diff --git a/data/03/85/87/038587945A52FFFC10F6FF78FB59FF3B.xml b/data/03/85/87/038587945A52FFFC10F6FF78FB59FF3B.xml new file mode 100644 index 00000000000..1d038031683 --- /dev/null +++ b/data/03/85/87/038587945A52FFFC10F6FF78FB59FF3B.xml @@ -0,0 +1,377 @@ + + + +A new species of Archithosia Birket-Smith, 1965 in the subgenus Eurythosia Krüger, 2015 from Uganda (Lepidoptera: Erebidae: Arctiinae: Lithosiini) + + + +Author + +Durante, Antonio +Museo di Storia naturale del Salento. S. P. Calimera-Borgagne, Km 1, 73021 Calimera, Italy + + + +Author + +Potenza, Luigi +Museo di Storia naturale del Salento. S. P. Calimera-Borgagne, Km 1, 73021 Calimera, Italy + + + +Author + +Pellegrino, Gabriele +Museo di Storia naturale del Salento. S. P. Calimera-Borgagne, Km 1, 73021 Calimera, Italy + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +356 +368 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.4 + +journal article +10.11646/zootaxa.5492.3.4 +1175-5326 +13234883 +54F5ECF4-C148-4AB3-9F2C-F3B066B55825 + + + + + + + +Archithosia henricus +Durante, Potenza & Pellegrino + +n. sp. +( +Figs. 5 +, +6 +, +7 +, +8 +) + + + + +https://zoobank.org/NomenclaturalActs/ +740DB914-0CDB-468B-8423-1E70A9D86C45 + + + + + + +Holotype + +: +♂ +, +UGANDA +, +Kibale +forest, m 1500, +0°33’40” N +30°21’24” E +, 19/ + +24-10-2014 + +, +Durante +leg., g. sl. n. 1159 +MAD +; in coll. +MSNS +. + + + + + +Paratypes + +: +1 ♂ +, +UGANDA +, same data as the holotype, g. sl. n. 1175 +MAD + +; + +1 ♀ +, +UGANDA +, same data as the holotype, g. sl. n. 1176 +MAD + +; in coll. +MSNS +. + + + + +Diagnosis +. The species of the subgenus + +Eurythosia + +are all very similar in terms of their external features, so definite discrimination is impossible without examination of the genitalia (see Discussion). + + + +FIGURE 5. + +Archithosia henricus + + +n. sp. + +, holotype, male, gen sl. n. 1159 MAD. + + + + +FIGURE 6. + +Archithosia henricus + + +n. sp. + +, paratype, female, gen. sl. n. 1176 MAD. + + + +The male genitalia of + +Archithosia henricus + +n. sp. +are closely related to those of + +Archithosia makomensis + +, from which they differ in (i) the length of the ala valvae distal transverse process, which is about 1/4 of the entire length of the ala valvae in + +henricus + +n. sp. +and about 1/ +3 in + +makomensis + +; (ii) the width of the supravalva, which in + +henricus + +n. sp. +is about half what it is in + +makomensis + +; (iii) the dorsal sclerotisation at the base of the supravalva (i.e. proximal half of costa), which is more accentuated in + +henricus + +n. sp. +than in + +makomensis + +; the valvellae, which are sturdier in + +henricus + +n. sp. +than + +makomensis + +and are directed outwards in the former and are mesad or directed inwards in the latter. The sturdy process of the supravalva in + +angulifascia + +and that of the ala valvae in + +kinuthiae + +easily distinguish these species from both + +henricus + +n. sp. +and + +derelicta + +due to absence of the distal process of the ala valvae in the latter. In the female genitalia, the oblong shape of the corpus bursae in + +angulifascia + +and + +kinuthiae + +is distinctive (roundish in + +henricus + +n. sp. +). In + +makomensis + +the V-shaped (convex) lamella antevaginalis and the star-shaped signum are diagnostic (arched, concave lamella antevaginalis and round, cup-shaped signum in + +henricus + +n. sp. +). + + + + +Description +. ( +Figs. 5 +, +6 +). Wingspan +♂ +36–37 mm +; +♀ +35. +5 mm +. Head with vertex beige and frontoclypeus from dark beige to almost black with margins darker; antennae ciliated (cilia denser in male), dorsally same colour as vertex, ventrally unpigmented; labial palpi porrect, not extending beyond frons, same colour as frontoclypeus, with distal segment always blackish. + +Patagia, tegulae and thorax same colour as forewings, sometimes slightly darker; patagia and tegulae somewhat darker in the middle. Legs same colour as thorax but forelegs darker, mid- and hindlegs internally lighter; spurs formula 0-2-4. Forewing upperside ground colour brown-dusted beige with angled brown band from middle of costa towards space between M1 and M3, afterwards turning to inner margin almost in middle. Forewing underside unmarked, lighter beige than upperside. Hindwing upperside unmarked, ground colour light beige; underside same colour as upperside. + +Abdomen slightly lighter than thorax. A2 sternum apodemes ( +Fig. 8B +) long (plesiomorphic character according to +Bendib & Minet (1999: 243) +, but see also Holloway (2001: 281)). + +Female seventh abdominal sternite about 1/3 shorter than sixth; seventh tergite slightly longer than sixth. Eighth abdominal sternite swollen, about 1/2 as long as sixth sternite; eighth tergite about 1/5 as long as seventh tergite, and completely fused to it (no trace of intersegmental membrane). + +Genitalia. +♂ +( +Fig. 7 +). Uncus (UC) falcate, larger at base, ending in sharp hook ( +Fig. 7B +); tegumen (TE) triangular in dorsal view; tuba analis (TA) short, membranous with very thin rod-like scaphium (SC) along its basal 4/5 ( +Fig. 7B +); vinculum (VI) weak, large-u-shaped (ventro-caudal view), not forming evident saccus; juxta (JU) arched ( +Fig. 7C +); valvellae (VA) sturdy, thorn-like, directed outwards, firmly connected to bases of ala valvae; manica penis (MP) membranous, with membranous thin tape-like dorsal transtilla (TR) connecting two plicae centripetalis ( +Fig. 7D +). + + +Ala valvae (AV) rectangularish, fairly sclerotised, with distal conical process (CP) covered in small spines ( +Fig. 7A +) and ending in sturdy thorn; supravalva (SV) only slightly sclerotised, digitiform, exceeding ala valvae by about ¼ of its total length, bearing a swollen process (SP in +Fig. 7A +) arising from dorsal part of basis valvae and extending along costa up to middle of total length. + + +Aedeagus ( +Figs. 7E, 7F +) almost as long as ala valvae, tubular, fairly sclerotised, with dorsal entrance of ductus ejaculatorius (DE) and short coecum penis (CE, about 1/6 of total length of aedeagus); vesica with distal main lobe (ML), bearing sturdy thorn-like cornutus (CO), second small lobe (SL) with right side covered in small granicula, third dorsal medium lobe (DL), membranous and dome-shaped, and fourth small lobe (LL) on left, also covered in granicula ( +Fig. 7F +). + + +♀ +( +Fig.8 +). Seventh sternite very short,quite swollen, slightly sclerotised in its central portion;lamella antevaginalis (LA) made up of strongly sclerotised arch, prolonged laterally into two rectangularish flat sclerotisations (LS) partly entering, funnel-like, ostium bursae. Dorsal portion of ostium completely membranous. Eighth sternite swollen, sclerotised, bent over sterigma and seventh sternum, covering ostium; central caudal portion (CP) membranous and scobinate with tiny spines. Papillae anales (PA) smoothly squarish (in pictures appearing to taper caudally due to rolling of external caudal corners); anterior apophyses slightly shorter than papillae anales; posterior apophyses almost twice as long as anterior apophyses; dorsal pheromone glands (PG; +Fig. 8C +) unpaired, with one caudal opening at intersegmental membrane between A8 and A9, anteriorly deeply bifurcated into two triangularish lobes as long as papillae anales. + + +Ductus bursae (DB) half as long as corpus bursae, quite large, swollen leftward, with dorso-caudal small dome-shaped appendix (DA; +Fig. 8D +), from which ductus seminalis (DS) arises (clearly visible in dorsal view). Corpus bursae (CB) spheroid, membranous, internally covered in tiny spines; signum (SI) roundish, deeply concave, strongly sclerotised, scobinate with small spines. + + + + +FIGURE 7. + +Archithosia henricus + + +n. sp. + +, male genitalia. +A +. Genitalia, ventral view. +B +. Uncus, lateral view. +C +. Juxta, ventral view. +D +. Transtilla, dorsal view. +E +. Aedeagus, right side. +F +. Aedeagus, left side. AV ala valvae; CE coecum penis; CO cornutus; CP conical process of ala valvae; DE ductus ejaculatorius entrance; DL dorsal medium lobe of vesica; JU juxta; LL small left lateral lobe of vesica; ML main lobe of vesica; MP manica penis; SC scaphium; SL small right lateral lobe of vesica; SP swollen process of supravalva; SV supravalva; TA tuba analis; TE tegumen; TR transtilla; UC uncus; VA valvella; VI vinculum. + + + + +FIGURE 8. + +Archithosia henricus + +n. sp. +, female genitalia. +A +. Genitalia, ventral view. +B +. Apodemes. +C +. papillae anales and dorsal pheromone glands, dorsal view. +D +. Appendix and ductus bursae, dorsal view. AP apodemes of second sternite; CB corpus bursae; CP membranous caudal portion of ostium bursae; DA dome-shaped appendix; DB ductus bursae; DS ductus seminalis; LA lamella antevaginalis; LS lateral extensions of lamella antevaginalis; PA papillae anales; PG dorsal pheromone glands; SI signum; VII st seventh sternite; VIII st eighth sternite. + + + + +Etymology. +The authors wish to dedicate the new species to the young researcher of the MSNS, Enrico Panzera, who recently passed away suddenly at the age of 30. Noun in apposition (ICZN 11.9.1.2). + + + + \ No newline at end of file diff --git a/data/03/9C/A1/039CA1771B4EA808FF26FD0B8FF4B9EB.xml b/data/03/9C/A1/039CA1771B4EA808FF26FD0B8FF4B9EB.xml new file mode 100644 index 00000000000..cf1d509052e --- /dev/null +++ b/data/03/9C/A1/039CA1771B4EA808FF26FD0B8FF4B9EB.xml @@ -0,0 +1,110 @@ + + + +A new intriguing species of the genus Pseudachorutella Stach, 1949 (Collembola: Neanuridae: Pseudachorutinae) from Kyrgyzstan with the supplementation of generic definition and the description of the first instar larva + + + +Author + +Stryczek, Beata +0009-0005-4460-5700 +University of Wrocław, Faculty of Biological Sciences, Department of Invertebrate Biology, Evolution and Conservation, Przybyszewskiego 65, 51 - 148 Wrocław, Poland +317366@uwr.edu.pl + + + +Author + +Smolis, Adrian +University of Wrocław, Faculty of Biological Sciences, Department of Invertebrate Biology, Evolution and Conservation, Przybyszewskiego 65, 51 - 148 Wrocław, Poland + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +421 +430 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.8 + +journal article +10.11646/zootaxa.5492.3.8 +1175-5326 +13234985 +CB539C52-9416-450D-AF15-70092ADB602B + + + + + + +Genus + +Pseudachorutella +Stach, 1949 + + + + + + + + + + +Pseudachorutella +Stach, 1949: 79 + + +. + + + + + +Type +species: + +Pseudachorutes asigillatus +Börner, 1901 + +, by subsequent designation by +Stach 1949: 80 +. + + + + +New diagnosis. +Colour in alive bluish grey, blue violet or dark blue. +Pseudachorutinae +of medium to large size, length of adults usually more than +1 mm +. Postantennal organ (PAO) absent. 8+8 dark pigmented eyes. Dorsal tubercles or reticulations absent. Ant. III and IV fused dorsally, sensorial equipment of dorsal side of Ant. IV with 6–8 cylindrical sensilla S. Ms of Ant. IV developed as microsensillum. Ventral side of ant. III usually with numerous modified setae (ventral “file”). Buccal cone usually elongated, line of labial setae C–D parallel to A–B. Moth part present but reduced: maxilla usually styliform and with or without apical hook/hooks, mandible slender or rarely strong, with 2–8 teeth. Dens with 5–7 setae, mucro present or rarely absent. Male organ present or absent. Tibiotarsi without or rarely with capitate setae. Claw with inner tooth or teeth, sometimes lateral teeth also present. Empodial appendage absent. + + + + +Remarks +. Since, the new species described here has distinct and unknown characters within the genus, we decided to modify its diagnosis. + + + + \ No newline at end of file diff --git a/data/03/9C/A1/039CA1771B4EA80FFF26FA968B32BA44.xml b/data/03/9C/A1/039CA1771B4EA80FFF26FA968B32BA44.xml new file mode 100644 index 00000000000..832b5c88f4d --- /dev/null +++ b/data/03/9C/A1/039CA1771B4EA80FFF26FA968B32BA44.xml @@ -0,0 +1,388 @@ + + + +A new intriguing species of the genus Pseudachorutella Stach, 1949 (Collembola: Neanuridae: Pseudachorutinae) from Kyrgyzstan with the supplementation of generic definition and the description of the first instar larva + + + +Author + +Stryczek, Beata +0009-0005-4460-5700 +University of Wrocław, Faculty of Biological Sciences, Department of Invertebrate Biology, Evolution and Conservation, Przybyszewskiego 65, 51 - 148 Wrocław, Poland +317366@uwr.edu.pl + + + +Author + +Smolis, Adrian +University of Wrocław, Faculty of Biological Sciences, Department of Invertebrate Biology, Evolution and Conservation, Przybyszewskiego 65, 51 - 148 Wrocław, Poland + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +421 +430 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.8 + +journal article +10.11646/zootaxa.5492.3.8 +1175-5326 +13234985 +CB539C52-9416-450D-AF15-70092ADB602B + + + + + + + +Pseudachorutella intricata + +sp. nov. + + + + + + +Figs 1–17 + + + + +Type material. + +Holotype +, adult female on slide, under bark of spruce trees, +Dzhety Oguz +, +Karakol Region +, Issyk- +Kul District +, +Kyrgyzstan +, + +11.VI.2006 + +, leg. +R +. +J. Pomorski + +. + +Paratypes +: +2 juveniles +and +4 adult +females, under bark of spruce trees, in decaying wood, same data as holotype + +; + +2 adult +females, same locality as holotype + +, + +mosses on waterfall, + +25.VII.2005 + +, leg. +R +. +J. Pomorski + +. + + + + +Diagnosis +. A large-sized species. Ant. IV with eight cylindrical sensilla S in adults. Labrum with 5/4,2,4 setae, labium with 12–13 ordinary setae and subapical spine, labial organites absent. Maxillae styliform without apical hooks, mandibles with two basal and three small apical teeth. Dorsal chaetotaxy of abdomen characterized by the presence of supplementary setae, especially in dorso-external parts of abdominal terga. Abd. V lacking setae p +2 +. Furca reduced, dens short and globular in shape with 5–6 + 5–6 setae, mucro absent. + + + + +Description of the first instar larva. +Body length: 1.45 mm Colour of the body bluish-grey. Granulation relatively coarse and uniform. Dorsal organ of Th. I present ( +Fig. 6 +). + + +Antennae equal to head. Ant. I with 7 chaetae, Ant. II with 12–13 chaetae. Chaetotaxy of Ant. III–IV as in +Figs 1, 2 +. AOIII with two small internal curved sensilla, one ventral microsensillum and two cylindrical guard sensilla, ventral one longer than dorsal ( +Figs 1–2 +). Ant. IV with trilobed apical vesicle, subapical organite, microsensillum and chaeta i. Ant. IV with two cylindrical sensilla S and six sensilla mou, without ventral “file”. + + +8 + 8 ocelli dark pigmented. Buccal cone elongated, labral sclerifications ogival ( +Figs 5, 7 +). Labium with 12 + 12 setae ( +Fig. 5 +). Maxilla and mandible as in adults. Ventral side of head with 6+6 setae Vi ( +Fig. 5 +). + + +Dorsal chaetotaxy as in +Fig. 6 +, setae relatively short and smooth differentiated into macro- and mesosetae, body sensilla slightly shorter than ordinary setae, fine and smooth. Sensillar formula: 022/11111. Microsensillum on Th. II present. Head with dorsal setae d +1 +present and a +0 +absent ( +Fig. 6 +). Th. II with setae a +2 +present and m +3 +absent. Th. III with setae a +2 +absent. Dorsal side of body without additional setae, plurichaetosis absent. + + + +FIGURES 1–4. + +Pseudachorutella intricata + + +sp. nov +. + +: 1, dorsal chaetotaxy of Ant. III–IV (I instar larva); 2, ventral chaetotaxy of Ant. III–IV (I instar larva); 3, dorsal chaetotaxy of Ant. III–IV (adult); 4, ventral chaetotaxy of Ant. IV (adult). + + + + +FIGURES 5–8. + +Pseudachorutella intricata + + +sp. nov +. + +(I instar larva): 5, labium and group Vi; 6, dorsal chaetotaxy of head, thorax and abdomen; 7, labrum; 8, ventral chaetotaxy of abdomen. + + + + +FIGURES 9–13. + +Pseudachorutella intricata + + +sp. nov +. + +(adult): 9, labium and group Vi; 10, maxilla; 11, mandible; 12, mandible (front view); 13, retinaculum and furca. + + + + +FIGURES 14–17. + +Pseudachorutella intricata + + +sp. nov +. + +(adult): 14, dorsal chaetotaxy of head and Th.I–II; 15, dorsal chaetotaxy of Abd. III–VI; 16, ventral chaetotaxy of abdomen; 17, tibiotarsus and claw of leg I. + + +Thoracic sterna without setae. + +Ventral chaetotaxy of abdomen as in +Fig. 8 +. Ventral tube with 4 + 4 setae. Each anal valve with 2 small setae hr. Retinaculum with 3 + 3 teeth, furca as in +Fig. 8 +. + +Tibiotarsi I, II, III with 19, 19, 18 setae respectively, seta M present. Femora I, II, III with 13, 12, 11 setae respectively, trochanters with 6 setae each, coxae I, II, III with 3, 8, 8 setae, subcoxae 2 of legs I, II and III with 0, 2 and 2 setae, subcoxae 1 of legs I, II and III with 1, 2 and 2 setae. Claws with one inner tooth and without lateral teeth. + +Description of adults. +Body length: 2.85 mm–3.45 mm ( +holotype +: 3.25 mm). Colour of the body dark bluish-grey. Granulation relatively coarse and uniform. + + +Antennae equal to head. Ant. I with 7 chaetae, Ant. II with 12–13 chaetae. Chaetotaxy of Ant. III–IV as in +Figs 3, 4 +. AOIII with two small internal curved sensilla, one ventral microsensillum and two cylindrical guard sensilla, ventral one longer than dorsal. Ant. IV with eight cylindrical sensilla S and numerous sensilla mou ( +Fig. 3 +), without ventral “file” ( +Fig. 4 +). + + +8 + 8 ocelli dark pigmented. Buccal cone elongated, labral sclerifications ogival ( +Fig. 9 +). Labium 13+13 setae, additional setae p +1 +present ( +Fig. 9 +). Maxilla and mandible as in +Figs 10–12 +. Ventral side of head with 6+6 setae Vi ( +Fig. 9 +). + + +Dorsal chaetotaxy as in +Figs 14–15 +, setae relatively short and smooth not differentiated into macro- and mesosetae, body sensilla slightly shorter than ordinary setae, fine and smooth. Sensillar formula: 022/11111. Microsensillum on Th. II present. Head with dorsal setae d +1 +present and a +0 +absent ( +Fig. 14 +). Th. II with setae a +2 +and m +1 +present. Th. III with setae a +2 +absent. Dorsal side of body with some additional setae, moderate plurichaetosis present especially in groups De ( +Fig. 15 +). + + +Thoracic sterna without setae. Ventral chaetotaxy of abdomen as in +Fig. 16 +. Moderate plurichaetosis present. Ventral tube with 8 + 8 setae ( +Fig. 16 +). Each anal valve with 2 small setae hr. Retinaculum with 3 + 3 teeth ( +Fig. 13 +). Furca as in +Figs 13 +and +16 +. + + +Tibiotarsi I, II, III with 19, 19, 18 setae respectively, seta M present ( +Fig. 17 +). Femora I, II, III with 13, 12, 11 setae respectively, trochanters with 6 setae each, coxae I, II, III with 3, 8, 8 setae, subcoxae 2 of legs I, II and III with 0, 2 and 2 setae, subcoxae 1 of legs I, II and III with 1, 2 and 2 setae. Claws with one inner tooth and without lateral teeth ( +Fig. 17 +). + + +Note. +The comparison of morphology of the first instar larva and adults of + +Pseudachorutella intricata + + +sp. nov. + +provides important data in the context of the taxonomy of the analyzed genus and even the subfamily. In general, the chaetotaxy of the first instar larva, both the antennae and other parts of the body, is highly incomplete compared to the adults. For example, antennal segment IV the first instar larva lacks six sensilla S and many sensilla mou. Similarly, the dorsal chaetoataxy of thorax and abdomen of this stage is devoid of numerous setae found in adults. Thus, during development, a transition from a non plurichaetotic to a moderate plurichaetotic form can be observed. Therefore, to avoid incorrect taxonomic decisions, the description of new species of the genus + +Pseudachorutella + +should be based on material containing adult stages. Although knowledge of first-instar larval morphology is of less importance for generic taxonomy, it may be helpful in classification and phylogenetic considerations within subfamily and family (see Discussion). + + + + +Etymology. +The name of the new species