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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.56.519" ID-GBIF-Dataset="5cd1bde1-9049-456c-83cf-1b3faa7fa9cc" ID-PMC="PMC3088330" ID-Pensoft-Pub="1313-2970-56-65" ID-PubMed="21594173" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-56-65" ModsDocOrigin="ZooKeys 56" ModsDocTitle="New data on Neotropical Scolytus Geoffroy, 1762 with description of five new species from Peru (Coleoptera, Curculionidae, Scolytinae)" checkinTime="1451251007590" checkinUser="pensoft" docAuthor="V. Petrov, Alexander &amp; Y. Mandelshtam, Michail" docDate="2010" docId="3E6D5698D2A93F4C4F3D9EB200244706" docLanguage="en" docName="ZooKeys 56: 65-104" docOrigin="ZooKeys 56" docSource="http://dx.doi.org/10.3897/zookeys.56.519" docTitle="Scolytus vagabundus Petrov &amp; Mandelshtam, 2010, sp. n." docType="treatment" docVersion="5" lastPageNumber="93" masterDocId="FF8FFF9ED926FFB6DE0FA13AA40FCA11" masterDocTitle="New data on Neotropical Scolytus Geoffroy, 1762 with description of five new species from Peru (Coleoptera, Curculionidae, Scolytinae)" masterLastPageNumber="104" masterPageNumber="65" pageNumber="92" updateTime="1668162715365" updateUser="ExternalLinkService">
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<mods:title>New data on Neotropical Scolytus Geoffroy, 1762 with description of five new species from Peru (Coleoptera, Curculionidae, Scolytinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>V. Petrov, Alexander</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Y. Mandelshtam, Michail</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>56</mods:number>
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<mods:extent unit="page">
<mods:start>65</mods:start>
<mods:end>104</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.56.519</mods:url>
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<treatment ID-GBIF-Taxon="159359797" LSID="urn:lsid:zoobank.org:act:113AFEB7-9AB8-4E8A-8068-E0ECE0A00633" httpUri="http://treatment.plazi.org/id/3E6D5698D2A93F4C4F3D9EB200244706" lastPageId="28" lastPageNumber="93" pageId="27" pageNumber="92">
<subSubSection pageId="27" pageNumber="92" type="nomenclature">
<paragraph pageId="27" pageNumber="92">
<taxonomicName LSID="urn:lsid:zoobank.org:act:113AFEB7-9AB8-4E8A-8068-E0ECE0A00633" class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus vagabundus" order="Coleoptera" pageId="27" pageNumber="92" phylum="Arthropoda" rank="species" species="vagabundus">Scolytus vagabundus</taxonomicName>
<taxonomicNameLabel pageId="27" pageNumber="92">sp. n.</taxonomicNameLabel>
Figs 2425
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="92" type="type locality">
<paragraph pageId="27" pageNumber="92">Type locality.</paragraph>
<paragraph pageId="27" pageNumber="92">Peru, Loreto province, left bank of Amazon River.</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="92" type="type material">
<paragraph pageId="27" pageNumber="92">Type material.</paragraph>
<paragraph pageId="27" pageNumber="92">Holotype ♀ (ZMM): PERU: LORETO PROVINCE: left bank of Amazon River, 70 km SSW from Iquitos to Nauta, 130 m a.s.l., 1.03.2008, leg.. A.V. Petrov.</paragraph>
<caption pageId="27" pageNumber="92">
<paragraph pageId="27" pageNumber="92">
Figure 24. Habitus of
<taxonomicName class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus vagabundus" order="Coleoptera" pageId="27" pageNumber="92" phylum="Arthropoda" rank="species" species="vagabundus">Scolytus vagabundus</taxonomicName>
, female
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="27" pageNumber="92" type="diagnosis">
<paragraph pageId="27" pageNumber="92">Diagnosis.</paragraph>
<paragraph pageId="27" pageNumber="92">
Species closely related to
<taxonomicName class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus carveli" order="Coleoptera" pageId="27" pageNumber="92" phylum="Arthropoda" rank="species" species="carveli">Scolytus carveli</taxonomicName>
and
<taxonomicName class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus pinnatus" order="Coleoptera" pageId="27" pageNumber="92" phylum="Arthropoda" rank="species" species="pinnatus">Scolytus pinnatus</taxonomicName>
Eggers, from which it can be distinguished by body size, by second sternite tubercle form and its position on sternite, by presence of tubercle on fifth sternite and by abdominal vestiture. From
<taxonomicName class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus pinnatus" order="Coleoptera" pageId="27" pageNumber="92" phylum="Arthropoda" rank="species" species="pinnatus">Scolytus pinnatus</taxonomicName>
new species can be distinguished also by absence of tubercles and denticles on fourth sternite.
</paragraph>
</subSubSection>
<subSubSection lastPageId="28" lastPageNumber="93" pageId="27" pageNumber="92" type="description">
<paragraph pageId="27" pageNumber="92">Description.</paragraph>
<paragraph pageId="27" pageNumber="92">Female: body length 2.2 mm, 2.2 times as long as wide; colour dark brown. Head dark grayish-brown, nearly black. Frons convex from eye to eye from epistoma to vertex; surface aciculate from vertex to epistoma. Vestiture scant, hairs short and sparse, conspicuous in lateral parts of front. Antenna brown, densely covered by short golden hairs, club elliptical with evenly rounded apex. Pronotum 1.1 times as long as wide, its maximal width at the half of its length, from center to apex pronotum seems more elongated. Pronotal surface shining, punctured by deep points at base and in central part of disk; punctures of lateral sides of pronotum and its apical portion slightly larger. Apical portion of pronotum with few short dark hairs. Pronotum has a well defined acute lateral margin separating it from prothorax (propleura). Lateral sides of prothorax (propleura) are abundantly and evenly punctured by punctures of moderate size.</paragraph>
<paragraph pageId="27" pageNumber="92">Scutellum triangular, deeply set in scutellar impression.</paragraph>
<paragraph lastPageId="28" lastPageNumber="93" pageId="27" pageNumber="92">
Elytra dark grayish brown with reddish brown declivity. Elytra 1.1 times as long as wide, 1.0 times as long as pronotum; lateral elytral margins slightly widened from base to middle of their length, these sides are evenly rounded then towards elytral declivity. Elytral surface shining, punctured with sparse punctures in striae. Striae and interstriae are deepened, interstriae with few and sparse punctures, evident only in posterior part of elytra. In the posterior part of elytra and on declivity, interstriae with rows of pale sparse erect hairs. Abdomen reddish brown. Second sternite set vertically, perpendicular to first sternite, anterior margin of second sternite weakly rounded, not costate, sternite two armed by a large, laterally compressed median spine, this spine basis runs through second sternite from its base and up to posterior margin. On dorsum of the spine a sharpened denticle directed by its apex downwards; this denticle located closer to base of second sternite. Lateral sides of second and third sternites with small shar
<pageBreakToken pageId="28" pageNumber="93" start="start">pened</pageBreakToken>
denticles. Fifth sternite with a median tubercle. Surface of second sternite smooth and shining, without puncturation, other sternites with minute dense puncturation. Abdomen densely covered by yellow hairs of moderate length, hair apices directed toward tubercle of fifth sternite. Legs reddish brown, with golden hairs.
</paragraph>
<paragraph pageId="28" pageNumber="93">Male: Unknown.</paragraph>
<caption pageId="28" pageNumber="93">
<paragraph pageId="28" pageNumber="93">
Figure 25. Sternites of abdomen of
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, female
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="28" pageNumber="93" type="distribution">
<paragraph pageId="28" pageNumber="93">Distribution.</paragraph>
<paragraph pageId="28" pageNumber="93">Known only from the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="93" type="etymology">
<paragraph pageId="28" pageNumber="93">Etymology.</paragraph>
<paragraph pageId="28" pageNumber="93">
Species name originates from the Hispan word
<normalizedToken originalValue="“vagabundo”">&quot;vagabundo&quot;</normalizedToken>
(vagabond).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="84B0EAE78041FFAEFF3E7617DDADFA85" bold="true" box="[151,457,1377,1404]" italics="true" pageId="16" pageNumber="321">Lepidodactylus dialeukos</emphasis>
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.
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, mature female, collected by
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[
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,
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,
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1938.
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<emphasis id="84B0EAE78041FFAEFF6E7546DD5DF9B0" box="[199,313,1584,1609]" italics="true" pageId="16" pageNumber="321">Diagnosis</emphasis>
. A medium-sized species (adult SVL
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) species of
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<emphasis id="84B0EAE78041FFAEFC767546D8EEF9B3" box="[991,1162,1584,1610]" italics="true" pageId="16" pageNumber="321">Lepidodactylus</emphasis>
</taxonomicName>
having a subcylindrical tail without a lateral fringe of enlarged scales; one divided subterminal scansor on digits; 14 enlarged scales of the pore-bearing series limited to precloacal region; 910 T4 lamellae, 89 T1 lamellae; toes relatively long (T4/SVL = 0.12) and wide (T4W/T4L = 0.36), poorly webbed (T3T4webL/T4L = 0.20), with lamellae covering only about half their length (T4lamellaeL/4TL = 0.53); nine small precloacal scales between apex of pore-bearing series and cloaca; dorsum uniformly gray with a row of dorsolateral white spots on each side, lacking dark spots in this region; white postocular stripe; narrow bands or patches of white scales on dorsal surfaces of forearms, wrists, shanks, ankles, and digits; and black punctations widely scattered ventrally.
</paragraph>
<paragraph id="B67B36F58041FFAFFF6E7426D8D3FD9A" blockId="16.[151,1437,1511,2042]" lastBlockId="17.[151,1437,152,2031]" lastPageId="17" lastPageNumber="322" pageId="16" pageNumber="321">
<emphasis id="84B0EAE78041FFAEFF6E7426DE4EF890" box="[199,554,1872,1898]" italics="true" pageId="16" pageNumber="321">Comparisons with other species</emphasis>
. The subcylindrical tail without a lateral fringe of enlarged scales and the single divided subterminal scansors under the toes place
<taxonomicName id="71C44D768041FFAEFD607402DF83F877" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[713,999,1908,1934]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="321" phylum="Chordata" rank="species" species="dialeukos" status="sp. nov.">
<emphasis id="84B0EAE78041FFAEFD607402DF83F877" box="[713,999,1908,1934]" italics="true" pageId="16" pageNumber="321">Lepidodactylus dialeukos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8041FFAEFC477402D823F877" box="[1006,1095,1908,1934]" pageId="16" pageNumber="321" rank="species">
<emphasis id="84B0EAE78041FFAEFC477402D823F877" bold="true" box="[1006,1095,1908,1934]" pageId="16" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
in
<bibRefCitation id="D2554B048041FFAEFBC27402D9F8F877" author="Brown, W. C. &amp; Parker, F." box="[1131,1436,1908,1934]" pageId="16" pageNumber="321" pagination="253 - 265" refId="ref18821" refString="Brown, W. C. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Sciences, 41, 253 - 265." type="journal article" year="1977">Brown and Parkers (1977)</bibRefCitation>
Group II.
<taxonomicName id="71C44D768041FFAEFEA174EEDE42F84B" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[264,550,1944,1970]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="321" phylum="Chordata" rank="species" species="dialeukos" status="sp. nov.">
<emphasis id="84B0EAE78041FFAEFEA174EEDE42F84B" box="[264,550,1944,1970]" italics="true" pageId="16" pageNumber="321">Lepidodactylus dialeukos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8041FFAEFD8674EEDEEDF84B" box="[559,649,1944,1970]" pageId="16" pageNumber="321" rank="species">
<emphasis id="84B0EAE78041FFAEFD8674EEDEEDF84B" bold="true" box="[559,649,1944,1970]" pageId="16" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from all other Melanesian members of this group in being the only species with only a single divided subterminal scansor (
<emphasis id="84B0EAE78041FFAEFCEC74CADFE9F82C" box="[837,909,1980,2005]" italics="true" pageId="16" pageNumber="321">versus</emphasis>
having
<quantity id="713C9B108041FFAEFC4174CAD84BF82F" box="[1000,1071,1980,2006]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="16" pageNumber="321" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">24 in</quantity>
the other species) and extensive white markings. It further differs from other Melanesian members of this group as follows: from
<taxonomicName id="71C44D768041FFAEFB707496D927F800" box="[1241,1347,2016,2041]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="321" phylum="Chordata" rank="species" species="guppyi">
<emphasis id="84B0EAE78041FFAEFB707496D927F800" box="[1241,1347,2016,2041]" italics="true" pageId="16" pageNumber="321">L. guppyi</emphasis>
</taxonomicName>
,
<emphasis id="84B0EAE78041FFAFFAF87496DCADFF4A" italics="true" lastPageId="17" lastPageNumber="322" pageId="16" pageNumber="321">L. pul- cher</emphasis>
, and
<taxonomicName id="71C44D768040FFAFFEA373ECDD13FF4A" baseAuthorityName="Brown &amp; Tanner" baseAuthorityYear="1949" box="[266,375,153,179]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="shebae">
<emphasis id="84B0EAE78040FFAFFEA373ECDD13FF4A" box="[266,375,153,179]" italics="true" pageId="17" pageNumber="322">L. shebae</emphasis>
</taxonomicName>
in having fewer enlarged precloacal scales (14
<emphasis id="84B0EAE78040FFAFFC3073ECDF85FF4A" box="[921,993,154,179]" italics="true" pageId="17" pageNumber="322">versus</emphasis>
3952, 1820, and 34, respectively, in the other species), and further from
<taxonomicName id="71C44D768040FFAFFD8473C8DEFEFF2E" baseAuthorityName="Brown &amp; Tanner" baseAuthorityYear="1949" box="[557,666,189,215]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="shebae">
<emphasis id="84B0EAE78040FFAFFD8473C8DEFEFF2E" box="[557,666,189,215]" italics="true" pageId="17" pageNumber="322">L. shebae</emphasis>
</taxonomicName>
in its larger adult size (SVL
<quantity id="713C9B108040FFAFFC4873CAD822FF2E" box="[993,1094,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="17" pageNumber="322" unit="mm" value="46.5">46.5 mm</quantity>
<emphasis id="84B0EAE78040FFAFFBE473C8D8F1FF2E" box="[1101,1173,190,215]" italics="true" pageId="17" pageNumber="322">versus</emphasis>
~
<quantity id="713C9B108040FFAFFB0273CBD89FFF2E" box="[1195,1275,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="17" pageNumber="322" unit="mm" value="36.0">36 mm</quantity>
in
<taxonomicName id="71C44D768040FFAFFA8973C8D9E8FF2E" baseAuthorityName="Brown &amp; Tanner" baseAuthorityYear="1949" box="[1312,1420,189,215]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="shebae">
<emphasis id="84B0EAE78040FFAFFA8973C8D9E8FF2E" box="[1312,1420,189,215]" italics="true" pageId="17" pageNumber="322">L. shebae</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="71C44D768040FFAFFEEC7394DE65FF02" box="[325,513,226,251]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE78040FFAFFEEC7394DE65FF02" box="[325,513,226,251]" italics="true" pageId="17" pageNumber="322">L. novaeguineae</emphasis>
</taxonomicName>
in having less coverage of the fourth toe by lamellae (T4lamellaeL/4TL = 0.53
<emphasis id="84B0EAE78040FFAFFF3E7270DCBBFEE6" box="[151,223,262,287]" italics="true" pageId="17" pageNumber="322">versus</emphasis>
<quantity id="713C9B108040FFAFFF4F7272DD10FEE6" box="[230,372,260,287]" metricMagnitude="-2" metricUnit="m" metricValue="2.0193" metricValueMax="2.3876" metricValueMin="1.651" pageId="17" pageNumber="322" unit="in" value="0.7949999999999999" valueMax="0.94" valueMin="0.65">0.650.94 in</quantity>
<taxonomicName id="71C44D768040FFAFFED27270DE50FEE6" box="[379,564,262,287]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE78040FFAFFED27270DE50FEE6" box="[379,564,262,287]" italics="true" pageId="17" pageNumber="322">L. novaeguineae</emphasis>
</taxonomicName>
), a longer fourth toe (T4/SVL = 0.12
<emphasis id="84B0EAE78040FFAFFC487270D84DFEE6" box="[993,1065,262,287]" italics="true" pageId="17" pageNumber="322">versus</emphasis>
<quantity id="713C9B108040FFAFFB997273D8AFFEE6" box="[1072,1227,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="2.4892" metricValueMax="2.7939999999999996" metricValueMin="2.1844" pageId="17" pageNumber="322" unit="in" value="0.098" valueMax="0.11" valueMin="0.086">0.0860.11 in</quantity>
<taxonomicName id="71C44D768040FFAFFB7B7270D9EFFEE6" box="[1234,1419,262,287]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE78040FFAFFB7B7270D9EFFEE6" box="[1234,1419,262,287]" italics="true" pageId="17" pageNumber="322">L. novaeguineae</emphasis>
</taxonomicName>
), a white postocular stripe (absent in
<taxonomicName id="71C44D768040FFAFFD82725CDE82FEBA" box="[555,742,298,323]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE78040FFAFFD82725CDE82FEBA" box="[555,742,298,323]" italics="true" pageId="17" pageNumber="322">L. novaeguineae</emphasis>
</taxonomicName>
), dorsum with a dorsolateral row of white spots (absent in
<taxonomicName id="71C44D768040FFAFFA2F725CDD56FE9E" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE78040FFAFFA2F725CDD56FE9E" italics="true" pageId="17" pageNumber="322">L. novaeguineae</emphasis>
</taxonomicName>
), and bands or patches of white scales on dorsal surfaces of digits (absent in
<taxonomicName id="71C44D768040FFAFFB257238D921FE9E" box="[1164,1349,334,359]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE78040FFAFFB257238D921FE9E" box="[1164,1349,334,359]" italics="true" pageId="17" pageNumber="322">L. novaeguineae</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="71C44D768040FFAFFEAC7204DDE1FE72" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[261,389,369,395]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFEAC7204DDE1FE72" box="[261,389,369,395]" italics="true" pageId="17" pageNumber="322">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFE257207DD81FE72" box="[396,485,369,395]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFE257207DD81FE72" bold="true" box="[396,485,369,395]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
in its larger adult size (SVL =
<quantity id="713C9B108040FFAFFC977206DFC6FE72" box="[830,930,368,395]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="17" pageNumber="322" unit="mm" value="46.5">46.5 mm</quantity>
<emphasis id="84B0EAE78040FFAFFC017204DF94FE72" box="[936,1008,370,395]" italics="true" pageId="17" pageNumber="322">versus</emphasis>
mean 40.3, range =
<quantity id="713C9B108040FFAFFB7C7206D907FE72" box="[1237,1379,368,395]" metricMagnitude="-2" metricUnit="m" metricValue="4.025" metricValueMax="4.55" metricValueMin="3.5" pageId="17" pageNumber="322" unit="mm" value="40.25" valueMax="45.5" valueMin="35.0">3545.5 mm</quantity>
in
<taxonomicName id="71C44D768040FFAFFA2F7204DC9DFE56" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFA2F7204DC9DFE56" italics="true" pageId="17" pageNumber="322">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFEA972E3DD3DFE56" box="[256,345,405,431]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFEA972E3DD3DFE56" bold="true" box="[256,345,405,431]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
), white postocular stripe (absent in
<taxonomicName id="71C44D768040FFAFFD4172E0DF03FE56" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[744,871,405,431]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFD4172E0DF03FE56" box="[744,871,405,431]" italics="true" pageId="17" pageNumber="322">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFCC772E3DFA3FE56" box="[878,967,405,431]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFCC772E3DFA3FE56" bold="true" box="[878,967,405,431]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
), dorsum with a dorsolateral row of white spots (absent in
<taxonomicName id="71C44D768040FFAFFEE572CCDDAFFE2A" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[332,459,441,467]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFEE572CCDDAFFE2A" box="[332,459,441,467]" italics="true" pageId="17" pageNumber="322">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFE7B72CFDE4FFE2A" box="[466,555,441,467]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFE7B72CFDE4FFE2A" bold="true" box="[466,555,441,467]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
), and bands or patches of white scales on dorsal surfaces of digits (absent in
<taxonomicName id="71C44D768040FFAFFA2F72CCDC9DFE0E" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFA2F72CCDC9DFE0E" italics="true" pageId="17" pageNumber="322">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFEA972ABDD3DFE0E" box="[256,345,477,503]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFEA972ABDD3DFE0E" bold="true" box="[256,345,477,503]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
). It differs from
<taxonomicName id="71C44D768040FFAFFDBD72A8DED7FE0E" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[532,691,477,503]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFDBD72A8DED7FE0E" box="[532,691,477,503]" italics="true" pageId="17" pageNumber="322">L. kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFD1072ABDF77FE0E" box="[697,787,477,503]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFD1072ABDF77FE0E" bold="true" box="[697,787,477,503]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
in having fewer T4 lamellae (10
<emphasis id="84B0EAE78040FFAFFB2F72A8D8AAFE0E" box="[1158,1230,478,503]" italics="true" pageId="17" pageNumber="322">versus</emphasis>
<quantity id="713C9B108040FFAFFB7D72ABD953FE0E" box="[1236,1335,477,503]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="5.08" metricValueMin="3.048" pageId="17" pageNumber="322" unit="in" value="16.0" valueMax="20.0" valueMin="12.0">1220 in</quantity>
<taxonomicName id="71C44D768040FFAFFA9772A8DC84FDE2" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFA9772A8DC84FDE2" italics="true" pageId="17" pageNumber="322">L. kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFF4E7177DD24FDE2" box="[231,320,513,539]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFF4E7177DD24FDE2" bold="true" box="[231,320,513,539]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
), dorsum uniform gray (boldly patterned with brown in
<taxonomicName id="71C44D768040FFAFFC187174D82BFDE2" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[945,1103,513,539]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFC187174D82BFDE2" box="[945,1103,513,539]" italics="true" pageId="17" pageNumber="322">L. kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFBFF7177D8CBFDE2" box="[1110,1199,513,539]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFBFF7177D8CBFDE2" bold="true" box="[1110,1199,513,539]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
), bands or patches of white scales on dorsal surfaces of digits (absent in
<taxonomicName id="71C44D768040FFAFFD7B7150DF15FDC6" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[722,881,549,575]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFD7B7150DF15FDC6" box="[722,881,549,575]" italics="true" pageId="17" pageNumber="322">L. kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFCD17153DFB6FDC6" box="[888,978,549,575]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFCD17153DFB6FDC6" bold="true" box="[888,978,549,575]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
), and black punctations widely scattered ventrally (many posteroventral and plantar scales uniformly brown in
<taxonomicName id="71C44D768040FFAFFC01713CD822FD9A" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[936,1094,585,611]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="322" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE78040FFAFFC01713CD822FD9A" box="[936,1094,585,611]" italics="true" pageId="17" pageNumber="322">L. kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8040FFAFFBE4713FD8C3FD9A" box="[1101,1191,585,611]" pageId="17" pageNumber="322" rank="species">
<emphasis id="84B0EAE78040FFAFFBE4713FD8C3FD9A" bold="true" box="[1101,1191,585,611]" pageId="17" pageNumber="322">sp. nov.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
<paragraph id="B67B36F58040FFAFFF6E7118D9F6FBCD" blockId="17.[151,1437,152,2031]" pageId="17" pageNumber="322">
<emphasis id="84B0EAE78040FFAFFF6E7118DDA9FD7E" box="[199,461,621,647]" italics="true" pageId="17" pageNumber="322">
Description of
<typeStatus id="697F88578040FFAFFEC4711BDDA9FD7E" box="[365,461,621,647]" pageId="17" pageNumber="322" type="holotype">holotype</typeStatus>
</emphasis>
. A female of medium size (SVL =
<quantity id="713C9B108040FFAFFCE6711ADFD0FD7E" box="[847,948,620,647]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="17" pageNumber="322" unit="mm" value="46.5">46.5 mm</quantity>
, TrL = 16.0 mm). Head relatively long (HL/ SVL = 0.19) and wide (HW/HL = 0.85), distinct from neck. Loreal region inflated; no distinct canthus rostralis. Top of snout, area between nares, and area posterior to nares concave. Snout tapered and rounded at tip, long (SN/HL = 0.60), almost twice as long as eye diameter (SN/EY = 1.9). Eye relatively large (EY/HL = 0.32, EY/EN = 0.70); pupil vertical, constricted into series of four lobes on right, rounded on left; supraciliaries same size as adjacent granules. Ear opening small (Ear/HL = 0.080), approximately round; distance between ear and eye larger than eye diameter (EE/EY = 1.4). Rostral wider (
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) than high (
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), highest just medial to nares, lower between these points; length (
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). Scales on snout tip undifferentiable due to loss of keratin layer. External nares circular. Mental shallowly triangular, wider (
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) than long (
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), with a short, medial suture posteriorly. Mental bordered posteriorly by three enlarged postmentals, these bordered posteriorly by three rows of enlarged gulars that are followed by smaller scales decreasing in size to granular chin scales. First four infralabials bordered below by somewhat enlarged scales; remaining scales below infralabials of approximately twice size of central throat scales. Supralabials to mid-orbital position ten on each side; to angle of jaw 13 on each side. Infralabials 11 on each side.
</paragraph>
<paragraph id="B67B36F58040FFAFFF6E774BD817FB65" blockId="17.[151,1437,152,2031]" pageId="17" pageNumber="322">Body of modest habitus (TrL/SVL = 0.34), slightly depressed. Dorsal scales on head, body, limbs, and throat tiny, juxtaposed granules, larger on sides and snout; tubercles absent. Ventral scales larger, flatter but somewhat beaded, smooth, subimbricate, gradually decreasing in size laterally to become granular.</paragraph>
<paragraph id="B67B36F58040FFAFFF6E77D3D8BDFADE" blockId="17.[151,1437,152,2031]" pageId="17" pageNumber="322">Enlarged precloacal scales in single series of 14, all but one bearing a tiny indentation but lacking true pores with sebaceous plugs; no enlarged scales on thighs. Enlarged scales form a pubic patch between the precloacal series and vent; no tiny scales inserted between the precloacal series and the pubic patch on either side; ten scales in a row between apex of enlarged precloacal series and vent. Scales on palms and soles rounded, smooth.</paragraph>
<paragraph id="B67B36F58040FFAFFF6E7646DF21F9C6" blockId="17.[151,1437,152,2031]" pageId="17" pageNumber="322">
Fore- and hindlimbs relatively small but well-developed (FA/SVL = 0.11, CS/SVL = 0.11). Digits well-developed, fairly widely dilated throughout their length (T4W/T4L = 0.36), all but first fingers and toes with recurved claws; clawed phalanges laterally compressed, free above and extending slightly beyond terminal scansors. Subdigital lamellae narrow and smooth, all digits with single divided subterminal scansor; lamellae extend for approximately half length of each toe (T4 scansor L/T4L = 0.53). Lamellae of manus
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8 on right,
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98 on left; of pes
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?7 on right,
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7 on left; fourth toe on right pes missing tip. Relative lengths of digits on manus and pes I &lt;II &lt;V &lt;III &lt;IV. Webbing present between all digits, basal between most digits but more extensive between T3 and T4 (T3T4webL/T4L = 0.20).
</paragraph>
<paragraph id="B67B36F58040FFAFFF6E753FD928F936" blockId="17.[151,1437,152,2031]" pageId="17" pageNumber="322">Tail appears to be entirely original, subcylindrical, somewhat compressed, relatively narrow (TW/SVL = 0.080); lateral margins without skin flanges or spines. Scales of tail small, flat, smooth, subimbricate, slightly larger ventrally than dorsally. Cloacal sacs swollen, with small external orifices situated near lateral margins of vent; no postcloacal spur on either side of tailbase; midventral scales of sac slightly larger than those ventrolaterally.</paragraph>
<paragraph id="B67B36F58040FFAFFF6E75AFD8ACF816" blockId="17.[151,1437,152,2031]" pageId="17" pageNumber="322">
<emphasis id="84B0EAE78040FFAFFF6E75AFDDDCF90A" box="[199,440,1753,1779]" italics="true" pageId="17" pageNumber="322">Color in preservative</emphasis>
: Dorsal ground color pale gray with white spots laterally and dorsolaterally, largely arrayed into rows (
<figureCitation id="2EFF2A708040FFAFFEF0758BDDFBF8E1" box="[345,415,1789,1816]" captionStart="FIGURE 6" captionStartId="18.[151,250,1894,1919]" captionTargetBox="[283,1296,434,1735]" captionTargetId="figure@18.[273,1313,378,1869]" captionTargetPageId="18" captionText="FIGURE 6. Dorsal view of holotype of Lepidodactylus dialeukos sp. nov., BMNH 1974.3028." httpUri="https://zenodo.org/record/3363257/files/figure.png" pageId="17" pageNumber="322">Fig. 6</figureCitation>
). Narrow (13 granules wide) white dorsal crossbands before and after forearm insertion; another vaguely suggested on back of neck. Narrow white postocular stripe, another running behind ear opening, another short white line from corner of jaw to throat. Labials white spotted with black. Narrow white crossbands on forearm, wrist, shank, ankle, and each digit. Narrow white bands on tail bordered anteriorly with one or two dark-brown spots, first band behind legs connecting to short white vertebral stripe that extends to above hindlimb insertion, where it is margined anteriorly with a short, broken black chevron. White spots scattered on thighs. Venter pale gray, heavily stippled with black punctations, denser laterally. Iris silver veined with black.
</paragraph>
<paragraph id="B67B36F58043FFACFF6E73ECD913FF2E" blockId="18.[151,1436,152,359]" pageId="18" pageNumber="323">
<emphasis id="84B0EAE78043FFACFF6E73ECDDACFF4A" box="[199,456,153,179]" italics="true" pageId="18" pageNumber="323">Measurements (in mm)</emphasis>
. SVL = 46.5, TrL = 16.0, TW = 3.7, FA = 5.2, CS = 5.3, HL = 8.8, HW = 7.5, Ear = 0.7, EE = 3.9, EY = 2.8, SN = 5.3, EN = 4.0, IN = 2.3, T4L = 5.5, T4W = 2.0, T4 scansor L = 2.9, T3T4webL = 1.1.
</paragraph>
<paragraph id="B67B36F58043FFACFF6E7397DFABFEE6" blockId="18.[151,1436,152,359]" pageId="18" pageNumber="323">
<emphasis id="84B0EAE78043FFACFF6E7397DD5BFF02" box="[199,319,225,251]" italics="true" pageId="18" pageNumber="323">Etymology</emphasis>
. The name is a masculine possessive Greek adjective meaning
<emphasis id="84B0EAE78043FFACFBBB7397D880FF02" box="[1042,1252,225,251]" italics="true" pageId="18" pageNumber="323">marked with white</emphasis>
. The name is in reference to the numerous narrow white markings that typify this species.
</paragraph>
<paragraph id="B67B36F58043FFACFF6E725CDE8AFE9E" blockId="18.[151,1436,152,359]" pageId="18" pageNumber="323">
<emphasis id="84B0EAE78043FFACFF6E725CDD6BFEBA" box="[199,271,298,323]" italics="true" pageId="18" pageNumber="323">Range</emphasis>
. Known only from the
<typeStatus id="697F88578043FFACFDB1725FDE2CFEBA" box="[536,584,297,323]" pageId="18" pageNumber="323">type</typeStatus>
locality, but likely ranges widely across Yapen Island (
<figureCitation id="2EFF2A708043FFACFB1F725FD89FFEBA" box="[1206,1275,297,323]" captionStart="FIGURE 2" captionStartId="7.[151,248,874,899]" captionTargetBox="[162,1379,249,813]" captionTargetId="figure@7.[162,1379,249,813]" captionTargetPageId="7" captionText="FIGURE 2. Map of New Guinea and surrounding islands showing type localities for Lepidodactylus aignanus sp. nov. (diamond), L. dialeukos sp. nov. (triangle), L. kwasnickae sp. nov. (square), L. mitchelli sp. nov. (circle), and L. zweifeli sp. nov. (star)." httpUri="https://zenodo.org/record/3363249/files/figure.png" pageId="18" pageNumber="323">Fig. 2</figureCitation>
), although the species has apparently not been collected in 80 years.
</paragraph>
<caption id="E2BB667D8043FFACFF3E7410D838F886" box="[151,1116,1894,1919]" httpUri="https://zenodo.org/record/3363257/files/figure.png" pageId="18" pageNumber="323" startId="18.[151,250,1894,1919]" targetBox="[283,1296,434,1735]" targetPageId="18">
<paragraph id="B67B36F58043FFACFF3E7410D838F886" blockId="18.[151,1116,1894,1919]" box="[151,1116,1894,1919]" pageId="18" pageNumber="323">
<emphasis id="84B0EAE78043FFACFF3E7410DD77F886" bold="true" box="[151,275,1894,1919]" pageId="18" pageNumber="323">FIGURE 6.</emphasis>
Dorsal view of holotype of
<taxonomicName id="71C44D768043FFACFD987411DF57F886" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[561,819,1895,1919]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="323" phylum="Chordata" rank="species" species="dialeukos" status="sp. nov.">
<emphasis id="84B0EAE78043FFACFD987411DF57F886" box="[561,819,1895,1919]" italics="true" pageId="18" pageNumber="323">Lepidodactylus dialeukos</emphasis>
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<emphasis id="84B0EAE78043FFACFC907411DFEEF886" bold="true" box="[825,906,1895,1919]" pageId="18" pageNumber="323">sp. nov.</emphasis>
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, BMNH 1974.3028.
</paragraph>
</caption>
<paragraph id="B67B36F58042FFADFF6E73EFD953FF4A" blockId="19.[151,1437,153,395]" box="[199,1335,153,179]" pageId="19" pageNumber="324">
<emphasis id="84B0EAE78042FFADFF6E73EFDD47FF4A" box="[199,291,153,179]" italics="true" pageId="19" pageNumber="324">Ecology</emphasis>
. The elevation of the sole specimen suggests that it is an inhabitant of interior forested areas.
</paragraph>
<paragraph id="B67B36F58042FFADFF6E73CBD93CFE72" blockId="19.[151,1437,153,395]" pageId="19" pageNumber="324">
<emphasis id="84B0EAE78042FFADFF6E73CBDD4DFF2E" box="[199,297,189,215]" italics="true" pageId="19" pageNumber="324">Remarks</emphasis>
. In recognizing their phenetic Group II
<bibRefCitation id="D2554B048042FFADFD4473CBD868FF2E" author="Brown, W. C. &amp; Parker, F." box="[749,1036,189,215]" pageId="19" pageNumber="324" pagination="253 - 265" refId="ref18821" refString="Brown, W. C. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Sciences, 41, 253 - 265." type="journal article" year="1977">Brown and Parker (1977)</bibRefCitation>
defined it as comprising those species having 14 divided subterminal lamellae. Above I noted that
<taxonomicName id="71C44D768042FFADFC237394D870FF02" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[906,1044,225,251]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="324" phylum="Chordata" rank="species" species="dialeukos" status="sp. nov.">
<emphasis id="84B0EAE78042FFADFC237394D870FF02" box="[906,1044,225,251]" italics="true" pageId="19" pageNumber="324">L. dialeukos</emphasis>
</taxonomicName>
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<emphasis id="84B0EAE78042FFADFBB77397D81DFF02" bold="true" box="[1054,1145,225,251]" pageId="19" pageNumber="324">sp. nov.</emphasis>
</taxonomicNameLabel>
is unique, so far as I can determine, in having consistently only a single divided subterminal scansor. The observations of single divided subterminal scansors made by Brown and Parker in other species represent only occasional outliers on single digits in species for which the large majority of digits are characterized by 24 divided lamellae. Hence, the consistent condition of only a single divided subterminal scansor across all digits seems unique to
<taxonomicName id="71C44D768042FFADFBC77204D893FE72" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1134,1271,369,395]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="324" phylum="Chordata" rank="species" species="dialeukos" status="sp. nov.">
<emphasis id="84B0EAE78042FFADFBC77204D893FE72" box="[1134,1271,369,395]" italics="true" pageId="19" pageNumber="324">L. dialeukos</emphasis>
</taxonomicName>
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<emphasis id="84B0EAE78042FFADFB577207D93CFE72" bold="true" box="[1278,1368,369,395]" pageId="19" pageNumber="324">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
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<emphasis id="84B0EAE78053FFBCFF3E7443DDA1F8B6" bold="true" box="[151,453,1845,1872]" italics="true" pageId="2" pageNumber="307">Lepidodactylus aignanus</emphasis>
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<paragraph id="B67B36F58053FFBCFF3E74C3D9FEF80A" blockId="2.[151,1436,1973,2035]" pageId="2" pageNumber="307">
<materialsCitation id="06AC3CA88053FFBCFF3E74C3D9FEF80A" ID-GBIF-Occurrence="2313474471" collectingDate="2003-01-13" collectionCode="BPBM" collectorName="F. Kraus &amp; Liak" country="Papua New Guinea" latitude="-10.6594" location="Misima Island" longLatPrecision="7" longitude="152.6941" pageId="2" pageNumber="307" specimenCode="BPBM 17229" specimenCount="1" stateProvince="Milne Bay" typeStatus="holotype">
<typeStatus id="697F88578053FFBCFF3E74C3DC9AF836" box="[151,254,1973,1999]" pageId="2" pageNumber="307" type="holotype">
<emphasis id="84B0EAE78053FFBCFF3E74C3DC9AF836" box="[151,254,1973,1999]" italics="true" pageId="2" pageNumber="307">Holotype</emphasis>
</typeStatus>
.
<specimenCode id="E6629E8E8053FFBCFEA674C3DDC8F836" box="[271,428,1973,1999]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="307" type="Museum">BPBM 17229</specimenCode>
(field tag FK 6989), mature female, obtained by
<collectorName id="1B3153238053FFBCFC4474C3D82AF836" box="[1005,1102,1973,1999]" pageId="2" pageNumber="307">F. Kraus</collectorName>
from local collector at
<collectorName id="1B3153238053FFBCFACB74C3D9FCF836" box="[1378,1432,1973,1999]" pageId="2" pageNumber="307">Liak</collectorName>
,
<geoCoordinate id="D3F050328053FFBCFF3E74AEDD74F80A" box="[151,272,2008,2035]" degrees="10.6594" direction="south" orientation="latitude" pageId="2" pageNumber="307" precision="5" value="-10.6594">10.6594°S</geoCoordinate>
,
<geoCoordinate id="D3F050328053FFBCFEB274AEDDC7F80A" box="[283,419,2008,2035]" degrees="152.6941" direction="east" orientation="longitude" pageId="2" pageNumber="307" precision="5" value="152.6941">152.6941°E</geoCoordinate>
, near sea level,
<location id="B31B602E8053FFBCFDFC74AFDE9CF80A" LSID="urn:lsid:plazi:treatment:3E6D87E38053FFBAFF3E7443DDAFFCEE:B31B602E8053FFBCFDFC74AFDE9CF80A" box="[597,760,2009,2035]" country="Papua New Guinea" latitude="-10.6594" longLatPrecision="7" longitude="152.6941" name="Misima Island" pageId="2" pageNumber="307" stateProvince="Milne Bay">Misima Island</location>
,
<collectingRegion id="7400F8178053FFBCFCAD74AFDF8CF80A" box="[772,1000,2009,2035]" country="Papua New Guinea" name="Milne Bay" pageId="2" pageNumber="307">Milne Bay Province</collectingRegion>
,
<collectingCountry id="CED376658053FFBCFC5A74AFD8AAF80A" box="[1011,1230,2009,2035]" name="Papua New Guinea" pageId="2" pageNumber="307">Papua New Guinea</collectingCountry>
,
<collectingDate id="D23EE9DD8053FFBCFB7074AFD9F2F80A" box="[1241,1430,2009,2035]" pageId="2" pageNumber="307" value="2003-01-13">
<date id="C27A10358053FFBCFB7074AFD9F2F80A" box="[1241,1430,2009,2035]" pageId="2" pageNumber="307" value="2003-01-13">13 January 2003</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="B67B36F58052FFBDFF6E73ECDFB3FEBA" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="308">
<emphasis id="84B0EAE78052FFBDFF6E73ECDD5DFF4A" box="[199,313,154,179]" italics="true" pageId="3" pageNumber="308">Diagnosis</emphasis>
. A small (adult SVL
<quantity id="713C9B108052FFBDFD8373EEDEF4FF4A" box="[554,656,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="3" pageNumber="308" unit="mm" value="37.5">37.5 mm</quantity>
) species of
<taxonomicName id="71C44D768052FFBDFCB173EFDFA7FF4A" authorityName="Kraus" authorityYear="2019" box="[792,963,153,179]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="genus">
<emphasis id="84B0EAE78052FFBDFCB173EFDFA7FF4A" box="[792,963,153,179]" italics="true" pageId="3" pageNumber="308">Lepidodactylus</emphasis>
</taxonomicName>
having a subcylindrical tail without a lateral fringe of enlarged scales; all subterminal scansors entire; 17 enlarged scales of pore-bearing series limited to precloacal region; 1112 T4 lamellae, 79 T1 lamellae; fairly wide toes (T4W/T4L = 0.33); toes approximately onequarter webbed (T3T4webL/T4L = 0.26); dorsum with several irregular dark-brown saddles, and many pale lateral and dorsolateral round spots, especially concentrated from forearm to eye.
</paragraph>
<paragraph id="B67B36F58052FFBDFF6E723BDE2AFD36" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="308">
<emphasis id="84B0EAE78052FFBDFF6E723BDE5DFE9E" box="[199,569,333,359]" italics="true" pageId="3" pageNumber="308">Comparisons with other species</emphasis>
. The subcylindrical tail without a lateral fringe of enlarged scales and the series of undivided scansors under all toes place
<taxonomicName id="71C44D768052FFBDFD6D7207DFBAFE72" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[708,990,369,395]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78052FFBDFD6D7207DFBAFE72" box="[708,990,369,395]" italics="true" pageId="3" pageNumber="308">Lepidodactylus aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8052FFBDFC4F7207D825FE72" box="[998,1089,369,395]" pageId="3" pageNumber="308" rank="species">
<emphasis id="84B0EAE78052FFBDFC4F7207D825FE72" bold="true" box="[998,1089,369,395]" pageId="3" pageNumber="308">sp. nov.</emphasis>
</taxonomicNameLabel>
in
<bibRefCitation id="D2554B048052FFBDFBCE7207D9F8FE72" author="Brown, W. C. &amp; Parker, F." box="[1127,1436,369,395]" pageId="3" pageNumber="308" pagination="253 - 265" refId="ref18821" refString="Brown, W. C. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Sciences, 41, 253 - 265." type="journal article" year="1977">Brown and Parkers (1977)</bibRefCitation>
Group I.
<taxonomicName id="71C44D768052FFBDFF5672E3DE7DFE56" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[255,537,405,431]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78052FFBDFF5672E3DE7DFE56" box="[255,537,405,431]" italics="true" pageId="3" pageNumber="308">Lepidodactylus aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8052FFBDFD8972E3DE1FFE56" box="[544,635,405,431]" pageId="3" pageNumber="308" rank="species">
<emphasis id="84B0EAE78052FFBDFD8972E3DE1FFE56" bold="true" box="[544,635,405,431]" pageId="3" pageNumber="308">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from other Melanesian members of this group as follows: from
<taxonomicName id="71C44D768052FFBDFF3E72CCDE0EFE2A" authority="Brown, McCoy &amp; Rodda" authorityName="Brown, McCoy &amp; Rodda" box="[151,618,441,467]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="84B0EAE78052FFBDFF3E72CCDD27FE2A" box="[151,323,441,467]" italics="true" pageId="3" pageNumber="308">L. flaviocularis</emphasis>
Brown, McCoy &amp; Rodda
</taxonomicName>
,
<taxonomicName id="71C44D768052FFBDFDDF72CCDFCBFE2A" authority="Brown &amp; Parker" authorityName="Brown &amp; Parker" authorityYear="1977" box="[630,943,441,467]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="magnus">
<emphasis id="84B0EAE78052FFBDFDDF72CCDE8FFE2A" box="[630,747,442,467]" italics="true" pageId="3" pageNumber="308">L. magnus</emphasis>
Brown &amp; Parker
</taxonomicName>
,
<taxonomicName id="71C44D768052FFBDFC1372CCD88EFE2A" authority="Brown &amp; Parker" authorityName="Brown &amp; Parker" authorityYear="1977" box="[954,1258,441,467]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="mutahi">
<emphasis id="84B0EAE78052FFBDFC1372CCD842FE2A" box="[954,1062,441,467]" italics="true" pageId="3" pageNumber="308">L. mutahi</emphasis>
Brown &amp; Parker
</taxonomicName>
, and
<taxonomicName id="71C44D768052FFBDFA8C72CCD9F9FE2A" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[1317,1437,441,467]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="84B0EAE78052FFBDFA8C72CCD9F9FE2A" box="[1317,1437,441,467]" italics="true" pageId="3" pageNumber="308">L. pumilus</emphasis>
</taxonomicName>
in having 17 enlarged precloacal scales (
<emphasis id="84B0EAE78052FFBDFDF672A8DEC3FE0E" box="[607,679,478,503]" italics="true" pageId="3" pageNumber="308">versus</emphasis>
2650 enlarged precloacal/femoral scales in the other species) limited to the precloacal region (extending far onto the thighs in the other species); from
<taxonomicName id="71C44D768052FFBDFBCC7174D9F8FDE2" authority="Pernetta &amp; Black" authorityName="Pernetta &amp; Black" authorityYear="1983" box="[1125,1436,513,539]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="browni">
<emphasis id="84B0EAE78052FFBDFBCC7174D8B5FDE2" box="[1125,1233,513,539]" italics="true" pageId="3" pageNumber="308">L. browni</emphasis>
Pernetta &amp; Black
</taxonomicName>
and
<taxonomicName id="71C44D768052FFBDFF6E7150DE74FDC6" authority="Brown &amp; Parker" authorityName="Brown &amp; Parker" authorityYear="1977" box="[199,528,549,575]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="84B0EAE78052FFBDFF6E7150DD2BFDC6" box="[199,335,549,575]" italics="true" pageId="3" pageNumber="308">L. orientalis</emphasis>
Brown &amp; Parker
</taxonomicName>
in having 17 enlarged precloacal scales (
<emphasis id="84B0EAE78052FFBDFC777150D842FDC6" box="[990,1062,550,575]" italics="true" pageId="3" pageNumber="308">versus</emphasis>
<quantity id="713C9B108052FFBDFB847153D8F4FDC6" box="[1069,1168,549,575]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.858" metricValueMin="4.826" pageId="3" pageNumber="308" unit="in" value="23.0" valueMax="27.0" valueMin="19.0">1927 in</quantity>
<taxonomicName id="71C44D768052FFBDFB3E7150D97BFDC6" authorityName="Brown &amp; Parker" authorityYear="1977" box="[1175,1311,549,575]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="84B0EAE78052FFBDFB3E7150D97BFDC6" box="[1175,1311,549,575]" italics="true" pageId="3" pageNumber="308">L. orientalis</emphasis>
</taxonomicName>
and
<quantity id="713C9B108052FFBDFAFF7153DCC9FD9A" metricMagnitude="-1" metricUnit="m" metricValue="7.747" metricValueMax="9.906" metricValueMin="5.588" pageId="3" pageNumber="308" unit="in" value="30.5" valueMax="39.0" valueMin="22.0">2239 in</quantity>
<taxonomicName id="71C44D768052FFBDFF1C713CDD45FD9A" authorityName="Pernetta &amp; Black" authorityYear="1983" box="[181,289,585,611]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="browni">
<emphasis id="84B0EAE78052FFBDFF1C713CDD45FD9A" box="[181,289,585,611]" italics="true" pageId="3" pageNumber="308">L. browni</emphasis>
</taxonomicName>
), toes with more webbing (T3T4webL/T4L = 0.26
<emphasis id="84B0EAE78052FFBDFCC1713CDFD4FD9A" box="[872,944,586,611]" italics="true" pageId="3" pageNumber="308">versus</emphasis>
<quantity id="713C9B108052FFBDFC1E713FD820FD9A" box="[951,1092,585,611]" metricMagnitude="-3" metricUnit="m" metricValue="3.429" metricValueMax="4.064" metricValueMin="2.7939999999999996" pageId="3" pageNumber="308" unit="in" value="0.135" valueMax="0.16" valueMin="0.11">0.110.16 in</quantity>
<taxonomicName id="71C44D768052FFBDFBE5713CD8DCFD9A" authorityName="Pernetta &amp; Black" authorityYear="1983" box="[1100,1208,585,611]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="browni">
<emphasis id="84B0EAE78052FFBDFBE5713CD8DCFD9A" box="[1100,1208,585,611]" italics="true" pageId="3" pageNumber="308">L. browni</emphasis>
</taxonomicName>
and
<quantity id="713C9B108052FFBDFB59713FD91AFD9A" box="[1264,1406,585,611]" metricMagnitude="-3" metricUnit="m" metricValue="3.048" metricValueMax="3.556" metricValueMin="2.54" pageId="3" pageNumber="308" unit="in" value="0.12" valueMax="0.14" valueMin="0.1">0.100.14 in</quantity>
<taxonomicName id="71C44D768052FFBDFA2F713CDD66FD7E" authorityName="Brown &amp; Parker" authorityYear="1977" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="84B0EAE78052FFBDFA2F713CDD66FD7E" italics="true" pageId="3" pageNumber="308">L. orientalis</emphasis>
</taxonomicName>
,
<figureCitation id="2EFF2A708052FFBDFEB8711BDD03FD7E" box="[273,359,621,647]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="3" pageNumber="308">Fig. 1C</figureCitation>
), and dorsum with many pale lateral and dorsolateral round spots (
<figureCitation id="2EFF2A708052FFBDFBF4711BD8D2FD7E" box="[1117,1206,621,647]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="3" pageNumber="308">Fig. 1A</figureCitation>
), especially concentrated from forearm to eye (
<emphasis id="84B0EAE78052FFBDFE6271E4DE77FD52" box="[459,531,658,683]" italics="true" pageId="3" pageNumber="308">versus</emphasis>
absent in
<taxonomicName id="71C44D768052FFBDFD2B71E4DE89FD52" authorityName="Pernetta &amp; Black" authorityYear="1983" box="[642,749,657,683]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="browni">
<emphasis id="84B0EAE78052FFBDFD2B71E4DE89FD52" box="[642,749,657,683]" italics="true" pageId="3" pageNumber="308">L. browni</emphasis>
</taxonomicName>
and
<taxonomicName id="71C44D768052FFBDFC8B71E4DFCEFD52" authorityName="Brown &amp; Parker" authorityYear="1977" box="[802,938,657,683]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="84B0EAE78052FFBDFC8B71E4DFCEFD52" box="[802,938,657,683]" italics="true" pageId="3" pageNumber="308">L. orientalis</emphasis>
</taxonomicName>
, or with few vague dorsolateral spots behind forearm in one specimen of
<taxonomicName id="71C44D768052FFBDFE7B71C0DE5AFD36" authorityName="Pernetta &amp; Black" authorityYear="1983" box="[466,574,693,719]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="308" phylum="Chordata" rank="species" species="browni">
<emphasis id="84B0EAE78052FFBDFE7B71C0DE5AFD36" box="[466,574,693,719]" italics="true" pageId="3" pageNumber="308">L. browni</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="B67B36F58052FFBDFF6E71ACDF67FB3E" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="308">
<emphasis id="84B0EAE78052FFBDFF6E71ACDDA9FD0A" box="[199,461,729,755]" italics="true" pageId="3" pageNumber="308">
Description of
<typeStatus id="697F88578052FFBDFEC471AFDDA9FD0A" box="[365,461,729,755]" pageId="3" pageNumber="308" type="holotype">holotype</typeStatus>
</emphasis>
. A mature female, with right-lateral incision. Animal small (SVL =
<quantity id="713C9B108052FFBDFB1171AED978FD0A" box="[1208,1308,728,755]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="3" pageNumber="308" unit="mm" value="37.5">37.5 mm</quantity>
, TrL =
<quantity id="713C9B108052FFBDFAC271AFDCA3FCEE" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="3" pageNumber="308" unit="mm" value="18.3">18.3 mm</quantity>
). Head relatively long (HL/SVL = 0.23) and wide (HW/HL = 0.80), distinct from neck. Loreal region slightly inflated; no distinct canthus rostralis. Interorbital region and top of snout concave. Snout tapered and rounded at tip, relatively long (SN/HL = 0.41), longer than eye diameter (SN/EY = 1.6). Eye relatively large (EY/HL = 0.26, EY/EN = 0.76); pupil vertical, constricted into series of four lobes; supraciliaries subequal in size to adjacent granules. Ear opening small (Ear/HL = 0.070), somewhat compressed, oriented diagonally from posterodorsal to anteroventral; distance between ear and eye larger than eye diameter (EE/EY = 1.4). Rostral wider (
<quantity id="713C9B108052FFBDFB6E70C7D945FC32" box="[1223,1313,945,971]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="308" unit="mm" value="1.8">1.8 mm</quantity>
) than high (
<quantity id="713C9B108052FFBDFF3670A3DC9FFC16" box="[159,251,981,1007]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="308" unit="mm" value="0.8">0.8 mm</quantity>
), highest just medial to nares, lower between these points; length
<quantity id="713C9B108052FFBDFC7770A3D85CFC16" box="[990,1080,981,1007]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="308" unit="mm" value="0.4">0.4 mm</quantity>
. Supranasals separated by three internasals. Rostral in contact with first supralabials, two supranasals, and three internasals. External nares circular; each bordered by rostral, two supranasals, first supralabial, and one postnasal. Mental shallowly triangular, slightly longer (
<quantity id="713C9B108052FFBDFF477736DD32FBA2" box="[238,342,1088,1115]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="308" unit="mm" value="0.65">0.65 mm</quantity>
) than wide (
<quantity id="713C9B108052FFBDFE4A7737DE2AFBA2" box="[483,590,1089,1115]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="308" unit="mm" value="0.6">0.60 mm</quantity>
). Mental bordered posteriorly by two enlarged postmentals, these bordered posteriorly by two rows of slightly enlarged scales prior to granular chin scales. Infralabials bordered by enlarged scales that gradually decrease in size posteriorly. Supralabials to mid-orbital position nine on each side; to angle of jaw 13 on right, 12 on left. Infralabials 11 on each side.
</paragraph>
<paragraph id="B67B36F58052FFBDFF6E77A7D940FACA" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="308">Body of modest habitus (TrL/SVL = 0.49), slightly depressed. Dorsal scales on head, body, and limbs tiny, juxtaposed granules; tubercles absent. Ventral scales flat and smooth; those on throat smaller and juxtaposed; those on abdomen larger and subimbricate, gradually decreasing in size laterally and on chin to become granular.</paragraph>
<paragraph id="B67B36F58052FFBDFF6E764BDDADFA3A" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="308">Enlarged precloacal scales 17, in a single short series; no enlarged scales on thighs. Enlarged scales form a pubic patch between the precloacal series and vent; no row of tiny scales between the precloacal series and either side of the pubic patch; ten scales in a row between apex of enlarged precloacal series and vent. Scales on palms and soles rounded, smooth.</paragraph>
<paragraph id="B67B36F58052FFBDFF6E76BBDE91F91A" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="308">
Fore- and hindlimbs relatively small (FA/SVL = 0.083, CS/SVL = 0.12). Digits well-developed, moderately dilated throughout their length (T4W/T4L = 0.33), all but first fingers and toes with recurved claws; clawed phalanges laterally compressed, free above and extending slightly beyond terminal scansors. Subdigital scansors narrow and smooth, all undivided except for three terminal scansors divided on left third toe; lamellae extend for more than half length of each toe (T4 scansor L/T4L = 0.62,
<figureCitation id="2EFF2A708052FFBDFD3D752BDE8EF98E" box="[660,746,1629,1655]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="3" pageNumber="308">Fig. 1C</figureCitation>
). Lamellae of manus
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9 on right,
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8 on left; of pes
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10 on right, 9
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on left. Relative lengths of digits on manus and pes I &lt;II &lt;V &lt;III &lt;IV. Webbing present between all digits except first and second on both manus and pes; toes approximately one-quarter webbed (T3T4webL/T4L = 0.26,
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)
</paragraph>
<paragraph id="B67B36F58052FFBDFF6E759BD8F9F8B6" blockId="3.[151,1437,152,2015]" pageId="3" pageNumber="308">
Tail missing except for
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behind vent, relatively wide (TW/SVL = 0.10), with slight constriction approximately
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behind vent; lateral margins without skin flanges or spines. Scales of tail remnant small, flat, smooth, subimbricate ventrally; granular dorsally; no enlarged postcloacal spurs on sides of tailbase.
</paragraph>
<paragraph id="B67B36F58052FFBAFF6E742FDD21FF02" blockId="3.[151,1437,152,2015]" lastBlockId="4.[151,1436,153,791]" lastPageId="4" lastPageNumber="309" pageId="3" pageNumber="308">
<emphasis id="84B0EAE78052FFBDFF6E742FDDDDF88A" box="[199,441,1881,1907]" italics="true" pageId="3" pageNumber="308">Color in preservative</emphasis>
: Dorsal ground color medium brown with small, irregular, vague, darker-brown markings scattered dorsally and laterally, these coalescing into a short scapular line on each side (
<figureCitation id="2EFF2A708052FFBDFB08740BD89EF86E" box="[1185,1274,1917,1943]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="3" pageNumber="308">Fig. 1A</figureCitation>
). A lateral and dorsolateral row of pale-straw spots on each side, extending from in front of forearm insertion to just before groin (lateral row) and to base of tail (dorsolateral row). Other, more vaguely defined pale spots scattered about behind head. Venter pale straw with scattered brown punctations concentrated toward sides. Under limbs, palms, and soles same but with all scales having brown punctations. Labials also pale straw with brown punctations. Iris dark brown, pupil pale gray.
</paragraph>
<paragraph id="B67B36F58055FFBAFF6E7270D910FEBA" blockId="4.[151,1436,153,791]" pageId="4" pageNumber="309">
<emphasis id="84B0EAE78055FFBAFF6E7270DDACFEE6" box="[199,456,261,287]" italics="true" pageId="4" pageNumber="309">Measurements (in mm)</emphasis>
. SVL = 37.5, TrL = 18.3, TW = 3.8, FA = 3.1, CS = 4.6, HL = 8.6, HW = 6.9, Ear = 0.6, EE = 3.0, EY = 2.2, SN = 3.5, EN = 2.9, IN = 1.8, T4L = 3.9, T4W = 1.3, T4lamellaeL = 2.4, T3T4webL = 1.0.
</paragraph>
<paragraph id="B67B36F58055FFBAFF6E723BDDB3FE72" blockId="4.[151,1436,153,791]" pageId="4" pageNumber="309">
<emphasis id="84B0EAE78055FFBAFF6E723BDD5BFE9E" box="[199,319,333,359]" italics="true" pageId="4" pageNumber="309">Etymology</emphasis>
. The name is a masculine adjective referring to the
<typeStatus id="697F88578055FFBAFC3F723BDFA2FE9E" box="[918,966,333,359]" pageId="4" pageNumber="309">type</typeStatus>
locality of Misima Island, whose former name was St. Aignan Island.
</paragraph>
<paragraph id="B67B36F58055FFBAFF6E72E0DC96FE2A" blockId="4.[151,1436,153,791]" pageId="4" pageNumber="309">
<emphasis id="84B0EAE78055FFBAFF6E72E0DD6BFE56" box="[199,271,406,431]" italics="true" pageId="4" pageNumber="309">Range</emphasis>
. Known only from the
<typeStatus id="697F88578055FFBAFDB572E3DE28FE56" box="[540,588,405,431]" pageId="4" pageNumber="309">type</typeStatus>
locality, but no doubt ranges across the lower elevations of Misima Island (
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).
</paragraph>
<paragraph id="B67B36F58055FFBAFF6E72ABDE71FD7E" blockId="4.[151,1436,153,791]" pageId="4" pageNumber="309">
<emphasis id="84B0EAE78055FFBAFF6E72ABDD47FE0E" box="[199,291,477,503]" italics="true" pageId="4" pageNumber="309">Ecology</emphasis>
. The area around Liak is mixed gardens with a fringe of coastal trees. Common trees in the area include
<taxonomicName id="71C44D768055FFBAFF3E7177DD39FDE2" box="[151,349,513,539]" class="Magnoliopsida" family="Moraceae" genus="Artocarpus" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="309" phylum="Tracheophyta" rank="species" species="altilis">
<emphasis id="84B0EAE78055FFBAFF3E7177DD39FDE2" box="[151,349,513,539]" italics="true" pageId="4" pageNumber="309">Artocarpus altilis</emphasis>
</taxonomicName>
,
<taxonomicName id="71C44D768055FFBAFEC57177DEE7FDE2" box="[364,643,513,539]" class="Magnoliopsida" family="Calophyllaceae" genus="Calophyllum" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="309" phylum="Tracheophyta" rank="species" species="inophyllum">
<emphasis id="84B0EAE78055FFBAFEC57177DEE7FDE2" box="[364,643,513,539]" italics="true" pageId="4" pageNumber="309">Calophyllum inophyllum</emphasis>
</taxonomicName>
,
<taxonomicName id="71C44D768055FFBAFD3B7177DF59FDE2" box="[658,829,513,539]" class="Liliopsida" family="Arecaceae" genus="Cocos" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="309" phylum="Tracheophyta" rank="species" species="nucifera">
<emphasis id="84B0EAE78055FFBAFD3B7177DF59FDE2" box="[658,829,513,539]" italics="true" pageId="4" pageNumber="309">Cocos nucifera</emphasis>
</taxonomicName>
, and
<taxonomicName id="71C44D768055FFBAFCD47174DFDFFDE2" box="[893,955,514,539]" class="Liliopsida" family="Musaceae" genus="Musa" kingdom="Plantae" order="Zingiberales" pageId="4" pageNumber="309" phylum="Tracheophyta" rank="genus">
<emphasis id="84B0EAE78055FFBAFCD47174DFDFFDE2" box="[893,955,514,539]" italics="true" pageId="4" pageNumber="309">Musa</emphasis>
</taxonomicName>
spp. It is uncertain how close to shore the specimen was collected, but
<taxonomicName id="71C44D768055FFBAFE757150DE33FDC6" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" box="[476,599,549,575]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="84B0EAE78055FFBAFE757150DE33FDC6" box="[476,599,549,575]" italics="true" pageId="4" pageNumber="309">L. lugubris</emphasis>
</taxonomicName>
was obtained at the same location and often dominates shoreline communities, so
<taxonomicName id="71C44D768055FFBAFEAD713CDDEEFD9A" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[260,394,586,611]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFEAD713CDDEEFD9A" box="[260,394,586,611]" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFE38713FDD88FD9A" box="[401,492,585,611]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFE38713FDD88FD9A" bold="true" box="[401,492,585,611]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
may occur somewhat inland from that. It remains unknown as well what the upper elevation of this species might be.
</paragraph>
<paragraph id="B67B36F58055FFBAFF6E71E7DDAFFCEE" blockId="4.[151,1436,153,791]" pageId="4" pageNumber="309">
<emphasis id="84B0EAE78055FFBAFF6E71E7DD4DFD52" box="[199,297,657,683]" italics="true" pageId="4" pageNumber="309">Remarks</emphasis>
. This species was included in the phylogeny of
<bibRefCitation id="D2554B048055FFBAFCEE71E7D849FD52" author="Oliver, P. M. &amp; Brown, R. &amp; Kraus, F. &amp; Rittmeyer, E. &amp; Travers, S. L. &amp; Siler, C. D." box="[839,1069,657,683]" pageId="4" pageNumber="309" pagination="20171760" refId="ref19729" refString="Oliver, P. M., Brown, R., Kraus, F., Rittmeyer, E., Travers, S. L. &amp; Siler, C. D. (2018 a) Lizards of the lost arcs: mid-Cenozoic diversification, persistence and ecological marginalization around the western Pacific. Proceedings of the Royal Society of London B, 285 (1871), 20171760. https: // doi. org / 10.1098 / rspb. 2017.1760" type="journal article" year="2018">
Oliver
<emphasis id="84B0EAE78055FFBAFC3F71E4DFABFD52" box="[918,975,657,683]" italics="true" pageId="4" pageNumber="309">et al.</emphasis>
(2018a)
</bibRefCitation>
as a member of the
<taxonomicName id="71C44D768055FFBAFABA71E4D9F8FD52" authorityName="Brown &amp; Parker" authorityYear="1977" box="[1299,1436,657,683]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="84B0EAE78055FFBAFABA71E4D9F8FD52" box="[1299,1436,657,683]" italics="true" pageId="4" pageNumber="309">L. orientalis</emphasis>
</taxonomicName>
species group. Its sister taxon in that tree was a population of
<taxonomicName id="71C44D768055FFBAFCFA71C3DF9AFD36" authorityName="Kraus" authorityYear="2019" box="[851,1022,693,719]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="84B0EAE78055FFBAFCFA71C3DF9AFD36" box="[851,1022,693,719]" italics="true" pageId="4" pageNumber="309">Lepidodactylus</emphasis>
</taxonomicName>
from nearby Sudest Island, which is known from only two specimens, has proven difficult to distinguish from
<taxonomicName id="71C44D768055FFBAFC7F71ACD83BFD0A" authorityName="Brown &amp; Parker" authorityYear="1977" box="[982,1119,729,755]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="84B0EAE78055FFBAFC7F71ACD83BFD0A" box="[982,1119,729,755]" italics="true" pageId="4" pageNumber="309">L. orientalis</emphasis>
</taxonomicName>
based on that small sample, and is not described herein.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

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<heading id="ED3381998055FFBAFF3E701FDE71FC7A" bold="true" box="[151,533,873,900]" fontSize="11" level="1" pageId="4" pageNumber="309" reason="1">
<emphasis id="84B0EAE78055FFBAFF3E701FDE71FC7A" bold="true" box="[151,533,873,900]" pageId="4" pageNumber="309">
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<emphasis id="84B0EAE78055FFBAFF3E701FDDD4FC7D" bold="true" box="[151,432,873,900]" italics="true" pageId="4" pageNumber="309">Lepidodactylus zweifeli</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="B67B36F58055FFBAFF3E70DCDFB0FC3D" blockId="4.[151,980,873,964]" box="[151,980,938,964]" pageId="4" pageNumber="309">
<uri id="C2553AF78055FFBAFF3E70DCDFB0FC3D" box="[151,980,938,964]" pageId="4" pageNumber="309">
urn:lsid:zoobank.org:act:
<uuid id="C2620C208055FFBAFE0E70DCDFB0FC3D" box="[423,980,938,964]" pageId="4" pageNumber="309">7722DE9A-EA83-44EF-96D4-C010B041DBA8</uuid>
</uri>
</paragraph>
<paragraph id="B67B36F58055FFBAFF3E709FD8F7FBDE" blockId="4.[151,1437,1000,2035]" pageId="4" pageNumber="309">
<materialsCitation id="06AC3CA88055FFBAFF3E709FD8F7FBDE" ID-GBIF-Occurrence="2313474472" collectingDate="1969-08-05" collectionCode="AMNH" collectorName="R. Zweifel &amp; Wanuma" country="Papua New Guinea" elevation="671" latitude="-4.9" location="Adelbert Mts." longLatPrecision="783" longitude="145.33" pageId="4" pageNumber="309" specimenCode="AMNH 105088" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="697F88578055FFBAFF3E709FDC9AFBFA" box="[151,254,1001,1027]" pageId="4" pageNumber="309" type="holotype">
<emphasis id="84B0EAE78055FFBAFF3E709FDC9AFBFA" box="[151,254,1001,1027]" italics="true" pageId="4" pageNumber="309">Holotype</emphasis>
</typeStatus>
.
