, 1635 m, 21.V.2009, coll. Liang Hongbin and Wang Zhiliang, 1♂2♀; Batang, 1975, 9♂; Dukou, Pingdi, 5-22.VI.1987, coll. Zhang Baolin, 13♂; Huili, 23-29.VII.1974, coll. Han Yinheng, 2♂1♀; Yanyuan, Jinhe, 1230 m, 28.VI.1984, coll. Chen Yixin, 1♂. Guizhou (IZCAS): Meiyun, 6.IV.1978, coll. Xia
<normalizedToken originalValue="Huaien">Huai'en</normalizedToken>
, 1♀. Yunnan (IZCAS): Lijiang, 22-23.V.1980, 7♂1♀; Lijiang, Yushuizhai, 2680 m, 21.VI.2009, coll. Qi Feng, 1♂; Qujing, 3-20.VII.1982, coll. Wang Linyao and Song Shimei, 4♂; Luoci, 21.VI.1982, coll. Song Shimei, 1♂; Yongsheng, Liude, 2250m, 9.VII.1984, coll. Liu Dayun, 1♂. More material from Hubei, Sichuan, Yunnan, Tibet in coll. ZFMK.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="71" type="distribution">
<paragraph pageId="26" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="26" pageNumber="71">China (Henan, Zhejiang, Hubei, Hunan, Fujian, Guangxi, Sichuan, Guizhou, Yunnan, Tibet).</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="71" type="remarks">
<paragraph pageId="26" pageNumber="71">Remarks.</paragraph>
<paragraph pageId="26" pageNumber="71">
The female which
<bibRefCitation author="Oberthuer, C" journalOrPublisher="Etudes d'Entomologie" pageId="30" pageNumber="75" pagination="13 - 38" title="Nouveaux Lepidopteres du Thibet." volume="11" year="1886">
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
(1886
</bibRefCitation>
, pl. 5, fig.30) figured, is generally considered to be the holotype of thibetaria. However, he indirectly mentioned a larger number of specimens in his original description, by writing: &quot;some specimens have the wings crossed by curved medial lines which cut through the discoidal spots&quot; (translated from French). The female is part of the ZFMK collection, as already mentioned by
<bibRefCitation pageId="26" pageNumber="71">Wehrli (1941)</bibRefCitation>
, but all other syntypes are not. It is well possible that they have been transferred to The Natural History Museum, London, which keeps a large part of the Charles
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
collection. If these specimens will be found existing, the female will lose its holotype status. Eight specimens of thibetaria from several Sichuan localities in the ZFMK collection bearing the typical printed labels of
<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
have been collected later, in the years following the description of thibetaria, thus they do not belong to the syntype series.
</paragraph>
<caption pageId="26" pageNumber="71">
<paragraph pageId="26" pageNumber="71">
Figures 54-63. Adults of
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="genus">Biston</taxonomicName>
. 54
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston perclara" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="perclara">Biston perclara</taxonomicName>
, male 55-57
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston thibetaria" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="thibetaria">Biston thibetaria</taxonomicName>
. 55 male (syntype) 56 male 57 female 58-63
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston panterinaria subsp. panterinaria" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="panterinaria" subSpecies="panterinaria">Biston panterinaria panterinaria</taxonomicName>
. 58 male (syntype of
<taxonomicName class="Insecta" family="Geometridae" genus="Culcula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Culcula" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="genus">Culcula</taxonomicName>
panterinaria lienpingensis) 59 female (syntype of
<taxonomicName class="Insecta" family="Geometridae" genus="Culcula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Culcula panterinaria subsp. szechuanensis" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="panterinaria" subSpecies="szechuanensis">Culcula panterinaria szechuanensis</taxonomicName>
) 60 male (Dayu, Jiangxi) 61 male (Jiulianshan, Jiangxi) 62 male (Shixing, Guangdong) 63 male (Pengshui, Sichuan). Scale bar = 1 cm.