referring to its intricate characters i.e. strong reduction of furca, unknown in other members of the genus. + + + + +Remarks. + +Pseudachorutella intricata + + +sp. nov. + +is unique, distinguished from other species of the genus by presence of strong reduction of furca i.e. dens shortened and globular in shape, mucro absent. Nevertheless, the new species appears to be most similar to + +P. plurichaetosa +Babenko & Kuznetsova, 2024 + +, species recently described from the Caucasus ( +Babenko & Kuznetsova 2024 +). Besides these unique characters of the new species, they can additionally be distinguished by the following features: ventral “file” on Ant. IV (absent in + +P. intricata + + +sp. nov +. + +, present in + +P. plurichaetosa + +), apical hooks on maxillae (absent in + +P. intricata + + +sp. nov +. + +, present in + +P. plurichaetosa + +), labral formula (5/4,2,4 setae in + +P. intricata + + +sp. nov +. + +, 4/2,3,3,4 in + +P. plurichaetosa + +) and lateral teeth on claw (absent in + +P. intricata + + +sp. nov +. + +, present in + +P. plurichaetosa + +). + +In addition, the differences between all known Palearctic species of the genus are provided in the key. + + + \ No newline at end of file diff --git a/data/03/A9/87/03A987C2FFD0DA49FF23458AFAC73C1C.xml b/data/03/A9/87/03A987C2FFD0DA49FF23458AFAC73C1C.xml new file mode 100644 index 00000000000..27156fa0155 --- /dev/null +++ b/data/03/A9/87/03A987C2FFD0DA49FF23458AFAC73C1C.xml @@ -0,0 +1,496 @@ + + + +Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India + + + +Author + +Naik, Santhosh +National Pusa Collection, Division of Entomology, ICAR- Indian Agricultural Research Institute, New Delhi, 110012, India + + + +Author + +Shashank, P. R. +0000-0002-8177-6091 +spathour@gmail.com + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +409 +420 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.7 + +journal article +10.11646/zootaxa.5492.3.7 +1175-5326 +13234967 +47F51BFA-AC3B-42BB-BBBC-E5ACE29C3F2F + + + + + + + +Terthreutis subrectangulus + +sp. nov. +Shashank & Santhosh + + + + + + +( +Fig. 1F +, +2E +, +3I–J +, +5G–H +) + + +Type locality: +India +(Umiam, +Meghalaya +) + + + +Holotype +: +INPC + + + + + +Diagnosis. + +Terthreutis subrectangulus + + +sp. nov. + +is closely related to + +Terthreutis chiangmaiana + +Razowski, +2008 + + +in adult morphological resemblance but differs in male genitalia characters. + +Terthreutis subrectangulus + +has transtilla incomplete, valva subrectangular, and smooth phallus without any serration whereas + +T. chiangmaiana + +has transtilla complete and continuous, valva rounded, and serrated phallus with the presence of some denticles in the distal portion. Also, it is found that the transtilla is non-continuous or incomplete with two sclerotized lobes and this +type +of transtilla is present in some species of + +Terthreutis + +such as + +T. bipunctata + +, + +T. orbicularis + +, + +T. xanthocycla + +, + +T. sphaerocosma + +and + +T. series + +. However, there are thirteen species described worldwide ( +Buchsbaum & Chen, 2013 +; + +Gilligan +et al. +, 2018 + +), out of which three species have been described from +India +: + +T. argentea +, +T. bulligera +and + + +T. sphaerocosma +( + +Pathania +et al. +, 2020 + +) + +. However, most of the species of + +Terthreutis + +are different but present some elements similar to the new species for example an incomplete transtilla, elongated and well sclerotised uncus, gnathal arms long and wide and well sclerotised sacculus about as long as 1/2 of the length of valva. This study adds one more species to the world fauna of + +Terthreutis + +and increases the number of Indian species to four. + + + + +FIGURE 3. +Wing venation; + +Choristoneura ribhoiensis + + +sp. nov. + +(holotype, male) ( +A. +Forewing, +B. +Hindwing); + +Clepsis laetornata + +( +C. +Forewing, +D. +Hindwing)); + +Diplocalyptis ferruginimixta + +( +E. +Forewing, +F. +Hindwing); + +Diplocalyptis operosa + +( +G. +Forewing, +H. +Hindwing); + +Terthreutis subrectangulus + + +sp. nov +. + +(holotype, male) ( +I. +Forewing, +J. +Hindwing). Scale bar = 1 mm. + + + + +FIGURE 4. + +Choristoneura ribhoiensis + + +sp. nov. + +( +A. +Male genitalia, +B. +Uncus, +C. +Transtilla process, +D. +Phallus); + +Clepsis laetornata + +( +E. +Male genitalia, +F. +Female genitalia). Scale bar A, E, F= 0.5 mm; B, C, D= 0.1 mm. + + + + +FIGURE 5. + +Diplocalyptis ferruginimixta + +( +A. +Male genitalia, +B. +Uncus, +C. +Phallus); + +Diplocalyptis operosa + +( +D. +Male genitalia, +E. +Uncus, +F. +Phallus); + +Terthreutis subrectangulus + + +sp. nov. + +(holotype, male) ( +G. +Male genitalia, +H. +Uncus & Socii). Scale bar A, D, G= 0.5 mm; B, C, E, F, H= 0.1 mm. + + + + +Description. +Head. Ocellus small, brown compound eyes. Head dusted cream white, face cream white. Vertex and dorsal part of frons clothed with long pale cream white rough, upwardly projected scales. Ventral part of frons region covered with small, flat, upwardly raised pale ochreous white scales. Labial palpi slender, clothed with forwardly projected whitish cream scales; first segment short, cream white, tinged with brownish cream; second segment curved, brownish white, interspersed with pale ochreous along dorsal margin from base to near apex; small third segment, porrect with large ochreous white scale. Antenna filiform, pale ochreous white with short cilia. Chaetosemata present. + + +Thorax. Pronotal collar, tegulae, mesonotum, and posterior crest brownish cream mixed with ochreous yellow scales; legs unmodified, middle legs with a pair of tibial spurs; hind tibia covered with dense, cream white scales interspersed with light cream scales, inner margin with tuft of whitish cream scales. Wings: Forewing (wingspan +20 mm +; n= 5) uniformly broad, costa tinged ochreous, termen weakly oblique, gently sinuate; cream brownish ground color; tornal spot small, brown accompanied by more anterior cream brown elongate blotch; postmedian blotch rounded, brownish cream; some brown dots in apical and terminal area. Hindwing yellowish cream with concolor cilia along the apical and anal margin. Wing venation: Forewing with all the veins separated beyond the discal cell, R4 and R5 stalked, M3 and CuA1 arched basally, CuP indistinct. Hindwing Sc reach up to Rs and M1 stalked, M3 and CuA1 stalked, CuP distinct. + +Abdomen. Clothed with yellowish cream. Male genitalia: Uncus long, slender, strongly scelerotised, uniform width narrow towards apex; tegumen broad, rectangular, well seclerotised with strong shoulders; socius not prominent; gnathos strong with small lateral prominence and large terminal lobe; gnathal hooks highly sclerotized, from apex of tegumen shoulder; valva large, subrectangular, symmetrical, lightly sclerotised with minute hairs; transtilla incomplete, with two sclerotised lobes; sacculus strongly sclerotized, uniform width, reach 1/2 of valva; cucullus membranous with tufts of hairs; juxta medium, dumbbell- shaped with broad and rounded base; anellus cup shaped and fused with phallus; vinculum medium well sclerotized. Phallus simple, long and slender with uniform width. No cornuti detected. + +Female. +Unknown. + + + + +Material studied. + +India +: +Meghalaya +: +Ri Bhoi +: CPGSAS, +Umiam +( +25°03’22’’N +, +91°54’42’’E +, 1520 MSL), +2♂ +, + +17.iii.2021 + +, +Mercury +vapour lamp coll. +Santhosh Naik G +& +Sunil + +; + +India +: +Mizoram +: +Kolasib +( +24°11’51’’N +, +92°41’40’’E +, + +493m + +), +1♂ +, + +10.ii.2020 + +, +Mercury +vapour lamp coll. +Niranjana G N +& +Tahseen +R + + +H; +Mizoram +: +Kolasib +( +24°11’51’’N +, +92°41’40’’E +, + +493m + +), +1♂ +, + +11.ii.2020 + +, +Mercury +vapour lamp coll. +Niranjana G N +& +Tahseen +R + +H; + +India +: +Himachal Pradesh +: +Hamirpur +: +Neri +( +31°40’56’’N +, +76°29’15’’E +, 733 MSL), +1♀ +, + +19.xi.2019 + +, +Mercury +vapour lamp coll. +Shivanshu G + +; genitalia slide no. L00050946 ( +♂ +). + + + + +Etymology. +The new species + +Terthreutis subrectangulus + + +sp. nov. + +is named after the subrectangular shape of the valva, which is distinct from all other described species of + +Terthreutis + +. + + + + +Distribution. +India +( +Himachal Pradesh +, +Meghalaya +, +Mizoram +) + + + + +Remarks. +Genus + +Terthreutis + +is described by Meyrick (1918), and species of the genus were studied by + +Kawabe +et al. +(1992) + +and +Razowski (2008) +. + +Terthreutis subrectangulus + + +sp. nov. + +resembles + +Terthreutis chiangmaiana + +Razowski, +2008 + + +in the appearance, but differs in the male genitalia, transtilla incomplete, valva subrectangular, and phallus is smooth without any serration; whereas in + +T. chiangmaiana + +the transtilla is complete and continuous, the valva is rounded, and the phallus is serrated with the presence of some denticles in the distal portion. + + + + \ No newline at end of file diff --git a/data/03/A9/87/03A987C2FFD2DA46FF2346CEFC473C44.xml b/data/03/A9/87/03A987C2FFD2DA46FF2346CEFC473C44.xml new file mode 100644 index 00000000000..073db56c89d --- /dev/null +++ b/data/03/A9/87/03A987C2FFD2DA46FF2346CEFC473C44.xml @@ -0,0 +1,388 @@ + + + +Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India + + + +Author + +Naik, Santhosh +National Pusa Collection, Division of Entomology, ICAR- Indian Agricultural Research Institute, New Delhi, 110012, India + + + +Author + +Shashank, P. R. +0000-0002-8177-6091 +spathour@gmail.com + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +409 +420 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.7 + +journal article +10.11646/zootaxa.5492.3.7 +1175-5326 +13234967 +47F51BFA-AC3B-42BB-BBBC-E5ACE29C3F2F + + + + + + + +Diplocalyptis ferruginimixta +Razowski 2009 + + + + + + + +( +Fig. 1D +, +2C +, +3E–F +, +5A–C +) + + +Type locality: +Vietnam + + + +Holotype +: +MNHU +( +♂ +) + + + +Description. +Head. Clothed brownish cream ochreous scales. Ocellus well developed. Vertex and dorsal part of frons clothed with anterior and forward projected cream brownish ochreous scales, base of the frons with small appressed upwardly projected light brown ochreous coloured scales. Labial upcurved covered with ochreous brown scales; first segment brownish ochreous; second segment long and curved, cream interspersed with dark brown scales along dorsal margin from base to middle apex; small third segment with pale brown, concealed within second segment. Antenna filiform covered with brownish orange scale with intersegmental short cilia. Chaetosemata present. + + +Thorax. Pronotal collar, tegulae, mesonotum, and posterior crest cream mixed with brown cream ochreous interspersed with brown scales; middle legs with a pair of tibial spurs; hind tibia covered with dense, whitish ochreous scales in male, inner margin with a tuft of long, brownish ochreous coloured long hair pencil, originating from base of hindtibia. Wings: Forewing (wingspan +12–14 mm +; n= 13) subrectangular, pale brownish grey ground color pale with cream costal and terminal areas; in dorsal third of wing brownish; strigulation brownish grey, costal fold medium, clothed with brownish ochreous scales, costal terminal areas more cream, basal half of wing with ochreous scale; termen fairly oblique, straight; dorsal third of wing brownish; median fascia slender, uniformly broad, ferruginous medially, edged brown; subapical blotch brownish with grey brown spots; terminal suffusion weak marked by brown median veins. Cilia cream, brown at tornus. Hindwing brownish grey, cream towards base, with brownish venation; cilia paler than wing. Wing venation: Forewing with all the veins separated beyond the discal cell, CuP indistinct. Hindwing with Rs and M1 connate basally, M3 and CuA2 stalked, CuP distinct. + +Abdomen. Covered with dark brownish ochreous scale dorsally, ochreous grey ventrally and tufts of hairs apically. Male genitalia. Uncus long, broad base, apex bilobed; tegumen subrectangular, dorsolaterally with small angled shoulders; gnathos strongly sclerotized bands, arising near middle of tegumen, slightly widened towards apex and gnathal hooks well developed; valva membranous, symmetrical; transtilla sclerotized, spinose plate; juxta medium, subtriangular, bilobed apex; sacculus moderately sclerotized, basely broad and narrow apex, with moderately dense sockets basally, continued as row of short setae reaching base of cucullus; vinculum membranous. Phallus broad with curved, slender ventral termination with an apical spine, with leaf-like deciduous cornuti, cornuti sockets are visible. + +Female. +Unknown. + + + + +Material studied. + +India +: +Karnataka +: +Shivamogga +: +Bhadravathi +( +31°51’14’’ N +, +75°41’20’’ E +, 600 MSL), +1♂ +, + +21.x.2020 + +, +Mercury +vapour lamp coll. +Santhosh Naik G. + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’ N +, +77°44’52’’E +, 901 MSL), +6♂ +, + +10.ix.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’ N +, +77°44’52’’ E +, 901 MSL), +1♂ +, + +21.viii.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’N +, +77°44’52’’E +, 901 MSL), +1♂ +, + +1.ix.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’ N +, +77°44’52’’ E +, 901 MSL), +1♂ +, + +2.ix.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’ N +, +77°44’52’’ E +, 901 MSL), +1♂ +, + +22.xi.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Chikkavaderapura +( +12°53’31’’N +, +77°44’40’’ E +, 914 MSL), +1♂ +, + +6.vii.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Chikkavaderapura +( +12°53’31’’N +, +77°44’40’’ E +, 914 MSL), +1♂ +, + +15.vii.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Chamarajanagar +(11°92’61’’N, 76°94’37’’E, + +717m + +), +1♂ +, 18.xii.217, +Mercury +vapour lamp coll. +Shashank P +R + +; + +India +: +Karnataka +: +Chikkamagaluru +: N B +Colony +( +13°35’00’’N +, +75°46’51’’E +, 680 MSL), +1♂ +, + +18.ix.2020 + +, +Mercury +vapour lamp coll. +Santhosh Naik G + +; genitalia slide no. L00051302 ( +♂ +). + + + + +Distribution. +India +( +Karnataka +); +Vietnam +( + +Pathania +et al. +, 2020 + +). + + + + +Remarks. + +Diplocalyptis ferruginimixta + +looks similar to + +D. tennuicula +Razowski + +from +Zhejiang +, +China +but can be distinguished by the broad uncus, the dorsal process of the labis, and the long, curved terminal process of the phallus ( +Razowski, 2009 +). This species is newly recorded in +India +. + + + + \ No newline at end of file diff --git a/data/03/A9/87/03A987C2FFD3DA45FF2346AAFB0C3F25.xml b/data/03/A9/87/03A987C2FFD3DA45FF2346AAFB0C3F25.xml new file mode 100644 index 00000000000..e220042c5b5 --- /dev/null +++ b/data/03/A9/87/03A987C2FFD3DA45FF2346AAFB0C3F25.xml @@ -0,0 +1,322 @@ + + + +Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India + + + +Author + +Naik, Santhosh +National Pusa Collection, Division of Entomology, ICAR- Indian Agricultural Research Institute, New Delhi, 110012, India + + + +Author + +Shashank, P. R. +0000-0002-8177-6091 +spathour@gmail.com + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +409 +420 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.7 + +journal article +10.11646/zootaxa.5492.3.7 +1175-5326 +13234967 +47F51BFA-AC3B-42BB-BBBC-E5ACE29C3F2F + + + + + + + +Diplocalyptis operosa +( +Meyrick, 1908 +) + + + + + + + +( +Fig. 1E +, +2D +, +3G–H +, +5D–F +) + + +Type locality: +Sri Lanka +(Maskeliya) + + + +Lectotype +: +BMNH +( +♂ +) + + + +Description. +Head. Clothed brownish cream ochreous scales. Ocellus well developed. Vertex and dorsal part of frons clothed with anterior and forward projected cream brownish scales, base of the frons with small appressed upwardly projected light brownish coloured scales. Labial palpi brownish; first segment brownish ochreous scale; second segment long and curved, cream interspersed with dark brown scales along dorsal margin from base to middle apex; small third segment with pale brown concealed within second segment antenna filiform covered with brownish orange scale with intersegmental short cilia. Chaetosemata present. + + +Thorax. Pronotal collar, tegulae, mesonotum, and posterior crest cream mixed with dark brown interspersed with dark gray scales; middle legs with a pair of tibial spurs; hind tibia covered with dense, brownish grey scales in male, inner margin with a tuft of long, brownish orange coloured long hair pencil, originating from base of hindtibia. Wings: Forewing (wingspan +14–16 mm +; n=8) subrectangular, pale brownish grey ground color with deep dark brownish band medially and subapically, tinged with dark greyish scales; costal fold broad, clothed with brownish ochreous scales; cilia cream, brown at tornus. Hindwing brownish grey, cream towards base, with brownish venation; cilia paler than wing. Wing venation: Forewing with all the veins separated beyond the discal cell, CuP indistinct. Hindwing with Rs and M1 connate basally, M3 and CuA2 stalked, CuP distinct. + +Abdomen. Covered with dark brownish ochreous scale dorsally, ochreous grey ventrally and tufts of hairs apically. Male genitalia. Uncus long, broad base, apex bilobed; tegumen subrectangular, dorsolaterally with small angled shoulders; socii small; gnathos strongly sclerotized bands, arising near middle of tegumen, slightly widened towards apex and gnathal hooks well developed; valva membranous, symmetrical; transtilla sclerotized, spinose plate; juxta medium, subtriangular, bilobed apex; sacculus moderately sclerotized, basely broad and narrow apex, with moderately dense sockets basally, continued as row of short setae reaching base of cucullus; vinculum membranous, lateral lobes of vinculum rather long, pointed. Phallus broad with curved, slender ventral termination with apical spine, with bunch of deciduous cornuti. + +Female. +Unknown. + + + + +Material studied. + +India +: +Karnataka +: +Chikkamagaluru +: N B +Colony +( +13°35’00’’ N +, +75°46’51’’E +, 680 MSL), +1♂ +, + +18.ix.2020 + +, +Mercury +vapour lamp coll. +Santhosh Naik G + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’N +, +77°44’52’’E +, 901 MSL), +2♂ +, + +10.ix.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’N +, +77°44’52’’E +, 901 MSL), +3♂ +, + +10.ix.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’ N +, +77°44’52’’E +, 901 MSL), +1♂ +, + +1.ix.