<specimenCode id="E6629E8E8055FFBAFEA0709FDDDCFBFA" box="[265,440,1000,1027]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="309">AMNH 105088</specimenCode>
(field tag RZ 08430), mature male, collected by
<collectorName id="1B3153238055FFBAFC66709FD821FBFA" box="[975,1093,1001,1027]" pageId="4" pageNumber="309">R. Zweifel</collectorName>
at
<collectorName id="1B3153238055FFBAFBCD709FD8AEFBFA" box="[1124,1226,1001,1027]" pageId="4" pageNumber="309">Wanuma</collectorName>
,
<geoCoordinate id="D3F050328055FFBAFB7D709FD945FBFA" box="[1236,1313,1001,1027]" degrees="4.90" direction="south" orientation="latitude" pageId="4" pageNumber="309" precision="555" value="-4.9">4.90°S</geoCoordinate>
,
<geoCoordinate id="D3F050328055FFBAFA85709ED9F3FBFA" box="[1324,1431,1000,1027]" degrees="145.33" direction="east" orientation="longitude" pageId="4" pageNumber="309" precision="555" value="145.33">145.33°E</geoCoordinate>
,
<elevation id="3DE9D1C68055FFBAFF3E777BDD64FBDE" box="[151,256,1037,1063]" metricMagnitude="2" metricUnit="m" metricValue="6.7056" pageId="4" pageNumber="309" unit="ft" value="2200.0">
<quantity id="713C9B108055FFBAFF3E777BDD64FBDE" box="[151,256,1037,1063]" metricMagnitude="2" metricUnit="m" metricValue="6.7056" pageId="4" pageNumber="309" unit="ft" value="2200.0">2200 feet</quantity>
</elevation>
[=
<elevation id="3DE9D1C68055FFBAFE8E777BDD16FBDE" box="[295,370,1036,1063]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="309" unit="m" value="670.0">
<quantity id="713C9B108055FFBAFE8E777BDD16FBDE" box="[295,370,1036,1063]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="309" unit="m" value="670.0">670 m</quantity>
</elevation>
],
<location id="B31B602E8055FFBAFE2D777BDE46FBDE" LSID="urn:lsid:plazi:treatment:3E6D87E38055FFB9FF3E701FDFA2FF2E:B31B602E8055FFBAFE2D777BDE46FBDE" box="[388,546,1037,1063]" country="Papua New Guinea" latitude="-4.9" longLatPrecision="783" longitude="145.33" name="Adelbert Mts." pageId="4" pageNumber="309" stateProvince="Madang">Adelbert Mts.</location>
,
<collectingRegion id="7400F8178055FFBAFD84777BDE92FBDE" box="[557,758,1037,1063]" country="Papua New Guinea" name="Madang" pageId="4" pageNumber="309">Madang Province</collectingRegion>
,
<collectingCountry id="CED376658055FFBAFCAB777BDFB9FBDE" box="[770,989,1037,1063]" name="Papua New Guinea" pageId="4" pageNumber="309">Papua New Guinea</collectingCountry>
,
<collectingDate id="D23EE9DD8055FFBAFC41777AD8EAFBDE" box="[1000,1166,1036,1063]" pageId="4" pageNumber="309" value="1969-08-05">
<date id="C27A10358055FFBAFC41777AD8EAFBDE" box="[1000,1166,1036,1063]" pageId="4" pageNumber="309" value="1969-08-05">5 August 1969</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="B67B36F58055FFBAFF6E7744DF6DFBB2" blockId="4.[151,1437,1000,2035]" box="[199,777,1072,1099]" pageId="4" pageNumber="309">
<materialsCitation id="06AC3CA88055FFBAFF6E7744DF61FBB2" ID-GBIF-Occurrence="2313474475" box="[199,773,1072,1099]" collectionCode="AMNH" pageId="4" pageNumber="309" specimenCode="AMNH 105087" specimenCount="2" typeStatus="paratype">
<typeStatus id="697F88578055FFBAFF6E7744DD4AFBB2" box="[199,302,1074,1099]" pageId="4" pageNumber="309" type="paratype">
<emphasis id="84B0EAE78055FFBAFF6E7744DD4AFBB2" box="[199,302,1074,1099]" italics="true" pageId="4" pageNumber="309">Paratype</emphasis>
</typeStatus>
. Same data as
<typeStatus id="697F88578055FFBAFE7C7747DE53FBB2" box="[469,567,1073,1099]" pageId="4" pageNumber="309" type="holotype">holotype</typeStatus>
(
<specimenCode id="E6629E8E8055FFBAFDEC7747DE99FBB2" box="[581,765,1072,1099]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="309">AMNH 105087</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B67B36F58055FFBAFF6E7720D950FADA" blockId="4.[151,1437,1000,2035]" pageId="4" pageNumber="309">
<emphasis id="84B0EAE78055FFBAFF6E7720DD5DFB96" box="[199,313,1110,1135]" italics="true" pageId="4" pageNumber="309">Diagnosis</emphasis>
. A medium-sized (adult SVL
<quantity id="713C9B108055FFBAFD3A7722DF41FB96" box="[659,805,1108,1135]" metricMagnitude="-2" metricUnit="m" metricValue="4.275" metricValueMax="4.45" metricValueMin="4.1" pageId="4" pageNumber="309" unit="mm" value="42.75" valueMax="44.5" valueMin="41.0">4144.5 mm</quantity>
) species of
<taxonomicName id="71C44D768055FFBAFC187723D838FB96" authorityName="Kraus" authorityYear="2019" box="[945,1116,1109,1135]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="genus">
<emphasis id="84B0EAE78055FFBAFC187723D838FB96" box="[945,1116,1109,1135]" italics="true" pageId="4" pageNumber="309">Lepidodactylus</emphasis>
</taxonomicName>
having a subcylindrical tail without a lateral fringe of enlarged scales; all subterminal scansors entire; 40 enlarged precloacal/femoral scales in three series extending along entire femoral region, 38 precloacal/femoral pores in three series in the sole male, with the lateral series on each thigh separated by 23 scales from the precloacal series; 912 T4 lamellae, 78 T1 lamellae; fairly narrow toes (T4W/T4L = 0.2427) with basal webbing (T3T4webL/T4L = 0.100.13, T4T5webL/T4L = 0.080.12); narrow snout (EN/IN = 1.721.79); and dorsum with dark-brown markings but no pale spots.
</paragraph>
<paragraph id="B67B36F58055FFBAFF6E765BDE22F87E" blockId="4.[151,1437,1000,2035]" pageId="4" pageNumber="309">
<emphasis id="84B0EAE78055FFBAFF6E765BDE4FFABE" box="[199,555,1325,1351]" italics="true" pageId="4" pageNumber="309">Comparisons with other species</emphasis>
. The subcylindrical tail without a lateral fringe of enlarged scales and the series of undivided scansors under all toes place
<taxonomicName id="71C44D768055FFBAFDDC7627DF1AFA92" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[629,894,1361,1387]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="zweifeli" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFDDC7627DF1AFA92" box="[629,894,1361,1387]" italics="true" pageId="4" pageNumber="309">Lepidodactylus zweifeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFC2C7627DFBBFA92" box="[901,991,1361,1387]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFC2C7627DFBBFA92" bold="true" box="[901,991,1361,1387]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
in
<bibRefCitation id="D2554B048055FFBAFBAA7627D951FA92" author="Brown, W. C. &amp; Parker, F." box="[1027,1333,1361,1387]" pageId="4" pageNumber="309" pagination="253 - 265" refId="ref18821" refString="Brown, W. C. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Sciences, 41, 253 - 265." type="journal article" year="1977">Brown and Parkers (1977)</bibRefCitation>
Group I.
<taxonomicName id="71C44D768055FFBAFF3E7603DDC4FA76" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[151,416,1397,1423]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="zweifeli" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFF3E7603DDC4FA76" box="[151,416,1397,1423]" italics="true" pageId="4" pageNumber="309">Lepidodactylus zweifeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFE0E7603DE65FA76" box="[423,513,1397,1423]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFE0E7603DE65FA76" bold="true" box="[423,513,1397,1423]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from other Melanesian members of this group as follows: from
<taxonomicName id="71C44D768055FFBAFA807600D9F1FA76" authorityName="Pernetta &amp; Black" authorityYear="1983" box="[1321,1429,1397,1423]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="browni">
<emphasis id="84B0EAE78055FFBAFA807600D9F1FA76" box="[1321,1429,1397,1423]" italics="true" pageId="4" pageNumber="309">L. browni</emphasis>
</taxonomicName>
,
<taxonomicName id="71C44D768055FFBAFF3E76ECDD20FA4A" box="[151,324,1433,1459]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="flaviocularis">
<emphasis id="84B0EAE78055FFBAFF3E76ECDD20FA4A" box="[151,324,1433,1459]" italics="true" pageId="4" pageNumber="309">L. flaviocularis</emphasis>
</taxonomicName>
,
<taxonomicName id="71C44D768055FFBAFEFA76ECDDA4FA4A" authorityName="Brown &amp; Parker" authorityYear="1977" box="[339,448,1433,1459]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="mutahi">
<emphasis id="84B0EAE78055FFBAFEFA76ECDDA4FA4A" box="[339,448,1433,1459]" italics="true" pageId="4" pageNumber="309">L. mutahi</emphasis>
</taxonomicName>
, and
<taxonomicName id="71C44D768055FFBAFDA976ECDEEEFA4A" authorityName="Brown &amp; Parker" authorityYear="1977" box="[512,650,1433,1459]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="orientalis">
<emphasis id="84B0EAE78055FFBAFDA976ECDEEEFA4A" box="[512,650,1433,1459]" italics="true" pageId="4" pageNumber="309">L. orientalis</emphasis>
</taxonomicName>
in having the precloacal/femoral pore series arrayed into three series (
<emphasis id="84B0EAE78055FFBAFF0876C8DC8DFA2E" box="[161,233,1470,1495]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
a continuous row in the other species); from
<taxonomicName id="71C44D768055FFBAFD7676C8DF30FA2E" authorityName="Brown &amp; Parker" authorityYear="1977" box="[735,852,1470,1495]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="magnus">
<emphasis id="84B0EAE78055FFBAFD7676C8DF30FA2E" box="[735,852,1470,1495]" italics="true" pageId="4" pageNumber="309">L. magnus</emphasis>
</taxonomicName>
in its much smaller size (SVL =
<quantity id="713C9B108055FFBAFB6876CAD92BFA2E" box="[1217,1359,1468,1495]" metricMagnitude="-2" metricUnit="m" metricValue="4.275" metricValueMax="4.45" metricValueMin="4.1" pageId="4" pageNumber="309" unit="mm" value="42.75" valueMax="44.5" valueMin="41.0">4144.5 mm</quantity>
<emphasis id="84B0EAE78055FFBAFAFC76C8D9F9FA2E" box="[1365,1437,1470,1495]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFF3E7696DD6BFA02" box="[151,271,1504,1531]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="309" unit="mm" value="60.0" valueMax="70.0" valueMin="50.0">5070 mm</quantity>
in
<taxonomicName id="71C44D768055FFBAFE987694DDC1FA02" authorityName="Brown &amp; Parker" authorityYear="1977" box="[305,421,1506,1531]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="magnus">
<emphasis id="84B0EAE78055FFBAFE987694DDC1FA02" box="[305,421,1506,1531]" italics="true" pageId="4" pageNumber="309">L. magnus</emphasis>
</taxonomicName>
), fewer lamellae on T1 (78
<emphasis id="84B0EAE78055FFBAFD4E7694DF4BFA02" box="[743,815,1506,1531]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFC9C7697DFEDFA02" box="[821,905,1505,1531]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="4" pageNumber="309" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">912 in</quantity>
<taxonomicName id="71C44D768055FFBAFC267694D867FA02" authorityName="Brown &amp; Parker" authorityYear="1977" box="[911,1027,1506,1531]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="magnus">
<emphasis id="84B0EAE78055FFBAFC267694D867FA02" box="[911,1027,1506,1531]" italics="true" pageId="4" pageNumber="309">L. magnus</emphasis>
</taxonomicName>
), and in having the precloacal/femoral pores arrayed in three series (
<emphasis id="84B0EAE78055FFBAFDA47570DE31F9E6" box="[525,597,1542,1567]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
usually in a single series in
<taxonomicName id="71C44D768055FFBAFC337570D86BF9E6" authorityName="Brown &amp; Parker" authorityYear="1977" box="[922,1039,1542,1567]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="magnus">
<emphasis id="84B0EAE78055FFBAFC337570D86BF9E6" box="[922,1039,1542,1567]" italics="true" pageId="4" pageNumber="309">L. magnus</emphasis>
</taxonomicName>
, which may, however, have one or a few scales disrupting the series); and from
<taxonomicName id="71C44D768055FFBAFD39755CDF63F9BA" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[656,775,1577,1603]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="84B0EAE78055FFBAFD39755CDF63F9BA" box="[656,775,1577,1603]" italics="true" pageId="4" pageNumber="309">L. pumilus</emphasis>
</taxonomicName>
in having the femoral and precloacal pore series separated by only 35 scales (
<emphasis id="84B0EAE78055FFBAFED47538DDA1F99E" box="[381,453,1614,1639]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
712 scales in
<taxonomicName id="71C44D768055FFBAFDDF7538DE8AF99E" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[630,750,1613,1639]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="84B0EAE78055FFBAFDDF7538DE8AF99E" box="[630,750,1613,1639]" italics="true" pageId="4" pageNumber="309">L. pumilus</emphasis>
</taxonomicName>
), and less toe webbing (T3T4webL/T4L = 0.100.13
<emphasis id="84B0EAE78055FFBAFAFC7538D9F9F99E" box="[1365,1437,1614,1639]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFF3E7507DD42F972" box="[151,294,1649,1675]" metricMagnitude="-3" metricUnit="m" metricValue="4.699" metricValueMax="4.826" metricValueMin="4.572" pageId="4" pageNumber="309" unit="in" value="0.185" valueMax="0.19" valueMin="0.18">0.180.19 in</quantity>
<taxonomicName id="71C44D768055FFBAFE877504DDC3F972" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[302,423,1649,1675]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="84B0EAE78055FFBAFE877504DDC3F972" box="[302,423,1649,1675]" italics="true" pageId="4" pageNumber="309">L. pumilus</emphasis>
</taxonomicName>
,
<figureCitation id="2EFF2A708055FFBAFE1F7507DE6BF972" box="[438,527,1649,1675]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="4" pageNumber="309">Fig. 1D</figureCitation>
), especially between T4 and T5 (T4T5webL/T4L = 0.080.12
<emphasis id="84B0EAE78055FFBAFB747504D941F972" box="[1245,1317,1650,1675]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFA847507D919F972" box="[1325,1405,1649,1675]" metricMagnitude="-3" metricUnit="m" metricValue="5.842" pageId="4" pageNumber="309" unit="in" value="0.23">0.23 in</quantity>
<taxonomicName id="71C44D768055FFBAFA2F7504DC94F956" baseAuthorityName="Boulenger" baseAuthorityYear="1885" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="84B0EAE78055FFBAFA2F7504DC94F956" italics="true" pageId="4" pageNumber="309">L. pumilus</emphasis>
</taxonomicName>
).
<taxonomicName id="71C44D768055FFBAFEA075E3DE77F956" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[265,531,1685,1711]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="zweifeli" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFEA075E3DE77F956" box="[265,531,1685,1711]" italics="true" pageId="4" pageNumber="309">Lepidodactylus zweifeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFDB575E3DE1CF956" box="[540,632,1685,1711]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFDB575E3DE1CF956" bold="true" box="[540,632,1685,1711]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="71C44D768055FFBAFCBB75E0DFFCF956" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[786,920,1686,1711]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFCBB75E0DFFCF956" box="[786,920,1686,1711]" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFC0875E3DF99F956" box="[929,1021,1685,1711]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFC0875E3DF99F956" bold="true" box="[929,1021,1685,1711]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
in its larger size (SVL
<quantity id="713C9B108055FFBAFAA475E2D9F9F956" box="[1293,1437,1684,1711]" metricMagnitude="-2" metricUnit="m" metricValue="4.275" metricValueMax="4.45" metricValueMin="4.1" pageId="4" pageNumber="309" unit="mm" value="42.75" valueMax="44.5" valueMin="41.0">4144.5 mm</quantity>
<emphasis id="84B0EAE78055FFBAFF3E75CCDCBBF92A" box="[151,223,1722,1747]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFF4E75CEDD28F92A" box="[231,332,1720,1747]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="4" pageNumber="309" unit="mm" value="37.5">37.5 mm</quantity>
in
<taxonomicName id="71C44D768055FFBAFEDB75CCDD9CF92A" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[370,504,1722,1747]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFEDB75CCDD9CF92A" box="[370,504,1722,1747]" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFDA975CFDE3EF92A" box="[512,602,1721,1747]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFDA975CFDE3EF92A" bold="true" box="[512,602,1721,1747]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
), in having enlarged femoral scales (absent in
<taxonomicName id="71C44D768055FFBAFBC675CCD891F92A" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1135,1269,1722,1747]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFBC675CCD891F92A" box="[1135,1269,1722,1747]" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFB5475CFD933F92A" box="[1277,1367,1721,1747]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFB5475CFD933F92A" bold="true" box="[1277,1367,1721,1747]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
) such that the total number of enlarged precloacal/femoral scales is 40 (
<emphasis id="84B0EAE78055FFBAFCDE75A8DFDBF90E" box="[887,959,1758,1783]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFC6F75ABDF9BF90E" box="[966,1023,1757,1783]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="4" pageNumber="309" unit="in" value="17.0">17 in</quantity>
<taxonomicName id="71C44D768055FFBAFBAF75A8D8EFF90E" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1030,1163,1758,1783]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFBAF75A8D8EFF90E" box="[1030,1163,1758,1783]" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFB3B75ABD88FF90E" box="[1170,1259,1757,1783]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFB3B75ABD88FF90E" bold="true" box="[1170,1259,1757,1783]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
), narrower toes (T4W/T4L = 0.2427
<emphasis id="84B0EAE78055FFBAFE397474DDBCF8E2" box="[400,472,1794,1819]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFE777477DE48F8E2" box="[478,556,1793,1819]" metricMagnitude="-3" metricUnit="m" metricValue="8.382" pageId="4" pageNumber="309" unit="in" value="0.33">0.33 in</quantity>
<taxonomicName id="71C44D768055FFBAFD9A7474DED3F8E2" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[563,695,1794,1819]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFD9A7474DED3F8E2" box="[563,695,1794,1819]" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFD147477DF73F8E2" box="[701,791,1793,1819]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFD147477DF73F8E2" bold="true" box="[701,791,1793,1819]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<figureCitation id="2EFF2A708055FFBAFC8D7477DF1CF8E2" box="[804,888,1793,1819]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="4" pageNumber="309">Fig. 1C</figureCitation>
<emphasis id="84B0EAE78055FFBAFCD67474DFA3F8E2" box="[895,967,1794,1819]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
1D), less toe webbing (T3T4webL/T4L = 0.100.13
<emphasis id="84B0EAE78055FFBAFEA77450DD32F8C6" box="[270,342,1830,1855]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFEF57453DDCEF8C6" box="[348,426,1829,1855]" metricMagnitude="-3" metricUnit="m" metricValue="6.604" pageId="4" pageNumber="309" unit="in" value="0.26">0.26 in</quantity>
<taxonomicName id="71C44D768055FFBAFE187450DE51F8C6" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[433,565,1830,1855]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFE187450DE51F8C6" box="[433,565,1830,1855]" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFD957453DEF1F8C6" box="[572,661,1829,1855]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFD957453DEF1F8C6" bold="true" box="[572,661,1829,1855]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<figureCitation id="2EFF2A708055FFBAFD0A7453DE92F8C6" box="[675,758,1829,1855]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="4" pageNumber="309">Fig. 1C</figureCitation>
<emphasis id="84B0EAE78055FFBAFD577450DF22F8C6" box="[766,838,1830,1855]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
1D), narrower snout (EN/IN = 1.721.79
<emphasis id="84B0EAE78055FFBAFAB47450D901F8C6" box="[1309,1381,1830,1855]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
<quantity id="713C9B108055FFBAFAC27453DCC9F89A" metricMagnitude="-2" metricUnit="m" metricValue="4.0894" pageId="4" pageNumber="309" unit="in" value="1.61">1.61 in</quantity>
<taxonomicName id="71C44D768055FFBAFF1A743CDD53F89A" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[179,311,1866,1891]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFF1A743CDD53F89A" box="[179,311,1866,1891]" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFE94743FDDF2F89A" box="[317,406,1865,1891]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFE94743FDDF2F89A" bold="true" box="[317,406,1865,1891]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
), and dorsum without pale spots (
<emphasis id="84B0EAE78055FFBAFCA6743CDF33F89A" box="[783,855,1866,1891]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
with lateral and dorsolateral rows of pale spots in
<taxonomicName id="71C44D768055FFBAFA2F743CDC9AF87E" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="309" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78055FFBAFA2F743CDC9AF87E" italics="true" pageId="4" pageNumber="309">L. aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8055FFBAFEAC741BDD3AF87E" box="[261,350,1901,1927]" pageId="4" pageNumber="309" rank="species">
<emphasis id="84B0EAE78055FFBAFEAC741BDD3AF87E" bold="true" box="[261,350,1901,1927]" pageId="4" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<figureCitation id="2EFF2A708055FFBAFEC5741BDDA4F87E" box="[364,448,1901,1927]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="4" pageNumber="309">Fig. 1A</figureCitation>
<emphasis id="84B0EAE78055FFBAFE6E7418DE6BF87E" box="[455,527,1902,1927]" italics="true" pageId="4" pageNumber="309">versus</emphasis>
1B).
</paragraph>
<paragraph id="B67B36F58055FFBBFF6E74E4DF93FDC6" blockId="4.[151,1437,1000,2035]" lastBlockId="5.[151,1437,152,2015]" lastPageId="5" lastPageNumber="310" pageId="4" pageNumber="309">
<emphasis id="84B0EAE78055FFBAFF6E74E4DDA9F852" box="[199,461,1937,1963]" italics="true" pageId="4" pageNumber="309">
Description of
<typeStatus id="697F88578055FFBAFEC474E7DDA9F852" box="[365,461,1937,1963]" pageId="4" pageNumber="309" type="holotype">holotype</typeStatus>
</emphasis>
. A mature male of medium size (SVL = 41.0 mm, TrL = 21.0 mm). Head relatively long (HL/SVL = 0.23) and wide (HW/HL = 0.76), distinct from neck. Loreal region slightly inflated; no distinct canthus rostralis. Top of snout, area between nares, and area posterior to nares concave. Snout tapered and rounded at tip, relatively long (SN/HL = 0.43), longer than eye diameter (SN/EY = 1.5). Eye relatively large (EY/HL = 0.28, EY/ EN = 0.84); pupil vertical, constricted into series of four lobes; supraciliaries slightly larger than adjacent granules. Ear opening small (Ear/HL = 0.065), compressed, oriented diagonally from posterodorsal to anteroventral; distance between ear and eye slightly larger than eye diameter (EE/EY = 1.1). Rostral wider (
<quantity id="713C9B108054FFBBFBFA7273D8C8FEE6" box="[1107,1196,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="310" unit="mm" value="1.7">1.7 mm</quantity>
) than high (
<quantity id="713C9B108054FFBBFA9D7273D9F4FEE6" box="[1332,1424,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="310" unit="mm" value="0.9">0.9 mm</quantity>
), highest just medial to nares, lower between these points; length
<quantity id="713C9B108054FFBBFCCB725EDFDFFEBA" box="[866,955,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="310" unit="mm" value="0.5">0.5 mm</quantity>
; dorsal portion with short groove on posterior
<quantity id="713C9B108054FFBBFF6E723BDD45FE9E" box="[199,289,333,359]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="310" unit="mm" value="0.2">0.2 mm</quantity>
. Supranasals separated by three internasals. Rostral in contact with first supralabials, two supranasals, and three internasals. External nares circular; each bordered by rostral, two supranasals, first supralabial, and one postnasal. Mental shallowly triangular, shorter (
<quantity id="713C9B108054FFBBFD0072E3DF6BFE56" box="[681,783,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="310" unit="mm" value="0.6">0.60 mm</quantity>
) than wide (
<quantity id="713C9B108054FFBBFC3E72E2D864FE56" box="[919,1024,404,431]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="310" unit="mm" value="0.75">0.75 mm</quantity>
). Mental bordered posteriorly by two enlarged postmentals, these bordered posteriorly by slightly enlarged scales that progressively decrease in size posteriorly prior to granular chin scales. First two infralabials bordered below by somewhat enlarged scales; remaining scales below infralabials of approximately same size as throat scales. Supralabials to mid-orbital position eight on each side, to angle of jaw 14 on each side. Infralabials 11 on right, 10 on left.
</paragraph>
<paragraph id="B67B36F58054FFBBFF6E713FDF15FD52" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">Body of modest habitus (TrL/SVL = 0.51), slightly depressed. Dorsal scales on head, body, limbs, and throat tiny, juxtaposed granules, larger on sides and snout; tubercles absent. Ventral scales larger, flat and smooth, subimbricate, gradually decreasing in size laterally to become granular.</paragraph>
<paragraph id="B67B36F58054FFBBFF6E71C3DFF4FCA6" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">
Enlarged precloacal/femoral scales in three series of
<date id="C27A10358054FFBBFCB471C3DFE0FD36" box="[797,900,693,719]" pageId="5" pageNumber="310" value="2012-12-16">12-16-12</date>
scales separated by four slightly smaller scales on right, and three on left; those on thighs only slightly larger than adjacent scales. Precloacal/femoral pores in three series of
<date id="C27A10358054FFBBFF52718BDD0DFCEE" box="[251,361,765,791]" pageId="5" pageNumber="310" value="2012-12-14">12-14-12.</date>
Enlarged scales form a pubic patch between the precloacal series and vent; no row of tiny scales between the precloacal series and either side of the pubic patch; nine scales in a row between apex of enlarged precloacal series and vent. Scales on palms and soles rounded, smooth.
</paragraph>
<paragraph id="B67B36F58054FFBBFF6E701FDF5DFB86" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">
Fore- and hindlimbs relatively small but well-developed (FA/SVL = 0.095, CS/SVL = 0.12). Digits well-developed, narrowly dilated throughout their length (T4W/T4L = 0.27), all but first fingers and toes with recurved claws; clawed phalanges laterally compressed, free above and extending slightly beyond terminal scansors. Subdigital lamellae narrow and smooth, all undivided except for second scansor divided on right first toe; lamellae extend for only slightly more than half length of each toe (T4 scansor L/T4L = 0.54,
<figureCitation id="2EFF2A708054FFBBFC60708FD845FBEA" box="[969,1057,1017,1043]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="5" pageNumber="310">Fig. 1D</figureCitation>
). Lamellae of manus
<date id="C27A10358054FFBBFAB9708FD907FBEA" box="[1296,1379,1017,1043]" pageId="5" pageNumber="310" value="2010-08-06">6810</date>
97 on right, 78997 on left; of pes
<date id="C27A10358054FFBBFD84776BDEFAFBCE" box="[557,670,1053,1079]" pageId="5" pageNumber="310" value="2010-08-09" valueMax="2010-08-09" valueMin="2010-08-09">89910</date>
8 on right, 79998 on left. Relative lengths of digits on manus and pes I &lt;II &lt;V &lt;III &lt;IV. Webbing present between all digits except fingers 13 on manus and toes 12 on pes; toes only basally webbed (T3T4webL/T4L = 0.13,
<figureCitation id="2EFF2A708054FFBBFD7A7713DF48FB86" box="[723,812,1125,1151]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="5" pageNumber="310">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="B67B36F58054FFBBFF6E77FFD8A7FAF6" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">Tail complete, subcylindrical, relatively narrow (TW/SVL = 0.076); lateral margins without skin flanges or spines. Scales of tail small, flat, smooth, subimbricate, larger ventrally than dorsally. Cloacal sacs swollen, with small external orifices situated near lateral margins of vent, cut on right side; two enlarged, blunt postcloacal spurs on each side of tailbase; midventral scales of sac not distinctly larger than those ventrolaterally.</paragraph>
<paragraph id="B67B36F58054FFBBFF6E766FDE76FA3A" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">
<emphasis id="84B0EAE78054FFBBFF6E766FDDDCFACA" box="[199,440,1305,1331]" italics="true" pageId="5" pageNumber="310">Color in preservative</emphasis>
: Dorsal ground color yellowish brown with small, dark-brown markings dorsally, laterally, and on limbs; dark markings connecting to form weak dorsolateral stripe in scapular region and extending posterior to forearm insertion (
<figureCitation id="2EFF2A708054FFBBFE597617DE23FA82" box="[496,583,1377,1403]" captionStart="FIGURE 1" captionStartId="6.[151,250,1719,1744]" captionTargetBox="[170,1373,201,1630]" captionTargetId="figure@6.[170,1417,181,1693]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal views of holotypes of (A) Lepidodactylus aignanus sp. nov., BPBM 17229, and (B) L. zweifeli sp. nov., AMNH 105088; right foot of holotypes of (C) Lepidodactylus aignanus sp. nov., BPBM 17229, and (D) L. zweifeli sp. nov., AMNH 105088. Scale = 2 mm." httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="5" pageNumber="310">Fig. 1B</figureCitation>
). Dark-brown markings on tail forming vague, narrow bands. Snout spotted with small dark-brown markings. Venter pale straw with many brown scales scattered throughout; palms and soles brown. Iris black; pupil pale gray.
</paragraph>
<paragraph id="B67B36F58054FFBBFF6E76B8DCBCF9D6" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">
<emphasis id="84B0EAE78054FFBBFF6E76B8DDA3FA1E" box="[199,455,1485,1511]" italics="true" pageId="5" pageNumber="310">Measurements (in mm)</emphasis>
. SVL = 41.0, TrL = 21.0, TL = 37.0, TW = 3.1, FA = 3.9, CS = 5.1, HL = 9.3, HW = 7.1, Ear = 0.6, EE = 2.9, EY = 2.6, SN = 4.0, EN = 3.1, IN = 1.8, T4L = 4.8, T4W = 1.3, T4lamellaeL = 2.6, T3T4webL = 0.6.
</paragraph>
<paragraph id="B67B36F58054FFBBFF6E754CD882F8D2" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">
<emphasis id="84B0EAE78054FFBBFF6E754CDD4BF9AA" box="[199,303,1594,1619]" italics="true" pageId="5" pageNumber="310">Variation</emphasis>
. The enlarged precloacal and femoral scales are in three series in the female
<typeStatus id="697F88578054FFBBFBD4754FD8B9F9AA" box="[1149,1245,1593,1619]" pageId="5" pageNumber="310" type="paratype">paratype</typeStatus>
(
<date id="C27A10358054FFBBFB45754FD937F9AA" box="[1260,1363,1593,1619]" pageId="5" pageNumber="310" value="2012-12-16">12-16-12</date>
scales right to left), with five small scales intervening between these. The
<typeStatus id="697F88578054FFBBFC2D752BDF80F98E" box="[900,996,1629,1655]" pageId="5" pageNumber="310" type="paratype">paratype</typeStatus>
has a greater number of entire lamellae (12 on each fourth toe), ten scales in a row between the apex of the enlarged precloacal series and the vent, narrower toes (T4W/T4L = 0.24), less webbing (T3T4webL/T4L = 0.10), and a relatively smaller eye (EY/SVL = 0.056, EY/ EN = 0.84) than the
<typeStatus id="697F88578054FFBBFED575BFDD86F91A" box="[380,482,1737,1763]" pageId="5" pageNumber="310" type="holotype">holotype</typeStatus>
. It is more heavily dusted with dark brown laterally and dorsolaterally, with the darkbrown markings forming series of short dashes dorsally and four short lines behind head and in scapular region. The head is more densely spotted with brown markings, and the brown markings are denser on the tail.
</paragraph>
<paragraph id="B67B36F58054FFBBFF6E7440DDE9F86E" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">
<emphasis id="84B0EAE78054FFBBFF6E7440DE37F8B6" box="[199,595,1845,1871]" italics="true" pageId="5" pageNumber="310">
Measurements of
<typeStatus id="697F88578054FFBBFE267440DD97F8B6" box="[399,499,1846,1871]" pageId="5" pageNumber="310" type="paratype">paratype</typeStatus>
(in mm)
</emphasis>
. SVL = 44.5, TrL = 22.0, TL = 45.0, TW = 3.1, FA = 3.9, CS = 5.2, HL = 9.9, HW = 7.2, Ear = 0.6, EE = 3.1, EY = 2.5, SN = 4.5, EN = 3.4, IN = 1.9, T4L = 5.0, T4W = 1.2, T4lamellae L = 3.1, T3T4webL = 0.5.
</paragraph>
<paragraph id="B67B36F58054FFBBFF6E74D7D94DF826" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="310">
<emphasis id="84B0EAE78054FFBBFF6E74D7DD5BF842" box="[199,319,1953,1979]" italics="true" pageId="5" pageNumber="310">Etymology</emphasis>
. The name is a genitive honorific for the collector, Richard Zweifel, whose contributions to knowledge of the herpetofauna of New
<collectingCountry id="CED376658054FFBBFDA774B3DE3BF826" box="[526,607,1989,2015]" name="Guinea-Bissau" pageId="5" pageNumber="310">Guinea</collectingCountry>
and nearby islands were foundational to current understanding.
</paragraph>
<caption id="E2BB667D8057FFB8FF3E75C1DDB3F8E1" httpUri="https://zenodo.org/record/3363247/files/figure.png" pageId="6" pageNumber="311" startId="6.[151,250,1719,1744]" targetBox="[170,1373,201,1630]" targetPageId="6">
<paragraph id="B67B36F58057FFB8FF3E75C1DDB3F8E1" blockId="6.[151,1436,1719,1816]" pageId="6" pageNumber="311">
<emphasis id="84B0EAE78057FFB8FF3E75C1DD71F929" bold="true" box="[151,277,1719,1744]" pageId="6" pageNumber="311">FIGURE 1.</emphasis>
Dorsal views of holotypes of (A)
<taxonomicName id="71C44D768057FFB8FDD075CEDF1DF929" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[633,889,1720,1744]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="311" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78057FFB8FDD075CEDF1DF929" box="[633,889,1720,1744]" italics="true" pageId="6" pageNumber="311">Lepidodactylus aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8057FFB8FC2975CEDFB6F929" box="[896,978,1720,1744]" pageId="6" pageNumber="311" rank="species">
<emphasis id="84B0EAE78057FFB8FC2975CEDFB6F929" bold="true" box="[896,978,1720,1744]" pageId="6" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
, BPBM 17229, and (B)
<taxonomicName id="71C44D768057FFB8FB7B75CFD95FF929" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1234,1339,1720,1744]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="311" phylum="Chordata" rank="species" species="zweifeli" status="sp. nov.">
<emphasis id="84B0EAE78057FFB8FB7B75CFD95FF929" box="[1234,1339,1720,1744]" italics="true" pageId="6" pageNumber="311">L. zweifeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8057FFB8FAED75CED9F1F929" box="[1348,1429,1720,1744]" pageId="6" pageNumber="311" rank="species">
<emphasis id="84B0EAE78057FFB8FAED75CED9F1F929" bold="true" box="[1348,1429,1720,1744]" pageId="6" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
, AMNH 105088; right foot of holotypes of (C)
<taxonomicName id="71C44D768057FFB8FDD275AADF1EF90D" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[635,890,1756,1780]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="311" phylum="Chordata" rank="species" species="aignanus" status="sp. nov.">
<emphasis id="84B0EAE78057FFB8FDD275AADF1EF90D" box="[635,890,1756,1780]" italics="true" pageId="6" pageNumber="311">Lepidodactylus aignanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8057FFB8FC2875AADFB7F90D" box="[897,979,1756,1780]" pageId="6" pageNumber="311" rank="species">
<emphasis id="84B0EAE78057FFB8FC2875AADFB7F90D" bold="true" box="[897,979,1756,1780]" pageId="6" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
, BPBM 17229, and (D)
<taxonomicName id="71C44D768057FFB8FB7A75ABD958F90D" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1235,1340,1756,1780]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="311" phylum="Chordata" rank="species" species="zweifeli" status="sp. nov.">
<emphasis id="84B0EAE78057FFB8FB7A75ABD958F90D" box="[1235,1340,1756,1780]" italics="true" pageId="6" pageNumber="311">L. zweifeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8057FFB8FAED75AAD9F1F90D" box="[1348,1429,1756,1780]" pageId="6" pageNumber="311" rank="species">
<emphasis id="84B0EAE78057FFB8FAED75AAD9F1F90D" bold="true" box="[1348,1429,1756,1780]" pageId="6" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
, AMNH 105088. Scale = 2 mm.
</paragraph>
</caption>
<paragraph id="B67B36F58057FFB8FF6E7431DC96F87C" blockId="6.[151,1437,1862,2033]" pageId="6" pageNumber="311">
<emphasis id="84B0EAE78057FFB8FF6E7431DD6BF899" box="[199,271,1863,1888]" italics="true" pageId="6" pageNumber="311">Range</emphasis>
. Known only from the
<typeStatus id="697F88578057FFB8FDB37431DE2EF898" box="[538,586,1863,1889]" pageId="6" pageNumber="311">type</typeStatus>
locality, but likely to occur throughout the hill forests of the Adelbert Mts. (
<figureCitation id="2EFF2A708057FFB8FF09741CDC81F87C" box="[160,229,1898,1925]" captionStart="FIGURE 2" captionStartId="7.[151,248,874,899]" captionTargetBox="[162,1379,249,813]" captionTargetId="figure@7.[162,1379,249,813]" captionTargetPageId="7" captionText="FIGURE 2. Map of New Guinea and surrounding islands showing type localities for Lepidodactylus aignanus sp. nov. (diamond), L. dialeukos sp. nov. (triangle), L. kwasnickae sp. nov. (square), L. mitchelli sp. nov. (circle), and L. zweifeli sp. nov. (star)." httpUri="https://zenodo.org/record/3363249/files/figure.png" pageId="6" pageNumber="311">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="B67B36F58057FFB9FF6E74F8DFA2FF2E" blockId="6.[151,1437,1862,2033]" lastBlockId="7.[151,1437,153,215]" lastPageId="7" lastPageNumber="312" pageId="6" pageNumber="311">
<emphasis id="84B0EAE78057FFB8FF6E74F8DD4DF851" box="[199,297,1934,1960]" italics="true" pageId="6" pageNumber="311">Remarks</emphasis>
. These specimens were briefly discussed by
<bibRefCitation id="D2554B048057FFB8FCB274F9D841F850" author="Brown, W. C. &amp; Parker, F." box="[795,1061,1934,1961]" pageId="6" pageNumber="311" pagination="253 - 265" refId="ref18821" refString="Brown, W. C. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Sciences, 41, 253 - 265." type="journal article" year="1977">Brown &amp; Parker (1977)</bibRefCitation>
, who noted that they were similar in webbing but smaller than
<taxonomicName id="71C44D768057FFB8FE7574C5DE35F835" authorityName="Brown &amp; Parker" authorityYear="1977" box="[476,593,1971,1996]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="311" phylum="Chordata" rank="species" species="magnus">
<emphasis id="84B0EAE78057FFB8FE7574C5DE35F835" box="[476,593,1971,1996]" italics="true" pageId="6" pageNumber="311">L. magnus</emphasis>
</taxonomicName>
and were more like
<taxonomicName id="71C44D768057FFB8FC9574C5DFD7F835" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[828,947,1970,1996]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="311" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="84B0EAE78057FFB8FC9574C5DFD7F835" box="[828,947,1970,1996]" italics="true" pageId="6" pageNumber="311">L. pumilus</emphasis>
</taxonomicName>
in size and possibly pore pattern. As noted above, in addition to size,
<taxonomicName id="71C44D768057FFB8FE6874A1DE51F809" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[449,565,2006,2032]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="311" phylum="Chordata" rank="species" species="zweifeli" status="sp. nov.">
<emphasis id="84B0EAE78057FFB8FE6874A1DE51F809" box="[449,565,2006,2032]" italics="true" pageId="6" pageNumber="311">L. zweifeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8057FFB8FD9574A1DEF2F808" box="[572,662,2007,2033]" pageId="6" pageNumber="311" rank="species">
<emphasis id="84B0EAE78057FFB8FD9574A1DEF2F808" bold="true" box="[572,662,2007,2033]" pageId="6" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="71C44D768057FFB8FC8574A1DFC5F809" authorityName="Brown &amp; Parker" authorityYear="1977" box="[812,929,2007,2032]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="311" phylum="Chordata" rank="species" species="magnus">
<emphasis id="84B0EAE78057FFB8FC8574A1DFC5F809" box="[812,929,2007,2032]" italics="true" pageId="6" pageNumber="311">L. magnus</emphasis>
</taxonomicName>
in its tripartite distribution of the precloacal/ femoral pores and in having fewer T4 lamellae. The easiest way to distinguish
<taxonomicName id="71C44D768056FFB9FBA473ECD8E4FF4A" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1037,1152,153,179]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="312" phylum="Chordata" rank="species" species="zweifeli" status="sp. nov.">
<emphasis id="84B0EAE78056FFB9FBA473ECD8E4FF4A" box="[1037,1152,153,179]" italics="true" pageId="7" pageNumber="312">L. zweifeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8056FFB9FB2E73EFD885FF4A" box="[1159,1249,153,179]" pageId="7" pageNumber="312" rank="species">
<emphasis id="84B0EAE78056FFB9FB2E73EFD885FF4A" bold="true" box="[1159,1249,153,179]" pageId="7" pageNumber="312">sp. nov.</emphasis>
</taxonomicNameLabel>
from
<taxonomicName id="71C44D768056FFB9FA8F73ECD9F9FF4A" baseAuthorityName="Boulenger" baseAuthorityYear="1885" box="[1318,1437,153,179]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="312" phylum="Chordata" rank="species" species="pumilus">
<emphasis id="84B0EAE78056FFB9FA8F73ECD9F9FF4A" box="[1318,1437,153,179]" italics="true" pageId="7" pageNumber="312">L. pumilus</emphasis>
</taxonomicName>
is by the extensive webbing of the latter (especially between T4 and T5).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="79710B62E6708D84BDA110067A060A85">New species of Lepidodactylus (Squamata: Gekkonidae) from New Guinea and adjacent islands</mods:title>
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<treatment id="3E6D87E3805AFFAEFF3E7098DE7CFAF6" ID-GBIF-Taxon="159043707" LSID="urn:lsid:zoobank.org:act:9D4E2932-2478-437E-8F2E-EF67C78D19EB" httpUri="http://treatment.plazi.org/id/3E6D87E3805AFFAEFF3E7098DE7CFAF6" lastPageId="16" lastPageNumber="321" pageId="11" pageNumber="316">
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<heading id="ED338199805AFFB5FF3E7098DE22FBF0" bold="true" box="[151,582,1006,1033]" fontSize="11" level="1" pageId="11" pageNumber="316" reason="1">
<emphasis id="84B0EAE7805AFFB5FF3E7098DE22FBF0" bold="true" box="[151,582,1006,1033]" pageId="11" pageNumber="316">
<taxonomicName id="71C44D76805AFFB5FF3E7098DD85FBF0" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[151,481,1006,1033]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE7805AFFB5FF3E7098DD85FBF0" bold="true" box="[151,481,1006,1033]" italics="true" pageId="11" pageNumber="316">Lepidodactylus kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805AFFB5FE417099DE22FBF0" box="[488,582,1007,1033]" pageId="11" pageNumber="316" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<figureCitation id="2EFF2A70805AFFB5FF3E7764DD68FBD4" box="[151,268,1042,1069]" captionStart="FIGURE 3" captionStartId="10.[151,250,1876,1901]" captionTargetBox="[197,1391,727,1826]" captionTargetId="figure@10.[154,1433,719,1851]" captionTargetPageId="10" captionText="FIGURE 3. Ventral views of (A) holotype of L. mitchelli sp. nov. (UMMZ 244058), (B) paratype of L. kwasnickae sp. nov. (BPBM 39884), and (C) paratype of L. novaeguineae (AMNH 100210) showing ventral pigmentation differences among these species; scale bars = 5 mm. Ventral views of (D) left foot of holotype of L. mitchelli sp. nov. (UMMZ 244058), (E) left foot of paratype of L. kwasnickae sp. nov. (BPBM 39884), and (F) right foot of paratype of L. novaeguineae (AMNH 100210) showing toe-width and pigmentation differences among these species; scale bars = 2 mm." httpUri="https://zenodo.org/record/3363251/files/figure.png" pageId="11" pageNumber="316">Fig. 3B, E</figureCitation>
,
<figureCitation id="2EFF2A70805AFFB5FEB17764DD18FBD4" box="[280,380,1042,1069]" captionStart="FIGURE 4" captionStartId="14.[151,250,1805,1830]" captionTargetBox="[151,1436,928,1786]" captionTargetId="figure@14.[151,1436,928,1786]" captionTargetPageId="14" captionText="FIGURE 4. Portraits of new Lepidodactylus species in life. (A) holotype of L. mitchelli sp. nov. (UMMZ 244058), (B) holo- type of L. kwasnickae sp. nov. (BPBM 39880), (C) paratype of L. kwasnickae sp. nov. (BPBM 39879), and (D) paratype of L. kwasnickae sp. nov. (BPBM 39151)." httpUri="https://zenodo.org/record/3363253/files/figure.png" pageId="11" pageNumber="316">4B, C, D</figureCitation>
</paragraph>
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<materialsCitation id="06AC3CA8805AFFB5FF3E7703D937FB4A" ID-GBIF-Occurrence="2313474479" collectingDate="2011-10-12" collectionCode="BPBM" collectorName="F. Kraus &amp; Gisabwai" country="Papua New Guinea" elevation="110" latitude="-9.076" location="Woodlark Island" longLatPrecision="7" longitude="152.7748" pageId="11" pageNumber="316" specimenCode="BPBM 39880" specimenCount="1" stateProvince="Milne Bay" typeStatus="holotype">
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<emphasis id="84B0EAE7805AFFB5FF3E7703DC9AFB76" box="[151,254,1141,1167]" italics="true" pageId="11" pageNumber="316">Holotype</emphasis>
</typeStatus>
.
<specimenCode id="E6629E8E805AFFB5FEA47703DDCCFB76" box="[269,424,1141,1167]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="316" type="Museum">BPBM 39880</specimenCode>
(field tag FK 15809), mature male, collected by
<collectorName id="1B315323805AFFB5FC7A7703D856FB76" box="[979,1074,1141,1167]" pageId="11" pageNumber="316">F. Kraus</collectorName>
and local villagers at
<collectorName id="1B315323805AFFB5FA827703D9FCFB76" box="[1323,1432,1141,1167]" pageId="11" pageNumber="316">Gisabwai</collectorName>
,
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,
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,
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<quantity id="713C9B10805AFFB5FE0977EFDD8CFB4A" box="[416,488,1177,1203]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="316" unit="m" value="110.0">110 m</quantity>
</elevation>
,
<location id="B31B602E805AFFB5FE5D77EFDECBFB4A" LSID="urn:lsid:plazi:treatment:3E6D87E3805AFFAEFF3E7098DE7CFAF6:B31B602E805AFFB5FE5D77EFDECBFB4A" box="[500,687,1177,1203]" country="Papua New Guinea" latitude="-9.076" longLatPrecision="7" longitude="152.7748" name="Woodlark Island" pageId="11" pageNumber="316" stateProvince="Milne Bay">Woodlark Island</location>
,
<collectingRegion id="7400F817805AFFB5FD1277EFDFFBFB4A" box="[699,927,1177,1203]" country="Papua New Guinea" name="Milne Bay" pageId="11" pageNumber="316">Milne Bay Province</collectingRegion>
,
<collectingCountry id="CED37665805AFFB5FC0377EFD8E2FB4A" box="[938,1158,1177,1203]" name="Papua New Guinea" pageId="11" pageNumber="316">Papua New Guinea</collectingCountry>
,
<date id="C27A1035805AFFB5FB3977EFD92BFB4A" box="[1168,1359,1177,1203]" pageId="11" pageNumber="316" value="2011-10-12">
<collectingDate id="D23EE9DD805AFFB5FB3977EFD92BFB4A" box="[1168,1359,1177,1203]" pageId="11" pageNumber="316" value="2011-10-12">12 October 2011</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="B67B36F5805AFFB5FF6E77C8DD88FABA" blockId="11.[151,1437,1140,2031]" pageId="11" pageNumber="316">
<materialsCitation id="06AC3CA8805AFFB5FF6E77C8DFDBFB2E" ID-GBIF-Occurrence="2313474481" box="[199,959,1213,1239]" collectionCode="BPBM" location="Same" pageId="11" pageNumber="316" specimenCode="BPBM 39879, 39881-82" specimenCount="2" typeStatus="paratype">
<emphasis id="84B0EAE7805AFFB5FF6E77C8DDE5FB2E" box="[199,385,1213,1239]" italics="true" pageId="11" pageNumber="316">
<typeStatus id="697F8857805AFFB5FF6E77C8DD5DFB2E" box="[199,313,1214,1239]" pageId="11" pageNumber="316" type="paratype">Paratypes</typeStatus>
(n=9)
</emphasis>
.
<location id="B31B602E805AFFB5FE3977CBDDAAFB2E" LSID="urn:lsid:plazi:treatment:3E6D87E3805AFFAEFF3E7098DE7CFAF6:B31B602E805AFFB5FE3977CBDDAAFB2E" box="[400,462,1213,1239]" name="Same" pageId="11" pageNumber="316">Same</location>
data as
<typeStatus id="697F8857805AFFB5FD8077CBDEEFFB2E" box="[553,651,1213,1239]" pageId="11" pageNumber="316" type="holotype">holotype</typeStatus>
(
<specimenCode id="E6629E8E805AFFB5FD3377CBDF5EFB2E" box="[666,826,1213,1239]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="316" type="Museum">BPBM 39879</specimenCode>
,
<specimenCode id="E6629E8E805AFFB5FCEC77CBDFD3FB2E" box="[837,951,1213,1239]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="316" type="Museum">3988182</specimenCode>
)
</materialsCitation>
<materialsCitation id="06AC3CA8805AFFB5FC6F77CBDD70FB02" ID-GBIF-Occurrence="2313474478" collectingDate="2011-10-17" collectionCode="BPBM" pageId="11" pageNumber="316" specimenCode="BPBM 39883-84" specimenCount="1" typeStatus="paratype">
except collected
<collectingDate id="D23EE9DD805AFFB5FB2B77CBD95AFB2E" box="[1154,1342,1213,1239]" pageId="11" pageNumber="316" value="2011-10-17">
<date id="C27A1035805AFFB5FB2B77CBD95AFB2E" box="[1154,1342,1213,1239]" pageId="11" pageNumber="316" value="2011-10-17">17 October 2011</date>
</collectingDate>
(
<specimenCode id="E6629E8E805AFFB5FAE477CBDD68FB02" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="316" type="Museum">BPBM 3988384</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="06AC3CA8805AFFB5FE8C7797D97EFB02" ID-GBIF-Occurrence="2313474480" box="[293,1306,1248,1275]" collectingDate="2010-10-17" collectionCode="BPBM" elevation="67" latitude="-9.1216" location="Upper Muniai River" longLatPrecision="7" longitude="152.7486" pageId="11" pageNumber="316" specimenCode="BPBM 39149" specimenCount="1" typeStatus="paratype">
<location id="B31B602E805AFFB5FE8C7797DE72FB02" LSID="urn:lsid:plazi:treatment:3E6D87E3805AFFAEFF3E7098DE7CFAF6:B31B602E805AFFB5FE8C7797DE72FB02" box="[293,534,1249,1275]" latitude="-9.1216" longLatPrecision="7" longitude="152.7486" name="Upper Muniai River" pageId="11" pageNumber="316">Upper Muniai River</location>
,
<geoCoordinate id="D3F05032805AFFB5FD8E7797DEF5FB02" box="[551,657,1249,1275]" degrees="9.1216" direction="south" orientation="latitude" pageId="11" pageNumber="316" precision="5" value="-9.1216">9.1216°S</geoCoordinate>
,
<geoCoordinate id="D3F05032805AFFB5FD0A7796DF4FFB02" box="[675,811,1248,1275]" degrees="152.7486" direction="east" orientation="longitude" pageId="11" pageNumber="316" precision="5" value="152.7486">152.7486°E</geoCoordinate>
,
<elevation id="3DE9D1C6805AFFB5FC957797DF19FB02" box="[828,893,1249,1275]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="11" pageNumber="316" unit="m" value="67.0">
<quantity id="713C9B10805AFFB5FC957797DF19FB02" box="[828,893,1249,1275]" metricMagnitude="1" metricUnit="m" metricValue="6.7" pageId="11" pageNumber="316" unit="m" value="67.0">67 m</quantity>
</elevation>
,
<collectingDate id="D23EE9DD805AFFB5FC277797D832FB02" box="[910,1110,1249,1275]" pageId="11" pageNumber="316" value="2010-10-17">
<date id="C27A1035805AFFB5FC277797D832FB02" box="[910,1110,1249,1275]" pageId="11" pageNumber="316" value="2010-10-17">17 October 2010</date>
</collectingDate>
(
<specimenCode id="E6629E8E805AFFB5FBC27797D976FB02" box="[1131,1298,1249,1275]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="316" type="Museum">BPBM 39149</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="06AC3CA8805AFFB5FA827797DFC4FAE6" ID-GBIF-Occurrence="2313474473" collectingDate="2010-10-19" collectionCode="BPBM" elevation="105" latitude="-9.0745" location="Gisabwai" longLatPrecision="5" longitude="152.77264" pageId="11" pageNumber="316" specimenCode="BPBM 39150-51" specimenCount="1" typeStatus="paratype">
<location id="B31B602E805AFFB5FA827797D9FCFB02" LSID="urn:lsid:plazi:treatment:3E6D87E3805AFFAEFF3E7098DE7CFAF6:B31B602E805AFFB5FA827797D9FCFB02" box="[1323,1432,1249,1275]" latitude="-9.0745" longLatPrecision="5" longitude="152.77264" name="Gisabwai" pageId="11" pageNumber="316">Gisabwai</location>
,
<geoCoordinate id="D3F05032805AFFB5FF3E7672DD65FAE6" box="[151,257,1284,1311]" degrees="9.0745" direction="south" orientation="latitude" pageId="11" pageNumber="316" precision="5" value="-9.0745">9.0745°S</geoCoordinate>
,
<geoCoordinate id="D3F05032805AFFB5FEA47672DDC7FAE6" box="[269,419,1284,1311]" degrees="152.77265" direction="east" orientation="longitude" pageId="11" pageNumber="316" precision="1" value="152.77264">152.77265°E</geoCoordinate>
,
<elevation id="3DE9D1C6805AFFB5FE077672DD93FAE6" box="[430,503,1284,1311]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="11" pageNumber="316" unit="m" value="105.0">
<quantity id="713C9B10805AFFB5FE077672DD93FAE6" box="[430,503,1284,1311]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="11" pageNumber="316" unit="m" value="105.0">105 m</quantity>
</elevation>
,
<collectingDate id="D23EE9DD805AFFB5FDAA7673DEDBFAE6" box="[515,703,1285,1311]" pageId="11" pageNumber="316" value="2010-10-19">
<date id="C27A1035805AFFB5FDAA7673DEDBFAE6" box="[515,703,1285,1311]" pageId="11" pageNumber="316" value="2010-10-19">19 October 2010</date>
</collectingDate>
(
<specimenCode id="E6629E8E805AFFB5FD647673DFFDFAE6" box="[717,921,1284,1311]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="316" type="Museum">BPBM 3915051</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="06AC3CA8805AFFB5FC027673DD8DFABA" ID-GBIF-Occurrence="2313474477" collectingDate="2010-10-22" collectionCode="BPBM" elevation="60" latitude="-9.1153" location="Piak Track" longLatPrecision="7" longitude="152.7484" pageId="11" pageNumber="316" specimenCode="BPBM 39152" specimenCount="1" typeStatus="paratype">
<location id="B31B602E805AFFB5FC027673D843FAE6" LSID="urn:lsid:plazi:treatment:3E6D87E3805AFFAEFF3E7098DE7CFAF6:B31B602E805AFFB5FC027673D843FAE6" box="[939,1063,1285,1311]" latitude="-9.1153" longLatPrecision="7" longitude="152.7484" name="Piak Track" pageId="11" pageNumber="316">Piak Track</location>
,
<geoCoordinate id="D3F05032805AFFB5FB9B7672D8FFFAE6" box="[1074,1179,1284,1311]" degrees="9.1153" direction="south" orientation="latitude" pageId="11" pageNumber="316" precision="5" value="-9.1153">9.1153°S</geoCoordinate>
,
<geoCoordinate id="D3F05032805AFFB5FB0F7672D94BFAE6" box="[1190,1327,1284,1311]" degrees="152.7484" direction="east" orientation="longitude" pageId="11" pageNumber="316" precision="5" value="152.7484">152.7484°E</geoCoordinate>
,
<elevation id="3DE9D1C6805AFFB5FA937673D911FAE6" box="[1338,1397,1285,1311]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="316" unit="m" value="60.0">
<quantity id="713C9B10805AFFB5FA937673D911FAE6" box="[1338,1397,1285,1311]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="316" unit="m" value="60.0">60 m</quantity>
</elevation>
,
<collectingDate id="D23EE9DD805AFFB5FA297673DD54FABA" pageId="11" pageNumber="316" value="2010-10-22">
<date id="C27A1035805AFFB5FA297673DD54FABA" pageId="11" pageNumber="316" value="2010-10-22">22 October 2010</date>
</collectingDate>
(
<specimenCode id="E6629E8E805AFFB5FE96765FDD85FABA" box="[319,481,1320,1347]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="316" type="Museum">BPBM 39152</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B67B36F5805AFFB5FF6E7638D9E2F99A" blockId="11.[151,1437,1140,2031]" pageId="11" pageNumber="316">
<emphasis id="84B0EAE7805AFFB5FF6E7638DD5DFA9E" box="[199,313,1358,1383]" italics="true" pageId="11" pageNumber="316">Diagnosis</emphasis>
. A medium-sized species (adult SVL
<quantity id="713C9B10805AFFB5FD73763ADF19FA9E" box="[730,893,1356,1383]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="4.95" metricValueMin="4.15" pageId="11" pageNumber="316" unit="mm" value="45.5" valueMax="49.5" valueMin="41.5">41.549.5 mm</quantity>
) species of
<taxonomicName id="71C44D76805AFFB5FC56763BD8CEFA9E" box="[1023,1194,1357,1383]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="genus">
<emphasis id="84B0EAE7805AFFB5FC56763BD8CEFA9E" box="[1023,1194,1357,1383]" italics="true" pageId="11" pageNumber="316">Lepidodactylus</emphasis>
</taxonomicName>
having a subcylindrical tail without a lateral fringe of enlarged scales; 23 divided subterminal scansors on T4; 1215 enlarged scales of pore-bearing series limited to precloacal region, 1214 precloacal pores in males; 1220 T4 lamellae, 812 T1 lamellae; lamellae occupying most of the length of the toes (T4lamellaeL/T4L = 0.670.93); toes relatively long (T4L/SVL = 0.110.13) and of moderate width (T4W/T4L = 0.260.34); toes approximately one-quarter webbed (T3T4webL/T4L = 0.180.26); numerous (910) small precloacal scales between apex of pore-bearing series and the cloaca in adults; no row of tiny scales between the precloacal series and either side of the pubic patch; row of dorsolateral white spots, lacking dark spots in this region; and many posteroventral and plantar scales uniformly brown.
</paragraph>
<paragraph id="B67B36F5805AFFB2FF6E751BDDE5FE2A" blockId="11.[151,1437,1140,2031]" lastBlockId="12.[151,1437,153,2015]" lastPageId="12" lastPageNumber="317" pageId="11" pageNumber="316">
<emphasis id="84B0EAE7805AFFB5FF6E751BDE5DF97E" box="[199,569,1645,1671]" italics="true" pageId="11" pageNumber="316">Comparisons with other species</emphasis>
. The subcylindrical tail without a lateral fringe of enlarged scales and the 23 divided subterminal scansors under the toes place
<taxonomicName id="71C44D76805AFFB5FCA975E7D850F952" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[768,1076,1681,1707]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE7805AFFB5FCA975E7D850F952" box="[768,1076,1681,1707]" italics="true" pageId="11" pageNumber="316">Lepidodactylus kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805AFFB5FB9575E7D8F2F952" box="[1084,1174,1681,1707]" pageId="11" pageNumber="316" rank="species">
<emphasis id="84B0EAE7805AFFB5FB9575E7D8F2F952" bold="true" box="[1084,1174,1681,1707]" pageId="11" pageNumber="316">sp. nov.</emphasis>
</taxonomicNameLabel>
in
<bibRefCitation id="D2554B04805AFFB5FB1275E7DC86F936" author="Brown, W. C. &amp; Parker, F." pageId="11" pageNumber="316" pagination="253 - 265" refId="ref18821" refString="Brown, W. C. &amp; Parker, F. (1977) Lizards of the genus Lepidodactylus (Gekkonidae) from the Indo-Australian Archipelago and the islands of the Pacific, with descriptions of new species. Proceedings of the California Academy of Sciences, 41, 253 - 265." type="journal article" year="1977">Brown and Parkers (1977)</bibRefCitation>
Group II.