</paragraph>
</caption>
<caption pageId="26" pageNumber="71">
<paragraph pageId="26" pageNumber="71">
Figures 64-69. Adults of
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="genus">Biston</taxonomicName>
. 64-65
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston panterinaria subsp. panterinaria" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="panterinaria" subSpecies="panterinaria">Biston panterinaria panterinaria</taxonomicName>
. 64 female (Fangshan, Beijing) 65 female (Yingtaogou, Beijing); 66-69
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston panterinaria subsp. exanthemata" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="panterinaria" subSpecies="exanthemata">Biston panterinaria exanthemata</taxonomicName>
. 66 male 67 ditto, underside 68 female 69 ditto, underside. Scale bar = 1 cm.
</paragraph>
</caption>
<caption pageId="26" pageNumber="71">
<paragraph pageId="26" pageNumber="71">
Figures 70-75. Male genitalia of
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="genus">Biston</taxonomicName>
. 70
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston melacron" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="melacron">Biston melacron</taxonomicName>
71
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston marginata" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="marginata">Biston marginata</taxonomicName>
72
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston thoracicaria" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="thoracicaria">Biston thoracicaria</taxonomicName>
73
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston betularia subsp. parva" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="betularia" subSpecies="parva">Biston betularia parva</taxonomicName>
74
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston betularia subsp. nepalensis" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="betularia" subSpecies="nepalensis">Biston betularia nepalensis</taxonomicName>
75
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston robustum" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="robustum">Biston robustum</taxonomicName>
. Scale bar = 1 mm.
</paragraph>
</caption>
<caption pageId="26" pageNumber="71">
<paragraph pageId="26" pageNumber="71">
Figures 76-81. Male genitalia of
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="genus">Biston</taxonomicName>
. 76
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston mediolata" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="mediolata">Biston mediolata</taxonomicName>
sp. n. 77
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston contectaria" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="contectaria">Biston contectaria</taxonomicName>
78
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston bengaliaria" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="bengaliaria">Biston bengaliaria</taxonomicName>
79
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston pustulata" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="pustulata">Biston pustulata</taxonomicName>
80
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston suppressaria" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="suppressaria">Biston suppressaria</taxonomicName>
81
<taxonomicName class="Insecta" family="Geometridae" genus="Biston" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Biston regalis" order="Lepidoptera" pageId="26" pageNumber="71" phylum="Arthropoda" rank="species" species="regalis">Biston regalis</taxonomicName>
. Scale bar = 1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. &amp; Tishechkin, Alexey K." docDate="2013" docId="3EF3E2F914DE2DD2840BF3AB0D69737B" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus hamistrius Schmidt 1893, comb. n." docType="treatment" docVersion="3" lastPageNumber="210" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="207" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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<treatment ID-GBIF-Taxon="152041013" LSID="urn:lsid:plazi:treatment:3EF3E2F914DE2DD2840BF3AB0D69737B" httpUri="http://treatment.plazi.org/id/3EF3E2F914DE2DD2840BF3AB0D69737B" lastPageId="209" lastPageNumber="210" pageId="206" pageNumber="207">
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<paragraph pageId="206" pageNumber="207">
<taxonomicName LSID="http://species-id.net/wiki/Operclipygus_hamistrius" authority="Schmidt, 1893" authorityName="Schmidt" authorityYear="1893" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="206" pageNumber="207" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius (Schmidt, 1893)</taxonomicName>
<taxonomicNameLabel pageId="206" pageNumber="207">comb. n.</taxonomicNameLabel>
Figs 4B, 6F56B,
<normalizedToken originalValue="DF57AD">D-F57A-D</normalizedToken>
, IMap 20
</paragraph>
</subSubSection>
<subSubSection pageId="206" pageNumber="207" type="reference_group">
<paragraph pageId="206" pageNumber="207">
<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister hamistrius" order="Coleoptera" pageId="206" pageNumber="207" phylum="Arthropoda" rank="species" species="hamistrius">Phelister hamistrius</taxonomicName>
Schmidt, 1893: 13.