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; + +India +: +Karnataka +: +Bengaluru +: +Dommasandra lake +( +12°52’26’’N +, +77°44’52’’E +, 901 MSL), +1♂ +, + +10.xi.2021 + +, +Mercury +vapour lamp coll. +Karthik Reddy M + +; genitalia slide no. L00051084 ( +♂ +). + + + + +Distribution. +India +( +Karnataka +), +Sri Lanka +(Maskeliya) ( +Meyrick, 1908 +). + + + + +Remarks. +In the present study, + +D +. +ferruginimixta +Razowski, 2009 + +and + +D +. +operosa +( +Meyrick, 1908 +) + +are new additions to the Indian + +Diplocalyptis + +fauna. + +D +. +operosa + +is similar to + +D +. +ferruginimixta +Razowski + +but the uncus is longer in + +D. ferruginimixta + +. Eight species of + +Diplocalyptis +species + +are currently known worldwide ( + +Gilligan +et al. +, 2018 + +), with only one species ( + +Diplocalyptis congruentana +(Kennel, 1901)) + +previously recorded from +India +( + +Pathania +et al. +, 2020 + +). This study increases the number of Indian + +Diplocalyptis +species + +to three. + + + + \ No newline at end of file diff --git a/data/03/A9/87/03A987C2FFD5DA47FF234746FAED3C60.xml b/data/03/A9/87/03A987C2FFD5DA47FF234746FAED3C60.xml new file mode 100644 index 00000000000..1b3bb9da6a0 --- /dev/null +++ b/data/03/A9/87/03A987C2FFD5DA47FF234746FAED3C60.xml @@ -0,0 +1,334 @@ + + + +Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India + + + +Author + +Naik, Santhosh +National Pusa Collection, Division of Entomology, ICAR- Indian Agricultural Research Institute, New Delhi, 110012, India + + + +Author + +Shashank, P. R. +0000-0002-8177-6091 +spathour@gmail.com + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +409 +420 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.7 + +journal article +10.11646/zootaxa.5492.3.7 +1175-5326 +13234967 +47F51BFA-AC3B-42BB-BBBC-E5ACE29C3F2F + + + + + + + +Clepsis laetornata +(Wang Li & +Wang, 2003 +) + + + + + + + +( +Fig. 1B–C +, +2B +, +3C–D +, +4E–F +) + + +Type locality: +China +, ( +Yunnan Province +, Weishan County) + + + +Holotype +: +NUTC +( +♂ +) + + + + + +Diagnosis. + +Clepsis laetornata +(Wang Li & +Wang, 2003 +) + +is closely related to + +Clepsis rurinana +(Linnaeus) + +, but can be distinguished from it by basal blotch on the surface of the forewing very large and distinct; uncus shorter and much stronger than that of the latter species; phallus much slender, laterally without subterminal denticle. + + + + +Description. +Head. Ocellus moderate, well developed and brownish black compound eyes. Head brownish dusted with pale brownish, face brownish. Vertex and dorsal part of frons clothed with long pale ochreous brownish rough, upwardly projected scales.Ventral part of frons region covered with small, flat, upwardly raised pale brownish ochreous scales. Labial palpi slender, clothed with forwardly projected pale yellowish scales, first segment clothed with ochreous cream, tinged with whitish cream scales; second segment long and curved, ochreous interspersed with long pale brownish yellow along dorsal margin from base to near apex; small third segment, porrect with long ochreous brown scale. Antenna filiform, pale ochreous, cilia not visible. Chaetosemata present. + + +Thorax: Pronotal collar, tegulae, mesonotum, and posterior crest brownish mixed with pale yellowish orange scales; legs unmodified, middle legs with a pair of tibial spurs; hind tibia covered with dense, white scales interspersed with light brown scales, inner margin with tuft of brown scales. Wings: Forewing (wingspan +14–16 mm +; n= 7) sub rectangular, slightly widening distally, strongly sinuate and slightly convex from basal 1/3; pale brown ground color with three prominent dark brown band at basally, medially and apically. Hindwing pale brown with concolor cilia along the apical and anal margin. Wing venation: Forewing with all the veins separated beyond the discal cell, R4 and R5 stalked, CuP remain at margin. Hindwing with Rs and M1 connate basally, M3 and CuA2 connate basally, CuP distinct. + +Abdomen: Dorsal surface pale brown, ventral surface pale yellow with ochreous brown annual tufts. Male genitalia: Uncus medium, moderately sclerotized, distinctly broadened at middle, obtuse apically; tegumen broad, sub rectangular; socius small; gnathos sclerotized indistinct lateral plate; gnathal hooks sclerotized band from tegumen shoulder join medially and pointed; sacculus narrow, slightly convex near base with minute setae; valva subrectangular, with densely setose inner margin; transtilla lobes large, spinose; juxta medium, heart shaped, rounded base; anellus cup-shaped and fused with phallus; vinculum medium, slightly sclerotized. Phallus elongated and narrow towards apex. Female genitalia: Papillae anales medium, narrow with setae; posterior apophyses almost as long as anterior apophyses; sterigma large, slightly sclerotized, forming a large cup shaped plate; ostium small oval shaped slit opening sclerotized; ductus bursae broad, membranous with scelerotised band run along with its length; corpus bursae rounded sac like membrane with signum large, horn-shaped, inner side with denticles. + + + +Material studied. + +India +: CPGSAS, +Umiam +: +Meghalaya +; ( +25°40’42’’ N +, +91°54’42’’E +, 1520 MSL), +3♂ +and +1♀ +, + +16.iii.2021 + +, +Mercury +vapour lamp coll. +Santhosh Naik G +& +Sunil + +; + +India +: +Karnataka +: +Chikkamagaluru +: N B +Colony +( +13°35’00’’N +, +75°46’51’’E +, 12 MSL), +1♀ +, + +18.ix.2020 + +, +Mercury +vapour lamp coll. +Santhosh Naik G + +; + +India +: +Himachal Pradesh +: +Kinnaur +: +Kalpa +(31°53’77’’ N, +78°27’54’’E +, 2960 MSL), +1♂ +, + +15.viii.2020 + +, +Mercury +vapour lamp coll. +Shashank P +R + +; + +India +: +Kerala +: +Ernakulum +: +Pindimana +( +10°10’51’’ N +, 76°62’68’’E, 680 MSL), +1♂ +, + +18xi.2016 + +, +Mercury +vapour lamp coll. +Shashank P +R + +; genitalia slide no. L00051156 ( +♂ +), L00051157 ( +♀ +). + + + + +Distribution. +India +( +Meghalaya +, +Karnataka +, +Himachal Pradesh +, +Kerala +). + + + + +Remarks. +The specific name comes from the Latin word, laetus (distinct) and ornatus (ornate), corresponding to the distinct basal blotch, median fascia and subapical blotch in the forewing ( + +Wang +et al. +, 2003 + +). Until now, ten species of + +Clepsis + +have been described from +India +( + +Pathania +et al. +, 2020 + +) namely: + +C. gemina +Razowski, 1979 + +; + +C. humana +(Meyrick, 1912) + +; + +C. ingenua +(Meyrick, 1912) + +; + +C. kupwari +Ganai & Khan, 2017 + +[nomen nudum]; + +C. leptograpta +(Meyrick, 1924) + +; + +C. melissa +( +Meyrick, 1908 +) + +; + +C. neomelissa +Rose & Pooni, 2004 + +; + +C. provocata +(Meyrick, 1912) + +; + +C. rurinana +(Linnaeus, 1758) + +and + +C. translucida +( +Meyrick, 1908 +) + +. This study adds one more species to the Indian fauna of + +Clepsis +, + +including a description of the female and the first illustrations of Indian + +Clepsis + +. + + + + \ No newline at end of file diff --git a/data/03/A9/87/03A987C2FFD7DA40FF23461BFC0D3900.xml b/data/03/A9/87/03A987C2FFD7DA40FF23461BFC0D3900.xml new file mode 100644 index 00000000000..2207f54d185 --- /dev/null +++ b/data/03/A9/87/03A987C2FFD7DA40FF23461BFC0D3900.xml @@ -0,0 +1,304 @@ + + + +Discovery of two new species and three new records of the tribe Archipini (Lepidoptera: Tortricidae) from India + + + +Author + +Naik, Santhosh +National Pusa Collection, Division of Entomology, ICAR- Indian Agricultural Research Institute, New Delhi, 110012, India + + + +Author + +Shashank, P. R. +0000-0002-8177-6091 +spathour@gmail.com + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +409 +420 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.7 + +journal article +10.11646/zootaxa.5492.3.7 +1175-5326 +13234967 +47F51BFA-AC3B-42BB-BBBC-E5ACE29C3F2F + + + + + + + +Choristoneura ribhoiensis + +sp. nov. +Shashank & Santhosh + + + + + + +( +Figs. 1A +, +2A +, +3A–B +, +4A–D +) + + +Type locality: +India +(Ri Bhoi, +Meghalaya +) + + + +Holotype +: +INPC +( +♂ +) + + + + + +Diagnosis. + +Choristoneura ribhoiensis + + +sp. nov. + +, is closely related to + +Choristoneura chapana +Razowski, 2008 + +but differ in genitalia characters. Male genitalia of + +C. ribhoiensis + +have the uncus longer, tegumen relatively smaller, well developed transtilla process, and sacculus not prominent. Whereas genitalia of + +C. chapana + +have the uncus relatively smaller, tegumen wider and bigger, poorly developed transtilla process and prominent sacculus running up to the outer margin of valva. Fifty-three species of + +Choristoneura + +are currently recognised globally ( + +Gilligan +et al. +, 2018 + +), out of which six species are recorded from +India +, namely: + +C. colyma +Razowski, 2006 + +; + +C. ferrugininotata +Obraztsov, 1968 + +; + +C. griseicoma +(Meyrick, 1924) + +; + +C. neurophaea +(Meyrick, 1932) + +; + +C. propensa +Razowski, 1992 + +and + +C. pseudofumiferana +Ganai & Khan, 2017 + +[nomen nudum] ( + +Pathania +et al. +, 2020 + +). These species are different from + +C. ribhoiensis + +sp. nov +but present some similar elements, such as an an incomplete transtilla divided in two labies and gnathos moderately sclerotized lateral bands with well-developed sclerotised gnathal hooks arises from tegumen shoulder and are connected medially. + + + + +FIGURE 1. +Dorsal habitus illustrations. +A. + +Choristoneura ribhoiensis + + +sp. nov. + +(holotype, male); +B. + +Clepsis laetornata + +(male); +C. + +Clepsis laetornata + +(female); +D. + +Diplocalyptis ferruginimixta + +(male); +E. + +Diplocalyptis operosa + +(male); +F. + +Terthreutis subrectangulus + + +sp. nov. + +(holotype, male). Scale bar = 2 mm. + + + + +Description. +Head: Ocellus moderate, well developed and brownish compound eyes. Head whitish dusted with light grey, face fuscous. Vertex and dorsal part of frons clothed with long pale ochreous rough, upwardly projected scales. Ventral part of frons region covered with small, flat, upwardly raised pale ochreous scales. Labial palpi clothed with forwardly projected pale yellowish ochreous scales; first segment short, ochreous, tinged with whitish cream; second segment long and broad, ochreous interspersed with pale yellow along dorsal margin from base to near apex; small third segment, porrect with long ochreous scale. Antenna filiform, pale ochreous, cilia not visible. Chaetosemata present. + + +Thorax: Smooth, pronotal collar, tegulae, mesonotum, and posterior crest grey mixed with dark brown scales; legs unmodified, middle legs with a pair of tibial spurs; hind tibia covered with dense white scales interspersed with light grey scales, inner margin with tuft of ochreous grey scales. Wings: Forewing (wingspan +14 mm +) sub rectangular, slightly widening distally, slightly sinuate and slightly convex in outer half, apex falcate, with cream ochreous scales as background interspersed with dark grey, tinged with brown ochreous and orange-grey scales; Hindwing subtriangular, dark brown, with pale brownish grey apex and greyish brown cilia along the apical and anal margin. Wing venation: Forewing with all the veins separated beyond the discal cell, R4 and R5 stalked, CuP remain at margin. Hindwing with Rs and M1 connate basally, M3 and CuA2 connate basally, CuP distinct. + +Abdomen: Light grey ochreous brown. Male genitalia: Uncus long, subtriangular, broad base obtuse apically; tegumen broad, subrectangular; socius not prominent; gnathos moderately sclerotized, lateral bands; gnathal hooks sclerotized, from tegumen shoulder, connected medially; valva medium, ovate, spinose; sacculus small sclerotized with minute setae; transtilla incomplete, formed by two sclerotized lobes with minute spines, connected medially; juxta rather small, round basally; vinculum medium sclerotized. Phallus broad base tapering towards apex; vesica usually with three lanceolate cornuti, caulis short. + +Female. +Unknown. + + + + +Material studied. + +India +: +Meghalaya +: +Ri Bhoi +: CPGSAS, +Umiam +( +25°03’22’’N +, +91°54’42’’E +, 1520 MSL), +1♂ +, + +16.iii.2021 + + +, Mercury vapour lamp coll. Santhosh Naik G & Sunil; genitalia slide no. L00051352 ( +♂ +). + + + + +Etymology. +The new species + +Choristoneura ribhoiensis + + +sp. nov. + +, is named after the +type +locality, Ri Bhoi, +Meghalaya +, +India +. + + + + +Distribution: +India +( +Meghalaya +). + + + + +Remarks: +This new species is placed in + +Choristoneura + +based on adult resemblance and male genitalia characters. The +holotype +was collected from a deciduous forest in +Meghalaya +, +India +. + + + + \ No newline at end of file diff --git a/data/03/F0/0D/03F00D6BFF8FFFF14994F9D7AAC7FB03.xml b/data/03/F0/0D/03F00D6BFF8FFFF14994F9D7AAC7FB03.xml new file mode 100644 index 00000000000..2da46bd0370 --- /dev/null +++ b/data/03/F0/0D/03F00D6BFF8FFFF14994F9D7AAC7FB03.xml @@ -0,0 +1,1199 @@ + + + +First report of the family Karschiidae (Arachnida, Solifugae) from Gansu Province, China, with description of two new species + + + +Author + +Fan, Wenlong +0009-0004-6854-6330 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + + + +Author + +Zhang, Chao +0000-0003-1702-1206 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + + + +Author + +Zhang, Feng +0000-0002-3347-1031 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +369 +394 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.5 + +journal article +10.11646/zootaxa.5492.3.5 +1175-5326 +13234925 +8CBBFFB6-24C8-45EB-B71B-CA636A8AD05F + + + + + + + +Karschia (Rhinokarschia) liui + +sp. nov. + + + + + + +Figures 1 +, +12–15 +, +16A–E +, +17C +. +Table 1 +, +2 +. + + + + +Type material. + + +Holotype + +♂ +(MHBU-Sol-GS2023043001), +CHINA +: +Gansu Province +, +Linze County +, +Pingchuan Town +, +Yigongcheng Village +, +39.3358°N +, +100.0976°E +, ca + +1300m + +elev., + +30.IV.2023 + +, leg. +Jiliang Liu. + + + + + + +Paratypes + +. + +1♂ +(MHBU-Sol-GS2023043002), same data as holotype + +; + +2♂♂ +3♀♀ +(MHBU-Sol-GS2016051901– 03, SWUC-Sol-GS2016051901–02), +CHINA +: +Gansu Province +, +Linze County +, +Pingchuan Town +, +Yigongcheng Village +, +39.3358°N +, +100.0976°E +, ca + +1320m + +elev., + +19.Ⅴ.2016 + +, leg. +Jiliang Liu. + + + +Other material. + +9 juveniles +(MHBU-Sol-GS2016091301-09) +CHINA +: +Gansu Province +, +Linze County +, +Pingchuan Town +, +Yigongcheng Village +, +39.3358°N +, +100.0976°E +, ca + +1300m + +elev., + +13.IX.2016 + +, leg. +Jiliang Liu + +; + +3 juveniles +(MHBU-Sol-GS2023070301-04), +CHINA +: +Gansu Province +, +Linze County +, +Pingchuan Town +, +Yigongcheng Village +, +39.3358°N +, +100.0976°E +, ca + +1300m + +elev., + +3.VII.2023 + +, leg. +Wenlong Fan. + + + + + +FIGURE 12. + +Karschia (Rhinokarschia) liui + + +sp. nov. + +Habitus. +A, B. +Holotype male (MHBU-Sol-GS2023043001). +C, D. +Paratype female (MHBU-Sol-GS2023043002). Scale bars = 10 mm. + + + + +FIGURE 13. + +Karschia (Rhinokarschia) liui + + +sp. nov. + +, holotype male (MHBU-Solifugae-GS2023043001). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Flagellum and +fcs +, prolateral aspect. +D. +Propeltidium, dorsal aspect. +E. +Ctenidia on sternite III, ventral aspect. +F. +Ctenidia on sternite IV, ventral aspect. Scale bars = 1.0 mm (A, B, D); 0.5 mm (E, F); 0.2 mm (C). + + + + +Etymology. +Named after Dr. Jiliang Liu (the Northwest Institute of Eco-Environment and Resources, CAS, +China +), who collected over 100 solifuges from Zhangye City, +Gansu Province +. + + + + +Diagnosis. +Males of + +Karschia (Rhinokarschia) liui + + +sp. nov. + +should belong to the subgenus + +Karschia +( +Rhinokarschia +) + +by having a coiled flagellum and presence of notedly hornlike process on fixed finger ( +Gromov 2003 +; +Figs 13A, B +, +15 A, B +). + +K +. ( +R +.) +liui + +differs from other + +Karschia +( +Rhinokarschia +) + +species except + +Karschia (Rhinokarschia) zarudnyi +Birula, 1922 + +by the presence of a small triangular hornlike process on cheliceral fixed finger ( +Fig. 13A +). This new species differs from + +K. (R.) zarudnyi + +by the shorter flagellum (extending not further than the FD tooth, where it extends nearly to the FT in + +K. (R.) zarudnyi + +), flagellum lateral apophysis triangular-shaped ( +Figs 13C +, +17C +), the small straight (slightly curved posteriorly in + +K +. ( +R +.) +zarudnyi + +) fixed finger hornlike process ( +Fig. 13C +), the non-broadened apex of ctenidia on sternite III ( +Figs 13E +, +15G +), the presence of more short spines on pedipalpal tarsus (8–9), and the absence of papillae on pedipalpal metatarsus ( +Figs 15F +, +16B +). Females differ from other + +Karschia +( +Rhinokarschia +) + +species by the presence of spines on pedipalpal metatarsus and tarsus ( +Fig. 16D, E +), the short, needle-like ctenidia on sternite IV (shorter than other species in this subgenus, ctenidia in + +K. (R.) liui + +reach the distal border of the sternite IV, but ctenidia in other species reach the following sternite) ( +Figs 14F +, +15J +), and the genital operculum not subtriangular, distally not widened and genital opening visible between plates. ( +Figs 14E +, +15I +). + + + + +FIGURE 14. + +Karschia (Rhinokarschia) liui + + +sp. nov. + +, paratype female (MHBU-Solifugae-GS2023043002). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Dentition on left cheliceral fixed finger, ventral aspect. +D. +Propeltidium, dorsal aspect. +E. +Genital operculum, ventral aspect. +F. +Ctenidia on sternite IV, ventral aspect. Scale bars = 1.0 mm (A, B, D); 0.5 mm (C, E); 0.4 mm (F). + + + + +FIGURE 15. + +Karschia (Rhinokarschia) liui + + +sp. nov. + +, holotype male (MHBU-Solifugae-GS2023043002) (A, B, E–H), paratype female (C, D, I, J). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Left chelicera, retrolateral aspect. +D. +Left chelicera, prolateral aspect. +E. +Pedipalpal metatarsus and tarsus, dorsal aspect. +F. +Pedipalpal metatarsus and tarsus, ventral aspect. +G. +Ctenidia on sternite III, ventral aspect. +H. +Ctenidia on sternite IV, ventral aspect. +I. +Genital operculum, ventral aspect. +J. +Ctenidia on sternite IV, ventral aspect. + + + + +FIGURE 16. +Pedipalpal metatarsus and tarsus. +A, B. + +K. (R.) liui + + +sp. nov. + +, holotype male (MHBU-Solifugae-GS2023043001). +C–E. + +K. (R.) liui + + +sp. nov. + +, paratype female (MHBU-Solifugae-GS2023043002). +F, G. + +E. turkestana +Kraepelin, 1899 + +., male (MHBU-Solifugae-GS2019071101). +H, I. + +E. dentapicalis + + +sp. nov. + +, holotype male (MHBU-Solifugae-GS2019071101). +Scale bars = 1.0 mm (A–D, F–I); 0.5mm (E) +. + + + + + +Description. Male. +Holotype + +(MHBU-Sol-GS2023043001). + + +Measurements. +Total body length 16.91, +CH +1.74, CL 4.66, CW 1.54, PL 2.20, PW 3.05, A/CP 8.12, CL/ +CH +2.68. Pedipalp 17.4 (4.75, 5.16, 3.63, 1.03), leg Ⅰ 14.57 (4.43, 3.88, 2.51, 1.17, 0.18, leg II 15.83 (2.56, 4.37, 3.24, 0.78, 1.01), leg III 20.64 (3.91,5.60, 3.95, 1.37, 1.26), leg IV 23.64 (5.57, 5.86, 4.23, 1.20, 1.05). + + +Coloration. +In 75% ethanol-preserved specimens ( +Figs 12A, B +). General background light yellow. Propeltidium yellow, densely covered with short yellowish-brown setae and sparsely with long tawny setae; ocular tubercle black ( +Fig. 13D +). Opisthosoma densely covered with yellowish-brown setae dorsally, except for black abdominal tergites and sclerites. Chelicera yellow brown, covered with long brownish setae; each terminal tooth of fixed finger and movable finger reddish ( +Figs 13A, B +), pedipalps and legs yellowish brown expect coxa and trochanter, covered with brownish setae. Malleoli white. + + +Propeltidium. +Wider than long ( +Fig. 13D +), with a dense pubescence of thin, short, anteriorly directed fine setae; median eyes separated by less than one diameter; propeltidium lobe without eyespot. Anterior, posterior, and lateral edges with several long, curved spiniform setae that perpendicular to the surface of the propeltidium ( +Fig. 13D +). Ocular tubercle with four middle distal spiniform setae. + + +Chelicerae. + +Fixed +finger with median teeth series comprising well-developed primary teeth ( +FP +, +FM +and +FD +), +FD +tooth size greater, while +FM +and +FP +similar in size ( +Figs 13A–C +, +15A, B +); moreover, median teeth series comprising all secondary teeth (2 +FSD +and 2 +FSM +); secondary teeth overall size smaller than primary teeth, distal +FSD +and +FSM +smaller than proximal +FSD +and +FSM +. +Retrofondal +teeth series uninterrupted with six teeth (2 +RFSP +, 1 +RFP +, 1 +RFM +, 2 +RFA +); profondal teeth series consisting of five teeth (2 +PFSP +, 1 +PFP +, 1 +PFSM +, 1 +PFM +) ( +Figs 13A, B +, +15A, B +). +Fixed +finger asetose area sinuous, with ventral margin predominantly linear in lateral aspect; prodorsal carina not sharp, not elevated in lateral aspect, with small and straight hornlike process; fixed finger mucron without subterminal ( +FST +) teeth, and apex ( +FT +tooth) curved, hook-shaped. +Movable +finger with median teeth series comprising well-developed +MP +and +MM +primary teeth, the former of which larger, and one +MSM +secondary tooth smaller than +MM +(i.e., +MP +> +MM +> +MSM +), movable finger +MP +and +MSM +teeth close to one another, +MSM +upright and triangular-shaped; movable finger without subproximal ( +MSP +) or subterminal ( +MST +) teeth; movable finger mucron moderately long, with obvious prolateral flange in front of the +MM +tooth ( +Figs 13A–C +, +15A, B +). +Retrolateral +and dorsal surfaces with abundant retrolateral manus ( +rlm +) and retrolateral finger setae ( +rlf +), of different sizes, which predominantly straight and