<taxonomicName id="71C44D76805AFFB5FEF375C3DEE9F936" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[346,653,1717,1743]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE7805AFFB5FEF375C3DEE9F936" box="[346,653,1717,1743]" italics="true" pageId="11" pageNumber="316">Lepidodactylus kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805AFFB5FD3C75C3DE8BF936" box="[661,751,1717,1743]" pageId="11" pageNumber="316" rank="species">
<emphasis id="84B0EAE7805AFFB5FD3C75C3DE8BF936" bold="true" box="[661,751,1717,1743]" pageId="11" pageNumber="316">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from other Melanesian members of this group as follows: from
<taxonomicName id="71C44D76805AFFB5FEB275ACDDE2F90A" box="[283,390,1754,1779]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="guppyi">
<emphasis id="84B0EAE7805AFFB5FEB275ACDDE2F90A" box="[283,390,1754,1779]" italics="true" pageId="11" pageNumber="316">L. guppyi</emphasis>
</taxonomicName>
,
<taxonomicName id="71C44D76805AFFB5FE3C75ACDE6EF90A" authorityName="Boulenger" authorityYear="1885" box="[405,522,1753,1779]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="84B0EAE7805AFFB5FE3C75ACDE6EF90A" box="[405,522,1753,1779]" italics="true" pageId="11" pageNumber="316">L. pulcher</emphasis>
</taxonomicName>
, and
<taxonomicName id="71C44D76805AFFB5FDE375ACDED2F90A" baseAuthorityName="Brown &amp; Tanner" baseAuthorityYear="1949" box="[586,694,1753,1779]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="shebae">
<emphasis id="84B0EAE7805AFFB5FDE375ACDED2F90A" box="[586,694,1753,1779]" italics="true" pageId="11" pageNumber="316">L. shebae</emphasis>
</taxonomicName>
in having fewer enlarged precloacal scales (1215
<emphasis id="84B0EAE7805AFFB5FB5675ACD923F90A" box="[1279,1351,1754,1779]" italics="true" pageId="11" pageNumber="316">versus</emphasis>
3952, 1820, and 34, respectively) and fewer precloacal pores (1214
<emphasis id="84B0EAE7805AFFB5FCC97588DFCCF8EE" box="[864,936,1790,1815]" italics="true" pageId="11" pageNumber="316">versus</emphasis>
3952, 16, and 32, respectively); and further differs from
<taxonomicName id="71C44D76805AFFB5FE817454DDFAF8C2" authorityName="Boulenger" authorityYear="1885" box="[296,414,1825,1851]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="84B0EAE7805AFFB5FE817454DDFAF8C2" box="[296,414,1825,1851]" italics="true" pageId="11" pageNumber="316">L. pulcher</emphasis>
</taxonomicName>
in having the lamellae occupy less area on the fourth toe (T4lamellaeL/T4L = 0.670.93
<emphasis id="84B0EAE7805AFFB5FF3E7430DCBBF8A6" box="[151,223,1862,1887]" italics="true" pageId="11" pageNumber="316">versus</emphasis>
1.0 in
<taxonomicName id="71C44D76805AFFB5FE877430DDC6F8A6" authorityName="Boulenger" authorityYear="1885" box="[302,418,1861,1887]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="84B0EAE7805AFFB5FE877430DDC6F8A6" box="[302,418,1861,1887]" italics="true" pageId="11" pageNumber="316">L. pulcher</emphasis>
</taxonomicName>
) and from
<taxonomicName id="71C44D76805AFFB5FD897430DEE9F8A6" baseAuthorityName="Brown &amp; Tanner" baseAuthorityYear="1949" box="[544,653,1861,1887]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="shebae">
<emphasis id="84B0EAE7805AFFB5FD897430DEE9F8A6" box="[544,653,1861,1887]" italics="true" pageId="11" pageNumber="316">L. shebae</emphasis>
</taxonomicName>
in its larger adult size (SVL
<quantity id="713C9B10805AFFB5FC7A7432D812F8A6" box="[979,1142,1860,1887]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="4.95" metricValueMin="4.15" pageId="11" pageNumber="316" unit="mm" value="45.5" valueMax="49.5" valueMin="41.5">41.549.5 mm</quantity>
<emphasis id="84B0EAE7805AFFB5FBD47430D8A1F8A6" box="[1149,1221,1862,1887]" italics="true" pageId="11" pageNumber="316">versus</emphasis>
~
<quantity id="713C9B10805AFFB5FB727433D94FF8A6" box="[1243,1323,1861,1887]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="11" pageNumber="316" unit="mm" value="36.0">36 mm</quantity>
in
<taxonomicName id="71C44D76805AFFB5FAE67430DCDBF87A" baseAuthorityName="Brown &amp; Tanner" baseAuthorityYear="1949" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="shebae">
<emphasis id="84B0EAE7805AFFB5FAE67430DCDBF87A" italics="true" pageId="11" pageNumber="316">L. shebae</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="71C44D76805AFFB5FED7741CDE5CF87A" box="[382,568,1898,1923]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE7805AFFB5FED7741CDE5CF87A" box="[382,568,1898,1923]" italics="true" pageId="11" pageNumber="316">L. novaeguineae</emphasis>
</taxonomicName>
in having more T4 lamellae (1220
<emphasis id="84B0EAE7805AFFB5FC7C741CD879F87A" box="[981,1053,1898,1923]" italics="true" pageId="11" pageNumber="316">versus</emphasis>
<quantity id="713C9B10805AFFB5FB8D741ED81EF87A" box="[1060,1146,1896,1923]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.81" metricValueMin="2.286" pageId="11" pageNumber="316" unit="in" value="12.0" valueMax="15.0" valueMin="9.0">915 in</quantity>
<taxonomicName id="71C44D76805AFFB5FB28741CD95EF87A" box="[1153,1338,1898,1923]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE7805AFFB5FB28741CD95EF87A" box="[1153,1338,1898,1923]" italics="true" pageId="11" pageNumber="316">L. novaeguineae</emphasis>
</taxonomicName>
,
<figureCitation id="2EFF2A70805AFFB5FAE0741FD9F9F87A" box="[1353,1437,1897,1923]" captionStart="FIGURE 3" captionStartId="10.[151,250,1876,1901]" captionTargetBox="[197,1391,727,1826]" captionTargetId="figure@10.[154,1433,719,1851]" captionTargetPageId="10" captionText="FIGURE 3. Ventral views of (A) holotype of L. mitchelli sp. nov. (UMMZ 244058), (B) paratype of L. kwasnickae sp. nov. (BPBM 39884), and (C) paratype of L. novaeguineae (AMNH 100210) showing ventral pigmentation differences among these species; scale bars = 5 mm. Ventral views of (D) left foot of holotype of L. mitchelli sp. nov. (UMMZ 244058), (E) left foot of paratype of L. kwasnickae sp. nov. (BPBM 39884), and (F) right foot of paratype of L. novaeguineae (AMNH 100210) showing toe-width and pigmentation differences among these species; scale bars = 2 mm." httpUri="https://zenodo.org/record/3363251/files/figure.png" pageId="11" pageNumber="316">Fig. 3E</figureCitation>
<emphasis id="84B0EAE7805AFFB5FF3E74F8DCBBF85E" box="[151,223,1934,1959]" italics="true" pageId="11" pageNumber="316">versus</emphasis>
3F), a longer fourth toe (T4L/SVL = 0.110.13
<emphasis id="84B0EAE7805AFFB5FCAC74F8DF29F85E" box="[773,845,1934,1959]" italics="true" pageId="11" pageNumber="316">versus</emphasis>
<quantity id="713C9B10805AFFB5FCFD74FBDFC0F85E" box="[852,932,1933,1959]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="316" unit="in" value="0.1">0.10 in</quantity>
<taxonomicName id="71C44D76805AFFB5FC0574F8D801F85E" box="[940,1125,1934,1959]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE7805AFFB5FC0574F8D801F85E" box="[940,1125,1934,1959]" italics="true" pageId="11" pageNumber="316">L. novaeguineae</emphasis>
</taxonomicName>
), dorsum and sides usually rufescent and with dorsolateral series of white spots (
<emphasis id="84B0EAE7805AFFB5FD4674C4DF53F832" box="[751,823,1970,1995]" italics="true" pageId="11" pageNumber="316">versus</emphasis>
dorsal pattern with longitudinal dark-brown streaks or dorsolateral series of bold dark-brown spots in
<taxonomicName id="71C44D76805AFFB5FD0774A0DF03F816" box="[686,871,2006,2031]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="316" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE7805AFFB5FD0774A0DF03F816" box="[686,871,2006,2031]" italics="true" pageId="11" pageNumber="316">L. novaeguineae</emphasis>
</taxonomicName>
), and many posteroventral and plantar scales uni- formly brown (
<emphasis id="84B0EAE7805DFFB2FEED73ECDDE8FF4A" box="[324,396,154,179]" italics="true" pageId="12" pageNumber="317">versus</emphasis>
only with minute black punctations in
<taxonomicName id="71C44D76805DFFB2FCE073ECD866FF4A" box="[841,1026,154,179]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="317" phylum="Chordata" rank="species" species="novaeguineae">
<emphasis id="84B0EAE7805DFFB2FCE073ECD866FF4A" box="[841,1026,154,179]" italics="true" pageId="12" pageNumber="317">L. novaeguineae</emphasis>
</taxonomicName>
,
<figureCitation id="2EFF2A70805DFFB2FBBB73EFD802FF4A" box="[1042,1126,153,179]" captionStart="FIGURE 3" captionStartId="10.[151,250,1876,1901]" captionTargetBox="[197,1391,727,1826]" captionTargetId="figure@10.[154,1433,719,1851]" captionTargetPageId="10" captionText="FIGURE 3. Ventral views of (A) holotype of L. mitchelli sp. nov. (UMMZ 244058), (B) paratype of L. kwasnickae sp. nov. (BPBM 39884), and (C) paratype of L. novaeguineae (AMNH 100210) showing ventral pigmentation differences among these species; scale bars = 5 mm. Ventral views of (D) left foot of holotype of L. mitchelli sp. nov. (UMMZ 244058), (E) left foot of paratype of L. kwasnickae sp. nov. (BPBM 39884), and (F) right foot of paratype of L. novaeguineae (AMNH 100210) showing toe-width and pigmentation differences among these species; scale bars = 2 mm." httpUri="https://zenodo.org/record/3363251/files/figure.png" pageId="12" pageNumber="317">Fig. 3E</figureCitation>
<emphasis id="84B0EAE7805DFFB2FBC473ECD8D1FF4A" box="[1133,1205,154,179]" italics="true" pageId="12" pageNumber="317">versus</emphasis>
3F)). It differs from
<taxonomicName id="71C44D76805DFFB2FF3E73C8DD73FF2E" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[151,279,189,215]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="317" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805DFFB2FF3E73C8DD73FF2E" box="[151,279,189,215]" italics="true" pageId="12" pageNumber="317">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805DFFB2FEB773CBDD13FF2E" box="[286,375,189,215]" pageId="12" pageNumber="317" rank="species">
<emphasis id="84B0EAE7805DFFB2FEB773CBDD13FF2E" bold="true" box="[286,375,189,215]" pageId="12" pageNumber="317">sp. nov.</emphasis>
</taxonomicNameLabel>
in its larger adult size (SVL = mean
<quantity id="713C9B10805DFFB2FCBC73CADF1FFF2E" box="[789,891,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="4.57" pageId="12" pageNumber="317" unit="mm" value="45.7">45.7 mm</quantity>
, range
<quantity id="713C9B10805DFFB2FC6573CAD80BFF2E" box="[972,1135,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="4.95" metricValueMin="4.15" pageId="12" pageNumber="317" unit="mm" value="45.5" valueMax="49.5" valueMin="41.5">41.549.5 mm</quantity>
<emphasis id="84B0EAE7805DFFB2FBDF73C8D8DAFF2E" box="[1142,1214,190,215]" italics="true" pageId="12" pageNumber="317">versus</emphasis>
mean 40.3, range =
<quantity id="713C9B10805DFFB2FF3E7396DD41FF02" box="[151,293,224,251]" metricMagnitude="-2" metricUnit="m" metricValue="4.025" metricValueMax="4.55" metricValueMin="3.5" pageId="12" pageNumber="317" unit="mm" value="40.25" valueMax="45.5" valueMin="35.0">3545.5 mm</quantity>
in
<taxonomicName id="71C44D76805DFFB2FEEE7394DDA2FF02" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[327,454,225,251]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="317" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805DFFB2FEEE7394DDA2FF02" box="[327,454,225,251]" italics="true" pageId="12" pageNumber="317">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805DFFB2FE647397DE42FF02" box="[461,550,225,251]" pageId="12" pageNumber="317" rank="species">
<emphasis id="84B0EAE7805DFFB2FE647397DE42FF02" bold="true" box="[461,550,225,251]" pageId="12" pageNumber="317">sp. nov.</emphasis>
</taxonomicNameLabel>
, mass = mean
<quantity id="713C9B10805DFFB2FD657397DF77FF02" box="[716,787,225,251]" metricMagnitude="-3" metricUnit="kg" metricValue="2.36" pageId="12" pageNumber="317" unit="g" value="2.36">2.36 g</quantity>
, range
<quantity id="713C9B10805DFFB2FCCA7396DF8CFF02" box="[867,1000,224,251]" metricMagnitude="-3" metricUnit="kg" metricValue="2.4" metricValueMax="2.95" metricValueMin="1.85" pageId="12" pageNumber="317" unit="g" value="2.4" valueMax="2.95" valueMin="1.85">1.852.95 g</quantity>
<emphasis id="84B0EAE7805DFFB2FC467394D853FF02" box="[1007,1079,226,251]" italics="true" pageId="12" pageNumber="317">versus</emphasis>
mean
<quantity id="713C9B10805DFFB2FB297397D8A3FF02" box="[1152,1223,225,251]" metricMagnitude="-3" metricUnit="kg" metricValue="1.63" pageId="12" pageNumber="317" unit="g" value="1.63">1.63 g</quantity>
, range
<quantity id="713C9B10805DFFB2FAB17396D9F8FF02" box="[1304,1436,224,251]" metricMagnitude="-3" metricUnit="kg" metricValue="1.65" metricValueMax="2.15" metricValueMin="1.15" pageId="12" pageNumber="317" unit="g" value="1.65" valueMax="2.15" valueMin="1.15">1.152.15 g</quantity>
in
<taxonomicName id="71C44D76805DFFB2FF1D7270DD50FEE6" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[180,308,261,287]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="317" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805DFFB2FF1D7270DD50FEE6" box="[180,308,261,287]" italics="true" pageId="12" pageNumber="317">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805DFFB2FE927273DDF1FEE6" box="[315,405,261,287]" pageId="12" pageNumber="317" rank="species">
<emphasis id="84B0EAE7805DFFB2FE927273DDF1FEE6" bold="true" box="[315,405,261,287]" pageId="12" pageNumber="317">sp. nov.</emphasis>
</taxonomicNameLabel>
); longer fourth toe (T4L/SVL = 0.110.13
<emphasis id="84B0EAE7805DFFB2FCD37270DFA6FEE6" box="[890,962,262,287]" italics="true" pageId="12" pageNumber="317">versus</emphasis>
<quantity id="713C9B10805DFFB2FC607273D807FEE6" box="[969,1123,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="2.5273000000000003" metricValueMax="2.7939999999999996" metricValueMin="2.2605999999999997" pageId="12" pageNumber="317" unit="in" value="0.0995" valueMax="0.11" valueMin="0.089">0.0890.11 in</quantity>
<taxonomicName id="71C44D76805DFFB2FBC37270D88EFEE6" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1130,1258,261,287]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="317" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805DFFB2FBC37270D88EFEE6" box="[1130,1258,261,287]" italics="true" pageId="12" pageNumber="317">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805DFFB2FB587273D92EFEE6" box="[1265,1354,261,287]" pageId="12" pageNumber="317" rank="species">
<emphasis id="84B0EAE7805DFFB2FB587273D92EFEE6" bold="true" box="[1265,1354,261,287]" pageId="12" pageNumber="317">sp. nov.</emphasis>
</taxonomicNameLabel>
); more and smaller scales between the enlarged precloacal series and the opening of the cloaca in adults (910 vs
<quantity id="713C9B10805DFFB2FAFD725FD9F8FEBA" box="[1364,1436,297,323]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="12" pageNumber="317" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">68 in</quantity>
<taxonomicName id="71C44D76805DFFB2FF3E7238DD73FE9E" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[151,279,333,359]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="317" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805DFFB2FF3E7238DD73FE9E" box="[151,279,333,359]" italics="true" pageId="12" pageNumber="317">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805DFFB2FEB7723BDD1CFE9E" box="[286,376,333,359]" pageId="12" pageNumber="317" rank="species">
<emphasis id="84B0EAE7805DFFB2FEB7723BDD1CFE9E" bold="true" box="[286,376,333,359]" pageId="12" pageNumber="317">sp. nov.</emphasis>
</taxonomicNameLabel>
); tiny scales between the precloacal series and either side of the pubic patch absent (present in
<taxonomicName id="71C44D76805DFFB2FF3E7204DD73FE72" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[151,279,369,395]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="317" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805DFFB2FF3E7204DD73FE72" box="[151,279,369,395]" italics="true" pageId="12" pageNumber="317">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805DFFB2FEB77207DD1CFE72" box="[286,376,369,395]" pageId="12" pageNumber="317" rank="species">
<emphasis id="84B0EAE7805DFFB2FEB77207DD1CFE72" bold="true" box="[286,376,369,395]" pageId="12" pageNumber="317">sp. nov.</emphasis>
</taxonomicNameLabel>
); and dorsum and sides more boldly patterned, usually rufescent, and with row of dorsolateral white spots (
<emphasis id="84B0EAE7805DFFB2FE8C72E0DD09FE56" box="[293,365,406,431]" italics="true" pageId="12" pageNumber="317">versus</emphasis>
dorsal pattern obscure, brown, and without dorsolateral white spots in
<taxonomicName id="71C44D76805DFFB2FB2D72E0D967FE56" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1156,1283,405,431]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="317" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805DFFB2FB2D72E0D967FE56" box="[1156,1283,405,431]" italics="true" pageId="12" pageNumber="317">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805DFFB2FAA072E3D906FE56" box="[1289,1378,405,431]" pageId="12" pageNumber="317" rank="species">
<emphasis id="84B0EAE7805DFFB2FAA072E3D906FE56" bold="true" box="[1289,1378,405,431]" pageId="12" pageNumber="317">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<figureCitation id="2EFF2A70805DFFB2FAD972E3DCDCFE2A" captionStart="FIGURE 4" captionStartId="14.[151,250,1805,1830]" captionTargetBox="[151,1436,928,1786]" captionTargetId="figure@14.[151,1436,928,1786]" captionTargetPageId="14" captionText="FIGURE 4. Portraits of new Lepidodactylus species in life. (A) holotype of L. mitchelli sp. nov. (UMMZ 244058), (B) holo- type of L. kwasnickae sp. nov. (BPBM 39880), (C) paratype of L. kwasnickae sp. nov. (BPBM 39879), and (D) paratype of L. kwasnickae sp. nov. (BPBM 39151)." httpUri="https://zenodo.org/record/3363253/files/figure.png" pageId="12" pageNumber="317">Fig. 4A</figureCitation>
<emphasis id="84B0EAE7805DFFB2FF1672CCDD63FE2A" box="[191,263,442,467]" italics="true" pageId="12" pageNumber="317">versus</emphasis>
4B, C, D).
</paragraph>
<paragraph id="B67B36F5805DFFB2FF6E72A8DFDBFC16" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="317">
<emphasis id="84B0EAE7805DFFB2FF6E72A8DDB7FE0E" box="[199,467,477,503]" italics="true" pageId="12" pageNumber="317">
Description of
<typeStatus id="697F8857805DFFB2FEDA72ABDDB7FE0E" box="[371,467,477,503]" pageId="12" pageNumber="317" type="holotype">holotype</typeStatus>
</emphasis>
. A mature male of medium size (SVL =
<quantity id="713C9B10805DFFB2FC0772AAD872FE0E" box="[942,1046,476,503]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" pageId="12" pageNumber="317" unit="mm" value="41.5">41.5 mm</quantity>
, TrL =
<quantity id="713C9B10805DFFB2FBC672ABD8BDFE0E" box="[1135,1241,477,503]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="12" pageNumber="317" unit="mm" value="19.3">19.3 mm</quantity>
). Head relatively long (HL/SVL = 0.23) and wide (HW/HL = 0.88), distinct from neck. Loreal region slightly inflated; no distinct canthus rostralis. Top of snout, area between nares, and area posterior to nares concave. Snout tapered and rounded at tip, relatively long (SN/HL = 0.42), longer than eye diameter (SN/EY = 1.6). Eye relatively large (EY/HL = 0.26, EY/EN = 0.71); pupil vertical, constricted into series of four lobes; supraciliaries same size as adjacent granules. Ear opening small (Ear/HL = 0.083), compressed, oriented diagonally from posterodorsal to anteroventral on right, semicircular on left; distance between ear and eye larger than eye diameter (EE/EY = 1.4). Rostral wider (
<quantity id="713C9B10805DFFB2FA9571C3D9F0FD36" box="[1340,1428,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="317" unit="mm" value="1.9">1.9 mm</quantity>
) than high (
<quantity id="713C9B10805DFFB2FEBC71AEDDE6FD0A" box="[277,386,728,755]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="317" unit="mm" value="0.85">0.85 mm</quantity>
), highest just medial to nares, lower between these points; length (
<quantity id="713C9B10805DFFB2FB2571AFD88FFD0A" box="[1164,1259,729,755]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="317" unit="mm" value="0.2">0.2 mm</quantity>
); barely visible from above. Supranasals separated by three internasals. Rostral in contact with first supralabials, two supranasals, and three internasals. External nares circular; each bordered by rostral, two supranasals, first supralabial, and one postnasal. Mental shallowly triangular, wider (
<quantity id="713C9B10805DFFB2FD087033DF6CFCA6" box="[673,776,837,863]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="317" unit="mm" value="0.9">0.90 mm</quantity>
) than long (
<quantity id="713C9B10805DFFB2FC397032DF9EFCA6" box="[912,1018,836,863]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="317" unit="mm" value="0.75">0.75 mm</quantity>
). Mental bordered posteriorly by two enlarged postmentals, these bordered posteriorly by three rows of enlarged gulars that are followed by smaller, granular chin scales. First three infralabials bordered below by somewhat enlarged scales; remaining scales below infralabials of approximately twice size of central throat scales. Supralabials to mid-orbital position ten on each side; to angle of jaw 12 on each side. Infralabials 12 on right, 13 on left.
</paragraph>
<paragraph id="B67B36F5805DFFB2FF6E708FD872FBA2" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="317">Body of modest habitus (TrL/SVL = 0.47), slightly depressed. Dorsal scales on head, body, limbs, and throat tiny, juxtaposed granules, larger on sides and snout; tubercles absent. Ventral scales flatter but somewhat beaded, smooth, subimbricate, gradually decreasing in size laterally to become granular.</paragraph>
<paragraph id="B67B36F5805DFFB2FF6E7713DE77FB12" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="317">Enlarged precloacal scales in single series of 14, each bearing a pore; no enlarged scales on thighs. Enlarged scales form a pubic patch between the precloacal series and vent; no tiny scales between the precloacal series and either side of the pubic patch; ten scales in a row between apex of enlarged precloacal series and vent. Scales on palms and soles rounded, smooth.</paragraph>
<paragraph id="B67B36F5805DFFB2FF6E7783D8CBFA1E" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="317">
Fore- and hindlimbs relatively small but well-developed (FA/SVL = 0.11, CS/SVL = 0.14). Digits well-developed, fairly narrowly dilated throughout their length (T4W/T4L = 0.30), all but first fingers and toes with recurved claws; clawed phalanges laterally compressed, free above and extending slightly beyond terminal scansors. Subdigital lamellae narrow and smooth, all digits with 23 divided subterminal scansors; lamellae extend for most of length of each toe (T4 scansor L/T4L = 0.74). Lamellae of manus
<date id="C27A1035805DFFB2FCDB76F3DF9BFA66" box="[882,1023,1413,1439]" pageId="12" pageNumber="317" value="2014-10-07" valueMax="2014-12-07" valueMin="2014-10-07">7101214</date>
10 on right,
<date id="C27A1035805DFFB2FB3A76F3D979FA66" box="[1171,1309,1413,1439]" pageId="12" pageNumber="317" value="2013-08-11" valueMax="2013-08-13" valueMin="2013-08-11">8111313</date>
11 on left; of pes
<date id="C27A1035805DFFB2FF4876DEDD1DFA3A" box="[225,377,1448,1475]" pageId="12" pageNumber="317" value="2015-11-14" valueMax="2015-11-16" valueMin="2015-11-14">11141615</date>
9 on right,
<date id="C27A1035805DFFB2FDAB76DFDE1BFA3A" box="[514,639,1449,1475]" pageId="12" pageNumber="317" value="2014-09-09" valueMax="2014-09-16" valueMin="2014-09-09">991614</date>
9 on left. Relative lengths of digits on manus and pes I &lt;II &lt;V &lt;III &lt;IV. Webbing present between all digits; toes only one-fifth webbed (T3T4webL/T4L = 0.21).
</paragraph>
<paragraph id="B67B36F5805DFFB2FF6E7687DD24F962" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="317">Tail regenerated behind cloacal sacs, subcylindrical, not as long as body (TL/SVL = 0.95), relatively wide (TW/SVL = 0.11); lateral margins without skin flanges or spines. Scales of tail small, flat, smooth, subimbricate, slightly larger ventrally than dorsally. Cloacal sacs swollen, with small external orifices situated near lateral margins of vent; one small, blunt postcloacal spur on each side of tailbase; midventral scales of sac slightly larger than those ventrolaterally.</paragraph>
<paragraph id="B67B36F5805DFFB2FF6E75D3D9E5F88A" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="317">
<emphasis id="84B0EAE7805DFFB2FF6E75D3DDDEF946" box="[199,442,1701,1727]" italics="true" pageId="12" pageNumber="317">Color in preservative</emphasis>
: Dorsal ground color medium brown, heavily mottled laterally and dorsolaterally with darker brown arrayed into five or six approximate bands divided mid-dorsally; top of head and snout uniformly dark brown. Body with dorsolateral and lateral rows of obscure white dots, each comprising only 12 granules. A short, poorly demarcated, pale postocular stripe present. Tail same color as body but with dark markings arrayed into narrow diagonal lines. Venter white with few scattered brown scales, these denser on chin, under limbs, and on palmar and plantar surfaces; under tail brown. Iris copper colored and heavily veined with dark copper; pupil light gray.
</paragraph>
<paragraph id="B67B36F5805DFFB2FF6E7408DDF1F826" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="317">
<emphasis id="84B0EAE7805DFFB2FF6E7408DDA3F86E" box="[199,455,1917,1943]" italics="true" pageId="12" pageNumber="317">Measurements (in mm)</emphasis>
. SVL = 41.5, TrL = 19.3, TL = 39.5, TW = 4.5, FA = 4.5, CS = 5.8, HL = 9.6, HW = 8.4, Ear = 0.8, EE = 3.4, EY = 2.5, SN = 4.0, EN = 3.5, IN = 2.0, T4L = 4.7, T4W = 1.4, T4 scansor L = 3.5, T3T4webL = 1.0, mass (g) = 1.85.
</paragraph>
<paragraph id="B67B36F5805CFFB3FF6E73ECDE2CFDE2" blockId="13.[151,1437,152,539]" pageId="13" pageNumber="318">
<emphasis id="84B0EAE7805CFFB3FF6E73ECDD4BFF4A" box="[199,303,154,179]" italics="true" pageId="13" pageNumber="318">Variation</emphasis>
. The enlarged precloacal scales vary from 1215, and the number of precloacal pores from 1214. Number of T4 lamellae range from 1220, with individuals varying considerably in the extent to which the proximal scales under the digits are subdivided. Based on the small sample available, females appear to have more lamellae than males (
<tableCitation id="FB46034E805CFFB3FEE07273DDC5FEE6" box="[329,417,261,287]" captionStart="TABLE 2" captionStartId="13.[151,239,586,610]" captionTargetPageId="13" captionText="TABLE 2. Summary meristic and mensural data for type series of ten Lepidodactylus kwasnickae sp. nov.. Backslashes distinguish between right and left sides." httpUri="http://table.plazi.org/id/E2BB667D805CFFB3FF3E713CDE24FD7E" pageId="13" pageNumber="318" tableUuid="E2BB667D805CFFB3FF3E713CDE24FD7E">Table 2</tableCitation>
). Number of divided subterminal scansors varies from 23 and is most often two; divided subterminal scansors are lacking on the first digit of the manus in only BPBM 39151 (both sides) and BPBM 39881 (left side). Lamellae occupy most of the length of each digit (T4 scansor L/T4L = 0.670.93, mean = 0.83). The modal number of supralabials to under the pupil is nine, but two specimens have eight and three have ten. Toes are relatively narrow (T4W/T4L = 0.260.34, mean = 0.31) and have relatively little webbing (T3T4webL/T4L = 0.180.26, mean = 0.22). Otherwise, mensural features show relatively little variation except that females seem to have smaller ears, and the juvenile (BPBM 39150) has a relatively shorter trunk, wider head, wider snout, larger eye, and larger ear than do adults (
<tableCitation id="FB46034E805CFFB3FE4A7177DE5FFDE2" box="[483,571,513,539]" captionStart="TABLE 2" captionStartId="13.[151,239,586,610]" captionTargetPageId="13" captionText="TABLE 2. Summary meristic and mensural data for type series of ten Lepidodactylus kwasnickae sp. nov.. Backslashes distinguish between right and left sides." httpUri="http://table.plazi.org/id/E2BB667D805CFFB3FF3E713CDE24FD7E" pageId="13" pageNumber="318" tableUuid="E2BB667D805CFFB3FF3E713CDE24FD7E">Table 2</tableCitation>
).
</paragraph>
<caption id="E2BB667D805CFFB3FF3E713CDE24FD7E" ID-Table-UUID="E2BB667D805CFFB3FF3E713CDE24FD7E" httpUri="http://table.plazi.org/id/E2BB667D805CFFB3FF3E713CDE24FD7E" pageId="13" pageNumber="318" targetBox="[166,1327,658,1961]" targetIsTable="true" targetPageId="13">
<paragraph id="B67B36F5805CFFB3FF3E713CDE24FD7E" blockId="13.[151,1436,586,647]" pageId="13" pageNumber="318">
<emphasis id="84B0EAE7805CFFB3FF3E713CDD6EFD9A" bold="true" box="[151,266,586,611]" pageId="13" pageNumber="318">TABLE 2.</emphasis>
Summary meristic and mensural data for type series of ten
<taxonomicName id="71C44D76805CFFB3FC22713DD8D4FD9A" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[907,1200,587,611]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="318" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE7805CFFB3FC22713DD8D4FD9A" box="[907,1200,587,611]" italics="true" pageId="13" pageNumber="318">Lepidodactylus kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805CFFB3FB1E713DD968FD9A" box="[1207,1292,587,611]" pageId="13" pageNumber="318" rank="species">
<emphasis id="84B0EAE7805CFFB3FB1E713DD968FD9A" bold="true" box="[1207,1292,587,611]" pageId="13" pageNumber="318">sp. nov.</emphasis>
</taxonomicNameLabel>
. Backslashes distinguish between right and left sides.
</paragraph>
</caption>
<paragraph id="B67B36F5805CFFB3FF0F71E4D970F850" pageId="13" pageNumber="318">
<table id="C4C4C455805C0041FF0F71E4D94BF850" box="[166,1327,658,1961]" colsContinueIn="14.[166,1327,197,903]" gridcols="6" gridrows="34" pageId="13" pageNumber="318">
<tr id="08F434B7805C0041FF0F71E4D94BFD52" box="[166,1327,658,683]" gridrow="0" pageId="13" pageNumber="318" rowspan-2="1" rowspan-4="1">
<th id="4B255DCB805C0041FF0F71E4DD12FD52" box="[166,374,658,683]" gridcol="0" gridrow="0" pageId="13" pageNumber="318">Character</th>
<th id="4B255DCB805C0041FE0871E4DE4EFD52" box="[417,554,658,683]" gridcol="1" gridrow="0" pageId="13" pageNumber="318">Males (n = 3)</th>
<th id="4B255DCB805C0041FC9771E4DFBAFD52" box="[830,990,658,683]" gridcol="3" gridrow="0" pageId="13" pageNumber="318">Females (n = 6)</th>
<th id="4B255DCB805C0041FB7271E4D94BFD52" box="[1243,1327,658,683]" gridcol="5" gridrow="0" pageId="13" pageNumber="318">Juvenile</th>
</tr>
<tr id="08F434B7805C0041FF0F71CFD94BFD28" box="[166,1327,697,721]" gridrow="1" pageId="13" pageNumber="318" rowspan-0="1" rowspan-5="1">
<td id="4B255DCB805C0041FE0871CFDE4EFD28" box="[417,554,697,721]" gridcol="1" gridrow="1" pageId="13" pageNumber="318">Mean</td>
<td id="4B255DCB805C0041FDC671CFDE8AFD28" box="[623,750,697,721]" gridcol="2" gridrow="1" pageId="13" pageNumber="318">Range</td>
<td id="4B255DCB805C0041FC9771CFDFBAFD28" box="[830,990,697,721]" gridcol="3" gridrow="1" pageId="13" pageNumber="318">Mean</td>
<td id="4B255DCB805C0041FBA571CFD8EFFD28" box="[1036,1163,697,721]" gridcol="4" gridrow="1" pageId="13" pageNumber="318">Range</td>
</tr>
<tr id="08F434B7805C0041FF0F71A8D94BFD0E" box="[166,1327,734,759]" gridrow="2" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F71A8DD12FD0E" box="[166,374,734,759]" gridcol="0" gridrow="2" pageId="13" pageNumber="318">mass (g)</th>
<td id="4B255DCB805C0041FE0871A8DE4EFD0E" box="[417,554,734,759]" gridcol="1" gridrow="2" pageId="13" pageNumber="318">2.33</td>
<td id="4B255DCB805C0041FDC671A8DE8AFD0E" box="[623,750,734,759]" gridcol="2" gridrow="2" pageId="13" pageNumber="318">1.852.95</td>
<td id="4B255DCB805C0041FC9771A8DFBAFD0E" box="[830,990,734,759]" gridcol="3" gridrow="2" pageId="13" pageNumber="318">2.38</td>
<td id="4B255DCB805C0041FBA571A8D8EFFD0E" box="[1036,1163,734,759]" gridcol="4" gridrow="2" pageId="13" pageNumber="318">2.002.70</td>
<td id="4B255DCB805C0041FB7271A8D94BFD0E" box="[1243,1327,734,759]" gridcol="5" gridrow="2" pageId="13" pageNumber="318">0.20</td>
</tr>
<tr id="08F434B7805C0041FF0F7072D94BFCE4" box="[166,1327,772,797]" gridrow="3" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7072DD12FCE4" box="[166,374,772,797]" gridcol="0" gridrow="3" pageId="13" pageNumber="318">SVL</th>
<td id="4B255DCB805C0041FE087072DE4EFCE4" box="[417,554,772,797]" gridcol="1" gridrow="3" pageId="13" pageNumber="318">45.7</td>
<td id="4B255DCB805C0041FDC67072DE8AFCE4" box="[623,750,772,797]" gridcol="2" gridrow="3" pageId="13" pageNumber="318">41.549.5</td>
<td id="4B255DCB805C0041FC977072DFBAFCE4" box="[830,990,772,797]" gridcol="3" gridrow="3" pageId="13" pageNumber="318">45.7</td>
<td id="4B255DCB805C0041FBA57072D8EFFCE4" box="[1036,1163,772,797]" gridcol="4" gridrow="3" pageId="13" pageNumber="318">43.547.0</td>
<td id="4B255DCB805C0041FB727072D94BFCE4" box="[1243,1327,772,797]" gridcol="5" gridrow="3" pageId="13" pageNumber="318">18.5</td>
</tr>
<tr id="08F434B7805C0041FF0F705FD94BFCBB" box="[166,1327,809,834]" gridrow="4" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F705FDD12FCBB" box="[166,374,809,834]" gridcol="0" gridrow="4" pageId="13" pageNumber="318">TrL</th>
<td id="4B255DCB805C0041FE08705FDE4EFCBB" box="[417,554,809,834]" gridcol="1" gridrow="4" pageId="13" pageNumber="318">22.5</td>
<td id="4B255DCB805C0041FDC6705FDE8AFCBB" box="[623,750,809,834]" gridcol="2" gridrow="4" pageId="13" pageNumber="318">19.324.3</td>
<td id="4B255DCB805C0041FC97705FDFBAFCBB" box="[830,990,809,834]" gridcol="3" gridrow="4" pageId="13" pageNumber="318">23.3</td>
<td id="4B255DCB805C0041FBA5705FD8EFFCBB" box="[1036,1163,809,834]" gridcol="4" gridrow="4" pageId="13" pageNumber="318">22.324.2</td>
<td id="4B255DCB805C0041FB72705FD94BFCBB" box="[1243,1327,809,834]" gridcol="5" gridrow="4" pageId="13" pageNumber="318">8.4</td>
</tr>
<tr id="08F434B7805C0041FF0F7039D94BFC91" box="[166,1327,847,872]" gridrow="5" pageId="13" pageNumber="318" rowspan-5="1">
<th id="4B255DCB805C0041FF0F7039DD12FC91" box="[166,374,847,872]" gridcol="0" gridrow="5" pageId="13" pageNumber="318">TL</th>
<td id="4B255DCB805C0041FE087039DE4EFC91" box="[417,554,847,872]" gridcol="1" gridrow="5" pageId="13" pageNumber="318">39.5</td>
<td id="4B255DCB805C0041FDC67039DE8AFC91" box="[623,750,847,872]" gridcol="2" gridrow="5" pageId="13" pageNumber="318">39.5</td>
<td id="4B255DCB805C0041FC977039DFBAFC91" box="[830,990,847,872]" gridcol="3" gridrow="5" pageId="13" pageNumber="318">45.3</td>
<td id="4B255DCB805C0041FBA57039D8EFFC91" box="[1036,1163,847,872]" gridcol="4" gridrow="5" pageId="13" pageNumber="318">38.049.5</td>
</tr>
<tr id="08F434B7805C0041FF0F7003D94BFC77" box="[166,1327,885,910]" gridrow="6" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7003DD12FC77" box="[166,374,885,910]" gridcol="0" gridrow="6" pageId="13" pageNumber="318">TW</th>
<td id="4B255DCB805C0041FE087003DE4EFC77" box="[417,554,885,910]" gridcol="1" gridrow="6" pageId="13" pageNumber="318">4.9</td>
<td id="4B255DCB805C0041FDC67003DE8AFC77" box="[623,750,885,910]" gridcol="2" gridrow="6" pageId="13" pageNumber="318">4.55.3</td>
<td id="4B255DCB805C0041FC977003DFBAFC77" box="[830,990,885,910]" gridcol="3" gridrow="6" pageId="13" pageNumber="318">4.7</td>
<td id="4B255DCB805C0041FBA57003D8EFFC77" box="[1036,1163,885,910]" gridcol="4" gridrow="6" pageId="13" pageNumber="318">4.45.0</td>
<td id="4B255DCB805C0041FB727003D94BFC77" box="[1243,1327,885,910]" gridcol="5" gridrow="6" pageId="13" pageNumber="318">1.7</td>
</tr>
<tr id="08F434B7805C0041FF0F70EDD94BFC4D" box="[166,1327,923,948]" gridrow="7" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F70EDDD12FC4D" box="[166,374,923,948]" gridcol="0" gridrow="7" pageId="13" pageNumber="318">CL</th>
<td id="4B255DCB805C0041FE0870EDDE4EFC4D" box="[417,554,923,948]" gridcol="1" gridrow="7" pageId="13" pageNumber="318">6.3</td>
<td id="4B255DCB805C0041FDC670EDDE8AFC4D" box="[623,750,923,948]" gridcol="2" gridrow="7" pageId="13" pageNumber="318">5.87.0</td>
<td id="4B255DCB805C0041FC9770EDDFBAFC4D" box="[830,990,923,948]" gridcol="3" gridrow="7" pageId="13" pageNumber="318">6.2</td>
<td id="4B255DCB805C0041FBA570EDD8EFFC4D" box="[1036,1163,923,948]" gridcol="4" gridrow="7" pageId="13" pageNumber="318">5.66.6</td>
<td id="4B255DCB805C0041FB7270EDD94BFC4D" box="[1243,1327,923,948]" gridcol="5" gridrow="7" pageId="13" pageNumber="318">2.6</td>
</tr>
<tr id="08F434B7805C0041FF0F70B7D94BFC23" box="[166,1327,961,986]" gridrow="8" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F70B7DD12FC23" box="[166,374,961,986]" gridcol="0" gridrow="8" pageId="13" pageNumber="318">FL</th>
<td id="4B255DCB805C0041FE0870B7DE4EFC23" box="[417,554,961,986]" gridcol="1" gridrow="8" pageId="13" pageNumber="318">5.0</td>
<td id="4B255DCB805C0041FDC670B7DE8AFC23" box="[623,750,961,986]" gridcol="2" gridrow="8" pageId="13" pageNumber="318">4.55.3</td>
<td id="4B255DCB805C0041FC9770B7DFBAFC23" box="[830,990,961,986]" gridcol="3" gridrow="8" pageId="13" pageNumber="318">4.9</td>
<td id="4B255DCB805C0041FBA570B7D8EFFC23" box="[1036,1163,961,986]" gridcol="4" gridrow="8" pageId="13" pageNumber="318">4.75.2</td>
<td id="4B255DCB805C0041FB7270B7D94BFC23" box="[1243,1327,961,986]" gridcol="5" gridrow="8" pageId="13" pageNumber="318">2.2</td>
</tr>
<tr id="08F434B7805C0041FF0F7090D94BFC06" box="[166,1327,998,1023]" gridrow="9" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7090DD12FC06" box="[166,374,998,1023]" gridcol="0" gridrow="9" pageId="13" pageNumber="318">HL</th>
<td id="4B255DCB805C0041FE087090DE4EFC06" box="[417,554,998,1023]" gridcol="1" gridrow="9" pageId="13" pageNumber="318">10.5</td>
<td id="4B255DCB805C0041FDC67090DE8AFC06" box="[623,750,998,1023]" gridcol="2" gridrow="9" pageId="13" pageNumber="318">9.611.5</td>
<td id="4B255DCB805C0041FC977090DFBAFC06" box="[830,990,998,1023]" gridcol="3" gridrow="9" pageId="13" pageNumber="318">10.6</td>
<td id="4B255DCB805C0041FBA57090D8EFFC06" box="[1036,1163,998,1023]" gridcol="4" gridrow="9" pageId="13" pageNumber="318">9.810.8</td>
<td id="4B255DCB805C0041FB727090D94BFC06" box="[1243,1327,998,1023]" gridcol="5" gridrow="9" pageId="13" pageNumber="318">4.8</td>
</tr>
<tr id="08F434B7805C0041FF0F777BD94BFBDC" box="[166,1327,1037,1061]" gridrow="10" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F777BDD12FBDC" box="[166,374,1037,1061]" gridcol="0" gridrow="10" pageId="13" pageNumber="318">HW</th>
<td id="4B255DCB805C0041FE08777BDE4EFBDC" box="[417,554,1037,1061]" gridcol="1" gridrow="10" pageId="13" pageNumber="318">9.4</td>
<td id="4B255DCB805C0041FDC6777BDE8AFBDC" box="[623,750,1037,1061]" gridcol="2" gridrow="10" pageId="13" pageNumber="318">8.410.1</td>
<td id="4B255DCB805C0041FC97777BDFBAFBDC" box="[830,990,1037,1061]" gridcol="3" gridrow="10" pageId="13" pageNumber="318">9.0</td>
<td id="4B255DCB805C0041FBA5777BD8EFFBDC" box="[1036,1163,1037,1061]" gridcol="4" gridrow="10" pageId="13" pageNumber="318">8.79.3</td>
<td id="4B255DCB805C0041FB72777BD94BFBDC" box="[1243,1327,1037,1061]" gridcol="5" gridrow="10" pageId="13" pageNumber="318">4.3</td>
</tr>
<tr id="08F434B7805C0041FF0F7744D94BFBB2" box="[166,1327,1074,1099]" gridrow="11" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7744DD12FBB2" box="[166,374,1074,1099]" gridcol="0" gridrow="11" pageId="13" pageNumber="318">EY</th>
<td id="4B255DCB805C0041FE087744DE4EFBB2" box="[417,554,1074,1099]" gridcol="1" gridrow="11" pageId="13" pageNumber="318">2.6</td>
<td id="4B255DCB805C0041FDC67744DE8AFBB2" box="[623,750,1074,1099]" gridcol="2" gridrow="11" pageId="13" pageNumber="318">2.52.7</td>
<td id="4B255DCB805C0041FC977744DFBAFBB2" box="[830,990,1074,1099]" gridcol="3" gridrow="11" pageId="13" pageNumber="318">2.6</td>
<td id="4B255DCB805C0041FBA57744D8EFFBB2" box="[1036,1163,1074,1099]" gridcol="4" gridrow="11" pageId="13" pageNumber="318">2.42.7</td>
<td id="4B255DCB805C0041FB727744D94BFBB2" box="[1243,1327,1074,1099]" gridcol="5" gridrow="11" pageId="13" pageNumber="318">1.6</td>
</tr>
<tr id="08F434B7805C0041FF0F772ED94BFB88" box="[166,1327,1112,1137]" gridrow="12" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F772EDD12FB88" box="[166,374,1112,1137]" gridcol="0" gridrow="12" pageId="13" pageNumber="318">EN</th>
<td id="4B255DCB805C0041FE08772EDE4EFB88" box="[417,554,1112,1137]" gridcol="1" gridrow="12" pageId="13" pageNumber="318">3.8</td>
<td id="4B255DCB805C0041FDC6772EDE8AFB88" box="[623,750,1112,1137]" gridcol="2" gridrow="12" pageId="13" pageNumber="318">3.54.1</td>
<td id="4B255DCB805C0041FC97772EDFBAFB88" box="[830,990,1112,1137]" gridcol="3" gridrow="12" pageId="13" pageNumber="318">3.7</td>
<td id="4B255DCB805C0041FBA5772ED8EFFB88" box="[1036,1163,1112,1137]" gridcol="4" gridrow="12" pageId="13" pageNumber="318">3.33.9</td>
<td id="4B255DCB805C0041FB72772ED94BFB88" box="[1243,1327,1112,1137]" gridcol="5" gridrow="12" pageId="13" pageNumber="318">1.5</td>
</tr>
<tr id="08F434B7805C0041FF0F7708D94BFB6F" box="[166,1327,1150,1174]" gridrow="13" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7708DD12FB6F" box="[166,374,1150,1174]" gridcol="0" gridrow="13" pageId="13" pageNumber="318">IN</th>
<td id="4B255DCB805C0041FE087708DE4EFB6F" box="[417,554,1150,1174]" gridcol="1" gridrow="13" pageId="13" pageNumber="318">2.2</td>
<td id="4B255DCB805C0041FDC67708DE8AFB6F" box="[623,750,1150,1174]" gridcol="2" gridrow="13" pageId="13" pageNumber="318">2.02.4</td>
<td id="4B255DCB805C0041FC977708DFBAFB6F" box="[830,990,1150,1174]" gridcol="3" gridrow="13" pageId="13" pageNumber="318">2.3</td>
<td id="4B255DCB805C0041FBA57708D8EFFB6F" box="[1036,1163,1150,1174]" gridcol="4" gridrow="13" pageId="13" pageNumber="318">2.22.3</td>
<td id="4B255DCB805C0041FB727708D94BFB6F" box="[1243,1327,1150,1174]" gridcol="5" gridrow="13" pageId="13" pageNumber="318">1.2</td>
</tr>
<tr id="08F434B7805C0041FF0F77D5D94BFB45" box="[166,1327,1187,1212]" gridrow="14" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F77D5DD12FB45" box="[166,374,1187,1212]" gridcol="0" gridrow="14" pageId="13" pageNumber="318">SN</th>
<td id="4B255DCB805C0041FE0877D5DE4EFB45" box="[417,554,1187,1212]" gridcol="1" gridrow="14" pageId="13" pageNumber="318">4.4</td>
<td id="4B255DCB805C0041FDC677D5DE8AFB45" box="[623,750,1187,1212]" gridcol="2" gridrow="14" pageId="13" pageNumber="318">4.04.8</td>
<td id="4B255DCB805C0041FC9777D5DFBAFB45" box="[830,990,1187,1212]" gridcol="3" gridrow="14" pageId="13" pageNumber="318">4.4</td>
<td id="4B255DCB805C0041FBA577D5D8EFFB45" box="[1036,1163,1187,1212]" gridcol="4" gridrow="14" pageId="13" pageNumber="318">4.14.6</td>
<td id="4B255DCB805C0041FB7277D5D94BFB45" box="[1243,1327,1187,1212]" gridcol="5" gridrow="14" pageId="13" pageNumber="318">1.8</td>
</tr>
<tr id="08F434B7805C0041FF0F77BFD94BFB1B" box="[166,1327,1225,1250]" gridrow="15" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F77BFDD12FB1B" box="[166,374,1225,1250]" gridcol="0" gridrow="15" pageId="13" pageNumber="318">Ear</th>
<td id="4B255DCB805C0041FE0877BFDE4EFB1B" box="[417,554,1225,1250]" gridcol="1" gridrow="15" pageId="13" pageNumber="318">0.8</td>
<td id="4B255DCB805C0041FDC677BFDE8AFB1B" box="[623,750,1225,1250]" gridcol="2" gridrow="15" pageId="13" pageNumber="318">0.80.9</td>
<td id="4B255DCB805C0041FC9777BFDFBAFB1B" box="[830,990,1225,1250]" gridcol="3" gridrow="15" pageId="13" pageNumber="318">0.7</td>
<td id="4B255DCB805C0041FBA577BFD8EFFB1B" box="[1036,1163,1225,1250]" gridcol="4" gridrow="15" pageId="13" pageNumber="318">0.60.7</td>
<td id="4B255DCB805C0041FB7277BFD94BFB1B" box="[1243,1327,1225,1250]" gridcol="5" gridrow="15" pageId="13" pageNumber="318">0.4</td>
</tr>
<tr id="08F434B7805C0041FF0F7799D94BFAF1" box="[166,1327,1263,1288]" gridrow="16" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7799DD12FAF1" box="[166,374,1263,1288]" gridcol="0" gridrow="16" pageId="13" pageNumber="318">EE</th>
<td id="4B255DCB805C0041FE087799DE4EFAF1" box="[417,554,1263,1288]" gridcol="1" gridrow="16" pageId="13" pageNumber="318">3.8</td>
<td id="4B255DCB805C0041FDC67799DE8AFAF1" box="[623,750,1263,1288]" gridcol="2" gridrow="16" pageId="13" pageNumber="318">3.44.0</td>
<td id="4B255DCB805C0041FC977799DFBAFAF1" box="[830,990,1263,1288]" gridcol="3" gridrow="16" pageId="13" pageNumber="318">3.8</td>
<td id="4B255DCB805C0041FBA57799D8EFFAF1" box="[1036,1163,1263,1288]" gridcol="4" gridrow="16" pageId="13" pageNumber="318">3.63.9</td>
<td id="4B255DCB805C0041FB727799D94BFAF1" box="[1243,1327,1263,1288]" gridcol="5" gridrow="16" pageId="13" pageNumber="318">1.7</td>
</tr>
<tr id="08F434B7805C0041FF0F7663D94BFAD7" box="[166,1327,1301,1326]" gridrow="17" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7663DD12FAD7" box="[166,374,1301,1326]" gridcol="0" gridrow="17" pageId="13" pageNumber="318">T4L</th>
<td id="4B255DCB805C0041FE087663DE4EFAD7" box="[417,554,1301,1326]" gridcol="1" gridrow="17" pageId="13" pageNumber="318">5.2</td>
<td id="4B255DCB805C0041FDC67663DE8AFAD7" box="[623,750,1301,1326]" gridcol="2" gridrow="17" pageId="13" pageNumber="318">4.75.6</td>
<td id="4B255DCB805C0041FC977663DFBAFAD7" box="[830,990,1301,1326]" gridcol="3" gridrow="17" pageId="13" pageNumber="318">5.4</td>
<td id="4B255DCB805C0041FBA57663D8EFFAD7" box="[1036,1163,1301,1326]" gridcol="4" gridrow="17" pageId="13" pageNumber="318">5.05.8</td>
<td id="4B255DCB805C0041FB727663D94BFAD7" box="[1243,1327,1301,1326]" gridcol="5" gridrow="17" pageId="13" pageNumber="318">2.10</td>
</tr>
<tr id="08F434B7805C0041FF0F764DD94BFAAD" box="[166,1327,1339,1364]" gridrow="18" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F764DDD12FAAD" box="[166,374,1339,1364]" gridcol="0" gridrow="18" pageId="13" pageNumber="318">T4 width</th>
<td id="4B255DCB805C0041FE08764DDE4EFAAD" box="[417,554,1339,1364]" gridcol="1" gridrow="18" pageId="13" pageNumber="318">1.6</td>
<td id="4B255DCB805C0041FDC6764DDE8AFAAD" box="[623,750,1339,1364]" gridcol="2" gridrow="18" pageId="13" pageNumber="318">1.41.8</td>
<td id="4B255DCB805C0041FC97764DDFBAFAAD" box="[830,990,1339,1364]" gridcol="3" gridrow="18" pageId="13" pageNumber="318">1.6</td>
<td id="4B255DCB805C0041FBA5764DD8EFFAAD" box="[1036,1163,1339,1364]" gridcol="4" gridrow="18" pageId="13" pageNumber="318">1.51.7</td>
<td id="4B255DCB805C0041FB72764DD94BFAAD" box="[1243,1327,1339,1364]" gridcol="5" gridrow="18" pageId="13" pageNumber="318">0.70</td>
</tr>
<tr id="08F434B7805C0041FF0F7616D94BFA80" box="[166,1327,1376,1401]" gridrow="19" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7616DD12FA80" box="[166,374,1376,1401]" gridcol="0" gridrow="19" pageId="13" pageNumber="318">T3T4 web L</th>
<td id="4B255DCB805C0041FE087616DE4EFA80" box="[417,554,1376,1401]" gridcol="1" gridrow="19" pageId="13" pageNumber="318">1.2</td>
<td id="4B255DCB805C0041FDC67616DE8AFA80" box="[623,750,1376,1401]" gridcol="2" gridrow="19" pageId="13" pageNumber="318">1.01.4</td>
<td id="4B255DCB805C0041FC977616DFBAFA80" box="[830,990,1376,1401]" gridcol="3" gridrow="19" pageId="13" pageNumber="318">1.1</td>
<td id="4B255DCB805C0041FBA57616D8EFFA80" box="[1036,1163,1376,1401]" gridcol="4" gridrow="19" pageId="13" pageNumber="318">1.01.3</td>
<td id="4B255DCB805C0041FB727616D94BFA80" box="[1243,1327,1376,1401]" gridcol="5" gridrow="19" pageId="13" pageNumber="318">0.40</td>
</tr>
<tr id="08F434B7805C0041FF0F76F0D94BFA66" box="[166,1327,1414,1439]" gridrow="20" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F76F0DD12FA66" box="[166,374,1414,1439]" gridcol="0" gridrow="20" pageId="13" pageNumber="318">T4scansor L</th>
<td id="4B255DCB805C0041FE0876F0DE4EFA66" box="[417,554,1414,1439]" gridcol="1" gridrow="20" pageId="13" pageNumber="318">4.1</td>
<td id="4B255DCB805C0041FDC676F0DE8AFA66" box="[623,750,1414,1439]" gridcol="2" gridrow="20" pageId="13" pageNumber="318">3.54.6</td>
<td id="4B255DCB805C0041FC9776F0DFBAFA66" box="[830,990,1414,1439]" gridcol="3" gridrow="20" pageId="13" pageNumber="318">4.6</td>
<td id="4B255DCB805C0041FBA576F0D8EFFA66" box="[1036,1163,1414,1439]" gridcol="4" gridrow="20" pageId="13" pageNumber="318">3.95.3</td>
<td id="4B255DCB805C0041FB7276F0D94BFA66" box="[1243,1327,1414,1439]" gridcol="5" gridrow="20" pageId="13" pageNumber="318">1.80</td>
</tr>
<tr id="08F434B7805C0041FF0F76DAD94BFA3C" box="[166,1327,1452,1477]" gridrow="21" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F76DADD12FA3C" box="[166,374,1452,1477]" gridcol="0" gridrow="21" pageId="13" pageNumber="318">T4 scansors</th>
<td id="4B255DCB805C0041FE0876DADE4EFA3C" box="[417,554,1452,1477]" gridcol="1" gridrow="21" pageId="13" pageNumber="318">14.7</td>
<td id="4B255DCB805C0041FDC676DADE8AFA3C" box="[623,750,1452,1477]" gridcol="2" gridrow="21" pageId="13" pageNumber="318">1216</td>
<td id="4B255DCB805C0041FC9776DADFBAFA3C" box="[830,990,1452,1477]" gridcol="3" gridrow="21" pageId="13" pageNumber="318">17.1</td>
<td id="4B255DCB805C0041FBA576DAD8EFFA3C" box="[1036,1163,1452,1477]" gridcol="4" gridrow="21" pageId="13" pageNumber="318">1520</td>
<td id="4B255DCB805C0041FB7276DAD94BFA3C" box="[1243,1327,1452,1477]" gridcol="5" gridrow="21" pageId="13" pageNumber="318">15/14</td>
</tr>
<tr id="08F434B7805C0041FF0F76A4D94BFA12" box="[166,1327,1490,1515]" gridrow="22" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F76A4DD12FA12" box="[166,374,1490,1515]" gridcol="0" gridrow="22" pageId="13" pageNumber="318">Divided T4 scansors</th>
<td id="4B255DCB805C0041FE0876A4DE4EFA12" box="[417,554,1490,1515]" gridcol="1" gridrow="22" pageId="13" pageNumber="318">2.5</td>
<td id="4B255DCB805C0041FDC676A4DE8AFA12" box="[623,750,1490,1515]" gridcol="2" gridrow="22" pageId="13" pageNumber="318">23</td>
<td id="4B255DCB805C0041FC9776A4DFBAFA12" box="[830,990,1490,1515]" gridcol="3" gridrow="22" pageId="13" pageNumber="318">2.3</td>
<td id="4B255DCB805C0041FBA576A4D8EFFA12" box="[1036,1163,1490,1515]" gridcol="4" gridrow="22" pageId="13" pageNumber="318">23</td>
<td id="4B255DCB805C0041FB7276A4D94BFA12" box="[1243,1327,1490,1515]" gridcol="5" gridrow="22" pageId="13" pageNumber="318">2/2</td>
</tr>
<tr id="08F434B7805C0041FF0F768ED94BF9E8" box="[166,1327,1528,1553]" gridrow="23" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F768EDD12F9E8" box="[166,374,1528,1553]" gridcol="0" gridrow="23" pageId="13" pageNumber="318">Enlarged precloacal</th>
<td id="4B255DCB805C0041FE08768EDE4EF9E8" box="[417,554,1528,1553]" gridcol="1" gridrow="23" pageId="13" pageNumber="318">13.3</td>
<td id="4B255DCB805C0041FDC6768EDE8AF9E8" box="[623,750,1528,1553]" gridcol="2" gridrow="23" pageId="13" pageNumber="318">1314</td>
<td id="4B255DCB805C0041FC97768EDFBAF9E8" box="[830,990,1528,1553]" gridcol="3" gridrow="23" pageId="13" pageNumber="318">13.5</td>
<td id="4B255DCB805C0041FBA5768ED8EFF9E8" box="[1036,1163,1528,1553]" gridcol="4" gridrow="23" pageId="13" pageNumber="318">1215</td>
<td id="4B255DCB805C0041FB72768ED94BF9E8" box="[1243,1327,1528,1553]" gridcol="5" gridrow="23" pageId="13" pageNumber="318">13</td>
</tr>
<tr id="08F434B7805C0041FF0F756AD94BF9AC" box="[166,1327,1564,1621]" gridrow="24" pageId="13" pageNumber="318" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="4B255DCB805C0041FF0F756ADD12F9AC" box="[166,374,1564,1621]" gridcol="0" gridrow="24" pageId="13" pageNumber="318">scales Precloacal pores</th>
<td id="4B255DCB805C0041FE08756ADE4EF9AC" box="[417,554,1564,1621]" gridcol="1" gridrow="24" pageId="13" pageNumber="318">13.0</td>
<td id="4B255DCB805C0041FDC6756ADE8AF9AC" box="[623,750,1564,1621]" gridcol="2" gridrow="24" pageId="13" pageNumber="318">1214</td>
</tr>
<tr id="08F434B7805C0041FF0F7514D94BF982" box="[166,1327,1634,1659]" gridrow="25" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7514DD12F982" box="[166,374,1634,1659]" gridcol="0" gridrow="25" pageId="13" pageNumber="318">Rows cloacal scales</th>
<td id="4B255DCB805C0041FE087514DE4EF982" box="[417,554,1634,1659]" gridcol="1" gridrow="25" pageId="13" pageNumber="318">9.7</td>
<td id="4B255DCB805C0041FDC67514DE8AF982" box="[623,750,1634,1659]" gridcol="2" gridrow="25" pageId="13" pageNumber="318">910</td>
<td id="4B255DCB805C0041FC977514DFBAF982" box="[830,990,1634,1659]" gridcol="3" gridrow="25" pageId="13" pageNumber="318">9</td>
<td id="4B255DCB805C0041FBA57514D8EFF982" box="[1036,1163,1634,1659]" gridcol="4" gridrow="25" pageId="13" pageNumber="318">9</td>
<td id="4B255DCB805C0041FB727514D94BF982" box="[1243,1327,1634,1659]" gridcol="5" gridrow="25" pageId="13" pageNumber="318">8</td>
</tr>
<tr id="08F434B7805C0041FF0F75FED94BF958" box="[166,1327,1672,1697]" gridrow="26" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F75FEDD12F958" box="[166,374,1672,1697]" gridcol="0" gridrow="26" pageId="13" pageNumber="318">TrunkL/SVL</th>
<td id="4B255DCB805C0041FE0875FEDE4EF958" box="[417,554,1672,1697]" gridcol="1" gridrow="26" pageId="13" pageNumber="318">0.49</td>
<td id="4B255DCB805C0041FDC675FEDE8AF958" box="[623,750,1672,1697]" gridcol="2" gridrow="26" pageId="13" pageNumber="318">0.470.52</td>
<td id="4B255DCB805C0041FC9775FEDFBAF958" box="[830,990,1672,1697]" gridcol="3" gridrow="26" pageId="13" pageNumber="318">0.51</td>
<td id="4B255DCB805C0041FBA575FED8EFF958" box="[1036,1163,1672,1697]" gridcol="4" gridrow="26" pageId="13" pageNumber="318">0.490.53</td>
<td id="4B255DCB805C0041FB7275FED94BF958" box="[1243,1327,1672,1697]" gridcol="5" gridrow="26" pageId="13" pageNumber="318">0.45</td>
</tr>
<tr id="08F434B7805C0041FF0F75D8D94BF93E" box="[166,1327,1710,1735]" gridrow="27" pageId="13" pageNumber="318" rowspan-5="1">
<th id="4B255DCB805C0041FF0F75D8DD12F93E" box="[166,374,1710,1735]" gridcol="0" gridrow="27" pageId="13" pageNumber="318">TailL/SVL</th>
<td id="4B255DCB805C0041FE0875D8DE4EF93E" box="[417,554,1710,1735]" gridcol="1" gridrow="27" pageId="13" pageNumber="318">0.95</td>
<td id="4B255DCB805C0041FDC675D8DE8AF93E" box="[623,750,1710,1735]" gridcol="2" gridrow="27" pageId="13" pageNumber="318">0.95</td>
<td id="4B255DCB805C0041FC9775D8DFBAF93E" box="[830,990,1710,1735]" gridcol="3" gridrow="27" pageId="13" pageNumber="318">0.98</td>
<td id="4B255DCB805C0041FBA575D8D8EFF93E" box="[1036,1163,1710,1735]" gridcol="4" gridrow="27" pageId="13" pageNumber="318">0.0830.11</td>
</tr>
<tr id="08F434B7805C0041FF0F75A5D94BF915" box="[166,1327,1747,1772]" gridrow="28" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F75A5DD12F915" box="[166,374,1747,1772]" gridcol="0" gridrow="28" pageId="13" pageNumber="318">TailW/SVL</th>
<td id="4B255DCB805C0041FE0875A5DE4EF915" box="[417,554,1747,1772]" gridcol="1" gridrow="28" pageId="13" pageNumber="318">0.11</td>
<td id="4B255DCB805C0041FDC675A5DE8AF915" box="[623,750,1747,1772]" gridcol="2" gridrow="28" pageId="13" pageNumber="318">0.11</td>
<td id="4B255DCB805C0041FC9775A5DFBAF915" box="[830,990,1747,1772]" gridcol="3" gridrow="28" pageId="13" pageNumber="318">0.10</td>
<td id="4B255DCB805C0041FBA575A5D8EFF915" box="[1036,1163,1747,1772]" gridcol="4" gridrow="28" pageId="13" pageNumber="318">0.100.11</td>
<td id="4B255DCB805C0041FB7275A5D94BF915" box="[1243,1327,1747,1772]" gridcol="5" gridrow="28" pageId="13" pageNumber="318">0.09</td>
</tr>
<tr id="08F434B7805C0041FF0F758CD94BF8EB" box="[166,1327,1786,1810]" gridrow="29" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F758CDD12F8EB" box="[166,374,1786,1810]" gridcol="0" gridrow="29" pageId="13" pageNumber="318">CrusL/SVL</th>
<td id="4B255DCB805C0041FE08758CDE4EF8EB" box="[417,554,1786,1810]" gridcol="1" gridrow="29" pageId="13" pageNumber="318">0.14</td>
<td id="4B255DCB805C0041FDC6758CDE8AF8EB" box="[623,750,1786,1810]" gridcol="2" gridrow="29" pageId="13" pageNumber="318">0.130.14</td>
<td id="4B255DCB805C0041FC97758CDFBAF8EB" box="[830,990,1786,1810]" gridcol="3" gridrow="29" pageId="13" pageNumber="318">0.14</td>
<td id="4B255DCB805C0041FBA5758CD8EFF8EB" box="[1036,1163,1786,1810]" gridcol="4" gridrow="29" pageId="13" pageNumber="318">0.130.14</td>
<td id="4B255DCB805C0041FB72758CD94BF8EB" box="[1243,1327,1786,1810]" gridcol="5" gridrow="29" pageId="13" pageNumber="318">0.14</td>
</tr>
<tr id="08F434B7805C0041FF0F7456D94BF8C1" box="[166,1327,1824,1848]" gridrow="30" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7456DD12F8C1" box="[166,374,1824,1848]" gridcol="0" gridrow="30" pageId="13" pageNumber="318">ForearmL/SVL</th>
<td id="4B255DCB805C0041FE087456DE4EF8C1" box="[417,554,1824,1848]" gridcol="1" gridrow="30" pageId="13" pageNumber="318">0.11</td>
<td id="4B255DCB805C0041FDC67456DE8AF8C1" box="[623,750,1824,1848]" gridcol="2" gridrow="30" pageId="13" pageNumber="318">0.11</td>
<td id="4B255DCB805C0041FC977456DFBAF8C1" box="[830,990,1824,1848]" gridcol="3" gridrow="30" pageId="13" pageNumber="318">0.11</td>
<td id="4B255DCB805C0041FBA57456D8EFF8C1" box="[1036,1163,1824,1848]" gridcol="4" gridrow="30" pageId="13" pageNumber="318">0.100.11</td>
<td id="4B255DCB805C0041FB727456D94BF8C1" box="[1243,1327,1824,1848]" gridcol="5" gridrow="30" pageId="13" pageNumber="318">0.12</td>
</tr>
<tr id="08F434B7805C0041FF0F7433D94BF8A7" box="[166,1327,1861,1886]" gridrow="31" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F7433DD12F8A7" box="[166,374,1861,1886]" gridcol="0" gridrow="31" pageId="13" pageNumber="318">HL/SVL</th>
<td id="4B255DCB805C0041FE087433DE4EF8A7" box="[417,554,1861,1886]" gridcol="1" gridrow="31" pageId="13" pageNumber="318">0.23</td>
<td id="4B255DCB805C0041FDC67433DE8AF8A7" box="[623,750,1861,1886]" gridcol="2" gridrow="31" pageId="13" pageNumber="318">0.23</td>
<td id="4B255DCB805C0041FC977433DFBAF8A7" box="[830,990,1861,1886]" gridcol="3" gridrow="31" pageId="13" pageNumber="318">0.23</td>
<td id="4B255DCB805C0041FBA57433D8EFF8A7" box="[1036,1163,1861,1886]" gridcol="4" gridrow="31" pageId="13" pageNumber="318">0.230.24</td>
<td id="4B255DCB805C0041FB727433D94BF8A7" box="[1243,1327,1861,1886]" gridcol="5" gridrow="31" pageId="13" pageNumber="318">0.26</td>
</tr>
<tr id="08F434B7805C0041FF0F741DD94BF87A" box="[166,1327,1899,1923]" gridrow="32" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F741DDD12F87A" box="[166,374,1899,1923]" gridcol="0" gridrow="32" pageId="13" pageNumber="318">HW/SVL</th>
<td id="4B255DCB805C0041FE08741DDE4EF87A" box="[417,554,1899,1923]" gridcol="1" gridrow="32" pageId="13" pageNumber="318">0.21</td>
<td id="4B255DCB805C0041FDC6741DDE8AF87A" box="[623,750,1899,1923]" gridcol="2" gridrow="32" pageId="13" pageNumber="318">0.200.21</td>
<td id="4B255DCB805C0041FC97741DDFBAF87A" box="[830,990,1899,1923]" gridcol="3" gridrow="32" pageId="13" pageNumber="318">0.20</td>
<td id="4B255DCB805C0041FBA5741DD8EFF87A" box="[1036,1163,1899,1923]" gridcol="4" gridrow="32" pageId="13" pageNumber="318">0.190.20</td>
<td id="4B255DCB805C0041FB72741DD94BF87A" box="[1243,1327,1899,1923]" gridcol="5" gridrow="32" pageId="13" pageNumber="318">0.23</td>
</tr>
<tr id="08F434B7805C0041FF0F74E6D94BF850" box="[166,1327,1936,1961]" gridrow="33" pageId="13" pageNumber="318">
<th id="4B255DCB805C0041FF0F74E6DD12F850" box="[166,374,1936,1961]" gridcol="0" gridrow="33" pageId="13" pageNumber="318">EY/SVL</th>
<td id="4B255DCB805C0041FE0874E6DE4EF850" box="[417,554,1936,1961]" gridcol="1" gridrow="33" pageId="13" pageNumber="318">0.057</td>
<td id="4B255DCB805C0041FDC674E6DE8AF850" box="[623,750,1936,1961]" gridcol="2" gridrow="33" pageId="13" pageNumber="318">0.0550.060</td>
<td id="4B255DCB805C0041FC9774E6DFBAF850" box="[830,990,1936,1961]" gridcol="3" gridrow="33" pageId="13" pageNumber="318">0.056</td>
<td id="4B255DCB805C0041FBA574E6D8EFF850" box="[1036,1163,1936,1961]" gridcol="4" gridrow="33" pageId="13" pageNumber="318">0.0520.060</td>
<td id="4B255DCB805C0041FB7274E6D94BF850" box="[1243,1327,1936,1961]" gridcol="5" gridrow="33" pageId="13" pageNumber="318">0.086</td>
</tr>
</table>
</paragraph>
<paragraph id="B67B36F5805CFFB3FBF074CED9EBF836" blockId="13.[1113,1423,1976,2000]" box="[1113,1423,1976,2000]" pageId="13" pageNumber="318">
<tableNote id="4B22377B805CFFB3FBF074CED9EBF836" box="[1113,1423,1976,2000]" pageId="13" pageNumber="318" targetBox="[166,1327,658,1961]" targetPageId="13">......continued on the next page</tableNote>
</paragraph>
<caption id="E2BB667D805FFFB0FF0F73D6DDF0FF40" ID-Table-UUID="E2BB667D805FFFB0FF0F73D6DDF0FF40" box="[166,404,160,185]" httpUri="http://table.plazi.org/id/E2BB667D805FFFB0FF0F73D6DDF0FF40" pageId="14" pageNumber="319" startId="14.[166,250,160,185]" targetBox="[166,1327,197,903]" targetIsTable="true" targetPageId="14">
<paragraph id="B67B36F5805FFFB0FF0F73D6DDF0FF40" blockId="14.[166,404,160,185]" box="[166,404,160,185]" pageId="14" pageNumber="319">
<emphasis id="84B0EAE7805FFFB0FF0F73D6DD77FF40" bold="true" box="[166,275,160,185]" pageId="14" pageNumber="319">TABLE 2.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="B67B36F5805FFFB0FF0F73B3D963FC7E" pageId="14" pageNumber="319">
<table id="C4C4C455805F0041FF0F73B3D94BFC7E" box="[166,1327,197,903]" gridcols="6" gridrows="19" pageId="14" pageNumber="319">
<tr id="08F434B7805F0041FF0F73B3D94BFF27" box="[166,1327,197,222]" gridrow="0" pageId="14" pageNumber="319" rowspan-2="1" rowspan-4="1">
<th id="4B255DCB805F0041FF0F73B3DD3DFF27" box="[166,345,197,222]" gridcol="0" gridrow="0" pageId="14" pageNumber="319">Character</th>