</paragraph>
</subSubSection>
<subSubSection pageId="206" pageNumber="207" type="type locality">
<paragraph pageId="206" pageNumber="207">Type locality.</paragraph>
<paragraph pageId="206" pageNumber="207">Not specified beyond Mexico.</paragraph>
</subSubSection>
<subSubSection lastPageId="207" lastPageNumber="208" pageId="206" pageNumber="207" type="type material">
<paragraph pageId="206" pageNumber="207">Type material.</paragraph>
<paragraph pageId="206" pageNumber="207">
Lectotype, here designated (ZMHB):
<normalizedToken originalValue="“Mexico.”/”hamistrius”">&quot;Mexico.&quot;/&quot;hamistrius&quot;</normalizedToken>
/ &quot;LECTOTYPE
<taxonomicName class="Insecta" family="Histeridae" genus="Phelister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phelister hamistrius" order="Coleoptera" pageId="206" pageNumber="207" phylum="Arthropoda" rank="species" species="hamistrius">Phelister hamistrius</taxonomicName>
Schmidt M.S.Caterino &amp; A.K.Tishechkin des. 2010&quot;. This species was described from an unspecified number of specimens, and the lectotype designation fixes primary type status on the only known original specimen.
</paragraph>
<paragraph lastPageId="207" lastPageNumber="208" pageId="206" pageNumber="207">
Other material. BELIZE: Cayo: 1: Las Cuevas Research Station,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="16.733334">16°44'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-88.98333">88°59'W</geoCoordinate>
, vi.2006, J. Kitson (BMNH), 1: 550m, v.1997, D. Inward (BMNH); 1: Chiquibul Forest Reserve, San Pastor, 13.xi.1994, FIT (BMNH); 1: Gran de Oro, 6.i.1995, FIT (BMNH). GUATEMALA: Baja Verapaz: 1: 7km E Purulha, 1600m, 23.v.1991, cloud forest litter, R. Anderson (SEMC); 1: 17km N Salama Hy 5, 1700m, 29.
<normalizedToken originalValue="vi">vi-</normalizedToken>
3.vii.1993, FIT, J. Ashe &amp; R. Brooks (SEMC); Zacapa: 2: 3.5km SE La Union, 1500m, 25-27.vi.1993, FIT, J. Ashe &amp; R. Brooks (SEMC, FMNH); 3: 23-25.vi.1993 (SEMC, MSCC, AKTC). HONDURAS: Atlantida: 2: 15km W La Ceiba, 15-19.vi.1996, FIT, tropical rainforest, R. Lehman (TAMU), 1: 9-30.vii.1996, Malaise trap, tropical rainforest, R. Lehman (TAMU);
<normalizedToken originalValue="Cortés">Cortes</normalizedToken>
: 2: Parque Nac. Cusuco, El Cortecito, 361198, 1717050, 1250m, 4-7.vi.2006, FIT, mature broadleaf forest, J. Nunez-Mino (OUMNH); 1: Parque Nac. Cusuco,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="15.483334">15°29'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-88.21667">88°13'W</geoCoordinate>
, 30.viii.1995, Malaise trap, oak/pine cloud forest, R. Cave, (AKTC); 1: 15.vii.1995, FIT, R. Cordero (AKTC); 1: Parque Nac. Cerro Azul-Meambar, Los Pinos,
<geoCoordinate direction="north" orientation="latitude" precision="92" value="14.873333">14°52.4'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-87.91167">87°54.7'W</geoCoordinate>
, 800m, 10-16.v.2002, FIT, secondary forest, S. Peck (CHND); Francisco
<normalizedToken originalValue="Morazán">Morazan</normalizedToken>
: 1: Yuscaran Cerro Montserrat, 28.i.1994, W. Morjan (AKTC); Olancho: 1: Parque Nac. La Muralla, 15km N La Union,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="15.116667">15°07'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-86.75">86°45'W</geoCoordinate>
, ii.1995, Malaise trap, high elevation rainforest, R. Cave (AKTC); 1: La Union, Parque Nac. La Muralla,
<pageBreakToken pageId="207" pageNumber="208" start="start">x</pageBreakToken>
.1994, R. Cordero (AKTC), 2: xi.1994, I. Maradiaga (AKTC), 2: 20-31.viii.1994, R. Cordero (AKTC). NICARAGUA, Granada: 1: Volcan Mombacho, San Joaquin #3, 21.vi.1998, malaise trap, J.M. Maes (MEL); 1: Volcan Mombacho, El Progreso #3, 15.iv.1998, malaise trap, J.M. Maes (MEL); Matagalpa: 1: 6 km N Matagalpa, Selva Negra Hotel,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="12.999833">12°59.99'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-85.90884">85°54.53'W</geoCoordinate>
, 1530m, 20-22.v.2002, FIT, R. Brooks, Z. Falin, S. Chatzimanolis (SEMC); 6: Selva Negra Reserve,
<geoCoordinate direction="north" orientation="latitude" precision="92" value="13.005">13°0.3'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-85.903336">85°54.2'W</geoCoordinate>