rigid; some of these setae arranged in bilaterally symmetrical pattern, as some principal retrolateral finger (principal +rlf +) setae that more flexible than others ( +Figs 13A +, +15A +); prolateral surface with array of setal +types +, as follows: proventral distal ( +pvd +) setae consisting of row of plumose setae, starting at level of the flagellum and ending near level of +RFSP +tooth; proventral subdistal ( +pvsd +) setae arranged in rather disorganized pattern, +pvsd +comb not markedly differentiated; two +pvsd +elongated called flagellar complex plumose ( +fcp +) setae, moreover two +fcs +behind flagellum above the +fcp +; carpet-like field of bristle-like promedial ( +pm +) setae broad. Stridulatory plate longer than high, occupying most of the prolateral surface of manus, above stridulatory plate two rows of +pdp +existing; stridulatory apparatus not obvious. Distal limit of the prolateral setose area of movable finger reaching midpoint between +MSM +and +MM +teeth; movable finger prodorsal ( +mpd +) setal series consisting of spinous setae arranged in a rather staggered row, adjacent to abundant non-plumose setae of the movable finger promedial ( +mpm +) and proventral ( +mpv +) setal series ( +Figs 13B +, +15B +). Flagellum coiled, fringed and sessile, with a small triangular lateral apophysis. The flagellar complex includes two +fcp +and two short, thick +fcs +( +Figs 13B, C +, +15B +, +17C +). Flagellar basal peg unexpanded, tapering from the base ( +Fig 13C +, +17C +) + +. + + + +FIGURE 17. +Flagellum of male. +A. + +Eusimonia turkestana +Kraepelin, 1899 + +B. + +Eusimonia dentapicalis + + +sp. nov. +C. + + +Karschia (Rhinokarschia) liui + + +sp. nov. +Scale bars = 0.2 mm + +. + + + + +Opisthosoma +. + +The entire surface covered with almost adpressed fine setae, and numerous long, curved, bifurcate setae, and tergites with abundant setae. Sternite III with two posterior paramedian groups of ctenidia, being gradually larger to posterior, ctenidia coarse needle-like gradually sharp from the base to the spire ( +Figs 13E +, +15G +); sternite IV with a row of 13 (partial shedding) long and thin cylindrical ctenidia ( +Figs 13F +, +15H +). + + + +Pedipalps +. + +Entirely covered with short fine setae and long, thick setae. Tarsus with eight ventral spines not symmetrical arrangement; metatarsus with 12 ventral spines (partial shedding) not arranged in pairs and without papillae ( +Figs 15E, F +, +16A, B +). + + +Legs. +Legs totally covered with long, thick, setae and short fine setae. Leg Ⅰ spineless with two small claws. Tibiae II, III, and IV with a pair of distal spiniform setae ventrally; tibiae II and III with a single dorsal spiniform setae. Metatarsi II and III with a series of three dorsal spiniform setae, a pair of distal spiniform setae ventrally, and some paired short, thick, spiniform setae over their entire ventral surface; metatarsus IV also with these paired spiniform setae over its entire ventral surface and two distal spine-like setae ventrally. + + + +TABLE 1. +DNA voucher specimen metadata. + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
+Species + +DNA Voucher Code + +Sex + +GenBank accession no. + +Sequence length + +Collection localities +
+1 + + +K(R). +liui + + +sp. nov. + +FWL068MalePP627402682Yigongcheng Village, PingchuanTown, Linze County, Gansu Province, China
+2 + + +K(R). +liui + + +sp. nov. + +FWL079FemalePP627401695Yigongcheng Village, PingchuanTown, Linze County, Gansu Province, China
+3 + + +E. turkestana + +FWL058FemalePP627404694Annamba wild camel National Nature Reserve, Akesay County, Gansu Province, China
+4 + + +E. turkestana + +FWL071MalePP627407685Yigongcheng Village, PingchuanTown, Linze County, Gansu Province, China
+5 + + +E. turkestana + +FWL113FemalePP627406683Yigongcheng Village, PingchuanTown, Linze County, Gansu Province, China
+6 + + +E. turkestana + +FWL065FemalePP627408693Suwu Desert, Lanhu Town, Minqin County, Gansu Province, China
+7 + + +E. turkestana + +FWL067MalePP627409686Gansu Anxi Extreme Drought Desert National Nature Reserve, Guazhou County, Gansu Province, China
+8 + + +E. +dentapicalis + + +sp. nov. + +FWL074MalePP627410683Jinta Town, Jinta County, Gansu Province, China
+9 + + +E. +dentapicalis + + +sp. nov. + +FWL075MalePP627411695Jinta Town, Jinta County, Gansu Province, China
+10 + + +E. +dentapicalis + + +sp. nov. + +FWL117FemalePP627412683Tangwan Villag, Pingchuan Town, Linze County, Gansu Province, China
+ + + +TABLE 2. +Genetic distance among the three species ( + +K. (R.) liui + + +sp. nov. +, + + +E. turkestana + +and + +E. dentapicalis + + +sp. nov. + +). + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + + +
+1 + +2 + +3 + +4 + +5 + +6 + +7 + +8 + +9 + +10 +
+1 +
+2 +0.0014
+3 +0.21440.2138
+4 +0.20820.20760.0439
+5 +0.20670.20610.04690.0029
+6 +0.20230.20200.04110.02500.0234
+7 +0.21410.213500.04390.04680.0410
+8 +0.20700.20560.14260.14390.14240.13800.1424
+9 +0.20820.20610.14060.14180.14040.13620.14040.0015
+10 +0.21000.20850.14120.14240.14100.14100.14100.02790.0279
+ + +Female. +Paratype +female (MHBU-Sol-GS2016051901). + + +Measurements. +Total body length 19.87, +CH +1.69, CL 4.94, CW 1.52, PL 1.91, PW 3.27, A/CP 5.90, CL/ +CH +2.92. Pedipalp 12.81 (3.13, 3.80, 2.53, 0.80), leg Ⅰ 10.08 (2.20, 2.52, 1.70, 0.85, 0.14), leg II 8.97 (1.40, 2.15, 1.58, 0.55, 0.58), leg III 12.04 (2.37, 2.639, 2.06, 0.49, 0.65), leg IV 17.49(3.79, 3.34, 2.98, 1.02, 1.09). + + +Coloration. +In 75% ethanol-preserved specimens ( +Figs 12C, D +). Coloration mostly as in the male, but may be due to preservation differences, female darker overall. + + +Propeltidium. +Same as in the male. + + + +Chelicera +. + +Fixed finger with median teeth series comprising well-developed primary teeth (FP, FM and FD), FD tooth size smaller, while FM and FP similar in size ( +Figs 14A–C +, +15C, D +); moreover, median teeth series comprising all secondary teeth (2FSD and 2 +FSM +); secondary teeth overall size smaller than primary teeth, distal FSD and +FSM +smaller than proximal FSD and +FSM +. Retrofondal teeth series uninterrupted with nine teeth (3RFSP, 1RFP, 1RFSM, 1RFM, 3RFA); profondal teeth series consisting of six teeth (3PFSP, 1PFP, 1PFSM, 1PFM). Fondal teeth graded as II, IV, VI, VIII, IX, tiny I, tiny III, tiny V, tiny VII retrolaterally; I, II, III, IV, V tiny VI prolaterally ( +Fig. 14C +). Movable finger with median teeth series comprising well-developed MP and MM primary teeth, the former of which larger, and two MSM, four MST and four MSP secondary tooth that smaller than MM (i.e., MP> MM> MSM> MST> MSP), movable finger MSM teeth located MM and MP center, MST teeth located MM front, and MSP located MP after; all secondary teeth upright and triangular-shaped. Fixed finger and movable finger mucron without subterminal (FST and MST) teeth. Cheliceral setose areas and stridulatory plate mostly like male, but female without +fcp, fcs, +and +pdp +only one row ( +Figs 14B +, +15D +). + + +Opisthosoma. +The entire surface covered with almost adpressed fine setae and numerous long, curved, bifurcate setae. Genital operculum not triangular-shape, not widened distally and genital opening visible between plates. Genital plate resembling a trapezoid-shaped with a slight sunken in the middle structure, and with slightly chitinized folds distal to plates ( +Figs 14E +, +15I +). 13 short needle-like ctenidia on genital sternite IV which extending the succeeding sternite edge ( +Figs 14F +, +15J +). + + +Pedipalps. +Entirely covered with short fine setae and long, thick setae. Tarsus with three ventral spines not symmetrical arrangement, and metatarsus with five ventral spines (partial shedding) not arranged in pairs ( +Figs 16D, E +). + + + +Legs +. + +Same as in the male. + + + + +Distribution and habitat. + +K. (R). +liui + + +sp. nov. + +is known only from the +type +locality ( +Fig. 1 +). Habitat: Gobi Desert ( +Fig. 2A +). In contrast to the majority of mostly + +Karschia +species + +, which are typically found in hills and mountains as noted by +Gromov (2003) +, + +K. (R.) liui + + +sp. nov. + +exhibits a preference for low-altitude, flat regions of the Gobi Desert. + + +Collection method. +During the day by hand collecting from under stones. Given their nocturnal habits, we also collect specimens using pitfall traps. + + +Variability. +Males. Total length 16.39–17.23. Body coloration light yellow to yellow. Chelicera with manus yellowish to brown. Pedipalpal tarsus with 8–9 spines and metatarsus with 10–13 spines. The number of cheliceral fixed finger fondal teeth 9–11 (profondal teeth 4–5; retrofondal teeth 5–6). The number of ctenidia on sternite IV 13–14. Females. Total length 18.56–21.26. Variability of body coloration as in males. Pedipalpal tarsus with 3–4 spines, metatarsus with 4–6 spines. The number of cheliceral fixed finger fondal teeth 14–16 (profondal teeth 4–6; retrofondal teeth 10–11). MST 3–4, MSP 3–4. The number of ctenidia on sternite IV 12–13. + + + + +Remark. +Based on the comparison of genetic distances, with a genetic distance of 0.14% ( +Table 2 +) between males and females collected from the same locations, we believe that they are the same species. + + + + \ No newline at end of file diff --git a/data/03/F0/0D/03F00D6BFF95FFF94994F9D6AA5EFA07.xml b/data/03/F0/0D/03F00D6BFF95FFF94994F9D6AA5EFA07.xml new file mode 100644 index 00000000000..305497a504b --- /dev/null +++ b/data/03/F0/0D/03F00D6BFF95FFF94994F9D6AA5EFA07.xml @@ -0,0 +1,804 @@ + + + +First report of the family Karschiidae (Arachnida, Solifugae) from Gansu Province, China, with description of two new species + + + +Author + +Fan, Wenlong +0009-0004-6854-6330 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + + + +Author + +Zhang, Chao +0000-0003-1702-1206 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + + + +Author + +Zhang, Feng +0000-0002-3347-1031 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +369 +394 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.5 + +journal article +10.11646/zootaxa.5492.3.5 +1175-5326 +13234925 +8CBBFFB6-24C8-45EB-B71B-CA636A8AD05F + + + + + + + +Eusimonia dentapicalis + +sp. nov. + + + + + + +Figures 1 +, +8–11 +, +16H, I +, +17B +. +Table 1 +, +2 +. + + + + +Type Material. + + +Holotype + +♂ +(MHBU-Sol-GS2023070401), +CHINA +: +Gansu Province +, +Jinta County +, +Jinta Town +, +39.7921°N +, +99.0911°E +, ca + +1280m + +elev., + +4.VII.2023 + +, leg. +Wenlong Fan. + + + + + +Paratypes +: + +7♀♀ +(MHBU-Sol-GS2023070402–09), same data as holotype + +; +1♂ +1♀ +(MHBU-Sol-GS2023050701– 02), + +1♂ +2♀ +(SWUC-Sol-GS2023050701–03), +CHINA +: +Gansu Province +, +Linze County +, +Pingchuan Town +, +Tangwan Village +, +39.4500°N +, +100.2667°E +, ca + +1240m + +elev., + +7.Ⅴ.2023 + +, leg. +Jiliang Liu + +; + +1♂ +2♀♀ +(MHBU-Sol-GS2023060801– 03), +CHINA +: +Gansu Province +, +Linze County +, +Dongxiaokouzi +, +39.4000°N +, +100.1833°E +, ca + +1320 m + +elev., + +8.VI.2023 + +, leg. +Jiliang Liu + +; + +2♀♀ +(SWUC-Sol-GS2023030701–02), +Gansu Province +, +Linze County +, +Pingchuan Town +, +39.4019°N +, +100.1215°E +, ca + +1350m + +elev., + +3.VII.2023 + +, leg. +Wenlong Fan. + + + + +FIGURE 8. + +Eusimonia dentapicalis + + +sp. nov. + +Habitus. +A, B. +Holotype male (MHBU-Sol-GS2023070401). +C, D. +Paratype female (MHBU-Sol-GS2023070402). Scale bars = 10 mm. + + + + +FIGURE 9. + +Eusimonia dentapicalis + + +sp. nov. + +, holotype male (MHBU-Sol-GS2023070401). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Flagellum, prolateral aspect. +D. +Propeltidium, dorsal aspect. +E. +Ctenidia on sternite III, ventral aspect. +F. +Ctenidia on sternite IV, ventral aspect. Scale bars = 1.0 mm. + + + +Other material. + +5 juveniles +(MHBU-Sol-GS2023070410-15), +CHINA +: +Gansu Province +, +Jinta County +, +Jinta Town +, +39.7921°N +, +99.0911°E +, ca + +1280m + +elev., + +4.VII.2023 + +, leg. +Wenlong Fan + +; + +4 juveniles +(MHBU-SolGS2023060804–07), +CHINA +: +Gansu Province +, +Linze County +, +Dongxiaokouzi +, +39.4000°N +, +100.1833°E +, ca + +1320 m + +elev., + +8.VI.2023 + +, leg. +Jiliang Liu. + + + + + +Etymology. +The specific name is derived from a combination of the Latin words “ +apical +” and “ +dentata +”, meaning terminal tooth, referring to the presence of subterminal (MST) teeth on cheliceral fixed finger. + + + + +Diagnosis. +Males of + +Eusimonia dentapicalis + + +sp. nov. + +differ from other + +Eusimonia +species + +except + +E. turkestana + +by having a triangular-shaped dorsal secondary flagellum ( +Figs 9C +, +17B +) and the absence of spines on pedipalpal tarsus ( +Figs 16H, I +). + +E. dentapicalis + + +sp. nov. + +differs from + +E. turkestana + +by the dorsal secondary flagellum overall shape wider and the tubelike ventral primary flagellum blunt at the apex ( +Figs 9C +, +17B +), by having weaker spiniform setae on the anterior edge of the propeltidium ( +Fig. 9D +), by a shorter, curved hornlike process (extending not further than the FSD tooth, where it extends nearly to the FD in + +E. turkestana + +) and the absence of tooth on hornlike process ( +Fig. 9C +); by the presence of 4–5 tiny subterminal (FST) teeth on fixed finger mucron in all males examined ( +Fig. 9C +), and by the longer ctenidia on sternite III than + +E. turkestana + +(ctenidia in + +E. dentapicalis + +extend past the distal border of the sternite III, but ctenidia in + +E. turkestana + +not extend past) ( +Figs 9E +, +11E +). Females differ from other + +Eusimonia +species + +by the presence of tiny FST on the cheliceral fixed finger, the ( +Figs 10A +, +11G +), genital operculum triangular in shape with clear demarcation between the plates, genital operculum distally widened and genital opening visible between plates ( +Fig 10E +, +11I +). + + + + +FIGURE 10. + +Eusimonia dentapicalis + + +sp. nov. + +, paratype female (MHBU-Sol-GS2023070402). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Dentition on left cheliceral fixed finger, ventral aspect. +D. +Propeltidium, dorsal aspect. +E. +Genital operculum, ventral aspect. +F. +Ctenidia on sternite IV, ventral aspect. +Scale bars = 1.0 mm (A, B, D); 0.5 mm (C, E,); 0.4 mm (F) +. + + + + + +Description. Male. +Holotype + +(MHBU-Sol-GS2023070401). +Measurements. +Total body length 13.21, +CH +1.22, CL 4.74, CW 1.13, PL 1.92, PW 2.86, A/CP 6.01, CL/ +CH + +3.87. Pedipalp 13.38 (3.19, 3.45, 3.28, 1.21), leg Ⅰ 10.19 (2.05, 2.73, 1.89, 0.82, 0.05), leg II 9.92 (1.80, 2.67, 1.22, +0.84, 0.84), leg III 11.85 (2.40, 3.18, 1.50, 0.68, 0.73), leg IV 16.42 (3.43, 3.19, 3.73, 0.63, 1.04). + + +FIGURE 11. + +Eusimonia dentapicalis + + +sp. nov. + +, holotype male (A–F), paratype female (G–J). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Pedipalpal metatarsus and tarsus, dorsal aspect. +D. +Pedipalpal metatarsus and tarsus, ventral aspect. +E. +Ctenidia on sternite III, ventral aspect. +F. +Ctenidia on sternite IV, ventral aspect. +G. +Left chelicera, retrolateral aspect. +H. +Left chelicera, prolateral aspect. +I. +Genital operculum, ventral aspect. +J. +Ctenidia on sternite IV, ventral aspect. + + + +Coloration. +In 75% ethanol-preserved specimens ( +Figs 8A, B +). Prosoma general background yellow, and opisthosoma’s general background yellow-green. Propeltidium yellow with brown on either side of ocular tubercle, densely covered with short yellowish-brown setae and sparsely with long tawny setae; ocular tubercle black ( +Fig. 9D +). Opisthosoma densely covered with yellowish-brown setae dorsally, abdominal tergites and sclerites dark yellow-green. Chelicera yellow, covered with long brownish setae; each terminal tooth of fixed finger and movable finger reddish ( +Figs. 9A, B +), pedipalps and legs yellowish-brown except coxa and trochanter, covered with brownish setae. Malleoli white. + + +Propeltidium. +Wider than long ( +Fig. 9D +), with a dense pubescence of thin, short, anteriorly directed fine setae; median eyes separated by less than one diameter; propeltidium lobe without eyespot. Anterior, posterior, and lateral edges with several long, curved spiniform setae perpendicular to the surface of the propeltidium. Ocular tubercle with a row of proximal spiniform setae, four short and two long middle distal spiniform setae. On each side of ocular tubercle, propeltidium with longer spiniform setae ( +Fig. 9D +). + + +Chelicerae. + +Fixed +finger with median teeth series comprising well-developed primary teeth ( +FP +, +FM +and +FD +), +FD +tooth size lesser, while +FM +and +FP +similar in size ( +Figs 9A–C +, +11A, B +); moreover, median teeth series comprising all secondary teeth (2 +FSD +and 2 +FSM +); secondary teeth overall size smaller than primary teeth, distal +FSD +and +FSM +smaller than proximal +FSD +and +FSM +( +Fig. 9C +). +Retrofondal +teeth series uninterrupted with five teeth (1 +RFP +, 1 +RFM +, 3 +RFA +), and profondal teeth series consisting of five teeth (2 +PFSP +, 1 +PFP +, 1 +PFSM +, 1 +PFM +) ( +Fig. 9C +). +Fixed +finger asetose area sinuous, with ventral margin predominantly linear in lateral aspect; prodorsal carina not sharp, not elevated in lateral aspect ( +Figs 9A–C +). +Cheliceral +fixed finger hornlike process slightly curved without teeth ( +Fig. 9C +), mucron with four tiny subterminal ( +FST +) teeth, and apex ( +FT +tooth) curved, hook-shaped ( +Fig. 9C +). +Movable +finger with median teeth series comprising well-developed +MP +and +MM +primary teeth, the former of which larger, and two +MSM +secondary tooth smaller than +MM +(i.e., +MP +> +MM +> +MSM +), movable finger one +MSM +teeth close to +MP +, another one close to +MM +, +MSM +upright and triangular-shaped; movable finger without subproximal ( +MSP +) or subterminal ( +MST +) teeth; movable finger mucron moderately long, with obvious prolateral flange in front of the +MM +tooth. +Retrolateral +and dorsal surfaces with abundant retrolateral manus ( +rlm +) and retrolateral finger ( +rlf +) setae, of different sizes,, which are predominantly straight and rigid; some of these setae arranged in bilaterally symmetrical patterns, as some principal retrolateral finger (principal +rlf +) setae that more flexible than others ( +Figs 9A +, +11A +); prolateral surface with array of setal +types +( +Figs 9B +, +11B +), as follows: proventral distal ( +pvd +) setae consisting of row of strong setae, starting at level of the flagellum and ending near level of +PFM +tooth; proventral subdistal ( +pvsd +) setae arranged in rather disorganized pattern, +pvsd +comb not markedly differentiated; carpet-like field of bristle-like promedial ( +pm +) setae broad. Stridulatory plate longer than high, occupying most of the prolateral surface of manus, above stridulatory plate two prodorsal proximal ( +pdp +) setae existing; stridulatory apparatus not obvious. Distal limit of the prolateral setose area of movable finger reaching midpoint between +MSM +and +MM +teeth; movable finger prodorsal (mpd) setal series consisting of spinous setae arranged in a rather staggered row, adjacent to abundant non-plumose setae of the movable finger promedial ( +mpm +) and proventral ( +mpv +) setal series ( +Figs 9B +, +11B +). Flagellum typical of + +Eusimonia + +, bear two distinct, membranous or weakly sclerotized, sessile flagella: a broad, thin, dorsal secondary flagellum and a tubelike ventral primary flagellum ( +Figs 9A–C +, +11A, B +, +17B +); flagellum without flagellar complex subspiniform to spiniform ( +fcs +) setae; membranous dorsal flagellum (secondary flagellum) with a slight central depression ( +Figs 9C +, +17B +) + +. + + +Opisthosoma. +The entire surface covered with almost adpressed fine setae, and numerous long, curved, bifurcate setae, and tergites with abundant setae. Sternite III with 4+5 coarse and spiny ctenidia extend past the distal border of the sternite III ( +Figs 9E +, +11E +), sternite IV with