<th id="4B255DCB805F0041FE0873B3DE4EFF27" box="[417,554,197,222]" gridcol="1" gridrow="0" pageId="14" pageNumber="319">Males (n = 3)</th>
<th id="4B255DCB805F0041FC9773B3DFBAFF27" box="[830,990,197,222]" gridcol="3" gridrow="0" pageId="14" pageNumber="319">Females (n = 6)</th>
<th id="4B255DCB805F0041FB7273B3D94BFF27" box="[1243,1327,197,222]" gridcol="5" gridrow="0" pageId="14" pageNumber="319">Juvenile</th>
</tr>
<tr id="08F434B7805F0041FF0F739AD94BFEFD" box="[166,1327,236,260]" gridrow="1" pageId="14" pageNumber="319" rowspan-0="1" rowspan-5="1">
<td id="4B255DCB805F0041FE08739ADE4EFEFD" box="[417,554,236,260]" gridcol="1" gridrow="1" pageId="14" pageNumber="319">Mean</td>
<td id="4B255DCB805F0041FDC6739ADE8AFEFD" box="[623,750,236,260]" gridcol="2" gridrow="1" pageId="14" pageNumber="319">Range</td>
<td id="4B255DCB805F0041FC97739ADFBAFEFD" box="[830,990,236,260]" gridcol="3" gridrow="1" pageId="14" pageNumber="319">Mean</td>
<td id="4B255DCB805F0041FBA5739AD8EFFEFD" box="[1036,1163,236,260]" gridcol="4" gridrow="1" pageId="14" pageNumber="319">Range</td>
</tr>
<tr id="08F434B7805F0041FF0F7264D94BFED2" box="[166,1327,274,299]" gridrow="2" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F7264DD3DFED2" box="[166,345,274,299]" gridcol="0" gridrow="2" pageId="14" pageNumber="319">EN/SVL</th>
<td id="4B255DCB805F0041FE087264DE4EFED2" box="[417,554,274,299]" gridcol="1" gridrow="2" pageId="14" pageNumber="319">0.083</td>
<td id="4B255DCB805F0041FDC67264DE8AFED2" box="[623,750,274,299]" gridcol="2" gridrow="2" pageId="14" pageNumber="319">0.0800.084</td>
<td id="4B255DCB805F0041FC977264DFBAFED2" box="[830,990,274,299]" gridcol="3" gridrow="2" pageId="14" pageNumber="319">0.080</td>
<td id="4B255DCB805F0041FBA57264D8EFFED2" box="[1036,1163,274,299]" gridcol="4" gridrow="2" pageId="14" pageNumber="319">0.0760.084</td>
<td id="4B255DCB805F0041FB727264D94BFED2" box="[1243,1327,274,299]" gridcol="5" gridrow="2" pageId="14" pageNumber="319">0.081</td>
</tr>
<tr id="08F434B7805F0041FF0F7241D94BFEA9" box="[166,1327,311,336]" gridrow="3" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F7241DD3DFEA9" box="[166,345,311,336]" gridcol="0" gridrow="3" pageId="14" pageNumber="319">IN/SVL</th>
<td id="4B255DCB805F0041FE087241DE4EFEA9" box="[417,554,311,336]" gridcol="1" gridrow="3" pageId="14" pageNumber="319">0.049</td>
<td id="4B255DCB805F0041FDC67241DE8AFEA9" box="[623,750,311,336]" gridcol="2" gridrow="3" pageId="14" pageNumber="319">0.0480.050</td>
<td id="4B255DCB805F0041FC977241DFBAFEA9" box="[830,990,311,336]" gridcol="3" gridrow="3" pageId="14" pageNumber="319">0.049</td>
<td id="4B255DCB805F0041FBA57241D8EFFEA9" box="[1036,1163,311,336]" gridcol="4" gridrow="3" pageId="14" pageNumber="319">0.0480.051</td>
<td id="4B255DCB805F0041FB727241D94BFEA9" box="[1243,1327,311,336]" gridcol="5" gridrow="3" pageId="14" pageNumber="319">0.065</td>
</tr>
<tr id="08F434B7805F0041FF0F722BD94BFE8F" box="[166,1327,349,374]" gridrow="4" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F722BDD3DFE8F" box="[166,345,349,374]" gridcol="0" gridrow="4" pageId="14" pageNumber="319">SN/SVL</th>
<td id="4B255DCB805F0041FE08722BDE4EFE8F" box="[417,554,349,374]" gridcol="1" gridrow="4" pageId="14" pageNumber="319">0.096</td>
<td id="4B255DCB805F0041FDC6722BDE8AFE8F" box="[623,750,349,374]" gridcol="2" gridrow="4" pageId="14" pageNumber="319">0.0930.097</td>
<td id="4B255DCB805F0041FC97722BDFBAFE8F" box="[830,990,349,374]" gridcol="3" gridrow="4" pageId="14" pageNumber="319">0.097</td>
<td id="4B255DCB805F0041FBA5722BD8EFFE8F" box="[1036,1163,349,374]" gridcol="4" gridrow="4" pageId="14" pageNumber="319">0.0940.10</td>
<td id="4B255DCB805F0041FB72722BD94BFE8F" box="[1243,1327,349,374]" gridcol="5" gridrow="4" pageId="14" pageNumber="319">0.10</td>
</tr>
<tr id="08F434B7805F0041FF0F72F5D94BFE65" box="[166,1327,387,412]" gridrow="5" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F72F5DD3DFE65" box="[166,345,387,412]" gridcol="0" gridrow="5" pageId="14" pageNumber="319">EE/SVL</th>
<td id="4B255DCB805F0041FE0872F5DE4EFE65" box="[417,554,387,412]" gridcol="1" gridrow="5" pageId="14" pageNumber="319">0.083</td>
<td id="4B255DCB805F0041FDC672F5DE8AFE65" box="[623,750,387,412]" gridcol="2" gridrow="5" pageId="14" pageNumber="319">0.0810.085</td>
<td id="4B255DCB805F0041FC9772F5DFBAFE65" box="[830,990,387,412]" gridcol="3" gridrow="5" pageId="14" pageNumber="319">0.083</td>
<td id="4B255DCB805F0041FBA572F5D8EFFE65" box="[1036,1163,387,412]" gridcol="4" gridrow="5" pageId="14" pageNumber="319">0.0790.085</td>
<td id="4B255DCB805F0041FB7272F5D94BFE65" box="[1243,1327,387,412]" gridcol="5" gridrow="5" pageId="14" pageNumber="319">0.092</td>
</tr>
<tr id="08F434B7805F0041FF0F72DFD94BFE3B" box="[166,1327,425,450]" gridrow="6" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F72DFDD3DFE3B" box="[166,345,425,450]" gridcol="0" gridrow="6" pageId="14" pageNumber="319">HW/HL</th>
<td id="4B255DCB805F0041FE0872DFDE4EFE3B" box="[417,554,425,450]" gridcol="1" gridrow="6" pageId="14" pageNumber="319">0.89</td>
<td id="4B255DCB805F0041FDC672DFDE8AFE3B" box="[623,750,425,450]" gridcol="2" gridrow="6" pageId="14" pageNumber="319">0.880.92</td>
<td id="4B255DCB805F0041FC9772DFDFBAFE3B" box="[830,990,425,450]" gridcol="3" gridrow="6" pageId="14" pageNumber="319">0.85</td>
<td id="4B255DCB805F0041FBA572DFD8EFFE3B" box="[1036,1163,425,450]" gridcol="4" gridrow="6" pageId="14" pageNumber="319">0.830.89</td>
<td id="4B255DCB805F0041FB7272DFD94BFE3B" box="[1243,1327,425,450]" gridcol="5" gridrow="6" pageId="14" pageNumber="319">0.90</td>
</tr>
<tr id="08F434B7805F0041FF0F72B9D94BFE11" box="[166,1327,463,488]" gridrow="7" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F72B9DD3DFE11" box="[166,345,463,488]" gridcol="0" gridrow="7" pageId="14" pageNumber="319">EY/EN</th>
<td id="4B255DCB805F0041FE0872B9DE4EFE11" box="[417,554,463,488]" gridcol="1" gridrow="7" pageId="14" pageNumber="319">0.69</td>
<td id="4B255DCB805F0041FDC672B9DE8AFE11" box="[623,750,463,488]" gridcol="2" gridrow="7" pageId="14" pageNumber="319">0.660.71</td>
<td id="4B255DCB805F0041FC9772B9DFBAFE11" box="[830,990,463,488]" gridcol="3" gridrow="7" pageId="14" pageNumber="319">0.70</td>
<td id="4B255DCB805F0041FBA572B9D8EFFE11" box="[1036,1163,463,488]" gridcol="4" gridrow="7" pageId="14" pageNumber="319">0.620.79</td>
<td id="4B255DCB805F0041FB7272B9D94BFE11" box="[1243,1327,463,488]" gridcol="5" gridrow="7" pageId="14" pageNumber="319">1.07</td>
</tr>
<tr id="08F434B7805F0041FF0F7282D94BFDF4" box="[166,1327,500,525]" gridrow="8" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F7282DD3DFDF4" box="[166,345,500,525]" gridcol="0" gridrow="8" pageId="14" pageNumber="319">EN/IN</th>
<td id="4B255DCB805F0041FE087282DE4EFDF4" box="[417,554,500,525]" gridcol="1" gridrow="8" pageId="14" pageNumber="319">1.7</td>
<td id="4B255DCB805F0041FDC67282DE8AFDF4" box="[623,750,500,525]" gridcol="2" gridrow="8" pageId="14" pageNumber="319">1.61.8</td>
<td id="4B255DCB805F0041FC977282DFBAFDF4" box="[830,990,500,525]" gridcol="3" gridrow="8" pageId="14" pageNumber="319">1.6</td>
<td id="4B255DCB805F0041FBA57282D8EFFDF4" box="[1036,1163,500,525]" gridcol="4" gridrow="8" pageId="14" pageNumber="319">1.51.7</td>
<td id="4B255DCB805F0041FB727282D94BFDF4" box="[1243,1327,500,525]" gridcol="5" gridrow="8" pageId="14" pageNumber="319">1.25</td>
</tr>
<tr id="08F434B7805F0041FF0F716DD94BFDCA" box="[166,1327,539,563]" gridrow="9" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F716DDD3DFDCA" box="[166,345,539,563]" gridcol="0" gridrow="9" pageId="14" pageNumber="319">SN/HL</th>
<td id="4B255DCB805F0041FE08716DDE4EFDCA" box="[417,554,539,563]" gridcol="1" gridrow="9" pageId="14" pageNumber="319">0.41</td>
<td id="4B255DCB805F0041FDC6716DDE8AFDCA" box="[623,750,539,563]" gridcol="2" gridrow="9" pageId="14" pageNumber="319">0.410.42</td>
<td id="4B255DCB805F0041FC97716DDFBAFDCA" box="[830,990,539,563]" gridcol="3" gridrow="9" pageId="14" pageNumber="319">0.42</td>
<td id="4B255DCB805F0041FBA5716DD8EFFDCA" box="[1036,1163,539,563]" gridcol="4" gridrow="9" pageId="14" pageNumber="319">0.410.43</td>
<td id="4B255DCB805F0041FB72716DD94BFDCA" box="[1243,1327,539,563]" gridcol="5" gridrow="9" pageId="14" pageNumber="319">0.38</td>
</tr>
<tr id="08F434B7805F0041FF0F7136D94BFDA0" box="[166,1327,576,601]" gridrow="10" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F7136DD3DFDA0" box="[166,345,576,601]" gridcol="0" gridrow="10" pageId="14" pageNumber="319">IN/EN</th>
<td id="4B255DCB805F0041FE087136DE4EFDA0" box="[417,554,576,601]" gridcol="1" gridrow="10" pageId="14" pageNumber="319">0.59</td>
<td id="4B255DCB805F0041FDC67136DE8AFDA0" box="[623,750,576,601]" gridcol="2" gridrow="10" pageId="14" pageNumber="319">0.570.62</td>
<td id="4B255DCB805F0041FC977136DFBAFDA0" box="[830,990,576,601]" gridcol="3" gridrow="10" pageId="14" pageNumber="319">0.61</td>
<td id="4B255DCB805F0041FBA57136D8EFFDA0" box="[1036,1163,576,601]" gridcol="4" gridrow="10" pageId="14" pageNumber="319">0.590.67</td>
<td id="4B255DCB805F0041FB727136D94BFDA0" box="[1243,1327,576,601]" gridcol="5" gridrow="10" pageId="14" pageNumber="319">0.80</td>
</tr>
<tr id="08F434B7805F0041FF0F7110D94BFD86" box="[166,1327,614,639]" gridrow="11" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F7110DD3DFD86" box="[166,345,614,639]" gridcol="0" gridrow="11" pageId="14" pageNumber="319">SN/EY</th>
<td id="4B255DCB805F0041FE087110DE4EFD86" box="[417,554,614,639]" gridcol="1" gridrow="11" pageId="14" pageNumber="319">1.7</td>
<td id="4B255DCB805F0041FDC67110DE8AFD86" box="[623,750,614,639]" gridcol="2" gridrow="11" pageId="14" pageNumber="319">1.61.8</td>
<td id="4B255DCB805F0041FC977110DFBAFD86" box="[830,990,614,639]" gridcol="3" gridrow="11" pageId="14" pageNumber="319">1.7</td>
<td id="4B255DCB805F0041FBA57110D8EFFD86" box="[1036,1163,614,639]" gridcol="4" gridrow="11" pageId="14" pageNumber="319">1.61.9</td>
<td id="4B255DCB805F0041FB727110D94BFD86" box="[1243,1327,614,639]" gridcol="5" gridrow="11" pageId="14" pageNumber="319">1.1</td>
</tr>
<tr id="08F434B7805F0041FF0F71FAD94BFD5C" box="[166,1327,652,677]" gridrow="12" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F71FADD3DFD5C" box="[166,345,652,677]" gridcol="0" gridrow="12" pageId="14" pageNumber="319">EY/HL</th>
<td id="4B255DCB805F0041FE0871FADE4EFD5C" box="[417,554,652,677]" gridcol="1" gridrow="12" pageId="14" pageNumber="319">0.25</td>
<td id="4B255DCB805F0041FDC671FADE8AFD5C" box="[623,750,652,677]" gridcol="2" gridrow="12" pageId="14" pageNumber="319">0.230.26</td>
<td id="4B255DCB805F0041FC9771FADFBAFD5C" box="[830,990,652,677]" gridcol="3" gridrow="12" pageId="14" pageNumber="319">0.24</td>
<td id="4B255DCB805F0041FBA571FAD8EFFD5C" box="[1036,1163,652,677]" gridcol="4" gridrow="12" pageId="14" pageNumber="319">0.220.27</td>
<td id="4B255DCB805F0041FB7271FAD94BFD5C" box="[1243,1327,652,677]" gridcol="5" gridrow="12" pageId="14" pageNumber="319">0.33</td>
</tr>
<tr id="08F434B7805F0041FF0F71C7D94BFD33" box="[166,1327,689,714]" gridrow="13" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F71C7DD3DFD33" box="[166,345,689,714]" gridcol="0" gridrow="13" pageId="14" pageNumber="319">EarL/HL</th>
<td id="4B255DCB805F0041FE0871C7DE4EFD33" box="[417,554,689,714]" gridcol="1" gridrow="13" pageId="14" pageNumber="319">0.079</td>
<td id="4B255DCB805F0041FDC671C7DE8AFD33" box="[623,750,689,714]" gridcol="2" gridrow="13" pageId="14" pageNumber="319">0.0760.083</td>
<td id="4B255DCB805F0041FC9771C7DFBAFD33" box="[830,990,689,714]" gridcol="3" gridrow="13" pageId="14" pageNumber="319">0.063</td>
<td id="4B255DCB805F0041FBA571C7D8EFFD33" box="[1036,1163,689,714]" gridcol="4" gridrow="13" pageId="14" pageNumber="319">0.0560.071</td>
<td id="4B255DCB805F0041FB7271C7D94BFD33" box="[1243,1327,689,714]" gridcol="5" gridrow="13" pageId="14" pageNumber="319">0.083</td>
</tr>
<tr id="08F434B7805F0041FF0F71A1D94BFD09" box="[166,1327,727,752]" gridrow="14" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F71A1DD3DFD09" box="[166,345,727,752]" gridcol="0" gridrow="14" pageId="14" pageNumber="319">EE/EY</th>
<td id="4B255DCB805F0041FE0871A1DE4EFD09" box="[417,554,727,752]" gridcol="1" gridrow="14" pageId="14" pageNumber="319">1.4</td>
<td id="4B255DCB805F0041FDC671A1DE8AFD09" box="[623,750,727,752]" gridcol="2" gridrow="14" pageId="14" pageNumber="319">1.41.5</td>
<td id="4B255DCB805F0041FC9771A1DFBAFD09" box="[830,990,727,752]" gridcol="3" gridrow="14" pageId="14" pageNumber="319">1.5</td>
<td id="4B255DCB805F0041FBA571A1D8EFFD09" box="[1036,1163,727,752]" gridcol="4" gridrow="14" pageId="14" pageNumber="319">1.31.6</td>
<td id="4B255DCB805F0041FB7271A1D94BFD09" box="[1243,1327,727,752]" gridcol="5" gridrow="14" pageId="14" pageNumber="319">1.1</td>
</tr>
<tr id="08F434B7805F0041FF0F7188D94BFCEF" box="[166,1327,766,790]" gridrow="15" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F7188DD3DFCEF" box="[166,345,766,790]" gridcol="0" gridrow="15" pageId="14" pageNumber="319">T4L/SVL</th>
<td id="4B255DCB805F0041FE087188DE4EFCEF" box="[417,554,766,790]" gridcol="1" gridrow="15" pageId="14" pageNumber="319">0.11</td>
<td id="4B255DCB805F0041FDC67188DE8AFCEF" box="[623,750,766,790]" gridcol="2" gridrow="15" pageId="14" pageNumber="319">0.110.12</td>
<td id="4B255DCB805F0041FC977188DFBAFCEF" box="[830,990,766,790]" gridcol="3" gridrow="15" pageId="14" pageNumber="319">0.12</td>
<td id="4B255DCB805F0041FBA57188D8EFFCEF" box="[1036,1163,766,790]" gridcol="4" gridrow="15" pageId="14" pageNumber="319">0.110.13</td>
<td id="4B255DCB805F0041FB727188D94BFCEF" box="[1243,1327,766,790]" gridcol="5" gridrow="15" pageId="14" pageNumber="319">0.11</td>
</tr>
<tr id="08F434B7805F0041FF0F7055D94BFCC5" box="[166,1327,803,828]" gridrow="16" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F7055DD3DFCC5" box="[166,345,803,828]" gridcol="0" gridrow="16" pageId="14" pageNumber="319">T4width/T4L</th>
<td id="4B255DCB805F0041FE087055DE4EFCC5" box="[417,554,803,828]" gridcol="1" gridrow="16" pageId="14" pageNumber="319">0.31</td>
<td id="4B255DCB805F0041FDC67055DE8AFCC5" box="[623,750,803,828]" gridcol="2" gridrow="16" pageId="14" pageNumber="319">0.290.33</td>
<td id="4B255DCB805F0041FC977055DFBAFCC5" box="[830,990,803,828]" gridcol="3" gridrow="16" pageId="14" pageNumber="319">0.30</td>
<td id="4B255DCB805F0041FBA57055D8EFFCC5" box="[1036,1163,803,828]" gridcol="4" gridrow="16" pageId="14" pageNumber="319">0.260.34</td>
<td id="4B255DCB805F0041FB727055D94BFCC5" box="[1243,1327,803,828]" gridcol="5" gridrow="16" pageId="14" pageNumber="319">0.33</td>
</tr>
<tr id="08F434B7805F0041FF0F703FD94BFC9B" box="[166,1327,841,866]" gridrow="17" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F703FDD3DFC9B" box="[166,345,841,866]" gridcol="0" gridrow="17" pageId="14" pageNumber="319">webL/T4L</th>
<td id="4B255DCB805F0041FE08703FDE4EFC9B" box="[417,554,841,866]" gridcol="1" gridrow="17" pageId="14" pageNumber="319">0.23</td>
<td id="4B255DCB805F0041FDC6703FDE8AFC9B" box="[623,750,841,866]" gridcol="2" gridrow="17" pageId="14" pageNumber="319">0.210.26</td>
<td id="4B255DCB805F0041FC97703FDFBAFC9B" box="[830,990,841,866]" gridcol="3" gridrow="17" pageId="14" pageNumber="319">0.21</td>
<td id="4B255DCB805F0041FBA5703FD8EFFC9B" box="[1036,1163,841,866]" gridcol="4" gridrow="17" pageId="14" pageNumber="319">0.180.26</td>
<td id="4B255DCB805F0041FB72703FD94BFC9B" box="[1243,1327,841,866]" gridcol="5" gridrow="17" pageId="14" pageNumber="319">0.19</td>
</tr>
<tr id="08F434B7805F0041FF0F7018D94BFC7E" box="[166,1327,878,903]" gridrow="18" pageId="14" pageNumber="319">
<th id="4B255DCB805F0041FF0F7018DD3DFC7E" box="[166,345,878,903]" gridcol="0" gridrow="18" pageId="14" pageNumber="319">T4lamellaeL/T4L</th>
<td id="4B255DCB805F0041FE087018DE4EFC7E" box="[417,554,878,903]" gridcol="1" gridrow="18" pageId="14" pageNumber="319">0.78</td>
<td id="4B255DCB805F0041FDC67018DE8AFC7E" box="[623,750,878,903]" gridcol="2" gridrow="18" pageId="14" pageNumber="319">0.740.82</td>
<td id="4B255DCB805F0041FC977018DFBAFC7E" box="[830,990,878,903]" gridcol="3" gridrow="18" pageId="14" pageNumber="319">0.85</td>
<td id="4B255DCB805F0041FBA57018D8EFFC7E" box="[1036,1163,878,903]" gridcol="4" gridrow="18" pageId="14" pageNumber="319">0.670.93</td>
<td id="4B255DCB805F0041FB727018D94BFC7E" box="[1243,1327,878,903]" gridcol="5" gridrow="18" pageId="14" pageNumber="319">0.86</td>
</tr>
</table>
</paragraph>
<caption id="E2BB667D805FFFB0FF3E747BDE6AF897" httpUri="https://zenodo.org/record/3363253/files/figure.png" pageId="14" pageNumber="319" startId="14.[151,250,1805,1830]" targetBox="[151,1436,928,1786]" targetPageId="14">
<paragraph id="B67B36F5805FFFB0FF3E747BDE6AF897" blockId="14.[151,1437,1805,1902]" pageId="14" pageNumber="319">
<emphasis id="84B0EAE7805FFFB0FF3E747BDD70F8DF" bold="true" box="[151,276,1805,1830]" pageId="14" pageNumber="319">FIGURE 4.</emphasis>
Portraits of new
<taxonomicName id="71C44D76805FFFB0FE6F747BDE05F8DC" box="[454,609,1805,1829]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="319" phylum="Chordata" rank="genus">
<emphasis id="84B0EAE7805FFFB0FE6F747BDE05F8DC" box="[454,609,1805,1829]" italics="true" pageId="14" pageNumber="319">Lepidodactylus</emphasis>
</taxonomicName>
species in life. (A) holotype of
<taxonomicName id="71C44D76805FFFB0FC007478D879F8DC" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[937,1053,1805,1829]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="319" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805FFFB0FC007478D879F8DC" box="[937,1053,1805,1829]" italics="true" pageId="14" pageNumber="319">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805FFFB0FB8D7478D812F8DF" box="[1060,1142,1806,1830]" pageId="14" pageNumber="319" rank="species">
<emphasis id="84B0EAE7805FFFB0FB8D7478D812F8DF" bold="true" box="[1060,1142,1806,1830]" pageId="14" pageNumber="319">sp. nov.</emphasis>
</taxonomicNameLabel>
(UMMZ 244058), (B) holo- type of
<taxonomicName id="71C44D76805FFFB0FF4C7444DD11F8B0" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[229,373,1841,1865]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="319" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE7805FFFB0FF4C7444DD11F8B0" box="[229,373,1841,1865]" italics="true" pageId="14" pageNumber="319">L. kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805FFFB0FED57444DDA9F8B3" box="[380,461,1842,1866]" pageId="14" pageNumber="319" rank="species">
<emphasis id="84B0EAE7805FFFB0FED57444DDA9F8B3" bold="true" box="[380,461,1842,1866]" pageId="14" pageNumber="319">sp. nov.</emphasis>
</taxonomicNameLabel>
(BPBM 39880), (C) paratype of
<taxonomicName id="71C44D76805FFFB0FC897444DFD4F8B0" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[800,944,1841,1865]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="319" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE7805FFFB0FC897444DFD4F8B0" box="[800,944,1841,1865]" italics="true" pageId="14" pageNumber="319">L. kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805FFFB0FC1E7444D86CF8B3" box="[951,1032,1842,1866]" pageId="14" pageNumber="319" rank="species">
<emphasis id="84B0EAE7805FFFB0FC1E7444D86CF8B3" bold="true" box="[951,1032,1842,1866]" pageId="14" pageNumber="319">sp. nov.</emphasis>
</taxonomicNameLabel>
(BPBM 39879), and (D) paratype of
<taxonomicName id="71C44D76805FFFB0FA217444DD68F894" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="319" phylum="Chordata" rank="species" species="kwasnickae" status="sp. nov.">
<emphasis id="84B0EAE7805FFFB0FA217444DD68F894" italics="true" pageId="14" pageNumber="319">L. kwasnickae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805FFFB0FEBB7420DD07F897" box="[274,355,1878,1902]" pageId="14" pageNumber="319" rank="species">
<emphasis id="84B0EAE7805FFFB0FEBB7420DD07F897" bold="true" box="[274,355,1878,1902]" pageId="14" pageNumber="319">sp. nov.</emphasis>
</taxonomicNameLabel>
(BPBM 39151).
</paragraph>
</caption>
<paragraph id="B67B36F5805FFFB1FF6E74E0D8C2FE9E" blockId="14.[151,1436,1942,2041]" lastBlockId="15.[151,1436,153,359]" lastPageId="15" lastPageNumber="320" pageId="14" pageNumber="319">
With one important exception, dorsal color pattern in the adult
<typeStatus id="697F8857805FFFB0FC2074E1DF90F848" box="[905,1012,1943,1969]" pageId="14" pageNumber="319" type="paratype">paratypes</typeStatus>
is much like that seen in the
<typeStatus id="697F8857805FFFB0FA9374E0D9F8F849" box="[1338,1436,1942,1968]" pageId="14" pageNumber="319" type="holotype">holotype</typeStatus>
except that BPBM 39883 has the crossbands better developed and BPBM 38881 is paler overall. However, BPBM 39151 differs considerably from all other adult specimens, having a wide orange-brown mid-dorsal stripe below which the sides are evenly dark brown (
<figureCitation id="2EFF2A70805EFFB1FDF873EFDECFFF4A" box="[593,683,153,179]" captionStart="FIGURE 4" captionStartId="14.[151,250,1805,1830]" captionTargetBox="[151,1436,928,1786]" captionTargetId="figure@14.[151,1436,928,1786]" captionTargetPageId="14" captionText="FIGURE 4. Portraits of new Lepidodactylus species in life. (A) holotype of L. mitchelli sp. nov. (UMMZ 244058), (B) holo- type of L. kwasnickae sp. nov. (BPBM 39880), (C) paratype of L. kwasnickae sp. nov. (BPBM 39879), and (D) paratype of L. kwasnickae sp. nov. (BPBM 39151)." httpUri="https://zenodo.org/record/3363253/files/figure.png" pageId="15" pageNumber="320">Fig. 4D</figureCitation>
). Lateral and dorsolateral rows of pale spots are present in all speci- mens but vary in their distinctiveness. The venters are white with a few brown scales (as seen in the
<typeStatus id="697F8857805EFFB1FABA73CBD913FF21" box="[1299,1399,189,216]" pageId="15" pageNumber="320" type="holotype">holotype</typeStatus>
) in BPBM 39149 and BPBM 39879 but are more spotted with brown in the other specimens, and are heavily spotted with brown scales in BPBM 39152 and BPBM 39882. All specimens have the pale postocular stripe and copper iris heavily veined with dark copper as seen in the
<typeStatus id="697F8857805EFFB1FD34725FDF67FEBA" box="[669,771,297,323]" pageId="15" pageNumber="320" type="holotype">holotype</typeStatus>
. The juvenile is brown with a wide mid-dorsal stripe of pale brown narrowly margined posteriorly with dark brown; venter heavily peppered with brown.
</paragraph>
<caption id="E2BB667D805EFFB1FF3E74EFDE43F803" httpUri="https://zenodo.org/record/3363255/files/figure.png" pageId="15" pageNumber="320" startId="15.[151,250,1945,1970]" targetBox="[185,1401,381,1926]" targetPageId="15">
<paragraph id="B67B36F5805EFFB1FF3E74EFDE43F803" blockId="15.[151,1436,1945,2042]" pageId="15" pageNumber="320">
<emphasis id="84B0EAE7805EFFB1FF3E74EFDD70F84B" bold="true" box="[151,276,1945,1970]" pageId="15" pageNumber="320">FIGURE 5.</emphasis>
(A) Boiaboiawaga Island, Milne Bay Province, Papua New Guinea, habitat of
<taxonomicName id="71C44D76805EFFB1FBEB74ECD959F84B" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[1090,1341,1946,1970]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="320" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE7805EFFB1FBEB74ECD959F84B" box="[1090,1341,1946,1970]" italics="true" pageId="15" pageNumber="320">Lepidodactylus mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C805EFFB1FAED74ECD9F1F84B" box="[1348,1429,1946,1970]" pageId="15" pageNumber="320" rank="species">
<emphasis id="84B0EAE7805EFFB1FAED74ECD9F1F84B" bold="true" box="[1348,1429,1946,1970]" pageId="15" pageNumber="320">sp. nov.</emphasis>
</taxonomicNameLabel>
, photo by D. Mitchell; and (B) interior forest on Woodlark Island, Milne Bay Province, Papua New Guinea, habitat of
<emphasis id="84B0EAE7805EFFB1FAEF74C9DCBDF803" italics="true" pageId="15" pageNumber="320">L. kwas- nickae</emphasis>
<emphasis id="84B0EAE7805EFFB1FF497494DD54F803" bold="true" box="[224,304,2018,2042]" pageId="15" pageNumber="320">sp. nov.</emphasis>
, photo by D. Polhemus.
</paragraph>
</caption>
<paragraph id="B67B36F58041FFAEFF6E73EFDF90FD52" blockId="16.[151,1437,153,1295]" pageId="16" pageNumber="321">
<emphasis id="84B0EAE78041FFAEFF6E73EFDD30FF4A" box="[199,340,153,179]" italics="true" pageId="16" pageNumber="321">Color in life</emphasis>
. This species shows considerable variation in color pattern. Field notes for BPBM 39149 stated “Dorsum tan with six irregular dark-brown crossbands, each partially or entirely divided by ground color vertebrally. Four more brown bands on tail, each margined posteriorly in dark brown and then white. Lips and face with faint yellowish cast. Venter white, heavily speckled with brown, but with faint yellow wash on throat (but not on chin). Iris tan reticulated with dark brown. Brown dorsal bands almost speckled in appearance because of interspersed tan scales.” BPBM 39879 was brown with irregular darker-brown bands; sides darker than mid-dorsum; bands on the tail dark brown on a tan ground; with a light-brown postocular stripe (
<figureCitation id="2EFF2A708041FFAEFC107207D877FE72" box="[953,1043,369,395]" captionStart="FIGURE 4" captionStartId="14.[151,250,1805,1830]" captionTargetBox="[151,1436,928,1786]" captionTargetId="figure@14.[151,1436,928,1786]" captionTargetPageId="14" captionText="FIGURE 4. Portraits of new Lepidodactylus species in life. (A) holotype of L. mitchelli sp. nov. (UMMZ 244058), (B) holo- type of L. kwasnickae sp. nov. (BPBM 39880), (C) paratype of L. kwasnickae sp. nov. (BPBM 39879), and (D) paratype of L. kwasnickae sp. nov. (BPBM 39151)." httpUri="https://zenodo.org/record/3363253/files/figure.png" pageId="16" pageNumber="321">Fig. 4C</figureCitation>
); venter white with brown punctations, yellower around vent. The
<typeStatus id="697F88578041FFAEFDA672E3DE15FE56" box="[527,625,405,431]" pageId="16" pageNumber="321" type="holotype">holotype</typeStatus>
(BPBM 39880) was similar but with a row of white spots on each side (
<figureCitation id="2EFF2A708041FFAEFF0972CFDC9CFE2A" box="[160,248,441,467]" captionStart="FIGURE 4" captionStartId="14.[151,250,1805,1830]" captionTargetBox="[151,1436,928,1786]" captionTargetId="figure@14.[151,1436,928,1786]" captionTargetPageId="14" captionText="FIGURE 4. Portraits of new Lepidodactylus species in life. (A) holotype of L. mitchelli sp. nov. (UMMZ 244058), (B) holo- type of L. kwasnickae sp. nov. (BPBM 39880), (C) paratype of L. kwasnickae sp. nov. (BPBM 39879), and (D) paratype of L. kwasnickae sp. nov. (BPBM 39151)." httpUri="https://zenodo.org/record/3363253/files/figure.png" pageId="16" pageNumber="321">Fig. 4B</figureCitation>
); BPBM 39881 was the same but with tan spots on sides. An alternate color pattern was found in the juvenile BPBM 39150: “Dark-orange vertebral stripe bordered by charcoal-gray stripes, below which everything is dark gray. Mid-venter with pink cast.” BPBM 39151 had a similar striped pattern but the colors differed: “Light orange vertebral stripe and light orange on top of head. Vertebral stripe bordered by black stripe on each side, below which is tan with dark-brown spots, some of these elongated and arrayed in a broken lateral stripe. Venter white, heavily speckled with brown scales. Iris tan reticulated with dark brown. Tongue pink.” BPBM 39152 was dark brown above with an orange-brown vertebral zigzag and scattered tan dots laterally.
</paragraph>
<paragraph id="B67B36F58041FFAEFF6E71C3DF0FFD0A" blockId="16.[151,1437,153,1295]" pageId="16" pageNumber="321">
<emphasis id="84B0EAE78041FFAEFF6E71C3DD5BFD36" box="[199,319,693,719]" italics="true" pageId="16" pageNumber="321">Etymology</emphasis>
. The name is a genitive honorific for my friend Gretta Kwasnicka who has long provided gracious hospitality and support for my fieldwork in
<collectingRegion id="7400F8178041FFAEFD2B71AFDF03FD0A" box="[642,871,729,755]" country="Papua New Guinea" name="Milne Bay" pageId="16" pageNumber="321">Milne Bay Province</collectingRegion>
.
</paragraph>
<paragraph id="B67B36F58041FFAEFF6E7188DE25FCC2" blockId="16.[151,1437,153,1295]" pageId="16" pageNumber="321">
<emphasis id="84B0EAE78041FFAEFF6E7188DD6BFCEE" box="[199,271,766,791]" italics="true" pageId="16" pageNumber="321">Range</emphasis>
. Known from central Woodlark Island, but it no doubt ranges across the entire extent of this low (maximum elevation
<quantity id="713C9B108041FFAEFEEC7057DDE9FCC2" box="[325,397,801,827]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="16" pageNumber="321" unit="m" value="240.0">240 m</quantity>
) island (
<figureCitation id="2EFF2A708041FFAEFE467057DE50FCC2" box="[495,564,801,827]" captionStart="FIGURE 2" captionStartId="7.[151,248,874,899]" captionTargetBox="[162,1379,249,813]" captionTargetId="figure@7.[162,1379,249,813]" captionTargetPageId="7" captionText="FIGURE 2. Map of New Guinea and surrounding islands showing type localities for Lepidodactylus aignanus sp. nov. (diamond), L. dialeukos sp. nov. (triangle), L. kwasnickae sp. nov. (square), L. mitchelli sp. nov. (circle), and L. zweifeli sp. nov. (star)." httpUri="https://zenodo.org/record/3363249/files/figure.png" pageId="16" pageNumber="321">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="B67B36F58041FFAEFF6E7033D8C6FC32" blockId="16.[151,1437,153,1295]" pageId="16" pageNumber="321">
<emphasis id="84B0EAE78041FFAEFF6E7033DD47FCA6" box="[199,291,837,863]" italics="true" pageId="16" pageNumber="321">Ecology</emphasis>
. We collected this species from
<taxonomicName id="71C44D768041FFAEFD3A7033DF67FCA6" box="[659,771,837,863]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="16" pageNumber="321" phylum="Tracheophyta" rank="genus">
<emphasis id="84B0EAE78041FFAEFD3A7033DF67FCA6" box="[659,771,837,863]" italics="true" pageId="16" pageNumber="321">Pandanus</emphasis>
</taxonomicName>
axils in densely shaded second-growth forest (
<figureCitation id="2EFF2A708041FFAEFABE7033D914FCA6" box="[1303,1392,836,863]" captionStart="FIGURE 5" captionStartId="15.[151,250,1945,1970]" captionTargetBox="[185,1401,381,1926]" captionTargetId="figure@15.[185,1401,381,1926]" captionTargetPageId="15" captionText="FIGURE 5. (A) Boiaboiawaga Island, Milne Bay Province, Papua New Guinea, habitat of Lepidodactylus mitchelli sp. nov., photo by D. Mitchell; and (B) interior forest on Woodlark Island, Milne Bay Province, Papua New Guinea, habitat of L. kwas- nickae sp. nov., photo by D. Polhemus." httpUri="https://zenodo.org/record/3363255/files/figure.png" pageId="16" pageNumber="321">Fig. 5B</figureCitation>
). In nearby areas that had been largely cleared of trees,
<taxonomicName id="71C44D768041FFAEFD7C701CDF2BFC7A" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" box="[725,847,873,899]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="321" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="84B0EAE78041FFAEFD7C701CDF2BFC7A" box="[725,847,873,899]" italics="true" pageId="16" pageNumber="321">L. lugubris</emphasis>
</taxonomicName>
occupied the remaining
<taxonomicName id="71C44D768041FFAEFBCD701FD8B0FC7A" box="[1124,1236,873,899]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="16" pageNumber="321" phylum="Tracheophyta" rank="genus">
<emphasis id="84B0EAE78041FFAEFBCD701FD8B0FC7A" box="[1124,1236,873,899]" italics="true" pageId="16" pageNumber="321">Pandanus</emphasis>
</taxonomicName>
trees. Hence, this species appears to be an obligate forest dweller. Much of Woodlark Island is currently covered by recovering secondary rainforest, so habitat for this apparently obligate forest dweller is presently adequate.
</paragraph>
<paragraph id="B67B36F58041FFAEFF6E70A3DFEFFC16" blockId="16.[151,1437,153,1295]" box="[199,907,981,1007]" pageId="16" pageNumber="321">BPBM 39883 had a few mites between the lamellae of one toe.</paragraph>
<paragraph id="B67B36F58041FFAEFF6E708FD841FB86" blockId="16.[151,1437,153,1295]" pageId="16" pageNumber="321">
<emphasis id="84B0EAE78041FFAEFF6E708FDD4DFBEA" box="[199,297,1017,1043]" italics="true" pageId="16" pageNumber="321">Remarks</emphasis>
. This species was found to be the sister taxon to
<taxonomicName id="71C44D768041FFAEFCE2708CDFAEFBEA" authority="Kraus, 2019" authorityName="Kraus" authorityYear="2019" box="[843,970,1017,1043]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="321" phylum="Chordata" rank="species" species="mitchelli" status="sp. nov.">
<emphasis id="84B0EAE78041FFAEFCE2708CDFAEFBEA" box="[843,970,1017,1043]" italics="true" pageId="16" pageNumber="321">L. mitchelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F83579C8041FFAEFC78708FD84EFBEA" box="[977,1066,1017,1043]" pageId="16" pageNumber="321" rank="species">
<emphasis id="84B0EAE78041FFAEFC78708FD84EFBEA" bold="true" box="[977,1066,1017,1043]" pageId="16" pageNumber="321">sp. nov.</emphasis>
</taxonomicNameLabel>
in the phylogenetic study of
<bibRefCitation id="D2554B048041FFAEFAC6708FDD39FBCE" author="Oliver, P. M. &amp; Brown, R. &amp; Kraus, F. &amp; Rittmeyer, E. &amp; Travers, S. L. &amp; Siler, C. D." pageId="16" pageNumber="321" pagination="20171760" refId="ref19729" refString="Oliver, P. M., Brown, R., Kraus, F., Rittmeyer, E., Travers, S. L. &amp; Siler, C. D. (2018 a) Lizards of the lost arcs: mid-Cenozoic diversification, persistence and ecological marginalization around the western Pacific. Proceedings of the Royal Society of London B, 285 (1871), 20171760. https: // doi. org / 10.1098 / rspb. 2017.1760" type="journal article" year="2018">
Oliver
<emphasis id="84B0EAE78041FFAEFF6A7768DC98FBCE" box="[195,252,1053,1079]" italics="true" pageId="16" pageNumber="321">et al.</emphasis>
(2018a)
</bibRefCitation>
, and these two species were estimated by them to have been separated for approximately 4 MY, which accords well with the timing of the opening of the Woodlark Rift, which has moved Woodlark Island away from the other islands of Milne Bay Province since that time (
<bibRefCitation id="D2554B048041FFAEFCE57713D87CFB86" author="Taylor, B. &amp; Goodliffe, A. M. &amp; Martinez, F." box="[844,1048,1125,1151]" pageId="16" pageNumber="321" pagination="7497 - 7512" refId="ref20642" refString="Taylor, B., Goodliffe, A. M. &amp; Martinez, F. (1999) How continents break up: Insights from Papua New Guinea. Journal of Geophysical Research, 104, 7497 - 7512. https: // doi. org / 10.1029 / 1998 JB 900115" type="journal article" year="1999">
Taylor
<emphasis id="84B0EAE78041FFAEFC347710DFB2FB86" box="[925,982,1125,1151]" italics="true" pageId="16" pageNumber="321">et al.</emphasis>
1999
</bibRefCitation>
).
</paragraph>
<paragraph id="B67B36F58041FFAEFF6E77FFDE7CFAF6" blockId="16.[151,1437,153,1295]" pageId="16" pageNumber="321">
I have earlier (
<bibRefCitation id="D2554B048041FFAEFEC377FFDD8BFB5A" author="Kraus, F." box="[362,495,1161,1187]" pageId="16" pageNumber="321" pagination="552 - 558" refId="ref19684" refString="Kraus, F. (2017) A new insular species of Oreophryne (Anura, Microhylidae) from Papua New Guinea. Journal of Herpetology, 51, 552 - 558. https: // doi. org / 10.1670 / 17 - 002" type="journal article" year="2017">Kraus 2017</bibRefCitation>
) discussed conservation threats to the forests of Woodlark, which have included exploratory gold mining and proposals to convert much of the island to oil palm. Should outside pressures for resource exploitation continue, much of this species habitat could be lost, as would be true for the many other forest-dwelling endemic species on Woodlark.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,175 @@
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<mods:title id="28AAD652C0A2847346542CA1E2211D66">Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)</mods:title>
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<subSubSection id="FEDE6561FFCBFFDCFF69A547AB1BF9E4" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="B67B36EAFFCBFFDCFF69A547AB92F9C5" blockId="16.[151,407,1590,1649]" box="[151,407,1590,1616]" pageId="16" pageNumber="17">
<heading id="ED338186FFCBFFDCFF69A547AB92F9C5" bold="true" box="[151,407,1590,1616]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="84B0EAF8FFCBFFDCFF69A547AB92F9C5" bold="true" box="[151,407,1590,1616]" pageId="16" pageNumber="17">
<taxonomicName id="71C44D69FFCBFFDCFF69A547AB38F9C5" ID-CoL="8ZZ76" box="[151,317,1590,1616]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFCBFFDCFF69A547AB38F9C5" bold="true" box="[151,317,1590,1616]" italics="true" pageId="16" pageNumber="17">Agrionympha</emphasis>
</taxonomicName>
male 1
</emphasis>
</heading>
</paragraph>
<paragraph id="B67B36EAFFCBFFDCFF69A528AB1BF9E4" blockId="16.[151,407,1590,1649]" box="[151,286,1625,1649]" pageId="16" pageNumber="17">
<figureCitation id="2EFF2A6FFFCBFFDCFF69A528AAEAF9E4" box="[151,239,1625,1649]" captionStart="FIGURE 10" captionStartId="18.[151,250,1913,1936]" captionTargetBox="[313,1332,581,1845]" captionTargetId="figure@18.[244,1340,573,1969]" captionTargetPageId="18" captionText="FIGURE 10. Agrionympha male genitalia. A B, A. pseliacma, Meyrick, Karkloof Falls: A, lateral view, from slide 3830, Transvaal Museum, Pretoria; B, detail of phallus, lateral. C G, A. sagittella, sp. nov. Hogsback: C, lateral; D, dorsal; E, ventral; F, phallus, lateral; G, phallus dorsal. H J, Agrionympha species 1 , Diepwalle Forest: H, lateral; I, phallus lateral; J, phallus dorsal." httpUri="https://zenodo.org/record/203563/files/figure.png" pageId="16" pageNumber="17">Figs. 10</figureCitation>
HJ
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFCBFFDDFF69A5D0A974F916" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="B67B36EAFFCBFFDDFF69A5D0AF3AF9CA" blockId="16.[151,1436,1697,1830]" lastBlockId="17.[151,1437,1606,2028]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="84B0EAF8FFCBFFDCFF69A5D0AB85F92F" bold="true" box="[151,384,1697,1722]" pageId="16" pageNumber="17">Material examined.</emphasis>
<quantity id="713C9B0FFFCBFFDCFE78A5D0ABDEF92C" box="[390,475,1697,1721]" metricMagnitude="0" metricUnit="m" metricValue="5.9182" pageId="16" pageNumber="17" unit="in" value="233.0">233 in</quantity>
alcohol.
<collectingCountry id="CED3767AFFCBFFDCFDBCA5D0A8D1F92C" box="[578,724,1697,1721]" name="South Africa" pageId="16" pageNumber="17">South Africa</collectingCountry>
, Western Cape, Knysna district, Diepwalle Forest near Forestry Station,
<quantity id="713C9B0FFFCBFFDCFF0BA5B4AB45F948" box="[245,320,1733,1757]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="16" pageNumber="17" unit="m" value="520.0">520 m</quantity>
,
<date id="C27A102AFFCBFFDCFEB0A5B4A80BF948" box="[334,526,1733,1757]" pageId="16" pageNumber="17" value="1979-12-12" valueMax="1979-12-16" valueMin="1979-12-12">1216 Dec 1979</date>
, S. &amp; J. Peck, in malaise trap. Specimen 1 (
<collectionCode id="D0D5AE2FFFCBFFDCFBF7A5B4AE61F948" box="[1033,1124,1733,1757]" pageId="16" pageNumber="17">BMNH</collectionCode>
): Male genitalia prep. G.W. Gibbs G1001 (
<collectionCode id="D0D5AE2FFFCBFFDCFEBAA598AB99F894" box="[324,412,1769,1793]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="16" pageNumber="17">ZMUC</collectionCode>
). Attempted
<collectionCode id="D0D5AE2FFFCBFFDCFDCDA59BA875F897" LSID="urn:lsid:biocol.org:col:15536" box="[563,624,1770,1794]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="16" pageNumber="17">DNA</collectionCode>
extraction of metathorax (by D.C. Lees):
<collectionCode id="D0D5AE2FFFCBFFDCFB9DA598AEBFF894" box="[1123,1210,1769,1793]" pageId="16" pageNumber="17">BMNH</collectionCode>
(E) # 678474, plate MSL079: well E2. Specimen 1: (
<collectionCode id="D0D5AE2FFFCBFFDCFDF9A47CA85AF8B0" box="[519,607,1805,1829]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="16" pageNumber="17">ZMUC</collectionCode>
): Attempted
<collectionCode id="D0D5AE2FFFCBFFDCFD11A47FA929F8B3" LSID="urn:lsid:biocol.org:col:15536" box="[751,812,1806,1830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="16" pageNumber="17">DNA</collectionCode>
extraction of thoracic muscles (by Y. Kobayashi): AG1
<emphasis id="84B0EAF8FFCAFFDDFF39A537AB5CF9CA" bold="true" box="[199,345,1606,1631]" pageId="17" pageNumber="18">Description.</emphasis>
The only available specimens are preserved in alcohol and lack scales and pigmentation.
</paragraph>
<caption id="E2BB6662FFCAFFDDFF69A6B7A8A0F98F" httpUri="https://zenodo.org/record/203562/files/figure.png" pageId="17" pageNumber="18" targetBox="[259,1325,201,1478]" targetPageId="17">
<paragraph id="B67B36EAFFCAFFDDFF69A6B7A8A0F98F" blockId="17.[151,1436,1478,1562]" pageId="17" pageNumber="18">
<emphasis id="84B0EAF8FFCAFFDDFF69A6B7AB17FA49" bold="true" box="[151,274,1478,1501]" pageId="17" pageNumber="18">FIGURE 9.</emphasis>
<taxonomicName id="71C44D69FFCAFFDDFEE6A6B6ABA7FA48" box="[280,418,1479,1501]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFCAFFDDFEE6A6B6ABA7FA48" box="[280,418,1479,1501]" italics="true" pageId="17" pageNumber="18">Agrionympha</emphasis>
</taxonomicName>
male genitalia. AD,
<taxonomicName id="71C44D69FFCAFFDDFD7FA6B9A8D2FA48" box="[641,727,1479,1501]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="84B0EAF8FFCAFFDDFD7FA6B9A8D2FA48" box="[641,727,1479,1501]" italics="true" pageId="17" pageNumber="18">A. karoo</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="9F835783FFCAFFDDFD1AA6B6A931FA48" box="[740,820,1479,1501]" pageId="17" pageNumber="18" rank="species">
<emphasis id="84B0EAF8FFCAFFDDFD1AA6B6A92BFA49" bold="true" box="[740,814,1479,1500]" pageId="17" pageNumber="18">sp. nov</emphasis>
.
</taxonomicNameLabel>
Toorberg: A, lateral, inset ventral view of median plate; B, dorsal; C, phallus lateral; D, phallus dorsal. EH,
<taxonomicName id="71C44D69FFCAFFDDFD6AA697A914FA69" box="[660,785,1510,1532]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kroonella" status="sp. nov.">
<emphasis id="84B0EAF8FFCAFFDDFD6AA697A914FA69" box="[660,785,1510,1532]" italics="true" pageId="17" pageNumber="18">A. kroonella</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="9F835783FFCAFFDDFCE0A697A96AFA6E" box="[798,879,1510,1531]" pageId="17" pageNumber="18" rank="species">
<emphasis id="84B0EAF8FFCAFFDDFCE0A697A96CFA6E" bold="true" box="[798,873,1510,1531]" pageId="17" pageNumber="18">sp. nov</emphasis>
.
</taxonomicNameLabel>
Mariepskop: E, lateral, inset ventral view of median plate; F, dorsal; G, phallus lateral; H, phallus dorsal.
</paragraph>
</caption>
<paragraph id="B67B36EAFFCAFFDDFF39A51AA974F916" blockId="17.[151,1437,1606,2028]" box="[199,881,1643,1667]" pageId="17" pageNumber="18">
<emphasis id="84B0EAF8FFCAFFDDFF39A51AAB01F916" box="[199,260,1643,1667]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
: Antennae: decolourised; 4344 flagellomeres in male.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFCAFFDDFF39A5FEAF64F852" pageId="17" pageNumber="18" type="description">
<paragraph id="B67B36EAFFCAFFDDFF39A5FEA859F932" blockId="17.[151,1437,1606,2028]" box="[199,604,1679,1703]" pageId="17" pageNumber="18">
<emphasis id="84B0EAF8FFCAFFDDFF39A5FEAB1DF932" box="[199,280,1679,1703]" italics="true" pageId="17" pageNumber="18">Thorax</emphasis>
: FW length
<quantity id="713C9B0FFFCAFFDDFE5AA5FEABFFF932" box="[420,506,1679,1703]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="17" pageNumber="18" unit="mm" value="3.1">3.1 mm</quantity>
in male.
</paragraph>
<paragraph id="B67B36EAFFCAFFDDFF39A5C2A8B2F8EA" blockId="17.[151,1437,1606,2028]" pageId="17" pageNumber="18">
<emphasis id="84B0EAF8FFCAFFDDFF39A5C2ABA7F95E" box="[199,418,1715,1739]" italics="true" pageId="17" pageNumber="18">Male postabdomen</emphasis>
: (
<figureCitation id="2EFF2A6FFFCAFFDDFE41A5C2A802F95E" box="[447,519,1715,1739]" captionStart="FIGURE 10" captionStartId="18.[151,250,1913,1936]" captionTargetBox="[313,1332,581,1845]" captionTargetId="figure@18.[244,1340,573,1969]" captionTargetPageId="18" captionText="FIGURE 10. Agrionympha male genitalia. A B, A. pseliacma, Meyrick, Karkloof Falls: A, lateral view, from slide 3830, Transvaal Museum, Pretoria; B, detail of phallus, lateral. C G, A. sagittella, sp. nov. Hogsback: C, lateral; D, dorsal; E, ventral; F, phallus, lateral; G, phallus dorsal. H J, Agrionympha species 1 , Diepwalle Forest: H, lateral; I, phallus lateral; J, phallus dorsal." httpUri="https://zenodo.org/record/203563/files/figure.png" pageId="17" pageNumber="18">Fig.10</figureCitation>
HJ) Sternum VIII remnants irregularly oval, bearing about 12 minute setae. Anteroventral margin of IX sclerotisation broadly rounded, unthickened. Median plate narrow, tapered anteriorly, lacking a keel and unpigmented. Valvae moderately long (length:width ratio 4.6). About 35 hair-sensilla located in membraneous dorsum of segment IX. Phallus short, ratio length:height 4.9; neck region not constricted; gonopore semicircular, a little over half (0.59) width between lappets with proximal margin straight, a distinctive flap-like overhang in the mid-line; ventral bulb moderate.
</paragraph>
<paragraph id="B67B36EAFFCAFFDDFF39A4FAA867F836" blockId="17.[151,1437,1606,2028]" box="[199,610,1931,1955]" pageId="17" pageNumber="18">
<emphasis id="84B0EAF8FFCAFFDDFF39A4FAABB2F836" box="[199,439,1931,1955]" italics="true" pageId="17" pageNumber="18">Female postabdomen</emphasis>
: sex unknown.
</paragraph>
<paragraph id="B67B36EAFFCAFFDDFF3BA4DFAF64F852" blockId="17.[151,1437,1606,2028]" box="[197,1377,1966,1991]" pageId="17" pageNumber="18">
<emphasis id="84B0EAF8FFCAFFDDFF3BA4DFAB4DF852" bold="true" box="[197,328,1966,1991]" pageId="17" pageNumber="18">Bionomics.</emphasis>
Malaise-trapped in montane tall indigenous forest with
<taxonomicName id="71C44D69FFCAFFDDFC42A4DEAEA1F852" box="[956,1188,1967,1991]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="falcatus">
<emphasis id="84B0EAF8FFCAFFDDFC42A4DEAEA1F852" box="[956,1188,1967,1991]" italics="true" pageId="17" pageNumber="18">Podocarpus falcatus</emphasis>
</taxonomicName>
and
<emphasis id="84B0EAF8FFCAFFDDFB24A4C1AF64F852" box="[1242,1377,1967,1991]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="71C44D69FFCAFFDDFB24A4C1AF59F852" box="[1242,1372,1967,1991]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="latifolius">P. latifolius</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFCAFFDDFF3BA4A3AECBF87E" box="[197,1230,2002,2028]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="B67B36EAFFCAFFDDFF3BA4A3AECBF87E" blockId="17.[151,1437,1606,2028]" box="[197,1230,2002,2028]" pageId="17" pageNumber="18">
<emphasis id="84B0EAF8FFCAFFDDFF3BA4A3AB5BF87E" bold="true" box="[197,350,2002,2027]" pageId="17" pageNumber="18">Distribution.</emphasis>
<collectingCountry id="CED3767AFFCAFFDDFE9BA4A2ABFDF87E" box="[357,504,2003,2027]" name="South Africa" pageId="17" pageNumber="18">South Africa</collectingCountry>
, Western Cape, near Knysna. Only known from a single locality.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFC9FFDEFF3BA3E9A834FD8D" pageId="18" pageNumber="19" type="discussion">
<paragraph id="B67B36EAFFC9FFDEFF3BA3E9A834FD8D" blockId="18.[151,1436,151,536]" pageId="18" pageNumber="19">
<emphasis id="84B0EAF8FFC9FFDEFF3BA3E9AB3CFF25" bold="true" box="[197,313,152,176]" pageId="18" pageNumber="19">Remarks.</emphasis>
The male genitalia are separable from those of the named species by the modest number (but&gt;30) of hair-sensilla on dorsum IX, combined with the large gonopore size and absence of constricted neck on distal phallus; pertinent phallus characters cannot be compared in
<taxonomicName id="71C44D69FFC9FFDEFCBAA391A9B3FF6D" box="[836,950,224,248]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFC9FFDEFCBAA391A9B3FF6D" box="[836,950,224,248]" italics="true" pageId="18" pageNumber="19">pseliacma</emphasis>
</taxonomicName>
, from which male 1 appears to differ (in lateral projection) by the more narrow/elongate sternum VIII remnants, the contour of the segment IX front margin (concave rather than convex towards venter), the more parallel-sided valves and the more regular outline of the dorsum X lobe. However, we cannot on morphological grounds ascertain that these completely denuded male specimens are not conspecific with one of the female-onlyspecies
<taxonomicName id="71C44D69FFC9FFDEFC8EA201A9CDFE1D" box="[880,968,368,392]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="jansella">
<emphasis id="84B0EAF8FFC9FFDEFC8EA201A9CDFE1D" box="[880,968,368,392]" italics="true" pageId="18" pageNumber="19">jansella</emphasis>
</taxonomicName>
,
<taxonomicName id="71C44D69FFC9FFDEFC27A200AE03FE1D" box="[985,1030,369,392]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFC9FFDEFC27A200AE03FE1D" box="[985,1030,369,392]" italics="true" pageId="18" pageNumber="19">vari</emphasis>
</taxonomicName>
and
<taxonomicName id="71C44D69FFC9FFDEFBBFA201AEB8FE1D" box="[1089,1213,368,392]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pseudovari">
<emphasis id="84B0EAF8FFC9FFDEFBBFA201AEB8FE1D" box="[1089,1213,368,392]" italics="true" pageId="18" pageNumber="19">pseudovari</emphasis>
</taxonomicName>
(of which only the last-mentioned is also recorded from Western Cape). Moreover, even if molecular markers render/could render such conspecificity unlikely, we decided against naming a taxon on the basis of this material. It is an important background for this decision that the specimens in question do not add significantly to the known spectrum of structural diversity within the genus.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,160 @@
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<mods:title id="4D605440AB566E3F79B1DADA16431919">Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)</mods:title>
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<treatment id="3E6D87FCFFD0FFC0FF69A477A8B3FABA" ID-DOI="http://doi.org/10.5281/zenodo.6189663" ID-GBIF-Taxon="119388876" ID-Zenodo-Dep="6189663" LSID="urn:lsid:plazi:treatment:3E6D87FCFFD0FFC0FF69A477A8B3FABA" httpUri="http://treatment.plazi.org/id/3E6D87FCFFD0FFC0FF69A477A8B3FABA" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="FEDE6561FFD0FFC7FF69A477ABDBF8D4" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="B67B36EAFFD0FFC7FF69A477A86AF8B5" blockId="11.[151,623,1798,1857]" box="[151,623,1798,1824]" pageId="11" pageNumber="12">
<heading id="ED338186FFD0FFC7FF69A477A86AF8B5" bold="true" box="[151,623,1798,1824]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="84B0EAF8FFD0FFC7FF69A477A86AF8B5" bold="true" box="[151,623,1798,1824]" pageId="11" pageNumber="12">
<taxonomicName id="71C44D69FFD0FFC7FF69A477A80FF8B5" authority="Gibbs" authorityName="Gibbs" box="[151,522,1798,1824]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kroonella" status="sp. nov.">
<emphasis id="84B0EAF8FFD0FFC7FF69A477ABBDF8B5" bold="true" box="[151,440,1798,1824]" italics="true" pageId="11" pageNumber="12">Agrionympha kroonella</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="9F835783FFD0FFC7FDEFA477A86AF8B5" box="[529,623,1798,1824]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B67B36EAFFD0FFC7FF69A458ABDBF8D4" blockId="11.[151,623,1798,1857]" box="[151,478,1833,1857]" pageId="11" pageNumber="12">
(
<figureCitation id="2EFF2A6FFFD0FFC7FF61A458AAECF8D4" box="[159,233,1833,1857]" captionStart="FIGURE 1" captionStartId="7.[151,250,1973,1996]" captionTargetBox="[206,1390,1043,1943]" captionTargetId="figure@7.[196,1401,1035,1968]" captionTargetPageId="7" captionText="FIGURE 1. Adult moths from life: A, Agrionympha capensis Whalley, Storms River; B, A. kroonella, sp. nov. Mariepskop; C, A. fuscoapicella, sp. nov. Hogsback; D, A. sagittella, sp. nov. Hogsback. Scale bar = 1 mm." httpUri="https://zenodo.org/record/203554/files/figure.png" pageId="11" pageNumber="12">Figs. 1</figureCitation>
B, 2E, 4D, 9EH, 11)
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD0FFC7FF69A41DAE28F865" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="B67B36EAFFD0FFC7FF69A41DAE28F865" blockId="11.[151,1436,1899,2032]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF69A41DAB87F810" bold="true" box="[151,386,1900,1925]" pageId="11" pageNumber="12">Material examined.</emphasis>
<emphasis id="84B0EAF8FFD0FFC7FE72A41DABF7F811" box="[396,498,1900,1924]" italics="true" pageId="11" pageNumber="12">
<typeStatus id="697F8848FFD0FFC7FE72A41DABF7F811" box="[396,498,1900,1924]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
3,
<collectingCountry id="CED3767AFFD0FFC7FDDCA41DA8B2F811" box="[546,695,1900,1924]" name="South Africa" pageId="11" pageNumber="12">South Africa</collectingCountry>
, Transvaal, Mariepskop, 24º34ʹS 32º52ʹE,
<quantity id="713C9B0FFFD0FFC7FB5FA41DAEFFF811" box="[1185,1274,1900,1924]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="11" pageNumber="12" unit="m" value="1680.0">1680 m</quantity>
,
<date id="C27A102AFFD0FFC7FAF7A41DAF9DF811" box="[1289,1432,1900,1924]" pageId="11" pageNumber="12" value="1986-01-29">29 Jan 1986</date>
, G.W. Gibbs &amp; D.M. Kroon (
<collectionCode id="D0D5AE2FFFD0FFC7FE2DA4E1A816F83D" box="[467,531,1936,1960]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="11" pageNumber="12">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFD0FFC7FDDAA4E0A89AF83D" box="[548,671,1937,1960]" italics="true" pageId="11" pageNumber="12">
<typeStatus id="697F8848FFD0FFC7FDDAA4E0A89CF83D" box="[548,665,1937,1960]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
:
</emphasis>
63 2Ƥ, same data as above (
<collectionCode id="D0D5AE2FFFD0FFC7FC18A4E1AE3EF83D" box="[998,1083,1936,1960]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="11" pageNumber="12">ZMUC</collectionCode>
,
<collectionCode id="D0D5AE2FFFD0FFC7FBB8A4E1AE81F83D" box="[1094,1156,1936,1960]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="11" pageNumber="12">TMP</collectionCode>
). Male genital preps N.P. Kristensen NPK1033 (
<collectionCode id="D0D5AE2FFFD0FFC7FE6DA4C5ABEFF859" box="[403,490,1972,1996]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="11" pageNumber="12">ZMUC</collectionCode>
); G.W. Gibbs G1003, G1007 (
<collectionCode id="D0D5AE2FFFD0FFC7FCC9A4C5A973F859" box="[823,886,1972,1996]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="11" pageNumber="12">TMP</collectionCode>
); Female genital preps N.P.Kristensen NPK1029; G.W. Gibbs G1022 (
<collectionCode id="D0D5AE2FFFD0FFC7FE84A4A9ABBFF865" box="[378,442,2008,2032]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="11" pageNumber="12">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFD0FFC7FE32A4A9A876F865" box="[460,627,2008,2032]" italics="true" pageId="11" pageNumber="12">Other material</emphasis>
: 13 1Ƥ, from same locality in alcohol.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD7FFC0FF3BA3E6AF16FB9A" pageId="12" pageNumber="13" type="description">
<paragraph id="B67B36EAFFD7FFC0FF3BA3E6AF17FE89" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF3BA3E6AB52FF25" bold="true" box="[197,343,151,176]" pageId="12" pageNumber="13">Description.</emphasis>
<emphasis id="84B0EAF8FFD7FFC0FEA3A3E9AB9CFF25" box="[349,409,152,176]" italics="true" pageId="12" pageNumber="13">Head</emphasis>
: Antennae pale yellowish-brown with a grey tip of 15 flagellomeres in male,
<quantity id="713C9B0FFFD7FFC0FAFDA3E6AF2BFF3A" box="[1283,1326,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="13" unit="in" value="2.0">2 in</quantity>
female; a total of 3940 flagellomeres in male with basal 2 scale-covered,
<quantity id="713C9B0FFFD7FFC0FC74A3CDA9F5FF41" box="[906,1008,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="7.239" metricValueMax="7.366" metricValueMin="7.112" pageId="12" pageNumber="13" unit="in" value="28.5" valueMax="29.0" valueMin="28.0">2829 in</quantity>
female with basal 5 scaled. Piliform scales on scape, pedicel and scales of first few flagellomeres dark blackish-brown, strongly contrasting with main antennal colour. Head piliform scales creamy-white, paler than antennae, darkening along the midline.
</paragraph>
<paragraph id="B67B36EAFFD7FFC0FF39A259AB26FD82" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF39A259AB1DFED5" box="[199,280,296,320]" italics="true" pageId="12" pageNumber="13">Thorax</emphasis>
: Tegulae with silvery-white scales; mesonotum black-scaled with purple iridescence. Legs with all femora silvery-white, darker at distal joints; foretibia white with a black anterior face, midtibia white, hindtibia mainly black; tarsi virtually all black but with proximal ends of tarsomeres 1 and 2 of midtarsus whitish. FW length of male and female
<quantity id="713C9B0FFFD7FFC0FE89A2E5ABC8FE39" box="[375,461,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="13" unit="mm" value="3.2">3.2 mm</quantity>
(3.13.3). FW maculation (
<figureCitation id="2EFF2A6FFFD7FFC0FCF9A2E5A94FFE39" box="[775,842,404,428]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
E) with a strong arrow-shaped claval mark, broadening distally and terminating obliquely at about one third; a small triangular remnant of the median band on the dorsum; an uninterrupted postmedian band, tapering towards the dorsum end; a broad apical band which is slightly subterminal.
</paragraph>
<paragraph id="B67B36EAFFD7FFC0FF39A155ABE9FCA2" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF39A155ABA4FDA9" box="[199,417,548,572]" italics="true" pageId="12" pageNumber="13">Male postabdomen</emphasis>
(
<figureCitation id="2EFF2A6FFFD7FFC0FE4BA155ABF9FDA9" box="[437,508,548,572]" captionStart="FIGURE 9" captionStartId="17.[151,250,1478,1501]" captionTargetBox="[259,1325,201,1478]" captionTargetId="figure@17.[254,1332,193,1527]" captionTargetPageId="17" captionText="FIGURE 9. Agrionympha male genitalia. A D, A. karoo, sp. nov. Toorberg: A, lateral, inset ventral view of median plate; B, dorsal; C, phallus lateral; D, phallus dorsal. E H, A. kroonella, sp. nov. Mariepskop: E, lateral, inset ventral view of median plate; F, dorsal; G, phallus lateral; H, phallus dorsal." httpUri="https://zenodo.org/record/203562/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
E, F): Sternum VIII remnants irregularly delimited, somewhat rhomboidal with about 4 small setae. Anteroventral margin of IX sclerotisation broadly rounded and thickened. Median plate broad, subrectangular with rounded corners, anterior part lightly sclerotised. Valvae of moderate length (ratio length:width 4.0), gently curved upward and tapering. Dorsum of IX with about 80 hair-sensilla between the dorsal arms of the vinculum. Phallus very thick and broad, ratio length:height 4.0; distal phallus lacking a constricted neck region in dorsal view; gonopore orientated anterodorsally, very wide and gaping, almost as wide as widest part of phallus; a distinctive heavily darkened body within centre of aedeagus at the level of the lappets; ventral bulb large (male genital preps. G1003, G1007).
</paragraph>
<paragraph id="B67B36EAFFD7FFC0FF39A035AF16FB9A" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF39A035ABC7FCC9" box="[199,450,836,860]" italics="true" pageId="12" pageNumber="13">Female postabdomen:</emphasis>
Segment V gland
<quantity id="713C9B0FFFD7FFC0FD66A035A8EAFCC9" box="[664,751,836,860]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="13" unit="mm" value="0.4">0.4 mm</quantity>
diam., with about 10 fluted sales, three of which arise from the discal area of the protuberance. Segment X approximately equidimensional, its sclerites rounded. Papilla forming a thickened collar region between genital chamber and bursa, its length equalling its width, anterior and posterior margins not strongly reflexed but dorsal side deeply cut away for passage of the thick-walled duct of spermatheca; the latter with its bulbous distal end about 2/3 diameter of ductus length, thin-walled mid section of utriculus shorter than ductus, anterior sac of utriculus
<quantity id="713C9B0FFFD7FFC0FD0DA086A952FB85" box="[755,855,1015,1040]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="12" pageNumber="13" unit="mm" value="0.07">0.07 mm</quantity>
diameter (female genital prep. G1022).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD7FFC0FF3BA76AAE89FBA0" box="[197,1164,1051,1077]" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="B67B36EAFFD7FFC0FF3BA76AAE89FBA0" blockId="12.[151,1437,151,1329]" box="[197,1164,1051,1077]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF3BA76AAB3AFBA1" bold="true" box="[197,319,1051,1076]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
Presence of forewing median band only as a costal spot is unique in genus.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD7FFC0FF3BA731A9E4FB7D" pageId="12" pageNumber="13" type="etymology">
<paragraph id="B67B36EAFFD7FFC0FF3BA731A93AFBE9" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF3BA731AB49FBCD" bold="true" box="[197,332,1088,1112]" pageId="12" pageNumber="13">Etymology.</emphasis>
Named in honour of Dr Doug Kroon whose knowledge and enthusiasm for South African
<taxonomicName id="71C44D69FFD7FFC0FAA1A731AAEEFBE9" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
helped uncover the diversity of micropterigids there.