, 1395m, FIT, 19-27.v.2012, A. Derunkov, DNA Extracts MSC-2334, MSC-2349, MSC-2350, MSC-2351 (SBMNH, MSCC, AKTC).
</paragraph>
</subSubSection>
<subSubSection lastPageId="209" lastPageNumber="210" pageId="207" pageNumber="208" type="description">
<paragraph pageId="207" pageNumber="208">Diagnostic description.</paragraph>
<paragraph lastPageId="209" lastPageNumber="210" pageId="207" pageNumber="208">
Length: 1.78-2.25 mm, width: 1.44-1.84 mm; body rufescent, elongate oval, outline slightly interrupted at humeri, pronotum more nearly parallel-sided, moderately convex; frons shallowly depressed at middle, with conspicuous, fine ground punctation; frontal stria weakly divergent between eyes, anterior portion varied from complete to absent between antennal bases, occasionally represented only by isolated fragments; apical margin of labrum distinctly emarginate; left mandible untoothed, right mandible with small, acute basal tooth; pronotum lacking prescutellar impression, disk with ~15 small coarse punctures at sides; marginal stria of pronotum descending from dorsal margin anteriorly to hypomeron about one-fourth behind anterior corners, markedly disrupting margin; lateral submarginal stria very close to sides, frequently weak, fragmented toward base, either continuous across front or interrupted, the central portion barely detached with its ends faintly recurved posterad; median pronotal gland openings about two-thirds pronotal length from anterior margin; elytra with two complete epipleural striae, outer subhumeral stria present in apical half, inner subhumeral stria absent, striae 1-4 complete, 5th stria present in apical third, sutural stria present in apical half; nearly entire venter bearing waves of microsculpture; prosternal keel weakly emarginate at base, carinal striae complete, narrowed between coxae, with faint microsculpture between, meeting in bulbous anterior arch, weak secondary striae present behind prosternal gland openings; prosternal lobe with marginal stria abbreviated at sides; anterior margin of mesoventrite weakly projecting, marginal stria complete or interrupted; mesometaventral stria strongly arched forward very close to anterior mesoventral margin, continued by lateral metaventral stria toward metacoxa, frequently interrupted in posterior half of metaventral disk; central portion of metaventral disk with conspicuous ground punctation, particularly toward posterior margin; 1st abdominal ventrite with single, incomplete lateral stria, punctures of sides of disk simple, separate, not coalescing or forming strigosity; propygidium and pygidium with very conspicuous transversely reticulate microscultpure; propygidium with sparse small, shallow punctures, separated 2
<normalizedToken originalValue="3×">-3x</normalizedToken>
their diameters; pygidium with smaller punctures separated by only about 1.5
<normalizedToken originalValue="×">x</normalizedToken>
their diameters; marginal pygidial stria fine, present only around apical half of margin, occasionally interrupted near apex. Male genitalia (Figs 57
<normalizedToken originalValue="AD">A-D</normalizedToken>
, I): accessory sclerites well developed; T8 with sides straight, weakly convergent to near apex, with faint desclerotization and abrupt step to apex (especially evident in lateral view), basal emargination narrow, rather shallow, reaching only halfway to basal membrane attachment line, ventrolateral apodemes symmetrical along underside nearly meeting at midline; S8 rather
<pageBreakToken pageId="208" pageNumber="209" start="start">short</pageBreakToken>
, with sides weakly convergent toward apex, apical guides gradually more strongly developed toward apex, apices obliquely subacute, ventral surface distinctly wrinkled, halves approximate near base, weakly diverging toward apex; T9 with sides moderately
<pageBreakToken pageId="209" pageNumber="210" start="start">rounded</pageBreakToken>
, apices convergent, subacute; T10 with halves separate; S9 with broad, weakly arcuate base, narrowed apically, apex shallowly arcuate but without fine median emargination, apical flange entire; tegmen narrow, elongate, widest just beyond midpoint, apex subacute, with medioventral process thin but well sclerotized, narrowly
<normalizedToken originalValue="U">'U'</normalizedToken>