a row of 15 long and thin peg-like ctenidia, almost reach to the distal edge of the sternite Ⅴ ( +Figs 9F +, +11F +). + + + +Pedipalps +. + +Entirely covered with short fine setae and long, thick setae. Tarsus without spines; posterior half of metatarsus thickens inward, resembling a hump, with six quite strong, distally diminishing, forward-inclined spines which thicker and flattened at distal ends ( +Figs 11C, D +, +16H, I +). + + +Legs. +Covered with abundant short and a few long setae, similar to those on pedipalps. Leg I with two small claws but without spine-like setae. Tibiae II, III, and IV with a pair of distal spine-like setae ventrally; tibiae II and III with a single dorsal spine-like setae. Metatarsi II and III with a series of three dorsal spine-like setae, a pair of distal spine-like setae ventrally, and some paired short, thick, spine-shaped bristles over their entire ventral surface; metatarsus IV also with these paired bristles over its entire ventral surface and two distal spine-like setae ventrally. + + +Female. +Paratype +female (MHBU-Sol-GS2023070402). + + +Measurements. +Total body length 12.72, +CH +1.35, CL 4.30, CW 1.10, PL 1.89, PW 2.50, A/CP 4.07, CL/ +CH +3.20. Pedipalp 11.13 (3.00, 2.76, 2.78, 1.00), leg Ⅰ 7.08 (1.37, 1.76, 1.49, 0.61, 0.04), leg II 7.95 (1.39, 1.29, 2.10, 0.66, 0.58), leg III 10.95 (2.03, 2.75, 2.24, 0.74, 0.66), leg IV 14.84 (3.11, 3.26, 3.03, 0.81, 0.77). + + +Coloration. +In 95% ethanol-preserved specimens ( +Figs 8C, D +). Coloration as in the male. + + +Propeltidium. +Mostly the same as male, but female ocular tubercle spiniform setae on each side of ocular tubercle not as strong as in male ( +Fig. 10D +). + + + +Chelicera +. + +Fixed finger with median teeth series comprising well-developed primary teeth (FP, FM and FD), FD tooth size smaller, while FM and FP similar in size ( +Figs 10A–C +, +11G, H +); moreover, median teeth series comprising all secondary teeth (2FSD and 2 +FSM +); secondary teeth overall size smaller than primary teeth, distal FSD and +FSM +smaller than proximal FSD and +FSM +. Retrofondal teeth series uninterrupted with seven teeth (3RFSP, 1RFP, 1RFM, 2RFA); profondal teeth series consisting of three teeth (1PFP, 1PFSM, 1PFM) ( +Fig. 10C +). Fondal teeth graded as II, IV, III, V, I, VI, VII retrolaterally; I, II, III prolaterally ( +Fig. 10C +). Fixed finger mucron with two tiny subterminal (FST) teeth. Movable finger with median teeth series comprising well-developed MP and MM primary teeth, the former of which larger, and four MSM, four MST and three MSP secondary teeth that smaller than MM (i.e., MP> MM> MSM> MSP> MST), movable finger MSM teeth located MM and MP center, MST teeth located MM front, and MSP located MP after; all secondary teeth upright and triangular-shaped ( +Figs 10A, B +, +11G, H +). Cheliceral setose areas and stridulatory plate mostly like male, but female proventral distal ( +pvd +) setae only consist of row of plumose setae ( +Figs 10B +, +11H +). + + +Opisthosoma. +The entire surface covered with almost adpressed fine setae and numerous long, curved, bifurcate setae. Genital operculum with clear demarcation between the plates, distal border widened, resembling a triangle shape. Genital opening visible between plates. ( +Figs 10E +, +11I +). 14 short needle-like ctenidia on sternite IV extending 3/4 the length of the following sternite ( +Figs 10F +, +11J +). + + +Pedipalps. +Entirely covered with short fine setae and long, thick setae and without spines. + + + +Legs +. + +Same as in the male. + + + + +Distribution and habitat. + +E. dentapicalis + + +sp. nov. + +is known from the northwest part of +Gansu Province +, +China +( +Fig. 1 +). Habitat: desert ( +Figs 2C–E +). + +E. dentapicalis + + +sp. nov. + +also live in clayey, stony or sandy loam deserts at low altitudes. + + +Collection method. +During the night using an ultra-violet lamp. Under ultraviolet light, +Solifugae +reflect a faint blue glow, making it easy to spot and catch. Given their nocturnal habits, we also collect specimens using pitfall traps. + + +Variability. +Males. Total length 11.32–16.64. Body coloration light yellow to yellow. Chelicerae with manus yellowish to brown. The number of cheliceral fixed finger fondal teeth 9–11 (profondal teeth 4–5; retrofondal teeth 5–6), FST 4–5. The number of ctenidia on sternite IV 12–15. Females. Total length 12.32–15.39. Variability of body coloration as in males. The number of cheliceral fixed finger fondal teeth 10–11 (profondal teeth 3–5; retrofondal teeth 6–7). FST 2–4, MST 4–5, MSP 3–4. The number of ctenidia on sternite IV 13–15. + + + + +Remark. +Based on the comparison of genetic distances, with a genetic distance of 0.15% ( +Table 2 +) between male and female collected from same locations, we believe that they are same species. + + + + \ No newline at end of file diff --git a/data/03/F0/0D/03F00D6BFF9DFFE34994FA2AAC94F9CF.xml b/data/03/F0/0D/03F00D6BFF9DFFE34994FA2AAC94F9CF.xml new file mode 100644 index 00000000000..2fbb2fadd2d --- /dev/null +++ b/data/03/F0/0D/03F00D6BFF9DFFE34994FA2AAC94F9CF.xml @@ -0,0 +1,740 @@ + + + +First report of the family Karschiidae (Arachnida, Solifugae) from Gansu Province, China, with description of two new species + + + +Author + +Fan, Wenlong +0009-0004-6854-6330 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + + + +Author + +Zhang, Chao +0000-0003-1702-1206 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + + + +Author + +Zhang, Feng +0000-0002-3347-1031 +Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China & Hebei Basic Science Center for Biotic Interaction, Hebei University, Baoding, Hebei 071002, P. R. China + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +369 +394 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.5 + +journal article +10.11646/zootaxa.5492.3.5 +1175-5326 +13234925 +8CBBFFB6-24C8-45EB-B71B-CA636A8AD05F + + + + + + + +Eusimonia turkestana +Kraepelin, 1899 + + + + + + + +Figures 1 +, +4–7 +, +16F, G +, +17A +. +Table 1 +, +2 +. + + + + +Type material. + +Holotype +♂ +, +Central Asia +(as +Turkestana +in the original description), stored at +ZMB +(the +Zoological Museum +, +Museum +for +Naturkunde +, +Berlin +, +Germany +), not examined. + + + +Material examined: + +1♂ +(MHBU-Sol-GS2019071101), +CHINA +: +Gansu Province +, +Guazhou County +, +Gansu +Anxi Extreme Drought Desert National Nature Reserve +, +40.1317°N +, +95.8667°E +, ca + +1610m + +elev., + +11.VII.2019 + +, leg. +Xinglong Bai + +; + +1♂ +(MHBU-Sol-GS2023070601), +CHINA +: +Gansu Province +, +Guazhou County +, +Gansu +Anxi Extreme Drought Desert National Nature Reserve +, +40.1307°N +, +95.8652°E +, ca + +1530m + +elev., + +6.VII.2023 + +, leg. +Wenlong Fan + +; + +3♂♂ +1♀ +(SWUC-Sol-GS2023070101–04), +CHINA +: +Gansu Province +, +Minqin County +, +Lanhu Town +, +Suwu Desert +, +38.2687°N +, +103.2717°E +, ca + +1440m + +elev., + +1.VII.2023 + +, leg. +Wenlong Fan + +; + +9♂♂ +10♀♀ +(MHBU-Sol-GSJLL01–19) +CHINA +: +Gansu Province +, +Linze County +, +Pingchuan Town +, +Yigongcheng Village +, +39.3359°N +, +100.0977°E +, ca + +1320m + +elev., leg. +Jiliang Liu + +; + +3♀♀ +(SWUC-Sol-GS2019071601–03), +CHINA +: +Gansu Province +, +Aksay Kazakh Autonomous County +, +Annanba Wild Camel National Nature Reserve +, +39.2361°N +, +92.3392°E +, ca + +2240m + +elev., + +16.VII.2019 + +, leg. +Xinglong Bai. + + + + + +Description. Male +(MHBU-Sol-GS2019071101). + + + +FIGURE 1. +Map plotting known locality records. + + + +Measurements. +Total body length 11.48, +CH +1.13, CL 4.07, CW 0.71, PL 1.56, PW 2.23, A/CP 6.55, CL/ +CH +3.60. Pedipalp 11.73 (2.68, 3.24, 2.83, 1.04), leg Ⅰ 9.62 (2.16, 2.52, 1.71, 0.94, 0.06), leg II 9.63 (1.79, 2.25, 1.82, 0.63, 0.71), leg III 11.47 (2.44, 2.56, 1.96, 0.82, 0.73), leg IV 15.53(3.86, 4.68, 1.37, 0.86, 0.82). + + +Coloration. +In 95% ethanol-preserved specimens ( +Figs 4A, B +). Prosoma general background yellow, and opisthosoma general background black. Propeltidium tan with brown on either side of ocular tubercle, densely covered with short yellowish-brown setae and sparsely with long tawny setae; ocular tubercle black ( +Fig. 5D +). Opisthosoma densely covered with yellowish-brown setae dorsally, abdominal tergites and sclerites black. Chelicera yellow, covered with long brownish setae; each terminal tooth of fixed finger and movable finger reddish ( +Figs 5A, B +), pedipalps and legs yellowish-brown except coxa and trochanter, covered with brownish setae. Malleoli white. + + + +FIGURE 2. +Habitats where + +Karschia (Rhinokarschia) liui + + +sp. nov. + +( +A +), + +Eusimonia +. +dentapicalis + + +sp. nov. + +( +C–E +), and + +E. turkestana +Kraepelin, 1899 + +. ( +B, F +) were found. +A. +Linze County, Pingchuan Town, Yigongcheng Village. +B. +Jiuquan City, Aksay Kazakh Autonomous County, Annanba Wild Camel National Nature Reserve. +C. +Linze County, Dongxiaokouzi. +D. +Linze County, Pingchuan Town, Tangwan Village. +E. +Jinta County, Jinta Town. +F. +Gansu Province, Guazhou County, Gansu Anxi Extreme Drought Desert National Nature Reserve. + + + +Propeltidium. +Wider than long ( +Fig. 5D +), with a dense pubescence of thin, short, anteriorly directed fine setae; median eyes separated by less than one diameter; propeltidium lobe without eyespot. Anterior, posterior, and lateral edges with several long, curved spiniform setae perpendicular to the surface of the propeltidium. Ocular tubercle with a row of proximal spiniform setae, a row of short and two long middle distal spiniform setae. On each side of ocular tubercle, propeltidium with strong spiniform setae ( +Fig. 5D +). + + +Chelicerae. + +Fixed +finger with median teeth series comprising well-developed primary teeth ( +FP +, +FM +and +FD +), +FD +tooth size lesser, while +FM +and +FP +similar in size ( +Figs 5A–C +, +7A, B +); moreover, median teeth series comprising all secondary teeth (1 +FSD +and 2 +FSM +); secondary teeth overall size smaller than primary teeth, distal +FSM +smaller than proximal +FSM +. +Retrofondal +teeth series uninterrupted with five teeth (1 +RFP +, 1 +RFM +, 3 +RFA +), and profondal teeth series consisting of four teeth (1 +PFSP +, 1 +PFP +, 1 +PFSM +, 1 +PFM +) ( +Fig. 5C +). +Fixed +finger asetose area sinuous, with ventral margin predominantly linear in lateral aspect; prodorsal carina not sharp, not elevated in lateral aspect ( +Figs 5A–C +). +Cheliceral +fixed finger hornlike process curved with two teeth ( +Fig. 5C +), mucron without subterminal ( +FST +) teeth, and apex ( +FT +tooth) curved, hook-shaped ( +Fig. 5C +). +Movable +finger with median teeth series comprising well-developed +MP +and +MM +primary teeth, the former of which larger, and two +MSM +secondary tooth smaller than +MM +(i.e., +MP +> +MM +> +MSM +), movable finger one +MSM +teeth close to +MP +, another one close to +MM +, +MSM +upright and triangular-shaped; movable finger without subproximal ( +MSP +) or subterminal ( +MST +) teeth. ( +Figs 5A, B +) +Retrolateral +and dorsal surfaces with abundant retrolateral manus ( +rlm +) and retrolateral finger ( +rlf +) setae, of different sizes, which are predominantly straight and rigid; some of these setae arranged in bilaterally symmetrical pattern, as some principal retrolateral finger (principal +rlf +) setae that more flexible than others ( +Figs 5A +, +7A +); prolateral surface with array of setal +types +( +Figs 5B +, +7B +), as follows: proventral distal ( +pvd +) setae consisting of row of strong setae, starting at level of the flagellum and ending near level of +PFM +tooth; proventral subdistal ( +pvsd +) setae arranged in rather disorganized pattern, +pvsd +comb not markedly differentiated; carpet-like field of bristle-like promedial ( +pm +) setae broad. Stridulatory plate longer than high, occupying most of the prolateral surface of manus, above stridulatory plate two prodorsal proximal ( +pdp +) setae exist; stridulatory apparatus not obvious. Distal limit of the prolateral setose area of movable finger reaching midpoint between +MSM +and +MM +teeth; movable finger prodorsal ( +mpd +) setal series consisting of spinous setae arranged in a rather staggered row, adjacent to abundant non-plumose setae of the movable finger promedial ( +mpm +) and proventral ( +mpv +) setal series ( +Figs 5B +, +7B +). Flagellum typical of + +Eusimonia + +, bear two distinct, membranous or weakly sclerotized, sessile flagella: a broad, thin, dorsal secondary flagellum and a tubelike ventral primary flagellum ( +Figs 5A–C +, +7A, B +, +17A +); flagellum without flagellar complex subspiniform to spiniform ( +fcs +) setae; membranous dorsal flagellum (secondary flagellum) narrow triangular-shaped without central depression ( +Fig. 5C +) + +. + + + +FIGURE 3. +Phylogenetic results ML tree (A) and BI tree (B) from the COⅠ sequence. + + + + +FIGURE 4. + +Eusimonia turkestana +Kraepelin, 1899 + +. Habitus. +A, B. +Male (MHBU-Sol-GS2019071101). +C, D. +Female (MHBUSol-GS2023070101). Scale bars = 10 mm. + + + + +FIGURE 5. + +Eusimonia turkestana +Kraepelin, 1899 + +, male (MHBU-Sol-GS2023070601). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Flagellum, prolateral aspect. +D. +Propeltidium, dorsal aspect. +E. +Ctenidia on sternite III, ventral aspect. +F. +Ctenidia on sternite IV, ventral aspect. Scale bars = 1.0 mm (A, B, C); 0.5 mm (D, E, F). + + + +Opisthosoma. +The entire surface covered with almost adpressed fine setae, and numerous long, curved, bifurcate setae, and tergites with abundant setae. Sternite III with 5+5 coarse and spiny ctenidia reach the distal border of the sternite III ( +Figs 5E +, +7E +), sternite IV with a row of 13 long and thin peg-like ctenidia, almost reach the distal border of the sternite Ⅴ ( +Figs 5F +, +7F +). + + + +Pedipalps +. + +Entirely covered with short fine setae and long, thick setae. Tarsus without spines; posterior half of metatarsus thickens inward, resembling a hump, with six quite strong, distally diminishing, forward-inclined spines which thicker and flattened at distal ends ( +Figs 7C, D +, +16F, G +). + + + +FIGURE 6. + +Eusimonia turkestana +Kraepelin, 1899 + +, female (MHBU-Sol-GS2023070101). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Dentition on left cheliceral fixed finger, ventral aspect. +D. +Propeltidium, dorsal aspect. +E. +Genital operculum, ventral aspect. +F. +Ctenidia on sternite IV, ventral aspect. Scale bars = 1.0 mm (A, B, D); 0.5 mm (C, E, F). + + + +Legs. +Leg I coated with abundant short setae similar to those on pedipalps, without stout or spiniform setae; tibia and metatarsus with few long thin setae, tarsus without spiniform setae; with two small claws. Tibiae II and III with one distal spiniform setae dorsally and a pair of distal spiniform setae ventrally. Metatarsi II and III with a series of three dorsal spiniform setae and a pair of distal spiniform setae ventrally; metatarsus IV ventrally with two distal spiniform setae and three to five thick, spine-shaped bristles arranged in a prolateral row. Legs covered with long, thick setae and short fine setae. Distal part of claw short, occupying approximately 1/5 of the claw length. + + +Female +(MHBU-Sol-GS2023070101) + + +Measurements. +Total body length 12.78, +CH +1.62, CL 4.43, CW 1.46, PL 2.20, PW 3.06, A/CP 5.43, CL/ +CH +2.73. Pedipalp 11.80 (2.91, 3.13, 2.848, 1.19), leg Ⅰ 8.43 (2.48, 1.86, 1.47, 0.83, 0.05), leg II 8.69 (1.67, 1.76, 1.67, 0.47, 0.54), leg III 10.24 (2.23, 2.74, 1.34, 0.37, 0.51), leg IV 15.77 (3.72, 4.08, 1.56, 0.67, 1.05). + + +Coloration. +In 95% ethanol-preserved specimens ( +Figs 4C, D +). Coloration mostly like male, but opisthosoma general background light chartreuse. + + + +FIGURE 7. + +Eusimonia turkestana +Kraepelin, 1899 + +., male (A–F), female (G–J). +A. +Left chelicera, retrolateral aspect. +B. +Left chelicera, prolateral aspect. +C. +Pedipalpal metatarsus and tarsus, dorsal aspect. +D. +Pedipalpal metatarsus and tarsus, ventral aspect. +E. +Ctenidia on sternite III, ventral aspect. +F. +Ctenidia on sternite IV, ventral aspect. +G. +Left chelicera, retrolateral aspect. +H. +Left chelicera, prolateral aspect. +I. +Genital operculum, ventral aspect. +J. +Ctenidia on sternite IV, ventral aspect. + + + +Propeltidium. +Mostly the same as male, but female ocular tubercle spiniform setae on each side of ocular tubercle not as strong as in male ( +Fig. 6D +). + + + +Chelicera +. + +Fixed finger with median teeth series comprising well-developed primary teeth (FP, FM and FD), FD tooth size smaller, while FM and FP similar in size ( +Figs 6A–C +, +7G, H +); moreover, median teeth series comprising all secondary teeth (2FSD and 2 +FSM +); secondary teeth overall size smaller than primary teeth, distal FSD and +FSM +smaller than proximal FSD and +FSM +. Retrofondal teeth series uninterrupted with eight teeth (4RFSP, 1RFP, 1RFM, 2RFA); profondal teeth series consist of five teeth (1PFP, 1PFSM, 2PFSP, 1PFM) ( +Fig. 6C +). Fondal teeth graded as II, III, IV, I, VII, VI, VIII, V retrolaterally; II, I, III, V, IV prolaterally ( +Fig. 6C +). Fixed finger mucron without subterminal (FST) teeth. Movable finger with median teeth series comprising well-developed MP and MM primary teeth, the former of which larger, and three MSM and two MSP secondary teeth that smaller than MM (i.e., MP> MM> MSM> MSP), movable finger MSM teeth located MM and MP center, MST teeth located MM front, and MSP located MP after; all secondary teeth upright and triangular-shaped ( +Figs 6A, B +, +7G, H +). Cheliceral setose areas and stridulatory plate mostly like male, but female proventral distal ( +pvd +) setae only consisting of row of plumose setae ( +Figs 6B +, +7H +). + + +Opisthosoma. +The entire surface covered with almost adpressed fine setae and numerous long, curved, bifurcate setae. Genital operculum with clear demarcation between the plates, distal border widened, resembling a triangle shape. Genital opening not visible between plates or distal to plates. ( +Figs 6E +, +7I +). 