</paragraph>
<paragraph id="B67B36EAFFD7FFC0FF3BA7F6A9E4FB7D" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF3BA7F6AB4DFB35" bold="true" box="[197,328,1159,1184]" pageId="12" pageNumber="13">Bionomics.</emphasis>
This species occurred amongst moist ferns and low shrubs on a sunny roadside bank in the transition zone between taller forest and higher altitude low fynbos-type vegetation. It was found while GWG and DMK were unsuccessfully searching for
<taxonomicName id="71C44D69FFD7FFC0FDE4A7A1A863FB7D" box="[538,614,1232,1256]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFD7FFC0FDE4A7A1A863FB7D" box="[538,614,1232,1256]" italics="true" pageId="12" pageNumber="13">A. vari</emphasis>
</taxonomicName>
in the vicinity of its
<typeStatus id="697F8848FFD7FFC0FCAEA7A0A985FB7C" box="[848,896,1233,1257]" pageId="12" pageNumber="13">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD7FFC0FF3BA782A8B3FABA" pageId="12" pageNumber="13" type="distribution">
<paragraph id="B67B36EAFFD7FFC0FF3BA782A8B3FABA" blockId="12.[151,1437,151,1329]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF3BA782AB5BFA99" bold="true" box="[197,350,1267,1292]" pageId="12" pageNumber="13">Distribution.</emphasis>
<collectingCountry id="CED3767AFFD7FFC0FE9BA785ABFCFA99" box="[357,505,1268,1292]" name="South Africa" pageId="12" pageNumber="13">South Africa</collectingCountry>
, Mpumalanga Province, formerly Transvaal. Only known from the
<typeStatus id="697F8848FFD7FFC0FB0DA784AF26FA98" box="[1267,1315,1269,1293]" pageId="12" pageNumber="13">type</typeStatus>
locality in the Drakensberg Ranges at 24º34ʹS and 32º52ʹE.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,157 @@
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<mods:title id="F6DF89AE8F732D488522B206A0CCB6F2">Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)</mods:title>
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<treatment id="3E6D87FCFFD1FFC7FF69A456AF8BFB29" ID-DOI="http://doi.org/10.5281/zenodo.6189661" ID-GBIF-Taxon="119388875" ID-Zenodo-Dep="6189661" LSID="urn:lsid:plazi:treatment:3E6D87FCFFD1FFC7FF69A456AF8BFB29" httpUri="http://treatment.plazi.org/id/3E6D87FCFFD1FFC7FF69A456AF8BFB29" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="FEDE6561FFD1FFC6FF69A456AB8EF8F7" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="B67B36EAFFD1FFC6FF69A456A844F8D4" blockId="10.[151,577,1831,1890]" box="[151,577,1831,1857]" pageId="10" pageNumber="11">
<heading id="ED338186FFD1FFC6FF69A456A844F8D4" bold="true" box="[151,577,1831,1857]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="84B0EAF8FFD1FFC6FF69A456A844F8D4" bold="true" box="[151,577,1831,1857]" pageId="10" pageNumber="11">
<taxonomicName id="71C44D69FFD1FFC6FF69A456ABD9F8D4" authority="Gibbs" authorityName="Gibbs" box="[151,476,1831,1857]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="84B0EAF8FFD1FFC6FF69A456AB8FF8D4" bold="true" box="[151,394,1831,1857]" italics="true" pageId="10" pageNumber="11">Agrionympha karoo</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="9F835783FFD1FFC6FE1AA456A844F8D4" box="[484,577,1831,1857]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B67B36EAFFD1FFC6FF69A43BAB8EF8F7" blockId="10.[151,577,1831,1890]" box="[151,395,1866,1890]" pageId="10" pageNumber="11">
(
<figureCitation id="2EFF2A6FFFD1FFC6FF61A43BAAE8F8F7" box="[159,237,1866,1890]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="10" pageNumber="11">Figs. 2</figureCitation>
B, 9AD, 11.)
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD1FFC6FF69A4E3A960F867" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="B67B36EAFFD1FFC6FF69A4E3A960F867" blockId="10.[151,1436,1938,2035]" pageId="10" pageNumber="11">
<emphasis id="84B0EAF8FFD1FFC6FF69A4E3AB85F83E" bold="true" box="[151,384,1938,1963]" pageId="10" pageNumber="11">Material examined.</emphasis>
<emphasis id="84B0EAF8FFD1FFC6FE78A4E2ABE8F83E" box="[390,493,1939,1963]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="697F8848FFD1FFC6FE78A4E2ABE8F83E" box="[390,493,1939,1963]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
3,
<collectingCountry id="CED3767AFFD1FFC6FDE8A4E3A8ACF83F" box="[534,681,1938,1962]" name="South Africa" pageId="10" pageNumber="11">South Africa</collectingCountry>
, Eastern Cape, Great
<taxonomicName id="71C44D69FFD1FFC6FC67A4E2A9DAF83F" box="[921,991,1939,1962]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="20" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">Karoo</taxonomicName>
near Graaff Reinet, Swart River Valley, SW slope of Toorberg, 32º 11ʹS, 24º 0 3ʹE,
<quantity id="713C9B0FFFD1FFC6FD7AA4C6A8DBF85A" box="[644,734,1975,1999]" metricMagnitude="3" metricUnit="m" metricValue="1.607" pageId="10" pageNumber="11" unit="m" value="1607.0">1607 m</quantity>
,
<date id="C27A102AFFD1FFC6FD15A4C6A974F85A" box="[747,881,1975,1999]" pageId="10" pageNumber="11" value="2004-02-09">9 Feb 2004</date>
, G.W. Gibbs &amp; D.M. Kroon. (male genital prep. G1024,
<collectionCode id="D0D5AE2FFFD1FFC6FF0FA4ABAB2AF867" box="[241,303,2010,2034]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="10" pageNumber="11">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFD1FFC6FEBCA4ADABACF866" box="[322,425,2012,2035]" italics="true" pageId="10" pageNumber="11">
<typeStatus id="697F8848FFD1FFC6FEBCA4ADABACF866" box="[322,425,2012,2035]" pageId="10" pageNumber="11" type="paratype">Paratype</typeStatus>
</emphasis>
Ƥ from same site,
<date id="C27A102AFFD1FFC6FD80A4ABA90FF867" box="[638,778,2010,2034]" pageId="10" pageNumber="11" value="2004-02-10">10 Feb 2004</date>
(
<collectionCode id="D0D5AE2FFFD1FFC6FCE7A4ABA95CF867" box="[793,857,2010,2034]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="10" pageNumber="11">TMP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD0FFC7FF39A3E6A852FCC2" pageId="11" pageNumber="12" type="description">
<paragraph id="B67B36EAFFD0FFC7FF39A3E6AB69FF62" blockId="11.[151,1437,151,1213]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF39A3E6AB5CFF25" bold="true" box="[199,345,151,176]" pageId="11" pageNumber="12">Description.</emphasis>
<emphasis id="84B0EAF8FFD0FFC7FE9FA3E9AB98FF25" box="[353,413,152,176]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
. Antennae black throughout; 36 flagellomeres in male, basal 1 or 2 with black scales; 28 flagellomeres in female, basal 8 with scales. Piliform scales on scape and pedicel black. Head piliform scales offwhite, grey-tipped.
</paragraph>
<paragraph id="B67B36EAFFD0FFC7FF39A272AB0CFD80" blockId="11.[151,1437,151,1213]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF39A272AB1DFE8E" box="[199,280,259,283]" italics="true" pageId="11" pageNumber="12">Thorax</emphasis>
. Tegulae and dorsum black-scaled with coppery reflections. Sides of thorax silvery-white scaled, forecoxa dark grey anteriorly but with a few silver scales laterally; femora pale grey with dark band distally; tibia and tarsi entirely black except for a pale grey band on midtarsomere 3. FW length
<quantity id="713C9B0FFFD0FFC7FBF3A23BAE66FEF7" box="[1037,1123,330,354]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="12" unit="mm" value="2.5">2.5 mm</quantity>
in male,
<quantity id="713C9B0FFFD0FFC7FB33A23BAF21FEF7" box="[1229,1316,330,354]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="11" pageNumber="12" unit="mm" value="2.6">2.6 mm</quantity>
in female. FWs (
<figureCitation id="2EFF2A6FFFD0FFC7FF25A21FAB18FE13" box="[219,285,366,390]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="11" pageNumber="12">Fig. 2</figureCitation>
B) iridescent coppery dark-grey, maculation with all 4 silvery-white bands present, bordered with black scales: a slightly crescentic oblique claval mark; a strongly kinked continuous median band, widest at costa, interrupted on R and M by thin lines of black scales, most prominent in female; a near-straight postmedian line, broad, uininterrupted in male but with black scales along M in female; an equally broad subapical line; apex black. HWs dark-grey.
</paragraph>
<paragraph id="B67B36EAFFD0FFC7FF39A150AAF1FCA6" blockId="11.[151,1437,151,1213]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF39A150ABA5FDAC" box="[199,416,545,569]" italics="true" pageId="11" pageNumber="12">Male postabdomen</emphasis>
(
<figureCitation id="2EFF2A6FFFD0FFC7FE4AA150ABFFFDAC" box="[436,506,545,569]" captionStart="FIGURE 9" captionStartId="17.[151,250,1478,1501]" captionTargetBox="[259,1325,201,1478]" captionTargetId="figure@17.[254,1332,193,1527]" captionTargetPageId="17" captionText="FIGURE 9. Agrionympha male genitalia. A D, A. karoo, sp. nov. Toorberg: A, lateral, inset ventral view of median plate; B, dorsal; C, phallus lateral; D, phallus dorsal. E H, A. kroonella, sp. nov. Mariepskop: E, lateral, inset ventral view of median plate; F, dorsal; G, phallus lateral; H, phallus dorsal." httpUri="https://zenodo.org/record/203562/files/figure.png" pageId="11" pageNumber="12">Fig. 9</figureCitation>
AD): Sternum VIII remnants roughly trapezoidal, bearing about 6 small setae. Anteroventral margin of IX sclerotisation extended into a very broad rounded apex, markedly thickened but not darkened. Median plate tapering to a sharp point, unevenly sclerotised with a central keel. Valvae of moderate length (ratio length:width 3.8), angled upwards and distinctly elbowed on mesal aspect in ventral view. A row of about 11 hair-sensilla on the membraneous dorsum of IX. Phallus moderately elongate, ratio length:height 4.5; aedeagus with a constricted neck region in dorsal view; gonopore moderate, slightly less than half width of phallus at lappets, heart-shaped with its proximal border raised on a prominence; ventral bulb large (male genital prep. G1024).
</paragraph>
<paragraph id="B67B36EAFFD0FFC7FF39A04EA852FCC2" blockId="11.[151,1437,151,1213]" box="[199,599,831,855]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF39A04EABB2FCC2" box="[199,439,831,855]" italics="true" pageId="11" pageNumber="12">Female postabdomen</emphasis>
not dissected.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD0FFC7FF3BA013ABC1FC0A" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="B67B36EAFFD0FFC7FF3BA013ABC1FC0A" blockId="11.[151,1437,151,1213]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF3BA013AB3AFCEE" bold="true" box="[197,319,866,891]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Forewing with coexistence of discrete and complete median band with similarly complete postmedian band unique in genus.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD0FFC7FF3BA0DBA814FB0D" pageId="11" pageNumber="12" type="etymology">
<paragraph id="B67B36EAFFD0FFC7FF3BA0DBA998FC57" blockId="11.[151,1437,151,1213]" box="[197,925,938,962]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF3BA0DBAB49FC57" bold="true" box="[197,332,938,962]" pageId="11" pageNumber="12">Etymology.</emphasis>
Named after the semi-arid region in which it occurs.
</paragraph>
<paragraph id="B67B36EAFFD0FFC7FF3BA0BCA814FB0D" blockId="11.[151,1437,151,1213]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF3BA0BCAB4DFC73" bold="true" box="[197,328,973,998]" pageId="11" pageNumber="12">Bionomics.</emphasis>
This smallest species of
<taxonomicName id="71C44D69FFD0FFC7FD8BA0BFA908FC73" box="[629,781,974,998]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFD0FFC7FD8BA0BFA908FC73" box="[629,781,974,998]" italics="true" pageId="11" pageNumber="12">Agrionympha</emphasis>
</taxonomicName>
has occurred amongst
<taxonomicName id="71C44D69FFD0FFC7FBDEA0BFAE88FC73" box="[1056,1165,974,998]" class="Polypodiopsida" family="Pteridaceae" genus="Adiantum" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="84B0EAF8FFD0FFC7FBDEA0BFAE88FC73" box="[1056,1165,974,998]" italics="true" pageId="11" pageNumber="12">Adiantum</emphasis>
</taxonomicName>
and other ferns on the rocky bank of stream where seepage maintained very limited growth of thallose liverworts and mosses in an otherwise hot and arid environment. The adults were taken flying at 6.307.00 am while dew remained on the grass. Since all other
<taxonomicName id="71C44D69FFD0FFC7FEBFA748ABDFFBC4" box="[321,474,1081,1105]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFD0FFC7FEBFA748ABDFFBC4" box="[321,474,1081,1105]" italics="true" pageId="11" pageNumber="12">Agrionympha</emphasis>
</taxonomicName>
species, and indeed micropterigids in general, are essentially species of moist forests or forest margins, the existance of
<taxonomicName id="71C44D69FFD0FFC7FDE5A72CA87FFBE0" box="[539,634,1117,1141]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="karoo">
<emphasis id="84B0EAF8FFD0FFC7FDE5A72CA87FFBE0" box="[539,634,1117,1141]" italics="true" pageId="11" pageNumber="12">A. karoo</emphasis>
</taxonomicName>
within this semi-arid region suggests a relictual ecological status in the face of long-term climate change.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD0FFC7FF3BA7D5AF8BFB29" box="[197,1422,1188,1213]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="B67B36EAFFD0FFC7FF3BA7D5AF8BFB29" blockId="11.[151,1437,151,1213]" box="[197,1422,1188,1213]" pageId="11" pageNumber="12">
<emphasis id="84B0EAF8FFD0FFC7FF3BA7D5AB5BFB28" bold="true" box="[197,350,1188,1213]" pageId="11" pageNumber="12">Distribution.</emphasis>
<collectingCountry id="CED3767AFFD0FFC7FE9BA7D5ABFDFB29" box="[357,504,1188,1212]" name="South Africa" pageId="11" pageNumber="12">South Africa</collectingCountry>
, known only from the
<typeStatus id="697F8848FFD0FFC7FD0BA7D4A920FB28" box="[757,805,1189,1213]" pageId="11" pageNumber="12">type</typeStatus>
locality in Eastern Cape, near Graaff Reinet at
<quantity id="713C9B0FFFD0FFC7FAC7A7D5AF8EFB29" box="[1337,1419,1188,1212]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="m" value="1600.0">1600m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,181 @@
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<mods:title id="125E03F33CD0DC61BAFBADE7ED1CE173">Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)</mods:title>
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<mods:namePart id="22855666A5C8BAF921E7EBDDB9C93769">Kristensen, Niels P.</mods:namePart>
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<treatment id="3E6D87FCFFD2FFC5FF69A3E6A9E3F993" ID-DOI="http://doi.org/10.5281/zenodo.6189655" ID-GBIF-Taxon="119388874" ID-Zenodo-Dep="6189655" LSID="urn:lsid:plazi:treatment:3E6D87FCFFD2FFC5FF69A3E6A9E3F993" httpUri="http://treatment.plazi.org/id/3E6D87FCFFD2FFC5FF69A3E6A9E3F993" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="FEDE6561FFD2FFC5FF69A3E6ABB1FF47" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="B67B36EAFFD2FFC5FF69A3E6A89EFF24" blockId="9.[151,667,151,210]" box="[151,667,151,177]" pageId="9" pageNumber="10">
<heading id="ED338186FFD2FFC5FF69A3E6A89EFF24" bold="true" box="[151,667,151,177]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="84B0EAF8FFD2FFC5FF69A3E6A89EFF24" bold="true" box="[151,667,151,177]" pageId="9" pageNumber="10">
<taxonomicName id="71C44D69FFD2FFC5FF69A3E6A833FF24" authority="Gibbs" authorityName="Gibbs" box="[151,566,151,177]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fuscoapicella" status="sp. nov.">
<emphasis id="84B0EAF8FFD2FFC5FF69A3E6ABE1FF24" bold="true" box="[151,484,151,177]" italics="true" pageId="9" pageNumber="10">Agrionympha fuscoapicella</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="9F835783FFD2FFC5FDC3A3E6A89EFF24" box="[573,667,151,177]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B67B36EAFFD2FFC5FF69A3CBABB1FF47" blockId="9.[151,667,151,210]" box="[151,436,186,210]" pageId="9" pageNumber="10">
(
<figureCitation id="2EFF2A6FFFD2FFC5FF61A3CBAAECFF47" box="[159,233,186,210]" captionStart="FIGURE 1" captionStartId="7.[151,250,1973,1996]" captionTargetBox="[206,1390,1043,1943]" captionTargetId="figure@7.[196,1401,1035,1968]" captionTargetPageId="7" captionText="FIGURE 1. Adult moths from life: A, Agrionympha capensis Whalley, Storms River; B, A. kroonella, sp. nov. Mariepskop; C, A. fuscoapicella, sp. nov. Hogsback; D, A. sagittella, sp. nov. Hogsback. Scale bar = 1 mm." httpUri="https://zenodo.org/record/203554/files/figure.png" pageId="9" pageNumber="10">Figs. 1</figureCitation>
C, 2D, 8AD, 11)
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD2FFC5FF69A273AF97FE67" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="B67B36EAFFD2FFC5FF69A273AF97FE67" blockId="9.[151,1437,258,1543]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF69A273AB82FE8E" bold="true" box="[151,391,258,283]" pageId="9" pageNumber="10">Material examined.</emphasis>
<emphasis id="84B0EAF8FFD2FFC5FE6AA272ABFEFE8E" box="[404,507,259,283]" italics="true" pageId="9" pageNumber="10">
<typeStatus id="697F8848FFD2FFC5FE6AA272ABFEFE8E" box="[404,507,259,283]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
</emphasis>
3,
<collectingCountry id="CED3767AFFD2FFC5FDCCA273A8C9FE8F" box="[562,716,258,282]" name="South Africa" pageId="9" pageNumber="10">South Africa</collectingCountry>
, Eastern Cape, Hogsback, Madonna &amp; Child Falls,
<geoCoordinate id="D3F0502DFFD2FFC5FABDA273AF99FE8F" box="[1347,1436,258,282]" direction="south" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="-32.6">32º36´S</geoCoordinate>
<geoCoordinate id="D3F0502DFFD2FFC5FF69A257AAF0FEAB" box="[151,245,294,318]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="26.966667">26º58´E</geoCoordinate>
,
<quantity id="713C9B0FFFD2FFC5FEFCA257AB59FEAB" box="[258,348,294,318]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="10" unit="m" value="1050.0">1050 m</quantity>
,
<date id="C27A102AFFD2FFC5FE94A257ABFBFEAB" box="[362,510,294,318]" pageId="9" pageNumber="10" value="1986-02-12">12 Feb 1986</date>
, G.W. Gibbs (
<collectionCode id="D0D5AE2FFFD2FFC5FD5EA257A8DAFEAB" box="[672,735,294,318]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="9" pageNumber="10">TMP</collectionCode>
). Male genital prep. NPK1035 (
<collectionCode id="D0D5AE2FFFD2FFC5FBAFA257AEADFEAB" box="[1105,1192,294,318]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="10">ZMUC</collectionCode>
).
<typeStatus id="697F8848FFD2FFC5FB43A259AF2AFEAA" box="[1213,1327,296,319]" pageId="9" pageNumber="10" type="paratype">
<emphasis id="84B0EAF8FFD2FFC5FB43A259AF2AFEAA" box="[1213,1327,296,319]" italics="true" pageId="9" pageNumber="10">Paratypes</emphasis>
</typeStatus>
13 11Ƥ,
<collectingCountry id="CED3767AFFD2FFC5FF69A23BAB2DFEF7" box="[151,296,330,354]" name="South Africa" pageId="9" pageNumber="10">South Africa</collectingCountry>
(EC), Hogsback: 2Ƥ, Madonna &amp; Child Falls,
<quantity id="713C9B0FFFD2FFC5FCBBA23BA998FEF7" box="[837,925,330,354]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="10" unit="m" value="1050.0">1050 m</quantity>
,
<date id="C27A102AFFD2FFC5FC55A23BAE39FEF7" box="[939,1084,330,354]" pageId="9" pageNumber="10" value="1986-02-12">12 Feb 1986</date>
, G.W. Gibbs (
<collectionCode id="D0D5AE2FFFD2FFC5FB25A23BAF37FEF7" box="[1243,1330,330,354]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="10">ZMUC</collectionCode>
); 13 7Ƥ, Auckland Reserve track,
<quantity id="713C9B0FFFD2FFC5FE4AA21FA846FE13" box="[436,579,366,390]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="9" pageNumber="10" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">9001000 m</quantity>
,
<date id="C27A102AFFD2FFC5FDAEA21EA8D7FE13" box="[592,722,366,390]" pageId="9" pageNumber="10" value="2004-02-07">7 Feb 2004</date>
, G.W. Gibbs (
<collectionCode id="D0D5AE2FFFD2FFC5FC93A21FA9ADFE13" box="[877,936,366,390]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="9" pageNumber="10">TMP</collectionCode>
,
<collectionCode id="D0D5AE2FFFD2FFC5FC4AA21FAE0DFE13" box="[948,1032,366,390]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="10">ZMUC</collectionCode>
,
<collectionCode id="D0D5AE2FFFD2FFC5FBEAA21FAE68FE13" box="[1044,1133,366,390]" pageId="9" pageNumber="10">BMNH</collectionCode>
); 2Ƥ, Hogsback Pass road,
<quantity id="713C9B0FFFD2FFC5FF69A2E3AB2CFE3F" box="[151,297,402,426]" metricMagnitude="2" metricUnit="m" metricValue="9.9" metricValueMax="10.4" metricValueMin="9.4" pageId="9" pageNumber="10" unit="m" value="990.0" valueMax="1040.0" valueMin="940.0">9401040 m</quantity>
,
<date id="C27A102AFFD2FFC5FEC6A2E3ABF2FE3F" box="[312,503,402,426]" pageId="9" pageNumber="10" value="1986-02-11" valueMax="1986-02-13" valueMin="1986-02-11">1113 Feb 1986</date>
, G.W. Gibbs. Female genital prep. G1016 (
<collectionCode id="D0D5AE2FFFD2FFC5FC0EA2E3AE2AFE3F" box="[1008,1071,402,426]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="9" pageNumber="10">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFD2FFC5FBBBA2E2AEF5FE3E" box="[1093,1264,403,427]" italics="true" pageId="9" pageNumber="10">Other material</emphasis>
:
<collectingCountry id="CED3767AFFD2FFC5FAFCA2E3AF92FE3F" box="[1282,1431,402,426]" name="South Africa" pageId="9" pageNumber="10">South Africa</collectingCountry>
, Eastern Cape. Specimens cited above plus alcohol preserved material from Hogsback: Auckland Reserve (ca
<quantity id="713C9B0FFFD2FFC5FA9AA2C7AAABFE67" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="m" value="1000.0">1000 m</quantity>
) and Kettlespout track (
<quantity id="713C9B0FFFD2FFC5FE45A2ABA811FE67" box="[443,532,474,498]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="10" unit="m" value="1200.0">1200 m</quantity>
),
<date id="C27A102AFFD2FFC5FDD6A2AAA8ACFE67" box="[552,681,474,498]" pageId="9" pageNumber="10" value="2004-02-07">7 Feb 2004</date>
, G.W. Gibbs. Male genital prep. G1017, wing slide G1027 (
<collectionCode id="D0D5AE2FFFD2FFC5FAB9A2ABAF83FE67" box="[1351,1414,474,498]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="9" pageNumber="10">TMP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD2FFC5FF3BA28FAE29FA9F" pageId="9" pageNumber="10" type="description">
<paragraph id="B67B36EAFFD2FFC5FF3BA28FAAC2FDCB" blockId="9.[151,1437,258,1543]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF3BA28FAB52FD82" bold="true" box="[197,343,510,535]" pageId="9" pageNumber="10">Description.</emphasis>
<emphasis id="84B0EAF8FFD2FFC5FEA3A28FAB9CFD83" box="[349,409,510,534]" italics="true" pageId="9" pageNumber="10">Head</emphasis>
: Antennae black throughout; 43 flagellomeres in male with basal 3 scale-covered,
<quantity id="713C9B0FFFD2FFC5FAC7A28FAF99FD83" box="[1337,1436,510,534]" metricMagnitude="-1" metricUnit="m" metricValue="8.254999999999999" metricValueMax="8.382" metricValueMin="8.128" pageId="9" pageNumber="10" unit="in" value="32.5" valueMax="33.0" valueMin="32.0">3233 in</quantity>
female with basal 10 scaled. Piliform scales on scape and pedicel black. Head piliform scales ochreous with darker tips.
</paragraph>
<paragraph id="B67B36EAFFD2FFC5FF39A11AA96BFCA3" blockId="9.[151,1437,258,1543]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF39A11AAB1DFD16" box="[199,280,619,643]" italics="true" pageId="9" pageNumber="10">Thorax</emphasis>
: Tegulae black-scaled, also ventrally on all coxae; mesonotum black with gold iridescence. Hindfemora greyish; fore and midtibiae black on dorsal surface, white beneath with a white band at 0.5; all tarsi black with whitish bands at proximal ends of tarsomeres 1, 2 and 3; except that tarsomere 1 is entirely black on hindleg. FW length 3.0 mm in male,
<quantity id="713C9B0FFFD2FFC5FE53A1A7A803FD7B" box="[429,518,726,750]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="10" unit="mm" value="3.5">3.5 mm</quantity>
in female (3.43.5). FW maculation (
<figureCitation id="2EFF2A6FFFD2FFC5FC42A1A7AE05FD7B" box="[956,1024,726,750]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="9" pageNumber="10">Fig. 2</figureCitation>
D) with a slightly bent claval mark, angled towards dorsum; an almost straight, uninterrupted median band and a small triangular remnant of the postmedian band from the costa to about half; subapical band absent.
</paragraph>
<paragraph id="B67B36EAFFD2FFC5FF39A032AEC0FBA7" blockId="9.[151,1437,258,1543]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF39A032AB98FCCE" box="[199,413,835,859]" italics="true" pageId="9" pageNumber="10">Male postabdomen</emphasis>
(
<figureCitation id="2EFF2A6FFFD2FFC5FE50A033ABE2FCCF" box="[430,487,834,858]" captionStart="FIGURE 8" captionStartId="16.[151,250,1415,1438]" captionTargetBox="[311,1355,490,1414]" captionTargetId="figure@16.[223,1363,483,1474]" captionTargetPageId="16" captionText="FIGURE 8. Agrionympha genitalia. A D, A. fuscoapicella sp. nov. male, Hogsback: A, lateral, inset - ventral view of median plate; B, dorsal, C, phallus lateral; D, phallus dorsal. E F, A. jansella sp. nov. female, Karkloof Falls: E, extended terminal segments of abdomen; F, internal female genitalia." httpUri="https://zenodo.org/record/203561/files/figure.png" pageId="9" pageNumber="10">Fig.8</figureCitation>
AD): Sternum VIII remnants large, trapezoidal, lying closely adjacent to anterolateral edge of vinculum, bearing about 16 small setae. Anteroventral margin of IX sclerotisation broadly rounded and thickened. Median plate narrow, tapered at anterior end with a distinct ventral keel, strongly darkened anteriorly. Valvae of moderate length, with their curvature markedly angled (ratio length:width 4.0). Tegumen lobes with an irregular un-sclerotised patch anteriorly along the mid-dorsal line. A cluster of about 40 hair-sensilla on the membraneous dorsum of IX. Phallus elongate, ratio length:height 5.6; aedeagus with a constricted neck region in dorsal view; gonopore heart-shaped, small, less than half width of aedeagus; ventral bulb moderate
</paragraph>
<paragraph id="B67B36EAFFD2FFC5FF69A74FA858FBC3" blockId="9.[151,1437,258,1543]" box="[151,605,1086,1110]" pageId="9" pageNumber="10">(male genital preps. NPK 1035, G1023).</paragraph>
<paragraph id="B67B36EAFFD2FFC5FF39A712AE29FA9F" blockId="9.[151,1437,258,1543]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF39A712ABC7FBEE" box="[199,450,1123,1147]" italics="true" pageId="9" pageNumber="10">Female postabdomen:</emphasis>
Segment V gland
<quantity id="713C9B0FFFD2FFC5FD67A713A8F4FBEF" box="[665,753,1122,1146]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="10" unit="mm" value="0.4">0.4 mm</quantity>
diam., with about 12 long fluted scales. Segment X equidimensional, the sclerites rounded, often emarginated along the anterior edge. Papilla goblet-shaped with its thick walls strongly reflexed around the anterior margin; thick-walled proximal section of spermatheca with its bulbous end about half the diameter of ductus length, thin-walled mid section of utriculus about the same length as ductus, anterior sac of utriculus
<quantity id="713C9B0FFFD2FFC5FE59A783A80EFA9F" box="[423,523,1266,1290]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="10" unit="mm" value="0.08">0.08 mm</quantity>
diameter (female genital preps. G1015, G1016).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD2FFC5FF3BA667AE17FABB" box="[197,1042,1302,1327]" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="B67B36EAFFD2FFC5FF3BA667AE17FABB" blockId="9.[151,1437,258,1543]" box="[197,1042,1302,1327]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF3BA667AB3AFABA" bold="true" box="[197,319,1302,1327]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Immediately distinctive by absence of forewing subapical band.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD2FFC5FF3BA64AAB8FFA77" pageId="9" pageNumber="10" type="etymology">
<paragraph id="B67B36EAFFD2FFC5FF3BA64AAB1AFAE3" blockId="9.[151,1437,258,1543]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF3BA64AAB49FAC6" bold="true" box="[197,332,1339,1363]" pageId="9" pageNumber="10">Etymology.</emphasis>
From
<emphasis id="84B0EAF8FFD2FFC5FE67A64AABDAFAC6" box="[409,479,1339,1363]" italics="true" pageId="9" pageNumber="10">fuscus</emphasis>
(dark) and
<emphasis id="84B0EAF8FFD2FFC5FD9AA64DA89CFAC6" box="[612,665,1340,1363]" italics="true" pageId="9" pageNumber="10">apex</emphasis>
(tip), referring to the blackish-brown wing apex which lacks a subapical band.
</paragraph>
<paragraph id="B67B36EAFFD2FFC5FF3BA6F3AB8FFA77" blockId="9.[151,1437,258,1543]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF3BA6F3AB4DFA0E" bold="true" box="[197,328,1410,1435]" pageId="9" pageNumber="10">Bionomics.</emphasis>
Occurs in tall dense rainforest, under the canopy but in light wells and margins where forest floor is damp and periphyton present. Sweep-netted by day from low soft annuals and ferns. Females predominate in collections (90%, n=30).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD2FFC5FF3BA69FA9E3F993" box="[197,998,1518,1543]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="B67B36EAFFD2FFC5FF3BA69FA9E3F993" blockId="9.[151,1437,258,1543]" box="[197,998,1518,1543]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF3BA69FAB5BF992" bold="true" box="[197,350,1518,1543]" pageId="9" pageNumber="10">Distribution.</emphasis>
<collectingCountry id="CED3767AFFD2FFC5FE9BA69FABFDF993" box="[357,504,1518,1542]" name="South Africa" pageId="9" pageNumber="10">South Africa</collectingCountry>
, known only from Hogsback, Eastern Cape.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,160 @@
<document id="77DB460BCACC3FC9141454340E0418B9" ID-DOI="10.5281/zenodo.203553" ID-GBIF-Dataset="dd1ad33b-e90a-4c23-a6b7-08d77c43e95f" ID-ISSN="1175-5326" ID-Zenodo-Dep="203553" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460237623643" checkinUser="plazi" docAuthor="Gibbs, George W. &amp; Kristensen, Niels P." docDate="2011" docId="3E6D87FCFFD2FFC6FF69A524AF91FD11" docLanguage="en" docName="zt02764p021.pdf" docOrigin="Zootaxa 2764" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Agrionympha jansella Gibbs, sp. nov." docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="C254FF84FFDBFFCCFFFEA371AA05FF95" masterDocTitle="Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="10" updateTime="1698239897964" updateUser="plazi">
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<mods:title id="DE10A7FE8CE599D5CF203FFA54FFF278">Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)</mods:title>
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<mods:namePart id="7EB2373E46693ED11A86EDC7E5BCFA56">Gibbs, George W.</mods:namePart>
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<mods:namePart id="7AC6D0E30C87F111C696D9A7A8AEB202">Kristensen, Niels P.</mods:namePart>
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<mods:date id="637E8253F53097B259480DA18C708B8C">2011</mods:date>
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<treatment id="3E6D87FCFFD2FFC6FF69A524AF91FD11" ID-DOI="http://doi.org/10.5281/zenodo.6189659" ID-GBIF-Taxon="119388873" ID-Zenodo-Dep="6189659" LSID="urn:lsid:plazi:treatment:3E6D87FCFFD2FFC6FF69A524AF91FD11" httpUri="http://treatment.plazi.org/id/3E6D87FCFFD2FFC6FF69A524AF91FD11" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="FEDE6561FFD2FFC5FF69A524ABADF91A" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="B67B36EAFFD2FFC5FF69A524A85FF9FA" blockId="9.[151,602,1621,1679]" box="[151,602,1621,1647]" pageId="9" pageNumber="10">
<heading id="ED338186FFD2FFC5FF69A524A85FF9FA" bold="true" box="[151,602,1621,1647]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="84B0EAF8FFD2FFC5FF69A524A85FF9FA" bold="true" box="[151,602,1621,1647]" pageId="9" pageNumber="10">
<taxonomicName id="71C44D69FFD2FFC5FF69A524ABF0F9FA" authority="Gibbs" authorityName="Gibbs" box="[151,501,1621,1647]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jansella" status="sp. nov.">
<emphasis id="84B0EAF8FFD2FFC5FF69A524ABA7F9FA" bold="true" box="[151,418,1621,1647]" italics="true" pageId="9" pageNumber="10">Agrionympha jansella</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="9F835783FFD2FFC5FE02A524A85FF9FA" box="[508,602,1621,1647]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B67B36EAFFD2FFC5FF69A506ABADF91A" blockId="9.[151,602,1621,1679]" box="[151,424,1655,1679]" pageId="9" pageNumber="10">
(
<figureCitation id="2EFF2A6FFFD2FFC5FF61A506AAEEF91A" box="[159,235,1655,1679]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="9" pageNumber="10">Figs. 2</figureCitation>
G, 3D, 8E, F, 11)
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD2FFC5FF69A5CBAF4DF963" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="B67B36EAFFD2FFC5FF69A5CBAF4DF963" blockId="9.[151,1436,1721,2034]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF69A5CBAB87F946" bold="true" box="[151,386,1722,1747]" pageId="9" pageNumber="10">Material examined.</emphasis>
<emphasis id="84B0EAF8FFD2FFC5FE74A5CAABF4F946" box="[394,497,1723,1747]" italics="true" pageId="9" pageNumber="10">
<typeStatus id="697F8848FFD2FFC5FE74A5CAABF4F946" box="[394,497,1723,1747]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
</emphasis>
Ƥ,
<collectingCountry id="CED3767AFFD2FFC5FDE0A5CBA8B6F947" box="[542,691,1722,1746]" name="South Africa" pageId="9" pageNumber="10">South Africa</collectingCountry>
, Kwazulu
<collectingCountry id="CED3767AFFD2FFC5FCD3A5CBA969F947" box="[813,876,1722,1746]" name="South Africa" pageId="9" pageNumber="10">Natal</collectingCountry>
, Karkloof Falls, 29º24ʹS 30º17ʹE,
<quantity id="713C9B0FFFD2FFC5FB0AA5CBAF48F947" box="[1268,1357,1722,1746]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="9" pageNumber="10" unit="m" value="1020.0">1020 m</quantity>
,
<date id="C27A102AFFD2FFC5FAA5A5CBAAD7F963" pageId="9" pageNumber="10" value="1986-02-05">5 Feb 1986</date>
, G.W. Gibbs (
<collectionCode id="D0D5AE2FFFD2FFC5FE93A5AFABA9F963" box="[365,428,1758,1782]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="9" pageNumber="10">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFD2FFC5FE40A591A82AF962" box="[446,559,1760,1783]" italics="true" pageId="9" pageNumber="10">
<typeStatus id="697F8848FFD2FFC5FE40A591A82AF962" box="[446,559,1760,1783]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
</emphasis>
3Ƥ, data as above (
<collectionCode id="D0D5AE2FFFD2FFC5FCF3A5AFA94DF963" box="[781,840,1758,1782]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="9" pageNumber="10">TMP</collectionCode>
,
<collectionCode id="D0D5AE2FFFD2FFC5FCAAA5AFA9AFF963" box="[852,938,1758,1782]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="10">ZMUC</collectionCode>
). Female genital prep G1021 (
<collectionCode id="D0D5AE2FFFD2FFC5FB02A5AFAF3EF963" box="[1276,1339,1758,1782]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="9" pageNumber="10">TMP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD2FFC6FF39A473A99EFE12" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="B67B36EAFFD2FFC5FF39A473AE55F83F" blockId="9.[151,1436,1721,2034]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF39A473AB5DF88E" bold="true" box="[199,344,1794,1819]" pageId="9" pageNumber="10">Description.</emphasis>
<emphasis id="84B0EAF8FFD2FFC5FE9FA472AB98F88E" box="[353,413,1795,1819]" italics="true" pageId="9" pageNumber="10">Head</emphasis>
: Antennae black throughout; 3334 flagellomeres in female with basal 11 scale-covered. Piliform scales on scape and pedicel black. Head piliform scales black on dorsum but creamy-white on frons. Thorax tegulae black-scaled and with black scales ventrally on all coxae; mesonotum black-scaled with bronzy iridescence. Hindfemora greyish; fore and midtibiae black with whitish band at 0.5; tarsi black with whitish bands at proximal ends of tarsomeres 1, 2 and 3, except that hindtarsomere 1 is entirely black.
</paragraph>
<paragraph id="B67B36EAFFD2FFC5FF39A4C7AE9AF867" blockId="9.[151,1436,1721,2034]" pageId="9" pageNumber="10">
<emphasis id="84B0EAF8FFD2FFC5FF39A4C7AB1AF85B" box="[199,287,1974,1998]" italics="true" pageId="9" pageNumber="10">Thorax.</emphasis>
FW length (female)
<quantity id="713C9B0FFFD2FFC5FDF3A4C7A862F85B" box="[525,615,1974,1998]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="9" pageNumber="10" unit="mm" value="3.2">3.2 mm</quantity>
. FW maculation (
<figureCitation id="2EFF2A6FFFD2FFC5FCCCA4C7A970F85B" box="[818,885,1974,1998]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="9" pageNumber="10">Fig. 2</figureCitation>
G) lacking a claval mark; a very narrow, slightly convex median band; an incomplete postmedian band from costa to half; a large apical spot.
</paragraph>
<paragraph id="B67B36EAFFD1FFC6FF39A3E9A84DFF3A" blockId="10.[151,1437,151,645]" box="[199,584,151,176]" pageId="10" pageNumber="11">
<emphasis id="84B0EAF8FFD1FFC6FF39A3E9AB99FF25" box="[199,412,152,176]" italics="true" pageId="10" pageNumber="11">Male postabdomen</emphasis>
: sex unknown.
</paragraph>
<paragraph id="B67B36EAFFD1FFC6FF39A3CDA99EFE12" blockId="10.[151,1437,151,645]" pageId="10" pageNumber="11">
<emphasis id="84B0EAF8FFD1FFC6FF39A3CDABC6FF41" box="[199,451,188,212]" italics="true" pageId="10" pageNumber="11">Female postabdomen:</emphasis>
Segment V gland
<quantity id="713C9B0FFFD1FFC6FD61A3CDA8FDFF41" box="[671,760,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="11" unit="mm" value="0.4">0.4 mm</quantity>
diam., with about 10 long fluted scales. Genital segments (
<figureCitation id="2EFF2A6FFFD1FFC6FF61A3AEAAE7FF62" box="[159,226,223,247]" captionStart="FIGURE 8" captionStartId="16.[151,250,1415,1438]" captionTargetBox="[311,1355,490,1414]" captionTargetId="figure@16.[223,1363,483,1474]" captionTargetPageId="16" captionText="FIGURE 8. Agrionympha genitalia. A D, A. fuscoapicella sp. nov. male, Hogsback: A, lateral, inset - ventral view of median plate; B, dorsal, C, phallus lateral; D, phallus dorsal. E F, A. jansella sp. nov. female, Karkloof Falls: E, extended terminal segments of abdomen; F, internal female genitalia." httpUri="https://zenodo.org/record/203561/files/figure.png" pageId="10" pageNumber="11">Fig. 8</figureCitation>
E) with segment X approximately equidimensional, lateral sclerites C-shaped with anterior edge emarginated. Papilla large (
<quantity id="713C9B0FFFD1FFC6FE8DA275ABDFFE89" box="[371,474,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="mm" value="0.12">0.12 mm</quantity>
deep), approximately equidimensional with its antero-ventral edge strongly reflexed into the bursa, dorsal margin deeply cut away; thick-walled proximal ductus section of spermatheca with its bulbous end about half diameter of ductus length, thin-walled mid-section of utriculus shorter than the ductus, anterior sac of utriculus
<quantity id="713C9B0FFFD1FFC6FEB7A21EAB9AFE12" box="[329,415,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="0.1">0.1 mm</quantity>
diameter (female genital prep G1021, TMP).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD1FFC6FF3BA2E2A9B0FE39" box="[197,949,403,428]" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="B67B36EAFFD1FFC6FF3BA2E2A9B0FE39" blockId="10.[151,1437,151,645]" box="[197,949,403,428]" pageId="10" pageNumber="11">
<emphasis id="84B0EAF8FFD1FFC6FF3BA2E2AB3AFE39" bold="true" box="[197,319,403,428]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
The only
<taxonomicName id="71C44D69FFD1FFC6FE4FA2E5A84CFE39" box="[433,585,404,428]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFD1FFC6FE4FA2E5A84CFE39" box="[433,585,404,428]" italics="true" pageId="10" pageNumber="11">Agrionympha</emphasis>
</taxonomicName>
lacking a forewing claval mark.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD1FFC6FF3BA2C9AB2DFDCA" pageId="10" pageNumber="11" type="etymology">
<paragraph id="B67B36EAFFD1FFC6FF3BA2C9A9FDFE61" blockId="10.[151,1437,151,645]" pageId="10" pageNumber="11">
<emphasis id="84B0EAF8FFD1FFC6FF3BA2C9AB49FE45" bold="true" box="[197,332,440,464]" pageId="10" pageNumber="11">Etymology.</emphasis>
Named in honour of A.J.T. Janse, who revealed the presence of micropterigids in southern Africa when he collected
<taxonomicName id="71C44D69FFD1FFC6FE96A2ADABFCFE61" box="[360,505,476,500]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFD1FFC6FE96A2ADABFCFE61" box="[360,505,476,500]" italics="true" pageId="10" pageNumber="11">A. pseliacma</emphasis>
</taxonomicName>
at the same locality (Karkloof Falls) in 1917.
</paragraph>
<paragraph id="B67B36EAFFD1FFC6FF3BA28EAB2DFDCA" blockId="10.[151,1437,151,645]" pageId="10" pageNumber="11">
<emphasis id="84B0EAF8FFD1FFC6FF3BA28EAB4DFD8D" bold="true" box="[197,328,511,536]" pageId="10" pageNumber="11">Bionomics.</emphasis>
This species was discovered amongst ferns and seedlings on a steep slope under a thin, low, exotic canopy, while one of us (GWG) was searching for additional specimens of
<taxonomicName id="71C44D69FFD1FFC6FC1AA155AEBCFDA9" authority="Janse" authorityName="Janse" box="[996,1209,548,572]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFD1FFC6FC1AA155AE71FDA9" box="[996,1140,548,572]" italics="true" pageId="10" pageNumber="11">A. pseliacma</emphasis>
Janse
</taxonomicName>
in the vicinity of its
<typeStatus id="697F8848FFD1FFC6FF69A139AAC2FDF5" box="[151,199,584,608]" pageId="10" pageNumber="11">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD1FFC6FF3BA11AAF91FD11" box="[197,1428,619,645]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="B67B36EAFFD1FFC6FF3BA11AAF91FD11" blockId="10.[151,1437,151,645]" box="[197,1428,619,645]" pageId="10" pageNumber="11">
<emphasis id="84B0EAF8FFD1FFC6FF3BA11AAB5BFD11" bold="true" box="[197,350,619,644]" pageId="10" pageNumber="11">Distribution.</emphasis>
<collectingCountry id="CED3767AFFD1FFC6FE9BA11DABFDFD11" box="[357,504,620,644]" name="South Africa" pageId="10" pageNumber="11">South Africa</collectingCountry>
, Eastern Cape. Known only from the
<typeStatus id="697F8848FFD1FFC6FC60A11CA9CBFD10" box="[926,974,621,645]" pageId="10" pageNumber="11">type</typeStatus>
locality at Karkloof Falls, near Howick.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="12E9C84F718F1DD11DA7E81065717897">Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)</mods:title>
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<paragraph id="B67B36EAFFD4FFC3FF69A6D3A834FA29" blockId="15.[151,561,1442,1534]" box="[151,561,1442,1468]" pageId="15" pageNumber="16">
<heading id="ED338186FFD4FFC3FF69A6D3A834FA29" bold="true" box="[151,561,1442,1468]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="84B0EAF8FFD4FFC3FF69A6D3A834FA29" bold="true" box="[151,561,1442,1468]" pageId="15" pageNumber="16">
<taxonomicName id="71C44D69FFD4FFC3FF69A6D3A828FA29" ID-CoL="B7SJ" authority="Whalley, 1978" authorityName="Whalley" authorityYear="1978" box="[151,557,1442,1468]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFD4FFC3FF69A6D3AB76FA29" bold="true" box="[151,371,1442,1468]" italics="true" pageId="15" pageNumber="16">Agrionympha vari</emphasis>
<bibRefCitation id="D2554B1BFFD4FFC3FE84A6D3A828FA29" author="Whalley" box="[378,557,1442,1468]" pageId="15" pageNumber="21" refString="Whalley, P. E. S. (1978) New taxa of fossil and recent Micropterigidae with a discussion of their evolution and a comment on the evolution of Lepidoptera (Insecta). Annals of the Transvaal Museum, 31, 71 - 86." type="journal article" year="1978">Whalley, 1978</bibRefCitation>
</taxonomicName>
.
</emphasis>
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</paragraph>
<paragraph id="B67B36EAFFD4FFC3FF69A6B4AB2DFA6B" blockId="15.[151,561,1442,1534]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF69A6B4AB67FA48" box="[151,354,1477,1501]" italics="true" pageId="15" pageNumber="16">Ann. Transv. Mus.</emphasis>
31: 76. (
<figureCitation id="2EFF2A6FFFD4FFC3FF61A697AAE8FA6B" box="[159,237,1510,1534]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="15" pageNumber="16">
Figs.
<date id="C27A102AFFD4FFC3FF20A697AAE8FA6B" box="[222,237,1510,1534]" pageId="15" pageNumber="16">2</date>
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I, 11)
</paragraph>
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<paragraph id="B67B36EAFFD4FFC3FF69A55AA83CF9F2" blockId="15.[151,1437,1579,2036]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF69A55AAB86F9D1" bold="true" box="[151,387,1579,1604]" pageId="15" pageNumber="16">Material examined.</emphasis>
<typeStatus id="697F8848FFD4FFC3FE73A55DABF0F9D1" box="[397,501,1580,1604]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
Ƥ, labelled: Mariepskop,
<date id="C27A102AFFD4FFC3FCDBA55DA9DAF9D1" box="[805,991,1580,1604]" pageId="15" pageNumber="16" value="1956-01-24" valueMax="1956-01-25" valueMin="1956-01-24">2425 Jan 1956</date>
,
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. HT 4855,
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<emphasis id="84B0EAF8FFD4FFC3FB32A55DAF99F9D1" box="[1228,1436,1580,1604]" italics="true" pageId="15" pageNumber="16">Agrionympha vari</emphasis>
Whalley
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(
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). A perfect specimen.
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<emphasis id="84B0EAF8FFD4FFC3FF3BA502AB52F919" bold="true" box="[197,343,1651,1676]" pageId="15" pageNumber="16">Description.</emphasis>
<emphasis id="84B0EAF8FFD4FFC3FE9AA505ABA5F919" box="[356,416,1652,1676]" italics="true" pageId="15" pageNumber="16">Head</emphasis>
: Antennae dark greyish-brown throughout; 24 flagellomeres in female, lacking basal scales. Piliform scales on scape, pedicel and head ochreous-brown. Cranial piliform scales ochreous with brown tinge.
</paragraph>
<paragraph id="B67B36EAFFD4FFC3FF39A591ABE6F839" blockId="15.[151,1437,1579,2036]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF39A591AB1DF96D" box="[199,280,1760,1784]" italics="true" pageId="15" pageNumber="16">Thorax</emphasis>
: Tegulae silvery-white scaled. Dorsum of thorax dark greyish-brown with purplish sheen. Pro- and mesofemora pale ochreous, hindfemur shiny ochreous-grey; all tibiae and tarsi similar to but darker than femora; no obvious banding. FW length
<quantity id="713C9B0FFFD4FFC3FE00A456A852F8D5" box="[510,599,1831,1856]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="mm" value="3.5">3.5 mm</quantity>
. in female. FW maculation (
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Fig
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I) with bands not distinctly dark-edged; a narrow claval mark, kinked toward dorsum; an uninterrupted median band; a narrow post-median band represented by a costal bar extending to the middle of the wing and an almost circular spot on the dorsum; a narrow apical band which is slightly subterminal.
</paragraph>
<paragraph id="B67B36EAFFD4FFC3FF39A4C9A84DF845" blockId="15.[151,1437,1579,2036]" box="[199,584,1976,2000]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF39A4C9AB99F845" box="[199,412,1976,2000]" italics="true" pageId="15" pageNumber="16">Male postabdomen</emphasis>
: sex unknown.
</paragraph>
<paragraph id="B67B36EAFFD4FFC3FF39A4ADAE38F861" blockId="15.[151,1437,1579,2036]" box="[199,1085,2012,2036]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF39A4ADABB2F861" box="[199,439,2012,2036]" italics="true" pageId="15" pageNumber="16">Female postabdomen</emphasis>
: Not examined (see remarks under
<taxonomicName id="71C44D69FFD4FFC3FCB9A4ADA9E7F861" box="[839,994,2012,2036]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pseudovari">
<emphasis id="84B0EAF8FFD4FFC3FCB9A4ADA9E7F861" box="[839,994,2012,2036]" italics="true" pageId="15" pageNumber="16">A. pseudovari</emphasis>
</taxonomicName>
above).
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<subSubSection id="FEDE6561FFCBFFDCFF3BA3E6A922FEF1" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="B67B36EAFFCBFFDCFF3BA3E6A922FEF1" blockId="16.[151,1436,151,429]" pageId="16" pageNumber="17">
<emphasis id="84B0EAF8FFCBFFDCFF3BA3E6AB3AFF25" bold="true" box="[197,319,151,176]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Like
<taxonomicName id="71C44D69FFCBFFDCFE7CA3E9A81BFF25" box="[386,542,152,176]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pseudovari">
<emphasis id="84B0EAF8FFCBFFDCFE7CA3E9A81BFF25" box="[386,542,152,176]" italics="true" pageId="16" pageNumber="17">A. pseudovari</emphasis>
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distinctive by forewing postmedian band being interrupted in central part; distinguishable from
<taxonomicName id="71C44D69FFCBFFDCFE9AA3CDABE5FF41" box="[356,480,188,212]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pseudovari">
<emphasis id="84B0EAF8FFCBFFDCFE9AA3CDABE5FF41" box="[356,480,188,212]" italics="true" pageId="16" pageNumber="17">pseudovari</emphasis>
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by un-lined bands, and more parallel-sided costal remnant of postmedian band.
<emphasis id="84B0EAF8FFCBFFDCFF3BA391AB3CFF6D" bold="true" box="[197,313,224,248]" pageId="16" pageNumber="17">Remarks.</emphasis>
D.M. Kroon and GWG searched for
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<emphasis id="84B0EAF8FFCBFFDCFD22A391A92DFF6D" box="[732,808,224,248]" italics="true" pageId="16" pageNumber="17">A. vari</emphasis>
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at the
<typeStatus id="697F8848FFCBFFDCFC8AA391A9A1FF6D" box="[884,932,224,248]" pageId="16" pageNumber="17">type</typeStatus>
locality, 30 years after its discovery, without success, but instead finding yet another species of
<emphasis id="84B0EAF8FFCBFFDCFD2EA275A9E9FE89" box="[720,1004,260,284]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="71C44D69FFCBFFDCFD2EA275A96DFE89" box="[720,872,260,284]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Agrionympha</taxonomicName>
(
<taxonomicName id="71C44D69FFCBFFDCFC86A275A9E1FE89" box="[888,996,260,284]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="kroonella">kroonella</taxonomicName>
)
</emphasis>
. With this experience at two localities (also at Karkloof Falls) we can only presume that the diversity of micropterigids in
<collectingCountry id="CED3767AFFCBFFDCFBC7A256AECDFEAA" box="[1081,1224,295,319]" name="South Africa" pageId="16" pageNumber="17">South Africa</collectingCountry>
could be considerably higher than the current systematic study has revealed.
</paragraph>
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<subSubSection id="FEDE6561FFCBFFDCFF3BA21EABF2FE39" pageId="16" pageNumber="17" type="distribution">
<paragraph id="B67B36EAFFCBFFDCFF3BA21EABF2FE39" blockId="16.[151,1436,151,429]" pageId="16" pageNumber="17">
<emphasis id="84B0EAF8FFCBFFDCFF3BA21EAB5BFE1D" bold="true" box="[197,350,367,392]" pageId="16" pageNumber="17">Distribution.</emphasis>
<collectingCountry id="CED3767AFFCBFFDCFE90A21EA80FFE12" box="[366,522,367,391]" name="South Africa" pageId="16" pageNumber="17">South Africa</collectingCountry>
, Mpumalanga Province, formerly Transvaal, Mariepskop, 24º34ʹS, 32º52ʹE. Known only from
<typeStatus id="697F8848FFCBFFDCFE98A2E4AB93FE38" box="[358,406,405,429]" pageId="16" pageNumber="17">type</typeStatus>
locality.
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<mods:identifier id="EA3788EC1A91B89F987D7A438EBCD668" type="Zenodo-Dep">203553</mods:identifier>
</mods:mods>
<treatment id="3E6D87FCFFD6FFC3FF69A6E8AF54FAC1" ID-DOI="http://doi.org/10.5281/zenodo.6189667" ID-GBIF-Taxon="119388871" ID-Zenodo-Dep="6189667" LSID="urn:lsid:plazi:treatment:3E6D87FCFFD6FFC3FF69A6E8AF54FAC1" httpUri="http://treatment.plazi.org/id/3E6D87FCFFD6FFC3FF69A6E8AF54FAC1" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="FEDE6561FFD6FFC1FF69A6E8AB34FA41" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="B67B36EAFFD6FFC1FF69A6E8A87AFA26" blockId="13.[151,639,1433,1492]" box="[151,639,1433,1459]" pageId="13" pageNumber="14">
<heading id="ED338186FFD6FFC1FF69A6E8A87AFA26" bold="true" box="[151,639,1433,1459]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="84B0EAF8FFD6FFC1FF69A6E8A87AFA26" bold="true" box="[151,639,1433,1459]" pageId="13" pageNumber="14">
<taxonomicName id="71C44D69FFD6FFC1FF69A6E8A81FFA26" authority="Gibbs" authorityName="Gibbs" box="[151,538,1433,1459]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pseudovari" status="sp. nov.">
<emphasis id="84B0EAF8FFD6FFC1FF69A6E8ABCDFA26" bold="true" box="[151,456,1433,1459]" italics="true" pageId="13" pageNumber="14">Agrionympha pseudovari</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="9F835783FFD6FFC1FDDCA6E8A87AFA26" box="[546,639,1433,1459]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B67B36EAFFD6FFC1FF69A6CDAB34FA41" blockId="13.[151,639,1433,1492]" box="[151,305,1468,1492]" pageId="13" pageNumber="14">
(
<figureCitation id="2EFF2A6FFFD6FFC1FF61A6CDAAE8FA41" box="[159,237,1468,1492]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="13" pageNumber="14">Figs. 2</figureCitation>
C, 11)
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD6FFC1FF69A575AE26F9D5" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="B67B36EAFFD6FFC1FF69A575AE26F9D5" blockId="13.[151,1436,1540,2032]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF69A575AB7AF988" bold="true" box="[151,383,1540,1565]" pageId="13" pageNumber="14">Material examined.</emphasis>
<emphasis id="84B0EAF8FFD6FFC1FE78A575ABE8F989" box="[390,493,1540,1564]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="697F8848FFD6FFC1FE78A575ABE8F989" box="[390,493,1540,1564]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
Ƥ, labelled: Worcester, Fairy Glen,
<date id="C27A102AFFD6FFC1FC80A575AE5EF989" box="[894,1115,1540,1564]" pageId="13" pageNumber="14" value="1966-10-15" valueMax="1966-10-19" valueMin="1966-10-15">15 and 19 Oct 1966</date>
,
<taxonomicName id="71C44D69FFD6FFC1FB9BA575AE92F989" box="[1125,1175,1540,1564]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="20" phylum="Arthropoda" rank="species" species="vari">Vári</taxonomicName>
and Potgieter (=
<taxonomicName id="71C44D69FFD6FFC1FAAEA575AAF2F9D5" authority="Whalley" authorityName="Whalley" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFD6FFC1FAAEA575AF99F989" box="[1360,1436,1540,1564]" italics="true" pageId="13" pageNumber="14">A. vari</emphasis>
Whalley
</taxonomicName>
,
<typeStatus id="697F8848FFD6FFC1FEFCA558AB67F9D4" box="[258,354,1577,1601]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
4856
<collectionCode id="D0D5AE2FFFD6FFC1FE56A559ABE3F9D5" box="[424,486,1576,1600]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="13" pageNumber="14">TMP</collectionCode>
). RH FW missing. Abdomen slide G10440 (
<collectionCode id="D0D5AE2FFFD6FFC1FC26A559AE12F9D5" box="[984,1047,1576,1600]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="13" pageNumber="14">TMP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD6FFC1FF3BA53DA96EF83D" pageId="13" pageNumber="14" type="description">
<paragraph id="B67B36EAFFD6FFC1FF3BA53DAF1FF91D" blockId="13.[151,1436,1540,2032]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF3BA53DAB52F9F0" bold="true" box="[197,343,1612,1637]" pageId="13" pageNumber="14">Description.</emphasis>
<emphasis id="84B0EAF8FFD6FFC1FEA1A53DAB99F9F1" box="[351,412,1612,1636]" italics="true" pageId="13" pageNumber="14">Head</emphasis>
: Antennae dark grey-brown throughout; 24 flagellomeres in female, lacking basal scales. Piliform scales on scape and pedicel ochreous-brown. Head piliform scales ochreous with brown tinge.
</paragraph>
<paragraph id="B67B36EAFFD6FFC1FF39A5E5A8D7F8A9" blockId="13.[151,1436,1540,2032]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF39A5E5AB24F939" box="[199,289,1684,1708]" italics="true" pageId="13" pageNumber="14">Thorax:</emphasis>
Tegulae silvery-white scaled. Dorsum of thorax ochreous-grey. Legs with femora pale ochreous. FW length
<quantity id="713C9B0FFFD6FFC1FF18A5C9AB3BF945" box="[230,318,1720,1744]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="14" unit="mm" value="3.5">3.5 mm</quantity>
in female. FWs (
<figureCitation id="2EFF2A6FFFD6FFC1FDF7A5C9A84BF945" box="[521,590,1720,1744]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
C) with bands distinctly edged with black scales; a lozenge-shaped claval mark, slightly angled toward the dorsum; a broad uninterrupted median band, almost straight; postmedian band represented by two separate almost circular spots, the larger on costal margin, the smaller on dorsum; a slightly kinked apical band which is distinctly subterminal.
</paragraph>
<paragraph id="B67B36EAFFD6FFC1FF39A439A849F8F5" blockId="13.[151,1436,1540,2032]" box="[199,588,1864,1888]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF39A439AB99F8F5" box="[199,412,1864,1888]" italics="true" pageId="13" pageNumber="14">Male postabdomen</emphasis>
: Sex unknown.
</paragraph>
<paragraph id="B67B36EAFFD6FFC1FF39A41DA96EF83D" blockId="13.[151,1436,1540,2032]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF39A41DABBDF811" box="[199,440,1900,1924]" italics="true" pageId="13" pageNumber="14">Female postabdomen</emphasis>
: The abdomen slide preparation has not been re-examined here, since we do not expect to find taxonomically decisive characters in the female genitalia.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD6FFC1FF3BA4C5AED5F865" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="B67B36EAFFD6FFC1FF3BA4C5AED5F865" blockId="13.[151,1436,1540,2032]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF3BA4C5AB3AF858" bold="true" box="[197,319,1972,1997]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
Like
<taxonomicName id="71C44D69FFD6FFC1FE7FA4C5ABC8F859" box="[385,461,1972,1996]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFD6FFC1FE7FA4C5ABC8F859" box="[385,461,1972,1996]" italics="true" pageId="13" pageNumber="14">A. vari</emphasis>
</taxonomicName>
distinctive by forewing postmedian band being interrupted in central part; distinguishable from
<taxonomicName id="71C44D69FFD6FFC1FEF4A4A9AB4AF865" box="[266,335,2008,2032]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFD6FFC1FEF4A4A9AB4AF865" box="[266,335,2008,2032]" italics="true" pageId="13" pageNumber="14">A.vari</emphasis>
</taxonomicName>
by dark lining of bands, and broader, ovoid costal remnant of postmedian band.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD5FFC2FF3BA3E9AF01FF3A" box="[197,1284,151,176]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="B67B36EAFFD5FFC2FF3BA3E9AF01FF3A" blockId="14.[151,1436,151,463]" box="[197,1284,151,176]" pageId="14" pageNumber="15">
<emphasis id="84B0EAF8FFD5FFC2FF3BA3E9AB49FF25" bold="true" box="[197,332,152,176]" pageId="14" pageNumber="15">Etymology.</emphasis>
The name conveys the superficial similarity of this species to
<taxonomicName id="71C44D69FFD5FFC2FBF8A3E9AF05FF3A" authority="Whalley, 1978" authorityName="Whalley" authorityYear="1978" box="[1030,1280,151,176]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFD5FFC2FBF8A3E9AE57FF25" box="[1030,1106,152,176]" italics="true" pageId="14" pageNumber="15">A. vari</emphasis>
<bibRefCitation id="D2554B1BFFD5FFC2FBA4A3E6AF05FF3A" author="Whalley" box="[1114,1280,151,175]" pageId="14" pageNumber="21" refString="Whalley, P. E. S. (1978) New taxa of fossil and recent Micropterigidae with a discussion of their evolution and a comment on the evolution of Lepidoptera (Insecta). Annals of the Transvaal Museum, 31, 71 - 86." type="journal article" year="1978">Whalley, 1978</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD5FFC2FF3BA3CDA82EFE1D" pageId="14" pageNumber="15" type="discussion">
<paragraph id="B67B36EAFFD5FFC2FF3BA3CDA82EFE1D" blockId="14.[151,1436,151,463]" pageId="14" pageNumber="15">
<emphasis id="84B0EAF8FFD5FFC2FF3BA3CDAB3CFF41" bold="true" box="[197,313,188,212]" pageId="14" pageNumber="15">Remarks.</emphasis>
Although believed to be conspecific with
<taxonomicName id="71C44D69FFD5FFC2FCE7A3CDA963FF41" box="[793,870,188,212]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFD5FFC2FCE7A3CDA963FF41" box="[793,870,188,212]" italics="true" pageId="14" pageNumber="15">A. vari</emphasis>
</taxonomicName>
by
<bibRefCitation id="D2554B1BFFD5FFC2FC6FA3CDAE41FF41" author="Whalley" box="[913,1092,188,212]" pageId="14" pageNumber="21" refString="Whalley, P. E. S. (1978) New taxa of fossil and recent Micropterigidae with a discussion of their evolution and a comment on the evolution of Lepidoptera (Insecta). Annals of the Transvaal Museum, 31, 71 - 86." type="journal article" year="1978">Whalley (1978)</bibRefCitation>
, we consider the different FW pattern to be convincing evidence that this specimen represents a distinct species. The taxons ocurrence in the semi-arid Klein
<taxonomicName id="71C44D69FFD5FFC2FEB3A274AB93FE89" box="[333,406,261,284]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="20" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">Karoo</taxonomicName>
, in October, is a far cry from the montane rainforest of the northern Drakensberg Ranges in January, where
<taxonomicName id="71C44D69FFD5FFC2FEBAA259AB9DFEAA" box="[324,408,295,320]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="20" phylum="Arthropoda" rank="species" species="vari">A. Vári</taxonomicName>
collected the designated
<typeStatus id="697F8848FFD5FFC2FD4EA256A917FEAA" box="[688,786,295,319]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
of Whalleys species, over
<quantity id="713C9B0FFFD5FFC2FBBBA256AEA2FEAA" box="[1093,1191,295,319]" metricMagnitude="6" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="15" unit="km" value="1400.0">1400 km</quantity>
away. The
<typeStatus id="697F8848FFD5FFC2FADBA259AF50FED5" box="[1317,1365,296,320]" pageId="14" pageNumber="15">type</typeStatus>
locality of
<taxonomicName id="71C44D69FFD5FFC2FF26A23DAB76FEF1" box="[216,371,332,356]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pseudovari">
<emphasis id="84B0EAF8FFD5FFC2FF26A23DAB76FEF1" box="[216,371,332,356]" italics="true" pageId="14" pageNumber="15">A. pseudovari</emphasis>
</taxonomicName>
, in a deep sandstone kloof, where damp seepage has permitted growth of liverworts, is reminiscent of the habitat of
<taxonomicName id="71C44D69FFD5FFC2FE7BA201ABE1FE1D" box="[389,484,368,392]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="84B0EAF8FFD5FFC2FE7BA201ABE1FE1D" box="[389,484,368,392]" italics="true" pageId="14" pageNumber="15">A. karoo</emphasis>
</taxonomicName>
<emphasis id="84B0EAF8FFD5FFC2FE12A201A82EFE1D" bold="true" box="[492,555,368,392]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="9F835783FFD5FFC2FE12A201A82EFE1D" box="[492,555,368,392]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD5FFC2FF3BA2E2A853F96D" pageId="14" pageNumber="15" type="distribution">
<paragraph id="B67B36EAFFD5FFC2FF3BA2E2AAF4FE5A" blockId="14.[151,1436,151,463]" pageId="14" pageNumber="15">
<emphasis id="84B0EAF8FFD5FFC2FF3BA2E2AB5BFE39" bold="true" box="[197,350,403,428]" pageId="14" pageNumber="15">Distribution.</emphasis>
<collectingCountry id="CED3767AFFD5FFC2FE97A2E5ABFAFE39" box="[361,511,404,428]" name="South Africa" pageId="14" pageNumber="15">South Africa</collectingCountry>
, Western Cape, near Worcester in the Klein
<taxonomicName id="71C44D69FFD5FFC2FBF2A2E4AE53FE39" box="[1036,1110,405,428]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="20" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">Karoo</taxonomicName>
. Known only from the
<typeStatus id="697F8848FFD5FFC2FA92A2E4AF99FE38" box="[1388,1436,405,429]" pageId="14" pageNumber="15">type</typeStatus>
locality.
</paragraph>
<caption id="E2BB6662FFD5FFC2FF69A6C0AACEF9E5" httpUri="https://zenodo.org/record/203560/files/figure.png" pageId="14" pageNumber="15" targetBox="[160,1416,529,1421]" targetPageId="14">
<paragraph id="B67B36EAFFD5FFC2FF69A6C0AACEF9E5" blockId="14.[151,1436,1457,1648]" pageId="14" pageNumber="15">
<emphasis id="84B0EAF8FFD5FFC2FF69A6C0AB11FA52" bold="true" box="[151,276,1457,1480]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
<taxonomicName id="71C44D69FFD5FFC2FEE2A6C3A866FA5D" authority="Whalley" authorityName="Whalley" box="[284,611,1458,1480]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFD5FFC2FEE2A6C3A801FA5D" box="[284,516,1458,1480]" italics="true" pageId="14" pageNumber="15">Agrionympha capensis</emphasis>
Whalley
</taxonomicName>
, genitalia: AE, male, Storms River: A, ventral; B, lateral; C, dorsal; D, phallus lateral; E, phallus dorsal. FG, female, Bloukrans Pass: F, extended terminal segments of abdomen; G, internal female genitalia.