-shaped, projecting beneath, about one-fourth from tegmen base; basal piece about one-third tegmen length; median lobe thin, no more than one-third tegmen length.
</paragraph>
</subSubSection>
<subSubSection pageId="209" pageNumber="210" type="remarks">
<paragraph pageId="209" pageNumber="210">Remarks.</paragraph>
<paragraph pageId="209" pageNumber="210">The nominate species in this group can be recognized by the combination of: simple punctures on sides of the 1st abdominal ventrite (Fig. 56E), rather than elongate gouges; the descent of the marginal pronotal stria to the hypomeron about one-fourth behind the anterior corner, usually conspicuously disrupting the margin (Fig. 56F); the frequently complete frontal stria; and the usually continuous lateral and anterior submarginal striae (Fig. 56D). The unique type specimen, from an unspecified locality in Mexico, does not correspond extremely well with specimens from parts of northern Central America, having the pronotal striae broken, in particular, but they are certainly close. More material from Mexico would help substantiate the differences, and perhaps justify the separation of populations from most of these other areas as a distinct species.</paragraph>
<caption pageId="209" pageNumber="210">
<paragraph pageId="209" pageNumber="210">
Figure 56.
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
group. A Pygidial microsculpture of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus arquus" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="arquus">Operclipygus arquus</taxonomicName>
B Pygidial microsculpture of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
C Pronotum of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus arquus" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="arquus">Operclipygus arquus</taxonomicName>
D Dorsal habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
E Ventral habitus of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
F Lateral view of pronotum of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="209" pageNumber="210">
<paragraph pageId="209" pageNumber="210">
Figure 57. Male genitalia of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
group. A T8 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
B S8 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
C T9 &amp; T10 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
D S9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
E T8 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus quinquestriatus" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="quinquestriatus">Operclipygus quinquestriatus</taxonomicName>
F S8 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus quinquestriatus" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="quinquestriatus">Operclipygus quinquestriatus</taxonomicName>
G T9 &amp; T10 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus quinquestriatus" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="quinquestriatus">Operclipygus quinquestriatus</taxonomicName>
H S9 of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus quinquestriatus" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="quinquestriatus">Operclipygus quinquestriatus</taxonomicName>
I Aedeagus, dorsal and lateral views, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hamistrius" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="hamistrius">Operclipygus hamistrius</taxonomicName>
J Aedeagus, dorsal and lateral views, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus geometricus" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="geometricus">Operclipygus geometricus</taxonomicName>
K Aedeagus, dorsal and lateral views, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus arquus" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="arquus">Operclipygus arquus</taxonomicName>
L Aedeagus, lateral view, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus rufescens" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="rufescens">Operclipygus rufescens</taxonomicName>
M Aedeagus, dorsal and lateral views, of
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus chiapensis" order="Coleoptera" pageId="209" pageNumber="210" phylum="Arthropoda" rank="species" species="chiapensis">Operclipygus chiapensis</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>