15 long needle-like ctenidia on sternite IV extending the length of the following sternite ( +Figs 6F +, +7J +). + + +Pedipalps. +Entirely covered with short fine setae and long, thick setae and without spines. + + + +Legs +. + +Same as in the male. + + + + +Distribution and habitat. + +E. turkestana + +is widely distributed in the arid regions of northwest +China +( +Nei Mongol +, +Xinjiang +), +Kazakhstan +and +Mongolia +. Habitat: Gobi Desert ( +Fig. 2F +) and desert steppe ( +Fig. 2B +). As +Gromov (2003) +mentioned, + +E. turkestana + +live in clayey, stony or sandy loam deserts at low altitudes. + + +Collection method. +During the night using an ultra-violet lamp. Under ultraviolet light, +Solifugae +reflects a faint blue glow, making it easy to spot and catch. Given their nocturnal habits, we also collect specimens using pitfall traps. + + +Variability. +Males. Total length 10.56–15.64. Body coloration light yellow to yellow. Chelicerae with manus yellowish to brown. The number of cheliceral fixed finger fondal teeth 9–11 (profondal teeth 4–5; retrofondal teeth 5–6). The number of ctenidia on sternite IV 13–15. Females. Total length 11.72–16.36. Variability of body coloration as in males. The number of cheliceral fixed finger fondal teeth 12–14 (profondal teeth 4–5; retrofondal teeth 7–9), MST 0–2, MSP 2–3. The number of ctenidia on sternite IV 14–15. + + + + +Remark. +We matched both sexes based on DNA barcoding data, with a genetic distance between male and female voucher specimens ranging from 0%–4.11% ( +Table 1 +). No distinct morphological differences were found among individuals of the same sex. After examining +30 adult +specimens, we discovered a high intraspecific mitochondrial divergence in + +E. turkestana + +. Given that this species has a widespread distribution across Central Asia, this variability is especially prominent among specimens from various locations, indicating potential longterm adaptation and evolutionary processes within distinct geographical regions. The identification of this genetic diversity holds significant implications for comprehending the species' biogeographical distribution, population genetic structure, and evolutionary trajectory. + + + + \ No newline at end of file diff --git a/data/87/4E/87/874E87E1413EFF929FA76F8FFD8AE140.xml b/data/87/4E/87/874E87E1413EFF929FA76F8FFD8AE140.xml new file mode 100644 index 00000000000..0545e1fabe2 --- /dev/null +++ b/data/87/4E/87/874E87E1413EFF929FA76F8FFD8AE140.xml @@ -0,0 +1,355 @@ + + + +A revision of Chinese endemic Neptis hesione Leech, 1890 with a new species described from Hainan Island (Lepidoptera, Nymphalidae) + + + +Author + +Li, Hua-Zhao +The People’s Government of Dusong Township, Jinchuan County, Aba, Sichuan, P. R. China + + + +Author + +Liu, Zhe +Nanfanghangkongcheng, Sanya, Hainan, P. R. China + + + +Author + +Li, Jia-Ling +0000-0003-3560-4798 +19071300110003@hainanu.edu.cn + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +431 +439 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.9 + +journal article +10.11646/zootaxa.5492.3.9 +1175-5326 +30D78760-7670-49BE-8C8E-87B48E117A83 + + + + + + + +Neptis kejiaae +H.Z. Li & Z. Liu + +sp. nov. +(Chinese name: 可佳ųợø) + + + + + +urn:lsid:zoobank.org:act: +7A667364-D3C5-4F79-BA10-5182DF6E4270 + + + + + +( +Figs. 1 +for adults, 3 for genitalia, 4 for diagnosis, 5 for distribution, 6 for phylogeny) + + + + + + +Holotype +: + +♂ +, +Mt. Wuzhi +, +Wuzhishan +, +Hainan +, +China +, ca. + +800 m + +, + +17.IV.2024 + +, +Z. Liu +leg. (NK1; will be deposited at +CMNH +) + +. + + +Paratype +: + +♂ +, collection data same as the holotype (NK2; +CZL +) + +. + + + + +Description: Male. +( +Fig. 1 +) Ground color black on upperside and brown on underside. Wing markings yellow, tinged with orange in upperside and lighter in underside. Forewing upperside: postdiscal spots composed of three bands, lower clavate band from basal part of spaces 1a to 1b, middle round band from spaces 1b to 3, upper ellipse band from spaces 4 to 6; lower and midle postdiscal bands almost contact with each other; subcostal small spots present in spaces 8 and 9, beyond upper postdiscal brand; cell streak with basal part laterally dusted and with distal part pointed. Hindwing upperside: discal band wide, basally whitish and turned into gray when passing through speculum; postdiscal band narrow, interrupted by black veins. Forewing underside: wing patterns similar to upperside but yellow markings broader; a submarginal yellow fascia running from spaces 8 to 3, ending at middle postdiscal brand; another submarginal fascia, outside preceding one, passing from costa to dorsum, tinged with lilac. Hindwing underside: subbase decorated with a yellow streak of an irregular shape; discal band wider, with a black line embedded in its lower edge; postdiscal band wider accompanied by yellow submarginal fascia, with their distal portions overlaid with lilac; ground color between discal and postdiscal bands dark-brown. Speculum on hindwing upperside and corresponding polished area on forewing underside dark. Fringes chequered black and white. +Male Genitalia. +( +Fig. 3 +) Vinculum: slender, triple as long as saccus, ventrally broad. Saccus: from the part connected to vinculum, continuously narrowed forward, ending with a sharp point. Tegumen: dorsally elliptical, cephalic part protruding. Gnathos: vertically elongated, ventral portion integrated and protruding caudally. Uncus: straight, elongated into a pointed hook, slightly down-curved. Valva: more or less tongue-shaped with ampulla and harpe terminally differentiated; harpe wide, blade-like, gradually curved upward into a blunt end; ampulla fused with costa, a hump present in middle rear, caudal end of ampulla narrower and somewhat shorter than harpe with inconspicuous upper teeth. Juxta: bilateral parts expansive posteriorly, elliptical. Phallus: almost straight; basally expansive and terminal end forming a sharp upward hook; cornutus with tiny spins in expanded caudal part. + + + + +Diagnosis. +The new species can be distinguished from other members in the + +N. hesione + +-group by the following characters ( +Fig. 4 +): + + + +FIGURE 2. +Holotype of + +Neptis hesione + +and the related literatures. + + + + +FIGURE 3. +Male genitalia of + +Neptis kejiaae + + +sp. nov. + +le = lateral left view, ds = dorsal view, vs = ventral view. + + + + +FIGURE 4. +Diagnosis of + +Neptis kejiaae + + +sp. nov. + +and other species in + +N. hesione + +-group. + + +(1) Wing markings are more developed than other species in this group. + +(2) The color of yellow markings is more intense, tinged with orange, but lighter in + +N. hesione + +and whiter in + +N. podarces + + +stat. rev. + + +(3) Male brands on the hindwing speculum are darker than other species in this group. + +(4) Terminal hook of the ampulla is almost absent as in + +N. podarces + + +stat. rev. + +, whereas it is present in + +N. hesione + +. + + + + +Distribution. +Endemic to +Hainan +Is., +China +. + + + + +Etymology. +The specific name is dedicated to Ms. Ke-Jia Sun, the wife of the second author, who encourages him to continue surveys on the butterfly fauna of +Hainan +. + + + + +Ecology. +Both the new species and + +N. h. +hesione + +are sympatric with + +Pantoporia bieti +( +Oberthür, 1894 +) + +, a smaller species from a different genus but with a similar appearance ( +Oberthür 1894 +). + + + + +FIGURE 5. +Distributional map of + +Neptis hesione + +-group. The type localities are represented by the letter T. Source from references in Material and methods. + + + + +FIGURE 6. +Phylogenetic tree of + +Neptis hesione + +-group inferred from ML analysis of +COI +with bootstrap support values (SHaLRT support/ultrafast bootstrap support). Mainly modified from + +Wu +et al. +(2016) + +and + +Ma +et al. +(2020) + +. + + + + +Remarks. +At present, a total of seven yellow-type + +Neptis +species + +are known from +Hainan +, six of which are described as endemic, including + +N. ananta yanagisawai +Sugiyama, 1992 + +, + +N. noyala qionga +Gu & Wang, 1994 + +, + +N. namba leechuanlungi +Wang & Lang, 2009 + +, + +N. manasa neohainana +Lang, Xue & Han, 2009 + +, + +N. armandia leishuaimini +Li & Liu, 2023 + +, and + +N. kejiaae + + +sp. nov. + +( +Sugiyama 1992 +; +Chou 1994 +; Lang 2009; 2012; + +Li +et al. +2023 + +). All of those yellow populations are extremely rare and hardly rediscovered, appearing to be threatened except +N. a. yanagisawai +(Lang +et al. +2009). + + + + \ No newline at end of file diff --git a/data/B3/6D/06/B36D0606FFD0AC1CFF28F940FD20FB11.xml b/data/B3/6D/06/B36D0606FFD0AC1CFF28F940FD20FB11.xml new file mode 100644 index 00000000000..167986085ce --- /dev/null +++ b/data/B3/6D/06/B36D0606FFD0AC1CFF28F940FD20FB11.xml @@ -0,0 +1,306 @@ + + + +Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec & Ollivier) with notable male genital morphology (Carabidae: Trechini) + + + +Author + +Schmidt, Joachim +General and Systematic Zoology, University of Rostock, Germany + + + +Author + +Merene, Yeshitla +Department of Zoological Sciences, Addis Ababa University, Ethiopia & Amhara Agricultural Research Institute, Bahir Dar, Ethiopia + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +343 +355 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.3 + +journal article +10.11646/zootaxa.5492.3.3 +1175-5326 +13234851 +8F81F053-DE78-4684-BCC9-875AE0C41615 + + + + + + + +Trechus +( +Abyssinotus +) +kniphofia +Schmidt + +, +sp. n. + + + + + + +Fig. 2C +, +3B, E, J + + + + +Type material. + +Holotype +: +♂ +, with label data: +Ethiopia +, +Amhara +, E-slope +Mt. Choke +, +Wondasha Guskuam +, alt. + +3650 m + +, +10°41‘05“N +37°59‘21“E +, + +2.V.2022 + +, leg. +J. Schmidt +, +Yeshitla, M. +( +CSCHM +) + +. + +Paratypes +: +4 ♂♂ +, +5 ♀♀ +, with same data as holotype ( +CSCHM +) + +. + +1 ♂ +, +3 ♀♀ +, +Mt. Choke +, eastern crater valley, alt. + +3700–3800 m + +, +10°42‘59“N +37°54‘13“E +, + +06. +V + + +.2022, leg. +J. Schmidt +, +Yeshitla M. +( +CSCHM +); + +3 ♂♂ +, +2 ♀♀ +, NE-slope +Mt. Choke +, above +Felege Birhan +, alt. + +3750–3850 m + +, +10°42‘13“N +37°56‘32“E +, + +30.IV.2022 + +, leg. +J. Schmidt +, +Yeshitla M. +( +CSCHM +) + +. + + + + +Etymology +. The new species is named after the beautiful red-hot poker +Kniphofia foliosa +which is a common +Asphodelaceae +at sites where + +T. kniphofia + +sp. n. +occurs. + + + + +Description +. Body length: +2.3–2.8 mm +. + +Proportions (n = 10): PW/HW = 1.36–1.44 (Ø = 1.40); PW/PL = 1.32–1.44 (Ø = 1.39); PW/PBW = 1.18–1.22 (Ø = 1.20); PBW/PAW = 1.16–1.29 (Ø = 1.22); EW/PW = 1.41–1.51 (Ø = 1.46); EL/EW = 1.31–1.38 (Ø = 1.35). + +Colour: Head, pronotum and elytra blackish brown; mandibles, palps and legs yellowish brown; antennae usually monochrome, in some specimens, antennae slightly darkened beginning from second or third antennomere; in most specimens 2 +nd +maxillary palpomere darkened. + +Microsculpture: Sculpticells on frons and supraorbital area large, moderately impressed, almost isodiametric, on clypeus more finely impressed. Sculpticells on pronotal disc moderately impressed, slightly transverse. Elytra with large and deeply impressed, almost isodiametric or somewhat irregularly formed sculpticells. + +Head: As described in + +T. yitbareki + +sp. n. + + +Prothorax: As described in + +T. yitbareki + +sp. n. + + +Pterothorax: Striae 1–3 very slightly impressed or indistinct, smooth, first stria finer impressed than second or absent. In all other characters as described in + +T. yitbareki + +sp. n. + + +Legs: Two basal protarsomeres of males moderately dilated and dentoid at the inner apical border ( +Fig. 3B +). In all other characters as described in + +T. yitbareki + +sp. n. + + +Male genitalia: EL/AL = 2.02–2.38 (Ø = 2.23; n = 8). Aedeagal median lobe robust, in lateral view almost evenly bent throughout, straight near apex, with short apical lamella which is hook-like bent dorsally ( +Fig. 3E +); in dorsal view with side margins almost parallel, very slightly widened in apical quarter and shortly narrowed towards slender apical lamella ( +Fig. 3J +). Dorsal opening extends basally near to the apical margin of basal bulb. Sagittal aileron absent. Endophallus symmetrical in dorsal view, without distinct copulatory pieces, densely covered by tiny scales, in the inactive position with a sac-like folding structure near its base and an area covered by large sclerotized scales on its dorsal side ( +Fig. 3F +). Parameres similar in size and shape, rather slender, each with four apical setae. + + + + +Differential diagnosis +. In habitus and male genital characters very similar to + +T. reebae + +and + +T. yitbareki + +sp. n. +, but differs in the shape of the basal two protarsomeres of males which are moderately dilated ( +Fig. 3B +; markedly dilated in the latter species, see +Fig. 3A, C +). The median lobe of the aedeagus is shorter and more evenly bent than in + +T. yitbareki + +sp. n. +, and is characterized by a shorter apical lamella. + +T. kniphofia + +sp. n. +differs from the likewise similar + +T. igori + +sp. n. +in the more robust and, on an average, larger body, and in the larger aedeagus with longer, hook-like bent apical lamella. + + + + +Distribution +. Endemic to Mt. Choke in northern +Ethiopia +. The new species is probably the eastern vicariant of + +T. reebae + +sp. n. +, and was found along slopes on eastern margins of the crater rim ( +Fig. 4 +). + + +Habitat preferences +. As described in + +T. reebae + +. + + + + \ No newline at end of file diff --git a/data/B3/6D/06/B36D0606FFD1AC12FF28FAE0FE37FDE1.xml b/data/B3/6D/06/B36D0606FFD1AC12FF28FAE0FE37FDE1.xml new file mode 100644 index 00000000000..6300f2acaab --- /dev/null +++ b/data/B3/6D/06/B36D0606FFD1AC12FF28FAE0FE37FDE1.xml @@ -0,0 +1,321 @@ + + + +Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec & Ollivier) with notable male genital morphology (Carabidae: Trechini) + + + +Author + +Schmidt, Joachim +General and Systematic Zoology, University of Rostock, Germany + + + +Author + +Merene, Yeshitla +Department of Zoological Sciences, Addis Ababa University, Ethiopia & Amhara Agricultural Research Institute, Bahir Dar, Ethiopia + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +343 +355 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.3 + +journal article +10.11646/zootaxa.5492.3.3 +1175-5326 +13234851 +8F81F053-DE78-4684-BCC9-875AE0C41615 + + + + + + + +Trechus +( +Abyssinotus +) +igori +Schmidt + +, +sp. n. + + + + + + +Fig. 2D +, +3G, K + + + + +Type material. + +Holotype +: +♂ +, with label data: +Ethiopia +, +Amhara +, W-slope +Mt.Choke +, + +3370 m + +, + +23.II.2019 + +, +10°38’07”N +37°45’51”E +, leg. +D. Hauth +, J. Schmidt, +Yeshitla M. +, +Yitbarek, W. +( +CSCHM +) + +. + +Paratypes +: +8 ♂♂ +, +7 ♀♀ +, with same data as holotype ( +CSCHM +) + +. + + + + +Etymology +. The new species is dedicated to my dear friend and colleague, the distinguished carabidologist and explorer of the high altitude environments, Igor A. Belousov, St. Petersburg. + + + + +Description +. Body length: +2.2–2.3 mm +. + +Proportions (n = 10): PW/HW = 1.31–1.41 (Ø = 1.36); PW/PL = 1.41–1.50 (Ø = 1.44); PW/PBW = 1.23–1.30 (Ø = 1.27); PBW/PAW = 1.07–1.12 (Ø = 1.10); EW/PW = 1.44–1.53 (Ø = 1.50); EL/EW = 1.30–1.35 (Ø = 1.32). +Colour: Head capsule, pronotum and elytra dark brown, pronotum paler in some specimens; clypeus, labrum, mandibles, palps, antennae and legs yellowish brown. +Microsculpture: Sculpticells on frons and supraorbital area large, moderately impressed, almost isodiametric, slightly transverse on clypeus. Sculpticells on pronotal disc and elytra moderately impressed, transverse. + +Head: As described in + +T. yitbareki + +sp. n. + + +Prothorax: Pronotum moderately large, transverse, sub-discoidal, with base slightly more markedly narrowed towards base than in the proceeding species. In all other characters as described in + +T. yitbareki + +sp. n. + + + +FIGURE 3. +Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of + +Trechus +( +Abyssinotus +) + +species. +A +, +D +, +I +, + +T. reebae +( +Quéinnec & Ollivier, 2021 +) + +, specimen from western crater valley. +B +, +E +, +J +, + +T. kniphofia +Schmidt + +, +sp. n. +, paratype. +C +, +F +, +H +, + +T. yitbareki +Schmidt + +, +sp. n. +, paratype. +G +, +K +, + +T. igori +Schmidt + +, +sp. n. +, paratype. + + + +Pterothorax: First stria absent in anterior 2/3, suggestively impressed before its apex, striae 2 and 3 very finely impressed, indistinct, external striae absent apart from the apical portion of stria 8. In all other characters as described in + +T. yitbareki + +sp. n. + + +Legs: As in + +T. kniphofia + +sp. n. + + +Male genitalia: EL/AL = 2.20–2.30 (Ø = 2.24; n = 8). Aedeagal median lobe moderately robust, rather stout, in lateral view evenly curved throughout, with apical lamella very short, very slightly sclerotized, very shortly bent upwardly ( +Fig. 3G +). Median lobe in dorsal view with side margins almost parallel, shortly narrowed towards apical lamella ( +Fig. 3K +). Dorsal opening extends basally near to the apical margin of basal bulb. Sagittal aileron absent. Endophallus as described in + +T. kniphofia + +sp. n. +Parameres similar in size and shape, slender, each with 3–4 apical setae. + + + + +Differential diagnosis +. On average, + +T. igori + +sp. n. +is the smallest species of the + +T. reebae + +group. It differs from all other species of the group by the transverse pattern of elytral microsculpture (isodiametric or somewhat irregularly shaped in the other species) and the very slightly sclerotized apex of the aedeagal median lobe. It additionally differs from + +T. niloticus + +, + +T. reebae + +and + +T. yitbareki + +sp. n. +in the less markedly dilated basal protarsomeres in the male sex. Due to its small body length and shallower elytral striae, + +T. igori + +sp. n. +is similar to + +T. abyssinicus + +and + +T. afroalpinus + +, however, it is easily distinguished by two basal protarsomeres dilated in males (one in the latter species). + + + + +Distribution +. Endemic to Mt. Choke in northern +Ethiopia +. The new species was only found near the top of a mountain ridge that extends from the crater rim to the west were it occurs sympatric with + +T. apertus + +sp. n. +and + +T. yitbareki + +sp. n. +( +Fig. 4 +). + + +Habitat preferences +. The new species was found syntopic with + +T. apertus + +sp. n. +and + +T. yitbareki + +sp. n. +; see above for details of the habitat. + + + + \ No newline at end of file diff --git a/data/B3/6D/06/B36D0606FFD2AC1DFF28FB45FF28F9A9.xml b/data/B3/6D/06/B36D0606FFD2AC1DFF28FB45FF28F9A9.xml new file mode 100644 index 00000000000..847d263649f --- /dev/null +++ b/data/B3/6D/06/B36D0606FFD2AC1DFF28FB45FF28F9A9.xml @@ -0,0 +1,350 @@ + + + +Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec & Ollivier) with notable male genital morphology (Carabidae: Trechini) + + + +Author + +Schmidt, Joachim +General and Systematic Zoology, University of Rostock, Germany + + + +Author + +Merene, Yeshitla +Department of Zoological Sciences, Addis Ababa University, Ethiopia & Amhara Agricultural Research Institute, Bahir Dar, Ethiopia + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +343 +355 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.3 + +journal article +10.11646/zootaxa.5492.3.3 +1175-5326 +13234851 +8F81F053-DE78-4684-BCC9-875AE0C41615 + + + + + + + +Trechus +( +Abyssinotus +) +yitbareki +Schmidt + +, +sp. n. + + + + + + +Fig. 2B +, +3C, F, H + + + + +Type material. + +Holotype +: +♂ +, with label data: +Ethiopia +, +Amhara +, W-slope +Mt.Choke +, + +3370 m + +, + +23.II.2019 + +, +10°38‘07“N +37°45‘51“E +, leg. +D. Hauth +, J. Schmidt, +Yeshitla M. +, +Yitbarek, W. +( +CSCHM +) + +. + +Paratypes +: +33 ♂♂ +, +17 ♀♀ +, with same data as holotype ( +NHMAA +, +ZSM +, +CAF +, +CSCHM +) + +. + +48 ♂♂ +, +31 ♀♀ +, W-slope +Mt. Choke +, + +3450 m + +, + +22.II.2019 + +, +10°38‘09“N +37°46‘06“E +, leg. +D. Hauth +, +J. Schmidt +, +Yeshitla M. +, +Yitbarek, W. +( +NHMAA +, +ZSM +, +CSCHM +) + +; of these, + +1 ♂ +(DNA voucher +SMNS-L1821 +) + +was used for molecular genetic study ( + +Faille +et al. +2023 + +). + + + + +Etymology +. The new species is dedicated to one of its collectors, Yitbarek Woldehawariat, professor and entomologist at the Department of Zoological Science, University of +Addis Ababa +. + + + + +Description +. Body length: +2.4–2.8 mm +. + +Proportions (n = 10): PW/HW = 1.39–1.47 (Ø = 1.43); PW/PL = 1.33–1.42 (Ø = 1.37); PW/PBW = 1.18–1.22 (Ø = 1.20); PBW/PAW = 1.21–1.29 (Ø = 1.24); EW/PW = 1.41–1.51 (Ø = 1.46); EL/EW = 1.28–1.35 (Ø = 1.31). + +Colour: Head, pronotum and elytra blackish brown, pronotum sometimes more reddish; mandibles, palps, antennal base and legs yellowish brown; antennae darkened beginning from second or third antennomere; in most specimens 2 +nd +maxillary palpomere darkened. + +Microsculpture: Sculpticells on frons and supraorbital area large, moderately impressed, almost isodiametric, on clypeus very finely impressed, slightly transverse. Sculpticells on pronotal disc moderately impressed, slightly transverse. Elytra with large and deeply impressed, almost isodiametric or somewhat irregularly formed sculpticells. + +Head: Compound eyes rather small, moderately convex; tempora about 4/5 of length of eyes. In all other characters as described in + +T. apertus + +sp. n. + + + +FIGURE 2. +Habitus photographs of + +Trechus +( +Abyssinotus +) + +species; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. +A +, + +T. reebae +( +Quéinnec & Ollivier, 2021 +) + +, specimen from western crater valley. +B +, + +T. yitbareki +Schmidt + +, +sp. n. +, paratype. +C +, + +T. kniphofia +Schmidt + +, +sp. n. +, paratype. +D +, + +T. igori +Schmidt + +, +sp. n. +, paratype. + + +Pronotum: Pronotum large and transverse, sub-discoidal, broadest slightly before middle, with base distinctly broader than apex; disc moderately convex.Anterior margin slightly concave, with anterior angles slightly protruded anteriorly, rounded; basal margin convex, with laterobasal angles markedly shifted anteriorly; lateral margin almost evenly rounded throughout; laterobasal angles obtuse, sometimes indistinct. Marginal gutter narrow, extending towards laterobasal foveae, slightly widened between laterobasal angle and foveae. Median longitudinal impression finely incised, disappearing near apex and base; anterior transverse impression indistinct, smooth, posterior transverse impression broad and very shallow, smooth; laterobasal foveae small, moderately deep, smooth. Lateral and laterobasal setae present, with the former situated at the end of anterior pronotal third. +Pro- and mesothorax: Pro- and mesepisternum glabrous and smooth. + +Pterothorax: Elytra short, ovate, with disc convex, in dorsal view broadest in middle, humerus rounded, subapical sinuation indistinct, apex rounded. Striae 1–3 slightly impressed, indistinct in some specimens, smooth or suggestively punctate, external striae absent apart from the apical portion of stria 8 which is distinctly deepened before apex; parascutellar stria absent. Recurrent preapical stria short, deep, with outer curvature directed towards the fifth stria. Parascutellar seta present; three discal setae located in the 3rd interval, adjoined to the 3rd stria: anterior seta located near the end of the anterior elytral 8th; middle seta located anterior of elytral middle, near the end of elytral 3/7; posterior seta (= subapical seta) located about 1/8 of elytral length from elytral apex; seta of the recurrent stria isolated, removed from this stria by distance of the diameter of the pore. Number and positions of the setae of the marginal umbilicate series as normal for + +Trechus + +s. l. +Metepisternum very short, glabrous and smooth, with outer margin about as long as anterior margin. + + +Legs: Moderately short and robust. Protibia dilated towards apex, straight on external margin, its dorsal surface with longitudinal groove developed but very flat, without micro-setae on anterior surface. Two basal protarsomeres of males markedly dilated and dentoid at the inner apical border ( +Fig. 3C +). + + +Male genitalia: EL/AL = 1.77–1.89 (Ø = 1.82; n = 8). Aedeagal median lobe robust, in lateral view markedly bent in basal half, almost straight towards apex, with moderately long apical lamella hook-like bent dorsally ( +Fig. 3F +); in dorsal view with side margin almost parallel, very slightly widened in apical quarter and shortly narrowed towards slender apical lamella ( +Fig. 3H +). Dorsal opening extends basally near to the apical margin of basal bulb. Sagittal aileron absent. Endophallus symmetrical in dorsal view, without a distinct copulatory piece, densely covered by tiny scales, in the inactive position with a sac-like folding structure near its base and an area covered by large sclerotized scales on its dorsal side ( +Fig. 3F +). Parameres similar in size and shape, rather slender, each with four apical setae. + + + + +Differential diagnosis +. Externally, + +T. yitbareki + +sp. n. +is very similar to + +T. reebae + +, but differs in the proportionally longer median lobe of the aedeagus (EL/AL < +1.9 in + +T. yitbareki + +sp. n. +,> 2.0 in = + +T. reebae + +) and in the longer apical lamella ( +Fig. 3F +). The new species differs from the likewise similar + +T. igori + +sp. n. +and + +T. kniphofia + +sp. n. +, prominently in the more markedly dilated protarsomeres 1 and 2 ( +Fig. 3C +), and additionally in the more robust body which is, on an average larger, and in the larger aedeagus with longer apical lamella which is more markedly hook-like bent upwardly. + + + + +Distribution +. Endemic to Mt. Choke in northern +Ethiopia +. The new species was only found near the top of a mountain ridge that extends from the crater rim to the west were it occurs sympatric with + +T. apertus + +sp. n. +and + +T. igori + +sp. n. +( +Fig. 4 +). + + +Habitat preferences +. The new species was found syntopic with + +T. apertus + +sp. n. +; see above for details of the habitat. + + + + \ No newline at end of file diff --git a/data/B3/6D/06/B36D0606FFD2AC1FFF28FEFCFB9AFBB6.xml b/data/B3/6D/06/B36D0606FFD2AC1FFF28FEFCFB9AFBB6.xml new file mode 100644 index 00000000000..1cdc2eddb04 --- /dev/null +++ b/data/B3/6D/06/B36D0606FFD2AC1FFF28FEFCFB9AFBB6.xml @@ -0,0 +1,288 @@ + + + +Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec & Ollivier) with notable male genital morphology (Carabidae: Trechini) + + + +Author + +Schmidt, Joachim +General and Systematic Zoology, University of Rostock, Germany + + + +Author + +Merene, Yeshitla +Department of Zoological Sciences, Addis Ababa University, Ethiopia & Amhara Agricultural Research Institute, Bahir Dar, Ethiopia + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +343 +355 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.3 + +journal article +10.11646/zootaxa.5492.3.3 +1175-5326 +13234851 +8F81F053-DE78-4684-BCC9-875AE0C41615 + + + + + + + +Trechus +( +Abyssinotus +) +reebae +( +Quéinnec & Ollivier, 2021 +) + + + + + + + +Fig 2A +, +3A, F, I + + +Citations. + +Nilotrechus reebae +Quéinnec & Ollivier + +: + +Quéinnec +et al. +(2021) + +: 55. + + + + + + +Trechus +( +Abyssinotus +) +reebae +(Quéinnec & Ollivier) + +: + + +Merene +et al. +(2023) + +: 49 + +. + + + + + + +Type +material. + +Not studied. Identification is based on the original description, including habitus and male genital figures ( + +Quéinnec +et al. +2021 + +) as well as on several +additional specimens +collected close to the +type +locality. + + +Additional material. + +12 ♂♂ +, +14 ♀♀ +, +Ethiopia +, +Amhara +, +Mt. Choke +, western crater valley, alt. + +3700–3800 m + +, +10°41’14”N +37°50’07”E +, + +24.II.2019 + +, leg. +D. Hauth +, +J. Schmidt +, +Yeshitla M. +, +Yitbarek, W. +( +NHMAA +, +CAF +, +CSCHM +) + +. + +22 ♂♂ +, +11 ♀♀ +, +Mt. Choke +, western crater valley, alt. + +3500–3600 m + +, +10°41’00”N +37°50’35”E +, + +01. +V + + +.2022, leg. +J. Schmidt +, +Yeshitla M. +( +CSCHM +). + +11 ♂♂ +, +4 ♀♀ +, N-slope +Mt. Choke +, +N of Waber +, alt. + +3450–3600 m + +, +10°44’48”N +37°46’22”E +, + +07. +V + + +.2022, leg. +J. Schmidt +, +Yeshitla M. +( +CSCHM +). + + + + +Description. +See + +Quéinnec +et al. +(2021: 55–56) + +. Proportion EL/AL = 2.01–2.12 (Ø = 2.05; n = 8). + + + + +Diagnosis. +See key to species, below. + + + + +Distribution +. Endemic to Mt. Choke in northern +Ethiopia +. Based on + +Quéinnec +et al. +(2021) + +and new sampling data, the species occurs on top of the western crater rim and on western slope of the crater valley, as well as on the north-facing mountain range north of the village Waber ( +Fig. 4 +). + + +Habitat preferences +. + +Quéinnec +et al. +(2021) + +collected most specimens of the +type +series under clay at the base of a small cliff. The additional specimens of + +T. reebae + +were collected under stones and sifted from the humus-rich soil under dense vegetation alongside small streams, and no specimens were found on open pastures and in dry bushland areas. For the vertical distribution altitudes of +3500 to 3900 m +were noted (see also + +Quéinnec +et al. +2021 + +). Based on these data, the new species can be considered hygrophilic and cold-adapted. + + + + \ No newline at end of file diff --git a/data/B3/6D/06/B36D0606FFD7AC1FFF28FC99FC78FF2D.xml b/data/B3/6D/06/B36D0606FFD7AC1FFF28FC99FC78FF2D.xml new file mode 100644 index 00000000000..a33e46d2e93 --- /dev/null +++ b/data/B3/6D/06/B36D0606FFD7AC1FFF28FC99FC78FF2D.xml @@ -0,0 +1,388 @@ + + + +Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec & Ollivier) with notable male genital morphology (Carabidae: Trechini) + + + +Author + +Schmidt, Joachim +General and Systematic Zoology, University of Rostock, Germany + + + +Author + +Merene, Yeshitla +Department of Zoological Sciences, Addis Ababa University, Ethiopia & Amhara Agricultural Research Institute, Bahir Dar, Ethiopia + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +343 +355 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.3 + +journal article +10.11646/zootaxa.5492.3.3 +1175-5326 +13234851 +8F81F053-DE78-4684-BCC9-875AE0C41615 + + + + + + + +Trechus +( +Abyssinotus +) +apertus +Schmidt + +, +sp. n. + + + + + + +Fig. 1 + + + + +Type material. + +Holotype +: +♂ +, with label data: +Ethiopia +, +Amhara +, W-slope +Mt.Choke +, + +3370 m + +, + +23.II.2019 + +, +10°38’07”N +37°45’51”E +, leg. +D. Hauth +, J. Schmidt, +Yeshitla M. +, +Yitbarek, W. +( +CSCHM +) + +. + +Paratypes +: +8 ♂♂ +, +7 ♀♀ +, with same data as holotype ( +CSCHM +) + +. + +4 ♂♂ +, +5 ♀♀ +, W-slope +Mt. Choke +, + +3450 m + +, + +22.II.2019 + +, +10°38’09”N +37°46’06”E +, leg. +D. Hauth +, J. Schmidt, +Yeshitla M. +, +Yitbarek, W. +( +NHMAA +, +CAF +, +CSCHM +) + +. + + + + +Etymology +. The species’ epithet refers to the extraordinary wide dorsal opening of the aedeagal median lobe which is a morphological peculiarity of the new species. + + + + +Description +. Body length: +2.5–2.8 mm +. + +Proportions (n = 12): PW/HW = 1.35–1.44 (Ø = 1.41); PW/PL = 1.37–1.45 (Ø = 1.40); PW/PBW = 1.18–1.22 (Ø = 1.20); PBW/PAW = 1.18–1.24 (Ø = 1.21); EW/PW = 1.38–1.46 (Ø = 1.43); EL/EW = 1.24–1.30 (Ø = 1.27). + +Colour: Head, pronotum and elytra blackish brown; mandibles, palps, antennal base and legs yellowish brown; antennae slightly darkened beginning from second antennomere; in most specimens 2 +nd +maxillary palpomere somewhat darkened; elytral apical margin very narrowly yellowish. + +Microsculpture: Sculpticells on frons and supraorbital area large, moderately impressed, almost isodiametric; sculpticells on clypeus very finely impressed, slightly transverse. Sculpticells on pronotal disc smaller and finer impressed than on head disc, slightly transverse. Elytra with large and deeply impressed, almost isodiametric sculpticells throughout. + +Head: Moderately robust. Mandibles short, with right mandible tridentate where all teeth are almost the same length, without diastema, without recognizable separation of premolar and retinaculum ( +Fig. 1B +); tooth of left mandible with two blunt cusps ( +Fig. 1C +). Penultimate labial palpomere with three setae ( +Fig. 1F +). Labrum with apical margin moderately emarginate, with six setae near apical margin ( +Fig. 1D +). Clypeus each side with two setae. Compound eyes rather small, slightly convex; tempora about ¾ of length of eyes, markedly convex, distinctly wrinkled to the neck. Dorsal surface of head with supraorbital area and frons markedly convex, and with supraorbital furrows deeply impressed, evenly curved. Two supraorbital setae and one suborbital seta each side. Tempora distinctly pubescent. Antennae short, with second antennomere about as long as scape, and with third antennomere shorter than second. + + + +FIGURE 1. + +Trechus +( +Abyssinotus +) +apertus +Schmidt + +, +sp. n. +, holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). +A +, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. +B +, right mandible, ventral aspect. +C +, left mandible, ventral aspect. +D +, labrum, dorsal aspect. +E +, left maxilla. +F +, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. +G +, aedeagus, left lateral view. +H +, aedeagus, dorsal view. + + + +Pronotum: Large and transverse, sub-discoidal, broadest slightly before middle, with base distinctly broader than apex; disc moderately convex. Anterior margin slightly concave (almost straight in middle), with anterior angles slightly protruded anteriorly, rounded; basal margin convex, with laterobasal angles markedly shifted anteriorly; lateral margin almost evenly rounded throughout; laterobasal angles very obtuse, indistinct ( +Fig. 1A +). Marginal gutter narrow, slightly widened near laterobasal angle, extending towards laterobasal foveae. Median longitudinal impression very finely incised, disappearing near apex; anterior and posterior transverse impressions indistinct, smooth; laterobasal foveae small, moderately deep, smooth. Lateral and laterobasal setae present, with the former situated at the end of anterior pronotal third. + +Pro- and mesepisternum: Glabrous and smooth. + +Pterothorax: Elytra short, ovate, with disc convex, in dorsal view broadest in middle, humerus rounded but distinct, subapical sinuation indistinct, apex rounded. Striae 1–3 and anterior portion of stria 8 slightly impressed, punctate, external striae finer, 6–7 sometimes indistinct, stria 8 distinctly deepened before apex; parascutellar stria very short, deep, isolated. Recurrent preapical stria short, deep, with outer curvature directed towards the fifth stria. Parascutellar seta present; three discal setae located in the 3rd interval, adjoined to the 3rd stria ( +Fig. 1A +): anterior seta located near the end of the anterior elytral 7th; middle seta located anterior of elytral middle, near the end of elytral 3/7; posterior seta (= subapical seta) located about 1/8 of elytral length from elytral apex; seta of the recurrent stria isolated, removed from this stria by distance of the diameter of the pore ( +Fig. 1A +). Number and positions of the setae of the marginal umbilicate series typical of + +Trechus + +s. l. +( +Fig. 1A +). Metepisternum very short, glabrous and smooth, with outer margin about as long as anterior margin. + + +Legs: Moderately short and robust. Protibia dilated towards apex, straight on external margin, its dorsal surface with very flat longitudinal groove without micro-setae on anterior surface. Two basal protarsomeres of males dilated and dentoid at the inner apical border ( +Fig. 1C +). + + +Male genitalia: EL/AL= 2.00–2.23 (Ø = 2.11; n = 8). Aedeagal median lobe robust, in lateral view markedly bent in basal third, almost straight towards apex, with ventral margin slightly convex in middle, ventrally bent towards apex, the latter with a small upwardly bent hook ( +Fig. 1G +); in dorsal view broad in basal half, sinusoidal, narrowed from middle to apex ( +Fig. 1H +). Dorsal surface of median lobe not sclerotized apart from the basal quarter of the basal bulb near the basal opening which is slightly sclerotized; sagittal aileron absent. Endophallus symmetrical in dorsal view, without distinct copulatory pieces, densely covered by rows of tiny scales along dorsal side of the inactive internal sac from its base to apex, and with a complex folding structure and large sclerotized scales near base of internal sac. Parameres similar in size and shape, moderately short, each with three or four apical setae. + + + + +Differential diagnosis +. + +Trechus apertus + +sp. n. +is distinguished from all previously described + +Trechus + +(s. l.) species of +Ethiopia +, by the aedeagal median lobe ostium which extends basally across three quarters of the basal bulb. Not any Ethiopian Trechina species was known so far that has a dorsal opening that includes parts of the basal bulb. In external characters, the new species differs additionally from all the small to medium-sized trechine species from Mt. Choke as follows: from + +Trechus abyssinicus +(Quéinnec & Ollivier) + +, + +T. afroalpinus +(Quéinnec & Ollivier) + +, + +T. amharicus +Ortuño & Novoa + +and + +T. lobeliae +(Quéinnec & Ollivier) + +in two basal protarsomeres dilated in males (one in all the latter species), from + +T. amharicus + +additionally in the presence of the middle seta of the elytral discal series (absent in + +T. amharicus + +), from + +T. lobeliae + +additionally in the smaller body size and the almost completely rounded pronotal laterobasal angles (rectangular in + +T. lobelia + +), and from + +T. abyssinicus + +, + +T. afroalpinus + +, + +T. igori + +sp. n. +, + +T. kniphofia + +sp. n. +, + +T. reebae +(Quéinnec & Ollivier) + +and + +T. yitbareki + +sp. n. +in the more deeply impressed and distinctly punctate elytral striae (finely impressed and smooth or indistinctly punctate in all the latter species). In habitus, + +T. apertus + +sp. n. +is very similar to + +T. niloticus +Quéinnec & Ollivier + +which was described from the western crater rim of Mt. Choke. However, in + +T. niloticus + +the dorsal opening of the aedeagal median lobe do not reach the basal bulb (see + +Quéinnec +et al. +2021 + +: Fig. 26B, C). In addition, in + +T. niloticus + +, the median lobe is more elongated, with ventral margin straight in middle (convex in + +T. apertus + +sp. n. +), its apical lamella is much longer, markedly bent ventrally (short and slightly bent ventrally in + +T. apertus + +sp. n. +), the parameres are slenderer. + + + + +Distribution +. Endemic to Mt. Choke in northern +Ethiopia +. The new species was only found near the top of a mountain ridge that extends from the crater rim to the west ( +Fig. 4 +). + + +Habitat preferences +. All specimens of + +T. apertus + +sp. n. +were sifted from the humus-rich soil under dense vegetation alongside a small brook. The brook has its source near the top of the mountain ridge and flows to the north, at an elevation of +3370–3450 m +. No specimens were found on the pastures and in the bushland which bordered the brook. Based on these collecting circumstances, the new species can be considered hygrophilic and cold-adapted. As potentially natural vegetation, a moist afromontane forest would have formed at the site where the new species was found, but here it has given way to a cultivated steppe. + + + + \ No newline at end of file diff --git a/data/BE/17/A3/BE17A334FF91192588DBFB53FDA8F938.xml b/data/BE/17/A3/BE17A334FF91192588DBFB53FDA8F938.xml new file mode 100644 index 00000000000..a67354767c0 --- /dev/null +++ b/data/BE/17/A3/BE17A334FF91192588DBFB53FDA8F938.xml @@ -0,0 +1,131 @@ + + + +Description of four new species of Lamiinae (Coleoptera: Cerambycidae) from Panama + + + +Author + +Lanuza-Garay, Alfredo +Universidad de Panamá, Centro Regional Universitario de Colón, Escuela de Biología, Departamento de Zoología, Cristobal County, Randolph Avenue, Rainbow City, Colon City, PANAMA & Museo de Invertebrados Graham Bell Fairchild, Universidad de Panamá, Manuel Espinosa Batista Avenue, Panama City, PANAMA. + + + +Author + +Rivera, Marleny +Smithsonian Tropical Research Institute, Ancon, Panama City, PANAMA + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +440 +448 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.10 + +journal article +10.11646/zootaxa.5492.3.10 +1175-5326 +13234786 +170F815F-A836-462F-A99B-7F1066CFE1AB + + + + + + + +Piriana aielloae + +sp. nov. + + + + + + +( +Figs. 21–25 +) + + + +Holotype +female, +Panama +, +Panama province +, +Cerro Campana +, + +26.xi.1977 + +, +H. P. Stockwell +(STRI_ENT_0063016). + + + + +Description. +Holotype +male. + +Integument mostly dark brown; head, gulamentum, mouthparts, and apex of tarsal claws reddish brown; maxillary and labial palpi and antennomere IV yellowish, dark brown toward apex; peduncle of pro- and mesofemora and base of tarsal claws light-brown; prothorax reddish brown dorsally, dark brown ventrally; pronotum orangish with wide longitudinal dark-brown area centrally. Scutellum dark brown. Elytra with anterior 2/3 orangish except brownish wide longitudinal band close to suture. + + +Head. +Frons transverse, slightly convex anteriorly, finely punctate with yellowish pubescence not obscuring integument; area between antennal tubercles with wide yellowish pubescent band on each side of median groove; eyes deeply emarginate; distance between upper eye lobes 0.3 times distance between outer margins of eyes; frontally, distance between lower eye lobes 0.6 times distance