<emphasis id="84B0EAF8FFD5FFC2FF69A684AAB5F99F" bold="true" box="[151,176,1525,1546]" pageId="14" pageNumber="15">cb</emphasis>
, corpus bursae;
<emphasis id="84B0EAF8FFD5FFC2FEA6A684AB87F99F" bold="true" box="[344,386,1525,1546]" pageId="14" pageNumber="15">dph</emphasis>
, distal portion of phallus;
<emphasis id="84B0EAF8FFD5FFC2FD6EA684A8B3F99F" bold="true" box="[656,694,1525,1546]" pageId="14" pageNumber="15">dsp</emphasis>
, ductus of spermatheca;
<emphasis id="84B0EAF8FFD5FFC2FC4AA684A9C8F99F" bold="true" box="[948,973,1525,1546]" pageId="14" pageNumber="15">go</emphasis>
, gonopore with radial folds;
<emphasis id="84B0EAF8FFD5FFC2FB08A684AF13F99F" bold="true" box="[1270,1302,1525,1546]" pageId="14" pageNumber="15">lag</emphasis>
, lagena;
<emphasis id="84B0EAF8FFD5FFC2FA8AA684AF93F99F" bold="true" box="[1396,1430,1525,1546]" pageId="14" pageNumber="15">lap</emphasis>
, lappet,
<emphasis id="84B0EAF8FFD5FFC2FF1AA567AB0BF9BE" bold="true" box="[228,270,1558,1579]" pageId="14" pageNumber="15">mpl</emphasis>
, median plate;
<emphasis id="84B0EAF8FFD5FFC2FE4DA567ABD9F9BE" bold="true" box="[435,476,1558,1579]" pageId="14" pageNumber="15">pap</emphasis>
, papilla;
<emphasis id="84B0EAF8FFD5FFC2FDBEA567A86FF9BE" bold="true" box="[576,618,1558,1579]" pageId="14" pageNumber="15">pph</emphasis>
, proximal portion of phallus;
<emphasis id="84B0EAF8FFD5FFC2FC56A567A9E4F9B9" bold="true" box="[936,993,1557,1580]" pageId="14" pageNumber="15">scl ix</emphasis>
, ninth sclerite;
<emphasis id="84B0EAF8FFD5FFC2FB78A567AEBCF9BE" bold="true" box="[1158,1209,1558,1579]" pageId="14" pageNumber="15">scl x</emphasis>
, tenth sclerite;
<emphasis id="84B0EAF8FFD5FFC2FAA2A564AF93F9B9" bold="true" box="[1372,1430,1557,1580]" pageId="14" pageNumber="15">S viii</emphasis>
, eighth sternite;
<emphasis id="84B0EAF8FFD5FFC2FEC8A549AB66F9D8" bold="true" box="[310,355,1592,1613]" pageId="14" pageNumber="15">spm</emphasis>
, spermatheca;
<emphasis id="84B0EAF8FFD5FFC2FE00A549A83DF9DB" bold="true" box="[510,568,1591,1614]" pageId="14" pageNumber="15">T viii</emphasis>
, eighth tergite;
<emphasis id="84B0EAF8FFD5FFC2FD25A549A8FAF9D8" bold="true" box="[731,767,1592,1613]" pageId="14" pageNumber="15">T x</emphasis>
, tenth tergum;
<emphasis id="84B0EAF8FFD5FFC2FC63A549A9BBF9D8" bold="true" box="[925,958,1592,1613]" pageId="14" pageNumber="15">utr</emphasis>
, distal bulb of utriculus;
<emphasis id="84B0EAF8FFD5FFC2FB3FA549AEDFF9D8" bold="true" box="[1217,1242,1592,1613]" pageId="14" pageNumber="15">va</emphasis>
, valva;
<emphasis id="84B0EAF8FFD5FFC2FAD3A549AF4DF9D8" bold="true" box="[1325,1352,1592,1613]" pageId="14" pageNumber="15">vb</emphasis>
, ventral bulb.
</paragraph>
</caption>
<paragraph id="B67B36EAFFD5FFC2FF69A5CEA853F96D" blockId="14.[151,598,1727,1784]" pageId="14" pageNumber="15">
<taxonomicName id="71C44D69FFD5FFC2FF69A5CEABEAF942" authority="Gibbs" authorityName="Gibbs" box="[151,495,1727,1752]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sagittella" status="sp. nov.">
<emphasis id="84B0EAF8FFD5FFC2FF69A5CEABA0F94D" bold="true" box="[151,421,1727,1752]" italics="true" pageId="14" pageNumber="15">Agrionympha sagittella</emphasis>
Gibbs
</taxonomicName>
<emphasis id="84B0EAF8FFD5FFC2FE0BA5CEA84AF942" bold="true" box="[501,591,1727,1751]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="9F835783FFD5FFC2FE0BA5CEA84AF942" box="[501,591,1727,1751]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="2EFF2A6FFFD5FFC2FF61A591AAECF96D" box="[159,233,1760,1784]" captionStart="FIGURE 1" captionStartId="7.[151,250,1973,1996]" captionTargetBox="[206,1390,1043,1943]" captionTargetId="figure@7.[196,1401,1035,1968]" captionTargetPageId="7" captionText="FIGURE 1. Adult moths from life: A, Agrionympha capensis Whalley, Storms River; B, A. kroonella, sp. nov. Mariepskop; C, A. fuscoapicella, sp. nov. Hogsback; D, A. sagittella, sp. nov. Hogsback. Scale bar = 1 mm." httpUri="https://zenodo.org/record/203554/files/figure.png" pageId="14" pageNumber="15">Figs. 1</figureCitation>
D, 2F, 3A, C, E, 4C, 10CG, 11)
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD5FFC3FF69A452A9CBFF41" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="B67B36EAFFD5FFC3FF69A452A9CBFF41" blockId="14.[151,1437,1827,2031]" lastBlockId="15.[151,1437,151,1364]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="84B0EAF8FFD5FFC2FF69A452AB87F8A9" bold="true" box="[151,386,1827,1852]" pageId="14" pageNumber="15">Material examined.</emphasis>
<typeStatus id="697F8848FFD5FFC2FE75A452ABF7F8AE" box="[395,498,1827,1851]" pageId="14" pageNumber="15" type="holotype">
<emphasis id="84B0EAF8FFD5FFC2FE75A452ABF7F8AE" box="[395,498,1827,1851]" italics="true" pageId="14" pageNumber="15">Holotype</emphasis>
</typeStatus>
3, labelled:
<collectingCountry id="CED3767AFFD5FFC2FD76A452A95DF8AE" box="[648,856,1827,1851]" name="South Africa" pageId="14" pageNumber="15">SOUTH AFRICA</collectingCountry>
, CP,
<geoCoordinate id="D3F0502DFFD5FFC2FC6DA452A9E9F8AE" box="[915,1004,1827,1851]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-32.583332">32º35´S</geoCoordinate>
<geoCoordinate id="D3F0502DFFD5FFC2FC0BA452AE56F8AE" box="[1013,1107,1827,1851]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="26.95">26º57´E</geoCoordinate>
, Kettlespout,
<quantity id="713C9B0FFFD5FFC2FB0FA452AF4FF8AE" box="[1265,1354,1827,1851]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="14" pageNumber="15" unit="m" value="1260.0">1260 m</quantity>
, Hogsback S.F.
<date id="C27A102AFFD5FFC2FEFAA436ABBCF8CA" box="[260,441,1863,1887]" pageId="14" pageNumber="15" value="1986-02-11" valueMax="1986-02-13" valueMin="1986-02-11">1113 Feb 1986</date>
, G.W. Gibbs (
<collectionCode id="D0D5AE2FFFD5FFC2FDADA436A897F8CA" box="[595,658,1863,1887]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="14" pageNumber="15">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFD5FFC2FD5BA439A912F8CA" box="[677,791,1864,1887]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="697F8848FFD5FFC2FD5BA439A912F8CA" box="[677,791,1864,1887]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
</emphasis>
: 83 5Ƥ,
<collectingCountry id="CED3767AFFD5FFC2FC80A436AE0BF8CA" box="[894,1038,1863,1887]" name="South Africa" pageId="14" pageNumber="15">South Africa</collectingCountry>
(EC), Hogsback, Kettlespout track,
<quantity id="713C9B0FFFD5FFC2FF69A41AAAF4F816" box="[151,241,1899,1923]" metricMagnitude="3" metricUnit="m" metricValue="1.26" pageId="14" pageNumber="15" unit="m" value="1260.0">1260 m</quantity>
,
<geoCoordinate id="D3F0502DFFD5FFC2FEFEA41AAB5FF816" box="[256,346,1899,1923]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="-32.416668">32º25´S</geoCoordinate>
<geoCoordinate id="D3F0502DFFD5FFC2FE9BA41AABC4F816" box="[357,449,1899,1923]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="26.95">26º57´E</geoCoordinate>
,
<date id="C27A102AFFD5FFC2FE2FA41AA895F816" box="[465,656,1899,1923]" pageId="14" pageNumber="15" value="1986-02-11" valueMax="1986-02-13" valueMin="1986-02-11">1113 Feb 1986</date>
, N. Duke &amp; G.W. Gibbs (male genital preps, N.P. Kristensen 1034 (
<collectionCode id="D0D5AE2FFFD5FFC2FF61A4FEAAF3F832" box="[159,246,1935,1959]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="14" pageNumber="15">ZMUC</collectionCode>
), G.W. Gibbs G1008 (
<collectionCode id="D0D5AE2FFFD5FFC2FE0BA4FEA831F832" box="[501,564,1935,1959]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="14" pageNumber="15">TMP</collectionCode>
)); 33,
<collectingCountry id="CED3767AFFD5FFC2FD76A4FEA91FF832" box="[648,794,1935,1959]" name="South Africa" pageId="14" pageNumber="15">South Africa</collectingCountry>
(EC), Hogsback, Auckland Nature Reserve track,
<quantity id="713C9B0FFFD5FFC2FA9AA4FEAAEAF85E" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="14" pageNumber="15" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900 1000 m</quantity>
,
<date id="C27A102AFFD5FFC2FF02A4C2AB99F85E" box="[252,412,1971,1995]" pageId="14" pageNumber="15" value="2004-02-05" valueMax="2004-02-07" valueMin="2004-02-05">57 Feb 2004</date>
, G.W. Gibbs; 23 2Ƥ,
<collectingCountry id="CED3767AFFD5FFC2FD66A4C2A92CF85E" box="[664,809,1971,1995]" name="South Africa" pageId="14" pageNumber="15">South Africa</collectingCountry>
(EC), Ngadu Forest, 10 k N Umtata,
<quantity id="713C9B0FFFD5FFC2FB26A4C2AF35F85E" box="[1240,1328,1971,1995]" metricMagnitude="3" metricUnit="m" metricValue="1.304" pageId="14" pageNumber="15" unit="m" value="1304.0">1304 m</quantity>
, 23 &amp;
<date id="C27A102AFFD5FFC2FA7EA4C2AB0CF87A" pageId="14" pageNumber="15" value="1986-11-30">30 Nov 1986</date>
, N. Duke (male genital prep G1004,
<collectionCode id="D0D5AE2FFFD5FFC2FD59A4A6A8E0F87A" box="[679,741,2007,2031]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="14" pageNumber="15">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFD5FFC2FD06A4A6A9A5F87A" box="[760,928,2007,2031]" italics="true" pageId="14" pageNumber="15">Other material</emphasis>
: Alcohol preserved material from Hogsback: Kettlespout track (
<quantity id="713C9B0FFFD4FFC3FE99A3E6ABC5FF3A" box="[359,448,151,175]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="16" unit="m" value="1200.0">1200 m</quantity>
), Hogsback Pass roadside (
<quantity id="713C9B0FFFD4FFC3FD0AA3E6A945FF3A" box="[756,832,151,175]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="15" pageNumber="16" unit="m" value="940.0">940 m</quantity>
), Auckland Nature Reserve track (ca.
<quantity id="713C9B0FFFD4FFC3FB11A3E6AF4DFF3A" box="[1263,1352,151,175]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="1000.0">1000 m</quantity>
),
<date id="C27A102AFFD4FFC3FAA3A3E9AAD7FF41" pageId="15" pageNumber="16" value="2004-02-07">7 Feb 2004</date>
, G.W. Gibbs (female genital prep. G1020, wing slide G1026,
<collectionCode id="D0D5AE2FFFD4FFC3FC7DA3CDA9C4FF41" box="[899,961,188,212]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="15" pageNumber="16">TMP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD4FFC3FF3BA3AEA9A1FBCD" pageId="15" pageNumber="16" type="description">
<paragraph id="B67B36EAFFD4FFC3FF3BA3AEA9BEFEAA" blockId="15.[151,1437,151,1364]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF3BA3AEAB52FF6D" bold="true" box="[197,343,223,248]" pageId="15" pageNumber="16">Description.</emphasis>
<emphasis id="84B0EAF8FFD4FFC3FE9CA391AB9BFF6D" box="[354,414,224,248]" italics="true" pageId="15" pageNumber="16">Head</emphasis>
: Antennae grey-brown to dark brown-black throughout; 45 flagellomeres in male with basal 3 scale-covered;
<quantity id="713C9B0FFFD4FFC3FE69A275ABFFFE89" box="[407,506,260,284]" metricMagnitude="-1" metricUnit="m" metricValue="8.254999999999999" metricValueMax="8.382" metricValueMin="8.128" pageId="15" pageNumber="16" unit="in" value="32.5" valueMax="33.0" valueMin="32.0">3233 in</quantity>
female with basal 7 scaled. Piliform scales on scape and pedicel grey-black. Head piliform scales ochreous, greyish in mid-line on dorsum, paler on frons.
</paragraph>
<paragraph id="B67B36EAFFD4FFC3FF39A23DA96BFDCA" blockId="15.[151,1437,151,1364]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF39A23DAB1DFEF1" box="[199,280,332,356]" italics="true" pageId="15" pageNumber="16">Thorax</emphasis>
: Tegulae creamy-white scaled; mesonotum with black scales. Legs with all coxae and femora creamywhite, distal joints of coxae greyish; fore- and midtibiae mainly black but with whitish band in middle; fore- and midtarsi black with whitish bands at proximal ends of tarsomeres 1, 2 and 3, hindtarsus entirely black. FW length
<quantity id="713C9B0FFFD4FFC3FF69A2C6AAF5FE5A" box="[151,240,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="15" pageNumber="16" unit="mm" value="3.4">3.4 mm</quantity>
in male (3.23.5),
<quantity id="713C9B0FFFD4FFC3FE2CA2C6A82EFE5A" box="[466,555,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="15" pageNumber="16" unit="mm" value="3.7">3.7 mm</quantity>
in female (3.63.8). HW venation with Sc doubling sometimes more pronounced than in any other micropterigid examined (
<figureCitation id="2EFF2A6FFFD4FFC3FD26A2ADA91FFE61" box="[728,794,476,500]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="15" pageNumber="16">Fig. 3</figureCitation>
E). FW maculation (
<figureCitation id="2EFF2A6FFFD4FFC3FC00A2ADAE4DFE61" box="[1022,1096,476,500]" captionStart="FIGURE 1" captionStartId="7.[151,250,1973,1996]" captionTargetBox="[206,1390,1043,1943]" captionTargetId="figure@7.[196,1401,1035,1968]" captionTargetPageId="7" captionText="FIGURE 1. Adult moths from life: A, Agrionympha capensis Whalley, Storms River; B, A. kroonella, sp. nov. Mariepskop; C, A. fuscoapicella, sp. nov. Hogsback; D, A. sagittella, sp. nov. Hogsback. Scale bar = 1 mm." httpUri="https://zenodo.org/record/203554/files/figure.png" pageId="15" pageNumber="16">Figs. 1</figureCitation>
D, 2F) with very conspicuous, broad, T-shaped basal band resulting from confluence of claval mark with median band; a postmedian band, kinked at dorsum and interrupted by lines of dark scales on major veins; a postmedian band, also kinked toward dorsum and broken by lines of dark scales along veins; apex dark-scaled.
</paragraph>
<paragraph id="B67B36EAFFD4FFC3FF39A11DA9B3FCEA" blockId="15.[151,1437,151,1364]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF39A11DAB99FD11" box="[199,412,620,644]" italics="true" pageId="15" pageNumber="16">Male postabdomen</emphasis>
(
<figureCitation id="2EFF2A6FFFD4FFC3FE52A11DABFEFD11" box="[428,507,620,644]" captionStart="FIGURE 10" captionStartId="18.[151,250,1913,1936]" captionTargetBox="[313,1332,581,1845]" captionTargetId="figure@18.[244,1340,573,1969]" captionTargetPageId="18" captionText="FIGURE 10. Agrionympha male genitalia. A B, A. pseliacma, Meyrick, Karkloof Falls: A, lateral view, from slide 3830, Transvaal Museum, Pretoria; B, detail of phallus, lateral. C G, A. sagittella, sp. nov. Hogsback: C, lateral; D, dorsal; E, ventral; F, phallus, lateral; G, phallus dorsal. H J, Agrionympha species 1 , Diepwalle Forest: H, lateral; I, phallus lateral; J, phallus dorsal." httpUri="https://zenodo.org/record/203563/files/figure.png" pageId="15" pageNumber="16">Fig. 10</figureCitation>
CG): Sternum VIII remnants small, irregularly oval, bearing about 14 small setae. Anteroventral margin of IX sclerotisation pointed and slightly thickened. Median plate narrow, rectangular, not pigmented anteriorly. Valvae long and slender (ratio length:width 5.7), gently curved upwards and tapering. Approximately 50 hair-sensilla present, uniquely located along the anterior border of tergum VIII rather than on dorsum IX. About two thirds of the sensilla are partially covered by a field of comb-edged micro-scales (
<figureCitation id="2EFF2A6FFFD4FFC3FAB8A18DAF8CFC81" box="[1350,1417,764,788]" captionStart="FIGURE 4" captionStartId="10.[151,250,1611,1634]" captionTargetBox="[198,1390,705,1611]" captionTargetId="figure@10.[197,1390,700,1611]" captionTargetPageId="10" captionText="FIGURE 4. Structural detail of Agrionympha species: A, A. capensis, antennal flagellomere with ascoid sensilla (SEM); B, A. capensis, segment V gland (SEM); C, A. sagittella, hair-sensilla and micro-scales on dorsum of male tergum VIII (SEM); D, A. kroonella, hair sensilla on dorsum of segment IX (SEM). asb, branch of ascoid sensilla; as, central disc of ascoid sensilla; ped, raised peduncle of segment V gland, S iv, fourth sternite; S v, fifth sternite; sc, long fluted piliform scale. Scale bars: A, B = 0.01 mm; C, D = 0.005 mm." httpUri="https://zenodo.org/record/203557/files/figure.png" pageId="15" pageNumber="16">Fig. 4</figureCitation>
C SEM). Lateroventral X plates considerably smaller than tergum X. Phallus ratio length:height 5.3, with only a hint of a neck constriction in dorsal view; gonopore dorsal, heart-shaped, approximately half the width of the phallus between the lappets; ventral bulb moderate (male genital prep. G1008).
</paragraph>
<paragraph id="B67B36EAFFD4FFC3FF39A0FDA9A1FBCD" blockId="15.[151,1437,151,1364]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF39A0FDABC1FC31" box="[199,452,908,932]" italics="true" pageId="15" pageNumber="16">Female postabdomen:</emphasis>
Segment V gland
<quantity id="713C9B0FFFD4FFC3FD5AA0FDA8FBFC31" box="[676,766,908,932]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="16" unit="mm" value="0.4">0.4 mm</quantity>
diam., with 1213 long piliform scales. Segment X very slightly elongated, lateral sclerites about 1.1x longer than high, anterior edge slightly emarginated in some specimens. Papilla cup-shaped, roughly equidimensional, parallel sided with only the posterior margin reflexed. Thickwalled proximal section of spermatheca with its bulbous distal end about 2/3 diameter of ductus length, thin-walled mid-section of utriculus with proximal bulb poorly defined, this section about equal to the ductus in length, anterior distal sac of utriculus
<quantity id="713C9B0FFFD4FFC3FE72A731ABF5FBCD" box="[396,496,1088,1112]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="16" unit="mm" value="0.11">0.11 mm</quantity>
diameter (female genital prep G1020).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD4FFC3FF3BA712AB8AFB35" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="B67B36EAFFD4FFC3FF3BA712AB8AFB35" blockId="15.[151,1437,151,1364]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF3BA712AB3AFBE9" bold="true" box="[197,319,1123,1148]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
The arrow-shaped or T-configuration of the confluent forewing median and claval bands is unique to this species.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD4FFC3FF3BA7DDAF0AFAA5" pageId="15" pageNumber="16" type="etymology">
<paragraph id="B67B36EAFFD4FFC3FF3BA7DDAEF2FB51" blockId="15.[151,1437,151,1364]" box="[197,1271,1196,1220]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF3BA7DDAB49FB51" bold="true" box="[197,332,1196,1220]" pageId="15" pageNumber="16">Etymology.</emphasis>
From
<emphasis id="84B0EAF8FFD4FFC3FE68A7DCABE7FB51" box="[406,482,1197,1220]" italics="true" pageId="15" pageNumber="16">sagitta</emphasis>
(arrow), referring to the above-mentioned distinctive maculation trait.
</paragraph>
<paragraph id="B67B36EAFFD4FFC3FF3BA7BEAF0AFAA5" blockId="15.[151,1437,151,1364]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF3BA7BEAB4DFB7D" bold="true" box="[197,328,1231,1256]" pageId="15" pageNumber="16">Bionomics.</emphasis>
This montane species may coexist with
<taxonomicName id="71C44D69FFD4FFC3FCF3A7A1A9C4FB7D" box="[781,961,1232,1256]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fuscoapicella">
<emphasis id="84B0EAF8FFD4FFC3FCF3A7A1A9C4FB7D" box="[781,961,1232,1256]" italics="true" pageId="15" pageNumber="16">A. fuscoapicella</emphasis>
</taxonomicName>
along the margins of tall forest but is usually in sites that are more open to the sky. It has occurred amongst ferns, grasses and shrubs on damp sunny banks where liverworts are visible and along sunny roadsides or tracks at altitudes between 940 and
<quantity id="713C9B0FFFD4FFC3FB4DA669AF0FFAA5" box="[1203,1290,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="16" unit="m" value="1300.0">1300 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD4FFC3FF3BA64AAF54FAC1" box="[197,1361,1339,1364]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="B67B36EAFFD4FFC3FF3BA64AAF54FAC1" blockId="15.[151,1437,151,1364]" box="[197,1361,1339,1364]" pageId="15" pageNumber="16">
<emphasis id="84B0EAF8FFD4FFC3FF3BA64AAB5BFAC1" bold="true" box="[197,350,1339,1364]" pageId="15" pageNumber="16">Distribution.</emphasis>
<collectingCountry id="CED3767AFFD4FFC3FE9BA64DABFDFAC1" box="[357,504,1340,1364]" name="South Africa" pageId="15" pageNumber="16">South Africa</collectingCountry>
, Eastern Cape, Hogsback and Ngadu Forests, which are about
<quantity id="713C9B0FFFD4FFC3FB48A64DAF0EFAC1" box="[1206,1291,1340,1364]" metricMagnitude="5" metricUnit="m" metricValue="3.4" pageId="15" pageNumber="16" unit="km" value="340.0">340 km</quantity>
apart.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,152 @@
<document id="F610C0877A1D4A411682057BB8777B37" ID-DOI="10.5281/zenodo.203553" ID-GBIF-Dataset="dd1ad33b-e90a-4c23-a6b7-08d77c43e95f" ID-ISSN="1175-5326" ID-Zenodo-Dep="203553" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460237623643" checkinUser="plazi" docAuthor="Gibbs, George W. &amp; Kristensen, Niels P." docDate="2011" docId="3E6D87FCFFD7FFC1FF69A6F2AF6EFC81" docLanguage="en" docName="zt02764p021.pdf" docOrigin="Zootaxa 2764" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Agrionympha pseliacma Meyrick 1921" docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="C254FF84FFDBFFCCFFFEA371AA05FF95" masterDocTitle="Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="13" updateTime="1698239897964" updateUser="plazi">
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<mods:title id="BC2D483F069708225F7B29206CF580B2">Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)</mods:title>
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<heading id="ED338186FFD7FFC0FF69A6F2A864FA09" bold="true" box="[151,609,1411,1436]" fontSize="10" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="71C44D69FFD7FFC0FF69A6F2A864FA09" ID-CoL="B7SH" authority="Meyrick, 1921" authorityName="Meyrick" authorityYear="1921" box="[151,609,1411,1436]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFD7FFC0FF69A6F2A864FA09" bold="true" box="[151,609,1411,1436]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF69A6F2ABAAFA09" bold="true" box="[151,431,1411,1436]" italics="true" pageId="12" pageNumber="13">Agrionympha pseliacma</emphasis>
Meyrick, 1921
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<emphasis id="84B0EAF8FFD7FFC0FF69A6D9AB5FFA55" box="[151,346,1448,1472]" italics="true" pageId="12" pageNumber="13">Ann. Transv. Mus</emphasis>
. 8: 144.
</paragraph>
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<subSubSection id="FEDE6561FFD7FFC0FF69A6BDAB97FA71" box="[151,402,1484,1508]" pageId="12" pageNumber="13" type="description">
<paragraph id="B67B36EAFFD7FFC0FF69A6BDAB97FA71" blockId="12.[151,609,1411,1508]" box="[151,402,1484,1508]" pageId="12" pageNumber="13">
(
<figureCitation id="2EFF2A6FFFD7FFC0FF61A6BDAAE9FA71" box="[159,236,1484,1508]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="12" pageNumber="13">Figs. 2</figureCitation>
H, 10A, B, 11)
</paragraph>
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<subSubSection id="FEDE6561FFD7FFC0FF69A562AB84F902" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="B67B36EAFFD7FFC0FF69A562AB84F902" blockId="12.[151,1437,1554,2012]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF69A562AB87F9B9" bold="true" box="[151,386,1555,1580]" pageId="12" pageNumber="13">Material examined.</emphasis>
<emphasis id="84B0EAF8FFD7FFC0FE75A565ABF7F9B9" box="[395,498,1556,1580]" italics="true" pageId="12" pageNumber="13">
<typeStatus id="697F8848FFD7FFC0FE75A565ABF7F9B9" box="[395,498,1556,1580]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
Ƥ,
<collectingCountry id="CED3767AFFD7FFC0FDE1A565A8B1F9B9" box="[543,692,1556,1580]" name="South Africa" pageId="12" pageNumber="13">South Africa</collectingCountry>
, Kwazulu-Natal, Karkloof, 29º24ʹS, 30º17ʹE,
<date id="C27A102AFFD7FFC0FB41A565AF49F9B9" box="[1215,1356,1556,1580]" pageId="12" pageNumber="13" value="1917-01-11">11 Jan 1917</date>
, A.J.T. Janse (HT 77,
<collectionCode id="D0D5AE2FFFD7FFC0FEC0A546AB79F9DA" box="[318,380,1591,1615]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="12" pageNumber="13">TMP</collectionCode>
). Abdomen slide 7874, wings, antenna, fore leg, slide 1582 (
<collectionCode id="D0D5AE2FFFD7FFC0FBD1A546AE6BF9DA" box="[1071,1134,1591,1615]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="12" pageNumber="13">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFD7FFC0FB7DA548AEEFF9C5" box="[1155,1258,1593,1616]" italics="true" pageId="12" pageNumber="13">
<typeStatus id="697F8848FFD7FFC0FB7DA548AEEFF9C5" box="[1155,1258,1593,1616]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
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13, from same location,
<date id="C27A102AFFD7FFC0FEFEA52CAB75F9E1" box="[256,368,1628,1652]" pageId="12" pageNumber="13" value="1930-12">Dec 1930</date>
, A.J.T. Janse (
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). Male genital slide 3830, wings slide 3275 (
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); head slide M300, N.P. Kristensen (
<collectionCode id="D0D5AE2FFFD7FFC0FEE3A50EAB71F902" box="[285,372,1663,1687]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="12" pageNumber="13">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD7FFC1FF3BA5D2A860FE5A" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="B67B36EAFFD7FFC0FF3BA5D2ABC9F891" blockId="12.[151,1437,1554,2012]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF3BA5D2AB52F929" bold="true" box="[197,343,1699,1724]" pageId="12" pageNumber="13">Description.</emphasis>
<emphasis id="84B0EAF8FFD7FFC0FE9EA5D5AB99F929" box="[352,412,1700,1724]" italics="true" pageId="12" pageNumber="13">Head</emphasis>
: Antennae dark greyish-brown with faint purplish sheen, becoming slightly paler toward tip. Flagellar segments in male unknown, 30 segments in female with basal 8 scaled; piliform scales on scape, pedicel and head pale ochreous.
</paragraph>
<paragraph id="B67B36EAFFD7FFC0FF39A461AF11F801" blockId="12.[151,1437,1554,2012]" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF39A461AB1DF8BD" box="[199,280,1808,1832]" italics="true" pageId="12" pageNumber="13">Thorax</emphasis>
: dark bronzy-fuscus, tegulae lost. Legs damaged and largely descaled. FW length 3.0 mm in male,
<quantity id="713C9B0FFFD7FFC0FA87A47EAAC6F8D9" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="13" unit="mm" value="3.4">3.4 mm</quantity>
in female. FWs (
<figureCitation id="2EFF2A6FFFD7FFC0FE78A445ABCDF8D9" box="[390,456,1844,1868]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
H) coppery-golden-bronze with purplish reflections, all bands silvery-white edged with black scales; a narrow lozenge-shaped claval mark, angled toward the dorsum; a straight median band; a short postmedian band from costa reaching to middle of wing; a small apical band which is slightly subterminal.
</paragraph>
<paragraph id="B67B36EAFFD7FFC1FF39A4D1ABB6FE39" blockId="12.[151,1437,1554,2012]" lastBlockId="13.[151,1437,151,789]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="84B0EAF8FFD7FFC0FF39A4D1AB9BF82D" box="[199,414,1952,1976]" italics="true" pageId="12" pageNumber="13">Male postabdomen</emphasis>
(
<figureCitation id="2EFF2A6FFFD7FFC0FE51A4D1A804F82D" box="[431,513,1952,1976]" captionStart="FIGURE 10" captionStartId="18.[151,250,1913,1936]" captionTargetBox="[313,1332,581,1845]" captionTargetId="figure@18.[244,1340,573,1969]" captionTargetPageId="18" captionText="FIGURE 10. Agrionympha male genitalia. A B, A. pseliacma, Meyrick, Karkloof Falls: A, lateral view, from slide 3830, Transvaal Museum, Pretoria; B, detail of phallus, lateral. C G, A. sagittella, sp. nov. Hogsback: C, lateral; D, dorsal; E, ventral; F, phallus, lateral; G, phallus dorsal. H J, Agrionympha species 1 , Diepwalle Forest: H, lateral; I, phallus lateral; J, phallus dorsal." httpUri="https://zenodo.org/record/203563/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
A, description prepared from unique lateral slide mount 3830 Janse, TMP): Sternum VIII remnants roughly triangular with about 7 microsetae. Anterior margin of IX distinctly darkened and thickened, the thickened band becoming submarginal ventrally. Median plate roughly rectangular, 3.5 times longer than wide, more strongly sclerotised anteriorly then posteriorly. Valvae short (ratio length:width 3.6); their upward curvature markedly angled just beyond mid-length and tapering from there to a rounded apex; the postero-ventral face distinctly but irregularly flattened (note: this could possibly be an artifact of slide preparation). Form of tegumen lobes not clear in the preparation but appearing to conform with generic description. A cluster of about 33 hair-sensilla on the membraneous dorsum of IX. Phallus ratio length:height 4.2; ventral bulb moderate; status of neck region and shape of gonopore cannot be determined from the slide mount but there appears to be a flap-like overhang in the mid-line.
</paragraph>
<paragraph id="B67B36EAFFD6FFC1FF39A2C9A860FE5A" blockId="13.[151,1437,151,789]" box="[199,613,439,464]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF39A2C9ABB2FE45" box="[199,439,440,464]" italics="true" pageId="13" pageNumber="14">Female postabdomen</emphasis>
: not examined.
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<subSubSection id="FEDE6561FFD6FFC1FF3BA2AAAB85FD7A" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="B67B36EAFFD6FFC1FF3BA2AAA968FDA9" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF3BA2AAAB3AFE61" bold="true" box="[197,319,475,500]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
Shares with
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<emphasis id="84B0EAF8FFD6FFC1FE27A2ADA895FE61" box="[473,656,476,500]" italics="true" pageId="13" pageNumber="14">A. fuscoapicella</emphasis>
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and
<taxonomicName id="71C44D69FFD6FFC1FD32A2ADA942FE61" box="[716,839,476,500]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="jansella">
<emphasis id="84B0EAF8FFD6FFC1FD32A2ADA942FE61" box="[716,839,476,500]" italics="true" pageId="13" pageNumber="14">A. jansella</emphasis>
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the short forewing postmedian band represented as only on the costal end, but retains the complete set of bands, i.e. distinguishable from the former by the presence of an apical patch, and the latter by the presence of the claval mark.
</paragraph>
<paragraph id="B67B36EAFFD6FFC1FF3BA136AB85FD7A" blockId="13.[151,1437,151,789]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF3BA136AB4DFDF5" bold="true" box="[197,328,583,608]" pageId="13" pageNumber="14">Bionomics.</emphasis>
Found under the shade of very high trees with little patches of sunshine in the entrance to a kloof.
<bibRefCitation id="D2554B1BFFD6FFC1FF69A11CAB2DFD11" author="Janse" box="[151,296,620,644]" pageId="13" pageNumber="21" refString="Janse, A. J. T. (1942) The Moths of South Africa. Vol IV (1): 75 - 78. Pl LVII, figs 4 - 10. Published by the author, Pretoria." type="journal article" year="1942">Janse (1942)</bibRefCitation>
noted that in the 14 years between capture of the female and the male, the original bush had been cleared away and replaced by a wattle plantation. The male was found nearer the mountain side where the original vegetation was still intact. Clearly additional sites in the district should be examined to determine whether
<taxonomicName id="71C44D69FFD6FFC1FAB0A1C5AAE6FD65" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFD6FFC1FAB0A1C5AAE6FD65" italics="true" pageId="13" pageNumber="14">A. pseliacma</emphasis>
</taxonomicName>
still survives.
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<paragraph id="B67B36EAFFD6FFC1FF3BA18AAF6EFC81" blockId="13.[151,1437,151,789]" box="[197,1387,763,789]" pageId="13" pageNumber="14">
<emphasis id="84B0EAF8FFD6FFC1FF3BA18AAB5BFC81" bold="true" box="[197,350,763,788]" pageId="13" pageNumber="14">Distribution.</emphasis>
<collectingCountry id="CED3767AFFD6FFC1FE9BA18DABFDFC81" box="[357,504,764,788]" name="South Africa" pageId="13" pageNumber="14">South Africa</collectingCountry>
, Kwazulu-Natal, Karkloof Falls, near Howick. Known only from
<typeStatus id="697F8848FFD6FFC1FB24A18CAF0FFC80" box="[1242,1290,765,789]" pageId="13" pageNumber="14">type</typeStatus>
locality.
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<mods:identifier id="2A2FCF78B4D7FE362F7BDBC9C7A7E041" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="72347157C32E9369DB113ABBC768749B" type="Zenodo-Dep">203553</mods:identifier>
</mods:mods>
<treatment id="3E6D87FCFFDAFFC9FF69A0EEAE1BFCC9" ID-DOI="http://doi.org/10.5281/zenodo.6189649" ID-GBIF-Taxon="119388867" ID-Zenodo-Dep="6189649" LSID="urn:lsid:plazi:treatment:3E6D87FCFFDAFFC9FF69A0EEAE1BFCC9" httpUri="http://treatment.plazi.org/id/3E6D87FCFFDAFFC9FF69A0EEAE1BFCC9" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2">
<subSubSection id="FEDE6561FFDAFFCDFF69A0EEABEDFC2D" box="[151,488,927,952]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="B67B36EAFFDAFFCDFF69A0EEABEDFC2D" blockId="1.[151,488,927,952]" box="[151,488,927,952]" pageId="1" pageNumber="2">
<heading id="ED338186FFDAFFCDFF69A0EEABEDFC2D" bold="true" box="[151,488,927,952]" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
<taxonomicName id="71C44D69FFDAFFCDFF69A0EEABEDFC2D" ID-CoL="8ZZ76" authority="Meyrick, 1921" authorityName="Meyrick" authorityYear="1921" box="[151,488,927,952]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDAFFCDFF69A0EEABEDFC2D" bold="true" box="[151,488,927,952]" pageId="1" pageNumber="2">
<emphasis id="84B0EAF8FFDAFFCDFF69A0EEAB33FC2D" bold="true" box="[151,310,927,952]" italics="true" pageId="1" pageNumber="2">Agrionympha</emphasis>
Meyrick, 1921
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDAFFCDFF69A099A84DFC68" box="[151,584,999,1022]" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="B67B36EAFFDAFFCDFF69A099A84DFC68" blockId="1.[151,584,999,1022]" box="[151,584,999,1022]" pageId="1" pageNumber="2">
<emphasis id="84B0EAF8FFDAFFCDFF69A099AB1EFC6B" bold="true" box="[151,283,999,1022]" pageId="1" pageNumber="2">
<typeStatus id="697F8848FFDAFFCDFF69A099AACFFC68" box="[151,202,1000,1021]" pageId="1" pageNumber="2">Type</typeStatus>
species
</emphasis>
<taxonomicName id="71C44D69FFDAFFCDFEDDA098A841FC68" authority="Meyrick, 1921" authorityName="Meyrick" authorityYear="1921" box="[291,580,1000,1022]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFDAFFCDFEDDA098ABA3FC6B" box="[291,422,1000,1022]" italics="true" pageId="1" pageNumber="2">A. pseliacma</emphasis>
Meyrick, 1921
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDAFFCFFF69A75BAB8AFA57" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph id="B67B36EAFFDAFFCDFF69A75BAEDBFA13" blockId="1.[151,1437,1066,2026]" pageId="1" pageNumber="2">
<emphasis id="84B0EAF8FFDAFFCDFF69A75BAB24FBD6" bold="true" box="[151,289,1066,1091]" pageId="1" pageNumber="2">Description</emphasis>
.
<emphasis id="84B0EAF8FFDAFFCDFECFA75AAB6BFBD6" box="[305,366,1067,1091]" italics="true" pageId="1" pageNumber="2">Head</emphasis>
. Vertex between ocelli and compound eyes densely microtrichiated. Anterior tentorial arm at base with a high dorsolateral crest, the invagination line of which is demarcated as a curved sulcus extending upwards/outwards from the tentorial pit; dorsal tentorial arm small but distinct, situated some distance behind/ above the inner end of the crest. Antennal scape (
<figureCitation id="2EFF2A6FFFDAFFCDFD2EA7E7A916FB3B" box="[720,787,1174,1198]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
F) elongate and swollen (length ~1.5x first flagellomere) with constriction near midlength; pedicel swollen, width equal to scape, equidimensional (length 1.0x first flagellomere). Flagellum in male of 4045 flagellomeres, scale-covered on 23 basal flagellomeres. Female antennae shorter, 2633 flagellomeres with 310 scaled. Fully developed ascoids (
<figureCitation id="2EFF2A6FFFDAFFCDFBE3A673AE66FA8F" box="[1053,1123,1282,1306]" captionStart="FIGURE 4" captionStartId="10.[151,250,1611,1634]" captionTargetBox="[198,1390,705,1611]" captionTargetId="figure@10.[197,1390,700,1611]" captionTargetPageId="10" captionText="FIGURE 4. Structural detail of Agrionympha species: A, A. capensis, antennal flagellomere with ascoid sensilla (SEM); B, A. capensis, segment V gland (SEM); C, A. sagittella, hair-sensilla and micro-scales on dorsum of male tergum VIII (SEM); D, A. kroonella, hair sensilla on dorsum of segment IX (SEM). asb, branch of ascoid sensilla; as, central disc of ascoid sensilla; ped, raised peduncle of segment V gland, S iv, fourth sternite; S v, fifth sternite; sc, long fluted piliform scale. Scale bars: A, B = 0.01 mm; C, D = 0.005 mm." httpUri="https://zenodo.org/record/203557/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
A) with about 15 radiating branches, equally spaced around the base. Fenced sensilla coeloconica not conspicuous, probably generally absent. Mandibles and armature of labral epipharynx of well-developed micropterigid type repeatedly described, in most detail for
<taxonomicName id="71C44D69FFDAFFCDFEFAA61EAB8CFA13" box="[260,393,1391,1414]" class="Insecta" family="Micropterigidae" genus="Micropterix" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDAFFCDFEFAA61EAB8CFA13" box="[260,393,1391,1414]" italics="true" pageId="1" pageNumber="2">Micropterix</emphasis>
</taxonomicName>
(
<bibRefCitation id="D2554B1BFFDAFFCDFE69A61FA86CFA13" author="Hannemann" box="[407,617,1390,1414]" pageId="1" pageNumber="20" refString="Hannemann, H. J. (1956) Die Kopfmuskulatur von Micropteryx calthella (L.) (Lep.). Morphologie und Funktion. Zoologische Jahrbucher (Anatomie), 75, 177 - 206." type="journal article" year="1956">Hannemann, 1956</bibRefCitation>
;
<bibRefCitation id="D2554B1BFFDAFFCDFD8BA61FA9ABFA13" author="Chauvin" box="[629,942,1390,1414]" pageId="1" pageNumber="20" refString="Chauvin, G. &amp; Faucheux, M. (1981) Les pieces buccales et leurs recepteurs sensoriels chez l'imago de Micropterix calthella L. (Lepidoptera: Micropterigidae). International Journal of Insect Morphology and Embryology, 10, 425 - 439. Gibbs, G. W., Kobayashi, Y., Suzuki, H., Hashimoto, S., Lees, D. C., Sugimoto, M. &amp; Saigusa, T. (2004) Molecular phylogeny of Micropterigidae. Proceedings of XXII International Congress of Entomology. 2004, Brisbane, Australia. Abstract only. Gibbs, G. W. (2010) Micropterigidae (Lepidoptera) of SW Pacific: a revision with the establishment of five new genera from Australia, New Caledonia and New Zealand. Zootaxa, 2520, 1 - 48." type="journal article" year="1981">Chauvin &amp; Faucheux, 1981</bibRefCitation>
). Labial palp 2-segmented.
</paragraph>
<paragraph id="B67B36EAFFDAFFCDFF39A6E5A8A2FA3F" blockId="1.[151,1437,1066,2026]" box="[199,679,1426,1451]" pageId="1" pageNumber="2">
<emphasis id="84B0EAF8FFDAFFCDFF39A6E5AAF9FA3E" box="[199,252,1428,1451]" italics="true" pageId="1" pageNumber="2">Legs</emphasis>
. Fore tibial epiphysis well developed.
</paragraph>
<paragraph id="B67B36EAFFDAFFCDFF39A6C6AE4DF87F" blockId="1.[151,1437,1066,2026]" pageId="1" pageNumber="2">
<emphasis id="84B0EAF8FFDAFFCDFF39A6C6AB68FA5A" box="[199,365,1463,1487]" italics="true" pageId="1" pageNumber="2">Wing structure</emphasis>
(
<figureCitation id="2EFF2A6FFFDAFFCDFE83A6C7ABC6FA5B" box="[381,451,1462,1486]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
). Most species have the + rounded-lanceolate wings typical of
<taxonomicName id="71C44D69FFDAFFCDFB6FA6C7AF4CFA5B" box="[1169,1353,1462,1486]" class="Insecta" family="Micropterigidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Micropterigidae</taxonomicName>
; a very narrow and pointed wing shape in
<taxonomicName id="71C44D69FFDAFFCDFDE2A6AAA899FA66" box="[540,668,1499,1523]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDAFFCDFDE2A6AAA899FA66" box="[540,668,1499,1523]" italics="true" pageId="1" pageNumber="2">A. capensis</emphasis>
</taxonomicName>
(males in particular) is extreme in the family. Forewing (FW) venation with Sc and R forked; Rs3 and 4 sessile (free) in all but
<taxonomicName id="71C44D69FFDAFFCDFC65A68FAE1AF983" box="[923,1055,1534,1558]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDAFFCDFC65A68FAE1AF983" box="[923,1055,1534,1558]" italics="true" pageId="1" pageNumber="2">A. capensis</emphasis>
</taxonomicName>
(
<figureCitation id="2EFF2A6FFFDAFFCDFBCCA68FAE7DF983" box="[1074,1144,1534,1558]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
B) where these veins are stalked; Rs4 to apex or termen/dorsum. In the four examined
<taxonomicName id="71C44D69FFDAFFCDFC98A552A9E2F9AE" box="[870,999,1571,1595]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDAFFCDFC98A552A9E2F9AE" box="[870,999,1571,1595]" italics="true" pageId="1" pageNumber="2">A. capensis</emphasis>
</taxonomicName>
males (and in one of the [four] examined more broad-winged females of that species) the anterior M branch anastomoses with the stem of Rs3&amp;4, and the free terminal M1 hence is stalked on the latter. A M3-CuA cross-vein may or may not be present. It can be variably developed within a single species (also within the same sex), and so can a Sc-R cross-vein as well as the crossvein/vein which connect the base of CuA to the bases of M and CuP, respectively. In the hindwing (HW) venation the intriguing double configuration of Sc first described from micropterigids by
<bibRefCitation id="D2554B1BFFDAFFCDFBB1A5A7AEF9F97B" author="Philpott" box="[1103,1276,1750,1774]" pageId="1" pageNumber="21" refString="Philpott, A. (1923) A study of the venation of the New Zealand species of Micropterygidae. Transactions and Proceedings of the New Zealand Institute, 54, 155 - 161." type="journal article" year="1923">Philpott (1923)</bibRefCitation>
is sometimes very prominent (
<figureCitation id="2EFF2A6FFFDAFFCDFEAAA58BAB93F887" box="[340,406,1786,1810]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
E), and before the apex the two, now diverging, Sc branches may be united by an apparent cross-vein (
<figureCitation id="2EFF2A6FFFDAFFCDFEE6A46FAB5AF8A3" box="[280,351,1822,1846]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="1" pageNumber="2">Figs 3</figureCitation>
C, D). Exceptionally, however, in one male specimen [of three of which slide mounts were examined] of
<taxonomicName id="71C44D69FFDAFFCDFF0AA432AB71F8CE" box="[244,372,1859,1883]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDAFFCDFF0AA432AB71F8CE" box="[244,372,1859,1883]" italics="true" pageId="1" pageNumber="2">A. capensis</emphasis>
</taxonomicName>
the anterior Sc branch becomes obliterated just beyond the bifurcation. Discrete stem of vein R at most present in the form of short spur (referred to as recurrent R vein by
<bibRefCitation id="D2554B1BFFDAFFCDFBEFA417AEB4F8EB" author="Philpott" box="[1041,1201,1894,1918]" pageId="1" pageNumber="21" refString="Philpott, A. (1923) A study of the venation of the New Zealand species of Micropterygidae. Transactions and Proceedings of the New Zealand Institute, 54, 155 - 161." type="journal article" year="1923">Philpott, 1923</bibRefCitation>
) immediately before its anastomosis with Sc2. 13 frenulum bristles. Following veins overall similar to FW counterparts. The putative base of A1 is indistinguishably fused with CuP, and a free distal part of this vein may or may not be developed; when present it is markedly crossvein-like, oblique and soon anastomosing with A2.
</paragraph>
<paragraph id="B67B36EAFFD9FFCEFF39A3E8A9EEFEAA" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="3">
<emphasis id="84B0EAF8FFD9FFCEFF39A3E8AB43FF25" box="[199,326,153,176]" italics="true" pageId="2" pageNumber="3">FW pattern</emphasis>
(
<figureCitation id="2EFF2A6FFFD9FFCEFEABA3E6AB9FFF3A" box="[341,410,151,175]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). Very conservative and reminiscent of many European and Australian species in the family. Somewhat uniform throughout the genus, but with clear specific differences in details. Ground colour dark brown to black, with purplish-coppery iridescence and brilliant silvery-white maculation which, when most complete (e.g.
<figureCitation id="2EFF2A6FFFD9FFCEFF69A275AADCFE89" box="[151,217,260,284]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
B), consist of four components: a claval mark (along the claval furrow, i.e., the concavity within which CuP is lying), a median band, a postmedian band, and a subapical band (or spot).
</paragraph>
<paragraph id="B67B36EAFFD9FFCEFF39A23DAEACFDA9" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="3">
<emphasis id="84B0EAF8FFD9FFCEFF39A23DABB3FEF1" box="[199,438,332,356]" italics="true" pageId="2" pageNumber="3">Scale colour on body</emphasis>
. Head variable on vertex, frons and antennae (see descriptions) but with a black lateral streak extending from the base of the antenna to the area below the compound eyes in all species; vertex with long tufts of piliform scales. Dorsum of thorax with flat black or brown-black scales, usually with metallic iridescence (see descriptions), lateral walls with silvery white scales, concolorous with fore and midfemora. Tegulae either black (in 3 species) or whitish (in 5 species) clothed in a mix of elongate flat scales and long piliform scales. Legs variable (see descriptions); but fore and mid femora glistening white with grey distal joints; hindtibiae uniformly dark brown-black, but fore and midtibiae and tarsi with variable pale bands. Abdomen black.
</paragraph>
<paragraph id="B67B36EAFFD9FFCEFF39A139ABC2FD59" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="3">
<emphasis id="84B0EAF8FFD9FFCEFF39A139ABAEFDF5" box="[199,427,584,608]" italics="true" pageId="2" pageNumber="3">Pregenital abdomen</emphasis>
. Dorsum I with or without a trace of a narrow transverse sclerite, unscaled. Sternum V gland (
<figureCitation id="2EFF2A6FFFD9FFCEFF18A11DAB32FD11" box="[230,311,620,644]" captionStart="FIGURE 4" captionStartId="10.[151,250,1611,1634]" captionTargetBox="[198,1390,705,1611]" captionTargetId="figure@10.[197,1390,700,1611]" captionTargetPageId="10" captionText="FIGURE 4. Structural detail of Agrionympha species: A, A. capensis, antennal flagellomere with ascoid sensilla (SEM); B, A. capensis, segment V gland (SEM); C, A. sagittella, hair-sensilla and micro-scales on dorsum of male tergum VIII (SEM); D, A. kroonella, hair sensilla on dorsum of segment IX (SEM). asb, branch of ascoid sensilla; as, central disc of ascoid sensilla; ped, raised peduncle of segment V gland, S iv, fourth sternite; S v, fifth sternite; sc, long fluted piliform scale. Scale bars: A, B = 0.01 mm; C, D = 0.005 mm." httpUri="https://zenodo.org/record/203557/files/figure.png" pageId="2" pageNumber="3">Figs. 4</figureCitation>
B, D) orifice on a relatively large (about
<quantity id="713C9B0FFFD9FFCEFCF4A11DA964FD11" box="[778,865,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="3" unit="mm" value="0.4">0.4 mm</quantity>
diam.), modestly elevated, near-circular protuberance; markedly microtrichiated on discal area around the orifice, with 812 long fluted piliform scales concentrated around its posterior border.
</paragraph>
<paragraph id="B67B36EAFFD9FFCEFF39A1A9A819F90D" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="3">
<emphasis id="84B0EAF8FFD9FFCEFF39A1A9AB69FD65" box="[199,364,728,752]" italics="true" pageId="2" pageNumber="3">Male genitalia</emphasis>
(
<figureCitation id="2EFF2A6FFFD9FFCEFE83A1A6ABFFFD7A" box="[381,506,727,751]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="14.[151,250,1457,1480]" captionStartId-1="16.[151,250,1415,1438]" captionStartId-2="17.[151,250,1478,1501]" captionStartId-3="18.[151,250,1913,1936]" captionTargetBox-0="[160,1416,529,1421]" captionTargetBox-1="[311,1355,490,1414]" captionTargetBox-2="[259,1325,201,1478]" captionTargetBox-3="[313,1332,581,1845]" captionTargetId-0="figure@14.[160,1427,519,1454]" captionTargetId-1="figure@16.[223,1363,483,1474]" captionTargetId-2="figure@17.[254,1332,193,1527]" captionTargetId-3="figure@18.[244,1340,573,1969]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIGURE 7. Agrionympha capensis Whalley, genitalia: A E, male, Storms River: A, ventral; B, lateral; C, dorsal; D, phallus lateral; E, phallus dorsal. F G, female, Bloukrans Pass: F, extended terminal segments of abdomen; G, internal female genitalia. cb, corpus bursae; dph, distal portion of phallus; dsp, ductus of spermatheca; go, gonopore with radial folds; lag, lagena; lap, lappet, mpl, median plate; pap, papilla; pph, proximal portion of phallus; scl ix, ninth sclerite; scl x, tenth sclerite; S viii, eighth sternite; spm, spermatheca; T viii, eighth tergite; T x, tenth tergum; utr, distal bulb of utriculus; va, valva; vb, ventral bulb." captionText-1="FIGURE 8. Agrionympha genitalia. A D, A. fuscoapicella sp. nov. male, Hogsback: A, lateral, inset - ventral view of median plate; B, dorsal, C, phallus lateral; D, phallus dorsal. E F, A. jansella sp. nov. female, Karkloof Falls: E, extended terminal segments of abdomen; F, internal female genitalia." captionText-2="FIGURE 9. Agrionympha male genitalia. A D, A. karoo, sp. nov. Toorberg: A, lateral, inset ventral view of median plate; B, dorsal; C, phallus lateral; D, phallus dorsal. E H, A. kroonella, sp. nov. Mariepskop: E, lateral, inset ventral view of median plate; F, dorsal; G, phallus lateral; H, phallus dorsal." captionText-3="FIGURE 10. Agrionympha male genitalia. A B, A. pseliacma, Meyrick, Karkloof Falls: A, lateral view, from slide 3830, Transvaal Museum, Pretoria; B, detail of phallus, lateral. C G, A. sagittella, sp. nov. Hogsback: C, lateral; D, dorsal; E, ventral; F, phallus, lateral; G, phallus dorsal. H J, Agrionympha species 1 , Diepwalle Forest: H, lateral; I, phallus lateral; J, phallus dorsal." httpUri-0="https://zenodo.org/record/203560/files/figure.png" httpUri-1="https://zenodo.org/record/203561/files/figure.png" httpUri-2="https://zenodo.org/record/203562/files/figure.png" httpUri-3="https://zenodo.org/record/203563/files/figure.png" pageId="2" pageNumber="3">Figs. 710</figureCitation>
). Sternum VIII remnants present as a pair of small, but distinct, isolated and variably shaped sclerotisations bearing some minute setae. Segment IX sclerite (= vinculum) forming a dorsally open ring, its thickened anteroventral margin extended forward, telescoping into sternum of segment VII; anterodorsal margins straight or very slightly concave and neither pigmented nor markedly thickened; posterior margin strongly convex, overlapping base of valve. A cluster of specialised sensilla (circular elevations surrounding a small central hair-like process,
<figureCitation id="2EFF2A6FFFD9FFCEFE9EA0FDABA6FC31" box="[352,419,908,932]" captionStart="FIGURE 4" captionStartId="10.[151,250,1611,1634]" captionTargetBox="[198,1390,705,1611]" captionTargetId="figure@10.[197,1390,700,1611]" captionTargetPageId="10" captionText="FIGURE 4. Structural detail of Agrionympha species: A, A. capensis, antennal flagellomere with ascoid sensilla (SEM); B, A. capensis, segment V gland (SEM); C, A. sagittella, hair-sensilla and micro-scales on dorsum of male tergum VIII (SEM); D, A. kroonella, hair sensilla on dorsum of segment IX (SEM). asb, branch of ascoid sensilla; as, central disc of ascoid sensilla; ped, raised peduncle of segment V gland, S iv, fourth sternite; S v, fifth sternite; sc, long fluted piliform scale. Scale bars: A, B = 0.01 mm; C, D = 0.005 mm." httpUri="https://zenodo.org/record/203557/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
C) is present on dorsum IX in all species except
<taxonomicName id="71C44D69FFD9FFCEFC34A0FDAE4FFC31" box="[970,1098,908,932]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFD9FFCEFC34A0FDAE4FFC31" box="[970,1098,908,932]" italics="true" pageId="2" pageNumber="3">A. capensis</emphasis>
</taxonomicName>
and
<taxonomicName id="71C44D69FFD9FFCEFB7DA0FDAF0EFC31" box="[1155,1291,908,932]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sagittella">
<emphasis id="84B0EAF8FFD9FFCEFB7DA0FDAF0EFC31" box="[1155,1291,908,932]" italics="true" pageId="2" pageNumber="3">A. sagittella</emphasis>
</taxonomicName>
; in the latter what may be serial homologues occur on an anterior zone of tergum VIII (
<figureCitation id="2EFF2A6FFFD9FFCEFC13A0DEAE36FC52" box="[1005,1075,943,967]" captionStart="FIGURE 4" captionStartId="10.[151,250,1611,1634]" captionTargetBox="[198,1390,705,1611]" captionTargetId="figure@10.[197,1390,700,1611]" captionTargetPageId="10" captionText="FIGURE 4. Structural detail of Agrionympha species: A, A. capensis, antennal flagellomere with ascoid sensilla (SEM); B, A. capensis, segment V gland (SEM); C, A. sagittella, hair-sensilla and micro-scales on dorsum of male tergum VIII (SEM); D, A. kroonella, hair sensilla on dorsum of segment IX (SEM). asb, branch of ascoid sensilla; as, central disc of ascoid sensilla; ped, raised peduncle of segment V gland, S iv, fourth sternite; S v, fifth sternite; sc, long fluted piliform scale. Scale bars: A, B = 0.01 mm; C, D = 0.005 mm." httpUri="https://zenodo.org/record/203557/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
C). Valvae simple, slender, bent upwards, weakly tapering toward a rounded apex; ratio of length:width at mid-length 3.65.7; lacking any medial projections or processes but with an area of stiff retro-bristles near the apex. Median plate well developed, sometimes with anterior part more or less strongly darkened. Tergum X plate completely divided in mid-line, posteriorly produced into a pair of rounded sclerotised uncus lobes. Lateroventral walls of anal cone with a pair of irregular sclerotised plates immediately below the tergal plates and largely concealed by the latter, bearing numerous large microtrichia but no setae; membranous distal part of anal cone small, inconspicuous. Phallus short, 1.31.7 of ventromedial length of segment IX sclerotisation. Distal/posterior part of phallus tube equal to or shorter (1.00.6x) than anterior part, enclosed in sclerotised phallocrypt, subdivided into head, neck and body regions, the latter with heavily-sclerotised walls produced into a pair of small arrowhead-like lateral lappets level with the proximal margin of the gonopore; neck more or less constricted in dorsal view but not apparent in lateral view; slightly swollen ventral bulb projecting beyond gonopore, rather thickwalled but unsclerotised (mesocuticular), not appearing as a separate branch; gonopore generally heart-shaped or circular, orientated dorsally or tilted slightly towards apex, bordered by lightly sclerotised radial folds. Just inside the gonopore the cuticular floor of the ejaculatory duct may be markedly thickened and textured, and may be produced into a small sclerotised apodemal formation (
<figureCitation id="2EFF2A6FFFD9FFCEFF61A6BDAAE1FA71" box="[159,228,1484,1508]" captionStart="FIGURE 5" captionStartId="11.[151,250,1629,1652]" captionTargetBox="[195,1374,1262,1585]" captionTargetId="figure@11.[195,1391,1246,1608]" captionTargetPageId="11" captionText="FIGURE 5. Structural detail of Agrionympha capensis: A, ventral valve muscle (*) from IX sclerotisation, medial view. Whole mount preparation of specimen sectioned sagittally, chlorazol black staining; B. Agrionympha capensis, sagittal section of phallic apex, showing ejaculatory duct with thickened cuticular intima (*) and apodemal seclerotization (arrow)." httpUri="https://zenodo.org/record/203558/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
B); retractor muscle fibres inserting on this posterior duct portion, such as have been identified in a
<collectingCountry id="CED3767AFFD9FFCEFAA7A6BDAACEF99C" name="Madagascar" pageId="2" pageNumber="3">Malagasy</collectingCountry>
micropterigid genus, were not observable in histological sections. Posterior phallocrypt with scaly texture, raised free edges of the micro-scales with fine comb-like margins. Valve musculature originating on (here incomplete) segment IX ring comprising both the unusual (in micropterigids) ventral (
<figureCitation id="2EFF2A6FFFD9FFCEFBE8A549AE52F9C5" box="[1046,1111,1592,1616]" captionStart="FIGURE 5" captionStartId="11.[151,250,1629,1652]" captionTargetBox="[195,1374,1262,1585]" captionTargetId="figure@11.[195,1391,1246,1608]" captionTargetPageId="11" captionText="FIGURE 5. Structural detail of Agrionympha capensis: A, ventral valve muscle (*) from IX sclerotisation, medial view. Whole mount preparation of specimen sectioned sagittally, chlorazol black staining; B. Agrionympha capensis, sagittal section of phallic apex, showing ejaculatory duct with thickened cuticular intima (*) and apodemal seclerotization (arrow)." httpUri="https://zenodo.org/record/203558/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
A) and the more widespread, but also not ubiquitous, dorsal set; dorsal muscle inserting on membranous body wall just proximad to/above valve base rather than on the latter itself.
</paragraph>
<paragraph id="B67B36EAFFD9FFCFFF39A5D5AAFDFE3A" blockId="2.[151,1437,151,2012]" lastBlockId="3.[151,1437,151,2014]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="84B0EAF8FFD9FFCEFF39A5D5AB82F929" box="[199,391,1700,1724]" italics="true" pageId="2" pageNumber="3">Female genitalia</emphasis>
(
<figureCitation id="2EFF2A6FFFD9FFCEFE69A5D5ABDEF929" box="[407,475,1700,1724]" captionStart="FIGURE 7" captionStartId="14.[151,250,1457,1480]" captionTargetBox="[160,1416,529,1421]" captionTargetId="figure@14.[160,1427,519,1454]" captionTargetPageId="14" captionText="FIGURE 7. Agrionympha capensis Whalley, genitalia: A E, male, Storms River: A, ventral; B, lateral; C, dorsal; D, phallus lateral; E, phallus dorsal. F G, female, Bloukrans Pass: F, extended terminal segments of abdomen; G, internal female genitalia. cb, corpus bursae; dph, distal portion of phallus; dsp, ductus of spermatheca; go, gonopore with radial folds; lag, lagena; lap, lappet, mpl, median plate; pap, papilla; pph, proximal portion of phallus; scl ix, ninth sclerite; scl x, tenth sclerite; S viii, eighth sternite; spm, spermatheca; T viii, eighth tergite; T x, tenth tergum; utr, distal bulb of utriculus; va, valva; vb, ventral bulb." httpUri="https://zenodo.org/record/203560/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
F). Relatively uniform among the species. Segment VIII tergite and sternite independent and unmodified; segments IX and X capable of extreme elongation but normally IX is telescoped inside segment VIII; segment IX sclerite a discontinuous ring, with a gap along mid-dorsal line, the gap being wider anteriorly than posteriorly, the sclerite slightly higher than long. Melanised terminal plates (lateral sclerites of segment X) vary from equidimensional to longer than high (about 1.6x), sometimes appearing emarginated along anterior margin. In
<taxonomicName id="71C44D69FFD9FFCEFE91A429A823F8E5" box="[367,550,1880,1904]" class="Insecta" family="Micropterigidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Micropterigidae</taxonomicName>
, the spermatheca arises from the genital chamber inside a thickened and highly malleable, collar-like papilla, often with complexly folded latero-ventral walls which can be distorted according to the reproductive status of the individual. It is situated adjacent to the entrance to corpus bursae which, in
<taxonomicName id="71C44D69FFD9FFCEFA9FA4D1AAFBF849" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFD9FFCEFA9FA4D1AAFBF849" italics="true" pageId="2" pageNumber="3">Agrionympha</emphasis>
</taxonomicName>
, is not constricted at this point. The spermatheca itself (
<figureCitation id="2EFF2A6FFFD9FFCEFC7FA4B5A9C0F849" box="[897,965,1988,2012]" captionStart="FIGURE 7" captionStartId="14.[151,250,1457,1480]" captionTargetBox="[160,1416,529,1421]" captionTargetId="figure@14.[160,1427,519,1454]" captionTargetPageId="14" captionText="FIGURE 7. Agrionympha capensis Whalley, genitalia: A E, male, Storms River: A, ventral; B, lateral; C, dorsal; D, phallus lateral; E, phallus dorsal. F G, female, Bloukrans Pass: F, extended terminal segments of abdomen; G, internal female genitalia. cb, corpus bursae; dph, distal portion of phallus; dsp, ductus of spermatheca; go, gonopore with radial folds; lag, lagena; lap, lappet, mpl, median plate; pap, papilla; pph, proximal portion of phallus; scl ix, ninth sclerite; scl x, tenth sclerite; S viii, eighth sternite; spm, spermatheca; T viii, eighth tergite; T x, tenth tergum; utr, distal bulb of utriculus; va, valva; vb, ventral bulb." httpUri="https://zenodo.org/record/203560/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
G) is subdivided into three parts; a thickwalled, heavily staining basal duct, a more voluminous thin-walled utriculus, and a terminal lagena. In
<taxonomicName id="71C44D69FFD8FFCFFA9FA3E9AAFBFF41" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFD8FFCFFA9FA3E9AAFBFF41" italics="true" pageId="3" pageNumber="4">Agrionympha</emphasis>
</taxonomicName>
, the duct is narrow, slightly sinuous (both papilla and basal duct stain heavily with chlorazol black), its distal end abruptly bulbous; the utriculus of three sections but relatively short overall (no longer than 8th tergite), broadly connected to duct bulb through a thin-walled bladder-like proximal section, a short generalised middle section, leading through a constricted, often convoluted junction to its most distinctive feature—a large distal spherical bulb 2
<date id="C27A102AFFD8FFCFFEE8A23EAB27FEF2" box="[278,290,335,359]" pageId="3" pageNumber="4">3</date>
x diameter of middle section of utriculus but often collapsed so as to appear crescentic; a short lagena extends from the distal utricular bulb. Corpus bursae small, simple, without spines, signa, or other sclerotisations in the wall.
</paragraph>
<paragraph id="B67B36EAFFD8FFCFFF39A2CDAB8AFA57" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="4">
<emphasis id="84B0EAF8FFD8FFCFFF39A2CDAB84FE46" box="[199,385,444,467]" italics="true" pageId="3" pageNumber="4">Immature stages</emphasis>
(
<figureCitation id="2EFF2A6FFFD8FFCFFE6CA2CBABDDFE47" box="[402,472,442,466]" captionStart="FIGURE 6" captionStartId="13.[151,250,1305,1328]" captionTargetBox="[252,1339,848,1247]" captionTargetId="figure@13.[245,1342,844,1302]" captionTargetPageId="13" captionText="FIGURE 6. Larva of Agrionympha sp. nov. Mariepskop: A, habitus of second instar larva; B, trunk chaetotaxy. Scale bar 1 mm." httpUri="https://zenodo.org/record/203559/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
). Early stage (probably second instar) larvae have been found (by GWG) at Mariepskop, Mpumalanga Province (formerly Transvaal), and Kogel Bay, Western Cape, almost certainly representing two species. Both are of the sabatincoid morphotype (Gibbs, 2010) characterised by the possession of eight pairs of abdominal spiracles; the trunk broadly hexagonal in cross-section; and a cryptic pigment pattern (pigmentation disappearing after several years of storage in 70% ethanol) integument with a honeycomb of raised ridges. The head can be totally recessed into the prothorax as in all micropterigid larvae. The cranial ecdysial line is distinct, a cluster of 5 stemmata occurs on each side within a melanised cage, and a medial melanisation is present on the hypostomal bridge. The thoracic legs are short, three-segmented, lack a distinct melanised coxa and the femur carries an inflatable membraneous sac on its inner side near the base (see Gibbs, 2010). Abdominal prolegs are lacking. Cranial chaetotaxy was not examined. Macro-setae of the trunk could be scored with accuracy from both taxa but confirming the presence or absence of short micro-setae was not always possible. The macro-setae are relatively short, thickened, with clubbed apices, D1 arising from low integumental flanges on the conspicuously honey-combed integument. These larvae of
<taxonomicName id="71C44D69FFD8FFCFFD75A01AA921FC16" box="[651,804,875,899]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFD8FFCFFD75A01AA921FC16" box="[651,804,875,899]" italics="true" pageId="3" pageNumber="4">Agrionympha</emphasis>
</taxonomicName>
are notable for the reduction of D2 setae on abdominal segments to minute spherical knobs. Prothorax with 9 pairs of setae, directed forward, six aligned around the margin of the head recess fold: dorsal and lateral setae as long as any on the trunk; five pairs on dorsum, with two forming a longitudinal row either side of mid-dorsal line (D1 &amp; D2, with D2 on the anterior head recess margin), and three forming a discrete oblique row along a dorso-lateral ridge (anterior-most are XD1, XD2, posterior L1), the two remaining L setae are aligned vertically at the level of the spiracle (L2 &amp; L3) close to the margin of the head recess; a single short SV seta is situated dorsad to the tibiotarsus insertion and a small thinner seta is located on the margin of the head recess at the level of the leg base. No
<date id="C27A102AFFD8FFCFFC90A716A995FBEA" box="[878,912,1127,1151]" pageId="3" pageNumber="4">V1</date>
seta could be found associated with the coxae on these larvae. Mesothorax with five pairs of setae, aligned around the middle of the segment: the D setae longer than L setae, a much smaller spherical SV seta; D1 and D
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vertical line dorsally; L1 and L2 just above the lateral flange of the trunk at level of spiracles, more or less vertically aligned with the more ventral of the pair slightly forward; the single SV seta dorsad of the leg. Four pairs of setae on the metathorax are similarly arranged except that the D2 seta is absent. Abdominal segments 18 each with only three pairs of setae aligned around the middle of the segments: two D setae, the minute spherical D2 seta is repeated along all trunk segments to A8; a single L setae, about half the length of D1 on the lateral flange slightly below and posterior to the spiracle. Two pairs of clubbed setae occur on A
<quantity id="713C9B0FFFD8FFCFFEADA6F6AB7BFA0A" box="[339,382,1415,1439]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="4" unit="in" value="9.0">9 in</quantity>
the dorsal region, the more ventral one slightly forward and shorter; A10 with a single pair of minute globular setae.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFD8FFC8FF39A6BEAE65FCA6" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="discussion">
<paragraph id="B67B36EAFFD8FFCFFF39A6BEA8C5F8E7" blockId="3.[151,1437,151,2014]" pageId="3" pageNumber="4">
<emphasis id="84B0EAF8FFD8FFCFFF39A6BEABEBFA72" bold="true" box="[199,494,1487,1511]" pageId="3" pageNumber="4">Remarks on morphology</emphasis>
. The subcosto-radial region of the micropterigid hindwing has long attracted particular attention, and a configuration observed in some specimens of
<taxonomicName id="71C44D69FFD8FFCFFC7EA682A9FCF99E" box="[896,1017,1523,1547]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jansella">
<emphasis id="84B0EAF8FFD8FFCFFC7EA682A9FCF99E" box="[896,1017,1523,1547]" italics="true" pageId="3" pageNumber="4">A. jansella</emphasis>
</taxonomicName>
and
<taxonomicName id="71C44D69FFD8FFCFFBD1A682AEB3F99E" box="[1071,1206,1523,1547]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sagittella">
<emphasis id="84B0EAF8FFD8FFCFFBD1A682AEB3F99E" box="[1071,1206,1523,1547]" italics="true" pageId="3" pageNumber="4">A. sagittella</emphasis>
</taxonomicName>
proved to be of particular note. Here the two branches of the doubled Sc are connected by a short, oblique apparent cross-vein. Had it not been for the presence of the R spur (distinct in the
<taxonomicName id="71C44D69FFD8FFCFFCE2A54AA996F9C6" box="[796,915,1595,1619]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="jansella">
<emphasis id="84B0EAF8FFD8FFCFFCE2A54AA996F9C6" box="[796,915,1595,1619]" italics="true" pageId="3" pageNumber="4">A. jansella</emphasis>
</taxonomicName>
specimen illustrated in
<figureCitation id="2EFF2A6FFFD8FFCFFB63A54BAED8F9C7" box="[1181,1245,1594,1618]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
D, only just indicated in the
<taxonomicName id="71C44D69FFD8FFCFFEE3A52EABA1F9E2" box="[285,420,1631,1655]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sagittella">
<emphasis id="84B0EAF8FFD8FFCFFEE3A52EABA1F9E2" box="[285,420,1631,1655]" italics="true" pageId="3" pageNumber="4">A. sagittella</emphasis>
</taxonomicName>
specimen illustrated in
<figureCitation id="2EFF2A6FFFD8FFCFFD53A52EA8E8F9E2" box="[685,749,1631,1655]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
C) diverging from the posterior Sc branch immediately distad from the cross-vein, it would in this case have appeared straightforward to interpret the posterior Sc branch as vein R. Similarly, it is the presence of the spur in conspecific/congeneric moths which prompts us to reject the interpretation of the unusually discrete Sc2, itself devoid of any R-spur, of the
<taxonomicName id="71C44D69FFD8FFCFFC21A5BAAE63F976" box="[991,1126,1739,1763]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sagittella">
<emphasis id="84B0EAF8FFD8FFCFFC21A5BAAE63F976" box="[991,1126,1739,1763]" italics="true" pageId="3" pageNumber="4">A. sagittella</emphasis>
</taxonomicName>
specimen illustrated in
<figureCitation id="2EFF2A6FFFD8FFCFFA8EA5BBAAA6F892" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
E as vein R. Moreover: where a vestigial cross-vein between Sc and the R+Rs-stem is identifiable in the basal part of the wing, it joins vein Sc distad from the level where the doubling of the latter has become evident; this evidently speaks against the said cross-vein actually being the basal part of R, and the apparent posterior component of the doubled Sc being the more distal part of R.
</paragraph>
<paragraph id="B67B36EAFFD8FFC8FF39A4F1ABADFE64" blockId="3.[151,1437,151,2014]" lastBlockId="4.[151,1437,151,2035]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<bibRefCitation id="D2554B1BFFD8FFCFFF39A4F1ABAFF802" box="[199,426,1919,1944]" pageId="3" pageNumber="21" refString="Huang, D., Nel, A. &amp; Minet, J. (2010) A new family of moths from the Middle Jurassic. Acta Geologica Sinica (English Edition), 84, 874 - 885." type="journal article">
Huang
<emphasis id="84B0EAF8FFD8FFCFFEE5A4F1AB4AF802" box="[283,335,1919,1943]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. (2010)
</bibRefCitation>
suggested that the hindwing Sc is unforked in all non-agathiphagid
<taxonomicName id="71C44D69FFD8FFCFFB3CA40EAF4CF802" box="[1218,1353,1919,1943]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
(a few scattered exoporians excepted), and that what has been interpreted as a Sc
<quantity id="713C9B0FFFD8FFCFFC2EA4D3A9FFF82F" box="[976,1018,1954,1978]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="4" unit="in" value="2.0">2 in</quantity>
micropterigids is consistently a Sc-R crossvein. The cuticle thickening which
<bibRefCitation id="D2554B1BFFD8FFCFFD94A4B7A9B3F84B" author="Kristensen" box="[618,950,1990,2014]" pageId="3" pageNumber="21" refString="Kristensen, N. P. &amp; Nielsen, E. S. (1982) South American micropterigid moths: two new genera of the Sabatinca group. Entomologica Scandinavica, 13, 513 - 529." type="journal article" year="1982">Kristensen &amp; Nielsen (1982)</bibRefCitation>
interpreted as a true Sc-R cross-vein in a specimen of
<taxonomicName id="71C44D69FFDFFFC8FED6A3E9A8C5FF3A" authority="Meyrick, 1912" authorityName="Meyrick" authorityYear="1912" box="[296,704,151,176]" class="Insecta" family="Micropterigidae" genus="Sabatinca" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="calliarcha">
<emphasis id="84B0EAF8FFDFFFC8FED6A3E9A817FF25" box="[296,530,152,176]" italics="true" pageId="4" pageNumber="5">Sabatinca calliarcha</emphasis>
Meyrick, 1912
</taxonomicName>
may indeed likely be a teratological formation as maintained by Huang
<emphasis id="84B0EAF8FFDFFFC8FF17A3CCAB27FF41" box="[233,290,188,212]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
, but we do not consider these authors evidence (flat/slightly concave lower surface, absence of erect setae) for the apically-posteriorly-curving branch from Sc
<quantity id="713C9B0FFFDFFFC8FCD3A3AEA950FF62" box="[813,853,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="5" unit="in" value="1.0">1 in</quantity>
this taxon being a cross-vein rather than Sc2 to be compelling. Because the posterior component of the double Sc may be discrete as far basad as it occasionally is (afore-mentioned
<taxonomicName id="71C44D69FFDFFFC8FE9FA256ABEFFEAA" box="[353,490,295,319]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sagittella">
<emphasis id="84B0EAF8FFDFFFC8FE9FA256ABEFFEAA" box="[353,490,295,319]" italics="true" pageId="4" pageNumber="5">A. sagittella</emphasis>
</taxonomicName>
specimen in
<figureCitation id="2EFF2A6FFFDFFFC8FD7CA256A8B8FEAA" box="[642,701,295,319]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="4" pageNumber="5">Fig 3</figureCitation>
E), we consider it most unlikely that it can be a Sc-R cross-vein, even though in less extreme cases (
<taxonomicName id="71C44D69FFDFFFC8FDE3A23BA890FEF7" box="[541,661,330,354]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="jansella">A. jansella</taxonomicName>
specimen in
<figureCitation id="2EFF2A6FFFDFFFC8FCD6A23BA962FEF7" box="[808,871,330,354]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
D) its diverging apical part does have some resemblance (also position-wise) with the fore wing Sc-R cross-vein in certain micropterigids, as noted by Huang et al. Seemingly unquestionable forking (even twice in one case) of Sc in some members of the family was documented by
<bibRefCitation id="D2554B1BFFDFFFC8FF42A2C7ABE6FE5B" author="Hashimoto" box="[188,483,438,462]" pageId="4" pageNumber="21" refString="Hashimoto, S. (2006) A taxonomic study of the family Micropterigidae (Lepidoptera, Micropterigoidea) of Japan, with the phylogenetic relationship among the Northern Hemisphere genera. Bulletin of the Kitakyushu Museum of Natural History and Human History (Series A) Series A, 4, 39 - 109." type="journal article" year="2006">
Hashimoto (2006,
<figureCitation id="2EFF2A6FFFDFFFC8FE6DA2C7ABDCFE5B" box="[403,473,438,462]" captionStart="FIGURE 6" captionStartId="13.[151,250,1305,1328]" captionTargetBox="[252,1339,848,1247]" captionTargetId="figure@13.[245,1342,844,1302]" captionTargetPageId="13" captionText="FIGURE 6. Larva of Agrionympha sp. nov. Mariepskop: A, habitus of second instar larva; B, trunk chaetotaxy. Scale bar 1 mm." httpUri="https://zenodo.org/record/203559/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
)
</bibRefCitation>
; the 'doubling' of the hindwing Sc is in any case an intriguing phenomenon which deserves closer scrutiny.
</paragraph>
<paragraph id="B67B36EAFFDFFFC8FF39A28FA9E8FD15" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="5">It remains uncertain whether the oblique vein between the apical Sc branches here referred to as an apparent cross-vein (i.e., a Sc1-Sc2 cross-vein), or represents an anterior branch (R1) of a forked vein R, the posterior branch (R2) of which has, then, coalesced with Sc2. The position of the vestige (spur) of the R-stem is so close to the lower end of this apparent cross-vein that both possibilities seem open.</paragraph>
<paragraph id="B67B36EAFFDFFFC8FF39A1FDAE65FCA6" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="5">
The anterior portion of the micropterigid phallus is tightly surrounded by the sclerotised phallocrypt (and hence appearing double-walled), and the posterior portion beyond the sclerotised phallocrypt have been referred to as phallobase and aedeagus, respectively (
<bibRefCitation id="D2554B1BFFDFFFC8FD6CA1A2A950FD7E" author="Kristensen" box="[658,853,723,747]" pageId="4" pageNumber="21" refString="Kristensen, N. P. (1984) Skeletomuscular anatomy of the male genitalia of Epimartyria (Lepidoptera: Micropterigidae). Entomologica Scandinavica, 15, 97 - 112." type="journal article" year="1984">Kristensen, 1984</bibRefCitation>
). However, the two together are apparently homologous with the phallobase (or phallotheca) ascribed to the lepidopteran ground plan, while a true aedeagus is undeveloped in micropterigids as in most non-agathiphagid
<taxonomicName id="71C44D69FFDFFFC8FCFFA06AAE59FCA6" authority="Kristensen 2003" authorityName="Kristensen" authorityYear="2003" box="[769,1116,795,819]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="order">
Lepidoptera (
<bibRefCitation id="D2554B1BFFDFFFC8FC69A06AAE51FCA6" author="Kristensen" box="[919,1108,795,819]" pageId="4" pageNumber="21" refString="Kristensen, N. P. (2003) Skeleton and muscles: adults. In: Kristensen, N. P (Ed.) Lepidoptera: Moths and butterflies 2. Handbuch der Zoologie / Handbook of Zoology IV / 36, 39 - 131. Walter de Gruyter, Berlin &amp; New York." type="journal article" year="2003">Kristensen 2003</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDFFFC9FF39A04FAE2CFD32" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="B67B36EAFFDFFFC8FF39A04FAEF9FBE1" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="5">
<emphasis id="84B0EAF8FFDFFFC8FF39A04FAB44FCC2" bold="true" box="[199,321,830,855]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
<taxonomicName id="71C44D69FFDFFFC8FEB4A04EABE7FCC2" box="[330,482,831,855]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FEB4A04EABE7FCC2" box="[330,482,831,855]" italics="true" pageId="4" pageNumber="5">Agrionympha</emphasis>
</taxonomicName>
is unique within the family by the combined presence of a forked FW vein R (plesiomorphy shared with all S. Hemisphere genera except
<taxonomicName id="71C44D69FFDFFFC8FCF0A012AE4CFCEF" authority="Gibbs, 2010" authorityName="Gibbs" authorityYear="2010" box="[782,1097,866,891]" class="Insecta" family="Micropterigidae" genus="Austromartyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FCF0A012A9BFFCEE" box="[782,954,867,891]" italics="true" pageId="4" pageNumber="5">Austromartyria</emphasis>
Gibbs, 2010
</taxonomicName>
and an undescribed
<collectingCountry id="CED3767AFFDFFFC8FAD1A013AF99FCEF" box="[1327,1436,866,890]" name="Madagascar" pageId="4" pageNumber="5">Malagasy</collectingCountry>
genus), absence of a discrete full-length hindwing vein R and of a R/Rs fork (apomorphy shared with some SW Pacific genera), and complete longitudinal division of bilobed male tergum X plate (apomorphy elsewhere ocurring only in the S. American
<taxonomicName id="71C44D69FFDFFFC8FE53A0BEA849FC73" box="[429,588,975,998]" class="Insecta" family="Micropterigidae" genus="Hypomartyria" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FE53A0BEA849FC73" box="[429,588,975,998]" italics="true" pageId="4" pageNumber="5">Hypomartyria</emphasis>
</taxonomicName>
and one Australian
<taxonomicName id="71C44D69FFDFFFC8FCCCA0BEA9A8FC73" box="[818,941,975,998]" class="Insecta" family="Micropterigidae" genus="Tasmantrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FCCCA0BEA9A8FC73" box="[818,941,975,998]" italics="true" pageId="4" pageNumber="5">Tasmantrix</emphasis>
</taxonomicName>
species). The cluster of hair-sensilla on the membranous IX region may be a generic groundplan autapomorphy, and its absence in
<taxonomicName id="71C44D69FFDFFFC8FB9FA080AEE5FB9C" box="[1121,1248,1009,1033]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDFFFC8FB9FA080AEE5FB9C" box="[1121,1248,1009,1033]" italics="true" pageId="4" pageNumber="5">A. capensis</emphasis>
</taxonomicName>
and
<taxonomicName id="71C44D69FFDFFFC8FAEBA080AF99FB9C" box="[1301,1436,1009,1033]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sagittella">
<emphasis id="84B0EAF8FFDFFFC8FAEBA080AF99FB9C" box="[1301,1436,1009,1033]" italics="true" pageId="4" pageNumber="5">A. sagittella</emphasis>
</taxonomicName>
hence secondary; the dorsum IX location is suggestive of homology with the dorsum IX sensilla in
<taxonomicName id="71C44D69FFDFFFC8FB03A767AF99FBB8" box="[1277,1436,1046,1069]" class="Insecta" family="Micropterigidae" genus="Hypomartyria" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FB03A767AF99FBB8" box="[1277,1436,1046,1069]" italics="true" pageId="4" pageNumber="5">Hypomartyria</emphasis>
</taxonomicName>
which, however, are more close-set, have more prominent marginal rims, and are devoid of the central hair-like process. Larvae with D2 setae of trunk segments A1A8 minute and spherical, as in
<taxonomicName id="71C44D69FFDFFFC8FBB4A72CAEF0FBE0" box="[1098,1269,1117,1141]" class="Insecta" family="Micropterigidae" genus="Austromartyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FBB4A72CAEF0FBE0" box="[1098,1269,1117,1141]" italics="true" pageId="4" pageNumber="5">Austromartyria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B67B36EAFFDFFFC8FF39A7F1AB86F96D" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="5">
<emphasis id="84B0EAF8FFDFFFC8FF39A7F1A8BDFB0C" bold="true" box="[199,696,1152,1177]" pageId="4" pageNumber="5">Affinities and intrageneric relationships.</emphasis>
Findings emerging from preliminary analyses of 16S and 28S rRNA (Gibbs
<emphasis id="84B0EAF8FFDFFFC8FECBA7D4AB62FB29" box="[309,359,1188,1212]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2004, Gibbs 2010) place
<taxonomicName id="71C44D69FFDFFFC8FD70A7D5A923FB29" box="[654,806,1188,1212]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FD70A7D5A923FB29" box="[654,806,1188,1212]" italics="true" pageId="4" pageNumber="5">Agrionympha</emphasis>
</taxonomicName>
in a (weakly supported) clade, referred to loosely as the southern sabatincoid lineage, which also includes undescribed genera from
<collectingCountry id="CED3767AFFDFFFC8FC00A7B9AE97FB75" box="[1022,1170,1224,1248]" name="South Africa" pageId="4" pageNumber="5">South Africa</collectingCountry>
,
<collectingCountry id="CED3767AFFDFFFC8FB63A7B9AF21FB75" box="[1181,1316,1224,1248]" name="Madagascar" pageId="4" pageNumber="5">Madagascar</collectingCountry>
and
<collectingCountry id="CED3767AFFDFFFC8FAA2A7B9AACFFA96" name="Costa Rica" pageId="4" pageNumber="5">Costa Rica</collectingCountry>
as well as
<taxonomicName id="71C44D69FFDFFFC8FEBAA79DABEAFA91" box="[324,495,1260,1284]" class="Insecta" family="Micropterigidae" genus="Austromartyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FEBAA79DABEAFA91" box="[324,495,1260,1284]" italics="true" pageId="4" pageNumber="5">Austromartyria</emphasis>
</taxonomicName>
(N. Qld,
<collectingCountry id="CED3767AFFDFFFC8FDA7A79AA8C7FA96" box="[601,706,1259,1283]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
) and
<taxonomicName id="71C44D69FFDFFFC8FD00A79CA9F3FA96" authority="(Chile)" baseAuthorityName="Chile" box="[766,1014,1259,1284]" class="Insecta" family="Micropterigidae" genus="Hypomartyria" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FD00A79CA998FA91" box="[766,925,1261,1284]" italics="true" pageId="4" pageNumber="5">Hypomartyria</emphasis>
(
<collectingCountry id="CED3767AFFDFFFC8FC52A79AA9EAFA96" box="[940,1007,1259,1283]" name="Chile" pageId="4" pageNumber="5">Chile</collectingCountry>
)
</taxonomicName>
. The structure of the sternum V gland protuberance with its marginal piliform scales, the presence of both dorsal and ventral valve muscles from the segment IX ring, and the morphology (including chaetotaxy) of the early-instar larva is compatible with this placement. So too is the structure of the spermatheca which closely resembles
<taxonomicName id="71C44D69FFDFFFC8FBF9A626AEB6FAFA" box="[1031,1203,1367,1391]" class="Insecta" family="Micropterigidae" genus="Austromartyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FBF9A626AEB6FAFA" box="[1031,1203,1367,1391]" italics="true" pageId="4" pageNumber="5">Austromartyria</emphasis>
</taxonomicName>
from
<collectingCountry id="CED3767AFFDFFFC8FAFFA626AF6DFAFA" box="[1281,1384,1367,1391]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
and
<taxonomicName id="71C44D69FFDFFFC8FF69A60AAB31FA07" box="[151,308,1403,1426]" class="Insecta" family="Micropterigidae" genus="Squamicornia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDFFFC8FF69A60AAB31FA07" box="[151,308,1403,1426]" italics="true" pageId="4" pageNumber="5">Squamicornia</emphasis>
</taxonomicName>
from
<collectingCountry id="CED3767AFFDFFFC8FE82A60BABD9FA07" box="[380,476,1402,1426]" name="Ecuador" pageId="4" pageNumber="5">Ecuador</collectingCountry>
. However, in all these cases the resemblances are due to traits which occur in other micropterigids as well and which may be plesiomorphic within the family. Comprehensive sets of morphological and molecular data on all Micropterigid genera are currently being developed, and combined analyses of these will likely provide a more robust generic phylogeny in due course. A preliminary morphological analysis of the known larvae of
<taxonomicName id="71C44D69FFDFFFC8FEF4A578ABBAF9B4" box="[266,447,1545,1569]" class="Insecta" family="Micropterigidae" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Micropterigidae</taxonomicName>
(GWG unpublished) emphasised the clear distinction between two morphotypes: 1)
<emphasis id="84B0EAF8FFDFFFC8FF5EA55FAB2BF9D0" box="[160,302,1582,1605]" italics="true" pageId="4" pageNumber="5">Micropterix-</emphasis>
type known from genera in Europe,
<collectingCountry id="CED3767AFFDFFFC8FD2DA55CA93FF9D0" box="[723,826,1581,1605]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
and
<collectingCountry id="CED3767AFFDFFFC8FC8CA55FAE0CF9D0" box="[882,1033,1581,1605]" name="New Zealand" pageId="4" pageNumber="5">New Zealand</collectingCountry>
(see Gibbs 2010 for further detail), and 2) the sabatincoid-
<typeStatus id="697F8848FFDFFFC8FE55A520ABDAF9FC" box="[427,479,1617,1641]" pageId="4" pageNumber="5">type</typeStatus>
. Within the latter group larval characters prove useful for distinguishing between the northern hemisphere members (e.g.
<emphasis id="84B0EAF8FFDFFFC8FDC6A507AF96F918" box="[568,1427,1653,1677]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="71C44D69FFDFFFC8FDC6A507A981F918" authority="Neomicropterix" authorityName="Neomicropterix" box="[568,900,1654,1677]" class="Insecta" family="Micropterigidae" genus="Epimartyria" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Epimartyria, Neomicropterix</taxonomicName>
,
<taxonomicName id="71C44D69FFDFFFC8FC6CA507AF96F918" authority="Issikiomartyria, Kurokopteryx" authorityName="Issikiomartyria, Kurokopteryx" box="[914,1427,1653,1677]" class="Insecta" family="Micropterigidae" genus="Paramartyria" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Paramartyria, Issikiomartyria, Kurokopteryx</taxonomicName>
</emphasis>
) found in North
<collectingCountry id="CED3767AFFDFFFC8FEAEA5E9ABB6F925" box="[336,435,1688,1712]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
,
<collectingCountry id="CED3767AFFDFFFC8FE3DA5E8A803F925" box="[451,518,1689,1712]" name="Japan" pageId="4" pageNumber="5">Japan</collectingCountry>
,
<collectingCountry id="CED3767AFFDFFFC8FDEBA5E9A86FF925" box="[533,618,1688,1712]" name="Taiwan" pageId="4" pageNumber="5">Taiwan</collectingCountry>
,
<collectingCountry id="CED3767AFFDFFFC8FD87A5E9A8DEF925" box="[633,731,1688,1712]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
, and the remaining genera which occur around the southern hemisphere (
<emphasis id="84B0EAF8FFDFFFC8FED3A5CDA9DFF941" box="[301,986,1724,1748]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="71C44D69FFDFFFC8FED3A5CDABCDF941" box="[301,456,1724,1748]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Agrionympha</taxonomicName>
,
<taxonomicName id="71C44D69FFDFFFC8FE26A5CDA883F941" box="[472,646,1724,1748]" class="Insecta" family="Micropterigidae" genus="Austromartyria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Austromartyria</taxonomicName>
,
<taxonomicName id="71C44D69FFDFFFC8FD69A5CDA90FF941" box="[663,778,1724,1748]" class="Insecta" family="Micropterigidae" genus="Sabatinca" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Sabatinca</taxonomicName>
,
<taxonomicName id="71C44D69FFDFFFC8FCE7A5CCA9BDF941" box="[793,952,1725,1748]" class="Insecta" family="Micropterigidae" genus="Hypomartyria" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Hypomartyria</taxonomicName>
in
</emphasis>
Africa,
<collectingCountry id="CED3767AFFDFFFC8FBBEA5CDAEAFF941" box="[1088,1194,1724,1748]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
,
<collectingCountry id="CED3767AFFDFFFC8FB44A5CCAF52F941" box="[1210,1367,1724,1748]" name="New Zealand" pageId="4" pageNumber="5">New Zealand</collectingCountry>
,
<collectingCountry id="CED3767AFFDFFFC8FA99A5CCAB0CF96D" name="New Caledonia" pageId="4" pageNumber="5">New Caledonia</collectingCountry>
and
<collectingCountry id="CED3767AFFDFFFC8FEC1A591AB85F96D" box="[319,384,1760,1784]" name="Chile" pageId="4" pageNumber="5">Chile</collectingCountry>
.
</paragraph>
<paragraph id="B67B36EAFFDFFFC8FF39A475A8A0F866" blockId="4.[151,1437,151,2035]" pageId="4" pageNumber="5">
We consider the morphological characters so far studied and found useful for species discrimination to be an insufficient basis for making suggestions about intrageneric phylogeny. Suffice it here to note that
<taxonomicName id="71C44D69FFDFFFC8FB12A456AF68F8AA" box="[1260,1389,1831,1855]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDFFFC8FB12A456AF68F8AA" box="[1260,1389,1831,1855]" italics="true" pageId="4" pageNumber="5">A. capensis</emphasis>
</taxonomicName>
and
<taxonomicName id="71C44D69FFDFFFC8FF69A43AAB25F8F6" box="[151,288,1867,1891]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sagittella">
<emphasis id="84B0EAF8FFDFFFC8FF69A43AAB25F8F6" box="[151,288,1867,1891]" italics="true" pageId="4" pageNumber="5">A. sagittella</emphasis>
</taxonomicName>
both in their way stand particularly out from the rest: both lack the dorsum IX sensillum clusters in males, the former is moreover distinctive by the extreme wing shape and the Rs3&amp;Rs4 stalk (from which, probably at least consistently in the male, M1 issues as well) and the latter by the presence of dorsum VIII sensilla. We are unable to make well-substantiated decisions about character polarity in these cases, and both species may well prove to be quite subordinate within the genus.
</paragraph>
<paragraph id="B67B36EAFFDEFFC9FF39A3E6A964FF62" blockId="5.[151,1436,151,860]" pageId="5" pageNumber="6">
Species discrimination in
<taxonomicName id="71C44D69FFDEFFC9FE0DA3E9A88EFF25" box="[499,651,152,176]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDEFFC9FE0DA3E9A88EFF25" box="[499,651,152,176]" italics="true" pageId="5" pageNumber="6">Agrionympha</emphasis>
</taxonomicName>
is best achieved from the forewing maculation and confirmed with male genitalic details. Female morphological details contribute little to taxonomic precision, as in most micropterigid genera, but are presented here for comparative purposes.
</paragraph>
<paragraph id="B67B36EAFFDEFFC9FF39A272A9D4FDA9" blockId="5.[151,1436,151,860]" pageId="5" pageNumber="6">
<emphasis id="84B0EAF8FFDEFFC9FF39A272AB4FFE89" bold="true" box="[199,330,259,284]" pageId="5" pageNumber="6">Bionomics.</emphasis>
<taxonomicName id="71C44D69FFDEFFC9FEADA275ABEEFE89" box="[339,491,260,284]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDEFFC9FEADA275ABEEFE89" box="[339,491,260,284]" italics="true" pageId="5" pageNumber="6">Agrionympha</emphasis>
</taxonomicName>
species occur in moist forest or fynbos shrubland from sea level to
<quantity id="713C9B0FFFDEFFC9FB0BA275AF4BFE89" box="[1269,1358,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="6" unit="m" value="1700.0">1700 m</quantity>
, wherever there is sufficient seepage and a shade canopy to promote the growth of liverworts at some time of the year and provide the sheltered humidity necessary for adult moths. The day-flying adults are generally found amongst low vegetation (usually ferns) on cool roadside banks or alongside streams. In the hot arid climate of parts of southern Africa, these flight conditions are only fulfilled for a very limited part of the day, leading to an interesting variation of diurnal activity times in which these moisture-sensitive moths appear to be active only for an hour or so after dawn, prior to the evaporation of overnight dew. This behaviour has been observed at Toorberg in the
<taxonomicName id="71C44D69FFDEFFC9FAA8A2ACAF99FE61" box="[1366,1436,477,500]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">Karoo</taxonomicName>
and similarly at Storms River, Eastern Cape, where
<taxonomicName id="71C44D69FFDEFFC9FD26A171A952FD8D" box="[728,855,512,536]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDEFFC9FD26A171A952FD8D" box="[728,855,512,536]" italics="true" pageId="5" pageNumber="6">A. capensis</emphasis>
</taxonomicName>
can be taken at night (females are attracted to light) or amongst the low fynbos shrubland before the heat of the day builds up.
</paragraph>
<paragraph id="B67B36EAFFDEFFC9FF39A136A869FD11" blockId="5.[151,1436,151,860]" pageId="5" pageNumber="6">Adults have been taken between mid-October and March but each species appears to have a relatively restricted flight period at any one locality.</paragraph>
<paragraph id="B67B36EAFFDEFFC9FF39A1FEAE2CFD32" blockId="5.[151,1436,151,860]" box="[199,1065,655,679]" pageId="5" pageNumber="6">The known larvae, putatively representing two species, are liverwort-feeders.</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDEFFC9FF39A1C2AE1BFCC9" pageId="5" pageNumber="6" type="distribution">
<paragraph id="B67B36EAFFDEFFC9FF39A1C2AEDDFC81" blockId="5.[151,1436,151,860]" pageId="5" pageNumber="6">
<emphasis id="84B0EAF8FFDEFFC9FF39A1C2AB5CFD59" bold="true" box="[199,345,691,716]" pageId="5" pageNumber="6">Distribution</emphasis>
(
<figureCitation id="2EFF2A6FFFDEFFC9FE96A1C5ABBCFD59" box="[360,441,692,716]" captionStart="FIGURE 11" captionStartId="19.[151,250,924,947]" captionTargetBox="[218,1371,201,865]" captionTargetId="figure@19.[208,1379,193,921]" captionTargetPageId="19" captionText="FIGURE 11. Distribution of known Agrionympha species in southern Africa: open circles, A. capensis Whalley; open star, A. fuscoapicella sp. nov; open square, A kroonella sp. nov. and A. vari Whalley; solid 5 - point star, A. sagittella sp. nov.; multipoint star, A. karoo sp. nov.; solid square, A. jansella sp. nov. &amp; A. pseliacma Janse; solid triangle, A. pseudovari sp. nov.; solid circle, male species 1 , Diepwalle." httpUri="https://zenodo.org/record/203564/files/figure.png" pageId="5" pageNumber="6">Fig. 11</figureCitation>
). The presently-known species are restricted to the eastern escarpment ranges and coastal fringe of
<collectingCountry id="CED3767AFFDEFFC9FEFEA1A6A81EFD7A" box="[256,539,727,751]" name="South Africa" pageId="5" pageNumber="6">Republic of South Africa</collectingCountry>
from
<geoCoordinate id="D3F0502DFFDEFFC9FD9FA1A6A8C7FD7A" box="[609,706,727,751]" direction="south" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="-24.566668">24º 34´S</geoCoordinate>
in Mpumalanga (formerly Transvaal) to
<geoCoordinate id="D3F0502DFFDEFFC9FB6EA1A6AEF6FD7A" box="[1168,1267,727,751]" direction="south" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="-34.266666">34º 16´S</geoCoordinate>
, Western Cape at Cape Town, with outliers in the
<taxonomicName id="71C44D69FFDEFFC9FDE7A18CA85AFC81" box="[537,607,765,788]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">Karoo</taxonomicName>
near Graaff Reinet (
<geoCoordinate id="D3F0502DFFDEFFC9FCBBA18DA9A3FC81" box="[837,934,764,788]" direction="south" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="-32.183334">32º 11´S</geoCoordinate>
) and Worcester (
<geoCoordinate id="D3F0502DFFDEFFC9FB98A18DAECEFC81" box="[1126,1227,764,788]" direction="south" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="-33.633335">33º 38´S</geoCoordinate>
).
</paragraph>
<paragraph id="B67B36EAFFDEFFC9FF39A06EAE1BFCC9" blockId="5.[151,1436,151,860]" pageId="5" pageNumber="6">
Associated taxa of plants and animals indicate that
<taxonomicName id="71C44D69FFDEFFC9FCE2A051A9B1FCAD" box="[796,948,800,824]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDEFFC9FCE2A051A9B1FCAD" box="[796,948,800,824]" italics="true" pageId="5" pageNumber="6">Agrionympha</emphasis>
</taxonomicName>
is a component of the well-known “Cape Biota”. It is not found in the lowland Afrotropical rainforests of southern Africa.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,223 @@
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<treatment id="3E6D87FCFFDDFFCBFF69A72FA9E4FC70" ID-DOI="http://doi.org/10.5281/zenodo.6189653" ID-GBIF-Taxon="119388869" ID-Zenodo-Dep="6189653" LSID="urn:lsid:plazi:treatment:3E6D87FCFFDDFFCBFF69A72FA9E4FC70" httpUri="http://treatment.plazi.org/id/3E6D87FCFFDDFFCBFF69A72FA9E4FC70" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="FEDE6561FFDDFFCAFF69A72FA84BFB0C" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="B67B36EAFFDDFFCAFF69A72FA867FBED" blockId="6.[151,610,1118,1210]" box="[151,610,1118,1144]" pageId="6" pageNumber="7">
<heading id="ED338186FFDDFFCAFF69A72FA867FBED" bold="true" box="[151,610,1118,1144]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="71C44D69FFDDFFCAFF69A72FA867FBED" ID-CoL="B7SG" authority="Whalley, 1978" authorityName="Whalley" authorityYear="1978" box="[151,610,1118,1144]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDDFFCAFF69A72FA867FBED" bold="true" box="[151,610,1118,1144]" pageId="6" pageNumber="7">
<emphasis id="84B0EAF8FFDDFFCAFF69A72FABAEFBED" bold="true" box="[151,427,1118,1144]" italics="true" pageId="6" pageNumber="7">Agrionympha capensis</emphasis>
<bibRefCitation id="D2554B1BFFDDFFCAFE4CA72FA867FBED" author="Whalley" box="[434,610,1118,1144]" pageId="6" pageNumber="21" refString="Whalley, P. E. S. (1978) New taxa of fossil and recent Micropterigidae with a discussion of their evolution and a comment on the evolution of Lepidoptera (Insecta). Annals of the Transvaal Museum, 31, 71 - 86." type="journal article" year="1978">Whalley, 1978</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="B67B36EAFFDDFFCAFF69A7F0A84BFB0C" blockId="6.[151,610,1118,1210]" box="[151,590,1153,1177]" pageId="6" pageNumber="7">
<emphasis id="84B0EAF8FFDDFFCAFF69A7F0AB67FB0C" box="[151,354,1153,1177]" italics="true" pageId="6" pageNumber="7">Ann. Transv. Mus.</emphasis>
31: 76, Pl 14,
<figureCitation id="2EFF2A6FFFDDFFCAFDFBA7F0A84CFB0C" box="[517,585,1153,1177]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDDFFCAFF69A7D3A864FB2F" box="[151,609,1186,1210]" pageId="6" pageNumber="7" type="description">
<paragraph id="B67B36EAFFDDFFCAFF69A7D3A864FB2F" blockId="6.[151,610,1118,1210]" box="[151,609,1186,1210]" pageId="6" pageNumber="7">
(
<figureCitation id="2EFF2A6FFFDDFFCAFF61A7D3AAEDFB2F" box="[159,232,1186,1210]" captionStart="FIGURE 1" captionStartId="7.[151,250,1973,1996]" captionTargetBox="[206,1390,1043,1943]" captionTargetId="figure@7.[196,1401,1035,1968]" captionTargetPageId="7" captionText="FIGURE 1. Adult moths from life: A, Agrionympha capensis Whalley, Storms River; B, A. kroonella, sp. nov. Mariepskop; C, A. fuscoapicella, sp. nov. Hogsback; D, A. sagittella, sp. nov. Hogsback. Scale bar = 1 mm." httpUri="https://zenodo.org/record/203554/files/figure.png" pageId="6" pageNumber="7">Figs. 1</figureCitation>
A, 2A, 3B, F, G), 4A, B, 5, 7, 11)
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDDFFCAFF69A794A96BF913" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="B67B36EAFFDDFFCAFF69A794A96BF913" blockId="6.[151,1436,1253,2028]" pageId="6" pageNumber="7">
<emphasis id="84B0EAF8FFDDFFCAFF69A794AB84FB6B" bold="true" box="[151,385,1253,1278]" pageId="6" pageNumber="7">Material examined.</emphasis>
<emphasis id="84B0EAF8FFDDFFCAFE76A794ABEAFB68" box="[392,495,1253,1277]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="697F8848FFDDFFCAFE76A794ABEAFB68" box="[392,495,1253,1277]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
Ƥ,
<collectingCountry id="CED3767AFFDDFFCAFDE4A794A8A8FB68" box="[538,685,1253,1277]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
, Western Cape, Knysna, Garden of Eden.
<date id="C27A102AFFDDFFCAFB77A794AF3AFB68" box="[1161,1343,1253,1277]" pageId="6" pageNumber="7" value="1955-01-16" valueMax="1955-01-20" valueMin="1955-01-16">1620 Jan 1955</date>
, L.
<taxonomicName id="71C44D69FFDDFFCAFA94A794AF99FB68" box="[1386,1436,1253,1277]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="20" phylum="Arthropoda" rank="species" species="vari">Vári</taxonomicName>
(
<collectionCode id="D0D5AE2FFFDDFFCAFF61A678AADAFAB4" box="[159,223,1289,1313]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="6" pageNumber="7">TMP</collectionCode>
).
<emphasis id="84B0EAF8FFDDFFCAFF0EA67BAB67FAB4" box="[240,354,1290,1313]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="697F8848FFDDFFCAFF0EA67BAB67FAB4" box="[240,354,1290,1313]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
</emphasis>
: 3Ƥ, Jonkershoek (WC),
<date id="C27A102AFFDDFFCAFD7EA678A911FAB4" box="[640,788,1289,1313]" pageId="6" pageNumber="7" value="1977-11-19">19 Nov 1977</date>
, L. &amp; G.
<taxonomicName id="71C44D69FFDDFFCAFC86A678A9AFFAB4" box="[888,938,1289,1313]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="20" phylum="Arthropoda" rank="species" species="vari">Vári</taxonomicName>
(
<collectionCode id="D0D5AE2FFFDDFFCAFC46A678AE17FAB4" box="[952,1042,1289,1313]" pageId="6" pageNumber="7">BMNH</collectionCode>
); Hermanus (WC),
<date id="C27A102AFFDDFFCAFB12A678AF7CFAB4" box="[1260,1401,1289,1313]" pageId="6" pageNumber="7" value="1977-02-26">26 Feb 1977</date>
, L. &amp; G.
<taxonomicName id="71C44D69FFDDFFCAFF2CA65DAB01FAD1" box="[210,260,1324,1348]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="20" phylum="Arthropoda" rank="species" species="vari">Vári</taxonomicName>
(
<collectionCode id="D0D5AE2FFFDDFFCAFEEDA65DAB56FAD1" box="[275,339,1324,1348]" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="6" pageNumber="7">TMP</collectionCode>
). O
<emphasis id="84B0EAF8FFDDFFCAFE87A65CA809FAD0" box="[377,524,1325,1349]" italics="true" pageId="6" pageNumber="7">ther material</emphasis>
:
<collectingCountry id="CED3767AFFDDFFCAFDE4A65DA8A8FAD1" box="[538,685,1324,1348]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
, Western Cape: 13, 7Ƥ Kogel Bay,
<geoCoordinate id="D3F0502DFFDDFFCAFBBEA65DAE9FFAD1" box="[1088,1178,1324,1348]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-34.266666">34º16´S</geoCoordinate>
<geoCoordinate id="D3F0502DFFDDFFCAFB5EA65DAEFBFAD1" box="[1184,1278,1324,1348]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="18.85">18º51´E</geoCoordinate>
,
<quantity id="713C9B0FFFDDFFCAFAF7A65DAF41FAD1" box="[1289,1348,1324,1348]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="20.0">20 m</quantity>
,
<date id="C27A102AFFDDFFCAFAB1A65DAAD7FAFD" pageId="6" pageNumber="7" value="1986-02-19">19 Feb 1986</date>
, G.W. Gibbs; 133 3Ƥ, Knysna, Goudveld Forest, Jubilee Creek,
<quantity id="713C9B0FFFDDFFCAFC3EA621AE0EFAFD" box="[960,1035,1360,1384]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="m" value="200.0">200 m</quantity>
,
<date id="C27A102AFFDDFFCAFBE7A621AED6FAFD" box="[1049,1235,1360,1384]" pageId="6" pageNumber="7" value="2004-01-27" valueMax="2004-01-31" valueMin="2004-01-27">2731 Jan 2004</date>
; 13 4Ƥ, Knysna, Diepwalle Forest, Spitskop,
<quantity id="713C9B0FFFDDFFCAFE29A605A85CFA19" box="[471,601,1396,1420]" metricMagnitude="2" metricUnit="m" metricValue="7.8" metricValueMax="8.0" metricValueMin="7.6" pageId="6" pageNumber="7" unit="m" value="780.0" valueMax="800.0" valueMin="760.0">760800 m</quantity>
,
<date id="C27A102AFFDDFFCAFD98A605A8F7FA19" box="[614,754,1396,1420]" pageId="6" pageNumber="7" value="2004-01-31">31 Jan 2004</date>
; 1Ƥ, 3? (abdomen missing), Du Toits Kloof,
<date id="C27A102AFFDDFFCAFB0CA605AF70FA19" box="[1266,1397,1396,1420]" pageId="6" pageNumber="7" value="1989-02-06">6 Feb 1989</date>
, H. Geertsema. Eastern Cape: 33 7Ƥ, Storms River NP,
<geoCoordinate id="D3F0502DFFDDFFCAFD0BA6E6A94AFA3A" box="[757,847,1431,1455]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-33.966667">33º58´S</geoCoordinate>
<geoCoordinate id="D3F0502DFFDDFFCAFCA6A6E6A9B3FA3A" box="[856,950,1431,1455]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="23.933332">23º56´E</geoCoordinate>
,
<quantity id="713C9B0FFFDDFFCAFC3DA6E6AE0BFA3A" box="[963,1038,1431,1455]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="7" unit="m" value="240.0">240 m</quantity>
,
<date id="C27A102AFFDDFFCAFBE5A6E6AEDDFA3A" box="[1051,1240,1431,1455]" pageId="6" pageNumber="7" value="1986-02-14" valueMax="1986-02-15" valueMin="1986-02-14">1415 Feb 1986</date>
, G.W. Gibbs; 7Ƥ same locality,
<quantity id="713C9B0FFFDDFFCAFEC9A6CAAB7AFA46" box="[311,383,1467,1491]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="7" unit="m" value="240.0">240 m</quantity>
, at UV light,
<date id="C27A102AFFDDFFCAFDE8A6CAA8CBFA46" box="[534,718,1467,1491]" pageId="6" pageNumber="7" value="1986-02-14" valueMax="1986-02-15" valueMin="1986-02-14">1415 Feb 1986</date>
, D.M. Kroon; 13 1Ƥ, Bloukrans River, Bloukrans Pass road,
<quantity id="713C9B0FFFDDFFCAFA7EA6CAAAAAFA62" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="7" unit="m" value="32.0">32 m</quantity>
,
<date id="C27A102AFFDDFFCAFF42A6AEAB4BFA62" box="[188,334,1503,1527]" pageId="6" pageNumber="7" value="1986-02-15">15 Feb 1986</date>
, G.W. Gibbs; 2Ƥ, Natures Valley camp,
<geoCoordinate id="D3F0502DFFDDFFCAFCDDA6AEA979FA62" box="[803,892,1503,1527]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-33.966667">33º58´S</geoCoordinate>
<geoCoordinate id="D3F0502DFFDDFFCAFC7BA6AEA9E6FA62" box="[901,995,1503,1527]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="23.566668">23º34´E</geoCoordinate>
,
<quantity id="713C9B0FFFDDFFCAFC0EA6AEAE1BFA62" box="[1008,1054,1503,1527]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="m" value="4.0">4 m</quantity>
, at UV light,
<date id="C27A102AFFDDFFCAFB42A6AEAF51FA62" box="[1212,1364,1503,1527]" pageId="6" pageNumber="7" value="1986-11-15">15 Nov 1986</date>
, D.M. Kroon; 13 1Ƥ, Umtata, Langeni Forest,
<quantity id="713C9B0FFFDDFFCAFDA1A572A8B3F98E" box="[607,694,1539,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="m" value="1500.0">1500 m</quantity>
,
<date id="C27A102AFFDDFFCAFD3FA572A977F98E" box="[705,882,1539,1563]" pageId="6" pageNumber="7" value="1986-11-02" valueMax="1986-11-16" valueMin="1986-11-02">216 Nov 1986</date>
, N. Duke. Male genital preps. G.W. Gibbs G1002 Storms R.), G1006 (Storms R.). Female genital preps. G.W. Gibbs G1009 (Bloukrans Pass), G1010 (Jubilee Ck), G1011 (Storms R.). Wing slide G1025 (Jubilee Ck). One of Gibbs (1999) specimens from Storms R., an entire female, extracted for
<collectionCode id="D0D5AE2FFFDDFFCAFE78A51EABC6F912" LSID="urn:lsid:biocol.org:col:15536" box="[390,451,1647,1671]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="6" pageNumber="7">DNA</collectionCode>
by Kobayashi (AG2; r63; see below).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDDFFCBFF3BA5E0A87AFD52" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="B67B36EAFFDDFFCAFF3BA5E0AF57F964" blockId="6.[151,1436,1253,2028]" pageId="6" pageNumber="7">
<emphasis id="84B0EAF8FFDDFFCAFF3BA5E0AB53F93F" bold="true" box="[197,342,1681,1706]" pageId="6" pageNumber="7">Description.</emphasis>
<emphasis id="84B0EAF8FFDDFFCAFE9CA5E3AB9AF93F" box="[354,415,1682,1706]" italics="true" pageId="6" pageNumber="7">Head</emphasis>
. Antennae gradually darkening from yellowish-brown to dark grey over 17 basalmost flagellomeres in male,
<quantity id="713C9B0FFFDDFFCAFE67A5C7ABD6F95B" box="[409,467,1718,1742]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="6" pageNumber="7" unit="in" value="12.0">12 in</quantity>
female; total of 4042 flagellomeres in male with basal 3 scale-covered,
<quantity id="713C9B0FFFDDFFCAFAF1A5C7AF4CF95B" box="[1295,1353,1718,1742]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="6" pageNumber="7" unit="in" value="26.0">26 in</quantity>
female with basal 3 scaled. Piliform scales on scape, pedicel and head yellow-brown, same colour as flagellomeres.
</paragraph>
<paragraph id="B67B36EAFFDDFFCBFF39A58CAF79FF62" blockId="6.[151,1436,1253,2028]" lastBlockId="7.[151,1436,151,997]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="84B0EAF8FFDDFFCAFF39A58CAB1DF880" box="[199,280,1789,1813]" italics="true" pageId="6" pageNumber="7">Thorax</emphasis>
. Tegulae creamy-white scaled; mesonotum black-scaled. Legs with forecoxae black, mid- and hindcoxae ochreous; femora all with glistening white scales; tibiae black; tarsi black but with whitish bands at proximal end of all tarsomeres except hindtarsomere 1. FW length male
<quantity id="713C9B0FFFDDFFCAFCA5A435A9B4F8C9" box="[859,945,1860,1884]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="7" unit="mm" value="2.8">2.8 mm</quantity>
(2.63.0), female
<quantity id="713C9B0FFFDDFFCAFB7FA435AEDDF8C9" box="[1153,1240,1860,1884]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="7" unit="mm" value="3.1">3.1 mm</quantity>
(2.93.4). Wings markedly elongate (FW ratio length:width 3.6, HW 4.64.9) with apex attenuated, wing shape most extreme in male (
<figureCitation id="2EFF2A6FFFDDFFCAFF20A4FDAB24F831" box="[222,289,1932,1956]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
B). FW and HW with Rs3 and Rs4 stalked; in some specimens (all four males of which slide mounts were examined, and one of four females) M1 anastomosing with stem, and hence eventually issuing from Rs3+4 (
<figureCitation id="2EFF2A6FFFDDFFCAFF61A4A2AAE6F87E" box="[159,227,2003,2027]" captionStart="FIGURE 3" captionStartId="8.[151,250,1803,1826]" captionTargetBox="[245,1340,882,1771]" captionTargetId="figure@8.[241,1346,878,1782]" captionTargetPageId="8" captionText="FIGURE 3. Wing venation and head appendages, Agrionympha species: A, A. sagittella male, Hogsback; B, A. capensis male, Jubilee Ck; C, A. sagittella, hindwing front margin with apparent cross-vein between Sc 1 and Sc 2, R spur identifiable only as a minute process on Sc 2; D, A. jansella, hindwing front margin with cross vein between Sc 1 and Sc 2, R spur forming a sizable, proximally-directed process on Sc 2; E, A. sagittella, hindwing front margin with Sc 1 and Sc 2 separate for an unusually long distance, but unconnected by apparent cross-vein, no spur identifiable; F, A. capensis female, scape and pedicel of antenna; G, A. capensis female maxillary palp. Scale line for F and G = 0.5 mm." httpUri="https://zenodo.org/record/203556/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
B). Forewing maculation (
<figureCitation id="2EFF2A6FFFDDFFCAFDF0A4A2A854F87E" box="[526,593,2003,2027]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
A) distinctive, with very broad silvery-white bands; a prominent L-shaped figure resulting from confluence of claval mark with costal part of median band; median band broken in middle with posterior part distally displaced on dorsum; postmedian band very broad in anterior half, interrupted by line of dark scales across M2, not reaching to dorsum; apical band subterminal, also interrupted by lines of dark scales.
</paragraph>
<paragraph id="B67B36EAFFDCFFCBFF39A272ABC3FE5B" blockId="7.[151,1436,151,997]" pageId="7" pageNumber="8">
<emphasis id="84B0EAF8FFDCFFCBFF39A272AB9AFE8E" box="[199,415,259,283]" italics="true" pageId="7" pageNumber="8">Male postabdomen</emphasis>
(
<figureCitation id="2EFF2A6FFFDCFFCBFE4FA272A804FE8E" box="[433,513,259,283]" captionStart="FIGURE 7" captionStartId="14.[151,250,1457,1480]" captionTargetBox="[160,1416,529,1421]" captionTargetId="figure@14.[160,1427,519,1454]" captionTargetPageId="14" captionText="FIGURE 7. Agrionympha capensis Whalley, genitalia: A E, male, Storms River: A, ventral; B, lateral; C, dorsal; D, phallus lateral; E, phallus dorsal. F G, female, Bloukrans Pass: F, extended terminal segments of abdomen; G, internal female genitalia. cb, corpus bursae; dph, distal portion of phallus; dsp, ductus of spermatheca; go, gonopore with radial folds; lag, lagena; lap, lappet, mpl, median plate; pap, papilla; pph, proximal portion of phallus; scl ix, ninth sclerite; scl x, tenth sclerite; S viii, eighth sternite; spm, spermatheca; T viii, eighth tergite; T x, tenth tergum; utr, distal bulb of utriculus; va, valva; vb, ventral bulb." httpUri="https://zenodo.org/record/203560/files/figure.png" pageId="7" pageNumber="8">Figs. 7</figureCitation>
AE): Sternum VIII remnants irregularly triangular, bearing about 5 small setae. Anteroventral margin of IX sclerotisation broadly rounded, unthickened. Median plate narrow, keeled but unpigmented. Valvae of moderate length, curved upward (length:width ratio 4.0). Hair-sensilla absent. Phallus elongate and nearly straight, ratio length:height 5.6; aedeagus with very constricted neck region in dorsal view; gonopore egg-shaped, widest distally, less than half (0.37) width of aedeagus between the lappets; ventral bulb very reduced. (male genital prep. G1006)
</paragraph>
<paragraph id="B67B36EAFFDCFFCBFF2EA2A8A87AFD52" blockId="7.[151,1436,151,997]" pageId="7" pageNumber="8">
<emphasis id="84B0EAF8FFDCFFCBFF2EA2A8ABC7FE64" box="[208,450,473,497]" italics="true" pageId="7" pageNumber="8">Female postabdomen</emphasis>
(
<figureCitation id="2EFF2A6FFFDCFFCBFE2DA2A8A812FE64" box="[467,535,473,497]" captionStart="FIGURE 7" captionStartId="14.[151,250,1457,1480]" captionTargetBox="[160,1416,529,1421]" captionTargetId="figure@14.[160,1427,519,1454]" captionTargetPageId="14" captionText="FIGURE 7. Agrionympha capensis Whalley, genitalia: A E, male, Storms River: A, ventral; B, lateral; C, dorsal; D, phallus lateral; E, phallus dorsal. F G, female, Bloukrans Pass: F, extended terminal segments of abdomen; G, internal female genitalia. cb, corpus bursae; dph, distal portion of phallus; dsp, ductus of spermatheca; go, gonopore with radial folds; lag, lagena; lap, lappet, mpl, median plate; pap, papilla; pph, proximal portion of phallus; scl ix, ninth sclerite; scl x, tenth sclerite; S viii, eighth sternite; spm, spermatheca; T viii, eighth tergite; T x, tenth tergum; utr, distal bulb of utriculus; va, valva; vb, ventral bulb." httpUri="https://zenodo.org/record/203560/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
FG): Abdomen elongate; segment V gland
<quantity id="713C9B0FFFDCFFCBFBEAA2A8AE69FE64" box="[1044,1132,473,497]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="mm" value="0.4">0.4 mm</quantity>
diam., with about 10 long fluted scales; segment VIII unmodified. Segment X elongate, the sclerites longer than high (about 1.6x) with darkening often emarginated along anterior edge. Papilla small (
<quantity id="713C9B0FFFDCFFCBFCBFA150A9ADFDAC" box="[833,936,545,569]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="7" pageNumber="8" unit="mm" value="0.07">0.07 mm</quantity>
deep), approximately equidimensional with complex walls reflexed back from the anterior opening on either side. Thick-walled proximal section of spermatheca with its bulbous end about 0.8x diameter of ductus length and with a small lateral bulb adjacent to large bulb; thin-walled mid section of utriculus about half the length of the thick-walled ductus, anterior sac of utriculus
<quantity id="713C9B0FFFDCFFCBFA87A1FDAAC6FD52" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="mm" value="0.1">0.1 mm</quantity>
diameter (female genital prep. G1009).
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDCFFCBFF3BA1A2AE79FC0A" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="B67B36EAFFDCFFCBFF3BA1A2A9A9FC9A" blockId="7.[151,1436,151,997]" pageId="7" pageNumber="8">
<emphasis id="84B0EAF8FFDCFFCBFF3BA1A2AB3AFD79" bold="true" box="[197,319,723,748]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
The L-shaped patch on the FW base is distinctive. Structurally unique by extremely slender/ pointed wings with stalked Rs3/Rs4 and absence of male hair-sensilla.
</paragraph>
<paragraph id="B67B36EAFFDCFFCBFF3BA06BAE79FC0A" blockId="7.[151,1436,151,997]" pageId="7" pageNumber="8">
<emphasis id="84B0EAF8FFDCFFCBFF3BA06BAB44FCA6" bold="true" box="[197,321,794,819]" pageId="7" pageNumber="8">Bionomics</emphasis>
.Occurs close to damp seepage areas in fynbos, with ferns, grasses or shrubs exposed to full sunlight, or around forest margins. This species has occurred over a wide range of altitudes from
<quantity id="713C9B0FFFDCFFCBFB40A04FAEEFFCC3" box="[1214,1258,830,854]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="m" value="4.0">4 m</quantity>
to
<quantity id="713C9B0FFFDCFFCBFAF1A04FAF69FCC3" box="[1295,1388,830,854]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="8" unit="m" value="1500.0">1,500 m</quantity>
and seasonally between early November and early March, and is thus probably double-brooded. Females are attracted to MV light traps, often in considerable numbers, but no males have been taken this way.
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDCFFCBFF3BA0D8A9E4FC70" pageId="7" pageNumber="8" type="distribution">
<paragraph id="B67B36EAFFDCFFCBFF3BA0D8A9E4FC70" blockId="7.[151,1436,151,997]" pageId="7" pageNumber="8">
<emphasis id="84B0EAF8FFDCFFCBFF3BA0D8AB5BFC57" bold="true" box="[197,350,937,962]" pageId="7" pageNumber="8">Distribution.</emphasis>
<collectingCountry id="CED3767AFFDCFFCBFE9BA0DBABFCFC57" box="[357,505,938,962]" name="South Africa" pageId="7" pageNumber="8">South Africa</collectingCountry>
, Western and Eastern Cape districts; the most widespread species of
<taxonomicName id="71C44D69FFDCFFCBFB02A0DBAF91FC57" box="[1276,1428,938,962]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDCFFCBFB02A0DBAF91FC57" box="[1276,1428,938,962]" italics="true" pageId="7" pageNumber="8">Agrionympha</emphasis>
</taxonomicName>
, extending from
<geoCoordinate id="D3F0502DFFDCFFCBFEB7A0BCABA1FC70" box="[329,420,973,997]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="18.85">18º51´E</geoCoordinate>
near Cape Town to
<geoCoordinate id="D3F0502DFFDCFFCBFD78A0BCA8E4FC70" box="[646,737,973,997]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="28.85">28º51´E</geoCoordinate>
near Umtata (
<figureCitation id="2EFF2A6FFFDCFFCBFC7FA0BCA9D6FC70" box="[897,979,973,997]" captionStart="FIGURE 11" captionStartId="19.[151,250,924,947]" captionTargetBox="[218,1371,201,865]" captionTargetId="figure@19.[208,1379,193,921]" captionTargetPageId="19" captionText="FIGURE 11. Distribution of known Agrionympha species in southern Africa: open circles, A. capensis Whalley; open star, A. fuscoapicella sp. nov; open square, A kroonella sp. nov. and A. vari Whalley; solid 5 - point star, A. sagittella sp. nov.; multipoint star, A. karoo sp. nov.; solid square, A. jansella sp. nov. &amp; A. pseliacma Janse; solid triangle, A. pseudovari sp. nov.; solid circle, male species 1 , Diepwalle." httpUri="https://zenodo.org/record/203564/files/figure.png" pageId="7" pageNumber="8">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,367 @@
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<mods:title id="F0D59E2EA8B9F805C8EB78C335F5B5FC">Agrionympha, the long-known South African jaw moths: a revision with descriptions of new species (Lepidoptera, Micropterigidae)</mods:title>
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<treatment id="3E6D87FCFFDEFFCAFF69A0DEAF9EFB86" ID-DOI="http://doi.org/10.5281/zenodo.6189651" ID-GBIF-Taxon="119388866" ID-Zenodo-Dep="6189651" LSID="urn:lsid:plazi:treatment:3E6D87FCFFDEFFCAFF69A0DEAF9EFB86" httpUri="http://treatment.plazi.org/id/3E6D87FCFFDEFFCAFF69A0DEAF9EFB86" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="FEDE6561FFDEFFC9FF69A0DEA833FC5C" box="[151,566,943,969]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="B67B36EAFFDEFFC9FF69A0DEA833FC5C" blockId="5.[151,566,943,969]" box="[151,566,943,969]" pageId="5" pageNumber="6">
<heading id="ED338186FFDEFFC9FF69A0DEA833FC5C" bold="true" box="[151,566,943,969]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="84B0EAF8FFDEFFC9FF69A0DEA833FC5C" bold="true" box="[151,566,943,969]" pageId="5" pageNumber="6">
Checklist of
<taxonomicName id="71C44D69FFDEFFC9FECDA0DEABDCFC5C" ID-CoL="8ZZ76" box="[307,473,943,969]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDEFFC9FECDA0DEABDCFC5C" bold="true" box="[307,473,943,969]" italics="true" pageId="5" pageNumber="6">Agrionympha</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDEFFC9FF69A087AB8FFABB" pageId="5" pageNumber="6" type="description">
<paragraph id="B67B36EAFFDEFFC9FF69A087ABA2FBEE" blockId="5.[151,508,1014,1327]" pageId="5" pageNumber="6">
<emphasis id="84B0EAF8FFDEFFC9FF69A087AAA9FB9A" bold="true" box="[151,172,1014,1039]" pageId="5" pageNumber="6">1.</emphasis>
<taxonomicName id="71C44D69FFDEFFC9FF4DA087ABBAFB9B" authority="Whalley, 1978" authorityName="Whalley" authorityYear="1978" box="[179,447,1014,1039]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDEFFC9FF4DA087AB10FB9A" bold="true" box="[179,277,1014,1039]" italics="true" pageId="5" pageNumber="6">capensis</emphasis>
<bibRefCitation id="D2554B1BFFDEFFC9FEE2A087ABBAFB9B" author="Whalley" box="[284,447,1014,1038]" pageId="5" pageNumber="21" refString="Whalley, P. E. S. (1978) New taxa of fossil and recent Micropterigidae with a discussion of their evolution and a comment on the evolution of Lepidoptera (Insecta). Annals of the Transvaal Museum, 31, 71 - 86." type="journal article" year="1978">Whalley, 1978</bibRefCitation>
</taxonomicName>
<emphasis id="84B0EAF8FFDEFFC9FF69A76BAB4EFBA6" bold="true" box="[151,331,1050,1075]" pageId="5" pageNumber="6">
2.
<taxonomicName id="71C44D69FFDEFFC9FF4DA76BAB4EFBA6" authority="Gibbs" authorityName="Gibbs" box="[179,331,1050,1075]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="fuscoapicella" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FF4DA76BAB4EFBA6" bold="true" box="[179,331,1050,1075]" italics="true" pageId="5" pageNumber="6">fuscoapicella</emphasis>
</taxonomicName>
</emphasis>
Gibbs,
<emphasis id="84B0EAF8FFDEFFC9FE5DA76AAB08FBC2" bold="true" pageId="5" pageNumber="6">
<taxonomicNameLabel id="9F835783FFDEFFC9FE5DA76AABF9FBA6" box="[419,508,1051,1075]" pageId="5" pageNumber="6" rank="species">
sp.
<date id="C27A102AFFDEFFC9FE35A76AABF9FBA6" box="[459,508,1051,1075]" pageId="5" pageNumber="6">nov.</date>
</taxonomicNameLabel>
3.
<taxonomicName id="71C44D69FFDEFFC9FF4DA74FAB08FBC2" authority="Gibbs" authorityName="Gibbs" box="[179,269,1086,1111]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jansella" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FF4DA74FAB08FBC2" bold="true" box="[179,269,1086,1111]" italics="true" pageId="5" pageNumber="6">jansella</emphasis>
</taxonomicName>
</emphasis>
Gibbs,
<taxonomicNameLabel id="9F835783FFDEFFC9FE9BA74EABBBFBC2" box="[357,446,1087,1111]" pageId="5" pageNumber="6" rank="species">
<emphasis id="84B0EAF8FFDEFFC9FE9BA74EAB7AFBC2" bold="true" box="[357,383,1087,1111]" pageId="5" pageNumber="6">sp</emphasis>
.
<emphasis id="84B0EAF8FFDEFFC9FE73A74EABBBFBC2" bold="true" box="[397,446,1087,1111]" pageId="5" pageNumber="6">
<date id="C27A102AFFDEFFC9FE73A74EABBBFBC2" box="[397,446,1087,1111]" pageId="5" pageNumber="6">nov.</date>
</emphasis>
</taxonomicNameLabel>
4.
<taxonomicName id="71C44D69FFDEFFC9FF4DA713AB47FBEF" authority="Gibbs" authorityName="Gibbs" box="[179,322,1122,1147]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FF4DA713AAF3FBEE" bold="true" box="[179,246,1122,1147]" italics="true" pageId="5" pageNumber="6">karoo</emphasis>
Gibbs
</taxonomicName>
,
<taxonomicNameLabel id="9F835783FFDEFFC9FEB3A712ABA2FBEE" box="[333,423,1123,1147]" pageId="5" pageNumber="6" rank="species">
<emphasis id="84B0EAF8FFDEFFC9FEB3A712AB62FBEE" bold="true" box="[333,359,1123,1147]" pageId="5" pageNumber="6">sp</emphasis>
.
<emphasis id="84B0EAF8FFDEFFC9FE88A712ABA2FBEE" bold="true" box="[374,423,1123,1147]" pageId="5" pageNumber="6">nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="B67B36EAFFDEFFC9FF69A7F6AB8FFABB" blockId="5.[151,508,1014,1327]" pageId="5" pageNumber="6">
<emphasis id="84B0EAF8FFDEFFC9FF69A7F6AB24FB0A" bold="true" box="[151,289,1158,1183]" pageId="5" pageNumber="6">
5.
<taxonomicName id="71C44D69FFDEFFC9FF4DA7F7AB24FB0A" authority="Gibbs" authorityName="Gibbs" box="[179,289,1158,1183]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kroonella" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FF4DA7F7AB24FB0A" bold="true" box="[179,289,1158,1183]" italics="true" pageId="5" pageNumber="6">kroonella</emphasis>
</taxonomicName>
</emphasis>
Gibbs,
<taxonomicNameLabel id="9F835783FFDEFFC9FE87A7F6ABD6FB0A" box="[377,467,1159,1183]" pageId="5" pageNumber="6" rank="species">
<emphasis id="84B0EAF8FFDEFFC9FE87A7F6AB96FB0A" bold="true" box="[377,403,1159,1183]" pageId="5" pageNumber="6">sp</emphasis>
.
<emphasis id="84B0EAF8FFDEFFC9FE5CA7F6ABD6FB0A" bold="true" box="[418,467,1159,1183]" pageId="5" pageNumber="6">
<date id="C27A102AFFDEFFC9FE5CA7F6ABD6FB0A" box="[418,467,1159,1183]" pageId="5" pageNumber="6">nov.</date>
</emphasis>
</taxonomicNameLabel>
6.
<taxonomicName id="71C44D69FFDEFFC9FF4DA7DBABD7FB57" authority="Meyrick, 1921" authorityName="Meyrick" authorityYear="1921" box="[179,466,1194,1219]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFDEFFC9FF4DA7DBAB23FB56" bold="true" box="[179,294,1194,1219]" italics="true" pageId="5" pageNumber="6">pseliacma</emphasis>
Meyrick, 1921
</taxonomicName>
<emphasis id="84B0EAF8FFDEFFC9FF69A7BEAB34FB72" bold="true" box="[151,305,1230,1255]" pageId="5" pageNumber="6">
7.