between outer margins of eyes; space between upper eye lobe and base of antenna with yellowish pubescence. Genae 0.75 times shorter than lower eye lobe. Central area of postclypeus with a pair of short, erect black setae, and one long, erect black seta on each side; sides of postclypeus with one short erect black setae. Anterior margin of labrum with short yellow setae. Gulamentum smooth and glabrous. Antennae slender, reaching elytral apex at base of antennomere VI; finely punctate, dorsally covered with grayish pubescence not obscuring integument; scape elongated and cylindrical, slightly expanded apically, reaching mesocoxae in lateral view; scape and antennomere III with long erect black setae ventrally; antennomere IV with sparse, long, erect black setae ventrally; antennomeres V–VI with black setae apically. Antennal formula based on length of antennomere III:—Scape = 1.20;—Pedicel = 0.07;—IV = 0.87;—V = 0.72;—VI = 0.47;—VII = 0.30;—VIII = 0.70;—IX = 0.27;—X = 0.12;—XI = 0.20. + + + +FIGURES 21–25. + +Piriana aielloae + +, + +sp. nov. + +, holotype female: 19. dorsal habitus; 20. ventral habitus; 21. lateral habitus; 22. head, frontal view; 23. Close-up of pro legs + + + +Thorax. +Prothorax 0.75 times wider than long, lateral tubercles feebly developed, obtuse. Pronotum moderately convex on disc, depressed posteriorly; with sparse whitish pubescence; finely punctate, punctures abundant, coarser posteriorly. Prosternum with grayish pubescence not obscuring integument; prosternal process narrow, not laminiform, expanded apically, centrally 0.2 times procoxal width. Mesanepisternum and mesepimeron with sparse grayish pubescence not obscuring integument; mesoventral process narrow, not laminiform, centrally 0.16 times mesocoxal width; metanepisternum with sparse grayish pubescence; meso and metaventrite with sparse grayish pubescence not obscuring the integument; metathoracic discrimen reaching middle of metaventrite. + + +Elytra. +Coarsely punctate on anterior half and laterally on anterior fifth, punctures gradually sparser toward posterior region; light area with dense orangish pubescence and dark area with sparse grayish pubescence, except area close to suture on anterior 2/3 with orangish pubescence partially obscuring integument; with abundant, long, erect dark setae; humeral carina feebly marked, extending from humeri to beginning of posterior third; apex emarginate, sutural and outer angle feebly acute. + + +Legs. +Covered with grayish pubescence not obscuring integument; meso- and metatibiae with a few long, erect black setae from basal quarter ventrally and from middle dorsally. Protibiae curved, armed with one pre-medial rounded tubercle. Protarsomere I about as long as II–III together; meso- and metatarsomere I twice length of II–III together. + + +Abdomen. +Ventral surface with grayish pubescence not obscuring integument. Ventrite 5 not extending beyond elytral apices, sides and apex rounded. + + + + \ No newline at end of file diff --git a/data/BE/17/A3/BE17A334FF93192488DBF8FEFD25FEF2.xml b/data/BE/17/A3/BE17A334FF93192488DBF8FEFD25FEF2.xml new file mode 100644 index 00000000000..8ed16a0446b --- /dev/null +++ b/data/BE/17/A3/BE17A334FF93192488DBF8FEFD25FEF2.xml @@ -0,0 +1,135 @@ + + + +Description of four new species of Lamiinae (Coleoptera: Cerambycidae) from Panama + + + +Author + +Lanuza-Garay, Alfredo +Universidad de Panamá, Centro Regional Universitario de Colón, Escuela de Biología, Departamento de Zoología, Cristobal County, Randolph Avenue, Rainbow City, Colon City, PANAMA & Museo de Invertebrados Graham Bell Fairchild, Universidad de Panamá, Manuel Espinosa Batista Avenue, Panama City, PANAMA. + + + +Author + +Rivera, Marleny +Smithsonian Tropical Research Institute, Ancon, Panama City, PANAMA + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +440 +448 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.10 + +journal article +10.11646/zootaxa.5492.3.10 +1175-5326 +13234786 +170F815F-A836-462F-A99B-7F1066CFE1AB + + + + + + + +Pseudosparna carmenae + +sp. nov. + + + + + + +( +Figs. 15–20 +) + + + +Holotype +male, +Panama +, +Colon province +, +Pipeline Road +, + +20.vi.1982 + +, +H. P. Stockwell +col. (STRI_ENT_0063017). + + + + +FIGURES 15–20. + +Pseudosparna carmenae +, + + +sp. nov. + +, holotype male: 15. dorsal habitus; 16. ventral habitus; 17. lateral habitus; 18. head, frontal view. 19. Ventral view showing prosternal and mesoventral processes; + +Pseudosparna dimitrisi + +male: Ventral view showing prosternal and mesoventral processes + + + + +Description. +Holotype +male. + +Integument mostly dark brown, nearly black on some areas; frons (except dark brown area under antennal tubercles), genae, and prothoracic tubercles reddish brown; mouthparts, coxae, and base and ventral surface of femora light brown; antennomere IV light brown, dark brown toward apex; gulamentum yellowish; prothorax reddish brown except lateral tubercles, which are light brown; pronotum orangish with a central black band. Scutellum dark brown. Elytra with anterior half orangish except blackish wide longitudinal band close to suture and lateral margin. + + +Head. +Frons transverse, slightly convex anteriorly; finely punctate, with yellowish pubescence covering surface, pubescence denser laterally and close to median groove, which is deep, narrow and reaching clypeus. Eyes deeply emarginate; distance between upper eye lobes 0.3 times distance between outer margins of eyes; frontally, distance between lower eye lobes 0.6 times distance between outer margins of eyes. Area between upper eye lobe and base of antenna with yellowish pubescence. Genae 1.3 times longer than lower eye lobe. Gulamentum smooth and glabrous. Dorsal sides of occiput with a patch of yellowish pubescence not obscuring integument. Surface of antennal tubercles with sparse grayish pubescence not obscuring integument. Antennae slender, reaching elytral apices at base of antennomere VIII, finely punctate; scape elongated and cylindrical, slightly expanded apically, reaching mesocoxae in lateral view; with long black setae ventrally, antennomere III with moderately long black setae ventrally; IV slightly longer than V; V–XI each similar in size and pubescence. Antennal formula based on length of antennomere III: –Scape = 1.57;—Pedicel = 0.08;—IV = 0.97;—V = 0.82;—VI = 0.80;—VII = 0.82;— VIII = 0.91;—IX = 1.00;—X = 1.10;—XI = 0.60. + + +Thorax. +Prothorax 1.2 times wider than long; lateral tubercles short and acute, with a shallow dorsal depression basally. Pronotum moderately convex anteriorly, depressed posteriorly; finely punctate, punctures more abundant and coarser posteriorly. Prosternum with moderately dense pale setae; prosternal process narrow, not laminiform, expanded apically, narrowest area 0.22 times procoxal width; mesanepisternum and mesepimeron with sparse grayish pubescence not obscuring integument; mesoventral process truncate apically, narrowest area 0.1 times mesocoxal width; metanepisternum with sparse grayish pubescence not obscuring integument; meso and metaventrites with sparse grayish pubescence not obscuring the integument; metathoracic discrimen complete. + + +Elytra. +Abundantly, coarsely, punctate; apex emarginate, sinuous toward rounded sutural angle, outer angle spiniform;with long,erect,sparse black setae throughout.On elytral anterior half, integument obscured by dense orangish pubescence, centroapical area and suture covered with sparse grayish pubescence, not obscuring integument. + + +Legs. +With abundant yellowish pubescence not obscuring integument; mesotibiae with short, erect black setae from basal quarter ventrally and from middle dorsally, setae distinctly denser about posterior third, metatibiae with short, erect, black setae from basal quarter ventrally and from base dorsally to posterior third. Metatarsomere I slightly longer than twice length of II–III together. + + +Abdomen. +Ventrites with grayish pubescence not obscuring integument; intersegment of ventrites 1 and 2 with yellowish pubescence; apex of ventrite 5 concave. + + + + \ No newline at end of file diff --git a/data/BE/17/A3/BE17A334FF94192688DBFD5BFEE3FBCF.xml b/data/BE/17/A3/BE17A334FF94192688DBFD5BFEE3FBCF.xml new file mode 100644 index 00000000000..149196aa7d7 --- /dev/null +++ b/data/BE/17/A3/BE17A334FF94192688DBFD5BFEE3FBCF.xml @@ -0,0 +1,141 @@ + + + +Description of four new species of Lamiinae (Coleoptera: Cerambycidae) from Panama + + + +Author + +Lanuza-Garay, Alfredo +Universidad de Panamá, Centro Regional Universitario de Colón, Escuela de Biología, Departamento de Zoología, Cristobal County, Randolph Avenue, Rainbow City, Colon City, PANAMA & Museo de Invertebrados Graham Bell Fairchild, Universidad de Panamá, Manuel Espinosa Batista Avenue, Panama City, PANAMA. + + + +Author + +Rivera, Marleny +Smithsonian Tropical Research Institute, Ancon, Panama City, PANAMA + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +440 +448 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.10 + +journal article +10.11646/zootaxa.5492.3.10 +1175-5326 +13234786 +170F815F-A836-462F-A99B-7F1066CFE1AB + + + + + + + +Trichotithonus barrocoloradensis + +sp. nov. + + + + + + +( +Figs. 7–10 +) + + + +Holotype +female, +Panama +, +Panama province +, +Barro Colorado Island +, + +8.x.1977 + +, +H. Wolda +col. (STRI_ENT_0030820). + +Description + + + +. +Holotype +female + +. Integument mostly brownish; scape and pedicel pale brown; pro- and mesocoxae, gulamentum, and mouthparts yellowish brown; legs and antennomeres III–XI yellowish brown basally, then progressively brown to dark brown toward apex. Pronotum with six whitish maculae, four on posterior third and two laterally on anterior third, descending obliquely from anterolateral angles to central disc. Elytra with pre-medial, rounded whitish macula laterally. + + + +FIGURES 7–14. + +Trichotithonus +spp. 7 + +–10. + +Trichotithonus barrocoloradensis +, + + +sp. nov. + +, holotype female: 7. dorsal habitus; 8. ventral habitus; 9. lateral habitus; 10. head, frontal view; 11–14. + +Trichotithonus giuglarisi + +, holotype female: 11. head, frontal view; 12. lateral habitus; 13. dorsal habitus; 14. ventral habitus + + + +Head. +Frons with yellowish pubescence not obscuring integument, pubescence denser close to eyes. Central area between antennal tubercles with grayish pubescent band, nearly obscuring integument. Distance between upper eyes lobes 0.21 times distance between outer margins of eyes; frontally, distance between lower eye lobes 0.50 times distance between outer margins of eyes. Genae 0.67 times shorter than lower eye lobes. Gulamentum smooth and glabrous. Antennae slender, reaching elytral apices at base of antennomere VI; with a few long, erect setae on ventral side of antennomeres, from posterior area of scape to antennomere IV, then only apically in the remaining antennomeres. Scape elongate and cylindrical, finely punctate. Antennal formula based on length of antennomere III: –Scape = 1.10;—Pedicel = 0.12;—IV = 1.00;—V = 1.00;—VI = 0.97;—VII = 0.93;—VIII = 0.87;—IX = 0.81;—X = 0.81;—XI = 0.50. + + +Thorax. +Prothorax 1.2 times wider than long; lateral tubercles acute, obliquely directed backward, located on posterior third. Maculae on pronotum densely whitish pubescent, remaining pronotal surface covered with sparse whitish pubescence not obscuring integument; center of pronotum with a small elevation on each side of anterior half and another centrally; punctation quite coarse near basal margin. Prosternal process laminiform centrally; margins of mesanepisternum and mesepimeron with dense grayish pubescence nearly obscuring integument, remaining surface covered with sparse grayish pubescence not obscuring integument; mesoventral process narrow, 0.13 times mesocoxal width; anterior third of metanepisternum with dense grayish pubescence, remaining surface with sparse grayish pubescence not obscuring integument, meso and metaventrite with grayish pubescence not obscuring the integument. Scutellum sub-rounded apically, with whitish pubescence not obscuring integument. + + +Elytra. +Densely, coarsely punctate on anterior half, then finely punctate on posterior half; surface with many long, erect black setae; centrobasal crest with a small tuft of black setae directed backward; with dark-brown pubescence centrolaterally and dense grayish pubescence surrounding suture obscuring integument; remaining elytral surface covered with sparse whitish pubescence, not obscuring integument; apex obliquely emarginate, outer angle extended into a long spine. + + +Legs. +Femora and tibiae with long erect setae. Length of metatarsomere I subequal to length of metatarsomere II–III together. + + +Abdomen. +Ventrites with whitish pubescence not obscuring integument; ventrite 5 concave with outer angles spiniform. + + + + \ No newline at end of file diff --git a/data/BE/17/A3/BE17A334FF96192088DBFC2FFD55F802.xml b/data/BE/17/A3/BE17A334FF96192088DBFC2FFD55F802.xml new file mode 100644 index 00000000000..a82c6baf6f1 --- /dev/null +++ b/data/BE/17/A3/BE17A334FF96192088DBFC2FFD55F802.xml @@ -0,0 +1,132 @@ + + + +Description of four new species of Lamiinae (Coleoptera: Cerambycidae) from Panama + + + +Author + +Lanuza-Garay, Alfredo +Universidad de Panamá, Centro Regional Universitario de Colón, Escuela de Biología, Departamento de Zoología, Cristobal County, Randolph Avenue, Rainbow City, Colon City, PANAMA & Museo de Invertebrados Graham Bell Fairchild, Universidad de Panamá, Manuel Espinosa Batista Avenue, Panama City, PANAMA. + + + +Author + +Rivera, Marleny +Smithsonian Tropical Research Institute, Ancon, Panama City, PANAMA + +text + + +Zootaxa + + +2024 + +2024-08-06 + + +5492 + + +3 + + +440 +448 + + + + +http://dx.doi.org/10.11646/zootaxa.5492.3.10 + +journal article +10.11646/zootaxa.5492.3.10 +1175-5326 +13234786 +170F815F-A836-462F-A99B-7F1066CFE1AB + + + + + + + +Sternacutus akersae + +sp. nov. + + + + + + +( +Figs. 1–6 +) + + + +Holotype +female, +Panama +, +San Blas region +, +Nusagandi +, + +9–11.xi.1991 + +, +M. Akers +col. (MACO). + + + + +FIGURES 1–6. + +Sternacutus akersae + +, + +sp. nov. + +, holotype female: 1. dorsal habitus; 2. Close-up of pronotum and anterior third of pronotum; 3. head, frontal view; 4. ventral habitus showing prosternal and mesoventral process; 5. abdomen; 6. lateral habitus. + + + + +Description. +Holotype +female + +. Integument mostly dark brown; scape pale brown; antennomeres III–VI yellowish brown except dark-brown apex; antennomere VIII yellowish on basal third, dark-brown on remaining surface; + +antennomeres VII, IX–XI dark brown; mouthparts yellowish brown; mandibles dark brown. Pronotum with two rounded blackish maculae centrally and one moderate and elongated blackish macula near base of lateral tubercles of prothorax. Elytral anterior third dark brown; premedial to posterior third yellowish with two transverse, irregular dark-brown macula, one postmedial, another anteapical; with sparse dark spots along humeral carina, between postmedial and apical maculae, and along elytral suture; epipleura with dark-brown macula basally. Coxae pale brown; femora yellowish brown basally, then progressively brown to dark brown toward apex; tibial medial third yellowish, basal and apical thirds dark brown; tarsomeres I yellowish, rest of tarsomeres dark brown. + +Head. +Frons abundantly, minutely punctate; with sparse pale-yellowish pubescence. Area between antennal tubercles with sparse yellowish pubescence not obscuring integument; area between antennal tubercles and upper eye lobes with yellowish pubescence obscuring integument. Distance between upper eyes lobes 0.15 times distance between outer margins of eyes; frontally, distance between lower eye lobes 0.44 times distance between outer margins of eyes. Genae 0.68 times shorter than lower eye lobe, with dense yellowish pubescence.Antennal tubercles with abundant pale-yellow pubescence frontally, sparse yellowish-brown pubescence dorsally, except apex with dense yellow pubescence, and abundant yellow pubescence posteriorly. Clypeus with one long, erect brownish seta on each side, clypeal surface near fronto-clypeal suture with short yellowish setae. Gulamentum smooth, glabrous. Antennae slender, reaching elytral apex at base of antennomere VII. Scape slightly sinuous ventrally; with abundant yellowish-white pubescence not obscuring integument, with a few long, erect brown setae interspersed near apex of ventral surface. Pedicel with abundant yellowish pubescence not obscuring integument and one long, erect yellowish-brown seta ventrally; antennomeres III–VIII with abundant yellowish-white pubescence on light area, sparse on dark area; antennomeres VII, IX–XI with sparse yellowish-white pubescence; antennomeres III and IV with short, erect, thick black setae ventrally; antennomere V with two short, thick black setae ventrally; antennomeres VI and VIII with short, thick black seta ventrally. Antennal formula based on length of antennomere III:—Scape = 0.95.—Pedicel = 0.10.—IV = 0.88.—V = 0.75.—VI = 0.55.—VII = 0.40.—VIII = 0.55.—IX = 0.65.—X = 0.60.—XI = 0.60. + + +Thorax. +Prothorax 1.75 times wider than long; lateral tubercles short, with acute apex directed backward. Pronotum with yellowish pubescence not obscuring integument; coarsely punctate, punctures abundant on central disc and posteriorly; central disc maculae with sparse yellowish pubescence not obscuring the integument, each one of them surrounded by yellowish pubescence not obscuring the integument. Sides of prothorax with abundant pale-yellow pubescence. Prosternum with pale-yellow pubescence, distinctly sparser on wide central region. Prosternal process narrow, not laminiform, expanded apically, narrowest area 0.23 times procoxal width. Margins of mesanepisternum and mesepimeron with dense grayish pubescence, remaining surface with sparse grayish pubescence not obscuring integument; mesoventral process not laminiform, narrowest area 0.37 times mesocoxal width; metanepisternum and metaventrite with dense yellow pubescence, pubescence paler and distinctly sparser on wide central region of metaventrite. Scutellum with abundant yellowish-brown pubescence not obscuring integument. + + +Elytra. +Elytral surface abundantly, coarsely punctate on anterior half, punctures gradually finer, sparser toward apex on posterior half; centrobasal crest elevated, directed backward; apex of elytron emarginated with outer angle triangularly acuminate; humeral carina moderately well marked from humerus to posterior quarter; dorsal surface with two moderately well-marked carinae, outermost oblique, from near humeri to posterior third, innermost straight, from apex of centrobasal crest to posterior third; with abundant grayish pubescence partially obscuring integument, except dark areas with dark-brown pubescence not obscuring integument. + + +Legs. +Femora with abundant whitish pubescence not obscuring integument, pubescence slightly denser and whiter on some areas of femoral club, especially on meso- and metafemora, and dense and longer on apex. Tibiae with abundant yellowish pubescence not obscuring integument, except sparser pubescence on dorsal and lateral surfaces of posterior third, and dense, bristly yellowish-brown pubescence on posterior third of ventral surface; dorsal surface of posterior third of mesotibiae with abundant, short, thick, erect blackish setae; dorsal surface of posterior third of metatibiae with a few short, thick, erect dark-brown setae. Tarsomeres with abundant whitish pubescence dorsally not obscuring integument; metatarsomere I distinctly longer than II–III together. + + +Abdomen. +Ventrites with abundant yellowish and whitish pubescence not obscuring integument; apex of ventrite 5 slightly concave, with yellowish setae. + + + + \ No newline at end of file