<taxonomicName id="71C44D69FFDEFFC9FF4DA7BFAB34FB72" authority="Gibbs" authorityName="Gibbs" box="[179,305,1230,1255]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pseudovari" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FF4DA7BFAB34FB72" bold="true" box="[179,305,1230,1255]" italics="true" pageId="5" pageNumber="6">pseudovari</emphasis>
</taxonomicName>
</emphasis>
Gibbs,
<taxonomicNameLabel id="9F835783FFDEFFC9FE77A7BEABE7FB72" box="[393,482,1231,1255]" pageId="5" pageNumber="6" rank="species">
<emphasis id="84B0EAF8FFDEFFC9FE77A7BEABA6FB72" bold="true" box="[393,419,1231,1255]" pageId="5" pageNumber="6">sp</emphasis>
.
<emphasis id="84B0EAF8FFDEFFC9FE4FA7BEABE7FB72" bold="true" box="[433,482,1231,1255]" pageId="5" pageNumber="6">
<date id="C27A102AFFDEFFC9FE4FA7BEABE7FB72" box="[433,482,1231,1255]" pageId="5" pageNumber="6">nov.</date>
</emphasis>
</taxonomicNameLabel>
8.
<taxonomicName id="71C44D69FFDEFFC9FF4DA783AB6DFA9F" authority="Gibbs" authorityName="Gibbs" box="[179,360,1266,1291]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sagittella" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FF4DA783AB1EFA9E" bold="true" box="[179,283,1266,1291]" italics="true" pageId="5" pageNumber="6">sagittella</emphasis>
Gibbs
</taxonomicName>
,
<taxonomicNameLabel id="9F835783FFDEFFC9FE8DA782ABC9FA9E" box="[371,460,1267,1291]" pageId="5" pageNumber="6" rank="species">
<emphasis id="84B0EAF8FFDEFFC9FE8DA782AB88FA9E" bold="true" box="[371,397,1267,1291]" pageId="5" pageNumber="6">sp</emphasis>
.
<emphasis id="84B0EAF8FFDEFFC9FE65A782ABC9FA9E" bold="true" box="[411,460,1267,1291]" pageId="5" pageNumber="6">
<date id="C27A102AFFDEFFC9FE65A782ABC9FA9E" box="[411,460,1267,1291]" pageId="5" pageNumber="6">nov.</date>
</emphasis>
</taxonomicNameLabel>
9.
<taxonomicName id="71C44D69FFDEFFC9FF4DA667AB8FFABB" authority="Whalley, 1978" authorityName="Whalley" authorityYear="1978" box="[179,394,1302,1327]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFDEFFC9FF4DA667AAE5FABA" bold="true" box="[179,224,1302,1327]" italics="true" pageId="5" pageNumber="6">vari</emphasis>
<bibRefCitation id="D2554B1BFFDEFFC9FF19A667AB8FFABB" author="Whalley" box="[231,394,1302,1326]" pageId="5" pageNumber="21" refString="Whalley, P. E. S. (1978) New taxa of fossil and recent Micropterigidae with a discussion of their evolution and a comment on the evolution of Lepidoptera (Insecta). Annals of the Transvaal Museum, 31, 71 - 86." type="journal article" year="1978">Whalley, 1978</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDEFFC9FF69A6F2AB0DF9B0" pageId="5" pageNumber="6" type="key">
<paragraph id="B67B36EAFFDEFFC9FF69A6F2A9D3FA0F" blockId="5.[151,982,1410,1435]" box="[151,982,1410,1435]" pageId="5" pageNumber="6">
<emphasis id="84B0EAF8FFDEFFC9FF69A6F2A940FA0E" bold="true" box="[151,837,1410,1435]" pageId="5" pageNumber="6">
Key to species of
<taxonomicName id="71C44D69FFDEFFC9FE9AA6F3A806FA0E" ID-CoL="8ZZ76" box="[356,515,1410,1435]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDEFFC9FE9AA6F3A806FA0E" bold="true" box="[356,515,1410,1435]" italics="true" pageId="5" pageNumber="6">Agrionympha</emphasis>
</taxonomicName>
based on adult maculation
</emphasis>
(
<figureCitation id="2EFF2A6FFFDEFFC9FCAAA6F3A9A4FA0F" box="[852,929,1410,1434]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
AI)
</paragraph>
<paragraph id="B67B36EAFFDEFFC9FF69A6B4AB0DF9B0" blockId="5.[151,1436,1477,1681]" pageId="5" pageNumber="6">Although a highly conservative genus, the species can be confidently diagnosed from FW maculation and certain other colour differences. In the key, the nomenclature for the 4 band marks follows that outlined in the generic diagnosis.</paragraph>
</subSubSection>
<subSubSection id="FEDE6561FFDEFFCAFF39A540AF9EFB86" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="B67B36EAFFDEFFC9FF39A540A85FF904" blockId="5.[151,1436,1477,1681]" pageId="5" pageNumber="6">
NOTE: males are not yet known for
<taxonomicName id="71C44D69FFDEFFC9FDA0A543A88EF9DC" box="[606,651,1586,1609]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFDEFFC9FDA0A543A88EF9DC" box="[606,651,1586,1609]" italics="true" pageId="5" pageNumber="6">vari</emphasis>
</taxonomicName>
,
<emphasis id="84B0EAF8FFDEFFC9FD67A540A8FDF9DC" box="[665,760,1585,1609]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="71C44D69FFDEFFC9FD67A540A8F0F9DC" box="[665,757,1585,1609]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jansella">jansella</taxonomicName>
,
</emphasis>
and
<taxonomicName id="71C44D69FFDEFFC9FCD0A540A9AFF9DC" box="[814,938,1585,1609]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pseudovari">
<emphasis id="84B0EAF8FFDEFFC9FCD0A540A9AFF9DC" box="[814,938,1585,1609]" italics="true" pageId="5" pageNumber="6">pseudovari</emphasis>
</taxonomicName>
; while sexual dimorphism in wing pattern is minimal or non-existent in the great majority of micropterigids, it does occur in a few taxa, and its possible occurrence here must always be kept in mind.
</paragraph>
<paragraph id="B67B36EAFFDEFFC9FF69A5CDAF8EF8BC" blockId="5.[151,1436,1724,2038]" pageId="5" pageNumber="6">
1 Claval mark confluent with median band (e.g.
<figureCitation id="2EFF2A6FFFDEFFC9FD61A5CDA8D3F944" box="[671,726,1724,1745]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A or F)................................................... 2 Claval mark discrete or absent (e.g.
<figureCitation id="2EFF2A6FFFDEFFC9FDC5A5A8A876F97B" box="[571,627,1753,1774]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
B or G)............................................................ 3 2(1) Median band complete, forming with the claval mark an arrowhead-like figure (
<figureCitation id="2EFF2A6FFFDEFFC9FC1EA587AE1CF89E" box="[992,1049,1782,1803]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
F)...............
<taxonomicName id="71C44D69FFDEFFC9FB0EA587AF4CF89E" box="[1264,1353,1782,1803]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sagittella" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FB0EA587AF4CF89E" box="[1264,1353,1782,1803]" italics="true" pageId="5" pageNumber="6">sagittella</emphasis>
</taxonomicName>
<emphasis id="84B0EAF8FFDEFFC9FAB1A587AF9EF89E" bold="true" box="[1359,1435,1782,1803]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="9F835783FFDEFFC9FAB1A587AF9EF89E" box="[1359,1435,1782,1803]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Median band interrupted in middle: anterior part forming a broad L-shape with the claval mark; posterior part appearing dis-
</paragraph>
<paragraph id="B67B36EAFFDEFFCAFF1FA440A962FD89" blockId="5.[151,1436,1724,2038]" lastBlockId="6.[151,1142,482,540]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
tally displaced, forming a discrete spot on dorsum (
<figureCitation id="2EFF2A6FFFDEFFC9FD33A440A903F8D3" box="[717,774,1841,1862]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
A)..............................................
<taxonomicName id="71C44D69FFDEFFC9FAB7A440AF9EF8D3" box="[1353,1435,1841,1862]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDEFFC9FAB7A440AF9EF8D3" box="[1353,1435,1841,1862]" italics="true" pageId="5" pageNumber="6">capensis</emphasis>
</taxonomicName>
3(1) Median band complete from costa to dorsum (e.g.
<figureCitation id="2EFF2A6FFFDEFFC9FD3FA43FA8FFF8F6" box="[705,762,1870,1891]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
C), postmedian band either complete (e.g.
<figureCitation id="2EFF2A6FFFDEFFC9FB70A43FAEC3F8F6" box="[1166,1222,1870,1891]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
B) or incomplete (e.g.
<figureCitation id="2EFF2A6FFFDEFFC9FF1FA41DAB1CF814" box="[225,281,1900,1921]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
C............................................................................................. 4 Median band represented by only a spot on dorsum, postmedian band complete from costa to termen (
<figureCitation id="2EFF2A6FFFDEFFC9FB47A4F8AEF7F80B" box="[1209,1266,1929,1950]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
E)..................................................................................................
<taxonomicName id="71C44D69FFDEFFC9FB10A4D7AF4CF82E" box="[1262,1353,1958,1979]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kroonella" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FB10A4D7AF4CF82E" box="[1262,1353,1958,1979]" italics="true" pageId="5" pageNumber="6">kroonella</emphasis>
</taxonomicName>
<emphasis id="84B0EAF8FFDEFFC9FAB1A4D7AF9EF82E" bold="true" box="[1359,1435,1958,1979]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="9F835783FFDEFFC9FAB1A4D7AF9EF82E" box="[1359,1435,1958,1979]" pageId="5" pageNumber="6" rank="species">
sp.
<date id="C27A102AFFDEFFC9FA8CA4D7AF9EF82E" box="[1394,1435,1958,1979]" pageId="5" pageNumber="6">nov.</date>
</taxonomicNameLabel>
</emphasis>
4(3) Claval mark absent (
<figureCitation id="2EFF2A6FFFDEFFC9FE59A4B5ABE5F84C" box="[423,480,1988,2009]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
G)...............................................................
<taxonomicName id="71C44D69FFDEFFC9FB03A4B5AF4CF84C" box="[1277,1353,1988,2009]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jansella" status="sp. nov.">
<emphasis id="84B0EAF8FFDEFFC9FB03A4B5AF4CF84C" box="[1277,1353,1988,2009]" italics="true" pageId="5" pageNumber="6">jansella</emphasis>
</taxonomicName>
<emphasis id="84B0EAF8FFDEFFC9FAB1A4B2AF9EF84D" bold="true" box="[1359,1435,1987,2008]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="9F835783FFDEFFC9FAB1A4B2AF9EF84D" box="[1359,1435,1987,2008]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Claval mark distinct (e.g.
<figureCitation id="2EFF2A6FFFDEFFC9FE24A490A817F863" box="[474,530,2017,2038]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
D)........................................................................ 5 5(4) Apical band a distinctive, usually dark-edged spot or short transverse band (e.g.
<figureCitation id="2EFF2A6FFFDDFFCAFC1DA3E6AE1EFF39" box="[995,1051,151,172]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
H)........................... 6 Apical band exceedingly reduced, represented by a few light scales, or completely absent (
<figureCitation id="2EFF2A6FFFDDFFCAFBB8A3C4AE7BFF5F" box="[1094,1150,181,202]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
D)...
<taxonomicName id="71C44D69FFDDFFCAFB37A3C4AF4CFF5F" box="[1225,1353,181,202]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="fuscoapicella" status="sp. nov.">
<emphasis id="84B0EAF8FFDDFFCAFB37A3C4AF4CFF5F" box="[1225,1353,181,202]" italics="true" pageId="6" pageNumber="7">fuscoapicella</emphasis>
</taxonomicName>
<emphasis id="84B0EAF8FFDDFFCAFAB1A3C4AF99FF5F" bold="true" box="[1359,1436,181,202]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="9F835783FFDDFFCAFAB1A3C4AF99FF5F" box="[1359,1436,181,202]" pageId="6" pageNumber="7" rank="species">
sp.
<date id="C27A102AFFDDFFCAFA8CA3C4AF99FF5F" box="[1394,1436,181,202]" pageId="6" pageNumber="7">nov.</date>
</taxonomicNameLabel>
</emphasis>
6(5) Postmedian band interrupted i.e. not a continuous band from costa to termen (e.g.
<figureCitation id="2EFF2A6FFFDDFFCAFC0AA3A3AE29FF72" box="[1012,1068,210,231]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
C)........................... 7 Postmedian band complete from costa to termen (
<figureCitation id="2EFF2A6FFFDDFFCAFD43A39EA8F2FE91" box="[701,759,239,260]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
B)..........................................
<taxonomicName id="71C44D69FFDDFFCAFAECA381AF4CFE90" box="[1298,1353,240,261]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="karoo" status="sp. nov.">
<emphasis id="84B0EAF8FFDDFFCAFAECA381AF4CFE90" box="[1298,1353,240,261]" italics="true" pageId="6" pageNumber="7">karoo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F835783FFDDFFCAFAB1A39EAF9EFE91" box="[1359,1435,239,260]" pageId="6" pageNumber="7" rank="species">
<emphasis id="84B0EAF8FFDDFFCAFAB1A39EAF9EFE91" bold="true" box="[1359,1435,239,260]" pageId="6" pageNumber="7">
sp.
<date id="C27A102AFFDDFFCAFA8CA39EAF9EFE91" box="[1394,1435,239,260]" pageId="6" pageNumber="7">nov.</date>
</emphasis>
</taxonomicNameLabel>
7(6) Postmedian band interrupted in middle, but present as distinctive spots on both costa and termen (e.g.
<figureCitation id="2EFF2A6FFFDDFFCAFB4AA27CAEEEFEB7" box="[1204,1259,269,290]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
C).......... 8 Postmedian band present only as a narrow band from costa to the middle of the wing (
<figureCitation id="2EFF2A6FFFDDFFCAFBE5A25BAE56FEAA" box="[1051,1107,298,319]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
H)...............
<taxonomicName id="71C44D69FFDDFFCAFAC5A25BAF99FEAA" box="[1339,1436,298,319]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFDDFFCAFAC5A25BAF99FEAA" box="[1339,1436,298,319]" italics="true" pageId="6" pageNumber="7">pseliacma</emphasis>
</taxonomicName>
7(6) All silvery bands distinctly edged with black scales; costal part of postmedian band almost circular (
<figureCitation id="2EFF2A6FFFDDFFCAFB5FA236AEDFFEC9" box="[1185,1242,327,348]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
C)..................................................................................................
<taxonomicName id="71C44D69FFDDFFCAFB20A214AF4DFEEF" box="[1246,1352,357,378]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pseudovari" status="sp. nov.">
<emphasis id="84B0EAF8FFDDFFCAFB20A214AF4DFEEF" box="[1246,1352,357,378]" italics="true" pageId="6" pageNumber="7">pseudovari</emphasis>
</taxonomicName>
<emphasis id="84B0EAF8FFDDFFCAFAB1A214AF9EFEEF" bold="true" box="[1359,1435,357,378]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="9F835783FFDDFFCAFAB1A214AF9EFEEF" box="[1359,1435,357,378]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Silvery bands not edged with black scales; costal part of postmedian band nearly 3 times longer than wide (
<figureCitation id="2EFF2A6FFFDDFFCAFB18A2F3AF1AFE02" box="[1254,1311,386,407]" captionStart="FIGURE 2" captionStartId="8.[151,250,775,798]" captionTargetBox="[269,1419,198,724]" captionTargetId="figure@8.[159,1427,193,772]" captionTargetPageId="8" captionText="FIGURE 2. Forewing maculation of Agrionympha species: A, A. capensis; B, A. karoo, C, A. pseudovari, D, A. fuscoapicella; E, A. kroonella, F, A. sagittella; G, A. jansella; H, A. pseliacma; I, A. vari." httpUri="https://zenodo.org/record/203555/files/figure.png" pageId="6" pageNumber="7">
Fig.
<date id="C27A102AFFDDFFCAFAEDA2F3AF1AFE02" box="[1299,1311,386,407]" pageId="6" pageNumber="7">2</date>
</figureCitation>
I).....
<taxonomicName id="71C44D69FFDDFFCAFA8FA2F3AF99FE02" box="[1393,1436,386,407]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vari">
<emphasis id="84B0EAF8FFDDFFCAFA8FA2F3AF99FE02" box="[1393,1436,386,407]" italics="true" pageId="6" pageNumber="7">vari</emphasis>
</taxonomicName>
<emphasis id="84B0EAF8FFDDFFCAFF69A292A9E3FE6E" bold="true" box="[151,998,482,507]" pageId="6" pageNumber="7">
Key to species of
<taxonomicName id="71C44D69FFDDFFCAFE9AA293A806FE6E" box="[356,515,482,507]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="84B0EAF8FFDDFFCAFE9AA293A806FE6E" bold="true" box="[356,515,482,507]" italics="true" pageId="6" pageNumber="7">Agrionympha</emphasis>
</taxonomicName>
based on male postabdomen &amp; genitalia
</emphasis>
(
<figureCitation id="2EFF2A6FFFDDFFCAFC08A293AE6AFE6F" box="[1014,1135,482,506]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="14.[151,250,1457,1480]" captionStartId-1="16.[151,250,1415,1438]" captionStartId-2="17.[151,250,1478,1501]" captionStartId-3="18.[151,250,1913,1936]" captionTargetBox-0="[160,1416,529,1421]" captionTargetBox-1="[311,1355,490,1414]" captionTargetBox-2="[259,1325,201,1478]" captionTargetBox-3="[313,1332,581,1845]" captionTargetId-0="figure@14.[160,1427,519,1454]" captionTargetId-1="figure@16.[223,1363,483,1474]" captionTargetId-2="figure@17.[254,1332,193,1527]" captionTargetId-3="figure@18.[244,1340,573,1969]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionText-0="FIGURE 7. Agrionympha capensis Whalley, genitalia: A E, male, Storms River: A, ventral; B, lateral; C, dorsal; D, phallus lateral; E, phallus dorsal. F G, female, Bloukrans Pass: F, extended terminal segments of abdomen; G, internal female genitalia. cb, corpus bursae; dph, distal portion of phallus; dsp, ductus of spermatheca; go, gonopore with radial folds; lag, lagena; lap, lappet, mpl, median plate; pap, papilla; pph, proximal portion of phallus; scl ix, ninth sclerite; scl x, tenth sclerite; S viii, eighth sternite; spm, spermatheca; T viii, eighth tergite; T x, tenth tergum; utr, distal bulb of utriculus; va, valva; vb, ventral bulb." captionText-1="FIGURE 8. Agrionympha genitalia. A D, A. fuscoapicella sp. nov. male, Hogsback: A, lateral, inset - ventral view of median plate; B, dorsal, C, phallus lateral; D, phallus dorsal. E F, A. jansella sp. nov. female, Karkloof Falls: E, extended terminal segments of abdomen; F, internal female genitalia." captionText-2="FIGURE 9. Agrionympha male genitalia. A D, A. karoo, sp. nov. Toorberg: A, lateral, inset ventral view of median plate; B, dorsal; C, phallus lateral; D, phallus dorsal. E H, A. kroonella, sp. nov. Mariepskop: E, lateral, inset ventral view of median plate; F, dorsal; G, phallus lateral; H, phallus dorsal." captionText-3="FIGURE 10. Agrionympha male genitalia. A B, A. pseliacma, Meyrick, Karkloof Falls: A, lateral view, from slide 3830, Transvaal Museum, Pretoria; B, detail of phallus, lateral. C G, A. sagittella, sp. nov. Hogsback: C, lateral; D, dorsal; E, ventral; F, phallus, lateral; G, phallus dorsal. H J, Agrionympha species 1 , Diepwalle Forest: H, lateral; I, phallus lateral; J, phallus dorsal." httpUri-0="https://zenodo.org/record/203560/files/figure.png" httpUri-1="https://zenodo.org/record/203561/files/figure.png" httpUri-2="https://zenodo.org/record/203562/files/figure.png" httpUri-3="https://zenodo.org/record/203563/files/figure.png" pageId="6" pageNumber="7">Figs. 710</figureCitation>
) Note: males unknown for
<emphasis id="84B0EAF8FFDDFFCAFE45A175A88BFD89" box="[443,654,516,540]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="71C44D69FFDDFFCAFE45A175A80FFD89" box="[443,522,516,540]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vari">A. vari</taxonomicName>
,
<taxonomicName id="71C44D69FFDDFFCAFDEBA175A88BFD89" box="[533,654,516,540]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="jansella">A. jansella</taxonomicName>
</emphasis>
and
<emphasis id="84B0EAF8FFDDFFCAFD3AA175A962FD89" box="[708,871,516,540]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="71C44D69FFDDFFCAFD3AA175A966FD89" box="[708,867,516,540]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pseudovari">A. pseudovari</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="B67B36EAFFDDFFCAFF69A137AF99FD03" blockId="6.[151,1436,582,1043]" pageId="6" pageNumber="7">1 Specialized hair-sensilla present on membranous dorsum between tergum VIII and tergum X lobes; valvae length:width ratio 4.0 or less.......................................................................................... 3 Hair-sensilla absent in said area; valvae length:width ratio 4.05.7............................................... 2</paragraph>
<paragraph id="B67B36EAFFDDFFCAFF69A1EFAF9EFD7B" blockId="6.[151,1436,582,1043]" pageId="6" pageNumber="7">
2(1) Hair-sensilla present on anterior border of tergum VIII; valvae elongate, length:width ratio greater than 5 (
<figureCitation id="2EFF2A6FFFDDFFCAFB22A1EFAF24FD26" box="[1244,1313,670,691]" captionStart="FIGURE 10" captionStartId="18.[151,250,1913,1936]" captionTargetBox="[313,1332,581,1845]" captionTargetId="figure@18.[244,1340,573,1969]" captionTargetPageId="18" captionText="FIGURE 10. Agrionympha male genitalia. A B, A. pseliacma, Meyrick, Karkloof Falls: A, lateral view, from slide 3830, Transvaal Museum, Pretoria; B, detail of phallus, lateral. C G, A. sagittella, sp. nov. Hogsback: C, lateral; D, dorsal; E, ventral; F, phallus, lateral; G, phallus dorsal. H J, Agrionympha species 1 , Diepwalle Forest: H, lateral; I, phallus lateral; J, phallus dorsal." httpUri="https://zenodo.org/record/203563/files/figure.png" pageId="6" pageNumber="7">Fig. 10</figureCitation>
C)....................................................................................................
<taxonomicName id="71C44D69FFDDFFCAFABCA1CDAB2EFD7B" authority="Tergum" authorityName="Tergum" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sagittella">
<emphasis id="84B0EAF8FFDDFFCAFABCA1CDAF9EFD44" box="[1346,1435,700,721]" italics="true" pageId="6" pageNumber="7">sagittella</emphasis>
Tergum
</taxonomicName>
VIII devoid of hair-sensilla; length:width ratio of valvae 4.0 (
<figureCitation id="2EFF2A6FFFDDFFCAFC83A1A8A9B3FD7B" box="[893,950,729,750]" captionStart="FIGURE 7" captionStartId="14.[151,250,1457,1480]" captionTargetBox="[160,1416,529,1421]" captionTargetId="figure@14.[160,1427,519,1454]" captionTargetPageId="14" captionText="FIGURE 7. Agrionympha capensis Whalley, genitalia: A E, male, Storms River: A, ventral; B, lateral; C, dorsal; D, phallus lateral; E, phallus dorsal. F G, female, Bloukrans Pass: F, extended terminal segments of abdomen; G, internal female genitalia. cb, corpus bursae; dph, distal portion of phallus; dsp, ductus of spermatheca; go, gonopore with radial folds; lag, lagena; lap, lappet, mpl, median plate; pap, papilla; pph, proximal portion of phallus; scl ix, ninth sclerite; scl x, tenth sclerite; S viii, eighth sternite; spm, spermatheca; T viii, eighth tergite; T x, tenth tergum; utr, distal bulb of utriculus; va, valva; vb, ventral bulb." httpUri="https://zenodo.org/record/203560/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
B)...............................
<taxonomicName id="71C44D69FFDDFFCAFAB7A1A8AF9EFD7B" box="[1353,1435,729,750]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="84B0EAF8FFDDFFCAFAB7A1A8AF9EFD7B" box="[1353,1435,729,750]" italics="true" pageId="6" pageNumber="7">capensis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="B67B36EAFFDDFFCAFF69A187AF99FCF6" blockId="6.[151,1436,582,1043]" pageId="6" pageNumber="7">
3(1) With less than 45 hair-sensilla; median plate narrow, tapered anteriorly, at least
<date id="C27A102AFFDDFFCAFC27A187A9F4FC9E" box="[985,1009,758,779]" pageId="6" pageNumber="7">3x</date>
longer than wide; phallus without conspicuous internal sclerotised body........................................................................... 4 With about 80 hair-sensilla; median plate broad and rectangular, approx.
<date id="C27A102AFFDDFFCAFC62A040A9B1FCD3" box="[924,948,817,838]" pageId="6" pageNumber="7">2x</date>
longer than wide; ductus ejaculatorius thickening just inside gonopore conspicuous, roughly textured and somewhat sclerotised (
<figureCitation id="2EFF2A6FFFDDFFCAFC2AA03FAE08FCF6" box="[980,1037,846,867]" captionStart="FIGURE 9" captionStartId="17.[151,250,1478,1501]" captionTargetBox="[259,1325,201,1478]" captionTargetId="figure@17.[254,1332,193,1527]" captionTargetPageId="17" captionText="FIGURE 9. Agrionympha male genitalia. A D, A. karoo, sp. nov. Toorberg: A, lateral, inset ventral view of median plate; B, dorsal; C, phallus lateral; D, phallus dorsal. E H, A. kroonella, sp. nov. Mariepskop: E, lateral, inset ventral view of median plate; F, dorsal; G, phallus lateral; H, phallus dorsal." httpUri="https://zenodo.org/record/203562/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
E).......................
<taxonomicName id="71C44D69FFDDFFCAFABFA03FAF99FCF6" box="[1345,1436,846,867]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kroonella">
<emphasis id="84B0EAF8FFDDFFCAFABFA03FAF99FCF6" box="[1345,1436,846,867]" italics="true" pageId="6" pageNumber="7">kroonella</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="B67B36EAFFDDFFCAFF69A01DAEFEFC4C" blockId="6.[151,1436,582,1043]" pageId="6" pageNumber="7">
4.(3) Gonopore small, less than half the width of the Phallus between lappets; neck of phallus constricted in dorsal view..... 5 Gonopore large, well over half width of aedeagus between lappets; neck of phallus not constricted.. undescribed species 1 NOTE: while
<taxonomicName id="71C44D69FFDDFFCAFE96A0D7ABCEFC2E" box="[360,459,934,955]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pseliacma">
<emphasis id="84B0EAF8FFDDFFCAFE96A0D7ABCEFC2E" bold="true" box="[360,459,934,955]" italics="true" pageId="6" pageNumber="7">pseliacma</emphasis>
</taxonomicName>
will run to this couplet, the unique slide preparation made by Janse does not allow observation of the phallus character used here; the species has more than 30 hair-sensilla like both species 1 and
<taxonomicName id="71C44D69FFDDFFCAFB8BA0B5AEF0FC4C" box="[1141,1269,964,985]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="fuscoapicella">
<emphasis id="84B0EAF8FFDDFFCAFB8BA0B5AEF0FC4C" box="[1141,1269,964,985]" italics="true" pageId="6" pageNumber="7">fuscoapicella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B67B36EAFFDDFFCAFF69A090AF9EFB86" blockId="6.[151,1436,582,1043]" pageId="6" pageNumber="7">
5(4) More than 30 hair-sensilla on dorsum IX; gonopore plain, without a dorsal prominence (
<figureCitation id="2EFF2A6FFFDDFFCAFBC8A090AE68FC63" box="[1078,1133,993,1014]" captionStart="FIGURE 8" captionStartId="16.[151,250,1415,1438]" captionTargetBox="[311,1355,490,1414]" captionTargetId="figure@16.[223,1363,483,1474]" captionTargetPageId="16" captionText="FIGURE 8. Agrionympha genitalia. A D, A. fuscoapicella sp. nov. male, Hogsback: A, lateral, inset - ventral view of median plate; B, dorsal, C, phallus lateral; D, phallus dorsal. E F, A. jansella sp. nov. female, Karkloof Falls: E, extended terminal segments of abdomen; F, internal female genitalia." httpUri="https://zenodo.org/record/203561/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
C)............
<taxonomicName id="71C44D69FFDDFFCAFAE2A090AB18FB86" authority="Fewer" authorityName="Fewer" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="fuscoapicella">
<emphasis id="84B0EAF8FFDDFFCAFAE2A090AF99FC63" box="[1308,1436,993,1014]" italics="true" pageId="6" pageNumber="7">fuscoapicella</emphasis>
Fewer
</taxonomicName>
than 20 hair-sensilla on dorsum IX; gonopore with its proximal margin raised on a prominence (
<figureCitation id="2EFF2A6FFFDDFFCAFB42A08FAEF0FB86" box="[1212,1269,1022,1043]" captionStart="FIGURE 9" captionStartId="17.[151,250,1478,1501]" captionTargetBox="[259,1325,201,1478]" captionTargetId="figure@17.[254,1332,193,1527]" captionTargetPageId="17" captionText="FIGURE 9. Agrionympha male genitalia. A D, A. karoo, sp. nov. Toorberg: A, lateral, inset ventral view of median plate; B, dorsal; C, phallus lateral; D, phallus dorsal. E H, A. kroonella, sp. nov. Mariepskop: E, lateral, inset ventral view of median plate; F, dorsal; G, phallus lateral; H, phallus dorsal." httpUri="https://zenodo.org/record/203562/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
C).......
<taxonomicName id="71C44D69FFDDFFCAFA9AA08FAF9EFB86" box="[1380,1435,1022,1043]" class="Insecta" family="Micropterigidae" genus="Agrionympha" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="karoo">
<emphasis id="84B0EAF8FFDDFFCAFA9AA08FAF9EFB86" box="[1380,1435,1022,1043]" italics="true" pageId="6" pageNumber="7">karoo</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

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<taxonomicName id="8AF2BFECF0CB26AD954FD5EC9207C0DA" ID-CoL="56JK8" authority="Thomas 1917" authorityName="Thomas" authorityYear="1917" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageNumber="1181" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B8989CB72832CF99492C3D8281BA18BC" italics="true" pageNumber="1181">Thomasomys gracilis</emphasis>
Thomas 1917
</taxonomicName>
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<subSubSection id="542EAD7C2820861D0B6B980161ACB7F7" pageNumber="1181" type="reference_group">
<paragraph id="48E53C96EA84722A775F6CADDF2933A6" pageNumber="1181">
<treatmentCitation id="2862B806BC8E4E2549D08C03B779ACFC" author="Thomas" authority="Thomas 1917" authorityName="Thomas" authorityYear="1917" class="Mammalia" family="Cricetidae" genus="Thomasomys" issue="4" journal="Smithson. Misc. Coll." kingdom="Animalia" order="Rodentia" page="2" pageNumber="1181" phylum="Chordata" rank="species" species="gracilis" title="Thomasomys gracilis" volume="68" year="1917">
<taxonomicName id="0CEEFF18F9E03354FF371B3FD46EDCC8" ID-CoL="56JK8" authority="Thomas 1917" authorityName="Thomas" authorityYear="1917" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" pageNumber="1181" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="51B5BF0AE8ACAC366EBADA324C1D1656" italics="true" pageNumber="1181">Thomasomys gracilis</emphasis>
Thomas 1917
</taxonomicName>
,
<bibCitation id="29BCB045B0B11FE6A030A8013AD5772F" author="Thomas" issue="4" journal="Smithson. Misc. Coll." pageNumber="1181" pagination="2" title="Thomasomys gracilis" volume="68" year="1917">Smithson. Misc. Coll., 68 (4): 2</bibCitation>
</treatmentCitation>
.
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<paragraph id="9CC16D036D9D60707CD204294F05B99C" pageNumber="1181">
<emphasis id="EAF1928F3D1C61897EDC3C2A5D8FED6C" inLineHeading="true" pageNumber="1181">Type Locality:</emphasis>
<materialsCitation id="6A1B995D08849D1AA8027CD137698F15" ID-GBIF-Occurrence="4867286435" country="Peru" elevation="3658.0" pageNumber="1181" stateProvince="Cusco">
<collectingCountry id="9158342D4DC9BB795EBB15260C83DB08" name="Peru">Perú</collectingCountry>
,
<collectingRegion id="1DDDFF6F543975217B6DC36D2BA35674" country="Peru" name="Cusco">Cuzco</collectingRegion>
Dept., Machu Picchu,
<quantity id="E649E38593AE3C08DF6689B15F8A01C7" metricMagnitude="0" metricUnit="m" metricValue="3.6576" unit="ft" value="12.0">12,000 ft</quantity>
(
<quantity id="2FF1BA55AD4B3FA3EB0B42274BDCB0CB" metricMagnitude="3" metricUnit="m" metricValue="3.658" unit="m" value="3658.0">
<elevation id="6D3EAEEC350FB669C0DF0E3D515EEB4A">3658 m</elevation>
</quantity>
).
</materialsCitation>
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<subSubSection id="2FD5B68C165C04B99AFAE49B5670E154" pageNumber="1181" type="vernacular_names">
<paragraph id="A960CE8D33D4C5A662696064C3315111" pageNumber="1181">
<emphasis id="B87ACE71D76530B84DC665C46295D3AA" inLineHeading="true" pageNumber="1181">Vernacular Names:</emphasis>
<vernacularName id="81B2E6C7757647724BB2516F2BF6EB2D" language="eng" pageNumber="1181">Gracile Thomasomys</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="A336C6A2D166260E97F788BF162109C6" pageNumber="1181" type="distribution">
<paragraph id="9B7860B63F94FF6767985D0C18031F12" pageNumber="1181">
<emphasis id="FF3F57BCFF4F44FD59B13695B000131F" inLineHeading="true" pageNumber="1181">Distribution:</emphasis>
Andes of SE
<collectingCountry id="22FF45AC66EF84FCF33F96675A82F541" name="Peru">Perú</collectingCountry>
, about
<quantity id="21F2740F5CC6E1C760881C5284DBF68E" metricMagnitude="3" metricUnit="m" metricValue="3.525" metricValueMax="4.3" metricValueMin="2.75" unit="m" value="3525.0" valueMax="4300.0" valueMin="2750.0">2750-4300 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="BEE6EA780DCCE295D807BBFA0346D437" pageNumber="1181" type="conservation">
<paragraph id="22F11B2215892A333E8B829CCC8365D7" pageNumber="1181">
<emphasis id="583DD90467BA2A4F9539536EFFE0106A" inLineHeading="true" pageNumber="1181">Conservation:</emphasis>
<collectionCode id="AC1C9B19D34B83B102ADBDA62FCFBB27">IUCN</collectionCode>
Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="5C5726873FA0ADA0E40A36F940246A93" pageNumber="1181" type="discussion">
<paragraph id="DADE9112E9F410E3ACDE4A9DFF5C763C" pageNumber="1181">
<emphasis id="14F4D365CCC47055E3AAD8646B7174A8" inLineHeading="true" pageNumber="1181">Discussion:</emphasis>
Formerly included
<taxonomicName id="AF4BDCD0FCF59AF09C9D5301B000BB31" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="cinnameus">
<emphasis id="3C122663B65F6BECD13C49EA15886636" italics="true" pageNumber="1181">cinnameus</emphasis>
</taxonomicName>
and
<taxonomicName id="BE5368FF3FA6E7039E47B52259AA724D" authorityName="Anthony" authorityYear="1923" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="hudsoni">
<emphasis id="C1D797674C39E677A966561A991663A2" italics="true" pageNumber="1181">hudsoni</emphasis>
</taxonomicName>
as subspecies or referred synonyms (Cabrera, 1961;
<bibRefCitation id="807422F6D0D73D906E6D09F3C39A8C52" author="Musser, G. G. &amp; M. D. Carleton" refId="ref252333" refString="Musser, G. G., and M. D. Carleton. 1993. Family Muridae. Pp. 501 - 755, in: Mammal species of the world, a taxonomic and geographic reference, Second ed. (D. E. Wilson and D. M. Reeder, eds.). Smithsonian Institution Press, Washington D. C., xviii + 1206 pp." year="1993">Musser and Carleton, 1993</bibRefCitation>
, respectively), forms reinstated to species by
<bibRefCitation id="8E07437D3613E6BE6515A0DCE6B3EB85" author="Voss, R. S." refId="ref378425" refString="Voss, R. S. 2003. A new species of Thomasomys (Rodentia: Muridae) from eastern Ecuador, with remarks on mammalian diversity and biogeography in the Cordillera Oriental. American Museum Novitates, 3421: 1 - 47." year="2003">Voss (2003)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,215 @@
<document ID-DOI="10.25221/fee.389.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7164791" approvalRequired="24" approvalRequired_for_textStreams="24" checkinTime="1665244883012" checkinUser="felipe" docAuthor="Legalov, A. A." docDate="2019" docId="3E6D9136FFA2817B12CF79D9AA5B1F12" docLanguage="en" docName="FarEastEntomol.389.1-6.pdf" docOrigin="Far Eastern Entomologist 389" docSource="http://dx.doi.org/10.25221/fee.389.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Auletobius (Auletobius) siberutensis Legalov 2019, sp. n." docType="treatment" docVersion="2" lastPageNumber="4" masterDocId="C254E94EFFA38178122C7D28A9441A20" masterDocTitle="A new species of the genus Auletobius Desbrochers des Loges, 1869 (Coleoptera: Rhynchitidae) from Indonesia" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1665257860076" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of the genus Auletobius Desbrochers des Loges, 1869 (Coleoptera: Rhynchitidae) from Indonesia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Legalov, A. A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-08-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>389</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>6</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.389.1</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7164791</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:3E6D9136FFA2817B12CF79D9AA5B1F12" httpUri="http://treatment.plazi.org/id/3E6D9136FFA2817B12CF79D9AA5B1F12" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection pageId="1" pageNumber="2" type="nomenclature">
<paragraph blockId="1.[227,796,1265,1290]" box="[227,796,1265,1290]" pageId="1" pageNumber="2">
<heading box="[227,796,1265,1290]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis bold="true" box="[227,796,1265,1290]" pageId="1" pageNumber="2">
<taxonomicName authority="Legalov" authorityName="Legalov" authorityYear="2019" box="[227,724,1265,1290]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="siberutensis" status="sp. n." subGenus="Auletobius">
<emphasis bold="true" box="[227,346,1265,1289]" italics="true" pageId="1" pageNumber="2">Auletobius</emphasis>
(
<emphasis bold="true" box="[361,479,1265,1289]" italics="true" pageId="1" pageNumber="2">Auletobius</emphasis>
)
<emphasis bold="true" box="[495,625,1265,1289]" italics="true" pageId="1" pageNumber="2">siberutensis</emphasis>
Legalov
</taxonomicName>
,
<taxonomicNameLabel box="[736,796,1266,1290]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="1.[227,1053,1306,1328]" box="[227,1053,1306,1328]" pageId="1" pageNumber="2">
http://zoobank.org/NomenclaturalActs/
<uuid box="[612,1053,1306,1328]" pageId="1" pageNumber="2">CC4834D1-C429-481B-9809-123BCC086208</uuid>
</paragraph>
</subSubSection>
<subSubSection box="[227,319,1346,1370]" pageId="1" pageNumber="2" type="description">
<paragraph blockId="1.[227,319,1346,1370]" box="[227,319,1346,1370]" pageId="1" pageNumber="2">
<figureCitation box="[227,319,1346,1370]" captionStart="Figs" captionStartId="2.[265,309,1535,1557]" captionTargetBox="[246,1090,483,1476]" captionTargetId="figure-88@2.[227,1123,472,1502]" captionTargetPageId="2" captionText="Figs. 13. Auletobius siberutensis, sp. n., male. 1 holotype, habitus, dorsal view; 2 " figureDoi="http://doi.org/10.5281/zenodo.7164793" httpUri="https://zenodo.org/record/7164793/files/figure.png" pageId="1" pageNumber="2">Figs 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="materials_examined">
<paragraph blockId="1.[227,1127,1409,1590]" box="[265,1127,1409,1433]" pageId="1" pageNumber="2">
<materialsCitation box="[265,1127,1409,1433]" collectionCode="TYPE, MATERIAL" country="Indonesia" location="Mentawai Islands" municipality="Siberut Island" pageId="1" pageNumber="2" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode box="[265,481,1409,1433]" pageId="1" pageNumber="2">TYPE MATERIAL</collectionCode>
.
<typeStatus box="[492,589,1409,1433]" pageId="1" pageNumber="2">Holotype</typeStatus>
<specimenCount box="[617,639,1409,1432]" pageId="1" pageNumber="2" type="male"></specimenCount>
,
<emphasis bold="true" box="[651,760,1409,1433]" pageId="1" pageNumber="2">
<collectingCountry box="[651,760,1409,1433]" name="Indonesia" pageId="1" pageNumber="2">Indonesia</collectingCountry>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:3E6D9136FFA2817B12CF79D9AA5B1F12:B31B76FBFFA28179112A78A9AA871FB9" box="[774,963,1409,1433]" country="Indonesia" municipality="Siberut Island" name="Mentawai Islands" pageId="1" pageNumber="2">Mentawai Islands</location>
,
<collectingMunicipality box="[974,1123,1409,1433]" pageId="1" pageNumber="2">Siberut Island</collectingMunicipality>
,
</materialsCitation>
</paragraph>
<paragraph blockId="1.[227,1127,1409,1590]" pageId="1" pageNumber="2">
<materialsCitation collectingDate="2007-01" collectionCode="ISEA" elevation="75" location="Salappa" municipality="St. Jakl" pageId="1" pageNumber="2" specimenCount="75" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:3E6D9136FFA2817B12CF79D9AA5B1F12:B31B76FBFFA2817912CF7888A87C1F98" box="[227,312,1440,1464]" municipality="St. Jakl" name="Salappa" pageId="1" pageNumber="2">Salappa</location>
vill.,
<quantity box="[374,484,1440,1464]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="2" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
<elevation box="[374,484,1440,1464]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="1" pageNumber="2" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50100 m</elevation>
</quantity>
,
<date box="[495,640,1440,1465]" pageId="1" pageNumber="2" value="2007-01">
<collectingDate box="[495,640,1440,1465]" pageId="1" pageNumber="2" value="2007-01">January 2007</collectingDate>
</date>
,
<collectingMunicipality box="[651,730,1440,1464]" pageId="1" pageNumber="2">St. Jakl</collectingMunicipality>
(
<collectionCode box="[744,809,1440,1464]" pageId="1" pageNumber="2">ISEA</collectionCode>
).
<typeStatus box="[827,935,1441,1465]" pageId="1" pageNumber="2">Paratypes</typeStatus>
: the same label as in
<typeStatus box="[254,350,1471,1495]" pageId="1" pageNumber="2">holotype</typeStatus>
,
<specimenCount box="[361,505,1471,1495]" pageId="1" pageNumber="2" type="generic">75 specimens</specimenCount>
(
<collectionCode box="[520,585,1472,1496]" pageId="1" pageNumber="2">ISEA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
<paragraph blockId="1.[227,1127,1409,1590]" pageId="1" pageNumber="2">
DESCRIPTION. MALE. Body length (without rostrum)
<quantity box="[888,1022,1503,1527]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="1" pageNumber="2" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.61.9 mm</quantity>
. Rostrum length
<quantity box="[301,434,1534,1558]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="2" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
. Body brown, covered with decumbent hair with metallic lustre.
</paragraph>
<paragraph blockId="1.[227,1127,1409,1590]" box="[227,936,1565,1590]" pageId="1" pageNumber="2">Antennomeres 27 and procoxae, profemora and protibiae yellow.</paragraph>
<paragraph blockId="2.[227,1127,264,441]" box="[265,1126,264,288]" pageId="2" pageNumber="3">Head subparallel behind eye. Mandibles externally dentate. Rostrum quite long,</paragraph>
<paragraph blockId="2.[227,1127,264,441]" box="[227,1127,294,319]" pageId="2" pageNumber="3">almost straight, 1.3 times as long as pronotum, 5.3 times as long as wide at apex, 5.8</paragraph>
<paragraph blockId="2.[227,1127,264,441]" pageId="2" pageNumber="3">times as long as wide in middle, 5.0 times as long as wide at base, smooth, without carina or stria. Eyes large, stark convex, rounded. Forehead convex, finely punctate,</paragraph>
<paragraph blockId="2.[227,1127,264,441]" lastBlockId="2.[227,1127,1535,1612]" pageId="2" pageNumber="3">
1.9 times as wide as rostrum base width. Temples very short. Antennae inserted at base of rostrum, reaching elytral base. Antennomeres 1 and 2 almost oval. Antennomere aedeagus,
<typeStatus box="[330,415,1563,1584]" pageId="2" pageNumber="3">paratype</typeStatus>
, dorsal view; 3 armament of endophallus, dorsal view. Scale bar 0.5
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7164793" ID-Zenodo-Dep="7164793" box="[265,1127,1535,1557]" httpUri="https://zenodo.org/record/7164793/files/figure.png" pageId="2" pageNumber="3" startId="2.[265,309,1535,1557]" targetBox="[246,1090,483,1476]" targetPageId="2">
<paragraph blockId="2.[227,1127,1535,1612]" box="[265,1127,1535,1557]" pageId="2" pageNumber="3">
Figs. 13.
<taxonomicName authority="Legalov, 2019" authorityName="Legalov" authorityYear="2019" box="[371,597,1535,1556]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="siberutensis" status="sp. n.">
<emphasis box="[371,597,1535,1556]" italics="true" pageId="2" pageNumber="3">Auletobius siberutensis</emphasis>
</taxonomicName>
,
<emphasis bold="true" box="[611,668,1535,1556]" pageId="2" pageNumber="3">
<taxonomicNameLabel box="[611,668,1535,1556]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male. 1 holotype, habitus, dorsal view; 2
</paragraph>
</caption>
<paragraph blockId="2.[670,683,1650,1674]" box="[670,683,1650,1674]" pageId="2" pageNumber="3">3</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,1127,264,288]" pageId="3" pageNumber="4">1 slightly longer than antennomere 2. Antennomere 3-5 long-conical. Antennomere 3</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,1127,294,319]" pageId="3" pageNumber="4">longer and narrower than antennomere 2. Antennomere 4 shorter than antennomere</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" pageId="3" pageNumber="4">3. Antennomere 5 subequal to antennomere 4. Antennomeres 6 and 7 oval. Antennomere 6 shorter than antennomere 5. Antennomere 7 slightly shorter and slightly wider than antennomere 6. Antennomere 8 almost rounded, shorter and wider than antennomere 7. Antennal club loose, about 0.5 times as long as antennomeres 18</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,1126,448,472]" pageId="3" pageNumber="4">combined. Antennomere 9 transverse. Antennomere 10 subequal to antennomere 9.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,998,478,503]" pageId="3" pageNumber="4">Antennomere 11 slightly shorter than antennomeres 9 and 10 combined.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" pageId="3" pageNumber="4">Pronotum almost campaniform, 1.5 times as long as wide at apex, 1.4 times as narrow as wide in middle and 1.5 times as narrow as wide at base. Disk weakly flattened, densely punctate. Scutellum small, almost rectangular.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[265,1126,601,625]" pageId="3" pageNumber="4">Elytra 1.7 times as long as wide at base, 1.3 times as long as wide in middle, 2.1</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" pageId="3" pageNumber="4">times as long as wide at apical fourth, 2.8 times as long as pronotum, densely punctate. Humeri slightly flattened. Elytral striae absent. Epipleura narrow.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,697,693,717]" pageId="3" pageNumber="4">Rounded apex of elytra when both together.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[265,1127,724,748]" pageId="3" pageNumber="4">Pre- and postcoxal portions of prosternum short. Procoxal cavities contiguous.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,1127,754,778]" pageId="3" pageNumber="4">Metanepisternum punctate. Metaventrite weakly convex, punctate. Abdomen convex,</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,325,786,809]" pageId="3" pageNumber="4">punctate.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[265,1127,816,840]" pageId="3" pageNumber="4">Procoxae conical. Metacoxae transverse. Trochanters small. Femora thickened,</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" pageId="3" pageNumber="4">lacking teeth. Tibiae almost straight, without costate dorsal margin and two apical spurs. Tarsi long. Tarsomere 1 long-conical. Tarsomere 2 wide-conical. Tarsomere</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,1039,908,932]" pageId="3" pageNumber="4">3 bilobed. Tarsomere 5 elongate. Tarsal claws slightly divergent, with teeth.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[265,1126,938,963]" pageId="3" pageNumber="4">
FEMALE. Body length (without rostrum)
<quantity box="[727,860,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.9" metricValueMin="1.4" pageId="3" pageNumber="4" unit="mm" value="1.65" valueMax="1.9" valueMin="1.4">1.41.9 mm</quantity>
. Rostrum length 0.50.6
</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[227,839,969,993]" pageId="3" pageNumber="4">mm. Rostrum little longer and thicker. Elytra little wider.</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" box="[265,1127,1000,1024]" pageId="3" pageNumber="4">
COMPARISION. The new species is close to
<taxonomicName authority="(Roelofs, 1874)" baseAuthorityName="Roelofs" baseAuthorityYear="1874" box="[756,1068,1000,1024]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="fumigatus">
<emphasis box="[756,772,1000,1023]" italics="true" pageId="3" pageNumber="4">A</emphasis>
.
<emphasis box="[786,891,1000,1024]" italics="true" pageId="3" pageNumber="4">fumigatus</emphasis>
(Roelofs, 1874)
</taxonomicName>
from
</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" pageId="3" pageNumber="4">
<collectingCountry box="[227,326,1030,1054]" name="China" pageId="3" pageNumber="4">East Asia</collectingCountry>
but differs in the thinner and smooth rostrum, narrower pronotum, extended basally basal sclerite of the endophallus, and smaller body sizes. It is similar to
<taxonomicName box="[1103,1127,1061,1085]" pageId="3" pageNumber="4">
<emphasis box="[1103,1119,1061,1084]" italics="true" pageId="3" pageNumber="4">A</emphasis>
.
</taxonomicName>
</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" pageId="3" pageNumber="4">
<emphasis box="[227,310,1092,1115]" italics="true" pageId="3" pageNumber="4">acaciae</emphasis>
Voss, 1935 from
<collectingRegion box="[525,575,1092,1115]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="4">Java</collectingRegion>
, but is distinguished in the longer (considerably longer than head) narrow rostrum, narrow pronotum, wider elytra and narrowed to the apex basal sclerite of the endophallus. From
<taxonomicName authority="Legalov, 2007" authorityName="Legalov" authorityYear="2007" box="[771,1126,1153,1177]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sumatranensis">
<emphasis box="[771,787,1153,1176]" italics="true" pageId="3" pageNumber="4">A</emphasis>
.
<emphasis box="[805,959,1154,1177]" italics="true" pageId="3" pageNumber="4">sumatranensis</emphasis>
Legalov, 2007
</taxonomicName>
</paragraph>
<paragraph blockId="3.[227,1127,264,1330]" pageId="3" pageNumber="4">
from
<collectingRegion box="[288,380,1184,1208]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="4">Sumatra</collectingRegion>
, it is differs in the dark body with yellow procoxae, profemora and protibiae, a rather narrow pronotum, sparse punctate forehead and body covered with sparse hairs.
</paragraph>
</subSubSection>
<subSubSection box="[265,825,1276,1300]" pageId="3" pageNumber="4" type="distribution">
<paragraph blockId="3.[227,1127,264,1330]" box="[265,825,1276,1300]" pageId="3" pageNumber="4">
DISTRIBUTION.
<collectingCountry box="[465,573,1276,1300]" name="Indonesia" pageId="3" pageNumber="4">Indonesia</collectingCountry>
: Siberut Island (
<figureCitation box="[746,812,1276,1300]" captionStart="Fig" captionStartId="4.[265,300,826,848]" captionTargetBox="[247,1107,257,811]" captionTargetId="figure-0@4.[249,1104,260,808]" captionTargetPageId="4" captionText="Fig. 4. Distribution of species of the genus Auletobius from Sunda Islands. star A." figureDoi="http://doi.org/10.5281/zenodo.7164795" httpUri="https://zenodo.org/record/7164795/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[265,799,1306,1330]" pageId="3" pageNumber="4" type="etymology">
<paragraph blockId="3.[227,1127,264,1330]" box="[265,799,1306,1330]" pageId="3" pageNumber="4">ETYMOLOGY. From name of the island Siberut.</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="2CE5BFF1ABCCBBF297723F66782B5D6F" ID-DOI="10.25221/fee.389.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7164791" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665244883012" checkinUser="felipe" docAuthor="Legalov, A. A." docDate="2019" docId="3E6D9136FFA7817D12CF7E85AD211B5C" docLanguage="en" docName="FarEastEntomol.389.1-6.pdf" docOrigin="Far Eastern Entomologist 389" docSource="http://dx.doi.org/10.25221/fee.389.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.2:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Auletobius DESBROCHERS DES LOGES 1869" docType="key" docVersion="4" lastPageNumber="6" masterDocId="C254E94EFFA38178122C7D28A9441A20" masterDocTitle="A new species of the genus Auletobius Desbrochers des Loges, 1869 (Coleoptera: Rhynchitidae) from Indonesia" masterLastPageNumber="6" masterPageNumber="1" pageNumber="5" updateTime="1699368761949" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4A3D946A23C1547A3D0D11C000CF4424">A new species of the genus Auletobius Desbrochers des Loges, 1869 (Coleoptera: Rhynchitidae) from Indonesia</mods:title>
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<subSubSection id="FEDE73ABFFA7817C12CF7E85AD2719E5" box="[227,1123,941,965]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="B67B2020FFA7817C12CF7E85AD2719E5" blockId="4.[227,1123,941,965]" box="[227,1123,941,965]" pageId="4" pageNumber="5">
<heading id="ED33974CFFA7817C12CF7E85AD2719E5" box="[227,1123,941,965]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="5" reason="2">
<emphasis id="84B0FC32FFA7817C12CF7E85AD2719E5" bold="true" box="[227,1123,941,965]" pageId="4" pageNumber="5">
Key to species of the genus
<taxonomicName id="71C45BA3FFA7817C10347E85ABCB19E5" ID-CoL="8HCX7" authorityName="DESBROCHERS DES LOGES" authorityYear="1869" box="[536,655,941,965]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="84B0FC32FFA7817C10347E85ABCB19E5" bold="true" box="[536,655,941,965]" italics="true" pageId="4" pageNumber="5">Auletobius</emphasis>
</taxonomicName>
from Sunda Isl. and neighboring regions
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="B67B2020FFA7817C12CF7EC2AD1F1E22" blockId="4.[227,1127,1002,1609]" box="[227,1115,1002,1026]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF7EC2AD1F1E22" box="[227,1115,1002,1026]" pageId="4" pageNumber="5">1. Body yellow or yellow-brown ............................................................................. 2</keyLead>
</paragraph>
<paragraph id="B67B2020FFA7817C12CF7921AD1D1E01" blockId="4.[227,1127,1002,1609]" box="[227,1113,1033,1057]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF7921AD1D1E01" box="[227,1113,1033,1057]" pageId="4" pageNumber="5"> Body black or brown ........................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D303885FFA7817C12CF7900AD1E1E7E" pageId="4" pageNumber="5">
<paragraph id="B67B2020FFA7817C12CF7900AD1D1E60" blockId="4.[227,1127,1002,1609]" box="[227,1113,1063,1088]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF7900AD1D1E60" box="[227,1113,1063,1088]" pageId="4" pageNumber="5">
2. Body yellow.
<collectingRegion id="7400EEC2FFA7817C13B8790FAB551E1F" box="[404,529,1063,1087]" country="Indonesia" name="Kalimantan Utara" pageId="4" pageNumber="5">Kalimantan</collectingRegion>
.....................................
<taxonomicName id="71C45BA3FFA7817C11387900AD1D1E60" authority="Legalov, 2018" authorityName="Legalov" authorityYear="2018" box="[788,1113,1063,1088]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="crockerensis">
<emphasis id="84B0FC32FFA7817C11387900AA601E1F" box="[788,804,1064,1087]" italics="true" pageId="4" pageNumber="5">A</emphasis>
.
<emphasis id="84B0FC32FFA7817C111E790FAAFD1E1F" box="[818,953,1063,1087]" italics="true" pageId="4" pageNumber="5">crockerensis</emphasis>
Legalov, 2018
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B67B2020FFA7817C12CF796EAD1E1E7E" blockId="4.[227,1127,1002,1609]" box="[227,1114,1094,1118]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF796EAD1E1E7E" box="[227,1114,1094,1118]" pageId="4" pageNumber="5"> Body yellow-brown .............................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D303885FFA7817C12CF794DAD1E1EBC" pageId="4" pageNumber="5">
<paragraph id="B67B2020FFA7817C12CF794DAD1A1E5D" blockId="4.[227,1127,1002,1609]" box="[227,1118,1125,1149]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF794DAD1A1E5D" box="[227,1118,1125,1149]" pageId="4" pageNumber="5">
3. Forehead densely punctate.
<collectingRegion id="7400EEC2FFA7817C1004794DABC51E5D" box="[552,641,1125,1149]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="5">Sumatra</collectingRegion>
..................
<taxonomicName id="71C45BA3FFA7817C112A794DAD1A1E5D" ID-CoL="JS2Q" authority="Legalov, 2007" authorityName="Legalov" authorityYear="2007" box="[774,1118,1125,1149]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sumatranensis">
<emphasis id="84B0FC32FFA7817C112A794DAA521E5C" box="[774,790,1125,1148]" italics="true" pageId="4" pageNumber="5">A</emphasis>
.
<emphasis id="84B0FC32FFA7817C110F794DAAF91E5C" box="[803,957,1125,1148]" italics="true" pageId="4" pageNumber="5">sumatranensis</emphasis>
Legalov, 2007
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B67B2020FFA7817C12CF79ACAD1E1EBC" blockId="4.[227,1127,1002,1609]" box="[227,1114,1155,1180]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF79ACAD1E1EBC" box="[227,1114,1155,1180]" pageId="4" pageNumber="5">
Forehead sparsely punctate.
<collectingRegion id="7400EEC2FFA7817C100B79ABABE01EBB" box="[551,676,1155,1179]" country="Indonesia" name="Kalimantan Utara" pageId="4" pageNumber="5">Kalimantan</collectingRegion>
............
<taxonomicName id="71C45BA3FFA7817C112C79ACAD1E1EBC" ID-CoL="JRZY" authority="Legalov, 2007" authorityName="Legalov" authorityYear="2007" box="[768,1114,1155,1180]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sandakanensis">
<emphasis id="84B0FC32FFA7817C112C79ACAA541EBB" box="[768,784,1156,1179]" italics="true" pageId="4" pageNumber="5">A</emphasis>
.
<emphasis id="84B0FC32FFA7817C113279ABAAFD1EBB" box="[798,953,1155,1179]" italics="true" pageId="4" pageNumber="5">sandakanensis</emphasis>
Legalov, 2007
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D303885FFA7817C12CF798AAD1A1EF8" pageId="4" pageNumber="5">
<paragraph id="B67B2020FFA7817C12CF798AAD181E9A" blockId="4.[227,1127,1002,1609]" box="[227,1116,1186,1210]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF798AAD181E9A" box="[227,1116,1186,1210]" pageId="4" pageNumber="5">4 Procoxae, profemora and protibiae yellow ........................................................... 5</keyLead>
</paragraph>
<paragraph id="B67B2020FFA7817C12CF79E9AD1A1EF8" blockId="4.[227,1127,1002,1609]" box="[227,1118,1217,1241]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF79E9AD1A1EF8" box="[227,1118,1217,1241]" pageId="4" pageNumber="5"> All legs brown ....................................................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D303885FFA7817C12CF79F7AD251F15" pageId="4" pageNumber="5">
<paragraph id="B67B2020FFA7817C12CF79F7AD1F1F36" blockId="4.[227,1127,1002,1609]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF79F7AD1F1F36" pageId="4" pageNumber="5">
5. Profemora, base of meso- and metafemora, tibiae and tarsi yellow.
<collectingCountry id="CED360B0FFA7817C11FF79F7AD0F1ED7" box="[979,1099,1247,1271]" name="Philippines" pageId="4" pageNumber="5">Philippines</collectingCountry>
... .........................................................................
<taxonomicName id="71C45BA3FFA7817C10DA79D6AD1F1F36" authority="Legalov, 2018" authorityName="Legalov" authorityYear="2018" box="[758,1115,1278,1302]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kapataganensis">
<emphasis id="84B0FC32FFA7817C10DA79D6AA421F35" box="[758,774,1278,1301]" italics="true" pageId="4" pageNumber="5">A</emphasis>
.
<emphasis id="84B0FC32FFA7817C113879D6AAFE1F36" box="[788,954,1278,1302]" italics="true" pageId="4" pageNumber="5">kapataganensis</emphasis>
Legalov, 2018
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B67B2020FFA7817C12CF7835AD251F15" blockId="4.[227,1127,1002,1609]" box="[227,1121,1309,1333]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF7835AD251F15" box="[227,1121,1309,1333]" pageId="4" pageNumber="5"> Only procoxae, profemora and protibiae yellow ................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D303885FFA7817C12CF7813AD181F8F" pageId="4" pageNumber="5">
<paragraph id="B67B2020FFA7817C12CF7813AD241F52" blockId="4.[227,1127,1002,1609]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF7813AD241F52" pageId="4" pageNumber="5">
6. Rostrum short, equal in length to head, thicker. Pronotum wider. Elytra narrower. Basal sclerite widened at apex.
<collectingRegion id="7400EEC2FFA7817C10767872ABCD1F51" box="[602,649,1370,1393]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="5">Java</collectingRegion>
.................................
<taxonomicName id="71C45BA3FFA7817C115E7872AD241F52" ID-CoL="JRSW" authority="Voss, 1935" authorityName="Voss" authorityYear="1935" box="[882,1120,1370,1394]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="acaciae">
<emphasis id="84B0FC32FFA7817C115E7872AAC61F51" box="[882,898,1370,1393]" italics="true" pageId="4" pageNumber="5">A</emphasis>
.
<emphasis id="84B0FC32FFA7817C11A37873AAA61F52" box="[911,994,1371,1394]" italics="true" pageId="4" pageNumber="5">acaciae</emphasis>
Voss, 1935
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B67B2020FFA7817C12CF7851AD181F8F" blockId="4.[227,1127,1002,1609]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF7851AD181F8F" pageId="4" pageNumber="5">
Rostrum longer, much longer than head, narrow. Pronotum narrow. Elytra wider. Basal sclerite narrowed at apex ...........................
<taxonomicName id="71C45BA3FFA7817C113B78B0AD511F8F" authority="Legalov" authorityName="Legalov" authorityYear="2019" box="[791,1045,1431,1455]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="siberutensis" status="sp. nov.">
<emphasis id="84B0FC32FFA7817C113B78B0AAF01F8F" box="[791,948,1431,1455]" italics="true" pageId="4" pageNumber="5">A. siberutensis</emphasis>
Legalov
</taxonomicName>
,
<emphasis id="84B0FC32FFA7817C160C78BFAD181F8F" bold="true" box="[1056,1116,1431,1455]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="9F834149FFA7817C160C78BFAD181F8F" box="[1056,1116,1431,1455]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D303885FFA7817C12CF789EAD1E1FCD" pageId="4" pageNumber="5">
<paragraph id="B67B2020FFA7817C12CF789EAD1F1FEE" blockId="4.[227,1127,1002,1609]" box="[227,1115,1462,1486]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF789EAD1F1FEE" box="[227,1115,1462,1486]" pageId="4" pageNumber="5">
7. Pronotum cordiform.
<collectingRegion id="7400EEC2FFA7817C13C9789EAB511FED" box="[485,533,1462,1485]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="5">Java</collectingRegion>
....................................
<taxonomicName id="71C45BA3FFA7817C113E789EAD1F1FEE" ID-CoL="JS2D" authority="Voss, 1935" authorityName="Voss" authorityYear="1935" box="[786,1115,1462,1486]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="subcordaticollis">
<emphasis id="84B0FC32FFA7817C113E789EAA661FED" box="[786,802,1462,1485]" italics="true" pageId="4" pageNumber="5">A</emphasis>
.
<emphasis id="84B0FC32FFA7817C111C789EAA981FEE" box="[816,988,1462,1486]" italics="true" pageId="4" pageNumber="5">subcordaticollis</emphasis>
Voss, 1935
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B67B2020FFA7817C12CF78FDAD1E1FCD" blockId="4.[227,1127,1002,1609]" box="[227,1114,1493,1517]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF78FDAD1E1FCD" box="[227,1114,1493,1517]" pageId="4" pageNumber="5"> Pronotum campaniform ........................................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D303885FFA7817C12CF78DBAD251C69" pageId="4" pageNumber="5">
<paragraph id="B67B2020FFA7817C12CF78DBAD191C0A" blockId="4.[227,1127,1002,1609]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF78DBAD191C0A" pageId="4" pageNumber="5">
8. Elytra more elongated. Body brown.
<collectingCountry id="CED360B0FFA7817C10BD78DBABB91C2B" box="[657,765,1523,1547]" name="Singapore" pageId="4" pageNumber="5">Singapore</collectingCountry>
..................................................... ............................................................................
<taxonomicName id="71C45BA3FFA7817C11267B3AAD191C0A" ID-CoL="JS26" authority="Legalov, 2007" authorityName="Legalov" authorityYear="2007" box="[778,1117,1554,1578]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="singaporensis">
<emphasis id="84B0FC32FFA7817C11267B3AAA5E1C09" box="[778,794,1554,1577]" italics="true" pageId="4" pageNumber="5">A</emphasis>
.
<emphasis id="84B0FC32FFA7817C11047B3BAAF81C0A" box="[808,956,1555,1578]" italics="true" pageId="4" pageNumber="5">singaporensis</emphasis>
Legalov, 2007
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B67B2020FFA7817C12CF7B19AD251C69" blockId="4.[227,1127,1002,1609]" box="[227,1121,1585,1609]" pageId="4" pageNumber="5">
<keyLead id="0D358315FFA7817C12CF7B19AD251C69" box="[227,1121,1585,1609]" pageId="4" pageNumber="5"> Elytra wider. Body black and brown ..................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="0D303885FFA6817D12CF7C20AD211B5C" pageId="5" pageNumber="6">
<paragraph id="B67B2020FFA6817D12CF7C20AD201B1F" blockId="5.[227,1127,264,380]" pageId="5" pageNumber="6">
<keyLead id="0D358315FFA6817D12CF7C20AD201B1F" pageId="5" pageNumber="6">
9. Rostrum almost equal to head and pronotum combined.
<collectingRegion id="7400EEC2FFA6817D114D7C20AAD41B3F" box="[865,912,264,287]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="6">Java</collectingRegion>
............................... .........................................................................................
<taxonomicName id="71C45BA3FFA6817D114D7C0FAD201B1F" ID-CoL="JRTS" authority="Voss, 1940" authorityName="Voss" authorityYear="1940" box="[865,1124,294,319]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="blawanus">
<emphasis id="84B0FC32FFA6817D114D7C0FAA351B1E" box="[865,881,295,318]" italics="true" pageId="5" pageNumber="6">A</emphasis>
.
<emphasis id="84B0FC32FFA6817D11527C0EAAA01B1E" box="[894,996,294,318]" italics="true" pageId="5" pageNumber="6">blawanus</emphasis>
Voss, 1940
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B67B2020FFA6817D12CF7C6EAD211B5C" blockId="5.[227,1127,264,380]" pageId="5" pageNumber="6">
<keyLead id="0D358315FFA6817D12CF7C6EAD211B5C" pageId="5" pageNumber="6">
Rostrum distinctly shorter than head and pronotum combined.
<collectingCountry id="CED360B0FFA6817D11BD7C6DAD4D1B7D" box="[913,1033,325,349]" name="Philippines" pageId="5" pageNumber="6">Philippines</collectingCountry>
............. ..............................................................................................
<taxonomicName id="71C45BA3FFA6817D11AE7C4CAD211B5C" ID-CoL="JRYT" authority="Voss, 1922" authorityName="Voss" authorityYear="1922" box="[898,1125,356,380]" class="Insecta" family="Attelabidae" genus="Auletobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="84B0FC32FFA6817D11AE7C4CAAD61B5B" box="[898,914,356,379]" italics="true" pageId="5" pageNumber="6">A</emphasis>
.
<emphasis id="84B0FC32FFA6817D118C7C4DAAA21B5C" box="[928,998,357,380]" italics="true" pageId="5" pageNumber="6">ovatus</emphasis>
Voss, 1922
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>